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<document id="29A5AEA3884930EEE3C6CB16C1951950" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626381872300" checkinUser="felipe" docAuthor="Oláh, János, Johanson, Kjell Arne &amp; Barnard, Peter C." docDate="2008" docId="627D87E1FF7FF798FF7EFC41FBE07CAA" docLanguage="en" docName="zt01738p171.pdf" docOrigin="Zootaxa 1738" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Cheumatopsyche akisena Olah &amp; Johanson 2008, new species" docType="treatment" docVersion="2" lastPageNumber="131" masterDocId="9E44FF99FFFEF71AFFE9FF9BFFD07D08" masterDocTitle="Revision of the Oriental and Afrotropical species of Cheumatopsyche Wallengren (Hydropsychidae, Trichoptera)" masterLastPageNumber="171" masterPageNumber="1" pageNumber="130" updateTime="1699011095215" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="1451C9224706B458321F8A773A6CAA59" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="73BBB12A12FEEDE4884189DBE974B184">
<mods:title id="30673ED0F21EECEFD1CEF90CF6AC225A">Revision of the Oriental and Afrotropical species of Cheumatopsyche Wallengren (Hydropsychidae, Trichoptera)</mods:title>
</mods:titleInfo>
<mods:name id="CC2CC60A22A8F0213B4716B03A42F992" type="personal">
<mods:role id="99E0940FBC4CBAFCCB379B77B3261E0A">
<mods:roleTerm id="C29C9B1B5EA1CB7BDD4C8D400723BB90">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0A1C20556281DE4CBF93881D01A79D27">Oláh, János</mods:namePart>
</mods:name>
<mods:name id="2B0035DA5D43B080CD2AA55EB3C050C1" type="personal">
<mods:role id="330B493405C435BBA5294AF2ECFE6A9A">
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</mods:role>
<mods:namePart id="D199D0A0CB19C03775E45458BCA022EC">Johanson, Kjell Arne</mods:namePart>
</mods:name>
<mods:name id="94D596AA417848A36E5E878997E48139" type="personal">
<mods:role id="6ACA89E0873E1F03FFE914E02D630C45">
<mods:roleTerm id="2A2C79A99E3811057B4A9019451866CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C888661D133D723F3D19C6E88C4D28ED">Barnard, Peter C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="E82481930C5E4CEAE584DAC6DE3C836F">text</mods:typeOfResource>
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<mods:title id="64A75A94D5999BEDB9019D1590F04295">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D137F67004DE948CBF9B1237E9C36A24">
<mods:date id="D28CBAAE0C329179DF38943A33F0BE8E">2008</mods:date>
<mods:detail id="417A2B3B136EB2F4F3BDFFD763125AED" type="pubDate">
<mods:number id="6B94DC26AA09FA97B74F076573CD3A68">2008-03-31</mods:number>
</mods:detail>
<mods:detail id="D5300B2366CFAFA8C1A317EECBA1E796" type="volume">
<mods:number id="15FE639615198B08C8087EBC3740B0B1">1738</mods:number>
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<mods:start id="BB96C757647000327B0401BF79A88BF6">1</mods:start>
<mods:end id="AA1863D2E30FCE5A774EC74E6AE053B3">171</mods:end>
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<mods:classification id="50751D967E8E318630F9BE208062CEDD">journal article</mods:classification>
<mods:identifier id="31BF5F9BD5453F7F92B3B42A9974147A" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="627D87E1FF7FF798FF7EFC41FBE07CAA" LSID="urn:lsid:plazi:treatment:627D87E1FF7FF798FF7EFC41FBE07CAA" httpUri="http://treatment.plazi.org/id/627D87E1FF7FF798FF7EFC41FBE07CAA" lastPageId="130" lastPageNumber="131" pageId="129" pageNumber="130">
<subSubSection id="A2CE657CFF7FF79BFF7EFC41FC9B7EFC" box="[151,843,986,1012]" pageId="129" pageNumber="130" type="nomenclature">
<paragraph id="EA6B36F7FF7FF79BFF7EFC41FC9B7EFC" blockId="129.[151,843,986,1052]" box="[151,843,986,1012]" pageId="129" pageNumber="130">
