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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.886.31163" ID-GBIF-Dataset="ddbbd7eb-5dd3-4d91-a5b0-c8ccbc76fe05" ID-PMC="PMC6851204" ID-Pensoft-Pub="1313-2970-886-61" ID-Pensoft-UUID="E3765D62384D5A6B9CB55987C04EA58E" ID-PubMed="31736622" ID-ZooBank="512BB3C93B884A34977061F3845770BA" ModsDocID="1313-2970-886-61" checkinTime="1573015829836" checkinUser="pensoft" docAuthor="Liu, Ping, Irfan, Muhammad, Yang, Su-Fang &amp; Peng, Xian-Jin" docDate="2019" docId="B600173E8375588CBED9767C047BE915" docLanguage="en" docName="ZooKeys 886: 61-77" docOrigin="ZooKeys 886" docSource="http://dx.doi.org/10.3897/zookeys.886.31163" docTitle="Araneus conexus Liu, Irfan, Yang &amp; Peng, 2019, sp. nov." docType="treatment" docUuid="08B788A4-1CEF-4673-9BCE-56E4CA49A456" docUuidSource="ZooBank" docVersion="5" id="E3765D62384D5A6B9CB55987C04EA58E" lastPageNumber="61" masterDocId="E3765D62384D5A6B9CB55987C04EA58E" masterDocTitle="Two new species of Araneus Clerck, 1757 (Araneae, Araneidae) and first description of A. wulongensis male from China" masterLastPageNumber="77" masterPageNumber="61" pageNumber="61" updateTime="1668168025371" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of Araneus Clerck, 1757 (Araneae, Araneidae) and first description of A. wulongensis male from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Ping</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Irfan, Muhammad</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Su-Fang</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Peng, Xian-Jin</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>886</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>61</mods:start>
<mods:end>77</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.886.31163</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.886.31163</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-886-61</mods:identifier>
<mods:identifier type="ZooBank">512BB3C93B884A34977061F3845770BA</mods:identifier>
<mods:identifier type="Pensoft-UUID">E3765D62384D5A6B9CB55987C04EA58E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="160140627" LSID="urn:lsid:zoobank.org:act:08B788A4-1CEF-4673-9BCE-56E4CA49A456" httpUri="http://treatment.plazi.org/id/B600173E8375588CBED9767C047BE915" lastPageNumber="61" pageId="0" pageNumber="61">
<subSubSection pageId="0" pageNumber="61" type="nomenclature">
<paragraph pageId="0" pageNumber="61">
<taxonomicName LSID="B600173E-8375-588C-BED9-767C047BE915" class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araneus conexus" order="Araneae" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="conexus">Araneus conexus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="61">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Araneus conexus sp. nov., male (holotype, A-C); female (paratype collected together with the holotype, D-G). A, D habitus, dorsal view B, F habitus, ventral view E habitus, lateral view C, G leg I and II, prolateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.886.31163.figure1" httpUri="https://binary.pensoft.net/fig/352762" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure1">Figs 1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Araneus conexus sp. nov., male holotype palp A, D prolateral view B, E ventral view C, F median apophysis. Abbreviations: C = conductor; E = embolus; MA = median apophysis; STA = subterminal apophysis; TA = terminal apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.31163.figure2" httpUri="https://binary.pensoft.net/fig/352763" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure2">2</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Araneus conexus sp. nov., male (paratype palp expanded A, B; endaparatus C-E) A-C prolateral view D lateral view E ventral view. Abbreviations: C = conductor; E = embolus; EL = embolic lamella; MA = median apophysis; STA = subterminal apophysis; TA = terminal apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.31163.figure3" httpUri="https://binary.pensoft.net/fig/352764" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure3">3</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Araneus conexus sp. nov., female (epigyne, A-D) A, B ventral view C, D posterior view. Abbreviations: CO = copulatory opening; FD = fertilization duct; LP = lateral plate; MP = median plate; S = scape; SP = spermatheca. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.31163.figure4" httpUri="https://binary.pensoft.net/fig/352765" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure4">4</figureCitation>
,
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Araneus conexus sp. nov., male (paratype, A, C GKJ 026); female (paratype, B, D GKJ 026) A, B habitus, dorsal view C palp, prolaterior view D epigyne, ventral view. Scale bars: 0.5 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zookeys.886.31163.figure5" httpUri="https://binary.pensoft.net/fig/352766" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure5">5</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Collection localities for Araneus conexus sp. nov., Araneus digitatus sp. nov., and Araneus wulongensis Song &amp; Zhu, 1992 in China." figureDoi="10.3897/zookeys.886.31163.figure12" httpUri="https://binary.pensoft.net/fig/352773" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure12">12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="type material">
<paragraph pageId="0" pageNumber="61">Type material.</paragraph>
<paragraph pageId="0" pageNumber="61">
Holotype, ♂,
<emphasis bold="true" pageId="0" pageNumber="61">China, Yunnan Province</emphasis>
: Tengchong County, Jietou Township, Datang Village: Longtang River, papaya orchard,
<geoCoordinate degrees="25.75720" direction="north" orientation="latitude" precision="1" value="25.7572">25.75720N</geoCoordinate>
,
<geoCoordinate degrees="98.69459" direction="east" orientation="longitude" precision="1" value="98.69459">98.69459E</geoCoordinate>
, 2078 m, 16.05.2006, X. J. Peng, X.P. Wang and P. Hu leg. (Peng060516). Paratypes: 1♂ 2♀, same data as holotype (Peng060516); 4♂ 3♀, Dahe Ridge,
<geoCoordinate degrees="25.42018" direction="north" orientation="latitude" precision="1" value="25.42018">25.42018N</geoCoordinate>
,
<geoCoordinate degrees="98.40946" direction="east" orientation="longitude" precision="1" value="98.40946">98.40946E</geoCoordinate>
, 1878 m, 19.05.2006, X.J. Peng, X.P. Wang and P. Hu leg. (Peng060519); 1♂ 2♀, Longling County, Longjiang Township, Xiaoheishan Nature Reserve,
<geoCoordinate degrees="24.82886" direction="north" orientation="latitude" precision="1" value="24.82886">24.82886N</geoCoordinate>
,
<geoCoordinate degrees="98.75917" direction="east" orientation="longitude" precision="1" value="98.75917">98.75917E</geoCoordinate>
, 2010 m, 26.05.2005, H.M. Yan leg. (GKJ026); 2♀, Longyang District, Bawan Village, Nankang Valley,
<geoCoordinate degrees="24.82587" direction="north" orientation="latitude" precision="1" value="24.82587">24.82587N</geoCoordinate>
,
<geoCoordinate degrees="98.76832" direction="east" orientation="longitude" precision="1" value="98.76832">98.76832E</geoCoordinate>
, 2148 m, 26.05.2005, K.J. Guo leg. (GKJ027).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="etymology">
<paragraph pageId="0" pageNumber="61">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="61">
The specific name from Latin adjective
<emphasis italics="true" pageId="0" pageNumber="61">conexus</emphasis>
(joined together), referring to the abdominal humps joined together in female.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="diagnosis">
<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="61">
The new species resembles
<taxonomicName lsidName="A. stella" pageId="0" pageNumber="61" rank="species" species="stella">
<emphasis italics="true" pageId="0" pageNumber="61">A. stella</emphasis>
</taxonomicName>
(Karsch, 1879) (
<bibRefCitation author="Tanikawa, A" editor="Ono, H" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="61" refId="B8" refString="Tanikawa, A, Ono, H, 2009. Hersiliidae. Nephilidae, Tetragnathidae, Araneidae. In: OnoH (Ed.) The spiders of Japan with Keys to the Families and Genera and Illustrations of the Species.Tokai University Press, Kanagawa, 149, 403-463." title="Hersiliidae. Nephilidae, Tetragnathidae, Araneidae. In: OnoH (Ed.) The spiders of Japan with Keys to the Families and Genera and Illustrations of the Species. Tokai University Press, Kanagawa, 149, 403 - 463." year="2009">Tanikawa 2009</bibRefCitation>
: figs 218, 219;
<bibRefCitation author="Kim, ST" journalOrPublisher="Invertebrate Fauna of Korea" pageId="0" pageNumber="61" pagination="1 - 146" refId="B2" refString="Kim, ST, Lee, SY, 2012. Arthropoda: Arachnida: Araneae: Araneidae. Araneid spiders. Invertebrate Fauna of Korea 21: 1 - 146" title="Arthropoda: Arachnida: Araneae: Araneidae. Araneid spiders." volume="21" year="2012">Kim and Lee 2012</bibRefCitation>
: fig. 18), but can be distinguished by: 1) median apophysis with a spur and 4-6 teeth in
<taxonomicName lsidName="A. conexus" pageId="0" pageNumber="61" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">A. conexus</emphasis>
</taxonomicName>
sp. nov. vs with 2 large teeth in
<taxonomicName lsidName="A. stella" pageId="0" pageNumber="61" rank="species" species="stella">
<emphasis italics="true" pageId="0" pageNumber="61">A. stella</emphasis>
</taxonomicName>
; 2) subterminal apophysis longer than wide, with blunt tip in
<taxonomicName lsidName="A. conexus" pageId="0" pageNumber="61" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">A. conexus</emphasis>
</taxonomicName>
sp. nov. vs almost as long as wide, with pointed tip in
<taxonomicName lsidName="A. stella" pageId="0" pageNumber="61" rank="species" species="stella">
<emphasis italics="true" pageId="0" pageNumber="61">A. stella</emphasis>
</taxonomicName>
; 3) terminal apophysis almost parallel to the embolus and subterminal apophysis in prolateral view in
<taxonomicName lsidName="A. conexus" pageId="0" pageNumber="61" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">A. conexus</emphasis>
</taxonomicName>
sp. nov. vs almost perpendicular, tip overlapping the conductor in
<taxonomicName lsidName="A. stella" pageId="0" pageNumber="61" rank="species" species="stella">
<emphasis italics="true" pageId="0" pageNumber="61">A. stella</emphasis>
</taxonomicName>
; 4) distal part of embolus hook-shaped in
<taxonomicName lsidName="A. conexus" pageId="0" pageNumber="61" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">A. conexus</emphasis>
</taxonomicName>
sp. nov. vs straight in
<taxonomicName lsidName="A. stella" pageId="0" pageNumber="61" rank="species" species="stella">
<emphasis italics="true" pageId="0" pageNumber="61">A. stella</emphasis>
</taxonomicName>
; 5) scape almost as long as epigyne in
<taxonomicName lsidName="A. conexus" pageId="0" pageNumber="61" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">A. conexus</emphasis>
</taxonomicName>
sp. nov. vs about 2 times longer than epigyne in
<taxonomicName lsidName="A. stella" pageId="0" pageNumber="61" rank="species" species="stella">
<emphasis italics="true" pageId="0" pageNumber="61">A. stella</emphasis>
</taxonomicName>
; 6) abdomen with a pair of humps at the anterior part, almost merged with each other in
<taxonomicName lsidName="A. conexus" pageId="0" pageNumber="61" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">A. conexus</emphasis>
</taxonomicName>
sp. nov. vs humps not merged and located at the lateral sides in
<taxonomicName lsidName="A. stella" pageId="0" pageNumber="61" rank="species" species="stella">
<emphasis italics="true" pageId="0" pageNumber="61">A. stella</emphasis>
</taxonomicName>
. The female of new species resembles
<taxonomicName lsidName="A. bicavus" pageId="0" pageNumber="61" rank="species" species="bicavus">
<emphasis italics="true" pageId="0" pageNumber="61">A. bicavus</emphasis>
</taxonomicName>
Zhu &amp; Wang, 1994 (
<bibRefCitation author="Yin, CM" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="61" refId="B11" refString="Yin, CM, Wang, JF, Zhu, MS, Xie, LP, Peng, XJ, Bao, YH, 1997. Fauna Sinica: Arachnida: Araneae: Araneidae. Science Press, Beijing" title="Fauna Sinica: Arachnida: Araneae: Araneidae." year="1997">Yin et al. 1997</bibRefCitation>
: fig. 77), but can be distinguished by: 1) the abdomen with a pair of large humps at the anterior part, almost merged with each other in
<taxonomicName lsidName="A. conexus" pageId="0" pageNumber="61" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">A. conexus</emphasis>
</taxonomicName>
sp. nov. vs small humps not merged and located at the lateral sides in
<taxonomicName lsidName="A. bicavus" pageId="0" pageNumber="61" rank="species" species="bicavus">
<emphasis italics="true" pageId="0" pageNumber="61">A. bicavus</emphasis>
</taxonomicName>
; 2) the epigyne with a depression positioned vertically on each side of the scape in
<taxonomicName lsidName="A. conexus" pageId="0" pageNumber="61" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">A. conexus</emphasis>
</taxonomicName>
sp. nov. vs with a circular depression in
<taxonomicName lsidName="A. bicavus" pageId="0" pageNumber="61" rank="species" species="bicavus">
<emphasis italics="true" pageId="0" pageNumber="61">A. bicavus</emphasis>
</taxonomicName>
. The female also resembles
<taxonomicName lsidName="A. boesenbergi" pageId="0" pageNumber="61" rank="species" species="boesenbergi">
<emphasis italics="true" pageId="0" pageNumber="61">A. boesenbergi</emphasis>
</taxonomicName>
(Fox, 1938) (
<bibRefCitation author="Yin, CM" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="61" refId="B11" refString="Yin, CM, Wang, JF, Zhu, MS, Xie, LP, Peng, XJ, Bao, YH, 1997. Fauna Sinica: Arachnida: Araneae: Araneidae. Science Press, Beijing" title="Fauna Sinica: Arachnida: Araneae: Araneidae." year="1997">Yin et al. 1997</bibRefCitation>
: fig. 79), but can be distinguished by: furrow of epigynal plate semicircular in
<taxonomicName lsidName="A. conexus" pageId="0" pageNumber="61" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">A. conexus</emphasis>
</taxonomicName>
sp. nov. vs almost circular in
<taxonomicName lsidName="A. boesenbergi" pageId="0" pageNumber="61" rank="species" species="boesenbergi">
<emphasis italics="true" pageId="0" pageNumber="61">A. boesenbergi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">Description.</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Male</emphasis>
(holotype) (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Araneus conexus sp. nov., male (holotype, A-C); female (paratype collected together with the holotype, D-G). A, D habitus, dorsal view B, F habitus, ventral view E habitus, lateral view C, G leg I and II, prolateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.886.31163.figure1" httpUri="https://binary.pensoft.net/fig/352762" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure1">
Fig. 1
<normalizedToken originalValue="A–C">A-C</normalizedToken>
</figureCitation>
): total length 3.04. Carapace 1.62 long, 1.30 wide, yellow; fovea, cervical, and radial grooves distinct. Eye sizes and interdistances: AME 0.08; ALE 0.09; PME 0.10; PLE 0.08;
<normalizedToken originalValue="AME–AME">AME-AME</normalizedToken>
0.06;
<normalizedToken originalValue="AME–ALE">AME-ALE</normalizedToken>
0.23;
<normalizedToken originalValue="PME–PME">PME-PME</normalizedToken>
0.14;
<normalizedToken originalValue="PME–PLE">PME-PLE</normalizedToken>
0.24; MOL 0.24; MOA 0.23; MOP 0.27. Sternum yellowish brown, with transverse light band anteriorly. Chelicerae yellow. Endites yellow, distal end pale. Labium brown, distal part pale yellow. Legs yellow, with dark-brown annuli. Tibia I slightly curved, with several strong spines: 3d, 1v, 7p, 1r; tibia II spines: 3d, 3v, 6p, 1r. Leg lengths: I, 6.54 (1.86, 2.25, 1.51, 0.92); II, 5.21 (1.56, 1.77, 1.28, 0.60); III, 3.04 (1.03, 1.02, 0.60, 0.39); IV, 4.22 (1.33, 1.44, 0.98, 0.47). Abdomen 2.63 long, 1.30 wide, oval, dorsum grayish yellow, anterior part dark and slightly bulged in the middle, posterior part with 4 dark transverse bands, 4 pairs of sigillae; ventral side with a longitudinal brown band, lateral sides grayish yellow. Spinnerets yellowish brown.
</paragraph>
<caption doi="10.3897/zookeys.886.31163.figure1" httpUri="https://binary.pensoft.net/fig/352762" pageId="0" pageNumber="61" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 1.</emphasis>
<taxonomicName class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araneus conexus" order="Araneae" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">Araneus conexus</emphasis>
</taxonomicName>
sp. nov., male (holotype,
<emphasis bold="true" pageId="0" pageNumber="61">
<normalizedToken originalValue="A–C">A-C</normalizedToken>
</emphasis>
); female (paratype collected together with the holotype,
<emphasis bold="true" pageId="0" pageNumber="61">
<normalizedToken originalValue="D–G">D-G</normalizedToken>
</emphasis>
).
<emphasis bold="true" pageId="0" pageNumber="61">A, D</emphasis>
habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="61">B, F</emphasis>
habitus, ventral view
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="61">C, G</emphasis>
leg I and II, prolateral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
Palp (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Araneus conexus sp. nov., male holotype palp A, D prolateral view B, E ventral view C, F median apophysis. Abbreviations: C = conductor; E = embolus; MA = median apophysis; STA = subterminal apophysis; TA = terminal apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.31163.figure2" httpUri="https://binary.pensoft.net/fig/352763" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure2">Figs 2</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Araneus conexus sp. nov., male (paratype palp expanded A, B; endaparatus C-E) A-C prolateral view D lateral view E ventral view. Abbreviations: C = conductor; E = embolus; EL = embolic lamella; MA = median apophysis; STA = subterminal apophysis; TA = terminal apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.31163.figure3" httpUri="https://binary.pensoft.net/fig/352764" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure3">3</figureCitation>
). Patella with 2 macrosetae. Tibia wider than long, ventral side bulging in prolateral view. Tegulum slightly grooved ventrally. Median apophysis with a prolateral spur, with 1 large and 3-5 small retrolateral teeth. Conductor membranous, gear lever-shaped, with swollen tip (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Araneus conexus sp. nov., male (paratype palp expanded A, B; endaparatus C-E) A-C prolateral view D lateral view E ventral view. Abbreviations: C = conductor; E = embolus; EL = embolic lamella; MA = median apophysis; STA = subterminal apophysis; TA = terminal apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.31163.figure3" httpUri="https://binary.pensoft.net/fig/352764" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure3">Fig. 3A, B</figureCitation>
). Subterminal apophysis sclerotized, longer than wide, terminal margin with several teeth. Terminal apophysis sword-shaped, with pointed tip. Embolus longer than wide, tip hooked, directed anti-clockwise; embolic lamella membranous, finger-shaped in ventral view, distally grooved (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Araneus conexus sp. nov., male (paratype palp expanded A, B; endaparatus C-E) A-C prolateral view D lateral view E ventral view. Abbreviations: C = conductor; E = embolus; EL = embolic lamella; MA = median apophysis; STA = subterminal apophysis; TA = terminal apophysis. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.31163.figure3" httpUri="https://binary.pensoft.net/fig/352764" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure3">Fig. 3</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.886.31163.figure2" httpUri="https://binary.pensoft.net/fig/352763" pageId="0" pageNumber="61" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 2.</emphasis>
<taxonomicName class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araneus conexus" order="Araneae" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">Araneus conexus</emphasis>
</taxonomicName>
sp. nov., male holotype palp
<emphasis bold="true" pageId="0" pageNumber="61">A, D</emphasis>
prolateral view
<emphasis bold="true" pageId="0" pageNumber="61">B, E</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="61">C, F</emphasis>
median apophysis. Abbreviations: C = conductor; E = embolus; MA = median apophysis; STA = subterminal apophysis; TA = terminal apophysis. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.886.31163.figure3" httpUri="https://binary.pensoft.net/fig/352764" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
<taxonomicName class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araneus conexus" order="Araneae" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">Araneus conexus</emphasis>
</taxonomicName>
sp. nov., male (paratype palp expanded
<emphasis bold="true" pageId="0" pageNumber="61">A, B</emphasis>
; endaparatus
<emphasis bold="true" pageId="0" pageNumber="61">
<normalizedToken originalValue="C–E">C-E</normalizedToken>
</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="61">
<normalizedToken originalValue="A–C">A-C</normalizedToken>
</emphasis>
prolateral view
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
ventral view. Abbreviations: C= conductor; E = embolus; EL = embolic lamella; MA = median apophysis; STA = subterminal apophysis; TA = terminal apophysis. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Female</emphasis>
(allotype) (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Araneus conexus sp. nov., male (holotype, A-C); female (paratype collected together with the holotype, D-G). A, D habitus, dorsal view B, F habitus, ventral view E habitus, lateral view C, G leg I and II, prolateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.886.31163.figure1" httpUri="https://binary.pensoft.net/fig/352762" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure1">
Fig. 1
<normalizedToken originalValue="C–F">C-F</normalizedToken>
</figureCitation>
): total length 4.42. Carapace 1.79 long, 1.65 wide, yellowish brown; cervical and radial groves distinct. Eye sizes and interdistances: AME, 0.07; ALE, 0.08; PME, 0.10; PLE, 0.07;
<normalizedToken originalValue="AME–AME">AME-AME</normalizedToken>
, 0.12;
<normalizedToken originalValue="AME–ALE">AME-ALE</normalizedToken>
, 0.35;
<normalizedToken originalValue="PME–PME">PME-PME</normalizedToken>
, 0.12;
<normalizedToken originalValue="PME–PLE">PME-PLE</normalizedToken>
, 0.29; MOL, 0.26; MOA, 0.26; MOP, 0.30. Sternum dark brown with transverse light band anteriorly. Chelicerae dark brown. Endites brown, distal end pale. Labium brown, distal third part pale yellow. Legs yellow, annuli indistinct. Tibia I straight with few spines: 2d, 2p, 1r; tibia II: 2d, 2r. Leg lengths: I, 5.45 (1.68, 1.99, 1.20, 0.58); II, 4.70 (1.50, 1.65, 1.02, 0.53); III, 2.89 (0.92, 0.98, 0.59, 0.40); IV, 4.26 (1.33, 1.45, 1.00, 0.48). Abdomen 3.70 long, 2.67 wide, with 2 anterior humps, almost merged with each other, paler and all other morphological pattern similar to the male.
</paragraph>
<paragraph pageId="0" pageNumber="61">
Epigyne (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Araneus conexus sp. nov., female (epigyne, A-D) A, B ventral view C, D posterior view. Abbreviations: CO = copulatory opening; FD = fertilization duct; LP = lateral plate; MP = median plate; S = scape; SP = spermatheca. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.886.31163.figure4" httpUri="https://binary.pensoft.net/fig/352765" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure4">Fig. 4</figureCitation>
). Epigyne wider than long; scape almost straight, tip slightly curved ventrally; lateral plates longer than wide, expanded laterally in ventral view; median plate longer than wide, tongue-shaped, grooved posteriorly; basal lamellae absent. Copulatory openings facing ventrally, in the slit between median and lateral plates. Copulatory ducts inconspicuous. Spermathecae oval.
</paragraph>
<caption doi="10.3897/zookeys.886.31163.figure4" httpUri="https://binary.pensoft.net/fig/352765" pageId="0" pageNumber="61" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 4.</emphasis>
<taxonomicName class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araneus conexus" order="Araneae" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">Araneus conexus</emphasis>
</taxonomicName>
sp. nov., female (epigyne,
<emphasis bold="true" pageId="0" pageNumber="61">
<normalizedToken originalValue="A–D">A-D</normalizedToken>
</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="61">A, B</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="61">C, D</emphasis>
posterior view. Abbreviations: CO = copulatory opening; FD = fertilization duct; LP = lateral plate; MP = median plate; S = scape; SP = spermatheca. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Variation</emphasis>
(paratype male and female from GKJ026) (
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Araneus conexus sp. nov., male (paratype, A, C GKJ 026); female (paratype, B, D GKJ 026) A, B habitus, dorsal view C palp, prolaterior view D epigyne, ventral view. Scale bars: 0.5 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zookeys.886.31163.figure5" httpUri="https://binary.pensoft.net/fig/352766" pageId="0" pageNumber="61" tableDoi="10.3897/zookeys.886.31163.figure5">Fig. 5</figureCitation>
): abdomen with a pair of large dark lateral spots, and pair of herringbones spots in the middle area of dorsum. The structure of copulatory organs as in holotype and paratypes of Peng060516.
</paragraph>
<caption doi="10.3897/zookeys.886.31163.figure5" httpUri="https://binary.pensoft.net/fig/352766" pageId="0" pageNumber="61" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 5.</emphasis>
<taxonomicName class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araneus conexus" order="Araneae" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="conexus">
<emphasis italics="true" pageId="0" pageNumber="61">Araneus conexus</emphasis>
</taxonomicName>
sp. nov., male (paratype,
<emphasis bold="true" pageId="0" pageNumber="61">A, C</emphasis>
GKJ026); female (paratype,
<emphasis bold="true" pageId="0" pageNumber="61">B, D</emphasis>
GKJ026)
<emphasis bold="true" pageId="0" pageNumber="61">A, B</emphasis>
habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
palp, prolaterior view
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
epigyne, ventral view. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="61">A, B</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="61">C, D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="distribution">
<paragraph pageId="0" pageNumber="61">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="61">Known only from the type locality.</paragraph>
</subSubSection>
</treatment>
</document>

119
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<mods:classification id="72BDB30E23DE5DAFA1C12667C22AA411">journal article</mods:classification>
<mods:identifier id="9DA04EAD9F3D6D9A4B33EF4A30292F80" type="DOI">http://dx.doi.org/10.3897/zookeys.1193.115831</mods:identifier>
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<treatment id="B6013E519C3C559DB467F2A8DDA955B8" LSID="urn:lsid:zoobank.org:act:7F8C078F-94F4-4D71-BB49-3DB46F8C947E" httpUri="http://treatment.plazi.org/id/B6013E519C3C559DB467F2A8DDA955B8" lastPageNumber="219" pageId="0" pageNumber="219">
<subSubSection id="B494ABB78CC0ACACEF81391D873BED46" pageId="0" pageNumber="219" type="nomenclature">
<paragraph id="EE7AF35424C9DF272482A9F2B8239EB2" pageId="0" pageNumber="219">
<taxonomicName id="70457CA18E108590A598A45C72698C6E" LSID="https://zoobank.org/7F8C078F-94F4-4D71-BB49-3DB46F8C947E" authority="Han &amp; van Achterberg" authorityName="Han &amp; van Achterberg" authorityYear="2024" class="Insecta" family="Braconidae" genus="Xynobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xynobius duoferus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="duoferus" status="sp. nov.">Xynobius duoferus Han &amp; van Achterberg</taxonomicName>
<taxonomicNameLabel id="E86C751921E16E3484D70501D882BB19" pageId="0" pageNumber="219">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="68AD1D95FC44C85E5A59E8306C32B9B2" pageId="0" pageNumber="219" type="description">
<paragraph id="E94C7B6710AF1A738EF60EEB1E5E8D09" pageId="0" pageNumber="219">
<figureCitation id="65518FE272F7CAAA8FE7C4F2F1B0230F" captionStart="Figure 29" captionStartId="F6" captionText="Figure 29. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀, habitus, lateral." figureDoi="10.3897/zookeys.1193.115831.figure29" httpUri="https://binary.pensoft.net/fig/997931" pageId="0" pageNumber="219">Figs 29</figureCitation>
<figureCitation id="AE4521831A2A1DBCC9724CCE3342BC36" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">, 30-41</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FCDC543DC64A832CBF19C9B0EC9548E1" pageId="0" pageNumber="219" type="type material">
<paragraph id="A77CFBC1DEF7787C5223F31BA861A880" pageId="0" pageNumber="219">Type material.</paragraph>
<paragraph id="B34D9DF71703E980299DC463069400D5" pageId="0" pageNumber="219">
<emphasis id="E1558394913D8BDF99415827ED0B4F69" bold="true" pageId="0" pageNumber="219">
<emphasis id="D0B355ACEFEF567A95626863BEE2E850" italics="true" pageId="0" pageNumber="219">Holotype</emphasis>
.
</emphasis>
♀ (NIBR), &quot;South Korea: Jangam Cave, Pyeongchang-gun, Gangwon prov.,
<geoCoordinate id="70BD63C6B82964BE06806E79F7B09E39" degrees="37" direction="north" minutes="23" orientation="latitude" precision="1" seconds="54.2" value="37.39839">37°23'54.2&quot;N</geoCoordinate>
,
<geoCoordinate id="142FFD84EC4451F6FB90EE8A485FCF8A" degrees="128" direction="east" minutes="25" orientation="longitude" precision="1" seconds="24.2" value="128.4234">128°25'24.2&quot;E</geoCoordinate>
, 11.vii.2020, SW [= collected by sweeping], Hyojoong Kim leg., KSNU&quot;.
</paragraph>
<caption id="6C373C29E4082723A2BEEFD4FBA95C16" doi="10.3897/zookeys.1193.115831.figure29" httpUri="https://binary.pensoft.net/fig/997931" pageId="0" pageNumber="219" start="Figure 29" startId="F6">
<paragraph id="2385E9D392755FFF237784D23AF54AF0" pageId="0" pageNumber="219">
<emphasis id="53B768050BB6CC5711290DBAD74DC63F" bold="true" pageId="0" pageNumber="219">Figure 29.</emphasis>
<taxonomicName id="6F5EAB6C08CE3FCF46EFD995C5A878F3" authorityName="Han &amp; van Achterberg" authorityYear="2024" class="Insecta" family="Braconidae" genus="Xynobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xynobius duoferus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="duoferus">
<emphasis id="F2B07CE3639C02070F9089D6F5A6C887" italics="true" pageId="0" pageNumber="219">Xynobius duoferus</emphasis>
</taxonomicName>
Han &amp; van Achterberg, sp. nov., holotype, ♀, habitus, lateral.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B16F46A4AC846CDFC319C4B8E4D013E3" pageId="0" pageNumber="219" type="diagnosis">
<paragraph id="063485AC5E3C75C38616C8B0E46519EB" pageId="0" pageNumber="219">Diagnosis.</paragraph>
<paragraph id="EB49CBEE0F3582D245896A59F44197F3" pageId="0" pageNumber="219">
Dorsope distinct (Fig.
<figureCitation id="A6E085C72EDBA68032FBB15641A0DAFB" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">37</figureCitation>
); first metasomal tergite with straight longitudinal striae; notauli complete and narrowly crenulate (Fig.
<figureCitation id="FF0EF813EF41B5E7932E3C9BA050E7E1" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">32</figureCitation>
); mesoscutum largely smooth and sparsely setose medially; second tergite striate-rugose medially; 20th to 26th antennal segments of ♀ white followed by two dark apical segments.
</paragraph>
<caption id="318A2F98AA14E85FEE83CF94D8373DD7" doi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219" start="Figures 30–41" startId="F7">
<paragraph id="39CDDEF8E65A7E793D25A1FE74ABF89B" pageId="0" pageNumber="219">
<emphasis id="9BEB8D60895C4C6C7C1E025DF8FC7F9B" bold="true" pageId="0" pageNumber="219">Figures 30-41.</emphasis>
<taxonomicName id="8BC846BF9BEF6FEDA511F35FA5A0E4B6" authorityName="Han &amp; van Achterberg" authorityYear="2024" class="Insecta" family="Braconidae" genus="Xynobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xynobius duoferus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="duoferus">
<emphasis id="E4D49312024885A92DDA28D4D6064C86" italics="true" pageId="0" pageNumber="219">Xynobius duoferus</emphasis>
</taxonomicName>
Han &amp; van Achterberg, sp. nov., holotype, ♀
<emphasis id="60A148367BF34E0C80F77FDF004ED2F7" bold="true" pageId="0" pageNumber="219">30</emphasis>
wings
<emphasis id="9C0F67FD61B1D81CF71BD9DD42F5A117" bold="true" pageId="0" pageNumber="219">31</emphasis>
mesosoma, lateral view
<emphasis id="5E5D13520009B6F00BC56A2FDAFA3FBE" bold="true" pageId="0" pageNumber="219">32</emphasis>
mesosoma, dorsal view
<emphasis id="076248704AF09A4B89E201048EFA870D" bold="true" pageId="0" pageNumber="219">33</emphasis>
metasoma, dorsal view
<emphasis id="001E77184E064CB0967F724EF79E2378" bold="true" pageId="0" pageNumber="219">34</emphasis>
head, anterior view
<emphasis id="384C08CCCC0AF096C1612FD918C676FD" bold="true" pageId="0" pageNumber="219">35</emphasis>
head, dorsal view
<emphasis id="955023E3DDC8AF08BD5A84ACEB5DD705" bold="true" pageId="0" pageNumber="219">36</emphasis>
propodeum, dorsal view
<emphasis id="333D413CAD86FD87BA267C31830D906A" bold="true" pageId="0" pageNumber="219">37</emphasis>
1st metasomal tergite, dorsal view
<emphasis id="FCF09B994A33C24401E3A2B1FD2B1061" bold="true" pageId="0" pageNumber="219">38</emphasis>
ovipositor and its sheath, lateral view
<emphasis id="0C5A96EAF3D8A71BEC721833F07FEE46" bold="true" pageId="0" pageNumber="219">39</emphasis>
basal part of antenna
<emphasis id="787EFD15310FDCDA735DE1CBA7C65777" bold="true" pageId="0" pageNumber="219">40</emphasis>
apical part of antenna
<emphasis id="B2D1550D737D69AEB10C00C80B178BC2" bold="true" pageId="0" pageNumber="219">41</emphasis>
antenna.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="59EAA8300F95DEE45E599A7DBE89AE1D" pageId="0" pageNumber="219" type="description">
<paragraph id="73B3C143D22DFB37F1E16678A6BB2AA4" pageId="0" pageNumber="219">Description.</paragraph>
<paragraph id="55944F948C9250675C97AC5CB6275813" pageId="0" pageNumber="219">Female; length of body 2.0 mm, of fore wing 2.5 mm.</paragraph>
<paragraph id="1D46600B863B7BDC7F6C55CAA050C83A" pageId="0" pageNumber="219">
<emphasis id="813F4D31E13AE3725E2EFBFDD24CF021" bold="true" pageId="0" pageNumber="219">
<emphasis id="EBE89E7FA51330D8C3D56407DA276DDC" italics="true" pageId="0" pageNumber="219">Head</emphasis>
.
</emphasis>
Antenna with 28 segments and 1.6
<normalizedToken id="977E1513E5D519BADD70CB1386E4DD15" originalValue="×">x</normalizedToken>
longer than body; third segment of antenna 5.8
<normalizedToken id="9E8E8AAFE71F9700A1E931000ECA1B38" originalValue="×">x</normalizedToken>
longer than wide and 1.1
<normalizedToken id="2D32053E64AF2B7121DF1A0974ED007A" originalValue="×">x</normalizedToken>
longer than fourth segment (Figs
<figureCitation id="49174E715197F70884EF8FDA39213742" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">39</figureCitation>
,
<figureCitation id="A970963C91E006B9CCE0658076D36A06" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">41</figureCitation>
); depression of frons present near antennal sockets (Fig.
<figureCitation id="E3B2AD1AE5A0F7EEEB19F1322DAF4AE8" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">34</figureCitation>
); eye ~ 2.5
<normalizedToken id="FE76DE7D5230DC0BC324B37243B1731F" originalValue="×">x</normalizedToken>
longer than temple in dorsal view (Fig.
<figureCitation id="A810E085DD5157ECE59A723DB3643F00" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">35</figureCitation>
); frons and vertex smooth, glabrous and moderately setose; face largely shiny, smooth and densely setose, but granulate latero-dorsally; median keel present; clypeus twice wider than its maximum height (Fig.
<figureCitation id="0E1709E8D3F5BC3EAE42B5CFF22D2F4F" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">34</figureCitation>
); clypeus semi-circular, moderately setose, and ventral margin of clypeus straight and above upper level of condyles of mandible; hypoclypeal depression present; length of maxillary palp nearly 0.9
<normalizedToken id="053D585558919022B4B9AAC4EBDF1520" originalValue="×">x</normalizedToken>
as long as height of head; malar sulcus absent; occipital carina absent dorsally; mandible triangular in lateral view, hardly twisted and gradually widened basally (Fig.
<figureCitation id="81C4AD8EE787B49E38CEBE270201D421" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">34</figureCitation>
).
</paragraph>
<paragraph id="18CBFDD94F3936D4D79057C9132FDAEA" pageId="0" pageNumber="219">
<emphasis id="C53A950B1525A8F7353709A045EAB6A4" bold="true" pageId="0" pageNumber="219">
<emphasis id="8EE1F72EA7181F92FA30C03E1CB92FDF" italics="true" pageId="0" pageNumber="219">Mesosoma</emphasis>
.
</emphasis>
Mesosoma 1.4
<normalizedToken id="657ADBA0CA66D242046E6F3C4E301166" originalValue="×">x</normalizedToken>
longer than its height; pronotal side largely smooth and smooth groove present along its ventral margin; propleuron smooth and sparsely setose, without transverse carinae; mesopleuron largely smooth and sparsely setose antero-dorsally and postero-ventrally, but precoxal sulcus oblique, medium-sized and densely crenulate; epicnemial area smooth (Fig.
<figureCitation id="F947D4E2A886BEA3AC7EAF9851E0770A" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">31</figureCitation>
); mesopleural sulcus smooth; anterior groove of metapleuron smooth; metapleuron reticulate-rugose and moderately setose (Fig.
<figureCitation id="000A971CEDEB6D72A11329F9FA34909A" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">31</figureCitation>
); notauli complete on disc of mesoscutum and narrowly crenulate; mesoscutum smooth, largely glabrous but middle lobe sparsely setose (Fig.
<figureCitation id="9FDEF31E901036B1E5FA960909E9701D" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">32</figureCitation>
); medio-posterior depression of mesoscutum round and shallow; scutellar sulcus wide and crenulate (Fig.
<figureCitation id="B6B025389402BF6FE4F04111824CFC6F" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">32</figureCitation>
); scutellum largely smooth and glabrous, rather flat in lateral view and protruding above level of mesoscutum; propodeum rugose with long medio-longitudinal carina, transverse carinae, and areola, remainder of propodeum largely smooth (Figs
<figureCitation id="2184F812BAB538C41E2E85F4C981847B" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">32</figureCitation>
,
<figureCitation id="6312ED94A9941143C385A478EFADE15A" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">36</figureCitation>
).
</paragraph>
<paragraph id="CA52B5ACC38F749C2C12AFCBEF2A9FBB" pageId="0" pageNumber="219">
<emphasis id="04619E8AB84F58F811232E8047A80FCA" bold="true" pageId="0" pageNumber="219">
<emphasis id="4177D10205A972F2451E614EAE600825" italics="true" pageId="0" pageNumber="219">Wings</emphasis>
.
</emphasis>
Fore wing (Fig.
<figureCitation id="98E4FB45658DEA4B1D79A95AEFB77D7F" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">30</figureCitation>
): pterostigma triangular and rather directly narrowed apically; vein 1-SR+M almost straight; vein 3-SR sublinear with vein r, converging with vein 2-M and 1.6
<normalizedToken id="32AC1B9E91E9757A869292C1AA2DE143" originalValue="×">x</normalizedToken>
longer than vein 2-SR; vein 2-SR almost straight; vein SR1 straight, 2.0
<normalizedToken id="696626BE52A19DFC6EA2D5F48A8C5C98" originalValue="×">x</normalizedToken>
longer than vein 3-SR; vein 1-M straight; r: 3-SR: SR1 = 5: 40: 84; vein m-cu distinctly postfurcal, converging to vein 1-M posteriorly and angled with vein 2-M; first subdiscal cell transverse and closed; vein CU1b present. Hind wing (Fig.
<figureCitation id="243A830E68E312A3191931E188F3CDFE" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">30</figureCitation>
): vein m-cu absent; vein 1r-m 0.8
<normalizedToken id="DDC2FCE876FAED1DBFF6ADDC697A7C66" originalValue="×">x</normalizedToken>
as long as vein 1-M; vein 2-M pigmented.
</paragraph>
<paragraph id="5E2277F84E32BCE764A5D47CF2EBC485" pageId="0" pageNumber="219">
<emphasis id="AD8745A378FEEF36BEA82A5310C5E568" bold="true" pageId="0" pageNumber="219">
<emphasis id="AA0B94CBCD28F26FE587E006B83A8B3E" italics="true" pageId="0" pageNumber="219">Legs</emphasis>
.
</emphasis>
Hind femur 4.5
<normalizedToken id="64FB7A74E839BA14CDBF7D31F15AB4C5" originalValue="×">x</normalizedToken>
longer than its maximum width (Fig.
<figureCitation id="92FAE19197D0ADAF29AAAD16D2E9B7D5" captionStart="Figure 29" captionStartId="F6" captionText="Figure 29. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀, habitus, lateral." figureDoi="10.3897/zookeys.1193.115831.figure29" httpUri="https://binary.pensoft.net/fig/997931" pageId="0" pageNumber="219">29</figureCitation>
).
</paragraph>
<paragraph id="A6280C7A9D7E3962F65EC3172AB2427B" pageId="0" pageNumber="219">
<emphasis id="807293015B926CC0244E23F58D293A10" bold="true" pageId="0" pageNumber="219">
<emphasis id="75CF2BAA8935E9CFB09E1217948FE6B2" italics="true" pageId="0" pageNumber="219">Metasoma</emphasis>
.
</emphasis>
First metasomal tergite 1.5
<normalizedToken id="54984672BDAB5214D9F44E6CF0F6B078" originalValue="×">x</normalizedToken>
longer than its apical width; first tergite gradually widened apically and its surface densely longitudinally striate postero-medially, and remainder of tergite shiny and smooth (Fig.
<figureCitation id="DB2739016A5FD54303F4B82F0EEFD3D8" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">37</figureCitation>
); dorsope present and surrounded by strongly curved dorsal carinae (Fig.
<figureCitation id="905CE5F095A3F8DB4D1639F2AE3AB142" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">37</figureCitation>
); second metasomal suture absent dorsally (Fig.
<figureCitation id="B542C54B63CB880D92CF9A166694A60E" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">33</figureCitation>
); second tergite striate-rugose medially except a pair of droplet-shaped impressions anteriorly; following tergites shiny, smooth and moderately setose posteriorly (Fig.
<figureCitation id="58697D915E8520210952454BA222E6CD" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">33</figureCitation>
); setose part of ovipositor sheath ~ 1.2
<normalizedToken id="8DEAF9F745F9339B82EF96FD542F8598" originalValue="×">x</normalizedToken>
longer than first tergite and 0.1
<normalizedToken id="B0B4E5886BF11A0917DD6F59B49B3011" originalValue="×">x</normalizedToken>
as long as fore wing (Fig.
<figureCitation id="1AFC6135A9A4E616E74A9D143C0CFC7D" captionStart="Figures 30–41" captionStartId="F7" captionText="Figures 30 - 41. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀ 30 wings 31 mesosoma, lateral view 32 mesosoma, dorsal view 33 metasoma, dorsal view 34 head, anterior view 35 head, dorsal view 36 propodeum, dorsal view 37 1 st metasomal tergite, dorsal view 38 ovipositor and its sheath, lateral view 39 basal part of antenna 40 apical part of antenna 41 antenna." figureDoi="10.3897/zookeys.1193.115831.figures30-41" httpUri="https://binary.pensoft.net/fig/997932" pageId="0" pageNumber="219">38</figureCitation>
).
</paragraph>
<paragraph id="3912D2ED90B1D5E36F4590A306842FA9" pageId="0" pageNumber="219">
<emphasis id="223EFCE018D86FF622362D307A02F82E" bold="true" pageId="0" pageNumber="219">
<emphasis id="D234B0A9B569F30C1834126D0E10275A" italics="true" pageId="0" pageNumber="219">Colour</emphasis>
.
</emphasis>
Body generally brown to black (Fig.
<figureCitation id="65CA3D94037A39AF4CB5E2FDA64BBF58" captionStart="Figure 29" captionStartId="F6" captionText="Figure 29. Xynobius duoferus Han &amp; van Achterberg, sp. nov., holotype, ♀, habitus, lateral." figureDoi="10.3897/zookeys.1193.115831.figure29" httpUri="https://binary.pensoft.net/fig/997931" pageId="0" pageNumber="219">29</figureCitation>
); entire head, transverse band below the precoxal sulcus, tarsi, inside of dorsope, and first and second tergites dorsally dark brown; legs, remaining tergites, ovipositor, and basal segments (1st-6th) of antenna brown; palp pale yellowish or white; apical segments (20th-26th) of antenna white to white-brown; pterostigma and veins of wings pale brown; wings subhyaline.
</paragraph>
</subSubSection>
<subSubSection id="FCB9A982BF5A240A43E81C2270751031" pageId="0" pageNumber="219" type="distribution">
<paragraph id="D4DE0054FCF12FD4CEEF3B038AC6BFD8" pageId="0" pageNumber="219">Distribution.</paragraph>
<paragraph id="AFDE5CDD0A35817EDD20B93DCBCED7EF" pageId="0" pageNumber="219">South Korea.</paragraph>
</subSubSection>
<subSubSection id="0069E5A449482AFB3887359A0F9CD150" pageId="0" pageNumber="219" type="biology_ecology">
<paragraph id="108A3707BDF6EDB6C44E11317FF8DA87" pageId="0" pageNumber="219">Biology.</paragraph>
<paragraph id="4EB89B81009E0E9053D6E43A67E743C4" pageId="0" pageNumber="219">Unknown.</paragraph>
</subSubSection>
<subSubSection id="339B0225C6C4140A7990715844C70FEE" pageId="0" pageNumber="219" type="etymology">
<paragraph id="BEADDA9237C56FEABCEB293CF1B827DE" pageId="0" pageNumber="219">Etymology.</paragraph>
<paragraph id="5C40DCE5D28226FD7EF0E3C7FC2291FE" pageId="0" pageNumber="219">
Name derived from
<emphasis id="23794B89BEF582C6754C2F166A875DB7" italics="true" pageId="0" pageNumber="219">duo</emphasis>
(Latin for two) and -
<emphasis id="BF566BD004F6BA400BED78288100A778" italics="true" pageId="0" pageNumber="219">fero</emphasis>
(suffix in Latin meaning carrying or having), because of the two apical dark antennal segments.
</paragraph>
</subSubSection>
<subSubSection id="01D6C9BA47B548082D7B5C0159023742" pageId="0" pageNumber="219" type="remarks">
<paragraph id="14C55A6253A35580EB3FA3797450C108" pageId="0" pageNumber="219">Remarks.</paragraph>
<paragraph id="77CFB7E502DA3BD93D46C71CCD43BB3D" pageId="0" pageNumber="219">
This species runs to
<taxonomicName id="D8015C3507B1001683CE73FDD868D5C1" class="Insecta" family="Braconidae" genus="Xynobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xynobius notauliferus" order="Hymenoptera" pageId="0" pageNumber="219" phylum="Arthropoda" rank="species" species="notauliferus">
<emphasis id="7FE580CBBA379D0A9BE45BCA48BA156E" italics="true" pageId="0" pageNumber="219">Xynobius notauliferus</emphasis>
</taxonomicName>
Li &amp; van Achterberg, 2013 in the key by
<bibRefCitation id="993900F0CBFD85FF97B502F8BF14D267" DOI="https://doi.org/10.3897/zookeys.326.5911" author="Li, X-Y" journalOrPublisher="ZooKeys" pageId="0" pageNumber="219" pagination="1 - 186" refId="B9" refString="Li, X-Y, van, Achterberg C, Tan, J-C, 2013. Revision of the subfamily Opiinae (Hymenoptera, Braconidae) from Hunan (China), including thirty-six new species and two new genera. ZooKeys 268: 1 - 186, DOI: https://doi.org/10.3897/zookeys.326.5911" title="Revision of the subfamily Opiinae (Hymenoptera, Braconidae) from Hunan (China), including thirty-six new species and two new genera." url="https://doi.org/10.3897/zookeys.326.5911" volume="268" year="2013">Li et al. (2013)</bibRefCitation>
. It differs by having the length of maxillary palp 0.9
<normalizedToken id="BC50B0E99E4177E31C47B6C75FAF3BF3" originalValue="×">x</normalizedToken>
height of head (1.4
<normalizedToken id="5AA690D30FE4AF563B4CCD1C02520EBC" originalValue="×">x</normalizedToken>
in
<taxonomicName id="0904A452D295A28FF16C481E0D78FB18" lsidName="X. notauliferus" pageId="0" pageNumber="219" rank="species" species="notauliferus">
<emphasis id="BC9C93617EE34AA7750C7A65164943DD" italics="true" pageId="0" pageNumber="219">X. notauliferus</emphasis>
</taxonomicName>
), face smooth, but granulate latero-dorsally (smooth), clypeus twice wider than high (1.6
<normalizedToken id="4CB9F5685D49F1D4AB59E7C184042C2E" originalValue="×">x</normalizedToken>
wider than its maximum height), first metasomal tergite 1.5
<normalizedToken id="F4B3635F607230EB8B778ABEFDF7285A" originalValue="×">x</normalizedToken>
longer than its apical width (length 1.3
<normalizedToken id="C5A5A1474489F3A2137D59AF24B5B5DF" originalValue="×">x</normalizedToken>
), antenna of ♀ with two apical antennal segments dark brown (6 or 7 such segments), pterostigma directly narrowed distally (gradually narrowed) and outer side of hind femur without brownish patch (with patch).
</paragraph>
</subSubSection>
</treatment>
</document>

296
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@ -0,0 +1,296 @@
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<mods:title>Codonoboea personatiflora (Gesneriaceae), a new species from Peninsular Malaysia</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kiew, Ruth</mods:namePart>
</mods:name>
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<mods:role>
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<mods:namePart>Sam, Yen-Yen</mods:namePart>
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<treatment ID-GBIF-Taxon="152024466" LSID="urn:lsid:plazi:treatment:B60178BA659CF41B041E376443D47DD8" httpUri="http://treatment.plazi.org/id/B60178BA659CF41B041E376443D47DD8" lastPageId="3" lastPageNumber="64" pageId="0" pageNumber="61">
<subSubSection pageId="0" pageNumber="61" type="nomenclature">
<paragraph pageId="0" pageNumber="61">
<taxonomicName LSID="6A84C2F4-3DC8-5C54-B813-34C9D5D1A8F3" authority="Kiew &amp; Y. Y. Sam" authorityName="Kiew &amp; Y. Y. Sam" authorityYear="2012" class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Codonoboea personatiflora" order="Lamiales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="personatiflora" status="sp. nov.">Codonoboea personatiflora Kiew &amp; Y.Y.Sam</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="61">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Codonoboea personatiflora Kiew &amp; Y. Y. Sam. A habit B coloured young leaves C pair-flowered cymose inflorescences D flower (top view) E flower (from below) G longitudinal section with upper lip removed." httpUri="https://binary.pensoft.net/fig/10188" pageId="0" pageNumber="61">Figure 1</figureCitation>
<figureCitation captionStart="Map 1" captionStartId="F2" captionText="Map 1. Distribution of Codonoboea personatiflora in Peninsular Malaysia." httpUri="https://binary.pensoft.net/fig/10189" pageId="0" pageNumber="61">Map 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="diagnosis">
<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="61">
Different from all other species of
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="61">Codonoboea</emphasis>
</taxonomicName>
Ridl. in its personate flower where the lower lip is directed upwards and completely closes the mouth of the corolla tube.
</paragraph>
</subSubSection>
<subSubSection lastPageId="1" lastPageNumber="62" pageId="0" pageNumber="61" type="type">
<paragraph pageId="0" pageNumber="61">Type.</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Peninsular Malaysia.</emphasis>
Terengganu: Kemaman, Sungai Nipah, Bukit Kajang,
<geoCoordinate degrees="4" direction="north" minutes="20" orientation="latitude" precision="925" value="4.3333335">4°20'N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="07" orientation="longitude" precision="925" value="103.11667">103°07'E</geoCoordinate>
, 22 Nov 1935, E.J.H.Corner SFN 30540 (holotype: SING!; isotypes: K!, L, E, SAR!).
</paragraph>
<paragraph pageId="0" pageNumber="61">
Robust, erect, unbranched herb.
<emphasis bold="true" pageId="0" pageNumber="61">Stem</emphasis>
woody to 65 cm tall, flowering at ca 15 cm tall, bare stem below the leaves 4-9 mm diam.
<emphasis bold="true" pageId="0" pageNumber="61">Leaves</emphasis>
opposite, pairs equal, spaced 8-17 mm apart, glossy when young, reddish towards the base and bright yellowish green toward apex, older leaves uniformly dull greyish green above; petioles 1.3-3 cm long, slightly grooved above, minutely pubescent; lamina oblanceolate, thinly leathery, 15-34
<normalizedToken originalValue="×">x</normalizedToken>
4-8 cm, lateral veins whitish or yellowish green, sometimes with a silvery band along the midrib, paler beneath, base narrowed then slightly rounded, margin serrate in the upper half, teeth ca 1 mm long, apex narrowly acuminate, acumen to 1
<normalizedToken originalValue="–2.5(–">-2.5(-</normalizedToken>
4) cm long; midrib and veins impressed above, prominent beneath in life, lateral veins 9-20 pairs, ferrugineous beneath.
</paragraph>
<paragraph lastPageId="1" lastPageNumber="62" pageId="0" pageNumber="61">
<taxonomicName genus="Inflorescence" lsidName="" pageId="0" pageNumber="61" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Inflorescence</emphasis>
</taxonomicName>
a pair-flowered dichasial cyme with up to 5th order branching, erect from the upper leaf axils, 12-34 cm long, peduncle purple or reddish brown, 7.5-29 cm long, bracts green, linear, 2-10
<normalizedToken originalValue="×">x</normalizedToken>
ca 0.5 mm, pedicels 2-5 mm long. Inflorescence, pedicel and calyx with minute, dense, non-glandular pubescence.
<emphasis bold="true" pageId="0" pageNumber="61">Flowers</emphasis>
pendent, to 20 mm long, almost hanging vertically down, buds green or only at the base; calyx brownish purple or green, ca 1.5-2
<normalizedToken originalValue="×">x</normalizedToken>
0.75 mm long, 5-lobed divided almost to the base, apex acute; corolla pale cream with pale pink lobes, personate, densely glandular pubescent outside, tube 9-13
<normalizedToken originalValue="×">x</normalizedToken>
3-5 mm, lower half of tube cylindric, the upper half gradually dilating to ca twice the width of the basal half, mouth 6-9 mm diam., upper lip erect, glabrous, ca 7 mm long with 2 strongly reflexed lobes, each ca 2.5
<normalizedToken originalValue="×">x</normalizedToken>
2.5 mm with a raised patch of long glandular hairs between lobes, lower lip positioned upwards and appressed against the upper lip closing the corolla mouth, 8-10
<normalizedToken originalValue="×">x</normalizedToken>
6.5-9 mm long, the 3 rounded, glabrous lobes reflexed to cucullate, raised and forming a rim, lateral lobes ca 2
<normalizedToken originalValue="×">x</normalizedToken>
2 mm, central lobe ca 3
<normalizedToken originalValue="×">x</normalizedToken>
2.5 mm, throat with dense patch of deep pink unbranched long multiseriate glandular hairs, nectar guides 2, strongly raised, concolorous with the tube; stamens 2, joined 4-5 mm from corolla base, filaments white, slightly bowed inwards, 7-13 mm long with 2 blunt horns at the attachment to the anther, anthers purple or pale cream, ca 2
<normalizedToken originalValue="×">x</normalizedToken>
0.75 mm, cohering face-to-face, staminodes 2, slender ca 3 mm long, hooked at apex; nectary cylindric, margin unequally lobed, upper lobes 2-3 mm long, lower lobes 2-3.5 mm long; ovary and style slender, with
<pageBreakToken pageId="1" pageNumber="62" start="start">dense</pageBreakToken>
brownish glandular hairs, ovary white, 5-8
<normalizedToken originalValue="×">x</normalizedToken>
0.75-1 mm long, style purple-brown, 5-6 mm long, stigma globose, almost 0.5 mm diam. The anthers and stigma are positioned at the same level just below the upper lip.
<emphasis bold="true" pageId="1" pageNumber="62">Fruits</emphasis>
slender, cylindric, 22-45
<normalizedToken originalValue="×">x</normalizedToken>
1-1.5 mm, dehiscing along the upper suture.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10188" pageId="1" pageNumber="62" start="Figure 1" startId="F1">
<paragraph pageId="1" pageNumber="62">
<emphasis bold="true" pageId="1" pageNumber="62">Figure 1.</emphasis>
<taxonomicName authorityName="Kiew &amp; Y. Y. Sam" authorityYear="2012" class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="1" pageNumber="62" phylum="Magnoliophyta" rank="species" species="personatiflora">
<emphasis italics="true" pageId="1" pageNumber="62">Codonoboea personatiflora</emphasis>
</taxonomicName>
Kiew &amp; Y.Y.Sam.
<emphasis bold="true" pageId="1" pageNumber="62">A</emphasis>
habit
<emphasis bold="true" pageId="1" pageNumber="62">B</emphasis>
coloured young leaves
<emphasis bold="true" pageId="1" pageNumber="62">C</emphasis>
pair-flowered cymose inflorescences
<emphasis bold="true" pageId="1" pageNumber="62">D</emphasis>
flower (top view)
<emphasis bold="true" pageId="1" pageNumber="62">E</emphasis>
flower (from below)
<emphasis bold="true" pageId="1" pageNumber="62">G</emphasis>
longitudinal section with upper lip removed.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="1" pageNumber="62" type="distribution">
<paragraph pageId="1" pageNumber="62">Distribution.</paragraph>
<paragraph pageId="1" pageNumber="62">Peninsular Malaysia, endemic in the state of Terengganu.</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10189" pageId="1" pageNumber="62" start="Map 1" startId="F2">
<paragraph pageId="1" pageNumber="62">
<emphasis bold="true" pageId="1" pageNumber="62">Map 1.</emphasis>
Distribution of
<taxonomicName authorityName="Kiew &amp; Y. Y. Sam" authorityYear="2012" class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="1" pageNumber="62" phylum="Magnoliophyta" rank="species" species="personatiflora">
<emphasis italics="true" pageId="1" pageNumber="62">Codonoboea personatiflora</emphasis>
</taxonomicName>
in Peninsular Malaysia.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="1" pageNumber="62" type="ecology">
<paragraph pageId="1" pageNumber="62">Ecology.</paragraph>
<paragraph pageId="1" pageNumber="62">In primary or logged-over lowland mixed dipterocarp forest at low altitudes (below 100 m), on shaded hillsides or slopes, often above streams.</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="62" type="etymology">
<paragraph pageId="1" pageNumber="62">Etymology.</paragraph>
<paragraph pageId="1" pageNumber="62">Named for its personate (closed) flower that is unique for the genus.</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62" type="conservation status">
<paragraph pageId="1" pageNumber="62">Conservation status.</paragraph>
<paragraph lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
EN B2a,b(ii,iii). Following the IUCN Criteria and Categories (
<bibRefCitation author="IUCN," pageId="4" pageNumber="65" refId="B3" refString="IUCN, , 2001. Red list categories and criteria version 3.1. IUCN Species Survival Commission, IUCN, Gland, Switzerland &amp; Cambridge, UK." title="Red list categories and criteria version 3.1. IUCN Species Survival Commission, IUCN, Gland, Switzerland &amp; Cambridge, UK." year="2001">IUCN 2001</bibRefCitation>
;
<bibRefCitation author="Chua, LSL" pageId="4" pageNumber="65" refId="B2" refString="Chua, LSL, 2010. Species assessment and conservation in Peninsular Malaysia. In: Kiew R, Chung RCK, Saw LG, Soepadmo E, Boyce PC (Eds) Flora of Peninsular Malaysia 2,1: 47-54." title="Species assessment and conservation in Peninsular Malaysia. In: Kiew R, Chung RCK, Saw LG, Soepadmo E, Boyce PC (Eds) Flora of Peninsular Malaysia 2,1: 47 - 54." year="2010">Chua 2010</bibRefCitation>
), this species falls within the Endangered Category because it is known from five localities, has in total an area of occupancy of 28 sq km. In addition, it is nowhere found within Peninsular
<normalizedToken originalValue="Malaysia’s">Malaysia's</normalizedToken>
network of Totally Protect
<pageBreakToken pageId="2" pageNumber="63" start="start">ed</pageBreakToken>
Areas and it grows at low altitudes in lowland forest, a habitat that is threatened by or already has been logged or is threatened by forest clearance for oil palm plantations.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="63" type="other specimens examined">
<paragraph pageId="2" pageNumber="63">Other specimens examined.</paragraph>
<paragraph pageId="2" pageNumber="63">
<emphasis bold="true" pageId="2" pageNumber="63">Peninsular Malaysia</emphasis>
. Terengganu: Dungun, Jengai Forest Reserve, Compartment 71, 15 Mar 1998, Anon. s.n. (SING!), Compartment 52, 65 m alt.,
<geoCoordinate degrees="4" direction="north" minutes="32.76" orientation="latitude" precision="9" value="4.546">4°32.76'N</geoCoordinate>
,
<geoCoordinate degrees="102" direction="east" minutes="58.34" orientation="longitude" precision="9" value="102.972336">102°58.34'E</geoCoordinate>
, 17 Oct 2002, Y.Y.Sam FRI 47153 (KEP!, SAN!); Hulu Terengganu, Ulu Telemong Forest Reserve,
<geoCoordinate degrees="5" direction="north" minutes="12" orientation="latitude" precision="925" value="5.2">5°12'N</geoCoordinate>
,
<geoCoordinate degrees="102" direction="east" minutes="46" orientation="longitude" precision="925" value="102.76667">102°46'E</geoCoordinate>
, 25 Jul 2006, R.Kiew RK 5339 (KEP!, K!, SAR!); Hulu Terengganu, Ladang Ternakan TERSAT, 62 m alt.,
<geoCoordinate degrees="5" direction="north" minutes="01.07" orientation="latitude" precision="9" value="5.017833">5°01.07'N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="01.17" orientation="longitude" precision="9" value="103.0195">103°01.17'E</geoCoordinate>
, 31 Oct 2009, M.Kamarul-Hisham FRI 67168 (KEP! SING!); Kemaman, Sungai Nipah Forest Reserve, Jeram Tanduk, 81 m alt.,
<geoCoordinate degrees="4" direction="north" minutes="18.93" orientation="latitude" precision="9" value="4.3155">4°18.93'N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="10.42" orientation="longitude" precision="9" value="103.17367">103°10.42'E</geoCoordinate>
, Y.Y.Sam FRI 47197 (KEP! SING!); probably from Kemaman, Sungai Nipah, collector unknown, flowered in Forest Research Institute Malaysia Nursery on 9 Apr 2012, FRI 75314(KEP!); Setiu, Ulu Setiu Forest Reserve,
<geoCoordinate degrees="5" direction="north" minutes="26" orientation="latitude" precision="925" value="5.4333334">5°26'N</geoCoordinate>
,
<geoCoordinate degrees="102" direction="east" minutes="44" orientation="longitude" precision="925" value="102.73333">102°44'E</geoCoordinate>
, 7 Mar 2002, Y.Y.Sam FRI 44395 (KEP!), flowered in Forest Research Institute Malaysia Nursery on 9 Sept 2003, Y.Y.Sam FRI 44483 (KEP!).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63" type="discussion">
<paragraph pageId="2" pageNumber="63">Discussion.</paragraph>
<paragraph lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
This new species belongs to
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="2" pageNumber="63" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="63">Codonoboea</emphasis>
</taxonomicName>
sect.
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" genus="Didymanthus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Caryophyllales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="63">Didymanthus</emphasis>
</taxonomicName>
in its erect habit, petiolate leaves in distant pairs, cymose inflorescences with a long peduncle and
<pageBreakToken pageId="3" pageNumber="64" start="start">inconspicuous</pageBreakToken>
bracts, and medium-sized (to 2 cm long) flowers (
<bibRefCitation author="Ridley, HN" journalOrPublisher="Gardens' Bulletin Singapore" pageId="4" pageNumber="65" refId="B6" refString="Ridley, HN, 1923. Gesneriaceae. Flora of Malay Peninsula.2: 495-547." title="Gesneriaceae. Flora of Malay Peninsula. 2: 495 - 547." year="1923">Ridley 1923</bibRefCitation>
). However, it differs from the other species in this section, and indeed from all other known
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="64">Codonoboea</emphasis>
</taxonomicName>
species, in its personate flower. The special feature of this flower is the lower lip where the recurved lobes form a rim that is pressed upwards against the upper lip and so closes the corolla mouth, unlike the usual open funnel- or trumpet-shaped or campanulate corolla of
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="64">Codonoboea</emphasis>
</taxonomicName>
species. In addition, the aperture is filled by the long hairs, a feature not seen in other
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="64">Codonoboea</emphasis>
</taxonomicName>
species. The lower lip appears to be hinged because it is readily bends downward when pressure is applied so opening up the mouth of the corolla tube.
</paragraph>
<paragraph pageId="3" pageNumber="64">
The majority of
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="64">Codonoboea</emphasis>
</taxonomicName>
species are nectar flowers (the cylindrical nectary surrounding the base of the ovary supplies nectar as the reward for the pollinator) with a narrow tubular flower that is strongly dilated to produce an open funnel- or trumpet-shaped flower, often with the lower lobes projecting and forming a landing platform for the insect pollinator. Usually there are yellow or orange nectar guides on the floor of the tube. The flower is therefore open to any insect small enough to enter the tube. Although there are very few observations on pollination in this genus, bumblebees (
<taxonomicName class="Hexapoda" family="Apidae" genus="Bombus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="64">Bombus</emphasis>
</taxonomicName>
sp.) have been observed visiting two species,
<taxonomicName authorityName="Kiew" authorityYear="2011" baseAuthorityName="Ridl." class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="species" species="hispida">
<emphasis italics="true" pageId="3" pageNumber="64">Codonoboea hispida</emphasis>
</taxonomicName>
(Ridl.) Kiewand
<taxonomicName authorityName="Kiew" authorityYear="2011" baseAuthorityName="Ridl." class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="species" species="robinsonii">
<emphasis italics="true" pageId="3" pageNumber="64">Codonoboea robinsonii</emphasis>
</taxonomicName>
(Ridl.) Kiew, both belonging to
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="64">Codonoboea</emphasis>
</taxonomicName>
sect.
<taxonomicName class="Magnoliopsida" family="Amaranthaceae" genus="Didymanthus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Caryophyllales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="64">Didymanthus</emphasis>
</taxonomicName>
(
<bibRefCitation author="Kiew, R" journalOrPublisher="Malayan Nature Journal" pageId="4" pageNumber="65" pagination="257 - 265" refId="B4" refString="Kiew, R, 2009. The natural history of Malaysian Gesneriaceae. Malayan Nature Journal 61 (3): 257 - 265" title="The natural history of Malaysian Gesneriaceae." volume="61" year="2009">Kiew 2009</bibRefCitation>
) and it is likely, based on floral morphology, that bees are the pollinators of most of these
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="64">Codonoboea</emphasis>
</taxonomicName>
species. The other type of flower seen in the genus is the smaller, short-tubed, often campanulate pollen flower where the anthers are large and conspicuous in the mouth of the corolla. These flowers either have very small nectaries or none at all and pollen is offered as the reward.
</paragraph>
<paragraph pageId="3" pageNumber="64">
This new species is obviously a nectar flower but is unique in
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Codonoboea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="64">Codonoboea</emphasis>
</taxonomicName>
in that the upper and lower lips fit closely together so that the mouth is closed and in addition there is a tangle of long hairs just inside the mouth that might prevent small insects from squeezing between the lips. This personate flower closely resembles that of the ornamental snapdragon,
<taxonomicName class="Eudicots" family="Plantaginaceae" genus="Antirrhinum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="species" species="majus">
<emphasis italics="true" pageId="3" pageNumber="64">Antirrhinum majus</emphasis>
</taxonomicName>
L. (
<taxonomicName authorityName="de Jussieu" authorityYear="1789" class="Magnoliopsida" family="Scrophulariaceae" genus="Scrophulariaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Lamiales" pageId="3" pageNumber="64" phylum="Magnoliophyta" rank="genus">Scrophulariaceae</taxonomicName>
) that is pollinated by bumblebees that on landing on the flower are sufficiently heavy to depress and open the lower lip.
</paragraph>
<paragraph pageId="3" pageNumber="64">
There are only two other examples of this type of personate flower in Malaysian
<taxonomicName genus="Gesneriaceae" lsidName="" pageId="3" pageNumber="64" rank="genus">Gesneriaceae</taxonomicName>
, namely in
<taxonomicName class="Dicotyledoneae" family="Gesneriaceae" genus="Didymocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Tubiflorae" pageId="3" pageNumber="64" phylum="Angiospermae" rank="species" species="antirrhinoides">
<emphasis italics="true" pageId="3" pageNumber="64">Didymocarpus antirrhinoides</emphasis>
</taxonomicName>
A.Weber (
<bibRefCitation author="Weber, A" journalOrPublisher="Blumea" pageId="4" pageNumber="65" pagination="291 - 309" refId="B7" refString="Weber, A, Burtt, BL, 1983. Didymocarpus corchorifolius and its allies (Gesneriaceae). Blumea 28 (2): 291 - 309" title="Didymocarpus corchorifolius and its allies (Gesneriaceae)." volume="28" year="1983">Weber and Burtt 1983</bibRefCitation>
) that has a flower of similar size (15-23 mm long) and
<taxonomicName genus="Rhyncholossum" lsidName="" pageId="3" pageNumber="64" rank="species" species="medusothrix">
<emphasis italics="true" pageId="3" pageNumber="64">Rhyncholossum medusothrix</emphasis>
</taxonomicName>
B.L.Burtt (
<bibRefCitation author="Burtt, BL" pageId="4" pageNumber="65" refId="B1" refString="Burtt, BL, 1962. Studies in the Gesneriaceae of the Old World. XXIII: Rhynchoglossum and Klugia.24: 167-171." title="Studies in the Gesneriaceae of the Old World. XXIII: Rhynchoglossum and Klugia. 24: 167 - 171." year="1962">Burtt 1962</bibRefCitation>
) with a corolla tube 10-25 mm long that in addition has
<normalizedToken originalValue="‘medusoid’">'medusoid'</normalizedToken>
hairs in the throat. However, for none of these personate flowers is the pollinator known.
</paragraph>
</subSubSection>
</treatment>
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Genus
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<taxonomicName id="F9A84D18245DC62DFF04FF4DFE34FF55" ID-CoL="8NXHR" authority="Zeuner, 1940" authorityName="Zeuner" authorityYear="1940" box="[151,419,186,207]" class="Insecta" family="Tettigoniidae" genus="Phlugiolopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="0CDCEA89245DC62DFF04FF4DFE8CFF55" box="[151,283,186,207]" italics="true" pageId="1" pageNumber="296">Phlugiolopsis</emphasis>
<bibRefCitation id="5A394B6A245DC62DFEB2FF4DFE34FF55" author="Zeuner, F. E." box="[289,419,186,207]" pageId="1" pageNumber="296" pagination="76 - 84" refId="ref3868" refString="Zeuner, F. E. (1940) Phlugiolopsis henryi, n. g., n. sp., a new tettigoniid, and other Saltatoria (Orthop.) from the Royal Botanic Gardens, Kew. Journal of the Society for British Entomology, 2, 76 - 84." type="journal article" year="1940">Zeuner, 1940</bibRefCitation>
</taxonomicName>
,
<emphasis id="0CDCEA89245DC62DFE3EFF4DFCF3FF55" box="[429,868,186,207]" italics="true" pageId="1" pageNumber="296">Journal of the Society for British Entomology</emphasis>
, 2: 77;
<bibRefCitation id="5A394B6A245DC62DFC3DFF4DFBDDFF55" author="Yamasaki, T." box="[942,1098,186,207]" pageId="1" pageNumber="296" pagination="353 - 358" refId="ref3748" refString="Yamasaki, T. (1986) Discovery of Phlugiolopsis (Orthoptera, Tettigoniidae, Meconematinae) in the Ryukyu Islands. Kontyu, Tokyo, 54 (2), 353 - 358." type="journal article" year="1986">Yamasaki, 1986</bibRefCitation>
,
<emphasis id="0CDCEA89245DC62DFBC7FF4DFB0EFF55" box="[1108,1177,186,207]" italics="true" pageId="1" pageNumber="296">Kontyû</emphasis>
,
<emphasis id="0CDCEA89245DC62DFB35FF4DFB49FF55" box="[1190,1246,186,207]" italics="true" pageId="1" pageNumber="296">Tokyo</emphasis>
, 54(2): 353;
<bibRefCitation id="5A394B6A245DC62DFACEFF4DFEACFF76" author="Kevan, D. K. M. &amp; Jin, X. B." pageId="1" pageNumber="296" pagination="253 - 274" refId="ref3070" refString="Kevan, D. K. M. &amp; Jin, X. B. (1993) New species of the Xiphidiosis - group from the Indian region (Grylloptera Tettigonioidea Meconematidae). Tropical Zoology, 6 (2), 253 - 274. https: // doi. org / 10.1080 / 03946975.1993.10539226" type="journal article" year="1993">Kevan &amp; Jin, 1993</bibRefCitation>
,
<emphasis id="0CDCEA89245DC62DFED6FF20FE7EFF76" box="[325,489,215,236]" italics="true" pageId="1" pageNumber="296">Tropical Zoology</emphasis>
, 6: 254;
<bibRefCitation id="5A394B6A245DC62DFDD2FF20FD6DFF76" author="Kevan, D. K. M. &amp; Jin, X. B." box="[577,762,215,236]" pageId="1" pageNumber="296" pagination="253 - 274" refId="ref3070" refString="Kevan, D. K. M. &amp; Jin, X. B. (1993) New species of the Xiphidiosis - group from the Indian region (Grylloptera Tettigonioidea Meconematidae). Tropical Zoology, 6 (2), 253 - 274. https: // doi. org / 10.1080 / 03946975.1993.10539226" type="journal article" year="1993">Kevan &amp; Jin, 1993</bibRefCitation>
,
<emphasis id="0CDCEA89245DC62DFC96FF20FC75FF76" box="[773,994,215,236]" italics="true" pageId="1" pageNumber="296">Invertebrate Taxonomy</emphasis>
, 7: 1591; Bian, Shi &amp; Chang, 2012,
<emphasis id="0CDCEA89245DC62DFAD4FF20FA02FF76" box="[1351,1429,215,236]" italics="true" pageId="1" pageNumber="296">Zootaxa</emphasis>
, 3281: 22;
<bibRefCitation id="5A394B6A245DC62DFEBFFF02FD9EFE90" author="Wang, H. Q. &amp; Li, K. &amp; Liu, X. W." box="[300,521,245,266]" pageId="1" pageNumber="296" pagination="27 - 48" refId="ref3601" refString="Wang, H. Q., Li, K. &amp; Liu, X. W. (2012) A taxonomic study on the species of the genus Phlugiolopsis Zeuner (Orthoptera, Tettigoniidae, Meconematinae). Zootaxa, 3332, 27 - 48." type="journal article" year="2012">Wang, Li &amp; Liu, 2012</bibRefCitation>
,
<emphasis id="0CDCEA89245DC62DFD87FF02FDF4FE90" box="[532,611,245,266]" italics="true" pageId="1" pageNumber="296">Zootaxa</emphasis>
, 3384: 68; Bian, Shi &amp; Chang, 2012,
<emphasis id="0CDCEA89245DC62DFC4FFF02FBBDFE90" box="[988,1066,245,266]" italics="true" pageId="1" pageNumber="296">Zootaxa</emphasis>
, 3411: 55;
<bibRefCitation id="5A394B6A245DC62DFB0FFF02FA0FFE90" author="Bian, X. &amp; Shi, F. M. &amp; Chang, Y. L." box="[1180,1432,245,266]" pageId="1" pageNumber="296" pagination="159 - 191" refId="ref2332" refString="Bian, X., Shi, F. M. &amp; Chang, Y. L. (2013) Second supplement for the genus Phlugiolopsis Zeuner, 1940 (Orthoptera: Tettigoniidae: Meconematinae) from China, with eight new species. Zootaxa, 3701 (2), 159 - 191. https: // doi. org / 10.11646 / zootaxa. 3701.2.3" type="journal article" year="2013">Bian, Shi &amp; Chang, 2013</bibRefCitation>
,
<emphasis id="0CDCEA89245DC62DFF54FEE5FE81FEBD" box="[199,278,274,295]" italics="true" pageId="1" pageNumber="296">Zootaxa</emphasis>
, 3701(2): 159;
<bibRefCitation id="5A394B6A245DC62DFE38FEE5FDC3FEBD" author="Bian, X. &amp; Shi, F. M." box="[427,596,274,295]" pageId="1" pageNumber="296" pagination="341 - 366" refId="ref2400" refString="Bian, X. &amp; Shi, F. M. (2018) New taxa of the genus Phlugiolopsis (Orthoptera: Tettigoniidae: Meconematinae) from Yunnan, China, with comments on the importance to taxonomy of the left tegmen. Zootaxa, 4532 (3), 341 - 366. https: // doi. org / 10.11646 / zootaxa. 4532.3.2" type="journal article" year="2018">Bian &amp; Shi, 2018</bibRefCitation>
,
<emphasis id="0CDCEA89245DC62DFDCDFEE5FD3BFEBD" box="[606,684,274,295]" italics="true" pageId="1" pageNumber="296">Zootaxa</emphasis>
, 4532(3): 341.
</paragraph>
</subSubSection>
<subSubSection id="76B26510245DC62DFF04FED8FDFDFEDE" box="[151,618,303,324]" pageId="1" pageNumber="296" type="materials_examined">
<paragraph id="3E17369B245DC62DFF04FED8FDFDFEDE" blockId="1.[151,1436,151,324]" box="[151,618,303,324]" pageId="1" pageNumber="296">
<typeStatus id="E1138839245DC62DFF04FED8FF50FEDE" box="[151,199,303,324]" pageId="1" pageNumber="296">Type</typeStatus>
species:
<taxonomicName id="F9A84D18245DC62DFE8DFED8FDFDFEDE" authority="Zeuner, 1940" authorityName="Zeuner" authorityYear="1940" box="[286,618,303,324]" class="Insecta" family="Tettigoniidae" genus="Phlugiolopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="species" species="henryi">
<emphasis id="0CDCEA89245DC62DFE8DFED8FE72FEDE" box="[286,485,303,324]" italics="true" pageId="1" pageNumber="296">Phlugiolopsis henryi</emphasis>
<bibRefCitation id="5A394B6A245DC62DFE78FED8FDFDFEDE" author="Zeuner, F. E." box="[491,618,303,324]" pageId="1" pageNumber="296" pagination="76 - 84" refId="ref3868" refString="Zeuner, F. E. (1940) Phlugiolopsis henryi, n. g., n. sp., a new tettigoniid, and other Saltatoria (Orthop.) from the Royal Botanic Gardens, Kew. Journal of the Society for British Entomology, 2, 76 - 84." type="journal article" year="1940">Zeuner, 1940</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
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<heading id="655F81F7245DC62DFF04FE65FCFCFE36" bold="true" box="[151,875,402,428]" fontSize="11" level="1" pageId="1" pageNumber="296" reason="1">
<emphasis id="0CDCEA89245DC62DFF04FE65FCFCFE36" bold="true" box="[151,875,402,428]" pageId="1" pageNumber="296">
<taxonomicName id="F9A84D18245DC62DFF04FE65FEABFE36" ID-CoL="8NXHR" authorityName="Zeuner" authorityYear="1940" box="[151,316,402,428]" class="Insecta" family="Tettigoniidae" genus="Phlugiolopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="0CDCEA89245DC62DFF04FE65FEABFE36" bold="true" box="[151,316,402,428]" italics="true" pageId="1" pageNumber="296">Phlugiolopsis</emphasis>
</taxonomicName>
(
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<emphasis id="0CDCEA89245DC62DFEDFFE65FE67FE36" bold="true" box="[332,496,402,428]" italics="true" pageId="1" pageNumber="296">Phlugiolopsis</emphasis>
</taxonomicName>
?)
<emphasis id="0CDCEA89245DC62DFD83FE65FD0AFE36" bold="true" box="[528,669,402,428]" italics="true" pageId="1" pageNumber="296">taiwanensis</emphasis>
sp. nov. (
<figureCitation id="A6932A1E245DC62DFC89FE65FCF6FE36" box="[794,865,402,428]" captionStart="FIGURE 1" captionStartId="1.[151,244,1975,1997]" captionTargetBox="[150,1436,1103,1961]" captionTargetId="figure@1.[150,1436,1103,1961]" captionTargetPageId="1" captionText="FIGURE 1. Phlugiolopsis (Phlugiolopsis?) taiwanensis sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F. apex of abdomen: C. lateral view; D. dorsal view; E. ventral view; F. dorso-lateral view." httpUri="https://zenodo.org/record/2588609/files/figure.png" pageId="1" pageNumber="296">Fig. 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="76B26510245DC62EFF04FE42FE0FFEA3" lastPageId="2" lastPageNumber="297" pageId="1" pageNumber="296" type="description">
<paragraph id="3E17369B245DC62DFF04FE42FD61FD8B" blockId="1.[151,1436,402,1049]" pageId="1" pageNumber="296">
<emphasis id="0CDCEA89245DC62DFF04FE42FEFDFE56" bold="true" box="[151,362,437,460]" pageId="1" pageNumber="296">Description. Male.</emphasis>
Body size relatively small. Fastigium verticis conical, narrower than antennal scape, apex obtusely rounded, with a longitudinal sulcus on dorsal surface. Eyes oval, protruding forward. Apical segment of maxillary palpus slightly longer than subapical one, apex slightly swollen.
</paragraph>
<paragraph id="3E17369B245DC62DFF54FDEAFC7AFDCD" blockId="1.[151,1436,402,1049]" pageId="1" pageNumber="296">
Pronotum elongate, anterior margin slightly straight while posterior margin arc-shaped (
<figureCitation id="A6932A1E245DC62DFB1AFDEAFB77FDAE" box="[1161,1248,541,564]" captionStart="FIGURE 1" captionStartId="1.[151,244,1975,1997]" captionTargetBox="[150,1436,1103,1961]" captionTargetId="figure@1.[150,1436,1103,1961]" captionTargetPageId="1" captionText="FIGURE 1. Phlugiolopsis (Phlugiolopsis?) taiwanensis sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F. apex of abdomen: C. lateral view; D. dorsal view; E. ventral view; F. dorso-lateral view." httpUri="https://zenodo.org/record/2588609/files/figure.png" pageId="1" pageNumber="296">Fig. 1A</figureCitation>
); pronotal lateral lobe longer than deep, posterior margin oblique, humeral sinus absent (
<figureCitation id="A6932A1E245DC62DFC1FFDB7FC48FDCD" box="[908,991,576,599]" captionStart="FIGURE 1" captionStartId="1.[151,244,1975,1997]" captionTargetBox="[150,1436,1103,1961]" captionTargetId="figure@1.[150,1436,1103,1961]" captionTargetPageId="1" captionText="FIGURE 1. Phlugiolopsis (Phlugiolopsis?) taiwanensis sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F. apex of abdomen: C. lateral view; D. dorsal view; E. ventral view; F. dorso-lateral view." httpUri="https://zenodo.org/record/2588609/files/figure.png" pageId="1" pageNumber="296">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="3E17369B245DC62DFF54FD95FD40FD06" blockId="1.[151,1436,402,1049]" pageId="1" pageNumber="296">
Tegmina short, surpassing posterior margin of pronotum and reaching posterior margin of 2nd abdominal tergite, apices obtusely rounded; hind wings absent (
<figureCitation id="A6932A1E245DC62DFDE6FD72FD5DFD06" box="[629,714,645,668]" captionStart="FIGURE 1" captionStartId="1.[151,244,1975,1997]" captionTargetBox="[150,1436,1103,1961]" captionTargetId="figure@1.[150,1436,1103,1961]" captionTargetPageId="1" captionText="FIGURE 1. Phlugiolopsis (Phlugiolopsis?) taiwanensis sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F. apex of abdomen: C. lateral view; D. dorsal view; E. ventral view; F. dorso-lateral view." httpUri="https://zenodo.org/record/2588609/files/figure.png" pageId="1" pageNumber="296">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="3E17369B245DC62DFF54FD5FFEF3FCD3" blockId="1.[151,1436,402,1049]" pageId="1" pageNumber="296">All femora unarmed ventrally, genicular lobes with apices obtuse. Fore coxae with 1 small spine; fore tibiae with 5– 6 spines respectively on inner and outer sides of ventral surface, tympanal organs open on both sides. Middle tibiae with 4–6 spines on both sides of ventral surface. Hind tibiae with 2 internal spines and 5 external spines on ventral surface, and dorsal surface with 22–27 spines separately on both sides, bearing two pairs of ventral apical spurs and one pair of dorsal apical spurs.</paragraph>
<paragraph id="3E17369B245DC62DFF54FCA2FF53FB83" blockId="1.[151,1436,402,1049]" pageId="1" pageNumber="296">
Centre of 10th abdominal tergite concave (
<figureCitation id="A6932A1E245DC62DFD06FCA2FD7DFCF6" box="[661,746,853,876]" captionStart="FIGURE 1" captionStartId="1.[151,244,1975,1997]" captionTargetBox="[150,1436,1103,1961]" captionTargetId="figure@1.[150,1436,1103,1961]" captionTargetPageId="1" captionText="FIGURE 1. Phlugiolopsis (Phlugiolopsis?) taiwanensis sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F. apex of abdomen: C. lateral view; D. dorsal view; E. ventral view; F. dorso-lateral view." httpUri="https://zenodo.org/record/2588609/files/figure.png" pageId="1" pageNumber="296">Fig. 1D</figureCitation>
). Epiproct tongue-shaped, hidden under 10th abdominal tergite. Basal area of cercus stout, subsequently divided into two lobes, dorsal lobe slightly short, nearly rectangular, somewhat depressed, outer lateral angle triangular projecting backwards, terminal acute; ventral lobe broad basally, tapering, 1/3 apical area narrow, up-curved and twisted, terminal obtusely rounded (
<figureCitation id="A6932A1E245DC62DFC1CFC4AFC5DFC4E" box="[911,970,957,980]" captionStart="FIGURE 1" captionStartId="1.[151,244,1975,1997]" captionTargetBox="[150,1436,1103,1961]" captionTargetId="figure@1.[150,1436,1103,1961]" captionTargetPageId="1" captionText="FIGURE 1. Phlugiolopsis (Phlugiolopsis?) taiwanensis sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F. apex of abdomen: C. lateral view; D. dorsal view; E. ventral view; F. dorso-lateral view." httpUri="https://zenodo.org/record/2588609/files/figure.png" pageId="1" pageNumber="296">Fig. 1</figureCitation>
C–F). Basal area of subgenital plate broad, apical area somewhat narrow, posterior margin obtusely rounded, with conical styli on ventral surface of subapex (
<figureCitation id="A6932A1E245DC62DFAE3FC17FF21FB83" captionStart="FIGURE 1" captionStartId="1.[151,244,1975,1997]" captionTargetBox="[150,1436,1103,1961]" captionTargetId="figure@1.[150,1436,1103,1961]" captionTargetPageId="1" captionText="FIGURE 1. Phlugiolopsis (Phlugiolopsis?) taiwanensis sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F. apex of abdomen: C. lateral view; D. dorsal view; E. ventral view; F. dorso-lateral view." httpUri="https://zenodo.org/record/2588609/files/figure.png" pageId="1" pageNumber="296">Fig. 1E</figureCitation>
).
</paragraph>
<caption id="6AD76613245DC62DFF04F840FC0DF874" httpUri="https://zenodo.org/record/2588609/files/figure.png" pageId="1" pageNumber="296" startId="1.[151,244,1975,1997]" targetBox="[150,1436,1103,1961]" targetPageId="1">
<paragraph id="3E17369B245DC62DFF04F840FC0DF874" blockId="1.[151,1436,1975,2031]" pageId="1" pageNumber="296">
<emphasis id="0CDCEA89245DC62DFF04F840FE9BF857" bold="true" box="[151,268,1975,1997]" pageId="1" pageNumber="296">FIGURE 1.</emphasis>
<taxonomicName id="F9A84D18245DC62DFE81F840FE01F856" authorityName="Zeuner" authorityYear="1940" box="[274,406,1975,1996]" class="Insecta" family="Tettigoniidae" genus="Phlugiolopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="0CDCEA89245DC62DFE81F840FE01F856" box="[274,406,1975,1996]" italics="true" pageId="1" pageNumber="296">Phlugiolopsis</emphasis>
</taxonomicName>
(
<taxonomicName id="F9A84D18245DC62DFE37F840FDBFF856" authorityName="Zeuner" authorityYear="1940" box="[420,552,1975,1996]" class="Insecta" family="Tettigoniidae" genus="Phlugiolopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="0CDCEA89245DC62DFE37F840FDBFF856" box="[420,552,1975,1996]" italics="true" pageId="1" pageNumber="296">Phlugiolopsis</emphasis>
</taxonomicName>
?)
<emphasis id="0CDCEA89245DC62DFDD3F840FD26F856" box="[576,689,1975,1996]" italics="true" pageId="1" pageNumber="296">taiwanensis</emphasis>
<emphasis id="0CDCEA89245DC62DFD24F840FC94F857" bold="true" box="[695,771,1975,1997]" pageId="1" pageNumber="296">sp. nov.</emphasis>
, male:
<emphasis id="0CDCEA89245DC62DFCD8F840FCEAF857" bold="true" box="[843,893,1975,1997]" pageId="1" pageNumber="296">A–B.</emphasis>
pronotum:
<emphasis id="0CDCEA89245DC62DFC7EF840FB92F857" bold="true" box="[1005,1029,1975,1997]" pageId="1" pageNumber="296">A.</emphasis>
dorsal view;
<emphasis id="0CDCEA89245DC62DFB15F840FB0BF857" bold="true" box="[1158,1180,1975,1997]" pageId="1" pageNumber="296">B.</emphasis>
lateral view;
<emphasis id="0CDCEA89245DC62DFA8CF840FAD8F857" bold="true" box="[1311,1359,1975,1997]" pageId="1" pageNumber="296">C–F.</emphasis>
apex of abdomen:
<emphasis id="0CDCEA89245DC62DFF68F82EFE84F875" bold="true" box="[251,275,2009,2031]" pageId="1" pageNumber="296">C.</emphasis>
lateral view;
<emphasis id="0CDCEA89245DC62DFE05F82EFE3AF875" bold="true" box="[406,429,2009,2031]" pageId="1" pageNumber="296">D.</emphasis>
dorsal view;
<emphasis id="0CDCEA89245DC62DFDBCF82EFDD2F875" bold="true" box="[559,581,2009,2031]" pageId="1" pageNumber="296">E.</emphasis>
ventral view;
<emphasis id="0CDCEA89245DC62DFD5DF82EFD49F875" bold="true" box="[718,734,2009,2031]" pageId="1" pageNumber="296">F.</emphasis>
dorso-lateral view.
</paragraph>
</caption>
<paragraph id="3E17369B245EC62EFF54FF6FFDC9FE8D" blockId="2.[151,1436,152,591]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54FF6FFEDEFF35" bold="true" box="[199,329,152,175]" pageId="2" pageNumber="297">Coloration.</emphasis>
Body yellowish brown. Dorsal area of head with 4 longitudinal black brown stripes. Disc of pronotum with a broad longitudinal brown stripe, apical half slightly expended, 2/3 anterior area of stripe black brown on lateral margins. Spines of all tibiae light brown. Apices of all femora and all tarsi light brown. Lateral margins of abdomen with one brown longitudinal stripe respectively.
</paragraph>
<paragraph id="3E17369B245EC62EFF54FED5FE0FFEA3" blockId="2.[151,1436,152,591]" box="[199,408,290,313]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54FED5FEB6FEA3" bold="true" box="[199,289,290,313]" pageId="2" pageNumber="297">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="76B26510245EC62EFF54FEB2FEDBFEE5" pageId="2" pageNumber="297" type="materials_examined">
<paragraph id="3E17369B245EC62EFF54FEB2FEDBFEE5" blockId="2.[151,1436,152,591]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54FEB2FE3FFEC6" bold="true" box="[199,424,325,348]" pageId="2" pageNumber="297">Material examined.</emphasis>
<materialsCitation id="8EC03CC6245EC62EFE22FEB2FB0BFEC6" ID-GBIF-Occurrence="2012743362" box="[433,1180,325,348]" collectingDate="1931-08" collectorName="Powei Chen." country="Taiwan" location="Yilan" pageId="2" pageNumber="297" specimenCount="1" stateProvince="Yilan" typeStatus="holotype">
<typeStatus id="E1138839245EC62EFE22FEB2FD8FFEC6" box="[433,536,325,348]" pageId="2" pageNumber="297" type="holotype">Holotype</typeStatus>
: male,
<collectingRegion id="FC6CF879245EC62EFDFAFEB2FD32FEC6" box="[617,677,325,348]" country="Taiwan" name="Yilan" pageId="2" pageNumber="297">Yilan</collectingRegion>
,
<collectingCountry id="46BF760B245EC62EFD20FEB2FC93FEC6" box="[691,772,325,348]" name="Taiwan" pageId="2" pageNumber="297">Taiwan</collectingCountry>
,
<collectingDate id="5A52E9B3245EC62EFC82FEB2FC12FEC6" box="[785,901,325,348]" pageId="2" pageNumber="297" value="1931-08">
<date id="4A16105B245EC62EFC82FEB2FC12FEC6" box="[785,901,325,348]" pageId="2" pageNumber="297" value="1931-08">31 August</date>
</collectingDate>
, 2018, coll.
<collectorName id="935D534D245EC62EFB83FEB2FB0BFEC6" box="[1040,1180,325,348]" pageId="2" pageNumber="297">Powei Chen.</collectorName>
</materialsCitation>
<typeStatus id="E1138839245EC62EFB37FEB2FA9AFEC6" box="[1188,1293,325,348]" pageId="2" pageNumber="297" type="paratype">Paratype</typeStatus>
:
<specimenCount id="28AEFD12245EC62EFA82FEB2FABAFEC6" box="[1297,1325,325,348]" pageId="2" pageNumber="297" type="male">1♂</specimenCount>
, the other data as
<typeStatus id="E1138839245EC62EFF74FE9FFEDFFEE5" box="[231,328,360,383]" pageId="2" pageNumber="297" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="76B26510245EC62EFF54FE7DFA80FE3B" box="[199,1303,393,417]" pageId="2" pageNumber="297" type="description">
<paragraph id="3E17369B245EC62EFF54FE7DFA80FE3B" blockId="2.[151,1436,152,591]" box="[199,1303,393,417]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54FE7DFE24FE3B" bold="true" box="[199,435,394,417]" pageId="2" pageNumber="297">Measurements(mm).</emphasis>
Body:
<specimenCount id="28AEFD12245EC62EFD92FE7DFD82FE3B" box="[513,533,394,417]" pageId="2" pageNumber="297" type="male">♂</specimenCount>
5.8–6.6; pronotum:
<specimenCount id="28AEFD12245EC62EFD7CFE7DFC94FE3B" box="[751,771,394,417]" pageId="2" pageNumber="297" type="male">♂</specimenCount>
4.1; tegmina:
<specimenCount id="28AEFD12245EC62EFC0EFE7DFC26FE3B" box="[925,945,394,417]" pageId="2" pageNumber="297" type="male">♂</specimenCount>
2.2–2.4; hind femora:
<specimenCount id="28AEFD12245EC62EFB36FE7DFB2EFE3B" box="[1189,1209,394,417]" pageId="2" pageNumber="297" type="male">♂</specimenCount>
7.9–8.3.
</paragraph>
</subSubSection>
<subSubSection id="76B26510245EC62EFF54FE5AFD6CFDB6" pageId="2" pageNumber="297" type="discussion">
<paragraph id="3E17369B245EC62EFF54FE5AFD6CFDB6" blockId="2.[151,1436,152,591]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54FE5AFED1FE5E" bold="true" box="[199,326,429,452]" pageId="2" pageNumber="297">Discussion.</emphasis>
This new species can be distinguished from known species of
<taxonomicName id="F9A84D18245EC62EFB9BFE5AFB0DFE5E" authorityName="Zeuner" authorityYear="1940" box="[1032,1178,429,452]" class="Insecta" family="Tettigoniidae" genus="Phlugiolopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="0CDCEA89245EC62EFB9BFE5AFB0DFE5E" box="[1032,1178,429,452]" italics="true" pageId="2" pageNumber="297">Phlugiolopsis</emphasis>
</taxonomicName>
by: basal area of male cercus stout, subsequently divided into two lobes, dorsal lobe slightly short, nearly rectangular, while ventral lobe broad basally, tapering, 1/3 apical area narrow, bent upwards and twisted; posterior margin of subgenital plate obtusely rounded, with conical styli on ventral surface of subapex.
</paragraph>
</subSubSection>
<subSubSection id="76B26510245EC62EFF54FDCFFC74FDD5" box="[199,995,568,591]" pageId="2" pageNumber="297" type="etymology">
<paragraph id="3E17369B245EC62EFF54FDCFFC74FDD5" blockId="2.[151,1436,152,591]" box="[199,995,568,591]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54FDCFFED0FDD5" bold="true" box="[199,327,568,591]" pageId="2" pageNumber="297">Etymology.</emphasis>
The name of the new species is derived from the
<typeStatus id="E1138839245EC62EFCCAFDCFFC10FDD5" box="[857,903,568,591]" pageId="2" pageNumber="297">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

171
data/B6/01/87/B601878D245EC62EFF04FC8AFDCAF81B.xml Normal file
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<mods:title id="D0F997CA40E8F67F8DBD87B4C8295D9D">Two new species of the tribe Meconematini (Orthoptera: Tettigoniidae: Meconematinae) from Taiwan</mods:title>
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<treatment id="B601878D245EC62EFF04FC8AFDCAF81B" ID-DOI="http://doi.org/10.5281/zenodo.5933310" ID-GBIF-Taxon="154366339" ID-Zenodo-Dep="5933310" LSID="urn:lsid:plazi:treatment:B601878D245EC62EFF04FC8AFDCAF81B" httpUri="http://treatment.plazi.org/id/B601878D245EC62EFF04FC8AFDCAF81B" lastPageNumber="297" pageId="2" pageNumber="297">
<subSubSection id="76B26510245EC62EFF04FC8AFC83FC0D" box="[151,788,893,919]" pageId="2" pageNumber="297" type="nomenclature">
<paragraph id="3E17369B245EC62EFF04FC8AFC83FC0D" blockId="2.[151,1436,893,1921]" box="[151,788,893,919]" pageId="2" pageNumber="297">
<heading id="655F81F7245EC62EFF04FC8AFC83FC0D" bold="true" box="[151,788,893,919]" fontSize="11" level="1" pageId="2" pageNumber="297" reason="1">
<emphasis id="0CDCEA89245EC62EFF04FC8AFC83FC0D" bold="true" box="[151,788,893,919]" pageId="2" pageNumber="297">
<taxonomicName id="F9A84D18245EC62EFF04FC8AFDD9FC0D" ID-CoL="8PWFC" authority="Wang &amp; Chen &amp; Shi, 2019" authorityName="Wang &amp; Chen &amp; Shi" authorityYear="2019" box="[151,590,893,919]" class="Insecta" family="Tettigoniidae" genus="Xiphidiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="subGenus" status="sp. nov." subGenus="Xiphidiopsis">
<emphasis id="0CDCEA89245EC62EFF04FC8AFEBAFC0D" bold="true" box="[151,301,893,919]" italics="true" pageId="2" pageNumber="297">Xiphidiopsis</emphasis>
(
<emphasis id="0CDCEA89245EC62EFEADFC8AFE44FC0D" bold="true" box="[318,467,893,919]" italics="true" pageId="2" pageNumber="297">Xiphidiopsis</emphasis>
)
<emphasis id="0CDCEA89245EC62EFE77FC8AFDD9FC0D" bold="true" box="[484,590,893,919]" italics="true" pageId="2" pageNumber="297">trifoliata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17EF57F2245EC62EFDC6FC8AFD25FC0D" box="[597,690,893,919]" pageId="2" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="A6932A1E245EC62EFD51FC8AFC9CFC0D" box="[706,779,893,919]" captionStart="FIGURE 2" captionStartId="3.[151,244,1435,1457]" captionTargetBox="[155,1432,191,1413]" captionTargetId="figure@3.[155,1432,191,1413]" captionTargetPageId="3" captionText="FIGURE 2. Xiphidiopsis (Xiphidiopsis) trifoliata sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F, I apex of abdomen: C. lateral view; D. apico-lateral view; E. ventral view; F. apical view; I. latero-ventral view; G. apical area of left cercus in lateral-anterior view; H. subgenital plate in ventral view." httpUri="https://zenodo.org/record/2588611/files/figure.png" pageId="2" pageNumber="297">Fig. 2</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="76B26510245EC62EFF04FC57FE0FF9F6" pageId="2" pageNumber="297" type="description">
<paragraph id="3E17369B245EC62EFF04FC57FB1DFC66" blockId="2.[151,1436,893,1921]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF04FC57FEFBFC2D" bold="true" box="[151,364,928,951]" pageId="2" pageNumber="297">Description. Male.</emphasis>
Body small, medium size for the tribe
<taxonomicName id="F9A84D18245EC62EFCBFFC57FC5EFC2D" authorityName="Burmeister" authorityYear="1838" box="[812,969,928,951]" class="Insecta" family="Tettigoniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="tribe" tribe="Meconematini">Meconematini</taxonomicName>
. Fastigium verticis conical, narrower than antennal scape, broadly blunt apically, with a small sulcus on dorsal surface. Eyes oval and distinctly protruding forward. Apical segment of maxillary palpus feebly longer than subapical one, slightly swelled apically.
</paragraph>
<paragraph id="3E17369B245EC62EFF54FBFFFC39FBDB" blockId="2.[151,1436,893,1921]" pageId="2" pageNumber="297">
Anterior margin of pronotum straight, posterior margin obtusely rounded (
<figureCitation id="A6932A1E245EC62EFC61FBFFFBDCFB85" box="[1010,1099,1032,1055]" captionStart="FIGURE 2" captionStartId="3.[151,244,1435,1457]" captionTargetBox="[155,1432,191,1413]" captionTargetId="figure@3.[155,1432,191,1413]" captionTargetPageId="3" captionText="FIGURE 2. Xiphidiopsis (Xiphidiopsis) trifoliata sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F, I apex of abdomen: C. lateral view; D. apico-lateral view; E. ventral view; F. apical view; I. latero-ventral view; G. apical area of left cercus in lateral-anterior view; H. subgenital plate in ventral view." httpUri="https://zenodo.org/record/2588611/files/figure.png" pageId="2" pageNumber="297">Fig. 2A</figureCitation>
); lateral lobe higher than long, posterior margin undulating and oblique, humeral sinus shallow (
<figureCitation id="A6932A1E245EC62EFCDDFBDDFC36FBDB" box="[846,929,1066,1089]" captionStart="FIGURE 2" captionStartId="3.[151,244,1435,1457]" captionTargetBox="[155,1432,191,1413]" captionTargetId="figure@3.[155,1432,191,1413]" captionTargetPageId="3" captionText="FIGURE 2. Xiphidiopsis (Xiphidiopsis) trifoliata sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F, I apex of abdomen: C. lateral view; D. apico-lateral view; E. ventral view; F. apical view; I. latero-ventral view; G. apical area of left cercus in lateral-anterior view; H. subgenital plate in ventral view." httpUri="https://zenodo.org/record/2588611/files/figure.png" pageId="2" pageNumber="297">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="3E17369B245EC62EFF54FBBAFAC1FBFE" blockId="2.[151,1436,893,1921]" box="[199,1366,1101,1124]" pageId="2" pageNumber="297">Tegmina narrow and long, surpassing apices of hind femora; hind wings distinctly extending behind tegmina.</paragraph>
<paragraph id="3E17369B245EC62EFF54FB87FD68FB75" blockId="2.[151,1436,893,1921]" pageId="2" pageNumber="297">All femora without spines on ventral surface, genicular lobes blunt apically. Fore coxae with a small spine; fore tibiae with 5 inner and 6 outer spines on ventral surface; tympanal organs open on both sides, ovoid. Middle tibiae with 5–6 spines respectively on both sides of ventral surface. Hind tibiae with 26–30 on both dorsal sides as well as two pairs of ventral apical spurs and one pair of dorsal apical spurs.</paragraph>
<paragraph id="3E17369B245EC62EFF54FB0DFEAAFA7B" blockId="2.[151,1436,893,1921]" pageId="2" pageNumber="297">
10th abdominal tergite with a distinct notch on posterior margin, in the middle bearing a robust asymmetric posterior process which bent downwards, apical half slightly extended, posterior margin blunted (
<figureCitation id="A6932A1E245EC62EFBD0FAEAFB03FAAE" box="[1091,1172,1309,1332]" captionStart="FIGURE 2" captionStartId="3.[151,244,1435,1457]" captionTargetBox="[155,1432,191,1413]" captionTargetId="figure@3.[155,1432,191,1413]" captionTargetPageId="3" captionText="FIGURE 2. Xiphidiopsis (Xiphidiopsis) trifoliata sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F, I apex of abdomen: C. lateral view; D. apico-lateral view; E. ventral view; F. apical view; I. latero-ventral view; G. apical area of left cercus in lateral-anterior view; H. subgenital plate in ventral view." httpUri="https://zenodo.org/record/2588611/files/figure.png" pageId="2" pageNumber="297">Fig. 2F</figureCitation>
); left lateral margin near middle area with a spine-shaped process, terminal obtusely rounded (
<figureCitation id="A6932A1E245EC62EFCEAFAB7FC58FACD" box="[889,975,1344,1367]" captionStart="FIGURE 2" captionStartId="3.[151,244,1435,1457]" captionTargetBox="[155,1432,191,1413]" captionTargetId="figure@3.[155,1432,191,1413]" captionTargetPageId="3" captionText="FIGURE 2. Xiphidiopsis (Xiphidiopsis) trifoliata sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F, I apex of abdomen: C. lateral view; D. apico-lateral view; E. ventral view; F. apical view; I. latero-ventral view; G. apical area of left cercus in lateral-anterior view; H. subgenital plate in ventral view." httpUri="https://zenodo.org/record/2588611/files/figure.png" pageId="2" pageNumber="297">Fig. 2D</figureCitation>
); cercus robust at base, feebly compressed; 2/3 basal area bent inner-dorsally distinctly (
<figureCitation id="A6932A1E245EC62EFDEFFA95FD47FAE3" box="[636,720,1378,1401]" captionStart="FIGURE 2" captionStartId="3.[151,244,1435,1457]" captionTargetBox="[155,1432,191,1413]" captionTargetId="figure@3.[155,1432,191,1413]" captionTargetPageId="3" captionText="FIGURE 2. Xiphidiopsis (Xiphidiopsis) trifoliata sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F, I apex of abdomen: C. lateral view; D. apico-lateral view; E. ventral view; F. apical view; I. latero-ventral view; G. apical area of left cercus in lateral-anterior view; H. subgenital plate in ventral view." httpUri="https://zenodo.org/record/2588611/files/figure.png" pageId="2" pageNumber="297">Fig. 2E</figureCitation>
), 1/3 apical area cylindrical, curved dorsad, apical area extending bilaterally, forming three lobes (
<figureCitation id="A6932A1E245EC62EFE69FA72FDC4FA06" box="[506,595,1413,1436]" captionStart="FIGURE 2" captionStartId="3.[151,244,1435,1457]" captionTargetBox="[155,1432,191,1413]" captionTargetId="figure@3.[155,1432,191,1413]" captionTargetPageId="3" captionText="FIGURE 2. Xiphidiopsis (Xiphidiopsis) trifoliata sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F, I apex of abdomen: C. lateral view; D. apico-lateral view; E. ventral view; F. apical view; I. latero-ventral view; G. apical area of left cercus in lateral-anterior view; H. subgenital plate in ventral view." httpUri="https://zenodo.org/record/2588611/files/figure.png" pageId="2" pageNumber="297">Fig. 2G</figureCitation>
). Subgenital plate slightly trapezoidal, basal area broad, apical area concave dorsally, posterior margin narrowly rounded (
<figureCitation id="A6932A1E245EC62EFD02FA5FFD7CFA25" box="[657,747,1448,1471]" captionStart="FIGURE 2" captionStartId="3.[151,244,1435,1457]" captionTargetBox="[155,1432,191,1413]" captionTargetId="figure@3.[155,1432,191,1413]" captionTargetPageId="3" captionText="FIGURE 2. Xiphidiopsis (Xiphidiopsis) trifoliata sp. nov., male: A–B. pronotum: A. dorsal view; B. lateral view; C–F, I apex of abdomen: C. lateral view; D. apico-lateral view; E. ventral view; F. apical view; I. latero-ventral view; G. apical area of left cercus in lateral-anterior view; H. subgenital plate in ventral view." httpUri="https://zenodo.org/record/2588611/files/figure.png" pageId="2" pageNumber="297">Fig. 2H</figureCitation>
). Subapex of subgenital plate with stout, compressed styli on lateral margins.
</paragraph>
<paragraph id="3E17369B245EC62EFF54FA1AFE35F9D3" blockId="2.[151,1436,893,1921]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54FA1AFEDEF99E" bold="true" box="[199,329,1517,1540]" pageId="2" pageNumber="297">Coloration.</emphasis>
Body yellowish brown, probably green when alive. Eyes brown. All third tarsi and genicular lobes of all femora with apices black. Spines of fore and middle tibiae black brown on ventral surface while spines of hind tibiae with apices black brown.
</paragraph>
<paragraph id="3E17369B245EC62EFF54F9A2FE0FF9F6" blockId="2.[151,1436,893,1921]" box="[199,408,1621,1644]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54F9A2FEB6F9F6" bold="true" box="[199,289,1621,1644]" pageId="2" pageNumber="297">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="76B26510245EC62EFF54F98FFE8FF92B" pageId="2" pageNumber="297" type="materials_examined">
<paragraph id="3E17369B245EC62EFF54F98FFE8FF92B" blockId="2.[151,1436,893,1921]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54F98FFE32F915" bold="true" box="[199,421,1656,1679]" pageId="2" pageNumber="297">Material examined.</emphasis>
<materialsCitation id="8EC03CC6245EC62EFE3FF98FFBFCF915" ID-GBIF-Occurrence="2012743363" box="[428,1131,1656,1679]" collectingDate="1922-07" collectorName="Tien Hsieh." country="Taiwan" location="Taidong" pageId="2" pageNumber="297" specimenCount="1" stateProvince="Taitung" typeStatus="holotype">
<typeStatus id="E1138839245EC62EFE3FF98FFD84F915" box="[428,531,1656,1679]" pageId="2" pageNumber="297" type="holotype">Holotype</typeStatus>
: male,
<collectingRegion id="FC6CF879245EC62EFDCDF98FFD2FF915" box="[606,696,1656,1679]" country="Taiwan" name="Taitung" pageId="2" pageNumber="297">Taidong</collectingRegion>
,
<collectingCountry id="46BF760B245EC62EFD51F98FFC84F915" box="[706,787,1656,1679]" name="Taiwan" pageId="2" pageNumber="297">Taiwan</collectingCountry>
,
<collectingDate id="5A52E9B3245EC62EFC8EF98FFCFAF915" box="[797,877,1656,1679]" pageId="2" pageNumber="297" value="1922-07">
<date id="4A16105B245EC62EFC8EF98FFCFAF915" box="[797,877,1656,1679]" pageId="2" pageNumber="297" value="1922-07">22 July</date>
</collectingDate>
, 2016, coll.
<collectorName id="935D534D245EC62EFC7DF98FFBFCF915" box="[1006,1131,1656,1679]" pageId="2" pageNumber="297">Tien Hsieh.</collectorName>
</materialsCitation>
<typeStatus id="E1138839245EC62EFBE2F98FFB45F915" box="[1137,1234,1656,1679]" pageId="2" pageNumber="297" type="paratype">Paratype</typeStatus>
:
<specimenCount id="28AEFD12245EC62EFB4FF98FFB68F915" box="[1244,1279,1656,1679]" pageId="2" pageNumber="297" type="male">1♂</specimenCount>
, the other data as
<typeStatus id="E1138839245EC62EFF27F96DFE82F92B" box="[180,277,1690,1713]" pageId="2" pageNumber="297" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="76B26510245EC62EFF54F94AFAA5F94E" box="[199,1330,1724,1748]" pageId="2" pageNumber="297" type="description">
<paragraph id="3E17369B245EC62EFF54F94AFAA5F94E" blockId="2.[151,1436,893,1921]" box="[199,1330,1724,1748]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54F94AFE24F94E" bold="true" box="[199,435,1725,1748]" pageId="2" pageNumber="297">Measurements(mm).</emphasis>
Body:
<specimenCount id="28AEFD12245EC62EFD92F94AFD82F94E" box="[513,533,1725,1748]" pageId="2" pageNumber="297" type="male">♂</specimenCount>
10–10.3; pronotum:
<specimenCount id="28AEFD12245EC62EFD65F94AFC9DF94E" box="[758,778,1725,1748]" pageId="2" pageNumber="297" type="male">♂</specimenCount>
3.8; tegmina:
<specimenCount id="28AEFD12245EC62EFC37F94AFC2FF94E" box="[932,952,1725,1748]" pageId="2" pageNumber="297" type="male">♂</specimenCount>
16.3–18.2; hind femora:
<specimenCount id="28AEFD12245EC62EFB54F94AFB4CF94E" box="[1223,1243,1725,1748]" pageId="2" pageNumber="297" type="male">♂</specimenCount>
9.6–10.
</paragraph>
</subSubSection>
<subSubSection id="76B26510245EC62EFF54F917FB13F8A6" pageId="2" pageNumber="297" type="discussion">
<paragraph id="3E17369B245EC62EFF54F917FB13F8A6" blockId="2.[151,1436,893,1921]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54F917FED1F96D" bold="true" box="[199,326,1760,1783]" pageId="2" pageNumber="297">Discussion.</emphasis>
This new species differs from known species of
<taxonomicName id="F9A84D18245EC62EFCC5F917FC4CF96D" authorityName="Redtenbacher" authorityYear="1891" box="[854,987,1760,1783]" class="Insecta" family="Tettigoniidae" genus="Xiphidiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="0CDCEA89245EC62EFCC5F917FC4CF96D" box="[854,987,1760,1783]" italics="true" pageId="2" pageNumber="297">Xiphidiopsis</emphasis>
</taxonomicName>
in: 10th abdominal tergite of male with a robust asymmetric posterior process; basal area of cercus slightly compressed, 2/3 basal area bent inner-dorsally distinctly, 1/3 apical area curved dorsad, apical area extending bilaterally, forming three lobes.
</paragraph>
</subSubSection>
<subSubSection id="76B26510245EC62EFF54F8BFFDCAF81B" pageId="2" pageNumber="297" type="etymology">
<paragraph id="3E17369B245EC62EFF54F8BFFDCAF81B" blockId="2.[151,1436,893,1921]" pageId="2" pageNumber="297">
<emphasis id="0CDCEA89245EC62EFF54F8BFFED0F8C5" bold="true" box="[199,327,1864,1887]" pageId="2" pageNumber="297">Etymology.</emphasis>
The name of the new species is derived from apex of male cercus divided into three lobes. Latin “
<emphasis id="0CDCEA89245EC62EFAFDF8BFFA07F8C5" box="[1390,1424,1864,1887]" italics="true" pageId="2" pageNumber="297">tri-</emphasis>
” means three, and Latin “
<emphasis id="0CDCEA89245EC62EFE0FF89DFE28F81B" box="[412,447,1898,1921]" italics="true" pageId="2" pageNumber="297">foli</emphasis>
” means foliar.
</paragraph>
</subSubSection>
</treatment>
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<heading id="655F81F7245EC62EFF04FD57FD15FD20" bold="true" box="[151,642,671,698]" fontSize="11" level="1" pageId="2" pageNumber="297" reason="1">
<emphasis id="0CDCEA89245EC62EFF04FD57FD15FD20" bold="true" box="[151,642,671,698]" pageId="2" pageNumber="297">
Genus
<taxonomicName id="F9A84D18245EC62EFF7EFD68FD15FD20" ID-CoL="8P2XH" authority="Redtenbacher, 1891" authorityName="Redtenbacher" authorityYear="1891" box="[237,642,671,698]" class="Insecta" family="Tettigoniidae" genus="Xiphidiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="0CDCEA89245EC62EFF7EFD68FE14FD23" bold="true" box="[237,387,671,697]" italics="true" pageId="2" pageNumber="297">Xiphidiopsis</emphasis>
<bibRefCitation id="5A394B6A245EC62EFE18FD57FD15FD20" author="Redtenbacher, J." box="[395,642,672,698]" pageId="2" pageNumber="297" pagination="315 - 562" refId="ref3281" refString="Redtenbacher, J. (1891) Monographie der Conocephaliden. Ferhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 41, 315 - 562." type="journal article" year="1891">Redtenbacher, 1891</bibRefCitation>
</taxonomicName>
</emphasis>
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<taxonomicName id="F9A84D18245EC62EFF04FD35FE46FD4D" ID-CoL="8P2XH" authority="Redtenbacher, 1891" authorityName="Redtenbacher" authorityYear="1891" box="[151,465,706,727]" class="Insecta" family="Tettigoniidae" genus="Xiphidiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="0CDCEA89245EC62EFF04FD35FE98FD4D" box="[151,271,706,727]" italics="true" pageId="2" pageNumber="297">Xiphidiopsis</emphasis>
<bibRefCitation id="5A394B6A245EC62EFE85FD35FE46FD4D" author="Redtenbacher, J." box="[278,465,706,727]" pageId="2" pageNumber="297" pagination="315 - 562" refId="ref3281" refString="Redtenbacher, J. (1891) Monographie der Conocephaliden. Ferhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 41, 315 - 562." type="journal article" year="1891">Redtenbacher,1891</bibRefCitation>
</taxonomicName>
,
<emphasis id="0CDCEA89245EC62EFE4FFD35FAA9FD4D" box="[476,1342,706,727]" italics="true" pageId="2" pageNumber="297">Ƒerhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien</emphasis>
, 41: 333;
</paragraph>
<paragraph id="3E17369B245EC62EFF54FD28FA0BFD6E" blockId="2.[151,1436,671,815]" box="[199,1436,735,756]" pageId="2" pageNumber="297">
<bibRefCitation id="5A394B6A245EC62EFF54FD28FE15FD6E" author="Bey-Bienko, G. Y." box="[199,386,735,756]" pageId="2" pageNumber="297" pagination="472 - 483" refId="ref2202" refString="Bey-Bienko, G. Y. (1971) A revision of the bush-crickets of the genus Xiphidiopsis Redtenbacher (Orthoptera, Tettigonioidea). Entomological Review, 50, 472 - 483." type="journal article" year="1971">Bey-Bienko, 1971</bibRefCitation>
,
<emphasis id="0CDCEA89245EC62EFE07FD28FDE2FD6E" box="[404,629,735,756]" italics="true" pageId="2" pageNumber="297">Entomological Review</emphasis>
, 50: 479;
<bibRefCitation id="5A394B6A245EC62EFD62FD28FC54FD6E" author="Kevan, D. K. M. &amp; Jin, X. B." box="[753,963,735,756]" pageId="2" pageNumber="297" pagination="253 - 274" refId="ref3070" refString="Kevan, D. K. M. &amp; Jin, X. B. (1993) New species of the Xiphidiosis - group from the Indian region (Grylloptera Tettigonioidea Meconematidae). Tropical Zoology, 6 (2), 253 - 274. https: // doi. org / 10.1080 / 03946975.1993.10539226" type="journal article" year="1993">Kevan &amp; Jin, 1993</bibRefCitation>
,
<emphasis id="0CDCEA89245EC62EFC45FD28FB15FD6E" box="[982,1154,735,756]" italics="true" pageId="2" pageNumber="297">Tropical Zoology</emphasis>
, 6: 253;
<bibRefCitation id="5A394B6A245EC62EFB60FD28FA0FFD6E" author="Gorochov, A. V." box="[1267,1432,735,756]" pageId="2" pageNumber="297" pagination="63 - 92" refId="ref2581" refString="Gorochov, A. V. (1993) A contribution to the knowledge of the tribe Meconematini (Orthoptera: Tettigoniidae). Zoosystematica Rossica, 2 (1), 63 - 92." type="journal article" year="1993">Gorochov, 1993</bibRefCitation>
,
</paragraph>
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<emphasis id="0CDCEA89245EC62EFF54FD0AFE3AFC88" box="[199,429,765,786]" italics="true" pageId="2" pageNumber="297">Zoosystematica Rossica</emphasis>
, 2(1): 64;
<bibRefCitation id="5A394B6A245EC62EFD81FD0AFD39FC88" author="Gorochov, A. V." box="[530,686,765,786]" pageId="2" pageNumber="297" pagination="101 - 131" refId="ref2617" refString="Gorochov, A. V. (1998) New and little known Meconematinae of the tribes Meconematini and Phlugidini (Orthoptera, Tettigoniidae). Zoosystematica Rossica, 7 (1), 101 - 131." type="journal article" year="1998">Gorochov, 1998</bibRefCitation>
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, 7(1): 101;
<bibRefCitation id="5A394B6A245EC62EFB9CFD0AFB3DFC88" author="Han, L. &amp; Di, J. X. &amp; Chang, Y. L. &amp; Shi, F. M." box="[1039,1194,765,786]" pageId="2" pageNumber="297" pagination="553 - 562" refId="ref2892" refString="Han, L., Di, J. X., Chang, Y. L. &amp; Shi, F. M. (2015) Three new species of the genus Xiphidiopsis Redtenbacher, 1891 (Orthoptera: Tettigoniidae: Meconematinae) in China. Zootaxa, 4018 (4), 553 - 562. https: // doi. org / 10.11646 / zootaxa. 4018.4.4" type="journal article" year="2015">
Han
<emphasis id="0CDCEA89245EC62EFBAFFD0AFBF0FC88" box="[1084,1127,765,786]" italics="true" pageId="2" pageNumber="297">et al</emphasis>
., 2015
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,
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, 4018(4): 553.
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specie:
<taxonomicName id="F9A84D18245EC62EFE86FCEDFD06FCB5" authority="Redtenbacher, 1891" authorityName="Redtenbacher" authorityYear="1891" box="[277,657,794,815]" class="Insecta" family="Tettigoniidae" genus="Xiphidiopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="0CDCEA89245EC62EFE86FCEDFE5EFCB5" box="[277,457,794,815]" italics="true" pageId="2" pageNumber="297">Xiphidiopsis fallax</emphasis>
<bibRefCitation id="5A394B6A245EC62EFE5CFCEDFD06FCB5" author="Redtenbacher, J." box="[463,657,794,815]" pageId="2" pageNumber="297" pagination="315 - 562" refId="ref3281" refString="Redtenbacher, J. (1891) Monographie der Conocephaliden. Ferhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 41, 315 - 562." type="journal article" year="1891">Redtenbacher, 1891</bibRefCitation>
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<taxonomicName LSID="urn:lsid:zoobank.org:act:80CFF96F-2331-45EB-B4B5-DF5814A57660" class="Diplopoda" family="Paradoxosomatidae" genus="Aponedyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aponedyopus similis" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="similis">Aponedyopus similis</taxonomicName>
<taxonomicNameLabel pageId="8" pageNumber="9">sp. n.</taxonomicNameLabel>
Figs 24-3144, 4554 &amp; 55
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<paragraph pageId="8" pageNumber="9">Material examined:</paragraph>
<paragraph pageId="8" pageNumber="9">Holotype ♂ (TFRI), Taiwan (R. O. C.), Taichung County (台中縣), HePing (和平鄉), Shengguang (勝光), ca 2,200 m a.s.l., 26 March - 25 April, 2003, leg. W. C. Yeh.</paragraph>
<paragraph pageId="8" pageNumber="9">Paratype ♂ (NSYSUB-DI 75), Taiwan (R. O. C.), Hsinchu County (新竹縣), Wufeng Township (五峰鄉), GuanU (觀霧), 24.5 km from entrance to national park, ca 2,000 m a.s.l., 13 August 2002, leg. C. C. Chen, Y. H. Lin &amp; J. N. Huang.</paragraph>
<caption pageId="8" pageNumber="9">
<paragraph pageId="8" pageNumber="9">
Figures 24-31.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Aponedyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aponedyopus similis" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="similis">Aponedyopus similis</taxonomicName>
sp. n., ♂ holotype (24) ♂ paratype (25-31). 24 Entire body, dorsal view 25 Anterior body portion, lateral view 26-27 Midbody segments, dorsal and lateral views, respectively 28-29 Epiproct, dorsal and lateral views, respectively 30 Hypoproct, ventral view 31 Sternal cones near gonopod aperture. Scale bars: 1.0 mm for 24-29, 0.5 mm for 30, 31. cal: calluses; pap: pre-apical papillae.
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<paragraph pageId="8" pageNumber="9">Name:</paragraph>
<paragraph pageId="8" pageNumber="9">To emphasize the close resemblance to the next new species.</paragraph>
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<paragraph pageId="8" pageNumber="9">Diagnosis:</paragraph>
<paragraph pageId="8" pageNumber="9">
Being apparently the most similar to
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sp. n., based both on several peripheral characters (shorter legs, mostly a smaller body size etc.) and gonopod conformation, it is distinguished by the gonopod lobe b being membranous and lobiform, the terminal branches of the solenophore differing in length and crossing each other, with branch x carrying an inconspicuous lobe (see also Key below).
</paragraph>
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<paragraph pageId="8" pageNumber="9">Description:</paragraph>
<paragraph pageId="8" pageNumber="9">Length ca 22 mm (♂, n=2); width of pro- and metazona 10 ca 1.8 and 2.0 mm, respectively.</paragraph>
<paragraph lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
General coloration in alcohol brown to dark brown (Figs 24-27), with a clear pattern of a lighter brown to yellow brown axial stripe consisting of narrower subtriang
<pageBreakToken pageId="9" pageNumber="10" start="start">ular</pageBreakToken>
spots on proterga and twice as wide central spots on metaterga, these spots growing slightly infuscate, to blackish both towards stricture and posterior half of metaterga; prozona slightly darker than metazona, thus providing a vague cingulate pattern as well; paraterga, legs and venter slightly lighter than background, light grey-brown; head marbled brown, especially well so in vertigial region, genae contrastingly yellowish, a square median spot above antennal sockets contrastingly dark brown; antennae increasingly infuscate, up to blackish distad, distinctly darker at margins, marbled and lighter centrally, only tip contrastingly pallid; both collum and segment 2 with a very
<pageBreakToken pageId="10" pageNumber="11" start="start">faint</pageBreakToken>
, yellow-brown, axial line; epiproct uniformly light brown, only very slightly infuscate near base.
</paragraph>
<paragraph pageId="10" pageNumber="11">Postcollum constriction evident; in width, segment 2 = 3 &lt;4 &lt;collum &lt;head = segments 5-15; thereafter body gradually and gently tapering towards telson both in width and height. Antennae (Fig. 25) medium-sized, slender, reaching behind stricture of tergite 3. Paraterga (Figs 26 &amp; 27) very poorly developed, very evident and low only on segment 2, calluses (cal) (nearly) completely delimited by a sulcus dorsally, in caudal 1/3 also ventrally only on pore-bearing segments. Transverse sulcus (Figs 26, 27) developed on segments 5-17, traceable on segment 18, wanting on 19th, narrow, shallow, neither beaded at bottom nor reaching bases of paraterga. Surface smooth throughout, slightly granulated only below paraterga 2-4. Limbus thin, caudal margin entire. Stricture dividing pro- and metazona shallow, narrow, not beaded at bottom (Figs 26 &amp; 27). Pleurosternal carinae present only on segments 2 and 3 (Fig. 25). Tergal setae almost fully abraded, 2+1 retained only at anterior edge of collum; pattern untraceable. Ozopores lateral, lying on calluses ca 1/2 metatergal length in front of caudal edge (Figs 26 &amp; 27). Epiproct long (Figs 28 &amp; 29), flattened dorsoventrally, straight, not curved caudoventrad in lateral view, ratio of epiproct length to pre-epiproct length of telson 1: 1.3, tip of epiproct slightly concave; pre-apical papillae (pap) evident, close to apex. Hypoproct (Fig. 30) rounded, subtrapeziform, 1+1 setae at caudal corners situated on well-separated knobs, sides slightly concave.</paragraph>
<paragraph pageId="10" pageNumber="11">Sterna sparsely setose; lamina between coxae 4 setose and emarginate (Fig. 31); segment 7 with a pair of prominent sternal cones (= spiracles) flanking gonopod aperture; each cross-impression with a transverse sulcus, but without axial groove. Legs (Fig. 27) moderately long and slender, legs 1 to anterior legs of segment 17 with tarsal brushes, thereafter legs broken off in both available ♂♂, each midbody leg ca 1.2 times as long as body height, coxa 2 with a small apical process supporting a gonopore.</paragraph>
<paragraph pageId="10" pageNumber="11">Gonopods (Figs 44, 45, 54 &amp; 55) with process b at base of postfemoral part lobe-shaped, membranous, not like a distinct process; l at base of solenophore rather vague; distal part of gonopod deeply bifid, divided into a longer solenomere (sl), more complex at end and bearing a low terminal lobe, and a slightly shorter, simple, nearly pointed solenophore branch (sph); ends of both branches crossing.</paragraph>
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<paragraph pageId="10" pageNumber="11">Distribution:</paragraph>
<paragraph pageId="10" pageNumber="11">This species seems to be local, occurring only rather high (2,000-2,200 m a.s.l.) in the mountains of northern Taiwan (Map).</paragraph>
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Wheeler.
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<paragraph id="811FA1AECDE67AF95EF6171099960C40" pageNumber="294">[[ queen ]] (non decrite) Long. 6,5 mm. Couleur, pilosite comme chez la. [[ worker ]]. Le pronotum rugueux, non strie. Les stries des mesonotum et scutellum longitudinales et plus fortes que chez la [[ worker ]]. Metanotum en bourrelet transversal, enfonce, lisse, et luisant. Les stries de la face basale de l'epinotum divergent en arriere et en dehors, celles de la face declive sont concentriques a concavite inferieure. Le gastre plus luisant, pour le reste comme la [[ worker ]].</paragraph>
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<taxonomicName id="9850BC58E8D677FBE338EE588B805369" ID-CoL="7X5FZ" ID-ENA="1775724" authority="de Beaumont, 1945" authorityName="de Beaumont" authorityYear="1945" class="Insecta" family="Crabronidae" genus="Trypoxylon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trypoxylon minus" order="Hymenoptera" pageId="0" pageNumber="8050" phylum="Arthropoda" rank="species" species="minus">Trypoxylon minus de Beaumont, 1945</taxonomicName>
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<taxonomicName id="7598D8D468991351C974FCBAFE47D708" genus="Apoidea" lsidName="Apoidea koma" pageId="0" pageNumber="8050" rank="species" species="koma">koma</taxonomicName>
Tsuneki, 1956
</paragraph>
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<paragraph id="B15C7D91A9796A5F60665A0329EA3300" pageId="0" pageNumber="8050">Distribution</paragraph>
<paragraph id="EAE13106F9B8F619D4D843E08E02DDDB" pageId="0" pageNumber="8050">England</paragraph>
</subSubSection>
<subSubSection id="B1A8CB790DB231E1A9C3B15CC6594567" pageId="0" pageNumber="8050" type="notes">
<paragraph id="F676495BAAB169EC7B78EE7F7C931FF7" pageId="0" pageNumber="8050">Notes</paragraph>
<paragraph id="1C3965FFDE9583EC693CB35D2BA50127" pageId="0" pageNumber="8050">
added by
<bibRefCitation id="C7FF98FB9D43A9CB462B58B2FB502DF9" author="Felton, J. C." journalOrPublisher="Entomologist's Monthly Magazine" pageId="0" pageNumber="8050" pagination="221 - 224" title="The genus Trypoxylon Latreille (Hym., Sphecidae) in Kent, with a first record for Trypoxylonminus de Beaumont" volume="124" year="1988">Felton (1988)</bibRefCitation>
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82
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<mods:namePart>Cassar, Thomas</mods:namePart>
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<mods:namePart>Mifsud, David</mods:namePart>
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<paragraph pageId="0" pageNumber="201">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="201">None.</paragraph>
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<paragraph pageId="0" pageNumber="201">Notes.</paragraph>
<paragraph pageId="0" pageNumber="201">
This species was recorded by
<bibRefCitation author="Erlandsson, S" journalOrPublisher="Eos Madrid" pageId="0" pageNumber="201" pagination="11 - 93" refId="B6" refString="Erlandsson, S, 1972. Hymenoptera Aculeata from the European parts of the Mediterranean Countries. Eos Madrid 48: 11 - 93" title="Hymenoptera Aculeata from the European parts of the Mediterranean Countries." volume="48" year="1972">Erlandsson (1972)</bibRefCitation>
and
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. Its distribution range includes Sicily, Malta and Southeast Europe (including Rhodes); it is also found in Cyprus and the Middle East. Individuals of this species may vary somewhat in tergite morphology (
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Eos Madrid" pageId="0" pageNumber="201" refId="B12" refString="Gusenleitner, J, 1998b. Vespoidea und Sapygidae gesammelt auf der Insel Samos (Hymenoptera: Eumenidae, Masaridae, Sapygidae). Linzer Biologische Beitraege 30/1: 155-161." title="Vespoidea und Sapygidae gesammelt auf der Insel Samos (Hymenoptera: Eumenidae, Masaridae, Sapygidae). Linzer Biologische Beitraege 30 / 1: 155 - 161." year="1998 b">Gusenleitner 1998b</bibRefCitation>
).
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<paragraph pageId="10" pageNumber="11">Taxon classification Animalia Hymenoptera Chrysididae</paragraph>
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<taxonomicName authority="du Buysson, 1892" authorityName="du Buysson" authorityYear="1892" class="Insecta" family="Chrysididae" genus="Omalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Omalus biaccinctus" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="biaccinctus">Omalus biaccinctus (du Buysson, 1892)</taxonomicName>
Figs 20, 21
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="reference_group">
<paragraph pageId="10" pageNumber="11">
<taxonomicName class="Insecta" family="Chrysididae" genus="Ellampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ellampus biaccinctus" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="biaccinctus">Ellampus biaccinctus</taxonomicName>
du Buysson (in
<normalizedToken originalValue="André">Andre</normalizedToken>
), 1892: 152.
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<paragraph pageId="10" pageNumber="11">
<taxonomicName class="Insecta" family="Chrysididae" genus="Omalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Omalus biaccinctus" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="biaccinctus">Omalus biaccinctus</taxonomicName>
:
<bibRefCitation author="Trautmann, W" journalOrPublisher="Uschman, Weimar" pageId="99" pageNumber="100" title="Die Goldwespen Europas." year="1927">Trautmann 1927</bibRefCitation>
: 41.
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<paragraph pageId="10" pageNumber="11">Diagnosis.</paragraph>
<paragraph lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Length 3-5 mm. Both sexes differ from
<taxonomicName class="Insecta" family="Chrysididae" genus="Omalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Omalus aeneus" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aeneus">Omalus aeneus</taxonomicName>
and
<taxonomicName class="Insecta" family="Chrysididae" genus="Omalus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Omalus puncticollis" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="puncticollis">Omalus puncticollis</taxonomicName>
by having a bicoloured body (head and mesosoma violet or black, mesosoma reddish or
<pageBreakToken pageId="11" pageNumber="12" start="start">greenish</pageBreakToken>
) and a dull and wrinkled mesoscutum without punctures (Fig. 20). The apical notch of T3 is also more deeply triangular (Fig. 21) than in the other two species. The colour of the mesosoma is violet in the female, but dorsally black or dark violet in the male. The metasoma is greenish, golden or reddish in the female, whereas it is greenish, rarely golden or reddish, and dorsally usually black in the male.
</paragraph>
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<paragraph pageId="11" pageNumber="12">Distribution.</paragraph>
<paragraph pageId="11" pageNumber="12">
Denmark, Estonia, Finland, Latvia, Norway, Sweden. Relatively rare. - West Palearctic: from western Europe to western Asia (
<bibRefCitation author="Linsenmaier, W" journalOrPublisher="Mitteilungen der Schweizerischen Entomologischen Gesellschaft" pageId="95" pageNumber="96" pagination="1 - 232" title="Revision der Familie Chrysididae (Hymenoptera) mit besonderer Bruecksichtigung der europaeischen Spezies." volume="32" year="1959">Linsenmaier 1959</bibRefCitation>
).
</paragraph>
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<paragraph pageId="11" pageNumber="12">Biology.</paragraph>
<paragraph pageId="11" pageNumber="12">
Habitat: pine forest margins and clearings, semi-open sandy areas. Occasionally found on flowers of
<taxonomicName family="Apiaceae" lsidName="Symphyta" pageId="11" pageNumber="12" rank="family">Apiaceae</taxonomicName>
and
<taxonomicName family="Asteraceae" lsidName="Symphyta" pageId="11" pageNumber="12" rank="family">Asteraceae</taxonomicName>
(
<bibRefCitation author="Kofler, A" journalOrPublisher="Carinthia II" pageId="94" pageNumber="95" pagination="343 - 356" title="Die Goldwespen Osttirols (Insecta: Hymenoptera, Chrysididae)." volume="85" year="1975">Kofler 1975</bibRefCitation>
,
<bibRefCitation author="Rosa, P" journalOrPublisher="Giornale Italiano di Entomologia" pageId="97" pageNumber="98" pagination="79 - 90" title="Alcune osservazioni sulle relazioni tra Vegetazione e Crisidi (Hymenoptera, Chrysididae) in Italia." volume="11" year="2004">Rosa 2004</bibRefCitation>
). Flight period: June to August. Host:
<taxonomicName class="Insecta" family="Crabronidae" genus="Passaloecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passaloecus turionum" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="turionum">Passaloecus turionum</taxonomicName>
Dahlbom,
<taxonomicName class="Insecta" family="Crabronidae" genus="Passaloecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passaloecus gracilis" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gracilis">Passaloecus gracilis</taxonomicName>
(Curtis) and
<taxonomicName class="Insecta" family="Crabronidae" genus="Passaloecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passaloecus eremita" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="eremita">Passaloecus eremita</taxonomicName>
Kohl (
<taxonomicName family="Crabronidae" lsidName="" pageId="11" pageNumber="12" rank="family">Crabronidae</taxonomicName>
) (
<bibRefCitation author="Lomholdt, O" journalOrPublisher="Fauna Entomologica Scandinavica" pageId="95" pageNumber="96" pagination="1 - 452" title="The Sphecidae (Hymenoptera) of Fennoscandia and Denmark." volume="4" year="1975 - 1976">Lomholdt 1975</bibRefCitation>
,
<bibRefCitation author="Tormos, J" journalOrPublisher="Florida Entomologist" pageId="99" pageNumber="100" pagination="56 - 63" title="Description of the mature larvae of Chrysisgracillima and Omalusbiaccinctus and new data on the biology of Trichrysiscyanea (HymenopteraChrysididae)." url="10.2307/3495754" volume="79" year="1996">Tormos et al. 1996</bibRefCitation>
,
<bibRefCitation author="Wickl, K-H" journalOrPublisher="Galathea" pageId="100" pageNumber="101" pagination="57 - 72" title="Goldwespen der Oberpfalz (Hymenoptera: Chrysididae)." volume="17" year="2001">Wickl 2001</bibRefCitation>
, our own obs.). Adults have been reared from old resin-galls of
<taxonomicName genus="Retinea" lsidName="Retinea resinella" pageId="11" pageNumber="12" rank="species" species="resinella">Retinea resinella</taxonomicName>
(Linnaeus) (
<taxonomicName family="Tortricidae" lsidName="" pageId="11" pageNumber="12" rank="family">Tortricidae</taxonomicName>
) (V. Vikberg, pers. obs.) and pieces of pine wood (
<bibRefCitation author="Kofler, A" journalOrPublisher="Carinthia II" pageId="94" pageNumber="95" pagination="343 - 356" title="Die Goldwespen Osttirols (Insecta: Hymenoptera, Chrysididae)." volume="85" year="1975">Kofler 1975</bibRefCitation>
) with host nests inside. The females oviposit in living aphids at the hunting site of their host, and the egg is brought into the
<normalizedToken originalValue="host’s">host's</normalizedToken>
nest concealed in the aphid prey (
<bibRefCitation author="Winterhagen, P" journalOrPublisher="European Journal of Entomology" pageId="100" pageNumber="101" pagination="557 - 559" title="Strategy for sneaking into a host's home: The cuckoo wasp Omalusbiaccinctus (Hymenoptera: Chrysididae) inserts its eggs into living aphids that are the prey of its host." url="10.14411/eje.2015.064" volume="112" year="2015">Winterhagen 2015</bibRefCitation>
). Thus, the females do not enter the nest of their host for oviposition.
</paragraph>
<caption pageId="11" pageNumber="12">
<paragraph pageId="11" pageNumber="12">
Figures 19-25. Mesosoma, lateral view (arrow indicating mesopleuron): 19
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♀. Pronotum and mesoscutum, dorsal view: 20
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♀ 22
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♀ 24
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♀. T3, postero-dorsal view: 21
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♀ 23
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♀ 25
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♀. Scale 1 mm.
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142
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<taxonomicName LSID="B60342D2-EE04-5D0C-AC64-5E596AF4A37A" authority="Munster, 1924" authorityName="Munster" authorityYear="1924" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Contacyphon kongsbergensis" order="Coleoptera" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="kongsbergensis">Contacyphon kongsbergensis (Munster, 1924)</taxonomicName>
<figureCitation captionStart="Figures 22–24" captionStartId="F12" captionText="Figures 22 - 24. 22 Contacyphon fuscescens (Klausnitzer) 22 A aedeagus, dorsal plate 22 B aedeagus, ventral plate 22 C male tergite 8 and 9 23 Contacyphon kongsbergensis (Munster) 23 A aedeagus, dorsal plate 23 B aedeagus, ventral plate 23 C male sternite 9 and accessory sclerite 24 Contacyphon obscurellus (Klausnitzer) 24 A aedeagus, dorsal plate 24 B aedeagus, ventral plate 24 C male sternite 9 24 D male tergite 9. Scale bars: 0.2 mm (22 A-C; 23 A-C), 0.5 mm (24 A-D)." figureDoi="10.3897/zookeys.894.37862.figures22-24" httpUri="https://binary.pensoft.net/fig/362148" pageId="0" pageNumber="53" tableDoi="10.3897/zookeys.894.37862.figures22-24">Figure 23</figureCitation>
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<paragraph pageId="0" pageNumber="53">Distribution.</paragraph>
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Holarctic. Recorded from across the Palaearctic region, from Western Europe to the Russian Far East (
<bibRefCitation author="Klausnitzer, B" editor="Loebl, I" journalOrPublisher="Polskie Pismo Entomologiczne" pageId="0" pageNumber="53" refId="B105" refString="Klausnitzer, B, Loebl, I, Smetana, A, 2006. Scirtidae. In: LoeblISmetanaA (Eds) Catalogue of Palaearctic Coleoptera (Vol.3). Apollo Books, Stenstrup, 316-323." title="Scirtidae. In: LoeblISmetanaA (Eds) Catalogue of Palaearctic Coleoptera (Vol. 3). Apollo Books, Stenstrup, 316 - 323." year="2006">Klausnitzer 2006</bibRefCitation>
). In North America, apparently previously known only from Alaska (Kalsin Bay, Kodiak (
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)). This record was overlooked by
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.360.4742" author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="0" pageNumber="53" pagination="1 - 44" refId="B27" refString="Bousquet, Y, Bouchard, P, Davies, A, Sikes, D, 2013. Checklist of beetles (Coleoptera) of Canada and Alaska. Second edition. ZooKeys 360: 1 - 44" title="Checklist of beetles (Coleoptera) of Canada and Alaska. Second edition." url="https://doi.org/10.3897/zookeys.360.4742" volume="360" year="2013">Bousquet et al. (2013)</bibRefCitation>
.
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<paragraph pageId="0" pageNumber="53">Canadian records.</paragraph>
<paragraph pageId="0" pageNumber="53">British Columbia: Smithers, 28-Jul-2014 to 05-Aug-2014 (4 exx, CBG). Alberta: Jasper National Park, 02-Aug-2010 to 05-Aug-2010 (1 ex, CBG). Manitoba: Churchill, 05-Aug-2005 (1 ex, CBG); Churchill, 18-Aug-2006 (1 ex, JBWM); Churchill, 21-Jul-2007 (2 exx, CBG); Riding Mountain National Park, 16-Jul-2012 to 23-Jul-2012 (1 ex, CBG).</paragraph>
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<paragraph pageId="0" pageNumber="53">Diagnostic information</paragraph>
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(based on
<bibRefCitation author="Nyholm, T" journalOrPublisher="Entomologica Scandinavica Supplementum" pageId="0" pageNumber="53" pagination="1 - 100" refId="B154" refString="Nyholm, T, 1972. Die nordeuropaeischen Arten der Gattung Cyphon Paykull (Coleoptera). Entomologica Scandinavica Supplementum 3: 1 - 100" title="Die nordeuropaeischen Arten der Gattung Cyphon Paykull (Coleoptera)." volume="3" year="1972">Nyholm 1972</bibRefCitation>
and
<bibRefCitation author="Lohse, GA" editor="Freude, H" journalOrPublisher="Wiley-Blackwell, Chichester" pageId="0" pageNumber="53" refId="B138" refString="Lohse, GA, Freude, H, Harde, KW, Lohse, GA, 1979. Helodidae. In: FreudeHHardeKWLohseGA (Eds) Die Kaefer Mitteleuropas, Band 6.Goecke &amp; Evers, Krefeld, 249-263." title="Helodidae. In: FreudeHHardeKWLohseGA (Eds) Die Kaefer Mitteleuropas, Band 6. Goecke &amp; Evers, Krefeld, 249 - 263." year="1979">Lohse 1979</bibRefCitation>
). Body length 2.7-3.2 mm. Reddish or yellowish brown, head usually darker than pronotum and elytra. Head and pronotum with dense, coarse, granulate punctation. Elytra densely punctate. Male sternite IX and accessory sclerite as in
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. Aedeagus as in
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.
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This species is known from acidic
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bogs (
<bibRefCitation author="Nyholm, T" journalOrPublisher="Entomologica Scandinavica Supplementum" pageId="0" pageNumber="53" pagination="1 - 100" refId="B154" refString="Nyholm, T, 1972. Die nordeuropaeischen Arten der Gattung Cyphon Paykull (Coleoptera). Entomologica Scandinavica Supplementum 3: 1 - 100" title="Die nordeuropaeischen Arten der Gattung Cyphon Paykull (Coleoptera)." volume="3" year="1972">Nyholm 1972</bibRefCitation>
). Most Canadian specimens were collected with Malaise traps; also collected with light traps and by sweep netting.
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is morphologically most reliably identified by its genitalia. The lack of a modern revision of North American
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prevents detailed comparison with related species.
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<bibRefCitation id="5A3B4B3DD360FF81FE6C380EFD4AFD91" author="Osawa, M. &amp; Okuno, J." box="[434,718,584,610]" pageId="10" pageNumber="473" pagination="29 - 46" refId="ref12922" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12 (1), 29 - 46. https: // doi. org / 10.12782 / specdiv. 12.29" type="journal article" year="2007">Osawa &amp; Okuno, 2007</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="76B06547D360FF81FF49382CFF7BFD70" box="[151,255,618,643]" pageId="10" pageNumber="473" type="description">
<paragraph id="3E1536CCD360FF81FF49382CFF7BFD70" blockId="10.[151,718,583,643]" box="[151,255,618,643]" pageId="10" pageNumber="473">
(
<figureCitation id="A6912A49D360FF81FF41382CFF73FD70" box="[159,247,618,643]" captionStart="FIGURE 3" captionStartId="11.[151,250,1378,1400]" captionTargetBox="[151,1436,195,1356]" captionTargetId="figure@11.[151,1436,195,1356]" captionTargetPageId="11" captionText="FIGURE 3. Entire animal, dorsal view. A, Pylopaguropsis bellula Osawa &amp; Okuno, 2007, male (sl 4.1 mm), RUMF-ZC-6042 (IE17); B, Pylopaguropsis fimbriata McLaughlin &amp; Haig, 1989, ovigerous female (sl 4.4 mm), RUMF-ZC-6046 (IE179); C, Pylopaguropsis granulata Asakura, 2000, female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31); D, Pylopaguropsis keijii McLaughlin &amp; Haig, 1989, male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87); E, Pylopaguropsis speciosa McLaughlin &amp; Haig, 1989, ovigerous female (sl 3.7 mm), RUMF-ZC-6058 (SH104)." httpUri="https://zenodo.org/record/2627824/files/figure.png" pageId="10" pageNumber="473">Fig. 3A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="76B06547D360FF81FF4938F4FAE0FD3B" box="[151,1380,690,712]" pageId="10" pageNumber="473" type="reference_group">
<paragraph id="3E1536CCD360FF81FF4938F4FAE0FD3B" blockId="10.[151,1380,690,712]" box="[151,1380,690,712]" pageId="10" pageNumber="473">
<treatmentCitationGroup id="1EBA11E2D360FF81FF4938F4FAE0FD3B" box="[151,1380,690,712]" pageId="10" pageNumber="473">
<emphasis id="0CDEEADED360FF81FF4938F4FEB1FD3B" box="[151,309,690,712]" italics="true" pageId="10" pageNumber="473">Pylopaguropsis</emphasis>
<taxonomicName id="F9AA4D4FD360FF81FEE538F4FD1DFD3B" authority="Osawa &amp; Okuno, 2007: 31" authorityName="Osawa &amp; Okuno" authorityPageNumber="31" authorityYear="2007" box="[315,665,690,712]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="0CDEEADED360FF81FEE538F4FE05FD3B" box="[315,385,690,712]" italics="true" pageId="10" pageNumber="473">bellula</emphasis>
<treatmentCitation id="BF0B10DDD360FF81FE5638F4FD1DFD3B" author="Osawa, M. &amp; Okuno, J." box="[392,665,690,712]" page="31" pageId="10" pageNumber="473" year="2007">
<bibRefCitation id="5A3B4B3DD360FF81FE5638F4FDF0FD3B" author="Osawa, M. &amp; Okuno, J." box="[392,628,690,712]" pageId="10" pageNumber="473" pagination="29 - 46" refId="ref12922" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12 (1), 29 - 46. https: // doi. org / 10.12782 / specdiv. 12.29" type="journal article" year="2007">Osawa &amp; Okuno, 2007</bibRefCitation>
: 31
</treatmentCitation>
</taxonomicName>
, figs. 1–5, 6A, B, 7A, B (
<typeStatus id="E111886ED360FF81FC4238F4FC4DFD3B" box="[924,969,690,712]" pageId="10" pageNumber="473">type</typeStatus>
locality: Kume Island, Ryukyu Islands).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="76B06547D360FF81FF4938B3FD22FC31" pageId="10" pageNumber="473" type="materials_examined">
<paragraph id="3E1536CCD360FF81FF4938B3FD22FC31" blockId="10.[151,1437,757,1286]" pageId="10" pageNumber="473">
<emphasis id="0CDEEADED360FF81FF4938B3FCCFFCFD" bold="true" box="[151,843,757,782]" pageId="10" pageNumber="473">
Material examined. Ie
<materialsCitation id="8EC23C91D360FF81FE7038B3FCCFFCFD" ID-GBIF-Occurrence="2234219653" box="[430,843,757,782]" collectingDate="2016" collectingDateMax="2067" collectingDateMin="2016" collectionCode="IE" country="Iceland" county="Akuma-no-Yakata cave" location="Okinawa Island" municipality="Shimoji Island" pageId="10" pageNumber="473" specimenCode="IE13, IE17, IE59" specimenCount="6" specimenCount-male="6">
<collectingCountry id="46BD765CD360FF81FE7038B3FE7EFCFD" box="[430,506,757,782]" name="Iceland" pageId="10" pageNumber="473">Island</collectingCountry>
, unnamed submarine cave:
</materialsCitation>
</emphasis>
main tube,
<collectingDate id="5A50E9E4D360FF81FC0738B0FB2AFCFD" box="[985,1198,757,782]" pageId="10" pageNumber="473" value="2016-09-07">
<date id="4A14100CD360FF81FC0738B0FB2AFCFD" box="[985,1198,757,782]" pageId="10" pageNumber="473" value="2016-09-07">7 September 2016</date>
</collectingDate>
,
<specimenCount id="28ACFD45D360FF81FB6238B0FA91FCFD" box="[1212,1301,757,782]" pageId="10" pageNumber="473" type="male">2 males</specimenCount>
(sl 2.9,
<quantity id="F9529B29D360FF81FAA738B0FF43FCC1" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="10" pageNumber="473" unit="mm" value="3.3">3.3 mm</quantity>
), 1 ovigerous female (sl
<quantity id="F9529B29D360FF81FE3C395FFDBAFCC1" box="[482,574,793,818]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="10" pageNumber="473" unit="mm" value="4.6">4.6 mm</quantity>
), RUMF-ZC-6041 (
<specimenCode id="6E0C9EB7D360FF81FCFF395CFCD8FCC1" box="[801,860,794,818]" collectionCode="IE" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="10" pageNumber="473">
<collectionCode id="58BBAE09D360FF81FCFF395CFCC4FCC1" box="[801,832,794,818]" pageId="10" pageNumber="473">IE</collectionCode>
13
</specimenCode>
); main tube, near bifurcation of tube,
<collectingDate id="5A50E9E4D360FF81FAD0395CFF56FCA5" pageId="10" pageNumber="473" value="2016-09-07">
<date id="4A14100CD360FF81FAD0395CFF56FCA5" pageId="10" pageNumber="473" value="2016-09-07">7 September 2016</date>
</collectingDate>
,
<specimenCount id="28ACFD45D360FF81FF033978FEAEFCA5" box="[221,298,829,854]" pageId="10" pageNumber="473" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D360FF81FE8A3978FE2AFCA5" box="[340,430,830,854]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="10" pageNumber="473" unit="mm" value="4.1">4.1 mm</quantity>
), RUMF-ZC-6042 (
<specimenCode id="6E0C9EB7D360FF81FD4E3978FD4FFCA5" box="[656,715,830,854]" collectionCode="IE" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="10" pageNumber="473">
<collectionCode id="58BBAE09D360FF81FD4E3978FD2BFCA5" box="[656,687,830,854]" pageId="10" pageNumber="473">IE</collectionCode>
17
</specimenCode>
); main tube, near bifurcation of tube,
<collectingDate id="5A50E9E4D360FF81FBAB3978FAC1FCA5" box="[1141,1349,829,854]" pageId="10" pageNumber="473" value="2016-09-08">
<date id="4A14100CD360FF81FBAB3978FAC1FCA5" box="[1141,1349,829,854]" pageId="10" pageNumber="473" value="2016-09-08">8 September 2016</date>
</collectingDate>
,
<specimenCount id="28ACFD45D360FF81FA8E3978FA18FCA5" box="[1360,1436,829,854]" pageId="10" pageNumber="473" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D360FF81FF643927FE91FC89" box="[186,277,865,890]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="10" pageNumber="473" unit="mm" value="3.6">3.6 mm</quantity>
), RUMF-ZC-6043 (
<specimenCode id="6E0C9EB7D360FF81FE283927FDB5FC89" box="[502,561,865,890]" collectionCode="IE" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="10" pageNumber="473">
<collectionCode id="58BBAE09D360FF81FE283927FD90FC89" box="[502,532,865,890]" pageId="10" pageNumber="473">IE</collectionCode>
59
</specimenCode>
).
<emphasis id="0CDEEADED360FF81FD9B3927FC61FC89" bold="true" box="[581,997,865,890]" pageId="10" pageNumber="473">
<location id="3B756017D360FF81FD9B3927FC82FC89" LSID="urn:lsid:plazi:treatment:B60387DAD360FF81FF493801FAF8FAF5:3B756017D360FF81FD9B3927FC82FC89" box="[581,774,865,890]" country="Iceland" county="Akuma-no-Yakata cave" municipality="Shimoji Island" name="Okinawa Island" pageId="10" pageNumber="473">Okinawa Island</location>
,
<location id="3B756017D360FF81FCCC3927FC64FC89" LSID="urn:lsid:plazi:treatment:B60387DAD360FF81FF493801FAF8FAF5:3B756017D360FF81FCCC3927FC64FC89" box="[786,992,865,890]" country="Iceland" county="Akuma-no-Yakata cave" municipality="Shimoji Island" name="Hedo Dome cave" pageId="10" pageNumber="473">Hedo Dome cave</location>
:
</emphasis>
main tube,
<collectingDate id="5A50E9E4D360FF81FBB53924FAA5FC89" box="[1131,1313,865,890]" pageId="10" pageNumber="473" value="2016-08-11">
<date id="4A14100CD360FF81FBB53924FAA5FC89" box="[1131,1313,865,890]" pageId="10" pageNumber="473" value="2016-08-11">11 August 2016</date>
</collectingDate>
,
<specimenCount id="28ACFD45D360FF81FAF33924FAFCFC89" box="[1325,1400,865,890]" pageId="10" pageNumber="473" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D360FF81FF4939C3FF76FC6D" box="[151,242,901,926]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="10" pageNumber="473" unit="mm" value="2.6">2.6 mm</quantity>
), RUMF-ZC-6044 (
<collectionCode id="58BBAE09D360FF81FE0D39C3FE7AFC6D" box="[467,510,901,926]" pageId="10" pageNumber="473">HD</collectionCode>
<collectingDate id="5A50E9E4D360FF81FE2039C3FDDBFC6D" box="[510,607,901,926]" pageId="10" pageNumber="473" value="2016" valueMax="2067">
<date id="4A14100CD360FF81FE2039C3FDDBFC6D" box="[510,607,901,926]" pageId="10" pageNumber="473" value="2016" valueMax="2067">2016-67</date>
</collectingDate>
).
<emphasis id="0CDEEADED360FF81FDAC39C3FBD6FC6D" bold="true" box="[626,1106,901,926]" pageId="10" pageNumber="473">
<collectingMunicipality id="DE71ACB6D360FF81FDAC39C3FCA7FC6D" box="[626,803,901,926]" pageId="10" pageNumber="473">Shimoji Island</collectingMunicipality>
,
<collectingCounty id="D7744E40D360FF81FCEE39C3FBC9FC6D" box="[816,1101,901,926]" pageId="10" pageNumber="473">Akuma-no-Yakata cave</collectingCounty>
:
</emphasis>
entrance,
<collectingDate id="5A50E9E4D360FF81FB1639C0FA1CFC6D" box="[1224,1432,901,926]" pageId="10" pageNumber="473" value="2017-09-01">
<date id="4A14100CD360FF81FB1639C0FA1CFC6D" box="[1224,1432,901,926]" pageId="10" pageNumber="473" value="2017-09-01">1 September 2017</date>
</collectingDate>
,
<specimenCount id="28ACFD45D360FF81FF4939ECFF66FC31" box="[151,226,937,962]" pageId="10" pageNumber="473" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D360FF81FED239ECFEE2FC31" box="[268,358,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="473" unit="mm" value="3.8">3.8 mm</quantity>
), RUMF-ZC-6045 (
<collectionCode id="58BBAE09D360FF81FD9839ECFDEAFC31" box="[582,622,938,962]" pageId="10" pageNumber="473">SH</collectionCode>
101).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D360FF81FF19398BFAD9FBDD" pageId="10" pageNumber="473" type="distribution">
<paragraph id="3E1536CCD360FF81FF19398BFC8CFBF9" blockId="10.[151,1437,757,1286]" pageId="10" pageNumber="473">
<emphasis id="0CDEEADED360FF81FF19398BFEE4FC15" bold="true" box="[199,352,973,998]" pageId="10" pageNumber="473">Distribution.</emphasis>
Ryukyu Islands (Ie
<collectingCountry id="46BD765CD360FF81FD97398BFD14FC15" box="[585,656,973,998]" name="Iceland" pageId="10" pageNumber="473">Island</collectingCountry>
, Okinawa Island, Kume Island, and Shimoji Island) in southwestern
<collectingCountry id="46BD765CD360FF81FF4939B4FF5EFBFA" box="[151,218,1010,1033]" name="Japan" pageId="10" pageNumber="473">Japan</collectingCountry>
;
<quantity id="F9529B29D360FF81FF3B39B7FECCFBFA" box="[229,328,1009,1034]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="10" pageNumber="473" unit="m" value="25.0" valueMax="35.0" valueMin="15.0">15–35 m</quantity>
(
<bibRefCitation id="5A3B4B3DD360FF81FE8939B4FD7FFBF9" author="Osawa, M. &amp; Okuno, J." box="[343,763,1009,1034]" pageId="10" pageNumber="473" pagination="29 - 46" refId="ref12922" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12 (1), 29 - 46. https: // doi. org / 10.12782 / specdiv. 12.29" type="journal article" year="2007">Osawa &amp; Okuno 2007; present study</bibRefCitation>
).
</paragraph>
<paragraph id="3E1536CCD360FF81FF193E53FAD9FBDD" blockId="10.[151,1437,757,1286]" box="[199,1373,1045,1070]" pageId="10" pageNumber="473">
<emphasis id="0CDEEADED360FF81FF193E53FEAEFBDD" bold="true" box="[199,298,1045,1070]" pageId="10" pageNumber="473">Habitat.</emphasis>
Coral reefs; found in submarine caves and crevices of fore-reef slopes (
<bibRefCitation id="5A3B4B3DD360FF81FB913E50FAD5FBDD" author="Osawa, M. &amp; Okuno, J." box="[1103,1361,1045,1070]" pageId="10" pageNumber="473" pagination="29 - 46" refId="ref12922" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12 (1), 29 - 46. https: // doi. org / 10.12782 / specdiv. 12.29" type="journal article" year="2007">Osawa &amp; Okuno 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D360FF81FF193E7FFAF8FAF5" pageId="10" pageNumber="473" type="discussion">
<paragraph id="3E1536CCD360FF81FF193E7FFAF8FAF5" blockId="10.[151,1437,757,1286]" pageId="10" pageNumber="473">
<emphasis id="0CDEEADED360FF81FF193E7FFEBFFBA1" bold="true" box="[199,315,1081,1106]" pageId="10" pageNumber="473">Remarks.</emphasis>
Five species of the genus
<emphasis id="0CDEEADED360FF81FDAD3E7FFCA6FBA2" box="[627,802,1081,1105]" italics="true" pageId="10" pageNumber="473">Pylopaguropsis</emphasis>
<bibRefCitation id="5A3B4B3DD360FF81FCF53E7FFC43FBA1" author="Alcock, A." box="[811,967,1081,1106]" pageId="10" pageNumber="473" pagination="1 - 197" refId="ref9291" refString="Alcock, A. (1905) Anomura. Fasc. I. Pagurides. In: Catalogue of the Indian Decapod Crustacea in the Collections of the Indian Museum. Fol. 2. Indian Museum, Calcutta, pp. 1 - 197." type="book chapter" year="1905">Alcock, 1905</bibRefCitation>
,
<taxonomicName id="F9AA4D4FD360FF81FC083E7CFBBAFBA2" authorityName="Osawa &amp; Okuno" authorityYear="2007" box="[982,1086,1081,1105]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="0CDEEADED360FF81FC083E7CFBBAFBA2" box="[982,1086,1081,1105]" italics="true" pageId="10" pageNumber="473">P. bellula</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AA4D4FD360FF81FB8D3E7CFB5DFBA2" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[1107,1241,1081,1105]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="0CDEEADED360FF81FB8D3E7CFB5DFBA2" box="[1107,1241,1081,1105]" italics="true" pageId="10" pageNumber="473">P. fimbriata</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AA4D4FD360FF81FB343E7CFAF2FBA2" authorityName="Asakura" authorityYear="2000" box="[1258,1398,1081,1105]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="granulata">
<emphasis id="0CDEEADED360FF81FB343E7CFAF2FBA2" box="[1258,1398,1081,1105]" italics="true" pageId="10" pageNumber="473">P. granulata</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AA4D4FD360FF81FA593E7CFF4BFB86" authorityName="McLaughlin &amp; Haig" authorityYear="1989" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="keijii">
<emphasis id="0CDEEADED360FF81FA593E7CFF4BFB86" italics="true" pageId="10" pageNumber="473">P. keijii</emphasis>
</taxonomicName>
, and
<taxonomicName id="F9AA4D4FD360FF81FECD3E18FE16FB86" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[275,402,1118,1141]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="0CDEEADED360FF81FECD3E18FE16FB86" box="[275,402,1118,1141]" italics="true" pageId="10" pageNumber="473">P. speciosa</emphasis>
</taxonomicName>
, have been recorded from the submarine caves in the Ryukyu Islands (
<bibRefCitation id="5A3B4B3DD360FF81FB0D3E18FF57FB69" author="Osawa, M. &amp; Takeda, M." pageId="10" pageNumber="473" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD360FF81FF3C3EC4FE6BFB69" author="Osawa, M. &amp; Okuno, J." box="[226,495,1153,1178]" pageId="10" pageNumber="473" pagination="29 - 46" refId="ref12922" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12 (1), 29 - 46. https: // doi. org / 10.12782 / specdiv. 12.29" type="journal article" year="2007">Osawa &amp; Okuno 2007</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD360FF81FE213EC4FC84FB69" author="Osawa, M. &amp; Fujita, Y." box="[511,768,1153,1178]" pageId="10" pageNumber="473" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita 2016</bibRefCitation>
; present study;
<tableCitation id="73280377D360FF81FC633EC7FB9CFB69" box="[957,1048,1153,1178]" captionStart="TABLE 2" captionStartId="6.[147,233,596,618]" captionText="TABLE 2. Submarine caves surveyed and hermit crab species collected in three islands of the Ryukyu Islands, southwestern Japan." pageId="10" pageNumber="473">Table 2</tableCitation>
). The dactyli and propodi of the ambulatory legs are solid-colored in
<taxonomicName id="F9AA4D4FD360FF81FD9C3EE0FD2EFB4E" authorityName="Osawa &amp; Okuno" authorityYear="2007" box="[578,682,1189,1213]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="0CDEEADED360FF81FD9C3EE0FD2EFB4E" box="[578,682,1189,1213]" italics="true" pageId="10" pageNumber="473">P. bellula</emphasis>
</taxonomicName>
and
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<emphasis id="0CDEEADED360FF81FD343EE0FCC5FB4E" box="[746,833,1189,1213]" italics="true" pageId="10" pageNumber="473">P. keijii</emphasis>
</taxonomicName>
, whereas those of
<taxonomicName id="F9AA4D4FD360FF81FBC13EE0FB21FB4E" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[1055,1189,1189,1213]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="0CDEEADED360FF81FBC13EE0FB21FB4E" box="[1055,1189,1189,1213]" italics="true" pageId="10" pageNumber="473">P. fimbriata</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AA4D4FD360FF81FB693EE0FAC7FB4E" authorityName="Asakura" authorityYear="2000" box="[1207,1347,1189,1213]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="granulata">
<emphasis id="0CDEEADED360FF81FB693EE0FAC7FB4E" box="[1207,1347,1189,1213]" italics="true" pageId="10" pageNumber="473">P. granulata</emphasis>
</taxonomicName>
, and
<taxonomicName id="F9AA4D4FD360FF81FA593EE0FF7CFB12" authorityName="McLaughlin &amp; Haig" authorityYear="1989" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="0CDEEADED360FF81FA593EE0FF7CFB12" italics="true" pageId="10" pageNumber="473">P. speciosa</emphasis>
</taxonomicName>
have white stripes (
<figureCitation id="A6912A49D360FF81FE3B3E8FFDA8FB11" box="[485,556,1225,1250]" captionStart="FIGURE 3" captionStartId="11.[151,250,1378,1400]" captionTargetBox="[151,1436,195,1356]" captionTargetId="figure@11.[151,1436,195,1356]" captionTargetPageId="11" captionText="FIGURE 3. Entire animal, dorsal view. A, Pylopaguropsis bellula Osawa &amp; Okuno, 2007, male (sl 4.1 mm), RUMF-ZC-6042 (IE17); B, Pylopaguropsis fimbriata McLaughlin &amp; Haig, 1989, ovigerous female (sl 4.4 mm), RUMF-ZC-6046 (IE179); C, Pylopaguropsis granulata Asakura, 2000, female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31); D, Pylopaguropsis keijii McLaughlin &amp; Haig, 1989, male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87); E, Pylopaguropsis speciosa McLaughlin &amp; Haig, 1989, ovigerous female (sl 3.7 mm), RUMF-ZC-6058 (SH104)." httpUri="https://zenodo.org/record/2627824/files/figure.png" pageId="10" pageNumber="473">Fig. 3</figureCitation>
). Although such a solid coloration of the ambulatory legs is also known in another congener
<taxonomicName id="F9AA4D4FD360FF81FE813EA8FDF2FAF5" authority="Asakura, 2000" authorityName="Asakura" authorityYear="2000" box="[351,630,1261,1286]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="furusei">
<emphasis id="0CDEEADED360FF81FE813EA8FE4DFAF6" box="[351,457,1261,1285]" italics="true" pageId="10" pageNumber="473">P. furusei</emphasis>
<bibRefCitation id="5A3B4B3DD360FF81FE0E3EABFDF2FAF5" author="Asakura, A." box="[464,630,1261,1286]" pageId="10" pageNumber="473" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura, 2000</bibRefCitation>
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, it is uncommon and usually striped in other
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species.
</paragraph>
</subSubSection>
</treatment>
</document>

234
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<emphasis id="0CDEEADED360FF81FE863F1FFE4CFA80" bold="true" box="[344,456,1369,1395]" italics="true" pageId="10" pageNumber="473">fimbriata</emphasis>
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(
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)
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, figs. 1f, l, 2f, 4d, 6d, 8e, 10d, 12d,
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(
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locality:
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Island,
<collectingCountry id="46BD765CD360FF81FF193FA4FEEEFA0B" box="[199,362,1506,1528]" name="Northern Mariana Islands" pageId="10" pageNumber="473">Mariana Islands</collectingCountry>
).—
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: 85
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, figs. 1C–E, 8, 9, 19D, 20.
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<emphasis id="0CDEEADED360FF81FF493C63FCCEF9CD" bold="true" box="[151,842,1573,1598]" pageId="10" pageNumber="473">
Material examined. Ie
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entrance,
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, 1 ovigerous female (sl
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), RUMF-ZC-6046 (
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<collectionCode id="58BBAE09D360FF81FE6A3C0CFE56F991" box="[436,466,1610,1634]" pageId="10" pageNumber="473">IE</collectionCode>
179
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); main tube,
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<date id="4A14100CD360FF81FD7D3C0CFCCCF991" box="[675,840,1610,1634]" pageId="10" pageNumber="473" value="2017-06-24">24 June 2017</date>
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,
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(sl
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), RUMF-ZC-6047 (
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<collectionCode id="58BBAE09D360FF81FA983C0CFAE0F991" box="[1350,1380,1610,1634]" pageId="10" pageNumber="473">IE</collectionCode>
183
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,
<location id="3B756017D360FF81FEBA3C2BFDB6F975" LSID="urn:lsid:plazi:treatment:B60387DAD360FF81FF493F1FFAF3F839:3B756017D360FF81FEBA3C2BFDB6F975" box="[356,562,1645,1670]" country="Iceland" county="Akuma-no-Yakata cave" municipality="Shimoji Island" name="Hedo Dome cave" pageId="10" pageNumber="473">Hedo Dome cave</location>
:
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main tube,
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<date id="4A14100CD360FF81FD633C28FCF7F975" box="[701,883,1645,1670]" pageId="10" pageNumber="473" value="2016-08-11">11 August 2016</date>
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,
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(sl 4.0 mm), RUMF-ZC-6048 (
<collectionCode id="58BBAE09D360FF81FAEF3C2BFADEF975" box="[1329,1370,1645,1670]" pageId="10" pageNumber="473">HD</collectionCode>
<collectingDate id="5A50E9E4D360FF81FA843C2BFF33F959" pageId="10" pageNumber="473" value="2016" valueMax="2052">
<date id="4A14100CD360FF81FA843C2BFF33F959" pageId="10" pageNumber="473" value="2016" valueMax="2052">2016- 52</date>
</collectingDate>
); same data as RUMF-ZC-6048,
<specimenCount id="28ACFD45D360FF81FDF03CD4FDFEF959" box="[558,634,1681,1706]" pageId="10" pageNumber="473" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D360FF81FD7B3CD4FC84F959" box="[677,768,1682,1706]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="10" pageNumber="473" unit="mm" value="4.1">4.1 mm</quantity>
), RUMF-ZC-6049 (
<collectionCode id="58BBAE09D360FF81FC3F3CD7FB88F959" box="[993,1036,1681,1706]" pageId="10" pageNumber="473">HD</collectionCode>
<collectingDate id="5A50E9E4D360FF81FBD23CD7FBE8F959" box="[1036,1132,1681,1706]" pageId="10" pageNumber="473" value="2016" valueMax="2066">
<date id="4A14100CD360FF81FBD23CD7FBE8F959" box="[1036,1132,1681,1706]" pageId="10" pageNumber="473" value="2016" valueMax="2066">2016-66</date>
</collectingDate>
); main tube,
<collectingDate id="5A50E9E4D360FF81FB213CD4FA1CF959" box="[1279,1432,1682,1706]" pageId="10" pageNumber="473" value="2017-05-21">
<date id="4A14100CD360FF81FB213CD4FA1CF959" box="[1279,1432,1682,1706]" pageId="10" pageNumber="473" value="2017-05-21">21 May 2017</date>
</collectingDate>
,
<specimenCount id="28ACFD45D360FF81FF493CF0FF66F93D" box="[151,226,1717,1742]" pageId="10" pageNumber="473" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D360FF81FED33CF0FEE3F93D" box="[269,359,1718,1742]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="10" pageNumber="473" unit="mm" value="4.4">4.4 mm</quantity>
), RUMF-ZC-6050 (
<collectionCode id="58BBAE09D360FF81FD993CF0FDF6F93D" box="[583,626,1718,1742]" pageId="10" pageNumber="473">HD</collectionCode>
<collectingDate id="5A50E9E4D360FF81FDAF3CF0FD54F93D" box="[625,720,1718,1742]" pageId="10" pageNumber="473" value="2017" valueMax="2084">
<date id="4A14100CD360FF81FDAF3CF0FD54F93D" box="[625,720,1718,1742]" pageId="10" pageNumber="473" value="2017" valueMax="2084">2017-84</date>
</collectingDate>
, addition).
<emphasis id="0CDEEADED360FF81FC913CF3FAAAF93D" bold="true" box="[847,1326,1717,1742]" pageId="10" pageNumber="473">
<collectingMunicipality id="DE71ACB6D360FF81FC913CF3FB84F93D" box="[847,1024,1717,1742]" pageId="10" pageNumber="473">Shimoji Island</collectingMunicipality>
,
<collectingCounty id="D7744E40D360FF81FBD33CF3FAAEF93D" box="[1037,1322,1717,1742]" pageId="10" pageNumber="473">Akuma-no-Yakata cave</collectingCounty>
:
</emphasis>
entrance,
<collectingDate id="5A50E9E4D360FF81FF493C9CFEE2F901" box="[151,358,1753,1778]" pageId="10" pageNumber="473" value="2017-09-01">
<date id="4A14100CD360FF81FF493C9CFEE2F901" box="[151,358,1753,1778]" pageId="10" pageNumber="473" value="2017-09-01">1 September 2017</date>
</collectingDate>
,
<specimenCount id="28ACFD45D360FF81FEAF3C9CFE38F901" box="[369,444,1753,1778]" pageId="10" pageNumber="473" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D360FF81FE383C9CFDC4F901" box="[486,576,1754,1778]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="10" pageNumber="473" unit="mm" value="4.8">4.8 mm</quantity>
), RUMF-ZC-6051 (
<collectionCode id="58BBAE09D360FF81FCFE3C9CFCC3F901" box="[800,839,1754,1778]" pageId="10" pageNumber="473">SH</collectionCode>
103).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D360FF81FF193CBBFE34F871" pageId="10" pageNumber="473" type="distribution">
<paragraph id="3E1536CCD360FF81FF193CBBFD64F8C9" blockId="10.[151,1436,1573,1994]" pageId="10" pageNumber="473">
<emphasis id="0CDEEADED360FF81FF193CBBFEE4F8E5" bold="true" box="[199,352,1789,1814]" pageId="10" pageNumber="473">Distribution.</emphasis>
<collectingCountry id="46BD765CD360FF81FEB43CBBFE57F8E5" box="[362,467,1789,1814]" name="Malaysia" pageId="10" pageNumber="473">Malaysia</collectingCountry>
,
<collectingCountry id="46BD765CD360FF81FE3F3CBBFDD5F8E5" box="[481,593,1789,1814]" name="Indonesia" pageId="10" pageNumber="473">Indonesia</collectingCountry>
,
<collectingCountry id="46BD765CD360FF81FD803CB8FD21F8E5" box="[606,677,1790,1814]" name="Guam" pageId="10" pageNumber="473">Guam</collectingCountry>
, and
<collectingCountry id="46BD765CD360FF81FD3B3CB8FCA1F8E6" box="[741,805,1790,1813]" name="Japan" pageId="10" pageNumber="473">Japan</collectingCountry>
(Ryukyu and Izu Islands); subtidal to
<quantity id="F9529B29D360FF81FB3D3CBBFA9AF8E6" box="[1251,1310,1789,1814]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="473" unit="m" value="35.0">35 m</quantity>
(
<bibRefCitation id="5A3B4B3DD360FF81FAF13CB8FEAEF8C9" author="Osawa, M. &amp; Takeda, M." pageId="10" pageNumber="473" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD360FF81FEEB3D67FDABF8C9" author="Okuno, J. &amp; Arima, H." box="[309,559,1825,1850]" pageId="10" pageNumber="473" pagination="29 - 43" refId="ref12530" refString="Okuno, J. &amp; Arima, H. (2006) Additional records of inshore hermit crabs (Crustacea: Decapoda: Anomura) from Izu-ohshima Island, the northern Izu Islands, Japan. Bulletin of the Biogeographical Society of Japan, 61, 29 - 43. [in Japanese with English abstract]" type="journal article" year="2006">Okuno &amp; Arima 2006</bibRefCitation>
; present study).
</paragraph>
<paragraph id="3E1536CCD360FF81FF193D03FE34F871" blockId="10.[151,1436,1573,1994]" pageId="10" pageNumber="473">
<emphasis id="0CDEEADED360FF81FF193D03FEAEF8AD" bold="true" box="[199,298,1861,1886]" pageId="10" pageNumber="473">Habitat.</emphasis>
Coral reefs; often found in small crevices and horizontal holes, or submarine caves in fore-reef slopes (
<bibRefCitation id="5A3B4B3DD360FF81FF403D2CFE20F871" author="Osawa, M. &amp; Takeda, M." box="[158,420,1897,1922]" pageId="10" pageNumber="473" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D360FF81FF193DCBFAF3F839" pageId="10" pageNumber="473" type="discussion">
<paragraph id="3E1536CCD360FF81FF193DCBFAF3F839" blockId="10.[151,1436,1573,1994]" pageId="10" pageNumber="473">
<emphasis id="0CDEEADED360FF81FF193DCBFEBFF855" bold="true" box="[199,315,1933,1958]" pageId="10" pageNumber="473">Remarks.</emphasis>
<emphasis id="0CDEEADED360FF81FE9F3DCBFDDBF856" box="[321,607,1933,1957]" italics="true" pageId="10" pageNumber="473">
Pylopaguropsis
<taxonomicName id="F9AA4D4FD360FF81FE293DCBFDDBF856" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[503,607,1933,1957]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="fimbriata">fimbriata</taxonomicName>
</emphasis>
is the most commonly found species of the genus in coral reefs, including submarine caves in the Ryukyu Islands (
<bibRefCitation id="5A3B4B3DD360FF81FD843DF7FD7FF839" author="Asakura, A." box="[602,763,1969,1994]" pageId="10" pageNumber="473" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD360FF81FCD93DF4FB8EF839" author="Osawa, M. &amp; Takeda, M." box="[775,1034,1969,1994]" pageId="10" pageNumber="473" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD360FF81FBCB3DF4FA8DF839" author="Osawa, M. &amp; Fujita, Y." box="[1045,1289,1969,1994]" pageId="10" pageNumber="473" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita 2016</bibRefCitation>
;
<tableCitation id="73280377D360FF81FACA3DF7FAEDF839" box="[1300,1385,1969,1994]" captionStart="TABLE 2" captionStartId="6.[147,233,596,618]" captionText="TABLE 2. Submarine caves surveyed and hermit crab species collected in three islands of the Ryukyu Islands, southwestern Japan." pageId="10" pageNumber="473">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

