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<mods:title id="B889E3C3EB839DF1B3A6B7A4E6B4E681">Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)</mods:title>
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<taxonomicName id="8AFC4D5CFFD2B82286C9CD74F192FAFC" authority="Fain 1989" authorityName="Fain" authorityYear="1989" box="[151,494,1301,1328]" class="Arachnida" family="Heterozerconidae" genus="Afroheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis id="7F88EACDFFD2B82286C9CD74F117FAE3" bold="true" box="[151,363,1301,1328]" italics="true" pageId="28" pageNumber="29">Afroheterozercon</emphasis>
<bibRefCitation id="296D4B2EFFD2B822872CCD74F192FAFC" author="Fain, A." box="[370,494,1301,1327]" pageId="28" pageNumber="29" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989</bibRefCitation>
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<taxonomicName id="8AFC4D5CFFD2B82286C9CD3BF199FAA1" authority="Fain 1989: 147" authorityName="Fain" authorityPageNumber="147" authorityYear="1989" box="[151,485,1370,1394]" class="Arachnida" family="Heterozerconidae" genus="Afroheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFD2B82286C9CD3BF139FAA1" box="[151,325,1370,1394]" italics="true" pageId="28" pageNumber="29">Afroheterozercon</emphasis>
<bibRefCitation id="296D4B2EFFD2B8228712CD3BF199FAA1" author="Fain, A." box="[332,485,1370,1394]" pageId="28" pageNumber="29" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989: 147</bibRefCitation>
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[
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species.
<taxonomicName id="8AFC4D5CFFD2B8228422CD3AF387FAA1" authority="Fain 1988" authorityName="Fain" authorityYear="1988" box="[636,1019,1370,1394]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="spirostreptus">
<emphasis id="7F88EACDFFD2B8228422CD3AF3F0FAA2" box="[636,908,1370,1394]" italics="true" pageId="28" pageNumber="29">Heterozercon spirostreptus</emphasis>
<bibRefCitation id="296D4B2EFFD2B82285CDCD3BF387FAA1" author="Fain, A." box="[915,1019,1370,1394]" pageId="28" pageNumber="29" pagination="237 - 242" refId="ref37668" refString="Fain, A. (1988) Notes on mites associated with Myriapoda. III. Two new species of the genus Heterozercon Berlese, 1888 (Acari, Mesostigmata) from Afrotropical Myriapods. Bulletin et Annales de la Societe royale belge d'Entomologie, 124, 237 - 242." type="journal article" year="1988">Fain 1988</bibRefCitation>
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, by original designation].
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<taxonomicName id="8AFC4D5CFFD2B82286C9CD19F139FA43" authorityName="Fain" authorityYear="1989" box="[151,325,1400,1424]" class="Arachnida" family="Heterozerconidae" genus="Afroheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFD2B82286C9CD19F139FA43" box="[151,325,1400,1424]" italics="true" pageId="28" pageNumber="29">Afroheterozercon</emphasis>
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.—
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<bibRefCitation id="296D4B2EFFD2B822873ACD18F26FFA43" author="Evans, G. O." box="[356,531,1400,1425]" pageId="28" pageNumber="29" refId="ref37637" refString="Evans, G. O. (1992) Principles of acarology. CAB International, Wallingford, 563 pp. https: // doi. org / 10.1079 / 9780851988221.0000" type="book" year="1992">Evans 1992: 426</bibRefCitation>
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;
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Klompen
<emphasis id="7F88EACDFFD2B82284DCCD18F2D2FA43" box="[642,686,1400,1424]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
. 2013: 302
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.
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</paragraph>
</subSubSection>
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Diagnosis. Spermatodactyl of male of compressed
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(7, except in
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<emphasis id="7F88EACDFFD2B82285CACDDEF3D1FA0A" box="[916,941,1471,1497]" italics="true" pageId="28" pageNumber="29">Af</emphasis>
.
<emphasis id="7F88EACDFFD2B82285E4CDDEF436FA0B" box="[954,1098,1471,1496]" italics="true" pageId="28" pageNumber="29">spirostreptus</emphasis>
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); cornicula with a distal notch (24;
<figureCitation id="D5C72A5AFFD2B8228690CD82F15EFA2E" box="[206,290,1507,1533]" captionStart="FIGURES 3134" captionStartId="43.[151,264,1909,1933]" captionTargetBox="[204,1384,184,1882]" captionTargetId="figure-21@43.[201,1387,181,1885]" captionTargetPageId="43" captionText="FIGURES 3134. Heterozerconidae, gnathosoma, adults: 31, Narceoheterozercon ohioensis, female (OSAL 001054); 32, Narceoheterozercon ohioensis, male (redrawn from Gerdeman &amp; Klompen (2003)); 33, Afroheterozercon mahsbergi, female (OSAL 003037); 34, Afroheterozercon mahsbergi, male (OSAL 003042; red arrow: notch in corniculus). Inset: gnathotectum (female and male) and labrum + salivary stylets. Abbreviations: LL, lateral lips." figureDoi="http://doi.org/10.5281/zenodo.8203681" httpUri="https://zenodo.org/record/8203681/files/figure.png" pageId="28" pageNumber="29">Fig. 34</figureCitation>
, arrow); sclerotized areas near insertion sternal setae
<emphasis id="7F88EACDFFD2B82285D2CD82F3D0FA2E" box="[908,940,1507,1533]" italics="true" pageId="28" pageNumber="29">
st
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in female fused to endopodal shields (38); sclerotized areas near sternal setae
<emphasis id="7F88EACDFFD2B8228440CE66F242F9F2" box="[542,574,1543,1569]" italics="true" pageId="28" pageNumber="29">st2</emphasis>
and
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st
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in male completely fused (51.0); endopodal and sternitiventral shield in male fused (58, shared with
<taxonomicName id="8AFC4D5CFFD2B82287B2CE4AF2F2F997" box="[492,654,1579,1604]" class="Arachnida" family="Heterozerconidae" genus="Maracazercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFD2B82287B2CE4AF2F2F997" box="[492,654,1579,1604]" italics="true" pageId="28" pageNumber="29">Maracazercon</emphasis>
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); metapodal and sternitiventral shields in male fused (59); sternitiventral and anal shields in male fused, but line of fusion still visible (66.1, shared with several species in other genera); marginal opisthosomal setae anchor-like (78.2); lyrifissures
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iv
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in male absent (79).
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFD2B8228699CEF6F35EF87E" pageId="28" pageNumber="29" type="discussion">
<paragraph id="4D4336DFFFD2B8228699CEF6F5EEF92A" blockId="28.[151,1437,1471,1965]" pageId="28" pageNumber="29">
Remarks. Of the 12 characters that can be distinguished in
<bibRefCitation id="296D4B2EFFD2B8228505CEF6F38DF962" author="Fain, A." box="[859,1009,1687,1713]" pageId="28" pageNumber="29" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fains (1989)</bibRefCitation>
diagnosis of the genus, two (marginal opisthosomal region with numerous anchor-like spines (78), metapodal and sternitiventral shields fused (59)) are unique derived characters, the remainder are either primitive at this level, or are subject to multiple optimizations.
</paragraph>
<paragraph id="4D4336DFFFD2B8228699CF62F35EF87E" blockId="28.[151,1437,1471,1965]" pageId="28" pageNumber="29">
Distinct structures directly posterior to coxae IV of the females (
<figureCitation id="D5C72A5AFFD2B82285FACF62F38BF8CE" box="[932,1015,1795,1821]" captionStart="FIGURES 6768" captionStartId="52.[151,264,1957,1981]" captionTargetBox="[264,1325,1167,1929]" captionTargetId="figure-44@52.[258,1328,1161,1934]" captionTargetPageId="52" captionText="FIGURES 6768. Heterozerconidae, Heterozercon calakmulensis n. sp., idiosoma venter: 67, female (CNAC 012444); 68, male (CNAC 012443)." figureDoi="http://doi.org/10.5281/zenodo.8203717" httpUri="https://zenodo.org/record/8203717/files/figure.png" pageId="28" pageNumber="29">Fig. 67</figureCitation>
, arrow) are assumed to be secondary genital openings (solenostomes). They are present in females of most species of
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<emphasis id="7F88EACDFFD2B822826ECF46F48DF892" box="[1072,1265,1831,1857]" italics="true" pageId="28" pageNumber="29">Afroheterozercon</emphasis>
</taxonomicName>
examined (see
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Klompen
<emphasis id="7F88EACDFFD2B8228759CF2AF145F8B6" box="[263,313,1867,1893]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
. 2013
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) and absent in all males. The exception is
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<emphasis id="7F88EACDFFD2B822853ACF2AF301F8B6" box="[868,893,1867,1893]" italics="true" pageId="28" pageNumber="29">Af</emphasis>
.
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</taxonomicName>
where females lack the structures posterior to coxae IV. This is also the only species where the male has a spermatodactyl that resembles the “straight”, rather than the “compressed”,
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(
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<taxonomicName id="8AFC4D5CFFD4B82486C9C945F208FEE8" authority="Finnegan 1931: 1349" authorityName="Finnegan" authorityPageNumber="1349" authorityYear="1931" box="[151,628,291,315]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="oudemansi">
<emphasis id="7F88EACDFFD4B82486C9C945F1E8FEE8" box="[151,404,291,315]" italics="true" pageId="26" pageNumber="27">Heterozercon oudemansi</emphasis>
<bibRefCitation id="296D4B2EFFD4B82487C4C942F208FEE8" author="Finnegan, S." box="[410,628,291,315]" pageId="26" pageNumber="27" pagination="1349 - 1357" refId="ref37764" refString="Finnegan, S. (1931) On a new species of mite of the family Heterozerconidae parasitic on a snake. Proceedings of the Zoological Society of London, 4, 1349 - 1357. https: // doi. org / 10.1111 / j. 1096 - 3642.1931. tb 01066. x" type="journal article" year="1931">Finnegan 1931: 1349</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4D4336DFFFD4B82486C9C922F213FEAA" blockId="26.[151,1437,291,407]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="6DEC11F1FFD4B82486C9C922F255FE89" box="[151,553,322,346]" pageId="26" pageNumber="27">
<treatmentCitation id="CC5D10CEFFD4B82486C9C922F255FE89" author="Lizaso, N. M." box="[151,553,322,346]" page="140" pageId="26" pageNumber="27" year="1979">
<taxonomicName id="8AFC4D5CFFD4B82486C9C922F255FE89" authority="Lizaso 1979: 140" authorityName="Lizaso" authorityPageNumber="140" authorityYear="1979" box="[151,553,322,346]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="7F88EACDFFD4B82486C9C922F10EFE89" box="[151,370,322,346]" italics="true" pageId="26" pageNumber="27">Heterozercon elegans</emphasis>
<bibRefCitation id="296D4B2EFFD4B8248727C923F255FE89" author="Lizaso, N. M." box="[377,553,322,346]" pageId="26" pageNumber="27" pagination="139 - 144" refId="ref39295" refString="Lizaso, N. M. (1979) Um novo acaro da familia Heterozerconidae coletado sobre serpentes Brasileiras. Descric &quot; o de Heterozercon elegans sp. n. (Acarina: Mesostigmata). Memorias do Instituto Butantan, Sao Paulo, 42 - 43, 139 - 144." type="journal article" year="1979">Lizaso 1979: 140</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
[junior synonym of
<treatmentCitation id="CC5D10CEFFD4B82484A7C923F459FE89" author="Fain, A." box="[761,1061,322,346]" page="155" pageId="26" pageNumber="27" year="1989">
<taxonomicName id="8AFC4D5CFFD4B82484A7C923F459FE89" authority="Fain 1989: 155" authorityName="Fain" authorityPageNumber="155" authorityYear="1989" box="[761,1061,322,346]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="oudemansi">
<emphasis id="7F88EACDFFD4B82484A7C923F374FE8A" box="[761,776,322,345]" italics="true" pageId="26" pageNumber="27">A</emphasis>
.
<emphasis id="7F88EACDFFD4B824854BC923F3FFFE89" box="[789,899,322,346]" italics="true" pageId="26" pageNumber="27">oudemansi</emphasis>
<bibRefCitation id="296D4B2EFFD4B82485D7C923F459FE89" author="Fain, A." box="[905,1061,322,346]" pageId="26" pageNumber="27" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989: 155</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<treatmentCitation id="CC5D10CEFFD4B8248271C923F5EDFE89" author="Flechtmann, C. H. W. &amp; Johnston, D. E." box="[1071,1425,322,346]" page="143" pageId="26" pageNumber="27" year="1990">
<bibRefCitation id="296D4B2EFFD4B8248271C923F5EDFE89" author="Flechtmann, C. H. W. &amp; Johnston, D. E." box="[1071,1425,322,346]" pageId="26" pageNumber="27" pagination="143 - 148" refId="ref37819" refString="Flechtmann, C. H. W. &amp; Johnston, D. E. (1990) Zeterohercon, a new genus of Heterozerconidae (Acari: Mesostigmata) and the description of Zeterohercon amphisbaenae n. sp. from Brasil. International Journal of Acarology, 16, 143 - 148. https: // doi. org / 10.1080 / 01647959008683526" type="journal article" year="1990">Flechtmann &amp; Johnston 1990: 143</bibRefCitation>
</treatmentCitation>
].
<treatmentCitationGroup id="6DEC11F1FFD4B82486C9C901F213FEAA" box="[151,623,352,377]" pageId="26" pageNumber="27">
<taxonomicName id="8AFC4D5CFFD4B82486C9C901F1CCFEAB" baseAuthorityName="Finnegan" baseAuthorityYear="1931" box="[151,432,352,376]" class="Arachnida" family="Heterozerconidae" genus="Amheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="oudemansi">
<emphasis id="7F88EACDFFD4B82486C9C901F1CCFEAB" box="[151,432,352,376]" italics="true" pageId="26" pageNumber="27">Amheterozercon oudemansi</emphasis>
</taxonomicName>
.—
<treatmentCitation id="CC5D10CEFFD4B8248793C901F210FEAA" author="Fain, A." box="[461,620,352,377]" page="148" pageId="26" pageNumber="27" year="1989">
<bibRefCitation id="296D4B2EFFD4B8248793C901F210FEAA" author="Fain, A." box="[461,620,352,377]" pageId="26" pageNumber="27" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989: 148</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4D4336DFFFD4B82486C9C91EF36BFE44" blockId="26.[151,1437,291,407]" box="[151,791,383,407]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="6DEC11F1FFD4B82486C9C91EF36BFE44" box="[151,791,383,407]" pageId="26" pageNumber="27">
<taxonomicName id="8AFC4D5CFFD4B82486C9C91EF1EEFE44" authority="(Finnegan 1931)" box="[151,402,383,407]" class="Arachnida" family="Heterozerconidae" genus="Zeterohercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="oudemansi">
<emphasis id="7F88EACDFFD4B82486C9C91EF1EEFE44" box="[151,402,383,407]" italics="true" pageId="26" pageNumber="27">Zeterohercon oudemansi</emphasis>
</taxonomicName>
.—
<treatmentCitation id="CC5D10CEFFD4B82487F1C91EF36FFE44" author="Flechtmann, C. H. W. &amp; Johnston, D. E." box="[431,787,383,407]" page="143" pageId="26" pageNumber="27" year="1990">
<bibRefCitation id="296D4B2EFFD4B82487F1C91EF36FFE44" author="Flechtmann, C. H. W. &amp; Johnston, D. E." box="[431,787,383,407]" pageId="26" pageNumber="27" pagination="143 - 148" refId="ref37819" refString="Flechtmann, C. H. W. &amp; Johnston, D. E. (1990) Zeterohercon, a new genus of Heterozerconidae (Acari: Mesostigmata) and the description of Zeterohercon amphisbaenae n. sp. from Brasil. International Journal of Acarology, 16, 143 - 148. https: // doi. org / 10.1080 / 01647959008683526" type="journal article" year="1990">Flechtmann &amp; Johnston 1990: 143</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFD4B82486C9C9A7F1DDFD40" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="4D4336DFFFD4B82486C9C9A7F55EFE0C" blockId="26.[151,1436,453,659]" box="[151,1314,453,480]" pageId="26" pageNumber="27">
Type depositories.
