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data/03/92/54/03925419FFF5FFF213E9F901FA51AB56.xml
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<subSubSection id="C321B684FFF5FFF113E9F901FD5CACAC" box="[151,657,1764,1791]" pageId="9" pageNumber="393" type="nomenclature">
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@ -71,7 +73,7 @@ Key to species of
) or at most with pale-brown humeral spot (
<figureCitation id="1300F98AFFF5FFF11729F8CDFB6CAD6D" box="[1111,1185,1833,1855]" captionStart="FIGURE 28–32" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 28–32. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="9" pageNumber="393">Fig. 28</figureCitation>
). Pronotum parallel-sided (
<figureCitation id="1300F98AFFF5FFF11397F8A3FEF3AD0E" box="[233,318,1863,1885]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="9" pageNumber="393">Figs 7, 8</figureCitation>
<figureCitation id="1300F98AFFF5FFF11397F8A3FEF3AD0E" box="[233,318,1863,1885]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="9" pageNumber="393">Figs 7, 8</figureCitation>
). Space between eyes relatively broad, HW / FW &lt;4................................................. 2
</keyLead>
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@ -80,7 +82,7 @@ Key to species of
- Elytra bicoloured with anterior half black and posterior half pale brown. (
<figureCitation id="1300F98AFFF5FFF110E7F880FC2DAD2B" box="[921,992,1892,1914]" captionStart="FIGURE 33–37" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 33–37. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="9" pageNumber="393">Fig. 34</figureCitation>
). Sides of pronotum not parallel, widest before middle (
<figureCitation id="1300F98AFFF5FFF1124CF865FEA0ADC5" box="[306,365,1921,1943]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="9" pageNumber="393">Fig. 9</figureCitation>
<figureCitation id="1300F98AFFF5FFF1124CF865FEA0ADC5" box="[306,365,1921,1943]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="9" pageNumber="393">Fig. 9</figureCitation>
). Space between eyes very narrow, HW / FW&gt; 4. Bituberculate foveae lacking in apical third of elytra.........................................................................................
<emphasis id="B94F391DFFF5FFF117AFF87BFA51ADE6" bold="true" box="[1233,1436,1950,1972]" pageId="9" pageNumber="393">
<taxonomicName id="4C3B9E8CFFF5FFF117AFF87BFA84ADE6" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1233,1353,1950,1972]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">

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@ -61,7 +63,7 @@
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<subSubSection id="C321B684FFF9FFFD13E9FA9AFE5EAFCA" box="[151,403,1406,1432]" pageId="5" pageNumber="389" type="description">
<paragraph id="8B84E50FFFF9FFFD13E9FA9AFE5EAFCA" blockId="5.[151,543,1369,1432]" box="[151,403,1406,1432]" pageId="5" pageNumber="389">
<figureCitation id="1300F98AFFF9FFFD13E9FA9AFE8DAFCA" box="[151,320,1406,1432]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="5" pageNumber="389">Figs 3, 6, 9, 12</figureCitation>
<figureCitation id="1300F98AFFF9FFFD13E9FA9AFE8DAFCA" box="[151,320,1406,1432]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Figs 3, 6, 9, 12</figureCitation>
, 33–37
</paragraph>
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@ -174,13 +176,13 @@ and
<emphasis id="B94F391DFFF9FFFD1105F893FCC1ADC2" box="[635,780,1910,1936]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
</taxonomicName>
(
<figureCitation id="1300F98AFFF9FFFD1067F892FCB6ADC2" box="[793,891,1910,1936]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="5" pageNumber="389">Figs 4, 6</figureCitation>
<figureCitation id="1300F98AFFF9FFFD1067F892FCB6ADC2" box="[793,891,1910,1936]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Figs 4, 6</figureCitation>
) whereas it is elongate-oval in
<taxonomicName id="4C3B9E8CFFF9FFFD17AFF893FAACADC2" authorityName="Menier" authorityYear="1979" box="[1233,1377,1911,1936]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF9FFFD17AFF893FAACADC2" box="[1233,1377,1911,1936]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
</taxonomicName>
(
<figureCitation id="1300F98AFFF9FFFD1611F892FF6AADE7" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="5" pageNumber="389">Fig. 5</figureCitation>
<figureCitation id="1300F98AFFF9FFFD1611F892FF6AADE7" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Fig. 5</figureCitation>
). The new species most resembles
