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<document id="714E766A32429317447B0453A35FCFFD" ID-DOI="10.5281/zenodo.5164903" ID-ISSN="1942-1354" ID-Zenodo-Dep="5164903" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628190592502" checkinUser="felipe" docAuthor="Ratcliffe, Brett C. &amp; Cave, Ronald D." docDate="2010" docId="B65987E9FF872A2FFF07DD28FD5AF95A" docLanguage="en" docName="InsectaMundi.0139.1-15.pdf" docOrigin="Insecta Mundi 2010 (139)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Tomarus adoceteus Ratcliffe and Cave 2010, new species" docType="treatment" docVersion="3" lastPageNumber="8" masterDocId="4A60FF91FF8F2A26FFC7D90CFFECFF81" masterDocTitle="The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini)" masterLastPageNumber="15" masterPageNumber="1" pageNumber="7" updateTime="1699035530847" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="4A063EB245AED4A5E37774244B6E0FED" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AA535044A56B49125EA752B11BEDF957">
<mods:title id="B57F6E0C8101284A32224A04ECC50363">The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini)</mods:title>
</mods:titleInfo>
<mods:name id="1217967AA1A8F8176767F79BE99AEEEC" type="personal">
<mods:role id="8BD69695A974645DB18DACFEBFED1BA6">
<mods:roleTerm id="F185B855F2D8586980E2D194ECC402D3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CC09B885F05DEE8EEB2FEE3792CD0B9B">Ratcliffe, Brett C.</mods:namePart>
</mods:name>
<mods:name id="005A13343C6D71DC8D091FB4EC78A001" type="personal">
<mods:role id="1D73D3FBEA509600EBD36F92A2C795A1">
<mods:roleTerm id="6E85BD92F2A5B99D6B675CC6EEC571B3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E8FACA75C70D7FBC9D81B03270E8CBF3">Cave, Ronald D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B50CBEAC31C04D168C62931916CF1876">text</mods:typeOfResource>
<mods:relatedItem id="5EF8C0DFA7453E56A553BFDF3D226C8D" type="host">
<mods:titleInfo id="B62BB74B1992FC9D12EB8FBE787C672E">
<mods:title id="FECF73522D366DE1AA489A5215AB4197">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="61A98CAB6A689EEC09A7961F6EB5311D">
<mods:date id="75352B6FA5126D2329FE2B67A760AF51">2010</mods:date>
<mods:detail id="525BE68E6037664F708A75074138F1FA" type="pubDate">
<mods:number id="E6394A6362450D0D406FFD2A3C47D886">2010-09-24</mods:number>
</mods:detail>
<mods:detail id="19E09172093230A727E71E012703D8FC" type="volume">
<mods:number id="F7F3978E2E864F790794D0D5951F0280">2010</mods:number>
</mods:detail>
<mods:detail id="2F9015E6949E0B23FDF32025F24AFFFA" type="issue">
<mods:number id="32EEC4C8E7C58E287407A3A58B032181">139</mods:number>
</mods:detail>
<mods:extent id="B79C1606570B62C463BAA744F30DD780" unit="page">
<mods:start id="8D44D956A9E0A30AD38FE09EA62CB6C6">1</mods:start>
<mods:end id="9766421E80DB9AA58E90E077DD3910F4">15</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="356E7477213DEAC452ACC02916B568FF">journal article</mods:classification>
<mods:identifier id="8C0CF56AC41901050AE9E519BF5F8FB2" type="DOI">10.5281/zenodo.5164903</mods:identifier>