<typeStatus id="9247887DFFD4B8248735C9A4F1AFFE0C" box="[363,467,453,479]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
<taxonomicName id="8AFC4D5CFFD4B8248784C9A7F209FE0C" box="[474,629,453,480]" class="Arachnida" family="Heterozerconidae" genus="Zeterohercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="oudemansi">
<emphasis id="7F88EACDFFD4B8248784C9A7F192FE0C" box="[474,494,454,479]" italics="true" pageId="26" pageNumber="27">H</emphasis>
.
<emphasis id="7F88EACDFFD4B82487A2C9A4F209FE0C" box="[508,629,453,479]" italics="true" pageId="26" pageNumber="27">oudemansi</emphasis>
</taxonomicName>
deposited at
<collectionCode id="2BEDAE1AFFD4B8248554C9A7F31EFE33" box="[778,866,454,480]" pageId="26" pageNumber="27">BMNH</collectionCode>
;
<typeStatus id="9247887DFFD4B8248533C9A4F3B3FE0C" box="[877,975,453,479]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
<taxonomicName id="8AFC4D5CFFD4B8248588C9A7F432FE0C" authorityName="Lizaso" authorityYear="1979" box="[982,1102,453,480]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="7F88EACDFFD4B8248588C9A7F396FE0C" box="[982,1002,454,479]" italics="true" pageId="26" pageNumber="27">H</emphasis>
.
<emphasis id="7F88EACDFFD4B82485A6C9A4F432FE0C" box="[1016,1102,453,479]" italics="true" pageId="26" pageNumber="27">elegans</emphasis>
</taxonomicName>
deposited at
<collectionCode id="2BEDAE1AFFD4B82482BCC9A4F562FE0C" box="[1250,1310,453,479]" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="26" pageNumber="27">IBSP</collectionCode>
.
</paragraph>
<paragraph id="4D4336DFFFD4B8248699C988F190FDD7" blockId="26.[151,1436,453,659]" box="[199,492,489,516]" pageId="26" pageNumber="27">Material examined. None.</paragraph>
<paragraph id="4D4336DFFFD4B8248699CA6CF1DDFD40" blockId="26.[151,1436,453,659]" pageId="26" pageNumber="27">
Remarks. Based on females and males on
<taxonomicName id="8AFC4D5CFFD4B82484EBCA6FF400FDF4" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[693,1148,525,551]" class="Squamata" family="Boidae" genus="Epicrates" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="cenchria">
<emphasis id="7F88EACDFFD4B82484EBCA6FF3F1FDF4" box="[693,909,525,551]" italics="true" pageId="26" pageNumber="27">Epicrates cenchria</emphasis>
(Linnaeus) (Boidae)
</taxonomicName>
from upper Amazon (
<taxonomicName id="8AFC4D5CFFD4B82483DFCA6FF16CFD98" class="Arachnida" family="Heterozerconidae" genus="Zeterohercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="oudemansi">
<emphasis id="7F88EACDFFD4B82483DFCA6FF5E9FDF4" box="[1409,1429,526,551]" italics="true" pageId="26" pageNumber="27">H</emphasis>
.
<emphasis id="7F88EACDFFD4B82486C9CA50F16CFD98" box="[151,272,561,587]" italics="true" pageId="26" pageNumber="27">oudemansi</emphasis>
</taxonomicName>
) (
<bibRefCitation id="296D4B2EFFD4B8248777CA50F1AAFD98" author="Finnegan, S." box="[297,470,561,587]" pageId="26" pageNumber="27" pagination="1349 - 1357" refId="ref37764" refString="Finnegan, S. (1931) On a new species of mite of the family Heterozerconidae parasitic on a snake. Proceedings of the Zoological Society of London, 4, 1349 - 1357. https: // doi. org / 10.1111 / j. 1096 - 3642.1931. tb 01066. x" type="journal article" year="1931">Finnegan 1931</bibRefCitation>
); on
<taxonomicName id="8AFC4D5CFFD4B8248450CA50F2E5FD98" box="[526,665,561,587]" class="Reptilia" family="Colubridae" genus="Waglerophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="7F88EACDFFD4B8248450CA50F2E5FD98" box="[526,665,561,587]" italics="true" pageId="26" pageNumber="27">Waglerophis</emphasis>
</taxonomicName>
(now
<taxonomicName id="8AFC4D5CFFD4B82484BCCA50F33AFD98" box="[738,838,561,587]" class="Squamata" family="Colubridae" genus="Xenodon" kingdom="Animalia" order="Tetraodontiformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="7F88EACDFFD4B82484BCCA50F33AFD98" box="[738,838,561,587]" italics="true" pageId="26" pageNumber="27">Xenodon</emphasis>
</taxonomicName>
)
<emphasis id="7F88EACDFFD4B8248508CA53F3C0FD98" box="[854,956,562,587]" italics="true" pageId="26" pageNumber="27">merremii</emphasis>
(Wagler),
<taxonomicName id="8AFC4D5CFFD4B824826ACA50F4B2FD98" authorityName="Amaral" authorityYear="1935" box="[1076,1230,561,587]" class="Squamata" family="Colubridae" genus="Mastigodryas" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="7F88EACDFFD4B824826ACA50F4B2FD98" box="[1076,1230,561,587]" italics="true" pageId="26" pageNumber="27">Mastigodryas</emphasis>
</taxonomicName>
(now
<taxonomicName id="8AFC4D5CFFD4B8248348CA50F5EEFD98" authorityName="Montingelli, Grazziotin, Battilana, Murphy, Zhang &amp; Zaher" authorityYear="2019" box="[1302,1426,561,587]" class="Squamata" family="Colubridae" genus="Palusophis" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="7F88EACDFFD4B8248348CA50F5EEFD98" box="[1302,1426,561,587]" italics="true" pageId="26" pageNumber="27">Palusophis</emphasis>
</taxonomicName>
)
<emphasis id="7F88EACDFFD4B82486C9CA34F17BFDBC" box="[151,263,597,623]" italics="true" pageId="26" pageNumber="27">bifossatus</emphasis>
(Raddi), and
<taxonomicName id="8AFC4D5CFFD4B82487F5CA34F323FDBC" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[427,863,597,623]" class="Squamata" family="Colubridae" genus="Erythrolamprus" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="aesculapii">
<emphasis id="7F88EACDFFD4B82487F5CA34F2A6FDBC" box="[427,730,597,623]" italics="true" pageId="26" pageNumber="27">Erythrolamprus aesculapii</emphasis>
(Linnaeus)
</taxonomicName>
(all
<taxonomicName id="8AFC4D5CFFD4B82485C6CA34F465FDBC" authorityName="Oppell" authorityYear="1811" box="[920,1049,597,623]" class="Squamata" family="Colubridae" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Chordata" rank="family">Colubridae</taxonomicName>
), from various sites in
<collectingCountry id="35EB764FFFD4B8248377CA34F511FDBC" box="[1321,1389,597,623]" name="Brazil" pageId="26" pageNumber="27">Brazil</collectingCountry>
(
<taxonomicName id="8AFC4D5CFFD4B82483DFCA37F091FD40" authorityName="Lizaso" authorityYear="1979" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="7F88EACDFFD4B82483DFCA37F5E9FDBC" box="[1409,1429,598,623]" italics="true" pageId="26" pageNumber="27">H</emphasis>
.
<emphasis id="7F88EACDFFD4B82486C9CA18F091FD40" box="[151,237,633,659]" italics="true" pageId="26" pageNumber="27">elegans</emphasis>
</taxonomicName>
) (
<bibRefCitation id="296D4B2EFFD4B8248758CA18F1E8FD40" author="Lizaso, N. M." box="[262,404,633,659]" pageId="26" pageNumber="27" pagination="139 - 144" refId="ref39295" refString="Lizaso, N. M. (1979) Um novo acaro da familia Heterozerconidae coletado sobre serpentes Brasileiras. Descric &quot; o de Heterozercon elegans sp. n. (Acarina: Mesostigmata). Memorias do Instituto Butantan, Sao Paulo, 42 - 43, 139 - 144." type="journal article" year="1979">Lizaso 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,54 +1,54 @@
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@ -150,7 +150,7 @@ on tarsi II spine-like (111, shared with
); all remaining setae on legs II of male setiform.
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Remarks. Of the characters listed in
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@ -201,9 +201,6 @@ Paramegistidae (
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<subSubSection id="05E66554FFD8B8288699CEC2F1C8F89E" pageId="22" pageNumber="23" type="discussion">
<paragraph id="4D4336DFFFD8B8288699CEC2F1F4F8D6" blockId="22.[151,1436,1411,1869]" pageId="22" pageNumber="23">
Remarks. The main difference between this genus and
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@ -132,25 +132,6 @@ Remarks. The main difference between this genus and
is the presence of well-developed ventral suckers. In many other characteristics, e.g., the shape of the female chelicera and elongate dorsal setae, it closely resembles that genus.
</paragraph>
<paragraph id="4D4336DFFFD8B8288699CF6EF1C8F89E" blockId="22.[151,1436,1411,1869]" pageId="22" pageNumber="23">An examination of multiple females using light microscopy did not reveal any indication of secondary genital openings (solenostomes).</paragraph>
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<paragraph id="4D4336DFFFD9B82986C9CFF6F391F800" blockId="23.[151,1436,1943,2004]" pageId="23" pageNumber="24">
<emphasis id="7F88EACDFFD9B82986C9CFF6F144F87C" bold="true" box="[151,312,1943,1967]" pageId="23" pageNumber="24">FIGURES 47</emphasis>
.
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, chelicera males, photograph (top) and composite drawing (bottom): 45,
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<emphasis id="7F88EACDFFD9B8298348CFF9F17AF800" italics="true" pageId="23" pageNumber="24">Amyzozercon chocoensis</emphasis>
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(OSAL 106788); 67,
<taxonomicName id="8AFC4D5CFFD9B82987AFCFDDF289F800" authorityName="Klompen &amp; Gerdeman" authorityYear="2023" box="[497,757,1979,2003]" class="Arachnida" family="Heterozerconidae" genus="Ecuazercon" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="cushuimensis">
<emphasis id="7F88EACDFFD9B82987AFCFDDF289F800" box="[497,757,1979,2003]" italics="true" pageId="23" pageNumber="24">Ecuazercon cushuimensis</emphasis>
</taxonomicName>
(FMNH-INS 4449621).
</paragraph>
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<emphasis id="7F88EACDFFD6B82686C9C8F7F113FF63" bold="true" box="[151,367,150,176]" pageId="24" pageNumber="25">Included species </emphasis>
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@ -124,7 +124,7 @@ Etymology. The generic name combines “a”, Greek for not, without, with “my
).
</paragraph>
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Remarks. “Undescribed
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@ -152,9 +152,6 @@ is easily recognizable by the absence of opisthogastral suckers (62), the presen
(62).
</paragraph>
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<emphasis id="7F88EACDFFDBB82B86C9C8F6F113FF62" bold="true" box="[151,367,150,177]" pageId="21" pageNumber="22">Included species </emphasis>
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).
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Diagnosis: Anterior dorsal setae
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<emphasis id="7F88EACDFFDCB82C86C9CF01F107F8A9" bold="true" box="[151,379,1887,1914]" italics="true" pageId="18" pageNumber="19">Discozercon derric</emphasis>
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Material examined. Two females and
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Remarks. On
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from
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(
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), on scolopendrid centipedes from two localities in
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,
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(
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@ -118,7 +118,7 @@ species:
).
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Diagnosis: Sclerotized areas around sternal setae
<emphasis id="7F88EACDFFDDB82D84E6CE1CF2A5F944" box="[696,729,1661,1687]" italics="true" pageId="19" pageNumber="20">
@ -152,9 +152,6 @@ examined. In fact, we are not aware of such a structure in any other
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.
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<emphasis id="7F88EACDFFDAB82A86C9C8F6F113FF62" bold="true" box="[151,367,150,177]" pageId="20" pageNumber="21">Included species </emphasis>
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Diagnosis: Movable digit in adults with thin, strap-like dorsal extension (2); gnathotectum without prominent points (15); median dorsal setae minute (31); genitiventral and anal shields not fused (66, reversed within some
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<heading id="160B81B3FFDFB82F86C9CCE6F24EFB73" bold="true" box="[151,562,1158,1185]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="4">
@ -120,7 +120,7 @@ in male fused to both sternitiventral and endopodal shields (48); opisthogastral
) setae (73); equally spaced long marginal setae present (76).
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFDFB82F8699CD88F113F8F0" pageId="17" pageNumber="18" type="discussion">
<subSubSection id="05E66554FFDFB82F8699CD88F3AEF964" pageId="17" pageNumber="18" type="discussion">
<paragraph id="4D4336DFFFDFB82F8699CD88F3AEF964" blockId="17.[151,1437,1297,1719]" pageId="17" pageNumber="18">
Remarks:
<bibRefCitation id="296D4B2EFFDFB82F8765CD88F233F9D0" author="Seeman, O. D. &amp; Baker, M. R." box="[315,591,1513,1539]" pageId="17" pageNumber="18" pagination="130 - 142" refId="ref39622" refString="Seeman, O. D. &amp; Baker, M. R. (2013) A new genus and species of Discozerconidae (Acari: Mesostigmata) from carabid beetles (Coleoptera: Carabidae) in New Zealand. Zootaxa, 3750 (2), 130 - 142. https: // doi. org / 10.11646 / zootaxa. 3750.2.2" type="journal article" year="2013">Seeman &amp; Baker (2013)</bibRefCitation>
@ -145,9 +145,6 @@ and
<taxonomicName id="8AFC4D5CFFDFB82F8298CE18F5ECF940" authorityName="Berlese" authorityYear="1892" box="[1222,1424,1657,1683]" class="Arachnida" family="Heterozerconidae" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Heterozerconidae</taxonomicName>
), lacking the medial posterior extension of the genital / genitiventral shield.
</paragraph>
<paragraph id="4D4336DFFFDFB82F86C9CF68F113F8F0" blockId="17.[151,367,1801,1827]" box="[151,367,1801,1827]" pageId="17" pageNumber="18">
<emphasis id="7F88EACDFFDFB82F86C9CF68F113F8F0" bold="true" box="[151,367,1801,1827]" pageId="17" pageNumber="18">Included species </emphasis>
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</subSubSection>
</treatment>
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View file

@ -0,0 +1,150 @@
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<heading id="160B81B3FFE9B81986C9CAD3F4B3FD1E" bold="true" box="[151,1231,690,717]" fontSize="11" level="2" pageId="39" pageNumber="40" reason="4">
<emphasis id="7F88EACDFFE9B81986C9CAD3F4B3FD1E" bold="true" box="[151,1231,690,717]" pageId="39" pageNumber="40">
<taxonomicName id="8AFC4D5CFFE9B81986C9CAD3F421FD1F" authority="(Gerdeman, Garcia, Herczak &amp; Klompen 2018)" baseAuthorityName="Gerdeman, Garcia, Herczak &amp; Klompen" baseAuthorityYear="2018" box="[151,1117,690,717]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="39" pageNumber="40" phylum="Arthropoda" rank="subGenus" subGenus="Philippinozercon">
<emphasis id="7F88EACDFFE9B81986C9CAD3F165FD1E" bold="true" box="[151,281,690,717]" italics="true" pageId="39" pageNumber="40">Allozercon</emphasis>
(
<emphasis id="7F88EACDFFE9B8198775CAD3F186FD1E" bold="true" box="[299,506,690,717]" italics="true" pageId="39" pageNumber="40">Philippinozercon</emphasis>
) (
<bibRefCitation id="296D4B2EFFE9B819844DCAD3F429FD1F" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[531,1109,690,717]" pageId="39" pageNumber="40" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">Gerdeman, Garcia, Herczak &amp; Klompen 2018</bibRefCitation>
)
</taxonomicName>
n. comb.
