289 lines
36 KiB
XML
289 lines
36 KiB
XML
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<mods:title id="D051264598DE76BDEEF1F149EAA19B36">Revision of Madagascan Cleridae (Coleoptera: Cleroidea). Part I. - The genus Biflabelotillus Pic, 1949 (Tillinae)</mods:title>
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<mods:title id="B4566AFF3EDD9289AA5CFB40C82C0356">Zootaxa</mods:title>
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<paragraph id="8B84E50FFFF9FFFD13E9FF73FDA7AAE3" blockId="5.[151,618,151,214]" box="[151,618,151,178]" pageId="5" pageNumber="389">
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<heading id="D0CC5263FFF9FFFD13E9FF73FDA7AAE3" bold="true" box="[151,618,151,178]" fontSize="11" level="1" pageId="5" pageNumber="389" reason="1">
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<taxonomicName id="4C3B9E8CFFF9FFFD13E9FF73FDA7AAE3" authority="Menier, 1979" authorityName="Menier" authorityYear="1979" box="[151,618,151,178]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
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<emphasis id="B94F391DFFF9FFFD13E9FF73FDA7AAE3" bold="true" box="[151,618,151,178]" pageId="5" pageNumber="389">
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<emphasis id="B94F391DFFF9FFFD13E9FF73FE0CAAE3" bold="true" box="[151,449,151,178]" italics="true" pageId="5" pageNumber="389">Biflabelotillus peyrierasi</emphasis>
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<bibRefCitation id="EFAA98FEFFF9FFFD12B6FF73FDA7AAE3" author="Menier, J. J." box="[456,618,151,177]" pageId="5" pageNumber="389" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier, 1979</bibRefCitation>
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</emphasis>
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</taxonomicName>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C321B684FFF9FFFD13E9FF5FFE5CAA87" box="[151,401,187,214]" pageId="5" pageNumber="389" type="description">
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<paragraph id="8B84E50FFFF9FFFD13E9FF5FFE5CAA87" blockId="5.[151,618,151,214]" box="[151,401,187,214]" pageId="5" pageNumber="389">
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<figureCitation id="1300F98AFFF9FFFD13E9FF5FFEF2AA87" box="[151,319,187,214]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Figs 2, 5, 8, 11</figureCitation>
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, 21–32
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C321B684FFF9FFFD13E9FEE4FCA2AB4A" box="[151,879,255,280]" pageId="5" pageNumber="389" type="reference_group">
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<paragraph id="8B84E50FFFF9FFFD13E9FEE4FCA2AB4A" blockId="5.[151,879,255,280]" box="[151,879,255,280]" pageId="5" pageNumber="389">
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<treatmentCitationGroup id="AB2BC221FFF9FFFD13E9FEE4FCA2AB4A" box="[151,879,255,280]" pageId="5" pageNumber="389">
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<treatmentCitation id="0A9AC31EFFF9FFFD13E9FEE4FD46AB45" author="Menier, J. J." box="[151,651,255,280]" page="75" pageId="5" pageNumber="389" year="1979">
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<taxonomicName id="4C3B9E8CFFF9FFFD13E9FEE4FD46AB45" authority="Menier, 1979: 75" authorityName="Menier" authorityPageNumber="75" authorityYear="1979" box="[151,651,255,280]" class="Insecta" family="Cleridae" genus="Biflabellotillus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
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||
<emphasis id="B94F391DFFF9FFFD13E9FEE4FEE3AB4A" box="[151,302,256,280]" italics="true" pageId="5" pageNumber="389">Biflabellotillus</emphasis>
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||
[sic!]
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<emphasis id="B94F391DFFF9FFFD120EFEE4FE1AAB45" box="[368,471,256,279]" italics="true" pageId="5" pageNumber="389">peyrierasi</emphasis>
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<bibRefCitation id="EFAA98FEFFF9FFFD12A3FEE4FD46AB45" author="Menier, J. J." box="[477,651,255,280]" pageId="5" pageNumber="389" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier, 1979: 75</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
(original description).