<heading id="B123819BFF7FF79BFF7EFC41FC9B7EFC" bold="true" box="[151,843,986,1012]" fontSize="11" level="1" pageId="129" pageNumber="130" reason="1">
<emphasis id="D8A0EAE5FF7FF79BFF7EFC41FC9B7EFC" bold="true" box="[151,843,986,1012]" pageId="129" pageNumber="130">
<taxonomicName id="2DD44D74FF7FF79BFF7EFC41FD7F7EFC" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[151,687,986,1012]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="akisena" status="sp. nov.">
<emphasis id="D8A0EAE5FF7FF79BFF7EFC41FE1A7EFC" bold="true" box="[151,458,986,1012]" italics="true" pageId="129" pageNumber="130">Cheumatopsyche akisena</emphasis>
Oláh &amp; Johanson
</taxonomicName>
,
<taxonomicNameLabel id="C393579EFF7FF79BFD52FC41FC9B7EFC" box="[699,843,986,1012]" pageId="129" pageNumber="130" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFF7FF798FF7EFB99FE227C0A" lastPageId="130" lastPageNumber="131" pageId="129" pageNumber="130" type="description">
<paragraph id="EA6B36F7FF7FF79BFF7EFB99FEE37914" blockId="129.[151,843,986,1052]" box="[151,307,1026,1052]" pageId="129" pageNumber="130">
<figureCitation id="72EF2A72FF7FF79BFF7EFB99FEE37914" box="[151,307,1026,1052]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 313–316</figureCitation>
</paragraph>
<paragraph id="EA6B36F7FF7FF79BFF7EFBC9FB4179B4" blockId="129.[151,1436,1106,2012]" pageId="129" pageNumber="130">
This species is similar to
<taxonomicName id="2DD44D74FF7FF79BFE54FBC8FD1D7964" authority="Ulmer" authorityName="Ulmer" box="[445,717,1106,1132]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="D8A0EAE5FF7FF79BFE54FBC8FDAB7964" box="[445,635,1107,1132]" italics="true" pageId="129" pageNumber="130">C. maculipennis</emphasis>
Ulmer
</taxonomicName>
from Borneo, but
<taxonomicName id="2DD44D74FF7FF79BFC4AFBC8FBCE7964" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[931,1054,1107,1132]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="akisena">
<emphasis id="D8A0EAE5FF7FF79BFC4AFBC8FBCE7964" box="[931,1054,1107,1132]" italics="true" pageId="129" pageNumber="130">C. akisena</emphasis>
</taxonomicName>
is easily separated from
<taxonomicName id="2DD44D74FF7FF79BFAA9FBC9FED2799C" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="maculipennis">C. maculipennis</taxonomicName>
by the strongly tapering apex of segment X. In addition, the harpagones of
<taxonomicName id="2DD44D74FF7FF79BFB6DFBE0FAD0799C" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[1156,1280,1147,1172]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="akisena">
<emphasis id="D8A0EAE5FF7FF79BFB6DFBE0FAD0799C" box="[1156,1280,1147,1172]" italics="true" pageId="129" pageNumber="130">C. akisena</emphasis>
</taxonomicName>
have a deep, subapical excision on the dorsal margin, partly encircled by a small digitiform process.
</paragraph>
<paragraph id="EA6B36F7FF7FF79BFF2FFB51FF2A788C" blockId="129.[151,1436,1106,2012]" pageId="129" pageNumber="130">
Male. Body and wings light brown with brown pubescence; wings with no spots. Maxillary palp segment I shortest, segment II longer, segments III and IV equally long, segment V as long as sum of segments I–IV. Head dorsum brown, with 5 lighter warts. Proepisternum without swollen setal wart. Protarsal claws asymmetrical, laterally flanked by setal bundle. Forewing length 6.0 mm, hind wing length
<quantity id="2D2C9B12FF7FF79BFB68FAD9FB0E7854" box="[1153,1246,1346,1372]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="129" pageNumber="130" unit="mm" value="4.6">4.6 mm</quantity>
. Hind wing fork I absent.