</emphasis>
</heading>
</paragraph>
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<paragraph id="4D4336DFFFE9B81986C9CA96F32EFCDC" blockId="39.[151,1437,759,814]" box="[151,850,759,783]" pageId="39" pageNumber="40">
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<taxonomicName id="8AFC4D5CFFE9B81986C9CA96F34CFCDC" authority="Gerdeman, Garcia, Herczak &amp; Klompen (2018)" authorityName="Gerdeman, Garcia, Herczak &amp; Klompen" authorityYear="2018" box="[151,816,759,783]" class="Arachnida" family="Heterozerconidae" genus="Philippinozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFE9B81986C9CA96F139FCDC" box="[151,325,759,783]" italics="true" pageId="39" pageNumber="40">Philippinozercon</emphasis>
<bibRefCitation id="296D4B2EFFE9B8198715CA96F34CFCDC" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[331,816,759,783]" pageId="39" pageNumber="40" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">Gerdeman, Garcia, Herczak &amp; Klompen (2018)</bibRefCitation>
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: 10
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<paragraph id="4D4336DFFFE9B8198506CA99F355FCFD" blockId="39.[151,1437,759,814]" pageId="39" pageNumber="40">
[
<typeStatus id="9247887DFFE9B8198500CA99F3F0FCC3" box="[862,908,760,784]" pageId="39" pageNumber="40">type</typeStatus>
species:
<taxonomicName id="8AFC4D5CFFE9B81985B6CA96F25AFCFD" authority="Gerdeman, Garcia, Herczak &amp; Klompen 2018" authorityName="Gerdeman, Garcia, Herczak &amp; Klompen" authorityYear="2018" class="Arachnida" family="Heterozerconidae" genus="Philippinozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="makilingensis">
<emphasis id="7F88EACDFFE9B81985B6CA96F55AFCDC" box="[1000,1318,759,783]" italics="true" pageId="39" pageNumber="40">Philippinozercon makilingensis</emphasis>
<bibRefCitation id="296D4B2EFFE9B8198372CA96F25AFCFD" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." pageId="39" pageNumber="40" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">Gerdeman, Garcia, Herczak &amp; Klompen 2018</bibRefCitation>
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, by original designation].
</paragraph>
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<paragraph id="4D4336DFFFE9B81986C9CB3DF4D4FC30" blockId="39.[151,1437,860,1210]" pageId="39" pageNumber="40">
Diagnosis. Axial outgrowth of palp trochanter in male distinct and membranous (9, reversal?); palp genu seta
<emphasis id="7F88EACDFFE9B81983D8CB3DF5E0FCA5" box="[1414,1436,860,886]" italics="true" pageId="39" pageNumber="40">pl</emphasis>
absent in adults (13); anterior dorsal setae
<emphasis id="7F88EACDFFE9B819842ECBE1F2F9FC49" box="[624,645,896,922]" italics="true" pageId="39" pageNumber="40">j1</emphasis>
minute (&lt;5 µm) (28.2, with few reversals); distinct curved, sclerotized ridge on anterior margin of female genitiventral shield present (46; shared with
<taxonomicName id="8AFC4D5CFFE9B819824FCBC4F4DBFC6D" authorityName="Berlese" authorityYear="1888" box="[1041,1191,933,958]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFE9B819824FCBC4F4DBFC6D" box="[1041,1191,933,958]" italics="true" pageId="39" pageNumber="40">Heterozercon</emphasis>
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); postero-lateral margin of metapodal shields appears cut off, with sharp angle (61), setae
<emphasis id="7F88EACDFFE9B81985FECBA9F3C0FC31" box="[928,956,968,994]" italics="true" pageId="39" pageNumber="40">
S
<quantity id="8A049B3AFFE9B81985F0CBA9F3C0FC31" box="[942,956,968,994]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="39" pageNumber="40" unit="in" value="5.0">5</quantity>
</emphasis>
in male minute (75).
</paragraph>
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<subSubSection id="05E66554FFE9B8198699CB8DF148FB69" pageId="39" pageNumber="40" type="discussion">
<paragraph id="4D4336DFFFE9B8198699CB8DF267FB9C" blockId="39.[151,1437,860,1210]" pageId="39" pageNumber="40">
Remarks. In terms of the number of unique characters,
<taxonomicName id="8AFC4D5CFFE9B8198575CB8DF397FBD5" authorityName="Gerdeman, Garcia, Herczak &amp; Klompen" authorityYear="2018" box="[811,1003,1004,1030]" class="Arachnida" family="Heterozerconidae" genus="Philippinozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFE9B8198575CB8DF397FBD5" box="[811,1003,1004,1030]" italics="true" pageId="39" pageNumber="40">Philippinozercon</emphasis>
</taxonomicName>
has more apomorphies than the majority of heterozerconid genera. Even so, it consistently appears as nested within
<taxonomicName id="8AFC4D5CFFE9B819824FCC71F4F5FBF9" authorityName="Vitzthum" authorityYear="1926" box="[1041,1161,1040,1066]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFE9B819824FCC71F4F5FBF9" box="[1041,1161,1040,1066]" italics="true" pageId="39" pageNumber="40">Allozercon</emphasis>
</taxonomicName>
, and is therefore reclassified as a subgenus of
<taxonomicName id="8AFC4D5CFFE9B81987C2CC55F268FB9D" authorityName="Vitzthum" authorityYear="1926" box="[412,532,1076,1102]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFE9B81987C2CC55F268FB9D" box="[412,532,1076,1102]" italics="true" pageId="39" pageNumber="40">Allozercon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4D4336DFFFE9B8198699CC39F148FB69" blockId="39.[151,1437,860,1210]" pageId="39" pageNumber="40">
<taxonomicName id="8AFC4D5CFFE9B8198699CC39F143FBA1" authorityName="Vitzthum" authorityYear="1926" box="[199,319,1112,1138]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFE9B8198699CC39F143FBA1" box="[199,319,1112,1138]" italics="true" pageId="39" pageNumber="40">Allozercon</emphasis>
</taxonomicName>
<taxonomicName id="8AFC4D5CFFE9B8198716CC39F26AFBA1" authorityName="Gerdeman, Garcia, Herczak &amp; Klompen" authorityYear="2018" box="[328,534,1112,1138]" class="Arachnida" family="Heterozerconidae" genus="Philippinozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="makilingensis" subGenus="Philippinozercon">
(
<emphasis id="7F88EACDFFE9B819870CCC38F11FFBA1" box="[338,355,1113,1138]" italics="true" pageId="39" pageNumber="40">P</emphasis>
.)
<emphasis id="7F88EACDFFE9B8198722CC39F26AFBA1" box="[380,534,1112,1138]" italics="true" pageId="39" pageNumber="40">makilingensis</emphasis>
</taxonomicName>
is one of only two species of
<taxonomicName id="8AFC4D5CFFE9B819852ACC39F445FBA1" authorityName="Berlese" authorityYear="1892" box="[884,1081,1112,1138]" class="Arachnida" family="Heterozerconidae" kingdom="Animalia" order="Mesostigmata" pageId="39" pageNumber="40" phylum="Arthropoda" rank="family">Heterozerconidae</taxonomicName>
for which all immature instars have been described (
<bibRefCitation id="296D4B2EFFE9B81987D6CC1DF2FCFB45" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[392,640,1148,1175]" pageId="39" pageNumber="40" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">
Gerdeman
<emphasis id="7F88EACDFFE9B819845BCC1CF24AFB45" box="[517,566,1148,1174]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
. 2018
</bibRefCitation>
). Basic phenology of this species was described in Gerdeman &amp; Garcia (2009(2010)).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,52 +1,52 @@
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@ -85,7 +85,7 @@ in male absent.
</paragraph>
<paragraph id="4D4336DFFFEAB81A8699CF79F319F885" blockId="36.[151,1436,1744,2023]" pageId="36" pageNumber="37">
Description. Female idiosoma length 1530 (152), width 1287 (186) (N=5); male idiosoma length 1311 (73), width 1058 (32) (N= 5). Complete measurements in
<tableCitation id="007E0364FFEAB81A84BACF5DF31CF885" box="[740,864,1852,1878]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="15.[152,245,151,175]" captionStartId-1="16.[158,251,151,175]" captionText-0="TABLE 3. Comparative measurements (in µm) of newly described Heterozerconidae: females." captionText-1="TABLE 4. Comparative measurements (in µm) of newly described Heterozerconidae: males." pageId="36" pageNumber="37">Tables 34</tableCitation>
<tableCitation id="007E0364FFEAB81A84BACF5DF31CF885" box="[740,864,1852,1878]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="15.[152,245,151,175]" captionStartId-1="16.[158,251,151,175]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="TABLE 3. Comparative measurements (in µm) of newly described Heterozerconidae: females." captionText-1="TABLE 4. Comparative measurements (in µm) of newly described Heterozerconidae: males." pageId="36" pageNumber="37">Tables 34</tableCitation>
.
</paragraph>
<paragraph id="4D4336DFFFEAB81A8699CF01F2C7F834" blockId="36.[151,1436,1744,2023]" pageId="36" pageNumber="37">
@ -287,7 +287,7 @@ absent. Tarsi IV setae
and
<emphasis id="7F88EACDFFE8B8188407CE31F2FDF9B9" box="[601,641,1616,1642]" italics="true" pageId="38" pageNumber="39">pv4</emphasis>
absent. Complete chaetotaxy in
<tableCitation id="007E0364FFE8B81885B6CE31F440F9B9" box="[1000,1084,1616,1642]" captionStart="TABLE 2" captionStartId="14.[152,245,152,176]" captionText="TABLE 2. Leg chaetotaxy in adult Heterozerconoidea." pageId="38" pageNumber="39">Table 2</tableCitation>
<tableCitation id="007E0364FFE8B81885B6CE31F440F9B9" box="[1000,1084,1616,1642]" captionStart="TABLE 2" captionStartId="14.[152,245,152,176]" captionTargetPageId="14" captionText="TABLE 2. Leg chaetotaxy in adult Heterozerconoidea." pageId="38" pageNumber="39">Table 2</tableCitation>
. Coxae I setae in male setiform. Femora I setae
<emphasis id="7F88EACDFFE8B818871CCE12F11AF95E" box="[322,358,1651,1677]" italics="true" pageId="38" pageNumber="39">al1</emphasis>
and
@ -327,15 +327,14 @@ and
setiform.
</paragraph>
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<materialsCitation id="FD943C82FFE8B8188699CF46F2C8F892" box="[199,692,1830,1857]" collectionCode="UPLB" pageId="38" pageNumber="39" specimenCount="1" typeStatus="holotype">
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Type depositories:
<typeStatus id="9247887DFFE8B81887C0CF47F27AF893" box="[414,518,1830,1856]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
male at
<collectionCode id="2BEDAE1AFFE8B8188434CF46F2C8F892" box="[618,692,1831,1857]" country="Philippines" name="Museum of Natural History, University of the Philippines" pageId="38" pageNumber="39">UPLB</collectionCode>
, accession number UPLB MNH ACA-01335.
</materialsCitation>
, accession number UPLB
<materialsCitation id="FD943C82FFE8B81885BCCF46F16CF8B7" collectionCode="UPLB, OSAL" pageId="38" pageNumber="39" specimenCount="1" typeStatus="paratype">
MNH ACA-01335.
<materialsCitation id="FD943C82FFE8B818829DCF46F16CF8B7" collectionCode="UPLB, OSAL" pageId="38" pageNumber="39" specimenCount="1" typeStatus="paratype">
<typeStatus id="9247887DFFE8B818829DCF46F54CF892" box="[1219,1328,1831,1857]" pageId="38" pageNumber="39" type="paratype">Paratypes</typeStatus>
at
<collectionCode id="2BEDAE1AFFE8B818830BCF46F5E1F892" box="[1365,1437,1831,1857]" country="Philippines" name="Museum of Natural History, University of the Philippines" pageId="38" pageNumber="39">UPLB</collectionCode>
@ -345,50 +344,42 @@ and
</paragraph>
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Material examined.
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,
<collectingRegion id="8F38F83DFFE8B818846CCF0FF2C9F85B" box="[562,693,1902,1928]" country="Philippines" name="Southern Leyte" pageId="38" pageNumber="39">Leyte Prov.</collectingRegion>
,
<collectingRegion id="8F38F83DFFE8B81884E1CF0FF283F85B" box="[703,767,1902,1928]" country="Philippines" name="Southern Leyte" pageId="38" pageNumber="39">Leyte</collectingRegion>
Is., VISCA, Hubasan Creek,
, Leyte Is., VISCA, Hubasan Creek,
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<geoCoordinate id="28C85018FFE8B818829CCF0FF539F85B" box="[1218,1349,1902,1928]" degrees="124.8109" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="5" value="124.8109">124.8109 E</geoCoordinate>
,
<date id="3942101FFFE8B818830ECF0FF0AEF878" pageId="38" pageNumber="39" value="2000-06-07">7Jun- 2000</date>
, Gerdeman, Beverly S. &amp; Garcia, Rufino C., ex millipede, BSG 00060745,
<materialsCitation id="FD943C82FFE8B818821CCFF0F528F87F" box="[1090,1364,1937,1964]" collectionCode="UPLB" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="5BFAFD56FFE8B818821CCFF0F4ECF878" box="[1090,1168,1937,1963]" pageId="38" pageNumber="39" type="male">1 male</specimenCount>
,
<typeStatus id="9247887DFFE8B81882C2CFF0F57DF878" box="[1180,1281,1937,1963]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
,
, holotype,
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MNH ACA-01335
</materialsCitation>
MNH ACA-01335. Same data,
<materialsCitation id="FD943C82FFE8B81887E7CFD4F2ABF803" box="[441,727,1973,2000]" collectionCode="OSAL" pageId="38" pageNumber="39" specimenCode="OSAL 053240" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
.
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Same data,
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1F,
; 1F,
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same locality, date and collectors, ex big black millipede,
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00060713,14,15,16,
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<materialsCitation id="FD943C82FFE8B8188582CFBBF42DF820" box="[988,1105,2009,2036]" collectionCode="OSAL" pageId="38" pageNumber="39" specimenCount="1" typeStatus="holotype">same data</materialsCitation>
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same data,
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View file

@ -0,0 +1,193 @@
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<mods:title id="6B85AD406235AB752F409C711536CE32">Genus-level revision of the Heterozerconoidea (Parasitiformes: Mesostigmata)</mods:title>
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<emphasis id="7F88EACDFFEAB81A86C9CC36F110FBA1" bold="true" box="[151,364,1111,1138]" italics="true" pageId="36" pageNumber="37">Allozercon audax</emphasis>
(Berlese 1910)
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<emphasis id="7F88EACDFFEAB81A86C9CCFCF118FB67" box="[151,356,1180,1204]" italics="true" pageId="36" pageNumber="37">Heterozercon audax</emphasis>
<bibRefCitation id="296D4B2EFFEAB81A8734CCFDF24EFB66" author="Berlese, A." box="[362,562,1180,1205]" pageId="36" pageNumber="37" pagination="243 - 271" refId="ref37123" refString="Berlese, A. (1910 b) Lista di nuove specie e nuovi generi di Acari. Redia, Firenze, 6, 243 - 271." type="journal article" year="1910">Berlese 1910b: 247</bibRefCitation>
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.