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C321B684FFF9FFFD13E9FEA2FCD9A846" pageId="5" pageNumber="389" type="materials_examined">
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<paragraph id="8B84E50FFFF9FFFD13E9FEA2FC1DAB32" blockId="5.[151,1436,325,1288]" box="[151,976,325,352]" pageId="5" pageNumber="389">
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<materialsCitation id="3B53EF52FFF9FFFD13E9FEA2FC00AB32" box="[151,973,325,352]" county="Mananara Nord District" location="Antanambe" pageId="5" pageNumber="389" specimenCount="1" stateProvince="Analanjirofo Region">
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||
<emphasis id="B94F391DFFF9FFFD13E9FEA2FEFBAB0D" bold="true" box="[151,310,325,352]" pageId="5" pageNumber="389">Type locality.</emphasis>
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||
<location id="8EE4B3D4FFF9FFFD1245FEA2FE72AB32" LSID="urn:lsid:plazi:treatment:03925419FFF9FFFD13E9FF73FA40AF5A:8EE4B3D4FFF9FFFD1245FEA2FE72AB32" box="[315,447,326,352]" county="Mananara Nord District" municipality="Mananara Nord District" name="Antanambe" pageId="5" pageNumber="389" stateProvince="Analanjirofo Region">Antanambe</location>
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,
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||
<collectingCounty id="62E59D83FFF9FFFD12B4FEA2FD15AB32" box="[458,728,326,352]" pageId="5" pageNumber="389">Mananara Nord District</collectingCounty>
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,
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||
<collectingRegion id="49FF2BEDFFF9FFFD119CFEA2FC00AB32" box="[738,973,326,352]" pageId="5" pageNumber="389">Analanjirofo Region</collectingRegion>
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8B84E50FFFF9FFFD13B9FE8EFE69AB9E" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
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||
<emphasis id="B94F391DFFF9FFFD13B9FE8EFE16ABD1" bold="true" box="[199,475,361,388]" pageId="5" pageNumber="389">Type material studied.</emphasis>
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||
<materialsCitation id="3B53EF52FFF9FFFD1298FE8EFE21ABFA" collectionCode="MNHN" country="Madagascar" county="Mananara Nord District" location="Madagascar" pageId="5" pageNumber="389" specimenCount="1" specimenCount-male="1" stateProvince="Analanjirofo Region" typeStatus="holotype">
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<typeStatus id="54805BADFFF9FFFD1298FE8EFD91ABD6" box="[486,604,362,388]" pageId="5" pageNumber="389" type="holotype">HOLOTYPE</typeStatus>
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,
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||
<specimenCount id="9D3D2E86FFF9FFFD1110FE8EFD4EABD6" box="[622,643,362,388]" count="1" pageId="5" pageNumber="389" type="male">♂</specimenCount>
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(
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<collectionCode id="ED2A7DCAFFF9FFFD11EBFE8EFD3FABD6" box="[661,754,362,388]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="389">MNHN</collectionCode>
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): [white, printed]
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||
<collectingCountry id="F32CA59FFFF9FFFD10BBFE8EFB81ABD6" box="[965,1100,362,388]" name="Madagascar" pageId="5" pageNumber="389">Madagascar</collectingCountry>
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Est / distr. Mananara – N. / Antanambe // [red, printed] ♂
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</materialsCitation>
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||
<typeStatus id="54805BADFFF9FFFD128BFE6AFD44ABFA" box="[501,649,398,424]" pageId="5" pageNumber="389" type="holotype">HOLOTYPE</typeStatus>
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||
// [white, handwritten]
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<taxonomicName id="4C3B9E8CFFF9FFFD10E6FE6AFB62ABFA" authorityName="Menier" authorityYear="1979" box="[920,1199,398,424]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">Biflabelotillus peyrierasi</taxonomicName>
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<specimenCount id="9D3D2E86FFF9FFFD17C6FE6AFB00ABFA" box="[1208,1229,398,424]" count="1" pageId="5" pageNumber="389" type="male">♂</specimenCount>
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||
/ n.sp.
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||
<typeStatus id="54805BADFFF9FFFD165AFE6AFA41ABFA" box="[1316,1420,398,424]" pageId="5" pageNumber="389" type="holotype">Holotype</typeStatus>
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/ J.J. MENIER det. 1979.
