420 lines
77 KiB
XML
420 lines
77 KiB
XML
<document id="C4A6946FDB9A701B8F31737B77F3D539" ID-DOI="10.1093/zoolinnean/zlad140" ID-ISSN="0024-4082" ID-Zenodo-Dep="13220046" ID-ZooBank="F6E6171-16B7-43DA-A064-6D13AFD99FC1" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1722844718276" checkinUser="plazi" docAuthor="Han, Yuru, Zhou, Yadong, Chen, Chong & Wang, Yueyun" docDate="2024" docId="9C126744BA6DFFF3FC119224DC48C780" docLanguage="en" docName="zlad140.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad140" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.6:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="6" docTitle="Levensteiniella pettiboneae Han, Zhou, Chen & Wang, 2024, sp.nov." docType="treatment" docVersion="3" lastPageNumber="294" masterDocId="602B1F3CBA6FFFF7FFD39454DF36C622" masterDocTitle="Diversity and biogeography of scale worms in the subfamily Lepidonotopodinae (Annelida: Polynoidae) from Indian Ocean hydrothermal vents with descriptions of four new species" masterLastPageNumber="304" masterPageNumber="290" pageNumber="292" updateTime="1723569002591" updateUser="valdenar">
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<mods:titleInfo id="468FC7A1A97CBAC83B7DE0FD24FF5707">
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<mods:title id="28389956EFB1D26F3302E38D7DAEFB59">Diversity and biogeography of scale worms in the subfamily Lepidonotopodinae (Annelida: Polynoidae) from Indian Ocean hydrothermal vents with descriptions of four new species</mods:title>
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<mods:name id="664ECAECAC1A8627C9163F943E3614A6" type="personal">
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<mods:roleTerm id="A6B73718543B9F7204904030DADD3B22">Author</mods:roleTerm>
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<mods:namePart id="B9D65B6526C1DEE01137AC210BA03C89">Han, Yuru</mods:namePart>
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<mods:affiliation id="79E210A284AA6DB20DC2FA91DFC47DBF">Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou, China</mods:affiliation>
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<mods:roleTerm id="8BC8DB08A1747AC7B1137F7550EAAE5F">Author</mods:roleTerm>
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<mods:namePart id="DB2EAC98596478673B608910A235FEC1">Zhou, Yadong</mods:namePart>
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<mods:affiliation id="F90C18403BC9D79E7578FB93AF1629F3">Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou, China & Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519082, China</mods:affiliation>
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<mods:nameIdentifier id="BFBF547248DB82183EC732FF86548DCC" type="email">yadong_zhou@sio.org.cn</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="BC2EA4916930F82F9C8303B533DF1E28">Author</mods:roleTerm>
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<mods:namePart id="419C2BB256B7D6BD8E28AB753C71C9CF">Chen, Chong</mods:namePart>
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<mods:affiliation id="3E1896518AB5F97F796AC08D2922C743">X-STAR, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), 2 - 15 Natsushima-cho, Yokosuka, Kanagawa Prefecture 237 - 0061, Japan</mods:affiliation>
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</mods:role>
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<mods:namePart id="A1FBC7596DEAE559CFA25ACB4EA53BC3">Wang, Yueyun</mods:namePart>
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<mods:affiliation id="AC3B402957F12BB29810C1EABF7563E7">Key Laboratory of Marine Ecosystem Dynamics, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou, China</mods:affiliation>
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<mods:title id="230B420B2B20B2EB34C30CADAA5CCC20">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="DB2771E16E5C6E201923E0F66A0F0DF8">2024</mods:date>
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<mods:number id="D8964F58D361507F4950515111F9FEB0">2023-10-17</mods:number>
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<mods:url id="92B9A92D11844FC03E19694DEECAB592">http://dx.doi.org/10.1093/zoolinnean/zlad140</mods:url>
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<mods:classification id="CEC243E1731E2E85D9E0100C7D153FEA">journal article</mods:classification>
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<mods:identifier id="C0F68E6BEB36DE665BA68C5C38E7B71F" type="DOI">10.1093/zoolinnean/zlad140</mods:identifier>
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<treatment id="9C126744BA6DFFF3FC119224DC48C780" ID-DOI="http://doi.org/10.5281/zenodo.13220038" ID-Zenodo-Dep="13220038" LSID="urn:lsid:plazi:treatment:9C126744BA6DFFF3FC119224DC48C780" httpUri="http://treatment.plazi.org/id/9C126744BA6DFFF3FC119224DC48C780" lastPageId="4" lastPageNumber="294" pageId="2" pageNumber="292">
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<subSubSection id="5CA185D9BA6DFFF5FC119224DA2DC0A9" box="[962,1307,1648,1675]" pageId="2" pageNumber="292" type="nomenclature">
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<paragraph id="1404D652BA6DFFF5FC119224DA2DC0A9" blockId="2.[962,1307,1648,1675]" box="[962,1307,1648,1675]" pageId="2" pageNumber="292">
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<heading id="4F4C613EBA6DFFF5FC119224DA2DC0A9" bold="true" box="[962,1307,1648,1675]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="292" reason="2">
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<emphasis id="26CF0A40BA6DFFF5FC119224DA2DC0A9" bold="true" box="[962,1307,1648,1675]" pageId="2" pageNumber="292">
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<taxonomicName id="D3BBADD1BA6DFFF5FC119224DBFCC0A9" ID-CoL="BV3M5" authorityName="Han & Zhou & Chen & Wang" authorityYear="2024" box="[962,1226,1648,1675]" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="292" phylum="Annelida" rank="species" species="pettiboneae" status="sp. nov.">
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<emphasis id="26CF0A40BA6DFFF5FC119224DBFCC0A9" bold="true" box="[962,1226,1648,1675]" italics="true" pageId="2" pageNumber="292">Levensteiniella pettiboneae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3DFCB73BBA6DFFF5FB1C9225DA2DC0A9" box="[1231,1307,1649,1675]" pageId="2" pageNumber="292" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5CA185D9BA6DFFF3FB9492F7DC48C780" lastPageId="4" lastPageNumber="294" pageId="2" pageNumber="292" type="description">
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<paragraph id="1404D652BA6DFFF5FB9492F7DBA0C09F" blockId="2.[1095,1174,1699,1725]" box="[1095,1174,1699,1725]" pageId="2" pageNumber="292">
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<emphasis id="26CF0A40BA6DFFF5FB9492F7DBA0C09F" bold="true" box="[1095,1174,1699,1725]" pageId="2" pageNumber="292">
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||
(
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<figureCitation id="8C80CAD7BA6DFFF5FB8192F7DBBDC09F" box="[1106,1163,1699,1725]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="2" pageNumber="292">Fig. 2</figureCitation>
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)
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</emphasis>
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</paragraph>
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<paragraph id="1404D652BA6DFFF5FCF992B7DBBBC138" blockId="2.[810,1458,1763,1850]" pageId="2" pageNumber="292">
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<emphasis id="26CF0A40BA6DFFF5FCF992B7DB25C0D9" box="[810,1043,1763,1787]" italics="true" pageId="2" pageNumber="292">ZooBank registration:</emphasis>
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||
<uri id="602ADA50BA6DFFF5FBFC92B7DBBFC138" pageId="2" pageNumber="292">
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||
urn:lsid:zoobank.org:act:
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||
<uuid id="601DEC87BA6DFFF5FAE292B7DBBFC138" pageId="2" pageNumber="292">73D3C46A-9708-436F-8C70-022D17950010</uuid>
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||
</uri>
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||
.
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||
</paragraph>
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<paragraph id="1404D652BA6DFFF5FC969376DBEBC11B" blockId="2.[810,1458,1763,1850]" box="[837,1245,1826,1850]" pageId="2" pageNumber="292">
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<taxonomicName id="D3BBADD1BA6DFFF5FC969376DC8EC118" authorityName="Kinberg" authorityYear="1856" box="[837,952,1826,1850]" class="Polychaeta" family="Polynoidae" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="292" phylum="Annelida" rank="family">Polynoidae</taxonomicName>
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indet. #4—
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<bibRefCitation id="702AABA3BA6DFFF5FBE09376DBE1C118" author="Zhou Y & Chen C & Zhang D" box="[1075,1239,1826,1850]" pageId="2" pageNumber="292" pagination="2858 - 70" refId="ref13789" refString="Zhou Y, Chen C, Zhang D et al. Delineating biogeographic regions in Indian Ocean deep-sea vents and implications for conservation. Diversity and Distribution 2022; 28: 2858 - 70." type="journal article" year="2022">
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Zhou
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<emphasis id="26CF0A40BA6DFFF5FBA39377DBA9C118" box="[1136,1183,1826,1850]" italics="true" pageId="2" pageNumber="292">et al.</emphasis>
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2022
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</bibRefCitation>
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||
.
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</paragraph>
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<paragraph id="1404D652BA6DFFF4FCF99308DD2AC04F" blockId="2.[810,1459,1883,1970]" lastBlockId="3.[129,777,1589,1645]" lastPageId="3" lastPageNumber="293" pageId="2" pageNumber="292">
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<emphasis id="26CF0A40BA6DFFF5FCF99308DCA6C151" box="[810,912,1884,1907]" italics="true" pageId="2" pageNumber="292">Diagnosis:</emphasis>
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Moderate-sized
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<taxonomicName id="D3BBADD1BA6DFFF5FBEA930FDBF0C151" authorityName="Pettibone" authorityYear="1985" box="[1081,1222,1883,1907]" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="2" pageNumber="292" phylum="Annelida" rank="genus">
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<emphasis id="26CF0A40BA6DFFF5FBEA930FDBF0C151" box="[1081,1222,1883,1907]" italics="true" pageId="2" pageNumber="292">Levensteiniella</emphasis>
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</taxonomicName>
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with oval prostomium; iridescent elytra 11 pairs, with micropapillae on posterior and lateral border; 10 pairs of inconspicuous rounded ventral papillae on segments 11–20 on female; two pairs of long ventral papillae on segments 11 and 12 on male.
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</paragraph>
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<caption id="40C486DABA6CFFF4FF529111DBEAC024" ID-DOI="http://doi.org/10.5281/zenodo.13220050" ID-Zenodo-Dep="13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="3" pageNumber="293" startId="3.[129,194,1349,1373]" targetBox="[129,1473,142,1321]" targetPageId="3" targetType="figure">
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||
<paragraph id="1404D652BA6CFFF4FF529111DBEAC024" blockId="3.[129,1466,1349,1542]" pageId="3" pageNumber="293">
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<emphasis id="26CF0A40BA6CFFF4FF529111DFEEC37F" bold="true" box="[129,216,1349,1373]" pageId="3" pageNumber="293">Figure 2.</emphasis>
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<taxonomicName id="D3BBADD1BA6CFFF4FF0E9111DEFFC37F" authorityName="Han & Zhou & Chen & Wang" authorityYear="2024" box="[221,457,1349,1373]" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="293" phylum="Annelida" rank="species" species="pettiboneae" status="sp. nov.">
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<emphasis id="26CF0A40BA6CFFF4FF0E9111DEFFC37F" box="[221,457,1349,1373]" italics="true" pageId="3" pageNumber="293">Levensteiniella pettiboneae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3DFCB73BBA6CFFF4FE1D9112DD23C37F" box="[462,533,1350,1373]" pageId="3" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
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Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217. Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm.
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</paragraph>
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</caption>
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||
<paragraph id="1404D652BA6CFFF4FF5292DBDD1DC084" blockId="3.[129,555,1678,1702]" box="[129,555,1678,1702]" pageId="3" pageNumber="293">
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<materialsCitation id="A4D3DC0FBA6CFFF4FF5292DBDD11C084" box="[129,551,1678,1702]" collectionCode="SWIR" pageId="3" pageNumber="293" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="26CF0A40BA6CFFF4FF5292DBDE35C084" box="[129,259,1678,1702]" italics="true" pageId="3" pageNumber="293">
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||
<typeStatus id="CB0068F0BA6CFFF4FF5292DBDF87C084" box="[129,177,1679,1702]" pageId="3" pageNumber="293">Type</typeStatus>
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locality:
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</emphasis>
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Tiancheng vent field,
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<collectionCode id="72AA4E97BA6CFFF4FE3092DADD11C084" box="[483,551,1678,1702]" pageId="3" pageNumber="293">SWIR</collectionCode>
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||
</materialsCitation>
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||
.
