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<document id="CD62E3F5B83850A9DAA3F06EE424D724" ID-DOI="10.11646/zootaxa.5322.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8203657" ID-ZooBank="DD295EE6-AB3F-4377-900E-CC76F1C6BC2D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690885128246" checkinUser="plazi" docAuthor="Klompen, Hans &amp; Gerdeman, Beverly S." docDate="2023" docId="C55587C9FFD4B82286C9CEABF1AAFDEE" docLanguage="en" docName="zootaxa.5322.1.1.pdf" docOrigin="Zootaxa 5322 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5322.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Narceoheterozercon Gerdeman &amp; Klompen 2003" docType="treatment" docVersion="1" lastPageNumber="29" masterDocId="396CFFB1FFCEB83E865EC861F07CFFD3" masterDocTitle="Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)" masterLastPageNumber="66" masterPageNumber="1" pageNumber="27" updateTime="1726239911995" updateUser="tatiana">
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<mods:title id="B0C9B023FB43A89F7AEC056AC7956A0D">Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)</mods:title>
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<mods:namePart id="66963CC22618B012D2AE8006FCD0321C">Klompen, Hans</mods:namePart>
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<mods:namePart id="9C45936D216AE932EFB02AC4E720F5C1">Gerdeman, Beverly S.</mods:namePart>
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<mods:date id="B08129196400030F6ADA2EA1B168C792">2023</mods:date>
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<treatment id="C55587C9FFD4B82286C9CEABF1AAFDEE" ID-DOI="http://doi.org/10.5281/zenodo.8209449" ID-Zenodo-Dep="8209449" LSID="urn:lsid:plazi:treatment:C55587C9FFD4B82286C9CEABF1AAFDEE" httpUri="http://treatment.plazi.org/id/C55587C9FFD4B82286C9CEABF1AAFDEE" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<subSubSection id="05E66554FFD4B82486C9CEABF28AF937" box="[151,758,1738,1765]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="4D4336DFFFD4B82486C9CEABF28AF937" blockId="26.[151,758,1738,1765]" box="[151,758,1738,1765]" pageId="26" pageNumber="27">
<heading id="160B81B3FFD4B82486C9CEABF28AF937" bold="true" box="[151,758,1738,1765]" fontSize="11" level="2" pageId="26" pageNumber="27" reason="4">
<taxonomicName id="8AFC4D5CFFD4B82486C9CEABF28AF937" authority="Gerdeman &amp; Klompen 2003" authorityName="Gerdeman &amp; Klompen" authorityYear="2003" box="[151,758,1738,1765]" class="Arachnida" family="Heterozerconidae" genus="Narceoheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFD4B82486C9CEABF28AF937" bold="true" box="[151,758,1738,1765]" pageId="26" pageNumber="27">
<emphasis id="7F88EACDFFD4B82486C9CEABF1F7F936" bold="true" box="[151,395,1738,1765]" italics="true" pageId="26" pageNumber="27">Narceoheterozercon</emphasis>
<bibRefCitation id="296D4B2EFFD4B82487CDCEABF28AF937" author="Gerdeman, B. S. &amp; Klompen, H." box="[403,758,1738,1765]" pageId="26" pageNumber="27" pagination="351 - 370" refId="ref38078" refString="Gerdeman, B. S. &amp; Klompen, H. (2003) A new North American heterozerconid, Narceoheterozercon ohioensis, n. gen., n. sp., with first description of immatures of Heterozerconidae (Acari: Mesostigmata). International Journal of Acarology, 29, 351 - 370. https: // doi. org / 10.1080 / 01647950308684352" type="journal article" year="2003">Gerdeman &amp; Klompen 2003</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="05E66554FFD4B82486C9CF6EF2B2F8F4" box="[151,718,1807,1831]" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="4D4336DFFFD4B82486C9CF6EF2B2F8F4" blockId="26.[151,1436,1807,1862]" box="[151,718,1807,1831]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="6DEC11F1FFD4B82486C9CF6EF2B2F8F4" box="[151,718,1807,1831]" pageId="26" pageNumber="27">