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</paragraph>
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||
<paragraph id="8B84E50FFFF9FFFD13B9FE31FCD9A846" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
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||
<emphasis id="B94F391DFFF9FFFD13B9FE31FDF0ABBD" bold="true" box="[199,573,469,495]" pageId="5" pageNumber="389">Additional material examined.</emphasis>
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||
(
|
||
<specimenCount id="9D3D2E86FFF9FFFD112DFE32FD4FABA2" box="[595,642,470,496]" count="1" pageId="5" pageNumber="389" type="male">1 ♂</specimenCount>
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||
,
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||
<specimenCount id="9D3D2E86FFF9FFFD11ECFE32FD0CABA2" box="[658,705,470,496]" count="1" pageId="5" pageNumber="389" type="female">1 ♀</specimenCount>
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||
IJC):
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||
<materialsCitation id="3B53EF52FFF9FFFD106EFE32FCD9A846" collectingDate="1995-11-27" collectorName="Ivo Jenis" country="Madagascar" county="Moramanga" location="East" municipality="Moramanga District" pageId="5" pageNumber="389" specimenCount="1">
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||
<location id="8EE4B3D4FFF9FFFD106EFE32FC8DABA2" LSID="urn:lsid:plazi:treatment:03925419FFF9FFFD13E9FF73FA40AF5A:8EE4B3D4FFF9FFFD106EFE32FC8DABA2" box="[784,832,470,496]" country="Madagascar" county="Moramanga" municipality="Moramanga District" name="East" pageId="5" pageNumber="389">East</location>
|
||
<collectingCountry id="F32CA59FFFF9FFFD1035FE32FC19ABA2" box="[843,980,470,496]" name="Madagascar" pageId="5" pageNumber="389">Madagascar</collectingCountry>
|
||
,
|
||
<location id="8EE4B3D4FFF9FFFD109CFE32FACBABA2" LSID="urn:lsid:plazi:treatment:03925419FFF9FFFD13E9FF73FA40AF5A:8EE4B3D4FFF9FFFD109CFE32FACBABA2" box="[994,1286,470,496]" country="Madagascar" county="Moramanga" municipality="Moramanga District" name="Alaotra-Mangoro Region" pageId="5" pageNumber="389">Alaotra-Mangoro Region</location>
|
||
,
|
||
<collectingMunicipality id="6BE07F75FFF9FFFD166BFE32FF23A846" pageId="5" pageNumber="389">Moramanga District</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62E59D83FFF9FFFD1387FE1EFE4DA846" box="[249,384,506,532]" pageId="5" pageNumber="389">Moramanga</collectingCounty>
|
||
env., 25.–
|
||
<date id="FF85C3CFFFF9FFFD128EFE1EFDB9A847" box="[496,628,506,533]" pageId="5" pageNumber="389" value="1995-11-27">
|
||
<collectingDate id="EFC13A27FFF9FFFD128EFE1EFDB9A847" box="[496,628,506,533]" pageId="5" pageNumber="389" value="1995-11-27">27.11.1995</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26CE80D9FFF9FFFD1101FE1EFD2EA846" box="[639,739,506,532]" pageId="5" pageNumber="389">Ivo Jeniš</collectorName>
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||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321B684FFF9FFFD13B9FDFAFBEBA86A" box="[199,1062,542,568]" pageId="5" pageNumber="389" type="discussion">
|
||
<paragraph id="8B84E50FFFF9FFFD13B9FDFAFBEBA86A" blockId="5.[151,1436,325,1288]" box="[199,1062,542,568]" pageId="5" pageNumber="389">
|
||
<emphasis id="B94F391DFFF9FFFD13B9FDFAFEF6A86A" bold="true" box="[199,315,542,568]" pageId="5" pageNumber="389">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="54805BADFFF9FFFD120AFDFAFE1BA86A" box="[372,470,542,568]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
|
||
was fixed by original designation by
|
||
<bibRefCitation id="EFAA98FEFFF9FFFD1002FDFAFBEFA86A" author="Menier, J. J." box="[892,1058,542,568]" pageId="5" pageNumber="389" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier (1979)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321B684FFF9FFFD13B9FDA5FDE3A89A" pageId="5" pageNumber="389" type="diagnosis">
|
||
<paragraph id="8B84E50FFFF9FFFD13B9FDA5FDE3A89A" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
|
||
<emphasis id="B94F391DFFF9FFFD13B9FDA5FE1DA809" bold="true" box="[199,464,577,603]" pageId="5" pageNumber="389">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD12A6FDA7FDA6A80E" authorityName="Menier" authorityYear="1979" box="[472,619,579,604]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94F391DFFF9FFFD12A6FDA7FDA6A80E" box="[472,619,579,604]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from both