</paragraph>
<paragraph id="EA6B36F7FF7FF79BFF2EFA09FA4C7AD4" blockId="129.[151,1436,1106,2012]" pageId="129" pageNumber="130">
Male genitalia. Segment IX fused annularly; sternum about 2 times longer than tergum (
<figureCitation id="72EF2A72FF7FF79BFB25FA09FAE378A4" box="[1228,1331,1426,1452]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 313</figureCitation>
); ventrocaudal spiny lobes absent; anterior margins of segment IX straight; apical lobes on posterolateral margins right-angled, triangular, bordering wide and deep articulation cavity of coxopodites (
<figureCitation id="72EF2A72FF7FF79BFB6AFA79FB3B78F4" box="[1155,1259,1506,1532]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 313</figureCitation>
); spine row on posterior margins of segment IX interrupted; spines on apicodorsal spiny lobes stouter and 2 times longer than other spines. Intersegmental step between segment IX and segment X filled by elevated dorsum of segment X. Segment X long, with strongly dorsad, hook-shaped apex (
<figureCitation id="72EF2A72FF7FF79BFCB5F9C1FC137B7C" box="[860,963,1626,1652]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 313</figureCitation>
) formed by apicoventral setal lobes; in dorsal view (
<figureCitation id="72EF2A72FF7FF79BFEC7F919FE427B94" box="[302,402,1666,1692]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 314</figureCitation>
) apex strongly tapering. Setaless mesocaudal lobe reduced. Lateral setose areas forming pair of elevated warts located centrally on segment X, clearly visible in dorsal and lateral view (
<figureCitation id="72EF2A72FF7FF79BFAC6F931FF187BE4" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 313, 314</figureCitation>
). Dorsal and lateral interlobular gaps absent. Transverse sutures very broad, running obliquely above lateral setose areas (
<figureCitation id="72EF2A72FF7FF79BFE83F961FE037A1C" box="[362,467,1786,1812]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 313</figureCitation>
). Narrow, sclerotised longitudinal sutures forming continuation of dorsal margins of ventroapical setal lobes (
<figureCitation id="72EF2A72FF7FF79BFE0EF8B9FD817A34" box="[487,593,1826,1852]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 313</figureCitation>
); meeting transverse sutures at posterior margin of segment IX. Coxopodites narrowing immediately after bases (
<figureCitation id="72EF2A72FF7FF79BFD40F8D1FC9C7A6C" box="[681,844,1866,1892]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 313, 314</figureCitation>
), before strongly widening apically in lateral view (
<figureCitation id="72EF2A72FF7FF79BFF76F8E9FEDB7A84" box="[159,267,1906,1932]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 313</figureCitation>
), slightly widening apically in ventral view (
<figureCitation id="72EF2A72FF7FF79BFCD2F8E9FC777A84" box="[827,935,1906,1932]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 314</figureCitation>
); slightly exceeding apex of segment X. Harpagones basally broad before narrowing at mid-length; apices curving dorsad, with posterior incision (
<figureCitation id="72EF2A72FF7FF79BFA84F801FF177AD4" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 313</figureCitation>
); in ventral view nearly straight, with parallel-lateral margins (
<figureCitation id="72EF2A72FF7FF79BFC59F859FBC97AD4" box="[944,1049,1986,2012]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="129" pageNumber="130">Fig. 314</figureCitation>
). Phallotheca narrowing towards
</paragraph>
<paragraph id="EA6B36F7FF7CF798FF7EFF03FE227C0A" blockId="130.[151,1436,152,418]" pageId="130" pageNumber="131">
distal two-thirds (
<figureCitation id="72EF2A72FF7CF798FE80FF03FE1E7DBA" box="[361,462,152,178]" captionStart="FIGURES 313–316" captionStartId="130.[151,269,1946,1970]" captionTargetBox="[166,1415,487,1898]" captionTargetId="figure-104@130.[151,1436,474,1922]" captionTargetPageId="130" captionText="FIGURES 313–316. Cheumatopsyche akisena Oláh &amp; Johanson, new species, holotype. 309 — genitalia, lateral; 310 — genitalia, dorsal; 311 — gonocoxite, ventral; 312 — phallus, lateral." pageId="130" pageNumber="131">Fig. 316</figureCitation>
) before broadening with ventral keel from distal one-third; sclerotised endothecal process almost circular, somewhat elongated; phallotremal sclerites rounded; vestigial ventral endothecal membranous lobe not visible.
