Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae Author Zwick, Peter text Zootaxa 2016 2016-03-02 4085 2 151 198 journal article 31436 10.11646/zootaxa.4085.2.1 ba498adf-4dec-4fa3-8213-9d4d8c3b9dd3 1175-5326 1052535 E41CB99C-5177-47A7-A424-2453D27E48F0 The Austrocyphon tribulator- complex ( Figs 95–98 ) New material studied. 1♂ : 11.45S 142.35E QLD Heathlands dump 7June-25 July 1992 P.Zborowski , E.Nielsen Malaise #2 open forest ; 1♂ : 11.45S 142.35E QLD Heathlands- Dump site 18 Aug-18 Sep 1992 Malaise Trap P.Zborowski & L.Miller ; 3♂ : 13.44S 143.20E QLD 11km WbyN of Bald Hill McIlwraith range 27 June-12 July 1989 T.A.Weir 520m search party campsite / ex light trap . 1 ♂ : Bellender Ker NP, QLD 1.5km ENE Broken Nose 28 June 1992 C.Reid beating rainforest vegetation 500m (all ANIC ). FIGURES 90−94. Austrocyphon scissus n. sp. , male terminalia. 90, T8; 91, T9; 92, S9 and tegmen; 93, penis; 94, apex of penis, enlarged. 91−93 to the same scale. Abbreviations: ce, centema; pa, pala; pd, the fused parameroids; pe, paramere; te, tegmen; tb, transverse bridge; tr, trigonium. Austrocyphon tribulator Zwick, 2013 is a small (BL ~ 2 mm ) yellowish to light brown species that is widespread in Queensland . It differs from other members of the A. adelaidae -group by the wide, completely bare plate of T8 and mainly by the X-shaped arrangement of the sclerites representing T9. The basal apodemes converge and eventually meet, and from the meeting point issues a pair of divergent caudal sclerites. In the typical morph, the caudal sclerites are flat lanceolate blades with fine asperities on the surface ( Fig. 95 ). Some aberrant specimens were recorded when the species was first described. They differed mainly by the narrowness of the caudal T9 sclerites ( Fig. 96 ). Shapes of T8, S9 and the penis also differed but were not uniform among the aberrant individuals from several localities. Some more variations have now been observed ( Fig. 97 ), further increasing the diversity of morphs. The significance and cause of the heterogeneity are unknown. For the time being the specimens are regarded as members of an unresolved complex, including a single individual (from Broken Nose) whose T9 sclerites have an approximately circular cross section and appear club shaped ( Fig. 98 ).