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<mods:title id="2EA1742CB39BFE7789EA35EE768C02DC">Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae</mods:title>
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<heading id="2653395AFF89D363EBE6F890FD9CF869" bold="true" box="[151,613,1894,1920]" fontSize="11" level="1" pageId="37" pageNumber="188" reason="1">
<emphasis id="4FD05224FF89D363EBE6F890FD9CF869" bold="true" box="[151,613,1894,1920]" pageId="37" pageNumber="188">
The
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<taxonomicName id="BAA4F5B5FF89D363EBBEF890FE81F869" authorityName="Zwick" authorityYear="2013" box="[207,376,1894,1920]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">Austrocyphon</taxonomicName>
tribulator-
</emphasis>
complex
</emphasis>
</heading>
</paragraph>
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(
<figureCitation id="E59F92B3FF89D363EBEEF87FFEE6F84B" box="[159,287,1929,1954]" captionStart="FIGURES 95 98" captionStartId="39.[151,264,1213,1235]" captionTargetBox="[196,1389,473,1161]" captionTargetId="figure@39.[189,1397,469,1169]" captionTargetPageId="39" captionText="FIGURES 95 98. Austrocyphon tribulator- complex, morphs of male T 9. 95, typical morph (near Cape Tribulation); aberrant morphs: 96, 97 from Heathlands; 98, from Broken Nose. Not to scale; 95, 96 from Zwick (2013 c)." httpUri="https://zenodo.org/record/1052577/files/figure.png" pageId="37" pageNumber="188">Figs 9598</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="35BEDDBDFF89D361EBE6F827FC50FE61" lastPageId="39" lastPageNumber="190" pageId="37" pageNumber="188" type="materials_examined">
<paragraph id="7D1B8E36FF89D360EBE6F827FC60FEF5" blockId="37.[151,1436,2001,2026]" lastBlockId="38.[151,1436,151,285]" lastPageId="38" lastPageNumber="189" pageId="37" pageNumber="188">
<materialsCitation id="CDCC846BFF89D360EBE6F827FE5FFF59" ID-GBIF-Occurrence="1701200976" collectingDate="1992-06-07" collectingDateMax="1992-07-25" collectingDateMin="1992-06-07" collectorName="P. Zborowski &amp; E. Nielsen Malaise" lastPageId="38" lastPageNumber="189" latitude="-11.45" location="New" longLatPrecision="777" longitude="142.35" municipality="Heathlands" pageId="37" pageNumber="188" specimenCount="1" specimenCount-male="1">
<emphasis id="4FD05224FF89D363EBE6F827FE59F803" bold="true" box="[151,416,2001,2026]" pageId="37" pageNumber="188">
<location id="787BD8EDFF89D363EBE6F827FF35F803" LSID="urn:lsid:plazi:treatment:F50D3F20FF89D361EBE6F890FC50FE61:787BD8EDFF89D363EBE6F827FF35F803" box="[151,204,2001,2026]" latitude="-11.45" longLatPrecision="777" longitude="142.35" municipality="Heathlands" name="New" pageId="37" pageNumber="188">New</location>
material studied.
</emphasis>
<specimenCount id="6BA245BFFF89D363EAD9F824FE37F803" box="[424,462,2002,2026]" pageId="37" pageNumber="188" type="male">1♂</specimenCount>
:
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<geoCoordinate id="1890E8F1FF89D363E940F827FD76F803" box="[561,655,2001,2026]" direction="east" orientation="longitude" pageId="37" pageNumber="188" precision="555" value="142.35">142.35E</geoCoordinate>
<emphasis id="4FD05224FF89D363E9E9F827FD2CF803" bold="true" box="[664,725,2001,2026]" pageId="37" pageNumber="188">QLD</emphasis>
<collectingMunicipality id="9D7F144CFF89D363E9ACF827FCA2F803" box="[733,859,2001,2026]" pageId="37" pageNumber="188">Heathlands</collectingMunicipality>
dump
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<collectingDate id="195E511EFF89D363E8DDF827FB73F803" box="[940,1162,2001,2026]" pageId="37" pageNumber="188" value="1992-06-07" valueMax="1992-07-25" valueMin="1992-06-07">7June-25 July 1992</collectingDate>
</date>
<collectorName id="D051EBE0FF89D363EFE3F827FADBF803" box="[1170,1314,2001,2026]" pageId="37" pageNumber="188">P.Zborowski</collectorName>
,
<collectorName id="D051EBE0FF89D360EE5EF827FF09FF59" lastPageId="38" lastPageNumber="189" pageId="37" pageNumber="188">E.Nielsen Malaise</collectorName>
#2 open forest
</materialsCitation>
;
<materialsCitation id="CDCC846BFF8AD360EAC4FF6EFE54FF3C" ID-GBIF-Occurrence="1701200972" collectingDate="1992-08-18" collectingDateMax="1992-09-18" collectingDateMin="1992-08-18" collectorName="P. Zborowski &amp; L. Miller" latitude="-11.45" location="Dump" longLatPrecision="777" longitude="142.35" pageId="38" pageNumber="189" specimenCount="1" specimenCount-male="1">
<specimenCount id="6BA245BFFF8AD360EAC4FF6EFE22FF59" box="[437,475,152,176]" pageId="38" pageNumber="189" type="male">1♂</specimenCount>
:
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<geoCoordinate id="1890E8F1FF8AD360E932FF61FD58FF59" box="[579,673,151,176]" direction="east" orientation="longitude" pageId="38" pageNumber="189" precision="555" value="142.35">142.35E</geoCoordinate>
QLD Heathlands-
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site
<date id="091AA8F6FF8AD360EF77FF6EFB03FF59" box="[1030,1274,152,176]" pageId="38" pageNumber="189" value="1992-08-18" valueMax="1992-09-18" valueMin="1992-08-18">
<collectingDate id="195E511EFF8AD360EF77FF6EFB03FF59" box="[1030,1274,152,176]" pageId="38" pageNumber="189" value="1992-08-18" valueMax="1992-09-18" valueMin="1992-08-18">18 Aug-18 Sep 1992</collectingDate>
</date>
<collectingMethod id="A4E5F621FF8AD360EE74FF61FA65FF46" box="[1285,1436,151,176]" pageId="38" pageNumber="189">Malaise Trap</collectingMethod>
<collectorName id="D051EBE0FF8AD360EBE6FF4AFEDAFF3C" box="[151,291,188,213]" pageId="38" pageNumber="189">P.Zborowski</collectorName>
&amp;
<collectorName id="D051EBE0FF8AD360EA3AFF4AFE54FF3C" box="[331,429,188,213]" pageId="38" pageNumber="189">L.Miller</collectorName>
</materialsCitation>
;
<materialsCitation id="CDCC846BFF8AD360EAC8FF4AFCDFFF1E" ID-GBIF-Occurrence="1701200971" collectingDate="1989-06-27" collectingDateMax="1989-07-12" collectingDateMin="1989-06-27" collectorName="T. A. Weir" elevation="520" latitude="-13.44" location="Bald Hill McIlwraith" longLatPrecision="774" longitude="143.2" pageId="38" pageNumber="189" specimenCount="3" specimenCount-male="3">
<specimenCount id="6BA245BFFF8AD360EAC8FF4AFE19FF3D" box="[441,480,188,212]" pageId="38" pageNumber="189" type="male">3♂</specimenCount>
:
<geoCoordinate id="1890E8F1FF8AD360EA9CFF4AFDC2FF3D" box="[493,571,188,212]" direction="south" orientation="latitude" pageId="38" pageNumber="189" precision="555" value="-13.44">13.44S</geoCoordinate>
<geoCoordinate id="1890E8F1FF8AD360E935FF4AFD5BFF3D" box="[580,674,188,212]" direction="east" orientation="longitude" pageId="38" pageNumber="189" precision="555" value="143.2">143.20E</geoCoordinate>
QLD
<quantity id="BA5C23D3FF8AD360E99CFF4AFCD5FF3C" box="[749,812,188,213]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="38" pageNumber="189" unit="km" value="11.0">11km</quantity>
WbyN of
<location id="787BD8EDFF8AD360E8D9FF4AFB61FF3C" LSID="urn:lsid:plazi:treatment:F50D3F20FF89D361EBE6F890FC50FE61:787BD8EDFF8AD360E8D9FF4AFB61FF3C" box="[936,1176,188,213]" latitude="-13.44" longLatPrecision="774" longitude="143.2" name="Bald Hill McIlwraith" pageId="38" pageNumber="189">Bald Hill McIlwraith</location>
range
<date id="091AA8F6FF8AD360EF96FF4AFF36FF11" pageId="38" pageNumber="189" value="1989-06-27" valueMax="1989-07-12" valueMin="1989-06-27">
<collectingDate id="195E511EFF8AD360EF96FF4AFF36FF11" pageId="38" pageNumber="189" value="1989-06-27" valueMax="1989-07-12" valueMin="1989-06-27">27 June-12 July 1989</collectingDate>
</date>
<collectorName id="D051EBE0FF8AD360EBA6FF29FEC7FF11" box="[215,318,223,248]" pageId="38" pageNumber="189">T.A.Weir</collectorName>
<quantity id="BA5C23D3FF8AD360EA34FF29FE7CFF11" box="[325,389,223,248]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="38" pageNumber="189" unit="m" value="520.0">
<elevation id="F6896905FF8AD360EA34FF29FE7CFF11" box="[325,389,223,248]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="38" pageNumber="189" unit="m" value="520.0">520m</elevation>
</quantity>
search party campsite /
<collectingMethod id="A4E5F621FF8AD360E9E8FF16FCDFFF1E" box="[665,806,223,248]" pageId="38" pageNumber="189">ex light trap</collectingMethod>
</materialsCitation>
.
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<specimenCount id="6BA245BFFF8AD360E840FF16FCADFF11" box="[817,852,223,248]" pageId="38" pageNumber="189" type="male">
1
<emphasis id="4FD05224FF8AD360E84EFF29FCADFF11" bold="true" box="[831,852,223,248]" pageId="38" pageNumber="189"></emphasis>
</specimenCount>
:
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NP, QLD
<quantity id="BA5C23D3FF8AD360EF0AFF29FB3BFF11" box="[1147,1218,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="189" unit="km" value="1.5">1.5km</quantity>
ENE
<collectingMunicipality id="9D7F144CFF8AD360EE79FF29FA65FF1E" box="[1288,1436,223,248]" pageId="38" pageNumber="189">Broken Nose</collectingMunicipality>
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<collectingDate id="195E511EFF8AD360EBE6FEF2FED5FEF5" box="[151,300,260,284]" pageId="38" pageNumber="189" value="1992-06-28">28 June 1992</collectingDate>
</date>
<collectorName id="D051EBE0FF8AD360EA42FEF2FE7BFEF4" box="[307,386,260,285]" pageId="38" pageNumber="189">C.Reid</collectorName>
beating rainforest vegetation
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</quantity>
(all
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).
</materialsCitation>
</paragraph>
<caption id="29DBDEBEFF8AD360EBE6F93AFDA9F8F6" httpUri="https://zenodo.org/record/1052575/files/figure.png" pageId="38" pageNumber="189" targetBox="[164,1420,326,1722]" targetPageId="38">
<paragraph id="7D1B8E36FF8AD360EBE6F93AFDA9F8F6" blockId="38.[151,1436,1740,1823]" pageId="38" pageNumber="189">
<emphasis id="4FD05224FF8AD360EBE6F93AFEAEF90B" bold="true" box="[151,343,1740,1762]" pageId="38" pageNumber="189">FIGURES 9094.</emphasis>
<taxonomicName id="BAA4F5B5FF8AD360EA11F93AFDC3F90B" authority="Zwick, 2016" authorityName="Zwick" authorityYear="2016" box="[352,570,1740,1762]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="scissus" status="sp. nov.">
<emphasis id="4FD05224FF8AD360EA11F93AFDC3F90B" box="[352,570,1740,1762]" italics="true" pageId="38" pageNumber="189">Austrocyphon scissus</emphasis>
</taxonomicName>
<emphasis id="4FD05224FF8AD360E932F93AFD86F90B" bold="true" box="[579,639,1740,1762]" pageId="38" pageNumber="189">
<taxonomicNameLabel id="54E3EF5FFF8AD360E932F93AFD86F90B" box="[579,639,1740,1762]" pageId="38" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male terminalia. 90, T8; 91, T9; 92, S9 and tegmen; 93, penis; 94, apex of penis, enlarged. 9193 to the same scale. Abbreviations: ce, centema; pa, pala; pd, the fused parameroids; pe, paramere; te, tegmen; tb, transverse bridge; tr, trigonium.
</paragraph>
</caption>
<paragraph id="7D1B8E36FF8AD360EBB6F8BAFB80F81C" blockId="38.[151,1436,1868,2037]" pageId="38" pageNumber="189">
<taxonomicName id="BAA4F5B5FF8AD360EBB6F8BAFD84F88D" authority="Zwick, 2013" authorityName="Zwick: Zwick" authorityYear="2013" box="[199,637,1868,1893]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="tribulator">
<emphasis id="4FD05224FF8AD360EBB6F8BAFE22F88D" box="[199,475,1868,1892]" italics="true" pageId="38" pageNumber="189">Austrocyphon tribulator</emphasis>
Zwick, 2013
</taxonomicName>
is a small (BL ~
<quantity id="BA5C23D3FF8AD360E815F8BAFC54F88D" box="[868,941,1868,1892]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="189" unit="mm" value="2.0">2 mm</quantity>
) yellowish to light brown species that is widespread in
<collectingRegion id="BF6040D4FF8AD360EA4DF899FE3DF861" box="[316,452,1903,1928]" country="Australia" name="Queensland" pageId="38" pageNumber="189">Queensland</collectingRegion>
. It differs from other members of the
<taxonomicName id="BAA4F5B5FF8AD360E803F886FBF9F861" box="[882,1024,1904,1928]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="189" phylum="Arthropoda" rank="species" species="adelaidae">
<emphasis id="4FD05224FF8AD360E803F886FBF9F861" box="[882,1024,1904,1928]" italics="true" pageId="38" pageNumber="189">A. adelaidae</emphasis>
</taxonomicName>
-group by the wide, completely bare plate of T8 and mainly by the X-shaped arrangement of the sclerites representing T9. The basal apodemes converge and eventually meet, and from the meeting point issues a pair of divergent caudal sclerites. In the typical morph, the caudal sclerites are flat lanceolate blades with fine asperities on the surface (
<figureCitation id="E59F92B3FF8AD360EF69F82AFB92F81C" box="[1048,1131,2012,2037]" captionStart="FIGURES 95 98" captionStartId="39.[151,264,1213,1235]" captionTargetBox="[196,1389,473,1161]" captionTargetId="figure@39.[189,1397,469,1169]" captionTargetPageId="39" captionText="FIGURES 95 98. Austrocyphon tribulator- complex, morphs of male T 9. 95, typical morph (near Cape Tribulation); aberrant morphs: 96, 97 from Heathlands; 98, from Broken Nose. Not to scale; 95, 96 from Zwick (2013 c)." httpUri="https://zenodo.org/record/1052577/files/figure.png" pageId="38" pageNumber="189">Fig. 95</figureCitation>
).
</paragraph>
<paragraph id="7D1B8E36FF8BD361EBB6FF6EFD89FEF4" blockId="39.[151,1436,151,392]" pageId="39" pageNumber="190">
Some aberrant specimens were recorded when the species was first described. They differed mainly by the narrowness of the caudal T9 sclerites (
<figureCitation id="E59F92B3FF8BD361E93BFF4AFD66FF3C" box="[586,671,188,213]" captionStart="FIGURES 95 98" captionStartId="39.[151,264,1213,1235]" captionTargetBox="[196,1389,473,1161]" captionTargetId="figure@39.[189,1397,469,1169]" captionTargetPageId="39" captionText="FIGURES 95 98. Austrocyphon tribulator- complex, morphs of male T 9. 95, typical morph (near Cape Tribulation); aberrant morphs: 96, 97 from Heathlands; 98, from Broken Nose. Not to scale; 95, 96 from Zwick (2013 c)." httpUri="https://zenodo.org/record/1052577/files/figure.png" pageId="39" pageNumber="190">Fig. 96</figureCitation>
). Shapes of T8, S9 and the penis also differed but were not uniform among the aberrant individuals from several localities. Some more variations have now been observed (
<figureCitation id="E59F92B3FF8BD361EE4BFF29FA76FF11" box="[1338,1423,223,248]" captionStart="FIGURES 95 98" captionStartId="39.[151,264,1213,1235]" captionTargetBox="[196,1389,473,1161]" captionTargetId="figure@39.[189,1397,469,1169]" captionTargetPageId="39" captionText="FIGURES 95 98. Austrocyphon tribulator- complex, morphs of male T 9. 95, typical morph (near Cape Tribulation); aberrant morphs: 96, 97 from Heathlands; 98, from Broken Nose. Not to scale; 95, 96 from Zwick (2013 c)." httpUri="https://zenodo.org/record/1052577/files/figure.png" pageId="39" pageNumber="190">Fig. 97</figureCitation>
), further increasing the diversity of morphs.
</paragraph>
<paragraph id="7D1B8E36FF8BD361EBB6FED1FC50FE61" blockId="39.[151,1436,151,392]" pageId="39" pageNumber="190">
The significance and cause of the heterogeneity are unknown. For the time being the specimens are regarded as members of an unresolved complex, including a single individual (from Broken Nose) whose T9 sclerites have an approximately circular cross section and appear club shaped (
<figureCitation id="E59F92B3FF8BD361E839FE99FC64FE61" box="[840,925,367,392]" captionStart="FIGURES 95 98" captionStartId="39.[151,264,1213,1235]" captionTargetBox="[196,1389,473,1161]" captionTargetId="figure@39.[189,1397,469,1169]" captionTargetPageId="39" captionText="FIGURES 95 98. Austrocyphon tribulator- complex, morphs of male T 9. 95, typical morph (near Cape Tribulation); aberrant morphs: 96, 97 from Heathlands; 98, from Broken Nose. Not to scale; 95, 96 from Zwick (2013 c)." httpUri="https://zenodo.org/record/1052577/files/figure.png" pageId="39" pageNumber="190">Fig. 98</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>