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<treatmentCitationGroup id="6DEC11F1FFEAB81A86C9CCDDF293FB00" box="[151,751,1211,1235]" pageId="36" pageNumber="37">
<taxonomicName id="8AFC4D5CFFEAB81A86C9CCDDF118FB00" authorityName="Berlese" authorityYear="1910" box="[151,356,1211,1235]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="audax">
<emphasis id="7F88EACDFFEAB81A86C9CCDDF118FB00" box="[151,356,1211,1235]" italics="true" pageId="36" pageNumber="37">Heterozercon audax</emphasis>
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.—
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<bibRefCitation id="296D4B2EFFEAB81A87DDCCDAF238FB00" author="Vitzthum, H. &amp; Graf" box="[387,580,1211,1235]" pageId="36" pageNumber="37" pagination="37 - 48" refId="ref39822" refString="Vitzthum, H., Graf. (1925) Fauna sumatrensis (Beitrag No. 5) Acarinae. Supplementa Entomologica, 11, 37 - 48." type="journal article" year="1925">Vitzthum 1925: 37</bibRefCitation>
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;
<treatmentCitation id="CC5D10CEFFEAB81A8410CCDAF297FB00" author="Fain, A." box="[590,747,1211,1235]" page="145" pageId="36" pageNumber="37" year="1989">
<bibRefCitation id="296D4B2EFFEAB81A8410CCDAF297FB00" author="Fain, A." box="[590,747,1211,1235]" pageId="36" pageNumber="37" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989: 145</bibRefCitation>
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<treatmentCitationGroup id="6DEC11F1FFEAB81A86C9CCBAF23EFB21" box="[151,578,1242,1266]" pageId="36" pageNumber="37">
<treatmentCitation id="CC5D10CEFFEAB81A86C9CCBAF23EFB21" author="Vitzthum, H. &amp; Graf" box="[151,578,1242,1266]" page="106" pageId="36" pageNumber="37" year="1926">
<taxonomicName id="8AFC4D5CFFEAB81A86C9CCBAF23EFB21" authority="Vitzthum 1926: 106" authorityName="Vitzthum" authorityPageNumber="106" authorityYear="1926" box="[151,578,1242,1266]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="elapsus">
<emphasis id="7F88EACDFFEAB81A86C9CCBAF10EFB21" box="[151,370,1242,1266]" italics="true" pageId="36" pageNumber="37">Heterozercon elapsus</emphasis>
<bibRefCitation id="296D4B2EFFEAB81A8729CCBBF23EFB21" author="Vitzthum, H. &amp; Graf" box="[375,578,1242,1266]" pageId="36" pageNumber="37" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926: 106</bibRefCitation>
</taxonomicName>
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[junior synonym of
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<emphasis id="7F88EACDFFEAB81A854CCCBAF3A3FB21" box="[786,991,1242,1266]" italics="true" pageId="36" pageNumber="37">Heterozercon audax</emphasis>
Berlese 1910
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by
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<bibRefCitation id="296D4B2EFFEAB81A82D1CCBBF52EFB21" author="Vitzthum, H. &amp; Graf" box="[1167,1362,1242,1266]" pageId="36" pageNumber="37" pagination="37 - 48" refId="ref39822" refString="Vitzthum, H., Graf. (1925) Fauna sumatrensis (Beitrag No. 5) Acarinae. Supplementa Entomologica, 11, 37 - 48." type="journal article" year="1925">Vitzthum 1925: 37</bibRefCitation>
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].
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Type depository.
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<typeStatus id="9247887DFFEAB81A8700CD41F1BAFAE9" box="[350,454,1312,1338]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="8AFC4D5CFFEAB81A87AFCD40F226FAE9" authorityName="Berlese" authorityYear="1910" box="[497,602,1312,1339]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="audax">
<emphasis id="7F88EACDFFEAB81A87AFCD40F279FAE9" box="[497,517,1313,1338]" italics="true" pageId="36" pageNumber="37">H</emphasis>
.
<emphasis id="7F88EACDFFEAB81A8448CD41F226FAE9" box="[534,602,1312,1338]" italics="true" pageId="36" pageNumber="37">audax</emphasis>
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at
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, accession number
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.
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<paragraph id="4D4336DFFFEAB81A85B6CD41F124FA51" blockId="36.[151,1437,1312,1554]" pageId="36" pageNumber="37">
The
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and unique specimen of
<taxonomicName id="8AFC4D5CFFEAB81A86C9CD24F173FA8D" box="[151,271,1348,1375]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="elapsus">
<emphasis id="7F88EACDFFEAB81A86C9CD24F0D7FA8D" box="[151,171,1349,1374]" italics="true" pageId="36" pageNumber="37">H</emphasis>
.
<emphasis id="7F88EACDFFEAB81A86E5CD25F173FA8D" box="[187,271,1348,1374]" italics="true" pageId="36" pageNumber="37">elapsus</emphasis>
</taxonomicName>
was lost before completion of the description and the description was prepared entirely from memory (
<bibRefCitation id="296D4B2EFFEAB81A86FFCD09F130FA51" author="Vitzthum, H. &amp; Graf" box="[161,332,1384,1410]" pageId="36" pageNumber="37" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926</bibRefCitation>
).
</paragraph>
<paragraph id="4D4336DFFFEAB81A8699CDEDF08CF9C1" blockId="36.[151,1437,1312,1554]" pageId="36" pageNumber="37">
Remarks. Based on females and males from “
<taxonomicName id="8AFC4D5CFFEAB81A8491CDEDF321FA75" box="[719,861,1420,1446]" class="Diplopoda" family="Polyxenidae" genus="Scolopendra" kingdom="Animalia" order="Polyxenida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A8491CDEDF321FA75" box="[719,861,1420,1446]" italics="true" pageId="36" pageNumber="37">Scolopendra</emphasis>
</taxonomicName>
” spp. and “
<emphasis id="7F88EACDFFEAB81A85BBCDEDF4BFFA75" box="[997,1219,1420,1446]" italics="true" pageId="36" pageNumber="37">Spirostreptus depak</emphasis>
” spp. from at least three different localities on
<collectingRegion id="8F38F83DFFEAB81A8789CDD0F277FA18" box="[471,523,1457,1483]" country="Indonesia" name="Jawa Timur" pageId="36" pageNumber="37">Java</collectingRegion>
,
<collectingCountry id="35EB764FFFEAB81A8447CDD1F2FAFA19" box="[537,646,1456,1482]" name="Indonesia" pageId="36" pageNumber="37">Indonesia</collectingCountry>
(
<bibRefCitation id="296D4B2EFFEAB81A84C9CDD1F33DFA19" author="Berlese, A." box="[663,833,1456,1482]" pageId="36" pageNumber="37" pagination="243 - 271" refId="ref37123" refString="Berlese, A. (1910 b) Lista di nuove specie e nuovi generi di Acari. Redia, Firenze, 6, 243 - 271." type="journal article" year="1910">Berlese 1910b</bibRefCitation>
);
<taxonomicName id="8AFC4D5CFFEAB81A8509CDD0F3ACFA19" box="[855,976,1456,1483]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="elapsus">
<emphasis id="7F88EACDFFEAB81A8509CDD0F317FA19" box="[855,875,1457,1482]" italics="true" pageId="36" pageNumber="37">H</emphasis>
.
<emphasis id="7F88EACDFFEAB81A8522CDD1F3ACFA19" box="[892,976,1456,1482]" italics="true" pageId="36" pageNumber="37">elapsus</emphasis>
</taxonomicName>
on
<taxonomicName id="8AFC4D5CFFEAB81A85A1CDD1F4D0FA18" box="[1023,1196,1456,1483]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="7F88EACDFFEAB81A85A1CDD1F4FEFA19" box="[1023,1154,1456,1482]" italics="true" pageId="36" pageNumber="37">Thyropygus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="8AFC4D5CFFEAB81A82E3CDD1F51CFA19" authorityName="Brandt" authorityYear="1833" box="[1213,1376,1456,1482]" class="Diplopoda" kingdom="Animalia" order="Spirostreptida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="order">Spirostreptida</taxonomicName>
:
<taxonomicName id="8AFC4D5CFFEAB81A8330CDD1F161FA3D" authorityName="Pocock" authorityYear="1894" class="Diplopoda" family="Spirostreptidae" kingdom="Animalia" order="Spirostreptida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="family">Spirostreptidae</taxonomicName>
) collected on
<collectingRegion id="8F38F83DFFEAB81A87E1CDB5F263FA3D" box="[447,543,1492,1518]" country="Indonesia" name="Sumatera Utara" pageId="36" pageNumber="37">Sumatra</collectingRegion>
,
<collectingCountry id="35EB764FFFEAB81A8472CDB5F2E5FA3D" box="[556,665,1492,1518]" name="Indonesia" pageId="36" pageNumber="37">Indonesia</collectingCountry>
(
<bibRefCitation id="296D4B2EFFEAB81A84F5CDB5F324FA3D" author="Vitzthum, H. &amp; Graf" box="[683,856,1492,1518]" pageId="36" pageNumber="37" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926</bibRefCitation>
).
<bibRefCitation id="296D4B2EFFEAB81A8532CDB5F38DFA3D" author="Fain, A." box="[876,1009,1492,1518]" pageId="36" pageNumber="37" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain (1989)</bibRefCitation>
noted that
<taxonomicName id="8AFC4D5CFFEAB81A822DCDB4F497FA3D" box="[1139,1259,1492,1519]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="elapsus">
<emphasis id="7F88EACDFFEAB81A822DCDB4F4FBFA3D" box="[1139,1159,1493,1518]" italics="true" pageId="36" pageNumber="37">H</emphasis>
.
<emphasis id="7F88EACDFFEAB81A82C9CDB5F497FA3D" box="[1175,1259,1492,1518]" italics="true" pageId="36" pageNumber="37">elapsus</emphasis>
</taxonomicName>
may be a valid species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,353 @@
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<taxonomicName id="8AFC4D5CFFEDB81D86C9CC7BF1A5FBE7" authority="Vitzthum 1926" authorityName="Vitzthum" authorityYear="1926" box="[151,473,1050,1077]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D86C9CC7BF1A5FBE7" bold="true" box="[151,473,1050,1077]" pageId="35" pageNumber="36">
<emphasis id="7F88EACDFFEDB81D86C9CC7BF165FBE6" bold="true" box="[151,281,1050,1077]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
<bibRefCitation id="296D4B2EFFEDB81D877ECC7BF1A5FBE7" author="Vitzthum, H. &amp; Graf" box="[288,473,1050,1076]" pageId="35" pageNumber="36" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926</bibRefCitation>
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<taxonomicName id="8AFC4D5CFFEDB81D86C9CC3EF1A9FBA4" authority="Vitzthum 1926: 104" authorityName="Vitzthum" authorityPageNumber="104" authorityYear="1926" box="[151,469,1119,1143]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D86C9CC3EF178FBA4" box="[151,260,1119,1143]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
<bibRefCitation id="296D4B2EFFEDB81D8754CC3EF1A9FBA4" author="Vitzthum, H. &amp; Graf" box="[266,469,1119,1143]" pageId="35" pageNumber="36" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926: 104</bibRefCitation>
</taxonomicName>
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<subSubSection id="05E66554FFEDB81D8785CC01F56CFBA4" box="[475,1296,1119,1144]" pageId="35" pageNumber="36" type="type_taxon">
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[
<typeStatus id="9247887DFFEDB81D87BFCC01F273FBAB" box="[481,527,1120,1144]" pageId="35" pageNumber="36">type</typeStatus>
species.
<taxonomicName id="8AFC4D5CFFEDB81D8435CC3EF470FBA4" authority="Vitzthum 1926" authorityName="Vitzthum" authorityYear="1926" box="[619,1036,1119,1143]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="fecundissimus">
<emphasis id="7F88EACDFFEDB81D8435CC3EF311FBA4" box="[619,877,1119,1143]" italics="true" pageId="35" pageNumber="36">Allozercon fecundissimus</emphasis>
<bibRefCitation id="296D4B2EFFEDB81D852DCC3EF470FBA4" author="Vitzthum, H. &amp; Graf" box="[883,1036,1119,1143]" pageId="35" pageNumber="36" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926</bibRefCitation>
</taxonomicName>
, by original designation].