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD10A1FDA7FBBEA80E" authorityName="Pic" authorityYear="1949" box="[991,1139,578,604]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B94F391DFFF9FFFD10A1FDA7FBBEA80E" box="[991,1139,578,604]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD17D3FDA7FAF5A80E" authorityName="Pavel & Purchart" authorityYear="2024" box="[1197,1336,578,604]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFF9FFFD17D3FDA7FAF5A80E" box="[1197,1336,578,604]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F391DFFF9FFFD163FFDA6FA51A80E" bold="true" box="[1345,1436,578,604]" pageId="5" pageNumber="389">
|
||
<taxonomicNameLabel id="A27C8466FFF9FFFD163FFDA6FA51A80E" box="[1345,1436,578,604]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
mainly by black parallel sided elytra and tuberculate elytral interstriae, whereas the elytra are brown in
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD1675FD83FA51A8D2" authorityName="Pic" authorityYear="1949" box="[1291,1436,614,640]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B94F391DFFF9FFFD1675FD83FA51A8D2" box="[1291,1436,614,640]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
and bicoloured in
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD1224FD6FFE2CA8F6" authorityName="Pavel & Purchart" authorityYear="2024" box="[346,481,650,676]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
|
||
<emphasis id="B94F391DFFF9FFFD1224FD6FFE2CA8F6" box="[346,481,650,676]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94F391DFFF9FFFD129BFD6EFDF1A8F6" bold="true" box="[485,572,650,676]" pageId="5" pageNumber="389">
|
||
<taxonomicNameLabel id="A27C8466FFF9FFFD129BFD6EFDF1A8F6" box="[485,572,650,676]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, subparallel, slightly widened with smooth elytral interstriae in both mentioned species (
|
||
<figureCitation id="1300F98AFFF9FFFD1389FD4AFEB5A89A" box="[247,376,686,712]" captionStart="FIGURE 13–20" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 13–20. Biflabelotillus foveolatus Pic, 1949—13–15 (lectotype), 16–20 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="5" pageNumber="389">Figs 14, 17</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFF9FFFD12FAFD4AFE01A89A" box="[388,460,686,712]" captionStart="FIGURE 21–27" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 21–27. Biflabelotillus peyrierasi Menier, 1979—21–23 (holotype), 24–27 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="5" pageNumber="389">22, 24</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFF9FFFD12A6FD4AFE3BA89A" box="[472,502,686,712]" captionStart="FIGURE 28–32" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 28–32. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="5" pageNumber="389">28</figureCitation>
|
||
,
|
||
<figureCitation id="1300F98AFFF9FFFD117CFD4AFDEFA89A" box="[514,546,686,712]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="5" pageNumber="389">34</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321B684FFF9FFFD13B9FD35FD9DA9BA" pageId="5" pageNumber="389" type="description">
|
||
<paragraph id="8B84E50FFFF9FFFD13B9FD35FAD5A8BF" blockId="5.[151,1436,325,1288]" box="[199,1304,721,749]" pageId="5" pageNumber="389">
|
||
<emphasis id="B94F391DFFF9FFFD13B9FD35FEB8A8B9" bold="true" box="[199,373,721,747]" pageId="5" pageNumber="389">Redescription.</emphasis>
|
||
Size of
|
||
<typeStatus id="54805BADFFF9FFFD12AFFD36FDFEA8BE" box="[465,563,722,748]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
|
||
8.1 x
|
||
<quantity id="4CC348EAFFF9FFFD1107FD36FD1FA8BE" box="[633,722,722,748]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="389" unit="mm" value="2.2">2.2 mm</quantity>
|
||
. Winged, black, with tarsi pale brown (
|
||
<figureCitation id="1300F98AFFF9FFFD17F6FD36FAC1A8BF" box="[1160,1292,722,749]" captionStart="FIGURE 21–27" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 21–27. Biflabelotillus peyrierasi Menier, 1979—21–23 (holotype), 24–27 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="5" pageNumber="389">Figs 21–25</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFF9FFFD13B9FD12FADBA967" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
|
||