<mods:identifier id="F1C9189DE3263A7C2E0C16A2BC840060" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="3825CC0266AB5C1305B28D1857022A3E" type="Zenodo-Dep">5164903</mods:identifier>
</mods:mods>
<treatment id="B65987E9FF872A2FFF07DD28FD5AF95A" LSID="urn:lsid:plazi:treatment:B65987E9FF872A2FFF07DD28FD5AF95A" httpUri="http://treatment.plazi.org/id/B65987E9FF872A2FFF07DD28FD5AF95A" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<subSubSection id="76EA6574FF872A2EFF07DD28FEA4FBFA" pageId="8" pageNumber="7" type="nomenclature">
<paragraph id="3E4F36FFFF872A2EFF07DD28FE8EFBDA" blockId="8.[192,870,1038,1147]" pageId="8" pageNumber="7">
<heading id="65078193FF872A2EFF07DD28FE8EFBDA" bold="true" fontSize="10" level="2" pageId="8" pageNumber="7" reason="3">
<emphasis id="0C84EAEDFF872A2EFF07DD28FE8EFBDA" bold="true" pageId="8" pageNumber="7">
<taxonomicName id="F9F04D7CFF872A2EFF07DD28FD2AFBBA" authority="Ratcliffe and Cave" authorityName="Ratcliffe and Cave" authorityYear="2010" box="[192,710,1058,1083]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="adoceteus" status="sp. nov.">
<emphasis id="0C84EAEDFF872A2EFF07DD28FE2AFBBA" bold="true" box="[192,454,1058,1083]" italics="true" pageId="8" pageNumber="7">Tomarus adoceteus</emphasis>
Ratcliffe and Cave
</taxonomicName>
,
<taxonomicNameLabel id="17B75796FF872A2EFF07DD4FFE8EFBDA" box="[192,354,1091,1115]" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DD6FFEA4FBFA" blockId="8.[192,870,1038,1147]" box="[192,328,1123,1147]" pageId="8" pageNumber="7">(Fig. 10-12)</paragraph>
</subSubSection>
<subSubSection id="76EA6574FF872A2FFF07DDAFFCA0FA5A" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7" type="materials_examined">
<paragraph id="3E4F36FFFF872A2EFF07DDAFFCFCFB3A" blockId="8.[192,1056,1187,1787]" box="[192,784,1187,1211]" pageId="8" pageNumber="7">
<emphasis id="0C84EAEDFF872A2EFF07DDAFFE69FB3A" bold="true" box="[192,389,1187,1211]" pageId="8" pageNumber="7">Type Material.</emphasis>
<typeStatus id="E14B885DFF872A2EFE4DDDAFFE1BFB3A" box="[394,503,1187,1211]" pageId="8" pageNumber="7" type="holotype">Holotype</typeStatus>
male, labeled “Roy Soc-
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DDCFFCFCFB5A" blockId="8.[192,1056,1187,1787]" box="[192,784,1219,1243]" pageId="8" pageNumber="7">CIG Expdn./Little Cayman, BWI/Pirates’ Point/</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DDEFFCFDFB7A" blockId="8.[192,1056,1187,1787]" box="[192,785,1251,1275]" pageId="8" pageNumber="7">
house light/
<date id="4A4E103FFF872A2EFE8CDDEFFE2FFB7A" box="[331,451,1251,1275]" pageId="8" pageNumber="7" value="1975-07-31">31.7.1975</date>
. R. R. Askew//BM 1981/342”
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DC0FFCFCFA9A" blockId="8.[192,1056,1187,1787]" box="[192,784,1283,1307]" pageId="8" pageNumber="7">
(hand written)/ our red
<typeStatus id="E14B885DFF872A2EFE12DC0FFDD0FA9A" box="[469,572,1283,1307]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
label.
<typeStatus id="E14B885DFF872A2EFD4FDC0FFD01FA9A" box="[648,749,1283,1307]" pageId="8" pageNumber="7" type="allotype">Allotype</typeStatus>
fe-
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DC2FFCFCFABA" blockId="8.[192,1056,1187,1787]" box="[192,784,1315,1339]" pageId="8" pageNumber="7">
male with same data but date of
<date id="4A4E103FFF872A2EFD81DC2FFD42FABA" box="[582,686,1315,1339]" pageId="8" pageNumber="7" value="1975-08-09">9.8.1975</date>
and our
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DC4FFCFCFADA" blockId="8.[192,1056,1187,1787]" box="[192,784,1347,1372]" pageId="8" pageNumber="7">
red
<typeStatus id="E14B885DFF872A2EFF2BDC4FFEA7FADA" box="[236,331,1347,1371]" pageId="8" pageNumber="7" type="allotype">allotype</typeStatus>
label. Single male
<typeStatus id="E14B885DFF872A2EFDE0DC48FD7EFADD" box="[551,658,1348,1372]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
and single
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DC6FFCFCFAFA" blockId="8.[192,1056,1187,1787]" box="[192,784,1379,1404]" pageId="8" pageNumber="7">
female
<typeStatus id="E14B885DFF872A2EFEDDDC68FE65FAFD" box="[282,393,1380,1404]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
with same data as
<typeStatus id="E14B885DFF872A2EFDB0DC6FFD36FAFA" box="[631,730,1379,1403]" pageId="8" pageNumber="7" type="allotype">allotype</typeStatus>
and
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DC8FFCFCFA1D" blockId="8.[192,1056,1187,1787]" box="[192,784,1411,1436]" pageId="8" pageNumber="7">
with our yellow
<typeStatus id="E14B885DFF872A2EFEB9DC88FE07FA1D" box="[382,491,1412,1436]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
labels. Female
<typeStatus id="E14B885DFF872A2EFD64DC88FCFCFA1D" box="[675,784,1412,1436]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DCAFFCFCFA3A" blockId="8.[192,1056,1187,1787]" box="[192,784,1443,1467]" pageId="8" pageNumber="7">labeled “WEST INDIES: Cayman Is./Grand Cay-</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DCCFFCFCFA5A" blockId="8.[192,1056,1187,1787]" box="[192,784,1475,1499]" pageId="8" pageNumber="7">
man/
<date id="4A4E103FFF872A2EFF39DCCFFE8EFA5A" box="[254,354,1475,1499]" pageId="8" pageNumber="7" value="1992-06">VI-1992</date>
coll. P. Fitzgerald/blacklight trap”
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DCEFFCFCFA7A" blockId="8.[192,1056,1187,1787]" box="[192,784,1507,1532]" pageId="8" pageNumber="7">
and with our yellow
<typeStatus id="E14B885DFF872A2EFE1BDCE8FDBCFA7D" box="[476,592,1508,1532]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
label. Female
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DF08FCFCF99A" blockId="8.[192,1056,1187,1787]" box="[192,784,1539,1564]" pageId="8" pageNumber="7">
<typeStatus id="E14B885DFF872A2EFF07DF08FEC3F99D" box="[192,303,1540,1564]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
labeled “Grand Cayman Is. B. W. I./
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DF2FFCFCF9BA" blockId="8.[192,1056,1187,1787]" box="[192,784,1571,1595]" pageId="8" pageNumber="7">
<collectingRegion id="FC34F81DFF872A2EFF07DF2FFEB8F9BA" box="[192,340,1571,1595]" country="Cayman Islands" name="George Town" pageId="8" pageNumber="7">Georgetown</collectingRegion>
/
<date id="4A4E103FFF872A2EFE9ADF2FFE11F9BA" box="[349,509,1571,1595]" pageId="8" pageNumber="7" value="1973-09-16">16 Sept. 1973</date>
/E. J. Gerberg//at black
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DF4FFCFCF9DA" blockId="8.[192,1056,1187,1787]" box="[192,784,1603,1628]" pageId="8" pageNumber="7">
light” and with our yellow
<typeStatus id="E14B885DFF872A2EFDCCDF48FD96F9DD" box="[523,634,1604,1628]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
label. Holo-
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DF68FCFCF9FA" blockId="8.[192,1056,1187,1787]" box="[192,784,1635,1660]" pageId="8" pageNumber="7">
type and
<typeStatus id="E14B885DFF872A2EFEF4DF6FFE7AF9FA" box="[307,406,1635,1659]" pageId="8" pageNumber="7" type="allotype">allotype</typeStatus>
deposited at The Natural His-
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DF88FCFCF91A" blockId="8.[192,1056,1187,1787]" box="[192,784,1667,1692]" pageId="8" pageNumber="7">
tory Museum, London (
<collectionCode id="58E1AE3AFF872A2EFE12DF8FFDDDF91A" box="[469,561,1667,1691]" pageId="8" pageNumber="7">BMNH</collectionCode>
).
<typeStatus id="E14B885DFF872A2EFD85DF88FD51F91D" box="[578,701,1668,1692]" pageId="8" pageNumber="7" type="paratype">Paratypes</typeStatus>
depos-
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DFAFFCFCF93A" blockId="8.[192,1056,1187,1787]" box="[192,784,1699,1723]" pageId="8" pageNumber="7">
ited in the
<collectingRegion id="FC34F81DFF872A2EFEF8DFAFFE33F93A" box="[319,479,1699,1723]" country="United States of America" name="Florida" pageId="8" pageNumber="7">Florida State</collectingRegion>
Collection of Arthropods,
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DFCFFCFCF95A" blockId="8.[192,1056,1187,1787]" box="[192,784,1731,1755]" pageId="8" pageNumber="7">
Gainesville, FL (
<collectionCode id="58E1AE3AFF872A2EFE42DFCFFE23F95A" box="[389,463,1731,1755]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="8" pageNumber="7" type="Museum">FSCA</collectionCode>
) and the B. C. Ratcliffe col-
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DFEFFD84F97A" blockId="8.[192,1056,1187,1787]" box="[192,616,1763,1787]" pageId="8" pageNumber="7">
lection, Lincoln,
<collectingRegion id="FC34F81DFF872A2EFE41DFEFFE11F97A" box="[390,509,1763,1787]" country="United States of America" name="Nebraska" pageId="8" pageNumber="7">Nebraska</collectingRegion>
(
<collectionCode id="58E1AE3AFF872A2EFDCCDFEFFDB1F97A" box="[523,605,1763,1787]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="8" pageNumber="7">BCRC</collectionCode>
).
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DE2FFCFCF8BA" blockId="8.[192,1440,1827,1916]" box="[192,784,1827,1851]" pageId="8" pageNumber="7">
<emphasis id="0C84EAEDFF872A2EFF07DE2FFEADF8BA" bold="true" box="[192,321,1827,1851]" pageId="8" pageNumber="7">
<typeStatus id="E14B885DFF872A2EFF07DE2FFED0F8BA" box="[192,316,1827,1851]" pageId="8" pageNumber="7" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Length
<quantity id="F9089B1AFF872A2EFE2FDE2FFDBCF8BA" box="[488,592,1827,1851]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="8" pageNumber="7" unit="mm" value="22.1">22.1 mm</quantity>
; width
<quantity id="F9089B1AFF872A2EFD64DE2FFCE0F8BA" box="[675,780,1827,1851]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="8" pageNumber="7" unit="mm" value="12.1">12.1 mm</quantity>
.
</paragraph>
<paragraph id="3E4F36FFFF872A2EFF07DE4FFB15F8FD" blockId="8.[192,1440,1827,1916]" pageId="8" pageNumber="7">
Color black.
<emphasis id="0C84EAEDFF872A2EFE9EDE4EFE45F8DA" bold="true" box="[345,425,1858,1883]" pageId="8" pageNumber="7">
<emphasis id="0C84EAEDFF872A2EFE9EDE4EFE4EF8DA" bold="true" box="[345,418,1858,1883]" italics="true" pageId="8" pageNumber="7">Head</emphasis>
:
</emphasis>
Frons coarsely rugose, rugae
<emphasis id="0C84EAEDFF872A2EFCF7DE44FC27F8DF" bold="true" box="[816,971,1864,1886]" pageId="8" pageNumber="7">Figure 10-12.</emphasis>
<taxonomicName id="F9F04D7CFF872A2EFC17DE46FA86F8DF" authority="Ratcliffe and Cave" authorityName="Ratcliffe and Cave" box="[976,1386,1865,1887]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="adoceteus">
<emphasis id="0C84EAEDFF872A2EFC17DE46FB73F8DF" box="[976,1183,1865,1887]" italics="true" pageId="8" pageNumber="7">Tomarus adoceteus</emphasis>
Ratcliffe and Cave
</taxonomicName>
, new transversely curved; vertex at center with small, species.
<emphasis id="0C84EAEDFF872A2EFC49DE69FC5FF8FA" bold="true" box="[910,947,1893,1915]" pageId="8" pageNumber="7">10)</emphasis>
Habitus.
<emphasis id="0C84EAEDFF872A2EFBE2DE69FB82F8FA" bold="true" box="[1061,1134,1893,1915]" pageId="8" pageNumber="7">11-12)</emphasis>
Parameres.
</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07D9CFFCFCFF5A" blockId="9.[192,1393,195,1308]" box="[192,784,195,219]" pageId="9" pageNumber="8">subtriangular, smooth, shining area. Frontoclypeal</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07D9EFFCFCFF7A" blockId="9.[192,1393,195,1308]" box="[192,784,227,251]" pageId="9" pageNumber="8">region with low, transverse tubercle either side of</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07D80FFCFCFE9A" blockId="9.[192,1393,195,1308]" box="[192,784,259,283]" pageId="9" pageNumber="8">middle, tubercles broadly separated, laterad of</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07D82FFCFCFEBA" blockId="9.[192,1393,195,1308]" box="[192,784,291,315]" pageId="9" pageNumber="8">clypeal teeth. Clypeus with surface sparsely, trans-</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07D84FFCFCFEDA" blockId="9.[192,1393,195,1308]" box="[192,784,323,348]" pageId="9" pageNumber="8">versely rugose; apex broadly truncate, with 2 small</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07D86FFCFCFEFA" blockId="9.[192,1393,195,1308]" box="[192,784,355,379]" pageId="9" pageNumber="8">teeth, teeth weakly reflexed. Interocular width</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07D88FFCFCFE1A" blockId="9.[192,1393,195,1308]" box="[192,784,387,412]" pageId="9" pageNumber="8">equals 3.5 transverse eye diameters. Antenna 10-</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07D8AFFCFCFE3D" blockId="9.[192,1393,195,1308]" box="[192,784,419,444]" pageId="9" pageNumber="8">segmented, club subequal in length to segments</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07D8CFFCFCFE5A" blockId="9.[192,1393,195,1308]" box="[192,784,451,475]" pageId="9" pageNumber="8">2-7. Mandibles with 2 apical teeth and a rounded</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07D8EFFCFCFE7A" blockId="9.[192,1393,195,1308]" box="[192,784,483,507]" pageId="9" pageNumber="8">
lobe at base.
<emphasis id="0C84EAEDFF862A2FFEABD8E8FE11FE7A" bold="true" box="[364,509,483,507]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFEABD8E8FE1AFE7A" bold="true" box="[364,502,484,507]" italics="true" pageId="9" pageNumber="8">Pronotum</emphasis>
:
</emphasis>
Surface on disc with
</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DB08FCFCFD9A" blockId="9.[192,1393,195,1308]" box="[192,784,515,539]" pageId="9" pageNumber="8">sparse, small punctures; punctures becoming mod-</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DB28FCFCFDBA" blockId="9.[192,1393,195,1308]" box="[192,784,547,571]" pageId="9" pageNumber="8">erate in size and density on sides and anterior</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DB4FFCFCFDDA" blockId="9.[192,1393,195,1308]" box="[192,784,579,603]" pageId="9" pageNumber="8">fourth. Anterior margin with small, median tu-</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DB6FFCFCFDFA" blockId="9.[192,1393,195,1308]" box="[192,784,611,635]" pageId="9" pageNumber="8">bercle. A small, longitudinally oblong fovea present</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DB8FFCFCFD1A" blockId="9.[192,1393,195,1308]" box="[192,784,643,667]" pageId="9" pageNumber="8">behind tubercle, fovea about as wide as distance</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DBAFFCFCFD3A" blockId="9.[192,1393,195,1308]" box="[192,784,675,699]" pageId="9" pageNumber="8">between frontoclypeal tubercles, surface within</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DBCFFCFCFD5A" blockId="9.[192,1393,195,1308]" box="[192,784,706,732]" pageId="9" pageNumber="8">
fovea rugose. Base lacking marginal bead.
<emphasis id="0C84EAEDFF862A2FFD73DBCEFCFCFD5A" bold="true" box="[692,784,706,731]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFD73DBCEFCE4FD5A" bold="true" box="[692,776,706,731]" italics="true" pageId="9" pageNumber="8">Elytra</emphasis>
:
</emphasis>
</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DBEFFCFCFD7A" blockId="9.[192,1393,195,1308]" box="[192,784,739,763]" pageId="9" pageNumber="8">Surface shagreened, with moderate to large, weakly</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DA0FFCFCFC9A" blockId="9.[192,1393,195,1308]" box="[192,784,771,795]" pageId="9" pageNumber="8">ocellate punctures, punctures becoming larger lat-</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DA2FFCFCFCBA" blockId="9.[192,1393,195,1308]" box="[192,784,803,827]" pageId="9" pageNumber="8">erally; 3 pairs of double rows of punctures poorly</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DA4FFCFCFCDA" blockId="9.[192,1393,195,1308]" box="[192,784,834,859]" pageId="9" pageNumber="8">
defined.
<taxonomicName id="F9F04D7CFF862A2FFEE3DA4EFDE0FCDA" authority=": Surface" authorityName="Surface" box="[292,524,834,859]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="0C84EAEDFF862A2FFEE3DA4EFE47FCDA" bold="true" box="[292,427,834,859]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFEE3DA4EFE48FCDA" bold="true" box="[292,420,834,859]" italics="true" pageId="9" pageNumber="8">Pygidium</emphasis>
:
</emphasis>
Surface
</taxonomicName>
on center third nearly
</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DA6FFCFCFCFA" blockId="9.[192,1393,195,1308]" box="[192,784,867,891]" pageId="9" pageNumber="8">smooth, with small, sparse punctures; surface else-</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DA8FFCFCFC1A" blockId="9.[192,1393,195,1308]" box="[192,784,899,924]" pageId="9" pageNumber="8">where densely rugopunctate. In lateral view, sur-</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DAAFFCFCFC3A" blockId="9.[192,1393,195,1308]" box="[192,784,931,955]" pageId="9" pageNumber="8">
face convex.
<emphasis id="0C84EAEDFF862A2FFE93DAA8FE74FC3A" bold="true" box="[340,408,931,955]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFE93DAA8FE7DFC3A" bold="true" box="[340,401,932,955]" italics="true" pageId="9" pageNumber="8">Legs</emphasis>
:
</emphasis>
Protibia tridentate, basal tooth
</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DACFFCFCFC5A" blockId="9.[192,1393,195,1308]" box="[192,784,963,987]" pageId="9" pageNumber="8">slightly removed from others. Protarsus simple,</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DAE8FCFCFC7A" blockId="9.[192,1393,195,1308]" box="[192,784,995,1019]" pageId="9" pageNumber="8">not enlarged. Apex of metatibia subtruncate, with</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DD0FFCFCFB9A" blockId="9.[192,1393,195,1308]" box="[192,784,1027,1051]" pageId="9" pageNumber="8">10 stout, short spinules (some broken off, sockets</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DD2FFCFCFBBA" blockId="9.[192,1393,195,1308]" box="[192,784,1059,1083]" pageId="9" pageNumber="8">visible). Apex of first tarsomere of metatarsus</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DD4FFCFCFBDA" blockId="9.[192,1393,195,1308]" box="[192,784,1091,1115]" pageId="9" pageNumber="8">
subtruncate.
<emphasis id="0C84EAEDFF862A2FFEACDD48FE22FBDA" bold="true" box="[363,462,1091,1115]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFEACDD48FE2BFBDA" bold="true" box="[363,455,1092,1115]" italics="true" pageId="9" pageNumber="8">Venter</emphasis>
:
</emphasis>
Prosternal process long,
</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DD6FFCFCFBFA" blockId="9.[192,1393,195,1308]" box="[192,784,1123,1147]" pageId="9" pageNumber="8">thick, columnar, setigerous at apex, apex longitu-</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DD8FFCFCFB1A" blockId="9.[192,1393,195,1308]" box="[192,784,1155,1179]" pageId="9" pageNumber="8">dinally oval; setae long, reddish brown.</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DDA8FCFCFB3A" blockId="9.[192,1393,195,1308]" box="[192,784,1187,1211]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFF07DDA8FEB1FB3A" bold="true" box="[192,349,1187,1211]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFF07DDA8FEBAFB3A" bold="true" box="[192,342,1188,1211]" italics="true" pageId="9" pageNumber="8">Parameres</emphasis>
:
</emphasis>
Fig. 11-12. The combination of nar-
</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DDC8FCFCFB7A" blockId="9.[192,1393,195,1308]" pageId="9" pageNumber="8">
row form, a long tooth projecting laterally at the
<emphasis id="0C84EAEDFF862A2FFCF7DDC2FC40FB65" bold="true" box="[816,940,1230,1252]" pageId="9" pageNumber="8">Figure 13.</emphasis>
<taxonomicName id="F9F04D7CFF862A2FFC75DDDCFAE7FB64" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[946,1291,1231,1253]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="cuniculus">
<emphasis id="0C84EAEDFF862A2FFC75DDDCFB64FB65" box="[946,1160,1231,1253]" italics="true" pageId="9" pageNumber="8">Tomarus cuniculus</emphasis>
(Fabricius)
</taxonomicName>
, habitus. midpoint of the shaft, and a narrowly elongate apex
</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DC0FFD8EFA9A" blockId="9.[192,1393,195,1308]" box="[192,610,1283,1308]" pageId="9" pageNumber="8">
is unique among
<taxonomicName id="F9F04D7CFF862A2FFE49DC08FD8EFA9A" box="[398,610,1283,1307]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="0C84EAEDFF862A2FFE49DC08FE11FA9A" box="[398,509,1284,1307]" italics="true" pageId="9" pageNumber="8">Tomarus</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="3E4F36FFFF862A2FFF07DC4FFCA0FA5A" blockId="9.[192,1441,1347,1499]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFF07DC4FFED1FADA" bold="true" box="[192,317,1347,1371]" pageId="9" pageNumber="8">
<typeStatus id="E14B885DFF862A2FFF07DC4FFEDBFADA" box="[192,311,1347,1371]" pageId="9" pageNumber="8" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Length
<quantity id="F9089B1AFF862A2FFDCBDC4FFD96FADA" box="[524,634,1347,1371]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="9" pageNumber="8" unit="mm" value="21.7">21.7 mm</quantity>
; width
<quantity id="F9089B1AFF862A2FFD15DC4FFCACFADA" box="[722,832,1347,1371]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="9" pageNumber="8" unit="mm" value="11.7">11.7 mm</quantity>
. Color dark reddish brown. As
<typeStatus id="E14B885DFF862A2FFB7DDC4FFACBFADA" box="[1210,1319,1347,1371]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
, except in the following respects:
<emphasis id="0C84EAEDFF862A2FFE27DC6EFDDDFAFA" bold="true" box="[480,561,1378,1403]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFE27DC6EFDC5FAFA" bold="true" box="[480,553,1378,1403]" italics="true" pageId="9" pageNumber="8">Head</emphasis>
:
</emphasis>
Entire surface transversely rugose. Occiput at center lacking smooth, shiny area.
<emphasis id="0C84EAEDFF862A2FFE8ADC88FE31FA1A" bold="true" box="[333,477,1411,1435]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFE8ADC88FE3AFA1A" bold="true" box="[333,470,1412,1435]" italics="true" pageId="9" pageNumber="8">Pronotum</emphasis>
:
</emphasis>
Fovea reduced to a shallow, short, longitudinal depression.
<taxonomicName id="F9F04D7CFF862A2FFB76DC8EFA4CFA1A" authority=": Surface" authorityName="Surface" box="[1201,1440,1410,1435]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="0C84EAEDFF862A2FFB76DC8EFAD0FA1A" bold="true" box="[1201,1340,1410,1435]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFB76DC8EFAD9FA1A" bold="true" box="[1201,1333,1410,1435]" italics="true" pageId="9" pageNumber="8">Pygidium</emphasis>
:
</emphasis>
Surface
</taxonomicName>
on center third with sparse, large punctures. In lateral view, surface nearly flat.
<emphasis id="0C84EAEDFF862A2FFB69DCA8FB1FFA3A" bold="true" box="[1198,1267,1443,1467]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFB69DCA8FB00FA3A" bold="true" box="[1198,1260,1444,1467]" italics="true" pageId="9" pageNumber="8">Legs</emphasis>
:
</emphasis>
Protibia with trace of fourth, basal tooth. Left posterior leg missing.
</paragraph>
</subSubSection>
<subSubSection id="76EA6574FF862A2FFF07DF0FFDDEF9FA" pageId="9" pageNumber="8" type="description">
<paragraph id="3E4F36FFFF862A2FFF07DF0FFDD5F9BA" blockId="9.[192,1440,1539,1659]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFF07DF0FFEA1F99A" bold="true" box="[192,333,1539,1563]" pageId="9" pageNumber="8">Variation.</emphasis>
Males (
<specimenCount id="28F6FD76FF862A2FFE74DF0FFDD7F99A" box="[435,571,1539,1563]" pageId="9" pageNumber="8" type="generic" typeStatus="paratype">1 paratype</specimenCount>
). Length
<quantity id="F9089B1AFF862A2FFD76DF0FFCCDF99A" box="[689,801,1539,1563]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="9" pageNumber="8" unit="mm" value="22.5">22.5 mm</quantity>
; width
<quantity id="F9089B1AFF862A2FFCBADF0FFC01F99A" box="[893,1005,1539,1563]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="9" pageNumber="8" unit="mm" value="12.4">12.4 mm</quantity>
. The male
<typeStatus id="E14B885DFF862A2FFBB1DF08FB09F99D" box="[1142,1253,1540,1564]" pageId="9" pageNumber="8" type="paratype">paratype</typeStatus>
does not differ significantly from the
<typeStatus id="E14B885DFF862A2FFE0EDF2FFDD9F9BA" box="[457,565,1571,1595]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="3E4F36FFFF862A2FFF37DF4FFDDEF9FA" blockId="9.[192,1440,1539,1659]" pageId="9" pageNumber="8">
Females (
<specimenCount id="28F6FD76FF862A2FFEA3DF4FFE18F9DA" box="[356,500,1603,1627]" pageId="9" pageNumber="8" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
). Length 20.0-
<quantity id="F9089B1AFF862A2FFD5BDF4FFCEAF9DA" box="[668,774,1603,1627]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="9" pageNumber="8" unit="mm" value="22.4">22.4 mm</quantity>
; width
<quantity id="F9089B1AFF862A2FFC9CDF4FFBEDF9DA" box="[859,1025,1603,1627]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" metricValueMax="1.22" metricValueMin="1.03" pageId="9" pageNumber="8" unit="mm" value="11.25" valueMax="12.2" valueMin="10.3">10.3-12.2 mm</quantity>
. The female
<typeStatus id="E14B885DFF862A2FFB54DF48FAE7F9DD" box="[1171,1291,1604,1628]" pageId="9" pageNumber="8" type="paratype">paratypes</typeStatus>
do not differ significantly from the
<typeStatus id="E14B885DFF862A2FFE0EDF6FFDC2F9FA" box="[457,558,1635,1659]" pageId="9" pageNumber="8" type="allotype">allotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="76EA6574FF862A2FFF07DFAFFD5AF95A" pageId="9" pageNumber="8" type="etymology">
<paragraph id="3E4F36FFFF862A2FFF07DFAFFD5AF95A" blockId="9.[192,1440,1699,1755]" pageId="9" pageNumber="8">
<emphasis id="0C84EAEDFF862A2FFF07DFAFFEB1F93A" bold="true" box="[192,349,1699,1723]" pageId="9" pageNumber="8">Etymology.</emphasis>
From the Greek
<emphasis id="0C84EAEDFF862A2FFDEDDFAFFD7BF93A" box="[554,663,1699,1723]" italics="true" pageId="9" pageNumber="8">adoketus</emphasis>
, meaning surprising or unusual. Used here as a noun in apposition, hence,
<taxonomicName id="F9F04D7CFF862A2FFE94DFC8FE2EF95A" authorityName="Erichson" authorityYear="1847" box="[339,450,1732,1755]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="0C84EAEDFF862A2FFE94DFC8FE2EF95A" box="[339,450,1732,1755]" italics="true" pageId="9" pageNumber="8">Tomarus</emphasis>
</taxonomicName>
, the surprising one.
</paragraph>
</subSubSection>
</treatment>
</document>
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