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<emphasis id="7F88EACDFFEDB81D86C9CC1FF178FB45" box="[151,260,1150,1174]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
</taxonomicName>
.—
<treatmentCitation id="CC5D10CEFFEDB81D877FCC1FF27BFB45" author="Womersley, H." box="[289,519,1150,1174]" page="129" pageId="35" pageNumber="36" year="1958">
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</treatmentCitation>
;
<treatmentCitation id="CC5D10CEFFEDB81D844DCC1FF30BFB45" author="Flechtmann, C. H. W. &amp; Johnston, D. E." box="[531,887,1150,1174]" page="147" pageId="35" pageNumber="36" year="1990">
<bibRefCitation id="296D4B2EFFEDB81D844DCC1FF30BFB45" author="Flechtmann, C. H. W. &amp; Johnston, D. E." box="[531,887,1150,1174]" pageId="35" pageNumber="36" pagination="143 - 148" refId="ref37819" refString="Flechtmann, C. H. W. &amp; Johnston, D. E. (1990) Zeterohercon, a new genus of Heterozerconidae (Acari: Mesostigmata) and the description of Zeterohercon amphisbaenae n. sp. from Brasil. International Journal of Acarology, 16, 143 - 148. https: // doi. org / 10.1080 / 01647959008683526" type="journal article" year="1990">Flechtmann &amp; Johnston 1990: 147</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="CC5D10CEFFEDB81D85DCCC1FF4C9FB45" author="Gerdeman, B. S. &amp; Garcia, R." box="[898,1205,1150,1174]" page="93" pageId="35" pageNumber="36" year="2010">
<bibRefCitation id="296D4B2EFFEDB81D85DCCC1FF4C9FB45" author="Gerdeman, B. S. &amp; Garcia, R." box="[898,1205,1150,1174]" pageId="35" pageNumber="36" pagination="93 - 96" refId="ref37991" refString="Gerdeman, B. S. &amp; Garcia, R. (2010) Heterozerconidae: A comparison between a temperate and a tropical species. In: Sabelis, M. W. &amp; Bruin, J. (Eds.), Trends in Acarology, Proceedings of the 12 th International Congress. Springer Science, Dordrecht, pp. 93 - 96. https: // doi. org / 10.1007 / 978 - 90 - 481 - 9837 - 5 _ 14" type="book chapter" year="2010">Gerdeman &amp; Garcia 2010: 93</bibRefCitation>
</treatmentCitation>
;
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<bibRefCitation id="296D4B2EFFEDB81D829FCC1FF08CFB67" author="Di Palma, A. &amp; de Moraes, G. J. &amp; Gerdeman, B. S. &amp; Huber, S. &amp; Kitajima, E. W. &amp; Alberti, G." pageId="35" pageNumber="36" pagination="639 - 655" refId="ref37329" refString="Di Palma, A., de Moraes, G. J., Gerdeman, B. S., Huber, S., Kitajima, E. W. &amp; Alberti, G. (2015) Ultrastructural and functional adaptations of the female reproductive system in the family Heterozerconidae (Acari, Anactinotrichida, Gamasida, Heterozerconina) and implications for the systematic position of the group. Arthropod Structure &amp; Development, 44, 639 - 655. https: // doi. org / 10.1016 / j. asd. 2015.09.002" type="journal article" year="2015">
Di Palma
<emphasis id="7F88EACDFFEDB81D8376CC1EF529FB45" box="[1320,1365,1150,1174]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. 2015: 640
</bibRefCitation>
</treatmentCitation>
;
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<bibRefCitation id="296D4B2EFFEDB81D86A5CCFDF18EFB66" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[251,498,1180,1205]" pageId="35" pageNumber="36" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">
Gerdeman
<emphasis id="7F88EACDFFEDB81D8734CCFCF1EBFB67" box="[362,407,1180,1204]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. 2018: 9
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.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4D4336DFFFEDB81D86C9CCDAF190FB00" blockId="35.[151,1437,1119,1297]" box="[151,492,1211,1235]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="6DEC11F1FFEDB81D86C9CCDAF190FB00" box="[151,492,1211,1235]" pageId="35" pageNumber="36">
<treatmentCitation id="CC5D10CEFFEDB81D86C9CCDAF190FB00" author="Fain, A." box="[151,492,1211,1235]" page="146" pageId="35" pageNumber="36" year="1989">
<taxonomicName id="8AFC4D5CFFEDB81D86C9CCDAF190FB00" authority="Fain 1989: 146" authorityName="Fain" authorityPageNumber="146" authorityYear="1989" box="[151,492,1211,1235]" class="Arachnida" family="Heterozerconidae" genus="Asioheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D86C9CCDAF13AFB00" box="[151,326,1211,1235]" italics="true" pageId="35" pageNumber="36">Asioheterozercon</emphasis>
<bibRefCitation id="296D4B2EFFEDB81D8711CCDAF190FB00" author="Fain, A." box="[335,492,1211,1235]" pageId="35" pageNumber="36" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989: 146</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEDB81D87ABCCDAF3B9FB21" pageId="35" pageNumber="36" type="multiple">
<paragraph id="4D4336DFFFEDB81D87ABCCDAF3B9FB21" blockId="35.[151,1437,1119,1297]" pageId="35" pageNumber="36">
(
<typeStatus id="9247887DFFEDB81D87A2CCDAF255FB00" box="[508,553,1211,1235]" pageId="35" pageNumber="36">type</typeStatus>
species
<taxonomicName id="8AFC4D5CFFEDB81D84DDCCDDF39CFB00" authority="Berlese 1910" authorityName="Berlese" authorityYear="1910" box="[643,992,1211,1235]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="audax">
<emphasis id="7F88EACDFFEDB81D84DDCCDDF32DFB00" box="[643,849,1211,1235]" italics="true" pageId="35" pageNumber="36">Heterozercon audax</emphasis>
Berlese 1910
</taxonomicName>
by original designation);
<treatmentCitation id="CC5D10CEFFEDB81D82B0CCDDF5E0FB00" author="Evans, G. O." box="[1262,1436,1211,1235]" page="426" pageId="35" pageNumber="36" year="1992">
<bibRefCitation id="296D4B2EFFEDB81D82B0CCDDF5E0FB00" author="Evans, G. O." box="[1262,1436,1211,1235]" pageId="35" pageNumber="36" refId="ref37637" refString="Evans, G. O. (1992) Principles of acarology. CAB International, Wallingford, 563 pp. https: // doi. org / 10.1079 / 9780851988221.0000" type="book" year="1992">Evans 1992: 426</bibRefCitation>
</treatmentCitation>
[junior synonym of
<taxonomicName id="8AFC4D5CFFEDB81D87CFCCBBF2E7FB21" authority="Vitzthum 1926" authorityName="Vitzthum" authorityYear="1926" box="[401,667,1242,1266]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D87CFCCBBF182FB21" box="[401,510,1242,1266]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
<bibRefCitation id="296D4B2EFFEDB81D845ACCBBF2E7FB21" author="Vitzthum, H. &amp; Graf" box="[516,667,1242,1266]" pageId="35" pageNumber="36" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926</bibRefCitation>
</taxonomicName>
by
<treatmentCitation id="CC5D10CEFFEDB81D849FCCBBF3C6FB21" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[705,954,1242,1266]" page="9" pageId="35" pageNumber="36" year="2018">
<bibRefCitation id="296D4B2EFFEDB81D849FCCBBF3C6FB21" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[705,954,1242,1266]" pageId="35" pageNumber="36" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">
Gerdeman
<emphasis id="7F88EACDFFEDB81D856FCCBAF321FB21" box="[817,861,1242,1266]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. 2018: 9
</bibRefCitation>
</treatmentCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEDB81D86C9CC99F382FAC3" box="[151,1022,1272,1297]" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="4D4336DFFFEDB81D86C9CC99F382FAC3" blockId="35.[151,1437,1119,1297]" box="[151,1022,1272,1297]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="6DEC11F1FFEDB81D86C9CC99F382FAC3" box="[151,1022,1272,1297]" pageId="35" pageNumber="36">
<taxonomicName id="8AFC4D5CFFEDB81D86C9CC99F138FAC3" authority="Fain 1989" box="[151,324,1272,1296]" class="Arachnida" family="Heterozerconidae" genus="Alloheterozercon" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D86C9CC99F138FAC3" box="[151,324,1272,1296]" italics="true" pageId="35" pageNumber="36">Alloheterozercon</emphasis>
</taxonomicName>
.—
<bibRefCitation id="296D4B2EFFEDB81D873FCC99F2C4FAC3" author="Gerdeman, B. S. &amp; Klompen, H." box="[353,696,1272,1297]" pageId="35" pageNumber="36" pagination="351 - 370" refId="ref38078" refString="Gerdeman, B. S. &amp; Klompen, H. (2003) A new North American heterozerconid, Narceoheterozercon ohioensis, n. gen., n. sp., with first description of immatures of Heterozerconidae (Acari: Mesostigmata). International Journal of Acarology, 29, 351 - 370. https: // doi. org / 10.1080 / 01647950308684352" type="journal article" year="2003">Gerdeman &amp; Klompen 2003: 353</bibRefCitation>
[incorrect subsequent spelling].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEDB81D86C9CD5EF1F4F9DE" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="4D4336DFFFEDB81D86C9CD5EF1F4F9DE" blockId="35.[151,1437,1343,2017]" pageId="35" pageNumber="36">
Diagnosis. Axial outgrowth of palp trochanter in male distinct and sclerotized, solid (9); setae
<emphasis id="7F88EACDFFEDB81D8297CD5EF483FA8A" box="[1225,1279,1343,1369]" italics="true" pageId="35" pageNumber="36">hyp2</emphasis>
at least twice as long as setae
<emphasis id="7F88EACDFFEDB81D870ACD02F1F6FAAE" box="[340,394,1379,1405]" italics="true" pageId="35" pageNumber="36">hyp3</emphasis>
(22); insertion sternal setae
<emphasis id="7F88EACDFFEDB81D848CCD02F28EFAAE" box="[722,754,1379,1405]" italics="true" pageId="35" pageNumber="36">st2</emphasis>
and
<emphasis id="7F88EACDFFEDB81D8573CD02F332FAAE" box="[813,846,1379,1405]" italics="true" pageId="35" pageNumber="36">
st
<quantity id="8A049B3AFFEDB81D851ECD02F332FAAE" box="[832,846,1379,1405]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="35" pageNumber="36" unit="in" value="3.0">3</quantity>
</emphasis>
in female and
<emphasis id="7F88EACDFFEDB81D85A3CD02F462FAAE" box="[1021,1054,1379,1405]" italics="true" pageId="35" pageNumber="36">
st
<quantity id="8A049B3AFFEDB81D824ECD02F462FAAE" box="[1040,1054,1379,1405]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="35" pageNumber="36" unit="in" value="3.0">3</quantity>
</emphasis>
in male on isolated platelets (38, 40, 52); insertion sternal setae
<emphasis id="7F88EACDFFEDB81D87A8CDE6F26BFA72" box="[502,535,1415,1441]" italics="true" pageId="35" pageNumber="36">
st
<quantity id="8A049B3AFFEDB81D8455CDE6F26BFA72" box="[523,535,1415,1441]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="35" pageNumber="36" unit="in" value="1.0">1</quantity>
</emphasis>
in male on isolated platelets (48); opisthogastral setae
<emphasis id="7F88EACDFFEDB81D82D4CDE6F4CDFA72" box="[1162,1201,1415,1441]" italics="true" pageId="35" pageNumber="36">Jv5</emphasis>
inserted posterior to ventrianal fusion line (71); all marginal opisthosomal setae setiform (77, reversal); femora II setae
<emphasis id="7F88EACDFFEDB81D82B3CDCAF57FFA16" box="[1261,1283,1451,1477]" italics="true" pageId="35" pageNumber="36">al</emphasis>
,
<emphasis id="7F88EACDFFEDB81D834FCDCAF55BFA16" box="[1297,1319,1451,1477]" italics="true" pageId="35" pageNumber="36">pl</emphasis>
, and
<emphasis id="7F88EACDFFEDB81D833BCDCAF503FA17" box="[1381,1407,1451,1476]" italics="true" pageId="35" pageNumber="36">av</emphasis>
in male spines (105107, shared with
<taxonomicName id="8AFC4D5CFFEDB81D8479CDAEF2B5FA3B" authorityName="Fain" authorityYear="1989" box="[551,713,1487,1512]" class="Arachnida" family="Heterozerconidae" genus="Maracazercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D8479CDAEF2B5FA3B" box="[551,713,1487,1512]" italics="true" pageId="35" pageNumber="36">Maracazercon</emphasis>
</taxonomicName>
and
<taxonomicName id="8AFC4D5CFFEDB81D855FCDAEF3EBFA3B" authorityName="Berlese" authorityYear="1888" box="[769,919,1487,1512]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D855FCDAEF3EBFA3B" box="[769,919,1487,1512]" italics="true" pageId="35" pageNumber="36">Heterozercon</emphasis>
</taxonomicName>
); genual and tibial setae
<emphasis id="7F88EACDFFEDB81D82EDCDAEF4B1FA3B" box="[1203,1229,1487,1512]" italics="true" pageId="35" pageNumber="36">pv</emphasis>
on legs II in male spine-like (109, 110).
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEDB81A8699CE76F12AFE26" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="discussion">
<paragraph id="4D4336DFFFEDB81D8699CE76F450F86E" blockId="35.[151,1437,1343,2017]" pageId="35" pageNumber="36">
Remarks. A re-evaluation of the 11 characters listed by
<bibRefCitation id="296D4B2EFFEDB81D856DCE76F3CAF9E2" author="Fain, A." box="[819,950,1559,1585]" pageId="35" pageNumber="36" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain (1989)</bibRefCitation>
for the genus
<taxonomicName id="8AFC4D5CFFEDB81D8208CE76F564F9E2" authorityName="Fain" authorityYear="1989" box="[1110,1304,1559,1585]" class="Arachnida" family="Heterozerconidae" genus="Asioheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D8208CE76F564F9E2" box="[1110,1304,1559,1585]" italics="true" pageId="35" pageNumber="36">Asioheterozercon</emphasis>
</taxonomicName>
, shows that two are included in the updated diagnosis of
<taxonomicName id="8AFC4D5CFFEDB81D84D2CE5AF378F986" authorityName="Vitzthum" authorityYear="1926" box="[652,772,1595,1621]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D84D2CE5AF378F986" box="[652,772,1595,1621]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
</taxonomicName>
listed above: palpal trochanter with its apico-internal angle strongly produced and bearing 2 setae (our character 9) and anal shield with five setae (71). Three of the remaining characters are variable among the S.E. Asian populations: tarsus and tibia of legs I much narrower than the other segments of that leg (81.2); anterior region of the dorsum with 2040 long and stiff setae (29.1); antero-lateral margin of dorsal shield in male with numerous small spines (30.1). One is incorrect: palp trochanter in female bears only one seta. Seta
<emphasis id="7F88EACDFFEDB81D8764CE8EF128F8DA" box="[314,340,1775,1801]" italics="true" pageId="35" pageNumber="36">v2</emphasis>
is in fact present and is inserted on the membranous axial outgrowth of the trochanter. However, it is highly reduced (
<figureCitation id="D5C72A5AFFEDB81D87DACF72F1A6F8FE" box="[388,474,1811,1837]" captionStart="FIGURES 3942" captionStartId="45.[151,264,1909,1933]" captionTargetBox="[226,1361,184,1882]" captionTargetId="figure-21@45.[223,1364,181,1885]" captionTargetPageId="45" captionText="FIGURES 3942. Heterozerconidae, gnathosoma, adults: 39, Allozercon leytensis n. sp., female (OSAL 053243); 40, Allozercon leytensis n. sp., male (UPLB MNH ACA-01335; red arrow: sclerotized axial extension of palp trochanter); 41, Al. (Philippinozercon) makilingensis, female (OSAL 053271); 42, Al. (Philippinozercon) makilingensis male (OSAL 102161). Inset: gnathotectum and labrum + salivary stylets." figureDoi="http://doi.org/10.5281/zenodo.8203685" httpUri="https://zenodo.org/record/8203685/files/figure.png" pageId="35" pageNumber="36">Fig. 39</figureCitation>
). The remaining characters are imprecise or refer to primitive character states at this taxonomic level. To further clarify the status of
<taxonomicName id="8AFC4D5CFFEDB81D84E6CF56F34CF882" authorityName="Vitzthum" authorityYear="1926" box="[696,816,1847,1873]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEDB81D84E6CF56F34CF882" box="[696,816,1847,1873]" italics="true" pageId="35" pageNumber="36">Allozercon</emphasis>
</taxonomicName>
a complete description is provided for one of the nine geographically separate populations for which we have specimens available. These nine populations originate in the
<collectingCountry id="35EB764FFFEDB81D869ECF1EF142F84A" box="[192,318,1919,1945]" name="Philippines" pageId="35" pageNumber="36">Philippines</collectingCountry>
(
<collectingRegion id="8F38F83DFFEDB81D8710CF1EF1EBF84A" box="[334,407,1919,1945]" country="Philippines" name="Bohol" pageId="35" pageNumber="36">Bohol</collectingRegion>
,
<collectingRegion id="8F38F83DFFEDB81D87FFCF1EF198F84A" box="[417,484,1919,1945]" country="Philippines" name="Southern Leyte" pageId="35" pageNumber="36">Leyte</collectingRegion>
, Mindoro,
<collectingRegion id="8F38F83DFFEDB81D8401CF1EF2D4F84A" box="[607,680,1919,1945]" country="Philippines" name="Samar" pageId="35" pageNumber="36">Samar</collectingRegion>
, and
<collectingRegion id="8F38F83DFFEDB81D84BDCF1EF341F84A" box="[739,829,1919,1945]" country="Philippines" name="Siquijor" pageId="35" pageNumber="36">Siquijor</collectingRegion>
islands),
<collectingCountry id="35EB764FFFEDB81D85F7CF1EF471F84A" box="[937,1037,1919,1945]" name="Thailand" pageId="35" pageNumber="36">Thailand</collectingCountry>
(
<collectingRegion id="8F38F83DFFEDB81D8245CF1EF400F84A" box="[1051,1148,1919,1945]" country="Thailand" name="Sara Buri" pageId="35" pageNumber="36">Saraburi</collectingRegion>
and
<collectingRegion id="8F38F83DFFEDB81D82EDCF1EF561F84A" box="[1203,1309,1919,1945]" country="Thailand" name="Krung Thep Mahanakhon" pageId="35" pageNumber="36">Bangkok</collectingRegion>
),
<collectingCountry id="35EB764FFFEDB81D8371CF1EF514F84A" box="[1327,1384,1919,1945]" name="Laos" pageId="35" pageNumber="36">Laos</collectingCountry>
, and
<collectingCountry id="35EB764FFFEDB81D86C9CFC2F0A8F86E" box="[151,212,1955,1981]" name="India" pageId="35" pageNumber="36">India</collectingCountry>
. The remaining taxa will be described in an upcoming revision of the genus.
