303 lines
35 KiB
XML
303 lines
35 KiB
XML
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<mods:title id="39EB79D885FF160BB7BFB6E19F29A477">A review of the Oriental Copelatus latipes complex, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="1975861FC8F28215A947A8CF3786FFB5">Hájek, Jiří</mods:namePart>
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<mods:affiliation id="490E31DEEACB742B93BBEFF090553810">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Poc ̌ ernice, Czech Republic</mods:affiliation>
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<mods:namePart id="E334652726E89C0BF48B5948BA680941">Sheth, Sayali D.</mods:namePart>
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<mods:affiliation id="3EADC1D7AAA85700EAC51ACA7BF4B182">Department of Zoology, Prof. Ramkrishna More Arts, Commerce & Science College, Akurdi, Pune - 411044, Maharashtra, India</mods:affiliation>
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<mods:date id="764B8EEB11E5822121620428996132F5">2024</mods:date>
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<taxonomicName id="4C784D28062A4917FF272223AB34DF76" ID-CoL="Y3B3" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[151,515,798,825]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="latipes">
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<emphasis id="B90CEAB9062A4917FF272223A852DF77" bold="true" box="[151,357,798,825]" italics="true" pageId="9" pageNumber="128">Copelatus latipes</emphasis>
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<bibRefCitation id="EFE94B5A062A4917FEDD2223AB34DF76" author="Sharp, D." box="[365,515,798,824]" pageId="9" pageNumber="128" pagination="179 - 1003" refId="ref7398" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18. https: // doi. org / 10.5962 / bhl. title. 9530" type="journal article" year="1882">Sharp, 1882</bibRefCitation>
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(
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<figureCitation id="13432A2E062A4917FF2F227EA9DFDF13" box="[159,232,835,861]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="9" pageNumber="128">Figs 6</figureCitation>
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,
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<figureCitation id="13432A2E062A4917FF44227EA80DDF13" box="[244,314,835,861]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="9" pageNumber="128">17–18</figureCitation>
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)
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<emphasis id="B90CEAB9062A4917FF2722BAA87FDFD1" box="[151,328,903,927]" italics="true" pageId="9" pageNumber="128">Copelatus latipes</emphasis>
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<bibRefCitation id="EFE94B5A062A4917FEFE22BAA8C8DFD1" author="Sharp, D." box="[334,511,903,927]" pageId="9" pageNumber="128" pagination="179 - 1003" refId="ref7398" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18. https: // doi. org / 10.5962 / bhl. title. 9530" type="journal article" year="1882">Sharp, 1882: 580</bibRefCitation>
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;
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<treatmentCitation id="0AD910BA062A4917FDB922BAABD3DFD1" author="Regimbart, M." box="[521,740,903,927]" page="296" pageId="9" pageNumber="128" year="1899">
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<bibRefCitation id="EFE94B5A062A4917FDB922BAABD3DFD1" author="Regimbart, M." box="[521,740,903,927]" pageId="9" pageNumber="128" pagination="186 - 367" refId="ref7333" refString="Regimbart, M. (1899) Revision des Dytiscidae de la region Indo-Sino-Malaise. Annales de la Societe Entomologique de France, 68, 186 - 367." type="journal article" year="1899">Régimbart 1899: 296</bibRefCitation>
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;
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<bibRefCitation id="EFE94B5A062A4917FD5E22BAAA86DFD1" author="Vazirani, T. G." box="[750,945,903,927]" pageId="9" pageNumber="128" pagination="303 - 362" refId="ref7524" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4 (3), 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article" year="1970">Vazirani 1970: 307</bibRefCitation>
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.
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<typeStatus id="54C38809062A4917FF2722F0A9E5DFA9" box="[151,210,973,999]" pageId="9" pageNumber="128">Type</typeStatus>
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locality.
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“Malacca” [
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State]
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.