<treatmentCitation id="CC5D10CEFFD4B82486C9CF6EF2B2F8F4" author="Gerdeman, B. S. &amp; Klompen, H." box="[151,718,1807,1831]" page="352" pageId="26" pageNumber="27" year="2003">
<taxonomicName id="8AFC4D5CFFD4B82486C9CF6EF2B2F8F4" authority="Gerdeman &amp; Klompen 2003: 352" authorityName="Gerdeman &amp; Klompen" authorityPageNumber="352" authorityYear="2003" box="[151,718,1807,1831]" class="Arachnida" family="Heterozerconidae" genus="Narceoheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFD4B82486C9CF6EF11FF8F4" box="[151,355,1807,1831]" italics="true" pageId="26" pageNumber="27">Narceoheterozercon</emphasis>
<bibRefCitation id="296D4B2EFFD4B8248733CF6EF2B2F8F4" author="Gerdeman, B. S. &amp; Klompen, H." box="[365,718,1807,1831]" pageId="26" pageNumber="27" pagination="351 - 370" refId="ref38078" refString="Gerdeman, B. S. &amp; Klompen, H. (2003) A new North American heterozerconid, Narceoheterozercon ohioensis, n. gen., n. sp., with first description of immatures of Heterozerconidae (Acari: Mesostigmata). International Journal of Acarology, 29, 351 - 370. https: // doi. org / 10.1080 / 01647950308684352" type="journal article" year="2003">Gerdeman &amp; Klompen 2003: 352</bibRefCitation>
</taxonomicName>
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="05E66554FFD4B8248486CF71F27CF895" pageId="26" pageNumber="27" type="type_taxon">
<paragraph id="4D4336DFFFD4B8248486CF71F27CF895" blockId="26.[151,1436,1807,1862]" pageId="26" pageNumber="27">
[
<typeStatus id="9247887DFFD4B8248483CF71F377F8FB" box="[733,779,1808,1832]" pageId="26" pageNumber="27">type</typeStatus>
species:
<taxonomicName id="8AFC4D5CFFD4B8248531CF6EF080F895" authority="Gerdeman &amp; Klompen 2003" authorityName="Gerdeman &amp; Klompen" authorityYear="2003" class="Arachnida" family="Heterozerconidae" genus="Narceoheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ohioensis">
<emphasis id="7F88EACDFFD4B8248531CF6EF4D8F8F4" box="[879,1188,1807,1831]" italics="true" pageId="26" pageNumber="27">Narceoheterozercon ohioensis</emphasis>
<bibRefCitation id="296D4B2EFFD4B82482F0CF6EF080F895" author="Gerdeman, B. S. &amp; Klompen, H." pageId="26" pageNumber="27" pagination="351 - 370" refId="ref38078" refString="Gerdeman, B. S. &amp; Klompen, H. (2003) A new North American heterozerconid, Narceoheterozercon ohioensis, n. gen., n. sp., with first description of immatures of Heterozerconidae (Acari: Mesostigmata). International Journal of Acarology, 29, 351 - 370. https: // doi. org / 10.1080 / 01647950308684352" type="journal article" year="2003">Gerdeman &amp; Klompen 2003</bibRefCitation>
</taxonomicName>
, by original designation].
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFD4B82586C9CF15F163FF2A" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="4D4336DFFFD4B82586C9CF15F163FF2A" blockId="26.[151,1436,1908,2043]" lastBlockId="27.[151,1436,150,249]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Diagnosis. Spermatodactyl long, recurved, and largely smooth (7, except in
<taxonomicName id="8AFC4D5CFFD4B82485A0CF15F4A3F85D" authorityName="Gerdeman &amp; Klompen" authorityYear="2003" box="[1022,1247,1908,1934]" class="Arachnida" family="Heterozerconidae" genus="Narceoheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFD4B82485A0CF15F4A3F85D" box="[1022,1247,1908,1934]" italics="true" pageId="26" pageNumber="27">Narceoheterozercon</emphasis>
</taxonomicName>
ex Alabama-C); anterior dorsal setae
<emphasis id="7F88EACDFFD4B82487DDCFF9F1E4F861" box="[387,408,1944,1970]" italics="true" pageId="26" pageNumber="27">j1</emphasis>
minute (&lt;5 µm) (28.2); sternal lyrifissures
<emphasis id="7F88EACDFFD4B82485D6CFF9F3D6F861" box="[904,938,1944,1970]" italics="true" pageId="26" pageNumber="27">
iv
<quantity id="8A049B3AFFD4B82485C0CFF9F3D6F861" box="[926,938,1944,1970]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="26" pageNumber="27" unit="in" value="1.0">1</quantity>