<emphasis id="B94F391DFFF9FFFD13B9FD12FEC3A942" bold="true" box="[199,270,758,784]" pageId="5" pageNumber="389">Head.</emphasis>
|
||
Head including eyes broader than pronotum (DV) in ratio 1:0.8 (HW =
|
||
<quantity id="4CC348EAFFF9FFFD1756FD12FB4CA942" box="[1064,1153,758,784]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="389" unit="mm" value="1.7">1.7 mm</quantity>
|
||
). The space between eyes (frons) narrow (HW / FW = 2.8). Ultimate antennomere elongate-oval, 8.8x longer then broad (
|
||
<figureCitation id="1300F98AFFF9FFFD17BDFCFEFAC5A967" box="[1219,1288,794,821]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFF9FFFD13B9FCDAFB16A92E" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
|
||
<emphasis id="B94F391DFFF9FFFD13B9FCDAFE8AA90A" bold="true" box="[199,327,830,856]" pageId="5" pageNumber="389">Pronotum.</emphasis>
|
||
Longer than wide in ratio 1:1.3 (PL =
|
||
<quantity id="4CC348EAFFF9FFFD1070FCDAFCA5A90A" box="[782,872,830,856]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="389" unit="mm" value="1.8">1.8 mm</quantity>
|
||
, PW =
|
||
<quantity id="4CC348EAFFF9FFFD10BCFCDAFBD2A90A" box="[962,1055,830,856]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="389" unit="mm" value="1.4">1.4 mm</quantity>
|
||
), cylindrical, apical two thirds of pronotum parallel sided and slightly trangled in apical third, basal third strongly trangled (
|
||
<figureCitation id="1300F98AFFF9FFFD17F7FC86FB03A92E" box="[1161,1230,866,892]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFF9FFFD13B9FC62FB39A996" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
|
||
<emphasis id="B94F391DFFF9FFFD13B9FC62FED7A9F2" bold="true" box="[199,282,902,928]" pageId="5" pageNumber="389">Elytra.</emphasis>
|
||
Parallel, 2.3x longer than their width (EL =
|
||
<quantity id="4CC348EAFFF9FFFD1058FC62FC4CA9F2" box="[806,897,902,929]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="389" unit="mm" value="5.1">5.1 mm</quantity>
|
||
, EW =
|
||
<quantity id="4CC348EAFFF9FFFD10A0FC62FBF6A9F2" box="[990,1083,902,928]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="389" unit="mm" value="2.2">2.2 mm</quantity>
|
||
). Almost whole elytral surface covered with bituberculate foveae (
|
||
<figureCitation id="1300F98AFFF9FFFD115DFC4EFDBBA996" box="[547,630,938,964]" captionStart="FIGURE 21–27" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 21–27. Biflabelotillus peyrierasi Menier, 1979—21–23 (holotype), 24–27 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="5" pageNumber="389">Fig. 21</figureCitation>
|
||
). Space between foveae smaller than diameter of foveae.
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFF9FFFD13B9FC29FD9DA9BA" blockId="5.[151,1436,325,1288]" box="[199,592,973,1000]" pageId="5" pageNumber="389">
|
||
<emphasis id="B94F391DFFF9FFFD13B9FC29FE8BA9B5" bold="true" box="[199,326,973,999]" pageId="5" pageNumber="389">Scutellum.</emphasis>
|
||
Heart shaped (
|
||
<figureCitation id="1300F98AFFF9FFFD1291FC2AFD89A9BA" box="[495,580,974,1000]" captionStart="FIGURE 21–27" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 21–27. Biflabelotillus peyrierasi Menier, 1979—21–23 (holotype), 24–27 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="5" pageNumber="389">Fig. 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321B684FFF9FFFD13B9FC16FCE6AEB6" pageId="5" pageNumber="389" type="discussion">
|
||
<paragraph id="8B84E50FFFF9FFFD13B9FC16FBC8AE2A" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
|
||
<emphasis id="B94F391DFFF9FFFD13B9FC16FEFDAE5E" bold="true" box="[199,304,1010,1036]" pageId="5" pageNumber="389">Remark.</emphasis>
|
||
The species was described based on a single male. The female is unknown so far. Fortunately, we had the opportunity to examine the female of
|
||
<taxonomicName id="4C3B9E8CFFF9FFFD111BFBF3FD38AE62" authorityName="Menier" authorityYear="1979" box="[613,757,1047,1072]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
|
||
<emphasis id="B94F391DFFF9FFFD111BFBF3FD38AE62" box="[613,757,1047,1072]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
|
||
</taxonomicName>
|
||
and found out that it fully agrees with the male
|
||
<typeStatus id="54805BADFFF9FFFD1675FBF2FAA0AE62" box="[1291,1389,1046,1072]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
|
||