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<emphasis id="B90CEAB9062A4917FF7722CCA842D845" bold="true" box="[199,373,1009,1035]" pageId="9" pageNumber="128">Type material.</emphasis>
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<materialsCitation id="3B103CF6062A4917FECC22CFA9D7D81D" collectionCode="BMNH" country="Malaysia" location="Type" municipality="Type" pageId="9" pageNumber="128" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
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<typeStatus id="54C38809062A4917FECC22CFA8DBD842" box="[380,492,1010,1036]" pageId="9" pageNumber="128" type="lectotype">Lectotype</typeStatus>
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(designated by
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<bibRefCitation id="EFE94B5A062A4917FD0B22CCAAA3D845" author="Vazirani, T. G." box="[699,916,1009,1035]" pageId="9" pageNumber="128" pagination="303 - 362" refId="ref7524" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4 (3), 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article" year="1970">Vazirani 1970: 307</bibRefCitation>
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;
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<collectionCode id="ED69AE6E062A4917FC1022CCAACFD845" box="[928,1016,1009,1035]" pageId="9" pageNumber="128">BMNH</collectionCode>
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), labelled: “
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[p, round label with red frame] //
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/ 691 [hw] // Sharp Coll. / 1905-313. [p] //
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691 /
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<taxonomicName id="4C784D28062A4917FBA22528ADB5D861" authorityName="Erichson" authorityYear="1832" box="[1042,1154,1045,1071]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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/ latipes
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/
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<collectingCountry id="F36F763B062A4917FA8D2528ACABD861" box="[1341,1436,1045,1071]" name="Malaysia" pageId="9" pageNumber="128">Malacca</collectingCountry>
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[hw]”.
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<paragraph id="8BC736AB062A4917FF772560ABA3D965" blockId="9.[151,1437,973,1971]" pageId="9" pageNumber="128">
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<emphasis id="B90CEAB9062A4917FF772560AB0CD839" bold="true" box="[199,571,1117,1143]" pageId="9" pageNumber="128">Additional material examined.</emphasis>
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<materialsCitation id="3B103CF6062A4917FDF52560A9CAD8D5" collectingDate="2002-11-01" collectingDateMax="2002-11-11" collectingDateMin="2002-11-01" collectionCode="NMPC" collectorName="P. Kocarek" country="Cambodia" location="Siem Reap" pageId="9" pageNumber="128" specimenCount="1" specimenCount-female="1" stateProvince="Siem Reap">
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<emphasis id="B90CEAB9062A4917FDF52560ABDAD839" bold="true" box="[581,749,1117,1143]" pageId="9" pageNumber="128">
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<collectingCountry id="F36F763B062A4917FDF52560ABDFD839" box="[581,744,1117,1143]" name="Cambodia" pageId="9" pageNumber="128">CAMBODIA</collectingCountry>
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:
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</emphasis>
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,
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,
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<collectingMethod id="52394EBC062A4917FC762560AD05D836" box="[966,1074,1117,1144]" pageId="9" pageNumber="128">light trap</collectingMethod>
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,
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<date id="FFC6106B062A4917FBF12560ADEBD839" box="[1089,1244,1117,1143]" pageId="9" pageNumber="128" value="2002-11-01" valueMax="2002-11-11" valueMin="2002-11-01">
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<collectingDate id="EF82E983062A4917FBF12560ADEBD839" box="[1089,1244,1117,1143]" pageId="9" pageNumber="128" value="2002-11-01" valueMax="2002-11-11" valueMin="2002-11-01">1.–11.xi.2002</collectingDate>
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</date>
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,
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<collectorName id="268D537D062A4917FB5C2563AC5ED839" box="[1260,1385,1117,1144]" pageId="9" pageNumber="128">P. Kočárek</collectorName>
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leg. (
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<collectionCode id="ED69AE6E062A4917FF2F25BCA9C2D8D5" box="[159,245,1153,1179]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="128">NMPC</collectionCode>
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)
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</materialsCitation>
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.