168
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<paragraph id="3E1536CCD361FF80FF493C78FD0FF9AB" blockId="11.[151,651,1598,1657]" box="[151,651,1598,1624]" pageId="11" pageNumber="474">
<heading id="655D81A0D361FF80FF493C78FD0FF9AB" bold="true" box="[151,651,1598,1624]" fontSize="11" level="1" pageId="11" pageNumber="474" reason="1">
<emphasis id="0CDEEADED361FF80FF493C78FD0FF9AB" bold="true" box="[151,651,1598,1624]" pageId="11" pageNumber="474">
<emphasis id="0CDEEADED361FF80FF493C78FED5F9AB" bold="true" box="[151,337,1598,1624]" italics="true" pageId="11" pageNumber="474">Pylopaguropsis</emphasis>
<taxonomicName id="F9AA4D4FD361FF80FE863C78FD0FF9AB" authority="Asakura, 2000" authorityName="Asakura" authorityYear="2000" box="[344,651,1598,1624]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="474" phylum="Arthropoda" rank="species" species="granulata">
<emphasis id="0CDEEADED361FF80FE863C78FE4AF9AB" bold="true" box="[344,462,1598,1624]" italics="true" pageId="11" pageNumber="474">granulata</emphasis>
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<subSubSection id="76B06547D361FF80FF493C26FED2F98A" box="[151,342,1632,1657]" pageId="11" pageNumber="474" type="description">
<paragraph id="3E1536CCD361FF80FF493C26FED2F98A" blockId="11.[151,651,1598,1657]" box="[151,342,1632,1657]" pageId="11" pageNumber="474">
(
<figureCitation id="A6912A49D361FF80FF413C26FE85F98A" box="[159,257,1632,1657]" captionStart="FIGURE 3" captionStartId="11.[151,250,1378,1400]" captionTargetBox="[151,1436,195,1356]" captionTargetId="figure@11.[151,1436,195,1356]" captionTargetPageId="11" captionText="FIGURE 3. Entire animal, dorsal view. A, Pylopaguropsis bellula Osawa &amp; Okuno, 2007, male (sl 4.1 mm), RUMF-ZC-6042 (IE17); B, Pylopaguropsis fimbriata McLaughlin &amp; Haig, 1989, ovigerous female (sl 4.4 mm), RUMF-ZC-6046 (IE179); C, Pylopaguropsis granulata Asakura, 2000, female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31); D, Pylopaguropsis keijii McLaughlin &amp; Haig, 1989, male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87); E, Pylopaguropsis speciosa McLaughlin &amp; Haig, 1989, ovigerous female (sl 3.7 mm), RUMF-ZC-6058 (SH104)." httpUri="https://zenodo.org/record/2627824/files/figure.png" pageId="11" pageNumber="474">Figs. 3C</figureCitation>
,
<figureCitation id="A6912A49D361FF80FED33C26FE9DF98A" box="[269,281,1632,1657]" captionStart="FIGURE 5" captionStartId="13.[151,250,1477,1499]" captionTargetBox="[254,1336,832,1406]" captionTargetId="figure@13.[151,1436,807,1455]" captionTargetPageId="13" captionText="FIGURE 5. Pagurixus nomurai Komai &amp; Asakura, 1995: female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65) (A, B); Pylopaguropsis granulata Asakura, 2000: female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31) (C–E); Pylopaguropsis keijii McLaughlin &amp; Haig, 1989: male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87) (F). A, carpus of left cheliped, dorsal view; B, dactylus and propodus of left third pereopod, lateral view; C, right chela, dorsal view; D, carpus and merus of left second pereopod, lateral view; E, carpus and merus of left third pereopod, lateral view; F, propodus of right third pereopod, lateral view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627828/files/figure.png" pageId="11" pageNumber="474">5</figureCitation>
C–E)
</paragraph>
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<subSubSection id="76B06547D361FF80FF493CEFFA9CF94C" box="[151,1304,1705,1727]" pageId="11" pageNumber="474" type="reference_group">
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<treatmentCitationGroup id="1EBA11E2D361FF80FF493CEFFA9CF94C" box="[151,1304,1705,1727]" pageId="11" pageNumber="474">
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<emphasis id="0CDEEADED361FF80FF493CEFFEB1F94C" box="[151,309,1705,1727]" italics="true" pageId="11" pageNumber="474">Pylopaguropsis</emphasis>
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<emphasis id="0CDEEADED361FF80FEE53CEFFE1BF94C" box="[315,415,1705,1727]" italics="true" pageId="11" pageNumber="474">granulata</emphasis>
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<bibRefCitation id="5A3B4B3DD361FF80FE7B3CEFFDB9F94C" author="Asakura, A." box="[421,573,1705,1727]" pageId="11" pageNumber="474" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura, 2000</bibRefCitation>
: 100
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, figs. 16–18, 19F, 20 (type locality: Kume Island, Ryukyu Islands)
</materialsCitation>
.
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</paragraph>
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<subSubSection id="76B06547D361FF80FF493CADFE27F8DB" pageId="11" pageNumber="474" type="materials_examined">
<paragraph id="3E1536CCD361FF80FF493CADFE27F8DB" blockId="11.[151,1437,1771,2012]" pageId="11" pageNumber="474">
<materialsCitation id="8EC23C91D361FF80FF493CADFE1BF8DB" ID-GBIF-Occurrence="2465041899" collectingDate="2017-05-21" collectionCode="HD" country="Japan" location="silty room" pageId="11" pageNumber="474" specimenCode="RUMF- ZC-6052" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa Island, Hedo Dome cave">
<emphasis id="0CDEEADED361FF80FF493CADFCA0F8F7" bold="true" box="[151,804,1771,1796]" pageId="11" pageNumber="474">
Material examined.
<collectingRegion id="FC6EF82ED361FF80FE583CADFC9BF8F7" box="[390,799,1771,1796]" pageId="11" pageNumber="474">Okinawa Island, Hedo Dome cave</collectingRegion>
:
</emphasis>
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,
<date id="4A14100CD361FF80FC743CAAFBC6F8F7" box="[938,1090,1772,1796]" pageId="11" pageNumber="474" value="2017-05-21">
<collectingDate id="5A50E9E4D361FF80FC743CAAFBC6F8F7" box="[938,1090,1772,1796]" pageId="11" pageNumber="474" value="2017-05-21">21 May 2017</collectingDate>
</date>
,
<specimenCount id="28ACFD45D361FF80FB933CAAFB29F8F7" box="[1101,1197,1771,1796]" pageId="11" pageNumber="474" type="female">1 female</specimenCount>
(sl
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),
<specimenCode id="6E0C9EB7D361FF80FA9D3CAAFF78F8DB" pageId="11" pageNumber="474">RUMF- ZC-6052</specimenCode>
(HD
<date id="4A14100CD361FF80FEE93D56FE13F8DB" box="[311,407,1808,1832]" pageId="11" pageNumber="474" value="2017" valueMax="2031">2017-31</date>
)
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.
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<subSubSection id="76B06547D361FF80FF193D75FDEDF867" pageId="11" pageNumber="474" type="distribution">
<paragraph id="3E1536CCD361FF80FF193D75FE07F883" blockId="11.[151,1437,1771,2012]" pageId="11" pageNumber="474">
<emphasis id="0CDEEADED361FF80FF193D75FEE4F8BF" bold="true" box="[199,352,1843,1868]" pageId="11" pageNumber="474">Distribution.</emphasis>
Ryukyu Islands (Okinawa Island and Kume Island) in southwestern Japan;
<quantity id="F9529B29D361FF80FB163D75FAA9F8B8" box="[1224,1325,1843,1868]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="11" pageNumber="474" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10–15 m</quantity>
(
<bibRefCitation id="5A3B4B3DD361FF80FAE33D75FEF2F883" author="Asakura, A." pageId="11" pageNumber="474" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000; present study</bibRefCitation>
).
</paragraph>
<paragraph id="3E1536CCD361FF80FF193D3DFDEDF867" blockId="11.[151,1437,1771,2012]" box="[199,617,1915,1940]" pageId="11" pageNumber="474">
<emphasis id="0CDEEADED361FF80FF193D3DFEAEF867" bold="true" box="[199,298,1915,1940]" pageId="11" pageNumber="474">Habitat.</emphasis>
Coral reefs (
<bibRefCitation id="5A3B4B3DD361FF80FE653D3DFDDAF867" author="Asakura, A." box="[443,606,1915,1940]" pageId="11" pageNumber="474" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D361FF87FF193DD9FA12FE5E" lastPageId="12" lastPageNumber="475" pageId="11" pageNumber="474" type="discussion">
<paragraph id="3E1536CCD361FF80FF193DD9FA18F82F" blockId="11.[151,1437,1771,2012]" pageId="11" pageNumber="474">
<emphasis id="0CDEEADED361FF80FF193DD9FEBFF84B" bold="true" box="[199,315,1951,1976]" pageId="11" pageNumber="474">Remarks.</emphasis>
<emphasis id="0CDEEADED361FF80FE983DD9FDF4F844" box="[326,624,1951,1975]" italics="true" pageId="11" pageNumber="474">
Pylopaguropsis
<taxonomicName id="F9AA4D4FD361FF80FDDF3DD9FDF4F844" authorityName="Asakura" authorityYear="2000" box="[513,624,1951,1975]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="474" phylum="Arthropoda" rank="species" species="granulata">granulata</taxonomicName>
</emphasis>
was described by
<bibRefCitation id="5A3B4B3DD361FF80FC8C3DD9FB81F84B" author="Asakura, A." box="[850,1029,1951,1976]" pageId="11" pageNumber="474" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura (2000)</bibRefCitation>
based only on the male
<typeStatus id="E111886ED361FF80FAED3DD9FA1DF84B" box="[1331,1433,1951,1976]" pageId="11" pageNumber="474" type="holotype">holotype</typeStatus>
. McLaughlin
<emphasis id="0CDEEADED361FF80FEF23D82FEE4F82F" box="[300,352,1988,2012]" italics="true" pageId="11" pageNumber="474">et al</emphasis>
. (2010) showed a color photograph of the second specimen of the species from the Ryukyu
</paragraph>
<paragraph id="3E1536CCD366FF87FF493AD1FF6BFF26" blockId="12.[151,1437,151,429]" pageId="12" pageNumber="475">Islands. The specimen herein reported represents the third record and a first female collected of this poorly known species.</paragraph>
<paragraph id="3E1536CCD366FF87FF193A99FA12FE5E" blockId="12.[151,1437,151,429]" pageId="12" pageNumber="475">
There are some differences between the
<typeStatus id="E111886ED366FF87FD573A99FD6FFF0B" box="[649,747,223,248]" pageId="12" pageNumber="475" type="holotype">holotype</typeStatus>
and the specimen examined. The right chela is more elongate in the
<typeStatus id="E111886ED366FF87FF013B42FEC5FEEE" box="[223,321,260,285]" pageId="12" pageNumber="475" type="holotype">holotype</typeStatus>
than the present specimen; measured on the lateral margin, 1.5 versus 1.4 times longer than broad (cf.
<bibRefCitation id="5A3B4B3DD366FF87FF1A3B61FEE1FEB3" author="Asakura, A." box="[196,357,295,320]" pageId="12" pageNumber="475" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
: fig. 17A;
<figureCitation id="A6912A49D366FF87FE033B61FDB0FEB3" box="[477,564,295,320]" captionStart="FIGURE 5" captionStartId="13.[151,250,1477,1499]" captionTargetBox="[254,1336,832,1406]" captionTargetId="figure@13.[151,1436,807,1455]" captionTargetPageId="13" captionText="FIGURE 5. Pagurixus nomurai Komai &amp; Asakura, 1995: female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65) (A, B); Pylopaguropsis granulata Asakura, 2000: female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31) (C–E); Pylopaguropsis keijii McLaughlin &amp; Haig, 1989: male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87) (F). A, carpus of left cheliped, dorsal view; B, dactylus and propodus of left third pereopod, lateral view; C, right chela, dorsal view; D, carpus and merus of left second pereopod, lateral view; E, carpus and merus of left third pereopod, lateral view; F, propodus of right third pereopod, lateral view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627828/files/figure.png" pageId="12" pageNumber="475">Fig. 5C</figureCitation>
). The ventrolateral margin of the merus of each ambulatory leg is unarmed in the
<typeStatus id="E111886ED366FF87FF1F3B0AFEA2FE96" box="[193,294,332,357]" pageId="12" pageNumber="475" type="holotype">holotype</typeStatus>
, whereas it has two (second pereopod) and one (third pereopod) small spines on the distal half in the present specimen (
<figureCitation id="A6912A49D366FF87FEB73B29FE60FE7B" box="[361,484,367,392]" captionStart="FIGURE 5" captionStartId="13.[151,250,1477,1499]" captionTargetBox="[254,1336,832,1406]" captionTargetId="figure@13.[151,1436,807,1455]" captionTargetPageId="13" captionText="FIGURE 5. Pagurixus nomurai Komai &amp; Asakura, 1995: female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65) (A, B); Pylopaguropsis granulata Asakura, 2000: female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31) (C–E); Pylopaguropsis keijii McLaughlin &amp; Haig, 1989: male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87) (F). A, carpus of left cheliped, dorsal view; B, dactylus and propodus of left third pereopod, lateral view; C, right chela, dorsal view; D, carpus and merus of left second pereopod, lateral view; E, carpus and merus of left third pereopod, lateral view; F, propodus of right third pereopod, lateral view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627828/files/figure.png" pageId="12" pageNumber="475">Fig. 5D, E</figureCitation>
). The dorsal margin of the carpus of the third pereopod is also unarmed except for a distal spine in the
<typeStatus id="E111886ED366FF87FEBF3BD2FE42FE5E" box="[353,454,404,429]" pageId="12" pageNumber="475" type="holotype">holotype</typeStatus>
, but it has two very small spines on the proximal half in the present specimen (
<figureCitation id="A6912A49D366FF87FAEC3BD2FA0DFE5E" box="[1330,1417,404,429]" captionStart="FIGURE 5" captionStartId="13.[151,250,1477,1499]" captionTargetBox="[254,1336,832,1406]" captionTargetId="figure@13.[151,1436,807,1455]" captionTargetPageId="13" captionText="FIGURE 5. Pagurixus nomurai Komai &amp; Asakura, 1995: female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65) (A, B); Pylopaguropsis granulata Asakura, 2000: female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31) (C–E); Pylopaguropsis keijii McLaughlin &amp; Haig, 1989: male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87) (F). A, carpus of left cheliped, dorsal view; B, dactylus and propodus of left third pereopod, lateral view; C, right chela, dorsal view; D, carpus and merus of left second pereopod, lateral view; E, carpus and merus of left third pereopod, lateral view; F, propodus of right third pereopod, lateral view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627828/files/figure.png" pageId="12" pageNumber="475">Fig. 5E</figureCitation>
).
</paragraph>
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<emphasis id="0CDEEADED363FF82FF493952FE90FCDD" bold="true" box="[151,276,788,814]" italics="true" pageId="9" pageNumber="472">Pagurixus</emphasis>
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<paragraph id="3E1536CCD363FF82FF493970FEF8FCBC" blockId="9.[151,700,788,847]" box="[151,380,822,847]" pageId="9" pageNumber="472">
(
<figureCitation id="A6912A49D363FF82FF413970FEA0FCBD" box="[159,292,822,847]" captionStart="FIGURE 2" captionStartId="8.[151,250,1799,1821]" captionTargetBox="[151,1436,519,1777]" captionTargetId="figure@8.[151,1436,519,1777]" captionTargetPageId="8" captionText="FIGURE 2. Entire animal, dorsal view. A, Catapagurus tuberculosus (Asakura, 1999), male (sl 4.9 mm), RUMF-ZC-6031 (IE188); B, Pagurixus longipes Osawa, Fujita &amp; Okuno, 2006, male (sl 2.0 mm), RUMF-ZC-6033 (HD2017-133); C, Pagurixus nomurai Komai &amp; Asakura, 1995, female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65); D, Pagurixus nomurai Komai &amp; Asakura, 1995, ovigerous female (sl 4.4 mm), RUMF-ZC-6040 (SH102)." httpUri="https://zenodo.org/record/2627822/files/figure.png" pageId="9" pageNumber="472">Figs. 2C, D</figureCitation>
,
<figureCitation id="A6912A49D363FF82FEF13970FEF6FCBC" box="[303,370,822,847]" captionStart="FIGURE 5" captionStartId="13.[151,250,1477,1499]" captionTargetBox="[254,1336,832,1406]" captionTargetId="figure@13.[151,1436,807,1455]" captionTargetPageId="13" captionText="FIGURE 5. Pagurixus nomurai Komai &amp; Asakura, 1995: female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65) (A, B); Pylopaguropsis granulata Asakura, 2000: female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31) (C–E); Pylopaguropsis keijii McLaughlin &amp; Haig, 1989: male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87) (F). A, carpus of left cheliped, dorsal view; B, dactylus and propodus of left third pereopod, lateral view; C, right chela, dorsal view; D, carpus and merus of left second pereopod, lateral view; E, carpus and merus of left third pereopod, lateral view; F, propodus of right third pereopod, lateral view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627828/files/figure.png" pageId="9" pageNumber="472">5A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="76B06547D363FF82FF4939C6FD36FC21" pageId="9" pageNumber="472" type="reference_group">
<paragraph id="3E1536CCD363FF82FF4939C6FD36FC21" blockId="9.[151,1436,894,978]" pageId="9" pageNumber="472">
<treatmentCitationGroup id="1EBA11E2D363FF82FF4939C6FD36FC21" pageId="9" pageNumber="472">
<emphasis id="0CDEEADED363FF82FF4939C6FF7BFC66" box="[151,255,896,917]" italics="true" pageId="9" pageNumber="472">Pagurixus</emphasis>
<taxonomicName id="F9AA4D4FD363FF82FED839C6FD10FC67" authority="Komai &amp; Asakura, 1995: 341" authorityName="Komai &amp; Asakura" authorityPageNumber="341" authorityYear="1995" box="[262,660,894,917]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="0CDEEADED363FF82FED839C6FED8FC66" box="[262,348,896,917]" italics="true" pageId="9" pageNumber="472">nomurai</emphasis>
<treatmentCitation id="BF0B10DDD363FF82FEBD3938FD10FC67" author="Komai, T. &amp; Asakura, A." box="[355,660,894,916]" page="341" pageId="9" pageNumber="472" year="1995">
<bibRefCitation id="5A3B4B3DD363FF82FEBD3938FDE6FC67" author="Komai, T. &amp; Asakura, A." box="[355,610,894,916]" pageId="9" pageNumber="472" pagination="341 - 354" refId="ref10719" refString="Komai, T. &amp; Asakura, A. (1995) Pagurixus nomurai, new species, and addition al record of Pagurixus maorus (Nobili, 1906), hermit crabs from Kume-jima Island, the Ryukyus, Japan (Decapoda: Anomura: Paguridae). Journal of Crustacean Biology, 15 (2), 341 - 354." type="journal article" year="1995">Komai &amp; Asakura, 1995</bibRefCitation>
: 341
</treatmentCitation>
</taxonomicName>
, figs. 1–3 (
<typeStatus id="E111886ED363FF82FCD93938FCB0FC67" box="[775,820,894,916]" pageId="9" pageNumber="472">type</typeStatus>
locality: Kume Island, Ryukyu Islands).—
<treatmentCitation id="BF0B10DDD363FF82FB343938FEBEFC40" author="Osawa, M. &amp; Takeda, M." page="1109" pageId="9" pageNumber="472" year="2004">
<bibRefCitation id="5A3B4B3DD363FF82FB343938FF7AFC40" author="Osawa, M. &amp; Takeda, M." pageId="9" pageNumber="472" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
: 1109
</treatmentCitation>
.—
<treatmentCitation id="BF0B10DDD363FF82FE8639DBFDEAFC40" author="Komai, T. &amp; Osawa, M." box="[344,622,925,947]" page="32" pageId="9" pageNumber="472" year="2006">
<bibRefCitation id="5A3B4B3DD363FF82FE8639DBFDC3FC40" author="Komai, T. &amp; Osawa, M." box="[344,583,925,947]" pageId="9" pageNumber="472" pagination="1 - 107" refId="ref10916" refString="Komai, T. &amp; Osawa, M. (2006) A review of the Pagurixus boninensis species group, with descriptions of six new species (Crustacea: Decapoda: Anomura: Paguridae). Zootaxa, 1214, 1 - 107." type="journal article" year="2006">Komai &amp; Osawa 2006</bibRefCitation>
: 32
</treatmentCitation>
, figs. 12–14, 44C, 47.—
<treatmentCitation id="BF0B10DDD363FF82FCB539DBFBDCFC40" author="Poupin, J. &amp; Bouchard, J. - M. &amp; Dinhut, V. &amp; Cleva, R. &amp; Dumas, J." box="[875,1112,925,947]" page="45" pageId="9" pageNumber="472" year="2013">
<bibRefCitation id="5A3B4B3DD363FF82FCB539DBFBB4FC40" author="Poupin, J. &amp; Bouchard, J. - M. &amp; Dinhut, V. &amp; Cleva, R. &amp; Dumas, J." box="[875,1072,925,947]" pageId="9" pageNumber="472" pagination="1 - 73" refId="ref13047" refString="Poupin, J., Bouchard, J. - M., Dinhut, V., Cleva, R. &amp; Dumas, J. (2013) Anomura (Crustacea Decapoda) from the Mayotte region, western Indian Ocean. Atoll Research Bulletin, 593, i-iv, 1 - 73. https: // doi. org / 10.5479 / si. 00775630.593" type="journal article" year="2013">
Poupin
<emphasis id="0CDEEADED363FF82FC6039D8FC68FC40" box="[958,1004,925,947]" italics="true" pageId="9" pageNumber="472">et al</emphasis>
. 2013
</bibRefCitation>
: 45
</treatmentCitation>
, fig. 23D, E.—
<treatmentCitation id="BF0B10DDD363FF82FB2839DBFEA7FC21" author="Osawa, M. &amp; Fujita, Y." page="45" pageId="9" pageNumber="472" year="2016">
<bibRefCitation id="5A3B4B3DD363FF82FB2839DBFF79FC21" author="Osawa, M. &amp; Fujita, Y." pageId="9" pageNumber="472" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita 2016</bibRefCitation>
: 45
</treatmentCitation>
, fig. 6B.—
<treatmentCitation id="BF0B10DDD363FF82FE5139FAFDDBFC21" author="Tan, H. H. &amp; Naruse, T. &amp; Fujita, Y. &amp; Tan, S. K." box="[399,607,956,978]" page="411" pageId="9" pageNumber="472" year="2014">
<bibRefCitation id="5A3B4B3DD363FF82FE5139FAFDABFC21" author="Tan, H. H. &amp; Naruse, T. &amp; Fujita, Y. &amp; Tan, S. K." box="[399,559,956,978]" pageId="9" pageNumber="472" pagination="406 - 418" refId="ref13379" refString="Tan, H. H., Naruse, T., Fujita, Y. &amp; Tan, S. K. (2014) Observations on the fauna from submarine and associated anchialine caves in Christmas Island, Indian Ocean Territory, Australia. In: Tan, H. H., Orchard, M., Davie, P. J. F. &amp; Ng, P. K. L. (Eds.), Christmas Island and Cocos (Keeling) Islands: Biodiversity and Management Challenges. Raffles Bulletin of Zoology, Supplement, 30, pp. 406 - 418." type="journal article" year="2014">
Tan
<emphasis id="0CDEEADED363FF82FE6039FBFE6FFC21" box="[446,491,956,978]" italics="true" pageId="9" pageNumber="472">et al</emphasis>
. 2014
</bibRefCitation>
: 411
</treatmentCitation>
, fig. 12.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="76B06547D363FF82FF4939B8FA9CFB1C" pageId="9" pageNumber="472" type="materials_examined">
<paragraph id="3E1536CCD363FF82FF4939B8FA9CFB1C" blockId="9.[151,1437,1022,2019]" pageId="9" pageNumber="472">
<emphasis id="0CDEEADED363FF82FF4939B8FCAEFBE4" bold="true" box="[151,810,1022,1047]" pageId="9" pageNumber="472">Material examined. Okinawa Island, Hedo Dome cave:</emphasis>
main tube,
<date id="4A14100CD363FF82FC6C39B9FBEDFBE4" box="[946,1129,1022,1047]" pageId="9" pageNumber="472" value="2016-08-10">10 August 2016</date>
, 1 ovigerous female (sl
<quantity id="F9529B29D363FF82FAA739B8FF43FBC8" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="472" unit="mm" value="5.4">5.4 mm</quantity>
), RUMF-ZC-6034, (
<collectionCode id="58BBAE09D363FF82FE6C3E64FE5AFBC8" box="[434,478,1058,1083]" pageId="9" pageNumber="472">HD</collectionCode>
<date id="4A14100CD363FF82FE033E64FDB9FBC8" box="[477,573,1058,1083]" pageId="9" pageNumber="472" value="2016" valueMax="2051">2016-51</date>
); main tube,
<date id="4A14100CD363FF82FD0B3E65FC0BFBC8" box="[725,911,1058,1083]" pageId="9" pageNumber="472" value="2016-08-12">12 August 2016</date>
, 1 ovigerous female (sl
<quantity id="F9529B29D363FF82FB7B3E65FA85FBC8" box="[1189,1281,1059,1083]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="9" pageNumber="472" unit="mm" value="4.1">4.1 mm</quantity>
), RUMF-ZC- 6035 (
<collectionCode id="58BBAE09D363FF82FF013E00FE8EFBAC" box="[223,266,1094,1119]" pageId="9" pageNumber="472">HD</collectionCode>
<date id="4A14100CD363FF82FED43E00FEE8FBAC" box="[266,364,1094,1119]" pageId="9" pageNumber="472" value="2016" valueMax="2084">2016-84</date>
); main tube,
<quantity id="F9529B29D363FF82FDDE3E01FDBFFBAD" box="[512,571,1095,1119]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="472" unit="m" value="30.0">30 m</quantity>
from entrance, area of rocks,
<date id="4A14100CD363FF82FC4A3E01FBC9FBAC" box="[916,1101,1094,1119]" pageId="9" pageNumber="472" value="2016-08-12">12 August 2016</date>
,
<specimenCount id="28ACFD45D363FF82FB843E01FB22FBAC" box="[1114,1190,1094,1119]" pageId="9" pageNumber="472" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D363FF82FB0D3E00FAABFBAC" box="[1235,1327,1094,1119]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="472" unit="mm" value="4.5">4.5 mm</quantity>
), RUMF- ZC-6036 (
<collectionCode id="58BBAE09D363FF82FED03E2CFEBDFB70" box="[270,313,1130,1155]" pageId="9" pageNumber="472">HD</collectionCode>
<date id="4A14100CD363FF82FEE43E2CFE1EFB70" box="[314,410,1130,1155]" pageId="9" pageNumber="472" value="2016" valueMax="2085">2016-85</date>
); main tube,
<date id="4A14100CD363FF82FDEA3E2DFD75FB70" box="[564,753,1130,1155]" pageId="9" pageNumber="472" value="2016-08-13">13 August 2016</date>
,
<specimenCount id="28ACFD45D363FF82FD213E2DFCDCFB70" box="[767,856,1130,1155]" pageId="9" pageNumber="472" type="male">2 males</specimenCount>
(sl 2.3,
<quantity id="F9529B29D363FF82FC633E2CFB9EFB70" box="[957,1050,1130,1155]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="472" unit="mm" value="4.5">4.5 mm</quantity>
), RUMF-ZC-6037 (
<collectionCode id="58BBAE09D363FF82FADE3E2CFAAFFB70" box="[1280,1323,1130,1155]" pageId="9" pageNumber="472">HD</collectionCode>
2016-103, 104); main tube,
<date id="4A14100CD363FF82FE873EC9FE76FB54" box="[345,498,1167,1191]" pageId="9" pageNumber="472" value="2017-05-22">22 May 2017</date>
,
<specimenCount id="28ACFD45D363FF82FE213EC9FDE5FB54" box="[511,609,1166,1191]" pageId="9" pageNumber="472" type="female">1 female</specimenCount>
(sl 5.0 mm), RUMF-ZC-6038 (
<collectionCode id="58BBAE09D363FF82FC173EC8FC70FB54" box="[969,1012,1166,1191]" pageId="9" pageNumber="472">HD</collectionCode>
<date id="4A14100CD363FF82FC2B3EC8FBD0FB54" box="[1013,1108,1166,1191]" pageId="9" pageNumber="472" value="2017" valueMax="2065">2017-65</date>
); main tube,
<date id="4A14100CD363FF82FB363EC9FA06FB54" box="[1256,1410,1167,1191]" pageId="9" pageNumber="472" value="2017-05-21">21 May 2017</date>
,
<materialsCitation id="8EC23C91D363FF82FA503EC9FDF6FB1C" ID-GBIF-Occurrence="2234219655" collectingDate="2017" collectingDateMax="2085" collectingDateMin="2017" location="Shimoji Island" municipality="Yakata cave" pageId="9" pageNumber="472" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="28ACFD45D363FF82FA503EC9FF49FB38" pageId="9" pageNumber="472" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D363FF82FF233EF5FEDDFB38" box="[253,345,1203,1227]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="9" pageNumber="472" unit="mm" value="4.8">4.8 mm</quantity>
),
<specimenCount id="28ACFD45D363FF82FEB13EF5FE5AFB38" box="[367,478,1202,1227]" pageId="9" pageNumber="472" type="female">2 females</specimenCount>
(sl 3.7,
<quantity id="F9529B29D363FF82FD9F3EF4FD1AFB38" box="[577,670,1202,1227]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="472" unit="mm" value="5.5">5.5 mm</quantity>
), RUMF-ZC-6039 (
<collectionCode id="58BBAE09D363FF82FC5D3EF4FC2AFB38" box="[899,942,1202,1227]" pageId="9" pageNumber="472">HD</collectionCode>
<collectingDate id="5A50E9E4D363FF82FC703EF4FB88FB38" box="[942,1036,1202,1227]" pageId="9" pageNumber="472" value="2017" valueMax="2085">
<date id="4A14100CD363FF82FC703EF4FB88FB38" box="[942,1036,1202,1227]" pageId="9" pageNumber="472" value="2017" valueMax="2085">2017-85</date>
</collectingDate>
, 90).
<emphasis id="0CDEEADED363FF82FB8E3EF4FEB4FB1C" bold="true" pageId="9" pageNumber="472">
<location id="3B756017D363FF82FB8E3EF4FA87FB38" LSID="urn:lsid:plazi:treatment:B60387DAD363FF81FF493952FD05FE06:3B756017D363FF82FB8E3EF4FA87FB38" box="[1104,1283,1202,1227]" municipality="Yakata cave" name="Shimoji Island" pageId="9" pageNumber="472">
Shimoji
<collectingCountry id="46BD765CD363FF82FB693EF4FA87FB38" box="[1207,1283,1202,1227]" name="Iceland" pageId="9" pageNumber="472">Island</collectingCountry>
</location>
, Akuma-no-
<collectingMunicipality id="DE71ACB6D363FF82FF493E90FEAFFB1C" box="[151,299,1238,1263]" pageId="9" pageNumber="472">Yakata cave</collectingMunicipality>
:
</emphasis>
entrance,
<collectingDate id="5A50E9E4D363FF82FE7A3E91FDF6FB1C" box="[420,626,1238,1263]" pageId="9" pageNumber="472" value="2017-09-01">
<date id="4A14100CD363FF82FE7A3E91FDF6FB1C" box="[420,626,1238,1263]" pageId="9" pageNumber="472" value="2017-09-01">1 September 2017</date>
</collectingDate>
</materialsCitation>
, 1 ovigerous female (sl
<quantity id="F9529B29D363FF82FCA03E91FC5CFB1C" box="[894,984,1239,1263]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="472" unit="mm" value="4.4">4.4 mm</quantity>
), RUMF-ZC-6040 (
<collectionCode id="58BBAE09D363FF82FB663E91FB5BFB1C" box="[1208,1247,1239,1263]" pageId="9" pageNumber="472">SH</collectionCode>
102).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D363FF82FF193EBCFCC5FA50" pageId="9" pageNumber="472" type="distribution">
<paragraph id="3E1536CCD363FF82FF193EBCFDB1FAA8" blockId="9.[151,1437,1022,2019]" pageId="9" pageNumber="472">
<emphasis id="0CDEEADED363FF82FF193EBCFEE4FAE0" bold="true" box="[199,352,1274,1299]" pageId="9" pageNumber="472">Distribution.</emphasis>
Widely distributed in the Indo-West Pacific:
<collectingCountry id="46BD765CD363FF82FCA03EBDFC5AFAE0" box="[894,990,1275,1299]" name="Mayotte" pageId="9" pageNumber="472">Mayotte</collectingCountry>
,
<collectingCountry id="46BD765CD363FF82FC303EBCFBD4FAE0" box="[1006,1104,1274,1299]" name="Réunion" pageId="9" pageNumber="472">Réunion</collectingCountry>
,
<collectingCountry id="46BD765CD363FF82FB813EBCFB64FAE0" box="[1119,1248,1274,1299]" name="Philippines" pageId="9" pageNumber="472">Philippines</collectingCountry>
,
<collectingCountry id="46BD765CD363FF82FB2E3EBCFAC1FAE0" box="[1264,1349,1274,1299]" name="Taiwan" pageId="9" pageNumber="472">Taiwan</collectingCountry>
,
<collectingCountry id="46BD765CD363FF82FA8B3EBDFA1CFAE1" box="[1365,1432,1275,1298]" name="Japan" pageId="9" pageNumber="472">Japan</collectingCountry>
,
<collectingCountry id="46BD765CD363FF82FF493F59FF5AFAC4" box="[151,222,1311,1335]" name="Guam" pageId="9" pageNumber="472">Guam</collectingCountry>
, northwestern
<collectingCountry id="46BD765CD363FF82FE5F3F58FE6FFAC4" box="[385,491,1310,1335]" name="Australia" pageId="9" pageNumber="472">Australia</collectingCountry>
,
<collectingCountry id="46BD765CD363FF82FE2B3F59FDC0FAC4" box="[501,580,1311,1335]" name="Samoa" pageId="9" pageNumber="472">Samoa</collectingCountry>
, Cook, and
<collectingCountry id="46BD765CD363FF82FD163F58FC08FAC4" box="[712,908,1310,1335]" name="French Polynesia" pageId="9" pageNumber="472">French Polynesia</collectingCountry>
; subtidal to
<quantity id="F9529B29D363FF82FBCC3F58FBCEFAC5" box="[1042,1098,1310,1335]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="472" unit="m" value="60.0">60 m</quantity>
(Osawa &amp; Chan 2010;
<bibRefCitation id="5A3B4B3DD363FF82FA8E3F58FEC8FAA8" author="Komai, T. &amp; Rahayu, D. L." pageId="9" pageNumber="472" pagination="133 - 141" refId="ref11057" refString="Komai, T. &amp; Rahayu, D. L. (2013) Records of the hermit crab genus Pagurixus Melin, 1939 (Decapoda: Anomura: Paguridae) from shallow coral reefs in the Panglao Islands, the Philippines, with description of a new species. The Raffles Bulletin of Zoology, 61 (1), 133 - 141." type="journal article" year="2013">Komai &amp; Rahayu 2013</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD363FF82FE893F04FDADFAA8" author="Poupin, J. &amp; Bouchard, J. - M. &amp; Dinhut, V. &amp; Cleva, R. &amp; Dumas, J." box="[343,553,1346,1371]" pageId="9" pageNumber="472" pagination="1 - 73" refId="ref13047" refString="Poupin, J., Bouchard, J. - M., Dinhut, V., Cleva, R. &amp; Dumas, J. (2013) Anomura (Crustacea Decapoda) from the Mayotte region, western Indian Ocean. Atoll Research Bulletin, 593, i-iv, 1 - 73. https: // doi. org / 10.5479 / si. 00775630.593" type="journal article" year="2013">
Poupin
<emphasis id="0CDEEADED363FF82FE733F05FE5AFAA8" box="[429,478,1347,1371]" italics="true" pageId="9" pageNumber="472">et al</emphasis>
. 2013
</bibRefCitation>
).
</paragraph>
<paragraph id="3E1536CCD363FF82FF193F20FCC5FA50" blockId="9.[151,1437,1022,2019]" pageId="9" pageNumber="472">
<emphasis id="0CDEEADED363FF82FF193F20FEAEFA8C" bold="true" box="[199,298,1382,1407]" pageId="9" pageNumber="472">Habitat.</emphasis>
Coral reefs; found among branches or plates of dead corals in moats or in submarine caves in forereef slopes (
<bibRefCitation id="5A3B4B3DD363FF82FEC03FCDFDADFA50" author="Osawa, M. &amp; Takeda, M." box="[286,553,1418,1443]" pageId="9" pageNumber="472" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda, 2004</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD363FF82FDEB3FCCFCB1FA50" author="Komai, T. &amp; Osawa, M." box="[565,821,1418,1443]" pageId="9" pageNumber="472" pagination="1 - 107" refId="ref10916" refString="Komai, T. &amp; Osawa, M. (2006) A review of the Pagurixus boninensis species group, with descriptions of six new species (Crustacea: Decapoda: Anomura: Paguridae). Zootaxa, 1214, 1 - 107." type="journal article" year="2006">Komai &amp; Osawa 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D363FF81FF193FE8FD05FE06" lastPageId="10" lastPageNumber="473" pageId="9" pageNumber="472" type="discussion">
<paragraph id="3E1536CCD363FF82FF193FE8FF67F884" blockId="9.[151,1437,1022,2019]" pageId="9" pageNumber="472">
<emphasis id="0CDEEADED363FF82FF193FE8FEBFFA34" bold="true" box="[199,315,1454,1479]" pageId="9" pageNumber="472">Remarks.</emphasis>
In their redescription of
<taxonomicName id="F9AA4D4FD363FF82FDAA3FE9FB27FA34" authority=", Komai &amp; Osawa (2006, fig. 44 C)" authorityName=", Komai &amp; Osawa" authorityYear="2006" box="[628,1187,1454,1479]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="0CDEEADED363FF82FDAA3FE9FD71FA35" box="[628,757,1455,1478]" italics="true" pageId="9" pageNumber="472">P. nomurai</emphasis>
,
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noted that the fresh coloration of the pereopods is “generally brown, olive or bluish gray” and gave a photograph of bluish gray-colored specimen.
<bibRefCitation id="5A3B4B3DD363FF82FED03FB0FE62F9FC" author="Poupin, J. &amp; Bouchard, J. - M. &amp; Dinhut, V. &amp; Cleva, R. &amp; Dumas, J." box="[270,486,1526,1551]" pageId="9" pageNumber="472" pagination="1 - 73" refId="ref13047" refString="Poupin, J., Bouchard, J. - M., Dinhut, V., Cleva, R. &amp; Dumas, J. (2013) Anomura (Crustacea Decapoda) from the Mayotte region, western Indian Ocean. Atoll Research Bulletin, 593, i-iv, 1 - 73. https: // doi. org / 10.5479 / si. 00775630.593" type="journal article" year="2013">
Poupin
<emphasis id="0CDEEADED363FF82FEBA3FB1FE10F9FD" box="[356,404,1526,1550]" italics="true" pageId="9" pageNumber="472">et al</emphasis>
. (2013
</bibRefCitation>
, fig. 23D, E) and
<bibRefCitation id="5A3B4B3DD363FF82FD723FB0FB91F9FC" author="Arima, H." box="[684,1045,1526,1551]" pageId="9" pageNumber="472" refId="ref9388" refString="Arima, H. (2014) Hermit Crabs Illustrated. Seibundo-Shinkosha Co. Ltd, Tokyo, 223 pp. [in Japanese]" type="book" year="2014">Arima (2014, unnumbered figs.)</bibRefCitation>
also showed similarly bluish graycolored specimens of
<taxonomicName id="F9AA4D4FD363FF82FE4E3C5DFD88F9C1" authorityName=", Komai &amp; Osawa" authorityYear="2006" box="[400,524,1563,1586]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="0CDEEADED363FF82FE4E3C5DFD88F9C1" box="[400,524,1563,1586]" italics="true" pageId="9" pageNumber="472">P. nomurai</emphasis>
</taxonomicName>
. All the specimens have been recorded from rather open, coral and rocky reefs, although they usually hided in rubbles or crevices on the reefs. In the present specimens collected from submarine caves, the antennal flagella and pereopods are usually vivid reddish brown (
<figureCitation id="A6912A49D363FF82FBC33C24FBFEF988" box="[1053,1146,1634,1659]" captionStart="FIGURE 2" captionStartId="8.[151,250,1799,1821]" captionTargetBox="[151,1436,519,1777]" captionTargetId="figure@8.[151,1436,519,1777]" captionTargetPageId="8" captionText="FIGURE 2. Entire animal, dorsal view. A, Catapagurus tuberculosus (Asakura, 1999), male (sl 4.9 mm), RUMF-ZC-6031 (IE188); B, Pagurixus longipes Osawa, Fujita &amp; Okuno, 2006, male (sl 2.0 mm), RUMF-ZC-6033 (HD2017-133); C, Pagurixus nomurai Komai &amp; Asakura, 1995, female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65); D, Pagurixus nomurai Komai &amp; Asakura, 1995, ovigerous female (sl 4.4 mm), RUMF-ZC-6040 (SH102)." httpUri="https://zenodo.org/record/2627822/files/figure.png" pageId="9" pageNumber="472">Fig. 2C</figureCitation>
), whereas those of three ovigerous female specimens (sl
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, RUMF-ZC-6034, HD
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; sl
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, RUMF-ZC-6035, HD
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; sl
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, RUMF-ZC-6040, SH102) are rather pale, light brown (
<figureCitation id="A6912A49D363FF82FCA53CECFC52F930" box="[891,982,1706,1731]" captionStart="FIGURE 2" captionStartId="8.[151,250,1799,1821]" captionTargetBox="[151,1436,519,1777]" captionTargetId="figure@8.[151,1436,519,1777]" captionTargetPageId="8" captionText="FIGURE 2. Entire animal, dorsal view. A, Catapagurus tuberculosus (Asakura, 1999), male (sl 4.9 mm), RUMF-ZC-6031 (IE188); B, Pagurixus longipes Osawa, Fujita &amp; Okuno, 2006, male (sl 2.0 mm), RUMF-ZC-6033 (HD2017-133); C, Pagurixus nomurai Komai &amp; Asakura, 1995, female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65); D, Pagurixus nomurai Komai &amp; Asakura, 1995, ovigerous female (sl 4.4 mm), RUMF-ZC-6040 (SH102)." httpUri="https://zenodo.org/record/2627822/files/figure.png" pageId="9" pageNumber="472">Fig. 2D</figureCitation>
). The coloration of the ovigerous female (sl
<quantity id="F9529B29D363FF82FF643C89FE96F915" box="[186,274,1743,1767]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="472" unit="mm" value="4.4">4.4 mm</quantity>
, RUMF-ZC-6040, SH102) generally agrees well with that of a male reported by
<bibRefCitation id="5A3B4B3DD363FF82FB453C89FF73F8F8" author="Osawa, M. &amp; Fujita, Y." pageId="9" pageNumber="472" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita (2016: fig. 6B)</bibRefCitation>
; both specimens were collected from same locality, Akuma-no-Yakata cave in Shimoji Island. The “reddish brown” individuals were often observed by investigations in Hedo Dome cave.
<bibRefCitation id="5A3B4B3DD363FF82FB5B3D51FAB8F8DC" author="Tan, H. H. &amp; Naruse, T. &amp; Fujita, Y. &amp; Tan, S. K." box="[1157,1340,1814,1839]" pageId="9" pageNumber="472" pagination="406 - 418" refId="ref13379" refString="Tan, H. H., Naruse, T., Fujita, Y. &amp; Tan, S. K. (2014) Observations on the fauna from submarine and associated anchialine caves in Christmas Island, Indian Ocean Territory, Australia. In: Tan, H. H., Orchard, M., Davie, P. J. F. &amp; Ng, P. K. L. (Eds.), Christmas Island and Cocos (Keeling) Islands: Biodiversity and Management Challenges. Raffles Bulletin of Zoology, Supplement, 30, pp. 406 - 418." type="journal article" year="2014">
Tan
<emphasis id="0CDEEADED363FF82FB683D51FB6CF8DD" box="[1206,1256,1814,1838]" italics="true" pageId="9" pageNumber="472">et al</emphasis>
. (2014
</bibRefCitation>
: fig. 12) also gave a photograph of “reddish brown” specimen referred to
<taxonomicName id="F9AA4D4FD363FF82FCB53D7DFC61F8A1" authorityName=", Komai &amp; Osawa" authorityYear="2006" box="[875,997,1851,1874]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="0CDEEADED363FF82FCB53D7DFC61F8A1" box="[875,997,1851,1874]" italics="true" pageId="9" pageNumber="472">P. nomurai</emphasis>
</taxonomicName>
from Thunderdome Cave in
<collectingCountry id="46BD765CD363FF82FAF43D7CFF5BF884" name="Christmas Island" pageId="9" pageNumber="472">Christmas Island</collectingCountry>
.
</paragraph>
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In comparison with the redescription of
<taxonomicName id="F9AA4D4FD363FF82FD4E3DC5FC88F869" authorityName=", Komai &amp; Osawa" authorityYear="2006" box="[656,780,1923,1946]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="0CDEEADED363FF82FD4E3DC5FC88F869" box="[656,780,1923,1946]" italics="true" pageId="9" pageNumber="472">P. nomurai</emphasis>
</taxonomicName>
by
<bibRefCitation id="5A3B4B3DD363FF82FCE63DC4FBC9F868" author="Komai, T. &amp; Osawa, M." box="[824,1101,1922,1947]" pageId="9" pageNumber="472" pagination="1 - 107" refId="ref10916" refString="Komai, T. &amp; Osawa, M. (2006) A review of the Pagurixus boninensis species group, with descriptions of six new species (Crustacea: Decapoda: Anomura: Paguridae). Zootaxa, 1214, 1 - 107." type="journal article" year="2006">Komai &amp; Osawa (2006)</bibRefCitation>
, the present specimens of the “reddish brown” and “pale light brown” morphs have proportionally more slender carpus of the left cheliped in general; 4.4–5.4 (reddish brown morph;
<figureCitation id="A6912A49D363FF82FD843D8CFD34F810" box="[602,688,1994,2019]" captionStart="FIGURE 5" captionStartId="13.[151,250,1477,1499]" captionTargetBox="[254,1336,832,1406]" captionTargetId="figure@13.[151,1436,807,1455]" captionTargetPageId="13" captionText="FIGURE 5. Pagurixus nomurai Komai &amp; Asakura, 1995: female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65) (A, B); Pylopaguropsis granulata Asakura, 2000: female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31) (C–E); Pylopaguropsis keijii McLaughlin &amp; Haig, 1989: male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87) (F). A, carpus of left cheliped, dorsal view; B, dactylus and propodus of left third pereopod, lateral view; C, right chela, dorsal view; D, carpus and merus of left second pereopod, lateral view; E, carpus and merus of left third pereopod, lateral view; F, propodus of right third pereopod, lateral view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627828/files/figure.png" pageId="9" pageNumber="472">Fig. 5A</figureCitation>
) and 4.3–5.0 (pale light brown morph) versus 3.0–4.3 length of the width. Additionally, the chelipeds of the present specimens are less setose with shorter setae, particularly in females, than those of the
<bibRefCitation id="5A3B4B3DD360FF81FE1E3AFAFD67FF26" author="Komai, T. &amp; Osawa, M." box="[448,739,188,213]" pageId="10" pageNumber="473" pagination="1 - 107" refId="ref10916" refString="Komai, T. &amp; Osawa, M. (2006) A review of the Pagurixus boninensis species group, with descriptions of six new species (Crustacea: Decapoda: Anomura: Paguridae). Zootaxa, 1214, 1 - 107." type="journal article" year="2006">Komai &amp; Osawa’s (2006)</bibRefCitation>
material, although the authors cited the presence of relatively numerous tufts of stiff setae on the appendages is observed in large specimens, regardless of sex.
</paragraph>
<paragraph id="3E1536CCD360FF81FF193B42FD05FE06" blockId="10.[151,1436,151,501]" pageId="10" pageNumber="473">
The differences in fresh coloration and elongation of the left cheliped of the present specimens could be attributed to the degree of adaptation for the dark habitat in submarine caves, although the morphometric range of the ambulatory legs in the present specimens agrees with that in the material of
<taxonomicName id="F9AA4D4FD360FF81FBFA3B0BFB18FE97" authorityName=", Komai &amp; Osawa" authorityYear="2006" box="[1060,1180,333,356]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="0CDEEADED360FF81FBFA3B0BFB18FE97" box="[1060,1180,333,356]" italics="true" pageId="10" pageNumber="473">P. nomurai</emphasis>
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reported by
<bibRefCitation id="5A3B4B3DD360FF81FAEC3B0AFEBDFE7B" author="Komai, T. &amp; Osawa, M." pageId="10" pageNumber="473" pagination="1 - 107" refId="ref10916" refString="Komai, T. &amp; Osawa, M. (2006) A review of the Pagurixus boninensis species group, with descriptions of six new species (Crustacea: Decapoda: Anomura: Paguridae). Zootaxa, 1214, 1 - 107." type="journal article" year="2006">Komai &amp; Osawa (2006)</bibRefCitation>
; for example on the propodi, 5.4–6.5 (
<figureCitation id="A6912A49D360FF81FD343B29FCC4FE7B" box="[746,832,367,392]" captionStart="FIGURE 5" captionStartId="13.[151,250,1477,1499]" captionTargetBox="[254,1336,832,1406]" captionTargetId="figure@13.[151,1436,807,1455]" captionTargetPageId="13" captionText="FIGURE 5. Pagurixus nomurai Komai &amp; Asakura, 1995: female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65) (A, B); Pylopaguropsis granulata Asakura, 2000: female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31) (C–E); Pylopaguropsis keijii McLaughlin &amp; Haig, 1989: male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87) (F). A, carpus of left cheliped, dorsal view; B, dactylus and propodus of left third pereopod, lateral view; C, right chela, dorsal view; D, carpus and merus of left second pereopod, lateral view; E, carpus and merus of left third pereopod, lateral view; F, propodus of right third pereopod, lateral view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627828/files/figure.png" pageId="10" pageNumber="473">Fig. 5B</figureCitation>
) versus 5.5–6.5 times longer than high. To determine as if these distinctions are interspecific variations and a cryptic species exists under the name
<taxonomicName id="F9AA4D4FD360FF81FACA3BD3FA11FE5F" authorityName=", Komai &amp; Osawa" authorityYear="2006" box="[1300,1429,405,428]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="0CDEEADED360FF81FACA3BD3FA11FE5F" box="[1300,1429,405,428]" italics="true" pageId="10" pageNumber="473">P. nomurai</emphasis>
</taxonomicName>
, molecular data and more careful morphological comparison of the present specimens and other material referred to
<taxonomicName id="F9AA4D4FD360FF81FF493B9BFE95FE07" authorityName=", Komai &amp; Osawa" authorityYear="2006" box="[151,273,477,500]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="0CDEEADED360FF81FF493B9BFE95FE07" box="[151,273,477,500]" italics="true" pageId="10" pageNumber="473">P. nomurai</emphasis>
</taxonomicName>
are needed and now in progress.
</paragraph>
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180
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<emphasis id="0CDEEADED363FF82FF493AD1FE90FF42" bold="true" box="[151,276,151,177]" italics="true" pageId="9" pageNumber="472">Pagurixus</emphasis>
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<emphasis id="0CDEEADED363FF82FEC23AD1FE05FF42" bold="true" box="[284,385,151,177]" italics="true" pageId="9" pageNumber="472">longipes</emphasis>
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(
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)
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<emphasis id="0CDEEADED363FF82FF493B45FF7BFEEB" box="[151,255,259,280]" italics="true" pageId="9" pageNumber="472">Pagurixus</emphasis>
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<emphasis id="0CDEEADED363FF82FEDB3B44FEDFFEEB" box="[261,347,258,280]" italics="true" pageId="9" pageNumber="472">longipes</emphasis>
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: 36
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</taxonomicName>
, figs. 4–6, 7C (type locality: Kume Island, Ryukyu Islands).—
<treatmentCitation id="BF0B10DDD363FF82FAE43B44FE00FEC4" author="Komai, T. &amp; Rahayu, D. L." page="134" pageId="9" pageNumber="472" year="2013">
<bibRefCitation id="5A3B4B3DD363FF82FAE43B44FED5FEC4" author="Komai, T. &amp; Rahayu, D. L." pageId="9" pageNumber="472" pagination="133 - 141" refId="ref11057" refString="Komai, T. &amp; Rahayu, D. L. (2013) Records of the hermit crab genus Pagurixus Melin, 1939 (Decapoda: Anomura: Paguridae) from shallow coral reefs in the Panglao Islands, the Philippines, with description of a new species. The Raffles Bulletin of Zoology, 61 (1), 133 - 141." type="journal article" year="2013">Komai &amp; Rahayu 2013</bibRefCitation>
: 134
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.
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</paragraph>
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<subSubSection id="76B06547D363FF82FF493B22FE00FE37" pageId="9" pageNumber="472" type="materials_examined">
<paragraph id="3E1536CCD363FF82FF493B22FE00FE37" blockId="9.[151,1436,356,704]" pageId="9" pageNumber="472">
<materialsCitation id="8EC23C91D363FF82FF493B22FC8BFE53" ID-GBIF-Occurrence="2465041916" collectingDate="2017-05-21" collectionCode="HD" country="Japan" location="main tube" pageId="9" pageNumber="472" specimenCode="RUMF-ZC-6032" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa Island, Hedo Dome cave">
<emphasis id="0CDEEADED363FF82FF493B22FCC8FE8E" bold="true" box="[151,844,356,381]" pageId="9" pageNumber="472">
Material examined.
<collectingRegion id="FC6EF82ED363FF82FE4A3B22FCC3FE8E" box="[404,839,356,381]" pageId="9" pageNumber="472">Okinawa Island, Hedo Dome cave</collectingRegion>
:
</emphasis>
<location id="3B756017D363FF82FC873B22FC55FE8E" LSID="urn:lsid:plazi:treatment:B60387DAD363FF82FF493AD1FEF1FD33:3B756017D363FF82FC873B22FC55FE8E" box="[857,977,356,381]" country="Japan" name="main tube" pageId="9" pageNumber="472" stateProvince="Okinawa Island, Hedo Dome cave">main tube</location>
,
<date id="4A14100CD363FF82FC3D3B22FB0CFE8F" box="[995,1160,356,381]" pageId="9" pageNumber="472" value="2017-05-21">
<collectingDate id="5A50E9E4D363FF82FC3D3B22FB0CFE8F" box="[995,1160,356,381]" pageId="9" pageNumber="472" value="2017-05-21">21 May 2017</collectingDate>
</date>
,
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parasitized by peltogastrid (sl
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),
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(HD
<date id="4A14100CD363FF82FD793BCEFC83FE53" box="[679,775,392,416]" pageId="9" pageNumber="472" value="2017" valueMax="2088">2017-88</date>
)
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;
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<location id="3B756017D363FF82FCC43BC1FC0FFE53" LSID="urn:lsid:plazi:treatment:B60387DAD363FF82FF493AD1FEF1FD33:3B756017D363FF82FCC43BC1FC0FFE53" box="[794,907,391,416]" country="Japan" name="main tube" pageId="9" pageNumber="472" stateProvince="Okinawa Island, Hedo Dome Cave">main tube</location>
,
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<collectingDate id="5A50E9E4D363FF82FC483BCEFBA9FE53" box="[918,1069,392,416]" pageId="9" pageNumber="472" value="2017-05-23">23 May 2017</collectingDate>
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(sl 2.0 mm),
<specimenCode id="6E0C9EB7D363FF82FAC83BCEFF4BFE36" pageId="9" pageNumber="472">RUMF-ZC- 6033</specimenCode>
(HD2017-133)
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.
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<paragraph id="3E1536CCD363FF82FF193B89FC28FDFE" blockId="9.[151,1436,356,704]" pageId="9" pageNumber="472">
<materialsCitation id="8EC23C91D363FF82FF193B89FEF1FDFF" ID-GBIF-Occurrence="2465041910" country="Japan" pageId="9" pageNumber="472" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island And Kume Island)">
<emphasis id="0CDEEADED363FF82FF193B89FEE4FE1B" bold="true" box="[199,352,463,488]" pageId="9" pageNumber="472">Distribution.</emphasis>
Philippines (Panglao Island) and Ryukyu Islands (Okinawa Island and Kume Island) in southwestern Japan
</materialsCitation>
;
<materialsCitation id="8EC23C91D363FF82FE5E3BB2FD7FFDFF" ID-GBIF-Occurrence="2465041907" box="[384,763,500,525]" country="Japan" pageId="9" pageNumber="472" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island And Kume Island)">
<quantity id="F9529B29D363FF82FE5E3BB2FE67FDFF" box="[384,483,500,525]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="9" pageNumber="472" unit="m" value="25.0" valueMax="35.0" valueMin="15.0">15–35 m</quantity>
(
<bibRefCitation id="5A3B4B3DD363FF82FE2C3BB2FD7FFDFF" author="Komai, T. &amp; Rahayu, D. L." box="[498,763,500,525]" pageId="9" pageNumber="472" pagination="133 - 141" refId="ref11057" refString="Komai, T. &amp; Rahayu, D. L. (2013) Records of the hermit crab genus Pagurixus Melin, 1939 (Decapoda: Anomura: Paguridae) from shallow coral reefs in the Panglao Islands, the Philippines, with description of a new species. The Raffles Bulletin of Zoology, 61 (1), 133 - 141." type="journal article" year="2013">Komai &amp; Rahayu 2013</bibRefCitation>
</materialsCitation>
;
<materialsCitation id="8EC23C91D363FF82FCD93BB3FC2CFDFE" ID-GBIF-Occurrence="2465041915" box="[775,936,500,525]" country="Japan" pageId="9" pageNumber="472" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island And Kume Island)">present study)</materialsCitation>
.
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<paragraph id="3E1536CCD363FF82FF193851FA9DFDC3" blockId="9.[151,1436,356,704]" box="[199,1305,535,560]" pageId="9" pageNumber="472">
<emphasis id="0CDEEADED363FF82FF193851FEAEFDC3" bold="true" box="[199,298,535,560]" pageId="9" pageNumber="472">Habitat.</emphasis>
Coral reefs; often found in submarine caves (
<bibRefCitation id="5A3B4B3DD363FF82FCF7385EFC7CFDC3" author="Osawa, M. &amp; Fujita, Y. &amp; Okuno, J." box="[809,1016,535,560]" pageId="9" pageNumber="472" pagination="27 - 45" refId="ref12872" refString="Osawa, M., Fujita, Y. &amp; Okuno, J. (2006) Two new species of Pagurixus (Crustacea: Decapoda: Anomura: Paguridae) from submarine caves of the Ryukyu Islands, southwestern Japan. Zootaxa, 1148, 27 - 45." type="journal article" year="2006">
Osawa
<emphasis id="0CDEEADED363FF82FCA0385FFC2BFDC3" box="[894,943,536,560]" italics="true" pageId="9" pageNumber="472">et al</emphasis>
. 2006
</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD363FF82FBDA3851FA89FDC3" author="Komai, T. &amp; Rahayu, D. L." box="[1028,1293,535,560]" pageId="9" pageNumber="472" pagination="133 - 141" refId="ref11057" refString="Komai, T. &amp; Rahayu, D. L. (2013) Records of the hermit crab genus Pagurixus Melin, 1939 (Decapoda: Anomura: Paguridae) from shallow coral reefs in the Panglao Islands, the Philippines, with description of a new species. The Raffles Bulletin of Zoology, 61 (1), 133 - 141." type="journal article" year="2013">Komai &amp; Rahayu 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D363FF82FF19387AFEF1FD33" pageId="9" pageNumber="472" type="discussion">
<paragraph id="3E1536CCD363FF82FF19387AFEF1FD33" blockId="9.[151,1436,356,704]" pageId="9" pageNumber="472">
<emphasis id="0CDEEADED363FF82FF19387AFEBFFDA6" bold="true" box="[199,315,572,597]" pageId="9" pageNumber="472">Remarks.</emphasis>
In the Ryukyu Islands,
<taxonomicName id="F9AA4D4FD363FF82FD90387BFD4EFDA7" authorityName="Osawa, Fujita &amp; Okuno" authorityYear="2006" box="[590,714,572,596]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="0CDEEADED363FF82FD90387BFD4EFDA7" box="[590,714,572,596]" italics="true" pageId="9" pageNumber="472">P. longipes</emphasis>
</taxonomicName>
was previously recorded from only a submarine cave in Kume Island (
<bibRefCitation id="5A3B4B3DD363FF82FF353826FE3AFD8B" author="Osawa, M. &amp; Fujita, Y. &amp; Okuno, J." box="[235,446,607,632]" pageId="9" pageNumber="472" pagination="27 - 45" refId="ref12872" refString="Osawa, M., Fujita, Y. &amp; Okuno, J. (2006) Two new species of Pagurixus (Crustacea: Decapoda: Anomura: Paguridae) from submarine caves of the Ryukyu Islands, southwestern Japan. Zootaxa, 1148, 27 - 45." type="journal article" year="2006">
Osawa
<emphasis id="0CDEEADED363FF82FE9F3827FEF0FD8B" box="[321,372,608,632]" italics="true" pageId="9" pageNumber="472">et al</emphasis>
. 2006
</bibRefCitation>
;
<tableCitation id="73280377D363FF82FE153819FDA5FD8B" box="[459,545,607,632]" captionStart="TABLE 2" captionStartId="6.[147,233,596,618]" captionText="TABLE 2. Submarine caves surveyed and hermit crab species collected in three islands of the Ryukyu Islands, southwestern Japan." pageId="9" pageNumber="472">Table 2</tableCitation>
), although the species has been also known from reef walls in Panglao Island, the Philippines (
<bibRefCitation id="5A3B4B3DD363FF82FE8E38C2FDE5FD6F" author="Komai, T. &amp; Rahayu, D. L." box="[336,609,644,669]" pageId="9" pageNumber="472" pagination="133 - 141" refId="ref11057" refString="Komai, T. &amp; Rahayu, D. L. (2013) Records of the hermit crab genus Pagurixus Melin, 1939 (Decapoda: Anomura: Paguridae) from shallow coral reefs in the Panglao Islands, the Philippines, with description of a new species. The Raffles Bulletin of Zoology, 61 (1), 133 - 141." type="journal article" year="2013">Komai &amp; Rahayu 2013</bibRefCitation>
). The present specimens represent the second record of the species from the Ryukyu Islands.
</paragraph>
</subSubSection>
</treatment>
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265
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<treatment id="B60387DAD366FF86FF493CC6FD9DFD03" ID-DOI="http://doi.org/10.5281/zenodo.5308572" ID-GBIF-Taxon="154533464" ID-Zenodo-Dep="5308572" LSID="urn:lsid:plazi:treatment:B60387DAD366FF86FF493CC6FD9DFD03" httpUri="http://treatment.plazi.org/id/B60387DAD366FF86FF493CC6FD9DFD03" lastPageId="13" lastPageNumber="476" pageId="12" pageNumber="475">
<subSubSection id="76B06547D366FF87FF493CC6FD67F969" box="[151,739,1664,1690]" pageId="12" pageNumber="475" type="nomenclature">
<paragraph id="3E1536CCD366FF87FF493CC6FD67F969" blockId="12.[151,739,1664,1724]" box="[151,739,1664,1690]" pageId="12" pageNumber="475">
<heading id="655D81A0D366FF87FF493CC6FD67F969" bold="true" box="[151,739,1664,1690]" fontSize="11" level="1" pageId="12" pageNumber="475" reason="1">
<emphasis id="0CDEEADED366FF87FF493CC6FD67F969" bold="true" box="[151,739,1664,1690]" pageId="12" pageNumber="475">
<emphasis id="0CDEEADED366FF87FF493CC6FED5F969" bold="true" box="[151,337,1664,1690]" italics="true" pageId="12" pageNumber="475">Pylopaguropsis</emphasis>
<taxonomicName id="F9AA4D4FD366FF87FE863CC6FD67F969" authority="McLaughlin &amp; Haig, 1989" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[344,739,1664,1690]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="475" phylum="Arthropoda" rank="species" species="keijii">
<emphasis id="0CDEEADED366FF87FE863CC6FE10F969" bold="true" box="[344,404,1664,1690]" italics="true" pageId="12" pageNumber="475">keijii</emphasis>
<bibRefCitation id="5A3B4B3DD366FF87FE453CC6FD67F969" author="McLaughlin, P. A. &amp; Haig, J." box="[411,739,1664,1690]" pageId="12" pageNumber="475" pagination="123 - 171" refId="ref11585" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra Henderson, P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22 (2), 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig, 1989</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="76B06547D366FF87FF493CE5FEB1F94F" box="[151,309,1699,1724]" pageId="12" pageNumber="475" type="description">
<paragraph id="3E1536CCD366FF87FF493CE5FEB1F94F" blockId="12.[151,739,1664,1724]" box="[151,309,1699,1724]" pageId="12" pageNumber="475">
(
<figureCitation id="A6912A49D366FF87FF413CE5FE87F948" box="[159,259,1699,1724]" captionStart="FIGURE 3" captionStartId="11.[151,250,1378,1400]" captionTargetBox="[151,1436,195,1356]" captionTargetId="figure@11.[151,1436,195,1356]" captionTargetPageId="11" captionText="FIGURE 3. Entire animal, dorsal view. A, Pylopaguropsis bellula Osawa &amp; Okuno, 2007, male (sl 4.1 mm), RUMF-ZC-6042 (IE17); B, Pylopaguropsis fimbriata McLaughlin &amp; Haig, 1989, ovigerous female (sl 4.4 mm), RUMF-ZC-6046 (IE179); C, Pylopaguropsis granulata Asakura, 2000, female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31); D, Pylopaguropsis keijii McLaughlin &amp; Haig, 1989, male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87); E, Pylopaguropsis speciosa McLaughlin &amp; Haig, 1989, ovigerous female (sl 3.7 mm), RUMF-ZC-6058 (SH104)." httpUri="https://zenodo.org/record/2627824/files/figure.png" pageId="12" pageNumber="475">Figs. 3D</figureCitation>
,
<figureCitation id="A6912A49D366FF87FED03CE5FEA9F94F" box="[270,301,1699,1724]" captionStart="FIGURE 5" captionStartId="13.[151,250,1477,1499]" captionTargetBox="[254,1336,832,1406]" captionTargetId="figure@13.[151,1436,807,1455]" captionTargetPageId="13" captionText="FIGURE 5. Pagurixus nomurai Komai &amp; Asakura, 1995: female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65) (A, B); Pylopaguropsis granulata Asakura, 2000: female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31) (C–E); Pylopaguropsis keijii McLaughlin &amp; Haig, 1989: male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87) (F). A, carpus of left cheliped, dorsal view; B, dactylus and propodus of left third pereopod, lateral view; C, right chela, dorsal view; D, carpus and merus of left second pereopod, lateral view; E, carpus and merus of left third pereopod, lateral view; F, propodus of right third pereopod, lateral view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627828/files/figure.png" pageId="12" pageNumber="475">5F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="76B06547D366FF87FF493CADFD55F8EC" pageId="12" pageNumber="475" type="reference_group">
<paragraph id="3E1536CCD366FF87FF493CADFD55F8EC" blockId="12.[151,1436,1771,1823]" pageId="12" pageNumber="475">
<treatmentCitationGroup id="1EBA11E2D366FF87FF493CADFD55F8EC" infered="true" pageId="12" pageNumber="475">
<emphasis id="0CDEEADED366FF87FF493CADFEB1F8F2" box="[151,309,1771,1793]" italics="true" pageId="12" pageNumber="475">Pylopaguropsis</emphasis>
<taxonomicName id="F9AA4D4FD366FF87FEE23CADFD3AF8F2" authority="McLaughlin &amp; Haig, 1989: 150" authorityName="McLaughlin &amp; Haig" authorityPageNumber="150" authorityYear="1989" box="[316,702,1771,1793]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="475" phylum="Arthropoda" rank="species" species="keijii">
<emphasis id="0CDEEADED366FF87FEE23CADFEEBF8F2" box="[316,367,1771,1793]" italics="true" pageId="12" pageNumber="475">keijii</emphasis>
<treatmentCitation id="BF0B10DDD366FF87FEA83CADFD3AF8F2" author="McLaughlin, P. A. &amp; Haig, J." box="[374,702,1771,1793]" page="150" pageId="12" pageNumber="475" year="1989">
<bibRefCitation id="5A3B4B3DD366FF87FEA83CADFD0FF8F2" author="McLaughlin, P. A. &amp; Haig, J." box="[374,651,1771,1793]" pageId="12" pageNumber="475" pagination="123 - 171" refId="ref11585" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra Henderson, P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22 (2), 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig, 1989</bibRefCitation>
: 150
</treatmentCitation>
</taxonomicName>
, figs.
<date id="4A14100CD366FF87FD203CADFC97F8F2" box="[766,787,1771,1793]" pageId="12" pageNumber="475">2i</date>
, 3d, 5d, 7e, 9d, 11d, 13d (
<typeStatus id="E111886ED366FF87FBFC3CADFBCBF8F2" box="[1058,1103,1771,1793]" pageId="12" pageNumber="475">type</typeStatus>
locality: Oahu Island, Hawaiian Islands).—
<treatmentCitation id="BF0B10DDD366FF87FEED3D4FFE69F8EC" author="Asakura, A." box="[307,493,1801,1823]" page="82" pageId="12" pageNumber="475" year="2000">
<bibRefCitation id="5A3B4B3DD366FF87FEED3D4FFE4CF8EC" author="Asakura, A." box="[307,456,1801,1823]" pageId="12" pageNumber="475" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
: 82
</treatmentCitation>
, figs. 6, 7, 19B, C, 20.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="76B06547D366FF87FF493D0AFE7FF822" pageId="12" pageNumber="475" type="materials_examined">
<paragraph id="3E1536CCD366FF87FF493D0AFE7FF822" blockId="12.[151,1436,1868,2037]" pageId="12" pageNumber="475">
<emphasis id="0CDEEADED366FF87FF493D0AFCB3F896" bold="true" box="[151,823,1868,1893]" pageId="12" pageNumber="475">
Material examined. Ie
<materialsCitation id="8EC23C91D366FF87FE7A3D0AFCB3F896" ID-GBIF-Occurrence="2234219647" box="[420,823,1868,1893]" collectingDate="2017-06-24" country="Iceland" location="Island" pageId="12" pageNumber="475" specimenCount="1" specimenCount-female="1">
<collectingCountry id="46BD765CD366FF87FE7A3D0AFE75F896" box="[420,497,1868,1893]" name="Iceland" pageId="12" pageNumber="475">Island</collectingCountry>
, unnamed submarine cave:
</materialsCitation>
</emphasis>
entrance,
<collectingDate id="5A50E9E4D366FF87FC753D0BFBC6F896" box="[939,1090,1869,1893]" pageId="12" pageNumber="475" value="2017-06-24">
<date id="4A14100CD366FF87FC753D0BFBC6F896" box="[939,1090,1869,1893]" pageId="12" pageNumber="475" value="2017-06-24">24 June 2017</date>
</collectingDate>
,
<specimenCount id="28ACFD45D366FF87FB933D0BFB2AF896" box="[1101,1198,1868,1893]" pageId="12" pageNumber="475" type="female">1 female</specimenCount>
(sl
<quantity id="F9529B29D366FF87FB093D0BFAB5F896" box="[1239,1329,1869,1893]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="475" unit="mm" value="2.1">2.1 mm</quantity>
),
<materialsCitation id="8EC23C91D366FF87FA9D3D0BFAF9F87A" ID-GBIF-Occurrence="2234219657" collectingDate="2016-08-12" collectionCode="IE" country="Iceland" location="Okinawa Island" municipality="Hedo Dome cave" pageId="12" pageNumber="475" specimenCode="IE172" specimenCount="1" specimenCount-male="1">
RUMF- ZC-6053 (
<specimenCode id="6E0C9EB7D366FF87FECF3D37FEDFF87A" box="[273,347,1905,1929]" collectionCode="IE" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="12" pageNumber="475">
<collectionCode id="58BBAE09D366FF87FECF3D37FEABF87A" box="[273,303,1905,1929]" pageId="12" pageNumber="475">IE</collectionCode>
172
</specimenCode>
).
<emphasis id="0CDEEADED366FF87FEAD3D36FCACF87A" bold="true" box="[371,808,1904,1929]" pageId="12" pageNumber="475">
<location id="3B756017D366FF87FEAD3D36FDBDF87A" LSID="urn:lsid:plazi:treatment:B60387DAD366FF86FF493CC6FD9DFD03:3B756017D366FF87FEAD3D36FDBDF87A" box="[371,569,1904,1929]" country="Iceland" municipality="Hedo Dome cave" name="Okinawa Island" pageId="12" pageNumber="475">
Okinawa
<collectingCountry id="46BD765CD366FF87FE333D36FDBDF87A" box="[493,569,1904,1929]" name="Iceland" pageId="12" pageNumber="475">Island</collectingCountry>
</location>
,
<collectingMunicipality id="DE71ACB6D366FF87FD953D36FCA7F87A" box="[587,803,1904,1929]" pageId="12" pageNumber="475">Hedo Dome cave</collectingMunicipality>
:
</emphasis>
main tube,
<collectingDate id="5A50E9E4D366FF87FC633D37FBFBF87A" box="[957,1151,1904,1929]" pageId="12" pageNumber="475" value="2016-08-12">
<date id="4A14100CD366FF87FC633D37FBFBF87A" box="[957,1151,1904,1929]" pageId="12" pageNumber="475" value="2016-08-12">12 August 2016</date>
</collectingDate>
,
<specimenCount id="28ACFD45D366FF87FB513D37FB64F87A" box="[1167,1248,1904,1929]" pageId="12" pageNumber="475" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D366FF87FAC83D37FAF2F87A" box="[1302,1398,1905,1929]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="475" unit="mm" value="2.8">2.8 mm</quantity>
)
</materialsCitation>
, 1 ovigerous female (sl 2.0 mm), RUMF-ZC-6054 (
<collectionCode id="58BBAE09D366FF87FD183DD2FD75F85E" box="[710,753,1940,1965]" pageId="12" pageNumber="475">HD</collectionCode>
<date id="4A14100CD366FF87FD2F3DD2FCCBF85E" box="[753,847,1940,1965]" pageId="12" pageNumber="475" value="2016" valueMax="2081">2016-81</date>
, 82); main tube,
<date id="4A14100CD366FF87FBCC3DD3FB2AF85E" box="[1042,1198,1941,1965]" pageId="12" pageNumber="475" value="2017-05-21">21 May 2017</date>
,
<specimenCount id="28ACFD45D366FF87FB643DD3FA83F85E" box="[1210,1287,1940,1965]" pageId="12" pageNumber="475" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D366FF87FAEB3DD3FA14F85E" box="[1333,1424,1941,1965]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="475" unit="mm" value="1.9">1.9 mm</quantity>
), RUMF-ZC-6055 (
<collectionCode id="58BBAE09D366FF87FEBA3DFFFE14F822" box="[356,400,1977,2001]" pageId="12" pageNumber="475">HD</collectionCode>
<date id="4A14100CD366FF87FE513DFFFE6BF822" box="[399,495,1977,2001]" pageId="12" pageNumber="475" value="2017" valueMax="2087">2017-87</date>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D366FF86FF193D9AFCC2FF0B" lastPageId="13" lastPageNumber="476" pageId="12" pageNumber="475" type="distribution">
<paragraph id="3E1536CCD366FF87FF193D9AFA18F806" blockId="12.[151,1436,1868,2037]" box="[199,1436,2012,2037]" pageId="12" pageNumber="475">
<emphasis id="0CDEEADED366FF87FF193D9AFEE4F806" bold="true" box="[199,352,2012,2037]" pageId="12" pageNumber="475">Distribution.</emphasis>
Wide ranging in the Indo-West Pacific:
<collectingCountry id="46BD765CD366FF87FCEF3D9AFC12F806" box="[817,918,2012,2037]" name="Tanzania" pageId="12" pageNumber="475">Zanzibar</collectingCountry>
,
<collectingCountry id="46BD765CD366FF87FC7D3D9BFB87F806" box="[931,1027,2013,2037]" name="Mayotte" pageId="12" pageNumber="475">Mayotte</collectingCountry>
,
<collectingCountry id="46BD765CD366FF87FBCE3D9AFBFFF806" box="[1040,1147,2012,2037]" name="Maldives" pageId="12" pageNumber="475">Maldives</collectingCountry>
,
<collectingCountry id="46BD765CD366FF87FB563D9BFB4BF806" box="[1160,1231,2013,2037]" name="Guam" pageId="12" pageNumber="475">Guam</collectingCountry>
,
<collectingCountry id="46BD765CD366FF87FB023D9AFA1CF806" box="[1244,1432,2012,2037]" name="Micronesia" pageId="12" pageNumber="475">Caroline Islands</collectingCountry>
,
</paragraph>
<paragraph id="3E1536CCD367FF86FF493ADEFE38FF27" blockId="13.[151,1437,151,752]" pageId="13" pageNumber="476">
<collectingCountry id="46BD765CD367FF86FF493ADEFF53FF5C" box="[151,215,152,175]" name="Japan" pageId="13" pageNumber="476">Japan</collectingCountry>
(Ryukyu and Izu Islands), and Hawaii;
<quantity id="F9529B29D367FF86FD4C3AD1FD78FF5C" box="[658,764,151,176]" metricMagnitude="1" metricUnit="m" metricValue="1.625" metricValueMax="3.0" metricValueMin="0.25" pageId="13" pageNumber="476" unit="m" value="16.25" valueMax="30.0" valueMin="2.5">2.5–30 m</quantity>
(
<bibRefCitation id="5A3B4B3DD367FF86FCD73AD1FBB4FF43" author="McLaughlin, P. A. &amp; Haig, J." box="[777,1072,151,176]" pageId="13" pageNumber="476" pagination="123 - 171" refId="ref11585" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra Henderson, P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22 (2), 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig 1989</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD367FF86FBE53AD1FB5EFF43" author="Asakura, A." box="[1083,1242,151,176]" pageId="13" pageNumber="476" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD367FF86FB383AD1FF57FF26" author="Okuno, J. &amp; Arima, H." pageId="13" pageNumber="476" pagination="29 - 43" refId="ref12530" refString="Okuno, J. &amp; Arima, H. (2006) Additional records of inshore hermit crabs (Crustacea: Decapoda: Anomura) from Izu-ohshima Island, the northern Izu Islands, Japan. Bulletin of the Biogeographical Society of Japan, 61, 29 - 43. [in Japanese with English abstract]" type="journal article" year="2006">Okuno &amp; Arima 2006</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD367FF86FF033AFAFE34FF27" author="Poupin, J. &amp; Bouchard, J. - M. &amp; Dinhut, V. &amp; Cleva, R. &amp; Dumas, J." box="[221,432,188,213]" pageId="13" pageNumber="476" pagination="1 - 73" refId="ref13047" refString="Poupin, J., Bouchard, J. - M., Dinhut, V., Cleva, R. &amp; Dumas, J. (2013) Anomura (Crustacea Decapoda) from the Mayotte region, western Indian Ocean. Atoll Research Bulletin, 593, i-iv, 1 - 73. https: // doi. org / 10.5479 / si. 00775630.593" type="journal article" year="2013">
Poupin
<emphasis id="0CDEEADED367FF86FEEA3AFBFEE1FF27" box="[308,357,188,212]" italics="true" pageId="13" pageNumber="476">et al</emphasis>
. 2013
</bibRefCitation>
).
</paragraph>
<paragraph id="3E1536CCD367FF86FF193A99FCC2FF0B" blockId="13.[151,1437,151,752]" box="[199,838,223,248]" pageId="13" pageNumber="476">
<emphasis id="0CDEEADED367FF86FF193A99FEAEFF0B" bold="true" box="[199,298,223,248]" pageId="13" pageNumber="476">Habitat.</emphasis>
Coral reefs (
<bibRefCitation id="5A3B4B3DD367FF86FE653A99FDD9FF0B" author="Asakura, A." box="[443,605,223,248]" pageId="13" pageNumber="476" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD367FF86FDB63A99FCBEFF0B" author="Poupin, J. &amp; Bouchard, J. - M. &amp; Dinhut, V. &amp; Cleva, R. &amp; Dumas, J." box="[616,826,223,248]" pageId="13" pageNumber="476" pagination="1 - 73" refId="ref13047" refString="Poupin, J., Bouchard, J. - M., Dinhut, V., Cleva, R. &amp; Dumas, J. (2013) Anomura (Crustacea Decapoda) from the Mayotte region, western Indian Ocean. Atoll Research Bulletin, 593, i-iv, 1 - 73. https: // doi. org / 10.5479 / si. 00775630.593" type="journal article" year="2013">
Poupin
<emphasis id="0CDEEADED367FF86FD603AA7FD6BFF0B" box="[702,751,224,248]" italics="true" pageId="13" pageNumber="476">et al</emphasis>
. 2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D367FF86FF193B42FB58FDCE" pageId="13" pageNumber="476" type="discussion">
<paragraph id="3E1536CCD367FF86FF193B42FB58FDCE" blockId="13.[151,1437,151,752]" pageId="13" pageNumber="476">
<emphasis id="0CDEEADED367FF86FF193B42FEBFFEEE" bold="true" box="[199,315,260,285]" pageId="13" pageNumber="476">Remarks.</emphasis>
Variations of coloration on the right cheliped are observed in the present material. The carpus and chela are purple in general and becomes paler or has white tint distally in three small specimens (male, sl
<quantity id="F9529B29D367FF86FAE13B6EFA1CFECC" box="[1343,1432,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="476" unit="mm" value="1.9">1.9 mm</quantity>
, RUMF-ZC-6055, HD
<date id="4A14100CD367FF86FE503B0AFE69FE97" box="[398,493,332,356]" pageId="13" pageNumber="476" value="2017" valueMax="2087">2017-87</date>
; female, sl
<quantity id="F9529B29D367FF86FDB03B0AFD42FE97" box="[622,710,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="13" pageNumber="476" unit="mm" value="2.1">2.1 mm</quantity>
, RUMF-ZC-6053, IE172; ovigerous female, sl 2.0 mm, RUMF- ZC-6054, HD
<date id="4A14100CD367FF86FEEA3B29FE10FE7B" box="[308,404,367,392]" pageId="13" pageNumber="476" value="2016" valueMax="2082">2016-82</date>
), whereas they are whitish for most parts and have purple tint only on the mesial surfaces in the largest specimen (male, sl
<quantity id="F9529B29D367FF86FE343BD2FDC7FE5F" box="[490,579,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="476" unit="mm" value="2.8">2.8 mm</quantity>
, RUMF-ZC-6054, HD
<date id="4A14100CD367FF86FC983BD2FC21FE5E" box="[838,933,404,429]" pageId="13" pageNumber="476" value="2016" valueMax="2081">2016-81</date>
).
<bibRefCitation id="5A3B4B3DD367FF86FC663BD2FB16FE5F" author="Poupin, J. &amp; Bouchard, J. - M. &amp; Dinhut, V. &amp; Cleva, R. &amp; Dumas, J." box="[952,1170,404,429]" pageId="13" pageNumber="476" pagination="1 - 73" refId="ref13047" refString="Poupin, J., Bouchard, J. - M., Dinhut, V., Cleva, R. &amp; Dumas, J. (2013) Anomura (Crustacea Decapoda) from the Mayotte region, western Indian Ocean. Atoll Research Bulletin, 593, i-iv, 1 - 73. https: // doi. org / 10.5479 / si. 00775630.593" type="journal article" year="2013">
Poupin
<emphasis id="0CDEEADED367FF86FBD03BD3FBBAFE5F" box="[1038,1086,404,428]" italics="true" pageId="13" pageNumber="476">et al</emphasis>
. (2013
</bibRefCitation>
, fig. 26) also reported a male specimen (sl 3.0 mm), which agrees well with the present largest specimen in coloration of the right cheliped.
<bibRefCitation id="5A3B4B3DD367FF86FF493B9AFEC2FE07" author="Asakura, A." box="[151,326,476,501]" pageId="13" pageNumber="476" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura (2000)</bibRefCitation>
cited that the right cheliped of
<taxonomicName id="F9AA4D4FD367FF86FD763B9BFD78FE07" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[680,764,476,500]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="476" phylum="Arthropoda" rank="species" species="keijii">
<emphasis id="0CDEEADED367FF86FD763B9BFD78FE07" box="[680,764,476,500]" italics="true" pageId="13" pageNumber="476">P. keijii</emphasis>
</taxonomicName>
is generally fawn and gradually becomes white on the both fingers based on his larger material (sl 3.2,
<quantity id="F9529B29D367FF86FD7D3846FC86FDEB" box="[675,770,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="13" pageNumber="476" unit="mm" value="3.4">3.4 mm</quantity>
). These suggest that the coloration of the right cheliped gradually changes from purple to fawn or white in accordance with the increase of specimen size.
</paragraph>
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<emphasis id="0CDEEADED367FF86FF19380EFEF2FD93" box="[199,374,584,608]" italics="true" pageId="13" pageNumber="476">Pylopaguropsis</emphasis>
<taxonomicName id="F9AA4D4FD367FF86FEA3380EFC84FD93" authority="Asakura &amp; Paulay, 2003" authorityName="Asakura &amp; Paulay" authorityYear="2003" box="[381,768,583,608]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="476" phylum="Arthropoda" rank="species" species="lemaitrei">
<emphasis id="0CDEEADED367FF86FEA3380EFE65FD93" box="[381,481,584,608]" italics="true" pageId="13" pageNumber="476">lemaitrei</emphasis>
<bibRefCitation id="5A3B4B3DD367FF86FE373801FC84FD93" author="Asakura, A. &amp; Paulay, G." box="[489,768,583,608]" pageId="13" pageNumber="476" pagination="13 - 25" refId="ref9611" refString="Asakura, A. &amp; Paulay, G. (2003) Pylopaguropsis lemaitrei, a new species of hermit crab (Decapoda: Anomura: Paguridae) from French Polynesia. Crustacean Research, 32, 13 - 25. https: // doi. org / 10.18353 / crustacea. 32.0 _ 13" type="journal article" year="2003">Asakura &amp; Paulay, 2003</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="46BD765CD367FF86FC603801FB07FD93" box="[958,1155,583,608]" name="French Polynesia" pageId="13" pageNumber="476">French Polynesia</collectingCountry>
, is a sibling species with overall coloration similar to
<taxonomicName id="F9AA4D4FD367FF86FE06382BFDA8FD77" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[472,556,620,644]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="476" phylum="Arthropoda" rank="species" species="keijii">
<emphasis id="0CDEEADED367FF86FE06382BFDA8FD77" box="[472,556,620,644]" italics="true" pageId="13" pageNumber="476">P. keijii</emphasis>
</taxonomicName>
.
<bibRefCitation id="5A3B4B3DD367FF86FDE5382AFCD8FD77" author="Asakura, A. &amp; Paulay, G." box="[571,860,620,645]" pageId="13" pageNumber="476" pagination="13 - 25" refId="ref9611" refString="Asakura, A. &amp; Paulay, G. (2003) Pylopaguropsis lemaitrei, a new species of hermit crab (Decapoda: Anomura: Paguridae) from French Polynesia. Crustacean Research, 32, 13 - 25. https: // doi. org / 10.18353 / crustacea. 32.0 _ 13" type="journal article" year="2003">Asakura &amp; Paulay (2003)</bibRefCitation>
morphologically distinguished
<taxonomicName id="F9AA4D4FD367FF86FB1F382BFAC6FD77" authorityName="Asakura &amp; Paulay" authorityYear="2003" box="[1217,1346,620,644]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="476" phylum="Arthropoda" rank="species" species="lemaitrei">
<emphasis id="0CDEEADED367FF86FB1F382BFAC6FD77" box="[1217,1346,620,644]" italics="true" pageId="13" pageNumber="476">P. lemaitrei</emphasis>
</taxonomicName>
from
<taxonomicName id="F9AA4D4FD367FF86FA59382BFF4BFD5B" authorityName="McLaughlin &amp; Haig" authorityYear="1989" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="476" phylum="Arthropoda" rank="species" species="keijii">
<emphasis id="0CDEEADED367FF86FA59382BFF4BFD5B" italics="true" pageId="13" pageNumber="476">P. keijii</emphasis>
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by three instead of one, longitudinal sulci on the lateral surface of the propodus of the right third pereopod. The present specimens with color variations of the right cheliped undoubtedly belong to
<taxonomicName id="F9AA4D4FD367FF86FB7938F3FB7AFD3F" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[1191,1278,692,716]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="476" phylum="Arthropoda" rank="species" species="keijii">
<emphasis id="0CDEEADED367FF86FB7938F3FB7AFD3F" box="[1191,1278,692,716]" italics="true" pageId="13" pageNumber="476">P. keijii</emphasis>
</taxonomicName>
based on the morphological character (
<figureCitation id="A6912A49D367FF86FE693891FD89FD03" box="[439,525,727,752]" captionStart="FIGURE 5" captionStartId="13.[151,250,1477,1499]" captionTargetBox="[254,1336,832,1406]" captionTargetId="figure@13.[151,1436,807,1455]" captionTargetPageId="13" captionText="FIGURE 5. Pagurixus nomurai Komai &amp; Asakura, 1995: female (sl 5.0 mm), RUMF-ZC-6038 (HD2017-65) (A, B); Pylopaguropsis granulata Asakura, 2000: female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31) (C–E); Pylopaguropsis keijii McLaughlin &amp; Haig, 1989: male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87) (F). A, carpus of left cheliped, dorsal view; B, dactylus and propodus of left third pereopod, lateral view; C, right chela, dorsal view; D, carpus and merus of left second pereopod, lateral view; E, carpus and merus of left third pereopod, lateral view; F, propodus of right third pereopod, lateral view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627828/files/figure.png" pageId="13" pageNumber="476">Fig. 5F</figureCitation>
).
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(
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, figs. 1c, i, 2c, 3e, 5e, 7f, 9e, 11e, 13e (
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locality: Okinawa Island, Ryukyu Islands).—
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, figs. 1A, B, 4, 5, 19A, 20.—
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: 1111
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.
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<emphasis id="0CDEEADED367FF86FF493D28FF69F870" box="[151,237,1902,1923]" italics="true" pageId="13" pageNumber="476">Pagurus</emphasis>
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sp.—
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–5, 124, unnumbered figs.
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<paragraph id="3E1536CCD367FF86FF493DE9FA18F81F" blockId="13.[151,1436,1967,2029]" pageId="13" pageNumber="476">
<materialsCitation id="8EC23C91D367FF86FF493DE9FC08F81E" ID-GBIF-Occurrence="2465041897" collectingDate="2016-08-12" collectionCode="HD" country="Japan" location="main tube, 30 m from entrance, area of rocks" pageId="13" pageNumber="476" specimenCode="RUMF-ZC-6056" specimenCount="3" specimenCount-male="3" stateProvince="Okinawa Island, Hedo Dome cave">
<emphasis id="0CDEEADED367FF86FF493DE9FCA6F83B" bold="true" box="[151,802,1967,1992]" pageId="13" pageNumber="476">
Material examined.
<collectingRegion id="FC6EF82ED367FF86FE583DE9FC99F83B" box="[390,797,1967,1992]" pageId="13" pageNumber="476">Okinawa Island, Hedo Dome cave</collectingRegion>
:
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main tube,
<quantity id="F9529B29D367FF86FC7A3DF6FC59F834" box="[932,989,1968,1992]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="476" unit="m" value="30.0">30 m</quantity>
from entrance, area of rocks
</location>
,
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</date>
,
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(sl
<quantity id="F9529B29D367FF86FEBF3D92FE69F81F" box="[353,493,2004,2028]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="13" pageNumber="476" unit="mm" value="2.55" valueMax="2.9" valueMin="2.2">2.2–2.9 mm</quantity>
),
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(HD
<date id="4A14100CD367FF86FD273D92FCDDF81E" box="[761,857,2004,2029]" pageId="13" pageNumber="476" value="2016" valueMax="2086">2016-86</date>
–88)
</materialsCitation>
;
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<location id="3B756017D367FF86FC463D92FB8FF81E" LSID="urn:lsid:plazi:treatment:B60387DAD367FF85FF493C82FC63FE5F:3B756017D367FF86FC463D92FB8FF81E" box="[920,1035,2004,2029]" country="Japan" name="main tube" pageId="13" pageNumber="476" stateProvince="Okinawa Island, Hedo Dome Cave">main tube</location>
,
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<collectingDate id="5A50E9E4D367FF86FBC93D92FB35F81F" box="[1047,1201,2004,2029]" pageId="13" pageNumber="476" value="2017-05-21">21 May 2017</collectingDate>
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,
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(sl
<quantity id="F9529B29D367FF86FAEB3D92FA14F81F" box="[1333,1424,2004,2028]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="13" pageNumber="476" unit="mm" value="3.1">3.1 mm</quantity>
)
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,
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<paragraph id="3E1536CCD364FF85FF493AD1FC98FF27" blockId="14.[151,1437,151,429]" pageId="14" pageNumber="477">
<materialsCitation id="8EC23C91D364FF85FF493AD1FC9CFF27" ID-GBIF-Occurrence="2465041896" collectingDate="2017-09-01" collectionCode="HD" country="Japan" location="entrance" pageId="14" pageNumber="477" specimenCount="1" stateProvince="Shimoji Island, Akuma-no-Yakata cave">
RUMF-ZC-6057 (HD
<date id="4A14100CD364FF85FE473AD1FE7DFF43" box="[409,505,151,176]" pageId="14" pageNumber="477" value="2017" valueMax="2086">2017-86</date>
).
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<collectingRegion id="FC6EF82ED364FF85FDCB3AD1FB8FFF43" box="[533,1035,151,176]" pageId="14" pageNumber="477">Shimoji Island, Akuma-no-Yakata cave</collectingRegion>
:
</emphasis>
<location id="3B756017D364FF85FBFF3ADEFB07FF5C" LSID="urn:lsid:plazi:treatment:B60387DAD367FF85FF493C82FC63FE5F:3B756017D364FF85FBFF3ADEFB07FF5C" box="[1057,1155,152,175]" country="Japan" name="entrance" pageId="14" pageNumber="477" stateProvince="Shimoji Island, Akuma-no-Yakata cave">entrance</location>
,
<date id="4A14100CD364FF85FB463ADEFAFDFF43" box="[1176,1401,151,176]" pageId="14" pageNumber="477" value="2017-09-01">
<collectingDate id="5A50E9E4D364FF85FB463ADEFAFDFF43" box="[1176,1401,151,176]" pageId="14" pageNumber="477" value="2017-09-01">1 September 2017</collectingDate>
</date>
,
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(sl
<quantity id="F9529B29D364FF85FE5C3AFAFE59FF27" box="[386,477,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="477" unit="mm" value="3.7">3.7 mm</quantity>
), RUMF-ZC-6058 (SH104)
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.
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<paragraph id="3E1536CCD364FF85FF193A99FA19FF0B" blockId="14.[151,1437,151,429]" box="[199,1437,223,248]" pageId="14" pageNumber="477">
<emphasis id="0CDEEADED364FF85FF193A99FEE4FF0B" bold="true" box="[199,352,223,248]" pageId="14" pageNumber="477">Distribution.</emphasis>
Japan (Ryukyu and Izu Islands) and Australia;
<quantity id="F9529B29D364FF85FCAF3A99FC50FF04" box="[881,980,223,248]" metricMagnitude="1" metricUnit="m" metricValue="3.55" metricValueMax="6.1" metricValueMin="1.0" pageId="14" pageNumber="477" unit="m" value="35.5" valueMax="61.0" valueMin="10.0">10–61 m</quantity>
(
<bibRefCitation id="5A3B4B3DD364FF85FC3F3A99FB06FF0B" author="Asakura, A." box="[993,1154,223,248]" pageId="14" pageNumber="477" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD364FF85FB503AA6FA15FF0B" author="Osawa, M. &amp; Takeda, M." box="[1166,1425,223,248]" pageId="14" pageNumber="477" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
).
</paragraph>
<paragraph id="3E1536CCD364FF85FF193B42FA08FEEF" blockId="14.[151,1437,151,429]" box="[199,1420,260,285]" pageId="14" pageNumber="477">
<emphasis id="0CDEEADED364FF85FF193B42FEAEFEEE" bold="true" box="[199,298,260,285]" pageId="14" pageNumber="477">Habitat.</emphasis>
Coral reefs, often in rocky areas or submarine caves (
<bibRefCitation id="5A3B4B3DD364FF85FC583B42FBF5FEEF" author="Kato, S. &amp; Okuno, J." box="[902,1137,260,285]" pageId="14" pageNumber="477" refId="ref10558" refString="Kato, S. &amp; Okuno, J. (2001) Shrimps and crabs of Hachijo Island. TBS-Britannica Co. Ltd, Tokyo, 160 pp. [in Japanese]" type="book" year="2001">Kato &amp; Okuno 2001</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD364FF85FBA53B42FA04FEEF" author="Osawa, M. &amp; Takeda, M." box="[1147,1408,260,285]" pageId="14" pageNumber="477" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
).
</paragraph>
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<subSubSection id="76B06547D364FF85FF193B61FC63FE5F" pageId="14" pageNumber="477" type="discussion">
<paragraph id="3E1536CCD364FF85FF193B61FC63FE5F" blockId="14.[151,1437,151,429]" pageId="14" pageNumber="477">
<emphasis id="0CDEEADED364FF85FF193B61FEBFFEB3" bold="true" box="[199,315,295,320]" pageId="14" pageNumber="477">Remarks.</emphasis>
<emphasis id="0CDEEADED364FF85FE9C3B6EFDDCFECC" box="[322,600,296,320]" italics="true" pageId="14" pageNumber="477">
Pylopaguropsis
<taxonomicName id="F9AA4D4FD364FF85FE293B6EFDDCFECC" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[503,600,296,319]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="477" phylum="Arthropoda" rank="species" species="speciosa">speciosa</taxonomicName>
</emphasis>
has been often recorded from coral reefs including submarine caves in the Ryukyu Islands (
<bibRefCitation id="5A3B4B3DD364FF85FE843B0AFE7BFE97" author="Asakura, A." box="[346,511,332,357]" pageId="14" pageNumber="477" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD364FF85FDD33B0AFC9FFE97" author="Osawa, M. &amp; Takeda, M." box="[525,795,332,357]" pageId="14" pageNumber="477" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD364FF85FCF73B0AFBA3FE96" author="Osawa, M. &amp; Fujita, Y." box="[809,1063,332,357]" pageId="14" pageNumber="477" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita 2016</bibRefCitation>
;
<tableCitation id="73280377D364FF85FBE83B0AFB0BFE97" box="[1078,1167,332,357]" captionStart="TABLE 2" captionStartId="6.[147,233,596,618]" captionText="TABLE 2. Submarine caves surveyed and hermit crab species collected in three islands of the Ryukyu Islands, southwestern Japan." pageId="14" pageNumber="477">Table 2</tableCitation>
). The coloration of the species is very characteristic and readily distinguished from other congeners; the right chela is magenta and the left cheliped and ambulatory legs are bright orange with white stripes (
<figureCitation id="A6912A49D364FF85FC5D3BD2FC5FFE5F" box="[899,987,404,429]" captionStart="FIGURE 3" captionStartId="11.[151,250,1378,1400]" captionTargetBox="[151,1436,195,1356]" captionTargetId="figure@11.[151,1436,195,1356]" captionTargetPageId="11" captionText="FIGURE 3. Entire animal, dorsal view. A, Pylopaguropsis bellula Osawa &amp; Okuno, 2007, male (sl 4.1 mm), RUMF-ZC-6042 (IE17); B, Pylopaguropsis fimbriata McLaughlin &amp; Haig, 1989, ovigerous female (sl 4.4 mm), RUMF-ZC-6046 (IE179); C, Pylopaguropsis granulata Asakura, 2000, female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31); D, Pylopaguropsis keijii McLaughlin &amp; Haig, 1989, male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87); E, Pylopaguropsis speciosa McLaughlin &amp; Haig, 1989, ovigerous female (sl 3.7 mm), RUMF-ZC-6058 (SH104)." httpUri="https://zenodo.org/record/2627824/files/figure.png" pageId="14" pageNumber="477">Fig. 3E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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Family
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Diogenidae
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<emphasis id="0CDEEADED368FF89FF49397AFDB1FCA5" bold="true" box="[151,565,828,854]" pageId="2" pageNumber="465">
<emphasis id="0CDEEADED368FF89FF49397AFEDFFCA5" bold="true" box="[151,347,828,854]" italics="true" pageId="2" pageNumber="465">Aniculus retipes</emphasis>
<bibRefCitation id="5A3B4B3DD368FF89FEBC397AFDB1FCA5" author="Lewinsohn, C." box="[354,565,828,854]" pageId="2" pageNumber="465" pagination="76 - 82" refId="ref11235" refString="Lewinsohn, C. (1982) Aniculus retipes n. sp. ein neuer Einsiedlerkrebs aus dem Roten Meer (Decapoda, Anomura). Crustaceana, 42 (1 - 2), 76 - 82. https: // doi. org / 10.1163 / 156854082 X 00713" type="journal article" year="1982">Lewinsohn, 1982</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="76B06547D368FF89FF493918FBFDFC28" pageId="2" pageNumber="465" type="description">
<paragraph id="3E1536CCD368FF89FF493918FF7BFC84" blockId="2.[151,565,828,887]" box="[151,255,862,887]" pageId="2" pageNumber="465">
(
<figureCitation id="A6912A49D368FF89FF413918FF72FC84" box="[159,246,862,887]" captionStart="FIGURE 1" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@5.[151,1436,193,1705]" captionTargetPageId="5" captionText="FIGURE 1. Entire animal, dorsal view. A, Aniculus retipes Lewinsohn, 1982, male (sl 11.9 mm), RUMF-ZC-6006 (IE141); B, Paguristes jalur Morgan, 1992, male (sl 5.4 mm), RUMF-ZC-6012 (IE69); C, Pseudopaguristes shidarai Asakura, 2004, male (sl 2.6 mm), RUMF-ZC-6020 (HD2017-140); D, Catapaguroides foresti McLaughlin, 2002, male (sl 2.0 mm), RUMFZC-6023 (IE173); E, Catapaguroides iejimensis Osawa &amp; Takeda, 2004, male (sl 1.8 mm), RUMF-ZC-6026 (IE177); F, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, male (sl 2.1 mm), RUMF-ZC-6027 (IE60); G, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107)." httpUri="https://zenodo.org/record/2627820/files/figure.png" pageId="2" pageNumber="465">Fig. 1A</figureCitation>
)
</paragraph>
<paragraph id="3E1536CCD368FF89FF4939E0FBFDFC28" blockId="2.[151,1436,934,987]" pageId="2" pageNumber="465">
<emphasis id="0CDEEADED368FF89FF4939E0FEBDFC4E" box="[151,313,934,957]" italics="true" pageId="2" pageNumber="465">Aniculus retipes</emphasis>
<bibRefCitation id="5A3B4B3DD368FF89FE9E39E0FE76FC4F" author="Lewinsohn, C." box="[320,498,934,956]" pageId="2" pageNumber="465" pagination="76 - 82" refId="ref11235" refString="Lewinsohn, C. (1982) Aniculus retipes n. sp. ein neuer Einsiedlerkrebs aus dem Roten Meer (Decapoda, Anomura). Crustaceana, 42 (1 - 2), 76 - 82. https: // doi. org / 10.1163 / 156854082 X 00713" type="journal article" year="1982">Lewinsohn, 1982</bibRefCitation>
: 76, figs. 1, 2 (
<typeStatus id="E111886ED368FF89FD5639E0FD31FC4F" box="[648,693,934,956]" pageId="2" pageNumber="465">type</typeStatus>
locality: Mersa Bereika, Sinai Peninsula).—
<bibRefCitation id="5A3B4B3DD368FF89FBA939E0FB70FC4F" author="Forest, J." box="[1143,1268,934,956]" pageId="2" pageNumber="465" pagination="1 - 91" refId="ref9819" refString="Forest, J. (1984) Revision du genre Aniculus. Crustaceana, Supplement, 8: 1 - 91." type="journal article" year="1984">Forest 1984</bibRefCitation>
: 51, figs. 13, 20, 51–58.—McLaughlin
<emphasis id="0CDEEADED368FF89FE773980FE52FC28" box="[425,470,965,987]" italics="true" pageId="2" pageNumber="465">et al</emphasis>
. 2007: 103, 2 unnumbered figs.—
<bibRefCitation id="5A3B4B3DD368FF89FCEE3983FC6BFC28" author="Poupin, J. &amp; Bouchard, J. - M. &amp; Dinhut, V. &amp; Cleva, R. &amp; Dumas, J." box="[816,1007,965,987]" pageId="2" pageNumber="465" pagination="1 - 73" refId="ref13047" refString="Poupin, J., Bouchard, J. - M., Dinhut, V., Cleva, R. &amp; Dumas, J. (2013) Anomura (Crustacea Decapoda) from the Mayotte region, western Indian Ocean. Atoll Research Bulletin, 593, i-iv, 1 - 73. https: // doi. org / 10.5479 / si. 00775630.593" type="journal article" year="2013">
Poupin
<emphasis id="0CDEEADED368FF89FC5E3980FC29FC28" box="[896,941,965,987]" italics="true" pageId="2" pageNumber="465">et al</emphasis>
. 2013
</bibRefCitation>
: 26, fig. 13B.
</paragraph>
</subSubSection>
<subSubSection id="76B06547D368FF89FF493E41FF75FB0B" pageId="2" pageNumber="465" type="materials_examined">
<paragraph id="3E1536CCD368FF89FF493E41FF75FB0B" blockId="2.[151,1436,1031,1525]" pageId="2" pageNumber="465">
<emphasis id="0CDEEADED368FF89FF493E41FCCCFBD3" bold="true" box="[151,840,1031,1056]" pageId="2" pageNumber="465">
Material examined. Ie
<materialsCitation id="8EC23C91D368FF89FE723E41FCCCFBD3" ID-GBIF-Occurrence="2234219536" box="[428,840,1031,1056]" collectingDate="2017-06-23" country="Iceland" pageId="2" pageNumber="465" specimenCount="1">Island, unnamed submarine cave:</materialsCitation>
</emphasis>
entrance, roof,
<collectingDate id="5A50E9E4D368FF89FBDE3E4EFB19FBD3" box="[1024,1181,1032,1056]" pageId="2" pageNumber="465" value="2017-06-23">
<date id="4A14100CD368FF89FBDE3E4EFB19FBD3" box="[1024,1181,1032,1056]" pageId="2" pageNumber="465" value="2017-06-23">23 June 2017</date>
</collectingDate>
,
<materialsCitation id="8EC23C91D368FF89FB743E4EFA1CFBB6" ID-GBIF-Occurrence="2234219639" collectingDate="2016-08-10" collectionCode="IE" country="Iceland" elevation="5" location="entrance, 5 m from entrance, roof" municipality="Hedo Dome cave" pageId="2" pageNumber="465" specimenCode="RUMF-ZC-6006" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa Island, Hedo Dome cave">
<specimenCount id="28ACFD45D368FF89FB743E4EFB7CFBD3" box="[1194,1272,1031,1056]" pageId="2" pageNumber="465" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D368FF89FAF93E4EFA14FBD3" box="[1319,1424,1032,1056]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="2" pageNumber="465" unit="mm" value="11.9">11.9 mm</quantity>
),
<specimenCode id="6E0C9EB7D368FF89FF493E6AFED0FBB6" box="[151,340,1068,1093]" pageId="2" pageNumber="465">RUMF-ZC-6006</specimenCode>
(IE141).
<emphasis id="0CDEEADED368FF89FE613E6AFCDFFBB6" bold="true" box="[447,859,1068,1093]" pageId="2" pageNumber="465">
<collectingRegion id="FC6EF82ED368FF89FE613E6AFCD2FBB6" box="[447,854,1068,1093]" pageId="2" pageNumber="465">Okinawa Island, Hedo Dome cave</collectingRegion>
:
</emphasis>
<location id="3B756017D368FF89FCBC3E6BFB5CFBB6" LSID="urn:lsid:plazi:treatment:B60387DAD368FF89FF4938B0FABEFA07:3B756017D368FF89FCBC3E6BFB5CFBB6" box="[866,1240,1068,1093]" country="Iceland" municipality="Hedo Dome cave" name="entrance, 5 m from entrance, roof" pageId="2" pageNumber="465" stateProvince="Okinawa Island, Hedo Dome cave">
entrance,
<quantity id="F9529B29D368FF89FC103E6AFC7CFBB7" box="[974,1016,1068,1093]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="465" unit="m" value="5.0">5 m</quantity>
from entrance, roof
</location>
,
<collectingDate id="5A50E9E4D368FF89FB3D3E6AFA1CFBB6" box="[1251,1432,1068,1093]" pageId="2" pageNumber="465" value="2016-08-10">
<date id="4A14100CD368FF89FB3D3E6AFA1CFBB6" box="[1251,1432,1068,1093]" pageId="2" pageNumber="465" value="2016-08-10">10 August 2016</date>
</collectingDate>
</materialsCitation>
,
<specimenCount id="28ACFD45D368FF89FF493E16FF65FB9B" box="[151,225,1103,1128]" pageId="2" pageNumber="465" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D368FF89FED43E16FEE0FB9B" box="[266,356,1104,1128]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="465" unit="mm" value="9.1">9.1 mm</quantity>
), RUMF-ZC-6007 (
<collectionCode id="58BBAE09D368FF89FD9C3E09FDE9FB9B" box="[578,621,1103,1128]" pageId="2" pageNumber="465">HD</collectionCode>
<date id="4A14100CD368FF89FDB33E09FD4AFB9B" box="[621,718,1103,1128]" pageId="2" pageNumber="465" value="2016" valueMax="2047">2016-47</date>
); entrance,
<quantity id="F9529B29D368FF89FC933E09FCF3FB94" box="[845,887,1103,1128]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="465" unit="m" value="5.0">5 m</quantity>
from entrance, rocks,
<date id="4A14100CD368FF89FBAE3E16FAA1FB9B" box="[1136,1317,1103,1128]" pageId="2" pageNumber="465" value="2016-08-10">10 August 2016</date>
,
<specimenCount id="28ACFD45D368FF89FAF13E16FAFEFB9B" box="[1327,1402,1103,1128]" pageId="2" pageNumber="465" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D368FF89FF493E32FF72FB7F" box="[151,246,1140,1164]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="465" unit="mm" value="7.1">7.1 mm</quantity>
), RUMF-ZC-6008 (
<collectionCode id="58BBAE09D368FF89FE3E3E32FD8FFB7E" box="[480,523,1140,1165]" pageId="2" pageNumber="465">HD</collectionCode>
<date id="4A14100CD368FF89FDD53E32FDEFFB7E" box="[523,619,1140,1165]" pageId="2" pageNumber="465" value="2016" valueMax="2049">2016-49</date>
); main tube,
<date id="4A14100CD368FF89FCD23E32FC2AFB7F" box="[780,942,1140,1165]" pageId="2" pageNumber="465" value="2017-05-21">21 May 2017</date>
,
<materialsCitation id="8EC23C91D368FF89FC613E32FD2EFB43" ID-GBIF-Occurrence="2234219654" collectingDate="2017" collectingDateMax="2083" collectingDateMin="2017" collectionCode="IE" country="Iceland" elevation="5" location="entrance, 5 m from entrance, roof" municipality="Hedo Dome cave" pageId="2" pageNumber="465" specimenCode="RUMF-ZC-6009" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa Island, Hedo Dome cave">
<specimenCount id="28ACFD45D368FF89FC613E32FBA1FB7E" box="[959,1061,1140,1165]" pageId="2" pageNumber="465" type="female">1 female</specimenCount>
(sl 17.0 mm),
<specimenCode id="6E0C9EB7D368FF89FB003E32FA18FB7E" box="[1246,1436,1140,1165]" pageId="2" pageNumber="465">RUMF-ZC-6009</specimenCode>
(HD
<date id="4A14100CD368FF89FF153EDEFEAFFB43" box="[203,299,1176,1200]" pageId="2" pageNumber="465" value="2017" valueMax="2083">2017-83</date>
); same data as RUMF-ZC-6009
</materialsCitation>
,
<materialsCitation id="8EC23C91D368FF89FD653EDEFC3AFB27" ID-GBIF-Occurrence="2234219510" collectingDate="2017-02-06" collectingDateMax="2094" collectingDateMin="2017" country="Iceland" location="middle zone, roof" municipality="Akuma-no-Yakata cave" pageId="2" pageNumber="465" specimenCode="RUMF-ZC-6010" specimenCount="1" specimenCount-male="1" stateProvince="Shimoji Island, Akuma-no-Yakata cave">
<specimenCount id="28ACFD45D368FF89FD653EDEFC88FB43" box="[699,780,1175,1200]" pageId="2" pageNumber="465" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D368FF89FC9C3ED1FC34FB43" box="[834,944,1175,1200]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="2" pageNumber="465" unit="mm" value="13.6">13.6 mm</quantity>
),
<specimenCode id="6E0C9EB7D368FF89FC163ED1FB02FB43" box="[968,1158,1175,1200]" pageId="2" pageNumber="465">RUMF-ZC-6010</specimenCode>
(HD
<date id="4A14100CD368FF89FB183EDEFAA2FB43" box="[1222,1318,1176,1200]" pageId="2" pageNumber="465" value="2017" valueMax="2094">2017-94</date>
).
<emphasis id="0CDEEADED368FF89FAE03ED1FD92FB26" bold="true" pageId="2" pageNumber="465">
<collectingRegion id="FC6EF82ED368FF89FAE03ED1FD96FB26" pageId="2" pageNumber="465">Shimoji Island, Akuma-no-Yakata cave</collectingRegion>
:
</emphasis>
<location id="3B756017D368FF89FDFF3EFAFD74FB26" LSID="urn:lsid:plazi:treatment:B60387DAD368FF89FF4938B0FABEFA07:3B756017D368FF89FDFF3EFAFD74FB26" box="[545,752,1212,1237]" country="Iceland" municipality="Akuma-no-Yakata cave" name="middle zone, roof" pageId="2" pageNumber="465" stateProvince="Shimoji Island, Akuma-no-Yakata cave">middle zone, roof</location>
,
<collectingDate id="5A50E9E4D368FF89FD233EFAFC3AFB27" box="[765,958,1212,1237]" pageId="2" pageNumber="465" value="2017-02-06">
<date id="4A14100CD368FF89FD233EFAFC3AFB27" box="[765,958,1212,1237]" pageId="2" pageNumber="465" value="2017-02-06">6 February 2017</date>
</collectingDate>
</materialsCitation>
,
<specimenCount id="28ACFD45D368FF89FC123EFAFBB4FB26" box="[972,1072,1212,1237]" pageId="2" pageNumber="465" type="female">1 female</specimenCount>
(sl
<quantity id="F9529B29D368FF89FB813EFAFB4EFB27" box="[1119,1226,1212,1236]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="465" unit="mm" value="12.7">12.7 mm</quantity>
), RUMF-ZC-6011 (
<collectionCode id="58BBAE09D368FF89FF413E99FF42FB0B" box="[159,198,1247,1272]" pageId="2" pageNumber="465">SH</collectionCode>
55).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D368FF89FF193F42FC7DFA7B" pageId="2" pageNumber="465" type="distribution">
<paragraph id="3E1536CCD368FF89FF193F42FA18FA96" blockId="2.[151,1436,1031,1525]" pageId="2" pageNumber="465">
<emphasis id="0CDEEADED368FF89FF193F42FEE4FAEE" bold="true" box="[199,352,1284,1309]" pageId="2" pageNumber="465">Distribution.</emphasis>
Wide ranging in the Indo-West Pacific: Red Sea, eastern Africa (
<collectingCountry id="46BD765CD368FF89FB873F42FB40FAEE" box="[1113,1220,1284,1309]" name="Tanzania" pageId="2" pageNumber="465">Tanzania</collectingCountry>
), Glorioso Islands,
<collectingCountry id="46BD765CD368FF89FF493F61FED2FAB3" box="[151,342,1319,1344]" name="Christmas Island" pageId="2" pageNumber="465">Christmas Island</collectingCountry>
,
<collectingCountry id="46BD765CD368FF89FEBF3F6EFDF5FAB3" box="[353,625,1319,1344]" name="Cocos (Keeling) Islands" pageId="2" pageNumber="465">Cocos (Keeling) Islands</collectingCountry>
,
<collectingCountry id="46BD765CD368FF89FDA23F61FD68FAB3" box="[636,748,1319,1344]" name="Indonesia" pageId="2" pageNumber="465">Indonesia</collectingCountry>
,
<collectingCountry id="46BD765CD368FF89FD293F61FCDDFAB3" box="[759,857,1319,1344]" name="Vietnam" pageId="2" pageNumber="465">Vietnam</collectingCountry>
,
<collectingCountry id="46BD765CD368FF89FCBD3F61FC3CFAB3" box="[867,952,1319,1344]" name="Taiwan" pageId="2" pageNumber="465">Taiwan</collectingCountry>
,
<collectingCountry id="46BD765CD368FF89FC1D3F6EFB82FACC" box="[963,1030,1320,1343]" name="Japan" pageId="2" pageNumber="465">Japan</collectingCountry>
, eastern
<collectingCountry id="46BD765CD368FF89FBB83F61FB4BFAB3" box="[1126,1231,1319,1344]" name="Australia" pageId="2" pageNumber="465">Australia</collectingCountry>
,
<collectingCountry id="46BD765CD368FF89FB043F61FA1CFAB3" box="[1242,1432,1319,1344]" name="Solomon Islands" pageId="2" pageNumber="465">Solomon Islands</collectingCountry>
,
<collectingCountry id="46BD765CD368FF89FF493F0AFF65FA96" box="[151,225,1356,1381]" name="United Kingdom" pageId="2" pageNumber="465">Wallis</collectingCountry>
,
<collectingCountry id="46BD765CD368FF89FF323F0BFEB9FA97" box="[236,317,1357,1380]" name="Wallis and Futuna" pageId="2" pageNumber="465">Futuna</collectingCountry>
,
<collectingCountry id="46BD765CD368FF89FE993F0AFE12FA97" box="[327,406,1356,1380]" name="Samoa" pageId="2" pageNumber="465">Samoa</collectingCountry>
, and
<collectingCountry id="46BD765CD368FF89FE0E3F0AFD11FA96" box="[464,661,1356,1381]" name="French Polynesia" pageId="2" pageNumber="465">French Polynesia</collectingCountry>
;
<quantity id="F9529B29D368FF89FD7F3F0AFD72FA97" box="[673,758,1356,1381]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="3.5" metricValueMin="0.0" pageId="2" pageNumber="465" unit="m" value="17.5" valueMax="35.0" valueMin="0.0">0–35 m</quantity>
(McLaughlin
<emphasis id="0CDEEADED368FF89FC483F0BFC4CFA97" box="[918,968,1356,1380]" italics="true" pageId="2" pageNumber="465">et al</emphasis>
. 2007;
<bibRefCitation id="5A3B4B3DD368FF89FBC23F0AFB6FFA97" author="Poupin, J. &amp; Bouchard, J. - M. &amp; Dinhut, V. &amp; Cleva, R. &amp; Dumas, J." box="[1052,1259,1356,1381]" pageId="2" pageNumber="465" pagination="1 - 73" refId="ref13047" refString="Poupin, J., Bouchard, J. - M., Dinhut, V., Cleva, R. &amp; Dumas, J. (2013) Anomura (Crustacea Decapoda) from the Mayotte region, western Indian Ocean. Atoll Research Bulletin, 593, i-iv, 1 - 73. https: // doi. org / 10.5479 / si. 00775630.593" type="journal article" year="2013">
Poupin
<emphasis id="0CDEEADED368FF89FBAC3F0BFB26FA97" box="[1138,1186,1356,1380]" italics="true" pageId="2" pageNumber="465">et al</emphasis>
. 2013
</bibRefCitation>
; present study).
</paragraph>
<paragraph id="3E1536CCD368FF89FF193F29FC7DFA7B" blockId="2.[151,1436,1031,1525]" box="[199,1017,1391,1416]" pageId="2" pageNumber="465">
<emphasis id="0CDEEADED368FF89FF193F29FEAEFA7B" bold="true" box="[199,298,1391,1416]" pageId="2" pageNumber="465">Habitat.</emphasis>
Coral and rocky reefs (
<bibRefCitation id="5A3B4B3DD368FF89FDEF3F36FC8BFA7B" author="Nomura, K. &amp; Miyake, S. &amp; Kamezaki, N." box="[561,783,1391,1416]" pageId="2" pageNumber="465" pagination="2 - 7" refId="ref12411" refString="Nomura, K., Miyake, S. &amp; Kamezaki, N. (1996) Revision of the genus Aniculus (Anomura, Diogenidae) in Japan. I. O. P. Diving News, 8, 2 - 7. [in Japanese with English abstract]" type="journal article" year="1996">
Nomura
<emphasis id="0CDEEADED368FF89FD4B3F37FD43FA7B" box="[661,711,1392,1416]" italics="true" pageId="2" pageNumber="465">et al</emphasis>
. 1996
</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD368FF89FCC53F29FC6AFA7B" author="Poupin, J. &amp; Bouchard, J. - M. &amp; Dinhut, V. &amp; Cleva, R. &amp; Dumas, J." box="[795,1006,1391,1416]" pageId="2" pageNumber="465" pagination="1 - 73" refId="ref13047" refString="Poupin, J., Bouchard, J. - M., Dinhut, V., Cleva, R. &amp; Dumas, J. (2013) Anomura (Crustacea Decapoda) from the Mayotte region, western Indian Ocean. Atoll Research Bulletin, 593, i-iv, 1 - 73. https: // doi. org / 10.5479 / si. 00775630.593" type="journal article" year="2013">
Poupin
<emphasis id="0CDEEADED368FF89FCAC3F37FC26FA7B" box="[882,930,1392,1416]" italics="true" pageId="2" pageNumber="465">et al</emphasis>
. 2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D368FF89FF193FD2FABEFA07" pageId="2" pageNumber="465" type="discussion">
<paragraph id="3E1536CCD368FF89FF193FD2FABEFA07" blockId="2.[151,1436,1031,1525]" pageId="2" pageNumber="465">
<emphasis id="0CDEEADED368FF89FF193FD2FEBFFA5E" bold="true" box="[199,315,1428,1453]" pageId="2" pageNumber="465">Remarks.</emphasis>
Two species of the genus
<emphasis id="0CDEEADED368FF89FDBB3FD2FD43FA5F" box="[613,711,1428,1452]" italics="true" pageId="2" pageNumber="465">Aniculus</emphasis>
<bibRefCitation id="5A3B4B3DD368FF89FD113FD3FCD7FA5E" author="Dana, J. D." box="[719,851,1428,1453]" pageId="2" pageNumber="465" pagination="6 - 28" refId="ref9666" refString="Dana, J. D. (1852) Conspectus crustaceorum, etc., Conspectus of the Crustacea of the Exploring Expedition under Capt. Wilkes, U. S. N., including the Paguridea, continued, the Megalopidea, and the Macroura. Paguridea, continued, and subtribe Megalopidea. (Preprint from) Proceedings of the Academy of Natural Sciences, Philadelphia, 6, 6 - 28." type="journal article" year="1852">Dana, 1852</bibRefCitation>
,
<emphasis id="0CDEEADED368FF89FC813FD2FC66FA5F" box="[863,994,1428,1452]" italics="true" pageId="2" pageNumber="465">A. maximus</emphasis>
<bibRefCitation id="5A3B4B3DD368FF89FC373FD2FB32FA5E" author="Edmondson, C. H." box="[1001,1206,1428,1453]" pageId="2" pageNumber="465" pagination="67 - 86" refId="ref9742" refString="Edmondson, C. H. (1952) Additional central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 21 (6), 67 - 86." type="journal article" year="1952">Edmondson, 1952</bibRefCitation>
and
<emphasis id="0CDEEADED368FF89FB333FD2FADCFA5F" box="[1261,1368,1428,1452]" italics="true" pageId="2" pageNumber="465">A. retipes</emphasis>
, have been recorded from submarine caves in the Ryukyu Islands (
<bibRefCitation id="5A3B4B3DD368FF89FC9A3FF1FBCBFA23" author="Rahayu, D. L. &amp; Osawa, M." box="[836,1103,1463,1488]" pageId="2" pageNumber="465" pagination="176 - 190" refId="ref13115" refString="Rahayu, D. L. &amp; Osawa, M. (2012) Hermit crabs (Crustacea: Decapoda: Diogenidae) from the KUMEJIMA 2009 Expedition, Japan. Zootaxa, 3367, 176 - 190." type="journal article" year="2012">Rahayu &amp; Osawa 2012</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD368FF89FB823FFEFAD7FA23" author="Osawa, M. &amp; Fujita, Y." box="[1116,1363,1463,1488]" pageId="2" pageNumber="465" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita 2016</bibRefCitation>
;
<tableCitation id="73280377D368FF89FA803FF1FF2CFA07" captionStart="TABLE 2" captionStartId="6.[147,233,596,618]" captionText="TABLE 2. Submarine caves surveyed and hermit crab species collected in three islands of the Ryukyu Islands, southwestern Japan." pageId="2" pageNumber="465">Table 2</tableCitation>
). In general,
<emphasis id="0CDEEADED368FF89FEE53F9AFE21FA07" box="[315,421,1500,1524]" italics="true" pageId="2" pageNumber="465">A. retipes</emphasis>
is more commonly found in cryptic environments of coral reefs than
<emphasis id="0CDEEADED368FF89FB6E3F9AFAB7FA07" box="[1200,1331,1500,1524]" italics="true" pageId="2" pageNumber="465">A. maximus</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

545
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<taxonomicName id="F9AA4D4FD368FF89FF493C01FD8AF991" ID-CoL="cc706566-9604-402c-9fd5-534ca11bb9d6" authority="Morgan, 1992" authorityName="Morgan" authorityYear="1992" box="[151,526,1607,1634]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="jalur">
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<emphasis id="0CDEEADED368FF89FF493C01FEDCF992" bold="true" box="[151,344,1607,1633]" italics="true" pageId="2" pageNumber="465">Paguristes jalur</emphasis>
<bibRefCitation id="5A3B4B3DD368FF89FE813C0EFD8AF991" author="Morgan, G. J." box="[351,526,1608,1634]" pageId="2" pageNumber="465" pagination="163 - 174" refId="ref12214" refString="Morgan, G. J. (1992) The hermit crabs (Crustacea: Decapoda: Coenobitidae, Diogenidae, Paguridae) of Christmas and Cocos (Keeling) Islands, Indian Ocean, with description of a new species of Paguristes. Raffles Bulletin of Zoology, 49 (2), 163 - 174." type="journal article" year="1992">Morgan, 1992</bibRefCitation>
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<paragraph id="3E1536CCD368FF89FF493C2CFF7AF970" blockId="2.[151,526,1607,1667]" box="[151,254,1642,1667]" pageId="2" pageNumber="465">
(
<figureCitation id="A6912A49D368FF89FF413C2CFF70F970" box="[159,244,1642,1667]" captionStart="FIGURE 1" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@5.[151,1436,193,1705]" captionTargetPageId="5" captionText="FIGURE 1. Entire animal, dorsal view. A, Aniculus retipes Lewinsohn, 1982, male (sl 11.9 mm), RUMF-ZC-6006 (IE141); B, Paguristes jalur Morgan, 1992, male (sl 5.4 mm), RUMF-ZC-6012 (IE69); C, Pseudopaguristes shidarai Asakura, 2004, male (sl 2.6 mm), RUMF-ZC-6020 (HD2017-140); D, Catapaguroides foresti McLaughlin, 2002, male (sl 2.0 mm), RUMFZC-6023 (IE173); E, Catapaguroides iejimensis Osawa &amp; Takeda, 2004, male (sl 1.8 mm), RUMF-ZC-6026 (IE177); F, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, male (sl 2.1 mm), RUMF-ZC-6027 (IE60); G, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107)." httpUri="https://zenodo.org/record/2627820/files/figure.png" pageId="2" pageNumber="465">Fig. 1B</figureCitation>
)
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<emphasis id="0CDEEADED368FF89FF493CF5FEB8F93B" box="[151,316,1714,1736]" italics="true" pageId="2" pageNumber="465">Paguristes jalur</emphasis>
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: 167
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, figs. 1, 2 (
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locality: Horburgh Island,
<collectingCountry id="46BD765CD368FF89FC143CF4FB47F93B" box="[970,1219,1714,1736]" name="Cocos (Keeling) Islands" pageId="2" pageNumber="465">Cocos (Keeling) Islands</collectingCountry>
).—
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<bibRefCitation id="5A3B4B3DD368FF89FB373CF4FF79F914" author="Osawa, M. &amp; Takeda, M." pageId="2" pageNumber="465" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
: 1099
</treatmentCitation>
, figs. 1–4.—McLaughlin 2007: 216, fig. 11.
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</subSubSection>
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<paragraph id="3E1536CCD368FF89FF193D52FA18F887" blockId="2.[151,1436,1812,1909]" pageId="2" pageNumber="465">
<emphasis id="0CDEEADED368FF89FF193D52FCEAF8DE" bold="true" box="[199,878,1812,1837]" pageId="2" pageNumber="465">
Material examined. Ie
<materialsCitation id="8EC23C91D368FF89FE063D52FCEAF8DE" ID-GBIF-Occurrence="2234219652" box="[472,878,1812,1837]" collectingDate="2016" collectingDateMax="2069" collectingDateMin="2016" collectionCode="IE" country="Iceland" municipality="Hedo Dome cave" pageId="2" pageNumber="465" specimenCode="IE69" specimenCount="1" specimenCount-male="5">Island, unnamed submarine cave:</materialsCitation>
</emphasis>
main tube, near bifurcation of tube,
<collectingDate id="5A50E9E4D368FF89FAD03D52FF56F8A3" pageId="2" pageNumber="465" value="2016-09-09">
<date id="4A14100CD368FF89FAD03D52FF56F8A3" pageId="2" pageNumber="465" value="2016-09-09">9 September 2016</date>
</collectingDate>
,
<specimenCount id="28ACFD45D368FF89FF013D7EFEA9F8A3" box="[223,301,1847,1872]" pageId="2" pageNumber="465" type="male">1 male</specimenCount>
(sl
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), RUMF-ZC-6012 (
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<collectionCode id="58BBAE09D368FF89FD413D71FD39F8A3" box="[671,701,1847,1872]" pageId="2" pageNumber="465">IE</collectionCode>
69
</specimenCode>
).
<emphasis id="0CDEEADED368FF89FD2E3D71FB1DF8A3" bold="true" box="[752,1177,1847,1872]" pageId="2" pageNumber="465">
<location id="3B756017D368FF89FD2E3D71FC37F8A3" LSID="urn:lsid:plazi:treatment:B60387DAD368FF8FFF493C01FCAFFD76:3B756017D368FF89FD2E3D71FC37F8A3" box="[752,947,1847,1872]" country="Iceland" municipality="Hedo Dome cave" name="Okinawa Island" pageId="2" pageNumber="465">Okinawa Island</location>
,
<collectingMunicipality id="DE71ACB6D368FF89FC1C3D71FB10F8A3" box="[962,1172,1847,1872]" pageId="2" pageNumber="465">Hedo Dome cave</collectingMunicipality>
:
</emphasis>
main tube,
<collectingDate id="5A50E9E4D368FF89FAFB3D7EFF56F886" pageId="2" pageNumber="465" value="2016-08-12">
<date id="4A14100CD368FF89FAFB3D7EFF56F886" pageId="2" pageNumber="465" value="2016-08-12">12 August 2016</date>
</collectingDate>
,
<specimenCount id="28ACFD45D368FF89FF023D1AFEA3F886" box="[220,295,1884,1909]" pageId="2" pageNumber="465" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D368FF89FE8F3D1AFE2EF887" box="[337,426,1884,1909]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="465" unit="mm" value="5.5">5.5 mm</quantity>
), RUMF-ZC-6015 (
<collectionCode id="58BBAE09D368FF89FD573D1AFD30F886" box="[649,692,1884,1909]" pageId="2" pageNumber="465">HD</collectionCode>
<collectingDate id="5A50E9E4D368FF89FD6A3D1AFC91F886" box="[692,789,1884,1909]" pageId="2" pageNumber="465" value="2016" valueMax="2069">
<date id="4A14100CD368FF89FD6A3D1AFC91F886" box="[692,789,1884,1909]" pageId="2" pageNumber="465" value="2016" valueMax="2069">2016-69</date>
</collectingDate>
); same data as RUMF-ZC-6015,
<specimenCount id="28ACFD45D368FF89FB593D1AFB58F886" box="[1159,1244,1884,1909]" pageId="2" pageNumber="465" type="male">3 males</specimenCount>
(sl
<quantity id="F9529B29D368FF89FAD83D1AFA14F887" box="[1286,1424,1884,1909]" metricMagnitude="-3" metricUnit="m" metricValue="5.449999999999999" metricValueMax="6.3" metricValueMin="4.6" pageId="2" pageNumber="465" unit="mm" value="5.449999999999999" valueMax="6.3" valueMin="4.6">4.6–6.3 mm</quantity>
),
</paragraph>
<caption id="6AD56644D369FF88FF4D3847FE70FDB0" pageId="3" pageNumber="466">
<paragraph id="3E1536CCD369FF88FF4D3847FE70FDB0" blockId="3.[147,1911,513,579]" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3847FE86FDE4" bold="true" box="[147,258,513,535]" pageId="3" pageNumber="466">TABLE 1.</emphasis>
Hermit crab species reported from submarine caves in the Ryukyu Islands, southwestern Japan. Collection data of specimens reported by Rahayu &amp; Osawa (2012: table 1) is given by Naruse (2012).
</paragraph>
</caption>
<paragraph id="3E1536CCD369FF88FF4D382FF9A0F946" pageId="3" pageNumber="466">
<table id="4CAAC46CD3690074FF4D381CF8E6F946" box="[147,1890,602,1717]" gridcols="3" gridrows="29" pageId="3" pageNumber="466">
<tr id="809A348ED3690074FF4D381CF8E6FD7D" box="[147,1890,602,654]" gridrow="0" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D381CFD7AFD7D" box="[147,766,602,654]" gridcol="0" gridrow="0" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D382FFF65FD8C" bold="true" box="[147,225,617,639]" pageId="3" pageNumber="466">Species</emphasis>
</th>
<th id="C34B5DF2D3690074FCA8381CFA86FD7D" box="[886,1282,602,654]" gridcol="1" gridrow="0" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FCA8381CFBEAFD7D" bold="true" pageId="3" pageNumber="466">Locality (based on collection records from submarine caves)</emphasis>
</th>
<th id="C34B5DF2D3690074FAE4381CF8E6FD7D" box="[1338,1890,602,654]" gridcol="2" gridrow="0" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FAE4382FFA22FD8C" bold="true" box="[1338,1446,617,639]" pageId="3" pageNumber="466">Reference</emphasis>
</th>
</tr>
<tr id="809A348ED3690074FF4D38DFF8E6FD5C" box="[147,1890,665,687]" gridrow="1" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D38DFF8E6FD5C" box="[147,1890,665,687]" colspan="3" colspanRight="2" gridcol="0" gridrow="1" pageId="3" pageNumber="466">
<taxonomicName id="F9AA4D4FD369FF88FF4D38DFFE88FD5C" authorityName="Ortmann" authorityYear="1892" box="[147,268,665,687]" class="Malacostraca" family="Diogenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="family">
<emphasis id="0CDEEADED369FF88FF4D38DFFE88FD5C" bold="true" box="[147,268,665,687]" pageId="3" pageNumber="466">Diogenidae</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="809A348ED3690074FF4D38FBF8E6FD20" box="[147,1890,701,723]" gridrow="2" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D38FBFD7AFD20" box="[147,766,701,723]" gridcol="0" gridrow="2" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D38FBFEC8FD20" box="[147,332,701,723]" italics="true" pageId="3" pageNumber="466">Aniculus maximus</emphasis>
Edmondson, 1952
</th>
<td id="C34B5DF2D3690074FCA838FBFA86FD20" box="[886,1282,701,723]" gridcol="1" gridrow="2" pageId="3" pageNumber="466">Shimoji Island</td>
<td id="C34B5DF2D3690074FAE438FBF8E6FD20" box="[1338,1890,701,723]" gridcol="2" gridrow="2" pageId="3" pageNumber="466">Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D38A7F8E6FD04" box="[147,1890,737,759]" gridrow="3" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D38A7FD7AFD04" box="[147,766,737,759]" gridcol="0" gridrow="3" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D38A7FEB3FD04" box="[147,311,737,759]" italics="true" pageId="3" pageNumber="466">Aniculus retipes</emphasis>
Lewinsohn, 1982
</th>
<td id="C34B5DF2D3690074FCA838A7FA86FD04" box="[886,1282,737,759]" gridcol="1" gridrow="3" pageId="3" pageNumber="466">Kume Island, Shimoji Island</td>
<td id="C34B5DF2D3690074FAE438A7F8E6FD04" box="[1338,1890,737,759]" gridcol="2" gridrow="3" pageId="3" pageNumber="466">Rahayu &amp; Osawa (2012), Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D3943F8E6FCE8" box="[147,1890,773,795]" gridrow="4" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3943FD7AFCE8" box="[147,766,773,795]" gridcol="0" gridrow="4" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3943FED3FCE8" box="[147,343,773,795]" italics="true" pageId="3" pageNumber="466">Calcinus gaimardii</emphasis>
(H. Milne Edwards, 1848)
</th>
<td id="C34B5DF2D3690074FCA83943FA86FCE8" box="[886,1282,773,795]" gridcol="1" gridrow="4" pageId="3" pageNumber="466">Kume Island</td>
<td id="C34B5DF2D3690074FAE43943F8E6FCE8" box="[1338,1890,773,795]" gridcol="2" gridrow="4" pageId="3" pageNumber="466">Rahayu &amp; Osawa (2012)</td>
</tr>
<tr id="809A348ED3690074FF4D396FF8E6FCCC" box="[147,1890,809,831]" gridrow="5" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D396FFD7AFCCC" box="[147,766,809,831]" gridcol="0" gridrow="5" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D396FFEDAFCCC" box="[147,350,809,831]" italics="true" pageId="3" pageNumber="466">Calcinus guamensis</emphasis>
Wooster, 1984
</th>
<td id="C34B5DF2D3690074FCA8396FFA86FCCC" box="[886,1282,809,831]" gridcol="1" gridrow="5" pageId="3" pageNumber="466">Kume Island</td>
<td id="C34B5DF2D3690074FAE4396FF8E6FCCC" box="[1338,1890,809,831]" gridcol="2" gridrow="5" pageId="3" pageNumber="466">Rahayu &amp; Osawa (2012)</td>
</tr>
<tr id="809A348ED3690074FF4D390BF8E6FC90" box="[147,1890,845,867]" gridrow="6" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D390BFD7AFC90" box="[147,766,845,867]" gridcol="0" gridrow="6" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D390BFE02FC90" box="[147,390,845,867]" italics="true" pageId="3" pageNumber="466">Calcinus lineapropodus</emphasis>
Morgan &amp; Forest, 1991
</th>
<td id="C34B5DF2D3690074FCA8390BFA86FC90" box="[886,1282,845,867]" gridcol="1" gridrow="6" pageId="3" pageNumber="466">Kume Island</td>
<td id="C34B5DF2D3690074FAE4390BF8E6FC90" box="[1338,1890,845,867]" gridcol="2" gridrow="6" pageId="3" pageNumber="466">Rahayu &amp; Osawa (2012)</td>
</tr>
<tr id="809A348ED3690074FF4D3937F8E6FC74" box="[147,1890,881,903]" gridrow="7" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3937FD7AFC74" box="[147,766,881,903]" gridcol="0" gridrow="7" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3937FEC7FC74" box="[147,323,881,903]" italics="true" pageId="3" pageNumber="466">Calcinus vachoni</emphasis>
Forest, 1958
</th>
<td id="C34B5DF2D3690074FCA83937FA86FC74" box="[886,1282,881,903]" gridcol="1" gridrow="7" pageId="3" pageNumber="466">Kume Island</td>
<td id="C34B5DF2D3690074FAE43937F8E6FC74" box="[1338,1890,881,903]" gridcol="2" gridrow="7" pageId="3" pageNumber="466">Rahayu &amp; Osawa (2012)</td>
</tr>
<tr id="809A348ED3690074FF4D39D3F8E6FC58" box="[147,1890,917,939]" gridrow="8" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D39D3FD7AFC58" box="[147,766,917,939]" gridcol="0" gridrow="8" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D39D3FEFCFC58" box="[147,376,917,939]" italics="true" pageId="3" pageNumber="466">Ciliopagurus strigatus</emphasis>
(Herbst, 1804)
</th>
<td id="C34B5DF2D3690074FCA839D3FA86FC58" box="[886,1282,917,939]" gridcol="1" gridrow="8" pageId="3" pageNumber="466">Kume Island</td>
<td id="C34B5DF2D3690074FAE439D3F8E6FC58" box="[1338,1890,917,939]" gridcol="2" gridrow="8" pageId="3" pageNumber="466">Rahayu &amp; Osawa (2012)</td>
</tr>
<tr id="809A348ED3690074FF4D39FFF8E6FC3C" box="[147,1890,953,975]" gridrow="9" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D39FFFD7AFC3C" box="[147,766,953,975]" gridcol="0" gridrow="9" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D39FFFED4FC3C" box="[147,336,953,975]" italics="true" pageId="3" pageNumber="466">Dardanus guttatus</emphasis>
(Olivier, 1812)
</th>
<td id="C34B5DF2D3690074FCA839FFFA86FC3C" box="[886,1282,953,975]" gridcol="1" gridrow="9" pageId="3" pageNumber="466">Kume Island</td>
<td id="C34B5DF2D3690074FAE439FFF8E6FC3C" box="[1338,1890,953,975]" gridcol="2" gridrow="9" pageId="3" pageNumber="466">Rahayu &amp; Osawa (2012)</td>
</tr>
<tr id="809A348ED3690074FF4D399BF8E6FC00" box="[147,1890,989,1011]" gridrow="10" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D399BFD7AFC00" box="[147,766,989,1011]" gridcol="0" gridrow="10" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D399BFEE2FC00" box="[147,358,989,1011]" italics="true" pageId="3" pageNumber="466">Dardanus lagopodes</emphasis>
(Forskål, 1775)
</th>
<td id="C34B5DF2D3690074FCA8399BFA86FC00" box="[886,1282,989,1011]" gridcol="1" gridrow="10" pageId="3" pageNumber="466">Kume Island, Shimoji Island</td>
<td id="C34B5DF2D3690074FAE4399BF8E6FC00" box="[1338,1890,989,1011]" gridcol="2" gridrow="10" pageId="3" pageNumber="466">Rahayu &amp; Osawa (2012), Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D3E47F8E6FBE4" box="[147,1890,1025,1047]" gridrow="11" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3E47FD7AFBE4" box="[147,766,1025,1047]" gridcol="0" gridrow="11" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3E47FED3FBE4" box="[147,343,1025,1047]" italics="true" pageId="3" pageNumber="466">Dardanus megistos</emphasis>
(Herbst, 1804)
</th>
<td id="C34B5DF2D3690074FCA83E47FA86FBE4" box="[886,1282,1025,1047]" gridcol="1" gridrow="11" pageId="3" pageNumber="466">Kume Island</td>
<td id="C34B5DF2D3690074FAE43E47F8E6FBE4" box="[1338,1890,1025,1047]" gridcol="2" gridrow="11" pageId="3" pageNumber="466">Rahayu &amp; Osawa (2012)</td>
</tr>
<tr id="809A348ED3690074FF4D3E63F8E6FBC8" box="[147,1890,1061,1083]" gridrow="12" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3E63FD7AFBC8" box="[147,766,1061,1083]" gridcol="0" gridrow="12" pageId="3" pageNumber="466">
<taxonomicName id="F9AA4D4FD369FF88FF4D3E63FE49FBC8" authority="Morgan, 1992" authorityName="Morgan" authorityYear="1992" box="[147,461,1061,1083]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="jalur">
<emphasis id="0CDEEADED369FF88FF4D3E63FEB2FBC8" box="[147,310,1061,1083]" italics="true" pageId="3" pageNumber="466">Paguristes jalur</emphasis>
Morgan, 1992
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83E63FA86FBC8" box="[886,1282,1061,1083]" gridcol="1" gridrow="12" pageId="3" pageNumber="466">Shimoji Island</td>
<td id="C34B5DF2D3690074FAE43E63F8E6FBC8" box="[1338,1890,1061,1083]" gridcol="2" gridrow="12" pageId="3" pageNumber="466">Osawa &amp; Takeda (2004), Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D3E0FF8E6FBAC" box="[147,1890,1097,1119]" gridrow="13" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3E0FFD7AFBAC" box="[147,766,1097,1119]" gridcol="0" gridrow="13" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3E0FFE27FBAC" box="[147,419,1097,1119]" italics="true" pageId="3" pageNumber="466">Pseudopaguristes janetkae</emphasis>
McLaughlin, 2002a
</th>
<td id="C34B5DF2D3690074FCA83E0FFA86FBAC" box="[886,1282,1097,1119]" gridcol="1" gridrow="13" pageId="3" pageNumber="466">Shimoji Island</td>
<td id="C34B5DF2D3690074FAE43E0FF8E6FBAC" box="[1338,1890,1097,1119]" gridcol="2" gridrow="13" pageId="3" pageNumber="466">Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D3E2BF8E6FB70" box="[147,1890,1133,1155]" gridrow="14" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3E2BFD7AFB70" box="[147,766,1133,1155]" gridcol="0" gridrow="14" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3E2BFEC3FB70" box="[147,327,1133,1155]" italics="true" pageId="3" pageNumber="466">Pseudopaguristes</emphasis>
<taxonomicName id="F9AA4D4FD369FF88FE933E2BFDB8FB70" authority="Asakura, 2004" authorityName="Asakura" authorityYear="2004" box="[333,572,1133,1155]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="shidarai">
<emphasis id="0CDEEADED369FF88FE933E2BFE25FB70" box="[333,417,1133,1155]" italics="true" pageId="3" pageNumber="466">shidarai</emphasis>
Asakura, 2004
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83E2BFA86FB70" box="[886,1282,1133,1155]" gridcol="1" gridrow="14" pageId="3" pageNumber="466">Shimoji Island</td>
<td id="C34B5DF2D3690074FAE43E2BF8E6FB70" box="[1338,1890,1133,1155]" gridcol="2" gridrow="14" pageId="3" pageNumber="466">Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D3EF3F8E6FB38" box="[147,1890,1205,1227]" gridrow="15" pageId="3" pageNumber="466" rowspan-1="1" rowspan-2="1">
<th id="C34B5DF2D3690074FF4D3EF3FD7AFB38" box="[147,766,1205,1227]" gridcol="0" gridrow="15" pageId="3" pageNumber="466">
<taxonomicName id="F9AA4D4FD369FF88FF4D3EF3FE86FB38" authorityName="Latreille" authorityYear="1802" box="[147,258,1205,1227]" class="Malacostraca" family="Paguridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="family">
<emphasis id="0CDEEADED369FF88FF4D3EF3FE86FB38" bold="true" box="[147,258,1205,1227]" pageId="3" pageNumber="466">Paguridae</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="809A348ED3690074FF4D3E93F8E6FAFB" box="[147,1890,1237,1288]" gridrow="16" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3E93FD7AFAFB" box="[147,766,1237,1288]" gridcol="0" gridrow="16" pageId="3" pageNumber="466">
<taxonomicName id="F9AA4D4FD369FF88FF4D3EA2FDC9FB09" authority="McLaughlin, 2002 b" authorityName="McLaughlin" authorityYear="2002" box="[147,589,1252,1274]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="0CDEEADED369FF88FF4D3EA2FEF9FB09" box="[147,381,1252,1274]" italics="true" pageId="3" pageNumber="466">Catapaguroides foresti</emphasis>
McLaughlin, 2002b
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83E93FA86FAFB" box="[886,1282,1237,1288]" gridcol="1" gridrow="16" pageId="3" pageNumber="466">Ie Island, Okinawa Island</td>
<td id="C34B5DF2D3690074FAE43E93F8E6FAFB" box="[1338,1890,1237,1288]" gridcol="2" gridrow="16" pageId="3" pageNumber="466">
Osawa &amp; Takeda (2004, as
<taxonomicName id="F9AA4D4FD369FF88F98A3E93FAF4FAFB" authority="Osawa &amp; Takeda, 2004" authorityName="Osawa &amp; Takeda" authorityYear="2004" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="kasei">
<emphasis id="0CDEEADED369FF88F98A3E93F921FB18" box="[1620,1701,1237,1259]" italics="true" pageId="3" pageNumber="466">C. kasei</emphasis>
Osawa &amp; Takeda, 2004
</taxonomicName>
), Komai
<emphasis id="0CDEEADED369FF88FA133EB4F984FAFB" box="[1485,1536,1266,1288]" italics="true" pageId="3" pageNumber="466">et al.</emphasis>
(2010)
</td>
</tr>
<tr id="809A348ED3690074FF4D3F55F8E6FADA" box="[147,1890,1299,1321]" gridrow="17" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3F55FD7AFADA" box="[147,766,1299,1321]" gridcol="0" gridrow="17" pageId="3" pageNumber="466">
<taxonomicName id="F9AA4D4FD369FF88FF4D3F55FD1CFADA" authority="Osawa &amp; Takeda, 2004" authorityName="Osawa &amp; Takeda" authorityYear="2004" box="[147,664,1299,1321]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="iejimensis">
<emphasis id="0CDEEADED369FF88FF4D3F55FE25FADA" box="[147,417,1299,1321]" italics="true" pageId="3" pageNumber="466">Catapaguroides iejimensis</emphasis>
Osawa &amp; Takeda, 2004
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83F55FA86FADA" box="[886,1282,1299,1321]" gridcol="1" gridrow="17" pageId="3" pageNumber="466">Ie Island, Kume Island</td>
<td id="C34B5DF2D3690074FAE43F55F8E6FADA" box="[1338,1890,1299,1321]" gridcol="2" gridrow="17" pageId="3" pageNumber="466">Osawa &amp; Takeda (2004), Osawa &amp; Okuno (2007)</td>
</tr>
<tr id="809A348ED3690074FF4D3F71F8E6FABE" box="[147,1890,1335,1357]" gridrow="18" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3F71FD7AFABE" box="[147,766,1335,1357]" gridcol="0" gridrow="18" pageId="3" pageNumber="466">
<taxonomicName id="F9AA4D4FD369FF88FF4D3F71FD7AFABE" authority="Komai, Yamada &amp; Shirakawa, 2010" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[147,766,1335,1357]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="0CDEEADED369FF88FF4D3F71FE03FABE" box="[147,391,1335,1357]" italics="true" pageId="3" pageNumber="466">Catapaguroides longior</emphasis>
Komai, Yamada &amp; Shirakawa, 2010
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83F71FA86FABE" box="[886,1282,1335,1357]" gridcol="1" gridrow="18" pageId="3" pageNumber="466">Okinawa Island</td>
<td id="C34B5DF2D3690074FAE43F71F8E6FABE" box="[1338,1890,1335,1357]" gridcol="2" gridrow="18" pageId="3" pageNumber="466">
Komai
<emphasis id="0CDEEADED369FF88FA5B3F71FA3DFABE" box="[1413,1465,1335,1357]" italics="true" pageId="3" pageNumber="466">et al.</emphasis>
(2010)
</td>
</tr>
<tr id="809A348ED3690074FF4D3F1DF8E6FA82" box="[147,1890,1371,1393]" gridrow="19" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3F1DFD7AFA82" box="[147,766,1371,1393]" gridcol="0" gridrow="19" pageId="3" pageNumber="466">
<taxonomicName id="F9AA4D4FD369FF88FF4D3F1DFDCEFA82" authority="(Asakura, 1999)" baseAuthorityName="Asakura" baseAuthorityYear="1999" box="[147,586,1371,1393]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="tuberculosus">
<emphasis id="0CDEEADED369FF88FF4D3F1DFE24FA82" box="[147,416,1371,1393]" italics="true" pageId="3" pageNumber="466">Catapagurus tuberculosus</emphasis>
(Asakura, 1999)
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83F1DFA86FA82" box="[886,1282,1371,1393]" gridcol="1" gridrow="19" pageId="3" pageNumber="466">Ie Island, Shimoji Island</td>
<td id="C34B5DF2D3690074FAE43F1DF8E6FA82" box="[1338,1890,1371,1393]" gridcol="2" gridrow="19" pageId="3" pageNumber="466">Osawa &amp; Takeda (2004), Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D3F39F8E6FA66" box="[147,1890,1407,1429]" gridrow="20" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3F39FD7AFA66" box="[147,766,1407,1429]" gridcol="0" gridrow="20" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3F39FE2DFA66" box="[147,425,1407,1429]" italics="true" pageId="3" pageNumber="466">Kumepagurus cavernicolus</emphasis>
Komai &amp; Osawa, 2012
</th>
<td id="C34B5DF2D3690074FCA83F39FA86FA66" box="[886,1282,1407,1429]" gridcol="1" gridrow="20" pageId="3" pageNumber="466">Kume Island</td>
<td id="C34B5DF2D3690074FAE43F39F8E6FA66" box="[1338,1890,1407,1429]" gridcol="2" gridrow="20" pageId="3" pageNumber="466">Komai &amp; Osawa (2012)</td>
</tr>
<tr id="809A348ED3690074FF4D3FE5F8E6FA4A" box="[147,1890,1443,1465]" gridrow="21" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3FE5FD7AFA4A" box="[147,766,1443,1465]" gridcol="0" gridrow="21" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3FE5FEF0FA4A" box="[147,372,1443,1465]" italics="true" pageId="3" pageNumber="466">Nematopagurus lipkei</emphasis>
McLaughlin &amp; Okuno, 2010
</th>
<td id="C34B5DF2D3690074FCA83FE5FA86FA4A" box="[886,1282,1443,1465]" gridcol="1" gridrow="21" pageId="3" pageNumber="466">Okinawa Island</td>
<td id="C34B5DF2D3690074FAE43FE5F8E6FA4A" box="[1338,1890,1443,1465]" gridcol="2" gridrow="21" pageId="3" pageNumber="466">McLaughlin &amp; Okuno (2010)</td>
</tr>
<tr id="809A348ED3690074FF4D3F81F8E6FA2E" box="[147,1890,1479,1501]" gridrow="22" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3F81FD7AFA2E" box="[147,766,1479,1501]" gridcol="0" gridrow="22" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3F81FF7FFA2E" box="[147,251,1479,1501]" italics="true" pageId="3" pageNumber="466">Pagurixus</emphasis>
<taxonomicName id="F9AA4D4FD369FF88FEDF3F81FD0BFA2E" authority="Osawa, Fujita &amp; Okuno, 2006" authorityName="Osawa, Fujita &amp; Okuno" authorityYear="2006" box="[257,655,1479,1501]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="0CDEEADED369FF88FEDF3F81FED3FA2E" box="[257,343,1479,1501]" italics="true" pageId="3" pageNumber="466">longipes</emphasis>
Osawa, Fujita &amp; Okuno, 2006
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83F81FA86FA2E" box="[886,1282,1479,1501]" gridcol="1" gridrow="22" pageId="3" pageNumber="466">Kume Island</td>
<td id="C34B5DF2D3690074FAE43F81F8E6FA2E" box="[1338,1890,1479,1501]" gridcol="2" gridrow="22" pageId="3" pageNumber="466">
Osawa
<emphasis id="0CDEEADED369FF88FA5B3F81FA3DFA2E" box="[1413,1465,1479,1501]" italics="true" pageId="3" pageNumber="466">et al.</emphasis>
(2006)
</td>
</tr>
<tr id="809A348ED3690074FF4D3FADF8E6F9F2" box="[147,1890,1515,1537]" gridrow="23" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3FADFD7AF9F2" box="[147,766,1515,1537]" gridcol="0" gridrow="23" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3FADFF7FF9F2" box="[147,251,1515,1537]" italics="true" pageId="3" pageNumber="466">Pagurixus</emphasis>
<taxonomicName id="F9AA4D4FD369FF88FEDF3FADFDD3F9F2" authority="Komai &amp; Asakura, 1995" authorityName="Komai &amp; Asakura" authorityYear="1995" box="[257,599,1515,1537]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="0CDEEADED369FF88FEDF3FADFED3F9F2" box="[257,343,1515,1537]" italics="true" pageId="3" pageNumber="466">nomurai</emphasis>
Komai &amp; Asakura, 1995
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83FADFA86F9F2" box="[886,1282,1515,1537]" gridcol="1" gridrow="23" pageId="3" pageNumber="466">Ie Island, Irabu Island, Shimoji Island</td>
<td id="C34B5DF2D3690074FAE43FADF8E6F9F2" box="[1338,1890,1515,1537]" gridcol="2" gridrow="23" pageId="3" pageNumber="466">Osawa &amp; Takeda (2004), Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D3C49F8E6F9D6" box="[147,1890,1551,1573]" gridrow="24" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3C49FD7AF9D6" box="[147,766,1551,1573]" gridcol="0" gridrow="24" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3C49FECBF9D6" box="[147,335,1551,1573]" italics="true" pageId="3" pageNumber="466">Pagurixus pulcher</emphasis>
Osawa, Fujita &amp; Okuno, 2006
</th>
<td id="C34B5DF2D3690074FCA83C49FA86F9D6" box="[886,1282,1551,1573]" gridcol="1" gridrow="24" pageId="3" pageNumber="466">Kume Island</td>
<td id="C34B5DF2D3690074FAE43C49F8E6F9D6" box="[1338,1890,1551,1573]" gridcol="2" gridrow="24" pageId="3" pageNumber="466">
Osawa
<emphasis id="0CDEEADED369FF88FA5B3C49FA3DF9D6" box="[1413,1465,1551,1573]" italics="true" pageId="3" pageNumber="466">et al.</emphasis>
(2006)
</td>
</tr>
<tr id="809A348ED3690074FF4D3C75F8E6F9BA" box="[147,1890,1587,1609]" gridrow="25" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3C75FD7AF9BA" box="[147,766,1587,1609]" gridcol="0" gridrow="25" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3C75FEB6F9BA" box="[147,306,1587,1609]" italics="true" pageId="3" pageNumber="466">Pylopaguropsis</emphasis>
<taxonomicName id="F9AA4D4FD369FF88FEE63C75FDEBF9BA" authority="Osawa &amp; Okuno, 2007" authorityName="Osawa &amp; Okuno" authorityYear="2007" box="[312,623,1587,1609]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="0CDEEADED369FF88FEE63C75FEFAF9BA" box="[312,382,1587,1609]" italics="true" pageId="3" pageNumber="466">bellula</emphasis>
Osawa &amp; Okuno, 2007
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83C75FA86F9BA" box="[886,1282,1587,1609]" gridcol="1" gridrow="25" pageId="3" pageNumber="466">Kume Island, Shimoji Island</td>
<td id="C34B5DF2D3690074FAE43C75F8E6F9BA" box="[1338,1890,1587,1609]" gridcol="2" gridrow="25" pageId="3" pageNumber="466">Osawa &amp; Okuno (2007), Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D3C11F8E6F99E" box="[147,1890,1623,1645]" gridrow="26" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3C11FD7AF99E" box="[147,766,1623,1645]" gridcol="0" gridrow="26" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3C11FEB6F99E" box="[147,306,1623,1645]" italics="true" pageId="3" pageNumber="466">Pylopaguropsis</emphasis>
<taxonomicName id="F9AA4D4FD369FF88FEE63C11FD2FF99E" authority="McLaughlin &amp; Haig, 1989" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[312,683,1623,1645]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="0CDEEADED369FF88FEE63C11FE12F99E" box="[312,406,1623,1645]" italics="true" pageId="3" pageNumber="466">fimbriata</emphasis>
McLaughlin &amp; Haig, 1989
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83C11FA86F99E" box="[886,1282,1623,1645]" gridcol="1" gridrow="26" pageId="3" pageNumber="466">Ie Island, Irabu Island, Shimoji Island</td>
<td id="C34B5DF2D3690074FAE43C11F8E6F99E" box="[1338,1890,1623,1645]" gridcol="2" gridrow="26" pageId="3" pageNumber="466">Osawa &amp; Takeda (2004), Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D3C3DF8E6F962" box="[147,1890,1659,1681]" gridrow="27" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3C3DFD7AF962" box="[147,766,1659,1681]" gridcol="0" gridrow="27" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3C3DFEB6F962" box="[147,306,1659,1681]" italics="true" pageId="3" pageNumber="466">Pylopaguropsis</emphasis>
<taxonomicName id="F9AA4D4FD369FF88FEE63C3DFD20F962" authority="McLaughlin &amp; Haig, 1989" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[312,676,1659,1681]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="0CDEEADED369FF88FEE63C3DFE0BF962" box="[312,399,1659,1681]" italics="true" pageId="3" pageNumber="466">speciosa</emphasis>
McLaughlin &amp; Haig, 1989
</taxonomicName>
</th>
<td id="C34B5DF2D3690074FCA83C3DFA86F962" box="[886,1282,1659,1681]" gridcol="1" gridrow="27" pageId="3" pageNumber="466">Irabu Island, Shimoji Island</td>
<td id="C34B5DF2D3690074FAE43C3DF8E6F962" box="[1338,1890,1659,1681]" gridcol="2" gridrow="27" pageId="3" pageNumber="466">Osawa &amp; Takeda (2004), Osawa &amp; Fujita (2016)</td>
</tr>
<tr id="809A348ED3690074FF4D3CD9F8E6F946" box="[147,1890,1695,1717]" gridrow="28" pageId="3" pageNumber="466">
<th id="C34B5DF2D3690074FF4D3CD9FD7AF946" box="[147,766,1695,1717]" gridcol="0" gridrow="28" pageId="3" pageNumber="466">
<emphasis id="0CDEEADED369FF88FF4D3CD9FEC7F946" box="[147,323,1695,1717]" italics="true" pageId="3" pageNumber="466">Turleania saliens</emphasis>
Osawa &amp; Fujita, 2008
</th>
<td id="C34B5DF2D3690074FCA83CD9FA86F946" box="[886,1282,1695,1717]" gridcol="1" gridrow="28" pageId="3" pageNumber="466">Kume Island, Irabu Island</td>
<td id="C34B5DF2D3690074FAE43CD9F8E6F946" box="[1338,1890,1695,1717]" gridcol="2" gridrow="28" pageId="3" pageNumber="466">Osawa &amp; Fujita (2008)</td>
</tr>
</table>
</paragraph>
<paragraph id="3E1536CCD36EFF8FFF493ADEFBDDFE96" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="467">
<specimenCount id="28ACFD45D36EFF8FFF493ADEFE86FF43" box="[151,258,151,176]" pageId="4" pageNumber="467" type="female">4 females</specimenCount>
(sl
<quantity id="F9529B29D36EFF8FFEF23AD1FE32FF43" box="[300,438,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="5.2" metricValueMin="2.6" pageId="4" pageNumber="467" unit="mm" value="3.9" valueMax="5.2" valueMin="2.6">2.6–5.2 mm</quantity>
), 4 ovigerous females (sl
<quantity id="F9529B29D36EFF8FFD0C3AD1FCD8FF43" box="[722,860,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.5" metricValueMin="3.7" pageId="4" pageNumber="467" unit="mm" value="4.6" valueMax="5.5" valueMin="3.7">3.7–5.5 mm</quantity>
), RUMF-ZC-6016 (
<collectionCode id="58BBAE09D36EFF8FFBE53AD1FBE1FF43" box="[1083,1125,151,176]" pageId="4" pageNumber="467">HD</collectionCode>
<date id="4A14100CD36EFF8FFBBB3AD1FB41FF43" box="[1125,1221,151,176]" pageId="4" pageNumber="467" value="2016" valueMax="2070">2016-70</date>
–80); main tube,
<quantity id="F9529B29D36EFF8FFA5E3ADEFF29FF27" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="467" unit="m" value="30.0">30 m</quantity>
from entrance, area of rocks,
<date id="4A14100CD36EFF8FFE253AFAFD34FF26" box="[507,688,188,213]" pageId="4" pageNumber="467" value="2016-08-12">12 August 2016</date>
,
<specimenCount id="28ACFD45D36EFF8FFD653AFAFC94FF26" box="[699,784,188,213]" pageId="4" pageNumber="467" type="male">3 males</specimenCount>
(sl 4.0–5.0 mm), 1 ovigerous female (sl
<quantity id="F9529B29D36EFF8FFB093AFAFAB5FF27" box="[1239,1329,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="4" pageNumber="467" unit="mm" value="5.8">5.8 mm</quantity>
), RUMF- ZC-6017 (
<collectionCode id="58BBAE09D36EFF8FFED53A99FEB2FF0B" box="[267,310,223,248]" pageId="4" pageNumber="467">HD</collectionCode>
<date id="4A14100CD36EFF8FFEE83A99FE11FF0B" box="[310,405,223,248]" pageId="4" pageNumber="467" value="2016" valueMax="2089">2016-89</date>
–92); main tube,
<date id="4A14100CD36EFF8FFD8A3AA6FD6AFF0B" box="[596,750,224,248]" pageId="4" pageNumber="467" value="2017-05-21">21 May 2017</date>
,
<specimenCount id="28ACFD45D36EFF8FFD273AA6FCD5FF0B" box="[761,849,223,248]" pageId="4" pageNumber="467" type="male">3 males</specimenCount>
(sl
<quantity id="F9529B29D36EFF8FFCA23A99FB83FF0B" box="[892,1031,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="5.9" metricValueMin="2.5" pageId="4" pageNumber="467" unit="mm" value="4.2" valueMax="5.9" valueMin="2.5">2.5–5.9 mm</quantity>
),
<materialsCitation id="8EC23C91D36EFF8FFBC53AA6FD72FEEE" ID-GBIF-Occurrence="2234219640" collectingDate="2017" collectingDateMax="2084" collectingDateMin="2017" pageId="4" pageNumber="467" specimenCode="RUMF-ZC- 6018" specimenCount="1" specimenCount-female="1">
<specimenCount id="28ACFD45D36EFF8FFBC53AA6FBF9FF0B" box="[1051,1149,223,248]" pageId="4" pageNumber="467" type="female">1 female</specimenCount>
(sl
<quantity id="F9529B29D36EFF8FFB763A99FA87FF0B" box="[1192,1283,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="4" pageNumber="467" unit="mm" value="5.1">5.1 mm</quantity>
),
<specimenCode id="6E0C9EB7D36EFF8FFAC83AA6FF4BFEEE" pageId="4" pageNumber="467">RUMF-ZC- 6018</specimenCode>
(HD
<date id="4A14100CD36EFF8FFECE3B42FEF5FEEF" box="[272,369,260,284]" pageId="4" pageNumber="467" value="2017" valueMax="2084">2017-84</date>
); same data as RUMF-ZC-6018
</materialsCitation>
,
<materialsCitation id="8EC23C91D36EFF8FFCD73B42FC46FEB3" ID-GBIF-Occurrence="2234219559" collectingDate="2017-02-04" collectingDateMax="2089" collectingDateMin="2017" location="entrance" municipality="Akuma-no-Yakata cave" pageId="4" pageNumber="467" specimenCode="RUMF-ZC-6019" specimenCount="1" specimenCount-female="1" stateProvince="Shimoji Island, Akuma-no-Yakata cave">
<specimenCount id="28ACFD45D36EFF8FFCD73B42FCF9FEEE" box="[777,893,260,285]" pageId="4" pageNumber="467" type="female">1 females</specimenCount>
(sl
<quantity id="F9529B29D36EFF8FFC6B3B42FB93FEEF" box="[949,1047,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="467" unit="mm" value="5.3">5.3 mm</quantity>
),
<specimenCode id="6E0C9EB7D36EFF8FFBEF3B42FB6AFEEE" box="[1073,1262,260,285]" pageId="4" pageNumber="467">RUMF-ZC-6019</specimenCode>
(HD
<date id="4A14100CD36EFF8FFAEE3B42FA14FEEF" box="[1328,1424,260,284]" pageId="4" pageNumber="467" value="2017" valueMax="2089">2017-89</date>
).
<emphasis id="0CDEEADED36EFF8FFF493B61FD06FEB3" bold="true" box="[151,642,295,320]" pageId="4" pageNumber="467">
<collectingRegion id="FC6EF82ED36EFF8FFF493B61FDF9FEB3" box="[151,637,295,320]" pageId="4" pageNumber="467">Shimoji Island, Akuma-no-Yakata cave</collectingRegion>
:
</emphasis>
<location id="3B756017D36EFF8FFD533B6EFD6BFECC" LSID="urn:lsid:plazi:treatment:B60387DAD368FF8FFF493C01FCAFFD76:3B756017D36EFF8FFD533B6EFD6BFECC" box="[653,751,296,319]" municipality="Akuma-no-Yakata cave" name="entrance" pageId="4" pageNumber="467" stateProvince="Shimoji Island, Akuma-no-Yakata cave">entrance</location>
,
<collectingDate id="5A50E9E4D36EFF8FFD203B6EFC46FEB3" box="[766,962,295,320]" pageId="4" pageNumber="467" value="2017-02-04">
<date id="4A14100CD36EFF8FFD203B6EFC46FEB3" box="[766,962,295,320]" pageId="4" pageNumber="467" value="2017-02-04">4 February 2017</date>
</collectingDate>
</materialsCitation>
,
<materialsCitation id="8EC23C91D36EFF8FFC0C3B6EFDC9FE96" ID-GBIF-Occurrence="2234219656" collectingDate="2017" collectingDateMax="2089" collectingDateMin="2017" location="entrance" municipality="Akuma-no-Yakata cave" pageId="4" pageNumber="467" specimenCode="RUMF-ZC-6013" specimenCount="1" specimenCount-female="1" stateProvince="Shimoji Island, Akuma-no-Yakata cave">
<specimenCount id="28ACFD45D36EFF8FFC0C3B6EFBB3FEB3" box="[978,1079,295,320]" pageId="4" pageNumber="467" type="female">1 female</specimenCount>
(sl
<quantity id="F9529B29D36EFF8FFBB73B61FB43FEB3" box="[1129,1223,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="4" pageNumber="467" unit="mm" value="6.3">6.3 mm</quantity>
),
<specimenCode id="6E0C9EB7D36EFF8FFB003B61FA18FEB3" box="[1246,1436,295,320]" pageId="4" pageNumber="467">RUMF-ZC-6013</specimenCode>
(SH53); same data as RUMF-ZC-6013
</materialsCitation>
,
<specimenCount id="28ACFD45D36EFF8FFD863B0AFD27FE96" box="[600,675,332,357]" pageId="4" pageNumber="467" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D36EFF8FFD103B0AFCACFE97" box="[718,808,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="4" pageNumber="467" unit="mm" value="5.7">5.7 mm</quantity>
), RUMF-ZC-6014 (
<collectionCode id="58BBAE09D36EFF8FFBD63B0AFBABFE96" box="[1032,1071,332,357]" pageId="4" pageNumber="467">SH</collectionCode>
54).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D36EFF8FFF193B29FEB3FE07" pageId="4" pageNumber="467" type="distribution">
<paragraph id="3E1536CCD36EFF8FFF193B29FC45FE5F" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="467">
<emphasis id="0CDEEADED36EFF8FFF193B29FEE4FE7B" bold="true" box="[199,352,367,392]" pageId="4" pageNumber="467">Distribution.</emphasis>
Eastern Indian Ocean:
<collectingCountry id="46BD765CD36EFF8FFDAD3B36FC0CFE7B" box="[627,904,367,392]" name="Cocos (Keeling) Islands" pageId="4" pageNumber="467">Cocos (Keeling) Islands</collectingCountry>
and
<collectingCountry id="46BD765CD36EFF8FFC1B3B29FB0CFE7B" box="[965,1160,367,392]" name="Christmas Island" pageId="4" pageNumber="467">Christmas Island</collectingCountry>
; western Pacific:
<collectingCountry id="46BD765CD36EFF8FFA893B36FA18FE7B" box="[1367,1436,368,392]" name="Guam" pageId="4" pageNumber="467">Guam</collectingCountry>
and Ryukyu Islands in southwestern
<collectingCountry id="46BD765CD36EFF8FFDED3BD3FDF3FE5F" box="[563,631,405,428]" name="Japan" pageId="4" pageNumber="467">Japan</collectingCountry>
;
<quantity id="F9529B29D36EFF8FFD5F3BD2FD52FE5F" box="[641,726,404,428]" metricMagnitude="1" metricUnit="m" metricValue="1.95" metricValueMax="3.7" metricValueMin="0.2" pageId="4" pageNumber="467" unit="m" value="19.5" valueMax="37.0" valueMin="2.0">2–37 m</quantity>
(McLaughlin 2007).
</paragraph>
<paragraph id="3E1536CCD36EFF8FFF193BF1FEB3FE07" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="467">
<emphasis id="0CDEEADED36EFF8FFF193BF1FEAEFE23" bold="true" box="[199,298,439,464]" pageId="4" pageNumber="467">Habitat.</emphasis>
Coral reefs; submarine caves and small crevices or horizontal holes on fore-reef slopes (
<bibRefCitation id="5A3B4B3DD36EFF8FFAF13BFEFEAFFE07" author="Osawa, M. &amp; Takeda, M." pageId="4" pageNumber="467" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D36EFF8FFF193BB9FCAFFD76" pageId="4" pageNumber="467" type="discussion">
<paragraph id="3E1536CCD36EFF8FFF193BB9FCAFFD76" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="467">
<emphasis id="0CDEEADED36EFF8FFF193BB9FEBFFDEB" bold="true" box="[199,315,511,536]" pageId="4" pageNumber="467">Remarks.</emphasis>
<taxonomicName id="F9AA4D4FD36EFF8FFE9B3846FE79FDEB" authorityName="Morgan" authorityYear="1992" box="[325,509,512,536]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="jalur">
<emphasis id="0CDEEADED36EFF8FFE9B3846FE79FDEB" box="[325,509,512,536]" italics="true" pageId="4" pageNumber="467">Paguristes jalur</emphasis>
</taxonomicName>
is the sole species of the genus recorded from shallow coral reef environments including submarine caves in the Ryukyu Islands (
<bibRefCitation id="5A3B4B3DD36EFF8FFD043862FBCEFDCF" author="Osawa, M. &amp; Takeda, M." box="[730,1098,548,573]" pageId="4" pageNumber="467" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">
Osawa &amp; Takeda 2004;
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). This species has been often found in small crevices or horizontal holes on fore-reef slopes of the islands, accompanied by
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Pylopaguropsis
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with similarly red stripes on the ambulatory legs.
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<emphasis id="0CDEEADED36DFF8CFF493C92FE51F91D" bold="true" box="[151,469,1748,1774]" italics="true" pageId="7" pageNumber="470">Catapagurus tuberculosus</emphasis>
(
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)
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(
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<emphasis id="0CDEEADED36DFF8CFF493D78FE99F8A7" box="[151,285,1854,1876]" italics="true" pageId="7" pageNumber="470">Icelopagurus</emphasis>
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<emphasis id="0CDEEADED36DFF8CFEFD3D78FE20F8A7" box="[291,420,1854,1876]" italics="true" pageId="7" pageNumber="470">tuberculosus</emphasis>
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: 382
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, figs. 1–4 (
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locality: Kume Island, Ryukyu Islands).
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<emphasis id="0CDEEADED36DFF8CFF493D18FE25F880" box="[151,417,1885,1907]" italics="true" pageId="7" pageNumber="470">Catapagurus tuberculosus</emphasis>
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.—
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: 830
</treatmentCitation>
, figs. 1D, G, 51.—
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: 1112
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<emphasis id="0CDEEADED36DFF8CFF493DD9FE05F84B" bold="true" box="[151,385,1951,1976]" pageId="7" pageNumber="470">Material examined.</emphasis>
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Ie
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, unnamed submarine cave:
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entrance,
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<date id="4A14100CD36DFF8CFC683DD9FBD4F84B" box="[950,1104,1951,1976]" pageId="7" pageNumber="470" value="2017-06-25">25 June 2017</date>
</collectingDate>
,
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(sl
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188
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<emphasis id="0CDEEADED362FF83FF193AD1FEE4FF43" bold="true" box="[199,352,151,176]" pageId="8" pageNumber="471">Distribution.</emphasis>
Ryukyu Islands (Ie
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, Kume Island, and Shimoji Island) in southwestern
<collectingCountry id="46BD765CD362FF83FB393ADEFAAFFF5C" box="[1255,1323,152,175]" name="Japan" pageId="8" pageNumber="471">Japan</collectingCountry>
;
<quantity id="F9529B29D362FF83FAE93AD1FA18FF5C" box="[1335,1436,151,176]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="3.5" metricValueMin="1.0" pageId="8" pageNumber="471" unit="m" value="22.5" valueMax="35.0" valueMin="10.0">10–35 m</quantity>
(
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;
<bibRefCitation id="5A3B4B3DD362FF83FE703AFAFD26FF26" author="Osawa, M. &amp; Fujita, Y." box="[430,674,188,213]" pageId="8" pageNumber="471" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita 2016</bibRefCitation>
; present study).
</paragraph>
<paragraph id="3E1536CCD362FF83FF193A99FA8CFF0B" blockId="8.[151,1436,151,464]" box="[199,1288,223,248]" pageId="8" pageNumber="471">
<emphasis id="0CDEEADED362FF83FF193A99FEAEFF0B" bold="true" box="[199,298,223,248]" pageId="8" pageNumber="471">Habitat.</emphasis>
Coral reefs; often found on fine sediments in submarine caves (
<bibRefCitation id="5A3B4B3DD362FF83FC283AA6FB78FF0B" author="Osawa, M. &amp; Takeda, M." box="[1014,1276,223,248]" pageId="8" pageNumber="471" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D362FF83FF193B42FDCAFE23" pageId="8" pageNumber="471" type="discussion">
<paragraph id="3E1536CCD362FF83FF193B42FDCAFE23" blockId="8.[151,1436,151,464]" pageId="8" pageNumber="471">
<emphasis id="0CDEEADED362FF83FF193B42FEBFFEEE" bold="true" box="[199,315,260,285]" pageId="8" pageNumber="471">Remarks.</emphasis>
This species was originally described as
<emphasis id="0CDEEADED362FF83FCC13B42FBCFFEEF" box="[799,1099,260,284]" italics="true" pageId="8" pageNumber="471">
Icelopagurus
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</emphasis>
by
<bibRefCitation id="5A3B4B3DD362FF83FBA23B42FAB4FEEF" author="Asakura, A." box="[1148,1328,260,285]" pageId="8" pageNumber="471" pagination="381 - 395" refId="ref9413" refString="Asakura, A. (1999) Icelopagurus tuberculosus, a distinct new hermit crab species (Crustacea: Decapoda: Paguridae) from Japan. Proceedings of the Biological Society of Washington, 112 (2), 381 - 395." type="journal article" year="1999">Asakura (1999)</bibRefCitation>
.
<bibRefCitation id="5A3B4B3DD362FF83FAE13B42FF65FEB3" author="Asakura, A." pageId="8" pageNumber="471" pagination="823 - 891" refId="ref9504" refString="Asakura, A. (2001) A revision of the hermit crabs of the genera Catapagurus A. Milne-Edwards and Hemipagurus Smith from the Indo-West Pacific (Crustacea: Decapoda: Anomura: Paguridae). Invertebrate Taxonomy, 15, 823 - 891. https: // doi. org / 10.1071 / IT 00034" type="journal article" year="2001">Asakura (2001)</bibRefCitation>
subsequently transferred the species to the genus
<taxonomicName id="F9AA4D4FD362FF83FCCD3B6EFB47FEB3" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" box="[787,1219,295,320]" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="0CDEEADED362FF83FCCD3B6EFC21FECC" box="[787,933,296,319]" italics="true" pageId="8" pageNumber="471">Catapagurus</emphasis>
<bibRefCitation id="5A3B4B3DD362FF83FC723B6EFB47FEB3" author="Milne-Edwards, A." box="[940,1219,295,320]" pageId="8" pageNumber="471" pagination="1 - 68" refId="ref12024" refString="Milne-Edwards, A. (1880) Report on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, 78, 79, by the United States Coast Survey steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., commanding. VIII. Etudes preliminaires sur les Crustaces. Bulletin of the Museum of Comparative Zoology, Harvard College, 8, 1 - 68." type="journal article" year="1880">A. Milne-Edwards, 1880</bibRefCitation>
</taxonomicName>
from
<emphasis id="0CDEEADED362FF83FAD63B6EFA18FEB3" box="[1288,1436,296,320]" italics="true" pageId="8" pageNumber="471">Icelopagurus</emphasis>
<bibRefCitation id="5A3B4B3DD362FF83FF493B0AFEF4FE97" author="McLaughlin, P. A." box="[151,368,332,357]" pageId="8" pageNumber="471" pagination="433 - 572" refId="ref11331" refString="McLaughlin, P. A. (1997) Crustacea Decapoda: Hermit crabs of the family Paguridae from the KARUBAR cruise in Indonesia. In: Crosnier, A. &amp; Bouchet, P. (Eds.), Resultats des Campagnes MUSORSTOM. Fol. 16. Memoires du Museum national d'Histoire naturelle, Paris, 172, pp. 433 - 572." type="journal article" year="1997">McLaughlin, 1997</bibRefCitation>
.
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<emphasis id="0CDEEADED362FF83FE5F3B0AFD2AFE97" box="[385,686,332,356]" italics="true" pageId="8" pageNumber="471">Catapagurus tuberculosus</emphasis>
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has been often recorded from submarine caves in the Ryukyu Islands (
<bibRefCitation id="5A3B4B3DD362FF83FF2B3B36FE7DFE7B" author="Osawa, M. &amp; Takeda, M." box="[245,505,367,392]" pageId="8" pageNumber="471" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
,
<bibRefCitation id="5A3B4B3DD362FF83FDDA3B36FD73FE7B" author="Osawa, M. &amp; Fujita, Y." box="[516,759,367,392]" pageId="8" pageNumber="471" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita 2016</bibRefCitation>
;
<tableCitation id="73280377D362FF83FCDD3B29FCDDFE7B" box="[771,857,367,392]" captionStart="TABLE 2" captionStartId="6.[147,233,596,618]" captionText="TABLE 2. Submarine caves surveyed and hermit crab species collected in three islands of the Ryukyu Islands, southwestern Japan." pageId="8" pageNumber="471">Table 2</tableCitation>
). As noted by
<bibRefCitation id="5A3B4B3DD362FF83FC223B36FA94FE7B" author="Osawa, M. &amp; Takeda, M." box="[1020,1296,367,392]" pageId="8" pageNumber="471" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda (2004)</bibRefCitation>
, this species carries a gastropod shell, which protects only the abdomen and posterior carapace, being too small for the hermit crab to retract the thoracic appendages.
</paragraph>
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275
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<emphasis id="0CDEEADED36DFF8CFF49388AFE30FD15" bold="true" box="[151,436,716,742]" italics="true" pageId="7" pageNumber="470">Catapaguroides longior</emphasis>
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(
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E–J)
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, figs. 1–4, 5A (
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locality: Okinawa Island, Ryukyu Islands).
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Ie
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main tube, near bifurcation of tube,
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main tube,
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<quantity id="F9529B29D36DFF8CFB7639FAFA87FC27" box="[1192,1283,956,980]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="7" pageNumber="470" unit="mm" value="2.1">2.1 mm</quantity>
), RUMF-ZC- 6028 (
<specimenCode id="6E0C9EB7D36DFF8CFF3E3999FE98FC0B" box="[224,284,991,1016]" collectionCode="IE" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="7" pageNumber="470">
<collectionCode id="58BBAE09D36DFF8CFF3E3999FF7AFC0B" box="[224,254,991,1016]" pageId="7" pageNumber="470">IE</collectionCode>
65
</specimenCode>
).
</materialsCitation>
<materialsCitation id="8EC23C91D36DFF8CFEEC3999FAC4FBEF" ID-GBIF-Occurrence="2465041912" collectingDate="2017-05-22" collectionRegion="Okinawa Island, Hedo Dome cave" country="Japan" location="silty room" pageId="7" pageNumber="470" specimenCode="RUMF-ZC-6029, RUMF-ZC-6030" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<emphasis id="0CDEEADED36DFF8CFEEC3999FD5DFC0B" bold="true" box="[306,729,991,1016]" pageId="7" pageNumber="470">
<location id="3B756017D36DFF8CFEEC3999FE71FC0B" LSID="urn:lsid:plazi:treatment:B60387DAD36DFF8CFF49388AFC35F973:3B756017D36DFF8CFEEC3999FE71FC0B" box="[306,501,991,1016]" country="Iceland" municipality="Hedo Dome cave" name="Okinawa Island" pageId="7" pageNumber="470">Okinawa Island</location>
,
<collectingMunicipality id="DE71ACB6D36DFF8CFDDD3999FD50FC0B" box="[515,724,991,1016]" pageId="7" pageNumber="470">Hedo Dome cave</collectingMunicipality>
:
</emphasis>
silty room,
<collectingDate id="5A50E9E4D36DFF8CFCB639A6FB81FC0B" box="[872,1029,992,1016]" pageId="7" pageNumber="470" value="2017-05-22">
<date id="4A14100CD36DFF8CFCB639A6FB81FC0B" box="[872,1029,992,1016]" pageId="7" pageNumber="470" value="2017-05-22">22 May 2017</date>
</collectingDate>
,
<specimenCount id="28ACFD45D36DFF8CFBCC39A6FBF2FC0B" box="[1042,1142,991,1016]" pageId="7" pageNumber="470" type="female">1 female</specimenCount>
(sl
<quantity id="F9529B29D36DFF8CFB7B39A6FA85FC0B" box="[1189,1281,992,1016]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="470" unit="mm" value="1.2">1.2 mm</quantity>
), RUMF-ZC- 6029 (
<collectionCode id="58BBAE09D36DFF8CFF003E42FE8DFBEF" box="[222,265,1028,1052]" pageId="7" pageNumber="470">HD</collectionCode>
2017-107); same data as RUMF-ZC-6029,
<specimenCount id="28ACFD45D36DFF8CFD353E42FCB2FBEE" box="[747,822,1028,1053]" pageId="7" pageNumber="470" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D36DFF8CFCBF3E42FC3FFBEF" box="[865,955,1028,1052]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="470" unit="mm" value="1.4">1.4 mm</quantity>
), RUMF-ZC-6030 (
<collectionCode id="58BBAE09D36DFF8CFB453E42FB43FBEF" box="[1179,1223,1028,1052]" pageId="7" pageNumber="470">HD</collectionCode>
2017-141).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="76B06547D36DFF8CFF193E61FAA6FB7B" pageId="7" pageNumber="470" type="distribution">
<paragraph id="3E1536CCD36DFF8CFF193E61FE07FB96" blockId="7.[151,1437,919,1664]" pageId="7" pageNumber="470">
<emphasis id="0CDEEADED36DFF8CFF193E61FEE4FBB3" bold="true" box="[199,352,1063,1088]" pageId="7" pageNumber="470">Distribution.</emphasis>
Ryukyu Islands (Ie
<collectingCountry id="46BD765CD36DFF8CFD973E61FD09FBB3" box="[585,653,1063,1088]" name="Iceland" pageId="7" pageNumber="470">Island</collectingCountry>
and Okinawa Island) in southwestern
<collectingCountry id="46BD765CD36DFF8CFB993E6EFB0EFBCC" box="[1095,1162,1064,1087]" name="Japan" pageId="7" pageNumber="470">Japan</collectingCountry>
;
<quantity id="F9529B29D36DFF8CFB493E61FB78FBCC" box="[1175,1276,1063,1088]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="7" pageNumber="470" unit="m" value="22.5" valueMax="30.0" valueMin="15.0">15–30 m</quantity>
(Komai
<emphasis id="0CDEEADED36DFF8CFABC3E6FFA11FBB3" box="[1378,1429,1064,1088]" italics="true" pageId="7" pageNumber="470">et al</emphasis>
. 2010; present study).
</paragraph>
<paragraph id="3E1536CCD36DFF8CFF193E29FAA6FB7B" blockId="7.[151,1437,919,1664]" box="[199,1314,1135,1160]" pageId="7" pageNumber="470">
<emphasis id="0CDEEADED36DFF8CFF193E29FEAEFB7B" bold="true" box="[199,298,1135,1160]" pageId="7" pageNumber="470">Habitat.</emphasis>
Coral reefs; inside of a submarine cave about
<quantity id="F9529B29D36DFF8CFCEC3E36FCEFFB74" box="[818,875,1136,1160]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="470" unit="m" value="10.0">10 m</quantity>
from the entrance (Komai
<emphasis id="0CDEEADED36DFF8CFB443E37FB4EFB7B" box="[1178,1226,1136,1160]" italics="true" pageId="7" pageNumber="470">et al</emphasis>
. 2010).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D36DFF8CFF193ED2FC35F973" pageId="7" pageNumber="470" type="discussion">
<paragraph id="3E1536CCD36DFF8CFF193ED2FACFFB23" blockId="7.[151,1437,919,1664]" pageId="7" pageNumber="470">
<emphasis id="0CDEEADED36DFF8CFF193ED2FEBFFB5E" bold="true" box="[199,315,1172,1197]" pageId="7" pageNumber="470">Remarks.</emphasis>
<taxonomicName id="F9AA4D4FD36DFF8CFE9F3ED2FDC9FB5F" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[321,589,1172,1196]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="0CDEEADED36DFF8CFE9F3ED2FDC9FB5F" box="[321,589,1172,1196]" italics="true" pageId="7" pageNumber="470">Catapaguroides longior</emphasis>
</taxonomicName>
was described by Komai
<emphasis id="0CDEEADED36DFF8CFCB23ED3FC19FB5F" box="[876,925,1172,1196]" italics="true" pageId="7" pageNumber="470">et al</emphasis>
. (2010) based only on the male
<typeStatus id="E111886ED36DFF8CFADF3ED2FAE3FB5E" box="[1281,1383,1172,1197]" pageId="7" pageNumber="470" type="holotype">holotype</typeStatus>
. The present material includes a first female of the species (sl
<quantity id="F9529B29D36DFF8CFCCD3EFEFCEFFB3C" box="[787,875,1208,1232]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="470" unit="mm" value="1.2">1.2 mm</quantity>
, RUMF-ZC-6029, HD2017-107;
<figureCitation id="A6912A49D36DFF8CFB3B3EF1FAB9FB23" box="[1253,1341,1207,1232]" captionStart="FIGURE 1" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@5.[151,1436,193,1705]" captionTargetPageId="5" captionText="FIGURE 1. Entire animal, dorsal view. A, Aniculus retipes Lewinsohn, 1982, male (sl 11.9 mm), RUMF-ZC-6006 (IE141); B, Paguristes jalur Morgan, 1992, male (sl 5.4 mm), RUMF-ZC-6012 (IE69); C, Pseudopaguristes shidarai Asakura, 2004, male (sl 2.6 mm), RUMF-ZC-6020 (HD2017-140); D, Catapaguroides foresti McLaughlin, 2002, male (sl 2.0 mm), RUMFZC-6023 (IE173); E, Catapaguroides iejimensis Osawa &amp; Takeda, 2004, male (sl 1.8 mm), RUMF-ZC-6026 (IE177); F, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, male (sl 2.1 mm), RUMF-ZC-6027 (IE60); G, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107)." httpUri="https://zenodo.org/record/2627820/files/figure.png" pageId="7" pageNumber="470">Fig. 1G</figureCitation>
).
</paragraph>
<paragraph id="3E1536CCD36DFF8CFF193E9AFB6BFA93" blockId="7.[151,1437,919,1664]" pageId="7" pageNumber="470">
The
<typeStatus id="E111886ED36DFF8CFF253E9AFED9FB06" box="[251,349,1244,1269]" pageId="7" pageNumber="470" type="holotype">holotype</typeStatus>
has a small but distinct dorsomedian spine on the carpus of the right cheliped (cf. Komai
<emphasis id="0CDEEADED36DFF8CFABD3E9BFA11FB07" box="[1379,1429,1244,1268]" italics="true" pageId="7" pageNumber="470">et al</emphasis>
. 2010: fig. 2 A, D). In the present material, the spine is prominent only in a female and it is minute or absent in other male specimens (
<figureCitation id="A6912A49D36DFF8CFEBF3F62FE5FFACF" box="[353,475,1316,1341]" captionStart="FIGURE 4" captionStartId="12.[151,250,1444,1466]" captionTargetBox="[297,1299,511,1375]" captionTargetId="figure@12.[151,1436,483,1422]" captionTargetPageId="12" captionText="FIGURE 4. Catapaguroides foresti McLaughlin, 2002: male (sl 2.0 mm), RUMF-ZC-6023 (IE173) (A–C), male (sl 1.5 mm), RUMF-ZC-6024 (IE184) (D); Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010: male (sl 2.1 mm), RUMF-ZC- 6028 (IE65) (E–H), female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107) (I, J). A, E, shield and ocular peduncles, dorsal view; B, distal part of right ocular peduncle, lateral view; C, D, G, I, right chela, dorsal view; F, left ocular peduncle, lateral view; H, J, carpus of right cheliped, dorsal view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627826/files/figure.png" pageId="7" pageNumber="470">Fig. 4H, J</figureCitation>
). Additionally, the ventrolateral distal angle of the carpus of the right cheliped is unarmed in the
<typeStatus id="E111886ED36DFF8CFE983F01FE2FFA93" box="[326,427,1351,1376]" pageId="7" pageNumber="470" type="holotype">holotype</typeStatus>
, whereas it has a minute to small spine in the present material (
<figureCitation id="A6912A49D36DFF8CFBB13F01FB65FA93" box="[1135,1249,1351,1376]" captionStart="FIGURE 4" captionStartId="12.[151,250,1444,1466]" captionTargetBox="[297,1299,511,1375]" captionTargetId="figure@12.[151,1436,483,1422]" captionTargetPageId="12" captionText="FIGURE 4. Catapaguroides foresti McLaughlin, 2002: male (sl 2.0 mm), RUMF-ZC-6023 (IE173) (A–C), male (sl 1.5 mm), RUMF-ZC-6024 (IE184) (D); Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010: male (sl 2.1 mm), RUMF-ZC- 6028 (IE65) (E–H), female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107) (I, J). A, E, shield and ocular peduncles, dorsal view; B, distal part of right ocular peduncle, lateral view; C, D, G, I, right chela, dorsal view; F, left ocular peduncle, lateral view; H, J, carpus of right cheliped, dorsal view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627826/files/figure.png" pageId="7" pageNumber="470">Fig. 4H, J</figureCitation>
).
</paragraph>
<paragraph id="3E1536CCD36DFF8CFF193F2AFC35F973" blockId="7.[151,1437,919,1664]" pageId="7" pageNumber="470">
The fresh coloration superficially resembles between
<taxonomicName id="F9AA4D4FD36DFF8CFCE53F2AFC2CFA77" authorityName="McLaughlin" authorityYear="2002" box="[827,936,1388,1412]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="0CDEEADED36DFF8CFCE53F2AFC2CFA77" box="[827,936,1388,1412]" italics="true" pageId="7" pageNumber="470">C. foresti</emphasis>
</taxonomicName>
and
<taxonomicName id="F9AA4D4FD36DFF8CFC393F2AFBDBFA77" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[999,1119,1388,1412]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="0CDEEADED36DFF8CFC393F2AFBDBFA77" box="[999,1119,1388,1412]" italics="true" pageId="7" pageNumber="470">C. longior</emphasis>
</taxonomicName>
(
<figureCitation id="A6912A49D36DFF8CFBAD3F2AFA9CFA77" box="[1139,1304,1388,1413]" captionStart="FIGURE 1" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@5.[151,1436,193,1705]" captionTargetPageId="5" captionText="FIGURE 1. Entire animal, dorsal view. A, Aniculus retipes Lewinsohn, 1982, male (sl 11.9 mm), RUMF-ZC-6006 (IE141); B, Paguristes jalur Morgan, 1992, male (sl 5.4 mm), RUMF-ZC-6012 (IE69); C, Pseudopaguristes shidarai Asakura, 2004, male (sl 2.6 mm), RUMF-ZC-6020 (HD2017-140); D, Catapaguroides foresti McLaughlin, 2002, male (sl 2.0 mm), RUMFZC-6023 (IE173); E, Catapaguroides iejimensis Osawa &amp; Takeda, 2004, male (sl 1.8 mm), RUMF-ZC-6026 (IE177); F, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, male (sl 2.1 mm), RUMF-ZC-6027 (IE60); G, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107)." httpUri="https://zenodo.org/record/2627820/files/figure.png" pageId="7" pageNumber="470">Fig. 1D, F, G</figureCitation>
), but some morphological characters easily distinguish the two species. The ocular peduncles are comparatively longer and more slender in
<taxonomicName id="F9AA4D4FD36DFF8CFE953FF2FE36FA3F" authorityName="McLaughlin" authorityYear="2002" box="[331,434,1460,1484]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="0CDEEADED36DFF8CFE953FF2FE36FA3F" box="[331,434,1460,1484]" italics="true" pageId="7" pageNumber="470">C. foresti</emphasis>
</taxonomicName>
than
<taxonomicName id="F9AA4D4FD36DFF8CFE313FF2FDE4FA3F" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[495,608,1460,1484]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="0CDEEADED36DFF8CFE313FF2FDE4FA3F" box="[495,608,1460,1484]" italics="true" pageId="7" pageNumber="470">C. longior</emphasis>
</taxonomicName>
, and the corneas are conical in
<taxonomicName id="F9AA4D4FD36DFF8CFC603FF2FBA2FA3F" authorityName="McLaughlin" authorityYear="2002" box="[958,1062,1460,1484]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="0CDEEADED36DFF8CFC603FF2FBA2FA3F" box="[958,1062,1460,1484]" italics="true" pageId="7" pageNumber="470">C. foresti</emphasis>
</taxonomicName>
rather than rounded in
<taxonomicName id="F9AA4D4FD36DFF8CFAF73FF2FA18FA3F" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[1321,1436,1460,1484]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="0CDEEADED36DFF8CFAF73FF2FA18FA3F" box="[1321,1436,1460,1484]" italics="true" pageId="7" pageNumber="470">C. longior</emphasis>
</taxonomicName>
(
<figureCitation id="A6912A49D36DFF8CFF413F91FEDBFA03" box="[159,351,1495,1520]" captionStart="FIGURE 4" captionStartId="12.[151,250,1444,1466]" captionTargetBox="[297,1299,511,1375]" captionTargetId="figure@12.[151,1436,483,1422]" captionTargetPageId="12" captionText="FIGURE 4. Catapaguroides foresti McLaughlin, 2002: male (sl 2.0 mm), RUMF-ZC-6023 (IE173) (A–C), male (sl 1.5 mm), RUMF-ZC-6024 (IE184) (D); Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010: male (sl 2.1 mm), RUMF-ZC- 6028 (IE65) (E–H), female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107) (I, J). A, E, shield and ocular peduncles, dorsal view; B, distal part of right ocular peduncle, lateral view; C, D, G, I, right chela, dorsal view; F, left ocular peduncle, lateral view; H, J, carpus of right cheliped, dorsal view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627826/files/figure.png" pageId="7" pageNumber="470">Fig. 4A, B, E, F</figureCitation>
). In the right cheliped, the distomesial angle of the palm and the proximomesial margin of the dactylus have a small spine respectively in
<taxonomicName id="F9AA4D4FD36DFF8CFDA63FBAFD5AF9E7" authorityName="McLaughlin" authorityYear="2002" box="[632,734,1532,1556]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="0CDEEADED36DFF8CFDA63FBAFD5AF9E7" box="[632,734,1532,1556]" italics="true" pageId="7" pageNumber="470">C. foresti</emphasis>
</taxonomicName>
, instead of being unarmed in
<taxonomicName id="F9AA4D4FD36DFF8CFBF73FBAFB1EF9E7" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[1065,1178,1532,1556]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="0CDEEADED36DFF8CFBF73FBAFB1EF9E7" box="[1065,1178,1532,1556]" italics="true" pageId="7" pageNumber="470">C. longior</emphasis>
</taxonomicName>
, and the fixed finger is stouter in
<taxonomicName id="F9AA4D4FD36DFF8CFED83C66FEE8F9CB" authorityName="McLaughlin" authorityYear="2002" box="[262,364,1568,1592]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="0CDEEADED36DFF8CFED83C66FEE8F9CB" box="[262,364,1568,1592]" italics="true" pageId="7" pageNumber="470">C. foresti</emphasis>
</taxonomicName>
than
<taxonomicName id="F9AA4D4FD36DFF8CFE743C66FD98F9CB" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[426,540,1568,1592]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="0CDEEADED36DFF8CFE743C66FD98F9CB" box="[426,540,1568,1592]" italics="true" pageId="7" pageNumber="470">C. longior</emphasis>
</taxonomicName>
, particularly in males (
<figureCitation id="A6912A49D36DFF8CFCC03C59FC4FF9CB" box="[798,971,1567,1592]" captionStart="FIGURE 4" captionStartId="12.[151,250,1444,1466]" captionTargetBox="[297,1299,511,1375]" captionTargetId="figure@12.[151,1436,483,1422]" captionTargetPageId="12" captionText="FIGURE 4. Catapaguroides foresti McLaughlin, 2002: male (sl 2.0 mm), RUMF-ZC-6023 (IE173) (A–C), male (sl 1.5 mm), RUMF-ZC-6024 (IE184) (D); Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010: male (sl 2.1 mm), RUMF-ZC- 6028 (IE65) (E–H), female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107) (I, J). A, E, shield and ocular peduncles, dorsal view; B, distal part of right ocular peduncle, lateral view; C, D, G, I, right chela, dorsal view; F, left ocular peduncle, lateral view; H, J, carpus of right cheliped, dorsal view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627826/files/figure.png" pageId="7" pageNumber="470">Fig. 4C, D, G, I</figureCitation>
). The male sexual tube on the coxa of the right fifth pereopod is also much stouter and shorter in
<taxonomicName id="F9AA4D4FD36DFF8CFCD93C02FCEAF9AF" authorityName="McLaughlin" authorityYear="2002" box="[775,878,1604,1628]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="0CDEEADED36DFF8CFCD93C02FCEAF9AF" box="[775,878,1604,1628]" italics="true" pageId="7" pageNumber="470">C. foresti</emphasis>
</taxonomicName>
than
<taxonomicName id="F9AA4D4FD36DFF8CFC703C02FBA4F9AF" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[942,1056,1604,1628]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="0CDEEADED36DFF8CFC703C02FBA4F9AF" box="[942,1056,1604,1628]" italics="true" pageId="7" pageNumber="470">C. longior</emphasis>
</taxonomicName>
. (cf.
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: fig. 1H;
<bibRefCitation id="5A3B4B3DD36DFF8CFF493C2EFE1FF973" author="Osawa, M. &amp; Takeda, M." box="[151,411,1639,1664]" pageId="7" pageNumber="470" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
: fig. 9F, as
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<emphasis id="0CDEEADED36DFF8CFDC43C2EFDF0F973" box="[538,628,1640,1664]" italics="true" pageId="7" pageNumber="470">C. kasei</emphasis>
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; Komai
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. 2010: fig.
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).
</paragraph>
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153
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<emphasis id="0CDEEADED36DFF8CFF493AD1FE50FF42" bold="true" box="[151,468,151,177]" italics="true" pageId="7" pageNumber="470">Catapaguroides iejimensis</emphasis>
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(
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)
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<emphasis id="0CDEEADED36DFF8CFF493B44FE2DFEEB" box="[151,425,258,280]" italics="true" pageId="7" pageNumber="470">Catapaguroides iejimensis</emphasis>
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: 1113
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, figs. 5–7 (
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locality: Ie
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, Ryukyu Islands).—
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: 44
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, fig. 7D.
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<emphasis id="0CDEEADED36DFF8CFF493B22FE05FE8E" bold="true" box="[151,385,356,381]" pageId="7" pageNumber="470">Material examined.</emphasis>
<materialsCitation id="8EC23C91D36DFF8CFE573B22FEE5FE53" ID-GBIF-Occurrence="2465041895" collectingDate="2017-06-24" collectionCode="IE" collectionRegion="Ie Island, unnamed submarine cave" country="Japan" location="entrance" pageId="7" pageNumber="470" specimenCode="RUMF-ZC-6026" specimenCount="1" specimenCount-male="1">
<emphasis id="0CDEEADED36DFF8CFE573B22FCC4FE8E" bold="true" box="[393,832,356,381]" pageId="7" pageNumber="470">
Ie
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, unnamed submarine cave:
</emphasis>
entrance,
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<date id="4A14100CD36DFF8CFC683B22FBD4FE8F" box="[950,1104,356,380]" pageId="7" pageNumber="470" value="2017-06-24">24 June 2017</date>
</collectingDate>
,
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(sl
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), RUMF- ZC-6026 (
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<collectionCode id="58BBAE09D36DFF8CFED53BCEFEADFE53" box="[267,297,392,416]" pageId="7" pageNumber="470">IE</collectionCode>
177
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<emphasis id="0CDEEADED36DFF8CFF193BEAFEE4FE36" bold="true" box="[199,352,428,453]" pageId="7" pageNumber="470">Distribution.</emphasis>
Ryukyu Islands (Ie
<collectingCountry id="46BD765CD36DFF8CFD9B3BEAFD0DFE36" box="[581,649,428,453]" name="Iceland" pageId="7" pageNumber="470">Island</collectingCountry>
and Kume Island) in southwestern
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;
<quantity id="F9529B29D36DFF8CFBB43BEAFB4AFE37" box="[1130,1230,428,453]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="7" pageNumber="470" unit="m" value="22.5" valueMax="30.0" valueMin="15.0">15–30 m</quantity>
(
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).
</paragraph>
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<emphasis id="0CDEEADED36DFF8CFF193BB2FEAEFDFE" bold="true" box="[199,298,500,525]" pageId="7" pageNumber="470">Habitat.</emphasis>
Coral reefs; often found in submarine caves (
<bibRefCitation id="5A3B4B3DD36DFF8CFCF73BB2FBAFFDFF" author="Osawa, M. &amp; Okuno, J." box="[809,1067,500,525]" pageId="7" pageNumber="470" pagination="29 - 46" refId="ref12922" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12 (1), 29 - 46. https: // doi. org / 10.12782 / specdiv. 12.29" type="journal article" year="2007">Osawa &amp; Okuno 2007</bibRefCitation>
).
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<emphasis id="0CDEEADED36DFF8CFF193851FEBFFDC3" bold="true" box="[199,315,535,560]" pageId="7" pageNumber="470">Remarks.</emphasis>
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<emphasis id="0CDEEADED36DFF8CFE9C385EFDE9FDDC" box="[322,621,536,560]" italics="true" pageId="7" pageNumber="470">Catapaguroides iejimensis</emphasis>
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has been recorded primary from submarine caves in the Ryukyu Islands (
<bibRefCitation id="5A3B4B3DD36DFF8CFF40387AFE2DFDA7" author="Osawa, M. &amp; Takeda, M." box="[158,425,572,597]" pageId="7" pageNumber="470" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD36DFF8CFE6B387AFD3DFDA7" author="Osawa, M. &amp; Okuno, J." box="[437,697,572,597]" pageId="7" pageNumber="470" pagination="29 - 46" refId="ref12922" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12 (1), 29 - 46. https: // doi. org / 10.12782 / specdiv. 12.29" type="journal article" year="2007">Osawa &amp; Okuno 2007</bibRefCitation>
;
<tableCitation id="73280377D36DFF8CFD18387AFC98FDA7" box="[710,796,572,597]" captionStart="TABLE 2" captionStartId="6.[147,233,596,618]" captionText="TABLE 2. Submarine caves surveyed and hermit crab species collected in three islands of the Ryukyu Islands, southwestern Japan." pageId="7" pageNumber="470">Table 2</tableCitation>
), but the specimens hitherto obtained are rather limited. Only one specimen was collected during the present investigations.
</paragraph>
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174
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<mods:title id="F5E735135105B2DA12E17473EA1685A4">Submarine cave hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from three islands of the Ryukyu Islands, southwestern Japan</mods:title>
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<emphasis id="0CDEEADED36EFF8FFF493891FEEFFD02" bold="true" box="[151,363,727,753]" italics="true" pageId="4" pageNumber="467">Pseudopaguristes</emphasis>
<taxonomicName id="F9AA4D4FD36EFF8FFEAC3891FD15FD01" authority="Asakura, 2004" authorityName="Asakura" authorityYear="2004" box="[370,657,727,754]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="shidarai">
<emphasis id="0CDEEADED36EFF8FFEAC3891FE52FD02" bold="true" box="[370,470,727,753]" italics="true" pageId="4" pageNumber="467">shidarai</emphasis>
<bibRefCitation id="5A3B4B3DD36EFF8FFE02389EFD15FD01" author="Asakura, A." box="[476,657,728,754]" pageId="4" pageNumber="467" pagination="153 - 168" refId="ref9561" refString="Asakura, A. (2004) Pseudopaguristes shidarai, a new species of hermit crab (Crustacea: Decapoda: Diogenidae) from Japan, the fourth species of the genus. Proceedings of the Biological Society of Washington, 117 (2), 153 - 168." type="journal article" year="2004">Asakura, 2004</bibRefCitation>
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<paragraph id="3E1536CCD36EFF8FFF4938BCFF79FCE0" blockId="4.[151,657,727,787]" box="[151,253,762,787]" pageId="4" pageNumber="467">
(
<figureCitation id="A6912A49D36EFF8FFF4138BCFF70FCE0" box="[159,244,762,787]" captionStart="FIGURE 1" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@5.[151,1436,193,1705]" captionTargetPageId="5" captionText="FIGURE 1. Entire animal, dorsal view. A, Aniculus retipes Lewinsohn, 1982, male (sl 11.9 mm), RUMF-ZC-6006 (IE141); B, Paguristes jalur Morgan, 1992, male (sl 5.4 mm), RUMF-ZC-6012 (IE69); C, Pseudopaguristes shidarai Asakura, 2004, male (sl 2.6 mm), RUMF-ZC-6020 (HD2017-140); D, Catapaguroides foresti McLaughlin, 2002, male (sl 2.0 mm), RUMFZC-6023 (IE173); E, Catapaguroides iejimensis Osawa &amp; Takeda, 2004, male (sl 1.8 mm), RUMF-ZC-6026 (IE177); F, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, male (sl 2.1 mm), RUMF-ZC-6027 (IE60); G, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107)." httpUri="https://zenodo.org/record/2627820/files/figure.png" pageId="4" pageNumber="467">Fig. 1C</figureCitation>
)
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<emphasis id="0CDEEADED36EFF8FFF493904FECFFCAB" box="[151,331,834,856]" italics="true" pageId="4" pageNumber="467">Pseudopaguristes</emphasis>
<taxonomicName id="F9AA4D4FD36EFF8FFE8F3904FDF1FCAB" authority="Asakura, 2004: 153" authorityName="Asakura" authorityPageNumber="153" authorityYear="2004" box="[337,629,834,856]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="shidarai">
<emphasis id="0CDEEADED36EFF8FFE8F3904FE22FCAB" box="[337,422,834,856]" italics="true" pageId="4" pageNumber="467">shidarai</emphasis>
<treatmentCitation id="BF0B10DDD36EFF8FFE723904FDF1FCAB" author="Asakura, A." box="[428,629,834,856]" page="153" pageId="4" pageNumber="467" year="2004">
<bibRefCitation id="5A3B4B3DD36EFF8FFE723904FDC0FCAB" author="Asakura, A." box="[428,580,834,856]" pageId="4" pageNumber="467" pagination="153 - 168" refId="ref9561" refString="Asakura, A. (2004) Pseudopaguristes shidarai, a new species of hermit crab (Crustacea: Decapoda: Diogenidae) from Japan, the fourth species of the genus. Proceedings of the Biological Society of Washington, 117 (2), 153 - 168." type="journal article" year="2004">Asakura, 2004</bibRefCitation>
: 153
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, fig. 1–12 (type locality: Miyako Island, Ryukyu Islands)
</materialsCitation>
.
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<materialsCitation id="8EC23C91D36EFF8FFF4939C3FE29FC31" ID-GBIF-Occurrence="2465041902" collectingDate="2017-05-22" collectionCode="HD" country="Japan" location="main tube" pageId="4" pageNumber="467" specimenCode="RUMF- ZC-6020" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa Island, Hedo Dome cave">
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Material examined.
<collectingRegion id="FC6EF82ED36EFF8FFE5739C3FCA3FC6D" box="[393,807,901,926]" pageId="4" pageNumber="467">Okinawa Island, Hedo Dome cave</collectingRegion>
:
</emphasis>
<location id="3B756017D36EFF8FFCEB39C3FC2CFC6D" LSID="urn:lsid:plazi:treatment:B60387DAD36EFF8FFF493891FEC5FB4D:3B756017D36EFF8FFCEB39C3FC2CFC6D" box="[821,936,901,926]" country="Japan" name="main tube" pageId="4" pageNumber="467" stateProvince="Okinawa Island, Hedo Dome cave">main tube</location>
,
<date id="4A14100CD36EFF8FFC6A39C0FBCBFC6D" box="[948,1103,902,926]" pageId="4" pageNumber="467" value="2017-05-22">
<collectingDate id="5A50E9E4D36EFF8FFC6A39C0FBCBFC6D" box="[948,1103,902,926]" pageId="4" pageNumber="467" value="2017-05-22">22 May 2017</collectingDate>
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,
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(sl
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(HD2017-140)
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.
<materialsCitation id="8EC23C91D36EFF8FFE6739EFFD86FC15" ID-GBIF-Occurrence="2465041906" collectingDate="2017-09-01" collectionCode="SH" country="Japan" location="entrance" pageId="4" pageNumber="467" specimenCode="RUMF-ZC-6021" specimenCount="1" specimenCount-female="1" stateProvince="Shimoji Island, Akuma-no-Yakata cave">
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<collectingRegion id="FC6EF82ED36EFF8FFE6739EFFC11FC31" box="[441,917,937,962]" pageId="4" pageNumber="467">Shimoji Island, Akuma-no-Yakata cave</collectingRegion>
:
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<location id="3B756017D36EFF8FFC7C39ECFB80FC32" LSID="urn:lsid:plazi:treatment:B60387DAD36EFF8FFF493891FEC5FB4D:3B756017D36EFF8FFC7C39ECFB80FC32" box="[930,1028,938,961]" country="Japan" name="entrance" pageId="4" pageNumber="467" stateProvince="Shimoji Island, Akuma-no-Yakata cave">entrance</location>
,
<date id="4A14100CD36EFF8FFBD139ECFB64FC31" box="[1039,1248,937,962]" pageId="4" pageNumber="467" value="2017-09-01">
<collectingDate id="5A50E9E4D36EFF8FFBD139ECFB64FC31" box="[1039,1248,937,962]" pageId="4" pageNumber="467" value="2017-09-01">1 September 2017</collectingDate>
</date>
,
<specimenCount id="28ACFD45D36EFF8FFB3539ECFAC9FC31" box="[1259,1357,937,962]" pageId="4" pageNumber="467" type="female">1 female</specimenCount>
(sl
<quantity id="F9529B29D36EFF8FFAA739ECFF43FC15" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="467" unit="mm" value="2.3">2.3 mm</quantity>
),
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(SH105)
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.
</paragraph>
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<paragraph id="3E1536CCD36EFF8FFF1939B7FBD3FBDD" blockId="4.[151,1436,901,1214]" pageId="4" pageNumber="467">
<materialsCitation id="8EC23C91D36EFF8FFF1939B7FEF1FBDE" ID-GBIF-Occurrence="2465041908" country="Japan" pageId="4" pageNumber="467" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island, Kume Island, Miyako Island, And Shimoji Island)">
<emphasis id="0CDEEADED36EFF8FFF1939B7FEE4FBF9" bold="true" box="[199,352,1009,1034]" pageId="4" pageNumber="467">Distribution.</emphasis>
Ryukyu Islands (Okinawa Island, Kume Island, Miyako Island, and Shimoji Island) in southwestern Japan
</materialsCitation>
;
<materialsCitation id="8EC23C91D36EFF8FFE5E3E50FD17FBDD" ID-GBIF-Occurrence="2465041904" box="[384,659,1045,1070]" country="Japan" pageId="4" pageNumber="467" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island, Kume Island, Miyako Island, And Shimoji Island)">
<quantity id="F9529B29D36EFF8FFE5E3E50FE67FBDE" box="[384,483,1046,1070]" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="4" pageNumber="467" unit="m" value="30.0" valueMax="40.0" valueMin="20.0">20–40 m</quantity>
(
<bibRefCitation id="5A3B4B3DD36EFF8FFE2F3E53FD17FBDD" author="Asakura, A." box="[497,659,1045,1070]" pageId="4" pageNumber="467" pagination="153 - 168" refId="ref9561" refString="Asakura, A. (2004) Pseudopaguristes shidarai, a new species of hermit crab (Crustacea: Decapoda: Diogenidae) from Japan, the fourth species of the genus. Proceedings of the Biological Society of Washington, 117 (2), 153 - 168." type="journal article" year="2004">Asakura 2004</bibRefCitation>
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;
<materialsCitation id="8EC23C91D36EFF8FFD403E53FC23FBDD" ID-GBIF-Occurrence="2465041901" box="[670,935,1045,1070]" country="Japan" pageId="4" pageNumber="467" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island, Kume Island, Miyako Island, And Shimoji Island)">
<bibRefCitation id="5A3B4B3DD36EFF8FFD403E53FC23FBDD" author="Rahayu, D. L. &amp; Osawa, M." box="[670,935,1045,1070]" pageId="4" pageNumber="467" pagination="176 - 190" refId="ref13115" refString="Rahayu, D. L. &amp; Osawa, M. (2012) Hermit crabs (Crustacea: Decapoda: Diogenidae) from the KUMEJIMA 2009 Expedition, Japan. Zootaxa, 3367, 176 - 190." type="journal article" year="2012">Rahayu &amp; Osawa 2012</bibRefCitation>
</materialsCitation>
;
<materialsCitation id="8EC23C91D36EFF8FFC6C3E50FBD7FBDD" ID-GBIF-Occurrence="2465041917" box="[946,1107,1045,1070]" country="Japan" pageId="4" pageNumber="467" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island, Kume Island, Miyako Island, And Shimoji Island)">present study)</materialsCitation>
.
</paragraph>
<paragraph id="3E1536CCD36EFF8FFF193E7FFC14FBA1" blockId="4.[151,1436,901,1214]" box="[199,912,1081,1106]" pageId="4" pageNumber="467">
<emphasis id="0CDEEADED36EFF8FFF193E7FFEAEFBA1" bold="true" box="[199,298,1081,1106]" pageId="4" pageNumber="467">Habitat.</emphasis>
Coral reefs; fore-reef slopes (
<bibRefCitation id="5A3B4B3DD36EFF8FFDA63E7FFC00FBA1" author="Rahayu, D. L. &amp; Osawa, M." box="[632,900,1081,1106]" pageId="4" pageNumber="467" pagination="176 - 190" refId="ref13115" refString="Rahayu, D. L. &amp; Osawa, M. (2012) Hermit crabs (Crustacea: Decapoda: Diogenidae) from the KUMEJIMA 2009 Expedition, Japan. Zootaxa, 3367, 176 - 190." type="journal article" year="2012">Rahayu &amp; Osawa 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D36EFF8FFF193E1BFEC5FB4D" pageId="4" pageNumber="467" type="discussion">
<paragraph id="3E1536CCD36EFF8FFF193E1BFEC5FB4D" blockId="4.[151,1436,901,1214]" pageId="4" pageNumber="467">
<emphasis id="0CDEEADED36EFF8FFF193E1BFEBFFB85" bold="true" box="[199,315,1117,1142]" pageId="4" pageNumber="467">Remarks.</emphasis>
In the Ryukyu Islands,
<taxonomicName id="F9AA4D4FD36EFF8FFD9C3E18FD33FB86" authorityName="Asakura" authorityYear="2004" box="[578,695,1117,1141]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="shidarai">
<emphasis id="0CDEEADED36EFF8FFD9C3E18FD33FB86" box="[578,695,1117,1141]" italics="true" pageId="4" pageNumber="467">P. shidarai</emphasis>
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has been recorded from cryptic environments of the shallow coral reefs such as small crevices and horizontal holes, and submarine caves (
<bibRefCitation id="5A3B4B3DD36EFF8FFC133EC7FB5FFB69" author="Rahayu, D. L. &amp; Osawa, M." box="[973,1243,1153,1178]" pageId="4" pageNumber="467" pagination="176 - 190" refId="ref13115" refString="Rahayu, D. L. &amp; Osawa, M. (2012) Hermit crabs (Crustacea: Decapoda: Diogenidae) from the KUMEJIMA 2009 Expedition, Japan. Zootaxa, 3367, 176 - 190." type="journal article" year="2012">Rahayu &amp; Osawa 2012</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD36EFF8FFB363EC4FF57FB4D" author="Osawa, M. &amp; Fujita, Y." pageId="4" pageNumber="467" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita 2016</bibRefCitation>
;
<tableCitation id="73280377D36EFF8FFF033EE3FEB7FB4D" box="[221,307,1189,1214]" captionStart="TABLE 2" captionStartId="6.[147,233,596,618]" captionText="TABLE 2. Submarine caves surveyed and hermit crab species collected in three islands of the Ryukyu Islands, southwestern Japan." pageId="4" pageNumber="467">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
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236
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<mods:title id="FBE0D43C52DD0DCE9DFE1292E4639A0C">Submarine cave hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from three islands of the Ryukyu Islands, southwestern Japan</mods:title>
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<paragraph id="3E1536CCD36EFF8FFF493F10FD10FA83" blockId="4.[151,660,1366,1426]" box="[151,660,1366,1392]" pageId="4" pageNumber="467">
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<taxonomicName id="F9AA4D4FD36EFF8FFF493F10FD10FA83" ID-CoL="11d53467-c2ce-47c1-9dd2-1fed683ea53a" authority="McLaughlin, 2002" authorityName="McLaughlin" authorityYear="2002" box="[151,660,1366,1392]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="foresti">
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<emphasis id="0CDEEADED36EFF8FFF493F10FE2DFA83" bold="true" box="[151,425,1366,1392]" italics="true" pageId="4" pageNumber="467">Catapaguroides foresti</emphasis>
McLaughlin, 2002
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(
<figureCitation id="A6912A49D36EFF8FFF413F3FFE86FA61" box="[159,258,1401,1426]" captionStart="FIGURE 1" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@5.[151,1436,193,1705]" captionTargetPageId="5" captionText="FIGURE 1. Entire animal, dorsal view. A, Aniculus retipes Lewinsohn, 1982, male (sl 11.9 mm), RUMF-ZC-6006 (IE141); B, Paguristes jalur Morgan, 1992, male (sl 5.4 mm), RUMF-ZC-6012 (IE69); C, Pseudopaguristes shidarai Asakura, 2004, male (sl 2.6 mm), RUMF-ZC-6020 (HD2017-140); D, Catapaguroides foresti McLaughlin, 2002, male (sl 2.0 mm), RUMFZC-6023 (IE173); E, Catapaguroides iejimensis Osawa &amp; Takeda, 2004, male (sl 1.8 mm), RUMF-ZC-6026 (IE177); F, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, male (sl 2.1 mm), RUMF-ZC-6027 (IE60); G, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107)." httpUri="https://zenodo.org/record/2627820/files/figure.png" pageId="4" pageNumber="467">Figs. 1D</figureCitation>
,
<figureCitation id="A6912A49D36EFF8FFED03F3CFE98FA61" box="[270,284,1402,1426]" captionStart="FIGURE 4" captionStartId="12.[151,250,1444,1466]" captionTargetBox="[297,1299,511,1375]" captionTargetId="figure@12.[151,1436,483,1422]" captionTargetPageId="12" captionText="FIGURE 4. Catapaguroides foresti McLaughlin, 2002: male (sl 2.0 mm), RUMF-ZC-6023 (IE173) (A–C), male (sl 1.5 mm), RUMF-ZC-6024 (IE184) (D); Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010: male (sl 2.1 mm), RUMF-ZC- 6028 (IE65) (E–H), female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107) (I, J). A, E, shield and ocular peduncles, dorsal view; B, distal part of right ocular peduncle, lateral view; C, D, G, I, right chela, dorsal view; F, left ocular peduncle, lateral view; H, J, carpus of right cheliped, dorsal view. Setae omitted from all illustrations. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/2627826/files/figure.png" pageId="4" pageNumber="467">4</figureCitation>
A–D)
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<emphasis id="0CDEEADED36EFF8FFF493F87FEFBFA24" box="[151,383,1473,1495]" italics="true" pageId="4" pageNumber="467">Catapaguroides foresti</emphasis>
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<bibRefCitation id="5A3B4B3DD36EFF8FFE5B3F87FDD6FA24" author="McLaughlin, P. A." box="[389,594,1473,1495]" pageId="4" pageNumber="467" pagination="495 - 504" refId="ref11437" refString="McLaughlin, P. A. (2002 b) A new species of the genus Catapaguroides (Decapoda: Anomura: Paguroidea: Paguridae) from Guam, Micronesia. Crustaceana, 75 (3 - 4), 495 - 504. https: // doi. org / 10.1163 / 156854002760095543" type="journal article" year="2002">McLaughlin, 2002b</bibRefCitation>
: 499
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, fig. 1 (
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locality:
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).—Komai
<emphasis id="0CDEEADED36EFF8FFBD13F84FBB8FA24" box="[1039,1084,1473,1495]" italics="true" pageId="4" pageNumber="467">et al</emphasis>
. 2010: 38, fig. 5B.
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<emphasis id="0CDEEADED36EFF8FFF493FA6FEF7FA05" box="[151,371,1504,1526]" italics="true" pageId="4" pageNumber="467">Catapaguroides kasei</emphasis>
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<bibRefCitation id="5A3B4B3DD36EFF8FFEA43FA6FDEFFA05" author="Osawa, M. &amp; Takeda, M." box="[378,619,1504,1526]" pageId="4" pageNumber="467" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda, 2004</bibRefCitation>
: 1121
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, figs. 8–10 (
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locality: Ie
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, Ryukyu Islands).
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<emphasis id="0CDEEADED36EFF8FFF493C64FE06F9C8" bold="true" box="[151,386,1570,1595]" pageId="4" pageNumber="467">Material examined.</emphasis>
<materialsCitation id="8EC23C91D36EFF8FFE523C64FDACF9AC" ID-GBIF-Occurrence="2465041909" collectingDate="2016-09-09" collectionCode="IE" collectionRegion="Ie Island, unname submarine cave" country="Japan" location="main tube" pageId="4" pageNumber="467" specimenCode="RUMF-ZC-6022" specimenCount="1" specimenCount-male="1">
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Ie
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, unnamed submarine cave:
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main tube,
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<date id="4A14100CD36EFF8FFC0B3C65FB2DF9C8" box="[981,1193,1570,1595]" pageId="4" pageNumber="467" value="2016-09-09">9 September 2016</date>
</collectingDate>
,
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(sl
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), RUMF-ZC-6022 (
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<collectionCode id="58BBAE09D36EFF8FFEB83C00FE07F9AC" box="[358,387,1606,1631]" pageId="4" pageNumber="467">IE</collectionCode>
65
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#, addition);
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<materialsCitation id="8EC23C91D36EFF8FFDEC3C01FADAF9AC" ID-GBIF-Occurrence="2465041914" box="[562,1374,1606,1631]" collectingDate="2017-06-24" collectionCode="IE" collectionRegion="Ie Island, unnamed submarine cave" country="Japan" location="entrance" pageId="4" pageNumber="467" specimenCode="RUMF-ZC-6023" specimenCount="1" specimenCount-male="1">
entrance,
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<date id="4A14100CD36EFF8FFD7F3C01FCBAF9AC" box="[673,830,1607,1631]" pageId="4" pageNumber="467" value="2017-06-24">24 June 2017</date>
</collectingDate>
,
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(sl 2.0 mm), RUMF-ZC-6023 (
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<collectionCode id="58BBAE09D36EFF8FFAD63C01FAA2F9AC" box="[1288,1318,1607,1631]" pageId="4" pageNumber="467">IE</collectionCode>
173
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);
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<materialsCitation id="8EC23C91D36EFF8FFAB93C00FCF3F970" ID-GBIF-Occurrence="2465041898" collectingDate="2017-06-24" collectionCode="IE" collectionRegion="Ie Island, unnamed submarine cave" country="Japan" location="right hall" pageId="4" pageNumber="467" specimenCode="RUMF-ZC-6024" specimenCount="1" specimenCount-male="1">
right hall,
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<date id="4A14100CD36EFF8FFF113C2DFEE3F970" box="[207,359,1643,1667]" pageId="4" pageNumber="467" value="2017-06-24">24 June 2017</date>
</collectingDate>
,
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(sl
<quantity id="F9529B29D36EFF8FFE393C2CFDC5F970" box="[487,577,1642,1667]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="467" unit="mm" value="1.5">1.5 mm</quantity>
), RUMF-ZC-6024 (
<specimenCode id="6E0C9EB7D36EFF8FFCFF3C2DFCEEF970" box="[801,874,1643,1667]" collectionCode="IE" country="Mexico" name="Cepario de Hongos del Instituto de Ecologia" pageId="4" pageNumber="467">
<collectionCode id="58BBAE09D36EFF8FFCFF3C2DFCBBF970" box="[801,831,1643,1667]" pageId="4" pageNumber="467">IE</collectionCode>
184
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).
</materialsCitation>
<materialsCitation id="8EC23C91D36EFF8FFCA33C2CFC0BF954" ID-GBIF-Occurrence="2465041905" collectingDate="2017-05-22" collectionRegion="Okinawa Island, Hedo Dome cave" country="Japan" location="silty room" pageId="4" pageNumber="467" specimenCode="RUMF-ZC-6025" specimenCount="1" specimenCount-male="1">
<emphasis id="0CDEEADED36EFF8FFCA33C2CFA9FF970" bold="true" box="[893,1307,1642,1667]" pageId="4" pageNumber="467">
<location id="3B756017D36EFF8FFCA33C2CFBB9F970" LSID="urn:lsid:plazi:treatment:B60387DAD36EFF8FFF493F10FB87F88C:3B756017D36EFF8FFCA33C2CFBB9F970" box="[893,1085,1642,1667]" country="Iceland" municipality="Hedo Dome cave" name="Okinawa Island" pageId="4" pageNumber="467">Okinawa Island</location>
,
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:
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silty room,
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<date id="4A14100CD36EFF8FFF493CC9FEABF954" box="[151,303,1679,1703]" pageId="4" pageNumber="467" value="2017-05-22">22 May 2017</date>
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,
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(sl
<quantity id="F9529B29D36EFF8FFE713CC9FD8EF954" box="[431,522,1679,1703]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="467" unit="mm" value="1.1">1.1 mm</quantity>
), RUMF-ZC-6025 (
<collectionCode id="58BBAE09D36EFF8FFD373CC9FC90F954" box="[745,788,1679,1703]" pageId="4" pageNumber="467">HD</collectionCode>
2017-108).
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</paragraph>
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<emphasis id="0CDEEADED36EFF8FFF193CF4FEE4F938" bold="true" box="[199,352,1714,1739]" pageId="4" pageNumber="467">Distribution.</emphasis>
<collectingCountry id="46BD765CD36EFF8FFEB73CF5FE2AF938" box="[361,430,1715,1739]" name="Guam" pageId="4" pageNumber="467">Guam</collectingCountry>
and southern
<collectingCountry id="46BD765CD36EFF8FFD8E3CF5FD14F939" box="[592,656,1715,1738]" name="Japan" pageId="4" pageNumber="467">Japan</collectingCountry>
(
<collectingRegion id="FC6EF82ED36EFF8FFD7E3CF4FCE1F938" box="[672,869,1714,1739]" country="Japan" name="Kochi" pageId="4" pageNumber="467">Kochi Prefecture</collectingRegion>
,
<collectingRegion id="FC6EF82ED36EFF8FFCAC3CF4FC53F938" box="[882,983,1714,1739]" country="Japan" name="Okinawa" pageId="4" pageNumber="467">Okinawa</collectingRegion>
Island, and Ie
<collectingCountry id="46BD765CD36EFF8FFB5D3CF4FB48F938" box="[1155,1228,1714,1739]" name="Iceland" pageId="4" pageNumber="467">Island</collectingCountry>
); subtidal to
<quantity id="F9529B29D36EFF8FFABF3CF5FA18F939" box="[1377,1436,1715,1739]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="467" unit="m" value="30.0">30 m</quantity>
(Komai
<emphasis id="0CDEEADED36EFF8FFF2D3C91FEA0F91D" box="[243,292,1750,1774]" italics="true" pageId="4" pageNumber="467">et al</emphasis>
. 2010; present study).
</paragraph>
<paragraph id="3E1536CCD36EFF8FFF193CBCFB81F8E0" blockId="4.[151,1436,1570,1919]" box="[199,1029,1786,1811]" pageId="4" pageNumber="467">
<emphasis id="0CDEEADED36EFF8FFF193CBCFEAEF8E0" bold="true" box="[199,298,1786,1811]" pageId="4" pageNumber="467">Habitat.</emphasis>
Coral reefs; often found in submarine caves (Komai
<emphasis id="0CDEEADED36EFF8FFCA03CBDFC2BF8E0" box="[894,943,1787,1811]" italics="true" pageId="4" pageNumber="467">et al</emphasis>
. 2010).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D36EFF8FFF193D58FB87F88C" pageId="4" pageNumber="467" type="discussion">
<paragraph id="3E1536CCD36EFF8FFF193D58FB87F88C" blockId="4.[151,1436,1570,1919]" pageId="4" pageNumber="467">
<emphasis id="0CDEEADED36EFF8FFF193D58FEBFF8C4" bold="true" box="[199,315,1822,1847]" pageId="4" pageNumber="467">Remarks.</emphasis>
Komai
<emphasis id="0CDEEADED36EFF8FFE483D59FE43F8C5" box="[406,455,1822,1846]" italics="true" pageId="4" pageNumber="467">et al</emphasis>
. (2010) synonymized
<taxonomicName id="F9AA4D4FD36EFF8FFD183D59FBB5F8C4" authority="Osawa &amp; Takeda, 2004" authorityName="Osawa &amp; Takeda" authorityYear="2004" box="[710,1073,1822,1847]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="kasei">
<emphasis id="0CDEEADED36EFF8FFD183D59FCA5F8C5" box="[710,801,1822,1846]" italics="true" pageId="4" pageNumber="467">C. kasei</emphasis>
<bibRefCitation id="5A3B4B3DD36EFF8FFCF63D59FBB5F8C4" author="Osawa, M. &amp; Takeda, M." box="[808,1073,1822,1847]" pageId="4" pageNumber="467" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda, 2004</bibRefCitation>
</taxonomicName>
with
<taxonomicName id="F9AA4D4FD36EFF8FFBAC3D59FB5DF8C4" authorityName="McLaughlin" authorityYear="2002" box="[1138,1241,1823,1847]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="0CDEEADED36EFF8FFBAC3D59FB5DF8C4" box="[1138,1241,1823,1847]" italics="true" pageId="4" pageNumber="467">C. foresti</emphasis>
</taxonomicName>
.
<taxonomicName id="F9AA4D4FD36EFF8FFB363D58FF5AF8A8" authorityName="McLaughlin" authorityYear="2002" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="0CDEEADED36EFF8FFB363D58FF5AF8A8" italics="true" pageId="4" pageNumber="467">Catapaguroides foresti</emphasis>
</taxonomicName>
and
<taxonomicName id="F9AA4D4FD36EFF8FFEC83D05FE0DF8A8" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[278,393,1859,1883]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="0CDEEADED36EFF8FFEC83D05FE0DF8A8" box="[278,393,1859,1883]" italics="true" pageId="4" pageNumber="467">C. longior</emphasis>
</taxonomicName>
, both collected by the present investigations, have similar coloration in general (
<figureCitation id="A6912A49D36EFF8FFAFF3D04FF2AF88C" captionStart="FIGURE 1" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@5.[151,1436,193,1705]" captionTargetPageId="5" captionText="FIGURE 1. Entire animal, dorsal view. A, Aniculus retipes Lewinsohn, 1982, male (sl 11.9 mm), RUMF-ZC-6006 (IE141); B, Paguristes jalur Morgan, 1992, male (sl 5.4 mm), RUMF-ZC-6012 (IE69); C, Pseudopaguristes shidarai Asakura, 2004, male (sl 2.6 mm), RUMF-ZC-6020 (HD2017-140); D, Catapaguroides foresti McLaughlin, 2002, male (sl 2.0 mm), RUMFZC-6023 (IE173); E, Catapaguroides iejimensis Osawa &amp; Takeda, 2004, male (sl 1.8 mm), RUMF-ZC-6026 (IE177); F, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, male (sl 2.1 mm), RUMF-ZC-6027 (IE60); G, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107)." httpUri="https://zenodo.org/record/2627820/files/figure.png" pageId="4" pageNumber="467">Fig. 1D, F, G</figureCitation>
), but their morphologies are very different (see
<emphasis id="0CDEEADED36EFF8FFD1B3D20FCB6F88C" bold="true" box="[709,818,1894,1919]" pageId="4" pageNumber="467">Remarks</emphasis>
under
<taxonomicName id="F9AA4D4FD36EFF8FFC5E3D21FC76F88D" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[896,1010,1894,1918]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="0CDEEADED36EFF8FFC5E3D21FC76F88D" box="[896,1010,1894,1918]" italics="true" pageId="4" pageNumber="467">C. longior</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

54
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<bibRefCitation id="90DED4219A093815AAEFCD10C7129390" pageId="0" pageNumber="4567">Gaffigan et al. 2014</bibRefCitation>
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<emphasis id="0CDEB0106D70FF9D01DE8AA2FE1EA5CB" box="[224,438,1453,1477]" italics="true" pageId="24" pageNumber="25">Oecomys mamorae</emphasis>
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(
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)
<emphasis id="0CDEB0106D70FF9D00798AE3FE31A5F1" box="[327,409,1516,1535]" italics="true" pageId="24" pageNumber="25">mamorae</emphasis>
<materialsCitation id="8EC2665F6D70FF9D00A18AE3FDF1A5F1" ID-GBIF-Occurrence="3400112302" box="[415,601,1516,1535]" collectionCode="BMNH" country="Bolivia" county="Yungas" location="Mosetenes" municipality="upper Rio Mamore'" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Cochabamba" typeStatus="holotype">
<bibRefCitation id="5A3B11F36D70FF9D00A18AE3FDF1A5F1" author="Thomas, O." box="[415,601,1516,1535]" pageId="24" pageNumber="25" pagination="445 - 448" refId="ref17999" refString="Thomas, O. 1906 b. Notes on South American rodents. III. A new Oecomys and two new species of Holochilus. Annals and Magazine of Natural History (7) 18: 445 - 448." type="journal article" year="1906">Thomas, 1906b: 445</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(type locality—
<collectingCountry id="46BD2C926D70FF9D01CA890BFE9DA619" box="[244,309,1540,1559]" name="Bolivia" pageId="24" pageNumber="25">Bolivia</collectingCountry>
,
<collectingRegion id="FC6EA2E06D70FF9D007B890BFE68A619" box="[325,448,1540,1559]" country="Bolivia" name="Cochabamba" pageId="24" pageNumber="25">Cochabamba</collectingRegion>
,
<collectingCounty id="D774148E6D70FF9D00EF890BFDB1A619" box="[465,537,1540,1559]" pageId="24" pageNumber="25">Yungas</collectingCounty>
,
<collectingMunicipality id="DE71F6786D70FF9D0314890BFF43A620" pageId="24" pageNumber="25">upper Río Mamore´</collectingMunicipality>
,
<location id="3B753AD96D70FF9D01CB8914FEFFA620" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D70FF9D01CB8914FEFFA620" box="[245,343,1563,1582]" country="Bolivia" county="Yungas" municipality="upper Rio Mamore'" name="Mosetenes" pageId="24" pageNumber="25" stateProvince="Cochabamba">Mosetenes</location>
;
<typeStatus id="E111D2A06D70FF9D005F8914FE18A620" box="[353,432,1563,1582]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
—BMNH 1900.8.3.21).
</treatmentCitationGroup>
</paragraph>
<paragraph id="3E156C026D70FF9D01BE893AFF64A651" blockId="24.[128,654,1516,1753]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="1EBA4B2C6D70FF9D01BE893AFF64A651" pageId="24" pageNumber="25">
<treatmentCitation id="BF0B4A136D70FF9D01BE893AFE44A649" author="Osgood, W. H." box="[128,492,1588,1608]" page="206" pageId="24" pageNumber="25" year="1916">
<taxonomicName id="F9AA17816D70FF9D01BE893AFE44A649" authority=": Osgood, 1916: 206" authorityName="Osgood" authorityPageNumber="206" authorityYear="1916" box="[128,492,1588,1608]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D70FF9D01BE893AFE84A646" box="[128,300,1589,1608]" italics="true" pageId="24" pageNumber="25">Oecomys mamorae</emphasis>
:
<bibRefCitation id="5A3B11F36D70FF9D0002893BFE44A649" author="Osgood, W. H." box="[316,492,1588,1607]" pageId="24" pageNumber="25" pagination="199 - 216" refId="ref17335" refString="Osgood, W. H. 1916. Mammals of the Collins-Day South American Expedition. Field Museum of Natural History Publications Zoological Series 10 (14): 199 - 216." type="journal article" year="1916">Osgood, 1916: 206</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(name combination).
</treatmentCitationGroup>
</paragraph>
<paragraph id="3E156C026D70FF9D01BE896AFF64A69E" blockId="24.[128,654,1516,1753]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="1EBA4B2C6D70FF9D01BE896AFF64A69E" pageId="24" pageNumber="25">
<treatmentCitation id="BF0B4A136D70FF9D01BE896AFE5CA676" author="Ellerman, J. R." box="[128,500,1637,1656]" page="358" pageId="24" pageNumber="25" year="1941">
<taxonomicName id="F9AA17816D70FF9D01BE896AFE5CA676" authority=": Ellerman, 1941: 358" authorityName=": Ellerman" authorityPageNumber="358" authorityYear="1941" box="[128,500,1637,1656]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D70FF9D01BE896AFE9AA676" box="[128,306,1637,1656]" italics="true" pageId="24" pageNumber="25">Oryzomys mamorae</emphasis>
:
<bibRefCitation id="5A3B11F36D70FF9D0000896AFE5CA676" author="Ellerman, J. R." box="[318,500,1637,1656]" pageId="24" pageNumber="25" refId="ref16569" refString="Ellerman, J. R. 1941. The families and genera of living rodents. Vol. II. Family Muridae. London: British Museum (Natural History), 690 pp." type="book" year="1941">Ellerman, 1941: 358</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(name combination).
</treatmentCitationGroup>
</paragraph>
<paragraph id="3E156C026D70FF9D01BE8999FF4EA6D7" blockId="24.[128,654,1516,1753]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="1EBA4B2C6D70FF9D01BE8999FF4EA6D7" pageId="24" pageNumber="25">
<treatmentCitation id="BF0B4A136D70FF9D01BE8999FD26A6A7" author="Cabrera, A." box="[128,654,1686,1705]" page="405" pageId="24" pageNumber="25" year="1961">
<taxonomicName id="F9AA17816D70FF9D01BE8999FD26A6A7" authority="mamorae: Cabrera, 1961: 405" authorityName="Cabrera" authorityPageNumber="405" authorityYear="1961" box="[128,654,1686,1705]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="subSpecies" species="mamorae" subSpecies="mamorae">
<emphasis id="0CDEB0106D70FF9D01BE8999FE00A6A7" box="[128,424,1686,1705]" italics="true" pageId="24" pageNumber="25">Oryzomys mamorae mamorae</emphasis>
:
<bibRefCitation id="5A3B11F36D70FF9D00FD8999FD26A6A7" author="Cabrera, A." box="[451,654,1686,1705]" pageId="24" pageNumber="25" pagination="309 - 732" refId="ref16249" refString="Cabrera, A. 1961. Catalogo de los mamiferos de America del Sur. Revista del Museo Argentino de Ciencias Naturales '' Bernardino Rivadavia' ' Ciencias Zoologicas 4 (2): v - xxii, 309 - 732." type="journal article" year="1961">Cabrera, 1961: 405</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(retained as species, ranked as nominate subspecies).
</treatmentCitationGroup>
</paragraph>
<paragraph id="3E156C026D70FF9D03838B26FB3AA45A" blockId="24.[701,1228,1065,1158]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="1EBA4B2C6D70FF9D03838B26FB3AA45A" pageId="24" pageNumber="25">
<treatmentCitation id="BF0B4A136D70FF9D03838B26FB39A432" author="Hershkovitz, P." box="[701,1169,1065,1084]" page="559" pageId="24" pageNumber="25" year="1960">
<taxonomicName id="F9AA17816D70FF9D03838B26FB39A432" authority=": Hershkovitz, 1960: 559" authorityName="Hershkovitz" authorityPageNumber="559" authorityYear="1960" box="[701,1169,1065,1084]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="subSpecies" species="concolor" subSpecies="roberti">
<emphasis id="0CDEB0106D70FF9D03838B26FC07A432" box="[701,943,1065,1084]" italics="true" pageId="24" pageNumber="25">Oryzomys concolor roberti</emphasis>
:
<bibRefCitation id="5A3B11F36D70FF9D02828B26FB39A432" author="Hershkovitz, P." box="[956,1169,1065,1084]" pageId="24" pageNumber="25" pagination="513 - 568" refId="ref16974" refString="Hershkovitz, P. 1960. Mammals of northern Colombia, preliminary report no. 8: Arboreal rice rats, a systematic revision of the subgenus Oecomys, genus Oryzomys. Proceedings of the United States National Museum 110: 513 - 568." type="journal article" year="1960">Hershkovitz, 1960: 559</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(part,
<taxonomicName id="F9AA17816D70FF9D03E68B4DFC82A45B" authorityName=": Ellerman" authorityYear="1941" box="[728,810,1090,1109]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D70FF9D03E68B4DFC82A45B" box="[728,810,1090,1109]" italics="true" pageId="24" pageNumber="25">mamorae</emphasis>
</taxonomicName>
allocated as full subspecific synonym).
</treatmentCitationGroup>
</paragraph>
<paragraph id="3E156C026D70FF9D03838B54FBA7A488" blockId="24.[701,1228,1065,1158]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="1EBA4B2C6D70FF9D03838B54FBA7A488" pageId="24" pageNumber="25">
<treatmentCitation id="BF0B4A136D70FF9D03838B54FB22A463" author="Anderson, S." box="[701,1162,1114,1134]" page="12" pageId="24" pageNumber="25" year="1985">
<taxonomicName id="F9AA17816D70FF9D03838B54FB22A463" authority=": Anderson, 1985: 12" authorityName="Anderson" authorityPageNumber="12" authorityYear="1985" box="[701,1162,1114,1134]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="subSpecies" species="concolor" subSpecies="roberti">
<emphasis id="0CDEB0106D70FF9D03838B54FC1BA460" box="[701,947,1115,1134]" italics="true" pageId="24" pageNumber="25">Oecomys concolor roberti</emphasis>
:
<bibRefCitation id="5A3B11F36D70FF9D02F68B55FB22A463" author="Anderson, S." box="[968,1162,1114,1133]" pageId="24" pageNumber="25" pagination="5 - 16" refId="ref16167" refString="Anderson, S. 1985. Lista preliminar de mamiferos bolivianos. Academia Nacional de Ciencias de Bolivia Cuadernos 65: 5 - 16." type="journal article" year="1985">Anderson, 1985: 12</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(part,
<emphasis id="0CDEB0106D70FF9D03E68B7CFC9AA488" box="[728,818,1139,1158]" italics="true" pageId="24" pageNumber="25">marmorae</emphasis>
listed as full synonym).
</treatmentCitationGroup>
</paragraph>
<paragraph id="3E156C026D70FF9D03E68BACFBADA699" blockId="24.[701,1228,1187,1747]" pageId="24" pageNumber="25">
EMENDED DIAGNOSIS: A species of
<taxonomicName id="F9AA17816D70FF9D05B78BACFD41A4D7" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="0CDEB0106D70FF9D05B78BACFD41A4D7" italics="true" pageId="24" pageNumber="25">Oecomys</emphasis>
</taxonomicName>
(
<taxonomicName id="F9AA17816D70FF9D03C18BCFFC06A4D6" ID-CoL="93N26" authorityName="Wagner" authorityYear="1843" box="[767,942,1216,1240]" class="Mammalia" family="Cricetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="subFamily" subFamily="Sigmodontinae">Sigmodontinae</taxonomicName>
:
<taxonomicName id="F9AA17816D70FF9D02818BCFFBEEA4D6" ID-CoL="L3X" box="[959,1094,1216,1240]" class="Mammalia" family="Cricetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="tribe" tribe="Oryzomyini">Oryzomyini</taxonomicName>
) characterized by a combination of medium-large size (HBL
<emphasis id="0CDEB0106D70FF9D022E8BF0FC8EA51A" bold="true" box="[784,806,1279,1300]" italics="true" pageId="24" pageNumber="25">&lt;</emphasis>
<quantity id="F952C1E76D70FF9D02088BF3FC62A51A" box="[822,970,1276,1300]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="24" pageNumber="25" unit="mm" value="140.0" valueMax="150.0" valueMin="130.0">130–150 mm</quantity>
, HFL
<emphasis id="0CDEB0106D70FF9D05198BF0FB95A51A" bold="true" box="[1063,1085,1279,1300]" italics="true" pageId="24" pageNumber="25">&lt;</emphasis>
<quantity id="F952C1E76D70FF9D05708BF3FB60A51A" box="[1102,1224,1276,1300]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="24" pageNumber="25" unit="mm" value="27.0" valueMax="29.0" valueMin="25.0">25–29 mm</quantity>
, ONL
<emphasis id="0CDEB0106D70FF9D022F8A13FC8FA53F" bold="true" box="[785,807,1308,1329]" italics="true" pageId="24" pageNumber="25">&lt;</emphasis>
<quantity id="F952C1E76D70FF9D027E8A15FC12A53C" box="[832,954,1306,1330]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="24" pageNumber="25" unit="mm" value="32.0" valueMax="33.0" valueMin="31.0">31–33 mm</quantity>
), relatively long tail (TL
<emphasis id="0CDEB0106D70FF9D03C98A35FCA5A541" bold="true" box="[759,781,1338,1359]" italics="true" pageId="24" pageNumber="25">&lt;</emphasis>
<quantity id="F952C1E76D70FF9D02258A38FC07A541" box="[795,943,1335,1359]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="24" pageNumber="25" unit="mm" value="160.0" valueMax="170.0" valueMin="150.0">150–170 mm</quantity>
), narrow interorbit and weakly developed supraorbital ridges, long and narrow incisive foramina, relatively short bony palate and simple posterolateral palatal pits, alisphenoid struts typically absent, and a derived carotid circulatory pattern (skull lacking squamosal-alisphenoid groove, sphenofrontal foramen, and posterolateral groove on the parapterygoid plate; posterior opening to the alisphenoid canal compressed; stapedial foramen absent; groove dorsally crossing the parapterygoid plate present).
</paragraph>
</subSubSection>
<subSubSection id="76B03F896D70FF9C03E68992FD94A14A" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="distribution">
<paragraph id="3E156C026D70FF9C03E68992FD94A14A" blockId="24.[701,1228,1187,1747]" lastBlockId="25.[88,614,211,1753]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<materialsCitation id="8EC2665F6D70FF9C03E68992FF07A14A" ID-GBIF-Occurrence="3400112361" country="Bolivia" county="Known" elevation="2100" lastPageId="25" lastPageNumber="26" location="Subhumid" municipality="Chaco" pageId="24" pageNumber="25" specimenCount="1">
DISTRIBUTION:
<location id="3B753AD96D70FF9D02AE8992FBADA6BB" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D70FF9D02AE8992FBADA6BB" box="[912,1029,1693,1717]" country="Bolivia" county="Known" municipality="Chaco" name="Subhumid" pageId="24" pageNumber="25">Subhumid</location>
and gallery forests in savanna and
<collectingMunicipality id="DE71F6786D70FF9D028989B4FBA8A6DD" box="[951,1024,1723,1747]" pageId="24" pageNumber="25">Chaco</collectingMunicipality>
zones of central and eastern
<collectingCountry id="46BD2C926D71FF9C01CC8FDCFEEDA0E5" box="[242,325,211,235]" name="Bolivia" pageId="25" pageNumber="26">Bolivia</collectingCountry>
, contiguous westcentral
<collectingCountry id="46BD2C926D71FF9C01668FFEFF36A107" box="[88,158,241,265]" name="Brazil" pageId="25" pageNumber="26">Brazil</collectingCountry>
, and northern and eastern
<collectingCountry id="46BD2C926D71FF9C00C48FFEFDCDA107" box="[506,613,241,265]" name="Paraguay" pageId="25" pageNumber="26">Paraguay</collectingCountry>
(fig. 6).
<collectingCounty id="D774148E6D71FF9C01888E01FEA3A128" box="[182,267,270,294]" pageId="25" pageNumber="26">Known</collectingCounty>
elevational range sea level to
<quantity id="F952C1E76D71FF9C01668E23FF07A14A" box="[88,175,300,324]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="25" pageNumber="26" unit="m" value="2100.0">
<elevation id="B5878B316D71FF9C01668E23FF07A14A" box="[88,175,300,324]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="25" pageNumber="26" unit="m" value="2100.0">2100 m</elevation>
</quantity>
</materialsCitation>
, most localities within
<quantity id="F952C1E76D71FF9C00878E23FD90A14A" box="[441,568,300,324]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="25" pageNumber="26" unit="m" value="350.0" valueMax="500.0" valueMin="200.0">200–500 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="76B03F896D71FF9F014C8E45FDC9A393" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="discussion">
<paragraph id="3E156C026D71FF9C014C8E45FEE6A444" blockId="25.[88,614,211,1753]" pageId="25" pageNumber="26">
REMARKS: The cranium of the
<typeStatus id="E111D2A06D71FF9C00D58E45FDB3A16C" box="[491,539,330,354]" pageId="25" pageNumber="26">type</typeStatus>
specimen of
<taxonomicName id="F9AA17816D71FF9C01918E67FEBEA18E" authorityName=": Ellerman" authorityYear="1941" box="[175,278,360,384]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C01918E67FEBEA18E" box="[175,278,360,384]" italics="true" pageId="25" pageNumber="26">mamorae</emphasis>
</taxonomicName>
(BMNH 1900.8.3.21), an old adult female, exhibits the essential traits of a derived carotid pattern (no sphenofrontal foramen, stapedial foramen minute), yet it does retain shallow traces of the squamosalalisphenoid groove on the inner walls of the braincase. As in most specimens of
<taxonomicName id="F9AA17816D71FF9C03748D14FF17A25F" authorityName=": Ellerman" authorityYear="1941" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C03748D14FF17A25F" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
that we have examined (
<tableCitation id="732859B96D71FF9C03398D37FDFFA25E" box="[519,599,568,592]" captionStart="TABLE 5" captionStartId="18.[340,422,209,231]" captionTargetBox="[128,621,351,787]" captionText="TABLE 5 Occurrence of Alisphenoid Struts (2/2 5 struts absent both sides; 2/+ 5 strut present on one side; +/+ 5 struts present both sides.)" httpUri="http://table.plazi.org/id/6AD53C8A6D7AFF97006A8FDEFD25A139" pageId="25" pageNumber="26" tableUuid="6AD53C8A6D7AFF97006A8FDEFD25A139">table 5</tableCitation>
), the
<typeStatus id="E111D2A06D71FF9C01B38D59FF15A260" box="[141,189,598,622]" pageId="25" pageNumber="26">type</typeStatus>
lacks alisphenoid struts.
<bibRefCitation id="5A3B11F36D71FF9C03378D59FF4BA282" author="Thomas, O." pageId="25" pageNumber="26" pagination="445 - 448" refId="ref17999" refString="Thomas, O. 1906 b. Notes on South American rodents. III. A new Oecomys and two new species of Holochilus. Annals and Magazine of Natural History (7) 18: 445 - 448." type="journal article" year="1906">Thomas (1906b: 446)</bibRefCitation>
critically contrasted his new form
<taxonomicName id="F9AA17816D71FF9C01668D9DFF4CA2A4" authorityName=": Ellerman" authorityYear="1941" box="[88,228,658,682]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C01668D9DFF4CA2A4" box="[88,228,658,682]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
to
<taxonomicName id="F9AA17816D71FF9C00338D9DFE3CA2A4" authorityName="Thomas" authorityYear="1899" box="[269,404,658,682]" class="Mammalia" family="Muridae" genus="Rhipidomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="marmosurus">
<emphasis id="0CDEB0106D71FF9C00338D9DFE3CA2A4" box="[269,404,658,682]" italics="true" pageId="25" pageNumber="26">marmosurus</emphasis>
</taxonomicName>
, here allocated as a junior synonym of
<taxonomicName id="F9AA17816D71FF9C007C8DBFFE6BA2C6" authorityName=": Musser and Carleton" authorityYear="1993" box="[322,451,688,712]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="concolor">
<emphasis id="0CDEB0106D71FF9C007C8DBFFE6BA2C6" box="[322,451,688,712]" italics="true" pageId="25" pageNumber="26">O. concolor</emphasis>
</taxonomicName>
, and captured the consistent proportional differences that we have observed in the larger samples of each species now available: ‘‘the distinction of this animal [
<taxonomicName id="F9AA17816D71FF9C018A8C28FE96A331" authorityName=": Ellerman" authorityYear="1941" box="[180,318,807,831]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C018A8C28FE96A331" box="[180,318,807,831]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
], which may be separated from its only equal in size,
<taxonomicName id="F9AA17816D71FF9C009D8C4AFF39A374" authorityName="Thomas" authorityYear="1899" class="Mammalia" family="Muridae" genus="Rhipidomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="marmosurus" subGenus="Oe.">
<emphasis id="0CDEB0106D71FF9C009D8C4AFE17A353" box="[419,447,837,861]" italics="true" pageId="25" pageNumber="26">O.</emphasis>
(
<emphasis id="0CDEB0106D71FF9C00ED8C4AFE53A353" box="[467,507,837,861]" italics="true" pageId="25" pageNumber="26">Oe.</emphasis>
)
<emphasis id="0CDEB0106D71FF9C03318C4AFF39A374" italics="true" pageId="25" pageNumber="26">marmosurus</emphasis>
</taxonomicName>
, by its narrower interorbital region, less developed orbital ledges, and larger palatal foramina.’’ The interorbital constriction (IOB) and long incisive foramina (LIF) correlate strongly with the first canonical variate that discriminates specimens of
<taxonomicName id="F9AA17816D71FF9C00A88CF8FD8BA401" authorityName=": Ellerman" authorityYear="1941" box="[406,547,1015,1039]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C00A88CF8FD8BA401" box="[406,547,1015,1039]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
from those of
<taxonomicName id="F9AA17816D71FF9C01F18B1AFEF3A423" authorityName=": Musser and Carleton" authorityYear="1993" box="[207,347,1045,1069]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="concolor">
<emphasis id="0CDEB0106D71FF9C01F18B1AFEF3A423" box="[207,347,1045,1069]" italics="true" pageId="25" pageNumber="26">O. concolor</emphasis>
</taxonomicName>
and
<taxonomicName id="F9AA17816D71FF9C00978B1AFDCEA423" box="[425,614,1045,1069]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="sydandersoni">
<emphasis id="0CDEB0106D71FF9C00978B1AFDCEA423" box="[425,614,1045,1069]" italics="true" pageId="25" pageNumber="26">O. sydandersoni</emphasis>
</taxonomicName>
(fig. 3;
<tableCitation id="732859B96D71FF9C01958B3DFEADA444" box="[171,261,1074,1098]" captionStart="TABLE 3" captionStartId="11.[300,382,209,231]" captionTargetBox="[88,594,404,1015]" captionText="TABLE 3 Results of Three-group Discriminant Function Analysis of Adult Oecomys sp. novum, O. concolor, and O. mamorae (Based on 17 log-transformed craniodental variables; see Materials and Methods and fig. 3.)" httpUri="http://table.plazi.org/id/6AD53C8A6D63FF8E00128FDEFDF4A162" pageId="25" pageNumber="26" tableUuid="6AD53C8A6D63FF8E00128FDEFDF4A162">tables 3</tableCitation>
,
<tableCitation id="732859B96D71FF9C002A8B3DFE8CA444" box="[276,292,1074,1098]" captionStart="TABLE 4" captionStartId="13.[587,669,209,231]" captionTargetBox="[88,1153,351,1705]" captionText="TABLE 4 External and Craniodental Measurements for the Type Series of O. sydandersoni, New Species, and Samples of O. concolor and O. mamorae (Sample statistics include the mean, ± 1 standard deviation, and the observed range)." httpUri="http://table.plazi.org/id/6AD53C8A6D65FF8803758FDEFB92A139" pageId="25" pageNumber="26" tableUuid="6AD53C8A6D65FF8803758FDEFB92A139">4</tableCitation>
,
<tableCitation id="732859B96D71FF9C000F8B3DFEE9A444" box="[305,321,1074,1098]" captionStart="TABLE 7" captionStartId="24.[626,708,209,231]" captionTargetBox="[128,1163,320,945]" captionText="TABLE 7 External and Craniodental Measurements (in mm) of the Type Specimens of Hesperomys concolor Wagner, Rhipidomys marmosurus Thomas, and Oryzomys mamorae Thomas" httpUri="http://table.plazi.org/id/6AD53C8A6D70FF9D034C8FDEFC52A113" pageId="25" pageNumber="26" tableUuid="6AD53C8A6D70FF9D034C8FDEFC52A113">7</tableCitation>
).
</paragraph>
<paragraph id="3E156C026D71FF9C014C8B5FFC8EA14D" blockId="25.[88,614,211,1753]" lastBlockId="25.[662,1188,211,912]" pageId="25" pageNumber="26">
Following Thomas,
<taxonomicName id="F9AA17816D71FF9C005F8B5EFE47A467" authorityName=": Ellerman" authorityYear="1941" box="[353,495,1105,1129]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C005F8B5EFE47A467" box="[353,495,1105,1129]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
had been retained as a distinctive species by most authors until
<bibRefCitation id="5A3B11F36D71FF9C01C08B83FE75A4AA" author="Hershkovitz, P." box="[254,477,1164,1188]" pageId="25" pageNumber="26" pagination="513 - 568" refId="ref16974" refString="Hershkovitz, P. 1960. Mammals of northern Colombia, preliminary report no. 8: Arboreal rice rats, a systematic revision of the subgenus Oecomys, genus Oryzomys. Proceedings of the United States National Museum 110: 513 - 568." type="journal article" year="1960">Hershkovitz (1960)</bibRefCitation>
submerged it within his expansive view of
<taxonomicName id="F9AA17816D71FF9C00828BA5FD96A4CC" authorityName=": Musser and Carleton" authorityYear="1993" box="[444,574,1194,1218]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="concolor">
<emphasis id="0CDEB0106D71FF9C00828BA5FD96A4CC" box="[444,574,1194,1218]" italics="true" pageId="25" pageNumber="26">O. concolor</emphasis>
</taxonomicName>
, as a full synonym of
<emphasis id="0CDEB0106D71FF9C00138BC7FE11A4EE" box="[301,441,1224,1248]" italics="true" pageId="25" pageNumber="26">
O. c.
<taxonomicName id="F9AA17816D71FF9C00508BC7FE11A4EE" baseAuthorityName="Thomas" baseAuthorityYear="1904" box="[366,441,1224,1248]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="roberti">roberti</taxonomicName>
</emphasis>
. Hershkovitz’s definition of the subspecies confused two distinctive species,
<taxonomicName id="F9AA17816D71FF9C00068A0CFDCEA515" authority="Thomas (1903)" authorityName="Thomas" authorityYear="1903" box="[312,614,1283,1307]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="roberti">
<emphasis id="0CDEB0106D71FF9C00068A0CFE02A515" box="[312,426,1283,1307]" italics="true" pageId="25" pageNumber="26">O. roberti</emphasis>
Thomas (1903)
</taxonomicName>
and
<taxonomicName id="F9AA17816D71FF9C01B38A2EFE4DA537" authority="Thomas (1906 b)" authorityName="Thomas" authorityYear="1906" box="[141,485,1313,1337]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C01B38A2EFEB3A537" box="[141,283,1313,1337]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
<bibRefCitation id="5A3B11F36D71FF9C001B8A2EFE4DA537" author="Thomas, O." box="[293,485,1313,1337]" pageId="25" pageNumber="26" pagination="445 - 448" refId="ref17999" refString="Thomas, O. 1906 b. Notes on South American rodents. III. A new Oecomys and two new species of Holochilus. Annals and Magazine of Natural History (7) 18: 445 - 448." type="journal article" year="1906">Thomas (1906b)</bibRefCitation>
</taxonomicName>
, which are easily separable from one another and from
<taxonomicName id="F9AA17816D71FF9C03748A30FE99A57A" authority="(Wagner)" authorityName="Mouth" baseAuthorityName="Wagner" baseAuthorityYear="1845" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="concolor">
<emphasis id="0CDEB0106D71FF9C03748A30FF1EA57B" italics="true" pageId="25" pageNumber="26">O. concolor</emphasis>
(Wagner)
</taxonomicName>
proper. Examples of
<taxonomicName id="F9AA17816D71FF9C03748A52FF0BA59C" authorityName="Thomas" authorityYear="1903" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="roberti">
<emphasis id="0CDEB0106D71FF9C03748A52FF0BA59C" italics="true" pageId="25" pageNumber="26">O. roberti</emphasis>
</taxonomicName>
, like most other
<taxonomicName id="F9AA17816D71FF9C00448A75FE77A59C" box="[378,479,1402,1426]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis id="0CDEB0106D71FF9C00448A75FE77A59C" box="[378,479,1402,1426]" italics="true" pageId="25" pageNumber="26">Oecomys</emphasis>
</taxonomicName>
, retain the complete carotid arterial pattern in contrast to the derived condition found in
<taxonomicName id="F9AA17816D71FF9C00EE8AB9FDF7A5C0" authorityName="Thomas" authorityYear="1906" box="[464,607,1462,1486]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C00EE8AB9FDF7A5C0" box="[464,607,1462,1486]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
. Although the dorsal pelage of both species tends to be pale ochraceous-tawny and the venter mostly white, the cover hairs are typically shorter in
<taxonomicName id="F9AA17816D71FF9C007D8922FE1EA64B" authorityName="Thomas" authorityYear="1903" box="[323,438,1581,1605]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="roberti">
<emphasis id="0CDEB0106D71FF9C007D8922FE1EA64B" box="[323,438,1581,1605]" italics="true" pageId="25" pageNumber="26">O. roberti</emphasis>
</taxonomicName>
(
<quantity id="F952C1E76D71FF9C00F48922FD80A64B" box="[458,552,1581,1605]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="25" pageNumber="26" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
over the rump) than
<taxonomicName id="F9AA17816D71FF9C001B8944FE1FA66D" authorityName="Thomas" authorityYear="1906" box="[293,439,1611,1635]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C001B8944FE1FA66D" box="[293,439,1611,1635]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
(
<quantity id="F952C1E76D71FF9C00F18945FD99A66C" box="[463,561,1610,1634]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="25" pageNumber="26" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
). In addition, the supraorbital shelf and posterolateral palatal pits are more pronounced in
<taxonomicName id="F9AA17816D71FF9C03748989FF0BA6B2" authorityName="Thomas" authorityYear="1903" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="roberti">
<emphasis id="0CDEB0106D71FF9C03748989FF0BA6B2" italics="true" pageId="25" pageNumber="26">O. roberti</emphasis>
</taxonomicName>
. Specimens of
<taxonomicName id="F9AA17816D71FF9C005989ABFE75A6B2" authorityName="Thomas" authorityYear="1903" box="[359,477,1700,1724]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="roberti">
<emphasis id="0CDEB0106D71FF9C005989ABFE75A6B2" box="[359,477,1700,1724]" italics="true" pageId="25" pageNumber="26">O. roberti</emphasis>
</taxonomicName>
have been recorded in northern and easternmost
<collectingCountry id="46BD2C926D71FF9C032B89CEFDCDA6D7" box="[533,613,1729,1753]" name="Bolivia" pageId="25" pageNumber="26">Bolivia</collectingCountry>
(
<bibRefCitation id="5A3B11F36D71FF9C03A38FDCFCF1A0E5" author="Anderson, S." box="[669,857,211,235]" pageId="25" pageNumber="26" pagination="1 - 652" refId="ref16192" refString="Anderson, S. 1997. Mammals of Bolivia, taxonomy and distribution. Bulletin of the American Museum of Natural History 231: 1 - 652." type="journal article" year="1997">Anderson, 1997</bibRefCitation>
;
<bibRefCitation id="5A3B11F36D71FF9C02578FDCFBCAA0E5" author="Emmons, L. H. &amp; V. Chavez &amp; N. Rocha &amp; B. Phillips &amp; I. Phillips &amp; M. J. Swarner" box="[873,1122,211,235]" pageId="25" pageNumber="26" pagination="23 - 46" refId="ref16634" refString="Emmons, L. H., V. Chavez, N. Rocha, B. Phillips, I. Phillips, L. F. d. Aguila, and M. J. Swarner. 2006. The non-flying mammals of Noel Kempff Mercado National Park (Bolivia). Revista Boliviana de Ecologia y Conservacion Ambiental 19: 23 - 46." type="journal article" year="2006">Emmons et al., 2006</bibRefCitation>
), just peripheral to the range of
<taxonomicName id="F9AA17816D71FF9C02818FFFFBE1A106" authorityName="Thomas" authorityYear="1906" box="[959,1097,240,264]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C02818FFFFBE1A106" box="[959,1097,240,264]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
(fig. 6), but the two species have yet to be discovered in sympatry.
</paragraph>
<paragraph id="3E156C026D71FF9C038E8E46FBC9A381" blockId="25.[662,1188,211,912]" pageId="25" pageNumber="26">
Specimens of
<taxonomicName id="F9AA17816D71FF9C02718E46FC1BA16F" box="[847,947,329,353]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis id="0CDEB0106D71FF9C02718E46FC1BA16F" box="[847,947,329,353]" italics="true" pageId="25" pageNumber="26">Oecomys</emphasis>
</taxonomicName>
have been recovered from owl pellets in northeastern
<collectingCountry id="46BD2C926D71FF9C05158E68FB37A171" box="[1067,1183,359,383]" name="Argentina" pageId="25" pageNumber="26">Argentina</collectingCountry>
, from the provinces of
<collectingRegion id="FC6EA2E06D71FF9C02928E8BFC5DA192" box="[940,1013,388,412]" country="Argentina" name="Chaco" pageId="25" pageNumber="26">Chaco</collectingRegion>
(
<bibRefCitation id="5A3B11F36D71FF9C05358E8BFC99A1B7" author="Massoia, E. &amp; A. Fornes" pageId="25" pageNumber="26" pagination="319 - 324" refId="ref17019" refString="Massoia, E., and A. Fornes. 1965. Oryzomys (Oecomys) Thomas, 1906, Nuevo subgenero de cricetidos para la Republica Argentina (Rodentia). Physis Seccion C 25: 319 - 324." type="journal article" year="1965">Massoia and Fornes, 1965</bibRefCitation>
, as
<taxonomicName id="F9AA17816D71FF9C02568EADFC47A1B4" authorityName="Mouth" baseAuthorityName="Wagner" baseAuthorityYear="1845" box="[872,1007,418,442]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="concolor">
<emphasis id="0CDEB0106D71FF9C02568EADFC47A1B4" box="[872,1007,418,442]" italics="true" pageId="25" pageNumber="26">O. concolor</emphasis>
</taxonomicName>
) and
<collectingCountry id="46BD2C926D71FF9C05038EAEFB0BA1B7" box="[1085,1187,417,441]" name="Taiwan" pageId="25" pageNumber="26">Formosa</collectingCountry>
(
<bibRefCitation id="5A3B11F36D71FF9C03A38EB0FBF9A1D9" author="Pardinas, U. F. J. &amp; P. Ramirez-Llorens" box="[669,1105,446,471]" pageId="25" pageNumber="26" pagination="103 - 107" refId="ref17368" refString="Pardinas, U. F. J., and P. Ramirez-Llorens. 2005. The genus Oecomys (Rodentia, Sigmodontinae) in Argentina. Mammalia 69: 103 - 107." type="journal article" year="2005">Pardiñas and Ramírez-Llorens, 2005</bibRefCitation>
, as
<emphasis id="0CDEB0106D71FF9C05B68EB0FB0CA1D9" box="[1160,1188,447,471]" italics="true" pageId="25" pageNumber="26">O.</emphasis>
sp.). By geographic proximity alone, these samples are plausibly referable to
<taxonomicName id="F9AA17816D71FF9C052A8EF5FB34A21C" authorityName="Thomas" authorityYear="1906" box="[1044,1180,506,530]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C052A8EF5FB34A21C" box="[1044,1180,506,530]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
, but certain measurements reported for the fragmented crania of
<collectingCountry id="46BD2C926D71FF9C02B58D3BFB8DA242" box="[907,1061,564,588]" name="Argentina" pageId="25" pageNumber="26">the Argentine</collectingCountry>
form seem to run larger than
<taxonomicName id="F9AA17816D71FF9C02448D5DFBA0A264" authorityName="Thomas" authorityYear="1906" box="[890,1032,594,618]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C02448D5DFBA0A264" box="[890,1032,594,618]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
proper from
<collectingCountry id="46BD2C926D71FF9C03A88D60FD4EA289" box="[662,742,623,647]" name="Bolivia" pageId="25" pageNumber="26">Bolivia</collectingCountry>
(notably IOB, LIF, BIF, LD, WM1—
<bibRefCitation id="5A3B11F36D71FF9C03A88D83FB93A2AA" author="Pardinas, U. F. J. &amp; P. Ramirez-Llorens" box="[662,1083,652,677]" pageId="25" pageNumber="26" pagination="103 - 107" refId="ref17368" refString="Pardinas, U. F. J., and P. Ramirez-Llorens. 2005. The genus Oecomys (Rodentia, Sigmodontinae) in Argentina. Mammalia 69: 103 - 107." type="journal article" year="2005">Pardiñas and Ramírez-Llorens, 2005</bibRefCitation>
: table 1). Critical review of variation within nominal
<taxonomicName id="F9AA17816D71FF9C05B68DA5FD54A2D1" authorityName="Thomas" authorityYear="1906" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C05B68DA5FD54A2D1" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
and comparisons with
<collectingCountry id="46BD2C926D71FF9C05368DC8FB0CA2D1" box="[1032,1188,711,735]" name="Argentina" pageId="25" pageNumber="26">the Argentine</collectingCountry>
populations are required to verify their specific assignment. Such a review should include those populations documented in eastern
<collectingCountry id="46BD2C926D71FF9C050C8C10FB08A339" box="[1074,1184,799,823]" name="Paraguay" pageId="25" pageNumber="26">Paraguay</collectingCountry>
, to the east of the
<collectingCountry id="46BD2C926D71FF9C025C8C32FC65A35B" box="[866,973,829,853]" name="Paraguay" pageId="25" pageNumber="26">Paraguay</collectingCountry>
River, reported as
<taxonomicName id="F9AA17816D71FF9C03A88C55FC12A37C" authority="(Myers, 1982)" baseAuthorityName="Myers" baseAuthorityYear="1982" box="[662,954,858,882]" class="Mammalia" family="Muridae" genus="Oecomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="concolor">
<emphasis id="0CDEB0106D71FF9C03A88C55FCBEA37C" box="[662,790,858,882]" italics="true" pageId="25" pageNumber="26">O. concolor</emphasis>
(
<bibRefCitation id="5A3B11F36D71FF9C02188C55FC1BA37C" author="Myers, P." box="[806,947,858,882]" pageId="25" pageNumber="26" pagination="85 - 93" refId="ref17304" refString="Myers, P. 1982. Origins and affinities of the mammal fauna of Paraguay. University of Pittsburgh Pymatuning Laboratory of Ecology Special Publication 6: 85 - 93." type="journal article" year="1982">Myers, 1982</bibRefCitation>
)
</taxonomicName>
and here referred to
<taxonomicName id="F9AA17816D71FF9C03A88C77FCB7A39E" authorityName="Thomas" authorityYear="1906" box="[662,799,888,912]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C03A88C77FCB7A39E" box="[662,799,888,912]" italics="true" pageId="25" pageNumber="26">O. mamorae</emphasis>
</taxonomicName>
based on our examinations.
</paragraph>
<paragraph id="3E156C026D71FF9F038E8CADFDC9A393" blockId="25.[662,1188,930,1754]" lastBlockId="26.[128,654,210,925]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<materialsCitation id="8EC2665F6D71FF9C038E8CADFBCAA3DD" ID-GBIF-Occurrence="3400112342" collectionCode="FMNH" collectorName="Baures" country="Bolivia" location="Beni" pageId="25" pageNumber="26" specimenCode="FMNH 117063-117066" specimenCount="1" stateProvince="Beni">
SPECIMENS EXAMINED: 128, as follows.
<emphasis id="0CDEB0106D71FF9C054D8CADFD1BA3DD" bold="true" pageId="25" pageNumber="26">
<collectingCountry id="46BD2C926D71FF9C054D8CADFD1BA3DD" name="Bolivia" pageId="25" pageNumber="26">Bolivia</collectingCountry>
</emphasis>
—
<emphasis id="0CDEB0106D71FF9C03F48CB2FD5FA3DC" box="[714,759,957,978]" italics="true" pageId="25" pageNumber="26">
<collectingRegion id="FC6EA2E06D71FF9C03F48CB2FD5FA3DC" box="[714,759,957,978]" country="Bolivia" name="Beni" pageId="25" pageNumber="26">Beni</collectingRegion>
</emphasis>
,
<collectorName id="935F09D46D71FF9C02368CB2FCE6A3DD" box="[776,846,957,979]" pageId="25" pageNumber="26">Baures</collectorName>
(
<specimenCode id="6E0CC4796D71FF9C025E8CB2FBF3A3DD" box="[864,1115,957,979]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26" type="Museum">FMNH 117063–117066</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C054E8CB2FBBAA3E0" ID-GBIF-Occurrence="3400112340" collectionCode="AMNH" country="Bolivia" location="Boca del Rio Ibare" pageId="25" pageNumber="26" specimenCode="AMNH 211749" specimenCount="1" stateProvince="Beni">
<location id="3B753AD96D71FF9C054E8CB2FC93A3E0" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C054E8CB2FC93A3E0" country="Bolivia" name="Boca del Rio Ibare" pageId="25" pageNumber="26" stateProvince="Beni">Boca del Río Ibaré</location>
(
<specimenCode id="6E0CC4796D71FF9C02668CD7FBA4A3E0" box="[856,1036,984,1006]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH 211749</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C05158CD8FC93A42D" ID-GBIF-Occurrence="3400112311" collectionCode="AMNH" country="Bolivia" location="Rio Ibare'" pageId="25" pageNumber="26" specimenCode="AMNH 211718, 211719, 211750-211752" specimenCount="1" stateProvince="Beni">
<location id="3B753AD96D71FF9C05158CD8FB08A3E0" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C05158CD8FB08A3E0" box="[1067,1184,983,1006]" country="Bolivia" name="Rio Ibare'" pageId="25" pageNumber="26" stateProvince="Beni">Río Ibare´</location>
,
<quantity id="F952C1E76D71FF9C03A88CFDFD7EA406" box="[662,726,1010,1032]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="25" pageNumber="26" unit="km" value="26.0">26 km</quantity>
from mouth (
<specimenCode id="6E0CC4796D71FF9C02B38CFDFB97A406" box="[909,1087,1010,1032]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH 211718</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C056B8CFDFB08A406" box="[1109,1184,1010,1032]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">211719</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C03A88B02FC9EA42D" box="[662,822,1037,1059]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">211750–211752</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C02648B02FC3AA430" ID-GBIF-Occurrence="3400112306" collectionCode="AMNH" country="Bolivia" elevation="200" location="Yacuma Prov." municipality="Busurucucu" pageId="25" pageNumber="26" specimenCode="AMNH 263491" specimenCount="1" stateProvince="Beni">
<collectingMunicipality id="DE71F6786D71FF9C02648B02FC7BA42D" box="[858,979,1037,1059]" pageId="25" pageNumber="26">Busurucucú</collectingMunicipality>
,
<location id="3B753AD96D71FF9C02CB8B02FB08A42D" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C02CB8B02FB08A42D" box="[1013,1184,1037,1059]" country="Bolivia" municipality="Busurucucu" name="Yacuma Prov." pageId="25" pageNumber="26" stateProvince="Beni">Yacuma Prov.</location>
,
<quantity id="F952C1E76D71FF9C03A88B27FD7DA430" box="[662,725,1064,1086]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="m" value="200.0">
<elevation id="B5878B316D71FF9C03A88B27FD7DA430" box="[662,725,1064,1086]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="m" value="200.0">200 m</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D71FF9C03DA8B27FC23A430" box="[740,907,1064,1086]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH 263491</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C02A08B27FC18A456" ID-GBIF-Occurrence="3400112356" collectionCode="USNM" collectorName="S San Ramon" country="Bolivia" location="Lucuma" pageId="25" pageNumber="26" specimenCode="USNM 460430" specimenCount="1" stateProvince="Beni">
<location id="3B753AD96D71FF9C02A08B27FC5DA430" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C02A08B27FC5DA430" box="[926,1013,1064,1086]" country="Bolivia" name="Lucuma" pageId="25" pageNumber="26" stateProvince="Beni">Lúcuma</location>
, 6 leagues
<collectorName id="935F09D46D71FF9C05568B27FD4AA456" pageId="25" pageNumber="26">S San Ramón</collectorName>
(
<specimenCode id="6E0CC4796D71FF9C03C28B4DFC01A456" box="[764,937,1090,1112]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26" type="Museum">USNM 460430</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C02F88B4DFC94A47D" ID-GBIF-Occurrence="3400112362" collectionCode="FMNH" country="Bolivia" location="Magdalena" pageId="25" pageNumber="26" specimenCode="FMNH 117055-117058" specimenCount="1" stateProvince="Beni">
<location id="3B753AD96D71FF9C02F88B4DFB91A456" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C02F88B4DFB91A456" box="[966,1081,1090,1112]" country="Bolivia" name="Magdalena" pageId="25" pageNumber="26" stateProvince="Beni">Magdalena</location>
(
<specimenCode id="6E0CC4796D71FF9C056A8B4DFC9DA47D" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26" type="Museum">FMNH 117055–117058</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C02718B52FC8EA480" ID-GBIF-Occurrence="3400112322" collectionCode="NHMS" collectorName="El Consuelo" country="Bolivia" elevation="196" location="Rio" pageId="25" pageNumber="26" specimenCode="NHMS 153" specimenCount="1" stateProvince="Beni">
<location id="3B753AD96D71FF9C02718B52FCDDA47D" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C02718B52FCDDA47D" box="[847,885,1117,1139]" country="Bolivia" name="Rio" pageId="25" pageNumber="26" stateProvince="Beni">Río</location>
<collectingRegion id="FC6EA2E06D71FF9C02BA8B52FC1DA47D" box="[900,949,1117,1139]" country="Bolivia" name="Beni" pageId="25" pageNumber="26">Beni</collectingRegion>
,
<collectorName id="935F09D46D71FF9C02F68B52FBF9A47D" box="[968,1105,1117,1139]" pageId="25" pageNumber="26">El Consuelo</collectorName>
,
<quantity id="F952C1E76D71FF9C055A8B52FB0CA47D" box="[1124,1188,1117,1139]" metricMagnitude="2" metricUnit="m" metricValue="1.96" pageId="25" pageNumber="26" unit="m" value="196.0">
<elevation id="B5878B316D71FF9C055A8B52FB0CA47D" box="[1124,1188,1117,1139]" metricMagnitude="2" metricUnit="m" metricValue="1.96" pageId="25" pageNumber="26" unit="m" value="196.0">196 m</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D71FF9C03A38B77FC88A480" box="[669,800,1144,1166]" collectionCode="NHMS" pageId="25" pageNumber="26">NHMS 153</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C02048B77FCE7A4CD" ID-GBIF-Occurrence="3400112304" collectionCode="AMNH" country="Bolivia" county="Rio" elevation="226" location="Mamore River" municipality="Puerto" pageId="25" pageNumber="26" specimenCode="AMNH 211753" specimenCount="1" stateProvince="Beni">
<collectingCounty id="D774148E6D71FF9C02048B77FCC8A480" box="[826,864,1144,1166]" pageId="25" pageNumber="26">Río</collectingCounty>
<collectingRegion id="FC6EA2E06D71FF9C02518B77FC08A480" box="[879,928,1144,1166]" country="Bolivia" name="Beni" pageId="25" pageNumber="26">Beni</collectingRegion>
,
<collectingMunicipality id="DE71F6786D71FF9C028D8B77FC5FA480" box="[947,1015,1144,1166]" pageId="25" pageNumber="26">Puerto</collectingMunicipality>
Salinas,
<quantity id="F952C1E76D71FF9C055A8B77FB0CA480" box="[1124,1188,1144,1166]" metricMagnitude="2" metricUnit="m" metricValue="2.26" pageId="25" pageNumber="26" unit="m" value="226.0">
<elevation id="B5878B316D71FF9C055A8B77FB0CA480" box="[1124,1188,1144,1166]" metricMagnitude="2" metricUnit="m" metricValue="2.26" pageId="25" pageNumber="26" unit="m" value="226.0">226 m</elevation>
</quantity>
(NHMS 62–64, 74–77, 79, 80);
<location id="3B753AD96D71FF9C053D8B9CFB0BA4A7" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C053D8B9CFB0BA4A7" box="[1027,1187,1171,1193]" country="Bolivia" county="Rio" municipality="Puerto" name="Mamore River" pageId="25" pageNumber="26" stateProvince="Beni">Mamoré River</location>
(
<specimenCode id="6E0CC4796D71FF9C03A38BA2FCE0A4CD" box="[669,840,1197,1219]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH 211753</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C02618BA2FC0FA4D0" ID-GBIF-Occurrence="3400112345" collectionCode="AMNH" country="Bolivia" elevation="240" location="Marban" municipality="Rio Mamore'" pageId="25" pageNumber="26" specimenCode="AMNH 129254, 129255" specimenCount="1" stateProvince="Beni">
<collectingMunicipality id="DE71F6786D71FF9C02618BA2FC45A4CD" box="[863,1005,1197,1219]" pageId="25" pageNumber="26">Río Mamore´</collectingMunicipality>
,
<location id="3B753AD96D71FF9C02C38BA2FBFBA4CD" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C02C38BA2FBFBA4CD" box="[1021,1107,1197,1219]" country="Bolivia" municipality="Rio Mamore'" name="Marban" pageId="25" pageNumber="26" stateProvince="Beni">Marbán</location>
,
<quantity id="F952C1E76D71FF9C055A8BA2FB0CA4CD" box="[1124,1188,1197,1219]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="25" pageNumber="26" unit="m" value="240.0">
<elevation id="B5878B316D71FF9C055A8BA2FB0CA4CD" box="[1124,1188,1197,1219]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="25" pageNumber="26" unit="m" value="240.0">240 m</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D71FF9C03A38BC7FCEDA4D0" box="[669,837,1224,1246]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH 129254</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C026A8BC7FC08A4D0" box="[852,928,1224,1246]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">129255</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C02888BC7FD40A51D" ID-GBIF-Occurrence="3400112351" collectionCode="FMNH, USNM" country="Bolivia" location="San" municipality="Mamore'" pageId="25" pageNumber="26" specimenCode="FMNH 117053, 117059, 117067, 117068, USNM 391302" specimenCount="1" stateProvince="Beni">
<collectingMunicipality id="DE71F6786D71FF9C02888BC7FBB9A4D0" box="[950,1041,1224,1246]" pageId="25" pageNumber="26">Mamore´</collectingMunicipality>
,
<location id="3B753AD96D71FF9C051E8BC7FBEEA4D0" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C051E8BC7FBEEA4D0" box="[1056,1094,1224,1246]" country="Bolivia" municipality="Mamore'" name="San" pageId="25" pageNumber="26" stateProvince="Beni">San</location>
Joaquín (
<specimenCode id="6E0CC4796D71FF9C03A38BECFCE9A4F7" box="[669,833,1251,1273]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26" type="Museum">FMNH 117053</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C02718BECFC32A4F7" box="[847,922,1251,1273]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26" type="Museum">117059</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C02998BECFC5AA4F7" box="[935,1010,1251,1273]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26" type="Museum">117067</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C02C18BECFBE2A4F7" box="[1023,1098,1251,1273]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26" type="Museum">117068</specimenCode>
;
<specimenCode id="6E0CC4796D71FF9C05698BECFD4AA51D" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26" type="Museum">USNM 391302</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C03CC8BF2FB08A51D" ID-GBIF-Occurrence="3400112371" box="[754,1184,1277,1299]" collectionCode="AMNH" country="Bolivia" location="Rio Tijamuchi" pageId="25" pageNumber="26" specimenCode="AMNH 262012" specimenCount="1" stateProvince="Beni">
<location id="3B753AD96D71FF9C03CC8BF2FC22A51D" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C03CC8BF2FC22A51D" box="[754,906,1277,1299]" country="Bolivia" name="Rio Tijamuchi" pageId="25" pageNumber="26" stateProvince="Beni">Río Tijamuchi</location>
, sea level (
<specimenCode id="6E0CC4796D71FF9C02CA8BF2FB31A51D" box="[1012,1177,1277,1299]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH 262012</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C03A88A17FC5AA520" ID-GBIF-Occurrence="3400112346" box="[662,1010,1304,1326]" collectionCode="FMNH" country="Bolivia" location="Yuatre" pageId="25" pageNumber="26" specimenCode="FMNH 117060, 117061" specimenCount="1" stateProvince="Beni">
<location id="3B753AD96D71FF9C03A88A17FD75A520" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C03A88A17FD75A520" box="[662,733,1304,1326]" country="Bolivia" name="Yuatre" pageId="25" pageNumber="26" stateProvince="Beni">Yuatre</location>
(
<specimenCode id="6E0CC4796D71FF9C03D08A17FC3AA520" box="[750,914,1304,1326]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26" type="Museum">FMNH 117060</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C02A18A17FC42A520" box="[927,1002,1304,1326]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26" type="Museum">117061</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C053E8A17FB08A547" ID-GBIF-Occurrence="3400112341" accessionNumber="MSB 68481" collectionCode="MSB" country="Bolivia" elevation="253" location="Km" pageId="25" pageNumber="26" specimenCode="AMNH 264769" specimenCount="1" stateProvince="Beni">
<location id="3B753AD96D71FF9C053E8A17FB8FA520" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C053E8A17FB8FA520" box="[1024,1063,1304,1326]" country="Bolivia" name="Km" pageId="25" pageNumber="26" stateProvince="Beni">Km</location>
35,
<locationDeviation id="2FA775206D71FF9C05678A17FD59A547" location="Yucumo" pageId="25" pageNumber="26">NW of Yucumo</locationDeviation>
,
<quantity id="F952C1E76D71FF9C023D8A3CFCEBA547" box="[771,835,1331,1353]" metricMagnitude="2" metricUnit="m" metricValue="2.53" pageId="25" pageNumber="26" unit="m" value="253.0">
<elevation id="B5878B316D71FF9C023D8A3CFCEBA547" box="[771,835,1331,1353]" metricMagnitude="2" metricUnit="m" metricValue="2.53" pageId="25" pageNumber="26" unit="m" value="253.0">253 m</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D71FF9C02668A3CFBACA547" box="[856,1028,1331,1353]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH 264769</specimenCode>
;
<collectionCode id="58BBF4C76D71FF9C05288A3CFBE3A547" box="[1046,1099,1331,1353]" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="25" pageNumber="26" type="Museum">MSB</collectionCode>
68481)
</materialsCitation>
.
<materialsCitation id="8EC2665F6D71FF9C03A88A41FD74A570" ID-GBIF-Occurrence="3400112352" accessionNumber="MSB 63355" collectionCode="MSB" country="Bolivia" elevation="1200" location="2 km E Chuhuayacu" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Chuquisaca">
<emphasis id="0CDEB0106D71FF9C03A88A41FCA1A56D" box="[662,777,1358,1379]" italics="true" pageId="25" pageNumber="26">
<collectingRegion id="FC6EA2E06D71FF9C03A88A41FCA1A56D" box="[662,777,1358,1379]" country="Bolivia" name="Chuquisaca" pageId="25" pageNumber="26">Chuquisaca</collectingRegion>
</emphasis>
,
<location id="3B753AD96D71FF9C02228A41FC56A56A" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C02228A41FC56A56A" box="[796,1022,1358,1380]" country="Bolivia" name="2 km E Chuhuayacu" pageId="25" pageNumber="26" stateProvince="Chuquisaca">
<quantity id="F952C1E76D71FF9C02228A41FCF9A56A" box="[796,849,1358,1380]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="km" value="2.0">2 km</quantity>
E Chuhuayacu
</location>
,
<quantity id="F952C1E76D71FF9C05318A41FBF2A56A" box="[1039,1114,1358,1380]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="26" unit="m" value="1200.0">
<elevation id="B5878B316D71FF9C05318A41FBF2A56A" box="[1039,1114,1358,1380]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="25" pageNumber="26" unit="m" value="1200.0">1200 m</elevation>
</quantity>
(
<collectionCode id="58BBF4C76D71FF9C05508A41FB0CA56A" box="[1134,1188,1358,1380]" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="25" pageNumber="26" type="Museum">MSB</collectionCode>
63355)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C03D88A67FD40A5BA" ID-GBIF-Occurrence="3400112330" collectionCode="USNM" country="Bolivia" county="Tomina Province" elevation="2100" location="Tola Orko" municipality="Padilla" pageId="25" pageNumber="26" specimenCode="USNM 271581, 271582, 271584- 271587" specimenCount="1" stateProvince="Chuquisaca">
<collectingCounty id="D774148E6D71FF9C03D88A67FC30A570" box="[742,920,1384,1406]" pageId="25" pageNumber="26">Tomina Province</collectingCounty>
,
<quantity id="F952C1E76D71FF9C029C8A67FC4BA570" box="[930,995,1384,1406]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="26" unit="km" value="40.0">40 km</quantity>
from
<collectingMunicipality id="DE71F6786D71FF9C051F8A67FBC3A570" box="[1057,1131,1384,1406]" pageId="25" pageNumber="26">Padilla</collectingMunicipality>
,
<location id="3B753AD96D71FF9C054A8A67FD67A597" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C054A8A67FD67A597" country="Bolivia" county="Tomina Province" municipality="Padilla" name="Tola Orko" pageId="25" pageNumber="26" stateProvince="Chuquisaca">Tola Orko</location>
,
<quantity id="F952C1E76D71FF9C03E18A8CFC83A597" box="[735,811,1411,1433]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="25" pageNumber="26" unit="m" value="2100.0">
<elevation id="B5878B316D71FF9C03E18A8CFC83A597" box="[735,811,1411,1433]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="25" pageNumber="26" unit="m" value="2100.0">2100 m</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D71FF9C02018A8CFC4CA597" box="[831,996,1411,1433]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26" type="Museum">USNM 271581</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C02CA8A8CFB97A597" box="[1012,1087,1411,1433]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26" type="Museum">271582</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C056E8A8CFD4AA5BA" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26" type="Museum">271584– 271587</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C03CC8A91FBE2A5BA" ID-GBIF-Occurrence="3400112315" accessionNumber="MSB 63354" box="[754,1098,1438,1460]" collectionCode="MSB" country="Bolivia" elevation="1300" location="Rio Limon" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Chuquisaca">
<location id="3B753AD96D71FF9C03CC8A91FCCDA5BA" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C03CC8A91FCCDA5BA" box="[754,869,1438,1460]" country="Bolivia" name="Rio Limon" pageId="25" pageNumber="26" stateProvince="Chuquisaca">Río Limón</location>
,
<quantity id="F952C1E76D71FF9C02518A91FC12A5BA" box="[879,954,1438,1460]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="26" unit="m" value="1300.0">
<elevation id="B5878B316D71FF9C02518A91FC12A5BA" box="[879,954,1438,1460]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="26" unit="m" value="1300.0">1300 m</elevation>
</quantity>
(
<collectionCode id="58BBF4C76D71FF9C02F98A91FC55A5BA" box="[967,1021,1438,1460]" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="25" pageNumber="26" type="Museum">MSB</collectionCode>
63354)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C056D8A91FBBFA5C0" ID-GBIF-Occurrence="3400112365" collectionCode="USNM" collectorName="Tihumayu" country="Bolivia" location="Tomina Province" pageId="25" pageNumber="26" specimenCode="USNM 290906" specimenCount="1" stateProvince="Chuquisaca">
<location id="3B753AD96D71FF9C056D8A91FD5AA5C0" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C056D8A91FD5AA5C0" country="Bolivia" name="Tomina Province" pageId="25" pageNumber="26" stateProvince="Chuquisaca">Tomina Province</location>
,
<collectorName id="935F09D46D71FF9C03C58AB7FCCDA5C0" box="[763,869,1464,1486]" pageId="25" pageNumber="26">Tihumayu</collectorName>
(
<specimenCode id="6E0CC4796D71FF9C024F8AB7FBB8A5C0" box="[881,1040,1464,1486]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26" type="Museum">USNM 290906</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C051E8AB7FC04A5E7" ID-GBIF-Occurrence="3400112321" collectionCode="FMNH" country="Bolivia" location="Rio Capirenda" municipality="Ticucha" pageId="25" pageNumber="26" specimenCode="FMNH 72890" specimenCount="1" stateProvince="Chuquisaca">
<collectingMunicipality id="DE71F6786D71FF9C051E8AB7FBDCA5C0" box="[1056,1140,1464,1486]" pageId="25" pageNumber="26">Ticucha</collectingMunicipality>
,
<location id="3B753AD96D71FF9C05438AB7FCA9A5E7" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C05438AB7FCA9A5E7" country="Bolivia" municipality="Ticucha" name="Rio Capirenda" pageId="25" pageNumber="26" stateProvince="Chuquisaca">Río Capirenda</location>
(
<specimenCode id="6E0CC4796D71FF9C02308ADCFC0DA5E7" box="[782,933,1491,1513]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26" type="Museum">FMNH 72890</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="8EC2665F6D71FF9C028B8ADCFC49A60A" ID-GBIF-Occurrence="3400112329" collectionCode="ANSP" country="Bolivia" elevation="251" location="Rio Chapare" pageId="25" pageNumber="26" specimenCode="ANSP 19405" specimenCount="1" stateProvince="Cochabamba">
<emphasis id="0CDEB0106D71FF9C028B8ADCFB9FA5E6" box="[949,1079,1491,1512]" italics="true" pageId="25" pageNumber="26">
<collectingRegion id="FC6EA2E06D71FF9C028B8ADCFB9FA5E6" box="[949,1079,1491,1512]" country="Bolivia" name="Cochabamba" pageId="25" pageNumber="26">Cochabamba</collectingRegion>
</emphasis>
, Boca
<location id="3B753AD96D71FF9C05438ADCFD58A60A" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C05438ADCFD58A60A" country="Bolivia" name="Rio Chapare" pageId="25" pageNumber="26" stateProvince="Cochabamba">Río Chapare</location>
,
<quantity id="F952C1E76D71FF9C03C08AE1FC92A60A" box="[766,826,1518,1540]" metricMagnitude="2" metricUnit="m" metricValue="2.5145999999999997" pageId="25" pageNumber="26" unit="ft" value="825.0">
<elevation id="B5878B316D71FF9C03C08AE1FC92A60A" box="[766,826,1518,1540]" metricMagnitude="2" metricUnit="m" metricValue="2.5145999999999997" pageId="25" pageNumber="26" unit="ft" value="825.0">825 ft</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D71FF9C02728AE1FC71A60A" box="[844,985,1518,1540]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="26">ANSP 19405</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C02CE8AE1FC8CA661" ID-GBIF-Occurrence="3400112309" collectionCode="AMNH, FMNH" collectorName="Inst. Roy. Sc. Nat. Belgium" country="Bolivia" elevation="396" location="Todos Santos" municipality="Rio Moile Ichilo" pageId="25" pageNumber="26" specimenCode="AMNH 38520-38523, 38561, 38563-38566, 38568-38578, 40782-40786, FMNH 21520-21524" specimenCount="1" stateProvince="Cochabamba">
<collectingMunicipality id="DE71F6786D71FF9C02CE8AE1FB0CA60A" box="[1008,1188,1518,1540]" pageId="25" pageNumber="26">Río Moile Ichilo</collectingMunicipality>
(
<collectorName id="935F09D46D71FF9C03A38906FC16A611" box="[669,958,1545,1567]" pageId="25" pageNumber="26">Inst. Roy. Sc. Nat. Belgium</collectorName>
20103, 25297);
<location id="3B753AD96D71FF9C055D8906FD75A637" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C055D8906FD75A637" country="Bolivia" municipality="Rio Moile Ichilo" name="Todos Santos" pageId="25" pageNumber="26" stateProvince="Cochabamba">Todos Santos</location>
,
<quantity id="F952C1E76D71FF9C03C8892CFC96A637" box="[758,830,1571,1593]" metricMagnitude="2" metricUnit="m" metricValue="3.9624" pageId="25" pageNumber="26" unit="ft" value="1300.0">
<elevation id="B5878B316D71FF9C03C8892CFC96A637" box="[758,830,1571,1593]" metricMagnitude="2" metricUnit="m" metricValue="3.9624" pageId="25" pageNumber="26" unit="ft" value="1300.0">1300 ft</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D71FF9C0264892CFBE0A637" box="[858,1096,1571,1593]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">AMNH 38520–38523</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C055F892CFB08A637" box="[1121,1184,1571,1593]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">38561</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C03A88931FCB4A65A" box="[662,796,1598,1620]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">38563–38566</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C02148931FC19A65A" box="[810,945,1598,1620]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">38568–38578</specimenCode>
,
<specimenCode id="6E0CC4796D71FF9C02FE8931FBEEA65A" box="[960,1094,1598,1620]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="25" pageNumber="26">40782–40786</specimenCode>
;
<specimenCode id="6E0CC4796D71FF9C056B8931FCB6A661" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26" type="Museum">FMNH 21520–21524</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9C020F8956FC0CA6AA" ID-GBIF-Occurrence="3400112369" collectionCode="BMNH" collectorName="Mission Mosetenes" country="Bolivia" location="upper Rio Mamore'" municipality="Yungas Prov." pageId="25" pageNumber="26" specimenCode="BMNH 1900.8" specimenCount="1" stateProvince="Cochabamba" typeStatus="holotype">
<collectingMunicipality id="DE71F6786D71FF9C020F8956FC6BA661" box="[817,963,1625,1647]" pageId="25" pageNumber="26">Yungas Prov.</collectingMunicipality>
,
<location id="3B753AD96D71FF9C02EE8956FB08A661" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C02EE8956FB08A661" box="[976,1184,1624,1647]" country="Bolivia" municipality="Yungas Prov." name="upper Rio Mamore'" pageId="25" pageNumber="26" stateProvince="Cochabamba">upper Río Mamore´</location>
,
<collectorName id="935F09D46D71FF9C03A8897BFCF5A684" box="[662,861,1652,1674]" pageId="25" pageNumber="26">Mission Mosetenes</collectorName>
(
<specimenCode id="6E0CC4796D71FF9C0253897BFBA4A684" box="[877,1036,1652,1674]" collectionCode="BMNH" pageId="25" pageNumber="26">BMNH 1900.8</specimenCode>
.3.21 [
<typeStatus id="E111D2A06D71FF9C0577897BFB0BA684" box="[1097,1187,1652,1674]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
of
<taxonomicName id="F9AA17816D71FF9C038D8980FCA7A6AA" authorityName="Thomas" authorityYear="1906" box="[691,783,1679,1700]" class="Mammalia" family="Muridae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="mamorae">
<emphasis id="0CDEB0106D71FF9C038D8980FCA7A6AA" box="[691,783,1679,1700]" italics="true" pageId="25" pageNumber="26">mamorae</emphasis>
</taxonomicName>
], 1900.8.3.24)
</materialsCitation>
.
<materialsCitation id="8EC2665F6D71FF9C02918980FCD4A6B1" ID-GBIF-Occurrence="3400112347" accessionNumber="MSU 33018" collectionCode="MSU" country="Bolivia" location="1 mi W Puerto Linares" pageId="25" pageNumber="26" specimenCount="1" stateProvince="La Paz" typeStatus="holotype">
<emphasis id="0CDEB0106D71FF9C02918980FC51A6AA" box="[943,1017,1679,1700]" italics="true" pageId="25" pageNumber="26">
<collectingRegion id="FC6EA2E06D71FF9C02918980FC51A6AA" box="[943,1017,1679,1700]" country="Bolivia" name="La Paz" pageId="25" pageNumber="26">La Paz</collectingRegion>
</emphasis>
,
<location id="3B753AD96D71FF9C05368981FD4AA6B1" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C05368981FD4AA6B1" country="Bolivia" name="1 mi W Puerto Linares" pageId="25" pageNumber="26" stateProvince="La Paz">
<quantity id="F952C1E76D71FF9C05368981FB9EA6AA" box="[1032,1078,1678,1700]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="25" pageNumber="26" unit="mi" value="1.0">1 mi</quantity>
W Puerto Linares
</location>
(
<collectionCode id="58BBF4C76D71FF9C03CC89A6FC83A6B1" box="[754,811,1705,1727]" country="USA" httpUri="http://grbio.org/cool/53w1-8r3k" name="Michigan State University Museum" pageId="25" pageNumber="26" type="University or college">MSU</collectionCode>
33018)
</materialsCitation>
.
<materialsCitation id="8EC2665F6D71FF9C02B689A6FB8BA6D4" ID-GBIF-Occurrence="3400112353" collectionCode="USNM" collectorName="Ayacucho" country="Bolivia" location="Andres Ibanez Prov." pageId="25" pageNumber="26" specimenCode="USNM 390655" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<emphasis id="0CDEB0106D71FF9C02B689A6FC55A6B0" box="[904,1021,1705,1726]" italics="true" pageId="25" pageNumber="26">
<collectingRegion id="FC6EA2E06D71FF9C02B689A6FC55A6B0" box="[904,1021,1705,1726]" country="Bolivia" name="Santa Cruz" pageId="25" pageNumber="26">Santa Cruz</collectingRegion>
</emphasis>
,
<location id="3B753AD96D71FF9C053289A6FD78A6D4" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9C053289A6FD78A6D4" country="Bolivia" name="Andres Ibanez Prov." pageId="25" pageNumber="26" stateProvince="Santa Cruz">Andrés Ibáñez Prov.</location>
,
<collectorName id="935F09D46D71FF9C03D789CBFCF8A6D4" box="[745,848,1732,1754]" pageId="25" pageNumber="26">Ayacucho</collectorName>
(
<specimenCode id="6E0CC4796D71FF9C025389CBFBB4A6D4" box="[877,1052,1732,1754]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26" type="Museum">USNM 390655</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D71FF9F050289CBFE6FA0E6" ID-GBIF-Occurrence="3400112305" collectionCode="USNM" collectorName="Basilio" country="Bolivia" lastPageId="26" lastPageNumber="27" location="Cordillera Prov." pageId="25" pageNumber="26" specimenCode="USNM 390654" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<location id="3B753AD96D71FF9F050289CBFF12A0E6" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D71FF9F050289CBFF12A0E6" country="Bolivia" lastPageId="26" lastPageNumber="27" name="Cordillera Prov." pageId="25" pageNumber="26" stateProvince="Santa Cruz">Cordillera Prov.</location>
,
<collectorName id="935F09D46D72FF9F01F98FDDFEA5A0E6" box="[199,269,210,232]" pageId="26" pageNumber="27">Basilio</collectorName>
(
<specimenCode id="6E0CC4796D72FF9F00238FDDFE68A0E6" box="[285,448,210,232]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27" type="Museum">USNM 390654</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F00EA8FDDFE05A136" ID-GBIF-Occurrence="3400112326" collectionCode="AMNH, BMNH" country="Bolivia" elevation="500" location="Buena Vista" pageId="26" pageNumber="27" specimenCode="AMNH 61776, BMNH 26.12" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<location id="3B753AD96D72FF9F00EA8FDDFDFDA0E6" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F00EA8FDDFDFDA0E6" box="[468,597,210,232]" country="Bolivia" name="Buena Vista" pageId="26" pageNumber="27" stateProvince="Santa Cruz">Buena Vista</location>
, 350, 450, and
<quantity id="F952C1E76D72FF9F01E38FE2FEB5A10D" box="[221,285,237,259]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="m" value="500.0">
<elevation id="B5878B316D72FF9F01E38FE2FEB5A10D" box="[221,285,237,259]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="m" value="500.0">500 m</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D72FF9F00148FE2FE69A10D" box="[298,449,237,259]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="26" pageNumber="27">AMNH 61776</specimenCode>
;
<specimenCode id="6E0CC4796D72FF9F00F58FE2FDF2A10D" box="[459,602,237,259]" collectionCode="BMNH" pageId="26" pageNumber="27">BMNH 26.12</specimenCode>
.4.52, 28.2.9.39–28.2.9.42, 51.5.23.6; FMNH 25267, 25268, 51907, 51913, 51915)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F00858E2DFE9CA163" ID-GBIF-Occurrence="3400112360" accessionNumber="CAS 13805" collectionCode="CAS" country="Bolivia" elevation="1000" location="Camiri" municipality="Cordillera Province" pageId="26" pageNumber="27" specimenCode="USNM 276602" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<collectingMunicipality id="DE71F6786D72FF9F00858E2DFD22A136" box="[443,650,290,312]" pageId="26" pageNumber="27">Cordillera Province</collectingMunicipality>
,
<locationDeviation id="2FA775206D72FF9F01BE8E32FE9BA15D" box="[128,307,317,339]" location="Choreti" pageId="26" pageNumber="27">
<quantity id="F952C1E76D72FF9F01BE8E32FF1CA15D" box="[128,180,317,339]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="km" value="5.0">5 km</quantity>
S Choreti
</locationDeviation>
,
<location id="3B753AD96D72FF9F00768E32FE3AA15D" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F00768E32FE3AA15D" box="[328,402,317,339]" country="Bolivia" municipality="Cordillera Province" name="Camiri" pageId="26" pageNumber="27" stateProvince="Santa Cruz">Camiri</location>
,
<quantity id="F952C1E76D72FF9F00988E32FE5AA15D" box="[422,498,317,339]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="m" value="1000.0">
<elevation id="B5878B316D72FF9F00988E32FE5AA15D" box="[422,498,317,339]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="m" value="1000.0">1000 m</elevation>
</quantity>
(
<collectionCode id="58BBF4C76D72FF9F03378E32FD93A15D" box="[521,571,317,339]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="26" pageNumber="27" type="Museum">CAS</collectionCode>
13805;
<specimenCode id="6E0CC4796D72FF9F01BE8E58FE85A163" box="[128,301,343,365]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27" type="Museum">USNM 276602</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F00738E58FDABA186" ID-GBIF-Occurrence="3400112358" accessionNumber="MSB 55313" collectionCode="MSB" country="Bolivia" elevation="300" location="Estacion Pailon" pageId="26" pageNumber="27" specimenCode="AMNH 260420" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
3.5 km W
<location id="3B753AD96D72FF9F00E98E58FD21A163" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F00E98E58FD21A163" box="[471,649,343,365]" country="Bolivia" name="Estacion Pailon" pageId="26" pageNumber="27" stateProvince="Santa Cruz">Estación Pailón</location>
,
<quantity id="F952C1E76D72FF9F01BE8E7DFF17A186" box="[128,191,370,392]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="27" unit="m" value="300.0">
<elevation id="B5878B316D72FF9F01BE8E7DFF17A186" box="[128,191,370,392]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="27" unit="m" value="300.0">300 m</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D72FF9F01F08E7DFEDCA186" box="[206,372,370,392]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="26" pageNumber="27">AMNH 260420</specimenCode>
;
<collectionCode id="58BBF4C76D72FF9F00418E7DFE1CA186" box="[383,436,370,392]" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="26" pageNumber="27" type="Museum">MSB</collectionCode>
55313)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F03308E7DFF7AA1B3" ID-GBIF-Occurrence="3400112319" collectionCode="AMNH" country="Bolivia" elevation="535" location="Ingeniero Mora" pageId="26" pageNumber="27" specimenCode="AMNH 247757" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<quantity id="F952C1E76D72FF9F03308E7DFDEBA186" box="[526,579,370,392]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="27" unit="km" value="3.0">3 km</quantity>
N and
<quantity id="F952C1E76D72FF9F01BE8E82FF1CA1AD" box="[128,180,397,419]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="26" pageNumber="27" unit="km" value="7.0">7 km</quantity>
E
<location id="3B753AD96D72FF9F01D48E82FE33A1AD" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F01D48E82FE33A1AD" box="[234,411,397,419]" country="Bolivia" name="Ingeniero Mora" pageId="26" pageNumber="27" stateProvince="Santa Cruz">Ingeniero Mora</location>
,
<quantity id="F952C1E76D72FF9F008C8E82FD89A1AD" box="[434,545,397,419]" metricMagnitude="2" metricUnit="m" metricValue="5.35" metricValueMax="5.8" metricValueMin="4.9" pageId="26" pageNumber="27" unit="m" value="535.0" valueMax="580.0" valueMin="490.0">
<elevation id="B5878B316D72FF9F008C8E82FD89A1AD" box="[434,545,397,419]" metricMagnitude="2" metricUnit="m" metricValue="5.35" metricValueMax="5.8" metricValueMin="4.9" pageId="26" pageNumber="27" unit="m" value="535.0" valueMax="580.0" valueMin="490.0">490–580 m</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D72FF9F03058E82FF64A1B3" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="26" pageNumber="27">AMNH 247757</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F01D88EA8FF7BA1D6" ID-GBIF-Occurrence="3400112344" collectionCode="AMNH" country="Bolivia" location="Punta Rieles" pageId="26" pageNumber="27" specimenCode="AMNH 263101-263104, 263366" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<location id="3B753AD96D72FF9F01D88EA8FED9A1B3" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F01D88EA8FED9A1B3" box="[230,369,423,445]" country="Bolivia" name="Punta Rieles" pageId="26" pageNumber="27" stateProvince="Santa Cruz">Punta Rieles</location>
(
<specimenCode id="6E0CC4796D72FF9F00B78EA8FD22A1B3" box="[393,650,423,445]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="26" pageNumber="27">AMNH 263101–263104</specimenCode>
,
<specimenCode id="6E0CC4796D72FF9F01BE8ECDFF64A1D6" box="[128,204,450,472]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="26" pageNumber="27">263366</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F01DE8ECDFD22A1FD" ID-GBIF-Occurrence="3400112301" accessionNumber="MSB 56072" collectionCode="MSB" country="Bolivia" elevation="400" location="San Rafael de Amboro" pageId="26" pageNumber="27" specimenCode="AMNH 262013, 262014, 262119, 262099" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<location id="3B753AD96D72FF9F01DE8ECDFE71A1D6" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F01DE8ECDFE71A1D6" box="[224,473,450,472]" country="Bolivia" name="San Rafael de Amboro" pageId="26" pageNumber="27" stateProvince="Santa Cruz">San Rafael de Amboró</location>
,
<quantity id="F952C1E76D72FF9F00D48ECDFD82A1D6" box="[490,554,450,472]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="m" value="400.0">
<elevation id="B5878B316D72FF9F00D48ECDFD82A1D6" box="[490,554,450,472]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="m" value="400.0">400 m</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D72FF9F03058ECDFF63A1FD" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="26" pageNumber="27">AMNH 262013</specimenCode>
,
<specimenCode id="6E0CC4796D72FF9F01E18ED2FE82A1FD" box="[223,298,477,499]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="26" pageNumber="27">262014</specimenCode>
,
<specimenCode id="6E0CC4796D72FF9F00008ED2FE21A1FD" box="[318,393,477,499]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="26" pageNumber="27">262119</specimenCode>
,
<specimenCode id="6E0CC4796D72FF9F00A08ED2FE41A1FD" box="[414,489,477,499]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="26" pageNumber="27">262099</specimenCode>
;
<collectionCode id="58BBF4C76D72FF9F00C38ED2FD9AA1FD" box="[509,562,477,499]" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="26" pageNumber="27" type="Museum">MSB</collectionCode>
56072)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F01BE8EF8FF7AA226" ID-GBIF-Occurrence="3400112338" collectionCode="USNM" collectorName="Santa Ana" country="Bolivia" location="Velasco Prov." pageId="26" pageNumber="27" specimenCode="USNM 390656, 391301" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<location id="3B753AD96D72FF9F01BE8EF8FE8AA203" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F01BE8EF8FE8AA203" box="[128,290,503,525]" country="Bolivia" name="Velasco Prov." pageId="26" pageNumber="27" stateProvince="Santa Cruz">Velasco Prov.</location>
,
<collectorName id="935F09D46D72FF9F00008EF8FE13A203" box="[318,443,503,525]" pageId="26" pageNumber="27">Santa Ana</collectorName>
(
<specimenCode id="6E0CC4796D72FF9F00E58EF8FD22A203" box="[475,650,503,525]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27" type="Museum">USNM 390656</specimenCode>
,
<specimenCode id="6E0CC4796D72FF9F01BE8D1DFF63A226" box="[128,203,530,552]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27" type="Museum">391301</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F01E28D1DFD22A226" ID-GBIF-Occurrence="3400112348" accessionNumber="MSB 58648" box="[220,650,530,552]" collectionCode="MSB" country="Bolivia" elevation="400" location="Santa Cruz" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<quantity id="F952C1E76D72FF9F01E28D1DFEB5A226" box="[220,285,530,552]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="km" value="15.0">15 km</quantity>
S
<collectingRegion id="FC6EA2E06D72FF9F00078D1DFE06A226" box="[313,430,530,552]" country="Bolivia" name="Santa Cruz" pageId="26" pageNumber="27">Santa Cruz</collectingRegion>
,
<quantity id="F952C1E76D72FF9F00848D1DFE51A226" box="[442,505,530,552]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="m" value="400.0">
<elevation id="B5878B316D72FF9F00848D1DFE51A226" box="[442,505,530,552]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="m" value="400.0">400 m</elevation>
</quantity>
(
<collectionCode id="58BBF4C76D72FF9F03398D1DFD94A226" box="[519,572,530,552]" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="26" pageNumber="27" type="Museum">MSB</collectionCode>
58648)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F01BE8D22FDF7A24D" ID-GBIF-Occurrence="3400112354" box="[128,607,557,579]" collectionCode="CM" country="Bolivia" elevation="410" location="Santa Cruz de la Sierra" pageId="26" pageNumber="27" specimenCode="CM 2146, 2749" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<collectingRegion id="FC6EA2E06D72FF9F01BE8D22FEC0A24D" box="[128,360,557,579]" country="Bolivia" name="Santa Cruz" pageId="26" pageNumber="27">Santa Cruz de la Sierra</collectingRegion>
,
<quantity id="F952C1E76D72FF9F004E8D22FE18A24D" box="[368,432,557,579]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="26" pageNumber="27" unit="m" value="410.0">
<elevation id="B5878B316D72FF9F004E8D22FE18A24D" box="[368,432,557,579]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="26" pageNumber="27" unit="m" value="410.0">410 m</elevation>
</quantity>
(
<specimenCode id="6E0CC4796D72FF9F00828D22FDB4A24D" box="[444,540,557,579]" collectionCode="CM" country="China" name="Chongqing Museum" pageId="26" pageNumber="27">CM 2146</specimenCode>
,
<specimenCode id="6E0CC4796D72FF9F031B8D22FDF0A24D" box="[549,600,557,579]" collectionCode="CM" country="China" name="Chongqing Museum" pageId="26" pageNumber="27">2749</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F03568D23FE32A253" ID-GBIF-Occurrence="3400112332" collectionCode="FMNH" country="Bolivia" location="Rio Yapacani" pageId="26" pageNumber="27" specimenCode="FMNH 51914" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<location id="3B753AD96D72FF9F03568D23FF49A253" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F03568D23FF49A253" country="Bolivia" name="Rio Yapacani" pageId="26" pageNumber="27" stateProvince="Santa Cruz">Río Yapacaní</location>
(
<specimenCode id="6E0CC4796D72FF9F01CB8D48FE3BA253" box="[245,403,583,605]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="26" pageNumber="27" type="Museum">FMNH 51914</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="8EC2665F6D72FF9F00948D48FEC1A276" ID-GBIF-Occurrence="3400112312" collectionCode="NMW" collectorName="Caicara" country="Brazil" location="Mato Grosso" pageId="26" pageNumber="27" specimenCode="B475" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
<emphasis id="0CDEB0106D72FF9F00948D48FE4EA253" bold="true" box="[426,486,583,605]" pageId="26" pageNumber="27">
<collectingCountry id="46BD2C926D72FF9F00948D48FE4EA253" box="[426,486,583,605]" name="Brazil" pageId="26" pageNumber="27">Brazil</collectingCountry>
</emphasis>
—
<emphasis id="0CDEB0106D72FF9F00C08D47FD2FA253" box="[510,647,584,605]" italics="true" pageId="26" pageNumber="27">
<collectingRegion id="FC6EA2E06D72FF9F00C08D47FD2FA253" box="[510,647,584,605]" country="Brazil" name="Mato Grosso" pageId="26" pageNumber="27">Mato Grosso</collectingRegion>
</emphasis>
,
<collectorName id="935F09D46D72FF9F01BE8D6DFF66A276" box="[128,206,610,632]" pageId="26" pageNumber="27">Caiçara</collectorName>
(
<collectionCode id="58BBF4C76D72FF9F01E08D6DFE8AA276" box="[222,290,610,632]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">NMW</collectionCode>
<specimenCode id="6E0CC4796D72FF9F00158D6DFECAA276" box="[299,354,610,632]" collectionCode="NMW" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="27">B475</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F00488D6DFF6EA29D" ID-GBIF-Occurrence="3400112372" collectionCode="FMNH" country="Brazil" location="Uca" municipality="Descalvados" pageId="26" pageNumber="27" specimenCode="FMNH 26643" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
<collectingMunicipality id="DE71F6786D72FF9F00488D6DFE5FA276" box="[374,503,610,632]" pageId="26" pageNumber="27">Descalvados</collectingMunicipality>
,
<location id="3B753AD96D72FF9F033A8D6DFD86A276" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F033A8D6DFD86A276" box="[516,558,610,632]" country="Brazil" municipality="Descalvados" name="Uca" pageId="26" pageNumber="27" stateProvince="Mato Grosso">Uca</location>
(
<specimenCode id="6E0CC4796D72FF9F03008D6DFF68A29D" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="26" pageNumber="27" type="Museum">FMNH 26643</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="8EC2665F6D72FF9F01E78D72FE94A2A3" ID-GBIF-Occurrence="3400112323" collectionCode="USNM" collectorName="Fazenda Acurizal" country="Brazil" location="Mato Grosso do Sul" pageId="26" pageNumber="27" specimenCode="USNM 531278" specimenCount="1" stateProvince="Mato Grosso do Sul" typeStatus="holotype">
<emphasis id="0CDEB0106D72FF9F01E78D72FE14A29C" box="[217,444,637,658]" italics="true" pageId="26" pageNumber="27">
<collectingRegion id="FC6EA2E06D72FF9F01E78D72FE14A29C" box="[217,444,637,658]" country="Brazil" name="Mato Grosso do Sul" pageId="26" pageNumber="27">Mato Grosso do Sul</collectingRegion>
</emphasis>
,
<collectorName id="935F09D46D72FF9F00EF8D72FD26A29D" box="[465,654,637,659]" pageId="26" pageNumber="27">Fazenda Acurizal</collectorName>
(
<specimenCode id="6E0CC4796D72FF9F01B88D98FE9DA2A3" box="[134,309,663,685]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27" type="Museum">USNM 531278</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F00688D97FE31A2C6" ID-GBIF-Occurrence="3400112370" collectionCode="FMNH" country="Brazil" location="Urucum de Corumba" municipality="Rio Paraguai" pageId="26" pageNumber="27" specimenCode="FMNH 26811" specimenCount="1" stateProvince="Mato Grosso do Sul" typeStatus="holotype">
<collectingMunicipality id="DE71F6786D72FF9F00688D97FE58A2A3" box="[342,496,663,686]" pageId="26" pageNumber="27">Río Paraguai</collectingMunicipality>
,
<location id="3B753AD96D72FF9F03348D98FF49A2C6" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F03348D98FF49A2C6" country="Brazil" municipality="Rio Paraguai" name="Urucum de Corumba" pageId="26" pageNumber="27" stateProvince="Mato Grosso do Sul">Urúcum de Corumbá</location>
(
<specimenCode id="6E0CC4796D72FF9F01CB8DBDFE3BA2C6" box="[245,403,690,712]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="26" pageNumber="27" type="Museum">FMNH 26811</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="8EC2665F6D72FF9F00948DBDFE42A2F3" ID-GBIF-Occurrence="3400112327" accessionNumber="MSB 70699" collectionCode="MSB" collectorName="E Rio Apa" country="Paraguay" location="Colonia Sargento Dure'" municipality="Bella Vista" pageId="26" pageNumber="27" specimenCode="70739" specimenCount="1" stateProvince="Amambay" typeStatus="holotype">
<emphasis id="0CDEB0106D72FF9F00948DBDFDA2A2C6" bold="true" box="[426,522,690,712]" pageId="26" pageNumber="27">
<collectingCountry id="46BD2C926D72FF9F00948DBDFDA2A2C6" box="[426,522,690,712]" name="Paraguay" pageId="26" pageNumber="27">Paraguay</collectingCountry>
</emphasis>
—
<emphasis id="0CDEB0106D72FF9F031C8DBDFD2FA2C9" box="[546,647,690,711]" italics="true" pageId="26" pageNumber="27">
<collectingRegion id="FC6EA2E06D72FF9F031C8DBDFD2FA2C9" box="[546,647,690,711]" country="Paraguay" name="Amambay" pageId="26" pageNumber="27">Amambay</collectingRegion>
</emphasis>
,
<collectingMunicipality id="DE71F6786D72FF9F01BE8DC2FF58A2ED" box="[128,240,717,739]" pageId="26" pageNumber="27">Bella Vista</collectingMunicipality>
,
<location id="3B753AD96D72FF9F01C48DC2FE42A2ED" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F01C48DC2FE42A2ED" box="[250,490,717,739]" country="Paraguay" municipality="Bella Vista" name="Colonia Sargento Dure'" pageId="26" pageNumber="27" stateProvince="Amambay">Colonia Sargento Dure´</location>
,
<quantity id="F952C1E76D72FF9F00CA8DC2FD81A2ED" box="[500,553,717,739]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="27" unit="km" value="3.0">3 km</quantity>
(by road)
<collectorName id="935F09D46D72FF9F01BE8DE8FF55A2F3" box="[128,253,743,765]" pageId="26" pageNumber="27">E Río Apa</collectorName>
(
<collectionCode id="58BBF4C76D72FF9F002E8DE8FEEDA2F3" box="[272,325,743,765]" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="26" pageNumber="27" type="Museum">MSB</collectionCode>
70699,
<specimenCode id="6E0CC4796D72FF9F009D8DE8FE4BA2F3" box="[419,483,743,765]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="26" pageNumber="27" type="Museum">70739</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="8EC2665F6D72FF9F00C48DE7FF7AA33D" ID-GBIF-Occurrence="3400112366" collectionCode="UMMZ" country="Paraguay" location="Cerro Leon" municipality="Chaco" pageId="26" pageNumber="27" specimenCode="UMMZ 125456" specimenCount="1" stateProvince="Amambay" typeStatus="holotype">
<emphasis id="0CDEB0106D72FF9F00C48DE7FD91A2F3" box="[506,569,744,765]" italics="true" pageId="26" pageNumber="27">
<collectingMunicipality id="DE71F6786D72FF9F00C48DE7FD91A2F3" box="[506,569,744,765]" pageId="26" pageNumber="27">Chaco</collectingMunicipality>
</emphasis>
,
<locationDeviation id="2FA775206D72FF9F03738DE8FE25A316" location="Fortin Madrejon" pageId="26" pageNumber="27">
<quantity id="F952C1E76D72FF9F03738DE8FD26A2F3" box="[589,654,743,765]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="km" value="50.0">50 km</quantity>
WNW Fortín Madrejón
</locationDeviation>
,
<location id="3B753AD96D72FF9F009C8C0DFD8AA316" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F009C8C0DFD8AA316" box="[418,546,770,792]" country="Paraguay" municipality="Chaco" name="Cerro Leon" pageId="26" pageNumber="27" stateProvince="Amambay">Cerro León</location>
(
<specimenCode id="6E0CC4796D72FF9F03048C0DFF63A33D" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="26" pageNumber="27" type="University or college">UMMZ 125456</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8EC2665F6D72FF9F01E18C12FE29A366" ID-GBIF-Occurrence="3400112310" collectionCode="UCONN" country="Paraguay" county="Rio La Gorenza" location="San" municipality="Rio Timane" pageId="26" pageNumber="27" specimenCode="UCONN 19187-19189" specimenCount="1" stateProvince="Amambay" typeStatus="holotype">
<locationDeviation id="2FA775206D72FF9F01E18C12FD22A33D" box="[223,650,797,819]" location="Mayor Pablo La Gorenza" pageId="26" pageNumber="27">
<quantity id="F952C1E76D72FF9F01E18C12FEB7A33D" box="[223,287,797,819]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="26" pageNumber="27" unit="km" value="28.0">28 km</quantity>
WNW Mayor Pablo La Gorenza
</locationDeviation>
, edge
<collectingCounty id="D774148E6D72FF9F018A8C38FEF2A343" box="[180,346,823,845]" pageId="26" pageNumber="27">Río La Gorenza</collectingCounty>
(
<emphasis id="0CDEB0106D72FF9F00578C36FED5A343" bold="true" box="[361,381,825,845]" pageId="26" pageNumber="27">5</emphasis>
<collectingMunicipality id="DE71F6786D72FF9F00BA8C38FDA9A343" box="[388,513,823,845]" pageId="26" pageNumber="27">Río Timane</collectingMunicipality>
),
<location id="3B753AD96D72FF9F032C8C38FD90A343" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F032C8C38FD90A343" box="[530,568,823,845]" country="Paraguay" county="Rio La Gorenza" municipality="Rio Timane" name="San" pageId="26" pageNumber="27" stateProvince="Amambay">San</location>
Alfredo (
<specimenCode id="6E0CC4796D72FF9F01B88C5DFED2A366" box="[134,378,850,872]" collectionCode="UCONN" country="USA" httpUri="http://grbio.org/cool/jrf7-y871" name="University of Connecticut Biodiversity Research Collections" pageId="26" pageNumber="27">UCONN 19187–19189</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="8EC2665F6D72FF9F00AC8C5DFE3BA38D" ID-GBIF-Occurrence="3400112336" collectionCode="AMNH" country="Paraguay" location="40 km S San Ignacio" pageId="26" pageNumber="27" specimenCode="AMNH 234787" specimenCount="1" stateProvince="Misiones" typeStatus="holotype">
<emphasis id="0CDEB0106D72FF9F00AC8C5DFE42A369" box="[402,490,850,871]" italics="true" pageId="26" pageNumber="27">
<collectingRegion id="FC6EA2E06D72FF9F00AC8C5DFE42A369" box="[402,490,850,871]" country="Paraguay" name="Misiones" pageId="26" pageNumber="27">Misiones</collectingRegion>
</emphasis>
,
<location id="3B753AD96D72FF9F00C08C5DFF64A38D" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F00C08C5DFF64A38D" country="Paraguay" name="40 km S San Ignacio" pageId="26" pageNumber="27" stateProvince="Misiones">
<quantity id="F952C1E76D72FF9F00C08C5DFD97A366" box="[510,575,850,872]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="km" value="40.0">40 km</quantity>
S San Ignacio
</location>
(
<specimenCode id="6E0CC4796D72FF9F01DE8C62FE24A38D" box="[224,396,877,899]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="26" pageNumber="27">AMNH 234787</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="8EC2665F6D72FF9F009D8C62FDF6A393" ID-GBIF-Occurrence="3400112313" collectionCode="UMMZ" country="Paraguay" location="Saltos de Pirareta" pageId="26" pageNumber="27" specimenCode="UMMZ 133793" specimenCount="1" stateProvince="Paraguari" typeStatus="holotype">
<emphasis id="0CDEB0106D72FF9F009D8C62FDAEA38C" box="[419,518,877,898]" italics="true" pageId="26" pageNumber="27">
<collectingRegion id="FC6EA2E06D72FF9F009D8C62FDAEA38C" box="[419,518,877,898]" country="Paraguay" name="Paraguari" pageId="26" pageNumber="27">Paraguari</collectingRegion>
</emphasis>
,
<locationDeviation id="2FA775206D72FF9F03258C62FF4CA393" location="Piribebuy" pageId="26" pageNumber="27">
<quantity id="F952C1E76D72FF9F03258C62FDF3A38D" box="[539,603,877,899]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="26" pageNumber="27" unit="km" value="17.0">17 km</quantity>
SW Piribebuy
</locationDeviation>
,
<location id="3B753AD96D72FF9F01D18C88FE0BA393" LSID="urn:lsid:plazi:treatment:B603DD146D70FF9F01DE8AA2FDC9A393:3B753AD96D72FF9F01D18C88FE0BA393" box="[239,419,903,925]" country="Paraguay" name="Saltos de Pirareta" pageId="26" pageNumber="27" stateProvince="Paraguari">Saltos de Pirareta</location>
(
<specimenCode id="6E0CC4796D72FF9F008F8C88FDFFA393" box="[433,599,903,925]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="26" pageNumber="27" type="University or college">UMMZ 133793</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

1107
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<document ID-CLB-Dataset="291589" ID-GBIF-Dataset="1534c351-525b-451c-89df-0ff55b818363" ID-ISBN="978-3-258-08047-5" checkinTime="1706569713375" checkinUser="plazi" docAuthor="Konrad Lauber, Gerhart Wagner &amp; Andreas Gygax" docDate="2018" docId="B603F0CDA91BF07A3BD299ED116948FD" docLanguage="de" docName="FloraHelv.Fabaceae.xml" docOrigin="Flora Helvetica, Bern: Haupt Verlag" docTitle="Lotus corniculatus L." docType="treatment" docVersion="7" id="99615BCDA2F8E13E6BA123A303EE2544" lastPageNumber="318" masterDocId="8EE5A9E62DF2672ECBC27C54C8BB369F" masterDocTitle="Flora Helvetica - Fabaceae" masterLastPageNumber="400" masterPageNumber="37" pageNumber="318" updateTime="1711157890863" updateUser="ExternalLinkService">
<mods:mods id="32603080286B86A6DE6D407F441B2FF4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EF024BDD8DCAD7FA183A9A8FD48C3B47">
<mods:title id="F5E88F0CAF9A7CA423380267D1D94E3F">Flora Helvetica - Fabaceae</mods:title>
</mods:titleInfo>
<mods:name id="37F466B6667ABDC396C168213BF756EB" type="personal">
<mods:role id="CFB5294BA08580F255911D167547FD8D">
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</mods:role>
<mods:namePart id="9D3DD0FD202D67391630ED4F10AE4BA9">Konrad Lauber</mods:namePart>
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<mods:namePart id="03C9A020CF55211AB889447070626280">Gerhart Wagner</mods:namePart>
</mods:name>
<mods:name id="315B7282CA9F997AFCFC5382A85CAC1A" type="personal">
<mods:role id="32A4A2FE6F1E4EFC0D76D54841A68674">
<mods:roleTerm id="3D5727AE708E6A7BF5A05F5F6B2C5F08">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3DE0FFD7D0723571149FAF66E6F5D0F3">Andreas Gygax</mods:namePart>
</mods:name>
<mods:typeOfResource id="0C2ABF361B14CA1D2E03F70BC39457FC">text</mods:typeOfResource>
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<mods:dateIssued id="427F92505A91D83550DE8F812C5C65B3">2018</mods:dateIssued>
<mods:publisher id="19B0C450F09087C397AE29D61C333E50">Haupt Verlag</mods:publisher>
<mods:place id="9B33454D197786FE6D1FDCFCC56675A7">
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<mods:titleInfo id="0C03704F25794C01E99B4A2BA5A1296F">
<mods:title id="D8883D292B8A65C3CA008E8A6B27CA45">Flora Helvetica</mods:title>
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<treatment id="B603F0CDA91BF07A3BD299ED116948FD" ID-DOI="http://doi.org/10.5281/zenodo.10858763" ID-GBIF-Taxon="224054224" ID-Zenodo-Dep="10858763" LSID="urn:lsid:plazi:treatment:B603F0CDA91BF07A3BD299ED116948FD" httpUri="http://treatment.plazi.org/id/B603F0CDA91BF07A3BD299ED116948FD" lastPageNumber="318" pageNumber="318">
<subSubSection id="13A3CB5919C43B9D38166DC94184060A" pageNumber="318" type="nomenclature">
<paragraph id="450DE2A287A416FFA8A2713CCB75E285" pageNumber="318">
<taxonomicName id="67BFD3139F5504EFFE843A5BE030AB42" ID-CoL="3W7ZK" authority="L." authorityName="L." class="Magnoliopsida" family="Fabaceae" genus="Lotus" kingdom="Plantae" order="Fabales" pageNumber="390" phylum="Tracheophyta" rank="species" species="corniculatus">
<emphasis id="E9D87E7AE0E05E8127513F283FE30670" italics="true" pageNumber="318">Lotus corniculatus</emphasis>
L.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="03336399D277B370322BA8A902C33ADC" pageNumber="318" type="description">
<paragraph id="31E9E5A467A6686370789F48248447E2" pageNumber="318">
Artbeschreibung:
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hoch, aufsteigend bis aufrecht,
<emphasis id="68097E24E55066D5AEC8DAC0F343E886" bold="true" pageNumber="318">+/- kahl</emphasis>
(
<taxonomicName id="211C9DE14FE2C6BC036033CBBDC1ADDA" pageNumber="390" rank="variety" variety="corniculatus">
var.
<emphasis id="DB22126FBE0A8204870FA862366B5AF0" italics="true" pageNumber="318">corniculatus</emphasis>
</taxonomicName>
)
<emphasis id="93C21C37C9DFBF3882DAF1F5B3D3A188" bold="true" pageNumber="318">bis lang und dicht abstehend behaart</emphasis>
(
<taxonomicName id="E2944044B23C89CF3AB73D6554443AEF" pageNumber="390" rank="variety" variety="hirsutus">
var.
<emphasis id="00310C30DE1814BECC2952D9CFF747E3" italics="true" pageNumber="318">hirsutus</emphasis>
</taxonomicName>
),
<normalizedToken id="B419ED46AD4389318C28A26235D7AEF3" originalValue="Teilblätter" pageNumber="390">Teilblaetter</normalizedToken>
bis
<quantity id="02BFA7EE01E0BA7D868989F7AE7E580B" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageNumber="390" unit="cm" value="2.0">2 cm</quantity>
lang, 1-3mal so lang wie breit.
<emphasis id="B8BCB7D22D39887EDA046FDB972A6B46" bold="true" pageNumber="318">
<normalizedToken id="1B2B723074D326EAAA7E1A8CEC17EC05" originalValue="Blütenstand" pageNumber="390">Bluetenstand</normalizedToken>
2-8
<normalizedToken id="60D8B316264CA771A7888E773A2620BF" originalValue="blütig" pageNumber="390">bluetig</normalizedToken>
</emphasis>
. Kelch
<quantity id="FAD5E2847A28B5036916A77C2B20C0F6" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageNumber="390" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5-7 mm</quantity>
lang.
<normalizedToken id="7F7C52DA38BB67E3366D93EAEE677F4D" originalValue="Blüten" pageNumber="390">Blueten</normalizedToken>
<quantity id="692083125B0561686C8B9FD864A576EE" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.6" metricValueMin="0.7" pageNumber="390" unit="mm" value="11.5" valueMax="16.0" valueMin="7.0">7-16 mm</quantity>
lang.
<emphasis id="B07C34FABEF8AEC9AF628D8D136DEF9D" bold="true" pageNumber="318">Schiffchenspitze hell</emphasis>
, gelb oder
<normalizedToken id="F38DBAB35864AFE228A9694121699BAD" originalValue="rötlich" pageNumber="390">roetlich</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="74D3B041D0BA7833823F6EC9A3619AF7" pageNumber="318" type="biology_ecology">
<paragraph id="0ED0B639CD28D3F6E5D8627D455D90AA" pageNumber="318">
<normalizedToken id="0BF6DB0CA2DF93D9BEFDFE31A7332A7A" originalValue="Blütezeit" pageNumber="390">Bluetezeit</normalizedToken>
: 5-7
</paragraph>
<paragraph id="E3A7DD3C1EA96731264321D54CC0BE26" pageNumber="318">
Standort und Verbreitung in der Schweiz: Fettwiesen,
<normalizedToken id="CB783EAB2A7AB57DB6DDBA51D986C612" originalValue="Wegränder" pageNumber="390">Wegraender</normalizedToken>
/ kollin-subalpin(-alpin) / CH
</paragraph>
</subSubSection>
<subSubSection id="47A17A6AC9B3557114E3D7C227229510" pageNumber="318" type="distribution">
<paragraph id="0BD299BB9AD5A1D49717F42DF147E3BD" pageNumber="318">Verbreitung global: Eurosibirisch</paragraph>
</subSubSection>
<subSubSection id="0CF953D8F4CE84CC1CF81C5B7FB506EF" pageNumber="318" type="biology_ecology">
<paragraph id="41840AECF54A3E1846B65074776767E0" pageNumber="318">
<normalizedToken id="90F031A709417D288DE670BE84619E07" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="EC3E9CFBA349B84BD7CB50F4C2A6D5B4" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." pageNumber="318" publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="027043401874F61E35AD14B0098F88F6" pageNumber="318">
<table id="3B3DA8311978227ECF5D133C443A82FE" class="table" inLine="true" pageNumber="318">
<tr id="6C02E727940A884C061C745C20F93938" pageNumber="318">
<th id="B78015322089B91E78BCE68E5AC46CB3" colspan="2" pageNumber="318">
<emphasis id="04CC698A6DCD3FC0E0480462591C0380" bold="true" pageNumber="318">Bodenfaktoren</emphasis>
</th>
<th id="48783D371FF70140FF0CD70AB06747A5" colspan="2" pageNumber="318">
<emphasis id="9CB6923A350D5E5F12C4330065429649" bold="true" pageNumber="318">Klimafaktoren</emphasis>
</th>
<th id="669EB367F5995EA707A6A9FA3BBE2C29" colspan="2" pageNumber="318">
<emphasis id="ACAF146A81BA1F68A0BCE4968E70C728" bold="true" pageNumber="318">Salztoleranz</emphasis>
</th>
</tr>
<tr id="3A7A10A811B1EB29C60A0A7BF9E2169D" pageNumber="318">
<td id="00A299CF261BBA22F05C36056252D426" pageNumber="318">Feuchtezahl F</td>
<td id="1B93613F714E80CB19B5CEA7FAC50716" pageNumber="318" style="text-align:left;width:100px">--</td>
<td id="0E9F89AF389D9DA6B0B60B98FDD464D5" pageNumber="318">Lichtzahl L</td>
<td id="0C6D78AB497CEBB0CC80CE7DB9D1F8B8" pageNumber="318" style="text-align:left;width:60px">--</td>
<td id="98E6F1D263AD7B3745DB7C1E6B0A3E2D" pageNumber="318">Salzzeichen</td>
<td id="6BF93FEF270FB70669B277642060A050" pageNumber="318" style="text-align:left;width:60px">--</td>
</tr>
<tr id="3FFCD25928A9C80FB027691F8F71E229" pageNumber="318">
<td id="21775F36CFFF5E0842CB8D574F044B3A" pageNumber="318">Reaktionszahl R</td>
<td id="E99D4D7D02E89FBE23C8D53667819428" pageNumber="318" style="text-align:left">--</td>
<td id="C7CBE3E87356E9F17701CAB28AA07783" pageNumber="318">Temperaturzahl T</td>
<td id="21B87C2997D63009C574C9988C937B18" pageNumber="318" style="text-align:left">--</td>
</tr>
<tr id="9D4EC412F4B64600126D4FA3EA5EFE4E" pageNumber="318">
<td id="DA4AE881611B1744414DDBB0421FDA5E" pageNumber="318">
<normalizedToken id="BB31BCC31F7989DF5D3FF9F7871CA6EF" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="3D0A6042606DD27899F21243A372E986" pageNumber="318" style="text-align:left">--</td>
<td id="2F41EF447DE45E423108A3ED87459A81" pageNumber="318">
<normalizedToken id="DC577176A8F34AE68347480B24AC0EE8" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="666C64BCD526FFAD98D4C04C0FB14DCD" pageNumber="318" style="text-align:left">--</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="1341891F9835F6B68639B6AAE9B79AA5" pageNumber="318" type="vernacular_names">
<paragraph id="586EAB187025FA0A0506C37BCBFFC1AE" pageNumber="318">
Volksname Deutscher Name:
<vernacularName id="0A6A3CD40B53E7226DDBF04116E983B8" language="deu" pageNumber="318">
<normalizedToken id="C5DA3C03BBADAA6ECF991EFF8D08A2FE" originalValue="Gewöhnlicher">Gewoehnlicher</normalizedToken>
Hornklee
</vernacularName>
Nom
<normalizedToken id="9DBB315AE6B6882C3F4E5010856197C2" originalValue="français">francais</normalizedToken>
:
<vernacularName id="852FFD46091BB0258F2759908B924B18" language="fra" pageNumber="318">
Lotier
<normalizedToken id="67428DC17604B1B19CA8FA4DB9B726AE" originalValue="corniculé">cornicule</normalizedToken>
</vernacularName>
</paragraph>
</subSubSection>
</treatment>
</document>

225
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<treatment id="B604186B967C4EB5C6D6E70B00B563B9" ID-DOI="http://doi.org/10.5281/zenodo.10923456" ID-GBIF-Taxon="224778719" ID-Zenodo-Dep="10923456" LSID="urn:lsid:plazi:treatment:B604186B967C4EB5C6D6E70B00B563B9" httpUri="http://treatment.plazi.org/id/B604186B967C4EB5C6D6E70B00B563B9">
<subSubSection id="40E91E19286E9475BBA291D9638796C1" type="nomenclature">
<paragraph id="2B0421117FAB39C8B25BF83735F855E7">
<taxonomicName id="217CE2F005C87D76E1EA7CBFD7666A62" ID-CoL="6HTY9" authority="(Rohlena) Zielonk." authorityName="Zielonk." baseAuthorityName="Rohlena" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="subSpecies" species="pallens" subSpecies="scabrifolia">
<emphasis id="54D1A0B8318AE73A2DFF15B296062A9C" italics="true">Festuca pallens</emphasis>
subsp.
<emphasis id="EB3E13B2D5188EEE1F25C774C9FF978C" italics="true">scabrifolia</emphasis>
(Rohlena) Zielonk.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="6255856E1872FD00A3147D8FA8424506" type="multiple">
<paragraph id="80F7B775E4A24DD8F579237B35935861">
Unterart ISFS: 168900 Checklist: 1019350
<taxonomicName id="CAEB0AA9EE3CFF3525A3FDBD909739FA" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
<taxonomicName id="7FA825790010F3C009C60B241FDFCF87" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="genus">Festuca</taxonomicName>
<taxonomicName id="0E345349A13EC63A14D1A5300F31363E" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="ovina">Festuca ovina</taxonomicName>
superaggr.
<taxonomicName id="9015FAC9B3FB7B146FF1ADEBED93FD3A" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="ovina">Festuca ovina</taxonomicName>
aggr.
<taxonomicName id="4DAFD41534B1FC67ED5358C12BE1D1AC" authority="Host" authorityName="Host" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="pallens">Festuca pallens Host</taxonomicName>
<taxonomicName id="86A7FAD1D3639178E5B70318B1E067BC" authority="(Rohlena) Zielonk." authorityName="Zielonk." baseAuthorityName="Rohlena" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="subSpecies" species="pallens" subSpecies="scabrifolia">Festuca pallens subsp. scabrifolia (Rohlena) Zielonk.</taxonomicName>
</paragraph>
<paragraph id="B91093A9BB6DB057D6E0A0C2BE232C28">
<heading id="6A827FE280031D40A502F786CB33F40F">
<normalizedToken id="066084DCCE1A48C4B5A9444A42C97165" originalValue="Bestimmungsschlüssel">Bestimmungsschluessel</normalizedToken>
</heading>
</paragraph>
<paragraph id="BCEA60B9368F42F1B6C4EBAF5E12AC0B">
<heading id="6EE280931DBB66F2C8D1872BB0472085">Zusammenfassung</heading>
KEINE ANGABE
</paragraph>
<paragraph id="99E46FE59FCF17DDEF50F6DA7F186EC5">
<heading id="E9B5A411039C84B61EF41FA83AA7B489">
<normalizedToken id="8B21FA3627A56ECFFD44A4891D1EC846" originalValue="Ökologie">Oekologie</normalizedToken>
</heading>
</paragraph>
<paragraph id="9D4218B42FAA5B0E69A338D7381E8A95">
<heading id="262BE9119AF56BCE33E52118FC816DA0">Lebensraum Lebensraum</heading>
nach
<bibRefCitation id="C0698A28D62C5E4EF8F016BA9744230A" author="Delarze, R. &amp; Gonseth, Y. &amp; Eggenberger, S. &amp; Vust, M." location="Bad Hersfeld, Germany" publisher="Ott Verlag" refString="Delarze, R., Gonseth, Y., Eggenberger, S., Vust, M. 2015. Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition). Ott Verlag, Bad Hersfeld, Germany, 456 pp." title="Lebensräume der Schweiz: Oekologie – Gefährdung – Kennarten (3rd edition)" year="2015">Delarze &amp; al. 2015</bibRefCitation>
</paragraph>
<paragraph id="CED16DC04D6CCBF5FA100A8D548A1A4A">
<table id="6476D00EFDDFDCDCE7FA310897F91A72" class="table" inLine="true">
<tbody id="74218FA7427DEB06D3731EF371F69917">
<tr id="36E25AAF88320E5906A45A9FE007CBDC">
<td id="E0D3901ACF788D87C377D9B4B7DE7740">KEINE ANGABE</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="D987DD9F07889A6C334FC41B03832967" type="biology_ecology">
<paragraph id="C5831DF46509C518EFB9F4855F63A147">
<normalizedToken id="19731B95EBC67576702E01EE3D29D8C2" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="60B00982197004D81B177B504F61D0F0" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="B952A60D64BC036B8E244AD28A3A5F50">
<table id="E0C7CBF0E66A86FC520E3720422D78DA" class="table" inLine="true">
<tbody id="4C8408211CD5EA8EE53FA3AA98518C45">
<tr id="2F66FE9599F28DF23500929EE06DF46C">
<th id="0C96C34FC5DA11DD2133BAA16AF90CEF" colspan="2">
<emphasis id="00FAD077424CE24172BAF991639FDC57" bold="true">Bodenfaktoren</emphasis>
</th>
<th id="3C92ED4FE5C19DD4C1038CDF42FE9870" colspan="2">
<emphasis id="3D482D3DB6F00F031F773F49A89E829B" bold="true">Klimafaktoren</emphasis>
</th>
<th id="3D0E2957CBF2F7D50FA52AF75073BB48" colspan="2">
<emphasis id="ED4CBA0B7259E7692174C64F3D58B4DD" bold="true">Salztoleranz</emphasis>
</th>
</tr>
<tr id="0A08C1DC8FE4AD17DEBAB1052051DFB8">
<td id="87460D5BBDB5C25F1BFE1E357A9C0214">Feuchtezahl F</td>
<td id="1B7BE7BC3A2232318C3DC786F6BC6FF9" style="text-align:left;width:100px">--</td>
<td id="B221C6C46F85193982070CAFFB1A0086">Lichtzahl L</td>
<td id="E1C35BE478B7D8321C3B41DD86EE2F3E" style="text-align:left;width:60px">--</td>
<td id="D90E6BBBBE0C767AB94236CB0077511A">Salzzeichen</td>
<td id="56BC08657FF880556F90AF90B9C52698" style="text-align:left;width:60px">--</td>
</tr>
<tr id="28692E3EE6B9E1AF76C0DF6F97935D01">
<td id="5CA84E03F8B7DB40E0A5F53973F6C2A2">Reaktionszahl R</td>
<td id="CCE20280AB63355CBE1070008CE80AAF" style="text-align:left">--</td>
<td id="E3DC0B311CD640E1CF34422D8CE1FF09">Temperaturzahl T</td>
<td id="8DEAA87FE6730BFE2BE5C0418216E72A" style="text-align:left">--</td>
</tr>
<tr id="102EA18663446DD5E411F7CD670DBCE2">
<td id="1C648E90B6ED312175415264B59080AF">
<normalizedToken id="D7267EA45C54B36D15F3579ADB46A0DA" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="AE99B1E30562C3894421F8534D67EFA6" style="text-align:left">--</td>
<td id="CFF7ED919F84673DB8460CF0675D1633">
<normalizedToken id="C26EE1F862C27571DAB097273642CAED" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="2E36358CEB9F9F461CFDE77FD190539D" style="text-align:left">--</td>
</tr>
</tbody>
</table>
</paragraph>
</subSubSection>
<subSubSection id="1EE60C06821197DB25FCFDD9E774047A" type="reference_group">
<paragraph id="205E4B4BDA4687B3926A5B6EB1947EDA">
<heading id="6DDE735CCE05DDF9292E04259D3B6B4A">Nomenklatur</heading>
</paragraph>
<paragraph id="0B0D99B3C7DFDF24C6199C3C90424C40">
<treatmentCitation id="3C02CD92C21910B4E25C29E3962EC479">
<heading id="0714FBAFDB8305A2EA49669BDA1DCFAA">
<normalizedToken id="5A0EA706D0E36699D5B1674C8EEAB707" originalValue="Gültiger">Gueltiger</normalizedToken>
Name (
<bibRefCitation id="87E81DE202F84D28D816D73EBF4DEB41" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</heading>
:
<taxonomicName id="C56DEA5FBA03D1E6496D14F0406D06D1" ID-CoL="6HTY9" authority="(Rohlena) Zielonk." authorityName="Zielonk." baseAuthorityName="Rohlena" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="subSpecies" species="pallens" subSpecies="scabrifolia">
<emphasis id="E0209B80059651E496D8D32FC749AF6B" italics="true">Festuca pallens</emphasis>
subsp.
<emphasis id="FF86C3B562D604A96B9AE77814F34ACF" italics="true">scabrifolia</emphasis>
(Rohlena) Zielonk.
</taxonomicName>
</treatmentCitation>
</paragraph>
</subSubSection>
<subSubSection id="FE8CEE6A352005B7DF17B895D521DBAD" type="vernacular_names">
<paragraph id="FC7F290BC853445FA1F9F0DBFBB9B3CF">Volksname</paragraph>
</subSubSection>
<subSubSection id="0D6DBEA6F105B045D4C88A023A7CC8D7" type="multiple">
<paragraph id="7E14DCF145E61AD1F6AFE0F8BFFAC7B1">
Deutscher Name: -- Nom
<normalizedToken id="BDCF9BA2AA0E89729206AF258F882860" originalValue="français">francais</normalizedToken>
: -- Nome italiano: --
</paragraph>
</subSubSection>
<subSubSection id="38F4523F71FF79FEC8282E0238385CEF" type="reference_group">
<paragraph id="6E88C3CF0789C79FA2A2B8F4AF21F7A6">
<normalizedToken id="545C6B2192EF740C1BEEF8CE976F4DE3" originalValue="Übereinstimmung">Uebereinstimmung</normalizedToken>
mit anderen Referenzwerken
</paragraph>
<paragraph id="ACD31D80D9F396052616036F50FBAE2C">
<table id="49FB4D14379F134D9208E155BC78B952" class="table table-condensed" inLine="true">
<tbody id="59B66F434A0399A19E839A21C26DCFC2">
<tr id="BF146ED5B5307F5921CF40695083CD5B">
<th id="2D269CAC2522D1BDB5B2744CAEE77F6E" width="80px">Relation</th>
<th id="2FE6454B8A7EDC3C09567BA2C5C159AC">Nom</th>
<th id="8FF5EC946836E10481155DCF2906F035">Referenzwerke</th>
<th id="865B345BEA8890B8984F687A8724A610" width="90px">No</th>
</tr>
<tr id="057D9DCB69E50C320DD221BF654CD0B1">
<td id="9E144F45452A7B4E3DCACBB4D7840E80" class="taxon-relation">=</td>
<td id="AF0D7FA3321F082B8D67817EB4BA048E">
<taxonomicName id="2B4500AC655C8A823EAB14C7A3BE8E85" authority="(Rohlena) Zielonk." authorityName="Zielonk." baseAuthorityName="Rohlena" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="subSpecies" species="pallens" subSpecies="scabrifolia">Festuca pallens subsp. scabrifolia (Rohlena) Zielonk.</taxonomicName>
</td>
<td id="573721A4D66C459D0244BCF544B4CA7D">
<treatmentCitation id="A788EDDA62A05935A0C51A1305206BB5">
<bibRefCitation id="2EC5B078548F37842406514693F8B6FB">SISF/ISFS 2</bibRefCitation>
</treatmentCitation>
</td>
<td id="3D14F52F34037DACE276C94AAFEC9394">168900</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="BD37DC6E2D5EF88B6D6174200367E85E">
<tableNote id="C3460E43B5C2684172410C4767DB9819">
= Taxon stimmt mit akzeptiertem Taxon
<normalizedToken id="CDAB6997511C89E88158FCFF70674229" originalValue="überein">ueberein</normalizedToken>
(
<bibRefCitation id="566B1B9EF664DB8B5403EB640C200904" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) &lt;Taxon ist im akzeptierten Taxon (
<bibRefCitation id="4A2646A773B5BBD13FD8297ACED4E7A6" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
) enthalten&gt; Taxon
<normalizedToken id="86B93FFAA1CC722842BB0594356002AE" originalValue="enthält">enthaelt</normalizedToken>
(neben anderen) auch das akzeptierte Taxon (
<bibRefCitation id="F05F2D86C048ECEBC2A45C82532AFC5E" author="Juillerat, P. &amp; Bäumler, B. &amp; Bornand, C. &amp; Gygax, A. &amp; Jutzi, M. &amp; Möhl, A. &amp; Nyffeler, R. &amp; Sager, L. &amp; Santiago, H. &amp; Eggenberg, S." publicationUrl="https://doi.org/10.5281/zenodo.10680266" publisher="Info Flora" refString="Juillerat, P., Bäumler, B., Bornand, C., Gygax, A., Jutzi, M., Möhl, A., Nyffeler, R., Sager, L., Santiago, H., and Eggenberg, S. (2017). Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera. Info Flora. DOI: https://doi.org/10.5281/zenodo.10680266" title="Checklist 2017 der Gefässpflanzenflora der Schweiz / de la flore vasculaire de la Suisse / della flora vascolare della Svizzera" year="2017">Checklist 2017</bibRefCitation>
)
</tableNote>
</paragraph>
</subSubSection>
<subSubSection id="AEE7B8AD435E7F2266B8E616A60B5067" type="conservation">
<paragraph id="202CF513D026C0DE794EE83F9C8B512A">
<heading id="9B6FEBFBCBF6A4B344A6E6A9AE527F15">Status Indigenat</heading>
: -
</paragraph>
<paragraph id="98FA7465903F05763AC0EB7DF97FE1CC">
<emphasis id="A558A3ADC0487F5172C69F45FE057060" bold="true">
<heading id="EC2D764E3E5D0B8F22B98798E78469BC">
Liste der
<normalizedToken id="72D4B59A150C75CF226584AA8B3292AD" originalValue="gefährdeten">gefaehrdeten</normalizedToken>
Pflanzen IUCN
</heading>
(nach
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):
</emphasis>
Nein
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="3E5153555556418FBFBDC125C293CA71" authority="Hellen, 1948" authorityName="Hellen" authorityYear="1948" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora bifida" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="bifida">
Pristiphora bifida (
<normalizedToken id="9CA20AB520A0A17EC588886C954A5333" originalValue="Hellén">Hellen</normalizedToken>
, 1948)
</taxonomicName>
Figs 9, 25, 199, 267
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<subSubSection id="13CF444D617B1FB585F1DA207713CC72" pageId="64" pageNumber="65" type="reference_group">
<paragraph id="ED6AD92453380E2ABF5942FEBBE100F5" pageId="64" pageNumber="65">
<taxonomicName id="A7951B18551B99DDA28720156514302F" class="Insecta" family="Tenthredinidae" genus="Nematus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nematus (Pristiphora) bifidus" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="bifidus" subGenus="Pristiphora">Nematus (Pristiphora) bifidus</taxonomicName>
<normalizedToken id="619850561400A2CE3412CB7F373A314B" originalValue="Hellén">Hellen</normalizedToken>
, 1948b: 116-117. Lectotype ♀ (http://id.luomus.fi/GL.5214; designated by
<bibRefCitation id="81AE905C8B876F03D08CA9155E6B1149" author="Prous, M" journalOrPublisher="Journal of Hymenoptera Research" pageId="112" pageNumber="113" pagination="1 - 54" title="North-Western Palaearctic species of the Pristiphoraruficornis group (Hymenoptera, Tenthredinidae)." url="https://doi.org/10.3897/jhr.51.9162" volume="51" year="2016">Prous et al. 2016</bibRefCitation>
) in MZH, examined. Type locality: Malla,
<normalizedToken id="15E0A90E1091B799EF11D11F7775E34A" originalValue="Kilpisjärvi">Kilpisjaervi</normalizedToken>
,
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, Finland.
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<subSubSection id="11E007717BD2878684168FF813CCAC5A" pageId="64" pageNumber="65" type="similar species">
<paragraph id="EC17B320C2A7747AAD2AAFDA72D503C2" pageId="64" pageNumber="65">Similar species.</paragraph>
<paragraph id="307B31E1C413671620A8C5822F5624D6" pageId="64" pageNumber="65">
Externally, perhaps the most similar species is
<taxonomicName id="75D86CA0DB258BEC8F337B361853E7B2" lsidName="P. frigida" pageId="64" pageNumber="65" rank="species" species="frigida">P. frigida</taxonomicName>
, from which it can be distinguished by having pale hind trochanters, trochantelli, and tibiae (black or brown in
<taxonomicName id="771E6218DFDD277F363ECC84219ACEF2" lsidName="P. frigida" pageId="64" pageNumber="65" rank="species" species="frigida">P. frigida</taxonomicName>
). In addition, antennae of males have numerous and clearly visible stout black setae among finer paler ones (Fig. 18), while in
<taxonomicName id="9D59B6C93699151043D810F98B961A01" lsidName="P. frigida" pageId="64" pageNumber="65" rank="species" species="frigida">P. frigida</taxonomicName>
only a few black setae are barely visible (Fig. 17). The lancets (Figs 199-200) and penis valves (Figs 267, 274) are also different. The apical serrulae are somewhat shorter and more protruding, and the tangium of the lancet tends to be longer and narrower (Fig. 199) than in
<taxonomicName id="D070B7E5349AB14C2678EB61FC8D70BF" lsidName="P. frigida" pageId="64" pageNumber="65" rank="species" species="frigida">P. frigida</taxonomicName>
(Fig. 200). The penis valve lacks (Fig. 267) a membranous fold near the tip of the valvispina (present in
<taxonomicName id="7F1DD2B8D4F3B023B2E4A36F1693CD2F" lsidName="P. frigida" pageId="64" pageNumber="65" rank="species" species="frigida">P. frigida</taxonomicName>
; Fig. 274) and the pseudoceps has a distinct dorsal depression in the middle or basal part (absent in
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).
</paragraph>
</subSubSection>
<subSubSection id="EF34D4562C7079FE799D1C5324951DC7" pageId="64" pageNumber="65" type="genetic data">
<paragraph id="E4622AB7747C4BB0022C61EE12DA5746" pageId="64" pageNumber="65">Genetic data.</paragraph>
<paragraph id="C8D977A7ABB25B0B98C655A94588CF96" pageId="64" pageNumber="65">
Based on COI barcode sequences,
<taxonomicName id="CE610EC477FFAD67DD4A148AD64C2E1D" lsidName="P. bifida" pageId="64" pageNumber="65" rank="species" species="bifida">P. bifida</taxonomicName>
belongs to the same BIN cluster (BOLD:AAG3568) as
<taxonomicName id="E54E43C2F85576AC6971CCECAA330FE5" lsidName="P. aphantoneura" pageId="64" pageNumber="65" rank="species" species="aphantoneura">P. aphantoneura</taxonomicName>
,
<taxonomicName id="CAF6DC65499BE7B21C8126683AE8957F" lsidName="P. confusa" pageId="64" pageNumber="65" rank="species" species="confusa">P. confusa</taxonomicName>
,
<taxonomicName id="B7156697029D690E77095A162FCC759E" lsidName="P. luteipes" pageId="64" pageNumber="65" rank="species" species="luteipes">P. luteipes</taxonomicName>
,
<taxonomicName id="403A30D7C53BAB1BC93561C5AD324087" lsidName="P. opaca" pageId="64" pageNumber="65" rank="species" species="opaca">P. opaca</taxonomicName>
,
<taxonomicName id="7B7F5A32A6B28B81F2629948B4F8D65F" lsidName="P. pusilla" pageId="64" pageNumber="65" rank="species" species="pusilla">P. pusilla</taxonomicName>
,
<taxonomicName id="50DCE182F2D52A4BC5E48396DA7DF969" lsidName="P. staudingeri" pageId="64" pageNumber="65" rank="species" species="staudingeri">P. staudingeri</taxonomicName>
, and
<taxonomicName id="17287EF68D22A98287E1DE99ED01D75C" lsidName="P. subopaca" pageId="64" pageNumber="65" rank="species" species="subopaca">P. subopaca</taxonomicName>
(Fig. 4). Maximum distance within the BIN is 3.33% and minimum between species distance is 0.00%. The nearest neighbour to BOLD:AAG3568, diverging by minimum of 2.76%, is BOLD:AAQ2302 (
<taxonomicName id="CAB852D872BE5241356070D57B67A4AE" lsidName="P. armata" pageId="64" pageNumber="65" rank="species" species="armata">P. armata</taxonomicName>
and
<taxonomicName id="71A6B49720561A598A6B3BC65459A4EF" lsidName="P. leucopus" pageId="64" pageNumber="65" rank="species" species="leucopus">P. leucopus</taxonomicName>
). Based on nuclear data, maximum within species divergence is 0.1% (based on two specimens and both genes combined) and the nearest neighbour is 0.0% different (
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,
<taxonomicName id="53DEE75C3A471CBC067730399F0DDD02" lsidName="P. luteipes" pageId="64" pageNumber="65" rank="species" species="luteipes">P. luteipes</taxonomicName>
, or
<taxonomicName id="0D8C222B24F2F802BA2EE7B9B7D6049B" lsidName="P. beaumonti" pageId="64" pageNumber="65" rank="species" species="beaumonti">P. beaumonti</taxonomicName>
, only TPI). When including TPI introns, the nearest neighbour is still 0.0% different (
<taxonomicName id="6F5F6E638B3613731E2B4A0D003E7A73" lsidName="P. confusa" pageId="64" pageNumber="65" rank="species" species="confusa">P. confusa</taxonomicName>
or
<taxonomicName id="7FB1C43B59AAECCAF70C7C179BB99418" lsidName="P. luteipes" pageId="64" pageNumber="65" rank="species" species="luteipes">P. luteipes</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="0EAFBAEF395E23022E6A3817CA97417C" pageId="64" pageNumber="65" type="host">
<paragraph id="130B0FF4AD83D0FB3A3E73A8D2EB0CB4" pageId="64" pageNumber="65">Host plants.</paragraph>
<paragraph id="4BFEE6D4E00C2092E7ABBAA02FF107E8" pageId="64" pageNumber="65">
<taxonomicName id="6E196DA684F8139B9FFE756324E70470" class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix viminalis" order="Malpighiales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="viminalis">Salix viminalis</taxonomicName>
L. (
<bibRefCitation id="EEAAD8A339B0B04D3CEFCCEBE9BF2D15" author="Liston, AD" journalOrPublisher="Nachrichtenblatt der Bayerischen Entomologen" pageId="109" pageNumber="110" pagination="34 - 37" title="Recent records of remarkable sawflies in Bavaria." volume="58" year="2009">Liston and Burger 2009</bibRefCitation>
). In
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(Finland) some other species must be the host, as
<taxonomicName id="FD8F6E4003D7952934B001206F8238A7" lsidName="S. viminalis" pageId="64" pageNumber="65" rank="species" species="viminalis">S. viminalis</taxonomicName>
does not occur there.
</paragraph>
</subSubSection>
<subSubSection id="D1506A0CACA0A5143FFE4B2BB86289D8" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="C1BF7A01D10761B20612D8BE2759A1AA" pageId="64" pageNumber="65">Distribution and material examined.</paragraph>
<paragraph id="C0D33ED4E182403D2F6372DEB4D55D5E" pageId="64" pageNumber="65">West Palaearctic. Specimens studied are from Finland, Germany, Norway, and Sweden.</paragraph>
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Type status:
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. Occurrence: recordedBy:
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; individualCount:
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; sex:
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; Location: country:
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; stateProvince: Nylandia; municipality: Helsinki; locality:
<location LSID="urn:lsid:plazi:treatment:B6051F4B46BBC8315A2DB73065C04070:B945D86088FE4BFD04B33B655687B32E" country="Finland" latitude="60.261" longitude="24.965" name="Tuomarinkylae" pageId="0" pageNumber="1068">
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; eventDate:
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; habitat: Wood-storage areas in Helsinki; Record Level: institutionCode:
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<paragraph pageId="0" pageNumber="1068">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1068">
European.
<taxonomicName class="Insecta" family="Mycetophilidae" genus="Mycetophila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mycetophila distigma" order="Diptera" pageId="0" pageNumber="1068" phylum="Arthropoda" rank="species" species="distigma">Mycetophila distigma</taxonomicName>
(Fig. 21) is recorded from central and northern Europe (
<bibRefCitation pageId="0" pageNumber="1068">Chandler 2004</bibRefCitation>
). In Fennoscandia known from Sweden (Lule Lapmark,
<bibRefCitation author="Kjaerandsen, J." journalOrPublisher="Insect Systematics &amp; Evolution, Supplement" pageId="0" pageNumber="1068" pagination="1 - 165" title="Annotated checklist of fungus gnats from Sweden (Diptera: Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae and Mycetophilidae)" volume="65" year="2007">Kjaerandsen et al. 2007</bibRefCitation>
) and Norway (Akershus,
<bibRefCitation author="Soli, G." journalOrPublisher="Norwegian Journal of Entomology" pageId="0" pageNumber="1068" pagination="69 - 73" title="New records of fungus gnats for Norway (Diptera, Mycetophilidae)" volume="56" year="2009">Soli et al. 2009</bibRefCitation>
). New for the Finnish fauna.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1068" type="ecology">
<paragraph pageId="0" pageNumber="1068">Ecology</paragraph>
<paragraph pageId="0" pageNumber="1068">
Finnish specimens were collected in wood-storage areas in the city parks of Helsinki. Probably a saproxylic species, reared by
<normalizedToken originalValue="Ševčík">Sevcik</normalizedToken>
(
<bibRefCitation author="Sevcik, J." journalOrPublisher="Silesian Museum, Opava" pageId="0" pageNumber="1068" title="Czech and Slovak Diptera associated with fungi" year="2010">
<normalizedToken originalValue="Ševčík">Sevcik</normalizedToken>
2010
</bibRefCitation>
) from the polypore fungus
<taxonomicName class="Agaricomycetes" family="Meruliaceae" genus="Bjerkandera" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Bjerkandera adusta" order="Polyporales" pageId="0" pageNumber="1068" phylum="Basidiomycota" rank="species" species="adusta">Bjerkandera adusta</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

905
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and
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<emphasis id="0CD8EA85FFAF114E1B76CE2A673F6D27" box="[106,313,1034,1056]" italics="true" pageId="5" pageNumber="180">Phyllopertha campestris</emphasis>
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<emphasis id="0CD8EA85FFAF114E1B76CE07673F6D3A" box="[106,313,1063,1085]" italics="true" pageId="5" pageNumber="180">Phyllopertha campestris</emphasis>
var.
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(
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locality: Lyon),
<emphasis id="0CD8EA85FFAF114E1859CE07659E6D3B" bold="true" box="[837,920,1063,1084]" pageId="5" pageNumber="180">syn. nov.</emphasis>
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<treatmentCitation id="BF0D1086FFAF114E1B76CE6464856D5D" author="MULSANT, E." box="[106,643,1092,1114]" page="496" pageId="5" pageNumber="180" year="1842">
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<emphasis id="0CD8EA85FFAF114E1B76CE64673F6D5D" box="[106,313,1092,1114]" italics="true" pageId="5" pageNumber="180">Phyllopertha campestris</emphasis>
var.
<emphasis id="0CD8EA85FFAF114E1A79CE6467B16D5E" box="[357,439,1092,1113]" italics="true" pageId="5" pageNumber="180">maculata</emphasis>
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(
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<treatmentCitationGroup id="1EBC11B9FFAF114E1B76CE4165AE6D71" box="[106,936,1121,1143]" pageId="5" pageNumber="180">
<treatmentCitation id="BF0D1086FFAF114E1B76CE41648F6D70" author="MULSANT, E." box="[106,649,1121,1143]" page="496" pageId="5" pageNumber="180" year="1842">
<taxonomicName id="F9AC4D14FFAF114E1B76CE41648F6D70" authority="MULSANT, 1842: 496" authorityName="MULSANT" authorityPageNumber="496" authorityYear="1842" box="[106,649,1121,1143]" class="Insecta" family="Rutelidae" genus="Phyllopertha" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="variety" species="campestris" variety="pauperata">
<emphasis id="0CD8EA85FFAF114E1B76CE41673F6D71" box="[106,313,1121,1142]" italics="true" pageId="5" pageNumber="180">Phyllopertha campestris</emphasis>
var.
<emphasis id="0CD8EA85FFAF114E1A79CE4167BB6D71" box="[357,445,1121,1142]" italics="true" pageId="5" pageNumber="180">pauperata</emphasis>
<bibRefCitation id="5A3D4B66FFAF114E1ADFCE41648F6D70" author="MULSANT, E." box="[451,649,1121,1143]" pageId="5" pageNumber="180" refId="ref7062" refString="MULSANT, E. 1842: Histoire naturelle des Coleopteres de France. Lamellicornes. - Paris: Maison Libraire, Lyon: Imprimerie de Dumoulin, Ronet et Sibuet: viii + 626 pp., 3 pls." type="book" year="1842">MULSANT, 1842: 496</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="E1178835FFAF114E1989CE4164B86D71" box="[661,702,1121,1142]" pageId="5" pageNumber="180">type</typeStatus>
locality: Lyon),
<emphasis id="0CD8EA85FFAF114E1849CE4165AE6D71" bold="true" box="[853,936,1121,1142]" pageId="5" pageNumber="180">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="76B6651CFFAF11431B76CEAE623B6F10" lastPageId="8" lastPageNumber="183" pageId="5" pageNumber="180" type="materials_examined">
<paragraph id="3E133697FFAF114E1B76CEAE67246DA3" blockId="5.[106,290,1166,1188]" box="[106,290,1166,1188]" pageId="5" pageNumber="180">
<emphasis id="0CD8EA85FFAF114E1B76CEAE67246DA3" bold="true" box="[106,290,1166,1188]" pageId="5" pageNumber="180">Examined material:</emphasis>
</paragraph>
<paragraph id="3E133697FFAF114E1B76CE93623A6DE1" blockId="5.[106,1097,1202,1657]" pageId="5" pageNumber="180">
<materialsCitation id="8EC43CCAFFAF114E1B76CE9364336DCE" box="[106,565,1202,1225]" collectionCode="MHNG" country="France" location="Savoie" pageId="5" pageNumber="180" specimenCount="1">
<emphasis id="0CD8EA85FFAF114E1B76CE9366AB6DCF" bold="true" box="[106,173,1203,1224]" pageId="5" pageNumber="180">
<collectingCountry id="46BB7607FFAF114E1B76CE9366A16DCF" box="[106,167,1203,1224]" name="France" pageId="5" pageNumber="180">France</collectingCountry>
:
</emphasis>
„
<location id="3B73604CFFAF114E1BA6CE9366F06DCE" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1BA6CE9366F06DCE" box="[186,246,1203,1225]" country="France" name="Savoie" pageId="5" pageNumber="180">Savoie</location>
, [illegible:] Voison“ (1 ,
<collectionCode id="58BDAE52FFAF114E1AC2CE9364286DCE" box="[478,558,1203,1225]" country="Switzerland" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="180" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1952CE93654B6DCF" box="[590,845,1203,1225]" collectionCode="MNHB" country="France" location="Frankreich" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E194ACE9364BF6DCE" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E194ACE9364BF6DCE" box="[598,697,1203,1225]" country="France" name="Frankreich" pageId="5" pageNumber="180">Frankreich</location>
“ (1 ,
<collectionCode id="58BDAE52FFAF114E19E5CE9365406DCF" box="[761,838,1203,1224]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E187ACE9362436DCF" box="[870,1093,1203,1225]" collectionCode="MNHB" country="France" location="Lyon" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E1872CE93659B6DCF" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1872CE93659B6DCF" box="[878,925,1203,1224]" country="France" name="Lyon" pageId="5" pageNumber="180">Lyon</location>
“ (3,
<collectionCode id="58BDAE52FFAF114E18EDCE9362386DCF" box="[1009,1086,1203,1224]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1B67CEF0623E6DE1" box="[123,1080,1232,1254]" collectingDate="1966-06" collectionCode="SMTD" collectorName="Fehse" country="France" location="Basses Alpes" municipality="St. Andire" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E1B9FCEF066F36DE1" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1B9FCEF066F36DE1" box="[131,245,1232,1254]" country="France" municipality="St. Andire" name="Basses Alpes" pageId="5" pageNumber="180">Basses Alpes</location>
,
<collectingMunicipality id="DE77ACEDFFAF114E1BE2CEF0675A6DE1" box="[254,348,1232,1254]" pageId="5" pageNumber="180">St. Andire</collectingMunicipality>
,
<date id="4A121057FFAF114E1A79CEF067B96DE1" box="[357,447,1232,1254]" pageId="5" pageNumber="180" value="1966-06">
<collectingDate id="5A56E9BFFFAF114E1A79CEF067B96DE1" box="[357,447,1232,1254]" pageId="5" pageNumber="180" value="1966-06">Juni 1966</collectingDate>
</date>
“ (3, 6, CJS). – „Gall. mer.“ (3, 1 , coll.
<collectorName id="93595341FFAF114E18B1CEF065E46DE1" box="[941,994,1232,1254]" pageId="5" pageNumber="180">Fehse</collectorName>
,
<collectionCode id="58BDAE52FFAF114E18F6CEF062376DE1" box="[1002,1073,1232,1254]" pageId="5" pageNumber="180">SMTD</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="3E133697FFAF114E1B76CECD67EF6F7E" blockId="5.[106,1097,1202,1657]" pageId="5" pageNumber="180">
<materialsCitation id="8EC43CCAFFAF114E1B76CECD64C96C05" box="[106,719,1260,1282]" collectionCode="MNHB" country="Italy" location="Tirol" pageId="5" pageNumber="180" specimenCount="1">
<emphasis id="0CD8EA85FFAF114E1B76CECD669C6C05" bold="true" box="[106,154,1261,1282]" pageId="5" pageNumber="180">
<collectingCountry id="46BB7607FFAF114E1B76CECD66936C05" box="[106,149,1261,1282]" name="Italy" pageId="5" pageNumber="180">Italy</collectingCountry>
:
</emphasis>
„
<location id="3B73604CFFAF114E1BB7CECC66DC6C05" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1BB7CECC66DC6C05" box="[171,218,1260,1282]" country="Italy" name="Tirol" pageId="5" pageNumber="180">Tirol</location>
mer. | 24645“ (3 , historical collection,
<collectionCode id="58BDAE52FFAF114E1966CECD64CE6C05" box="[634,712,1261,1282]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E19F1CECC65426C18" collectingDate="1931-06" collectionCode="MNHB" collectorName="Schadewald" country="Italy" location="Sudtirol" pageId="5" pageNumber="180" specimenCount="2">
„
<location id="3B73604CFFAF114E19EACECC65456C05" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E19EACECC65456C05" box="[758,835,1260,1282]" country="Italy" name="Sudtirol" pageId="5" pageNumber="180">Südtirol</location>
,
<date id="4A121057FFAF114E1852CECD65996C05" box="[846,927,1261,1282]" pageId="5" pageNumber="180" value="1931-06">
<collectingDate id="5A56E9BFFFAF114E1852CECD65996C05" box="[846,927,1261,1282]" pageId="5" pageNumber="180" value="1931-06">VI.1931</collectingDate>
</date>
, leg.
<collectorName id="93595341FFAF114E18CDCECC62396C05" box="[977,1087,1260,1282]" pageId="5" pageNumber="180">Schadewald</collectorName>
“ (
<specimenCount id="28AAFD1EFFAF114E1B6DCF2966A16C18" box="[113,167,1289,1311]" pageId="5" pageNumber="180" type="generic">2 Ex.</specimenCount>
, CAW). – „Tirol | 24645“ (2, 1 , historical collection,
<collectionCode id="58BDAE52FFAF114E19F3CF2A653B6C18" box="[751,829,1290,1311]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1878CF2967B86C3B" collectingDate="1931-06" collectionCode="MNHB" collectorName="Schadewald" country="Italy" isTailEnumeration="true" location="Tyrol. Stentz." pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E1871CF2965EB6C19" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1871CF2965EB6C19" box="[877,1005,1289,1311]" country="Italy" name="Tyrol. Stentz." pageId="5" pageNumber="180">Tyrol. Stentz.</location>
| 24645“ (1 , historical collection,
<collectionCode id="58BDAE52FFAF114E1A74CF0767B06C3B" box="[360,438,1319,1340]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. – „24645“ (5 , 4 , historical collection,
<collectionCode id="58BDAE52FFAF114E1897CF0765DF6C3B" box="[907,985,1319,1340]" pageId="5" pageNumber="180">MNHB</collectionCode>
). –
<materialsCitation id="8EC43CCAFFAF114E1F1DCF0667576C5F" collectionCode="MNHB" country="Italy" location="Schmidt" municipality="Trient" pageId="5" pageNumber="180" specimenCount="1">
„
<collectingMunicipality id="DE77ACEDFFAF114E1F15CF0662426C3B" box="[1033,1092,1318,1340]" pageId="5" pageNumber="180">Trient</collectingMunicipality>
,
<location id="3B73604CFFAF114E1B76CF6366BF6C5E" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1B76CF6366BF6C5E" box="[106,185,1347,1369]" country="Italy" municipality="Trient" name="Schmidt" pageId="5" pageNumber="180">Schmidt</location>
“ (1 ,
<collectionCode id="58BDAE52FFAF114E1BE0CF63674C6C5F" box="[252,330,1347,1368]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1A76CF6365DD6C5F" box="[362,987,1347,1369]" collectionCode="MNHB" collectorName="Coll. Thieme" country="Italy" location="Trient" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E1A6FCF6367AA6C5E" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1A6FCF6367AA6C5E" box="[371,428,1347,1369]" country="Italy" name="Trient" pageId="5" pageNumber="180">Trient</location>
| 89946 |
<collectorName id="93595341FFAF114E1914CF6364826C5E" box="[520,644,1347,1369]" pageId="5" pageNumber="180">Coll. Thieme</collectorName>
“ (1 , historical collection,
<collectionCode id="58BDAE52FFAF114E1899CF6365D56C5F" box="[901,979,1347,1368]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E18E8CF63673F6C72" collectionCode="MNHB" country="Italy" location="Verona" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E18E0CF6362396C5F" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E18E0CF6362396C5F" box="[1020,1087,1347,1368]" country="Italy" name="Verona" pageId="5" pageNumber="180">Verona</location>
“ (3 , 1 ,
<collectionCode id="58BDAE52FFAF114E1BF8CF4067346C72" box="[228,306,1376,1397]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1A44CF4065186C72" box="[344,798,1375,1398]" collectionCode="MNHB" country="Italy" location="Toscana" pageId="5" pageNumber="180" specimenCount="1" stateProvince="Toscana">
„
<collectingRegion id="FC68F875FFAF114E1A7DCF4067A86C72" box="[353,430,1376,1397]" country="Italy" name="Toscana" pageId="5" pageNumber="180">Toscana</collectingRegion>
, [illegible:] Sinca | 69“ (1 ,
<collectionCode id="58BDAE52FFAF114E19D4CF4065106C72" box="[712,790,1376,1397]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1821CF4062376C72" box="[829,1073,1376,1397]" collectionCode="MNHB" country="Italy" location="Bozen" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E185ACF4065876C72" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E185ACF4065876C72" box="[838,897,1376,1397]" country="Italy" name="Bozen" pageId="5" pageNumber="180">Bozen</location>
“ (3 ,
<collectionCode id="58BDAE52FFAF114E18C0CF40622C6C72" box="[988,1066,1376,1397]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1B76CF5D643D6C94" box="[106,571,1404,1427]" collectionCode="NSMW" collectorName="Kollektion Dr. Fuchs" country="Italy" location="Bozen" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E1B6ECF5D66AA6C95" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1B6ECF5D66AA6C95" box="[114,172,1405,1426]" country="Italy" name="Bozen" pageId="5" pageNumber="180">Bozen</location>
|
<collectorName id="93595341FFAF114E1BDECF5C678F6C95" box="[194,393,1404,1426]" pageId="5" pageNumber="180">Kollektion Dr. Fuchs</collectorName>
“ (3 ,
<collectionCode id="58BDAE52FFAF114E1AF8CF5D64326C94" box="[484,564,1405,1427]" country="Germany" httpUri="http://grbio.org/cool/ahw9-s8iw" name="Naturwissenschftlich Sammlung, Museum Wiesbaden" pageId="5" pageNumber="180">NSMW</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1945CF5D66FD6CA8" collectionCode="NSMW" collectorName="W. Maus" country="Italy" county="Atzwang" location="Renon" municipality="Sudtirol" pageId="5" pageNumber="180" specimenCount="1">
„
<collectingCounty id="D7724E1BFFAF114E197ECF5D64B56C95" box="[610,691,1405,1426]" pageId="5" pageNumber="180">Atzwang</collectingCounty>
[
<collectingMunicipality id="DE77ACEDFFAF114E19DCCF5C65096C95" box="[704,783,1404,1426]" pageId="5" pageNumber="180">Südtirol</collectingMunicipality>
, zu
<location id="3B73604CFFAF114E1824CF5C657E6C95" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1824CF5C657E6C95" box="[824,888,1404,1426]" country="Italy" county="Atzwang" municipality="Sudtirol" name="Renon" pageId="5" pageNumber="180">Renon</location>
] 7.1910, g.
<collectorName id="93595341FFAF114E18F7CF5D62396C95" box="[1003,1087,1405,1426]" pageId="5" pageNumber="180">W. Maus</collectorName>
“ (1 ,
<collectionCode id="58BDAE52FFAF114E1BBFCFB966F26CA8" box="[163,244,1433,1455]" country="Germany" httpUri="http://grbio.org/cool/ahw9-s8iw" name="Naturwissenschftlich Sammlung, Museum Wiesbaden" pageId="5" pageNumber="180">NSMW</collectionCode>
)
</materialsCitation>
.-
<materialsCitation id="8EC43CCAFFAF114E1A12CFB9659F6CA8" box="[270,921,1433,1455]" collectionCode="MNHB" collectorName="A. Fiori" country="Italy" location="Emilia" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E1A0ACFB967506CA8" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1A0ACFB967506CA8" box="[278,342,1433,1455]" country="Italy" name="Emilia" pageId="5" pageNumber="180">Emilia</location>
, Fiumalbo, VI.[18]94,
<collectorName id="93595341FFAF114E1925CFB964856CA8" box="[569,643,1433,1455]" pageId="5" pageNumber="180">A. Fiori</collectorName>
| 110616“ (3 ,
<collectionCode id="58BDAE52FFAF114E1858CFBA65946CA8" box="[836,914,1434,1455]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E18A7CFB964366CCB" collectionCode="MNHB" collectorName="A. Fiori" country="Italy" location="Reno" municipality="Emilia" pageId="5" pageNumber="180" specimenCount="1">
„
<collectingMunicipality id="DE77ACEDFFAF114E18D8CFB962026CA8" box="[964,1028,1433,1455]" pageId="5" pageNumber="180">Emilia</collectingMunicipality>
,
<location id="3B73604CFFAF114E1F0DCFBA62426CA8" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1F0DCFBA62426CA8" box="[1041,1092,1434,1455]" country="Italy" municipality="Emilia" name="Reno" pageId="5" pageNumber="180">Reno</location>
, 6.VI.[18]90,
<collectorName id="93595341FFAF114E1BF4CF9667346CCB" box="[232,306,1462,1484]" pageId="5" pageNumber="180">A. Fiori</collectorName>
| 110617“ (1 ,
<collectionCode id="58BDAE52FFAF114E1AC7CF97642F6CCB" box="[475,553,1463,1484]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E194DCF9666B96CEF" collectionCode="MNHB" collectorName="A. Fiori" country="Italy" location="Rimini" municipality="Emilia" pageId="5" pageNumber="180" specimenCount="1">
„
<collectingMunicipality id="DE77ACEDFFAF114E1945CF96649F6CCB" box="[601,665,1462,1484]" pageId="5" pageNumber="180">Emilia</collectingMunicipality>
,
<location id="3B73604CFFAF114E19B9CF9664EF6CCB" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E19B9CF9664EF6CCB" box="[677,745,1462,1484]" country="Italy" municipality="Emilia" name="Rimini" pageId="5" pageNumber="180">Rimini</location>
, 6. [19]11,
<collectorName id="93595341FFAF114E1842CF9665A16CCB" box="[862,935,1462,1484]" pageId="5" pageNumber="180">A. Fiori</collectorName>
| 110618“ (1 ,
<collectionCode id="58BDAE52FFAF114E1B76CFF366BE6CEF" box="[106,184,1491,1512]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1BC7CFF365A96CEF" box="[219,943,1490,1513]" collectionCode="MNHB" collectorName="A. Fiori" country="Italy" location="Piemonte" pageId="5" pageNumber="180" specimenCount="1" stateProvince="Piemonte">
„
<collectingRegion id="FC68F875FFAF114E1BF8CFF367466CEE" box="[228,320,1491,1513]" country="Italy" name="Piemonte" pageId="5" pageNumber="180">Piemonte</collectingRegion>
, [illegible:] Garearel, VI.[18]90,
<collectorName id="93595341FFAF114E196DCFF364BE6CEE" box="[625,696,1491,1513]" pageId="5" pageNumber="180">A. Fiori</collectorName>
| 110619“ (1 ,
<collectionCode id="58BDAE52FFAF114E1845CFF365A16CEF" box="[857,935,1491,1512]" pageId="5" pageNumber="180">MNHB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E18D7CFF367556F01" collectionCode="SMTD" country="Italy" location="Trient" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E18CFCFF362086CEE" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E18CFCFF362086CEE" box="[979,1038,1491,1513]" country="Italy" name="Trient" pageId="5" pageNumber="180">Trient</location>
, [leg.] Kirsch“ (2,
<collectionCode id="58BDAE52FFAF114E1A18CFD0674A6F01" box="[260,332,1520,1542]" pageId="5" pageNumber="180">SMTD</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1A68CFD065206F01" box="[372,806,1519,1542]" collectionCode="SMTD" country="Italy" location="Tirol" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E1A61CFD067A86F01" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1A61CFD067A86F01" box="[381,430,1520,1542]" country="Italy" name="Tirol" pageId="5" pageNumber="180">Tirol</location>
, [leg.] Märkel“ (3, 3 ,
<collectionCode id="58BDAE52FFAF114E19CBCFD065196F01" box="[727,799,1520,1542]" pageId="5" pageNumber="180">SMTD</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1854CFD0679D6F24" collectingDate="1903-06" collectionCode="SMTD" collectorName="Dr. Noesske" country="Italy" location="Bozen" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E184CCFD0658A6F02" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E184CCFD0658A6F02" box="[848,908,1520,1541]" country="Italy" name="Bozen" pageId="5" pageNumber="180">Bozen</location>
,
<date id="4A121057FFAF114E1885CFD065EE6F02" box="[921,1000,1520,1541]" pageId="5" pageNumber="180" value="1903-06">
<collectingDate id="5A56E9BFFFAF114E1885CFD065EE6F02" box="[921,1000,1520,1541]" pageId="5" pageNumber="180" value="1903-06">VI.1903</collectingDate>
</date>
[leg.]
<collectorName id="93595341FFAF114E1F35CFD066B16F25" pageId="5" pageNumber="180">Dr. Noesske</collectorName>
“ (2 , 2 ,
<collectionCode id="58BDAE52FFAF114E1A50CC2D67926F24" box="[332,404,1549,1571]" pageId="5" pageNumber="180">SMTD</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1AA9CC2C65136F24" box="[437,789,1548,1571]" collectionCode="SMTD" country="Italy" location="Italia" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E1AA2CC2C67E96F25" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1AA2CC2C67E96F25" box="[446,495,1548,1570]" country="Italy" name="Italia" pageId="5" pageNumber="180">Italia</location>
, [leg.] Märkel“ (2,
<collectionCode id="58BDAE52FFAF114E19DACC2D65086F24" box="[710,782,1549,1571]" pageId="5" pageNumber="180">SMTD</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1833CC2C66DC6F38" collectionCode="ZSM" country="Italy" location="Tirolis" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E1824CC2C65736F25" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1824CC2C65736F25" box="[824,885,1548,1570]" country="Italy" name="Tirolis" pageId="5" pageNumber="180">Tirolis</location>
mer. | alte Sammlung“ (1 ,
<collectionCode id="58BDAE52FFAF114E1B82CC0966D56F38" box="[158,211,1577,1599]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="180" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1BEFCC0A64A76F38" box="[243,673,1577,1599]" collectionCode="ZSM" collectorName="Sammlung Cl. Muller" country="Italy" location="Bozen" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E1BE0CC0A67306F38" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1BE0CC0A67306F38" box="[252,310,1578,1599]" country="Italy" name="Bozen" pageId="5" pageNumber="180">Bozen</location>
|
<collectorName id="93595341FFAF114E1A5ACC0964176F38" box="[326,529,1577,1599]" pageId="5" pageNumber="180">Sammlung Cl. Müller</collectorName>
“ (2,
<collectionCode id="58BDAE52FFAF114E197ACC09649D6F38" box="[614,667,1577,1599]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="180" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E19A6CC0A672D6F5B" collectionCode="ZSM" collectorName="Sammlung J. Daniel" country="Italy" location="Bozen" pageId="5" pageNumber="180" specimenCount="1">
„
<location id="3B73604CFFAF114E19DFCC0A64FB6F38" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E19DFCC0A64FB6F38" box="[707,765,1578,1599]" country="Italy" name="Bozen" pageId="5" pageNumber="180">Bozen</location>
4.6.[18]84 J |
<collectorName id="93595341FFAF114E189FCC0962396F38" box="[899,1087,1577,1599]" pageId="5" pageNumber="180">Sammlung J. Daniel</collectorName>
“ (2 , 2 ,
<collectionCode id="58BDAE52FFAF114E1BF3CC6667226F5B" box="[239,292,1606,1628]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="180" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFAF114E1A5ECC6667E36F7E" collectingDate="1992-05-29" collectingDateMax="1992-05-31" collectingDateMin="1992-05-29" collectionCode="ZSM, CCZ" collectorName="Zorn" country="Italy" elevation="700" location="Fundortetiketten" pageId="5" pageNumber="180" specimenCount="1">
ohne
<location id="3B73604CFFAF114E1A68CC6664136F5B" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAF114E1A68CC6664136F5B" box="[372,533,1606,1628]" country="Italy" name="Fundortetiketten" pageId="5" pageNumber="180">Fundortetiketten</location>
: 4 , 5 ,
<collectionCode id="58BDAE52FFAF114E1985CC6664CB6F5C" box="[665,717,1606,1627]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="180" type="Museum">ZSM</collectionCode>
. – „Italien, Meeralpen, Entraque&gt;
<quantity id="F9549B72FFAF114E1F1ACC6662426F5B" box="[1030,1092,1606,1628]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="180" unit="m" value="700.0">
<elevation id="B581D1A4FFAF114E1F1ACC6662426F5B" box="[1030,1092,1606,1628]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="180" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="4A121057FFAF114E1B76CC4366F56F7F" box="[106,243,1635,1656]" pageId="5" pageNumber="180" value="1992-05-29" valueMax="1992-05-31" valueMin="1992-05-29">
<collectingDate id="5A56E9BFFFAF114E1B76CC4366F56F7F" box="[106,243,1635,1656]" pageId="5" pageNumber="180" value="1992-05-29" valueMax="1992-05-31" valueMin="1992-05-29">29.-31.V.1992</collectingDate>
</date>
, leg.
<collectorName id="93595341FFAF114E1A3FCC4367556F7F" box="[291,339,1635,1656]" pageId="5" pageNumber="180">Zorn</collectorName>
“ (2 ,
<collectionCode id="58BDAE52FFAF114E1AB6CC4367D86F7E" box="[426,478,1635,1657]" pageId="5" pageNumber="180">CCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="3E133697FFAC114D1BA1CA8F66FB69E5" blockId="6.[189,1180,175,341]" pageId="6" pageNumber="181">
<emphasis id="0CD8EA85FFAC114D1BA1CA8F673469C3" bold="true" box="[189,306,175,197]" pageId="6" pageNumber="181">
<collectingCountry id="46BB7607FFAC114D1BA1CA8F672A69C2" box="[189,300,175,197]" name="Switzerland" pageId="6" pageNumber="181">Switzerland</collectingCountry>
:
</emphasis>
„Helvet. | 24645“ (1 , historical collection,
<collectionCode id="58BDAE52FFAC114D19F0CA8F653C69C3" box="[748,826,175,196]" pageId="6" pageNumber="181">MNHB</collectionCode>
). – „Helvetia | alte Sammlung“ (1 ,
<collectionCode id="58BDAE52FFAC114D1BA1CAEC66F469E5" box="[189,242,204,226]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="181" type="Museum">ZSM</collectionCode>
).
</paragraph>
<paragraph id="3E133697FFAC114D1BA1CAC964D16852" blockId="6.[189,1180,175,341]" pageId="6" pageNumber="181">
<emphasis id="0CD8EA85FFAC114D1BA1CAC965DF69F9" bold="true" box="[189,985,232,254]" pageId="6" pageNumber="181">[We have also seen the following specimens that we believe were wrongly labelled:</emphasis>
<materialsCitation id="8EC43CCAFFAC114D18FDCAC964C86852" collectingDate="1985-06-22" collectionCode="MNHB, CER, ZSM" collectorName="W. Weber &amp; F. Wolf &amp; Sammlung Cl. Muller" country="France" county="Kamtschia" location="Bordeaux" municipality="Erlangen" pageId="6" pageNumber="181" specimenCount="1">
<collectingCountry id="46BB7607FFAC114D18FDCAC9622469F9" box="[993,1058,233,254]" name="France" pageId="6" pageNumber="181">France</collectingCountry>
: „
<location id="3B73604CFFAC114D1F29CAC8628869F9" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAC114D1F29CAC8628869F9" box="[1077,1166,232,254]" country="France" county="Kamtschia" municipality="Erlangen" name="Bordeaux" pageId="6" pageNumber="181">Bordeaux</location>
| ex museo
<collectorName id="93595341FFAC114D1A06CB26677E681C" box="[282,376,261,283]" pageId="6" pageNumber="181">W. Weber</collectorName>
“ (1 ,
<collectionCode id="58BDAE52FFAC114D1AA2CB26640A681C" box="[446,524,262,283]" pageId="6" pageNumber="181">MNHB</collectionCode>
). –
<collectingCountry id="46BB7607FFAC114D192CCB256461681C" box="[560,615,261,283]" name="Spain" pageId="6" pageNumber="181">Spain</collectingCountry>
: „Spanien,
<location id="3B73604CFFAC114D19CCCB256512681C" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFAC114D19CCCB256512681C" box="[720,788,261,283]" country="France" county="Kamtschia" municipality="Erlangen" name="Malaga" pageId="6" pageNumber="181">Malaga</location>
“ (1 ,
<collectionCode id="58BDAE52FFAC114D1846CB2665AE681C" box="[858,936,262,283]" pageId="6" pageNumber="181">MNHB</collectionCode>
). –
<collectingCountry id="46BB7607FFAC114D18D0CB256220681D" box="[972,1062,261,282]" name="Germany" pageId="6" pageNumber="181">Germany</collectingCountry>
: „Germania,
<collectingMunicipality id="DE77ACEDFFAC114D1BA1CB026717683F" box="[189,273,290,312]" pageId="6" pageNumber="181">Erlangen</collectingMunicipality>
“ (5 ,
<collectionCode id="58BDAE52FFAC114D1A76CB0367BE683F" box="[362,440,291,312]" pageId="6" pageNumber="181">MNHB</collectionCode>
). –
<collectingCountry id="46BB7607FFAC114D1AC1CB02642B683F" box="[477,557,290,312]" name="Bulgaria" pageId="6" pageNumber="181">Bulgaria</collectingCountry>
: „
<collectingCountry id="46BB7607FFAC114D1923CB02648A683F" box="[575,652,290,312]" name="Bulgaria" pageId="6" pageNumber="181">Bulgaria</collectingCountry>
oc.,
<collectingCounty id="D7724E1BFFAC114D19A0CB026526683F" box="[700,800,290,312]" pageId="6" pageNumber="181">Kamtschia</collectingCounty>
,
<date id="4A121057FFAC114D1837CB02659F683F" box="[811,921,290,312]" pageId="6" pageNumber="181" value="1985-06-22">
<collectingDate id="5A56E9BFFFAC114D1837CB02659F683F" box="[811,921,290,312]" pageId="6" pageNumber="181" value="1985-06-22">22.VI.1985</collectingDate>
</date>
, leg.
<collectorName id="93595341FFAC114D18D0CB036217683F" box="[972,1041,290,312]" pageId="6" pageNumber="181">F. Wolf</collectorName>
“ (1 ,
<collectionCode id="58BDAE52FFAC114D1F44CB01628A683F" box="[1112,1164,289,312]" pageId="6" pageNumber="181">CER</collectionCode>
).] –
<collectingCountry id="46BB7607FFAC114D1BD3CB1F670B6852" box="[207,269,319,341]" name="Russia" pageId="6" pageNumber="181">Russia</collectingCountry>
: „Ross. m. |
<collectorName id="93595341FFAC114D1A9CCB1F644B6852" box="[384,589,319,341]" pageId="6" pageNumber="181">Sammlung Cl. Müller</collectorName>
“ (1 ,
<collectionCode id="58BDAE52FFAC114D198DCB1E64CE6852" box="[657,712,318,341]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="181" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.]
</paragraph>
<caption id="6AD3661FFFAC114D1BF0CC0164C16F77" ID-DOI="http://doi.org/10.5281/zenodo.4779387" ID-Zenodo-Dep="4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="6" pageNumber="181" startId="6.[236,274,1569,1591]" targetBox="[286,1068,393,1546]" targetPageId="6">
<paragraph id="3E133697FFAC114D1BF0CC0164C16F77" blockId="6.[236,1145,1569,1648]" pageId="6" pageNumber="181">
<emphasis id="0CD8EA85FFAC114D1BF0CC01674D6F31" bold="true" box="[236,331,1569,1591]" pageId="6" pageNumber="181">Figs 6-10:</emphasis>
<taxonomicName id="F9AC4D14FFAC114D1A4BCC0164076F30" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" box="[343,513,1569,1591]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="succincta">
<emphasis id="0CD8EA85FFAC114D1A4BCC0164076F30" box="[343,513,1569,1591]" italics="true" pageId="6" pageNumber="181">Exomala succincta</emphasis>
</taxonomicName>
, France, Basses Alpes, St. Andire, . –
<emphasis id="0CD8EA85FFAC114D18BDCC0165B26F31" bold="true" box="[929,948,1569,1590]" pageId="6" pageNumber="181">6:</emphasis>
Aedeagus, dorsal. –
<emphasis id="0CD8EA85FFAC114D1BF0CC1E66F86F54" bold="true" box="[236,254,1598,1619]" pageId="6" pageNumber="181">7:</emphasis>
Aedeagus, ventral. –
<emphasis id="0CD8EA85FFAC114D1ADACC1E67DE6F54" bold="true" box="[454,472,1598,1619]" pageId="6" pageNumber="181">8:</emphasis>
Aedeagus, lateral. –
<emphasis id="0CD8EA85FFAC114D1985CC1E64AD6F54" bold="true" box="[665,683,1598,1619]" pageId="6" pageNumber="181">9:</emphasis>
Last joint and inner claw (outer claw not shown) of right fore tarsus. –
<emphasis id="0CD8EA85FFAC114D1AA8CC7A67D56F77" bold="true" box="[436,467,1626,1648]" pageId="6" pageNumber="181">10:</emphasis>
Pronotum and scutellum.
</paragraph>
</caption>
<caption id="6AD3661FFFAD114C1B86CFB1649A6F30" ID-DOI="http://doi.org/10.5281/zenodo.4779389" ID-Zenodo-Dep="4779389" httpUri="https://zenodo.org/record/4779389/files/figure.png" pageId="7" pageNumber="182" startId="7.[154,192,1425,1447]" targetBox="[242,910,200,1402]" targetPageId="7">
<paragraph id="3E133697FFAD114C1B86CFB1649A6F30" blockId="7.[154,1063,1425,1591]" pageId="7" pageNumber="182">
<emphasis id="0CD8EA85FFAD114C1B86CFB167036CA1" bold="true" box="[154,261,1425,1447]" pageId="7" pageNumber="182">Figs 11-22:</emphasis>
Colour pattern of left elytron. –
<emphasis id="0CD8EA85FFAD114C192ACFB164726CA1" bold="true" box="[566,628,1425,1446]" pageId="7" pageNumber="182">11-16:</emphasis>
<taxonomicName id="F9AC4D14FFAD114C1966CFB1652E6CA0" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[634,808,1425,1447]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFAD114C1966CFB1652E6CA0" box="[634,808,1425,1447]" italics="true" pageId="7" pageNumber="182">Exomala campestris</emphasis>
</taxonomicName>
–
<emphasis id="0CD8EA85FFAD114C1823CFB165786CA1" bold="true" box="[831,894,1425,1446]" pageId="7" pageNumber="182">17-22:</emphasis>
<taxonomicName id="F9AC4D14FFAD114C1898CFB162206CA0" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" box="[900,1062,1425,1447]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="succincta">
<emphasis id="0CD8EA85FFAD114C1898CFB162206CA0" box="[900,1062,1425,1447]" italics="true" pageId="7" pageNumber="182">Exomala succincta</emphasis>
</taxonomicName>
–
<emphasis id="0CD8EA85FFAD114C1BB7CF8E67296CC4" bold="true" box="[171,303,1454,1475]" pageId="7" pageNumber="182">11-13, 17-19:</emphasis>
. –
<emphasis id="0CD8EA85FFAD114C1A6BCF8E67FD6CC4" bold="true" box="[375,507,1454,1475]" pageId="7" pageNumber="182">14-16, 20-22:</emphasis>
. –
<emphasis id="0CD8EA85FFAD114C195FCF8E64C96CC4" bold="true" box="[579,719,1454,1475]" pageId="7" pageNumber="182">11, 14, 17, 20:</emphasis>
Specimens with reduced dark marks. –
<emphasis id="0CD8EA85FFAD114C1BB0CFEB673C6CE7" bold="true" box="[172,314,1483,1504]" pageId="7" pageNumber="182">12, 15, 18, 21:</emphasis>
Specimens with moderately developed dark marks. –
<emphasis id="0CD8EA85FFAD114C182CCFEB65B96CE7" bold="true" box="[816,959,1483,1504]" pageId="7" pageNumber="182">13, 16, 19, 22:</emphasis>
Specimens with well developed dark marks. –
<emphasis id="0CD8EA85FFAD114C1AFBCFC764006CFB" bold="true" box="[487,518,1511,1532]" pageId="7" pageNumber="182">11:</emphasis>
Gallia mer. –
<emphasis id="0CD8EA85FFAD114C198ECFC764B66CFB" bold="true" box="[658,688,1511,1532]" pageId="7" pageNumber="182">12:</emphasis>
France. –
<emphasis id="0CD8EA85FFAD114C180BCFC765306CFB" bold="true" box="[791,822,1511,1532]" pageId="7" pageNumber="182">13:</emphasis>
Gironde: La Teste. –
<emphasis id="0CD8EA85FFAD114C1F1BCFC762206CFB" bold="true" box="[1031,1062,1511,1532]" pageId="7" pageNumber="182">14:</emphasis>
France. –
<emphasis id="0CD8EA85FFAD114C1BEFCC2467146F1E" bold="true" box="[243,274,1540,1561]" pageId="7" pageNumber="182">15:</emphasis>
France. –
<emphasis id="0CD8EA85FFAD114C1A6DCC2467896F1E" bold="true" box="[369,399,1540,1561]" pageId="7" pageNumber="182">16:</emphasis>
Wironde: Soulac-sur-Mer. –
<emphasis id="0CD8EA85FFAD114C1984CC2464B06F1E" bold="true" box="[664,694,1540,1561]" pageId="7" pageNumber="182">17:</emphasis>
Tirol. –
<emphasis id="0CD8EA85FFAD114C1819CC2465226F1E" bold="true" box="[773,804,1540,1561]" pageId="7" pageNumber="182">18:</emphasis>
Tirol mer. –
<emphasis id="0CD8EA85FFAD114C1886CC2465BF6F1E" bold="true" box="[922,953,1540,1561]" pageId="7" pageNumber="182">19:</emphasis>
Tirol. –
<emphasis id="0CD8EA85FFAD114C1F1BCC2462206F1E" bold="true" box="[1031,1062,1540,1561]" pageId="7" pageNumber="182">20:</emphasis>
Emilia, Reno. –
<emphasis id="0CD8EA85FFAD114C1A2DCC0167566F31" bold="true" box="[305,336,1569,1590]" pageId="7" pageNumber="182">21:</emphasis>
Piemonte: Garearel. –
<emphasis id="0CD8EA85FFAD114C193ACC0164426F31" bold="true" box="[550,580,1569,1590]" pageId="7" pageNumber="182">22:</emphasis>
Bolzano.
</paragraph>
</caption>
<paragraph id="3E133697FFA211431BC6CFAE623B6F10" blockId="8.[218,1125,1422,1559]" pageId="8" pageNumber="183">
<emphasis id="0CD8EA85FFA211431BC6CFAE67436CA4" bold="true" box="[218,325,1422,1444]" pageId="8" pageNumber="183">Figs 23-34:</emphasis>
<materialsCitation id="8EC43CCAFFA211431A50CFAE679A6CD9" collectionCode="SMTD" collectorName="Male" country="Spain" location="Ponferrada" pageId="8" pageNumber="183" specimenCount="1">
<collectorName id="93595341FFA211431A50CFAE677D6CA3" box="[332,379,1422,1444]" pageId="8" pageNumber="183">Male</collectorName>
genitalia. –
<emphasis id="0CD8EA85FFA211431AECCFAE64CC6CA4" bold="true" box="[496,714,1422,1443]" pageId="8" pageNumber="183">23, 25, 27, 29, 31, 33:</emphasis>
parameres in dorsal view. –
<emphasis id="0CD8EA85FFA2114318CACFAE671B6CC7" bold="true" pageId="8" pageNumber="183">24, 26, 28, 30, 32, 34:</emphasis>
parameres in ventral view and ventral plate. –
<emphasis id="0CD8EA85FFA2114319D6CF8B650F6CC7" bold="true" box="[714,777,1451,1472]" pageId="8" pageNumber="183">23-26:</emphasis>
<taxonomicName id="F9AC4D14FFA211431812CF8B65BA6CC6" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[782,956,1451,1473]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFA211431812CF8B65BA6CC6" box="[782,956,1451,1473]" italics="true" pageId="8" pageNumber="183">Exomala campestris</emphasis>
</taxonomicName>
. –
<emphasis id="0CD8EA85FFA2114318C5CF8B621A6CC7" bold="true" box="[985,1052,1451,1472]" pageId="8" pageNumber="183">23, 24:</emphasis>
[
<collectingCountry id="46BB7607FFA211431F3BCF8A62596CC7" box="[1063,1119,1450,1472]" name="Spain" pageId="8" pageNumber="183">Spain</collectingCountry>
]
<location id="3B73604CFFA211431BC6CFE867386CD9" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFA211431BC6CFE867386CD9" box="[218,318,1480,1502]" country="Spain" name="Ponferrada" pageId="8" pageNumber="183">Ponferrada</location>
(
<collectionCode id="58BDAE52FFA211431A50CFE867936CD9" box="[332,405,1480,1502]" pageId="8" pageNumber="183">SMTD</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFA211431AA6CFE865B56CD9" box="[442,947,1479,1502]" collectionCode="SMNS" country="France" location="Hautes Pyrenees" pageId="8" pageNumber="183" specimenCount="1">
<emphasis id="0CD8EA85FFA211431AA6CFE867F96CDA" bold="true" box="[442,511,1480,1501]" pageId="8" pageNumber="183">25, 26:</emphasis>
[
<collectingCountry id="46BB7607FFA211431917CFE7644B6CDA" box="[523,589,1479,1501]" name="France" pageId="8" pageNumber="183">France</collectingCountry>
] Cauterets,
<location id="3B73604CFFA2114319A3CFE8655F6CD9" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFA2114319A3CFE8655F6CD9" box="[703,857,1480,1502]" country="France" name="Hautes Pyrenees" pageId="8" pageNumber="183">Hautes Pyrenées</location>
(
<collectionCode id="58BDAE52FFA21143187BCFE865AA6CD9" box="[871,940,1480,1502]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="8" pageNumber="183">SMNS</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFA2114318CDCFE864756CFC" collectionCode="ZMHUB" country="Italy" location="Tirol" pageId="8" pageNumber="183" specimenCount="1">
<emphasis id="0CD8EA85FFA2114318CDCFE862166CDA" bold="true" box="[977,1040,1480,1501]" pageId="8" pageNumber="183">27-30:</emphasis>
<taxonomicName id="F9AC4D14FFA211431F0BCFE8672F6CFD" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="succincta">
<emphasis id="0CD8EA85FFA211431F0BCFE8672F6CFD" italics="true" pageId="8" pageNumber="183">Exomala succincta</emphasis>
</taxonomicName>
. –
<emphasis id="0CD8EA85FFA211431A5BCFC4678D6CFE" bold="true" box="[327,395,1508,1529]" pageId="8" pageNumber="183">27, 28:</emphasis>
[
<collectingCountry id="46BB7607FFA211431A8ACFC467CF6CFD" box="[406,457,1508,1530]" name="Italy" pageId="8" pageNumber="183">Italia</collectingCountry>
]
<location id="3B73604CFFA211431ACFCFC564076CFC" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFA211431ACFCFC564076CFC" box="[467,513,1509,1531]" country="Italy" name="Tirol" pageId="8" pageNumber="183">Tirol</location>
(
<collectionCode id="58BDAE52FFA211431913CFC5646B6CFC" box="[527,621,1509,1531]" pageId="8" pageNumber="183">ZMHUB</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFA211431993CFC465B46CFC" box="[655,946,1508,1531]" collectionCode="SMTD" country="Italy" location="Bozen" pageId="8" pageNumber="183" specimenCount="1">
<emphasis id="0CD8EA85FFA211431993CFC464D46CFE" bold="true" box="[655,722,1508,1529]" pageId="8" pageNumber="183">29, 30:</emphasis>
[
<collectingCountry id="46BB7607FFA2114319C2CFC465176CFD" box="[734,785,1508,1530]" name="Italy" pageId="8" pageNumber="183">Italia</collectingCountry>
]
<location id="3B73604CFFA211431800CFC565536CFD" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFA211431800CFC565536CFD" box="[796,853,1509,1530]" country="Italy" name="Bozen" pageId="8" pageNumber="183">Bozen</location>
(
<collectionCode id="58BDAE52FFA21143187ECFC565AD6CFC" box="[866,939,1509,1531]" pageId="8" pageNumber="183">SMTD</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFA2114318D1CFC4642E6F10" collectionCode="SMTD" country="Albania" location="Valona" pageId="8" pageNumber="183" specimenCount="1" stateProvince="Vlore">
<emphasis id="0CD8EA85FFA2114318D1CFC462176CFE" bold="true" box="[973,1041,1508,1529]" pageId="8" pageNumber="183">31, 32:</emphasis>
<taxonomicName id="F9AC4D14FFA211431F0BCFC5672C6F11" authorityName="PETROVITZ" authorityYear="1968" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="adriatica">
<emphasis id="0CD8EA85FFA211431F0BCFC5672C6F11" italics="true" pageId="8" pageNumber="183">Exomala adriatica</emphasis>
</taxonomicName>
[
<collectingCountry id="46BB7607FFA211431A29CC2167876F10" box="[309,385,1537,1559]" name="Albania" pageId="8" pageNumber="183">Albania</collectingCountry>
]
<collectingRegion id="FC68F875FFA211431A90CC2167CA6F10" box="[396,460,1537,1559]" country="Albania" name="Vlore" pageId="8" pageNumber="183">Valona</collectingRegion>
(
<collectionCode id="58BDAE52FFA211431AC4CC2164276F10" box="[472,545,1537,1559]" pageId="8" pageNumber="183">SMTD</collectionCode>
)
</materialsCitation>
. –
<materialsCitation id="8EC43CCAFFA21143195FCC21623F6F10" box="[579,1081,1537,1559]" collectionCode="CER" country="Turkey" location="Buglan Gecidi" pageId="8" pageNumber="183" specimenCount="1">
<emphasis id="0CD8EA85FFA21143195FCC2164806F11" bold="true" box="[579,646,1537,1558]" pageId="8" pageNumber="183">33, 34:</emphasis>
<taxonomicName id="F9AC4D14FFA211431990CC2165176F11" authorityName="PETROVITZ" authorityYear="1968" box="[652,785,1537,1558]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="bileki">
<emphasis id="0CD8EA85FFA211431990CC2165176F11" box="[652,785,1537,1558]" italics="true" pageId="8" pageNumber="183">Exomala bileki</emphasis>
</taxonomicName>
[
<collectingCountry id="46BB7607FFA211431801CC2165656F10" box="[797,867,1537,1559]" name="Turkey" pageId="8" pageNumber="183">Turkey</collectingCountry>
]
<location id="3B73604CFFA211431871CC2165F46F10" LSID="urn:lsid:plazi:treatment:B6058781FFAF11401BA9C99F674A6A8F:3B73604CFFA211431871CC2165F46F10" box="[877,1010,1537,1559]" country="Turkey" name="Buglan Gecidi" pageId="8" pageNumber="183">Buglan Gecidi</location>
(
<collectionCode id="58BDAE52FFA2114318E3CC2162346F10" box="[1023,1074,1537,1559]" pageId="8" pageNumber="183">CER</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="6AD3661FFFA311421B8CCE0167386D53" ID-DOI="http://doi.org/10.5281/zenodo.4779393" ID-Zenodo-Dep="4779393" httpUri="https://zenodo.org/record/4779393/files/figure.png" pageId="9" pageNumber="184" startId="9.[144,176,1057,1079]" targetBox="[154,1024,184,1044]" targetPageId="9">
<paragraph id="3E133697FFA311421B8CCE0167386D53" blockId="9.[144,1050,1057,1108]" pageId="9" pageNumber="184">
<emphasis id="0CD8EA85FFA311421B8CCE0166DE6D31" bold="true" box="[144,216,1057,1079]" pageId="9" pageNumber="184">Fig. 35:</emphasis>
Collecting localities of the examined material of
<taxonomicName id="F9AC4D14FFA3114219BCCE0165486D31" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[672,846,1057,1078]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFA3114219BCCE0165486D31" box="[672,846,1057,1078]" italics="true" pageId="9" pageNumber="184">Exomala campestris</emphasis>
</taxonomicName>
(circles) und
<taxonomicName id="F9AC4D14FFA3114218D0CE0166D96D54" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="184" phylum="Arthropoda" rank="species" species="succincta">
<emphasis id="0CD8EA85FFA3114218D0CE0166D96D54" italics="true" pageId="9" pageNumber="184">Exomala succincta</emphasis>
</taxonomicName>
(squares).
</paragraph>
</caption>
<subSubSection id="76B6651CFFA311411B76CE4F66F36AC5" lastPageId="10" lastPageNumber="185" pageId="9" pageNumber="184" type="description">
<paragraph id="3E133697FFA311421B76CE4F66EB6D80" blockId="9.[106,237,1135,1159]" box="[106,237,1135,1159]" pageId="9" pageNumber="184">
<emphasis id="0CD8EA85FFA311421B76CE4F66EB6D80" bold="true" box="[106,237,1135,1159]" pageId="9" pageNumber="184">Description:</emphasis>
</paragraph>
<paragraph id="3E133697FFA311421B76CEB567C16C0E" blockId="9.[106,1096,1173,1289]" pageId="9" pageNumber="184">
Head, pronotum and scutellum black, shining, often with weak metallic lustre. Elytra yellowish brown, moderately shining; dark marks almost always well developed (
<figureCitation id="A6972A12FFA311421832CE94659C6DCB" box="[814,922,1204,1228]" captionStart="Figs 11-22" captionStartId="7.[154,192,1425,1447]" captionTargetBox="[242,910,200,1402]" captionTargetId="figure-124@7.[155,1062,174,1425]" captionTargetPageId="7" captionText="Figs 11-22: Colour pattern of left elytron. – 11-16: Exomala campestris – 17-22: Exomala succincta – 11-13, 17-19: . – 14-16, 20-22: . – 11, 14, 17, 20: Specimens with reduced dark marks. – 12, 15, 18, 21: Specimens with moderately developed dark marks. – 13, 16, 19, 22: Specimens with well developed dark marks. – 11: Gallia mer. – 12: France. – 13: Gironde: La Teste. – 14: France. – 15: France. – 16:Wironde: Soulac-sur-Mer.– 17: Tirol. – 18:Tirol mer. – 19: Tirol. – 20: Emilia, Reno. – 21: Piemonte: Garearel. – 22: Bolzano." figureDoi="http://doi.org/10.5281/zenodo.4779389" httpUri="https://zenodo.org/record/4779389/files/figure.png" pageId="9" pageNumber="184">figs 17-22</figureCitation>
); usually more or less broadly blackened along the periphery, on the umbone and on the area around the scutellum, and with a transversal black band.
</paragraph>
<paragraph id="3E133697FFA311421B76CF3765246C6A" blockId="9.[106,1096,1303,1389]" pageId="9" pageNumber="184">Clypeus broadly rectangular; sides parallel; fore edge reflexed and weakly convex; anterior angles rounded; punctation transversely rugose; clypeus sparsely, forehead and vertex densely covered with erect, light-coloured setae; with very dense, rugose punctation.</paragraph>
<paragraph id="3E133697FFA311421B76CF5B657C6C94" blockId="9.[106,890,1403,1427]" box="[106,890,1403,1427]" pageId="9" pageNumber="184">Antennal club rather short, outer face of last antennomere distinctly convex.</paragraph>
<paragraph id="3E133697FFA311421B76CF82649C6CF1" blockId="9.[106,1096,1441,1527]" pageId="9" pageNumber="184">
Pronotum laterally with an obtuse, blunt angle (
<figureCitation id="A6972A12FFA311421976CF8164B66CBE" box="[618,688,1441,1465]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="9" pageNumber="184">fig. 10</figureCitation>
); basal margin disappearing before the postero-lateral angles; base straight in front of scutellum (
<figureCitation id="A6972A12FFA3114219B4CFE064E86CDF" box="[680,750,1472,1496]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="9" pageNumber="184">fig. 10</figureCitation>
); with dense transverse, somewhat rugose punctation and long, erect light-coloured setae.
</paragraph>
<paragraph id="3E133697FFA311421B76CC25650A6F1B" blockId="9.[106,780,1541,1565]" box="[106,780,1541,1565]" pageId="9" pageNumber="184">
Scutellum (
<figureCitation id="A6972A12FFA311421BFCCC2567226F1A" box="[224,292,1541,1565]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="9" pageNumber="184">fig. 10</figureCitation>
) broadly semicircular; densely punctate, setose.
</paragraph>
<paragraph id="3E133697FFA311421B76CC0B657C6F65" blockId="9.[106,1096,1579,1634]" pageId="9" pageNumber="184">
Elytra (
<figureCitation id="A6972A12FFA311421BA8CC0B67186F44" box="[180,286,1579,1603]" captionStart="Figs 11-22" captionStartId="7.[154,192,1425,1447]" captionTargetBox="[242,910,200,1402]" captionTargetId="figure-124@7.[155,1062,174,1425]" captionTargetPageId="7" captionText="Figs 11-22: Colour pattern of left elytron. – 11-16: Exomala campestris – 17-22: Exomala succincta – 11-13, 17-19: . – 14-16, 20-22: . – 11, 14, 17, 20: Specimens with reduced dark marks. – 12, 15, 18, 21: Specimens with moderately developed dark marks. – 13, 16, 19, 22: Specimens with well developed dark marks. – 11: Gallia mer. – 12: France. – 13: Gironde: La Teste. – 14: France. – 15: France. – 16:Wironde: Soulac-sur-Mer.– 17: Tirol. – 18:Tirol mer. – 19: Tirol. – 20: Emilia, Reno. – 21: Piemonte: Garearel. – 22: Bolzano." figureDoi="http://doi.org/10.5281/zenodo.4779389" httpUri="https://zenodo.org/record/4779389/files/figure.png" pageId="9" pageNumber="184">figs 17-22</figureCitation>
) broadest approximately in the middle; odd intervals feebly convex and weakly setose; all intervals with more or less dinstinct transverse, rugose punctation.
</paragraph>
<caption id="6AD3661FFFA011411BDDC92662036A1C" ID-DOI="http://doi.org/10.5281/zenodo.4779395" ID-Zenodo-Dep="4779395" box="[193,1029,774,796]" httpUri="https://zenodo.org/record/4779395/files/figure.png" pageId="10" pageNumber="185" startId="10.[193,225,774,796]" targetBox="[203,1167,196,758]" targetPageId="10">
<paragraph id="3E133697FFA011411BDDC92662036A1C" blockId="10.[193,1029,774,796]" box="[193,1029,774,796]" pageId="10" pageNumber="185">
<emphasis id="0CD8EA85FFA011411BDDC926670F6A1C" bold="true" box="[193,265,774,796]" pageId="10" pageNumber="185">Fig. 36:</emphasis>
Boxplots of the body length of males and females of
<taxonomicName id="F9AC4D14FFA0114119E9C927656E6A1B" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[757,872,775,796]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFA0114119E9C927656E6A1B" box="[757,872,775,796]" italics="true" pageId="10" pageNumber="185">E. campestris</emphasis>
</taxonomicName>
and
<taxonomicName id="F9AC4D14FFA01141188AC92765F96A1B" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" box="[918,1023,775,796]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="succincta">
<emphasis id="0CD8EA85FFA01141188AC92765F96A1B" box="[918,1023,775,796]" italics="true" pageId="10" pageNumber="185">E. succincta</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="3E133697FFA011411BA1C91F62666A50" blockId="10.[189,1120,831,855]" box="[189,1120,831,855]" pageId="10" pageNumber="185">Pygidium densely transversely rugose; with moderately dense, long, light-coloured pilosity.</paragraph>
<paragraph id="3E133697FFA011411BA1C94565336A7B" blockId="10.[189,821,869,893]" box="[189,821,869,893]" pageId="10" pageNumber="185">Ventral face and femora with rather long light-coloured setae.</paragraph>
<paragraph id="3E133697FFA011411BA1C9AC66F36AC5" blockId="10.[189,1179,907,962]" pageId="10" pageNumber="185">
Inner claw of fore tarsi in males (
<figureCitation id="A6972A12FFA011411907C9AB64546AA4" box="[539,594,907,931]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="10" pageNumber="185">fig. 9</figureCitation>
) relatively long, approximately half as long as last tarsal joint.
</paragraph>
</subSubSection>
<subSubSection id="76B6651CFFA011411BA1C9FF62296CA3" pageId="10" pageNumber="185" type="distribution">
<paragraph id="3E133697FFA011411BA1C9FF674E6AF0" blockId="10.[189,328,991,1015]" box="[189,328,991,1015]" pageId="10" pageNumber="185">
<emphasis id="0CD8EA85FFA011411BA1C9FF674E6AF0" bold="true" box="[189,328,991,1015]" pageId="10" pageNumber="185">Distribution:</emphasis>
</paragraph>
<paragraph id="3E133697FFA011411BA1CE26628A6D3B" blockId="10.[189,1180,1029,1084]" pageId="10" pageNumber="185">
In fig. 23 the locality data of the examined material are mapped. According to our data and recent literature, the distribution patterns of
<taxonomicName id="F9AC4D14FFA011411958CE0564C26D3B" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[580,708,1061,1084]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFA011411958CE0564C26D3B" box="[580,708,1061,1084]" italics="true" pageId="10" pageNumber="185">E. campestris</emphasis>
</taxonomicName>
and
<taxonomicName id="F9AC4D14FFA0114119EBCE05656D6D3B" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" box="[759,875,1061,1084]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="succincta">
<emphasis id="0CD8EA85FFA0114119EBCE05656D6D3B" box="[759,875,1061,1084]" italics="true" pageId="10" pageNumber="185">E. succincta</emphasis>
</taxonomicName>
can be described as follows.
</paragraph>
<paragraph id="3E133697FFA011411BA1CE6A67816DDA" blockId="10.[189,1179,1098,1245]" pageId="10" pageNumber="185">
The distribution area of
<taxonomicName id="F9AC4D14FFA011411AA4CE6A647F6D65" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[440,633,1098,1122]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFA011411AA4CE6A647F6D65" box="[440,633,1098,1122]" italics="true" pageId="10" pageNumber="185">Exomala campestris</emphasis>
</taxonomicName>
lies roughly in the triangle between the Pyrenees, the Cantabric mountains and the Iberian Central sierras (Sistema Central). In the south the species reaches
<collectingRegion id="FC68F875FFA011411A12CEA8675A6DA7" box="[270,348,1160,1184]" country="Spain" name="Madrid" pageId="10" pageNumber="185">Madrid</collectingRegion>
and Guadalajara (
<bibRefCitation id="5A3D4B66FFA011411907CEA864F66DA7" author="BAGUENA, L. C." box="[539,752,1160,1187]" pageId="10" pageNumber="185" refId="ref5705" refString="BAGUENA, L. C. 1967: Scarabaeoidea de la fauna Ibero-Balear y Pirenaica. - Instituto Espanol de Entomologia. Madrid: 576 pp." type="book" year="1967">BÁGUENA 1967: 435</bibRefCitation>
, sub:
<taxonomicName id="F9AC4D14FFA011411830CEA862006D98" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[812,1030,1160,1184]" class="Insecta" family="Rutelidae" genus="Blitopertha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFA011411830CEA862006D98" box="[812,1030,1160,1184]" italics="true" pageId="10" pageNumber="185">Blitopertha campestris</emphasis>
</taxonomicName>
). In the north its distribution area includes the French Pyrenees and the lowlands of Landes and Médoc up to the Atlantic Ocean.
</paragraph>
<paragraph id="3E133697FFA011411BA1CECC67E46C46" blockId="10.[189,1179,1259,1345]" pageId="10" pageNumber="185">
In general terms
<taxonomicName id="F9AC4D14FFA011411A76CECC67DB6C04" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" box="[362,477,1260,1283]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="succincta">
<emphasis id="0CD8EA85FFA011411A76CECC67DB6C04" box="[362,477,1260,1283]" italics="true" pageId="10" pageNumber="185">E. succincta</emphasis>
</taxonomicName>
is a species of the southern Alp valleys, distributed from Lyon in the west to
<collectingCountry id="46BB7607FFA011411A17CF2A67666C25" box="[267,352,1290,1314]" name="Slovenia" pageId="10" pageNumber="185">Slovenia</collectingCountry>
in the east. In
<collectingCountry id="46BB7607FFA011411AEACF2A64226C25" box="[502,548,1290,1314]" name="Italy" pageId="10" pageNumber="185">Italy</collectingCountry>
its distribution area does also include the northern Apennines (
<collectingRegion id="FC68F875FFA011411BD8CF0967716C46" box="[196,375,1321,1345]" country="Italy" name="Emilia-Romagna" pageId="10" pageNumber="185">Emilia-Romagna</collectingRegion>
,
<collectingRegion id="FC68F875FFA011411A9CCF0967D16C46" box="[384,471,1321,1345]" country="Italy" name="Toscana" pageId="10" pageNumber="185">Toscana</collectingRegion>
).
</paragraph>
<paragraph id="3E133697FFA011411BA1CF6F62296CA3" blockId="10.[189,1179,1359,1444]" pageId="10" pageNumber="185">
Therefore, according to the available data,
<taxonomicName id="F9AC4D14FFA01141198DCF7065106C60" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[657,790,1360,1383]" class="Insecta" family="Rutelidae" genus="Blitopertha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFA01141198DCF7065106C60" box="[657,790,1360,1383]" italics="true" pageId="10" pageNumber="185">E. campestris</emphasis>
</taxonomicName>
and
<taxonomicName id="F9AC4D14FFA01141184FCF7065CD6C60" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" box="[851,971,1360,1383]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="succincta">
<emphasis id="0CD8EA85FFA01141184FCF7065CD6C60" box="[851,971,1360,1383]" italics="true" pageId="10" pageNumber="185">E. succincta</emphasis>
</taxonomicName>
show an allopatric distribution, their respective geographic ranges are neither overlapping nor connected. They are separated by the Rhône valley, the Massif Central and the Cévennes in central
<collectingCountry id="46BB7607FFA0114118FFCFAD622D6CA3" box="[995,1067,1421,1444]" name="France" pageId="10" pageNumber="185">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="76B6651CFFA011401BA1CFE1674A6A8F" lastPageId="11" lastPageNumber="186" pageId="10" pageNumber="185" type="diagnosis">
<paragraph id="3E133697FFA011411BA1CFE167A06CDE" blockId="10.[189,422,1473,1497]" box="[189,422,1473,1497]" pageId="10" pageNumber="185">
<emphasis id="0CD8EA85FFA011411BA1CFE167A06CDE" bold="true" box="[189,422,1473,1497]" pageId="10" pageNumber="185">Differential diagnosis:</emphasis>
</paragraph>
<paragraph id="3E133697FFA011401BA1CFC862286855" blockId="10.[189,1179,1512,1566]" lastBlockId="11.[116,1086,222,904]" lastPageId="11" lastPageNumber="186" pageId="10" pageNumber="185">
The differences on which the very similar species
<taxonomicName id="F9AC4D14FFA0114119DCCFC865476CF8" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[704,833,1512,1535]" class="Insecta" family="Rutelidae" genus="Blitopertha" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFA0114119DCCFC865476CF8" box="[704,833,1512,1535]" italics="true" pageId="10" pageNumber="185">E. campestris</emphasis>
</taxonomicName>
and
<taxonomicName id="F9AC4D14FFA01141186ACFC865EC6CF8" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" box="[886,1002,1512,1535]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="185" phylum="Arthropoda" rank="species" species="succincta">
<emphasis id="0CD8EA85FFA01141186ACFC865EC6CF8" box="[886,1002,1512,1535]" italics="true" pageId="10" pageNumber="185">E. succincta</emphasis>
</taxonomicName>
can be separated are given in the table below.
<emphasis id="0CD8EA85FFA111401B90CAFE658969F4" bold="true" box="[140,911,222,243]" pageId="11" pageNumber="186">
character
<emphasis id="0CD8EA85FFA111401A66CAFE658969F4" bold="true" box="[378,911,222,243]" italics="true" pageId="11" pageNumber="186">
<taxonomicName id="F9AC4D14FFA111401A66CAFE67F569F4" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[378,499,222,243]" class="Insecta" family="Rutelidae" genus="Blitopertha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="campestris">E. campestris</taxonomicName>
<taxonomicName id="F9AC4D14FFA11140183ECAFE658969F4" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" box="[802,911,222,243]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="succincta">E. succincta</taxonomicName>
</emphasis>
</emphasis>
inner claw of shorter than half the length of last as long as half the length of last tarsal joint fore tarsi () tarsal joint (
<figureCitation id="A6972A12FFA111401A60CB3B67A86836" box="[380,430,283,305]" captionStart="Figs 1-5" captionStartId="4.[219,257,1515,1537]" captionTargetBox="[311,1068,331,1488]" captionTargetId="figure-115@4.[219,1126,270,1510]" captionTargetPageId="4" captionText="Figs 1-5: Exomala campestris, France, Gironde, La Teste, . – 1: Aedeagus, dorsal. – 2: Aedeagus, ventral. – 3: Aedeagus, lateral. – 4: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 5: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779383" httpUri="https://zenodo.org/record/4779383/files/figure.png" pageId="11" pageNumber="186">fig. 4</figureCitation>
) (
<figureCitation id="A6972A12FFA111401966CB3B64AD6836" box="[634,683,283,305]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="11" pageNumber="186">fig. 9</figureCitation>
) body length 7.0-9.5 mm (mean 8.48 mm) (
<figureCitation id="A6972A12FFA111401935CB1C64C56856" box="[553,707,316,338]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="11" pageNumber="186">fig. 6, 8.2-10.4</figureCitation>
mm (mean 9.13 mm) (
<figureCitation id="A6972A12FFA111401883CB1C65D86855" box="[927,990,316,338]" captionStart="Fig" captionStartId="10.[193,225,774,796]" captionTargetBox="[203,1167,196,758]" captionTargetId="figure-268@10.[193,1176,176,768]" captionTargetPageId="10" captionText="Fig. 36: Boxplots of the body length of males and females of E. campestris and E. succincta." figureDoi="http://doi.org/10.5281/zenodo.4779395" httpUri="https://zenodo.org/record/4779395/files/figure.png" pageId="11" pageNumber="186">fig. 36</figureCitation>
,
<tableCitation id="732E032CFFA1114018F4CB1C622E6855" box="[1000,1064,316,338]" captionStart="Tab" captionStartId="11.[106,144,950,971]" captionText="Tab. 2: Body length of males and females of E. campestris and E. succincta (minimum, maximum, mean, standard deviation, median). The differences in mean between E. campestris and E. succincta (males, females and overall) are highly significant (T-test, p&lt;0,01)." pageId="11" pageNumber="186">table 2</tableCitation>
)
</paragraph>
<caption id="6AD3661FFFA111401B76CA8F650B69C2" box="[106,781,175,197]" pageId="11" pageNumber="186" startId="11.[106,144,175,196]">
<paragraph id="3E133697FFA111401B76CA8F650B69C2" blockId="11.[106,781,175,197]" box="[106,781,175,197]" pageId="11" pageNumber="186">
<emphasis id="0CD8EA85FFA111401B76CA8F66AB69C3" bold="true" box="[106,173,175,196]" pageId="11" pageNumber="186">Tab. 1:</emphasis>
Morphological differences between
<taxonomicName id="F9AC4D14FFA111401AE7CA90646969C2" baseAuthorityName="LATREILLE" baseAuthorityYear="1804" box="[507,623,176,197]" class="Insecta" family="Rutelidae" genus="Blitopertha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFA111401AE7CA90646969C2" box="[507,623,176,197]" italics="true" pageId="11" pageNumber="186">E. campestris</emphasis>
</taxonomicName>
and
<taxonomicName id="F9AC4D14FFA111401981CA90650F69C2" baseAuthorityName="CASTELNAU" baseAuthorityYear="1840" box="[669,777,176,197]" class="Insecta" family="Rutelidae" genus="Exomala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="succincta">E. succincta</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="3E133697FFA111401A11CB7865FF68CB" blockId="11.[116,1086,222,904]" pageId="11" pageNumber="186">
<tableCitation id="732E032CFFA111401A11CB78674B6868" box="[269,333,344,367]" captionStart="Tab" captionStartId="11.[106,144,950,971]" captionText="Tab. 2: Body length of males and females of E. campestris and E. succincta (minimum, maximum, mean, standard deviation, median). The differences in mean between E. campestris and E. succincta (males, females and overall) are highly significant (T-test, p&lt;0,01)." pageId="11" pageNumber="186">table 2</tableCitation>
) antennal club elongate, outer face of last antenno- less elongate, outer face of last antennomere () mere weakly convex distinctly convex pronotum sides evenly rounded (
<figureCitation id="A6972A12FFA111401AC6CB96640A68CB" box="[474,524,438,460]" captionStart="Figs 1-5" captionStartId="4.[219,257,1515,1537]" captionTargetBox="[311,1068,331,1488]" captionTargetId="figure-115@4.[219,1126,270,1510]" captionTargetPageId="4" captionText="Figs 1-5: Exomala campestris, France, Gironde, La Teste, . – 1: Aedeagus, dorsal. – 2: Aedeagus, ventral. – 3: Aedeagus, lateral. – 4: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 5: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779383" httpUri="https://zenodo.org/record/4779383/files/figure.png" pageId="11" pageNumber="186">fig. 5</figureCitation>
); sides with an obtuse, blunt angle (
<figureCitation id="A6972A12FFA1114018B2CB9665EB68CB" box="[942,1005,438,460]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="11" pageNumber="186">fig. 10</figureCitation>
);
</paragraph>
<paragraph id="3E133697FFA111401A11CBF3623868EE" blockId="11.[116,1086,222,904]" box="[269,1086,467,489]" pageId="11" pageNumber="186">basal margin reaching postero-lateral basal margin disappearing before the postero-</paragraph>
<paragraph id="3E133697FFA111401A11CBD0623B6B01" blockId="11.[116,1086,222,904]" box="[269,1085,496,518]" pageId="11" pageNumber="186">angles, sometimes very faint there; lateral angles; base infront of scutellum straight</paragraph>
<paragraph id="3E133697FFA111401A11C82D64B86B24" blockId="11.[116,1086,222,904]" box="[269,702,525,547]" pageId="11" pageNumber="186">
base in front of scutellum evenly (
<figureCitation id="A6972A12FFA111401966C82D64B16B24" box="[634,695,525,547]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="11" pageNumber="186">fig. 10</figureCitation>
)
</paragraph>
<paragraph id="3E133697FFA111401A11C80A656D6B67" blockId="11.[116,1086,222,904]" pageId="11" pageNumber="186">
rounded, convex (
<figureCitation id="A6972A12FFA111401AA8C80A67E36B38" box="[436,485,553,576]" captionStart="Figs 1-5" captionStartId="4.[219,257,1515,1537]" captionTargetBox="[311,1068,331,1488]" captionTargetId="figure-115@4.[219,1126,270,1510]" captionTargetPageId="4" captionText="Figs 1-5: Exomala campestris, France, Gironde, La Teste, . – 1: Aedeagus, dorsal. – 2: Aedeagus, ventral. – 3: Aedeagus, lateral. – 4: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 5: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779383" httpUri="https://zenodo.org/record/4779383/files/figure.png" pageId="11" pageNumber="186">fig. 5</figureCitation>
) scutellum mostly narrow and elongate, rarely wide, semicircular (
<figureCitation id="A6972A12FFA11140183BC86A65626B67" box="[807,868,586,608]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="11" pageNumber="186">fig. 10</figureCitation>
)
</paragraph>
<paragraph id="3E133697FFA111401A11C847623B6B9A" blockId="11.[116,1086,222,904]" pageId="11" pageNumber="186">
semicircular (
<figureCitation id="A6972A12FFA111401A95C84767BC6B7A" box="[393,442,615,637]" captionStart="Figs 1-5" captionStartId="4.[219,257,1515,1537]" captionTargetBox="[311,1068,331,1488]" captionTargetId="figure-115@4.[219,1126,270,1510]" captionTargetPageId="4" captionText="Figs 1-5: Exomala campestris, France, Gironde, La Teste, . – 1: Aedeagus, dorsal. – 2: Aedeagus, ventral. – 3: Aedeagus, lateral. – 4: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 5: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779383" httpUri="https://zenodo.org/record/4779383/files/figure.png" pageId="11" pageNumber="186">fig. 5</figureCitation>
) elytra dark marks usually reduced, dark marks, including the transversal band, almost
</paragraph>
<paragraph id="3E133697FFA111401A11C884623B6BBE" blockId="11.[116,1086,222,904]" box="[269,1085,676,698]" pageId="11" pageNumber="186">
transversal band rarely distinct always well developed (
<figureCitation id="A6972A12FFA111401845C88465BE6BBD" box="[857,952,676,698]" captionStart="Figs 11-22" captionStartId="7.[154,192,1425,1447]" captionTargetBox="[242,910,200,1402]" captionTargetId="figure-124@7.[155,1062,174,1425]" captionTargetPageId="7" captionText="Figs 11-22: Colour pattern of left elytron. – 11-16: Exomala campestris – 17-22: Exomala succincta – 11-13, 17-19: . – 14-16, 20-22: . – 11, 14, 17, 20: Specimens with reduced dark marks. – 12, 15, 18, 21: Specimens with moderately developed dark marks. – 13, 16, 19, 22: Specimens with well developed dark marks. – 11: Gallia mer. – 12: France. – 13: Gironde: La Teste. – 14: France. – 15: France. – 16:Wironde: Soulac-sur-Mer.– 17: Tirol. – 18:Tirol mer. – 19: Tirol. – 20: Emilia, Reno. – 21: Piemonte: Garearel. – 22: Bolzano." figureDoi="http://doi.org/10.5281/zenodo.4779389" httpUri="https://zenodo.org/record/4779389/files/figure.png" pageId="11" pageNumber="186">figs 17-22</figureCitation>
) [cf. ROBERT
</paragraph>
<paragraph id="3E133697FFA111401A11C8E162346BF0" blockId="11.[116,1086,222,904]" pageId="11" pageNumber="186">
(
<figureCitation id="A6972A12FFA111401A0FC8E167746BD0" box="[275,370,705,727]" captionStart="Figs 11-22" captionStartId="7.[154,192,1425,1447]" captionTargetBox="[242,910,200,1402]" captionTargetId="figure-124@7.[155,1062,174,1425]" captionTargetPageId="7" captionText="Figs 11-22: Colour pattern of left elytron. – 11-16: Exomala campestris – 17-22: Exomala succincta – 11-13, 17-19: . – 14-16, 20-22: . – 11, 14, 17, 20: Specimens with reduced dark marks. – 12, 15, 18, 21: Specimens with moderately developed dark marks. – 13, 16, 19, 22: Specimens with well developed dark marks. – 11: Gallia mer. – 12: France. – 13: Gironde: La Teste. – 14: France. – 15: France. – 16:Wironde: Soulac-sur-Mer.– 17: Tirol. – 18:Tirol mer. – 19: Tirol. – 20: Emilia, Reno. – 21: Piemonte: Garearel. – 22: Bolzano." figureDoi="http://doi.org/10.5281/zenodo.4779389" httpUri="https://zenodo.org/record/4779389/files/figure.png" pageId="11" pageNumber="186">figs 11-16</figureCitation>
) (1992):
<figureCitation id="A6972A12FFA1114019A3C8E164F66BD1" box="[703,752,705,727]" captionStart="Figs 1-5" captionStartId="4.[219,257,1515,1537]" captionTargetBox="[311,1068,331,1488]" captionTargetId="figure-115@4.[219,1126,270,1510]" captionTargetPageId="4" captionText="Figs 1-5: Exomala campestris, France, Gironde, La Teste, . – 1: Aedeagus, dorsal. – 2: Aedeagus, ventral. – 3: Aedeagus, lateral. – 4: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 5: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779383" httpUri="https://zenodo.org/record/4779383/files/figure.png" pageId="11" pageNumber="186">fig. 2</figureCitation>
, sub:
<taxonomicName id="F9AC4D14FFA11140183BC8E162276BD1" authority="LATR." authorityName="LATR." box="[807,1057,704,727]" class="Insecta" family="Rutelidae" genus="Blitopertha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="186" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="0CD8EA85FFA11140183BC8E165EC6BD0" box="[807,1002,705,727]" italics="true" pageId="11" pageNumber="186">Blitopertha campestris</emphasis>
LATR.
</taxonomicName>
] male genitalia sides of parameres straight in dorsal sides of parameres sinuate in dorsal view (
<figureCitation id="A6972A12FFA1114018E8C8C162206BF0" box="[1012,1062,737,759]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="11" pageNumber="186">fig. 6</figureCitation>
);
</paragraph>
<paragraph id="3E133697FFA111401A11C8DE62386A13" blockId="11.[116,1086,222,904]" box="[269,1086,766,788]" pageId="11" pageNumber="186">
view (
<figureCitation id="A6972A12FFA111401A58C8DE67726A13" box="[324,372,766,788]" captionStart="Figs 1-5" captionStartId="4.[219,257,1515,1537]" captionTargetBox="[311,1068,331,1488]" captionTargetId="figure-115@4.[219,1126,270,1510]" captionTargetPageId="4" captionText="Figs 1-5: Exomala campestris, France, Gironde, La Teste, . – 1: Aedeagus, dorsal. – 2: Aedeagus, ventral. – 3: Aedeagus, lateral. – 4: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 5: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779383" httpUri="https://zenodo.org/record/4779383/files/figure.png" pageId="11" pageNumber="186">fig. 1</figureCitation>
); each paramere broadened before the apex, in
</paragraph>
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each paramere evenly narrow before ventral view (
<figureCitation id="A6972A12FFA1114019F3C93B65276A36" box="[751,801,795,817]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="11" pageNumber="186">fig. 7</figureCitation>
);
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the apex, in ventral view (
<figureCitation id="A6972A12FFA111401AE7C91864286A49" box="[507,558,824,846]" captionStart="Figs 1-5" captionStartId="4.[219,257,1515,1537]" captionTargetBox="[311,1068,331,1488]" captionTargetId="figure-115@4.[219,1126,270,1510]" captionTargetPageId="4" captionText="Figs 1-5: Exomala campestris, France, Gironde, La Teste, . – 1: Aedeagus, dorsal. – 2: Aedeagus, ventral. – 3: Aedeagus, lateral. – 4: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 5: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779383" httpUri="https://zenodo.org/record/4779383/files/figure.png" pageId="11" pageNumber="186">fig. 2</figureCitation>
); ventral plate trapezoidal, very wide basally (
<figureCitation id="A6972A12FFA111401F19C91862306A4A" box="[1029,1078,824,846]" captionStart="Figs 6-10" captionStartId="6.[236,274,1569,1591]" captionTargetBox="[286,1068,393,1546]" captionTargetId="figure-147@6.[236,1143,362,1566]" captionTargetPageId="6" captionText="Figs 6-10: Exomala succincta, France, Basses Alpes, St. Andire, . – 6: Aedeagus, dorsal. – 7: Aedeagus, ventral. – 8: Aedeagus, lateral. – 9: Last joint and inner claw (outer claw not shown) of right fore tarsus. – 10: Pronotum and scutellum." figureDoi="http://doi.org/10.5281/zenodo.4779387" httpUri="https://zenodo.org/record/4779387/files/figure.png" pageId="11" pageNumber="186">fig. 7</figureCitation>
)
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<paragraph id="3E133697FFA111401A11C97564666A6C" blockId="11.[116,1086,222,904]" box="[269,608,853,875]" pageId="11" pageNumber="186">ventral plate narrow and elongate</paragraph>
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(
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)
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species complex
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This species complex is the largest within the
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ala
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The forecoxae are usually completely or mostly unsculptured, smooth, and shining, although sculpture is sometimes present. What sculpture is present is mostly superficial and generally does not consist of strongly longitudinally arranged rugae. Instead, it is reticulate-punctate with few superimposed rugulae or traces of rugulae. In a few species (e.g.
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the forecoxae are partly rugulose, but the sculpture is comparatively weak, and never covers the whole coxa as in the
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species complex. The sculpture may appear longitudinal at first glance, but this is due to sections of linearly arranged reticulate-punctate sculpture, not true rugae/rugulae (e.g.
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The first gastral tergite is always completely devoid of any sculpture, and fairly smooth and shining.
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<mods:date id="F93667D126E80985087EDA8B327E899C">2021</mods:date>
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<mods:number id="0A4737CA4FF101B1F34E8E9FC3D3E57F">9</mods:number>
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<mods:classification id="6E9BE3CCAB55FBA79ED8E0EB1E0E4676">journal article</mods:classification>
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<paragraph id="868DF261C60F1CB8801D7D95E9B3CA89" pageId="0" pageNumber="62736">
<taxonomicName id="2C0F38AA894C6FB9CBEACBB57774CF2A" ID-CoL="3Y88R" LSID="B605950B-1D0C-5B6C-BA28-6F112DE17601" authority="Fauchald, 1970" authorityName="Fauchald" authorityYear="1970" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Marphysa angelensis" order="Eunicida" pageId="0" pageNumber="62736" phylum="Annelida" rank="species" species="angelensis">Marphysa angelensis Fauchald, 1970</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="90A37F5EFEFE6A4D4BF62736C32F81F8" pageId="0" pageNumber="62736" type="materials_examined">
<paragraph id="44E21C7A4080046ACF610A19A51F4383" pageId="0" pageNumber="62736">Materials</paragraph>
<paragraph id="B0ABD833EB2C0354EABEF0A8D54CF46C" pageId="0" pageNumber="62736">
<materialsCitation id="2F5B863AC06CC68095C06A5BBE37DC92" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" pageId="0" pageNumber="62736" specimenCount="1" typeStatus="Other material">
<emphasis id="8307199329C14B73D9BE852F790642C4" bold="true" pageId="0" pageNumber="62736">Type status:</emphasis>
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.
<emphasis id="D646BCADB82F3D521AE764927843395E" bold="true" pageId="0" pageNumber="62736">Occurrence:</emphasis>
catalogNumber: CZAP-153; recordedBy: G. da Silva, D. Costa; individualCount:
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;
<emphasis id="6E709340CF21382BEBB2AC59C3E3A550" bold="true" pageId="0" pageNumber="62736">Location:</emphasis>
locality: Seixas Beach; verbatimDepth:
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</paragraph>
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<subSubSection id="6F43E046DD8563E20853E67496413A73" pageId="0" pageNumber="62736" type="distribution">
<paragraph id="23D827117E9E917BF5CDD69452F5141D" pageId="0" pageNumber="62736">Distribution</paragraph>
<paragraph id="7E64C701048C54D16576CBBDB1FB1A3D" pageId="0" pageNumber="62736">
Gulf of California, Gulf of Mexico and Brazilian coast (
<normalizedToken id="4FFEA482466213F9B620B1B4927F76D4" originalValue="Paraíba">Paraiba</normalizedToken>
and
<normalizedToken id="020A267F8C1FD3B4EAE06B76ECCC676D" originalValue="São">Sao</normalizedToken>
Paulo States) (
<bibRefCitation id="13C93054B72AAAE8A34D59D0148196CC" author="Amaral, A. Cecilia Z." journalOrPublisher="UNICAMP, Campinas" pageId="0" pageNumber="62736" refId="B6420637" refString="Amaral, A. Cecilia Z., Nallin, Silvana A. Henriques, Steiner, Tatiana Menchini, Forroni, Thalita Oliveira, Filho, Decio Gomes, 2013. Catalogo das especies de Annelida Polychaeta do Brasil. UNICAMP, Campinas" title="Catalogo das especies de Annelida Polychaeta do Brasil" year="2013">Amaral et al. 2013</bibRefCitation>
,
<bibRefCitation id="D3F038761DBCC709B54B46D132CDF28F" author="Costa, Dimitri Araujo" journalOrPublisher="Revista Brasileira de Gestao Ambiental e Sustentabilidade" pageId="0" pageNumber="62736" pagination="313 - 320" refId="B6421986" refString="Costa, Dimitri Araujo, Fernandes, Helder Formiga, Silva, Francisco Assis, Christoffersen, Martin Lindsey, 2017. Checklist of species of Polychaeta (Annelida) of the Seixas Beach, Joao Pessoa, state of Paraiba, Northeast Brazil. Revista Brasileira de Gestao Ambiental e Sustentabilidade 4 (8): 313 - 320" title="Checklist of species of Polychaeta (Annelida) of the Seixas Beach, Joao Pessoa, state of Paraiba, Northeast Brazil" volume="4" year="2017">Costa et al. 2017</bibRefCitation>
,
<bibRefCitation id="91069D32B58DCEF8B61FAC618ECF4731" author="Prata, Jessica" journalOrPublisher="Biota Neotropica" pageId="0" pageNumber="62736" refId="B6423190" refString="Read Geoffrey Fauchald Kristian Marphysa angelensis Fauchald, 1970. World Register of Marine Species (WoRMS), World Polychaeta database http://www.marinespecies.org/aphia.php?p=taxdetails&amp;id=329228 2020-11-07T00:00:00+02:00" year="2017">Read and Fauchald 2020f</bibRefCitation>
).
</paragraph>
<paragraph id="A50AB1924A37CA4A4F87FEB212E103E6" pageId="0" pageNumber="62736">
<emphasis id="6012FF9AF91469BEFB4F143BC3F366AA" bold="true" pageId="0" pageNumber="62736">
Distribution in
<normalizedToken id="DA3AFCE9D5D171CF22F92432844F2B73" originalValue="Paraíba">Paraiba</normalizedToken>
</emphasis>
: Seixas Beach (
<bibRefCitation id="0CFE2C3BDE8F37D2C94BAFB67768FC6F" author="Costa, Dimitri Araujo" journalOrPublisher="Revista Brasileira de Gestao Ambiental e Sustentabilidade" pageId="0" pageNumber="62736" pagination="313 - 320" refId="B6421986" refString="Costa, Dimitri Araujo, Fernandes, Helder Formiga, Silva, Francisco Assis, Christoffersen, Martin Lindsey, 2017. Checklist of species of Polychaeta (Annelida) of the Seixas Beach, Joao Pessoa, state of Paraiba, Northeast Brazil. Revista Brasileira de Gestao Ambiental e Sustentabilidade 4 (8): 313 - 320" title="Checklist of species of Polychaeta (Annelida) of the Seixas Beach, Joao Pessoa, state of Paraiba, Northeast Brazil" volume="4" year="2017">Costa et al. 2017</bibRefCitation>
; and this study).
</paragraph>
</subSubSection>
<subSubSection id="A574F521C955F80F27C8FC1A6E4392C8" pageId="0" pageNumber="62736" type="notes">
<paragraph id="C158B242149B76A99342E09F908405E6" pageId="0" pageNumber="62736">Notes</paragraph>
<paragraph id="E9285F47F7F3CE7B2171110CC4C91DF4" pageId="0" pageNumber="62736">Found inside the rhodoliths.</paragraph>
</subSubSection>
<subSubSection id="35E8D33A306F377E92EB44CF898A3F31" pageId="0" pageNumber="62736" type="diagnosis">
<paragraph id="1480771CE400C43FF5EF0D9B86920847" pageId="0" pageNumber="62736">Diagnosis</paragraph>
<paragraph id="15C47E34055E10F9EC1C5CA4468E1AC3" pageId="0" pageNumber="62736">
(
<bibRefCitation id="EF7E5D534BE4CB465424E399A9C331B5" author="Fauchald, Kristian" journalOrPublisher="Revista Colombiana de Ciencia Animal - RECIA" pageId="0" pageNumber="62736" pagination="1 - 335" refId="B6422101" refString="Fauchald, Kristian, 1970. Polychaetous annelids of the families Eunicidae, Lumbrineridae, Iphitimidae, Arabellidae, Lysaretidae and Dorvilleidae from western Mexico. In: Allan Hancock Monographs in Marine Biology.: 1 - 335" title="Polychaetous annelids of the families Eunicidae, Lumbrineridae, Iphitimidae, Arabellidae, Lysaretidae and Dorvilleidae from western Mexico" volumeTitle="Allan Hancock Monographs in Marine Biology." year="1970">Fauchald 1970</bibRefCitation>
,
<bibRefCitation id="3637005A081725A16FBB24FEEC515AC7" author="Paxton, Hannelore" journalOrPublisher="Zoosymposia" pageId="0" pageNumber="62736" pagination="241 - 264" refId="B6423113" refString="Paxton, Hannelore, 2009. Phylogeny of Eunicida (Annelida) based on morphology of jaws. Zoosymposia 2 (1): 241 - 264" title="Phylogeny of Eunicida (Annelida) based on morphology of jaws" volume="2" year="2009">Paxton 2009</bibRefCitation>
,
<bibRefCitation id="E29C07AEBAE9BE239D5E35934B919E72" author="Salazar-Vallejo, Sergio I." journalOrPublisher="Revista de Biologia Tropical" pageId="0" pageNumber="62736" pagination="1481 - 1498" refId="B6426677" refString="Salazar-Vallejo, Sergio I., Carrera-Parra, Luis F., 1997. Eunicidos (Polychaeta) del Caribe mexicano con claves para las especies del Gran Caribe: Fauchaldius, Lysidice, Marphysa, Nematonereis y Palola. Revista de Biologia Tropical 45 (4): 1481 - 1498" title="Eunicidos (Polychaeta) del Caribe mexicano con claves para las especies del Gran Caribe: Fauchaldius, Lysidice, Marphysa, Nematonereis y Palola" volume="45" year="1997">Salazar-Vallejo and Carrera-Parra 1997</bibRefCitation>
): Prostomium with a short anterior incision, with two eyes and five smooth antennae. Jaws eulabidognath-type (asymmetrical, posterior parts dentate to
<taxonomicName id="68A929795AD03C3564A5E6DA6F77A403" authorityName="Milne-Edwards" authorityYear="1875" class="Polychaeta" family="Paguridae" genus="Mithraculus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Mithraculus forceps" order="Phyllodocida" pageId="0" pageNumber="62736" phylum="Annelida" rank="species" species="forceps">Mithraculus forceps</taxonomicName>
-like, short carriers). Formula (maxillae): 1+1, 3+5, 6+0, 3+8 and 1+1. Peristomial cirri absent. Ramified branchiae (1 to 3 filaments) from chaetiger 9 (Fig.
<figureCitation id="A7780F9B55F73320674D1982D0812A41" captionStartId="F6439127" pageId="0" pageNumber="62736">3</figureCitation>
<figureCitation id="A1B3693CA462FB3420957E92A6A3EB37" captionStartId="F6439133" pageId="0" pageNumber="62736">a</figureCitation>
). Anterior notopodial cirri enlarged and posterior ones digitiform, both types longer than ventral ones. Neuropodia with anterior cirri globular, posterior ones conical; limbate, pectinates, falcigers and spinigers chaetae; 1-3 dark aciculae by chaetiger and dark subacicular bidentate hooks.
</paragraph>
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</treatment>
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(
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Figures 1
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3
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)
</paragraph>
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<emphasis id="0CD8D70BFF98283A51CDB138C40F3CCD" bold="true" box="[151,316,1659,1683]" pageId="0" pageNumber="597">Female. Head.</emphasis>
Anterior half of frons with white and brown pubescence, posterior half with sparse and fine pubescence not obscuring integument. Vertex concave, with white pubescence and some black sparse setae. Antennal tubercles, scape and pedicel with white pubescence; basal half of antennomere III with dense white pubescence, distal half with short and sparse pubescence, not obscuring integument, apex with a black tuft, denser ventrally; basal half of antennomere IV with white pubescence and distal half with short and sparse pubescence, not obscuring integument; antennomeres V and VI with basal third with white pubescence and remainder of surface with short and sparse pubescence, not obscuring integument. Antennomeres VII to XI with short, sparse golden pubescence, not obscuring surface. Distance between upper eye lobes 0.50 times length of scape; in frontal view, distance between lower eye lobes 0.71 times length of scape. Eyes divided, upper eye lobes separated from lower lobes. Genae black, with decumbent white setae. Clypeus brownishorange, truncate, glabrous. Labrum brownish-orange, truncate, with long setae at base and with short brownish-orange setae at apex. Mandibles black, with microrugosities on base. Maxillary and palpomere light black. Antennal formula (ratio) based on length of antennomere III: scape = 0.61; pedicel = 0.19; IV = 0.47; V = 0.32; VI = 0.23; VII = 0.22; VIII = 0.17; IX = 0.15; X = 0.12; XI = 0.16.
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<emphasis id="0CD8D70BFF99283B519DB7A1C4163AA4" bold="true" box="[199,293,226,250]" pageId="1" pageNumber="598">Thorax:</emphasis>
Prothorax wider than longer. Pronotum with sparse long black setae, median region of anterior margin with a black spot; with three parallels glabrous and elongated tubercles, lateral tubercles occupy half of the length of pronotum. Lateral tubercles of prothorax acute. Prosternum black, with some sparse white setae. Mesoventrite with some punctures, and abundant long white setae. Metaventrite finely punctate centrally, with abundant white setae laterally. Metanepisternum with abundant white setae. Epipleura with abundant long white hairs.
<emphasis id="0CD8D70BFF99283B529AB631C1233BD4" bold="true" box="[960,1040,370,394]" pageId="1" pageNumber="598">Elytra.</emphasis>
Surface with fine white pubescence, interspersed with black spots, and some sparse long black setae. Each elytron with a longitudinal carina, which is curved at base and surrounds a brownish depression, extending parallel but not reaching elytral apex. Apex of elytra rounded.
<emphasis id="0CD8D70BFF99283B51CDB69DC5E03BA8" bold="true" box="[151,211,478,502]" pageId="1" pageNumber="598">Legs.</emphasis>
Femora pedunculate, with abundant white pubescence dorsally, reddish ventrally, integument black, with some white setae, at apex with abundant white pubescence. Tibiae clothed in setae of alternating colors, between black and white pubescence.
<emphasis id="0CD8D70BFF99283B503BB566C4E33863" bold="true" box="[353,464,549,573]" pageId="1" pageNumber="598">Abdomen</emphasis>
. Ventrites black with white setae. Apex of ventrite V truncate.
</paragraph>
<caption id="6AD35B91FF99283B51CDB111C6113CD0" httpUri="https://zenodo.org/record/2702423/files/figure.png" pageId="1" pageNumber="598" startId="1.[151,270,1618,1642]" targetBox="[168,1416,661,1585]" targetPageId="1">
<paragraph id="3E130B19FF99283B51CDB111C6113CD0" blockId="1.[151,1436,1618,1678]" pageId="1" pageNumber="598">
<emphasis id="0CD8D70BFF99283B51CDB111C4773C34" bold="true" box="[151,324,1618,1642]" pageId="1" pageNumber="598">FIGURES 1–6.</emphasis>
<taxonomicName id="F9AC709AFF99283B5016B111C73B3C34" authorityName="Marinoni &amp; Martins" authorityYear="1978" box="[332,520,1618,1642]" class="Insecta" family="Cerambycidae" genus="Anasillus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="crinitus">
<emphasis id="0CD8D70BFF99283B5016B111C73B3C34" box="[332,520,1618,1642]" italics="true" pageId="1" pageNumber="598">Anasillus crinitus</emphasis>
</taxonomicName>
.
<emphasis id="0CD8D70BFF99283B534DB111C7753C34" bold="true" box="[535,582,1618,1642]" pageId="1" pageNumber="598">1–3,</emphasis>
female;
<emphasis id="0CD8D70BFF99283B53FFB111C78A3C34" bold="true" box="[677,697,1618,1642]" pageId="1" pageNumber="598">1,</emphasis>
dorsal view;
<emphasis id="0CD8D70BFF99283B5216B111C6523C34" bold="true" box="[844,865,1618,1642]" pageId="1" pageNumber="598">2,</emphasis>
lateral view;
<emphasis id="0CD8D70BFF99283B52AFB111C13A3C34" bold="true" box="[1013,1033,1618,1642]" pageId="1" pageNumber="598">3,</emphasis>
ventral view.
<emphasis id="0CD8D70BFF99283B55F9B111C1E23C34" bold="true" box="[1187,1233,1618,1642]" pageId="1" pageNumber="598">4–6,</emphasis>
holotype, male;
<emphasis id="0CD8D70BFF99283B54D2B111C0AF3C34" bold="true" box="[1416,1436,1618,1642]" pageId="1" pageNumber="598">4,</emphasis>
dorsal view;
<emphasis id="0CD8D70BFF99283B507BB135C4063CD0" bold="true" box="[289,309,1654,1678]" pageId="1" pageNumber="598">5,</emphasis>
lateral view;
<emphasis id="0CD8D70BFF99283B509DB135C4E83CD0" bold="true" box="[455,475,1654,1678]" pageId="1" pageNumber="598">6,</emphasis>
detail of apical antennomeres.
</paragraph>
</caption>
<paragraph id="3E130B19FF99283B519DB1FDC1523CA4" blockId="1.[151,1437,1726,2038]" pageId="1" pageNumber="598">
<emphasis id="0CD8D70BFF99283B519DB1FDC4613C88" bold="true" box="[199,338,1726,1750]" pageId="1" pageNumber="598">Dimensions.</emphasis>
Female. Total length, 10.30; prothoracic length, 2.01; anterior prothoracic width, 2.70; posterior prothoracic width, 2.81; widest prothoracic width, 3.83; humeral width, 4.52; elytral length, 6.50.
</paragraph>
</subSubSection>
<subSubSection id="76B65892FF99283B519DB045C7D23D1C" pageId="1" pageNumber="598" type="materials_examined">
<paragraph id="3E130B19FF99283B519DB045C7D23D1C" blockId="1.[151,1437,1726,2038]" pageId="1" pageNumber="598">
<emphasis id="0CD8D70BFF99283B519DB045C4993D40" bold="true" box="[199,426,1798,1822]" pageId="1" pageNumber="598">Material examined.</emphasis>
<materialsCitation id="8EC40144FF99283B50EFB045C7EE3D1C" ID-GBIF-Occurrence="2239654644" collectingDate="2005-10" collectionCode="MEPB" country="Colombia" elevation="2300" location="Municipio de Guarne" pageId="1" pageNumber="598" specimenCount="1" specimenCount-female="1" stateProvince="Antioquia">
<collectingCountry id="46BB4B89FF99283B50EFB045C7113D40" box="[437,546,1798,1822]" name="Colombia" pageId="1" pageNumber="598">Colombia</collectingCountry>
: department of
<collectingRegion id="FC68C5FBFF99283B538FB045C6713D40" box="[725,834,1798,1822]" country="Colombia" name="Antioquia" pageId="1" pageNumber="598">Antioquia</collectingRegion>
,
<location id="3B735DC2FF99283B5208B045C1713D40" LSID="urn:lsid:plazi:treatment:B605BA0FFF98283851CDB172C1E73A8B:3B735DC2FF99283B5208B045C1713D40" box="[850,1090,1798,1822]" country="Colombia" name="Municipio de Guarne" pageId="1" pageNumber="598" stateProvince="Antioquia">Municipio de Guarne</location>
, Parque Piedras Blancas.
<quantity id="F954A6FCFF99283B543DB045C5823D1C" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="598" unit="m" value="2300.0">
<elevation id="B581EC2AFF99283B543DB045C5823D1C" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="598" unit="m" value="2300.0">2300 m</elevation>
</quantity>
.s.n.m.
<specimenCount id="28AAC090FF99283B51A0B069C46A3D1C" box="[250,345,1834,1858]" pageId="1" pageNumber="598" type="female">1 female</specimenCount>
, “colecta manual”.
<date id="4A122DD9FF99283B5373B069C74A3D1C" box="[553,633,1834,1858]" pageId="1" pageNumber="598" value="2005-10">
<collectingDate id="5A56D431FF99283B5373B069C74A3D1C" box="[553,633,1834,1858]" pageId="1" pageNumber="598" value="2005-10">X.2005</collectingDate>
</date>
(
<collectionCode id="58BD93DCFF99283B53DDB069C7E53D1C" box="[647,726,1834,1858]" collectionName="MEPB" pageId="1" pageNumber="598">MEPB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B65892FF992838519DB00DC1E73A8B" lastPageId="2" lastPageNumber="599" pageId="1" pageNumber="598" type="discussion">
<paragraph id="3E130B19FF992838519DB00DC1E73A8B" blockId="1.[151,1437,1726,2038]" lastBlockId="2.[151,1436,154,214]" lastPageId="2" lastPageNumber="599" pageId="1" pageNumber="598">
<emphasis id="0CD8D70BFF99283B519DB00DC4053D38" bold="true" box="[199,310,1870,1894]" pageId="1" pageNumber="598">Remarks.</emphasis>
<taxonomicName id="F9AC709AFF99283B5067B00DC4CB3D38" authorityName="Marinoni &amp; Martins" authorityYear="1978" box="[317,504,1870,1894]" class="Insecta" family="Cerambycidae" genus="Anasillus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="species" species="crinitus">
<emphasis id="0CD8D70BFF99283B5067B00DC4CB3D38" box="[317,504,1870,1894]" italics="true" pageId="1" pageNumber="598">Anasillus crinitus</emphasis>
</taxonomicName>
was known only by the
<typeStatus id="E117B5BBFF99283B525BB00DC66D3D38" box="[769,862,1870,1894]" pageId="1" pageNumber="598" type="holotype">holotype</typeStatus>
male (
<figureCitation id="A697179CFF99283B52F3B00DC13D3D38" box="[937,1038,1870,1894]" captionStart="FIGURES 1–6" captionStartId="1.[151,270,1618,1642]" captionTargetBox="[168,1416,661,1585]" captionTargetId="figure@1.[151,1436,592,1594]" captionTargetPageId="1" captionText="FIGURES 1–6. Anasillus crinitus. 1–3, female; 1, dorsal view; 2, lateral view; 3, ventral view. 4–6, holotype, male; 4, dorsal view; 5, lateral view; 6, detail of apical antennomeres." httpUri="https://zenodo.org/record/2702423/files/figure.png" pageId="1" pageNumber="598">
Figs. 4
<emphasis id="0CD8D70BFF99283B52A9B00DC1333D38" bold="true" box="[1011,1024,1870,1894]" pageId="1" pageNumber="598">–</emphasis>
6
</figureCitation>
) deposited at MZSP collection, from
<collectingRegion id="FC68C5FBFF99283B51CDB031C43A3DD4" box="[151,265,1906,1930]" country="Peru" name="Cajamarca" pageId="1" pageNumber="598">Cajamarca</collectingRegion>
department,
<collectingCountry id="46BB4B89FF99283B50C2B030C4FF3DD4" box="[408,460,1907,1930]" name="Peru" pageId="1" pageNumber="598">Peru</collectingCountry>
. Examination of the
<typeStatus id="E117B5BBFF99283B53F4B031C6383DD4" box="[686,779,1906,1930]" pageId="1" pageNumber="598" type="holotype">holotype</typeStatus>
revealed that the main differences between male and female are: presence of a tuft of black hairs at apex of antennomere XI (
<figureCitation id="A697179CFF99283B520BB0D5C6A13DF0" box="[849,914,1942,1966]" captionStart="FIGURES 1–6" captionStartId="1.[151,270,1618,1642]" captionTargetBox="[168,1416,661,1585]" captionTargetId="figure@1.[151,1436,592,1594]" captionTargetPageId="1" captionText="FIGURES 1–6. Anasillus crinitus. 1–3, female; 1, dorsal view; 2, lateral view; 3, ventral view. 4–6, holotype, male; 4, dorsal view; 5, lateral view; 6, detail of apical antennomeres." httpUri="https://zenodo.org/record/2702423/files/figure.png" pageId="1" pageNumber="598">Fig. 6</figureCitation>
) and orange pubescence on femora (
<figureCitation id="A697179CFF99283B5445B0D5C0533DF0" box="[1311,1376,1942,1966]" captionStart="FIGURES 1–6" captionStartId="1.[151,270,1618,1642]" captionTargetBox="[168,1416,661,1585]" captionTargetId="figure@1.[151,1436,592,1594]" captionTargetPageId="1" captionText="FIGURES 1–6. Anasillus crinitus. 1–3, female; 1, dorsal view; 2, lateral view; 3, ventral view. 4–6, holotype, male; 4, dorsal view; 5, lateral view; 6, detail of apical antennomeres." httpUri="https://zenodo.org/record/2702423/files/figure.png" pageId="1" pageNumber="598">Fig. 5</figureCitation>
). According to
<bibRefCitation id="5A3D76E8FF99283B5050B0F9C7013D8C" author="Marinoni, R. &amp; Martins, U." box="[266,562,1978,2002]" pageId="1" pageNumber="598" pagination="173 - 193" refId="ref1890" refString="Marinoni, R. C &amp; Martins, U. (1978) Notas sinonimicas e novas especies em Acanthoderini (Coleoptera: Cerambycidae: Lamiinae). Papeis Avulsos de Zoologia, 31 (11), 173 - 193." type="journal article" year="1978">Marinoni &amp; Martins (1978)</bibRefCitation>
,
<taxonomicName id="F9AC709AFF99283B5367B0F9C7933D8C" authorityName="Marinoni &amp; Martins" authorityYear="1978" box="[573,672,1978,2002]" class="Insecta" family="Cerambycidae" genus="Anasillus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="genus">
<emphasis id="0CD8D70BFF99283B5367B0F9C7933D8C" box="[573,672,1978,2002]" italics="true" pageId="1" pageNumber="598">Anasillus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="F9AC709AFF99283B5271B0F8C10C3D8C" authority="Thomson, 1864" authorityName="Thomson" authorityYear="1864" box="[811,1087,1978,2002]" class="Insecta" family="Cerambycidae" genus="Discopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="genus">
<emphasis id="0CD8D70BFF99283B5271B0F8C6BD3D8C" box="[811,910,1979,2002]" italics="true" pageId="1" pageNumber="598">Discopus</emphasis>
Thomson, 1864
</taxonomicName>
;
<taxonomicName id="F9AC709AFF99283B5510B0F9C0AB3D8C" authority="Schwarzer, 1930" authorityName="Schwarzer" authorityYear="1930" box="[1098,1432,1978,2002]" class="Insecta" family="Cerambycidae" genus="Paradiscopus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="genus">
<emphasis id="0CD8D70BFF99283B5510B0F9C1EE3D8C" box="[1098,1245,1978,2002]" italics="true" pageId="1" pageNumber="598">Paradiscopus</emphasis>
Schwarzer, 1930
</taxonomicName>
;
<taxonomicName id="F9AC709AFF99283B51CDB09CC4B83DA8" authority="Bates, 1880" authorityName="Bates" authorityYear="1880" box="[151,395,2014,2038]" class="Insecta" family="Cerambycidae" genus="Tetrasarus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="genus">
<emphasis id="0CD8D70BFF99283B51CDB09CC43B3DA8" box="[151,264,2015,2038]" italics="true" pageId="1" pageNumber="598">Tetrasarus</emphasis>
Bates, 1880
</taxonomicName>
and
<taxonomicName id="F9AC709AFF99283B50E4B09DC7F13DA8" authority="Bates, 1880" authorityName="Bates" authorityYear="1880" box="[446,706,2014,2038]" class="Insecta" family="Cerambycidae" genus="Plagiosarus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="598" phylum="Arthropoda" rank="genus">
<emphasis id="0CD8D70BFF99283B50E4B09DC70C3DA8" box="[446,575,2014,2038]" italics="true" pageId="1" pageNumber="598">Plagiosarus</emphasis>
Bates, 1880
</taxonomicName>
by the presence of a tuft of setae at antennomere III, but differs from them by the presence of a strip of setae at the epipleura and at the middle of the tibiae and by the presence of a tuft of setae at apex of antennomere XI. The specimen examined represents a
<emphasis id="0CD8D70BFF9A28385215B7FDC1E73A8B" bold="true" box="[847,1236,189,214]" pageId="2" pageNumber="599">
new country record for
<collectingCountry id="46BB4B89FF9A28385507B7FEC1FD3A8B" box="[1117,1230,189,213]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

115
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<paragraph id="3E130B19FF9A283851CDB075C6A93D10" blockId="2.[151,1437,1810,2014]" box="[151,922,1846,1870]" pageId="2" pageNumber="599">
<subSubSection id="76B65892FF9A283851CDB075C6053D10" box="[151,822,1846,1870]" pageId="2" pageNumber="599" type="nomenclature">
<taxonomicName id="F9AC709AFF9A283851CDB075C6053D10" ID-CoL="8VXG" authority="Chemsak &amp; Hovore, 2002" authorityName="Chemsak &amp; Hovore" authorityYear="2002" box="[151,822,1846,1870]" class="Insecta" family="Cerambycidae" genus="Acanthoderes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="ariasi" subGenus="Acanthoderes">
<emphasis id="0CD8D70BFF9A283851CDB075C41A3D10" box="[151,297,1846,1870]" italics="true" pageId="2" pageNumber="599">Acanthoderes</emphasis>
(
<emphasis id="0CD8D70BFF9A28385062B075C4F93D10" box="[312,458,1846,1870]" italics="true" pageId="2" pageNumber="599">Acanthoderes</emphasis>
)
<emphasis id="0CD8D70BFF9A28385080B075C72B3D13" box="[474,536,1846,1869]" italics="true" pageId="2" pageNumber="599">ariasi</emphasis>
<bibRefCitation id="5A3D76E8FF9A28385344B075C6053D10" author="Chemsak, J. A. &amp; Hovore, F. T." box="[542,822,1846,1870]" pageId="2" pageNumber="599" pagination="1 - 32" refId="ref1791" refString="Chemsak, J. A. &amp; Hovore, F. T. (2002) New Mexican and Central American species of Acanthoderini with notes on others, part I. (Coleoptera, Cerambycidae, Lamiinae). Les Cahiers Magellanes, 15, 1 - 32." type="journal article" year="2002">Chemsak &amp; Hovore, 2002</bibRefCitation>
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)
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<paragraph id="3E130B19FF9A283851CDB019C7823D2C" blockId="2.[151,1437,1810,2014]" box="[151,689,1882,1906]" pageId="2" pageNumber="599">
Distribution:
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,
<collectingCountry id="46BB4B89FF9A283850F2B019C4CF3D2F" box="[424,508,1882,1905]" name="Panama" pageId="2" pageNumber="599">Panama</collectingCountry>
(Monné 2018b).
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Material examined:
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<collectingCountry id="46BB4B89FF9A28385031B03DC4E43DC8" box="[363,471,1918,1942]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
,
<collectingRegion id="FC68C5FBFF9A28385085B03DC77F3DC8" box="[479,588,1918,1942]" country="Colombia" name="Antioquia" pageId="2" pageNumber="599">Antioquia</collectingRegion>
,
<collectingMunicipality id="DE779163FF9A2838530CB03DC7DA3DC8" box="[598,745,1918,1942]" pageId="2" pageNumber="599">San Jerónimo</collectingMunicipality>
,
<location id="3B735DC2FF9A283853A9B03DC1643DC8" LSID="urn:lsid:plazi:treatment:B605BA0FFF9A283851CDB075C7133D83:3B735DC2FF9A283853A9B03DC1643DC8" box="[755,1111,1918,1942]" country="Colombia" municipality="San Jeronimo" name="Parque recreativo los Tamarindos" pageId="2" pageNumber="599" stateProvince="Antioquia">Parque recreativo los Tamarindos</location>
.
<quantity id="F954A6FCFF9A2838553AB03EC1963DCB" box="[1120,1189,1917,1941]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="2" pageNumber="599" unit="m" value="925.0">
<elevation id="B581EC2AFF9A2838553AB03EC1BE3DCB" box="[1120,1165,1917,1941]" pageId="2" pageNumber="599">925</elevation>
m
</quantity>
.s.n.m. “
<collectingMethod id="E7ED730EFF9A283855A9B03DC4423DE4" pageId="2" pageNumber="599">Atraído por luz. Colecta manual</collectingMethod>
”.
<date id="4A122DD9FF9A283850DEB0E1C4CD3DE4" box="[388,510,1954,1978]" pageId="2" pageNumber="599" value="2006-04-27">
<collectingDate id="5A56D431FF9A283850DEB0E1C4CD3DE4" box="[388,510,1954,1978]" pageId="2" pageNumber="599" value="2006-04-27">27.IV.2006</collectingDate>
</date>
.
<collectorName id="93596ECFFF9A2838535CB0E1C75B3DE4" box="[518,616,1954,1978]" pageId="2" pageNumber="599">A. Vélez</collectorName>
. (
<collectionCode id="58BD93DCFF9A28385323B0E1C7FB3DE4" box="[633,712,1954,1978]" collectionName="MEPB" pageId="2" pageNumber="599">MEPB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="76B65892FF9A283851CDB085C7133D83" box="[151,544,1989,2014]" pageId="2" pageNumber="599" type="distribution">
<paragraph id="3E130B19FF9A283851CDB085C7133D83" blockId="2.[151,1437,1810,2014]" box="[151,544,1989,2014]" pageId="2" pageNumber="599">
<emphasis id="0CD8D70BFF9A283851CDB085C7133D83" bold="true" box="[151,544,1989,2014]" pageId="2" pageNumber="599">
New country record for
<collectingCountry id="46BB4B89FF9A283850F0B086C7283D83" box="[426,539,1989,2013]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
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113
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<emphasis id="0CD8D70BFF9A283851CDB179C45C3C0C" box="[151,367,1594,1618]" italics="true" pageId="2" pageNumber="599">Alcidion sulphurifer</emphasis>
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<paragraph id="3E130B19FF9A283851CDB11DC6283C28" blockId="2.[151,1437,1521,1762]" box="[151,795,1630,1654]" pageId="2" pageNumber="599">
Distribution:
<collectingCountry id="46BB4B89FF9A2838507CB11DC4693C2B" box="[294,346,1630,1653]" name="Peru" pageId="2" pageNumber="599">Peru</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A2838503EB11DC49B3C28" box="[356,424,1630,1654]" name="Brazil" pageId="2" pageNumber="599">Brazil</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A283850E9B11DC73D3C28" box="[435,526,1630,1654]" name="Ecuador" pageId="2" pageNumber="599">Ecuador</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A28385342B11DC7543C28" box="[536,615,1630,1654]" name="Bolivia" pageId="2" pageNumber="599">Bolivia</collectingCountry>
(Monné 2018b).
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Material examined:
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<collectingCountry id="46BB4B89FF9A28385029B1C1C4EC3CC4" box="[371,479,1666,1690]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
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,
<collectingMunicipality id="DE779163FF9A28385309B1C1C7F63CC4" box="[595,709,1666,1690]" pageId="2" pageNumber="599">Quimbaya</collectingMunicipality>
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.
<quantity id="F954A6FCFF9A28385280B1C1C1033CC7" box="[986,1072,1666,1690]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="599" unit="m" value="1100.0">
<elevation id="B581EC2AFF9A28385280B1C1C13D3CC4" box="[986,1038,1666,1690]" pageId="2" pageNumber="599">1100</elevation>
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<date id="4A122DD9FF9A2838552CB1C1C1C13CC4" box="[1142,1266,1666,1690]" pageId="2" pageNumber="599" value="2003-09-27">
<collectingDate id="5A56D431FF9A2838552CB1C1C1C13CC4" box="[1142,1266,1666,1690]" pageId="2" pageNumber="599" value="2003-09-27">27.IX.2003</collectingDate>
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<collectorName id="93596ECFFF9A283855A5B1C1C0553CC7" box="[1279,1382,1666,1690]" pageId="2" pageNumber="599">García L.</collectorName>
Col. (
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<emphasis id="0CD8D70BFF9A283851CDB189C7133CBF" bold="true" box="[151,544,1737,1762]" pageId="2" pageNumber="599">
New country record for
<collectingCountry id="46BB4B89FF9A283850F0B18AC7283CBF" box="[426,539,1737,1761]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
.
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113
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<treatment id="B605BA0FFF9A283851CDB259C7133F9F" ID-DOI="http://doi.org/10.5281/zenodo.5931331" ID-GBIF-Taxon="157119698" ID-Zenodo-Dep="5931331" LSID="urn:lsid:plazi:treatment:B605BA0FFF9A283851CDB259C7133F9F" httpUri="http://treatment.plazi.org/id/B605BA0FFF9A283851CDB259C7133F9F" lastPageNumber="599" pageId="2" pageNumber="599">
<paragraph id="3E130B19FF9A283851CDB259C74A3F6C" blockId="2.[151,1436,1270,1474]" box="[151,633,1306,1330]" pageId="2" pageNumber="599">
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<taxonomicName id="F9AC709AFF9A283851CDB259C7253F6C" ID-CoL="XH6D" authority="(Bates, 1870)" baseAuthorityName="Bates" baseAuthorityYear="1870" box="[151,534,1306,1330]" class="Insecta" family="Cerambycidae" genus="Compsibidion" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="charile">
<emphasis id="0CD8D70BFF9A283851CDB259C4B33F6C" box="[151,384,1306,1330]" italics="true" pageId="2" pageNumber="599">Compsibidion charile</emphasis>
(Bates, 1870)
</taxonomicName>
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<subSubSection id="76B65892FF9A28385346B259C74A3F6C" box="[540,633,1306,1330]" pageId="2" pageNumber="599" type="description">
(
<figureCitation id="A697179CFF9A2838537EB259C7433F6C" box="[548,624,1306,1330]" captionStart="FIGURES 7–13. 7" captionStartId="3.[151,270,1106,1130]" captionTargetBox="[153,1432,190,1052]" captionTargetId="figure@3.[151,1436,165,1082]" captionTargetPageId="3" captionText="FIGURES 7–13. 7, Coleoxestia vittata (Thomson, 1861), dorsal view; 8, Sphallenopsis pilosovittata (Bates, 1872), dorsal view; 9, Aneflus (Protaneflus) minutivestis Chemsak &amp; Linsley, 1963, dorsal view; 10, Mallocera amazonica Bates, 1870, dorsal view; 11, Compsibidion charile (Bates, 1870), dorsal view; 12, Alcidion sulphurifer (White, 1855), dorsal view; 13, Acanthoderes (Acanthoderes) ariasi Chemsak &amp; Hovore, 2002, dorsal view." httpUri="https://zenodo.org/record/2702425/files/figure.png" pageId="2" pageNumber="599">Fig. 11</figureCitation>
)
</subSubSection>
</paragraph>
<subSubSection id="76B65892FF9A283851CDB27DC6183F08" box="[151,811,1342,1366]" pageId="2" pageNumber="599" type="distribution">
<paragraph id="3E130B19FF9A283851CDB27DC6183F08" blockId="2.[151,1436,1270,1474]" box="[151,811,1342,1366]" pageId="2" pageNumber="599">
Distribution:
<collectingCountry id="46BB4B89FF9A2838507CB27DC4B23F08" box="[294,385,1342,1366]" name="Ecuador" pageId="2" pageNumber="599">Ecuador</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A283850D1B27DC71F3F08" box="[395,556,1342,1366]" name="French Guiana" pageId="2" pageNumber="599">French Guiana</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A2838536CB27DC7443F08" box="[566,631,1342,1366]" name="Brazil" pageId="2" pageNumber="599">Brazil</collectingCountry>
(Monné 2018a).
</paragraph>
</subSubSection>
<subSubSection id="76B65892FF9A283851CDB221C65E3FC0" pageId="2" pageNumber="599" type="materials_examined">
<paragraph id="3E130B19FF9A283851CDB221C65E3FC0" blockId="2.[151,1436,1270,1474]" pageId="2" pageNumber="599">
Material examined:
<materialsCitation id="8EC40144FF9A2838502EB221C65E3FC0" ID-GBIF-Occurrence="2239654645" collectedFrom="Borde de bosque" collectingDate="2005-09-25" collectingMethod="Trampa de luz negra" collectionCode="MEPB" collectorName="A. Velez" country="Colombia" elevation="450" location="Vereda las cristalinas, Reserva U. N. A." municipality="Puerto Berrio" pageId="2" pageNumber="599" specimenCount="1" stateProvince="Antioquia">
<collectingCountry id="46BB4B89FF9A2838502EB221C4D33F24" box="[372,480,1378,1402]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
,
<collectingRegion id="FC68C5FBFF9A283850B6B221C76A3F24" box="[492,601,1378,1402]" country="Colombia" name="Antioquia" pageId="2" pageNumber="599">Antioquia</collectingRegion>
,
<collectingMunicipality id="DE779163FF9A2838533DB221C7CD3F24" box="[615,766,1378,1402]" pageId="2" pageNumber="599">Puerto Berrio</collectingMunicipality>
,
<location id="3B735DC2FF9A28385251B221C1983F27" LSID="urn:lsid:plazi:treatment:B605BA0FFF9A283851CDB259C7133F9F:3B735DC2FF9A28385251B221C1983F27" box="[779,1195,1377,1402]" country="Colombia" municipality="Puerto Berrio" name="Vereda las cristalinas, Reserva U. N. A." pageId="2" pageNumber="599" stateProvince="Antioquia">Vereda las cristalinas, Reserva U.N.A.</location>
<quantity id="F954A6FCFF9A283855F1B222C0183F27" box="[1195,1323,1377,1401]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="599" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
<elevation id="B581EC2AFF9A283855F1B222C0183F27" box="[1195,1323,1377,1401]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="599" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400-500 m</elevation>
</quantity>
.
<collectedFrom id="8DB048E4FF9A28385462B221C4203FC0" pageId="2" pageNumber="599">Borde de bosque</collectedFrom>
, “
<collectingMethod id="E7ED730EFF9A2838507DB2C5C73A3FC0" box="[295,521,1414,1438]" pageId="2" pageNumber="599">Trampa de luz negra</collectingMethod>
”.
<date id="4A122DD9FF9A28385347B2C6C7A93FC3" box="[541,666,1413,1437]" pageId="2" pageNumber="599" value="2005-09-25">
<collectingDate id="5A56D431FF9A28385347B2C6C7A93FC3" box="[541,666,1413,1437]" pageId="2" pageNumber="599" value="2005-09-25">25.IX.2005</collectingDate>
</date>
.
<collectorName id="93596ECFFF9A283853F9B2C5C6313FC0" box="[675,770,1414,1438]" pageId="2" pageNumber="599">A. Veléz</collectorName>
. (
<collectionCode id="58BD93DCFF9A28385249B2C5C6513FC0" box="[787,866,1414,1438]" collectionName="MEPB" pageId="2" pageNumber="599">MEPB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="76B65892FF9A283851CDB2E9C7133F9F" box="[151,544,1449,1474]" pageId="2" pageNumber="599" type="distribution">
<paragraph id="3E130B19FF9A283851CDB2E9C7133F9F" blockId="2.[151,1436,1270,1474]" box="[151,544,1449,1474]" pageId="2" pageNumber="599">
<emphasis id="0CD8D70BFF9A283851CDB2E9C7133F9F" bold="true" box="[151,544,1449,1474]" pageId="2" pageNumber="599">
New country record for
<collectingCountry id="46BB4B89FF9A283850F0B2EAC7283F9F" box="[426,539,1449,1473]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

108
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@ -0,0 +1,108 @@
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<mods:title id="EE1C6AFAD3F113F382A14461AA314815">Description of the female of Anasillus crinitus Marinoni &amp; Martins 1978 (Coleoptera: Lamiinae: Acanthoderini) and new distribution records of longhorned beetles for Colombia</mods:title>
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<treatment id="B605BA0FFF9A283851CDB301C7133E9B" ID-DOI="http://doi.org/10.5281/zenodo.5931329" ID-GBIF-Taxon="157119696" ID-Zenodo-Dep="5931329" LSID="urn:lsid:plazi:treatment:B605BA0FFF9A283851CDB301C7133E9B" httpUri="http://treatment.plazi.org/id/B605BA0FFF9A283851CDB301C7133E9B" lastPageNumber="599" pageId="2" pageNumber="599">
<paragraph id="3E130B19FF9A283851CDB301C75A3E04" blockId="2.[151,1405,1090,1222]" box="[151,617,1090,1114]" pageId="2" pageNumber="599">
<subSubSection id="76B65892FF9A283851CDB301C7363E04" box="[151,517,1090,1114]" pageId="2" pageNumber="599" type="nomenclature">
<taxonomicName id="F9AC709AFF9A283851CDB301C7363E04" ID-CoL="734TC" authority="Bates, 1870" authorityName="Bates" authorityYear="1870" box="[151,517,1090,1114]" class="Insecta" family="Cerambycidae" genus="Mallocera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="599" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="0CD8D70BFF9A283851CDB301C4B23E07" box="[151,385,1090,1114]" italics="true" pageId="2" pageNumber="599">Mallocera amazonica</emphasis>
Bates, 1870
</taxonomicName>
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<subSubSection id="76B65892FF9A28385356B301C75A3E04" box="[524,617,1090,1114]" pageId="2" pageNumber="599" type="description">
(
<figureCitation id="A697179CFF9A2838534EB301C7523E04" box="[532,609,1090,1114]" captionStart="FIGURES 7–13. 7" captionStartId="3.[151,270,1106,1130]" captionTargetBox="[153,1432,190,1052]" captionTargetId="figure@3.[151,1436,165,1082]" captionTargetPageId="3" captionText="FIGURES 7–13. 7, Coleoxestia vittata (Thomson, 1861), dorsal view; 8, Sphallenopsis pilosovittata (Bates, 1872), dorsal view; 9, Aneflus (Protaneflus) minutivestis Chemsak &amp; Linsley, 1963, dorsal view; 10, Mallocera amazonica Bates, 1870, dorsal view; 11, Compsibidion charile (Bates, 1870), dorsal view; 12, Alcidion sulphurifer (White, 1855), dorsal view; 13, Acanthoderes (Acanthoderes) ariasi Chemsak &amp; Hovore, 2002, dorsal view." httpUri="https://zenodo.org/record/2702425/files/figure.png" pageId="2" pageNumber="599">Fig. 10</figureCitation>
)
</subSubSection>
</paragraph>
<subSubSection id="76B65892FF9A283851CDB325C1AD3E20" box="[151,1182,1126,1150]" pageId="2" pageNumber="599" type="distribution">
<paragraph id="3E130B19FF9A283851CDB325C1AD3E20" blockId="2.[151,1405,1090,1222]" box="[151,1182,1126,1150]" pageId="2" pageNumber="599">
Distribution:
<collectingCountry id="46BB4B89FF9A2838507CB325C4C73E20" box="[294,500,1126,1150]" name="Trinidad and Tobago" pageId="2" pageNumber="599">Trinidad &amp; Tobago</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A283850A5B325C7933E20" box="[511,672,1126,1150]" name="French Guiana" pageId="2" pageNumber="599">French Guiana</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A283853F0B325C7DD3E20" box="[682,750,1126,1150]" name="Brazil" pageId="2" pageNumber="599">Brazil</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A283853A3B325C6673E20" box="[761,852,1126,1150]" name="Ecuador" pageId="2" pageNumber="599">Ecuador</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A28385204B325C6A13E23" box="[862,914,1126,1149]" name="Peru" pageId="2" pageNumber="599">Peru</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A283852C6B325C6D83E20" box="[924,1003,1126,1150]" name="Bolivia" pageId="2" pageNumber="599">Bolivia</collectingCountry>
(Monné 2018a).
</paragraph>
</subSubSection>
<paragraph id="3E130B19FF9A283851CDB3C9C7133E9B" blockId="2.[151,1405,1090,1222]" pageId="2" pageNumber="599">
<subSubSection id="76B65892FF9A283851CDB3C9C04E3EFC" box="[151,1405,1162,1186]" pageId="2" pageNumber="599" type="materials_examined">
Material examined:
<materialsCitation id="8EC40144FF9A28385034B3C9C04E3EFC" ID-GBIF-Occurrence="2239654646" box="[366,1405,1162,1186]" collectingDate="2017-04-28" collectionCode="CEO" country="Colombia" location="Universidad Nacional de Colombia-Sede Arauca" municipality="Arauca" pageId="2" pageNumber="599" specimenCount="1" stateProvince="Arauca">
<collectingCountry id="46BB4B89FF9A28385034B3C9C4E93EFC" box="[366,474,1162,1186]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
,
<collectingRegion id="FC68C5FBFF9A283850B9B3C9C7003EFC" box="[483,563,1162,1186]" country="Colombia" name="Arauca" pageId="2" pageNumber="599">Arauca</collectingRegion>
,
<collectingMunicipality id="DE779163FF9A28385366B3C9C7BF3EFC" box="[572,652,1162,1186]" pageId="2" pageNumber="599">Arauca</collectingMunicipality>
,
<location id="3B735DC2FF9A283853CCB3C9C1913EFC" LSID="urn:lsid:plazi:treatment:B605BA0FFF9A283851CDB301C7133E9B:3B735DC2FF9A283853CCB3C9C1913EFC" box="[662,1186,1162,1186]" country="Colombia" municipality="Arauca" name="Universidad Nacional de Colombia-Sede Arauca" pageId="2" pageNumber="599" stateProvince="Arauca">Universidad Nacional de Colombia-Sede Arauca</location>
.
<date id="4A122DD9FF9A283855F1B3C9C0173EFC" box="[1195,1316,1162,1186]" pageId="2" pageNumber="599" value="2017-04-28">
<collectingDate id="5A56D431FF9A283855F1B3C9C0173EFC" box="[1195,1316,1162,1186]" pageId="2" pageNumber="599" value="2017-04-28">28.IV.2017</collectingDate>
</date>
. (
<collectionCode id="58BD93DCFF9A2838546CB3C9C0423EFC" box="[1334,1393,1162,1186]" collectionName="CEO" pageId="2" pageNumber="599">CEO</collectionCode>
).
</materialsCitation>
</subSubSection>
<subSubSection id="76B65892FF9A283851CDB3EDC7133E9B" box="[151,544,1197,1222]" pageId="2" pageNumber="599" type="distribution">
<emphasis id="0CD8D70BFF9A283851CDB3EDC7133E9B" bold="true" box="[151,544,1197,1222]" pageId="2" pageNumber="599">
New country record for
<collectingCountry id="46BB4B89FF9A283850F0B3EEC7283E9B" box="[426,539,1197,1221]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
.
</emphasis>
</subSubSection>
</paragraph>
</treatment>
</document>

119
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Distribution:
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(Monné 2018a).
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Material examined:
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<collectingDate id="5A56D431FF9A283855A3B4F2C04B3997" box="[1273,1400,945,969]" pageId="2" pageNumber="599" value="2005-09-25">25/IX/2005</collectingDate>
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.
<collectorName id="93596ECFFF9A283854D8B4F1C43439B0" pageId="2" pageNumber="599">A. Vélez</collectorName>
. (
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New country record for
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.
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Distribution:
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,
<collectingCountry id="46BB4B89FF9A28385379B5D1C74438F7" box="[547,631,658,681]" name="Panama" pageId="2" pageNumber="599">Panama</collectingCountry>
(Monné 2018a).
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Material examined:
<materialsCitation id="8EC40144FF9A28385028B5F5C48E38AC" ID-GBIF-Occurrence="2239654647" collectingDate="2009-11-07" collectingMethod="Colecta manual" collectionCode="CIUQ" country="Colombia" elevation="100" location="Buenaventura" pageId="2" pageNumber="599" specimenCount="1" stateProvince="Valle del Cauca">
<collectingCountry id="46BB4B89FF9A28385028B5F5C4ED3890" box="[370,478,694,718]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
,
<collectingRegion id="FC68C5FBFF9A283850B0B5F5C7A43890" box="[490,663,694,718]" country="Colombia" name="Valle del Cauca" pageId="2" pageNumber="599">Valle del Cauca</collectingRegion>
,
<collectingCountry id="46BB4B89FF9A283853FEB5F5C6263890" box="[676,789,694,718]" name="Saint Pierre and Miquelon" pageId="2" pageNumber="599">San Pedro</collectingCountry>
,
<location id="3B735DC2FF9A28385278B5F5C6893893" LSID="urn:lsid:plazi:treatment:B605BA0FFF9A283851CDB52DC713394B:3B735DC2FF9A28385278B5F5C6893893" box="[802,954,694,717]" country="Colombia" name="Buenaventura" pageId="2" pageNumber="599" stateProvince="Valle del Cauca">Buenaventura</location>
.
<quantity id="F954A6FCFF9A2838529FB5F5C1233893" box="[965,1040,694,718]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="599" unit="m" value="100.0">
<elevation id="B581EC2AFF9A2838529FB5F5C1233893" box="[965,1040,694,718]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="599" unit="m" value="100.0">100 m</elevation>
</quantity>
.s.n.m. “
<collectingMethod id="E7ED730EFF9A2838553EB5F5C0273890" box="[1124,1300,694,718]" pageId="2" pageNumber="599">Colecta manual</collectingMethod>
”.
<date id="4A122DD9FF9A28385473B5F5C0AA3890" box="[1321,1433,694,718]" pageId="2" pageNumber="599" value="2009-11-07">
<collectingDate id="5A56D431FF9A28385473B5F5C0AA3890" box="[1321,1433,694,718]" pageId="2" pageNumber="599" value="2009-11-07">7.XI.2009</collectingDate>
</date>
.
<location id="3B735DC2FF9A2838519FB599C46F38AC" LSID="urn:lsid:plazi:treatment:B605BA0FFF9A283851CDB52DC713394B:3B735DC2FF9A2838519FB599C46F38AC" box="[197,348,730,754]" country="Colombia" name="Lopez G. Col." pageId="2" pageNumber="599" stateProvince="Valle del Cauca">López G. Col.</location>
(
<collectionCode id="58BD93DCFF9A28385030B599C48238AC" box="[362,433,730,754]" collectionName="CIUQ" pageId="2" pageNumber="599">CIUQ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
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<emphasis id="0CD8D70BFF9A283851CDB5BDC713394B" bold="true" box="[151,544,765,790]" pageId="2" pageNumber="599">
New country record for
<collectingCountry id="46BB4B89FF9A283850F0B5BEC728394B" box="[426,539,765,789]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

116
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<mods:title id="A8B2FD3FD1C5DEE7C10105E22A03EAD9">Description of the female of Anasillus crinitus Marinoni &amp; Martins 1978 (Coleoptera: Lamiinae: Acanthoderini) and new distribution records of longhorned beetles for Colombia</mods:title>
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<emphasis id="0CD8D70BFF9A283851CDB6D5C4533BF3" box="[151,352,406,430]" italics="true" pageId="2" pageNumber="599">Coleoxestia vittata</emphasis>
(Thomson, 1861)
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(
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)
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Distribution:
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,
<collectingCountry id="46BB4B89FF9A283850EEB6F9C72D3B8C" box="[436,542,442,466]" name="Honduras" pageId="2" pageNumber="599">Honduras</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A28385375B6F9C7AC3B8C" box="[559,671,442,466]" name="Nicaragua" pageId="2" pageNumber="599">Nicaragua</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A283853EAB6F9C61D3B8C" box="[688,814,442,466]" name="Costa Rica" pageId="2" pageNumber="599">Costa Rica</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A28385264B6F9C6A63B8F" box="[830,917,442,465]" name="Panama" pageId="2" pageNumber="599">Panama</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A283852FCB6F9C6D93B8C" box="[934,1002,442,466]" name="Brazil" pageId="2" pageNumber="599">Brazil</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A283852A1B6F9C17F3B8C" box="[1019,1100,442,466]" name="Bolivia" pageId="2" pageNumber="599">Bolivia</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A28385507B6F9C1F23B8C" box="[1117,1217,442,466]" name="Paraguay" pageId="2" pageNumber="599">Paraguay</collectingCountry>
,
<collectingCountry id="46BB4B89FF9A2838558BB6F9C0083B8C" box="[1233,1339,442,466]" name="Argentina" pageId="2" pageNumber="599">Argentina</collectingCountry>
(Monné 2018a)
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Material examined:
<materialsCitation id="8EC40144FF9A28385034B541C10F3844" ID-GBIF-Occurrence="2239654650" box="[366,1084,513,538]" collectingDate="1991-10-05" collectionCode="MENT-UT" collectorName="A. O. Col" country="Colombia" location="Bajo Calima" pageId="2" pageNumber="599" specimenCount="1" stateProvince="Tolima">
<collectingCountry id="46BB4B89FF9A28385034B541C4E93844" box="[366,474,514,538]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
,
<collectingRegion id="FC68C5FBFF9A283850BEB541C7013844" box="[484,562,514,538]" country="Colombia" name="Tolima" pageId="2" pageNumber="599">Tolima</collectingRegion>
,
<location id="3B735DC2FF9A28385366B541C7F53844" LSID="urn:lsid:plazi:treatment:B605BA0FFF9A283851CDB6D5C7133863:3B735DC2FF9A28385366B541C7F53844" box="[572,710,514,538]" country="Colombia" name="Bajo Calima" pageId="2" pageNumber="599" stateProvince="Tolima">Bajo Calima</location>
.
<date id="4A122DD9FF9A2838538AB542C6053847" box="[720,822,513,537]" pageId="2" pageNumber="599" value="1991-10-05">
<collectingDate id="5A56D431FF9A2838538AB542C6053847" box="[720,822,513,537]" pageId="2" pageNumber="599" value="1991-10-05">5.X.1991</collectingDate>
</date>
.
<collectorName id="93596ECFFF9A28385265B541C6903844" box="[831,931,514,538]" pageId="2" pageNumber="599">A.O. Col</collectorName>
. (
<collectionCode id="58BD93DCFF9A283852EEB541C1033844" box="[948,1072,514,538]" pageId="2" pageNumber="599">MENT-UT</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="76B65892FF9A283851CDB565C7133863" box="[151,544,549,574]" pageId="2" pageNumber="599" type="distribution">
<paragraph id="3E130B19FF9A283851CDB565C7133863" blockId="2.[151,1436,296,574]" box="[151,544,549,574]" pageId="2" pageNumber="599">
<emphasis id="0CD8D70BFF9A283851CDB565C7133863" bold="true" box="[151,544,549,574]" pageId="2" pageNumber="599">
New country record for
<collectingCountry id="46BB4B89FF9A283850F0B566C7283863" box="[426,539,549,573]" name="Colombia" pageId="2" pageNumber="599">Colombia</collectingCountry>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

122
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Linnaeus
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,
<volumeTitle id="0BDCC486CA28ECF11B5742354D2DBF90">
<emphasis id="EB01512B9088A9698B1CA506CFA210AE" italics="true" pageNumber="423">Systema Naturae</emphasis>
, ed. 12, 2
</volumeTitle>
: 508. 1767
</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0842B4D21E8EB5D310655BEA712BDD57" pageNumber="423" type="multiple">
<paragraph id="7DB36CE13C1FD75779ABB1B20431522F" pageNumber="423">RCN: 5728.</paragraph>
</subSubSection>
<subSubSection id="F4752216BE636B433345EF8E15B03243" pageNumber="423" type="reference_group">
<paragraph id="315276FAB1BC1CB4E053E074160E5644" pageNumber="423">
Replaced synonym:
<taxonomicName id="AFBD25BFD865BCCD8D09EEE6BBA764AC" authority="L. (1763)" authorityName="Linnaeus" authorityYear="1763" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="variety" species="aurantium" variety="decumana">
<emphasis id="8367061B625C6FB2F27CA55E83B4FF4E" italics="true" pageNumber="423">Citrus aurantium</emphasis>
L. var.
<emphasis id="58A35B288AE7BC384597253747C7362D" italics="true" pageNumber="423">decumana</emphasis>
L. (1763)
</taxonomicName>
,
<emphasis id="C6693964C9E5D7ABC2E3C9B86FE98561" italics="true" pageNumber="423">nom. illeg.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="712C895714F524A82F9975832A25295F" pageNumber="423" type="materials_examined">
<paragraph id="21DAF6B0FF24224132DF033EAA5B07BA" pageNumber="423">
<typeStatus id="AAB28C6A2EB1E7E3A7CF8B9DCFE476E3">Lectotype</typeStatus>
(Mabberley in
<emphasis id="84549EA44DC3DCFB4F86288B6AE35B47" italics="true" pageNumber="423">Telopea</emphasis>
7: 170. 1997): [icon] &quot;
<emphasis id="FE6A2E6C85A4D539060C4B020C320857" italics="true" pageNumber="423">Malus arantia, fructu rotundo maximo pallescente humanum caput excedente</emphasis>
&quot; in Sloane, Voy. Jamaica 1: 41, t. 12, f. 2, 3. 1707. -
<materialsCitation id="99147F5FEB962FE0AF79C804F61CDF90" ID-GBIF-Occurrence="2612072693" collectionCode="BM-SL" specimenCode="115" typeStatus="typotype">
<typeStatus id="ECA2EA514225094A79AB8A331AAB9B61">Typotype</typeStatus>
: Herb. Sloane 7: 115 (
<collectionCode id="D3D0CF5DF7C5DB3BB2878BF64C609902">BM-SL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4B017884D09E07D593C2B0ADF1BCF9A0" pageNumber="423" type="nomenclature">
<paragraph id="895DD6D9D9B1C717CC2878979C59BB77" pageNumber="423">
Current name:
<taxonomicName id="429F537AB1C8B0AD6DEA87562ADF6494" ID-CoL="VMLB" authority="(Burm.) Merr." authorityName="Merr." baseAuthorityName="Burm." class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="species" species="maxima">
<emphasis id="BCC1A7220448FB5759CEF366ACEAC862" italics="true" pageNumber="423">
<emphasis id="4CD1CDE2B14E33EE4C4547266C0A9F36" bold="true" italics="true" pageNumber="423">Citrus maxima</emphasis>
</emphasis>
(Burm.) Merr.
</taxonomicName>
(
<taxonomicName id="6EEDE7A9A3BE289EF895A4BD2A1EA580" class="Magnoliopsida" family="Rutaceae" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="family">Rutaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="F37173F61B29AA423904C4B4ADFEC83F" pageNumber="423" type="discussion">
<paragraph id="A90A21A6AFD3FA155D3DF86AEC3B642E" pageNumber="423">
<emphasis id="8C6CF187B698DB6C1F084AF97827BCB9" italics="true" pageNumber="423">Note:</emphasis>
Specific epithet spelled
<normalizedToken id="2A83BAC484AA0F91D2232CBE957ADA23" originalValue="“decumanus”">&quot;decumanus&quot;</normalizedToken>
in the protologue. A
<emphasis id="77C2D4BF7D339BA20469FAA8BCF6943D" italics="true" pageNumber="423">nomen novum</emphasis>
for the illegitimate
<taxonomicName id="57675B43BF350B7A786BC777D671FBC0" authority="L. (1763)" authorityName="Linnaeus" authorityYear="1763" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" phylum="Tracheophyta" rank="variety" species="aurantium" variety="decumana">
<emphasis id="F03EC9683E576C8DB7055045A0CF9EC8" italics="true" pageNumber="423">Citrus aurantium</emphasis>
var.
<emphasis id="627E732F71DDE1C1F3A136973788A452" italics="true" pageNumber="423">decumana</emphasis>
L. (1763)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

308
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@ -0,0 +1,308 @@
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<mods:title id="2B9BFB326350518507E1346BBC3385C7">A new lithophilous species of Gesneriaceae, Petrocodon rubrostriatus, from the karst area of South Yunnan, China</mods:title>
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<mods:nameIdentifier id="4EFEAB17B4FD4A2DCAAE2B424485E3FA" type="ORCID">https://orcid.org/0000-0002-4707-2656</mods:nameIdentifier>
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<paragraph id="E600C86663BFE7322DCF25486C4031A3" pageId="0" pageNumber="289">
<taxonomicName id="69D0822DE84803150CEDCDA73DFF8C40" LSID="B60679A4-07DF-5610-BF9E-8AA0C736B5E5" authority="K. Tan, X. Q. Song &amp; M. X. Ren" authorityName="K. Tan, X. Q. Song &amp; M. X. Ren" authorityYear="2023" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon rubrostriatus" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="species" species="rubrostriatus" status="sp. nov.">Petrocodon rubrostriatus K.Tan, X.Q.Song &amp; M.X.Ren</taxonomicName>
<taxonomicNameLabel id="68FE06251DA22D632444D0F542F3E0CE" pageId="0" pageNumber="289">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6CE1D6B313F98FAFEFE2820C607CFC4C" pageId="0" pageNumber="289" type="description">
<paragraph id="3E2DBE812CAEEF5D37662007A711AEF2" pageId="0" pageNumber="289">
<figureCitation id="6D5FC660694E6BC6CF65BB20DD95D494" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Petrocodon rubrostriatus K. Tan, X. Q. Song &amp; M. X. Ren, sp. nov. A habitat B habit C underside of plant D oblique side view of flower and buds E cyme F bracts and flower bud G dissected calyx lobes, pistil with opened corolla H front view of flower I side view of flower J stamens with coherent anthers K dehisced capsule (Photographs by D. C. Meng)." figureDoi="10.3897/phytokeys.230.106358.figure2" httpUri="https://binary.pensoft.net/fig/894189" pageId="0" pageNumber="289">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="62E245A5174CEEB022CD2F45F8DAF31E" pageId="0" pageNumber="289" type="diagnosis">
<paragraph id="6A0E86CEC379E9FDA6D2B63603A93672" pageId="0" pageNumber="289">Diagnosis.</paragraph>
<paragraph id="41BE250966BF46FBF7F918EAE881DB3D" pageId="0" pageNumber="289">
The new species resembles
<taxonomicName id="D093C74BDEB380F172C5688B720EE149" authorityName="A.Weber &amp; Mich.Moller" authorityYear="2011" baseAuthorityName="W.T.Wang" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon mollifolius" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="species" species="mollifolius">
<emphasis id="7F4A2CCA527C9E5ABB48B819DE0C1A97" italics="true" pageId="0" pageNumber="289">Petrocodon mollifolius</emphasis>
</taxonomicName>
(W.T.Wang) A.Weber &amp; Mich.
<normalizedToken id="1435F5E1CFB5B64CCE1E34E310C14CFE" originalValue="Möller">Moeller</normalizedToken>
in leaf blade shape and size, flower base colour and size (Figs
<figureCitation id="A95DD2A805BED33C8A228C6D6477FF08" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Petrocodon rubrostriatus K. Tan, X. Q. Song &amp; M. X. Ren, sp. nov. A habitat B habit C underside of plant D oblique side view of flower and buds E cyme F bracts and flower bud G dissected calyx lobes, pistil with opened corolla H front view of flower I side view of flower J stamens with coherent anthers K dehisced capsule (Photographs by D. C. Meng)." figureDoi="10.3897/phytokeys.230.106358.figure2" httpUri="https://binary.pensoft.net/fig/894189" pageId="0" pageNumber="289">2</figureCitation>
,
<figureCitation id="B3C59B5B45A000A5DFCAA34630658AE0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Petrocodon mollifolius (W. T. Wang) A. Weber &amp; Mich. Moeller A adaxial view of flower B opened corolla with two fertile stamens and three staminodes C pistil with disc D dissected calyx lobes E stamens with coherent anthers (Photographs by D. C. Meng)." figureDoi="10.3897/phytokeys.230.106358.figure3" httpUri="https://binary.pensoft.net/fig/894190" pageId="0" pageNumber="289">3</figureCitation>
), but can be easily distinguished from the latter by corolla lobes with red to brownish longitudinal stripes (
<emphasis id="7E3F1B91FDFF17A650689751A714B264" italics="true" pageId="0" pageNumber="289">vs.</emphasis>
corolla purely yellow), shorter lanceolate bracts, ca. 4.5 mm long (
<emphasis id="430D3B0D58F1AB783A356D94A28E25A4" italics="true" pageId="0" pageNumber="289">vs.</emphasis>
linear, 12-20 mm long); longer style length ca. 10 mm (
<emphasis id="F19D7913AF3656A7B23F96856B3F9840" italics="true" pageId="0" pageNumber="289">vs.</emphasis>
6-8 mm), and staminode insertion at 2.5-3 mm from corolla base (
<emphasis id="646DEDCA47257ED12B1942871C46D327" italics="true" pageId="0" pageNumber="289">vs.</emphasis>
inserted at the corolla base).
</paragraph>
<caption id="3A994E280F3D5F6A82ADCC898B07831E" doi="10.3897/phytokeys.230.106358.figure3" httpUri="https://binary.pensoft.net/fig/894190" pageId="0" pageNumber="289" start="Figure 3" startId="F3">
<paragraph id="DECD07CF9557C5CE49154475D8EBFFD1" pageId="0" pageNumber="289">
<emphasis id="65277523C8F692E1970070415F4D924A" bold="true" pageId="0" pageNumber="289">Figure 3.</emphasis>
<taxonomicName id="D125D6A1F5781E60FDB716E9A96C6902" authorityName="A.Weber &amp; Mich.Moller" authorityYear="2011" baseAuthorityName="W.T.Wang" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon mollifolius" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="species" species="mollifolius">
<emphasis id="AA01A2E428F987D27791263519B2C8A3" italics="true" pageId="0" pageNumber="289">Petrocodon mollifolius</emphasis>
</taxonomicName>
(W.T.Wang) A.Weber &amp; Mich.
<normalizedToken id="F5F6E1BA24B05979486E9943CA304CE6" originalValue="Möller">Moeller</normalizedToken>
<emphasis id="FD40D64E5B4B4F2E9E22D834CBB87C10" bold="true" pageId="0" pageNumber="289">A</emphasis>
adaxial view of flower
<emphasis id="35D79556275390B24088ABF8C17B78E5" bold="true" pageId="0" pageNumber="289">B</emphasis>
opened corolla with two fertile stamens and three staminodes
<emphasis id="DC1FD2B721DBF10C89F026CABDC3168E" bold="true" pageId="0" pageNumber="289">C</emphasis>
pistil with disc
<emphasis id="9238A41677EFEB783940EDD169B63689" bold="true" pageId="0" pageNumber="289">D</emphasis>
dissected calyx lobes
<emphasis id="770303064DA02C67FF35F2839CF03852" bold="true" pageId="0" pageNumber="289">E</emphasis>
stamens with coherent anthers (Photographs by D.C. Meng).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="A746340309A8AAF3561C327EC16E70B2" pageId="0" pageNumber="289" type="materials_examined">
<paragraph id="D4B96C69B5AC9F32F3269BDFB6965687" pageId="0" pageNumber="289">
<typeStatus id="277E4E579D14CE5B6BFEE9266BB6C12B">Type</typeStatus>
.
</paragraph>
<paragraph id="5944C0EF4B724FB71254F5B2F768727D" pageId="0" pageNumber="289">
<materialsCitation id="D7293F4C9BAD1DEC8E4302B8C155F60A" collectingDate="2020-08-13" country="China" county="Lvchun county" elevation="390" latitude="22.6" location="Emaluoba" longLatPrecision="1259" longitude="102.26667" municipality="Banpo township" specimenCount="1" stateProvince="Yunnan Province" typeStatus="Holotype">
<collectingCountry id="B9608987F8065D192F0272AD39D665CF" name="China">China</collectingCountry>
.
<collectingRegion id="ED35847E009CE10490E2043BCE9CF532" country="China" name="Yunnan">Yunnan Province</collectingRegion>
:
<collectingCounty id="F086A9C3BBD25D3C1C42BBC4A9797AAA">Lvchun county</collectingCounty>
,
<collectingMunicipality id="4B5F93B47FA8CBAA05EDC7F71BC0CB84">Banpo township</collectingMunicipality>
,
<location id="86B5E606EA27655DADAF786ECE07CFDD" LSID="urn:lsid:plazi:treatment:B60679A407DF5610BF9E8AA0C736B5E5:86B5E606EA27655DADAF786ECE07CFDD" country="China" county="Lvchun county" latitude="22.6" longLatPrecision="1259" longitude="102.26667" municipality="Banpo township" name="Emaluoba" stateProvince="Yunnan Province">Emaluoba</location>
community,
<geoCoordinate id="D9DB58A9D2FE4D90834D9B20EEA6CFED" degrees="22" direction="north" minutes="36" orientation="latitude" precision="925" value="22.6">22°36'N</geoCoordinate>
,
<geoCoordinate id="0A66862FBAD35DF90D268C1E0E41727C" degrees="102" direction="east" minutes="16" orientation="longitude" precision="925" value="102.26667">102°16'E</geoCoordinate>
, altitude ca.
<quantity id="B9D854F1A0B9A91EA7C1FDF2628E3D28" metricMagnitude="2" metricUnit="m" metricValue="3.9" unit="m" value="390.0">
<elevation id="92F2DF255909D79A06D8C75CDB73B52B" metricMagnitude="2" metricUnit="m" metricValue="3.9" unit="m" value="390.0">390 m</elevation>
</quantity>
,
<collectingDate id="5E9DBB8E20E5FE323EFA87B4717EE0AD" value="2020-08-13">August 13, 2020</collectingDate>
,
<emphasis id="84AC34C03C26903A670E8C4483C2CD20" italics="true" pageId="0" pageNumber="289">Ke Tan 20TK0811008</emphasis>
(
<typeStatus id="4E5851E1C33242AD8D1AA122BBDE1B0E">Holotype</typeStatus>
: IBK! IBK00449896; Isotypes: HUTB, IBK! IBK00449897)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="09D7529B8090B0CFCFB21EAA2F7DF44A" pageId="0" pageNumber="289" type="description">
<paragraph id="1CB885E2820408AD6287F81686845D21" pageId="0" pageNumber="289">Description.</paragraph>
<paragraph id="B592A02ADD265E45EB560C6E42EC1050" pageId="0" pageNumber="289">
Perennial herb.
<emphasis id="4A2923EB9F83DF9D0EA6C86444DA4C94" bold="true" italics="true" pageId="0" pageNumber="289">Rhizomatous stem</emphasis>
1-2 cm long, 5-8 mm in diam., sometimes inconspicuous.
<emphasis id="FF3A0358427918616C0AE77491D45B56" bold="true" italics="true" pageId="0" pageNumber="289">Roots</emphasis>
fibrous, numerous, pale brown to brown.
<emphasis id="B5CB596EEA968C3C868876AC2FBAF656" bold="true" italics="true" pageId="0" pageNumber="289">Leaves</emphasis>
in a densely crowded basal rosette, (6-)9-12; petioles 1.5-4 cm long, densely whitish villous; lamina adaxial surface dark green, abaxial surface green, both surfaces whitish pubescent, herbaceous, elliptic, broadly elliptic to ovate, or rhombic ovate, 5.5-11.5
<normalizedToken id="9D3CFE32A48F7E6315AE9B8E457A35E6" originalValue="×">x</normalizedToken>
2.3-3.5 cm, margin serrate; 4-5 pairs of lateral veins on each side, ascending, tertiary venation also distinctive; apex acute, occasionally obtuse, base cuneate, occasionally asymmetric.
<emphasis id="4CC059486432C352E81D1A0976FEE6D8" bold="true" italics="true" pageId="0" pageNumber="289">Inflorescences</emphasis>
1-4 per plant, axillary, cymose, 3-5-flowered; peduncles green to pale brownish green, puberulent, interspersed with a few longer hairs, 6.5-8.5 cm long; bracts 2, opposite, green to pale green, lanceolate, ca. 4.5 mm long, ca. 1 mm across at base, abaxially pubescent, adaxially nearly glabrous, margin entire; bracteoles 2, opposite, narrowly triangular, colour and hairs same as bracts; pedicels 8-12 mm long, ca. 1 mm in diam., hairs same as peduncle. Calyx 5-sect to base, lobes green to pale green, lanceolate, ca. 2.5
<normalizedToken id="E52BC812B3882949B67F3BEB6B05B011" originalValue="×">x</normalizedToken>
0.5 mm, abaxially pubescent, adaxially nearly glabrous, margin entire.
<emphasis id="399582FCD4702657EF5853FFCFBAA2C3" bold="true" italics="true" pageId="0" pageNumber="289">Corolla</emphasis>
pale yellow to bright yellow, 1.9-2.2 cm long, base near-spherical, outside pubescent interspersed with few glandular hairs, inside glabrous; tube infundibuliform, tubular at the base and widening around the middle, ca. 15 mm long, ca. 4 mm in diam. at the base, ca. 7.5 mm in diam. at orifice, abaxial lip much longer than adaxial lip, adaxial lip 2-lobed to near base, slightly obliquely semicircular, lobes ca. 8 mm long, ca. 6 mm wide at base, margin entire, apex rounded, each lobe with two to three red to brownish-red longitudinal stripes, abaxial lip 3-lobed to more than middle, elliptical, lateral ones slightly oblique and smaller than the central one, lobes ca. 13 mm long, ca. 5 mm wide at base, margin entire, apex rounded, each lobe with one red to brownish-red longitudinal stripe.
<emphasis id="832155F253B74208CE35D2B7CAC086EF" bold="true" italics="true" pageId="0" pageNumber="289">Stamens</emphasis>
2, inserted in tube ca. 6 mm from corolla base; filaments pale green, ca. 8 mm long, straight, sparsely eglandular and glandular puberulent from the middle to the top and with glands from the middle to the bottom; anthers pale brownish yellow, elliptic, ca. 1.5
<normalizedToken id="EC6F80A48357871C0A4747F6C3DF5B2A" originalValue="×">x</normalizedToken>
2.2 mm, glabrous, cohering face to face; staminodes 3, inserted 2.5-3 mm from corolla base, ca. 2 mm long, glabrous.
<emphasis id="3A917754537A1562366ACFB7E746847B" bold="true" italics="true" pageId="0" pageNumber="289">Disc</emphasis>
brownish green, ca. 0.9 mm high, glabrous, margin sinuate.
<emphasis id="67979DA64C55A3A4F82DE2E852230A22" bold="true" italics="true" pageId="0" pageNumber="289">Pistil</emphasis>
ca. 21 mm long; ovary pale green, densely puberulent, ca. 10 mm long, ca. 1 mm wide; style whitish green, ca. 10 mm long, sparsely eglandular pubescent; stigma pale green, ca. 1 mm long, ca. 0.7 mm wide, disc-like.
<emphasis id="9CCFD285718843111868EA062B983298" bold="true" italics="true" pageId="0" pageNumber="289">Capsule</emphasis>
cylindrical, green when young, 1.7-2.1 cm long, ca. 1.5 mm wide, pubescent, becoming grey-brown and dehiscing into four valves when mature.
<emphasis id="A951DB8366B361752369E57020BC80E9" bold="true" italics="true" pageId="0" pageNumber="289">Seeds</emphasis>
unappendaged, long ellipsoid, ca. 0.4 mm long, ca. 0.2 mm wide.
</paragraph>
</subSubSection>
<subSubSection id="1C7BDB2F34FAB770C0FDEB5C656EB3DD" pageId="0" pageNumber="289" type="phenology">
<paragraph id="BA81871FED7C25F5F89BAC1B8D042F2B" pageId="0" pageNumber="289">Phenology.</paragraph>
<paragraph id="90F47B3AFC6D2387CF0592715BE4795F" pageId="0" pageNumber="289">Flowering in August, fruiting from October to November based on field observations.</paragraph>
</subSubSection>
<subSubSection id="149AA3CB1030ACF5D84C725776B8B1F6" pageId="0" pageNumber="289" type="etymology">
<paragraph id="F3DC30C51ED2A5F2877F74C476C70AE9" pageId="0" pageNumber="289">Etymology.</paragraph>
<paragraph id="278F2BEEF79547E9BB667D4F88EE341B" pageId="0" pageNumber="289">
The name
<taxonomicName id="68122323B2EDF27761F836A7805EB6D3" authorityName="K. Tan, X. Q. Song &amp; M. X. Ren" authorityYear="2023" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon rubrostriatus" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="species" species="rubrostriatus">
<emphasis id="3D2F6BD0E0B3363876FDAC3822EEFBF3" italics="true" pageId="0" pageNumber="289">Petrocodon rubrostriatus</emphasis>
</taxonomicName>
refers to the bright red to brownish stripes in the yellow corolla. This is noticeably different from the corolla colours of previously published
<taxonomicName id="A3A14ADF7E5DF3471B3DD5A4B1A62ECB" authorityName="Hance" authorityYear="1883" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="genus">
<emphasis id="EC3D6B5EEDDA30C7AC5805261C68C91B" italics="true" pageId="0" pageNumber="289">Petrocodon</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="3FB8F3EF1C08B2803DF546C1866AA878" pageId="0" pageNumber="289" type="vernacular name">
<paragraph id="D2735BD967FF4CEA8A4E1E8A28505806" pageId="0" pageNumber="289">Vernacular name.</paragraph>
<paragraph id="171E4D63F0DABED25A071D4ECD892ED3" pageId="0" pageNumber="289">
<normalizedToken id="698FD028F10D48B72F51B576B250A492" originalValue="Hóng">Hong</normalizedToken>
<normalizedToken id="D5883679F5494C02E6B7F78747044871" originalValue="Wén">Wen</normalizedToken>
<normalizedToken id="02778CD5FA9822C8646B7F422A53F922" originalValue="Shí">Shi</normalizedToken>
<normalizedToken id="E35A7A214F2B6F605726DB53C6525409" originalValue="Shān">Shan</normalizedToken>
<normalizedToken id="2135E11DB02F3F6CFF04341AFBBA3F42" originalValue="Jù">Ju</normalizedToken>
<normalizedToken id="9922FE479E64854A5D1C343630D007EB" originalValue="Tái">Tai</normalizedToken>
(红纹石山苣苔). The first two words,
<normalizedToken id="061D217CC4DB9107E15FC7A876CF1CE1" originalValue="“Hóng">&quot;Hong</normalizedToken>
<normalizedToken id="E889AD8C3BD9F1E67F7B7DDDD78B0E41" originalValue="Wén”">Wen&quot;</normalizedToken>
, mean red stripes of the corolla, and the following four words,
<normalizedToken id="34C74E4ADB5A79C4BEA2C5539A488E02" originalValue="“Shí">&quot;Shi</normalizedToken>
<normalizedToken id="715F23DD21AD34002B69A9B174048A6E" originalValue="Shān">Shan</normalizedToken>
<normalizedToken id="B483F8466C2201A18E1A377624A31F66" originalValue="Jù">Ju</normalizedToken>
<normalizedToken id="6006727260CA34DD6B352AA4F7C1079C" originalValue="Tái”">Tai&quot;</normalizedToken>
, mean
<taxonomicName id="2332DEE9CD4EBA350AEC9D84850D60CE" authorityName="Hance" authorityYear="1883" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="genus">
<emphasis id="3DCCB54CA0864A512CF0D1158AE68AE1" italics="true" pageId="0" pageNumber="289">Petrocodon</emphasis>
</taxonomicName>
in Chinese.
</paragraph>
</subSubSection>
<subSubSection id="99EC1A86284C105AE2B21358CA423362" pageId="0" pageNumber="289" type="distribution">
<paragraph id="20BB348A98590967612666E2A7B9633C" pageId="0" pageNumber="289">Distribution and habitat.</paragraph>
<paragraph id="54B171714413325ECF3B5D26EEA0A9F8" pageId="0" pageNumber="289">
<taxonomicName id="C2C89C96A80068C59275C04CDAC8C560" authorityName="K. Tan, X. Q. Song &amp; M. X. Ren" authorityYear="2023" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon rubrostriatus" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="species" species="rubrostriatus">
<emphasis id="C6608621E723E16BA9F9EB0046F168A9" italics="true" pageId="0" pageNumber="289">Petrocodon rubrostriatus</emphasis>
</taxonomicName>
is only known from the type locality, near Lixianjiang river, Emaluoba community, Banpo township, Lvchun county, Yunnan. The species grows on moist, shady tufa surfaces of a limestone cliff in a monsoon rainforest at ca. 400 m. Thus, it is exposed to a warm environment with high air humidity in a moderately shaded monsoon rainforest.
</paragraph>
</subSubSection>
<subSubSection id="4C18AFDDA296BB9B09A1FF6282E6EC9C" pageId="0" pageNumber="289" type="preliminary conservation assessment">
<paragraph id="CC8D9E531386797186CD33ADBD488017" pageId="0" pageNumber="289">Preliminary conservation assessment.</paragraph>
<paragraph id="6D9461745C9EABA3498A6CF05339F696" pageId="0" pageNumber="289">
<taxonomicName id="810B92E5D0280124C320039B8B357B2E" authorityName="K. Tan, X. Q. Song &amp; M. X. Ren" authorityYear="2023" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon rubrostriatus" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="species" species="rubrostriatus">
<emphasis id="C2B240D972B5672C473029387CB669AF" italics="true" pageId="0" pageNumber="289">Petrocodon rubrostriatus</emphasis>
</taxonomicName>
is currently only known from the type locality in the Lixianjiang river basin, where only one small population was observed. In total in 2020, there were fewer than 300 mature individuals in five separate subpopulations, clustered together in a fairly small site of ca. 100 m2, on a moist cliff on the rock surface of an unnamed limestone hill close to the Lixianjiang river. The area of occupancy (AOO), is significantly smaller than the smallest AOO unit of IUCN (10 km2 for Critically Endangered under B2). In 2022, we revisited the type locality and observed a decline in habitat quality caused by a prolonged drought in Southwest China, and a reduction in mature plants to only about 100. This suggests that the new species is extremely vulnerable and easily disturbed by the persistent drought and also the activities of local people. According to the IUCN red list criteria (
<bibRefCitation id="66773E2FE11B90521D85186A570FC978" author="Huang, H" journalOrPublisher="Guihaia" pageId="0" pageNumber="289" publicationUrl="https://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B7" refString="2022. . https://www.iucnredlist.org/documents/RedListGuidelines.pdf" url="https://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2022">IUCN 2022</bibRefCitation>
), the category of 'Critically
<normalizedToken id="FF921F7553407655582AD2EA91E865CF" originalValue="Endangered’">Endangered'</normalizedToken>
[CR, B2a,b (iii,iv,v)] is proposed here.
</paragraph>
</subSubSection>
<subSubSection id="4122F6C4AACD6C90392C1DF6DF796C97" pageId="0" pageNumber="289" type="taxonomic notes">
<paragraph id="BAABB139F19D333F4BFD766962D10378" pageId="0" pageNumber="289">Taxonomic notes.</paragraph>
<paragraph id="BECA5C5C47A73BDB3E19DD9188052598" pageId="0" pageNumber="289">
The new species is morphologically similar to
<taxonomicName id="F69880649FDBED48B8A969FB6F189CF4" authorityName="A.Weber &amp; Mich.Moller" authorityYear="2011" baseAuthorityName="W.T.Wang" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon mollifolius" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="species" species="mollifolius">
<emphasis id="445CF4016BC53C931366424658FB3C71" italics="true" pageId="0" pageNumber="289">Petrocodon mollifolius</emphasis>
</taxonomicName>
, but most easily distinguished by the longitudinal stripes of red to brownish-red on the petal lobes. With the new species here, there are now 50 species and one variety in
<taxonomicName id="4B8845D784A3D62778D7C6524E82B1FB" authorityName="Hance" authorityYear="1883" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="genus">
<emphasis id="081904EFD6C894912153A12F7515C882" italics="true" pageId="0" pageNumber="289">Petrocodon</emphasis>
</taxonomicName>
, and 47 species are distributed in China. South and Southwest China are the distribution and diversity centres of
<taxonomicName id="CBF512794F31847A644C327C28319919" authorityName="Hance" authorityYear="1883" class="Magnoliopsida" family="Gesneriaceae" genus="Petrocodon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Petrocodon" order="Lamiales" pageId="0" pageNumber="289" phylum="Tracheophyta" rank="genus">
<emphasis id="72DE9FEA1CF2190F1E7F16A919087A46" italics="true" pageId="0" pageNumber="289">Petrocodon</emphasis>
</taxonomicName>
. The distribution pattern of a few species extends to the northern Indo-China Peninsula, namely
<taxonomicName id="79ACCDDBDC9DAEB05F7C0F8391F977CE" lsidName="P. coccineus" pageId="0" pageNumber="289" rank="species" species="coccineus">
<emphasis id="9DDA12121DD31E3EDC1CE690C24849FD" italics="true" pageId="0" pageNumber="289">P. coccineus</emphasis>
</taxonomicName>
(C.Y.Wu ex H.W.Li) Yin Z.Wang and
<taxonomicName id="A8915530F2717BF90756F52F5975539C" lsidName="P. hispidus" pageId="0" pageNumber="289" rank="species" species="hispidus">
<emphasis id="8FF45EAEAC5CF67A584CF15A2A1F2500" italics="true" pageId="0" pageNumber="289">P. hispidus</emphasis>
</taxonomicName>
(W.T.Wang) A.Weber &amp; Mich.
<normalizedToken id="570CA313184DEDB851806B91014260E7" originalValue="Möller">Moeller</normalizedToken>
which are both distributed in South China and Northern Vietnam (
<bibRefCitation id="EEF25DE170352C7067C00F7D2B12F00A" author="Wei, YG" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="0" pageNumber="289" refId="B23" refString="Wei, YG, 2018. The distribution and conservation status of native plants in Guangxi, China. China Forestry Publishing House, Beijing" title="The distribution and conservation status of native plants in Guangxi, China." year="2018">Wei 2018</bibRefCitation>
;
<bibRefCitation id="82028889BFF665024083EA8BA5DEF73E" author="Wei, YG" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="0" pageNumber="289" refId="B24" refString="Wei, YG, Do, VT, Wen, F, 2022. A Checklist to the Plants of Northern Vietnam. China Forestry Publishing House, Beijing" title="A Checklist to the Plants of Northern Vietnam." year="2022">Wei et al. 2022</bibRefCitation>
),
<taxonomicName id="F9E429E7F42DE0F6E8FC2AD05B780950" lsidName="P. vietnamensis" pageId="0" pageNumber="289" rank="species" species="vietnamensis">
<emphasis id="5679117831F80817ED56B38D69D2624A" italics="true" pageId="0" pageNumber="289">P. vietnamensis</emphasis>
</taxonomicName>
Z.B.Xin, T.V.Do &amp; F.Wen is endemic of Vietnam (
<bibRefCitation id="A3DA3DA584DE9032F3AAD8D1BC0A2B17" DOI="https://doi.org/10.1111/njb.02996" author="Xin, ZB" journalOrPublisher="Taiwania" pageId="0" pageNumber="289" refId="B26" refString="Xin, ZB, Do, TV, Fu, LF, Huang, ZJ, Maciejewski, S, Wen, F, 2021. Petrocodon vietnamensis sp. nov. (Gesneriaceae) from karst caves in northern Vietnam. Nordic Journal of Botany 39(4): njb.02996. https://doi.org/10.1111/njb.02996" title="Petrocodon vietnamensis sp. nov. (Gesneriaceae) from karst caves in northern Vietnam. Nordic Journal of Botany 39 (4): njb. 02996." url="https://doi.org/10.1111/njb.02996" year="2021">Xin et al. 2021</bibRefCitation>
;
<bibRefCitation id="110CB2C1BC972013052D2BC8D037E77C" author="Wei, YG" journalOrPublisher="China Forestry Publishing House, Beijing" pageId="0" pageNumber="289" refId="B24" refString="Wei, YG, Do, VT, Wen, F, 2022. A Checklist to the Plants of Northern Vietnam. China Forestry Publishing House, Beijing" title="A Checklist to the Plants of Northern Vietnam." year="2022">Wei et al. 2022</bibRefCitation>
),
<taxonomicName id="2DC84A29B29CEFC66A2C9203C7D42559" lsidName="P. bonii" pageId="0" pageNumber="289" rank="species" species="bonii">
<emphasis id="0D171A89EBA92ADA6697873ECADF1CCE" italics="true" pageId="0" pageNumber="289">P. bonii</emphasis>
</taxonomicName>
(Pellegr.) A.Weber &amp; Mich.
<normalizedToken id="54A092406880382DEC5AC48897EF6E76" originalValue="Möller">Moeller</normalizedToken>
is distributed from Thailand to Vietnam (GRC 2023,
<bibRefCitation id="FC5B1813C15579E5B874DA9CA7B74604" author="Nguyen, LT" journalOrPublisher="Molecular Biology and Evolution" pageId="0" pageNumber="289" publicationUrl="http://www.plantsoftheworldonline.org/" refId="B15" refString="2023. . http://www.plantsoftheworldonline.org/" url="http://www.plantsoftheworldonline.org/" year="2023">POWO 2023</bibRefCitation>
), and
<taxonomicName id="1237A849D8BC63567886C7CB418E6877" lsidName="P. flavus" pageId="0" pageNumber="289" rank="species" species="flavus">
<emphasis id="BC8FB2DF8580A3EF7510BB3C622A3EAD" italics="true" pageId="0" pageNumber="289">P. flavus</emphasis>
</taxonomicName>
D.J.Middleton &amp; Sangvir. is an endemic of North Thailand (
<bibRefCitation id="27D741F91CD1C3DAFC6BFC261B7276EC" author="Middleton, DJ" journalOrPublisher="Thai Forest Bulletin" pageId="0" pageNumber="289" pagination="15 - 17" refId="B11" refString="Middleton, DJ, Sangvirotjanapat, S, La-Ongsri, W, 2015. A new species of Petrocodon (Gesneriaceae) from Thailand. Thai Forest Bulletin 43: 15 - 17" title="A new species of Petrocodon (Gesneriaceae) from Thailand." volume="43" year="2015">Middleton et al. 2015</bibRefCitation>
). The new species is found in the karst region near the Lixianjiang river, the boundary of China with Vietnam and Laos, and thus, the new species may also extend to these countries. Currently, the new species is tentatively considered an endemic to China until detailed field investigations are carried out in its neigbouring countries.
</paragraph>
</subSubSection>
</treatment>
</document>

459
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<mods:title id="8A076CE41DCDA7D7DDBB1C251D875B2F">A new species of Aplectana (Nematoda: Cosmocercidae) parasite of Pleurodema nebulosum (Anura: Leptodactylidae) from the Monte desert, Argentina, with a key to Neotropical species of the genus Aplectana</mods:title>
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<treatment id="B60687ACFFC16B70FF70FB5EFA8312CD" ID-DOI="http://doi.org/10.5281/zenodo.6043261" ID-GBIF-Taxon="127852502" ID-Zenodo-Dep="6043261" LSID="urn:lsid:plazi:treatment:B60687ACFFC16B70FF70FB5EFA8312CD" httpUri="http://treatment.plazi.org/id/B60687ACFFC16B70FF70FB5EFA8312CD" lastPageId="4" lastPageNumber="125" pageId="1" pageNumber="122">
<subSubSection id="76B56531FFC16B75FF70FB5EFEA811BE" pageId="1" pageNumber="122" type="nomenclature">
<paragraph id="3E1036BAFFC16B75FF70FB5EFE50119D" blockId="1.[151,490,1194,1254]" box="[151,490,1194,1221]" pageId="1" pageNumber="122">
<heading id="655881D6FFC16B75FF70FB5EFE50119D" bold="true" box="[151,490,1194,1221]" fontSize="11" level="1" pageId="1" pageNumber="122" reason="1">
<emphasis id="0CDBEAA8FFC16B75FF70FB5EFE50119D" bold="true" box="[151,490,1194,1221]" pageId="1" pageNumber="122">
<taxonomicName id="F9AF4D39FFC16B75FF70FB5EFE3F119C" box="[151,389,1194,1220]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="1" pageNumber="122" phylum="Nematoda" rank="species" species="nebulosa" status="sp. nov.">
<emphasis id="0CDBEAA8FFC16B75FF70FB5EFE3F119C" bold="true" box="[151,389,1194,1220]" italics="true" pageId="1" pageNumber="122">Aplectana nebulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17E857D3FFC16B75FE6BFB5FFE50119D" box="[396,490,1195,1221]" pageId="1" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3E1036BAFFC16B75FF70FB39FEA811BE" blockId="1.[151,490,1194,1254]" box="[151,274,1229,1254]" pageId="1" pageNumber="122">
(
<figureCitation id="A6942A3FFFC16B75FF78FB39FEB211BE" box="[159,264,1229,1254]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1910,1932]" captionStartId-1="3.[151,250,1685,1707]" captionTargetBox-0="[289,1283,207,1884]" captionTargetBox-1="[151,1436,193,1663]" captionTargetId-0="figure@2.[267,1308,195,1894]" captionTargetId-1="figure@3.[151,1436,193,1663]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Aplectana nebulosa sp. nov. from P. nebulosum. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view." captionText-1="FIGURE 2. Aplectana nebulosa sp. nov. from P. nebulosum. A. Posterior end of male, detail of precloacal papillae. B. Posterior end of male, detail of adcloacal papillae and spicules. C. Posterior end of male, detail of postcloacal papillae. Abbreviations: ap: adcloacal papillae; up: unpaired papilla. In figure A, closed arrows indicate precloacal papillae; open arrows indicate lateral alae; in figure C, closed arrows indicate postcloacal papillae." httpUri-0="https://zenodo.org/record/438218/files/figure.png" httpUri-1="https://zenodo.org/record/438219/files/figure.png" pageId="1" pageNumber="122">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="76B56531FFC16B70FF70FAE1FD101445" lastPageId="4" lastPageNumber="125" pageId="1" pageNumber="122" type="description">
<paragraph id="3E1036BAFFC16B75FF70FAE1FE8410BA" blockId="1.[151,1436,1301,2010]" pageId="1" pageNumber="122">
<emphasis id="0CDBEAA8FFC16B75FF70FAE1FE2F1076" bold="true" box="[151,405,1301,1326]" pageId="1" pageNumber="122">General morphology.</emphasis>
Small, slender nematodes. Cuticle transversely striated. Slight sexual dimorphism, females larger than males (
<figureCitation id="A6942A3FFFC16B75FE89FACDFE15100A" box="[366,431,1337,1362]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1932]" captionTargetBox="[289,1283,207,1884]" captionTargetId="figure@2.[267,1308,195,1894]" captionTargetPageId="2" captionText="FIGURE 1. Aplectana nebulosa sp. nov. from P. nebulosum. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view." httpUri="https://zenodo.org/record/438218/files/figure.png" pageId="1" pageNumber="122">Fig. 1</figureCitation>
A, B). Narrow lateral alae present in both sexes, beginning at region of nerve ring and ending at the middle of tail in females and at level of first pair of precloacal papillae in males (
<figureCitation id="A6942A3FFFC16B75FB1DFAA9FAFC102E" box="[1274,1350,1373,1398]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1932]" captionTargetBox="[289,1283,207,1884]" captionTargetId="figure@2.[267,1308,195,1894]" captionTargetPageId="2" captionText="FIGURE 1. Aplectana nebulosa sp. nov. from P. nebulosum. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view." httpUri="https://zenodo.org/record/438218/files/figure.png" pageId="1" pageNumber="122">Figs. 1</figureCitation>
E–1K). Numerous small somatic papillae present. Mouth with three lips; dorsal lip with two cephalic papillae and each ventrolateral lip with one cephalic papilla and one large amphid (
<figureCitation id="A6942A3FFFC16B75FC90FA51FC0D10E6" box="[887,951,1445,1470]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1932]" captionTargetBox="[289,1283,207,1884]" captionTargetId="figure@2.[267,1308,195,1894]" captionTargetPageId="2" captionText="FIGURE 1. Aplectana nebulosa sp. nov. from P. nebulosum. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view." httpUri="https://zenodo.org/record/438218/files/figure.png" pageId="1" pageNumber="122">Fig. 1</figureCitation>
C). Excretory pore anterior of oesophageal bulb (
<figureCitation id="A6942A3FFFC16B75FF3FFA3DFEAD10BA" box="[216,279,1481,1506]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1932]" captionTargetBox="[289,1283,207,1884]" captionTargetId="figure@2.[267,1308,195,1894]" captionTargetPageId="2" captionText="FIGURE 1. Aplectana nebulosa sp. nov. from P. nebulosum. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view." httpUri="https://zenodo.org/record/438218/files/figure.png" pageId="1" pageNumber="122">Fig. 1</figureCitation>
D).
</paragraph>
<paragraph id="3E1036BAFFC16B75FF20FA19FC7212CA" blockId="1.[151,1436,1301,2010]" pageId="1" pageNumber="122">
<emphasis id="0CDBEAA8FFC16B75FF20FA19FEBB135E" box="[199,257,1517,1542]" italics="true" pageId="1" pageNumber="122">Male</emphasis>
(based on
<typeStatus id="E1148818FFC16B75FE65FA19FE5E135E" box="[386,484,1517,1542]" pageId="1" pageNumber="122" type="holotype">holotype</typeStatus>
and
<specimenCount id="28A9FD33FFC16B75FDC7FA19FD0C135E" box="[544,694,1517,1542]" pageId="1" pageNumber="122" type="generic" typeStatus="paratypes">15 paratypes</specimenCount>
): Total length 2.9 ±
<quantity id="F9579B5FFFC16B75FC44FA1AFBB0135D" box="[931,1034,1518,1542]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="122" unit="mm" value="0.41">0.41 mm</quantity>
(
<quantity id="F9579B5FFFC16B75FBFDFA19FB27135E" box="[1050,1181,1517,1542]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.5" metricValueMin="2.2" pageId="1" pageNumber="122" unit="mm" value="2.85" valueMax="3.5" valueMin="2.2">2.2–3.5mm</quantity>
) [
<quantity id="F9579B5FFFC16B75FB52FA1AFAAF135D" box="[1205,1301,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="122" unit="mm" value="2.9">2.9 mm</quantity>
]; maximum width 136.5 ± 25.4 (100–178) [125]. Pharynx length 40.7 ± 4.2 (35–49.5) [46], width 34.3 ± 5.1 (25–41) [37.7]; oesophagus: corpus length 320.2 ± 31.4 (280–392) [310] width 34.8 ± 5.3 (25–40) [25]; isthmus length 43.7 ± 8.4 (30–56.5) [30], isthmus width 23.8 ± 5.1 (18–35.5) [25]; bulb length 54.2 ± 9 (35.5–68) [60], bulb width 53.3 ± 5.8 (39–61) [55]. Nerve ring 223.0 ± 23.4 (175–255) [240] and excretory pore 331.7 ± 72.9 (191–502.5) [345], from anterior extremity, respectively. Tail length 167.9 ± 31.3 (105–220) [202.2]. Cloacal papillae arranged as follows (
<figureCitation id="A6942A3FFFC16B75FF78F931FF5A1386" box="[159,224,1733,1758]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1932]" captionTargetBox="[289,1283,207,1884]" captionTargetId="figure@2.[267,1308,195,1894]" captionTargetPageId="2" captionText="FIGURE 1. Aplectana nebulosa sp. nov. from P. nebulosum. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view." httpUri="https://zenodo.org/record/438218/files/figure.png" pageId="1" pageNumber="122">Fig. 1</figureCitation>
E, F): Two rows of 6–7 pairs of precloacal papillae, the last two paris located laterally to the cloaca (
<figureCitation id="A6942A3FFFC16B75FA88F931FF1C125A" captionStart="FIGURE 2" captionStartId="3.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1663]" captionTargetId="figure@3.[151,1436,193,1663]" captionTargetPageId="3" captionText="FIGURE 2. Aplectana nebulosa sp. nov. from P. nebulosum. A. Posterior end of male, detail of precloacal papillae. B. Posterior end of male, detail of adcloacal papillae and spicules. C. Posterior end of male, detail of postcloacal papillae. Abbreviations: ap: adcloacal papillae; up: unpaired papilla. In figure A, closed arrows indicate precloacal papillae; open arrows indicate lateral alae; in figure C, closed arrows indicate postcloacal papillae." httpUri="https://zenodo.org/record/438219/files/figure.png" pageId="1" pageNumber="122">Fig. 2</figureCitation>
A); 2 pairs of adcloacal papillae in anterior margin of cloaca and one middle, large unpaired papilla between the two anterior pairs (
<figureCitation id="A6942A3FFFC16B75FE89F8F9FE09127E" box="[366,435,1805,1830]" captionStart="FIGURE 2" captionStartId="3.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1663]" captionTargetId="figure@3.[151,1436,193,1663]" captionTargetPageId="3" captionText="FIGURE 2. Aplectana nebulosa sp. nov. from P. nebulosum. A. Posterior end of male, detail of precloacal papillae. B. Posterior end of male, detail of adcloacal papillae and spicules. C. Posterior end of male, detail of postcloacal papillae. Abbreviations: ap: adcloacal papillae; up: unpaired papilla. In figure A, closed arrows indicate precloacal papillae; open arrows indicate lateral alae; in figure C, closed arrows indicate postcloacal papillae." httpUri="https://zenodo.org/record/438219/files/figure.png" pageId="1" pageNumber="122">Fig. 2</figureCitation>
B); 5 pairs of postcloacal papillae (first and second pairs ventral, third pair lateral, fourth pair ventral, fifth pair lateral) (
<figureCitation id="A6942A3FFFC16B75FE10F8C5FD861212" box="[503,572,1841,1866]" captionStart="FIGURE 2" captionStartId="3.[151,250,1685,1707]" captionTargetBox="[151,1436,193,1663]" captionTargetId="figure@3.[151,1436,193,1663]" captionTargetPageId="3" captionText="FIGURE 2. Aplectana nebulosa sp. nov. from P. nebulosum. A. Posterior end of male, detail of precloacal papillae. B. Posterior end of male, detail of adcloacal papillae and spicules. C. Posterior end of male, detail of postcloacal papillae. Abbreviations: ap: adcloacal papillae; up: unpaired papilla. In figure A, closed arrows indicate precloacal papillae; open arrows indicate lateral alae; in figure C, closed arrows indicate postcloacal papillae." httpUri="https://zenodo.org/record/438219/files/figure.png" pageId="1" pageNumber="122">Fig. 2</figureCitation>
C); papillae pattern: 6–7:2:5+1. Spicules slightly equal, length 104.2 ± 12.4 (85–125.9) [110], pointed at distal end and with variably shaped capitulum (
<figureCitation id="A6942A3FFFC16B75FC0BF8A1FB961236" box="[1004,1068,1877,1902]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1932]" captionTargetBox="[289,1283,207,1884]" captionTargetId="figure@2.[267,1308,195,1894]" captionTargetPageId="2" captionText="FIGURE 1. Aplectana nebulosa sp. nov. from P. nebulosum. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view." httpUri="https://zenodo.org/record/438218/files/figure.png" pageId="1" pageNumber="122">Fig. 1</figureCitation>
G). Gubernaculum length 52.2 ± 9.1 (37.4–62.9) [55], weakly sclerotized, ovate in ventral view (
<figureCitation id="A6942A3FFFC16B75FC86F88DFC1B12CA" box="[865,929,1913,1938]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1932]" captionTargetBox="[289,1283,207,1884]" captionTargetId="figure@2.[267,1308,195,1894]" captionTargetPageId="2" captionText="FIGURE 1. Aplectana nebulosa sp. nov. from P. nebulosum. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view." httpUri="https://zenodo.org/record/438218/files/figure.png" pageId="1" pageNumber="122">Fig. 1</figureCitation>
H).
</paragraph>
<paragraph id="3E1036BAFFC16B70FF20F869FB40158C" blockId="1.[151,1436,1301,2010]" lastBlockId="4.[151,1437,151,1941]" lastPageId="4" lastPageNumber="125" pageId="1" pageNumber="122">
<emphasis id="0CDBEAA8FFC16B75FF20F869FEA612EE" box="[199,284,1949,1974]" italics="true" pageId="1" pageNumber="122">Female</emphasis>
(based on
<typeStatus id="E1148818FFC16B75FE72F869FE5512EE" box="[405,495,1949,1974]" pageId="1" pageNumber="122" type="allotype">allotype</typeStatus>
and
<specimenCount id="28A9FD33FFC16B75FDC1F86AFD0012EE" box="[550,698,1950,1974]" pageId="1" pageNumber="122" type="generic" typeStatus="paratypes">13 paratypes</specimenCount>
): Total length 4.6 ±
<quantity id="F9579B5FFFC16B75FC7DF86AFC4412ED" box="[922,1022,1950,1974]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="122" unit="mm" value="0.28">0.28 mm</quantity>
(
<quantity id="F9579B5FFFC16B75FBEBF869FB2C12EE" box="[1036,1174,1949,1974]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="5.8" metricValueMin="2.9" pageId="1" pageNumber="122" unit="mm" value="4.35" valueMax="5.8" valueMin="2.9">2.9–5.8 mm</quantity>
) [
<quantity id="F9579B5FFFC16B75FB4CF869FAA212ED" box="[1195,1304,1949,1974]" metricMagnitude="-3" metricUnit="m" metricValue="4.54" pageId="1" pageNumber="122" unit="mm" value="4.54">4.54 mm</quantity>
]; maximum width 200 ± 60.1 (100–298) [235]. Pharynx length 58.2 ± 15 (30–83) [70], pharynx width 49.2 ± 12.5 (28–72) [60]; oesophagus: corpus length 395.4 ± 65.0 (277–485) [425], width 47.6 ± 13.2 (30–79.1) [60]; isthmus length 48.5 ± 5.3 (41–58) [50], isthmus width 29.5 ± 4.7 (21–38) [40], bulb length 68.4 ± 9.8 (52–84) [80], bulb width 62.9 ± 9.4 (48–75) [70]. Nerve ring 249.4 ± 40.3 (200–325) [285] and excretory pore 417.7 ± 70 (234–506) [450] from anterior extremity, respectively. Vulva post-equatorial, transverse slit, slightly salient, situated 1.7 ±
<quantity id="F9579B5FFFC36B77FB19F86EFAE112EA" box="[1278,1371,1946,1970]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="124" unit="mm" value="0.43">0.43mm</quantity>
(1.0– 2.5) [
<quantity id="F9579B5FFFC36B77FF32F84BFE8D128F" box="[213,311,1983,2007]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="124" unit="mm" value="1.4">1.40mm</quantity>
] from posterior extremity; muscular ovejector, directed anteriorly (
<figureCitation id="A6942A3FFFC36B77FBACF84AFB2A128F" box="[1099,1168,1982,2007]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1932]" captionTargetBox="[289,1283,207,1884]" captionTargetId="figure@2.[267,1308,195,1894]" captionTargetPageId="2" captionText="FIGURE 1. Aplectana nebulosa sp. nov. from P. nebulosum. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view." httpUri="https://zenodo.org/record/438218/files/figure.png" pageId="3" pageNumber="124">
Fig.
<date id="4A11107AFFC36B77FB63F84BFB2A128F" box="[1156,1168,1983,2007]" pageId="3" pageNumber="124">1</date>
</figureCitation>
I, J), joining 2 uteri, 1 directed anteriorly, 1 posteriorly. Tail, conical, 261.7 ± 41.1 (200–343) [215] in length (
<figureCitation id="A6942A3FFFC46B70FB3AFF63FA9C15E8" box="[1245,1318,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1932]" captionTargetBox="[289,1283,207,1884]" captionTargetId="figure@2.[267,1308,195,1894]" captionTargetPageId="2" captionText="FIGURE 1. Aplectana nebulosa sp. nov. from P. nebulosum. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view." httpUri="https://zenodo.org/record/438218/files/figure.png" pageId="4" pageNumber="125">Fig. 1</figureCitation>
K). Eggs, unembryonated, thin shelled oval, length 84.8 ± 15.0 (61.5–105) [90], width 67 ± 14.2 (49–90) [80].
</paragraph>
<caption id="6AD06632FFC26B76FF70F882FB1B12B0" httpUri="https://zenodo.org/record/438218/files/figure.png" pageId="2" pageNumber="123" targetBox="[289,1283,207,1884]" targetPageId="2">
<paragraph id="3E1036BAFFC26B76FF70F882FB1B12B0" blockId="2.[151,1436,1910,2024]" pageId="2" pageNumber="123">
<emphasis id="0CDBEAA8FFC26B76FF70F882FEAD12D3" bold="true" box="[151,279,1910,1932]" pageId="2" pageNumber="123">FIGURE 1.</emphasis>
<taxonomicName id="F9AF4D39FFC26B76FEC6F882FE5712D4" box="[289,493,1910,1932]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="123" phylum="Nematoda" rank="species" species="nebulosa" status="sp. nov.">
<emphasis id="0CDBEAA8FFC26B76FEC6F882FE5712D4" box="[289,493,1910,1932]" italics="true" pageId="2" pageNumber="123">Aplectana nebulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17E857D3FFC26B76FE10F882FDF112D3" box="[503,587,1910,1931]" pageId="2" pageNumber="123" rank="species">
<emphasis id="0CDBEAA8FFC26B76FE10F882FDF112D3" bold="true" box="[503,587,1910,1931]" pageId="2" pageNumber="123">sp. nov.</emphasis>
</taxonomicNameLabel>
from
<taxonomicName id="F9AF4D39FFC26B76FD75F883FCA612D4" box="[658,796,1910,1932]" class="Amphibia" family="Leiuperidae" genus="Pleurodema" kingdom="Animalia" order="Anura" pageId="2" pageNumber="123" phylum="Chordata" rank="species" species="nebulosum">
<emphasis id="0CDBEAA8FFC26B76FD75F883FCA612D4" box="[658,796,1910,1932]" italics="true" pageId="2" pageNumber="123">P. nebulosum</emphasis>
</taxonomicName>
. A. Female, general view. B. Male, General view. C. Male, anterior extremity, apical view. D. Male, anterior extremity, lateral view. E. Male, posterior extremity, ventral view. F. Male, posterior extremity, lateral view. G. Spicule, ventral view. H. Gubernaculum, ventral view. I. Female, vulva and ovejector, ventral view. J. Female, vulva and ovejector, lateral view. K. Female, posterior extremity, lateral view.
</paragraph>
</caption>
<caption id="6AD06632FFC36B77FF70F961FC21125E" httpUri="https://zenodo.org/record/438219/files/figure.png" pageId="3" pageNumber="124" targetBox="[151,1436,193,1663]" targetPageId="3">
<paragraph id="3E1036BAFFC36B77FF70F961FC21125E" blockId="3.[151,1437,1684,1798]" pageId="3" pageNumber="124">
<emphasis id="0CDBEAA8FFC36B77FF70F961FEAD13F2" bold="true" box="[151,279,1685,1707]" pageId="3" pageNumber="124">FIGURE 2.</emphasis>
<taxonomicName id="F9AF4D39FFC36B77FEC5F960FE5513F2" box="[290,495,1684,1706]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="3" pageNumber="124" phylum="Nematoda" rank="species" species="nebulosa" status="sp. nov.">
<emphasis id="0CDBEAA8FFC36B77FEC5F960FE5513F2" box="[290,495,1684,1706]" italics="true" pageId="3" pageNumber="124">Aplectana nebulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17E857D3FFC36B77FE1DF961FDF513F2" box="[506,591,1685,1706]" pageId="3" pageNumber="124" rank="species">
<emphasis id="0CDBEAA8FFC36B77FE1DF961FDF513F2" bold="true" box="[506,591,1685,1706]" pageId="3" pageNumber="124">sp. nov.</emphasis>
</taxonomicNameLabel>
from
<emphasis id="0CDBEAA8FFC36B77FD71F962FC9213F2" box="[662,808,1684,1707]" italics="true" pageId="3" pageNumber="124">
<taxonomicName id="F9AF4D39FFC36B77FD71F962FC9E13F2" box="[662,804,1684,1707]" class="Amphibia" family="Leiuperidae" genus="Pleurodema" kingdom="Animalia" order="Anura" pageId="3" pageNumber="124" phylum="Chordata" rank="species" species="nebulosum">P. nebulosum</taxonomicName>
.
</emphasis>
A. Posterior end of male, detail of precloacal papillae. B. Posterior end of male, detail of adcloacal papillae and spicules. C. Posterior end of male, detail of postcloacal papillae. Abbreviations: ap: adcloacal papillae; up: unpaired papilla. In figure A, closed arrows indicate precloacal papillae; open arrows indicate lateral alae; in figure C, closed arrows indicate postcloacal papillae.
</paragraph>
</caption>
<paragraph id="3E1036BAFFC46B70FF20FF2BFE0715A0" blockId="4.[151,1437,151,1941]" box="[199,445,223,248]" pageId="4" pageNumber="125">
<emphasis id="0CDBEAA8FFC46B70FF20FF2BFE0715A0" bold="true" box="[199,445,223,248]" pageId="4" pageNumber="125">Taxonomy summary</emphasis>
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20FEF0FD101445" blockId="4.[151,1437,151,1941]" box="[199,682,260,285]" pageId="4" pageNumber="125">
<emphasis id="0CDBEAA8FFC46B70FF20FEF0FE311445" bold="true" box="[199,395,260,285]" pageId="4" pageNumber="125">Site of infection:</emphasis>
small and large intestine.
</paragraph>
</subSubSection>
<subSubSection id="76B56531FFC46B70FF20FED3FD3B1488" pageId="4" pageNumber="125" type="materials_examined">
<paragraph id="3E1036BAFFC46B70FF20FED3FB091418" blockId="4.[151,1437,151,1941]" box="[199,1203,295,320]" pageId="4" pageNumber="125">
<materialsCitation id="8EC73CE7FFC46B70FF20FED3FB091418" ID-GBIF-Occurrence="1449840821" box="[199,1203,295,320]" country="Argentina" location="Matagusanos" municipality="Ullum Department" pageId="4" pageNumber="125" specimenCount="1" stateProvince="San Juan Province" typeStatus="holotype">
<emphasis id="0CDBEAA8FFC46B70FF20FED3FED31418" bold="true" box="[199,361,295,320]" pageId="4" pageNumber="125">
<typeStatus id="E1148818FFC46B70FF20FED3FEB81418" box="[199,258,295,320]" pageId="4" pageNumber="125">Type</typeStatus>
locality:
</emphasis>
<location id="3B706061FFC46B70FE88FEDCFDBC1418" LSID="urn:lsid:plazi:treatment:B60687ACFFC16B70FF70FB5EFA8312CD:3B706061FFC46B70FE88FEDCFDBC1418" box="[367,518,296,320]" country="Argentina" municipality="Ullum Department" name="Matagusanos" pageId="4" pageNumber="125" stateProvince="San Juan Province">Matagusanos</location>
,
<collectingMunicipality id="DE74ACC0FFC46B70FDF6FED3FD5D1467" box="[529,743,295,320]" pageId="4" pageNumber="125">Ullúm Department</collectingMunicipality>
,
<collectingRegion id="FC6BF858FFC46B70FD15FEDCFC791418" box="[754,963,295,320]" country="Argentina" name="San Juan" pageId="4" pageNumber="125">
San
<collectingRegion id="FC6BF858FFC46B70FCC4FEDCFC791418" box="[803,963,295,320]" inferredBySuffix="true" pageId="4" pageNumber="125">Juan Province</collectingRegion>
</collectingRegion>
,
<collectingCountry id="46B8762AFFC46B70FC28FED3FB841418" box="[975,1086,295,320]" name="Argentina" pageId="4" pageNumber="125">Argentina</collectingCountry>
(
<figureCitation id="A6942A3FFFC46B70FBA9FED3FB2A1418" box="[1102,1168,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1121,1143]" captionTargetBox="[460,1128,193,1100]" captionTargetId="figure@5.[460,1128,193,1100]" captionTargetPageId="5" captionText="FIGURE 3. A. Adult male of Pleurodema nebulosum from Matagusanos. B. Panoramic view of type locality." httpUri="https://zenodo.org/record/438220/files/figure.png" pageId="4" pageNumber="125">Fig. 3</figureCitation>
B).
</materialsCitation>
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20FEB8FD3B1488" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="125">
<emphasis id="0CDBEAA8FFC46B70FF20FEB8FE74143D" bold="true" box="[199,462,332,357]" pageId="4" pageNumber="125">Deposition of types:</emphasis>
Holotype (male), allotype (female) and paratypes (2 males, 2 females) in Helminthological Collection of the Museum of La Plata (Holotype: MLP-He 7156; Allotype: MLP-He 7157; Paratypes: MLP-He 7158); Paratypes (5 males, 5 females) in Helminthological Collection of Centro de Ecología Aplicada del Litoral (CECOAL
<date id="4A11107AFFC46B70FDE7FE43FDCF1488" box="[512,629,439,464]" pageId="4" pageNumber="125" value="2601-06-16">16062601</date>
).
</paragraph>
</subSubSection>
<subSubSection id="76B56531FFC46B70FF20FE28FB991740" pageId="4" pageNumber="125" type="description">
<paragraph id="3E1036BAFFC46B70FF20FE28FB991740" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="125">
<emphasis id="0CDBEAA8FFC46B70FF20FE28FEE414AD" bold="true" box="[199,350,476,501]" pageId="4" pageNumber="125">Symbiotype:</emphasis>
<taxonomicName id="F9AF4D39FFC46B70FE8EFE28FCE014AD" authority="Burmeister, 1861" authorityName="Burmeister" authorityYear="1861" box="[361,858,476,501]" class="Amphibia" family="Leiuperidae" genus="Pleurodema" kingdom="Animalia" order="Anura" pageId="4" pageNumber="125" phylum="Chordata" rank="species" species="nebulosum">
<emphasis id="0CDBEAA8FFC46B70FE8EFE28FDCC14AD" box="[361,630,476,501]" italics="true" pageId="4" pageNumber="125">Pleurodema nebulosum</emphasis>
(Burmeister, 1861)
</taxonomicName>
(
<taxonomicName id="F9AF4D39FFC46B70FC8AFE28FC0D14AC" box="[877,951,476,500]" class="Amphibia" kingdom="Animalia" order="Anura" pageId="4" pageNumber="125" phylum="Chordata" rank="order">Anura</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FC20FE28FBC714AD" box="[967,1149,476,501]" class="Amphibia" family="Leptodactylidae" kingdom="Animalia" order="Anura" pageId="4" pageNumber="125" phylum="Chordata" rank="family">Leptodactylidae</taxonomicName>
) (
<figureCitation id="A6942A3FFFC46B70FB7EFE28FB4D14AC" box="[1177,1271,476,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1121,1143]" captionTargetBox="[460,1128,193,1100]" captionTargetId="figure@5.[460,1128,193,1100]" captionTargetPageId="5" captionText="FIGURE 3. A. Adult male of Pleurodema nebulosum from Matagusanos. B. Panoramic view of type locality." httpUri="https://zenodo.org/record/438220/files/figure.png" pageId="4" pageNumber="125">Fig. 3A</figureCitation>
), Herpetology Collection of Fundación Miguel Lillo (Deposited number FML 29422 to 29433).
</paragraph>
</subSubSection>
<subSubSection id="76B56531FFC46B70FF20FDD0FAF41738" pageId="4" pageNumber="125" type="etymology">
<paragraph id="3E1036BAFFC46B70FF20FDD0FC3A1765" blockId="4.[151,1437,151,1941]" box="[199,896,548,573]" pageId="4" pageNumber="125">
<emphasis id="0CDBEAA8FFC46B70FF20FDD0FEF41765" bold="true" box="[199,334,548,573]" pageId="4" pageNumber="125">Etymology.</emphasis>
The new species is named in reference to its host.
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20FDB3FAF41738" blockId="4.[151,1437,151,1941]" box="[199,1358,583,608]" pageId="4" pageNumber="125">
<emphasis id="0CDBEAA8FFC46B70FF20FDB3FE111738" bold="true" box="[199,427,583,608]" pageId="4" pageNumber="125">Host-parasite data.</emphasis>
Prevalence 58.5%, twenty-four of the frogs were found to harbour 397 nematodes.
</paragraph>
</subSubSection>
<subSubSection id="76B56531FFC46B70FF20FD98FA8312CD" pageId="4" pageNumber="125" type="discussion">
<paragraph id="3E1036BAFFC46B70FF20FD98FDAB17A8" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="125">
<emphasis id="0CDBEAA8FFC46B70FF20FD98FE8117DD" bold="true" box="[199,315,620,645]" pageId="4" pageNumber="125">Remarks.</emphasis>
The genus
<taxonomicName id="F9AF4D39FFC46B70FE5BFD98FD9417DD" box="[444,558,620,645]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="genus">
<emphasis id="0CDBEAA8FFC46B70FE5BFD98FD9417DD" box="[444,558,620,645]" italics="true" pageId="4" pageNumber="125">Aplectana</emphasis>
</taxonomicName>
is characterized by possessing lateral alae and somatic papillae, males without rosettes or plectanes on the posterior end, and females with numerous thin-shelled eggs in the uteri, and 2 ovaries anterior to the vulva (
<bibRefCitation id="5A3E4B4BFFC46B70FE6CFD40FD8D1794" author="Gibbons" box="[395,567,692,717]" pageId="4" pageNumber="125" refString="Gibbons, L. M. (2010) Keys to the nematode parasites of vertebrates. Supplementary Folume. CAB International and Natural History Museum, London, 146 pp." type="book" year="2010">Gibbons, 2010</bibRefCitation>
). Nematodes of this genus parasitize mainly amphibians and some species of reptiles worldwide (
<bibRefCitation id="5A3E4B4BFFC46B70FE90FD23FDBE17A8" author="Baker" box="[375,516,727,752]" pageId="4" pageNumber="125" refString="Baker, M. R. (1987) Synopsis of the Nematoda parasitic in amphibians and reptiles. Memorial University of Newfoundland. Occasional papers in Biology, 11, 1 - 325." type="journal article" year="1987">Baker, 1987</bibRefCitation>
).
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20FD08FE421605" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="125">
The species of genus
<taxonomicName id="F9AF4D39FFC46B70FE5BFD08FD94164D" box="[444,558,764,789]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="genus">
<emphasis id="0CDBEAA8FFC46B70FE5BFD08FD94164D" box="[444,558,764,789]" italics="true" pageId="4" pageNumber="125">Aplectana</emphasis>
</taxonomicName>
are grouped mainly according to the presence and absence of gubernaculum; other features that distinguish these helminths are the number and arrangement of caudal papillae, and the size of the gubernaculum and spicules.
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20FC93FA2C1125" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="125">
According to
<bibRefCitation id="5A3E4B4BFFC46B70FE82FC9CFDF116D8" box="[357,587,871,896]" pageId="4" pageNumber="125" refString="Bursey, C. R., Goldberg, S. R. &amp; Kraus, F. (2011) New Species of Aplectana (Nematoda: Cosmocercidae) in Sphenomorphus pratti from Papua New Guinea. Journal of Parasitology, 97 (4), 654 - 660." type="journal article">
Bursey
<emphasis id="0CDBEAA8FFC46B70FE5BFC9DFE4C16D8" box="[444,502,871,896]" italics="true" pageId="4" pageNumber="125">et al.</emphasis>
(2011)
</bibRefCitation>
, there are 26 species reported for the Neotropical Realm, 19 of them with gubernaculum:
<taxonomicName id="F9AF4D39FFC46B70FEA1FC78FC7816FC" authority="Ramallo, Bursey and Goldberg, 2008" authorityName="Ramallo, Bursey and Goldberg" authorityYear="2008" box="[326,962,908,933]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="adaechevarriae">
<emphasis id="0CDBEAA8FFC46B70FEA1FC78FDAF16FD" box="[326,533,908,933]" italics="true" pageId="4" pageNumber="125">A. adaechevarriae</emphasis>
Ramallo, Bursey and Goldberg, 2008
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FC29FC78FAC316FC" authority="Adamson and Baccam, 1988" authorityName="Adamson and Baccam" authorityYear="1988" box="[974,1401,908,933]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="albae">
<emphasis id="0CDBEAA8FFC46B70FC29FC78FB9116FD" box="[974,1067,908,933]" italics="true" pageId="4" pageNumber="125">A. albae</emphasis>
Adamson and Baccam, 1988
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FA63FC78FDB01690" authority="Baker and Vaucher, 1986" authorityName="Baker and Vaucher" authorityYear="1986" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="elenae">
<emphasis id="0CDBEAA8FFC46B70FA63FC78FF5A1690" italics="true" pageId="4" pageNumber="125">A. elenae</emphasis>
<bibRefCitation id="5A3E4B4BFFC46B70FF0FFC5BFDB01690" author="Baker" box="[232,522,943,968]" pageId="4" pageNumber="125" refString="Baker, M. R. &amp; Vaucher, C. (1986) Parasitic helminths from Paraguay XII: Aplectana Raillet and Henry, 1916 (Nematoda: Cosmocercoidea) from frogs. Revue Suisse de Zoologie, 93, 607 - 616." type="journal article" year="1986">Baker and Vaucher, 1986</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FDF0FC44FC3A1690" authority="Walton, 1940" authorityName="Walton" authorityYear="1940" box="[535,896,943,968]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="hamatospicula">
<emphasis id="0CDBEAA8FFC46B70FDF0FC44FD671690" box="[535,733,943,968]" italics="true" pageId="4" pageNumber="125">A. hamatospicula</emphasis>
Walton, 1940
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FC6AFC44FF6816B5" authority="Bursey, Goldberg and Telford, 2006" authorityName="Bursey, Goldberg and Telford" authorityYear="2006" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="herediaensis">
<emphasis id="0CDBEAA8FFC46B70FC6AFC44FB801690" box="[909,1082,943,968]" italics="true" pageId="4" pageNumber="125">A. herediaensis</emphasis>
Bursey, Goldberg and Telford, 2006
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FF06FC20FD4E16B4" authority="(Baylis, 1927) Travassos, 1931" authorityName="(Baylis, 1927) Travassos" authorityYear="1931" box="[225,756,980,1005]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="hylambatis">
<emphasis id="0CDBEAA8FFC46B70FF06FC20FEC516B5" box="[225,383,980,1005]" italics="true" pageId="4" pageNumber="125">A. hylambatis</emphasis>
(Baylis, 1927) Travassos, 1931
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FCE3FC20FB8116B4" authority="Caballero, 1949" authorityName="Caballero" authorityYear="1949" box="[772,1083,980,1005]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="incerta">
<emphasis id="0CDBEAA8FFC46B70FCE3FC20FCCC16B4" box="[772,886,980,1004]" italics="true" pageId="4" pageNumber="125">A. incerta</emphasis>
Caballero, 1949
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FBADFC20FF681148" authority="Bravo Hollis, 1943" authorityName="Bravo Hollis" authorityYear="1943" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="itzocanensis">
<emphasis id="0CDBEAA8FFC46B70FBADFC20FB4C16B4" box="[1098,1270,980,1004]" italics="true" pageId="4" pageNumber="125">A. itzocanensis</emphasis>
Bravo Hollis, 1943
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FF06FC0CFE631148" authority="Silva, 1955" authorityName="Silva" authorityYear="1955" box="[225,473,1015,1040]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="lopesi">
<emphasis id="0CDBEAA8FFC46B70FF06FC0CFEFD1148" box="[225,327,1015,1040]" italics="true" pageId="4" pageNumber="125">A. lopesi</emphasis>
Silva, 1955
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FE0FFC0CFB931148" authority="(Schneider, 1866) Miranda, 1924" authorityName="(Schneider, 1866) Miranda" authorityYear="1924" box="[488,1065,1015,1040]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="membranosa">
<emphasis id="0CDBEAA8FFC46B70FE0FFC0CFD261148" box="[488,668,1015,1040]" italics="true" pageId="4" pageNumber="125">A. membranosa</emphasis>
(Schneider, 1866) Miranda, 1924
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FBDEFC0CFEAE116D" authority="(Caballero, 1933) Ballesteros" authorityName="(Caballero, 1933) Ballesteros" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="mexicana">
<emphasis id="0CDBEAA8FFC46B70FBDEFC0CFB7D1148" box="[1081,1223,1016,1040]" italics="true" pageId="4" pageNumber="125">A. mexicana</emphasis>
(Caballero, 1933) Ballesteros
</taxonomicName>
–Marquez, 1945,
<taxonomicName id="F9AF4D39FFC46B70FE38FBE8FCFB116D" authority="Travassos, 1925" authorityName="Travassos" authorityYear="1925" box="[479,833,1052,1077]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="micropenis">
<emphasis id="0CDBEAA8FFC46B70FE38FBE8FDC6116C" box="[479,636,1052,1076]" italics="true" pageId="4" pageNumber="125">A. micropenis</emphasis>
Travassos, 1925
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FCA8FBE8FA9A116D" authority="Baker and Vaucher, 1986" authorityName="Baker and Vaucher" authorityYear="1986" box="[847,1312,1052,1077]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="paraelenae">
<emphasis id="0CDBEAA8FFC46B70FCA8FBE8FC4A116D" box="[847,1008,1052,1077]" italics="true" pageId="4" pageNumber="125">A. paraelenae</emphasis>
<bibRefCitation id="5A3E4B4BFFC46B70FC1DFBE8FA9A116D" author="Baker" box="[1018,1312,1052,1077]" pageId="4" pageNumber="125" refString="Baker, M. R. &amp; Vaucher, C. (1986) Parasitic helminths from Paraguay XII: Aplectana Raillet and Henry, 1916 (Nematoda: Cosmocercoidea) from frogs. Revue Suisse de Zoologie, 93, 607 - 616." type="journal article" year="1986">Baker and Vaucher, 1986</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FACAFBE8FE861100" authority="Miranda 1924" authorityName="Miranda" authorityYear="1924" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="pusilla">
<emphasis id="0CDBEAA8FFC46B70FACAFBE8FA26116D" box="[1325,1436,1052,1077]" italics="true" pageId="4" pageNumber="125">A. pusilla</emphasis>
Miranda 1924
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FEACFBB4FD301100" authority="Travassos, 1925" authorityName="Travassos" authorityYear="1925" box="[331,650,1087,1112]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="raillieti">
<emphasis id="0CDBEAA8FFC46B70FEACFBB4FE781100" box="[331,450,1087,1112]" italics="true" pageId="4" pageNumber="125">A. raillieti</emphasis>
Travassos, 1925
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FD7EFBB4FBDA1100" authority="Barus and Coy Otero, 1969" authorityName="Barus and Coy Otero" authorityYear="1969" box="[665,1120,1087,1112]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="rysavyi">
<emphasis id="0CDBEAA8FFC46B70FD7EFBB4FCB41100" box="[665,782,1088,1112]" italics="true" pageId="4" pageNumber="125">A. ryšavýi</emphasis>
Barus and Coy Otero, 1969
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FB88FBB4FE031124" authority="(Gomes and Motta, 1967) Baker, 1980" authorityName="(Gomes and Motta, 1967) Baker" authorityYear="1980" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="travassosi">
<emphasis id="0CDBEAA8FFC46B70FB88FBB4FABE1100" box="[1135,1284,1088,1112]" italics="true" pageId="4" pageNumber="125">A. travassosi</emphasis>
(Gomes and Motta, 1967)
<bibRefCitation id="5A3E4B4BFFC46B70FEC8FB90FE031124" author="Baker" box="[303,441,1124,1149]" pageId="4" pageNumber="125" refString="Baker, M. R. (1980) Revision of Old World species of the genus Aplectana Raillet and Henry, 1916 (Nematoda, Cosmocercidae). Bulletin du Museum National d'Histoire Naturelle, Series 4, Sect. A, 2, 955 - 998." type="journal article" year="1980">Baker, 1980</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FE23FB90FB9F1124" authority="Ramallo, Bursey and Goldberg, 2008" authorityName="Ramallo, Bursey and Goldberg" authorityYear="2008" box="[452,1061,1124,1149]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="tucumanensis">
<emphasis id="0CDBEAA8FFC46B70FE23FB90FDC11124" box="[452,635,1124,1148]" italics="true" pageId="4" pageNumber="125">A. tucumanensis</emphasis>
Ramallo, Bursey and Goldberg, 2008
</taxonomicName>
, and
<taxonomicName id="F9AF4D39FFC46B70FBB9FB90FA2B1125" authority="Travassos, 1926" authorityName="Travassos" authorityYear="1926" box="[1118,1425,1124,1149]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="vellardi">
<emphasis id="0CDBEAA8FFC46B70FBB9FB90FB691125" box="[1118,1235,1124,1149]" italics="true" pageId="4" pageNumber="125">A. vellardi</emphasis>
Travassos, 1926
</taxonomicName>
.
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20FB7CFB5A1054" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="125">
Apart from this, two groups can be identified, following the characteristics of caudal papillae: those with a large unpaired papilla anterior to the cloaca, and those without this character. Of the above mentioned species with gubernaculum, eleven have an unpaired papilla:
<taxonomicName id="F9AF4D39FFC46B70FD00FB24FCF611B0" box="[743,844,1231,1256]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="albae">
<emphasis id="0CDBEAA8FFC46B70FD00FB24FCF611B0" box="[743,844,1231,1256]" italics="true" pageId="4" pageNumber="125">A. albae</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FC84FB24FC6911B0" box="[867,979,1231,1256]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="elenae">
<emphasis id="0CDBEAA8FFC46B70FC84FB24FC6911B0" box="[867,979,1231,1256]" italics="true" pageId="4" pageNumber="125">A. elenae</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FC0EFB24FB0F11B0" box="[1001,1205,1231,1256]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="hamatospicula">
<emphasis id="0CDBEAA8FFC46B70FC0EFB24FB0F11B0" box="[1001,1205,1231,1256]" italics="true" pageId="4" pageNumber="125">A. hamatospicula</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FB2BFB24FAD411B0" box="[1228,1390,1231,1256]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="hylambatis">
<emphasis id="0CDBEAA8FFC46B70FB2BFB24FAD411B0" box="[1228,1390,1231,1256]" italics="true" pageId="4" pageNumber="125">A. hylambatis</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FA63FB24FE9A1054" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="itzocanensis">
<emphasis id="0CDBEAA8FFC46B70FA63FB24FE9A1054" italics="true" pageId="4" pageNumber="125">A. itzocanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FEC9FB00FE641055" box="[302,478,1268,1293]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="membranosa">
<emphasis id="0CDBEAA8FFC46B70FEC9FB00FE641055" box="[302,478,1268,1293]" italics="true" pageId="4" pageNumber="125">A. membranosa</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FE0CFB00FD321055" box="[491,648,1268,1293]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="paraelenae">
<emphasis id="0CDBEAA8FFC46B70FE0CFB00FD321055" box="[491,648,1268,1293]" italics="true" pageId="4" pageNumber="125">A. paraelenae</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FD71FB00FCB31055" box="[662,777,1268,1293]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="raillieti">
<emphasis id="0CDBEAA8FFC46B70FD71FB00FCB31055" box="[662,777,1268,1293]" italics="true" pageId="4" pageNumber="125">A. raillieti</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FCF0FB00FC3D1054" box="[791,903,1268,1292]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="rysavyi">
<emphasis id="0CDBEAA8FFC46B70FCF0FB00FC3D1054" box="[791,903,1268,1292]" italics="true" pageId="4" pageNumber="125">A. ryšavýi</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FC72FB00FB9C1054" box="[917,1062,1268,1292]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="travassosi">
<emphasis id="0CDBEAA8FFC46B70FC72FB00FB9C1054" box="[917,1062,1268,1292]" italics="true" pageId="4" pageNumber="125">A. travassosi</emphasis>
</taxonomicName>
, and
<taxonomicName id="F9AF4D39FFC46B70FB83FB00FB631055" box="[1124,1241,1268,1293]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="vellardi">
<emphasis id="0CDBEAA8FFC46B70FB83FB00FB631055" box="[1124,1241,1268,1293]" italics="true" pageId="4" pageNumber="125">A. vellardi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20FAE3FB0210C5" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="125">
The new species described here belongs to the group of species that possess both a gubernaculum and an unpaired papilla anterior to the cloaca, being more related to
<taxonomicName id="F9AF4D39FFC46B70FCA3FAC8FC4E100D" box="[836,1012,1340,1365]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="membranosa">
<emphasis id="0CDBEAA8FFC46B70FCA3FAC8FC4E100D" box="[836,1012,1340,1365]" italics="true" pageId="4" pageNumber="125">A. membranosa</emphasis>
</taxonomicName>
,
<taxonomicName id="F9AF4D39FFC46B70FBE6FAC8FB16100D" box="[1025,1196,1340,1365]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="paraelaenae">
<emphasis id="0CDBEAA8FFC46B70FBE6FAC8FB16100D" box="[1025,1196,1340,1365]" italics="true" pageId="4" pageNumber="125">A. paraelaenae</emphasis>
</taxonomicName>
and
<taxonomicName id="F9AF4D39FFC46B70FB04FAC8FACF100C" box="[1251,1397,1340,1364]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="travassosi">
<emphasis id="0CDBEAA8FFC46B70FB04FAC8FACF100C" box="[1251,1397,1340,1364]" italics="true" pageId="4" pageNumber="125">A. travassosi</emphasis>
</taxonomicName>
for possessing four adcloacal papillae, pre-oesophageal excretory pore, males with tail tapering gradually to spike-like distal portion, and gubernaculum elongated, with posterior end pointed and thickened borders.
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20FA5CFD2A1374" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="125">
However, the new species differs from
<taxonomicName id="F9AF4D39FFC46B70FD6EFA5CFC811098" box="[649,827,1447,1472]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="membranosa">
<emphasis id="0CDBEAA8FFC46B70FD6EFA5CFC811098" box="[649,827,1447,1472]" italics="true" pageId="4" pageNumber="125">A. membranosa</emphasis>
</taxonomicName>
by the following characters: 1) number of precloacal papillae (6–7 pairs vs. 3 pairs); 2) number and arrangement of postcloacal papillae (1° and 2° ventral, 3° lateral, 4° ventral, 5° lateral vs. 1° and 2° ventral, 3° lateral, 4° ventral); 3) shape and size of spicules (no forked spicules, 85– 125.9µm vs. forked spicules, 227–234.3µm).
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20F9C3FC8D13B8" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="125">
<taxonomicName id="F9AF4D39FFC46B70FF20F9C3FE121308" box="[199,424,1591,1616]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="nebulosa" status="sp. nov.">
<emphasis id="0CDBEAA8FFC46B70FF20F9C3FE121308" box="[199,424,1591,1616]" italics="true" pageId="4" pageNumber="125">Aplectana nebulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17E857D3FFC46B70FE54F9C3FDAA1308" box="[435,528,1591,1616]" pageId="4" pageNumber="125" rank="species">
<emphasis id="0CDBEAA8FFC46B70FE54F9C3FDAA1308" bold="true" box="[435,528,1591,1616]" pageId="4" pageNumber="125">sp. nov.</emphasis>
</taxonomicNameLabel>
can also be distinguished from
<taxonomicName id="F9AF4D39FFC46B70FC6DF9CCFB901308" box="[906,1066,1591,1616]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="paraelenae">
<emphasis id="0CDBEAA8FFC46B70FC6DF9CCFB901308" box="[906,1066,1591,1616]" italics="true" pageId="4" pageNumber="125">A. paraelenae</emphasis>
</taxonomicName>
by the following characters: 1) number of precloacal papillae (6–7 pairs vs. 3 pairs, respectively); 2) number and arrangement of postcloacal papillae (1° and 2° ventral, 3° lateral, 4° ventral, 5° lateral vs. 1°, 2°, 3° subventral, 4° and 5° subdorsal, 6° lateral); 3) size of spicules (85–125.9 µm vs. 227µm); 4) position of end of lateral alae in males (precloacal, at the level of 3° pair of precloacal papillae vs. postcloacal, at middle of tail).
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20F918FCE51228" blockId="4.[151,1437,151,1941]" pageId="4" pageNumber="125">
Finally,
<taxonomicName id="F9AF4D39FFC46B70FEC1F918FDB2125D" box="[294,520,1772,1797]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="nebulosa" status="sp. nov.">
<emphasis id="0CDBEAA8FFC46B70FEC1F918FDB2125D" box="[294,520,1772,1797]" italics="true" pageId="4" pageNumber="125">Aplectana nebulosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17E857D3FFC46B70FDF4F918FDCB125D" box="[531,625,1772,1797]" pageId="4" pageNumber="125" rank="species">
<emphasis id="0CDBEAA8FFC46B70FDF4F918FDCB125D" bold="true" box="[531,625,1772,1797]" pageId="4" pageNumber="125">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="F9AF4D39FFC46B70FCF6F918FC1C125C" box="[785,934,1772,1796]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="species" species="travassosi">
<emphasis id="0CDBEAA8FFC46B70FCF6F918FC1C125C" box="[785,934,1772,1796]" italics="true" pageId="4" pageNumber="125">A. travassosi</emphasis>
</taxonomicName>
by the following characters: 1) number of precloacal papillae (6–7 pairs vs. 3 pairs); 2) number and arrangement of postcloacal papillae (1° and 2° ventral, 3° lateral, 4° ventral, 5° lateral vs. 1° pair ventral, 2°, 3° and 4° subventral); 3) size of spicules (85–125.9µm vs. 230– 245µm); 4) size of gubernaculum (37.4–62.9µm vs. 80–95µm).
</paragraph>
<paragraph id="3E1036BAFFC46B70FF20F889FA8312CD" blockId="4.[151,1437,151,1941]" box="[199,1337,1916,1941]" pageId="4" pageNumber="125">
We provide a key for the species of
<taxonomicName id="F9AF4D39FFC46B70FDBEF888FD7112CD" box="[601,715,1916,1941]" class="Chromadorea" family="Cosmocercidae" genus="Aplectana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="125" phylum="Nematoda" rank="genus">
<emphasis id="0CDBEAA8FFC46B70FDBEF888FD7112CD" box="[601,715,1916,1941]" italics="true" pageId="4" pageNumber="125">Aplectana</emphasis>
</taxonomicName>
from amphibians and reptiles from Neotropical Realm.
</paragraph>
</subSubSection>
</treatment>
</document>

97
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@ -0,0 +1,97 @@
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Figs 47
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17
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</subSubSection>
<subSubSection pageId="180" pageNumber="181" type="type locality">
<paragraph pageId="180" pageNumber="181">Type locality.</paragraph>
<paragraph pageId="180" pageNumber="181">
PERU: Loreto: 20 km NNE Iquitos, Gen Gen [
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.6166668">3°37'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-73.3">73°18'W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection pageId="180" pageNumber="181" type="type material">
<paragraph pageId="180" pageNumber="181">Type material.</paragraph>
<paragraph pageId="180" pageNumber="181">
Holotype male: &quot;PERU: Loreto, 20 km NNE Iquitos, Rio
<normalizedToken originalValue="Momón">Momon</normalizedToken>
, Gen Gen, 120m, 5-6 Febr 2007, A.V. Petrov&quot;/ &quot;Caterino/Tishechkin
<taxonomicName lsidName="" pageId="180" pageNumber="181" rank="tribe" tribe="Exosternini">Exosternini</taxonomicName>
Voucher EXO-00319&quot; (FMNH).
</paragraph>
</subSubSection>
<subSubSection pageId="180" pageNumber="181" type="description">
<paragraph pageId="180" pageNumber="181">Diagnostic description.</paragraph>
<paragraph pageId="180" pageNumber="181">
Length: 1.97 mm, width: 1.65 mm; body rufo-piceous, smooth and shining, ovoid, rather strongly convex; frons depressed at middle, frontal stria divergent between eyes, central portion detached from sides; supraorbital stria fine, more or less complete, connected to frontal stria; epistoma convex, weakly emarginate at apex; pronotal disk strongly flattened across middle of base, but not plicate; disk with very fine ground punctation and few faint coarser punctures at sides; marginal pronotal stria absent behind head; lateral submarginal stria continuous with anterior submarginal stria; anterior pronotal gland openings present behind eye and midway between median opening and anterior pronotal corner; elytron with one complete epipleural stria, outer subhumeral stria present in apical half, inner subhumeral stria absent, the interval between outer subhumeral and 1st dorsal stria strongly swollen, striae 1-3 complete, 4th and 5th striae short, present only in apical one-fifth, sutural striae present in apical two-thirds, more broadly impressed anteriorly; prosternal keel broad, flat, weakly emarginate at base, carinal striae complete, convergent, united in front; mesoventrite weakly projecting at middle, marginal stria interrupted; mesometaventral stria broadly arched forward beyond middle of mesoventral disk, continued at sides by lateral metaventral stria toward outer third of metacoxa; 1st abdominal ventrite with single complete lateral stria, bent mediad at apex; propygidium with faint ground punctation, with large, shallow, elongate oval, punctures separated by about half their diameters; pygidium with dense but rather shallow ground punctation, small, coarser punctures uniformly interspersed; marginal pygidial sulcus fine, weakly crenulate, nearly complete but usually obsolete in basal corners. Male genitalia indistinguishable from those of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
(see Figs 48
<normalizedToken originalValue="A–D">A-D</normalizedToken>
, K).
</paragraph>
</subSubSection>
<subSubSection pageId="180" pageNumber="181" type="remarks">
<paragraph pageId="180" pageNumber="181">Remarks.</paragraph>
<paragraph pageId="180" pageNumber="181">
While this species is very similar to
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus farctus" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="farctus">Operclipygus farctus</taxonomicName>
in many characters, it is easily distinguished by its darker, more strongly convex, and rounder body form (Fig. 47F). The large, shallow punctures of the propygidium (Fig. 47G) and the absence of pronotal plicae distinguish it from some of the even more strongly convex species.
</paragraph>
</subSubSection>
<subSubSection pageId="180" pageNumber="181" type="etymology">
<paragraph pageId="180" pageNumber="181">Etymology.</paragraph>
<paragraph pageId="180" pageNumber="181">We name this species for scolytine specialist Alexander Petrov, in recognition of his numerous contributions of specimens to this study, principally from Peru.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>Integrative taxonomic revision of the land snail genus Sarika Godwin-Austen, 1907 in Thailand, with descriptions of nine new species (Eupulmonata, Ariophantidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pholyotha, Arthit</mods:namePart>
<mods:affiliation>Biological Sciences Program, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tongkerd, Piyoros</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>976</mods:number>
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<mods:start>1</mods:start>
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<mods:classification>journal article</mods:classification>
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<treatment ID-GBIF-Taxon="168504529" LSID="urn:lsid:zoobank.org:act:B61773AB-0762-4B4D-A097-162ED9C43720" httpUri="http://treatment.plazi.org/id/B6073C8928CE513DB8B08369C717ED51" lastPageNumber="1" pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="B6073C89-28CE-513D-B8B0-8369C717ED51" authority="Pholyotha &amp; Panha" authorityName="Pholyotha &amp; Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi" status="sp. nov.">Sarika gratesi Pholyotha &amp; Panha</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Geographic distribution of Sarika gratesi sp. nov. based on the specimens examined herein." figureDoi="10.3897/zookeys.976.53859.figure49" httpUri="https://binary.pensoft.net/fig/464431" pageId="0" pageNumber="1">Figs 49</figureCitation>
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">, 50A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
CUMZ 7275 (Fig.
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">50A</figureCitation>
, width 16.5 mm, height 8.8 mm).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
CUMZ 7276 (10 shells; Fig.
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">50B</figureCitation>
, width 16.0 mm, height 8.3 mm), 7912 (30 shells), NHMUK 20200289 (two shells), SMF (two shells), ZRC.MOL.017031 (two shells).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Thailand-Northeastern.</emphasis>
Dry dipterocarp forest at Phu Lan Kha, Nong Bua Daeng, Chaiyaphum,
<geoCoordinate degrees="16" direction="north" minutes="00" orientation="latitude" precision="1" seconds="00.9" value="16.00025">16°00'00.9&quot;N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="52" orientation="longitude" precision="1" seconds="33.4" value="101.875946">101°52'33.4&quot;E</geoCoordinate>
: CUMZ 7277.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
The limestone outcrop with dry deciduous forest at Tham Phraya Nakarat (Cave), Chum Phae, Khon Kaen, Thailand,
<geoCoordinate degrees="16" direction="north" minutes="48" orientation="latitude" precision="1" seconds="30.3" value="16.808416">16°48'30.3&quot;N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="57" orientation="longitude" precision="1" seconds="13.7" value="101.953804">101°57'13.7&quot;E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell medium-sized, depressed to strongly depressed, and pale brown. Aperture irregular with peristome rather simple above then expanded middle with curved inside aperture and thickened below periphery.</paragraph>
<caption doi="10.3897/zookeys.976.53859.figure49" httpUri="https://binary.pensoft.net/fig/464431" pageId="0" pageNumber="1" start="Figure 49" startId="F49">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 49.</emphasis>
Geographic distribution of
<taxonomicName authorityName="Pholyotha &amp; Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
<emphasis italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
</taxonomicName>
sp. nov. based on the specimens examined herein.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Shell</emphasis>
.
</emphasis>
Shell depressed to strongly depressed or nearly flattened, medium-sized (shell width up to 17.4 mm, shell height up to 8.8 mm), rather thin, and slightly opaque. Shell surface smooth, polished and with thin growth lines; shell colour pale brown. Whorls 6-7, increasing regularly; body whorl large and well rounded. Spire little to moderately elevated; suture impressed. Aperture crescent-shaped and open obliquely; peristome irregular. Apertural lip at upper periphery simple; at periphery with invagination of triangular lip (beak-like); at below periphery rather thickened inside aperture and little expanded. Columellar margin straight, slightly thickened and expanded near umbilicus. Umbilicus narrowly opened (Fig.
<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909)." figureDoi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1">50A, B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.976.53859.figure50" httpUri="https://binary.pensoft.net/fig/464432" pageId="0" pageNumber="1" start="Figure 50" startId="F50">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 50.</emphasis>
Shells of
<taxonomicName authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
</taxonomicName>
spp.
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
<taxonomicName authorityName="Pholyotha &amp; Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
<emphasis italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
holotype CUMZ 7275 and
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
paratype CUMZ 7276
<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
<taxonomicName lsidName="S. pumicata" pageId="0" pageNumber="1" rank="species" species="pumicata">
<emphasis italics="true" pageId="0" pageNumber="1">S. pumicata</emphasis>
</taxonomicName>
syntype NHMUK 1893.2.4.980-981
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName lsidName="S. ochtogyra" pageId="0" pageNumber="1" rank="species" species="ochtogyra">
<emphasis italics="true" pageId="0" pageNumber="1">S. ochtogyra</emphasis>
</taxonomicName>
holotype SMF 227103/1
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName lsidName="S. benoiti" pageId="0" pageNumber="1" rank="species" species="benoiti">
<emphasis italics="true" pageId="0" pageNumber="1">S. benoiti</emphasis>
</taxonomicName>
modified from
<bibRefCitation author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="343 - 379" refId="B13" refString="Crosse, H, Fischer, P, 1863. Note sur la faune malacologique de Cochinchine, comprenant la description des especes nouvelles ou peu connues. Journal de Conchyliologie 11: 343 - 379" title="Note sur la faune malacologique de Cochinchine, comprenant la description des especes nouvelles ou peu connues." volume="11" year="1863">Crosse and Fischer (1863)</bibRefCitation>
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
<taxonomicName lsidName="S. rex" pageId="0" pageNumber="1" rank="species" species="rex">
<emphasis italics="true" pageId="0" pageNumber="1">S. rex</emphasis>
</taxonomicName>
modified from
<bibRefCitation author="Preston, HB" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="202 - 202" refId="B71" refString="Preston, HB, 1909. Descriptions of new species of Macrochlamys and Pseudodon from Siam. Proceedings of the Malacological Society of London 8: 202 - 202" title="Descriptions of new species of Macrochlamys and Pseudodon from Siam." volume="8" year="1909">Preston (1909)</bibRefCitation>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific name
<taxonomicName lsidName="gratesi" pageId="0" pageNumber="1" rank="species" species="gratesi">
<emphasis italics="true" pageId="0" pageNumber="1">gratesi</emphasis>
</taxonomicName>
is named in honour to Admiral Chorchat Gra-tes of the Royal Thai Navy, who made possible many fieldtrips especially the remote islands areas in Thailand.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pholyotha &amp; Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
<emphasis italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
</taxonomicName>
sp. nov. is currently known from the restricted area of the dry deciduous and dry dipterocarp forests in Khon Kaen and Chaiyaphum provinces (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Geographic distribution of Sarika gratesi sp. nov. based on the specimens examined herein." figureDoi="10.3897/zookeys.976.53859.figure49" httpUri="https://binary.pensoft.net/fig/464431" pageId="0" pageNumber="1">49</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
This new species is easy to distinguish from all known
<taxonomicName authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
</taxonomicName>
as well as
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Macrochlamys</emphasis>
</taxonomicName>
species by its unique beak-like lip, while all other species in these two genera have simple to little thickened lips. Only,
<taxonomicName baseAuthorityName="Benson" baseAuthorityYear="1863" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys aspides" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aspides">
<emphasis italics="true" pageId="0" pageNumber="1">Macrochlamys aspides</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1080/00222936308681435" author="Benson, WH" journalOrPublisher="Bulletin du Musee ďHistoire Naturelle de Marseille, Serie II" pageId="0" pageNumber="1" refId="B7" refString="Benson, WH, 1863. Characters of new land-shells from the Andaman Islands, Burmah, and Ceylon, and of the animal of Sophina. Annals and Magazine of Natural History Series 3, 11: 318-323. https://doi.org/10.1080/00222936308681435" title="Characters of new land-shells from the Andaman Islands, Burmah, and Ceylon, and of the animal of Sophina. Annals and Magazine of Natural History Series 3, 11: 318 - 323." url="https://doi.org/10.1080/00222936308681435" year="1863">Benson 1863</bibRefCitation>
) from Myanmar tends to have a slightly thickened and expanded lip at below periphery, but
<taxonomicName authorityName="Pholyotha &amp; Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
<emphasis italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
</taxonomicName>
sp. nov. has a projecting and curved triangular shape lip at the periphery.
<taxonomicName authorityName="Pholyotha &amp; Panha" authorityYear="2020" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika gratesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gratesi">
<emphasis italics="true" pageId="0" pageNumber="1">Sarika gratesi</emphasis>
</taxonomicName>
sp. nov. is identical to
<taxonomicName authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
</taxonomicName>
because its shells are matched with
<taxonomicName authorityName="Godwin-Austen" authorityYear="1907" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sarika</emphasis>
</taxonomicName>
more than the Siam
<taxonomicName class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Macrochlamys</emphasis>
</taxonomicName>
(
<bibRefCitation author="Pholyotha, A" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="763 - 781" refId="B66" refString="Pholyotha, A, Sutcharit, C, Panha, S, 2018. The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae). Raffles Bulletin of Zoology 66: 763 - 781" title="The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)." volume="66" year="2018">Pholyotha et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

86
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6260165" ID-GBIF-Taxon="100127690" ID-Zenodo-Dep="6260165" LSID="urn:lsid:plazi:treatment:B60784A68429631DE745BC25580EA5C1" httpUri="http://treatment.plazi.org/id/B60784A68429631DE745BC25580EA5C1" lastPageNumber="128" pageNumber="128">
<subSubSection type="nomenclature">
<paragraph pageNumber="128">
<taxonomicName class="Arachnida" family="Phthiracaridae" genus="Steganacarus" kingdom="Animalia" order="Sarcoptiformes" pageNumber="128" phylum="Arthropoda" rank="species" species="brevipilus" subGenus="Tropacarus">Steganacarus (Tropacarus) brevipilus</taxonomicName>
(Berlese, 1923) [66a,b]
</paragraph>
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<subSubSection type="reference_group">
<paragraph pageNumber="128">
Syn., Tax.:
<taxonomicName class="Arachnida" family="Phthiracaridae" genus="Phthiracarus" kingdom="Animalia" order="Sarcoptiformes" pageNumber="128" phylum="Arthropoda" rank="species" species="brevipilus" subGenus="Trachychoplophora">Phthiracarus (Trachychoplophora) b.</taxonomicName>
Berlese, 1923.
<taxonomicName class="Arachnida" family="Phthiracaridae" genus="Tropacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oribatida" pageNumber="128" phylum="Arthropoda" rank="species" species="brevipilus">Tropacarus b.</taxonomicName>
: Van der Hammen 1959.
<taxonomicName class="Arachnida" family="Phthiracaridae" genus="Steganacarus" kingdom="Animalia" order="Sarcoptiformes" pageNumber="128" phylum="Arthropoda" rank="species" species="brevipilus">Steganacarus b.</taxonomicName>
: Bernini 1971 (B); Mahunka 1991c (B); Niedbala 1992 (B).
<taxonomicName class="Arachnida" family="Phthiracaridae" genus="Steganacarus" kingdom="Animalia" order="Sarcoptiformes" pageNumber="128" phylum="Arthropoda" rank="species" species="brevipilus" subGenus="Tropacarus">St. (Tropacarus) b.</taxonomicName>
: Bernini &amp; Avanzati 1989b (B); Niedbala 1994b (B).
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</subSubSection>
<subSubSection type="synonymic_list">
<paragraph pageNumber="128">
-
<taxonomicName class="Arachnida" family="Phthiracaridae" genus="Tropacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oribatida" pageNumber="128" phylum="Arthropoda" rank="species" species="curtipilus">Tropacarus curtipilus</taxonomicName>
: Strenzke 1952.
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<paragraph pageNumber="128">
<normalizedToken originalValue="Ökologie">Oekologie</normalizedToken>
: In moorigen
<normalizedToken originalValue="Böden">Boeden</normalizedToken>
(selten).
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<subSubSection type="distribution">
<paragraph pageNumber="128">
Verbreitung:
<normalizedToken originalValue="Paläarktis">Palaearktis</normalizedToken>
, Asien.
</paragraph>
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</treatment>
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237
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<emphasis id="0CDAEADAFF80FFE2FF39D3ABFE9F62AF" bold="true" box="[151,350,655,681]" italics="true" pageId="11" pageNumber="454">Batraxis ziyueae</emphasis>
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,
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(
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A, 12G–I, 14D, 14T, 15)
</paragraph>
</subSubSection>
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<materialsCitation id="8EC63C95FF80FFE2FF39D3DEFEBF6359" ID-GBIF-Occurrence="1290996358" collectingDate="2016-04-16" collectionCode="SNUC" collectorName="Zi-Chun Xiong" country="China" county="Lincang City" elevation="1243" latitude="24.730001" location="Manwan Town" longLatPrecision="21" longitude="100.33361" municipality="Yun Hsien" pageId="11" pageNumber="454" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="0CDAEADAFF80FFE2FF39D3DEFEFC6315" bold="true" box="[151,317,762,787]" pageId="11" pageNumber="454">
<typeStatus id="E115886AFF80FFE2FF39D3DEFF136315" box="[151,210,762,787]" pageId="11" pageNumber="454">Type</typeStatus>
material
</emphasis>
(2 ƋƋ,
<specimenCount id="28A8FD41FF80FFE2FE35D3DFFE026315" box="[411,451,763,787]" pageId="11" pageNumber="454" type="female">1 ♀</specimenCount>
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<typeStatus id="E115886AFF80FFE2FE72D3DEFD8C6315" box="[476,589,762,787]" pageId="11" pageNumber="454" type="holotype">Holotype</typeStatus>
:
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:
</emphasis>
Ƌ, labeled ‘
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,
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,
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,
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leg.’ (
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)
</materialsCitation>
.
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<typeStatus id="E115886AFF80FFE2FE27D262FDC46359" box="[393,517,838,863]" pageId="11" pageNumber="454" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="46B97658FF80FFE2FDBFD262FDB26359" box="[529,627,838,863]" name="China" pageId="11" pageNumber="454">CHINA</collectingCountry>
:
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1 Ƌ,
<specimenCount id="28A8FD41FF80FFE2FD18D263FD216358" box="[694,736,839,863]" pageId="11" pageNumber="454" type="female">1 ♀</specimenCount>
, same label data as
<typeStatus id="E115886AFF80FFE2FC6ED262FBE36359" box="[960,1058,838,863]" pageId="11" pageNumber="454" type="holotype">holotype</typeStatus>
except ‘
<date id="4A101008FF80FFE2FBD0D262FB3C6359" box="[1150,1277,838,863]" pageId="11" pageNumber="454" value="2016-04-20">
<collectingDate id="5A54E9E0FF80FFE2FBD0D262FB3C6359" box="[1150,1277,838,863]" pageId="11" pageNumber="454" value="2016-04-20">20.iv.2016</collectingDate>
</date>
’ (
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)
</materialsCitation>
.
</paragraph>
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<subSubSection id="76B46543FF80FFE2FF69D24EFC056414" pageId="11" pageNumber="454" type="diagnosis">
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<emphasis id="0CDAEADAFF80FFE2FF69D24EFE626385" bold="true" box="[199,419,874,899]" pageId="11" pageNumber="454">Diagnosis of male.</emphasis>
Body almost glabrous; frons with complete subantennal transverse sulcus; antennomeres IX as long as wide, IX obliquely subconical; pronotum with small median antebasal fovea and shallow, complete basal impression; all tibiae slender; protrochanters protuberant ventrally, profemora simple, protibiae concaved and protuberant near apex, mesotibiae lacking apical spur; tergite IV with broad, abrupt basal sulcus, discal carinae short, widely separated, with complete marginal carinae; aedeagus stout.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF80FFE2FF69D53AFE446749" pageId="11" pageNumber="454" type="description">
<paragraph id="3E1136C8FF80FFE2FF69D53AFAB4660D" blockId="11.[151,1437,762,1943]" pageId="11" pageNumber="454">
<emphasis id="0CDAEADAFF80FFE2FF69D53AFE986431" bold="true" box="[199,345,1054,1079]" pageId="11" pageNumber="454">Description.</emphasis>
Male (
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A). Body almost glabrous, reddish brown, length
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. Head rectangular, slightly wider than long, HL
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, HW
<quantity id="F9569B2DFF80FFE2FD76D567FCBE645C" box="[728,895,1091,1115]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="11" pageNumber="454" unit="mm" value="0.41" valueMax="0.42" valueMin="0.4">0.40–0.42 mm</quantity>
, vertex moderately convex, with pair of small, distinct vertexal foveae; antennal tubercles low; frons flattened, subantennal transverse sulcus complete, with shallow vertexal sulcus merging with subantennal transverse sulcus and connecting foveae; each eye composed of about 30 facets; antennae relatively thick, antennomeres III slightly elongate, IV‒VIII as long as wide, IX and X transverse, XI largest, widest at middle; median gular carina relatively narrow, with two separated nude foveae. Pronotum slightly wider than long, PL
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, PW
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, with small median antebasal fovea and shallow but complete basal impression. Elytra wider than long, EL 0.4 8–
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, EW
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, each elytron with two distinct foveae at base. Protrochanters protuberant at ventral margin (
<figureCitation id="A6952A4DFF80FFE2FBEFD41AFB566551" box="[1089,1175,1342,1367]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="11" pageNumber="454">Fig. 14</figureCitation>
T), protibiae distinctly concave before apex, with acute at cavity (
<figureCitation id="A6952A4DFF80FFE2FDDDD446FD05657D" box="[627,708,1378,1403]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="11" pageNumber="454">Fig. 14</figureCitation>
D). Abdomen as long as wide, AL
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, AW
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, obliquely narrowed at base; tergite IV (first visible tergite) with broad, abrupt basal sulcus; discal carinae short, widely separated, extending to apical one-fifth of tergal length, width between carinae
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at apices; with complete marginal carinae. Aedeagus (
<figureCitation id="A6952A4DFF80FFE2FD5BD4EAFC8865E1" box="[757,841,1486,1511]" captionStart="FIGURE 12" captionStartId="18.[151,250,1877,1899]" captionTargetBox="[166,1433,201,1840]" captionTargetId="figure@18.[151,1435,193,1856]" captionTargetPageId="18" captionText="FIGURE 12. Aedeagi of Batraxis species. (A, B, D, E, G, H. in ventral view. C, F, I. in lateral view). A – C. B. nanlingensis. D – F. B. yifan. G – I. B. ziyueae. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256553/files/figure.png" pageId="11" pageNumber="454">Fig. 12</figureCitation>
G–I)
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in length, stout, parameres strongly narrowed apically, each with two long apical setae; endophallus complicated, with pairs of symmetric sclerites.
</paragraph>
<paragraph id="3E1136C8FF80FFE2FF69D732FAAA6671" blockId="11.[151,1437,762,1943]" pageId="11" pageNumber="454">
Female. Similar to male in general appearance; protrochanters and protibiae simples; each eye composed of about 30 facets. Measurements: BL
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; HL
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; HW
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; PL
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; PW
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; EL
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; EW
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; AL
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; AW
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; width between discal carinae of tergite IV
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at apices.
</paragraph>
<paragraph id="3E1136C8FF80FFE2FF69D7A6FE446749" blockId="11.[151,1437,762,1943]" pageId="11" pageNumber="454">
<emphasis id="0CDAEADAFF80FFE2FF69D7A6FE72669D" bold="true" box="[199,435,1666,1691]" pageId="11" pageNumber="454">Comparative notes.</emphasis>
<taxonomicName id="F9AE4D4BFF80FFE2FE13D7A7FDB8669C" box="[445,633,1667,1690]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="454" phylum="Arthropoda" rank="species" species="ziyueae">
<emphasis id="0CDAEADAFF80FFE2FE13D7A7FDB8669C" box="[445,633,1667,1690]" italics="true" pageId="11" pageNumber="454">Batraxis ziyueae</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="F9AE4D4BFF80FFE2FCB9D7A7FBE2669D" authority="Wang &amp; Yin" authorityName="Wang &amp; Yin" box="[791,1059,1666,1691]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="454" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="0CDAEADAFF80FFE2FCB9D7A7FC46669C" box="[791,903,1666,1690]" italics="true" pageId="11" pageNumber="454">B. tibialis</emphasis>
Wang &amp; Yin
</taxonomicName>
by the thick antennae, complete subantennal transverse sulcus of the head, preapically concaved protibiae with a small tubercle inside the cavity, abdomen obliquely narrowed at the base, and short discal carinae. These two species can be separated by the simple profemora, and aedeagal parameres strongly narrowed at the apices in
<taxonomicName id="F9AE4D4BFF80FFE2FBA2D7CBFB416700" box="[1036,1152,1775,1798]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="454" phylum="Arthropoda" rank="species" species="ziyueae">
<emphasis id="0CDAEADAFF80FFE2FBA2D7CBFBDC6700" box="[1036,1053,1775,1798]" italics="true" pageId="11" pageNumber="454">B</emphasis>
.
<emphasis id="0CDAEADAFF80FFE2FB82D7CBFB416700" box="[1068,1152,1775,1798]" italics="true" pageId="11" pageNumber="454">ziyueae</emphasis>
</taxonomicName>
.
<taxonomicName id="F9AE4D4BFF80FFE2FB21D7CBFA856700" box="[1167,1348,1774,1798]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="454" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="0CDAEADAFF80FFE2FB21D7CBFA856700" box="[1167,1348,1774,1798]" italics="true" pageId="11" pageNumber="454">Batraxis tibialis</emphasis>
</taxonomicName>
has the profemora with a small protuberance at the middle of the ventral margin, and the aedeagal parameres are much broader at the apices.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF80FFE2FF69D67EFDBF6775" box="[199,638,1882,1907]" pageId="11" pageNumber="454" type="distribution">
<paragraph id="3E1136C8FF80FFE2FF69D67EFDBF6775" blockId="11.[151,1437,762,1943]" box="[199,638,1882,1907]" pageId="11" pageNumber="454">
<emphasis id="0CDAEADAFF80FFE2FF69D67EFEA16775" bold="true" box="[199,352,1882,1907]" pageId="11" pageNumber="454">Distribution.</emphasis>
<collectingCountry id="46B97658FF80FFE2FEC9D67EFE6F6775" box="[359,430,1882,1907]" name="China" pageId="11" pageNumber="454">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF80FFE2FE17D67FFDCF6774" box="[441,526,1883,1906]" country="China" name="Yunnan" pageId="11" pageNumber="454">Yunnan</collectingRegion>
(
<figureCitation id="A6952A4DFF80FFE2FDB0D67EFDB06775" box="[542,625,1882,1907]" captionStart="FIGURE 15" captionStartId="21.[151,250,1023,1045]" captionTargetBox="[151,1436,193,1001]" captionTargetId="figure@21.[151,1436,193,1002]" captionTargetPageId="21" captionText="FIGURE 15. Distribution of Batraxis species in China." httpUri="https://zenodo.org/record/256556/files/figure.png" pageId="11" pageNumber="454">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF80FFE2FF69D65AFA8C6791" box="[199,1357,1918,1943]" pageId="11" pageNumber="454" type="etymology">
<paragraph id="3E1136C8FF80FFE2FF69D65AFA8C6791" blockId="11.[151,1437,762,1943]" box="[199,1357,1918,1943]" pageId="11" pageNumber="454">
<emphasis id="0CDAEADAFF80FFE2FF69D65AFE8F6791" bold="true" box="[199,334,1918,1943]" pageId="11" pageNumber="454">Etymology.</emphasis>
The specific epithet is dedicated to the collector’s sister of this new species, Zi-Yue Xiong.
</paragraph>
</subSubSection>
</treatment>
</document>

266
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<emphasis id="0CDAEADAFF82FFE0FF39D003FDFF6144" bold="true" box="[151,574,295,322]" pageId="9" pageNumber="452">
<taxonomicName id="F9AE4D4BFF82FFE0FF39D003FE5E6147" ID-CoL="L28Z" box="[151,415,295,321]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="452" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="0CDAEADAFF82FFE0FF39D003FE5E6147" bold="true" box="[151,415,295,321]" italics="true" pageId="9" pageNumber="452">Batraxis nanlingensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="17E957A1FF82FFE0FE00D00CFDFF6144" box="[430,574,296,322]" pageId="9" pageNumber="452" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3E1136C8FF82FFE0FF39D06EFE086165" blockId="9.[151,574,295,355]" box="[151,457,330,355]" pageId="9" pageNumber="452">
(
<figureCitation id="A6952A4DFF82FFE0FF31D06EFF246165" box="[159,229,330,355]" captionStart="FIGURE 5" captionStartId="10.[151,250,1949,1971]" captionTargetBox="[158,1416,638,1914]" captionTargetId="figure@10.[151,1435,625,1928]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal habitus of Batraxis species. A. B. nanlingensis (female). B. B. yifan (male). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256546/files/figure.png" pageId="9" pageNumber="452">Figs 5</figureCitation>
A, 12A–C, 14R, 15)
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF82FFE0FF39D0B2FEB662BF" pageId="9" pageNumber="452" type="materials_examined">
<paragraph id="3E1136C8FF82FFE0FF39D0B2FEB662BF" blockId="9.[151,1437,405,1382]" pageId="9" pageNumber="452">
<materialsCitation id="8EC63C95FF82FFE0FF39D0B2FCBD61FC" ID-GBIF-Occurrence="1290996380" collectingDate="2015-05-05" collectionCode="SNUC" collectorName="Ruyuan Hsien &amp; Nanling N. R. &amp; Disilindao &amp; Peng &amp; Tu &amp; Zhou" country="China" elevation="1500" latitude="24.929722" location="Guangdong Province" longLatPrecision="21" longitude="112.997215" pageId="9" pageNumber="452" specimenCount="6" specimenCount-female="6" stateProvince="Guangdong Province" typeStatus="holotype">
<emphasis id="0CDAEADAFF82FFE0FF39D0B2FEFC61A9" bold="true" box="[151,317,406,431]" pageId="9" pageNumber="452">
<typeStatus id="E115886AFF82FFE0FF39D0B2FF1361A9" box="[151,210,406,431]" pageId="9" pageNumber="452">Type</typeStatus>
material
</emphasis>
(1 Ƌ,
<specimenCount id="28A8FD41FF82FFE0FE28D0B2FE0261A9" box="[390,451,406,431]" pageId="9" pageNumber="452" type="female">6 ♀♀</specimenCount>
).
<emphasis id="0CDAEADAFF82FFE0FE72D0B2FD0161A9" bold="true" box="[476,704,406,431]" pageId="9" pageNumber="452">
<typeStatus id="E115886AFF82FFE0FE72D0B2FD8C61A9" box="[476,589,406,431]" pageId="9" pageNumber="452" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="46B97658FF82FFE0FDF7D0B2FD7A61A9" box="[601,699,406,431]" name="China" pageId="9" pageNumber="452">CHINA</collectingCountry>
:
</emphasis>
Ƌ, labeled ‘
<collectingCountry id="46B97658FF82FFE0FCE8D0B3FC6961A9" box="[838,936,407,431]" name="China" pageId="9" pageNumber="452">CHINA</collectingCountry>
:
<collectingRegion id="FC6AF82AFF82FFE0FC1CD0B2FB6261A9" box="[946,1187,406,431]" country="China" name="Guangdong" pageId="9" pageNumber="452">Guangdong Province</collectingRegion>
,
<collectorName id="935B531EFF82FFE0FB03D0B3FA8D61A9" box="[1197,1356,406,431]" pageId="9" pageNumber="452">Ruyuan Hsien</collectorName>
(乳源 县),
<collectorName id="935B531EFF82FFE0FF60D099FEA561D0" box="[206,356,445,470]" pageId="9" pageNumber="452">Nanling N.R.</collectorName>
(南岭自然保护区),
<collectorName id="935B531EFF82FFE0FDF9D099FD0D61D0" box="[599,716,445,470]" pageId="9" pageNumber="452">Disilindao</collectorName>
(第四林道),
<geoCoordinate id="5B9A500FFF82FFE0FCC5D099FC3061D0" box="[875,1009,445,470]" direction="north" orientation="latitude" pageId="9" pageNumber="452" precision="15" value="24.929722">24°55'47&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF82FFE0FC52D099FB4C61D0" box="[1020,1165,445,470]" direction="east" orientation="longitude" pageId="9" pageNumber="452" precision="15" value="112.997215">112°59'50&quot;E</geoCoordinate>
, mixed forest, leaf litter, wood, sifted,
<quantity id="F9569B2DFF82FFE0FE9FD0C6FE4861FC" box="[305,393,482,507]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="452" unit="m" value="1500.0">
<elevation id="B583D1FBFF82FFE0FE9FD0C6FE4861FC" box="[305,393,482,507]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="452" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="4A101008FF82FFE0FE38D0C6FDC761FD" box="[406,518,482,507]" pageId="9" pageNumber="452" value="2015-05-05">
<collectingDate id="5A54E9E0FF82FFE0FE38D0C6FDC761FD" box="[406,518,482,507]" pageId="9" pageNumber="452" value="2015-05-05">05.v.2015</collectingDate>
</date>
,
<collectorName id="935B531EFF82FFE0FDBDD0C7FD8F61FD" box="[531,590,483,507]" pageId="9" pageNumber="452">Peng</collectorName>
,
<collectorName id="935B531EFF82FFE0FDF4D0C7FDB961FC" box="[602,632,483,506]" pageId="9" pageNumber="452">Tu</collectorName>
&amp;
<collectorName id="935B531EFF82FFE0FD30D0C6FD1861FD" box="[670,729,482,507]" pageId="9" pageNumber="452">Zhou</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF82FFE0FC8DD0C6FCB561FC" box="[803,884,482,506]" pageId="9" pageNumber="452">SNUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8EC63C95FF82FFE0FC26D0C6FEA16224" ID-GBIF-Occurrence="1290996350" collectionCode="SNUC" country="China" location="CHINA" pageId="9" pageNumber="452" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="0CDAEADAFF82FFE0FC26D0C6FBB661FD" bold="true" box="[904,1143,482,507]" pageId="9" pageNumber="452">
<typeStatus id="E115886AFF82FFE0FC26D0C6FBC561FD" box="[904,1028,482,507]" pageId="9" pageNumber="452" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="46B97658FF82FFE0FBBFD0C6FBB261FD" box="[1041,1139,482,507]" name="China" pageId="9" pageNumber="452">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="28A8FD41FF82FFE0FBD1D0C6FB0161FC" box="[1151,1216,482,506]" pageId="9" pageNumber="452" type="female">3 ♀♀</specimenCount>
, same label data as
<typeStatus id="E115886AFF82FFE0FF39D32EFF386224" box="[151,249,522,546]" pageId="9" pageNumber="452" type="holotype">holotype</typeStatus>
(
<collectionCode id="58BFAE0DFF82FFE0FEA9D32EFE996224" box="[263,344,522,546]" pageId="9" pageNumber="452">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF82FFE0FEC5D32EFF366268" ID-GBIF-Occurrence="1290996353" collectingDate="2015-04-28" collectionCode="SNUC" collectorName="Ruyuan Hsien &amp; Nanling N. R. &amp; Walkway &amp; Peng &amp; Tu &amp; Zhou" country="China" elevation="1220" latitude="24.932499" location="Guangdong Province" longLatPrecision="21" longitude="113.005" pageId="9" pageNumber="452" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong Province" typeStatus="paratype">
<specimenCount id="28A8FD41FF82FFE0FEC5D32EFE556227" box="[363,404,522,546]" pageId="9" pageNumber="452" type="female">1 ♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF82FFE0FE53D32EFD9E6224" box="[509,607,522,546]" name="China" pageId="9" pageNumber="452">CHINA</collectingCountry>
:
<collectingRegion id="FC6AF82AFF82FFE0FDC6D32DFC996224" box="[616,856,521,546]" country="China" name="Guangdong" pageId="9" pageNumber="452">Guangdong Province</collectingRegion>
,
<collectorName id="935B531EFF82FFE0FCCCD32EFBC56224" box="[866,1028,521,546]" pageId="9" pageNumber="452">Ruyuan Hsien</collectorName>
,
<collectorName id="935B531EFF82FFE0FBA0D32DFB676227" box="[1038,1190,521,546]" pageId="9" pageNumber="452">Nanling N.R.</collectorName>
,
<collectorName id="935B531EFF82FFE0FB1FD32DFADB6224" box="[1201,1306,521,546]" pageId="9" pageNumber="452">Walkway</collectorName>
(步行道),
<geoCoordinate id="5B9A500FFF82FFE0FF39D309FEDC6240" box="[151,285,557,582]" direction="north" orientation="latitude" pageId="9" pageNumber="452" precision="15" value="24.932499">24°55'57&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF82FFE0FE80D30AFE7F6240" box="[302,446,558,582]" direction="east" orientation="longitude" pageId="9" pageNumber="452" precision="15" value="113.005">113°00'18&quot;E</geoCoordinate>
, mixed forest, leaf litter, wood, sifted,
<quantity id="F9569B2DFF82FFE0FC3AD30AFC316240" box="[916,1008,558,582]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="9" pageNumber="452" unit="m" value="1220.0">
<elevation id="B583D1FBFF82FFE0FC3AD30AFC316240" box="[916,1008,558,582]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="9" pageNumber="452" unit="m" value="1220.0">1220 m</elevation>
</quantity>
,
<date id="4A101008FF82FFE0FBAFD309FBBB6240" box="[1025,1146,557,582]" pageId="9" pageNumber="452" value="2015-04-28">
<collectingDate id="5A54E9E0FF82FFE0FBAFD309FBBB6240" box="[1025,1146,557,582]" pageId="9" pageNumber="452" value="2015-04-28">28.iv.2015</collectingDate>
</date>
,
<collectorName id="935B531EFF82FFE0FB25D30BFB076241" box="[1163,1222,559,583]" pageId="9" pageNumber="452">Peng</collectorName>
,
<collectorName id="935B531EFF82FFE0FB78D30BFB356240" box="[1238,1268,559,582]" pageId="9" pageNumber="452">Tu</collectorName>
&amp;
<collectorName id="935B531EFF82FFE0FA8CD309FA9C6240" box="[1314,1373,557,582]" pageId="9" pageNumber="452">Zhou</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF82FFE0FF31D372FF316268" box="[159,240,598,622]" pageId="9" pageNumber="452">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF82FFE0FEACD372FEB262BF" ID-GBIF-Occurrence="1290996352" collectingDate="2015-04-26" collectionCode="SNUC" collectorName="Chenzhou &amp; Yizhang Hsien &amp; Mangshan N. R. &amp; Peng &amp; Tu &amp; Zhou" country="China" elevation="1400" latitude="24.940556" location="Hunan" longLatPrecision="21" longitude="112.98833" pageId="9" pageNumber="452" specimenCount="1" specimenCount-female="1" stateProvince="Hunan" typeStatus="paratype">
<specimenCount id="28A8FD41FF82FFE0FEACD372FEEA626B" box="[258,299,598,622]" pageId="9" pageNumber="452" type="female">1 ♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF82FFE0FE3AD372FE376268" box="[404,502,598,622]" name="China" pageId="9" pageNumber="452">CHINA</collectingCountry>
:
<collectingRegion id="FC6AF82AFF82FFE0FDAED372FD8C626B" box="[512,589,598,621]" country="China" name="Hunan" pageId="9" pageNumber="452">Hunan</collectingRegion>
,
<collectorName id="935B531EFF82FFE0FDF6D371FD08626B" box="[600,713,597,621]" pageId="9" pageNumber="452">Chenzhou</collectorName>
(郴州),
<collectorName id="935B531EFF82FFE0FC82D371FC116268" box="[812,976,597,622]" pageId="9" pageNumber="452">Yizhang Hsien</collectorName>
(宜章县),
<collectorName id="935B531EFF82FFE0FBE0D371FB3F626B" box="[1102,1278,597,622]" pageId="9" pageNumber="452">Mangshan N.R.</collectorName>
(莽山自然保 护区),
<geoCoordinate id="5B9A500FFF82FFE0FF43D359FEB26290" box="[237,371,637,662]" direction="north" orientation="latitude" pageId="9" pageNumber="452" precision="15" value="24.940556">24°56'26&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF82FFE0FED1D359FDCF6290" box="[383,526,637,662]" direction="east" orientation="longitude" pageId="9" pageNumber="452" precision="15" value="112.98833">112°59'18&quot;E</geoCoordinate>
, mixed forest, leaf litter, wood, sifted &amp; beating,
<quantity id="F9569B2DFF82FFE0FB9BD35AFB4D6293" box="[1077,1164,638,662]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="452" unit="m" value="1400.0">
<elevation id="B583D1FBFF82FFE0FB9BD35AFB4D6293" box="[1077,1164,638,662]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="452" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="4A101008FF82FFE0FB39D359FAD16290" box="[1175,1296,637,662]" pageId="9" pageNumber="452" value="2015-04-26">
<collectingDate id="5A54E9E0FF82FFE0FB39D359FAD16290" box="[1175,1296,637,662]" pageId="9" pageNumber="452" value="2015-04-26">26.iv.2015</collectingDate>
</date>
,
<collectorName id="935B531EFF82FFE0FAB5D35AFA976290" box="[1307,1366,638,662]" pageId="9" pageNumber="452">Peng</collectorName>
,
<collectorName id="935B531EFF82FFE0FACFD35AFABE6293" box="[1377,1407,638,661]" pageId="9" pageNumber="452">Tu</collectorName>
&amp;
<collectorName id="935B531EFF82FFE0FF39D385FF1362BC" box="[151,210,673,698]" pageId="9" pageNumber="452">Zhou</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF82FFE0FEB5D385FEAD62BF" box="[283,364,673,697]" pageId="9" pageNumber="452">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF82FFE0FF69D3E1FCDC651B" pageId="9" pageNumber="452" type="description">
<paragraph id="3E1136C8FF82FFE0FF69D3E1FB5B646F" blockId="9.[151,1437,405,1382]" pageId="9" pageNumber="452">
<emphasis id="0CDAEADAFF82FFE0FF69D3E1FE9862D8" bold="true" box="[199,345,709,734]" pageId="9" pageNumber="452">Description.</emphasis>
Male. Body dark-brown, with sparse, minute pubescence in most areas. Head rectangular, vertex moderately convex, with pair of distinct vertexal foveae; antennal tubercles low; frons flattened, subantennal transverse sulcus complete, with shallow vertexal sulcus merging with subantennal transverse sulcus and connecting foveae; each eye composed of about 30 facets; antennae relatively thin, antennomeres III‒VIII elongate, IX as long as wide, X transverse, XI largest, widest at middle; median gular carina shallow, with two separated nude foveae. Pronotum with indistinct median antebasal fovea and complete basal impression. Each elytron with two shallow foveae at base. Protrochanters protuberant at ventral margin (
<figureCitation id="A6952A4DFF82FFE0FBF4D2B9FB6A63B0" box="[1114,1195,925,950]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="9" pageNumber="452">Fig. 14</figureCitation>
R). Abdomen slightly narrowed at base; tergite IV (first visible tergite) with narrow basal sulcus; discal carinae relatively long, close, extending to apical one-third of tergal length; short marginal carinae extending to one-seventh of tergal length. Aedeagus (
<figureCitation id="A6952A4DFF82FFE0FEB8D52DFEA66427" box="[278,359,1033,1058]" captionStart="FIGURE 12" captionStartId="18.[151,250,1877,1899]" captionTargetBox="[166,1433,201,1840]" captionTargetId="figure@18.[151,1435,193,1856]" captionTargetPageId="18" captionText="FIGURE 12. Aedeagi of Batraxis species. (A, B, D, E, G, H. in ventral view. C, F, I. in lateral view). A – C. B. nanlingensis. D – F. B. yifan. G – I. B. ziyueae. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256553/files/figure.png" pageId="9" pageNumber="452">Fig. 12</figureCitation>
A–C)
<quantity id="F9569B2DFF82FFE0FE00D52DFDD56427" box="[430,532,1033,1057]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="452" unit="mm" value="0.27">0.27 mm</quantity>
in length, relatively stout, parameres distinctly exceeding apex of median lobe, rounded apically, each with two long apical setae; endophallus symmetric, comprised with three sclerites, with lateral two widely separated at base, and numerous weakly-sclerotized setae-like structures.
</paragraph>
<paragraph id="3E1136C8FF82FFE0FF69D551FD5364FC" blockId="9.[151,1437,405,1382]" pageId="9" pageNumber="452">
Female (
<figureCitation id="A6952A4DFF82FFE0FE84D551FEAC6488" box="[298,365,1141,1166]" captionStart="FIGURE 5" captionStartId="10.[151,250,1949,1971]" captionTargetBox="[158,1416,638,1914]" captionTargetId="figure@10.[151,1435,625,1928]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal habitus of Batraxis species. A. B. nanlingensis (female). B. B. yifan (male). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256546/files/figure.png" pageId="9" pageNumber="452">Fig. 5</figureCitation>
A). Similar to male in general appearance; protrochanters simple; each eye composed of about 30 facets. Measurements: BL
<quantity id="F9569B2DFF82FFE0FE4AD5BDFD4864B7" box="[484,649,1177,1201]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.79" metricValueMin="1.71" pageId="9" pageNumber="452" unit="mm" value="1.75" valueMax="1.79" valueMin="1.71">1.71–1.79 mm</quantity>
; HL
<quantity id="F9569B2DFF82FFE0FD6ED5BDFCA464B7" box="[704,869,1177,1201]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="9" pageNumber="452" unit="mm" value="0.41" valueMax="0.42" valueMin="0.4">0.40–0.42 mm</quantity>
; HW
<quantity id="F9569B2DFF82FFE0FC0BD5BDFB8B64B7" box="[933,1098,1177,1201]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="9" pageNumber="452" unit="mm" value="0.4" valueMax="0.42" valueMin="0.38">0.38–0.42 mm</quantity>
; PL
<quantity id="F9569B2DFF82FFE0FBD2D5BDFAE364B7" box="[1148,1314,1177,1201]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" metricValueMax="3.3" metricValueMin="2.9" pageId="9" pageNumber="452" unit="mm" value="0.31" valueMax="0.33" valueMin="0.29">0.29–0.33 mm</quantity>
; PW
<quantity id="F9569B2DFF82FFE0FAF3D5BDFEC164D3" metricMagnitude="-4" metricUnit="m" metricValue="3.8" metricValueMax="3.9" metricValueMin="3.7" pageId="9" pageNumber="452" unit="mm" value="0.38" valueMax="0.39" valueMin="0.37">0.37– 0.39 mm</quantity>
; EL
<quantity id="F9569B2DFF82FFE0FE97D599FE2364D3" box="[313,482,1213,1238]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" metricValueMax="5.699999999999999" metricValueMin="5.5" pageId="9" pageNumber="452" unit="mm" value="0.56" valueMax="0.57" valueMin="0.55">0.55–0.57 mm</quantity>
; EW
<quantity id="F9569B2DFF82FFE0FD8AD599FD0C64D3" box="[548,717,1213,1238]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.8999999999999995" metricValueMin="6.4" pageId="9" pageNumber="452" unit="mm" value="0.665" valueMax="0.69" valueMin="0.64">0.64–0.69 mm</quantity>
; AL
<quantity id="F9569B2DFF82FFE0FCA7D59AFC7064D3" box="[777,945,1214,1238]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="4.8" metricValueMin="4.699999999999999" pageId="9" pageNumber="452" unit="mm" value="0.475" valueMax="0.48" valueMin="0.47">0.47–0.48 mm</quantity>
; AW 0.6 2–
<quantity id="F9569B2DFF82FFE0FB9DD599FB5C64D3" box="[1075,1181,1213,1238]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="9" pageNumber="452" unit="mm" value="0.64">0.64 mm</quantity>
; width between discal carinae of tergite IV
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at apices.
</paragraph>
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<emphasis id="0CDAEADAFF82FFE0FF69D421FE716518" bold="true" box="[199,432,1285,1310]" pageId="9" pageNumber="452">Comparative notes.</emphasis>
See that under
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<emphasis id="0CDAEADAFF82FFE0FDF0D422FDAE651B" box="[606,623,1286,1309]" italics="true" pageId="9" pageNumber="452">B</emphasis>
.
<emphasis id="0CDAEADAFF82FFE0FDD3D421FD1B651B" box="[637,730,1285,1309]" italics="true" pageId="9" pageNumber="452">gloriosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17E957A1FF82FFE0FD4CD422FCDC651B" box="[738,797,1286,1309]" pageId="9" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF82FFE0FF69D40DFD6A6544" box="[199,683,1321,1346]" pageId="9" pageNumber="452" type="distribution">
<paragraph id="3E1136C8FF82FFE0FF69D40DFD6A6544" blockId="9.[151,1437,405,1382]" box="[199,683,1321,1346]" pageId="9" pageNumber="452">
<emphasis id="0CDAEADAFF82FFE0FF69D40DFEA16544" bold="true" box="[199,352,1321,1346]" pageId="9" pageNumber="452">Distribution.</emphasis>
<collectingCountry id="46B97658FF82FFE0FEC9D40DFE6F6544" box="[359,430,1321,1346]" name="China" pageId="9" pageNumber="452">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF82FFE0FE17D40DFDFD6544" box="[441,572,1321,1346]" country="China" name="Guangdong" pageId="9" pageNumber="452">Guangdong</collectingRegion>
(
<figureCitation id="A6952A4DFF82FFE0FDE5D40DFD5F6544" box="[587,670,1321,1346]" captionStart="FIGURE 15" captionStartId="21.[151,250,1023,1045]" captionTargetBox="[151,1436,193,1001]" captionTargetId="figure@21.[151,1436,193,1002]" captionTargetPageId="21" captionText="FIGURE 15. Distribution of Batraxis species in China." httpUri="https://zenodo.org/record/256556/files/figure.png" pageId="9" pageNumber="452">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF82FFE0FF69D469FB116563" box="[199,1232,1357,1382]" pageId="9" pageNumber="452" type="etymology">
<paragraph id="3E1136C8FF82FFE0FF69D469FB116563" blockId="9.[151,1437,405,1382]" box="[199,1232,1357,1382]" pageId="9" pageNumber="452">
<emphasis id="0CDAEADAFF82FFE0FF69D469FE8F6560" bold="true" box="[199,334,1357,1382]" pageId="9" pageNumber="452">Etymology.</emphasis>
The specific epithet is taken after the
<typeStatus id="E115886AFF82FFE0FD57D46AFCE86560" box="[761,809,1358,1382]" pageId="9" pageNumber="452">type</typeStatus>
locality, i.e. Nanling Nature Reserve.
</paragraph>
</subSubSection>
</treatment>
</document>

262
data/B6/07/87/B60787DEFF82FFE2FF39D49CFBA8623B.xml Normal file
View file

@ -0,0 +1,262 @@
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<mods:title id="0F3A60E0C297B07CB299EF4CE43CB05D">New species and records of Batraxis Reitter (Coleoptera: Staphylinidae: Pselaphinae) in continental China</mods:title>
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<mods:namePart id="7F4D3BDF4D283738BD0971D51F397D48">Wang, Dan</mods:namePart>
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<mods:namePart id="E7053BEBAD93DD6B5C8B08664AF4588A">Yin, Zi-Wei</mods:namePart>
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<mods:date id="8CE0F0A0B4E46C507B881FB1B2249D62">2016</mods:date>
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<taxonomicName id="F9AE4D4BFF82FFE0FF39D49CFE8265D4" ID-CoL="L29X" box="[151,323,1464,1490]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="452" phylum="Arthropoda" rank="species" species="yifan" status="sp. nov.">
<emphasis id="0CDAEADAFF82FFE0FF39D49CFE8265D4" bold="true" box="[151,323,1464,1490]" italics="true" pageId="9" pageNumber="452">Batraxis yifan</emphasis>
</taxonomicName>
,
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(
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B, 12D–F, 14G, 14S, 15)
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF82FFE0FF39D703FC6766B6" pageId="9" pageNumber="452" type="materials_examined">
<paragraph id="3E1136C8FF82FFE0FF39D703FC6766B6" blockId="9.[151,1437,1575,2036]" pageId="9" pageNumber="452">
<materialsCitation id="8EC63C95FF82FFE0FF39D703FDA8668A" ID-GBIF-Occurrence="1290996351" collectingDate="2016-03-14" collectionCode="SNUC" collectorName="Zi-Chun Xiong" country="China" county="Lincang City" elevation="2417" latitude="24.67472" location="Manwan Town" longLatPrecision="21" longitude="100.31389" municipality="Yun Hsien" pageId="9" pageNumber="452" specimenCount="7" specimenCount-female="7" stateProvince="Yunnan Province" typeStatus="holotype">
<emphasis id="0CDAEADAFF82FFE0FF39D703FEFC6646" bold="true" box="[151,317,1575,1600]" pageId="9" pageNumber="452">
<typeStatus id="E115886AFF82FFE0FF39D703FF136646" box="[151,210,1575,1600]" pageId="9" pageNumber="452">Type</typeStatus>
material
</emphasis>
(6 ƋƋ,
<specimenCount id="28A8FD41FF82FFE0FE36D70CFE156646" box="[408,468,1576,1600]" pageId="9" pageNumber="452" type="female">7 ♀♀</specimenCount>
).
<emphasis id="0CDAEADAFF82FFE0FE42D703FD116646" bold="true" box="[492,720,1575,1600]" pageId="9" pageNumber="452">
<typeStatus id="E115886AFF82FFE0FE42D703FD9C6646" box="[492,605,1575,1600]" pageId="9" pageNumber="452" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="46B97658FF82FFE0FDC7D703FD0A6646" box="[617,715,1575,1600]" name="China" pageId="9" pageNumber="452">CHINA</collectingCountry>
:
</emphasis>
Ƌ, labeled ‘
<collectingCountry id="46B97658FF82FFE0FCFAD70CFC776646" box="[852,950,1576,1600]" name="China" pageId="9" pageNumber="452">CHINA</collectingCountry>
:
<collectingRegion id="FC6AF82AFF82FFE0FC6ED70CFB436646" box="[960,1154,1575,1600]" country="China" name="Yunnan" pageId="9" pageNumber="452">Yunnan Province</collectingRegion>
,
<collectingCounty id="D7704E44FF82FFE0FB23D703FADE6646" box="[1165,1311,1575,1600]" pageId="9" pageNumber="452">Lincang City</collectingCounty>
(临沧市),
<collectingMunicipality id="DE75ACB2FF82FFE0FF39D774FECD666E" box="[151,268,1615,1640]" pageId="9" pageNumber="452">Yun Hsien</collectingMunicipality>
(云县),
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(漫湾镇),
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(水坝头村),
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,
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,
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<collectingDate id="5A54E9E0FF82FFE0FF39D757FED2668A" box="[151,275,1651,1676]" pageId="9" pageNumber="452" value="2016-03-14">14.iii.2016</collectingDate>
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,
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leg.’ (
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)
</materialsCitation>
.
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:
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:
</emphasis>
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,
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, same label data as
<typeStatus id="E115886AFF82FFE0FB61D757FAF0668D" box="[1231,1329,1651,1675]" pageId="9" pageNumber="452" type="holotype">holotype</typeStatus>
(
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)
</materialsCitation>
;
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1 Ƌ,
<specimenCount id="28A8FD41FF82FFE0FF61D7BCFF3866A9" box="[207,249,1688,1712]" pageId="9" pageNumber="452" type="female">1 ♀</specimenCount>
, same data as previous material except ‘
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’ (
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)
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.
</paragraph>
</subSubSection>
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<emphasis id="0CDAEADAFF82FFE0FF69D79FFE6766D2" bold="true" box="[199,422,1723,1748]" pageId="9" pageNumber="452">Diagnosis of male.</emphasis>
Body almost glabrous, elytra and abdomen with sparse, minute pubescence; frons with complete subantennal transverse sulcus; antennomeres IX slightly elongate, XI obliquely narrowed at apex; pronotum with distinct median antebasal fovea, lacking basal impression; all tibiae slender; protrochanters protuberant ventrally, profemora simple, protibiae lacking protuberance, mesotibiae with apical spur; tergite IV with narrow basal sulcus, discal carinae relatively long, close, with relatively long marginal carinae; aedeagus relatively elongate.
</paragraph>
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<emphasis id="0CDAEADAFF82FFE0FF69D6B7FE9867AA" bold="true" box="[199,345,1939,1964]" pageId="9" pageNumber="452">Description.</emphasis>
Male (
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B). Body almost glabrous, dark-brown, length
<quantity id="F9569B2DFF82FFE0FBB4D6B0FB0567AD" box="[1050,1220,1940,1964]" metricMagnitude="-3" metricUnit="m" metricValue="2.195" metricValueMax="2.2" metricValueMin="2.19" pageId="9" pageNumber="452" unit="mm" value="2.195" valueMax="2.2" valueMin="2.19">2.19–2.20 mm</quantity>
. Head rectangular, slightly longer than wide, HL
<quantity id="F9569B2DFF82FFE0FDAFD69CFD6A67C9" box="[513,683,1976,2000]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="4.9" metricValueMin="4.699999999999999" pageId="9" pageNumber="452" unit="mm" value="0.48" valueMax="0.49" valueMin="0.47">0.47–0.49 mm</quantity>
, HW
<quantity id="F9569B2DFF82FFE0FD58D69CFCA067C9" box="[758,865,1976,2000]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="9" pageNumber="452" unit="mm" value="0.47">0.47 mm</quantity>
, vertex moderately convex, with pair of distinct vertexal foveae; antennal tubercles low; frons flattened, subantennal transverse sulcus abrupt, with indistinct vertexal sulcus merging with subantennal transverse sulcus and connecting foveae; each eye composed of about 30 facets; antennae thin, antennomeres III‒IX elongate, X slightly transverse, XI largest, widest at middle; median gular carina relatively narrow, with two separated nude foveae. Pronotum slightly wider than long, PL
<quantity id="F9569B2DFF81FFE3FA82D1C4FF04611A" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="10" pageNumber="453" unit="mm" value="0.41" valueMax="0.42" valueMin="0.4">0.40–0.42 mm</quantity>
, PW
<quantity id="F9569B2DFF81FFE3FEAED020FE65611A" box="[256,420,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="10" pageNumber="453" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.50 mm</quantity>
, with distinct median antebasal fovea, lacking basal impression. Elytra wider than long, EL
<quantity id="F9569B2DFF81FFE3FF39D003FE856139" box="[151,324,295,320]" metricMagnitude="-4" metricUnit="m" metricValue="6.799999999999999" metricValueMax="6.8999999999999995" metricValueMin="6.7" pageId="10" pageNumber="453" unit="mm" value="0.6799999999999999" valueMax="0.69" valueMin="0.67">0.67–0.69 mm</quantity>
, EW
<quantity id="F9569B2DFF81FFE3FE3CD00CFDFE6139" box="[402,575,296,320]" metricMagnitude="-4" metricUnit="m" metricValue="8.35" metricValueMax="8.4" metricValueMin="8.299999999999999" pageId="10" pageNumber="453" unit="mm" value="0.835" valueMax="0.84" valueMin="0.83">0.83–0.84 mm</quantity>
, each elytron with two shallow foveae at base. Protrochanters weakly protuberant at ventral margin (
<figureCitation id="A6952A4DFF81FFE3FE56D068FD8C6162" box="[504,589,332,357]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="10" pageNumber="453">Fig. 14</figureCitation>
S), mesotibiae with apical tooth (
<figureCitation id="A6952A4DFF81FFE3FC65D068FBDE6162" box="[971,1055,332,357]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="10" pageNumber="453">Fig. 14</figureCitation>
G). Abdomen slightly wider than long, AL
<quantity id="F9569B2DFF81FFE3FEACD04BFE666181" box="[258,423,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.3" metricValueMin="6.2" pageId="10" pageNumber="453" unit="mm" value="0.625" valueMax="0.63" valueMin="0.62">0.62–0.63 mm</quantity>
, AW
<quantity id="F9569B2DFF81FFE3FE4BD054FD4B6181" box="[485,650,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" metricValueMax="7.4" metricValueMin="7.199999999999999" pageId="10" pageNumber="453" unit="mm" value="0.73" valueMax="0.74" valueMin="0.72">0.72–0.74 mm</quantity>
, slightly expanded at base; tergite IV (first visible tergite) with narrow basal sulcus; discal carinae relatively long, close, extending to apical one-third of tergal length, width between carinae
<quantity id="F9569B2DFF81FFE3FF5ED09CFE5561C9" box="[240,404,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="1.7999999999999998" metricValueMin="1.7" pageId="10" pageNumber="453" unit="mm" value="0.175" valueMax="0.18" valueMin="0.17">0.17–0.18 mm</quantity>
at apices; marginal carinae extending to half of tergal length. Aedeagus (
<figureCitation id="A6952A4DFF81FFE3FB78D093FAE661D6" box="[1238,1319,439,464]" captionStart="FIGURE 12" captionStartId="18.[151,250,1877,1899]" captionTargetBox="[166,1433,201,1840]" captionTargetId="figure@18.[151,1435,193,1856]" captionTargetPageId="18" captionText="FIGURE 12. Aedeagi of Batraxis species. (A, B, D, E, G, H. in ventral view. C, F, I. in lateral view). A – C. B. nanlingensis. D – F. B. yifan. G – I. B. ziyueae. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256553/files/figure.png" pageId="10" pageNumber="453">Fig. 12</figureCitation>
D–F)
<quantity id="F9569B2DFF81FFE3FAC5D09CFF0261F2" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="453" unit="mm" value="0.34">0.34 mm</quantity>
in length, relatively elongate, parameres distinctly exceeding apex of median lobe, protuberant at apicolateral margins, each with three apical setae; endophallus with two long sclerites and numerous spine-like structures, inclining toward right.
</paragraph>
<caption id="6AD16640FF81FFE3FF39D6B9FAEF67B5" box="[151,1326,1949,1971]" httpUri="https://zenodo.org/record/256546/files/figure.png" pageId="10" pageNumber="453" targetBox="[158,1416,638,1914]" targetPageId="10">
<paragraph id="3E1136C8FF81FFE3FF39D6B9FAEF67B5" blockId="10.[151,1326,1949,1971]" box="[151,1326,1949,1971]" pageId="10" pageNumber="453">
<emphasis id="0CDAEADAFF81FFE3FF39D6B9FED367B5" bold="true" box="[151,274,1949,1971]" pageId="10" pageNumber="453">FIGURE 5.</emphasis>
Dorsal habitus of
<taxonomicName id="F9AE4D4BFF81FFE3FE63D6BAFDE267B5" box="[461,547,1950,1971]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="0CDAEADAFF81FFE3FE63D6BAFDE267B5" box="[461,547,1950,1971]" italics="true" pageId="10" pageNumber="453">Batraxis</emphasis>
</taxonomicName>
species.
<emphasis id="0CDAEADAFF81FFE3FDD1D6B9FD5967B5" bold="true" box="[639,664,1949,1971]" pageId="10" pageNumber="453">A.</emphasis>
<taxonomicName id="F9AE4D4BFF81FFE3FD30D6BAFCFA67B4" box="[670,827,1949,1971]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="453" phylum="Arthropoda" rank="species" species="nanlingensis">
<emphasis id="0CDAEADAFF81FFE3FD30D6BAFCFA67B4" box="[670,827,1949,1971]" italics="true" pageId="10" pageNumber="453">B. nanlingensis</emphasis>
</taxonomicName>
(female).
<emphasis id="0CDAEADAFF81FFE3FC0DD6B9FC7B67B5" bold="true" box="[931,954,1949,1971]" pageId="10" pageNumber="453">B.</emphasis>
<taxonomicName id="F9AE4D4BFF81FFE3FC6FD6BAFBD167B4" box="[961,1040,1949,1971]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="453" phylum="Arthropoda" rank="species" species="yifan">
<emphasis id="0CDAEADAFF81FFE3FC6FD6BAFBD167B4" box="[961,1040,1949,1971]" italics="true" pageId="10" pageNumber="453">B. yifan</emphasis>
</taxonomicName>
(male). Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="3E1136C8FF80FFE2FF69D1B3FD1B611B" blockId="11.[151,1437,151,573]" pageId="11" pageNumber="454">
Female. Similar to male in general appearance; protrochanters and mesotibiae simple; each eye composed of about 30 facets. Measurements: BL
<quantity id="F9569B2DFF80FFE2FD87D198FD0F60D2" box="[553,718,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.12" metricValueMax="2.13" metricValueMin="2.11" pageId="11" pageNumber="454" unit="mm" value="2.12" valueMax="2.13" valueMin="2.11">2.11–2.13 mm</quantity>
; HL
<quantity id="F9569B2DFF80FFE2FCAAD198FC6B60D2" box="[772,938,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.0" metricValueMin="4.8" pageId="11" pageNumber="454" unit="mm" value="0.49" valueMax="0.5" valueMin="0.48">0.48–0.50 mm</quantity>
; HW
<quantity id="F9569B2DFF80FFE2FC44D198FB5160D2" box="[1002,1168,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="4.8" metricValueMin="4.699999999999999" pageId="11" pageNumber="454" unit="mm" value="0.475" valueMax="0.48" valueMin="0.47">0.47–0.48 mm</quantity>
; PL
<quantity id="F9569B2DFF80FFE2FB6CD198FAA660D2" box="[1218,1383,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.0" metricValueMin="3.9" pageId="11" pageNumber="454" unit="mm" value="0.395" valueMax="0.4" valueMin="0.39">0.39–0.40 mm</quantity>
; PW
<quantity id="F9569B2DFF80FFE2FF39D1FBFE8160F1" box="[151,320,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.699999999999999" metricValueMin="4.5" pageId="11" pageNumber="454" unit="mm" value="0.45999999999999996" valueMax="0.47" valueMin="0.45">0.45–0.47 mm</quantity>
; EL
<quantity id="F9569B2DFF80FFE2FED5D1FBFDE560F1" box="[379,548,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="11" pageNumber="454" unit="mm" value="0.665" valueMax="0.67" valueMin="0.66">0.66–0.67 mm</quantity>
; EW
<quantity id="F9569B2DFF80FFE2FDC6D1C4FCD060F1" box="[616,785,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="8.200000000000001" metricValueMax="8.299999999999999" metricValueMin="8.100000000000001" pageId="11" pageNumber="454" unit="mm" value="0.82" valueMax="0.83" valueMin="0.81">0.81–0.83 mm</quantity>
; AL
<quantity id="F9569B2DFF80FFE2FCE1D1FBFC3960F1" box="[847,1016,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="5.7" metricValueMax="6.0" metricValueMin="5.4" pageId="11" pageNumber="454" unit="mm" value="0.57" valueMax="0.6" valueMin="0.54">0.54–0.60 mm</quantity>
; AW
<quantity id="F9569B2DFF80FFE2FB93D1C4FB2660F1" box="[1085,1255,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="7.35" metricValueMax="7.4" metricValueMin="7.3" pageId="11" pageNumber="454" unit="mm" value="0.735" valueMax="0.74" valueMin="0.73">0.73–0.74 mm</quantity>
; width between discal carinae of tergite IV
<quantity id="F9569B2DFF80FFE2FE66D020FDAA611A" box="[456,619,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="1.6999999999999997" metricValueMax="1.7999999999999998" metricValueMin="1.6" pageId="11" pageNumber="454" unit="mm" value="0.16999999999999998" valueMax="0.18" valueMin="0.16">0.16–0.18 mm</quantity>
at apices.
</paragraph>
<paragraph id="3E1136C8FF80FFE2FF69D003FB2C61F3" blockId="11.[151,1437,151,573]" pageId="11" pageNumber="454">
<emphasis id="0CDAEADAFF80FFE2FF69D003FE776146" bold="true" box="[199,438,295,320]" pageId="11" pageNumber="454">Comparative notes.</emphasis>
<taxonomicName id="F9AE4D4BFF80FFE2FE6DD00CFDA76146" box="[451,614,296,320]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="454" phylum="Arthropoda" rank="species" species="yifan">
<emphasis id="0CDAEADAFF80FFE2FE6DD00CFDA76146" box="[451,614,296,320]" italics="true" pageId="11" pageNumber="454">Batraxis yifan</emphasis>
</taxonomicName>
is close to
<taxonomicName id="F9AE4D4BFF80FFE2FD55D00DFBE46146" authority="Wang &amp; Yin" authorityName="Wang &amp; Yin" box="[763,1061,295,320]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="454" phylum="Arthropoda" rank="species" species="spinosas">
<emphasis id="0CDAEADAFF80FFE2FD55D00DFC416139" box="[763,896,296,320]" italics="true" pageId="11" pageNumber="454">B. spinosas</emphasis>
Wang &amp; Yin
</taxonomicName>
by the similar general habitus, presence of an apical tooth on the mesotibiae, and aedeagus with parameres distinctly exceeding the apex of the median lobe. These two species can be readily separated by the presence of a distinct pronotal median fovea, and tergite IV with parallel discal carinae in the new species, and the aedeagal parameres are protuberant at apicolateral margins. In comparison,
<taxonomicName id="F9AE4D4BFF80FFE2FE03D09DFDED61C9" box="[429,556,440,464]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="454" phylum="Arthropoda" rank="species" species="spinosas">
<emphasis id="0CDAEADAFF80FFE2FE03D09DFDED61C9" box="[429,556,440,464]" italics="true" pageId="11" pageNumber="454">B. spinosas</emphasis>
</taxonomicName>
has an indistinct, shallow median fovea on the pronotum, the discal carinae of tergite IV are distinctly divergent, and the aedeagal parameres lack protuberance at lateral margins.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF80FFE2FF69D0DBFDBF621E" box="[199,638,511,536]" pageId="11" pageNumber="454" type="distribution">
<paragraph id="3E1136C8FF80FFE2FF69D0DBFDBF621E" blockId="11.[151,1437,151,573]" box="[199,638,511,536]" pageId="11" pageNumber="454">
<emphasis id="0CDAEADAFF80FFE2FF69D0DBFEA1621E" bold="true" box="[199,352,511,536]" pageId="11" pageNumber="454">Distribution.</emphasis>
<collectingCountry id="46B97658FF80FFE2FEC9D0DBFE6F621E" box="[359,430,511,536]" name="China" pageId="11" pageNumber="454">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF80FFE2FE17D324FDCF6211" box="[441,526,512,535]" country="China" name="Yunnan" pageId="11" pageNumber="454">Yunnan</collectingRegion>
(
<figureCitation id="A6952A4DFF80FFE2FDB0D0DBFDB0621E" box="[542,625,511,536]" captionStart="FIGURE 15" captionStartId="21.[151,250,1023,1045]" captionTargetBox="[151,1436,193,1001]" captionTargetId="figure@21.[151,1436,193,1002]" captionTargetPageId="21" captionText="FIGURE 15. Distribution of Batraxis species in China." httpUri="https://zenodo.org/record/256556/files/figure.png" pageId="11" pageNumber="454">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF80FFE2FF69D300FBA8623B" box="[199,1129,548,573]" pageId="11" pageNumber="454" type="etymology">
<paragraph id="3E1136C8FF80FFE2FF69D300FBA8623B" blockId="11.[151,1437,151,573]" box="[199,1129,548,573]" pageId="11" pageNumber="454">
<emphasis id="0CDAEADAFF80FFE2FF69D300FE8F623B" bold="true" box="[199,334,548,573]" pageId="11" pageNumber="454">Etymology.</emphasis>
The specific epithet ‘
<emphasis id="0CDAEADAFF80FFE2FDEFD300FD43623A" box="[577,642,548,572]" italics="true" pageId="11" pageNumber="454">yifan’</emphasis>
is the nick name of the collector’s nephew.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="3E1136C8FF83FFE1FF39D2D3FDF06414" blockId="8.[151,561,1015,1075]" box="[151,561,1015,1042]" pageId="8" pageNumber="451">
<heading id="655981A4FF83FFE1FF39D2D3FDF06414" bold="true" box="[151,561,1015,1042]" fontSize="11" level="1" pageId="8" pageNumber="451" reason="1">
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<taxonomicName id="F9AE4D4BFF83FFE1FF39D2D3FE536417" ID-CoL="5WFV2" box="[151,402,1015,1041]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="451" phylum="Arthropoda" rank="species" species="longithorax" status="sp. nov.">
<emphasis id="0CDAEADAFF83FFE1FF39D2D3FE536417" bold="true" box="[151,402,1015,1041]" italics="true" pageId="8" pageNumber="451">Batraxis longithorax</emphasis>
</taxonomicName>
,
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</emphasis>
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(
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B, 11G–I, 14Q, 15)
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF83FFE1FF39D542FE3364CC" pageId="8" pageNumber="451" type="materials_examined">
<paragraph id="3E1136C8FF83FFE1FF39D542FE3364CC" blockId="8.[151,1437,1125,2019]" pageId="8" pageNumber="451">
<materialsCitation id="8EC63C95FF83FFE1FF39D542FE2F64CC" ID-GBIF-Occurrence="1290996369" collectingDate="2014-07-12" collectionCode="SNUC" collectorName="Peng &amp; Song &amp; Yu &amp; Yan" country="China" elevation="790" latitude="24.133888" location="Luohan Shan" longLatPrecision="21" longitude="110.12723" municipality="Jinxiu Country" pageId="8" pageNumber="451" specimenCount="2" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="0CDAEADAFF83FFE1FF39D542FEFC6479" bold="true" box="[151,317,1126,1151]" pageId="8" pageNumber="451">
<typeStatus id="E115886AFF83FFE1FF39D542FF136479" box="[151,210,1126,1151]" pageId="8" pageNumber="451">Type</typeStatus>
material
</emphasis>
(1 Ƌ).
<emphasis id="0CDAEADAFF83FFE1FE22D542FDAE6479" bold="true" box="[396,623,1126,1151]" pageId="8" pageNumber="451">
<typeStatus id="E115886AFF83FFE1FE22D542FE3C6479" box="[396,509,1126,1151]" pageId="8" pageNumber="451" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="46B97658FF83FFE1FDA7D542FDAA6479" box="[521,619,1126,1151]" name="China" pageId="8" pageNumber="451">CHINA</collectingCountry>
:
</emphasis>
Ƌ, labeled ‘
<collectingCountry id="46B97658FF83FFE1FD5DD543FC946479" box="[755,853,1127,1151]" name="China" pageId="8" pageNumber="451">CHINA</collectingCountry>
:
<collectingRegion id="FC6AF82AFF83FFE1FCF1D542FC026479" box="[863,963,1126,1151]" country="China" name="Guangxi" pageId="8" pageNumber="451">Guangxi</collectingRegion>
,
<collectingMunicipality id="DE75ACB2FF83FFE1FC63D542FBB56479" box="[973,1140,1126,1151]" pageId="8" pageNumber="451">Jinxiu Country</collectingMunicipality>
(金秀县),
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(罗汉山),
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,
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, beech forest, mixed leaf litter, grass, sifted, alt.
<quantity id="F9569B2DFF83FFE1FBDDD5AAFB7F64A0" box="[1139,1214,1166,1190]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="8" pageNumber="451" unit="m" value="790.0">
<elevation id="B583D1FBFF83FFE1FBDDD5AAFB7F64A0" box="[1139,1214,1166,1190]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="8" pageNumber="451" unit="m" value="790.0">790 m</elevation>
</quantity>
,
<date id="4A101008FF83FFE1FB63D5A9FA8E64A0" box="[1229,1359,1165,1190]" pageId="8" pageNumber="451" value="2014-07-12">
<collectingDate id="5A54E9E0FF83FFE1FB63D5A9FA8E64A0" box="[1229,1359,1165,1190]" pageId="8" pageNumber="451" value="2014-07-12">12.vii.2014</collectingDate>
</date>
,
<collectorName id="935B531EFF83FFE1FAF3D5ABFA5964A1" box="[1373,1432,1167,1191]" pageId="8" pageNumber="451">Peng</collectorName>
,
<collectorName id="935B531EFF83FFE1FF39D596FF1264CC" box="[151,211,1202,1226]" pageId="8" pageNumber="451">Song</collectorName>
,
<collectorName id="935B531EFF83FFE1FF70D597FF3C64CC" box="[222,253,1203,1226]" pageId="8" pageNumber="451">Yu</collectorName>
&amp;
<collectorName id="935B531EFF83FFE1FE8FD597FE8C64CC" box="[289,333,1203,1226]" pageId="8" pageNumber="451">Yan</collectorName>
leg.’ (
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF83FFE1FF69D5F2FD1F6579" pageId="8" pageNumber="451" type="diagnosis">
<paragraph id="3E1136C8FF83FFE1FF69D5F2FD1F6579" blockId="8.[151,1437,1125,2019]" pageId="8" pageNumber="451">
<emphasis id="0CDAEADAFF83FFE1FF69D5F2FE6964E9" bold="true" box="[199,424,1238,1263]" pageId="8" pageNumber="451">Diagnosis of male.</emphasis>
Body pubescent; frons with complete subantennal transverse sulcus; antennomeres IX strongly transverse, IX enlarged; pronotum with shallow median antebasal fovea and distinct basal impression; all tibiae slender; protrochanters strongly protuberant ventrally, profemora simple, protibiae lacking protuberance, mesotibiae lacking apical spur; tergite IV with narrow basal sulcus, discal carinae long, moderately close, with relatively long marginal carinae; aedeagus elongate.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF83FFE1FF69D4AEFB2367E4" pageId="8" pageNumber="451" type="description">
<paragraph id="3E1136C8FF83FFE1FF69D4AEFD0D6771" blockId="8.[151,1437,1125,2019]" pageId="8" pageNumber="451">
<emphasis id="0CDAEADAFF83FFE1FF69D4AEFE9865A5" bold="true" box="[199,345,1418,1443]" pageId="8" pageNumber="451">Description.</emphasis>
Male (
<figureCitation id="A6952A4DFF83FFE1FE1FD4AEFE3B65A4" box="[433,506,1418,1443]" captionStart="FIGURE 4" captionStartId="7.[151,250,1979,2001]" captionTargetBox="[158,1427,724,1948]" captionTargetId="figure@7.[151,1435,707,1958]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal habitus of Batraxis males. A. B. gesar. B. B. longithorax. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256545/files/figure.png" pageId="8" pageNumber="451">Fig. 4</figureCitation>
B). Body reddish-brown, pubescent, length
<quantity id="F9569B2DFF83FFE1FC56D4AEFBA565A4" box="[1016,1124,1418,1443]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" pageId="8" pageNumber="451" unit="mm" value="1.76">1.76 mm</quantity>
. Head slightly trapezoidal, slightly longer than wide, HL
<quantity id="F9569B2DFF83FFE1FE5BD48BFD9F65C0" box="[501,606,1455,1479]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="451" unit="mm" value="0.4">0.40 mm</quantity>
, HW
<quantity id="F9569B2DFF83FFE1FD0BD48BFCCC65C0" box="[677,781,1455,1479]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="451" unit="mm" value="0.37">0.37 mm</quantity>
, vertex moderately convex, with pair of distinct vertexal foveae; antennal tubercles low; frons flattened and narrow, subantennal transverse sulcus complete, anterior margin of V-shaped vertexal sulcus merging with subantennal sulcus and connecting foveae; each eye composed of about 30 facets; antennae thick, antennomeres III slightly elongate, IV as long as wide, V‒X transverse, XI largest, widest at basal fourth; median gular carina narrow, with two separated nude foveae. Pronotum as long as wide, PL
<quantity id="F9569B2DFF83FFE1FF39D746FF3E667C" box="[151,255,1634,1659]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="451" unit="mm" value="0.35">0.35 mm</quantity>
, PW
<quantity id="F9569B2DFF83FFE1FEEED746FE69667C" box="[320,424,1634,1658]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="451" unit="mm" value="0.34">0.34 mm</quantity>
, with shallow median antebasal fovea and complete basal impression. Elytra wider than long, EL
<quantity id="F9569B2DFF83FFE1FEA1D7A3FEBC6698" box="[271,381,1671,1695]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="451" unit="mm" value="0.48">0.48 mm</quantity>
, EW
<quantity id="F9569B2DFF83FFE1FE65D7A2FDF96698" box="[459,568,1670,1695]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="8" pageNumber="451" unit="mm" value="0.64">0.64 mm</quantity>
, each elytron with two shallow foveae at base. Protrochanters strongly protuberant at ventral margin (
<figureCitation id="A6952A4DFF83FFE1FE41D78EFD8066C4" box="[495,577,1706,1731]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="8" pageNumber="451">Fig. 14</figureCitation>
Q), mesotarsomeres II expanded at apex. Abdomen slightly wider than long, AL
<quantity id="F9569B2DFF83FFE1FF6BD7EAFEEC66E0" box="[197,301,1742,1767]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="8" pageNumber="451" unit="mm" value="0.53">0.53 mm</quantity>
, AW
<quantity id="F9569B2DFF83FFE1FEDED7EAFE1966E0" box="[368,472,1742,1767]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="451" unit="mm" value="0.6">0.60 mm</quantity>
, slightly narrowed at base; tergite IV (first visible tergite) with narrow basal sulcus; discal carinae long, moderately close, extending to half of tergal length, width between carinae
<quantity id="F9569B2DFF83FFE1FB66D7D6FAED670C" box="[1224,1324,1778,1802]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="451" unit="mm" value="0.2">0.20 mm</quantity>
at apices; marginal carinae extending to half of tergal length. Aedeagus (
<figureCitation id="A6952A4DFF83FFE1FCFBD632FC636729" box="[853,930,1814,1839]" captionStart="FIGURE 11" captionStartId="17.[151,250,1967,1989]" captionTargetBox="[190,1403,193,1944]" captionTargetId="figure@17.[178,1407,193,1953]" captionTargetPageId="17" captionText="FIGURE 11. Aedeagi of Batraxis species. (A, B, D, E, G, H. in ventral view. C, F, I. in lateral view). A – C. B. longicornis. D – F. B. gesar. G – I. B. longithorax. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256552/files/figure.png" pageId="8" pageNumber="451">Fig. 11</figureCitation>
G–I)
<quantity id="F9569B2DFF83FFE1FC4ED633FB846728" box="[992,1093,1815,1839]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="451" unit="mm" value="0.31">0.31 mm</quantity>
in length, elongate, parameres slightly exceeding apex of median lobe, rounded apically, each with two apical setae; endophallus with several weakly sclerotized and nearly symmetric sclerites.
</paragraph>
<paragraph id="3E1136C8FF83FFE1FF69D6A6FE5F679D" blockId="8.[151,1437,1125,2019]" box="[199,414,1922,1947]" pageId="8" pageNumber="451">Female. Unknown.</paragraph>
<paragraph id="3E1136C8FF83FFE1FF69D682FB2367E4" blockId="8.[151,1437,1125,2019]" pageId="8" pageNumber="451">
<emphasis id="0CDAEADAFF83FFE1FF69D682FE7167B9" bold="true" box="[199,432,1958,1983]" pageId="8" pageNumber="451">Comparative notes.</emphasis>
<taxonomicName id="F9AE4D4BFF83FFE1FE19D683FD6167B8" box="[439,672,1958,1982]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="451" phylum="Arthropoda" rank="species" species="longithorax">
<emphasis id="0CDAEADAFF83FFE1FE19D683FD6167B8" box="[439,672,1958,1982]" italics="true" pageId="8" pageNumber="451">Batraxis longithorax</emphasis>
</taxonomicName>
can be readily separated from all other congeners by several unique characters, i.e. the relatively elongate pronotum, thick antennae, and expanded mesotarsomeres II.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF82FFE0FF69D1B3FD4860B6" box="[199,649,151,176]" pageId="9" pageNumber="452" type="distribution">
<paragraph id="3E1136C8FF82FFE0FF69D1B3FD4860B6" blockId="9.[199,927,151,213]" box="[199,649,151,176]" pageId="9" pageNumber="452">
<emphasis id="0CDAEADAFF82FFE0FF69D1B3FEA160B6" bold="true" box="[199,352,151,176]" pageId="9" pageNumber="452">Distribution.</emphasis>
<collectingCountry id="46B97658FF82FFE0FEC9D1B3FE6F60B6" box="[359,430,151,176]" name="China" pageId="9" pageNumber="452">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF82FFE0FE17D1B3FDD860B6" box="[441,537,151,176]" country="China" name="Guangxi" pageId="9" pageNumber="452">Guangxi</collectingRegion>
(
<figureCitation id="A6952A4DFF82FFE0FD86D1B3FDBD60B6" box="[552,636,151,176]" captionStart="FIGURE 15" captionStartId="21.[151,250,1023,1045]" captionTargetBox="[151,1436,193,1001]" captionTargetId="figure@21.[151,1436,193,1002]" captionTargetPageId="21" captionText="FIGURE 15. Distribution of Batraxis species in China." httpUri="https://zenodo.org/record/256556/files/figure.png" pageId="9" pageNumber="452">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF82FFE0FF69D198FC5E60D2" box="[199,927,188,213]" pageId="9" pageNumber="452" type="etymology">
<paragraph id="3E1136C8FF82FFE0FF69D198FC5E60D2" blockId="9.[199,927,151,213]" box="[199,927,188,213]" pageId="9" pageNumber="452">
<emphasis id="0CDAEADAFF82FFE0FF69D198FE8F60D3" bold="true" box="[199,334,188,213]" pageId="9" pageNumber="452">Etymology.</emphasis>
The specific epithet refers to the elongate pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>

288
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<mods:identifier id="0F9308DCE5A3E1A26AC29BAD0F1C5AD8" type="ZooBank">49EE6E10-C40B-4988-A086-CEE330544A0B</mods:identifier>
</mods:mods>
<treatment id="B60787DEFF87FFE4FF39D73FFC2D6266" ID-DOI="http://doi.org/10.5281/zenodo.6077887" ID-GBIF-Taxon="123514485" ID-Zenodo-Dep="6077887" LSID="urn:lsid:plazi:treatment:B60787DEFF87FFE4FF39D73FFC2D6266" httpUri="http://treatment.plazi.org/id/B60787DEFF87FFE4FF39D73FFC2D6266" lastPageId="13" lastPageNumber="456" pageId="12" pageNumber="455">
<subSubSection id="76B46543FF87FFE5FF39D73FFDC56650" pageId="12" pageNumber="455" type="nomenclature">
<paragraph id="3E1136C8FF87FFE5FF39D73FFD9F6633" blockId="12.[151,606,1563,1622]" box="[151,606,1563,1589]" pageId="12" pageNumber="455">
<heading id="655981A4FF87FFE5FF39D73FFD9F6633" bold="true" box="[151,606,1563,1589]" fontSize="11" level="1" pageId="12" pageNumber="455" reason="1">
<taxonomicName id="F9AE4D4BFF87FFE5FF39D73FFD9F6633" ID-CoL="L27Z" authority="Wang and Yin, 2016" authorityName="Wang and Yin" authorityYear="2016" box="[151,606,1563,1589]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="455" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="0CDAEADAFF87FFE5FF39D73FFD9F6633" bold="true" box="[151,606,1563,1589]" pageId="12" pageNumber="455">
<emphasis id="0CDAEADAFF87FFE5FF39D73FFE9B6633" bold="true" box="[151,346,1563,1589]" italics="true" pageId="12" pageNumber="455">Batraxis bicolor</emphasis>
Wang and Yin, 2016
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="3E1136C8FF87FFE5FF39D719FDC56650" blockId="12.[151,606,1563,1622]" box="[151,516,1597,1622]" pageId="12" pageNumber="455">
(
<figureCitation id="A6952A4DFF87FFE5FF31D719FF276650" box="[159,230,1597,1622]" captionStart="FIGURE 6" captionStartId="12.[151,250,1460,1482]" captionTargetBox="[157,1423,209,1419]" captionTargetId="figure@12.[151,1436,195,1439]" captionTargetPageId="12" captionText="FIGURE 6. Dorsal habitus of Batraxis males. A. B. ziyueae. B. B. bicolor, population from Yunnan. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256547/files/figure.png" pageId="12" pageNumber="455">Figs 6</figureCitation>
B, 13A–C, 14E, 14U, 15)
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF87FFE5FF39D7A3FD11669A" box="[151,720,1670,1692]" pageId="12" pageNumber="455" type="reference_group">
<paragraph id="3E1136C8FF87FFE5FF39D7A3FD11669A" blockId="12.[151,1113,1670,1722]" box="[151,720,1670,1692]" pageId="12" pageNumber="455">
<taxonomicName id="F9AE4D4BFF87FFE5FF39D7A3FD0D669A" ID-CoL="L27Z" authority="Wang and Yin (in Wang et al. 2016: 2)" authorityName="Wang and Yin (in Wang et al." authorityPageNumber="2" authorityYear="2016" box="[151,716,1670,1692]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="455" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="0CDAEADAFF87FFE5FF39D7A3FEFD669D" box="[151,316,1670,1692]" italics="true" pageId="12" pageNumber="455">Batraxis bicolor</emphasis>
Wang and Yin (in Wang
<emphasis id="0CDAEADAFF87FFE5FD92D7A3FDA9669D" box="[572,616,1670,1692]" italics="true" pageId="12" pageNumber="455">et al</emphasis>
. 2016: 2)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF87FFE5FF39D781FC0C6795" pageId="12" pageNumber="455" type="materials_examined">
<paragraph id="3E1136C8FF87FFE5FF39D781FB9866BC" blockId="12.[151,1113,1670,1722]" box="[151,1113,1700,1722]" pageId="12" pageNumber="455">
<materialsCitation id="8EC63C95FF87FFE5FF39D781FB9866BC" ID-GBIF-Occurrence="1290996381" box="[151,1113,1700,1722]" country="China" elevation="625" latitude="19.167778" location="Limu Mt." longLatPrecision="21" longitude="109.74583" municipality="Qiongzhong Hsien" pageId="12" pageNumber="455" specimenCount="625" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="E115886AFF87FFE5FF39D781FF0866BC" box="[151,201,1701,1722]" pageId="12" pageNumber="455">Type</typeStatus>
locality:
<collectingCountry id="46B97658FF87FFE5FE87D780FEA966BC" box="[297,360,1700,1722]" name="China" pageId="12" pageNumber="455">China</collectingCountry>
,
<collectingRegion id="FC6AF82AFF87FFE5FEDCD780FE7F66BC" box="[370,446,1700,1722]" country="China" name="Hainan" pageId="12" pageNumber="455">Hainan</collectingRegion>
,
<collectingMunicipality id="DE75ACB2FF87FFE5FE66D780FD4866BC" box="[456,649,1700,1722]" pageId="12" pageNumber="455">Qiongzhong Hsien</collectingMunicipality>
,
<location id="3B716013FF87FFE5FD3CD780FD3766BC" LSID="urn:lsid:plazi:treatment:B60787DEFF87FFE4FF39D73FFC2D6266:3B716013FF87FFE5FD3CD780FD3766BC" box="[658,758,1700,1722]" country="China" latitude="19.167778" longLatPrecision="21" longitude="109.74583" municipality="Qiongzhong Hsien" name="Limu Mt." pageId="12" pageNumber="455" stateProvince="Hainan">Limu Mt.</location>
,
<geoCoordinate id="5B9A500FFF87FFE5FCAED780FCBB66BC" box="[768,890,1700,1722]" direction="north" orientation="latitude" pageId="12" pageNumber="455" precision="15" value="19.167778">19°10'04&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF87FFE5FC2AD780FBC666BC" box="[900,1031,1700,1722]" direction="east" orientation="longitude" pageId="12" pageNumber="455" precision="15" value="109.74583">109°44'45&quot;E</geoCoordinate>
,
<specimenCount id="28A8FD41FF87FFE5FBBCD780FB9866BC" box="[1042,1113,1700,1722]" pageId="12" pageNumber="455" type="generic">
<quantity id="F9569B2DFF87FFE5FBBCD780FB9466BC" box="[1042,1109,1700,1722]" metricMagnitude="2" metricUnit="m" metricValue="6.25" pageId="12" pageNumber="455" unit="m" value="625.0">
<elevation id="B583D1FBFF87FFE5FBBCD780FB9466BC" box="[1042,1109,1700,1722]" metricMagnitude="2" metricUnit="m" metricValue="6.25" pageId="12" pageNumber="455" unit="m" value="625.0">625 m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="3E1136C8FF87FFE5FF39D7C3FC0C6795" blockId="12.[151,1436,1767,2012]" pageId="12" pageNumber="455">
<materialsCitation id="8EC63C95FF87FFE5FF39D7C3FE2A674D" ID-GBIF-Occurrence="1290996386" collectingDate="2016-04-09" collectionCode="SNUC" collectorName="Zi-Chun Xiong" country="China" county="Lincang City" elevation="1371" latitude="24.708612" location="Manwan" longLatPrecision="21" longitude="100.37889" municipality="Yun Hsien" pageId="12" pageNumber="455" specimenCount="17" specimenCount-female="17" stateProvince="Yunnan Province">
<emphasis id="0CDAEADAFF87FFE5FF39D7C3FDC26706" bold="true" box="[151,515,1767,1792]" pageId="12" pageNumber="455">Additional material examined</emphasis>
(14 ƋƋ,
<specimenCount id="28A8FD41FF87FFE5FDD4D7CCFD096706" box="[634,712,1768,1792]" pageId="12" pageNumber="455" type="female">17 ♀♀</specimenCount>
). 1 Ƌ, labeled ‘
<collectingCountry id="46B97658FF87FFE5FC2DD7CCFC246706" box="[899,997,1768,1792]" name="China" pageId="12" pageNumber="455">CHINA</collectingCountry>
:
<collectingRegion id="FC6AF82AFF87FFE5FC5DD7CCFB7B6706" box="[1011,1210,1767,1792]" country="China" name="Yunnan" pageId="12" pageNumber="455">Yunnan Province</collectingRegion>
,
<collectingCounty id="D7704E44FF87FFE5FB66D7C3FAA16706" box="[1224,1376,1767,1792]" pageId="12" pageNumber="455">Lincang City</collectingCounty>
,
<collectingMunicipality id="DE75ACB2FF87FFE5FAC1D7CCFF1A672E" pageId="12" pageNumber="455">Yun Hsien</collectingMunicipality>
,
<location id="3B716013FF87FFE5FF49D634FE8D6721" LSID="urn:lsid:plazi:treatment:B60787DEFF87FFE4FF39D73FFC2D6266:3B716013FF87FFE5FF49D634FE8D6721" box="[231,332,1808,1831]" country="China" county="Lincang City" latitude="24.708612" longLatPrecision="21" longitude="100.37889" municipality="Yun Hsien" name="Manwan" pageId="12" pageNumber="455" stateProvince="Yunnan Province">Manwan</location>
,
<location id="3B716013FF87FFE5FEF9D634FE586721" LSID="urn:lsid:plazi:treatment:B60787DEFF87FFE4FF39D73FFC2D6266:3B716013FF87FFE5FEF9D634FE586721" box="[343,409,1808,1831]" country="China" county="Lincang City" latitude="24.708612" longLatPrecision="21" longitude="100.37889" municipality="Yun Hsien" name="Town" pageId="12" pageNumber="455" stateProvince="Yunnan Province">Town</location>
,
<location id="3B716013FF87FFE5FE08D62BFD90672E" LSID="urn:lsid:plazi:treatment:B60787DEFF87FFE4FF39D73FFC2D6266:3B716013FF87FFE5FE08D62BFD90672E" box="[422,593,1807,1832]" country="China" county="Lincang City" latitude="24.708612" longLatPrecision="21" longitude="100.37889" municipality="Yun Hsien" name="Manjiu Village" pageId="12" pageNumber="455" stateProvince="Yunnan Province">Manjiu Village</location>
(慢旧村),
<geoCoordinate id="5B9A500FFF87FFE5FD75D62BFCA16721" box="[731,864,1807,1831]" direction="north" orientation="latitude" pageId="12" pageNumber="455" precision="15" value="24.708612">24°42'31''N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF87FFE5FCC2D62BFC3C6721" box="[876,1021,1807,1831]" direction="east" orientation="longitude" pageId="12" pageNumber="455" precision="15" value="100.37889">100°22'44''E</geoCoordinate>
, rotten wood, alt.
<quantity id="F9569B2DFF87FFE5FB67D62BFAE06721" box="[1225,1313,1807,1831]" metricMagnitude="3" metricUnit="m" metricValue="1.371" pageId="12" pageNumber="455" unit="m" value="1371.0">
<elevation id="B583D1FBFF87FFE5FB67D62BFAE06721" box="[1225,1313,1807,1831]" metricMagnitude="3" metricUnit="m" metricValue="1.371" pageId="12" pageNumber="455" unit="m" value="1371.0">1371 m</elevation>
</quantity>
,
<date id="4A101008FF87FFE5FA83D62BFA56672E" box="[1325,1431,1807,1832]" pageId="12" pageNumber="455" value="2016-04-09">
<collectingDate id="5A54E9E0FF87FFE5FA83D62BFA56672E" box="[1325,1431,1807,1832]" pageId="12" pageNumber="455" value="2016-04-09">9.iv.2016</collectingDate>
</date>
,
<collectorName id="935B531EFF87FFE5FF39D617FE84674A" box="[151,325,1843,1868]" pageId="12" pageNumber="455">Zi-Chun Xiong</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF87FFE5FE3DD617FE25674D" box="[403,484,1843,1867]" pageId="12" pageNumber="455">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF87FFE5FE54D617FA436769" ID-GBIF-Occurrence="1290996367" collectingDate="2016-04-19" collectionCode="SNUC" collectorName="Zi-Chun Xiong" country="China" county="Lincang City" elevation="1243" latitude="24.730001" location="Manwan Town" longLatPrecision="21" longitude="100.33361" municipality="Yun Hsien" pageId="12" pageNumber="455" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan">
5 ƋƋ,
<specimenCount id="28A8FD41FF87FFE5FDE2D617FD4E674D" box="[588,655,1843,1867]" pageId="12" pageNumber="455" type="female">3 ♀♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF87FFE5FD50D617FC8B674A" box="[766,842,1843,1868]" name="China" pageId="12" pageNumber="455">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF87FFE5FCF9D610FC71674D" box="[855,944,1844,1867]" country="China" name="Yunnan" pageId="12" pageNumber="455">Yunnan</collectingRegion>
,
<collectingCounty id="D7704E44FF87FFE5FC13D617FB92674A" box="[957,1107,1843,1868]" pageId="12" pageNumber="455">Lincang City</collectingCounty>
,
<collectingMunicipality id="DE75ACB2FF87FFE5FBCCD610FB1D674A" box="[1122,1244,1843,1868]" pageId="12" pageNumber="455">Yun Hsien</collectingMunicipality>
,
<location id="3B716013FF87FFE5FB44D610FA59674D" LSID="urn:lsid:plazi:treatment:B60787DEFF87FFE4FF39D73FFC2D6266:3B716013FF87FFE5FB44D610FA59674D" box="[1258,1432,1844,1867]" country="China" county="Lincang City" latitude="24.730001" longLatPrecision="21" longitude="100.33361" municipality="Yun Hsien" name="Manwan Town" pageId="12" pageNumber="455" stateProvince="Yunnan">Manwan Town</location>
,
<location id="3B716013FF87FFE5FF39D673FE8A6776" LSID="urn:lsid:plazi:treatment:B60787DEFF87FFE4FF39D73FFC2D6266:3B716013FF87FFE5FF39D673FE8A6776" box="[151,331,1879,1904]" country="China" county="Lincang City" latitude="24.730001" longLatPrecision="21" longitude="100.33361" municipality="Yun Hsien" name="Guiziba Village" pageId="12" pageNumber="455" stateProvince="Yunnan">Guiziba Village</location>
,
<geoCoordinate id="5B9A500FFF87FFE5FEF9D673FE1C6769" box="[343,477,1879,1903]" direction="north" orientation="latitude" pageId="12" pageNumber="455" precision="15" value="24.730001">24°43'48&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF87FFE5FE44D673FDBA6769" box="[490,635,1879,1903]" direction="east" orientation="longitude" pageId="12" pageNumber="455" precision="15" value="100.33361">100°20'01&quot;E</geoCoordinate>
, rotten wood, alt.
<quantity id="F9569B2DFF87FFE5FCE6D673FC606769" box="[840,929,1879,1903]" metricMagnitude="3" metricUnit="m" metricValue="1.2429999999999999" pageId="12" pageNumber="455" unit="m" value="1243.0">
<elevation id="B583D1FBFF87FFE5FCE6D673FC606769" box="[840,929,1879,1903]" metricMagnitude="3" metricUnit="m" metricValue="1.2429999999999999" pageId="12" pageNumber="455" unit="m" value="1243.0">1243 m</elevation>
</quantity>
,
<date id="4A101008FF87FFE5FC03D673FBE46776" box="[941,1061,1879,1904]" pageId="12" pageNumber="455" value="2016-04-19">
<collectingDate id="5A54E9E0FF87FFE5FC03D673FBE46776" box="[941,1061,1879,1904]" pageId="12" pageNumber="455" value="2016-04-19">19.iv.2016</collectingDate>
</date>
,
<collectorName id="935B531EFF87FFE5FB9CD673FB1E6776" box="[1074,1247,1879,1904]" pageId="12" pageNumber="455">Zi-Chun Xiong</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF87FFE5FA87D673FABB6769" box="[1321,1402,1879,1903]" pageId="12" pageNumber="455">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF87FFE5FA20D673FC086795" ID-GBIF-Occurrence="1290996383" collectingDate="2016-04-20" collectionCode="SNUC" collectorName="Zi-Chun Xiong" country="China" county="Lincang City" elevation="1243" latitude="24.730001" location="Manwan Town" longLatPrecision="21" longitude="100.33361" municipality="Yun Hsien" pageId="12" pageNumber="455" specimenCount="14" specimenCount-female="14" stateProvince="Yunnan">
8 ƋƋ,
<specimenCount id="28A8FD41FF87FFE5FF61D65FFEDC6795" box="[207,285,1915,1939]" pageId="12" pageNumber="455" type="female">14 ♀♀</specimenCount>
, same data as previous material except ‘
<date id="4A101008FF87FFE5FD73D65FFC9D6792" box="[733,860,1915,1940]" pageId="12" pageNumber="455" value="2016-04-20">
<collectingDate id="5A54E9E0FF87FFE5FD73D65FFC9D6792" box="[733,860,1915,1940]" pageId="12" pageNumber="455" value="2016-04-20">20.iv.2016</collectingDate>
</date>
’ (
<collectionCode id="58BFAE0DFF87FFE5FCDED65FFC006795" box="[880,961,1915,1939]" pageId="12" pageNumber="455">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF87FFE4FF69D6BBFE0F623B" lastPageId="13" lastPageNumber="456" pageId="12" pageNumber="455" type="discussion">
<paragraph id="3E1136C8FF87FFE4FF69D6BBFDF06163" blockId="12.[151,1436,1767,2012]" lastBlockId="13.[151,1437,151,608]" lastPageId="13" lastPageNumber="456" pageId="12" pageNumber="455">
<emphasis id="0CDAEADAFF87FFE5FF69D6BBFE8E67BE" bold="true" box="[199,335,1951,1976]" pageId="12" pageNumber="455">Comments.</emphasis>
The population of
<taxonomicName id="F9AE4D4BFF87FFE5FD81D684FD2967B1" box="[559,744,1951,1975]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="455" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="0CDAEADAFF87FFE5FD81D684FD2967B1" box="[559,744,1951,1975]" italics="true" pageId="12" pageNumber="455">Batraxis bicolor</emphasis>
</taxonomicName>
from
<collectingRegion id="FC6AF82AFF87FFE5FC9CD6BBFC4267BE" box="[818,899,1951,1976]" country="China" name="Hainan" pageId="12" pageNumber="455">Hainan</collectingRegion>
has reddish brown head, pronotum, and elytra and a much darker abdomen (Wang
<emphasis id="0CDAEADAFF87FFE5FD98D6E0FDA967DD" box="[566,616,1987,2011]" italics="true" pageId="12" pageNumber="455">et al</emphasis>
. 2016) which is stable in all available individuals, while the population from
<collectingRegion id="FC6AF82AFF86FFE4FF74D1BCFEEE60A9" box="[218,303,152,175]" country="China" name="Yunnan" pageId="13" pageNumber="456">Yunnan</collectingRegion>
(
<figureCitation id="A6952A4DFF86FFE4FEEBD1B3FE4C60B6" box="[325,397,151,176]" captionStart="FIGURE 6" captionStartId="12.[151,250,1460,1482]" captionTargetBox="[157,1423,209,1419]" captionTargetId="figure@12.[151,1436,195,1439]" captionTargetPageId="12" captionText="FIGURE 6. Dorsal habitus of Batraxis males. A. B. ziyueae. B. B. bicolor, population from Yunnan. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256547/files/figure.png" pageId="13" pageNumber="456">Fig. 6</figureCitation>
B) are uniform in color (darkish brown). Furthermore, the aedeagal endophallus of the population from
<collectingRegion id="FC6AF82AFF86FFE4FEF9D198FE6960D3" box="[343,424,188,213]" country="China" name="Hainan" pageId="13" pageNumber="456">Hainan</collectingRegion>
is strictly symmetric, but in the population from
<collectingRegion id="FC6AF82AFF86FFE4FC4DD199FBFD60D2" box="[995,1084,189,212]" country="China" name="Yunnan" pageId="13" pageNumber="456">Yunnan</collectingRegion>
, it is slightly asymmetric (
<figureCitation id="A6952A4DFF86FFE4FAC1D198FF7360FE" captionStart="FIGURE 13" captionStartId="19.[151,250,1495,1517]" captionTargetBox="[185,1406,200,1454]" captionTargetId="figure@19.[174,1410,193,1463]" captionTargetPageId="19" captionText="FIGURE 13. Aedeagi of Batraxis species. (A, B, D, E. in ventral view. C, F. in lateral view). A – C. B. bicolor, population from Yunnan. D – F. B. tibialis, population from Jiangxi. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256554/files/figure.png" pageId="13" pageNumber="456">Fig. 13</figureCitation>
A–C). The above differences are interpreted as intraspecific variation of this species. Otherwise, the long protuberance of the protrochanters (
<figureCitation id="A6952A4DFF86FFE4FD9CD020FD45611A" box="[562,644,260,285]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="13" pageNumber="456">Fig. 14</figureCitation>
U) and small preapical tooth of the protibiae (
<figureCitation id="A6952A4DFF86FFE4FB37D020FB2D611A" box="[1177,1260,260,285]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="13" pageNumber="456">Fig. 14</figureCitation>
E) are stable in both populations.
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<emphasis id="0CDAEADAFF86FFE4FECCD00CFDDB6146" box="[354,538,296,320]" italics="true" pageId="13" pageNumber="456">Batraxis bicolor</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="F9AE4D4BFF86FFE4FD1ED00DFC606139" authority="Nomura" authorityName="Nomura" box="[688,929,296,320]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="456" phylum="Arthropoda" rank="species" species="splendida">
<emphasis id="0CDAEADAFF86FFE4FD1ED00DFD006146" box="[688,705,297,320]" italics="true" pageId="13" pageNumber="456">B</emphasis>
.
<emphasis id="0CDAEADAFF86FFE4FD7FD00CFCFF6146" box="[721,830,296,320]" italics="true" pageId="13" pageNumber="456">splendida</emphasis>
Nomura
</taxonomicName>
from
<collectingCountry id="46B97658FF86FFE4FC46D00CFBE96139" box="[1000,1064,296,319]" name="Japan" pageId="13" pageNumber="456">Japan</collectingCountry>
and
<collectingCountry id="46B97658FF86FFE4FBCED00CFB666139" box="[1120,1191,296,319]" name="South Korea" pageId="13" pageNumber="456">Korea</collectingCountry>
, but the latter species lack a protuberance on the protibiae.
</paragraph>
<paragraph id="3E1136C8FF86FFE4FF69D054FE0F623B" blockId="13.[151,1437,151,608]" pageId="13" pageNumber="456">
Measurements of the population from
<collectingRegion id="FC6AF82AFF86FFE4FDD2D054FD176181" box="[636,726,368,391]" country="China" name="Yunnan" pageId="13" pageNumber="456">Yunnan</collectingRegion>
: male—BL 2.00–
<quantity id="F9569B2DFF86FFE4FC0ED054FBC66181" box="[928,1031,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="2.17" pageId="13" pageNumber="456" unit="mm" value="2.17">2.17 mm</quantity>
, HL
<quantity id="F9569B2DFF86FFE4FBECD054FB296181" box="[1090,1256,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.3" metricValueMin="4.2" pageId="13" pageNumber="456" unit="mm" value="0.425" valueMax="0.43" valueMin="0.42">0.42–0.43 mm</quantity>
, HW
<quantity id="F9569B2DFF86FFE4FA82D054FF0461AA" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.3" metricValueMin="4.2" pageId="13" pageNumber="456" unit="mm" value="0.425" valueMax="0.43" valueMin="0.42">0.42–0.43 mm</quantity>
, PL
<quantity id="F9569B2DFF86FFE4FF57D0B0FE5F61AA" box="[249,414,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="13" pageNumber="456" unit="mm" value="0.405" valueMax="0.42" valueMin="0.39">0.39–0.42 mm</quantity>
, PW
<quantity id="F9569B2DFF86FFE4FE75D0B0FD4061AA" box="[475,641,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.699999999999999" metricValueMin="4.3" pageId="13" pageNumber="456" unit="mm" value="0.44999999999999996" valueMax="0.47" valueMin="0.43">0.43–0.47 mm</quantity>
, EL
<quantity id="F9569B2DFF86FFE4FD18D0B0FC9D61AA" box="[694,860,404,429]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.4" metricValueMin="5.0" pageId="13" pageNumber="456" unit="mm" value="0.52" valueMax="0.54" valueMin="0.5">0.50–0.54 mm</quantity>
, EW
<quantity id="F9569B2DFF86FFE4FC34D0B0FB8161AA" box="[922,1088,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="8.35" metricValueMax="8.4" metricValueMin="8.299999999999999" pageId="13" pageNumber="456" unit="mm" value="0.835" valueMax="0.84" valueMin="0.83">0.83–0.84 mm</quantity>
, AL
<quantity id="F9569B2DFF86FFE4FBD6D0B0FADF61AA" box="[1144,1310,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="7.9" metricValueMin="7.1" pageId="13" pageNumber="456" unit="mm" value="0.75" valueMax="0.79" valueMin="0.71">0.71–0.79 mm</quantity>
, AW
<quantity id="F9569B2DFF86FFE4FAF3D0B0FEC161C9" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.5" metricValueMin="7.8" pageId="13" pageNumber="456" unit="mm" value="0.815" valueMax="0.85" valueMin="0.78">0.78– 0.85 mm</quantity>
, width between discal carinae of tergite IV
<quantity id="F9569B2DFF86FFE4FD53D09CFC6261C9" box="[765,931,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.4" metricValueMin="3.8" pageId="13" pageNumber="456" unit="mm" value="0.41" valueMax="0.44" valueMin="0.38">0.38–0.44 mm</quantity>
at apices, length of the aedeagus
<quantity id="F9569B2DFF86FFE4FA80D09CFA5961C9" box="[1326,1432,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="13" pageNumber="456" unit="mm" value="0.49">0.49 mm</quantity>
; female—BL
<quantity id="F9569B2DFF86FFE4FE82D0F8FE1561F2" box="[300,468,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.94" metricValueMax="2.05" metricValueMin="1.83" pageId="13" pageNumber="456" unit="mm" value="1.94" valueMax="2.05" valueMin="1.83">1.83–2.05 mm</quantity>
; HL
<quantity id="F9569B2DFF86FFE4FDBED0F8FD7961F2" box="[528,696,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" metricValueMax="4.0" metricValueMin="3.8" pageId="13" pageNumber="456" unit="mm" value="0.39" valueMax="0.4" valueMin="0.38">0.38–0.40 mm</quantity>
; HW
<quantity id="F9569B2DFF86FFE4FD53D0F8FC6461F2" box="[765,933,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.0" metricValueMin="3.9" pageId="13" pageNumber="456" unit="mm" value="0.395" valueMax="0.4" valueMin="0.39">0.39–0.40 mm</quantity>
; PL
<quantity id="F9569B2DFF86FFE4FC72D0F8FB4561F2" box="[988,1156,476,501]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="3.9" metricValueMin="3.5" pageId="13" pageNumber="456" unit="mm" value="0.37" valueMax="0.39" valueMin="0.35">0.35–0.39 mm</quantity>
; PW
<quantity id="F9569B2DFF86FFE4FB6AD0F8FAAD61F2" box="[1220,1388,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" pageId="13" pageNumber="456" unit="mm" value="0.43" valueMax="0.44" valueMin="0.42">0.42–0.44 mm</quantity>
; EL
<quantity id="F9569B2DFF86FFE4FF39D0DBFEFD6211" box="[151,316,511,536]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="4.9" metricValueMin="4.5" pageId="13" pageNumber="456" unit="mm" value="0.47" valueMax="0.49" valueMin="0.45">0.45–0.49 mm</quantity>
; EW
<quantity id="F9569B2DFF86FFE4FED4D324FDE16211" box="[378,544,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="8.1" metricValueMin="7.4" pageId="13" pageNumber="456" unit="mm" value="0.775" valueMax="0.81" valueMin="0.74">0.74–0.81 mm</quantity>
; AL
<quantity id="F9569B2DFF86FFE4FDF9D0DBFD3C6211" box="[599,765,511,536]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" metricValueMax="7.7" metricValueMin="6.5" pageId="13" pageNumber="456" unit="mm" value="0.71" valueMax="0.77" valueMin="0.65">0.65–0.77 mm</quantity>
; AW
<quantity id="F9569B2DFF86FFE4FC95D324FC206211" box="[827,993,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" metricValueMax="8.2" metricValueMin="7.4" pageId="13" pageNumber="456" unit="mm" value="0.78" valueMax="0.82" valueMin="0.74">0.74–0.82 mm</quantity>
; width between discal carinae of tergite IV
<quantity id="F9569B2DFF86FFE4FF15D300FE9F623A" box="[187,350,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="13" pageNumber="456" unit="mm" value="0.4" valueMax="0.42" valueMin="0.38">0.38–0.42 mm</quantity>
at apices.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF86FFE4FF69D363FC2D6266" box="[199,1004,583,608]" pageId="13" pageNumber="456" type="distribution">
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<emphasis id="0CDAEADAFF86FFE4FF69D363FEA16266" bold="true" box="[199,352,583,608]" pageId="13" pageNumber="456">Distribution.</emphasis>
<collectingCountry id="46B97658FF86FFE4FEC9D363FE6F6266" box="[359,430,583,608]" name="China" pageId="13" pageNumber="456">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF86FFE4FE17D36CFDCF6259" box="[441,526,584,607]" country="China" name="Yunnan" pageId="13" pageNumber="456">Yunnan</collectingRegion>
(new provincial record),
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(
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).
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655
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@ -0,0 +1,655 @@
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(
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B, 9, 10A‒F, 14A‒B, 14J‒N, 15)
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material
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(29 ƋƋ,
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:
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:
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Ƌ, labeled ‘
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<collectingRegion id="FC6AF82AFF89FFEBFBA5D7FFFB9966F4" box="[1035,1112,1755,1778]" country="China" name="Hunan" pageId="2" pageNumber="445">Hunan</collectingRegion>
,
<collectingCounty id="D7704E44FF89FFEBFBC9D7FEFB3E66F5" box="[1127,1279,1754,1779]" pageId="2" pageNumber="445">Liuyang City</collectingCounty>
(浏阳市),
<collectingMunicipality id="DE75ACB2FF89FFEBFF39D626FE98671C" box="[151,345,1794,1819]" pageId="2" pageNumber="445">Daweishan Town</collectingMunicipality>
(大围山镇),
<location id="3B716013FF89FFEBFE52D626FD45671D" LSID="urn:lsid:plazi:treatment:B60787DEFF89FFEDFF39D74FFCB5611B:3B716013FF89FFEBFE52D626FD45671D" box="[508,644,1794,1819]" country="China" county="Liuyang City" latitude="28.426945" longLatPrecision="20" longitude="114.12862" municipality="Daweishan Town" name="Dawei Shan" pageId="2" pageNumber="445" stateProvince="Hunan">Dawei Shan</location>
(大围山),
<geoCoordinate id="5B9A500FFF89FFEBFCA2D626FC53671D" box="[780,914,1794,1819]" direction="north" orientation="latitude" pageId="2" pageNumber="445" precision="15" value="28.426945">28°25'37&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF89FFEBFC32D627FBEC671D" box="[924,1069,1795,1819]" direction="east" orientation="longitude" pageId="2" pageNumber="445" precision="15" value="114.12862">114°07'43&quot;E</geoCoordinate>
, mixed leaf litter, sifted,
<quantity id="F9569B2DFF89FFEBFAEFD627FA56671C" box="[1345,1431,1795,1819]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="2" pageNumber="445" unit="m" value="1440.0">
<elevation id="B583D1FBFF89FFEBFAEFD627FA56671C" box="[1345,1431,1795,1819]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="2" pageNumber="445" unit="m" value="1440.0">1440 m</elevation>
</quantity>
,
<date id="4A101008FF89FFEBFF39D602FED86739" box="[151,281,1830,1855]" pageId="2" pageNumber="445" value="2013-07-21">
<collectingDate id="5A54E9E0FF89FFEBFF39D602FED86739" box="[151,281,1830,1855]" pageId="2" pageNumber="445" value="2013-07-21">21.vii.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF89FFEBFE87D603FEA56739" box="[297,356,1831,1855]" pageId="2" pageNumber="445">Peng</collectorName>
,
<collectorName id="935B531EFF89FFEBFEDAD602FE5D6739" box="[372,412,1830,1855]" pageId="2" pageNumber="445">Dai</collectorName>
&amp;
<collectorName id="935B531EFF89FFEBFE67D602FE336739" box="[457,498,1830,1855]" pageId="2" pageNumber="445">Xie</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF89FFEBFDEDD603FD556739" box="[579,660,1831,1855]" pageId="2" pageNumber="445">SNUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8EC63C95FF89FFEBFD05D602FF366764" ID-GBIF-Occurrence="1290996379" collectionCode="SNUC" country="China" location="CHINA" pageId="2" pageNumber="445" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
<emphasis id="0CDAEADAFF89FFEBFD05D602FC5F6739" bold="true" box="[683,926,1830,1855]" pageId="2" pageNumber="445">
<typeStatus id="E115886AFF89FFEBFD05D602FCE96739" box="[683,808,1830,1855]" pageId="2" pageNumber="445" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="46B97658FF89FFEBFC99D602FC586739" box="[823,921,1830,1855]" name="China" pageId="2" pageNumber="445">CHINA</collectingCountry>
:
</emphasis>
4 ƋƋ,
<specimenCount id="28A8FD41FF89FFEBFBAED602FB856738" box="[1024,1092,1830,1855]" pageId="2" pageNumber="445" type="female">6 ♀♀</specimenCount>
, same label data as
<typeStatus id="E115886AFF89FFEBFA94D602FA5D6739" box="[1338,1436,1830,1855]" pageId="2" pageNumber="445" type="holotype">holotype</typeStatus>
(
<collectionCode id="58BFAE0DFF89FFEBFF31D66EFF316764" box="[159,240,1866,1890]" pageId="2" pageNumber="445">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF89FFEBFEAAD66EFC286764" ID-GBIF-Occurrence="1290996359" box="[260,1001,1866,1890]" collectingDate="2013-07-22" collectionCode="SNUC" country="China" location="CHINA" pageId="2" pageNumber="445" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="28A8FD41FF89FFEBFEAAD66EFEF16764" box="[260,304,1866,1890]" pageId="2" pageNumber="445" type="male">1 ♂</specimenCount>
, same data as previous material except ‘
<date id="4A101008FF89FFEBFD5DD66EFCBD6764" box="[755,892,1866,1890]" pageId="2" pageNumber="445" value="2013-07-22">
<collectingDate id="5A54E9E0FF89FFEBFD5DD66EFCBD6764" box="[755,892,1866,1890]" pageId="2" pageNumber="445" value="2013-07-22">22.vii.2013</collectingDate>
</date>
’ (
<collectionCode id="58BFAE0DFF89FFEBFC3ED66EFC206764" box="[912,993,1866,1890]" pageId="2" pageNumber="445">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF89FFEBFC5AD66EFD87678C" ID-GBIF-Occurrence="1290996368" collectingDate="2013-07-20" collectionCode="SNUC" country="China" elevation="1460" location="CHINA" pageId="2" pageNumber="445" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="28A8FD41FF89FFEBFC5AD66EFBF76764" box="[1012,1078,1866,1890]" pageId="2" pageNumber="445" type="male">2 ♂♂</specimenCount>
, same data as previous material except ‘
<quantity id="F9569B2DFF89FFEBFF42D656FE88678F" box="[236,329,1906,1930]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="445" unit="m" value="1460.0">
<elevation id="B583D1FBFF89FFEBFF42D656FE88678F" box="[236,329,1906,1930]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="2" pageNumber="445" unit="m" value="1460.0">1460 m</elevation>
</quantity>
,
<date id="4A101008FF89FFEBFEFBD656FE1B678C" box="[341,474,1906,1930]" pageId="2" pageNumber="445" value="2013-07-20">
<collectingDate id="5A54E9E0FF89FFEBFEFBD656FE1B678C" box="[341,474,1906,1930]" pageId="2" pageNumber="445" value="2013-07-20">20.vii.2013</collectingDate>
</date>
’ (
<collectionCode id="58BFAE0DFF89FFEBFE40D656FDFE678C" box="[494,575,1906,1930]" pageId="2" pageNumber="445">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF89FFEBFDFDD656FCD067D0" ID-GBIF-Occurrence="1290996382" collectingDate="2015-10-04" collectionCode="SNUC" collectorName="Yan &amp; Tu &amp; Shen &amp; Jiang &amp; Zhou" country="China" county="Wuyishan City" elevation="1242" latitude="27.733612" location="Guadun" longLatPrecision="20" longitude="117.63694" municipality="Tongmu Village" pageId="2" pageNumber="445" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Fujian Province" typeStatus="paratype">
<specimenCount id="28A8FD41FF89FFEBFDFDD656FD55678F" box="[595,660,1906,1930]" pageId="2" pageNumber="445" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="28A8FD41FF89FFEBFD0FD656FD0D678F" box="[673,716,1906,1930]" pageId="2" pageNumber="445" type="female">1 ♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF89FFEBFC96D656FC45678C" box="[824,900,1906,1930]" name="China" pageId="2" pageNumber="445">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF89FFEBFC21D656FB84678C" box="[911,1093,1906,1930]" country="China" name="Fujian" pageId="2" pageNumber="445">Fujian Province</collectingRegion>
,
<collectingCounty id="D7704E44FF89FFEBFBFED656FB39678C" box="[1104,1272,1906,1930]" pageId="2" pageNumber="445">Wuyishan City</collectingCounty>
(武夷山市),
<collectingMunicipality id="DE75ACB2FF89FFEBFF39D6BEFE8A67B4" box="[151,331,1945,1970]" pageId="2" pageNumber="445">Tongmu Village</collectingMunicipality>
(桐木村),
<location id="3B716013FF89FFEBFE7CD6BDFDEA67B4" LSID="urn:lsid:plazi:treatment:B60787DEFF89FFEDFF39D74FFCB5611B:3B716013FF89FFEBFE7CD6BDFDEA67B4" box="[466,555,1945,1970]" country="China" county="Wuyishan City" latitude="27.733612" longLatPrecision="20" longitude="117.63694" municipality="Tongmu Village" name="Guadun" pageId="2" pageNumber="445" stateProvince="Fujian Province">Guadun</location>
(挂墩),
<geoCoordinate id="5B9A500FFF89FFEBFD38D6BEFCDC67B4" box="[662,797,1946,1970]" direction="north" orientation="latitude" pageId="2" pageNumber="445" precision="15" value="27.733612">27°44'01&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF89FFEBFC89D6BEFC7967B4" box="[807,952,1946,1970]" direction="east" orientation="longitude" pageId="2" pageNumber="445" precision="15" value="117.63694">117°38'13&quot;E</geoCoordinate>
, mixed forest, leaf litter, wood, sifted,
<quantity id="F9569B2DFF89FFEBFACAD6BEFF6E67D3" metricMagnitude="3" metricUnit="m" metricValue="1.242" pageId="2" pageNumber="445" unit="m" value="1242.0">
<elevation id="B583D1FBFF89FFEBFACAD6BEFF6E67D3" metricMagnitude="3" metricUnit="m" metricValue="1.242" pageId="2" pageNumber="445" unit="m" value="1242.0">1242 m</elevation>
</quantity>
,
<date id="4A101008FF89FFEBFF15D699FEDE67D0" box="[187,287,1981,2006]" pageId="2" pageNumber="445" value="2015-10-04">
<collectingDate id="5A54E9E0FF89FFEBFF15D699FEDE67D0" box="[187,287,1981,2006]" pageId="2" pageNumber="445" value="2015-10-04">4.x.2015</collectingDate>
</date>
,
<collectorName id="935B531EFF89FFEBFE84D69AFE9867D3" box="[298,345,1982,2005]" pageId="2" pageNumber="445">Yan</collectorName>
,
<collectorName id="935B531EFF89FFEBFECAD69AFE4467D3" box="[356,389,1982,2005]" pageId="2" pageNumber="445">Tu</collectorName>
,
<collectorName id="935B531EFF89FFEBFE3ED699FE0A67D0" box="[400,459,1981,2006]" pageId="2" pageNumber="445">Shen</collectorName>
,
<collectorName id="935B531EFF89FFEBFE78D699FDD067D0" box="[470,529,1981,2006]" pageId="2" pageNumber="445">Jiang</collectorName>
&amp;
<collectorName id="935B531EFF89FFEBFD9BD699FDB167D0" box="[565,624,1981,2006]" pageId="2" pageNumber="445">Zhou</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF89FFEBFD16D69AFCC867D0" box="[696,777,1982,2006]" pageId="2" pageNumber="445">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF89FFEAFCB2D69AFCD060DE" ID-GBIF-Occurrence="1290996363" collectingDate="2013-06-21" collectionCode="SNUC" collectorName="Xie &amp; Lv &amp; Yu" country="China" elevation="650" lastPageId="3" lastPageNumber="446" latitude="29.248333" location="Gutian Shan National Reserve" longLatPrecision="20" longitude="118.12056" municipality="Quzhou City" pageId="2" pageNumber="445" specimenCount="3" specimenCount-female="3" stateProvince="Zhejiang" typeStatus="paratype">
4 ƋƋ,
<specimenCount id="28A8FD41FF89FFEBFCC7D69AFC6867D3" box="[873,937,1982,2006]" pageId="2" pageNumber="445" type="female">3 ♀♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF89FFEBFBBCD699FB9F67D0" box="[1042,1118,1981,2006]" name="China" pageId="2" pageNumber="445">China</collectingCountry>
: N.
<collectingRegion id="FC6AF82AFF89FFEBFB25D699FB3367D0" box="[1163,1266,1981,2006]" country="China" name="Zhejiang" pageId="2" pageNumber="445">Zhejiang</collectingRegion>
,
<collectingMunicipality id="DE75ACB2FF89FFEBFB52D699FA4D67D0" box="[1276,1420,1981,2006]" pageId="2" pageNumber="445">Quzhou City</collectingMunicipality>
(衢州市),
<location id="3B716013FF88FFEAFEA6D1BFFD9860B5" LSID="urn:lsid:plazi:treatment:B60787DEFF89FFEDFF39D74FFCB5611B:3B716013FF88FFEAFEA6D1BFFD9860B5" box="[264,601,155,180]" country="China" latitude="29.248333" longLatPrecision="20" longitude="118.12056" municipality="Quzhou City" name="Gutian Shan National Reserve" pageId="3" pageNumber="446" stateProvince="Zhejiang">Gutian Shan National Reserve</location>
(古田山自然保护区),
<geoCoordinate id="5B9A500FFF88FFEAFCC3D1BFFC3260B2" box="[877,1011,155,180]" direction="north" orientation="latitude" pageId="3" pageNumber="446" precision="15" value="29.248333">29°14'54&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF88FFEAFC50D1B8FB4F60B2" box="[1022,1166,156,180]" direction="east" orientation="longitude" pageId="3" pageNumber="446" precision="15" value="118.12056">118°07'14&quot;E</geoCoordinate>
, mixed leaf litter, sifted,
<quantity id="F9569B2DFF88FFEAFF39D19BFED860D1" box="[151,281,191,215]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="446" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<elevation id="B583D1FBFF88FFEAFF39D19BFED860D1" box="[151,281,191,215]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="446" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500-800 m</elevation>
</quantity>
,
<date id="4A101008FF88FFEAFE82D19BFE6760D1" box="[300,422,191,215]" pageId="3" pageNumber="446" value="2013-06-21">
<collectingDate id="5A54E9E0FF88FFEAFE82D19BFE6760D1" box="[300,422,191,215]" pageId="3" pageNumber="446" value="2013-06-21">21.vi.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF88FFEAFE19D19BFE2360D1" box="[439,482,191,215]" pageId="3" pageNumber="446">Xie</collectorName>
,
<collectorName id="935B531EFF88FFEAFE5AD1E4FDD260D1" box="[500,531,192,215]" pageId="3" pageNumber="446">Lv</collectorName>
&amp;
<collectorName id="935B531EFF88FFEAFDEAD1E4FDA260D1" box="[580,611,192,215]" pageId="3" pageNumber="446">Yu</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF88FFEAFD16D1E4FCC860DE" box="[696,777,192,216]" pageId="3" pageNumber="446">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF88FFEAFC8CD19BFB7A60FA" ID-GBIF-Occurrence="1290996372" collectingDate="2013-06-19" collectionCode="SNUC" collectorName="Hu &amp; Song &amp; Lv &amp; Xie &amp; Yu" country="China" elevation="450" latitude="29.245834" location="Gutian Shan National Reserve" longLatPrecision="20" longitude="118.11444" municipality="Quzhou City" pageId="3" pageNumber="446" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Zhejiang" typeStatus="paratype">
<specimenCount id="28A8FD41FF88FFEAFC8CD19BFCA860D1" box="[802,873,191,215]" pageId="3" pageNumber="446" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="28A8FD41FF88FFEAFCD2D19BFC0360D1" box="[892,962,191,215]" pageId="3" pageNumber="446" type="female">6 ♀♀</specimenCount>
, same data as previous material except ‘
<geoCoordinate id="5B9A500FFF88FFEAFF35D1C7FEE660FA" box="[155,295,227,252]" direction="north" orientation="latitude" pageId="3" pageNumber="446" precision="15" value="29.245834">29°14'45&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF88FFEAFE98D1C7FE0560FA" box="[310,452,227,252]" direction="east" orientation="longitude" pageId="3" pageNumber="446" precision="15" value="118.11444">118°06'52&quot;E</geoCoordinate>
,
<quantity id="F9569B2DFF88FFEAFE7BD1C7FD9760FD" box="[469,598,227,251]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="3" pageNumber="446" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
<elevation id="B583D1FBFF88FFEAFE7BD1C7FD9760FD" box="[469,598,227,251]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="3" pageNumber="446" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400-500 m</elevation>
</quantity>
,
<date id="4A101008FF88FFEAFDCBD1C0FD2160FA" box="[613,736,228,252]" pageId="3" pageNumber="446" value="2013-06-19">
<collectingDate id="5A54E9E0FF88FFEAFDCBD1C0FD2160FA" box="[613,736,228,252]" pageId="3" pageNumber="446" value="2013-06-19">19.vi.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF88FFEAFD41D1C0FCD560FD" box="[751,788,228,251]" pageId="3" pageNumber="446">Hu</collectorName>
,
<collectorName id="935B531EFF88FFEAFC8AD1C0FCA160FA" box="[804,864,228,252]" pageId="3" pageNumber="446">Song</collectorName>
,
<collectorName id="935B531EFF88FFEAFCC1D1C0FC4E60FD" box="[879,911,228,251]" pageId="3" pageNumber="446">Lv</collectorName>
,
<collectorName id="935B531EFF88FFEAFC31D1C0FC0960FA" box="[927,968,228,252]" pageId="3" pageNumber="446">Xie</collectorName>
&amp;
<collectorName id="935B531EFF88FFEAFC5DD1C0FBD360FD" box="[1011,1042,228,251]" pageId="3" pageNumber="446">Yu</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF88FFEAFBCDD1C0FB7560FA" box="[1123,1204,228,252]" pageId="3" pageNumber="446">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF88FFEAFB65D1C0FE9C6169" ID-GBIF-Occurrence="1290996361" collectingDate="2013-07-13" collectionCode="SNUC" collectorName="Song &amp; Yin &amp; Yu" country="China" elevation="925" location="Mingyue Shan National Park" municipality="Yichun City" pageId="3" pageNumber="446" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Jiangxi Province" typeStatus="paratype">
<specimenCount id="28A8FD41FF88FFEAFB65D1C0FB3A60FD" box="[1227,1275,228,252]" pageId="3" pageNumber="446" type="male">1 ♂</specimenCount>
,
<specimenCount id="28A8FD41FF88FFEAFAA5D1C0FAF860FD" box="[1291,1337,228,252]" pageId="3" pageNumber="446" type="female">1 ♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF88FFEAFF35D02FFF266122" box="[155,231,267,292]" name="China" pageId="3" pageNumber="446">China</collectingCountry>
: W.
<collectingRegion id="FC6AF82AFF88FFEAFEB9D02FFE146122" box="[279,469,267,292]" country="China" name="Jiangxi" pageId="3" pageNumber="446">Jiangxi Province</collectingRegion>
,
<collectingMunicipality id="DE75ACB2FF88FFEAFE4ED02FFDA66122" box="[480,615,267,292]" pageId="3" pageNumber="446">Yichun City</collectingMunicipality>
(宜春市),
<location id="3B716013FF88FFEAFD5FD02FFBF76122" LSID="urn:lsid:plazi:treatment:B60787DEFF89FFEDFF39D74FFCB5611B:3B716013FF88FFEAFD5FD02FFBF76122" box="[753,1078,267,292]" country="China" municipality="Yichun City" name="Mingyue Shan National Park" pageId="3" pageNumber="446" stateProvince="Jiangxi Province">Mingyue Shan National Park</location>
(明月山国家公园), 27°35'43- 41&quot;N, 114°16'29-05&quot;E, along path in mixed forest, bamboo, leaf litter, sifted,
<quantity id="F9569B2DFF88FFEAFBAFD00BFB4B6141" box="[1025,1162,303,328]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="11.5" metricValueMin="7.0" pageId="3" pageNumber="446" unit="m" value="925.0" valueMax="1150.0" valueMin="700.0">
<elevation id="B583D1FBFF88FFEAFBAFD00BFB4B6141" box="[1025,1162,303,328]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="11.5" metricValueMin="7.0" pageId="3" pageNumber="446" unit="m" value="925.0" valueMax="1150.0" valueMin="700.0">700-1150 m</elevation>
</quantity>
,
<date id="4A101008FF88FFEAFB39D00BFAD8614E" box="[1175,1305,303,328]" pageId="3" pageNumber="446" value="2013-07-13">
<collectingDate id="5A54E9E0FF88FFEAFB39D00BFAD8614E" box="[1175,1305,303,328]" pageId="3" pageNumber="446" value="2013-07-13">13.vii.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF88FFEAFA8BD014FAA0614E" box="[1317,1377,304,328]" pageId="3" pageNumber="446">Song</collectorName>
,
<collectorName id="935B531EFF88FFEAFAC3D00BFA59614E" box="[1389,1432,303,328]" pageId="3" pageNumber="446">Yin</collectorName>
,
<collectorName id="935B531EFF88FFEAFF39D07CFF776169" box="[151,182,344,367]" pageId="3" pageNumber="446">Yu</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF88FFEAFEAAD073FE946169" box="[260,341,343,367]" pageId="3" pageNumber="446">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF88FFEAFEC5D073FEA961BD" ID-GBIF-Occurrence="1290996390" collectingDate="2013-10-22" collectionCode="SNUC" collectorName="Peng &amp; Shen &amp; Yan" country="China" elevation="1600" latitude="27.586945" location="Mingyue Shan" longLatPrecision="20" longitude="114.281395" municipality="Yichun City" pageId="3" pageNumber="446" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Jiangxi Province" typeStatus="paratype">
<specimenCount id="28A8FD41FF88FFEAFEC5D073FE6E6169" box="[363,431,343,367]" pageId="3" pageNumber="446" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="28A8FD41FF88FFEAFE13D073FDC16169" box="[445,512,343,367]" pageId="3" pageNumber="446" type="female">3 ♀♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF88FFEAFDDAD073FD7D6176" box="[628,700,343,368]" name="China" pageId="3" pageNumber="446">China</collectingCountry>
: W.
<collectingRegion id="FC6AF82AFF88FFEAFD5FD073FC736176" box="[753,946,343,368]" country="China" name="Jiangxi" pageId="3" pageNumber="446">Jiangxi Province</collectingRegion>
,
<collectingMunicipality id="DE75ACB2FF88FFEAFC11D073FB8A6176" box="[959,1099,343,368]" pageId="3" pageNumber="446">Yichun City</collectingMunicipality>
,
<location id="3B716013FF88FFEAFBF7D073FAC16176" LSID="urn:lsid:plazi:treatment:B60787DEFF89FFEDFF39D74FFCB5611B:3B716013FF88FFEAFBF7D073FAC16176" box="[1113,1280,343,368]" country="China" latitude="27.586945" longLatPrecision="20" longitude="114.281395" municipality="Yichun City" name="Mingyue Shan" pageId="3" pageNumber="446" stateProvince="Jiangxi Province">Mingyue Shan</location>
(明月山),
<geoCoordinate id="5B9A500FFF88FFEAFF39D05FFEDC6192" box="[151,285,379,404]" direction="north" orientation="latitude" pageId="3" pageNumber="446" precision="15" value="27.586945">27°35'13&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF88FFEAFE84D05FFE7B6192" box="[298,442,379,404]" direction="east" orientation="longitude" pageId="3" pageNumber="446" precision="15" value="114.281395">114°16'53&quot;E</geoCoordinate>
, mixed forest, leaf litter, wood, sifted &amp; beating, ca.
<quantity id="F9569B2DFF88FFEAFBB7D05FFBB06195" box="[1049,1137,379,404]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="446" unit="m" value="1600.0">
<elevation id="B583D1FBFF88FFEAFBB7D05FFBB06195" box="[1049,1137,379,404]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="446" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<date id="4A101008FF88FFEAFBD1D05FFB336195" box="[1151,1266,379,403]" pageId="3" pageNumber="446" value="2013-10-22">
<collectingDate id="5A54E9E0FF88FFEAFBD1D05FFB336195" box="[1151,1266,379,403]" pageId="3" pageNumber="446" value="2013-10-22">22.x.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF88FFEAFB50D058FAF86192" box="[1278,1337,380,404]" pageId="3" pageNumber="446">Peng</collectorName>
,
<collectorName id="935B531EFF88FFEAFAE8D05FFABF6192" box="[1350,1406,379,404]" pageId="3" pageNumber="446">Shen</collectorName>
&amp;
<collectorName id="935B531EFF88FFEAFF39D087FF0261BC" box="[151,195,419,442]" pageId="3" pageNumber="446">Yan</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF88FFEAFEBED087FEA061BD" box="[272,353,419,443]" pageId="3" pageNumber="446">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF88FFEAFED8D087FF36620C" ID-GBIF-Occurrence="1290996375" collectingDate="2013-07-24" collectionCode="SNUC" collectorName="Hu &amp; Lv" country="China" county="Fengxin County" elevation="1000" latitude="28.820002" location="Niyang Shan" longLatPrecision="20" longitude="115.05723" municipality="Yichun City" pageId="3" pageNumber="446" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Jiangxi Province" typeStatus="paratype">
<specimenCount id="28A8FD41FF88FFEAFED8D087FE6561BC" box="[374,420,419,443]" pageId="3" pageNumber="446" type="male">1 ♂</specimenCount>
,
<specimenCount id="28A8FD41FF88FFEAFE1CD087FE3561BC" box="[434,500,419,443]" pageId="3" pageNumber="446" type="female">4 ♀♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF88FFEAFDCCD087FD6F61BD" box="[610,686,419,443]" name="China" pageId="3" pageNumber="446">China</collectingCountry>
: W.
<collectingRegion id="FC6AF82AFF88FFEAFD4DD087FC6561BD" box="[739,932,419,443]" country="China" name="Jiangxi" pageId="3" pageNumber="446">Jiangxi Province</collectingRegion>
,
<collectingMunicipality id="DE75ACB2FF88FFEAFC1FD087FBFD61BD" box="[945,1084,419,443]" pageId="3" pageNumber="446">Yichun City</collectingMunicipality>
,
<collectingCounty id="D7704E44FF88FFEAFBE4D087FAC061BD" box="[1098,1281,419,443]" pageId="3" pageNumber="446">Fengxin County</collectingCounty>
(奉新县),
<location id="3B716013FF88FFEAFF39D0EEFEE861E5" LSID="urn:lsid:plazi:treatment:B60787DEFF89FFEDFF39D74FFCB5611B:3B716013FF88FFEAFF39D0EEFEE861E5" box="[151,297,458,483]" country="China" county="Fengxin County" latitude="28.820002" longLatPrecision="20" longitude="115.05723" municipality="Yichun City" name="Niyang Shan" pageId="3" pageNumber="446" stateProvince="Jiangxi Province">Niyang Shan</location>
(泥洋山),
<geoCoordinate id="5B9A500FFF88FFEAFE6CD0EFFD8961E5" box="[450,584,459,483]" direction="north" orientation="latitude" pageId="3" pageNumber="446" precision="15" value="28.820002">28°49'12&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF88FFEAFDFAD0EEFD2561E5" box="[596,740,458,483]" direction="east" orientation="longitude" pageId="3" pageNumber="446" precision="15" value="115.05723">115°03'26&quot;E</geoCoordinate>
, mixed leaf litter, sifted,
<quantity id="F9569B2DFF88FFEAFC51D0EFFB9661E4" box="[1023,1111,459,483]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="446" unit="m" value="1000.0">
<elevation id="B583D1FBFF88FFEAFC51D0EFFB9661E4" box="[1023,1111,459,483]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="446" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="4A101008FF88FFEAFBCAD0EFFB2E61E5" box="[1124,1263,459,483]" pageId="3" pageNumber="446" value="2013-07-24">
<collectingDate id="5A54E9E0FF88FFEAFBCAD0EFFB2E61E5" box="[1124,1263,459,483]" pageId="3" pageNumber="446" value="2013-07-24">24.VII.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF88FFEAFB55D0EFFADC61E4" box="[1275,1309,459,482]" pageId="3" pageNumber="446">Hu</collectorName>
&amp;
<collectorName id="935B531EFF88FFEAFAEDD0EFFAA361E4" box="[1347,1378,459,482]" pageId="3" pageNumber="446">Lv</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF88FFEAFF31D0D6FF31620C" box="[159,240,498,522]" pageId="3" pageNumber="446">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF88FFEAFEACD0D6FF366254" ID-GBIF-Occurrence="1290996387" collectingDate="2013-10-24" collectionCode="SNUC" collectorName="Peng &amp; Shen &amp; Yan" country="China" county="Luxi County" elevation="1360" latitude="27.585278" location="Yangshimu" longLatPrecision="20" longitude="114.26139" pageId="3" pageNumber="446" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Jiangxi Province" typeStatus="paratype">
<specimenCount id="28A8FD41FF88FFEAFEACD0D6FEEC620C" box="[258,301,498,522]" pageId="3" pageNumber="446" type="male">1 ♂</specimenCount>
,
<specimenCount id="28A8FD41FF88FFEAFE97D0D6FEA3620C" box="[313,354,498,522]" pageId="3" pageNumber="446" type="female">1 ♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF88FFEAFE65D0D6FDD6620C" box="[459,535,498,522]" name="China" pageId="3" pageNumber="446">China</collectingCountry>
: W.
<collectingRegion id="FC6AF82AFF88FFEAFDE8D0D6FCC5620C" box="[582,772,498,522]" country="China" name="Jiangxi" pageId="3" pageNumber="446">Jiangxi Province</collectingRegion>
,
<collectingCounty id="D7704E44FF88FFEAFCA0D0D6FC5A620C" box="[782,923,498,522]" pageId="3" pageNumber="446">Luxi County</collectingCounty>
(芦溪县),
<location id="3B716013FF88FFEAFBB9D0D6FB54620D" LSID="urn:lsid:plazi:treatment:B60787DEFF89FFEDFF39D74FFCB5611B:3B716013FF88FFEAFBB9D0D6FB54620D" box="[1047,1173,498,523]" country="China" county="Luxi County" latitude="27.585278" longLatPrecision="20" longitude="114.26139" name="Yangshimu" pageId="3" pageNumber="446" stateProvince="Jiangxi Province">Yangshimu</location>
(羊狮幕),
<geoCoordinate id="5B9A500FFF88FFEAFABFD0D6FA56620D" box="[1297,1431,498,523]" direction="north" orientation="latitude" pageId="3" pageNumber="446" precision="15" value="27.585278">27°35'07&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF88FFEAFF39D332FEE66229" box="[151,295,534,559]" direction="east" orientation="longitude" pageId="3" pageNumber="446" precision="15" value="114.26139">114°15'41&quot;E</geoCoordinate>
, mixed forest, leaf litter, wood, sifted &amp; beating, ca.,
<quantity id="F9569B2DFF88FFEAFC33D332FC396228" box="[925,1016,534,559]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="3" pageNumber="446" unit="m" value="1360.0">
<elevation id="B583D1FBFF88FFEAFC33D332FC396228" box="[925,1016,534,559]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="3" pageNumber="446" unit="m" value="1360.0">1360 m</elevation>
</quantity>
,
<date id="4A101008FF88FFEAFBA9D333FBB86229" box="[1031,1145,535,559]" pageId="3" pageNumber="446" value="2013-10-24">
<collectingDate id="5A54E9E0FF88FFEAFBA9D333FBB86229" box="[1031,1145,535,559]" pageId="3" pageNumber="446" value="2013-10-24">24.x.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF88FFEAFB26D333FB026229" box="[1160,1219,535,559]" pageId="3" pageNumber="446">Peng</collectorName>
,
<collectorName id="935B531EFF88FFEAFB7FD332FAC86229" box="[1233,1289,534,559]" pageId="3" pageNumber="446">Shen</collectorName>
&amp;
<collectorName id="935B531EFF88FFEAFA9DD333FA9E6228" box="[1331,1375,535,558]" pageId="3" pageNumber="446">Yan</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF88FFEAFF31D31EFF316254" box="[159,240,570,594]" pageId="3" pageNumber="446">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF88FFEAFEADD31EFAA16254" ID-GBIF-Occurrence="1290996393" box="[259,1376,570,595]" collectingDate="2013-10-28" collectionCode="SNUC" collectorName="Peng &amp; Shen &amp; Yan" country="China" county="Luxi County" elevation="1050" latitude="27.466389" location="Yangshimu" longLatPrecision="20" longitude="114.165" pageId="3" pageNumber="446" specimenCount="1" specimenCount-male="1" stateProvince="Jiangxi Province" typeStatus="paratype">
<specimenCount id="28A8FD41FF88FFEAFEADD31EFEEF6254" box="[259,302,570,594]" pageId="3" pageNumber="446" type="male">1 ♂</specimenCount>
, same data as previous material except ‘
<geoCoordinate id="5B9A500FFF88FFEAFD42D31EFCB66255" box="[748,887,570,595]" direction="north" orientation="latitude" pageId="3" pageNumber="446" precision="15" value="27.466389">27°27'59&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF88FFEAFC2CD31EFBCE6255" box="[898,1039,570,595]" direction="east" orientation="longitude" pageId="3" pageNumber="446" precision="15" value="114.165">114°09'54&quot;E</geoCoordinate>
,
<quantity id="F9569B2DFF88FFEAFBB3D31EFBB56254" box="[1053,1140,570,594]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="446" unit="m" value="1050.0">
<elevation id="B583D1FBFF88FFEAFBB3D31EFBB56254" box="[1053,1140,570,594]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="3" pageNumber="446" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<date id="4A101008FF88FFEAFBD1D31EFB356254" box="[1151,1268,570,594]" pageId="3" pageNumber="446" value="2013-10-28">
<collectingDate id="5A54E9E0FF88FFEAFBD1D31EFB356254" box="[1151,1268,570,594]" pageId="3" pageNumber="446" value="2013-10-28">28.x.2013</collectingDate>
</date>
’ (
<collectionCode id="58BFAE0DFF88FFEAFAA9D31EFA996254" box="[1287,1368,570,594]" pageId="3" pageNumber="446">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF88FFEAFAC5D31EFF3662C0" ID-GBIF-Occurrence="1290996388" collectingDate="2013-07-23" collectionCode="SNUC" collectorName="Song &amp; Yin &amp; Yu" country="China" county="Gaozhou County" elevation="1025" latitude="27.3975" location="Pingxiang City" longLatPrecision="20" longitude="114.015" pageId="3" pageNumber="446" specimenCount="1" specimenCount-male="1" stateProvince="Jiangxi" typeStatus="paratype">
<specimenCount id="28A8FD41FF88FFEAFAC5D31EFA566254" box="[1387,1431,570,594]" pageId="3" pageNumber="446" type="male">1 ♂</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF88FFEAFF5BD346FE80627D" box="[245,321,610,635]" name="China" pageId="3" pageNumber="446">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF88FFEAFEE0D346FE63627D" box="[334,418,610,635]" country="China" name="Jiangxi" pageId="3" pageNumber="446">Jiangxi</collectingRegion>
,
<location id="3B716013FF88FFEAFE01D346FD9A627D" LSID="urn:lsid:plazi:treatment:B60787DEFF89FFEDFF39D74FFCB5611B:3B716013FF88FFEAFE01D346FD9A627D" box="[431,603,610,635]" country="China" county="Gaozhou County" latitude="27.3975" longLatPrecision="20" longitude="114.015" name="Pingxiang City" pageId="3" pageNumber="446" stateProvince="Jiangxi">Pingxiang City</location>
,
<collectingCounty id="D7704E44FF88FFEAFDC6D346FCEB627C" box="[616,810,610,635]" pageId="3" pageNumber="446">Gaozhou County</collectingCounty>
(高州县), Gao-Tian-Yan (高天岩),
<geoCoordinate id="5B9A500FFF88FFEAFABFD346FA56627D" box="[1297,1431,610,635]" direction="north" orientation="latitude" pageId="3" pageNumber="446" precision="15" value="27.3975">27°23'51&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF88FFEAFF39D3A1FEE66298" box="[151,295,645,670]" direction="east" orientation="longitude" pageId="3" pageNumber="446" precision="15" value="114.015">114°00'54&quot;E</geoCoordinate>
, broad-leaved &amp; pine forest, mixed litter, sifted, ca.
<quantity id="F9569B2DFF88FFEAFC32D3A1FC396298" box="[924,1016,645,670]" metricMagnitude="3" metricUnit="m" metricValue="1.025" pageId="3" pageNumber="446" unit="m" value="1025.0">
<elevation id="B583D1FBFF88FFEAFC32D3A1FC396298" box="[924,1016,645,670]" metricMagnitude="3" metricUnit="m" metricValue="1.025" pageId="3" pageNumber="446" unit="m" value="1025.0">1025 m</elevation>
</quantity>
,
<date id="4A101008FF88FFEAFBA7D3A1FB4A6298" box="[1033,1163,645,670]" pageId="3" pageNumber="446" value="2013-07-23">
<collectingDate id="5A54E9E0FF88FFEAFBA7D3A1FB4A6298" box="[1033,1163,645,670]" pageId="3" pageNumber="446" value="2013-07-23">23.vii.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF88FFEAFB32D3A2FB196298" box="[1180,1240,646,670]" pageId="3" pageNumber="446">Song</collectorName>
,
<collectorName id="935B531EFF88FFEAFB46D3A1FAD16298" box="[1256,1296,645,670]" pageId="3" pageNumber="446">Yin</collectorName>
&amp;
<collectorName id="935B531EFF88FFEAFA91D3A3FA9F6298" box="[1343,1374,647,670]" pageId="3" pageNumber="446">Yu</collectorName>
leg,’ (
<collectionCode id="58BFAE0DFF88FFEAFF31D38AFF3162C0" box="[159,240,686,710]" pageId="3" pageNumber="446">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF88FFEAFEACD38AFF366317" ID-GBIF-Occurrence="1290996366" collectingDate="2014-07-31" collectionCode="SNUC" collectorName="Peng &amp; Song &amp; Yu &amp; Yan" country="China" county="Xing'an County" elevation="1180" latitude="25.887222" location="Jiuniutang" longLatPrecision="20" longitude="110.48694" municipality="Maoer Shan" pageId="3" pageNumber="446" specimenCount="14" specimenCount-female="11" specimenCount-male="3" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="28A8FD41FF88FFEAFEACD38AFE8362C3" box="[258,322,686,710]" pageId="3" pageNumber="446" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="28A8FD41FF88FFEAFEE0D38AFE5B62C3" box="[334,410,686,710]" pageId="3" pageNumber="446" type="female">11 ♀♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF88FFEAFDACD389FD8F62C3" box="[514,590,685,709]" name="China" pageId="3" pageNumber="446">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF88FFEAFDF6D389FD7A62C3" box="[600,699,685,709]" country="China" name="Guangxi" pageId="3" pageNumber="446">Guangxi</collectingRegion>
,
<collectingCounty id="D7704E44FF88FFEAFD68D389FCBA62C0" box="[710,891,685,710]" pageId="3" pageNumber="446">Xing’an County</collectingCounty>
(兴安县),
<collectingMunicipality id="DE75ACB2FF88FFEAFC58D38AFBBF62C0" box="[1014,1150,685,710]" pageId="3" pageNumber="446">Maoer Shan</collectingMunicipality>
(猫儿山),
<location id="3B716013FF88FFEAFB55D389FAB162C0" LSID="urn:lsid:plazi:treatment:B60787DEFF89FFEDFF39D74FFCB5611B:3B716013FF88FFEAFB55D389FAB162C0" box="[1275,1392,685,710]" country="China" county="Xing'an County" latitude="25.887222" longLatPrecision="20" longitude="110.48694" municipality="Maoer Shan" name="Jiuniutang" pageId="3" pageNumber="446" stateProvince="Guangxi">Jiuniutang</location>
(九 牛塘),
<geoCoordinate id="5B9A500FFF88FFEAFF42D3F1FEB262E8" box="[236,371,725,750]" direction="north" orientation="latitude" pageId="3" pageNumber="446" precision="15" value="25.887222">25°53'14&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF88FFEAFED0D3F2FDCF62E8" box="[382,526,726,750]" direction="east" orientation="longitude" pageId="3" pageNumber="446" precision="15" value="110.48694">110°29'13&quot;E</geoCoordinate>
, mixed leaf litter, humus, sifted,
<quantity id="F9569B2DFF88FFEAFCD5D3F2FC1162EB" box="[891,976,726,750]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="3" pageNumber="446" unit="m" value="1180.0">
<elevation id="B583D1FBFF88FFEAFCD5D3F2FC1162EB" box="[891,976,726,750]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="3" pageNumber="446" unit="m" value="1180.0">1180 m</elevation>
</quantity>
,
<date id="4A101008FF88FFEAFC72D3F1FB9F62E8" box="[988,1118,725,750]" pageId="3" pageNumber="446" value="2014-07-31">
<collectingDate id="5A54E9E0FF88FFEAFC72D3F1FB9F62E8" box="[988,1118,725,750]" pageId="3" pageNumber="446" value="2014-07-31">31.vii.2014</collectingDate>
</date>
,
<collectorName id="935B531EFF88FFEAFBC7D3F2FB6562E8" box="[1129,1188,726,750]" pageId="3" pageNumber="446">Peng</collectorName>
,
<collectorName id="935B531EFF88FFEAFB00D3F2FB2B62E8" box="[1198,1258,726,750]" pageId="3" pageNumber="446">Song</collectorName>
,
<collectorName id="935B531EFF88FFEAFB5BD3F2FAD562EB" box="[1269,1300,726,749]" pageId="3" pageNumber="446">Yu</collectorName>
&amp;
<collectorName id="935B531EFF88FFEAFA99D3F2FAA262EB" box="[1335,1379,726,749]" pageId="3" pageNumber="446">Yan</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF88FFEAFF31D3DDFF316317" box="[159,240,761,785]" pageId="3" pageNumber="446">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF88FFEAFEA8D3DDFDF66330" ID-GBIF-Occurrence="1290996389" collectingDate="2014-07-28" collectionCode="SNUC" collectorName="Peng &amp; Song &amp; Yu &amp; Yan" country="China" county="Xing'an County" elevation="1020" latitude="25.867779" location="Jiuniutang" longLatPrecision="20" longitude="110.440834" municipality="Maoer Shan" pageId="3" pageNumber="446" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="28A8FD41FF88FFEAFEA8D3DDFEF26317" box="[262,307,761,785]" pageId="3" pageNumber="446" type="female">1 ♀</specimenCount>
, same data as previous material except ‘
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,
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, beech forest, mixed leaf litter, sifted,
<quantity id="F9569B2DFF88FFEAFF4DD23AFEFB6333" box="[227,314,798,822]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="446" unit="m" value="1020.0">
<elevation id="B583D1FBFF88FFEAFF4DD23AFEFB6333" box="[227,314,798,822]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="446" unit="m" value="1020.0">1020 m</elevation>
</quantity>
,
<date id="4A101008FF88FFEAFEE8D239FE0B6330" box="[326,458,797,822]" pageId="3" pageNumber="446" value="2014-07-28">
<collectingDate id="5A54E9E0FF88FFEAFEE8D239FE0B6330" box="[326,458,797,822]" pageId="3" pageNumber="446" value="2014-07-28">28.vii.2014</collectingDate>
</date>
’ (
<collectionCode id="58BFAE0DFF88FFEAFE70D23AFDEE6330" box="[478,559,798,822]" pageId="3" pageNumber="446">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF88FFEAFF69D265FA5C63EC" pageId="3" pageNumber="446" type="diagnosis">
<paragraph id="3E1136C8FF88FFEAFF69D265FA5C63EC" blockId="3.[151,1437,155,2010]" pageId="3" pageNumber="446">
<emphasis id="0CDAEADAFF88FFEAFF69D265FE61635C" bold="true" box="[199,416,833,858]" pageId="3" pageNumber="446">Diagnosis of male.</emphasis>
Body glabrous, elytra and abdomen with sparse, short setae; frons with abrupt subantennal transverse sulcus; antennomeres IX elongate, XI concaved anterolaterally; pronotum with shallow median antebasal fovea and distinct basal impression; all tibiae slender; protrochanters protuberant at ventral margin, profemora simple, protibiae sometimes with preapical protuberance, mesotibiae lacking apical spur; tergite IV with narrow basal sulcus, discal carinae relatively long, close, with short marginal carinae; aedeagus relatively elongate.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF88FFEAFF69D2D1FD456794" pageId="3" pageNumber="446" type="description">
<paragraph id="3E1136C8FF88FFEAFF69D2D1FF38662C" blockId="3.[151,1437,155,2010]" pageId="3" pageNumber="446">
<emphasis id="0CDAEADAFF88FFEAFF69D2D1FE986408" bold="true" box="[199,345,1013,1038]" pageId="3" pageNumber="446">Description.</emphasis>
Male (
<figureCitation id="A6952A4DFF88FFEAFE05D2D1FE306408" box="[427,497,1013,1038]" captionStart="FIGURE 2" captionStartId="4.[151,250,1611,1633]" captionTargetBox="[162,1431,349,1576]" captionTargetId="figure@4.[151,1435,339,1589]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal habitus of Batraxis males. A. B. robusta, population from Guangxi. B. B. gloriosa, population from Hunan. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256543/files/figure.png" pageId="3" pageNumber="446">Fig. 2</figureCitation>
B). Body reddish-brown, length
<quantity id="F9569B2DFF88FFEAFCCFD2D1FBC8640B" box="[865,1033,1013,1038]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" metricValueMax="1.74" metricValueMin="1.54" pageId="3" pageNumber="446" unit="mm" value="1.64" valueMax="1.74" valueMin="1.54">1.54–1.74 mm</quantity>
. Head rectangular, as long as wide, HL
<quantity id="F9569B2DFF88FFEAFF6BD53EFEAC6437" box="[197,365,1050,1074]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="4.0" metricValueMin="3.3" pageId="3" pageNumber="446" unit="mm" value="0.365" valueMax="0.4" valueMin="0.33">0.33–0.40 mm</quantity>
, HW
<quantity id="F9569B2DFF88FFEAFE1CD53EFD9B6437" box="[434,602,1050,1074]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.9" metricValueMin="3.4" pageId="3" pageNumber="446" unit="mm" value="0.365" valueMax="0.39" valueMin="0.34">0.34–0.39 mm</quantity>
, vertex glabrous, apical margin of frons and clypeus sparsely pubescent; vertex moderately convex, with pair of small vertexal foveae; antennal tubercles low; frons flattened, subantennal transverse sulcus abrupt, indistinct at middle, shallow vertexal sulcus merging with subantennal transverse sulcus and connecting foveae; each eye composed of about 20 facets; antennae relatively thin, antennomeres III–IX elongate, X slightly transverse, XI largest, widest at middle, concaved anterolaterally; median gular carina narrow, with two distinct nude gular foveae. Pronotum slightly wider than long, PL
<quantity id="F9569B2DFF88FFEAFBA8D5EAFB6F64E3" box="[1030,1198,1230,1254]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="3" pageNumber="446" unit="mm" value="0.315" valueMax="0.33" valueMin="0.3">0.30–0.33 mm</quantity>
, PW
<quantity id="F9569B2DFF88FFEAFB5ED5E9FA5964E3" box="[1264,1432,1229,1254]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="3.9" metricValueMin="3.5" pageId="3" pageNumber="446" unit="mm" value="0.37" valueMax="0.39" valueMin="0.35">0.35–0.39 mm</quantity>
, with indistinct median antebasal fovea and distinct basal impression. Elytra wider than long, EL
<quantity id="F9569B2DFF88FFEAFB5ED5D5FA59650F" box="[1264,1432,1265,1290]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="3" pageNumber="446" unit="mm" value="0.5" valueMax="0.55" valueMin="0.45">0.45–0.55 mm</quantity>
, EW
<quantity id="F9569B2DFF88FFEAFF61D431FEBB652B" box="[207,378,1301,1326]" metricMagnitude="-4" metricUnit="m" metricValue="6.35" metricValueMax="6.8" metricValueMin="5.8999999999999995" pageId="3" pageNumber="446" unit="mm" value="0.635" valueMax="0.68" valueMin="0.59">0.59–0.68 mm</quantity>
, dorsal surface with sparse, minute setae, each elytron with two distinct foveae at base. Protrochanters protuberant at ventral margin (
<figureCitation id="A6952A4DFF88FFEAFD33D41DFD316554" box="[669,752,1337,1362]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="3" pageNumber="446">Fig. 14</figureCitation>
J). Abdomen wider than long, AL
<quantity id="F9569B2DFF88FFEAFBD8D41DFADD6557" box="[1142,1308,1337,1362]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.699999999999999" metricValueMin="4.0" pageId="3" pageNumber="446" unit="mm" value="0.485" valueMax="0.57" valueMin="0.4">0.40–0.57 mm</quantity>
, AW
<quantity id="F9569B2DFF88FFEAFAF3D41DFEC36573" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.2" metricValueMin="5.5" pageId="3" pageNumber="446" unit="mm" value="0.585" valueMax="0.62" valueMin="0.55">0.55– 0.62 mm</quantity>
, slightly narrowed at base; tergite IV (first visible tergite) with narrow basal sulcus; discal carinae relatively long, close, extending to apical one-third of tergal length, width between carinae
<quantity id="F9569B2DFF88FFEAFB25D4A5FAEF659F" box="[1163,1326,1409,1434]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="3" pageNumber="446" unit="mm" value="0.13" valueMax="0.16" valueMin="0.1">0.10–0.16 mm</quantity>
at apices; short marginal carinae extending to apical one-ninth of tergal length. Aedeagus (
<figureCitation id="A6952A4DFF88FFEAFBE8D481FB4A65B8" box="[1094,1163,1445,1470]" captionStart="FIGURE 9" captionStartId="15.[151,250,1924,1946]" captionTargetBox="[192,1403,193,1899]" captionTargetId="figure@15.[181,1405,193,1903]" captionTargetPageId="15" captionText="FIGURE 9. Aedeagi of Batraxis gloriosa. (A, B, D, E, G, H. in ventral view. C, F, I. in lateral view). A – C. Population from Hunan. D – F. Population from Fujian. G – I. Population from Zhejiang. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256550/files/figure.png" pageId="3" pageNumber="446">Fig. 9</figureCitation>
A‒C)
<quantity id="F9569B2DFF88FFEAFB7BD482FABD65BB" box="[1237,1404,1446,1470]" metricMagnitude="-4" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="3" pageNumber="446" unit="mm" value="0.255" valueMax="0.27" valueMin="0.24">0.24–0.27 mm</quantity>
in length, relatively elongate, parameres distinctly exceeding apex of median lobe, apex flattened, each with two preapical setae; endophallus with forked sclerite at middle and numerous weakly sclerotized spine-like structures laterally.
</paragraph>
<paragraph id="3E1136C8FF88FFEAFF69D711FDDD66BC" blockId="3.[151,1437,155,2010]" pageId="3" pageNumber="446">
Female. Similar to male in general appearance; protrochanters simple; each eye composed of about 20 facets. Measurements: BL
<quantity id="F9569B2DFF88FFEAFEDDD77DFDD86677" box="[371,537,1625,1650]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" metricValueMax="1.7" metricValueMin="1.56" pageId="3" pageNumber="446" unit="mm" value="1.63" valueMax="1.7" valueMin="1.56">1.56–1.70 mm</quantity>
; HL
<quantity id="F9569B2DFF88FFEAFDFFD77EFD376677" box="[593,758,1626,1650]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.8" metricValueMin="3.4" pageId="3" pageNumber="446" unit="mm" value="0.36" valueMax="0.38" valueMin="0.34">0.34–0.38 mm</quantity>
; HW
<quantity id="F9569B2DFF88FFEAFC99D77DFC1C6677" box="[823,989,1625,1650]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="3" pageNumber="446" unit="mm" value="0.365" valueMax="0.38" valueMin="0.35">0.35–0.38 mm</quantity>
; PL
<quantity id="F9569B2DFF88FFEAFBBED77EFB776677" box="[1040,1206,1626,1650]" metricMagnitude="-4" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="3" pageNumber="446" unit="mm" value="0.305" valueMax="0.32" valueMin="0.29">0.29–0.32 mm</quantity>
; PW
<quantity id="F9569B2DFF88FFEAFB5CD77DFA596677" box="[1266,1432,1625,1650]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="3" pageNumber="446" unit="mm" value="0.375" valueMax="0.4" valueMin="0.35">0.35–0.40 mm</quantity>
; EL
<quantity id="F9569B2DFF88FFEAFF6CD75AFEAB6693" box="[194,362,1662,1686]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.9" metricValueMin="4.4" pageId="3" pageNumber="446" unit="mm" value="0.46499999999999997" valueMax="0.49" valueMin="0.44">0.44–0.49 mm</quantity>
; EW
<quantity id="F9569B2DFF88FFEAFE02D759FD946693" box="[428,597,1661,1686]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="6.3" metricValueMin="5.699999999999999" pageId="3" pageNumber="446" unit="mm" value="0.6" valueMax="0.63" valueMin="0.57">0.57–0.63 mm</quantity>
; AL
<quantity id="F9569B2DFF88FFEAFD3ED759FCF96693" box="[656,824,1661,1686]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.2" metricValueMin="4.4" pageId="3" pageNumber="446" unit="mm" value="0.48" valueMax="0.52" valueMin="0.44">0.44–0.52 mm</quantity>
; AW
<quantity id="F9569B2DFF88FFEAFCD5D759FBE56693" box="[891,1060,1661,1686]" metricMagnitude="-4" metricUnit="m" metricValue="5.7" metricValueMax="6.0" metricValueMin="5.4" pageId="3" pageNumber="446" unit="mm" value="0.57" valueMax="0.6" valueMin="0.54">0.54–0.60 mm</quantity>
; width between discal carinae of tergite IV
<quantity id="F9569B2DFF88FFEAFEA7D785FE6D66BF" box="[265,428,1697,1722]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="446" unit="mm" value="0.125" valueMax="0.15" valueMin="0.1">0.10–0.15 mm</quantity>
at apices.
</paragraph>
<paragraph id="3E1136C8FF88FFEAFF69D7E1FD456794" blockId="3.[151,1437,155,2010]" pageId="3" pageNumber="446">
<emphasis id="0CDAEADAFF88FFEAFF69D7E1FE7566D8" bold="true" box="[199,436,1733,1758]" pageId="3" pageNumber="446">Comparative notes.</emphasis>
<taxonomicName id="F9AE4D4BFF88FFEAFE10D7E2FD4466DB" box="[446,645,1733,1757]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="446" phylum="Arthropoda" rank="species" species="gloriosa">
<emphasis id="0CDAEADAFF88FFEAFE10D7E2FD4466DB" box="[446,645,1733,1757]" italics="true" pageId="3" pageNumber="446">Batraxis gloriosa</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="F9AE4D4BFF88FFEAFC88D7E2FC1766DB" box="[806,982,1733,1757]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="446" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="0CDAEADAFF88FFEAFC88D7E2FC1766DB" box="[806,982,1733,1757]" italics="true" pageId="3" pageNumber="446">B. nanlingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17E957A1FF88FFEAFC4FD7E2FBE066DB" box="[993,1057,1734,1757]" pageId="3" pageNumber="446" rank="species">sp. n.</taxonomicNameLabel>
(described below) and
<taxonomicName id="F9AE4D4BFF88FFEAFA98D7E2FEEA6704" authority="Wang &amp; Yin" authorityName="Wang &amp; Yin" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="446" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="0CDAEADAFF88FFEAFA98D7E2FA5D66DB" box="[1334,1436,1734,1757]" italics="true" pageId="3" pageNumber="446">B. minor</emphasis>
Wang &amp; Yin
</taxonomicName>
by the broad head which is as wide as the pronotum, and aedeagal endophallus with the median sclerites widely separated at the base. The new species can be separated from
<taxonomicName id="F9AE4D4BFF88FFEAFBAAD62AFB706723" box="[1028,1201,1805,1829]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="446" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="0CDAEADAFF88FFEAFBAAD62AFBD46723" box="[1028,1045,1806,1829]" italics="true" pageId="3" pageNumber="446">B</emphasis>
.
<emphasis id="0CDAEADAFF88FFEAFB8AD629FB706723" box="[1060,1201,1805,1829]" italics="true" pageId="3" pageNumber="446">nanlingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17E957A1FF88FFEAFB17D62AFB346723" box="[1209,1269,1806,1829]" pageId="3" pageNumber="446" rank="species">sp. n.</taxonomicNameLabel>
by the smaller body size (
<quantity id="F9569B2DFF88FFEAFEBFD615FE74674F" box="[273,437,1841,1866]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.59" metricValueMin="1.56" pageId="3" pageNumber="446" unit="mm" value="1.575" valueMax="1.59" valueMin="1.56">1.56‒1.59 mm</quantity>
vs
<quantity id="F9569B2DFF88FFEAFE72D616FDF7674C" box="[476,566,1842,1866]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="446" unit="mm" value="1.7">1.7 mm</quantity>
), the antennomeres XI are distinctly notched at the anterolateral margins, and the more slender aedeagus.
<taxonomicName id="F9AE4D4BFF88FFEAFE61D672FDB8676B" box="[463,633,1878,1901]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="446" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="0CDAEADAFF88FFEAFE61D672FDB8676B" box="[463,633,1878,1901]" italics="true" pageId="3" pageNumber="446">Batraxis minor</emphasis>
</taxonomicName>
has a body size range from
<quantity id="F9569B2DFF88FFEAFC16D671FBDD676B" box="[952,1052,1877,1902]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="3" pageNumber="446" unit="mm" value="1.35">1.35 mm</quantity>
to
<quantity id="F9569B2DFF88FFEAFBEFD672FB66676B" box="[1089,1191,1878,1902]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" pageId="3" pageNumber="446" unit="mm" value="1.37">1.37 mm</quantity>
, and the apices of the aedeagal parameres are distinctly narrowed.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF88FFEDFF69D6B9FEE460D3" lastPageId="4" lastPageNumber="447" pageId="3" pageNumber="446" type="discussion">
<paragraph id="3E1136C8FF88FFEDFF69D6B9FEE460D3" blockId="3.[151,1437,155,2010]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="447" pageId="3" pageNumber="446">
<emphasis id="0CDAEADAFF88FFEAFF69D6B9FE8E67B0" bold="true" box="[199,335,1949,1974]" pageId="3" pageNumber="446">Comments.</emphasis>
The populations from
<collectingRegion id="FC6AF82AFF88FFEAFDF5D6B9FD6367B0" box="[603,674,1949,1974]" country="China" name="Fujian" pageId="3" pageNumber="446">Fujian</collectingRegion>
and
<collectingRegion id="FC6AF82AFF88FFEAFD71D6B9FC8267B0" box="[735,835,1949,1974]" country="China" name="Zhejiang" pageId="3" pageNumber="446">Zhejiang</collectingRegion>
have the protibiae each with an indistinct preapical protuberance (
<figureCitation id="A6952A4DFF88FFEAFE95D6E5FE4F67DC" box="[315,398,1985,2010]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="3" pageNumber="446">Fig. 14</figureCitation>
A‒B), while those from
<collectingRegion id="FC6AF82AFF88FFEAFD05D6E5FD3E67DC" box="[683,767,1985,2010]" country="China" name="Jiangxi" pageId="3" pageNumber="446">Jiangxi</collectingRegion>
,
<collectingRegion id="FC6AF82AFF88FFEAFCA2D6E6FC9867DF" box="[780,857,1986,2009]" country="China" name="Hunan" pageId="3" pageNumber="446">Hunan</collectingRegion>
, and
<collectingRegion id="FC6AF82AFF88FFEAFC34D6E5FC3A67DC" box="[922,1019,1985,2010]" country="China" name="Guangxi" pageId="3" pageNumber="446">Guangxi</collectingRegion>
have simple protibiae. This and the slight differences in the aedeagal structures (
<figureCitation id="A6952A4DFF8FFFEDFD3CD1B3FD1C60B6" box="[658,733,151,176]" captionStart="FIGURE 9" captionStartId="15.[151,250,1924,1946]" captionTargetBox="[192,1403,193,1899]" captionTargetId="figure@15.[181,1405,193,1903]" captionTargetPageId="15" captionText="FIGURE 9. Aedeagi of Batraxis gloriosa. (A, B, D, E, G, H. in ventral view. C, F, I. in lateral view). A – C. Population from Hunan. D – F. Population from Fujian. G – I. Population from Zhejiang. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256550/files/figure.png" pageId="4" pageNumber="447">Figs 9</figureCitation>
,
<figureCitation id="A6952A4DFF8FFFEDFD44D1BCFCC460B6" box="[746,773,152,176]" captionStart="FIGURE 10" captionStartId="16.[151,250,1955,1977]" captionTargetBox="[168,1432,200,1927]" captionTargetId="figure@16.[151,1435,193,1933]" captionTargetPageId="16" captionText="FIGURE 10. Aedeagi of Batraxis species. (A, B, D, E, G, H. in ventral view. C, F, I. in lateral view). A – C. B. gloriosa, population from Jiangxi. D – F. B. gloriosa, population from Guangxi. G – I. B. antennata. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256551/files/figure.png" pageId="4" pageNumber="447">10</figureCitation>
A‒F) are both interpreted as intraspecific variations of the new species.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8FFFEDFF69D1FBFCB5611B" pageId="4" pageNumber="447" type="distribution">
<paragraph id="3E1136C8FF8FFFEDFF69D1FBFCB5611B" blockId="4.[151,1436,151,285]" pageId="4" pageNumber="447">
<emphasis id="0CDAEADAFF8FFFEDFF69D1FBFEA160FE" bold="true" box="[199,352,223,248]" pageId="4" pageNumber="447">Distribution.</emphasis>
<collectingCountry id="46B97658FF8FFFEDFEC9D1FBFE6F60FE" box="[359,430,223,248]" name="China" pageId="4" pageNumber="447">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF8FFFEDFE17D1FBFDE160FE" box="[441,544,223,248]" country="China" name="Zhejiang" pageId="4" pageNumber="447">Zhejiang</collectingRegion>
,
<collectingRegion id="FC6AF82AFF8FFFEDFD84D1FBFDBF60FE" box="[554,638,223,248]" country="China" name="Jiangxi" pageId="4" pageNumber="447">Jiangxi</collectingRegion>
,
<collectingRegion id="FC6AF82AFF8FFFEDFD27D1FBFD1560FE" box="[649,724,223,248]" country="China" name="Fujian" pageId="4" pageNumber="447">Fujian</collectingRegion>
,
<collectingRegion id="FC6AF82AFF8FFFEDFD71D1C4FCED60F1" box="[735,812,224,247]" country="China" name="Hunan" pageId="4" pageNumber="447">Hunan</collectingRegion>
,
<collectingRegion id="FC6AF82AFF8FFFEDFC96D1FBFC5960FE" box="[824,920,223,248]" country="China" name="Guangxi" pageId="4" pageNumber="447">Guangxi</collectingRegion>
(
<figureCitation id="A6952A4DFF8FFFEDFC09D1FBFC3B60FE" box="[935,1018,223,248]" captionStart="FIGURE 15" captionStartId="21.[151,250,1023,1045]" captionTargetBox="[151,1436,193,1001]" captionTargetId="figure@21.[151,1436,193,1002]" captionTargetPageId="21" captionText="FIGURE 15. Distribution of Batraxis species in China." httpUri="https://zenodo.org/record/256556/files/figure.png" pageId="4" pageNumber="447">Fig. 15</figureCitation>
).
<emphasis id="0CDAEADAFF8FFFEDFF69D020FE8F611B" bold="true" box="[199,334,260,285]" pageId="4" pageNumber="447">Etymology.</emphasis>
The specific epithet ‘
<taxonomicName id="F9AE4D4BFF8FFFEDFDEFD020FD5F611A" box="[577,670,260,284]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="447" phylum="Arthropoda" rank="species" species="gloriosa">
<emphasis id="0CDAEADAFF8FFFEDFDEFD020FD5F611A" box="[577,670,260,284]" italics="true" pageId="4" pageNumber="447">gloriosa</emphasis>
</taxonomicName>
’ means ‘glorious’.
</paragraph>
</subSubSection>
</treatment>
</document>

330
data/B6/07/87/B60787DEFF8AFFE8FF39D041FC4E67EA.xml Normal file
View file

@ -0,0 +1,330 @@
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<mods:title id="B50DF617EE7F24EB59E175277B46A851">New species and records of Batraxis Reitter (Coleoptera: Staphylinidae: Pselaphinae) in continental China</mods:title>
</mods:titleInfo>
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<mods:namePart id="01EC4D3664A355E334BDDF7228D0A17D">Wang, Dan</mods:namePart>
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<mods:namePart id="28AFCE573A47341C97B2ABB0E74A0ACE">Yin, Zi-Wei</mods:namePart>
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<mods:typeOfResource id="7C80E180C5F3E34E0838160673900269">text</mods:typeOfResource>
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<mods:date id="B37533F9476C672AF1DC041255822865">2016</mods:date>
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<treatment id="B60787DEFF8AFFE8FF39D041FC4E67EA" ID-DOI="http://doi.org/10.5281/zenodo.6077850" ID-GBIF-Taxon="123514483" ID-Zenodo-Dep="6077850" LSID="urn:lsid:plazi:treatment:B60787DEFF8AFFE8FF39D041FC4E67EA" httpUri="http://treatment.plazi.org/id/B60787DEFF8AFFE8FF39D041FC4E67EA" lastPageNumber="444" pageId="1" pageNumber="444">
<subSubSection id="76B46543FF8AFFE8FF39D041FEA461A4" pageId="1" pageNumber="444" type="nomenclature">
<paragraph id="3E1136C8FF8AFFE8FF39D041FE2F6178" blockId="1.[151,494,356,418]" box="[151,494,356,382]" pageId="1" pageNumber="444">
<heading id="655981A4FF8AFFE8FF39D041FE2F6178" bold="true" box="[151,494,356,382]" fontSize="10" level="2" pageId="1" pageNumber="444" reason="2">
<emphasis id="0CDAEADAFF8AFFE8FF39D041FE2F6178" bold="true" box="[151,494,356,382]" pageId="1" pageNumber="444">
<taxonomicName id="F9AE4D4BFF8AFFE8FF39D041FE96617B" ID-CoL="L29F" box="[151,343,356,381]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="0CDAEADAFF8AFFE8FF39D041FE96617B" bold="true" box="[151,343,356,381]" italics="true" pageId="1" pageNumber="444">Batraxis robusta</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="17E957A1FF8AFFE8FECAD041FE2F6178" box="[356,494,357,382]" pageId="1" pageNumber="444" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3E1136C8FF8AFFE8FF39D0ADFEA461A4" blockId="1.[151,494,356,418]" box="[151,357,393,418]" pageId="1" pageNumber="444">
(
<figureCitation id="A6952A4DFF8AFFE8FF31D0ADFF2461A7" box="[159,229,393,418]" captionStart="FIGURE 1" captionStartId="2.[151,250,1516,1538]" captionTargetBox="[161,1432,212,1483]" captionTargetId="figure@2.[151,1435,195,1494]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Batraxis robusta, male. A. Population from Jiangxi. B. Population from Guizhou. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256542/files/figure.png" pageId="1" pageNumber="444">Figs 1</figureCitation>
,
<figureCitation id="A6952A4DFF8AFFE8FF5DD0ADFEC061A7" box="[243,257,393,417]" captionStart="FIGURE 2" captionStartId="4.[151,250,1611,1633]" captionTargetBox="[162,1431,349,1576]" captionTargetId="figure@4.[151,1435,339,1589]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal habitus of Batraxis males. A. B. robusta, population from Guangxi. B. B. gloriosa, population from Hunan. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256543/files/figure.png" pageId="1" pageNumber="444">2</figureCitation>
A, 8, 15)
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8AFFE8FF39D0F0FEBC62F9" pageId="1" pageNumber="444" type="materials_examined">
<paragraph id="3E1136C8FF8AFFE8FF39D0F0FEBC62F9" blockId="1.[151,1437,468,2028]" pageId="1" pageNumber="444">
<materialsCitation id="8EC63C95FF8AFFE8FF39D0F0FB4D623B" ID-GBIF-Occurrence="1290996373" collectingDate="2013-07-19" collectionCode="SNUC" collectorName="Song &amp; Yin &amp; Yu" country="China" county="Pingxiang City" elevation="1370" latitude="27.460835" location="Tai" longLatPrecision="20" longitude="114.167496" municipality="Wugong Shan National Park" pageId="1" pageNumber="444" specimenCount="3" specimenCount-female="3" stateProvince="Jiangxi" typeStatus="holotype">
<emphasis id="0CDAEADAFF8AFFE8FF39D0F0FE8161EB" bold="true" box="[151,320,468,493]" pageId="1" pageNumber="444">
<typeStatus id="E115886AFF8AFFE8FF39D0F0FF1361EB" box="[151,210,468,493]" pageId="1" pageNumber="444">Type</typeStatus>
material
</emphasis>
(14 ƋƋ,
<specimenCount id="28A8FD41FF8AFFE8FE1AD0F1FE3261EB" box="[436,499,469,493]" pageId="1" pageNumber="444" type="female">3 ♀♀</specimenCount>
).
<emphasis id="0CDAEADAFF8AFFE8FDA1D0F0FD3761EB" bold="true" box="[527,758,468,493]" pageId="1" pageNumber="444">
<typeStatus id="E115886AFF8AFFE8FDA1D0F0FD4161EB" box="[527,640,468,493]" pageId="1" pageNumber="444" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="46B97658FF8AFFE8FD21D0F0FD3061EB" box="[655,753,468,493]" name="China" pageId="1" pageNumber="444">CHINA</collectingCountry>
:
</emphasis>
Ƌ, labeled ‘
<collectingCountry id="46B97658FF8AFFE8FC2AD0F1FC2761EB" box="[900,998,469,493]" name="China" pageId="1" pageNumber="444">CHINA</collectingCountry>
:
<collectingRegion id="FC6AF82AFF8AFFE8FC5DD0F0FB8661EB" box="[1011,1095,468,493]" country="China" name="Jiangxi" pageId="1" pageNumber="444">Jiangxi</collectingRegion>
,
<collectingCounty id="D7704E44FF8AFFE8FBFBD0F0FAC161EB" box="[1109,1280,468,493]" pageId="1" pageNumber="444">Pingxiang City</collectingCounty>
(萍乡市),
<collectingMunicipality id="DE75ACB2FF8AFFE8FF39D0D9FE146213" box="[151,469,508,533]" pageId="1" pageNumber="444">Wugong Shan National Park</collectingMunicipality>
(武功山国家森林公园),
<geoCoordinate id="5B9A500FFF8AFFE8FCA8D0D9FC4C6213" box="[774,909,509,533]" direction="north" orientation="latitude" pageId="1" pageNumber="444" precision="15" value="27.460835">27°27'39&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF8AFFE8FC36D0D9FBE96213" box="[920,1064,509,533]" direction="east" orientation="longitude" pageId="1" pageNumber="444" precision="15" value="114.167496">114°10'03&quot;E</geoCoordinate>
, cable way station to Dian-Jiang-
<location id="3B716013FF8AFFE8FF39D300FF7A623B" LSID="urn:lsid:plazi:treatment:B60787DEFF8AFFE8FF39D041FC4E67EA:3B716013FF8AFFE8FF39D300FF7A623B" box="[151,187,548,573]" country="China" county="Pingxiang City" latitude="27.460835" longLatPrecision="20" longitude="114.167496" municipality="Wugong Shan National Park" name="Tai" pageId="1" pageNumber="444" stateProvince="Jiangxi">Tai</location>
(点将台), broad leaf, sifted,
<quantity id="F9569B2DFF8AFFE8FDBCD301FD71623A" box="[530,688,549,573]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" metricValueMax="1.4" metricValueMin="1.34" pageId="1" pageNumber="444" unit="m" value="1370.0" valueMax="1400.0" valueMin="1340.0">
<elevation id="B583D1FBFF8AFFE8FDBCD301FD71623A" box="[530,688,549,573]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" metricValueMax="1.4" metricValueMin="1.34" pageId="1" pageNumber="444" unit="m" value="1370.0" valueMax="1400.0" valueMin="1340.0">1340–1400 m</elevation>
</quantity>
,
<date id="4A101008FF8AFFE8FD12D300FCFF623B" box="[700,830,548,573]" pageId="1" pageNumber="444" value="2013-07-19">
<collectingDate id="5A54E9E0FF8AFFE8FD12D300FCFF623B" box="[700,830,548,573]" pageId="1" pageNumber="444" value="2013-07-19">19.vii.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF8AFFE8FCE5D301FC46623B" box="[843,903,549,573]" pageId="1" pageNumber="444">Song</collectorName>
,
<collectorName id="935B531EFF8AFFE8FC3DD300FC7E623B" box="[915,959,548,573]" pageId="1" pageNumber="444">Yin</collectorName>
,
<collectorName id="935B531EFF8AFFE8FC64D301FC28623A" box="[970,1001,549,572]" pageId="1" pageNumber="444">Yu</collectorName>
leg. (
<collectionCode id="58BFAE0DFF8AFFE8FB84D301FBBE623B" box="[1066,1151,549,573]" pageId="1" pageNumber="444">SNUC</collectionCode>
)’
</materialsCitation>
.
<emphasis id="0CDAEADAFF8AFFE8FB39D300FA47623B" bold="true" box="[1175,1414,548,573]" pageId="1" pageNumber="444">
Paratypes:
<materialsCitation id="8EC63C95FF8AFFE8FAB1D300FA47623B" ID-GBIF-Occurrence="1290996392" box="[1311,1414,548,573]" collectionCode="SNUC" country="China" location="CHINA" pageId="1" pageNumber="444" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="holotype">
<collectingCountry id="46B97658FF8AFFE8FAB1D300FA41623B" box="[1311,1408,548,573]" name="China" pageId="1" pageNumber="444">CHINA</collectingCountry>
:
</materialsCitation>
</emphasis>
<specimenCount id="28A8FD41FF8AFFE8FA20D301FF026262" pageId="1" pageNumber="444" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="28A8FD41FF8AFFE8FF61D368FF3B6262" box="[207,250,588,612]" pageId="1" pageNumber="444" type="female">1 ♀</specimenCount>
, same label data as
<typeStatus id="E115886AFF8AFFE8FE70D368FD816262" box="[478,576,588,612]" pageId="1" pageNumber="444" type="holotype">holotype</typeStatus>
(
<collectionCode id="58BFAE0DFF8AFFE8FDFED368FD606262" box="[592,673,588,612]" pageId="1" pageNumber="444">SNUC</collectionCode>
);
<materialsCitation id="8EC63C95FF8AFFE8FD1AD368FD6C62B2" ID-GBIF-Occurrence="1290996357" collectingDate="2015-07-24" collectionCode="SNUC" collectorName="Jiuwan Shan N. R. &amp; Li &amp; Zhao" country="China" elevation="1200" latitude="25.195" location="Yangmei'ao" longLatPrecision="21" longitude="108.6475" municipality="Liuzhou City" pageId="1" pageNumber="444" specimenCount="8" specimenCount-female="2" specimenCount-male="6" stateProvince="Guangxi" typeStatus="holotype">
<specimenCount id="28A8FD41FF8AFFE8FD1AD368FD366262" box="[692,759,588,612]" pageId="1" pageNumber="444" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="28A8FD41FF8AFFE8FCADD368FC826262" box="[771,835,588,612]" pageId="1" pageNumber="444" type="female">2 ♀♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF8AFFE8FC00D368FC3B6262" box="[942,1018,588,612]" name="China" pageId="1" pageNumber="444">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF8AFFE8FBABD368FBA96262" box="[1029,1128,588,612]" country="China" name="Guangxi" pageId="1" pageNumber="444">Guangxi</collectingRegion>
,
<collectingMunicipality id="DE75ACB2FF8AFFE8FBDBD368FACB6262" box="[1141,1290,588,612]" pageId="1" pageNumber="444">Liuzhou City</collectingMunicipality>
(柳州市),
<collectorName id="935B531EFF8AFFE8FF39D350FEA1628A" box="[151,352,628,653]" pageId="1" pageNumber="444">Jiuwan Shan N.R.</collectorName>
(九万山自然保护区),
<location id="3B716013FF8AFFE8FDC9D350FD30628B" LSID="urn:lsid:plazi:treatment:B60787DEFF8AFFE8FF39D041FC4E67EA:3B716013FF8AFFE8FDC9D350FD30628B" box="[615,753,628,653]" country="China" latitude="25.195" longLatPrecision="21" longitude="108.6475" municipality="Liuzhou City" name="Yangmei'ao" pageId="1" pageNumber="444" stateProvince="Guangxi">Yangmei’ao</location>
(杨梅坳),
<geoCoordinate id="5B9A500FFF8AFFE8FCC3D350FC33628B" box="[877,1010,628,653]" direction="north" orientation="latitude" pageId="1" pageNumber="444" precision="15" value="25.195">25°11'42&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF8AFFE8FC53D350FB4F628B" box="[1021,1166,628,653]" direction="east" orientation="longitude" pageId="1" pageNumber="444" precision="15" value="108.6475">108°38'51&quot;E</geoCoordinate>
, mixed leaf litter, sifted,
<quantity id="F9569B2DFF8AFFE8FF39D3B8FF2F62B5" box="[151,238,668,692]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="444" unit="m" value="1200.0">
<elevation id="B583D1FBFF8AFFE8FF39D3B8FF2F62B5" box="[151,238,668,692]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="444" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="4A101008FF8AFFE8FF55D3BFFE4762B5" box="[251,390,667,691]" pageId="1" pageNumber="444" value="2015-07-24">
<collectingDate id="5A54E9E0FF8AFFE8FF55D3BFFE4762B5" box="[251,390,667,691]" pageId="1" pageNumber="444" value="2015-07-24">24.VII.2015</collectingDate>
</date>
,
<collectorName id="935B531EFF8AFFE8FE3DD3B8FE6D62B2" box="[403,428,668,692]" pageId="1" pageNumber="444">Li</collectorName>
&amp;
<collectorName id="935B531EFF8AFFE8FE7FD3B8FDCA62B2" box="[465,523,668,692]" pageId="1" pageNumber="444">Zhao</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF8AFFE8FDFAD3B8FD6462B2" box="[596,677,668,692]" pageId="1" pageNumber="444">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF8AFFE8FD17D3B8FEB862F9" ID-GBIF-Occurrence="1290996377" collectingDate="2015-07-20" collectionCode="SNUC" collectorName="Maolan N. R. &amp; He &amp; Hu &amp; Wang" country="China" elevation="800" latitude="25.26361" location="Libo Hsien" longLatPrecision="21" longitude="107.90723" pageId="1" pageNumber="444" specimenCount="4" specimenCount-male="4" stateProvince="Guizhou" typeStatus="holotype">
<specimenCount id="28A8FD41FF8AFFE8FD17D3B8FD3B62B5" box="[697,762,668,692]" pageId="1" pageNumber="444" type="male">4 ♂♂</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF8AFFE8FCCAD3B8FC7162B2" box="[868,944,668,692]" name="China" pageId="1" pageNumber="444">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF8AFFE8FC12D3B8FBDE62B2" box="[956,1055,668,692]" country="China" name="Guizhou" pageId="1" pageNumber="444">Guizhou</collectingRegion>
,
<location id="3B716013FF8AFFE8FB85D3B8FB6962B2" LSID="urn:lsid:plazi:treatment:B60787DEFF8AFFE8FF39D041FC4E67EA:3B716013FF8AFFE8FB85D3B8FB6962B2" box="[1067,1192,668,692]" country="China" latitude="25.26361" longLatPrecision="21" longitude="107.90723" name="Libo Hsien" pageId="1" pageNumber="444" stateProvince="Guizhou">Libo Hsien</location>
(荔波县),
<collectorName id="935B531EFF8AFFE8FAE8D3BFFF0D62DD" pageId="1" pageNumber="444">Maolan N.R.</collectorName>
(茂兰自然保护区),
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,
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, mixed leaf litter, sifted,
<quantity id="F9569B2DFF8AFFE8FBA9D3E7FB4862DD" box="[1031,1161,707,732]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="1" pageNumber="444" unit="m" value="800.0" valueMax="850.0" valueMin="750.0">
<elevation id="B583D1FBFF8AFFE8FBA9D3E7FB4862DD" box="[1031,1161,707,732]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="1" pageNumber="444" unit="m" value="800.0" valueMax="850.0" valueMin="750.0">750–850 m</elevation>
</quantity>
,
<date id="4A101008FF8AFFE8FB3BD3E7FAE162DA" box="[1173,1312,707,732]" pageId="1" pageNumber="444" value="2015-07-20">
<collectingDate id="5A54E9E0FF8AFFE8FB3BD3E7FAE162DA" box="[1173,1312,707,732]" pageId="1" pageNumber="444" value="2015-07-20">20.VII.2015</collectingDate>
</date>
,
<collectorName id="935B531EFF8AFFE8FA83D3E0FA9162DD" box="[1325,1360,708,731]" pageId="1" pageNumber="444">He</collectorName>
,
<collectorName id="935B531EFF8AFFE8FAF2D3E0FABF62DD" box="[1372,1406,708,731]" pageId="1" pageNumber="444">Hu</collectorName>
&amp;
<collectorName id="935B531EFF8AFFE8FF39D3CCFF196306" box="[151,216,744,768]" pageId="1" pageNumber="444">Wang</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF8AFFE8FE8ED3C3FEB062F9" box="[288,369,743,767]" pageId="1" pageNumber="444">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8AFFE8FF69D22FFCAA6389" pageId="1" pageNumber="444" type="diagnosis">
<paragraph id="3E1136C8FF8AFFE8FF69D22FFCAA6389" blockId="1.[151,1437,468,2028]" pageId="1" pageNumber="444">
<emphasis id="0CDAEADAFF8AFFE8FF69D22FFE6D6322" bold="true" box="[199,428,779,804]" pageId="1" pageNumber="444">Diagnosis of male.</emphasis>
Body covered with distinct pubescence; frons with complete subantennal transverse sulcus; antennomeres IX as long as wide, XI subconical; pronotum with distinct median antebasal fovea and basal impression; legs simple, all tibiae slender; tergite IV with narrow basal sulcus, discal carinae long, moderately close, with complete marginal carinae; aedeagus relatively stout.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8AFFE8FF69D2BFFB586715" pageId="1" pageNumber="444" type="description">
<paragraph id="3E1136C8FF8AFFE8FF69D2BFFCEC658E" blockId="1.[151,1437,468,2028]" pageId="1" pageNumber="444">
<emphasis id="0CDAEADAFF8AFFE8FF69D2BFFE9863B2" bold="true" box="[199,345,923,948]" pageId="1" pageNumber="444">Description.</emphasis>
Male (
<figureCitation id="A6952A4DFF8AFFE8FE02D2BFFE2C63B2" box="[428,493,923,948]" captionStart="FIGURE 1" captionStartId="2.[151,250,1516,1538]" captionTargetBox="[161,1432,212,1483]" captionTargetId="figure@2.[151,1435,195,1494]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Batraxis robusta, male. A. Population from Jiangxi. B. Population from Guizhou. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256542/files/figure.png" pageId="1" pageNumber="444">Fig. 1</figureCitation>
A). Body dark-brown, pubescent, length
<quantity id="F9569B2DFF8AFFE8FC6DD2BFFBAA63B5" box="[963,1131,923,948]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.69" metricValueMin="2.31" pageId="1" pageNumber="444" unit="mm" value="2.5" valueMax="2.69" valueMin="2.31">2.31‒2.69 mm</quantity>
. Head rectangular, slightly longer than wide, HL
<quantity id="F9569B2DFF8AFFE8FE3ED29BFDF463D1" box="[400,565,959,984]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="6.2" metricValueMin="4.9" pageId="1" pageNumber="444" unit="mm" value="0.5549999999999999" valueMax="0.62" valueMin="0.49">0.49‒0.62 mm</quantity>
, HW
<quantity id="F9569B2DFF8AFFE8FDD6D29BFCDC63D1" box="[632,797,959,984]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="1" pageNumber="444" unit="mm" value="0.51" valueMax="0.54" valueMin="0.48">0.48‒0.54 mm</quantity>
, vertex moderately convex, with pair of distinct vertexal foveae; antennal tubercles slightly raised; frons flattened, transverse subantennal sulcus complete, anterior margin of U-shaped vertexal sulcus merging with subantennal sulcus and connecting foveae; each eye composed of about 30 facets; antennae relatively thin, antennomeres III–VII elongate, VIII slightly elongate, IX as long as wide, X transverse, XI largest, widest at basal 1/3; median gular carina narrow, with two nude, separated gular foveae. Pronotum slightly wider than long, PL
<quantity id="F9569B2DFF8AFFE8FDF5D550FCC2648D" box="[603,771,1140,1164]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.9" metricValueMin="4.4" pageId="1" pageNumber="444" unit="mm" value="0.46499999999999997" valueMax="0.49" valueMin="0.44">0.44‒0.49 mm</quantity>
, PW
<quantity id="F9569B2DFF8AFFE8FCEAD557FC2A648D" box="[836,1003,1139,1164]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="5.8" metricValueMin="5.2" pageId="1" pageNumber="444" unit="mm" value="0.55" valueMax="0.58" valueMin="0.52">0.52‒0.58 mm</quantity>
, with distinct median antebasal fovea and basal impression. Elytra wider than long, EL
<quantity id="F9569B2DFF8AFFE8FD11D5B3FCA564A9" box="[703,868,1175,1200]" metricMagnitude="-4" metricUnit="m" metricValue="7.05" metricValueMax="7.7" metricValueMin="6.4" pageId="1" pageNumber="444" unit="mm" value="0.705" valueMax="0.77" valueMin="0.64">0.64‒0.77 mm</quantity>
, EW
<quantity id="F9569B2DFF8AFFE8FC0FD5B3FB8764A9" box="[929,1094,1175,1199]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" metricValueMax="9.8" metricValueMin="8.4" pageId="1" pageNumber="444" unit="mm" value="0.9099999999999999" valueMax="0.98" valueMin="0.84">0.84‒0.98 mm</quantity>
, each elytron with two distinct foveae at base. All legs simple, tibiae slender. Abdomen wider than long, AL
<quantity id="F9569B2DFF8AFFE8FBA5D59FFB7064D5" box="[1035,1201,1211,1236]" metricMagnitude="-4" metricUnit="m" metricValue="7.35" metricValueMax="8.8" metricValueMin="5.8999999999999995" pageId="1" pageNumber="444" unit="mm" value="0.735" valueMax="0.88" valueMin="0.59">0.59‒0.88 mm</quantity>
, AW
<quantity id="F9569B2DFF8AFFE8FB5CD598FA5964D5" box="[1266,1432,1212,1236]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" metricValueMax="9.2" metricValueMin="8.2" pageId="1" pageNumber="444" unit="mm" value="0.87" valueMax="0.92" valueMin="0.82">0.82‒0.92 mm</quantity>
, slightly narrowed at base; tergite IV (first visible tergite) with narrow basal sulcus, discal carinae long, relatively close, extending to apical two-thirds of tergal length, width between carinae
<quantity id="F9569B2DFF8AFFE8FBBDD427FB7B651D" box="[1043,1210,1283,1308]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="1" pageNumber="444" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.25‒0.30 mm</quantity>
at apices, marginal carinae complete. Aedeagus (
<figureCitation id="A6952A4DFF8AFFE8FE42D403FDF16539" box="[492,560,1319,1344]" captionStart="FIGURE 8" captionStartId="14.[151,250,1960,1982]" captionTargetBox="[191,1401,193,1911]" captionTargetId="figure@14.[169,1405,193,1918]" captionTargetPageId="14" captionText="FIGURE 8. Aedeagi of Batraxis robusta. (A, B, D, E, G, H. in ventral view. C, F, I. in lateral view). A – C. Population from Jiangxi. D – F. Population from Guizhou. G – I. Population from Guangxi. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256549/files/figure.png" pageId="1" pageNumber="444">Fig. 8</figureCitation>
A‒C)
<quantity id="F9569B2DFF8AFFE8FDD5D403FD226539" box="[635,739,1319,1343]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="444" unit="mm" value="0.4">0.40 mm</quantity>
long, relatively stout, parameres slightly exceeding apex of median lobe, rounded apically, each with three preapical setae; endophallus complicated, with series of symmetric sclerites, and numerous spine-like structures distomedially.
</paragraph>
<paragraph id="3E1136C8FF8AFFE8FF69D4B7FEF3661E" blockId="1.[151,1437,468,2028]" pageId="1" pageNumber="444">
Female. Similar to male in general appearance; each eye composed of about 15 facets. Measurements: BL
<quantity id="F9569B2DFF8AFFE8FF39D49CFEFC65C9" box="[151,317,1464,1488]" metricMagnitude="-3" metricUnit="m" metricValue="2.485" metricValueMax="2.7" metricValueMin="2.27" pageId="1" pageNumber="444" unit="mm" value="2.485" valueMax="2.7" valueMin="2.27">2.27‒2.70 mm</quantity>
; HL 0.5 5‒
<quantity id="F9569B2DFF8AFFE8FE1CD493FDDA65C9" box="[434,539,1463,1488]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="444" unit="mm" value="0.6">0.60 mm</quantity>
; HW
<quantity id="F9569B2DFF8AFFE8FDF2D493FCC365C9" box="[604,770,1463,1488]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="1" pageNumber="444" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50‒0.53 mm</quantity>
; PL
<quantity id="F9569B2DFF8AFFE8FC9BD49CFC1A65C9" box="[821,987,1464,1488]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="1" pageNumber="444" unit="mm" value="0.45999999999999996" valueMax="0.48" valueMin="0.44">0.44‒0.48 mm</quantity>
; PW
<quantity id="F9569B2DFF8AFFE8FBB9D49CFB7C65C9" box="[1047,1213,1463,1488]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="5.5" metricValueMin="5.2" pageId="1" pageNumber="444" unit="mm" value="0.535" valueMax="0.55" valueMin="0.52">0.52‒0.55 mm</quantity>
; EL
<quantity id="F9569B2DFF8AFFE8FB5CD493FA5965C9" box="[1266,1432,1463,1488]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="7.1" metricValueMin="6.2" pageId="1" pageNumber="444" unit="mm" value="0.665" valueMax="0.71" valueMin="0.62">0.62‒0.71 mm</quantity>
; EW
<quantity id="F9569B2DFF8AFFE8FF65D4FFFEB365F5" box="[203,370,1499,1524]" metricMagnitude="-4" metricUnit="m" metricValue="8.95" metricValueMax="9.3" metricValueMin="8.6" pageId="1" pageNumber="444" unit="mm" value="0.895" valueMax="0.93" valueMin="0.86">0.86‒0.93 mm</quantity>
; AL
<quantity id="F9569B2DFF8AFFE8FE02D4FFFD9265F5" box="[428,595,1499,1524]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="9.1" metricValueMin="6.7" pageId="1" pageNumber="444" unit="mm" value="0.79" valueMax="0.91" valueMin="0.67">0.67‒0.91 mm</quantity>
; AW
<quantity id="F9569B2DFF8AFFE8FD3AD4F8FCFA65F5" box="[660,827,1500,1524]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="8.8" metricValueMin="8.7" pageId="1" pageNumber="444" unit="mm" value="0.875" valueMax="0.88" valueMin="0.87">0.87‒0.88 mm</quantity>
; width between discal carinae of tergite IV
<quantity id="F9569B2DFF8AFFE8FA82D4F8FF026611" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="1" pageNumber="444" unit="mm" value="0.29" valueMax="0.3" valueMin="0.28">0.28‒0.30 mm</quantity>
at apices.
</paragraph>
<paragraph id="3E1136C8FF8AFFE8FF69D707FB586715" blockId="1.[151,1437,468,2028]" pageId="1" pageNumber="444">
<emphasis id="0CDAEADAFF8AFFE8FF69D707FE73663A" bold="true" box="[199,434,1571,1596]" pageId="1" pageNumber="444">Comparative notes.</emphasis>
<taxonomicName id="F9AE4D4BFF8AFFE8FE14D700FDB6663D" box="[442,631,1571,1595]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="0CDAEADAFF8AFFE8FE14D700FDB6663D" box="[442,631,1571,1595]" italics="true" pageId="1" pageNumber="444">Batraxis robusta</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="F9AE4D4BFF8AFFE8FCA1D700FBCC663A" authority="Wang &amp; Yin" authorityName="Wang &amp; Yin" box="[783,1037,1571,1596]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="0CDAEADAFF8AFFE8FCA1D700FCE1663D" box="[783,800,1572,1595]" italics="true" pageId="1" pageNumber="444">B</emphasis>
.
<emphasis id="0CDAEADAFF8AFFE8FC9ED707FCB5663D" box="[816,884,1571,1595]" italics="true" pageId="1" pageNumber="444">pilosa</emphasis>
Wang &amp; Yin
</taxonomicName>
by the long body pubescence. The new species can be distinguished from
<taxonomicName id="F9AE4D4BFF8AFFE8FDF5D76CFD016659" box="[603,704,1607,1631]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="0CDAEADAFF8AFFE8FDF5D76CFDAD6659" box="[603,620,1608,1631]" italics="true" pageId="1" pageNumber="444">B</emphasis>
.
<emphasis id="0CDAEADAFF8AFFE8FDD2D763FD016659" box="[636,704,1607,1631]" italics="true" pageId="1" pageNumber="444">pilosa</emphasis>
</taxonomicName>
by its large body (
<quantity id="F9569B2DFF8AFFE8FC33D763FBC86666" box="[925,1033,1607,1632]" metricMagnitude="-3" metricUnit="m" metricValue="2.61" pageId="1" pageNumber="444" unit="mm" value="2.61">2.61 mm</quantity>
), slightly raised antennal tubercles, more slender antennae, close discal carinae of tergite IV, simple protrochanters and protibiae, and complicated structures of the aedeagal endophallus. In
<taxonomicName id="F9AE4D4BFF8AFFE8FD2BD7B4FD2D66A1" box="[645,748,1679,1703]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="444" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="0CDAEADAFF8AFFE8FD2BD7B4FD5766A1" box="[645,662,1680,1703]" italics="true" pageId="1" pageNumber="444">B</emphasis>
.
<emphasis id="0CDAEADAFF8AFFE8FD06D7ABFD2D66A1" box="[680,748,1679,1703]" italics="true" pageId="1" pageNumber="444">pilosa</emphasis>
</taxonomicName>
, the body is smaller (
<quantity id="F9569B2DFF8AFFE8FC5ED7ABFB9C66AE" box="[1008,1117,1679,1704]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="1" pageNumber="444" unit="mm" value="2.05">2.05 mm</quantity>
), the antennal tubercles are distinctly raised, the antennae are thicker, the discal carinae of the tergite IV are widely separated, the ventral margin of each protrochanter is strongly protuberant, each protibia has a small preapical tooth, the aedeagus is more slender, and the aedeagal endophallus lacks distinct distomedian spine-like structures.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8AFFE8FF69D63BFDFC67A2" pageId="1" pageNumber="444" type="discussion">
<paragraph id="3E1136C8FF8AFFE8FF69D63BFDFC67A2" blockId="1.[151,1437,468,2028]" pageId="1" pageNumber="444">
<emphasis id="0CDAEADAFF8AFFE8FF69D63BFE8E673E" bold="true" box="[199,335,1823,1848]" pageId="1" pageNumber="444">Comments.</emphasis>
Males of the populations from
<collectingRegion id="FC6AF82AFF8AFFE8FD1CD63BFCC2673E" box="[690,771,1823,1848]" country="China" name="Jiangxi" pageId="1" pageNumber="444">Jiangxi</collectingRegion>
(
<figureCitation id="A6952A4DFF8AFFE8FCBCD63BFC9D673E" box="[786,860,1823,1848]" captionStart="FIGURE 1" captionStartId="2.[151,250,1516,1538]" captionTargetBox="[161,1432,212,1483]" captionTargetId="figure@2.[151,1435,195,1494]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Batraxis robusta, male. A. Population from Jiangxi. B. Population from Guizhou. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256542/files/figure.png" pageId="1" pageNumber="444">Figs. 1</figureCitation>
A, 8A‒C),
<collectingRegion id="FC6AF82AFF8AFFE8FC73D63BFBFC673E" box="[989,1085,1823,1848]" country="China" name="Guizhou" pageId="1" pageNumber="444">Guizhou</collectingRegion>
(
<figureCitation id="A6952A4DFF8AFFE8FBE3D63BFB59673E" box="[1101,1176,1823,1848]" captionStart="FIGURE 1" captionStartId="2.[151,250,1516,1538]" captionTargetBox="[161,1432,212,1483]" captionTargetId="figure@2.[151,1435,195,1494]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Batraxis robusta, male. A. Population from Jiangxi. B. Population from Guizhou. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256542/files/figure.png" pageId="1" pageNumber="444">Figs. 1</figureCitation>
B, 8D‒F) and
<collectingRegion id="FC6AF82AFF8AFFE8FA92D63BFA5D673E" box="[1340,1436,1823,1848]" country="China" name="Guangxi" pageId="1" pageNumber="444">Guangxi</collectingRegion>
(
<figureCitation id="A6952A4DFF8AFFE8FF31D667FF2D675A" box="[159,236,1859,1884]" captionStart="FIGURE 2" captionStartId="4.[151,250,1611,1633]" captionTargetBox="[162,1431,349,1576]" captionTargetId="figure@4.[151,1435,339,1589]" captionTargetPageId="4" captionText="FIGURE 2. Dorsal habitus of Batraxis males. A. B. robusta, population from Guangxi. B. B. gloriosa, population from Hunan. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256543/files/figure.png" pageId="1" pageNumber="444">Figs. 2</figureCitation>
A, 8G‒I) are slightly different in the form of the pronotal basal impression, in the width of the discal carinae on tergite IV, and in the lengths of the median sclerites of the aedeagal endophallus. These differences are interpreted as intraspecific variations.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8AFFE8FF69D68BFC9767CE" box="[199,854,1967,1992]" pageId="1" pageNumber="444" type="distribution">
<paragraph id="3E1136C8FF8AFFE8FF69D68BFC9767CE" blockId="1.[151,1437,468,2028]" box="[199,854,1967,1992]" pageId="1" pageNumber="444">
<emphasis id="0CDAEADAFF8AFFE8FF69D68BFEA167CE" bold="true" box="[199,352,1967,1992]" pageId="1" pageNumber="444">Distribution.</emphasis>
<collectingCountry id="46B97658FF8AFFE8FEC9D68BFE6F67CE" box="[359,430,1967,1992]" name="China" pageId="1" pageNumber="444">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF8AFFE8FE17D68BFDCC67CE" box="[441,525,1967,1992]" country="China" name="Jiangxi" pageId="1" pageNumber="444">Jiangxi</collectingRegion>
,
<collectingRegion id="FC6AF82AFF8AFFE8FDB6D68BFDBA67CE" box="[536,635,1967,1992]" country="China" name="Guizhou" pageId="1" pageNumber="444">Guizhou</collectingRegion>
,
<collectingRegion id="FC6AF82AFF8AFFE8FD28D68BFD2767CE" box="[646,742,1967,1992]" country="China" name="Guangxi" pageId="1" pageNumber="444">Guangxi</collectingRegion>
(
<figureCitation id="A6952A4DFF8AFFE8FD58D68BFC8867CE" box="[758,841,1967,1992]" captionStart="FIGURE 15" captionStartId="21.[151,250,1023,1045]" captionTargetBox="[151,1436,193,1001]" captionTargetId="figure@21.[151,1436,193,1002]" captionTargetPageId="21" captionText="FIGURE 15. Distribution of Batraxis species in China." httpUri="https://zenodo.org/record/256556/files/figure.png" pageId="1" pageNumber="444">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8AFFE8FF69D6F7FC4E67EA" box="[199,911,2003,2028]" pageId="1" pageNumber="444" type="etymology">
<paragraph id="3E1136C8FF8AFFE8FF69D6F7FC4E67EA" blockId="1.[151,1437,468,2028]" box="[199,911,2003,2028]" pageId="1" pageNumber="444">
<emphasis id="0CDAEADAFF8AFFE8FF69D6F7FE8F67EA" bold="true" box="[199,334,2003,2028]" pageId="1" pageNumber="444">Etymology.</emphasis>
The specific epithet refers to the robust body form.
</paragraph>
</subSubSection>
</treatment>
</document>

255
data/B6/07/87/B60787DEFF8CFFE1FF39D1B3FB8163A3.xml Normal file
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@ -0,0 +1,255 @@
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<mods:title id="49334A0C5A12AFB1120F11884B236056">New species and records of Batraxis Reitter (Coleoptera: Staphylinidae: Pselaphinae) in continental China</mods:title>
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<taxonomicName id="F9AE4D4BFF8CFFEEFF39D1B3FE8260B7" ID-CoL="68G73" box="[151,323,151,177]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="450" phylum="Arthropoda" rank="species" species="gesar" status="sp. nov.">
<emphasis id="0CDAEADAFF8CFFEEFF39D1B3FE8260B7" bold="true" box="[151,323,151,177]" italics="true" pageId="7" pageNumber="450">Batraxis gesar</emphasis>
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,
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(
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A, 11D–F, 14C, 14H, 14P, 15)
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<subSubSection id="76B46543FF8CFFEEFF39D022FD9F61B0" pageId="7" pageNumber="450" type="materials_examined">
<paragraph id="3E1136C8FF8CFFEEFF39D022FD9F61B0" blockId="7.[151,1436,261,654]" pageId="7" pageNumber="450">
<materialsCitation id="8EC63C95FF8CFFEEFF39D022FAC86145" ID-GBIF-Occurrence="1290996365" collectingDate="2010-08-17" collectionCode="SNUC" collectorName="Jian-Qing Zhu" country="China" county="Cuona County" elevation="2500" latitude="27.825556" location="Lebu" longLatPrecision="20" longitude="91.75972" pageId="7" pageNumber="450" specimenCount="2" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="0CDAEADAFF8CFFEEFF39D022FEFC6119" bold="true" box="[151,317,262,287]" pageId="7" pageNumber="450">
<typeStatus id="E115886AFF8CFFEEFF39D022FF136119" box="[151,210,262,287]" pageId="7" pageNumber="450">Type</typeStatus>
material
</emphasis>
(3 ƋƋ).
<emphasis id="0CDAEADAFF8CFFEEFE0DD022FD466119" bold="true" box="[419,647,262,287]" pageId="7" pageNumber="450">
<typeStatus id="E115886AFF8CFFEEFE0DD022FDD56119" box="[419,532,262,287]" pageId="7" pageNumber="450" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="46B97658FF8CFFEEFD8ED022FD406119" box="[544,641,262,287]" name="China" pageId="7" pageNumber="450">CHINA</collectingCountry>
:
</emphasis>
Ƌ, labeled ‘
<collectingCountry id="46B97658FF8CFFEEFCA2D023FCAF6119" box="[780,878,263,287]" name="China" pageId="7" pageNumber="450">CHINA</collectingCountry>
:
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,
<collectingCounty id="D7704E44FF8CFFEEFC79D023FBBA6119" box="[983,1147,263,287]" pageId="7" pageNumber="450">Cuona County</collectingCounty>
(错那县),
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(勒布),
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,
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, leaf litter, sifted,
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,
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<collectingDate id="5A54E9E0FF8CFFEEFCBED00EFC5B6145" box="[784,922,298,323]" pageId="7" pageNumber="450" value="2010-08-17">17.viii.2010</collectingDate>
</date>
,
<collectorName id="935B531EFF8CFFEEFC03D00EFB966145" box="[941,1111,298,323]" pageId="7" pageNumber="450">Jian-Qing Zhu</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF8CFFEEFB1ED00FFAC06145" box="[1200,1281,299,323]" pageId="7" pageNumber="450">SNUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8EC63C95FF8CFFEEFAB5D00EFE556194" ID-GBIF-Occurrence="1290996378" collectingDate="2013-06-18" collectingDateMax="2013-06-22" collectingDateMin="2013-06-18" collectionCode="SNUC" collectorName="Xuan Bi" country="China" county="Cuona County" elevation="2750" location="Lexiang" municipality="China" pageId="7" pageNumber="450" specimenCount="1" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="0CDAEADAFF8CFFEEFAB5D00EFF3C616D" bold="true" pageId="7" pageNumber="450">
<typeStatus id="E115886AFF8CFFEEFAB5D00EFA566145" box="[1307,1431,298,323]" pageId="7" pageNumber="450" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="46B97658FF8CFFEEFF39D076FF38616D" box="[151,249,338,363]" name="China" pageId="7" pageNumber="450">CHINA</collectingCountry>
:
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1 Ƌ, labeled ‘
<collectingMunicipality id="DE75ACB2FF8CFFEEFE32D076FE26616D" box="[412,487,338,363]" pageId="7" pageNumber="450">China</collectingMunicipality>
:
<collectingRegion id="FC6AF82AFF8CFFEEFE5DD076FD86616D" box="[499,583,338,363]" country="China" name="Xizang" pageId="7" pageNumber="450">Xizang</collectingRegion>
,
<collectingCounty id="D7704E44FF8CFFEEFDFCD076FD36616C" box="[594,759,338,362]" pageId="7" pageNumber="450">Cuona County</collectingCounty>
,
<location id="3B716013FF8CFFEEFCACD076FC9F616D" LSID="urn:lsid:plazi:treatment:B60787DEFF8CFFE1FF39D1B3FB8163A3:3B716013FF8CFFEEFCACD076FC9F616D" box="[770,862,338,363]" country="China" county="Cuona County" municipality="China" name="Lexiang" pageId="7" pageNumber="450" stateProvince="Xizang">Lexiang</location>
(勒乡), alt.
<quantity id="F9569B2DFF8CFFEEFBABD076FB62616C" box="[1029,1187,338,362]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="7" pageNumber="450" unit="m" value="2750.0" valueMax="2800.0" valueMin="2700.0">
<elevation id="B583D1FBFF8CFFEEFBABD076FB62616C" box="[1029,1187,338,362]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="7" pageNumber="450" unit="m" value="2750.0" valueMax="2800.0" valueMin="2700.0">2700–2800 m</elevation>
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,
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</date>
, Wen-
<collectorName id="935B531EFF8CFFEEFF39D05EFF346194" box="[151,245,378,402]" pageId="7" pageNumber="450">Xuan Bi</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF8CFFEEFE95D05EFE4D6194" box="[315,396,378,402]" pageId="7" pageNumber="450">SNUC</collectionCode>
)
</materialsCitation>
;
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<specimenCount id="28A8FD41FF8CFFEEFE31D05EFE0B6197" box="[415,458,378,402]" pageId="7" pageNumber="450" type="male">1 ♂</specimenCount>
, labeled ‘
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:
<collectingRegion id="FC6AF82AFF8CFFEEFD29D05DFD196194" box="[647,728,377,402]" country="China" name="Xizang" pageId="7" pageNumber="450">Xizang</collectingRegion>
A.R.,
<collectingCounty id="D7704E44FF8CFFEEFC8ED05DFC0E6194" box="[800,975,377,402]" pageId="7" pageNumber="450">Yadong County</collectingCounty>
(亚东县),
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(下亚东),
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</quantity>
,
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<collectingDate id="5A54E9E0FF8CFFEEFF39D0B9FED261B0" box="[151,275,413,438]" pageId="7" pageNumber="450" value="2010-08-08">8.viii.2010</collectingDate>
</date>
,
<collectorName id="935B531EFF8CFFEEFEB0D0BBFE7961B0" box="[286,440,413,438]" pageId="7" pageNumber="450">Wen-Xuan Bi</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF8CFFEEFDAFD0BAFD9361B0" box="[513,594,414,438]" pageId="7" pageNumber="450">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8CFFEEFF69D0E5FDD26288" pageId="7" pageNumber="450" type="diagnosis">
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<emphasis id="0CDAEADAFF8CFFEEFF69D0E5FE7561DC" bold="true" box="[199,436,449,474]" pageId="7" pageNumber="450">Diagnosis of male.</emphasis>
Body almost glabrous, abdomen sparsely with short setae; frons with complete subantennal transverse sulcus; antennomeres IX transverse, XI protuberant at inner basal margins; pronotum with distinct median antebasal fovea and basal impression; all tibiae slender; protrochanters protuberant ventrally, profemora with large spine at ventral margins, protibiae strongly projected before middle, mesotibiae simple; tergite IV with broad, abrupt basal sulcus, discal carinae short, relatively widely separated, with complete marginal carinae; aedeagus relatively stout.
</paragraph>
</subSubSection>
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<paragraph id="3E1136C8FF8CFFEEFF39D69FFBB767D7" blockId="7.[151,1142,1979,2001]" box="[151,1142,1979,2001]" pageId="7" pageNumber="450">
<emphasis id="0CDAEADAFF8CFFEEFF39D69FFED367D7" bold="true" box="[151,274,1979,2001]" pageId="7" pageNumber="450">FIGURE 4.</emphasis>
Dorsal habitus of
<taxonomicName id="F9AE4D4BFF8CFFEEFE63D69FFDE267D6" box="[461,547,1979,2000]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="0CDAEADAFF8CFFEEFE63D69FFDE267D6" box="[461,547,1979,2000]" italics="true" pageId="7" pageNumber="450">Batraxis</emphasis>
</taxonomicName>
males.
<emphasis id="0CDAEADAFF8CFFEEFDDFD69FFD4B67D7" bold="true" box="[625,650,1979,2001]" pageId="7" pageNumber="450">A.</emphasis>
<taxonomicName id="F9AE4D4BFF8CFFEEFD3ED698FD2267D7" box="[656,739,1980,2001]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="450" phylum="Arthropoda" rank="species" species="gesar">
<emphasis id="0CDAEADAFF8CFFEEFD3ED698FD2267D7" box="[656,739,1980,2001]" italics="true" pageId="7" pageNumber="450">B. gesar</emphasis>
</taxonomicName>
.
<emphasis id="0CDAEADAFF8CFFEEFD5ED69FFCC767D7" bold="true" box="[752,774,1979,2001]" pageId="7" pageNumber="450">B.</emphasis>
<taxonomicName id="F9AE4D4BFF8CFFEEFCA3D698FC6167D6" box="[781,928,1979,2001]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="450" phylum="Arthropoda" rank="species" species="longithorax">
<emphasis id="0CDAEADAFF8CFFEEFCA3D698FC6167D6" box="[781,928,1979,2001]" italics="true" pageId="7" pageNumber="450">B. longithorax</emphasis>
</taxonomicName>
. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="76B46543FF83FFE1FF69D1B3FE90635B" pageId="8" pageNumber="451" type="description">
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<emphasis id="0CDAEADAFF83FFE1FF69D1B3FE9860B6" bold="true" box="[199,345,151,176]" pageId="8" pageNumber="451">Description.</emphasis>
Male (
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A). Body almost glabrous, reddish-brown, length
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. Head rectangular, slightly longer than wide, HL
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, HW
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, vertex glabrous, apical margin of frons and clypeus sparsely pubescent; vertex moderately convex, with pair of distinct vertexal foveae; antennal tubercles low; frons flattened and broad, subantennal transverse sulcus straight and complete, lacking vertexal sulcus; each eye composed of about 30 facets; antennae thick, antennomeres III elongate, IV and V as long as wise, VI‒X transverse, XI largest, protuberant at base; median gular carina shallow, with two separated nude foveae. Pronotum slightly wider than long, PL
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, PW
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, with distinct median antebasal fovea and complete basal impression. Elytra wider than long, EL
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, EW
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, each elytron with two distinct foveae at base. Protrochanters protuberant at ventral margin (
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P), protibiae strongly projected before middle (
<figureCitation id="A6952A4DFF83FFE1FAC1D093FF7561F2" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="8" pageNumber="451">Fig. 14</figureCitation>
C), profemora with large middle spine at ventral margins (
<figureCitation id="A6952A4DFF83FFE1FCF7D0F8FC6C61F2" box="[857,941,476,501]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="8" pageNumber="451">Fig. 14</figureCitation>
H). Abdomen slightly wider than long, AL
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, AW
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, slightly narrowed at base; tergite IV (first visible tergite) sparsely pubescent, with broad, abrupt basal sulcus; discal carinae short, relatively widely separated, extending to apical one-fifth of tergal length, width between carinae
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at apices; with complete marginal carinae. Aedeagus (
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D–F)
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in length, relatively stout, parameres slightly exceeding apex of median lobe, rounded apically, each with two apical setae and hook-like protuberance; endophallus with pair of symmetric sclerites, and weakly sclerotized structures at apex.
</paragraph>
<paragraph id="3E1136C8FF83FFE1FF69D3F3FE5F62F6" blockId="8.[151,1437,151,933]" box="[199,414,727,752]" pageId="8" pageNumber="451">Female. Unknown.</paragraph>
<paragraph id="3E1136C8FF83FFE1FF69D3D8FE90635B" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="451">
<emphasis id="0CDAEADAFF83FFE1FF69D3D8FE736313" bold="true" box="[199,434,764,789]" pageId="8" pageNumber="451">Comparative notes.</emphasis>
<taxonomicName id="F9AE4D4BFF83FFE1FE15D3D9FDA06312" box="[443,609,765,788]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="451" phylum="Arthropoda" rank="species" species="gesar">
<emphasis id="0CDAEADAFF83FFE1FE15D3D9FDA06312" box="[443,609,765,788]" italics="true" pageId="8" pageNumber="451">Batraxis gesar</emphasis>
</taxonomicName>
is the only known species with strongly modified antennomeres XI and fore legs in the male, when combined with the aedeagal structures and distribution, it can be readily separated from other congeners.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF83FFE1FF69D243FDBB6386" box="[199,634,871,896]" pageId="8" pageNumber="451" type="distribution">
<paragraph id="3E1136C8FF83FFE1FF69D243FDBB6386" blockId="8.[151,1437,151,933]" box="[199,634,871,896]" pageId="8" pageNumber="451">
<emphasis id="0CDAEADAFF83FFE1FF69D243FEA16386" bold="true" box="[199,352,871,896]" pageId="8" pageNumber="451">Distribution.</emphasis>
<collectingCountry id="46B97658FF83FFE1FEC9D243FE6F6386" box="[359,430,871,896]" name="China" pageId="8" pageNumber="451">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF83FFE1FE17D243FDCB6386" box="[441,522,871,896]" country="China" name="Xizang" pageId="8" pageNumber="451">Xizang</collectingRegion>
(
<figureCitation id="A6952A4DFF83FFE1FDB7D243FDAD6386" box="[537,620,871,896]" captionStart="FIGURE 15" captionStartId="21.[151,250,1023,1045]" captionTargetBox="[151,1436,193,1001]" captionTargetId="figure@21.[151,1436,193,1002]" captionTargetPageId="21" captionText="FIGURE 15. Distribution of Batraxis species in China." httpUri="https://zenodo.org/record/256556/files/figure.png" pageId="8" pageNumber="451">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF83FFE1FF69D2A8FB8163A3" box="[199,1088,908,933]" pageId="8" pageNumber="451" type="etymology">
<paragraph id="3E1136C8FF83FFE1FF69D2A8FB8163A3" blockId="8.[151,1437,151,933]" box="[199,1088,908,933]" pageId="8" pageNumber="451">
<emphasis id="0CDAEADAFF83FFE1FF69D2A8FE8F63A3" bold="true" box="[199,334,908,933]" pageId="8" pageNumber="451">Etymology.</emphasis>
The specific epithet refers to King
<taxonomicName id="F9AE4D4BFF83FFE1FD74D2A8FCDC63A2" box="[730,797,908,932]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="464" phylum="Arthropoda" rank="species" species="gesar" status="sp. nov.">Gesar</taxonomicName>
, a culture hero of
<collectingRegion id="FC6AF82AFF83FFE1FC46D2A8FBFC63A3" box="[1000,1085,908,933]" country="China" name="Xizang" pageId="8" pageNumber="451">Xizang</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

253
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<emphasis id="0CDAEADAFF8DFFEFFF39D397FE4C62CB" bold="true" box="[151,397,691,717]" italics="true" pageId="6" pageNumber="449">Batraxis longicornis</emphasis>
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,
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(
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B, 11A‒C, 14O, 15)
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<typeStatus id="E115886AFF8DFFEFFF39D23AFF136331" box="[151,210,798,823]" pageId="6" pageNumber="449">Type</typeStatus>
material
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(7 ƋƋ,
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:
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:
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Ƌ, labeled ‘
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,
<location id="3B716013FF8DFFEFFF56D266FE04635C" LSID="urn:lsid:plazi:treatment:B60787DEFF8DFFEFFF39D397FC3367DC:3B716013FF8DFFEFFF56D266FE04635C" box="[248,453,834,859]" country="China" county="Tengchong County" latitude="25.29611" longLatPrecision="21" longitude="98.799446" municipality="Baoshan City" name="Mingguang Town" pageId="6" pageNumber="449" stateProvince="Yunnan">Mingguang Town</location>
,
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,
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,
<geoCoordinate id="5B9A500FFF8DFFEFFCA4D266FC4C635D" box="[778,909,834,859]" direction="east" orientation="longitude" pageId="6" pageNumber="449" precision="15" value="98.799446">98°47'58&quot;E</geoCoordinate>
,
<quantity id="F9569B2DFF8DFFEFFC32D266FC37635C" box="[924,1014,834,859]" metricMagnitude="3" metricUnit="m" metricValue="1.61" pageId="6" pageNumber="449" unit="m" value="1610.0">
<elevation id="B583D1FBFF8DFFEFFC32D266FC37635C" box="[924,1014,834,859]" metricMagnitude="3" metricUnit="m" metricValue="1.61" pageId="6" pageNumber="449" unit="m" value="1610.0">1610 m</elevation>
</quantity>
,
<date id="4A101008FF8DFFEFFBABD266FBBC635D" box="[1029,1149,834,859]" pageId="6" pageNumber="449" value="2013-04-20">
<collectingDate id="5A54E9E0FF8DFFEFFBABD266FBBC635D" box="[1029,1149,834,859]" pageId="6" pageNumber="449" value="2013-04-20">20.iv.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF8DFFEFFB25D266FB77635D" box="[1163,1206,834,859]" pageId="6" pageNumber="449">Dai</collectorName>
,
<collectorName id="935B531EFF8DFFEFFB6BD267FB3E635D" box="[1221,1279,835,859]" pageId="6" pageNumber="449">Song</collectorName>
&amp;
<collectorName id="935B531EFF8DFFEFFA86D267FAA1635D" box="[1320,1376,835,859]" pageId="6" pageNumber="449">Peng</collectorName>
leg.’ (
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)
</materialsCitation>
.
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<emphasis id="0CDAEADAFF8DFFEFFEAFD242FE316379" bold="true" box="[257,496,870,895]" pageId="6" pageNumber="449">
<typeStatus id="E115886AFF8DFFEFFEAFD242FEBC6379" box="[257,381,870,895]" pageId="6" pageNumber="449" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="46B97658FF8DFFEFFE27D242FE2A6379" box="[393,491,870,895]" name="China" pageId="6" pageNumber="449">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="28A8FD41FF8DFFEFFE58D242FDF76378" box="[502,566,870,894]" pageId="6" pageNumber="449" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="28A8FD41FF8DFFEFFDEFD243FD4E6378" box="[577,655,871,895]" pageId="6" pageNumber="449" type="female">14 ♀♀</specimenCount>
, same label data as
<typeStatus id="E115886AFF8DFFEFFCC5D242FC106378" box="[875,977,870,894]" pageId="6" pageNumber="449" type="holotype">holotype</typeStatus>
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;
<materialsCitation id="8EC63C95FF8DFFEFFC75D243FEF063C1" ID-GBIF-Occurrence="1290996370" collectingDate="2013-04-22" collectionCode="SNUC" collectorName="Dai &amp; Song &amp; Peng" country="China" county="Baoshan City" elevation="1220" latitude="25.2775" location="Baihua Village" longLatPrecision="21" longitude="98.79806" municipality="Baihualing" pageId="6" pageNumber="449" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="28A8FD41FF8DFFEFFC75D243FBDA6378" box="[987,1051,871,895]" pageId="6" pageNumber="449" type="female">3 ♀♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF8DFFEFFB2DD242FB0E6378" box="[1155,1231,870,894]" name="China" pageId="6" pageNumber="449">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF8DFFEFFB77D243FAF36378" box="[1241,1330,871,894]" country="China" name="Yunnan" pageId="6" pageNumber="449">Yunnan</collectingRegion>
,
<collectingCounty id="D7704E44FF8DFFEFFA92D242FF0963A5" pageId="6" pageNumber="449">Baoshan City</collectingCounty>
,
<collectingMunicipality id="DE75ACB2FF8DFFEFFF78D2AEFE9563A5" box="[214,340,906,931]" pageId="6" pageNumber="449">Baihualing</collectingMunicipality>
,
<location id="3B716013FF8DFFEFFECCD2AEFDCE63A5" LSID="urn:lsid:plazi:treatment:B60787DEFF8DFFEFFF39D397FC3367DC:3B716013FF8DFFEFFECCD2AEFDCE63A5" box="[354,527,906,931]" country="China" county="Baoshan City" latitude="25.2775" longLatPrecision="21" longitude="98.79806" municipality="Baihualing" name="Baihua Village" pageId="6" pageNumber="449" stateProvince="Yunnan">Baihua Village</location>
, near stream,
<geoCoordinate id="5B9A500FFF8DFFEFFD1CD2AEFCF963A5" box="[690,824,906,931]" direction="north" orientation="latitude" pageId="6" pageNumber="449" precision="15" value="25.2775">25°16'39&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF8DFFEFFCE9D2AEFC0B63A5" box="[839,970,906,931]" direction="east" orientation="longitude" pageId="6" pageNumber="449" precision="15" value="98.79806">98°47'53&quot;E</geoCoordinate>
,
<quantity id="F9569B2DFF8DFFEFFC77D2AFFBF363A4" box="[985,1074,907,931]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="6" pageNumber="449" unit="m" value="1220.0">
<elevation id="B583D1FBFF8DFFEFFC77D2AFFBF363A4" box="[985,1074,907,931]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="6" pageNumber="449" unit="m" value="1220.0">1220 m</elevation>
</quantity>
,
<date id="4A101008FF8DFFEFFBEFD2AEFB7B63A5" box="[1089,1210,906,931]" pageId="6" pageNumber="449" value="2013-04-22">
<collectingDate id="5A54E9E0FF8DFFEFFBEFD2AEFB7B63A5" box="[1089,1210,906,931]" pageId="6" pageNumber="449" value="2013-04-22">22.iv.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF8DFFEFFB66D2AEFB3263A5" box="[1224,1267,906,931]" pageId="6" pageNumber="449">Dai</collectorName>
,
<collectorName id="935B531EFF8DFFEFFAAFD2AFFAFA63A5" box="[1281,1339,907,931]" pageId="6" pageNumber="449">Song</collectorName>
&amp;
<collectorName id="935B531EFF8DFFEFFACAD2AFFA5D63A5" box="[1380,1436,907,931]" pageId="6" pageNumber="449">Peng</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF8DFFEFFF76D28BFEE863C1" box="[216,297,943,967]" pageId="6" pageNumber="449">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8DFFEFFF69D2F6FF3C6499" pageId="6" pageNumber="449" type="diagnosis">
<paragraph id="3E1136C8FF8DFFEFFF69D2F6FF3C6499" blockId="6.[151,1437,798,2011]" pageId="6" pageNumber="449">
<emphasis id="0CDAEADAFF8DFFEFFF69D2F6FE7363ED" bold="true" box="[199,434,978,1003]" pageId="6" pageNumber="449">Diagnosis of male.</emphasis>
Body almost glabrous, abdomen covered with sparse setae; frons with complete subantennal transverse sulcus; antennomeres IX as long as wide, XI lengthily elongate; pronotum with shallow median antebasal fovea and distinct basal impression; all tibiae slender; protrochanters protuberant ventrally, profemora simple, protibiae lacking protuberance, mesotibiae lacking apical spur; tergite IV with narrow basal sulcus, discal carinae relatively short, moderately close, with relatively short marginal carinae; aedeagus relatively elongate.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8DFFEFFF69D58EFD266795" pageId="6" pageNumber="449" type="description">
<paragraph id="3E1136C8FF8DFFEFFF69D58EFA4566BC" blockId="6.[151,1437,798,2011]" pageId="6" pageNumber="449">
<emphasis id="0CDAEADAFF8DFFEFFF69D58EFE9864C5" bold="true" box="[199,345,1194,1219]" pageId="6" pageNumber="449">Description.</emphasis>
Male (
<figureCitation id="A6952A4DFF8DFFEFFE04D58EFE2C64C5" box="[426,493,1194,1219]" captionStart="FIGURE 3" captionStartId="5.[151,250,1435,1457]" captionTargetBox="[159,1425,198,1391]" captionTargetId="figure@5.[151,1435,193,1413]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus of Batraxis males. A. B. antennata. B. B. longicornis. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256544/files/figure.png" pageId="6" pageNumber="449">Fig. 3</figureCitation>
B). Body yellowish brown, length
<quantity id="F9569B2DFF8DFFEFFCD8D58EFBDF64C4" box="[886,1054,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="2.04" metricValueMax="2.06" metricValueMin="2.02" pageId="6" pageNumber="449" unit="mm" value="2.04" valueMax="2.06" valueMin="2.02">2.02‒2.06 mm</quantity>
. Head rectangular, slightly longer than wide, HL
<quantity id="F9569B2DFF8DFFEFFEE8D5EAFE2C64E0" box="[326,493,1230,1255]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.699999999999999" metricValueMin="4.5" pageId="6" pageNumber="449" unit="mm" value="0.45999999999999996" valueMax="0.47" valueMin="0.45">0.45‒0.47 mm</quantity>
, HW
<quantity id="F9569B2DFF8DFFEFFD9DD5EBFD1B64E0" box="[563,730,1231,1255]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.4" metricValueMin="4.3" pageId="6" pageNumber="449" unit="mm" value="0.435" valueMax="0.44" valueMin="0.43">0.43‒0.44 mm</quantity>
, vertex glabrous, apical margin of frons and clypeus sparsely pubescent; vertex moderately convex, with pair of distinct vertexal foveae; antennal tubercles low; frons flattened and broad, subantennal transverse sulcus complete, anterior margin of U-shaped vertexal sulcus merging with subantennal transverse sulcus and connecting foveae; each eye composed of about 30 facets; antennae relatively thin, antennomeres III–VII slightly elongate, VIII and IX as long as wide, X slightly transverse, XI largest and elongate, widest at basal third; median gular carina narrow, with two separated nude gular foveae. Pronotum slightly wider than long, PL
<quantity id="F9569B2DFF8DFFEFFE7AD483FDFB65B8" box="[468,570,1447,1471]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="449" unit="mm" value="0.38">0.38 mm</quantity>
, PW
<quantity id="F9569B2DFF8DFFEFFDDBD482FCDB65B8" box="[629,794,1446,1471]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.699999999999999" metricValueMin="4.5" pageId="6" pageNumber="449" unit="mm" value="0.45999999999999996" valueMax="0.47" valueMin="0.45">0.45‒0.47 mm</quantity>
, with shallow median antebasal fovea and complete basal impression. Elytra wider than long, EL
<quantity id="F9569B2DFF8DFFEFFDE1D4EEFD7465E4" box="[591,693,1482,1507]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="449" unit="mm" value="0.55">0.55 mm</quantity>
, EW
<quantity id="F9569B2DFF8DFFEFFD5CD4EFFC5765E4" box="[754,918,1483,1507]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" metricValueMax="7.4" metricValueMin="7.199999999999999" pageId="6" pageNumber="449" unit="mm" value="0.73" valueMax="0.74" valueMin="0.72">0.72‒0.74 mm</quantity>
, each elytron with two shallow foveae at base. Protrochanters protuberant at ventral margin (
<figureCitation id="A6952A4DFF8DFFEFFD0BD4CAFD386601" box="[677,761,1518,1543]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="6" pageNumber="449">Fig. 14</figureCitation>
O). Abdomen slightly wider than long, AL
<quantity id="F9569B2DFF8DFFEFFB5ED4CAFA596600" box="[1264,1432,1518,1543]" metricMagnitude="-4" metricUnit="m" metricValue="6.45" metricValueMax="6.6" metricValueMin="6.3" pageId="6" pageNumber="449" unit="mm" value="0.645" valueMax="0.66" valueMin="0.63">0.63‒0.66 mm</quantity>
, AW
<quantity id="F9569B2DFF8DFFEFFF64D736FEAF662C" box="[202,366,1554,1579]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" metricValueMax="6.7" metricValueMin="6.5" pageId="6" pageNumber="449" unit="mm" value="0.66" valueMax="0.67" valueMin="0.65">0.65‒0.67 mm</quantity>
, slightly expanded at base; tergite IV (first visible tergite) sparsely pubescent, with narrow basal sulcus; discal carinae relatively short and close, extending to one-third of tergal length, width between carinae
<quantity id="F9569B2DFF8DFFEFFAC5D712FF026674" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="449" unit="mm" value="0.25">0.25 mm</quantity>
at apices; marginal carinae extending to apical one-third of tergal length. Aedeagus (
<figureCitation id="A6952A4DFF8DFFEFFB2AD77EFB126675" box="[1156,1235,1626,1651]" captionStart="FIGURE 11" captionStartId="17.[151,250,1967,1989]" captionTargetBox="[190,1403,193,1944]" captionTargetId="figure@17.[178,1407,193,1953]" captionTargetPageId="17" captionText="FIGURE 11. Aedeagi of Batraxis species. (A, B, D, E, G, H. in ventral view. C, F, I. in lateral view). A – C. B. longicornis. D – F. B. gesar. G – I. B. longithorax. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256552/files/figure.png" pageId="6" pageNumber="449">Fig. 11</figureCitation>
A‒C)
<quantity id="F9569B2DFF8DFFEFFAB5D77FFABE6674" box="[1307,1407,1627,1651]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="449" unit="mm" value="0.31">0.31 mm</quantity>
in length, relatively elongate, parameres distinctly exceeding apex of median lobe, apex flattened, each with three thick apical setae; endophallus with two pairs of symmetric, elongate sclerites and weakly sclerotized structures.
</paragraph>
<paragraph id="3E1136C8FF8DFFEFFF69D7E2FDDD674D" blockId="6.[151,1437,798,2011]" pageId="6" pageNumber="449">
Female. Similar to male in general appearance; protrochanters simple; each eye composed of about 30 facets. Measurements: BL
<quantity id="F9569B2DFF8DFFEFFEDDD7CEFDD86704" box="[371,537,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="2.195" metricValueMax="2.35" metricValueMin="2.04" pageId="6" pageNumber="449" unit="mm" value="2.195" valueMax="2.35" valueMin="2.04">2.04‒2.35 mm</quantity>
; HL
<quantity id="F9569B2DFF8DFFEFFDFFD7CFFD376704" box="[593,758,1771,1795]" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.699999999999999" metricValueMin="4.4" pageId="6" pageNumber="449" unit="mm" value="0.45499999999999996" valueMax="0.47" valueMin="0.44">0.44‒0.47 mm</quantity>
; HW
<quantity id="F9569B2DFF8DFFEFFC99D7CEFC1C6704" box="[823,989,1770,1795]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="6" pageNumber="449" unit="mm" value="0.44" valueMax="0.45" valueMin="0.43">0.43‒0.45 mm</quantity>
; PL
<quantity id="F9569B2DFF8DFFEFFBBED7CFFB776704" box="[1040,1206,1771,1795]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="6" pageNumber="449" unit="mm" value="0.405" valueMax="0.42" valueMin="0.39">0.39‒0.42 mm</quantity>
; PW
<quantity id="F9569B2DFF8DFFEFFB5CD7CEFA596704" box="[1266,1432,1770,1795]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="449" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.40‒0.50 mm</quantity>
; EL
<quantity id="F9569B2DFF8DFFEFFF6CD62AFEAB6720" box="[194,362,1806,1831]" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.5" metricValueMin="5.2" pageId="6" pageNumber="449" unit="mm" value="0.585" valueMax="0.65" valueMin="0.52">0.52‒0.65 mm</quantity>
; EW
<quantity id="F9569B2DFF8DFFEFFE02D62AFD946720" box="[428,597,1806,1831]" metricMagnitude="-4" metricUnit="m" metricValue="7.949999999999999" metricValueMax="8.5" metricValueMin="7.4" pageId="6" pageNumber="449" unit="mm" value="0.7949999999999999" valueMax="0.85" valueMin="0.74">0.74‒0.85 mm</quantity>
; AL
<quantity id="F9569B2DFF8DFFEFFD3ED62AFCF96720" box="[656,824,1806,1831]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.1" metricValueMin="6.8999999999999995" pageId="6" pageNumber="449" unit="mm" value="0.75" valueMax="0.81" valueMin="0.69">0.69‒0.81 mm</quantity>
; AW
<quantity id="F9569B2DFF8DFFEFFCD5D62AFBE56720" box="[891,1060,1806,1831]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.9" metricValueMin="6.6" pageId="6" pageNumber="449" unit="mm" value="0.725" valueMax="0.79" valueMin="0.66">0.66‒0.79 mm</quantity>
; width between discal carinae of tergite IV
<quantity id="F9569B2DFF8DFFEFFEA7D616FE6D674C" box="[265,428,1842,1867]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="6" pageNumber="449" unit="mm" value="0.275" valueMax="0.3" valueMin="0.25">0.25‒0.30 mm</quantity>
at apices.
</paragraph>
<paragraph id="3E1136C8FF8DFFEFFF69D672FD266795" blockId="6.[151,1437,798,2011]" pageId="6" pageNumber="449">
<emphasis id="0CDAEADAFF8DFFEFFF69D672FE726769" bold="true" box="[199,435,1878,1903]" pageId="6" pageNumber="449">Comparative notes.</emphasis>
<taxonomicName id="F9AE4D4BFF8DFFEFFE12D673FD656768" box="[444,676,1878,1902]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="449" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="0CDAEADAFF8DFFEFFE12D673FD656768" box="[444,676,1878,1902]" italics="true" pageId="6" pageNumber="449">Batraxis longicornis</emphasis>
</taxonomicName>
can be readily separated from all other congeners by the strongly elongate antennomeres XI and form of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8DFFEFFF69D6BAFDBF67B1" box="[199,638,1950,1975]" pageId="6" pageNumber="449" type="distribution">
<paragraph id="3E1136C8FF8DFFEFFF69D6BAFDBF67B1" blockId="6.[151,1437,798,2011]" box="[199,638,1950,1975]" pageId="6" pageNumber="449">
<emphasis id="0CDAEADAFF8DFFEFFF69D6BAFEA167B1" bold="true" box="[199,352,1950,1975]" pageId="6" pageNumber="449">Distribution.</emphasis>
<collectingCountry id="46B97658FF8DFFEFFEC9D6BAFE6F67B1" box="[359,430,1950,1975]" name="China" pageId="6" pageNumber="449">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF8DFFEFFE17D6BBFDCF67B0" box="[441,526,1951,1974]" country="China" name="Yunnan" pageId="6" pageNumber="449">Yunnan</collectingRegion>
(
<figureCitation id="A6952A4DFF8DFFEFFDB0D6BAFDB067B1" box="[542,625,1950,1975]" captionStart="FIGURE 15" captionStartId="21.[151,250,1023,1045]" captionTargetBox="[151,1436,193,1001]" captionTargetId="figure@21.[151,1436,193,1002]" captionTargetPageId="21" captionText="FIGURE 15. Distribution of Batraxis species in China." httpUri="https://zenodo.org/record/256556/files/figure.png" pageId="6" pageNumber="449">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8DFFEFFF69D6E6FC3367DC" box="[199,1010,1986,2011]" pageId="6" pageNumber="449" type="etymology">
<paragraph id="3E1136C8FF8DFFEFFF69D6E6FC3367DC" blockId="6.[151,1437,798,2011]" box="[199,1010,1986,2011]" pageId="6" pageNumber="449">
<emphasis id="0CDAEADAFF8DFFEFFF69D6E6FE8F67DD" bold="true" box="[199,334,1986,2011]" pageId="6" pageNumber="449">Etymology.</emphasis>
The specific epithet refers to the elongate antennomeres XI.
</paragraph>
</subSubSection>
</treatment>
</document>

271
data/B6/07/87/B60787DEFF8FFFEFFF39D7EEFC3C6259.xml Normal file
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@ -0,0 +1,271 @@
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<mods:title id="A1F365C16089ED2DA85B823FA24DEA65">New species and records of Batraxis Reitter (Coleoptera: Staphylinidae: Pselaphinae) in continental China</mods:title>
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<paragraph id="3E1136C8FF8FFFEDFF39D7EEFDDC66E3" blockId="4.[151,541,1738,1798]" box="[151,541,1738,1765]" pageId="4" pageNumber="447">
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<taxonomicName id="F9AE4D4BFF8FFFEDFF39D7EEFEBE66E2" ID-CoL="L27X" box="[151,383,1738,1764]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="447" phylum="Arthropoda" rank="species" species="antennata" status="sp. nov.">
<emphasis id="0CDAEADAFF8FFFEDFF39D7EEFEBE66E2" bold="true" box="[151,383,1738,1764]" italics="true" pageId="4" pageNumber="447">Batraxis antennata</emphasis>
</taxonomicName>
,
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(
<figureCitation id="A6952A4DFF8FFFEDFF31D7C9FF256700" box="[159,228,1773,1798]" captionStart="FIGURE 3" captionStartId="5.[151,250,1435,1457]" captionTargetBox="[159,1425,198,1391]" captionTargetId="figure@5.[151,1435,193,1413]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal habitus of Batraxis males. A. B. antennata. B. B. longicornis. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256544/files/figure.png" pageId="4" pageNumber="447">Figs 3</figureCitation>
A, 10G‒I, 15)
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8FFFEDFF39D61DFD4D67EF" pageId="4" pageNumber="447" type="materials_examined">
<paragraph id="3E1136C8FF8FFFEDFF39D61DFD4D67EF" blockId="4.[151,1436,1849,2026]" pageId="4" pageNumber="447">
<materialsCitation id="8EC63C95FF8FFFEDFF39D61DFD006798" ID-GBIF-Occurrence="1290996385" collectingDate="2013-04-26" collectionCode="SNUC" collectorName="Dai &amp; Song &amp; Peng" country="China" county="Tengchong County" elevation="1900" latitude="25.729445" location="Mingguang Town" longLatPrecision="21" longitude="98.57139" municipality="Baoshan City" pageId="4" pageNumber="447" specimenCount="4" specimenCount-female="4" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="0CDAEADAFF8FFFEDFF39D61DFE836754" bold="true" box="[151,322,1849,1874]" pageId="4" pageNumber="447">
<typeStatus id="E115886AFF8FFFEDFF39D61DFF136754" box="[151,210,1849,1874]" pageId="4" pageNumber="447">Type</typeStatus>
material
</emphasis>
(2 ƋƋ,
<specimenCount id="28A8FD41FF8FFFEDFE03D61EFE2F6754" box="[429,494,1850,1874]" pageId="4" pageNumber="447" type="female">4 ♀♀</specimenCount>
).
<emphasis id="0CDAEADAFF8FFFEDFDA2D61DFD346754" bold="true" box="[524,757,1849,1874]" pageId="4" pageNumber="447">
<typeStatus id="E115886AFF8FFFEDFDA2D61DFDBC6754" box="[524,637,1849,1874]" pageId="4" pageNumber="447" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="46B97658FF8FFFEDFD20D61DFD2E6754" box="[654,751,1849,1874]" name="China" pageId="4" pageNumber="447">CHINA</collectingCountry>
:
</emphasis>
Ƌ, labeled ‘
<collectingCountry id="46B97658FF8FFFEDFC27D61EFC2A6754" box="[905,1003,1850,1874]" name="China" pageId="4" pageNumber="447">CHINA</collectingCountry>
:
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,
<collectingMunicipality id="DE75ACB2FF8FFFEDFBCCD61DFB3F6754" box="[1122,1278,1849,1874]" pageId="4" pageNumber="447">Baoshan City</collectingMunicipality>
(保山市),
<collectingCounty id="D7704E44FF8FFFEDFF39D645FEAD677C" box="[151,364,1889,1914]" pageId="4" pageNumber="447">Tengchong County</collectingCounty>
(腾冲县),
<location id="3B716013FF8FFFEDFE45D645FD73677F" LSID="urn:lsid:plazi:treatment:B60787DEFF8FFFEFFF39D7EEFC3C6259:3B716013FF8FFFEDFE45D645FD73677F" box="[491,690,1889,1914]" country="China" county="Tengchong County" latitude="25.729445" longLatPrecision="21" longitude="98.57139" municipality="Baoshan City" name="Mingguang Town" pageId="4" pageNumber="447" stateProvince="Yunnan">Mingguang Town</location>
(明光镇),
<location id="3B716013FF8FFFEDFC9DD645FC04677C" LSID="urn:lsid:plazi:treatment:B60787DEFF8FFFEFFF39D7EEFC3C6259:3B716013FF8FFFEDFC9DD645FC04677C" box="[819,965,1889,1914]" country="China" county="Tengchong County" latitude="25.729445" longLatPrecision="21" longitude="98.57139" municipality="Baoshan City" name="Zizhi Village" pageId="4" pageNumber="447" stateProvince="Yunnan">Zizhi Village</location>
(自治村),
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,
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,
<quantity id="F9569B2DFF8FFFEDFACAD646FF6E679B" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="447" unit="m" value="1900.0">
<elevation id="B583D1FBFF8FFFEDFACAD646FF6E679B" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="447" unit="m" value="1900.0">1900 m</elevation>
</quantity>
,
<date id="4A101008FF8FFFEDFF10D6A1FEF76798" box="[190,310,1925,1950]" pageId="4" pageNumber="447" value="2013-04-26">
<collectingDate id="5A54E9E0FF8FFFEDFF10D6A1FEF76798" box="[190,310,1925,1950]" pageId="4" pageNumber="447" value="2013-04-26">26.iv.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF8FFFEDFEEBD6A1FEB16798" box="[325,368,1925,1950]" pageId="4" pageNumber="447">Dai</collectorName>
,
<collectorName id="935B531EFF8FFFEDFED0D6A2FE796798" box="[382,440,1926,1950]" pageId="4" pageNumber="447">Song</collectorName>
&amp;
<collectorName id="935B531EFF8FFFEDFE4CD6A2FDDB6798" box="[482,538,1926,1950]" pageId="4" pageNumber="447">Peng</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF8FFFEDFDC6D6A2FD786798" box="[616,697,1926,1950]" pageId="4" pageNumber="447">SNUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8EC63C95FF8FFFEDFD61D6A1FF3667C3" ID-GBIF-Occurrence="1290996391" collectionCode="SNUC" country="China" location="CHINA" pageId="4" pageNumber="447" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="0CDAEADAFF8FFFEDFD61D6A1FC016798" bold="true" box="[719,960,1925,1950]" pageId="4" pageNumber="447">
<typeStatus id="E115886AFF8FFFEDFD61D6A1FC8A6798" box="[719,843,1925,1950]" pageId="4" pageNumber="447" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="46B97658FF8FFFEDFCF4D6A1FC7A6798" box="[858,955,1925,1950]" name="China" pageId="4" pageNumber="447">CHINA</collectingCountry>
:
</emphasis>
1 Ƌ,
<specimenCount id="28A8FD41FF8FFFEDFBA7D6A2FB8D679B" box="[1033,1100,1926,1950]" pageId="4" pageNumber="447" type="female">3 ♀♀</specimenCount>
, same label data as
<typeStatus id="E115886AFF8FFFEDFA94D6A1FA5D6798" box="[1338,1436,1925,1950]" pageId="4" pageNumber="447" type="holotype">holotype</typeStatus>
(
<collectionCode id="58BFAE0DFF8FFFEDFF31D689FF3167C3" box="[159,240,1965,1989]" pageId="4" pageNumber="447">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8EC63C95FF8FFFEDFEADD689FD4967EF" ID-GBIF-Occurrence="1290996384" collectingDate="2013-04-20" collectionCode="SNUC" collectorName="Song &amp; Peng &amp; Dai" country="China" elevation="1700" latitude="25.295" location="Baihualing" longLatPrecision="21" longitude="98.79694" municipality="Baoshan City" pageId="4" pageNumber="447" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="28A8FD41FF8FFFEDFEADD689FEED67C2" box="[259,300,1965,1989]" pageId="4" pageNumber="447" type="female">1 ♀</specimenCount>
, labeled ‘
<collectingCountry id="46B97658FF8FFFEDFE38D689FE2367C3" box="[406,482,1965,1989]" name="China" pageId="4" pageNumber="447">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF8FFFEDFE42D689FD8467C2" box="[492,581,1965,1988]" country="China" name="Yunnan" pageId="4" pageNumber="447">Yunnan</collectingRegion>
,
<collectingMunicipality id="DE75ACB2FF8FFFEDFDE1D689FD2967C3" box="[591,744,1965,1989]" pageId="4" pageNumber="447">Baoshan City</collectingMunicipality>
,
<location id="3B716013FF8FFFEDFD5DD689FCAF67C3" LSID="urn:lsid:plazi:treatment:B60787DEFF8FFFEFFF39D7EEFC3C6259:3B716013FF8FFFEDFD5DD689FCAF67C3" box="[755,878,1965,1989]" country="China" latitude="25.295" longLatPrecision="21" longitude="98.79694" municipality="Baoshan City" name="Baihualing" pageId="4" pageNumber="447" stateProvince="Yunnan">Baihualing</location>
(百花岭), alt.
<quantity id="F9569B2DFF8FFFEDFB8ED689FBB667C3" box="[1056,1143,1965,1989]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="447" unit="m" value="1700.0">
<elevation id="B583D1FBFF8FFFEDFB8ED689FBB667C3" box="[1056,1143,1965,1989]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="447" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
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,
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,
<date id="4A101008FF8FFFEDFF39D6F5FECE67EC" box="[151,271,2001,2026]" pageId="4" pageNumber="447" value="2013-04-20">
<collectingDate id="5A54E9E0FF8FFFEDFF39D6F5FECE67EC" box="[151,271,2001,2026]" pageId="4" pageNumber="447" value="2013-04-20">20.iv.2013</collectingDate>
</date>
,
<collectorName id="935B531EFF8FFFEDFEB5D6F5FE9667EF" box="[283,343,2001,2025]" pageId="4" pageNumber="447">Song</collectorName>
,
<collectorName id="935B531EFF8FFFEDFECCD6F6FE5B67EC" box="[354,410,2002,2026]" pageId="4" pageNumber="447">Peng</collectorName>
&amp;
<collectorName id="935B531EFF8FFFEDFE10D6F5FE2767EC" box="[446,486,2001,2026]" pageId="4" pageNumber="447">Dai</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF8FFFEDFD81D6F5FD4167EF" box="[559,640,2001,2025]" pageId="4" pageNumber="447">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="6AD16640FF8EFFECFF39D4BFFB6065B7" box="[151,1185,1435,1457]" httpUri="https://zenodo.org/record/256544/files/figure.png" pageId="5" pageNumber="448" targetBox="[159,1425,198,1391]" targetPageId="5">
<paragraph id="3E1136C8FF8EFFECFF39D4BFFB6065B7" blockId="5.[151,1185,1435,1457]" box="[151,1185,1435,1457]" pageId="5" pageNumber="448">
<emphasis id="0CDAEADAFF8EFFECFF39D4BFFED365B7" bold="true" box="[151,274,1435,1457]" pageId="5" pageNumber="448">FIGURE 3.</emphasis>
Dorsal habitus of
<taxonomicName id="F9AE4D4BFF8EFFECFE63D4B8FDE265B7" box="[461,547,1436,1457]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="0CDAEADAFF8EFFECFE63D4B8FDE265B7" box="[461,547,1436,1457]" italics="true" pageId="5" pageNumber="448">Batraxis</emphasis>
</taxonomicName>
males.
<emphasis id="0CDAEADAFF8EFFECFDDFD4BFFD4B65B7" bold="true" box="[625,650,1435,1457]" pageId="5" pageNumber="448">A.</emphasis>
<taxonomicName id="F9AE4D4BFF8EFFECFD3ED4B8FCD065B7" box="[656,785,1436,1457]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="antennata">
<emphasis id="0CDAEADAFF8EFFECFD3ED4B8FCD065B7" box="[656,785,1436,1457]" italics="true" pageId="5" pageNumber="448">B. antennata</emphasis>
</taxonomicName>
.
<emphasis id="0CDAEADAFF8EFFECFCB0D4BFFCF465B7" bold="true" box="[798,821,1435,1457]" pageId="5" pageNumber="448">B.</emphasis>
<taxonomicName id="F9AE4D4BFF8EFFECFC92D4B8FC0A65B6" box="[828,971,1435,1457]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="448" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="0CDAEADAFF8EFFECFC92D4B8FC0A65B6" box="[828,971,1435,1457]" italics="true" pageId="5" pageNumber="448">B. longicornis</emphasis>
</taxonomicName>
. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="76B46543FF8EFFECFF69D4FCFAD9665A" pageId="5" pageNumber="448" type="diagnosis">
<paragraph id="3E1136C8FF8EFFECFF69D4FCFAD9665A" blockId="5.[151,1437,1496,2025]" pageId="5" pageNumber="448">
<emphasis id="0CDAEADAFF8EFFECFF69D4FCFE6A65F7" bold="true" box="[199,427,1496,1521]" pageId="5" pageNumber="448">Diagnosis of male.</emphasis>
Body almost glabrous, elytra with sparse, short setae; frons with abrupt subantennal transverse sulcus; antennomeres IX slightly elongate, XI concaved at apicolateral margins; pronotum with distinct median antebasal fovea and basal impression; all legs simple, tibiae slender; tergite IV with narrow basal sulcus, discal carinae relatively short, moderately close, with relatively short marginal carinae; aedeagus stout.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8EFFEFFF69D74CFEBF6211" lastPageId="6" lastPageNumber="449" pageId="5" pageNumber="448" type="description">
<paragraph id="3E1136C8FF8EFFEFFF69D74CFC8060F1" blockId="5.[151,1437,1496,2025]" lastBlockId="6.[151,1437,151,608]" lastPageId="6" lastPageNumber="449" pageId="5" pageNumber="448">
<emphasis id="0CDAEADAFF8EFFECFF69D74CFE986687" bold="true" box="[199,345,1640,1665]" pageId="5" pageNumber="448">Description.</emphasis>
Male (
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A). Body dark-brown, length
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. Head rectangular, slightly wider than long, HL
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, HW
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, vertex glabrous, apical margin of frons and clypeus sparsely pubescent; vertex moderately convex, with pair of small but distinct vertexal foveae; antennal tubercles low; frons flattened, subantennal transverse sulcus abrupt, vertexal sulcus short; each eye composed of about 30 facets; antennae thin, antennomeres III–IX elongate, X about as long as wide, XI largest, widest at middle, distinctly concaved at anterolateral margins; median gular carina narrow, with two separated nude gular foveae. Pronotum slightly wider than long, PL
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, PW
<quantity id="F9569B2DFF8EFFECFDAFD665FDAC675E" box="[513,621,1857,1881]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="448" unit="mm" value="0.43">0.43 mm</quantity>
, with distinct median antebasal fovea and complete V-shaped basal impression. Elytra wider than long, EL
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, EW
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, dorsal surface with sparse, minute setae, each elytron with two distinct foveae at base. All legs simple. Abdomen slightly wider than long, AL
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, AW
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, slightly narrowed at base; tergite IV (first visible tergite) with narrow basal sulcus; discal carinae relatively short, close, extending to one-fourth of tergal length, width between carinae
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at apices; marginal carinae extending to apical one-third of tergal length. Aedeagus (
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G‒I)
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in length, stout, parameres slightly shorter than median lobe, each with two short apical setae; endophallus without distinct sclerites, comprising a series of symmetric, weakly-sclerotized setae-like structures.
</paragraph>
<paragraph id="3E1136C8FF8DFFEFFF69D020FEF3618E" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="449">
Female. Similar to male in general appearance; each eye composed of about 30 facets. Measurements: BL
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; HL 0.4 3‒
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; HW
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; PL
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; PW
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; EL
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; EW
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; AL
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; AW
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; width between discal carinae of tergite IV
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at apices.
</paragraph>
<paragraph id="3E1136C8FF8DFFEFFF69D0B0FEBF6211" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="449">
<emphasis id="0CDAEADAFF8DFFEFFF69D0B0FE7561AB" bold="true" box="[199,436,404,429]" pageId="6" pageNumber="449">Comparative notes.</emphasis>
<taxonomicName id="F9AE4D4BFF8DFFEFFE11D0B1FD5861AA" box="[447,665,405,428]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="449" phylum="Arthropoda" rank="species" species="antennata">
<emphasis id="0CDAEADAFF8DFFEFFE11D0B1FD5861AA" box="[447,665,405,428]" italics="true" pageId="6" pageNumber="449">Batraxis antennata</emphasis>
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is a very distinct species among its Chinese congeners. It can be readily separated from all other species by the antennomeres XI distinctly concaved at the apicolateral margins, the presence of a V-shaped sulcus on the pronotal base, and the aedeagal endophallus has numerous weakly-sclerotized setae-like structures.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8DFFEFFF69D300FDBF623B" box="[199,638,548,573]" pageId="6" pageNumber="449" type="distribution">
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<emphasis id="0CDAEADAFF8DFFEFFF69D300FEA1623B" bold="true" box="[199,352,548,573]" pageId="6" pageNumber="449">Distribution.</emphasis>
<collectingCountry id="46B97658FF8DFFEFFEC9D300FE6F623B" box="[359,430,548,573]" name="China" pageId="6" pageNumber="449">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF8DFFEFFE17D301FDCF623A" box="[441,526,549,572]" country="China" name="Yunnan" pageId="6" pageNumber="449">Yunnan</collectingRegion>
(
<figureCitation id="A6952A4DFF8DFFEFFDB0D300FDB0623B" box="[542,625,548,573]" captionStart="FIGURE 15" captionStartId="21.[151,250,1023,1045]" captionTargetBox="[151,1436,193,1001]" captionTargetId="figure@21.[151,1436,193,1002]" captionTargetPageId="21" captionText="FIGURE 15. Distribution of Batraxis species in China." httpUri="https://zenodo.org/record/256556/files/figure.png" pageId="6" pageNumber="449">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF8DFFEFFF69D363FC3C6259" box="[199,1021,583,608]" pageId="6" pageNumber="449" type="etymology">
<paragraph id="3E1136C8FF8DFFEFFF69D363FC3C6259" blockId="6.[151,1437,151,608]" box="[199,1021,583,608]" pageId="6" pageNumber="449">
<emphasis id="0CDAEADAFF8DFFEFFF69D363FE8F6266" bold="true" box="[199,334,583,608]" pageId="6" pageNumber="449">Etymology.</emphasis>
The specific epithet refers to the concaved antennomeres XI.
</paragraph>
</subSubSection>
</treatment>
</document>

215
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<heading id="655981A4FF98FFFAFF39D775FD9A666A" bold="true" box="[151,603,1617,1644]" fontSize="11" level="1" pageId="19" pageNumber="462" reason="1">
<taxonomicName id="F9AE4D4BFF98FFFAFF39D775FD9A666A" ID-CoL="L29N" authority="Wang and Yin, 2016" authorityName="Wang and Yin" authorityYear="2016" box="[151,603,1617,1644]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="462" phylum="Arthropoda" rank="species" species="tibialis">
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<emphasis id="0CDAEADAFF98FFFAFF39D775FE97666D" bold="true" box="[151,342,1617,1643]" italics="true" pageId="19" pageNumber="462">Batraxis tibialis</emphasis>
Wang and Yin, 2016
</emphasis>
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<paragraph id="3E1136C8FF98FFFAFF39D750FDD9668B" blockId="19.[151,603,1617,1677]" box="[151,536,1652,1677]" pageId="19" pageNumber="462">
(
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,
<figureCitation id="A6952A4DFF98FFFAFF5DD751FECD668B" box="[243,268,1653,1677]" captionStart="FIGURE 13" captionStartId="19.[151,250,1495,1517]" captionTargetBox="[185,1406,200,1454]" captionTargetId="figure@19.[174,1410,193,1463]" captionTargetPageId="19" captionText="FIGURE 13. Aedeagi of Batraxis species. (A, B, D, E. in ventral view. C, F. in lateral view). A – C. B. bicolor, population from Yunnan. D – F. B. tibialis, population from Jiangxi. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256554/files/figure.png" pageId="19" pageNumber="462">13</figureCitation>
D–F, 14F,
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,
<date id="4A101008FF98FFFAFE19D751FE26668B" box="[439,487,1653,1677]" pageId="19" pageNumber="462" value="2014-05">14V</date>
, 15)
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<emphasis id="0CDAEADAFF98FFFAFF39D799FEF866D7" box="[151,313,1724,1746]" italics="true" pageId="19" pageNumber="462">Batraxis tibialis</emphasis>
Wang and Yin (in Wang
<emphasis id="0CDAEADAFF98FFFAFD97D799FDA766D7" box="[569,614,1724,1746]" italics="true" pageId="19" pageNumber="462">et al</emphasis>
. 2016: 4)
</taxonomicName>
.
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locality:
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,
<location id="3B716013FF98FFFAFD3DD7FFFD3766F7" LSID="urn:lsid:plazi:treatment:B60787DEFF98FFFDFF39D775FC296163:3B716013FF98FFFAFD3DD7FFFD3766F7" box="[659,758,1755,1777]" country="China" latitude="19.167778" longLatPrecision="21" longitude="109.74583" municipality="Qiongzhong Hsien" name="Limu Mt." pageId="19" pageNumber="462" stateProvince="Hainan">Limu Mt.</location>
,
<geoCoordinate id="5B9A500FFF98FFFAFCAED7FFFCBB66F7" box="[768,890,1755,1777]" direction="north" orientation="latitude" pageId="19" pageNumber="462" precision="15" value="19.167778">19°10'04&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF98FFFAFC2AD7FFFBC666F7" box="[900,1031,1755,1777]" direction="east" orientation="longitude" pageId="19" pageNumber="462" precision="15" value="109.74583">109°44'45&quot;E</geoCoordinate>
,
<specimenCount id="28A8FD41FF98FFFAFBBCD7FFFB9866F7" box="[1042,1113,1755,1777]" pageId="19" pageNumber="462" type="generic">
<quantity id="F9569B2DFF98FFFAFBBCD7FFFB9466F7" box="[1042,1109,1755,1777]" metricMagnitude="2" metricUnit="m" metricValue="6.25" pageId="19" pageNumber="462" unit="m" value="625.0">
<elevation id="B583D1FBFF98FFFAFBBCD7FFFB9466F7" box="[1042,1109,1755,1777]" metricMagnitude="2" metricUnit="m" metricValue="6.25" pageId="19" pageNumber="462" unit="m" value="625.0">625 m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="3E1136C8FF98FFFAFF39D605FF3D6780" blockId="19.[151,1437,1825,2034]" pageId="19" pageNumber="462">
<materialsCitation id="8EC63C95FF98FFFAFF39D605FF3D6780" ID-GBIF-Occurrence="1290996354" collectingDate="2014-07-24" collectionCode="SNUC" collectorName="Chen &amp; Hu &amp; Lv &amp; Yu" country="China" county="Ji'an City" elevation="580" latitude="26.5175" location="Bijia Shan" longLatPrecision="20" longitude="114.18806" municipality="Jinggang Shan" pageId="19" pageNumber="462" specimenCount="1" stateProvince="Jiangxi">
<emphasis id="0CDAEADAFF98FFFAFF39D605FDC4673C" bold="true" box="[151,517,1825,1850]" pageId="19" pageNumber="462">Additional material examined.</emphasis>
2 ƋƋ, labeled ‘
<collectingCountry id="46B97658FF98FFFAFD16D606FCDB673C" box="[696,794,1826,1850]" name="China" pageId="19" pageNumber="462">CHINA</collectingCountry>
: W.
<collectingRegion id="FC6AF82AFF98FFFAFCE5D605FC5E673C" box="[843,927,1825,1850]" country="China" name="Jiangxi" pageId="19" pageNumber="462">Jiangxi</collectingRegion>
,
<collectingCounty id="D7704E44FF98FFFAFC05D605FBDB673C" box="[939,1050,1825,1850]" pageId="19" pageNumber="462">Ji’an City</collectingCounty>
,
<collectingMunicipality id="DE75ACB2FF98FFFAFB88D605FB0A673C" box="[1062,1227,1825,1850]" pageId="19" pageNumber="462">Jinggang Shan</collectingMunicipality>
(井冈山),
<location id="3B716013FF98FFFAFACBD605FF0E6764" LSID="urn:lsid:plazi:treatment:B60787DEFF98FFFDFF39D775FC296163:3B716013FF98FFFAFACBD605FF0E6764" country="China" county="Ji'an City" latitude="26.5175" longLatPrecision="20" longitude="114.18806" municipality="Jinggang Shan" name="Bijia Shan" pageId="19" pageNumber="462" stateProvince="Jiangxi">Bijia Shan</location>
(笔架山),
<geoCoordinate id="5B9A500FFF98FFFAFECDD66DFE286764" box="[355,489,1865,1890]" direction="north" orientation="latitude" pageId="19" pageNumber="462" precision="15" value="26.5175">26°31'03&quot;N</geoCoordinate>
,
<geoCoordinate id="5B9A500FFF98FFFAFE5BD66DFD456767" box="[501,644,1865,1889]" direction="east" orientation="longitude" pageId="19" pageNumber="462" precision="15" value="114.18806">114°11'17&quot;E</geoCoordinate>
, mixed leaf litter, sifted,
<quantity id="F9569B2DFF98FFFAFC31D66DFC286767" box="[927,1001,1865,1890]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="19" pageNumber="462" unit="m" value="580.0">
<elevation id="B583D1FBFF98FFFAFC31D66DFC286767" box="[927,1001,1865,1890]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="19" pageNumber="462" unit="m" value="580.0">580 m</elevation>
</quantity>
,
<date id="4A101008FF98FFFAFC58D66DFBB96764" box="[1014,1144,1865,1890]" pageId="19" pageNumber="462" value="2014-07-24">
<collectingDate id="5A54E9E0FF98FFFAFC58D66DFBB96764" box="[1014,1144,1865,1890]" pageId="19" pageNumber="462" value="2014-07-24">24.vii.2014</collectingDate>
</date>
,
<collectorName id="935B531EFF98FFFAFB2AD66DFB036764" box="[1156,1218,1865,1890]" pageId="19" pageNumber="462">Chen</collectorName>
,
<collectorName id="935B531EFF98FFFAFB60D66EFB326767" box="[1230,1267,1866,1889]" pageId="19" pageNumber="462">Hu</collectorName>
,
<collectorName id="935B531EFF98FFFAFB51D66EFADF6767" box="[1279,1310,1866,1889]" pageId="19" pageNumber="462">Lv</collectorName>
&amp;
<collectorName id="935B531EFF98FFFAFAEDD66EFAA36767" box="[1347,1378,1866,1889]" pageId="19" pageNumber="462">Yu</collectorName>
leg.’ (
<collectionCode id="58BFAE0DFF98FFFAFF31D64AFF316780" box="[159,240,1902,1926]" pageId="19" pageNumber="462">SNUC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF98FFFDFF69D6B5FD1F6139" lastPageId="20" lastPageNumber="463" pageId="19" pageNumber="462" type="discussion">
<paragraph id="3E1136C8FF98FFFDFF69D6B5FD7860D3" blockId="19.[151,1437,1825,2034]" lastBlockId="20.[151,1437,151,357]" lastPageId="20" lastPageNumber="463" pageId="19" pageNumber="462">
<emphasis id="0CDAEADAFF98FFFAFF69D6B5FE8E67AC" bold="true" box="[199,335,1937,1962]" pageId="19" pageNumber="462">Comments.</emphasis>
The glabrous body (
<figureCitation id="A6952A4DFF98FFFAFD94D6B5FDBC67AF" box="[570,637,1937,1962]" captionStart="FIGURE 7" captionStartId="13.[151,250,1908,1930]" captionTargetBox="[162,1423,678,1868]" captionTargetId="figure@13.[151,1435,663,1886]" captionTargetPageId="13" captionText="FIGURE 7. Dorsal habitus of Batraxis tibialis. A. Population from Jiangxi. B. Population from Hainan. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256548/files/figure.png" pageId="19" pageNumber="462">Fig. 7</figureCitation>
A), obliquely narrowed abdomen at base, distinctly concaved protibiae (
<figureCitation id="A6952A4DFF98FFFAFF31D691FF3067C8" box="[159,241,1973,1998]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="19" pageNumber="462">Fig. 14</figureCitation>
F), presence of small ventral protuberance at the middle of profemora (
<figureCitation id="A6952A4DFF98FFFAFBA0D691FBA067C8" box="[1038,1121,1973,1998]" captionStart="FIGURE 14" captionStartId="20.[151,250,1889,1911]" captionTargetBox="[185,1407,412,1869]" captionTargetId="figure@20.[171,1420,403,1869]" captionTargetPageId="20" captionText="FIGURE 14. Legs of male Batraxis species. (A – F. Protibia. G. Mesotibia. H, I. Profemur. J – V. Protrochanter). A, K. B. gloriosa, population from Fujian. B, L. B. gloriosa, population from Zhejiang. C, H, P. B. gesar. D, T. B. ziyueae. E, U. B. bicolor, population from Yunnan. F, I, V. B. tibialis, population from Jiangxi. G, S. B. yifan. J. B. gloriosa, population from Hunan. M. B. gloriosa, population from Jiangxi. N. B. gloriosa, population from Guangxi. O. B. longicornis. Q. B. longithorax. R. B. nanlingensis. Scale bars: A ‒ I = 0.1 mm; J ‒ V = 0.05 mm." httpUri="https://zenodo.org/record/256555/files/figure.png" pageId="19" pageNumber="462">
Fig.
<date id="4A101008FF98FFFAFBEDD692FBA067C8" box="[1091,1121,1974,1998]" pageId="19" pageNumber="462" value="2014-01">14</date>
</figureCitation>
I), and the form of aedeagus (
<figureCitation id="A6952A4DFF98FFFAFF31D6FDFF3367F7" box="[159,242,2009,2034]" captionStart="FIGURE 13" captionStartId="19.[151,250,1495,1517]" captionTargetBox="[185,1406,200,1454]" captionTargetId="figure@19.[174,1410,193,1463]" captionTargetPageId="19" captionText="FIGURE 13. Aedeagi of Batraxis species. (A, B, D, E. in ventral view. C, F. in lateral view). A – C. B. bicolor, population from Yunnan. D – F. B. tibialis, population from Jiangxi. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/256554/files/figure.png" pageId="19" pageNumber="462">Figs 13</figureCitation>
D–F) of the population from
<collectingRegion id="FC6AF82AFF98FFFAFD90D6FDFD4E67F4" box="[574,655,2009,2034]" country="China" name="Jiangxi" pageId="19" pageNumber="462">Jiangxi</collectingRegion>
readily lead its identity to
<taxonomicName id="F9AE4D4BFF98FFFAFC6ED6FEFBEF67F7" box="[960,1070,2009,2033]" class="Insecta" family="Staphylinidae" genus="Batraxis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="462" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="0CDAEADAFF98FFFAFC6ED6FEFC1067F7" box="[960,977,2010,2033]" italics="true" pageId="19" pageNumber="462">B</emphasis>
.
<emphasis id="0CDAEADAFF98FFFAFC4ED6FDFBEF67F7" box="[992,1070,2009,2033]" italics="true" pageId="19" pageNumber="462">tibialis</emphasis>
</taxonomicName>
. The differences in the forms of the protibial cavity and aedeagal endophallus between the populations from
<collectingRegion id="FC6AF82AFF9FFFFDFB8CD1B3FBB260B6" box="[1058,1139,151,176]" country="China" name="Hainan" pageId="20" pageNumber="463">Hainan</collectingRegion>
(
<figureCitation id="A6952A4DFF9FFFFDFB26D1B3FB0E60B6" box="[1160,1231,151,176]" captionStart="FIGURE 7" captionStartId="13.[151,250,1908,1930]" captionTargetBox="[162,1423,678,1868]" captionTargetId="figure@13.[151,1435,663,1886]" captionTargetPageId="13" captionText="FIGURE 7. Dorsal habitus of Batraxis tibialis. A. Population from Jiangxi. B. Population from Hainan. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/256548/files/figure.png" pageId="20" pageNumber="463">Fig. 7</figureCitation>
B) and
<collectingRegion id="FC6AF82AFF9FFFFDFA82D1B3FABC60B6" box="[1324,1405,151,176]" country="China" name="Jiangxi" pageId="20" pageNumber="463">Jiangxi</collectingRegion>
is considered intraspecific variation of this species.
</paragraph>
<paragraph id="3E1136C8FF9FFFFDFF69D1C4FD1F6139" blockId="20.[151,1437,151,357]" pageId="20" pageNumber="463">
Measurements of the population from
<collectingRegion id="FC6AF82AFF9FFFFDFDD1D1FBFD1560FE" box="[639,724,223,248]" country="China" name="Jiangxi" pageId="20" pageNumber="463">Jiangxi</collectingRegion>
: BL
<quantity id="F9569B2DFF9FFFFDFCA3D1C4FCB460F1" box="[781,885,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="20" pageNumber="463" unit="mm" value="1.93">1.93 mm</quantity>
, HL
<quantity id="F9569B2DFF9FFFFDFC1FD1C4FBD960F1" box="[945,1048,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="20" pageNumber="463" unit="mm" value="0.42">0.42 mm</quantity>
, HW
<quantity id="F9569B2DFF9FFFFDFBF0D1C4FB0760F1" box="[1118,1222,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="20" pageNumber="463" unit="mm" value="0.43">0.43 mm</quantity>
, PL
<quantity id="F9569B2DFF9FFFFDFB53D1C4FAA460F1" box="[1277,1381,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="463" unit="mm" value="0.38">0.38 mm</quantity>
, PW
<quantity id="F9569B2DFF9FFFFDFF39D020FF3C611A" box="[151,253,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="20" pageNumber="463" unit="mm" value="0.42">0.42 mm</quantity>
, EL
<quantity id="F9569B2DFF9FFFFDFE9CD020FE59611A" box="[306,408,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="20" pageNumber="463" unit="mm" value="0.53">0.53 mm</quantity>
, EW
<quantity id="F9569B2DFF9FFFFDFE78D020FDFC611A" box="[470,573,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="20" pageNumber="463" unit="mm" value="0.72">0.72 mm</quantity>
, AL
<quantity id="F9569B2DFF9FFFFDFDDBD020FD1A611A" box="[629,731,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="463" unit="mm" value="0.6">0.60 mm</quantity>
, AW
<quantity id="F9569B2DFF9FFFFDFCB4D020FC40611A" box="[794,897,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="20" pageNumber="463" unit="mm" value="0.69">0.69 mm</quantity>
, mm, width between discal carinae of tergite IV
<quantity id="F9569B2DFF9FFFFDFF39D00CFF3A6139" box="[151,251,296,320]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="463" unit="mm" value="0.3">0.30 mm</quantity>
at apices, length of the aedeagus
<quantity id="F9569B2DFF9FFFFDFDDDD00CFD1B6139" box="[627,730,296,320]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="463" unit="mm" value="0.24">0.24 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="76B46543FF9FFFFDFF69D068FC296163" box="[199,1000,332,357]" pageId="20" pageNumber="463" type="distribution">
<paragraph id="3E1136C8FF9FFFFDFF69D068FC296163" blockId="20.[151,1437,151,357]" box="[199,1000,332,357]" pageId="20" pageNumber="463">
<emphasis id="0CDAEADAFF9FFFFDFF69D068FEA16163" bold="true" box="[199,352,332,357]" pageId="20" pageNumber="463">Distribution.</emphasis>
<collectingCountry id="46B97658FF9FFFFDFEC9D068FE6F6163" box="[359,430,332,357]" name="China" pageId="20" pageNumber="463">China</collectingCountry>
:
<collectingRegion id="FC6AF82AFF9FFFFDFE17D068FDCB6163" box="[441,522,332,357]" country="China" name="Jiangxi" pageId="20" pageNumber="463">Jiangxi</collectingRegion>
(new provincial record),
<collectingRegion id="FC6AF82AFF9FFFFDFC89D068FCB96163" box="[807,888,332,357]" country="China" name="Hainan" pageId="20" pageNumber="463">Hainan</collectingRegion>
(
<figureCitation id="A6952A4DFF9FFFFDFC29D068FC1A6163" box="[903,987,332,357]" captionStart="FIGURE 15" captionStartId="21.[151,250,1023,1045]" captionTargetBox="[151,1436,193,1001]" captionTargetId="figure@21.[151,1436,193,1002]" captionTargetPageId="21" captionText="FIGURE 15. Distribution of Batraxis species in China." httpUri="https://zenodo.org/record/256556/files/figure.png" pageId="20" pageNumber="463">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

107
data/B6/07/A7/B607A7B0D63AA99A84344E377E890DB3.xml Normal file
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<mods:title id="B4FFCEEC7E87E85F53342C5AEB60705D">Flora der Schweiz und angrenzender Gebiete. Band 2. Nymphaceae bis Primulaceae (2 nd edition) (p. 956): Papaveraceae</mods:title>
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<paragraph id="3866B1BA06FA2466B105E07BC51A86D4" pageId="null" pageNumber="116">
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Glaucium
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(
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) Rudolph
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<paragraph id="B9686457B698AD3AF2539CC64320C8A9" pageId="null" pageNumber="116">Roter Hornmohn</paragraph>
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</title>
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<paragraph id="4D139526617277B052EFA16D53192C34" pageId="null" pageNumber="116">
Meist 1
<normalizedToken id="7A0CAEF55EA3B01E665A108E7EEA78D7" originalValue="jährig" pageId="null" pageNumber="116">jaehrig</normalizedToken>
; 15-50 cm hoch;
<normalizedToken id="2DCD77E80BB54B0C589DC1A0BA13D87F" originalValue="blaugrün" pageId="null" pageNumber="116">blaugruen</normalizedToken>
bereift. Stengel aufrecht, verzweigt, zerstreut behaart.
<normalizedToken id="99CA9DF64F7B0B51EDBB9CCBD78EB5DE" originalValue="Blätter" pageId="null" pageNumber="116">Blaetter</normalizedToken>
dick, fiederteilig, mit gerundeten Buchten und eher schmalen,
<normalizedToken id="D7A4F1F083FA8DD7843C8EDC73BA6F96" originalValue="unregelmäßig" pageId="null" pageNumber="116">unregelmaessig</normalizedToken>
<normalizedToken id="64582B22C03347998B7923CE3C3513EA" originalValue="gezähnten" pageId="null" pageNumber="116">gezaehnten</normalizedToken>
Abschnitten, zerstreut behaart bis kahl, die untern gestielt, die obern mit schmalen, spitzen Zipfeln etwas umfassend.
<normalizedToken id="A896FCFB0087E29BA193EF4EFC34B52F" originalValue="Kelchblätter" pageId="null" pageNumber="116">Kelchblaetter</normalizedToken>
mit bis 4 mm langen Gliederhaaren.
<emphasis id="BF218E4461C1B7476EADEB05A6E93976" italics="true" pageId="null" pageNumber="116">
<normalizedToken id="1D6EF3DE67EE9BF9BA2A68AE72CE8843" originalValue="Kronblätter" pageId="null" pageNumber="116">Kronblaetter</normalizedToken>
1,5-3 cm lang, scharlachrot bis orangegelb
</emphasis>
, oft mit dunklem Fleck am Grunde. Frucht 10-20 cm lang,
<emphasis id="5DEBEB1D778C088C4600342CDA00C16D" italics="true" pageId="null" pageNumber="116">gerade oder nur wenig gebogen, mit nach vorn gerichteten Haaren besetzt.</emphasis>
Samen fast kugelig, 1,2-1,5 mm lang. -
<normalizedToken id="2E87B2213CBF983BB35C0833FCF4B86E" originalValue="Blüte" pageId="null" pageNumber="116">Bluete</normalizedToken>
: Sommer.
</paragraph>
<paragraph id="518086D4B98B482E13388639762347C0" pageId="null" pageNumber="116">
<emphasis id="B18A0775194E6D71CC9578B3268D300F" italics="true" pageId="null" pageNumber="116">Zytologische Angaben. 2n</emphasis>
=
<emphasis id="C5C469BD34F14147317344EAE4281F53" italics="true" pageId="null" pageNumber="116">12:</emphasis>
Ohne Herkunftsangabe des Materials (Sugiura 1940).
</paragraph>
<paragraph id="E524401AE88B0F0972CA17DD3C982C32" pageId="null" pageNumber="116">
<emphasis id="B4404E8CF8738301A31487F8AFC58A21" italics="true" pageId="null" pageNumber="116">Standort.</emphasis>
Kollin. Lockere,
<normalizedToken id="8BA5C31B48F255F61E251BA4DF2F7B38" originalValue="nährstoffreiche" pageId="null" pageNumber="116">naehrstoffreiche</normalizedToken>
, kalkhaltige
<normalizedToken id="901FFFEC9920955DFC6D62EC30168CA9" originalValue="Böden" pageId="null" pageNumber="116">Boeden</normalizedToken>
in warmen Lagen. Getreidefelder,
<normalizedToken id="399F077AC68D14552EE118C147D2FB48" originalValue="Schuttplätze" pageId="null" pageNumber="116">Schuttplaetze</normalizedToken>
.
</paragraph>
<paragraph id="30A78575DCCC25F680140E989AF4B5A1" pageId="null" pageNumber="116">
<emphasis id="BD6E613BD9059D91FE5C7B640109D17F" italics="true" pageId="null" pageNumber="116">Verbreitung. Mediterrane Pflanze:</emphasis>
<normalizedToken id="7F1AC10A171135D441EEA538E02072E7" originalValue="Nordwärts" pageId="null" pageNumber="116">Nordwaerts</normalizedToken>
bis
<normalizedToken id="C4BF674DD2D4EC189960C3010C5E12E7" originalValue="Südfrankreich" pageId="null" pageNumber="116">Suedfrankreich</normalizedToken>
, Mittelitalien,
<normalizedToken id="AD145D9D05E0913165F46231E949CCBC" originalValue="Südbalkan" pageId="null" pageNumber="116">Suedbalkan</normalizedToken>
; Kleinasien, Nordwestafrika. - Im Gebiet gelegentlich eingeschleppt,
<normalizedToken id="24FEF3672E7DCD87D89B9245B3A3D833" originalValue="früher" pageId="null" pageNumber="116">frueher</normalizedToken>
z.B. im Wallis.
</paragraph>
</subSubSection>
</treatment>
</document>

193
data/B6/07/AF/B607AF452F29579F8EA8EE9CCF53646E.xml Normal file
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@ -0,0 +1,193 @@
<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.6.87158" ID-Pensoft-Pub="2535-0730-2-143" ID-Pensoft-UUID="DB80A4F877A753509B09A89648156DB4" ID-ZooBank="F5A5F50EC380414F838DA60AD251B2C6" ModsDocID="2535-0730-6-2-143" checkinTime="1658525555661" checkinUser="pensoft" docAuthor="Zamani, Alireza, Al-Yacoub, Ghassan A. Ali &amp; Abdullah Najim, Shurooq" docDate="2022" docId="B607AF452F29579F8EA8EE9CCF53646E" docLanguage="en" docName="EvolutSyst 6(2): 143-150" docOrigin="Evolutionary Systematics 6 (2)" docPubDate="2022-07-22" docSource="http://dx.doi.org/10.3897/evolsyst.6.87158" docTitle="Zelotes fagei Denis 1955" docType="treatment" docVersion="1" id="DB80A4F877A753509B09A89648156DB4" lastPageNumber="143" masterDocId="DB80A4F877A753509B09A89648156DB4" masterDocTitle="New data on Gnaphosidae (Arachnida, Araneae) of Iraq" masterLastPageNumber="150" masterPageNumber="143" pageNumber="143" updateTime="1658525555661" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>New data on Gnaphosidae (Arachnida, Araneae) of Iraq</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamani, Alireza</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, Turku FI- 20014, Finland</mods:affiliation>
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</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Al-Yacoub, Ghassan A. Ali</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4931-855X</mods:nameIdentifier>
<mods:affiliation>Department of Biology, College of Education for Pure Sciences, University of Thi-Qar, Thi-Qar, Iraq</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Abdullah Najim, Shurooq</mods:namePart>
<mods:affiliation>Department of Ecology, College of Science, University of Basrah, Basrah, Iraq</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Evolutionary Systematics</mods:title>
</mods:titleInfo>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-07-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>6</mods:number>
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<mods:number>2</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.6.87158</mods:identifier>
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<subSection pageId="0" pageNumber="143" type="results">
<treatment LSID="urn:lsid:plazi:treatment:B607AF452F29579F8EA8EE9CCF53646E" httpUri="http://treatment.plazi.org/id/B607AF452F29579F8EA8EE9CCF53646E" lastPageNumber="143" pageId="0" pageNumber="143">
<subSubSection pageId="0" pageNumber="143" type="nomenclature">
<paragraph pageId="0" pageNumber="143">
<taxonomicName LSID="B607AF45-2F29-579F-8EA8-EE9CCF53646E" authority="Denis, 1955" authorityName="Denis" authorityYear="1955" class="Arachnida" family="Gnaphosidae" genus="Zelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelotes fagei" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="fagei">Zelotes fagei Denis, 1955</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="description">
<paragraph pageId="0" pageNumber="143">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Ventral view of epigynes (B, C, D) and dorsal view of vulvae (A, E) of gnaphosids. A. Haplodrassus dalmatensis (L. Koch, 1866); B. Minosiella intermedia Denis, 1958; C. Marinarozelotes jaxartensis (Kroneberg, 1875); D, E. Zelotes fagei Denis, 1955." httpUri="https://binary.pensoft.net/fig/718432" pageId="0" pageNumber="143">Fig. 4D, E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="identification">
<paragraph pageId="0" pageNumber="143">Identification.</paragraph>
<paragraph pageId="0" pageNumber="143">
<bibRefCitation author="Levy, G" journalOrPublisher="Israel Journal of Zoology" pageId="0" pageNumber="143" pagination="93 - 158" refId="B19" refString="Levy, G, 1998. The ground-spider genera Setaphis, Trachyzelotes, Zelotes, and Drassyllus (Araneae: Gnaphosidae) in Israel. Israel Journal of Zoology 44: 93 - 158" title="The ground-spider genera Setaphis, Trachyzelotes, Zelotes, and Drassyllus (Araneae: Gnaphosidae) in Israel." volume="44" year="1998">Levy (1998)</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.13156/arac.2011.14.3.97" author="FitzPatrick, MJ" journalOrPublisher="Bulletin of the British Arachnological Society" pageId="0" pageNumber="143" pagination="97 - 172" refId="B15" refString="FitzPatrick, MJ, 2007. A taxonomic revision of the Afrotropical species of Zelotes (Arachnida: Araneae: Gnaphosidae). Bulletin of the British Arachnological Society 14 (3): 97 - 172, DOI: https://doi.org/10.13156/arac.2011.14.3.97" title="A taxonomic revision of the Afrotropical species of Zelotes (Arachnida: Araneae: Gnaphosidae)." url="https://doi.org/10.13156/arac.2011.14.3.97" volume="14" year="2007">FitzPatrick (2007)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="materials_examined">
<paragraph pageId="0" pageNumber="143">Material.</paragraph>
<paragraph pageId="0" pageNumber="143">
<materialsCitation collectingDate="1972-12-20" collectorName="O. Jakes" country="IRAQ" elevation="260" latitude="30.8" location="Ash Shabakah" longLatPrecision="1220" longitude="43.65" specimenCount="1" specimenCount-female="1" stateProvince="Najaf Province">
<collectingCountry name="Iraq">IRAQ</collectingCountry>
:
<emphasis bold="true" italics="true" pageId="0" pageNumber="143">
<collectingRegion country="Iraq" name="An Najaf">Najaf Province</collectingRegion>
</emphasis>
:
<specimenCount type="female">1♀</specimenCount>
(MMBC),
<location LSID="urn:lsid:plazi:treatment:B607AF452F29579F8EA8EE9CCF53646E:AFFE6F4593402948C11DB98F5C6AA466" country="IRAQ" latitude="30.8" longLatPrecision="1220" longitude="43.65" name="Ash Shabakah" stateProvince="Najaf Province">Ash Shabakah</location>
camp,
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.5" unit="km" value="150.0">150 km</quantity>
SW of
<collectingRegion country="Iraq" name="An Najaf">Najaf</collectingRegion>
, [
<geoCoordinate degrees="30" direction="north" minutes="48" orientation="latitude" precision="925" value="30.8">30°48'N</geoCoordinate>
,
<geoCoordinate degrees="43" direction="east" minutes="39" orientation="longitude" precision="925" value="43.65">43°39'E</geoCoordinate>
],
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.6" unit="m" value="260.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.6" unit="m" value="260.0">260 m</elevation>
</quantity>
, stone desert,
<collectingDate value="1972-12-20">20.12.1972</collectingDate>
(
<collectorName>
O.
<normalizedToken originalValue="Jakeš">Jakes</normalizedToken>
</collectorName>
)
</materialsCitation>
.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/718432" pageId="0" pageNumber="143" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="143">
<emphasis bold="true" pageId="0" pageNumber="143">Figure 4.</emphasis>
Ventral view of epigynes (
<emphasis bold="true" pageId="0" pageNumber="143">B, C, D</emphasis>
) and dorsal view of vulvae (
<emphasis bold="true" pageId="0" pageNumber="143">A, E</emphasis>
) of gnaphosids.
<emphasis bold="true" pageId="0" pageNumber="143">A.</emphasis>
<taxonomicName baseAuthorityName="L. Koch" baseAuthorityYear="1866" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus dalmatensis" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="dalmatensis">
<emphasis italics="true" pageId="0" pageNumber="143">Haplodrassus dalmatensis</emphasis>
</taxonomicName>
(L. Koch, 1866);
<emphasis bold="true" pageId="0" pageNumber="143">B.</emphasis>
<taxonomicName authorityName="Denis" authorityYear="1958" class="Arachnida" family="Gnaphosidae" genus="Minosiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Minosiella intermedia" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis italics="true" pageId="0" pageNumber="143">Minosiella intermedia</emphasis>
</taxonomicName>
Denis, 1958;
<emphasis bold="true" pageId="0" pageNumber="143">C.</emphasis>
<taxonomicName baseAuthorityName="Kroneberg" baseAuthorityYear="1875" family="Gnaphosidae" genus="Marinarozelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Marinarozelotes jaxartensis" order="Aranei" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="jaxartensis">
<emphasis italics="true" pageId="0" pageNumber="143">Marinarozelotes jaxartensis</emphasis>
</taxonomicName>
(Kroneberg, 1875);
<emphasis bold="true" pageId="0" pageNumber="143">D, E.</emphasis>
<taxonomicName authorityName="Denis" authorityYear="1955" class="Arachnida" family="Gnaphosidae" genus="Zelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelotes fagei" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="fagei">
<emphasis italics="true" pageId="0" pageNumber="143">Zelotes fagei</emphasis>
</taxonomicName>
Denis, 1955.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="distribution">
<paragraph pageId="0" pageNumber="143">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="143">Previously known from Niger, Egypt and Israel.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="comments">
<paragraph pageId="0" pageNumber="143">Comment.</paragraph>
<paragraph pageId="0" pageNumber="143">New record for Iraq, with the current material representing the easternmost record of the species across its known range.</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/718433" pageId="0" pageNumber="143" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="143">
<emphasis bold="true" pageId="0" pageNumber="143">Figure 5.</emphasis>
Epigynes of
<taxonomicName authorityName="Fage" authorityYear="1929" class="Arachnida" family="Gnaphosidae" genus="Pterotricha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterotricha dalmasi" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="dalmasi">
<emphasis italics="true" pageId="0" pageNumber="143">Pterotricha dalmasi</emphasis>
</taxonomicName>
Fage, 1929 (
<emphasis bold="true" pageId="0" pageNumber="143">A-C</emphasis>
) and
<taxonomicName authorityName="Zamani &amp; Marusik" authorityYear="2018" class="Arachnida" family="Gnaphosidae" genus="Pterotricha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterotricha kovblyuki" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="kovblyuki">
<emphasis italics="true" pageId="0" pageNumber="143">Pterotricha kovblyuki</emphasis>
</taxonomicName>
Zamani &amp; Marusik, 2018 (
<emphasis bold="true" pageId="0" pageNumber="143">D-F</emphasis>
).
<emphasis bold="true" pageId="0" pageNumber="143">A, D.</emphasis>
intact epigyne, ventral view;
<emphasis bold="true" pageId="0" pageNumber="143">B, E.</emphasis>
macerated epigyne, ventral view;
<emphasis bold="true" pageId="0" pageNumber="143">C, F.</emphasis>
vulva, dorsal view. Abbreviations:
<emphasis italics="true" pageId="0" pageNumber="143">Ag</emphasis>
- accessorial gland;
<emphasis italics="true" pageId="0" pageNumber="143">Ah</emphasis>
- anterior hood;
<emphasis italics="true" pageId="0" pageNumber="143">Ld</emphasis>
- lateral duct;
<emphasis italics="true" pageId="0" pageNumber="143">Lm</emphasis>
- lateral margin;
<emphasis italics="true" pageId="0" pageNumber="143">Re</emphasis>
- receptacle.
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>

89
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</mods:name>
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<mods:role>
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</mods:role>
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<taxonomicNameLabel pageId="15" pageNumber="88">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="88" type="reference_group">
<paragraph pageId="15" pageNumber="88">
≡
<taxonomicName authorityName="Jahn &amp; Kusber &amp; Cocquyt" authorityYear="2017" baseAuthorityName="Jahn &amp; Kusber &amp; Cocquyt" baseAuthorityYear="2017" class="Bacillariophyceae" family="Surirellaceae" genus="Surirella" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Surirella spiraloides" order="Surirellales" pageId="15" pageNumber="88" phylum="Bacillariophyta" rank="species" species="spiraloides" status="comb. nov.">Surirella spiraloides</taxonomicName>
Hust. in Huber-Pestalozzi, Phytoplankt.
<normalizedToken originalValue="Süsswass">Suesswass</normalizedToken>
. vol. 2 (2), 507, fig. 617. 1942.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="88" type="lectotype">
<paragraph pageId="15" pageNumber="88">Lectotype</paragraph>
<paragraph pageId="15" pageNumber="88">
(designated by
<bibRefCitation author="Simonsen, R" journalOrPublisher="Atlas and catalogue of the diatom types of Friedrich Hustedt" pageId="28" pageNumber="101" refId="B90" refString="Simonsen, R, 1987. . Atlas and catalogue of the diatom types of Friedrich Hustedt 1" volume="1" year="1987">Simonsen 1987</bibRefCitation>
). BRM X4/45 Lake Tanganyika &quot;Tanganika See. 6&quot;.
</paragraph>
<paragraph pageId="15" pageNumber="88">http://phycobank.org/100083</paragraph>
<paragraph pageId="15" pageNumber="88">
-
<taxonomicName authorityName="Jahn &amp; Kusber &amp; Cocquyt" authorityYear="2017" baseAuthorityName="Jahn &amp; Kusber &amp; Cocquyt" baseAuthorityYear="2017" class="Bacillariophyceae" family="Surirellaceae" genus="Surirella" kingdom="Plantae" lsidName="Surirella spiraloides" order="Surirellales" pageId="15" pageNumber="88" phylum="Bacillariophyta" rank="species" species="spiraloides">
<emphasis italics="true" pageId="15" pageNumber="88">Surirella spiraloides</emphasis>
</taxonomicName>
Hust. in A.W.F. Schmidt, Atlas Diatom.-Kunde, pl. 353: fig. 2, 3. 1922, nom. inval.
</paragraph>
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<taxonomicName id="5D81BBE99248E0F446F9B23F6B6E920C" ID-CoL="79NWK" authority="Pfeiffer, 1856" authorityName="Pfeiffer" authorityYear="1856" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika hainesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">Sarika hainesi (Pfeiffer, 1856)</taxonomicName>
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<taxonomicName id="91A578F4587305D8C6377E6A06D2EC1A" class="Gastropoda" family="Ariophantidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix hainesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">Helix hainesi</taxonomicName>
Pfeiffer, 1856a: 32. Type locality: Siam [Thailand].
<bibRefCitation id="98856DA93CAB8C14C5056B0B2CE74483" pageId="0" pageNumber="1">Pfeiffer 1856b</bibRefCitation>
: 75, 76, pl. 21, figs 1-3.
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<paragraph id="C3699181856C4D2589A72CA8EBB47E69" pageId="0" pageNumber="1">
<taxonomicName id="997EBB571BFFE44FDDEF36ED8870EC80" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrochlamys hainesi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesi">Macrochlamys hainesi</taxonomicName>
:
<bibRefCitation id="D0B848D2295CF57B5B402F7937F17506" author="Saurin, E" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="113 - 120" title="Coquilles nouvelles de ľIndochine." volume="93" year="1953">Saurin 1953</bibRefCitation>
: 113.
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<taxonomicName id="2352E324A8F8B360F6882E8E17CAB18E" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika hainesii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="hainesii">Sarika aff. hainesii</taxonomicName>
[sic]:
<bibRefCitation id="38733C2BE788673B7ACED0F2BB1A51DF" author="Solem, A" journalOrPublisher="Spolia Zoologia Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" title="Some non-marine mollusks from Thailand." volume="24" year="1966">Solem 1966</bibRefCitation>
: 38, 39, fig. 5a.
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<paragraph id="7147C49BC438800CC7206481759CCEB3" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="ABDAA3AC35998D3A96447DCBB21E6E4F" pageId="0" pageNumber="1">
Syntypes NMHUK ex. Cuming collection from
<normalizedToken id="703250536C53CB7EA14BDC8063D9D668" originalValue="“Siam”">&quot;Siam&quot;</normalizedToken>
(3 shells; Fig. 38F).
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<paragraph id="87B5C3CB829FA968808DEC736A60ED29" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="444F3FF2B3E05BBA5C568CD5AD3C97AC" pageId="0" pageNumber="1">
Laos and Thailand (
<bibRefCitation id="0DD55083EF150D0F303BE74A60B1A16E" author="Saurin, E" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="113 - 120" title="Coquilles nouvelles de ľIndochine." volume="93" year="1953">Saurin 1953</bibRefCitation>
,
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).
</paragraph>
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<paragraph id="0EF6F850F07143D7A213B676CD18D942" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="C94087EC237EE8E0E917F2BBE87C103B" pageId="0" pageNumber="1">No material of this species was found, and only the type specimens were examined.</paragraph>
<caption id="F63298477DAE0E5A005A76AC68651B24" pageId="0" pageNumber="1">
<paragraph id="4D7670B507A40A7E04D6BF339EB307FC" pageId="0" pageNumber="1">
Figure 38. A, B
<taxonomicName id="4763C4B66A17B249B5C71ADD9D0B5FAC" class="Gastropoda" family="Ariophantidae" genus="Microcystina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcystina messageri" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="messageri">Microcystina messageri</taxonomicName>
A syntype MNHN-IM-2000-9658 and B CUMZ collection C
<taxonomicName id="EB90F936482AA9C58AE5EA2E6FEA714D" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sarika benoiti" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="benoiti">Sarika benoiti</taxonomicName>
, CUMZ collection D, E
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D syntype MNHN-IM-2000-27882 and E CUMZ collection F
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, syntype NMHUK ex. Cuming collection.
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Anthicus melancholicus
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, 1848**
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Map 21
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New Brunswick, Sunbury Co. 9.5 km NE jct. 101 &amp; 645,
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,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-66.6755">66.6755°W</geoCoordinate>
, 30.VIII.2008, R. P. Webster, old field with open sandy areas, sweeping foliage (1, RWC).
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<paragraph pageId="15" pageNumber="294">Collection and habitat data.</paragraph>
<paragraph pageId="15" pageNumber="294">This species was swept from foliage in an old field with open sandy areas. The adult was captured during late August.</paragraph>
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<paragraph pageId="15" pageNumber="294">Distribution in Canada and Alaska.</paragraph>
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ON, QC, NB (
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).
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<paragraph pageId="15" pageNumber="294">
Map 21. Collection localities in New Brunswick, Canada of
<taxonomicName class="Insecta" family="Anthicidae" genus="Anthicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthicus melancholicus" order="Coleoptera" pageId="15" pageNumber="294" phylum="Arthropoda" rank="species" species="melancholicus">Anthicus melancholicus</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1095.77996" ID-GBIF-Dataset="591e49b3-50ad-4f6c-8718-865ef7394b39" ID-PMC="PMC9021157" ID-Pensoft-Pub="1313-2970-1095-13" ID-Pensoft-UUID="EAEF3E50B7C054788C2706EB70256E09" ID-PubMed="35836690" ID-ZooBank="94B1536FF07C4595BE95A795A861D9E9" ModsDocID="1313-2970-1095-13" checkinTime="1649881565219" checkinUser="pensoft" docAuthor="Pham, Huyen T. M. &amp; Karanovic, Ivana" docDate="2022" docId="B608A8581A0553D3BC12BFA8435D5C36" docLanguage="en" docName="ZooKeys 1095: 13-42" docOrigin="ZooKeys 1095" docPubDate="2022-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1095.77996" docTitle="Parasterope singula Pham &amp; Karanovic 2022, sp. nov." docType="treatment" docUuid="4C34D735-E105-40E7-83A4-67BA3BD2C9EF" docUuidSource="ZooBank" docVersion="4" id="EAEF3E50B7C054788C2706EB70256E09" lastPageNumber="13" masterDocId="EAEF3E50B7C054788C2706EB70256E09" masterDocTitle="Four new Parasterope (Ostracoda, Myodocopina) from the Northwest Pacific and their phylogeny based on 16 S rRNA" masterLastPageNumber="42" masterPageNumber="13" pageNumber="13" updateTime="1668151662464" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Four new Parasterope (Ostracoda, Myodocopina) from the Northwest Pacific and their phylogeny based on 16 S rRNA</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pham, Huyen T. M.</mods:namePart>
<mods:affiliation>Department of Life Science, Research Institute for Convergence of Basic Science, College of Natural Sciences, Hanyang University, Seoul, 04763, Republic of Korea</mods:affiliation>
<mods:nameIdentifier type="email">minhhuyen.ks.nb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Karanovic, Ivana</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9002-9952</mods:nameIdentifier>
<mods:affiliation>Department of Life Science, Research Institute for Convergence of Basic Science, College of Natural Sciences, Hanyang University, Seoul, 04763, Republic of Korea &amp; Institute for Marine and Antarctic Studies, University of Tasmania, Private Bag 49, 7001, Hobart, Tasmania, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-04-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1095</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>13</mods:start>
<mods:end>42</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1095.77996</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1095.77996</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1095-13</mods:identifier>
<mods:identifier type="ZooBank">94B1536FF07C4595BE95A795A861D9E9</mods:identifier>
<mods:identifier type="Pensoft-UUID">EAEF3E50B7C054788C2706EB70256E09</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="194594626" LSID="urn:lsid:zoobank.org:act:4C34D735-E105-40E7-83A4-67BA3BD2C9EF" httpUri="http://treatment.plazi.org/id/B608A8581A0553D3BC12BFA8435D5C36" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection pageId="0" pageNumber="13" type="nomenclature">
<paragraph pageId="0" pageNumber="13">
<taxonomicName LSID="http://zoobank.org/4C34D735-E105-40E7-83A4-67BA3BD2C9EF" authority="Pham &amp; Karanovic, 2022" authorityName="Pham &amp; Karanovic" authorityYear="2022" class="Ostracoda" family="Cylindroleberididae" genus="Parasterope" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Parasterope singula" order="Myodocopida" pageId="0" pageNumber="13" phylum="Cnidaria" rank="species" species="singula" status="sp. nov.">Parasterope singula</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="description">
<paragraph pageId="0" pageNumber="13">
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Light microscopy and scanning electron microscopy images of Parasterope singula sp. nov. holotype female adult: a Lateral view from left valve b Internal view from left valve c External view." figureDoi="10.3897/zookeys.1095.77996.figure8" httpUri="https://binary.pensoft.net/fig/672096" pageId="0" pageNumber="13">Figs 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">, 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="materials_examined">
<paragraph pageId="0" pageNumber="13">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" italics="true" pageId="0" pageNumber="13">
<typeStatus>Holotype</typeStatus>
</emphasis>
female dissected on one slide, shells on SEM stub (NIBR IV 0000887835).
<materialsCitation collectingDate="2012-11-29" country="South Korea" county="Ho Young Soh" location="The" municipality="Chuja Island" specimenCount="two" specimenCount-female="1" typeStatus="Paratypes">
<emphasis bold="true" italics="true" pageId="0" pageNumber="13">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
<specimenCount type="female">two females</specimenCount>
dissected on one slide each, shells on SEM stub (NIBR IV 0000879835_2&amp;3).
<location LSID="urn:lsid:plazi:treatment:B608A8581A0553D3BC12BFA8435D5C36:FD50CB0DEBE4FDDE967C7FB7BFE93AB8" country="South Korea" county="Ho Young Soh" municipality="Chuja Island" name="The">The</location>
sample was collected from the type locality:
<collectingCountry name="South Korea">South Korea</collectingCountry>
,
<collectingMunicipality>Chuja Island</collectingMunicipality>
by
<collectingCounty>Ho Young Soh</collectingCounty>
on
<collectingDate value="2012-11-29">29 November 2012</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="etymology">
<paragraph pageId="0" pageNumber="13">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="13">
The name is a Latin noun,
<taxonomicName authorityName="Pham &amp; Karanovic" authorityYear="2022" class="Ostracoda" family="Cylindroleberididae" genus="Parasterope" kingdom="Animalia" lsidName="Parasterope singula" order="Myodocopida" pageId="0" pageNumber="13" phylum="Cnidaria" rank="species" species="singula">
<emphasis italics="true" pageId="0" pageNumber="13">Parasterope singula</emphasis>
</taxonomicName>
, because only a female has been collected. The name is in nominative, feminine singular, agreeing in gender with the genus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="diagnosis">
<paragraph pageId="0" pageNumber="13">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="13">Surface of the shell completely smooth. Anterior end with a deep incisure and posterior infold of the carapace with a broad shelf. Posterior end wider than anterior. Dorsal margin rounded. Lateral eye well developed with black pigmented ommatidia. Uropodal lamellae with nine claws.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="description">
<paragraph pageId="0" pageNumber="13">Description.</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Female.</emphasis>
Shell (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Light microscopy and scanning electron microscopy images of Parasterope singula sp. nov. holotype female adult: a Lateral view from left valve b Internal view from left valve c External view." figureDoi="10.3897/zookeys.1095.77996.figure8" httpUri="https://binary.pensoft.net/fig/672096" pageId="0" pageNumber="13">8</figureCitation>
). Carapace oval, broadening at posterior, greatest height near middle, carapace length 1.25 mm, height 0.9 mm. Carapace smooth and without ornamentation.
</paragraph>
<caption doi="10.3897/zookeys.1095.77996.figure8" httpUri="https://binary.pensoft.net/fig/672096" pageId="0" pageNumber="13" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 8.</emphasis>
Light microscopy and scanning electron microscopy images of
<taxonomicName authorityName="Pham &amp; Karanovic" authorityYear="2022" class="Ostracoda" family="Cylindroleberididae" genus="Parasterope" kingdom="Animalia" lsidName="Parasterope singula" order="Myodocopida" pageId="0" pageNumber="13" phylum="Cnidaria" rank="species" species="singula">
<emphasis italics="true" pageId="0" pageNumber="13">Parasterope singula</emphasis>
</taxonomicName>
sp. nov. holotype female adult:
<emphasis bold="true" pageId="0" pageNumber="13">a</emphasis>
Lateral view from left valve
<emphasis bold="true" pageId="0" pageNumber="13">b</emphasis>
Internal view from left valve
<emphasis bold="true" pageId="0" pageNumber="13">c</emphasis>
External view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="13">
A1 (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Light microscopy and scanning electron microscopy images of Parasterope singula sp. nov. holotype female adult: a Lateral view from left valve b Internal view from left valve c External view." figureDoi="10.3897/zookeys.1095.77996.figure8" httpUri="https://binary.pensoft.net/fig/672096" pageId="0" pageNumber="13">8a</figureCitation>
). 1st joint: with no hair on broad surface. 2nd joint: one spinous dorsal seta, one short lateral bristle, and small proximal dorsal spines. 3rd joint: seven setae (one short ventral and six dorsal). 4th joint: three bristles (one long dorsal bristle and two short ventral bristles). 5th joint: sensory bristle with six filaments without short proximal terminal. 6th joint: one medial seta, attached to bottom of 7th joint boundary. 7th joint: a-bristle claw-like, bare; b-seta with marginal filaments; c-seta with marginal filaments. 8th joint: d-bristle absent; e-bristle bare with blunt tip; f- and g-bristles with marginal filaments.
</paragraph>
<paragraph pageId="0" pageNumber="13">
Bellonci orange (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9b</figureCitation>
). Elongate with rounded tip with unclear constriction.
</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">
<emphasis italics="true" pageId="0" pageNumber="13">Eyes</emphasis>
.
</emphasis>
Lateral eye with 18 ommatidia.
</paragraph>
<paragraph pageId="0" pageNumber="13">
A2 (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9c, d</figureCitation>
). Protopodite: rounded without medial bristle. Endopodite (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9c</figureCitation>
): not strongly jointed, with long terminal bristle. Exopodite (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9d</figureCitation>
): 9-jointed; bristle of 2nd segment along ventral margin and spines along dorsal margin reaching 8th joint; 3rd-8th joints with natatory hairs and spines along proximal part of ventral margin. 9th article: four bristles (two long natatory and two short bristles). 4th-8th joints with minute spines at inner terminal corner.
</paragraph>
<caption doi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 9.</emphasis>
<taxonomicName authorityName="Pham &amp; Karanovic" authorityYear="2022" class="Ostracoda" family="Cylindroleberididae" genus="Parasterope" kingdom="Animalia" lsidName="Parasterope singula" order="Myodocopida" pageId="0" pageNumber="13" phylum="Cnidaria" rank="species" species="singula">
<emphasis italics="true" pageId="0" pageNumber="13">Parasterope singula</emphasis>
</taxonomicName>
sp. nov. holotype female adult:
<emphasis bold="true" pageId="0" pageNumber="13">a</emphasis>
A1
<emphasis bold="true" pageId="0" pageNumber="13">b</emphasis>
bellonci organ
<emphasis bold="true" pageId="0" pageNumber="13">c</emphasis>
A2 endopodite
<emphasis bold="true" pageId="0" pageNumber="13">d</emphasis>
A2 exopodite
<emphasis bold="true" pageId="0" pageNumber="13">e</emphasis>
Md basale and coxale endite
<emphasis bold="true" pageId="0" pageNumber="13">f</emphasis>
Md basale endite
<emphasis bold="true" pageId="0" pageNumber="13">g</emphasis>
Md endopodite
<emphasis bold="true" pageId="0" pageNumber="13">h</emphasis>
L5
<emphasis bold="true" pageId="0" pageNumber="13">i</emphasis>
L6
<emphasis bold="true" pageId="0" pageNumber="13">k</emphasis>
L7
<emphasis bold="true" pageId="0" pageNumber="13">l</emphasis>
F. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="13">
Md (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9e-g</figureCitation>
). Coxale endite (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9e</figureCitation>
) same as that on
<taxonomicName lsidName="P. busanensis" pageId="0" pageNumber="13" rank="species" species="busanensis">
<emphasis italics="true" pageId="0" pageNumber="13">P. busanensis</emphasis>
</taxonomicName>
. Basale (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9f</figureCitation>
): Basale endite with three spinous end bristles, four triaenid bristles with three spines excluding terminal pair, and two equal-length bare bristles. Basale dorsal margin with two long terminal setae, without mid-bristle; five rows of small spines on the broad surface near dorsal margin. Exopodite with hirsute tip and two small subterminal setae, exopodite almost same length as dorsal margin of first endopodite article. Endopodite (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9g</figureCitation>
): 1st joint: ventral margin with three bristles (two long with long marginal spines, and one shorter without spine or hair); 2nd joint: ventral margin: with three terminal bristles; dorsal margin: with stout a-, b-, c-, and d-bristles and without bristle proximal to a-bristle; one long e-bristle between b- and c-bristles; four medial bristles forming an oblique row adjacent to c-bristle; f-bristle between c- and d-bristles, long, bare; g-bristle longer than f-bristle. 3rd segment with a strong dorsal claw-like bristle without marginal spines, three juxtaposed stout bristles of equal length, and two thin bristles.
</paragraph>
<paragraph pageId="0" pageNumber="13">
Mxl. Endite I with three bristles. Endite II with three bristles. Basale: one dorsal medial distal bristle, two ventral medial proximal bristles. Endopodite: 1st with one short
<normalizedToken originalValue="α-seta">α-seta</normalizedToken>
and one hairy
<normalizedToken originalValue="β-seta">β-seta</normalizedToken>
; 2nd article with two terminal bristles.
</paragraph>
<paragraph pageId="0" pageNumber="13">
L5 (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9h</figureCitation>
). Ventral section with fan of long setae. Comb with spinous exopodal bristle just reaching past distal end of comb, one short slender bristle just ventral to base of stout bristle, one bristle near exopodal bristle stem, and four lateral bristles set back from edge at comb mid length.
</paragraph>
<paragraph pageId="0" pageNumber="13">
L6 (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9i</figureCitation>
). Unidentified number of endite bristle on the anterior margin. Lateral flap with hairs but no bristles. Anterior tip of skirt with two small bristles. Ventral and postero-ventral margin with 17 plumose bristles (one missing).
</paragraph>
<paragraph pageId="0" pageNumber="13">
L7 (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9k</figureCitation>
). Each limb with ten bristles. Four proximal and six distal bristles.
</paragraph>
<paragraph pageId="0" pageNumber="13">
F (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Parasterope singula sp. nov. holotype female adult: a A 1 b bellonci organ c A 2 endopodite d A 2 exopodite e Md basale and coxale endite f Md basale endite g Md endopodite h L 5 i L 6 k L 7 l F. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1095.77996.figure9" httpUri="https://binary.pensoft.net/fig/672097" pageId="0" pageNumber="13">9l</figureCitation>
). Each lamella with six claws plus three posterior claws bristle-like. A total of nine claws and bristles.
</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Male.</emphasis>
Not collected.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="remarks">
<paragraph pageId="0" pageNumber="13">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="13">
The presence of rows of small spines on the broad surface near the dorsal margin of the Md basale is a unique feature of
<taxonomicName authorityName="Pham &amp; Karanovic" authorityYear="2022" class="Ostracoda" family="Cylindroleberididae" genus="Parasterope" kingdom="Animalia" lsidName="Parasterope singula" order="Myodocopida" pageId="0" pageNumber="13" phylum="Cnidaria" rank="species" species="singula">
<emphasis italics="true" pageId="0" pageNumber="13">Parasterope singula</emphasis>
</taxonomicName>
. Additionally, the number of bristles on the L7 (ten bristles: four on the terminal segment (two on each side), six on the proximal segments (three on each side) is less than in all other
<taxonomicName class="Ostracoda" family="Cylindroleberididae" genus="Parasterope" kingdom="Animalia" lsidName="Parasterope" order="Myodocopida" pageId="0" pageNumber="13" phylum="Cnidaria" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Parasterope</emphasis>
</taxonomicName>
representatives. The new species is similar to
<taxonomicName lsidName="P. mckenziei" pageId="0" pageNumber="13" rank="species" species="mckenziei">
<emphasis italics="true" pageId="0" pageNumber="13">P. mckenziei</emphasis>
</taxonomicName>
, described from Samar Province, Philippines (McKenzie, 1970) but differs in the following characters: the Md endopodite is without a proximal bristle next to the a-bristle, a character not known in any other
<taxonomicName class="Ostracoda" family="Cylindroleberididae" genus="Parasterope" kingdom="Animalia" lsidName="Parasterope" order="Myodocopida" pageId="0" pageNumber="13" phylum="Cnidaria" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="13">Parasterope</emphasis>
</taxonomicName>
, and the number of claws on UL (nine vs. seven).
</paragraph>
<paragraph pageId="0" pageNumber="13">
The 2nd joint of the Md endopodite in
<taxonomicName lsidName="P. singula" pageId="0" pageNumber="13" rank="species" species="singula">
<emphasis italics="true" pageId="0" pageNumber="13">P. singula</emphasis>
</taxonomicName>
carries one oblique row of cleaning setae, also recorded in
<taxonomicName lsidName="P. antyx" pageId="0" pageNumber="13" rank="species" species="antyx">
<emphasis italics="true" pageId="0" pageNumber="13">P. antyx</emphasis>
</taxonomicName>
Kornicker, 1989 from Bay of Biscay (Kornicker, 1989),
<taxonomicName lsidName="P. gamma" pageId="0" pageNumber="13" rank="species" species="gamma">
<emphasis italics="true" pageId="0" pageNumber="13">P. gamma</emphasis>
</taxonomicName>
Kornicker, Harrison-Nelson &amp; Coles, 2007 from Hawaiian Islands (
<bibRefCitation author="Kornicker, LS" journalOrPublisher="Bishop Museum bulletin in Zoology" pageId="0" pageNumber="13" pagination="1 - 128" refId="B23" refString="Kornicker, LS, Harrison-Nelson, E, Coles, SL, 2007. Ostracoda (Myodocopina) from O'ahu and French Frigate Shoals, Hawaiian Islands. Bishop Museum bulletin in Zoology 8: 1 - 128" title="Ostracoda (Myodocopina) from O'ahu and French Frigate Shoals, Hawaiian Islands." volume="8" year="2007">Kornicker et al. 2007</bibRefCitation>
),
<taxonomicName lsidName="P. hulingsi" pageId="0" pageNumber="13" rank="species" species="hulingsi">
<emphasis italics="true" pageId="0" pageNumber="13">P. hulingsi</emphasis>
</taxonomicName>
Baker, 1978 from California (
<bibRefCitation DOI="https://doi.org/10.1163/156854078X00042" author="Baker, JH" journalOrPublisher="Crustaceana" pageId="0" pageNumber="13" pagination="139 - 151" refId="B1" refString="Baker, JH, 1978. Two new species of Parasterope (Myodocopina, Ostracoda) from southern California. Crustaceana 35 (2): 139 - 151, DOI: https://doi.org/10.1163/156854078X00042" title="Two new species of Parasterope (Myodocopina, Ostracoda) from southern California." url="https://doi.org/10.1163/156854078X00042" volume="35" year="1978">Baker 1978</bibRefCitation>
),
<taxonomicName lsidName="P. jenseni" pageId="0" pageNumber="13" rank="species" species="jenseni">
<emphasis italics="true" pageId="0" pageNumber="13">P. jenseni</emphasis>
</taxonomicName>
from Sagami Sea, Japan (
<bibRefCitation author="Poulsen, EM" journalOrPublisher="Marine Biodiversity" pageId="0" pageNumber="13" refId="B33" refString="Poulsen, EM, 1965. Ostracoda - Myodocopa, Part II: Cypridiniformes - Rutidermatidae, Sarsiellidae and Asteropidae. Dana Report Carlsberg Foundation (65): 1-484." title="Ostracoda - Myodocopa, Part II: Cypridiniformes - Rutidermatidae, Sarsiellidae and Asteropidae. Dana Report Carlsberg Foundation (65): 1 - 484." year="1965">Poulsen 1965</bibRefCitation>
),
<taxonomicName lsidName="P. lagunicola" pageId="0" pageNumber="13" rank="species" species="lagunicola">
<emphasis italics="true" pageId="0" pageNumber="13">P. lagunicola</emphasis>
</taxonomicName>
Hartmann, 1984 from French Polynesia (
<bibRefCitation author="Hartmann, G" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="0" pageNumber="13" pagination="117 - 169" refId="B10" refString="Hartmann, G, 1984. Zur Kenntnis der Ostracoden der polynesischen Inseln Huahine (Gesellschaftsinseln) und Rangiroa (Tuamotu-Inseln). Mit Bemerkungen zur Verbreitung und Ausbreitung litoraler Ostracoden und einer Uebersicht ueber die bislang auf den pazifischen Inseln gefunden. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 81: 117 - 169" title="Zur Kenntnis der Ostracoden der polynesischen Inseln Huahine (Gesellschaftsinseln) und Rangiroa (Tuamotu-Inseln). Mit Bemerkungen zur Verbreitung und Ausbreitung litoraler Ostracoden und einer Uebersicht ueber die bislang auf den pazifischen Inseln gefunden." volume="81" year="1984">Hartmann 1984</bibRefCitation>
),
<taxonomicName lsidName="P. mckenziei" pageId="0" pageNumber="13" rank="species" species="mckenziei">
<emphasis italics="true" pageId="0" pageNumber="13">P. mckenziei</emphasis>
</taxonomicName>
from the Philippines (
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.39" author="Kornicker, LS" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="13" pagination="1 - 32" refId="B15" refString="Kornicker, LS, 1970. Myodocopid Ostracoda (Cypridinacea) from the Philippine Islands. Smithsonian Contributions to Zoology 39: 1 - 32, DOI: https://doi.org/10.5479/si.00810282.39" title="Myodocopid Ostracoda (Cypridinacea) from the Philippine Islands." url="https://doi.org/10.5479/si.00810282.39" volume="39" year="1970">Kornicker 1970</bibRefCitation>
),
<taxonomicName lsidName="P. muelleri" pageId="0" pageNumber="13" rank="species" species="muelleri">
<emphasis italics="true" pageId="0" pageNumber="13">P. muelleri</emphasis>
</taxonomicName>
Skogsberg, 1920 from England (
<bibRefCitation author="Poulsen, EM" journalOrPublisher="Marine Biodiversity" pageId="0" pageNumber="13" refId="B33" refString="Poulsen, EM, 1965. Ostracoda - Myodocopa, Part II: Cypridiniformes - Rutidermatidae, Sarsiellidae and Asteropidae. Dana Report Carlsberg Foundation (65): 1-484." title="Ostracoda - Myodocopa, Part II: Cypridiniformes - Rutidermatidae, Sarsiellidae and Asteropidae. Dana Report Carlsberg Foundation (65): 1 - 484." year="1965">Poulsen 1965</bibRefCitation>
),
<taxonomicName lsidName="P. prolixa" pageId="0" pageNumber="13" rank="species" species="prolixa">
<emphasis italics="true" pageId="0" pageNumber="13">P. prolixa</emphasis>
</taxonomicName>
Kornicker, 1975 from Australia (
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.163" author="Kornicker, LS" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="13" pagination="375 - 720" refId="B16" refString="Kornicker, LS, 1975. Antarctic Ostracoda (Myodocopina), Part 2. Smithsonian Contributions to Zoology 163: 375 - 720, DOI: https://doi.org/10.5479/si.00810282.163" title="Antarctic Ostracoda (Myodocopina), Part 2." url="https://doi.org/10.5479/si.00810282.163" volume="163" year="1975">Kornicker 1975</bibRefCitation>
),
<taxonomicName lsidName="P. sequex" pageId="0" pageNumber="13" rank="species" species="sequex">
<emphasis italics="true" pageId="0" pageNumber="13">P. sequex</emphasis>
</taxonomicName>
Kornicker &amp; Poore, 1996 from Australia (
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.573" author="Kornicker, LS" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="13" pagination="1 - 186" refId="B22" refString="Kornicker, LS, Poore, GC, 1996. Ostracoda (Myodocopina) of the SE Australian Continental Slope: Part 3. Smithsonian Contributions to Zoology 573: 1 - 186, DOI: https://doi.org/10.5479/si.00810282.573" title="Ostracoda (Myodocopina) of the SE Australian Continental Slope: Part 3." url="https://doi.org/10.5479/si.00810282.573" volume="573" year="1996">Kornicker and Poore 1996</bibRefCitation>
),
<taxonomicName lsidName="P. styx" pageId="0" pageNumber="13" rank="species" species="styx">
<emphasis italics="true" pageId="0" pageNumber="13">P. styx</emphasis>
</taxonomicName>
from Chile (
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.163" author="Kornicker, LS" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="13" pagination="375 - 720" refId="B16" refString="Kornicker, LS, 1975. Antarctic Ostracoda (Myodocopina), Part 2. Smithsonian Contributions to Zoology 163: 375 - 720, DOI: https://doi.org/10.5479/si.00810282.163" title="Antarctic Ostracoda (Myodocopina), Part 2." url="https://doi.org/10.5479/si.00810282.163" volume="163" year="1975">Kornicker 1975</bibRefCitation>
), and
<taxonomicName lsidName="P. theta" pageId="0" pageNumber="13" rank="species" species="theta">
<emphasis italics="true" pageId="0" pageNumber="13">P. theta</emphasis>
</taxonomicName>
from
<normalizedToken originalValue="Waikïkï">Waikiki</normalizedToken>
(
<bibRefCitation author="Kornicker, LS" journalOrPublisher="Bishop Museum bulletin in Zoology" pageId="0" pageNumber="13" pagination="1 - 128" refId="B23" refString="Kornicker, LS, Harrison-Nelson, E, Coles, SL, 2007. Ostracoda (Myodocopina) from O'ahu and French Frigate Shoals, Hawaiian Islands. Bishop Museum bulletin in Zoology 8: 1 - 128" title="Ostracoda (Myodocopina) from O'ahu and French Frigate Shoals, Hawaiian Islands." volume="8" year="2007">Kornicker et al. 2007</bibRefCitation>
). The three other new species,
<taxonomicName lsidName="P. busanensis" pageId="0" pageNumber="13" rank="species" species="busanensis">
<emphasis italics="true" pageId="0" pageNumber="13">P. busanensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. sagami" pageId="0" pageNumber="13" rank="species" species="sagami">
<emphasis italics="true" pageId="0" pageNumber="13">P. sagami</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="P. sohi" pageId="0" pageNumber="13" rank="species" species="sohi">
<emphasis italics="true" pageId="0" pageNumber="13">P. sohi</emphasis>
</taxonomicName>
also have two oblique rows. The exopodite of the A2 in
<taxonomicName lsidName="P. singula" pageId="0" pageNumber="13" rank="species" species="singula">
<emphasis italics="true" pageId="0" pageNumber="13">P. singula</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. sohi" pageId="0" pageNumber="13" rank="species" species="sohi">
<emphasis italics="true" pageId="0" pageNumber="13">P. sohi</emphasis>
</taxonomicName>
carries minute spines at inner terminal corner on the 5th-8th joints, while these spines are present from the 1st-8th joints in
<taxonomicName lsidName="P. busanensis" pageId="0" pageNumber="13" rank="species" species="busanensis">
<emphasis italics="true" pageId="0" pageNumber="13">P. busanensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. sagami" pageId="0" pageNumber="13" rank="species" species="sagami">
<emphasis italics="true" pageId="0" pageNumber="13">P. sagami</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="13">GenBank numbers 16S: OK048686, OK048687; 18S OK048723.</paragraph>
</subSubSection>
</treatment>
</document>

104
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1
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&quot;Habitat in
<taxonomicName id="4D065CD197835869BA91A3929B176E34" authorityName="A.Nageswara Rao" authorityYear="1999" class="Liliopsida" family="Orchidaceae" genus="India" kingdom="Plantae" order="Asparagales" phylum="Tracheophyta" rank="genus">India</taxonomicName>
.&quot; RCN: 197.
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<paragraph id="F96C59D35EF916F7BDE8C08A64CECACA" pageNumber="812">
<materialsCitation id="0D76CAF0F6009D98C0C05B7D9D144FC8" ID-GBIF-Occurrence="2612020885" collectionCode="P-TOURN" specimenCode="1079" typeStatus="neotype">
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(Hedge in Jarvis &amp; al. in
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50: 517. 2001): Herb. Tournefort No. 1079 (
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Current name:
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L.
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(
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<emphasis id="78E3FF7814F6195D3EEEDE47C9B41306" italics="true" pageNumber="812">Note:</emphasis>
Hedge (in Davis,
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7: 451. 1982; in Rechinger,
<emphasis id="4CBCB0001999B1755D42AD937BFB63E5" italics="true" pageNumber="812">Fl. Iranica</emphasis>
150: 466. 1982) noted the absence of material in LINN and suggested that a Tournefort collection &quot;could possibly be selected as type&quot;. He subsequently (2001) formalised this suggestion by designating it as a
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64
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<taxonomicName authority="Warren 1896" authorityYear="1896" class="Insecta" family="Geometridae" genus="Eois" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eois (Psilocambogia) semirubra" order="Lepidoptera" pageId="0" pageNumber="5447" phylum="Arthropoda" rank="species" species="semirubra" subGenus="Psilocambogia">Eois (Psilocambogia) semirubra (Warren 1896)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="5447" type="reference_group">
<paragraph pageId="0" pageNumber="5447">
<taxonomicName class="Insecta" family="Geometridae" genus="Eois" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eois (Psilocambogia) semirubra" order="Lepidoptera" pageId="0" pageNumber="5447" phylum="Arthropoda" rank="species" species="semirubra" subGenus="Psilocambogia">Eois (Psilocambogia) semirubra</taxonomicName>
<bibRefCitation author="Warren, W." journalOrPublisher="Novitates Zoologicae" pageId="0" pageNumber="5447" pagination="335 - 419" title="New species of Drepanulidae, Thyrididae, Uraniidae, Epiplemidae, and Geometridae in the Tring Museum" volume="3" year="1896">Warren 1896</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="5447" type="materials_examined">
<paragraph pageId="0" pageNumber="5447">Materials</paragraph>
<paragraph pageId="0" pageNumber="5447">
<materialsCitation pageId="0" pageNumber="5447" specimenCount="1" typeStatus="Holotype">
Type status:
<typeStatus pageId="0" pageNumber="5447">Holotype</typeStatus>
. Occurrence: sex:
<specimenType pageId="0" pageNumber="5447">m</specimenType>
; Record Level: ownerInstitutionCode: NHM
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="5447" type="distribution">
<paragraph pageId="0" pageNumber="5447">Distribution</paragraph>
<paragraph pageId="0" pageNumber="5447">Type locality: [West Papua], Humboldt Bay [Yos Sudarso Bay]</paragraph>
</subSubSection>
</treatment>
</document>

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; Record Level: bibliographicCitation: Rambousek (1912: 99)
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<paragraph pageId="4" pageNumber="53">Taxon classification Animalia Coleoptera Cantharidae</paragraph>
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<taxonomicName authority="Pic, 1906" authorityName="Pic" authorityYear="1906" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius impressipennis" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="impressipennis">Paramaronius impressipennis (Pic, 1906)</taxonomicName>
Figs 10-13, 17, 19, 21, 25, 29, 33
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53" type="remarks">
<paragraph pageId="4" pageNumber="53">Remarks.</paragraph>
<paragraph lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53">
This species was briefly described by
<bibRefCitation author="Pic, M" journalOrPublisher="L'Echange, Revue Linneenne" pageId="13" pageNumber="62" pagination="51 - 52" title="Sur divers Maronius Gorh. et genres voisins de l'Amerique Meridionale." volume="22" year="1906">Pic (1906)</bibRefCitation>
based on a male from
<normalizedToken originalValue="Tucumán">Tucuman</normalizedToken>
, northern Argentina. However, some specimens have been collected in the
<pageBreakToken pageId="5" pageNumber="54" start="start">northeastern</pageBreakToken>
region of Brazil (states of Bahia and
<normalizedToken originalValue="Maranhão">Maranhao</normalizedToken>
), far away from the type locality. The comparison of the Brazilian specimens with the holotype (MNHN) shows there are no observable differences within them and they are, indeed, the same species.
</paragraph>
<paragraph pageId="5" pageNumber="54">
Even though these localities are so distant, they belong to Cerrado and Chaco provinces, adjacent areas of the same biogeographic subregion (Chaco dominion) (
<bibRefCitation pageId="5" pageNumber="54">Morrone 2000</bibRefCitation>
, 2006) (Fig. 44).
</paragraph>
<paragraph pageId="5" pageNumber="54">
A redescription of
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius impressipennis" order="Coleoptera" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="impressipennis">Paramaronius impressipennis</taxonomicName>
is presented as well as a supplementary description of the female for the first time to complete its morphological information after specimens from Brazil.
</paragraph>
</subSubSection>
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<paragraph pageId="5" pageNumber="54">Material examined.</paragraph>
<paragraph lastPageId="6" lastPageNumber="55" pageId="5" pageNumber="54">
HOLOTYPE ♂ (MNHN): ARGENTINA:
<normalizedToken originalValue="Tucumán">Tucuman</normalizedToken>
, 4.i.1900. BRAZIL:
<normalizedToken originalValue="Maranhão">Maranhao</normalizedToken>
, Mirador (Parque Estadual do Mirador, Base da Geraldina)
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-6.776944">6°46'37&quot;S</geoCoordinate>
;
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-45.109444">45°06'34&quot;W</geoCoordinate>
, 22.
<normalizedToken originalValue="ii–">ii-</normalizedToken>
01.iii.2009, armadilha luminosa [light trap], F. Limeira-de-Oliveira col. (4♂, 14♀ MZSP, 4♂, 15♀ CZMA); same locality, (Parque Estadual do Mirador, Base do Mosquito), 04-08.ii.2011, armadilha luminosa, F. Limeira-de-Oliveira col. (1♀) (CZMA); Bahia, Barreiras (Estrada Mata de Cachoei
<pageBreakToken pageId="6" pageNumber="55" start="start">ras</pageBreakToken>
, Acaba Vida, km 7),
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-11.872222">11°52'20&quot;S</geoCoordinate>
;
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-45.54861">45°32'55&quot;W</geoCoordinate>
, 23.i.2009, luz [light trap], Nihei, Figueiredo, Almeida &amp; Cezar col. (1♀) (MZSP).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="56" pageId="6" pageNumber="55" type="redescription">
<paragraph pageId="6" pageNumber="55">Redescription.</paragraph>
<paragraph lastPageId="7" lastPageNumber="56" pageId="6" pageNumber="55">
Head with vertex, frons, clypeus and bases of mandibles pale yellow, slightly translucent; apex of mandibles dark brown; occipital region with a V-shaped testaceous mark; labial and maxillary palpi pale yellow, last palpomere brown. Antennae light brown, three first antennomeres pale yellow. Pronotum pale yellow to light brown,
<pageBreakToken pageId="7" pageNumber="56" start="start">sometimes</pageBreakToken>
translucent. Scutellum pale yellow with apex translucent. Elytra testaceous to light brown with a lateral longitudinal darker brown band from the epipleura to the posterior third quarter of dorsal surface; apex of elytra with an oblique yellow band on outer margin; on less pigmented specimens, pronotum pale yellow and lateral and apical patches indistinct. Hind wings light brown. Legs testaceous to light brown; tarsi and dorsal surface of tibiae darker. Ventrally, thorax and abdomen pale yellow.
</paragraph>
<paragraph pageId="7" pageNumber="56">Male (Fig. 10): body covered by dense and fine pubescence. Lateral margins of head arcuate behind eyes; vertex slightly convex, frons flat. Eyes prominent. Antennae short, with one antennomere exceeding the apex of elytra. Antennomere 1 long and swollen, 2.7 times longer than wide; antennomere 2 short, third antennomere 1.7 times shorter than antennomere 1, forth to seventh subequal in length, slightly shorter than antennomere 1, the latter progressively shorter than seventh. Pronotum subrectangular, shiny, slightly narrower near fore angles; anterior margin slightly rounded. Scutellum triangular, apex truncate. Elytra short, 1.5 times longer than wide, covered by fine hairs and some sparse black hairs. Apical half modified, forming a longitudinal ridge and two sloped surfaces covered by very short hairs (Figs 11, 12). Legs slender; hind tibia as long as hind femur; tarsomeres gradually increasing in size from fore to hind legs; first metatarsomere 1.7 times longer than second and 2.7 times longer than third. Abdominal glandular pores slightly prominent. Seventh abdominal ventrite (Fig. 17) wider than long, deeply emarginate, forming two distal lobes, narrowed apicad. Aedeagus (Fig. 25) with right prolongation of tegmen large and arcuate on posterior margin, and covered with large setae; dorsal surface with a short and rounded apophysis; left setiferous prolongation very long and curved, clubbed apically, with few setae on its apex. Right paramere (Fig. 29) short, narrowing apicad, then divided at apex, forming an inclined prolongation, reaching apophysis of tegmen; left paramere flat, very short and wide, divided apically and partially covering dorsal surface of median lobe; median lobe long and curved to right, sometimes retracted. Ventral and lateral surfaces of tegmen (Fig. 33) with fine scratches.</paragraph>
<caption pageId="7" pageNumber="56">
<paragraph pageId="7" pageNumber="56">
Figures 10-13. 10-11
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius impressipennis" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="impressipennis">Paramaronius impressipennis</taxonomicName>
(Pic), holotype 10 habitus, dorsal view 11 elytra, dorsal view 12-13
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius impressipennis" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="impressipennis">Paramaronius impressipennis</taxonomicName>
(Pic) from Brazil 12 male elytra, dorsal view 13 female elytra, dorsal view. Scale bars: 2.0 mm.
</paragraph>
</caption>
<caption pageId="7" pageNumber="56">
<paragraph pageId="7" pageNumber="56">
Figures 14-21. 14-17 Seventh abdominal ventrite of male, ventral view 18-19 seventh abdominal ventrite of female, ventral view 20-21 female right coxite, ventral view 14, 18, 20
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius serranus" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="serranus">Paramaronius serranus</taxonomicName>
sp. n. 15
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius brancuccii" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="brancuccii">Paramaronius brancuccii</taxonomicName>
sp. n. 16
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius cavipennis" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="cavipennis">Paramaronius cavipennis</taxonomicName>
sp. n. 17, 19, 21
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius impressipennis" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="impressipennis">Paramaronius impressipennis</taxonomicName>
(Pic). Scale bars: 0.5 mm (14-19); 0.2 mm (20-21).
</paragraph>
</caption>
<caption pageId="7" pageNumber="56">
<paragraph pageId="7" pageNumber="56">
Figures 22-29. 22-25 Aedeagus, dorsal view 26-30 right paramere, dorsal view 22, 26
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius serranus" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="serranus">Paramaronius serranus</taxonomicName>
sp. n. 23, 27
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius brancuccii" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="brancuccii">Paramaronius brancuccii</taxonomicName>
sp. n. 24, 28
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius cavipennis" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="cavipennis">Paramaronius cavipennis</taxonomicName>
sp. n. 25, 30
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius impressipennis" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="impressipennis">Paramaronius impressipennis</taxonomicName>
(Pic). Scale bars: 0.5 mm (22-25); 0.2 mm (26-29).
</paragraph>
</caption>
<caption pageId="7" pageNumber="56">
<paragraph pageId="7" pageNumber="56">
Figures 30-33. Aedeagus, left view 30
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius serranus" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="serranus">Paramaronius serranus</taxonomicName>
sp. n. 31
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius brancuccii" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="brancuccii">Paramaronius brancuccii</taxonomicName>
sp. n. 32
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius cavipennis" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="cavipennis">Paramaronius cavipennis</taxonomicName>
sp. n. 33
<taxonomicName class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paramaronius impressipennis" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="impressipennis">Paramaronius impressipennis</taxonomicName>
(Pic). Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="7" pageNumber="56">Female (Fig. 13) slightly bigger than male; antennomere 1 slender, not swollen; dorsal surface of elytra without sculptural modification, brown lateral and yellow apical bands more distinct. Seventh abdominal ventrite (Fig. 19) trapezoidal, distal margin straight with a short, rounded notch. Coxites (Fig. 21) small and membranous; styles short, wider apically.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="56" type="distribution">
<paragraph pageId="7" pageNumber="56">Distribution.</paragraph>
<paragraph pageId="7" pageNumber="56">
Argentina (
<normalizedToken originalValue="Tucumán">Tucuman</normalizedToken>
) and Brazil (
<normalizedToken originalValue="Maranhão">Maranhao</normalizedToken>
and Bahia) (Fig. 44).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="56" type="biological data">
<paragraph pageId="7" pageNumber="56">Biological data.</paragraph>
<paragraph pageId="7" pageNumber="56">
The specimens from
<normalizedToken originalValue="Maranhão">Maranhao</normalizedToken>
and Bahia were collected on Brazilian savannah (cerrado) and were attracted by light traps.
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<taxonomicName LSID="urn:lsid:zoobank.org:act:9ACD0F86-341A-4855-925A-51104BB8C8F4" authority="Klimaszewski &amp; Godin" class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta (Dimetrota) whitehorsensis" order="Coleoptera" pageId="6" pageNumber="213" phylum="Arthropoda" rank="species" species="whitehorsensis" subGenus="Dimetrota">Atheta (Dimetrota) whitehorsensis Klimaszewski &amp; Godin</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="213">sp. n.</taxonomicNameLabel>
Figs 620, 2140-43
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<paragraph pageId="6" pageNumber="213">Holotype (male).</paragraph>
<paragraph pageId="6" pageNumber="213">Canada, Yukon, Whitehorse, Granger, 60.7078, -135.0971, 25.VIII.2007, 657 m, soil sifting, black spruce stand, AWT, B. Godin (LFC).</paragraph>
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<paragraph pageId="6" pageNumber="213">Paratype.</paragraph>
<paragraph pageId="6" pageNumber="213">Canada, Yukon, Whitehorse, Granger, 60.7078, -135.0971, 5.VIII.2007, 657 m, soil sifting, black spruce stand, AWT, B. Godin (ECW) 1 female.</paragraph>
</subSubSection>
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<paragraph pageId="6" pageNumber="213">Etymology.</paragraph>
<paragraph pageId="6" pageNumber="213">The specific name derives from the name of the type locality, which is Whitehorse, Yukon.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="213" type="diagnosis">
<paragraph pageId="6" pageNumber="213">Diagnosis.</paragraph>
<paragraph pageId="6" pageNumber="213">Body narrowly oval, dark brown to black, with bases of antennae and legs rust-brown, surface matte, with asperate dense punctation on forebody and strong meshed microsculpture (Fig. 6); length 1.9-2.0 mm; head narrower than pronotum and elytra, with short postocular area, eyes large and slightly protruding; antennae slender, slightly incrassate apically, article 4 subquadrate, 5 slightly elongate and 6-10 slightly to strongly transverse; pronotum strongly transverse and broadest in the middle; elytra transverse, longer than pronotum; abdomen broadly arcuate laterally (Fig. 6). MALE: tergite 8 transverse and truncate apically (Fig. 40); sternite 8 widely rounded apically (Fig. 41); median lobe of aedeagus with venter of tubus straight and short, and apex sharply produced (Fig. 20). FEMALE: tergite and sternite 8 truncate apically (Figs 42, 43); spermatheca with pipe-shaped capsule and long stem hooked posteriorly (Fig. 21).</paragraph>
<paragraph pageId="6" pageNumber="213">
This species is similar externally to
<taxonomicName class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta (Dimetrota) hampshirensis" order="Coleoptera" pageId="6" pageNumber="213" phylum="Arthropoda" rank="species" species="hampshirensis" subGenus="Dimetrota">Atheta (Dimetrota) hampshirensis</taxonomicName>
Bernhauer and
<taxonomicName class="Insecta" family="Staphylinidae" genus="Atheta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atheta (Datomicra) dadopora" order="Coleoptera" pageId="6" pageNumber="213" phylum="Arthropoda" rank="species" species="dadopora" subGenus="Datomicra">Atheta (Datomicra) dadopora</taxonomicName>
Thomson but differs in the shape of the spermatheca and median lobe of the aedeagus, and has a broader body than the latter species.
</paragraph>
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<paragraph pageId="6" pageNumber="213">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="213">This native Nearctic species is known only from the type locality in the Yukon Territory.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="213" type="bionomics">
<paragraph pageId="6" pageNumber="213">Bionomics.</paragraph>
<paragraph pageId="6" pageNumber="213">Adults were captured by sifting soil in a black spruce stand.</paragraph>
</subSubSection>
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28. Ranunculus calycibus patulis, pedunculis teretibus, foliis tripartito-multifidis: summis linearibus.
<emphasis id="D9E58117816DC14E718BC9905EEB3908" italics="true">Fl. suec. 466.</emphasis>
<emphasis id="1201B6E85C6838DFCD734F31BBBF15DD" italics="true">Dalib. paris. 164.</emphasis>
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Ranunculus foliis peltatis quiquangularibus multipartitis: laciniis linearibus, caule multifloro.
<emphasis id="A47F0C154566328DEE1E0A722B4DE69B" italics="true">Hort. cliff. 231.</emphasis>
<emphasis id="6046FC9F4314101C936FEBD54A9A6376" italics="true">Roy. lugdb. 492.</emphasis>
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Ranunculus pratensis erectus acris.
<emphasis id="6F46DF55F5C5105D0C229A5E5CA10355" italics="true">Bauh. pin. 178.</emphasis>
<emphasis id="995C24FE55600FF86110821E1F1D542D" italics="true">Fl. lapp. 228.</emphasis>
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Ranunculus hortensis.
<emphasis id="BCF06F64F99D3255137C2047DB7E676B" italics="true">Dod. pempt. 426.</emphasis>
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β. Ranunculus hortensis erectus, flore pleno.
<emphasis id="E9455EF9B216DEBDB9D48976D4D6CFF9" italics="true">Bauh. pin. 179.</emphasis>
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<emphasis id="CC1E7B92A4F98CD965887960BBA40204" italics="true">Habitat in</emphasis>
Europae
<emphasis id="F5D5600F074005D8B3D98C1D1DDD5A22" italics="true">pratis, pascuis. ♃</emphasis>
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<paragraph pageNumber="752">&quot;Habitat in Virginia, Canada. D. Kalm.&quot; RCN: 578.</paragraph>
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Current name:
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3.
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Plants forming semi-globose cushions to
<quantity id="F95AF8B4FFD3FF8DFDABCD267EE2FEF9" box="[610,681,302,328]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="52" unit="cm" value="30.0">30 cm</quantity>
large. Stems terete, woody,
<quantity id="F95AF8B4FFD3FF8DFC38CD267830FEF9" box="[1009,1147,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="5" pageNumber="52" unit="mm" value="1.7" valueMax="1.9" valueMin="1.5">1.5–1.9 mm</quantity>
in diameter near the base, the upper branches
<quantity id="F95AF8B4FFD3FF8DFEA2CD5A7DBEFEDD" box="[363,501,338,364]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.1" metricValueMin="1.6" pageId="5" pageNumber="52" unit="mm" value="1.85" valueMax="2.1" valueMin="1.6">1.6–2.1 mm</quantity>
in diameter including leaves. Leaves numerous, persistent, only the terminal ones green; petiole white to pale brown with violet streaks, flat, appressed, to
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long and
<quantity id="F95AF8B4FFD3FF8DFBB1CD7E7885FE21" box="[1144,1230,374,401]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="52" unit="mm" value="0.5">0.5 mm</quantity>
broad, ciliate along the margins; lamina erect to spreading, glossy-green, tubercled, flat to semi-terete, to
<quantity id="F95AF8B4FFD3FF8DFB9DCD9278E0FE05" box="[1108,1195,410,437]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="52" unit="mm" value="0.5">0.5 mm</quantity>
thick, narrowly ovate, 1.5–1.9 ×
<quantity id="F95AF8B4FFD3FF8DFF35CDB67DCEFE69" box="[252,389,446,472]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="5" pageNumber="52" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.6–0.9 mm</quantity>
, the apex obtuse, crowned with (4–)6–8(–11) spreading to erect pellucid trichomes to
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long. Inflorescences capitoid, 1–3-flowered, (sub)sessile with the peduncles to
<quantity id="F95AF8B4FFD3FF8DFC30CDEA7804FE4D" box="[1017,1103,482,509]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="52" unit="mm" value="0.5">0.5 mm</quantity>
long, the outer bracts leaf-like, ovate, 1.4–1.9 ×
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, the inner bracts oblong, 1.2–1.7 ×
<quantity id="F95AF8B4FFD3FF8DFCBECE0E7848FD91" box="[887,1027,518,545]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="5" pageNumber="52" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 mm</quantity>
, both
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of bracts ciliate along the margins and with a few pellucid trichomes at the apex. Flowers gynodioecious; calyx vestigial; corolla white, 3-lobed, funnelform, the tube
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long, the lobes spreading, triangular to ovate, obtuse, 0.9–1.3 ×
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; stamens exserted and spreading, the thecae oblong,
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long; pistil with the style exserted to
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, 2-parted, the branches to
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long. Fruits unknown.
</paragraph>
<paragraph id="3E1D5551FFD3FF8DFF74CEB27EA9FC1D" blockId="5.[136,1452,302,1300]" pageId="5" pageNumber="52">
Among the high-elevation species of
<taxonomicName id="F9A22ED2FFD3FF8DFDBDCEB27EAAFD65" box="[628,737,698,724]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="genus">
<emphasis id="0CD68943FFD3FF8DFDBDCEB27EAAFD65" box="[628,737,698,724]" italics="true" pageId="5" pageNumber="52">Valeriana</emphasis>
</taxonomicName>
from the tropical Andes, the cushion habit is also present in
<taxonomicName id="F9A22ED2FFD3FF8DFF41CED77E03FD49" authority="Graebner (1945: 37)" authorityName="Graebner" authorityPageNumber="37" authorityYear="1945" box="[136,584,734,761]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="species" species="aschersoniana">
<emphasis id="0CD68943FFD3FF8DFF41CED77D07FD49" box="[136,332,734,760]" italics="true" pageId="5" pageNumber="52">V. aschersoniana</emphasis>
Graebner (1945: 37)
</taxonomicName>
from
<collectingCountry id="46B515C1FFD3FF8DFD5ECED67E81FD49" box="[663,714,734,760]" name="Peru" pageId="5" pageNumber="52">Peru</collectingCountry>
and
<taxonomicName id="F9A22ED2FFD3FF8DFCC2CED778C8FD49" authority="Kunth (1819: 324)" authorityName="Kunth" authorityPageNumber="324" authorityYear="1819" box="[779,1155,734,760]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="species" species="aretioides">
<emphasis id="0CD68943FFD3FF8DFCC2CED77FD0FD49" box="[779,923,734,760]" italics="true" pageId="5" pageNumber="52">V. aretioides</emphasis>
<bibRefCitation id="5A3328A0FFD3FF8DFC6ECED678C8FD49" author="Kunth, K. S." box="[935,1155,734,760]" pageId="5" pageNumber="52" pagination="324" refId="ref4387" refString="Kunth, K. S. (1819) Nova Genera et Species Plantarum (quarto ed.) 3: 324." type="journal article" year="1819">Kunth (1819: 324)</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="46B515C1FFD3FF8DFB1BCED6797BFD49" box="[1234,1328,734,760]" name="Ecuador" pageId="5" pageNumber="52">Ecuador</collectingCountry>
–
<collectingCountry id="46B515C1FFD3FF8DFAF4CED679E0FD49" box="[1341,1451,734,760]" name="Colombia" pageId="5" pageNumber="52">Colombia</collectingCountry>
(
<bibRefCitation id="5A3328A0FFD3FF8DFF46CF0A7DF7FCAD" author="Weberling, F. &amp; Uhlarz, H." box="[143,444,770,796]" pageId="5" pageNumber="52" pagination="217 - 242" refId="ref4850" refString="Weberling, F. &amp; Uhlarz, H. (1977) Morphologische, anatomische und palynologische Untersuchungen an der Gattung Aretiastrum (Valerianaceae). Plant Systematics and Evolution 127: 217 - 242. https: // doi. org / 10.1007 / BF 00985989" type="journal article" year="1977">Weberling &amp; Uhlarz 1977</bibRefCitation>
, Eriksen 1989). However, both species have typically 5-lobed corollas, although flowers are occasionally 3–4-lobed in
<taxonomicName id="F9A22ED2FFD3FF8DFE2ECF2F7FBFFCF1" authority="(Weberling &amp; Stutzel 2006)" baseAuthorityName="Weberling &amp; Stutzel" baseAuthorityYear="2006" box="[487,1012,806,832]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="species" species="aschersoniana">
<emphasis id="0CD68943FFD3FF8DFE2ECF2F7EE3FCF1" box="[487,680,806,832]" italics="true" pageId="5" pageNumber="52">V. aschersoniana</emphasis>
(
<bibRefCitation id="5A3328A0FFD3FF8DFD73CF2E7FA6FCF1" author="Weberling, F. &amp; Stutzel, T." box="[698,1005,806,832]" pageId="5" pageNumber="52" pagination="193 - 205" refId="ref4815" refString="Weberling, F. &amp; Stutzel, T. (2006) Morphological and anatomical investigations of Aretiastrum magellanicum (Hombr. &amp; Jacq.) Skottsb. Wulfenia 13: 193 - 205." type="journal article" year="2006">Weberling &amp; Stützel 2006</bibRefCitation>
)
</taxonomicName>
and 4-lobed in
<taxonomicName id="F9A22ED2FFD3FF8DFB78CF2F7C87FCD5" authority="(Eriksen 1989)" baseAuthorityName="Eriksen" baseAuthorityYear="1989" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="species" species="aretioides">
<emphasis id="0CD68943FFD3FF8DFB78CF2F7889FCF1" box="[1201,1218,807,832]" italics="true" pageId="5" pageNumber="52">V</emphasis>
.
<emphasis id="0CD68943FFD3FF8DFB1ACF2E7909FCF1" box="[1235,1346,806,832]" italics="true" pageId="5" pageNumber="52">aretioides</emphasis>
(Eriksen 1989)
</taxonomicName>
, and leaf laminas lack the distinct crown of trichomes at the apex (
<bibRefCitation id="5A3328A0FFD3FF8DFC09CF4278A0FCD5" author="Weberling, F. &amp; Uhlarz, H." box="[960,1259,842,868]" pageId="5" pageNumber="52" pagination="217 - 242" refId="ref4850" refString="Weberling, F. &amp; Uhlarz, H. (1977) Morphologische, anatomische und palynologische Untersuchungen an der Gattung Aretiastrum (Valerianaceae). Plant Systematics and Evolution 127: 217 - 242. https: // doi. org / 10.1007 / BF 00985989" type="journal article" year="1977">Weberling &amp; Uhlarz 1977</bibRefCitation>
), which provides the cushions of
<taxonomicName id="F9A22ED2FFD3FF8DFEFDCF677DBDFC39" authorityName="Sklenar &amp; B. Eriksen" authorityYear="2023" box="[308,502,878,904]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="species" species="xenophylloides">
<emphasis id="0CD68943FFD3FF8DFEFDCF677DBDFC39" box="[308,502,878,904]" italics="true" pageId="5" pageNumber="52">V. xenophylloides</emphasis>
</taxonomicName>
a slightly silvery-white appearance. Moreover, the flowers of
<taxonomicName id="F9A22ED2FFD3FF8DFB6DCF677966FC39" baseAuthorityName="Eriksen" baseAuthorityYear="1989" box="[1188,1325,878,904]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="species" species="aretioides">
<emphasis id="0CD68943FFD3FF8DFB6DCF677966FC39" box="[1188,1325,878,904]" italics="true" pageId="5" pageNumber="52">V. aretioides</emphasis>
</taxonomicName>
are yellow. Confusion with other species of
<taxonomicName id="F9A22ED2FFD3FF8DFE3ACF9A7E2BFC1D" box="[499,608,914,940]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="genus">
<emphasis id="0CD68943FFD3FF8DFE3ACF9A7E2BFC1D" box="[499,608,914,940]" italics="true" pageId="5" pageNumber="52">Valeriana</emphasis>
</taxonomicName>
is unlikely.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD3FF8DFF74CFBD7D4AFB8D" pageId="5" pageNumber="52" type="distribution">
<paragraph id="3E1D5551FFD3FF8DFF74CFBD7D4AFB8D" blockId="5.[136,1452,302,1300]" pageId="5" pageNumber="52">
<emphasis id="0CD68943FFD3FF8DFF74CFBD7E42FC7E" bold="true" box="[189,521,949,976]" pageId="5" pageNumber="52">Distribution and habitat:—</emphasis>
<taxonomicName id="F9A22ED2FFD3FF8DFDC0CFBE7F6CFC61" authorityName="Sklenar &amp; B. Eriksen" authorityYear="2023" box="[521,807,950,976]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="species" species="xenophylloides">
<emphasis id="0CD68943FFD3FF8DFDC0CFBE7F6CFC61" box="[521,807,950,976]" italics="true" pageId="5" pageNumber="52">Valeriana xenophylloides</emphasis>
</taxonomicName>
was collected only once in a very humid bamboo-shrub páramo of southern
<collectingCountry id="46B515C1FFD3FF8DFEA3CFD27D81FC45" box="[362,458,986,1012]" name="Ecuador" pageId="5" pageNumber="52">Ecuador</collectingCountry>
. It grew in patches of open vegetation on wet slopes with a rich cover of bryophytes and lichens (
<figureCitation id="A69949D4FFD3FF8DFF2FCFF67D10FBA9" box="[230,347,1022,1048]" captionStart="FIGURE 3" captionStartId="4.[136,229,1913,1935]" captionTargetBox="[357,1231,749,1889]" captionTargetId="figure-243@4.[357,1231,749,1889]" captionTargetPageId="4" captionText="FIGURE 3. A. Habitat of Valeriana plateadensis at the summit of the Cerro Plateado Biological Reserve. Photograph by Álvaro J. Pérez. B. Habitat of Valeriana yacuriensis at the Lagunas Negras in the Páramo de Jimbura, Yacuri National Park. Photograph by Petr Sklenář. C. Habitat of Valeriana xenophylloides in the Páramo de Patococha. Photograph by Petr Sklenář." figureDoi="http://doi.org/10.5281/zenodo.7532117" httpUri="https://zenodo.org/record/7532117/files/figure.png" pageId="5" pageNumber="52">Figure 3C</figureCitation>
). Only a few individual cushions were observed but more plants can be expected to occur at higher elevations.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD3FF8DFF74C84D7909FB35" pageId="5" pageNumber="52" type="conservation">
<paragraph id="3E1D5551FFD3FF8DFF74C84D7909FB35" blockId="5.[136,1452,302,1300]" pageId="5" pageNumber="52">
<emphasis id="0CD68943FFD3FF8DFF74C84D7DFFFBD1" bold="true" box="[189,436,1093,1120]" pageId="5" pageNumber="52">Conservation status:</emphasis>
Data deficient (DD); as only a few individuals have been observed from one locality more field work is needed to assess the conservation status according to the IUCN criteria properly (
<bibRefCitation id="5A3328A0FFD3FF8DFB65C862797DFB35" author="IUCN" box="[1196,1334,1130,1156]" pageId="5" pageNumber="52" refId="ref4312" refString="IUCN (2022) Guidelines for using the IUCN Red List categories and criteria. Version 15.1. Prepared by the Standards and Petitions Committee. Available from: https: // www. iucnredlist. org / resources / redlistguidelines (accessed 12 December 2022)" type="url" year="2022">IUCN 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD3FF8DFF74C8867EA5FB19" box="[189,750,1166,1192]" pageId="5" pageNumber="52" type="biology_ecology">
<paragraph id="3E1D5551FFD3FF8DFF74C8867EA5FB19" blockId="5.[136,1452,302,1300]" box="[189,750,1166,1192]" pageId="5" pageNumber="52">
<emphasis id="0CD68943FFD3FF8DFF74C8867D08FB19" bold="true" box="[189,323,1166,1192]" pageId="5" pageNumber="52">Phenology:</emphasis>
Flowers were collected in November.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD3FF8DFF74C8BA7F4BFAA5" pageId="5" pageNumber="52" type="etymology">
<paragraph id="3E1D5551FFD3FF8DFF74C8BA7F4BFAA5" blockId="5.[136,1452,302,1300]" pageId="5" pageNumber="52">
<emphasis id="0CD68943FFD3FF8DFF74C8BA7D03FB7D" bold="true" box="[189,328,1202,1228]" pageId="5" pageNumber="52">Etymology:</emphasis>
The habit of
<taxonomicName id="F9A22ED2FFD3FF8DFE23C8BA7F43FB7D" authorityName="Sklenar &amp; B. Eriksen" authorityYear="2023" box="[490,776,1202,1228]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="species" species="xenophylloides">
<emphasis id="0CD68943FFD3FF8DFE23C8BA7F43FB7D" box="[490,776,1202,1228]" italics="true" pageId="5" pageNumber="52">Valeriana xenophylloides</emphasis>
</taxonomicName>
resembles
<taxonomicName id="F9A22ED2FFD3FF8DFC45C8BA7923FB7C" authority="(Kunth) (1820: 150)" baseAuthorityName="Kunth" baseAuthorityPageNumber="150" baseAuthorityYear="1820" box="[908,1384,1202,1229]" class="Magnoliopsida" family="Asteraceae" genus="Xenophyllum" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="species" species="humile">
<emphasis id="0CD68943FFD3FF8DFC45C8BA783EFB7D" box="[908,1141,1202,1228]" italics="true" pageId="5" pageNumber="52">Xenophyllum humile</emphasis>
(Kunth) (1820: 150)
</taxonomicName>
Funk (1997: 239), a distinct cushion-forming
<taxonomicName id="F9A22ED2FFD3FF8DFD80C8DE7E8FFB41" box="[585,708,1238,1264]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="52" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
species which is commonly found in the paramos of
<collectingCountry id="46B515C1FFD3FF8DFAD7C8DE7930FB41" box="[1310,1403,1238,1264]" name="Ecuador" pageId="5" pageNumber="52">Ecuador</collectingCountry>
and the epithet of the new species refers to this resemblance.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="1D1D58456DEDB5DA83B893051B372F80">Three new species of Valeriana (Valerianoideae, Caprifoliaceae) from southern Ecuador</mods:title>
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<mods:namePart id="594C2ECD67DC5F3CE5691F24D35D599E">Persson, Claes</mods:namePart>
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<mods:affiliation id="A6D329F2EFDA2750A80B24ACB0FA23C5">Department of Biological and Environmental Sciences, University of Gothenburg, P. O. Box 461, 40530 Göteborg, Sweden. &amp; claes. persson @ bioenv. gu. se; https: // orcid. org / 0000 - 0003 - 3499 - 3373</mods:affiliation>
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</mods:role>
<mods:namePart id="C47E1D37562214606C5466082783B1E8">Eriksen, Bente</mods:namePart>
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<mods:affiliation id="3FC9D6FDE72E9DA4EA52BDD7F33E7171">Botanical Garden, Lund University, Östra Vallgatan 20, 223 61 Lund, Sweden. &amp; bente. eriksen @ botan. lu. se; https: // orcid. org / 0000 - 0002 - 6797 - 5262</mods:affiliation>
<mods:nameIdentifier id="E9EFE85F6329F01195F0DC66D5E531FA" type="email">bente.eriksen@botan.lu.se</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="60B0FEA4BB0263B17439C66C3C747327">Pérez, Álvaro J.</mods:namePart>
<mods:nameIdentifier id="66E2F87BD4885CCBAB416409C05BBDBC" type="ORCID">0000-0002-0644-9111</mods:nameIdentifier>
<mods:affiliation id="8DF079BAAFE0E886FA27D7E932021391">Herbario QCA, Escuela de Ciencias Biológicas, Pontificia Universidad Católica del Ecuador, Apartado 17 - 01 - 2184, Quito, Ecuador. &amp; alvaro. perez. castaneda @ gmail. com; https: // orcid. org / 0000 - 0002 - 0644 - 9111</mods:affiliation>
<mods:nameIdentifier id="A064D3A763D578660BD988EAE3A49680" type="email">alvaro.perez.castaneda@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="F8DA7AAC849FC2C24B69D1E1E337DDF2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB60CB33127919A523C93B74116EE537">Zapata, J. Nicolás</mods:namePart>
<mods:nameIdentifier id="E8AFC82FE3827DF31DE18F4C70E2DB08" type="ORCID">0000-0002-2863-6524</mods:nameIdentifier>
<mods:affiliation id="84B971F3057BA986D4C7A7892D6FFAAB">Herbario QCA, Escuela de Ciencias Biológicas, Pontificia Universidad Católica del Ecuador, Apartado 17 - 01 - 2184, Quito, Ecuador. &amp; DIADE, Univ Montpellier, CIRAD, IRD, Montpellier, France. &amp; josenicozb @ gmail. com; https: // orcid. org / 0000 - 0002 - 2863 - 6524</mods:affiliation>
<mods:nameIdentifier id="4D68EE93FE3AA3EB46061472A6430CB5" type="email">josenicozb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="07EB97AF2F529A84ABD282A56910B055" type="personal">
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<mods:roleTerm id="F2AEB16B8B84D3FC42B7996461BBAE4F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D7B153CF3CED74B883CD516F0DE0621C">Couvreur, Thomas L. P.</mods:namePart>
<mods:nameIdentifier id="F17FCDAE15C51761EEA8472F676A826A" type="ORCID">0000-0002-8509-6587</mods:nameIdentifier>
<mods:affiliation id="BFBF48B17993DFA3B5161F9447FCD9FE">Herbario QCA, Escuela de Ciencias Biológicas, Pontificia Universidad Católica del Ecuador, Apartado 17 - 01 - 2184, Quito, Ecuador. &amp; DIADE, Univ Montpellier, CIRAD, IRD, Montpellier, France. &amp; thomas. couvreur @ ird. fr; https: // orcid. org / 0000 - 0002 - 8509 - 6587</mods:affiliation>
<mods:nameIdentifier id="2114302242395D8D3D40868AA9EC26B6" type="email">thomas.couvreur@ird.fr</mods:nameIdentifier>
</mods:name>
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<mods:role id="EE40C691DCCDA754C21B1DDC0D237618">
<mods:roleTerm id="C494F0439913D46CA972BD46C7208B68">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="208678DCD9130519477E09D331F5A392">Sklenář, Petr</mods:namePart>
<mods:nameIdentifier id="D4AC49A4754A78032C45D8435BFC627D" type="ORCID">0000-0003-0429-2621</mods:nameIdentifier>
<mods:affiliation id="13B9A759CF6BFD8F04844B975B699E54">Department of Botany, Faculty of Science, Charles University in Prague, Benátská 2, CZ- 128 01 Prague, Czech Republic. &amp; petr. sklenar @ natur. cuni. cz; https: // orcid. org / 0000 - 0003 - 0429 - 2621</mods:affiliation>
<mods:nameIdentifier id="C51C3F0BFAB1958C88A085054DA6006C" type="email">petr.sklenar@natur.cuni.cz</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="8D9E4A7128EF771C3B15C85546280C7B">text</mods:typeOfResource>
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<mods:titleInfo id="94794CA7000A30914B6CBF03EC2BF296">
<mods:title id="BA66319763651157E42502DFF1B41A80">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="CE67ABD8B632A5E643BB4DDB19A3B2E5">
<mods:date id="EBE8DDC54B55FF7EFB0A210DF74338B0">2023</mods:date>
<mods:detail id="B910D59DF740D39F8C7446AE7F9B6C97" type="pubDate">
<mods:number id="631D86BA358C19E8C5320DF894B99C28">2023-01-13</mods:number>
</mods:detail>
<mods:detail id="201B9958421C2855FE9BD703F278426D" type="volume">
<mods:number id="96CAFF2F613BB218CBA2C30DBBAB102B">579</mods:number>
</mods:detail>
<mods:detail id="28AA1E7F7500C01E9CEB4BAB3916A489" type="issue">
<mods:number id="D8AE5389E98A08AA0ADD1D62CA92F017">1</mods:number>
</mods:detail>
<mods:extent id="E1B3A4C8C1843185EBF481D884F39B14" unit="page">
<mods:start id="27461BC2580C3412E8823ED5729435FF">47</mods:start>
<mods:end id="54A9051413BC2D8FECCC143F4609DF21">53</mods:end>
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<mods:location id="D3E14295C63A0CAB547B350418793C76">
<mods:url id="AFD15BA6A3144FCECBF780D360CF2B12">http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification id="17FE779F9E17887D4D419740967AC3A9">journal article</mods:classification>
<mods:identifier id="6F1CF80CA1FE5C775EF953E19EE8FD92" type="CLB-Dataset">226695</mods:identifier>
<mods:identifier id="3ED69E27261F6A72DE743FEBAC344954" type="DOI">10.11646/phytotaxa.579.1.5</mods:identifier>
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<mods:identifier id="AC6B581D37139FEBE87A5A29A9C77C16" type="Zenodo-Dep">7532106</mods:identifier>
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<treatment id="B60BE447FFD4FF8CFF41CA77790FFD65" ID-DOI="http://doi.org/10.5281/zenodo.7544120" ID-GBIF-Taxon="208979581" ID-Zenodo-Dep="7544120" LSID="urn:lsid:plazi:treatment:B60BE447FFD4FF8CFF41CA77790FFD65" httpUri="http://treatment.plazi.org/id/B60BE447FFD4FF8CFF41CA77790FFD65" lastPageId="4" lastPageNumber="51" pageId="2" pageNumber="49">
<subSubSection id="76B806DAFFD4FF8AFF41CA777851F928" box="[136,1050,1662,1689]" pageId="2" pageNumber="49" type="nomenclature">
<paragraph id="3E1D5551FFD4FF8AFF41CA777851F928" blockId="2.[136,1452,1662,1796]" box="[136,1050,1662,1689]" pageId="2" pageNumber="49">
<heading id="6555E23DFFD4FF8AFF41CA777851F928" bold="true" box="[136,1050,1662,1689]" fontSize="11" level="1" pageId="2" pageNumber="49" reason="1">
<emphasis id="0CD68943FFD4FF8AFF41CA777851F928" bold="true" box="[136,1050,1662,1689]" pageId="2" pageNumber="49">
2.
<taxonomicName id="F9A22ED2FFD4FF8AFF6DCA767EEBF929" ID-CoL="B6C65" authority="Sklenar &amp; B.Eriksen" authorityName="Sklenar &amp; B. Eriksen" authorityYear="2023" box="[164,672,1662,1689]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="2" pageNumber="49" phylum="Tracheophyta" rank="species" species="yacuriensis" status="sp. nov.">
<emphasis id="0CD68943FFD4FF8AFF6DCA767DD5F928" bold="true" box="[164,414,1662,1689]" italics="true" pageId="2" pageNumber="49">Valeriana yacuriensis</emphasis>
Sklenář &amp; B.Eriksen
</taxonomicName>
,
<emphasis id="0CD68943FFD4FF8AFD65CA777F48F928" bold="true" box="[684,771,1663,1689]" italics="true" pageId="2" pageNumber="49">
<taxonomicNameLabel id="17E53438FFD4FF8AFD65CA777F48F928" box="[684,771,1663,1689]" pageId="2" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
—
<figureCitation id="A69949D4FFD4FF8AFCD6CA767F87F928" box="[799,972,1662,1689]" captionStart="FIGURE 1" captionStartId="2.[136,229,1558,1580]" captionTargetBox="[188,1402,203,1523]" captionTargetId="figure-21@2.[177,1410,190,1534]" captionTargetPageId="2" captionText="FIGURE 1. A–C. Valeriana plateadensis. D–E. V. yacuriensis. F–G. V. xenophylloides. A, D, F. Habit. B. Close up of branches. C, E, G. Close up of inflorescence, showing 3-merous flowers. Photographs by Álvaro J. Pérez (A, C), Claes Persson (B), Petr Sklenář (D–G)." figureDoi="http://doi.org/10.5281/zenodo.7532110" httpUri="https://zenodo.org/record/7532110/files/figure.png" pageId="2" pageNumber="49">Figures 1 D–E</figureCitation>
,
<figureCitation id="A69949D4FFD4FF8AFC11CA777851F928" box="[984,1050,1663,1689]" captionStart="FIGURE 2" captionStartId="3.[136,229,1880,1902]" captionTargetBox="[188,1378,298,1834]" captionTargetId="figure-67@3.[183,1404,281,1856]" captionTargetPageId="3" captionText="FIGURE 2. A–E. Valeriana plateadensis. A. Habit. B. Side view of shoot (left), top view with flowers (right). C. Leaves in top view (left) and side view (right). D. Flower and bracts. E. Longitudinal section of ovary with hanging ovule (left) and of flower (right). F–J. V. yacuriensis. F. Habit. G. Side view of shoot (up), top view with flowers (down). H. Leaves in top view. I. Flower and bracts (left), front view of flower (right). J. Longitudinal section of flower. K–O. V. xenophylloides. K. Habit. L. Side view of shoot. M. Leaves in top view (left), side view (right) and the crown of trichomes at the leaf apex (up left). N. Flower and bracts (left), front view of flower (right). O. Longitudinal section of flower. Line drawings by Bobbi Angell." figureDoi="http://doi.org/10.5281/zenodo.7532114" httpUri="https://zenodo.org/record/7532114/files/figure.png" pageId="2" pageNumber="49">2 F–J</figureCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD4FF8AFF41CAAF7D5FF8B5" pageId="2" pageNumber="49" type="materials_examined">
<paragraph id="3E1D5551FFD4FF8AFF41CAAF7D5FF8B5" blockId="2.[136,1452,1662,1796]" pageId="2" pageNumber="49">
<materialsCitation id="8ECA5F0CFFD4FF8AFF41CAAF7D5AF8B5" ID-GBIF-Occurrence="4015641306" collectingDate="2009-06-16" collectionCode="P, J" collectorName="de Sabanilla &amp; Parque Nacional Yacuri" country="Ecuador" elevation="3500" latitude="-4.71" location="Loja" longLatPrecision="21" longitude="-79.42889" pageId="2" pageNumber="49" specimenCount="1" stateProvince="Loja" typeStatus="holotype">
Type:
<collectingCountry id="46B515C1FFD4FF8AFF0CCAAF7D77F90D" box="[197,316,1703,1724]" name="Ecuador" pageId="2" pageNumber="49">ECUADOR</collectingCountry>
.
<collectingRegion id="FC669BB3FFD4FF8AFE8CCAAE7D3EF90D" box="[325,373,1702,1724]" country="Ecuador" name="Loja" pageId="2" pageNumber="49">Loja</collectingRegion>
: Cordillera las Lagunillas (
<collectorName id="93573087FFD4FF8AFDB1CAAE7EB8F90D" box="[632,755,1702,1724]" pageId="2" pageNumber="49">de Sabanilla</collectorName>
), paramo de las Lagunas Negras [
<collectorName id="93573087FFD4FF8AFBFECAAF7969F90A" box="[1079,1314,1702,1724]" pageId="2" pageNumber="49">Parque Nacional Yacurí</collectorName>
],
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,
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,
<quantity id="F95AF8B4FFD4FF8AFE9BCAC27DD6F951" box="[338,413,1738,1760]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="49" unit="m" value="3500.0">
<elevation id="B58FB262FFD4FF8AFE9BCAC27DD6F951" box="[338,413,1738,1760]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="49" unit="m" value="3500.0">3500 m</elevation>
</quantity>
,
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<collectingDate id="5A588A79FFD4FF8AFE6ECAC27E6CF951" box="[423,551,1738,1760]" pageId="2" pageNumber="49" value="2009-06-16">16 June 2009</collectingDate>
</date>
(fl),
<emphasis id="0CD68943FFD4FF8AFD91CAC37FBBF951" box="[600,1008,1738,1760]" italics="true" pageId="2" pageNumber="49">
<collectionCode id="58B3CD94FFD4FF8AFD91CAC37E2CF951" box="[600,615,1739,1760]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="49" type="Herbarium">P</collectionCode>
. Sklenar,
<collectionCode id="58B3CD94FFD4FF8AFD0ACAC37E9AF951" box="[707,721,1739,1760]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="2" pageNumber="49" type="Herbarium">J</collectionCode>
. Mackova, &amp;
<collectionCode id="58B3CD94FFD4FF8AFC9FCAC37F2EF951" box="[854,869,1739,1760]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="49" type="Herbarium">P</collectionCode>
. Macek 12053
</emphasis>
(
<typeStatus id="E119EBF3FFD4FF8AFC34CAC27818F951" box="[1021,1107,1738,1760]" pageId="2" pageNumber="49" type="holotype">holotype</typeStatus>
PRC; isotypes GB-0210434, QCA-188696)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD4FF8BFF41CB3B7F41FEB1" lastPageId="3" lastPageNumber="50" pageId="2" pageNumber="49" type="description">
<paragraph id="3E1D5551FFD4FF8BFF41CB3B7F41FEB1" blockId="2.[136,1452,1843,2085]" lastBlockId="3.[136,1451,158,257]" lastPageId="3" lastPageNumber="50" pageId="2" pageNumber="49">
Shrubs to
<quantity id="F95AF8B4FFD4FF8AFECBCB3B7D00F8FC" box="[258,331,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="49" unit="cm" value="40.0">40 cm</quantity>
tall, branched stems ascending to erect, terete, to
<quantity id="F95AF8B4FFD4FF8AFC56CB3B7FB2F8FC" box="[927,1017,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="49" unit="mm" value="4.1">4.1 mm</quantity>
in diameter. Leaves numerous, wellspaced basally but becoming densely clustered in the upper part of the branches; the petiole persistent, triangular, flat, sheathing,
<quantity id="F95AF8B4FFD4FF8AFECBCB737DC2F824" box="[258,393,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="2" pageNumber="49" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.6–1.7 mm</quantity>
broad basally, ca
<quantity id="F95AF8B4FFD4FF8AFD9DCB737EE0F824" box="[596,683,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="49" unit="mm" value="2.2">2.2 mm</quantity>
long, green distally, whitish-brown with violet streaks basally when young, sparsely ciliate along the margins; the lamina spreading, glossy-green, tubercled, glabrous, flat to shallowly canaliculate towards the petiole, grooved adaxially, ca
<quantity id="F95AF8B4FFD4FF8AFD31CBCB7F1BF86C" box="[760,848,1987,2013]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="49" unit="mm" value="0.3">0.3 mm</quantity>
thick, oblong to obovate, 4.6–6.3 ×
<quantity id="F95AF8B4FFD4FF8AFB38CBCB7937F86C" box="[1265,1404,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="2" pageNumber="49" unit="mm" value="1.9" valueMax="2.1" valueMin="1.7">1.7–2.1 mm</quantity>
, the apex obtuse, the margin entire, rarely with a few cilia basally. Inflorescence capitoid, 6–20-flowered, sessile, the bracts leaf-like, ovate, 0.8–1.6 x
<quantity id="F95AF8B4FFD4FF8AFE63C4037E79F794" box="[426,562,2059,2085]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.8" metricValueMin="2.8" pageId="2" pageNumber="49" unit="mm" value="3.3" valueMax="3.8" valueMin="2.8">2.8–3.8 mm</quantity>
, acute. Flowers gynodioecious; calyx vestigial; corolla white, 3-lobed, narrowly funnelform, the tube to
<quantity id="F95AF8B4FFD5FF8BFE50CC967DBBFF08" box="[409,496,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="50" unit="mm" value="4.2">4.2 mm</quantity>
long, the lobes triangular, 1.3–1.7 ×
<quantity id="F95AF8B4FFD5FF8BFC53CC96786EFF08" box="[922,1061,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="3" pageNumber="50" unit="mm" value="1.15" valueMax="1.4" valueMin="0.9">0.9–1.4 mm</quantity>
, spreading and recurved, the apex obtuse; stamens exserted to
<quantity id="F95AF8B4FFD5FF8BFE77CCCA7E5DFF6C" box="[446,534,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="50" unit="mm" value="2.1">2.1 mm</quantity>
, spreading, the thecae
<quantity id="F95AF8B4FFD5FF8BFCD9CCCA7FF9FF6C" box="[784,946,194,221]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.6" metricValueMin="3.5" pageId="3" pageNumber="50" unit="mm" value="0.405" valueMax="0.46" valueMin="0.35">0.35–0.46 mm</quantity>
long; pistil with the style exserted to
<quantity id="F95AF8B4FFD5FF8BFA99CCCA79ECFF6C" box="[1360,1447,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="50" unit="mm" value="2.7">2.7 mm</quantity>
, 3-parted, the branches ca
<quantity id="F95AF8B4FFD5FF8BFE6ECCEE7E40FEB0" box="[423,523,230,257]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="50" unit="mm" value="0.15">0.15 mm</quantity>
long. Fruits unknown.
</paragraph>
</subSubSection>
<caption id="6ADD05D9FFD5FF8BFF41CB507EA6F792" ID-DOI="http://doi.org/10.5281/zenodo.7532114" ID-Zenodo-Dep="7532114" httpUri="https://zenodo.org/record/7532114/files/figure.png" pageId="3" pageNumber="50" startId="3.[136,229,1880,1902]" targetBox="[188,1378,298,1834]" targetPageId="3">
<paragraph id="3E1D5551FFD5FF8BFF41CB507EA6F792" blockId="3.[136,1453,1880,2083]" pageId="3" pageNumber="50">
<emphasis id="0CD68943FFD5FF8BFF41CB507CB4F8DC" bold="true" box="[136,255,1880,1902]" pageId="3" pageNumber="50">FIGURE 2.</emphasis>
A–E.
<taxonomicName id="F9A22ED2FFD5FF8BFE89CB517E56F8DF" authorityName="A. J. Perez, C. Persson &amp; J. N. Zapata" authorityYear="2023" box="[320,541,1881,1902]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="3" pageNumber="50" phylum="Tracheophyta" rank="species" species="plateadensis">
<emphasis id="0CD68943FFD5FF8BFE89CB517E56F8DF" box="[320,541,1881,1902]" italics="true" pageId="3" pageNumber="50">Valeriana plateadensis</emphasis>
</taxonomicName>
. A. Habit. B. Side view of shoot (left), top view with flowers (right). C. Leaves in top view (left) and side view (right). D. Flower and bracts. E. Longitudinal section of ovary with hanging ovule (left) and of flower (right). F–J.
<taxonomicName id="F9A22ED2FFD5FF8BFA53CB757CBEF807" authorityName="Sklenar &amp; B. Eriksen" authorityYear="2023" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="3" pageNumber="50" phylum="Tracheophyta" rank="species" species="yacuriensis">
<emphasis id="0CD68943FFD5FF8BFA53CB757CBEF807" italics="true" pageId="3" pageNumber="50">V. yacuriensis</emphasis>
</taxonomicName>
. F. Habit. G. Side view of shoot (up), top view with flowers (down). H. Leaves in top view. I. Flower and bracts (left), front view of flower (right). J. Longitudinal section of flower. K–O.
<taxonomicName id="F9A22ED2FFD5FF8BFD2FCBCD7FC6F86B" authorityName="Sklenar &amp; B. Eriksen" authorityYear="2023" box="[742,909,1989,2010]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="3" pageNumber="50" phylum="Tracheophyta" rank="species" species="xenophylloides">
<emphasis id="0CD68943FFD5FF8BFD2FCBCD7FC6F86B" box="[742,909,1989,2010]" italics="true" pageId="3" pageNumber="50">V. xenophylloides</emphasis>
</taxonomicName>
. K. Habit. L. Side view of shoot. M. Leaves in top view (left), side view (right) and the crown of trichomes at the leaf apex (up left). N. Flower and bracts (left), front view of flower (right). O. Longitudinal section of flower. Line drawings by Bobbi Angell.
</paragraph>
</caption>
<subSubSection id="76B806DAFFD2FF8CFF74CC967D7AFEB1" pageId="4" pageNumber="51" type="materials_examined">
<paragraph id="3E1D5551FFD2FF8CFF74CC967D7AFEB1" blockId="4.[136,1451,158,725]" pageId="4" pageNumber="51">
<emphasis id="0CD68943FFD2FF8CFF74CC967E09FF09" bold="true" box="[189,578,158,184]" pageId="4" pageNumber="51">Additional specimens examined:</emphasis>
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<collectingCountry id="46B515C1FFD2FF8CFD83CC967E9EFF09" box="[586,725,158,184]" name="Ecuador" pageId="4" pageNumber="51">ECUADOR</collectingCountry>
.
<collectingRegion id="FC669BB3FFD2FF8CFD29CC967F53FF09" box="[736,792,158,184]" country="Ecuador" name="Loja" pageId="4" pageNumber="51">Loja</collectingRegion>
:
<collectingMunicipality id="DE79CF2BFFD2FF8CFCEACC97787AFF09" box="[803,1073,158,185]" pageId="4" pageNumber="51">Parque Nacional Yacurí</collectingMunicipality>
, around the
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,
<geoCoordinate id="5B963396FFD2FF8CFF41CCCA7D51FF6C" box="[136,282,194,221]" degrees="79" direction="west" minutes="25" orientation="longitude" pageId="4" pageNumber="51" precision="15" seconds="27" value="-79.424164">W79°25’27”</geoCoordinate>
,
<geoCoordinate id="5B963396FFD2FF8CFEEECCCA7DE5FF6C" box="[295,430,194,221]" degrees="04" direction="south" minutes="42" orientation="latitude" pageId="4" pageNumber="51" precision="15" seconds="51" value="-4.7141666">S04°42’51”</geoCoordinate>
,
<quantity id="F95AF8B4FFD2FF8CFE75CCCA7E5EFF6C" box="[444,533,194,221]" metricMagnitude="3" metricUnit="m" metricValue="3.325" pageId="4" pageNumber="51" unit="m" value="3325.0">
<elevation id="B58FB262FFD2FF8CFE75CCCA7E5EFF6C" box="[444,533,194,221]" metricMagnitude="3" metricUnit="m" metricValue="3.325" pageId="4" pageNumber="51" unit="m" value="3325.0">3325 m</elevation>
</quantity>
,
<date id="4A1C7391FFD2FF8CFDEACCCB7EBFFF6D" box="[547,756,194,221]" pageId="4" pageNumber="51" value="2018-11-07">
<collectingDate id="5A588A79FFD2FF8CFDEACCCB7EBFFF6D" box="[547,756,194,221]" pageId="4" pageNumber="51" value="2018-11-07">7 November 2018</collectingDate>
</date>
,
<emphasis id="0CD68943FFD2FF8CFCC8CCCA7916FF6D" box="[769,1373,194,221]" italics="true" pageId="4" pageNumber="51">
Sklenar
<collectionCode id="58B3CD94FFD2FF8CFCA9CCCB7F38FF6D" box="[864,883,195,220]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="51" type="Herbarium">P</collectionCode>
., Ptacek
<collectionCode id="58B3CD94FFD2FF8CFC10CCCB7FA0FF6D" box="[985,1003,195,220]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="51" type="Herbarium">J</collectionCode>
., Klimesova
<collectionCode id="58B3CD94FFD2FF8CFBB5CCCB78C5FF6D" box="[1148,1166,195,220]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="51" type="Herbarium">J</collectionCode>
., Klimes
<collectionCode id="58B3CD94FFD2FF8CFB3FCCCB7941FF6C" box="[1270,1290,195,221]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="51" type="Herbarium">A</collectionCode>
. 15684
</emphasis>
</materialsCitation>
(
<collectingCountry id="46B515C1FFD2FF8CFAA7CCCA79ECFF6D" box="[1390,1447,194,220]" name="China" pageId="4" pageNumber="51">PRC</collectingCountry>
, QCA-241671).
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD2FF8CFF74CD027F29FE21" pageId="4" pageNumber="51" type="diagnosis">
<paragraph id="3E1D5551FFD2FF8CFF74CD027F29FE21" blockId="4.[136,1451,158,725]" pageId="4" pageNumber="51">
<taxonomicName id="F9A22ED2FFD2FF8CFF74CD027DF9FE95" authorityName="Sklenar &amp; B. Eriksen" authorityYear="2023" box="[189,434,266,292]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="yacuriensis">
<emphasis id="0CD68943FFD2FF8CFF74CD027DF9FE95" box="[189,434,266,292]" italics="true" pageId="4" pageNumber="51">Valeriana yacuriensis</emphasis>
</taxonomicName>
is distinguished from the other 3-lobed species of
<taxonomicName id="F9A22ED2FFD2FF8CFC25CD027812FE95" box="[1004,1113,266,292]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="genus">
<emphasis id="0CD68943FFD2FF8CFC25CD027812FE95" box="[1004,1113,266,292]" italics="true" pageId="4" pageNumber="51">Valeriana</emphasis>
</taxonomicName>
in
<collectingCountry id="46B515C1FFD2FF8CFBB4CD027891FE95" box="[1149,1242,266,292]" name="Ecuador" pageId="4" pageNumber="51">Ecuador</collectingCountry>
and northern
<collectingCountry id="46B515C1FFD2FF8CFAB1CD0379E0FE94" box="[1400,1451,267,293]" name="Peru" pageId="4" pageNumber="51">Peru</collectingCountry>
by the shrubby habit and petiolate leaves which are distinctly clustered apically.
<taxonomicName id="F9A22ED2FFD2FF8CFC34CD2679E0FEF8" authority="Killip (1928: 501)" authorityName="Killip" authorityPageNumber="501" authorityYear="1928" box="[1021,1451,302,329]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="imbricata">
<emphasis id="0CD68943FFD2FF8CFC34CD267896FEF9" box="[1021,1245,302,328]" italics="true" pageId="4" pageNumber="51">Valeriana imbricata</emphasis>
<bibRefCitation id="5A3328A0FFD2FF8CFB2BCD2679E0FEF8" author="Killip, E. P." box="[1250,1451,302,329]" pageId="4" pageNumber="51" pagination="498 - 501" refId="ref4359" refString="Killip, E. P. (1928) Seven new species of Valeriana. Journal of the Washington Academy of Sciences 18: 498 - 501." type="journal article" year="1928">Killip (1928: 501)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="F9A22ED2FFD2FF8CFF7ECD5B7D16FEDD" authorityName="A. J. Perez, C. Persson &amp; J. N. Zapata" authorityYear="2023" box="[183,349,338,364]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="plateadensis">
<emphasis id="0CD68943FFD2FF8CFF7ECD5B7D16FEDD" box="[183,349,338,364]" italics="true" pageId="4" pageNumber="51">V. plateadensis</emphasis>
</taxonomicName>
are shrubs but have (sub)sessile, closely imbricate leaves which are persistent throughout the stem, whereas
<taxonomicName id="F9A22ED2FFD2FF8CFF22CD7F7DE5FE21" authorityName="Sklenar &amp; B. Eriksen" authorityYear="2023" box="[235,430,374,400]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="xenophylloides">
<emphasis id="0CD68943FFD2FF8CFF22CD7F7DE5FE21" box="[235,430,374,400]" italics="true" pageId="4" pageNumber="51">V. xenophylloides</emphasis>
</taxonomicName>
(described below) is cushion-forming.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD2FF8CFF74CD927960FE4D" pageId="4" pageNumber="51" type="distribution">
<paragraph id="3E1D5551FFD2FF8CFF74CD927960FE4D" blockId="4.[136,1451,158,725]" pageId="4" pageNumber="51">
<emphasis id="0CD68943FFD2FF8CFF74CD927DAAFE05" bold="true" box="[189,481,410,436]" pageId="4" pageNumber="51">Distribution and habitat</emphasis>
:—
<taxonomicName id="F9A22ED2FFD2FF8CFDCDCD927EB3FE05" authorityName="Sklenar &amp; B. Eriksen" authorityYear="2023" box="[516,760,410,436]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="yacuriensis">
<emphasis id="0CD68943FFD2FF8CFDCDCD927EB3FE05" box="[516,760,410,436]" italics="true" pageId="4" pageNumber="51">Valeriana yacuriensis</emphasis>
</taxonomicName>
was collected twice at the
<typeStatus id="E119EBF3FFD2FF8CFBECCD93781EFE04" box="[1061,1109,411,437]" pageId="4" pageNumber="51">type</typeStatus>
locality, i.e., in humid paramo grasslands with scattered shrubs around the Laguna Negra in the Parque Nacional Yacurí (
<figureCitation id="A69949D4FFD2FF8CFB67CDB6796CFE69" box="[1198,1319,446,472]" captionStart="FIGURE 3" captionStartId="4.[136,229,1913,1935]" captionTargetBox="[357,1231,749,1889]" captionTargetId="figure-243@4.[357,1231,749,1889]" captionTargetPageId="4" captionText="FIGURE 3. A. Habitat of Valeriana plateadensis at the summit of the Cerro Plateado Biological Reserve. Photograph by Álvaro J. Pérez. B. Habitat of Valeriana yacuriensis at the Lagunas Negras in the Páramo de Jimbura, Yacuri National Park. Photograph by Petr Sklenář. C. Habitat of Valeriana xenophylloides in the Páramo de Patococha. Photograph by Petr Sklenář." figureDoi="http://doi.org/10.5281/zenodo.7532117" httpUri="https://zenodo.org/record/7532117/files/figure.png" pageId="4" pageNumber="51">Figure 3B</figureCitation>
). Since the locality occurs at the Ecuador-Peruvian border, the occurrence of the species in the latter country is likely.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD2FF8CFF74CE0E7DA9FDD9" pageId="4" pageNumber="51" type="conservation">
<paragraph id="3E1D5551FFD2FF8CFF74CE0E7DA9FDD9" blockId="4.[136,1451,158,725]" pageId="4" pageNumber="51">
<emphasis id="0CD68943FFD2FF8CFF74CE0E7DFFFD91" bold="true" box="[189,436,518,544]" pageId="4" pageNumber="51">Conservation status:</emphasis>
Data deficient (DD); as
<taxonomicName id="F9A22ED2FFD2FF8CFD00CE0E7FF6FD91" authorityName="Sklenar &amp; B. Eriksen" authorityYear="2023" box="[713,957,518,544]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="4" pageNumber="51" phylum="Tracheophyta" rank="species" species="yacuriensis">
<emphasis id="0CD68943FFD2FF8CFD00CE0E7FF6FD91" box="[713,957,518,544]" italics="true" pageId="4" pageNumber="51">Valeriana yacuriensis</emphasis>
</taxonomicName>
has only been collected twice from the
<typeStatus id="E119EBF3FFD2FF8CFAB2CE0F79E0FD90" box="[1403,1451,519,545]" pageId="4" pageNumber="51">type</typeStatus>
locality it is obvious that more field work is needed in order to assess the conservation status according to the IUCN criteria properly (
<bibRefCitation id="5A3328A0FFD2FF8CFE85CE467D9DFDD9" author="IUCN" box="[332,470,590,616]" pageId="4" pageNumber="51" refId="ref4312" refString="IUCN (2022) Guidelines for using the IUCN Red List categories and criteria. Version 15.1. Prepared by the Standards and Petitions Committee. Available from: https: // www. iucnredlist. org / resources / redlistguidelines (accessed 12 December 2022)" type="url" year="2022">IUCN 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD2FF8CFF74CE7A7F5FFD01" pageId="4" pageNumber="51" type="biology_ecology">
<paragraph id="3E1D5551FFD2FF8CFF74CE7A7F5FFD01" blockId="4.[136,1451,158,725]" pageId="4" pageNumber="51">
<emphasis id="0CD68943FFD2FF8CFF74CE7A7D08FD3D" bold="true" box="[189,323,626,652]" pageId="4" pageNumber="51">Phenology:</emphasis>
Flowers were observed in July and November which indicates that the population of the species has a very extended (perhaps continuous) period of flowering.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD2FF8CFF74CEB2790FFD65" box="[189,1348,698,725]" pageId="4" pageNumber="51" type="etymology">
<paragraph id="3E1D5551FFD2FF8CFF74CEB2790FFD65" blockId="4.[136,1451,158,725]" box="[189,1348,698,725]" pageId="4" pageNumber="51">
<emphasis id="0CD68943FFD2FF8CFF74CEB27D03FD65" bold="true" box="[189,328,698,724]" pageId="4" pageNumber="51">Etymology:</emphasis>
The epithet refers to the name of the Parque Nacional Yacurí where the species was found.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="B33D3F17F3BDEB81C7CE696314E4DBA9">Three new species of Valeriana (Valerianoideae, Caprifoliaceae) from southern Ecuador</mods:title>
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<mods:namePart id="3BD4B1A4240FF8A9AD1F9E594CEF1082">Persson, Claes</mods:namePart>
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<mods:namePart id="C8EBC7C74C44D065D25358F778038609">Eriksen, Bente</mods:namePart>
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<mods:affiliation id="5B7AD6906674463518D0951F477DF8F7">Botanical Garden, Lund University, Östra Vallgatan 20, 223 61 Lund, Sweden. &amp; bente. eriksen @ botan. lu. se; https: // orcid. org / 0000 - 0002 - 6797 - 5262</mods:affiliation>
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<mods:affiliation id="8D116E7AC447C169308D6C334C8751DA">Herbario QCA, Escuela de Ciencias Biológicas, Pontificia Universidad Católica del Ecuador, Apartado 17 - 01 - 2184, Quito, Ecuador. &amp; alvaro. perez. castaneda @ gmail. com; https: // orcid. org / 0000 - 0002 - 0644 - 9111</mods:affiliation>
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<mods:namePart id="5EE6BA118399A08CEE47BF22564530FB">Couvreur, Thomas L. P.</mods:namePart>
<mods:nameIdentifier id="3AF0F77DA3EA2834737794D7E2F0D9C4" type="ORCID">0000-0002-8509-6587</mods:nameIdentifier>
<mods:affiliation id="727958979E3C8915FE93AEEA2B6F673B">Herbario QCA, Escuela de Ciencias Biológicas, Pontificia Universidad Católica del Ecuador, Apartado 17 - 01 - 2184, Quito, Ecuador. &amp; DIADE, Univ Montpellier, CIRAD, IRD, Montpellier, France. &amp; thomas. couvreur @ ird. fr; https: // orcid. org / 0000 - 0002 - 8509 - 6587</mods:affiliation>
<mods:nameIdentifier id="8D5F8282E2A0C6F29DE62CD198F2A2C7" type="email">thomas.couvreur@ird.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="2EF79D77F6D95A29891778F20C5F5B59" type="personal">
<mods:role id="1868937F1926DD049B09B6C3F7725A2A">
<mods:roleTerm id="8AC008BA99DCF1119EDBFD59F62B416A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C115C3FF5D1D218455A451141BC3B1F">Sklenář, Petr</mods:namePart>
<mods:nameIdentifier id="F90509F42C0357A622954C5B8A5C1DC3" type="ORCID">0000-0003-0429-2621</mods:nameIdentifier>
<mods:affiliation id="BAC5B6F700ED994BA5677CDC41BF1360">Department of Botany, Faculty of Science, Charles University in Prague, Benátská 2, CZ- 128 01 Prague, Czech Republic. &amp; petr. sklenar @ natur. cuni. cz; https: // orcid. org / 0000 - 0003 - 0429 - 2621</mods:affiliation>
<mods:nameIdentifier id="EF180B71E27B0504952C07592423DF4D" type="email">petr.sklenar@natur.cuni.cz</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0F183DF4E79A7A6D43EA0236DA1B95B6">text</mods:typeOfResource>
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<mods:titleInfo id="BF32FA2B76ED003D6FC2946588C1ACC1">
<mods:title id="AF92403D0F30C514BD73EAB6A9DE456F">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="8EDF058EE056309F38D40741030942EA">
<mods:date id="1D3684CB55919EC1A21818EA798C6791">2023</mods:date>
<mods:detail id="313CBE43204773C020CD869B545D3493" type="pubDate">
<mods:number id="68C789BE341D44BA454EB0DD7DC6CB91">2023-01-13</mods:number>
</mods:detail>
<mods:detail id="1B1367A7A01FCF2CC2C9D0C82856616E" type="volume">
<mods:number id="511D88EEF27F37822C764755C79F0ECD">579</mods:number>
</mods:detail>
<mods:detail id="D6D156256C5E59DEF2E0552D296B1216" type="issue">
<mods:number id="4E1E77D0AB857379D8744090A05985F8">1</mods:number>
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<mods:extent id="8B2502E598170E43169263F19D1B26D0" unit="page">
<mods:start id="670FF5EB5492E3BE76F47D58C7BA0057">47</mods:start>
<mods:end id="A543A44BA6AA5F2BB71CDBB5A6A1C48A">53</mods:end>
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<mods:location id="7D0806A4CFDC827BA60FB4A7255B1BB6">
<mods:url id="D7CA284F14BF34B58EAD026AD92A7506">http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification id="5A8C135A426C6BA9AB50C6E9B604BE12">journal article</mods:classification>
<mods:identifier id="DF4BADA1598A362E91867633028BEF3E" type="CLB-Dataset">226695</mods:identifier>
<mods:identifier id="1946F2DB26DB6746BF2597C4BE0689F2" type="DOI">10.11646/phytotaxa.579.1.5</mods:identifier>
<mods:identifier id="BD7ED8ACF94A85DDB3F463625782FCF8" type="GBIF-Dataset">554b4408-0afc-4d63-b3e1-e27d29df9eb6</mods:identifier>
<mods:identifier id="AEE6F32AD5DEB0024DAF3B858998627C" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="5C0BA15E821753CE6E336C866A773293" type="Zenodo-Dep">7532106</mods:identifier>
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<treatment id="B60BE447FFD7FF89FF41CF257F5AF7AC" ID-DOI="http://doi.org/10.5281/zenodo.7532104" ID-GBIF-Taxon="208979580" ID-Zenodo-Dep="7532104" LSID="urn:lsid:plazi:treatment:B60BE447FFD7FF89FF41CF257F5AF7AC" httpUri="http://treatment.plazi.org/id/B60BE447FFD7FF89FF41CF257F5AF7AC" lastPageNumber="48" pageId="1" pageNumber="48">
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<paragraph id="3E1D5551FFD7FF89FF41CF2578A1FCF6" blockId="1.[136,1452,812,947]" box="[136,1258,812,843]" pageId="1" pageNumber="48">
<heading id="6555E23DFFD7FF89FF41CF2578A1FCF6" bold="true" box="[136,1258,812,843]" fontSize="11" level="1" pageId="1" pageNumber="48" reason="1">
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1.
<taxonomicName id="F9A22ED2FFD7FF89FF6DCF257F21FCF9" ID-CoL="B6C63" authority="A.J. Perez, C. Persson &amp; J.N. Zapata" authorityName="A. J. Perez, C. Persson &amp; J. N. Zapata" authorityYear="2023" box="[164,874,812,843]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="1" pageNumber="48" phylum="Tracheophyta" rank="species" species="plateadensis" status="sp. nov.">
<emphasis id="0CD68943FFD7FF89FF6DCF257DE2FCF9" bold="true" box="[164,425,813,840]" italics="true" pageId="1" pageNumber="48">Valeriana plateadensis</emphasis>
Á.J. Pérez, C. Persson &amp; J.N. Zapata
</taxonomicName>
,
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<taxonomicNameLabel id="17E53438FFD7FF89FCBECF267F86FCF9" box="[887,973,814,840]" pageId="1" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
—
<figureCitation id="A69949D4FFD7FF89FC23CF2578DDFCF6" box="[1002,1174,813,839]" captionStart="FIGURE 1" captionStartId="2.[136,229,1558,1580]" captionTargetBox="[188,1402,203,1523]" captionTargetId="figure-21@2.[177,1410,190,1534]" captionTargetPageId="2" captionText="FIGURE 1. A–C. Valeriana plateadensis. D–E. V. yacuriensis. F–G. V. xenophylloides. A, D, F. Habit. B. Close up of branches. C, E, G. Close up of inflorescence, showing 3-merous flowers. Photographs by Álvaro J. Pérez (A, C), Claes Persson (B), Petr Sklenář (D–G)." figureDoi="http://doi.org/10.5281/zenodo.7532110" httpUri="https://zenodo.org/record/7532110/files/figure.png" pageId="1" pageNumber="48">Figures 1 A–C</figureCitation>
,
<figureCitation id="A69949D4FFD7FF89FB6BCF2578A1FCF6" box="[1186,1258,813,839]" captionStart="FIGURE 2" captionStartId="3.[136,229,1880,1902]" captionTargetBox="[188,1378,298,1834]" captionTargetId="figure-67@3.[183,1404,281,1856]" captionTargetPageId="3" captionText="FIGURE 2. A–E. Valeriana plateadensis. A. Habit. B. Side view of shoot (left), top view with flowers (right). C. Leaves in top view (left) and side view (right). D. Flower and bracts. E. Longitudinal section of ovary with hanging ovule (left) and of flower (right). F–J. V. yacuriensis. F. Habit. G. Side view of shoot (up), top view with flowers (down). H. Leaves in top view. I. Flower and bracts (left), front view of flower (right). J. Longitudinal section of flower. K–O. V. xenophylloides. K. Habit. L. Side view of shoot. M. Leaves in top view (left), side view (right) and the crown of trichomes at the leaf apex (up left). N. Flower and bracts (left), front view of flower (right). O. Longitudinal section of flower. Line drawings by Bobbi Angell." figureDoi="http://doi.org/10.5281/zenodo.7532114" httpUri="https://zenodo.org/record/7532114/files/figure.png" pageId="1" pageNumber="48">2 A–E</figureCitation>
</emphasis>
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</paragraph>
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<subSubSection id="76B806DAFFD7FF89FF41CF5D7FB8FC03" pageId="1" pageNumber="48" type="materials_examined">
<paragraph id="3E1D5551FFD7FF89FF41CF5D7FB8FC03" blockId="1.[136,1452,812,947]" pageId="1" pageNumber="48">
<materialsCitation id="8ECA5F0CFFD7FF89FF41CF5D7FA4FC03" ID-GBIF-Occurrence="4015641302" collectingDate="2021-08-08" collectionCode="A, J, N, T, L, P, C, R, W" country="Ecuador" county="Perez" elevation="2900" latitude="-4.6194444" location="Zamora-Chinchipe" longLatPrecision="1" longitude="-78.78306" municipality="Zapata" pageId="1" pageNumber="48" specimenCount="1" stateProvince="Zamora-Chinchipe" typeStatus="holotype">
Type:
<collectingCountry id="46B515C1FFD7FF89FF0FCF5D7D76FCDB" box="[198,317,853,874]" name="Ecuador" pageId="1" pageNumber="48">ECUADOR</collectingCountry>
.
<collectingRegion id="FC669BB3FFD7FF89FE8ECF5D7E4BFCDA" box="[327,512,853,875]" country="Ecuador" name="Zamora-Chinchipe" pageId="1" pageNumber="48">Zamora-Chinchipe</collectingRegion>
: Cantón Nangaritza, Parroquia Nuevo Paraíso. Reserva Biológica Cerro Plateado, [la cima de la meseta],
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,
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,
<quantity id="F95AF8B4FFD7FF89FDD0CF717E2DFC3F" box="[537,614,889,910]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="48" unit="m" value="2900.0">
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</quantity>
,
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<collectingDate id="5A588A79FFD7FF89FDBBCF717F4AFC3F" box="[626,769,889,910]" pageId="1" pageNumber="48" value="2021-08-08">8 August 2021</collectingDate>
</date>
(fl),
<emphasis id="0CD68943FFD7FF89FCFCCF717DAAFC03" italics="true" pageId="1" pageNumber="48">
<collectionCode id="58B3CD94FFD7FF89FCFCCF717F0DFC3F" box="[821,838,889,910]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="48" type="Herbarium">A</collectionCode>
.
<collectionCode id="58B3CD94FFD7FF89FC83CF717F13FC3F" box="[842,856,889,910]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="1" pageNumber="48" type="Herbarium">J</collectionCode>
.
<collectingCounty id="D77C2DDDFFD7FF89FCABCF717FD1FC3F" box="[866,922,889,910]" pageId="1" pageNumber="48">Pérez</collectingCounty>
,
<collectionCode id="58B3CD94FFD7FF89FC6FCF727FF2FC3E" box="[934,953,890,911]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="48" type="Herbarium">N</collectionCode>
.
<collectingMunicipality id="DE79CF2BFFD7FF89FC0DCF727841FC3E" box="[964,1034,890,911]" pageId="1" pageNumber="48">Zapata</collectingMunicipality>
,
<collectionCode id="58B3CD94FFD7FF89FBDFCF727869FC3E" box="[1046,1058,890,911]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="48">T</collectionCode>
.
<collectionCode id="58B3CD94FFD7FF89FBEDCF72787FFC3E" box="[1060,1076,890,911]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="1" pageNumber="48" type="Herbarium">L</collectionCode>
.
<collectionCode id="58B3CD94FFD7FF89FBF1CF727802FC3E" box="[1080,1097,890,911]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="48" type="Herbarium">P</collectionCode>
.
<location id="3B7D038AFFD7FF89FB9DCF7178E4FC3F" LSID="urn:lsid:plazi:treatment:B60BE447FFD7FF89FF41CF257F5AF7AC:3B7D038AFFD7FF89FB9DCF7178E4FC3F" box="[1108,1199,889,910]" country="China" county="Perez" latitude="-4.6194444" longLatPrecision="1" longitude="-78.78306" municipality="Zapata" name="Couvreur" pageId="1" pageNumber="48">Couvreur</location>
,
<collectionCode id="58B3CD94FFD7FF89FB73CF717886FC3F" box="[1210,1229,889,910]" country="Denmark" name="University of Copenhagen" pageId="1" pageNumber="48" type="Herbarium">C</collectionCode>
.
<location id="3B7D038AFFD7FF89FB11CF727963FC3E" LSID="urn:lsid:plazi:treatment:B60BE447FFD7FF89FF41CF257F5AF7AC:3B7D038AFFD7FF89FB11CF727963FC3E" box="[1240,1320,890,911]" country="China" county="Perez" latitude="-4.6194444" longLatPrecision="1" longitude="-78.78306" municipality="Zapata" name="Persson" pageId="1" pageNumber="48">Persson</location>
,
<collectionCode id="58B3CD94FFD7FF89FAFACF72790FFC3E" box="[1331,1348,890,911]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="48">R</collectionCode>
.
<location id="3B7D038AFFD7FF89FA86CF7279C0FC3E" LSID="urn:lsid:plazi:treatment:B60BE447FFD7FF89FF41CF257F5AF7AC:3B7D038AFFD7FF89FA86CF7279C0FC3E" box="[1359,1419,890,911]" country="China" county="Perez" latitude="-4.6194444" longLatPrecision="1" longitude="-78.78306" municipality="Zapata" name="Paqui" pageId="1" pageNumber="48">Paqui</location>
,
<collectionCode id="58B3CD94FFD7FF89FA5ECF7279E3FC3E" box="[1431,1448,890,911]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="48">R</collectionCode>
.
<location id="3B7D038AFFD7FF89FF74CF957D46FC03" LSID="urn:lsid:plazi:treatment:B60BE447FFD7FF89FF41CF257F5AF7AC:3B7D038AFFD7FF89FF74CF957D46FC03" box="[189,269,925,946]" country="China" county="Perez" latitude="-4.6194444" longLatPrecision="1" longitude="-78.78306" municipality="Zapata" name="Cabrera" pageId="1" pageNumber="48">Cabrera</location>
&amp;
<collectionCode id="58B3CD94FFD7FF89FEE4CF967D09FC02" box="[301,322,926,947]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="48" type="Herbarium">W</collectionCode>
.
<location id="3B7D038AFFD7FF89FE83CF957DEAFC03" LSID="urn:lsid:plazi:treatment:B60BE447FFD7FF89FF41CF257F5AF7AC:3B7D038AFFD7FF89FE83CF957DEAFC03" box="[330,417,925,946]" country="China" county="Perez" latitude="-4.6194444" longLatPrecision="1" longitude="-78.78306" municipality="Zapata" name="Santillan" pageId="1" pageNumber="48">Santillán</location>
11704
</emphasis>
(
<typeStatus id="E119EBF3FFD7FF89FE27CF957E0FFC02" box="[494,580,925,947]" pageId="1" pageNumber="48" type="holotype">holotype</typeStatus>
QCA-244410; isotypes GB-0210405, PRC)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD7FF89FF41CFD278A0FA2F" pageId="1" pageNumber="48" type="description">
<paragraph id="3E1D5551FFD7FF89FF41CFD278A0FA2F" blockId="1.[136,1452,986,2077]" pageId="1" pageNumber="48">
Shrub
<quantity id="F95AF8B4FFD7FF89FF1DCFD27D0FFC44" box="[212,324,986,1013]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="48" unit="cm" value="22.5" valueMax="25.0" valueMin="20.0">20–25 cm</quantity>
tall with ± erect branches. Stem and branches terete covered with dead leaves basally,
<quantity id="F95AF8B4FFD7FF89FAE0CFD279E0FC44" box="[1321,1451,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="48" unit="cm" value="0.75" valueMax="1.0" valueMin="0.5">0.5 to 1 cm</quantity>
wide (incl. leaves), slightly wider at the head-like top, branching rhythmic with 3–7 branches at each node. Leaves seemingly spirally arranged (but actually positioned in multiple ranks) and overlapping, sessile, glabrous, coriaceous, spatulate and sharply bent, proximal portion canaliculate, 5 ×
<quantity id="F95AF8B4FFD7FF89FCE7C84D7F24FBEE" box="[814,879,1093,1119]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="48" unit="mm" value="1.0">1 mm</quantity>
, adpressed tightly to the shoot, distal portion widely oblong 2.1–2.3 ×
<quantity id="F95AF8B4FFD7FF89FE9EC8607D85FB32" box="[343,462,1128,1155]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="1" pageNumber="48" unit="mm" value="2.1" valueMax="2.2" valueMin="2.0">2–2.2 mm</quantity>
, with rounded apex, above slightly concave with thickened margin, below sometimes with a raised narrowly triangular portion, fresh leaves generally spreading (forming an angle of 90° to the shoot), or in the uppermost portion of the shoots ascending to erect, dried and dead leaves the outer portion generally facing downwards. Inflorescence a loose cluster of up to 12 flowers at the top/end of the shoots, partial inflorescences of 2–3 flowers subtended by an opposite pair of involucral bracts that are entirely or fused in for 2/3. Flowers hermaphrodite? (neither pollen nor ovules seen); calyx vestigial; corolla funnelform, white, 3-lobed, the tube obconical,
<quantity id="F95AF8B4FFD7FF89FAFEC91179E7FA85" box="[1335,1452,1305,1332]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="1" pageNumber="48" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 mm</quantity>
long,
<quantity id="F95AF8B4FFD7FF89FF0EC9357D2BFAE6" box="[199,352,1341,1368]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="1" pageNumber="48" unit="mm" value="0.85" valueMax="1.5" valueMin="0.2">0.2 to 1.5 mm</quantity>
wide, the lobes widely ovate, ca 1.5 ×
<quantity id="F95AF8B4FFD7FF89FCC5C9357F05FAE6" box="[780,846,1341,1367]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="48" unit="mm" value="1.0">1 mm</quantity>
, apex obtuse; stamens 3, exserted, the filaments white, flattened,
<quantity id="F95AF8B4FFD7FF89FF33C9687D19FACA" box="[250,338,1376,1403]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="48" unit="mm" value="2.5">2.5 mm</quantity>
long, inserted in the upper portion of the tube, anthers 0.2 ×
<quantity id="F95AF8B4FFD7FF89FBD1C9687838FACA" box="[1048,1139,1376,1403]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="48" unit="mm" value="0.1">0.1 mm</quantity>
; style slightly exserted, the stigma capitate, the ovary cylindrical 0.7 ×
<quantity id="F95AF8B4FFD7FF89FDA7C98C7E8DFA2F" box="[622,710,1412,1438]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="48" unit="mm" value="0.3">0.3 mm</quantity>
, glabrous with 1 pendent ovule. Fruits unknown.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD7FF89FF74C9AF7FD1F9E1" pageId="1" pageNumber="48" type="materials_examined">
<paragraph id="3E1D5551FFD7FF89FF74C9AF7FD1F9E1" blockId="1.[136,1452,986,2077]" pageId="1" pageNumber="48">
<emphasis id="0CD68943FFD7FF89FF74C9AF7E09FA70" bold="true" box="[189,578,1447,1473]" pageId="1" pageNumber="48">Additional specimens examined:</emphasis>
<materialsCitation id="8ECA5F0CFFD7FF89FD83C9AF7F72F9B9" ID-GBIF-Occurrence="4015641301" collectingDate="2012-08-24" collectingDateMax="2016-09-27" collectingDateMin="2012-08-24" collectionCode="S, W, M, E, ECUAMZ, A, J, N" collectorName="D. Neill" country="Ecuador" county="Nangaritza" elevation="2915" latitude="-4.6194444" location="Reserva Biologica Cerro Plateado" longLatPrecision="1" longitude="-78.78306" municipality="Cordillera del Condor" pageId="1" pageNumber="48" specimenCount="1" stateProvince="Zamora-Chinchipe">
<collectingCountry id="46B515C1FFD7FF89FD83C9AF7E9EFA70" box="[586,725,1447,1473]" name="Ecuador" pageId="1" pageNumber="48">ECUADOR</collectingCountry>
.
<collectingRegion id="FC669BB3FFD7FF89FD29C9AF7FFCFA70" box="[736,951,1447,1473]" country="Ecuador" name="Zamora-Chinchipe" pageId="1" pageNumber="48">Zamora-Chinchipe</collectingRegion>
,
<collectingCounty id="D77C2DDDFFD7FF89FC0AC9AF780AFA70" box="[963,1089,1447,1473]" pageId="1" pageNumber="48">Nangaritza</collectingCounty>
,
<collectingMunicipality id="DE79CF2BFFD7FF89FB85C9AF790EFA70" box="[1100,1349,1447,1473]" pageId="1" pageNumber="48">Cordillera del Cóndor</collectingMunicipality>
,
<location id="3B7D038AFFD7FF89FA98C9A07DEEFA54" LSID="urn:lsid:plazi:treatment:B60BE447FFD7FF89FF41CF257F5AF7AC:3B7D038AFFD7FF89FA98C9A07DEEFA54" country="Ecuador" county="Nangaritza" latitude="-4.6194444" longLatPrecision="1" longitude="-78.78306" municipality="Cordillera del Condor" name="Reserva Biologica Cerro Plateado" pageId="1" pageNumber="48" stateProvince="Zamora-Chinchipe">Reserva Biológica Cerro Plateado</location>
, gently sloping summit area of
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, 4°37’10”
<collectionCode id="58B3CD94FFD7FF89FBE8C9C3787AFA54" box="[1057,1073,1483,1509]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="1" pageNumber="48" type="Herbarium">S</collectionCode>
, 78°46’59”
<collectionCode id="58B3CD94FFD7FF89FB67C9C37887FA57" box="[1198,1228,1483,1510]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="48" type="Herbarium">W</collectionCode>
,
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,
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(fl),
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<collectorName id="93573087FFD7FF89FF3EC9E77D1AF9B9" box="[247,337,1518,1544]" pageId="1" pageNumber="48">D. Neill</collectorName>
,
<collectionCode id="58B3CD94FFD7FF89FE92C9E77D3DF9B9" box="[347,374,1519,1544]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="48" type="Herbarium">M</collectionCode>
. Asanza &amp;
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. Cueva 17494
</emphasis>
(
<collectionCode id="58B3CD94FFD7FF89FD79C9E67F66F9B9" box="[688,813,1518,1544]" country="0" httpUri="http://grbio.org/cool/bky3-qnid" name="Universidad Estatal Amazónica" pageId="1" pageNumber="48" type="Herbarium">ECUAMZ</collectionCode>
);
</materialsCitation>
<materialsCitation id="8ECA5F0CFFD7FF89FCF6C9E7782BF99D" ID-GBIF-Occurrence="4015641303" collectingDate="2012-08-24" collectingDateMax="2016-09-27" collectingDateMin="2012-08-24" collectionCode="S, W, M, E, ECUAMZ, A, J, N" collectorName="D. Neill" country="Ecuador" county="Nangaritza" elevation="2915" latitude="-4.6194444" location="Reserva Biologica Cerro Plateado" longLatPrecision="1" longitude="-78.78306" municipality="Cordillera del Condor" pageId="1" pageNumber="48" specimenCount="1" stateProvince="Zamora-Chinchipe">
en la cima de la meseta,
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,
<geoCoordinate id="5B963396FFD7FF89FB11C9E67922F9B8" box="[1240,1385,1518,1545]" degrees="78.7830556" direction="west" orientation="longitude" pageId="1" pageNumber="48" precision="1" value="-78.78306">-78.7830556</geoCoordinate>
,
<quantity id="F95AF8B4FFD7FF89FABAC9E67CEBF99D" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="48" unit="m" value="2900.0">
<elevation id="B58FB262FFD7FF89FABAC9E67CEBF99D" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="48" unit="m" value="2900.0">2900 m</elevation>
</quantity>
,
<date id="4A1C7391FFD7FF89FF65CA1A7DCDF99D" box="[172,390,1554,1580]" pageId="1" pageNumber="48" value="2016-09-23">
<collectingDate id="5A588A79FFD7FF89FF65CA1A7DCDF99D" box="[172,390,1554,1580]" pageId="1" pageNumber="48" value="2016-09-23">23 September 2016</collectingDate>
</date>
(fl),
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<collectionCode id="58B3CD94FFD7FF89FE76CA1B7D98F99F" box="[447,467,1555,1582]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="48" type="Herbarium">Á</collectionCode>
.
<collectionCode id="58B3CD94FFD7FF89FE1FCA1B7DADF99F" box="[470,486,1555,1582]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="1" pageNumber="48" type="Herbarium">J</collectionCode>
. Pérez,
<collectionCode id="58B3CD94FFD7FF89FDF7CA1A7E1FF99A" box="[574,596,1554,1579]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="48" type="Herbarium">N</collectionCode>
. Zapata &amp;
<collectionCode id="58B3CD94FFD7FF89FD1BCA1A7EA1F99A" box="[722,746,1554,1579]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="48" type="Herbarium">W</collectionCode>
. Santillán 10175
</emphasis>
(QCA-244424);
</materialsCitation>
<materialsCitation id="8ECA5F0CFFD7FF89FBA1CA197FDCF9E1" ID-GBIF-Occurrence="4015641307" collectingDate="2012-08-24" collectingDateMax="2016-09-27" collectingDateMin="2012-08-24" collectionCode="S, W, M, E, ECUAMZ, A, J, N" collectorName="D. Neill" country="Ecuador" county="Nangaritza" elevation="2915" latitude="-4.6194444" location="Reserva Biologica Cerro Plateado" longLatPrecision="1" longitude="-78.78306" municipality="Cordillera del Condor" pageId="1" pageNumber="48" specimenCount="1" stateProvince="Zamora-Chinchipe">
<quantity id="F95AF8B4FFD7FF89FBA1CA19794EF99D" box="[1128,1285,1553,1580]" metricMagnitude="3" metricUnit="m" metricValue="2.875" metricValueMax="2.9" metricValueMin="2.85" pageId="1" pageNumber="48" unit="m" value="2875.0" valueMax="2900.0" valueMin="2850.0">
<elevation id="B58FB262FFD7FF89FBA1CA19794EF99D" box="[1128,1285,1553,1580]" metricMagnitude="3" metricUnit="m" metricValue="2.875" metricValueMax="2.9" metricValueMin="2.85" pageId="1" pageNumber="48" unit="m" value="2875.0" valueMax="2900.0" valueMin="2850.0">2850–2900 m</elevation>
</quantity>
,
<date id="4A1C7391FFD7FF89FAD8CA1A7C8BF9FE" pageId="1" pageNumber="48" value="2016-09-27">
<collectingDate id="5A588A79FFD7FF89FAD8CA1A7C8BF9FE" pageId="1" pageNumber="48" value="2016-09-27">27 September 2016</collectingDate>
</date>
(fl),
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<collectionCode id="58B3CD94FFD7FF89FF30CA3F7D46F9E3" box="[249,269,1591,1618]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="48" type="Herbarium">Á</collectionCode>
.
<collectionCode id="58B3CD94FFD7FF89FED8CA3F7D6AF9E3" box="[273,289,1591,1618]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="1" pageNumber="48" type="Herbarium">J</collectionCode>
. Pérez,
<collectionCode id="58B3CD94FFD7FF89FEB1CA3E7DC5F9FE" box="[376,398,1590,1615]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="48" type="Herbarium">N</collectionCode>
. Zapata &amp;
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. Santillán 10329
</emphasis>
(QCA-244532)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD7FF89FF74CA507925F927" pageId="1" pageNumber="48" type="diagnosis">
<paragraph id="3E1D5551FFD7FF89FF74CA507925F927" blockId="1.[136,1452,986,2077]" pageId="1" pageNumber="48">
<taxonomicName id="F9A22ED2FFD7FF89FF74CA507DF5F9C3" authorityName="A. J. Perez, C. Persson &amp; J. N. Zapata" authorityYear="2023" box="[189,446,1624,1650]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="1" pageNumber="48" phylum="Tracheophyta" rank="species" species="plateadensis">
<emphasis id="0CD68943FFD7FF89FF74CA507DF5F9C3" box="[189,446,1624,1650]" italics="true" pageId="1" pageNumber="48">Valeriana plateadensis</emphasis>
</taxonomicName>
is distinguished from other
<taxonomicName id="F9A22ED2FFD7FF89FD35CA507F8AF9C3" box="[764,961,1624,1650]" pageId="1" pageNumber="48">
<emphasis id="0CD68943FFD7FF89FD35CA507F22F9C3" box="[764,873,1624,1650]" italics="true" pageId="1" pageNumber="48">Valeriana</emphasis>
species
</taxonomicName>
of the high Andes with 3-lobed corollas by being small sparsely branched shrubs with small spatulate and sharply bent densely set leaves in multiple ranks.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD7FF89FF74CA977979F895" pageId="1" pageNumber="48" type="distribution">
<paragraph id="3E1D5551FFD7FF89FF74CA977979F895" blockId="1.[136,1452,986,2077]" pageId="1" pageNumber="48">
<emphasis id="0CD68943FFD7FF89FF74CA977E44F908" bold="true" box="[189,527,1695,1722]" pageId="1" pageNumber="48">Distribution and habitat:—</emphasis>
<taxonomicName id="F9A22ED2FFD7FF89FDC6CA977F5FF908" authorityName="A. J. Perez, C. Persson &amp; J. N. Zapata" authorityYear="2023" box="[527,788,1695,1721]" class="Magnoliopsida" family="Caprifoliaceae" genus="Valeriana" kingdom="Plantae" order="Dipsacales" pageId="1" pageNumber="48" phylum="Tracheophyta" rank="species" species="plateadensis">
<emphasis id="0CD68943FFD7FF89FDC6CA977F5FF908" box="[527,788,1695,1721]" italics="true" pageId="1" pageNumber="48">Valeriana plateadensis</emphasis>
</taxonomicName>
is an endemic species to Cerro Plateado, Cordillera del Cóndor in southeast
<collectingCountry id="46B515C1FFD7FF89FEBDCACB7D9AF96C" box="[372,465,1731,1757]" name="Ecuador" pageId="1" pageNumber="48">Ecuador</collectingCountry>
where it has been collected on sandstone soil between rocks on the top of the Andean tepui at
<quantity id="F95AF8B4FFD7FF89FF2ECAEF7D74F8B0" box="[231,319,1767,1793]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="48" unit="m" value="2900.0">2900 m</quantity>
(
<figureCitation id="A69949D4FFD7FF89FE99CAEE7D81F8B0" box="[336,458,1766,1793]" captionStart="FIGURE 3" captionStartId="4.[136,229,1913,1935]" captionTargetBox="[357,1231,749,1889]" captionTargetId="figure-243@4.[357,1231,749,1889]" captionTargetPageId="4" captionText="FIGURE 3. A. Habitat of Valeriana plateadensis at the summit of the Cerro Plateado Biological Reserve. Photograph by Álvaro J. Pérez. B. Habitat of Valeriana yacuriensis at the Lagunas Negras in the Páramo de Jimbura, Yacuri National Park. Photograph by Petr Sklenář. C. Habitat of Valeriana xenophylloides in the Páramo de Patococha. Photograph by Petr Sklenář." figureDoi="http://doi.org/10.5281/zenodo.7532117" httpUri="https://zenodo.org/record/7532117/files/figure.png" pageId="1" pageNumber="48">Figure 3A</figureCitation>
). Associated plants include
<taxonomicName id="F9A22ED2FFD7FF89FCC5CAEE78DDF8B1" authority="Stahl (2010: 86)" authorityName="Stahl" authorityPageNumber="86" authorityYear="2010" box="[780,1174,1766,1793]" class="Magnoliopsida" family="Symplocaceae" genus="Symplocos" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="48" phylum="Tracheophyta" rank="species" species="neillii">
<emphasis id="0CD68943FFD7FF89FCC5CAEE7F85F8B1" box="[780,974,1766,1792]" italics="true" pageId="1" pageNumber="48">Symplocos neillii</emphasis>
<bibRefCitation id="5A3328A0FFD7FF89FC11CAEE78DDF8B1" author="Stahl, E. B." box="[984,1174,1766,1793]" pageId="1" pageNumber="48" pagination="84 - 94" refId="ref4695" refString="Stahl, E. B. (2010) Additions to the knowledge of the genus Symplocos (Symplocaceae) in Ecuador and Peru. Novon: A Journal for Botanical Nomenclature 20 (1): 84 - 94. https: // doi. org / 10.3417 / 2008079" type="journal article" year="2010">Ståhl (2010: 86)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="F9A22ED2FFD7FF89FB6DCAEE7902F8B1" authority="Persson &amp; Eriksen &amp; Pérez &amp; Zapata &amp; Couvreur &amp; Sklenář, 2023" authorityName="Persson &amp; Eriksen &amp; Pérez &amp; Zapata &amp; Couvreur &amp; Sklenář" authorityYear="2023" box="[1188,1353,1766,1792]" class="Magnoliopsida" family="Asteraceae" genus="Diplostephium" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="48" phylum="Tracheophyta" rank="genus" status="sp. nov.">
<emphasis id="0CD68943FFD7FF89FB6DCAEE7902F8B1" box="[1188,1353,1766,1792]" italics="true" pageId="1" pageNumber="48">Diplostephium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="17E53438FFD7FF89FA9ACAEF79ECF8B0" box="[1363,1447,1767,1793]" pageId="1" pageNumber="48" rank="species">sp. nov</taxonomicNameLabel>
,
<taxonomicName id="F9A22ED2FFD7FF89FF41CB027E15F895" authority="(Clark) Clark (2009: 681)" authorityName="Clark" authorityPageNumber="681" authorityYear="2009" baseAuthorityName="Clark" box="[136,606,1802,1828]" class="Liliopsida" family="Poaceae" genus="Chusquea" kingdom="Plantae" order="Poales" pageId="1" pageNumber="48" phylum="Tracheophyta" rank="species" species="nana">
<emphasis id="0CD68943FFD7FF89FF41CB027D7CF895" box="[136,311,1802,1828]" italics="true" pageId="1" pageNumber="48">Chusquea nana</emphasis>
(Clark) Clark (2009: 681)
</taxonomicName>
and
<taxonomicName id="F9A22ED2FFD7FF89FD5CCB0378D4F894" authority="Silva &amp; Correa (2002: 543)" authorityName="Silva &amp; Correa" authorityPageNumber="543" authorityYear="2002" box="[661,1183,1802,1829]" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="48" phylum="Tracheophyta" rank="species" species="peruensis">
<emphasis id="0CD68943FFD7FF89FD5CCB037F28F895" box="[661,867,1803,1828]" italics="true" pageId="1" pageNumber="48">Drosera peruensis</emphasis>
<bibRefCitation id="5A3328A0FFD7FF89FCA2CB0278D4F894" author="Silva, T. R. S. &amp; Correa A." box="[875,1183,1802,1829]" pageId="1" pageNumber="48" pagination="543 - 545" refId="ref4637" refString="Silva, T. R. S. &amp; Correa A., M. D. (2002) Drosera peruensis (Droseraceae), a new species from Peru. Novon, A Journal for Botanical Nomenclature 12 (4): 543 - 545. https: // doi. org / 10.2307 / 3393136" type="journal article" year="2002">Silva &amp; Correa (2002: 543)</bibRefCitation>
</taxonomicName>
(
<figureCitation id="A69949D4FFD7FF89FB64CB02796EF895" box="[1197,1317,1802,1828]" captionStart="FIGURE 3" captionStartId="4.[136,229,1913,1935]" captionTargetBox="[357,1231,749,1889]" captionTargetId="figure-243@4.[357,1231,749,1889]" captionTargetPageId="4" captionText="FIGURE 3. A. Habitat of Valeriana plateadensis at the summit of the Cerro Plateado Biological Reserve. Photograph by Álvaro J. Pérez. B. Habitat of Valeriana yacuriensis at the Lagunas Negras in the Páramo de Jimbura, Yacuri National Park. Photograph by Petr Sklenář. C. Habitat of Valeriana xenophylloides in the Páramo de Patococha. Photograph by Petr Sklenář." figureDoi="http://doi.org/10.5281/zenodo.7532117" httpUri="https://zenodo.org/record/7532117/files/figure.png" pageId="1" pageNumber="48">Figure 3A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD7FF89FF74CB2578D4F803" pageId="1" pageNumber="48" type="conservation">
<paragraph id="3E1D5551FFD7FF89FF74CB2578D4F803" blockId="1.[136,1452,986,2077]" pageId="1" pageNumber="48">
<emphasis id="0CD68943FFD7FF89FF74CB257DFFF8F9" bold="true" box="[189,436,1837,1864]" pageId="1" pageNumber="48">Conservation status:</emphasis>
Only one population with ca. 40 mature individuals of this species were discovered at the
<typeStatus id="E119EBF3FFD7FF89FF41CB597CF3F8DA" box="[136,184,1873,1899]" pageId="1" pageNumber="48">type</typeStatus>
locality at the summit of the Cerro Plateado. It is an isolated area and difficult to access; nevertheless, climate change effects could affect the distribution and population size of this species negatively. According to the IUCN Red List criteria (
<bibRefCitation id="5A3328A0FFD7FF89FED1CB907DEBF803" author="IUCN" box="[280,416,1944,1970]" pageId="1" pageNumber="48" refId="ref4312" refString="IUCN (2022) Guidelines for using the IUCN Red List categories and criteria. Version 15.1. Prepared by the Standards and Petitions Committee. Available from: https: // www. iucnredlist. org / resources / redlistguidelines (accessed 12 December 2022)" type="url" year="2022">IUCN 2022</bibRefCitation>
) this species is therefore assessed as Vulnerable (VU, Criterion D2).
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD7FF89FF74CBB37F3EF864" box="[189,885,1979,2006]" pageId="1" pageNumber="48" type="biology_ecology">
<paragraph id="3E1D5551FFD7FF89FF74CBB37F3EF864" blockId="1.[136,1452,986,2077]" box="[189,885,1979,2006]" pageId="1" pageNumber="48">
<emphasis id="0CD68943FFD7FF89FF74CBB37D08F864" bold="true" box="[189,323,1979,2005]" pageId="1" pageNumber="48">Phenology:</emphasis>
Flowers were collected in August and September.
</paragraph>
</subSubSection>
<subSubSection id="76B806DAFFD7FF89FF74CBD77F5AF7AC" pageId="1" pageNumber="48" type="etymology">
<paragraph id="3E1D5551FFD7FF89FF74CBD77F5AF7AC" blockId="1.[136,1452,986,2077]" pageId="1" pageNumber="48">
<emphasis id="0CD68943FFD7FF89FF74CBD77D03F848" bold="true" box="[189,328,2015,2041]" pageId="1" pageNumber="48">Etymology:</emphasis>
The specific epithet of this species refers to Cerro Plateado in the Cordillera del Cóndor in southeastern
<collectingCountry id="46B515C1FFD7FF89FF41C40B7CAEF7AC" box="[136,229,2051,2077]" name="Ecuador" pageId="1" pageNumber="48">Ecuador</collectingCountry>
which is the only known locality for this species.
</paragraph>
</subSubSection>
</treatment>
</document>

85
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<mods:title id="468C0E6034C4DAF11471F0DC94A15E81">Pictorial key to species of the genus Ropalidia Guerin-Meneville, 1831 (Hymenoptera, Vespidae) from China, with description of one new species</mods:title>
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<taxonomicNameLabel id="DB703A5EB0DA1D6D028426C7B89A67CD" pageId="19" pageNumber="20">rec. n.</taxonomicNameLabel>
Figure 1J
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<taxonomicName id="038E16690F0A01D9B824995F79060133" class="Insecta" family="Vespidae" genus="Ropalidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ropalidia obscura" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="obscura">Ropalidia obscura</taxonomicName>
Gusenleitner, 1996: 15;
<bibRefCitation id="F0C1BE5732E5F7745770876510F6AE54" author="Kojima, J" journalOrPublisher="Natural History Bulletin of Ibaraki University" pageId="26" pageNumber="27" pagination="1 - 13" title="Social wasps collected by Malaise trapping in Southeast Asia, with a note on relative abundance of swarm-founding species (Insecta: Hymenoptera: Vespidae)." volume="1" year="1997">Kojima and van Achterberg 1997</bibRefCitation>
: 10. Type locality: Thailand.
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<paragraph id="6007FE6FA18FB96A40B23DBCEBFC97C6" pageId="19" pageNumber="20">Specimens examined.</paragraph>
<paragraph id="74533E51131C1D7D3ADD1E1ABCF9117F" pageId="19" pageNumber="20">
CHINA: 3 ♀ (NWUM), Mengla, Yaoqu, Yunnan, 27-30.ix.2010, JL Tan; 1 ♀ (NWUM), Longmen, Shangyong, Yunnan, 26.ix.2010, JL Tan; 1 ♀ (NWUM), Banna, Menglun, Yunnan, 5.x.2010, JL Tan. THAILAND: 1 ♀ (RMNH), 50 km SW Loei (
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,
<geoCoordinate id="B5778F0B8F2D9C53820823377857459D" direction="east" orientation="longitude" precision="925" value="101.333336">101°20'E</geoCoordinate>
), Phu Rua N. P, 14.vii.1986, R Hensen; (labeled
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Gusenleitner, 1996, by J Kojima in 1996).
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<paragraph id="31E2478FF6F73509B72C588381B424B3" pageId="19" pageNumber="20">Distribution.</paragraph>
<paragraph id="90D6D29AA66FA6AD560857807533F9C4" pageId="19" pageNumber="20">
China (Yunnan), Thailand. (
<bibRefCitation id="8EC06B17E114FB8A6A5FC6634A6BD25E" author="Kojima, J" journalOrPublisher="American Museum Novitates" pageId="26" pageNumber="27" pagination="1 - 96" title="Catalog of species in the polistine tribe Ropalidiini (Hymenoptera: Vespidae)." volume="3199" year="1997">Kojima and Carpenter 1997</bibRefCitation>
and updated to 2006).
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168
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Artbeschreibung:
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teils ungeteilt, teils mit 1-2 stumpfen seitlichen Abschnitten, oft asymmetrisch,
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lang,
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Verbreitung global: Mittel- und
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Zeigerwerte nach
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<emphasis id="4C58810089839BADA678B2372B9780ED" bold="true" pageNumber="212">Klimafaktoren</emphasis>
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<td id="4298255FC691F071AC13A40745014D72" pageNumber="212">Feuchtezahl F</td>
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feucht
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<td id="847592CD4B1E0A719A0AC417EC976D73" pageNumber="212">Salzzeichen</td>
<td id="3716C517B316A1C4CE4B6D0E840E8078" pageNumber="212" style="text-align:left;width:60px">--</td>
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<td id="D5BBE4AA6D0B4871A1B421CBBB8EA3EC" pageNumber="212" style="text-align:left">neutral bis basisch (pH 5.5-8.5)</td>
<td id="E9FC043770ECC4CADFD057C4B2A3C7B0" pageNumber="212">Temperaturzahl T</td>
<td id="2CBE730F1588B2F8FF3BB91485134A9B" pageNumber="212" style="text-align:left">alpin und nival (von der Baumgrenze bis zur Schneegrenze)</td>
</tr>
<tr id="87C112C8FFFDAA252CD1EF97E43618B5" pageNumber="212">
<td id="26A73A79ADE68B6509C0BE9F06236E8D" pageNumber="212">
<normalizedToken id="9BCC2475883BFBE4D51819063D03540C" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
<td id="550E3D692320CA141CB5E553BE5DE0C2" pageNumber="212" style="text-align:left">
sehr
<normalizedToken id="F4F3C024C57BDEAE91F42121160B794A" originalValue="nährstoffarm">naehrstoffarm</normalizedToken>
</td>
<td id="D7E2810D0AFB829F97220DB1F7FD6EED" pageNumber="212">
<normalizedToken id="4BAC9BA63C14EC95CF385BC99E46ED8F" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="13700872D54017AF2CA0AE61E772A6FA" pageNumber="212" style="text-align:left">subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter)</td>
</tr>
</table>
</paragraph>
</subSubSection>
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<paragraph id="B0A7AAA56322D2C2D6C9D3E823D29E3D" pageNumber="212">
Volksname Deutscher Name:
<vernacularName id="742C49E60FBEA6DE9A44A3A0A5E201A8" language="deu" pageNumber="212">Moschus-Steinbrech</vernacularName>
Nom
<normalizedToken id="C967D030D5DE2CE24065790C0E16AEB6" originalValue="français">francais</normalizedToken>
:
<vernacularName id="314B08533321A959FE3F28A487C469DD" language="fra" pageNumber="212">
Saxifrage
<normalizedToken id="C87F295448637CF13797FDB432BFED92" originalValue="musquée">musquee</normalizedToken>
</vernacularName>
Nome italiano:
<vernacularName id="9D201F679DC6A4BA22B27193E8B90100" language="ita" pageNumber="212">Sassifraga muschiata</vernacularName>
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125
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<paragraph id="292F657706AA98A79CCE73FA46D51FF5" pageId="3" pageNumber="48">Taxon classification Animalia Ephemeroptera Leptophlebiidae</paragraph>
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<taxonomicName id="0DA9CBDC42E8B9BB44E4EEBF70A66F11" ID-CoL="4ZTL" ID-ENA="1899174" authority="Polegatto &amp; Batista, 2007" authorityName="Polegatto &amp; Batista" authorityYear="2007" class="Insecta" family="Leptophlebiidae" genus="Hydromastodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydromastodon" order="Ephemeroptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="genus">Hydromastodon Polegatto &amp; Batista, 2007</taxonomicName>
Figures 3, 4, 7
</paragraph>
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<paragraph id="9DB14AFEA0938518998588129B5DD37A" pageId="4" pageNumber="49">
<pageBreakToken id="441BD149DC856B66FDCC1EA1A24D6EAB" pageId="4" pageNumber="49" start="start">Diagnosis</pageBreakToken>
.
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The male imago of
<taxonomicName id="C16E2EC5C7B053DCC477D13D484D60AD" class="Insecta" family="Leptophlebiidae" genus="Hydromastodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydromastodon" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="genus">Hydromastodon</taxonomicName>
can be distinguished from the other genera of the
<taxonomicName id="CD697FF791C978225A4C65E7C82AD17C" class="Insecta" family="Leptophlebiidae" genus="Hermanella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermanella" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="genus">Hermanella</taxonomicName>
complex by the following combination of characters: 1) Eyes meeting on meson of head (Fig. 3a); 2) Cross vein above fork of MA slanted (Fig. 4a); 3) Fork of MA asymmetrical and fork of MP slightly asymmetrical (MP2 connected to MP1 by a crossvein); 4) Styliger plate with a strong dorsally curved median projection (Fig. 4d, e); 5) Penis divided, each lobe with a long spine ventromedially directed (Fig. 4d, e).
</paragraph>
<caption id="CBD538C061B2803D8497C287C9D10AFA" pageId="4" pageNumber="49">
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Figure 3.
<taxonomicName id="EAFA388641711A6F409080B3DC49B32D" class="Insecta" family="Leptophlebiidae" genus="Hydromastodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydromastodon sallesi" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="sallesi">Hydromastodon sallesi</taxonomicName>
, male imago: a dorsal view b head and thorax, lateral view.
</paragraph>
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Figure 4.
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, male imago: a fore wing b hind wing c hind wing, enlarged d genitalia, ventral view e genitalia, lateral view.
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<paragraph id="2B5745E2380489F86D638A5E359BC020" pageId="4" pageNumber="49">Description of male imago</paragraph>
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(in alcohol). Head (Fig. 3a, b): Eyes meeting on meson of head, lower portion of eyes slightly &lt;
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length of upper portion.
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Thorax: Prosternum with rather wide, X-shaped median carina, with similar anterior and posterior arms; similar to
<taxonomicName id="C5313583700B0C50A5642CA543424EFE" class="Insecta" family="Leptophlebiidae" genus="Needhamella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Needhamella" order="Ephemeroptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="genus">Needhamella</taxonomicName>
, as shown by
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and Flowers (1989
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: fig. 18).
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Wings (Fig. 4a, b, c): Maximum width of fore wings 1/3 their maximum length; maximum width of hind wings about
<normalizedToken id="D92C3A0E75F75662BD56000B8D792D15" originalValue="½">1/2</normalizedToken>
their maximum length; maximum length of hind wings 1/6 maximum length of fore wings. Fore wings (Fig. 4a): vein Rs forked slightly&gt; 1/4 distance from base of vein to margin, fork of vein MA asymmetrical and forked at
<normalizedToken id="7BE8969A23A18A6FDD7F587FCC05E369" originalValue="½">1/2</normalizedToken>
distance from base of vein to margin, cross vein above fork of MA slanted; fork of vein MP slightly asymmetrical and forked 1/3 distance from base of vein to margin; vein ICu1 attached at base to vein CuA by crossvein; vein ICu2 free basally.
<pageBreakToken id="C0F2502527B3CC0C6A84F1E0C84A8931" pageId="5" pageNumber="50" start="start">Hind</pageBreakToken>
wings (Fig. 4b, c): costal projection well-developed, acute and located
<normalizedToken id="1E5CF2B91C21433BB6746C1430E6FD25" originalValue="½">1/2</normalizedToken>
distance from base to apex; vein MP unforked; apex of wings rounded; vein Sc
<normalizedToken id="C8E679B9C9F6412612EA1AB6F2C3FABF" originalValue="½">1/2</normalizedToken>
distance from base to wing margin, ending in crossvein; 5 cross veins present.
</paragraph>
<paragraph id="CF048A21BDA725AE443A600E5AE1318E" pageId="5" pageNumber="50">Legs. Ratio of segments of male forelegs, 0.6:1.0 (0.62 mm): 0.03:0.31:0.28:0.15: 0.08. Claws on each leg dissimilar, with one apically hooked and one blunt, pad-like.</paragraph>
<paragraph id="860B8634C733E0BF08FD5E9CC967E898" pageId="5" pageNumber="50">Abdomen: Genitalia (Fig. 4d, e) with segment II of forceps subequal to segment III; segment II of forceps 1/5 length of segment I; styliger plate with strong, dorsally curved, median projection. Penis divided, each lobe with long spine ventromedially directed. Caudal filaments broken off and lost.</paragraph>
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<taxonomicName id="BD49D4639F327CFB0FCD91902C8BD1CA" ID-CoL="8LZJ8" authority="Ellerman and Morrison-Scott 1951" authorityName="Ellerman and Morrison-Scott" authorityYear="1951" class="Mammalia" family="Vespertilionidae" genus="Eptesicus" kingdom="Animalia" order="Chiroptera" pageNumber="457" phylum="Chordata" rank="species" species="tatei" subGenus="Eptesicus">
<emphasis id="7FD6F3C94FE1E43DBB64CF333B8E8152" italics="true" pageNumber="457">Eptesicus (Eptesicus) tatei</emphasis>
<bibRefCitation id="82C5EE339E2C2A53B71C5B0FEF9FB653" author="Ellerman, J. R. &amp; T. C. S. Morrison-Scott" refId="ref93180" refString="Ellerman, J. R., and T. C. S. Morrison-Scott. 1951. Checklist of Palaearctic and Indian mammals 1758 to 1946. Trustees of the British Museum (Natural History), London, 810 pp." year="1951">Ellerman and Morrison-Scott 1951</bibRefCitation>
</taxonomicName>
,
<bibCitation id="CC893624178A582D25CAADF4B1833967" author="Ellerman and Morrison-Scott" pageNumber="457" pagination="158" title="Eptesicus (Eptesicus) tatei" volumeTitle="Checklist Palaearctic Indian Mammals" year="1951">Checklist Palaearctic Indian Mammals: 158</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9FAAF99C95458CBF4D1DE279C6EC9803" pageNumber="457" type="materials_examined">
<paragraph id="9D297DF36D80CD9BEFAE3DD5A1B3F577" pageNumber="457">
<emphasis id="C117AB2677FFB399A46B99D2D6301397" inLineHeading="true" pageNumber="457">Type Locality:</emphasis>
<materialsCitation id="84CAC17325077D787363BCCBCCD7073A" ID-GBIF-Occurrence="4867287336" country="India" pageNumber="457">
<collectingCountry id="D1644D1361B8E8109700D824A4A78856" name="India">India</collectingCountry>
, Darjeeling.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7D5678A722C004CC4A2283BCD8F42723" pageNumber="457" type="vernacular_names">
<paragraph id="3F819E6AAA0D5FDB871D72DB3BFC22B6" pageNumber="457">
<emphasis id="0560CE61A4A1DE6B4B9ACC387F309409" inLineHeading="true" pageNumber="457">Vernacular Names:</emphasis>
<vernacularName id="0A529263B01414B2FFC8F33A43977A36" language="eng" pageNumber="457">Sombre Bat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="D2191418DC3414EE78F932F3E69409FD" pageNumber="457" type="synonym_list">
<paragraph id="981C8CC3E4DBCD497E30F4AD1F8E2C1B" pageNumber="457">
<emphasis id="F441CB069203BD22FB76131B0980D1E1" inLineHeading="true" pageNumber="457">Synonyms:</emphasis>
<taxonomicName id="5698375180A73271AB3E77B6E8EB3B71" authority="Blyth 1863" authorityName="Blyth" authorityYear="1863" class="Mammalia" family="Vespertilionidae" genus="Eptesicus" kingdom="Animalia" order="Chiroptera" pageNumber="457" phylum="Chordata" rank="species" species="atratus" subGenus="Eptesicus">
<emphasis id="7D254F41636588A74CA951EE20A3206E" italics="true" pageNumber="457">Eptesicus (Eptesicus) atratus</emphasis>
Blyth 1863
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B9C0EBF3ECF10839D2391C5481747E22" pageNumber="457" type="distribution">
<paragraph id="B91501995EC78204D7D448F9E83AADB5" pageNumber="457">
<emphasis id="0EA5471F1B3B4096F054160B24957448" inLineHeading="true" pageNumber="457">Distribution:</emphasis>
NE
<collectingCountry id="1DB9626D7D070701304653702CA5458B" name="India">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="34A0DEE3724B354FCE3F010AF9FB6E75" pageNumber="457" type="conservation">
<paragraph id="814F154D23C90D49C005D303E683AAC7" pageNumber="457">
<emphasis id="C592C9C00DD69266F78752DFAC9C8C46" inLineHeading="true" pageNumber="457">Conservation:</emphasis>
<collectionCode id="DB1B2DB13BF24CE63DD336395C886B36">IUCN</collectionCode>
2003 and
<collectionCode id="779AB3EF9CBFE3A10B3245E11B687832">IUCN</collectionCode>
/
<collectionCode id="CFD20CF76A8AC7EC594D374368A5C7B9" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (2001) – Data Deficient.
</paragraph>
</subSubSection>
<subSubSection id="1C0A670E5677FAB9CBB06B74BAB140FF" pageNumber="457" type="discussion">
<paragraph id="10B50BD4997915C68700C27A2A00F990" pageNumber="457">
<emphasis id="B11C666FE5FBFC59024C949C3423D823" inLineHeading="true" pageNumber="457">Discussion:</emphasis>
Subgenus
<taxonomicName id="A2EF0D70934A6532F443FC56879717AF" authorityName="Rafinesque" authorityYear="1820" class="Mammalia" family="Vespertilionidae" genus="Eptesicus" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="genus">
<emphasis id="4559D36FA14504E63F112A38AD0C3E63" italics="true" pageNumber="457">Eptesicus</emphasis>
</taxonomicName>
.
<bibRefCitation id="B22DD2575350CB5C064D0C62F41E9FDE" author="Corbet, G. B. &amp; J. E. Hill" refId="ref67955" refString="Corbet, G. B., and J. E. Hill. 1992. Mammals of the Indomalayan region. A systematic review. Oxford University Press, Oxford, 488 pp." year="1992">Corbet and Hill (1992)</bibRefCitation>
noted that this species is known only from the
<typeStatus id="74F9AEB76A941A31D46E742CB654EA3C">holotype</typeStatus>
, but see
<bibRefCitation id="69BD2BC1F4B508074848DB5B407B2FEF" author="Agrawal, V. C. &amp; P. K. Das &amp; S. Chakraborty &amp; R. K. Ghose &amp; A. K. Mandal &amp; T. K. Chakraborty &amp; A. K. Poddar &amp; J. P. Lal &amp; T. P. Bhattachryya &amp; M. K. Ghosh" refId="ref2295" refString="Agrawal, V. C., P. K. Das, S. Chakraborty, R. K. Ghose, A. K. Mandal, T. K. Chakraborty, A. K. Poddar, J. P. Lal, T. P. Bhattachryya, and M. K. Ghosh. 1992. Mammalia. State Fauna Series 3: Fauna of West Bengal, Part 1: 27 - 169." year="1992">Agrawal et al. (1992)</bibRefCitation>
, who reported additional specimens.
</paragraph>
</subSubSection>
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82
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Santschi,
<taxonomicNameLabel id="B50843A0477E576C34A84C5A523D0B62">new status</taxonomicNameLabel>
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<taxonomicName id="089D6EAD57962312257A7092DA3398BC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142210" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole (Elasmopheidole) cavifrons subsp. longior" lsidName-HNS="Pheidole (Elasmopheidole) cavifrons st. longior Santschi" order="Hymenoptera" pageNumber="136" phylum="Arthropoda" rank="subSpecies" species="cavifrons" subGenus="Elasmopheidole" subSpecies="longior">Pheidole (Elasmopheidole) cavifrons st. longior</taxonomicName>
Santschi 1933f: 110. types Naturhist. Mus. Basel.
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<subSubSection id="B503A54A5EB7FC4B0DFF74FD78D8A94A" type="etymology">
<paragraph id="5DA4B835522F292BB64871B15F6DFC0F" pageNumber="136">
Etymology L
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, longer, referring to the head shape of the major.
</paragraph>
</subSubSection>
<subSubSection id="4D180C6CE3002B9C74246C2C32D10DD3" type="diagnosis">
<paragraph id="B69762992BB2695888FA7BD0C326C4A3" pageNumber="136">
diagnosis Closest to
<taxonomicName id="DE2F2E555F6A805034A4FC9B22B37E6D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33655" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole cavifrons Emery" lsidName-HNS="Pheidole cavifrons Emery" order="Hymenoptera" pageNumber="136" phylum="Arthropoda" rank="species" species="cavifrons">cavifrons</taxonomicName>
, somewhat less similar to
<taxonomicName id="386A71825E3DFA14AF39F019E1806BEE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33564" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole aberrans Mayr" lsidName-HNS="Pheidole aberrans Mayr" order="Hymenoptera" pageNumber="136" phylum="Arthropoda" rank="species" species="aberrans">aberrans</taxonomicName>
and
<taxonomicName id="33C239656DDBA644DDD07CED738329BB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33902" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole obnixa Forel" lsidName-HNS="Pheidole obnixa Forel" order="Hymenoptera" pageNumber="136" phylum="Arthropoda" rank="species" species="obnixa">obnixa</taxonomicName>
, distinguished as follows.
</paragraph>
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<subSubSection id="5517C346609711EBF4BA83FB378EDF56" type="description">
<paragraph id="95E3426AD67BD2884FF61F8354C4825B" pageNumber="136">
Major: head markedly elongate (Head Length/Head Width 1.47), and bicolored (see under Color below); dorsal posterior half of head profile flat; frontal lobes in full-face view taper anteriorly into blunt acute angles; carinulae originating on frontal lobes reach well past midpoint between eyes and occiput; mesosomal convexity in side view reduced, obliquely angular; propodeal spines denticulate; promesonotum smooth and shiny. Minor: nearly identical to minor of
<taxonomicName id="FD3568812159D5AA9AEE4D1017888EDF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33655" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole cavifrons Emery" lsidName-HNS="Pheidole cavifrons Emery" order="Hymenoptera" pageNumber="136" phylum="Arthropoda" rank="species" species="cavifrons">cavifrons</taxonomicName>
(q.v.).
</paragraph>
<paragraph id="AC70116994E42643A07A435AB6FCC5E8" pageNumber="136">Measurements (mm) Lectotype major: HW 0.72, HL 1.06, SL 0.30, EL 0.06, PW 0.50. Paralectotype minor: HW 0.50, HL 0.58, SL 0.36, EL 0.04, PW 0.32.</paragraph>
<paragraph id="D8CEB35A4BC8D8925C2234FEABFEC30D" pageNumber="136">color Major: strikingly bicolored, with anterior fourth of head dark brown, and rest of body medium yellow.</paragraph>
<paragraph id="6AA3747FAACE0DB625A9812902A67A3D" pageNumber="136">Minor: concolorous light yellow.</paragraph>
</subSubSection>
<subSubSection id="3B6E06E12AFE3FE10FD893164F51EEA4" type="distribution">
<paragraph id="2BA3ABC833C6483AD38DF1823026B3BA" pageNumber="136">range Known only from the type locality.</paragraph>
</subSubSection>
<subSubSection id="893EFF730E25CF50E0143EF68423096E" type="biology_ecology">
<paragraph id="801A6E3D39DC194669BB2BF9C0B6A482" pageNumber="136">Biology Unknown.</paragraph>
</subSubSection>
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<paragraph id="2CC20797CB59226EF2287F3F185DF4C1" pageNumber="136">
Figure Upper: lectotype, major. ARGENTTNA: Loreto, Misiones. The minor is nearly identical to the minor of
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(q.v.). Scale bar = 1 mm.
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372
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3.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Uvariastrum pierreanum. A Flowering branch B cauliflorous flower C Transversal cut of flower showing carpels and stamens D detail of sepals E detail of carpels and ovule placentation; stamen G fruit. Uvariastrum inscultptum H Flowering branch I flower bud J sepals K carpel L stamen M fruits. Drawing Helene Lamourdedieux, modified from La Flore du Gabon, Fig. 53. 1969." httpUri="https://binary.pensoft.net/fig/10562" pageId="13" pageNumber="14">Figure 9</figureCitation>
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<taxonomicName authorityName="Engler &amp; Diels" authorityYear="1899" class="Dicotyledoneae" family="Annonaceae" genus="Uvaria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Magnoliales" pageId="13" pageNumber="14" phylum="Angiospermae" rank="species" species="insculpta">Uvaria insculpta</taxonomicName>
Engl. &amp; Diels, Notizbl.
<normalizedToken originalValue="Königl">Koenigl</normalizedToken>
. Bot. Gart. Berlin 2: 295. 1899.
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<paragraph pageId="13" pageNumber="14">Type.</paragraph>
<paragraph pageId="13" pageNumber="14">
Cameroon. South-West Province:
<normalizedToken originalValue="Johann-Albrechtshöhe">Johann-Albrechtshoehe</normalizedToken>
[Kumba], 1896,
<emphasis italics="true" pageId="13" pageNumber="14">A. Staudt 740</emphasis>
(lectotype, designated here: B! [B100153111]; isolectotypes: COI!, EA!, G! 2-sheets [G00011729], K! 3-sheets [K000105338, K000105339, K000105340], P! 2-sheets [P00315828, P00315829], S!).
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="14" type="type">
<paragraph pageId="13" pageNumber="14">Type.</paragraph>
<paragraph pageId="13" pageNumber="14">
Based on
<taxonomicName authorityName="Engler &amp; Diels" authorityYear="1899" class="Dicotyledoneae" family="Annonaceae" genus="Uvaria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Magnoliales" pageId="13" pageNumber="14" phylum="Angiospermae" rank="species" species="insculpta">
<emphasis italics="true" pageId="13" pageNumber="14">Uvaria insculpta</emphasis>
</taxonomicName>
Engl. &amp; Diels.
</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
<paragraph pageId="13" pageNumber="14">Description.</paragraph>
<paragraph lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
Tree or shrub to 4-7 (15) m high, d.b.h. 2.5-5 cm; old branches spreading, glabrous, bark brown-grey; young branches pubescent, hairs ca. 0.5-1 mm long, erect, brown; leaf buds elongated, pubescent, hairs ca. 1 mm long, appressed, light brown. Petioles 1-2(4) mm long, 1 mm in diameter; pubescent, hairs ca. 0.4-0.8 mm long, erect, red brown, persisting in older leaves; leaf lamina inserted on top, weakly grooved adaxially. Leaf lamina 6-14 cm long, 2-4 cm wide, length:width ratio
<pageBreakToken pageId="14" pageNumber="15" start="start">2.5</pageBreakToken>
-4, narrowly elliptic to elliptic or narrowly obovate to obovate, papyraceous to sub coriaceous, glabrous to very sparsely pubescent adaxially, sparsely pubescent quickly becoming glabrous abaxially; dark green adaxially; lighter green abaxially; base rounded to subcordate, apex acuminate, acumen 1-2 cm long; midrib pubescent in young
<pageBreakToken pageId="15" pageNumber="16" start="start">leaves</pageBreakToken>
quickly becoming glabrous adaxially, hairs ca. 0.1 mm long, appressed, brown; densely to sparsely pubescent abaxially, hairs ca. 0.3 mm long, appressed, brown; secondary veins 8-12, straight to curving upwards, clearly anatomizing towards margins, glabrous, imprinted adaxially; sparsely pubescent, hairs ca. 0.3 mm long, appressed, brown, clearly visible abaxially. Rhipidia 1-2, on leafy branches, and one report of cauliflory. Flowering pedicels 0.8-1.5 cm long, 1-2 mm in diameter, densely pubescent, hairs 0.2-0.5 mm long, appressed, light brown. Bracts up to 3, ca. 4 mm long, 3-4 mm wide, basal, densely pubescent outside, hairs ca. 0.5 mm long, light to darker brown, glabrous inside. Sepals 0.7-1.5 cm long, 5-8 mm wide, length:width ratio 1.4-2.5, ovate to broadly ovate, base truncate, apex acute, margins slightly folded; densely pubescent, hairs ca. 0.5 mm long, appressed, light brown when dried; horizontally spreading in mature flowers, falling when in fruit, green to light green or pale yellow with darker margins in fresh flowers. Outer petals 2.3-3.5 cm long, 0.5-1 cm wide, length:width ratio 2-4, ovate to narrowly ovate, base narrowing, apex acute, densely pubescent to tomentose outside, hairs ca. 0.1 mm long, appressed, light brown, more densely pubescent along central vein, sparsely pubescent to tomentose inside, hairs ca. 0.1 mm long, appressed, light brown; pale yellow to white in fresh flowers. Inner petals smaller than outer ones, 1-2 cm long, 0.6-1 cm wide, length:width ratio 2-4, ovate to narrowly ovate, base narrowing, apex acute, pubescence and color same as outer petals; pale yellow to white in fresh flowers. Stamens 2-2.5 mm long, connective discoid, ca. 0.5 mm in diameter, densely pubescent; bright red. Carpels 6-7, falling off before stamens, 2-3 mm long, ca. 1 mm in diameter, densely pubescent, hairs 0.7-1 mm long, apprressed upwards, light brown; stigma ca. 1.5 mm in diameter, bilobed, glabrous, yellow to creamy; ovules 19-26, biseriate. Fruiting pedicels 1-2 cm long, 2-5 mm in diameter, woody, densely to sparsely pubescent, hairs ca. 0.5 mm long, appressed, light brown. Monocarps 2-8, 3-6 cm long, 1-2 cm in diameter, oblong to narrowly oblong, straight or bending, longitudinally ribbed, ribs ca. 4-6, resembling that of a peanut, densely pubescent all over, more so along ribs, hairs ca. 0.2 mm long, light brown; stipe 2-6 mm long, 2-5 mm in diameter; rostre 2-4 mm long. Seeds, ca. 20 per monocarp, 1-1.5 cm long, 0.7-0.9 cm wide, ellipsoid flat, 3-5 mm in depth, testa dark brown; raphe raised, slightly darker brown; hilum 2-4 mm long, 1-1.5 mm wide, narrowly elliptic to narrowly ovate.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10562" pageId="15" pageNumber="16" start="Figure 9" startId="F9">
<paragraph pageId="15" pageNumber="16">
<emphasis bold="true" pageId="15" pageNumber="16">Figure 9.</emphasis>
<taxonomicName class="Magnoliopsida" family="Annonaceae" genus="Uvariastrum" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Magnoliales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="pierreanum">
<emphasis italics="true" pageId="15" pageNumber="16">Uvariastrum pierreanum</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="15" pageNumber="16">A</emphasis>
Flowering branch
<emphasis bold="true" pageId="15" pageNumber="16">B</emphasis>
cauliflorous flower
<emphasis bold="true" pageId="15" pageNumber="16">C</emphasis>
Transversal cut of flower showing carpels and stamens
<emphasis bold="true" pageId="15" pageNumber="16">D</emphasis>
detail of sepals
<emphasis bold="true" pageId="15" pageNumber="16">E</emphasis>
detail of carpels and ovule placentation; stamen
<emphasis bold="true" pageId="15" pageNumber="16">G</emphasis>
fruit.
<taxonomicName class="Magnoliopsida" family="Annonaceae" genus="Uvariastrum" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Magnoliales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="inscultptum">
<emphasis italics="true" pageId="15" pageNumber="16">Uvariastrum inscultptum</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="15" pageNumber="16">H</emphasis>
Flowering branch
<emphasis bold="true" pageId="15" pageNumber="16">I</emphasis>
flower bud
<emphasis bold="true" pageId="15" pageNumber="16">J</emphasis>
sepals
<emphasis bold="true" pageId="15" pageNumber="16">K</emphasis>
carpel
<emphasis bold="true" pageId="15" pageNumber="16">L</emphasis>
stamen
<emphasis bold="true" pageId="15" pageNumber="16">M</emphasis>
fruits. Drawing
<normalizedToken originalValue="Helène">Helene</normalizedToken>
Lamourdedieux, modified from
<emphasis italics="true" pageId="15" pageNumber="16">La Flore du Gabon</emphasis>
, Fig. 53. 1969.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="distribution">
<paragraph pageId="15" pageNumber="16">Distribution.</paragraph>
<paragraph pageId="15" pageNumber="16">
Mainly a West African species, found in Liberia and Ivory Coast, two collections in Cameroon and two in Gabon;
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution of Uvariastrum insculptum." httpUri="https://binary.pensoft.net/fig/10563" pageId="15" pageNumber="16">Figure 10</figureCitation>
.
</paragraph>
<paragraph pageId="15" pageNumber="16">
<emphasis bold="true" pageId="15" pageNumber="16">Habitat and ecology.</emphasis>
Found in lowland primary and secondary rain forest 0-400 m.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="phenology">
<paragraph pageId="15" pageNumber="16">Phenology.</paragraph>
<paragraph pageId="15" pageNumber="16">Mature flowers found between Oct and Feb. Mature fruits found in Feb and from Apr till May and in Jul.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="preliminary iucn conservation status">
<paragraph pageId="15" pageNumber="16">Preliminary IUCN conservation status.</paragraph>
<paragraph pageId="15" pageNumber="16">
LC:
<taxonomicName class="Magnoliopsida" family="Annonaceae" genus="Uvariastrum" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Magnoliales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="insculptum">
<emphasis italics="true" pageId="15" pageNumber="16">Uvariastrum insculptum</emphasis>
</taxonomicName>
is well represented in herbaria and has a large distribution across West Africa, and a few specimens from Central Africa. It is also present in several protected areas such as national parks or protected areas (
<normalizedToken originalValue="Taï">Tai</normalizedToken>
National Park, Banco National Park, Monogaga Forest Reserve (Ivory Coast); Stubbs Creek Forest Reserve (Nigeria)). The Least Concern category thus seems appropriate.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="vernacular names">
<paragraph pageId="16" pageNumber="17">
<pageBreakToken pageId="16" pageNumber="17" start="start">Vernacular</pageBreakToken>
names.
</paragraph>
<paragraph pageId="16" pageNumber="17">None known.</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="uses">
<paragraph pageId="16" pageNumber="17">Uses.</paragraph>
<paragraph pageId="16" pageNumber="17">None known.</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="notes">
<paragraph pageId="16" pageNumber="17">Notes.</paragraph>
<paragraph pageId="16" pageNumber="17">This is the only largely West African species of the genus, although it occurs in the South-West region of Cameroon. It is easily distinguishable by its densely pubescent leaves and midrib and its impressed venation of the upper side of the leaves.</paragraph>
<paragraph pageId="16" pageNumber="17">
Two Staudt collections were cited in the protologue of
<taxonomicName authorityName="Engler &amp; Diels" authorityYear="1899" class="Dicotyledoneae" family="Annonaceae" genus="Uvaria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Magnoliales" pageId="16" pageNumber="17" phylum="Angiospermae" rank="species" species="insculpta">
<emphasis italics="true" pageId="16" pageNumber="17">Uvaria insculpta</emphasis>
</taxonomicName>
: n. 740 and n. 900. Although it would appear as if the former was already considered as the lectotype I found no publication formalizing this (e.g., Global Plants), so I have lectotypified it here using because it has several sheets compared to one located for #900.
</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="specimens examined">
<paragraph pageId="16" pageNumber="17">Specimens examined.</paragraph>
<paragraph pageId="16" pageNumber="17">
<emphasis bold="true" pageId="16" pageNumber="17">Cameroon.</emphasis>
<emphasis bold="true" pageId="16" pageNumber="17">South-West Province</emphasis>
:
<normalizedToken originalValue="Johann-Albrechtshöhe">Johann-Albrechtshoehe</normalizedToken>
Kumba,
<emphasis italics="true" pageId="16" pageNumber="17">Staudt, A. 900</emphasis>
(G); Likomba Pflanzung, 15-35 km NE von Victoria, Dec 1928,
<emphasis italics="true" pageId="16" pageNumber="17">Mildbraed, G.W.J. 10795</emphasis>
(A, K).
</paragraph>
<paragraph pageId="16" pageNumber="17">
<emphasis bold="true" pageId="16" pageNumber="17">Gabon.</emphasis>
<emphasis bold="true" pageId="16" pageNumber="17">
<normalizedToken originalValue="Ngounié">Ngounie</normalizedToken>
</emphasis>
: along a forestry road of chantier EFT (Exploitaion
<normalizedToken originalValue="Forestière">Forestiere</normalizedToken>
de Tsanba) starting at Ndjemba village on
<normalizedToken originalValue="Fougamou-Lambaréné">Fougamou-Lambarene</normalizedToken>
road, 29 Oct 2009,
<emphasis italics="true" pageId="16" pageNumber="17">Bissiengou, P. 644</emphasis>
(LBV, WAG); CFAD de Rimbunan Hijau, au Sud-Ouest du Parc National de la
<normalizedToken originalValue="Lopé">Lope</normalizedToken>
, 448m, 31 Jan 2009,
<emphasis italics="true" pageId="16" pageNumber="17">Dauby, G.V. 1510</emphasis>
(BRLU);
<emphasis bold="true" pageId="16" pageNumber="17">
<normalizedToken originalValue="Ogooué-Ivindo">Ogooue-Ivindo</normalizedToken>
</emphasis>
: 5km Sud du Petit Okano, 20 Mar 1979,
<emphasis italics="true" pageId="16" pageNumber="17">Florence, J. 1853</emphasis>
(P).
</paragraph>
<paragraph pageId="16" pageNumber="17">
<emphasis bold="true" pageId="16" pageNumber="17">Ghana.</emphasis>
<emphasis bold="true" pageId="16" pageNumber="17">Western Region</emphasis>
: Asientiem, 11 Jul 1912,
<emphasis italics="true" pageId="16" pageNumber="17">Chipp, T.F. 291</emphasis>
(K).
</paragraph>
<paragraph lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis bold="true" pageId="16" pageNumber="17">Ivory Coast.</emphasis>
<emphasis bold="true" pageId="16" pageNumber="17">Abidjan</emphasis>
: km 52 new road Abidjan-Ndouci, c. 10 km E of Sikensi, 19 Jul 1979,
<emphasis italics="true" pageId="16" pageNumber="17">Kruif, A.P.M. de 207</emphasis>
(FHO,WAG); Banco arboretum near Abidjan, 9 Oct 1961,
<emphasis italics="true" pageId="16" pageNumber="17">Wilde, J.J.F.E. de 3126</emphasis>
(K,WAG); Banco Forest Reserve, near Arboretum, along Banco River, 26 Feb 1975,
<emphasis italics="true" pageId="16" pageNumber="17">Koning, J. de 5412</emphasis>
(BR, C, E, G, MA, MO, WAG); Abid
<pageBreakToken pageId="17" pageNumber="18" start="start">jan</pageBreakToken>
, Banco Forest Reserve, Botanic Garden, 26 Feb 1976,
<emphasis italics="true" pageId="17" pageNumber="18">Koning, J. de 6583</emphasis>
(AAU, BR, C, EA, FR, G, GC, K, LMU, MA, MO, P, PRE, SL, WAG);
<normalizedToken originalValue="Adiopodoumé">Adiopodoume</normalizedToken>
, 5 Feb 1961,
<emphasis italics="true" pageId="17" pageNumber="18">Wit, H.C.D. de 9110</emphasis>
(WAG);
<normalizedToken originalValue="forêt">foret</normalizedToken>
du Banco, 20 Apr 1973,
<emphasis italics="true" pageId="17" pageNumber="18">
<normalizedToken originalValue="Aké">Ake</normalizedToken>
Assi, L. 12030
</emphasis>
(G); Arboretum du Banco, 19 Dec 1973,
<emphasis italics="true" pageId="17" pageNumber="18">
<normalizedToken originalValue="Aké">Ake</normalizedToken>
Assi, L. 12295
</emphasis>
(G);
<normalizedToken originalValue="forêt">foret</normalizedToken>
du Banco, 26 Mar 1980,
<emphasis italics="true" pageId="17" pageNumber="18">
<normalizedToken originalValue="Aké">Ake</normalizedToken>
Assi, L. 15146
</emphasis>
(G);
<normalizedToken originalValue="forêt">foret</normalizedToken>
du Banco, 1 Jun 1981,
<emphasis italics="true" pageId="17" pageNumber="18">
<normalizedToken originalValue="Aké">Ake</normalizedToken>
Assi, L. 15900
</emphasis>
(G); Arboretum du Banco, 5 Nov 1984,
<emphasis italics="true" pageId="17" pageNumber="18">
<normalizedToken originalValue="Aké">Ake</normalizedToken>
Assi, L. 16772
</emphasis>
(B, G); Banco, 10 Mar 1932,
<emphasis italics="true" pageId="17" pageNumber="18">
<normalizedToken originalValue="Aubréville">Aubreville</normalizedToken>
, A. SF 1331
</emphasis>
(P); Le Banco, Jun 1932,
<emphasis italics="true" pageId="17" pageNumber="18">
<normalizedToken originalValue="Aubréville">Aubreville</normalizedToken>
, A. SF 1341
</emphasis>
(P);
<emphasis bold="true" pageId="17" pageNumber="18">
<normalizedToken originalValue="Adzopé">Adzope</normalizedToken>
</emphasis>
: On border of
<normalizedToken originalValue="Comoé">Comoe</normalizedToken>
river, c. 15 km NW of Mbasso, c. 60 km NE of
<normalizedToken originalValue="Adzopé">Adzope</normalizedToken>
, 27 Jul 1963,
<emphasis italics="true" pageId="17" pageNumber="18">Wilde, W.J.J.O. de 570</emphasis>
(BR, K, P, WAG, Z);
<emphasis bold="true" pageId="17" pageNumber="18">
<normalizedToken originalValue="San-Pédro">San-Pedro</normalizedToken>
</emphasis>
:
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="Classée">Classee</normalizedToken>
Monogaga, just north of Sassandra - San Pedro road, 24 Mar 2000,
<emphasis italics="true" pageId="17" pageNumber="18">Jongkind, C.C.H. 4707</emphasis>
(MO, U, WAG); Classified forest of Monogaga, 26 Apr 1997, Breteler, F.J. 13761 (WAG);
<emphasis bold="true" pageId="17" pageNumber="18">Sassandra</emphasis>
: 79 km NNE of Sassandra, Lagako-Tokpeko, 7 May 1975,
<emphasis italics="true" pageId="17" pageNumber="18">Burg, W.J. van der 131</emphasis>
(WAG); 61 km N of Sassandra, W of Niapidou, 19 Jan 1959,
<emphasis italics="true" pageId="17" pageNumber="18">Leeuwenberg, A.J.M. 2492</emphasis>
(P, WAG); 61 km N of Sassandra, W of Niapidou, 21 Feb 1959,
<emphasis italics="true" pageId="17" pageNumber="18">Leeuwenberg, A.J.M. 2785</emphasis>
(P, WAG); 44 km Lakota-Sassandra, 28 Oct 1968,
<emphasis italics="true" pageId="17" pageNumber="18">Breteler, F.J. 5811</emphasis>
(BR, EA, HBG, K, LG, LISC, MA, MO, P, PRE, SRGH, U, WAG);
<emphasis bold="true" pageId="17" pageNumber="18">Tabou:</emphasis>
P.N.
<normalizedToken originalValue="Taï">Tai</normalizedToken>
, environ 0.5 Km
<normalizedToken originalValue="à">a</normalizedToken>
<normalizedToken originalValue="l’Est">l'Est</normalizedToken>
de la Station CRE,
<geoCoordinate degrees="5" direction="north" minutes="51" orientation="latitude" precision="925" value="5.85">5°51'N</geoCoordinate>
,
<geoCoordinate degrees="7" direction="west" minutes="21" orientation="longitude" precision="925" value="-7.35">7°21'W</geoCoordinate>
, Feb 1999,
<emphasis italics="true" pageId="17" pageNumber="18">Menzies, A. 107</emphasis>
(G); Entre Djiroutou et le mont
<normalizedToken originalValue="Niénoukoué">Nienoukoue</normalizedToken>
. Guiroutou, 50m au Sud du layon
<normalizedToken originalValue="‘Hana’">'Hana'</normalizedToken>
;
<normalizedToken originalValue="à">a</normalizedToken>
3.5 Km du campement
<normalizedToken originalValue="écotouristique">ecotouristique</normalizedToken>
, Mar 1999,
<emphasis italics="true" pageId="17" pageNumber="18">Menzies, A. 459</emphasis>
(G); P.N.
<normalizedToken originalValue="Taï">Tai</normalizedToken>
, station
<normalizedToken originalValue="d’écologie">d'ecologie</normalizedToken>
, 12 Jul 1996,
<emphasis italics="true" pageId="17" pageNumber="18">Chatelain, C. 1358</emphasis>
(CSRS).
</paragraph>
<paragraph pageId="17" pageNumber="18">
<emphasis bold="true" pageId="17" pageNumber="18">Liberia.</emphasis>
<emphasis bold="true" pageId="17" pageNumber="18">Grand Gedeh</emphasis>
: Putu Hills, E ridge, Mt. Jideh, 254m, 2 Dec 2010,
<emphasis italics="true" pageId="17" pageNumber="18">Putu Botanic Team EP 1307</emphasis>
(WAG).
</paragraph>
<paragraph pageId="17" pageNumber="18">
<emphasis bold="true" pageId="17" pageNumber="18">Nigeria.</emphasis>
<emphasis bold="true" pageId="17" pageNumber="18">Cross River State</emphasis>
: Calabar, Eket Distr, Stubbs Greek F.R. near Unyene, 10 Jan 1959,
<emphasis italics="true" pageId="17" pageNumber="18">Keay, R.W.J. 37719 (</emphasis>
K);
<emphasis bold="true" pageId="17" pageNumber="18">Lagos State</emphasis>
: Lagos Botanical Station, Atigere, Feb 1893,
<emphasis italics="true" pageId="17" pageNumber="18">Millen, H. 15</emphasis>
(K); Eba, 1931,
<emphasis italics="true" pageId="17" pageNumber="18">Kennedy, J.D. 1711</emphasis>
(FHO); road leading Murtala Muhamed botanical gardens, 26 Nov 1994,
<emphasis italics="true" pageId="17" pageNumber="18">Daramola, B.O. 94/ 581</emphasis>
(F, MO).
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/10563" pageId="17" pageNumber="18" start="Figure 10" startId="F10">
<paragraph pageId="17" pageNumber="18">
<emphasis bold="true" pageId="17" pageNumber="18">Figure 10.</emphasis>
Distribution of
<taxonomicName class="Magnoliopsida" family="Annonaceae" genus="Uvariastrum" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="" order="Magnoliales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="insculptum">
<emphasis italics="true" pageId="17" pageNumber="18">Uvariastrum insculptum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<paragraph id="874F536DD1FA20B89B10CDE51A805EAF" pageId="29" pageNumber="30">Taxon classification Animalia Lepidoptera Depressariidae</paragraph>
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<paragraph id="98A9E7B2A41A09D74D52606C913D4C5E" pageId="29" pageNumber="30">
<taxonomicName id="6B0CC7C2FFE43CAE9A2462DDE20E6829" LSID="http://zoobank.org/247A9873-10E1-478A-8D89-2FE77E58D686" authority="Phillips" class="Insecta" family="Elachistidae" genus="Ethmia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethmia johnpringlei" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="johnpringlei">Ethmia johnpringlei Phillips</taxonomicName>
<taxonomicNameLabel id="55AC02816E9743460684270D20C9B391" pageId="29" pageNumber="30">sp. n.</taxonomicNameLabel>
Figures 27, 72, 117, 160
</paragraph>
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<subSubSection id="8DCE1F1D88DFE3BC4B16EEA76241319E" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="28308408463024D6A891A544C29E3D1D" pageId="29" pageNumber="30">Diagnosis.</paragraph>
<paragraph id="1333C432C0C8DF8C183C84FF277249DD" pageId="29" pageNumber="30">This species can be distinguished by its smaller size and the presence of three curved large distal setae on the valva of male genitalia.</paragraph>
</subSubSection>
<subSubSection id="72FE139C77FC97D7D0169DE520AC7462" pageId="29" pageNumber="30" type="description">
<paragraph id="AEAF69AF61DE04FF0B3E070BEF0EFB99" pageId="29" pageNumber="30">Description.</paragraph>
<paragraph id="61381055E7CD7FF4F88DC2F86025CF21" pageId="29" pageNumber="30">
Male: FW length 7.1-7.8 mm (n = 3). Head: Labial palpus elongated surpassing base of antenna, grayish with black rings in II segment middle and subapical, a wide black ring subapical in III segment; front and crown grayish, occipital black tuft at mid-dorsum. Thorax: Grayish. FW ground color grayish, with a series of elongated dark markings, the most conspicuous an oblique line originating at costa before middle reaching a longitudinal line that goes from middle to near termen; three irregular black dots on posterior half: one at 0.20
<normalizedToken id="A174561308ADC08D4F0F1E7D8CDC9029" originalValue="×">x</normalizedToken>
from base, one close to midline and third near tornus. HW ground color white, costa simple. Abdomen: Brownish, genital scaling pale ochreous. Genitalia (Fig. 72) with uncus lacking; valva emarginate bellow apex, with apical
<normalizedToken id="7CDD4AEC3AD3BA633E498CA518AD1807" originalValue="“plume”">&quot;plume&quot;</normalizedToken>
, and distally three curved, strong, large setae.
</paragraph>
<paragraph id="82F77CEF730304C6C461D468C4A37F25" pageId="29" pageNumber="30">Female: FW length 7.9-8.3 mm (n = 3). Head and thorax: As described for male. Abdomen: Genitalia (Fig. 117) with sterigma simple; base of ductus sclerotized; antrum membranous; signum a short fold with irregular margin.</paragraph>
</subSubSection>
<subSubSection id="60F990226E10B426F2A5405B863C1ACF" pageId="29" pageNumber="30" type="holotype">
<paragraph id="36FD996FE04A02007E01FA7C21F2CA48" pageId="29" pageNumber="30">Holotype.</paragraph>
<paragraph id="811FBBA70837FE2E714BF8DD8BCE3152" pageId="29" pageNumber="30">
Male: 11-SRNP-104923, DNA Barcoded, Costa Rica: Guanacaste: ACG, Sector Maritza, Manta Mecate 587 m, 1.x.2011, R. Franco &amp; S.
<normalizedToken id="B0F716985DD709DFE18A3E88FD0E21E3" originalValue="Ríos">Rios</normalizedToken>
. Deposited in INBio. Paratypes: Costa Rica: Guanacaste: ACG: Sector Maritza, Manta Mecate 587 m, 1.x.2011, R. Franco &amp; S.
<normalizedToken id="79A65839E397866E7F2914C1FBE8C6AF" originalValue="Ríos">Rios</normalizedToken>
. Est. Maritza, Lado Oeste del
<normalizedToken id="565DA36337087405D4676B8509550AEF" originalValue="Volcán">Volcan</normalizedToken>
Orosi, 600 m, 2M 13.v.1988 Janzen and Hallwachs, 10M 11F 1.viii.1990, II curso
<normalizedToken id="7FF24760C6C770B1196A82BF02A4E4B3" originalValue="Parataxónomos">Parataxonomos</normalizedToken>
, 17M 27F 1.vii.1990, I Curso
<taxonomicName id="5A3E8F2D99E26431F01EEC07411CE773" above-genus="Microlepidoptera" genus="Microlepidoptera" lsidName="Microlepidoptera" pageId="29" pageNumber="30" rank="above-genus">Microlepidoptera</taxonomicName>
. 4 km W. Sta. Cecilia, 300 m 1M 17.iv.1983, Janzen &amp; Hallwachs (BMNH, INBio, EME, USNM).
</paragraph>
</subSubSection>
<subSubSection id="B6099E006FFF2E39C0E40DC4CBCA71FB" pageId="29" pageNumber="30" type="distribution and biology">
<paragraph id="EFA26FFA34A218881C663AD09C1C63E7" pageId="29" pageNumber="30">Distribution and biology.</paragraph>
<paragraph id="0BC865F32BC55A53023797FE6757B757" pageId="29" pageNumber="30">
<taxonomicName id="41A7483504AAFB8E4544687EA1D95274" class="Insecta" family="Elachistidae" genus="Ethmia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethmia johnpringlei" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="johnpringlei">Ethmia johnpringlei</taxonomicName>
has been collected in northern Costa Rica on the western sides of Cordillera
<normalizedToken id="A3453266E9B769CAEA31B201898F38EB" originalValue="Volcánica">Volcanica</normalizedToken>
de Guanacaste, from 300 to 600 m in the intersection between ACG dry forest and rain forest. The food plant and immature stages are unknown.
</paragraph>
</subSubSection>
<subSubSection id="F4171785960CE13F84E08592A4EE6D5A" pageId="30" pageNumber="31" type="etymology">
<paragraph id="19137F5F5A4207FC97195F56DDDD8233" pageId="30" pageNumber="31">
<pageBreakToken id="88A67B64E9BD870C246A0FC77FA32493" pageId="30" pageNumber="31" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph id="D4057EE62FED27A592FAD922110B5113" pageId="30" pageNumber="31">
<taxonomicName id="92B8609C4A4EFFD646FC6666FFE0E938" class="Insecta" family="Elachistidae" genus="Ethmia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethmia johnpringlei" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="johnpringlei">Ethmia johnpringlei</taxonomicName>
is named in honor of John Pringle of Stanford University, the father of Rob Pringle of Princeton University, and a member of the very small and select group of academic faculty who have supported ACG land purchase for permanent conservation.
</paragraph>
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</treatment>
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<mods:affiliation>Fisheries and Marine Science Faculty, Pattimura University, Ambon, Indonesia &amp; Maritime and Marine Science Center of Excellence, Pattimura University, Ambon, Indonesia</mods:affiliation>
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<taxonomicName LSID="B60CF750-DAA2-519A-8AB0-277B0391B762" authority="Randall et Eschmeyer, 2002" authorityName="Randall et Eschmeyer" authorityYear="2002" class="Actinopterygii" family="Scorpaenidae" genus="Scorpaenopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpaenopsis obtusa" order="Scorpaeniformes" pageId="0" pageNumber="245" phylum="Chordata" rank="species" species="obtusa">Scorpaenopsis obtusa Randall et Eschmeyer, 2002</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fresh specimen photograph of Scorpaenopsis obtusa, MZB. 26532, 91.1 mm SL, collected from Alor Island, Indonesia." figureDoi="10.3897/aiep.52.91145.figure1" httpUri="https://binary.pensoft.net/fig/770277" pageId="0" pageNumber="245">English common name: shortsnout scorpionfish (Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distributional records of Scorpaenopsis obtusa, based on previous studies (circles) and the presently reported study (star). Closed and open symbols indicate specimen- and underwater photograph-based records, respectively." figureDoi="10.3897/aiep.52.91145.figure2" httpUri="https://binary.pensoft.net/fig/770278" pageId="0" pageNumber="245">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Inner surface of pectoral fin (fresh condition) (A), head (showing line from posterior margin of orbit to posteroventral tip of retroarticular) (B), and caudal fin (C) of Scorpaenopsis obtusa, MZB. 26532, 91.1 mm SL, collected from Alor Island, Indonesia." figureDoi="10.3897/aiep.52.91145.figure3" httpUri="https://binary.pensoft.net/fig/770279" pageId="0" pageNumber="245">, 3)</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="245" type="materials_examined">
<paragraph pageId="0" pageNumber="245">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="245">
<materialsCitation collectingDate="2022-02-26" collectingMethod="hand net" collectorName="K. Wibowo" country="Indonesia" county="Alor Island" elevation="5" location="Ampera" municipality="Alor Barat Laut" specimenCount="1">
MZB.26532,
<specimenCount type="generic">1 specimen</specimenCount>
,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="9.11" unit="mm" value="91.1">91.1 mm</quantity>
SL,
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,
<collectingMunicipality>Alor Barat Laut</collectingMunicipality>
,
<collectingCounty>Alor Island</collectingCounty>
,
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
, 8°16ʹ03.7ʹʹS, 124°25ʹ33.1ʹʹE,
<elevation metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">
<quantity metricMagnitude="0" metricUnit="m" metricValue="5.0" unit="m" value="5.0">5 m</quantity>
depth
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,
<collectingMethod>hand net</collectingMethod>
,
<collectorName>K. Wibowo</collectorName>
,
<collectingDate value="2022-02-26">26 February 2022</collectingDate>
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.
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<subSubSection pageId="0" pageNumber="245" type="description">
<paragraph pageId="0" pageNumber="245">Description.</paragraph>
<paragraph pageId="0" pageNumber="245">Counts and proportional measurements [given below as percentages of SL and HL (for head characters; presented in parenthesis)] of presently reported specimen (91.1 mm SL) were as follows: dorsal fin with 12 spines and 9 soft rays; pectoral fin with 1 uppermost unbranched ray, 6 branched rays, 12 lower unbranched rays, total 19 rays; longitudinal scale rows 38; pored lateral-line scales 20; scales above lateral line 5, below 12; gill rakers on upper limb of first gill arch 4, lower limb 9, total 13; body depth 44.5% of SL; body width 30.2; head length 45.6; snout length 11.8 (26.0% of HL); orbit diameter 10.3 (22.7); interorbital width at posterior end of preocular spine bases 11.1 (24.3); caudal-peduncle depth 12.0; caudal-peduncle length 15.7; upper-jaw length 25.9 (57.0); pre-dorsal-fin length 38.5; pre-anal-fin length 71.5; pre-pelvic-fin length 39.3; first dorsal-fin spine length 7.8; second dorsal-fin spine length 11.9; third dorsal-fin spine broken; fourth dorsal-fin spine length 15.8; eleventh dorsal-fin spine length 10.8; twelfth dorsal-fin spine length 12.9; longest dorsal-fin soft ray length 19.2 (third ray); first anal-fin spine length 9.4; second anal-fin spine length 17.5; third anal-fin spine length 15.6; longest anal-fin soft ray length 23.0 (second ray); caudal-fin length 26.9; pectoral-fin length 39.0; pelvic-fin spine length 19.3; longest pelvic-fin soft ray length 29.2 (second ray); head width 21.5 (47.3); postorbital length 27.3 (59.9); maxillary depth 7.6 (16.6).</paragraph>
<paragraph pageId="0" pageNumber="245">Body wide anteriorly, progressively compressed posteriorly. Mouth large, oblique, positioned at angle of about 60° to horizontal axis of head and body. Posterior margin of maxilla extending beyond line from posterior margin of orbit to edge of retroarticular. Vomerine plate forming V-shaped patch, with rudimentary teeth. Palatine teeth absent. Nasal spine with 3 spinous points. Ascending process of premaxilla intruding into interorbital space, its posterior margin level with anterior margin of pupil. Median interorbital ridge absent. Interorbital ridges distinct; space between interorbital ridges shallow and broad, its width about half of orbit diameter. Preocular and supraocular spines small, directed upward. Postocular spine large, with 3 points, broadly joined to tympanic spine at base. Tympanic with 2 spinous points, located between postocular spines. Coronal spine absent. Occipital pit deep, its width greater than length. Parietal and nuchal spines about equal size, joined at base. Sphenotic with 2 small spines. Postorbital with 2 small spines. Pterotic spine simple, larger than supraocular spine. Upper posttemporal spine larger than lower spine. Space between parietal, nuchal, pterotic, and lower posttemporal spines with two small spines. Supracleithral spine asymmetric, with 3 and 2 pointed spines on left and right sides, respectively. Cleithral spine flattened, strongly pointed. Anterodorsal lacrimal spine present. Single small spines on middle and posterior end of lacrimal ridge. Anterior lacrimal spine directed anteriorly, its tip reaching dorsal margin of upper lip; two additional small points on posterior margin. Posterior lacrimal spine directed ventrally, larger than anterior spine, its tip not reaching dorsal margin of upper lip; an additional spine present on anterior margin; posterior lacrimal spine associated with short fimbriate flap. Lateral lacrimal spine present, its size approximately equal to anterodorsal lacrimal spine. Suborbital ridge with 3 spines. Suborbital pit present. Preopercle with 5 spines; uppermost spine largest, with supplemental preopercular spine on base; second spine sharp; third to fifth spines blunt. Upper opercular spine with 2 points; lower opercular spine simple, with median ridge.</paragraph>
<paragraph pageId="0" pageNumber="245">Lateral surface of body covered with weak ctenoid scales, becoming cycloid ventrally. Exposed cycloid scales covering pectoral-fin base. Cycloid scales embedded in thin skin covering anteroventral surface of body. Body scales not extending onto fin rays or membranes, except basally on seven uppermost pectoral fin rays. Lateral line complete, first two scales with spine-like projection at end of sensory tube. No scales on head.</paragraph>
<paragraph pageId="0" pageNumber="245">Numerous tiny papillae scattered on dorsal and lateral surfaces of head, including snout, interorbital space, outer margin of eye membrane, preopercle, and opercle. No papillae on occipital pit. Tentacle on upper posterior edge of low membranous tube associated with anterior nostril. Fleshy tentacle associated with posterior lacrimal spine. Several short, slender tentacles on lateral surface of maxilla. Broad, skin flap associated with each of third to fifth preopercular spines. Many small, round flaps on ventral surface of head and anteroventral surface of body. Many slender tentacles associated with posterior margin of opercle, pored lateral scales scattered on lateral and dorsal surfaces of body, and pectoral and caudal fins.</paragraph>
<paragraph pageId="0" pageNumber="245">Origin of first dorsal-fin spine above supracleithral spine. Posterior margin of opercular membrane level with posterior margin of third dorsal-fin spine base. Posterior tip of pectoral fin extending beyond third anal-fin spine base. Posterior tip of pelvic fin extending slightly beyond anus when depressed. Origin of last dorsal-fin spine just above origin of first anal-fin spine.</paragraph>
<paragraph pageId="0" pageNumber="245">
<emphasis bold="true" italics="true" pageId="0" pageNumber="245">Color when fresh</emphasis>
(Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Fresh specimen photograph of Scorpaenopsis obtusa, MZB. 26532, 91.1 mm SL, collected from Alor Island, Indonesia." figureDoi="10.3897/aiep.52.91145.figure1" httpUri="https://binary.pensoft.net/fig/770277" pageId="0" pageNumber="245">1</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Inner surface of pectoral fin (fresh condition) (A), head (showing line from posterior margin of orbit to posteroventral tip of retroarticular) (B), and caudal fin (C) of Scorpaenopsis obtusa, MZB. 26532, 91.1 mm SL, collected from Alor Island, Indonesia." figureDoi="10.3897/aiep.52.91145.figure3" httpUri="https://binary.pensoft.net/fig/770279" pageId="0" pageNumber="245">3A</figureCitation>
). Head and body mottled dark reddish-brown to white. Dorsal fin whitish to reddish-brown, distinct black blotch on base of spinous portion between fourth and eighth spines. Pectoral fin outer surface pinkish-white, blackish basally, with blackish band near tip; inner surface orange, with broad black blotch basally on each of uppermost four and lowermost seven rays, black submarginal band (progressively less distinct on lower unbranched rays), axil and base of membrane of fifth to fourteenth rays white with small black spots. Pelvic fin black, with white distal margin. Anal fin dark reddish with numerous tiny spots and white posterior margin. Caudal fin whitish, with broad pinkish band.
</paragraph>
<caption doi="10.3897/aiep.52.91145.figure2" httpUri="https://binary.pensoft.net/fig/770278" pageId="0" pageNumber="245" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="245">
<emphasis bold="true" pageId="0" pageNumber="245">Figure 2.</emphasis>
Distributional records of
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<emphasis italics="true" pageId="0" pageNumber="245">Scorpaenopsis obtusa</emphasis>
</taxonomicName>
, based on previous studies (circles) and the presently reported study (star). Closed and open symbols indicate specimen- and underwater photograph-based records, respectively.
</paragraph>
</caption>
<caption doi="10.3897/aiep.52.91145.figure3" httpUri="https://binary.pensoft.net/fig/770279" pageId="0" pageNumber="245" start="Figure 3" startId="F3">
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<emphasis bold="true" pageId="0" pageNumber="245">Figure 3.</emphasis>
Inner surface of pectoral fin (fresh condition) (
<emphasis bold="true" pageId="0" pageNumber="245">A</emphasis>
), head (showing line from posterior margin of orbit to posteroventral tip of retroarticular) (
<emphasis bold="true" pageId="0" pageNumber="245">B</emphasis>
), and caudal fin (
<emphasis bold="true" pageId="0" pageNumber="245">C</emphasis>
) of
<taxonomicName authorityName="Randall et Eschmeyer" authorityYear="2002" class="Actinopterygii" family="Scorpaenidae" genus="Scorpaenopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scorpaenopsis obtusa" order="Scorpaeniformes" pageId="0" pageNumber="245" phylum="Chordata" rank="species" species="obtusa">
<emphasis italics="true" pageId="0" pageNumber="245">Scorpaenopsis obtusa</emphasis>
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, MZB.26532, 91.1 mm SL, collected from Alor Island, Indonesia.
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Subfamily
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<taxonomicName id="D9FD5E80AFC0B30633A71C82F9892E60" class="Magnoliopsida" family="Malvaceae" genus="Carabici" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carabici" order="Malvales" pageId="69" pageNumber="70" phylum="Tracheophyta" rank="genus">Carabici</taxonomicName>
Latreille, 1802: 80 [stem: Carab-]. Type genus:
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Linnaeus, 1758.
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<taxonomicName id="9083DE36D4D4A983CEA78A49243FA7C8" ID-CoL="46KYR" class="Arachnida" family="Brachychthoniidae" genus="Neoliochthonius" kingdom="Animalia" order="Sarcoptiformes" pageNumber="87" phylum="Arthropoda" rank="species" species="piluliferus">Neoliochthonius piluliferus</taxonomicName>
(Forsslund, 1942) [44e,f]
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Syn., Tax.:
<taxonomicName id="1B00923C61D1164E3052644508585E7C" class="Arachnida" family="Brachychthoniidae" genus="Liochthonius" kingdom="Animalia" order="Sarcoptiformes" pageNumber="87" phylum="Arthropoda" rank="species" species="piluliferus">Liochthonius piluliferus Forsslund</taxonomicName>
, 1942: Niedbala 1972a (B).
<taxonomicName id="E3BBF06D78A2E43CAC12CBC350125B98" class="Arachnida" family="Brachychthoniidae" genus="Brachychthonius" kingdom="Animalia" order="Sarcoptiformes" pageNumber="87" phylum="Arthropoda" rank="species" species="piluliferus" species.outerRound="1">Brachychthonius p.</taxonomicName>
: Evans 1952b (B).
<taxonomicName id="738BAE8CABC963E93FBCD96386EE18B8" class="Arachnida" family="Chthoniidae" genus="Paraliochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageNumber="87" phylum="Arthropoda" rank="species" species="piluliferus" species.outerRound="1">Paraliochthonius p.</taxonomicName>
: Moritz 1976a (B); Balogh &amp; Mahunka 1983 (B).
<taxonomicName id="0EFBA7A3C501BBCAEB1363FDA658C856" class="Arachnida" family="Brachychthoniidae" genus="Neoliochthonius" kingdom="Animalia" order="Sarcoptiformes" pageNumber="87" phylum="Arthropoda" rank="species" species="piluliferus" species.outerRound="1">Neoliochthonius p.</taxonomicName>
: Lee 1982; Schatz 2004a.
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<paragraph id="799CFA64102F1CBE1FDE6E0E44885CE7" pageNumber="87">
<normalizedToken id="A52A337CB2C2D106638786996D2C57A7" originalValue="Ökologie">Oekologie</normalizedToken>
:
<normalizedToken id="09F217928B5EB6C91809CBF54AFE0426" originalValue="Waldböden">Waldboeden</normalizedToken>
.
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<paragraph id="883655751725C1ED4BF403722871C6E7" pageNumber="87">Verbreitung: Holarktis.</paragraph>
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<taxonomicName id="4622CAF93F42C2240928CD33254F6EBA" authority="Lindqvist, 1959" authorityName="Lindqvist" authorityYear="1959" class="Insecta" family="Tenthredinidae" genus="Pristiphora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristiphora tenuiserra" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="tenuiserra">Pristiphora tenuiserra (Lindqvist, 1959)</taxonomicName>
Figs 119, 123, 173, 302
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<taxonomicName id="0AB9A1B0ECCB2AFF9497556448A075EA" class="Insecta" family="Tenthredinidae" genus="Micronematus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Micronematus tenuiserra" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="tenuiserra">Micronematus tenuiserra</taxonomicName>
Lindqvist, 1959: 72. Holotype ♀ in MZLU (DEI-GISHym31572), examined. Type locality: Knivsta, Uppland, Sweden.
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<paragraph id="170CAFD16A192F47D5BA26C6E9B8997E" pageId="94" pageNumber="95">Similar species.</paragraph>
<paragraph id="161F5DB27663327EC743FB946E0B7820" pageId="94" pageNumber="95">Females can be distinguished from other species by the combination of the structure of valvula 3 (long and narrow and practically without scopa; Figs 119, 123) and (at least usually) relatively dark coloration (head dorsally black or with small pale spots, thorax black, abdomen at least dorsally black). Males are best recognised by examining penis valves (see the Key).</paragraph>
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<subSubSection id="DB92E31894405B23B0FAC17A77025F8B" pageId="94" pageNumber="95" type="genetic data">
<paragraph id="6737254ADB9CAC3B752277467F8CA40C" pageId="94" pageNumber="95">Genetic data.</paragraph>
<paragraph id="561291AA4590E8D478A01C89FB6B7730" pageId="94" pageNumber="95">
Based on COI barcode sequences (only one specimen in BOLD),
<taxonomicName id="B069482740EEE0114C8EF85EBD5F833D" lsidName="P. tenuiserra" pageId="94" pageNumber="95" rank="species" species="tenuiserra">P. tenuiserra</taxonomicName>
forms its own BIN cluster (BOLD:ADD2616) (Fig. 5). The nearest neighbour to BOLD:ADD2616, diverging by a minimum of 5.22%, is BOLD:ACO1401 (
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). No nuclear data are available.
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<paragraph id="19B02C31C5FA7AC421ADD83C899BB1A3" pageId="94" pageNumber="95">Host plants.</paragraph>
<paragraph id="68E2ED1D15D16DAC229F477112B8DDB1" pageId="94" pageNumber="95">Unknown, but the structure of valvula 3 suggests that the species is probably a conifer-feeder.</paragraph>
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<subSubSection id="BFF2F1A15C9759CF989777A97519BA1E" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="3AC3630090470FB918A90EBE4F339079" pageId="94" pageNumber="95">Distribution and material examined.</paragraph>
<paragraph id="92B72666C6858BD76DF24BF24362CDBA" pageId="94" pageNumber="95">West Palaearctic. Specimens studied are from Finland, Germany, Poland, and Sweden.</paragraph>
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<paragraph id="2013F7A75B734A2304ABDD418141B98C" pageId="99" pageNumber="100">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
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<taxonomicName id="4F63619876225922CA58CEAF6C950315" ID-CoL="SDCL" LSID="http://species-id.net/wiki/Cephaloleia_convexifrons" authority="Pic, 1923" authorityName="Pic" authorityYear="1923" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia convexifrons" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="convexifrons">Cephaloleia convexifrons Pic, 1923</taxonomicName>
Fig. 105
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<paragraph id="37055B92F9120E89CA7EE8BFFB9DDD89" pageId="99" pageNumber="100">
<taxonomicName id="41D1F75545C3FBAC79826E723089A2B4" class="Insecta" family="Chrysomelidae" genus="Cephalolia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cephalolia convexifrons" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="convexifrons">Cephalolia convexifrons</taxonomicName>
<bibRefCitation id="3F52116F68B8DE71ABA55253673C7BAD" author="Pic, M" journalOrPublisher="Melanges Exotico-Entomologiques" pageId="291" pageNumber="292" pagination="1 - 32" title="Nouveautes diverses." volume="38" year="1923">Pic 1923</bibRefCitation>
: 9.
</paragraph>
<paragraph id="05C41274E246E407D1A25605B8A91F29" pageId="99" pageNumber="100">
<taxonomicName id="F553F0C6DC110720DB651B1BEA07FABC" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia convexifrons" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="convexifrons">Cephaloleia convexifrons</taxonomicName>
Pic.
<bibRefCitation id="16AFB02E76DCFE4C53618EF888FF4817" author="Uhmann, E" journalOrPublisher="Beitraege zur Entomologie" pageId="296" pageNumber="297" pagination="35 - 42" title="Sechs neue Hispinen aus Suedamerika (Coleoptera: Chrysomelidae). 181. Beitrag zur Kenntnis der Hispinae." volume="7" year="1957 b">Uhmann 1957b</bibRefCitation>
: 17 (catalog),
<bibRefCitation id="505A39456648AB603314550CA04431DE" author="Uhmann, E" journalOrPublisher="W. Junk, Gravenhage" pageId="296" pageNumber="297" pagination="399 - 490" title="Coleopterorum Catalogus. Supplementa. Chrysomelidae: Hispinae, Corrigenda et Addenda" year="1964 a">1964a</bibRefCitation>
: 403 (catalog);
<bibRefCitation id="C7A925BD3F8B78F425C26C5809A6150E" author="Descarpentries, A" journalOrPublisher="Bulletin du Museum d'Histoire Naturelle Paris" pageId="286" pageNumber="287" pagination="137 - 154" title="Les types de la collection M. Pic. 1. Chrysomelinae et Hispinae du Nouveau-Monde." volume="31" year="1959">Descarpentries and Villiers 1959</bibRefCitation>
a: 139 (types).
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<paragraph id="9E8AA83D913F274CE5C3FD10E8012FDB" pageId="99" pageNumber="100">Description.</paragraph>
<paragraph id="B3BA7CE7CDB81B96C5DFD2DDBEFDBDF1" lastPageId="100" lastPageNumber="101" pageId="99" pageNumber="100">
Elongate; subconvex; antennae, head, and scutellum black; pronotum yellowish with broad medial longitudinal vitta from base to apex; elytra yellowish with broad black sutural vitta which narrows to apex and narrow black vitta from humerus to near apex; legs yellowish with darker joints and tarsi. Head: small; vertex densely punctate, medial sulcus absent; frons not projecting; not depressed between eyes. Antenna: reaches to humerus; slender; cylindrical; antennomere 1 short; 2 2
<normalizedToken id="621BF8865261571F02234E9AF890BE1E" originalValue="×">x</normalizedToken>
length of 1; 3 subequal in length to 2; 4-10 subequal in length, each shorter than 3; 11 2
<normalizedToken id="F0B4E5D25526CBB5645494E7B645ED4A" originalValue="×">x</normalizedToken>
length of 10, longest, acutely pointed at apex; 1-2 punctate; 3-11 setose. Pronotum: slightly transverse; lateral margin straight, slightly divergent for basal 4/5 then rounding to anterior angle, narrowly margined; anterior angle rounded, not produced; posterior angle acute; anterior margin emarginate behind head; disc subconvex; surface irregularly punctate; transverse basal impression present medially; pronotal length 0.8-1.0 mm;
<pageBreakToken id="35B6CC84CE30E4F33767ED056E648D40" pageId="100" pageNumber="101" start="start">pronotal</pageBreakToken>
width 1.7-1.9 mm. Scutellum: broadly pentagonal; impunctate. Elytron: lateral margin straight, slightly narrowing apically, smooth, margined; apex rounded; minute tooth present in sutural angle; humerus rounded, slightly produced; shallowly punctate-striate; pygidium exposed, punctate; elytral length 4.3-4.7 mm; elytral width 2.0-2.1 mm. Venter: pro-, meso-, and metasterna punctate; abdominal sterna punctate, each puncture with pale seta; suture between sterna 1 and 2 obsolete medially; last sternite with apical margin broadly emarginate medially in male, sinuate in female. Leg: slender; punctate; femur and tibia with pale seta in each puncture; tibia with fringe of setae on inner margin of apex. Total length: 5.9-6.1 mm.
</paragraph>
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<paragraph id="CAE5FBFDD7533E4FA3C4FB30F8BFEAB0" pageId="100" pageNumber="101">Diagnosis.</paragraph>
<paragraph id="E0EB99957D153D6F1643453CEC05772A" pageId="100" pageNumber="101">
This species is similar to
<taxonomicName id="FAF05E827ABBDE0C440684B60646B109" class="Insecta" family="Chrysomelidae" genus="Cephaloleia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cephaloleia chica" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="chica">Cephaloleia chica</taxonomicName>
sp. n. It can be distinguished by the larger size and by the medial longitudinal vitta on the pronotum.
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<paragraph id="2A99B9355F0C3F93B97FA503681EBC4C" pageId="100" pageNumber="101">Distribution.</paragraph>
<paragraph id="F10F22A91F47B5B5CDCCADA0250033A2" pageId="100" pageNumber="101">Bolivia, Peru.</paragraph>
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<subSubSection id="1C55C25BEBDE2686F2AF6AE40D01885A" pageId="100" pageNumber="101" type="type material examined">
<paragraph id="76711BD5E4AF3BD13D1A78F70EE51CF4" pageId="100" pageNumber="101">Type material examined.</paragraph>
<paragraph id="33176A34AF2A5B383D4DA2A2E9F242AA" pageId="100" pageNumber="101">
Holotype: Bolivie, Cochabamba, Germain [green printed label]/
<taxonomicName id="B82F5CD599510EDD8E9898C311D33F87" class="Insecta" family="Chrysomelidae" genus="Cephalolia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cephalolia convexifrons" order="Coleoptera" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="convexifrons">Cephalolia convexifrons</taxonomicName>
m [handwritten label]/ convexifrons Pic (1923) [handwritten label]/ type [white printed label with red border]/ Museum Paris Coll. \M. Pic [blue printed label]/ Holotype [red printed label]/ MNHN EC 2652 [printed label] (MNHN).
</paragraph>
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<subSubSection id="E21998D8BFB42C262555B51A7FDE4109" pageId="100" pageNumber="101" type="specimens examined">
<paragraph id="18A1DC70DB50658950164567B6DE8863" pageId="100" pageNumber="101">Specimens examined.</paragraph>
<paragraph id="C9A6E1119C753475A10A1BDA7E485D4C" pageId="100" pageNumber="101">No label data (USNM). Bolivia: Cochambamba- November 1953 (USNM). Peru: Huanuco- Tingo Maria region, 15-24 June 1937 (SEMC); Quiapicanchia- Cuzco, Quincemil, 11 June 1976 (USNM). Madre de Dios- Tambopata Wildlife Res., 30 km SW Pto Maldonado, 290 m, 15-30 November 1982 (USNM). Total: 5.</paragraph>
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<mods:namePart>Zahiri, Reza</mods:namePart>
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<mods:namePart>Rajaei, Hossein</mods:namePart>
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<mods:namePart>Husemann, Martin</mods:namePart>
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74.
<taxonomicName LSID="B60DFB78-8E8E-5E98-8A79-59D1B50BD512" authority="Bytinski-Salz" authorityName="Bytinski-Salz" class="Insecta" family="Hepialidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Mesembrynus) purpuralis subsp. v. var. asupinensis" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="variety" species="purpuralis" subGenus="Mesembrynus" subSpecies="v." variety="asupinensis">Zygaena (Mesembrynus) purpuralis v. asupinensis Bytinski-Salz</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="39" type="materials_examined">
<paragraph pageId="0" pageNumber="39">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="39">
<materialsCitation accessionNumber="ZMH 61536, ZMH 61544, ZMH 61537, ZMH 61538, ZMH 61539, ZMH 61540, ZMH 61541, ZMH 61542, ZMH 61543" collectingDate="1933-06-01" collectingDateMax="1934-05-28" collectingDateMin="1933-06-01" collectorName="Dr. H. Bytinski-Salz, Coll. Bytinski-Salz" specimenCount="9" specimenCount-male="9">
Labelled as
<normalizedToken originalValue="“Cotypes”">&quot;Cotypes&quot;</normalizedToken>
<specimenCount type="male">9♂</specimenCount>
<specimenCount>♀</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61536">ZMH 61536</accessionNumber>
-
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61544">ZMH 61544</accessionNumber>
) (Fig.
<figureCitation captionStart="Figures 67–74" captionStartId="F8" captionText="Figures 67 - 74. 67. Zygaena (Agrumenia) sogdiana altissima Burgeff, 1914; 68. Zygaena (Agrumenia) rosinae Korb, 1903; 69. Zygaena (Zygaena) transalpina sorrentina v. aureomaculata Stauder, 1921; 70. Zygaena exulans v. amarensis Diehl; 71. Zygaena (Zygaena) transalpina nigraltudinaria ab. flava Rocci, 1937; 72. Zygaena (Zygaena) transalpina sorrentina v. eranesceus Sicher, 1906; 73. Zygaena (Zygaena) transalpina litorea v. flava Stauder; 74. Zygaena (Mesembrynus) purpuralis v. asupinensis Byt. - S." httpUri="https://binary.pensoft.net/fig/522073" pageId="0" pageNumber="39">74</figureCitation>
). &quot;Rovigno
<normalizedToken originalValue="d´lstria">d'lstria</normalizedToken>
/ 6.1934 /
<collectorName>Dr. H. Bytinski-Salz</collectorName>
// Cotypus //
<collectorName>Coll. Bytinski-Salz</collectorName>
/ Eing. Nr. 20, 1960 //
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61536">ZMH 61536</accessionNumber>
&quot;; &quot;Rovigno
<normalizedToken originalValue="d´lstria">d'lstria</normalizedToken>
/ 6.1934 /
<collectorName>Dr.H.Bytinski-Salz</collectorName>
// Cotypus //
<collectorName>Coll. Bytinski-Salz</collectorName>
/ Eing. Nr. 20, 1960 //
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61537">ZMH 61537</accessionNumber>
&quot;; &quot;Rovigno
<normalizedToken originalValue="d´lstria">d'lstria</normalizedToken>
/ 6.1934 /
<collectorName>Dr. H. Bytinski-Salz</collectorName>
// Cotypus //
<collectorName>Coll. Bytinski-Salz</collectorName>
/ Eing. Nr. 20, 1960 //
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61538">ZMH 61538</accessionNumber>
&quot;; &quot;Rovigno
<normalizedToken originalValue="d´lstria">d'lstria</normalizedToken>
/ 6.1934 /
<collectorName>Dr.H.Bytinski-Salz</collectorName>
// Cotypus // var. asupinensis / By.5. //
<collectorName>Coll. Bytinski-Salz</collectorName>
/ Eing. Nr. 20, 1960 //
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61539">ZMH 61539</accessionNumber>
&quot;; &quot;Rovigno
<normalizedToken originalValue="d´lstria">d'lstria</normalizedToken>
/ 6.1933 /
<collectorName>Dr. H. Bytinski-Salz</collectorName>
// Cotypus //
<collectorName>Coll. Bytinski-Salz</collectorName>
/ Eing. Nr. 20, 1960 //
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61540">ZMH 61540</accessionNumber>
&quot;; &quot;Rovigno
<normalizedToken originalValue="d´lstria">d'lstria</normalizedToken>
/ 6.1934 /
<collectorName>Dr. H. Bytinski-Salz</collectorName>
// Cotypus //
<collectorName>Coll. Bytinski-Salz</collectorName>
/ Eing. Nr. 20, 1960 //
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61541">ZMH 61541</accessionNumber>
&quot;; &quot;Rovigno
<normalizedToken originalValue="d´lstria">d'lstria</normalizedToken>
/ 1.-
<collectingDate value="1933-06-06">6.6.1933</collectingDate>
/
<collectorName>Dr. H. Bytinski-Salz</collectorName>
// Cotypus // Coll. Bytinski-SalzColl. / Eing. Nr. 20, 1960 Eing. //
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61542">ZMH 61542</accessionNumber>
&quot;; &quot;Brioni / Istrien /
<collectingDate value="1934-05-28">28.V.1934</collectingDate>
/
<collectorName>Dr. H. Bytinski-Salz</collectorName>
// Cotypus //
<collectorName>Coll. Bytinski-Salz</collectorName>
/ Eing. Nr. 20, 1960 //
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61543">ZMH 61543</accessionNumber>
&quot;; &quot;Rovigno
<normalizedToken originalValue="d´lstria">d'lstria</normalizedToken>
/
<collectingDate value="1933-06-01" valueMax="1933-06-06" valueMin="1933-06-01">1.-6.VI.1933</collectingDate>
/
<collectorName>Dr. H. Bytinski-Salz</collectorName>
// Cotypus //
<collectorName>Coll. Bytinski-Salz</collectorName>
/ Eing. Nr. 20, 1960 //
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH61544">ZMH 61544</accessionNumber>
&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="current status">
<paragraph pageId="0" pageNumber="39">Current status.</paragraph>
<paragraph pageId="0" pageNumber="39">&quot;in litteris&quot; name and hence not available.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39" type="remarks">
<paragraph pageId="0" pageNumber="39">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="39">This taxon cannot be found in the literature.</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/522073" pageId="0" pageNumber="39" start="Figures 67–74" startId="F8">
<paragraph pageId="0" pageNumber="39">
<emphasis bold="true" pageId="0" pageNumber="39">Figures 67-74.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="39">67.</emphasis>
<taxonomicName authorityName="Burgeff" authorityYear="1914" class="Insecta" family="Hepialidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Agrumenia) sogdiana subsp. altissima" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="sogdiana" subGenus="Agrumenia" subSpecies="altissima">Zygaena (Agrumenia) sogdiana altissima</taxonomicName>
Burgeff, 1914;
<emphasis bold="true" pageId="0" pageNumber="39">68.</emphasis>
<taxonomicName authorityName="Korb" authorityYear="1903" class="Insecta" family="Hepialidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Agrumenia) rosinae" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="rosinae" subGenus="Agrumenia">Zygaena (Agrumenia) rosinae</taxonomicName>
Korb, 1903;
<emphasis bold="true" pageId="0" pageNumber="39">69.</emphasis>
<taxonomicName authorityName="Stauder" authorityYear="1921" class="Insecta" family="Hepialidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Zygaena) transalpina subsp. v. var. aureomaculata" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="variety" species="transalpina" subGenus="Zygaena" subSpecies="v." variety="aureomaculata">Zygaena (Zygaena) transalpina sorrentina v. aureomaculata</taxonomicName>
Stauder, 1921;
<emphasis bold="true" pageId="0" pageNumber="39">70.</emphasis>
<taxonomicName authorityName="Reiner &amp; Hohenwarth" authorityYear="1792" class="Insecta" family="Hepialidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena exulans" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="exulans">
<emphasis italics="true" pageId="0" pageNumber="39">Zygaena exulans</emphasis>
</taxonomicName>
v. amarensis Diehl;
<emphasis bold="true" pageId="0" pageNumber="39">71.</emphasis>
<taxonomicName authority="flava" class="Insecta" family="Hepialidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Zygaena) transalpina subsp. ab." order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="transalpina" subGenus="Zygaena" subSpecies="ab.">Zygaena (Zygaena) transalpina nigraltudinaria ab. flava</taxonomicName>
Rocci, 1937;
<emphasis bold="true" pageId="0" pageNumber="39">72.</emphasis>
<taxonomicName authorityName="Sicher" authorityYear="1906" class="Insecta" family="Hepialidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Zygaena) transalpina subsp. v. var. eranesceus" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="variety" species="transalpina" subGenus="Zygaena" subSpecies="v." variety="eranesceus">Zygaena (Zygaena) transalpina sorrentina v. eranesceus</taxonomicName>
Sicher, 1906;
<emphasis bold="true" pageId="0" pageNumber="39">73.</emphasis>
<taxonomicName class="Insecta" family="Hepialidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Zygaena) transalpina subsp. v. var. flava" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="variety" species="transalpina" subGenus="Zygaena" subSpecies="v." variety="flava">Zygaena (Zygaena) transalpina litorea v. flava</taxonomicName>
Stauder;
<emphasis bold="true" pageId="0" pageNumber="39">74.</emphasis>
<taxonomicName class="Insecta" family="Hepialidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Mesembrynus) purpuralis subsp. v. var. asupinensis" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="variety" species="purpuralis" subGenus="Mesembrynus" subSpecies="v." variety="asupinensis">Zygaena (Mesembrynus) purpuralis v. asupinensis</taxonomicName>
Byt.-S.
</paragraph>
</caption>
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<taxonomicName id="42B676A0EB95FEAF8CDF03066FAD6447" class="Insecta" family="Braconidae" genus="Apanteles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apanteles enephes" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="species" species="enephes">Apanteles enephes</taxonomicName>
Nixon, 1965
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<paragraph id="F52E6E9FF39B6C667E49C39546A9DD2C" pageId="0" pageNumber="8151">Distribution</paragraph>
<paragraph id="5EAFFB0E10720F2778AF8B496CAC075B" pageId="0" pageNumber="8151">England</paragraph>
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<emphasis id="B2A3C854D01B5FCB5FA415EAC7D0FAC2" italics="true" pageNumber="179">Hylobates albibarbis</emphasis>
Lyon 1911
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<taxonomicName id="E4F49E6F52C94699B3AB9EA43D85D5AE" ID-CoL="3NFRJ" authority="Lyon 1911" authorityName="Lyon" authorityYear="1911" class="Mammalia" family="Hylobatidae" genus="Hylobates" kingdom="Animalia" order="Primates" pageNumber="179" phylum="Chordata" rank="species" species="albibarbis">
<emphasis id="BD5DFCE898C1881B7BDAF124B5C96BBA" italics="true" pageNumber="179">Hylobates albibarbis</emphasis>
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,
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Proc.
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Natl. Mus., 40: 142
</bibCitation>
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.
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<emphasis id="19B376D0C47BE8333399D6E90EB01F55" inLineHeading="true" pageNumber="179">Type Locality:</emphasis>
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<collectingCountry id="2B24ECF61D4249F97B969ADC91288442" name="Indonesia">Indonesia</collectingCountry>
, W
<taxonomicName id="5E1B9D20B0117028456E003383F183CD" authority=", Sukadana." class="Mammalia" family="Cercopithecidae" genus="Erythrocebus" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="borneo">Borneo, Sukadana.</taxonomicName>
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<emphasis id="2BCADA83DB5929BDF2FD3183C4FFF2B5" inLineHeading="true" pageNumber="179">Vernacular Names:</emphasis>
<vernacularName id="4E51CBD2459165353FCA6B0AF38FDB1A" language="eng" pageNumber="179">Bornean White-bearded Gibbon</vernacularName>
.
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<paragraph id="CE45BBF8BD658E0D38BC238055B3472E" pageNumber="179">
<emphasis id="D81FD310CE9F8DE84630426B2BC3EBA5" inLineHeading="true" pageNumber="179">Distribution:</emphasis>
SW Borneo, south of Kapuas
<collectingRegion id="4EADC7E20892539BD1A99D5377AF5D3A" country="Gambia" name="Upper River">River</collectingRegion>
and W of Barito
<collectingRegion id="ED1A67DF867E42ED9A625923E99698F1" country="Gambia" name="Upper River">River</collectingRegion>
.
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<emphasis id="A7121F8AE82B7E6148CD76C27E8570AB" inLineHeading="true" pageNumber="179">Conservation:</emphasis>
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– Appendix I;
<collectingCountry id="49BE740B618A3460B29B14D0662524FD" name="United States of America">U.S.</collectingCountry>
<collectionCode id="E70AACBAD35C7B586C26FE0BF6178F7A" country="Brazil" lsid="urn:lsid:biocol.org:col:14319" name="Universidade de São Paulo" type="Herbarium">ESA</collectionCode>
– Endangered;
<collectionCode id="C41A03F347AA76633AD8E4E089AA857D">IUCN</collectionCode>
– Lower Risk (nt) as
<taxonomicName id="0707C2C10317A5568E8519ABD4294660" baseAuthorityName="Lyon" baseAuthorityYear="1911" class="Mammalia" family="Hylobatidae" genus="Hylobates" kingdom="Animalia" order="Primates" phylum="Chordata" rank="subSpecies" species="agilis" subSpecies="albibarbis">
<emphasis id="256A0784872A2ECEB2FBDADF7CF74E10" italics="true" pageNumber="179">H. agilis albibarbis</emphasis>
</taxonomicName>
.
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<paragraph id="29C3477C900749091837E97D03CF8A45" pageNumber="179">
<emphasis id="5E590F4460407CF4485DC1C718F2BFD5" inLineHeading="true" pageNumber="179">Discussion:</emphasis>
A subspecies of
<taxonomicName id="7A72C699F8FF981BDE0FE65124910A00" authorityName="F. Cuvier" authorityYear="1821" class="Mammalia" family="Hylobatidae" genus="Hylobates" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="agilis">
<emphasis id="E7F4B8D55E12E869872EAE6FFA46DA9F" italics="true" pageNumber="179">agilis</emphasis>
</taxonomicName>
according to
<bibRefCitation id="FB7C6B4D87BCFDFD9398CB2536CEAF27" author="Marshall, J. T., Jr. &amp; E. R. Marshall" refId="ref226350" refString="Marshall, J. T., Jr., and E. R. Marshall. 1976. Gibbons and their territorial songs. Science, 193: 235 - 237." year="1976">Marshall and Marshall (1976)</bibRefCitation>
, a subspecies of
<taxonomicName id="37B637A47D1BAF2EEC38D04CC757C96A" authorityName="Martin" authorityYear="1841" class="Mammalia" family="Hylobatidae" genus="Hylobates" kingdom="Animalia" order="Primates" phylum="Chordata" rank="species" species="muelleri">
<emphasis id="7F1E347AB30A96307F4829AD5BA60F04" italics="true" pageNumber="179">muelleri</emphasis>
</taxonomicName>
according to Groves (1974), a distinct species according to
<bibRefCitation id="C0FCA5B04614F8A77FC9155E0AD4CA7A" author="Groves, C. P." refId="ref132585" refString="Groves, C. P. 2001 c. Primate taxonomy. Smithsonian Institution Press, Washington, DC, 350 pp." year="2001">
Groves (2001
<emphasis id="142D1FC4E43A1EE3E0C5834F0F3F8C08" italics="true" pageNumber="179">c</emphasis>
)
</bibRefCitation>
.
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</heading>
</paragraph>
<paragraph id="3E1936FCFFDFFFB5FF01FA9CFEA0FAD0" blockId="1.[151,520,1255,1316]" box="[151,268,1290,1316]" pageId="1" pageNumber="60">
(
<figureCitation id="A69D2A79FFDFFFB5FF09FA9CFF49FAD0" box="[159,229,1290,1316]" captionStart="FIGURE 1" captionStartId="3.[151,255,1131,1155]" captionTargetBox="[256,1395,181,1034]" captionTargetId="figure@3.[178,1409,146,1114]" captionTargetPageId="3" captionText="FIGURE 1. Ranitomeya tolimense, new species. Lateral view showing color pattern. CZUT-A 989 adult female, SVL 17.94 mm. Photo: Manuel H. Bernal." httpUri="https://zenodo.org/record/179598/files/figure.png" pageId="1" pageNumber="60">Fig. 1</figureCitation>
,
<figureCitation id="A69D2A79FFDFFFB5FF65FA9CFEAFFAD0" box="[243,259,1290,1316]" captionStart="FIGURE 2" captionStartId="4.[151,255,1401,1425]" captionTargetBox="[185,1393,200,1365]" captionTargetId="figure@4.[180,1408,194,1377]" captionTargetPageId="4" captionText="FIGURE 2. Male nurse frog (CZUT-A 990) with a tadpole over his back at the type locality. Photo: Victor Luna." httpUri="https://zenodo.org/record/179599/files/figure.png" pageId="1" pageNumber="60">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="76BC6577FFDFFFB5FF01FACCFD89F9E0" pageId="1" pageNumber="60" type="materials_examined">
<paragraph id="3E1936FCFFDFFFB5FF01FACCFB12FA30" blockId="1.[151,1437,1370,2036]" pageId="1" pageNumber="60">
<emphasis id="0CD2EAEEFFDFFFB5FF01FACCFEBEFA80" bold="true" box="[151,274,1370,1396]" pageId="1" pageNumber="60">
<typeStatus id="E11D885EFFDFFFB5FF01FACCFEA2FA80" box="[151,270,1370,1396]" pageId="1" pageNumber="60" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="58B7AE39FFDFFFB5FE8DFACCFEE1FA80" box="[283,333,1370,1396]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="1" pageNumber="60">ICN</collectionCode>
53372, an adult male, collected by Oscar Gallego, Victor Luna and Manuel H. Bernal on
<date id="4A18103CFFDFFFB5FAE9FACCFEA3FA68" pageId="1" pageNumber="60" value="2007-05-19">19 May 2007</date>
near “Finca La Lulera”, vereda el Llano, Departamento del Tolima, Municipio de Falan, Cordillera Central of
<collectingCountry id="46B1766CFFDFFFB5FE82FA3CFE27FA30" box="[276,395,1450,1476]" name="Colombia" pageId="1" pageNumber="60">Colombia</collectingCountry>
, elevation
<quantity id="F95E9B19FFDFFFB5FD9FFA3CFDCEFA30" box="[521,610,1450,1476]" metricMagnitude="3" metricUnit="m" metricValue="1.8519999999999999" pageId="1" pageNumber="60" unit="m" value="1852.0">1852 m</quantity>
above sea level (
<geoCoordinate id="5B92503BFFDFFFB5FCBAFA3CFC02FA30" box="[812,942,1450,1476]" direction="north" orientation="latitude" pageId="1" pageNumber="60" precision="15" value="5.018889">5º01’08”N</geoCoordinate>
,
<geoCoordinate id="5B92503BFFDFFFB5FC2CFA3CFBE2FA30" box="[954,1102,1450,1476]" direction="west" orientation="longitude" pageId="1" pageNumber="60" precision="15" value="-75.04195">75º02’31”W</geoCoordinate>
) (
<figureCitation id="A69D2A79FFDFFFB5FBF1FA3CFB02FA30" box="[1127,1198,1450,1476]" captionStart="FIGURE 3" captionStartId="5.[151,255,1955,1979]" captionTargetBox="[292,1307,926,1900]" captionTargetId="figure@5.[226,1360,901,1932]" captionTargetPageId="5" captionText="FIGURE 3. Map of Colombia showing (arrow) the type locality of Ranitomeya tolimense." httpUri="https://zenodo.org/record/179600/files/figure.png" pageId="1" pageNumber="60">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="3E1936FCFFDFFFB5FF53FA44FD89F9E0" blockId="1.[151,1437,1370,2036]" pageId="1" pageNumber="60">
<emphasis id="0CD2EAEEFFDFFFB5FF53FA44FEE0FA18" bold="true" box="[197,332,1490,1516]" pageId="1" pageNumber="60">
<typeStatus id="E11D885EFFDFFFB5FF53FA44FEEBFA18" box="[197,327,1490,1516]" pageId="1" pageNumber="60" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="58B7AE39FFDFFFB5FEC3FA44FE2BFA18" box="[341,391,1490,1516]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="1" pageNumber="60">ICN</collectionCode>
53373, adult female; CZUT-A 988-989, adult females; CZUT-A 990-991, adult males; collected along with the
<typeStatus id="E11D885EFFDFFFB5FE21FA6CFD8EF9E0" box="[439,546,1530,1556]" pageId="1" pageNumber="60" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="76BC6577FFDFFFB5FF53F9B4FBB5F800" pageId="1" pageNumber="60" type="etymology">
<paragraph id="3E1936FCFFDFFFB5FF53F9B4FEE1F990" blockId="1.[151,1437,1370,2036]" pageId="1" pageNumber="60">
<emphasis id="0CD2EAEEFFDFFFB5FF53F9B4FEF9F9C8" bold="true" box="[197,341,1570,1596]" pageId="1" pageNumber="60">Etymology:</emphasis>
The specific epithet means “from Tolima”, the department of
<collectingCountry id="46B1766CFFDFFFB5FBAFF9B4FB01F9C8" box="[1081,1197,1570,1596]" name="Colombia" pageId="1" pageNumber="60">Colombia</collectingCountry>
where the new species was found.
</paragraph>
<paragraph id="3E1936FCFFDFFFB5FF53F9E4FBB5F800" blockId="1.[151,1437,1370,2036]" pageId="1" pageNumber="60">
<emphasis id="0CD2EAEEFFDFFFB5FF53F9E4FDAEF978" bold="true" box="[197,514,1650,1676]" pageId="1" pageNumber="60">Definition and diagnosis:</emphasis>
<taxonomicName id="F9A64D7FFFDFFFB5FD9DF9E5FCBDF978" box="[523,785,1651,1676]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="1" pageNumber="60" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFDFFFB5FD9DF9E5FCBDF978" box="[523,785,1651,1676]" italics="true" pageId="1" pageNumber="60">Ranitomeya tolimense</emphasis>
</taxonomicName>
is a dendrobatid having an adult SVL of
<quantity id="F95E9B19FFDFFFB5FA9FF9E4FF65F940" metricMagnitude="-2" metricUnit="m" metricValue="1.829" metricValueMax="1.892" metricValueMin="1.766" pageId="1" pageNumber="60" unit="mm" value="18.29" valueMax="18.92" valueMin="17.66">17.66–18.92 mm</quantity>
. Dorsal surfaces in life brown or dark brown, anteriorly with a moderate metallic yellow-bronze color that ends posterior to the head and gradually after the midbody. Ventral surfaces in life overall brown dark, with or without bluish green spots or speckling on belly, undersides of thighs and groins. The skin is granular on the dorsum, flanks, venter and the dorsal surface of the limbs, and comparatively less granular on the forelimbs. A brown dark stripe extends from the outer border of the eyes, crossing the tympanum, to the forelimbs insertions, which contrasts with the yellow color of the upper jaw. The proximodorsal surface of the upper arm is yellow. Fingers are webless; adpressed first finger shorter than second; third finger disc of adults about 1.5– 2.0 times wider than distal end of adjacent phalanx, discs on fingers notably expanded. Digits of feet lacking webbing or lateral fringe; toe I shorter than toe II, and unexpanded toe discs.
</paragraph>
</subSubSection>
<subSubSection id="76BC6577FFDCFFB0FF53FF0EFF46F9A2" lastPageId="4" lastPageNumber="63" pageId="2" pageNumber="61" type="description">
<paragraph id="3E1936FCFFDCFFB6FF53FF0EFE68FDED" blockId="2.[151,1437,152,978]" pageId="2" pageNumber="61">
<taxonomicName id="F9A64D7FFFDCFFB6FF53FF0EFE65FF45" box="[197,457,152,177]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFDCFFB6FF53FF0EFE65FF45" box="[197,457,152,177]" italics="true" pageId="2" pageNumber="61">Ranitomeya tolimense</emphasis>
</taxonomicName>
is considered to be related to
<emphasis id="0CD2EAEEFFDCFFB6FCB9FF0EFAA4FF45" box="[815,1288,152,177]" italics="true" pageId="2" pageNumber="61">
<taxonomicName id="F9A64D7FFFDCFFB6FCB9FF0EFC32FF45" box="[815,926,152,177]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="abdita">R. abdita</taxonomicName>
,
<taxonomicName id="F9A64D7FFFDCFFB6FC3DFF0EFB93FF45" box="[939,1087,152,177]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="bombetes">R. bombetes</taxonomicName>
,
<taxonomicName id="F9A64D7FFFDCFFB6FBDDFF0EFAA4FF45" box="[1099,1288,152,177]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="opisthomelas">R. opisthomelas</taxonomicName>
</emphasis>
and
<taxonomicName id="F9A64D7FFFDCFFB6FAD4FF0EFF5DFF2D" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="virolinensis">
<emphasis id="0CD2EAEEFFDCFFB6FAD4FF0EFF5DFF2D" italics="true" pageId="2" pageNumber="61">R. virolinensis</emphasis>
</taxonomicName>
based on shared larval characters, proposed to represent synapomorphies by
<bibRefCitation id="5A374B0DFFDCFFB6FB06FF56FA34FF2E" author="Myers" box="[1168,1432,192,218]" pageId="2" pageNumber="68" refString="Myers, C. W. &amp; Daly J. (1980) Taxonomy and ecology of Dendrobates bombetes, a new Andean poison frog with new skin toxins. American Museum Novitates, New York, 2694, 1 - 23." type="journal article" year="1980">Myers &amp; Daly (1980)</bibRefCitation>
. The character state is the median gap that interrupts the papillae fringe on the lower edge of the oral disc. The new species can be distinguished from these four species by the dorsal color:
<taxonomicName id="F9A64D7FFFDCFFB6FB8AFE86FB2BFEDD" box="[1052,1159,272,297]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="abdita">
<emphasis id="0CD2EAEEFFDCFFB6FB8AFE86FB2BFEDD" box="[1052,1159,272,297]" italics="true" pageId="2" pageNumber="61">R. abdita</emphasis>
</taxonomicName>
is unicolor dark bronzy brown with orange flash marks proximally on the arms and legs adjacent to axilla and groin;
<taxonomicName id="F9A64D7FFFDCFFB6FB79FEAEFA2CFEA5" box="[1263,1408,312,337]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="bombetes">
<emphasis id="0CD2EAEEFFDCFFB6FB79FEAEFA2CFEA5" box="[1263,1408,312,337]" italics="true" pageId="2" pageNumber="61">R. bombetes</emphasis>
</taxonomicName>
is black with dorsolateral stripes yellowish or orange golden;
<taxonomicName id="F9A64D7FFFDCFFB6FCD9FEF6FBA6FE8D" box="[847,1034,352,377]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="opisthomelas">
<emphasis id="0CD2EAEEFFDCFFB6FCD9FEF6FBA6FE8D" box="[847,1034,352,377]" italics="true" pageId="2" pageNumber="61">R. opisthomelas</emphasis>
</taxonomicName>
and
<taxonomicName id="F9A64D7FFFDCFFB6FBD5FEF6FB40FE8D" box="[1091,1260,352,377]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="virolinensis">
<emphasis id="0CD2EAEEFFDCFFB6FBD5FEF6FB40FE8D" box="[1091,1260,352,377]" italics="true" pageId="2" pageNumber="61">R. virolinensis</emphasis>
</taxonomicName>
are largely red turning gray distally on limbs and dark brown after two-third of body, respectively.
<taxonomicName id="F9A64D7FFFDCFFB6FB16FE1EFABFFE55" box="[1152,1299,392,417]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFDCFFB6FB16FE1EFABFFE55" box="[1152,1299,392,417]" italics="true" pageId="2" pageNumber="61">R. tolimense</emphasis>
</taxonomicName>
is syntopic with
<taxonomicName id="F9A64D7FFFDCFFB6FF43FE26FE19FE3D" box="[213,437,432,457]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="61" phylum="Chordata" rank="species" species="dorisswansonae">
<emphasis id="0CD2EAEEFFDCFFB6FF43FE26FE19FE3D" box="[213,437,432,457]" italics="true" pageId="2" pageNumber="61">R. dorisswansonae</emphasis>
</taxonomicName>
, but readily distinguished by color pattern and morphology since the latter is dark brown or black with irregular red blotches, and the toes I and II are partially or totally fused (not so in the new species described herein).
</paragraph>
<paragraph id="3E1936FCFFDCFFB6FF53FDB1FD70FD66" blockId="2.[151,1437,152,978]" pageId="2" pageNumber="61">
<emphasis id="0CD2EAEEFFDCFFB6FF53FDB1FDC7FDB5" bold="true" box="[197,619,551,577]" pageId="2" pageNumber="61">
Measurements of
<typeStatus id="E11D885EFFDCFFB6FE3EFDB1FDB8FDB5" box="[424,532,551,577]" pageId="2" pageNumber="61" type="holotype">holotype</typeStatus>
(mm):
</emphasis>
SVL 17.66; TL 7.98; GBW 8.13; greatest HW 5.97; IOD 2.35; HL 5.04; TSCN 1.55; NED 1.36;
<collectingCountry id="46B1766CFFDCFFB6FE6FFDC6FD81FD9E" box="[505,557,592,618]" name="India" pageId="2" pageNumber="61">IND</collectingCountry>
2.52; EL 2.32; HDT 1.18; MTD 1.02; HaL 4.65; WTFD 0.80; WTF 0.40; WTTD 0.64; WTT 0.61; WFTD 0.50; WFT 0.45.
</paragraph>
<paragraph id="3E1936FCFFDCFFB6FF53FD09FD0EFC26" blockId="2.[151,1437,152,978]" pageId="2" pageNumber="61">
<emphasis id="0CD2EAEEFFDCFFB6FF53FD09FC4BFD4D" bold="true" box="[197,999,671,697]" pageId="2" pageNumber="61">
Description of the
<typeStatus id="E11D885EFFDCFFB6FE20FD09FE40FD4D" box="[438,492,671,697]" pageId="2" pageNumber="61">type</typeStatus>
series (
<specimenCount id="28A0FD75FFDCFFB6FDC6FD09FD07FD4D" box="[592,683,671,697]" pageId="2" pageNumber="61" type="adult">3 adult</specimenCount>
males,
<specimenCount id="28A0FD75FFDCFFB6FC86FD09FCC6FD4D" box="[784,874,671,697]" pageId="2" pageNumber="61" type="adult">3 adult</specimenCount>
females):
</emphasis>
Size small, with maximum SVL of 18.91 (
<tableCitation id="73240347FFDCFFB6FF7EFD5EFEECFD16" box="[232,320,712,738]" captionStart="TABLE 1" captionStartId="2.[151,239,1031,1055]" captionTargetBox="[159,1400,1088,1956]" captionTargetPageId="2" captionText="TABLE 1. Measurements (in mm) for the type series of Ranitomeya tolimense, new species." httpUri="http://table.plazi.org/id/6AD96674FFDCFFB6FF01FB91FBDFFBEB" pageId="2" pageNumber="61" tableUuid="6AD96674FFDCFFB6FF01FB91FBDFFBEB">Table 1</tableCitation>
); females slightly larger than males (
<tableCitation id="73240347FFDCFFB6FD60FD5EFCFDFD16" box="[758,849,712,738]" captionStart="TABLE 2" captionStartId="5.[151,239,351,375]" captionTargetBox="[159,1380,408,817]" captionTargetPageId="5" captionText="TABLE 2. Size and proportions of adult Ranitomeya tolimense, New species sensu Myers &amp; Daly (1980)." httpUri="http://table.plazi.org/id/6AD96674FFDBFFB1FF01FEC9FAAFFE83" pageId="2" pageNumber="61" tableUuid="6AD96674FFDBFFB1FF01FEC9FAAFFE83">Table 2</tableCitation>
). Head narrower than body; greatest HW 73 % of GBW in males, and 79 % in females. Greatest HW averaging about 30 % of SVL (
<tableCitation id="73240347FFDCFFB6FB0AFD66FB57FCFE" box="[1180,1275,752,778]" captionStart="TABLE 2" captionStartId="5.[151,239,351,375]" captionTargetBox="[159,1380,408,817]" captionTargetPageId="5" captionText="TABLE 2. Size and proportions of adult Ranitomeya tolimense, New species sensu Myers &amp; Daly (1980)." httpUri="http://table.plazi.org/id/6AD96674FFDBFFB1FF01FEC9FAAFFE83" pageId="2" pageNumber="61" tableUuid="6AD96674FFDBFFB1FF01FEC9FAAFFE83">Table 2</tableCitation>
). Males with defined vocal slits, but with little or not external indication of the subgular vocal sac. Snout weakly pointed to truncate in dorsal and ventral aspects. Nares situated near tip of snout and visible from front and from below but not from above. Canthus rostralis rounded, loreal region flat, inclined slightly outward to lip. Eye longer than snout length; distance from center of naris to eye 65 % of EL in males and 64 % in females (
<tableCitation id="73240347FFDCFFB6FAA7FC06FA21FC5E" box="[1329,1421,912,938]" captionStart="TABLE 2" captionStartId="5.[151,239,351,375]" captionTargetBox="[159,1380,408,817]" captionTargetPageId="5" captionText="TABLE 2. Size and proportions of adult Ranitomeya tolimense, New species sensu Myers &amp; Daly (1980)." httpUri="http://table.plazi.org/id/6AD96674FFDBFFB1FF01FEC9FAAFFE83" pageId="2" pageNumber="61" tableUuid="6AD96674FFDBFFB1FF01FEC9FAAFFE83">Table 2</tableCitation>
). Tympanum vertically oval, smaller than eye.
</paragraph>
<caption id="6AD96674FFDCFFB6FF01FB91FBDFFBEB" ID-Table-UUID="6AD96674FFDCFFB6FF01FB91FBDFFBEB" box="[151,1139,1031,1055]" httpUri="http://table.plazi.org/id/6AD96674FFDCFFB6FF01FB91FBDFFBEB" pageId="2" pageNumber="61" targetBox="[159,1400,1088,1956]" targetIsTable="true" targetPageId="2">
<paragraph id="3E1936FCFFDCFFB6FF01FB91FBDFFBEB" blockId="2.[151,1139,1031,1055]" box="[151,1139,1031,1055]" pageId="2" pageNumber="61">
<emphasis id="0CD2EAEEFFDCFFB6FF01FB91FEA4FBEB" bold="true" box="[151,264,1031,1055]" pageId="2" pageNumber="61">TABLE 1.</emphasis>
Measurements (in mm) for the type series of
<taxonomicName id="F9A64D7FFFDCFFB6FD64FB9EFBBBFBEB" authority="New" authorityName="New" box="[754,1047,1032,1055]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="64" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFDCFFB6FD64FB9EFC72FBEB" box="[754,990,1032,1055]" italics="true" pageId="2" pageNumber="61">Ranitomeya tolimense</emphasis>
,
<taxonomicNameLabel id="17E15795FFDCFFB6FC7DFB9EFBBBFBEB" box="[1003,1047,1032,1055]" pageId="2" pageNumber="61" rank="species">new</taxonomicNameLabel>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="3E1936FCFFDCFFB6FEA2FBD6FAB5F850" pageId="2" pageNumber="61">
<table id="4CA6C45CFFDC004BFF09FBD6FAD4F850" box="[159,1400,1088,1956]" gridcols="7" gridrows="14" pageId="2" pageNumber="61">
<tr id="809634BEFFDC004BFF09FBD6FAD4FB8D" box="[159,1400,1088,1145]" gridrow="0" pageId="2" pageNumber="61" rowspan-0="1">
<th id="C3475DC2FFDC004BFEA5FBD6FE6CFB8D" box="[307,448,1088,1145]" gridcol="1" gridrow="0" pageId="2" pageNumber="61">ICN 53373 Adult female</th>
<th id="C3475DC2FFDC004BFE67FBD6FD2CFB8D" box="[497,640,1088,1145]" gridcol="2" gridrow="0" pageId="2" pageNumber="61">CZUT-A 988 Adult female</th>
<th id="C3475DC2FFDC004BFD26FBD6FC91FB8D" box="[688,829,1088,1145]" gridcol="3" gridrow="0" pageId="2" pageNumber="61">CZUT-A 989 Adult female</th>
<th id="C3475DC2FFDC004BFCF8FBD6FC49FB8D" box="[878,997,1088,1145]" gridcol="4" gridrow="0" pageId="2" pageNumber="61">ICN 53372 Adult male</th>
<th id="C3475DC2FFDC004BFBBAFBD6FB16FB8D" box="[1068,1210,1088,1145]" gridcol="5" gridrow="0" pageId="2" pageNumber="61">CZUT-A 990 Adult male</th>
<th id="C3475DC2FFDC004BFB7CFBD6FAD4FB8D" box="[1258,1400,1088,1145]" gridcol="6" gridrow="0" pageId="2" pageNumber="61">CZUT-A 991 Adult male</th>
</tr>
<tr id="809634BEFFDC004BFF09FB1BFAD4FB50" box="[159,1400,1165,1188]" gridrow="1" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09FB1BFF40FB50" box="[159,236,1165,1188]" gridcol="0" gridrow="1" pageId="2" pageNumber="61">SVL</th>
<td id="C3475DC2FFDC004BFEA5FB1BFE6CFB50" box="[307,448,1165,1188]" gridcol="1" gridrow="1" pageId="2" pageNumber="61">18.91</td>
<td id="C3475DC2FFDC004BFE67FB1BFD2CFB50" box="[497,640,1165,1188]" gridcol="2" gridrow="1" pageId="2" pageNumber="61">17.82</td>
<td id="C3475DC2FFDC004BFD26FB1BFC91FB50" box="[688,829,1165,1188]" gridcol="3" gridrow="1" pageId="2" pageNumber="61">18.11</td>
<td id="C3475DC2FFDC004BFCF8FB1BFC49FB50" box="[878,997,1165,1188]" gridcol="4" gridrow="1" pageId="2" pageNumber="61">17.66</td>
<td id="C3475DC2FFDC004BFBBAFB1BFB16FB50" box="[1068,1210,1165,1188]" gridcol="5" gridrow="1" pageId="2" pageNumber="61">17.95</td>
<td id="C3475DC2FFDC004BFB7CFB1BFAD4FB50" box="[1258,1400,1165,1188]" gridcol="6" gridrow="1" pageId="2" pageNumber="61">18.02</td>
</tr>
<tr id="809634BEFFDC004BFF09FB2EFAD4FB0D" box="[159,1400,1208,1273]" gridrow="2" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09FB2EFF40FB0D" box="[159,236,1208,1273]" gridcol="0" gridrow="2" pageId="2" pageNumber="61">TL GBW</th>
<td id="C3475DC2FFDC004BFEA5FB2EFE6CFB0D" box="[307,448,1208,1273]" gridcol="1" gridrow="2" pageId="2" pageNumber="61">7.94 8.16</td>
<td id="C3475DC2FFDC004BFE67FB2EFD2CFB0D" box="[497,640,1208,1273]" gridcol="2" gridrow="2" pageId="2" pageNumber="61">7.14 6.93</td>
<td id="C3475DC2FFDC004BFD26FB2EFC91FB0D" box="[688,829,1208,1273]" gridcol="3" gridrow="2" pageId="2" pageNumber="61">7.53 7.08</td>
<td id="C3475DC2FFDC004BFCF8FB2EFC49FB0D" box="[878,997,1208,1273]" gridcol="4" gridrow="2" pageId="2" pageNumber="61">7.98 8.13</td>
<td id="C3475DC2FFDC004BFBBAFB2EFB16FB0D" box="[1068,1210,1208,1273]" gridcol="5" gridrow="2" pageId="2" pageNumber="61">7.22 7.35</td>
<td id="C3475DC2FFDC004BFB7CFB2EFAD4FB0D" box="[1258,1400,1208,1273]" gridcol="6" gridrow="2" pageId="2" pageNumber="61">7.73 6.52</td>
</tr>
<tr id="809634BEFFDC004BFF09FA9BFAD4FAD0" box="[159,1400,1293,1316]" gridrow="3" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09FA9BFF40FAD0" box="[159,236,1293,1316]" gridcol="0" gridrow="3" pageId="2" pageNumber="61">HW</th>
<td id="C3475DC2FFDC004BFEA5FA9BFE6CFAD0" box="[307,448,1293,1316]" gridcol="1" gridrow="3" pageId="2" pageNumber="61">5.86</td>
<td id="C3475DC2FFDC004BFE67FA9BFD2CFAD0" box="[497,640,1293,1316]" gridcol="2" gridrow="3" pageId="2" pageNumber="61">5.42</td>
<td id="C3475DC2FFDC004BFD26FA9BFC91FAD0" box="[688,829,1293,1316]" gridcol="3" gridrow="3" pageId="2" pageNumber="61">5.64</td>
<td id="C3475DC2FFDC004BFCF8FA9BFC49FAD0" box="[878,997,1293,1316]" gridcol="4" gridrow="3" pageId="2" pageNumber="61">5.97</td>
<td id="C3475DC2FFDC004BFBBAFA9BFB16FAD0" box="[1068,1210,1293,1316]" gridcol="5" gridrow="3" pageId="2" pageNumber="61">5.81</td>
<td id="C3475DC2FFDC004BFB7CFA9BFAD4FAD0" box="[1258,1400,1293,1316]" gridcol="6" gridrow="3" pageId="2" pageNumber="61">5.35</td>
</tr>
<tr id="809634BEFFDC004BFF09FAAEFAD4FA8D" box="[159,1400,1336,1401]" gridrow="4" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09FAAEFF40FA8D" box="[159,236,1336,1401]" gridcol="0" gridrow="4" pageId="2" pageNumber="61">IOD HL</th>
<td id="C3475DC2FFDC004BFEA5FAAEFE6CFA8D" box="[307,448,1336,1401]" gridcol="1" gridrow="4" pageId="2" pageNumber="61">2.27 5.65</td>
<td id="C3475DC2FFDC004BFE67FAAEFD2CFA8D" box="[497,640,1336,1401]" gridcol="2" gridrow="4" pageId="2" pageNumber="61">2.25 4.97</td>
<td id="C3475DC2FFDC004BFD26FAAEFC91FA8D" box="[688,829,1336,1401]" gridcol="3" gridrow="4" pageId="2" pageNumber="61">2.36 5.14</td>
<td id="C3475DC2FFDC004BFCF8FAAEFC49FA8D" box="[878,997,1336,1401]" gridcol="4" gridrow="4" pageId="2" pageNumber="61">2.35 5.04</td>
<td id="C3475DC2FFDC004BFBBAFAAEFB16FA8D" box="[1068,1210,1336,1401]" gridcol="5" gridrow="4" pageId="2" pageNumber="61">2.16 5.08</td>
<td id="C3475DC2FFDC004BFB7CFAAEFAD4FA8D" box="[1258,1400,1336,1401]" gridcol="6" gridrow="4" pageId="2" pageNumber="61">2.19 5.46</td>
</tr>
<tr id="809634BEFFDC004BFF09FA1BFAD4FA50" box="[159,1400,1421,1444]" gridrow="5" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09FA1BFF40FA50" box="[159,236,1421,1444]" gridcol="0" gridrow="5" pageId="2" pageNumber="61">TSCN</th>
<td id="C3475DC2FFDC004BFEA5FA1BFE6CFA50" box="[307,448,1421,1444]" gridcol="1" gridrow="5" pageId="2" pageNumber="61">1.74</td>
<td id="C3475DC2FFDC004BFE67FA1BFD2CFA50" box="[497,640,1421,1444]" gridcol="2" gridrow="5" pageId="2" pageNumber="61">1.31</td>
<td id="C3475DC2FFDC004BFD26FA1BFC91FA50" box="[688,829,1421,1444]" gridcol="3" gridrow="5" pageId="2" pageNumber="61">1.57</td>
<td id="C3475DC2FFDC004BFCF8FA1BFC49FA50" box="[878,997,1421,1444]" gridcol="4" gridrow="5" pageId="2" pageNumber="61">1.55</td>
<td id="C3475DC2FFDC004BFBBAFA1BFB16FA50" box="[1068,1210,1421,1444]" gridcol="5" gridrow="5" pageId="2" pageNumber="61">1.38</td>
<td id="C3475DC2FFDC004BFB7CFA1BFAD4FA50" box="[1258,1400,1421,1444]" gridcol="6" gridrow="5" pageId="2" pageNumber="61">1.89</td>
</tr>
<tr id="809634BEFFDC004BFF09FA2EFAD4FA0D" box="[159,1400,1464,1529]" gridrow="6" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09FA2EFF40FA0D" box="[159,236,1464,1529]" gridcol="0" gridrow="6" pageId="2" pageNumber="61">NED IND</th>
<td id="C3475DC2FFDC004BFEA5FA2EFE6CFA0D" box="[307,448,1464,1529]" gridcol="1" gridrow="6" pageId="2" pageNumber="61">1.52 2.40</td>
<td id="C3475DC2FFDC004BFE67FA2EFD2CFA0D" box="[497,640,1464,1529]" gridcol="2" gridrow="6" pageId="2" pageNumber="61">1.44 2.22</td>
<td id="C3475DC2FFDC004BFD26FA2EFC91FA0D" box="[688,829,1464,1529]" gridcol="3" gridrow="6" pageId="2" pageNumber="61">1.35 2.34</td>
<td id="C3475DC2FFDC004BFCF8FA2EFC49FA0D" box="[878,997,1464,1529]" gridcol="4" gridrow="6" pageId="2" pageNumber="61">1.36 2.52</td>
<td id="C3475DC2FFDC004BFBBAFA2EFB16FA0D" box="[1068,1210,1464,1529]" gridcol="5" gridrow="6" pageId="2" pageNumber="61">1.65 2.33</td>
<td id="C3475DC2FFDC004BFB7CFA2EFAD4FA0D" box="[1258,1400,1464,1529]" gridcol="6" gridrow="6" pageId="2" pageNumber="61">1.38 2.19</td>
</tr>
<tr id="809634BEFFDC004BFF09F99BFAD4F9D0" box="[159,1400,1549,1572]" gridrow="7" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09F99BFF40F9D0" box="[159,236,1549,1572]" gridcol="0" gridrow="7" pageId="2" pageNumber="61">EL</th>
<td id="C3475DC2FFDC004BFEA5F99BFE6CF9D0" box="[307,448,1549,1572]" gridcol="1" gridrow="7" pageId="2" pageNumber="61">2.23</td>
<td id="C3475DC2FFDC004BFE67F99BFD2CF9D0" box="[497,640,1549,1572]" gridcol="2" gridrow="7" pageId="2" pageNumber="61">2.19</td>
<td id="C3475DC2FFDC004BFD26F99BFC91F9D0" box="[688,829,1549,1572]" gridcol="3" gridrow="7" pageId="2" pageNumber="61">2.24</td>
<td id="C3475DC2FFDC004BFCF8F99BFC49F9D0" box="[878,997,1549,1572]" gridcol="4" gridrow="7" pageId="2" pageNumber="61">2.32</td>
<td id="C3475DC2FFDC004BFBBAF99BFB16F9D0" box="[1068,1210,1549,1572]" gridcol="5" gridrow="7" pageId="2" pageNumber="61">2.20</td>
<td id="C3475DC2FFDC004BFB7CF99BFAD4F9D0" box="[1258,1400,1549,1572]" gridcol="6" gridrow="7" pageId="2" pageNumber="61">2.20</td>
</tr>
<tr id="809634BEFFDC004BFF09F9AEFAD4F98D" box="[159,1400,1592,1657]" gridrow="8" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09F9AEFF40F98D" box="[159,236,1592,1657]" gridcol="0" gridrow="8" pageId="2" pageNumber="61">HDT MTD</th>
<td id="C3475DC2FFDC004BFEA5F9AEFE6CF98D" box="[307,448,1592,1657]" gridcol="1" gridrow="8" pageId="2" pageNumber="61">1.44 0.85</td>
<td id="C3475DC2FFDC004BFE67F9AEFD2CF98D" box="[497,640,1592,1657]" gridcol="2" gridrow="8" pageId="2" pageNumber="61">1.13 0.64</td>
<td id="C3475DC2FFDC004BFD26F9AEFC91F98D" box="[688,829,1592,1657]" gridcol="3" gridrow="8" pageId="2" pageNumber="61">1.27 0.87</td>
<td id="C3475DC2FFDC004BFCF8F9AEFC49F98D" box="[878,997,1592,1657]" gridcol="4" gridrow="8" pageId="2" pageNumber="61">1.18 1.02</td>
<td id="C3475DC2FFDC004BFBBAF9AEFB16F98D" box="[1068,1210,1592,1657]" gridcol="5" gridrow="8" pageId="2" pageNumber="61">1.10 0.91</td>
<td id="C3475DC2FFDC004BFB7CF9AEFAD4F98D" box="[1258,1400,1592,1657]" gridcol="6" gridrow="8" pageId="2" pageNumber="61">1.33 0.94</td>
</tr>
<tr id="809634BEFFDC004BFF09F91BFAD4F950" box="[159,1400,1677,1700]" gridrow="9" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09F91BFF40F950" box="[159,236,1677,1700]" gridcol="0" gridrow="9" pageId="2" pageNumber="61">HaL</th>
<td id="C3475DC2FFDC004BFEA5F91BFE6CF950" box="[307,448,1677,1700]" gridcol="1" gridrow="9" pageId="2" pageNumber="61">4.57</td>
<td id="C3475DC2FFDC004BFE67F91BFD2CF950" box="[497,640,1677,1700]" gridcol="2" gridrow="9" pageId="2" pageNumber="61">4.30</td>
<td id="C3475DC2FFDC004BFD26F91BFC91F950" box="[688,829,1677,1700]" gridcol="3" gridrow="9" pageId="2" pageNumber="61">4.22</td>
<td id="C3475DC2FFDC004BFCF8F91BFC49F950" box="[878,997,1677,1700]" gridcol="4" gridrow="9" pageId="2" pageNumber="61">4.65</td>
<td id="C3475DC2FFDC004BFBBAF91BFB16F950" box="[1068,1210,1677,1700]" gridcol="5" gridrow="9" pageId="2" pageNumber="61">4.52</td>
<td id="C3475DC2FFDC004BFB7CF91BFAD4F950" box="[1258,1400,1677,1700]" gridcol="6" gridrow="9" pageId="2" pageNumber="61">4.10</td>
</tr>
<tr id="809634BEFFDC004BFF09F92EFAD4F90D" box="[159,1400,1720,1785]" gridrow="10" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09F92EFF40F90D" box="[159,236,1720,1785]" gridcol="0" gridrow="10" pageId="2" pageNumber="61">WTFD WTF</th>
<td id="C3475DC2FFDC004BFEA5F92EFE6CF90D" box="[307,448,1720,1785]" gridcol="1" gridrow="10" pageId="2" pageNumber="61">0.73 0.48</td>
<td id="C3475DC2FFDC004BFE67F92EFD2CF90D" box="[497,640,1720,1785]" gridcol="2" gridrow="10" pageId="2" pageNumber="61">0.64 0.40</td>
<td id="C3475DC2FFDC004BFD26F92EFC91F90D" box="[688,829,1720,1785]" gridcol="3" gridrow="10" pageId="2" pageNumber="61">0.65 0.39</td>
<td id="C3475DC2FFDC004BFCF8F92EFC49F90D" box="[878,997,1720,1785]" gridcol="4" gridrow="10" pageId="2" pageNumber="61">0.80 0.40</td>
<td id="C3475DC2FFDC004BFBBAF92EFB16F90D" box="[1068,1210,1720,1785]" gridcol="5" gridrow="10" pageId="2" pageNumber="61">0.75 0.46</td>
<td id="C3475DC2FFDC004BFB7CF92EFAD4F90D" box="[1258,1400,1720,1785]" gridcol="6" gridrow="10" pageId="2" pageNumber="61">0.78 0.40</td>
</tr>
<tr id="809634BEFFDC004BFF09F89BFAD4F8D0" box="[159,1400,1805,1828]" gridrow="11" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09F89BFF40F8D0" box="[159,236,1805,1828]" gridcol="0" gridrow="11" pageId="2" pageNumber="61">WTTD</th>
<td id="C3475DC2FFDC004BFEA5F89BFE6CF8D0" box="[307,448,1805,1828]" gridcol="1" gridrow="11" pageId="2" pageNumber="61">0.60</td>
<td id="C3475DC2FFDC004BFE67F89BFD2CF8D0" box="[497,640,1805,1828]" gridcol="2" gridrow="11" pageId="2" pageNumber="61">0.58</td>
<td id="C3475DC2FFDC004BFD26F89BFC91F8D0" box="[688,829,1805,1828]" gridcol="3" gridrow="11" pageId="2" pageNumber="61">0.56</td>
<td id="C3475DC2FFDC004BFCF8F89BFC49F8D0" box="[878,997,1805,1828]" gridcol="4" gridrow="11" pageId="2" pageNumber="61">0.64</td>
<td id="C3475DC2FFDC004BFBBAF89BFB16F8D0" box="[1068,1210,1805,1828]" gridcol="5" gridrow="11" pageId="2" pageNumber="61">0.60</td>
<td id="C3475DC2FFDC004BFB7CF89BFAD4F8D0" box="[1258,1400,1805,1828]" gridcol="6" gridrow="11" pageId="2" pageNumber="61">0.71</td>
</tr>
<tr id="809634BEFFDC004BFF09F8AEFAD4F88D" box="[159,1400,1848,1913]" gridrow="12" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09F8AEFF40F88D" box="[159,236,1848,1913]" gridcol="0" gridrow="12" pageId="2" pageNumber="61">WTT WFTD</th>
<td id="C3475DC2FFDC004BFEA5F8AEFE6CF88D" box="[307,448,1848,1913]" gridcol="1" gridrow="12" pageId="2" pageNumber="61">0.55 0.64</td>
<td id="C3475DC2FFDC004BFE67F8AEFD2CF88D" box="[497,640,1848,1913]" gridcol="2" gridrow="12" pageId="2" pageNumber="61">0.50 0.48</td>
<td id="C3475DC2FFDC004BFD26F8AEFC91F88D" box="[688,829,1848,1913]" gridcol="3" gridrow="12" pageId="2" pageNumber="61">0.44 0.46</td>
<td id="C3475DC2FFDC004BFCF8F8AEFC49F88D" box="[878,997,1848,1913]" gridcol="4" gridrow="12" pageId="2" pageNumber="61">0.61 0.50</td>
<td id="C3475DC2FFDC004BFBBAF8AEFB16F88D" box="[1068,1210,1848,1913]" gridcol="5" gridrow="12" pageId="2" pageNumber="61">0.54 0.44</td>
<td id="C3475DC2FFDC004BFB7CF8AEFAD4F88D" box="[1258,1400,1848,1913]" gridcol="6" gridrow="12" pageId="2" pageNumber="61">0.54 0.38</td>
</tr>
<tr id="809634BEFFDC004BFF09F81BFAD4F850" box="[159,1400,1933,1956]" gridrow="13" pageId="2" pageNumber="61">
<th id="C3475DC2FFDC004BFF09F81BFF40F850" box="[159,236,1933,1956]" gridcol="0" gridrow="13" pageId="2" pageNumber="61">WFT</th>
<td id="C3475DC2FFDC004BFEA5F81BFE6CF850" box="[307,448,1933,1956]" gridcol="1" gridrow="13" pageId="2" pageNumber="61">0.48</td>
<td id="C3475DC2FFDC004BFE67F81BFD2CF850" box="[497,640,1933,1956]" gridcol="2" gridrow="13" pageId="2" pageNumber="61">0.42</td>
<td id="C3475DC2FFDC004BFD26F81BFC91F850" box="[688,829,1933,1956]" gridcol="3" gridrow="13" pageId="2" pageNumber="61">0.36</td>
<td id="C3475DC2FFDC004BFCF8F81BFC49F850" box="[878,997,1933,1956]" gridcol="4" gridrow="13" pageId="2" pageNumber="61">0.45</td>
<td id="C3475DC2FFDC004BFBBAF81BFB16F850" box="[1068,1210,1933,1956]" gridcol="5" gridrow="13" pageId="2" pageNumber="61">0.40</td>
<td id="C3475DC2FFDC004BFB7CF81BFAD4F850" box="[1258,1400,1933,1956]" gridcol="6" gridrow="13" pageId="2" pageNumber="61">0.32</td>
</tr>
</table>
</paragraph>
<caption id="6AD96674FFDDFFB7FF01FBFDFD88FB51" httpUri="https://zenodo.org/record/179598/files/figure.png" pageId="3" pageNumber="62" targetBox="[256,1395,181,1034]" targetPageId="3">
<paragraph id="3E1936FCFFDDFFB7FF01FBFDFD88FB51" blockId="3.[151,1435,1131,1189]" pageId="3" pageNumber="62">
<emphasis id="0CD2EAEEFFDDFFB7FF01FBFDFEB7FB77" bold="true" box="[151,283,1131,1155]" pageId="3" pageNumber="62">FIGURE 1.</emphasis>
<taxonomicName id="F9A64D7FFFDDFFB7FEB5FBFAFDE0FB77" authority="New" authorityName="New" box="[291,588,1132,1155]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="64" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFDDFFB7FEB5FBFAFDBDFB77" box="[291,529,1132,1155]" italics="true" pageId="3" pageNumber="62">Ranitomeya tolimense</emphasis>
,
<taxonomicNameLabel id="17E15795FFDDFFB7FDB6FBFAFDE0FB77" box="[544,588,1132,1155]" pageId="3" pageNumber="62" rank="species">new</taxonomicNameLabel>
</taxonomicName>
species. Lateral view showing color pattern. CZUT-A 989 adult female, SVL 17.94 mm. Photo: Manuel H. Bernal.
</paragraph>
</caption>
<paragraph id="3E1936FCFFDDFFB7FF53FB4FFE52FA17" blockId="3.[151,1437,1241,1987]" pageId="3" pageNumber="62">
Hand (
<figureCitation id="A69D2A79FFDDFFB7FE82FB4FFEF6FB07" box="[276,346,1241,1267]" captionStart="FIGURE 4" captionStartId="6.[151,255,995,1019]" captionTargetBox="[237,1354,192,950]" captionTargetId="figure@6.[152,1435,176,990]" captionTargetPageId="6" captionText="FIGURE 4. Ventral view of hand and foot of Ranitomeya tolimense (CZUT-A 989). Scale = 1 mm." httpUri="https://zenodo.org/record/179601/files/figure.png" pageId="3" pageNumber="62">Fig. 4</figureCitation>
A) moderately large, its length (proximal edge of metacarpal tubercle to tip of longest finger) about 23.5–26.7 % of SVL. Relative length of adpressed fingers III&gt; IV&gt; II&gt; I; adpressed finger I shorter than finger II and its tip just reaching disc of second. Discs slightly expanded on finger I, but notably expanded on fingers II, III and IV; third finger disc in females and males averaging 1.59–1.86 times wider than distal end of adjacent phalanx. A large circular outer metacarpal tubercle on median base of palm, an smaller inner metacarpal tubercle on base of finger I, and one tubercle on base of fingers I and II, and two tubercles on fingers III and IV.
</paragraph>
<paragraph id="3E1936FCFFDDFFB7FF53FA67FECCF95F" blockId="3.[151,1437,1241,1987]" pageId="3" pageNumber="62">
<collectingCountry id="46B1766CFFDDFFB7FF53FA67FEACF9FF" box="[197,256,1521,1547]" name="India" pageId="3" pageNumber="62">Hind</collectingCountry>
limbs relatively long; tibia 40–45 % of SVL. Relative lengths of adpressed toes IV&gt; III&gt;V&gt;II&gt; I; toe I short (
<figureCitation id="A69D2A79FFDDFFB7FEB7F98FFEC5F9C7" box="[289,361,1561,1587]" captionStart="FIGURE 4" captionStartId="6.[151,255,995,1019]" captionTargetBox="[237,1354,192,950]" captionTargetId="figure@6.[152,1435,176,990]" captionTargetPageId="6" captionText="FIGURE 4. Ventral view of hand and foot of Ranitomeya tolimense (CZUT-A 989). Scale = 1 mm." httpUri="https://zenodo.org/record/179601/files/figure.png" pageId="3" pageNumber="62">Fig. 4</figureCitation>
B), reaching bottom of subarticular tubercle of toe II, and with unexpanded disc. Other toe discs slightly expanded. One to three nonprotuberant subarticular tubercles, one on toes I, II, two on toes III and V, and three on toe IV. Small outer and moderately-sized inner metatarsal tubercles. Feet lacking webbing or lateral fringes.
</paragraph>
<paragraph id="3E1936FCFFDDFFB7FF53F92EFAD2F8BF" blockId="3.[151,1437,1241,1987]" pageId="3" pageNumber="62">
<emphasis id="0CD2EAEEFFDDFFB7FF53F92EFEFAF926" bold="true" box="[197,342,1720,1746]" pageId="3" pageNumber="62">Coloration:</emphasis>
In life (
<figureCitation id="A69D2A79FFDDFFB7FE28F92FFDA9F927" box="[446,517,1721,1747]" captionStart="FIGURE 1" captionStartId="3.[151,255,1131,1155]" captionTargetBox="[256,1395,181,1034]" captionTargetId="figure@3.[178,1409,146,1114]" captionTargetPageId="3" captionText="FIGURE 1. Ranitomeya tolimense, new species. Lateral view showing color pattern. CZUT-A 989 adult female, SVL 17.94 mm. Photo: Manuel H. Bernal." httpUri="https://zenodo.org/record/179598/files/figure.png" pageId="3" pageNumber="62">Fig. 1</figureCitation>
), head yellow-bronze, upper jaw and anteroproximal part of the upper arm bright yellow; venter, throat, chest, sides of the body, hind limbs and hands brown or black brown. A dark stripe extends from the eyes to the arm insertion. Throat and venter with or without small bluish green blotches; flanks, groin and thighs (anterior and ventral) with or without a few bluish spots. Iris dark brown.
</paragraph>
<paragraph id="3E1936FCFFDDFFB7FF53F8CFFB2CF837" blockId="3.[151,1437,1241,1987]" pageId="3" pageNumber="62">In 70 % alcohol the dark or dark brown surfaces of the body maintain their color, as well as the bluish green of blotches or spots of ventral surfaces. However, the bronze color of head become dark whereas the yellow color of the upper arm and jaw change to a pale white and later to a dark gray.</paragraph>
<caption id="6AD96674FFDAFFB0FF01FAEFFAE3FA64" box="[151,1359,1401,1425]" httpUri="https://zenodo.org/record/179599/files/figure.png" pageId="4" pageNumber="63" targetBox="[185,1393,200,1365]" targetPageId="4">
<paragraph id="3E1936FCFFDAFFB0FF01FAEFFAE3FA64" blockId="4.[151,1359,1401,1425]" box="[151,1359,1401,1425]" pageId="4" pageNumber="63">
<emphasis id="0CD2EAEEFFDAFFB0FF01FAEFFEB5FA65" bold="true" box="[151,281,1401,1425]" pageId="4" pageNumber="63">FIGURE 2.</emphasis>
Male nurse frog (CZUT-A 990) with a tadpole over his back at the type locality. Photo: Victor Luna.
</paragraph>
</caption>
<paragraph id="3E1936FCFFDAFFB0FF50FA55FF46F9A2" blockId="4.[151,1437,1475,2022]" pageId="4" pageNumber="63">
<emphasis id="0CD2EAEEFFDAFFB0FF50FA55FE67FA29" bold="true" box="[198,459,1475,1501]" pageId="4" pageNumber="63">Sexual dimorphism:</emphasis>
The most remarkable external characteristics to differentiate adult males and females are the presence of vocal slits. Also, a wider disc on finger III and a smaller SVL was found in males (
<tableCitation id="73240347FFDAFFB0FF08F982FF57F9DA" box="[158,251,1556,1582]" captionStart="TABLE 2" captionStartId="5.[151,239,351,375]" captionTargetBox="[159,1380,408,817]" captionTargetPageId="5" captionText="TABLE 2. Size and proportions of adult Ranitomeya tolimense, New species sensu Myers &amp; Daly (1980)." httpUri="http://table.plazi.org/id/6AD96674FFDBFFB1FF01FEC9FAAFFE83" pageId="4" pageNumber="63" tableUuid="6AD96674FFDBFFB1FF01FEC9FAAFFE83">Table 2</tableCitation>
); however, there are a few data to be confident with this sexual difference. Testes and eggs are dark brown.
</paragraph>
</subSubSection>
<subSubSection id="76BC6577FFDAFFB3FF53F9F5FB89FE3E" lastPageId="7" lastPageNumber="66" pageId="4" pageNumber="63" type="distribution">
<paragraph id="3E1936FCFFDAFFB0FF53F9F5FB9FF8B2" blockId="4.[151,1437,1475,2022]" pageId="4" pageNumber="63">
<emphasis id="0CD2EAEEFFDAFFB0FF53F9F5FDC8F989" bold="true" box="[197,612,1635,1661]" pageId="4" pageNumber="63">Distribution and natural history:</emphasis>
So far,
<taxonomicName id="F9A64D7FFFDAFFB0FD54F9F2FCFFF989" box="[706,851,1636,1661]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFDAFFB0FD54F9F2FCFFF989" box="[706,851,1636,1661]" italics="true" pageId="4" pageNumber="63">R. tolimense</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="E11D885EFFDAFFB0FBEFF9F2FB07F98A" box="[1145,1195,1636,1662]" pageId="4" pageNumber="63">type</typeStatus>
locality on the eastern flank of the Cordillera Central, Municipio de Falan, Departamento de Tolima,
<collectingCountry id="46B1766CFFDAFFB0FBCDF91AFB7EF952" box="[1115,1234,1676,1702]" name="Colombia" pageId="4" pageNumber="63">Colombia</collectingCountry>
. Specimens were collected at an elevation of about
<quantity id="F95E9B19FFDAFFB0FDB2F922FDD1F93A" box="[548,637,1716,1742]" metricMagnitude="3" metricUnit="m" metricValue="1.8519999999999999" pageId="4" pageNumber="63" unit="m" value="1852.0">1852 m</quantity>
above sea level. They were found during day and the onset of night time in
<date id="4A18103CFFDAFFB0FF64F94AFEC7F902" box="[242,363,1756,1782]" pageId="4" pageNumber="63" value="2007-05">May 2007</date>
on the forest floor. On
<date id="4A18103CFFDAFFB0FD14F94AFC88F902" box="[642,804,1756,1782]" pageId="4" pageNumber="63" value="2007-05-19">19 May 2007</date>
, at 18:00 h (relative humidity: 75 %, air temperature: 20.4 ºC), we found a male nurse frog carrying a tadpole on his back on the ground near bromeliads and heliconias. When captured the frog expelled a milky secretion over his body (
<figureCitation id="A69D2A79FFDAFFB0FC4AF8BAFB88F8B2" box="[988,1060,1836,1862]" captionStart="FIGURE 2" captionStartId="4.[151,255,1401,1425]" captionTargetBox="[185,1393,200,1365]" captionTargetId="figure@4.[180,1408,194,1377]" captionTargetPageId="4" captionText="FIGURE 2. Male nurse frog (CZUT-A 990) with a tadpole over his back at the type locality. Photo: Victor Luna." httpUri="https://zenodo.org/record/179599/files/figure.png" pageId="4" pageNumber="63">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="3E1936FCFFDAFFB1FF53F8C2FACFFF2E" blockId="4.[151,1437,1475,2022]" lastBlockId="5.[151,1436,152,298]" lastPageId="5" lastPageNumber="64" pageId="4" pageNumber="63">
The
<typeStatus id="E11D885EFFDAFFB0FF6DF8C2FE82F89A" box="[251,302,1876,1902]" pageId="4" pageNumber="63">type</typeStatus>
locality of
<taxonomicName id="F9A64D7FFFDAFFB0FE2FF8C2FDE7F899" box="[441,587,1876,1901]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="63" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFDAFFB0FE2FF8C2FDE7F899" box="[441,587,1876,1901]" italics="true" pageId="4" pageNumber="63">R. tolimense</emphasis>
</taxonomicName>
is a small patch of a secondary forest dominated by trees with a maximum height of
<quantity id="F95E9B19FFDAFFB0FEDCF8EAFE02F862" box="[330,430,1916,1942]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="4" pageNumber="63" unit="m" value="5.0" valueMax="6.0" valueMin="4.0">4 to 6 m</quantity>
, covered by epiphytic plant, mainly of the families
<taxonomicName id="F9A64D7FFFDAFFB0FB91F8EAFBC6F862" box="[1031,1130,1916,1942]" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" pageId="4" pageNumber="63" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
,
<taxonomicName id="F9A64D7FFFDAFFB0FBE0F8EAFABBF862" box="[1142,1303,1916,1942]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="4" pageNumber="63" phylum="Tracheophyta" rank="family">Bromeliaceae</taxonomicName>
and
<taxonomicName id="F9A64D7FFFDAFFB0FAD9F8EAFF47F84A" class="Liliopsida" family="Orchidaceae" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="63" phylum="Tracheophyta" rank="family">Orchidaceae</taxonomicName>
. The canopy is dense allowing for little penetration of light. Humidity in the area is maintained by the constant cloud cover and frequent rainfall. The forest floor is covered by leaf litter, fallen tree branches and trunks, house to fungi, mosses and lichens. There are also smaller trees, palms and tree ferns. The mean temperature is about 19 ºC and the annual precipitation is between
<quantity id="F95E9B19FFDBFFB1FCEDFF56FB98FF2E" box="[891,1076,192,218]" metricMagnitude="0" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="5" pageNumber="64" unit="mm" value="2750.0" valueMax="3000.0" valueMin="2500.0">2500–3000 mm</quantity>
per year (
<bibRefCitation id="5A374B0DFFDBFFB1FB3CFF56FAFBFF2E" author="Gallego" box="[1194,1367,192,218]" pageId="5" pageNumber="67" refString="Gallego, O. (2003) Distribucion ecologica de la herpetofauna del Municipio de Falan, Tolima. Popayan-Cauca 2003. Trabajo de Grado (Ecologo). Fundacion Universitaria de Popayan." type="book chapter" year="2003">Gallego, 2003</bibRefCitation>
).
</paragraph>
<paragraph id="3E1936FCFFDBFFB1FF53FF7EFAC1FEDE" blockId="5.[151,1436,152,298]" pageId="5" pageNumber="64">
The general area is dominated by farms with coffee plantations, most likely not suitable habitat for
<emphasis id="0CD2EAEEFFDBFFB1FADDFF7EFF44FEDD" italics="true" pageId="5" pageNumber="64">
<taxonomicName id="F9A64D7FFFDBFFB1FADDFF7EFF48FEDD" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="tolimense">R. tolimense</taxonomicName>
.
</emphasis>
We recommend further studies to assess the distribution of this species and its conservation status.
</paragraph>
<caption id="6AD96674FFDBFFB1FF01FEC9FAAFFE83" ID-Table-UUID="6AD96674FFDBFFB1FF01FEC9FAAFFE83" box="[151,1283,351,375]" httpUri="http://table.plazi.org/id/6AD96674FFDBFFB1FF01FEC9FAAFFE83" pageId="5" pageNumber="64" targetBox="[159,1380,408,817]" targetIsTable="true" targetPageId="5">
<paragraph id="3E1936FCFFDBFFB1FF01FEC9FAAFFE83" blockId="5.[151,1283,351,375]" box="[151,1283,351,375]" pageId="5" pageNumber="64">
<emphasis id="0CD2EAEEFFDBFFB1FF01FEC9FEA4FE83" bold="true" box="[151,264,351,375]" pageId="5" pageNumber="64">TABLE 2.</emphasis>
Size and proportions of adult
<taxonomicName id="F9A64D7FFFDBFFB1FDDAFEF6FCD4FE83" authority="New" authorityName="New" box="[588,888,352,375]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFDBFFB1FDDAFEF6FC94FE83" box="[588,824,352,375]" italics="true" pageId="5" pageNumber="64">Ranitomeya tolimense</emphasis>
,
<taxonomicNameLabel id="17E15795FFDBFFB1FCD0FEF6FCD4FE83" box="[838,888,352,375]" pageId="5" pageNumber="64" rank="species">New</taxonomicNameLabel>
</taxonomicName>
species
<emphasis id="0CD2EAEEFFDBFFB1FC45FEF6FBA3FE83" box="[979,1039,352,375]" italics="true" pageId="5" pageNumber="64">sensu</emphasis>
Myers &amp; Daly (1980).
</paragraph>
</caption>
<paragraph id="3E1936FCFFDBFFB1FF09FE0EFAE5FCC5" pageId="5" pageNumber="64">
<table id="4CA6C45CFFDB004BFF09FE0EFAC8FCC5" box="[159,1380,408,817]" gridcols="6" gridrows="6" pageId="5" pageNumber="64">
<tr id="809634BEFFDB004BFF09FE0EFAC8FE5B" box="[159,1380,408,431]" gridrow="0" pageId="5" pageNumber="64">
<th id="C3475DC2FFDB004BFF09FE0EFE97FE5B" box="[159,315,408,431]" gridcol="0" gridrow="0" pageId="5" pageNumber="64">Measurements</th>
<th id="C3475DC2FFDB004BFE15FE0EFE46FE5B" box="[387,490,408,431]" gridcol="1" gridrow="0" pageId="5" pageNumber="64">N</th>
<th id="C3475DC2FFDB004BFDCCFE0EFCAEFE5B" box="[602,770,408,431]" gridcol="2" gridrow="0" pageId="5" pageNumber="64">Mean (+/- S.E.)</th>
<th id="C3475DC2FFDB004BFCDAFE0EFC17FE5B" box="[844,955,408,431]" gridcol="3" gridrow="0" pageId="5" pageNumber="64">Stand dev.</th>
<th id="C3475DC2FFDB004BFB83FE0EFBDFFE5B" box="[1045,1139,408,431]" gridcol="4" gridrow="0" pageId="5" pageNumber="64">C.V. (%)</th>
<th id="C3475DC2FFDB004BFB48FE0EFAC8FE5B" box="[1246,1380,408,431]" gridcol="5" gridrow="0" pageId="5" pageNumber="64">Range</th>
</tr>
<tr id="809634BEFFDB004BFF09FE54FAC8FE08" box="[159,1380,450,508]" gridrow="1" pageId="5" pageNumber="64">
<th id="C3475DC2FFDB004BFF09FE54FE97FE08" box="[159,315,450,508]" gridcol="0" gridrow="1" pageId="5" pageNumber="64">SVL</th>
<td id="C3475DC2FFDB004BFE15FE54FE46FE08" box="[387,490,450,508]" gridcol="1" gridrow="1" pageId="5" pageNumber="64">3 males 3 females</td>
<td id="C3475DC2FFDB004BFDCCFE54FCAEFE08" box="[602,770,450,508]" gridcol="2" gridrow="1" pageId="5" pageNumber="64">17.87 (0.11) 18.28 (0.22)</td>
<td id="C3475DC2FFDB004BFCDAFE54FC17FE08" box="[844,955,450,508]" gridcol="3" gridrow="1" pageId="5" pageNumber="64">0.190 0.564</td>
<td id="C3475DC2FFDB004BFB83FE54FBDFFE08" box="[1045,1139,450,508]" gridcol="4" gridrow="1" pageId="5" pageNumber="64">1.06 3.06</td>
<td id="C3475DC2FFDB004BFB48FE54FAC8FE08" box="[1246,1380,450,508]" gridcol="5" gridrow="1" pageId="5" pageNumber="64">17.66–18.02 17.82–18.91</td>
</tr>
<tr id="809634BEFFDB004BFF09FD86FAC8FDBD" box="[159,1380,528,585]" gridrow="2" pageId="5" pageNumber="64">
<th id="C3475DC2FFDB004BFF09FD86FE97FDBD" box="[159,315,528,585]" gridcol="0" gridrow="2" pageId="5" pageNumber="64">TL/SVL</th>
<td id="C3475DC2FFDB004BFE15FD86FE46FDBD" box="[387,490,528,585]" gridcol="1" gridrow="2" pageId="5" pageNumber="64">3 males 3 females</td>
<td id="C3475DC2FFDB004BFDCCFD86FCAEFDBD" box="[602,770,528,585]" gridcol="2" gridrow="2" pageId="5" pageNumber="64">0.42 (0.02) 0.41 (0.01)</td>
<td id="C3475DC2FFDB004BFCDAFD86FC17FDBD" box="[844,955,528,585]" gridcol="3" gridrow="2" pageId="5" pageNumber="64">0.010 0.005</td>
<td id="C3475DC2FFDB004BFB83FD86FBDFFDBD" box="[1045,1139,528,585]" gridcol="4" gridrow="2" pageId="5" pageNumber="64">4.76 2.42</td>
<td id="C3475DC2FFDB004BFB48FD86FAC8FDBD" box="[1246,1380,528,585]" gridcol="5" gridrow="2" pageId="5" pageNumber="64">0.45–0.40 0.40–0.41</td>
</tr>
<tr id="809634BEFFDB004BFF09FDCBFAC8FD63" box="[159,1380,605,663]" gridrow="3" pageId="5" pageNumber="64">
<th id="C3475DC2FFDB004BFF09FDCBFE97FD63" box="[159,315,605,663]" gridcol="0" gridrow="3" pageId="5" pageNumber="64">HL/SVL</th>
<td id="C3475DC2FFDB004BFE15FDCBFE46FD63" box="[387,490,605,663]" gridcol="1" gridrow="3" pageId="5" pageNumber="64">3 males 3 females</td>
<td id="C3475DC2FFDB004BFDCCFDCBFCAEFD63" box="[602,770,605,663]" gridcol="2" gridrow="3" pageId="5" pageNumber="64">0.30 (0.00) 0.30 (0.01)</td>
<td id="C3475DC2FFDB004BFCDAFDCBFC17FD63" box="[844,955,605,663]" gridcol="3" gridrow="3" pageId="5" pageNumber="64">0.011 0.002</td>
<td id="C3475DC2FFDB004BFB83FDCBFBDFFD63" box="[1045,1139,605,663]" gridcol="4" gridrow="3" pageId="5" pageNumber="64">3.66 0.66</td>
<td id="C3475DC2FFDB004BFB48FDCBFAC8FD63" box="[1246,1380,605,663]" gridcol="5" gridrow="3" pageId="5" pageNumber="64">0.28–0.32 0.30–0.31</td>
</tr>
<tr id="809634BEFFDB004BFF09FD3CFAC8FD10" box="[159,1380,682,740]" gridrow="4" pageId="5" pageNumber="64">
<th id="C3475DC2FFDB004BFF09FD3CFE97FD10" box="[159,315,682,740]" gridcol="0" gridrow="4" pageId="5" pageNumber="64">NED/EL</th>
<td id="C3475DC2FFDB004BFE15FD3CFE46FD10" box="[387,490,682,740]" gridcol="1" gridrow="4" pageId="5" pageNumber="64">3 males 3 females</td>
<td id="C3475DC2FFDB004BFDCCFD3CFCAEFD10" box="[602,770,682,740]" gridcol="2" gridrow="4" pageId="5" pageNumber="64">0.65 (0.04) 0.64 (0.02)</td>
<td id="C3475DC2FFDB004BFCDAFD3CFC17FD10" box="[844,955,682,740]" gridcol="3" gridrow="4" pageId="5" pageNumber="64">0.0.8 0.04</td>
<td id="C3475DC2FFDB004BFB83FD3CFBDFFD10" box="[1045,1139,682,740]" gridcol="4" gridrow="4" pageId="5" pageNumber="64">12.30 6.25</td>
<td id="C3475DC2FFDB004BFB48FD3CFAC8FD10" box="[1246,1380,682,740]" gridcol="5" gridrow="4" pageId="5" pageNumber="64">0.58–0.75 0.60–0.68</td>
</tr>
<tr id="809634BEFFDB004BFF09FD6EFAC8FCC5" box="[159,1380,760,817]" gridrow="5" pageId="5" pageNumber="64">
<th id="C3475DC2FFDB004BFF09FD6EFE97FCC5" box="[159,315,760,817]" gridcol="0" gridrow="5" pageId="5" pageNumber="64">WTFD/WTF</th>
<td id="C3475DC2FFDB004BFE15FD6EFE46FCC5" box="[387,490,760,817]" gridcol="1" gridrow="5" pageId="5" pageNumber="64">3 males 3 females</td>
<td id="C3475DC2FFDB004BFDCCFD6EFCAEFCC5" box="[602,770,760,817]" gridcol="2" gridrow="5" pageId="5" pageNumber="64">1.86 (0.11) 1.59 (0.04)</td>
<td id="C3475DC2FFDB004BFCDAFD6EFC17FCC5" box="[844,955,760,817]" gridcol="3" gridrow="5" pageId="5" pageNumber="64">0.20 0.07</td>
<td id="C3475DC2FFDB004BFB83FD6EFBDFFCC5" box="[1045,1139,760,817]" gridcol="4" gridrow="5" pageId="5" pageNumber="64">10.75 4.40</td>
<td id="C3475DC2FFDB004BFB48FD6EFAC8FCC5" box="[1246,1380,760,817]" gridcol="5" gridrow="5" pageId="5" pageNumber="64">1.63–2.00 1.52–1.66</td>
</tr>
</table>
</paragraph>
<caption id="6AD96674FFDBFFB1FF01F835FBF2F84F" box="[151,1118,1955,1979]" httpUri="https://zenodo.org/record/179600/files/figure.png" pageId="5" pageNumber="64" targetBox="[292,1307,926,1900]" targetPageId="5">
<paragraph id="3E1936FCFFDBFFB1FF01F835FBF2F84F" blockId="5.[151,1118,1955,1979]" box="[151,1118,1955,1979]" pageId="5" pageNumber="64">
<emphasis id="0CD2EAEEFFDBFFB1FF01F835FEB5F84F" bold="true" box="[151,281,1955,1979]" pageId="5" pageNumber="64">FIGURE 3.</emphasis>
Map of Colombia showing (arrow) the type locality of
<taxonomicName id="F9A64D7FFFDBFFB1FCFDF832FBF7F84F" box="[875,1115,1956,1979]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="5" pageNumber="64" phylum="Chordata" rank="species" species="tolimense">Ranitomeya tolimense</taxonomicName>
.
</paragraph>
</caption>
<caption id="6AD96674FFD8FFB2FF01FC75FB19FC0F" box="[151,1205,995,1019]" httpUri="https://zenodo.org/record/179601/files/figure.png" pageId="6" pageNumber="65" targetBox="[237,1354,192,950]" targetPageId="6">
<paragraph id="3E1936FCFFD8FFB2FF01FC75FB19FC0F" blockId="6.[151,1205,995,1019]" box="[151,1205,995,1019]" pageId="6" pageNumber="65">
<emphasis id="0CD2EAEEFFD8FFB2FF01FC75FEB5FC0F" bold="true" box="[151,281,995,1019]" pageId="6" pageNumber="65">FIGURE 4.</emphasis>
Ventral view of hand and foot of
<taxonomicName id="F9A64D7FFFD8FFB2FD17FC72FCC0FC0F" box="[641,876,996,1019]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFD8FFB2FD17FC72FCC0FC0F" box="[641,876,996,1019]" italics="true" pageId="6" pageNumber="65">Ranitomeya tolimense</emphasis>
</taxonomicName>
(CZUT-A 989). Scale = 1mm.
</paragraph>
</caption>
<paragraph id="3E1936FCFFD8FFB2FF53FBDAFC1CFB7A" blockId="6.[151,1436,1100,2006]" pageId="6" pageNumber="65">
<emphasis id="0CD2EAEEFFD8FFB2FF53FBDAFE9EFB92" bold="true" box="[197,306,1100,1126]" pageId="6" pageNumber="65">Tadpole:</emphasis>
The following description is based on a larva (CZUT-A: 991) obtained from the back of a nurse frog male (
<figureCitation id="A69D2A79FFD8FFB2FE8EFBE2FEC7FB7A" box="[280,363,1140,1166]" captionStart="FIGURE 2" captionStartId="4.[151,255,1401,1425]" captionTargetBox="[185,1393,200,1365]" captionTargetId="figure@4.[180,1408,194,1377]" captionTargetPageId="4" captionText="FIGURE 2. Male nurse frog (CZUT-A 990) with a tadpole over his back at the type locality. Photo: Victor Luna." httpUri="https://zenodo.org/record/179599/files/figure.png" pageId="6" pageNumber="65">Figs. 2</figureCitation>
,
<figureCitation id="A69D2A79FFD8FFB2FEE1FBE2FE24FB7A" box="[375,392,1140,1166]" captionStart="FIGURE 5" captionStartId="7.[151,255,1562,1586]" captionTargetBox="[196,1346,659,1561]" captionTargetId="figure@7.[196,1390,636,1586]" captionTargetPageId="7" captionText="FIGURE 5. Tadpole of Ranitomeya tolimense (CZUT-A 991). (A) Lateral view, stage 25 (Gosner, 1960). Scale equals 5 mm. (B) Mouth parts. Scale equals 1 mm." httpUri="https://zenodo.org/record/179602/files/figure.png" pageId="6" pageNumber="65">5</figureCitation>
). The larva is in stage 25 sensu
<bibRefCitation id="5A374B0DFFD8FFB2FD69FBE2FC01FB7A" author="Gosner" box="[767,941,1140,1166]" pageId="6" pageNumber="67" refString="Gosner, K. L. (1960) A simplified table for staging anuran embryos and larvae with notes on identification. Herpetologica, 16, 183 - 190." type="journal article" year="1960">Gosner (1960)</bibRefCitation>
.
</paragraph>
<paragraph id="3E1936FCFFD8FFB2FF53FB0AFC86FB2A" blockId="6.[151,1436,1100,2006]" pageId="6" pageNumber="65">Measurements in millimeters are: head-body length 4.4; total length 13.1; tail length 8.7; greatest depth from upper edge dorsal fin to lower edge ventral fin 1.3.</paragraph>
<paragraph id="3E1936FCFFD8FFB2FF53FB7AFA3DFA8A" blockId="6.[151,1436,1100,2006]" pageId="6" pageNumber="65">
The tadpole has a globular shape when viewed from above, with maximum body width of 61 % of head body length. The head and body are depressed, width much greater than depth; eyes and nostrils are dorsal, directed dorsolaterally; the spiracle is sinistral and low and the anus dextral (
<figureCitation id="A69D2A79FFD8FFB2FBB6FAAAFBCAFAA2" box="[1056,1126,1340,1366]" captionStart="FIGURE 5" captionStartId="7.[151,255,1562,1586]" captionTargetBox="[196,1346,659,1561]" captionTargetId="figure@7.[196,1390,636,1586]" captionTargetPageId="7" captionText="FIGURE 5. Tadpole of Ranitomeya tolimense (CZUT-A 991). (A) Lateral view, stage 25 (Gosner, 1960). Scale equals 5 mm. (B) Mouth parts. Scale equals 1 mm." httpUri="https://zenodo.org/record/179602/files/figure.png" pageId="6" pageNumber="65">Fig. 5</figureCitation>
A). The tail is 64 % of the total length and its maximum tail height is only 9.9 % of total length. The tip of the tail is rounded (
<figureCitation id="A69D2A79FFD8FFB2FAB3FAF2FAC7FA8A" box="[1317,1387,1380,1406]" captionStart="FIGURE 5" captionStartId="7.[151,255,1562,1586]" captionTargetBox="[196,1346,659,1561]" captionTargetId="figure@7.[196,1390,636,1586]" captionTargetPageId="7" captionText="FIGURE 5. Tadpole of Ranitomeya tolimense (CZUT-A 991). (A) Lateral view, stage 25 (Gosner, 1960). Scale equals 5 mm. (B) Mouth parts. Scale equals 1 mm." httpUri="https://zenodo.org/record/179602/files/figure.png" pageId="6" pageNumber="65">Fig. 5</figureCitation>
A).
</paragraph>
<paragraph id="3E1936FCFFD8FFB2FF53FA1AFC4CF8FA" blockId="6.[151,1436,1100,2006]" pageId="6" pageNumber="65">
The mouth is directed anteroventrally. There are two upper and three lower tooth rows, with the second upper row being broadly broken above the beak (
<figureCitation id="A69D2A79FFD8FFB2FD7FFA22FC9EFA3A" box="[745,818,1460,1486]" captionStart="FIGURE 5" captionStartId="7.[151,255,1562,1586]" captionTargetBox="[196,1346,659,1561]" captionTargetId="figure@7.[196,1390,636,1586]" captionTargetPageId="7" captionText="FIGURE 5. Tadpole of Ranitomeya tolimense (CZUT-A 991). (A) Lateral view, stage 25 (Gosner, 1960). Scale equals 5 mm. (B) Mouth parts. Scale equals 1 mm." httpUri="https://zenodo.org/record/179602/files/figure.png" pageId="6" pageNumber="65">Fig. 5</figureCitation>
B). The other rows are complete, although the third posterior one row is weakly pronounced, probably due to an ontogenetic difference with respect to other tadpoles found in the locality. The beak is massive, with serrate cutting edges. The oral disc is laterally indented and its posterior edge has a broad median gap (
<figureCitation id="A69D2A79FFD8FFB2FD5CF9BAFCBEF9B2" box="[714,786,1580,1606]" captionStart="FIGURE 5" captionStartId="7.[151,255,1562,1586]" captionTargetBox="[196,1346,659,1561]" captionTargetId="figure@7.[196,1390,636,1586]" captionTargetPageId="7" captionText="FIGURE 5. Tadpole of Ranitomeya tolimense (CZUT-A 991). (A) Lateral view, stage 25 (Gosner, 1960). Scale equals 5 mm. (B) Mouth parts. Scale equals 1 mm." httpUri="https://zenodo.org/record/179602/files/figure.png" pageId="6" pageNumber="65">Fig. 5</figureCitation>
B). This gap is shared with other related species of the genus
<taxonomicName id="F9A64D7FFFD8FFB2FF7EF9C2FED8F999" box="[232,372,1620,1645]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="0CD2EAEEFFD8FFB2FF7EF9C2FED8F999" box="[232,372,1620,1645]" italics="true" pageId="6" pageNumber="65">Ranitomeya</emphasis>
</taxonomicName>
as mentioned above and perhaps represents a synapomorphy (
<bibRefCitation id="5A374B0DFFD8FFB2FB10F9C2FA22F99A" author="Myers" box="[1158,1422,1620,1646]" pageId="6" pageNumber="68" refString="Myers, C. W. &amp; Daly J. (1980) Taxonomy and ecology of Dendrobates bombetes, a new Andean poison frog with new skin toxins. American Museum Novitates, New York, 2694, 1 - 23." type="journal article" year="1980">Myers &amp; Daly, 1980</bibRefCitation>
), although
<taxonomicName id="F9A64D7FFFD8FFB2FE92F9EAFE38F961" box="[260,404,1660,1685]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="bombetes">
<emphasis id="0CD2EAEEFFD8FFB2FE92F9EAFE38F961" box="[260,404,1660,1685]" italics="true" pageId="6" pageNumber="65">R. bombetes</emphasis>
</taxonomicName>
has a somewhat broader median gap and larger and fewer labial papillae than
<emphasis id="0CD2EAEEFFD8FFB2FABFF9EAFEFFF949" italics="true" pageId="6" pageNumber="65">
<taxonomicName id="F9A64D7FFFD8FFB2FABFF9EAFA34F961" box="[1321,1432,1660,1685]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="abdita">R. abdita</taxonomicName>
,
<taxonomicName id="F9A64D7FFFD8FFB2FF01F932FEFFF949" authority="Myers &amp; Daly, 1980" authorityName="Myers &amp; Daly" authorityYear="1980" box="[151,339,1700,1725]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="opisthomelas">R. opisthomelas</taxonomicName>
</emphasis>
(
<bibRefCitation id="5A374B0DFFD8FFB2FEF5F932FDF1F94A" author="Myers" box="[355,605,1700,1726]" pageId="6" pageNumber="68" refString="Myers, C. W. &amp; Daly J. (1980) Taxonomy and ecology of Dendrobates bombetes, a new Andean poison frog with new skin toxins. American Museum Novitates, New York, 2694, 1 - 23." type="journal article" year="1980">Myers &amp; Daly, 1980</bibRefCitation>
),
<taxonomicName id="F9A64D7FFFD8FFB2FDE5F932FAA2F94A" authority="Ruiz-Carranza &amp; Ramirez-Pinilla, 1992" authorityName="Ruiz-Carranza &amp; Ramirez-Pinilla" authorityYear="1992" box="[627,1294,1700,1726]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="virolinensis">
<emphasis id="0CD2EAEEFFD8FFB2FDE5F932FCB0F949" box="[627,796,1700,1725]" italics="true" pageId="6" pageNumber="65">R. virolinensis</emphasis>
(
<bibRefCitation id="5A374B0DFFD8FFB2FCBAF932FAAAF94A" author="Ruiz-Carranza" box="[812,1286,1700,1726]" pageId="6" pageNumber="68" refString="Ruiz-Carranza, P &amp; Ramirez-Pinilla M. P. (1992) Una nueva especie de Minyobates (Anura: Dendrobatidae) de Colombia, Lozania, 61, 1 - 16." type="journal article" year="1992">Ruiz-Carranza &amp; Ramirez-Pinilla, 1992</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="F9A64D7FFFD8FFB2FADFF932FF4CF911" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFD8FFB2FADFF932FF4CF911" italics="true" pageId="6" pageNumber="65">R. tolimense</emphasis>
</taxonomicName>
. Furthermore,
<taxonomicName id="F9A64D7FFFD8FFB2FE05F95AFD89F911" box="[403,549,1740,1765]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFD8FFB2FE05F95AFD89F911" box="[403,549,1740,1765]" italics="true" pageId="6" pageNumber="65">R. tolimense</emphasis>
</taxonomicName>
shares with
<taxonomicName id="F9A64D7FFFD8FFB2FD2BF95AFCCBF911" box="[701,871,1740,1765]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="virolinensis">
<emphasis id="0CD2EAEEFFD8FFB2FD2BF95AFCCBF911" box="[701,871,1740,1765]" italics="true" pageId="6" pageNumber="65">R. virolinensis</emphasis>
</taxonomicName>
one to two rows of labial papillae (
<figureCitation id="A69D2A79FFD8FFB2FA84F95AFAD4F912" box="[1298,1400,1740,1766]" captionStart="FIGURE 5" captionStartId="7.[151,255,1562,1586]" captionTargetBox="[196,1346,659,1561]" captionTargetId="figure@7.[196,1390,636,1586]" captionTargetPageId="7" captionText="FIGURE 5. Tadpole of Ranitomeya tolimense (CZUT-A 991). (A) Lateral view, stage 25 (Gosner, 1960). Scale equals 5 mm. (B) Mouth parts. Scale equals 1 mm." httpUri="https://zenodo.org/record/179602/files/figure.png" pageId="6" pageNumber="65">Figure 5</figureCitation>
B), whereas the illustrations of the other species shown apparently just one.
</paragraph>
<paragraph id="3E1936FCFFD8FFB2FF53F88AFDFAF872" blockId="6.[151,1436,1100,2006]" pageId="6" pageNumber="65">Life coloration was brown to dark brown, lighter on ventral side, turning clear towards the tail tip. In 10 % formalin, the body and tail are pale brown, uniformly pigmented, including fins. The tail tip is somewhat transparent and the venter translucent.</paragraph>
<paragraph id="3E1936FCFFD8FFB3FF50F802FE68FF2E" blockId="6.[151,1436,1100,2006]" lastBlockId="7.[151,1436,152,658]" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65">
<emphasis id="0CD2EAEEFFD8FFB2FF50F802FEC6F85A" bold="true" box="[198,362,1940,1966]" pageId="6" pageNumber="65">Vocalization:</emphasis>
The advertisement call of
<taxonomicName id="F9A64D7FFFD8FFB2FD32F802FC9AF859" box="[676,822,1940,1965]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="65" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFD8FFB2FD32F802FC9AF859" box="[676,822,1940,1965]" italics="true" pageId="6" pageNumber="65">R. tolimense</emphasis>
</taxonomicName>
(
<figureCitation id="A69D2A79FFD8FFB2FCD3F802FC27F85A" box="[837,907,1940,1966]" captionStart="FIGURE 6" captionStartId="8.[151,255,1340,1364]" captionTargetBox="[271,1264,235,1281]" captionTargetId="figure@8.[244,1344,194,1316]" captionTargetPageId="8" captionText="FIGURE 6. Advertisement call of Ranitomeya tolimense. A. Audiospectrogram of a call with three notes. B. Oscillogram of the three notes showing the pulses." httpUri="https://zenodo.org/record/179603/files/figure.png" pageId="6" pageNumber="65">Fig. 6</figureCitation>
) is a series of short buzzes of 0.84–0.99 seconds (N = 6). The call is pulsed throughout its length with a dominant frequency of about 4.73–5.22 kHz. The call is heard as a short, soft cricket-like buzz and sounds similar to the syntopic
<emphasis id="0CD2EAEEFFD9FFB3FBD2FF0EFA30FF45" box="[1092,1436,152,177]" italics="true" pageId="7" pageNumber="66">
<taxonomicName id="F9A64D7FFFD9FFB3FBD2FF0EFA34FF45" box="[1092,1432,152,177]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="dorisswansonae">Ranitomeya dorisswansonae</taxonomicName>
,
</emphasis>
although not as loud as it.
</paragraph>
<paragraph id="3E1936FCFFD9FFB3FF50FF7EFB89FE3E" blockId="7.[151,1436,152,658]" pageId="7" pageNumber="66">
The buzz call (sensu the defined of
<bibRefCitation id="5A374B0DFFD9FFB3FDF8FF7EFCFCFEF6" author="Lotters" box="[622,848,232,258]" pageId="7" pageNumber="67" refString="Lotters, S., Reichle, S. &amp; Jungfer, K. H. (2003) Advertisement calls of Neotropical poison frogs (Amphibia: Dendrobatidae) of the genera Colostethus, Dendrobate s, and Epipedobates, with notes on dendrobatid call classification. Journal of Natural History, 37, 1899 - 1911." type="journal article" year="2003">
Lötters
<emphasis id="0CD2EAEEFFD9FFB3FD5FFF7EFCA7FEF5" box="[713,779,232,257]" italics="true" pageId="7" pageNumber="66">et al.,</emphasis>
2003
</bibRefCitation>
) as described herein for
<taxonomicName id="F9A64D7FFFD9FFB3FBE0FF7EFAABFEF5" box="[1142,1287,232,257]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFD9FFB3FBE0FF7EFAABFEF5" box="[1142,1287,232,257]" italics="true" pageId="7" pageNumber="66">R. tolimense</emphasis>
</taxonomicName>
is similar to buzz calls described to species considered sharing larval characters with the new species, i.e.
<taxonomicName id="F9A64D7FFFD9FFB3FB72FE86FAD8FEDD" box="[1252,1396,272,297]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="bombetes">
<emphasis id="0CD2EAEEFFD9FFB3FB72FE86FAD8FEDD" box="[1252,1396,272,297]" italics="true" pageId="7" pageNumber="66">R. bombetes</emphasis>
</taxonomicName>
,
<taxonomicName id="F9A64D7FFFD9FFB3FA15FE86FDE8FEA6" authority="Myers &amp; Daly, 1980" authorityName="Myers &amp; Daly" authorityYear="1980" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="opisthomelas">
<emphasis id="0CD2EAEEFFD9FFB3FA15FE86FE9EFEA5" italics="true" pageId="7" pageNumber="66">R. opisthomelas</emphasis>
(
<bibRefCitation id="5A374B0DFFD9FFB3FED4FEAEFD97FEA6" author="Myers" box="[322,571,312,338]" pageId="7" pageNumber="68" refString="Myers, C. W. &amp; Daly J. (1980) Taxonomy and ecology of Dendrobates bombetes, a new Andean poison frog with new skin toxins. American Museum Novitates, New York, 2694, 1 - 23." type="journal article" year="1980">Myers &amp; Daly, 1980</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="F9A64D7FFFD9FFB3FDC6FEAEFC4FFEA6" authority="Amezquita, 1995" authorityName="Amezquita" authorityYear="1995" box="[592,995,312,338]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="virolinensis">
<emphasis id="0CD2EAEEFFD9FFB3FDC6FEAEFD54FEA5" box="[592,760,312,337]" italics="true" pageId="7" pageNumber="66">R. virolinensis</emphasis>
(
<bibRefCitation id="5A374B0DFFD9FFB3FC9CFEAEFC76FEA6" author="Amezquita" box="[778,986,312,338]" pageId="7" pageNumber="67" refString="Amezquita, A. (1995) Nachzucht von Minyobates virolinensis (Dendrobatidae) im Terrarium. Elaphe, Neue Folge, 3, 14 - 16." type="journal article" year="1995">Amézquita, 1995</bibRefCitation>
)
</taxonomicName>
, and likely
<taxonomicName id="F9A64D7FFFD9FFB3FBFBFEAEFF73FE8E" authority="Myers &amp; Daly, 1980" authorityName="Myers &amp; Daly" authorityYear="1980" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="adbita">
<emphasis id="0CD2EAEEFFD9FFB3FBFBFEAEFB76FEA5" box="[1133,1242,312,337]" italics="true" pageId="7" pageNumber="66">R. adbita</emphasis>
(
<bibRefCitation id="5A374B0DFFD9FFB3FB7CFEAEFF7AFE8E" author="Myers" pageId="7" pageNumber="68" refString="Myers, C. W. &amp; Daly J. (1980) Taxonomy and ecology of Dendrobates bombetes, a new Andean poison frog with new skin toxins. American Museum Novitates, New York, 2694, 1 - 23." type="journal article" year="1980">Myers &amp; Daly, 1980</bibRefCitation>
)
</taxonomicName>
. We will not rule out but can’t claim here that similarities in advertisement calls are due to close phylogenetic relationships. First, buzz calls have been reported in other dendrobatid frog species (see
<bibRefCitation id="5A374B0DFFD9FFB3FB69FE1EFF7AFE3E" author="Lotters" pageId="7" pageNumber="67" refString="Lotters, S., Reichle, S. &amp; Jungfer, K. H. (2003) Advertisement calls of Neotropical poison frogs (Amphibia: Dendrobatidae) of the genera Colostethus, Dendrobate s, and Epipedobates, with notes on dendrobatid call classification. Journal of Natural History, 37, 1899 - 1911." type="journal article" year="2003">
Lötters
<emphasis id="0CD2EAEEFFD9FFB3FACCFE1EFA21FE55" box="[1370,1421,392,417]" italics="true" pageId="7" pageNumber="66">et al</emphasis>
., 2003
</bibRefCitation>
), and second, anuran vocalizations are considered highly adaptive traits.
</paragraph>
</subSubSection>
<subSubSection id="76BC6577FFD9FFB3FF50FE41FB52FD66" pageId="7" pageNumber="66" type="discussion">
<paragraph id="3E1936FCFFD9FFB3FF50FE41FB52FD66" blockId="7.[151,1436,152,658]" pageId="7" pageNumber="66">
<emphasis id="0CD2EAEEFFD9FFB3FF50FE41FEEEFE05" bold="true" box="[198,322,471,497]" pageId="7" pageNumber="66">Remarks:</emphasis>
According to the possible larval synapomorphy suggested by
<bibRefCitation id="5A374B0DFFD9FFB3FBBAFE4EFA9CFE06" author="Myers" box="[1068,1328,472,498]" pageId="7" pageNumber="68" refString="Myers, C. W. &amp; Daly J. (1980) Taxonomy and ecology of Dendrobates bombetes, a new Andean poison frog with new skin toxins. American Museum Novitates, New York, 2694, 1 - 23." type="journal article" year="1980">Myers &amp; Daly (1980)</bibRefCitation>
and perhaps morphological and vocalization characters,
<taxonomicName id="F9A64D7FFFD9FFB3FD47FD96FCCDFDED" box="[721,865,512,537]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="tolimense">
<emphasis id="0CD2EAEEFFD9FFB3FD47FD96FCCDFDED" box="[721,865,512,537]" italics="true" pageId="7" pageNumber="66">R. tolimense</emphasis>
</taxonomicName>
is expected to be a member of an Andean monophyletic dendrobatid clade including
<emphasis id="0CD2EAEEFFD9FFB3FDC5FDBEFB9DFDB5" box="[595,1073,552,577]" italics="true" pageId="7" pageNumber="66">
<taxonomicName id="F9A64D7FFFD9FFB3FDC5FDBEFD68FDB5" box="[595,708,552,577]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="abdita">R. abdita</taxonomicName>
,
<taxonomicName id="F9A64D7FFFD9FFB3FD47FDBEFCC9FDB5" box="[721,869,552,577]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="bombetes">R. bombetes</taxonomicName>
,
<taxonomicName id="F9A64D7FFFD9FFB3FCE2FDBEFB9DFDB5" box="[884,1073,552,577]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="opisthomelas">R. opisthomelas</taxonomicName>
</emphasis>
and
<emphasis id="0CD2EAEEFFD9FFB3FBF8FDBEFA8CFDB5" box="[1134,1312,552,577]" italics="true" pageId="7" pageNumber="66">
<taxonomicName id="F9A64D7FFFD9FFB3FBF8FDBEFAB7FDB5" box="[1134,1307,552,577]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="virolinensis">R. virolinensis</taxonomicName>
.
</emphasis>
Larvae of other
<taxonomicName id="F9A64D7FFFD9FFB3FF4CFDC6FECAFD9D" box="[218,358,592,617]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="genus">
<emphasis id="0CD2EAEEFFD9FFB3FF4CFDC6FECAFD9D" box="[218,358,592,617]" italics="true" pageId="7" pageNumber="66">Ranitomeya</emphasis>
</taxonomicName>
lack a median gap in the papillae fringe on the lower edge of the oral disc (e.g.
<emphasis id="0CD2EAEEFFD9FFB3FA95FDC6FE12FD65" italics="true" pageId="7" pageNumber="66">
<taxonomicName id="F9A64D7FFFD9FFB3FA95FDC6FADBFD9D" box="[1283,1399,592,617]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="minuta">R. minuta</taxonomicName>
,
<taxonomicName id="F9A64D7FFFD9FFB3FA15FDC6FEADFD65" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="fulgurita">R. fulgurita</taxonomicName>
,
<taxonomicName id="F9A64D7FFFD9FFB3FE9AFDEEFE12FD65" box="[268,446,632,657]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="altobuyensis">R. altobuyensis</taxonomicName>
</emphasis>
) or remain unknown (
<emphasis id="0CD2EAEEFFD9FFB3FD51FDEEFB41FD65" box="[711,1261,632,657]" italics="true" pageId="7" pageNumber="66">
<taxonomicName id="F9A64D7FFFD9FFB3FD51FDEEFC98FD65" box="[711,820,632,657]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="viridis">R. viridis</taxonomicName>
,
<taxonomicName id="F9A64D7FFFD9FFB3FCD7FDEEFBA8FD65" box="[833,1028,632,657]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="daleswansoni">R. daleswansoni</taxonomicName>
,
<taxonomicName id="F9A64D7FFFD9FFB3FB99FDEEFB41FD65" box="[1039,1261,632,657]" class="Amphibia" family="Dendrobatidae" genus="Ranitomeya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="66" phylum="Chordata" rank="species" species="dorisswansonae">R. dorisswansonae</taxonomicName>
</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
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53
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3913739" ID-GBIF-Taxon="164811905" ID-Zenodo-Dep="3913739" LSID="urn:lsid:zoobank.org:act:7DD7E854-D86E-4952-AEC4-572383902672" httpUri="http://treatment.plazi.org/id/B60FE8BF8BA7B6E218068E13CCC26655" lastPageNumber="701" pageNumber="701" sourcePageUrl="https://www.biodiversitylibrary.org/page/727612">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:7DD7E854-D86E-4952-AEC4-572383902672" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Cicadellidae" genus="Anomia" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="electrica" status="spec. nov." zbkClass="Vermes" zbkKingdom="Animalia" zbkOrder="Testacea">Anomia electrica</taxonomicName>
[
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]
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<paragraph pageNumber="701">A. testa subrotunda flava laevi: altera convexo gibbosa.</paragraph>
<paragraph pageNumber="701">
<emphasis italics="true">Rumph. mus. t.</emphasis>
47.
<emphasis italics="true">f. L.</emphasis>
Ostreum electricum?
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<emphasis italics="true">Habitat in</emphasis>
M. Mediterraneo.
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Paragorgia stephencairnsi
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<taxonomicName id="4E85F25FB5CFBCEB7708DA7985CF57E7" class="Anthozoa" family="Paragorgiidae" genus="Paragorgia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paragorgia stephencairnsi" order="Alcyonacea" pageId="0" pageNumber="1" phylum="Cnidaria" rank="species" species="stephencairnsi">Paragorgia stephencairnsi</taxonomicName>
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, 2005: 57-60; figs 39-41.
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<paragraph id="9E03D79F45F204BB6BB34865C0A85222" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="C4D6B89EB33D314D6F556EE00A4A049A" pageId="0" pageNumber="1">No material in SBMNH collection (see Appendix 1: List of material examined).</paragraph>
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<paragraph id="774AFD9DCEFF53E0F99848AD81347125" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="C9F082B50E5A9F1737160B873E33DA2C" pageId="0" pageNumber="1">
A paratype specimen at NMNH (USNM 94437), identified as this species, was collected in proximity of the southern California Channel Islands, in an area lying close to the western boundary of the California Bight; range seems to extend northward to an area off British Columbia, Canada (location data for USNM 57982, the holotype; discussed in
<bibRefCitation id="B080F1138BCDF6A1D406DE546B7C9C48" author="Sanchez, JA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 72" title="Systematics of the bubblegum corals (Cnidaria: Octocorallia: Paragorgiidae) with description of new species from New Zealand and the Eastern Pacific." url="https://doi.org/10.11646/zootaxa.1014.1.1" volume="1014" year="2005">
<normalizedToken id="0B92D1DB002368C5450634B06A7D68E8" originalValue="Sánchez">Sanchez</normalizedToken>
2005
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). As this species was named in honor of Dr Stephen Cairns of the National Museum of Natural History, Smithsonian, it would be fitting to call this
<normalizedToken id="BDBF1565D573B34593FAE22147B30124" originalValue="“Stephen’s">&quot;Stephen's</normalizedToken>
bubblegum tree.&quot; Any similarities between this and other species in the genus were thoroughly discussed in
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<normalizedToken id="9F60B40DADE9DAEDBB6414A726697807" originalValue="Sánchez">Sanchez</normalizedToken>
(2005)
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.
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<paragraph pageId="0" pageNumber="27161">Collected from the Lewis and Clark County site (Table 1, Suppl. material 1)</paragraph>
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