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<mods:title id="D53529AAB97E8D343DD4F4D2E296FA0A">Distribution of intertidal rock oysters in the Pilbara, Western Australia</mods:title>
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<mods:namePart id="39974A45D37312604348BBBA20BF9D3B">Wells, Fred E.</mods:namePart>
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<mods:namePart id="5DCF55B9BFBC2AFCD7BA1A404E9C4A54">Lukehurst, Sherralee S.</mods:namePart>
<mods:affiliation id="418B90DEB3D2FDA149813428C73D6239">School of Molecular and Life Sciences, Curtin University, Kent Street, Bentley, WA 6102 Australia</mods:affiliation>
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<mods:namePart id="EA0DE7259D0C4247029EE8A4B849D157">Fullwood, Laura A. F.</mods:namePart>
<mods:affiliation id="5FEA3DEBF8E159BCC9BAA89AC1A1F948">School of Molecular and Life Sciences, Curtin University, Kent Street, Bentley, WA 6102 Australia</mods:affiliation>
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<mods:namePart id="FC2B75E35EBA2CED9C734A1C663BD9E7">Harvey, Euan S.</mods:namePart>
<mods:affiliation id="D8C95715BD2B3C38AF2856C0DC90389B">School of Molecular and Life Sciences, Curtin University, Kent Street, Bentley, WA 6102 Australia</mods:affiliation>
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<mods:date id="69D356CC591767755186E1FF67E6ED85">2024</mods:date>
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<treatment id="039A87ADD30EFFEEFD9CF88CFDDCF4EF" LSID="urn:lsid:plazi:treatment:039A87ADD30EFFEEFD9CF88CFDDCF4EF" httpUri="http://treatment.plazi.org/id/039A87ADD30EFFEEFD9CF88CFDDCF4EF" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137">
<subSubSection id="C3296530D30EFFEFFD9CF88CFAD0F7FE" pageId="6" pageNumber="137" type="nomenclature">
<paragraph id="8B8C36BBD30EFFEFFD9CF88CFA09F73B" blockId="6.[528,1432,1827,1856]" box="[528,1432,1827,1856]" pageId="6" pageNumber="137">
<heading id="D0C481D7D30EFFEFFD9CF88CFA09F73B" box="[528,1432,1827,1856]" fontSize="12" level="2" pageId="6" pageNumber="137" reason="4">
<taxonomicName id="4C334D38D30EFFEFFD9CF88CFBEAF73B" ID-CoL="6WYJ9" authority="(Peron and Lesueur, 1807)" baseAuthorityName="Peron and Lesueur" baseAuthorityYear="1807" box="[528,1147,1827,1856]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="6" pageNumber="137" phylum="Mollusca" rank="species" species="scyphophilla">
Saccostrea scyphophilla
<emphasis id="B947EAA9D30EFFEFFCB6F88BFBEAF73B" box="[826,1147,1827,1856]" italics="true" pageId="6" pageNumber="137">(Peron and Lesueur, 1807)</emphasis>
</taxonomicName>
<emphasis id="B947EAA9D30EFFEFFB0EF88BFA09F73B" box="[1154,1432,1827,1856]" italics="true" pageId="6" pageNumber="137">
(
<figureCitation id="13082A3ED30EFFEFFB06F88BFA9AF744" box="[1162,1291,1827,1856]" captionStart="Figure 2" captionStartId="2.[528,599,1212,1234]" captionTargetBox="[188,1453,43,1190]" captionTargetId="figure-287@2.[532,1454,188,1190]" captionTargetPageId="2" captionText="Figure 2. Intertidal rock oysters from the Pilbara, Western Australia. A: Saccostrea lineage A from a trial aquaculture farm (WAMS_117501; GenBank OR466892). B. Saccostrea scyphophilla (WAMS_117500; GenBank OR466930) C. Natural Saccostrea lineage A from a rocky shore (WAMS_117506; GenBank OR466894).D, E. Talonostrea sp. nov. (WAMS_117503; WAMS_117504; GenBank OR466939). Scale bars are 1 cm. Photographs by Fred Wells." figureDoi="http://doi.org/10.5281/zenodo.12628091" httpUri="https://zenodo.org/record/12628091/files/figure.png" pageId="6" pageNumber="137">Figures 2b</figureCitation>
,
<figureCitation id="13082A3ED30EFFEFFA9AF88CFAB4F73B" box="[1302,1317,1827,1855]" captionStart="Figure 3" captionStartId="3.[188,259,542,564]" captionTargetBox="[188,1454,43,504]" captionTargetId="figure-414@3.[192,1454,203,504]" captionTargetPageId="3" captionText="Figure 3. Saccostrea lineage A (left) and Saccostrea scyphophilla (center) and Talonostrea sp. nov. in their natural habitat in the Pilbara, Western Australia. Note that Talonostrea sp. nov. actually lived underwater attached to the bottom of the Avicennia marina branch but have been moved so they are visible. Photographs by Fred Wells." figureDoi="http://doi.org/10.5281/zenodo.12628093" httpUri="https://zenodo.org/record/12628093/files/figure.png" pageId="6" pageNumber="137">3</figureCitation>
;
<tableCitation id="C6B10300D30EFFEFFABFF88CFA1CF73B" box="[1331,1421,1827,1856]" captionStart="Table 1" captionStartId="5.[188,249,198,219]" captionTargetPageId="5" captionText="Table 1. Sites searched for intertidal oysters in the Pilbara, Western Australia. SA, Saccostrea lineage A; Ss, Saccostrea scyphophilla; Ta, Talonostrea sp. nov." pageId="6" pageNumber="137">Table 1</tableCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8C36BBD30EFFEFFD9CF8F2FAD0F7FE" blockId="6.[528,1454,1885,2042]" pageId="6" pageNumber="137">
<emphasis id="B947EAA9D30EFFEFFD9CF8F2FCDBF77E" box="[528,842,1885,1914]" italics="true" pageId="6" pageNumber="137">Brief description of shell:</emphasis>
Shell large, up to
<quantity id="4CCB9B5ED30EFFEFFBC8F8F2FB16F77E" box="[1092,1159,1885,1914]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="137" unit="cm" value="10.0">10cm</quantity>
in length, very heavy. Usually occurs as individuals that in dense assemblages do not extend over each other. Upper surface rounded. Shell off white, margins purplish. Interior whitish, adductor muscle scar on right valve may be a deep purple.
</paragraph>
</subSubSection>
<subSubSection id="C3296530D30FFFEEFDA2FF6AFDDCF4EF" pageId="7" pageNumber="138" type="description">
<paragraph id="8B8C36BBD30FFFEEFDA2FF6AFD4DF1E5" blockId="7.[528,1454,197,1259]" pageId="7" pageNumber="138">
<emphasis id="B947EAA9D30FFFEEFDA2FF6AFC3AF0E6" box="[558,939,197,226]" italics="true" pageId="7" pageNumber="138">Records in the present survey:</emphasis>
Ashburton Port area: Direction Island; East end Thevenard Island; Ashburton Island; Salt Creek; Old Onslow jetty, Beadon Point. Dampier Port area: Withnell Bay; Southern entrance to King Bay; Norbill Bay, Rosemary Island; Whalers Bay, Malus Island; Marney Bay, Malus Island; High Point, Whittaker Island; North side of East Intercourse Island. Port Hedland Port area: Public boat ramp; Finucane Island.
</paragraph>
<paragraph id="8B8C36BBD30FFFEEFDA2FE5FFBFAF289" blockId="7.[528,1454,197,1259]" pageId="7" pageNumber="138">
<emphasis id="B947EAA9D30FFFEEFDA2FE5FFABBF208" box="[558,1322,495,524]" italics="true" pageId="7" pageNumber="138">Previous records in the Pilbara verified by DNA sequences:</emphasis>
<bibRefCitation id="EFA24B4AD30FFFEEFAB5FE5FFD54F233" author="Lam K &amp; Morton B" pageId="7" pageNumber="138" pagination="235 - 245" refId="ref6310" refString="Lam K, Morton B (2006) Morphological and mitochondrial-DNA analysis of the Indo-West Pacific rock oysters (Ostreidae: Saccostrea species). Journal of Molluscan Studies 72: 235 - 245, https: // doi. org / 10.1016 / S 0044 - 8486 (03) 00215 - 1" type="journal article" year="2006">Lam and Morton (2006)</bibRefCitation>
: Vlamingh Head, north of Exmouth; Barrow Island; Withnell Bay, Burrup Peninsula (all as
<taxonomicName id="4C334D38D30FFFEEFC0DFDE9FB20F266" authority="(Gould, 1850)" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[897,1201,581,610]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="7" pageNumber="138" phylum="Mollusca" rank="species" species="mordax">
<emphasis id="B947EAA9D30FFFEEFC0DFDE9FC6CF266" box="[897,1021,581,610]" italics="true" pageId="7" pageNumber="138">S. mordax</emphasis>
(Gould, 1850)
</taxonomicName>
.
<bibRefCitation id="EFA24B4AD30FFFEEFB4FFDEAFA38F266" author="Snow M &amp; Fotedar S &amp; Wilson NG &amp; Kirkendale LA" box="[1219,1449,581,610]" pageId="7" pageNumber="138" refId="ref7132" refString="Snow M, Fotedar S, Wilson NG, Kirkendale LA (2023) Clarifying the natural distribution of Saccostrea Dollfus and Dautzenberg, 1920 (edible rock oyster) species in Western Australia to guide development of a fledgling aquaculture industry. Aquaculture 566: 739202, https: // doi. org / 10.1016 / j. aquaculture. 2022.739202" type="journal volume" year="2023">Snow et al. (2023)</bibRefCitation>
: Flying Foam Passage, north of Burrup Peninsula.
</paragraph>
<paragraph id="8B8C36BBD30FFFEEFDA2FD34FAC5F33F" blockId="7.[528,1454,197,1259]" pageId="7" pageNumber="138">
<emphasis id="B947EAA9D30FFFEEFDA2FD34FBE8F2BC" box="[558,1145,667,696]" italics="true" pageId="7" pageNumber="138">Published distribution (based on DNA analyses):</emphasis>
Okinawa,
<collectingCountry id="F324762BD30FFFEEFB71FD34FACAF2BC" box="[1277,1371,667,696]" name="Taiwan" pageId="7" pageNumber="138">Taiwan</collectingCountry>
,
<collectingCountry id="F324762BD30FFFEEFAEBFD34FDC7F2E7" name="Hong Kong" pageId="7" pageNumber="138">Hong Kong</collectingCountry>
,
<collectingCountry id="F324762BD30FFFEEFDE8FD68FD23F2E0" box="[612,690,711,740]" name="China" pageId="7" pageNumber="138">China</collectingCountry>
,
<collectingCountry id="F324762BD30FFFEEFD33FD68FCACF2E0" box="[703,829,711,740]" name="Singapore" pageId="7" pageNumber="138">Singapore</collectingCountry>
, northern
<collectingCountry id="F324762BD30FFFEEFC4EFD68FBA3F2E0" box="[962,1074,711,740]" name="Australia" pageId="7" pageNumber="138">Australia</collectingCountry>
(
<collectingRegion id="49F7F859D30FFFEEFBC8FD68FABAF2E0" box="[1092,1323,711,740]" country="Australia" name="Western Australia" pageId="7" pageNumber="138">Western Australia</collectingRegion>
,
<collectingRegion id="49F7F859D30FFFEEFAB4FD68FD15F30B" country="Australia" name="Northern Territory" pageId="7" pageNumber="138">Northern Territory</collectingRegion>
,
<collectingRegion id="49F7F859D30FFFEEFD18FD5DFCBBF30B" box="[660,810,754,783]" country="Australia" name="Queensland" pageId="7" pageNumber="138">Queensland</collectingRegion>
,
<collectingRegion id="49F7F859D30FFFEEFCB5FD5CFB8EF30B" box="[825,1055,754,783]" country="Australia" name="New South Wales" pageId="7" pageNumber="138">New South Wales</collectingRegion>
) (
<bibRefCitation id="EFA24B4AD30FFFEEFBB6FD5DFACEF30B" author="Lam K &amp; Morton B" box="[1082,1375,754,783]" pageId="7" pageNumber="138" pagination="235 - 245" refId="ref6310" refString="Lam K, Morton B (2006) Morphological and mitochondrial-DNA analysis of the Indo-West Pacific rock oysters (Ostreidae: Saccostrea species). Journal of Molluscan Studies 72: 235 - 245, https: // doi. org / 10.1016 / S 0044 - 8486 (03) 00215 - 1" type="journal article" year="2006">Lam and Morton 2006</bibRefCitation>
(as
<taxonomicName id="4C334D38D30FFFEEFA14FD5CFDFEF33F" baseAuthorityName="Gould" baseAuthorityYear="1850" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="7" pageNumber="138" phylum="Mollusca" rank="species" species="mordax">
<emphasis id="B947EAA9D30FFFEEFA14FD5CFDFEF33F" italics="true" pageId="7" pageNumber="138">S. mordax</emphasis>
</taxonomicName>
); McDougall 2020;
<bibRefCitation id="EFA24B4AD30FFFEEFCEEFCB1FBE0F33F" author="McDougall C &amp; Nenadic N &amp; Healy J" box="[866,1137,798,827]" pageId="7" pageNumber="138" refId="ref6497" refString="McDougall C, Nenadic N, Healy J (2020) Guide to Queensland's intertidal oysters. Griffith University, Brisbane, Queensland, Australia, 9 pp" type="book" year="2020">McDougall et al. 2020</bibRefCitation>
;
<bibRefCitation id="EFA24B4AD30FFFEEFBF0FCB1FAD9F33F" author="Snow M &amp; Fotedar S &amp; Wilson NG &amp; Kirkendale LA" box="[1148,1352,798,827]" pageId="7" pageNumber="138" refId="ref7132" refString="Snow M, Fotedar S, Wilson NG, Kirkendale LA (2023) Clarifying the natural distribution of Saccostrea Dollfus and Dautzenberg, 1920 (edible rock oyster) species in Western Australia to guide development of a fledgling aquaculture industry. Aquaculture 566: 739202, https: // doi. org / 10.1016 / j. aquaculture. 2022.739202" type="journal volume" year="2023">Snow et al. 2023</bibRefCitation>
).
</paragraph>
<paragraph id="8B8C36BBD30FFFEEFDA2FCE4FB10F412" blockId="7.[528,1454,197,1259]" pageId="7" pageNumber="138">
<emphasis id="B947EAA9D30FFFEEFDA2FCE4FC0AF363" box="[558,923,842,871]" italics="true" pageId="7" pageNumber="138">
Notes:
<taxonomicName id="4C334D38D30FFFEEFD0CFCE5FC0AF363" baseAuthorityName="Peron and Lesueur" baseAuthorityYear="1807" box="[640,923,842,871]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="7" pageNumber="138" phylum="Mollusca" rank="species" species="scyphophilla">Saccostrea scyphophilla</taxonomicName>
</emphasis>
also occurred on a wide variety of habitats, from isolated rocks, intertidal rock platforms, mangroves and artificial structures on both open and protected shores, sometimes at high densities. It frequently co-occurred with
<taxonomicName id="4C334D38D30FFFEEFC07FC62FB98F3ED" authorityName="Dollfus &amp; Dautzenberg" authorityYear="1920" box="[907,1033,973,1001]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="7" pageNumber="138" phylum="Mollusca" rank="genus">
<emphasis id="B947EAA9D30FFFEEFC07FC62FB98F3ED" box="[907,1033,973,1001]" italics="true" pageId="7" pageNumber="138">Saccostrea</emphasis>
</taxonomicName>
lineage A, but was more common on open, exposed shores than
<taxonomicName id="4C334D38D30FFFEEFC0FFC56FB91F411" authorityName="Dollfus &amp; Dautzenberg" authorityYear="1920" box="[899,1024,1017,1045]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="7" pageNumber="138" phylum="Mollusca" rank="genus">
<emphasis id="B947EAA9D30FFFEEFC0FFC56FB91F411" box="[899,1024,1017,1045]" italics="true" pageId="7" pageNumber="138">Saccostrea</emphasis>
</taxonomicName>
lineage A.
</paragraph>
<paragraph id="8B8C36BBD30FFFEEFDA2FB8AFDDCF4EF" blockId="7.[528,1454,197,1259]" pageId="7" pageNumber="138">
<bibRefCitation id="EFA24B4AD30FFFEEFDA2FB8AFC9BF445" author="Snow M &amp; Fotedar S &amp; Wilson NG &amp; Kirkendale LA" box="[558,778,1060,1090]" pageId="7" pageNumber="138" refId="ref7132" refString="Snow M, Fotedar S, Wilson NG, Kirkendale LA (2023) Clarifying the natural distribution of Saccostrea Dollfus and Dautzenberg, 1920 (edible rock oyster) species in Western Australia to guide development of a fledgling aquaculture industry. Aquaculture 566: 739202, https: // doi. org / 10.1016 / j. aquaculture. 2022.739202" type="journal volume" year="2023">Snow et al. (2023)</bibRefCitation>
commented that
<taxonomicName id="4C334D38D30FFFEEFC69FB8AFB06F445" baseAuthorityName="Peron and Lesueur" baseAuthorityYear="1807" box="[997,1175,1060,1089]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="7" pageNumber="138" phylum="Mollusca" rank="species" species="scyphophilla">
<emphasis id="B947EAA9D30FFFEEFC69FB8AFB06F445" box="[997,1175,1060,1089]" italics="true" pageId="7" pageNumber="138">S. scyphophilla</emphasis>
</taxonomicName>
in WA was previously reported as two lineages, A and B by
<bibRefCitation id="EFA24B4AD30FFFEEFC54FBFFFA97F468" author="Lam K &amp; Morton B" box="[984,1286,1103,1132]" pageId="7" pageNumber="138" pagination="235 - 245" refId="ref6310" refString="Lam K, Morton B (2006) Morphological and mitochondrial-DNA analysis of the Indo-West Pacific rock oysters (Ostreidae: Saccostrea species). Journal of Molluscan Studies 72: 235 - 245, https: // doi. org / 10.1016 / S 0044 - 8486 (03) 00215 - 1" type="journal article" year="2006">Lam and Morton (2006)</bibRefCitation>
, with B being very rare.
<bibRefCitation id="EFA24B4AD30FFFEEFD11FBD5FC02F493" author="Snow M &amp; Fotedar S &amp; Wilson NG &amp; Kirkendale LA" box="[669,915,1145,1175]" pageId="7" pageNumber="138" refId="ref7132" refString="Snow M, Fotedar S, Wilson NG, Kirkendale LA (2023) Clarifying the natural distribution of Saccostrea Dollfus and Dautzenberg, 1920 (edible rock oyster) species in Western Australia to guide development of a fledgling aquaculture industry. Aquaculture 566: 739202, https: // doi. org / 10.1016 / j. aquaculture. 2022.739202" type="journal volume" year="2023">Snow et al. (2023)</bibRefCitation>
considered that the differences between lineages were relatively small and combined the two, a practice followed here.
</paragraph>
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</treatment>
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<mods:title id="23C6FA8EB3AB0702D6848ED029B536F9">Distribution of intertidal rock oysters in the Pilbara, Western Australia</mods:title>
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<mods:namePart id="BA09FAD737943E136C7C2B985634B495">Wells, Fred E.</mods:namePart>
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<mods:affiliation id="61ED70AA5B644030772303C33172E8E0">School of Molecular and Life Sciences, Curtin University, Kent Street, Bentley, WA 6102 Australia</mods:affiliation>
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<mods:affiliation id="74000673AA0DB52C1475DDB2ECD622AC">School of Molecular and Life Sciences, Curtin University, Kent Street, Bentley, WA 6102 Australia</mods:affiliation>
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<paragraph id="8B8C36BBD30EFFEFFD9CFF68FBC6F0E7" blockId="6.[528,1111,198,228]" box="[528,1111,198,228]" pageId="6" pageNumber="137">
<heading id="D0C481D7D30EFFEFFD9CFF68FBC6F0E7" box="[528,1111,198,228]" centered="true" fontSize="12" level="2" pageId="6" pageNumber="137" reason="4">
<taxonomicName id="4C334D38D30EFFEFFD9CFF68FD07F0E0" ID-CoL="7PG3Q" authority="Dollfus &amp; Dautzenberg, 1920" authorityName="Dollfus &amp; Dautzenberg" authorityYear="1920" box="[528,662,199,228]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="6" pageNumber="137" phylum="Mollusca" rank="genus">Saccostrea</taxonomicName>
<emphasis id="B947EAA9D30EFFEFFD11FF69FBC6F0E7" box="[669,1111,198,227]" italics="true" pageId="6" pageNumber="137">
lineage A (
<figureCitation id="13082A3ED30EFFEFFCAEFF68FC52F0E7" box="[802,963,199,227]" captionStart="Figure 2" captionStartId="2.[528,599,1212,1234]" captionTargetBox="[188,1453,43,1190]" captionTargetId="figure-287@2.[532,1454,188,1190]" captionTargetPageId="2" captionText="Figure 2. Intertidal rock oysters from the Pilbara, Western Australia. A: Saccostrea lineage A from a trial aquaculture farm (WAMS_117501; GenBank OR466892). B. Saccostrea scyphophilla (WAMS_117500; GenBank OR466930) C. Natural Saccostrea lineage A from a rocky shore (WAMS_117506; GenBank OR466894).D, E. Talonostrea sp. nov. (WAMS_117503; WAMS_117504; GenBank OR466939). Scale bars are 1 cm. Photographs by Fred Wells." figureDoi="http://doi.org/10.5281/zenodo.12628091" httpUri="https://zenodo.org/record/12628091/files/figure.png" pageId="6" pageNumber="137">Figures 2a, c</figureCitation>
,
<figureCitation id="13082A3ED30EFFEFFC5CFF68FC4EF0E7" box="[976,991,199,227]" captionStart="Figure 3" captionStartId="3.[188,259,542,564]" captionTargetBox="[188,1454,43,504]" captionTargetId="figure-414@3.[192,1454,203,504]" captionTargetPageId="3" captionText="Figure 3. Saccostrea lineage A (left) and Saccostrea scyphophilla (center) and Talonostrea sp. nov. in their natural habitat in the Pilbara, Western Australia. Note that Talonostrea sp. nov. actually lived underwater attached to the bottom of the Avicennia marina branch but have been moved so they are visible. Photographs by Fred Wells." figureDoi="http://doi.org/10.5281/zenodo.12628093" httpUri="https://zenodo.org/record/12628093/files/figure.png" pageId="6" pageNumber="137">3</figureCitation>
;
<tableCitation id="C6B10300D30EFFEFFC62FF69FBDDF0E7" box="[1006,1100,198,227]" captionStart="Table 1" captionStartId="5.[188,249,198,219]" captionTargetPageId="5" captionText="Table 1. Sites searched for intertidal oysters in the Pilbara, Western Australia. SA, Saccostrea lineage A; Ss, Saccostrea scyphophilla; Ta, Talonostrea sp. nov." pageId="6" pageNumber="137">Table 1</tableCitation>
)
</emphasis>
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</paragraph>
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<paragraph id="8B8C36BBD30EFFEFFD9CFEAFFA08F1CC" blockId="6.[528,1454,256,1779]" pageId="6" pageNumber="137">
<emphasis id="B947EAA9D30EFFEFFD9CFEAFFCA6F119" box="[528,823,256,285]" italics="true" pageId="6" pageNumber="137">Brief description of shell:</emphasis>
Often densely packed so shells grow to fit available space. Medium size, oval, length
<quantity id="4CCB9B5ED30EFFEFFC2BFE84FC72F143" box="[935,995,299,328]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="137" unit="cm" value="5.0">5 cm</quantity>
or less, hinge to tip of shell. Margins crenulated, margins may have short lobes that can be worn off. Outer surface off white. Inner white, adductor scar on left valve purple. Oysters grown individually for aquaculture up to
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, elongate, left valve deeper.
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<subSubSection id="C3296530D30EFFEFFDA2FE79FAC5F376" pageId="6" pageNumber="137" type="distribution">
<paragraph id="8B8C36BBD30EFFEFFDA2FE79FB7DF2F6" blockId="6.[528,1454,256,1779]" pageId="6" pageNumber="137">
<emphasis id="B947EAA9D30EFFEFFDA2FE79FC0BF1F7" box="[558,922,470,499]" italics="true" pageId="6" pageNumber="137">Records in the present survey:</emphasis>
Ashburton Port area: East end Thevenard Island; Hooley Creek; Salt Creek; Beadon Creek jetty; Four Mile Creek. Dampier Port area: Withnell Bay, Burrup Peninsula; Southern entrance to King Bay; Eastern King Bay; Dampier boat landing; Watering Cove, Burrup Peninsula; Karratha Bay, West Lewis Island; North side of East Intercourse Island. Port Samson port area: Point Samson. Port Hedland Port area: Public boat ramp, Port Hedland reef, Pretty Pool.
</paragraph>
<paragraph id="8B8C36BBD30EFFEFFDA2FCAEFAC5F376" blockId="6.[528,1454,256,1779]" pageId="6" pageNumber="137">
<emphasis id="B947EAA9D30EFFEFFDA2FCAEFABBF319" box="[558,1322,768,797]" italics="true" pageId="6" pageNumber="137">Previous records in the Pilbara verified by DNA sequences:</emphasis>
<bibRefCitation id="EFA24B4AD30EFFEFFAB5FCAFFD41F34C" author="Lam K &amp; Morton B" pageId="6" pageNumber="137" pagination="235 - 245" refId="ref6310" refString="Lam K, Morton B (2006) Morphological and mitochondrial-DNA analysis of the Indo-West Pacific rock oysters (Ostreidae: Saccostrea species). Journal of Molluscan Studies 72: 235 - 245, https: // doi. org / 10.1016 / S 0044 - 8486 (03) 00215 - 1" type="journal article" year="2006">Lam and Morton (2006)</bibRefCitation>
: Exmouth; Barrow Island; Watering Cove; Withnell Bay.
<bibRefCitation id="EFA24B4AD30EFFEFFD9CFCFAFD7EF376" author="Snow M &amp; Fotedar S &amp; Wilson NG &amp; Kirkendale LA" box="[528,751,853,882]" pageId="6" pageNumber="137" refId="ref7132" refString="Snow M, Fotedar S, Wilson NG, Kirkendale LA (2023) Clarifying the natural distribution of Saccostrea Dollfus and Dautzenberg, 1920 (edible rock oyster) species in Western Australia to guide development of a fledgling aquaculture industry. Aquaculture 566: 739202, https: // doi. org / 10.1016 / j. aquaculture. 2022.739202" type="journal volume" year="2023">Snow et al. (2023)</bibRefCitation>
: Flying Foam Passage, north of Burrup Peninsula.
</paragraph>
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<paragraph id="8B8C36BBD30EFFEFFDA2FC2EFC98F476" blockId="6.[528,1454,256,1779]" pageId="6" pageNumber="137">
<emphasis id="B947EAA9D30EFFEFFDA2FC2EFC99F399" box="[558,776,897,925]" italics="true" pageId="6" pageNumber="137">
Notes:
<taxonomicName id="4C334D38D30EFFEFFD06FC2EFC99F399" authorityName="Dollfus &amp; Dautzenberg" authorityYear="1920" box="[650,776,897,925]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="6" pageNumber="137" phylum="Mollusca" rank="genus">Saccostrea</taxonomicName>
</emphasis>
lineage A occurred on a wide variety of habitats including lying loose on intertidal sandflats, and attached to isolated rocks, intertidal rock platforms, mangroves and artificial structures on both protected and open shores, sometimes at high densities. It frequently co-occurred with
<emphasis id="B947EAA9D30EFFEFFD4BFB84FC13F44C" box="[711,898,1067,1096]" italics="true" pageId="6" pageNumber="137">
<taxonomicName id="4C334D38D30EFFEFFD4BFB84FCEFF44C" baseAuthorityName="Peron and Lesueur" baseAuthorityYear="1807" box="[711,894,1067,1096]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="6" pageNumber="137" phylum="Mollusca" rank="species" species="scyphophilla">S. scyphophilla</taxonomicName>
,
</emphasis>
but was more common on protected shores than
<taxonomicName id="4C334D38D30EFFEFFDC3FBF9FC93F477" baseAuthorityName="Peron and Lesueur" baseAuthorityYear="1807" box="[591,770,1110,1139]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="6" pageNumber="137" phylum="Mollusca" rank="species" species="scyphophilla">
<emphasis id="B947EAA9D30EFFEFFDC3FBF9FC93F477" box="[591,770,1110,1139]" italics="true" pageId="6" pageNumber="137">S. scyphophilla</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8C36BBD30EFFEFFDA2FB2EFA9AF6F7" blockId="6.[528,1454,256,1779]" pageId="6" pageNumber="137">
<taxonomicName id="4C334D38D30EFFEFFDA2FB2EFC4AF49A" ID-CoL="6WYJ2" authority="(Born, 1778)" baseAuthorityName="Born" baseAuthorityYear="1778" box="[558,987,1153,1182]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="6" pageNumber="137" phylum="Mollusca" rank="species" species="cuccullata">
<emphasis id="B947EAA9D30EFFEFFDA2FB2EFCA0F49A" box="[558,817,1153,1182]" italics="true" pageId="6" pageNumber="137">Saccostrea cuccullata</emphasis>
(Born, 1778)
</taxonomicName>
was regarded as a widespread Indo- Pacific species with a variable shell morphology (
<bibRefCitation id="EFA24B4AD30EFFEFFB0CFB03FA84F4CC" author="Harry HW" box="[1152,1301,1195,1225]" pageId="6" pageNumber="137" pagination="121 - 158" refId="ref6059" refString="Harry HW (1985) Synopsis of the supraspecific classification of living oysters (Bivalvia, Gryphaeidae and Ostreidae). The Veliger 28: 121 - 158." type="journal article" year="1985">Harry 1985</bibRefCitation>
), but recent DNA sequencing has divided the “species” into a number of separate genetic lineages (e.g.
<bibRefCitation id="EFA24B4AD30EFFEFFD5CFAAEFB81F519" author="Lam K &amp; Morton B" box="[720,1040,1280,1309]" pageId="6" pageNumber="137" pagination="235 - 245" refId="ref6310" refString="Lam K, Morton B (2006) Morphological and mitochondrial-DNA analysis of the Indo-West Pacific rock oysters (Ostreidae: Saccostrea species). Journal of Molluscan Studies 72: 235 - 245, https: // doi. org / 10.1016 / S 0044 - 8486 (03) 00215 - 1" type="journal article" year="2006">Lam and Morton 2006</bibRefCitation>
;
<bibRefCitation id="EFA24B4AD30EFFEFFBABFAAFFA37F519" author="Sekino M &amp; Yamashita H" box="[1063,1446,1280,1309]" pageId="6" pageNumber="137" pagination="492 - 506" refId="ref6913" refString="Sekino M, Yamashita H (2016) Mitochondrial and nuclear DNA analyses of Saccostrea oysters in Japan highlight the confused taxonomy of the genus. Journal of Molluscan Studies 82: 492 - 506, https: // doi. org / 10.1093 / mollus / eyw 022" type="journal article" year="2016">Sekino and Yamashita 2016</bibRefCitation>
; McDougall 2020;
<bibRefCitation id="EFA24B4AD30EFFEFFD7EFA84FC58F54C" author="Snow M &amp; Fotedar S &amp; Wilson NG &amp; Kirkendale LA" box="[754,969,1323,1352]" pageId="6" pageNumber="137" refId="ref7132" refString="Snow M, Fotedar S, Wilson NG, Kirkendale LA (2023) Clarifying the natural distribution of Saccostrea Dollfus and Dautzenberg, 1920 (edible rock oyster) species in Western Australia to guide development of a fledgling aquaculture industry. Aquaculture 566: 739202, https: // doi. org / 10.1016 / j. aquaculture. 2022.739202" type="journal volume" year="2023">Snow et al. 2023</bibRefCitation>
). The taxonomic status of the various lineages is uncertain. Oysters previously identified as
<taxonomicName id="4C334D38D30EFFEFFB3EFAF9FADCF577" ID-CoL="6WYJ2" baseAuthorityName="Born" baseAuthorityYear="1778" box="[1202,1357,1366,1395]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="6" pageNumber="137" phylum="Mollusca" rank="species" species="cuccullata">
<emphasis id="B947EAA9D30EFFEFFB3EFAF9FADCF577" box="[1202,1357,1366,1395]" italics="true" pageId="6" pageNumber="137">S. cuccullata</emphasis>
</taxonomicName>
may in fact be a species complex, with each lineage representing a separate species (McDougall 2020).
<bibRefCitation id="EFA24B4AD30EFFEFFC94FA03FBF6F5CC" author="Lam K &amp; Morton B" box="[792,1127,1451,1480]" pageId="6" pageNumber="137" pagination="235 - 245" refId="ref6310" refString="Lam K, Morton B (2006) Morphological and mitochondrial-DNA analysis of the Indo-West Pacific rock oysters (Ostreidae: Saccostrea species). Journal of Molluscan Studies 72: 235 - 245, https: // doi. org / 10.1016 / S 0044 - 8486 (03) 00215 - 1" type="journal article" year="2006">Lam and Morton (2006)</bibRefCitation>
referred to the lineage occurring in the Pilbara as
<taxonomicName id="4C334D38D30EFFEFFC01FA78FBBEF5F7" ID-CoL="6WYJ2" baseAuthorityName="Born" baseAuthorityYear="1778" box="[909,1071,1494,1523]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="6" pageNumber="137" phylum="Mollusca" rank="species" species="cuccullata">
<emphasis id="B947EAA9D30EFFEFFC01FA78FBBEF5F7" box="[909,1071,1494,1523]" italics="true" pageId="6" pageNumber="137">S. cuccullata</emphasis>
</taxonomicName>
A, but more recent studies (McDougall 2020;
<bibRefCitation id="EFA24B4AD30EFFEFFD76F9AEFB80F61A" author="McDougall C &amp; Nenadic N &amp; Healy J" box="[762,1041,1537,1566]" pageId="6" pageNumber="137" refId="ref6497" refString="McDougall C, Nenadic N, Healy J (2020) Guide to Queensland's intertidal oysters. Griffith University, Brisbane, Queensland, Australia, 9 pp" type="book" year="2020">McDougall et al. 2020</bibRefCitation>
;
<bibRefCitation id="EFA24B4AD30EFFEFFB92F9AEFB7FF61A" author="Snow M &amp; Fotedar S &amp; Wilson NG &amp; Kirkendale LA" box="[1054,1262,1537,1566]" pageId="6" pageNumber="137" refId="ref7132" refString="Snow M, Fotedar S, Wilson NG, Kirkendale LA (2023) Clarifying the natural distribution of Saccostrea Dollfus and Dautzenberg, 1920 (edible rock oyster) species in Western Australia to guide development of a fledgling aquaculture industry. Aquaculture 566: 739202, https: // doi. org / 10.1016 / j. aquaculture. 2022.739202" type="journal volume" year="2023">Snow et al. 2023</bibRefCitation>
) have reported the lineages without a species name, a process followed here.
<bibRefCitation id="EFA24B4AD30EFFEFFAAEF984FDF1F677" author="Snow M &amp; Fotedar S &amp; Wilson NG &amp; Kirkendale LA" pageId="6" pageNumber="137" refId="ref7132" refString="Snow M, Fotedar S, Wilson NG, Kirkendale LA (2023) Clarifying the natural distribution of Saccostrea Dollfus and Dautzenberg, 1920 (edible rock oyster) species in Western Australia to guide development of a fledgling aquaculture industry. Aquaculture 566: 739202, https: // doi. org / 10.1016 / j. aquaculture. 2022.739202" type="journal volume" year="2023">Snow et al. (2023)</bibRefCitation>
reported that
<taxonomicName id="4C334D38D30EFFEFFC9DF9F8FC1FF677" ID-CoL="7PG3Q" authorityName="Dollfus &amp; Dautzenberg" authorityYear="1920" box="[785,910,1623,1651]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="6" pageNumber="137" phylum="Mollusca" rank="genus">
<emphasis id="B947EAA9D30EFFEFFC9DF9F8FC1FF677" box="[785,910,1623,1651]" italics="true" pageId="6" pageNumber="137">Saccostrea</emphasis>
</taxonomicName>
lineage A occurs only in
<collectingRegion id="49F7F859D30EFFEFFB4AF9F8FA38F677" box="[1222,1449,1622,1651]" country="Australia" name="Western Australia" pageId="6" pageNumber="137">Western Australia</collectingRegion>
, from Shark Bay to Broome. It is thus probably an undescribed species. Until this is done we have listed the WA specimens as
<taxonomicName id="4C334D38D30EFFEFFB38F903FAA3F6CC" ID-CoL="7PG3Q" authorityName="Dollfus &amp; Dautzenberg" authorityYear="1920" box="[1204,1330,1708,1736]" class="Bivalvia" family="Ostreidae" genus="Saccostrea" kingdom="Animalia" order="Ostreida" pageId="6" pageNumber="137" phylum="Mollusca" rank="genus">
<emphasis id="B947EAA9D30EFFEFFB38F903FAA3F6CC" box="[1204,1330,1708,1736]" italics="true" pageId="6" pageNumber="137">Saccostrea</emphasis>
</taxonomicName>
lineage A and deposited specimens in the Western Australian Museum..
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,127 @@
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<mods:title id="4828A2F1AC8A50ADCFE1A475D632F964">Distribution of intertidal rock oysters in the Pilbara, Western Australia</mods:title>
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<mods:namePart id="8E535C43F84D168F67B7FA3E01CF49AA">Lukehurst, Sherralee S.</mods:namePart>
<mods:affiliation id="700EFB54C014E22226071DEDA1F5D571">School of Molecular and Life Sciences, Curtin University, Kent Street, Bentley, WA 6102 Australia</mods:affiliation>
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<mods:namePart id="990C69B641B457B8B7D7E757F4C74D91">Fullwood, Laura A. F.</mods:namePart>
<mods:affiliation id="CB251EE0C97A6AE55288613A4AA37C75">School of Molecular and Life Sciences, Curtin University, Kent Street, Bentley, WA 6102 Australia</mods:affiliation>
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<mods:namePart id="20B044F0F2DBE2EB63A4BC396B036B3D">Harvey, Euan S.</mods:namePart>
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<treatment id="039A87ADD30FFFE1FD9CFAB4FAC1F1B3" LSID="urn:lsid:plazi:treatment:039A87ADD30FFFE1FD9CFAB4FAC1F1B3" httpUri="http://treatment.plazi.org/id/039A87ADD30FFFE1FD9CFAB4FAC1F1B3" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138">
<subSubSection id="C3296530D30FFFEEFD9CFAB4FC51F53C" box="[528,960,1307,1336]" pageId="7" pageNumber="138" type="nomenclature">
<paragraph id="8B8C36BBD30FFFEEFD9CFAB4FC51F53C" blockId="7.[528,960,1307,1336]" box="[528,960,1307,1336]" pageId="7" pageNumber="138">
<heading id="D0C481D7D30FFFEEFD9CFAB4FC51F53C" box="[528,960,1307,1336]" fontSize="12" level="2" pageId="7" pageNumber="138" reason="4">
<taxonomicName id="4C334D38D30FFFEEFD9CFAB4FD38F53C" ID-CoL="7PHZ3" authorityName="Wells &amp; Lukehurst &amp; Fullwood &amp; Harvey" authorityYear="2024" box="[528,681,1307,1336]" class="Bivalvia" family="Ostreidae" genus="Talonostrea" kingdom="Animalia" order="Ostreida" pageId="7" pageNumber="140" phylum="Mollusca" rank="genus" status="sp. nov.">Talonostrea</taxonomicName>
<emphasis id="B947EAA9D30FFFEEFD3DFAB3FC51F53C" box="[689,960,1307,1336]" italics="true" pageId="7" pageNumber="138">
<taxonomicNameLabel id="A27457D2D30FFFEEFD3DFAB3FC81F53C" box="[689,784,1308,1336]" pageId="7" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13082A3ED30FFFEEFCACFAB3FC27F53C" box="[800,950,1307,1336]" captionStart="Figure 2" captionStartId="2.[528,599,1212,1234]" captionTargetBox="[188,1453,43,1190]" captionTargetId="figure-287@2.[532,1454,188,1190]" captionTargetPageId="2" captionText="Figure 2. Intertidal rock oysters from the Pilbara, Western Australia. A: Saccostrea lineage A from a trial aquaculture farm (WAMS_117501; GenBank OR466892). B. Saccostrea scyphophilla (WAMS_117500; GenBank OR466930) C. Natural Saccostrea lineage A from a rocky shore (WAMS_117506; GenBank OR466894).D, E. Talonostrea sp. nov. (WAMS_117503; WAMS_117504; GenBank OR466939). Scale bars are 1 cm. Photographs by Fred Wells." figureDoi="http://doi.org/10.5281/zenodo.12628091" httpUri="https://zenodo.org/record/12628091/files/figure.png" pageId="7" pageNumber="138">Figure 2d, e</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3296530D30FFFEEFD9CFAFAFB7FF619" pageId="7" pageNumber="138" type="description">
<paragraph id="8B8C36BBD30FFFEEFD9CFAFAFB7FF619" blockId="7.[528,1454,1365,2034]" pageId="7" pageNumber="138">
<emphasis id="B947EAA9D30FFFEEFD9CFAFAFCA9F576" box="[528,824,1365,1394]" italics="true" pageId="7" pageNumber="138">Brief description of shell:</emphasis>
Shell small,
<quantity id="4CCB9B5ED30FFFEEFC5EFAF9FB97F577" box="[978,1030,1366,1395]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="138" unit="cm" value="4.0">4cm</quantity>
or less in length, flattened, fragile, lack chomata, outer margin of left valve may be extended upwards. Occur as single individuals, shape variable to elongate depending on substrate. Outer shell colour variable, white to deep purple. Interior of shell chalky white. Adductor scar usually white, but may be light purple.
</paragraph>
</subSubSection>
<subSubSection id="C3296530D30FFFEEFDA2F985FC42F7F6" pageId="7" pageNumber="138" type="distribution">
<paragraph id="8B8C36BBD30FFFEEFDA2F985FD45F699" blockId="7.[528,1454,1365,2034]" pageId="7" pageNumber="138">
<emphasis id="B947EAA9D30FFFEEFDA2F985FC28F643" box="[558,953,1578,1607]" italics="true" pageId="7" pageNumber="138">Records in the present survey:</emphasis>
Ashburton Port area: Hooley Creek. Dampier Port Area: Karratha Bay, West Lewis Island. Balla Balla port area: Intertidal creek.
</paragraph>
<paragraph id="8B8C36BBD30FFFEEFDA2F903FDC4F798" blockId="7.[528,1454,1365,2034]" pageId="7" pageNumber="138">
<emphasis id="B947EAA9D30FFFEEFDA2F903FAC4F6CC" box="[558,1365,1707,1736]" italics="true" pageId="7" pageNumber="138">Previous records in the Pilbara verified by DNA sequences:</emphasis>
None published. The WA Department of Primary Industry and Regional Development (
<collectionCode id="ED22AE7ED30FFFEEFD4BF8AFFCB9F719" box="[711,808,1792,1821]" pageId="7" pageNumber="138">DPIRD</collectionCode>
) has sequences from Flying Foam Passage, Dampier and Cone Bay, Kimberley collected between 2018-2020. The specimens were small and were obtained from spat collectors used during aquaculture trials.
</paragraph>
<paragraph id="8B8C36BBD30FFFEEFDA2F805FC42F7F6" blockId="7.[528,1454,1365,2034]" pageId="7" pageNumber="138">
<emphasis id="B947EAA9D30FFFEEFDA2F805FBEEF7C3" box="[558,1151,1962,1991]" italics="true" pageId="7" pageNumber="138">Published distribution (based on DNA analyses):</emphasis>
<location id="8EEC6060D30FFFEEFB04F804FABAF7C3" LSID="urn:lsid:plazi:treatment:039A87ADD30FFFE1FD9CFAB4FAC1F1B3:8EEC6060D30FFFEEFB04F804FABAF7C3" box="[1160,1323,1963,1991]" name="Moreton Bay" pageId="7" pageNumber="138" stateProvince="Queensland">Moreton Bay</location>
to Cairns,
<collectingRegion id="49F7F859D30FFFEEFD9CF87AFD35F7F6" box="[528,676,2005,2034]" country="Australia" name="Queensland" pageId="7" pageNumber="138">Queensland</collectingRegion>
(
<bibRefCitation id="EFA24B4AD30FFFEEFD3FF87AFC56F7F6" author="McDougall C &amp; Nenadic N &amp; Healy J" box="[691,967,2005,2034]" pageId="7" pageNumber="138" refId="ref6497" refString="McDougall C, Nenadic N, Healy J (2020) Guide to Queensland's intertidal oysters. Griffith University, Brisbane, Queensland, Australia, 9 pp" type="book" year="2020">McDougall et al. 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3296530D300FFE1FDA2FF69FAC1F1B3" pageId="8" pageNumber="139" type="discussion">
<paragraph id="8B8C36BBD300FFE1FDA2FF69FC3CF166" blockId="8.[528,1454,197,439]" pageId="8" pageNumber="139">
<emphasis id="B947EAA9D300FFE1FDA2FF69FDE7F0E6" box="[558,630,198,226]" italics="true" pageId="8" pageNumber="139">Notes:</emphasis>
<location id="8EEC6060D300FFE1FDF1FF6AFCC7F0E6" LSID="urn:lsid:plazi:treatment:039A87ADD30FFFE1FD9CFAB4FAC1F1B3:8EEC6060D300FFE1FDF1FF6AFCC7F0E6" box="[637,854,197,226]" municipality="The Karratha Bay" name="The Hooley Creek" pageId="8" pageNumber="139">The Hooley Creek</location>
specimen was collected on an intertidal sandflat in the creek.
<collectingMunicipality id="6BE8ACC1D300FFE1FD03FF40FCF9F108" box="[655,872,239,268]" pageId="8" pageNumber="139">The Karratha Bay</collectingMunicipality>
specimen was found on a steel support for the aquaculture facility.
<specimenCount id="9D35FD32D300FFE1FC80FEB5FC28F133" box="[780,953,282,311]" count="6" pageId="8" pageNumber="139" type="generic">Six specimens</specimenCount>
were collected at the base of mangroves (
<taxonomicName id="4C334D38D300FFE1FD97FEEAFD67F166" box="[539,758,325,354]" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="139" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="B947EAA9D300FFE1FD97FEEAFD67F166" box="[539,758,325,354]" italics="true" pageId="8" pageNumber="139">Avicennia marina</emphasis>
</taxonomicName>
) at Balla Balla.
</paragraph>
<paragraph id="8B8C36BBD300FFE1FDA2FEDFFAC1F1B3" blockId="8.[528,1454,197,439]" pageId="8" pageNumber="139">
<bibRefCitation id="EFA24B4AD300FFE1FDA2FEDFFCFBF189" author="McDougall C &amp; Nenadic N &amp; Healy J" box="[558,874,368,397]" pageId="8" pageNumber="139" refId="ref6497" refString="McDougall C, Nenadic N, Healy J (2020) Guide to Queensland's intertidal oysters. Griffith University, Brisbane, Queensland, Australia, 9 pp" type="book" year="2020">McDougall et al. (2020)</bibRefCitation>
believe this is an undescribed species. WA specimens have been deposited in the Western Australian Museum.
</paragraph>
</subSubSection>
</treatment>
</document>