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<document id= "8E01CF62069A122C0F82523645AEB019" ID-DOI= "10.25674/415" ID-ISSN= "2509-9523" ID-Zenodo-Dep= "13194032" IM.bibliography_approvedBy= "felipe" IM.illustrations_approvedBy= "felipe" IM.materialsCitations_approvedBy= "felipe" IM.metadata_approvedBy= "felipe" IM.tables_requiresApprovalFor= "GgImagineBatch" IM.taxonomicNames_approvedBy= "felipe" IM.treatments_approvedBy= "felipe" checkinTime= "1722631177589" checkinUser= "felipe" docAuthor= "Seifert, Bernhard" docDate= "2024" docId= "577887E6FF84FFDC68CF5D64FE2B358E" docLanguage= "en" docName= "SoilOrg.96.2.113-144.pdf" docOrigin= "Soil Organisms (London, England) 96 (2)" docSource= "http://dx.doi.org/10.25674/415" docStyle= "DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId= "EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName= "SoilOrg.2018-.journal_article" docStyleVersion= "2" docTitle= "Cardiocondyla heinzei Seifert, 2024, n.sp." docType= "treatment" docVersion= "2" lastPageNumber= "135" masterDocId= "AB41FF9EFF91FFCA681B5A5FFFA8347E" masterDocTitle= "A taxonomic revision of the ants of the Cardiocondyla wroughtonii group (Hymenoptera: Formicidae) with a checklist of the Cardiocondyla species of the world" masterLastPageNumber= "144" masterPageNumber= "113" pageNumber= "134" updateTime= "1722679537643" updateUser= "ExternalLinkService" zenodo-license-document= "CC-BY-4.0" >
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<mods:title id= "EC847B77A5E3BE69C809D498F1C1D826" > A taxonomic revision of the ants of the Cardiocondyla wroughtonii group (Hymenoptera: Formicidae) with a checklist of the Cardiocondyla species of the world</mods:title>
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<mods:namePart id= "F6428A4CCBF554A9142A9B40F7331B51" > Seifert, Bernhard</mods:namePart>
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<mods:date id= "E8AE146800D6973A75AC9F4E2D1B5D36" > 2024</mods:date>
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<treatment id= "577887E6FF84FFDC68CF5D64FE2B358E" LSID= "urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E" httpUri= "http://treatment.plazi.org/id/577887E6FF84FFDC68CF5D64FE2B358E" lastPageId= "22" lastPageNumber= "135" pageId= "21" pageNumber= "134" >
<subSubSection id= "97CB657BFF84FFDF68CF5D64FDE53329" box= "[212,589,1851,1879]" pageId= "21" pageNumber= "134" type= "nomenclature" >
<paragraph id= "DF6E36F0FF84FFDF68CF5D64FDE53329" blockId= "21.[212,589,1851,1879]" box= "[212,589,1851,1879]" pageId= "21" pageNumber= "134" >
<heading id= "8426819CFF84FFDF68CF5D64FDE53329" box= "[212,589,1851,1879]" fontSize= "11" level= "2" pageId= "21" pageNumber= "134" reason= "5" >
<emphasis id= "EDA5EAE2FF84FFDF68CF5D64FDE53329" bold= "true" box= "[212,589,1851,1879]" pageId= "21" pageNumber= "134" >
<taxonomicName id= "18D14D73FF84FFDF68CF5D64FDA83329" authorityName= "Seifert" authorityYear= "2024" box= "[212,512,1851,1879]" class= "Insecta" family= "Formicidae" genus= "Cardiocondyla" kingdom= "Animalia" order= "Hymenoptera" pageId= "21" pageNumber= "134" phylum= "Arthropoda" rank= "species" species= "heinzei" status= "sp. nov." >
<emphasis id= "EDA5EAE2FF84FFDF68CF5D64FDA83329" bold= "true" box= "[212,512,1851,1879]" italics= "true" pageId= "21" pageNumber= "134" > Cardiocondyla heinzei</emphasis>
</taxonomicName>
<taxonomicNameLabel id= "F6965799FF84FFDF6A115D64FDE53329" box= "[522,589,1851,1879]" pageId= "21" pageNumber= "134" rank= "species" > n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id= "97CB657BFF84FFDF68CF5D21FC783554" pageId= "21" pageNumber= "134" type= "etymology" >
<paragraph id= "DF6E36F0FF84FFDF68CF5D21FC8A33E9" blockId= "21.[189,802,1918,1975]" box= "[212,802,1918,1943]" pageId= "21" pageNumber= "134" > Etymology: the name is given in honor of Jürgen</paragraph>
<paragraph id= "DF6E36F0FF84FFDF68A65DC0FC783554" blockId= "21.[189,802,1918,1975]" lastBlockId= "21.[824,1437,241,1976]" pageId= "21" pageNumber= "134" >
Heinze wo has done and supervised outstanding research on the fascinating biology of
<taxonomicName id= "18D14D73FF84FFDF6C905AAEFA823577" authorityName= "Emery" authorityYear= "1869" box= "[1163,1322,241,265]" class= "Insecta" family= "Formicidae" genus= "Cardiocondyla" kingdom= "Animalia" order= "Hymenoptera" pageId= "21" pageNumber= "134" phylum= "Arthropoda" rank= "genus" >
<emphasis id= "EDA5EAE2FF84FFDF6C905AAEFA823577" box= "[1163,1322,241,265]" italics= "true" pageId= "21" pageNumber= "134" > Cardiocondyla</emphasis>
</taxonomicName>
ants over three decades.
</paragraph>
</subSubSection>
<subSubSection id= "97CB657BFF84FFDF6B545B6CFBE93747" pageId= "21" pageNumber= "134" type= "materials_examined" >
<paragraph id= "DF6E36F0FF84FFDF6B545B6CFC503535" blockId= "21.[824,1437,241,1976]" box= "[847,1016,307,331]" pageId= "21" pageNumber= "134" >
<emphasis id= "EDA5EAE2FF84FFDF6B545B6CFC503535" bold= "true" box= "[847,1016,307,331]" pageId= "21" pageNumber= "134" > Type material:</emphasis>
</paragraph>
<paragraph id= "DF6E36F0FF84FFDF6B545B0BFAB636CB" blockId= "21.[824,1437,241,1976]" pageId= "21" pageNumber= "134" >
<materialsCitation id= "6FB93CADFF84FFDF6B545B0BFAB636CB" collectingDate= "2019-04-04" collectingDateMax= "2019-04-07" collectingDateMin= "2019-04-04" collectionCode= "SMNG" county= "Res. Station" elevation= "191" latitude= "8.7703" location= "Iringo" longLatPrecision= "7" longitude= "-3.7897" municipality= "Park" pageId= "21" pageNumber= "134" specimenCode= "SMNG20240630" specimenCount= "9" specimenCount-worker= "8" stateProvince= "Comoe" typeStatus= "holotype" >
<typeStatus id= "006A8852FF84FFDF6B545B0BFC1B3512" box= "[847,947,340,364]" pageId= "21" pageNumber= "134" type= "holotype" > Holotype</typeStatus>
worker labelled ‘ IVO:
<geoCoordinate id= "BAE55037FF84FFDF6CDF5B0BFA803512" box= "[1220,1320,340,364]" degrees= "8.7703" direction= "north" orientation= "latitude" pageId= "21" pageNumber= "134" precision= "5" value= "8.7703" > 8.7703°N</geoCoordinate>
,
<geoCoordinate id= "BAE55037FF84FFDF6D375B0BFA303512" box= "[1324,1432,340,364]" degrees= "3.7897" direction= "west" orientation= "longitude" pageId= "21" pageNumber= "134" precision= "5" value= "-3.7897" > 3.7897°W</geoCoordinate>
,
<collectingRegion id= "1D15F812FF84FFDF6B235B2AFC2D35F3" box= "[824,901,373,397]" country= "Ivory Coast" name= "Comoe" pageId= "21" pageNumber= "134" > Comoé</collectingRegion>
<collectingCounty id= "360F4E7CFF84FFDF6B895B29FBB535F3" box= "[914,1053,373,397]" pageId= "21" pageNumber= "134" > Res. Station</collectingCounty>
,
<quantity id= "18299B15FF84FFDF6C345B2AFBC335F3" box= "[1071,1131,373,397]" metricMagnitude= "2" metricUnit= "m" metricValue= "1.91" pageId= "21" pageNumber= "134" unit= "m" value= "191.0" >
<elevation id= "54FCD1C3FF84FFDF6C345B2AFBC335F3" box= "[1071,1131,373,397]" metricMagnitude= "2" metricUnit= "m" metricValue= "1.91" pageId= "21" pageNumber= "134" unit= "m" value= "191.0" > 191m</elevation>
</quantity>
, foragers on path halfway between station and river, Heinze
<date id= "AB6F1030FF84FFDF6CD95BC9FA9E35D0" box= "[1218,1334,406,430]" pageId= "21" pageNumber= "134" value= "2019-04-04" >
<collectingDate id= "BB2BE9D8FF84FFDF6CD95BC9FA9E35D0" box= "[1218,1334,406,430]" pageId= "21" pageNumber= "134" value= "2019-04-04" > 2019.04.04</collectingDate>
</date>
– CI 18’ , ‘
<specimenCode id= "8F779E8BFF84FFDF6B275BE8FC5435B1" box= "[828,1020,439,463]" collectionCode= "SMNG" country= "Germany" httpUri= "http://grbio.org/cool/b2rf-iikk" name= "Senckenberg Museum fuer Naturkunde Goerlitz" pageId= "21" pageNumber= "134" > SMNG20240630</specimenCode>
’ [unique specimen identifier];
<specimenCount id= "C9D7FD79FF84FFDF6D7E5BE8FC2B358E" count= "3" pageId= "21" pageNumber= "134" type= "worker" > three worker</specimenCount>
<typeStatus id= "006A8852FF84FFDF6B945B87FC51358E" box= "[911,1017,472,496]" pageId= "21" pageNumber= "134" type= "paratype" > paratypes</typeStatus>
with same labelling on another pin;
<specimenCount id= "C9D7FD79FF84FFDF6B235BA7FC67366E" box= "[824,975,504,528]" count= "3" pageId= "21" pageNumber= "134" type= "worker" > three worker</specimenCount>
<typeStatus id= "006A8852FF84FFDF6BFE5BA6FBE7366F" box= "[997,1103,505,529]" pageId= "21" pageNumber= "134" type= "paratype" > paratypes</typeStatus>
labelled ‘ IVO:
<geoCoordinate id= "BAE55037FF84FFDF6D3A5BA6FA30366F" box= "[1313,1432,505,529]" degrees= "8.84028" direction= "north" orientation= "latitude" pageId= "21" pageNumber= "134" precision= "1" value= "8.84028" > 8.84028°N</geoCoordinate>
,
<geoCoordinate id= "BAE55037FF84FFDF6B235845FC01364C" box= "[824,937,538,562]" degrees= "3.77111" direction= "west" orientation= "longitude" pageId= "21" pageNumber= "134" precision= "1" value= "-3.77111" > 3.77111°W</geoCoordinate>
,
<collectingRegion id= "1D15F812FF84FFDF6BA15845FBAF364C" box= "[954,1031,538,562]" country= "Ivory Coast" name= "Comoe" pageId= "21" pageNumber= "134" > Comoé</collectingRegion>
<collectingMunicipality id= "3F0AAC8AFF84FFDF6C0F5846FBE0364F" box= "[1044,1096,537,561]" pageId= "21" pageNumber= "134" > Park</collectingMunicipality>
,
<location id= "DA0E602BFF84FFDF6C415846FB35364F" LSID= "urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6C415846FB35364F" box= "[1114,1181,537,561]" county= "Res. Station" latitude= "8.7703" longLatPrecision= "7" longitude= "-3.7897" municipality= "Park" name= "Iringo" pageId= "21" pageNumber= "134" stateProvince= "Comoe" > Iringo</location>
<collectingRegion id= "1D15F812FF84FFDF6CB25846FB4D364F" box= "[1193,1253,537,561]" country= "Nigeria" name= "Rivers" pageId= "21" pageNumber= "134" > River</collectingRegion>
<quantity id= "18299B15FF84FFDF6CE95845FA96364F" box= "[1266,1342,538,562]" metricMagnitude= "2" metricUnit= "m" metricValue= "2.02" pageId= "21" pageNumber= "134" unit= "m" value= "202.0" >
<elevation id= "54FCD1C3FF84FFDF6CE95845FA96364F" box= "[1266,1342,538,562]" metricMagnitude= "2" metricUnit= "m" metricValue= "2.02" pageId= "21" pageNumber= "134" unit= "m" value= "202.0" > 202 m</elevation>
</quantity>
,
<location id= "DA0E602BFF84FFDF6D4B5846FA35364F" LSID= "urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6D4B5846FA35364F" box= "[1360,1437,537,561]" county= "Res. Station" latitude= "8.7703" longLatPrecision= "7" longitude= "-3.7897" municipality= "Park" name= "Heinze" pageId= "21" pageNumber= "134" stateProvince= "Comoe" > Heinze</location>
<date id= "AB6F1030FF84FFDF6B235865FC04362C" box= "[824,940,570,594]" pageId= "21" pageNumber= "134" value= "2019-04-06" >
<collectingDate id= "BB2BE9D8FF84FFDF6B235865FC04362C" box= "[824,940,570,594]" pageId= "21" pageNumber= "134" value= "2019-04-06" > 2019.04.06</collectingDate>
</date>
– CI 45’ ;
<specimenCount id= "C9D7FD79FF84FFDF6C0A5864FB23362C" box= "[1041,1163,570,594]" count= "2" pageId= "21" pageNumber= "134" type= "worker" > two worker</specimenCount>
<typeStatus id= "006A8852FF84FFDF6C895864FB54362D" box= "[1170,1276,571,595]" pageId= "21" pageNumber= "134" type= "paratype" > paratypes</typeStatus>
labelled ‘ IVO:
<geoCoordinate id= "BAE55037FF84FFDF6B235804FC37360D" box= "[824,927,603,627]" degrees= "8.7703" direction= "north" orientation= "latitude" pageId= "21" pageNumber= "134" precision= "5" value= "8.7703" > 8.7703°N</geoCoordinate>
,
<geoCoordinate id= "BAE55037FF84FFDF6BB35804FBBB360D" box= "[936,1043,603,627]" degrees= "3.7897" direction= "west" orientation= "longitude" pageId= "21" pageNumber= "134" precision= "5" value= "-3.7897" > 3.7897°W</geoCoordinate>
,
<collectingRegion id= "1D15F812FF84FFDF6C075804FBC1360D" box= "[1052,1129,603,627]" country= "Ivory Coast" name= "Comoe" pageId= "21" pageNumber= "134" > Comoé</collectingRegion>
<location id= "DA0E602BFF84FFDF6C6B5803FB5B360D" LSID= "urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6C6B5803FB5B360D" box= "[1136,1267,603,627]" county= "Res. Station" latitude= "8.7703" longLatPrecision= "7" longitude= "-3.7897" municipality= "Park" name= "Res. Station" pageId= "21" pageNumber= "134" stateProvince= "Comoe" > Res. Station</location>
,
<quantity id= "18299B15FF84FFDF6CE55804FA92360D" box= "[1278,1338,603,627]" metricMagnitude= "2" metricUnit= "m" metricValue= "1.91" pageId= "21" pageNumber= "134" unit= "m" value= "191.0" >
<elevation id= "54FCD1C3FF84FFDF6CE55804FA92360D" box= "[1278,1338,603,627]" metricMagnitude= "2" metricUnit= "m" metricValue= "1.91" pageId= "21" pageNumber= "134" unit= "m" value= "191.0" > 191m</elevation>
</quantity>
, foragers on path halfway, between station and river,
<location id= "DA0E602BFF84FFDF6D545823FA3436EA" LSID= "urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6D545823FA3436EA" box= "[1359,1436,636,660]" county= "Res. Station" latitude= "8.7703" longLatPrecision= "7" longitude= "-3.7897" municipality= "Park" name= "Heinze" pageId= "21" pageNumber= "134" stateProvince= "Comoe" > Heinze</location>
<date id= "AB6F1030FF84FFDF6B2358C2FC0336CB" box= "[824,939,669,693]" pageId= "21" pageNumber= "134" value= "2019-04-07" >
<collectingDate id= "BB2BE9D8FF84FFDF6B2358C2FC0336CB" box= "[824,939,669,693]" pageId= "21" pageNumber= "134" value= "2019-04-07" > 2019.04.07</collectingDate>
</date>
– CI 55’ ; depository SMN Görlitz.
</materialsCitation>
</paragraph>
<paragraph id= "DF6E36F0FF84FFDF6B5458E1FB323689" blockId= "21.[824,1437,241,1976]" pageId= "21" pageNumber= "134" >
<emphasis id= "EDA5EAE2FF84FFDF6B5458E1FBFD36A8" bold= "true" box= "[847,1109,702,726]" pageId= "21" pageNumber= "134" > All material examined</emphasis>
.
<materialsCitation id= "6FB93CADFF84FFDF6C7F58E1FB3E3689" country= "Ivory Coast" location= "Ivory Coast" pageId= "21" pageNumber= "134" specimenCount= "3" typeStatus= "holotype" >
Only the
<specimenCount id= "C9D7FD79FF84FFDF6CD658E1FA9536A8" box= "[1229,1341,702,726]" count= "3" pageId= "21" pageNumber= "134" type= "generic" typeStatus= "type" > three type</specimenCount>
samples from
<collectingCountry id= "A7C67660FF84FFDF6B685880FC593689" box= "[883,1009,735,759]" name= "Ivory Coast" pageId= "21" pageNumber= "134" > Ivory Coast</collectingCountry>
were available
</materialsCitation>
.
</paragraph>
<paragraph id= "DF6E36F0FF84FFDF6B5458A0FBE93747" blockId= "21.[824,1437,241,1976]" pageId= "21" pageNumber= "134" >
<emphasis id= "EDA5EAE2FF84FFDF6B5458A0FB8A3766" bold= "true" box= "[847,1058,767,792]" pageId= "21" pageNumber= "134" > Geographic range</emphasis>
.
<materialsCitation id= "6FB93CADFF84FFDF6C2E595FFB953747" country= "Ivory Coast" location= "Ivory Coast" pageId= "21" pageNumber= "134" specimenCount= "2" typeStatus= "holotype" >
Only known from the
<specimenCount id= "C9D7FD79FF84FFDF6D23595FFA343766" box= "[1336,1436,768,792]" count= "2" pageId= "21" pageNumber= "134" type= "generic" typeStatus= "type" > two type</specimenCount>
localities in
<collectingCountry id= "A7C67660FF84FFDF6BA7597EFB953747" box= "[956,1085,801,825]" name= "Ivory Coast" pageId= "21" pageNumber= "134" > Ivory Coast</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id= "97CB657BFF84FFDC6B54591EFDFB35D0" lastPageId= "22" lastPageNumber= "135" pageId= "21" pageNumber= "134" type= "diagnosis" >
<paragraph id= "DF6E36F0FF84FFDC6B54591EFD953532" blockId= "21.[824,1437,241,1976]" lastBlockId= "22.[151,764,241,496]" lastPageId= "22" lastPageNumber= "135" pageId= "21" pageNumber= "134" >
<emphasis id= "EDA5EAE2FF84FFDF6B54591EFC163727" bold= "true" box= "[847,958,833,857]" pageId= "21" pageNumber= "134" > Diagnosis</emphasis>
: --Worker (
<tableCitation id= "9253034BFF84FFDF6C4C591EFB033727" box= "[1111,1195,833,857]" captionStart= "Table 2" captionStartId= "26.[150,203,1476,1497]" captionText= "Table 2. Workers of the C. wroughtonii group with mean FL/FR> 1.040 and originating from Africa and theArabian Peninsula. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId= "21" pageNumber= "134" > Tab. 2</tableCitation>
, Figs 44– 48, key). Extremely small, CS 377 µm. Head elongated, CL/CW 1.189. Median third of anterior clypeal margin slightly concave, median third of occipital margin straight. Postocular distance large, PoOc/CL 0.457. Frons very narrow (FRS/CS 0.219), widening frontad (FL/FR 1.063), caudal of FRS level parallel. Eye without any microsetae and rather smalll, EYE/CS 0.232. Scape moderately long, SL/CS 0.786. Metanotal groove in profile in absolute terms rather deep (MGr/CS 3.25 %) but with shallow slopes to mesonotum and propodeum. Prododeal spines long (SP/CS 0.201), thin and acute; in profile deviating from longitudinal mesosomal axis by only 25°, in dorsal view slightly diverging and and slightly incurved; their bases narrow (SPBA/CS 0.240). Petiole very narrow and moderately high (PeW/CS 0.234, PeH/CS 0.333); in lateral view with a concave anterior face and a long weakly convex dorsal profile; petiole in dorsal view very slender, its node 1.5 fold as long as wide and narrowing frontad. Postpetiole rather wide and low (PpW/CS 0.454, PpH/CS 0.276); the sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Postpetiole in dorsal view with a concave anterior margin and convex sides. Anterior clypeus with fine transverse rugulae. Vertex with densely-arranged, flat-bottomed and moderately large foveolae (dFov 16.7). The largest foveolae show a flat tubercle around the hair base of 6– 8 µm diameter which is sometimes connected with the outer ring through 1– 2 fine microcarinulae (
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<figureCitation id= "47EA2A75FF84FFDF6C405D62FB18332B" box= "[1115,1200,1853,1877]" captionStart= "Figure 47– 48" captionStartId= "14.[151,215,1787,1809]" captionTargetBox= "[151,1398,261,1741]" captionTargetPageId= "14" captionText= "Figure 47– 48. Cardiocondyla heinzei n.sp., holotype, Comoe / Ivory Coast, 4 April 2019. (47) head surface between inner eye margin and paramedian vertex. (48) anterior dorsum of 1st gaster tergite. Figure 49– 51: Cardiocondyla neferka, holotype, Mampong / Ghana, 10 February 1970; from www.antweb.org CASENT0901747, images by Ryan Perry. (49) head in dorsal view; (50) lateral view; (51) dorsal view." figureDoi= "http://doi.org/10.5281/zenodo.13194048" httpUri= "https://zenodo.org/record/13194048/files/figure.png" pageId= "21" pageNumber= "134" > Fig. 47</figureCitation>
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). Whole lateral area of mesosoma microreticulate, promesontum foveolate-reticulate. Petiole microreticulate, postpetiole more shiny. First gaster tergite with a rather long and dense pubescence (PLG/CS 7.12%, sqPDG 4.14) and a weak microreticulum (
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<figureCitation id= "47EA2A75FF87FFDC69545B4DFE373554" box= "[335,415,274,298]" captionStart= "Figure 47– 48" captionStartId= "14.[151,215,1787,1809]" captionTargetBox= "[151,1398,261,1741]" captionTargetPageId= "14" captionText= "Figure 47– 48. Cardiocondyla heinzei n.sp., holotype, Comoe / Ivory Coast, 4 April 2019. (47) head surface between inner eye margin and paramedian vertex. (48) anterior dorsum of 1st gaster tergite. Figure 49– 51: Cardiocondyla neferka, holotype, Mampong / Ghana, 10 February 1970; from www.antweb.org CASENT0901747, images by Ryan Perry. (49) head in dorsal view; (50) lateral view; (51) dorsal view." figureDoi= "http://doi.org/10.5281/zenodo.13194048" httpUri= "https://zenodo.org/record/13194048/files/figure.png" pageId= "22" pageNumber= "135" > Fig. 48</figureCitation>
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). All body parts yellowish with exception of the blackish brown gaster.
</paragraph>
<paragraph id= "DF6E36F0FF87FFDC68B55B0BFDFB35D0" blockId= "22.[151,764,241,496]" pageId= "22" pageNumber= "135" >
<emphasis id= "EDA5EAE2FF87FFDC68B55B0BFE063512" bold= "true" box= "[174,430,340,364]" pageId= "22" pageNumber= "135" > Taxonomic comments.</emphasis>
As a combination of large CL/ CW and small PeW/CS and FRS/CS not confuse with other members of the
<taxonomicName id= "18D14D73FF87FFDC69985BC9FDAE35D0" authorityName= "Forel" authorityYear= "1890" box= "[387,518,406,430]" class= "Insecta" family= "Formicidae" genus= "Cardiocondyla" kingdom= "Animalia" order= "Hymenoptera" pageId= "22" pageNumber= "135" phylum= "Arthropoda" rank= "species" species= "wroughtonii" >
<emphasis id= "EDA5EAE2FF87FFDC69985BC9FDAE35D0" box= "[387,518,406,430]" italics= "true" pageId= "22" pageNumber= "135" > wroughtonii</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id= "97CB657BFF87FFDC68B55BE8FE2B358E" pageId= "22" pageNumber= "135" type= "biology_ecology" >
<paragraph id= "DF6E36F0FF87FFDC68B55BE8FE2B358E" blockId= "22.[151,764,241,496]" pageId= "22" pageNumber= "135" >
<emphasis id= "EDA5EAE2FF87FFDC68B55BE8FEA335B1" bold= "true" box= "[174,267,439,463]" pageId= "22" pageNumber= "135" > Biology.</emphasis>
The workers foraged on ground in both open and woody situations.
</paragraph>
</subSubSection>
</treatment>
</document>