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<document id="8E01CF62069A122C0F82523645AEB019" ID-DOI="10.25674/415" ID-ISSN="2509-9523" ID-Zenodo-Dep="13194032" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722631177589" checkinUser="felipe" docAuthor="Seifert, Bernhard" docDate="2024" docId="577887E6FF84FFDC68CF5D64FE2B358E" docLanguage="en" docName="SoilOrg.96.2.113-144.pdf" docOrigin="Soil Organisms (London, England) 96 (2)" docSource="http://dx.doi.org/10.25674/415" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Cardiocondyla heinzei Seifert, 2024, n.sp." docType="treatment" docVersion="2" lastPageNumber="135" masterDocId="AB41FF9EFF91FFCA681B5A5FFFA8347E" masterDocTitle="A taxonomic revision of the ants of the Cardiocondyla wroughtonii group (Hymenoptera: Formicidae) with a checklist of the Cardiocondyla species of the world" masterLastPageNumber="144" masterPageNumber="113" pageNumber="134" updateTime="1722679537643" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EC847B77A5E3BE69C809D498F1C1D826">A taxonomic revision of the ants of the Cardiocondyla wroughtonii group (Hymenoptera: Formicidae) with a checklist of the Cardiocondyla species of the world</mods:title>
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<mods:namePart id="F6428A4CCBF554A9142A9B40F7331B51">Seifert, Bernhard</mods:namePart>
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<mods:date id="E8AE146800D6973A75AC9F4E2D1B5D36">2024</mods:date>
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<mods:number id="785AE7F8A2E1EA4CB2FB4B1392D9EC23">2024-08-01</mods:number>
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<mods:identifier id="3BE17B0D98AF8C7ABD0A16D04367216B" type="Zenodo-Dep">13194032</mods:identifier>
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<treatment id="577887E6FF84FFDC68CF5D64FE2B358E" LSID="urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E" httpUri="http://treatment.plazi.org/id/577887E6FF84FFDC68CF5D64FE2B358E" lastPageId="22" lastPageNumber="135" pageId="21" pageNumber="134">
<subSubSection id="97CB657BFF84FFDF68CF5D64FDE53329" box="[212,589,1851,1879]" pageId="21" pageNumber="134" type="nomenclature">
<paragraph id="DF6E36F0FF84FFDF68CF5D64FDE53329" blockId="21.[212,589,1851,1879]" box="[212,589,1851,1879]" pageId="21" pageNumber="134">
<heading id="8426819CFF84FFDF68CF5D64FDE53329" box="[212,589,1851,1879]" fontSize="11" level="2" pageId="21" pageNumber="134" reason="5">
<emphasis id="EDA5EAE2FF84FFDF68CF5D64FDE53329" bold="true" box="[212,589,1851,1879]" pageId="21" pageNumber="134">
<taxonomicName id="18D14D73FF84FFDF68CF5D64FDA83329" authorityName="Seifert" authorityYear="2024" box="[212,512,1851,1879]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="134" phylum="Arthropoda" rank="species" species="heinzei" status="sp. nov.">
<emphasis id="EDA5EAE2FF84FFDF68CF5D64FDA83329" bold="true" box="[212,512,1851,1879]" italics="true" pageId="21" pageNumber="134">Cardiocondyla heinzei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F6965799FF84FFDF6A115D64FDE53329" box="[522,589,1851,1879]" pageId="21" pageNumber="134" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF84FFDF68CF5D21FC783554" pageId="21" pageNumber="134" type="etymology">
<paragraph id="DF6E36F0FF84FFDF68CF5D21FC8A33E9" blockId="21.[189,802,1918,1975]" box="[212,802,1918,1943]" pageId="21" pageNumber="134">Etymology: the name is given in honor of Jürgen</paragraph>
<paragraph id="DF6E36F0FF84FFDF68A65DC0FC783554" blockId="21.[189,802,1918,1975]" lastBlockId="21.[824,1437,241,1976]" pageId="21" pageNumber="134">
Heinze wo has done and supervised outstanding research on the fascinating biology of
<taxonomicName id="18D14D73FF84FFDF6C905AAEFA823577" authorityName="Emery" authorityYear="1869" box="[1163,1322,241,265]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="EDA5EAE2FF84FFDF6C905AAEFA823577" box="[1163,1322,241,265]" italics="true" pageId="21" pageNumber="134">Cardiocondyla</emphasis>
</taxonomicName>
ants over three decades.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF84FFDF6B545B6CFBE93747" pageId="21" pageNumber="134" type="materials_examined">
<paragraph id="DF6E36F0FF84FFDF6B545B6CFC503535" blockId="21.[824,1437,241,1976]" box="[847,1016,307,331]" pageId="21" pageNumber="134">
<emphasis id="EDA5EAE2FF84FFDF6B545B6CFC503535" bold="true" box="[847,1016,307,331]" pageId="21" pageNumber="134">Type material:</emphasis>
</paragraph>
<paragraph id="DF6E36F0FF84FFDF6B545B0BFAB636CB" blockId="21.[824,1437,241,1976]" pageId="21" pageNumber="134">
<materialsCitation id="6FB93CADFF84FFDF6B545B0BFAB636CB" collectingDate="2019-04-04" collectingDateMax="2019-04-07" collectingDateMin="2019-04-04" collectionCode="SMNG" county="Res. Station" elevation="191" latitude="8.7703" location="Iringo" longLatPrecision="7" longitude="-3.7897" municipality="Park" pageId="21" pageNumber="134" specimenCode="SMNG20240630" specimenCount="9" specimenCount-worker="8" stateProvince="Comoe" typeStatus="holotype">
<typeStatus id="006A8852FF84FFDF6B545B0BFC1B3512" box="[847,947,340,364]" pageId="21" pageNumber="134" type="holotype">Holotype</typeStatus>
worker labelled IVO:
<geoCoordinate id="BAE55037FF84FFDF6CDF5B0BFA803512" box="[1220,1320,340,364]" degrees="8.7703" direction="north" orientation="latitude" pageId="21" pageNumber="134" precision="5" value="8.7703">8.7703°N</geoCoordinate>
,
<geoCoordinate id="BAE55037FF84FFDF6D375B0BFA303512" box="[1324,1432,340,364]" degrees="3.7897" direction="west" orientation="longitude" pageId="21" pageNumber="134" precision="5" value="-3.7897">3.7897°W</geoCoordinate>
,
<collectingRegion id="1D15F812FF84FFDF6B235B2AFC2D35F3" box="[824,901,373,397]" country="Ivory Coast" name="Comoe" pageId="21" pageNumber="134">Comoé</collectingRegion>
<collectingCounty id="360F4E7CFF84FFDF6B895B29FBB535F3" box="[914,1053,373,397]" pageId="21" pageNumber="134">Res. Station</collectingCounty>
,
<quantity id="18299B15FF84FFDF6C345B2AFBC335F3" box="[1071,1131,373,397]" metricMagnitude="2" metricUnit="m" metricValue="1.91" pageId="21" pageNumber="134" unit="m" value="191.0">
<elevation id="54FCD1C3FF84FFDF6C345B2AFBC335F3" box="[1071,1131,373,397]" metricMagnitude="2" metricUnit="m" metricValue="1.91" pageId="21" pageNumber="134" unit="m" value="191.0">191m</elevation>
</quantity>
, foragers on path halfway between station and river, Heinze
<date id="AB6F1030FF84FFDF6CD95BC9FA9E35D0" box="[1218,1334,406,430]" pageId="21" pageNumber="134" value="2019-04-04">
<collectingDate id="BB2BE9D8FF84FFDF6CD95BC9FA9E35D0" box="[1218,1334,406,430]" pageId="21" pageNumber="134" value="2019-04-04">2019.04.04</collectingDate>
</date>
CI 18,
<specimenCode id="8F779E8BFF84FFDF6B275BE8FC5435B1" box="[828,1020,439,463]" collectionCode="SMNG" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="21" pageNumber="134">SMNG20240630</specimenCode>
[unique specimen identifier];
<specimenCount id="C9D7FD79FF84FFDF6D7E5BE8FC2B358E" count="3" pageId="21" pageNumber="134" type="worker">three worker</specimenCount>
<typeStatus id="006A8852FF84FFDF6B945B87FC51358E" box="[911,1017,472,496]" pageId="21" pageNumber="134" type="paratype">paratypes</typeStatus>
with same labelling on another pin;
<specimenCount id="C9D7FD79FF84FFDF6B235BA7FC67366E" box="[824,975,504,528]" count="3" pageId="21" pageNumber="134" type="worker">three worker</specimenCount>
<typeStatus id="006A8852FF84FFDF6BFE5BA6FBE7366F" box="[997,1103,505,529]" pageId="21" pageNumber="134" type="paratype">paratypes</typeStatus>
labelled IVO:
<geoCoordinate id="BAE55037FF84FFDF6D3A5BA6FA30366F" box="[1313,1432,505,529]" degrees="8.84028" direction="north" orientation="latitude" pageId="21" pageNumber="134" precision="1" value="8.84028">8.84028°N</geoCoordinate>
,
<geoCoordinate id="BAE55037FF84FFDF6B235845FC01364C" box="[824,937,538,562]" degrees="3.77111" direction="west" orientation="longitude" pageId="21" pageNumber="134" precision="1" value="-3.77111">3.77111°W</geoCoordinate>
,
<collectingRegion id="1D15F812FF84FFDF6BA15845FBAF364C" box="[954,1031,538,562]" country="Ivory Coast" name="Comoe" pageId="21" pageNumber="134">Comoé</collectingRegion>
<collectingMunicipality id="3F0AAC8AFF84FFDF6C0F5846FBE0364F" box="[1044,1096,537,561]" pageId="21" pageNumber="134">Park</collectingMunicipality>
,
<location id="DA0E602BFF84FFDF6C415846FB35364F" LSID="urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6C415846FB35364F" box="[1114,1181,537,561]" county="Res. Station" latitude="8.7703" longLatPrecision="7" longitude="-3.7897" municipality="Park" name="Iringo" pageId="21" pageNumber="134" stateProvince="Comoe">Iringo</location>
<collectingRegion id="1D15F812FF84FFDF6CB25846FB4D364F" box="[1193,1253,537,561]" country="Nigeria" name="Rivers" pageId="21" pageNumber="134">River</collectingRegion>
<quantity id="18299B15FF84FFDF6CE95845FA96364F" box="[1266,1342,538,562]" metricMagnitude="2" metricUnit="m" metricValue="2.02" pageId="21" pageNumber="134" unit="m" value="202.0">
<elevation id="54FCD1C3FF84FFDF6CE95845FA96364F" box="[1266,1342,538,562]" metricMagnitude="2" metricUnit="m" metricValue="2.02" pageId="21" pageNumber="134" unit="m" value="202.0">202 m</elevation>
</quantity>
,
<location id="DA0E602BFF84FFDF6D4B5846FA35364F" LSID="urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6D4B5846FA35364F" box="[1360,1437,537,561]" county="Res. Station" latitude="8.7703" longLatPrecision="7" longitude="-3.7897" municipality="Park" name="Heinze" pageId="21" pageNumber="134" stateProvince="Comoe">Heinze</location>
<date id="AB6F1030FF84FFDF6B235865FC04362C" box="[824,940,570,594]" pageId="21" pageNumber="134" value="2019-04-06">
<collectingDate id="BB2BE9D8FF84FFDF6B235865FC04362C" box="[824,940,570,594]" pageId="21" pageNumber="134" value="2019-04-06">2019.04.06</collectingDate>
</date>
CI 45;
<specimenCount id="C9D7FD79FF84FFDF6C0A5864FB23362C" box="[1041,1163,570,594]" count="2" pageId="21" pageNumber="134" type="worker">two worker</specimenCount>
<typeStatus id="006A8852FF84FFDF6C895864FB54362D" box="[1170,1276,571,595]" pageId="21" pageNumber="134" type="paratype">paratypes</typeStatus>
labelled IVO:
<geoCoordinate id="BAE55037FF84FFDF6B235804FC37360D" box="[824,927,603,627]" degrees="8.7703" direction="north" orientation="latitude" pageId="21" pageNumber="134" precision="5" value="8.7703">8.7703°N</geoCoordinate>
,
<geoCoordinate id="BAE55037FF84FFDF6BB35804FBBB360D" box="[936,1043,603,627]" degrees="3.7897" direction="west" orientation="longitude" pageId="21" pageNumber="134" precision="5" value="-3.7897">3.7897°W</geoCoordinate>
,
<collectingRegion id="1D15F812FF84FFDF6C075804FBC1360D" box="[1052,1129,603,627]" country="Ivory Coast" name="Comoe" pageId="21" pageNumber="134">Comoé</collectingRegion>
<location id="DA0E602BFF84FFDF6C6B5803FB5B360D" LSID="urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6C6B5803FB5B360D" box="[1136,1267,603,627]" county="Res. Station" latitude="8.7703" longLatPrecision="7" longitude="-3.7897" municipality="Park" name="Res. Station" pageId="21" pageNumber="134" stateProvince="Comoe">Res. Station</location>
,
<quantity id="18299B15FF84FFDF6CE55804FA92360D" box="[1278,1338,603,627]" metricMagnitude="2" metricUnit="m" metricValue="1.91" pageId="21" pageNumber="134" unit="m" value="191.0">
<elevation id="54FCD1C3FF84FFDF6CE55804FA92360D" box="[1278,1338,603,627]" metricMagnitude="2" metricUnit="m" metricValue="1.91" pageId="21" pageNumber="134" unit="m" value="191.0">191m</elevation>
</quantity>
, foragers on path halfway, between station and river,
<location id="DA0E602BFF84FFDF6D545823FA3436EA" LSID="urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6D545823FA3436EA" box="[1359,1436,636,660]" county="Res. Station" latitude="8.7703" longLatPrecision="7" longitude="-3.7897" municipality="Park" name="Heinze" pageId="21" pageNumber="134" stateProvince="Comoe">Heinze</location>
<date id="AB6F1030FF84FFDF6B2358C2FC0336CB" box="[824,939,669,693]" pageId="21" pageNumber="134" value="2019-04-07">
<collectingDate id="BB2BE9D8FF84FFDF6B2358C2FC0336CB" box="[824,939,669,693]" pageId="21" pageNumber="134" value="2019-04-07">2019.04.07</collectingDate>
</date>
CI 55; depository SMN Görlitz.
</materialsCitation>
</paragraph>
<paragraph id="DF6E36F0FF84FFDF6B5458E1FB323689" blockId="21.[824,1437,241,1976]" pageId="21" pageNumber="134">
<emphasis id="EDA5EAE2FF84FFDF6B5458E1FBFD36A8" bold="true" box="[847,1109,702,726]" pageId="21" pageNumber="134">All material examined</emphasis>
.
<materialsCitation id="6FB93CADFF84FFDF6C7F58E1FB3E3689" country="Ivory Coast" location="Ivory Coast" pageId="21" pageNumber="134" specimenCount="3" typeStatus="holotype">
Only the
<specimenCount id="C9D7FD79FF84FFDF6CD658E1FA9536A8" box="[1229,1341,702,726]" count="3" pageId="21" pageNumber="134" type="generic" typeStatus="type">three type</specimenCount>
samples from
<collectingCountry id="A7C67660FF84FFDF6B685880FC593689" box="[883,1009,735,759]" name="Ivory Coast" pageId="21" pageNumber="134">Ivory Coast</collectingCountry>
were available
</materialsCitation>
.
</paragraph>
<paragraph id="DF6E36F0FF84FFDF6B5458A0FBE93747" blockId="21.[824,1437,241,1976]" pageId="21" pageNumber="134">
<emphasis id="EDA5EAE2FF84FFDF6B5458A0FB8A3766" bold="true" box="[847,1058,767,792]" pageId="21" pageNumber="134">Geographic range</emphasis>
.
<materialsCitation id="6FB93CADFF84FFDF6C2E595FFB953747" country="Ivory Coast" location="Ivory Coast" pageId="21" pageNumber="134" specimenCount="2" typeStatus="holotype">
Only known from the
<specimenCount id="C9D7FD79FF84FFDF6D23595FFA343766" box="[1336,1436,768,792]" count="2" pageId="21" pageNumber="134" type="generic" typeStatus="type">two type</specimenCount>
localities in
<collectingCountry id="A7C67660FF84FFDF6BA7597EFB953747" box="[956,1085,801,825]" name="Ivory Coast" pageId="21" pageNumber="134">Ivory Coast</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF84FFDC6B54591EFDFB35D0" lastPageId="22" lastPageNumber="135" pageId="21" pageNumber="134" type="diagnosis">
<paragraph id="DF6E36F0FF84FFDC6B54591EFD953532" blockId="21.[824,1437,241,1976]" lastBlockId="22.[151,764,241,496]" lastPageId="22" lastPageNumber="135" pageId="21" pageNumber="134">
<emphasis id="EDA5EAE2FF84FFDF6B54591EFC163727" bold="true" box="[847,958,833,857]" pageId="21" pageNumber="134">Diagnosis</emphasis>
: --Worker (
<tableCitation id="9253034BFF84FFDF6C4C591EFB033727" box="[1111,1195,833,857]" captionStart="Table 2" captionStartId="26.[150,203,1476,1497]" captionText="Table 2. Workers of the C. wroughtonii group with mean FL/FR&gt; 1.040 and originating from Africa and theArabian Peninsula. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="21" pageNumber="134">Tab. 2</tableCitation>
, Figs 4448, key). Extremely small, CS 377 µm. Head elongated, CL/CW 1.189. Median third of anterior clypeal margin slightly concave, median third of occipital margin straight. Postocular distance large, PoOc/CL 0.457. Frons very narrow (FRS/CS 0.219), widening frontad (FL/FR 1.063), caudal of FRS level parallel. Eye without any microsetae and rather smalll, EYE/CS 0.232. Scape moderately long, SL/CS 0.786. Metanotal groove in profile in absolute terms rather deep (MGr/CS 3.25 %) but with shallow slopes to mesonotum and propodeum. Prododeal spines long (SP/CS 0.201), thin and acute; in profile deviating from longitudinal mesosomal axis by only 25°, in dorsal view slightly diverging and and slightly incurved; their bases narrow (SPBA/CS 0.240). Petiole very narrow and moderately high (PeW/CS 0.234, PeH/CS 0.333); in lateral view with a concave anterior face and a long weakly convex dorsal profile; petiole in dorsal view very slender, its node 1.5 fold as long as wide and narrowing frontad. Postpetiole rather wide and low (PpW/CS 0.454, PpH/CS 0.276); the sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Postpetiole in dorsal view with a concave anterior margin and convex sides. Anterior clypeus with fine transverse rugulae. Vertex with densely-arranged, flat-bottomed and moderately large foveolae (dFov 16.7). The largest foveolae show a flat tubercle around the hair base of 68 µm diameter which is sometimes connected with the outer ring through 12 fine microcarinulae (
<figureCitation id="47EA2A75FF84FFDF6C405D62FB18332B" box="[1115,1200,1853,1877]" captionStart="Figure 4748" captionStartId="14.[151,215,1787,1809]" captionTargetBox="[151,1398,261,1741]" captionTargetPageId="14" captionText="Figure 4748. Cardiocondyla heinzei n.sp., holotype, Comoe / Ivory Coast, 4 April 2019. (47) head surface between inner eye margin and paramedian vertex. (48) anterior dorsum of 1st gaster tergite. Figure 4951: Cardiocondyla neferka, holotype, Mampong / Ghana, 10 February 1970; from www.antweb.org CASENT0901747, images by Ryan Perry. (49) head in dorsal view; (50) lateral view; (51) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13194048" httpUri="https://zenodo.org/record/13194048/files/figure.png" pageId="21" pageNumber="134">Fig. 47</figureCitation>
). Whole lateral area of mesosoma microreticulate, promesontum foveolate-reticulate. Petiole microreticulate, postpetiole more shiny. First gaster tergite with a rather long and dense pubescence (PLG/CS 7.12%, sqPDG 4.14) and a weak microreticulum (
<figureCitation id="47EA2A75FF87FFDC69545B4DFE373554" box="[335,415,274,298]" captionStart="Figure 4748" captionStartId="14.[151,215,1787,1809]" captionTargetBox="[151,1398,261,1741]" captionTargetPageId="14" captionText="Figure 4748. Cardiocondyla heinzei n.sp., holotype, Comoe / Ivory Coast, 4 April 2019. (47) head surface between inner eye margin and paramedian vertex. (48) anterior dorsum of 1st gaster tergite. Figure 4951: Cardiocondyla neferka, holotype, Mampong / Ghana, 10 February 1970; from www.antweb.org CASENT0901747, images by Ryan Perry. (49) head in dorsal view; (50) lateral view; (51) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13194048" httpUri="https://zenodo.org/record/13194048/files/figure.png" pageId="22" pageNumber="135">Fig. 48</figureCitation>
). All body parts yellowish with exception of the blackish brown gaster.
</paragraph>
<paragraph id="DF6E36F0FF87FFDC68B55B0BFDFB35D0" blockId="22.[151,764,241,496]" pageId="22" pageNumber="135">
<emphasis id="EDA5EAE2FF87FFDC68B55B0BFE063512" bold="true" box="[174,430,340,364]" pageId="22" pageNumber="135">Taxonomic comments.</emphasis>
As a combination of large CL/ CW and small PeW/CS and FRS/CS not confuse with other members of the
<taxonomicName id="18D14D73FF87FFDC69985BC9FDAE35D0" authorityName="Forel" authorityYear="1890" box="[387,518,406,430]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="135" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF87FFDC69985BC9FDAE35D0" box="[387,518,406,430]" italics="true" pageId="22" pageNumber="135">wroughtonii</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF87FFDC68B55BE8FE2B358E" pageId="22" pageNumber="135" type="biology_ecology">
<paragraph id="DF6E36F0FF87FFDC68B55BE8FE2B358E" blockId="22.[151,764,241,496]" pageId="22" pageNumber="135">
<emphasis id="EDA5EAE2FF87FFDC68B55BE8FEA335B1" bold="true" box="[174,267,439,463]" pageId="22" pageNumber="135">Biology.</emphasis>
The workers foraged on ground in both open and woody situations.
</paragraph>
</subSubSection>
</treatment>
</document>