|
||
<materialsCitation id="3B103CF6062A4917FEB825BCAB08D8F1" collectingDate="2001-05-22" collectingDateMax="2001-06-08" collectingDateMin="2001-05-22" collectionCode="NMPC" collectorName="E. Jendek & O. Sausa" country="Laos" elevation="550" latitude="17.716667" location="Nakai" longLatPrecision="1277" longitude="105.15" pageId="9" pageNumber="128" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Khammouan">
|
||
<emphasis id="B90CEAB9062A4917FEB825BCA86AD8D5" bold="true" box="[264,349,1153,1179]" pageId="9" pageNumber="128">
|
||
<collectingCountry id="F36F763B062A4917FEB825BCA86FD8D5" box="[264,344,1153,1179]" name="Laos" pageId="9" pageNumber="128">LAOS</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D7EFD22062A4917FED525BCA8A5D8D2" box="[357,402,1153,1180]" count="1" pageId="9" pageNumber="128" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D7EFD22062A4917FE2F25BCA8FED8D2" box="[415,457,1153,1180]" count="2" pageId="9" pageNumber="128" type="female">2 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="49BCF849062A4917FE6825BCABD4D8D5" box="[472,739,1153,1179]" country="Laos" name="Khammouan" pageId="9" pageNumber="128">Khammouane Province</collectingRegion>
|
||
,
|
||
<location id="8EA76070062A4917FD5F25BCAA05D8D5" LSID="urn:lsid:plazi:treatment:03D187BD062A4917FF272223A8E6DBFD:8EA76070062A4917FD5F25BCAA05D8D5" box="[751,818,1153,1179]" country="Laos" latitude="17.716667" longLatPrecision="1277" longitude="105.15" name="Nakai" pageId="9" pageNumber="128" stateProvince="Khammouan">Nakai</location>
|
||
env.,
|
||
<geoCoordinate id="EE4C506C062A4917FCC625BCAAE0D8D5" box="[886,983,1153,1179]" degrees="17" direction="north" minutes="43" orientation="latitude" pageId="9" pageNumber="128" precision="925" value="17.716667">17°43’N</geoCoordinate>
|
||
<geoCoordinate id="EE4C506C062A4917FC6F25BCAD7AD8D2" box="[991,1101,1153,1180]" degrees="105" direction="east" minutes="09" orientation="longitude" pageId="9" pageNumber="128" precision="925" value="105.15">105°09’E</geoCoordinate>
|
||
,
|
||
<quantity id="4C809B4E062A4917FBE925BCADECD8D2" box="[1113,1243,1153,1180]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="9" pageNumber="128" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">
|
||
<elevation id="0055D198062A4917FBE925BCADECD8D2" box="[1113,1243,1153,1180]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="9" pageNumber="128" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500–600 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFC6106B062A4917FB5725BCACAFD8D5" box="[1255,1432,1153,1179]" pageId="9" pageNumber="128" value="2001-05-22" valueMax="2001-06-08" valueMin="2001-05-22">
|
||
<collectingDate id="EF82E983062A4917FB5725BCACAFD8D5" box="[1255,1432,1153,1179]" pageId="9" pageNumber="128" value="2001-05-22" valueMax="2001-06-08" valueMin="2001-05-22">22.v.–8.vi.2001</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="268D537D062A4917FF27259BA83FD8F1" box="[151,264,1189,1216]" pageId="9" pageNumber="128">E. Jendek</collectorName>
|
||
&
|
||
<collectorName id="268D537D062A4917FE832598A8ADD88C" box="[307,410,1189,1218]" pageId="9" pageNumber="128">O. Šauša</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED69AE6E062A4917FE512598AB00D8F1" box="[481,567,1189,1215]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="128">NMPC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B103CF6062A4917FDFD2598ACAED8AD" collectingDate="2011-12-06" collectionCode="NMPC" collectorName="PT Silva Rimba Lestari & J. Hajek & J. Schneider & P. Votruba" country="Indonesia" elevation="60" latitude="0.123333335" location="Limbang" longLatPrecision="130" longitude="116.30167" pageId="9" pageNumber="128" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Utara">
|
||
<emphasis id="B90CEAB9062A4917FDFD2598ABC2D8F1" bold="true" box="[589,757,1189,1215]" pageId="9" pageNumber="128">
|
||
<collectingCountry id="F36F763B062A4917FDFD2598ABC7D8F1" box="[589,752,1189,1215]" name="Indonesia" pageId="9" pageNumber="128">INDONESIA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D7EFD22062A4917FCB02598AA18D88E" box="[768,815,1189,1216]" count="1" pageId="9" pageNumber="128" type="male">1 ♂</specimenCount>
|
||
, E
|
||
<collectingRegion id="49BCF849062A4917FCEB2598AAD6D8F1" box="[859,993,1189,1215]" country="Indonesia" name="Kalimantan Utara" pageId="9" pageNumber="128">Kalimantan</collectingRegion>
|
||
,
|
||
<collectorName id="268D537D062A4917FC5F259BAC35D8F1" box="[1007,1282,1189,1216]" pageId="9" pageNumber="128">PT Silva Rimba Lestari</collectorName>
|
||
[area], camp “
|
||
<location id="8EA76070062A4917FF1125F4A83AD8AD" LSID="urn:lsid:plazi:treatment:03D187BD062A4917FF272223A8E6DBFD:8EA76070062A4917FF1125F4A83AD8AD" box="[161,269,1225,1251]" country="Indonesia" latitude="0.123333335" longLatPrecision="130" longitude="116.30167" name="Limbang" pageId="9" pageNumber="128" stateProvince="Kalimantan Utara">Limbang</location>
|
||
”,
|
||
<geoCoordinate id="EE4C506C062A4917FE9725F4A8AAD8AD" box="[295,413,1225,1251]" degrees="00" direction="north" minutes="07.4" orientation="latitude" pageId="9" pageNumber="128" precision="92" value="0.123333335">00°07.4’N</geoCoordinate>
|
||
<geoCoordinate id="EE4C506C062A4917FE1825F4AB1ED8AD" box="[424,553,1225,1251]" degrees="116" direction="east" minutes="18.1" orientation="longitude" pageId="9" pageNumber="128" precision="92" value="116.30167">116°18.1’E</geoCoordinate>
|
||
,
|
||
<quantity id="4C809B4E062A4917FD8A25F4AB4ED8AA" box="[570,633,1225,1252]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="128" unit="m" value="60.0">
|
||
<elevation id="0055D198062A4917FD8A25F4AB4ED8AA" box="[570,633,1225,1252]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="128" unit="m" value="60.0">60 m</elevation>
|
||
</quantity>
|
||
, 3.–4.+
|
||
<date id="FFC6106B062A4917FD7E25F4AA74D8AD" box="[718,835,1225,1251]" pageId="9" pageNumber="128" value="2011-12-06">
|
||
<collectingDate id="EF82E983062A4917FD7E25F4AA74D8AD" box="[718,835,1225,1251]" pageId="9" pageNumber="128" value="2011-12-06">6.xii.2011</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="268D537D062A4917FCE325F7AA81D8AD" box="[851,950,1225,1252]" pageId="9" pageNumber="128">J. Hájek</collectorName>
|
||
,
|
||
<collectorName id="268D537D062A4917FC7525F7AD65D8AD" box="[965,1106,1225,1252]" pageId="9" pageNumber="128">J. Schneider</collectorName>
|
||
&
|
||
<collectorName id="268D537D062A4917FBCE25F7ADC3D8AD" box="[1150,1268,1225,1252]" pageId="9" pageNumber="128">P. Votruba</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED69AE6E062A4917FA8B25F4ACA6D8AD" box="[1339,1425,1225,1251]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="128">NMPC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B103CF6062A4917FF2725D0ABA7D965" collectingDate="2008-04-09" collectingDateMax="2008-04-30" collectingDateMin="2008-04-09" collectionCode="NMPC" collectorName="G. Keriung & Z. Linek" country="Malaysia" elevation="400" location="Kg. Tebu Hitnam" municipality="Endau Rompin" pageId="9" pageNumber="128" specimenCount="24" specimenCount-female="12" specimenCount-male="12" stateProvince="Pahang">
|
||
<emphasis id="B90CEAB9062A4917FF2725D0A800D949" bold="true" box="[151,311,1261,1287]" pageId="9" pageNumber="128">
|
||
<collectingCountry id="F36F763B062A4917FF2725D0A806D949" box="[151,305,1261,1287]" name="Malaysia" pageId="9" pageNumber="128">MALAYSIA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D7EFD22062A4917FE8E25D0A84FD946" box="[318,376,1261,1288]" count="12" pageId="9" pageNumber="128" type="male">12 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D7EFD22062A4917FE3525D0A88AD946" box="[389,445,1261,1288]" count="12" pageId="9" pageNumber="128" type="female">12 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="49BCF849062A4917FE7B25D0AB17D949" box="[459,544,1261,1287]" country="Malaysia" name="Pahang" pageId="9" pageNumber="128">Pahang</collectingRegion>
|
||
,
|
||
<locationDeviation id="9A752F89062A4917FD9C25D0AA78D949" box="[556,847,1261,1288]" location="Kuala Rompin" pageId="9" pageNumber="128">
|
||
<quantity id="4C809B4E062A4917FD9C25D0AB43D949" box="[556,628,1261,1288]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="128" unit="km" value="50.0">50 km</quantity>
|
||
NE Kuala Rompin
|
||
</locationDeviation>
|
||
,
|
||
<collectingMunicipality id="6BA3ACD1062A4917FCEA25D0AD34D949" box="[858,1027,1261,1287]" pageId="9" pageNumber="128">Endau Rompin</collectingMunicipality>
|
||
NP,
|
||
<collectorName id="268D537D062A4917FB8A25D0AD8FD949" box="[1082,1208,1261,1287]" pageId="9" pageNumber="128">G. Keriung</collectorName>
|
||
(
|
||
<location id="8EA76070062A4917FB7825D0ACA7D949" LSID="urn:lsid:plazi:treatment:03D187BD062A4917FF272223A8E6DBFD:8EA76070062A4917FB7825D0ACA7D949" box="[1224,1424,1261,1287]" country="Malaysia" municipality="Endau Rompin" name="Kg. Tebu Hitnam" pageId="9" pageNumber="128" stateProvince="Pahang">Kg. Tebu Hitnam</location>
|
||
),
|
||
<quantity id="4C809B4E062A4917FF27242CA9D7D962" box="[151,224,1297,1324]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="128" unit="m" value="400.0">
|
||
<elevation id="0055D198062A4917FF27242CA9D7D962" box="[151,224,1297,1324]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="128" unit="m" value="400.0">400 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFC6106B062A4917FF5C242CA8BFD965" box="[236,392,1297,1323]" pageId="9" pageNumber="128" value="2008-04-09" valueMax="2008-04-30" valueMin="2008-04-09">
|
||
<collectingDate id="EF82E983062A4917FF5C242CA8BFD965" box="[236,392,1297,1323]" pageId="9" pageNumber="128" value="2008-04-09" valueMax="2008-04-30" valueMin="2008-04-09">9.–30.iv.2008</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="268D537D062A4917FE23242FA8C4D965" box="[403,499,1297,1324]" pageId="9" pageNumber="128">Z. Linek</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED69AE6E062A4917FD82242CABBFD965" box="[562,648,1297,1323]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="9" pageNumber="128">NMPC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3626520062A4917FF772408AA69DB6D" pageId="9" pageNumber="128" type="diagnosis">
|
||
<paragraph id="8BC736AB062A4917FF772408ABD2DA95" blockId="9.[151,1437,973,1971]" pageId="9" pageNumber="128">
|
||
<emphasis id="B90CEAB9062A4917FF772408A875D901" bold="true" box="[199,322,1333,1359]" pageId="9" pageNumber="128">Diagnosis.</emphasis>
|
||
TL: 5.1–6.0 mm. Head black, clypeus paler, reddish; pronotum black, laterally brownish, pale colouration extending anteriorly; elytra black with reddish apex and sides laterally from elytral striae, and indistinct reddish pattern consisting of: two basal spots between striae 2–3 and 4–6, and longitudinal spot in middle third of elytra between striae 4–5. Pronotum with coarse punctures laterally. Each elytron with six discal striae and one submarginal stria: stria 1 largely absent, present only in apical fifth of elytral length; striae 2, 3 and 5 ending subapically; stria 6 shortest, ending at apical fifth; submarginal stria long, beginning at elytral mid-length and ending subapically at level of striae 2, 3 and 5 (
|
||
<figureCitation id="13432A2E062A4917FD122730ABD2DA69" box="[674,741,1549,1575]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="9" pageNumber="128">Fig. 6</figureCitation>
|
||
). Male longer spur on metatibia weakly sinuate in apical third. Median lobe of aedeagus in lateral view (
|
||
<figureCitation id="13432A2E062A4917FDDB270CAB8ADA05" box="[619,701,1585,1611]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="9" pageNumber="128">Fig. 17</figureCitation>
|
||
) broad and parallel-sided in basal 2/3; apical third sickle-shaped, almost regularly curved in middle, apex bifid; in 2/3 of its length with two distinct ‘teeth’ on ventral side—broad acute proximal one, and narrower, incised, apically obtuse distal one. Parameres with distinct lobe on inner margin (
|
||
<figureCitation id="13432A2E062A4917FF2F27A0A9C5DAF9" box="[159,242,1693,1719]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="9" pageNumber="128">Fig. 18</figureCitation>
|
||
). Females identical to males in habitus, punctation of elytra slightly coarser, weakly impressed short striolae present in middle part of elytra laterally from stria 3.
|
||
</paragraph>
|
||
<paragraph id="8BC736AB062A4917FF7727D8AA69DB6D" blockId="9.[151,1437,973,1971]" pageId="9" pageNumber="128">
|
||
<emphasis id="B90CEAB9062A4917FF7727D8A8CCDAB1" bold="true" box="[199,507,1765,1791]" pageId="9" pageNumber="128">Collecting circumstances.</emphasis>
|
||
The single specimen from
|
||
<collectingRegion id="49BCF849062A4917FC8E27D8AAF6DAB1" box="[830,961,1765,1791]" country="Indonesia" name="Kalimantan Utara" pageId="9" pageNumber="128">Kalimantan</collectingRegion>
|
||
was collected in a shallow pool in open oligotrophic wetland on sandy ground, surrounded with shrubs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3626520062A4917FF772610A8E6DBFD" pageId="9" pageNumber="128" type="distribution">
|
||
<paragraph id="8BC736AB062A4917FF772610A8E6DBFD" blockId="9.[151,1437,973,1971]" pageId="9" pageNumber="128">
|
||
<emphasis id="B90CEAB9062A4917FF772610A856DB09" bold="true" box="[199,353,1837,1863]" pageId="9" pageNumber="128">Distribution.</emphasis>
|
||
The species was so far known only from Peninsular
|
||
<collectingCountry id="F36F763B062A4917FC192610AD24DB09" box="[937,1043,1837,1863]" name="Malaysia" pageId="9" pageNumber="128">Malaysia</collectingCountry>
|
||
. Here we present new records from central
|
||
<collectingCountry id="F36F763B062A4917FF5A266FA814DB22" box="[234,291,1874,1900]" name="Laos" pageId="9" pageNumber="128">Laos</collectingCountry>
|
||
,
|
||
<collectingCountry id="F36F763B062A4917FE9F266CA895DB25" box="[303,418,1873,1899]" name="Cambodia" pageId="9" pageNumber="128">Cambodia</collectingCountry>
|
||
and Indonesian part of Borneo Island, documenting that
|
||
<taxonomicName id="4C784D28062A4917FB90266FADBBDB25" authorityName="Sharp" authorityYear="1882" box="[1056,1164,1873,1900]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="128" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B90CEAB9062A4917FB90266FADBBDB25" box="[1056,1164,1873,1900]" italics="true" pageId="9" pageNumber="128">C. latipes</emphasis>
|
||
</taxonomicName>
|
||
is relatively widespread in Southeast Asia (
|
||
<figureCitation id="13432A2E062A4917FEC32648A8FDDBC1" box="[371,458,1909,1935]" captionStart="FIGURE 19" captionStartId="6.[152,255,1288,1312]" captionTargetBox="[156,1437,273,1261]" captionTargetId="figure-66@6.[151,1437,273,1265]" captionTargetPageId="6" captionText="FIGURE 19. Map of distribution of Copelatus latipes species complex." figureDoi="http://doi.org/10.5281/zenodo.12742768" httpUri="https://zenodo.org/record/12742768/files/figure.png" pageId="9" pageNumber="128">Fig. 19</figureCitation>
|
||
).
|
||
<bibRefCitation id="EFE94B5A062A4917FE522648AB9ADBC1" author="Regimbart, M." box="[482,685,1909,1935]" pageId="9" pageNumber="128" pagination="186 - 367" refId="ref7333" refString="Regimbart, M. (1899) Revision des Dytiscidae de la region Indo-Sino-Malaise. Annales de la Societe Entomologique de France, 68, 186 - 367." type="journal article" year="1899">Régimbart (1899)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFE94B5A062A4917FD5A2648AAEFDBC1" author="Zimmermann, A." box="[746,984,1909,1935]" pageId="9" pageNumber="128" pagination="68 - 249" refId="ref7608" refString="Zimmermann, A. (1919) Die Schwimmkafer des Deutschen Entomologischen Museums in Berlin-Dahlem. Archiv fur Naturgeschichte, Abteilung A, 83 (12), 68 - 249. [1917]" type="journal article" year="1919">Zimmermann (1919)</bibRefCitation>
|
||
mentioned also
|
||
<collectingCountry id="F36F763B062A4917FB2B2648ADE0DBC1" box="[1179,1239,1909,1935]" name="India" pageId="9" pageNumber="128">India</collectingCountry>
|
||
, which was later doubted by
|
||
<bibRefCitation id="EFE94B5A062A4917FEAB26A4A8FADBFD" author="Vazirani, T. G." box="[283,461,1945,1971]" pageId="9" pageNumber="128" pagination="303 - 362" refId="ref7524" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4 (3), 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article" year="1970">Vazirani (1970)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |