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<document id="A33DC5F5324BB920B92240EF99F3CD8D" ID-DOI="10.11646/zootaxa.5481.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="12742760" ID-ZooBank="9ECB8E6A-5B6F-451F-AA95-7084B79AFE6B0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721033546162" checkinUser="plazi" docAuthor="Hájek, Jiří &amp; Sheth, Sayali D." docDate="2024" docId="03D187BD0622491CFF272632AB0DDBB8" docLanguage="en" docName="zootaxa.5481.1.7.pdf" docOrigin="Zootaxa 5481 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copelatus bezdeki Sheth, Ghate &amp; Hajek 2018" docType="treatment" docVersion="2" lastPageNumber="121" masterDocId="FFE8FFC50623491EFFB0213DA937DC4E" masterDocTitle="A review of the Oriental Copelatus latipes complex, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="130" masterPageNumber="119" pageNumber="120" updateTime="1721034136821" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D1198C11046CD01EB817AEB54F576548">A review of the Oriental Copelatus latipes complex, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="9C7FA642FFAE4C0C1B0C6963C76259B1">Hájek, Jiří</mods:namePart>
<mods:affiliation id="499B69D0E686D6637CA08C1AC76B38D1">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Poc ̌ ernice, Czech Republic</mods:affiliation>
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<mods:namePart id="008CA6FB27B308D7355BA0725696794D">Sheth, Sayali D.</mods:namePart>
<mods:affiliation id="726EDB0BB573ACC39D6F6B18767DF779">Department of Zoology, Prof. Ramkrishna More Arts, Commerce &amp; Science College, Akurdi, Pune - 411044, Maharashtra, India</mods:affiliation>
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<mods:date id="22BB0758D6940319F847717691F61C4B">2024</mods:date>
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<treatment id="03D187BD0622491CFF272632AB0DDBB8" ID-DOI="http://doi.org/10.5281/zenodo.12742789" ID-Zenodo-Dep="12742789" LSID="urn:lsid:plazi:treatment:03D187BD0622491CFF272632AB0DDBB8" httpUri="http://treatment.plazi.org/id/03D187BD0622491CFF272632AB0DDBB8" lastPageId="2" lastPageNumber="121" pageId="1" pageNumber="120">
<subSubSection id="C36265200622491FFF272632ABEFDB67" box="[151,728,1806,1834]" pageId="1" pageNumber="120" type="nomenclature">
<paragraph id="8BC736AB0622491FFF272632ABEFDB67" blockId="1.[151,728,1806,1869]" box="[151,728,1806,1834]" pageId="1" pageNumber="120">
<heading id="D08F81C70622491FFF272632ABEFDB67" bold="true" box="[151,728,1806,1834]" fontSize="11" level="1" pageId="1" pageNumber="120" reason="1">
<taxonomicName id="4C784D280622491FFF272632ABEFDB67" ID-CoL="8LZNR" authority="Sheth, Ghate &amp; Hajek, 2018" authorityName="Sheth, Ghate &amp; Hajek" authorityYear="2018" box="[151,728,1806,1834]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="bezdeki">
<emphasis id="B90CEAB90622491FFF272632ABEFDB67" bold="true" box="[151,728,1806,1834]" pageId="1" pageNumber="120">
<emphasis id="B90CEAB90622491FFF272632A846DB64" bold="true" box="[151,369,1807,1834]" italics="true" pageId="1" pageNumber="120">Copelatus bezdeki</emphasis>
<bibRefCitation id="EFE94B5A0622491FFEC82632ABEFDB67" author="Sheth, S. D. &amp; Ghate, H. V. &amp; Hajek, J." box="[376,728,1806,1833]" pageId="1" pageNumber="120" pagination="235 - 260" refId="ref7449" refString="Sheth, S. D., Ghate, H. V. &amp; Hajek, J. (2018) Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae). Zootaxa, 4459 (2), 235 - 260. https: // doi. org / 10.11646 / zootaxa. 4459.2.2" type="journal article" year="2018">Sheth, Ghate &amp; Hájek, 2018</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C36265200622491FFF27260EA810DB03" box="[151,295,1843,1869]" pageId="1" pageNumber="120" type="description">
<paragraph id="8BC736AB0622491FFF27260EA810DB03" blockId="1.[151,728,1806,1869]" box="[151,295,1843,1869]" pageId="1" pageNumber="120">
(
<figureCitation id="13432A2E0622491FFF2F260EA9D1DB03" box="[159,230,1843,1869]" captionStart="FIGURES 1–2" captionStartId="2.[152,270,1207,1231]" captionTargetBox="[202,1400,194,1163]" captionTargetId="figure-27@2.[170,1417,181,1183]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Copelatus latipes species complex. 1. C. bezdeki (holotype ♂; Maharashtra, India); 2. C. schereri (♂; Sri Lanka). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742762" httpUri="https://zenodo.org/record/12742762/files/figure.png" pageId="1" pageNumber="120">Figs 1</figureCitation>
,
<figureCitation id="13432A2E0622491FFF44260EA828DB03" box="[244,287,1843,1869]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="1" pageNumber="120">7–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36265200622491FFF272645AB87DBC1" box="[151,688,1911,1936]" pageId="1" pageNumber="120" type="reference_group">
<paragraph id="8BC736AB0622491FFF272645AB87DBC1" blockId="1.[151,688,1911,1936]" box="[151,688,1911,1936]" pageId="1" pageNumber="120">
<treatmentCitationGroup id="AB6811850622491FFF272645AB87DBC1" box="[151,688,1911,1936]" pageId="1" pageNumber="120">
<treatmentCitation id="0AD910BA0622491FFF272645AB9BDBC1" ID-DOI="http://doi.org/10.5281/zenodo.5966531" ID-Zenodo-Dep="5966531" author="Sheth, S. D. &amp; Ghate, H. V. &amp; Hajek, J." box="[151,684,1911,1936]" httpUri="http://treatment.plazi.org/id/03BC3747FF91FF9FFF5E1752FE66FC1A" page="245" pageId="1" pageNumber="120" year="2018">
<taxonomicName id="4C784D280622491FFF272645AB9BDBC1" ID-CoL="8LZNR" authority="Sheth, Ghate &amp; Hajek, 2018: 245" authorityName="Sheth, Ghate &amp; Hajek" authorityPageNumber="245" authorityYear="2018" box="[151,684,1911,1936]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="bezdeki">
<emphasis id="B90CEAB90622491FFF272645A867DBDE" box="[151,336,1912,1936]" italics="true" pageId="1" pageNumber="120">Copelatus bezdeki</emphasis>
<bibRefCitation id="EFE94B5A0622491FFEE72645AB9BDBC1" author="Sheth, S. D. &amp; Ghate, H. V. &amp; Hajek, J." box="[343,684,1911,1936]" pageId="1" pageNumber="120" pagination="235 - 260" refId="ref7449" refString="Sheth, S. D., Ghate, H. V. &amp; Hajek, J. (2018) Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae). Zootaxa, 4459 (2), 235 - 260. https: // doi. org / 10.11646 / zootaxa. 4459.2.2" type="journal article" year="2018">Sheth, Ghate &amp; Hájek, 2018: 245</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36265200622491CFF27268AAB06D9D7" lastPageId="2" lastPageNumber="121" pageId="1" pageNumber="120" type="materials_examined">
<paragraph id="8BC736AB0622491FFF27268AAB37DBBB" blockId="1.[151,1436,1975,2038]" pageId="1" pageNumber="120">
<materialsCitation id="3B103CF60622491FFF27268AAB37DBBB" country="India" elevation="640" location="Bhushi" municipality="Pune district" pageId="1" pageNumber="120" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
<emphasis id="B90CEAB90622491FFF27268AA805DB9F" bold="true" box="[151,306,1975,2001]" pageId="1" pageNumber="120">
<typeStatus id="54C388090622491FFF27268AA9E5DB9F" box="[151,210,1975,2001]" pageId="1" pageNumber="120">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F36F763B0622491FFEF3268AA848DB9F" box="[323,383,1975,2001]" name="India" pageId="1" pageNumber="120">India</collectingCountry>
,
<collectingRegion id="49BCF8490622491FFE3D268AAB2BDB9F" box="[397,540,1975,2001]" country="India" name="Maharashtra" pageId="1" pageNumber="120">Maharashtra</collectingRegion>
,
<collectingMunicipality id="6BA3ACD10622491FFD9A2685AB8BDB9F" box="[554,700,1975,2002]" pageId="1" pageNumber="120">Pune district</collectingMunicipality>
, ca.
<locationDeviation id="9A752F890622491FFD43268AAD6BDB9F" box="[755,1116,1975,2001]" location="Lonavala village" pageId="1" pageNumber="120">
<quantity id="4C809B4E0622491FFD43268AAA19DB9F" box="[755,814,1975,2001]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="120" unit="km" value="4.0">4 km</quantity>
SSW of Lonavala village
</locationDeviation>
,
<location id="8EA760700622491FFBDA268AAD8EDB9F" LSID="urn:lsid:plazi:treatment:03D187BD0622491CFF272632AB0DDBB8:8EA760700622491FFBDA268AAD8EDB9F" box="[1130,1209,1975,2001]" country="India" municipality="Pune district" name="Bhushi" pageId="1" pageNumber="120" stateProvince="Maharashtra">Bhushi</location>
dam, 18°43.2–4’N, 73°23.7–24.0’E, ca.
<elevation id="0055D1980622491FFECE26E6AB37DBBB" box="[382,512,2011,2038]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="120" unit="m" value="640.0">
<quantity id="4C809B4E0622491FFECE26E6A8F2DBB8" box="[382,453,2011,2038]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="120" unit="m" value="640.0">640 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
<caption id="DF0766230621491CFF28258AA8E9D8BD" ID-DOI="http://doi.org/10.5281/zenodo.12742762" ID-Zenodo-Dep="12742762" httpUri="https://zenodo.org/record/12742762/files/figure.png" pageId="2" pageNumber="121" startId="2.[152,270,1207,1231]" targetBox="[202,1400,194,1163]" targetPageId="2" targetType="figure">
<paragraph id="8BC736AB0621491CFF28258AA8E9D8BD" blockId="2.[151,1436,1207,1267]" pageId="2" pageNumber="121">
<emphasis id="B90CEAB90621491CFF28258AA876D881" bold="true" box="[152,321,1207,1231]" pageId="2" pageNumber="121">FIGURES 1–2.</emphasis>
Habitus of
<taxonomicName id="4C784D280621491CFE0A258AAB5BD881" authorityName="Sharp" authorityYear="1882" box="[442,620,1207,1231]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B90CEAB90621491CFE0A258AAB5BD881" box="[442,620,1207,1231]" italics="true" pageId="2" pageNumber="121">Copelatus latipes</emphasis>
</taxonomicName>
species complex. 1.
<taxonomicName id="4C784D280621491CFCF12585AA9CD881" authorityName="Sheth, Ghate &amp; Hajek" authorityYear="2018" box="[833,939,1207,1231]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="bezdeki">
<emphasis id="B90CEAB90621491CFCF12585AA9CD881" box="[833,939,1207,1231]" italics="true" pageId="2" pageNumber="121">C. bezdeki</emphasis>
</taxonomicName>
(holotype ♂; Maharashtra, India); 2.
<taxonomicName id="4C784D280621491CFA9C2585ACABD881" authorityName="Wewalka" authorityYear="1981" box="[1324,1436,1207,1231]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="schereri">
<emphasis id="B90CEAB90621491CFA9C2585ACABD881" box="[1324,1436,1207,1231]" italics="true" pageId="2" pageNumber="121">C. schereri</emphasis>
</taxonomicName>
(♂; Sri Lanka). Scale bar 2 mm.
</paragraph>
</caption>
<paragraph id="8BC736AB0621491CFF772405AB8BD91D" blockId="2.[151,1437,1336,2038]" box="[199,700,1336,1363]" pageId="2" pageNumber="121">
<emphasis id="B90CEAB90621491CFF772405A859D91C" bold="true" box="[199,366,1336,1362]" pageId="2" pageNumber="121">
<typeStatus id="54C388090621491CFF772405A835D91C" box="[199,258,1336,1362]" pageId="2" pageNumber="121">Type</typeStatus>
material
</emphasis>
. See
<bibRefCitation id="EFE94B5A0621491CFE1B2405AB80D91D" author="Sheth, S. D. &amp; Ghate, H. V. &amp; Hajek, J." box="[427,695,1336,1363]" pageId="2" pageNumber="121" pagination="235 - 260" refId="ref7449" refString="Sheth, S. D., Ghate, H. V. &amp; Hajek, J. (2018) Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae). Zootaxa, 4459 (2), 235 - 260. https: // doi. org / 10.11646 / zootaxa. 4459.2.2" type="journal article" year="2018">
Sheth
<emphasis id="B90CEAB90621491CFE422404AB1DD91C" box="[498,554,1336,1362]" italics="true" pageId="2" pageNumber="121">et al.</emphasis>
(2018: 245)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BC736AB0621491CFF772466AB06D9D7" blockId="2.[151,1437,1336,2038]" pageId="2" pageNumber="121">
<emphasis id="B90CEAB90621491CFF772466AB01D93B" bold="true" box="[199,566,1371,1397]" pageId="2" pageNumber="121">Additional material examined.</emphasis>
<materialsCitation id="3B103CF60621491CFD8D2466AB1AD9D7" collectingDate="1991-09-13" collectionCode="NHMW" collectorName="R. Schuh" country="India" location="Lonvala" pageId="2" pageNumber="121" specimenCount="11" specimenCount-female="5" specimenCount-male="6" stateProvince="Maharashtra">
<emphasis id="B90CEAB90621491CFD8D2466AA7ED938" bold="true" box="[573,841,1371,1398]" pageId="2" pageNumber="121">
<collectingCountry id="F36F763B0621491CFD8D2466ABA3D93B" box="[573,660,1371,1397]" name="India" pageId="2" pageNumber="121">INDIA</collectingCountry>
:
<collectingRegion id="49BCF8490621491CFD112461AA73D938" box="[673,836,1372,1398]" country="India" name="Maharashtra" pageId="2" pageNumber="121">Maharashtra</collectingRegion>
:
</emphasis>
<specimenCount id="9D7EFD220621491CFCE12466AA4AD938" box="[849,893,1371,1398]" count="6" pageId="2" pageNumber="121" type="male">6 ♂</specimenCount>
,
<specimenCount id="9D7EFD220621491CFC3A2466AA82D938" box="[906,949,1371,1398]" count="5" pageId="2" pageNumber="121" type="female">5 ♀</specimenCount>
,
<locationDeviation id="9A752F890621491CFC742461AD9CD938" box="[964,1195,1372,1398]" location="Bombay" pageId="2" pageNumber="121">
<quantity id="4C809B4E0621491CFC742461AD3BD938" box="[964,1036,1372,1398]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="121" unit="km" value="80.0">80 km</quantity>
E of Bombay
</locationDeviation>
,
<location id="8EA760700621491CFB072461AC24D938" LSID="urn:lsid:plazi:treatment:03D187BD0622491CFF272632AB0DDBB8:8EA760700621491CFB072461AC24D938" box="[1207,1299,1372,1398]" country="India" name="Lonvala" pageId="2" pageNumber="121" stateProvince="Maharashtra">Lonvala</location>
[Lonavala],
<date id="FFC6106B0621491CFF272442A826D9D7" box="[151,273,1407,1433]" pageId="2" pageNumber="121" value="1991-09-13">
<collectingDate id="EF82E9830621491CFF272442A826D9D7" box="[151,273,1407,1433]" pageId="2" pageNumber="121" value="1991-09-13">13.ix.1991</collectingDate>
</date>
,
<collectorName id="268D537D0621491CFEAD24BDA8B3D9D7" box="[285,388,1407,1434]" pageId="2" pageNumber="121">R. Schuh</collectorName>
leg. (
<collectionCode id="ED69AE6E0621491CFE732442AB11D9D7" box="[451,550,1407,1433]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="121" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36265200621491CFF77249FAA55DBE0" pageId="2" pageNumber="121" type="diagnosis">
<paragraph id="8BC736AB0621491CFF77249FAA55DBE0" blockId="2.[151,1437,1336,2038]" pageId="2" pageNumber="121">
<emphasis id="B90CEAB90621491CFF77249FA875D9F2" bold="true" box="[199,322,1442,1468]" pageId="2" pageNumber="121">Diagnosis.</emphasis>
TL:
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. Head dark brown, clypeus and anterior part of frons often paler, reddish; pronotum brown blackish, laterally broadly reddish; elytra yellowish orange, with blackish pattern consisting of elongate sutural spot extending laterally up to stria II in middle third of elytral length, small spot present subbasally between striae II–IV, and lines along elytral striae. Species variable in dorsal surface colouration: in one extreme, dark area on elytra reduced to single narrow oval spot along suture; in other extreme, sutural spot extended laterally and connected with subbasal spot, usually confluent to subbasal transverse band. Pronotum with coarse punctures laterally. Each elytron with six complete discal striae and one submarginal stria: stria 1 beginning more posteriorly than other striae, ending close to elytral apex; striae 2, 3 and 5 ending subapically; stria 6 shortest, ending at apical fourth; submarginal stria long, beginning at elytral mid-length and ending at apical fourth (
<figureCitation id="13432A2E0621491CFB032783ADC0DA96" box="[1203,1271,1726,1752]" captionStart="FIGURES 1–2" captionStartId="2.[152,270,1207,1231]" captionTargetBox="[202,1400,194,1163]" captionTargetId="figure-27@2.[170,1417,181,1183]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Copelatus latipes species complex. 1. C. bezdeki (holotype ♂; Maharashtra, India); 2. C. schereri (♂; Sri Lanka). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742762" httpUri="https://zenodo.org/record/12742762/files/figure.png" pageId="2" pageNumber="121">Fig. 1</figureCitation>
). Male longer spur on metatibia sinuate in apical third. Median lobe of aedeagus in lateral view (
<figureCitation id="13432A2E0621491CFB8527DFADB2DAB2" box="[1077,1157,1762,1788]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="2" pageNumber="121">Fig. 7a</figureCitation>
) broad in basal 3/5, then obtusely angled; apical 2/5 narrow, sinuous ventrally, almost straight dorsally until distinctly dorsally bent pointed apex; median lobe in 3/5 of its length with two distinct ‘teeth’ on ventral side—broad rectangular proximal one, and narrower acute distal one; a third, small, obtuse tooth visible in ventro-lateral view (
<figureCitation id="13432A2E0621491CFBDF2671ADF5DB28" box="[1135,1218,1868,1894]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="2" pageNumber="121">Fig. 7b</figureCitation>
). Parameres almost straight on inner margin (
<figureCitation id="13432A2E0621491CFE0C264DAB34DBC4" box="[444,515,1904,1930]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="2" pageNumber="121">Fig. 8</figureCitation>
). Female identical to male in habitus, short irregular strioles present laterally on pronotum and in middle third of elytra laterally from striae 3–4.
</paragraph>
</subSubSection>
<subSubSection id="C36265200621491CFF77268AAB0DDBB8" pageId="2" pageNumber="121" type="distribution">
<paragraph id="8BC736AB0621491CFF77268AAB0DDBB8" blockId="2.[151,1437,1336,2038]" pageId="2" pageNumber="121">
<emphasis id="B90CEAB90621491CFF77268AA856DB9F" bold="true" box="[199,353,1975,2001]" pageId="2" pageNumber="121">Distribution.</emphasis>
The species is so far known only from Pune district of
<collectingRegion id="49BCF8490621491CFC602685AD68DB9C" box="[976,1119,1976,2002]" country="India" name="Maharashtra" pageId="2" pageNumber="121">Maharashtra</collectingRegion>
,
<collectingCountry id="F36F763B0621491CFBDA2685AD91DB9C" box="[1130,1190,1976,2002]" name="India" pageId="2" pageNumber="121">India</collectingCountry>
, within the altitudinal range of
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a.s.l. (
<figureCitation id="13432A2E0621491CFE6926E1AB1BDBB8" box="[473,556,2012,2038]" captionStart="FIGURE 19" captionStartId="6.[152,255,1288,1312]" captionTargetBox="[156,1437,273,1261]" captionTargetId="figure-66@6.[151,1437,273,1265]" captionTargetPageId="6" captionText="FIGURE 19. Map of distribution of Copelatus latipes species complex." figureDoi="http://doi.org/10.5281/zenodo.12742768" httpUri="https://zenodo.org/record/12742768/files/figure.png" pageId="2" pageNumber="121">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

354
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@ -0,0 +1,354 @@
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<mods:title id="09393133FFE6429E3BF0C4B10BBB06AE">A review of the Oriental Copelatus latipes complex, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="CC4A0DAF32D5812D2AA591BC7D21AB51">Hájek, Jiří</mods:namePart>
<mods:affiliation id="95768F3C3A7CD4F5DEA86B8F4BDADDAB">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Poc ̌ ernice, Czech Republic</mods:affiliation>
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<mods:namePart id="1D6727D2F23CAAD0F469C3D3FC72E38A">Sheth, Sayali D.</mods:namePart>
<mods:affiliation id="BF975A2D8050ABF4B0FFA5AB85BE9B6E">Department of Zoology, Prof. Ramkrishna More Arts, Commerce &amp; Science College, Akurdi, Pune - 411044, Maharashtra, India</mods:affiliation>
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<paragraph id="8BC736AB06244919FF2727D7AB2BDB4B" blockId="7.[151,540,1770,1833]" box="[151,540,1770,1797]" pageId="7" pageNumber="126">
<heading id="D08F81C706244919FF2727D7AB2BDB4B" bold="true" box="[151,540,1770,1797]" fontSize="11" level="1" pageId="7" pageNumber="126" reason="1">
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<taxonomicName id="4C784D2806244919FF2727D7A880DB4B" authorityName="Hájek &amp; Sheth" authorityYear="2024" box="[151,439,1770,1797]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="lanxangensis" status="sp. nov.">
<emphasis id="B90CEAB906244919FF2727D7A880DB4B" bold="true" box="[151,439,1770,1797]" italics="true" pageId="7" pageNumber="126">Copelatus lanxangensis</emphasis>
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<taxonomicNameLabel id="A23F57C206244919FE0E27D6AB2BDB4B" box="[446,540,1771,1797]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C362652006244919FF272632A874DB67" box="[151,323,1806,1833]" pageId="7" pageNumber="126" type="description">
<paragraph id="8BC736AB06244919FF272632A874DB67" blockId="7.[151,540,1770,1833]" box="[151,323,1806,1833]" pageId="7" pageNumber="126">
(
<figureCitation id="13432A2E06244919FF2F2632A9DFDB67" box="[159,232,1806,1833]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="7" pageNumber="126">Figs 5</figureCitation>
,
<figureCitation id="13432A2E06244919FF442633A80CDB67" box="[244,315,1806,1833]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="7" pageNumber="126">15–16</figureCitation>
)
</paragraph>
</subSubSection>
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<typeStatus id="54C3880906244919FF27266AA9E5DB3F" box="[151,210,1879,1905]" pageId="7" pageNumber="126">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F36F763B06244919FE8C266AA842DB3F" box="[316,373,1879,1905]" name="Laos" pageId="7" pageNumber="126">Laos</collectingCountry>
,
<collectingRegion id="49BCF84906244919FE31266AABBCDB3F" box="[385,651,1879,1905]" country="Laos" name="Khammouan" pageId="7" pageNumber="126">Khammouane Province</collectingRegion>
,
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env.,
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.
</paragraph>
<paragraph id="8BC736AB06244919FF772646ACABDB93" blockId="7.[151,1436,1878,2013]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF772646A84DDBDA" bold="true" box="[199,378,1914,1941]" pageId="7" pageNumber="126">Type material.</emphasis>
<materialsCitation id="3B103CF606244919FE362646AA60DB93" collectingDate="2001-06-08" collectionCode="NMPC" collectorName="Sausa O." country="Laos" determinerName="Sheth" elevation="550" latitude="17.716667" location="Khammouan prov." longLatPrecision="1277" longitude="105.15" pageId="7" pageNumber="126" specimenCount="2" specimenCount-male="2" stateProvince="Khammouan" typeStatus="holotype">
<typeStatus id="54C3880906244919FE362646A8D9DBDB" box="[390,494,1915,1941]" pageId="7" pageNumber="126" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7EFD2206244919FE4A2646AB38DBDB" box="[506,527,1915,1941]" count="1" pageId="7" pageNumber="126" type="male">♂</specimenCount>
(
<collectionCode id="ED69AE6E06244919FD932646AB4FDBDB" box="[547,632,1915,1941]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="126">NMPC</collectionCode>
), labelled: “
<collectingCountry id="F36F763B06244919FCB72646AA64DBDB" box="[775,851,1915,1941]" name="Laos" pageId="7" pageNumber="126">LAOS</collectingCountry>
centr.22.5-
<date id="FFC6106B06244919FC662647AD0EDBDB" box="[982,1081,1914,1941]" pageId="7" pageNumber="126" value="2001-06-08">
<collectingDate id="EF82E98306244919FC662647AD0EDBDB" box="[982,1081,1914,1941]" pageId="7" pageNumber="126" value="2001-06-08">8.6.2001</collectingDate>
</date>
/
<collectingRegion id="49BCF84906244919FBE92646AC18DBDB" box="[1113,1327,1915,1941]" country="Laos" name="Khammouan" pageId="7" pageNumber="126">Khammouan prov.</collectingRegion>
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, / NAKAI env., /
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,
<geoCoordinate id="EE4C506C06244919FE4C26A2AB43DBF7" box="[508,628,1950,1977]" degrees="105" direction="east" minutes="09" orientation="longitude" pageId="7" pageNumber="126" precision="925" value="105.15">E 105°09’</geoCoordinate>
, / Jendek E. &amp;
<collectorName id="268D537D06244919FC9B269DAAA6DBF7" box="[811,913,1951,1979]" pageId="7" pageNumber="126">Šauša O.</collectorName>
leg. [p] //
<typeStatus id="54C3880906244919FBA126A2AD80DBF7" box="[1041,1207,1951,1977]" pageId="7" pageNumber="126" type="holotype">HOLOTYPUS</typeStatus>
<specimenCount id="9D7EFD2206244919FB7126A2ADE1DBF7" box="[1217,1238,1951,1977]" count="1" pageId="7" pageNumber="126" type="male">♂</specimenCount>
/
<taxonomicName id="4C784D2806244919FB4026A2ACBCDBF7" authorityName="Erichson" authorityYear="1832" box="[1264,1419,1951,1977]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB906244919FB4026A2ACBCDBF7" box="[1264,1419,1951,1977]" italics="true" pageId="7" pageNumber="126">COPELATUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C784D2806244919FF2726FEA81EDB93" authorityName="Hájek &amp; Sheth" authorityYear="2024" box="[151,297,1987,2013]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="lanxangensis" status="sp. nov.">
<emphasis id="B90CEAB906244919FF2726FEA81EDB93" box="[151,297,1987,2013]" italics="true" pageId="7" pageNumber="126">lanxangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57C206244919FE8026FEA8B0DB93" box="[304,391,1987,2013]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
/ Hájek &amp;
<determinerName id="7E1BDA6F06244919FDB226FEAB75DB93" box="[514,578,1987,2013]" pageId="7" pageNumber="126">Sheth</determinerName>
det. 2024 [red label, p]”
</materialsCitation>
.
<materialsCitation id="3B103CF606244919FCD226FEACABDB93" box="[866,1436,1987,2013]" collectingDate="2001-06-08" collectionCode="NMPC" collectorName="Sausa O." country="Laos" determinerName="Sheth" elevation="550" latitude="17.716667" location="Khammouan prov." longLatPrecision="1277" longitude="105.15" pageId="7" pageNumber="126" specimenCount="30" specimenCount-female="17" specimenCount-male="13" stateProvince="Khammouan" typeStatus="paratype">
<typeStatus id="54C3880906244919FCD226FEAAE4DB93" box="[866,979,1987,2013]" pageId="7" pageNumber="126" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7EFD2206244919FC6D26FEAD20DB93" box="[989,1047,1987,2013]" count="13" pageId="7" pageNumber="126" type="male">13 ♂</specimenCount>
,
<specimenCount id="9D7EFD2206244919FB9326FEAD6CDB93" box="[1059,1115,1987,2013]" count="17" pageId="7" pageNumber="126" type="female">17 ♀</specimenCount>
, same label data as holotype
</materialsCitation>
</paragraph>
<paragraph id="8BC736AB062B4916FF2721AAAC69DC9B" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="127">
(
<collectionCode id="ED69AE6E062B4916FF2F21AAA9D9DCFF" box="[159,238,151,177]" pageId="8" pageNumber="127">ASCG</collectionCode>
,
<collectionCode id="ED69AE6E062B4916FF4821AAA878DCFF" box="[248,335,151,177]" pageId="8" pageNumber="127">BMNH</collectionCode>
,
<collectionCode id="ED69AE6E062B4916FEE821AAA89ADCFF" box="[344,429,151,177]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="127">NMPC</collectionCode>
);
<materialsCitation id="3B103CF6062B4916FE0E21AAAC6DDC9B" collectingDate="2004-05-19" collectingDateMax="2004-05-22" collectingDateMin="2004-05-19" collectionCode="NMPC" collectorName="J. Bezdek &amp; All" country="Laos" elevation="200" latitude="18.158611" location="Vientiane prov." longLatPrecision="21" longitude="102.85777" pageId="8" pageNumber="127" specimenCount="1" specimenCount-male="1" stateProvince="Viangchan" typeStatus="paratype">
<specimenCount id="9D7EFD22062B4916FE0E21AAA8DFDCFF" box="[446,488,151,177]" count="1" pageId="8" pageNumber="127" type="male">1 ♂</specimenCount>
, “
<collectingCountry id="F36F763B062B4916FE4D21AAAB7CDCFF" box="[509,587,151,177]" name="Laos" pageId="8" pageNumber="127">LAOS</collectingCountry>
,
<collectingRegion id="49BCF849062B4916FDE421AAABCADCFF" box="[596,765,151,177]" country="Laos" name="Viangchan" pageId="8" pageNumber="127">Vientiane prov.</collectingRegion>
/ LAO PAKO env.,
<quantity id="4C809B4E062B4916FC6721AAAD2BDCFF" box="[983,1052,151,177]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="127" unit="m" value="200.0">
<elevation id="0055D198062B4916FC6721AAAD2BDCFF" box="[983,1052,151,177]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="127" unit="m" value="200.0">200 m</elevation>
</quantity>
/
<quantity id="4C809B4E062B4916FB9E21ABAD43DCFF" box="[1070,1140,150,177]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="127" unit="km" value="55.0">55 km</quantity>
NE
<collectingRegion id="49BCF849062B4916FB1321AAAC39DCFF" box="[1187,1294,151,177]" country="Laos" name="Viangchan" pageId="8" pageNumber="127">Vientiane</collectingRegion>
[
<geoCoordinate id="EE4C506C062B4916FAAB21AAACABDCFF" box="[1307,1436,151,177]" degrees="18" direction="north" minutes="9" orientation="latitude" pageId="8" pageNumber="127" precision="15" seconds="31" value="18.158611">18°9’31’’N</geoCoordinate>
<geoCoordinate id="EE4C506C062B4916FF272187A806DC9B" box="[151,305,186,213]" degrees="102" direction="east" minutes="51" orientation="longitude" pageId="8" pageNumber="127" precision="15" seconds="28" value="102.85777">102°51’28’’E</geoCoordinate>
] /
<date id="FFC6106B062B4916FEFE2186A8CBDC9B" box="[334,508,187,213]" pageId="8" pageNumber="127" value="2004-05-19" valueMax="2004-05-22" valueMin="2004-05-19">
<collectingDate id="EF82E983062B4916FEFE2186A8CBDC9B" box="[334,508,187,213]" pageId="8" pageNumber="127" value="2004-05-19" valueMax="2004-05-22" valueMin="2004-05-19">19.–22.V. 2004</collectingDate>
</date>
,
<collectorName id="268D537D062B4916FDB72186AB43DC9B" box="[519,628,187,213]" pageId="8" pageNumber="127">J. Bezděk</collectorName>
leg. (
<collectionCode id="ED69AE6E062B4916FD032186AA3EDC9B" box="[691,777,187,213]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="127">NMPC</collectionCode>
).
<collectorName id="268D537D062B4916FCAA2186AA09DC9B" box="[794,830,187,213]" pageId="8" pageNumber="127">All</collectorName>
<typeStatus id="54C38809062B4916FCF42186AA98DC9B" box="[836,943,187,213]" pageId="8" pageNumber="127" type="paratype">paratypes</typeStatus>
with the appropriate printed red label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3626520062B4917FF7721E3AC7FDD53" lastPageId="9" lastPageNumber="128" pageId="8" pageNumber="127" type="description">
<paragraph id="8BC736AB062B4916FF7721E3AB6CDD53" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF7721E3AB61DCB7" bold="true" box="[199,598,222,249]" pageId="8" pageNumber="127">
Description of
<typeStatus id="54C38809062B4916FE3121E2A8DFDCB7" box="[385,488,223,249]" pageId="8" pageNumber="127" type="holotype">holotype</typeStatus>
.
<emphasis id="B90CEAB9062B4916FE4A21E3AB61DCB7" bold="true" box="[506,598,222,249]" italics="true" pageId="8" pageNumber="127">Habitus</emphasis>
</emphasis>
oblong-oval, with continuous outline, broadest at mid-length of elytra, slightly convex. Dorsal surface submatt.
</paragraph>
<paragraph id="8BC736AB062B4916FF77201BAA14DDBB" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF77201BA86DDD0F" bold="true" box="[199,346,294,321]" italics="true" pageId="8" pageNumber="127">Colouration.</emphasis>
Head reddish brown, blackish around eyes and medially between eyes, lighter on clypeus, labrum and medially on vertex. Pronotum brown blackish, laterally reddish, pale colouration extending anteriorly; basal and anterior margin somewhat reddish translucent. Elytra brown blackish, with orange apex and sides laterally from elytral stria 6, and orangish pattern consisting of: two basal spots between striae 2–3 and 4–6; and longitudinal orange stripes beginning in second fourth between striae 3–5, and beginning subbasally between striae 5–6. Ventral surface reddish brown. Appendages orange to red (
<figureCitation id="13432A2E062B4916FD6120E6AA22DDBB" box="[721,789,474,501]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="8" pageNumber="127">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BC736AB062B4916FF7720C3AD68DEE7" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF7720C3A83BDE57" bold="true" box="[199,268,510,537]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF7720C3A832DE57" bold="true" box="[199,261,510,537]" italics="true" pageId="8" pageNumber="127">Head</emphasis>
.
</emphasis>
Moderately broad, ca. 0.62× width of pronotum, transversely elliptical. Anterior margin of clypeus indistinctly concave. Antenna with antennomeres long and slender. Reticulation consisting of fine, well impressed isodiametric polygonal meshes. Punctation double; several large setigerous punctures present in fronto-clypeal depressions, frontal depressions at level of anterior margin of eyes, and in depressions along inner margin of eyes; fine punctures smaller and sparser on clypeus, becoming larger and denser posteriorly.
</paragraph>
<paragraph id="8BC736AB062B4916FF77238EAB4BDF13" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF77238EA874DE83" bold="true" box="[199,323,691,717]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF77238EA80BDE83" bold="true" box="[199,316,691,717]" italics="true" pageId="8" pageNumber="127">Pronotum</emphasis>
.
</emphasis>
Transverse (width/length ratio = 2.8), broadest between posterior angles, lateral margins moderately curved. Lateral sides with beading thin but distinct, except at anterior angles. Reticulation and punctation similar to that of head; rows of coarse setigerous punctures present along anterior margin and laterally close to sides; several punctures present also in shallow basolateral depressions along basal margin.
<collectingRegion id="49BCF849062B4916FBE32222ADA9DF77" box="[1107,1182,799,825]" country="France" name="Centre" pageId="8" pageNumber="127">Centre</collectingRegion>
of disc with shallowly impressed medial longitudinal smooth line.
</paragraph>
<paragraph id="8BC736AB062B4916FF77225BA89CD817" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF77225BA822DFCF" bold="true" box="[199,277,870,897]" italics="true" pageId="8" pageNumber="127">Elytra.</emphasis>
Elytral striation consisting of six well impressed discal striae and one submarginal stria on each elytron: stria 1 beginning subbasally, striae 2–6 beginning at base of elytra; striae 1 and 4 ending close to apex, striae 2, 3 and 5 shorter than striae 1 and 4—not reaching apex of elytra; stria 6 shortest, ending approximately in apical fourth of elytral length; submarginal stria long, starting before elytral mid-length and terminating shortly before apex, at same level as dorsal striae 2, 3 and 5. Reticulation similar to that of head and pronotum, but less impressed. Punctation consisting of scattered very fine sparse punctures and coarse setigerous punctures present along elytral striae and lateral margins of elytra.
</paragraph>
<paragraph id="8BC736AB062B4916FF77255EAB37D88B" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF77255EA832D833" bold="true" box="[199,261,1123,1149]" italics="true" pageId="8" pageNumber="127">Legs.</emphasis>
Protibia modified, angled near base, distinctly broadened anteriorly, club shaped. Pro- and mesotarsomeres 1–3 distinctly broadened, with four rows of adhesive setae on their ventral side; claws simple. Longer spur on metatibia sinuate in apical third.
</paragraph>
<paragraph id="8BC736AB062B4916FF7725F3AC72DA47" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF7725F3A860D8A7" bold="true" box="[199,343,1230,1257]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF7725F3A867D8A7" bold="true" box="[199,336,1230,1257]" italics="true" pageId="8" pageNumber="127">Ventral side</emphasis>
.
</emphasis>
Prosternum sinuate anteriorly, obtusely keeled medially. Prosternal process shortly lanceolate, in cross-section convex, apex obtuse; process distinctly bordered laterally; reticulation not perceptible. Metaventrite with microsculpture consisting of polygonal meshes; lateral parts of metaventrite (‘metasternal wings’) tongueshaped, slender. Metacoxal lines incomplete, absent in ca. anterior fifth of metacoxal length. Metacoxal plates covered with long oblique strioles; reticulation consisting of oblique elongate polygonal meshes. Metacoxal processes rounded at posterior margin. Abdominal ventrites I–II with longitudinal strioles; ventrites III–IV with oblique strioles laterally. Tuft of setae present medially on ventrites III–V; ventrite VI with setigerous punctures laterally. Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I–II, oblique on ventrite III and transverse on ventrites IV–VI. Punctation consisting of fine, sparsely distributed punctures.
</paragraph>
<paragraph id="8BC736AB062B4916FF77272FAD3FDAF3" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF77272FA84EDA62" box="[199,377,1554,1581]" italics="true" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF77272FA845DA63" bold="true" box="[199,370,1554,1581]" italics="true" pageId="8" pageNumber="127">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view (
<figureCitation id="13432A2E062B4916FCDC272EAAFADA63" box="[876,973,1554,1581]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="8" pageNumber="127">Fig. 15a</figureCitation>
) broad in basal 2/3; apical third sickle-shaped, regularly curved and tapering distally, apex obtusely truncate; at 2/3 of its length with two distinct ‚teeth‘ on ventral side—broad rectangular proximal one, and medially incised and apically blunt distal one; third, small pointed tooth visible in ventro-lateral view (
<figureCitation id="13432A2E062B4916FD382742ABDDDAD7" box="[648,746,1662,1689]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="8" pageNumber="127">Fig. 15b</figureCitation>
). Parameres moderately broad, ‚D‘ shaped, with distinct lobe on inner margin, apex short and broad; apical lobe long, club-shaped (
<figureCitation id="13432A2E062B4916FC18279EAACCDAF3" box="[936,1019,1699,1725]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="8" pageNumber="127">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BC736AB062B4916FF7727FAA99BDB3F" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF7727FAA812DAAF" bold="true" box="[199,293,1735,1761]" pageId="8" pageNumber="127">Female.</emphasis>
Female identical to male in habitus. Protibia simple, not angled basally and only slightly broadened distally; pro- and mesotarsomeres not broadened, without adhesive setae; longer metatibial spur straight. Dorsal surface with reticulation slightly more impressed and thus appearing matt. Pronotum baso-laterally with several irregular striolae. Few weakly impressed short striolae present in middle part of elytra between elytral striae 5 and 6.
</paragraph>
<paragraph id="8BC736AB062B4917FF772647AD44DC9B" blockId="8.[151,1437,150,2013]" lastBlockId="9.[151,1437,151,717]" lastPageId="9" lastPageNumber="128" pageId="8" pageNumber="127">
<emphasis id="B90CEAB9062B4916FF772647A87ADBDA" bold="true" box="[199,333,1914,1940]" pageId="8" pageNumber="127">Variability.</emphasis>
Specimens of the
<typeStatus id="54C38809062B4916FD972646AB60DBDB" box="[551,599,1915,1941]" pageId="8" pageNumber="127">type</typeStatus>
series vary slightly in colouration, especially in infuscation of head and pronotum (from brown to nearly black) and elytra (from reddish brown to brown); some variation can be seen also in the extent of the orange colouration laterally on pronotum and elytra. In some specimens, elytral stria 1 is fragmented or absent in basal fourth of elytral length. The extent of the striolate elytral surface varies in females—in some specimens striolae are present in the middle part part of the elytra lateral to stria 2.
</paragraph>
<paragraph id="8BC736AB062A4917FF7721E2AC7FDD53" blockId="9.[151,1437,151,717]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FF7721E2A849DCB7" bold="true" box="[199,382,223,249]" pageId="9" pageNumber="128">Measurements.</emphasis>
TL:
<quantity id="4C809B4E062A4917FE0921E3AB74DCB7" box="[441,579,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" metricValueMax="6.2" metricValueMin="5.2" pageId="9" pageNumber="128" unit="mm" value="5.7" valueMax="6.2" valueMin="5.2">5.2–6.2 mm</quantity>
(mean value: 5.7 ±
<quantity id="4C809B4E062A4917FC9D21E2AABEDCB7" box="[813,905,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="128" unit="mm" value="0.2">0.2 mm</quantity>
);
<typeStatus id="54C38809062A4917FC2F21E2AD32DCB7" box="[927,1029,223,249]" pageId="9" pageNumber="128" type="holotype">holotype</typeStatus>
:
<quantity id="4C809B4E062A4917FBA221E3AD5ADCB7" box="[1042,1133,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="9" pageNumber="128" unit="mm" value="5.9">5.9 mm</quantity>
. TL-h:
<quantity id="4C809B4E062A4917FB7421E3AC7ADCB7" box="[1220,1357,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.6" metricValueMin="4.7" pageId="9" pageNumber="128" unit="mm" value="5.15" valueMax="5.6" valueMin="4.7">4.7–5.6 mm</quantity>
(mean value: 5.1 ±
<quantity id="4C809B4E062A4917FE93203EA84ADD53" box="[291,381,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="128" unit="mm" value="0.2">0.2 mm</quantity>
);
<typeStatus id="54C38809062A4917FE21203EA8C0DD53" box="[401,503,259,285]" pageId="9" pageNumber="128" type="holotype">holotype</typeStatus>
:
<quantity id="4C809B4E062A4917FDB1203FAB6DDD53" box="[513,602,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="128" unit="mm" value="5.3">5.3 mm</quantity>
. MW:
<quantity id="4C809B4E062A4917FD16203EAA1ADD53" box="[678,813,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="9" pageNumber="128" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7–3.1 mm</quantity>
(mean value: 2.9 ±
<quantity id="4C809B4E062A4917FBBC203EAD51DD53" box="[1036,1126,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="128" unit="mm" value="0.1">0.1 mm</quantity>
);
<typeStatus id="54C38809062A4917FBCA203EADD7DD53" box="[1146,1248,259,285]" pageId="9" pageNumber="128" type="holotype">holotype</typeStatus>
: 3.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="C3626520062A4917FF77201BAA0BDE2F" pageId="9" pageNumber="128" type="diagnosis">
<paragraph id="8BC736AB062A4917FF77201BAA0BDE2F" blockId="9.[151,1437,151,717]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FF77201BA8E7DD0E" bold="true" box="[199,464,294,320]" pageId="9" pageNumber="128">Differential diagnosis.</emphasis>
Based on the presence of six dorsal striae and a submarginal elytral stria,
<taxonomicName id="4C784D28062A4917FA9B201AA81EDD2B" authorityName="Hájek &amp; Sheth" authorityYear="2024" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="lanxangensis" status="sp. nov.">
<emphasis id="B90CEAB9062A4917FA9B201AA81EDD2B" italics="true" pageId="9" pageNumber="128">Copelatus lanxangensis</emphasis>
</taxonomicName>
<emphasis id="B90CEAB9062A4917FE812076A8BDDD2B" bold="true" box="[305,394,331,357]" pageId="9" pageNumber="128">
<taxonomicNameLabel id="A23F57C2062A4917FE812076A8BDDD2B" box="[305,394,331,357]" pageId="9" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be classified within the
<taxonomicName id="4C784D28062A4917FD7A2076AAB3DD2A" authorityName="Regimbart" authorityYear="1899" box="[714,900,331,357]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="irinus">
<emphasis id="B90CEAB9062A4917FD7A2076AAB3DD2A" box="[714,900,331,357]" italics="true" pageId="9" pageNumber="128">Copelatus irinus</emphasis>
</taxonomicName>
group sensu
<bibRefCitation id="EFE94B5A062A4917FBA92076ADFDDD2B" author="Guignot, F." box="[1049,1226,331,357]" pageId="9" pageNumber="128" pagination="659 - 995" refId="ref6814" refString="Guignot, F. (1961) Revision des hydrocanthares d'Afrique (Coleoptera Dityscoidea [sic!]). Troisieme partie. Annales du Musee Royal du Congo Belge, Serie 8 vo (Sciences Zoologiques), 90, 659 - 995." type="journal article" year="1961">Guignot (1961)</bibRefCitation>
. The characteristic shape of the median lobe with two ‘teeth’ on the ventral side (
<figureCitation id="13432A2E062A4917FCF52052AAA1DDC7" box="[837,918,366,393]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="9" pageNumber="128">Fig. 15</figureCitation>
) places the new species to the newly proposed
<taxonomicName id="4C784D28062A4917FF2720AEA833DDE3" authorityName="Sharp" authorityYear="1882" box="[151,260,403,429]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B90CEAB9062A4917FF2720AEA833DDE3" box="[151,260,403,429]" italics="true" pageId="9" pageNumber="128">C. latipes</emphasis>
</taxonomicName>
complex. The new species is most similar to the partly sympatric
<taxonomicName id="4C784D28062A4917FC4620AEAD54DDE3" authorityName="Sharp" authorityYear="1882" box="[1014,1123,403,429]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B90CEAB9062A4917FC4620AEAD54DDE3" box="[1014,1123,403,429]" italics="true" pageId="9" pageNumber="128">C. latipes</emphasis>
</taxonomicName>
. Both species can be easily distinguished by the elytral striation: while stria 1 begins subbasally or at most in basal fourth of elytral length in
<taxonomicName id="4C784D28062A4917FA33208AA81EDDBB" authorityName="Hájek &amp; Sheth" authorityYear="2024" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="lanxangensis" status="sp. nov.">
<emphasis id="B90CEAB9062A4917FA33208AA81EDDBB" italics="true" pageId="9" pageNumber="128">C. lanxangensis</emphasis>
</taxonomicName>
<emphasis id="B90CEAB9062A4917FE8120E6A8BCDDBB" bold="true" box="[305,395,475,501]" pageId="9" pageNumber="128">
<taxonomicNameLabel id="A23F57C2062A4917FE8120E6A8BCDDBB" box="[305,395,475,501]" pageId="9" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, it is largely absent and beginning in apical fifth in
<taxonomicName id="4C784D28062A4917FC6120E6AD0ADDBB" authorityName="Sharp" authorityYear="1882" box="[977,1085,475,501]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B90CEAB9062A4917FC6120E6AD0ADDBB" box="[977,1085,475,501]" italics="true" pageId="9" pageNumber="128">C. latipes</emphasis>
</taxonomicName>
(see
<figureCitation id="13432A2E062A4917FBC820E6AC33DDBB" box="[1144,1284,474,501]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="9" pageNumber="128">Figs 5 and 6</figureCitation>
). In addition,
<taxonomicName id="4C784D28062A4917FF2720C2A87CDE57" authorityName="Hájek &amp; Sheth" authorityYear="2024" box="[151,331,511,537]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="lanxangensis" status="sp. nov.">
<emphasis id="B90CEAB9062A4917FF2720C2A87CDE57" box="[151,331,511,537]" italics="true" pageId="9" pageNumber="128">C. lanxangensis</emphasis>
</taxonomicName>
<emphasis id="B90CEAB9062A4917FEE320C2A89ADE57" bold="true" box="[339,429,511,537]" pageId="9" pageNumber="128">
<taxonomicNameLabel id="A23F57C2062A4917FEE320C2A89ADE57" box="[339,429,511,537]" pageId="9" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from all other
<taxonomicName id="4C784D28062A4917FCF620C2AD27DE57" box="[838,1040,511,537]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FCF620C2AA8FDE57" box="[838,952,511,537]" italics="true" pageId="9" pageNumber="128">Copelatus</emphasis>
species
</taxonomicName>
by the shape of the male genitalia, especially by the regularly curved apical third of the median lobe, the broad rectangular proximal median ‘tooth’ and the medially incised and apically blunt distal tooth (
<figureCitation id="13432A2E062A4917FD6C237AAA18DE2F" box="[732,815,582,609]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="9" pageNumber="128">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3626520062A4917FF772356AA07DEE7" pageId="9" pageNumber="128" type="etymology">
<paragraph id="8BC736AB062A4917FF772356AA07DEE7" blockId="9.[151,1437,151,717]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FF772356A879DECB" bold="true" box="[199,334,619,645]" pageId="9" pageNumber="128">Etymology.</emphasis>
The species is named after Lan Xang or Lancang, a Lao kingdom that held the area of present-day
<collectingCountry id="F36F763B062A4917FF2723B2A9FADEE7" box="[151,205,655,681]" name="Laos" pageId="9" pageNumber="128">Laos</collectingCountry>
from 1353 to 1707. The specific name is an adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3626520062A4917FF77238FADF9DE83" box="[199,1230,690,717]" pageId="9" pageNumber="128" type="distribution">
<paragraph id="8BC736AB062A4917FF77238FADF9DE83" blockId="9.[151,1437,151,717]" box="[199,1230,690,717]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FF77238FA856DE82" bold="true" box="[199,353,690,716]" pageId="9" pageNumber="128">Distribution.</emphasis>
The species is so far only known from two localities in central
<collectingCountry id="F36F763B062A4917FB97238EAD6ADE83" box="[1063,1117,691,717]" name="Laos" pageId="9" pageNumber="128">Laos</collectingCountry>
(
<figureCitation id="13432A2E062A4917FBDD238EADF6DE83" box="[1133,1217,691,717]" captionStart="FIGURE 19" captionStartId="6.[152,255,1288,1312]" captionTargetBox="[156,1437,273,1261]" captionTargetId="figure-66@6.[151,1437,273,1265]" captionTargetPageId="6" captionText="FIGURE 19. Map of distribution of Copelatus latipes species complex." figureDoi="http://doi.org/10.5281/zenodo.12742768" httpUri="https://zenodo.org/record/12742768/files/figure.png" pageId="9" pageNumber="128">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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277
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<mods:title id="1EBBCF4387ED0429EF528B0297352B37">A review of the Oriental Copelatus latipes complex, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<paragraph id="8BC736AB06254918FF27244FA8E6D9C3" blockId="6.[151,465,1394,1457]" box="[151,465,1394,1421]" pageId="6" pageNumber="125">
<heading id="D08F81C706254918FF27244FA8E6D9C3" bold="true" box="[151,465,1394,1421]" fontSize="11" level="1" pageId="6" pageNumber="125" reason="1">
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<taxonomicName id="4C784D2806254918FF27244FA85BD9C3" authorityName="Hájek &amp; Sheth" authorityYear="2024" box="[151,364,1394,1421]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="kopetzi" status="sp. nov.">
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(
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,
<figureCitation id="13432A2E06254918FF4424AAA80CD9FF" box="[244,315,1431,1457]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="6" pageNumber="125">13–14</figureCitation>
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<materialsCitation id="3B103CF606254918FF2724E2AA26DA53" country="Nepal" county="Dsauda river" latitude="28.8975" location="Sudurpashchim Province" longLatPrecision="20" longitude="80.2275" municipality="Shuklaphanta Nature Reserve" pageId="6" pageNumber="125" specimenCount="1" typeStatus="holotype">
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locality.
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<location id="8EA7607006254918FE2824E2AB80D9B7" LSID="urn:lsid:plazi:treatment:03D187BD06254919FF27244FAC7DDAD7:8EA7607006254918FE2824E2AB80D9B7" box="[408,695,1503,1529]" country="Nepal" county="Dsauda river" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Shuklaphanta Nature Reserve" name="Sudurpashchim Province" pageId="6" pageNumber="125">Sudurpashchim Province</location>
,
<location id="8EA7607006254918FD7624E2AA87D9B7" LSID="urn:lsid:plazi:treatment:03D187BD06254919FF27244FAC7DDAD7:8EA7607006254918FD7624E2AA87D9B7" box="[710,944,1503,1529]" country="Nepal" county="Dsauda river" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Shuklaphanta Nature Reserve" name="Kanchanpur District" pageId="6" pageNumber="125">Kanchanpur District</location>
, ca.
<locationDeviation id="9A752F8906254918FC5924E2ADC1D9B7" box="[1001,1270,1503,1529]" location="Bhimdatta" pageId="6" pageNumber="125">
<quantity id="4C809B4E06254918FC5924E2AD11D9B7" box="[1001,1062,1503,1529]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="125" unit="km" value="9.0">9 km</quantity>
SE of Bhimdatta
</locationDeviation>
,
<collectingMunicipality id="6BA3ACD106254918FAB424E2A870DA53" pageId="6" pageNumber="125">Shuklaphanta Nature Reserve</collectingMunicipality>
,
<collectingCounty id="62A64E2706254918FEE3273EA8D2DA53" box="[339,485,1539,1565]" pageId="6" pageNumber="125">Dsauda river</collectingCounty>
,
<geoCoordinate id="EE4C506C06254918FE5F273EAB4ADA50" box="[495,637,1539,1566]" degrees="28" direction="north" minutes="53" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="51" value="28.8975">28°53’51’’N</geoCoordinate>
<geoCoordinate id="EE4C506C06254918FD34273EAA26DA53" box="[644,785,1539,1565]" degrees="80" direction="east" minutes="13" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="39" value="80.2275">80°13’39’’E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736AB06254918FF77271AADD7DAC7" blockId="6.[151,1437,1503,2033]" pageId="6" pageNumber="125">
<emphasis id="B90CEAB906254918FF77271AA8B6DA0F" bold="true" box="[199,385,1575,1601]" pageId="6" pageNumber="125">Type material.</emphasis>
<materialsCitation id="3B103CF606254918FE24271AADD7DAC7" collectingDate="2017-07-01" collectionCode="NMPC" collectorName="A. Kopetz" county="Kanchanpur" determinerName="Sheth" elevation="160" latitude="28.8975" location="Shuklaphanta" longLatPrecision="20" longitude="80.2275" municipality="Mahandranagar" pageId="6" pageNumber="125" specimenCount="2" specimenCount-male="2" stateProvince="Mahakali" typeStatus="holotype">
<typeStatus id="54C3880906254918FE24271AA8CBDA0F" box="[404,508,1575,1601]" pageId="6" pageNumber="125" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7EFD2206254918FDBF271AAB13DA0F" box="[527,548,1575,1601]" count="1" pageId="6" pageNumber="125" type="male">♂</specimenCount>
(
<collectionCode id="ED69AE6E06254918FD8F271AABA3DA0F" box="[575,660,1575,1601]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="6" pageNumber="125">NMPC</collectionCode>
), labelled: “NEP:
<collectingRegion id="49BCF84906254918FC32271AAADCDA0F" box="[898,1003,1575,1601]" country="Nepal" name="Mahakali" pageId="6" pageNumber="125">Mahakali</collectingRegion>
/
<collectingCounty id="62A64E2706254918FC44271AAD4ADA0F" box="[1012,1149,1575,1601]" pageId="6" pageNumber="125">Kanchanpur</collectingCounty>
vic. /
<collectingMunicipality id="6BA3ACD106254918FB57271AACAEDA0F" box="[1255,1433,1575,1601]" pageId="6" pageNumber="125">Mahandranagar</collectingMunicipality>
,
<location id="8EA7607006254918FF272776A818DA2B" LSID="urn:lsid:plazi:treatment:03D187BD06254919FF27244FAC7DDAD7:8EA7607006254918FF272776A818DA2B" box="[151,303,1611,1637]" county="Kanchanpur" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Mahandranagar" name="Shuklaphanta" pageId="6" pageNumber="125" stateProvince="Mahakali">Shuklaphanta</location>
/ Nature Res.,
<location id="8EA7607006254918FE792776AB68DA2B" LSID="urn:lsid:plazi:treatment:03D187BD06254919FF27244FAC7DDAD7:8EA7607006254918FE792776AB68DA2B" box="[457,607,1611,1637]" county="Kanchanpur" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Mahandranagar" name="Dsauda river" pageId="6" pageNumber="125" stateProvince="Mahakali">Dsauda river</location>
, LFF /
<geoCoordinate id="EE4C506C06254918FD1D2776AA0EDA28" box="[685,825,1611,1638]" degrees="28" direction="north" minutes="53" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="51" value="28.8975">N28°53’51”</geoCoordinate>
,
<geoCoordinate id="EE4C506C06254918FCF42776AAFBDA2B" box="[836,972,1611,1637]" degrees="80" direction="east" minutes="13" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="39" value="80.2275">E80°13’39”</geoCoordinate>
,
<quantity id="4C809B4E06254918FC672776AD20DA2B" box="[983,1047,1611,1637]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="125" unit="m" value="160.0">
<elevation id="0055D19806254918FC672776AD20DA2B" box="[983,1047,1611,1637]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="125" unit="m" value="160.0">160m</elevation>
</quantity>
/
<date id="FFC6106B06254918FB9B2776AD95DA2B" box="[1067,1186,1611,1637]" pageId="6" pageNumber="125" value="2017-07-01">
<collectingDate id="EF82E98306254918FB9B2776AD95DA2B" box="[1067,1186,1611,1637]" pageId="6" pageNumber="125" value="2017-07-01">1.VII.2017</collectingDate>
</date>
,leg.
<collectorName id="268D537D06254918FB7F2776AC74DA2B" box="[1231,1347,1611,1637]" pageId="6" pageNumber="125">A.Kopetz</collectorName>
#17–19 [p] //
<typeStatus id="54C3880906254918FF652752A84CDAC7" box="[213,379,1647,1673]" pageId="6" pageNumber="125" type="holotype">HOLOTYPUS</typeStatus>
<specimenCount id="9D7EFD2206254918FE332752A8AFDAC7" box="[387,408,1647,1673]" count="1" pageId="6" pageNumber="125" type="male">♂</specimenCount>
/
<taxonomicName id="4C784D2806254918FE1E2752AB7EDAC7" authorityName="Erichson" authorityYear="1832" box="[430,585,1647,1673]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAB906254918FE1E2752AB7EDAC7" box="[430,585,1647,1673]" italics="true" pageId="6" pageNumber="125">COPELATUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C784D2806254918FDEE2752AB9ADAC7" authorityName="Hájek &amp; Sheth" authorityYear="2024" box="[606,685,1647,1673]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="kopetzi" status="sp. nov.">
<emphasis id="B90CEAB906254918FDEE2752AB9ADAC7" box="[606,685,1647,1673]" italics="true" pageId="6" pageNumber="125">kopetzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F57C206254918FD042752AA3CDAC7" box="[692,779,1647,1673]" pageId="6" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
/ Hájek &amp;
<determinerName id="7E1BDA6F06254918FC372752AAF0DAC7" box="[903,967,1647,1673]" pageId="6" pageNumber="125">Sheth</determinerName>
det. 2024 [red label, p]”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C362652006254919FF7727AEAA6DD91B" lastPageId="7" lastPageNumber="126" pageId="6" pageNumber="125" type="description">
<paragraph id="8BC736AB06254918FF7727AEAB6CDA9F" blockId="6.[151,1437,1503,2033]" pageId="6" pageNumber="125">
<emphasis id="B90CEAB906254918FF7727AEAB61DAE3" bold="true" box="[199,598,1682,1709]" pageId="6" pageNumber="125">
Description of
<typeStatus id="54C3880906254918FE3127AEA8DFDAE3" box="[385,488,1683,1709]" pageId="6" pageNumber="125" type="holotype">holotype</typeStatus>
.
<emphasis id="B90CEAB906254918FE4A27AFAB61DAE3" bold="true" box="[506,598,1682,1709]" italics="true" pageId="6" pageNumber="125">Habitus</emphasis>
</emphasis>
oblong-oval, with continuous outline, broadest at mid-length of elytra, slightly convex. Dorsal surface submatt.
</paragraph>
<paragraph id="8BC736AB06254918FF7727E7AA13DBE7" blockId="6.[151,1437,1503,2033]" pageId="6" pageNumber="125">
<emphasis id="B90CEAB906254918FF7727E7A86DDABB" bold="true" box="[199,346,1754,1781]" italics="true" pageId="6" pageNumber="125">Colouration.</emphasis>
Head reddish brown, blackish around eyes and medially between eyes, lighter on clypeus, labrum and medially on vertex. Pronotum brown blackish, laterally reddish, pale colouration extending anteriorly. Elytra black, with orange apex and sides laterally from elytral stria 6, and orangish pattern consisting of: irregular basal transverse band split into three spots, somewhat extending between striae 1–2 and 4–5; and longitudinal orange stripes between striae 3–4, in posterior fourth between striae 4–5 and in posterior 2/3 between striae 5–6. Ventral surface reddish brown. Appendages orange to red (
<figureCitation id="13432A2E06254918FD6126B2AA21DBE7" box="[721,790,1935,1961]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="6" pageNumber="125">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BC736AB06254919FF77268FAD68DCB7" blockId="6.[151,1437,1503,2033]" lastBlockId="7.[151,1437,151,1689]" lastPageId="7" lastPageNumber="126" pageId="6" pageNumber="125">
<emphasis id="B90CEAB906254918FF77268FA83BDB83" bold="true" box="[199,268,1970,1997]" pageId="6" pageNumber="125">
<emphasis id="B90CEAB906254918FF77268FA832DB83" bold="true" box="[199,261,1970,1997]" italics="true" pageId="6" pageNumber="125">Head</emphasis>
.
</emphasis>
Moderately broad, ca. 0.63× width of pronotum, transversely elliptical. Anterior margin of clypeus slightly concave. Antenna with antennomeres long and slender. Reticulation consisting of fine, well impressed isodiametric polygonal meshes. Punctation double; several large setigerous punctures present in fronto-clypeal depressions, frontal depressions at level of anterior margin of eyes, and in depressions along inner margin of eyes; fine punctures smaller and sparser on clypeus, becoming larger and denser posteriorly.
</paragraph>
<paragraph id="8BC736AB06244919FF77203EAAD1DDE3" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF77203EA874DD53" bold="true" box="[199,323,259,285]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF77203EA80BDD53" bold="true" box="[199,316,259,285]" italics="true" pageId="7" pageNumber="126">Pronotum</emphasis>
.
</emphasis>
Transverse (width/length ratio = 2.9), broadest between posterior angles, lateral margins moderately curved. Lateral sides with beading thin but distinct, except at anterior angles. Reticulation similar to that of head. Punctation similar to that of head; rows of coarse setigerous punctures present along anterior margin, laterally close to sides, several punctures present also in shallow basolateral depressions along basal margin.
<collectingRegion id="49BCF84906244919FB732052AC39DDC7" box="[1219,1294,367,393]" country="France" name="Centre" pageId="7" pageNumber="126">Centre</collectingRegion>
of disc with shallowly impressed medial longitudinal smooth line (without reticulation).
</paragraph>
<paragraph id="8BC736AB06244919FF77208BAB18DEE7" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF77208BA822DD9F" bold="true" box="[199,277,438,465]" italics="true" pageId="7" pageNumber="126">Elytra.</emphasis>
Elytral striation consisting of six well impressed discal striae and one submarginal stria on each elytron: stria 1 beginning subbasally, striae 2–6 beginning at base of elytra; striae 1 and 4 ending close to apex; striae 2, 3 and 5 shorter than striae 1 and 4—not reaching apex of elytra; stria 6 shortest, ending approximately in apical fifth of elytral length; submarginal stria long, starting at about elytral mid-length and terminating shortly before apex, at same level as dorsal striae 2, 3 and 5. Reticulation similar to that of head and pronotum, but less impressed. Punctation consisting of coarse setigerous punctures present along elytral striae and lateral margins of elytra, and scattered very fine sparse punctures.
</paragraph>
<paragraph id="8BC736AB06244919FF77238EABB2DF5B" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF77238EA832DE83" bold="true" box="[199,261,691,717]" italics="true" pageId="7" pageNumber="126">Legs.</emphasis>
Protibia modified, angled near base, distinctly broadened anteriorly, club shaped. Pro- and mesotarsomeres 1–3 distinctly broadened, with four rows of adhesive setae on their ventral side; claws simple. Longer spur on metatibia indistinctly sinuate in apical third.
</paragraph>
<paragraph id="8BC736AB06244919FF772223A83BD833" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF772223A860DF77" bold="true" box="[199,343,798,825]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF772223A867DF77" bold="true" box="[199,336,798,825]" italics="true" pageId="7" pageNumber="126">Ventral side</emphasis>
.
</emphasis>
Prosternum sinuate anteriorly, obtusely keeled medially. Prosternal process shortly lanceolate, in cross-section convex, apex obtuse; process distinctly bordered laterally; reticulation not perceptible. Metaventrite with microsculpture consisting of polygonal meshes; lateral parts of metaventrite (‘metasternal wings’) tongueshaped, slender. Metacoxal lines incomplete, absent in ca. anterior fifth of metacoxal length. Metacoxal plates covered with long oblique strioles; reticulation consisting of elongate, oblique, polygonal meshes. Metacoxal processes rounded at posterior margin. Abdominal ventrites I–II with longitudinal strioles; ventrites III–IV with oblique strioles laterally. Tuft of setae present medially on ventrites III–V; ventrite VI with setigerous punctures laterally on either side. Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I–II, oblique on ventrite III and transverse on ventrites IV–VI. Punctation consisting of fine, sparsely distributed punctures.
</paragraph>
<paragraph id="8BC736AB06244919FF7725BBAAD1D943" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF7725BBA842D8EE" box="[199,373,1158,1185]" italics="true" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF7725BBA859D8EF" bold="true" box="[199,366,1158,1185]" italics="true" pageId="7" pageNumber="126">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view (
<figureCitation id="13432A2E06244919FCF525BAAAA2D8EF" box="[837,917,1159,1185]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="7" pageNumber="126">Fig. 13</figureCitation>
) broad in basal 2/3; apical third sickle-shaped, evenly curved, subapically narrowed to pointed apex; at 2/3 of its length with two distinct ‚teeth‘ on ventral side— broad blunt proximal one, and bilobed distal one. Parameres moderately broad, ‚D‘ shaped, with distinct lobe on inner margin, apex short and broad; apical lobe long, club-shaped (
<figureCitation id="13432A2E06244919FC3525CEAAEFD943" box="[901,984,1267,1293]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="7" pageNumber="126">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8BC736AB06244919FF77242AA895D97F" blockId="7.[151,1437,151,1689]" box="[199,418,1303,1329]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF77242AA812D97F" bold="true" box="[199,293,1303,1329]" pageId="7" pageNumber="126">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BC736AB06244919FF772406AA6DD91B" blockId="7.[151,1437,151,1689]" box="[199,858,1338,1365]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF772406A849D91B" bold="true" box="[199,382,1339,1365]" pageId="7" pageNumber="126">Measurements.</emphasis>
TL:
<quantity id="4C809B4E06244919FE052407AB39D91B" box="[437,526,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="7" pageNumber="126" unit="mm" value="5.7">5.7 mm</quantity>
; Tl-h:
<quantity id="4C809B4E06244919FDE82407AB86D91B" box="[600,689,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="126" unit="mm" value="5.1">5.1 mm</quantity>
; MW:
<quantity id="4C809B4E06244919FD4D2406AA61D91B" box="[765,854,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="126" unit="mm" value="2.9">2.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C362652006244919FF772463AD3EDA63" pageId="7" pageNumber="126" type="diagnosis">
<paragraph id="8BC736AB06244919FF772463AD3EDA63" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF772463A8E7D936" bold="true" box="[199,464,1374,1400]" pageId="7" pageNumber="126">Differential diagnosis.</emphasis>
Based on the presence of six dorsal striae and a submarginal elytral stria,
<taxonomicName id="4C784D2806244919FA9B2462A9D1D9D3" authorityName="Hájek &amp; Sheth" authorityYear="2024" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="kopetzi" status="sp. nov.">
<emphasis id="B90CEAB906244919FA9B2462A9D1D9D3" italics="true" pageId="7" pageNumber="126">Copelatus kopetzi</emphasis>
</taxonomicName>
<emphasis id="B90CEAB906244919FF5D24BEA870D9D3" bold="true" box="[237,327,1411,1437]" pageId="7" pageNumber="126">
<taxonomicNameLabel id="A23F57C206244919FF5D24BEA870D9D3" box="[237,327,1411,1437]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be classified within the
<taxonomicName id="4C784D2806244919FD3524BEAA08D9D2" authorityName="Regimbart" authorityYear="1899" box="[645,831,1411,1437]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="irinus">
<emphasis id="B90CEAB906244919FD3524BEAA08D9D2" box="[645,831,1411,1437]" italics="true" pageId="7" pageNumber="126">Copelatus irinus</emphasis>
</taxonomicName>
group
<emphasis id="B90CEAB906244919FC3E24BEAAFBD9D2" box="[910,972,1411,1436]" italics="true" pageId="7" pageNumber="126">sensu</emphasis>
<bibRefCitation id="EFE94B5A06244919FC6324BEADB3D9D3" author="Guignot, F." box="[979,1156,1411,1437]" pageId="7" pageNumber="126" pagination="659 - 995" refId="ref6814" refString="Guignot, F. (1961) Revision des hydrocanthares d'Afrique (Coleoptera Dityscoidea [sic!]). Troisieme partie. Annales du Musee Royal du Congo Belge, Serie 8 vo (Sciences Zoologiques), 90, 659 - 995." type="journal article" year="1961">Guignot (1961)</bibRefCitation>
. The characteristic shape of the median lobe with two ‘teeth’ on the ventral side (
<figureCitation id="13432A2E06244919FCA9249AAA5CD98F" box="[793,875,1447,1473]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="7" pageNumber="126">Fig. 13</figureCitation>
) places the new species into the newly proposed
<taxonomicName id="4C784D2806244919FF2724F6A833D9AB" authorityName="Sharp" authorityYear="1882" box="[151,260,1483,1509]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B90CEAB906244919FF2724F6A833D9AB" box="[151,260,1483,1509]" italics="true" pageId="7" pageNumber="126">C. latipes</emphasis>
</taxonomicName>
complex. Within this species complex, the new species is most similar to the sympatric
<taxonomicName id="4C784D2806244919FB5E24F6AC6CD9AB" authorityName="Rocchi" authorityYear="1976" box="[1262,1371,1483,1509]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="brivioi">
<emphasis id="B90CEAB906244919FB5E24F6AC6CD9AB" box="[1262,1371,1483,1509]" italics="true" pageId="7" pageNumber="126">C. brivioi</emphasis>
</taxonomicName>
; both species can be distinguished from each other by the shape of the median lobe of aedeagus—especially by the shape of the apex and the shape of processes on the ventral side (see
<figureCitation id="13432A2E06244919FCE4272EAACCDA63" box="[852,1019,1555,1581]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="7" pageNumber="126">Figs 11 and 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C362652006244919FF77270AADDCDA3B" pageId="7" pageNumber="126" type="etymology">
<paragraph id="8BC736AB06244919FF77270AADDCDA3B" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF77270AA879DA1F" bold="true" box="[199,334,1591,1617]" pageId="7" pageNumber="126">Etymology.</emphasis>
The new species is dedicated to its collector, Andreas Kopetz (Amt Wachsenburg,
<collectingCountry id="F36F763B06244919FB5D270AAC6DDA1F" box="[1261,1370,1591,1617]" name="Germany" pageId="7" pageNumber="126">Germany</collectingCountry>
), well known specialist on soldier beetles (
<taxonomicName id="4C784D2806244919FD9C2766AB8DDA3B" authorityName="Imhoff" authorityYear="1856" box="[556,698,1627,1653]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
). The specific name is a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C362652006244919FF772743AC7DDAD7" box="[199,1354,1662,1689]" pageId="7" pageNumber="126" type="distribution">
<paragraph id="8BC736AB06244919FF772743AC7DDAD7" blockId="7.[151,1437,151,1689]" box="[199,1354,1662,1689]" pageId="7" pageNumber="126">
<emphasis id="B90CEAB906244919FF772743A856DAD6" bold="true" box="[199,353,1662,1688]" pageId="7" pageNumber="126">Distribution.</emphasis>
The species is so far only known from the
<typeStatus id="54C3880906244919FCF42742AA43DAD7" box="[836,884,1663,1689]" pageId="7" pageNumber="126">type</typeStatus>
locality in south-western
<collectingCountry id="F36F763B06244919FB272742ADEDDAD7" box="[1175,1242,1663,1689]" name="Nepal" pageId="7" pageNumber="126">Nepal</collectingCountry>
(
<figureCitation id="13432A2E06244919FB592742AC0BDAD7" box="[1257,1340,1663,1689]" captionStart="FIGURE 19" captionStartId="6.[152,255,1288,1312]" captionTargetBox="[156,1437,273,1261]" captionTargetId="figure-66@6.[151,1437,273,1265]" captionTargetPageId="6" captionText="FIGURE 19. Map of distribution of Copelatus latipes species complex." figureDoi="http://doi.org/10.5281/zenodo.12742768" httpUri="https://zenodo.org/record/12742768/files/figure.png" pageId="7" pageNumber="126">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

269
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<mods:title id="A695A5ECAFE7436993D4D77DE21D0DE6">A review of the Oriental Copelatus latipes complex, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<heading id="D08F81C70627491AFF272528AB3AD87E" bold="true" box="[151,525,1045,1072]" fontSize="11" level="1" pageId="4" pageNumber="123" reason="1">
<taxonomicName id="4C784D280627491AFF272528AB3AD87E" ID-CoL="Y357" authority="Rocchi, 1976" authorityName="Rocchi" authorityYear="1976" box="[151,525,1045,1072]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="brivioi">
<emphasis id="B90CEAB90627491AFF272528AB3AD87E" bold="true" box="[151,525,1045,1072]" pageId="4" pageNumber="123">
<emphasis id="B90CEAB90627491AFF272528A852D87E" bold="true" box="[151,357,1045,1072]" italics="true" pageId="4" pageNumber="123">Copelatus brivioi</emphasis>
<bibRefCitation id="EFE94B5A0627491AFEDD2528AB3AD87E" author="Rocchi, S." box="[365,525,1045,1072]" pageId="4" pageNumber="123" pagination="177 - 180" refId="ref7362" refString="Rocchi, S. (1976) Ditiscidi del Bengla-desh con descrizione di Copelatus brivioi n. sp. (Coleoptera Dytiscidae). Bollettino della Societa Entomologica Italiana, 108, 177 - 180." type="journal article" year="1976">Rocchi, 1976</bibRefCitation>
</emphasis>
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<subSubSection id="C36265200627491AFF272507A875D81A" box="[151,322,1082,1108]" pageId="4" pageNumber="123" type="description">
<paragraph id="8BC736AB0627491AFF272507A875D81A" blockId="4.[151,525,1045,1108]" box="[151,322,1082,1108]" pageId="4" pageNumber="123">
(
<figureCitation id="13432A2E0627491AFF2F2507A9D0D81A" box="[159,231,1082,1108]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="4" pageNumber="123">Figs 3</figureCitation>
,
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)
</paragraph>
</subSubSection>
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<emphasis id="B90CEAB90627491AFF272543A87FD8D8" box="[151,328,1150,1174]" italics="true" pageId="4" pageNumber="123">Copelatus brivioi</emphasis>
<bibRefCitation id="EFE94B5A0627491AFEFE2543AB3BD8D8" author="Rocchi, S." box="[334,524,1150,1174]" pageId="4" pageNumber="123" pagination="177 - 180" refId="ref7362" refString="Rocchi, S. (1976) Ditiscidi del Bengla-desh con descrizione di Copelatus brivioi n. sp. (Coleoptera Dytiscidae). Bollettino della Societa Entomologica Italiana, 108, 177 - 180." type="journal article" year="1976">Rocchi, 1976: 177</bibRefCitation>
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.
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<emphasis id="B90CEAB90627491AFF2725F8A801D890" bold="true" box="[151,310,1220,1247]" pageId="4" pageNumber="123">
<typeStatus id="54C388090627491AFF2725F8A9E5D891" box="[151,210,1221,1247]" pageId="4" pageNumber="123">Type</typeStatus>
locality.
</emphasis>
“Dinajpur, Bengla-Desh” [
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,
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,
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, ca.
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.
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<emphasis id="B90CEAB90627491AFF7725D4A844D94C" bold="true" box="[199,371,1256,1283]" pageId="4" pageNumber="123">Type material.</emphasis>
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<typeStatus id="54C388090627491AFEC725D4A8EED94D" box="[375,473,1257,1283]" pageId="4" pageNumber="123" type="paratype">Paratype</typeStatus>
<specimenCount id="9D7EFD220627491AFE6E25D4A8C4D94D" box="[478,499,1257,1283]" count="1" pageId="4" pageNumber="123" type="male">♂</specimenCount>
(
<collectionCode id="ED69AE6E0627491AFE4F25D4AB61D94D" box="[511,598,1257,1283]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="4" pageNumber="123" type="Museum">PMNH</collectionCode>
), labelled: “
<collectingCountry id="F36F763B0627491AFD6525D4AA9FD94D" box="[725,936,1257,1283]" name="Bangladesh" pageId="4" pageNumber="123">EAST PAKISTAN</collectingCountry>
/ Dinajpur / XI-70 Barbe’ [p] // Ex Pontificum Institutum / Missionum Exterarum, / Detroit, Michigan; 1984. / Purch. from Carlo Brivio [p] //
<typeStatus id="54C388090627491AFB682430AC41D969" box="[1240,1398,1293,1319]" pageId="4" pageNumber="123" type="paratype">PARATYPUS</typeStatus>
[p, pink label with black frame] //
<taxonomicName id="4C784D280627491AFE40240CABB8D905" authorityName="Erichson" authorityYear="1832" box="[496,655,1329,1355]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="123" phylum="Arthropoda" rank="genus">COPELATUS</taxonomicName>
/ BRIVIOI / N. SP. [hw] /
<determinerName id="7E1BDA6F0627491AFC09240DAD12D905" box="[953,1061,1328,1355]" pageId="4" pageNumber="123">Rocchi S.</determinerName>
det 19 [p] 76 [hw] // YPM - ENT / 500554 [p]”
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736AB0627491AFF772445ABFED9B1" blockId="4.[151,1437,1220,2002]" pageId="4" pageNumber="123">
<emphasis id="B90CEAB90627491AFF772445AB09D9DC" bold="true" box="[199,574,1400,1426]" pageId="4" pageNumber="123">Additional material examined.</emphasis>
<materialsCitation id="3B103CF60627491AFDFA2445AB63D995" collectingDate="2017-07-01" collectionCode="ASCG" collectorName="A. Kopetz" country="Nepal" county="Sudurpashchim Province" elevation="160" latitude="28.8975" location="Mahandranagar" longLatPrecision="20" longitude="80.2275" municipality="Kanchanpur District" pageId="4" pageNumber="123" specimenCount="1" specimenCount-male="1">
<emphasis id="B90CEAB90627491AFDFA2445AB87D9DC" bold="true" box="[586,688,1400,1426]" pageId="4" pageNumber="123">
<collectingCountry id="F36F763B0627491AFDFA2445AB9CD9DC" box="[586,683,1400,1426]" name="Nepal" pageId="4" pageNumber="123">NEPAL</collectingCountry>
:
</emphasis>
<specimenCount id="9D7EFD220627491AFD0C2444ABDBD9DD" box="[700,748,1401,1427]" count="1" pageId="4" pageNumber="123" type="male">1 ♂</specimenCount>
,
<collectingCounty id="62A64E270627491AFD4D2445AD2AD9DC" box="[765,1053,1400,1426]" pageId="4" pageNumber="123">Sudurpashchim Province</collectingCounty>
,
<collectingMunicipality id="6BA3ACD10627491AFB9E2445AC2DD9DC" box="[1070,1306,1400,1426]" pageId="4" pageNumber="123">Kanchanpur District</collectingMunicipality>
, [ca.
<quantity id="4C809B4E0627491AFAEF2444ACABD9DC" box="[1375,1436,1400,1427]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="123" unit="km" value="9.0">9 km</quantity>
SE]
<location id="8EA760700627491AFF7B24A1A84CD9F8" LSID="urn:lsid:plazi:treatment:03D187BD06274918FF272528A89FDCB7:8EA760700627491AFF7B24A1A84CD9F8" box="[203,379,1436,1462]" country="Nepal" county="Sudurpashchim Province" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Kanchanpur District" name="Mahandranagar" pageId="4" pageNumber="123">Mahandranagar</location>
[= Bhimdatta],
<location id="8EA760700627491AFD8824A1AAB9D9F9" LSID="urn:lsid:plazi:treatment:03D187BD06274918FF272528A89FDCB7:8EA760700627491AFD8824A1AAB9D9F9" box="[568,910,1436,1463]" country="Nepal" county="Sudurpashchim Province" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Kanchanpur District" name="Shuklaphanta Nature Reserve" pageId="4" pageNumber="123">Shuklaphanta Nature Reserve</location>
,
<location id="8EA760700627491AFC2C24A1AD06D9F8" LSID="urn:lsid:plazi:treatment:03D187BD06274918FF272528A89FDCB7:8EA760700627491AFC2C24A1AD06D9F8" box="[924,1073,1436,1462]" country="Nepal" county="Sudurpashchim Province" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Kanchanpur District" name="Dsauda river" pageId="4" pageNumber="123">Dsauda river</location>
,
<geoCoordinate id="EE4C506C0627491AFB8F24A1ADFAD9F9" box="[1087,1229,1436,1463]" degrees="28" direction="north" minutes="53" orientation="latitude" pageId="4" pageNumber="123" precision="15" seconds="51" value="28.8975">28°53’51’’N</geoCoordinate>
<geoCoordinate id="EE4C506C0627491AFB6724A0AC54D9F9" box="[1239,1379,1437,1463]" degrees="80" direction="east" minutes="13" orientation="longitude" pageId="4" pageNumber="123" precision="15" seconds="39" value="80.2275">80°13’39’’E</geoCoordinate>
,
<quantity id="4C809B4E0627491AFAC224A1A998D995" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="123" unit="m" value="160.0">
<elevation id="0055D1980627491AFAC224A1A998D995" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="123" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<date id="FFC6106B0627491AFF0E24FDA805D994" box="[190,306,1472,1498]" pageId="4" pageNumber="123" value="2017-07-01">
<collectingDate id="EF82E9830627491AFF0E24FDA805D994" box="[190,306,1472,1498]" pageId="4" pageNumber="123" value="2017-07-01">1.vii.2017</collectingDate>
</date>
,
<collectorName id="268D537D0627491AFE8F24FCA881D995" box="[319,438,1473,1499]" pageId="4" pageNumber="123">A. Kopetz</collectorName>
leg. (
<collectionCode id="ED69AE6E0627491AFE4B24FCAB7BD995" box="[507,588,1473,1499]" pageId="4" pageNumber="123">ASCG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B103CF60627491AFDD224FDABF2D9B1" collectingDate="1985-12-16" collectingDateMax="1985-12-22" collectingDateMin="1985-12-16" collectionCode="NHMB, NMPC" collectorName="B. Bhakta" country="India" elevation="250" location="Bagrakote" municipality="Jalpaiguri District" pageId="4" pageNumber="123" specimenCount="2" specimenCount-male="2" stateProvince="West Bengal">
<emphasis id="B90CEAB90627491AFDD224FDAA5FD995" bold="true" box="[610,872,1472,1499]" pageId="4" pageNumber="123">
<collectingCountry id="F36F763B0627491AFDD224FDAB8ED994" box="[610,697,1472,1498]" name="India" pageId="4" pageNumber="123">INDIA</collectingCountry>
:
<collectingRegion id="49BCF8490627491AFD7724FCAA53D995" box="[711,868,1473,1499]" country="India" name="West Bengal" pageId="4" pageNumber="123">West Bengal</collectingRegion>
:
</emphasis>
<specimenCount id="9D7EFD220627491AFCC324FCAA95D995" box="[883,930,1473,1499]" count="2" pageId="4" pageNumber="123" type="male">2 ♂</specimenCount>
,
<collectingMunicipality id="6BA3ACD10627491AFC0124FDADB6D994" box="[945,1153,1472,1498]" pageId="4" pageNumber="123">Jalpaiguri District</collectingMunicipality>
,
<location id="8EA760700627491AFB3F24FDAC32D994" LSID="urn:lsid:plazi:treatment:03D187BD06274918FF272528A89FDCB7:8EA760700627491AFB3F24FDAC32D994" box="[1167,1285,1472,1498]" country="India" municipality="Jalpaiguri District" name="Bagrakote" pageId="4" pageNumber="123" stateProvince="West Bengal">Bagrakote</location>
,
<quantity id="4C809B4E0627491AFAA324FCACA0D995" box="[1299,1431,1473,1499]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="4" pageNumber="123" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="0055D1980627491AFAA324FCACA0D995" box="[1299,1431,1473,1499]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="4" pageNumber="123" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200–300 m</elevation>
</quantity>
,
<date id="FFC6106B0627491AFF2724D9A87DD9B1" box="[151,330,1508,1535]" pageId="4" pageNumber="123" value="1985-12-16" valueMax="1985-12-22" valueMin="1985-12-16">
<collectingDate id="EF82E9830627491AFF2724D9A87DD9B1" box="[151,330,1508,1535]" pageId="4" pageNumber="123" value="1985-12-16" valueMax="1985-12-22" valueMin="1985-12-16">16.–22.xii.1985</collectingDate>
</date>
,
<collectorName id="268D537D0627491AFEE624D8A8F2D9B0" box="[342,453,1508,1535]" pageId="4" pageNumber="123">B. Bhakta</collectorName>
leg. (
<collectionCode id="ED69AE6E0627491AFDB524D8AB6AD9B1" box="[517,605,1509,1535]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="4" pageNumber="123">NHMB</collectionCode>
,
<collectionCode id="ED69AE6E0627491AFDD824D8AB8AD9B1" box="[616,701,1509,1535]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="123">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36265200627491AFF772735A8E5DB9C" pageId="4" pageNumber="123" type="diagnosis">
<paragraph id="8BC736AB0627491AFF772735A8E5DB9C" blockId="4.[151,1437,1220,2002]" pageId="4" pageNumber="123">
<emphasis id="B90CEAB90627491AFF772735A875DA6C" bold="true" box="[199,322,1544,1570]" pageId="4" pageNumber="123">Diagnosis.</emphasis>
TL:
<quantity id="4C809B4E0627491AFEC82735AB35DA6D" box="[376,514,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.8" metricValueMin="5.3" pageId="4" pageNumber="123" unit="mm" value="5.55" valueMax="5.8" valueMin="5.3">5.3–5.8 mm</quantity>
. Head dark brown, clypeus paler, reddish; pronotum dark brown, laterally reddish, pale colouration extending anteriorly; elytra brown blackish with yellowish sides laterally from dorsal stria 6, and with variable yellowish pattern consisting of: two small basal spots anteriorly from elytral striae 2 and 5 respectively, yellowish stripes in posterior half of elytra between striae 3–5, and yellowish stripe starting at 1/5 or 1/4 of elytral length between striae 5–6. Pronotum with coarse punctures laterally. Each elytron with six discal striae and one submarginal stria: stria 1 beginning approximately at 1/5 of elytral length, ending close to elytral apex; striae 2, 3 and 5 ending subapically; stria 6 shortest, ending at apical fifth; submarginal stria long, beginning at elytral mid-length and ending subapically at level of striae 2, 3 and 5 (
<figureCitation id="13432A2E0627491AFC9E2639AA44DB51" box="[814,883,1796,1823]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="4" pageNumber="123">Fig. 3</figureCitation>
). Male longer spur on metatibia sinuate in apical third. Median lobe of aedeagus in lateral view (
<figureCitation id="13432A2E0627491AFD0E2615AA27DB0D" box="[702,784,1832,1859]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="4" pageNumber="123">Fig. 11</figureCitation>
) broad in basal 2/3; apical third sickle-shaped, regularly curved to obtuse apex; at 2/3 of its length with two distinct ‘teeth’ on ventral side—a broad rectangular proximal one, and a narrower, apically broadened and truncate distal one. Parameres with distinct lobe on inner margin (
<figureCitation id="13432A2E0627491AFADF264DA981DBE1" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="4" pageNumber="123">Fig. 12</figureCitation>
). Female identical to male in habitus, punctation of elytra slightly coarser, disc of elytra laterally from stria 2 with striolae (
<bibRefCitation id="EFE94B5A0627491AFE812685A8F2DB9C" author="Rocchi, S." box="[305,453,1976,2002]" pageId="4" pageNumber="123" pagination="177 - 180" refId="ref7362" refString="Rocchi, S. (1976) Ditiscidi del Bengla-desh con descrizione di Copelatus brivioi n. sp. (Coleoptera Dytiscidae). Bollettino della Societa Entomologica Italiana, 108, 177 - 180." type="journal article" year="1976">Rocchi 1976</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0766230626491BFF282649AABDDBB6" ID-DOI="http://doi.org/10.5281/zenodo.12742766" ID-Zenodo-Dep="12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="5" pageNumber="124" startId="5.[152,270,1908,1932]" targetBox="[195,1393,190,1873]" targetPageId="5" targetType="figure">
<paragraph id="8BC736AB0626491BFF282649AABDDBB6" blockId="5.[151,1437,1908,2041]" pageId="5" pageNumber="124">
<emphasis id="B90CEAB90626491BFF282649A867DBC2" bold="true" box="[152,336,1908,1932]" pageId="5" pageNumber="124">FIGURES 7–18.</emphasis>
Male genitalia of
<taxonomicName id="4C784D280626491BFDA52649ABFEDBC2" authorityName="Sharp" authorityYear="1882" box="[533,713,1908,1932]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B90CEAB90626491BFDA52649ABFEDBC2" box="[533,713,1908,1932]" italics="true" pageId="5" pageNumber="124">Copelatus latipes</emphasis>
</taxonomicName>
species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8.
<taxonomicName id="4C784D280626491BFCB726A4AA46DBFE" authorityName="Sheth, Ghate &amp; Hajek" authorityYear="2018" box="[775,881,1944,1968]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="bezdeki">
<emphasis id="B90CEAB90626491BFCB726A4AA46DBFE" box="[775,881,1944,1968]" italics="true" pageId="5" pageNumber="124">C. bezdeki</emphasis>
</taxonomicName>
(holotype; Maharashtra, India); 9–10.
<taxonomicName id="4C784D280626491BFB4F26A4AC58DBFE" authorityName="Wewalka" authorityYear="1981" box="[1279,1391,1944,1968]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="schereri">
<emphasis id="B90CEAB90626491BFB4F26A4AC58DBFE" box="[1279,1391,1944,1968]" italics="true" pageId="5" pageNumber="124">C. schereri</emphasis>
</taxonomicName>
(Sri Lanka); 11–12.
<taxonomicName id="4C784D280626491BFE852680A8A1DB9A" authorityName="Rocchi" authorityYear="1976" box="[309,406,1980,2004]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="brivioi">
<emphasis id="B90CEAB90626491BFE852680A8A1DB9A" box="[309,406,1980,2004]" italics="true" pageId="5" pageNumber="124">C. brivioi</emphasis>
</taxonomicName>
(paratype; Bangladesh); 13–14.
<taxonomicName id="4C784D280626491BFD6D2680AA75DB9A" authorityName="Hájek &amp; Sheth" authorityYear="2024" box="[733,834,1980,2004]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="kopetzi" status="sp. nov.">
<emphasis id="B90CEAB90626491BFD6D2680AA75DB9A" box="[733,834,1980,2004]" italics="true" pageId="5" pageNumber="124">C. kopetzi</emphasis>
</taxonomicName>
<emphasis id="B90CEAB90626491BFCF72680AAA1DB9B" bold="true" box="[839,918,1981,2005]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A23F57C20626491BFCF72680AAA1DB9B" box="[839,918,1981,2005]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype; Nepal); 15–16.
<taxonomicName id="4C784D280626491BFB162680AC70DB9A" authorityName="Hájek &amp; Sheth" authorityYear="2024" box="[1190,1351,1980,2004]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="lanxangensis" status="sp. nov.">
<emphasis id="B90CEAB90626491BFB162680AC70DB9A" box="[1190,1351,1980,2004]" italics="true" pageId="5" pageNumber="124">C. lanxangensis</emphasis>
</taxonomicName>
<emphasis id="B90CEAB90626491BFAFC2680ACABDB9B" bold="true" box="[1356,1436,1981,2005]" pageId="5" pageNumber="124">
<taxonomicNameLabel id="A23F57C20626491BFAFC2680ACABDB9B" box="[1356,1436,1981,2005]" pageId="5" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype; Laos); 17–18.
<taxonomicName id="4C784D280626491BFE2926DCA8CDDBB6" authorityName="Sharp" authorityYear="1882" box="[409,506,2016,2040]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B90CEAB90626491BFE2926DCA8CDDBB6" box="[409,506,2016,2040]" italics="true" pageId="5" pageNumber="124">C. latipes</emphasis>
</taxonomicName>
(Pahang, Malaysia). Scale bar 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C362652006254918FF7721ABA89FDCB7" pageId="6" pageNumber="125" type="distribution">
<paragraph id="8BC736AB06254918FF7721ABA89FDCB7" blockId="6.[151,1437,150,249]" pageId="6" pageNumber="125">
<emphasis id="B90CEAB906254918FF7721ABA856DCFE" bold="true" box="[199,353,150,176]" pageId="6" pageNumber="125">Distribution.</emphasis>
The species was so far only known from the
<typeStatus id="54C3880906254918FCD021AAAAA7DCFF" box="[864,912,151,177]" pageId="6" pageNumber="125">type</typeStatus>
locality in north-western
<collectingCountry id="F36F763B06254918FB0221AAAC0FDCFF" box="[1202,1336,151,177]" name="Bangladesh" pageId="6" pageNumber="125">Bangladesh</collectingCountry>
; here we present new records from western
<collectingCountry id="F36F763B06254918FD902186AB54DC9B" box="[544,611,187,213]" name="Nepal" pageId="6" pageNumber="125">Nepal</collectingCountry>
and north-eastern
<collectingCountry id="F36F763B06254918FC882186AA45DC9B" box="[824,882,187,213]" name="India" pageId="6" pageNumber="125">India</collectingCountry>
(
<collectingRegion id="49BCF84906254918FC312186AD21DC9B" box="[897,1046,187,213]" country="India" name="West Bengal" pageId="6" pageNumber="125">West Bengal</collectingRegion>
), all from a lowland strip south of the Himalayas (
<figureCitation id="13432A2E06254918FEF721E2A8ACDCB7" box="[327,411,223,249]" captionStart="FIGURE 19" captionStartId="6.[152,255,1288,1312]" captionTargetBox="[156,1437,273,1261]" captionTargetId="figure-66@6.[151,1437,273,1265]" captionTargetPageId="6" captionText="FIGURE 19. Map of distribution of Copelatus latipes species complex." figureDoi="http://doi.org/10.5281/zenodo.12742768" httpUri="https://zenodo.org/record/12742768/files/figure.png" pageId="6" pageNumber="125">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="AE34293025CEC836CA8606CF43AA0F07">A review of the Oriental Copelatus latipes complex, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="40F7C5803B6F7E206AE69A90271E101E">Hájek, Jiří</mods:namePart>
<mods:affiliation id="77731520735FBE0E3F8ABAE063C4D204">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Poc ̌ ernice, Czech Republic</mods:affiliation>
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<mods:namePart id="B18771964866F7214049104601E7808A">Sheth, Sayali D.</mods:namePart>
<mods:affiliation id="24AD4BC5841FE4676176B1F9E23D4C20">Department of Zoology, Prof. Ramkrishna More Arts, Commerce &amp; Science College, Akurdi, Pune - 411044, Maharashtra, India</mods:affiliation>
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<taxonomicName id="4C784D280627491AFF2721AAAB0ADCFF" ID-CoL="Y3FH" authority="Wewalka, 1981" authorityName="Wewalka" authorityYear="1981" box="[151,573,151,178]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="schereri">
<emphasis id="B90CEAB90627491AFF2721AAAB0ADCFF" bold="true" box="[151,573,151,178]" pageId="4" pageNumber="123">
<emphasis id="B90CEAB90627491AFF2721AAA84EDCFC" bold="true" box="[151,377,151,178]" italics="true" pageId="4" pageNumber="123">Copelatus schereri</emphasis>
<bibRefCitation id="EFE94B5A0627491AFE3021AAAB0ADCFF" author="Wewalka, G." box="[384,573,151,177]" pageId="4" pageNumber="123" pagination="65 - 70" refId="ref7577" refString="Wewalka, G. (1981) Drei neue Arten der Gattung Copelatus aus Indien (Dytiscidae, Col.). Koleopterologische Rundschau, 55, 65 - 70." type="journal article" year="1981">Wewalka, 1981</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C36265200627491AFF272186A802DC9B" box="[151,309,187,213]" pageId="4" pageNumber="123" type="description">
<paragraph id="8BC736AB0627491AFF272186A802DC9B" blockId="4.[151,573,151,213]" box="[151,309,187,213]" pageId="4" pageNumber="123">
(
<figureCitation id="13432A2E0627491AFF2F2186A9DFDC9B" box="[159,232,187,213]" captionStart="FIGURES 1–2" captionStartId="2.[152,270,1207,1231]" captionTargetBox="[202,1400,194,1163]" captionTargetId="figure-27@2.[170,1417,181,1183]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Copelatus latipes species complex. 1. C. bezdeki (holotype ♂; Maharashtra, India); 2. C. schereri (♂; Sri Lanka). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742762" httpUri="https://zenodo.org/record/12742762/files/figure.png" pageId="4" pageNumber="123">Figs 2</figureCitation>
,
<figureCitation id="13432A2E0627491AFF442186A81ADC9B" box="[244,301,187,213]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="4" pageNumber="123">9–10</figureCitation>
)
</paragraph>
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<paragraph id="8BC736AB0627491AFF27203DAA24DD56" blockId="4.[151,787,255,280]" box="[151,787,255,280]" pageId="4" pageNumber="123">
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<treatmentCitation id="0AD910BA0627491AFF27203DAB14DD59" author="Wewalka, G." box="[151,547,255,280]" page="65" pageId="4" pageNumber="123" year="1981">
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<emphasis id="B90CEAB90627491AFF27203DA861DD56" box="[151,342,256,280]" italics="true" pageId="4" pageNumber="123">Copelatus schereri</emphasis>
<bibRefCitation id="EFE94B5A0627491AFEEC203DAB14DD59" author="Wewalka, G." box="[348,547,255,280]" pageId="4" pageNumber="123" pagination="65 - 70" refId="ref7577" refString="Wewalka, G. (1981) Drei neue Arten der Gattung Copelatus aus Indien (Dytiscidae, Col.). Koleopterologische Rundschau, 55, 65 - 70." type="journal article" year="1981">Wewalka, 1981: 65</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0AD910BA0627491AFD9D203DAA38DD56" ID-DOI="http://doi.org/10.5281/zenodo.5966537" ID-Zenodo-Dep="5966537" author="Sheth, S. D. &amp; Ghate, H. V. &amp; Hajek, J." box="[557,783,256,280]" httpUri="http://treatment.plazi.org/id/03BC3747FF96FF9CFF5E1716FB1AFEB7" page="248" pageId="4" pageNumber="123" year="2018">
<bibRefCitation id="EFE94B5A0627491AFD9D203DAA38DD56" author="Sheth, S. D. &amp; Ghate, H. V. &amp; Hajek, J." box="[557,783,256,280]" pageId="4" pageNumber="123" pagination="235 - 260" refId="ref7449" refString="Sheth, S. D., Ghate, H. V. &amp; Hajek, J. (2018) Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae). Zootaxa, 4459 (2), 235 - 260. https: // doi. org / 10.11646 / zootaxa. 4459.2.2" type="journal article" year="2018">
Sheth
<emphasis id="B90CEAB90627491AFDDD203DAB96DD56" box="[621,673,256,280]" italics="true" pageId="4" pageNumber="123">et al.</emphasis>
2018: 248
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C36265200627491AFF27207BA800DD82" pageId="4" pageNumber="123" type="materials_examined">
<paragraph id="8BC736AB0627491AFF27207BAA3FDD2E" blockId="4.[151,1437,325,964]" box="[151,776,325,352]" pageId="4" pageNumber="123">
<materialsCitation id="3B103CF60627491AFF27207BAA34DD2E" box="[151,771,325,352]" country="India" location="Shimoga" municipality="Mysore" pageId="4" pageNumber="123" specimenCount="1" stateProvince="Karnataka" typeStatus="holotype">
<emphasis id="B90CEAB90627491AFF27207BA801DD11" bold="true" box="[151,310,325,352]" pageId="4" pageNumber="123">
<typeStatus id="54C388090627491AFF27207BA9E5DD2E" box="[151,210,326,352]" pageId="4" pageNumber="123">Type</typeStatus>
locality.
</emphasis>
“
<collectingCountry id="F36F763B0627491AFEF7207BA8A2DD2E" box="[327,405,326,352]" name="India" pageId="4" pageNumber="123">Indien</collectingCountry>
:
<collectingMunicipality id="6BA3ACD10627491AFE2F207BA8C2DD2E" box="[415,501,326,352]" pageId="4" pageNumber="123">Mysore</collectingMunicipality>
[
<collectingRegion id="49BCF8490627491AFDB2207BAB48DD2E" box="[514,639,326,352]" country="India" name="Karnataka" pageId="4" pageNumber="123">Karnataka</collectingRegion>
]:
<location id="8EA760700627491AFD20207BABCEDD2E" LSID="urn:lsid:plazi:treatment:03D187BD0627491AFF2721AAABE9DF8A:8EA760700627491AFD20207BABCEDD2E" box="[656,761,326,352]" country="India" municipality="Mysore" name="Shimoga" pageId="4" pageNumber="123" stateProvince="Karnataka">Shimoga</location>
”
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736AB0627491AFF772057AB8BDDCA" blockId="4.[151,1437,325,964]" box="[199,700,361,388]" pageId="4" pageNumber="123">
<emphasis id="B90CEAB90627491AFF772057A842DDCD" bold="true" box="[199,373,361,388]" pageId="4" pageNumber="123">
<typeStatus id="54C388090627491AFF772057A835DDCA" box="[199,258,362,388]" pageId="4" pageNumber="123">Type</typeStatus>
material.
</emphasis>
See
<bibRefCitation id="EFE94B5A0627491AFE1B2057AB80DDCA" author="Sheth, S. D. &amp; Ghate, H. V. &amp; Hajek, J." box="[427,695,362,388]" pageId="4" pageNumber="123" pagination="235 - 260" refId="ref7449" refString="Sheth, S. D., Ghate, H. V. &amp; Hajek, J. (2018) Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae). Zootaxa, 4459 (2), 235 - 260. https: // doi. org / 10.11646 / zootaxa. 4459.2.2" type="journal article" year="2018">
Sheth
<emphasis id="B90CEAB90627491AFE422056AB1DDDCA" box="[498,554,362,388]" italics="true" pageId="4" pageNumber="123">et al.</emphasis>
(2018: 248)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BC736AB0627491AFF7720B0A800DD82" blockId="4.[151,1437,325,964]" pageId="4" pageNumber="123">
<emphasis id="B90CEAB90627491AFF7720B0AB0FDDE9" bold="true" box="[199,568,397,423]" pageId="4" pageNumber="123">Additional material examined.</emphasis>
<materialsCitation id="3B103CF60627491AFDF020B0A804DD82" collectionCode="GWCV" collectorName="Ceylon &amp; Colombo Umg. &amp; M. Jach" country="Sri Lanka" location="SRI LANKA" pageId="4" pageNumber="123" specimenCount="1" specimenCount-male="1">
<emphasis id="B90CEAB90627491AFDF020B0ABD1DDE9" bold="true" box="[576,742,397,423]" pageId="4" pageNumber="123">
<collectingCountry id="F36F763B0627491AFDF020B0ABD6DDE9" box="[576,737,397,423]" name="Sri Lanka" pageId="4" pageNumber="123">SRI LANKA</collectingCountry>
:
</emphasis>
<specimenCount id="9D7EFD220627491AFD5F20B3AA2ADDE6" box="[751,797,398,424]" count="1" pageId="4" pageNumber="123" type="male">1 ♂</specimenCount>
,
<collectorName id="268D537D0627491AFC9B20B3AA48DDE6" box="[811,895,398,424]" pageId="4" pageNumber="123">Ceylon</collectorName>
,
<collectorName id="268D537D0627491AFC3B20B3AD0BDDE6" box="[907,1084,398,424]" pageId="4" pageNumber="123">Colombo Umg.</collectorName>
[= env.], 8.xi. [19]80,
<collectorName id="268D537D0627491AFAF220B3ACABDDE6" box="[1346,1436,398,424]" pageId="4" pageNumber="123">M. Jäch</collectorName>
leg. (
<collectionCode id="ED69AE6E0627491AFF7F208FA81CDD82" box="[207,299,434,460]" pageId="4" pageNumber="123">GWCV</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36265200627491AFF7720E8AC3DDF32" pageId="4" pageNumber="123" type="diagnosis">
<paragraph id="8BC736AB0627491AFF7720E8AC3DDF32" blockId="4.[151,1437,325,964]" pageId="4" pageNumber="123">
<emphasis id="B90CEAB90627491AFF7720E8A875DDA1" bold="true" box="[199,322,469,495]" pageId="4" pageNumber="123">Diagnosis.</emphasis>
TL: 5.0–
<quantity id="4C809B4E0627491AFE1A20EBAB35DDBE" box="[426,514,470,497]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="4" pageNumber="123" unit="mm" value="5.4">5.4 mm</quantity>
. Head reddish brown, darkened between eyes; pronotum dark brown, sides largely reddish; elytra yellowish orange with blackish pattern varying from vaguely delimited infuscation on disc between suture and elytral stria 2 and narrow blackish stripes along elytral striae, to presence of blackish subbasal transverse band extending somewhat posteriad between striae 4–6 and black sutural area between suture and stria 2. Pronotum with coarse punctures laterally, sometimes forming short striolae. Each elytron with six complete discal striae and one submarginal stria: stria 1 beginning more posteriorly than other striae, ending close to elytral apex; striae 2, 3 and 5 ending subapically; stria 6 shortest, ending at apical fourth; submarginal stria long, beginning at elytral mid-length and ending at apical fourth (
<figureCitation id="13432A2E0627491AFD9823EFAB58DEA2" box="[552,623,722,748]" captionStart="FIGURES 1–2" captionStartId="2.[152,270,1207,1231]" captionTargetBox="[202,1400,194,1163]" captionTargetId="figure-27@2.[170,1417,181,1183]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Copelatus latipes species complex. 1. C. bezdeki (holotype ♂; Maharashtra, India); 2. C. schereri (♂; Sri Lanka). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742762" httpUri="https://zenodo.org/record/12742762/files/figure.png" pageId="4" pageNumber="123">Fig. 2</figureCitation>
). Longer spur on metatibia almost straight. Median lobe of aedeagus in lateral view (
<figureCitation id="13432A2E0627491AFE9B23CBA84DDF5E" box="[299,378,758,784]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="4" pageNumber="123">Fig. 9a</figureCitation>
) broad in basal 3/5, narrow, almost regularly curved in apical half, with shallow emargination on ventral side before apex; apex beak-shaped; median lobe in 3/5 with two distinct ‘teeth’ on ventral side— broad rectangular proximal one, and narrower acute distal one; third, small, obtuse tooth visible in ventro-lateral view (
<figureCitation id="13432A2E0627491AFF2F225FA9C4DF32" box="[159,243,866,892]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="4" pageNumber="123">Fig. 9b</figureCitation>
). Parameres with distinct serration on inner margin in apical half (
<figureCitation id="13432A2E0627491AFC69225FAD1BDF32" box="[985,1068,866,892]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="4" pageNumber="123">Fig. 10</figureCitation>
). Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36265200627491AFF7722B8ABE9DF8A" pageId="4" pageNumber="123" type="distribution">
<paragraph id="8BC736AB0627491AFF7722B8ABE9DF8A" blockId="4.[151,1437,325,964]" pageId="4" pageNumber="123">
<emphasis id="B90CEAB90627491AFF7722B8A856DFD1" bold="true" box="[199,353,901,927]" pageId="4" pageNumber="123">Distribution.</emphasis>
A species previously known only from Western Ghats (
<collectingRegion id="49BCF8490627491AFC7522BBAD0DDFEE" box="[965,1082,902,928]" country="India" name="Karnataka" pageId="4" pageNumber="123">Karnataka</collectingRegion>
and
<collectingRegion id="49BCF8490627491AFBDE22BBADCBDFEE" box="[1134,1276,902,928]" country="India" name="Maharashtra" pageId="4" pageNumber="123">Maharashtra</collectingRegion>
) in
<collectingCountry id="F36F763B0627491AFA9522BBAC55DFEE" box="[1317,1378,902,928]" name="India" pageId="4" pageNumber="123">India</collectingCountry>
; here we present the first record from
<collectingCountry id="F36F763B0627491AFDB02297AB59DF8A" box="[512,622,938,964]" name="Sri Lanka" pageId="4" pageNumber="123">Sri Lanka</collectingCountry>
(
<figureCitation id="13432A2E0627491AFDCD2297ABE7DF8A" box="[637,720,938,964]" captionStart="FIGURE 19" captionStartId="6.[152,255,1288,1312]" captionTargetBox="[156,1437,273,1261]" captionTargetId="figure-66@6.[151,1437,273,1265]" captionTargetPageId="6" captionText="FIGURE 19. Map of distribution of Copelatus latipes species complex." figureDoi="http://doi.org/10.5281/zenodo.12742768" httpUri="https://zenodo.org/record/12742768/files/figure.png" pageId="4" pageNumber="123">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

303
data/03/D1/87/03D187BD062A4917FF272223A8E6DBFD.xml Normal file
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@ -0,0 +1,303 @@
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<mods:title id="39EB79D885FF160BB7BFB6E19F29A477">A review of the Oriental Copelatus latipes complex, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<paragraph id="8BC736AB062A4917FF272223AB34DF76" blockId="9.[151,515,798,861]" box="[151,515,798,825]" pageId="9" pageNumber="128">
<heading id="D08F81C7062A4917FF272223AB34DF76" bold="true" box="[151,515,798,825]" fontSize="11" level="1" pageId="9" pageNumber="128" reason="1">
<taxonomicName id="4C784D28062A4917FF272223AB34DF76" ID-CoL="Y3B3" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[151,515,798,825]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B90CEAB9062A4917FF272223AB34DF76" bold="true" box="[151,515,798,825]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FF272223A852DF77" bold="true" box="[151,357,798,825]" italics="true" pageId="9" pageNumber="128">Copelatus latipes</emphasis>
<bibRefCitation id="EFE94B5A062A4917FEDD2223AB34DF76" author="Sharp, D." box="[365,515,798,824]" pageId="9" pageNumber="128" pagination="179 - 1003" refId="ref7398" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18. https: // doi. org / 10.5962 / bhl. title. 9530" type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3626520062A4917FF27227EA874DF13" box="[151,323,835,861]" pageId="9" pageNumber="128" type="description">
<paragraph id="8BC736AB062A4917FF27227EA874DF13" blockId="9.[151,515,798,861]" box="[151,323,835,861]" pageId="9" pageNumber="128">
(
<figureCitation id="13432A2E062A4917FF2F227EA9DFDF13" box="[159,232,835,861]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="9" pageNumber="128">Figs 6</figureCitation>
,
<figureCitation id="13432A2E062A4917FF44227EA80DDF13" box="[244,314,835,861]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="9" pageNumber="128">17–18</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BC736AB062A4917FF2722BAAA82DFD1" blockId="9.[151,949,903,927]" box="[151,949,903,927]" pageId="9" pageNumber="128">
<treatmentCitationGroup id="AB681185062A4917FF2722BAAA82DFD1" box="[151,949,903,927]" pageId="9" pageNumber="128">
<treatmentCitation id="0AD910BA062A4917FF2722BAA8C8DFD1" author="Sharp, D." box="[151,511,903,927]" page="580" pageId="9" pageNumber="128" year="1882">
<taxonomicName id="4C784D28062A4917FF2722BAA8C8DFD1" ID-CoL="Y3B3" authority="Sharp, 1882: 580" authorityName="Sharp" authorityPageNumber="580" authorityYear="1882" box="[151,511,903,927]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B90CEAB9062A4917FF2722BAA87FDFD1" box="[151,328,903,927]" italics="true" pageId="9" pageNumber="128">Copelatus latipes</emphasis>
<bibRefCitation id="EFE94B5A062A4917FEFE22BAA8C8DFD1" author="Sharp, D." box="[334,511,903,927]" pageId="9" pageNumber="128" pagination="179 - 1003" refId="ref7398" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18. https: // doi. org / 10.5962 / bhl. title. 9530" type="journal article" year="1882">Sharp, 1882: 580</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AD910BA062A4917FDB922BAABD3DFD1" author="Regimbart, M." box="[521,740,903,927]" page="296" pageId="9" pageNumber="128" year="1899">
<bibRefCitation id="EFE94B5A062A4917FDB922BAABD3DFD1" author="Regimbart, M." box="[521,740,903,927]" pageId="9" pageNumber="128" pagination="186 - 367" refId="ref7333" refString="Regimbart, M. (1899) Revision des Dytiscidae de la region Indo-Sino-Malaise. Annales de la Societe Entomologique de France, 68, 186 - 367." type="journal article" year="1899">Régimbart 1899: 296</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD910BA062A4917FD5E22BAAA86DFD1" author="Vazirani, T. G." box="[750,945,903,927]" page="307" pageId="9" pageNumber="128" year="1970">
<bibRefCitation id="EFE94B5A062A4917FD5E22BAAA86DFD1" author="Vazirani, T. G." box="[750,945,903,927]" pageId="9" pageNumber="128" pagination="303 - 362" refId="ref7524" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4 (3), 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article" year="1970">Vazirani 1970: 307</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3626520062A4917FF2722F0ABA3D965" pageId="9" pageNumber="128" type="materials_examined">
<paragraph id="8BC736AB062A4917FF2722F0ABDFDFA9" blockId="9.[151,1437,973,1971]" box="[151,744,973,999]" pageId="9" pageNumber="128">
<materialsCitation id="3B103CF6062A4917FF2722F0ABD3DFA9" box="[151,740,973,999]" country="Malaysia, Malacca" location="Malaysia, Malacca" pageId="9" pageNumber="128" specimenCount="1" typeStatus="holotype">
<emphasis id="B90CEAB9062A4917FF2722F0A801DFA9" bold="true" box="[151,310,973,999]" pageId="9" pageNumber="128">
<typeStatus id="54C38809062A4917FF2722F0A9E5DFA9" box="[151,210,973,999]" pageId="9" pageNumber="128">Type</typeStatus>
locality.
</emphasis>
“Malacca” [
<collectingCountry id="F36F763B062A4917FE7122F0ABAFDFA9" box="[449,664,973,999]" isEnumeration="true" pageId="9" pageNumber="128">Malaysia, Malacca</collectingCountry>
State]
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736AB062A4917FF7722CCA9D7D81D" blockId="9.[151,1437,973,1971]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FF7722CCA842D845" bold="true" box="[199,373,1009,1035]" pageId="9" pageNumber="128">Type material.</emphasis>
<materialsCitation id="3B103CF6062A4917FECC22CFA9D7D81D" collectionCode="BMNH" country="Malaysia" location="Type" municipality="Type" pageId="9" pageNumber="128" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus id="54C38809062A4917FECC22CFA8DBD842" box="[380,492,1010,1036]" pageId="9" pageNumber="128" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D7EFD22062A4917FE4422CFAB3ED842" box="[500,521,1010,1036]" count="1" pageId="9" pageNumber="128" type="male">♂</specimenCount>
(designated by
<bibRefCitation id="EFE94B5A062A4917FD0B22CCAAA3D845" author="Vazirani, T. G." box="[699,916,1009,1035]" pageId="9" pageNumber="128" pagination="303 - 362" refId="ref7524" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4 (3), 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article" year="1970">Vazirani 1970: 307</bibRefCitation>
;
<collectionCode id="ED69AE6E062A4917FC1022CCAACFD845" box="[928,1016,1009,1035]" pageId="9" pageNumber="128">BMNH</collectionCode>
), labelled: “
<location id="8EA76070062A4917FBCD22CCAD80D845" LSID="urn:lsid:plazi:treatment:03D187BD062A4917FF272223A8E6DBFD:8EA76070062A4917FBCD22CCAD80D845" box="[1149,1207,1009,1035]" country="Malaysia" municipality="Type" name="Type" pageId="9" pageNumber="128">Type</location>
[p, round label with red frame] //
<collectingCountry id="F36F763B062A4917FE812528A8A7D861" box="[305,400,1045,1071]" name="Malaysia" pageId="9" pageNumber="128">Malacca</collectingCountry>
/ 691 [hw] // Sharp Coll. / 1905-313. [p] //
<collectingMunicipality id="6BA3ACD1062A4917FC3D252BAAF2D87E" box="[909,965,1046,1072]" pageId="9" pageNumber="128">Type</collectingMunicipality>
691 /
<taxonomicName id="4C784D28062A4917FBA22528ADB5D861" authorityName="Erichson" authorityYear="1832" box="[1042,1154,1045,1071]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
/ latipes
<taxonomicNameLabel id="A23F57C2062A4917FB5E252BAC14D87E" box="[1262,1315,1046,1072]" pageId="9" pageNumber="128" rank="species">n.sp.</taxonomicNameLabel>
/
<collectingCountry id="F36F763B062A4917FA8D2528ACABD861" box="[1341,1436,1045,1071]" name="Malaysia" pageId="9" pageNumber="128">Malacca</collectingCountry>
[hw]”.
</materialsCitation>
</paragraph>
<paragraph id="8BC736AB062A4917FF772560ABA3D965" blockId="9.[151,1437,973,1971]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FF772560AB0CD839" bold="true" box="[199,571,1117,1143]" pageId="9" pageNumber="128">Additional material examined.</emphasis>
<materialsCitation id="3B103CF6062A4917FDF52560A9CAD8D5" collectingDate="2002-11-01" collectingDateMax="2002-11-11" collectingDateMin="2002-11-01" collectionCode="NMPC" collectorName="P. Kocarek" country="Cambodia" location="Siem Reap" pageId="9" pageNumber="128" specimenCount="1" specimenCount-female="1" stateProvince="Siem Reap">
<emphasis id="B90CEAB9062A4917FDF52560ABDAD839" bold="true" box="[581,749,1117,1143]" pageId="9" pageNumber="128">
<collectingCountry id="F36F763B062A4917FDF52560ABDFD839" box="[581,744,1117,1143]" name="Cambodia" pageId="9" pageNumber="128">CAMBODIA</collectingCountry>
:
</emphasis>
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,
<collectingRegion id="49BCF849062A4917FC872560AA80D836" box="[823,951,1117,1144]" country="Cambodia" name="Siem Reap" pageId="9" pageNumber="128">Siem Reap</collectingRegion>
,
<collectingMethod id="52394EBC062A4917FC762560AD05D836" box="[966,1074,1117,1144]" pageId="9" pageNumber="128">light trap</collectingMethod>
,
<date id="FFC6106B062A4917FBF12560ADEBD839" box="[1089,1244,1117,1143]" pageId="9" pageNumber="128" value="2002-11-01" valueMax="2002-11-11" valueMin="2002-11-01">
<collectingDate id="EF82E983062A4917FBF12560ADEBD839" box="[1089,1244,1117,1143]" pageId="9" pageNumber="128" value="2002-11-01" valueMax="2002-11-11" valueMin="2002-11-01">1.–11.xi.2002</collectingDate>
</date>
,
<collectorName id="268D537D062A4917FB5C2563AC5ED839" box="[1260,1385,1117,1144]" pageId="9" pageNumber="128">P. Kočárek</collectorName>
leg. (
<collectionCode id="ED69AE6E062A4917FF2F25BCA9C2D8D5" box="[159,245,1153,1179]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="128">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B103CF6062A4917FEB825BCAB08D8F1" collectingDate="2001-05-22" collectingDateMax="2001-06-08" collectingDateMin="2001-05-22" collectionCode="NMPC" collectorName="E. Jendek &amp; O. Sausa" country="Laos" elevation="550" latitude="17.716667" location="Nakai" longLatPrecision="1277" longitude="105.15" pageId="9" pageNumber="128" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Khammouan">
<emphasis id="B90CEAB9062A4917FEB825BCA86AD8D5" bold="true" box="[264,349,1153,1179]" pageId="9" pageNumber="128">
<collectingCountry id="F36F763B062A4917FEB825BCA86FD8D5" box="[264,344,1153,1179]" name="Laos" pageId="9" pageNumber="128">LAOS</collectingCountry>
:
</emphasis>
<specimenCount id="9D7EFD22062A4917FED525BCA8A5D8D2" box="[357,402,1153,1180]" count="1" pageId="9" pageNumber="128" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D7EFD22062A4917FE2F25BCA8FED8D2" box="[415,457,1153,1180]" count="2" pageId="9" pageNumber="128" type="female">2 ♀</specimenCount>
,
<collectingRegion id="49BCF849062A4917FE6825BCABD4D8D5" box="[472,739,1153,1179]" country="Laos" name="Khammouan" pageId="9" pageNumber="128">Khammouane Province</collectingRegion>
,
<location id="8EA76070062A4917FD5F25BCAA05D8D5" LSID="urn:lsid:plazi:treatment:03D187BD062A4917FF272223A8E6DBFD:8EA76070062A4917FD5F25BCAA05D8D5" box="[751,818,1153,1179]" country="Laos" latitude="17.716667" longLatPrecision="1277" longitude="105.15" name="Nakai" pageId="9" pageNumber="128" stateProvince="Khammouan">Nakai</location>
env.,
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<geoCoordinate id="EE4C506C062A4917FC6F25BCAD7AD8D2" box="[991,1101,1153,1180]" degrees="105" direction="east" minutes="09" orientation="longitude" pageId="9" pageNumber="128" precision="925" value="105.15">105°09’E</geoCoordinate>
,
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<elevation id="0055D198062A4917FBE925BCADECD8D2" box="[1113,1243,1153,1180]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="9" pageNumber="128" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500–600 m</elevation>
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,
<date id="FFC6106B062A4917FB5725BCACAFD8D5" box="[1255,1432,1153,1179]" pageId="9" pageNumber="128" value="2001-05-22" valueMax="2001-06-08" valueMin="2001-05-22">
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</date>
,
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&amp;
<collectorName id="268D537D062A4917FE832598A8ADD88C" box="[307,410,1189,1218]" pageId="9" pageNumber="128">O. Šauša</collectorName>
leg. (
<collectionCode id="ED69AE6E062A4917FE512598AB00D8F1" box="[481,567,1189,1215]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="128">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B103CF6062A4917FDFD2598ACAED8AD" collectingDate="2011-12-06" collectionCode="NMPC" collectorName="PT Silva Rimba Lestari &amp; J. Hajek &amp; J. Schneider &amp; P. Votruba" country="Indonesia" elevation="60" latitude="0.123333335" location="Limbang" longLatPrecision="130" longitude="116.30167" pageId="9" pageNumber="128" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Utara">
<emphasis id="B90CEAB9062A4917FDFD2598ABC2D8F1" bold="true" box="[589,757,1189,1215]" pageId="9" pageNumber="128">
<collectingCountry id="F36F763B062A4917FDFD2598ABC7D8F1" box="[589,752,1189,1215]" name="Indonesia" pageId="9" pageNumber="128">INDONESIA</collectingCountry>
:
</emphasis>
<specimenCount id="9D7EFD22062A4917FCB02598AA18D88E" box="[768,815,1189,1216]" count="1" pageId="9" pageNumber="128" type="male">1 ♂</specimenCount>
, E
<collectingRegion id="49BCF849062A4917FCEB2598AAD6D8F1" box="[859,993,1189,1215]" country="Indonesia" name="Kalimantan Utara" pageId="9" pageNumber="128">Kalimantan</collectingRegion>
,
<collectorName id="268D537D062A4917FC5F259BAC35D8F1" box="[1007,1282,1189,1216]" pageId="9" pageNumber="128">PT Silva Rimba Lestari</collectorName>
[area], camp “
<location id="8EA76070062A4917FF1125F4A83AD8AD" LSID="urn:lsid:plazi:treatment:03D187BD062A4917FF272223A8E6DBFD:8EA76070062A4917FF1125F4A83AD8AD" box="[161,269,1225,1251]" country="Indonesia" latitude="0.123333335" longLatPrecision="130" longitude="116.30167" name="Limbang" pageId="9" pageNumber="128" stateProvince="Kalimantan Utara">Limbang</location>
”,
<geoCoordinate id="EE4C506C062A4917FE9725F4A8AAD8AD" box="[295,413,1225,1251]" degrees="00" direction="north" minutes="07.4" orientation="latitude" pageId="9" pageNumber="128" precision="92" value="0.123333335">00°07.4’N</geoCoordinate>
<geoCoordinate id="EE4C506C062A4917FE1825F4AB1ED8AD" box="[424,553,1225,1251]" degrees="116" direction="east" minutes="18.1" orientation="longitude" pageId="9" pageNumber="128" precision="92" value="116.30167">116°18.1’E</geoCoordinate>
,
<quantity id="4C809B4E062A4917FD8A25F4AB4ED8AA" box="[570,633,1225,1252]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="128" unit="m" value="60.0">
<elevation id="0055D198062A4917FD8A25F4AB4ED8AA" box="[570,633,1225,1252]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="128" unit="m" value="60.0">60 m</elevation>
</quantity>
, 3.–4.+
<date id="FFC6106B062A4917FD7E25F4AA74D8AD" box="[718,835,1225,1251]" pageId="9" pageNumber="128" value="2011-12-06">
<collectingDate id="EF82E983062A4917FD7E25F4AA74D8AD" box="[718,835,1225,1251]" pageId="9" pageNumber="128" value="2011-12-06">6.xii.2011</collectingDate>
</date>
,
<collectorName id="268D537D062A4917FCE325F7AA81D8AD" box="[851,950,1225,1252]" pageId="9" pageNumber="128">J. Hájek</collectorName>
,
<collectorName id="268D537D062A4917FC7525F7AD65D8AD" box="[965,1106,1225,1252]" pageId="9" pageNumber="128">J. Schneider</collectorName>
&amp;
<collectorName id="268D537D062A4917FBCE25F7ADC3D8AD" box="[1150,1268,1225,1252]" pageId="9" pageNumber="128">P. Votruba</collectorName>
leg. (
<collectionCode id="ED69AE6E062A4917FA8B25F4ACA6D8AD" box="[1339,1425,1225,1251]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="128">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B103CF6062A4917FF2725D0ABA7D965" collectingDate="2008-04-09" collectingDateMax="2008-04-30" collectingDateMin="2008-04-09" collectionCode="NMPC" collectorName="G. Keriung &amp; Z. Linek" country="Malaysia" elevation="400" location="Kg. Tebu Hitnam" municipality="Endau Rompin" pageId="9" pageNumber="128" specimenCount="24" specimenCount-female="12" specimenCount-male="12" stateProvince="Pahang">
<emphasis id="B90CEAB9062A4917FF2725D0A800D949" bold="true" box="[151,311,1261,1287]" pageId="9" pageNumber="128">
<collectingCountry id="F36F763B062A4917FF2725D0A806D949" box="[151,305,1261,1287]" name="Malaysia" pageId="9" pageNumber="128">MALAYSIA</collectingCountry>
:
</emphasis>
<specimenCount id="9D7EFD22062A4917FE8E25D0A84FD946" box="[318,376,1261,1288]" count="12" pageId="9" pageNumber="128" type="male">12 ♂</specimenCount>
,
<specimenCount id="9D7EFD22062A4917FE3525D0A88AD946" box="[389,445,1261,1288]" count="12" pageId="9" pageNumber="128" type="female">12 ♀</specimenCount>
,
<collectingRegion id="49BCF849062A4917FE7B25D0AB17D949" box="[459,544,1261,1287]" country="Malaysia" name="Pahang" pageId="9" pageNumber="128">Pahang</collectingRegion>
,
<locationDeviation id="9A752F89062A4917FD9C25D0AA78D949" box="[556,847,1261,1288]" location="Kuala Rompin" pageId="9" pageNumber="128">
<quantity id="4C809B4E062A4917FD9C25D0AB43D949" box="[556,628,1261,1288]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="128" unit="km" value="50.0">50 km</quantity>
NE Kuala Rompin
</locationDeviation>
,
<collectingMunicipality id="6BA3ACD1062A4917FCEA25D0AD34D949" box="[858,1027,1261,1287]" pageId="9" pageNumber="128">Endau Rompin</collectingMunicipality>
NP,
<collectorName id="268D537D062A4917FB8A25D0AD8FD949" box="[1082,1208,1261,1287]" pageId="9" pageNumber="128">G. Keriung</collectorName>
(
<location id="8EA76070062A4917FB7825D0ACA7D949" LSID="urn:lsid:plazi:treatment:03D187BD062A4917FF272223A8E6DBFD:8EA76070062A4917FB7825D0ACA7D949" box="[1224,1424,1261,1287]" country="Malaysia" municipality="Endau Rompin" name="Kg. Tebu Hitnam" pageId="9" pageNumber="128" stateProvince="Pahang">Kg. Tebu Hitnam</location>
),
<quantity id="4C809B4E062A4917FF27242CA9D7D962" box="[151,224,1297,1324]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="128" unit="m" value="400.0">
<elevation id="0055D198062A4917FF27242CA9D7D962" box="[151,224,1297,1324]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="128" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="FFC6106B062A4917FF5C242CA8BFD965" box="[236,392,1297,1323]" pageId="9" pageNumber="128" value="2008-04-09" valueMax="2008-04-30" valueMin="2008-04-09">
<collectingDate id="EF82E983062A4917FF5C242CA8BFD965" box="[236,392,1297,1323]" pageId="9" pageNumber="128" value="2008-04-09" valueMax="2008-04-30" valueMin="2008-04-09">9.–30.iv.2008</collectingDate>
</date>
,
<collectorName id="268D537D062A4917FE23242FA8C4D965" box="[403,499,1297,1324]" pageId="9" pageNumber="128">Z. Linek</collectorName>
leg. (
<collectionCode id="ED69AE6E062A4917FD82242CABBFD965" box="[562,648,1297,1323]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="128">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3626520062A4917FF772408AA69DB6D" pageId="9" pageNumber="128" type="diagnosis">
<paragraph id="8BC736AB062A4917FF772408ABD2DA95" blockId="9.[151,1437,973,1971]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FF772408A875D901" bold="true" box="[199,322,1333,1359]" pageId="9" pageNumber="128">Diagnosis.</emphasis>
TL: 5.1–6.0 mm. Head black, clypeus paler, reddish; pronotum black, laterally brownish, pale colouration extending anteriorly; elytra black with reddish apex and sides laterally from elytral striae, and indistinct reddish pattern consisting of: two basal spots between striae 2–3 and 4–6, and longitudinal spot in middle third of elytra between striae 4–5. Pronotum with coarse punctures laterally. Each elytron with six discal striae and one submarginal stria: stria 1 largely absent, present only in apical fifth of elytral length; striae 2, 3 and 5 ending subapically; stria 6 shortest, ending at apical fifth; submarginal stria long, beginning at elytral mid-length and ending subapically at level of striae 2, 3 and 5 (
<figureCitation id="13432A2E062A4917FD122730ABD2DA69" box="[674,741,1549,1575]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="9" pageNumber="128">Fig. 6</figureCitation>
). Male longer spur on metatibia weakly sinuate in apical third. Median lobe of aedeagus in lateral view (
<figureCitation id="13432A2E062A4917FDDB270CAB8ADA05" box="[619,701,1585,1611]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="9" pageNumber="128">Fig. 17</figureCitation>
) broad and parallel-sided in basal 2/3; apical third sickle-shaped, almost regularly curved in middle, apex bifid; in 2/3 of its length with two distinct ‘teeth’ on ventral side—broad acute proximal one, and narrower, incised, apically obtuse distal one. Parameres with distinct lobe on inner margin (
<figureCitation id="13432A2E062A4917FF2F27A0A9C5DAF9" box="[159,242,1693,1719]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="9" pageNumber="128">Fig. 18</figureCitation>
). Females identical to males in habitus, punctation of elytra slightly coarser, weakly impressed short striolae present in middle part of elytra laterally from stria 3.
</paragraph>
<paragraph id="8BC736AB062A4917FF7727D8AA69DB6D" blockId="9.[151,1437,973,1971]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FF7727D8A8CCDAB1" bold="true" box="[199,507,1765,1791]" pageId="9" pageNumber="128">Collecting circumstances.</emphasis>
The single specimen from
<collectingRegion id="49BCF849062A4917FC8E27D8AAF6DAB1" box="[830,961,1765,1791]" country="Indonesia" name="Kalimantan Utara" pageId="9" pageNumber="128">Kalimantan</collectingRegion>
was collected in a shallow pool in open oligotrophic wetland on sandy ground, surrounded with shrubs.
</paragraph>
</subSubSection>
<subSubSection id="C3626520062A4917FF772610A8E6DBFD" pageId="9" pageNumber="128" type="distribution">
<paragraph id="8BC736AB062A4917FF772610A8E6DBFD" blockId="9.[151,1437,973,1971]" pageId="9" pageNumber="128">
<emphasis id="B90CEAB9062A4917FF772610A856DB09" bold="true" box="[199,353,1837,1863]" pageId="9" pageNumber="128">Distribution.</emphasis>
The species was so far known only from Peninsular
<collectingCountry id="F36F763B062A4917FC192610AD24DB09" box="[937,1043,1837,1863]" name="Malaysia" pageId="9" pageNumber="128">Malaysia</collectingCountry>
. Here we present new records from central
<collectingCountry id="F36F763B062A4917FF5A266FA814DB22" box="[234,291,1874,1900]" name="Laos" pageId="9" pageNumber="128">Laos</collectingCountry>
,
<collectingCountry id="F36F763B062A4917FE9F266CA895DB25" box="[303,418,1873,1899]" name="Cambodia" pageId="9" pageNumber="128">Cambodia</collectingCountry>
and Indonesian part of Borneo Island, documenting that
<taxonomicName id="4C784D28062A4917FB90266FADBBDB25" authorityName="Sharp" authorityYear="1882" box="[1056,1164,1873,1900]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="latipes">
<emphasis id="B90CEAB9062A4917FB90266FADBBDB25" box="[1056,1164,1873,1900]" italics="true" pageId="9" pageNumber="128">C. latipes</emphasis>
</taxonomicName>
is relatively widespread in Southeast Asia (
<figureCitation id="13432A2E062A4917FEC32648A8FDDBC1" box="[371,458,1909,1935]" captionStart="FIGURE 19" captionStartId="6.[152,255,1288,1312]" captionTargetBox="[156,1437,273,1261]" captionTargetId="figure-66@6.[151,1437,273,1265]" captionTargetPageId="6" captionText="FIGURE 19. Map of distribution of Copelatus latipes species complex." figureDoi="http://doi.org/10.5281/zenodo.12742768" httpUri="https://zenodo.org/record/12742768/files/figure.png" pageId="9" pageNumber="128">Fig. 19</figureCitation>
).
<bibRefCitation id="EFE94B5A062A4917FE522648AB9ADBC1" author="Regimbart, M." box="[482,685,1909,1935]" pageId="9" pageNumber="128" pagination="186 - 367" refId="ref7333" refString="Regimbart, M. (1899) Revision des Dytiscidae de la region Indo-Sino-Malaise. Annales de la Societe Entomologique de France, 68, 186 - 367." type="journal article" year="1899">Régimbart (1899)</bibRefCitation>
and
<bibRefCitation id="EFE94B5A062A4917FD5A2648AAEFDBC1" author="Zimmermann, A." box="[746,984,1909,1935]" pageId="9" pageNumber="128" pagination="68 - 249" refId="ref7608" refString="Zimmermann, A. (1919) Die Schwimmkafer des Deutschen Entomologischen Museums in Berlin-Dahlem. Archiv fur Naturgeschichte, Abteilung A, 83 (12), 68 - 249. [1917]" type="journal article" year="1919">Zimmermann (1919)</bibRefCitation>
mentioned also
<collectingCountry id="F36F763B062A4917FB2B2648ADE0DBC1" box="[1179,1239,1909,1935]" name="India" pageId="9" pageNumber="128">India</collectingCountry>
, which was later doubted by
<bibRefCitation id="EFE94B5A062A4917FEAB26A4A8FADBFD" author="Vazirani, T. G." box="[283,461,1945,1971]" pageId="9" pageNumber="128" pagination="303 - 362" refId="ref7524" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4 (3), 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article" year="1970">Vazirani (1970)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>