</emphasis>
in adults absent (44); sclerotized areas near sternal setae
<emphasis id="7F88EACDFFD4B8248775CFDDF137F805" box="[299,331,1980,2006]" italics="true" pageId="26" pageNumber="27">st2</emphasis>
and
<emphasis id="7F88EACDFFD4B82487DACFDDF1D9F805" box="[388,421,1980,2006]" italics="true" pageId="26" pageNumber="27">
st
<quantity id="8A049B3AFFD4B82487C9CFDDF1D9F805" box="[407,421,1980,2006]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="26" pageNumber="27" unit="in" value="3.0">3</quantity>
</emphasis>
in male completely fused (51.0); females with prominent, well sclerotized, solenostomes arching anteriorly from the antero-median corner of the ventral opisthosomal suckers (64, except in
<taxonomicName id="8AFC4D5CFFD4B8258341CF81F179FF63" authorityName="Gerdeman &amp; Klompen" authorityYear="2003" class="Arachnida" family="Heterozerconidae" genus="Narceoheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="27" lastPageNumber="28" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFD4B8258341CF81F179FF63" italics="true" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">Narceoheterozercon</emphasis>
</taxonomicName>
ex Alabama-C); setae
<emphasis id="7F88EACDFFD5B8258457C8F6F259FF62" box="[521,549,151,177]" italics="true" pageId="27" pageNumber="28">
S
<quantity id="8A049B3AFFD5B8258449C8F6F259FF62" box="[535,549,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="27" pageNumber="28" unit="in" value="5.0">5</quantity>
</emphasis>
in adults minute (75), all marginal opisthosomal setae setiform (77, reversal); femora I seta
<emphasis id="7F88EACDFFD5B8258771C8DAF128FF06" box="[303,340,187,213]" italics="true" pageId="27" pageNumber="28">pl2</emphasis>
absent (85), femora II seta
<emphasis id="7F88EACDFFD5B82584D8C8DAF2DCFF07" box="[646,672,187,212]" italics="true" pageId="27" pageNumber="28">pv</emphasis>
in males setiform (108, reversal); remaining setae on legs II of male all setiform.
</paragraph>
</subSubSection>
<caption id="19836657FFD5B82586C9CFD6F46DF820" ID-DOI="http://doi.org/10.5281/zenodo.8203663" ID-Zenodo-Dep="8203663" httpUri="https://zenodo.org/record/8203663/files/figure.png" pageId="27" pageNumber="28" startId="27.[151,264,1975,1999]" targetBox="[154,1436,273,1949]" targetPageId="27" targetType="figure">
<paragraph id="4D4336DFFFD5B82586C9CFD6F46DF820" blockId="27.[151,1436,1975,2035]" pageId="27" pageNumber="28">
<emphasis id="7F88EACDFFD5B82586C9CFD6F13CF81C" bold="true" box="[151,320,1975,1999]" pageId="27" pageNumber="28">FIGURES 8–11</emphasis>
.
<taxonomicName id="8AFC4D5CFFD5B8258713CFD6F27EF81C" authorityName="Berlese" authorityYear="1892" box="[333,514,1975,1999]" class="Arachnida" family="Heterozerconidae" kingdom="Animalia" order="Mesostigmata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="family">Heterozerconidae</taxonomicName>
, chelicera males, photograph (top) and composite drawing (bottom): 8–9,
<taxonomicName id="8AFC4D5CFFD5B82582A6CFD6F155F820" baseAuthorityName="Flechtmann &amp; Johnston" baseAuthorityYear="1990" class="Arachnida" family="Heterozerconidae" genus="Amheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="amphisbaenae">
<emphasis id="7F88EACDFFD5B82582A6CFD6F155F820" italics="true" pageId="27" pageNumber="28">Amheterozercon amphisbaenae</emphasis>
</taxonomicName>
(OSAL 052760); 10–11,
<taxonomicName id="8AFC4D5CFFD5B8258473CFBAF323F820" authorityName="Gerdeman &amp; Klompen" authorityYear="2003" box="[557,863,2011,2035]" class="Arachnida" family="Heterozerconidae" genus="Narceoheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="ohioensis">
<emphasis id="7F88EACDFFD5B8258473CFBAF323F820" box="[557,863,2011,2035]" italics="true" pageId="27" pageNumber="28">Narceoheterozercon ohioensis</emphasis>
</taxonomicName>
(OSAL 001221).
</paragraph>
</caption>
<subSubSection id="05E66554FFD2B8228699C8F6F1AAFDEE" pageId="28" pageNumber="29" type="discussion">
<paragraph id="4D4336DFFFD2B8228699C8F6F240FE5A" blockId="28.[151,1437,151,573]" pageId="28" pageNumber="29">
Remarks. The diagnosis of the genus by
<bibRefCitation id="296D4B2EFFD2B82284FDC8F6F383FF62" author="Gerdeman, B. S. &amp; Klompen, H." box="[675,1023,151,177]" pageId="28" pageNumber="29" pagination="351 - 370" refId="ref38078" refString="Gerdeman, B. S. &amp; Klompen, H. (2003) A new North American heterozerconid, Narceoheterozercon ohioensis, n. gen., n. sp., with first description of immatures of Heterozerconidae (Acari: Mesostigmata). International Journal of Acarology, 29, 351 - 370. https: // doi. org / 10.1080 / 01647950308684352" type="journal article" year="2003">Gerdeman &amp; Klompen (2003)</bibRefCitation>
included four characters: 1) female with prominent, well sclerotized, solenostomes arching anteriorly from the antero- median corner of the ventral opisthosomal suckers (64.2); 2) male with long, recurved, and largely smooth spermatodactyl (7.2); 3) male lacking prominent spines on legs II (107–110); 4) adults lacking spine-like marginal opisthosomal setae (77). Of these, characters 1and 2, while unique, are not shared by some taxa included in the genus in this study (e.g., population Alabama-C). As a result, the diagnosis (and support) for this genus is not strong. Data on additional species may help clarify the concept of this genus.
</paragraph>
<paragraph id="4D4336DFFFD2B8228699C9F2F1AAFDEE" blockId="28.[151,1437,151,573]" pageId="28" pageNumber="29">
Most
<taxonomicName id="8AFC4D5CFFD2B8228754C9F2F23AFE7E" box="[266,582,403,429]" pageId="28" pageNumber="29">
<emphasis id="7F88EACDFFD2B8228754C9F2F197FE7E" box="[266,491,403,429]" italics="true" pageId="28" pageNumber="29">Narceoheterozercon</emphasis>
species
</taxonomicName>
have secondary genital openings (solenostomes) in the female (
<bibRefCitation id="296D4B2EFFD2B8228378C9F2F0A8FE02" author="Gerdeman, B. S." pageId="28" pageNumber="29" refId="ref37955" refString="Gerdeman, B. S. (2002) The biology of the Heterozerconidae. Ph. D. thesis, Department of Entomology, Ohio State University, Columbus, Ohio, 119 pp." type="book" year="2002">Gerdeman 2002</bibRefCitation>
). These are located at the antero-internal corners of the opisthogastral suckers and continue internally in the form of sclerotized tubes (
<figureCitation id="D5C72A5AFFD2B822879FC9BAF269FE26" box="[449,533,475,501]" captionStart="FIGURES 61–62" captionStartId="51.[151,264,1011,1035]" captionTargetBox="[247,1340,187,985]" captionTargetId="figure-21@51.[242,1345,181,988]" captionTargetPageId="51" captionText="FIGURES 61–62. Heterozerconidae, Narceoheterozercon ohioensis, idiosoma venter: 61, female; 62, male. Male redrawn from Gerdeman &amp; Klompen (2003)." figureDoi="http://doi.org/10.5281/zenodo.8203711" httpUri="https://zenodo.org/record/8203711/files/figure.png" pageId="28" pageNumber="29">Fig. 61</figureCitation>
, arrow). Evidence that these tubes serve as secondary genital openings includes the observation that these structures are limited to the females, and the observation of a broken-off spermatodactyl in one of the tubes (
<figureCitation id="D5C72A5AFFD2B822872BCA42F1B6FDEE" box="[373,458,546,573]" captionStart="FIGURES 48–52" captionStartId="48.[151,264,1906,1930]" captionTargetBox="[235,1353,181,1883]" captionTargetId="figure-25@48.[235,1353,181,1883]" captionTargetPageId="48" captionText="FIGURES 48–52. Heterozerconidae, details idiosoma: 48, Allozercon leytensis n. sp., female, hypertrichy of setae in j1 position (OSAL 053251); 49, same for male (OSAL 106780); 50, Allozercon leytensis n. sp., male, anterolateral spines (OSAL 106780); 50a, detail antero-lateral spines; 51, Heterozercon ex Cuba, female, venter (OSAL 125446); 52, Narceoheterozercon sp., broken off spermatodactyl (red arrow) in female secondary genital opening / sclerotized tube (OSAL 159712)." figureDoi="http://doi.org/10.5281/zenodo.8203697" httpUri="https://zenodo.org/record/8203697/files/figure.png" pageId="28" pageNumber="29">Fig. 52</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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