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFF7CF798FF2EFE94FE497C5A" pageId="130" pageNumber="131" type="materials_examined">
<paragraph id="EA6B36F7FF7CF798FF2EFE94FE497C5A" blockId="130.[151,1436,152,418]" pageId="130" pageNumber="131">
<materialsCitation id="5ABC3CAAFF7CF798FF2EFE94FE497C5A" collectingDate="1988-04-05" collectionCode="NRM" collectorName="S. Adebratt" country="Malaysia" county="Borneo" location="Mendolong Nersery" municipality="Sipitang" pageId="130" pageNumber="131" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="D8A0EAE5FF7CF798FF2EFE94FD777C21" bold="true" box="[199,679,271,297]" pageId="130" pageNumber="131">
<typeStatus id="356F8855FF7CF798FF2EFE94FEE97C21" box="[199,313,271,297]" pageId="130" pageNumber="131" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="92C37667FF7CF798FE7BFE94FDE47C21" box="[402,564,271,297]" name="Malaysia" pageId="130" pageNumber="131">MALAYSIA</collectingCountry>
:
<collectingCounty id="030A4E7BFF7CF798FDABFE94FD727C21" box="[578,674,271,297]" pageId="130" pageNumber="131">Borneo</collectingCounty>
:
</emphasis>
<collectingRegion id="2810F815FF7CF798FD58FE8BFD2C7C22" box="[689,764,272,298]" country="Malaysia" name="Sabah" pageId="130" pageNumber="131">Sabah</collectingRegion>
,
<collectingMunicipality id="0A0FAC8DFF7CF798FCE3FE8BFCBE7C22" box="[778,878,272,298]" pageId="130" pageNumber="131">Sipitang</collectingMunicipality>
,
<location id="EF0B602CFF7CF798FC94FE8BFBBB7C22" LSID="urn:lsid:plazi:treatment:627D87E1FF7FF798FF7EFC41FBE07CAA:EF0B602CFF7CF798FC94FE8BFBBB7C22" box="[893,1131,272,298]" country="Malaysia" county="Borneo" municipality="Sipitang" name="Mendolong Nersery" pageId="130" pageNumber="131" stateProvince="Sabah">Mendolong Nersery</location>
,
<date id="9E6A1037FF7CF798FB90FE8BFB387C22" box="[1145,1256,272,298]" pageId="130" pageNumber="131" value="1988-04-05">
<collectingDate id="8E2EE9DFFF7CF798FB90FE8BFB387C22" box="[1145,1256,272,298]" pageId="130" pageNumber="131" value="1988-04-05">5.iv.1988</collectingDate>
</date>
, light [
<collectorName id="47215321FF7CF798FAD6FE8BFF007C5A" pageId="130" pageNumber="131">S. Adebratt</collectorName>
] (
<collectionCode id="8CC5AE32FF7CF798FF0CFEA3FEFC7C5A" box="[229,300,312,338]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="130" pageNumber="131" type="Museum">NRM</collectionCode>
, alcohol)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFF7CF798FF2EFEC4FD7C7C72" box="[199,684,351,378]" pageId="130" pageNumber="131" type="distribution">
<paragraph id="EA6B36F7FF7CF798FF2EFEC4FD7C7C72" blockId="130.[151,1436,152,418]" box="[199,684,351,378]" pageId="130" pageNumber="131">
<emphasis id="D8A0EAE5FF7CF798FF2EFEC4FEB97C71" bold="true" box="[199,361,351,377]" pageId="130" pageNumber="131">Distribution:</emphasis>
<collectingCountry id="92C37667FF7CF798FE98FEFBFE0C7C72" box="[369,476,352,378]" name="Malaysia" pageId="130" pageNumber="131">Malaysia</collectingCountry>
(Borneo:
<collectingRegion id="2810F815FF7CF798FDBAFEFBFD707C72" box="[595,672,352,378]" country="Malaysia" name="Sabah" pageId="130" pageNumber="131">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFF7CF798FF2EFE1CFBE07CAA" box="[199,1072,391,418]" pageId="130" pageNumber="131" type="etymology">
<paragraph id="EA6B36F7FF7CF798FF2EFE1CFBE07CAA" blockId="130.[151,1436,152,418]" box="[199,1072,391,418]" pageId="130" pageNumber="131">
<emphasis id="D8A0EAE5FF7CF798FF2EFE1CFE877CA9" bold="true" box="[199,343,391,417]" pageId="130" pageNumber="131">Etymology:</emphasis>
after the hook-shaped segment X, hook “akisena” in Sanscrit.
</paragraph>
</subSubSection>
</treatment>
</document>
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