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||
</paragraph>
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<paragraph id="1404D652BA6CFFF4FF529285DBF3C1E7" blockId="3.[129,777,1744,1987]" lastBlockId="3.[825,1475,1589,1989]" pageId="3" pageNumber="293">
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||
<emphasis id="26CF0A40BA6CFFF4FF529285DE22C0CA" box="[129,276,1744,1768]" italics="true" pageId="3" pageNumber="293">Type material:</emphasis>
|
||
<materialsCitation id="A4D3DC0FBA6CFFF4FECF9284DD6DC164" collectingDate="2014-12-23" collectingDateMax="2017-03-14" collectingDateMin="2014-12-23" collectionCode="SWIR" elevation="2760" latitude="-23.88" longLatPrecision="752" longitude="69.6" pageId="3" pageNumber="293" specimenCode="RSIO35217, RSIO35219, RSIO35221, RSIO35222, RSIO35290, RSIO38129" specimenCount="7" typeStatus="paratype">
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||
Holytype (
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<specimenCode id="441D7E29BA6CFFF4FE439284DD2CC0CA" box="[400,538,1744,1768]" collectionCode="RSIO" pageId="3" pageNumber="293">RSIO35217</specimenCode>
|
||
): female, 25 segments,
|
||
<quantity id="D3437BB7BA6CFFF4FF5292BBDFC4C125" box="[129,242,1775,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.397" pageId="3" pageNumber="293" unit="mm" value="13.97">13.97 mm</quantity>
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||
in length,
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||
<quantity id="D3437BB7BA6CFFF4FEAA92BBDEE8C125" box="[377,478,1775,1799]" metricMagnitude="-3" metricUnit="m" metricValue="6.95" pageId="3" pageNumber="293" unit="mm" value="6.95">6.95 mm</quantity>
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in width, Tiancheng vent field (
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<geoCoordinate id="718FB095BA6CFFF4FF11935ADE21C104" box="[194,279,1806,1830]" degrees="27.95" direction="south" orientation="latitude" pageId="3" pageNumber="293" precision="555" value="-27.95">27.95°S</geoCoordinate>
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||
,
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||
<geoCoordinate id="718FB095BA6CFFF4FEFB935ADE4AC104" box="[296,380,1806,1830]" degrees="63.53" direction="east" orientation="longitude" pageId="3" pageNumber="293" precision="555" value="63.53">63.53°E</geoCoordinate>
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||
,
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||
<quantity id="D3437BB7BA6CFFF4FE5F935BDED5C104" box="[396,483,1807,1831]" metricMagnitude="3" metricUnit="m" metricValue="2.76" pageId="3" pageNumber="293" unit="m" value="2760.0">
|
||
<elevation id="9F963161BA6CFFF4FE5F935BDED5C104" box="[396,483,1807,1831]" metricMagnitude="3" metricUnit="m" metricValue="2.76" pageId="3" pageNumber="293" unit="m" value="2760.0">2760 m</elevation>
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||
</quantity>
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deep),
|
||
<collectionCode id="72AA4E97BA6CFFF4FDEE935BDDB7C105" box="[573,641,1807,1831]" pageId="3" pageNumber="293">SWIR</collectionCode>
|
||
, Dive87,
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||
<date id="6005F092BA6CFFF4FD3C935BDE06C164" pageId="3" pageNumber="293" value="2014-12-23">
|
||
<collectingDate id="7041097ABA6CFFF4FD3C935BDE06C164" pageId="3" pageNumber="293" value="2014-12-23">23 December 2014</collectingDate>
|
||
</date>
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. Paratype 1 (
|
||
<specimenCode id="441D7E29BA6CFFF4FE16937ADD79C164" box="[453,591,1838,1862]" collectionCode="RSIO" pageId="3" pageNumber="293">RSIO35219</specimenCode>
|
||
):
|
||
</materialsCitation>
|
||
<materialsCitation id="A4D3DC0FBA6CFFF4FDB6937ADD00C186" collectingDate="2014-12-23" collectingDateMax="2017-03-14" collectingDateMin="2014-12-23" collectionCode="SWIR" elevation="2760" latitude="-23.88" longLatPrecision="752" longitude="69.6" pageId="3" pageNumber="293" specimenCode="RSIO35217, RSIO35219, RSIO35221, RSIO35222, RSIO35290, RSIO38129" specimenCount="7" typeStatus="paratype">
|
||
female, 25 segments,
|
||
<quantity id="D3437BB7BA6CFFF4FF029319DE04C147" box="[209,306,1869,1893]" metricMagnitude="-3" metricUnit="m" metricValue="6.43" pageId="3" pageNumber="293" unit="mm" value="6.43">6.43 mm</quantity>
|
||
in length,
|
||
<quantity id="D3437BB7BA6CFFF4FE7C9319DD39C147" box="[431,527,1869,1893]" metricMagnitude="-3" metricUnit="m" metricValue="4.64" pageId="3" pageNumber="293" unit="mm" value="4.64">4.64 mm</quantity>
|
||
in width, Longqi vent field (
|
||
<geoCoordinate id="718FB095BA6CFFF4FF119339DE21C1A7" box="[194,279,1901,1925]" degrees="37.78" direction="south" orientation="latitude" pageId="3" pageNumber="293" precision="555" value="-37.78">37.78°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="718FB095BA6CFFF4FEFB9338DE4AC1A6" box="[296,380,1900,1924]" degrees="49.65" direction="east" orientation="longitude" pageId="3" pageNumber="293" precision="555" value="49.65">49.65°E</geoCoordinate>
|
||
,
|
||
<quantity id="D3437BB7BA6CFFF4FE5F9339DED5C1A6" box="[396,483,1901,1925]" metricMagnitude="3" metricUnit="m" metricValue="2.7399999999999998" pageId="3" pageNumber="293" unit="m" value="2740.0">
|
||
<elevation id="9F963161BA6CFFF4FE5F9339DED5C1A6" box="[396,483,1901,1925]" metricMagnitude="3" metricUnit="m" metricValue="2.7399999999999998" pageId="3" pageNumber="293" unit="m" value="2740.0">2740 m</elevation>
|
||
</quantity>
|
||
deep),
|
||
<collectionCode id="72AA4E97BA6CFFF4FDEE9339DDB7C1A7" box="[573,641,1901,1925]" pageId="3" pageNumber="293">SWIR</collectionCode>
|
||
, Dive100,
|
||
<date id="6005F092BA6CFFF4FD2F9339DE2AC186" pageId="3" pageNumber="293" value="2015-02-03">
|
||
<collectingDate id="7041097ABA6CFFF4FD2F9339DE2AC186" pageId="3" pageNumber="293" value="2015-02-03">3 February 2015</collectingDate>
|
||
</date>
|
||
. Paratype 2 (
|
||
<specimenCode id="441D7E29BA6CFFF4FE7293D8DD1CC186" box="[417,554,1932,1956]" collectionCode="RSIO" pageId="3" pageNumber="293">RSIO35221</specimenCode>
|
||
):
|
||
</materialsCitation>
|
||
<materialsCitation id="A4D3DC0FBA6CFFF4FDE893D8DA3EC04E" collectingDate="2014-12-23" collectingDateMax="2017-03-14" collectingDateMin="2014-12-23" collectionCode="SWIR" elevation="2760" latitude="-23.88" longLatPrecision="752" longitude="69.6" pageId="3" pageNumber="293" specimenCode="RSIO35217, RSIO35219, RSIO35221, RSIO35222, RSIO35290, RSIO38129" specimenCount="7" typeStatus="paratype">
|
||
male, part segments remaining,
|
||
<quantity id="D3437BB7BA6CFFF4FF2E93FFDE69C1E1" box="[253,351,1963,1987]" metricMagnitude="-3" metricUnit="m" metricValue="5.42" pageId="3" pageNumber="293" unit="mm" value="5.42">5.42 mm</quantity>
|
||
in length,
|
||
<quantity id="D3437BB7BA6CFFF4FE3393FFDD75C1E1" box="[480,579,1963,1987]" metricMagnitude="-3" metricUnit="m" metricValue="3.48" pageId="3" pageNumber="293" unit="mm" value="3.48">3.48 mm</quantity>
|
||
in width, Longqi vent field (
|
||
<geoCoordinate id="718FB095BA6CFFF4FC7C9261DB32C06F" box="[943,1028,1589,1613]" degrees="37.78" direction="south" orientation="latitude" pageId="3" pageNumber="293" precision="555" value="-37.78">37.78°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="718FB095BA6CFFF4FBC09261DB51C06F" box="[1043,1127,1589,1613]" degrees="49.65" direction="east" orientation="longitude" pageId="3" pageNumber="293" precision="555" value="49.65">49.65°E</geoCoordinate>
|
||
,
|
||
<quantity id="D3437BB7BA6CFFF4FBA69261DBFFC06F" box="[1141,1225,1589,1613]" metricMagnitude="3" metricUnit="m" metricValue="2.7399999999999998" pageId="3" pageNumber="293" unit="m" value="2740.0">
|
||
<elevation id="9F963161BA6CFFF4FBA69261DBFFC06F" box="[1141,1225,1589,1613]" metricMagnitude="3" metricUnit="m" metricValue="2.7399999999999998" pageId="3" pageNumber="293" unit="m" value="2740.0">2740 m</elevation>
|
||
</quantity>
|
||
deep),
|
||
<collectionCode id="72AA4E97BA6CFFF4FACD9261DA54C06F" box="[1310,1378,1589,1613]" pageId="3" pageNumber="293">SWIR</collectionCode>
|
||
, Dive90,
|
||
<date id="6005F092BA6CFFF4FCEA9201DCD6C04E" box="[825,992,1620,1645]" pageId="3" pageNumber="293" value="2015-01-03">
|
||
<collectingDate id="7041097ABA6CFFF4FCEA9201DCD6C04E" box="[825,992,1620,1645]" pageId="3" pageNumber="293" value="2015-01-03">3 January 2015</collectingDate>
|
||
</date>
|
||
. Paratype 3 (
|
||
<specimenCode id="441D7E29BA6CFFF4FBA19200DBCAC04E" box="[1138,1276,1620,1644]" collectionCode="RSIO" pageId="3" pageNumber="293">RSIO35222</specimenCode>
|
||
):
|
||
</materialsCitation>
|
||
<materialsCitation id="A4D3DC0FBA6CFFF4FAC29201DB60C12B" collectingDate="2014-12-23" collectingDateMax="2017-03-14" collectingDateMin="2014-12-23" collectionCode="SWIR" elevation="2760" latitude="-23.88" longLatPrecision="752" longitude="69.6" pageId="3" pageNumber="293" specimenCode="RSIO35217, RSIO35219, RSIO35221, RSIO35222, RSIO35290, RSIO38129" specimenCount="7" typeStatus="paratype">
|
||
female, part segments remaining,
|
||
<quantity id="D3437BB7BA6CFFF4FC249220DB62C0A9" box="[1015,1108,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="3.18" pageId="3" pageNumber="293" unit="mm" value="3.18">3.18 mm</quantity>
|
||
in length,
|
||
<quantity id="D3437BB7BA6CFFF4FB169220DA14C0A9" box="[1221,1314,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="3.43" pageId="3" pageNumber="293" unit="mm" value="3.43">3.43 mm</quantity>
|
||
in width, same collecting data as paratype 2. Paratype 4 (
|
||
<specimenCode id="441D7E29BA6CFFF4FB2492C7DAB4C089" box="[1271,1410,1683,1707]" collectionCode="RSIO" pageId="3" pageNumber="293">RSIO35290</specimenCode>
|
||
):
|
||
<quantity id="D3437BB7BA6CFFF4FA4692C7DC54C0E8" metricMagnitude="-3" metricUnit="m" metricValue="4.91" pageId="3" pageNumber="293" unit="mm" value="4.91">4.91 mm</quantity>
|
||
in length,
|
||
<quantity id="D3437BB7BA6CFFF4FC0492E6DB03C0E8" box="[983,1077,1714,1738]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="3" pageNumber="293" unit="mm" value="2.55">2.55 mm</quantity>
|
||
in width, Longqi vent field (
|
||
<geoCoordinate id="718FB095BA6CFFF4FABB92E7DA8BC0E9" box="[1384,1469,1715,1739]" degrees="37.78" direction="south" orientation="latitude" pageId="3" pageNumber="293" precision="555" value="-37.78">37.78°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="718FB095BA6CFFF4FCEA9286DCB8C0C8" box="[825,910,1746,1770]" degrees="49.65" direction="east" orientation="longitude" pageId="3" pageNumber="293" precision="555" value="49.65">49.65°E</geoCoordinate>
|
||
,
|
||
<quantity id="D3437BB7BA6CFFF4FC4D9286DCC3C0CB" box="[926,1013,1746,1770]" metricMagnitude="3" metricUnit="m" metricValue="2.7399999999999998" pageId="3" pageNumber="293" unit="m" value="2740.0">
|
||
<elevation id="9F963161BA6CFFF4FC4D9286DCC3C0CB" box="[926,1013,1746,1770]" metricMagnitude="3" metricUnit="m" metricValue="2.7399999999999998" pageId="3" pageNumber="293" unit="m" value="2740.0">2740 m</elevation>
|
||
</quantity>
|
||
deep),
|
||
<collectionCode id="72AA4E97BA6CFFF4FB839286DBA2C0C8" box="[1104,1172,1746,1770]" pageId="3" pageNumber="293">SWIR</collectionCode>
|
||
, Dive94,
|
||
<date id="6005F092BA6CFFF4FAD79286DA88C0C8" box="[1284,1470,1746,1770]" pageId="3" pageNumber="293" value="2015-01-11">
|
||
<collectingDate id="7041097ABA6CFFF4FAD79286DA88C0C8" box="[1284,1470,1746,1770]" pageId="3" pageNumber="293" value="2015-01-11">11 January 2015</collectingDate>
|
||
</date>
|
||
. Paratype 5 (
|
||
<specimenCode id="441D7E29BA6CFFF4FC1392A5DB7CC12B" box="[960,1098,1777,1801]" collectionCode="RSIO" pageId="3" pageNumber="293">RSIO38129</specimenCode>
|
||
):
|
||
</materialsCitation>
|
||
<materialsCitation id="A4D3DC0FBA6CFFF4FBB392A5DC8BC145" collectingDate="2014-12-23" collectingDateMax="2017-03-14" collectingDateMin="2014-12-23" collectionCode="SWIR" elevation="2760" latitude="-23.88" longLatPrecision="752" longitude="69.6" pageId="3" pageNumber="293" specimenCode="RSIO35217, RSIO35219, RSIO35221, RSIO35222, RSIO35290, RSIO38129" specimenCount="7" typeStatus="paratype">
|
||
female,
|
||
<quantity id="D3437BB7BA6CFFF4FB6692A5DA21C12B" box="[1205,1303,1777,1801]" metricMagnitude="-3" metricUnit="m" metricValue="7.58" pageId="3" pageNumber="293" unit="mm" value="7.58">7.58 mm</quantity>
|
||
in length,
|
||
<quantity id="D3437BB7BA6CFFF4FA4692A5DC54C10A" metricMagnitude="-3" metricUnit="m" metricValue="5.01" pageId="3" pageNumber="293" unit="mm" value="5.01">5.01 mm</quantity>
|
||
in width, Wocan vent field (
|
||
<geoCoordinate id="718FB095BA6CFFF4FB799344DBCFC10A" box="[1194,1273,1808,1832]" degrees="6.36" direction="north" orientation="latitude" pageId="3" pageNumber="293" precision="555" value="6.36">6.36°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="718FB095BA6CFFF4FADA9344DA6BC10A" box="[1289,1373,1808,1832]" degrees="69.60" direction="east" orientation="longitude" pageId="3" pageNumber="293" precision="555" value="69.6">69.60°E</geoCoordinate>
|
||
,
|
||
<quantity id="D3437BB7BA6CFFF4FABE9345DAF7C10A" box="[1389,1473,1809,1833]" metricMagnitude="3" metricUnit="m" metricValue="2.9699999999999998" pageId="3" pageNumber="293" unit="m" value="2970.0">
|
||
<elevation id="9F963161BA6CFFF4FABE9345DAF7C10A" box="[1389,1473,1809,1833]" metricMagnitude="3" metricUnit="m" metricValue="2.9699999999999998" pageId="3" pageNumber="293" unit="m" value="2970.0">2970 m</elevation>
|
||
</quantity>
|
||
deep), CR, Dive129,
|
||
<date id="6005F092BA6CFFF4FBFA9364DBE1C16A" box="[1065,1239,1840,1864]" pageId="3" pageNumber="293" value="2017-03-14">
|
||
<collectingDate id="7041097ABA6CFFF4FBFA9364DBE1C16A" box="[1065,1239,1840,1864]" pageId="3" pageNumber="293" value="2017-03-14">14 March 2017</collectingDate>
|
||
</date>
|
||
. Paratype 6 (NSMTPol_P-920):
|
||
</materialsCitation>
|
||
<materialsCitation id="A4D3DC0FBA6CFFF4FC12931BDBF7C1E7" collectingDate="2014-12-23" collectingDateMax="2017-03-14" collectingDateMin="2014-12-23" collectionCode="SWIR" elevation="2760" latitude="-23.88" longLatPrecision="752" longitude="69.6" pageId="3" pageNumber="293" specimenCode="RSIO35217, RSIO35219, RSIO35221, RSIO35222, RSIO35290, RSIO38129" specimenCount="7" typeStatus="paratype">
|
||
female, 25 segments,
|
||
<quantity id="D3437BB7BA6CFFF4FB4E931BDBCFC145" box="[1181,1273,1871,1895]" metricMagnitude="-3" metricUnit="m" metricValue="7.42" pageId="3" pageNumber="293" unit="mm" value="7.42">7.42 mm</quantity>
|
||
in length,
|
||
<quantity id="D3437BB7BA6CFFF4FAB5931BDAF4C145" box="[1382,1474,1871,1895]" metricMagnitude="-3" metricUnit="m" metricValue="4.66" pageId="3" pageNumber="293" unit="mm" value="4.66">4.66 mm</quantity>
|
||
in width, Edmond vent field (
|
||
<geoCoordinate id="718FB095BA6CFFF4FBBF933BDBF7C1A5" box="[1132,1217,1903,1927]" degrees="23.88" direction="south" orientation="latitude" pageId="3" pageNumber="293" precision="555" value="-23.88">23.88°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="718FB095BA6CFFF4FB1F933ADA16C1A4" box="[1228,1312,1902,1926]" degrees="69.60" direction="east" orientation="longitude" pageId="3" pageNumber="293" precision="555" value="69.6">69.60°E</geoCoordinate>
|
||
,
|
||
<quantity id="D3437BB7BA6CFFF4FAF9933BDA4CC1A4" box="[1322,1402,1903,1927]" metricMagnitude="3" metricUnit="m" metricValue="3.2769999999999997" pageId="3" pageNumber="293" unit="m" value="3277.0">
|
||
<elevation id="9F963161BA6CFFF4FAF9933BDA4CC1A4" box="[1322,1402,1903,1927]" metricMagnitude="3" metricUnit="m" metricValue="3.2769999999999997" pageId="3" pageNumber="293" unit="m" value="3277.0">3277 m</elevation>
|
||
</quantity>
|
||
deep), CIR, Dive1451,
|
||
<date id="6005F092BA6CFFF4FC3E93DADB84C184" box="[1005,1202,1934,1958]" pageId="3" pageNumber="293" value="2016-02-15">
|
||
<collectingDate id="7041097ABA6CFFF4FC3E93DADB84C184" box="[1005,1202,1934,1958]" pageId="3" pageNumber="293" value="2016-02-15">15 February 2016</collectingDate>
|
||
</date>
|
||
. All paratypes were fixed and stored in 95–100% (v/v) ethanol
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1404D652BA6BFFF3FFA294C5DEE0C767" blockId="4.[113,763,144,1421]" pageId="4" pageNumber="294">
|
||
<emphasis id="26CF0A40BA6BFFF3FFA294C5DFDEC68A" box="[113,232,145,168]" italics="true" pageId="4" pageNumber="294">Description:</emphasis>
|
||
Body short, tapering anteriorly and posteriorly (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FFAF94FBDFD5C6E5" box="[124,227,175,199]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2A, B</figureCitation>
|
||
). Elytra 11 pairs, subreniform (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FDE194FBDD48C6E5" box="[562,638,175,199]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2C</figureCitation>
|
||
), iridescent when fresh and pale white in ethanol, attached on prominent elytrophores on segments 2, 4, 5, 7, alternating to 21 (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FD7394BADDDFC724" box="[672,745,238,262]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2A</figureCitation>
|
||
), with micropapillae on posterior and lateral border, which can only be seen under SEM (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FEA99579DEF3C767" box="[378,453,301,325]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1404D652BA6BFFF3FF5E9518DE65C55B" blockId="4.[113,763,144,1421]" pageId="4" pageNumber="294">
|
||
Prostomium oval, deeply bilobed (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FE2C9518DD7FC746" box="[511,585,332,356]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2E</figureCitation>
|
||
). Anterior lobes subtriangle, with delicate frontal filaments (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FD95953FDDA7C7A1" box="[582,657,363,387]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2E</figureCitation>
|
||
). Median antenna with short, cylindrical ceratophore inserted into anterior notch; style subulate, with a long, terminal filament (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FFAF959DDFF1C7C3" box="[124,199,457,481]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2E</figureCitation>
|
||
). Palps stout, cylindrical, and tapering with terminal filaments. Eyes lacking (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FEA495BDDEE9C422" box="[375,479,489,513]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2E, F</figureCitation>
|
||
). First segment achaetous ventrally fused to prostomium, with two pairs of short, cylindrical tentaculophores; tentacular cirri conical, long, and stout with slender filaments (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FE7D9612DD23C47C" box="[430,533,582,607]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2E, F</figureCitation>
|
||
). Segment 2 with biramous parapodia bearing the first pair of elytra (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FD499632DDD1C45C" box="[666,743,614,638]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2E</figureCitation>
|
||
); ventral cirri morphologically similar to tentacular cirri and attached basally to neuropodia, much longer than ventral cirri on following segments (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FE509690DEE5C4FE" box="[387,467,708,732]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2F</figureCitation>
|
||
). Mouth ventrally located between segments 1 and 2 (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FE7396B7DEDDC4D9" box="[416,491,739,763]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2F</figureCitation>
|
||
). Pharynx when everted, surrounded by seven pairs of papillae on dorsal and ventral sides, as well as one pair of rounded papillae on lateral sides (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FFAF9715DFFDC57B" box="[124,203,833,857]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2G</figureCitation>
|
||
). Two pairs of curved jaws without teeth at opening of pharynx (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FF279734DE74C55B" box="[244,322,864,889]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1404D652BA6BFFF3FF5E97D4DEA7C2F3" blockId="4.[113,763,144,1421]" pageId="4" pageNumber="294">
|
||
Biramous parapodia with neuropodium about as long as notopodium (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FED297CBDE57C595" box="[257,353,927,951]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2K, L</figureCitation>
|
||
). Notopodium with rows of notochaetae (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FFAF97EADFD3C5F4" box="[124,229,958,983]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2H, L</figureCitation>
|
||
). Neuropodium with triangle projecting prechaetal lobe (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FF62978ADFCCC5D4" box="[177,250,990,1014]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2L</figureCitation>
|
||
) and short, rounded postchaetal lobe (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FD4E978ADDDCC5D4" box="[669,746,990,1014]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2K</figureCitation>
|
||
). Tapering dorsal cirri with long, cylindrical cirrophores attached on posterior dorsal side of notopodia on non-elytrigerous segments (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FF359068DE6DC271" box="[230,347,1084,1108]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2H, K</figureCitation>
|
||
). Rows of notochaetae stout, tapering to pointed tips with serrations in distal part (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FDA1900FDDDCC251" box="[626,746,1115,1139]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2L, M</figureCitation>
|
||
). Neurochaetae very numerous, forming fan-shaped bundle (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FD07902EDF9FC293" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2K, L</figureCitation>
|
||
) with two rows of widely-spaced acicular spines to hooked tips on both sides (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FEE790EDDEB6C2F3" box="[308,384,1209,1233]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1404D652BA6BFFF3FF5E908CDD57C36C" blockId="4.[113,763,144,1421]" pageId="4" pageNumber="294">
|
||
Pygidium small, rounded, anal cirri lost during collection (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FFAF90ACDFE8C32D" box="[124,222,1272,1296]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2A, B</figureCitation>
|
||
). Ten pairs of inconspicuous rounded papillae in segments
|
||
<quantity id="D3437BB7BA6BFFF3FF669143DE39C30D" box="[181,271,1303,1327]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="5.08" metricValueMin="2.794" pageId="4" pageNumber="294" unit="in" value="15.5" valueMax="20.0" valueMin="11.0">11–20 in</quantity>
|
||
<typeStatus id="CB0068F0BA6BFFF3FEC79143DE46C30D" box="[276,368,1303,1327]" pageId="4" pageNumber="294" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="8C80CAD7BA6BFFF3FE539143DEF6C30D" box="[384,448,1303,1327]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2I</figureCitation>
|
||
), which are difficult to observe on other female
|
||
<typeStatus id="CB0068F0BA6BFFF3FEC49163DE4CC36D" box="[279,378,1335,1359]" pageId="4" pageNumber="294" type="paratype">paratypes</typeStatus>
|
||
due to their small size.
|
||
</paragraph>
|
||
<paragraph id="1404D652BA6BFFF3FF5E9102DDC7C3AF" blockId="4.[113,763,144,1421]" pageId="4" pageNumber="294">
|
||
On male, two pairs of long, ventral papillae on segments 11 and 12, which extend to the following three segments (
|
||
<figureCitation id="8C80CAD7BA6BFFF3FD4C9121DDD6C3AF" box="[671,736,1397,1421]" captionStart="Figure 2" captionStartId="3.[129,194,1349,1373]" captionTargetBox="[129,1473,142,1321]" captionTargetId="figure-227@3.[129,1473,142,1321]" captionTargetPageId="3" captionText="Figure 2. Levensteiniella pettiboneae sp. nov. Holotype (RSIO35217, female) in dorsal (A) and ventral views (B); C, D, left elytron from segment 7 of the holotype RSIO35217 in dorsal view; head and anterior segments of paratype RSIO35219 in dorsal (E) view and ventral (F) view; anterior (G) view of pharynx of paratype RSIO35290; segments of paratype RSIO35219 in dorsal (H) view; ventral (I) view of female ventral papillae on segments 12–16 of holotype RSIO35217; ventral (J) view of male ventral papillae on segments 11 and 12 of paratype RSIO35221; posterior view (upper position) and anterior view (lower position) of right parapodia from holotype RSIO35217 segment 6 (K); anterior (L) view of segment 6 on holotype RSIO35217; M, notochaetae of segment 6 on holotype RSIO35217; N, neurochaetae of segment 6 on holotype RSIO35217.Scale bars: A, B = 5mm, C, E, F, H–L = 0.5 mm, D, M = 100 μm, G = 0.1 mm, N = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.13220050" httpUri="https://zenodo.org/record/13220050/files/figure.png" pageId="4" pageNumber="294">Fig. 2J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1404D652BA6BFFF3FFA291FADFEBC027" blockId="4.[113,763,1454,1541]" pageId="4" pageNumber="294">
|
||
<emphasis id="26CF0A40BA6BFFF3FFA291FADFD7C3E4" box="[113,225,1454,1478]" italics="true" pageId="4" pageNumber="294">Etymology:</emphasis>
|
||
The species is named in honour of the late Marian H. Pettibone, in recognition of her great contribution to polynoid taxonomy.
|
||
</paragraph>
|
||
<paragraph id="1404D652BA6BFFF3FFA29278DDC7C046" blockId="4.[113,763,1580,1636]" pageId="4" pageNumber="294">
|
||
<emphasis id="26CF0A40BA6BFFF3FFA29278DFD9C066" box="[113,239,1580,1604]" italics="true" pageId="4" pageNumber="294">Distribution:</emphasis>
|
||
Tiancheng vent field and Longqi vent field of SWIR, Edmond vent field of CIR, and Wocan vent field of CR.
|
||
</paragraph>
|
||
<paragraph id="1404D652BA6BFFF3FFA292DFDC48C780" blockId="4.[113,763,1675,1981]" lastBlockId="4.[810,1459,143,418]" pageId="4" pageNumber="294">
|
||
<emphasis id="26CF0A40BA6BFFF3FFA292DFDFE3C081" box="[113,213,1675,1699]" italics="true" pageId="4" pageNumber="294">Remarks:</emphasis>
|
||
Elytralandventralpapillaeareimportantdiagnostic characteristics within
|
||
<taxonomicName id="D3BBADD1BA6BFFF3FEBF92FEDD33C0E0" authorityName="Pettibone" authorityYear="1985" box="[364,517,1706,1730]" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="294" phylum="Annelida" rank="genus">
|
||
<emphasis id="26CF0A40BA6BFFF3FEBF92FEDD33C0E0" box="[364,517,1706,1730]" italics="true" pageId="4" pageNumber="294">Levensteiniella</emphasis>
|
||
</taxonomicName>
|
||
. The combination of these two features can separate
|
||
<taxonomicName id="D3BBADD1BA6BFFF3FE33929EDDCCC0C0" authorityName="Han & Zhou & Chen & Wang" authorityYear="2024" box="[480,762,1738,1762]" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="294" phylum="Annelida" rank="species" species="pettiboneae" status="sp. nov.">
|
||
<emphasis id="26CF0A40BA6BFFF3FE33929EDDCCC0C0" box="[480,762,1738,1762]" italics="true" pageId="4" pageNumber="294">Levensteiniella pettiboneae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3DFCB73BBA6BFFF3FFA292BDDFFCC122" box="[113,202,1769,1792]" pageId="4" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from most other congeners (e.g.
|
||
<taxonomicName id="D3BBADD1BA6BFFF3FDB192BDDEFFC102" authority="Wu and Xu, 2018" authorityName="Wu and Xu" authorityYear="2018" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="294" phylum="Annelida" rank="species" species="manusensis">
|
||
<emphasis id="26CF0A40BA6BFFF3FDB192BDDFD0C102" italics="true" pageId="4" pageNumber="294">Levensteiniella manusensis</emphasis>
|
||
<bibRefCitation id="702AABA3BA6BFFF3FF26935DDEFFC102" author="Wu X & Xu K" box="[245,457,1800,1824]" pageId="4" pageNumber="294" pagination="102 - 10" refId="ref13367" refString="Wu X, Xu K. Levensteiniella manusensis sp. nov., a new polychaete species (Annelida: Polynoidae) from deep-sea hydrothermal vents in the Manus Back-Arc Basin, Western Pacific. Zootaxa 2018; 4388: 102 - 10." type="journal article" year="2018">Wu and Xu, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D3BBADD1BA6BFFF3FDDE935CDEEAC162" authority="Hourdez and Desbruyeres, 2000" authorityName="Hourdez and Desbruyeres" authorityYear="2000" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="294" phylum="Annelida" rank="species" species="plicata">
|
||
<emphasis id="26CF0A40BA6BFFF3FDDE935CDDCDC102" box="[525,763,1800,1824]" italics="true" pageId="4" pageNumber="294">Levensteiniella plicata</emphasis>
|
||
<bibRefCitation id="702AABA3BA6BFFF3FFA2937CDEEAC162" author="Hourdez S & Desbruyeres D" box="[113,476,1832,1856]" pageId="4" pageNumber="294" pagination="97 - 102" refId="ref11682" refString="Hourdez S, Desbruyeres D. A new species of scale-worm (Polychaeta: Polynoidae), Levensteiniella plicata sp. nov., from the East Pacific Rise. Cahiers de Biologie Marine 2000; 41: 97 - 102." type="journal article" year="2000">Hourdez and Desbruyères, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
) except for
|
||
<taxonomicName id="D3BBADD1BA6BFFF3FDB1937CDEA8C17D" authority="(Pettibone 1985 a)" baseAuthorityName="Pettibone" baseAuthorityYear="1985" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="294" phylum="Annelida" rank="species" species="kincaidi">
|
||
<emphasis id="26CF0A40BA6BFFF3FDB1937CDFF2C17D" italics="true" pageId="4" pageNumber="294">Levensteiniella kincaidi</emphasis>
|
||
(
|
||
<bibRefCitation id="702AABA3BA6BFFF3FF089313DEA5C17D" author="Pettibone MH" box="[219,403,1863,1887]" pageId="4" pageNumber="294" pagination="740 - 57" refId="ref12446" refString="Pettibone MH. New genera and species of deep-sea Macellicephalinae and Harmothoinae (Polychaeta: Polynoidae) from the hydrothermal rift areas off the Galapagos and western Mexico at 21 ° N and from Santa Catalina Channel. Proceedings of the Biological Society of Washington 1985 a; 98: 740 - 57." type="journal article" year="1985">Pettibone 1985a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which resemble the new species in elytra-bearing micropapillae in posterior and lateral parts, and two pairs of long, ventral papillae on the male (
|
||
<bibRefCitation id="702AABA3BA6BFFF3FFAE93F1DE02C19F" author="Pettibone MH" box="[125,308,1957,1981]" pageId="4" pageNumber="294" pagination="740 - 57" refId="ref12446" refString="Pettibone MH. New genera and species of deep-sea Macellicephalinae and Harmothoinae (Polychaeta: Polynoidae) from the hydrothermal rift areas off the Galapagos and western Mexico at 21 ° N and from Santa Catalina Channel. Proceedings of the Biological Society of Washington 1985 a; 98: 740 - 57." type="journal article" year="1985">Pettibone 1985a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="702AABA3BA6BFFF3FE9593F1DE4BC19F" author="Pettibone MH" box="[326,381,1957,1981]" pageId="4" pageNumber="294" pagination="137 - 53" refId="ref12581" refString="Pettibone MH. New species of scale-worms (Polychaeta: Polynoidae) from the hydrothermal rift-area of the Mariana Back-arc Basin in the Western Central Pacific. Proceedings of the Biological Society of Washington 1989; 102: 137 - 53." type="journal article" year="1989">1989</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="702AABA3BA6BFFF3FE5E93F1DEF2C19F" author="Pettibone MH" box="[397,452,1957,1981]" pageId="4" pageNumber="294" pagination="825 - 38" refId="ref12623" refString="Pettibone MH. New species and new records of scaled polychaetes (Polychaeta: Polynoidae) fromtheAxialSeamountCalderaoftheJuan de Fuca ridge in the northeast Pacific Ocean off northern California. Proceedings of the Biological Society of Washington 1990; 103: 825 - 38." type="journal article" year="1990">1990</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="702AABA3BA6BFFF3FE0693F1DC57C68A" author="Hourdez S & Desbruyeres D" box="[469,865,144,1981]" pageId="4" pageNumber="294" pagination="97 - 102" refId="ref11682" refString="Hourdez S, Desbruyeres D. A new species of scale-worm (Polychaeta: Polynoidae), Levensteiniella plicata sp. nov., from the East Pacific Rise. Cahiers de Biologie Marine 2000; 41: 97 - 102." type="journal article" year="2000">Hourdez and Desbruyères 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="702AABA3BA6BFFF3FCBE94C4DC92C68A" author="Hourdez S & Desbruyeres D" box="[877,932,144,168]" pageId="4" pageNumber="294" pagination="13 - 21" refId="ref11724" refString="Hourdez S, Desbruyeres D. A new species of scale-worm (Polychaeta: Polynoidae), Levensteiniella iris sp nov., from the Rainbow and Lucky Strike vent fields (Mid-Atlantic Ridge). CBM-Cahiers de Biologie Marine 2003; 44: 13 - 21." type="journal article" year="2003">2003</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="702AABA3BA6BFFF3FC6394C4DB55C68A" author="Wu X & Xu K" box="[944,1123,143,168]" pageId="4" pageNumber="294" pagination="102 - 10" refId="ref13367" refString="Wu X, Xu K. Levensteiniella manusensis sp. nov., a new polychaete species (Annelida: Polynoidae) from deep-sea hydrothermal vents in the Manus Back-Arc Basin, Western Pacific. Zootaxa 2018; 4388: 102 - 10." type="journal article" year="2018">Wu and Xu 2018</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="702AABA3BA6BFFF3FBBC94DBDA02C685" author="Zhang Y & Chen C & Qiu JW" box="[1135,1332,143,167]" pageId="4" pageNumber="294" refId="ref13559" refString="Zhang Y, Chen C, Qiu JW. Sexually dimorphic scale worms (Annelida: Polynoidae) from hydrothermal vents in the Okinawa Trough: two new species and two new sex morphs. Frontiers in Marine Science 2018 b; 5: 112." type="journal volume" year="2018">
|
||
Zhang
|
||
<emphasis id="26CF0A40BA6BFFF3FB6994C4DBDAC685" box="[1210,1260,143,167]" italics="true" pageId="4" pageNumber="294">et al.</emphasis>
|
||
2018b
|
||
</bibRefCitation>
|
||
). However, the two species differ in the number of small, rounded papillae on the female (10 pairs in
|
||
<taxonomicName id="D3BBADD1BA6BFFF3FB48949ADA85C6C4" authorityName="Han & Zhou & Chen & Wang" authorityYear="2024" box="[1179,1459,206,230]" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="294" phylum="Annelida" rank="species" species="pettiboneae" status="sp. nov.">
|
||
<emphasis id="26CF0A40BA6BFFF3FB48949ADA85C6C4" box="[1179,1459,206,230]" italics="true" pageId="4" pageNumber="294">Levensteiniella pettiboneae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3DFCB73BBA6BFFF3FCF994BADC4AC727" box="[810,892,238,261]" pageId="4" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
|
||
vs. nine pairs in
|
||
<taxonomicName id="D3BBADD1BA6BFFF3FBE594B9DA1FC727" baseAuthorityName="Pettibone" baseAuthorityYear="1985" box="[1078,1321,237,261]" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="294" phylum="Annelida" rank="species" species="kincaidi">
|
||
<emphasis id="26CF0A40BA6BFFF3FBE594B9DA1FC727" box="[1078,1321,237,261]" italics="true" pageId="4" pageNumber="294">Levensteiniella kincaidi</emphasis>
|
||
</taxonomicName>
|
||
). Moreover, the number of pharyngeal papillae vary in
|
||
<taxonomicName id="D3BBADD1BA6BFFF3FAC99559DC4BC766" baseAuthorityName="Pettibone" baseAuthorityYear="1985" class="Polychaeta" family="Polynoidae" genus="Levensteiniella" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="294" phylum="Annelida" rank="species" species="kincaidi">
|
||
<emphasis id="26CF0A40BA6BFFF3FAC99559DC4BC766" italics="true" pageId="4" pageNumber="294">Levensteiniella kincaidi</emphasis>
|
||
</taxonomicName>
|
||
(13–16, mostly seven pairs), while there is an additional pair of lateral papillae in the new species in addition to the seven pairs on the dorsal and ventral sides (
|
||
<bibRefCitation id="702AABA3BA6BFFF3FA9A953FDC5BC780" author="Pettibone MH" pageId="4" pageNumber="294" pagination="740 - 57" refId="ref12446" refString="Pettibone MH. New genera and species of deep-sea Macellicephalinae and Harmothoinae (Polychaeta: Polynoidae) from the hydrothermal rift areas off the Galapagos and western Mexico at 21 ° N and from Santa Catalina Channel. Proceedings of the Biological Society of Washington 1985 a; 98: 740 - 57." type="journal article" year="1985">Pettibone 1985a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |