The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini) Author Ratcliffe, Brett C. Author Cave, Ronald D. text Insecta Mundi 2010 2010-09-24 2010 139 1 15 journal article 10.5281/zenodo.5164903 1942-1354 5164903 Tomarus adoceteus Ratcliffe and Cave , new species (Fig. 10-12) Type Material. Holotype male, labeled “Roy Soc- CIG Expdn./Little Cayman, BWI/Pirates’ Point/ house light/ 31.7.1975 . R. R. Askew//BM 1981/342” (hand written)/ our red holotype label. Allotype fe- male with same data but date of 9.8.1975 and our red allotype label. Single male paratype and single female paratype with same data as allotype and with our yellow paratype labels. Female paratype labeled “WEST INDIES: Cayman Is./Grand Cay- man/ VI-1992 coll. P. Fitzgerald/blacklight trap” and with our yellow paratype label. Female paratype labeled “Grand Cayman Is. B. W. I./ Georgetown / 16 Sept. 1973 /E. J. Gerberg//at black light” and with our yellow paratype label. Holo- type and allotype deposited at The Natural His- tory Museum, London ( BMNH ). Paratypes depos- ited in the Florida State Collection of Arthropods, Gainesville, FL ( FSCA ) and the B. C. Ratcliffe col- lection, Lincoln, Nebraska ( BCRC ). Holotype . Male. Length 22.1 mm ; width 12.1 mm . Color black. Head : Frons coarsely rugose, rugae Figure 10-12. Tomarus adoceteus Ratcliffe and Cave , new transversely curved; vertex at center with small, species. 10) Habitus. 11-12) Parameres. subtriangular, smooth, shining area. Frontoclypeal region with low, transverse tubercle either side of middle, tubercles broadly separated, laterad of clypeal teeth. Clypeus with surface sparsely, trans- versely rugose; apex broadly truncate, with 2 small teeth, teeth weakly reflexed. Interocular width equals 3.5 transverse eye diameters. Antenna 10- segmented, club subequal in length to segments 2-7. Mandibles with 2 apical teeth and a rounded lobe at base. Pronotum : Surface on disc with sparse, small punctures; punctures becoming mod- erate in size and density on sides and anterior fourth. Anterior margin with small, median tu- bercle. A small, longitudinally oblong fovea present behind tubercle, fovea about as wide as distance between frontoclypeal tubercles, surface within fovea rugose. Base lacking marginal bead. Elytra : Surface shagreened, with moderate to large, weakly ocellate punctures, punctures becoming larger lat- erally; 3 pairs of double rows of punctures poorly defined. Pygidium : Surface on center third nearly smooth, with small, sparse punctures; surface else- where densely rugopunctate. In lateral view, sur- face convex. Legs : Protibia tridentate, basal tooth slightly removed from others. Protarsus simple, not enlarged. Apex of metatibia subtruncate, with 10 stout, short spinules (some broken off, sockets visible). Apex of first tarsomere of metatarsus subtruncate. Venter : Prosternal process long, thick, columnar, setigerous at apex, apex longitu- dinally oval; setae long, reddish brown. Parameres : Fig. 11-12. The combination of nar- row form, a long tooth projecting laterally at the Figure 13. Tomarus cuniculus (Fabricius) , habitus. midpoint of the shaft, and a narrowly elongate apex is unique among Tomarus species. Allotype . Female. Length 21.7 mm ; width 11.7 mm . Color dark reddish brown. As holotype , except in the following respects: Head : Entire surface transversely rugose. Occiput at center lacking smooth, shiny area. Pronotum : Fovea reduced to a shallow, short, longitudinal depression. Pygidium : Surface on center third with sparse, large punctures. In lateral view, surface nearly flat. Legs : Protibia with trace of fourth, basal tooth. Left posterior leg missing. Variation. Males ( 1 paratype ). Length 22.5 mm ; width 12.4 mm . The male paratype does not differ significantly from the holotype . Females ( 3 paratypes ). Length 20.0- 22.4 mm ; width 10.3-12.2 mm . The female paratypes do not differ significantly from the allotype . Etymology. From the Greek adoketus , meaning surprising or unusual. Used here as a noun in apposition, hence, Tomarus , the surprising one.