<taxonomicName id="4C3B9E8CFFF9FFFD114FF87FFD0EADE6" authorityName="Pic" authorityYear="1949" box="[561,707,1946,1972]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFF9FFFD114FF87FFD0EADE6" box="[561,707,1946,1972]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
@ -193,13 +195,13 @@ but differs in following characteristics:
<taxonomicNameLabel id="A27C8466FFF9FFFD166BF87EFAA2ADE6" box="[1301,1391,1946,1972]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
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is a smaller species with bicoloured elytra and non-parallel sided pronotum, the latter widest before middle (
<figureCitation id="1300F98AFFF9FFFD165BF85AFAA0AD8A" box="[1317,1389,1982,2008]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="5" pageNumber="389">Figs 9</figureCitation>
<figureCitation id="1300F98AFFF9FFFD165BF85AFAA0AD8A" box="[1317,1389,1982,2008]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Figs 9</figureCitation>
, 33, 34) whereas
<taxonomicName id="4C3B9E8CFFFBFFFF1257F925FE70AC88" authorityName="Pic" authorityYear="1949" box="[297,445,1728,1754]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFBFFFF1257F925FE70AC88" box="[297,445,1728,1754]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
</taxonomicName>
is a larger species with uniformly coloured elytra and parallel sided pronotum (
<figureCitation id="1300F98AFFFBFFFF1633F924FA5AAC89" box="[1357,1431,1728,1755]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="7" pageNumber="391">Figs 7</figureCitation>
<figureCitation id="1300F98AFFFBFFFF1633F924FA5AAC89" box="[1357,1431,1728,1755]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Figs 7</figureCitation>
, 14–17). Additionally, these species can be distinguished as follows: in
<taxonomicName id="4C3B9E8CFFFBFFFF10B2F901FB95ACAC" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[972,1112,1764,1790]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFBFFFF10B2F901FB95ACAC" box="[972,1112,1764,1790]" italics="true" pageId="7" pageNumber="391">B. bicolorus</emphasis>
@ -208,13 +210,13 @@ is a larger species with uniformly coloured elytra and parallel sided pronotum (
<taxonomicNameLabel id="A27C8466FFFBFFFF171FF900FB73ACAC" box="[1121,1214,1764,1790]" pageId="7" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the eyes are larger and less emarginated anteriorly, the space between eyes (vertex) very narrow with HW / FW = 4.26 (
<figureCitation id="1300F98AFFFBFFFF1666F8ECFA43AD70" box="[1304,1422,1800,1826]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="7" pageNumber="391">Figs 9, 12</figureCitation>
<figureCitation id="1300F98AFFFBFFFF1666F8ECFA43AD70" box="[1304,1422,1800,1826]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Figs 9, 12</figureCitation>
), whereas in
<taxonomicName id="4C3B9E8CFFFBFFFF1262F8C9FE7DAD14" authorityName="Pic" authorityYear="1949" box="[284,432,1836,1862]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFBFFFF1262F8C9FE7DAD14" box="[284,432,1836,1862]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
</taxonomicName>
the eyes are smaller, more emarginated anteriorly and the space between eyes is much wider with HW / FW = 2.99–3.31 (
<figureCitation id="1300F98AFFFBFFFF115DF8B4FD5AAD38" box="[547,663,1872,1899]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="7" pageNumber="391">Figs 7, 10</figureCitation>
<figureCitation id="1300F98AFFFBFFFF115DF8B4FD5AAD38" box="[547,663,1872,1899]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Figs 7, 10</figureCitation>
). The terminal antennomere of
<taxonomicName id="4C3B9E8CFFFBFFFF1087F8B5FB4FAD38" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1017,1154,1872,1898]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFBFFFF1087F8B5FB4FAD38" box="[1017,1154,1872,1898]" italics="true" pageId="7" pageNumber="391">B. bicolorus</emphasis>
@ -223,19 +225,19 @@ the eyes are smaller, more emarginated anteriorly and the space between eyes is
<taxonomicNameLabel id="A27C8466FFFBFFFF17F7F8B4FB2EAD38" box="[1161,1251,1872,1898]" pageId="7" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
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is broadened on one side (2.75x longer than broad) and somewhat resembling the shape of a securiform shape of a palpus (
<figureCitation id="1300F98AFFFBFFFF1637F890FA43ADDC" box="[1353,1422,1908,1934]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="7" pageNumber="391">Fig. 6</figureCitation>
<figureCitation id="1300F98AFFFBFFFF1637F890FA43ADDC" box="[1353,1422,1908,1934]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Fig. 6</figureCitation>
), whereas in
<taxonomicName id="4C3B9E8CFFFBFFFF126BF87DFE6BADE0" authorityName="Pic" authorityYear="1949" box="[277,422,1944,1970]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFBFFFF126BF87DFE6BADE0" box="[277,422,1944,1970]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
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it is more elongate (3.25x longer than broad) and not securiform in shape (
<figureCitation id="1300F98AFFFBFFFF179CF87CFAEAADE0" box="[1250,1319,1944,1970]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="7" pageNumber="391">Fig. 4</figureCitation>
<figureCitation id="1300F98AFFFBFFFF179CF87CFAEAADE0" box="[1250,1319,1944,1970]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Fig. 4</figureCitation>
). Shape of antennomeres III-V is different, more tooth-like widened in
<taxonomicName id="4C3B9E8CFFFBFFFF104EF859FC0CAD84" authorityName="Pic" authorityYear="1949" box="[816,961,1980,2006]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFBFFFF104EF859FC0CAD84" box="[816,961,1980,2006]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
</taxonomicName>
(
<figureCitation id="1300F98AFFFBFFFF10B1F858FBFCAD84" box="[975,1073,1980,2006]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="7" pageNumber="391">Figs 1, 3</figureCitation>
<figureCitation id="1300F98AFFFBFFFF10B1F858FBFCAD84" box="[975,1073,1980,2006]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="7" pageNumber="391">Figs 1, 3</figureCitation>
). Scutellum almost squared with slightly rounded sides in
<taxonomicName id="4C3B9E8CFFFBFFFF12CDF805FDF0ADA8" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[435,573,2016,2042]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFBFFFF12CDF805FDF0ADA8" box="[435,573,2016,2042]" italics="true" pageId="7" pageNumber="391">B. bicolorus</emphasis>
@ -311,9 +313,9 @@ Size of
Head including eyes broader than pronotum (DV) in ratio 1:0.7 (HW =
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). The space between eyes (frons) very narrow (HW / FW = 4.26) (
<figureCitation id="1300F98AFFF4FFF011F4F86DFD11ADF1" box="[650,732,1929,1955]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="8" pageNumber="392">Fig. 12</figureCitation>
<figureCitation id="1300F98AFFF4FFF011F4F86DFD11ADF1" box="[650,732,1929,1955]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="8" pageNumber="392">Fig. 12</figureCitation>
). Antennae 1.47x longer than pronotum. Ultimate antennomere axe-like, widened (
<figureCitation id="1300F98AFFF4FFF01210F849FE7FAD95" box="[366,434,1965,1991]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="8" pageNumber="392">Fig. 6</figureCitation>
<figureCitation id="1300F98AFFF4FFF01210F849FE7FAD95" box="[366,434,1965,1991]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="8" pageNumber="392">Fig. 6</figureCitation>
).
</paragraph>
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@ -349,7 +351,7 @@ Longer than wide in ratio 1:1.5 (PL =
, PW =
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), subcylindrical but not parallel sided, widest before middle, sides of pronotum narrowing in basal third from pronotal base toward the middle part of pronotum, where it is distinctly widening and then slightly narrowing toward anterior edge of pronotum, surface roughly wrinkled, irregularly in dorsal part, more or less longitudinally coarsely wrinkled in lateral part (
<figureCitation id="1300F98AFFF5FFF11653FAD0FABFAF1D" box="[1325,1394,1332,1359]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="9" pageNumber="393">Fig. 9</figureCitation>
<figureCitation id="1300F98AFFF5FFF11653FAD0FABFAF1D" box="[1325,1394,1332,1359]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="9" pageNumber="393">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFF5FFF113B9FABDFE3DAC50" blockId="9.[151,1437,1224,1683]" pageId="9" pageNumber="393">

78
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<figureCitation id="1300F98AFFF9FFFD13E9FF5FFEF2AA87" box="[151,319,187,214]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="5" pageNumber="389">Figs 2, 5, 8, 11</figureCitation>
<figureCitation id="1300F98AFFF9FFFD13E9FF5FFEF2AA87" box="[151,319,187,214]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="5" pageNumber="389">Figs 2, 5, 8, 11</figureCitation>
, 21–32
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@ -203,7 +205,7 @@ Size of
Head including eyes broader than pronotum (DV) in ratio 1:0.8 (HW =
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). The space between eyes (frons) narrow (HW / FW = 2.8). Ultimate antennomere elongate-oval, 8.8x longer then broad (
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).
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@ -213,7 +215,7 @@ Longer than wide in ratio 1:1.3 (PL =
, PW =
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), cylindrical, apical two thirds of pronotum parallel sided and slightly trangled in apical third, basal third strongly trangled (
<figureCitation id="1300F98AFFF9FFFD17F7FC86FB03A92E" box="[1161,1230,866,892]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="5" pageNumber="389">Fig. 8</figureCitation>
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,
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, 41). It, however, can reliably be distinguished by biflabeloid antennae, with two flabelae situated at the beginning of each antennomere III–X. Moreover, the flabelae have the same diameter along their whole length (see also
<figureCitation id="1300F98AFFFDFFF91673FBFBFABDAE6B" box="[1293,1392,1055,1081]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="1" pageNumber="385">Figs 1–3</figureCitation>
<figureCitation id="1300F98AFFFDFFF91673FBFBFABDAE6B" box="[1293,1392,1055,1081]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 1–3</figureCitation>
). In other Madagascan genera which possess flabeloid or biflabeloid antennae, the flabelae are situated in the middle or at the end of antennomeres III–X and their diameter is not equal along their length but usually gradually widened toward their apex. Moreover, in some genera (e.g. in
<taxonomicName id="4C3B9E8CFFFDFFF91186FB6FFB8CAEF7" authority="Fairmaire, 1885" authorityName="Fairmaire" authorityYear="1885" box="[760,1089,1162,1189]" class="Insecta" family="Cleridae" genus="Diplocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFFDFFF91186FB6FFC4CAEF7" box="[760,897,1163,1189]" italics="true" pageId="1" pageNumber="385">Diplocladus</emphasis>
@ -187,19 +189,19 @@ Body cylindrical, shiny, sparsely pubescent. Body length
<paragraph id="8B84E50FFFFDFFF913B9FAFFFDEAAC2B" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913B9FAFFFEC3AF67" bold="true" box="[199,270,1307,1333]" pageId="1" pageNumber="385">Head.</emphasis>
Head including eyes broader than pronotum. Eyes large, with sparse erect setae, protruding laterally, anteriorly kidney-shaped (
<figureCitation id="1300F98AFFFDFFF912BDFADBFDF6AF0B" box="[451,571,1343,1369]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="1" pageNumber="385">Figs 7–12</figureCitation>
<figureCitation id="1300F98AFFFDFFF912BDFADBFDF6AF0B" box="[451,571,1343,1369]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 7–12</figureCitation>
). Frons and vertex wrinkled. Labrum bilobed, anterior margin with yellow setae and emarginate in middle (
<figureCitation id="1300F98AFFFDFFF9117EFA87FDB8AF2F" box="[512,629,1379,1405]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="1" pageNumber="385">Figs 7–12</figureCitation>
<figureCitation id="1300F98AFFFDFFF9117EFA87FDB8AF2F" box="[512,629,1379,1405]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 7–12</figureCitation>
). Terminal maxillary palpomeres cylindrical; terminal labial palpomeres securiform. Clypeus shiny, straight anteriorly, slightly dilated posteriad, sparsely pubescent.Antennae 11-segmented, biflabeloid, with flabelae situated at base of each antennomere III-X (
<figureCitation id="1300F98AFFFDFFF910C4FA4FFBEFAF97" box="[954,1058,1451,1477]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="1" pageNumber="385">Figs 1–3</figureCitation>
<figureCitation id="1300F98AFFFDFFF910C4FA4FFBEFAF97" box="[954,1058,1451,1477]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 1–3</figureCitation>
). Flabelae 2–3 times longer than length of respective antennomere. Flabelae equal in width along their whole length, divided, unjointed.Antennomere I thickened, approx. 2.5 longer than antennomere II, which is strongly shortened. Antennomere II with a small pronounced bump on inner side. Antennomeres III-X approx. equal in length. Dorsal side of antennomeres sparsely covered with long yellow recumbent hairs whereas inner side of antennomeres and flabelae is densely covered with shorter white erect hairs (
<figureCitation id="1300F98AFFFDFFF912CAF9BBFDD4AC2B" box="[436,537,1631,1657]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="1" pageNumber="385">Figs 1–3</figureCitation>
<figureCitation id="1300F98AFFFDFFF912CAF9BBFDD4AC2B" box="[436,537,1631,1657]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 1–3</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFDFFF913B9F967FCC4AC93" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913B9F967FE8AACCF" bold="true" box="[199,327,1667,1693]" pageId="1" pageNumber="385">Pronotum.</emphasis>
Shiny, longer than wide, cylindrical, surface roughly and irregularly wrinkled, whole surface covered with long erect and semi-erect hairs (
<figureCitation id="1300F98AFFFDFFF911E8F943FD30AC93" box="[662,765,1703,1729]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="1" pageNumber="385">Figs 7–9</figureCitation>
<figureCitation id="1300F98AFFFDFFF911E8F943FD30AC93" box="[662,765,1703,1729]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="1" pageNumber="385">Figs 7–9</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFDFFF913B9F92FFD09AD27" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">

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<subSubSection id="C321B684FFFEFFFA13E9FE8AFDF3ABDA" box="[151,574,366,393]" pageId="2" pageNumber="386" type="nomenclature">
<paragraph id="8B84E50FFFFEFFFA13E9FE8AFDF3ABDA" blockId="2.[151,574,366,429]" box="[151,574,366,393]" pageId="2" pageNumber="386">
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@ -61,7 +63,7 @@
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13E9FE77FE5FABFF" box="[151,402,403,429]" pageId="2" pageNumber="386" type="description">
<paragraph id="8B84E50FFFFEFFFA13E9FE77FE5FABFF" blockId="2.[151,574,366,429]" box="[151,402,403,429]" pageId="2" pageNumber="386">
<figureCitation id="1300F98AFFFEFFFA13E9FE77FE8DABFF" box="[151,320,403,429]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="2" pageNumber="386">Figs 1, 4, 7, 10</figureCitation>
<figureCitation id="1300F98AFFFEFFFA13E9FE77FE8DABFF" box="[151,320,403,429]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="2" pageNumber="386">Figs 1, 4, 7, 10</figureCitation>
, 13–20
</paragraph>
</subSubSection>
@ -304,7 +306,7 @@ in
<emphasis id="B94F391DFFFEFFFA13C8FAABFE84AF3A" box="[182,329,1359,1384]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
;
<figureCitation id="1300F98AFFFEFFFA1227FAABFE0DAF3B" box="[345,448,1358,1385]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="2" pageNumber="386">Figs 4, 5</figureCitation>
<figureCitation id="1300F98AFFFEFFFA1227FAABFE0DAF3B" box="[345,448,1358,1385]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="2" pageNumber="386">Figs 4, 5</figureCitation>
). From the new species
<taxonomicName id="4C3B9E8CFFFEFFFA11A6FAABFCAEAF3B" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[728,867,1359,1385]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA11A6FAABFCAEAF3B" box="[728,867,1359,1385]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
@ -324,7 +326,7 @@ it can also be distinguished by distinctly securiform ultimate antennomere (more
<taxonomicNameLabel id="A27C8466FFFEFFFA12E2FA73FE38AFE3" box="[412,501,1431,1457]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="1300F98AFFFEFFFA117DFA73FDA9AFE3" box="[515,612,1431,1457]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="2" pageNumber="386">Figs 7, 9</figureCitation>
<figureCitation id="1300F98AFFFEFFFA117DFA73FDA9AFE3" box="[515,612,1431,1457]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="2" pageNumber="386">Figs 7, 9</figureCitation>
) and dark brown abdomen (distinctly pale-brown in
<taxonomicName id="4C3B9E8CFFFEFFFA17D3FA73FAFBAFE3" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1197,1334,1431,1457]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA17D3FA73FAFBAFE3" box="[1197,1334,1431,1457]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
@ -355,7 +357,7 @@ Size of
Head including eyes broader than pronotum (DV) in ratio 1:0.8 (HW =
<quantity id="4CC348EAFFFEFFFA1756F9C3FB4CAC13" box="[1064,1153,1575,1601]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="386" unit="mm" value="2.1">2.1 mm</quantity>
). The space between eyes (frons) narrow (HW / FW = 3.5). Ultimate antennomere almost securifom, 3.4x longer then broad (
<figureCitation id="1300F98AFFFEFFFA1793F9AFFAFFAC37" box="[1261,1330,1611,1637]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="2" pageNumber="386">Fig. 4</figureCitation>
<figureCitation id="1300F98AFFFEFFFA1793F9AFFAFFAC37" box="[1261,1330,1611,1637]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="2" pageNumber="386">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F98BFC81ACFF" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
@ -365,7 +367,7 @@ Longer than wide in ratio 1:1.7 (PL =
, PW =
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), cylindrical, apical two thirds of pronotum parallel sided, basal third strongly trangled (
<figureCitation id="1300F98AFFFEFFFA1187F977FCF3ACFF" box="[761,830,1683,1709]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." pageId="2" pageNumber="386">Fig. 7</figureCitation>
<figureCitation id="1300F98AFFFEFFFA1187F977FCF3ACFF" box="[761,830,1683,1709]" captionStart="FIGURE 1–12" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 1–12. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 1–3—right antenna, antennomers II-VI; 4–6—right antenna, antennomers IX–XI (red arrows indicate last antennomere); 7–9—head and pronotum, dorsal view; 10–12—head, front view." figureDoi="http://doi.org/10.5281/zenodo.13939777" httpUri="https://zenodo.org/record/13939777/files/figure.png" pageId="2" pageNumber="386">Fig. 7</figureCitation>
).
</paragraph>
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