</paragraph>
<paragraph id="4D4336DFFFEDB81A8699CFA6F20CFE92" blockId="35.[151,1437,1343,2017]" lastBlockId="36.[151,1437,151,501]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<bibRefCitation id="296D4B2EFFEDB81D8699CFA6F1B9F832" author="Di Palma, A. &amp; de Moraes, G. J. &amp; Gerdeman, B. S. &amp; Huber, S. &amp; Kitajima, E. W. &amp; Alberti, G." box="[199,453,1990,2017]" pageId="35" pageNumber="36" pagination="639 - 655" refId="ref37329" refString="Di Palma, A., de Moraes, G. J., Gerdeman, B. S., Huber, S., Kitajima, E. W. &amp; Alberti, G. (2015) Ultrastructural and functional adaptations of the female reproductive system in the family Heterozerconidae (Acari, Anactinotrichida, Gamasida, Heterozerconina) and implications for the systematic position of the group. Arthropod Structure &amp; Development, 44, 639 - 655. https: // doi. org / 10.1016 / j. asd. 2015.09.002" type="journal article" year="2015">
Di Palma
<emphasis id="7F88EACDFFEDB81D8766CFA6F116F832" box="[312,362,1991,2017]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. (2015)
</bibRefCitation>
, in a study of the anatomy of the female genital system, found no indication of secondary genital openings (solenostomes) in females of an unidentified
<taxonomicName id="8AFC4D5CFFEAB81A852AC8F6F43FFF62" box="[884,1091,151,177]" pageId="36" pageNumber="37">
<emphasis id="7F88EACDFFEAB81A852AC8F6F390FF62" box="[884,1004,151,177]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
species
</taxonomicName>
from
<collectingRegion id="8F38F83DFFEAB81A82D8C8F6F4BDFF62" box="[1158,1217,151,177]" country="Philippines" name="Cebu" pageId="36" pageNumber="37">Cebu</collectingRegion>
island,
<collectingCountry id="35EB764FFFEAB81A8349C8F6F5E5FF62" box="[1303,1433,151,177]" name="Philippines" pageId="36" pageNumber="37">Philippines</collectingCountry>
. Examination of all available slide mounted
<taxonomicName id="8AFC4D5CFFEAB81A8429C8DAF293FF06" authorityName="Vitzthum" authorityYear="1926" box="[631,751,187,213]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A8429C8DAF293FF06" box="[631,751,187,213]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
</taxonomicName>
females also failed to find any indication of secondary genital openings either near the opisthogastral suckers (as in
<taxonomicName id="8AFC4D5CFFEAB81A84B7C8BEF303FF2B" authorityName="Berlese" authorityYear="1888" box="[745,895,223,248]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A84B7C8BEF303FF2B" box="[745,895,223,248]" italics="true" pageId="36" pageNumber="37">Heterozercon</emphasis>
</taxonomicName>
or
<taxonomicName id="8AFC4D5CFFEAB81A85FDC8BEF4F8FF2A" authorityName="Gerdeman &amp; Klompen" authorityYear="2003" box="[931,1156,223,249]" class="Arachnida" family="Heterozerconidae" genus="Narceoheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A85FDC8BEF4F8FF2A" box="[931,1156,223,249]" italics="true" pageId="36" pageNumber="37">Narceoheterozercon</emphasis>
</taxonomicName>
) or posterior to coxae IV (as in
<taxonomicName id="8AFC4D5CFFEAB81A8685C962F1E0FECE" authorityName="Fain" authorityYear="1989" box="[219,412,259,285]" class="Arachnida" family="Heterozerconidae" genus="Afroheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A8685C962F1E0FECE" box="[219,412,259,285]" italics="true" pageId="36" pageNumber="37">Afroheterozercon</emphasis>
</taxonomicName>
). In the absence of evidence to the contrary, we assume that
<taxonomicName id="8AFC4D5CFFEAB81A821DC962F56FFECE" box="[1091,1299,259,285]" pageId="36" pageNumber="37">
<emphasis id="7F88EACDFFEAB81A821DC962F4C7FECE" box="[1091,1211,259,285]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
species
</taxonomicName>
do not have a secondary genital system in the females.
</paragraph>
<paragraph id="4D4336DFFFEAB81A8699C92AF3C0FE5A" blockId="36.[151,1437,151,501]" pageId="36" pageNumber="37">
Immatures of an
<taxonomicName id="8AFC4D5CFFEAB81A87DFC92AF22CFEB6" box="[385,592,331,357]" pageId="36" pageNumber="37">
<emphasis id="7F88EACDFFEAB81A87DFC92AF185FEB6" box="[385,505,331,357]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
species
</taxonomicName>
from
<collectingRegion id="8F38F83DFFEAB81A84CFC92AF285FEB6" box="[657,761,331,357]" country="Thailand" name="Krung Thep Mahanakhon" pageId="36" pageNumber="37">Bangkok</collectingRegion>
,
<collectingCountry id="35EB764FFFEAB81A855CC92AF31AFEB6" box="[770,870,331,357]" name="Thailand" pageId="36" pageNumber="37">Thailand</collectingCountry>
were partially described in
<bibRefCitation id="296D4B2EFFEAB81A82CBC92AF5E4FEB6" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." box="[1173,1432,331,357]" pageId="36" pageNumber="37" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">
Gerdeman
<emphasis id="7F88EACDFFEAB81A8351C92AF542FEB6" box="[1295,1342,331,357]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. (2018)
</bibRefCitation>
, but a complete description of
<taxonomicName id="8AFC4D5CFFEAB81A87BBC90EF221FE5A" authorityName="Vitzthum" authorityYear="1926" box="[485,605,367,393]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A87BBC90EF221FE5A" box="[485,605,367,393]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
</taxonomicName>
immatures is not yet available.
</paragraph>
<paragraph id="4D4336DFFFEAB81A8699C9F2F12AFE26" blockId="36.[151,1437,151,501]" pageId="36" pageNumber="37">
<taxonomicName id="8AFC4D5CFFEAB81A8699C9F2F201FE7E" authority="Blaszak 1984" authorityName="Blaszak" authorityYear="1984" box="[199,637,403,429]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A8699C9F2F143FE7E" box="[199,319,403,429]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
<bibRefCitation id="296D4B2EFFEAB81A8717C9F2F19FFE7E" author="Blaszak, C." box="[329,483,403,429]" pageId="36" pageNumber="37" pagination="587 - 601" refId="ref37228" refString="Blaszak, C. (1984) Three new genera of zerconid mites (Acari, Mesostigmata, Zerconidae) from the United States of America with a key to the American genera. Polskie Pismo Entomologiczne, 53, 587 - 601." type="journal article" year="1984">Błaszak 1984</bibRefCitation>
(Zerconidae)
</taxonomicName>
is a junior homonym of
<taxonomicName id="8AFC4D5CFFEAB81A85FCC9F2F4B1FE7E" authority="Vitzthum 1926" authorityName="Vitzthum" authorityYear="1926" box="[930,1229,403,429]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A85FCC9F2F466FE7E" box="[930,1050,403,429]" italics="true" pageId="36" pageNumber="37">Allozercon</emphasis>
<bibRefCitation id="296D4B2EFFEAB81A827DC9F2F4B1FE7E" author="Vitzthum, H. &amp; Graf" box="[1059,1229,403,429]" pageId="36" pageNumber="37" pagination="1 - 198" refId="ref39851" refString="Vitzthum, H., Graf. (1926) Malayische Acari. Treubia, Buitenzorg, 8, 1 - 198." type="journal article" year="1926">Vitzthum 1926</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="296D4B2EFFEAB81A8280C9F2F0AEFE02" author="Kemal, M. &amp; Kocak, A. O." pageId="36" pageNumber="37" pagination="147 - 148" refId="ref38442" refString="Kemal, M. &amp; Kocak, A. O. (2009) A replacement name for Blaszakia Kocak &amp; Kemal, 2008 in the family Zerconidae (Acarina, Mesostigmata). Miscellaneous Papers of the Centre for Entomological Studies Ankara, 147 - 148, 10." type="book chapter" year="2009">Kemal &amp; Koçak 2009</bibRefCitation>
;
<bibRefCitation id="296D4B2EFFEAB81A8681C9D6F1A2FE02" author="Kocak, A. O. &amp; Kemal, M." box="[223,478,439,465]" pageId="36" pageNumber="37" pagination="1 - 6" refId="ref38707" refString="Kocak, A. O. &amp; Kemal, M. (2008) Nomenclatural notes on the genus group names of the order Acarina. Miscellaneous Papers of the Centre for Entomological Studies Ankara, 145, 1 - 6." type="journal article" year="2008">Koçak &amp; Kemal 2008</bibRefCitation>
). The zerconid genus name was replaced by
<taxonomicName id="8AFC4D5CFFEAB81A8584C9D6F12EFE26" authority="Kemal &amp; Kocak (Gerdeman et al. 2018)" class="Arachnida" family="Zerconidae" genus="Blaszakzercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEAB81A8584C9D6F407FE02" box="[986,1147,439,465]" italics="true" pageId="36" pageNumber="37">Blaszakzercon</emphasis>
Kemal &amp; Koçak (
<bibRefCitation id="296D4B2EFFEAB81A8313C9D6F136FE26" author="Gerdeman, B. S. &amp; Garcia, R. C. &amp; Herczak, A. &amp; Klompen, H." pageId="36" pageNumber="37" pagination="7 - 22" refId="ref38189" refString="Gerdeman, B. S., Garcia, R. C., Herczak, A. &amp; Klompen, H. (2018) Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars. Zootaxa, 4540 (1), 7 - 22. https: // doi. org / 10.11646 / zootaxa. 4540.1.4" type="journal article" year="2018">
Gerdeman
<emphasis id="7F88EACDFFEAB81A8690C9BAF17CFE26" box="[206,256,475,501]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. 2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,153 @@
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<paragraph id="4D4336DFFFEEB81E86C9C99BF1B4FDC7" blockId="32.[151,456,505,532]" box="[151,456,505,532]" pageId="32" pageNumber="33">
<heading id="160B81B3FFEEB81E86C9C99BF1B4FDC7" bold="true" box="[151,456,505,532]" fontSize="11" level="2" pageId="32" pageNumber="33" reason="4">
<taxonomicName id="8AFC4D5CFFEEB81E86C9C99BF1B4FDC7" authority="Fain 1989" authorityName="Fain" authorityYear="1989" box="[151,456,505,532]" class="Arachnida" family="Heterozerconidae" genus="Maracazercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E86C9C99BF1B4FDC7" bold="true" box="[151,456,505,532]" pageId="32" pageNumber="33">
<emphasis id="7F88EACDFFEEB81E86C9C99BF139FDC7" bold="true" box="[151,325,506,532]" italics="true" pageId="32" pageNumber="33">Maracazercon</emphasis>
<bibRefCitation id="296D4B2EFFEEB81E8713C998F1B4FDC7" author="Fain, A." box="[333,456,505,532]" pageId="32" pageNumber="33" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989</bibRefCitation>
</emphasis>
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<subSubSection id="05E66554FFEEB81E86C9CA5EF1C2FD85" box="[151,446,574,598]" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="4D4336DFFFEEB81E86C9CA5EF1C2FD85" blockId="32.[151,1180,574,599]" box="[151,446,574,598]" pageId="32" pageNumber="33">
<emphasis id="7F88EACDFFEEB81E86C9CA5EF163FD85" box="[151,287,575,598]" italics="true" pageId="32" pageNumber="33">Maracazeron</emphasis>
<bibRefCitation id="296D4B2EFFEEB81E877BCA5FF1C2FD85" author="Fain, A." box="[293,446,574,598]" pageId="32" pageNumber="33" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989: 148</bibRefCitation>
</paragraph>
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<paragraph id="4D4336DFFFEEB81E879BCA5EF4E0FD85" blockId="32.[151,1180,574,599]" box="[453,1180,574,599]" pageId="32" pageNumber="33">
[
<typeStatus id="9247887DFFEEB81E8795CA5EF185FD84" box="[459,505,575,599]" pageId="32" pageNumber="33">type</typeStatus>
species:
<emphasis id="7F88EACDFFEEB81E8408CA5EF2A2FD85" box="[598,734,575,598]" italics="true" pageId="32" pageNumber="33">Maracazeron</emphasis>
<taxonomicName id="8AFC4D5CFFEEB81E84BACA5FF3E5FD85" authority="Fain 1989" authorityName="Fain" authorityYear="1989" box="[740,921,574,598]" class="Arachnida" family="Heterozerconidae" genus="Maracazercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="joliveti">
<emphasis id="7F88EACDFFEEB81E84BACA5FF356FD85" box="[740,810,574,598]" italics="true" pageId="32" pageNumber="33">joliveti</emphasis>
<bibRefCitation id="296D4B2EFFEEB81E856ECA5FF3E5FD85" author="Fain, A." box="[816,921,574,598]" pageId="32" pageNumber="33" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989</bibRefCitation>
</taxonomicName>
, by original designation].
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEEB81E86C9CAE5F142FC80" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="4D4336DFFFEEB81E86C9CAE5F142FC80" blockId="32.[151,1436,644,995]" pageId="32" pageNumber="33">
Diagnosis. Distance from setae
<emphasis id="7F88EACDFFEEB81E845DCAE5F245FD4D" box="[515,569,644,670]" italics="true" pageId="32" pageNumber="33">hyp3</emphasis>
to
<emphasis id="7F88EACDFFEEB81E843FCAE4F204FD4D" box="[609,632,645,670]" italics="true" pageId="32" pageNumber="33">sc</emphasis>
distinctly smaller than distance from
<emphasis id="7F88EACDFFEEB81E8272CAE5F41EFD4D" box="[1068,1122,644,670]" italics="true" pageId="32" pageNumber="33">hyp3</emphasis>
to
<emphasis id="7F88EACDFFEEB81E82D4CAE5F4BDFD4D" box="[1162,1217,644,670]" italics="true" pageId="32" pageNumber="33">hyp2</emphasis>
(21.0); platelets of sternal setae
<emphasis id="7F88EACDFFEEB81E8770CAC8F132FD10" box="[302,334,681,707]" italics="true" pageId="32" pageNumber="33">st2</emphasis>
and
<emphasis id="7F88EACDFFEEB81E87D5CAC8F1D1FD10" box="[395,429,681,707]" italics="true" pageId="32" pageNumber="33">
st
<quantity id="8A049B3AFFEEB81E87C1CAC8F1D1FD10" box="[415,429,681,707]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="32" pageNumber="33" unit="in" value="3.0">3</quantity>
</emphasis>
in male completely fused (51); endopodal and sternitiventral shield in male fused (58, shared with
<taxonomicName id="8AFC4D5CFFEEB81E877CCAADF19FFD35" authorityName="Fain" authorityYear="1989" box="[290,483,716,742]" class="Arachnida" family="Heterozerconidae" genus="Afroheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E877CCAADF19FFD35" box="[290,483,716,742]" italics="true" pageId="32" pageNumber="33">Afroheterozercon</emphasis>
</taxonomicName>
); postanal seta less than 1/3 the length of paranal setae (73); marginal opisthosomal setae hook-like spines (78, shared with
<taxonomicName id="8AFC4D5CFFEEB81E8415CA90F29DFCD9" authorityName="Berlese" authorityYear="1892" box="[587,737,753,778]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E8415CA90F29DFCD9" box="[587,737,753,778]" italics="true" pageId="32" pageNumber="33">Heterozercon</emphasis>
</taxonomicName>
); femora II setae
<emphasis id="7F88EACDFFEEB81E85FDCA91F3C5FCD9" box="[931,953,752,778]" italics="true" pageId="32" pageNumber="33">al</emphasis>
,
<emphasis id="7F88EACDFFEEB81E8598CA91F3A0FCD9" box="[966,988,752,778]" italics="true" pageId="32" pageNumber="33">pl</emphasis>
, and
<emphasis id="7F88EACDFFEEB81E8249CA90F44DFCD9" box="[1047,1073,753,778]" italics="true" pageId="32" pageNumber="33">av</emphasis>
in male spines (105107, shared with
<taxonomicName id="8AFC4D5CFFEEB81E868ECB74F11AFCFD" authorityName="Berlese" authorityYear="1892" box="[208,358,789,814]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E868ECB74F11AFCFD" box="[208,358,789,814]" italics="true" pageId="32" pageNumber="33">Heterozercon</emphasis>
</taxonomicName>
and
<taxonomicName id="8AFC4D5CFFEEB81E87C0CB75F26AFCFD" box="[414,534,788,814]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E87C0CB75F26AFCFD" box="[414,534,788,814]" italics="true" pageId="32" pageNumber="33">Allozercon</emphasis>
</taxonomicName>
); setae
<emphasis id="7F88EACDFFEEB81E8430CB75F2EEFCFD" box="[622,658,788,814]" italics="true" pageId="32" pageNumber="33">pl1</emphasis>
on tarsi II spine-like (111; shared with
<taxonomicName id="8AFC4D5CFFEEB81E8210CB74F4ADFCFD" authorityName="Klompen &amp; Gerdeman" authorityYear="2023" box="[1102,1233,789,814]" class="Arachnida" family="Heterozerconidae" genus="Ecuazercon" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E8210CB74F4ADFCFD" box="[1102,1233,789,814]" italics="true" pageId="32" pageNumber="33">Ecuazercon</emphasis>
</taxonomicName>
,
<taxonomicName id="8AFC4D5CFFEEB81E82BECB75F5E9FCFD" authorityName="Fain" authorityYear="1989" box="[1248,1429,788,814]" class="Arachnida" family="Heterozerconidae" genus="Amheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E82BECB75F5E9FCFD" box="[1248,1429,788,814]" italics="true" pageId="32" pageNumber="33">Amheterozercon</emphasis>
</taxonomicName>
,
<taxonomicName id="8AFC4D5CFFEEB81E86C9CB58F151FC81" authorityName="Berlese" authorityYear="1892" box="[151,301,825,850]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E86C9CB58F151FC81" box="[151,301,825,850]" italics="true" pageId="32" pageNumber="33">Heterozercon</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEEB81E8699CB3DF349FC31" pageId="32" pageNumber="33" type="discussion">
<paragraph id="4D4336DFFFEEB81E8699CB3DF349FC31" blockId="32.[151,1436,644,995]" pageId="32" pageNumber="33">
Remarks. The above diagnosis adds some characteristics of the leg setation to
<bibRefCitation id="296D4B2EFFEEB81E8270CB3DF4BFFCA4" author="Fain, A." box="[1070,1219,860,887]" pageId="32" pageNumber="33" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fains (1989)</bibRefCitation>
diagnosis. This genus shares with
<taxonomicName id="8AFC4D5CFFEEB81E8716CBE1F275FC49" authorityName="Fain" authorityYear="1989" box="[328,521,896,922]" class="Arachnida" family="Heterozerconidae" genus="Afroheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E8716CBE1F275FC49" box="[328,521,896,922]" italics="true" pageId="32" pageNumber="33">Afroheterozercon</emphasis>
</taxonomicName>
the fusion of the endopodal and sternitiventral shields in the male, but differs from that genus by the lack of fusion between the metapodal and sternitiventral shields (58, 59). The resulting pattern of the male ventral shields is unique among
<taxonomicName id="8AFC4D5CFFEEB81E8439CBA9F34CFC31" authorityName="Berlese" authorityYear="1892" box="[615,816,968,994]" class="Arachnida" family="Heterozerconidae" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="family">Heterozerconidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,269 @@
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<paragraph id="4D4336DFFFEEB81E86C9CDBEF1A3FA2A" blockId="32.[151,479,1503,1529]" box="[151,479,1503,1529]" pageId="32" pageNumber="33">
<heading id="160B81B3FFEEB81E86C9CDBEF1A3FA2A" bold="true" box="[151,479,1503,1529]" fontSize="11" level="2" pageId="32" pageNumber="33" reason="4">
<taxonomicName id="8AFC4D5CFFEEB81E86C9CDBEF1A3FA2A" authority="Berlese 1892" authorityName="Berlese" authorityYear="1892" box="[151,479,1503,1529]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E86C9CDBEF1A3FA2A" bold="true" box="[151,479,1503,1529]" pageId="32" pageNumber="33">
<emphasis id="7F88EACDFFEEB81E86C9CDBEF146FA2A" bold="true" box="[151,314,1503,1529]" italics="true" pageId="32" pageNumber="33">Heterozercon</emphasis>
<bibRefCitation id="296D4B2EFFEEB81E871FCDBEF1A3FA2A" author="Berlese, A." box="[321,479,1503,1529]" pageId="32" pageNumber="33" refId="ref37026" refString="Berlese, A. (1892) Acari, Myriapoda et Scorpiones hucusque in Italia reperta. Ordo Mesostigmata (Gamasidae). Tipografia del Seminario, Padua, 143 pp., XI pls." type="book" year="1892">Berlese 1892</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEEB81E86C9CE45F1A2F9E8" box="[151,478,1571,1596]" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="4D4336DFFFEEB81E86C9CE45F1A2F9E8" blockId="32.[151,1197,1571,1626]" box="[151,478,1571,1596]" pageId="32" pageNumber="33">
<treatmentCitationGroup id="6DEC11F1FFEEB81E86C9CE45F1A2F9E8" box="[151,478,1571,1596]" pageId="32" pageNumber="33">
<treatmentCitation id="CC5D10CEFFEEB81E86C9CE45F1A2F9E8" author="Berlese, A." box="[151,478,1571,1596]" page="206" pageId="32" pageNumber="33" year="1888">
<taxonomicName id="8AFC4D5CFFEEB81E86C9CE45F1A2F9E8" authority="Berlese 1888: 206" authorityName="Berlese" authorityPageNumber="206" authorityYear="1888" box="[151,478,1571,1596]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E86C9CE45F163F9E8" box="[151,287,1572,1595]" italics="true" pageId="32" pageNumber="33">Heterozercon</emphasis>
<bibRefCitation id="296D4B2EFFEEB81E8778CE45F1A2F9E8" author="Berlese, A." box="[294,478,1571,1596]" pageId="32" pageNumber="33" pagination="171 - 222" refId="ref36987" refString="Berlese, A. (1888) Acari austro-Americani quos collegit Aloysius Balzan. Manipulus primus. Species novas circiter quinquaginta complectens. Bollettino della Societa Entomologica Italiana, 20, 171 - 222, pl. X." type="journal article" year="1888">Berlese 1888: 206</bibRefCitation>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEEB81E87BACE45F4D1F9EF" box="[484,1197,1572,1596]" pageId="32" pageNumber="33" type="type_taxon">
<paragraph id="4D4336DFFFEEB81E87BACE45F4D1F9EF" blockId="32.[151,1197,1571,1626]" box="[484,1197,1572,1596]" pageId="32" pageNumber="33">
[
<typeStatus id="9247887DFFEEB81E87B4CE45F264F9EF" box="[490,536,1572,1596]" pageId="32" pageNumber="33">type</typeStatus>
species:
<taxonomicName id="8AFC4D5CFFEEB81E842BCE45F471F9EF" authority="Berlese 1888" authorityName="Berlese" authorityYear="1888" box="[629,1037,1572,1596]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="degeneratus">
<emphasis id="7F88EACDFFEEB81E842BCE45F303F9EF" box="[629,895,1572,1596]" italics="true" pageId="32" pageNumber="33">Heterozercon degeneratus</emphasis>
<bibRefCitation id="296D4B2EFFEEB81E85D8CE45F471F9EF" author="Berlese, A." box="[902,1037,1572,1596]" pageId="32" pageNumber="33" pagination="171 - 222" refId="ref36987" refString="Berlese, A. (1888) Acari austro-Americani quos collegit Aloysius Balzan. Manipulus primus. Species novas circiter quinquaginta complectens. Bollettino della Societa Entomologica Italiana, 20, 171 - 222, pl. X." type="journal article" year="1888">Berlese 1888</bibRefCitation>
</taxonomicName>
, by monotypy].
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEEB81E86C9CE22F457F989" box="[151,1067,1602,1626]" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="4D4336DFFFEEB81E86C9CE22F457F989" blockId="32.[151,1197,1571,1626]" box="[151,1067,1602,1626]" pageId="32" pageNumber="33">
<taxonomicName id="8AFC4D5CFFEEB81E86C9CE22F163F989" authorityName="Berlese" authorityYear="1888" box="[151,287,1603,1626]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E86C9CE22F163F989" box="[151,287,1603,1626]" italics="true" pageId="32" pageNumber="33">Heterozercon</emphasis>
</taxonomicName>
.—
<bibRefCitation id="296D4B2EFFEEB81E8763CE23F1DBF989" author="Fain, A." box="[317,423,1602,1626]" pageId="32" pageNumber="33" pagination="145 - 156" refId="ref37719" refString="Fain, A. (1989) Notes on mites associated with Myriapoda. IV. New taxa in the Heterozerconidae (Acari, Mesostigmata). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Entomologie, 59, 145 - 156." type="journal article" year="1989">Fain 1989</bibRefCitation>
, 148;
<bibRefCitation id="296D4B2EFFEEB81E87BBCE22F293F989" author="Krantz, G. W. &amp; de Moraes, G. J." box="[485,751,1602,1626]" pageId="32" pageNumber="33" pagination="24 - 33" refId="ref38749" refString="Krantz, G. W. &amp; de Moraes, G. J. (2011) Discovery and description of nymphal stages of a heterozerconid mite (Acari: Mesostigmata: Heterozerconidae) from coastal forest litter in southeastern S &quot; o Paulo State, Brazil. In: de Moraes, G. J. &amp; Proctor, H. (Eds.), Acarology XIII: Proceedings of the International Congress, 2011, pp. 24 - 33. https: // doi. org / 10.11646 / zoosymposia. 6.1.5" type="journal article" year="2011">Krantz &amp; de Moraes 2011</bibRefCitation>
, 26;
<bibRefCitation id="296D4B2EFFEEB81E857ECE23F38FF989" author="Di Palma, A. &amp; de Moraes, G. J. &amp; Gerdeman, B. S. &amp; Huber, S. &amp; Kitajima, E. W. &amp; Alberti, G." box="[800,1011,1602,1626]" pageId="32" pageNumber="33" pagination="639 - 655" refId="ref37329" refString="Di Palma, A., de Moraes, G. J., Gerdeman, B. S., Huber, S., Kitajima, E. W. &amp; Alberti, G. (2015) Ultrastructural and functional adaptations of the female reproductive system in the family Heterozerconidae (Acari, Anactinotrichida, Gamasida, Heterozerconina) and implications for the systematic position of the group. Arthropod Structure &amp; Development, 44, 639 - 655. https: // doi. org / 10.1016 / j. asd. 2015.09.002" type="journal article" year="2015">
Di Palma
<emphasis id="7F88EACDFFEEB81E85DBCE22F3CEF989" box="[901,946,1602,1626]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. 2015
</bibRefCitation>
, 640.
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEEB81E86C9CEE9F20FF84C" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="4D4336DFFFEEB81E86C9CEE9F20FF84C" blockId="32.[151,1437,1672,2023]" pageId="32" pageNumber="33">
Diagnosis. Spermatodactyl shape flaccid (7); distance from setae
<emphasis id="7F88EACDFFEEB81E85DECEE9F3CBF971" box="[896,951,1672,1698]" italics="true" pageId="32" pageNumber="33">hyp3</emphasis>
to
<emphasis id="7F88EACDFFEEB81E8583CEE8F388F971" box="[989,1012,1673,1698]" italics="true" pageId="32" pageNumber="33">sc</emphasis>
distinctly smaller than distance from
<emphasis id="7F88EACDFFEEB81E86C9CECDF0B1F915" box="[151,205,1708,1734]" italics="true" pageId="32" pageNumber="33">hyp3</emphasis>
to
<emphasis id="7F88EACDFFEEB81E86ADCECDF156F915" box="[243,298,1708,1734]" italics="true" pageId="32" pageNumber="33">hyp2</emphasis>
(21.0, reversed in some species); anterior dorsal setae
<emphasis id="7F88EACDFFEEB81E85C6CECCF3D1F914" box="[920,941,1709,1735]" italics="true" pageId="32" pageNumber="33">j1</emphasis>
minute (&lt;5 µm) (28.2, few exceptions); female with a distinct curved, sclerotized ridge on anterior margin of female genitiventral shield (46; shared with
<emphasis id="7F88EACDFFEEB81E8322CEB1F5E9F939" box="[1404,1429,1744,1770]" italics="true" pageId="32" pageNumber="33">Al</emphasis>
. (
<taxonomicName id="8AFC4D5CFFEEB81E86FFCE95F11DF8DD" box="[161,353,1780,1806]" class="Arachnida" family="Heterozerconidae" genus="Philippinozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E86FFCE95F11DF8DD" box="[161,353,1780,1806]" italics="true" pageId="32" pageNumber="33">Philippinozercon</emphasis>
</taxonomicName>
)); sclerotized areas near sternal setae
<emphasis id="7F88EACDFFEEB81E8553CE94F351F8DC" box="[781,813,1781,1807]" italics="true" pageId="32" pageNumber="33">
st
<quantity id="8A049B3AFFEEB81E857ECE94F351F8DC" box="[800,813,1781,1807]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="32" pageNumber="33" unit="in" value="2.0">2</quantity>
</emphasis>
in male forming isolated platelets (50); marginal opisthosomal setae hook-like spines (78, shared with
<taxonomicName id="8AFC4D5CFFEEB81E84E5CF78F321F8E1" authorityName="Fain" authorityYear="1989" box="[699,861,1817,1842]" class="Arachnida" family="Heterozerconidae" genus="Maracazercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E84E5CF78F321F8E1" box="[699,861,1817,1842]" italics="true" pageId="32" pageNumber="33">Maracazercon</emphasis>
</taxonomicName>
); coxae I setae in male distinct spines (102, shared with
<taxonomicName id="8AFC4D5CFFEEB81E868ECF5DF1CDF885" authorityName="Gerdeman &amp; Klompen" authorityYear="2003" box="[208,433,1852,1878]" class="Arachnida" family="Heterozerconidae" genus="Narceoheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E868ECF5DF1CDF885" box="[208,433,1852,1878]" italics="true" pageId="32" pageNumber="33">Narceoheterozercon</emphasis>
</taxonomicName>
ex Alabama-C and a few
<taxonomicName id="8AFC4D5CFFEEB81E848DCF5DF38DF885" box="[723,1009,1852,1878]" pageId="32" pageNumber="33">
<emphasis id="7F88EACDFFEEB81E848DCF5DF3E8F885" box="[723,916,1852,1878]" italics="true" pageId="32" pageNumber="33">Afroheterozercon</emphasis>
species
</taxonomicName>
); femora II setae
<emphasis id="7F88EACDFFEEB81E82ECCF5DF4B4F885" box="[1202,1224,1852,1878]" italics="true" pageId="32" pageNumber="33">al</emphasis>
,
<emphasis id="7F88EACDFFEEB81E828BCF5DF497F885" box="[1237,1259,1852,1878]" italics="true" pageId="32" pageNumber="33">pl</emphasis>
, and
<emphasis id="7F88EACDFFEEB81E8376CF5CF53EF885" box="[1320,1346,1853,1878]" italics="true" pageId="32" pageNumber="33">av</emphasis>
in male spines (105107, shared with
<taxonomicName id="8AFC4D5CFFEEB81E87B8CF00F2F4F8A9" authorityName="Fain" authorityYear="1989" box="[486,648,1889,1914]" class="Arachnida" family="Heterozerconidae" genus="Maracazercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E87B8CF00F2F4F8A9" box="[486,648,1889,1914]" italics="true" pageId="32" pageNumber="33">Maracazercon</emphasis>
</taxonomicName>
and
<taxonomicName id="8AFC4D5CFFEEB81E84E0CF01F34AF8A9" box="[702,822,1888,1914]" class="Arachnida" family="Heterozerconidae" genus="Allozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E84E0CF01F34AF8A9" box="[702,822,1888,1914]" italics="true" pageId="32" pageNumber="33">Allozercon</emphasis>
</taxonomicName>
); setae
<emphasis id="7F88EACDFFEEB81E85D3CF01F3CDF8A9" box="[909,945,1888,1914]" italics="true" pageId="32" pageNumber="33">pl1</emphasis>
on tarsi II spine-like (111, shared with
<taxonomicName id="8AFC4D5CFFEEB81E8339CF00F093F84D" authorityName="Klompen &amp; Gerdeman" authorityYear="2023" class="Arachnida" family="Heterozerconidae" genus="Ecuazercon" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E8339CF00F093F84D" italics="true" pageId="32" pageNumber="33">Ecuazercon</emphasis>
</taxonomicName>
,
<taxonomicName id="8AFC4D5CFFEEB81E86A3CFE5F1CEF84D" authorityName="Fain" authorityYear="1989" box="[253,434,1924,1950]" class="Arachnida" family="Heterozerconidae" genus="Amheterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E86A3CFE5F1CEF84D" box="[253,434,1924,1950]" italics="true" pageId="32" pageNumber="33">Amheterozercon</emphasis>
</taxonomicName>
,
<taxonomicName id="8AFC4D5CFFEEB81E879ECFE4F21EF84D" authorityName="Fain" authorityYear="1989" box="[448,610,1925,1950]" class="Arachnida" family="Heterozerconidae" genus="Maracazercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E879ECFE4F21EF84D" box="[448,610,1925,1950]" italics="true" pageId="32" pageNumber="33">Maracazercon</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="05E66554FFEEB81F8699CFC9F325FDB2" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="discussion">
<paragraph id="4D4336DFFFEEB81F8699CFC9F5FBFEB6" blockId="32.[151,1437,1672,2023]" lastBlockId="33.[151,1437,151,609]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
Remarks. Available descriptions for the three described species are incomplete, a situation that is problematic especially given that
<taxonomicName id="8AFC4D5CFFEEB81E87D8CFACF260F835" authorityName="Berlese" authorityYear="1888" box="[390,540,1997,2022]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEEB81E87D8CFACF260F835" box="[390,540,1997,2022]" italics="true" pageId="32" pageNumber="33">Heterozercon</emphasis>
</taxonomicName>
is the
<typeStatus id="9247887DFFEEB81E8436CFACF2E4F834" box="[616,664,1997,2023]" pageId="32" pageNumber="33">type</typeStatus>
genus of the family. It suggests a need for a more complete description. We have examined specimens of undescribed populations that belong to the genus
<taxonomicName id="8AFC4D5CFFEFB81F8223C8F6F56FFF63" authorityName="Berlese" authorityYear="1888" box="[1149,1299,151,176]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="7F88EACDFFEFB81F8223C8F6F56FFF63" box="[1149,1299,151,176]" italics="true" pageId="33" pageNumber="34">Heterozercon</emphasis>
</taxonomicName>
from
<collectingCountry id="35EB764FFFEFB81F8304C8F6F5E4FF62" box="[1370,1432,151,177]" name="Cuba" pageId="33" pageNumber="34">Cuba</collectingCountry>
,
<collectingCountry id="35EB764FFFEFB81F86C9C8DAF171FF06" box="[151,269,187,213]" name="Venezuela" pageId="33" pageNumber="34">Venezuela</collectingCountry>
,
<collectingCountry id="35EB764FFFEFB81F8749C8DAF122FF06" box="[279,350,187,213]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
,
<collectingCountry id="35EB764FFFEFB81F8736C8DAF198FF06" box="[360,484,187,213]" name="Costa Rica" pageId="33" pageNumber="34">Costa Rica</collectingCountry>
,
<collectingCountry id="35EB764FFFEFB81F87B0C8DAF218FF06" box="[494,612,187,213]" name="Nicaragua" pageId="33" pageNumber="34">Nicaragua</collectingCountry>
,
<collectingCountry id="35EB764FFFEFB81F8430C8DAF2C4FF06" box="[622,696,187,213]" name="Belize" pageId="33" pageNumber="34">Belize</collectingCountry>
, and
<collectingCountry id="35EB764FFFEFB81F84AFC8DAF336FF06" box="[753,842,187,213]" name="Mexico" pageId="33" pageNumber="34">Mexico</collectingCountry>
. The specimens from
<collectingCountry id="35EB764FFFEFB81F8265C8DAF40AFF06" box="[1083,1142,187,213]" name="Cuba" pageId="33" pageNumber="34">Cuba</collectingCountry>
and
<collectingCountry id="35EB764FFFEFB81F82F4C8DAF561FF06" box="[1194,1309,187,213]" name="Venezuela" pageId="33" pageNumber="34">Venezuela</collectingCountry>
stand apart because of the heavy cuticular patterning on the venter (
<figureCitation id="D5C72A5AFFEFB81F8554C8BEF327FF2A" box="[778,859,222,249]" captionStart="FIGURES 4852" captionStartId="48.[151,264,1906,1930]" captionTargetBox="[235,1353,181,1883]" captionTargetId="figure-25@48.[235,1353,181,1883]" captionTargetPageId="48" captionText="FIGURES 4852. Heterozerconidae, details idiosoma: 48, Allozercon leytensis n. sp., female, hypertrichy of setae in j1 position (OSAL 053251); 49, same for male (OSAL 106780); 50, Allozercon leytensis n. sp., male, anterolateral spines (OSAL 106780); 50a, detail antero-lateral spines; 51, Heterozercon ex Cuba, female, venter (OSAL 125446); 52, Narceoheterozercon sp., broken off spermatodactyl (red arrow) in female secondary genital opening / sclerotized tube (OSAL 159712)." figureDoi="http://doi.org/10.5281/zenodo.8203697" httpUri="https://zenodo.org/record/8203697/files/figure.png" pageId="33" pageNumber="34">Fig. 51</figureCitation>
) and population
<collectingCountry id="35EB764FFFEFB81F8246C8BEF4EEFF2A" box="[1048,1170,223,249]" name="Costa Rica" pageId="33" pageNumber="34">Costa Rica</collectingCountry>
2 is unusual because of the extremely small opisthogastral suckers (much smaller than in
<taxonomicName id="8AFC4D5CFFEFB81F852FC962F42BFECE" authority="Fain" authorityName="Fain" authorityYear="1989" box="[881,1111,259,285]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="microsuctus">
<emphasis id="7F88EACDFFEFB81F852FC962F3F9FECF" box="[881,901,259,284]" italics="true" pageId="33" pageNumber="34">H</emphasis>
.
<emphasis id="7F88EACDFFEFB81F85CDC962F465FECF" box="[915,1049,259,284]" italics="true" pageId="33" pageNumber="34">microsuctus</emphasis>
Fain
</taxonomicName>
). For this reason, we selected specimens from
<collectingRegion id="8F38F83DFFEFB81F8713C946F180FE92" box="[333,508,295,321]" country="Mexico" name="Campeche" pageId="33" pageNumber="34">Campeche state</collectingRegion>
in
<collectingCountry id="35EB764FFFEFB81F8443C946F20FFE92" box="[541,627,295,321]" name="Mexico" pageId="33" pageNumber="34">Mexico</collectingCountry>
for description. This population is considered more representative for the genus. A more extensive revision of the
<taxonomicName id="8AFC4D5CFFEFB81F8407C92AF33BFEB6" box="[601,839,331,357]" pageId="33" pageNumber="34">
<emphasis id="7F88EACDFFEFB81F8407C92AF293FEB7" box="[601,751,331,356]" italics="true" pageId="33" pageNumber="34">Heterozercon</emphasis>
species
</taxonomicName>
of
<collectingCountry id="35EB764FFFEFB81F8532C92AF3BEFEB6" box="[876,962,331,357]" name="Mexico" pageId="33" pageNumber="34">Mexico</collectingCountry>
is in progress (Martinez
<emphasis id="7F88EACDFFEFB81F8284C92AF570FEB6" box="[1242,1292,331,357]" italics="true" pageId="33" pageNumber="34">et al</emphasis>
., in prep.).
</paragraph>
<paragraph id="4D4336DFFFEFB81F8699C90EF3CDFDCA" blockId="33.[151,1437,151,609]" pageId="33" pageNumber="34">
The female genital system of an undescribed
<taxonomicName id="8AFC4D5CFFEFB81F8494C90EF3C5FE5A" box="[714,953,367,393]" pageId="33" pageNumber="34">
<emphasis id="7F88EACDFFEFB81F8494C90EF31CFE5B" box="[714,864,367,392]" italics="true" pageId="33" pageNumber="34">Heterozercon</emphasis>
species
</taxonomicName>
from
<collectingCountry id="35EB764FFFEFB81F85A1C90EF43FFE5A" box="[1023,1091,367,393]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
was described by
<bibRefCitation id="296D4B2EFFEFB81F8348C90EF175FE7E" author="Di Palma, A. &amp; de Moraes, G. J. &amp; Gerdeman, B. S. &amp; Huber, S. &amp; Kitajima, E. W. &amp; Alberti, G." pageId="33" pageNumber="34" pagination="639 - 655" refId="ref37329" refString="Di Palma, A., de Moraes, G. J., Gerdeman, B. S., Huber, S., Kitajima, E. W. &amp; Alberti, G. (2015) Ultrastructural and functional adaptations of the female reproductive system in the family Heterozerconidae (Acari, Anactinotrichida, Gamasida, Heterozerconina) and implications for the systematic position of the group. Arthropod Structure &amp; Development, 44, 639 - 655. https: // doi. org / 10.1016 / j. asd. 2015.09.002" type="journal article" year="2015">
Di Palma
<emphasis id="7F88EACDFFEFB81F83D6C90EF0D1FE7E" italics="true" pageId="33" pageNumber="34">et al</emphasis>
. (2015)
</bibRefCitation>
. These authors noted the presence of secondary genital openings (solenostomes) in females on the axial side of the opisthogastral suckers. This observation was confirmed in four different species examined in this study (
<taxonomicName id="8AFC4D5CFFEFB81F86FFC9BAF11CFE26" authority="Klompen &amp; Gerdeman, 2023" authorityName="Klompen &amp; Gerdeman" authorityYear="2023" box="[161,352,475,501]" class="Arachnida" family="Heterozerconidae" genus="Heterozercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="calakmulensis" status="sp. nov.">
<emphasis id="7F88EACDFFEFB81F86FFC9BAF0C9FE27" box="[161,181,475,500]" italics="true" pageId="33" pageNumber="34">H</emphasis>
.
<emphasis id="7F88EACDFFEFB81F869FC9BAF11CFE26" box="[193,352,475,501]" italics="true" pageId="33" pageNumber="34">calakmulensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64BB57B6FFEFB81F8738C9BAF1DEFE26" box="[358,418,475,501]" pageId="33" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
, unidentified populations from
<collectingCountry id="35EB764FFFEFB81F84A0C9BAF339FE26" box="[766,837,475,501]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
,
<collectingCountry id="35EB764FFFEFB81F8511C9BAF3B7FE26" box="[847,971,475,501]" name="Costa Rica" pageId="33" pageNumber="34">Costa Rica</collectingCountry>
,
<collectingCountry id="35EB764FFFEFB81F858BC9BAF469FE26" box="[981,1045,475,501]" name="Cuba" pageId="33" pageNumber="34">Cuba</collectingCountry>
). The openings are not very distinct and may be associated with small, sclerotized patches (
<figureCitation id="D5C72A5AFFEFB81F855EC99EF32FFDCA" box="[768,851,511,537]" captionStart="FIGURES 6162" captionStartId="51.[151,264,1011,1035]" captionTargetBox="[247,1340,187,985]" captionTargetId="figure-21@51.[242,1345,181,988]" captionTargetPageId="51" captionText="FIGURES 6162. Heterozerconidae, Narceoheterozercon ohioensis, idiosoma venter: 61, female; 62, male. Male redrawn from Gerdeman &amp; Klompen (2003)." figureDoi="http://doi.org/10.5281/zenodo.8203711" httpUri="https://zenodo.org/record/8203711/files/figure.png" pageId="33" pageNumber="34">Fig. 61</figureCitation>
, arrow).
</paragraph>
<paragraph id="4D4336DFFFEFB81F8699CA42F325FDB2" blockId="33.[151,1437,151,609]" pageId="33" pageNumber="34">
Nymphs that probably belong to this genus have been described by
<bibRefCitation id="296D4B2EFFEFB81F8590CA42F575FDEE" author="Krantz, G. W. &amp; de Moraes, G. J." box="[974,1289,547,573]" pageId="33" pageNumber="34" pagination="24 - 33" refId="ref38749" refString="Krantz, G. W. &amp; de Moraes, G. J. (2011) Discovery and description of nymphal stages of a heterozerconid mite (Acari: Mesostigmata: Heterozerconidae) from coastal forest litter in southeastern S &quot; o Paulo State, Brazil. In: de Moraes, G. J. &amp; Proctor, H. (Eds.), Acarology XIII: Proceedings of the International Congress, 2011, pp. 24 - 33. https: // doi. org / 10.11646 / zoosymposia. 6.1.5" type="journal article" year="2011">Krantz &amp; de Moraes (2011)</bibRefCitation>
from
<collectingCountry id="35EB764FFFEFB81F830FCA42F5E4FDEE" box="[1361,1432,547,573]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
, although definitive associations with adults could not be made.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,52 +1,52 @@
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@ -92,7 +92,7 @@ minute, much smaller than
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Description. Female idiosoma length 834, width 698 (N=1); male idiosoma length 712, width 598 (N=1). All measurements in
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.
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@ -276,7 +276,7 @@ absent. Tarsi IV setae
and
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absent. Complete chaetotaxy in
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. Both coxae I setae in male distinct spines with rounded tips. Femora I setae
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and
@ -330,12 +330,15 @@ and
setiform.
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Type depository.
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and female
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,
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View file

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[
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Locality: Bogor (as Buitenzorg),
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Remarks. The description of this monotypic genus is based entirely on the dorsum and the pattern of long setae on the legs, as the venter in both available specimens was not visible (
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).
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noted some resemblance with
<taxonomicName id="8AFC4D5CFFF2B802873FCEFAF255F966" authorityName="Berlese" authorityYear="1892" box="[353,553,1691,1717]" class="Arachnida" family="Heterozerconidae" kingdom="Animalia" order="Mesostigmata" pageId="60" pageNumber="61" phylum="Arthropoda" rank="family">Heterozerconidae</taxonomicName>
, and this species has occasionally been grouped in
<taxonomicName id="8AFC4D5CFFF2B8028203CEFAF55AF966" authorityName="Berlese" authorityYear="1892" box="[1117,1318,1691,1717]" class="Arachnida" family="Heterozerconidae" kingdom="Animalia" order="Mesostigmata" pageId="60" pageNumber="61" phylum="Arthropoda" rank="family">Heterozerconidae</taxonomicName>
. However, the combination of relatively short cheliceral digits, presence of an added pair of long setae on the prodorsum (in addition to the setae referred to as
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j
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in this study), six pairs of long postero-lateral and posterior setae, the insertion pattern of the long filiform setae on legs I (multiple long setae on the genua, tibiae, and tarsi; shorter ones on the femora), and the absence of any indication of long spine-like setae on femora I, make it unlikely that this species belongs in
<taxonomicName id="8AFC4D5CFFF2B802874CCF2EF1A7F8BA" authorityName="Berlese" authorityYear="1892" box="[274,475,1871,1897]" class="Arachnida" family="Heterozerconidae" kingdom="Animalia" order="Mesostigmata" pageId="60" pageNumber="61" phylum="Arthropoda" rank="family">Heterozerconidae</taxonomicName>
. A more likely option would be in Trigynaspida, possibly
<taxonomicName id="8AFC4D5CFFF2B802823DCF2EF56EF8BA" baseAuthorityName="Klompen &amp; Austin" baseAuthorityYear="2007" box="[1123,1298,1871,1897]" class="Arachnida" family="Paramegistidae" kingdom="Animalia" order="Mesostigmata" pageId="60" pageNumber="61" phylum="Arthropoda" rank="family">Paramegistidae</taxonomicName>
.
</paragraph>
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</treatment>
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