and Menier´s description except the following characters: antennae serrate (
|
||
<figureCitation id="1300F98AFFF9FFFD10CBFBDEFBCBAE06" box="[949,1030,1082,1108]" captionStart="FIGURE 28–32" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 28–32. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="5" pageNumber="389">Fig. 30</figureCitation>
|
||
) (biflabeloid in males), each elytron with pale-brown humeral spot (whole elytral surface black in males) (
|
||
<figureCitation id="1300F98AFFF9FFFD10DAFBBAFC34AE2A" box="[932,1017,1118,1144]" captionStart="FIGURE 28–32" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 28–32. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="5" pageNumber="389">Fig. 28</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B84E50FFFF9FFFD13B9FB65FCE6AEB6" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
|
||
<emphasis id="B94F391DFFF9FFFD13B9FB65FE6FAEC9" bold="true" box="[199,418,1153,1179]" pageId="5" pageNumber="389">Variability of size.</emphasis>
|
||
Examined male: body length
|
||
<quantity id="4CC348EAFFF9FFFD1186FB66FC9CAECE" box="[760,849,1154,1181]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="389" unit="mm" value="7.5">7.5 mm</quantity>
|
||
, HW =
|
||
<quantity id="4CC348EAFFF9FFFD10D3FB66FBD9AECE" box="[941,1044,1154,1181]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="5" pageNumber="389" unit="mm" value="1.65">1.65 mm</quantity>
|
||
, HW / FW = 2.75, PL =
|
||
<quantity id="4CC348EAFFF9FFFD164FFB66FA55AECE" box="[1329,1432,1154,1181]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="5" pageNumber="389" unit="mm" value="1.75">1.75 mm</quantity>
|
||
, PW =
|
||
<quantity id="4CC348EAFFF9FFFD13A1FB42FEFAAE92" box="[223,311,1190,1216]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="389" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, EL =
|
||
<quantity id="4CC348EAFFF9FFFD12FFFB42FE17AE92" box="[385,474,1190,1217]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="389" unit="mm" value="4.5">4.5 mm</quantity>
|
||
, EW =
|
||
<quantity id="4CC348EAFFF9FFFD1150FB42FD4AAE92" box="[558,647,1190,1216]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="389" unit="mm" value="2.1">2.1 mm</quantity>
|
||
. Examined female body length 8.0 mm, HW = 1.65, HW / FW = 1.94, PL =
|
||
<quantity id="4CC348EAFFF9FFFD13ABFB2EFEE0AEB6" box="[213,301,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="389" unit="mm" value="1.9">1.9 mm</quantity>
|
||
, PW =
|
||
<quantity id="4CC348EAFFF9FFFD1201FB2EFE1AAEB6" box="[383,471,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="389" unit="mm" value="1.4">1.4 mm</quantity>
|
||
, EL =
|
||
<quantity id="4CC348EAFFF9FFFD115CFB2EFDB7AEB6" box="[546,634,1226,1253]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="5" pageNumber="389" unit="mm" value="5.4">5.4 mm</quantity>
|
||
, EW =
|
||
<quantity id="4CC348EAFFF9FFFD11B0FB2EFCEAAEB6" box="[718,807,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="389" unit="mm" value="2.4">2.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321B684FFF9FFFD13B9FB09FA40AF5A" box="[199,1421,1261,1288]" pageId="5" pageNumber="389" type="distribution">
|
||
<paragraph id="8B84E50FFFF9FFFD13B9FB09FA40AF5A" blockId="5.[151,1436,325,1288]" box="[199,1421,1261,1288]" pageId="5" pageNumber="389">
|
||
<emphasis id="B94F391DFFF9FFFD13B9FB09FEACAF55" bold="true" box="[199,353,1261,1287]" pageId="5" pageNumber="389">Distribution.</emphasis>
|
||
Eastern
|
||
<collectingCountry id="F32CA59FFFF9FFFD12BCFB0AFD86AF5A" box="[450,587,1262,1288]" name="Madagascar" pageId="5" pageNumber="389">Madagascar</collectingCountry>
|
||
, known only from Moramanga and Maroantsetra Districts so far (
|
||
<figureCitation id="1300F98AFFF9FFFD1653FB0AFA4CAF5A" box="[1325,1409,1262,1288]" captionStart="FIGURE 38" captionStartId="9.[151,250,1118,1142]" captionTargetBox="[489,1099,181,1094]" captionTargetId="figure-26@9.[489,1099,181,1094]" captionTargetPageId="9" captionText="FIGURE 38. Distribution map of the genus Biflabelotillus. B. foveolatus—red triangle, B. peyrierasi—white circle, B. bicolorus sp. nov.—yellow rhombus." figureDoi="http://doi.org/10.5281/zenodo.13915947" httpUri="https://zenodo.org/record/13915947/files/figure.png" pageId="5" pageNumber="389">Fig. 38</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |