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<document ID-DOI="http://dx.doi.org/10.3897/dez.70.97117" ID-Pensoft-Pub="1860-1324-1-69" ID-Pensoft-UUID="CA0CBE818BFA5DFA80DBD42521D5DBE3" ID-ZooBank="99BBA4C8ED2048879952B61CC25309D4" ModsDocID="1860-1324-70-1-69" checkinTime="1679031804748" checkinUser="pensoft" docAuthor="Tran, Anh Duc, Zettel, Herbert &amp; Sites, Robert W." docDate="2023" docId="62000292684555879D581EE06013402E" docLanguage="en" docName="DeutEntomolZeit 70(1): 69-111" docOrigin="Deutsche Entomologische Zeitschrift 70 (1)" docPubDate="2023-03-16" docSource="http://dx.doi.org/10.3897/dez.70.97117" docTitle="Eotrechus fuscus Basu, Chandra &amp; Venkatesan 2017" docType="treatment" docVersion="2" id="CA0CBE818BFA5DFA80DBD42521D5DBE3" lastPageNumber="69" masterDocId="CA0CBE818BFA5DFA80DBD42521D5DBE3" masterDocTitle="Revision of the genus Eotrechus Kirkaldy (Hemiptera, Heteroptera, Gerridae), with descriptions of six new species" masterLastPageNumber="111" masterPageNumber="69" pageNumber="69" updateTime="1679032168055" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of the genus Eotrechus Kirkaldy (Hemiptera, Heteroptera, Gerridae), with descriptions of six new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tran, Anh Duc</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9605-0739</mods:nameIdentifier>
<mods:affiliation>Faculty of Biology, University of Science, Vietnam National University, Hanoi, 334 Nguyen Trai, Thanh Xuan, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier type="email">tran.anhduc@hus.edu.vn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zettel, Herbert</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7760-0472</mods:nameIdentifier>
<mods:affiliation>Herbert Zettel, 2 nd Zoological Department (Entomology), Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sites, Robert W.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3895-813X</mods:nameIdentifier>
<mods:affiliation>Enns Entomology Museum, University of Missouri, Columbia, MO 65211, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>70</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>69</mods:start>
<mods:end>111</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.70.97117</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.70.97117</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-1-69</mods:identifier>
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</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:62000292684555879D581EE06013402E" httpUri="http://treatment.plazi.org/id/62000292684555879D581EE06013402E" lastPageNumber="69" pageId="0" pageNumber="69">
<subSubSection pageId="0" pageNumber="69" type="nomenclature">
<paragraph pageId="0" pageNumber="69">
<taxonomicName ID-CoL="6FM26" LSID="62000292-6845-5587-9D58-1EE06013402E" authority="Basu, Chandra &amp; Venkatesan, 2017" authorityName="Basu, Chandra &amp; Venkatesan" authorityYear="2017" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus fuscus" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="fuscus">Eotrechus fuscus Basu, Chandra &amp; Venkatesan, 2017</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="description">
<paragraph pageId="0" pageNumber="69">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eotrechus fuscus. A. Pygophore, lateral view; B. Proctiger, dorsal view; C. paramere, lateral view; D, E. Posterior part of abdomen, female, dorsal and lateral views (A, B. Same scale; D, E. Same scale)." figureDoi="10.3897/dez.70.97117.figure2" httpUri="https://binary.pensoft.net/fig/826091" pageId="0" pageNumber="69">Figs 2</figureCitation>
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Distribution records of species of Eotrechus. Squares represent E. kalidasa species group; circles represent E. pingae species group; triangles represent species unassigned to a group." figureDoi="10.3897/dez.70.97117.figure25" httpUri="https://binary.pensoft.net/fig/826114" pageId="0" pageNumber="69">, 25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="reference_group">
<paragraph pageId="0" pageNumber="69">
<taxonomicName authorityName="Basu, Chandra &amp; Venkatesan" authorityYear="2017" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus fuscus" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="fuscus">Eotrechus fuscus</taxonomicName>
Basu, Chandra &amp; Venkatesan, 2017: 392-397, figs 1-12 (type locality: Sikkim, India).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="materials_examined">
<paragraph pageId="0" pageNumber="69">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="69">
<materialsCitation collectingDate="2003-05-05" collectorName="Div., Alaungdaw Katthapa NP, Pagoda" country="Myanmar" elevation="350" latitude="22.31855" location="Sagaing" longLatPrecision="1" longitude="94.4753" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Sagaing">
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
•
<specimenCount type="male">1 ♂</specimenCount>
,
<specimenCount type="female">3 ♀♀</specimenCount>
(apterous);
<collectingRegion country="Myanmar" name="Sagaing">Sagaing</collectingRegion>
<collectorName>Div.</collectorName>
,
<collectorName>Alaungdaw Katthapa NP</collectorName>
,
<collectorName>Pagoda</collectorName>
stream;
<geoCoordinate degrees="22" direction="north" minutes="19.113" orientation="latitude" precision="1" value="22.31855">22°19.113'N</geoCoordinate>
,
<geoCoordinate degrees="94" direction="east" minutes="28.518" orientation="longitude" precision="1" value="94.4753">94°28.518'E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2003-05-05">5 May 2003</collectingDate>
; Boukal et al. leg.; (105); NHMW
</materialsCitation>
•
<materialsCitation collectingDate="2003-05-06" collectorName="Div., Alaungdaw Katthapa NP, Pagoda" country="Myanmar" elevation="350" latitude="22.318233" location="Sagaing" longLatPrecision="1" longitude="94.480385" specimenCount="1" specimenCount-male="1" stateProvince="Sagaing">
<specimenCount type="male">1 ♂</specimenCount>
(macropterous);
<collectingRegion country="Myanmar" name="Sagaing">Sagaing</collectingRegion>
<collectorName>Div.</collectorName>
,
<collectorName>Alaungdaw Katthapa NP</collectorName>
,
<collectorName>Pagoda</collectorName>
stream;
<geoCoordinate degrees="22" direction="north" minutes="19.094" orientation="latitude" precision="1" value="22.318233">22°19.094'N</geoCoordinate>
,
<geoCoordinate degrees="94" direction="east" minutes="28.823" orientation="longitude" precision="1" value="94.480385">94°28.823'E</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</quantity>
a.s.l.
</elevation>
;
<collectingDate value="2003-05-06">6 May 2003</collectingDate>
; Boukal et al. leg.; (110); NHMW
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="diagnosis">
<paragraph pageId="0" pageNumber="69">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="69">
Size: apterous male, length 10.30, width 2.64 (apterous); macropterous male, length 11.60, width 2.78; apterous females, length 10.80-11.70, width 2.72-2.82. Antenna: segments I and II, each distinctly longer than segments III, IV; segment III shortest, ca. 0.5-0.6
<normalizedToken originalValue="×">x</normalizedToken>
length of each of segments I, II, and ca. 0.9
<normalizedToken originalValue="×">x</normalizedToken>
length of segment IV. Mesosternum ca. 1.6-1.7
<normalizedToken originalValue="×">x</normalizedToken>
length of metasternum (male: 2.07: 1.19; female: 2.07: 1.30). Fore femur slender in both sexes. In apterous morph, posterior three quarters of mesonotum raised into a hump. Male: abdominal sterna III-VII with longitudinal median groove; sternum VII ca. 0.8
<normalizedToken originalValue="×">x</normalizedToken>
length of two preceding sterna combined, posterior margin median notch ca. 0.25
<normalizedToken originalValue="×">x</normalizedToken>
length of sternum VII. Male genitalia: pygophore with two long, relatively slender posterolateral projections (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eotrechus fuscus. A. Pygophore, lateral view; B. Proctiger, dorsal view; C. paramere, lateral view; D, E. Posterior part of abdomen, female, dorsal and lateral views (A, B. Same scale; D, E. Same scale)." figureDoi="10.3897/dez.70.97117.figure2" httpUri="https://binary.pensoft.net/fig/826091" pageId="0" pageNumber="69">2A</figureCitation>
); paramere with proximal half wider, twice as wide as distal half, distal half stout, slightly twisted, tapering to rounded apex (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eotrechus fuscus. A. Pygophore, lateral view; B. Proctiger, dorsal view; C. paramere, lateral view; D, E. Posterior part of abdomen, female, dorsal and lateral views (A, B. Same scale; D, E. Same scale)." figureDoi="10.3897/dez.70.97117.figure2" httpUri="https://binary.pensoft.net/fig/826091" pageId="0" pageNumber="69">2C</figureCitation>
); proctiger posterolaterally produced into rounded processes, posterior margin broadly rounded (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eotrechus fuscus. A. Pygophore, lateral view; B. Proctiger, dorsal view; C. paramere, lateral view; D, E. Posterior part of abdomen, female, dorsal and lateral views (A, B. Same scale; D, E. Same scale)." figureDoi="10.3897/dez.70.97117.figure2" httpUri="https://binary.pensoft.net/fig/826091" pageId="0" pageNumber="69">2B</figureCitation>
). Female: abdomen long and slender; genitalia visible in lateral view, proctiger slightly longer than wide, directed ventrad (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eotrechus fuscus. A. Pygophore, lateral view; B. Proctiger, dorsal view; C. paramere, lateral view; D, E. Posterior part of abdomen, female, dorsal and lateral views (A, B. Same scale; D, E. Same scale)." figureDoi="10.3897/dez.70.97117.figure2" httpUri="https://binary.pensoft.net/fig/826091" pageId="0" pageNumber="69">2D, E</figureCitation>
).
</paragraph>
<caption doi="10.3897/dez.70.97117.figure2" httpUri="https://binary.pensoft.net/fig/826091" pageId="0" pageNumber="69" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" pageId="0" pageNumber="69">Figure 2.</emphasis>
<taxonomicName authorityName="Basu, Chandra &amp; Venkatesan" authorityYear="2017" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus fuscus" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis italics="true" pageId="0" pageNumber="69">Eotrechus fuscus</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="69">A.</emphasis>
Pygophore, lateral view;
<emphasis bold="true" pageId="0" pageNumber="69">B.</emphasis>
Proctiger, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="69">C.</emphasis>
paramere, lateral view;
<emphasis bold="true" pageId="0" pageNumber="69">D, E.</emphasis>
Posterior part of abdomen, female, dorsal and lateral views (
<emphasis bold="true" pageId="0" pageNumber="69">A, B.</emphasis>
Same scale;
<emphasis bold="true" pageId="0" pageNumber="69">D, E.</emphasis>
Same scale).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="description">
<paragraph pageId="0" pageNumber="69">Supplemental description of apterous female.</paragraph>
<paragraph pageId="0" pageNumber="69">
Head width across eyes 1.99; interocular width 0.80; eye kidney-shaped in dorsal view, length 0.85. Antennae subequal to body length (12.14: 11.40), lengths of segments I-IV: 3.67: 3.38: 2.17: 2.92; segment I without black spines. Pronotum shorter than head length (1.40: 1.89). Posterior three quarters of mesonotum strongly swollen, forming a hump. Lengths of mesosternum and metasternum: 2.07 and 1.30. Lengths of leg segments (femur: tibia: tarsal segment I: tarsal segment II): fore leg: 4.75: 3.50: 0.62: 0.71; middle leg: 9.08: 8.83: 0.60: 0.91; hind leg: 10.0: 11.9: 0.68: 0.93. Fore trochanter with two or three long, fine setae on ventral side. Fore femur similar to males, length ca. 11.9
<normalizedToken originalValue="×">x</normalizedToken>
maximum width (4.75: 0.40). Fore tibia similar to males. Middle and hind legs similar to males: middle and hind femur with short spines scattered along ventral surface, middle and hind tibiae with scattered long, stout setae. Claws stout, lengths of fore, mid- and hind claws: 0.32: 0.42: 0.41. Total length of abdominal sterna II-VII ca. 0.4
<normalizedToken originalValue="×">x</normalizedToken>
body length (4.87: 11.4). Sternum VII ca. 0.9
<normalizedToken originalValue="×">x</normalizedToken>
length of two preceding sterna combined (1.36: 1.53), posterior margin straight. Connexival corners of sternum VII pointed and slightly curved mesad (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eotrechus fuscus. A. Pygophore, lateral view; B. Proctiger, dorsal view; C. paramere, lateral view; D, E. Posterior part of abdomen, female, dorsal and lateral views (A, B. Same scale; D, E. Same scale)." figureDoi="10.3897/dez.70.97117.figure2" httpUri="https://binary.pensoft.net/fig/826091" pageId="0" pageNumber="69">2D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="description">
<paragraph pageId="0" pageNumber="69">Supplemental description of macropterous male.</paragraph>
<paragraph pageId="0" pageNumber="69">
Head chiefly black, with a transverse yellow mark on posterior margin; pronotum chiefly black, with yellow stripe on anterior part of midline, not extending onto pronotal lobe; wings chiefly black, membrane brown. Pronotum length (including pronotal lobe) 3.66, humeral width 2.28. Fore wing length 7.25, wing veins similar to
<taxonomicName lsidName="E. kalidasa" pageId="0" pageNumber="69" rank="species" species="kalidasa">
<emphasis italics="true" pageId="0" pageNumber="69">E. kalidasa</emphasis>
</taxonomicName>
(see
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.15602" author="Matsuda, R" journalOrPublisher="The University of Kansas Science Bulletin" pageId="0" pageNumber="69" pagination="25 - 632" refId="B8" refString="Matsuda, R, 1960. Morphology, evolution and a classification of the Gerridae (Hemiptera - Heteroptera). The University of Kansas Science Bulletin 41 (2): 25 - 632, DOI: https://doi.org/10.5962/bhl.part.15602" title="Morphology, evolution and a classification of the Gerridae (Hemiptera - Heteroptera)." url="https://doi.org/10.5962/bhl.part.15602" volume="41" year="1960">Matsuda 1960</bibRefCitation>
: fig. 553;
<bibRefCitation author="Andersen, NM" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="69" pagination="1 - 25" refId="B2" refString="Andersen, NM, 1982. Semiterrestrial water striders of the genera Eotrechus Kirkaldy and Chimarrhometra Bianchi (Insecta, Hemiptera, Gerridae). Steenstrupia 9: 1 - 25" title="Semiterrestrial water striders of the genera Eotrechus Kirkaldy and Chimarrhometra Bianchi (Insecta, Hemiptera, Gerridae)." volume="9" year="1982">Andersen 1982</bibRefCitation>
: fig. 8). Other characteristics similar to apterous male.
</paragraph>
<paragraph pageId="0" pageNumber="69">Macropterous female: Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="remarks">
<paragraph pageId="0" pageNumber="69">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="69">
The original description by
<bibRefCitation DOI="https://doi.org/10.1515/aemnp-2017-0083" author="Basu, S" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="69" pagination="391 - 398" refId="B4" refString="Basu, S, Chandra, K, Venkatesan, T, 2017. Eotrechus fuscus sp. nov. from Northeastern India with a key to Indian species (Hemiptera: Heteroptera: Gerridae). Acta Entomologica Musei Nationalis Pragae 57 (2): 391 - 398, DOI: https://doi.org/10.1515/aemnp-2017-0083" title="Eotrechus fuscus sp. nov. from Northeastern India with a key to Indian species (Hemiptera: Heteroptera: Gerridae)." url="https://doi.org/10.1515/aemnp-2017-0083" volume="57" year="2017">Basu et al. (2017)</bibRefCitation>
was based on only a single apterous male. Thus, supplementary descriptions of apterous female and macropterous male are provided as above.
</paragraph>
<paragraph pageId="0" pageNumber="69">
<taxonomicName authorityName="Basu, Chandra &amp; Venkatesan" authorityYear="2017" class="Insecta" family="Gerridae" genus="Eotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eotrechus fuscus" order="Hemiptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis italics="true" pageId="0" pageNumber="69">Eotrechus fuscus</emphasis>
</taxonomicName>
can be confused with
<taxonomicName lsidName="E. kalidasa" pageId="0" pageNumber="69" rank="species" species="kalidasa">
<emphasis italics="true" pageId="0" pageNumber="69">E. kalidasa</emphasis>
</taxonomicName>
, and the two species have greatly overlapping distributions. The combination of differences in the relative lengths of antennal segments, the relative lengths of fore tarsal segments, and the shape of the parameres help to distinguish
<taxonomicName lsidName="E. fuscus" pageId="0" pageNumber="69" rank="species" species="fuscus">
<emphasis italics="true" pageId="0" pageNumber="69">E. fuscus</emphasis>
</taxonomicName>
from
<taxonomicName lsidName="E. kalidasa" pageId="0" pageNumber="69" rank="species" species="kalidasa">
<emphasis italics="true" pageId="0" pageNumber="69">E. kalidasa</emphasis>
</taxonomicName>
, and also from
<taxonomicName lsidName="E. steineri" pageId="0" pageNumber="69" rank="species" species="steineri">
<emphasis italics="true" pageId="0" pageNumber="69">E. steineri</emphasis>
</taxonomicName>
sp. nov. (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparative morphology of E. kalidasa, E. fuscus, and E. steineri sp. nov." httpUri="http://table.plazi.org/id/EF594DCD37796B0AEC53DFC488CF38A9" pageId="0" pageNumber="69" tableUuid="EF594DCD37796B0AEC53DFC488CF38A9">1</tableCitation>
).
</paragraph>
<caption ID-Table-UUID="EF594DCD37796B0AEC53DFC488CF38A9" httpUri="http://table.plazi.org/id/EF594DCD37796B0AEC53DFC488CF38A9" pageId="0" pageNumber="69" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="69">
<emphasis bold="true" pageId="0" pageNumber="69">Table 1.</emphasis>
Comparative morphology of
<taxonomicName lsidName="E. kalidasa" pageId="0" pageNumber="69" rank="species" species="kalidasa">
<emphasis italics="true" pageId="0" pageNumber="69">E. kalidasa</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="E. fuscus" pageId="0" pageNumber="69" rank="species" species="fuscus">
<emphasis italics="true" pageId="0" pageNumber="69">E. fuscus</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="E. steineri" pageId="0" pageNumber="69" rank="species" species="steineri">
<emphasis italics="true" pageId="0" pageNumber="69">E. steineri</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="69">
<table pageId="0" pageNumber="69">
<tr pageId="0" pageNumber="69">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="69" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="69" rowspan="1">
<taxonomicName lsidName="E. kalidasa" pageId="0" pageNumber="69" rank="species" species="kalidasa">
<emphasis italics="true" pageId="0" pageNumber="69">E. kalidasa</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="69" rowspan="1">
<taxonomicName lsidName="E. fuscus" pageId="0" pageNumber="69" rank="species" species="fuscus">
<emphasis italics="true" pageId="0" pageNumber="69">E. fuscus</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="69" rowspan="1">
<taxonomicName lsidName="E. steineri" pageId="0" pageNumber="69" rank="species" species="steineri">
<emphasis italics="true" pageId="0" pageNumber="69">E. steineri</emphasis>
</taxonomicName>
sp. nov.
</th>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="3">Antennal segments</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- all segments subequal in length</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- segments I and II, each distinctly longer than segments III and IV</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- segments I, II, and IV subequal in length</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- segments I and IV slightly longer than others</td>
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- segment III shortest, 0.5-0.6
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length of segments I, II, and ca. 0.9
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length of segment IV
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- segment III shortest, only 0.85-0.90
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length of each of other segments
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<td colspan="1" pageId="0" pageNumber="69" rowspan="1">segment I slightly longer than segment II (1.17-1.30: 1.00)</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">segment I shorter than segment II (ca. 0.7: 1.00)</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">segment I slightly longer than segment II (1.10-1.25: 1.00)</td>
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<td colspan="1" pageId="0" pageNumber="69" rowspan="1">Mesonotum of apterous form</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">transversely depressed in the middle</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">posterior three quarters with a hump</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">posterior two-thirds with a large hump</td>
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<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">Abdominal sterna of males</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">with narrow median groove from sterna III-VII</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">with narrow median groove from sterna III-VII</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">sterna II-VII weakly depressed medially, not forming distinct groove</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="3">Paramere</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- proximal third only slightly wider than distal part, convex dorsad</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- proximal half twice as wide as distal half</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- proximal third twice as wide as distal part</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- distal part slender, straight</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- distal half stout, slightly twisted</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- distal two-thirds slender, directed slightly dorsad</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- apex narrow</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- apex rounded</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">- apex rounded, slightly broadened</td>
</tr>
<tr pageId="0" pageNumber="69">
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">Abdominal apex of females</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">proctiger completely concealed, not visible</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">proctiger visible</td>
<td colspan="1" pageId="0" pageNumber="69" rowspan="1">proctiger almost completed concealed</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="69" type="distribution">
<paragraph pageId="0" pageNumber="69">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="69">
India: Sikkim (
<bibRefCitation DOI="https://doi.org/10.1515/aemnp-2017-0083" author="Basu, S" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="69" pagination="391 - 398" refId="B4" refString="Basu, S, Chandra, K, Venkatesan, T, 2017. Eotrechus fuscus sp. nov. from Northeastern India with a key to Indian species (Hemiptera: Heteroptera: Gerridae). Acta Entomologica Musei Nationalis Pragae 57 (2): 391 - 398, DOI: https://doi.org/10.1515/aemnp-2017-0083" title="Eotrechus fuscus sp. nov. from Northeastern India with a key to Indian species (Hemiptera: Heteroptera: Gerridae)." url="https://doi.org/10.1515/aemnp-2017-0083" volume="57" year="2017">Basu et al. 2017</bibRefCitation>
); Myanmar (new record): Sagaing Region (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Distribution records of species of Eotrechus. Squares represent E. kalidasa species group; circles represent E. pingae species group; triangles represent species unassigned to a group." figureDoi="10.3897/dez.70.97117.figure25" httpUri="https://binary.pensoft.net/fig/826114" pageId="0" pageNumber="69">25</figureCitation>
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<paragraph id="F11AE03D7A6FFE142264D56139ECE227" pageId="164" pageNumber="165">
<taxonomicName id="620F2DCF90D5660144A5D554DA9E92AA" ID-CoL="49M6M" authority="Genier, 1996" authorityName="Genier" authorityYear="1996" class="Insecta" family="Scarabaeidae" genus="Ontherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ontherus (Caelontherus) howdeni" order="Coleoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="howdeni" subGenus="Caelontherus">
Ontherus (Caelontherus) howdeni
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, 1996
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Plate 36B
</paragraph>
</subSubSection>
<subSubSection id="1C82E3FEC66D05969B4EDD4F8B742D06" pageId="164" pageNumber="165" type="reference_group">
<paragraph id="7B6E7A1D9FE4E74B2FAE4E0925D22954" pageId="164" pageNumber="165">
<taxonomicName id="76E0BD28212113DAAEA34038C0CA17E2" class="Insecta" family="Scarabaeidae" genus="Ontherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ontherus (Caelontherus) howdeni" order="Coleoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="howdeni" subGenus="Caelontherus">Ontherus (Caelontherus) howdeni</taxonomicName>
<normalizedToken id="2B8EC3A24B6D600DEB1210367833E4D3" originalValue="Génier">Genier</normalizedToken>
, 1996: 47 (original description. Type locality: Chanchamayo, Peru).
</paragraph>
<paragraph id="8A0C76741A984EBEAFA069F4D3700386" pageId="164" pageNumber="165">
<taxonomicName id="741D04121517DB7F05DB9AA4AD7268CB" class="Insecta" family="Scarabaeidae" genus="Ontherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ontherus howdeni" order="Coleoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="howdeni">Ontherus howdeni</taxonomicName>
:
<bibRefCitation id="6E299E5C37BC2C0BFF97D3DD4EFD6EE2" pageId="164" pageNumber="165">Krajcik 2012</bibRefCitation>
: 174 (complete list of species);
<bibRefCitation id="CFDEEFBE7BE903EBDD4C36BDEDB87AC2" author="Ratcliffe, BC" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="276" pageNumber="277" pagination="186 - 207" title="Beetles (Coleoptera) of Peru: A Survey of the Families." url="https://doi.org/10.2317/kent-88-02-186-207.1" volume="88" year="2015">Ratcliffe et al. 2015</bibRefCitation>
: 197 (cited for Peru).
</paragraph>
<paragraph id="D60F108C83569B14A54D44A7FD948B00" pageId="164" pageNumber="165">
<taxonomicName id="C4F9391E848750D3D27AEFF985AF4D73" class="Insecta" family="Scarabaeidae" genus="Ontherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ontherus (Caelontherus) howdeni" order="Coleoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="howdeni" subGenus="Caelontherus">Ontherus (Caelontherus) howdeni</taxonomicName>
:
<bibRefCitation id="292AEB37B104F9D08B8A26A541DADE15" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
: 318-319 (cited for Ecuador);
<bibRefCitation id="014299FDE69C18A0AAE000E9D048FBCB" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
: 96 (cited for Ecuador).
</paragraph>
</subSubSection>
<subSubSection id="A6EF6E4B1EB5CD02038A9BDA66B42AB6" pageId="164" pageNumber="165" type="type specimens">
<paragraph id="23D4CCB6812FE38350FD9528FA99494B" pageId="164" pageNumber="165">Type specimens.</paragraph>
<paragraph id="C79234FF6715DA4E134DB51A351E9193" pageId="164" pageNumber="165">
<taxonomicName id="F4B5A42A55A34F235D2021BE35CB8FDF" class="Insecta" family="Scarabaeidae" genus="Ontherus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ontherus (Caelontherus) howdeni" order="Coleoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="howdeni" subGenus="Caelontherus">Ontherus (Caelontherus) howdeni</taxonomicName>
<normalizedToken id="723E2EE245BC4EF0D7656CB5A654B13A" originalValue="Génier">Genier</normalizedToken>
, 1996. The holotype (♂) is deposited at the CMNC (see
<bibRefCitation id="488A345F916F490562B5B8A802422B36" author="Genier, F" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="267" pageNumber="268" pagination="1 - 169" title="A revision of the neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="170" year="1996">
<normalizedToken id="11F4D48DF260DF2741E00887A47D56B9" originalValue="Génier">Genier</normalizedToken>
1996
</bibRefCitation>
: 48). Locality: Chanchamayo, Peru, not examained.
</paragraph>
</subSubSection>
<subSubSection id="860811A28303B687D46A0260C381004B" pageId="164" pageNumber="165" type="distribution">
<paragraph id="7B45779F225C4CE8226D447DD3AD99AB" pageId="164" pageNumber="165">Distribution.</paragraph>
<paragraph id="CEFA4254BDFDB743603878833B69FA24" pageId="164" pageNumber="165">Ecuador and Peru.</paragraph>
</subSubSection>
<subSubSection id="B6101BEE29CA255D616D897C5A387BAF" pageId="164" pageNumber="165" type="records examined">
<paragraph id="788BD9EC7D4BD1EFCA4550B11EE614F5" pageId="164" pageNumber="165">Records examined.</paragraph>
<paragraph id="34B7B2BD217CF4AB54D285656E074D06" pageId="164" pageNumber="165">
<normalizedToken id="3FCCA3948C95F9DEE733A9AAC76EC75C" originalValue="SUCUMBÍOS">SUCUMBIOS</normalizedToken>
: La Bonita, 1800 m (3 specimens CEMT)
</paragraph>
</subSubSection>
<subSubSection id="1DBC50CE3B3974EA2814447D6389D24D" pageId="164" pageNumber="165" type="literature records">
<paragraph id="B277080EBF35F21B5DDF151C5C56E9AD" pageId="164" pageNumber="165">Literature records.</paragraph>
<paragraph id="A9AFB2E5BC90F6D914998F35F5142ADC" pageId="164" pageNumber="165">
Without specific locality (
<normalizedToken id="EE9EFF08103C2DAE9F132E2674D8D02A" originalValue="Génier">Genier</normalizedToken>
, 1996: 48)
</paragraph>
</subSubSection>
<subSubSection id="909BFB9FAAF36BC0A9CC09126AEE3EE9" pageId="164" pageNumber="165" type="temporal data">
<paragraph id="58BD174DC9509EF0FA8CF0CB0B7895FE" pageId="164" pageNumber="165">Temporal data.</paragraph>
<paragraph id="52113717DA267544C6ABB6B4A0A4494E" pageId="164" pageNumber="165">Collected in May.</paragraph>
</subSubSection>
<subSubSection id="4E923500CA689A4093CA6DBE05CC9E42" pageId="164" pageNumber="165" type="remarks">
<paragraph id="EC0B41290FEBC3B0AD3886CE52CFDFCB" pageId="164" pageNumber="165">Remarks.</paragraph>
<paragraph id="4148947ACC725DFD1DA94C044A4682F8" pageId="164" pageNumber="165">Inhabits the montane cloud forests of the Andean region at 1800 m a.s.l. The collection method is unknown.</paragraph>
</subSubSection>
</treatment>
</document>

123
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Subfamily
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Chapuis, 1874: 54 [stem: Aulacoscelid-]. Type genus:
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Duponchel et Chevrolat, 1842. Comment: original vernacular name available
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Art. 11.7.2): first used in latinized form by Heyne and Taschenberg (1907: 245, as
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), generally accepted as in
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<taxonomicName id="0A9AA4D181BE44C37428753DB544901C" ID-CoL="3GV8W" authority="Verrill, 1873" authorityName="Verrill" authorityYear="1873" class="Polychaeta" family="Goniadidae" genus="Goniadella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goniadella gracilis" order="Phyllodocida" pageId="0" pageNumber="20997" phylum="Annelida" rank="species" species="gracilis">Goniadella gracilis (Verrill, 1873)</taxonomicName>
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<paragraph id="EEF586C11CF6C1EBD8982CAB4A3CAB34" pageId="0" pageNumber="20997">Notes</paragraph>
<paragraph id="93CD52569020B919384202D8297AFDB0" pageId="0" pageNumber="20997">
Reported from Greece by
<bibRefCitation id="9CD45CC6CCB73BFEE4C68C6380216ADD" author="Bogdanos, Costantinos" journalOrPublisher="Thalassographica" pageId="0" pageNumber="20997" pagination="77 - 105" title="The macrozoobenthos of an Aegean Embayment" volume="6" year="1983">Bogdanos and Satsmadjis (1983)</bibRefCitation>
and
<bibRefCitation id="B0886F8209C7C77E2710083136BF9439" author="Bogdanos, Costantinos" journalOrPublisher="Thalassographica" pageId="0" pageNumber="20997" pagination="37 - 71" title="The Patraikos Gulf bottom fauna" volume="10" year="1987">Bogdanos and Satsmadjis (1987)</bibRefCitation>
. In the Mediterranean also known from Israel (
<bibRefCitation id="E420AC9440AABDB156F19376CB9D2572" author="Boeggemann, M." journalOrPublisher="Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg" pageId="0" pageNumber="20997" pagination="1 - 354" title="Revision of the Goniadidae (Annelida, Polychaeta)" volume="39" year="2005">
<normalizedToken id="AC7B005943192D13CA0F68BA6D343E92" originalValue="Böggemann">Boeggemann</normalizedToken>
2005
</bibRefCitation>
), otherwise reported from the North-West Atlantic, East Atlantic and South-West Indian Ocean (
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<normalizedToken id="A33E5169260EEAB95C2CD41370F5B44E" originalValue="Böggemann">Boeggemann</normalizedToken>
2005
</bibRefCitation>
).
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91
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<paragraph id="696A902EE5D4FB9948DC61F85F436C60" pageNumber="285">&quot;Habitat in America meridionali.&quot; RCN: 2333.</paragraph>
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<paragraph id="B638C60F573C7586BCDD6869A5AC4A3B" pageNumber="285">Type not designated.</paragraph>
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Original material:
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Herb. Linn. No. 416.4 (
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; [icon] in Dillenius, Hort. Eltham. 1: 195, t. 162, f. 196. 1732; [icon] in Linnaeus in Kongl. Swenska Wetensk. Acad. Handl. 3: 93, t. 6. 1742; [icon] in Rudbeck, Campi Elysii 2: 89, f. 10. 1701.
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Current name:
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/
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193
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<figureCitation id="729300EBFFA0FFDE769337F8FF3EE1C1" box="[151,222,967,992]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1268,1290]" captionTargetBox="[230,1360,680,1228]" captionTargetId="figure@2.[226,1360,665,1244]" captionTargetPageId="2" captionText="FIGURES 1–5. Dicraeus species, habitus. 1 D. crassispinus sp. nov.; 2 D. curvus sp. nov.; 3 D. dentatus sp. nov.; 4 D. liupanshanensis sp. nov.; 5 D. minutidentatus sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623715/files/figure.png" pageId="11" pageNumber="362">Figs 5</figureCitation>
,
<figureCitation id="729300EBFFA0FFDE76EE37F7FED0E1C1" box="[234,304,968,992]" captionStart="FIGURES 28–31" captionStartId="12.[151,264,1968,1990]" captionTargetBox="[196,1402,332,1902]" captionTargetId="figure@12.[151,1436,301,1946]" captionTargetPageId="12" captionText="FIGURES 28–31. Dicraeus minutidentatus sp. nov. male. 28 epandrium, posterior view; 29 epandrium, lateral view; 30 hypandrium and phallic complex, ventral view; 31 hypandrium and phallic complex, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623730/files/figure.png" pageId="11" pageNumber="362">28–31</figureCitation>
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<emphasis id="D8DCC07CFFA0FFDE76933030FEF1E609" bold="true" box="[151,273,1039,1064]" pageId="11" pageNumber="362">Diagnosis.</emphasis>
Gena as wide as first flagellomere. Ocellar triangle reaching anterior 2/3 of frons. Scutum yellow with 3 longitudinal stripes, coalesced posteiorly. Legs yellow except for tarsomeres 5 brown. Surstylus broad distally, some thickened setae on inner side of lateral and distal portions. Cercus broad basally, distal 2/3 narrow, fingerlike, directed medioventrally. Postgonite bearing a minute serrulation on ventrodistal margin.
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<emphasis id="D8DCC07CFFA0FFDE76C330A0FEB9E699" bold="true" box="[199,345,1183,1208]" pageId="11" pageNumber="362">Description.</emphasis>
Male (
<figureCitation id="729300EBFFA0FFDE77AC30A0FE0CE699" box="[424,492,1183,1208]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1268,1290]" captionTargetBox="[230,1360,680,1228]" captionTargetId="figure@2.[226,1360,665,1244]" captionTargetPageId="2" captionText="FIGURES 1–5. Dicraeus species, habitus. 1 D. crassispinus sp. nov.; 2 D. curvus sp. nov.; 3 D. dentatus sp. nov.; 4 D. liupanshanensis sp. nov.; 5 D. minutidentatus sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623715/files/figure.png" pageId="11" pageNumber="362">Fig. 5</figureCitation>
). Body length
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.
</paragraph>
<paragraph id="EA171C6EFFA0FFDE76C330FBFB79E794" blockId="11.[151,1437,1039,2037]" pageId="11" pageNumber="362">Head yellow, 1.1 times as wide as thorax, 0.8 times as long as high in profile, face somewhat concave in lateral view, facial carina indistinct; frons as long as wide, projecting slightly in front of eye; gena broad, as wide as first flagellomere, with few brown setulae on ventral margin; parafacial indistinct; clypeus light yellow; vibrissal angle obtuse. Ocellar triangle yellow, smooth, shiny, reaching anterior 2/3 of frons, with pointed apex; ocellar tubercle yellowish brown. Occiput black. Cephalic setae and setulae black. Antenna yellow with thick grey pruinescence except for basodorsal margin of first flagellomere brown; first flagellomere as long as wide; arista brown with short brown pubescence. Proboscis yellow with yellow setulae; palpus yellow with black setulae.</paragraph>
<paragraph id="EA171C6EFFA0FFDE76C331FFFCCDE53C" blockId="11.[151,1437,1039,2037]" pageId="11" pageNumber="362">Scutum 0.9 times as long as wide, yellow with 3 longitudinal reddish yellow stripes except for brownish lateral stripes basally, coalesced anteriorly. Thoracic pleuron shiny yellow except for anteroventral portion of anepisternum and posteroventral portion of meron brownish yellow. Scutellum yellow, 0.5 times as long as wide; apical scutellar seta 1.5 times as long as scutellum; subapical scutellar seta as long as scutellum. Postscutellum brown. Setae and setulae on thorax black; notopleurals 1+2, anterior notopleural seta as long as posterior notopleural seta; anterior postalar seta 3 times as long as posterior postalar seta; 1 pair of dorsocentral setae developed, as long as anterior postalar setae. Legs yellow except for tarsomeres 5 brown. Mid tibia with a short black subapical spur, which is shorter than diameter of tibia. Tibial organ long and narrow. Setulae on legs brown. Wing 2.5 times as long as wide, hyaline; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 6: 2.4: 1; r-m at basal 0.6 of discal medial cell. Halter pale yellow.</paragraph>
<paragraph id="EA171C6EFFA0FFDE76C33318FE05E544" blockId="11.[151,1437,1039,2037]" pageId="11" pageNumber="362">Abdomen yellowish brown except for tergite 1 yellow, tergites 1+2 with a pair of brown apico-lateral spots; venter yellow. Setulae brown.</paragraph>
<paragraph id="EA171C6EFFA0FFD976C33350FC6EE2D9" blockId="11.[151,1437,1039,2037]" lastBlockId="12.[151,1436,151,248]" lastPageId="12" lastPageNumber="363" pageId="11" pageNumber="362">
Male genitalia (
<figureCitation id="729300EBFFA0FFDE777A3350FDECE5A9" box="[382,524,1903,1928]" captionStart="FIGURES 28–31" captionStartId="12.[151,264,1968,1990]" captionTargetBox="[196,1402,332,1902]" captionTargetId="figure@12.[151,1436,301,1946]" captionTargetPageId="12" captionText="FIGURES 28–31. Dicraeus minutidentatus sp. nov. male. 28 epandrium, posterior view; 29 epandrium, lateral view; 30 hypandrium and phallic complex, ventral view; 31 hypandrium and phallic complex, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623730/files/figure.png" pageId="11" pageNumber="362">Figs. 28–31</figureCitation>
): Epandrium brown with long brown setulae, height 1.7 times width in lateral view; surstylus slightly shorter than depth of epandrium, nearly parallel-sided in lateral view, broad distally, some thickened setae on inner side of lateral and distal portions. Cercus broad basally, distal 2/3 narrow, finger-like, directed medioventrally, with a point apex, with some long setulae. Pre- and postgonites short, postgonite round distally, bearing a minute serrulation on ventrodistal margin, with few sensilla; basiphallus longer than wide, cylindrical; distiphallus shorter than basiphallus, membranous; phallapodeme sclerotized, long, with basal stalk broad in lateral view. Hypandrium closed, apically forming an arch.
</paragraph>
<caption id="BED74CE6FFA7FFD97693338FFAA5E5C4" httpUri="https://zenodo.org/record/2623730/files/figure.png" pageId="12" pageNumber="363" startId="12.[151,264,1968,1990]" targetBox="[196,1402,332,1902]" targetPageId="12">
<paragraph id="EA171C6EFFA7FFD97693338FFAA5E5C4" blockId="12.[151,1435,1968,2021]" pageId="12" pageNumber="363">
<emphasis id="D8DCC07CFFA7FFD97693338FFEB7E5E7" bold="true" box="[151,343,1968,1990]" pageId="12" pageNumber="363">FIGURES 28–31.</emphasis>
<taxonomicName id="2DA867EDFFA7FFD97766338EFD80E5E7" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[354,608,1968,1990]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="minutidentatus" status="sp. nov.">
<emphasis id="D8DCC07CFFA7FFD97766338EFD80E5E7" box="[354,608,1968,1990]" italics="true" pageId="12" pageNumber="363">Dicraeus minutidentatus</emphasis>
</taxonomicName>
<emphasis id="D8DCC07CFFA7FFD9746F338FFD5FE5E7" bold="true" box="[619,703,1968,1990]" pageId="12" pageNumber="363">
<taxonomicNameLabel id="C3EF7D07FFA7FFD9746F338FFD5FE5E7" box="[619,703,1968,1990]" pageId="12" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male. 28 epandrium, posterior view; 29 epandrium, lateral view; 30 hypandrium and phallic complex, ventral view; 31 hypandrium and phallic complex, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="EA171C6EFFA6FFD876C334A8FB28E291" blockId="13.[151,1437,151,608]" box="[199,1224,151,176]" pageId="13" pageNumber="364">
Female. Body length
<quantity id="2D50B18BFFA6FFD877BC34A7FDF0E28E" box="[440,528,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="364" unit="mm" value="2.2">2.2 mm</quantity>
, wing length
<quantity id="2D50B18BFFA6FFD874AC34A7FCE0E28E" box="[680,768,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="364" unit="mm" value="1.9">1.9 mm</quantity>
. External characteristics similar to male.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA6FFD876C33483FE31E33C" pageId="13" pageNumber="364" type="materials_examined">
<paragraph id="EA171C6EFFA6FFD876C33483FE31E33C" blockId="13.[151,1437,151,608]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD876C33483FDFCE2F4" bold="true" box="[199,540,188,213]" pageId="13" pageNumber="364">
Type material.
<materialsCitation id="5AC01633FFA6FFD877793483FDFCE2F4" ID-GBIF-Occurrence="2234214338" box="[381,540,188,213]" collectingDate="2007-05-28" collectingDateMax="2007-06-11" collectingDateMin="2007-05-28" collectionCode="CAU" country="China" location="Diaoluoshan Mountain" pageId="13" pageNumber="364" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="3513A2CCFFA6FFD877793483FDFCE2F4" box="[381,540,188,213]" pageId="13" pageNumber="364" type="holotype">HOLOTYPE</typeStatus>
</materialsCitation>
</emphasis>
<emphasis id="D8DCC07CFFA6FFD874273483FDD8E2F5" box="[547,568,188,212]" italics="true" pageId="13" pageNumber="364">6</emphasis>
,
<collectingCountry id="92BF5CFEFFA6FFD874433483FD43E2F5" box="[583,675,188,212]" name="China" pageId="13" pageNumber="364">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA6FFD874AA3483FCE2E2F4" box="[686,770,188,213]" country="China" name="Hainan" pageId="13" pageNumber="364">Hainan</collectingRegion>
,
<location id="EF774AB5FFA6FFD8750A3483FBF3E2F4" LSID="urn:lsid:plazi:treatment:6201AD78FFA0FFD87693379AFCBCE041:EF774AB5FFA6FFD8750A3483FBF3E2F4" box="[782,1043,188,213]" country="China" name="Diaoluoshan Mountain" pageId="13" pageNumber="364" stateProvince="Hainan">Diaoluoshan Mountain</location>
,
<date id="9E163AAEFFA6FFD8721B3483FB16E2F5" box="[1055,1270,188,212]" pageId="13" pageNumber="364" value="2007-05-28" valueMax="2007-06-11" valueMin="2007-05-28">
<collectingDate id="8E52C346FFA6FFD8721B3483FB16E2F5" box="[1055,1270,188,212]" pageId="13" pageNumber="364" value="2007-05-28" valueMax="2007-06-11" valueMin="2007-05-28">28. V-11. VI. 2007</collectingDate>
</date>
, Jie Zeng Leg. (CAU) (photo and abdomen dissected).
<materialsCitation id="5AC01633FFA6FFD8745134E0FE2DE33C" ID-GBIF-Occurrence="2234214324" collectingDate="2007-05-28" collectingDateMax="2007-06-11" collectingDateMin="2007-05-28" collectionCode="CAU" country="China" location="Diaoluoshan Mountain" pageId="13" pageNumber="364" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="paratype">
<emphasis id="D8DCC07CFFA6FFD8745134E0FD19E2D9" bold="true" box="[597,761,223,248]" pageId="13" pageNumber="364">
<typeStatus id="3513A2CCFFA6FFD8745134E0FD19E2D9" box="[597,761,223,248]" pageId="13" pageNumber="364" type="paratype">PARATYPES</typeStatus>
</emphasis>
1
<emphasis id="D8DCC07CFFA6FFD8751034DFFCC9E2D9" box="[788,809,224,248]" italics="true" pageId="13" pageNumber="364">6</emphasis>
,
<specimenCount id="FCAED7E7FFA6FFD8753334DFFC81E2D6" box="[823,865,224,248]" pageId="13" pageNumber="364" type="female">1 ♀</specimenCount>
, same data as holotype (
<collectionCode id="8CB984ABFFA6FFD8727734DFFB54E2D9" box="[1139,1204,224,248]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="13" pageNumber="364">CAU</collectionCode>
). All type specimens were stored in 75% ethanol
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA6FFD876C33518FDFCE361" box="[199,540,295,320]" pageId="13" pageNumber="364" type="distribution">
<paragraph id="EA171C6EFFA6FFD876C33518FDFCE361" blockId="13.[151,1437,151,608]" box="[199,540,295,320]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD876C33518FE80E361" bold="true" box="[199,352,295,320]" pageId="13" pageNumber="364">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFA6FFD877633518FE4AE361" box="[359,426,295,320]" name="China" pageId="13" pageNumber="364">China</collectingCountry>
(
<collectingRegion id="286CD28CFFA6FFD877BD3518FDF0E361" box="[441,528,295,320]" country="China" name="Hainan" pageId="13" pageNumber="364">Hainan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA6FFD876C33573FD57E039" pageId="13" pageNumber="364" type="discussion">
<paragraph id="EA171C6EFFA6FFD876C33573FD57E039" blockId="13.[151,1437,151,608]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD876C33573FEDBE344" bold="true" box="[199,315,332,357]" pageId="13" pageNumber="364">Remarks.</emphasis>
The new species is somewhat similar to
<taxonomicName id="2DA867EDFFA6FFD875163572FB88E345" authority="Ismay, 1984" authorityName="Ismay" authorityYear="1984" box="[786,1128,332,357]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="longisurstylus">
<emphasis id="D8DCC07CFFA6FFD875163572FC32E345" box="[786,978,332,356]" italics="true" pageId="13" pageNumber="364">D. longisurstylus</emphasis>
<bibRefCitation id="8E39619FFFA6FFD875DF3572FB88E345" author="Ismay, J. W." box="[987,1128,332,357]" pageId="13" pageNumber="364" pagination="5 - 17" refId="ref12100" refString="Ismay, J. W. (1984) New species of Dicraeus (Diptera: Chloropidae) from New Guinea and New Ireland. Esakia, 21, 5 - 17." type="journal article" year="1984">Ismay, 1984</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the following features: scutum with 3 longitudinal reddish yellow stripes except for lateral stripes brownish basally; first flagellomere round dorsodistally; cercus broad basally, distal 2/3 narrow, finger-like; postgonite bearing a minute serrulation on ventrodistal margin. In
<taxonomicName id="2DA867EDFFA6FFD8757E3586FBD9E3F1" authorityName="Ismay" authorityYear="1984" box="[890,1081,440,464]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="longisurstylus">
<emphasis id="D8DCC07CFFA6FFD8757E3586FBD9E3F1" box="[890,1081,440,464]" italics="true" pageId="13" pageNumber="364">D. longisurstylus</emphasis>
</taxonomicName>
, the scutum has 3 black stripes; the first flagellomere has a dorsodistal angle; the cercus is narrow and pointed; and the postgonite has not minute serrulation on ventrodistal margin (
<bibRefCitation id="8E39619FFFA6FFD87425363FFD4AE039" author="Ismay, J. W." box="[545,682,512,536]" pageId="13" pageNumber="364" pagination="5 - 17" refId="ref12100" refString="Ismay, J. W. (1984) New species of Dicraeus (Diptera: Chloropidae) from New Guinea and New Ireland. Esakia, 21, 5 - 17." type="journal article" year="1984">Ismay 1984</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA6FFD876C3361BFCBCE041" pageId="13" pageNumber="364" type="etymology">
<paragraph id="EA171C6EFFA6FFD876C3361BFCBCE041" blockId="13.[151,1437,151,608]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD876C3361BFEAEE01C" bold="true" box="[199,334,548,573]" pageId="13" pageNumber="364">Etymology.</emphasis>
The specific name is from the Latin
<emphasis id="D8DCC07CFFA6FFD874FE361AFCA3E01D" box="[762,835,549,572]" italics="true" pageId="13" pageNumber="364">minut-</emphasis>
(“small”) and
<taxonomicName id="2DA867EDFFA6FFD875EA361BFBADE01D" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[1006,1101,548,572]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="D8DCC07CFFA6FFD875EA361BFBADE01D" box="[1006,1101,548,572]" italics="true" pageId="13" pageNumber="364">dentatus</emphasis>
</taxonomicName>
(“denticulate”), and refers to the ventrodistal margin of postgonite with a minute serrulation.
</paragraph>
</subSubSection>
</treatment>
</document>

171
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<paragraph id="EA171C6EFFA1FFDF76933573FDD8E347" blockId="10.[151,568,332,391]" box="[151,568,332,358]" pageId="10" pageNumber="361">
<heading id="B15FAB02FFA1FFDF76933573FDD8E347" bold="true" box="[151,568,332,358]" fontSize="11" level="1" pageId="10" pageNumber="361" reason="1">
<taxonomicName id="2DA867EDFFA1FFDF76933573FDD8E347" ID-CoL="35QQR" authority="Nartshuk, 2000" authorityName="Nartshuk, 2000:" authorityYear="1325" box="[151,568,332,358]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="D8DCC07CFFA1FFDF76933573FDD8E347" bold="true" box="[151,568,332,358]" pageId="10" pageNumber="361">
<emphasis id="D8DCC07CFFA1FFDF76933573FE90E347" bold="true" box="[151,368,332,358]" italics="true" pageId="10" pageNumber="361">Dicraeus malaisei</emphasis>
<bibRefCitation id="8E39619FFFA1FFDF77733573FDD8E347" author="Nartshuk, E. P." box="[375,568,332,358]" pageId="10" pageNumber="361" pagination="1325 - 1330" refId="ref12522" refString="Nartshuk, E. P. (2000) Grass seed-eating insects of orientalis group of the genus Dicraeus (Diptera, Chloropidae). Zoologicheskii Zhurnal, 79, 1325 - 1330. [In Russian]" type="journal article" year="2000">Nartshuk, 2000</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="EA171C6EFFA1FFDF76933551FED0E3A6" blockId="10.[151,568,332,391]" box="[151,304,366,391]" pageId="10" pageNumber="361">
<figureCitation id="729300EBFFA1FFDF76933551FF3EE3A7" box="[151,222,366,391]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="10" pageNumber="361">Figs 7</figureCitation>
,
<figureCitation id="729300EBFFA1FFDF76EE3550FEE8E3A6" box="[234,264,367,391]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="10" pageNumber="361">33</figureCitation>
,
<figureCitation id="729300EBFFA1FFDF77103550FED0E3A6" box="[276,304,367,391]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="10" pageNumber="361">39</figureCitation>
</paragraph>
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<subSubSection id="A2B24FE5FFA1FFDF76933587FCFDE3ED" box="[151,797,438,461]" pageId="10" pageNumber="361" type="reference_group">
<paragraph id="EA171C6EFFA1FFDF76933587FCFDE3ED" blockId="10.[151,797,438,461]" box="[151,797,438,461]" pageId="10" pageNumber="361">
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<taxonomicName id="2DA867EDFFA1FFDF76933587FDD5E3ED" ID-CoL="35QQR" authority="Nartshuk, 2000: 1325" authorityName="Nartshuk, 2000:" authorityPageNumber="1325" authorityYear="1325" box="[151,565,438,461]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="D8DCC07CFFA1FFDF76933587FEB0E3ED" box="[151,336,438,461]" italics="true" pageId="10" pageNumber="361">Dicraeus malaisei</emphasis>
<treatmentCitation id="6B093A7FFFA1FFDF77523589FDD5E3ED" author="Nartshuk, E. P." box="[342,565,438,460]" page="1325" pageId="10" pageNumber="361" year="2000">
<bibRefCitation id="8E39619FFFA1FFDF77523589FE16E3ED" author="Nartshuk, E. P." box="[342,502,438,460]" pageId="10" pageNumber="361" pagination="1325 - 1330" refId="ref12522" refString="Nartshuk, E. P. (2000) Grass seed-eating insects of orientalis group of the genus Dicraeus (Diptera, Chloropidae). Zoologicheskii Zhurnal, 79, 1325 - 1330. [In Russian]" type="journal article" year="2000">Nartshuk, 2000</bibRefCitation>
: 1325
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="3513A2CCFFA1FFDF743B3588FD91E3ED" box="[575,625,439,460]" pageId="10" pageNumber="361">Type</typeStatus>
locality:
<collectingCountry id="92BF5CFEFFA1FFDF74D53588FCF9E3ED" box="[721,793,439,460]" name="Myanmar" pageId="10" pageNumber="361">Burma</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="BED74CE6FFA1FFDF7693339FFAA5E5F8" httpUri="https://zenodo.org/record/2623727/files/figure.png" pageId="10" pageNumber="361" startId="10.[151,264,1952,1974]" targetBox="[187,1399,550,1903]" targetPageId="10">
<paragraph id="EA171C6EFFA1FFDF7693339FFAA5E5F8" blockId="10.[151,1435,1952,2009]" pageId="10" pageNumber="361">
<emphasis id="D8DCC07CFFA1FFDF7693339FFEB7E597" bold="true" box="[151,343,1952,1974]" pageId="10" pageNumber="361">FIGURES 24–27.</emphasis>
<taxonomicName id="2DA867EDFFA1FFDF7764339EFD8AE597" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[352,618,1952,1974]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="liupanshanensis" status="sp. nov.">
<emphasis id="D8DCC07CFFA1FFDF7764339EFD8AE597" box="[352,618,1952,1974]" italics="true" pageId="10" pageNumber="361">Dicraeus liupanshanensis</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="C3EF7D07FFA1FFDF7470339FFD28E597" box="[628,712,1952,1974]" pageId="10" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male. 24 epandrium, posterior view; 25 epandrium, lateral view; 26 hypandrium and phallic complex, ventral view; 27 hypandrium and phallic complex, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="A2B24FE5FFA0FFDE769334A8FC02E089" pageId="11" pageNumber="362" type="description">
<paragraph id="EA171C6EFFA0FFDE769334A8FD5DE01C" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="362">
<emphasis id="D8DCC07CFFA0FFDE769334A8FEC9E291" bold="true" box="[151,297,151,176]" pageId="11" pageNumber="362">Description.</emphasis>
Head brown (
<figureCitation id="729300EBFFA0FFDE77CE34A8FDEEE291" box="[458,526,151,176]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="11" pageNumber="362">Fig. 7</figureCitation>
), 0.8 times as long as high in profile; frons brown with distal 1/4 yellow, as long as wide; gena yellow, broad, 0.8 times as wide as first flagellomere. Ocellar triangle smooth, shiny, distal 1/4 yellow, reaching anterior margin of frons, with pointed apex. Antenna yellow except for dorsal 1/2 of first flagellomere brown; first flagellomere 0.9 times as long as wide; arista blackish brown with short brown pubescence. Proboscis yellow and palpus yellowish brown. Scutum black, 1.1 times as long as wide. Thoracic pleuron shiny brown with a longitudinal yellow stripe running from posterior angle of postpronotal lobe to wing base; ventral 1/2 of katepisternum black. Scutellum brown, 0.5 times as long as wide. Legs yellow except for fore tarsomeres 2–5, mid and hind tarsomeres 4–5 yellowish brown. Wing hyaline, 2.7 times as long as wide; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 8: 2: 1. Halter pale yellow. Abdomen brown except for tergite 1 and distal portions of tergites 2–5 yellow. Male genitalia (
<figureCitation id="729300EBFFA0FFDE75A735E3FBE3E3D5" box="[931,1027,476,501]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="11" pageNumber="362">Figs. 33</figureCitation>
,
<figureCitation id="729300EBFFA0FFDE721635E3FBD2E3D5" box="[1042,1074,476,500]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="11" pageNumber="362">39</figureCitation>
): Epandrium brown, height 1.5 times width in profile; cercus small; surstylus 2 times as long as epandrium, nearly parallel, recurved at middle in profile, with some short setulae on distal portion.
</paragraph>
<paragraph id="EA171C6EFFA0FFDE76C33678FD09E041" blockId="11.[151,1437,151,861]" box="[199,745,583,608]" pageId="11" pageNumber="362">
<emphasis id="D8DCC07CFFA0FFDE76C33678FEFEE041" bold="true" box="[199,286,583,608]" pageId="11" pageNumber="362">Female</emphasis>
. External characteristics similar to male.
</paragraph>
<paragraph id="EA171C6EFFA0FFDE76C33653FC02E089" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="362">
<emphasis id="D8DCC07CFFA0FFDE76C33653FE2AE0A4" bold="true" box="[199,458,620,645]" pageId="11" pageNumber="362">Specimens examined.</emphasis>
<materialsCitation id="5AC01633FFA0FFDE77D03653FC3EE089" ID-GBIF-Occurrence="2234214308" collectingDate="2007-04-24" collectionCode="CAU" collectorName="Menglun, No." country="China" location="Yunnan" pageId="11" pageNumber="362" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan">
<specimenCount id="FCAED7E7FFA0FFDE77D03653FDF9E0A5" box="[468,537,620,644]" pageId="11" pageNumber="362" type="male">2 ♂♂</specimenCount>
,
<collectingCountry id="92BF5CFEFFA0FFDE742D3653FD65E0A5" box="[553,645,620,644]" name="China" pageId="11" pageNumber="362">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA0FFDE74903652FD0DE0A5" box="[660,749,621,644]" country="China" name="Yunnan" pageId="11" pageNumber="362">Yunnan</collectingRegion>
,
<collectorName id="475D79B8FFA0FFDE74FF3653FC7CE0A5" box="[763,924,620,645]" pageId="11" pageNumber="362">Menglun, No.</collectorName>
55,
<date id="9E163AAEFFA0FFDE75D03653FB89E0A5" box="[980,1129,620,644]" pageId="11" pageNumber="362" value="2007-04-24">
<collectingDate id="8E52C346FFA0FFDE75D03653FB89E0A5" box="[980,1129,620,644]" pageId="11" pageNumber="362" value="2007-04-24">24. IV. 2007</collectingDate>
</date>
, Wenliang Li Leg. (
<collectionCode id="8CB984ABFFA0FFDE73523653FA74E0A5" box="[1366,1428,620,644]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="11" pageNumber="362">CAU</collectionCode>
) (photo and abdomen dissected). All specimens were stored in 75% ethanol
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA0FFDE76C3368BFD9CE0ED" box="[199,636,692,717]" pageId="11" pageNumber="362" type="distribution">
<paragraph id="EA171C6EFFA0FFDE76C3368BFD9CE0ED" blockId="11.[151,1437,151,861]" box="[199,636,692,717]" pageId="11" pageNumber="362">
<emphasis id="D8DCC07CFFA0FFDE76C3368BFE80E0EC" bold="true" box="[199,352,692,717]" pageId="11" pageNumber="362">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFA0FFDE7763368BFE4AE0EC" box="[359,426,692,717]" name="China" pageId="11" pageNumber="362">China</collectingCountry>
(
<collectingRegion id="286CD28CFFA0FFDE77BD368BFDF5E0ED" box="[441,533,692,716]" country="China" name="Yunnan" pageId="11" pageNumber="362">Yunnan</collectingRegion>
);
<collectingCountry id="92BF5CFEFFA0FFDE742C368AFD98E0ED" box="[552,632,693,716]" name="Myanmar" pageId="11" pageNumber="362">Burma</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA0FFDE76C336E8FE01E17D" pageId="11" pageNumber="362" type="discussion">
<paragraph id="EA171C6EFFA0FFDE76C336E8FE01E17D" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="362">
<emphasis id="D8DCC07CFFA0FFDE76C336E8FEDBE0D1" bold="true" box="[199,315,727,752]" pageId="11" pageNumber="362">Remarks.</emphasis>
The new species is somewhat similar to
<taxonomicName id="2DA867EDFFA0FFDE751436E6FB89E0D1" authority="Kanmiya, 1971" authorityName="Kanmiya" authorityYear="1971" box="[784,1129,727,752]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="362" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="D8DCC07CFFA0FFDE751436E6FC51E0D1" box="[784,945,728,752]" italics="true" pageId="11" pageNumber="362">D. nartshukae</emphasis>
<bibRefCitation id="8E39619FFFA0FFDE75BC36E8FB89E0D1" author="Kanmiya, K." box="[952,1129,727,752]" pageId="11" pageNumber="362" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the propleuron black, and surstylus 2 times as long as epandrium, nearly parallel, recurved at middle in lateral view. In
<taxonomicName id="2DA867EDFFA0FFDE7740371EFE02E119" authorityName="Kanmiya" authorityYear="1971" box="[324,482,800,824]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="362" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="D8DCC07CFFA0FFDE7740371EFE02E119" box="[324,482,800,824]" italics="true" pageId="11" pageNumber="362">D. nartshukae</emphasis>
</taxonomicName>
, the propleuron is yellowish; and the surstylus is distinctly shorter than epandrium in lateral view (
<bibRefCitation id="8E39619FFFA0FFDE772D377BFE35E17D" author="Nartshuk, E. P." box="[297,469,836,861]" pageId="11" pageNumber="362" pagination="1325 - 1330" refId="ref12522" refString="Nartshuk, E. P. (2000) Grass seed-eating insects of orientalis group of the genus Dicraeus (Diptera, Chloropidae). Zoologicheskii Zhurnal, 79, 1325 - 1330. [In Russian]" type="journal article" year="2000">Nartshuk 2000</bibRefCitation>
).
</paragraph>
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171
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<emphasis id="D8DCC07CFFA2FFDC769335EEFDD5E3CA" bold="true" box="[151,565,465,491]" pageId="9" pageNumber="360">
<taxonomicName id="2DA867EDFFA2FFDC769335EEFE30E3CA" ID-CoL="9HS9M" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[151,464,465,491]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="360" phylum="Arthropoda" rank="species" species="liupanshanensis" status="sp. nov.">
<emphasis id="D8DCC07CFFA2FFDC769335EEFE30E3CA" bold="true" box="[151,464,465,491]" italics="true" pageId="9" pageNumber="360">Dicraeus liupanshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3EF7D07FFA2FFDC77D335EEFDD5E3CA" box="[471,565,465,491]" pageId="9" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA2FFDC769335CBFED0E02D" box="[151,304,500,525]" pageId="9" pageNumber="360" type="description">
<paragraph id="EA171C6EFFA2FFDC769335CBFED0E02D" blockId="9.[151,565,465,525]" box="[151,304,500,525]" pageId="9" pageNumber="360">
<figureCitation id="729300EBFFA2FFDC769335CBFF3EE02D" box="[151,222,500,525]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1268,1290]" captionTargetBox="[230,1360,680,1228]" captionTargetId="figure@2.[226,1360,665,1244]" captionTargetPageId="2" captionText="FIGURES 1–5. Dicraeus species, habitus. 1 D. crassispinus sp. nov.; 2 D. curvus sp. nov.; 3 D. dentatus sp. nov.; 4 D. liupanshanensis sp. nov.; 5 D. minutidentatus sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623715/files/figure.png" pageId="9" pageNumber="360">Figs 4</figureCitation>
,
<figureCitation id="729300EBFFA2FFDC76EE35CBFED0E02D" box="[234,304,500,524]" captionStart="FIGURES 24–27" captionStartId="10.[151,264,1952,1974]" captionTargetBox="[187,1399,550,1903]" captionTargetId="figure@10.[171,1428,513,1939]" captionTargetPageId="10" captionText="FIGURES 24–27. Dicraeus liupanshanensis sp. nov. male. 24 epandrium, posterior view; 25 epandrium, lateral view; 26 hypandrium and phallic complex, ventral view; 27 hypandrium and phallic complex, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623727/files/figure.png" pageId="9" pageNumber="360">24–27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA2FFDC76933603FD4FE0C4" pageId="9" pageNumber="360" type="diagnosis">
<paragraph id="EA171C6EFFA2FFDC76933603FD4FE0C4" blockId="9.[151,1437,572,2037]" pageId="9" pageNumber="360">
<emphasis id="D8DCC07CFFA2FFDC76933603FEF1E074" bold="true" box="[151,273,572,597]" pageId="9" pageNumber="360">Diagnosis.</emphasis>
Gena about 0.8 times as wide as first flagellomere. Ocellar triangle reaching anterior margin of frons. Scutum yellow with 5 longitudinal stripes. Scutellum yellow. Legs largely yellow, but hind femur and tibia brown with both ends yellow; fore and mid tarsomeres 2–5 and hind tarsus (sometimes tarsomeres 2–5) brown. Epandrium bearing with apical and subapical processes on ventral margin. Surstylus broad distally, some thickened setae on inner side of lateral and distal portions.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA2FFDC76C336D0FE7EE4D9" pageId="9" pageNumber="360" type="description">
<paragraph id="EA171C6EFFA2FFDC76C336D0FC3FE126" blockId="9.[151,1437,572,2037]" box="[199,991,751,776]" pageId="9" pageNumber="360">
<emphasis id="D8DCC07CFFA2FFDC76C336D0FEB9E129" bold="true" box="[199,345,751,776]" pageId="9" pageNumber="360">Description.</emphasis>
Male (
<figureCitation id="729300EBFFA2FFDC77AC36D0FE0DE129" box="[424,493,751,776]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1268,1290]" captionTargetBox="[230,1360,680,1228]" captionTargetId="figure@2.[226,1360,665,1244]" captionTargetPageId="2" captionText="FIGURES 1–5. Dicraeus species, habitus. 1 D. crassispinus sp. nov.; 2 D. curvus sp. nov.; 3 D. dentatus sp. nov.; 4 D. liupanshanensis sp. nov.; 5 D. minutidentatus sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623715/files/figure.png" pageId="9" pageNumber="360">Fig. 4</figureCitation>
). Body length
<quantity id="2D50B18BFFA2FFDC749636D0FD0AE126" box="[658,746,751,776]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="360" unit="mm" value="2.6">2.6 mm</quantity>
, wing length
<quantity id="2D50B18BFFA2FFDC758636CFFC3BE126" box="[898,987,752,776]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="360" unit="mm" value="2.8">2.8 mm</quantity>
.
</paragraph>
<paragraph id="EA171C6EFFA2FFDC76C3372BFBB6E609" blockId="9.[151,1437,572,2037]" pageId="9" pageNumber="360">Head yellow, as wide as thorax, 0.9 times as long as high in lateral view, face somewhat concave in lateral view, facial carina narrow; frons brown with anterior 1/2 yellow, as long as wide, projecting slightly in front of eye; gena broad, about 0.8 times as wide as first flagellomere, with few brown setulae on ventral margin; parafacial indistinct; clypeus light yellow; vibrissal angle obtuse. Ocellar triangle yellowish brown with distal 1/2 yellow, smooth, shiny, reaching anterior margin of frons, with pointed apex; ocellar tubercle black. Occiput brown. Cephalic setae and setulae black. Antenna yellow with thick grey pruinescence except for dorsal margin of first flagellomere yellowish brown; first flagellomere 0.9 times as long as wide; arista brown with short brown pubescence. Proboscis yellow with yellow setulae; palpus yellow with brown setulae.</paragraph>
<paragraph id="EA171C6EFFA2FFDC76C3300BFB80E7F9" blockId="9.[151,1437,572,2037]" pageId="9" pageNumber="360">Scutum as long as wide, yellow with 5 longitudinal stripes, medial stripe brown, broad and extending to posterior 1/3 of scutum, inner lateral stripes reddish yellow with distal 1/3 brown, broad, outer lateral stripes reddish yellow, short and narrow. Postpronotal lobe yellow with a brown spot. Thoracic pleuron shiny yellow except for anteroventral portion of anepisternum, ventral 3/4 of katepisternum and ventral 1/2 meron reddish yellow. Scutellum yellow, 0.7 times as long as wide; apical scutellar seta as long as scutellum; subapical scutellar seta 0.5 times as long as scutellum. Postscutellum black. Setae and setulae on thorax black; notopleurals 1+2, anterior notopleural seta as long as posterior notopleural seta; anterior postalar seta as long as posterior postalar seta; 1 pair of dorsocentral setae developed, 2 times as long as anterior postalar setae. Legs yellow except for hind femur and tibia brown with both ends yellow, fore and mid tarsomeres 2–5, hind tarsus (sometimes tarsomeres 2–5) brown. Mid tibia with a short black subapical spur, which is shorter than diameter of tibia. Tibial organ long and narrow. Setulae on legs black. Wing 2.5 times as long as wide, hyaline; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 8: 2: 1; r-m at basal 0.5 of discal medial cell. Halter pale yellow.</paragraph>
<paragraph id="EA171C6EFFA2FFDC76C331DBFAA6E7DC" blockId="9.[151,1437,572,2037]" box="[199,1350,1508,1533]" pageId="9" pageNumber="360">Abdomen yellow except for tergites 2–5 with a longitudinal brown spot; venter yellow. Setulae brown.</paragraph>
<paragraph id="EA171C6EFFA2FFDC76C33238FCA7E4F4" blockId="9.[151,1437,572,2037]" pageId="9" pageNumber="360">
Male genitalia (
<figureCitation id="729300EBFFA2FFDC777A3238FDECE401" box="[382,524,1543,1568]" captionStart="FIGURES 24–27" captionStartId="10.[151,264,1952,1974]" captionTargetBox="[187,1399,550,1903]" captionTargetId="figure@10.[171,1428,513,1939]" captionTargetPageId="10" captionText="FIGURES 24–27. Dicraeus liupanshanensis sp. nov. male. 24 epandrium, posterior view; 25 epandrium, lateral view; 26 hypandrium and phallic complex, ventral view; 27 hypandrium and phallic complex, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623727/files/figure.png" pageId="9" pageNumber="360">Figs. 24–27</figureCitation>
): Epandrium brown with long brown setulae, height 1.5 times width in lateral view, with apical and subapical processes on ventral margin, apical process distinctly curved inward in dorsal view; surstylus slightly shorter than depth of epandrium, nearly parallel-sided in lateral view, broad distally, some thickened setae on inner side of lateral and distal portions. Pre- and postgonites short, postgonite round distally, with few sensilla; basiphallus longer than wide, cylindrical; phallapodeme sclerotized, long, with basal stalk broad in lateral view. Hypandrium closed, apically forming an arch.
</paragraph>
<paragraph id="EA171C6EFFA2FFDC76C332E0FE7EE4D9" blockId="9.[151,1437,572,2037]" box="[199,414,1759,1784]" pageId="9" pageNumber="360">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA2FFDC76C3333BFCB0E544" pageId="9" pageNumber="360" type="materials_examined">
<paragraph id="EA171C6EFFA2FFDC76C3333BFCB0E544" blockId="9.[151,1437,572,2037]" pageId="9" pageNumber="360">
<emphasis id="D8DCC07CFFA2FFDC76C3333BFDFFE53C" bold="true" box="[199,543,1796,1821]" pageId="9" pageNumber="360">
Type material.
<materialsCitation id="5AC01633FFA2FFDC7784333BFDFFE53C" ID-GBIF-Occurrence="2234214323" box="[384,543,1796,1821]" collectingDate="2007-07-16" collectionCode="CAU" country="China" location="Botanical" municipality="Liupanshan Mountain" pageId="9" pageNumber="360" specimenCount="1" stateProvince="Ningxia" typeStatus="holotype">
<typeStatus id="3513A2CCFFA2FFDC7784333BFDFFE53C" box="[384,543,1796,1821]" pageId="9" pageNumber="360" type="holotype">HOLOTYPE</typeStatus>
</materialsCitation>
</emphasis>
<emphasis id="D8DCC07CFFA2FFDC742C333BFDDDE53D" box="[552,573,1796,1820]" italics="true" pageId="9" pageNumber="360">6</emphasis>
,
<collectingCountry id="92BF5CFEFFA2FFDC7448333BFD48E53D" box="[588,680,1796,1820]" name="China" pageId="9" pageNumber="360">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA2FFDC74B1333BFCF2E53C" box="[693,786,1796,1821]" country="China" name="Ningxia" pageId="9" pageNumber="360">Ningxia</collectingRegion>
,
<collectingMunicipality id="0A738614FFA2FFDC7524333BFBFCE53C" box="[800,1052,1796,1821]" pageId="9" pageNumber="360">Liupanshan Mountain</collectingMunicipality>
,
<location id="EF774AB5FFA2FFDC722D333BFB74E53C" LSID="urn:lsid:plazi:treatment:6201AD78FFA2FFDF769335EEFBDEE2D9:EF774AB5FFA2FFDC722D333BFB74E53C" box="[1065,1172,1796,1821]" country="China" municipality="Liupanshan Mountain" name="Botanical" pageId="9" pageNumber="360" stateProvince="Ningxia">Botanical</location>
garden,
<date id="9E163AAEFFA2FFDC72FE333BFA78E53D" box="[1274,1432,1796,1821]" pageId="9" pageNumber="360" value="2007-07-16">
<collectingDate id="8E52C346FFA2FFDC72FE333BFA78E53D" box="[1274,1432,1796,1821]" pageId="9" pageNumber="360" value="2007-07-16">16. VII. 2007</collectingDate>
</date>
, Qibiao Dong Leg. (CAU) (photo and abdomen dissected).
<materialsCitation id="5AC01633FFA2FFDC752C3318FCACE544" ID-GBIF-Occurrence="2234214328" collectingDate="2007-07-16" collectionCode="CAU" country="China" location="Botanical" municipality="Liupanshan Mountain" pageId="9" pageNumber="360" specimenCount="1" stateProvince="Ningxia" typeStatus="paratype">
<emphasis id="D8DCC07CFFA2FFDC752C3318FC2CE561" bold="true" box="[808,972,1831,1856]" pageId="9" pageNumber="360">
<typeStatus id="3513A2CCFFA2FFDC752C3318FC2CE561" box="[808,972,1831,1856]" pageId="9" pageNumber="360" type="paratype">PARATYPES</typeStatus>
</emphasis>
2
<emphasis id="D8DCC07CFFA2FFDC75E33317FBF1E561" box="[999,1041,1832,1856]" italics="true" pageId="9" pageNumber="360">66</emphasis>
, same data as holotype (
<collectionCode id="8CB984ABFFA2FFDC73203317FA85E561" box="[1316,1381,1832,1856]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="9" pageNumber="360">CAU</collectionCode>
). All type specimens dry mounted on label laterally on right thorax
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA2FFDC76C33350FDC5E5A9" box="[199,549,1903,1928]" pageId="9" pageNumber="360" type="distribution">
<paragraph id="EA171C6EFFA2FFDC76C33350FDC5E5A9" blockId="9.[151,1437,572,2037]" box="[199,549,1903,1928]" pageId="9" pageNumber="360">
<emphasis id="D8DCC07CFFA2FFDC76C33350FE80E5A9" bold="true" box="[199,352,1903,1928]" pageId="9" pageNumber="360">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFA2FFDC77633350FE4AE5A9" box="[359,426,1903,1928]" name="China" pageId="9" pageNumber="360">China</collectingCountry>
(
<collectingRegion id="286CD28CFFA2FFDC77BD3350FDFAE5A9" box="[441,538,1903,1928]" country="China" name="Ningxia" pageId="9" pageNumber="360">Ningxia</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA2FFDF76C333ABFA7CE2F5" lastPageId="10" lastPageNumber="361" pageId="9" pageNumber="360" type="discussion">
<paragraph id="EA171C6EFFA2FFDF76C333ABFA7CE2F5" blockId="9.[151,1437,572,2037]" lastBlockId="10.[151,1436,151,248]" lastPageId="10" lastPageNumber="361" pageId="9" pageNumber="360">
<emphasis id="D8DCC07CFFA2FFDC76C333ABFEDBE58C" bold="true" box="[199,315,1940,1965]" pageId="9" pageNumber="360">Remarks.</emphasis>
The new species is somewhat similar to
<taxonomicName id="2DA867EDFFA2FFDC751633AAFB88E58D" authority="Ismay, 1984" authorityName="Ismay" authorityYear="1984" box="[786,1128,1940,1965]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="360" phylum="Arthropoda" rank="species" species="longisurstylus">
<emphasis id="D8DCC07CFFA2FFDC751633AAFC32E58D" box="[786,978,1940,1964]" italics="true" pageId="9" pageNumber="360">D. longisurstylus</emphasis>
<bibRefCitation id="8E39619FFFA2FFDC75DF33AAFB88E58D" author="Ismay, J. W." box="[987,1128,1940,1965]" pageId="9" pageNumber="360" pagination="5 - 17" refId="ref12100" refString="Ismay, J. W. (1984) New species of Dicraeus (Diptera: Chloropidae) from New Guinea and New Ireland. Esakia, 21, 5 - 17." type="journal article" year="1984">Ismay, 1984</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the following features: ocellar triangle reaching anterior margin of frons; first flagellomere round dorsodistally; scutum with 5 brown to reddish yellow stripes; epandrium with apical and subapical processes on ventral margin. In
<taxonomicName id="2DA867EDFFA1FFDF776F34A6FDCCE291" authorityName="Ismay" authorityYear="1984" box="[363,556,152,176]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="longisurstylus">
<emphasis id="D8DCC07CFFA1FFDF776F34A6FDCCE291" box="[363,556,152,176]" italics="true" pageId="10" pageNumber="361">D. longisurstylus</emphasis>
</taxonomicName>
, the ocellar triangle reaches anterior 2/3 of frons; the first flagellomere has a dorsodistal angle; the scutum has 3 black stripes, fused basally; and the cercus is narrow and pointed (
<bibRefCitation id="8E39619FFFA1FFDF73033483FA70E2F5" author="Ismay, J. W." box="[1287,1424,188,212]" pageId="10" pageNumber="361" pagination="5 - 17" refId="ref12100" refString="Ismay, J. W. (1984) New species of Dicraeus (Diptera: Chloropidae) from New Guinea and New Ireland. Esakia, 21, 5 - 17." type="journal article" year="1984">Ismay 1984</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA1FFDF76C334E0FBDEE2D9" box="[199,1086,223,248]" pageId="10" pageNumber="361" type="etymology">
<paragraph id="EA171C6EFFA1FFDF76C334E0FBDEE2D9" blockId="10.[151,1436,151,248]" box="[199,1086,223,248]" pageId="10" pageNumber="361">
<emphasis id="D8DCC07CFFA1FFDF76C334E0FEA6E2D9" bold="true" box="[199,326,223,248]" pageId="10" pageNumber="361">Etymology</emphasis>
. The species is named after the
<typeStatus id="3513A2CCFFA1FFDF74A934DFFD3DE2D9" box="[685,733,224,248]" pageId="10" pageNumber="361">type</typeStatus>
locality Liupanshan Mountain.
</paragraph>
</subSubSection>
</treatment>
</document>

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<heading id="B15FAB02FFA3FFDD7693319EFDBAE79A" bold="true" box="[151,602,1441,1467]" fontSize="11" level="1" pageId="8" pageNumber="359" reason="1">
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<emphasis id="D8DCC07CFFA3FFDD7693319EFE75E79A" bold="true" box="[151,405,1441,1467]" italics="true" pageId="8" pageNumber="359">Dicraeus fuscipennis</emphasis>
<bibRefCitation id="8E39619FFFA3FFDD7799319EFDBAE79A" author="Kanmiya, K." box="[413,602,1441,1467]" pageId="8" pageNumber="359" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
</emphasis>
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<paragraph id="EA171C6EFFA3FFDD769331FBFED0E7FD" blockId="8.[151,602,1441,1501]" box="[151,304,1476,1501]" pageId="8" pageNumber="359">
<figureCitation id="729300EBFFA3FFDD769331FBFF3EE7FC" box="[151,222,1476,1501]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="8" pageNumber="359">Figs 6</figureCitation>
,
<figureCitation id="729300EBFFA3FFDD76EE31FBFEE9E7FD" box="[234,265,1476,1500]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="8" pageNumber="359">32</figureCitation>
,
<figureCitation id="729300EBFFA3FFDD771031FBFED0E7FD" box="[276,304,1476,1500]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="8" pageNumber="359">38</figureCitation>
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<treatmentCitationGroup id="CAB83B40FFA3FFDD76933232FCCBE403" box="[151,811,1548,1570]" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD76933232FF1EE403" box="[151,254,1549,1570]" italics="true" pageId="8" pageNumber="359">Discraeus</emphasis>
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<emphasis id="D8DCC07CFFA3FFDD77003233FE96E403" box="[260,374,1548,1570]" italics="true" pageId="8" pageNumber="359">fuscipennis</emphasis>
<treatmentCitation id="6B093A7FFFA3FFDD77783233FDAEE403" author="Kanmiya, K." box="[380,590,1548,1570]" page="173" pageId="8" pageNumber="359" year="1971">
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: 173
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.
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locality:
<collectingCountry id="92BF5CFEFFA3FFDD74EE3232FCC7E403" box="[746,807,1549,1570]" name="Japan" pageId="8" pageNumber="359">Japan</collectingCountry>
.
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<subSubSection id="A2B24FE5FFA3FFDC76933271FBD7E33C" lastPageId="9" lastPageNumber="360" pageId="8" pageNumber="359" type="description">
<paragraph id="EA171C6EFFA3FFDC76933271FE04E2F4" blockId="8.[151,1437,1614,2035]" lastBlockId="9.[151,1437,151,392]" lastPageId="9" lastPageNumber="360" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD76933271FEC9E446" bold="true" box="[151,297,1614,1639]" pageId="8" pageNumber="359">Description.</emphasis>
Head brown (
<figureCitation id="729300EBFFA3FFDD77CE3271FDEEE446" box="[458,526,1614,1639]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="8" pageNumber="359">Fig. 6</figureCitation>
), 0.8 times as long as high in profile; frons brown with distal portion yellow, 0.9 times as long as wide; gena yellow, broad, 0.7 times as wide as first flagellomere. Ocellar triangle smooth, shiny, distal 1/8 yellow, reaching anterior margin of frons, with pointed apex. Antenna yellow except for dorsal margin of first flagellomere brown; first flagellomere 0.9 times as long as wide; arista brown with short brown pubescence. Proboscis and palpus yellow. Scutum black, 1.1 times as long as wide. Thoracic pleuron shiny blackish brown except for propleuron largely yellow; ventral portion of katepimeron, dorsal and posterior margins of anepisternum narrowly yellow. Scutellum brown, 0.5 times as long as wide. Legs yellow except for fore and mid femora yellow to brownish yellow, hind femur brown with both ends yellow, middle 1/2 of hind tibia brown, all tarsomeres 3–5 brown. Wing tinged pale brown, 2.7 times as long as wide; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 6: 1.6: 1. Halter pale yellow. Abdomen brown except for tergite 1 yellow. Male genitalia (
<figureCitation id="729300EBFFA3FFDD730233ADFA83E58A" box="[1286,1379,1938,1963]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="8" pageNumber="359">Figs. 32</figureCitation>
,
<figureCitation id="729300EBFFA3FFDD736B33ACFA6FE58A" box="[1391,1423,1939,1963]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="8" pageNumber="359">38</figureCitation>
): Epandrium dark brown, with apical and subapical processes on ventral margin, apical process distinctly curved inward in dorsal view; surstylus as long as epandrium, nearly parallel-sided in profile, extended caudad, slightly broad, some thickened setae on inner side of lateral and distal portions. Postgonite round distally, bearing minute serrulations on apical margin.
</paragraph>
<paragraph id="EA171C6EFFA2FFDC76C334E0FBD7E33C" blockId="9.[151,1437,151,392]" pageId="9" pageNumber="360">
<emphasis id="D8DCC07CFFA2FFDC76C334E0FE2AE2D9" bold="true" box="[199,458,223,248]" pageId="9" pageNumber="360">Specimens examined.</emphasis>
<materialsCitation id="5AC01633FFA2FFDC77D034E0FBD3E33C" ID-GBIF-Occurrence="2234214334" collectingDate="2009-06-02" collectionCode="CAU" country="China" location="Songshan Mountain" municipality="Yanqing" pageId="9" pageNumber="360" specimenCount="6" specimenCount-male="6" stateProvince="Beijing">
<specimenCount id="FCAED7E7FFA2FFDC77D034E0FDF8E2D6" box="[468,536,223,248]" pageId="9" pageNumber="360" type="male">6 ♂♂</specimenCount>
,
<collectingCountry id="92BF5CFEFFA2FFDC742C34DFFD63E2D9" box="[552,643,224,248]" name="China" pageId="9" pageNumber="360">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA2FFDC749634E0FD08E2D9" box="[658,744,223,248]" country="China" name="Beijing" pageId="9" pageNumber="360">Beijing</collectingRegion>
,
<collectingMunicipality id="0A738614FFA2FFDC74F234E0FCB6E2D9" box="[758,854,223,248]" pageId="9" pageNumber="360">Yanqing</collectingMunicipality>
,
<location id="EF774AB5FFA2FFDC756234E0FBADE2D9" LSID="urn:lsid:plazi:treatment:6201AD78FFA3FFDC7693319EFEB6E3A9:EF774AB5FFA2FFDC756234E0FBADE2D9" box="[870,1101,223,248]" country="China" municipality="Yanqing" name="Songshan Mountain" pageId="9" pageNumber="360" stateProvince="Beijing">Songshan Mountain</location>
,
<date id="9E163AAEFFA2FFDC725834DFFB05E2D9" box="[1116,1253,224,248]" pageId="9" pageNumber="360" value="2009-06-02">
<collectingDate id="8E52C346FFA2FFDC725834DFFB05E2D9" box="[1116,1253,224,248]" pageId="9" pageNumber="360" value="2009-06-02">2. VI. 2009</collectingDate>
</date>
, Qifei Liu Leg. (
<collectionCode id="8CB984ABFFA2FFDC769A353BFF3CE33D" box="[158,220,260,284]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="9" pageNumber="360">CAU</collectionCode>
) (photo and abdomen dissected). All specimens were stored in 75% ethanol
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA2FFDC76C33518FD8CE31E" box="[199,620,295,320]" pageId="9" pageNumber="360" type="distribution">
<paragraph id="EA171C6EFFA2FFDC76C33518FD8CE31E" blockId="9.[151,1437,151,392]" box="[199,620,295,320]" pageId="9" pageNumber="360">
<emphasis id="D8DCC07CFFA2FFDC76C33518FE80E361" bold="true" box="[199,352,295,320]" pageId="9" pageNumber="360">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFA2FFDC77633518FE4AE361" box="[359,426,295,320]" name="China" pageId="9" pageNumber="360">China</collectingCountry>
(
<collectingRegion id="286CD28CFFA2FFDC77BD3518FDF2E361" box="[441,530,295,320]" country="China" name="Beijing" pageId="9" pageNumber="360">Beijing</collectingRegion>
);
<collectingCountry id="92BF5CFEFFA2FFDC74213517FD88E31E" box="[549,616,296,319]" name="Japan" pageId="9" pageNumber="360">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA2FFDC76C33573FEB6E3A9" pageId="9" pageNumber="360" type="discussion">
<paragraph id="EA171C6EFFA2FFDC76C33573FEB6E3A9" blockId="9.[151,1437,151,392]" pageId="9" pageNumber="360">
<emphasis id="D8DCC07CFFA2FFDC76C33573FEDBE344" bold="true" box="[199,315,332,357]" pageId="9" pageNumber="360">Remarks.</emphasis>
This species is distinguished from all the known species in its wing tinged with pale brown (
<bibRefCitation id="8E39619FFFA2FFDC769A3550FEAAE3A9" author="Kanmiya, K." box="[158,330,367,392]" pageId="9" pageNumber="360" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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117
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<taxonomicName id="2DA867EDFFA3FFDD7693377BFE1DE17F" ID-CoL="35QQ7" authority="Duda, 1930" authorityName="Duda" authorityYear="1930" box="[151,509,836,862]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="D8DCC07CFFA3FFDD7693377BFE1DE17F" bold="true" box="[151,509,836,862]" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD7693377BFE87E17F" bold="true" box="[151,359,836,862]" italics="true" pageId="8" pageNumber="359">Dicraeus flavipes</emphasis>
<bibRefCitation id="8E39619FFFA3FFDD776B377BFE1DE17F" author="Duda, O." box="[367,509,836,862]" pageId="8" pageNumber="359" pagination="278 - 304" refId="ref11917" refString="Duda, O. (1930) Neue und bekannte orientalische Chloropiden (Dipt.) des Deutschen Entomologischen Instituts in Berlin- Dahlem. Stettiner entomologische Zeitung, 91, 278 - 304." type="journal article" year="1930">Duda, 1930</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA3FFDD769337B4FCD9E181" box="[151,825,906,928]" pageId="8" pageNumber="359" type="reference_group">
<paragraph id="EA171C6EFFA3FFDD769337B4FCD9E181" blockId="8.[151,825,906,928]" box="[151,825,906,928]" pageId="8" pageNumber="359">
<taxonomicName id="2DA867EDFFA3FFDD769337B4FE2CE181" ID-CoL="35QQ7" authority="Duda, 1930" authorityName="Duda" authorityYear="1930" box="[151,460,906,928]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="D8DCC07CFFA3FFDD769337B4FEA9E181" box="[151,329,906,928]" italics="true" pageId="8" pageNumber="359">Dicraeus flavipes</emphasis>
<bibRefCitation id="8E39619FFFA3FFDD774B37B5FE2CE181" author="Duda, O." box="[335,460,906,928]" pageId="8" pageNumber="359" pagination="278 - 304" refId="ref11917" refString="Duda, O. (1930) Neue und bekannte orientalische Chloropiden (Dipt.) des Deutschen Entomologischen Instituts in Berlin- Dahlem. Stettiner entomologische Zeitung, 91, 278 - 304." type="journal article" year="1930">Duda, 1930</bibRefCitation>
</taxonomicName>
.: 295.
<typeStatus id="3513A2CCFFA3FFDD740F37B4FDDDE181" box="[523,573,907,928]" pageId="8" pageNumber="359">Type</typeStatus>
locality:
<collectingCountry id="92BF5CFEFFA3FFDD749937B5FD3DE181" box="[669,733,906,928]" name="China" pageId="8" pageNumber="359">China</collectingCountry>
:
<collectingCountry id="92BF5CFEFFA3FFDD74E337B5FCD5E181" box="[743,821,906,928]" name="Taiwan" pageId="8" pageNumber="359">Taiwan</collectingCountry>
.
</paragraph>
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<subSubSection id="A2B24FE5FFA3FFDD769337F2FDDBE6BB" pageId="8" pageNumber="359" type="description">
<paragraph id="EA171C6EFFA3FFDD769337F2FDDBE6BB" blockId="8.[151,1437,973,1358]" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD769337F2FEC9E1C7" bold="true" box="[151,297,973,998]" pageId="8" pageNumber="359">Description.</emphasis>
Head brown; frons slightly longer than wide, anterior 1/4 of frons yellow; gena broad, 0.25 times as deep as eye. Ocellar triangle smooth, shiny, reaching anterior margin of frons, with pointed apex. Palpus largely black. Thorax black. Legs yellow except for fore and mid tarsomeres 2–5 and hind tarsomeres 3–5 black. Wing hyaline, veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 6.6: 1.4: 1. Male genitalia: Epandrium yellow, surstylus 0.7 times as long as depth of epandrium in lateral view, clavate in form, gradually widened toward apex, round distally (
<bibRefCitation id="8E39619FFFA3FFDD778730BEFDCEE6BB" author="Kanmiya, K." box="[387,558,1153,1178]" pageId="8" pageNumber="359" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA3FFDD76C3309AFDFDE69F" box="[199,541,1189,1214]" pageId="8" pageNumber="359" type="distribution">
<paragraph id="EA171C6EFFA3FFDD76C3309AFDFDE69F" blockId="8.[151,1437,973,1358]" box="[199,541,1189,1214]" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD76C3309AFE80E69F" bold="true" box="[199,352,1189,1214]" pageId="8" pageNumber="359">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFA3FFDD7763309AFE4AE69F" box="[359,426,1189,1214]" name="China" pageId="8" pageNumber="359">China</collectingCountry>
(
<collectingRegion id="286CD28CFFA3FFDD77BD309AFDF9E69F" box="[441,537,1189,1214]" country="China" name="Taiwan)" pageId="8" pageNumber="359">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA3FFDD76C330F6FBDAE76F" pageId="8" pageNumber="359" type="discussion">
<paragraph id="EA171C6EFFA3FFDD76C330F6FBDAE76F" blockId="8.[151,1437,973,1358]" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD76C330F6FEDBE6C3" bold="true" box="[199,315,1225,1250]" pageId="8" pageNumber="359">Remarks.</emphasis>
The species is somewhat similar to
<taxonomicName id="2DA867EDFFA3FFDD74DA30F5FBD9E6C3" authority="Kanmiya, 1971" authorityName="Kanmiya" authorityYear="1971" box="[734,1081,1225,1250]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="D8DCC07CFFA3FFDD74DA30F5FC60E6C0" box="[734,896,1225,1249]" italics="true" pageId="8" pageNumber="359">D. nartshukae</emphasis>
<bibRefCitation id="8E39619FFFA3FFDD758C30F6FBD9E6C3" author="Kanmiya, K." box="[904,1081,1225,1250]" pageId="8" pageNumber="359" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the following features: gena 0.25 times as deep as eye in lateral view; palpus largely black; surstylus clavated in lateral view. In
<taxonomicName id="2DA867EDFFA3FFDD77D4312DFD90E708" authorityName="Kanmiya" authorityYear="1971" box="[464,624,1297,1321]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="D8DCC07CFFA3FFDD77D4312DFD90E708" box="[464,624,1297,1321]" italics="true" pageId="8" pageNumber="359">D. nartshukae</emphasis>
</taxonomicName>
, the gena is 0.3–0.4 times as deep as eye in lateral view; the palpus is entirely yellow; and the surstylus is nearly parallel in lateral view (
<bibRefCitation id="8E39619FFFA3FFDD7586310AFBCDE76F" author="Kanmiya, K." box="[898,1069,1333,1358]" pageId="8" pageNumber="359" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

686
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<paragraph id="EA171C6EFFA4FFDA7693326AFDB0E44E" blockId="15.[151,592,1621,1680]" box="[151,592,1621,1647]" pageId="15" pageNumber="366">
<heading id="B15FAB02FFA4FFDA7693326AFDB0E44E" bold="true" box="[151,592,1621,1647]" fontSize="11" level="1" pageId="15" pageNumber="366" reason="1">
<taxonomicName id="2DA867EDFFA4FFDA7693326AFDB0E44E" ID-CoL="8BSFM" authority="Stackelberg, 1955" authorityName="Stackelberg. Scale" authorityYear="1955" box="[151,592,1621,1647]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="rossicus">
<emphasis id="D8DCC07CFFA4FFDA7693326AFDB0E44E" bold="true" box="[151,592,1621,1647]" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA7693326AFE8DE44E" bold="true" box="[151,365,1621,1647]" italics="true" pageId="15" pageNumber="366">Dicraeus rossicus</emphasis>
<bibRefCitation id="8E39619FFFA4FFDA7770326AFDB0E44E" author="Stackelberg, A. A." box="[372,592,1621,1647]" pageId="15" pageNumber="366" pagination="328 - 333" refId="ref13003" refString="Stackelberg, A. A. (1955) New species of flies of the group Acalyptrata (Diptera) from the Leningrad province. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, 18, 328 - 333. [In Russian]" type="journal article" year="1955">Stackelberg, 1955</bibRefCitation>
</emphasis>
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<paragraph id="EA171C6EFFA4FFDA76933248FEDDE4B1" blockId="15.[151,592,1621,1680]" box="[151,317,1655,1680]" pageId="15" pageNumber="366">
<figureCitation id="729300EBFFA4FFDA76933248FF0BE4B1" box="[151,235,1655,1680]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="15" pageNumber="366">Figs 11</figureCitation>
,
<figureCitation id="729300EBFFA4FFDA76F33247FEF5E4B1" box="[247,277,1656,1680]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="15" pageNumber="366">37</figureCitation>
,
<figureCitation id="729300EBFFA4FFDA77253247FEDDE4B1" box="[289,317,1656,1680]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="15" pageNumber="366">43</figureCitation>
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<paragraph id="EA171C6EFFA4FFDA769332FEFBEEE4F7" blockId="15.[151,1038,1728,1750]" box="[151,1038,1728,1750]" pageId="15" pageNumber="366">
<treatmentCitationGroup id="CAB83B40FFA4FFDA769332FEFBEEE4F7" box="[151,1038,1728,1750]" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA769332FEFF1EE4F7" box="[151,254,1729,1750]" italics="true" pageId="15" pageNumber="366">Discraeus</emphasis>
<taxonomicName id="2DA867EDFFA4FFDA770032FEFDA8E4F7" ID-CoL="8BSFM" authority="Stackelberg, 1955: 332" authorityName="Stackelberg. Scale" authorityPageNumber="332" authorityYear="1955" box="[260,584,1728,1750]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="rossicus">
<emphasis id="D8DCC07CFFA4FFDA770032FEFEB6E4F7" box="[260,342,1729,1750]" italics="true" pageId="15" pageNumber="366">rossicus</emphasis>
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<bibRefCitation id="8E39619FFFA4FFDA775832FFFDF5E4F7" author="Stackelberg, A. A." box="[348,533,1728,1750]" pageId="15" pageNumber="366" pagination="328 - 333" refId="ref13003" refString="Stackelberg, A. A. (1955) New species of flies of the group Acalyptrata (Diptera) from the Leningrad province. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, 18, 328 - 333. [In Russian]" type="journal article" year="1955">Stackelberg, 1955</bibRefCitation>
: 332
</treatmentCitation>
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.
<typeStatus id="3513A2CCFFA4FFDA745632FEFD64E4F7" box="[594,644,1729,1750]" pageId="15" pageNumber="366">Type</typeStatus>
locality:
<collectingCountry id="92BF5CFEFFA4FFDA74E032FFFCC8E4F7" box="[740,808,1728,1750]" name="Russia" pageId="15" pageNumber="366">Russia</collectingCountry>
(
<collectingRegion id="286CD28CFFA4FFDA753032FFFBE3E4F7" box="[820,1027,1728,1750]" country="Russia" name="Leningrad" pageId="15" pageNumber="366">Leningrad Province</collectingRegion>
).
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<subSubSection id="A2B24FE5FFA4FFC47693333DFE31E119" lastPageId="17" lastPageNumber="368" pageId="15" pageNumber="366" type="description">
<paragraph id="EA171C6EFFA4FFC57693333DFAF3E31E" blockId="15.[151,1437,1794,2035]" lastBlockId="16.[151,1436,151,357]" lastPageId="16" lastPageNumber="367" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA7693333DFEC9E53A" bold="true" box="[151,297,1794,1819]" pageId="15" pageNumber="366">Description.</emphasis>
Head black (
<figureCitation id="729300EBFFA4FFDA77B8333DFDEDE53A" box="[444,525,1794,1819]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="15" pageNumber="366">Fig. 11</figureCitation>
), 0.85 times as long as high in profile; frons brown with distal 1/2 yellow, as long as wide; gena yellow, broad, 0.7 times as wide as first flagellomere; ocellar triangle black, reaching anterior 2/3 of frons, with pointed apex. Antenna yellow except for dorsal margin of first flagellomere black; first flagellomere 0.9 times as long as wide; arista brown with short brown pubescence. Proboscis brown to yellow, palpus brown. Scutum black, 1.1 times as long as wide. Thoracic pleuron shiny blackish brown. Scutellum black, 0.5 times as long as wide. Coxae and trochanters yellow, femora black except for distal 1/3 of fore femur, distal 1/4 of mid femur and distal portion of hind femur yellow, fore and mid tibiae yellow, hind tibia black with both ends yellow, tarsus yellow with fore tarsomeres 4–5, mid and hind tarsomeres 3–5 brown. Wing hyaline, 2.5 times as long as wide; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 6: 2: 1. Halter pale yellow. Abdomen brown except for tergite 1 yellow with a pair of brown apico-lateral spots. Male genitalia (
<figureCitation id="729300EBFFBBFFC5728034E0FB00E2D9" box="[1156,1248,223,248]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="16" pageNumber="367">Figs. 37</figureCitation>
,
<figureCitation id="729300EBFFBBFFC572E834DFFAECE2D9" box="[1260,1292,224,248]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="16" pageNumber="367">43</figureCitation>
): Epandrium brown, height 1.3 times width in profile; cercus narrowed, finger-like; surstylus as long as epandrium, distal 1/4 suddenly widened, round and narrow apically, with some long setulae on inside of distal widened part.
</paragraph>
<paragraph id="EA171C6EFFBBFFC576C33573FACAE344" blockId="16.[151,1436,151,357]" box="[199,1322,332,357]" pageId="16" pageNumber="367">
Female. Body length
<quantity id="2D50B18BFFBBFFC577BC3573FDA1E345" box="[440,577,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.4" metricValueMin="2.8" pageId="16" pageNumber="367" unit="mm" value="3.0999999999999996" valueMax="3.4" valueMin="2.8">2.8–3.4 mm</quantity>
, wing length
<quantity id="2D50B18BFFBBFFC574DD3573FC83E345" box="[729,867,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="16" pageNumber="367" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3–2.8 mm</quantity>
. External characteristics similar to male.
</paragraph>
<caption id="BED74CE6FFBBFFC576933399FA7CE5FA" httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="16" pageNumber="367" startId="16.[151,264,1958,1980]" targetBox="[189,1395,470,1879]" targetPageId="16">
<paragraph id="EA171C6EFFBBFFC576933399FA7CE5FA" blockId="16.[151,1436,1958,2011]" pageId="16" pageNumber="367">
<emphasis id="D8DCC07CFFBBFFC576933399FEB3E59D" bold="true" box="[151,339,1958,1980]" pageId="16" pageNumber="367">FIGURES 38–43.</emphasis>
<taxonomicName id="2DA867EDFFBBFFC5775D3398FE56E59D" authorityName="Loew" authorityYear="1873" box="[345,438,1959,1980]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFBBFFC5775D3398FE56E59D" box="[345,438,1959,1980]" italics="true" pageId="16" pageNumber="367">Dicraeus</emphasis>
</taxonomicName>
species, male. epandrium, lateral view. 38
<taxonomicName id="2DA867EDFFBBFFC575633398FBBEE59D" authority="Kanmiya" authorityName="Kanmiya" authorityYear="1971" box="[871,1118,1958,1980]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="D8DCC07CFFBBFFC575633398FC18E59D" box="[871,1016,1958,1980]" italics="true" pageId="16" pageNumber="367">D. fuscipennis</emphasis>
Kanmiya
</taxonomicName>
; 39
<taxonomicName id="2DA867EDFFBBFFC572833398FABAE59D" authority="Nartshuk" authorityName="Nartshuk" box="[1159,1370,1958,1980]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="malaise">
<emphasis id="D8DCC07CFFBBFFC572833398FB14E59D" box="[1159,1268,1958,1980]" italics="true" pageId="16" pageNumber="367">D. malaise</emphasis>
Nartshuk
</taxonomicName>
; 40
<taxonomicName id="2DA867EDFFBBFFC573873398FE8DE5FA" authority="Kanmiya" authorityName="Kanmiya" authorityYear="1971" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="D8DCC07CFFBBFFC573873398FEE8E5FA" italics="true" pageId="16" pageNumber="367">D. nartshukae</emphasis>
Kanmiya
</taxonomicName>
; 41
<taxonomicName id="2DA867EDFFBBFFC5779333F9FD88E5FA" authority="Becker" authorityName="Becker" authorityYear="1911" box="[407,616,1989,2011]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="D8DCC07CFFBBFFC5779333F9FDF6E5FA" box="[407,534,1989,2011]" italics="true" pageId="16" pageNumber="367">D. orientalis</emphasis>
Becker
</taxonomicName>
; 42
<taxonomicName id="2DA867EDFFBBFFC5749533F9FC4DE5FA" authority="Kanmiya" authorityName="Kanmiya" authorityYear="1971" box="[657,941,1989,2011]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="phyllostachyus">
<emphasis id="D8DCC07CFFBBFFC5749533F9FCA7E5FA" box="[657,839,1989,2011]" italics="true" pageId="16" pageNumber="367">D. phyllostachyus</emphasis>
Kanmiya
</taxonomicName>
; 43
<taxonomicName id="2DA867EDFFBBFFC575D233F9FAE7E5FA" authority="Stackelberg. Scale" authorityName="Stackelberg. Scale" authorityYear="1955" box="[982,1287,1989,2011]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="rossicus">
<emphasis id="D8DCC07CFFBBFFC575D233F9FC0FE5FA" box="[982,1007,1990,2011]" italics="true" pageId="16" pageNumber="367">D.</emphasis>
<emphasis id="D8DCC07CFFBBFFC575F133F9FBA7E5FA" box="[1013,1095,1990,2011]" italics="true" pageId="16" pageNumber="367">rossicus</emphasis>
Stackelberg. Scale
</taxonomicName>
bar = 0.1 mm.
</paragraph>
</caption>
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<emphasis id="D8DCC07CFFBAFFC476C334A8FE26E291" bold="true" box="[199,454,151,176]" pageId="17" pageNumber="368">Specimens examined.</emphasis>
1
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, Dakang Zhou Leg. (
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<collectingDate id="8E52C346FFBAFFC4750D3483FC4BE2F5" box="[777,939,188,212]" pageId="17" pageNumber="368" value="2003-07-23">23. VII. 2003</collectingDate>
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,
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<collectingDate id="8E52C346FFBAFFC474B034DFFCA4E2D9" box="[692,836,223,248]" pageId="17" pageNumber="368" value="2006-07-09">9. VII. 2006</collectingDate>
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, Xingyue Liu Leg. (
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,
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<collectingDate id="8E52C346FFBAFFC4742E353AFD5CE33D" box="[554,700,260,284]" pageId="17" pageNumber="368" value="2008-07-07">7. VII. 2008</collectingDate>
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(
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,
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,
<date id="9E163AAEFFBAFFC477F33517FD9DE361" box="[503,637,296,320]" pageId="17" pageNumber="368" value="2009-06-02">
<collectingDate id="8E52C346FFBAFFC477F33517FD9DE361" box="[503,637,296,320]" pageId="17" pageNumber="368" value="2009-06-02">2. VI. 2009</collectingDate>
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, Qifei Liu Leg. (
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) (photo and abdomen dissected); 1
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<collectingDate id="8E52C346FFBAFFC474AA3573FCA3E345" box="[686,835,332,357]" pageId="17" pageNumber="368" value="2009-06-25">25. VI. 2009</collectingDate>
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(
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♀♀
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, Sipei Liu Leg. (
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). All specimens were stored in 75% ethanol
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.
</paragraph>
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<subSubSection id="A2B24FE5FFBAFFC476C3377BFE0FE185" pageId="17" pageNumber="368" type="distribution">
<paragraph id="EA171C6EFFBAFFC476C3377BFE0FE185" blockId="17.[151,1437,151,933]" pageId="17" pageNumber="368">
<emphasis id="D8DCC07CFFBAFFC476C3377BFE80E17C" bold="true" box="[199,352,836,861]" pageId="17" pageNumber="368">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFBAFFC4776A377BFE51E17C" box="[366,433,836,861]" name="China" pageId="17" pageNumber="368">China</collectingCountry>
(
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,
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,
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,
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,
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,
<collectingRegion id="286CD28CFFBAFFC476933757FF03E1A1" box="[151,227,872,896]" country="China" name="Gansu" pageId="17" pageNumber="368">Gansu</collectingRegion>
);
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;
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(from European part to Far East);
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; Czech and Slovak Republics;
<collectingCountry id="92BF5CFEFFBAFFC472D83758FAD4E1A1" box="[1244,1332,871,896]" name="Estonia" pageId="17" pageNumber="368">Estonia</collectingCountry>
;
<collectingCountry id="92BF5CFEFFBAFFC4733B3758FA78E1A1" box="[1343,1432,871,896]" name="Finland" pageId="17" pageNumber="368">Finland</collectingCountry>
;
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;
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,
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.
</paragraph>
</subSubSection>
<caption id="BED74CE6FFBAFFC4769333F4FD1AE5C0" box="[151,762,1995,2017]" httpUri="https://zenodo.org/record/2623736/files/figure.png" pageId="17" pageNumber="368" startId="17.[151,250,1995,2017]" targetBox="[239,1342,1015,1957]" targetPageId="17">
<paragraph id="EA171C6EFFBAFFC4769333F4FD1AE5C0" blockId="17.[151,762,1995,2017]" box="[151,762,1995,2017]" pageId="17" pageNumber="368">
<emphasis id="D8DCC07CFFBAFFC4769333F4FEFFE5C0" bold="true" box="[151,287,1995,2017]" pageId="17" pageNumber="368">FIGURE 44.</emphasis>
The distribution of
<taxonomicName id="2DA867EDFFBAFFC477EE33F3FDA7E5C0" authorityName="Loew" authorityYear="1873" box="[490,583,1996,2017]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFBAFFC477EE33F3FDA7E5C0" box="[490,583,1996,2017]" italics="true" pageId="17" pageNumber="368">Dicraeus</emphasis>
</taxonomicName>
species in China.
</paragraph>
</caption>
<subSubSection id="A2B24FE5FFB9FFC776C334A8FD7BE33C" pageId="18" pageNumber="369" type="discussion">
<paragraph id="EA171C6EFFB9FFC776C334A8FD7BE33C" blockId="18.[151,1437,151,285]" pageId="18" pageNumber="369">
<emphasis id="D8DCC07CFFB9FFC776C334A8FEDBE291" bold="true" box="[199,315,151,176]" pageId="18" pageNumber="369">Remarks.</emphasis>
The species is somewhat similar to
<taxonomicName id="2DA867EDFFB9FFC774DD34A6FBDEE291" authority="Nartshuk, 1967" authorityName="Nartshuk" authorityYear="1967" box="[729,1086,151,176]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="369" phylum="Arthropoda" rank="species" species="stackelbergi">
<emphasis id="D8DCC07CFFB9FFC774DD34A6FC65E291" box="[729,901,152,176]" italics="true" pageId="18" pageNumber="369">D. stackelbergi</emphasis>
<bibRefCitation id="8E39619FFFB9FFC7758934A8FBDEE291" author="Nartshuk, E. P." box="[909,1086,151,176]" pageId="18" pageNumber="369" pagination="245 - 256" refId="ref12347" refString="Nartshuk, E. P. (1967) Chloropid flies of the genus Dicraeus Lw. (Diptera, Chloropidae). Entomologicheskoe Obozrenie, 46, 415 - 438. [In Russian, English translation in Entomological Review, 46, 245 - 256]" type="journal article" year="1967">Nartshuk, 1967</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the surstylus as long as epandrium in lateral view, with some long setulae on inside of distal widened part. In
<taxonomicName id="2DA867EDFFB9FFC776B234DEFE81E2D9" authorityName="Nartshuk" authorityYear="1967" box="[182,353,224,248]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="369" phylum="Arthropoda" rank="species" species="stackelbergi">
<emphasis id="D8DCC07CFFB9FFC776B234DEFE81E2D9" box="[182,353,224,248]" italics="true" pageId="18" pageNumber="369">D. stackelbergi</emphasis>
</taxonomicName>
, the surstylus is distinctly shorter than epandrium in lateral view, with some thickened setae on inside of distal widened part (
<bibRefCitation id="8E39619FFFB9FFC777E6353BFD6EE33C" author="Nartshuk, E. P." box="[482,654,260,285]" pageId="18" pageNumber="369" pagination="245 - 256" refId="ref12347" refString="Nartshuk, E. P. (1967) Chloropid flies of the genus Dicraeus Lw. (Diptera, Chloropidae). Entomologicheskoe Obozrenie, 46, 415 - 438. [In Russian, English translation in Entomological Review, 46, 245 - 256]" type="journal article" year="1967">Nartshuk 1967</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

214
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<mods:namePart id="D90A4A248E96BCACBAB6A63E97CD570A">Yang, Ding</mods:namePart>
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<mods:namePart id="81D6421110C0FADF622C0760663E00C2">Nartshuk, Emilia P.</mods:namePart>
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<treatment id="6201AD78FFA4FFDA769335E3FAE1E423" ID-DOI="http://doi.org/10.5281/zenodo.5942675" ID-GBIF-Taxon="154532343" ID-Zenodo-Dep="5942675" LSID="urn:lsid:plazi:treatment:6201AD78FFA4FFDA769335E3FAE1E423" httpUri="http://treatment.plazi.org/id/6201AD78FFA4FFDA769335E3FAE1E423" lastPageNumber="366" pageId="15" pageNumber="366">
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<paragraph id="EA171C6EFFA4FFDA769335E3FD62E3D7" blockId="15.[151,642,476,535]" box="[151,642,476,502]" pageId="15" pageNumber="366">
<heading id="B15FAB02FFA4FFDA769335E3FD62E3D7" bold="true" box="[151,642,476,502]" fontSize="11" level="1" pageId="15" pageNumber="366" reason="1">
<taxonomicName id="2DA867EDFFA4FFDA769335E3FD62E3D7" ID-CoL="35QRC" authority="Kanmiya, 1971" authorityName="Kanmiya" authorityYear="1971" box="[151,642,476,502]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="phyllostachyus">
<emphasis id="D8DCC07CFFA4FFDA769335E3FD62E3D7" bold="true" box="[151,642,476,502]" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA769335E3FE5EE3D7" bold="true" box="[151,446,476,502]" italics="true" pageId="15" pageNumber="366">Dicraeus phyllostachyus</emphasis>
<bibRefCitation id="8E39619FFFA4FFDA77C135E3FD62E3D7" author="Kanmiya, K." box="[453,642,476,502]" pageId="15" pageNumber="366" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="A2B24FE5FFA4FFDA769335C1FEDEE036" box="[151,318,510,535]" pageId="15" pageNumber="366" type="description">
<paragraph id="EA171C6EFFA4FFDA769335C1FEDEE036" blockId="15.[151,642,476,535]" box="[151,318,510,535]" pageId="15" pageNumber="366">
<figureCitation id="729300EBFFA4FFDA769335C1FF0CE036" box="[151,236,510,535]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="15" pageNumber="366">Figs 10</figureCitation>
,
<figureCitation id="729300EBFFA4FFDA76FC35C1FEF6E036" box="[248,278,510,535]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="15" pageNumber="366">36</figureCitation>
,
<figureCitation id="729300EBFFA4FFDA772635C0FEDEE036" box="[290,318,511,535]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="15" pageNumber="366">42</figureCitation>
</paragraph>
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<paragraph id="EA171C6EFFA4FFDA76933677FCA5E07D" blockId="15.[151,837,582,605]" box="[151,837,582,605]" pageId="15" pageNumber="366">
<treatmentCitationGroup id="CAB83B40FFA4FFDA76933677FCA5E07D" box="[151,837,582,605]" pageId="15" pageNumber="366">
<taxonomicName id="2DA867EDFFA4FFDA76933677FD89E07D" ID-CoL="35QRC" authority="Kanmiya, 1971: 166" authorityName="Kanmiya" authorityPageNumber="166" authorityYear="1971" box="[151,617,582,605]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="phyllostachyus">
<emphasis id="D8DCC07CFFA4FFDA76933677FE70E07D" box="[151,400,582,605]" italics="true" pageId="15" pageNumber="366">Dicraeus phyllostachyus</emphasis>
<treatmentCitation id="6B093A7FFFA4FFDA77933679FD89E07D" author="Kanmiya, K." box="[407,617,582,604]" page="166" pageId="15" pageNumber="366" year="1971">
<bibRefCitation id="8E39619FFFA4FFDA77933679FDD7E07D" author="Kanmiya, K." box="[407,567,582,604]" pageId="15" pageNumber="366" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
: 166
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</taxonomicName>
.
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locality:
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.
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<paragraph id="EA171C6EFFA4FFDA769336B6FED3E673" blockId="15.[151,1437,649,1538]" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA769336B6FEC9E083" bold="true" box="[151,297,649,674]" pageId="15" pageNumber="366">Description.</emphasis>
Head brown (
<figureCitation id="729300EBFFA4FFDA77CD36B6FDFCE083" box="[457,540,649,674]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="15" pageNumber="366">Fig. 10</figureCitation>
), 0.8 times as long as high in profile; distal 1/4 of frons yellow, frons as long as wide; gena yellow, broad, 0.7 times as wide as first flagellomere; ocellar triangle with pruinescence, distal 1/4 yellow, reaching anterior margin of frons, with pointed apex. Antenna yellow except for dorsal portion of first flagellomere brown; first flagellomere as long as wide; arista brown with short brown pubescence. Proboscis yellow, palpus yellowish brown. Scutum black with brownish pruinescence, 1.1 times as long as wide. Thoracic pleuron shiny brown with a longitudinal yellow stripe running from posterior angle of postpronotal lobe to wing base; ventral 1/2 of katepisternum black. Scutellum brown, 0.6 times as long as wide. Legs yellow except for hind femur and tibia brownish yellow on anterior surface, tarsomeres 4–5 brown. Wing hyaline, 2.6 times as long as wide; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 6.6: 2: 1. Halter pale yellow. Abdomen brown except for tergite 1 yellow but brown laterally. Male genitalia (
<figureCitation id="729300EBFFA4FFDA75B437F2FBEEE1C7" box="[944,1038,973,998]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="15" pageNumber="366">Figs. 36</figureCitation>
,
<figureCitation id="729300EBFFA4FFDA721F37F1FBDCE1C7" box="[1051,1084,974,998]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="15" pageNumber="366">42</figureCitation>
): Epandrium brown, height 1.7 times width in profile; cercus broad, as long as wide; surstylus much shorter than epandrium in profile, gently broadened basally, suddenly narrowed at middle and curved in a S-shaped form, with some thick setae on inside of distal portion.
</paragraph>
<paragraph id="EA171C6EFFA4FFDA76C33062FD09E657" blockId="15.[151,1437,649,1538]" box="[199,745,1117,1142]" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA76C33062FEC5E657" bold="true" box="[199,293,1117,1142]" pageId="15" pageNumber="366">Female.</emphasis>
External characteristics similar to male.
</paragraph>
<paragraph id="EA171C6EFFA4FFDA76C330BEFDA5E70B" blockId="15.[151,1437,649,1538]" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA76C330BEFE29E6BB" bold="true" box="[199,457,1153,1178]" pageId="15" pageNumber="366">Specimens examined.</emphasis>
<materialsCitation id="5AC01633FFA4FFDA77D630BDFDA5E70B" ID-GBIF-Occurrence="2234214302" collectingDate="1974-10-27" collectingDateMax="2014-08-16" collectingDateMin="1974-10-27" collectionCode="CAU, HZAU" country="China" county="Wuyishan Mountain" location="Emeishan Mountain" municipality="Dazhulan" pageId="15" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Fujian">
<specimenCount id="FCAED7E7FFA4FFDA77D630BDFE1FE6B8" box="[466,511,1154,1178]" pageId="15" pageNumber="366" type="female">1 ♀</specimenCount>
,
<collectingCountry id="92BF5CFEFFA4FFDA741430BDFD8CE6BB" box="[528,620,1154,1178]" name="China" pageId="15" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA4FFDA747D30BEFD23E6BB" box="[633,707,1153,1178]" country="China" name="Fujian" pageId="15" pageNumber="366">Fujian</collectingRegion>
,
<collectingCounty id="037664E2FFA4FFDA74D630BEFC5CE6BB" box="[722,956,1153,1178]" pageId="15" pageNumber="366">Wuyishan Mountain</collectingCounty>
,
<collectingMunicipality id="0A738614FFA4FFDA75CD30BEFBD7E6BB" box="[969,1079,1153,1178]" pageId="15" pageNumber="366">Dazhulan</collectingMunicipality>
,
<date id="9E163AAEFFA4FFDA724130BDFB32E6BB" box="[1093,1234,1154,1178]" pageId="15" pageNumber="366" value="1974-10-27">
<collectingDate id="8E52C346FFA4FFDA724130BDFB32E6BB" box="[1093,1234,1154,1178]" pageId="15" pageNumber="366" value="1974-10-27">27. X. 1974</collectingDate>
</date>
, Fasheng Li Leg. (
<collectionCode id="8CB984ABFFA4FFDA769B3099FF00E69F" box="[159,224,1190,1214]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="15" pageNumber="366">CAU</collectionCode>
); 2
<emphasis id="D8DCC07CFFA4FFDA770A309AFED8E69C" box="[270,312,1189,1213]" italics="true" pageId="15" pageNumber="366">66</emphasis>
,
<collectingCountry id="92BF5CFEFFA4FFDA774C3099FE44E69F" box="[328,420,1190,1214]" name="China" pageId="15" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA4FFDA77B6309AFDEFE69F" box="[434,527,1189,1214]" country="China" name="Sichuan" pageId="15" pageNumber="366">Sichuan</collectingRegion>
,
<location id="EF774AB5FFA4FFDA7419309AFCE5E69F" LSID="urn:lsid:plazi:treatment:6201AD78FFA4FFDA769335E3FAE1E423:EF774AB5FFA4FFDA7419309AFCE5E69F" box="[541,773,1189,1214]" country="China" county="Wuyishan Mountain" municipality="Dazhulan" name="Emeishan Mountain" pageId="15" pageNumber="366" stateProvince="Fujian">Emeishan Mountain</location>
,
<location id="EF774AB5FFA4FFDA7517309AFBE1E69F" LSID="urn:lsid:plazi:treatment:6201AD78FFA4FFDA769335E3FAE1E423:EF774AB5FFA4FFDA7517309AFBE1E69F" box="[787,1025,1189,1214]" country="China" county="Wuyishan Mountain" municipality="Dazhulan" name="Leidongping-Jinding" pageId="15" pageNumber="366" stateProvince="Fujian">Leidongping-Jinding</location>
,
<date id="9E163AAEFFA4FFDA720A3099FB56E69F" box="[1038,1206,1190,1214]" pageId="15" pageNumber="366" value="2009-08-11">
<collectingDate id="8E52C346FFA4FFDA720A3099FB56E69F" box="[1038,1206,1190,1214]" pageId="15" pageNumber="366" value="2009-08-11">11. VIII. 2009</collectingDate>
</date>
, JunchaoWang Leg. (
<collectionCode id="8CB984ABFFA4FFDA769A30F5FF3CE6C3" box="[158,220,1226,1250]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="15" pageNumber="366">CAU</collectionCode>
) (photo and abdomen dissected); 1
<emphasis id="D8DCC07CFFA4FFDA747730F6FD68E6C0" box="[627,648,1225,1249]" italics="true" pageId="15" pageNumber="366">6</emphasis>
,
<collectingCountry id="92BF5CFEFFA4FFDA749330F5FD13E6C3" box="[663,755,1226,1250]" name="China" pageId="15" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA4FFDA74FB30F6FCBCE6C3" box="[767,860,1225,1250]" country="China" name="Shaanxi" pageId="15" pageNumber="366">Shaanxi</collectingRegion>
,
<location id="EF774AB5FFA4FFDA756C30F6FC28E6C3" LSID="urn:lsid:plazi:treatment:6201AD78FFA4FFDA769335E3FAE1E423:EF774AB5FFA4FFDA756C30F6FC28E6C3" box="[872,968,1225,1250]" country="China" county="Wuyishan Mountain" municipality="Dazhulan" name="Zhouzhi" pageId="15" pageNumber="366" stateProvince="Fujian">Zhouzhi</location>
,
<location id="EF774AB5FFA4FFDA75D130F6FBB1E6C3" LSID="urn:lsid:plazi:treatment:6201AD78FFA4FFDA769335E3FAE1E423:EF774AB5FFA4FFDA75D130F6FBB1E6C3" box="[981,1105,1225,1250]" country="China" county="Wuyishan Mountain" municipality="Dazhulan" name="Houzhenzi" pageId="15" pageNumber="366" stateProvince="Fujian">Houzhenzi</location>
,
<date id="9E163AAEFFA4FFDA725930F6FB03E6C3" box="[1117,1251,1225,1250]" pageId="15" pageNumber="366" value="2009-05-15">
<collectingDate id="8E52C346FFA4FFDA725930F6FB03E6C3" box="[1117,1251,1225,1250]" pageId="15" pageNumber="366" value="2009-05-15">15. V. 2009</collectingDate>
</date>
, Maoling Sheng Leg. (
<collectionCode id="8CB984ABFFA4FFDA76DB30D1FED3E727" box="[223,307,1262,1286]" pageId="15" pageNumber="366">HZAU</collectionCode>
); 1
<emphasis id="D8DCC07CFFA4FFDA775830D2FE91E724" box="[348,369,1261,1285]" italics="true" pageId="15" pageNumber="366">6</emphasis>
,
<collectingCountry id="92BF5CFEFFA4FFDA778230D1FE01E727" box="[390,481,1262,1286]" name="China" pageId="15" pageNumber="366">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA4FFDA77F730D2FDB0E727" box="[499,592,1261,1286]" country="China" name="Shaanxi" pageId="15" pageNumber="366">Shaanxi</collectingRegion>
,
<location id="EF774AB5FFA4FFDA746730D2FD23E727" LSID="urn:lsid:plazi:treatment:6201AD78FFA4FFDA769335E3FAE1E423:EF774AB5FFA4FFDA746730D2FD23E727" box="[611,707,1261,1286]" country="China" county="Wuyishan Mountain" municipality="Dazhulan" name="Zhouzhi" pageId="15" pageNumber="366" stateProvince="Fujian">Zhouzhi</location>
,
<location id="EF774AB5FFA4FFDA74D130D2FCB1E727" LSID="urn:lsid:plazi:treatment:6201AD78FFA4FFDA769335E3FAE1E423:EF774AB5FFA4FFDA74D130D2FCB1E727" box="[725,849,1261,1286]" country="China" county="Wuyishan Mountain" municipality="Dazhulan" name="Houzhenzi" pageId="15" pageNumber="366" stateProvince="Fujian">Houzhenzi</location>
,
<date id="9E163AAEFFA4FFDA756730D2FBF5E727" box="[867,1045,1261,1286]" pageId="15" pageNumber="366" value="2014-08-16">
<collectingDate id="8E52C346FFA4FFDA756730D2FBF5E727" box="[867,1045,1261,1286]" pageId="15" pageNumber="366" value="2014-08-16">16. VIII. 2014</collectingDate>
</date>
, Xuankun Li Leg. (
<collectionCode id="8CB984ABFFA4FFDA730F30D1FABFE727" box="[1291,1375,1262,1286]" pageId="15" pageNumber="366">HZAU</collectionCode>
).
<location id="EF774AB5FFA4FFDA737C30D2FA7CE727" LSID="urn:lsid:plazi:treatment:6201AD78FFA4FFDA769335E3FAE1E423:EF774AB5FFA4FFDA737C30D2FA7CE727" box="[1400,1436,1261,1286]" country="China" county="Wuyishan Mountain" municipality="Dazhulan" name="All" pageId="15" pageNumber="366" stateProvince="Fujian">All</location>
specimens were stored in 75% ethanol
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA4FFDA76C3310AFCD2E76C" box="[199,818,1333,1358]" pageId="15" pageNumber="366" type="distribution">
<paragraph id="EA171C6EFFA4FFDA76C3310AFCD2E76C" blockId="15.[151,1437,649,1538]" box="[199,818,1333,1358]" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA76C3310AFE80E76F" bold="true" box="[199,352,1333,1358]" pageId="15" pageNumber="366">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFA4FFDA7763310AFE4AE76F" box="[359,426,1333,1358]" name="China" pageId="15" pageNumber="366">China</collectingCountry>
(
<collectingRegion id="286CD28CFFA4FFDA77BD310AFDE5E76F" box="[441,517,1333,1358]" country="China" name="Fujian" pageId="15" pageNumber="366">Fujian</collectingRegion>
,
<collectingRegion id="286CD28CFFA4FFDA7414310AFD8DE76F" box="[528,621,1333,1358]" country="China" name="Shaanxi" pageId="15" pageNumber="366">Shaanxi</collectingRegion>
,
<collectingRegion id="286CD28CFFA4FFDA747C310AFD37E76F" box="[632,727,1333,1358]" country="China" name="Sichuan" pageId="15" pageNumber="366">Sichuan</collectingRegion>
);
<collectingCountry id="92BF5CFEFFA4FFDA74EE3109FCCDE76C" box="[746,813,1334,1357]" name="Japan" pageId="15" pageNumber="366">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA4FFDA76C33166FAE1E423" pageId="15" pageNumber="366" type="discussion">
<paragraph id="EA171C6EFFA4FFDA76C33166FAE1E423" blockId="15.[151,1437,649,1538]" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA76C33166FEDBE753" bold="true" box="[199,315,1369,1394]" pageId="15" pageNumber="366">Remarks.</emphasis>
The new species is somewhat similar to
<taxonomicName id="2DA867EDFFA4FFDA75143165FBDFE753" authority="Becker, 1911" authorityName="Becker" authorityYear="1911" box="[784,1087,1369,1394]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="D8DCC07CFFA4FFDA75143165FC7FE750" box="[784,927,1369,1393]" italics="true" pageId="15" pageNumber="366">D. orientalis</emphasis>
<bibRefCitation id="8E39619FFFA4FFDA75A33166FBDFE753" author="Becker, T." box="[935,1087,1369,1394]" pageId="15" pageNumber="366" pagination="35 - 170" refId="ref11590" refString="Becker, T. (1911) Chloropidae. Eine monographische Studie. iii. Die indo-australische Region. Annales Historico-Naturales Musei Nationalis Hungarici, 9, 35 - 170." type="journal article" year="1911">Becker, 1911</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the ocellar triangle reaching the anterior margin of frons, antenna yellow except for the first flagellomere partly yellow, and surstylus gently broadened basally, suddenly narrowed at middle and curved in a S-like form. In
<taxonomicName id="2DA867EDFFA4FFDA769331F9FEC5E7FC" authorityName="Becker" authorityYear="1911" box="[151,293,1477,1501]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="D8DCC07CFFA4FFDA769331F9FEC5E7FC" box="[151,293,1477,1501]" italics="true" pageId="15" pageNumber="366">D. orientalis</emphasis>
</taxonomicName>
, the ocellar triangle reaches anterior 3/4 of frons; the antenna is entirely black; and the surstylus has two lobes, the dorsal one short and broadened, the ventral one elongate and slender (
<bibRefCitation id="8E39619FFFA4FFDA724C31D6FB14E423" author="Kanmiya, K." box="[1096,1268,1513,1538]" pageId="15" pageNumber="366" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

181
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<treatment id="6201AD78FFA6FFD87693368BFBEEE491" ID-DOI="http://doi.org/10.5281/zenodo.5942671" ID-GBIF-Taxon="154532342" ID-Zenodo-Dep="5942671" LSID="urn:lsid:plazi:treatment:6201AD78FFA6FFD87693368BFBEEE491" httpUri="http://treatment.plazi.org/id/6201AD78FFA6FFD87693368BFBEEE491" lastPageNumber="364" pageId="13" pageNumber="364">
<subSubSection id="A2B24FE5FFA6FFD87693368BFDB8E0EF" box="[151,600,692,718]" pageId="13" pageNumber="364" type="nomenclature">
<paragraph id="EA171C6EFFA6FFD87693368BFDB8E0EF" blockId="13.[151,600,692,751]" box="[151,600,692,718]" pageId="13" pageNumber="364">
<heading id="B15FAB02FFA6FFD87693368BFDB8E0EF" bold="true" box="[151,600,692,718]" fontSize="11" level="1" pageId="13" pageNumber="364" reason="1">
<taxonomicName id="2DA867EDFFA6FFD87693368BFDB8E0EF" ID-CoL="35QQX" authority="Kanmiya, 1971" authorityName="Kanmiya" authorityYear="1971" box="[151,600,692,718]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="D8DCC07CFFA6FFD87693368BFDB8E0EF" bold="true" box="[151,600,692,718]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD87693368BFE73E0EF" bold="true" box="[151,403,692,718]" italics="true" pageId="13" pageNumber="364">Dicraeus nartshukae</emphasis>
<bibRefCitation id="8E39619FFFA6FFD8779F368BFDB8E0EF" author="Kanmiya, K." box="[411,600,692,718]" pageId="13" pageNumber="364" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="A2B24FE5FFA6FFD8769336E9FCB1E115" pageId="13" pageNumber="364" type="description">
<paragraph id="EA171C6EFFA6FFD8769336E9FED0E0CE" blockId="13.[151,600,692,751]" box="[151,304,726,751]" pageId="13" pageNumber="364">
<figureCitation id="729300EBFFA6FFD8769336E9FF3DE0CE" box="[151,221,726,751]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="13" pageNumber="364">Figs 8</figureCitation>
,
<figureCitation id="729300EBFFA6FFD876EE36E8FEE9E0CE" box="[234,265,727,751]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="13" pageNumber="364">34</figureCitation>
,
<figureCitation id="729300EBFFA6FFD8771036E8FED0E0CE" box="[276,304,727,751]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="13" pageNumber="364">40</figureCitation>
</paragraph>
<paragraph id="EA171C6EFFA6FFD87693371FFCB1E115" blockId="13.[151,849,798,851]" box="[151,849,798,821]" pageId="13" pageNumber="364">
<taxonomicName id="2DA867EDFFA6FFD87693371FFE4CE115" authority="Nartshuk" authorityName="Nartshuk" box="[151,428,798,821]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="D8DCC07CFFA6FFD87693371FFEA9E115" box="[151,329,798,821]" italics="true" pageId="13" pageNumber="364">Dicraeus flavipes</emphasis>
Nartshuk
</taxonomicName>
nec Duda—misidentification. 1963: 108.
</paragraph>
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<subSubSection id="A2B24FE5FFA6FFD876933701FCC0E172" box="[151,800,829,851]" pageId="13" pageNumber="364" type="reference_group">
<paragraph id="EA171C6EFFA6FFD876933701FCC0E172" blockId="13.[151,849,798,851]" box="[151,800,829,851]" pageId="13" pageNumber="364">
<treatmentCitationGroup id="CAB83B40FFA6FFD876933701FCC0E172" box="[151,800,829,851]" pageId="13" pageNumber="364">
<taxonomicName id="2DA867EDFFA6FFD876933701FDA3E172" ID-CoL="35QQX" authority="Kanmiya, 1971: 169" authorityName="Kanmiya" authorityPageNumber="169" authorityYear="1971" box="[151,579,829,851]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="D8DCC07CFFA6FFD876933701FE8AE172" box="[151,362,829,851]" italics="true" pageId="13" pageNumber="364">Dicraeus nartshukae</emphasis>
<treatmentCitation id="6B093A7FFFA6FFD877753702FDA3E172" author="Kanmiya, K." box="[369,579,829,851]" page="169" pageId="13" pageNumber="364" year="1971">
<bibRefCitation id="8E39619FFFA6FFD877753702FDF0E172" author="Kanmiya, K." box="[369,528,829,851]" pageId="13" pageNumber="364" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
: 169
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="3513A2CCFFA6FFD874493701FD9FE172" box="[589,639,830,851]" pageId="13" pageNumber="364">Type</typeStatus>
locality:
<collectingCountry id="92BF5CFEFFA6FFD874DB3701FCFCE172" box="[735,796,830,851]" name="Japan" pageId="13" pageNumber="364">Japan</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA6FFD876933740FBD7E7F9" pageId="13" pageNumber="364" type="description">
<paragraph id="EA171C6EFFA6FFD876933740FEF9E74D" blockId="13.[151,1437,895,1712]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD876933740FEC9E1B9" bold="true" box="[151,297,895,920]" pageId="13" pageNumber="364">Description.</emphasis>
Head brown (
<figureCitation id="729300EBFFA6FFD877D43740FDF7E1B9" box="[464,535,895,920]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="13" pageNumber="364">Fig. 8</figureCitation>
), 0.9 times as long as high in profile; face yellow; frons blackish brown with distal 1/3 yellow, as long as wide; gena yellow, broad, 0.5 times as wide as first flagellomere. Parafacial yellow; ocellar triangle smooth, shiny, distal 1/3 yellow, reaching anterior margin of frons, with pointed apex. Antenna yellow except for dorsal 1/3 of first flagellomere brown; first flagellomere 0.8 times as long as wide; arista brown with short brown pubescence. Proboscis and palpus yellow. Scutum black, 1.1 times as long as wide. Thoracic pleuron shiny brown with a longitudinal yellow stripe running from posterior angle of postpronotal lobe to wing base; propleuron yellowish; anterioventral margins of anepisternum and katepisternum black; posterior portion of katepisternum and anterior portion of meron yellow. Scutellum brown, 0.6 times as long as wide. Legs yellow except for fore tarsomeres 2–5, mid and hind tarsomeres 4–5 blackish brown. Wing hyaline, 2.6 times as long as wide; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 5.4: 1.4: 1. Halter pale yellow. Abdomen brown except for tergite 1 yellow and tergites 2–5 yellow distally. Male genitalia (
<figureCitation id="729300EBFFA6FFD8722B30D8FB6CE721" box="[1071,1164,1255,1280]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="13" pageNumber="364">Figs. 34</figureCitation>
,
<figureCitation id="729300EBFFA6FFD8729C30D7FB59E721" box="[1176,1209,1256,1280]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="13" pageNumber="364">40</figureCitation>
): Epandrium yellow with many long setulae; cercus developed, 1.5 times wider than long in lateral view; surstylus slightly shorter than epandrium, nearly parallel-sided, round distally in lateral view, some thickened setae on inner side of lateral and distal parts.
</paragraph>
<paragraph id="EA171C6EFFA6FFD876C33148FD09E7B1" blockId="13.[151,1437,895,1712]" box="[199,745,1399,1424]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD876C33148FEFEE7B1" bold="true" box="[199,286,1399,1424]" pageId="13" pageNumber="364">Female</emphasis>
. External characteristics similar to male.
</paragraph>
<paragraph id="EA171C6EFFA6FFD876C331A3FBD7E7F9" blockId="13.[151,1437,895,1712]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD876C331A3FE28E794" bold="true" box="[199,456,1436,1461]" pageId="13" pageNumber="364">Specimens examined.</emphasis>
<materialsCitation id="5AC01633FFA6FFD877D531A3FBD3E7F9" ID-GBIF-Occurrence="2234214335" collectingDate="2010-07-05" collectionCode="CAU" country="China" location="Linggongli" municipality="Emeishan Mountain" pageId="13" pageNumber="364" specimenCount="3" specimenCount-male="3" stateProvince="Sichuan">
<specimenCount id="FCAED7E7FFA6FFD877D531A3FDF4E795" box="[465,532,1436,1460]" pageId="13" pageNumber="364" type="male">3 ♂♂</specimenCount>
,
<collectingCountry id="92BF5CFEFFA6FFD8742731A3FD9FE795" box="[547,639,1436,1460]" name="China" pageId="13" pageNumber="364">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA6FFD8748831A3FD09E794" box="[652,745,1436,1461]" country="China" name="Sichuan" pageId="13" pageNumber="364">Sichuan</collectingRegion>
,
<collectingMunicipality id="0A738614FFA6FFD874F231A3FC3EE794" box="[758,990,1436,1461]" pageId="13" pageNumber="364">Emeishan Mountain</collectingMunicipality>
,
<location id="EF774AB5FFA6FFD875EF31A3FB8AE794" LSID="urn:lsid:plazi:treatment:6201AD78FFA6FFD87693368BFBEEE491:EF774AB5FFA6FFD875EF31A3FB8AE794" box="[1003,1130,1436,1461]" country="China" municipality="Emeishan Mountain" name="Linggongli" pageId="13" pageNumber="364" stateProvince="Sichuan">Linggongli</location>
,
<date id="9E163AAEFFA6FFD8727331A3FAE8E795" box="[1143,1288,1436,1461]" pageId="13" pageNumber="364" value="2010-07-05">
<collectingDate id="8E52C346FFA6FFD8727331A3FAE8E795" box="[1143,1288,1436,1461]" pageId="13" pageNumber="364" value="2010-07-05">5. VII. 2010</collectingDate>
</date>
, Tao Li Leg. (
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) (photo and abdomen dissected). All specimens were stored in 75% ethanol
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA6FFD876C331DBFB22E7DD" box="[199,1218,1508,1533]" pageId="13" pageNumber="364" type="distribution">
<paragraph id="EA171C6EFFA6FFD876C331DBFB22E7DD" blockId="13.[151,1437,895,1712]" box="[199,1218,1508,1533]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD876C331DBFE80E7DC" bold="true" box="[199,352,1508,1533]" pageId="13" pageNumber="364">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFA6FFD8776331DBFE4AE7DC" box="[359,426,1508,1533]" name="China" pageId="13" pageNumber="364">China</collectingCountry>
(
<collectingRegion id="286CD28CFFA6FFD877BD31DBFDF9E7DC" box="[441,537,1508,1533]" country="China" name="Sichuan" pageId="13" pageNumber="364">Sichuan</collectingRegion>
);
<collectingCountry id="92BF5CFEFFA6FFD8742831DAFD8FE7DD" box="[556,623,1509,1532]" name="Japan" pageId="13" pageNumber="364">Japan</collectingCountry>
, Far East of
<collectingCountry id="92BF5CFEFFA6FFD874F831DBFCA7E7DC" box="[764,839,1508,1533]" name="Russia" pageId="13" pageNumber="364">Russia</collectingCountry>
(
<collectingRegion id="286CD28CFFA6FFD8755131DBFC39E7DC" box="[853,985,1508,1533]" country="Russia" name="Kamchatka" pageId="13" pageNumber="364">Kamchatka</collectingRegion>
,
<collectingRegion id="286CD28CFFA6FFD875E131DBFBAAE7DC" box="[997,1098,1508,1533]" country="Russia" name="Sakhalin" pageId="13" pageNumber="364">Sakhalin</collectingRegion>
, Kuril Is.).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA6FFD876C33238FBEEE491" pageId="13" pageNumber="364" type="discussion">
<paragraph id="EA171C6EFFA6FFD876C33238FBEEE491" blockId="13.[151,1437,895,1712]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD876C33238FEDBE401" bold="true" box="[199,315,1543,1568]" pageId="13" pageNumber="364">Remarks.</emphasis>
The new species is somewhat similar to
<taxonomicName id="2DA867EDFFA6FFD875073236FBE8E401" authority="Duda, 1930" authorityName="Duda" authorityYear="1930" box="[771,1032,1543,1568]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="D8DCC07CFFA6FFD875073236FC9DE401" box="[771,893,1544,1568]" italics="true" pageId="13" pageNumber="364">D. flavipes</emphasis>
<bibRefCitation id="8E39619FFFA6FFD875863238FBE8E401" author="Duda, O." box="[898,1032,1543,1568]" pageId="13" pageNumber="364" pagination="278 - 304" refId="ref11917" refString="Duda, O. (1930) Neue und bekannte orientalische Chloropiden (Dipt.) des Deutschen Entomologischen Instituts in Berlin- Dahlem. Stettiner entomologische Zeitung, 91, 278 - 304." type="journal article" year="1930">Duda, 1930</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the gena broad, 0.3–0.4 times as deep as eye in profile; palpus yellow; surstylus nearly parallel-sided, round distally in lateral view. In
<taxonomicName id="2DA867EDFFA6FFD877C4326EFDDBE449" authorityName="Duda" authorityYear="1930" box="[448,571,1616,1640]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="D8DCC07CFFA6FFD877C4326EFDDBE449" box="[448,571,1616,1640]" italics="true" pageId="13" pageNumber="364">D. flavipes</emphasis>
</taxonomicName>
, the gena is 0.25 times as deep as eye in profile; the palpus is largely black; and the surstylus is clavated in lateral view (
<bibRefCitation id="8E39619FFFA6FFD8748E324BFCD6E4AD" author="Kanmiya, K." box="[650,822,1652,1677]" pageId="13" pageNumber="364" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya 1971</bibRefCitation>
). The species is distributed to the North of Kamchatka and Island Karaginsky, host plant
<taxonomicName id="2DA867EDFFA6FFD8741032A6FBEAE491" authority="(Nartshuk &amp; Khruleva, 2010)" authorityName="Nartshuk &amp; Khruleva" authorityYear="2010" baseAuthorityName="Nartshuk &amp; Khruleva" baseAuthorityYear="2010" box="[532,1034,1687,1712]" class="Liliopsida" family="Poaceae" genus="Leymus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="13" pageNumber="364" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis id="D8DCC07CFFA6FFD8741032A6FD55E491" box="[532,693,1688,1712]" italics="true" pageId="13" pageNumber="364">Leymus mollis</emphasis>
(
<bibRefCitation id="8E39619FFFA6FFD874C732A8FBE2E491" author="Nartshuk, E. P. &amp; Khruleva, O. A." box="[707,1026,1687,1712]" pageId="13" pageNumber="364" pagination="654 - 656" refId="ref12710" refString="Nartshuk, E. P. &amp; Khruleva, O. A. (2010) On distribution range of the seed-eating Dicraeus nartshukae (Diptera, Chloropidae) and its host plant Leymus mollis. Zoologicheskii Zhurnal, 89 (7), 882 - 884. [In Russian, English translation in Entomological Review, 90 (5), 654 - 656]" type="journal article" year="2010">Nartshuk &amp; Khruleva, 2010</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

247
data/62/01/AD/6201AD78FFA6FFDA7693333BFF3FE3A9.xml Normal file
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<subSubSection id="A2B24FE5FFA6FFD87693333BFDC1E53F" box="[151,545,1796,1822]" pageId="13" pageNumber="364" type="nomenclature">
<paragraph id="EA171C6EFFA6FFD87693333BFDC1E53F" blockId="13.[151,545,1796,1855]" box="[151,545,1796,1822]" pageId="13" pageNumber="364">
<heading id="B15FAB02FFA6FFD87693333BFDC1E53F" bold="true" box="[151,545,1796,1822]" fontSize="11" level="1" pageId="13" pageNumber="364" reason="1">
<taxonomicName id="2DA867EDFFA6FFD87693333BFDC1E53F" ID-CoL="6D3VH" authority="Becker, 1911" authorityName="Becker" authorityYear="1911" box="[151,545,1796,1822]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="D8DCC07CFFA6FFD87693333BFDC1E53F" bold="true" box="[151,545,1796,1822]" pageId="13" pageNumber="364">
<emphasis id="D8DCC07CFFA6FFD87693333BFE9CE53F" bold="true" box="[151,380,1796,1822]" italics="true" pageId="13" pageNumber="364">Dicraeus orientalis</emphasis>
<bibRefCitation id="8E39619FFFA6FFD87780333BFDC1E53F" author="Becker, T." box="[388,545,1796,1822]" pageId="13" pageNumber="364" pagination="35 - 170" refId="ref11590" refString="Becker, T. (1911) Chloropidae. Eine monographische Studie. iii. Die indo-australische Region. Annales Historico-Naturales Musei Nationalis Hungarici, 9, 35 - 170." type="journal article" year="1911">Becker, 1911</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="A2B24FE5FFA6FFD876933319FED0E51E" box="[151,304,1830,1855]" pageId="13" pageNumber="364" type="description">
<paragraph id="EA171C6EFFA6FFD876933319FED0E51E" blockId="13.[151,545,1796,1855]" box="[151,304,1830,1855]" pageId="13" pageNumber="364">
<figureCitation id="729300EBFFA6FFD876933319FF3EE51E" box="[151,222,1830,1855]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="13" pageNumber="364">Figs 9</figureCitation>
,
<figureCitation id="729300EBFFA6FFD876EE3319FEE8E51E" box="[234,264,1830,1855]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="13" pageNumber="364">35</figureCitation>
,
<figureCitation id="729300EBFFA6FFD877103318FED0E51E" box="[276,304,1831,1855]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="13" pageNumber="364">41</figureCitation>
</paragraph>
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<paragraph id="EA171C6EFFA6FFD87693334FFC5DE5A5" blockId="13.[151,957,1902,1955]" box="[151,957,1902,1925]" pageId="13" pageNumber="364">
<treatmentCitationGroup id="CAB83B40FFA6FFD87693334FFC5DE5A5" box="[151,957,1902,1925]" pageId="13" pageNumber="364">
<taxonomicName id="2DA867EDFFA6FFD87693334FFDFCE5A5" ID-CoL="6D3VH" authority="Becker, 1911: 143" authorityName="Becker" authorityPageNumber="143" authorityYear="1911" box="[151,540,1902,1925]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="D8DCC07CFFA6FFD87693334FFEBBE5A5" box="[151,347,1902,1925]" italics="true" pageId="13" pageNumber="364">Dicraeus orientalis</emphasis>
<treatmentCitation id="6B093A7FFFA6FFD877653351FDFCE5A5" author="Becker, T." box="[353,540,1902,1924]" page="143" pageId="13" pageNumber="364" year="1911">
<bibRefCitation id="8E39619FFFA6FFD877653351FE0AE5A5" author="Becker, T." box="[353,490,1902,1924]" pageId="13" pageNumber="364" pagination="35 - 170" refId="ref11590" refString="Becker, T. (1911) Chloropidae. Eine monographische Studie. iii. Die indo-australische Region. Annales Historico-Naturales Musei Nationalis Hungarici, 9, 35 - 170." type="journal article" year="1911">Becker, 1911</bibRefCitation>
: 143
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="3513A2CCFFA6FFD874223350FDB8E5A5" box="[550,600,1903,1924]" pageId="13" pageNumber="364">Type</typeStatus>
locality:
<collectingCountry id="92BF5CFEFFA6FFD874BC3351FD18E5A5" box="[696,760,1902,1924]" name="China" pageId="13" pageNumber="364">China</collectingCountry>
:
<collectingCountry id="92BF5CFEFFA6FFD875063351FCAFE5A5" box="[770,847,1902,1924]" name="Taiwan" pageId="13" pageNumber="364">Taiwan</collectingCountry>
, Chipchip.
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA171C6EFFA6FFD8769333B2FC58E582" blockId="13.[151,957,1902,1955]" box="[151,952,1933,1955]" pageId="13" pageNumber="364">
<treatmentCitationGroup id="CAB83B40FFA6FFD8769333B2FC58E582" box="[151,952,1933,1955]" pageId="13" pageNumber="364">
<taxonomicName id="2DA867EDFFA6FFD8769333B2FDF8E582" authority="Becker, 1924: 122" authorityName="Becker" authorityPageNumber="122" authorityYear="1924" box="[151,536,1933,1955]" class="Insecta" family="Chloropidae" genus="Oscinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="364" phylum="Arthropoda" rank="species" species="giabrina">
<emphasis id="D8DCC07CFFA6FFD8769333B2FEB5E582" box="[151,341,1933,1955]" italics="true" pageId="13" pageNumber="364">Oscinella giabrina</emphasis>
<treatmentCitation id="6B093A7FFFA6FFD8775833B2FDF8E582" author="Becker, T." box="[348,536,1933,1955]" page="122" pageId="13" pageNumber="364" year="1924">
<bibRefCitation id="8E39619FFFA6FFD8775833B2FE05E582" author="Becker, T." box="[348,485,1933,1955]" pageId="13" pageNumber="364" pagination="117 - 124" refId="ref11670" refString="Becker, T. (1924) H. Sauter's Formosa-Ausbeute: Chloropidae (Diptera). Entomologische Mitteilungen, 13, 117 - 124." type="journal article" year="1924">Becker, 1924</bibRefCitation>
: 122
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="3513A2CCFFA6FFD8742633B1FDB4E582" box="[546,596,1934,1955]" pageId="13" pageNumber="364">Type</typeStatus>
locality:
<collectingCountry id="92BF5CFEFFA6FFD874B733B2FD13E582" box="[691,755,1933,1955]" name="China" pageId="13" pageNumber="364">China</collectingCountry>
:
<collectingCountry id="92BF5CFEFFA6FFD874F933B2FCAAE582" box="[765,842,1933,1955]" name="Taiwan" pageId="13" pageNumber="364">Taiwan</collectingCountry>
, Chipchip.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<emphasis id="D8DCC07CFFA6FFD8769333EFFEC9E5C8" bold="true" box="[151,297,2000,2025]" pageId="13" pageNumber="364">Description.</emphasis>
Head black (
<figureCitation id="729300EBFFA6FFD877BA33EFFDE3E5C9" box="[446,515,2000,2025]" captionStart="FIGURES 6–11" captionStartId="2.[151,264,1979,2001]" captionTargetBox="[246,1351,1402,1924]" captionTargetId="figure@2.[226,1360,1371,1958]" captionTargetPageId="2" captionText="FIGURES 6–11. Dicraeus species, habitus. 6 D. fuscipennis Kanmiya; 7 D. malaise Nartshuk; 8 D. nartshukae Kanmiya; 9 D. orientalis Becker; 10 D. Phyllostachyus Kanmiya; 11 D. rossicus Stackelberg. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623717/files/figure.png" pageId="13" pageNumber="364">Fig. 9</figureCitation>
), 0.8 times as long as high in profile; face yellow, distal 1/4 of frons yellow, frons
</paragraph>
<paragraph id="EA171C6EFFA5FFDB769334A7FEB1E041" blockId="14.[151,1437,151,789]" pageId="14" pageNumber="365">
1.4 times as long as wide; gena yellow with ventral margin black, broad, as wide as first flagellomere; ocellar triangle smooth, shiny, apex yellow, reaching anterior 3/4 of frons, with pointed apex. Antenna entirely black; first flagellomere 1.5 times as long as wide; arista black with short black pubescence. Proboscis brown, palpus yellowish brown. Scutum black, 1.3 times as long as wide. Thoracic pleuron shiny brown with a longitudinal yellow stripe running from lower 1/2 of postpronotal lobe to wing base. Scutellum black, 0.6 times as long as wide. Coxae and trochanters yellow, femora and tibiae brown except for distal tips of femora and basal tips of tibiae yellow, fore tarsus brown, mid and hind tarsi yellowish brown with tarsomeres 4–5 brown. Wing hyaline, 2.8 times as long as wide; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 6: 1.5: 1. Halter pale yellow. Abdomen brown but yellow basally. Male genitalia (
<figureCitation id="729300EBFFA5FFDB74F33588FCB7E3F1" box="[759,855,439,464]" captionStart="FIGURES 32–37" captionStartId="14.[151,264,1944,1966]" captionTargetBox="[174,1407,850,1907]" captionTargetId="figure@14.[151,1436,833,1930]" captionTargetPageId="14" captionText="FIGURES 32–37. Dicraeus species, male. epandrium, posterior view. 32 D. fuscipennis Kanmiya; 33 D. malaise Nartshuk; 34 D. nartshukae Kanmiya; 35 D. orientalis Becker; 36 D. phyllostachyus Kanmiya; 37 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="14" pageNumber="365">Figs. 35</figureCitation>
,
<figureCitation id="729300EBFFA5FFDB75623587FC66E3F1" box="[870,902,440,464]" captionStart="FIGURES 38–43" captionStartId="16.[151,264,1958,1980]" captionTargetBox="[189,1395,470,1879]" captionTargetId="figure@16.[151,1436,449,1902]" captionTargetPageId="16" captionText="FIGURES 38–43. Dicraeus species, male. epandrium, lateral view. 38 D. fuscipennis Kanmiya; 39 D. malaise Nartshuk; 40 D. nartshukae Kanmiya; 41 D. orientalis Becker; 42 D. phyllostachyus Kanmiya; 43 D. rossicus Stackelberg. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623734/files/figure.png" pageId="14" pageNumber="365">41</figureCitation>
): Epandrium brown, height 1.7 times width in lateral view; bearing a very long apical process, projected ventrally; cercus broad, as long as wide; surstylus peculiar, divided into two lobes, dorsal lobe short, broadened, with few short spinous setae on surface of middle part and with long thick setae on distal portion, ventral lobe elongate, slender, with some short setulae on inner side of distal portion.
</paragraph>
<paragraph id="EA171C6EFFA5FFDB76C33653FACAE0A4" blockId="14.[151,1437,151,789]" box="[199,1322,620,645]" pageId="14" pageNumber="365">
Female. Body length
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, wing length
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. External characteristics similar to male.
</paragraph>
<paragraph id="EA171C6EFFA5FFDB76C336B0FF06E134" blockId="14.[151,1437,151,789]" pageId="14" pageNumber="365">
<emphasis id="D8DCC07CFFA5FFDB76C336B0FE29E089" bold="true" box="[199,457,655,680]" pageId="14" pageNumber="365">Specimens examined.</emphasis>
<materialsCitation id="5AC01633FFA5FFDB77D736AFFD21E0EC" ID-GBIF-Occurrence="2234214337" collectingDate="2009-05-22" collectionCode="HZAU" country="China" location="Nanxi" municipality="Hekou" pageId="14" pageNumber="365" specimenCount="82" specimenCount-female="35" specimenCount-male="47" stateProvince="Yunnan">
<specimenCount id="FCAED7E7FFA5FFDB77D736AFFDC5E086" box="[467,549,656,680]" pageId="14" pageNumber="365" type="male">47 ♂♂</specimenCount>
,
<specimenCount id="FCAED7E7FFA5FFDB743036B0FD65E086" box="[564,645,655,680]" pageId="14" pageNumber="365" type="female">35 ♀♀</specimenCount>
,
<collectingCountry id="92BF5CFEFFA5FFDB749136AFFD11E089" box="[661,753,656,680]" name="China" pageId="14" pageNumber="365">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA5FFDB74FB36AFFCB8E086" box="[767,856,656,679]" country="China" name="Yunnan" pageId="14" pageNumber="365">Yunnan</collectingRegion>
,
<collectingMunicipality id="0A738614FFA5FFDB756136B0FC53E089" box="[869,947,655,680]" pageId="14" pageNumber="365">Hekou</collectingMunicipality>
,
<location id="EF774AB5FFA5FFDB75C536B0FBE8E089" LSID="urn:lsid:plazi:treatment:6201AD78FFA6FFDA7693333BFF3FE3A9:EF774AB5FFA5FFDB75C536B0FBE8E089" box="[961,1032,655,680]" country="China" municipality="Hekou" name="Nanxi" pageId="14" pageNumber="365" stateProvince="Yunnan">Nanxi</location>
,
<date id="9E163AAEFFA5FFDB721336AFFB40E089" box="[1047,1184,656,680]" pageId="14" pageNumber="365" value="2009-05-22">
<collectingDate id="8E52C346FFA5FFDB721336AFFB40E089" box="[1047,1184,656,680]" pageId="14" pageNumber="365" value="2009-05-22">22. V. 2009</collectingDate>
</date>
, Guoquan Wang Leg. (
<collectionCode id="8CB984ABFFA5FFDB769B368BFF13E0ED" box="[159,243,692,716]" pageId="14" pageNumber="365">HZAU</collectionCode>
), specimens were stored in 75% ethanol
</materialsCitation>
;
<materialsCitation id="5AC01633FFA5FFDB74CA368BFF03E134" ID-GBIF-Occurrence="2234214312" collectingDate="2009-09-18" collectionCode="CAU" country="China" location="Apple" municipality="Zhaotong" pageId="14" pageNumber="365" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="FCAED7E7FFA5FFDB74CA368BFD1CE0ED" box="[718,764,692,716]" pageId="14" pageNumber="365" type="male">1 ♂</specimenCount>
,
<collectingCountry id="92BF5CFEFFA5FFDB750D368BFC84E0ED" box="[777,868,692,716]" name="China" pageId="14" pageNumber="365">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA5FFDB7576368AFC2BE0ED" box="[882,971,693,716]" country="China" name="Yunnan" pageId="14" pageNumber="365">Yunnan</collectingRegion>
,
<collectingMunicipality id="0A738614FFA5FFDB75D3368BFBA5E0EC" box="[983,1093,692,717]" pageId="14" pageNumber="365">Zhaotong</collectingMunicipality>
,
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orchard,
<date id="9E163AAEFFA5FFDB7300368BFA78E0ED" box="[1284,1432,692,716]" pageId="14" pageNumber="365" value="2009-09-18">
<collectingDate id="8E52C346FFA5FFDB7300368BFA78E0ED" box="[1284,1432,692,716]" pageId="14" pageNumber="365" value="2009-09-18">18. IX. 2009</collectingDate>
</date>
, Tingting Zhang Leg. (
<collectionCode id="8CB984ABFFA5FFDB779C36E7FE36E0D1" box="[408,470,728,752]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="14" pageNumber="365">CAU</collectionCode>
) (photo and abdomen dissected), specimen dry mounted on label laterally on right thorax
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="BED74CE6FFA5FFDB769333A7FF26E5CA" httpUri="https://zenodo.org/record/2623732/files/figure.png" pageId="14" pageNumber="365" startId="14.[151,264,1944,1966]" targetBox="[174,1407,850,1907]" targetPageId="14">
<paragraph id="EA171C6EFFA5FFDB769333A7FF26E5CA" blockId="14.[151,1436,1944,2027]" pageId="14" pageNumber="365">
<emphasis id="D8DCC07CFFA5FFDB769333A7FEB4E58F" bold="true" box="[151,340,1944,1966]" pageId="14" pageNumber="365">FIGURES 32–37.</emphasis>
<taxonomicName id="2DA867EDFFA5FFDB775D33A6FE56E58F" authorityName="Loew" authorityYear="1873" box="[345,438,1945,1966]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFA5FFDB775D33A6FE56E58F" box="[345,438,1945,1966]" italics="true" pageId="14" pageNumber="365">Dicraeus</emphasis>
</taxonomicName>
species, male. epandrium, posterior view. 32
<taxonomicName id="2DA867EDFFA5FFDB758033A6FB9BE58F" authority="Kanmiya" authorityName="Kanmiya" authorityYear="1971" box="[900,1147,1944,1966]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="365" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="D8DCC07CFFA5FFDB758033A6FBF5E58F" box="[900,1045,1944,1966]" italics="true" pageId="14" pageNumber="365">D. fuscipennis</emphasis>
Kanmiya
</taxonomicName>
; 33
<taxonomicName id="2DA867EDFFA5FFDB72A133A6FA98E58F" authority="Nartshuk" authorityName="Nartshuk" box="[1189,1400,1944,1966]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="365" phylum="Arthropoda" rank="species" species="malaise">
<emphasis id="D8DCC07CFFA5FFDB72A133A6FAF3E58F" box="[1189,1299,1944,1966]" italics="true" pageId="14" pageNumber="365">D. malaise</emphasis>
Nartshuk
</taxonomicName>
; 34
<taxonomicName id="2DA867EDFFA5FFDB76933387FE6EE5EC" authority="Kanmiya" authorityName="Kanmiya" authorityYear="1971" box="[151,398,1975,1997]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="365" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="D8DCC07CFFA5FFDB76933387FEC8E5EC" box="[151,296,1975,1997]" italics="true" pageId="14" pageNumber="365">D. nartshukae</emphasis>
Kanmiya
</taxonomicName>
; 35
<taxonomicName id="2DA867EDFFA5FFDB77BE3387FD6EE5EC" authority="Becker" authorityName="Becker" authorityYear="1911" box="[442,654,1975,1997]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="365" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="D8DCC07CFFA5FFDB77BE3387FDDBE5EC" box="[442,571,1975,1997]" italics="true" pageId="14" pageNumber="365">D. orientalis</emphasis>
Becker
</taxonomicName>
; 36
<taxonomicName id="2DA867EDFFA5FFDB74BE3387FC37E5EC" authority="Kanmiya" authorityName="Kanmiya" authorityYear="1971" box="[698,983,1975,1997]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="365" phylum="Arthropoda" rank="species" species="phyllostachyus">
<emphasis id="D8DCC07CFFA5FFDB74BE3387FC8FE5EC" box="[698,879,1975,1997]" italics="true" pageId="14" pageNumber="365">D. phyllostachyus</emphasis>
Kanmiya
</taxonomicName>
; 37
<taxonomicName id="2DA867EDFFA5FFDB72073387FAD8E5EC" authority="Stackelberg. Scale" authorityName="Stackelberg. Scale" authorityYear="1955" box="[1027,1336,1975,1997]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="365" phylum="Arthropoda" rank="species" species="rossicus">
<emphasis id="D8DCC07CFFA5FFDB72073387FB95E5EC" box="[1027,1141,1976,1997]" italics="true" pageId="14" pageNumber="365">D. rossicus</emphasis>
Stackelberg. Scale
</taxonomicName>
bar = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="A2B24FE5FFA4FFDA76C334A8FD61E291" box="[199,641,151,176]" pageId="15" pageNumber="366" type="distribution">
<paragraph id="EA171C6EFFA4FFDA76C334A8FD61E291" blockId="15.[151,1437,151,392]" box="[199,641,151,176]" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA76C334A8FE80E291" bold="true" box="[199,352,151,176]" pageId="15" pageNumber="366">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFA4FFDA776334A8FE4AE291" box="[359,426,151,176]" name="China" pageId="15" pageNumber="366">China</collectingCountry>
(
<collectingRegion id="286CD28CFFA4FFDA77BD34A7FDF3E291" box="[441,531,152,176]" country="China" name="Yunnan" pageId="15" pageNumber="366">Yunnan</collectingRegion>
,
<collectingRegion id="286CD28CFFA4FFDA741A34A8FD9DE291" box="[542,637,151,176]" country="China" name="Taiwan)" pageId="15" pageNumber="366">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA4FFDA76C33483FF3FE3A9" pageId="15" pageNumber="366" type="discussion">
<paragraph id="EA171C6EFFA4FFDA76C33483FF3FE3A9" blockId="15.[151,1437,151,392]" pageId="15" pageNumber="366">
<emphasis id="D8DCC07CFFA4FFDA76C33483FEDBE2F4" bold="true" box="[199,315,188,213]" pageId="15" pageNumber="366">Remarks.</emphasis>
The new species is somewhat similar to
<taxonomicName id="2DA867EDFFA4FFDA751F3482FB41E2F5" authority="Kanmiya, 1971" authorityName="Kanmiya" authorityYear="1971" box="[795,1185,188,213]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="phyllostachyus">
<emphasis id="D8DCC07CFFA4FFDA751F3482FC06E2F5" box="[795,998,188,212]" italics="true" pageId="15" pageNumber="366">D. phyllostachyus</emphasis>
<bibRefCitation id="8E39619FFFA4FFDA75F43483FB41E2F5" author="Kanmiya, K." box="[1008,1185,188,213]" pageId="15" pageNumber="366" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the ocellar triangle reaching anterior 3/4 of frons, antenna entirely black and surstylus divided two lobes, dorsal lobe short, broadened, ventral lobe elongate, slender. In
<taxonomicName id="2DA867EDFFA4FFDA75EE353AFB55E33D" authorityName="Kanmiya" authorityYear="1971" box="[1002,1205,260,284]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="366" phylum="Arthropoda" rank="species" species="phyllostachyus">
<emphasis id="D8DCC07CFFA4FFDA75EE353AFB55E33D" box="[1002,1205,260,284]" italics="true" pageId="15" pageNumber="366">D. phyllostachyus</emphasis>
</taxonomicName>
, the ocellar triangle reaches the anterior margin of frons; the antenna is yellow with the first flagellomere partly yellow; and the surstylus is gently broadened basally, suddenly narrowed at middle and curved in a S-shaped form (
<bibRefCitation id="8E39619FFFA4FFDA73303573FF33E3A9" author="Kanmiya, K." pageId="15" pageNumber="366" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

175
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<treatment id="6201AD78FFA8FFD6769334A8FE58E5B2" ID-DOI="http://doi.org/10.5281/zenodo.5942653" ID-GBIF-Taxon="154532338" ID-Zenodo-Dep="5942653" LSID="urn:lsid:plazi:treatment:6201AD78FFA8FFD6769334A8FE58E5B2" httpUri="http://treatment.plazi.org/id/6201AD78FFA8FFD6769334A8FE58E5B2" lastPageNumber="354" pageId="3" pageNumber="354">
<subSubSection id="A2B24FE5FFA8FFD6769334A8FDE4E293" box="[151,516,151,178]" pageId="3" pageNumber="354" type="nomenclature">
<paragraph id="EA171C6EFFA8FFD6769334A8FDE4E293" blockId="3.[151,516,151,211]" box="[151,516,151,178]" pageId="3" pageNumber="354">
<heading id="B15FAB02FFA8FFD6769334A8FDE4E293" bold="true" box="[151,516,151,178]" fontSize="11" level="1" pageId="3" pageNumber="354" reason="1">
<emphasis id="D8DCC07CFFA8FFD6769334A8FDE4E293" bold="true" box="[151,516,151,178]" pageId="3" pageNumber="354">
<taxonomicName id="2DA867EDFFA8FFD6769334A8FE7FE290" ID-CoL="9HVHC" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[151,415,151,177]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="crassispinus" status="sp. nov.">
<emphasis id="D8DCC07CFFA8FFD6769334A8FE7FE290" bold="true" box="[151,415,151,177]" italics="true" pageId="3" pageNumber="354">Dicraeus crassispinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3EF7D07FFA8FFD677A334A7FDE4E293" box="[423,516,152,178]" pageId="3" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA8FFD676933485FED0E2F2" box="[151,304,186,211]" pageId="3" pageNumber="354" type="description">
<paragraph id="EA171C6EFFA8FFD676933485FED0E2F2" blockId="3.[151,516,151,211]" box="[151,304,186,211]" pageId="3" pageNumber="354">
<figureCitation id="729300EBFFA8FFD676933485FF3CE2F2" box="[151,220,186,211]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1268,1290]" captionTargetBox="[230,1360,680,1228]" captionTargetId="figure@2.[226,1360,665,1244]" captionTargetPageId="2" captionText="FIGURES 1–5. Dicraeus species, habitus. 1 D. crassispinus sp. nov.; 2 D. curvus sp. nov.; 3 D. dentatus sp. nov.; 4 D. liupanshanensis sp. nov.; 5 D. minutidentatus sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623715/files/figure.png" pageId="3" pageNumber="354">Figs 1</figureCitation>
,
<figureCitation id="729300EBFFA8FFD676EE3485FED0E2F2" box="[234,304,186,211]" captionStart="FIGURES 12–15" captionStartId="4.[151,264,1568,1590]" captionTargetBox="[180,1433,224,1509]" captionTargetId="figure@4.[151,1436,195,1546]" captionTargetPageId="4" captionText="FIGURES 12–15. Dicraeus crassispinus sp. nov. male. 12 epandrium, posterior view; 13 epandrium, lateral view; 14 hypandrium and phallic complex, ventral view; 15 hypandrium and phallic complex, lateral view. Abbreviations: bas, basiphallus; dis, distiphallus; ep, epandrium; hyp, hypandrium; phal, phallapodeme; pog, postgonite; prg, pregonite; sur, surstylus. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623719/files/figure.png" pageId="3" pageNumber="354">12–15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA8FFD67693353DFDBCE3A6" pageId="3" pageNumber="354" type="diagnosis">
<paragraph id="EA171C6EFFA8FFD67693353DFDBCE3A6" blockId="3.[151,1437,258,1939]" pageId="3" pageNumber="354">
<emphasis id="D8DCC07CFFA8FFD67693353DFEF1E33A" bold="true" box="[151,273,258,283]" pageId="3" pageNumber="354">Diagnosis.</emphasis>
Gena 0.5 times as wide as first flagellomere. Ocellar triangle reaching anterior margin of frons. Scutum brown with 5 longitudinal black stripes. Scutellum blackish brown. Legs yellow with tarsomeres 5 brown. Surstylus distinctly shorter than epandrium in lateral view, basal 2/3 nearly parallel-sided, recurved posteriorly, some thickened setae on middle surface.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA8FFD676C335ADFE7EE79E" pageId="3" pageNumber="354" type="description">
<paragraph id="EA171C6EFFA8FFD676C335ADFC3FE38B" blockId="3.[151,1437,258,1939]" box="[199,991,402,427]" pageId="3" pageNumber="354">
<emphasis id="D8DCC07CFFA8FFD676C335ADFEB9E38A" bold="true" box="[199,345,402,427]" pageId="3" pageNumber="354">Description.</emphasis>
Male (
<figureCitation id="729300EBFFA8FFD677AC35ADFE09E38A" box="[424,489,402,427]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1268,1290]" captionTargetBox="[230,1360,680,1228]" captionTargetId="figure@2.[226,1360,665,1244]" captionTargetPageId="2" captionText="FIGURES 1–5. Dicraeus species, habitus. 1 D. crassispinus sp. nov.; 2 D. curvus sp. nov.; 3 D. dentatus sp. nov.; 4 D. liupanshanensis sp. nov.; 5 D. minutidentatus sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623715/files/figure.png" pageId="3" pageNumber="354">Fig. 1</figureCitation>
). Body length
<quantity id="2D50B18BFFA8FFD6749635ADFD0AE38B" box="[658,746,402,427]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="354" unit="mm" value="3.5">3.5 mm</quantity>
, wing length
<quantity id="2D50B18BFFA8FFD6758635ACFC3BE38B" box="[898,987,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="354" unit="mm" value="1.8">1.8 mm</quantity>
.
</paragraph>
<paragraph id="EA171C6EFFA8FFD676C33589FBA1E0EA" blockId="3.[151,1437,258,1939]" pageId="3" pageNumber="354">Head yellow, as wide as thorax, 0.85 times as long as high in profile, face somewhat concave in lateral view, facial carina indistinct; frons yellowish brown with anterior 1/2 yellow, as long as wide, projecting slightly in front of eye; gena broad, about 0.5 times as wide as first flagellomere, with few brown setulae on ventral margin; parafacial indistinct; clypeus light yellow; vibrissal angle obtuse. Ocellar triangle yellowish brown with distal 1/4 yellow, smooth, shiny, reaching anterior margin of frons, with pointed apex; ocellar tubercle brown. Occiput black. Cephalic setae and setulae black. Antenna yellow with thick grey pruinescence except for dorsal 1/2 of first flagellomere yellowish brown; first flagellomere 0.8 times as long as wide; arista brown with short brown pubescence. Proboscis brown with brown setulae; palpus yellow with black setulae.</paragraph>
<paragraph id="EA171C6EFFA8FFD676C336E8FE8CE65A" blockId="3.[151,1437,258,1939]" pageId="3" pageNumber="354">Scutum 1.1 times as long as wide, brown with 5 longitudinal black stripes, medial stripe slightly broad and extending to posterior 4/5 of scutum, basal 1/3 of inner lateral stripes narrow, outer lateral stripes short and narrow. Postpronotal lobe yellow with a reddish yellow spot. Thoracic pleuron shiny yellow except for anteroventral portion of anepisternum brown; anterior 1/2 of anepumeron, ventral 1/2 of katepisternum and posteroventral portion of meron yellowish brown. Scutellum blackish brown, 0.5 times as long as wide; apical scutellar seta 1.5 times as long as scutellum; subapical scutellar seta as long as scutellum. Postscutellum black. Setae and setulae on thorax black; notopleurals 1+2, anterior notopleural seta as long as posterior notopleural seta; anterior postalar seta as long as posterior postalar seta; 1 pair of dorsocentral setae developed, 1.8 times as long as anterior postalar setae. Legs slender, yellow with tarsomeres 5 brown. Mid tibia with a short black subapical spur, which is longer than diameter of tibia. Tibial organ long and narrow. Setulae on legs black. Wing 2.6 times as long as wide, hyaline; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 8: 1.2: 1; r-m at basal 0.45 of discal medial cell. Halter pale yellow.</paragraph>
<paragraph id="EA171C6EFFA8FFD676C330B9FEDFE6E2" blockId="3.[151,1437,258,1939]" pageId="3" pageNumber="354">Abdomen brown except for tergite 1, distal portions of tergites 2–5 and tergites 7+8 yellow; venter yellow. Setulae brown.</paragraph>
<paragraph id="EA171C6EFFA8FFD676C330F1FEB1E7BA" blockId="3.[151,1437,258,1939]" pageId="3" pageNumber="354">
Male genitalia (
<figureCitation id="729300EBFFA8FFD6777230F1FDE0E6C6" box="[374,512,1230,1255]" captionStart="FIGURES 12–15" captionStartId="4.[151,264,1568,1590]" captionTargetBox="[180,1433,224,1509]" captionTargetId="figure@4.[151,1436,195,1546]" captionTargetPageId="4" captionText="FIGURES 12–15. Dicraeus crassispinus sp. nov. male. 12 epandrium, posterior view; 13 epandrium, lateral view; 14 hypandrium and phallic complex, ventral view; 15 hypandrium and phallic complex, lateral view. Abbreviations: bas, basiphallus; dis, distiphallus; ep, epandrium; hyp, hypandrium; phal, phallapodeme; pog, postgonite; prg, pregonite; sur, surstylus. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623719/files/figure.png" pageId="3" pageNumber="354">Figs. 12–15</figureCitation>
): Epandrium yellow with brown distally, with long brown setulae, height 1.7 times width in lateral view; surstylus distinctly shorter than depth of epandrium in lateral view, basal 2/3 nearly parallelsided, recurved posteriorly; some thickened setae on middle surface. Cercus absent. Pre- and postgonites short, postgonite round distally, with few sensilla; basiphallus longer than wide, cylindrical; distiphallus short, membranous; phallapodeme sclerotized, long, with basal stalk broad in lateral view. Hypandrium closed, apically forming an arch.
</paragraph>
<paragraph id="EA171C6EFFA8FFD676C33199FE7EE79E" blockId="3.[151,1437,258,1939]" box="[199,414,1446,1471]" pageId="3" pageNumber="354">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA8FFD676C331F5FD4FE40A" pageId="3" pageNumber="354" type="materials_examined">
<paragraph id="EA171C6EFFA8FFD676C331F5FD4FE40A" blockId="3.[151,1437,258,1939]" pageId="3" pageNumber="354">
<emphasis id="D8DCC07CFFA8FFD676C331F5FDC5E7C2" bold="true" box="[199,549,1482,1507]" pageId="3" pageNumber="354">
Type material.
<materialsCitation id="5AC01633FFA8FFD6778231F5FDC5E7C2" ID-GBIF-Occurrence="2234214343" box="[390,549,1482,1507]" collectingDate="2006-05-21" collectionCode="CAU" country="China" county="Honghe" location="Binlangzhai Reservoir" municipality="Hekou" pageId="3" pageNumber="354" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="3513A2CCFFA8FFD6778231F5FDC5E7C2" box="[390,549,1482,1507]" pageId="3" pageNumber="354" type="holotype">HOLOTYPE</typeStatus>
</materialsCitation>
</emphasis>
<emphasis id="D8DCC07CFFA8FFD6743531F4FDA6E7C2" box="[561,582,1483,1507]" italics="true" pageId="3" pageNumber="354">6</emphasis>
,
<collectingCountry id="92BF5CFEFFA8FFD6745D31F4FD55E7C2" box="[601,693,1483,1507]" name="China" pageId="3" pageNumber="354">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFA8FFD674C131F4FCFEE7C3" box="[709,798,1483,1506]" country="China" name="Yunnan" pageId="3" pageNumber="354">Yunnan</collectingRegion>
,
<collectingCounty id="037664E2FFA8FFD6752A31F5FC69E7C2" box="[814,905,1482,1507]" pageId="3" pageNumber="354">Honghe</collectingCounty>
,
<collectingMunicipality id="0A738614FFA8FFD6759E31F5FC08E7C2" box="[922,1000,1482,1507]" pageId="3" pageNumber="354">Hekou</collectingMunicipality>
,
<location id="EF774AB5FFA8FFD675FC31F5FB1AE7C2" LSID="urn:lsid:plazi:treatment:6201AD78FFA8FFD6769334A8FE58E5B2:EF774AB5FFA8FFD675FC31F5FB1AE7C2" box="[1016,1274,1482,1507]" country="China" county="Honghe" municipality="Hekou" name="Binlangzhai Reservoir" pageId="3" pageNumber="354" stateProvince="Yunnan">Binlangzhai Reservoir</location>
,
<date id="9E163AAEFFA8FFD6730E31F4FA78E7C2" box="[1290,1432,1482,1507]" pageId="3" pageNumber="354" value="2006-05-21">
<collectingDate id="8E52C346FFA8FFD6730E31F4FA78E7C2" box="[1290,1432,1482,1507]" pageId="3" pageNumber="354" value="2006-05-21">21. V. 2006</collectingDate>
</date>
, Xiushuai Yang Leg. (CAU) (photo and abdomen dissected).
<materialsCitation id="5AC01633FFA8FFD6754C31D1FD4AE40A" ID-GBIF-Occurrence="2234214313" collectingDate="2006-05-21" collectionCode="CAU" country="China" county="Honghe" location="Binlangzhai Reservoir" municipality="Hekou" pageId="3" pageNumber="354" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="D8DCC07CFFA8FFD6754C31D1FC0CE426" bold="true" box="[840,1004,1518,1543]" pageId="3" pageNumber="354">
<typeStatus id="3513A2CCFFA8FFD6754C31D1FC0CE426" box="[840,1004,1518,1543]" pageId="3" pageNumber="354" type="paratype">PARATYPES</typeStatus>
</emphasis>
2
<emphasis id="D8DCC07CFFA8FFD6720F31D1FBD5E427" box="[1035,1077,1518,1542]" italics="true" pageId="3" pageNumber="354">66</emphasis>
, same data as holotype (
<collectionCode id="8CB984ABFFA8FFD6734B31D0FA70E426" box="[1359,1424,1519,1543]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="3" pageNumber="354">CAU</collectionCode>
). All type specimens were stored in 75% ethanol
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA8FFD676C33209FDC1E46E" box="[199,545,1590,1615]" pageId="3" pageNumber="354" type="distribution">
<paragraph id="EA171C6EFFA8FFD676C33209FDC1E46E" blockId="3.[151,1437,258,1939]" box="[199,545,1590,1615]" pageId="3" pageNumber="354">
<emphasis id="D8DCC07CFFA8FFD676C33209FE80E46E" bold="true" box="[199,352,1590,1615]" pageId="3" pageNumber="354">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFA8FFD677633209FE4AE46E" box="[359,426,1590,1615]" name="China" pageId="3" pageNumber="354">China</collectingCountry>
(
<collectingRegion id="286CD28CFFA8FFD677BD3208FDF5E46E" box="[441,533,1591,1615]" country="China" name="Yunnan" pageId="3" pageNumber="354">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA8FFD676C33265FEB6E56A" pageId="3" pageNumber="354" type="discussion">
<paragraph id="EA171C6EFFA8FFD676C33265FEB6E56A" blockId="3.[151,1437,258,1939]" pageId="3" pageNumber="354">
<emphasis id="D8DCC07CFFA8FFD676C33265FEDBE452" bold="true" box="[199,315,1626,1651]" pageId="3" pageNumber="354">Remarks.</emphasis>
The new species is somewhat similar to
<taxonomicName id="2DA867EDFFA8FFD6750C3264FBA4E452" authority="Kanmiya, 1971" authorityName="Kanmiya" authorityYear="1971" box="[776,1092,1626,1651]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="ischaemi">
<emphasis id="D8DCC07CFFA8FFD6750C3264FC6EE452" box="[776,910,1627,1651]" italics="true" pageId="3" pageNumber="354">D. ischaemi</emphasis>
<bibRefCitation id="8E39619FFFA8FFD675913265FBA4E452" author="Kanmiya, K." box="[917,1092,1626,1651]" pageId="3" pageNumber="354" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the following features: ocellar triangle reaching anterior margin of frons; first flagellomere round dorsodistally; surstylus distinctly shorter than depth of epandrium in lateral view, basal 2/3 nearly parallel-sided, recurved posteriorly; some thickened setae on middle surface; cercus absent. In
<taxonomicName id="2DA867EDFFA8FFD6721F32F8FB41E4FF" authorityName="Kanmiya" authorityYear="1971" box="[1051,1185,1734,1758]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="ischaemi">
<emphasis id="D8DCC07CFFA8FFD6721F32F8FB41E4FF" box="[1051,1185,1734,1758]" italics="true" pageId="3" pageNumber="354">D. ischaemi</emphasis>
</taxonomicName>
, the ocellar triangle is extended to a little beyond middle of frons; the first flagellomere has a dorsodistal angle; the surstylus is slender, slightly concave at middle, broadly round apically, and has 5 long thick setae; and the cercus is slender, finger-like (
<bibRefCitation id="8E39619FFFA8FFD6769A330DFEAAE56A" author="Kanmiya, K." box="[158,330,1842,1867]" pageId="3" pageNumber="354" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA8FFD676C33369FE58E5B2" pageId="3" pageNumber="354" type="etymology">
<paragraph id="EA171C6EFFA8FFD676C33369FE58E5B2" blockId="3.[151,1437,258,1939]" pageId="3" pageNumber="354">
<emphasis id="D8DCC07CFFA8FFD676C33369FEAEE54E" bold="true" box="[199,334,1878,1903]" pageId="3" pageNumber="354">Etymology.</emphasis>
The specific name is from the Latin
<taxonomicName id="2DA867EDFFA8FFD674EC3368FC92E54F" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[744,882,1879,1902]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="crassispinus">
<emphasis id="D8DCC07CFFA8FFD674EC3368FC92E54F" box="[744,882,1879,1902]" italics="true" pageId="3" pageNumber="354">crassispinus</emphasis>
</taxonomicName>
(“crassispina”), referring to the thickened setulae on inner side of surstylus.
</paragraph>
</subSubSection>
</treatment>
</document>

284
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<subSubSection id="A2B24FE5FFAAFFD476933658FE77E0A3" box="[151,407,615,642]" pageId="1" pageNumber="352" type="nomenclature">
<paragraph id="EA171C6EFFAAFFD476933658FE77E0A3" blockId="1.[151,407,615,642]" box="[151,407,615,642]" pageId="1" pageNumber="352">
<heading id="B15FAB02FFAAFFD476933658FE77E0A3" bold="true" box="[151,407,615,642]" fontSize="11" level="1" pageId="1" pageNumber="352" reason="1">
<taxonomicName id="2DA867EDFFAAFFD476933658FE77E0A3" ID-CoL="62QZH" ID-ENA="540857" authority="Loew, 1873" authorityName="Loew" authorityYear="1873" box="[151,407,615,642]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD476933658FE77E0A3" bold="true" box="[151,407,615,642]" pageId="1" pageNumber="352">
<emphasis id="D8DCC07CFFAAFFD476933658FEE4E0A0" bold="true" box="[151,260,615,641]" italics="true" pageId="1" pageNumber="352">Dicraeus</emphasis>
<bibRefCitation id="8E39619FFFAAFFD4770F3657FE77E0A3" author="Loew, H." box="[267,407,616,642]" pageId="1" pageNumber="352" pagination="33 - 52" refId="ref12196" refString="Loew, H. (1873) Diptera nova, in Pannonia inferiori et in confinibus Daciae regionibus a Ferd. Kowarzio capta. Berliner Entomologische Zeitschrift, 17, 33 - 52." type="journal article" year="1873">Loew, 1873</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="A2B24FE5FFAAFFD47693368FFB39E1D6" pageId="1" pageNumber="352" type="reference_group">
<paragraph id="EA171C6EFFAAFFD47693368FFC19E0E5" blockId="1.[151,1436,686,1015]" box="[151,1017,686,709]" pageId="1" pageNumber="352">
<treatmentCitationGroup id="CAB83B40FFAAFFD47693368FFC19E0E5" box="[151,1017,686,709]" pageId="1" pageNumber="352">
<taxonomicName id="2DA867EDFFAAFFD47693368FFE7AE0E5" ID-CoL="62QZH" authority="Loew, 1873: 51" authorityName="Loew" authorityPageNumber="51" authorityYear="1873" box="[151,410,686,709]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD47693368FFF14E0E4" box="[151,244,688,709]" italics="true" pageId="1" pageNumber="352">Dicraeus</emphasis>
<treatmentCitation id="6B093A7FFFAAFFD476FE3690FE7AE0E5" author="Loew, H." box="[250,410,686,708]" page="51" pageId="1" pageNumber="352" year="1873">
<bibRefCitation id="8E39619FFFAAFFD476FE3690FE95E0E5" author="Loew, H." box="[250,373,686,708]" pageId="1" pageNumber="352" pagination="33 - 52" refId="ref12196" refString="Loew, H. (1873) Diptera nova, in Pannonia inferiori et in confinibus Daciae regionibus a Ferd. Kowarzio capta. Berliner Entomologische Zeitschrift, 17, 33 - 52." type="journal article" year="1873">Loew, 1873</bibRefCitation>
: 51
</treatmentCitation>
</taxonomicName>
.
<emphasis id="D8DCC07CFFAAFFD477A03691FDCBE0E5" bold="true" box="[420,555,686,708]" pageId="1" pageNumber="352">
<typeStatus id="3513A2CCFFAAFFD477A03691FE39E0E5" box="[420,473,686,708]" pageId="1" pageNumber="352">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="2DA867EDFFAAFFD4743C368FFC94E0E5" ID-CoL="35QR8" authority="Loew, 1873" authorityName="Loew" authorityYear="1873" box="[568,884,686,709]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="D8DCC07CFFAAFFD4743C368FFD16E0E5" box="[568,758,686,709]" italics="true" pageId="1" pageNumber="352">Dicraeus obscurus</emphasis>
<bibRefCitation id="8E39619FFFAAFFD474F83690FC94E0E5" author="Loew, H." box="[764,884,686,708]" pageId="1" pageNumber="352" pagination="33 - 52" refId="ref12196" refString="Loew, H. (1873) Diptera nova, in Pannonia inferiori et in confinibus Daciae regionibus a Ferd. Kowarzio capta. Berliner Entomologische Zeitschrift, 17, 33 - 52." type="journal article" year="1873">Loew, 1873</bibRefCitation>
</taxonomicName>
(monotypy).
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</paragraph>
<paragraph id="EA171C6EFFAAFFD4769336F1FB08E0C2" blockId="1.[151,1436,686,1015]" box="[151,1256,717,739]" pageId="1" pageNumber="352">
<treatmentCitationGroup id="CAB83B40FFAAFFD4769336F1FB08E0C2" box="[151,1256,717,739]" pageId="1" pageNumber="352">
<taxonomicName id="2DA867EDFFAAFFD4769336F1FE54E0C2" authority="Pandelle, 1898: 18" authorityName="Pandelle" authorityPageNumber="18" authorityYear="1898" box="[151,436,717,739]" class="Insecta" genus="Parastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD4769336F1FF0EE0C2" box="[151,238,718,739]" italics="true" pageId="1" pageNumber="352">Parastia</emphasis>
<treatmentCitation id="6B093A7FFFAAFFD476F036F2FE54E0C2" author="Pandelle, L." box="[244,436,717,739]" page="18" pageId="1" pageNumber="352" year="1898">
<bibRefCitation id="8E39619FFFAAFFD476F036F2FE6FE0C2" author="Pandelle, L." box="[244,399,717,739]" pageId="1" pageNumber="352" pagination="1 - 80" refId="ref12879" refString="Pandelle, L. (1898) Etudes sur les Muscides de France. III e. partie (part). Revue d'Entomologie, Caen. 17 (Supplement), 1 - 80." type="journal article" year="1898">Pandellé, 1898</bibRefCitation>
: 18
</treatmentCitation>
</taxonomicName>
.
<emphasis id="D8DCC07CFFAAFFD477BA36F2FDA4E0C2" bold="true" box="[446,580,717,739]" pageId="1" pageNumber="352">
<typeStatus id="3513A2CCFFAAFFD477BA36F2FE13E0C2" box="[446,499,717,739]" pageId="1" pageNumber="352">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="2DA867EDFFAAFFD4745536F1FC99E0C2" ID-CoL="8D4QC" authority="Haliday, 1838" authorityName="Haliday" authorityYear="1838" box="[593,889,717,739]" class="Insecta" family="Chloropidae" genus="Oscinis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="raptus">
<emphasis id="D8DCC07CFFAAFFD4745536F1FD05E0C2" box="[593,741,718,739]" italics="true" pageId="1" pageNumber="352">Oscinis raptus</emphasis>
Haliday, 1838
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(by designation of Sabrosky, 1964).
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<paragraph id="EA171C6EFFAAFFD4769336D3FBDAE123" blockId="1.[151,1436,686,1015]" box="[151,1082,748,770]" pageId="1" pageNumber="352">
<treatmentCitationGroup id="CAB83B40FFAAFFD4769336D3FBDAE123" box="[151,1082,748,770]" pageId="1" pageNumber="352">
<taxonomicName id="2DA867EDFFAAFFD4769336D3FE26E123" authority="Becker, 1910: 146" authorityName="Becker" authorityPageNumber="146" authorityYear="1910" box="[151,454,748,770]" class="Insecta" family="Chloropidae" genus="Oedesiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD4769336D3FEE3E123" box="[151,259,748,770]" italics="true" pageId="1" pageNumber="352">Oedesiella</emphasis>
<treatmentCitation id="6B093A7FFFAAFFD4770D36D3FE26E123" author="Becker, T." box="[265,454,748,770]" page="146" pageId="1" pageNumber="352" year="1910">
<bibRefCitation id="8E39619FFFAAFFD4770D36D3FE73E123" author="Becker, T." box="[265,403,748,770]" pageId="1" pageNumber="352" pagination="33 - 174" refId="ref11558" refString="Becker, T. (1910) Chloropidae. Eine monographische Studie. I. Teil. Palaarktische Region. Archivum Zoologicum, 1 (10), 33 - 174." type="journal article" year="1910">Becker, 1910</bibRefCitation>
: 146
</treatmentCitation>
</taxonomicName>
.
<emphasis id="D8DCC07CFFAAFFD477CB36D3FDB6E123" bold="true" box="[463,598,748,770]" pageId="1" pageNumber="352">
<typeStatus id="3513A2CCFFAAFFD477CB36D3FDE4E123" box="[463,516,748,770]" pageId="1" pageNumber="352">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="2DA867EDFFAAFFD4746736D3FC55E123" ID-CoL="8D32C" authority="Becker, 1910" authorityName="Becker" authorityYear="1910" box="[611,949,748,770]" class="Insecta" family="Chloropidae" genus="Oedesiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="D8DCC07CFFAAFFD4746736D3FCC8E123" box="[611,808,748,770]" italics="true" pageId="1" pageNumber="352">Oedesiella discolor</emphasis>
<bibRefCitation id="8E39619FFFAAFFD4752A36D3FC55E123" author="Becker, T." box="[814,949,748,770]" pageId="1" pageNumber="352" pagination="33 - 174" refId="ref11558" refString="Becker, T. (1910) Chloropidae. Eine monographische Studie. I. Teil. Palaarktische Region. Archivum Zoologicum, 1 (10), 33 - 174." type="journal article" year="1910">Becker, 1910</bibRefCitation>
</taxonomicName>
(monotypy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA171C6EFFAAFFD476933734FBAAE101" blockId="1.[151,1436,686,1015]" box="[151,1098,778,800]" pageId="1" pageNumber="352">
<treatmentCitationGroup id="CAB83B40FFAAFFD476933734FBAAE101" box="[151,1098,778,800]" pageId="1" pageNumber="352">
<taxonomicName id="2DA867EDFFAAFFD476933734FE21E101" authority="Becker, 1912: 250" authorityName="Becker" authorityPageNumber="250" authorityYear="1912" box="[151,449,778,800]" class="Insecta" family="Chloropidae" genus="Oxyapium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD476933734FF1FE101" box="[151,255,779,800]" italics="true" pageId="1" pageNumber="352">Oxyapium</emphasis>
<treatmentCitation id="6B093A7FFFAAFFD477013735FE21E101" author="Becker, T." box="[261,449,778,800]" page="250" pageId="1" pageNumber="352" year="1912">
<bibRefCitation id="8E39619FFFAAFFD477013735FE6FE101" author="Becker, T." box="[261,399,778,800]" pageId="1" pageNumber="352" pagination="21 - 256" refId="ref11621" refString="Becker, T. (1912) Chloropidae. Eine monographische Studie. IV. Teil. Nearktische Region. V. Neotropische Region. Nachtrag. I. Palaearktische Region, II. Aethiopische Region. Annales Historico-Naturales Musei Nationalis Hungarici, 10, 21 - 256." type="journal article" year="1912">Becker, 1912</bibRefCitation>
: 250
</treatmentCitation>
</taxonomicName>
.
<emphasis id="D8DCC07CFFAAFFD477CF3735FDB1E101" bold="true" box="[459,593,778,800]" pageId="1" pageNumber="352">
<typeStatus id="3513A2CCFFAAFFD477CF3735FDE0E101" box="[459,512,778,800]" pageId="1" pageNumber="352">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="2DA867EDFFAAFFD4745A3734FC26E101" authority="Becker, 1912" authorityName="Becker" authorityYear="1912" box="[606,966,778,800]" class="Insecta" family="Chloropidae" genus="Oxyapium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="longinerve">
<emphasis id="D8DCC07CFFAAFFD4745A3734FCD8E101" box="[606,824,778,800]" italics="true" pageId="1" pageNumber="352">Oxyapium longinerve</emphasis>
<bibRefCitation id="8E39619FFFAAFFD4753B3735FC26E101" author="Becker, T." box="[831,966,778,800]" pageId="1" pageNumber="352" pagination="21 - 256" refId="ref11621" refString="Becker, T. (1912) Chloropidae. Eine monographische Studie. IV. Teil. Nearktische Region. V. Neotropische Region. Nachtrag. I. Palaearktische Region, II. Aethiopische Region. Annales Historico-Naturales Musei Nationalis Hungarici, 10, 21 - 256." type="journal article" year="1912">Becker, 1912</bibRefCitation>
</taxonomicName>
(monotypy).
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</paragraph>
<paragraph id="EA171C6EFFAAFFD476933715FBDFE11E" blockId="1.[151,1436,686,1015]" box="[151,1087,809,831]" pageId="1" pageNumber="352">
<treatmentCitationGroup id="CAB83B40FFAAFFD476933715FBDFE11E" box="[151,1087,809,831]" pageId="1" pageNumber="352">
<taxonomicName id="2DA867EDFFAAFFD476933715FE05E11E" authority="Enderlein, 1936: 186" authorityName="Enderlein" authorityPageNumber="186" authorityYear="1936" box="[151,485,809,831]" class="Insecta" family="Chloropidae" genus="Dicraeinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD476933715FEE8E11E" box="[151,264,810,831]" italics="true" pageId="1" pageNumber="352">Dicraeinus</emphasis>
<treatmentCitation id="6B093A7FFFAAFFD477093716FE05E11E" author="Enderlein, G." box="[269,485,809,831]" page="186" pageId="1" pageNumber="352" year="1936">
<bibRefCitation id="8E39619FFFAAFFD477093716FE53E11E" author="Enderlein, G." box="[269,435,809,831]" pageId="1" pageNumber="352" pagination="1 - 259" refId="ref11951" refString="Enderlein, G. (1936) Zweiflugler, Diptera. In: Brohmer, P. et al. (Eds.), Tierwelt Mitteleuropas. FI. Insekten, 3 Teil, Leipzig, 1 - 259." type="book chapter" year="1936">Enderlein, 1936</bibRefCitation>
: 186
</treatmentCitation>
</taxonomicName>
.
<emphasis id="D8DCC07CFFAAFFD477EB3716FD96E11E" bold="true" box="[495,630,809,831]" pageId="1" pageNumber="352">
<typeStatus id="3513A2CCFFAAFFD477EB3716FDC4E11E" box="[495,548,809,831]" pageId="1" pageNumber="352">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="2DA867EDFFAAFFD474863716FC5AE11E" ID-CoL="888YH" authority="Loew, 1866" authorityName="Loew" authorityYear="1866" box="[642,954,809,831]" class="Insecta" family="Chloropidae" genus="Eutropha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="ingratus">
<emphasis id="D8DCC07CFFAAFFD474863716FCDCE11E" box="[642,828,809,831]" italics="true" pageId="1" pageNumber="352">Eutropha ingratus</emphasis>
Loew, 1866
</taxonomicName>
(monotypy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA171C6EFFAAFFD476933777FD2EE17F" blockId="1.[151,1436,686,1015]" box="[151,718,840,862]" pageId="1" pageNumber="352">
<treatmentCitationGroup id="CAB83B40FFAAFFD476933777FD2EE17F" box="[151,718,840,862]" pageId="1" pageNumber="352">
<taxonomicName id="2DA867EDFFAAFFD476933777FE1EE17F" authority="Enderlein, 1936: 187" authorityName="Enderlein" authorityPageNumber="187" authorityYear="1936" box="[151,510,840,862]" class="Insecta" family="Chloropidae" genus="Paroedesiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD476933777FEC3E17F" box="[151,291,840,862]" italics="true" pageId="1" pageNumber="352">Paroedesiella</emphasis>
<treatmentCitation id="6B093A7FFFAAFFD4772D3777FE1EE17F" author="Enderlein, G." box="[297,510,840,862]" page="187" pageId="1" pageNumber="352" year="1936">
<bibRefCitation id="8E39619FFFAAFFD4772D3777FE2FE17F" author="Enderlein, G." box="[297,463,840,862]" pageId="1" pageNumber="352" pagination="1 - 259" refId="ref11951" refString="Enderlein, G. (1936) Zweiflugler, Diptera. In: Brohmer, P. et al. (Eds.), Tierwelt Mitteleuropas. FI. Insekten, 3 Teil, Leipzig, 1 - 259." type="book chapter" year="1936">Enderlein, 1936</bibRefCitation>
: 187
</treatmentCitation>
</taxonomicName>
(unavailable name).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA171C6EFFAAFFD476933759FEA6E1BA" blockId="1.[151,1436,686,1015]" pageId="1" pageNumber="352">
<treatmentCitationGroup id="CAB83B40FFAAFFD476933759FEA6E1BA" pageId="1" pageNumber="352">
<taxonomicName id="2DA867EDFFAAFFD476933759FD99E15D" authority="Anonymous in Imperial Institute" authorityName="Anonymous in Imperial Institute" box="[151,633,870,893]" class="Insecta" family="Chloropidae" genus="Paroedesiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD476933759FEC3E15D" box="[151,291,870,892]" italics="true" pageId="1" pageNumber="352">Paroedesiella</emphasis>
Anonymous
<emphasis id="D8DCC07CFFAAFFD477A93757FE21E15C" box="[429,449,872,893]" italics="true" pageId="1" pageNumber="352">in</emphasis>
Imperial Institute
</taxonomicName>
of
<treatmentCitation id="6B093A7FFFAAFFD4749A3759FC72E15D" author="Imperial Institute of Entomology" box="[670,914,870,892]" page="394" pageId="1" pageNumber="352" year="1937">
<bibRefCitation id="8E39619FFFAAFFD4749A3759FCBDE15D" author="Imperial Institute of Entomology" box="[670,861,870,892]" pageId="1" pageNumber="352" pagination="1 - 445" refId="ref12049" refString="Imperial Institute of Entomology (1937) Insecta. In: Smith, M. (Ed.), The zoological record. Volume the seventy-third being the records of zoological literature relating chiefly to the year 1936. Zoological Society, London, pp. 1 - 445." type="book chapter" year="1937">Entomology, 1937</bibRefCitation>
: 394
</treatmentCitation>
.
<emphasis id="D8DCC07CFFAAFFD475983759FBC4E15D" bold="true" box="[924,1060,870,892]" pageId="1" pageNumber="352">
<typeStatus id="3513A2CCFFAAFFD475983759FC31E15D" box="[924,977,870,892]" pageId="1" pageNumber="352">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="2DA867EDFFAAFFD472373758FA7BE15D" ID-CoL="8D4QS" authority="Strobl, 1898: 263" authorityName="Strobl" authorityPageNumber="263" authorityYear="1898" box="[1075,1435,870,893]" class="Insecta" family="Chloropidae" genus="Oscinis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="styriacus">
<emphasis id="D8DCC07CFFAAFFD472373758FB02E15C" box="[1075,1250,871,893]" italics="true" pageId="1" pageNumber="352">Oscinis styriacus</emphasis>
Strobl, 1898: 263
</taxonomicName>
(monotypy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA171C6EFFAAFFD47693379BFEABE1F9" blockId="1.[151,1436,686,1015]" pageId="1" pageNumber="352">
<treatmentCitationGroup id="CAB83B40FFAAFFD47693379BFEABE1F9" pageId="1" pageNumber="352">
<taxonomicName id="2DA867EDFFAAFFD47693379BFE18E19B" authority="Sabrosky, 1941: 761" authorityName="Sabrosky" authorityPageNumber="761" authorityYear="1941" box="[151,504,932,954]" class="Insecta" family="Chloropidae" genus="Paroedesiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD47693379BFEC3E19B" box="[151,291,932,954]" italics="true" pageId="1" pageNumber="352">Paroedesiella</emphasis>
<treatmentCitation id="6B093A7FFFAAFFD4772D379BFE18E19B" author="Sabrosky, C. W." box="[297,504,932,954]" page="761" pageId="1" pageNumber="352" year="1941">
<bibRefCitation id="8E39619FFFAAFFD4772D379BFE28E19B" author="Sabrosky, C. W." box="[297,456,932,954]" pageId="1" pageNumber="352" pagination="735 - 765" refId="ref12915" refString="Sabrosky, C. W. (1941) An annotated list of genotypes of the Chloropidae of the world (Diptera). Annals of the Entomological Society of America, 34 (4), 735 - 765. https: // doi. org / 10.1093 / aesa / 34.4.735" type="journal article" year="1941">Sabrosky, 1941</bibRefCitation>
: 761
</treatmentCitation>
</taxonomicName>
(preoccupied by Anonymous, 1937).
<emphasis id="D8DCC07CFFAAFFD4757E379BFBE0E19B" bold="true" box="[890,1024,932,954]" pageId="1" pageNumber="352">
<typeStatus id="3513A2CCFFAAFFD4757E379BFC4FE19B" box="[890,943,932,954]" pageId="1" pageNumber="352">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="2DA867EDFFAAFFD47209379BFADFE19B" ID-CoL="8D4QS" authority="Strobl, 1898" authorityName="Strobl" authorityYear="1898" box="[1037,1343,932,954]" class="Insecta" family="Chloropidae" genus="Oscinis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="styriacus">
<emphasis id="D8DCC07CFFAAFFD47209379BFB5BE19B" box="[1037,1211,932,954]" italics="true" pageId="1" pageNumber="352">Oscinis styriacus</emphasis>
Strobl, 1898
</taxonomicName>
(original designation).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA171C6EFFAAFFD4769337DEFB39E1D6" blockId="1.[151,1436,686,1015]" box="[151,1241,993,1015]" pageId="1" pageNumber="352">
<treatmentCitationGroup id="CAB83B40FFAAFFD4769337DEFB39E1D6" box="[151,1241,993,1015]" pageId="1" pageNumber="352">
<taxonomicName id="2DA867EDFFAAFFD4769337DEFE03E1D6" authority="Nartshuk, 1967: 426" authorityName="Nartshuk" authorityPageNumber="426" authorityYear="1967" box="[151,483,993,1015]" class="Insecta" family="Chloropidae" genus="Eudicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD4769337DEFEEAE1D6" box="[151,266,993,1015]" italics="true" pageId="1" pageNumber="352">Eudicraeus</emphasis>
<treatmentCitation id="6B093A7FFFAAFFD4771437DEFE03E1D6" author="Nartshuk, E. P." box="[272,483,993,1015]" page="426" pageId="1" pageNumber="352" year="1967">
<bibRefCitation id="8E39619FFFAAFFD4771437DEFE50E1D6" author="Nartshuk, E. P." box="[272,432,993,1015]" pageId="1" pageNumber="352" pagination="245 - 256" refId="ref12347" refString="Nartshuk, E. P. (1967) Chloropid flies of the genus Dicraeus Lw. (Diptera, Chloropidae). Entomologicheskoe Obozrenie, 46, 415 - 438. [In Russian, English translation in Entomological Review, 46, 245 - 256]" type="journal article" year="1967">Nartshuk, 1967</bibRefCitation>
: 426
</treatmentCitation>
</taxonomicName>
.
<emphasis id="D8DCC07CFFAAFFD477E837DEFD93E1D6" bold="true" box="[492,627,993,1015]" pageId="1" pageNumber="352">
<typeStatus id="3513A2CCFFAAFFD477E837DEFDC1E1D6" box="[492,545,993,1015]" pageId="1" pageNumber="352">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="2DA867EDFFAAFFD4748437DDFC11E1D6" ID-CoL="8BSFM" authority="Stackelberg, 1955" authorityName="Stackelberg. Scale" authorityYear="1955" box="[640,1009,993,1015]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="rossicus">
<emphasis id="D8DCC07CFFAAFFD4748437DDFCD5E1D6" box="[640,821,994,1015]" italics="true" pageId="1" pageNumber="352">Dicraeus rossicus</emphasis>
<bibRefCitation id="8E39619FFFAAFFD4753F37DEFC11E1D6" author="Stackelberg, A. A." box="[827,1009,993,1015]" pageId="1" pageNumber="352" pagination="328 - 333" refId="ref13003" refString="Stackelberg, A. A. (1955) New species of flies of the group Acalyptrata (Diptera) from the Leningrad province. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, 18, 328 - 333. [In Russian]" type="journal article" year="1955">Stackelberg, 1955</bibRefCitation>
</taxonomicName>
(original designation).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFAAFFD47693301BFEA6E736" pageId="1" pageNumber="352" type="diagnosis">
<paragraph id="EA171C6EFFAAFFD47693301BFEA6E736" blockId="1.[151,1436,1060,1303]" pageId="1" pageNumber="352">
<emphasis id="D8DCC07CFFAAFFD47693301BFEF1E61C" bold="true" box="[151,273,1060,1085]" pageId="1" pageNumber="352">Diagnosis.</emphasis>
Gena broad, at least 0.5 times as wide as first flagellomere; ocellar setae reclinate, convergent to cruciate (rarely parallel); first flagellomere rounded or somewhat angulate apicodorsally; wing with C usually extending to tip of M
<subScript id="762C1E2BFFAAFFD4778A3044FE46E6A8" attach="left" box="[398,422,1147,1161]" fontSize="6" pageId="1" pageNumber="352">1+2</subScript>
, sometimes ending halfway between R
<subScript id="762C1E2BFFAAFFD4756D3044FC61E6A8" attach="left" box="[873,897,1147,1161]" fontSize="6" pageId="1" pageNumber="352">4+5</subScript>
and M
<subScript id="762C1E2BFFAAFFD475D13044FC0EE6A8" attach="left" box="[981,1006,1147,1161]" fontSize="6" pageId="1" pageNumber="352">1+2</subScript>
; R
<subScript id="762C1E2BFFAAFFD472153044FBC9E6A8" attach="left" box="[1041,1065,1147,1161]" fontSize="6" pageId="1" pageNumber="352">2+3</subScript>
long, second costal sector more than 2.5 times length of third (rarely 2.0–2.5 times length of third); male genitalia usually large, epandrium prominent, semicircular; surstylus developed, much diversified in shape; male cercus absent, reduced or present as a narrow projection; hypandrium closed posteriorly (
<bibRefCitation id="8E39619FFFAAFFD4750230E5FC27E6D2" author="Andersson, H." box="[774,967,1242,1267]" pageId="1" pageNumber="352" pagination="1 - 200" refId="ref11473" refString="Andersson, H. (1977) Taxonomic and phylogenetic studies on Chloropidae (Diptera) with special reference to Old World genera. Entomologica Scandinavica, Supplement, 8, 1 - 200." type="journal article" year="1977">Andersson 1977</bibRefCitation>
;
<bibRefCitation id="8E39619FFFAAFFD475DC30E5FB66E6D2" author="Kanmiya, K." box="[984,1158,1242,1267]" pageId="1" pageNumber="352" pagination="1 - 370" refId="ref12164" refString="Kanmiya, K. (1983) A systematic study of the Japanese Chloropidae (Diptera). Memoirs of the Entomological Society of Washington, 11, 1 - 370." type="journal article" year="1983">Kanmiya 1983</bibRefCitation>
;
<bibRefCitation id="8E39619FFFAAFFD4729330E4FAC3E6D2" author="Ismay, J. W." box="[1175,1315,1243,1267]" pageId="1" pageNumber="352" pagination="5 - 17" refId="ref12100" refString="Ismay, J. W. (1984) New species of Dicraeus (Diptera: Chloropidae) from New Guinea and New Ireland. Esakia, 21, 5 - 17." type="journal article" year="1984">Ismay 1984</bibRefCitation>
;
<bibRefCitation id="8E39619FFFAAFFD4733030E5FEDAE736" author="Barrie, C. L. &amp; Wheeler T. A." pageId="1" pageNumber="352" pagination="375 - 395" refId="ref11508" refString="Barrie, C. L. &amp; Wheeler T. A. (2016) Revision of the Nearctic species of Dicraeus Loew (Diptera: Chloropidae). Canadian Entomologiist, 148, 375 - 395. https: // doi. org / 10.4039 / tce. 2015.74" type="journal article" year="2016">Barrie &amp; Wheeler 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

243
data/62/01/AD/6201AD78FFAAFFD776933154FA7CE045.xml Normal file
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<subSubSection id="A2B24FE5FFAAFFD476933154FC1EE7A4" box="[151,1022,1386,1413]" pageId="1" pageNumber="352" type="nomenclature">
<paragraph id="EA171C6EFFAAFFD476933154FC1EE7A4" blockId="1.[151,1022,1386,1413]" box="[151,1022,1386,1413]" pageId="1" pageNumber="352">
<heading id="B15FAB02FFAAFFD476933154FC1EE7A4" bold="true" box="[151,1022,1386,1413]" fontSize="11" level="1" pageId="1" pageNumber="352" reason="1">
<emphasis id="D8DCC07CFFAAFFD476933154FC1EE7A4" bold="true" box="[151,1022,1386,1413]" pageId="1" pageNumber="352">
Key to species of
<taxonomicName id="2DA867EDFFAAFFD477693155FE3AE7A5" ID-CoL="62QZH" ID-ENA="540857" authorityName="Loew" authorityYear="1873" box="[365,474,1386,1412]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="D8DCC07CFFAAFFD477693155FE3AE7A5" bold="true" box="[365,474,1386,1412]" italics="true" pageId="1" pageNumber="352">Dicraeus</emphasis>
</taxonomicName>
from
<collectingCountry id="92BF5CFEFFAAFFD474233154FD94E7A4" box="[551,628,1387,1413]" name="China" pageId="1" pageNumber="352">China</collectingCountry>
(modified from
<bibRefCitation id="8E39619FFFAAFFD4753B3154FC16E7A4" author="Kanmiya, K." box="[831,1014,1387,1413]" pageId="1" pageNumber="352" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya 1971</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFAAFFD776933195FA7CE045" lastPageId="2" lastPageNumber="353" pageId="1" pageNumber="352" type="key">
<keyStep id="515C04CBFFAAFFD476933195FA7BE7FC" pageId="1" pageNumber="352">
<paragraph id="EA171C6EFFAAFFD476933195FA7CE79E" blockId="1.[151,1436,1450,2029]" box="[151,1436,1450,1471]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD476933195FA7CE79E" box="[151,1436,1450,1471]" pageId="1" pageNumber="352">1. Scutum background yellow or brown; thoracic pleuron yellow, partly reddish yellow or brown........................ 2</keyLead>
</paragraph>
<paragraph id="EA171C6EFFAAFFD4769331F7FA7BE7FC" blockId="1.[151,1436,1450,2029]" box="[151,1435,1480,1501]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD4769331F7FA7BE7FC" box="[151,1435,1480,1501]" pageId="1" pageNumber="352">-. Scutum entirely black; thoracic pleuron entirely or largely black................................................ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="515C04CBFFAAFFD4769331DAFA7CE439" pageId="1" pageNumber="352">
<paragraph id="EA171C6EFFAAFFD4769331DAFA7BE7DB" blockId="1.[151,1436,1450,2029]" box="[151,1435,1509,1530]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD4769331DAFA7BE7DB" box="[151,1435,1509,1530]" pageId="1" pageNumber="352">2. Scutellum entirely yellow............................................................................... 3</keyLead>
</paragraph>
<paragraph id="EA171C6EFFAAFFD47693323DFA7CE439" blockId="1.[151,1436,1450,2029]" box="[151,1436,1538,1560]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD47693323DFA7CE439" box="[151,1436,1538,1560]" pageId="1" pageNumber="352">
-. Scutellum entirely blackish brown (figs. 1, 12–15).........................................
<taxonomicName id="2DA867EDFFAAFFD472B1323DFAA9E436" ID-CoL="9HVHC" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[1205,1353,1538,1559]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="crassispinus" status="sp. nov.">
<emphasis id="D8DCC07CFFAAFFD472B1323DFAA9E436" box="[1205,1353,1538,1559]" italics="true" pageId="1" pageNumber="352">D. crassispinus</emphasis>
</taxonomicName>
<emphasis id="D8DCC07CFFAAFFD4734B323DFA7CE439" bold="true" box="[1359,1436,1538,1560]" pageId="1" pageNumber="352">
<taxonomicNameLabel id="C3EF7D07FFAAFFD4734B323DFA7CE439" box="[1359,1436,1538,1560]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="515C04CBFFAAFFD47693321FFA7CE451" pageId="1" pageNumber="352">
<paragraph id="EA171C6EFFAAFFD47693321FFA7BE414" blockId="1.[151,1436,1450,2029]" box="[151,1435,1568,1589]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD47693321FFA7BE414" box="[151,1435,1568,1589]" pageId="1" pageNumber="352">3. Frons yellow; ocellar triangle reaching anterior 2/3 of frons.................................................... 4</keyLead>
</paragraph>
<paragraph id="EA171C6EFFAAFFD476933202FA7CE451" blockId="1.[151,1436,1450,2029]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD476933202FA7CE451" pageId="1" pageNumber="352">
-. Frons brown with anterior 1/2 yellow; ocellar triangle reaching anterior margin of frons (figs. 4, 24–27).................................................................................................
<taxonomicName id="2DA867EDFFAAFFD472893265FAA8E44E" ID-CoL="9HS9M" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[1165,1352,1626,1647]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="liupanshanensis" status="sp. nov.">
<emphasis id="D8DCC07CFFAAFFD472893265FAA8E44E" box="[1165,1352,1626,1647]" italics="true" pageId="1" pageNumber="352">D. liupanshanensis</emphasis>
</taxonomicName>
<emphasis id="D8DCC07CFFAAFFD4734B3265FA7CE451" bold="true" box="[1359,1436,1626,1648]" pageId="1" pageNumber="352">
<taxonomicNameLabel id="C3EF7D07FFAAFFD4734B3265FA7CE451" box="[1359,1436,1626,1648]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="515C04CBFFAAFFD476933247FA7CE501" pageId="1" pageNumber="352">
<paragraph id="EA171C6EFFAAFFD476933247FA7CE4E9" blockId="1.[151,1436,1450,2029]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD476933247FA7CE4E9" pageId="1" pageNumber="352">
4. Scutum yellow with 5 longitudinal stripes; surstylus broad basally, with 3 spinous setae, suddenly narrowed at middle and curved; mesolobus long, 2 times as long as surstylus, bent toward venter of epandrium (figs. 2, 16–19).........................................................................................................
<taxonomicName id="2DA867EDFFAAFFD472EF328DFAA9E4E6" ID-CoL="9HW4Z" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[1259,1353,1714,1735]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="curvus" status="sp. nov.">
<emphasis id="D8DCC07CFFAAFFD472EF328DFAA9E4E6" box="[1259,1353,1714,1735]" italics="true" pageId="1" pageNumber="352">D. curvus</emphasis>
</taxonomicName>
<emphasis id="D8DCC07CFFAAFFD4734B328DFA7CE4E9" bold="true" box="[1359,1436,1714,1736]" pageId="1" pageNumber="352">
<taxonomicNameLabel id="C3EF7D07FFAAFFD4734B328DFA7CE4E9" box="[1359,1436,1714,1736]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="EA171C6EFFAAFFD4769332EFFA7CE501" blockId="1.[151,1436,1450,2029]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD4769332EFFA7CE501" pageId="1" pageNumber="352">
-. Scutum yellow with 3 longitudinal stripes; surstylus broad distally, with some thickened setae on inner side of lateral and distal portions; cercus broad basally, distal 2/3 narrow, finger-like, directed medioventrally (figs. 5, 28–31)....................................................................................................
<taxonomicName id="2DA867EDFFAAFFD4729D3335FAA8E53E" ID-CoL="9HY2V" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[1177,1352,1802,1823]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="minutidentatus" status="sp. nov.">
<emphasis id="D8DCC07CFFAAFFD4729D3335FAA8E53E" box="[1177,1352,1802,1823]" italics="true" pageId="1" pageNumber="352">D. minutidentatus</emphasis>
</taxonomicName>
<emphasis id="D8DCC07CFFAAFFD4734B3335FA7CE501" bold="true" box="[1359,1436,1802,1824]" pageId="1" pageNumber="352">
<taxonomicNameLabel id="C3EF7D07FFAAFFD4734B3335FA7CE501" box="[1359,1436,1802,1824]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="515C04CBFFAAFFD476933317FA7BE556" pageId="1" pageNumber="352">
<paragraph id="EA171C6EFFAAFFD476933317FA7CE51C" blockId="1.[151,1436,1450,2029]" box="[151,1436,1832,1853]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD476933317FA7CE51C" box="[151,1436,1832,1853]" pageId="1" pageNumber="352">5. Thoracic pleuron entirely black or blackish brown with anepisternum and propleuron partly yellow..................... 6</keyLead>
</paragraph>
<paragraph id="EA171C6EFFAAFFD47693337AFA7BE556" blockId="1.[151,1436,1450,2029]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD47693337AFA7BE556" pageId="1" pageNumber="352">-. Thoracic pleuron black with a longitudinal yellow stripe running from posterior angle or lower half of postpronotal lobe to wing base............................................................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="515C04CBFFAAFFD7769333BFFA7CE2EB" lastPageId="2" lastPageNumber="353" pageId="1" pageNumber="352">
<paragraph id="EA171C6EFFAAFFD4769333BFFA7BE5B4" blockId="1.[151,1436,1450,2029]" box="[151,1435,1920,1941]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD4769333BFFA7BE5B4" box="[151,1435,1920,1941]" pageId="1" pageNumber="352">6. Palpus yellow; wing entirely hyaline...................................................................... 7</keyLead>
</paragraph>
<paragraph id="EA171C6EFFAAFFD4769333A2FA7CE593" blockId="1.[151,1436,1450,2029]" box="[151,1436,1949,1970]" pageId="1" pageNumber="352">
<keyLead id="5159BF5BFFAAFFD4769333A2FA7CE593" box="[151,1436,1949,1970]" pageId="1" pageNumber="352">
-. Palpus brown; wing infuscated with pale brown (figs. 6, 32, 38)...............................
<taxonomicName id="2DA867EDFFAAFFD472B033A2FA7CE593" ID-CoL="6D47G" authority="Kanmiya" authorityName="Kanmiya" authorityYear="1971" box="[1204,1436,1949,1970]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="D8DCC07CFFAAFFD472B033A2FADEE593" box="[1204,1342,1949,1970]" italics="true" pageId="1" pageNumber="352">D. fuscipennis</emphasis>
Kanmiya
</taxonomicName>
</keyLead>
</paragraph>
<footnote id="89B30060FFAAFFD476933385FA7BE5CC" pageId="1" pageNumber="352">
<paragraph id="EA171C6EFFAAFFD476933385FA7BE5CC" blockId="1.[151,1436,1450,2029]" pageId="1" pageNumber="352">
7. Ocellar triangle reaching anterior 2/3 of frons; surstylus with distal portion suddenly widened in lateral view; cercus without dentation distally (figs. 11, 37, 43)......................................................
<taxonomicName id="2DA867EDFFAAFFD472BD33E7FA7BE5CC" ID-CoL="8BSFM" authority="Stackelberg" authorityName="Stackelberg. Scale" authorityYear="1955" box="[1209,1435,2008,2029]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="rossicus">
<emphasis id="D8DCC07CFFAAFFD472BD33E7FAC4E5CC" box="[1209,1316,2008,2029]" italics="true" pageId="1" pageNumber="352">D. rossicus</emphasis>
Stackelberg
</taxonomicName>
</paragraph>
</footnote>
<paragraph id="EA171C6EFFA9FFD7769334A8FA7CE2EB" blockId="2.[151,1436,151,612]" pageId="2" pageNumber="353">
<keyLead id="5159BF5BFFA9FFD7769334A8FA7CE2EB" pageId="2" pageNumber="353">
-. Ocellar triangle reaching anterior 9/10 of frons; surstylus nearly parallel-sided, narrowed distally in lateral view; cercus with a dentation distally (figs. 3, 20–23)..........................................................
<taxonomicName id="2DA867EDFFA9FFD772DD348AFAA8E2EB" ID-CoL="9HYLD" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[1241,1352,181,202]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
<emphasis id="D8DCC07CFFA9FFD772DD348AFAA8E2EB" box="[1241,1352,181,202]" italics="true" pageId="2" pageNumber="353">D. dentatus</emphasis>
</taxonomicName>
<emphasis id="D8DCC07CFFA9FFD7734B348BFA7CE2EB" bold="true" box="[1359,1436,180,202]" pageId="2" pageNumber="353">
<taxonomicNameLabel id="C3EF7D07FFA9FFD7734B348BFA7CE2EB" box="[1359,1436,180,202]" pageId="2" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
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<keyStep id="515C04CBFFA9FFD7769334EDFA7BE31E" pageId="2" pageNumber="353">
<paragraph id="EA171C6EFFA9FFD7769334EDFA7BE325" blockId="2.[151,1436,151,612]" pageId="2" pageNumber="353">
<keyLead id="5159BF5BFFA9FFD7769334EDFA7BE325" pageId="2" pageNumber="353">
8. Antenna entirely black; thoracic pleuron with a longitudinal yellow stripe running from lower half of postpronotal lobe to wing base; femora brown with yellow distally (figs. 9, 35 41)........................................
<taxonomicName id="2DA867EDFFA9FFD772D134D0FA7BE325" ID-CoL="6D3VH" authority="Becker" authorityName="Becker" authorityYear="1911" box="[1237,1435,239,260]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="D8DCC07CFFA9FFD772D134D0FAB1E325" box="[1237,1361,239,260]" italics="true" pageId="2" pageNumber="353">D. orientalis</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EA171C6EFFA9FFD776933532FA7BE31E" blockId="2.[151,1436,151,612]" pageId="2" pageNumber="353">
<keyLead id="5159BF5BFFA9FFD776933532FA7BE31E" pageId="2" pageNumber="353">-. Antenna yellow with dorsal portion of first flagellomere brown; thoracic pleuron with a longitudinal yellow stripe running from posterior angle of postpronotal lobe to wing base; femora entirely yellow..................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="515C04CBFFA9FFD776933578FA7BE395" pageId="2" pageNumber="353">
<paragraph id="EA171C6EFFA9FFD776933578FA7CE35B" blockId="2.[151,1436,151,612]" pageId="2" pageNumber="353">
<keyLead id="5159BF5BFFA9FFD776933578FA7CE35B" pageId="2" pageNumber="353">
9. Scutum subshining, entirely and sparsely covered with brown pruinescence; epandrium blackish brown; surstylus broadened basally, narrowed at middle and curved in a S-shaped form in lateral view (figs. 10, 36, 42)......
<taxonomicName id="2DA867EDFFA9FFD7728A355AFA7CE35B" ID-CoL="35QRC" authority="Kanmiya" authorityName="Kanmiya" authorityYear="1971" box="[1166,1436,357,378]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="phyllostachyus">
<emphasis id="D8DCC07CFFA9FFD7728A355AFADDE35B" box="[1166,1341,357,378]" italics="true" pageId="2" pageNumber="353">D. phyllostachyus</emphasis>
Kanmiya
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EA171C6EFFA9FFD7769335BDFA7BE395" blockId="2.[151,1436,151,612]" pageId="2" pageNumber="353">
<keyLead id="5159BF5BFFA9FFD7769335BDFA7BE395" pageId="2" pageNumber="353">-. Scutum almost entirely shining, without prominent pruinescence; epandrium yellow to brownish yellow; surstylus nearly parallel-sided in lateral view.............................................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="515C04CBFFA9FFD776933582FA7BE3CE" pageId="2" pageNumber="353">
<paragraph id="EA171C6EFFA9FFD776933582FA7CE3F3" blockId="2.[151,1436,151,612]" box="[151,1436,445,466]" pageId="2" pageNumber="353">
<keyLead id="5159BF5BFFA9FFD776933582FA7CE3F3" box="[151,1436,445,466]" pageId="2" pageNumber="353">
10. Surstylus 2 times as height as epanderium, curved at middle in lateral view (figs. 7, 33, 39)...........
<taxonomicName id="2DA867EDFFA9FFD772CB3582FA7CE3F3" ID-CoL="35QQR" authority="Nartshuk" authorityName="Nartshuk, 2000:" authorityYear="1325" box="[1231,1436,445,466]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="D8DCC07CFFA9FFD772CB3582FADDE3F3" box="[1231,1341,445,466]" italics="true" pageId="2" pageNumber="353">D. malaisei</emphasis>
Nartshuk
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EA171C6EFFA9FFD7769335E5FA7BE3CE" blockId="2.[151,1436,151,612]" box="[151,1435,474,495]" pageId="2" pageNumber="353">
<keyLead id="5159BF5BFFA9FFD7769335E5FA7BE3CE" box="[151,1435,474,495]" pageId="2" pageNumber="353">-. Surstylus shorter than height of epanderium, straight in lateral view............................................. 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="515C04CBFFA9FFD7769335C8FA7CE045" pageId="2" pageNumber="353">
<paragraph id="EA171C6EFFA9FFD7769335C8FA7CE00B" blockId="2.[151,1436,151,612]" pageId="2" pageNumber="353">
<keyLead id="5159BF5BFFA9FFD7769335C8FA7CE00B" pageId="2" pageNumber="353">
11. Gena 0.25 times as deep as eye; palpus partly or distal 1/2 brown; all tarsomeres 2–5 brown; surstylus clavate in lateral view........................................................................................
<taxonomicName id="2DA867EDFFA9FFD772FD362AFA7CE00B" ID-CoL="35QQ7" authority="Duda" authorityName="Duda" authorityYear="1930" box="[1273,1436,533,554]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="D8DCC07CFFA9FFD772FD362AFA82E00B" box="[1273,1378,533,554]" italics="true" pageId="2" pageNumber="353">D. flavipes</emphasis>
Duda
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EA171C6EFFA9FFD77693360DFA7CE045" blockId="2.[151,1436,151,612]" pageId="2" pageNumber="353">
<keyLead id="5159BF5BFFA9FFD77693360DFA7CE045" pageId="2" pageNumber="353">
-. Gena 0.3–0.4 times as deep as eye; palpus entirely yellow; tarsi entirely yellow or only tarsomere 5 brown; surstylus parallelsided in lateral view (figs. 8, 34, 40).....................................................
<taxonomicName id="2DA867EDFFA9FFD772B53670FA7CE045" ID-CoL="35QQX" authority="Kanmiya" authorityName="Kanmiya" authorityYear="1971" box="[1201,1436,591,612]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="nartshukae">
<emphasis id="D8DCC07CFFA9FFD772B53670FADDE045" box="[1201,1341,591,612]" italics="true" pageId="2" pageNumber="353">D. nartshukae</emphasis>
Kanmiya
</taxonomicName>
</keyLead>
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<emphasis id="D8DCC07CFFADFFD37693316AFE91E74E" bold="true" box="[151,369,1365,1391]" italics="true" pageId="6" pageNumber="357">Dicraeus dentatus</emphasis>
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</paragraph>
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<subSubSection id="A2B24FE5FFADFFD376933148FED0E7B1" box="[151,304,1399,1424]" pageId="6" pageNumber="357" type="description">
<paragraph id="EA171C6EFFADFFD376933148FED0E7B1" blockId="6.[151,471,1365,1424]" box="[151,304,1399,1424]" pageId="6" pageNumber="357">
<figureCitation id="729300EBFFADFFD376933148FF3DE7B1" box="[151,221,1399,1424]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1268,1290]" captionTargetBox="[230,1360,680,1228]" captionTargetId="figure@2.[226,1360,665,1244]" captionTargetPageId="2" captionText="FIGURES 1–5. Dicraeus species, habitus. 1 D. crassispinus sp. nov.; 2 D. curvus sp. nov.; 3 D. dentatus sp. nov.; 4 D. liupanshanensis sp. nov.; 5 D. minutidentatus sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623715/files/figure.png" pageId="6" pageNumber="357">Figs 3</figureCitation>
,
<figureCitation id="729300EBFFADFFD376EE3147FED0E7B1" box="[234,304,1400,1424]" captionStart="FIGURES 20–23" captionStartId="7.[151,264,1959,1981]" captionTargetBox="[172,1416,434,1894]" captionTargetId="figure@7.[151,1436,409,1938]" captionTargetPageId="7" captionText="FIGURES 20–23. Dicraeus dentatus sp. nov. male. 20 epandrium, posterior view; 21 epandrium, lateral view; 22 hypandrium and phallic complex, ventral view; 23 hypandrium and phallic complex, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623725/files/figure.png" pageId="6" pageNumber="357">20–23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFADFFD376933180FBC8E464" pageId="6" pageNumber="357" type="diagnosis">
<paragraph id="EA171C6EFFADFFD376933180FBC8E464" blockId="6.[151,1437,1471,2037]" pageId="6" pageNumber="357">
<emphasis id="D8DCC07CFFADFFD376933180FEF1E7F9" bold="true" box="[151,273,1471,1496]" pageId="6" pageNumber="357">Diagnosis.</emphasis>
Gena 0.5 times as wide as first flagellomere. Ocellar triangle reaching anterior 0.9 of frons. Scutum and scutellum black. Legs largely yellow except for femora and hind tibia black with both ends yellow, tarsomeres 5 blackish brown. Surstylus nearly parallel-sided, round distally. Cercus long, with a small odontoid process distally, directed medioventrally. Postgonite bearing minute serrulations on apical margin.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFADFFDD76C33270FE42E38C" lastPageId="8" lastPageNumber="359" pageId="6" pageNumber="357" type="description">
<paragraph id="EA171C6EFFADFFD376C33270FC3FE446" blockId="6.[151,1437,1471,2037]" box="[199,991,1615,1640]" pageId="6" pageNumber="357">
<emphasis id="D8DCC07CFFADFFD376C33270FEB9E449" bold="true" box="[199,345,1615,1640]" pageId="6" pageNumber="357">Description.</emphasis>
Male (
<figureCitation id="729300EBFFADFFD377AC3270FE0BE449" box="[424,491,1615,1640]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1268,1290]" captionTargetBox="[230,1360,680,1228]" captionTargetId="figure@2.[226,1360,665,1244]" captionTargetPageId="2" captionText="FIGURES 1–5. Dicraeus species, habitus. 1 D. crassispinus sp. nov.; 2 D. curvus sp. nov.; 3 D. dentatus sp. nov.; 4 D. liupanshanensis sp. nov.; 5 D. minutidentatus sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623715/files/figure.png" pageId="6" pageNumber="357">Fig. 3</figureCitation>
). Body length
<quantity id="2D50B18BFFADFFD374963270FD0AE446" box="[658,746,1615,1640]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="357" unit="mm" value="2.5">2.5 mm</quantity>
, wing length
<quantity id="2D50B18BFFADFFD37586326FFC3BE446" box="[898,987,1616,1640]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="357" unit="mm" value="2.4">2.4 mm</quantity>
.
</paragraph>
<paragraph id="EA171C6EFFADFFD376C3324BFA93E5A9" blockId="6.[151,1437,1471,2037]" pageId="6" pageNumber="357">Head black, 1.2 times as wide as thorax, 0.85 times as long as high in profile, face yellow with brown sides, somewhat concave in lateral view, facial carina indistinct; frons brown with yellow anterior 1/6, 0.9 times as long as wide, projecting slightly in front of eye; gena broad, about 0.5 times as wide as first flagellomere, with few brown setulae on ventral margin; parafacial indistinct; clypeus light brown; vibrissal angle obtuse. Ocellar triangle brown, smooth, shiny, reaching anterior 0.9 of frons, with pointed apex; ocellar tubercle black. Postgena and occiput black. Cephalic setae and setulae black. Antenna yellow with thick grey pruinescence except for dorsodistal portion of first flagellomere blackish brown; first flagellomere 0.9 times as long as wide; arista brown with short brown pubescence. Proboscis brown to yellow with brown setulae; palpus brown with black setulae.</paragraph>
<paragraph id="EA171C6EFFADFFD276C333ABFB10E344" blockId="6.[151,1437,1471,2037]" lastBlockId="7.[151,1436,151,357]" lastPageId="7" lastPageNumber="358" pageId="6" pageNumber="357">Scutum black, 1.1 times as long as wide. Thoracic pleuron shiny brown. Scutellum black, 0.6 times as long as wide; apical scutellar seta slightly longer than scutellum; subapical scutellar seta as long as scutellum. Postscutellum black. Setae and setulae on thorax black; notopleurals 1+2, anterior notopleural seta as long as posterior notopleural seta; anterior postalar seta slightly longer posterior postalar seta; 1 pair of dorsocentral setae developed, as long as anterior postalar setae. Legs slender, coxae yellow except basal portion of mid coxa black, femora and hind tibia black with both ends yellow, fore and mid tibiae yellow, tarsi yellow with tarsomeres 5 blackish brown. Mid tibia with a short black subapical spur, which is shorter than diameter of tibia. Tibial organ long and narrow. Setulae on legs black. Wing 2.6 times as long as wide, hyaline; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 5: 1.7: 3; r-m at basal 0.6 of discal medial cell. Halter pale yellow.</paragraph>
<caption id="BED74CE6FFACFFD276933398FB25E5FD" httpUri="https://zenodo.org/record/2623725/files/figure.png" pageId="7" pageNumber="358" startId="7.[151,264,1959,1981]" targetBox="[172,1416,434,1894]" targetPageId="7">
<paragraph id="EA171C6EFFACFFD276933398FB25E5FD" blockId="7.[151,1435,1959,2012]" pageId="7" pageNumber="358">
<emphasis id="D8DCC07CFFACFFD276933398FEB4E59C" bold="true" box="[151,340,1959,1981]" pageId="7" pageNumber="358">FIGURES 20–23.</emphasis>
<taxonomicName id="2DA867EDFFACFFD2775E3396FDF3E59C" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[346,531,1959,1982]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="dentatus" status="sp. nov.">
<emphasis id="D8DCC07CFFACFFD2775E3396FDF3E59C" box="[346,531,1959,1982]" italics="true" pageId="7" pageNumber="358">Dicraeus dentatus</emphasis>
</taxonomicName>
<emphasis id="D8DCC07CFFACFFD2741D3398FD89E59C" bold="true" box="[537,617,1959,1981]" pageId="7" pageNumber="358">
<taxonomicNameLabel id="C3EF7D07FFACFFD2741D3398FD89E59C" box="[537,617,1959,1981]" pageId="7" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male. 20 epandrium, posterior view; 21 epandrium, lateral view; 22 hypandrium and phallic complex, ventral view; 23 hypandrium and phallic complex, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="EA171C6EFFA3FFDD76C334A8FC03E291" blockId="8.[151,1437,151,752]" box="[199,995,151,176]" pageId="8" pageNumber="359">Abdomen brown except for tergite 1 and venter yellow. Setulae brown.</paragraph>
<paragraph id="EA171C6EFFA3FFDD76C33483FD7BE3A9" blockId="8.[151,1437,151,752]" pageId="8" pageNumber="359">
Male genitalia (
<figureCitation id="729300EBFFA3FFDD77793483FDEAE2F5" box="[381,522,188,213]" captionStart="FIGURES 20–23" captionStartId="7.[151,264,1959,1981]" captionTargetBox="[172,1416,434,1894]" captionTargetId="figure@7.[151,1436,409,1938]" captionTargetPageId="7" captionText="FIGURES 20–23. Dicraeus dentatus sp. nov. male. 20 epandrium, posterior view; 21 epandrium, lateral view; 22 hypandrium and phallic complex, ventral view; 23 hypandrium and phallic complex, lateral view. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623725/files/figure.png" pageId="8" pageNumber="359">Figs. 20–23</figureCitation>
): Epandrium brown with long brown setulae, height 1.3 times width in profile; surstylus slightly shorter than depth of epandrium in profile, nearly parallel-sided, round distally. Cercus long, 0.4 times as long as surstylus, with a small odontoid process distally, directed medioventrally. Pre- and postgonites short, postgonite with few sensilla, round distally, bearing minute serrulations on apical margin; basiphallus longer than wide, cylindrical; phallapodeme sclerotized, long, basal stalk broad with narrow basally in lateral view. Hypandrium closed, apically forming an arch.
</paragraph>
<paragraph id="EA171C6EFFA3FFDD76C335ABFE42E38C" blockId="8.[151,1437,151,752]" box="[199,418,404,429]" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD76C335ABFEC5E38C" bold="true" box="[199,293,404,429]" pageId="8" pageNumber="359">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA3FFDD76C33588FD64E039" pageId="8" pageNumber="359" type="materials_examined">
<paragraph id="EA171C6EFFA3FFDD76C33588FD64E039" blockId="8.[151,1437,151,752]" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD76C33588FDC8E3F1" bold="true" box="[199,552,439,464]" pageId="8" pageNumber="359">
Type material.
<materialsCitation id="5AC01633FFA3FFDD778E3588FDC8E3F1" ID-GBIF-Occurrence="2234214327" box="[394,552,439,464]" collectingDate="2011-06-19" collectionCode="CAU" country="China" county="Tunyuan" location="Taiwan" municipality="Lushan Mountain" pageId="8" pageNumber="359" specimenCount="1" typeStatus="holotype">
<typeStatus id="3513A2CCFFA3FFDD778E3588FDC8E3F1" box="[394,552,439,464]" pageId="8" pageNumber="359" type="holotype">HOLOTYPE</typeStatus>
</materialsCitation>
</emphasis>
<emphasis id="D8DCC07CFFA3FFDD74333587FDACE3F1" box="[567,588,440,464]" italics="true" pageId="8" pageNumber="359">6</emphasis>
,
<collectingCountry id="92BF5CFEFFA3FFDD74663587FD5EE3F1" box="[610,702,440,464]" name="China" pageId="8" pageNumber="359">CHINA</collectingCountry>
,
<location id="EF774AB5FFA3FFDD74D43588FCC5E3F1" LSID="urn:lsid:plazi:treatment:6201AD78FFADFFDD7693316AFACCE0CE:EF774AB5FFA3FFDD74D43588FCC5E3F1" box="[720,805,439,464]" country="China" county="Tunyuan" municipality="Lushan Mountain" name="Taiwan" pageId="8" pageNumber="359">Taiwan</location>
,
<location id="EF774AB5FFA3FFDD75333587FC6CE3EE" LSID="urn:lsid:plazi:treatment:6201AD78FFADFFDD7693316AFACCE0CE:EF774AB5FFA3FFDD75333587FC6CE3EE" box="[823,908,440,463]" country="China" county="Tunyuan" municipality="Lushan Mountain" name="Nantou" pageId="8" pageNumber="359">Nantou</location>
,
<collectingMunicipality id="0A738614FFA3FFDD759B3588FB90E3F1" box="[927,1136,439,464]" pageId="8" pageNumber="359">Lushan Mountain</collectingMunicipality>
,
<collectingCounty id="037664E2FFA3FFDD72863587FB08E3F1" box="[1154,1256,440,464]" pageId="8" pageNumber="359">Tunyuan</collectingCounty>
,
<date id="9E163AAEFFA3FFDD72FE3587FA77E3F1" box="[1274,1431,440,464]" pageId="8" pageNumber="359" value="2011-06-19">
<collectingDate id="8E52C346FFA3FFDD72FE3587FA77E3F1" box="[1274,1431,440,464]" pageId="8" pageNumber="359" value="2011-06-19">19. VI. 2011</collectingDate>
</date>
, Xiaoyan Liu Leg. (CAU) (photo and abdomen dissected).
<materialsCitation id="5AC01633FFA3FFDD752135E3FD60E039" ID-GBIF-Occurrence="2234214341" collectingDate="2011-06-19" collectionCode="CAU" country="China" county="Tunyuan" location="Taiwan" municipality="Lushan Mountain" pageId="8" pageNumber="359" specimenCount="1" typeStatus="paratype">
<emphasis id="D8DCC07CFFA3FFDD752135E3FC29E3D4" bold="true" box="[805,969,476,501]" pageId="8" pageNumber="359">
<typeStatus id="3513A2CCFFA3FFDD752135E3FC29E3D4" box="[805,969,476,501]" pageId="8" pageNumber="359" type="paratype">PARATYPES</typeStatus>
</emphasis>
2
<emphasis id="D8DCC07CFFA3FFDD75E235E3FBF0E3D5" box="[998,1040,476,500]" italics="true" pageId="8" pageNumber="359">66</emphasis>
, same data as holotype (
<collectionCode id="8CB984ABFFA3FFDD732035E3FA85E3D5" box="[1316,1381,476,500]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="8" pageNumber="359">CAU</collectionCode>
). All type specimens were stored in 75% ethanol
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA3FFDD76C3361BFDFDE01C" box="[199,541,548,573]" pageId="8" pageNumber="359" type="distribution">
<paragraph id="EA171C6EFFA3FFDD76C3361BFDFDE01C" blockId="8.[151,1437,151,752]" box="[199,541,548,573]" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD76C3361BFE80E01C" bold="true" box="[199,352,548,573]" pageId="8" pageNumber="359">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFA3FFDD7763361BFE4AE01C" box="[359,426,548,573]" name="China" pageId="8" pageNumber="359">China</collectingCountry>
(
<collectingRegion id="286CD28CFFA3FFDD77BD361BFDF9E01C" box="[441,537,548,573]" country="China" name="Taiwan)" pageId="8" pageNumber="359">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA3FFDD76C33678FAB5E0ED" pageId="8" pageNumber="359" type="discussion">
<paragraph id="EA171C6EFFA3FFDD76C33678FAB5E0ED" blockId="8.[151,1437,151,752]" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD76C33678FEDBE041" bold="true" box="[199,315,583,608]" pageId="8" pageNumber="359">Remarks.</emphasis>
The new species is somewhat similar to
<taxonomicName id="2DA867EDFFA3FFDD750F3676FB8CE041" authority="Kanmiya, 1971" authorityName="Kanmiya" authorityYear="1971" box="[779,1132,583,608]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="nitidiventris">
<emphasis id="D8DCC07CFFA3FFDD750F3676FC54E041" box="[779,948,584,608]" italics="true" pageId="8" pageNumber="359">D. nitidiventris</emphasis>
<bibRefCitation id="8E39619FFFA3FFDD75B83678FB8CE041" author="Kanmiya, K." box="[956,1132,583,608]" pageId="8" pageNumber="359" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the following features: all femora black with both ends yellow; abdomen brown except for tergite 1 yellow; cercus long, 0.4 times as long as surstylus, with a small odontoid process distally. In
<taxonomicName id="2DA867EDFFA3FFDD72B536AEFABBE089" authorityName="Kanmiya" authorityYear="1971" box="[1201,1371,656,680]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="nitidiventris">
<emphasis id="D8DCC07CFFA3FFDD72B536AEFABBE089" box="[1201,1371,656,680]" italics="true" pageId="8" pageNumber="359">D. nitidiventris</emphasis>
</taxonomicName>
, only the fore femur is black; the abdomen is entirely black; and the cercus is small and vestigial (
<bibRefCitation id="8E39619FFFA3FFDD7299368BFAA8E0ED" author="Kanmiya, K." box="[1181,1352,692,717]" pageId="8" pageNumber="359" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFA3FFDD769336E8FACCE0CE" box="[151,1324,727,752]" pageId="8" pageNumber="359" type="etymology">
<paragraph id="EA171C6EFFA3FFDD769336E8FACCE0CE" blockId="8.[151,1437,151,752]" box="[151,1324,727,752]" pageId="8" pageNumber="359">
<emphasis id="D8DCC07CFFA3FFDD769336E8FEFEE0D1" bold="true" box="[151,286,727,752]" pageId="8" pageNumber="359">Etymology.</emphasis>
The specific name is from the Latin
<taxonomicName id="2DA867EDFFA3FFDD74BF36E7FCFAE0D1" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[699,794,728,752]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="359" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="D8DCC07CFFA3FFDD74BF36E7FCFAE0D1" box="[699,794,728,752]" italics="true" pageId="8" pageNumber="359">dentatus</emphasis>
</taxonomicName>
(“dentiform”), referring to the shape of cercus.
</paragraph>
</subSubSection>
</treatment>
</document>

188
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</mods:relatedItem>
<mods:classification id="7E6B9692460617E15F3920D6BD3FF134">journal article</mods:classification>
<mods:identifier id="E933E472680082EAA7F1E022B8EFCD4F" type="CLB-Dataset">27572</mods:identifier>
<mods:identifier id="E390B49A33526A0F44378C0CF8DADDB2" type="DOI">10.11646/zootaxa.4554.2.2</mods:identifier>
<mods:identifier id="3FB9B0985FE4DB6CEA4A9C1C20ADEA65" type="GBIF-Dataset">0a20dc7b-5322-4d72-9536-6a80d2d56b22</mods:identifier>
<mods:identifier id="07E96312E26D49427D6F5F9C73BA033E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D9FA4B2CB12C6CFE727D43158BC13792" type="Zenodo-Dep">2623713</mods:identifier>
<mods:identifier id="F516E8AB1A71C14DE78C2F1D91CC971F" type="ZooBank">9C8F643F-7A2A-47D2-AEBF-A384BAB26B6B</mods:identifier>
</mods:mods>
<treatment id="6201AD78FFAFFFD3769332E8FAB6E72C" ID-DOI="http://doi.org/10.5281/zenodo.5942655" ID-GBIF-Taxon="154532334" ID-Zenodo-Dep="5942655" LSID="urn:lsid:plazi:treatment:6201AD78FFAFFFD3769332E8FAB6E72C" httpUri="http://treatment.plazi.org/id/6201AD78FFAFFFD3769332E8FAB6E72C" lastPageId="6" lastPageNumber="357" pageId="4" pageNumber="355">
<subSubSection id="A2B24FE5FFAFFFD1769332E8FE21E4D3" box="[151,449,1751,1778]" pageId="4" pageNumber="355" type="nomenclature">
<paragraph id="EA171C6EFFAFFFD1769332E8FE21E4D3" blockId="4.[151,449,1751,1811]" box="[151,449,1751,1778]" pageId="4" pageNumber="355">
<heading id="B15FAB02FFAFFFD1769332E8FE21E4D3" bold="true" box="[151,449,1751,1778]" fontSize="11" level="1" pageId="4" pageNumber="355" reason="1">
<emphasis id="D8DCC07CFFAFFFD1769332E8FE21E4D3" bold="true" box="[151,449,1751,1778]" pageId="4" pageNumber="355">
<taxonomicName id="2DA867EDFFAFFFD1769332E8FEBCE4D0" ID-CoL="9HW4Z" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[151,348,1751,1777]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="355" phylum="Arthropoda" rank="species" species="curvus" status="sp. nov.">
<emphasis id="D8DCC07CFFAFFFD1769332E8FEBCE4D0" bold="true" box="[151,348,1751,1777]" italics="true" pageId="4" pageNumber="355">Dicraeus curvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3EF7D07FFAFFFD1776032E7FE21E4D3" box="[356,449,1752,1778]" pageId="4" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFAFFFD1769332C5FED0E532" box="[151,304,1786,1811]" pageId="4" pageNumber="355" type="description">
<paragraph id="EA171C6EFFAFFFD1769332C5FED0E532" blockId="4.[151,449,1751,1811]" box="[151,304,1786,1811]" pageId="4" pageNumber="355">
<figureCitation id="729300EBFFAFFFD1769332C5FF3EE532" box="[151,222,1786,1811]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1268,1290]" captionTargetBox="[230,1360,680,1228]" captionTargetId="figure@2.[226,1360,665,1244]" captionTargetPageId="2" captionText="FIGURES 1–5. Dicraeus species, habitus. 1 D. crassispinus sp. nov.; 2 D. curvus sp. nov.; 3 D. dentatus sp. nov.; 4 D. liupanshanensis sp. nov.; 5 D. minutidentatus sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623715/files/figure.png" pageId="4" pageNumber="355">Figs 2</figureCitation>
,
<figureCitation id="729300EBFFAFFFD176EE32C5FED0E532" box="[234,304,1786,1811]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1615,1637]" captionTargetBox="[161,1399,227,1574]" captionTargetId="figure@5.[151,1436,193,1593]" captionTargetPageId="5" captionText="FIGURES 16–19. Dicraeus curvus sp. nov. male. 16 epandrium, posterior view; 17 epandrium, lateral view; 18 hypandrium and phallic complex, ventral view; 19 hypandrium and phallic complex, lateral view. Abbreviations: bas, basiphallus; cer, cercus; ep, epandrium; hyp, hypandrium; phal, phallapodeme; pog, postgonite; prg, pregonite; sur, surstylus. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623723/files/figure.png" pageId="4" pageNumber="355">16–19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFAFFFD17693337DFB25E5E6" pageId="4" pageNumber="355" type="diagnosis">
<paragraph id="EA171C6EFFAFFFD17693337DFB25E5E6" blockId="4.[151,1436,1858,2027]" pageId="4" pageNumber="355">
<emphasis id="D8DCC07CFFAFFFD17693337DFEF1E57A" bold="true" box="[151,273,1858,1883]" pageId="4" pageNumber="355">Diagnosis.</emphasis>
Gena 0.8 times as wide as first flagellomere. Ocellar triangle reaching anterior 2/3 of frons. Scutum yellow with 5 longitudinal stripes. Scutellum yellow. Legs yellow except for fore femur brownish on anterior surface, all tarsomeres 5 brown. Surstylus broad basally, with 3 spinous setae, suddenly narrowed at middle and curved. Mesolobus (fused cerci) long, 2 times as long as surstylus, bent to venter of epandrium.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFAFFFD376C333EDFE42E1A1" lastPageId="6" lastPageNumber="357" pageId="4" pageNumber="355" type="description">
<paragraph id="EA171C6EFFAFFFD176C333EDFC3FE5CB" blockId="4.[151,1436,1858,2027]" box="[199,991,2002,2027]" pageId="4" pageNumber="355">
<emphasis id="D8DCC07CFFAFFFD176C333EDFEB9E5CA" bold="true" box="[199,345,2002,2027]" pageId="4" pageNumber="355">Description.</emphasis>
Male (
<figureCitation id="729300EBFFAFFFD177AC33EDFE0DE5CA" box="[424,493,2002,2027]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1268,1290]" captionTargetBox="[230,1360,680,1228]" captionTargetId="figure@2.[226,1360,665,1244]" captionTargetPageId="2" captionText="FIGURES 1–5. Dicraeus species, habitus. 1 D. crassispinus sp. nov.; 2 D. curvus sp. nov.; 3 D. dentatus sp. nov.; 4 D. liupanshanensis sp. nov.; 5 D. minutidentatus sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2623715/files/figure.png" pageId="4" pageNumber="355">Fig. 2</figureCitation>
). Body length 2.0 mm, wing length
<quantity id="2D50B18BFFAFFFD1758633ECFC3BE5CB" box="[898,987,2003,2027]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="355" unit="mm" value="1.8">1.8 mm</quantity>
.
</paragraph>
<caption id="BED74CE6FFAEFFD076933270FF26E4E0" httpUri="https://zenodo.org/record/2623723/files/figure.png" pageId="5" pageNumber="356" startId="5.[151,264,1615,1637]" targetBox="[161,1399,227,1574]" targetPageId="5">
<paragraph id="EA171C6EFFAEFFD076933270FF26E4E0" blockId="5.[151,1436,1615,1729]" pageId="5" pageNumber="356">
<emphasis id="D8DCC07CFFAEFFD076933270FEB5E444" bold="true" box="[151,341,1615,1637]" pageId="5" pageNumber="356">FIGURES 16–19.</emphasis>
<taxonomicName id="2DA867EDFFAEFFD07758326FFDE4E444" authority="Liu &amp; Yang &amp; Nartshuk, 2019" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[348,516,1616,1637]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="curvus" status="sp. nov.">
<emphasis id="D8DCC07CFFAEFFD07758326FFDE4E444" box="[348,516,1616,1637]" italics="true" pageId="5" pageNumber="356">Dicraeus curvus</emphasis>
</taxonomicName>
<emphasis id="D8DCC07CFFAEFFD0740F3270FDBDE444" bold="true" box="[523,605,1615,1637]" pageId="5" pageNumber="356">
<taxonomicNameLabel id="C3EF7D07FFAEFFD0740F3270FDBDE444" box="[523,605,1615,1637]" pageId="5" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male. 16 epandrium, posterior view; 17 epandrium, lateral view; 18 hypandrium and phallic complex, ventral view; 19 hypandrium and phallic complex, lateral view. Abbreviations: bas, basiphallus; cer, cercus; ep, epandrium; hyp, hypandrium; phal, phallapodeme; pog, postgonite; prg, pregonite; sur, surstylus. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="EA171C6EFFAEFFD076C332D2FA60E5FF" blockId="5.[151,1437,1773,2014]" pageId="5" pageNumber="356">Head yellow, 1.1 times as wide as thorax, 0.7 times as long as high in profile, face somewhat concave in lateral view, facial carina indistinct; frons as long as wide, projecting slightly in front of eye; gena broad, about 0.8 times as wide as first flagellomere, with few black setulae on ventral margin; parafacial indistinct; clypeus light brown; vibrissal angle obtuse. Ocellar triangle smooth, shiny, reaching anterior 2/3 of frons, with pointed apex; ocellar tubercle black. Occiput black. Cephalic setae and setulae black. Antenna yellow with thick grey pruinescence except for basodorsal portion of first flagellomere brown; first flagellomere 0.9 times as long as wide; arista brown with short brown pubescence. Proboscis yellow with yellowish brown setulae; palpus yellow with black setulae.</paragraph>
<paragraph id="EA171C6EFFADFFD376C334A7FDDEE01C" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="357">Scutum as long as wide, largely brown except yellow laterally, with 5 longitudinal stripes, 3 central stripes blackish brown, nearly coalesced anteriorly, outer lateral stripes reddish yellow, short and narrow. Postpronotal lobe yellow with a brownish spot. Thoracic pleuron shiny yellow except for anteroventral portion of anepisternum, ventral 2/3 of katepisternum and posteroventral portion of meron brown; anteroventral margin of anepimeron yellowish brown. Scutellum yellow, 0.7 times as long as wide; apical scutellar seta longer than scutellum; subapical scutellar seta as long as scutellum. Postscutellum blackish brown. Setae and setulae on thorax black; notopleurals 1+2, anterior notopleural seta as long as posterior notopleural seta; anterior postalar seta as long as posterior postalar seta; 1 pair of dorsocentral setae developed, 2 times as long as anterior postalar setae. Legs yellow except fore femur brownish on anterior suface, all tarsomeres 5 brown. Mid tibia with a short black subapical spur, which is shorter than diameter of tibia. Tibial organ long and narrow. Setulae on legs brown. Wing 2.3 times as long as wide, hyaline; veins brown. Relative lengths of 2nd: 3rd: 4th costal sections = 6: 1.4: 1; r-m at basal 0.45 of discal medial cell. Halter pale yellow.</paragraph>
<paragraph id="EA171C6EFFADFFD376C33678FD59E0A4" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="357">Abdomen brown except for tergites 1+2 yellow, with brown distal and lateral margins; tergite 5 yellow with a black medial spot; venter yellow. Setulae brown.</paragraph>
<paragraph id="EA171C6EFFADFFD376C336B0FEB1E17C" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="357">
Male genitalia (
<figureCitation id="729300EBFFADFFD3777A36B0FDEBE089" box="[382,523,655,680]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1615,1637]" captionTargetBox="[161,1399,227,1574]" captionTargetId="figure@5.[151,1436,193,1593]" captionTargetPageId="5" captionText="FIGURES 16–19. Dicraeus curvus sp. nov. male. 16 epandrium, posterior view; 17 epandrium, lateral view; 18 hypandrium and phallic complex, ventral view; 19 hypandrium and phallic complex, lateral view. Abbreviations: bas, basiphallus; cer, cercus; ep, epandrium; hyp, hypandrium; phal, phallapodeme; pog, postgonite; prg, pregonite; sur, surstylus. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/2623723/files/figure.png" pageId="6" pageNumber="357">Figs. 16–19</figureCitation>
): Epandrium yellow with long brown setulae, height 1.3 times width in lateral view; surstylus slightly shorter than depth of epandrium in lateral view, basally broad with 3 spinous setae, suddenly narrowed at middle and curved. Mesolobus (fused cerci) long, 2 times as long as surstylus, bent to venter of epandrium. Pre- and postgonites short, postgonite round distally, with few sensilla; basiphallus longer than wide, cylindrical; phallapodeme sclerotized, long, with basal stalk broad in lateral view. Hypandrium closed, apically forming an arch.
</paragraph>
<paragraph id="EA171C6EFFADFFD376C33758FE42E1A1" blockId="6.[151,1437,151,1293]" box="[199,418,871,896]" pageId="6" pageNumber="357">
<emphasis id="D8DCC07CFFADFFD376C33758FEC5E1A1" bold="true" box="[199,293,871,896]" pageId="6" pageNumber="357">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFADFFD376C337B3FD64E1CC" pageId="6" pageNumber="357" type="materials_examined">
<paragraph id="EA171C6EFFADFFD376C337B3FD64E1CC" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="357">
<emphasis id="D8DCC07CFFADFFD376C337B3FDC3E184" bold="true" box="[199,547,908,933]" pageId="6" pageNumber="357">
Type material.
<materialsCitation id="5AC01633FFADFFD3778137B3FDC3E184" ID-GBIF-Occurrence="2234214330" box="[389,547,908,933]" collectingDate="2007-04-17" collectionCode="CAU" country="China" county="Xishuangbanna" location="Yexianggu" municipality="Mengyang" pageId="6" pageNumber="357" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="3513A2CCFFADFFD3778137B3FDC3E184" box="[389,547,908,933]" pageId="6" pageNumber="357" type="holotype">HOLOTYPE</typeStatus>
</materialsCitation>
</emphasis>
<emphasis id="D8DCC07CFFADFFD3742B37B3FDA4E185" box="[559,580,908,932]" italics="true" pageId="6" pageNumber="357">6</emphasis>
,
<collectingCountry id="92BF5CFEFFADFFD3745337B3FD53E185" box="[599,691,908,932]" name="China" pageId="6" pageNumber="357">CHINA</collectingCountry>
,
<collectingRegion id="286CD28CFFADFFD374C737B2FCFCE185" box="[707,796,909,932]" country="China" name="Yunnan" pageId="6" pageNumber="357">Yunnan</collectingRegion>
,
<collectingCounty id="037664E2FFADFFD3752F37B3FC3CE184" box="[811,988,908,933]" pageId="6" pageNumber="357">Xishuangbanna</collectingCounty>
,
<collectingMunicipality id="0A738614FFADFFD375E837B2FB87E184" box="[1004,1127,909,933]" pageId="6" pageNumber="357">Mengyang</collectingMunicipality>
,
<location id="EF774AB5FFADFFD3727237B3FB11E184" LSID="urn:lsid:plazi:treatment:6201AD78FFAFFFD3769332E8FAB6E72C:EF774AB5FFADFFD3727237B3FB11E184" box="[1142,1265,908,933]" country="China" county="Xishuangbanna" municipality="Mengyang" name="Yexianggu" pageId="6" pageNumber="357" stateProvince="Yunnan">Yexianggu</location>
,
<date id="9E163AAEFFADFFD3730537B3FA78E185" box="[1281,1432,908,932]" pageId="6" pageNumber="357" value="2007-04-17">
<collectingDate id="8E52C346FFADFFD3730537B3FA78E185" box="[1281,1432,908,932]" pageId="6" pageNumber="357" value="2007-04-17">17. IV. 2007</collectingDate>
</date>
, Wenliang Li Leg. (CAU).
<materialsCitation id="5AC01633FFADFFD377CC3790FD60E1CC" ID-GBIF-Occurrence="2234214329" collectingDate="2007-04-17" collectionCode="CAU" country="China" county="Xishuangbanna" location="Yexianggu" municipality="Mengyang" pageId="6" pageNumber="357" specimenCode="PARATYPE 1" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCode id="BA0EB415FFADFFD377CC3790FD94E1E9" box="[456,628,943,968]" collectionCode="CAU" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="6" pageNumber="357">
<emphasis id="D8DCC07CFFADFFD377CC3790FDBCE1E9" bold="true" box="[456,604,943,968]" pageId="6" pageNumber="357">PARATYPE</emphasis>
1
</specimenCode>
<emphasis id="D8DCC07CFFADFFD37479378FFD72E1E9" box="[637,658,944,968]" italics="true" pageId="6" pageNumber="357">6</emphasis>
, same data as holotype (
<collectionCode id="8CB984ABFFADFFD375B7378FFC11E1E9" box="[947,1009,944,968]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="6" pageNumber="357">CAU</collectionCode>
) (photo and abdomen dissected). All type specimens were stored in 75% ethanol
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFADFFD376C337C8FDC1E631" box="[199,545,1015,1040]" pageId="6" pageNumber="357" type="distribution">
<paragraph id="EA171C6EFFADFFD376C337C8FDC1E631" blockId="6.[151,1437,151,1293]" box="[199,545,1015,1040]" pageId="6" pageNumber="357">
<emphasis id="D8DCC07CFFADFFD376C337C8FE80E631" bold="true" box="[199,352,1015,1040]" pageId="6" pageNumber="357">Distribution.</emphasis>
<collectingCountry id="92BF5CFEFFADFFD3776337C8FE4AE631" box="[359,426,1015,1040]" name="China" pageId="6" pageNumber="357">China</collectingCountry>
(
<collectingRegion id="286CD28CFFADFFD377BD37C7FDF5E631" box="[441,533,1016,1040]" country="China" name="Yunnan" pageId="6" pageNumber="357">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFADFFD376C33023FF3FE6C9" pageId="6" pageNumber="357" type="discussion">
<paragraph id="EA171C6EFFADFFD376C33023FF3FE6C9" blockId="6.[151,1437,151,1293]" pageId="6" pageNumber="357">
<emphasis id="D8DCC07CFFADFFD376C33023FEDBE614" bold="true" box="[199,315,1052,1077]" pageId="6" pageNumber="357">Remarks.</emphasis>
The new species is somewhat similar to
<taxonomicName id="2DA867EDFFADFFD3750C3022FBA4E615" authority="Kanmiya, 1971" authorityName="Kanmiya" authorityYear="1971" box="[776,1092,1052,1077]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="ischaemi">
<emphasis id="D8DCC07CFFADFFD3750C3022FC6EE615" box="[776,910,1052,1076]" italics="true" pageId="6" pageNumber="357">D. ischaemi</emphasis>
<bibRefCitation id="8E39619FFFADFFD375913023FBA4E615" author="Kanmiya, K." box="[917,1092,1052,1077]" pageId="6" pageNumber="357" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya, 1971</bibRefCitation>
</taxonomicName>
, but can be separated from the latter by the following features: first flagellomere round dorsodistally; surstylus broad basally, with 3 spinous setae, suddenly narrowed at middle and curved; mesolobus (fused cerci) long, 2 times as long as surstylus, bent to venter of epandrium. In
<taxonomicName id="2DA867EDFFADFFD3776C30B6FE13E681" authorityName="Kanmiya" authorityYear="1971" box="[360,499,1160,1184]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="ischaemi">
<emphasis id="D8DCC07CFFADFFD3776C30B6FE13E681" box="[360,499,1160,1184]" italics="true" pageId="6" pageNumber="357">D. ischaemi</emphasis>
</taxonomicName>
, the first flagellomere has a dorsodistal angle; the surstylus is slender, slightly concave at middle, broadly round apically, and has 5 long thick setae; the cercus is slender, finger-like (
<bibRefCitation id="8E39619FFFADFFD373303093FF33E6C9" author="Kanmiya, K." pageId="6" pageNumber="357" pagination="157 - 180" refId="ref12133" refString="Kanmiya, K. (1971) Study on the genus Dicraeus Loew from Japan and Formosa (Diptera, Chloropidae). Mushi, 45, 157 - 180." type="journal article" year="1971">Kanmiya 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B24FE5FFADFFD376C330CBFAB6E72C" box="[199,1366,1268,1293]" pageId="6" pageNumber="357" type="etymology">
<paragraph id="EA171C6EFFADFFD376C330CBFAB6E72C" blockId="6.[151,1437,151,1293]" box="[199,1366,1268,1293]" pageId="6" pageNumber="357">
<emphasis id="D8DCC07CFFADFFD376C330CBFEAEE72C" bold="true" box="[199,334,1268,1293]" pageId="6" pageNumber="357">Etymology.</emphasis>
The specific name is from the Latin
<taxonomicName id="2DA867EDFFADFFD374EF30CAFCD6E72D" authorityName="Liu &amp; Yang &amp; Nartshuk" authorityYear="2019" box="[747,822,1269,1292]" class="Insecta" family="Chloropidae" genus="Dicraeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="curvus">
<emphasis id="D8DCC07CFFADFFD374EF30CAFCD6E72D" box="[747,822,1269,1292]" italics="true" pageId="6" pageNumber="357">curvus</emphasis>
</taxonomicName>
(“curved”), referring to the shape of mesolobus.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>New Coleoptera records from New Brunswick, Canada: Megalopodidae and Chrysomelidae</mods:title>
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Map 1
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<paragraph pageId="3" pageNumber="324">New Brunswick, Gloucester Co., Tracadie, 30.VII.1939, W. J. Brown (1, CNC). Kent Co., Kouchibouguac National Park, 5.VII.1977, S. J. Miller, 5786N (1, CNC); same locality, collector, and date, 5487A (1, CNC); same locality and collector, 9.VIII.1977, 5805B (1, CNC); same locality and collector, 16.VIII.1977, 6054V (2, CNC). Saint John Co., Saint John, Rockwood Park, 7.VIII.1953, J. F. Brimley (1, CNC). York Co. Fredericton, 16.VII.1928, W. J. Brown (4, CNC).</paragraph>
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No bionomic data were associated with the specimens. This species has been recorded from
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L.,
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus tremuloides" order="Malpighiales" pageId="3" pageNumber="324" phylum="Tracheophyta" rank="species" species="tremuloides">Populus tremuloides</taxonomicName>
Michx. and
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<bibRefCitation author="Clark, SM" journalOrPublisher=", The Coleopterists Society, United States, Sacramento, CA" pageId="15" pageNumber="336" title="Host plants of leaf beetles species occurring in the United States and Canada. Coleopterists Society Special Publication No. 2." year="2004">Clark et al. 2004</bibRefCitation>
).
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BC, AB, SK, MB, QC, NB (
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Map 1. Collection localities in New Brunswick, Canada of
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<mods:namePart>Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>905</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>235</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="161777011" LSID="urn:lsid:zoobank.org:act:E52BA14D-F9F4-429E-9A36-372006991B18" httpUri="http://treatment.plazi.org/id/62033938359C06EFCA33FEE332841ABE" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/E52BA14D-F9F4-429E-9A36-372006991B18" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mantadia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mantadia" status="sp. nov.">Pheidole mantadia</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">Figs 55A-F</figureCitation>
<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85G</figureCitation>
<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87K</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Madagascar. •1 major worker; Toamasina; Corridor Forestier Analamay-Mantadia, Tsaravoniana;
<geoCoordinate degrees="18.76124" direction="south" orientation="latitude" precision="1" value="-18.76124">-18.76124</geoCoordinate>
,
<geoCoordinate degrees="48.42134" direction="east" orientation="longitude" precision="1" value="48.42134">48.42134</geoCoordinate>
; alt. 939 m; 3 Dec 2012; B.L. Fisher et al. leg.; BLF30003, CASENT0297006 (CASC).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
.
</emphasis>
Madagascar. •1 w.; same data as for holotype; CASENT0923169 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
Head in full face view rectangular, slightly widening posteriorly; sides of the head with sparse, relatively short, erect pilosity; frons with thick, sparse, longitudinal rugae, interspaces smooth to indistinctly rugulose; inner hypostomal teeth distinct, low, thick, bulge-like, with rounded apex; outer hypostomal teeth higher, thick, with rounded tips, triangular; inner and outer teeth closely spaced and connected by indistinct concavity; propodeal spines moderately long, with base slightly wider than top; first gastral tergite shagreened.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
Whole head foveolate, frons and malar area with few additional, indistinct, longitudinal rugae, vertex with indistinct irregular rugae, genae with sculpture reduced to absent; promesonotum low, slightly convex, with steep posterior declivity; mesosoma foveolate, katepisternum, and mesonotum smooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 1): HL: 1.68; HW: 1.52; SL: 0.67; EL: 0.17; WL: 1.21; PSL: 0.24; MTL: 0.68; PNW: 0.7; PTW: 0.2; PPW: 0.61; CI: 90.5; SI: 43.9; PSLI: 14.5; PPI: 33.9; PNI: 46.1; MTI: 44.7.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view rectangular, slightly widening posteriorly, anterior of eyes relatively straight, posterior of eyes convex (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55B</figureCitation>
). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming indistinct transverse depression between frons and occipital lobes; inner hypostomal teeth invisible. Antennal scrobes indistinct and not delimited by carinulae. Sides of the head with sparse, relatively short, erect pilosity; whole head with dense, moderately long, suberect to erect pilosity. Occipital lobes shiny, with thick, sparse, irregular rugae, interspaces smooth to finely rugulose; frons with thick, sparse, longitudinal rugae, interspaces smooth to indistinctly rugulose; lateral sides of head with fine and sparse rugulae, area between rugulae finely foveolate; malar area with dense and thin longitudinal rugae, interspaces smooth; genae with fine, thin, and dense rugulae. Clypeus shiny and smooth, with thin, longitudinal rugulae on lateral sides; median notch present, narrow, and moderately deep; median longitudinal carina indistinct; lateral longitudinal carinae absent. Scape, when laid back, reaching the midlength of head; pilosity decumbent to erect (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55B, D</figureCitation>
). Inner hypostomal teeth distinct, low, thick, bulge-like, with rounded apex; outer hypostomal teeth higher, thick, with rounded tips, triangular (Fig.
<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85G</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum short, angular, and relatively low, posterior mesonotum steep, with small tubercle-like projections; promesonotal groove absent; metanotal groove absent or indistinct; propodeal spines moderately long, with base slightly wider than top, apex rounded; humeral area with small and flat tubercles (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D</figureCitation>
). Surface shiny, finely foveolate, katepisternum, anepisternum, and lateral surfaces of propodeum smooth, with few irregular, thick rugae. Pilosity moderately sparse, long, and erect (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny, finely foveolate; peduncle short, with small, rounded, horizontal lobes on its basal part; node moderately high and narrow, with convex apex, in rear view node relatively straight; pilosity moderately sparse and erect (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Shiny, with fine and sparse foveolae; short and rounded; in dorsal view sides with relatively long, acute, horn-like projections; pilosity long, moderately dense and erect (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
First gastral tergite shiny and shagreened; pilosity moderately dense, long, and erect (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, reddish brown to brown; legs yellow to brown (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55D, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1" start="Figure 55" startId="F55">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 55.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mantadia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mantadia">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mantadia</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0923169) and full-face view (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0297006).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 1): HL: 0.58; HW: 0.52; SL: 0.6; EL: 0.11; WL: 0.7; PSL: 0.1; MTL: 0.48; PNW: 0.37; PTW: 0.07; PPW: 0. 1; CI: 89.2; SI: 115.7; PSLI: 17.0; PPI: 64.4; PNI: 71.4; MTI: 91.7.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Occipital margin straight or indistinctly concave; occipital carina indistinct, weakly developed (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55A</figureCitation>
). Pilosity moderately sparse, long, and erect. Whole head foveolate, frons and malar area with few additional, indistinct, longitudinal rugae, vertex with indistinct irregular rugae, genae with sculpture reduced to absent. Clypeus shiny, foveolate to rugoreticulate; median longitudinal carina present; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity erect (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55A, C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum low, slightly convex, with steep posterior declivity; promesonotal groove absent; metanotal groove indistinct; propodeal spines short, triangular, apex acute (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C</figureCitation>
). Sculpture foveolate, katepisternum and mesonotum smooth. Pilosity sparse, long, and erect (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Peduncle moderately long and thin; node low, globular, and small; with few long, erect setae (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Short, low, and convex; with few short, erect setae (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
With sparse, erect pilosity (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, yellowish brown to brown (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Pheidole mantadia sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923169) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0297006)." figureDoi="10.3897/zookeys.905.39592.figure55" httpUri="https://binary.pensoft.net/fig/373102" pageId="0" pageNumber="1">55C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species was collected at 939 m in elevation, in rainforest. Nest was located in rotten log.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comments">
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mantadia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mantadia">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mantadia</emphasis>
</taxonomicName>
sp. nov. is most similar to
<taxonomicName lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="P. moramanaensis" pageId="0" pageNumber="1" rank="species" species="moramanaensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. moramanaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mantadia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mantadia">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mantadia</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName lsidName="P. curvistriat" pageId="0" pageNumber="1" rank="species" species="curvistriat">
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriat</emphasis>
</taxonomicName>
a sp. nov. by presence of fine foveolate sculpture on frons and mesosoma, low and thick inner hypostomal teeth, which are wider than high and thick, high outer hypostomal teeth; from
<taxonomicName lsidName="P. moramanaensis" pageId="0" pageNumber="1" rank="species" species="moramanaensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. moramanaensis</emphasis>
</taxonomicName>
sp. nov. by sparser and shorter pilosity on sides of head.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
<taxonomicName authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole mantadia" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mantadia">
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole mantadia</emphasis>
</taxonomicName>
sp. nov. can be distinguished from
<taxonomicName lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
</taxonomicName>
sp. nov. by frons and malar with indistinct, longitudinal rugae, and vertex with indistinct irregular and never arcuate rugae, absence of irregular rugae on mesosoma, and smooth mesonotum; from
<taxonomicName lsidName="P. moramanaensis" pageId="0" pageNumber="1" rank="species" species="moramanaensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. moramanaensis</emphasis>
</taxonomicName>
sp. nov. by frons and malar area with indistinct, longitudinal rugae, and never smooth clypeus.
</paragraph>
</subSubSection>
</treatment>
</document>

113
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<paragraph id="768DDF11582EAF1AD892ABF0D25D4984" pageNumber="425">
<taxonomicName id="CD5803A52A3E816C20A5B6590ADBC657" ID-CoL="7FDSK" authority="Thomas 1909" authorityName="Thomas" authorityYear="1909" class="Mammalia" family="Phyllostomidae" genus="Vampyressa" kingdom="Animalia" order="Chiroptera" pageNumber="425" phylum="Chordata" rank="species" species="nymphaea" subGenus="Metavampyressa">
<emphasis id="99A7EA8BC4EDB0A5FDF7E4764535EB5A" italics="true" pageNumber="425">Vampyressa (Metavampyressa) nymphaea</emphasis>
Thomas 1909
</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="C6F74F698B5788BF0E92D43743C06F05" pageNumber="425">
<treatmentCitation id="C4AC3DB22DF5080CD0667D1F28D5DEAB" author="Thomas" authority="Thomas 1909" authorityName="Thomas" authorityYear="1909" class="Mammalia" family="Phyllostomidae" genus="Vampyressa" journal="Ann. Mag. Nat. Hist., ser. 8" kingdom="Animalia" order="Chiroptera" page="230" pageNumber="425" phylum="Chordata" rank="species" species="nymphaea" subGenus="Metavampyressa" title="Vampyressa (Metavampyressa) nymphaea" volume="4" year="1909">
<taxonomicName id="F214C9E36749F04153BC2F0259E4771A" ID-CoL="7FDSK" authority="Thomas 1909" authorityName="Thomas" authorityYear="1909" class="Mammalia" family="Phyllostomidae" genus="Vampyressa" kingdom="Animalia" order="Chiroptera" pageNumber="425" phylum="Chordata" rank="species" species="nymphaea" subGenus="Metavampyressa">
<emphasis id="735B8620665E9016E7393C8E08700CDB" italics="true" pageNumber="425">Vampyressa (Metavampyressa) nymphaea</emphasis>
Thomas 1909
</taxonomicName>
,
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</treatmentCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="AC09778354D5B748AFEBD16CFA9AB20E" pageNumber="425">
<emphasis id="A08C1295C4C67185CC257878779CFE57" inLineHeading="true" pageNumber="425">Type Locality:</emphasis>
<materialsCitation id="C127B0585CC0F2D0877D0498E5004495" ID-GBIF-Occurrence="4867283403" country="Colombia" pageNumber="425" stateProvince="Choco">
<collectingCountry id="9FF2D08C52E3C67BE4152217CC37BEA6" name="Colombia">Colombia</collectingCountry>
,
<collectingRegion id="C40BCDC689E2598D1E8CC5B2BD3C10AF" country="Colombia" name="Choco">Chocó</collectingRegion>
, Novita (San Juan River).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D29D95B25B58A48BD92F5A09D4D73843" pageNumber="425" type="vernacular_names">
<paragraph id="1D3F8719A088DF04D1882A17510BC256" pageNumber="425">
<emphasis id="1D40609398FDFC2E7CB592D18A8E948C" inLineHeading="true" pageNumber="425">Vernacular Names:</emphasis>
<vernacularName id="19C6BC3BEAF7C30BB0389A1CAB56176B" language="eng" pageNumber="425">Striped Yellow-eared Bat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="67AB20B5D2D81C33C30DEA193971F11A" pageNumber="425" type="distribution">
<paragraph id="24EB344BFFC4D5327C972C443FBE4C6B" pageNumber="425">
<emphasis id="0A149C85AAF375BE1ADEFFC2BEF3D8AB" inLineHeading="true" pageNumber="425">Distribution:</emphasis>
W
<collectingCountry id="CB450C53CADBFA0F74067B8886BE6801" name="Ecuador">Ecuador</collectingCountry>
to
<collectingCountry id="3BC3DCDB85A63EDC05CFF4AB2D4C534C" name="Nicaragua">Nicaragua</collectingCountry>
. A record from SE
<collectingCountry id="81D3391F68C4F985477B47E304617C31" name="Peru">Peru</collectingCountry>
is suspect.
</paragraph>
</subSubSection>
<subSubSection id="8750857C969DCEFF2CBACEAAC00E3413" pageNumber="425" type="conservation">
<paragraph id="C2DC47459B99FA5120F2B55BBF6413FF" pageNumber="425">
<emphasis id="36F443246F19F25DD76E46280DCD0733" inLineHeading="true" pageNumber="425">Conservation:</emphasis>
<collectionCode id="6CFDFEDC81F81592CF3A650893DBFFF1">IUCN</collectionCode>
2003 and
<collectionCode id="9D5E2DDF4612AE4D9610ED6CFA3FC94E">IUCN</collectionCode>
/
<collectionCode id="A6E729EA97F30B4BF06C244BBEEEC32D" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (2001) – Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="9D8DDD6B541DCE5C360132D933BA1FD9" pageNumber="425" type="discussion">
<paragraph id="90726C377E89ECDBBE69CCD858322C02" pageNumber="425">
<emphasis id="C48F0E6F07F8086A6789DC0F2BF264D5" inLineHeading="true" pageNumber="425">Discussion:</emphasis>
Subgenus
<emphasis id="0F7DA1AD78B045FA880083BDDF3EA5B6" italics="true" pageNumber="425">Metvampyressa</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

86
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<treatmentCitation id="9114AA0221822F50A61FBCBF1B26FC3F" ID-DOI="http://doi.org/10.5281/zenodo.4356857" ID-Zenodo-Dep="4356857" author="Linnaeus" httpUri="http://treatment.plazi.org/id/05F7E523BCB9EFC51DD2586BA42A5849" pageNumber="463" year="1753">
<bibCitation id="9028F1902AFE44922750D75228A80FF5" author="Linnaeus" pagination="463" title="Euphorbia amygdaloides" volumeTitle="Species Plantarum 1" year="1753">
<taxonomicName id="F415FA2E7E8C012C8523D21DDC0AE0B1" ID-CoL="6GZBZ" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="amygdaloides">
<emphasis id="280AC1C3D5D922FDAF092288EFFACD8C" bold="true" pageNumber="510">Euphorbia amygdaloides</emphasis>
Linnaeus
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,
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<emphasis id="1201C9F1AE86A42B7021A741BAD13D2D" italics="true" pageNumber="510">Species Plantarum</emphasis>
1
</volumeTitle>
: 463. 1753
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</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7E5D5631B678A19107B03F25D4629787" pageNumber="510" type="distribution">
<paragraph id="DAFF40224862706E2475EA87EE7184AA" pageNumber="510">&quot;Habitat in Gallia, Germania.&quot; RCN: 3554.</paragraph>
</subSubSection>
<subSubSection id="DFC6BADB5F1B3BA7D3A2F0C8A41B0E7C" pageNumber="510" type="materials_examined">
<paragraph id="918F942AD55D51777FF53525A50A8148" pageNumber="510">
<materialsCitation id="E496BCC38860DD8885E69CA88C315A7A" ID-GBIF-Occurrence="2611986342" collectionCode="LINN" specimenCode="630.71" typeStatus="lectotype">
<typeStatus id="595ACD21AAA897A92B9D49CA083F85CE">Lectotype</typeStatus>
(Geltman in
<emphasis id="7B7376ADBFE42D621FEA10FE7FE78EF2" italics="true" pageNumber="510">Novosti Sist. Vyssh. Rast.</emphasis>
34: 106. 2003): Herb. Linn. No. 630.71 (
<collectionCode id="AB9AD6FF6B358BA0568C238DE37CE938">LINN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0E1FC2D1C01D2C79502C60354C522A84" pageNumber="510" type="nomenclature">
<paragraph id="61838BF2F3F5C34BE77E979B5F10CF0F" pageNumber="510">
Current name:
<taxonomicName id="B85A9BA8C460F40910ABD208BCCFCF17" ID-CoL="6GZBZ" authority="L." authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="species" species="amygdaloides">
<emphasis id="2943E6357584DE2529C3108A25568A1B" bold="true" italics="true" pageNumber="510">Euphorbia amygdaloides</emphasis>
L.
</taxonomicName>
(
<taxonomicName id="B25EE99D2E19AB51FFEAF4C76A7AB765" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
).
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72
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subsp.
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subsp.
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Lyon 1916
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<emphasis id="BE0ED09C851CC07260C1F3D4991FB282" italics="true" pageNumber="756">Ratufa bicolor</emphasis>
subsp.
<emphasis id="3ADDAA56205A55962149F9CCEB50AA4B" italics="true" pageNumber="756">laenata</emphasis>
Miller 1903
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<emphasis id="D8D9EA8674064838FF6BFC22FE6C05A8" bold="true" box="[228,442,965,994]" italics="true" pageId="1" pageNumber="5">Phaonia cincta</emphasis>
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<taxonomicName id="2DAD4D1774064838FF18FC0EFDE8024F" ID-CoL="9HWF6" authority="Sorokina, 2015" authorityName="Sorokina" authorityYear="2015" box="[151,574,1001,1030]" class="Insecta" family="Muscidae" genus="Phaonia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="5" phylum="Arthropoda" rank="species" species="juglans" status="syn. nov.">
<emphasis id="D8D9EA8674064838FF18FC0EFEBB024F" box="[151,365,1001,1030]" italics="true" pageId="1" pageNumber="5">Phaonia juglans</emphasis>
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<taxonomicNameLabel id="C3EA57FD74064838FDC5FC0EFD13024C" box="[586,709,1001,1029]" pageId="1" pageNumber="5">syn. nov.</taxonomicNameLabel>
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, “in the South Austria”,
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35.33
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</materialsCitation>
.
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,
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(
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.
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<collectingCountry id="92BA760474064838FF18FAF5FEDE0366" box="[151,264,1298,1327]" name="Russia" pageId="1" pageNumber="5">RUSSIA</collectingCountry>
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reg.,
<collectingMunicipality id="0A76ACEE74064838FE77FAF6FD950367" box="[504,579,1297,1326]" pageId="1" pageNumber="5">Sochi</collectingMunicipality>
env.,
<location id="EF72604F74064838FD17FAF6FF1F0318" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FD17FAF6FF1F0318" country="Russia" latitude="43.55" longLatPrecision="685" longitude="39.85" municipality="Sochi" name="Akhun Mt." pageId="1" pageNumber="5" stateProvince="Krasnodar">Akhun Mt.</location>
, (
<geoCoordinate id="8F99505374064838FF65FAD3FE8F0318" box="[234,345,1331,1361]" degrees="43.55" direction="north" orientation="latitude" pageId="1" pageNumber="5" precision="555" value="43.55">
43.55
<emphasis id="D8D9EA8674064838FEBCFAD4FE960318" box="[307,320,1331,1361]" italics="true" pageId="1" pageNumber="5">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="8F99505374064838FEE3FAD3FE03031B" box="[364,469,1331,1362]" degrees="39.85" direction="east" orientation="longitude" pageId="1" pageNumber="5" precision="555" value="39.85">
39.85
<emphasis id="D8D9EA8674064838FE3CFAD4FE160318" box="[435,448,1331,1361]" italics="true" pageId="1" pageNumber="5">°</emphasis>
E
</geoCoordinate>
),
<date id="9E13105474064838FE78FAD2FD39031B" box="[503,751,1333,1362]" pageId="1" pageNumber="5" value="2006-10-20">
<collectingDate id="8E57E9BC74064838FE78FAD2FD39031B" box="[503,751,1333,1362]" pageId="1" pageNumber="5" value="2006-10-20">20 October 2006</collectingDate>
</date>
,
<collectorName id="4758534274064838FF18FABFFEFA033C" box="[151,300,1368,1397]" pageId="1" pageNumber="5">N. Vikhrev</collectorName>
,
<specimenCount id="FCABFD1D74064838FEB3FABFFEB2033D" box="[316,356,1368,1397]" count="1" pageId="1" pageNumber="5" type="male">1♂</specimenCount>
;
<collectingRegion id="2869F87674064838FEF8FABFFE3A033C" box="[375,492,1368,1397]" country="Russia" name="Adygeya" pageId="1" pageNumber="5">Adygeya</collectingRegion>
,
<collectingMunicipality id="0A76ACEE74064838FE72FABFFD7B033C" box="[509,685,1368,1397]" pageId="1" pageNumber="5">Dakhovskaya</collectingMunicipality>
env.,
<location id="EF72604F74064838FF18FA9CFF3B03D1" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FF18FA9CFF3B03D1" box="[151,237,1403,1432]" country="Russia" municipality="Dakhovskaya" name="Belaya" pageId="1" pageNumber="5" stateProvince="Adygeya">Belaya</location>
<collectionCode id="8CBCAE5174064838FF75FA9BFEC403D1" box="[250,274,1404,1432]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="5">R</collectionCode>
. valley,
<geoCoordinate id="8F99505374064838FE0EFA9CFE2803D1" box="[385,510,1402,1432]" degrees="44.199" direction="north" orientation="latitude" pageId="1" pageNumber="5" precision="55" value="44.199">
44.199
<emphasis id="D8D9EA8674064838FE56FA9DFE3003D1" box="[473,486,1402,1432]" italics="true" pageId="1" pageNumber="5">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="8F99505374064838FD85FA9CFD5203D1" box="[522,644,1402,1432]" degrees="40.170" direction="east" orientation="longitude" pageId="1" pageNumber="5" precision="55" value="40.17">
40.170
<emphasis id="D8D9EA8674064838FDEDFA9DFDB903D1" box="[610,623,1402,1432]" italics="true" pageId="1" pageNumber="5">°</emphasis>
E
</geoCoordinate>
,
<quantity id="2D559B7174064838FD1AFA9CFD3903D1" box="[661,751,1403,1432]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="1" pageNumber="5" unit="m" value="465.0">
<elevation id="6180D1A774064838FD1AFA9CFD3903D1" box="[661,751,1403,1432]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="1" pageNumber="5" unit="m" value="465.0">465 m</elevation>
</quantity>
,
<date id="9E13105474064838FF18FA79FE7403F2" box="[151,418,1438,1467]" pageId="1" pageNumber="5" value="2009-08-18" valueMax="2009-08-31" valueMin="2009-08-18">
<collectingDate id="8E57E9BC74064838FF18FA79FE7403F2" box="[151,418,1438,1467]" pageId="1" pageNumber="5" value="2009-08-18" valueMax="2009-08-31" valueMin="2009-08-18">18–31 August 2009</collectingDate>
</date>
, K.
<collectorName id="4758534274064838FE57FA79FDA703F2" box="[472,625,1438,1467]" pageId="1" pageNumber="5">Tomkovich</collectorName>
,
<specimenCount id="FCABFD1D74064838FD0FFA79FD7E03F2" box="[640,680,1438,1467]" count="1" pageId="1" pageNumber="5" type="female">1♀</specimenCount>
(new records for
<location id="EF72604F74064838FEC8FA26FD450397" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FEC8FA26FD450397" box="[327,659,1473,1502]" country="Russia" latitude="44.199" longLatPrecision="67" longitude="40.17" name="South European Russia" pageId="1" pageNumber="5" stateProvince="Adygeya">South European Russia</location>
);
<emphasis id="D8D9EA8674064838FD3FFA25FF37004B" italics="true" pageId="1" pageNumber="5">
<collectingRegion id="2869F87674064838FD3FFA25FF37004B" country="Russia" name="Mordoviya" pageId="1" pageNumber="5">Mordovia</collectingRegion>
</emphasis>
reg.,
<collectingRegion id="2869F87674064838FEB8FA02FDDD004B" box="[311,523,1509,1538]" country="Russia" name="Mordoviya" pageId="1" pageNumber="5">Mordovia State</collectingRegion>
<location id="EF72604F74064838FD96FA02FD390048" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FD96FA02FD390048" box="[537,751,1509,1537]" country="Russia" latitude="54.72" longLatPrecision="641" longitude="43.23" name="Nature Reserve" pageId="1" pageNumber="5" stateProvince="Mordoviya">Nature Reserve</location>
,
<geoCoordinate id="8F99505374064838FF18F9EFFED3006D" box="[151,261,1542,1573]" degrees="54.72" direction="north" orientation="latitude" pageId="1" pageNumber="5" precision="555" value="54.72">
54.72
<emphasis id="D8D9EA8674064838FF50F9E1FF3A006D" box="[223,236,1542,1572]" italics="true" pageId="1" pageNumber="5">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="8F99505374064838FE80F9EFFEAE006D" box="[271,376,1542,1573]" degrees="43.23" direction="east" orientation="longitude" pageId="1" pageNumber="5" precision="555" value="43.23">
43.23
<emphasis id="D8D9EA8674064838FED9F9E1FEB5006D" box="[342,355,1542,1572]" italics="true" pageId="1" pageNumber="5">°</emphasis>
E
</geoCoordinate>
,
<date id="9E13105474064838FE07F9EFFDF4006C" box="[392,546,1544,1573]" pageId="1" pageNumber="5" value="2019-07-07">
<collectingDate id="8E57E9BC74064838FE07F9EFFDF4006C" box="[392,546,1544,1573]" pageId="1" pageNumber="5" value="2019-07-07">7 July 2019</collectingDate>
</date>
,
<collectorName id="4758534274064838FDBEF9EFFD6B006C" box="[561,701,1544,1573]" pageId="1" pageNumber="5">A. Ruchin</collectorName>
,
<specimenCount id="FCABFD1D74064838FD43F9EFFD22006D" box="[716,756,1544,1573]" count="1" pageId="1" pageNumber="5" type="male">1♂</specimenCount>
(all
<collectionCode id="8CBCAE5174064838FF5FF9CCFEEA0001" box="[208,316,1579,1608]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="1" pageNumber="5">ZMUM</collectionCode>
);
<emphasis id="D8D9EA8674064838FED5F9CBFE120001" box="[346,452,1580,1608]" italics="true" pageId="1" pageNumber="5">
<collectingRegion id="2869F87674064838FED5F9CBFE120001" box="[346,452,1580,1608]" country="Russia" name="Moscow" pageId="1" pageNumber="5">Moscow</collectingRegion>
</emphasis>
reg.:
<location id="EF72604F74064838FD94F9CCFDBB0001" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FD94F9CCFDBB0001" box="[539,621,1579,1608]" country="Russia" latitude="54.86" longLatPrecision="640" longitude="38.55" name="Ozery" pageId="1" pageNumber="5" stateProvince="Moscow">Ozery</location>
(
<geoCoordinate id="8F99505374064838FD0AF9CCFD220001" box="[645,756,1578,1608]" degrees="54.86" direction="north" orientation="latitude" pageId="1" pageNumber="5" precision="555" value="54.86">
54.86
<emphasis id="D8D9EA8674064838FD40F9CDFD0A0001" box="[719,732,1578,1608]" italics="true" pageId="1" pageNumber="5">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="8F99505374064838FF18F9A9FED70022" box="[151,257,1613,1643]" degrees="38.55" direction="east" orientation="longitude" pageId="1" pageNumber="5" precision="555" value="38.55">
38.55
<emphasis id="D8D9EA8674064838FF50F9AAFF3A0022" box="[223,236,1613,1643]" italics="true" pageId="1" pageNumber="5">°</emphasis>
E
</geoCoordinate>
), reared, 1974,
<collectionCode id="8CBCAE5174064838FE5FF9A8FE310022" box="[464,487,1615,1643]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="5" type="Museum">V</collectionCode>
.
<collectorName id="4758534274064838FE7BF9A9FD890022" box="[500,607,1614,1643]" pageId="1" pageNumber="5">Kovalev</collectorName>
,
<specimenCount id="FCABFD1D74064838FDE0F9A9FD410022" box="[623,663,1614,1643]" count="1" pageId="1" pageNumber="5" type="male">1♂</specimenCount>
(
<collectionCode id="8CBCAE5174064838FD27F9A9FD330022" box="[680,741,1614,1643]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="5" type="Museum">ZIN</collectionCode>
);
<collectorName id="4758534274064838FF16F995FEA300C7" box="[153,373,1649,1679]" pageId="1" pageNumber="5">SW Moskovsky</collectorName>
,
<geoCoordinate id="8F99505374064838FE02F996FE2F00C7" box="[397,505,1648,1678]" degrees="55.58" direction="north" orientation="latitude" pageId="1" pageNumber="5" precision="555" value="55.58">
55.58
<emphasis id="D8D9EA8674064838FE5CF997FE3600C7" box="[467,480,1648,1678]" italics="true" pageId="1" pageNumber="5">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="8F99505374064838FD83F995FDA300C7" box="[524,629,1648,1679]" degrees="37.34" direction="east" orientation="longitude" pageId="1" pageNumber="5" precision="555" value="37.34">
37.34
<emphasis id="D8D9EA8674064838FDDBF997FDB700C7" box="[596,609,1648,1678]" italics="true" pageId="1" pageNumber="5">°</emphasis>
E
</geoCoordinate>
,
<date id="9E13105474064838FD01F995FF0C00FB" pageId="1" pageNumber="5" value="2016-07-29">
<collectingDate id="8E57E9BC74064838FD01F995FF0C00FB" pageId="1" pageNumber="5" value="2016-07-29">29 July 2016</collectingDate>
</date>
,
<collectorName id="4758534274064838FF7FF972FE6100FB" box="[240,439,1685,1714]" pageId="1" pageNumber="5">K. Tomkovich</collectorName>
,
<specimenCount id="FCABFD1D74064838FE42F972FE2300F8" box="[461,501,1685,1714]" count="1" pageId="1" pageNumber="5" type="female">1♀</specimenCount>
(new records for
<location id="EF72604F74064838FF18F95FFE34009C" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FF18F95FFE34009C" box="[151,482,1720,1749]" country="Russia" latitude="55.58" longLatPrecision="638" longitude="37.34" name="Central European Russia" pageId="1" pageNumber="5" stateProvince="Moscow">Central European Russia</location>
) (
<collectionCode id="8CBCAE5174064838FE74F95FFDBF009C" box="[507,617,1720,1749]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="1" pageNumber="5">ZMUM</collectionCode>
).
</paragraph>
<paragraph id="EA12369474064838FF18F93CFEAD0176" blockId="1.[151,756,1086,2031]" pageId="1" pageNumber="5">
<materialsCitation id="5AC53CC974064838FF18F93CFDA70152" collectingDate="2009-03-25" collectionCode="V" country="Spain" elevation="925" latitude="37.0" location="Granada" longLatPrecision="5573" longitude="-3.08" municipality="Granada Valor" pageId="1" pageNumber="5" specimenCount="1">
<collectingCountry id="92BA760474064838FF18F93CFF2200B1" box="[151,244,1755,1784]" name="Spain" pageId="1" pageNumber="5">SPAIN</collectingCountry>
,
<emphasis id="D8D9EA8674064838FE8EF93CFEAC00B1" box="[257,378,1755,1784]" italics="true" pageId="1" pageNumber="5">
<location id="EF72604F74064838FE8EF93CFEAC00B1" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FE8EF93CFEAC00B1" box="[257,378,1755,1784]" country="Spain" latitude="37.0" longLatPrecision="5573" longitude="-3.08" municipality="Granada Valor" name="Granada" pageId="1" pageNumber="5">Granada</location>
</emphasis>
reg.,
<collectingMunicipality id="0A76ACEE74064838FE4DF93CFD5300B1" box="[450,645,1755,1784]" pageId="1" pageNumber="5">Granada Valor</collectingMunicipality>
(
<geoCoordinate id="8F99505374064838FD1AF93CFD2200B1" box="[661,756,1754,1784]" degrees="37.0" direction="north" orientation="latitude" pageId="1" pageNumber="5" precision="5555" value="37.0">
37.0
<emphasis id="D8D9EA8674064838FD40F93DFD0A00B1" box="[719,732,1754,1784]" italics="true" pageId="1" pageNumber="5">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="8F99505374064838FF18F919FF290152" box="[151,255,1789,1819]" degrees="3.08" direction="west" orientation="longitude" pageId="1" pageNumber="5" precision="555" value="-3.08">
3.08
<emphasis id="D8D9EA8674064838FF40F91AFF0A0152" box="[207,220,1789,1819]" italics="true" pageId="1" pageNumber="5">°</emphasis>
W
</geoCoordinate>
),
<quantity id="2D559B7174064838FE96F919FEB90152" box="[281,367,1790,1819]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="1" pageNumber="5" unit="m" value="925.0">
<elevation id="6180D1A774064838FE96F919FEB90152" box="[281,367,1790,1819]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="1" pageNumber="5" unit="m" value="925.0">925 m</elevation>
</quantity>
,
<date id="9E13105474064838FEF1F919FD9A0152" box="[382,588,1790,1819]" pageId="1" pageNumber="5" value="2009-03-25">
<collectingDate id="8E57E9BC74064838FEF1F919FD9A0152" box="[382,588,1790,1819]" pageId="1" pageNumber="5" value="2009-03-25">25 March 2009</collectingDate>
</date>
,
<collectionCode id="8CBCAE5174064838FDD5F918FDA70152" box="[602,625,1791,1819]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="5" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="5AC53CC974064838FDF1F919FEA00176" collectionCode="ZMUM" country="Spain" location="Michelsen" pageId="1" pageNumber="5" specimenCount="1" specimenCount-female="1">
<location id="EF72604F74064838FDF1F919FF6F0177" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FDF1F919FF6F0177" country="Spain" name="Michelsen" pageId="1" pageNumber="5">Michelsen</location>
,
<specimenCount id="FCABFD1D74064838FF49F8C5FF380177" box="[198,238,1826,1855]" count="1" pageId="1" pageNumber="5" type="female">1♀</specimenCount>
(
<collectionCode id="8CBCAE5174064838FF70F8C5FEBB0176" box="[255,365,1826,1855]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="1" pageNumber="5">ZMUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="EA12369474064838FF18F8A2FC4807C0" blockId="1.[151,756,1086,2031]" lastBlockId="1.[792,1399,153,640]" pageId="1" pageNumber="5">
<materialsCitation id="5AC53CC974064838FF18F8A2FB2E0690" collectingDate="1971-05-24" collectingDateMax="1971-06-18" collectingDateMin="1971-05-24" collectorName="N. Krivosheina" country="Turkmenistan" latitude="37.2" location="Ipay-Kala" longLatPrecision="7102" longitude="59.8" pageId="1" pageNumber="5" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="92BA760474064838FF18F8A2FE4F012B" box="[151,409,1861,1890]" name="Turkmenistan" pageId="1" pageNumber="5">TURKMENISTAN</collectingCountry>
:
<specimenCount id="FCABFD1D74064838FE27F8A2FE160128" box="[424,448,1861,1889]" count="1" pageId="1" pageNumber="5" type="male">♂</specimenCount>
<typeStatus id="3516883674064838FE44F8A2FDE8012B" box="[459,574,1861,1890]" pageId="1" pageNumber="5" type="holotype">holotype</typeStatus>
<taxonomicName id="2DAD4D1774064838FDC5F8A2FE7801CC" authority="Sorokina, 2015" authorityName="Sorokina" authorityYear="2015" class="Insecta" family="Muscidae" genus="Phaonia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="5" phylum="Arthropoda" rank="species" species="juglans">
<emphasis id="D8D9EA8674064838FDC5F8A2FF1B01CC" italics="true" pageId="1" pageNumber="5">Phaonia juglans</emphasis>
<bibRefCitation id="8E3C4B6574064838FF50F88FFE7801CC" author="Sorokina, V. S." box="[223,430,1896,1925]" pageId="1" pageNumber="5" pagination="571 - 587" refId="ref2967" refString="Sorokina, V. S. (2015) New species of the genus Phaonia R. - D., 1830 (Diptera, Muscidae) from Central Asia. Zootaxa, vol. 4013, no. 4, pp. 571 - 587. DOI: 10.11646 / zootaxa. 4013.4.7 (In English)" type="journal article" year="2015">Sorokina, 2015</bibRefCitation>
</taxonomicName>
,
<location id="EF72604F74064838FE4AF88FFD9401CC" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FE4AF88FFD9401CC" box="[453,578,1896,1925]" country="Turkmenistan" latitude="37.2" longLatPrecision="7102" longitude="59.8" name="Ipay-Kala" pageId="1" pageNumber="5">Ipay-Kala</location>
[Kopet Dag Range,
<geoCoordinate id="8F99505374064838FE8AF86CFEB701E1" box="[261,353,1930,1960]" degrees="37.2" direction="north" orientation="latitude" pageId="1" pageNumber="5" precision="5555" value="37.2">
37.2
<emphasis id="D8D9EA8674064838FEB3F86DFE9F01E1" box="[316,329,1930,1960]" italics="true" pageId="1" pageNumber="5">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="8F99505374064838FEF9F86CFE1901E1" box="[374,463,1930,1960]" degrees="59.8" direction="east" orientation="longitude" pageId="1" pageNumber="5" precision="5555" value="59.8">
59.8
<emphasis id="D8D9EA8674064838FE22F86DFE6C01E1" box="[429,442,1930,1960]" italics="true" pageId="1" pageNumber="5">°</emphasis>
E
</geoCoordinate>
], reared from larva found in a hollow of a nut tree, collected on
<date id="9E13105474064838FD6BF849FEC901A6" pageId="1" pageNumber="5" value="1971-06-06">
<collectingDate id="8E57E9BC74064838FD6BF849FEC901A6" pageId="1" pageNumber="5" value="1971-06-06">6 June 1971</collectingDate>
</date>
, emergence
<date id="9E13105474064838FE48F835FDA101A6" box="[455,631,2002,2031]" pageId="1" pageNumber="5" value="1971-06-18">
<collectingDate id="8E57E9BC74064838FE48F835FDA101A6" box="[455,631,2002,2031]" pageId="1" pageNumber="5" value="1971-06-18">18 June 1971</collectingDate>
</date>
,
<collectorName id="4758534274064838FD08F835FC5706FF" box="[647,897,153,2030]" pageId="1" pageNumber="5">N. Krivosheina</collectorName>
; the same data as above,
<collectorName id="4758534274064838FB4BFF7DFCEB0690" pageId="1" pageNumber="5">N. Krivosheina</collectorName>
, collected
<date id="9E13105474064838FC5AFF5BFB2E0690" box="[981,1272,188,217]" pageId="1" pageNumber="5" value="1971-05-24" valueMax="1971-06-09" valueMin="1971-05-24">
<collectingDate id="8E57E9BC74064838FC5AFF5BFB2E0690" box="[981,1272,188,217]" pageId="1" pageNumber="5" value="1971-05-24" valueMax="1971-06-09" valueMin="1971-05-24">24 May–9 June 1971</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="5AC53CC974064838FA84FF5BFAD0072F" collectingDate="1983-04-20" collectorName="N. Krivosheina" country="Turkmenistan" latitude="38.0" location="Lobanov's" longLatPrecision="706" longitude="58.15" municipality="Chuli" pageId="1" pageNumber="5" specimenCount="10" specimenCount-female="3" specimenCount-male="7" typeStatus="holotype">
<specimenCount id="FCABFD1D74064838FA84FF5BFAE50690" box="[1291,1331,188,217]" count="4" pageId="1" pageNumber="5" type="male">4♂</specimenCount>
,
<specimenCount id="FCABFD1D74064838FAC8FF5BFAB90690" box="[1351,1391,188,217]" count="3" pageId="1" pageNumber="5" type="female">3♀</specimenCount>
,
<specimenCount id="FCABFD1D74064838FC95FF38FC6F06B5" box="[794,953,223,252]" count="3" pageId="1" pageNumber="5" type="male">three males</specimenCount>
with
<location id="EF72604F74064838FB82FF38FB4506B5" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FB82FF38FB4506B5" box="[1037,1171,223,252]" country="Turkmenistan" latitude="38.0" longLatPrecision="706" longitude="58.15" municipality="Chuli" name="Lobanov's" pageId="1" pageNumber="5">Lobanov’s</location>
labels identified as
<taxonomicName id="2DAD4D1774064838FCB0FEE4FC780756" authorityName="ZETTERSTEDT" authorityYear="1846" box="[831,942,259,287]" class="Insecta" family="Muscidae" genus="Phaonia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="5" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="D8D9EA8674064838FCB0FEE4FC780756" box="[831,942,259,287]" italics="true" pageId="1" pageNumber="5">P. cincta</emphasis>
</taxonomicName>
;
<locationDeviation id="FBA02FB674064838FC30FEE4FB080756" box="[959,1246,258,288]" location="Geok Tepe" pageId="1" pageNumber="5">
<quantity id="2D559B7174064838FC30FEE4FBC50756" box="[959,1043,258,288]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="5" unit="km" value="20.0">20 km</quantity>
SE Geok Tepe
</locationDeviation>
,
<collectingMunicipality id="0A76ACEE74064838FB63FEE5FAE00756" box="[1260,1334,258,287]" pageId="1" pageNumber="5">Chuli</collectingMunicipality>
vill.,
<geoCoordinate id="8F99505374064838FC93FEC1FC51070B" box="[796,903,292,323]" degrees="38.00" direction="north" orientation="latitude" pageId="1" pageNumber="5" precision="555" value="38.0">
38.00
<emphasis id="D8D9EA8674064838FCEEFEC3FCB8070B" box="[865,878,292,322]" italics="true" pageId="1" pageNumber="5">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="8F99505374064838FC1EFEC2FC2D070B" box="[913,1019,292,322]" degrees="58.15" direction="east" orientation="longitude" pageId="1" pageNumber="5" precision="555" value="58.15">
58.15
<emphasis id="D8D9EA8674064838FC56FEC3FC30070B" box="[985,998,292,322]" italics="true" pageId="1" pageNumber="5">°</emphasis>
E
</geoCoordinate>
, reared from larvae collected on
<date id="9E13105474064838FCFFFEAEFBFD072F" box="[880,1067,329,358]" pageId="1" pageNumber="5" value="1983-04-20">
<collectingDate id="8E57E9BC74064838FCFFFEAEFBFD072F" box="[880,1067,329,358]" pageId="1" pageNumber="5" value="1983-04-20">20 April 1983</collectingDate>
</date>
,
<collectorName id="4758534274064838FBB5FEADFAD0072F" box="[1082,1286,329,358]" pageId="1" pageNumber="5">N. Krivosheina</collectorName>
</materialsCitation>
,
<materialsCitation id="5AC53CC974064838FA9AFEAEFC4C07C0" collectionCode="ZMUM" country="Turkmenistan" pageId="1" pageNumber="5" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="holotype">
<specimenCount id="FCABFD1D74064838FA9AFEAEFAEB072F" box="[1301,1341,329,358]" count="2" pageId="1" pageNumber="5" type="male">2♂</specimenCount>
,
<specimenCount id="FCABFD1D74064838FAC1FEAEFAA0072F" box="[1358,1398,329,358]" count="2" pageId="1" pageNumber="5" type="female">2♀</specimenCount>
(
<collectionCode id="8CBCAE5174064838FCACFE8BFC4707C0" box="[803,913,364,393]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="1" pageNumber="5">ZMUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="EA12369474064838FC95FE77FAF304C9" blockId="1.[792,1399,153,640]" pageId="1" pageNumber="5">
<collectingCountry id="92BA760474064838FC95FE77FC7D07E4" box="[794,939,400,429]" name="Ukraine" pageId="1" pageNumber="5">UKRAINE</collectingCountry>
:
<emphasis id="D8D9EA8674064838FC4BFE68FBFE07E5" box="[964,1064,399,428]" italics="true" pageId="1" pageNumber="5">
<collectingRegion id="2869F87674064838FC4BFE68FBFE07E5" box="[964,1064,399,428]" country="Ukraine" name="Poltava" pageId="1" pageNumber="5">Poltava</collectingRegion>
</emphasis>
reg.,
<location id="EF72604F74064838FB02FE68FB3D07E5" LSID="urn:lsid:plazi:treatment:6204878274064838FF6BFC22FAF304C9:EF72604F74064838FB02FE68FB3D07E5" box="[1165,1259,399,428]" country="Ukraine" latitude="49.84" longLatPrecision="660" longitude="33.9" name="Yareski" pageId="1" pageNumber="5" stateProvince="Poltava">Yareski</location>
(
<geoCoordinate id="8F99505374064838FA85FE68FAA107E5" box="[1290,1399,398,428]" degrees="49.84" direction="north" orientation="latitude" pageId="1" pageNumber="5" precision="555" value="49.84">
49.84
<emphasis id="D8D9EA8674064838FADEFE69FA8807E5" box="[1361,1374,398,428]" italics="true" pageId="1" pageNumber="5">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="8F99505374064838FC95FE54FC570799" box="[794,897,433,464]" degrees="33.90" direction="east" orientation="longitude" pageId="1" pageNumber="5" precision="555" value="33.9">
33.90
<emphasis id="D8D9EA8674064838FCEFFE56FCBB0786" box="[864,877,433,463]" italics="true" pageId="1" pageNumber="5">°</emphasis>
E
</geoCoordinate>
),
<date id="9E13105474064838FC29FE54FB620799" box="[934,1204,434,464]" pageId="1" pageNumber="5" value="1929-09-01">
<collectingDate id="8E57E9BC74064838FC29FE54FB620799" box="[934,1204,434,464]" pageId="1" pageNumber="5" value="1929-09-01">1 September 1929</collectingDate>
</date>
,
<collectorName id="4758534274064838FB5FFE55FAA70786" box="[1232,1393,434,463]" pageId="1" pageNumber="5">Gildebrandt</collectorName>
,
<specimenCount id="FCABFD1D74064838FC95FE31FC9707BB" box="[794,833,470,499]" count="1" pageId="1" pageNumber="5" type="male">1♂</specimenCount>
;
<emphasis id="D8D9EA8674064838FCDDFE31FC1607BA" box="[850,960,470,499]" italics="true" pageId="1" pageNumber="5">
<collectingRegion id="2869F87674064838FCDDFE31FC1607BA" box="[850,960,470,499]" country="Ukraine" name="Kharkiv" pageId="1" pageNumber="5">Kharkov</collectingRegion>
</emphasis>
reg.: vicinity of
<collectingRegion id="2869F87674064838FB17FE31FAD107BA" box="[1176,1287,470,499]" country="Ukraine" name="Kharkiv" pageId="1" pageNumber="5">Kharkov</collectingRegion>
(
<geoCoordinate id="8F99505374064838FA95FE32FAA107BB" box="[1306,1399,468,498]" degrees="50.0" direction="north" orientation="latitude" pageId="1" pageNumber="5" precision="5555" value="50.0">
50.0
<emphasis id="D8D9EA8674064838FADEFE33FA8807BB" box="[1361,1374,468,498]" italics="true" pageId="1" pageNumber="5">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="8F99505374064838FC95FE1EFCA7045F" box="[794,881,503,534]" degrees="36.2" direction="east" orientation="longitude" pageId="1" pageNumber="5" precision="5555" value="36.2">
36.2
<emphasis id="D8D9EA8674064838FCDFFE10FC8B045C" box="[848,861,503,533]" italics="true" pageId="1" pageNumber="5">°</emphasis>
E
</geoCoordinate>
),
<collectorName id="4758534274064838FC06FE1DFB8F045F" box="[905,1113,505,534]" pageId="1" pageNumber="5">W. Yaroshevsky</collectorName>
,
<date id="9E13105474064838FBE7FE1EFAC9045F" box="[1128,1311,505,534]" pageId="1" pageNumber="5" value="1882-04-25">
<collectingDate id="8E57E9BC74064838FBE7FE1EFAC9045F" box="[1128,1311,505,534]" pageId="1" pageNumber="5" value="1882-04-25">25 April 1882</collectingDate>
</date>
,
<specimenCount id="FCABFD1D74064838FAA1FE1EFA80045F" box="[1326,1366,505,534]" count="1" pageId="1" pageNumber="5" type="male">1♂</specimenCount>
;
<date id="9E13105474064838FAE9FE1EFC4B0470" pageId="1" pageNumber="5" value="1882-05-02">
<collectingDate id="8E57E9BC74064838FAE9FE1EFC4B0470" pageId="1" pageNumber="5" value="1882-05-02">2 May 1882</collectingDate>
</date>
,
<specimenCount id="FCABFD1D74064838FC27FDFBFC1B0470" box="[936,973,540,569]" count="1" pageId="1" pageNumber="5" type="male">1♂</specimenCount>
; uncertain localities, presumably the
<collectorName id="4758534274064838FCC7FDD8FC630415" box="[840,949,575,604]" pageId="1" pageNumber="5">Ukraine</collectorName>
:
<collectorName id="4758534274064838FC32FDA7FBB00415" box="[957,1126,575,604]" pageId="1" pageNumber="5">I. Porchinsky</collectorName>
,
<specimenCount id="FCABFD1D74064838FBFFFDD8FB4C0415" box="[1136,1178,575,604]" count="1" pageId="1" pageNumber="5" type="male">1♂</specimenCount>
;
<collectorName id="4758534274064838FB29FDA7FAA50415" box="[1190,1395,575,604]" pageId="1" pageNumber="5">W. Yaroshevsky</collectorName>
, on oak trunk,
<date id="9E13105474064838FC40FD84FBB90436" box="[975,1135,610,640]" pageId="1" pageNumber="5" value="1886-07-30">
<collectingDate id="8E57E9BC74064838FC40FD84FBB90436" box="[975,1135,610,640]" pageId="1" pageNumber="5" value="1886-07-30">30 July 1886</collectingDate>
</date>
,
<specimenCount id="FCABFD1D74064838FBF4FD84FB720436" box="[1147,1188,611,640]" count="1" pageId="1" pageNumber="5" type="male">1♂</specimenCount>
(all
<collectionCode id="8CBCAE5174064838FB54FD84FAC104C9" box="[1243,1303,611,640]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="5" type="Museum">ZIN</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>

100
data/62/05/33/6205333B3DFEFCF25352CA66311EBC64.xml Normal file
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@ -0,0 +1,100 @@
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<mods:title id="F55355672ED70B6A6121ACAB7B884B4F">Two new glyptosternine catfishes (Teleostei: Sisoridae) from Vietnam and China.</mods:title>
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<treatment id="6205333B3DFEFCF25352CA66311EBC64" ID-DOI="http://doi.org/10.5281/zenodo.6270273" ID-GBIF-Taxon="100124423" ID-Zenodo-Dep="6270273" LSID="urn:lsid:plazi:treatment:6205333B3DFEFCF25352CA66311EBC64" httpUri="http://treatment.plazi.org/id/6205333B3DFEFCF25352CA66311EBC64" lastPageNumber="2" pageNumber="2">
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<taxonomicName id="0C1A7A737CE535A8F8521CC54DE55336" ID-CoL="4DLDC" LSID-ZBK="urn:lsid:zoobank.org:act:AF241874-BEBF-4718-B795-BF5807DD0826" family="Sisoridae" genus="Pareuchiloglanis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pareuchiloglanis nebulifer Ng &amp; Kottelat 2000:11" order="Siluriformes" pageNumber="2" phylum="Chordata" rank="species" species="nebulifer">P. nebulifer</taxonomicName>
:
</paragraph>
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<subSubSection id="E4FDFC06CCC77B997D9FF9CCCBDF96D4" type="materials_examined">
<paragraph id="AE4C68E2567822A5E9C097EC8C419D69" pageNumber="2">
<materialsCitation id="A660A8037FBF46B6C1E60CC1DF7CB823" ID-GBIF-Occurrence="923902069" collectionCode="ZRC" country="Laos" latitude="20.326668" location="small creek, tributary of Houai Siam, upstream of Ban Kangpabong" longitude="104.41694" specimenCode="ZRC 45706" stateProvince="Houaphan province" typeStatus="holotype">
<specimenCode id="0D7F8F6FA803D931DE8F733D1AE3794E">
<collectionCode id="227464CABD3C6B4F1DEF33582E9855F5">ZRC</collectionCode>
45706
</specimenCode>
(
<typeStatus id="B30CEE703F148D43C2BDE6D4CA1A30C7">holotype</typeStatus>
), 94.8 mm SL;
</materialsCitation>
<materialsCitation id="01713E1181277B1D606423F70074A740" ID-GBIF-Occurrence="923902068" collectionCode="CMK" country="Laos" latitude="20.326668" location="small creek, tributary of Houai Siam, upstream of Ban Kangpabong" longitude="104.41694" specimenCode="CMK 15447" stateProvince="Houaphan province" typeStatus="paratype">
<specimenCode id="FF719DD63D64107C759BB74F9D183A14">
<collectionCode id="20A34903C7973BF11656157E9A54A9E0">CMK</collectionCode>
15447
</specimenCode>
(1
<typeStatus id="4470D92373146B3E462F0DFF6A732120">paratype</typeStatus>
), 62.3 mm SL;
<collectingCountry id="790890B4FB8AA943E41A07FA047F0464">Laos</collectingCountry>
:
<collectingRegion id="9831EC5A01F466BE48965219E5A681D2">Houaphan province</collectingRegion>
,
<location id="FEFB2392EEB126C6F72D4E59784438E2" LSID="urn:lsid:plazi:treatment:6205333B3DFEFCF25352CA66311EBC64:FEFB2392EEB126C6F72D4E59784438E2" country="Laos" latitude="20.326668" longitude="104.41694" name="small creek, tributary of Houai Siam, upstream of Ban Kangpabong" stateProvince="Houaphan province">small creek, tributary of Houai Siam, upstream of Ban Kangpabong</location>
,
<geoCoordinate id="E16481F0C9C18811046682ED91929365" direction="north" orientation="latitude" value="20.326668">20°19'36&quot;N</geoCoordinate>
<geoCoordinate id="FACDC1D7DA2863F91116D917F6F8A8CA" direction="east" orientation="longitude" value="104.41694">104°25'1&quot;E</geoCoordinate>
.
</materialsCitation>
<materialsCitation id="9C65D1C01D6BFC45FC1293121C03C4B2" ID-GBIF-Occurrence="923902060" collectionCode="CMK" country="Laos" latitude="20.133055" location="unnamed forest stream about 4 km SE from Ban Houatangoua on road to Xam Tai" longitude="104.56084" specimenCode="CMK 15342" stateProvince="Houaphan province" typeStatus="paratype">
<specimenCode id="5A6CACFCE84070BBAB4AC1F589FEE9B7">
<collectionCode id="D95D30FD4154F56E8D48449EA231E988">CMK</collectionCode>
15342
</specimenCode>
(1
<typeStatus id="7C021828C73E82DAB40B98A6ABC3EB3F">paratype</typeStatus>
) 86.1 mm SL;
</materialsCitation>
<materialsCitation id="F36283233292F8B9AA5506515E3F78FC" ID-GBIF-Occurrence="923902062" collectionCode="LARRI" country="Laos" latitude="20.133055" location="unnamed forest stream about 4 km SE from Ban Houatangoua on road to Xam Tai" longitude="104.56084" specimenCode="LARRI uncat." stateProvince="Houaphan province" typeStatus="paratype">
<specimenCode id="F19BB9A9C51BECA7644BF08659C16DC9">
<collectionCode id="809B0BE837339D7FEC8851CD3DC15CBB">LARRI</collectionCode>
uncat.
</specimenCode>
(1
<typeStatus id="BDC3FD5CF449C488AE8DAF4A4F864A5D">paratype</typeStatus>
), 75.2 mm SL;
<collectingCountry id="FB642230B77E3D577F940AC5E13DD15C">Laos</collectingCountry>
:
<collectingRegion id="892A5827D974BD88C6CB9CAFEAB00B76">Houaphan province</collectingRegion>
,
<location id="A46943FFE034152765F05F3084AAD7B1" LSID="urn:lsid:plazi:treatment:6205333B3DFEFCF25352CA66311EBC64:A46943FFE034152765F05F3084AAD7B1" country="Laos" latitude="20.133055" longitude="104.56084" name="unnamed forest stream about 4 km SE from Ban Houatangoua on road to Xam Tai" stateProvince="Houaphan province">unnamed forest stream about 4 km SE from Ban Houatangoua on road to Xam Tai</location>
,
<geoCoordinate id="51473C682243B0ED4353B33337F256D2" direction="north" orientation="latitude" value="20.133055">20°7'59&quot;N</geoCoordinate>
<geoCoordinate id="1DD20B6A8DD35737963AA87C50F7B71B" direction="east" orientation="longitude" value="104.56084">104°33'39&quot; E</geoCoordinate>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

59
data/62/05/4E/62054EFCCCE8FA359CCE1460632A25D3.xml Normal file
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@ -0,0 +1,59 @@
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<mods:namePart id="0B72835E7F3C4A9548E8F072FF34EF1A">Santschi, F.</mods:namePart>
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<mods:title id="3045CAD380347DB05E7E479FE453F515">Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale (1911 - 1912). Résultats scientifiques. Hyménoptères</mods:title>
</mods:titleInfo>
<mods:part id="58D71B8AB1AA8F17C86D1026453E0394">
<mods:date id="87D93EEC015B8D985F4E7F3153D7347A">1914</mods:date>
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<mods:number id="AF827107F2137A925500BCD34115DBB2">2</mods:number>
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<mods:identifier id="7B842597F38504073BBC7471E4164BCB" type="HNS-Pub">8111</mods:identifier>
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<treatment id="62054EFCCCE8FA359CCE1460632A25D3" ID-DOI="http://doi.org/10.5281/zenodo.5063690" ID-GBIF-Taxon="100126759" ID-Zenodo-Dep="5063690" LSID="urn:lsid:plazi:treatment:62054EFCCCE8FA359CCE1460632A25D3" httpUri="http://treatment.plazi.org/id/62054EFCCCE8FA359CCE1460632A25D3" lastPageNumber="109" pageNumber="108">
<subSubSection id="F2808A312B768325CFD9E6C71F6CD0FF" type="nomenclature">
<paragraph id="85B4AC3FCD7C8A8A8EACCBA6C6F4F912" pageNumber="108">
<taxonomicName id="3C75E6B26AF4B803598DD1A2913E9290" ID-CoL="RTCF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:27805" class="Insecta" family="Formicidae" genus="Cataulacus" kingdom="Animalia" lsidName="Cataulacus jeanneli Santschi" lsidName-HNS="Cataulacus jeanneli Santschi" order="Hymenoptera" pageNumber="108" phylum="Arthropoda" rank="species" species="jeanneli">Cataulacus Jeanneli</taxonomicName>
,
<taxonomicNameLabel id="4DFDEE121241E75D7931BEC54B4C0430" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="063E320DC07977E2AC710494059FA111" type="description">
<paragraph id="3F8B9B384081BA3248F8C35641CFF8BA" pageNumber="108">Fig. 16.</paragraph>
<paragraph id="2D7F09F2306BF8EBFF46D9B171448D22" lastPageNumber="109" pageNumber="108">
[[worker]]. - Long. 2,5 mill. Noir. Scape, genou, tibias et tarses jaune rougeatre. Pilosite tres courte, epaisse, tronquee et tres clairsemee sur le corps, plus longue sous le gastre. Quelques poils tres fins, aussi courts que les soies et aussi clairsemes, se voient en outre sur les pattes et les antennes. Mat. Tete lachement ridee- reticulee; les rides principales sont irregulierement longitudinales avec des anastomoses obliques ou transverses presque aussi fortes. Le fond, entre les mailles, est finement rugueux, presque lisse et assez luisant sur le dessus et le devant de la tete, tandis qu'en arriere il devient plutot ponctue en de a coudre. Cette derniere sculpture se retrouve sur tout le reste de l'insecte. Dos du thorax et pedicule fortement rides-canneles en long avec quelques anastomoses sur le devant du pronotum, et une zone sur le milieu du mesonotum, ou les rides s'attenuent un peu et laissent mieux voir la sculpture du fond. Gastre entierement ponctue-reti- cule en de a coudre, a peine quelques petites rides vers la base. Tete rectangulaire d'un quart a un cinquieme plus longue que large (aussi large que longue chez
<taxonomicName id="388D094728CBF392B33CBD05668605AB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:27827" class="Insecta" family="Formicidae" genus="Cataulacus" kingdom="Animalia" lsidName="Cataulacus pygmaeus André" lsidName-HNS="Cataulacus pygmaeus Andre" order="Hymenoptera" pageNumber="108" phylum="Arthropoda" rank="species" species="pygmaeus">C. pygmaeus Andre</taxonomicName>
). Les cotes legerement convexes, a peine denticules, le bord occipital presque droit avec les angles marques de deux dents mediocres. Les yeux ovales sont tres grands, plus longs que la distance qui les separe de l'angle anterieur de la tete. L'arete antennaire atteint le milieu de leur extremite anterieure et la penetre lege- rement de facon qu'une portion de l' oe il regarde en dessous dans la gouttiere antennaire. Bord de l'arete antennaire convexe, sans echancrure, avec une dent a ses deux extremites, la posterieure, preoculaire, moins indiquee. Mandibules finement reticulees. Thorax presque aussi large que la tete, a bords dentes, sans sutures, presque deux fois plus long que large. Le profil est a peine convexe sauf le devant du pronotum. Celui-ci aussi large en arriere qu'en avant. Epines epinotales aussi longues que les deux tiers de l'intervalle de leur extremite. Premier n oe ud cylindrique, un peu plus large en avant et aussi long que large avec un fort appendice en dessous. Deuxieme n oe ud plus large que long et que le prece- dent, retreci en arriere, presque deux fois plus haut que long. Gastre allonge, a cotes paralleles, pas beaucoup plus large que le thorax, etroitement et profondement echancre devant.
</paragraph>
<paragraph id="F00E6E83C9BCC722F3987946B1257823" pageNumber="109">
Celte petite espece, du groupe
<taxonomicName id="B0D8BE968CEA7579921D4DDED75F00BE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:27827" class="Insecta" family="Formicidae" genus="Cataulacus" kingdom="Animalia" lsidName="Cataulacus pygmaeus André" lsidName-HNS="Cataulacus pygmaeus Andre" order="Hymenoptera" pageNumber="109" phylum="Arthropoda" rank="species" species="pygmaeus">pygmaeus Andre</taxonomicName>
, est surtout caracterisee par ses tres grands yeux et sa tete allongee. Voisine du C. Weissi Santschi, dont la pilosite est bien plus longue et abondante, et du C. Lujae For. que je ne connais pas en nature.
</paragraph>
</subSubSection>
<subSubSection id="F9226A66A35CA77F55F6A76B92C74354" type="materials_examined">
<paragraph id="E6C95BA10066E96612C1C2F7C9E633B8" pageNumber="109">Afrique orientale anglaise: Gazi, dans la region cotiere, a 20 km. au sud de Mombasa (st. n° 6, nov. 1911), 1 [[worker]].</paragraph>
</subSubSection>
</treatment>
</document>

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<vernacularName id="64AF46A86368FFFB08A8F7DB71329F60" box="[237,576,2054,2100]" pageId="1" pageNumber="172">Banded Linsang</vernacularName>
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<heading id="B15B81EA6368FFFB0A34F7DB70A39F60" box="[625,977,2054,2100]" pageId="1" pageNumber="172">
<taxonomicName id="2DAC4D056368FFFB0A34F7DB70A39F60" baseAuthorityName="Hardwicke" baseAuthorityYear="1821" box="[625,977,2054,2100]" class="Bryopsida" family="Prionodontaceae" genus="Prionodon" kingdom="Plantae" order="Isobryales" pageId="1" pageNumber="172" phylum="Bryophyta" rank="species" species="linsang">
<emphasis id="D8D8EA946368FFFB0A34F7DB70A39F60" box="[625,977,2054,2100]" pageId="1" pageNumber="172">Prionodon linsang</emphasis>
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<heading id="B15B81EA6368FFFB08F9F79B70D69F0F" box="[188,932,2118,2139]" pageId="1" pageNumber="172">
<emphasis id="D8D8EA946368FFFB08F9F79B727B9F0F" box="[188,265,2118,2139]" pageId="1" pageNumber="172">French:</emphasis>
<vernacularName id="64AF46A86368FFFB0956F79B72FD9F0F" box="[275,399,2118,2139]" pageId="1" pageNumber="172">Linsang rayé</vernacularName>
/
<emphasis id="D8D8EA946368FFFB09E1F79B728D9F0F" box="[420,511,2118,2139]" pageId="1" pageNumber="172">German:</emphasis>
<vernacularName id="64AF46A86368FFFB0A4FF79B71E79F0F" box="[522,661,2118,2139]" pageId="1" pageNumber="172">Banderlinsang</vernacularName>
/
<emphasis id="D8D8EA946368FFFB0AEFF79B70779F0F" box="[682,773,2118,2139]" pageId="1" pageNumber="172">Spanish:</emphasis>
<vernacularName id="64AF46A86368FFFB0B55F79B70D69F0F" box="[784,932,2118,2139]" pageId="1" pageNumber="172">Linsang rayado</vernacularName>
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<emphasis id="D8D8EA946368FFFB08FEF75672249FFC" box="[187,342,2187,2216]" pageId="1" pageNumber="172">Taxonomy.</emphasis>
<taxonomicName id="2DAC4D056368FFFB092CF75670549FFC" authority="Hardwicke, 1821" authorityName="Hardwicke" authorityYear="1821" box="[361,806,2187,2216]" class="Mammalia" family="Viverridae" genus="Viverra" kingdom="Animalia" order="Carnivora" pageId="1" pageNumber="172" phylum="Chordata" rank="species" species="linsang">Viverra linsang Hardwicke, 1821</taxonomicName>
,
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<paragraph id="EA1336866368FFFB0B7CF75671E09F84" blockId="1.[183,1391,2187,3484]" pageId="1" pageNumber="172">
<materialsCitation id="5AC43CDB6368FFFB0B7CF75671E09F84" ID-GBIF-Occurrence="3710100301" pageId="1" pageNumber="172">
Malacca,
<collectingCountry id="92BB76166368FFFB0BFBF75677479FFC" box="[958,1077,2187,2216]" name="Malaysia" pageId="1" pageNumber="172">Malaysia</collectingCountry>
. [restricted by Robinson &amp; Kloss (1920) to “
<collectingRegion id="2868F8646368FFFB0A45F76E71089F84" box="[512,634,2227,2256]" pageId="1" pageNumber="172">Malacca</collectingRegion>
”].
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Two subspecies recognized,
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includes maculosus and gracilis includes hardwichaz, fredericae, and interliniurus.
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<caption id="BED3660E6368FFFB08F9F6FB77009EE8" ID-DOI="http://doi.org/10.5281/zenodo.5714316" ID-Zenodo-Dep="5714316" httpUri="https://zenodo.org/record/5714316/files/figure.png" inLine="true" pageId="1" pageNumber="172" targetBox="[1461,2043,1924,2332]" targetPageId="1">
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<emphasis id="D8D8EA946368FFFB08F9F6FB71399E13" box="[188,587,2342,2375]" pageId="1" pageNumber="172">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph id="EA1336866368FFFB08F8F68872379EC1" blockId="1.[183,1391,2187,3484]" pageId="1" pageNumber="172">
<taxonomicName id="2DAC4D056368FFFB08F8F68871119E3A" authority="Hardwicke, 1821" authorityName="Hardwicke" authorityYear="1821" box="[189,611,2389,2414]" class="Bryopsida" family="Prionodontaceae" genus="Prionodon" kingdom="Plantae" order="Isobryales" pageId="1" pageNumber="172" phylum="Bryophyta" rank="subSpecies" species="linsang" subSpecies="linsang">P. l. linsang Hardwicke, 1821</taxonomicName>
— S
<collectingCountry id="92BB76166368FFFB0AF6F68870489E3A" box="[691,826,2389,2414]" name="Myanmar" pageId="1" pageNumber="172">Myanmar</collectingCountry>
, Peninsular
<collectingCountry id="92BB76166368FFFB0BBCF68877019E3A" box="[1017,1139,2389,2414]" name="Malaysia" pageId="1" pageNumber="172">Malaysia</collectingCountry>
, Sumatra, and S
<collectingCountry id="92BB76166368FFFB08F9F6A572309EC1" box="[188,322,2424,2453]" name="Thailand" pageId="1" pageNumber="172">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="EA1336866368FFFB08F8F64277009EE8" blockId="1.[183,1391,2187,3484]" box="[189,1138,2463,2492]" pageId="1" pageNumber="172">
<taxonomicName id="2DAC4D056368FFFB08F8F642715D9EE8" authority="Horsfield, 1822" authorityName="Horsfield" authorityYear="1822" box="[189,559,2463,2492]" class="Bryopsida" family="Prionodontaceae" genus="Prionodon" kingdom="Plantae" order="Isobryales" pageId="1" pageNumber="172" phylum="Bryophyta" rank="subSpecies" species="linsang" subSpecies="gracilis">P. l. gracilis Horsfield, 1822</taxonomicName>
— Bangka I, Belitung I, Borneo, and
<collectingRegion id="2868F8646368FFFB0C75F642771C9EE8" box="[1072,1134,2463,2492]" country="Indonesia" name="Jawa Timur" pageId="1" pageNumber="172">Java</collectingRegion>
.
</paragraph>
</caption>
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<paragraph id="EA1336866368FFFB08F9F61F71629AC7" blockId="1.[183,1391,2187,3484]" pageId="1" pageNumber="172">
<emphasis id="D8D8EA946368FFFB08F9F61F72C99EB7" box="[188,443,2498,2531]" pageId="1" pageNumber="172">Descriptive notes.</emphasis>
Head-body 37.9-45 cm, tail 33-37.
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; weight 590-800 g. Ratio head-body/ tail slightly superior to one. Two pairs of mammae. Adult males may be slightly larger than females, but size overlap between the two genders is considerable. Has a genet-like aspect, with head and neck more elongated; pointed muzzle; nose pad brownish pink. Extremely slender; captive individual reported to escape through bars
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apart. Ears are medium-sized, broad at the base and rounded.Iris is dark brown to black, and pupil is vertical. Pelage is short and very soft. Coat color is relatively uniform across the body, and varies from pale yellowish to white. Individual variation exists in spot markings. Back exhibits two main pairs of lines of very large dark spots that fuse into five more or less clearly delineated transversal stripes. The characteristic pair of nucchal stripes in continuation with the second row of spots is wider than in Spotted Linsang. A line of thin spots starting behind the shoulders forms a broken medio-dorsal stripe thatis vertically crossing the transversal stripes. Smaller spots are irregularly distributed on the flanks, shoulders and thighs. The rest of the hindlimbs and the forepaws are not spotted. Five claws fully retractile; presence of sheaths. Hindfeet digitigrade, forefeet plantigrade; hallucal lobe small and remote; absence of metatarsal pads. Tail is almost as long as head and body; thickly furred, with short hair; exhibits seven to nine broad dark rings. Confused annealing pattern at the whitish tip, with the last bright ring often interspersed with a thin dark ring. Perineal glands are absent. Rostrum is moderately elongated; sagittal crest forms a large, flat stripe that fuses with the frontal bone. Caudal entotympanic bone more ventrally inflated—compared to the ectotympanic bone—than in Spotted Linsang. Dental formula: 13/3, C1/1,P 4/4, M 1/2 = 38. M2 may sometimes exist under vestigial condition. Chromosome number: 2n = 34. In gracilis, average body size and skull length are smaller.
</paragraph>
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<emphasis id="D8D8EA946368FFFB0050F8AA7BF690C0" box="[2069,2180,1911,1940]" pageId="1" pageNumber="172">Habitat.</emphasis>
Primary-secondary evergreen and mixed deciduous forests, up to
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(e.g. Jang Plateau,
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). May be found in ecotonal habitats and disturbed forests; few records in human-inhabited places. Its distribution could be shaped by moisture gradients and their topographical variations.
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D6368FFFB0050F770759B9DF3" pageId="1" pageNumber="172" type="food_feeding">
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<emphasis id="D8D8EA946368FFFB0050F7707A6D9F9A" box="[2069,2335,2221,2254]" pageId="1" pageNumber="172">Food and Feeding.</emphasis>
Primarily carnivorous. Stomach contents in continental
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and Borneo included rodents (long-tailed rats, spiny-furred rats, ground squirrels), lizards, frogs, birds and cockroaches. Captive individuals were fed with minced meat, mice, chicks, fresh fish, eggs, cottage cheese, and occasionally birds and ox liver;fruits were always refused. May hunt both in canopy and on the ground; direct observations always made on the ground. Great dexterity when chasing small birds. Rodents are killed with a bite to the neck; may combine grasping and killing into one movement, and proceed to a series of crunching bites from the seized part of the body to the head while the prey is maintained between the jaws. Forepaws are rarely used for manipulating preys during feeding; however, an individual was seen pulling off flesh pieces while maintaining the prey with its forepaws (head was eaten first).
</paragraph>
</subSubSection>
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<emphasis id="D8D8EA946368FFFB0DF6F57175EC9D99" box="[1459,1694,2732,2765]" pageId="1" pageNumber="172">Activity patterns.</emphasis>
Mostly nocturnal, but may be active during daytime. Presumably arboreal, but probably uses the ground to move and forage. Individuals in continental
<collectingCountry id="92BB76166368FFFB0DF3F526755B9C48" box="[1462,1577,2811,2844]" name="Malaysia" pageId="1" pageNumber="172">Malaysia</collectingCountry>
were seen moving among tree branches 3-8 m high but were always trapped on the ground. Nests in hollows of dead trees, under roots of large trees, or in tree holes above ground level. A nest entrance was around
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in diameter and
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in depth; the inside was covered with leaves and contained bits of dry sticks. Good climber and jumper; semi-digitigrade, with plantigrade forefeet and digitigrade hindfeet; uses a “head-first” vertical descent of trees. Has a cat-like gait, and often stands on its hindlegs. In captivity, observed sleeping with tail curled around forepaws and head; frequent sharpening of the claws; self grooming and face washing very similar to genets.
</paragraph>
</subSubSection>
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<emphasis id="D8D8EA946368FFFB0DF0F3E87B1F9B02" box="[1461,2157,3125,3158]" pageId="1" pageNumber="172">Movements, Home range and Social organization.</emphasis>
Almost unknown. Probably solitary. Newly-born captive kittens emit shrill and vibrating cries when handled; adults may snarl and make sharp, whistling noises when other animals are confronted. A captive male frequently urinated while walking, marking about
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around. Neck and shoulders, and subsequently flanks, are rubbed against objects for marking.
</paragraph>
</subSubSection>
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<emphasis id="D8D8EA946368FFFB0DF3F32775499A4F" box="[1462,1595,3322,3355]" pageId="1" pageNumber="172">Breeding.</emphasis>
Litters of two cubs, apparently once a year, between February and August. Dens reported in hollow trees; may raise young on the ground. Young with same coloration as adults except at nose (pink) and undersides (white). At birth, captive cubs weighed
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and measured
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from head to tail. Weight was doubled at day 18; eyes were wide open between 18 and 21 days. Young were
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and
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at eight weeks; equalled adult size at four months; at this age, they had the same color as their parents but their undersides were still whiter. The mother never carried her young in her mouth. In the wild, females stay with their mother until maturity, but males leave after weaning. A captive specimen lived up to ten years and eight months.
</paragraph>
</subSubSection>
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<emphasis id="D8D8EA94636BFFF808DCFE3C72839556" box="[153,497,481,514]" pageId="2" pageNumber="173">Status and Conservation.</emphasis>
CITES Appendix II. Classified as Least Concern on The [UCN Red List. Widespread distribution over the Sundaic region at all elevations but records are scarce, especially at northern edge (
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,
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) and in Java. Major threats are deforestation and,to a lesser extent, hunting/trade for fur. Most of the skins kept in museums were collected from natives. Protected by national laws in
<collectingCountry id="92BB7616636BFFF80C28FD5B778395CB" box="[1133,1265,646,671]" name="Myanmar" pageId="2" pageNumber="173">Myanmar</collectingCountry>
,
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,
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(only partially protected in
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) and
<collectingCountry id="92BB7616636BFFF80BEAFD7877499592" box="[943,1083,677,710]" name="Indonesia" pageId="2" pageNumber="173">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF808DDFD0877F19468" pageId="2" pageNumber="173" type="bibRefCitation_list">
<paragraph id="EA133686636BFFF808DDFD0877F19468" blockId="2.[151,1358,288,830]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF808DDFD08724095BA" box="[152,306,725,750]" pageId="2" pageNumber="173">Bibliography.</emphasis>
Azlan (2003), Banks (1949), Boitani et al. (2006), Davis (1958), Gangloff (1975), Hardwicke (1821), Jones (1982), Lim (1973, 1976), Louwman (1970), Nowak (1999), Pocock (1915d, 1933c, 1935), Sody (1936, 1949), Schreiber et al. (1989), Taylor (1988), Van Rompaey (1993), Wells et al. (2005).
</paragraph>
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</treatment>
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76
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Family
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• Moist and evergreen forests, including montane forests up to
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.
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238
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<mods:typeOfResource id="55F6BC7AE49753C20765AF9F2C9FAF7D">text</mods:typeOfResource>
<mods:relatedItem id="F8048ACCC79B8B37A7EB56AA62289F2F" type="host">
<mods:originInfo id="D3F46ABBF72EFE08F6848B2DCD3F5A31">
<mods:dateIssued id="1ABDFDDD21443486A11D19D702CE0B60">2009</mods:dateIssued>
<mods:dateOther id="E69A922E1CE50E347BF53A5581A03D03" type="pubDate">2009-01-31</mods:dateOther>
<mods:publisher id="4886E8D1A0161F133CA8699FA6F5C118">Lynx Edicions</mods:publisher>
<mods:place id="29D8B6581326834EA4804B11A4D5736C">
<mods:placeTerm id="12FEE66D8F21B7D4F9431DCF5301A075">Barcelona</mods:placeTerm>
</mods:place>
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<mods:titleInfo id="46994DF73D75A96CADE2C9251FD07340">
<mods:title id="867C00213F34F73D26975FEA1A4B0BFC">Handbook of the Mammals of the World – Volume 1 Carnivores</mods:title>
</mods:titleInfo>
<mods:part id="464CEBBFFA6558EE966640170B55C7EC">
<mods:extent id="05CAC0132C5433CD0E24A10ED7B9D2C9" unit="page">
<mods:start id="0B0658EB67D9ADA1F7547848F16B50E3">170</mods:start>
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<mods:classification id="5D4EBEADEF821AE0AB51DC13975B383F">book chapter</mods:classification>
<mods:identifier id="2E5DF266EA225A2881495E6DB6D797CB" type="CLB-Dataset">3404</mods:identifier>
<mods:identifier id="408C3B6734A201A5019BB20B792A169B" type="DOI">10.5281/zenodo.5714314</mods:identifier>
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<mods:identifier id="7F1880BC3C1B2D8B7F3299485F9AF91A" type="ISBN">978-84-96553-49-1</mods:identifier>
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<subSubSection id="A2B6650D636BFFF808DCFC5D73CA94E6" box="[153,184,896,946]" pageId="2" pageNumber="173" type="multiple">
<paragraph id="EA133686636BFFF808DCFC5D73CA94E6" blockId="2.[150,986,896,987]" box="[153,184,896,946]" pageId="2" pageNumber="173">
<heading id="B15B81EA636BFFF808DCFC5D73CA94E6" box="[153,184,896,946]" pageId="2" pageNumber="173">
<figureCitation id="72972A03636BFFF808DCFC5D73CA94E6" box="[153,184,896,946]" captionStart="Plate 11" captionStartId="1.[795,860,1704,1729]" captionTargetBox="[13,2796,16,1620]" captionTargetPageId="1" captionText="Plate 11 Species Accounts" figureDoi="http://doi.org/10.5281/zenodo.6554268" httpUri="https://zenodo.org/record/6554268/files/figure.png" pageId="2" pageNumber="173">2.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF8088CFC5D716E94E6" box="[201,540,896,946]" pageId="2" pageNumber="173" type="vernacular_names">
<paragraph id="EA133686636BFFF8088CFC5D716E94E6" blockId="2.[150,986,896,987]" box="[201,540,896,946]" pageId="2" pageNumber="173">
<heading id="B15B81EA636BFFF8088CFC5D716E94E6" box="[201,540,896,946]" pageId="2" pageNumber="173">
<vernacularName id="64AF46A8636BFFF8088CFC5D716E94E6" box="[201,540,896,946]" pageId="2" pageNumber="173">Spotted Linsang</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80A25FC5D70A894E6" box="[608,986,896,946]" pageId="2" pageNumber="173" type="nomenclature">
<paragraph id="EA133686636BFFF80A25FC5D70A894E6" blockId="2.[150,986,896,987]" box="[608,986,896,946]" pageId="2" pageNumber="173">
<heading id="B15B81EA636BFFF80A25FC5D70A894E6" box="[608,986,896,946]" pageId="2" pageNumber="173">
<taxonomicName id="2DAC4D05636BFFF80A25FC5D70A894E6" authorityName="Hodgson" authorityYear="1842" box="[608,986,896,946]" class="Bryopsida" family="Prionodontaceae" genus="Prionodon" kingdom="Plantae" order="Isobryales" pageId="2" pageNumber="173" phylum="Bryophyta" rank="species" species="pardicolor">
<emphasis id="D8D8EA94636BFFF80A25FC5D70A894E6" box="[608,986,896,946]" pageId="2" pageNumber="173">Prionodon pardicolor</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF808DDFC1970B9948D" box="[152,971,964,985]" pageId="2" pageNumber="173" type="vernacular_names">
<paragraph id="EA133686636BFFF808DDFC1970B9948D" blockId="2.[150,986,896,987]" box="[152,971,964,985]" pageId="2" pageNumber="173">
<heading id="B15B81EA636BFFF808DDFC1970B9948D" box="[152,971,964,985]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF808DDFC197396948D" box="[152,228,964,985]" pageId="2" pageNumber="173">French:</emphasis>
<vernacularName id="64AF46A8636BFFF808ABFC1972F9948D" box="[238,395,964,985]" pageId="2" pageNumber="173">Linsang tacheté</vernacularName>
/
<emphasis id="D8D8EA94636BFFF809E5FC197289948D" box="[416,507,964,985]" pageId="2" pageNumber="173">German:</emphasis>
<vernacularName id="64AF46A8636BFFF80A40FC1971E5948D" box="[517,663,964,985]" pageId="2" pageNumber="173">Fleckenlinsang</vernacularName>
/
<emphasis id="D8D8EA94636BFFF80AE9FC197075948D" box="[684,775,964,985]" pageId="2" pageNumber="173">Spanish:</emphasis>
<vernacularName id="64AF46A8636BFFF80B54FC1970B9948D" box="[785,971,964,985]" pageId="2" pageNumber="173">Linsang manchado</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80AB2FBD570329304" pageId="2" pageNumber="173" type="reference_group">
<paragraph id="EA133686636BFFF80AB2FBD570329304" blockId="2.[757,1351,1032,1458]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF80AB2FBD570E3937D" box="[759,913,1032,1065]" pageId="2" pageNumber="173">Taxonomy.</emphasis>
<taxonomicName id="2DAC4D05636BFFF80BE7FBD570499304" authority="Hodgson, 1842" authorityName="Hodgson" authorityYear="1842" class="Bryopsida" family="Prionodontaceae" genus="Prionodon" kingdom="Plantae" order="Isobryales" pageId="2" pageNumber="173" phylum="Bryophyta" rank="species" species="pardicolor">Prionodon pardicolor Hodgson, 1842</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80B25FBF277419323" pageId="2" pageNumber="173" type="materials_examined">
<paragraph id="EA133686636BFFF80B25FBF277419323" blockId="2.[757,1351,1032,1458]" pageId="2" pageNumber="173">
<materialsCitation id="5AC43CDB636BFFF80B25FBF277419323" ID-GBIF-Occurrence="3710100302" pageId="2" pageNumber="173">
“Sikim... Sub-Hemalayan mountains”. [
<collectingRegion id="2868F864636BFFF80B24FB8770B79323" box="[865,965,1114,1143]" country="India" name="Sikkim" pageId="2" pageNumber="173">Sikkim</collectingRegion>
,
<collectingCountry id="92BB7616636BFFF80B93FB8777549323" box="[982,1062,1114,1143]" name="India" pageId="2" pageNumber="173">India</collectingCountry>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80AB3FBA377B29392" pageId="2" pageNumber="173" type="discussion">
<paragraph id="EA133686636BFFF80AB3FBA377B29392" blockId="2.[757,1351,1032,1458]" pageId="2" pageNumber="173">
Two subspecies recognized,
<taxonomicName id="2DAC4D05636BFFF80CD3FBA3766593CB" authorityName="Hodgson" authorityYear="1842" box="[1174,1303,1150,1183]" class="Bryopsida" family="Prionodontaceae" genus="Prionodon" kingdom="Plantae" order="Isobryales" pageId="2" pageNumber="173" phylum="Bryophyta" rank="species" species="pardicolor">pardicolor</taxonomicName>
includes pardochrous, and perdicator.
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80AB2FB0D766992DF" pageId="2" pageNumber="173" type="distribution">
<caption id="BED3660E636BFFF80AB2FB0D766992DF" ID-DOI="http://doi.org/10.5281/zenodo.5714318" ID-Zenodo-Dep="5714318" httpUri="https://zenodo.org/record/5714318/files/figure.png" inLine="true" pageId="2" pageNumber="173" targetBox="[150,729,1043,1448]" targetPageId="2">
<paragraph id="EA133686636BFFF80AB2FB0D77F793B9" blockId="2.[757,1351,1032,1458]" box="[759,1157,1232,1261]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF80AB2FB0D77F793B9" box="[759,1157,1232,1261]" pageId="2" pageNumber="173">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph id="EA133686636BFFF80AB3FB2977A19268" blockId="2.[757,1351,1032,1458]" pageId="2" pageNumber="173">
<taxonomicName id="2DAC4D05636BFFF80AB3FB2977DA9241" authority="Hodgson, 1842" authorityName="Hodgson" authorityYear="1842" box="[758,1192,1268,1301]" class="Bryopsida" family="Prionodontaceae" genus="Prionodon" kingdom="Plantae" order="Isobryales" pageId="2" pageNumber="173" phylum="Bryophyta" rank="subSpecies" species="pardicolor" subSpecies="pardicolor">P. p. pardicolor Hodgson, 1842</taxonomicName>
—
<collectingCountry id="92BB7616636BFFF80C92FB29764D9241" box="[1239,1343,1268,1301]" name="Bhutan" pageId="2" pageNumber="173">Bhutan</collectingCountry>
, NE
<collectingCountry id="92BB7616636BFFF80B69FAFE70049268" box="[812,886,1315,1340]" name="India" pageId="2" pageNumber="173">India</collectingCountry>
, N
<collectingCountry id="92BB7616636BFFF80BEDFAFE775E9268" box="[936,1068,1315,1340]" name="Myanmar" pageId="2" pageNumber="173">Myanmar</collectingCountry>
, and
<collectingCountry id="92BB7616636BFFF80C3FFAFE77BD9268" box="[1146,1231,1315,1340]" name="Nepal" pageId="2" pageNumber="173">Nepal</collectingCountry>
.
</paragraph>
<paragraph id="EA133686636BFFF80AB3FA9F766992DF" blockId="2.[757,1351,1032,1458]" pageId="2" pageNumber="173">
<taxonomicName id="2DAC4D05636BFFF80AB3FA9F771B9237" authority="Thomas, 1925" authorityName="Thomas" authorityYear="1925" box="[758,1129,1346,1379]" class="Bryopsida" family="Prionodontaceae" genus="Prionodon" kingdom="Plantae" order="Isobryales" pageId="2" pageNumber="173" phylum="Bryophyta" rank="subSpecies" species="pardicolor" subSpecies="presina">P. p. presina Thomas, 1925</taxonomicName>
—
<collectingCountry id="92BB7616636BFFF80CD7FA9F76569237" box="[1170,1316,1346,1379]" name="Cambodia" pageId="2" pageNumber="173">Cambodia</collectingCountry>
, S
<collectingCountry id="92BB7616636BFFF80AB3FAB3703E92DF" box="[758,844,1390,1419]" name="China" pageId="2" pageNumber="173">China</collectingCountry>
,
<collectingCountry id="92BB7616636BFFF80B19FAB370EC92DF" box="[860,926,1390,1419]" name="Laos" pageId="2" pageNumber="173">Laos</collectingCountry>
, N
<collectingCountry id="92BB7616636BFFF80B8AFAB3772292DF" box="[975,1104,1390,1419]" name="Thailand" pageId="2" pageNumber="173">Thailand</collectingCountry>
, and
<collectingCountry id="92BB7616636BFFF80CD8FAB3766592DF" box="[1181,1303,1390,1419]" name="Vietnam" pageId="2" pageNumber="173">Vietnam</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80AB0FA4C7B979579" pageId="2" pageNumber="173" type="description">
<paragraph id="EA133686636BFFF80AB0FA4C7B979579" blockId="2.[757,1351,1032,1458]" lastBlockId="2.[1417,2627,289,2052]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF80AB0FA4C708192E6" box="[757,1011,1425,1458]" pageId="2" pageNumber="173">Descriptive notes.</emphasis>
Head-body 31-45 cm, tail 30-39.
<quantity id="2D549B63636BFFF80961FA6A7219928C" box="[292,363,1463,1496]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="173" unit="cm" value="9.0">9 cm</quantity>
; weight 550-1220 g. Ratio head-body/tail slightly superior to one. Two pairs of mammae. Adult males are larger than and may be twice as heavy as females. Has a genetlike aspect, with head and neck more elongated, pointed muzzle; nose pad brownish pink. Ears are medium-sized, broad at the base and rounded.Iris is dark brown to black. Pelage is short and very soft. Coat color is relatively uniform across the body, and varies from pale gray to yellow rufous on the back and flanks; ventral pelage pale gray to pale yellow. Individual variation exists in spot markings. Back exhibits two main pairs of lines of large dark spots rarely fused, and a characteristic pair of large nucchalstripes in continuation with the second row ofspots. A line of thin spots starting behind the shoulders forms a broken medio-dorsal stripe. Small spots are irregularly distributed on the flanks, shoulders and thighs. The rest of the hindlimbs and the forepaws are not spotted. Five claws fully retractile; presence of sheaths. Hindfeet digitigrade, forefeet plantigrade; hallucal lobe small and remote; absence of metatarsal pads. Tail is almost as long as head and body; thickly furred, with short hair; exhibits 7-9 broad dark rings. Confused annealing pattern at the whitish tip, with broad, distal rings often interspersed with thin ground-colored rings. Perineal glands are absent. Rostrum is moderately elongated; sagittal crest forms a large, flat stripe that fuses with the frontal bone. Caudal entotympanic bone poorly inflated ventrally when compared to the ectotympanic bone. Dental formula: 13/3, C1/1,P4/4,M 1/2 = 38. The brain weighs about
<quantity id="2D549B63636BFFF80DACFE637566968B" box="[1513,1556,446,479]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="2" pageNumber="173" unit="g" value="9.0">9 g</quantity>
. Geographic variations in coat pattern have been described. In presina, size is greater, dorsal spots are smaller and less regularly distributed, ground color—especially on the belly—is paler, and dark rings on the tail are narrower.
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80DD5FDE97546959D" pageId="2" pageNumber="173" type="biology_ecology">
<paragraph id="EA133686636BFFF80DD5FDE97546959D" blockId="2.[1417,2627,289,2052]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF80DD5FDE9768D9501" box="[1424,1535,564,597]" pageId="2" pageNumber="173">Habitat.</emphasis>
Primary and secondary moist forests, up to
<quantity id="2D549B63636BFFF8002AFDE97BA69501" box="[2159,2260,564,597]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="173" unit="m" value="2700.0">2700 m</quantity>
(e.g.
<collectingCountry id="92BB7616636BFFF8016CFDE97A0F9501" box="[2345,2429,564,597]" name="Nepal" pageId="2" pageNumber="173">Nepal</collectingCountry>
, Assam). Also recorded in mosaics of lowland riverine/sal forests and grassland (
<collectingCountry id="92BB7616636BFFF8016CFD827AF19528" box="[2345,2435,607,636]" name="Nepal" pageId="2" pageNumber="173">Nepal</collectingCountry>
), mixed bamboo forests along mountain rivers (
<collectingCountry id="92BB7616636BFFF80FCEFD5B7B7B95F7" box="[1931,2057,646,675]" name="Vietnam" pageId="2" pageNumber="173">Vietnam</collectingCountry>
), and disturbed evergreen forest (
<collectingCountry id="92BB7616636BFFF801B2FD5B76BD959D" name="Thailand" pageId="2" pageNumber="173">Thailand</collectingCountry>
,
<collectingCountry id="92BB7616636BFFF80DA5FD717555959D" box="[1504,1575,684,713]" name="Laos" pageId="2" pageNumber="173">Laos</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80DCAFD087A339378" pageId="2" pageNumber="173" type="food_feeding">
<paragraph id="EA133686636BFFF80DCAFD087A339378" blockId="2.[1417,2627,289,2052]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF80DCAFD0875E995A6" box="[1423,1691,725,754]" pageId="2" pageNumber="173">Food and Feeding.</emphasis>
Primarily carnivorous. Stomach contents in
<collectingCountry id="92BB7616636BFFF80166FD087AE895A6" box="[2339,2458,725,754]" name="Vietnam" pageId="2" pageNumber="173">Vietnam</collectingCountry>
showed remains of rodents, frogs, snakes and small birds. A captive female of
<quantity id="2D549B63636BFFF80115FD257AE9944D" box="[2384,2459,760,793]" metricMagnitude="-1" metricUnit="kg" metricValue="6.0" pageId="2" pageNumber="173" unit="g" value="600.0">600 g</quantity>
ate around
<quantity id="2D549B63636BFFF80DD7FCFD76A99415" box="[1426,1499,800,833]" metricMagnitude="-1" metricUnit="kg" metricValue="1.0" pageId="2" pageNumber="173" unit="g" value="100.0">100 g</quantity>
of meat per day, of which 76-5% was digested. Several authors have mentioned that the favorite food is passerine birds, which may be preyed upon either in trees or hiding in the grass. However,it is not known whether foraging more frequently occurs in trees or on the ground; a captive individual always ate on the ground. Small rodents are killed with a bite to the neck, whereas larger preys (e.g. rats) are leaped upon and finished whilst lying side-by-side on the ground. Reported to feed on carcasses killed by other predators. A captive specimen refused fish, eggs and fruits.
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80DC8FBEA7484926B" pageId="2" pageNumber="173" type="activity">
<paragraph id="EA133686636BFFF80DC8FBEA7484926B" blockId="2.[1417,2627,289,2052]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF80DC8FBEA75059300" box="[1421,1655,1079,1108]" pageId="2" pageNumber="173">Activity patterns.</emphasis>
Most of the active period (about eight hours) takes place during the night. Presumably arboreal, but from recent camera-trap records, may frequently use the ground to move and forage. A captive specimen was reported to spend most ofits time at heights up to
<quantity id="2D549B63636BFFF80EFDFB74759B939E" box="[1720,1769,1193,1226]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="173" unit="m" value="1.0">1 m</quantity>
, although wild individuals were sighted higher in the canopy. Shelter in trees. Good climber; springs and climbs in the manner of cats; semi-digitigrade, with plantigrade forefeet and digitigrade hindfeet; uses a “head-first” vertical descent of trees. Jumps may reach
<quantity id="2D549B63636BFFF80F32FAFF74D5926B" box="[1911,1959,1314,1343]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="173" unit="m" value="1.0">1 m</quantity>
high.
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80DC8FA977A9092E1" pageId="2" pageNumber="173" type="biology_ecology">
<paragraph id="EA133686636BFFF80DC8FA977A9092E1" blockId="2.[1417,2627,289,2052]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF80DC8FA977B379233" box="[1421,2117,1354,1383]" pageId="2" pageNumber="173">Movements, Home range and Social organization.</emphasis>
Almost unknown. Probably solitary. A female captive specimen marked territory with urine and feces. When scared, may emit a squeak accompanied by a “drumming” on the ground with one forepaw.
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80DC8FA6175C2917E" pageId="2" pageNumber="173" type="breeding">
<paragraph id="EA133686636BFFF80DC8FA6175C2917E" blockId="2.[1417,2627,289,2052]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF80DC8FA6175619289" box="[1421,1555,1468,1501]" pageId="2" pageNumber="173">Breeding.</emphasis>
Litters reported to be of two cubs, once or twice a year, between February and August (three litters in one year has been reported once in captivity). Dens are located in tree holes.
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80DC8F9EF74A29031" pageId="2" pageNumber="173" type="conservation">
<paragraph id="EA133686636BFFF80DC8F9EF74A29031" blockId="2.[1417,2627,289,2052]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF80DC8F9EF759A9107" box="[1421,1768,1586,1619]" pageId="2" pageNumber="173">Status and Conservation.</emphasis>
CITES Appendix I. Classified as Least Concern on The [UCN Red List. Widespread distribution over South-east Asia but individuals rarely sighted, especially in
<collectingCountry id="92BB7616636BFFF80E63F95D75DB91F5" box="[1574,1705,1664,1697]" name="Myanmar" pageId="2" pageNumber="173">Myanmar</collectingCountry>
,
<collectingCountry id="92BB7616636BFFF80EFFF95D744B91F5" box="[1722,1849,1664,1697]" name="Thailand" pageId="2" pageNumber="173">Thailand</collectingCountry>
and
<collectingCountry id="92BB7616636BFFF80FC1F95D7B6491F5" box="[1924,2070,1664,1697]" name="Cambodia" pageId="2" pageNumber="173">Cambodia</collectingCountry>
. Recent records from camera-trapping however suggest that the species is not uncommon in some parts of its range (e.g.
<collectingRegion id="2868F864636BFFF80DCCF90E768B91A4" box="[1417,1529,1747,1776]" country="China" name="Yunnan" pageId="2" pageNumber="173">Yunnan</collectingRegion>
,
<collectingCountry id="92BB7616636BFFF80E4DF90E751091A4" box="[1544,1634,1747,1776]" name="China" pageId="2" pageNumber="173">China</collectingCountry>
). Major threats are habitat loss (e.g. slash and burn shifting cultivation) and hunting for fur. Most of the skins kept in museums were collected from natives. Listed on Category II of the
<collectingCountry id="92BB7616636BFFF80F5EF8C3741D906B" box="[1819,1903,1822,1855]" name="China" pageId="2" pageNumber="173">China</collectingCountry>
Wildlife Protection Law; totally protected in
<collectingCountry id="92BB7616636BFFF801A9F8C37944906B" box="[2540,2614,1822,1855]" name="India" pageId="2" pageNumber="173">India</collectingCountry>
,
<collectingCountry id="92BB7616636BFFF80DC9F891757D9031" box="[1420,1551,1868,1893]" name="Myanmar" pageId="2" pageNumber="173">Myanmar</collectingCountry>
,
<collectingCountry id="92BB7616636BFFF80E65F89175069031" box="[1568,1652,1868,1893]" name="Nepal" pageId="2" pageNumber="173">Nepal</collectingCountry>
,
<collectingCountry id="92BB7616636BFFF80EC1F89174779031" box="[1668,1797,1868,1893]" name="Thailand" pageId="2" pageNumber="173">Thailand</collectingCountry>
, and
<collectingCountry id="92BB7616636BFFF80F17F89174BE9031" box="[1874,1996,1868,1893]" name="Vietnam" pageId="2" pageNumber="173">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B6650D636BFFF80DC9F8A475329F55" pageId="2" pageNumber="173" type="bibRefCitation_list">
<paragraph id="EA133686636BFFF80DC9F8A475329F55" blockId="2.[1417,2627,289,2052]" pageId="2" pageNumber="173">
<emphasis id="D8D8EA94636BFFF80DC9F8A4755190DA" box="[1420,1571,1913,1934]" pageId="2" pageNumber="173">Bibliography.</emphasis>
Boitani et al. (2006), Choudhury (1997a, 1997b, 2002), Duckworth (1997), Duckworth et al. (1999), Hodgson (1842, 1847), Kuznetzov &amp; Baranauskas (1993), Pham-chong-Ahn (1980), Pocock (1915¢, 1933c, 1935), Radinsky (1975), Ramakantha (1994), Schreiber et al. (1989), Sunquist (1982), Taylor (1988), Tizard (2002), Van Rompaey (1995).
</paragraph>
</subSubSection>
</treatment>
</document>

216
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@ -0,0 +1,216 @@
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<mods:title id="5959376E577A9624886335940C2845F5">Review of the genus Carlschoenherria (Coleoptera: Scarabaeidae: Melolonthinae) of the Philippines</mods:title>
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<subSubSection id="A2B66544B4309418FF2AFE68FD195C88" box="[151,720,476,503]" pageId="5" pageNumber="335" type="nomenclature">
<paragraph id="EA1336CFB4309418FF2AFE68FD195C88" blockId="5.[151,720,476,539]" box="[151,720,476,503]" pageId="5" pageNumber="335">
<heading id="B15B81A3B4309418FF2AFE68FD195C88" bold="true" box="[151,720,476,503]" fontSize="11" level="1" pageId="5" pageNumber="335" reason="1">
<emphasis id="D8D8EADDB4309418FF2AFE68FD195C88" bold="true" box="[151,720,476,503]" pageId="5" pageNumber="335">
<taxonomicName id="2DAC4D4CB4309418FF2AFE68FDF05C88" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[151,569,476,503]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="gapudi" status="sp. nov.">
<emphasis id="D8D8EADDB4309418FF2AFE68FE0E5C89" bold="true" box="[151,455,476,503]" italics="true" pageId="5" pageNumber="335">Carlschoenherria gapudi</emphasis>
Calcetas
</taxonomicName>
<taxonomicNameLabel id="C3EB57A6B4309418FDFDFE69FD195C88" box="[576,720,476,503]" pageId="5" pageNumber="335" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B4309418FF2AFDB4FE835F65" box="[151,330,512,539]" pageId="5" pageNumber="335" type="description">
<paragraph id="EA1336CFB4309418FF2AFDB4FE835F65" blockId="5.[151,720,476,539]" box="[151,330,512,539]" pageId="5" pageNumber="335">
(
<figureCitation id="72972A4AB4309418FF22FDB4FED05F65" box="[159,281,512,539]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1970,1995]" captionTargetBox="[273,1273,647,1916]" captionTargetId="figure@6.[376,1212,669,1951]" captionTargetPageId="6" captionText="FIGURES 9–12. Carlschoenherria gapudi Calcetas, new species, holotype, male. 9, habitus, dorsal aspect; 10, antenna, lateral aspect; 11, genitalia, dorsal aspect; 12, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3249663/files/figure.png" pageId="5" pageNumber="335">Figs. 9–12</figureCitation>
,
<figureCitation id="72972A4AB4309418FE98FDB4FE8B5F65" box="[293,322,512,539]" captionStart="FIGURE 36" captionStartId="15.[151,250,1710,1735]" captionTargetBox="[360,1227,424,1689]" captionTargetId="figure@15.[340,1247,416,1691]" captionTargetPageId="15" captionText="FIGURE 36. Geographical distribution map of Carlschoenherria Bezděk, 2016 in the Philippines." httpUri="https://zenodo.org/record/3249673/files/figure.png" pageId="5" pageNumber="335">36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B4309418FF2AFDFDFBC25FF9" pageId="5" pageNumber="335" type="materials_examined">
<paragraph id="EA1336CFB4309418FF2AFDFDFD445F1C" blockId="5.[151,1437,584,2015]" box="[151,653,584,611]" pageId="5" pageNumber="335">
<materialsCitation id="5AC43C92B4309418FF2AFDFDFD405F1C" ID-GBIF-Occurrence="2268317661" box="[151,649,584,611]" country="Philippines" location="Mindanao Island" pageId="5" pageNumber="335" specimenCount="1" typeStatus="holotype">
<emphasis id="D8D8EADDB4309418FF2AFDFDFEFF5F1D" bold="true" box="[151,310,584,611]" pageId="5" pageNumber="335">
<typeStatus id="3517886DB4309418FF2AFDFDFF1B5F1D" box="[151,210,585,611]" pageId="5" pageNumber="335">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="92BB765FB4309418FE81FDFCFE745F1C" box="[316,445,584,610]" name="Philippines" pageId="5" pageNumber="335">Philippines</collectingCountry>
,
<location id="EF736014B4309418FE75FDFCFD405F1C" LSID="urn:lsid:plazi:treatment:620587D9B430941BFF2AFE68FA8B5FF9:EF736014B4309418FE75FDFCFD405F1C" box="[456,649,584,610]" country="Philippines" name="Mindanao Island" pageId="5" pageNumber="335">Mindanao Island</location>
</materialsCitation>
.
</paragraph>
<paragraph id="EA1336CFB4309418FF7AFDD9FBC25FF9" blockId="5.[151,1437,584,2015]" box="[199,1035,620,647]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AFDD9FEBC5FF9" bold="true" box="[199,373,620,647]" pageId="5" pageNumber="335">Type material.</emphasis>
<materialsCitation id="5AC43C92B4309418FEC1FDD8FBC25FF9" ID-GBIF-Occurrence="2268317648" box="[380,1035,620,647]" collectingDate="1993-05" collectionCode="NMM" collectorName="V. Samarita" pageId="5" pageNumber="335" specimenCount="1" typeStatus="holotype">
<typeStatus id="3517886DB4309418FEC1FDD8FE2D5FF8" box="[380,484,620,646]" pageId="5" pageNumber="335" type="holotype">Holotype</typeStatus>
(male), labeled:
<collectingDate id="8E56E9E7B4309418FD1FFDD9FCDE5FF9" box="[674,791,621,647]" pageId="5" pageNumber="335" value="1993-05">
<date id="9E12100FB4309418FD1FFDD9FCDE5FF9" box="[674,791,621,647]" pageId="5" pageNumber="335" value="1993-05">May 1993</date>
</collectingDate>
,
<collectorName id="47595319B4309418FC9FFDD9FC6D5FF8" box="[802,932,620,647]" pageId="5" pageNumber="335">V. Samarita</collectorName>
(
<collectionCode id="8CBDAE0AB4309418FC0EFDD9FC365FF9" box="[947,1023,621,647]" pageId="5" pageNumber="335">NMM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B4309418FF7AFD24FB365AC5" pageId="5" pageNumber="335" type="description">
<paragraph id="EA1336CFB4309418FF7AFD24FC3F5F8D" blockId="5.[151,1437,584,2015]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AFD24FDEB5FD5" bold="true" box="[199,546,656,683]" pageId="5" pageNumber="335">
Description of
<typeStatus id="3517886DB4309418FEC8FD25FE145FD5" box="[373,477,657,683]" pageId="5" pageNumber="335" type="holotype">holotype</typeStatus>
, male
</emphasis>
. Length 21.3 mm, width 8.7 mm. Body dichromatic, head, pronotum and scutel- lum blackish brown; anterior half of elytra dark reddish brown with blackish tinge, posterior half with dark aluta- ceous tinge, covered with yellowish white setae; leg dark alutaceous (
<figureCitation id="72972A4AB4309418FC19FD6CFC215F8D" box="[932,1000,728,755]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1970,1995]" captionTargetBox="[273,1273,647,1916]" captionTargetId="figure@6.[376,1212,669,1951]" captionTargetPageId="6" captionText="FIGURES 9–12. Carlschoenherria gapudi Calcetas, new species, holotype, male. 9, habitus, dorsal aspect; 10, antenna, lateral aspect; 11, genitalia, dorsal aspect; 12, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3249663/files/figure.png" pageId="5" pageNumber="335">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="EA1336CFB4309418FF7AFD48FB395ED8" blockId="5.[151,1437,584,2015]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AFD48FECD5E68" box="[199,260,764,790]" italics="true" pageId="5" pageNumber="335">Head</emphasis>
. Clypeus with anterior margin nearly straight dorsally.Antennomere III with short stalk, widely expanded apically; antennomere IV distinctly shorter than the rest of lamellae antennomeres. Lamellae length 2.5 mm, nearly as long as anterior margin of the clypeus. Terminal maxillary palpomere with distinctly long, shallow lanceolate shaped groove or depression, as long as antennomere VI. Mentum with indistinct latitudinal mesal carina, with shal- low depression below it; surface subrugosely punctate, covered with few setae, acicular-like scales.
</paragraph>
<paragraph id="EA1336CFB4309418FF7AFC05FE255901" blockId="5.[151,1437,584,2015]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AFC05FEFF5EB4" box="[199,310,945,970]" italics="true" pageId="5" pageNumber="335">Pronotum</emphasis>
anterior margin concave, nearly straight mesally, weakly explanate, margin smooth mesally; margin towards anterior angle sloping upward, crenulate; anterior angle rounded, obtuse; margin crenulate, raised dorsally; lateral margin distinctly convex mesally, margin minutely crenulate, thinly carinate; posterior angle rounded, nearly at 90
<superScript id="1DD99B87B4309418FF70FBAFFF1C5957" attach="left" box="[205,213,1051,1065]" fontSize="6" pageId="5" pageNumber="335">0</superScript>
angle, margin crenulate; posterior margin deeply concave mesally, smooth; surface rugosely punctate mesally and minutely punctate on each side; with narrow, shallow longitudinal groove or depression mesally, groove with long yellowish setae (
<figureCitation id="72972A4AB4309418FE36FBD0FE165901" box="[395,479,1124,1151]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1970,1995]" captionTargetBox="[273,1273,647,1916]" captionTargetId="figure@6.[376,1212,669,1951]" captionTargetPageId="6" captionText="FIGURES 9–12. Carlschoenherria gapudi Calcetas, new species, holotype, male. 9, habitus, dorsal aspect; 10, antenna, lateral aspect; 11, genitalia, dorsal aspect; 12, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3249663/files/figure.png" pageId="5" pageNumber="335">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="EA1336CFB4309418FF7AFB3CFAE559B8" blockId="5.[151,1437,584,2015]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AFB3CFEFC59DC" box="[199,309,1160,1186]" italics="true" pageId="5" pageNumber="335">Scutellum</emphasis>
anterior margin nearly straight; posterior margin widely concave or rounded apically; with few rounded punctures; surface with rounded, sparsely spaced punctures mesally, slightly rugged posteriorly.
</paragraph>
<paragraph id="EA1336CFB4309418FF7AFB64FDD658E1" blockId="5.[151,1437,584,2015]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AFB64FED35994" box="[199,282,1232,1258]" italics="true" pageId="5" pageNumber="335">Elytron</emphasis>
with ten costae and ten intervals, each interval with dense, thick, stiff; setae longitudinally aligned along each interval; costae I–III wide, lustrous, impunctate; costae III–V coalesced apically; humeral margin distinctly wedge-shaped towards anterior angle; anterior angle rounded; lateral margin slightly convex mesally; apical margin widely rounded; sutural angle angulate, obtuse; sutural margin carinate near sutural angle and mesally, disappear- ing towards scutellum; surface covered with numerous, very narrow, elongate lanceolate to acicular-like, tapered apically white and yellowish setae.
</paragraph>
<paragraph id="EA1336CFB4309418FF7AFA1DFBCF5898" blockId="5.[151,1437,584,2015]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AFA1DFECD58BC" box="[199,260,1449,1474]" italics="true" pageId="5" pageNumber="335">Legs.</emphasis>
Foretibiae with two teeth. Metatibiae with numerous long, short whitish scales; with numerous, sparsely distributed long brownish stiff setae; posterior metatibiae with 15–17 spicules.
</paragraph>
<paragraph id="EA1336CFB4309418FF7AFA44FE9D5B08" blockId="5.[151,1437,584,2015]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AFA44FE0A5B74" box="[199,451,1520,1547]" italics="true" pageId="5" pageNumber="335">Ventral side of thorax.</emphasis>
Prosternal process spindle-shaped, slender longitudinally, truncate apically, tapered ba- sally. Metaventral process little prolonged, not reaching procoxae, attached mesally to the mesosternal process; metaventral process length 1.3 mm. Pro and mesosterna densely covered with long yellowish hairs except sparsely covered mesally.
</paragraph>
<paragraph id="EA1336CFB4309418FF7AF934FEC95A54" blockId="5.[151,1437,584,2015]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AF934FE605BE4" box="[199,425,1664,1690]" italics="true" pageId="5" pageNumber="335">Abdominal ventrites</emphasis>
. Covered mostly with long hairs, long elongate lanceolate to acicular-like scales; abdomi- nal ventrites not uniformly covered with pubescence, lacking in pubescence at least near each anterior and posterior margins or rather narrowly restricted in the distribution of pubescence (
<bibRefCitation id="8E3D4B3EB4309418FC00F97CFBF95B9D" author="Itoh, T." box="[957,1072,1736,1763]" pageId="5" pageNumber="335" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh 1993</bibRefCitation>
); sides of abdominal ventrites II and IV with large oval shaped patches of scales or abdominal maculation; abdominal ventrite V with small patch of scales.
</paragraph>
<paragraph id="EA1336CFB4309418FF7AF880FBF25A31" blockId="5.[151,1437,584,2015]" box="[199,1083,1844,1871]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AF880FEF35A30" box="[199,314,1844,1870]" italics="true" pageId="5" pageNumber="335">Pygidium.</emphasis>
Anterior margin arched or convex mesally, posterior margin narrow.
</paragraph>
<paragraph id="EA1336CFB4309418FF7AF8ECFB365AC5" blockId="5.[151,1437,584,2015]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AF8ECFEFD5A0C" box="[199,308,1880,1906]" italics="true" pageId="5" pageNumber="335">Aedeagus</emphasis>
. Posterior margin of parameres with elongate rounded protuberance, margin weakly concave towards lateral margin, longitudinally undulate towards apex. Length of apical process 1.0 mm from lateral view; apical margin bird-bill like laterally. Parameres and phallobase combined length 4.6 mm; as in
<figureCitation id="72972A4AB4309418FBC9F814FB335AC5" box="[1140,1274,1952,1979]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1970,1995]" captionTargetBox="[273,1273,647,1916]" captionTargetId="figure@6.[376,1212,669,1951]" captionTargetPageId="6" captionText="FIGURES 9–12. Carlschoenherria gapudi Calcetas, new species, holotype, male. 9, habitus, dorsal aspect; 10, antenna, lateral aspect; 11, genitalia, dorsal aspect; 12, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3249663/files/figure.png" pageId="5" pageNumber="335">Figs. 11, 12</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B4309418FF7AF870FD3F5AA1" box="[199,758,1988,2015]" pageId="5" pageNumber="335" type="distribution">
<paragraph id="EA1336CFB4309418FF7AF870FD3F5AA1" blockId="5.[151,1437,584,2015]" box="[199,758,1988,2015]" pageId="5" pageNumber="335">
<emphasis id="D8D8EADDB4309418FF7AF870FEA85AA1" bold="true" box="[199,353,1988,2015]" pageId="5" pageNumber="335">Distribution.</emphasis>
The
<collectingCountry id="92BB765FB4309418FE24F870FDDE5AA0" box="[409,535,1988,2014]" name="Philippines" pageId="5" pageNumber="335">Philippines</collectingCountry>
(Mindanao Island).
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B433941BFF7AFF2DFED35C61" pageId="6" pageNumber="336" type="etymology">
<paragraph id="EA1336CFB433941BFF7AFF2DFED35C61" blockId="6.[151,1437,153,647]" pageId="6" pageNumber="336">
<emphasis id="D8D8EADDB433941BFF7AFF2DFE875DCD" bold="true" box="[199,334,153,179]" pageId="6" pageNumber="336">Etymology.</emphasis>
This new species is named after the late Dr. Victor P. Gapud, Professor Emeritus of Taxonomy and Systematics at the Crop Protection Cluster, University of the
<collectingCountry id="92BB765FB433941BFCECFF09FC065DA9" box="[849,975,189,215]" name="Philippines" pageId="6" pageNumber="336">Philippines</collectingCountry>
Los Baños. He is a versatile taxonomist working on different groups of Philippine insects such as Collembola, Odonata, Hemiptera and selected groups of
<taxonomicName id="2DAC4D4CB433941BFF2AFEB1FEDF5C61" box="[151,278,261,287]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B433941BFF7AFE9CFA8B5FF9" pageId="6" pageNumber="336" type="diagnosis">
<paragraph id="EA1336CFB433941BFF7AFE9CFACD5F41" blockId="6.[151,1437,153,647]" pageId="6" pageNumber="336">
<emphasis id="D8D8EADDB433941BFF7AFE9CFE075C3D" bold="true" box="[199,462,296,323]" pageId="6" pageNumber="336">Differential diagnosis.</emphasis>
Based on the species-group classification of
<bibRefCitation id="8E3D4B3EB433941BFC74FE9DFB835C3D" author="Itoh, T." box="[969,1098,297,323]" pageId="6" pageNumber="336" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh (1993)</bibRefCitation>
, this species belongs to the
<taxonomicName id="2DAC4D4CB433941BFA3EFE9DFEDC5C19" authorityName="Laporte" authorityYear="1840" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="sulcipennis">
<emphasis id="D8D8EADDB433941BFA3EFE9DFEDC5C19" italics="true" pageId="6" pageNumber="336">C. sulcipennis</emphasis>
</taxonomicName>
species-group, the parameres are laterally compressed and bird-bill shaped at apex. Further, it can be classified into the
<taxonomicName id="2DAC4D4CB433941BFEDAFEC5FDC65CF5" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Brenske" baseAuthorityYear="1894" box="[359,527,369,395]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="philippinica">
<emphasis id="D8D8EADDB433941BFEDAFEC5FDC65CF5" box="[359,527,369,395]" italics="true" pageId="6" pageNumber="336">C. philippinica</emphasis>
</taxonomicName>
species-subgroup, its pronotum has a longitudinal groove along the median line and the abdominal ventrites are lacking in setation at least near each anterior and posterior margins or rather narrowly restricted in the distribution of setation. In addition, the shape of the parameres resembles that of
<taxonomicName id="2DAC4D4CB433941BFA8DFE0DFF145C89" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Brenske" baseAuthorityYear="1894" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="philippinica">
<emphasis id="D8D8EADDB433941BFA8DFE0DFF145C89" italics="true" pageId="6" pageNumber="336">C. philippinica</emphasis>
</taxonomicName>
. In comparison to
<taxonomicName id="2DAC4D4CB433941BFE08FE69FD975C89" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Brenske" baseAuthorityYear="1894" box="[437,606,477,503]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="philippinica">
<emphasis id="D8D8EADDB433941BFE08FE69FD975C89" box="[437,606,477,503]" italics="true" pageId="6" pageNumber="336">C. philippinica</emphasis>
</taxonomicName>
, antennomere III of
<taxonomicName id="2DAC4D4CB433941BFCF6FE69FBEC5C89" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[843,1061,477,503]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="gapudi" status="sp. nov.">
<emphasis id="D8D8EADDB433941BFCF6FE69FC725C89" box="[843,955,477,503]" italics="true" pageId="6" pageNumber="336">C. gapudi</emphasis>
Calcetas
</taxonomicName>
,
<taxonomicNameLabel id="C3EB57A6B433941BFB8CFE69FB705C89" box="[1073,1209,477,503]" pageId="6" pageNumber="336" rank="species">new species</taxonomicNameLabel>
is widely expanded apically and narrowly expanded in the former (
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). Also, the setation and placement of patches of setae on the abdominal ventrites of
<taxonomicName id="2DAC4D4CB433941BFE27FD91FD8B5F41" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Brenske" baseAuthorityYear="1894" box="[410,578,549,575]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="philippinica">
<emphasis id="D8D8EADDB433941BFE27FD91FD8B5F41" box="[410,578,549,575]" italics="true" pageId="6" pageNumber="336">C. philippinica</emphasis>
</taxonomicName>
and
<taxonomicName id="2DAC4D4CB433941BFDC4FD91FC995F41" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[633,848,549,575]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="336" phylum="Arthropoda" rank="species" species="gapudi" status="sp. nov.">
<emphasis id="D8D8EADDB433941BFDC4FD91FD2E5F41" box="[633,743,549,575]" italics="true" pageId="6" pageNumber="336">C. gapudi</emphasis>
Calcetas
</taxonomicName>
,
<taxonomicNameLabel id="C3EB57A6B433941BFCE6FD91FC2B5F41" box="[859,994,549,575]" pageId="6" pageNumber="336" rank="species">new species</taxonomicNameLabel>
are similar (
<figureCitation id="72972A4AB433941BFBD2FD91FB3E5F41" box="[1135,1271,549,575]" captionStart="FIGURES 13–15" captionStartId="7.[151,264,1257,1282]" captionTargetBox="[368,1220,181,1227]" captionTargetId="figure@7.[368,1220,181,1237]" captionTargetPageId="7" captionText="FIGURES 13–15. Abdomen, ventral aspect. 13, Carlschoenherria philippinica (Brenske), male; 14, Carlschoenherria gapudi Calcetas, new species, male; 15, Carlschoenherria sulcipennis (Laporte), male." httpUri="https://zenodo.org/record/3249665/files/figure.png" pageId="6" pageNumber="336">Figs. 13, 14</figureCitation>
).
</paragraph>
<paragraph id="EA1336CFB433941BFF7AFDFDFA8B5FF9" blockId="6.[151,1437,153,647]" pageId="6" pageNumber="336">The collector failed to cite the locality of the specimen. However, examining the other melolonthine specimens collected by Dr. Samarita, he was collecting somewhere in the island of Mindanao during that time period.</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="EA1336CFB432941AFF2AFACCFC5358C8" blockId="7.[151,1190,1400,1462]" pageId="7" pageNumber="337">
<heading id="B15B81A3B432941AFF2AFACCFC5358C8" bold="true" fontSize="11" level="1" pageId="7" pageNumber="337" reason="1">
<emphasis id="D8D8EADDB432941AFF2AFACCFC5358C8" bold="true" pageId="7" pageNumber="337">
Key to the species-group and species of
<taxonomicName id="2DAC4D4CB432941AFD38FACCFC9258ED" authorityName="sensu Itoh" authorityYear="1992" box="[645,859,1400,1427]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="D8D8EADDB432941AFD38FACCFC9258ED" bold="true" box="[645,859,1400,1427]" italics="true" pageId="7" pageNumber="337">Carlschoenherria</emphasis>
</taxonomicName>
(modified from
<bibRefCitation id="8E3D4B3EB432941AFB9AFACCFB5758EC" author="Itoh, T." box="[1063,1182,1400,1426]" pageId="7" pageNumber="337" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh 1993</bibRefCitation>
) from the
<collectingCountry id="92BB765FB432941AFEB7FA28FE5158C8" box="[266,408,1436,1462]" name="Philippines" pageId="7" pageNumber="337">Philippines</collectingCountry>
, the Malay Peninsula, Sumatra and Java
</emphasis>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="EA1336CFB432941AFF2AFA55FA555888" blockId="7.[151,1437,1504,2025]" box="[151,1436,1504,1526]" pageId="7" pageNumber="337">
<keyLead id="515D95FAB432941AFF2AFA55FA555888" box="[151,1436,1504,1526]" pageId="7" pageNumber="337">
1 Parameres compressed laterally, bird-bill shaped at apex (
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<emphasis id="D8D8EADDB432941AFCB1FA55FC5D5888" box="[780,916,1505,1526]" italics="true" pageId="7" pageNumber="337">C. sulcipennis</emphasis>
</taxonomicName>
species-group)............................. 2
</keyLead>
</paragraph>
<paragraph id="EA1336CFB432941AFF2AFA4AFA555B6D" blockId="7.[151,1437,1504,2025]" box="[151,1436,1534,1556]" pageId="7" pageNumber="337">
<keyLead id="515D95FAB432941AFF2AFA4AFA555B6D" box="[151,1436,1534,1556]" pageId="7" pageNumber="337">
- Parameres compressed dorsally, developed laterally (
<taxonomicName id="2DAC4D4CB432941AFD5DFA4AFC8A5B6D" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Arrow" baseAuthorityYear="1938" box="[736,835,1534,1555]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="hastata">
<emphasis id="D8D8EADDB432941AFD5DFA4AFC8A5B6D" box="[736,835,1534,1555]" italics="true" pageId="7" pageNumber="337">C. hastata</emphasis>
</taxonomicName>
species-group).................................... 7
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<keyLead id="515D95FAB432941AFF2AF9AFFA555B4E" box="[151,1436,1563,1585]" pageId="7" pageNumber="337">
2 Parameres with anterior margin wide, expanded laterally (the
<taxonomicName id="2DAC4D4CB432941AFC8EF9A8FC625B4E" authorityName="Calcetas" authorityYear="2019" box="[819,939,1563,1585]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="adoradae">
<emphasis id="D8D8EADDB432941AFC8EF9A8FC625B4E" box="[819,939,1563,1585]" italics="true" pageId="7" pageNumber="337">C. adoradae</emphasis>
</taxonomicName>
species-subgroup)......................... 3
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<keyLead id="515D95FAB432941AFF2AF98DFA555B30" box="[151,1436,1592,1614]" pageId="7" pageNumber="337">- Parameres with anterior margin narrow, not expanded laterally................................................. 4</keyLead>
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<paragraph id="EA1336CFB432941AFF2AF9E2FA555BBA" blockId="7.[151,1437,1504,2025]" pageId="7" pageNumber="337">
<keyLead id="515D95FAB432941AFF2AF9E2FA555BBA" pageId="7" pageNumber="337">
3 Anterior margin of parameres explanate, undulate dorsally towards phallobase; pronotum with anterior margin distinctly Vshaped, rounded mesally; anterior angle nearly at 90
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angle; pronotum with posterior margin entirely nearly straight, minutely concave mesally; anterior margin of clypeus nearly straight dorsally (Figs. 1–4).
<collectingCountry id="92BB765FB432941AFC57F924FB9F5BD8" box="[1002,1110,1680,1702]" name="Philippines" pageId="7" pageNumber="337">Philippines</collectingCountry>
...................................................................................................
<taxonomicName id="2DAC4D4CB432941AFBF3F91AFAE95BBA" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[1102,1312,1710,1732]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="adoradae" status="sp. nov.">
<emphasis id="D8D8EADDB432941AFBF3F91AFB0F5BBD" box="[1102,1222,1710,1731]" italics="true" pageId="7" pageNumber="337">C. adoradae</emphasis>
Calcetas
</taxonomicName>
,
<taxonomicNameLabel id="C3EB57A6B432941AFA94F91AFA555BBA" box="[1321,1436,1710,1732]" pageId="7" pageNumber="337" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="EA1336CFB432941AFF2AF978FA555A62" blockId="7.[151,1437,1504,2025]" pageId="7" pageNumber="337">
<keyLead id="515D95FAB432941AFF2AF978FA555A62" pageId="7" pageNumber="337">
- Anterior margin of parameres horse saddle-shaped laterally; pronotum with anterior margin concave, nearly straight mesally; anterior angle obtuse; pronotum posterior margin distinctly concave mesally; anterior margin of clypeus slightly concave dor- sally (
<figureCitation id="72972A4AB432941AFEA1F8B2FEB35A62" box="[284,378,1798,1820]" captionStart="FIGURES 5–8" captionStartId="4.[151,264,1930,1955]" captionTargetBox="[340,1247,705,1896]" captionTargetId="figure@4.[340,1247,705,1910]" captionText="FIGURES 5–8. Carlschoenherria hadsallae Calcetas, new species, holotype, male. 5, habitus, dorsal aspect; 6, pronotum, dorsal aspect; 7, genitalia, dorsal aspect; 8, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3407336/files/figure.png" pageId="7" pageNumber="337">Figs. 5–8</figureCitation>
).
<collectingCountry id="92BB765FB432941AFE36F8B2FE3E5A62" box="[395,503,1798,1820]" name="Philippines" pageId="7" pageNumber="337">Philippines</collectingCountry>
.................................................
<taxonomicName id="2DAC4D4CB432941AFBF1F8B2FAE95A62" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[1100,1312,1798,1820]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="hadsallae" status="sp. nov.">
<emphasis id="D8D8EADDB432941AFBF1F8B2FB0C5A65" box="[1100,1221,1798,1819]" italics="true" pageId="7" pageNumber="337">C. hadsallae</emphasis>
Calcetas
</taxonomicName>
,
<taxonomicNameLabel id="C3EB57A6B432941AFA94F8B2FA555A62" box="[1321,1436,1798,1820]" pageId="7" pageNumber="337" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51582E6AB432941AFF2AF897FA555A0A" pageId="7" pageNumber="337">
<paragraph id="EA1336CFB432941AFF2AF897FA545A29" blockId="7.[151,1437,1504,2025]" pageId="7" pageNumber="337">
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4 Pronotum without mesal longitudinal groove or depression (
<figureCitation id="72972A4AB432941AFC99F897FCA35A47" box="[804,874,1827,1849]" captionStart="FIGURES 16–19" captionStartId="8.[151,264,1979,2004]" captionTargetBox="[259,1424,674,1941]" captionTargetId="figure@8.[352,1235,697,1960]" captionTargetPageId="8" captionText="FIGURES 16–19. Carlschoenherria palawana (Moser), male from the Philippines, Palawan. 16, habitus, dorsal aspect; 17, pronotum, dorsal aspect; 18, genitalia, dorsal aspect; 19, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3249667/files/figure.png" pageId="7" pageNumber="337">Fig. 16</figureCitation>
) (
<taxonomicName id="2DAC4D4CB432941AFCC2F890FBCA5A46" baseAuthorityName="Brenske" baseAuthorityYear="1894" box="[895,1027,1827,1849]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="D8D8EADDB432941AFCC2F890FBCA5A46" box="[895,1027,1827,1849]" italics="true" pageId="7" pageNumber="337">C. borneensis</emphasis>
</taxonomicName>
species-subgroup).
<collectingCountry id="92BB765FB432941AFB79F897FAF95A47" box="[1220,1328,1827,1849]" name="Philippines" pageId="7" pageNumber="337">Philippines</collectingCountry>
...........................................................................................
<taxonomicName id="2DAC4D4CB432941AFB73F8F5FA545A29" authority="(Moser)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[1230,1437,1857,1879]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="palawana">
<emphasis id="D8D8EADDB432941AFB73F8F5FA805A28" box="[1230,1353,1857,1878]" italics="true" pageId="7" pageNumber="337">C. palawana</emphasis>
(Moser)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EA1336CFB432941AFF2AF8EAFA555A0A" blockId="7.[151,1437,1504,2025]" box="[151,1436,1886,1908]" pageId="7" pageNumber="337">
<keyLead id="515D95FAB432941AFF2AF8EAFA555A0A" box="[151,1436,1886,1908]" pageId="7" pageNumber="337">
- Pronotum with mesal longitudinal groove or depression (
<figureCitation id="72972A4AB432941AFCBBF8EAFC9F5A0D" box="[774,854,1886,1908]" captionStart="FIGURES 20–23" captionStartId="10.[151,264,1667,1692]" captionTargetBox="[356,1231,345,1644]" captionTargetId="figure@10.[356,1232,345,1647]" captionText="FIGURES 20–23. Carlschoenherria sulcipennis (Laporte), male from the Philippines, Luzon, Manila. 20, habitus, dorsal as- pect; 21, pronotum, dorsal aspect; 22, genitalia, dorsal aspect; 23, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3405255/files/figure.png" pageId="7" pageNumber="337">Figs. 21</figureCitation>
, 25)........................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51582E6AB432941AFF2AF8CFFA555A97" pageId="7" pageNumber="337">
<paragraph id="EA1336CFB432941AFF2AF8CFFA555AD1" blockId="7.[151,1437,1504,2025]" pageId="7" pageNumber="337">
<keyLead id="515D95FAB432941AFF2AF8CFFA555AD1" pageId="7" pageNumber="337">
5 Abdominal ventrites uniformly covered with pubescence; posterior angle of pronotum rounded, obtuse; prosternal process rectangular-shaped, narrow, elongate, rounded apically (
<figureCitation id="72972A4AB432941AFD40F82CFC8D5AD0" box="[765,836,1944,1966]" captionStart="FIGURES 13–15" captionStartId="7.[151,264,1257,1282]" captionTargetBox="[368,1220,181,1227]" captionTargetId="figure@7.[368,1220,181,1237]" captionTargetPageId="7" captionText="FIGURES 13–15. Abdomen, ventral aspect. 13, Carlschoenherria philippinica (Brenske), male; 14, Carlschoenherria gapudi Calcetas, new species, male; 15, Carlschoenherria sulcipennis (Laporte), male." httpUri="https://zenodo.org/record/3249665/files/figure.png" pageId="7" pageNumber="337">Fig. 15</figureCitation>
).
<collectingCountry id="92BB765FB432941AFCEBF82CFC0C5AD0" box="[854,965,1944,1966]" name="Philippines" pageId="7" pageNumber="337">Philippines</collectingCountry>
,
<collectingCountry id="92BB765FB432941AFC73F82CFBE25AD0" box="[974,1067,1944,1966]" name="Indonesia" pageId="7" pageNumber="337">Indonesia</collectingCountry>
...........
<taxonomicName id="2DAC4D4CB432941AFB09F82DFA555AD1" authority="(Laporte)" baseAuthorityName="Laporte" box="[1204,1436,1945,1967]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="sulcipennis">
<emphasis id="D8D8EADDB432941AFB09F82DFAF55AD0" box="[1204,1340,1945,1966]" italics="true" pageId="7" pageNumber="337">C. sulcipennis</emphasis>
(Laporte)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EA1336CFB432941AFF2AF802FA555A97" blockId="7.[151,1437,1504,2025]" pageId="7" pageNumber="337">
<keyLead id="515D95FAB432941AFF2AF802FA555A97" pageId="7" pageNumber="337">
- Abdominal ventrites not uniformly covered with pubescence (
<figureCitation id="72972A4AB432941AFC85F802FCB45AB2" box="[824,893,1974,1996]" captionStart="FIGURES 13–15" captionStartId="7.[151,264,1257,1282]" captionTargetBox="[368,1220,181,1227]" captionTargetId="figure@7.[368,1220,181,1237]" captionTargetPageId="7" captionText="FIGURES 13–15. Abdomen, ventral aspect. 13, Carlschoenherria philippinica (Brenske), male; 14, Carlschoenherria gapudi Calcetas, new species, male; 15, Carlschoenherria sulcipennis (Laporte), male." httpUri="https://zenodo.org/record/3249665/files/figure.png" pageId="7" pageNumber="337">Fig. 13</figureCitation>
) (
<taxonomicName id="2DAC4D4CB432941AFC2EF802FBEA5AB5" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Brenske" baseAuthorityYear="1894" box="[915,1059,1974,1995]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="337" phylum="Arthropoda" rank="species" species="philippinica">
<emphasis id="D8D8EADDB432941AFC2EF802FBEA5AB5" box="[915,1059,1974,1995]" italics="true" pageId="7" pageNumber="337">C. philippinica</emphasis>
</taxonomicName>
species-group); posterior angle of pro- notum rounded, nearly at 90
<superScript id="1DD99B87B432941AFE4CF866FE315AA0" attach="left" box="[497,504,2002,2014]" fontSize="5" pageId="7" pageNumber="337">0</superScript>
angle; prosternal process spindle-shaped, truncate apically, tapered basally................ 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51582E6AB43D9415FF2AFF2DFA555C3C" pageId="8" pageNumber="338">
<paragraph id="EA1336CFB43D9415FF2AFF2DFA545D94" blockId="8.[151,1437,153,674]" pageId="8" pageNumber="338">
<keyLead id="515D95FAB43D9415FF2AFF2DFA545D94" pageId="8" pageNumber="338">
6 Antennomere III narrowly expanded apically; lamella shorter than anterior margin of clypeus; parameres short; scutellum nar- rowly rounded apically; humeral margin of elytra slightly wedge-shaped towards anterior angle (
<figureCitation id="72972A4AB43D9415FB3EFF03FB305DB3" box="[1155,1273,183,205]" captionStart="FIGURES 24–28" captionStartId="11.[151,264,1654,1679]" captionTargetBox="[267,1223,359,1619]" captionTargetId="figure@11.[343,1230,380,1633]" captionTargetPageId="11" captionText="FIGURES 24–28. Carlschoenherria philippinica (Brenske), male from the Philippines. 24, habitus, dorsal aspect; 25, prono- tum, dorsal aspect; 26, antenna, lateral aspect; 27 genitalia, dorsal aspect; 28, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3249669/files/figure.png" pageId="8" pageNumber="338">Figs. 24–28</figureCitation>
).
<collectingCountry id="92BB765FB43D9415FAB4FF03FABC5DB3" box="[1289,1397,183,205]" name="Philippines" pageId="8" pageNumber="338">Philippines</collectingCountry>
..................................................................................
<taxonomicName id="2DAC4D4CB43D9415FB15FF60FA545D94" authority="(Brenske)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Brenske" baseAuthorityYear="1894" box="[1192,1437,212,234]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="338" phylum="Arthropoda" rank="species" species="philippinica">
<emphasis id="D8D8EADDB43D9415FB15FF60FAF15D97" box="[1192,1336,212,233]" italics="true" pageId="8" pageNumber="338">C. philippinica</emphasis>
(Brenske)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EA1336CFB43D9415FF2AFF46FA555C3C" blockId="8.[151,1437,153,674]" pageId="8" pageNumber="338">
<keyLead id="515D95FAB43D9415FF2AFF46FA555C3C" pageId="8" pageNumber="338">
- Antennomere III widely expanded apically; lamella nearly as long as anterior margin of clypeus; parameres long; scutellum widely rounded apically; humeral margin of elytra distinctly wedge-shaped towards anterior angle (
<figureCitation id="72972A4AB43D9415FB23FEBBFAC15C5B" box="[1182,1288,271,293]" captionStart="FIGURES 9–12" captionStartId="6.[151,264,1970,1995]" captionTargetBox="[273,1273,647,1916]" captionTargetId="figure@6.[376,1212,669,1951]" captionTargetPageId="6" captionText="FIGURES 9–12. Carlschoenherria gapudi Calcetas, new species, holotype, male. 9, habitus, dorsal aspect; 10, antenna, lateral aspect; 11, genitalia, dorsal aspect; 12, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3249663/files/figure.png" pageId="8" pageNumber="338">Figs. 9–12</figureCitation>
).
<collectingCountry id="92BB765FB43D9415FAA5FEBBFA4D5C5B" box="[1304,1412,271,293]" name="Philippines" pageId="8" pageNumber="338">Philippines</collectingCountry>
...........................................................................
<taxonomicName id="2DAC4D4CB43D9415FBDAFE98FAE95C3C" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[1127,1312,300,322]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="338" phylum="Arthropoda" rank="species" species="gapudi" status="sp. nov.">
<emphasis id="D8D8EADDB43D9415FBDAFE98FB0F5C3F" box="[1127,1222,300,321]" italics="true" pageId="8" pageNumber="338">C. gapudi</emphasis>
Calcetas
</taxonomicName>
,
<taxonomicNameLabel id="C3EB57A6B43D9415FA94FE99FA555C3C" box="[1321,1436,300,322]" pageId="8" pageNumber="338" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51582E6AB43D9415FF2AFEFEFA555C8F" pageId="8" pageNumber="338">
<paragraph id="EA1336CFB43D9415FF2AFEFEFA555CC9" blockId="8.[151,1437,153,674]" pageId="8" pageNumber="338">
<keyLead id="515D95FAB43D9415FF2AFEFEFA555CC9" pageId="8" pageNumber="338">
7 Ten elytral costae and ten elytral intervals incomplete, interrupted; elytra rugosely punctate, with irregular patch of setae; sides of abdominal ventrites with large patch of setae on abdominal ventrites II and V and with small patch of setae on abdominal ventrites I, III and IV; clypeus with anterior margin distinctly convex dorsally (
<figureCitation id="72972A4AB43D9415FC64FE30FB865CE4" box="[985,1103,388,410]" captionStart="FIGURES 29–31" captionStartId="12.[151,264,1726,1751]" captionTargetBox="[340,1247,380,1648]" captionTargetId="figure@12.[340,1247,380,1706]" captionText="FIGURES 29–31. Carlschoenherria argus (Burmeister), male from Indonesia, Java. 29, habitus, dorsal aspect; 30, genitalia, dorsal aspect; 31, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3405257/files/figure.png" pageId="8" pageNumber="338">Figs. 29–31</figureCitation>
). Malay Peninsula,
<collectingRegion id="2868F82DB43D9415FAB6FE30FA945CE7" box="[1291,1373,388,409]" country="Indonesia" name="Sumatera Utara" pageId="8" pageNumber="338">Sumatra</collectingRegion>
,
<collectingRegion id="2868F82DB43D9415FADAFE31FA5B5CE4" box="[1383,1426,389,410]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="338">Java</collectingRegion>
...................................................................................
<taxonomicName id="2DAC4D4CB43D9415FB77FE16FA555CC9" authority="(Burmeister)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Burmeister" baseAuthorityYear="1855" box="[1226,1436,417,439]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="338" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="D8D8EADDB43D9415FB77FE16FAD55CC9" box="[1226,1308,418,439]" italics="true" pageId="8" pageNumber="338">C. argus</emphasis>
(Burmeister)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EA1336CFB43D9415FF2AFE0BFA555C8F" blockId="8.[151,1437,153,674]" pageId="8" pageNumber="338">
<keyLead id="515D95FAB43D9415FF2AFE0BFA555C8F" pageId="8" pageNumber="338">
- Ten elytral costae and ten elytral intervals rows complete, uninterrupted; elytra not rugosely punctate, without patch of setae (
<figureCitation id="72972A4AB43D9415FF54FE68FEF05C8F" box="[233,313,476,498]" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1673,1698]" captionTargetBox="[342,1247,417,1630]" captionTargetId="figure@13.[340,1247,417,1653]" captionTargetPageId="13" captionText="FIGURES 32–34. Carlschoenherria hastata (Arrow), male from Malaysia, Perak, Larut Hills. 32, habitus, dorsal aspect; 33, genitalia, dorsal aspect; 34, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3405259/files/figure.png" pageId="8" pageNumber="338">Figs. 32</figureCitation>
, 35); clypeus with anterior margin not distinctly convex dorsally........................................ 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51582E6AB43D9415FF2AFE4EFA545FDC" pageId="8" pageNumber="338">
<paragraph id="EA1336CFB43D9415FF2AFE4EFA545F34" blockId="8.[151,1437,153,674]" pageId="8" pageNumber="338">
<keyLead id="515D95FAB43D9415FF2AFE4EFA545F34" pageId="8" pageNumber="338">
8 Metaventral process distinctly elongate; pygidium with apical margin finely serrate; sides of abdominal sternites without patch of hairs; apical margin of elytra nearly straight dorsally; clypeus with anterior margin nearly straight dorsally (
<figureCitation id="72972A4AB43D9415FAA6FDA3FA5B5F53" box="[1307,1426,535,557]" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1673,1698]" captionTargetBox="[342,1247,417,1630]" captionTargetId="figure@13.[340,1247,417,1653]" captionTargetPageId="13" captionText="FIGURES 32–34. Carlschoenherria hastata (Arrow), male from Malaysia, Perak, Larut Hills. 32, habitus, dorsal aspect; 33, genitalia, dorsal aspect; 34, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3405259/files/figure.png" pageId="8" pageNumber="338">Figs. 32–34</figureCitation>
). Malay Peninsula........................................................................
<taxonomicName id="2DAC4D4CB43D9415FB58FD80FA545F34" authority="(Arrow)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Arrow" baseAuthorityYear="1938" box="[1253,1437,564,586]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="338" phylum="Arthropoda" rank="species" species="hastata">
<emphasis id="D8D8EADDB43D9415FB58FD80FA815F37" box="[1253,1352,564,585]" italics="true" pageId="8" pageNumber="338">C. hastata</emphasis>
(Arrow)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EA1336CFB43D9415FF2AFDE6FA545FDC" blockId="8.[151,1437,153,674]" pageId="8" pageNumber="338">
<keyLead id="515D95FAB43D9415FF2AFDE6FA545FDC" pageId="8" pageNumber="338">
- Metaventral process short; pygidium with apical margin without serration; sides of abdominal sternites with patch of hairs on abdominal sternites II-IV; apical margin of elytra widely rounded dorsally; clypeus with anterior margin slightly convex dor- sally (
<figureCitation id="72972A4AB43D9415FEA1FD38FEAD5FDC" box="[284,356,652,674]" captionStart="FIGURE 35" captionStartId="14.[151,250,1983,2008]" captionTargetBox="[340,1247,181,1947]" captionTargetId="figure@14.[340,1247,181,1963]" captionTargetPageId="14" captionText="FIGURE 35. Carlschoenherria vervex (Sharp), female from Malaysia, Penang. 35, habitus, dorsal aspect." httpUri="https://zenodo.org/record/3249671/files/figure.png" pageId="8" pageNumber="338">Fig. 35</figureCitation>
). Malay Peninsula.............................................................
<taxonomicName id="2DAC4D4CB43D9415FB49FD38FA545FDC" authority="(Sharp)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Sharp" baseAuthorityYear="1876" box="[1268,1437,652,674]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="338" phylum="Arthropoda" rank="species" species="vervex">
<emphasis id="D8D8EADDB43D9415FB49FD38FA865FDC" box="[1268,1359,652,674]" italics="true" pageId="8" pageNumber="338">C. vervex</emphasis>
(Sharp)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

182
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<heading id="B15B81A3B434941CFF2AFA52FD3D5B7F" bold="true" box="[151,756,1510,1537]" fontSize="11" level="1" pageId="1" pageNumber="331" reason="1">
<emphasis id="D8D8EADDB434941CFF2AFA52FD3D5B7F" bold="true" box="[151,756,1510,1537]" pageId="1" pageNumber="331">
<taxonomicName id="2DAC4D4CB434941CFF2AFA52FD915B7F" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[151,600,1510,1537]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="331" phylum="Arthropoda" rank="species" species="adoradae" status="sp. nov.">
<emphasis id="D8D8EADDB434941CFF2AFA52FE2D5B7F" bold="true" box="[151,484,1510,1537]" italics="true" pageId="1" pageNumber="331">Carlschoenherria adoradae</emphasis>
Calcetas
</taxonomicName>
,
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</emphasis>
</heading>
</paragraph>
<paragraph id="EA1336CFB434941CFF2AF9BFFEF55B58" blockId="1.[151,756,1510,1574]" box="[151,316,1547,1574]" pageId="1" pageNumber="331">(Figs. 1–4, 36)</paragraph>
</subSubSection>
<subSubSection id="A2B66544B434941CFF2AF9E7FAB15BEF" pageId="1" pageNumber="331" type="materials_examined">
<paragraph id="EA1336CFB434941CFF2AF9E7FC915B13" blockId="1.[151,1436,1618,2041]" box="[151,856,1618,1645]" pageId="1" pageNumber="331">
<materialsCitation id="5AC43C92B434941CFF2AF9E7FC9D5B13" ID-GBIF-Occurrence="2268317642" box="[151,852,1618,1645]" country="Philippines" location="Luzon Island" municipality="The" pageId="1" pageNumber="331" specimenCount="1" stateProvince="Kalinga Apayao" typeStatus="holotype">
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<typeStatus id="3517886DB434941CFF2AF9E7FF1B5B13" box="[151,210,1619,1645]" pageId="1" pageNumber="331">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="0A77ACB5B434941CFE81F9E7FEA15B13" box="[316,360,1619,1645]" pageId="1" pageNumber="331">The</collectingMunicipality>
<collectingCountry id="92BB765FB434941CFED3F9E7FE265B13" box="[366,495,1619,1645]" name="Philippines" pageId="1" pageNumber="331">Philippines</collectingCountry>
,
<location id="EF736014B434941CFE47F9E7FD595B13" LSID="urn:lsid:plazi:treatment:620587D9B434941EFF2AFA52FEA85E69:EF736014B434941CFE47F9E7FD595B13" box="[506,656,1619,1645]" country="Philippines" municipality="The" name="Luzon Island" pageId="1" pageNumber="331" stateProvince="Kalinga Apayao">Luzon Island</location>
,
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<collectingRegion id="2868F82DB434941CFD26F9E7FD3D5B13" box="[667,756,1619,1645]" country="Philippines" name="Kalinga" pageId="1" pageNumber="331">Kalinga</collectingRegion>
<collectingRegion id="2868F82DB434941CFD44F9E7FC9D5B13" box="[761,852,1619,1645]" country="Philippines" name="Apayao" pageId="1" pageNumber="331">Apayao</collectingRegion>
</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="EA1336CFB434941CFF7AF9C3FAB15BEF" blockId="1.[151,1436,1618,2041]" box="[199,1400,1654,1682]" pageId="1" pageNumber="331">
<emphasis id="D8D8EADDB434941CFF7AF9C3FEBC5BEF" bold="true" box="[199,373,1654,1681]" pageId="1" pageNumber="331">Type material.</emphasis>
<materialsCitation id="5AC43C92B434941CFEC1F9C3FAB15BEF" ID-GBIF-Occurrence="2268317644" box="[380,1400,1654,1682]" collectingDate="1952-05-15" collectionCode="MNH" collectorName="S. M. Cendana" elevation="91" location="Bayog" pageId="1" pageNumber="331" specimenCount="1" stateProvince="Apayao" typeStatus="holotype">
<typeStatus id="3517886DB434941CFEC1F9C3FE2D5BEF" box="[380,484,1655,1681]" pageId="1" pageNumber="331" type="holotype">Holotype</typeStatus>
(male), labeled:
<collectingRegion id="2868F82DB434941CFD1DF9C3FD335BEF" box="[672,762,1655,1681]" country="Philippines" name="Apayao" pageId="1" pageNumber="331">Apayao</collectingRegion>
,
<location id="EF736014B434941CFCB8F9C3FC985BEF" LSID="urn:lsid:plazi:treatment:620587D9B434941EFF2AFA52FEA85E69:EF736014B434941CFCB8F9C3FC985BEF" box="[773,849,1655,1681]" name="Bayog" pageId="1" pageNumber="331" stateProvince="Apayao">Bayog</location>
, Elev.
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<quantity id="2D549B2AB434941CFC21F9C3FC2E5BEF" box="[924,999,1655,1681]" metricMagnitude="1" metricUnit="m" metricValue="9.144" pageId="1" pageNumber="331" unit="ft" value="300.0">300 ft.</quantity>
</elevation>
,
<collectingDate id="8E56E9E7B434941CFC4EF9C2FBA35BEF" box="[1011,1130,1654,1681]" pageId="1" pageNumber="331" value="1952-05-15">
<date id="9E12100FB434941CFC4EF9C2FBA35BEF" box="[1011,1130,1654,1681]" pageId="1" pageNumber="331" value="1952-05-15">15-v-1952</date>
</collectingDate>
,
<collectorName id="47595319B434941CFBCBF9C3FADC5BEF" box="[1142,1301,1655,1681]" pageId="1" pageNumber="331">S.M. Cendana</collectorName>
(
<collectionCode id="8CBDAE0AB434941CFA99F9C3FAA55BEF" box="[1316,1388,1655,1681]" country="Hungary" httpUri="http://grbio.org/cool/34nf-ps25" name="Musei Nacionalis Hungarici" pageId="1" pageNumber="331">MNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B434941EFF7AF92EFD375CAD" lastPageId="3" lastPageNumber="333" pageId="1" pageNumber="331" type="description">
<paragraph id="EA1336CFB434941CFF7AF92EFAAB5B83" blockId="1.[151,1436,1618,2041]" pageId="1" pageNumber="331">
<emphasis id="D8D8EADDB434941CFF7AF92EFDE15BCB" bold="true" box="[199,552,1690,1717]" pageId="1" pageNumber="331">
Description of
<typeStatus id="3517886DB434941CFEC5F92FFE295BCB" box="[376,480,1691,1717]" pageId="1" pageNumber="331" type="holotype">holotype</typeStatus>
, male
</emphasis>
. Length 20.5 mm, width 8.5 mm. Body dichromatic, head, pronotum and scu- tellum dark reddish brown; uniformly covered with short, brownish to whitish setae; elytron light reddish brown, covered with short brownish to yellowish setae, with few short whitish acicular-like scales; leg black (Fig. 1).
</paragraph>
<paragraph id="EA1336CFB434941CFF7AF8B3FCD05A87" blockId="1.[151,1436,1618,2041]" pageId="1" pageNumber="331">
<emphasis id="D8D8EADDB434941CFF7AF8B3FEC25A5F" box="[199,267,1799,1825]" italics="true" pageId="1" pageNumber="331">Head.</emphasis>
Mesal carina starting at junction of fronto-clypeal suture, terminating far behind vertex. Anterior margin of clypeus straight dorsally; surface subrugosely punctate anteriorly; with very narrow, smooth margin, impunctate, non-setose area adjacent to anterior margin; dorsal surface of clypeus sloping, deeply depressed adjacent to anterior margin. Antennomere III with short stalk, widely expanded apically. Lamellae length 2.3 mm. Terminal maxillary palpomere with shallow nearly flattened lanceolate shaped groove or depression, as long as antennomere III. Men- tum with distinct roof-like mesal carina, slanted on each side, with deep depression below it; surface subrugosely punctate, covered with few setae and acicular-like scales.
</paragraph>
<paragraph id="EA1336CFB436941EFF7AFF2DFA455D85" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="333">
<emphasis id="D8D8EADDB436941EFF7AFF2DFE055DCD" box="[199,460,153,179]" italics="true" pageId="3" pageNumber="333">Ventral side of thorax.</emphasis>
Prosternal process rectangulate, elongate, slender or narrow longitudinally, truncate apically. Prosternum and mesoventrum densely covered with long yellowish hairs. Metaventral process little prolonged, not reaching procoxae, sub-apically attached to mesoventral process; metaventral process length 1.5 mm.
</paragraph>
<paragraph id="EA1336CFB436941EFF7AFEB1FB725C19" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="333">
<emphasis id="D8D8EADDB436941EFF7AFEB1FE655C61" box="[199,428,261,287]" italics="true" pageId="3" pageNumber="333">Abdominal ventrites</emphasis>
not uniformly covered with setation, bare at least near each margin or rather narrowly restricted in distribution of setation; posterior part of abdominal ventrites II–IV narrow, smooth, bare; sides of II–V with large and small ovoid shaped patches of yellowish-white scales or abdominal maculation.
</paragraph>
<paragraph id="EA1336CFB436941EFF7AFEC5FC1D5CF5" blockId="3.[151,1437,153,791]" box="[199,980,369,395]" pageId="3" pageNumber="333">
<emphasis id="D8D8EADDB436941EFF7AFEC5FEFB5CF5" box="[199,306,369,395]" italics="true" pageId="3" pageNumber="333">Pygidium</emphasis>
with anterior margin nearly straight; posterior margin wide.
</paragraph>
<paragraph id="EA1336CFB436941EFF7AFE21FD375CAD" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="333">
<emphasis id="D8D8EADDB436941EFF7AFE21FEFD5CD1" box="[199,308,405,431]" italics="true" pageId="3" pageNumber="333">Aedeagus</emphasis>
. Posterior margin of parameres ribbon-like, longitudinally undulated towards apex. Length of param- eres and phallobase combined 6.4 mm; as in Figs. 3, 4.
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B436941EFF7AFE68FD495C89" box="[199,640,476,503]" pageId="3" pageNumber="333" type="distribution">
<paragraph id="EA1336CFB436941EFF7AFE68FD495C89" blockId="3.[151,1437,153,791]" box="[199,640,476,503]" pageId="3" pageNumber="333">
<emphasis id="D8D8EADDB436941EFF7AFE68FEA85C89" bold="true" box="[199,353,476,503]" pageId="3" pageNumber="333">Distribution.</emphasis>
The
<collectingCountry id="92BB765FB436941EFE24FE69FDDE5C89" box="[409,535,477,503]" name="Philippines" pageId="3" pageNumber="333">Philippines</collectingCountry>
(Luzon).
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B436941EFF7AFDB5FB005F41" pageId="3" pageNumber="333" type="etymology">
<paragraph id="EA1336CFB436941EFF7AFDB5FB005F41" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="333">
<emphasis id="D8D8EADDB436941EFF7AFDB5FE875F65" bold="true" box="[199,334,513,539]" pageId="3" pageNumber="333">Etymology.</emphasis>
This new species is named after Dr. Jessamyn R. Adorada, professor of taxonomy and systematics at the University of the
<collectingCountry id="92BB765FB436941EFE1DFD91FDE85F41" box="[416,545,549,575]" name="Philippines" pageId="3" pageNumber="333">Philippines</collectingCountry>
, Los Baños. She is also an expert on Philippine coccinellids.
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B436941EFF7AFDFCFEA85E69" pageId="3" pageNumber="333" type="diagnosis">
<paragraph id="EA1336CFB436941EFF7AFDFCFEA85E69" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="333">
<emphasis id="D8D8EADDB436941EFF7AFDFCFE195F1D" bold="true" box="[199,464,584,611]" pageId="3" pageNumber="333">Differential diagnosis.</emphasis>
Based on the species-group classification of
<bibRefCitation id="8E3D4B3EB436941EFC61FDFDFB965F1D" author="Itoh, T." box="[988,1119,585,611]" pageId="3" pageNumber="333" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh (1993)</bibRefCitation>
, this species belongs to the
<taxonomicName id="2DAC4D4CB436941EFF2AFDD9FEF05FF9" authorityName="Laporte" authorityYear="1840" box="[151,313,621,647]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="sulcipennis">
<emphasis id="D8D8EADDB436941EFF2AFDD9FEF05FF9" box="[151,313,621,647]" italics="true" pageId="3" pageNumber="333">C. sulcipennis</emphasis>
</taxonomicName>
group wherein the parameres are laterally compressed and bird-bill shaped at apex. However, in contrast to other members of
<taxonomicName id="2DAC4D4CB436941EFE58FD25FD4C5FD5" authorityName="Laporte" authorityYear="1840" box="[485,645,657,683]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="sulcipennis">
<emphasis id="D8D8EADDB436941EFE58FD25FD4C5FD5" box="[485,645,657,683]" italics="true" pageId="3" pageNumber="333">C. sulcipennis</emphasis>
</taxonomicName>
group, the anterior margin of the parameres of
<taxonomicName id="2DAC4D4CB436941EFB1CFD25FA515FD5" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[1185,1432,657,683]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="adoradae" status="sp. nov.">
<emphasis id="D8D8EADDB436941EFB1CFD25FAE75FD5" box="[1185,1326,657,683]" italics="true" pageId="3" pageNumber="333">C. adoradae</emphasis>
Calcetas
</taxonomicName>
,
<taxonomicNameLabel id="C3EB57A6B436941EFF2AFD01FED45FB1" box="[151,285,693,719]" pageId="3" pageNumber="333" rank="species">new species</taxonomicNameLabel>
and
<taxonomicName id="2DAC4D4CB436941EFEEEFD01FD815FB1" authority="Calcetas" authorityName="Calcetas" authorityYear="2019" box="[339,584,693,719]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="hadsallae" status="sp. nov.">
<emphasis id="D8D8EADDB436941EFEEEFD01FE295FB1" box="[339,480,693,719]" italics="true" pageId="3" pageNumber="333">C. hadsallae</emphasis>
Calcetas
</taxonomicName>
,
<taxonomicNameLabel id="C3EB57A6B436941EFDEEFD01FD105FB1" box="[595,729,693,719]" pageId="3" pageNumber="333" rank="species">new species</taxonomicNameLabel>
are wide and laterally expanded while it’s laterally compressed and bird-bill shaped at apex for all other species. Thus, they are classified under the newly proposed
<taxonomicName id="2DAC4D4CB436941EFAB2FD6DFA555F8D" authorityName="Calcetas" authorityYear="2019" box="[1295,1436,729,755]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="333" phylum="Arthropoda" rank="species" species="adoradae">
<emphasis id="D8D8EADDB436941EFAB2FD6DFA555F8D" box="[1295,1436,729,755]" italics="true" pageId="3" pageNumber="333">C. adoradae</emphasis>
</taxonomicName>
species-subgroup.
</paragraph>
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</treatment>
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<treatment id="620587D9B4389412FF2AF8A3FDE95CF5" ID-DOI="http://doi.org/10.5281/zenodo.5930694" ID-GBIF-Taxon="158297465" ID-Zenodo-Dep="5930694" LSID="urn:lsid:plazi:treatment:620587D9B4389412FF2AF8A3FDE95CF5" httpUri="http://treatment.plazi.org/id/620587D9B4389412FF2AF8A3FDE95CF5" lastPageId="15" lastPageNumber="345" pageId="13" pageNumber="343">
<subSubSection id="A2B66544B4389410FF2AF8A3FC995A4C" box="[151,848,1815,1842]" pageId="13" pageNumber="343" type="nomenclature">
<paragraph id="EA1336CFB4389410FF2AF8A3FC995A4C" blockId="13.[151,848,1815,1878]" box="[151,848,1815,1842]" pageId="13" pageNumber="343">
<heading id="B15B81A3B4389410FF2AF8A3FC995A4C" bold="true" box="[151,848,1815,1842]" fontSize="11" level="1" pageId="13" pageNumber="343" reason="1">
<emphasis id="D8D8EADDB4389410FF2AF8A3FC995A4C" bold="true" box="[151,848,1815,1842]" pageId="13" pageNumber="343">
<taxonomicName id="2DAC4D4CB4389410FF2AF8A3FDBA5A4F" authority="(Sharp, 1876)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Sharp" baseAuthorityYear="1876" box="[151,627,1815,1842]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="343" phylum="Arthropoda" rank="species" species="vervex" status="comb. nov.">
<emphasis id="D8D8EADDB4389410FF2AF8A3FE0B5A4C" bold="true" box="[151,450,1815,1842]" italics="true" pageId="13" pageNumber="343">Carlschoenherria vervex</emphasis>
(
<bibRefCitation id="8E3D4B3EB4389410FE6FF8A3FDA25A4F" author="Sharp, D." box="[466,619,1815,1841]" pageId="13" pageNumber="343" pagination="65 - 90" refId="ref7870" refString="Sharp, D. (1876) Description of some new species of Scarabaeidae from tropical Asia and Malaysia. Part III (Melolonthini). Coleopterologische Hefte, 15, 65 - 90." type="journal article" year="1876">Sharp, 1876</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C3EB57A6B4389410FDC7F8ACFC995A4C" box="[634,848,1816,1842]" pageId="13" pageNumber="343" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B4389410FF2AF888FF305A28" box="[151,249,1851,1878]" pageId="13" pageNumber="343" type="description">
<paragraph id="EA1336CFB4389410FF2AF888FF305A28" blockId="13.[151,848,1815,1878]" box="[151,249,1851,1878]" pageId="13" pageNumber="343">
(
<figureCitation id="72972A4AB4389410FF1DF888FF385A28" box="[160,241,1851,1878]" captionStart="FIGURE 35" captionStartId="14.[151,250,1983,2008]" captionTargetBox="[340,1247,181,1947]" captionTargetId="figure@14.[340,1247,181,1963]" captionTargetPageId="14" captionText="FIGURE 35. Carlschoenherria vervex (Sharp), female from Malaysia, Penang. 35, habitus, dorsal aspect." httpUri="https://zenodo.org/record/3249671/files/figure.png" pageId="13" pageNumber="343">Fig. 35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B4389410FF2AF834FB475AC9" pageId="13" pageNumber="343" type="reference_group">
<paragraph id="EA1336CFB4389410FF2AF834FCCF5AE6" blockId="13.[151,1166,1919,1975]" box="[151,774,1919,1944]" pageId="13" pageNumber="343">
<taxonomicName id="2DAC4D4CB4389410FF2AF834FDEB5AE6" authority="Sharp, 1876: 89 - 90" authorityName="Sharp" authorityPageNumber="89" authorityYear="1876" box="[151,546,1919,1944]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="343" phylum="Arthropoda" rank="species" species="vervex">
<emphasis id="D8D8EADDB4389410FF2AF834FE9D5AE6" box="[151,340,1920,1944]" italics="true" pageId="13" pageNumber="343">Melolontha vervex</emphasis>
<bibRefCitation id="8E3D4B3EB4389410FEE6F8CBFDEB5AE6" author="Sharp, D." box="[347,546,1919,1944]" pageId="13" pageNumber="343" pagination="65 - 90" refId="ref7870" refString="Sharp, D. (1876) Description of some new species of Scarabaeidae from tropical Asia and Malaysia. Part III (Melolonthini). Coleopterologische Hefte, 15, 65 - 90." type="journal article" year="1876">Sharp, 1876: 89–90</bibRefCitation>
</taxonomicName>
(original description).
</paragraph>
<paragraph id="EA1336CFB4389410FF2AF82BFB475AC9" blockId="13.[151,1166,1919,1975]" box="[151,1166,1950,1975]" pageId="13" pageNumber="343">
<treatmentCitationGroup id="CABC11E1B4389410FF2AF82BFB475AC9" box="[151,1166,1950,1975]" pageId="13" pageNumber="343">
<taxonomicName id="2DAC4D4CB4389410FF2AF82BFE375AC9" authority=": Itoh 1992: 10" authorityName="Itoh" authorityPageNumber="10" authorityYear="1992" box="[151,510,1950,1975]" class="Insecta" family="Curculionidae" genus="Schoenherria" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="343" phylum="Arthropoda" rank="species" species="vervex">
<emphasis id="D8D8EADDB4389410FF2AF82BFEA15AC8" box="[151,360,1951,1975]" italics="true" pageId="13" pageNumber="343">Schoenherria vervex</emphasis>
:
<treatmentCitation id="6B0D10DEB4389410FEC8F82AFE375AC9" author="Itoh, T." box="[373,510,1950,1975]" page="10" pageId="13" pageNumber="343" year="1992">
<bibRefCitation id="8E3D4B3EB4389410FEC8F82AFE375AC9" author="Itoh, T." box="[373,510,1950,1975]" pageId="13" pageNumber="343" pagination="7 - 14" refId="ref7594" refString="Itoh, T. (1992) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) I. Lamellicornia, 7, 7 - 14." type="journal article" year="1992">Itoh 1992: 10</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(new generic placement);
<bibRefCitation id="8E3D4B3EB4389410FCB1F82AFC725AC9" author="Itoh, T." box="[780,955,1950,1975]" pageId="13" pageNumber="343" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh 1993: 13–14</bibRefCitation>
(redescription, key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="BED36647B43B9413FF2AF80BFB055AA9" box="[151,1228,1983,2008]" httpUri="https://zenodo.org/record/3249671/files/figure.png" pageId="14" pageNumber="344" startId="14.[151,250,1983,2008]" targetBox="[340,1247,181,1947]" targetPageId="14">
<paragraph id="EA1336CFB43B9413FF2AF80BFB055AA9" blockId="14.[151,1228,1983,2008]" box="[151,1228,1983,2008]" pageId="14" pageNumber="344">
<emphasis id="D8D8EADDB43B9413FF2AF80BFEE95AA6" bold="true" box="[151,288,1983,2008]" pageId="14" pageNumber="344">FIGURE 35.</emphasis>
<taxonomicName id="2DAC4D4CB43B9413FE9BF874FDBE5AA6" authority="(Sharp)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Sharp" baseAuthorityYear="1876" box="[294,631,1983,2008]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="344" phylum="Arthropoda" rank="species" species="vervex">
<emphasis id="D8D8EADDB43B9413FE9BF874FDEA5AA6" box="[294,547,1984,2008]" italics="true" pageId="14" pageNumber="344">Carlschoenherria vervex</emphasis>
(Sharp)
</taxonomicName>
, female from Malaysia, Penang. 35, habitus, dorsal aspect.
</paragraph>
</caption>
<subSubSection id="A2B66544B43A9412FF2AFF2DFEBE5C61" pageId="15" pageNumber="345" type="materials_examined">
<paragraph id="EA1336CFB43A9412FF2AFF2DFDA65DCC" blockId="15.[151,1437,152,395]" box="[151,623,152,179]" pageId="15" pageNumber="345">
<materialsCitation id="5AC43C92B43A9412FF2AFF2DFDA25DCC" ID-GBIF-Occurrence="2268317696" box="[151,619,152,179]" country="Malaysia" location="Island" pageId="15" pageNumber="345" specimenCount="1" stateProvince="Pulau Pinang" typeStatus="holotype">
<emphasis id="D8D8EADDB43A9412FF2AFF2DFEFF5DCD" bold="true" box="[151,310,152,179]" pageId="15" pageNumber="345">
<typeStatus id="3517886DB43A9412FF2AFF2DFF1B5DCD" box="[151,210,153,179]" pageId="15" pageNumber="345">Type</typeStatus>
locality.
</emphasis>
“
<collectingCountry id="92BB765FB43A9412FEFBFF2CFE7B5DCC" box="[326,434,152,178]" name="Malaysia" pageId="15" pageNumber="345">Malaysia</collectingCountry>
,
<collectingRegion id="2868F82DB43A9412FE00FF2DFDC65DCD" box="[445,527,153,179]" country="Malaysia" name="Pulau Pinang" pageId="15" pageNumber="345">Penang</collectingRegion>
<location id="EF736014B43A9412FDAAFF2CFDA85DCC" LSID="urn:lsid:plazi:treatment:620587D9B4389412FF2AF8A3FDE95CF5:EF736014B43A9412FDAAFF2CFDA85DCC" box="[535,609,152,178]" country="Malaysia" name="Island" pageId="15" pageNumber="345" stateProvince="Pulau Pinang">Island</location>
”
</materialsCitation>
.
</paragraph>
<paragraph id="EA1336CFB43A9412FF7AFF09FDF35DA9" blockId="15.[151,1437,152,395]" box="[199,570,188,215]" pageId="15" pageNumber="345">
<emphasis id="D8D8EADDB43A9412FF7AFF09FE265DA9" bold="true" box="[199,495,188,215]" pageId="15" pageNumber="345">
<typeStatus id="3517886DB43A9412FF7AFF09FECB5DA9" box="[199,258,189,215]" pageId="15" pageNumber="345">Type</typeStatus>
material examined.
</emphasis>
None.
</paragraph>
<paragraph id="EA1336CFB43A9412FF7AFF54FEBE5C61" blockId="15.[151,1437,152,395]" pageId="15" pageNumber="345">
<emphasis id="D8D8EADDB43A9412FF7AFF54FDFA5D85" bold="true" box="[199,563,224,251]" pageId="15" pageNumber="345">Additional material examined.</emphasis>
<materialsCitation id="5AC43C92B43A9412FD84FF54FEBA5C61" ID-GBIF-Occurrence="2268317689" collectingDate="1998-02-16" collectionCode="OCPC" collectorName="Bukit Larut, E &amp; Cempirek." country="Malaysia" location="MALAYSIA" pageId="15" pageNumber="345" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="D8D8EADDB43A9412FD84FF54FD115D85" bold="true" box="[569,728,224,251]" pageId="15" pageNumber="345">
<collectingCountry id="92BB765FB43A9412FD84FF54FD1A5D85" box="[569,723,224,251]" name="Malaysia" pageId="15" pageNumber="345">MALAYSIA</collectingCountry>
:
</emphasis>
<specimenCount id="FCAAFD46B43A9412FD63FF55FCC85D85" box="[734,769,225,251]" pageId="15" pageNumber="345" type="female">1♀</specimenCount>
,
<collectorName id="47595319B43A9412FCB2FF54FC785D85" box="[783,945,224,251]" pageId="15" pageNumber="345">Bukit Larut, E</collectorName>
. Taiping,
<collectingDate id="8E56E9E7B43A9412FBA0FF54FB5F5D84" box="[1053,1174,224,250]" pageId="15" pageNumber="345" value="1998-02-16">
<date id="9E12100FB43A9412FBA0FF54FB5F5D84" box="[1053,1174,224,250]" pageId="15" pageNumber="345" value="1998-02-16">16-ii-1998</date>
</collectingDate>
, 1800 ASL,
<quantity id="2D549B2AB43A9412FA9DFF54FAA05D84" box="[1312,1385,224,251]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="345" unit="km" value="15.0">15 km</quantity>
, leg.
<collectorName id="47595319B43A9412FF2AFEB0FEC45C60" box="[151,269,260,286]" pageId="15" pageNumber="345">Cempírek.</collectorName>
(
<collectionCode id="8CBDAE0AB43A9412FEA1FEB1FEA55C61" box="[284,364,261,287]" pageId="15" pageNumber="345">OCPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43A9412FF7AFE9CFDE95CF5" pageId="15" pageNumber="345" type="distribution">
<paragraph id="EA1336CFB43A9412FF7AFE9CFDE95CF5" blockId="15.[151,1437,152,395]" pageId="15" pageNumber="345">
<emphasis id="D8D8EADDB43A9412FF7AFE9CFE905C3D" bold="true" box="[199,345,296,323]" pageId="15" pageNumber="345">Distribution</emphasis>
Originally described by
<bibRefCitation id="8E3D4B3EB43A9412FDD6FE9CFD345C3D" author="Sharp, D." box="[619,765,296,323]" pageId="15" pageNumber="345" pagination="65 - 90" refId="ref7870" refString="Sharp, D. (1876) Description of some new species of Scarabaeidae from tropical Asia and Malaysia. Part III (Melolonthini). Coleopterologische Hefte, 15, 65 - 90." type="journal article" year="1876">Sharp (1876)</bibRefCitation>
from
<collectingRegion id="2868F82DB43A9412FCFDFE9DFC5B5C3D" box="[832,914,297,323]" country="Malaysia" name="Pulau Pinang" pageId="15" pageNumber="345">Penang</collectingRegion>
Island (West of
<collectingCountry id="92BB765FB43A9412FBFBFE9CFB675C3D" box="[1094,1198,296,323]" name="Malaysia" pageId="15" pageNumber="345">Malaysia</collectingCountry>
) and reported by
<bibRefCitation id="8E3D4B3EB43A9412FAD2FE9CFF2B5C19" author="Itoh, T." pageId="15" pageNumber="345" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh (1993)</bibRefCitation>
from Tapah and
<collectingRegion id="2868F82DB43A9412FE19FEF8FE245C18" box="[420,493,332,358]" country="Malaysia" name="Kedah" pageId="15" pageNumber="345">Kedah</collectingRegion>
(Western
<collectingCountry id="92BB765FB43A9412FDDFFEF8FD035C19" box="[610,714,332,359]" name="Malaysia" pageId="15" pageNumber="345">Malaysia</collectingCountry>
) and
<collectingRegion id="2868F82DB43A9412FCB7FEF8FC955C18" box="[778,860,332,358]" country="Malaysia" name="Pahang" pageId="15" pageNumber="345">Pahang</collectingRegion>
(Central
<collectingCountry id="92BB765FB43A9412FC75FEF8FBFD5C19" box="[968,1076,332,359]" name="Malaysia" pageId="15" pageNumber="345">Malaysia</collectingCountry>
). The species is newly recorded in Bukit Larut (Western
<collectingCountry id="92BB765FB43A9412FE15FEC4FDDD5CF5" box="[424,532,368,395]" name="Malaysia" pageId="15" pageNumber="345">Malaysia</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>

110
data/62/05/87/620587D9B4399410FF2AF8F3FD175CF2.xml Normal file
View file

@ -0,0 +1,110 @@
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<paragraph id="EA1336CFB4399411FF2AF8F3FC975A1F" blockId="12.[151,862,1863,1926]" box="[151,862,1863,1890]" pageId="12" pageNumber="342">
<heading id="B15B81A3B4399411FF2AF8F3FC975A1F" bold="true" box="[151,862,1863,1890]" fontSize="11" level="1" pageId="12" pageNumber="342" reason="1">
<emphasis id="D8D8EADDB4399411FF2AF8F3FC975A1F" bold="true" box="[151,862,1863,1890]" pageId="12" pageNumber="342">
<taxonomicName id="2DAC4D4CB4399411FF2AF8F3FD485A1F" authority="(Arrow, 1938)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Arrow" baseAuthorityYear="1938" box="[151,641,1863,1890]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="342" phylum="Arthropoda" rank="species" species="hastata" status="comb. nov.">
<emphasis id="D8D8EADDB4399411FF2AF8F3FE055A1C" bold="true" box="[151,460,1863,1890]" italics="true" pageId="12" pageNumber="342">Carlschoenherria hastata</emphasis>
(
<bibRefCitation id="8E3D4B3EB4399411FE66F8F3FDB05A1F" author="Arrow, G. J." box="[475,633,1863,1889]" pageId="12" pageNumber="342" pagination="267 - 278" refId="ref7182" refString="Arrow, G. J. (1938) Notes on some Melolonthine Coleoptera from the Malay Peninsula and descriptions of a few new species. Journal of Federated Malay States Museum, 18, 267 - 278." type="journal article" year="1938">Arrow, 1938</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C3EB57A6B4399411FD34F8FCFC975A1F" box="[649,862,1863,1890]" pageId="12" pageNumber="342" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B4399411FF2AF8DFFEE75AF8" box="[151,302,1899,1926]" pageId="12" pageNumber="342" type="description">
<paragraph id="EA1336CFB4399411FF2AF8DFFEE75AF8" blockId="12.[151,862,1863,1926]" box="[151,302,1899,1926]" pageId="12" pageNumber="342">
(
<figureCitation id="72972A4AB4399411FF22F8DFFEEF5AF8" box="[159,294,1899,1926]" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1673,1698]" captionTargetBox="[342,1247,417,1630]" captionTargetId="figure@13.[340,1247,417,1653]" captionTargetPageId="13" captionText="FIGURES 32–34. Carlschoenherria hastata (Arrow), male from Malaysia, Perak, Larut Hills. 32, habitus, dorsal aspect; 33, genitalia, dorsal aspect; 34, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3405259/files/figure.png" pageId="12" pageNumber="342">Figs. 32–34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B4399411FF2AF81EFD305ABD" box="[151,761,1962,1987]" pageId="12" pageNumber="342" type="reference_group">
<paragraph id="EA1336CFB4399411FF2AF81EFD305ABD" blockId="12.[151,1175,1962,2017]" box="[151,761,1962,1987]" pageId="12" pageNumber="342">
<treatmentCitationGroup id="CABC11E1B4399411FF2AF81EFD305ABD" box="[151,761,1962,1987]" pageId="12" pageNumber="342">
<taxonomicName id="2DAC4D4CB4399411FF2AF81EFDDC5ABC" authority="Arrow, 1938: 278" authorityName="Arrow" authorityPageNumber="278" authorityYear="1938" box="[151,533,1962,1987]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="342" phylum="Arthropoda" rank="species" species="hastata">
<emphasis id="D8D8EADDB4399411FF2AF81EFE945ABC" box="[151,349,1962,1986]" italics="true" pageId="12" pageNumber="342">Melolontha hastata</emphasis>
<treatmentCitation id="6B0D10DEB4399411FEDFF81FFDDC5ABC" author="Arrow, G. J." box="[354,533,1962,1987]" page="278" pageId="12" pageNumber="342" year="1938">
<bibRefCitation id="8E3D4B3EB4399411FEDFF81FFDDC5ABC" author="Arrow, G. J." box="[354,533,1962,1987]" pageId="12" pageNumber="342" pagination="267 - 278" refId="ref7182" refString="Arrow, G. J. (1938) Notes on some Melolonthine Coleoptera from the Malay Peninsula and descriptions of a few new species. Journal of Federated Malay States Museum, 18, 267 - 278." type="journal article" year="1938">Arrow, 1938: 278</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(original description).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<paragraph id="EA1336CFB4399410FF2AF87DFC7B5DA9" blockId="12.[151,1175,1962,2017]" lastBlockId="13.[151,1437,188,396]" lastPageId="13" lastPageNumber="343" pageId="12" pageNumber="342">
<taxonomicName id="2DAC4D4CB4399411FF2AF87DFDCF5A9F" authority=": Itoh 1992: 10" authorityName="Itoh" authorityPageNumber="10" authorityYear="1992" box="[151,518,1992,2017]" class="Insecta" family="Curculionidae" genus="Schoenherria" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="342" phylum="Arthropoda" rank="species" species="hastata">
<emphasis id="D8D8EADDB4399411FF2AF87DFEB85A9F" box="[151,369,1993,2017]" italics="true" pageId="12" pageNumber="342">Schoenherria hastata</emphasis>
:
<treatmentCitation id="6B0D10DEB4399411FEC3F87CFDCF5A9F" author="Itoh, T." box="[382,518,1992,2017]" page="10" pageId="12" pageNumber="342" year="1992">
<bibRefCitation id="8E3D4B3EB4399411FEC3F87CFDCF5A9F" author="Itoh, T." box="[382,518,1992,2017]" pageId="12" pageNumber="342" pagination="7 - 14" refId="ref7594" refString="Itoh, T. (1992) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) I. Lamellicornia, 7, 7 - 14." type="journal article" year="1992">Itoh 1992: 10</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(new generic placement);
<bibRefCitation id="8E3D4B3EB4399411FCA8F87CFC0A5A9F" author="Itoh, T." box="[789,963,1992,2017]" pageId="12" pageNumber="342" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh 1993: 12–13</bibRefCitation>
(redescription, key)
<subSubSection id="A2B66544B4389410FF2AFF09FC7B5DA9" box="[151,946,188,215]" pageId="13" pageNumber="343" type="materials_examined">
<emphasis id="D8D8EADDB4389410FF2AFF09FEFF5DA9" bold="true" box="[151,310,188,215]" pageId="13" pageNumber="343">Type locality.</emphasis>
“Malaysia, Perak, Larut Hills; Pahang, Lubok Tamang”.
</subSubSection>
</paragraph>
<paragraph id="EA1336CFB4399411FB2EF87CFB5E5A9F" blockId="12.[151,1175,1962,2017]" box="[1171,1175,1992,2017]" pageId="12" pageNumber="342">.</paragraph>
<paragraph id="EA1336CFB4389410FF7AFF55FDF35D85" blockId="13.[151,1437,188,396]" box="[199,570,224,251]" pageId="13" pageNumber="343">
<emphasis id="D8D8EADDB4389410FF7AFF55FE265D85" bold="true" box="[199,495,224,251]" pageId="13" pageNumber="343">Type material examined.</emphasis>
None.
</paragraph>
<paragraph id="EA1336CFB4389410FF7AFEB0FEB95C3D" blockId="13.[151,1437,188,396]" pageId="13" pageNumber="343">
<emphasis id="D8D8EADDB4389410FF7AFEB0FD165C61" bold="true" box="[199,735,260,287]" pageId="13" pageNumber="343">Additional material examined. MALAYSIA:</emphasis>
1♂, 1♀, Tanah Rata, Cameron Highlands, 12-20-ii-1998, lgt. Cempírek (OCPC).
</paragraph>
<subSubSection id="A2B66544B4389410FF7AFEF8FD175CF2" pageId="13" pageNumber="343" type="distribution">
<paragraph id="EA1336CFB4389410FF7AFEF8FD175CF2" blockId="13.[151,1437,188,396]" pageId="13" pageNumber="343">
<emphasis id="D8D8EADDB4389410FF7AFEF8FEA85C19" bold="true" box="[199,353,332,359]" pageId="13" pageNumber="343">Distribution.</emphasis>
Originally described from Perak (Western Malaysia) and Pahang (Eastern Malaysia). Newly recorded in Tanah Rata, Pahang (Eastern Malaysia).
</paragraph>
</subSubSection>
</treatment>
</document>

182
data/62/05/87/620587D9B43C9414FF2AFF6EFC305FE4.xml Normal file
View file

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<mods:title id="5310B8897818BCB21DFAFBF2B6169B07">Review of the genus Carlschoenherria (Coleoptera: Scarabaeidae: Melolonthinae) of the Philippines</mods:title>
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</mods:mods>
<treatment id="620587D9B43C9414FF2AFF6EFC305FE4" ID-GBIF-Taxon="158297462" LSID="urn:lsid:plazi:treatment:620587D9B43C9414FF2AFF6EFC305FE4" httpUri="http://treatment.plazi.org/id/620587D9B43C9414FF2AFF6EFC305FE4" lastPageNumber="339" pageId="9" pageNumber="339">
<subSubSection id="A2B66544B43C9414FF2AFF6EFCB15D8B" box="[151,888,218,245]" pageId="9" pageNumber="339" type="nomenclature">
<paragraph id="EA1336CFB43C9414FF2AFF6EFCB15D8B" blockId="9.[151,888,218,282]" box="[151,888,218,245]" pageId="9" pageNumber="339">
<heading id="B15B81A3B43C9414FF2AFF6EFCB15D8B" bold="true" box="[151,888,218,245]" fontSize="11" level="1" pageId="9" pageNumber="339" reason="1">
<emphasis id="D8D8EADDB43C9414FF2AFF6EFCB15D8B" bold="true" box="[151,888,218,245]" pageId="9" pageNumber="339">
<taxonomicName id="2DAC4D4CB43C9414FF2AFF6EFD535D8A" authority="(Moser, 1915)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[151,666,218,245]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="339" phylum="Arthropoda" rank="species" species="palawana" status="comb. nov.">
<emphasis id="D8D8EADDB43C9414FF2AFF6EFE205D8B" bold="true" box="[151,489,218,245]" italics="true" pageId="9" pageNumber="339">Carlschoenherria palawana</emphasis>
(
<bibRefCitation id="8E3D4B3EB43C9414FE44FF6EFD5B5D8A" author="Moser, J." box="[505,658,218,244]" pageId="9" pageNumber="339" pagination="113 - 151" refId="ref7803" refString="Moser, J. (1915) Beitrag zur Kenntnis der Melolonthiden (Col.). IV. Deutsche Entomologische Zeitschrift, 1915, 113 - 151. https: // doi. org / 10.1002 / mmnd. 48019150204" type="journal article" year="1915">Moser, 1915</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C3EB57A6B43C9414FD1FFF6FFCB15D8B" box="[674,888,219,245]" pageId="9" pageNumber="339" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43C9414FF2AFF4BFE915C64" box="[151,344,255,282]" pageId="9" pageNumber="339" type="description">
<paragraph id="EA1336CFB43C9414FF2AFF4BFE915C64" blockId="9.[151,888,218,282]" box="[151,344,255,282]" pageId="9" pageNumber="339">
(
<figureCitation id="72972A4AB43C9414FF22FF4BFEEE5C67" box="[159,295,255,281]" captionStart="FIGURES 16–19" captionStartId="8.[151,264,1979,2004]" captionTargetBox="[259,1424,674,1941]" captionTargetId="figure@8.[352,1235,697,1960]" captionTargetPageId="8" captionText="FIGURES 16–19. Carlschoenherria palawana (Moser), male from the Philippines, Palawan. 16, habitus, dorsal aspect; 17, pronotum, dorsal aspect; 18, genitalia, dorsal aspect; 19, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3249667/files/figure.png" pageId="9" pageNumber="339">Figs. 16–19</figureCitation>
,
<figureCitation id="72972A4AB43C9414FE8EFF4BFE995C64" box="[307,336,255,282]" captionStart="FIGURE 36" captionStartId="15.[151,250,1710,1735]" captionTargetBox="[360,1227,424,1689]" captionTargetId="figure@15.[340,1247,416,1691]" captionTargetPageId="15" captionText="FIGURE 36. Geographical distribution map of Carlschoenherria Bezděk, 2016 in the Philippines." httpUri="https://zenodo.org/record/3249673/files/figure.png" pageId="9" pageNumber="339">36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43C9414FF2AFEF7FBE85C04" pageId="9" pageNumber="339" type="reference_group">
<paragraph id="EA1336CFB43C9414FF2AFEF7FB975C25" blockId="9.[151,1118,322,378]" box="[151,1118,322,347]" pageId="9" pageNumber="339">
<treatmentCitationGroup id="CABC11E1B43C9414FF2AFEF7FB975C25" box="[151,1118,322,347]" pageId="9" pageNumber="339">
<taxonomicName id="2DAC4D4CB43C9414FF2AFEF7FDFE5C25" authority="Moser, 1915: 149" authorityName="Moser" authorityPageNumber="149" authorityYear="1915" box="[151,567,322,347]" class="Insecta" family="Curculionidae" genus="Schonherria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="339" phylum="Arthropoda" rank="species" species="palawana">
<emphasis id="D8D8EADDB43C9414FF2AFEF7FEB65C25" box="[151,383,323,347]" italics="true" pageId="9" pageNumber="339">Schönherria palawana</emphasis>
<treatmentCitation id="6B0D10DEB43C9414FE38FEF7FDFE5C25" author="Moser, J." box="[389,567,322,347]" page="149" pageId="9" pageNumber="339" year="1915">
<bibRefCitation id="8E3D4B3EB43C9414FE38FEF7FDFE5C25" author="Moser, J." box="[389,567,322,347]" pageId="9" pageNumber="339" pagination="113 - 151" refId="ref7803" refString="Moser, J. (1915) Beitrag zur Kenntnis der Melolonthiden (Col.). IV. Deutsche Entomologische Zeitschrift, 1915, 113 - 151. https: // doi. org / 10.1002 / mmnd. 48019150204" type="journal article" year="1915">Moser, 1915: 149</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(original description);
<treatmentCitation id="6B0D10DEB43C9414FC9FFEF6FC2C5C25" author="Schultze, W. A." box="[802,997,322,347]" page="180" pageId="9" pageNumber="339" year="1916">
<bibRefCitation id="8E3D4B3EB43C9414FC9FFEF6FC2C5C25" author="Schultze, W. A." box="[802,997,322,347]" pageId="9" pageNumber="339" pagination="176 - 180" refId="ref7844" refString="Schultze, W. A. (1916) Catalogue of Philippine Coleoptera. The Philippine Journal of Science, 11, 176 - 180." type="journal article" year="1916">Schultze 1916: 180</bibRefCitation>
</treatmentCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA1336CFB43C9414FF2AFED6FBE85C04" blockId="9.[151,1118,322,378]" box="[151,1057,353,378]" pageId="9" pageNumber="339">
<treatmentCitationGroup id="CABC11E1B43C9414FF2AFED6FBE85C04" box="[151,1057,353,378]" pageId="9" pageNumber="339">
<taxonomicName id="2DAC4D4CB43C9414FF2AFED6FDE95C04" authority=": Itoh 1992: 10" authorityName=": Itoh" authorityPageNumber="10" authorityYear="1992" box="[151,544,353,378]" class="Insecta" family="Curculionidae" genus="Schoenherria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="339" phylum="Arthropoda" rank="species" species="palawana">
<emphasis id="D8D8EADDB43C9414FF2AFED6FE435C04" box="[151,394,354,378]" italics="true" pageId="9" pageNumber="339">Schoenherria palawana</emphasis>
:
<treatmentCitation id="6B0D10DEB43C9414FE2AFED5FDE95C04" author="Itoh, T." box="[407,544,353,378]" page="10" pageId="9" pageNumber="339" year="1992">
<bibRefCitation id="8E3D4B3EB43C9414FE2AFED5FDE95C04" author="Itoh, T." box="[407,544,353,378]" pageId="9" pageNumber="339" pagination="7 - 14" refId="ref7594" refString="Itoh, T. (1992) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) I. Lamellicornia, 7, 7 - 14." type="journal article" year="1992">Itoh 1992: 10</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(checklist);
<bibRefCitation id="8E3D4B3EB43C9414FD23FED5FC845C04" author="Itoh, T." box="[670,845,353,378]" pageId="9" pageNumber="339" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh 1993: 10–12</bibRefCitation>
(redescription, key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43C9414FF2AFE1CFC665F50" pageId="9" pageNumber="339" type="materials_examined">
<paragraph id="EA1336CFB43C9414FF2AFE1CFD8B5CBD" blockId="9.[151,1436,423,667]" box="[151,578,423,451]" pageId="9" pageNumber="339">
<materialsCitation id="5AC43C92B43C9414FF2AFE1CFDF75CBD" ID-GBIF-Occurrence="2268317695" box="[151,574,423,451]" location="Bimaluan" pageId="9" pageNumber="339" specimenCount="1" stateProvince="Palawan" typeStatus="holotype">
<emphasis id="D8D8EADDB43C9414FF2AFE1CFEFF5CBC" bold="true" box="[151,310,423,450]" pageId="9" pageNumber="339">
<typeStatus id="3517886DB43C9414FF2AFE1CFF1B5CBC" box="[151,210,424,450]" pageId="9" pageNumber="339">Type</typeStatus>
locality.
</emphasis>
“
<collectingRegion id="2868F82DB43C9414FEFBFE1CFE615CBC" box="[326,424,424,450]" country="Philippines" name="Palawan" pageId="9" pageNumber="339">Palawan</collectingRegion>
(
<location id="EF736014B43C9414FE0AFE1CFDE45CBD" LSID="urn:lsid:plazi:treatment:620587D9B43C9414FF2AFF6EFC305FE4:EF736014B43C9414FE0AFE1CFDE45CBD" box="[439,557,424,451]" name="Bimaluan" pageId="9" pageNumber="339" stateProvince="Palawan">Bimaluan</location>
)”
</materialsCitation>
.
</paragraph>
<paragraph id="EA1336CFB43C9414FF7AFE78FDF35C99" blockId="9.[151,1436,423,667]" box="[199,570,459,487]" pageId="9" pageNumber="339">
<emphasis id="D8D8EADDB43C9414FF7AFE78FE265C98" bold="true" box="[199,495,459,486]" pageId="9" pageNumber="339">
<typeStatus id="3517886DB43C9414FF7AFE78FECB5C98" box="[199,258,460,486]" pageId="9" pageNumber="339">Type</typeStatus>
material examined.
</emphasis>
None.
</paragraph>
<paragraph id="EA1336CFB43C9414FF7AFE5BFC665F50" blockId="9.[151,1436,423,667]" pageId="9" pageNumber="339">
<emphasis id="D8D8EADDB43C9414FF7AFE5BFDFD5F74" bold="true" box="[199,564,495,522]" pageId="9" pageNumber="339">Additional material examined.</emphasis>
<materialsCitation id="5AC43C92B43C9414FD86FE5BFC625F50" ID-GBIF-Occurrence="2268317664" collectingDate="1985-10-04" collectingDateMax="2013-05-23" collectingDateMin="1985-10-04" collectionCode="NMM, ER-PC" collectorName="V. Samarita" country="Philippines" location="Trident Mining" municipality="Narra" pageId="9" pageNumber="339" specimenCount="2" specimenCount-male="2" stateProvince="Palawan" typeStatus="holotype">
<emphasis id="D8D8EADDB43C9414FD86FE5BFC2B5F74" bold="true" box="[571,994,495,522]" pageId="9" pageNumber="339">
<collectingCountry id="92BB765FB43C9414FD86FE5BFD3B5F74" box="[571,754,495,522]" name="Philippines" pageId="9" pageNumber="339">PHILIPPINES</collectingCountry>
: LUZON:
<collectingRegion id="2868F82DB43C9414FCCFFE44FC145F74" box="[882,989,496,522]" country="Philippines" name="Palawan" pageId="9" pageNumber="339">Palawan</collectingRegion>
:
</emphasis>
<specimenCount id="FCAAFD46B43C9414FC54FE44FBC75F74" box="[1001,1038,496,522]" pageId="9" pageNumber="339" type="male">1♂</specimenCount>
,
<collectingMunicipality id="0A77ACB5B43C9414FBA7FE45FB945F75" box="[1050,1117,497,523]" pageId="9" pageNumber="339">Narra</collectingMunicipality>
,
<location id="EF736014B43C9414FBD5FE44FADC5F74" LSID="urn:lsid:plazi:treatment:620587D9B43C9414FF2AFF6EFC305FE4:EF736014B43C9414FBD5FE44FADC5F74" box="[1128,1301,496,522]" country="Philippines" municipality="Narra" name="Trident Mining" pageId="9" pageNumber="339" stateProvince="Palawan">Trident Mining</location>
,
<collectingDate id="8E56E9E7B43C9414FA9CFE5BFA515F74" box="[1313,1432,495,522]" pageId="9" pageNumber="339" value="1985-10-04">
<date id="9E12100FB43C9414FA9CFE5BFA515F74" box="[1313,1432,495,522]" pageId="9" pageNumber="339" value="1985-10-04">04-x-1985</date>
</collectingDate>
,
<collectorName id="47595319B43C9414FF2AFDA1FED35F50" box="[151,282,532,559]" pageId="9" pageNumber="339">V. Samarita</collectorName>
(
<collectionCode id="8CBDAE0AB43C9414FE95FDA0FEBD5F50" box="[296,372,532,558]" pageId="9" pageNumber="339">NMM</collectionCode>
);
<emphasis id="D8D8EADDB43C9414FE35FDA7FE375F50" bold="true" box="[392,510,531,558]" pageId="9" pageNumber="339">VISAYAS</emphasis>
:
<emphasis id="D8D8EADDB43C9414FDB1FDA7FDAC5F50" bold="true" box="[524,613,531,558]" pageId="9" pageNumber="339">
<collectingRegion id="2868F82DB43C9414FDB1FDA7FDA95F50" box="[524,608,531,558]" country="Philippines" name="Samar" pageId="9" pageNumber="339">Samar</collectingRegion>
:
</emphasis>
<specimenCount id="FCAAFD46B43C9414FDD1FDA0FD595F50" box="[620,656,532,558]" pageId="9" pageNumber="339" type="male">1♂</specimenCount>
,
<collectingDate id="8E56E9E7B43C9414FD20FDA0FCF35F50" box="[669,826,532,558]" pageId="9" pageNumber="339" value="2013-05-22" valueMax="2013-05-23" valueMin="2013-05-22">
<date id="9E12100FB43C9414FD20FDA0FCF35F50" box="[669,826,532,558]" pageId="9" pageNumber="339" value="2013-05-22" valueMax="2013-05-23" valueMin="2013-05-22">22-23-v-2013</date>
</collectingDate>
, (
<collectionCode id="8CBDAE0AB43C9414FCF0FDA0FC6A5F50" box="[845,931,532,558]" pageId="9" pageNumber="339">ER-PC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43C9414FF7AFD83FC305FE4" pageId="9" pageNumber="339" type="distribution">
<paragraph id="EA1336CFB43C9414FF7AFD83FC305FE4" blockId="9.[151,1436,423,667]" pageId="9" pageNumber="339">
<emphasis id="D8D8EADDB43C9414FF7AFD83FEA85F2C" bold="true" box="[199,353,567,594]" pageId="9" pageNumber="339">Distribution.</emphasis>
Moser (1905) described this species from
<collectingRegion id="2868F82DB43C9414FC88FD8CFC5D5F2C" box="[821,916,568,594]" country="Philippines" name="Palawan" pageId="9" pageNumber="339">Palawan</collectingRegion>
Island (Western
<collectingCountry id="92BB765FB43C9414FBF3FD8CFB185F2D" box="[1102,1233,568,595]" name="Philippines" pageId="9" pageNumber="339">Philippines</collectingCountry>
), later,
<bibRefCitation id="8E3D4B3EB43C9414FAA2FD8CFA555F2C" author="Itoh, T." box="[1311,1436,568,594]" pageId="9" pageNumber="339" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh (1993)</bibRefCitation>
added records from Luzon, Mindoro,
<collectingRegion id="2868F82DB43C9414FDFEFDE8FD6C5F08" box="[579,677,604,630]" country="Philippines" name="Palawan" pageId="9" pageNumber="339">Palawan</collectingRegion>
, Negros,
<collectingRegion id="2868F82DB43C9414FCADFDE9FC9A5F09" box="[784,851,605,631]" country="Philippines" name="Southern Leyte" pageId="9" pageNumber="339">Leyte</collectingRegion>
,
<collectingRegion id="2868F82DB43C9414FCE2FDE8FC0E5F08" box="[863,967,604,630]" country="Philippines" name="Romblon" pageId="9" pageNumber="339">Romblon</collectingRegion>
and Sibuyan Islands. Specimen from Sa- mar
<collectingCountry id="92BB765FB43C9414FF77FD34FEC75FE4" box="[202,270,640,666]" name="Iceland" pageId="9" pageNumber="339">Island</collectingCountry>
(Eastern Visayas) represents a new faunistic record for this island.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="DC4ED345AED1381A2B12B6F9BE90C64F">Review of the genus Carlschoenherria (Coleoptera: Scarabaeidae: Melolonthinae) of the Philippines</mods:title>
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<treatment id="620587D9B43E9411FF2AF8B0FD5F5C19" ID-DOI="http://doi.org/10.5281/zenodo.5930692" ID-GBIF-Taxon="184209185" ID-Zenodo-Dep="5930692" LSID="urn:lsid:plazi:treatment:620587D9B43E9411FF2AF8B0FD5F5C19" httpUri="http://treatment.plazi.org/id/620587D9B43E9411FF2AF8B0FD5F5C19" lastPageId="12" lastPageNumber="342" pageId="11" pageNumber="341">
<subSubSection id="A2B66544B43E9416FF2AF8B0FC4C5A60" box="[151,901,1796,1823]" pageId="11" pageNumber="341" type="nomenclature">
<paragraph id="EA1336CFB43E9416FF2AF8B0FC4C5A60" blockId="11.[151,901,1796,1859]" box="[151,901,1796,1823]" pageId="11" pageNumber="341">
<heading id="B15B81A3B43E9416FF2AF8B0FC4C5A60" bold="true" box="[151,901,1796,1823]" fontSize="11" level="1" pageId="11" pageNumber="341" reason="1">
<emphasis id="D8D8EADDB43E9416FF2AF8B0FC4C5A60" bold="true" box="[151,901,1796,1823]" pageId="11" pageNumber="341">
<taxonomicName id="2DAC4D4CB43E9416FF2AF8B0FD615A60" authority="(Burmeister, 1855)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Burmeister" baseAuthorityYear="1855" box="[151,680,1796,1823]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="341" phylum="Arthropoda" rank="species" species="argus" status="comb. nov.">
<emphasis id="D8D8EADDB43E9416FF2AF8B0FE715A61" bold="true" box="[151,440,1796,1823]" italics="true" pageId="11" pageNumber="341">Carlschoenherria argus</emphasis>
(
<bibRefCitation id="8E3D4B3EB43E9416FE75F8B0FD565A60" author="Burmeister, H. C. C." box="[456,671,1796,1822]" pageId="11" pageNumber="341" refId="ref7491" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Fortsetzung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Theod. Chr. Friedr. Enslin, Berlin: x + 569 pp." type="book" year="1855">Burmeister, 1855</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C3EB57A6B43E9416FD12F8B1FC4C5A60" box="[687,901,1796,1823]" pageId="11" pageNumber="341" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43E9416FF2AF89CFEE75A3D" box="[151,302,1832,1859]" pageId="11" pageNumber="341" type="description">
<paragraph id="EA1336CFB43E9416FF2AF89CFEE75A3D" blockId="11.[151,901,1796,1859]" box="[151,302,1832,1859]" pageId="11" pageNumber="341">
(
<figureCitation id="72972A4AB43E9416FF22F89CFEEF5A3D" box="[159,294,1832,1859]" captionStart="FIGURES 29–31" captionStartId="12.[151,264,1726,1751]" captionTargetBox="[340,1247,380,1648]" captionTargetId="figure@12.[340,1247,380,1706]" captionText="FIGURES 29–31. Carlschoenherria argus (Burmeister), male from Indonesia, Java. 29, habitus, dorsal aspect; 30, genitalia, dorsal aspect; 31, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3405257/files/figure.png" pageId="11" pageNumber="341">Figs. 29–31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43E9416FF2AF8D9FCB15ABD" pageId="11" pageNumber="341" type="reference_group">
<paragraph id="EA1336CFB43E9416FF2AF8D9FC825AFB" blockId="11.[151,1213,1900,2023]" box="[151,843,1900,1925]" pageId="11" pageNumber="341">
<taxonomicName id="2DAC4D4CB43E9416FF2AF8D9FDAF5AFB" authority="Burmeister, 1855: 419 - 420" authorityName="Burmeister" authorityPageNumber="419" authorityYear="1855" box="[151,614,1900,1925]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="341" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="D8D8EADDB43E9416FF2AF8D9FE825AFB" box="[151,331,1901,1925]" italics="true" pageId="11" pageNumber="341">Melolontha argus</emphasis>
<bibRefCitation id="8E3D4B3EB43E9416FEECF8D8FDAF5AFB" author="Burmeister, H. C. C." box="[337,614,1900,1925]" pageId="11" pageNumber="341" refId="ref7491" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Fortsetzung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Theod. Chr. Friedr. Enslin, Berlin: x + 569 pp." type="book" year="1855">Burmeister, 1855: 419–420</bibRefCitation>
</taxonomicName>
(original description).
</paragraph>
<paragraph id="EA1336CFB43E9416FF2AF838FB745ADA" blockId="11.[151,1213,1900,2023]" box="[151,1213,1931,1956]" pageId="11" pageNumber="341">
<treatmentCitationGroup id="CABC11E1B43E9416FF2AF838FB745ADA" box="[151,1213,1931,1956]" pageId="11" pageNumber="341">
<taxonomicName id="2DAC4D4CB43E9416FF2AF838FDDA5ADA" authority=": Brenske 1894: 27" authorityName="Brenske" authorityPageNumber="27" authorityYear="1894" box="[151,531,1931,1956]" class="Insecta" family="Curculionidae" genus="Schonherria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="341" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="D8D8EADDB43E9416FF2AF838FE9D5ADD" box="[151,340,1932,1956]" italics="true" pageId="11" pageNumber="341">Schönherria argus</emphasis>
:
<treatmentCitation id="6B0D10DEB43E9416FEDCF83FFDDA5ADA" author="Brenske, E." box="[353,531,1931,1956]" page="27" pageId="11" pageNumber="341" year="1894">
<bibRefCitation id="8E3D4B3EB43E9416FEDCF83FFDDA5ADA" author="Brenske, E." box="[353,531,1931,1956]" pageId="11" pageNumber="341" pagination="3 - 87" refId="ref7416" refString="Brenske, E. (1894) Die Melolonthiden der palaearctischen und orientalischen Region in Koniglichen Naturhistorischen Muse- um zu Brussel. Beschreibung neuer Arten und Bemerkungen zu bekannten. Memoires de la Societe Entomologique de Belgique, 2, 3 - 87." type="journal article" year="1894">Brenske 1894: 27</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(record from
<collectingCountry id="92BB765FB43E9416FD1CF83FFD325ADA" box="[673,763,1931,1956]" name="Malaysia" pageId="11" pageNumber="341">Malacca</collectingCountry>
);
<treatmentCitation id="6B0D10DEB43E9416FCB0F83FFC055ADA" author="Brenske, E." box="[781,972,1931,1956]" page="155" pageId="11" pageNumber="341" year="1900">
<bibRefCitation id="8E3D4B3EB43E9416FCB0F83FFC055ADA" author="Brenske, E." box="[781,972,1931,1956]" pageId="11" pageNumber="341" pagination="141 - 155" refId="ref7460" refString="Brenske, E. (1900) Melolonthiden gesammelt von J. - L. Weyers. Memoires de la Societe Entomologique de Belgique, 7, 141 - 155." type="journal article" year="1900">Brenske 1900: 155</bibRefCitation>
</treatmentCitation>
(Sumatra, new record).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA1336CFB43E9416FF2AF81EFCB15ABD" blockId="11.[151,1213,1900,2023]" box="[151,888,1962,1987]" pageId="11" pageNumber="341">
<treatmentCitationGroup id="CABC11E1B43E9416FF2AF81EFCB15ABD" box="[151,888,1962,1987]" pageId="11" pageNumber="341">
<taxonomicName id="2DAC4D4CB43E9416FF2AF81EFE235ABC" authority=": Itoh 1992: 9" authorityName="Itoh" authorityPageNumber="9" authorityYear="1992" box="[151,490,1962,1987]" class="Insecta" family="Curculionidae" genus="Schoenherria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="341" phylum="Arthropoda" rank="species" species="argus">
<emphasis id="D8D8EADDB43E9416FF2AF81EFE965ABC" box="[151,351,1962,1986]" italics="true" pageId="11" pageNumber="341">Schoenherria argus</emphasis>
:
<treatmentCitation id="6B0D10DEB43E9416FED1F81EFE235ABC" author="Itoh, T." box="[364,490,1962,1987]" page="9" pageId="11" pageNumber="341" year="1992">
<bibRefCitation id="8E3D4B3EB43E9416FED1F81EFE235ABC" author="Itoh, T." box="[364,490,1962,1987]" pageId="11" pageNumber="341" pagination="7 - 14" refId="ref7594" refString="Itoh, T. (1992) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) I. Lamellicornia, 7, 7 - 14." type="journal article" year="1992">Itoh 1992: 9</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<bibRefCitation id="8E3D4B3EB43E9416FE48F81EFD6D5ABD" author="Itoh, T." box="[501,676,1962,1987]" pageId="11" pageNumber="341" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh 1993: 14–15</bibRefCitation>
(redescription, key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43E9411FF2AF879FEE95C61" lastPageId="12" lastPageNumber="342" pageId="11" pageNumber="341" type="materials_examined">
<paragraph id="EA1336CFB43E9416FF2AF879FE425A99" blockId="11.[151,1213,1900,2023]" box="[151,395,1996,2023]" pageId="11" pageNumber="341">
<materialsCitation id="5AC43C92B43E9416FF2AF879FE425A99" ID-GBIF-Occurrence="2413160153" box="[151,395,1996,2023]" location="Java" pageId="11" pageNumber="341" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<emphasis id="D8D8EADDB43E9416FF2AF879FEFF5A99" bold="true" box="[151,310,1996,2023]" pageId="11" pageNumber="341">
<typeStatus id="3517886DB43E9416FF2AF879FF1B5A99" box="[151,210,1997,2023]" pageId="11" pageNumber="341">Type</typeStatus>
locality.
</emphasis>
“
<collectingRegion id="2868F82DB43E9416FEFAF879FE495A99" box="[327,384,1997,2023]" country="Indonesia" name="Jawa Timur" pageId="11" pageNumber="341">Java</collectingRegion>
”
</materialsCitation>
</paragraph>
<paragraph id="EA1336CFB43E9416FE37F879FE475A99" blockId="11.[151,1213,1900,2023]" box="[394,398,1997,2023]" pageId="11" pageNumber="341">.</paragraph>
<paragraph id="EA1336CFB4399411FF7AFF2DFDFA5DCD" blockId="12.[151,1437,152,359]" box="[199,563,152,179]" pageId="12" pageNumber="342">
<emphasis id="D8D8EADDB4399411FF7AFF2DFE265DCD" bold="true" box="[199,495,152,179]" pageId="12" pageNumber="342">Type material examined.</emphasis>
None
</paragraph>
<paragraph id="EA1336CFB4399411FF7AFF08FEE95C61" blockId="12.[151,1437,152,359]" pageId="12" pageNumber="342">
<emphasis id="D8D8EADDB4399411FF7AFF08FC8F5DA9" bold="true" box="[199,838,188,215]" pageId="12" pageNumber="342">Additional material examined. WEST MALAYSIA.</emphasis>
1♂, Pahang, Cameron Highlands, 2-9-iv-1997, D. Hauck Igt. (OCPC); 1♀, Pahang, Banlaran Benon Mts., K. Ulo Dong, SSE, 17-23-iv-1997, 10-15 km, D. Hauck Igt. (OCPC)
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B4399411FF7AFE9DFD5F5C19" pageId="12" pageNumber="342" type="discussion">
<paragraph id="EA1336CFB4399411FF7AFE9DFD5F5C19" blockId="12.[151,1437,152,359]" pageId="12" pageNumber="342">
<emphasis id="D8D8EADDB4399411FF7AFE9DFEF95C3D" bold="true" box="[199,304,297,323]" pageId="12" pageNumber="342">Remark.</emphasis>
A widely distributed species, originally described from Java, subsequently recorded from Malaysia and Sumatra where it is common (Itoh 1993).
</paragraph>
</subSubSection>
</treatment>
</document>

184
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@ -0,0 +1,184 @@
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<mods:title id="28159CB6EB36624C7FEBFBD902EBEAD4">Review of the genus Carlschoenherria (Coleoptera: Scarabaeidae: Melolonthinae) of the Philippines</mods:title>
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<paragraph id="EA1336CFB43F9417FF2AF8B8FC645A58" blockId="10.[151,941,1804,1867]" box="[151,941,1804,1831]" pageId="10" pageNumber="340">
<heading id="B15B81A3B43F9417FF2AF8B8FC645A58" bold="true" box="[151,941,1804,1831]" fontSize="11" level="1" pageId="10" pageNumber="340" reason="1">
<emphasis id="D8D8EADDB43F9417FF2AF8B8FC645A58" bold="true" box="[151,941,1804,1831]" pageId="10" pageNumber="340">
<taxonomicName id="2DAC4D4CB43F9417FF2AF8B8FD065A58" authority="(Brenske, 1894)" authorityName="Calcetas" authorityYear="2019" baseAuthorityName="Brenske" baseAuthorityYear="1894" box="[151,719,1804,1831]" class="Insecta" family="Melolonthidae" genus="Carlschoenherria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="340" phylum="Arthropoda" rank="species" species="philippinica" status="comb. nov.">
<emphasis id="D8D8EADDB43F9417FF2AF8B8FDCC5A59" bold="true" box="[151,517,1804,1831]" italics="true" pageId="10" pageNumber="340">Carlschoenherria philippinica</emphasis>
(
<bibRefCitation id="8E3D4B3EB43F9417FDA9F8B8FD0E5A58" author="Brenske, E." box="[532,711,1804,1830]" pageId="10" pageNumber="340" pagination="3 - 87" refId="ref7416" refString="Brenske, E. (1894) Die Melolonthiden der palaearctischen und orientalischen Region in Koniglichen Naturhistorischen Muse- um zu Brussel. Beschreibung neuer Arten und Bemerkungen zu bekannten. Memoires de la Societe Entomologique de Belgique, 2, 3 - 87." type="journal article" year="1894">Brenske, 1894</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C3EB57A6B43F9417FD6AF8B9FC645A58" box="[727,941,1804,1831]" pageId="10" pageNumber="340" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43F9417FF2AF884FE915A35" box="[151,344,1840,1867]" pageId="10" pageNumber="340" type="description">
<paragraph id="EA1336CFB43F9417FF2AF884FE915A35" blockId="10.[151,941,1804,1867]" box="[151,344,1840,1867]" pageId="10" pageNumber="340">
(
<figureCitation id="72972A4AB43F9417FF22F884FEEE5A34" box="[159,295,1840,1866]" captionStart="FIGURES 24–28" captionStartId="11.[151,264,1654,1679]" captionTargetBox="[267,1223,359,1619]" captionTargetId="figure@11.[343,1230,380,1633]" captionTargetPageId="11" captionText="FIGURES 24–28. Carlschoenherria philippinica (Brenske), male from the Philippines. 24, habitus, dorsal aspect; 25, prono- tum, dorsal aspect; 26, antenna, lateral aspect; 27 genitalia, dorsal aspect; 28, genitalia, lateral aspect." httpUri="https://zenodo.org/record/3249669/files/figure.png" pageId="10" pageNumber="340">Figs. 24–28</figureCitation>
,
<figureCitation id="72972A4AB43F9417FE8EF884FE995A35" box="[307,336,1840,1867]" captionStart="FIGURE 36" captionStartId="15.[151,250,1710,1735]" captionTargetBox="[360,1227,424,1689]" captionTargetId="figure@15.[340,1247,416,1691]" captionTargetPageId="15" captionText="FIGURE 36. Geographical distribution map of Carlschoenherria Bezděk, 2016 in the Philippines." httpUri="https://zenodo.org/record/3249673/files/figure.png" pageId="10" pageNumber="340">36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43F9417FF2AF8C1FBF05AAE" pageId="10" pageNumber="340" type="reference_group">
<paragraph id="EA1336CFB43F9417FF2AF8C1FE6F5AD2" blockId="10.[151,1436,1908,2036]" pageId="10" pageNumber="340">
<treatmentCitationGroup id="CABC11E1B43F9417FF2AF8C1FE6F5AD2" pageId="10" pageNumber="340">
<taxonomicName id="2DAC4D4CB43F9417FF2AF8C1FD2A5AF3" authority="Brenske, 1894: 27" authorityName="Brenske" authorityPageNumber="27" authorityYear="1894" box="[151,739,1908,1933]" class="Insecta" family="Curculionidae" genus="Schonherria" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="340" phylum="Arthropoda" rank="variety" species="hispida" variety="philippinica">
<emphasis id="D8D8EADDB43F9417FF2AF8C1FEA05AF3" box="[151,361,1909,1933]" italics="true" pageId="10" pageNumber="340">Schönherria hispida</emphasis>
var.
<emphasis id="D8D8EADDB43F9417FE1CF8C1FDD25AF3" box="[417,539,1909,1933]" italics="true" pageId="10" pageNumber="340">philippinica</emphasis>
<treatmentCitation id="6B0D10DEB43F9417FD99F8C0FD2A5AF3" author="Brenske, E." box="[548,739,1908,1933]" page="27" pageId="10" pageNumber="340" year="1894">
<bibRefCitation id="8E3D4B3EB43F9417FD99F8C0FD2A5AF3" author="Brenske, E." box="[548,739,1908,1933]" pageId="10" pageNumber="340" pagination="3 - 87" refId="ref7416" refString="Brenske, E. (1894) Die Melolonthiden der palaearctischen und orientalischen Region in Koniglichen Naturhistorischen Muse- um zu Brussel. Beschreibung neuer Arten und Bemerkungen zu bekannten. Memoires de la Societe Entomologique de Belgique, 2, 3 - 87." type="journal article" year="1894">Brenske, 1894: 27</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(original description);
<treatmentCitation id="6B0D10DEB43F9417FC6AF8C0FB095AF3" author="Dalla Torre, K. W." box="[983,1216,1908,1933]" page="286" pageId="10" pageNumber="340" year="1912">
<bibRefCitation id="8E3D4B3EB43F9417FC6AF8C0FB095AF3" author="Dalla Torre, K. W." box="[983,1216,1908,1933]" pageId="10" pageNumber="340" pagination="135 - 290" refId="ref7537" refString="Dalla Torre, K. W. (1912) Pars 49. Scarabaeidae: Melolonthinae III. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. XX. Pars. 49. W. Junk, Berlin, pp. 135 - 290." type="book chapter" year="1912">Dalla Torre 1912: 286</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="6B0D10DEB43F9417FAF8F8C0FEE45AD5" author="Schultze, W. A." page="180" pageId="10" pageNumber="340" year="1916">
<bibRefCitation id="8E3D4B3EB43F9417FAF8F8C0FEE45AD5" author="Schultze, W. A." pageId="10" pageNumber="340" pagination="176 - 180" refId="ref7844" refString="Schultze, W. A. (1916) Catalogue of Philippine Coleoptera. The Philippine Journal of Science, 11, 176 - 180." type="journal article" year="1916">Schultze 1916: 180</bibRefCitation>
</treatmentCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EA1336CFB43F9417FF2AF803FBF05AAE" blockId="10.[151,1436,1908,2036]" box="[151,1081,1975,2000]" pageId="10" pageNumber="340">
<treatmentCitationGroup id="CABC11E1B43F9417FF2AF803FBF05AAE" box="[151,1081,1975,2000]" pageId="10" pageNumber="340">
<taxonomicName id="2DAC4D4CB43F9417FF2AF803FDE35AB1" authority=": Itoh 1992: 9" authorityName="Itoh" authorityPageNumber="9" authorityYear="1992" box="[151,554,1975,2000]" class="Insecta" family="Curculionidae" genus="Schoenherria" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="340" phylum="Arthropoda" rank="species" species="philippinica">
<emphasis id="D8D8EADDB43F9417FF2AF803FE695AB1" box="[151,416,1975,1999]" italics="true" pageId="10" pageNumber="340">Schoenherria philippinica</emphasis>
:
<treatmentCitation id="6B0D10DEB43F9417FE13F803FDE35AB1" author="Itoh, T." box="[430,554,1975,2000]" page="9" pageId="10" pageNumber="340" year="1992">
<bibRefCitation id="8E3D4B3EB43F9417FE13F803FDE35AB1" author="Itoh, T." box="[430,554,1975,2000]" pageId="10" pageNumber="340" pagination="7 - 14" refId="ref7594" refString="Itoh, T. (1992) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) I. Lamellicornia, 7, 7 - 14." type="journal article" year="1992">Itoh 1992: 9</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(valid species);
<bibRefCitation id="8E3D4B3EB43F9417FD6DF803FCAF5AAE" author="Itoh, T." box="[720,870,1975,2000]" pageId="10" pageNumber="340" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh 1993: 6–7</bibRefCitation>
(redescription, key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43F9416FF2AF86EFC3B5D85" lastPageId="11" lastPageNumber="341" pageId="10" pageNumber="340" type="materials_examined">
<paragraph id="EA1336CFB43F9417FF2AF86EFE175A8A" blockId="10.[151,1436,1908,2036]" box="[151,478,2009,2036]" pageId="10" pageNumber="340">
<emphasis id="D8D8EADDB43F9417FF2AF86EFEFF5A8A" bold="true" box="[151,310,2009,2036]" pageId="10" pageNumber="340">
<typeStatus id="3517886DB43F9417FF2AF86EFF1B5A8A" box="[151,210,2010,2036]" pageId="10" pageNumber="340">Type</typeStatus>
locality.
</emphasis>
“
<collectingCountry id="92BB765FB43F9417FEFBF86EFE195A8A" box="[326,464,2010,2036]" name="Philippines" pageId="10" pageNumber="340">Philippinen</collectingCountry>
”.
</paragraph>
<paragraph id="EA1336CFB43E9416FF7AFF2DFDF35DCD" blockId="11.[151,1436,152,359]" box="[199,570,152,179]" pageId="11" pageNumber="341">
<emphasis id="D8D8EADDB43E9416FF7AFF2DFE265DCD" bold="true" box="[199,495,152,179]" pageId="11" pageNumber="341">
<typeStatus id="3517886DB43E9416FF7AFF2DFECB5DCD" box="[199,258,153,179]" pageId="11" pageNumber="341">Type</typeStatus>
material examined.
</emphasis>
None.
</paragraph>
<paragraph id="EA1336CFB43E9416FF7AFF08FC3B5D85" blockId="11.[151,1436,152,359]" pageId="11" pageNumber="341">
<emphasis id="D8D8EADDB43E9416FF7AFF08FDF55DA9" bold="true" box="[199,572,188,215]" pageId="11" pageNumber="341">Additional material examined.</emphasis>
<materialsCitation id="5AC43C92B43E9416FDFBFF08FA515DA9" ID-GBIF-Occurrence="2268317690" box="[582,1432,188,215]" collectionCode="UNSM" collectorName="Tangcolan" country="Philippines" location="Bukidnon" pageId="11" pageNumber="341" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Bukidnon" typeStatus="holotype">
<collectingCountry id="92BB765FB43E9416FDFBFF08FD305DA9" box="[582,761,188,215]" name="Philippines" pageId="11" pageNumber="341">
<emphasis id="D8D8EADDB43E9416FDFBFF08FD305DA9" bold="true" box="[582,761,188,215]" pageId="11" pageNumber="341">PHILIPPINES</emphasis>
</collectingCountry>
:
<emphasis id="D8D8EADDB43E9416FCB1FF08FC4B5DA9" bold="true" box="[780,898,188,215]" pageId="11" pageNumber="341">VISAYAS</emphasis>
:
<emphasis id="D8D8EADDB43E9416FC29FF08FBDD5DA9" bold="true" box="[916,1044,188,215]" pageId="11" pageNumber="341">
<collectingRegion id="2868F82DB43E9416FC29FF08FBC65DA9" box="[916,1039,188,215]" country="Philippines" name="Bukidnon" pageId="11" pageNumber="341">Bukidnon</collectingRegion>
:
</emphasis>
<specimenCount id="FCAAFD46B43E9416FBA2FF09FB935DA9" box="[1055,1114,189,215]" pageId="11" pageNumber="341" type="male">2♂♂</specimenCount>
,
<specimenCount id="FCAAFD46B43E9416FBD4FF09FB685DA9" box="[1129,1185,189,215]" pageId="11" pageNumber="341" type="female">3♀♀</specimenCount>
,
<collectorName id="47595319B43E9416FB0FFF08FAEF5DA8" box="[1202,1318,188,214]" pageId="11" pageNumber="341">Tangcolan</collectorName>
(
<collectionCode id="8CBDAE0AB43E9416FA84FF09FA595DA9" box="[1337,1424,189,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="11" pageNumber="341">UNSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5AC43C92B43E9416FF2AFF54FD975D85" ID-GBIF-Occurrence="2268317674" box="[151,606,224,251]" collectionCode="NMM" collectorName="V. Samarita" country="Philippines" pageId="11" pageNumber="341" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="D8D8EADDB43E9416FF2AFF54FE885D85" bold="true" box="[151,321,224,251]" pageId="11" pageNumber="341">MINDANAO:</emphasis>
<specimenCount id="FCAAFD46B43E9416FEF5FF55FEA55D85" box="[328,364,225,251]" pageId="11" pageNumber="341" type="male">1♂</specimenCount>
,
<collectorName id="47595319B43E9416FEC5FF55FE325D84" box="[376,507,224,251]" pageId="11" pageNumber="341">V. Samarita</collectorName>
(
<collectionCode id="8CBDAE0AB43E9416FDB4FF55FD9C5D85" box="[521,597,225,251]" pageId="11" pageNumber="341">NMM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="5AC43C92B43E9416FDD4FF54FC3B5D85" ID-GBIF-Occurrence="2413160286" box="[617,1010,224,251]" collectionCode="UNSM" collectorName="C. Baker" country="Philippines" location="Surigao" pageId="11" pageNumber="341" specimenCount="4" specimenCount-male="4" typeStatus="holotype">
<emphasis id="D8D8EADDB43E9416FDD4FF54FD065D85" bold="true" box="[617,719,224,251]" pageId="11" pageNumber="341">
<location id="EF736014B43E9416FDD4FF54FD025D85" LSID="urn:lsid:plazi:treatment:620587D9B43F9416FF2AF8B8FCDF5C19:EF736014B43E9416FDD4FF54FD025D85" box="[617,715,224,251]" country="Philippines" name="Surigao" pageId="11" pageNumber="341">Surigao</location>
:
</emphasis>
<specimenCount id="FCAAFD46B43E9416FD6BFF55FCD85D85" box="[726,785,225,251]" pageId="11" pageNumber="341" type="male">4♂♂</specimenCount>
,
<collectorName id="47595319B43E9416FCA1FF55FC495D84" box="[796,896,224,251]" pageId="11" pageNumber="341">C. Baker</collectorName>
(
<collectionCode id="8CBDAE0AB43E9416FC32FF55FC2F5D85" box="[911,998,225,251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="11" pageNumber="341">UNSM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2B66544B43E9416FF7AFEB0FCDF5C19" pageId="11" pageNumber="341" type="distribution">
<paragraph id="EA1336CFB43E9416FF7AFEB0FCDF5C19" blockId="11.[151,1436,152,359]" pageId="11" pageNumber="341">
<emphasis id="D8D8EADDB43E9416FF7AFEB0FEA85C61" bold="true" box="[199,353,260,287]" pageId="11" pageNumber="341">Distribution.</emphasis>
<bibRefCitation id="8E3D4B3EB43E9416FED5FEB0FDDC5C61" author="Brenske, E." box="[360,533,260,287]" pageId="11" pageNumber="341" pagination="3 - 87" refId="ref7416" refString="Brenske, E. (1894) Die Melolonthiden der palaearctischen und orientalischen Region in Koniglichen Naturhistorischen Muse- um zu Brussel. Beschreibung neuer Arten und Bemerkungen zu bekannten. Memoires de la Societe Entomologique de Belgique, 2, 3 - 87." type="journal article" year="1894">Brenske (1894)</bibRefCitation>
described this species from “
<collectingCountry id="92BB765FB43E9416FCE3FEB0FC2E5C60" box="[862,999,260,286]" name="Philippines" pageId="11" pageNumber="341">Philippinen</collectingCountry>
”, later,
<bibRefCitation id="8E3D4B3EB43E9416FB81FEB0FB725C61" author="Itoh, T." box="[1084,1211,260,287]" pageId="11" pageNumber="341" pagination="3 - 20" refId="ref7624" refString="Itoh, T. (1993) A taxonomical study on the genus Schoenherria Burmeister (Melolonthinae: Melolonthidae) II. Lamellicornia, 9, 3 - 20." type="journal article" year="1993">Itoh (1993)</bibRefCitation>
added records from Mindanao and
<collectingRegion id="2868F82DB43E9416FEFCFE9DFE485C3D" box="[321,385,297,323]" country="Philippines" name="Southern Leyte" pageId="11" pageNumber="341">Leyte</collectingRegion>
(Eastern Visayas). The species is newly recorded from the provinces of
<collectingRegion id="2868F82DB43E9416FB0AFE9CFAEC5C3C" box="[1207,1317,296,322]" country="Philippines" name="Bukidnon" pageId="11" pageNumber="341">Bukidnon</collectingRegion>
(Northern Mindanao) and Surigao (North-eastern tip of Mindanao).
</paragraph>
</subSubSection>
</treatment>
</document>

61
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<mods:title id="7448B044AD831B5032D67FA4E910BD88">Species plantarum: exhibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas</mods:title>
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<treatment id="6205BCE6D7463BEB2E4273938AF3F162" ID-DOI="http://doi.org/10.5281/zenodo.4358558" ID-GBIF-Taxon="164947349" ID-Zenodo-Dep="4358558" LSID="urn:lsid:plazi:treatment:6205BCE6D7463BEB2E4273938AF3F162" httpUri="http://treatment.plazi.org/id/6205BCE6D7463BEB2E4273938AF3F162" lastPageNumber="107" pageNumber="106">
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<taxonomicName id="73F15DDBEFDD3F18908E1C8334CB279D" ID-CoL="3F6CQ" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="pusillum" status="spec. nov.">Galium pusillum</taxonomicName>
,
<taxonomicNameLabel id="9EA7BF216C1FDB8F0BC8763B37C3EEA3" rank="species">spec. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="443AE3F6F651B8BD79EBE8E6CBA0543E" type="description">
<paragraph id="8A0EF754A69F0745BD89B07BF09781E1" pageNumber="106">10. Galium foliis octonis hispidis linearibus acuminaris subimbricatis, pedunculis dichotomis.</paragraph>
<paragraph id="C5473CF22CC22C11A22F1212662A100B" pageNumber="107">
<pageBreakToken id="96FF82D6DD555B3C6AEFC0ACBB9D09E3" pageNumber="107">Rubeola</pageBreakToken>
saxatilis.
<emphasis id="251F91346F5DD90FA47A8E67C6CAC901" italics="true">Bauh. pin. 334. prodr. 145.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="9447F006775D3910E5739CFFD1B055E3" type="distribution">
<paragraph id="359C2AE94577463B19D0E5D8AB8740E5" pageNumber="107">
<emphasis id="B16D751B2C80BADBF07DFF1BEE2CD6C7" italics="true">Habitat in montibus</emphasis>
Galloprovinciae
<emphasis id="D60330B9600791A6048C788F353F2AFF" italics="true">.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="9804659887AC850C65AB082EB839B168" type="description">
<paragraph id="404BA545DE5A44E135B103B2B556081C" pageNumber="107">
Caules
<emphasis id="B91C331C342B05BF2B6F51935F22217F" italics="true">numerosi, digitales, angulati;</emphasis>
Folia
<emphasis id="6B03887B721450E8ED21112F8AEF35BC" italics="true">verticillata foliolis 8. s.6, lanceolato-linearibus vel linearibus, acutis, uti caulis undique pilis patentibus hispida. Rami rariores, alterni; Foliorum verticilli saepe ita conferti, ut folia imbricatim digesta conspiciantur.</emphasis>
Panicula
<emphasis id="1FDE767C4EA566745F1FB71232A9C253" italics="true">rarior, fere terminalis ex pedunculis bis dichotomis.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

148
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<mods:title>Three additional new genera of acidocerine water scavenger beetles from the Guiana and Brazilian Shield regions of South America (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
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<mods:namePart>Giron, Jennifer C.</mods:namePart>
<mods:affiliation>Department of Ecology &amp; Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
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<mods:nameIdentifier type="email">entiminae@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Short, Andrew Edward Z.</mods:namePart>
<mods:affiliation>Department of Ecology &amp; Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
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<treatment ID-GBIF-Taxon="158217538" LSID="urn:lsid:zoobank.org:act:46ABEE79-02F3-4D51-BCD1-6D9BB5717CAA" httpUri="http://treatment.plazi.org/id/62062A63B719E9DFB8FAD0091E3414FE" lastPageNumber="144" pageId="35" pageNumber="144">
<subSubSection pageId="35" pageNumber="144" type="nomenclature">
<paragraph pageId="35" pageNumber="144">
<taxonomicName LSID="62062A63-B719-E9DF-B8FA-D0091E3414FE" authority="Girón &amp; Short, 2019" authorityName="Girón &amp; Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus petilus" order="Coleoptera" pageId="35" pageNumber="144" phylum="Arthropoda" rank="species" species="petilus" status="sp. nov.">Primocerus petilus</taxonomicName>
<taxonomicNameLabel pageId="35" pageNumber="144">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of Primocerus spp.: A-C Primocerus petilus: A dorsal view B lateral view C ventral view. D-F Primocerus striatolatus: D dorsal view E lateral view F ventral view. G-I Primocerus semipubescens: G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure13" httpUri="https://binary.pensoft.net/fig/310070" pageId="35" pageNumber="144">
Figs 13
<normalizedToken originalValue="A–C">A-C</normalizedToken>
</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="35" pageNumber="144">, 14J</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of Primocerus spp." figureDoi="10.3897/zookeys.855.33013.figure15" httpUri="https://binary.pensoft.net/fig/310072" pageId="35" pageNumber="144">, 15B</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Habitat of Primocerus spp. A habitat and type locality for P. cuspidis, Venezuela, Tobogan de la Selva, collecting event AS- 08 - 080 b B habitat and type locality for P. pijiguaense, Venezuela, Los Pijiguaos, collecting event AS- 07 - 015 C habitat and type locality for P. neutrum, Venezuela, along La Escalera, collecting event AS- 08 - 058 D habitat and type locality for P. petilus, Brazil, Vale do Paraiso, collecting event BR 18 - 0203 - 01 G." figureDoi="10.3897/zookeys.855.33013.figure16" httpUri="https://binary.pensoft.net/fig/310073" pageId="35" pageNumber="144">, 16D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="144" type="type material">
<paragraph pageId="35" pageNumber="144">Type material.</paragraph>
<paragraph pageId="35" pageNumber="144">
<emphasis bold="true" pageId="35" pageNumber="144">Holotype</emphasis>
(♂): &quot;BRAZIL:
<normalizedToken originalValue="Pará">Para</normalizedToken>
: Alenquer/
<geoCoordinate degrees="1.49292" direction="south" orientation="latitude" precision="1" value="-1.49292">1.49292S</geoCoordinate>
,
<geoCoordinate degrees="54.51566" direction="west" orientation="longitude" precision="1" value="-54.51566">54.51566W</geoCoordinate>
; 150 m/ Vale do
<normalizedToken originalValue="Paraíso">Paraiso</normalizedToken>
, ca. 55 km N. of Alenquer/ tiny wet rock/seepage on trail; 3.ii.2018/ leg. A. Short; BR18-0203-01G&quot; (INPA, DNA voucher specimen SLE 1498).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="144" type="differential diagnosis">
<paragraph pageId="35" pageNumber="144">Differential diagnosis.</paragraph>
<paragraph pageId="35" pageNumber="144">
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus petilus" order="Coleoptera" pageId="35" pageNumber="144" phylum="Arthropoda" rank="species" species="petilus">
<emphasis italics="true" pageId="35" pageNumber="144">Primocerus petilus</emphasis>
</taxonomicName>
can be recognized by the presence of sharply impressed elytral punctures, with serial punctures well differentiated (larger and deeper than remainder punctures), longitudinally aligned to form elytral striae. It is similar to
<taxonomicName lsidName="P. semipubescens" pageId="35" pageNumber="144" rank="species" species="semipubescens">
<emphasis italics="true" pageId="35" pageNumber="144">P. semipubescens</emphasis>
</taxonomicName>
, from which it can be differentiated by the hydrofuge pubescence of the metafemora covering basal 3/4 of the anterior surface (covering less than basal half in
<taxonomicName lsidName="P. semipubescens" pageId="35" pageNumber="144" rank="species" species="semipubescens">
<emphasis italics="true" pageId="35" pageNumber="144">P. semipubescens</emphasis>
</taxonomicName>
). It is also very similar to
<taxonomicName lsidName="P. striatolatus" pageId="35" pageNumber="144" rank="species" species="striatolatus">
<emphasis italics="true" pageId="35" pageNumber="144">P. striatolatus</emphasis>
</taxonomicName>
, from which it can be differentiated by the undefined elytral striae IX and X along the basal fourth of the elytra (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of Primocerus spp.: A-C Primocerus petilus: A dorsal view B lateral view C ventral view. D-F Primocerus striatolatus: D dorsal view E lateral view F ventral view. G-I Primocerus semipubescens: G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure13" httpUri="https://binary.pensoft.net/fig/310070" pageId="35" pageNumber="144">13B</figureCitation>
; elytral striae IX and X clearly impressed along their entire length in
<taxonomicName lsidName="P. striatolatus" pageId="35" pageNumber="144" rank="species" species="striatolatus">
<emphasis italics="true" pageId="35" pageNumber="144">P. striatolatus</emphasis>
</taxonomicName>
, Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of Primocerus spp.: A-C Primocerus petilus: A dorsal view B lateral view C ventral view. D-F Primocerus striatolatus: D dorsal view E lateral view F ventral view. G-I Primocerus semipubescens: G dorsal view H lateral view I ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure13" httpUri="https://binary.pensoft.net/fig/310070" pageId="35" pageNumber="144">13E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="144" type="description">
<paragraph pageId="35" pageNumber="144">Description.</paragraph>
<paragraph pageId="35" pageNumber="144">
Body length 3.4 mm, width 1.6 mm. Body elongate oval, moderately convex (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Primocerus spp.: A-D Primocerus maipure: A dorsal view B lateral view C ventral view D head, dorsal view. E-H Primocerus pijiguaense: E dorsal view F lateral view G ventral view H head, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.855.33013.figure12" httpUri="https://binary.pensoft.net/fig/310069" pageId="35" pageNumber="144">12A, B</figureCitation>
). General coloration dark brown. Elytra with ground punctures sharply marked, and well-defined rows of serial punctures (forming elytral striae); elytral striae very slightly impressed along posterior half of elytra. Posterior elevation of mesoventrite with simple, curved transverse ridge. Metafemora with hydrofuge pubescence covering basal 4/5 of anterior surface. Apex of fifth abdominal ventrite rounded. Aedeagus (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Thorax, abdomen and aedeagus of Primocerus spp.: A-D Primocerus neutrum: A ventral view of mesoventrite (white arrow pointing transverse ridge) B posterior view of metafurca C fifth abdominal ventrite D aedeagus. E-L aedeagus: E, F Primocerus maipure: E dorsal view F lateral view. G, H Primocerus pijiguaense: G dorsal view H lateral view I Primocerus gigas J Primocerus petilus K Primocerus striatolatus L Primocerus cuspidis. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.855.33013.figure14" httpUri="https://binary.pensoft.net/fig/310071" pageId="35" pageNumber="144">14J</figureCitation>
) with basal piece nearly 1.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than parameres; parameres nearly as long as median lobe (median lobe inserted further into basal piece, thus appearing shorter than parameres); apex of parameres narrowly rounded; apex of median lobe widely rounded.
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="144" type="etymology">
<paragraph pageId="35" pageNumber="144">Etymology.</paragraph>
<paragraph pageId="35" pageNumber="144">
Named with the Latin word
<taxonomicName lsidName="petilus" pageId="35" pageNumber="144" rank="species" species="petilus">
<emphasis italics="true" pageId="35" pageNumber="144">petilus</emphasis>
</taxonomicName>
meaning slender, in reference to the relative slenderness of the body in this species.
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="144" type="distribution">
<paragraph pageId="35" pageNumber="144">Distribution.</paragraph>
<paragraph pageId="35" pageNumber="144">
<taxonomicName authorityName="Girón &amp; Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus petilus" order="Coleoptera" pageId="35" pageNumber="144" phylum="Arthropoda" rank="species" species="petilus">
<emphasis italics="true" pageId="35" pageNumber="144">Primocerus petilus</emphasis>
</taxonomicName>
has only been collected at one locality in the north of Brazil, at an elevation of 150 m (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of Primocerus spp." figureDoi="10.3897/zookeys.855.33013.figure15" httpUri="https://binary.pensoft.net/fig/310072" pageId="35" pageNumber="144">15B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="144" type="remarks">
<paragraph pageId="35" pageNumber="144">Remarks.</paragraph>
<paragraph pageId="35" pageNumber="144">
The single known specimen is missing the maxillary palps. It was collected on a temporary wet spot on an exposed forested rock outcrop. The rock was wet when the specimen was collected due to recent rains but was dry by the following day (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Habitat of Primocerus spp. A habitat and type locality for P. cuspidis, Venezuela, Tobogan de la Selva, collecting event AS- 08 - 080 b B habitat and type locality for P. pijiguaense, Venezuela, Los Pijiguaos, collecting event AS- 07 - 015 C habitat and type locality for P. neutrum, Venezuela, along La Escalera, collecting event AS- 08 - 058 D habitat and type locality for P. petilus, Brazil, Vale do Paraiso, collecting event BR 18 - 0203 - 01 G." figureDoi="10.3897/zookeys.855.33013.figure16" httpUri="https://binary.pensoft.net/fig/310073" pageId="35" pageNumber="144">16D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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</mods:name>
<mods:name id="74AECB5C221D607656AEFBC86644EFE8" type="personal">
<mods:role id="5F24B7296C2A14E67D7BA596C1C2A145">
<mods:roleTerm id="33AB8752260DEE3FE8EC257931D17CF0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0AEB2A2483E7E8D689FC25C70FA71310">Pérez, Toni</mods:namePart>
</mods:name>
<mods:typeOfResource id="91EA405C28FD2C056D46D2481F1175E9">text</mods:typeOfResource>
<mods:relatedItem id="30997A5F0B244E778DC8870476A09E40" type="host">
<mods:titleInfo id="3ADA6F76908629E4E1FB1B0D68E931F3">
<mods:title id="17BA35F087BF2AF8580770C7CCEC4D26">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4EC85216D8890B9E7E32F84BE60DC3D6">
<mods:date id="B7C7894F9194CEED64CBFBCE164D8EB5">2013</mods:date>
<mods:detail id="9D20C6024626A76EDF30E1427786BB9C" type="volume">
<mods:number id="7DD17FE484AB2972A9C2078772EB4704">3700</mods:number>
</mods:detail>
<mods:detail id="3FC10BE28D5BCCB822C988EE7021B93C" type="issue">
<mods:number id="0FB2CDD05686499E34B0F7E4BF8D3F62">2</mods:number>
</mods:detail>
<mods:extent id="E044857C5A46CF89C8191DFAF5C63D51" unit="page">
<mods:start id="75B5D7E69F956B5594A5E0B1084202FB">201</mods:start>
<mods:end id="3055C4A8EF0D35EC6F80B2AF9DAA64F3">225</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="53D8EEDB29D20C5F46FF63552E6B7516">journal article</mods:classification>
<mods:identifier id="5E097FFC4766A413D2E6B1CBC46F7D99" type="DOI">10.11646/zootaxa.3700.2.1</mods:identifier>
<mods:identifier id="1B7BF2A79576C6E3A12AC4076325ADE6" type="GBIF-Dataset">3d28eeb6-88a2-4fb2-aebd-796c8b2c2287</mods:identifier>
<mods:identifier id="8D3A422572C664528F3B855763219E15" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5EA4D31DE4E0EE25F02BFB7E571CEAEA" type="Zenodo-Dep">218268</mods:identifier>
<mods:identifier id="9EE5E8F7163027F49844D46A6215B19F" type="ZooBank">8119DAED-D7AF-4858-BC63-F082151A4DAE</mods:identifier>
</mods:mods>
<treatment id="620687CFFFE2FF93EED61BF5FEF71079" ID-DOI="http://doi.org/10.5281/zenodo.6149880" ID-GBIF-Taxon="119574447" ID-Zenodo-Dep="6149880" LSID="urn:lsid:plazi:treatment:620687CFFFE2FF93EED61BF5FEF71079" httpUri="http://treatment.plazi.org/id/620687CFFFE2FF93EED61BF5FEF71079" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203">
<subSubSection id="A2B56552FFE2FF92EED61BF5FD4310F6" box="[151,707,1182,1209]" pageId="2" pageNumber="203" type="nomenclature">
<paragraph id="EA1036D9FFE2FF92EED61BF5FD4310F6" blockId="2.[151,707,1182,1209]" box="[151,707,1182,1209]" pageId="2" pageNumber="203">
<heading id="B15881B5FFE2FF92EED61BF5FD4310F6" bold="true" box="[151,707,1182,1209]" fontSize="11" level="1" pageId="2" pageNumber="203" reason="1">
<emphasis id="D8DBEACBFFE2FF92EED61BF5FD4310F6" bold="true" box="[151,707,1182,1209]" pageId="2" pageNumber="203">
<emphasis id="D8DBEACBFFE2FF92EED61BF5FECC10F7" bold="true" box="[151,332,1182,1208]" italics="true" pageId="2" pageNumber="203">
<taxonomicName id="2DAF4D5AFFE2FF92EED61BF5FE9610F7" ID-CoL="62LNY" box="[151,278,1182,1208]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="203" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
</emphasis>
<taxonomicName id="2DAF4D5AFFE2FF92EF121BF5FD4310F6" authority="(Preyssler, 1790)" authorityName="Preyssler" authorityYear="1790" baseAuthorityName="Preyssler" baseAuthorityYear="1790" box="[339,707,1182,1209]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="203" phylum="Arthropoda" rank="species" species="tetrachelatus">
<emphasis id="D8DBEACBFFE2FF92EF121BF5FE6F10F7" bold="true" box="[339,495,1182,1208]" italics="true" pageId="2" pageNumber="203">tetrachelatus</emphasis>
(Preyssler, 1790)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE2FF92EED61B8EFB87116D" pageId="2" pageNumber="203" type="materials_examined">
<paragraph id="EA1036D9FFE2FF92EED61B8EFB87116D" blockId="2.[151,1436,1253,2033]" pageId="2" pageNumber="203">
<emphasis id="D8DBEACBFFE2FF92EED61B8EFE8610B1" bold="true" box="[151,262,1253,1278]" pageId="2" pageNumber="203">Material.</emphasis>
<collectingCountry id="92B87649FFE2FF92EF5B1B8EFEDD10B1" box="[282,349,1253,1278]" name="Spain" pageId="2" pageNumber="203">Spain</collectingCountry>
, Andalusia, Jaén province,
<taxonomicName id="2DAF4D5AFFE2FF92EC851B8EFCCE10B1" box="[708,846,1253,1278]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="villacarrillo" status="sp. nov.">Villacarrillo</taxonomicName>
, Cueva de la Fuente del Tejo (38º3′24.32″N, 2º54′27.96″W;
<quantity id="2D579B3CFFE2FF92EF021A61FE1E116D" box="[323,414,1289,1314]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="2" pageNumber="203" unit="m" value="1490.0">1490 m</quantity>
.a.s.l.),
<date id="9E111019FFE2FF92EFAB1A61FDF6116D" box="[490,630,1290,1314]" pageId="2" pageNumber="203" value="2008-07-22">22.VII.2008</date>
, lgt. G.E.V.,
<specimenCount id="FCA9FD50FFE2FF92ED431A61FCAE116E" box="[770,814,1290,1314]" pageId="2" pageNumber="203" type="generic">4 ♂</specimenCount>
, 1 ♀, 1 T (DEUA).
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE2FF93EE861A46FCDD176F" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203" type="description">
<paragraph id="EA1036D9FFE2FF92EE861A46FB351109" blockId="2.[151,1436,1253,2033]" box="[199,1205,1325,1350]" pageId="2" pageNumber="203">
<emphasis id="D8DBEACBFFE2FF92EE861A46FED91109" bold="true" box="[199,345,1325,1350]" pageId="2" pageNumber="203">Description.</emphasis>
Epigean species, integument pigmented, without troglomorphic adaptations.
</paragraph>
<paragraph id="EA1036D9FFE2FF92EE861A39FDF21252" blockId="2.[151,1436,1253,2033]" pageId="2" pageNumber="203">
Carapace subquadrate, slightly longer than broad; anterior margin medially denticulate and weakly prominent; four eyes with lens and reflective tapeta, anterior eyes with convex lens, posterior eyes with weak lens; diameter of anterior eyes
<quantity id="2D579B3CFFE2FF92EF6F1AF2FE6211FE" box="[302,482,1433,1458]" metricMagnitude="-5" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="2" pageNumber="203" unit="mm" value="0.0475" valueMax="0.05" valueMin="0.045">0.045–0.05 mm</quantity>
, distance from anterior eyes to anterior margin of carapace
<quantity id="2D579B3CFFE2FF92EAC11AF2FAB411FE" box="[1152,1332,1433,1458]" metricMagnitude="-5" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="2" pageNumber="203" unit="mm" value="0.0325" valueMax="0.035" valueMin="0.03">0.03–0.035 mm</quantity>
, distance from anterior to posterior eyes
<quantity id="2D579B3CFFE2FF92EC401AD6FD37119A" box="[513,695,1469,1494]" metricMagnitude="-5" metricUnit="m" metricValue="4.249999999999999" metricValueMax="4.5" metricValueMin="4.0" pageId="2" pageNumber="203" unit="mm" value="0.042499999999999996" valueMax="0.045" valueMin="0.04">0.04–0.045 mm</quantity>
; chaetotaxy: 18–19 macrosetae, formula: m
<quantity id="2D579B3CFFE2FF92EAF21AD6FB5D1199" box="[1203,1245,1469,1494]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="203" unit="m" value="4.0">4m</quantity>
:6:4:2:
<quantity id="2D579B3CFFE2FF92EB5A1AD6FACA1199" box="[1307,1354,1469,1494]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="203" unit="in" value="2.0">2 in</quantity>
males, 4:6:4:2:
<quantity id="2D579B3CFFE2FF92EEAC1A8AFE9A11B5" box="[237,282,1505,1530]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="203" unit="in" value="3.0">3 in</quantity>
female, anteromedial seta
<quantity id="2D579B3CFFE2FF92EC0B1A89FD2D11B6" box="[586,685,1506,1530]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="203" unit="mm" value="0.11">0.11 mm</quantity>
long, ocular seta
<quantity id="2D579B3CFFE2FF92ED371A89FC5E11B6" box="[886,990,1506,1530]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="203" unit="mm" value="0.08">0.08 mm</quantity>
; Four or five lyrifissures in the anterior and ocular areas, two in the posterior area.
</paragraph>
<paragraph id="EA1036D9FFE2FF92EE861942FB411229" blockId="2.[151,1436,1253,2033]" pageId="2" pageNumber="203">
Pedipalpal coxa with 5 setae (including 2 on manducatory process), coxa
<date id="9E111019FFE2FF92EDBA1941FB99120D" box="[1019,1049,1578,1602]" pageId="2" pageNumber="203">I 3</date>
+ 3 marginal microsetae,
<date id="9E111019FFE2FF92EB7D1941FAE3120D" box="[1340,1379,1578,1602]" pageId="2" pageNumber="203">II 4</date>
+ 8– 9 bipinnate coxal spines,
<date id="9E111019FFE2FF92EFF21925FE641229" box="[435,484,1613,1638]" pageId="2" pageNumber="203">III 5</date>
+ 4 bipinnate coxal spines and
<date id="9E111019FFE2FF92ED071925FCFA1229" box="[838,890,1613,1638]" pageId="2" pageNumber="203">IV 6</date>
; intercoxal tubercle bisetose.
</paragraph>
<paragraph id="EA1036D9FFE2FF92EE86191AFDF812BA" blockId="2.[151,1436,1253,2033]" pageId="2" pageNumber="203">Chaetotaxy of tergites I–XI: 4:4:4:4:6:6:6:6:6:4:1T2T1. Genital area with 10 setae on sternite II in male and female; sternite III with (3)8(3) setae in females; sternite III in males with genital opening flanked by 6–8 setae on each side, 4+4 internal glandular setae and (3)10(3) marginal setae. Chaetotaxy of sternites IV–XI: (2)7(2):7:6:6:6:6:7:0. Anal cone 0+2 setae.</paragraph>
<paragraph id="EA1036D9FFE2FF92EE86186AFEF313E5" blockId="2.[151,1436,1253,2033]" pageId="2" pageNumber="203">
Chelicera with 6 setae on palm and 1 microseta laterally; seta
<emphasis id="D8DBEACBFFE2FF92ED3C186AFC131355" box="[893,915,1793,1818]" italics="true" pageId="2" pageNumber="203">gl</emphasis>
0.58–0.61 from base of movable finger. Fixed finger with 10–11 teeth, proximally decreasing in size. Movable finger without an isolated subapical tooth (
<emphasis id="D8DBEACBFFE2FF92EB20184EFAF71371" box="[1377,1399,1829,1854]" italics="true" pageId="2" pageNumber="203">di</emphasis>
), a large, single or bicuspid distal tooth and 5–6 teeth reduced in size proximally; spinneret low prominent and apically rounded, markedly more developed in female. Rallum with 11 blades, serrulae exterior and interior with 14 and 13 blades respectively.
</paragraph>
<paragraph id="EA1036D9FFE2FF93EE8618DEFD921512" blockId="2.[151,1436,1253,2033]" lastBlockId="3.[151,1437,151,1078]" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203">
Pedipalp with femoral chaetotaxy: 3:6:2:5:1, four lyrifissures: one antiaxial and three paraxial. Chelal hand very weakly depressed at level of
<emphasis id="D8DBEACBFFE2FF92EC5518B3FDD413BE" box="[532,596,2008,2033]" italics="true" pageId="2" pageNumber="203">ib-isb</emphasis>
, weak hump distad of
<emphasis id="D8DBEACBFFE2FF92ED1318B3FC1213BE" box="[850,914,2008,2033]" italics="true" pageId="2" pageNumber="203">ib-isb</emphasis>
. Fixed finger with 18–20 pointed teeth, one or two distal ones small, 5–6 basal teeth progressively decreasing in size, all with dental canals, 3–4 microtubercles at base; tip of finger with an accessory tooth (
<emphasis id="D8DBEACBFFE3FF93ECD11FD1FD26149C" box="[656,678,186,211]" italics="true" pageId="3" pageNumber="204">td</emphasis>
) on antiaxial face; four teeth at level of
<emphasis id="D8DBEACBFFE3FF93EA3E1FD0FB35149D" box="[1151,1205,187,210]" italics="true" pageId="3" pageNumber="204">est/it</emphasis>
occupying
<quantity id="2D579B3CFFE3FF93EB7C1FD0FA17149D" box="[1341,1431,187,211]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="204" unit="mm" value="0.1">0.1 mm</quantity>
, distance between apices
<quantity id="2D579B3CFFE3FF93EFF31FB5FDAA14BA" box="[434,554,222,246]" metricMagnitude="-5" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="204" unit="mm" value="0.033">0.033 mm</quantity>
. Distal half of movable finger with 7–8 pointed teeth with dental canals, the distal one small; proximal half without marginal lamina, with 7 low vestigial teeth reaching trichobothrium
<emphasis id="D8DBEACBFFE3FF93EB051F94FADD1557" box="[1348,1373,255,280]" italics="true" pageId="3" pageNumber="204">sb</emphasis>
, with dental canal; coupled sensilla
<emphasis id="D8DBEACBFFE3FF93EFA71E4FFD811574" box="[486,513,292,315]" italics="true" pageId="3" pageNumber="204">pc</emphasis>
between
<emphasis id="D8DBEACBFFE3FF93EC2D1E49FD051574" box="[620,645,290,315]" italics="true" pageId="3" pageNumber="204">sb</emphasis>
and
<emphasis id="D8DBEACBFFE3FF93ECFA1E49FD491574" box="[699,713,290,315]" italics="true" pageId="3" pageNumber="204">b</emphasis>
, slightly closer to
<emphasis id="D8DBEACBFFE3FF93EDD81E49FC321574" box="[921,946,290,315]" italics="true" pageId="3" pageNumber="204">sb</emphasis>
; basal apodeme long and apically indented. Trichobothrium
<emphasis id="D8DBEACBFFE3FF93EF0F1E2DFEE91512" box="[334,361,326,349]" italics="true" pageId="3" pageNumber="204">ist</emphasis>
level with
<emphasis id="D8DBEACBFFE3FF93EFA71E2EFD8B1511" box="[486,523,325,350]" italics="true" pageId="3" pageNumber="204">esb</emphasis>
.
</paragraph>
<paragraph id="EA1036D9FFE3FF93EE861E03FBAB15CF" blockId="3.[151,1437,151,1078]" box="[199,1067,359,384]" pageId="3" pageNumber="204">Leg IV. Tactile seta ratios: tibia 0.58–0.6, basitarsus 0.37–0.4 and tarsus 0.32.</paragraph>
<paragraph id="EA1036D9FFE3FF93EE861EE7FB78161F" blockId="3.[151,1437,151,1078]" pageId="3" pageNumber="204">
<emphasis id="D8DBEACBFFE3FF93EE861EE7FE7915EC" box="[199,505,394,419]" italics="true" pageId="3" pageNumber="204">Measurements and ratios.</emphasis>
Male. Body 1.82. Carapace 0.47/0.46 (1.0). Chelicera: palm 0.40/0.19 (2.2), movable finger 0.21. Pedipalp: femur 0.69/0.11 (6.5); patella 0.25/0.13 (2.0); chela 0.95/0.17 (5.5), depth 0.18 (5.2); hand 0.38 (2.2), length/depth (2.1); movable finger 0.58; ratio movable finger/hand 1.5; ratio femur/movable finger 1.2; ratio femur/carapace 1.5; ratio chela/carapace 2.0. Leg I: femur 0.41/0.07 (5.7); patella 0.20/0.07 (3.1); tibia 0.24/0.05 (4.7); tarsus 0.40/0.04 (9.4); ratio femur/patella 2.1. Leg IV: femur+patella 0.60/0.24 (2.6); tibia 0.40/0.09 (4.7); basitarsus 0.23/0.07 (3.5); telotarsus 0.42/0.04 (11.3); ratio telotarsus/basitarsus 1.8.
</paragraph>
<paragraph id="EA1036D9FFE3FF93EE861D31FCDD176F" blockId="3.[151,1437,151,1078]" pageId="3" pageNumber="204">
<emphasis id="D8DBEACBFFE3FF93EE861D31FE9B163C" box="[199,283,602,627]" italics="true" pageId="3" pageNumber="204">Female</emphasis>
. Body 1.83. Carapace 0.51/0.5 (1.0). Chelicera: palm 0.44/0.21 (2.1), movable finger 0.22. Pedipalp: femur 0.73/0.13 (5.8); patella 0.30/0.15 (2.0); chela 1.02/0.21 (4.7), depth 0.22 (4.5); hand 0.43 (2.0), length/depth (1.9); movable finger 0.59; ratio movable finger/hand 1.4; ratio femur/movable finger 1.2; ratio femur/carapace 1.4; ratio chela/carapace 2.0. Leg I: femur 0.44/0.08 (5.5); patella 0.21/0.07 (3.0); tibia 0.25/0.05 (4.9); tarsus 0.45/ 0.05 (9.9); ratio femur/patella 2.1. Leg IV: femur+patella 0.65/0.26 (2.5); tibia 0.41/0.09 (4.6); basitarsus 0.23/0.07 (3.3); telotarsus 0.44/0.04 (11.0); ratio telotarsus/basitarsus 1.9.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE3FF93EE861C41FD2D17E4" pageId="3" pageNumber="204" type="distribution">
<paragraph id="EA1036D9FFE3FF93EE861C41FD2D17E4" blockId="3.[151,1437,151,1078]" pageId="3" pageNumber="204">
<emphasis id="D8DBEACBFFE3FF93EE861C41FEE0170C" bold="true" box="[199,352,810,835]" pageId="3" pageNumber="204">Distribution.</emphasis>
Western Palaearctic Region, chiefly Mediterranean; introduced to eastern
<collectingCountry id="92B87649FFE3FF93EAA41C41FABC170C" box="[1253,1340,810,835]" name="Canada" pageId="3" pageNumber="204">Canada</collectingCountry>
,
<collectingCountry id="92B87649FFE3FF93EB0E1C40FA1C170C" box="[1359,1436,811,835]" name="United States of America" pageId="3" pageNumber="204">U.S.A.</collectingCountry>
(including Hawaii),
<collectingCountry id="92B87649FFE3FF93EF3F1C26FE3C1729" box="[382,444,845,870]" name="Cuba" pageId="3" pageNumber="204">Cuba</collectingCountry>
,
<collectingCountry id="92B87649FFE3FF93EF881C26FDBB1729" box="[457,571,845,870]" name="Argentina" pageId="3" pageNumber="204">Argentina</collectingCountry>
,
<collectingCountry id="92B87649FFE3FF93EC081C26FD411729" box="[585,705,845,870]" name="Seychelles" pageId="3" pageNumber="204">Seychelles</collectingCountry>
and southwestern
<collectingCountry id="92B87649FFE3FF93EDD81C26FC7F1729" box="[921,1023,845,870]" name="Australia" pageId="3" pageNumber="204">Australia</collectingCountry>
(Gardini 2013; Harvey 2011). New record for the province of Jaén, the second after Málaga for the Andalusian region; detailed locations for the Iberian Peninsula are given in Zaragoza (2007).
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE3FF93EE861CDEFEF71079" pageId="3" pageNumber="204" type="discussion">
<paragraph id="EA1036D9FFE3FF93EE861CDEFEF71079" blockId="3.[151,1437,151,1078]" pageId="3" pageNumber="204">
<emphasis id="D8DBEACBFFE3FF93EE861CDEFEBB1781" bold="true" box="[199,315,949,974]" pageId="3" pageNumber="204">Remarks.</emphasis>
These specimens fall within the wide range of variation described for
<emphasis id="D8DBEACBFFE3FF93EAC61CDDFAF41781" box="[1159,1396,949,974]" italics="true" pageId="3" pageNumber="204">
C. (E.)
<taxonomicName id="2DAF4D5AFFE3FF93EAA31CDEFAF41781" box="[1250,1396,949,974]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="tetrachelatus">tetrachelatus</taxonomicName>
</emphasis>
by Gardini (2009, 2013). Some chelal measures lie at the extremes of the ranges or exceed them (e.g. ratio movable finger/hand). We agree with Gardini (2013) that “the heterogeneous morphology of
<taxonomicName id="2DAF4D5AFFE3FF93EA7F1C90FB70105C" box="[1086,1264,1018,1043]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="tetrachelatus">
<emphasis id="D8DBEACBFFE3FF93EA7F1C90FB70105C" box="[1086,1264,1018,1043]" italics="true" pageId="3" pageNumber="204">C. tetrachelatus</emphasis>
</taxonomicName>
probably hides a species complex”.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="8A3DE24969F991B10D1842DC780CED3D" ID-DOI="10.11646/zootaxa.3700.2.1" ID-GBIF-Dataset="3d28eeb6-88a2-4fb2-aebd-796c8b2c2287" ID-ISSN="1175-5326" ID-Zenodo-Dep="218268" ID-ZooBank="8119DAED-D7AF-4858-BC63-F082151A4DAE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460662093739" checkinUser="plazi" docAuthor="Zaragoza, Juan A. &amp; Pérez, Toni" docDate="2013" docId="620687CFFFE3FF94EED61BEFFE11124A" docLanguage="en" docName="zt03700p225.pdf" docOrigin="Zootaxa 3700 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chthonius" docType="treatment" docVersion="8" lastPageNumber="205" masterDocId="9E3FFFB7FFE0FF90EE411F6BFF80144F" masterDocTitle="Hypogean pseudoscorpions (Arachnida) from Jaén province (Andalusia, Spain), with descriptions of four new species and a new synonymy" masterLastPageNumber="225" masterPageNumber="201" pageNumber="204" updateTime="1698347634734" updateUser="plazi">
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<mods:title id="998910E6B4420D4A97E05DDF53269A75">Hypogean pseudoscorpions (Arachnida) from Jaén province (Andalusia, Spain), with descriptions of four new species and a new synonymy</mods:title>
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<mods:namePart id="F770FA9B18FA94F7A272C84BAE29033E">Zaragoza, Juan A.</mods:namePart>
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<mods:namePart id="40F95D098088810984081D8372664B25">Pérez, Toni</mods:namePart>
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<mods:date id="CDE3BF159FB2A6F9257DD787E6B8BEC5">2013</mods:date>
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<treatment id="620687CFFFE3FF94EED61BEFFE11124A" ID-DOI="http://doi.org/10.5281/zenodo.6149882" ID-GBIF-Taxon="119574444" ID-Zenodo-Dep="6149882" LSID="urn:lsid:plazi:treatment:620687CFFFE3FF94EED61BEFFE11124A" httpUri="http://treatment.plazi.org/id/620687CFFFE3FF94EED61BEFFE11124A" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204">
<subSubSection id="A2B56552FFE3FF93EED61BEFFB66114C" pageId="3" pageNumber="204" type="nomenclature">
<paragraph id="EA1036D9FFE3FF93EED61BEFFAA710D0" blockId="3.[151,1319,1156,1183]" box="[151,1319,1156,1183]" pageId="3" pageNumber="204">
<heading id="B15881B5FFE3FF93EED61BEFFAA710D0" bold="true" box="[151,1319,1156,1183]" fontSize="11" level="1" pageId="3" pageNumber="204" reason="1">
<emphasis id="D8DBEACBFFE3FF93EED61BEFFAA710D0" bold="true" box="[151,1319,1156,1183]" pageId="3" pageNumber="204">
<emphasis id="D8DBEACBFFE3FF93EED61BEFFE5810D1" bold="true" box="[151,472,1156,1182]" italics="true" pageId="3" pageNumber="204">
<taxonomicName id="2DAF4D5AFFE3FF93EED61BEFFE9610D1" ID-CoL="62LNY" box="[151,278,1156,1182]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="204" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
<taxonomicName id="2DAF4D5AFFE3FF93EF121BEFFE5810D1" authority="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez, 2001" authorityName="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez" authorityYear="2001" box="[339,472,1156,1182]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="cazorlensis">cazorlensis</taxonomicName>
</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2001
</emphasis>
</heading>
</paragraph>
<paragraph id="EA1036D9FFE3FF93EED61BA1FB66114C" blockId="3.[151,1437,1226,1352]" pageId="3" pageNumber="204">
<taxonomicName id="2DAF4D5AFFE3FF93EED61BA1FD8F10AF" box="[151,527,1226,1248]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="verai" subGenus="Ephippiochthonius">
<emphasis id="D8DBEACBFFE3FF93EED61BA1FD8F10AF" box="[151,527,1226,1248]" italics="true" pageId="3" pageNumber="204">Chthonius (Ephippiochthonius) verai</emphasis>
</taxonomicName>
(not Zaragoza, 1985): Pérez Fernández &amp; Tinaut Ránera 2005: 1, 4, fig. 1; Barranco
<emphasis id="D8DBEACBFFE3FF93EB2B1BA0FA1D10AF" box="[1386,1437,1226,1248]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2005: 78; Pérez Fernández 2008: 341; Pérez Fernández &amp; Tinaut Ránera 2009: 108 (misidentifications).
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE3FF93EED61A7BFA991107" pageId="3" pageNumber="204" type="description">
<paragraph id="EA1036D9FFE3FF93EED61A7BFA991107" blockId="3.[151,1437,1226,1352]" pageId="3" pageNumber="204">
<taxonomicName id="2DAF4D5AFFE3FF93EED61A7BFB9A1169" authority="Carabajal Marquez et al. 2001: 180" authorityName="Carabajal Marquez et al." authorityPageNumber="180" authorityYear="2001" box="[151,1050,1296,1318]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="204" phylum="Arthropoda" rank="subSpecies" species="ventalloi" subGenus="Ephippiochthonius" subSpecies="cazorlensis">
<emphasis id="D8DBEACBFFE3FF93EED61A7BFD2C1169" box="[151,684,1296,1318]" italics="true" pageId="3" pageNumber="204">Chthonius (Ephippiochthonius) ventalloi cazorlensis</emphasis>
Carabajal Márquez
<emphasis id="D8DBEACBFFE3FF93ED3B1A7AFC2E1169" box="[890,942,1296,1318]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2001: 180
</taxonomicName>
–183, figs 14–18, table IV.
<taxonomicName id="2DAF4D5AFFE3FF93EED61A59FC0A1107" authority="Carabajal Marquez et al. 2001" authorityName="Carabajal Marquez et al." authorityYear="2001" box="[151,906,1330,1352]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="cazorlensis" subGenus="Ephippiochthonius">
<emphasis id="D8DBEACBFFE3FF93EED61A59FDCD1107" box="[151,589,1330,1352]" italics="true" pageId="3" pageNumber="204">Chthonius (Ephippiochthonius) cazorlensis</emphasis>
Carabajal Márquez
<emphasis id="D8DBEACBFFE3FF93ED5D1A58FCCF1107" box="[796,847,1330,1352]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2001
</taxonomicName>
: Pérez &amp; Zaragoza 2010: 17–22, fig. 1.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE3FF93EED61A1CFC6311FC" pageId="3" pageNumber="204" type="materials_examined">
<paragraph id="EA1036D9FFE3FF93EED61A1CFC6311FC" blockId="3.[151,1437,1399,2014]" pageId="3" pageNumber="204">
<emphasis id="D8DBEACBFFE3FF93EED61A1CFE8611DF" bold="true" box="[151,262,1399,1424]" pageId="3" pageNumber="204">Material.</emphasis>
<collectingCountry id="92B87649FFE3FF93EF591A1CFEDB11DF" box="[280,347,1399,1424]" name="Spain" pageId="3" pageNumber="204">Spain</collectingCountry>
, Andalusia, Jaén province, Hornos de Segura, Sima de la Fractura II (38º11′11.58″N, 2º47′17.89″W;
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<quantity id="2D579B3CFFE3FF93EF021AF1FE1111FC" box="[323,401,1434,1459]" metricMagnitude="2" metricUnit="m" metricValue="9.65" pageId="3" pageNumber="204" unit="m" value="965.0">965 m</quantity>
.
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a.s.l.),
<date id="9E111019FFE3FF93EF9D1AF0FDD011FC" box="[476,592,1435,1459]" pageId="3" pageNumber="204" value="2009-05-31">31.V.2009</date>
, lgt. G.E.V.,
<specimenCount id="FCA9FD50FFE3FF93EC9F1AF0FC8911FD" box="[734,777,1435,1459]" pageId="3" pageNumber="204" type="generic">4 ♂</specimenCount>
, 2 ♀, 1 T (DEUA).
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE3FF93EE861AD6FC7B12EA" pageId="3" pageNumber="204" type="diagnosis">
<paragraph id="EA1036D9FFE3FF93EE861AD6FC7B12EA" blockId="3.[151,1437,1399,2014]" pageId="3" pageNumber="204">
<emphasis id="D8DBEACBFFE3FF93EE861AD6FEC11199" bold="true" box="[199,321,1469,1494]" pageId="3" pageNumber="204">Diagnosis.</emphasis>
Weakly troglomorphic facies. Without eyes or eye-spots. Chelicera with 6 setae on palm and 0–2 microsetae laterally; movable finger without isolated subapical tooth; spinneret strongly reduced in males, prominent in females. Carapace with two macrosetae on posterior margin, preocular microsetae absent. Pedipalp: femur length in males 0.68–0.78 (ratio 6.3–7.1), females 0.72–0.80 (6.3–7.1×); chela length in males 0.93–1.03 (length/depth 6.2–6.6), females 1.06–1.08 (length/depth 5.9–6.1); fixed finger with 17–22 teeth, movable finger with 14–17 teeth with dental canals reaching distad
<emphasis id="D8DBEACBFFE3FF93ECB01901FC8A12CC" box="[753,778,1642,1667]" italics="true" pageId="3" pageNumber="204">sb</emphasis>
, closer to
<emphasis id="D8DBEACBFFE3FF93EDC71901FC1F12CC" box="[902,927,1642,1667]" italics="true" pageId="3" pageNumber="204">sb</emphasis>
than to
<emphasis id="D8DBEACBFFE3FF93EA431907FB9512CC" box="[1026,1045,1644,1667]" italics="true" pageId="3" pageNumber="204">st</emphasis>
, proximal raised lamina with 3–7 rounded vestigial teeth without canal; trichobothrium
<emphasis id="D8DBEACBFFE3FF93ECB319E5FC8D12EA" box="[754,781,1678,1701]" italics="true" pageId="3" pageNumber="204">ist</emphasis>
markedly distad
<emphasis id="D8DBEACBFFE3FF93ED8F19E6FC7312E9" box="[974,1011,1677,1702]" italics="true" pageId="3" pageNumber="204">esb</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE3FF94EE8619C4FE58107D" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204" type="description">
<paragraph id="EA1036D9FFE3FF93EE8619C4FE9C12A4" blockId="3.[151,1437,1399,2014]" pageId="3" pageNumber="204">
<emphasis id="D8DBEACBFFE3FF93EE8619C4FED91287" bold="true" box="[199,345,1711,1736]" pageId="3" pageNumber="204">Description.</emphasis>
<emphasis id="D8DBEACBFFE3FF93EF2319DBFCA71287" box="[354,807,1711,1736]" italics="true" pageId="3" pageNumber="204">Specimens from Sima de la Fractura II</emphasis>
. Weak troglomorphic facies; integument depigmented, pale brown.
</paragraph>
<paragraph id="EA1036D9FFE3FF93EE86199DFC5A131D" blockId="3.[151,1437,1399,2014]" pageId="3" pageNumber="204">
Carapace slightly longer than broad, slightly constricted posteriorly, subquadrate; anterior margin prominent and medially dentate, without epistome; without eyes or eye-spots; 18 setae, without preocular microsetae, formula 4:6:4:2:2, anteromedial seta
<quantity id="2D579B3CFFE3FF93EF971850FDF8131D" box="[470,632,1851,1875]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="3" pageNumber="204" unit="mm" value="0.12" valueMax="0.13" valueMin="0.11">0.11–0.13 mm</quantity>
long, ocular seta
<quantity id="2D579B3CFFE3FF93ED7C1850FC55131D" box="[829,981,1851,1875]" metricMagnitude="-5" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7" pageId="3" pageNumber="204" unit="mm" value="0.085" valueMax="0.1" valueMin="0.07">0.07–0.1 mm</quantity>
.
</paragraph>
<paragraph id="EA1036D9FFE3FF93EE861836FB1F13D7" blockId="3.[151,1437,1399,2014]" pageId="3" pageNumber="204">
Pedipalpal coxa 5 setae (including 2 on manducatory process), coxa
<date id="9E111019FFE3FF93ED931835FC711339" box="[978,1009,1886,1910]" pageId="3" pageNumber="204">I 3</date>
+ 3 marginal microsetae,
<date id="9E111019FFE3FF93EB5A1835FAC41339" box="[1307,1348,1886,1910]" pageId="3" pageNumber="204">II 4</date>
+ 7–12 bipinnate coxal spines,
<date id="9E111019FFE3FF93EFDF18EBFE4F13D7" box="[414,463,1919,1944]" pageId="3" pageNumber="204">III 5</date>
+ 3–6 bipinnate coxal spines,
<date id="9E111019FFE3FF93ED6518EBFCD913D7" box="[804,857,1919,1944]" pageId="3" pageNumber="204">IV 6</date>
; intercoxal tubercle bisetose.
</paragraph>
<paragraph id="EA1036D9FFE3FF94EE8618C9FD10149D" blockId="3.[151,1437,1399,2014]" lastBlockId="4.[151,1436,151,1542]" lastPageId="4" lastPageNumber="205" pageId="3" pageNumber="204">
Chaetotaxy of tergites
<date id="9E111019FFE3FF93EF8518C8FD9213F4" box="[452,530,1954,1979]" pageId="3" pageNumber="204">I–XI 4</date>
:4:4:4:6:6:6:6:6:4:1T2T1. Genital area with 9–10 setae on sternite II in males and
<quantity id="2D579B3CFFE3FF93EED618ADFF431391" box="[151,195,1989,2014]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="204" unit="in" value="9.0">9 in</quantity>
females; sternite III with (3)7(3) setae in females; sternite III in males with genital opening flanked by 6–10 setae on each side, 4+4 internal glandular setae and (3)9(3) marginal setae. Chaetotaxy of sternite IV–XI: (2)6– 7(2):7–8:6:6:6:6:6–8:0. Anal cone 0+2 setae.
</paragraph>
<paragraph id="EA1036D9FFE4FF94EE861FB6FE1315C9" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="205">
Chelicera with 6 setae on palm and one microseta laterally (one female with 1–2 microsetae); seta
<emphasis id="D8DBEACBFFE4FF94EB4E1FB6FAA514B9" box="[1295,1317,221,246]" italics="true" pageId="4" pageNumber="205">gl</emphasis>
0.53–0.56 from base of movable finger. Fixed finger with 11 teeth, two distal teeth distinctly larger than others, proximally reduced in size, three denticles at base. Movable finger without an isolated subapical tooth (
<emphasis id="D8DBEACBFFE4FF94EAE41E4EFB3B1571" box="[1189,1211,293,318]" italics="true" pageId="4" pageNumber="205">di</emphasis>
), with a large distal tooth and 7–8 contiguous teeth proximally decreasing in size, two denticles on base; spinneret strongly reduced in males, low in females.
</paragraph>
<paragraph id="EA1036D9FFE4FF94EE861EFAFBBB1686" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="205">
Pedipalp with femoral chaetotaxy 3:6–7:3:5:1, four lyrifissures: 1 antiaxial and 3 paraxial. Chelal hand markedly depressed at level of
<emphasis id="D8DBEACBFFE4FF94EFB41EDEFDB51581" box="[501,565,437,462]" italics="true" pageId="4" pageNumber="205">ib-isb</emphasis>
, with a short and prominent hump distad of
<emphasis id="D8DBEACBFFE4FF94EA681EDEFBE91581" box="[1065,1129,437,462]" italics="true" pageId="4" pageNumber="205">ib-isb</emphasis>
and distinct ventral hollow just before base of movable chelal finger. Fixed finger with 20–22 pointed teeth, all with dental canals except the two distal ones small; third distal tooth from tip a little wider than others; up to ten microtubercles at base; tip of finger with an accessory tooth (
<emphasis id="D8DBEACBFFE4FF94EC511D4AFDA61675" box="[528,550,545,570]" italics="true" pageId="4" pageNumber="205">td</emphasis>
) on antiaxial face; five teeth at level of
<emphasis id="D8DBEACBFFE4FF94EA511D49FBC71676" box="[1040,1095,546,569]" italics="true" pageId="4" pageNumber="205">est/it</emphasis>
occupying
<quantity id="2D579B3CFFE4FF94EA921D4AFAB01676" box="[1235,1328,545,569]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="205" unit="mm" value="0.1">0.1 mm</quantity>
, distance between apices
<quantity id="2D579B3CFFE4FF94EF091D2EFD8B1612" box="[328,523,581,606]" metricMagnitude="-5" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="4" pageNumber="205" unit="mm" value="0.0225" valueMax="0.025" valueMin="0.02">0.020–0.025 mm</quantity>
. Movable finger with 16–17 pointed teeth with dental canals, two distal ones tiny and two-three proximal ones reduced in size; proximally with slightly raised lamina with 4–7 rounded vestigial teeth, without dental canals, reaching to slightly proximad of trichobothrium
<emphasis id="D8DBEACBFFE4FF94EA4D1DE6FBA516E9" box="[1036,1061,653,678]" italics="true" pageId="4" pageNumber="205">sb</emphasis>
, level with end of dental row of fixed finger; five microtubercles at base. Trichobothrium
<emphasis id="D8DBEACBFFE4FF94ED5A1DD9FCB51686" box="[795,821,690,713]" italics="true" pageId="4" pageNumber="205">ist</emphasis>
markedly distal of
<emphasis id="D8DBEACBFFE4FF94EA4E1DDAFBB41685" box="[1039,1076,689,714]" italics="true" pageId="4" pageNumber="205">esb</emphasis>
.
</paragraph>
<paragraph id="EA1036D9FFE4FF94EE861DBDFD3E1731" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="205">
<emphasis id="D8DBEACBFFE4FF94EE861DBDFE6A16A2" box="[199,490,725,750]" italics="true" pageId="4" pageNumber="205">Measurements and ratios.</emphasis>
Males. Body 1.52–1.69. Carapace 0.43–0.49/0.41–0.48 (1.0). Chelicera: palm 0.44– 0.20 (2.2), movable finger 0.23. Pedipalp: femur 0.68–0.78/0.1–0.12 (6.8–7.1); patella 0.29–0.31/0.12–0.14 (2.2– 2.4); chela 0.93–1.03/0.14–0.16 (6.2–6.6), depth 0.14–0.17 (6.2–6.6); hand 0.37–0.41 (2.5–2.6), length/depth (2.5– 2.6); movable finger 0.56–0.60; ratio movable finger/hand 1.4–1.5; ratio femur/movable finger 1.2–1.3; ratio femur/carapace 1.6; ratio chela/carapace 2.1–2.2.
</paragraph>
<paragraph id="EA1036D9FFE4FF94EE861CE2FE58107D" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="205">Females. Body 1.58–1.78. Carapace 0.48–0.51/0.47–0.48 (1.0–1.1). Chelicera: palm 0.45–0.47/0.21–0.22 (2.1–2.2), movable finger 0.23–0.24. Pedipalp: femur 0.78–0.80/0.11–0.12 (6.9–7.1); patella 0.33–0.34/0.15 (2.2); chela 1.07–1.08/0.18 (5.9–6.1), depth 0.17–0.18 (5.9–6.1); hand 0.43–0.44 (2.4), length/depth 0.43–0.44 (2.4); movable finger 0.64–0.65; ratio movable finger/hand 1.5; ratio femur/movable finger 1.2; ratio femur/carapace 1.6; ratio chela/carapace 2.1–2.2.</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE4FF94EE861B56FDAE1019" box="[199,558,1085,1110]" pageId="4" pageNumber="205" type="distribution">
<paragraph id="EA1036D9FFE4FF94EE861B56FDAE1019" blockId="4.[151,1436,151,1542]" box="[199,558,1085,1110]" pageId="4" pageNumber="205">
<emphasis id="D8DBEACBFFE4FF94EE861B56FEE01019" bold="true" box="[199,352,1085,1110]" pageId="4" pageNumber="205">Distribution.</emphasis>
Andalusia,
<collectingCountry id="92B87649FFE4FF94EFA71B56FDAA1019" box="[486,554,1085,1110]" name="Spain" pageId="4" pageNumber="205">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE4FF94EE861B0AFE11124A" pageId="4" pageNumber="205" type="discussion">
<paragraph id="EA1036D9FFE4FF94EE861B0AFEE310A9" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="205">
<emphasis id="D8DBEACBFFE4FF94EE861B0AFEBB1035" bold="true" box="[199,315,1121,1146]" pageId="4" pageNumber="205">Remarks.</emphasis>
We have no hesitation in assigning these specimens to
<emphasis id="D8DBEACBFFE4FF94EDF81B09FB0D1035" box="[953,1165,1121,1146]" italics="true" pageId="4" pageNumber="205">
C. (E.)
<taxonomicName id="2DAF4D5AFFE4FF94EA4E1B0AFB0D1035" box="[1039,1165,1121,1146]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="cazorlensis">cazorlensis</taxonomicName>
</emphasis>
. The ratio of the chelal femur is distinctly slender in the studied sample (male 6.8–7.1, female 6.9–7.1), relative to the previously published data: male 6.27 (Carabajal Márquez
<emphasis id="D8DBEACBFFE4FF94ECEE1BC1FD69108D" box="[687,745,1193,1218]" italics="true" pageId="4" pageNumber="205">et al.</emphasis>
2010) and female 6.26 (Pérez &amp; Zaragoza 2010). The other data fit quite well.
</paragraph>
<paragraph id="EA1036D9FFE4FF94EE861B9AFED211D6" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="205">
The main difference between this species and
<emphasis id="D8DBEACBFFE4FF94EC881B99FCDD1146" box="[713,861,1266,1289]" italics="true" pageId="4" pageNumber="205">
C. (E.)
<taxonomicName id="2DAF4D5AFFE4FF94ED5B1B99FCDD1146" box="[794,861,1266,1289]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
, its close relative, is the length of the lamina on the chelal movable finger and the number of vestigial teeth without dental canals: short with 3–7 vestigial teeth (Pérez &amp; Zaragoza 2010; this paper) in
<emphasis id="D8DBEACBFFE4FF94EC461A51FD58111D" box="[519,728,1337,1362]" italics="true" pageId="4" pageNumber="205">
C. (E.)
<taxonomicName id="2DAF4D5AFFE4FF94EC1B1A52FD58111D" box="[602,728,1337,1362]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="cazorlensis">cazorlensis</taxonomicName>
</emphasis>
and long with 9–11 vestigial teeth in
<emphasis id="D8DBEACBFFE4FF94EAC31A51FA98111E" box="[1154,1304,1338,1361]" italics="true" pageId="4" pageNumber="205">
C. (E.)
<taxonomicName id="2DAF4D5AFFE4FF94EA941A51FA98111E" box="[1237,1304,1338,1361]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
. (Carabajal Márquez
<emphasis id="D8DBEACBFFE4FF94EF401A35FEB91139" box="[257,313,1373,1398]" italics="true" pageId="4" pageNumber="205">et al.</emphasis>
2011; this paper). The pedipalpal ratios in
<emphasis id="D8DBEACBFFE4FF94ED561A35FC661139" box="[791,998,1373,1398]" italics="true" pageId="4" pageNumber="205">
C. (E.)
<taxonomicName id="2DAF4D5AFFE4FF94ED291A36FC661139" box="[872,998,1373,1398]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="cazorlensis">cazorlensis</taxonomicName>
</emphasis>
are also significantly higher than those of
<emphasis id="D8DBEACBFFE4FF94EEF41AE9FECB11D6" box="[181,331,1410,1433]" italics="true" pageId="4" pageNumber="205">
C. (E.)
<taxonomicName id="2DAF4D5AFFE4FF94EF491AE9FECB11D6" box="[264,331,1410,1433]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
.
</paragraph>
<paragraph id="EA1036D9FFE4FF94EE861ACEFE11124A" blockId="4.[151,1436,151,1542]" pageId="4" pageNumber="205">
The records of
<emphasis id="D8DBEACBFFE4FF94EF341ACEFD9C11F2" box="[373,540,1445,1470]" italics="true" pageId="4" pageNumber="205">
<taxonomicName id="2DAF4D5AFFE4FF94EF341ACEFE6811F1" box="[373,488,1445,1470]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
</emphasis>
<taxonomicName id="2DAF4D5AFFE4FF94EC651ACDFC9511F1" authority="Zaragoza, 1985" authorityName="Zaragoza" authorityYear="1985" box="[548,789,1445,1470]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="verai">
<emphasis id="D8DBEACBFFE4FF94EC651ACDFDDE11F2" box="[548,606,1446,1469]" italics="true" pageId="4" pageNumber="205">verai</emphasis>
Zaragoza, 1985
</taxonomicName>
from this cave (Pérez Fernández &amp; Tinaut Ránera 2005; Barranco
<emphasis id="D8DBEACBFFE4FF94EF441AA1FEBD11AD" box="[261,317,1481,1506]" italics="true" pageId="4" pageNumber="205">et al.</emphasis>
2008; Pérez Fernández 2008; Pérez Fernández &amp; Tinaut Ránera 2009) are misidentifications based on
<emphasis id="D8DBEACBFFE4FF94EEFB1A85FE0A1249" box="[186,394,1517,1542]" italics="true" pageId="4" pageNumber="205">
C. (E.)
<taxonomicName id="2DAF4D5AFFE4FF94EF4D1A86FE0A1249" box="[268,394,1517,1542]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="cazorlensis">cazorlensis</taxonomicName>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,254 @@
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<mods:title id="789FA0D89743A127B39FCDEEE86C87EF">Hypogean pseudoscorpions (Arachnida) from Jaén province (Andalusia, Spain), with descriptions of four new species and a new synonymy</mods:title>
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<treatment id="620687CFFFE4FF96EED61933FD0F1788" ID-DOI="http://doi.org/10.5281/zenodo.6149884" ID-GBIF-Taxon="119574442" ID-Zenodo-Dep="6149884" LSID="urn:lsid:plazi:treatment:620687CFFFE4FF96EED61933FD0F1788" httpUri="http://treatment.plazi.org/id/620687CFFFE4FF96EED61933FD0F1788" lastPageId="6" lastPageNumber="207" pageId="4" pageNumber="205">
<subSubSection id="A2B56552FFE4FF94EED61933FF6B12DB" pageId="4" pageNumber="205" type="nomenclature">
<paragraph id="EA1036D9FFE4FF94EED61933FB67123C" blockId="4.[151,1255,1624,1685]" box="[151,1255,1624,1651]" pageId="4" pageNumber="205">
<heading id="B15881B5FFE4FF94EED61933FB67123C" bold="true" box="[151,1255,1624,1651]" fontSize="11" level="1" pageId="4" pageNumber="205" reason="1">
<emphasis id="D8DBEACBFFE4FF94EED61933FB67123C" bold="true" box="[151,1255,1624,1651]" pageId="4" pageNumber="205">
<emphasis id="D8DBEACBFFE4FF94EED61933FE1B123D" bold="true" box="[151,411,1624,1650]" italics="true" pageId="4" pageNumber="205">
<taxonomicName id="2DAF4D5AFFE4FF94EED61933FE96123D" ID-CoL="62LNY" box="[151,278,1624,1650]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
<taxonomicName id="2DAF4D5AFFE4FF94EF121933FE1B123D" authority="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez, 2011" authorityName="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez" authorityYear="2011" box="[339,411,1624,1650]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011
</emphasis>
</heading>
</paragraph>
<paragraph id="EA1036D9FFE4FF94EED61917FF6B12DB" blockId="4.[151,1255,1624,1685]" box="[151,235,1660,1685]" pageId="4" pageNumber="205">
(
<figureCitation id="72942A5CFFE4FF94EEDE1917FF6312DB" box="[159,227,1660,1685]" captionStart="FIGURES 2 – 5" captionStartId="5.[151,264,1280,1303]" captionTargetBox="[175,1412,308,1254]" captionTargetId="figure@5.[165,1421,301,1259]" captionTargetPageId="5" captionText="FIGURES 2 – 5. Habitus of living specimens. 2, Chthonius (E.) perezi Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011, Sistema de la Murcielaguina; 3, Chthonius (E.) espanyoli sp. nov., Complejo del Arroyo de la Rambla (PB – 4); 4, Neobisium (O.) perezi Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011, Sima del Campamento; 5, Allochernes masi (Navás, 1923), with the woodlouse Porcellio incanus Budde-Lund, 1879, Cueva de la Morciguilla." httpUri="https://zenodo.org/record/218270/files/figure.png" pageId="4" pageNumber="205">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE4FF94EED619AFFC341307" pageId="4" pageNumber="205" type="materials_examined">
<paragraph id="EA1036D9FFE4FF94EED619AFFC341307" blockId="4.[151,1437,1732,2009]" pageId="4" pageNumber="205">
<emphasis id="D8DBEACBFFE4FF94EED619AFFE861292" bold="true" box="[151,262,1732,1757]" pageId="4" pageNumber="205">Material.</emphasis>
<collectingCountry id="92B87649FFE4FF94EF5219AFFED61292" box="[275,342,1732,1757]" name="Spain" pageId="4" pageNumber="205">Spain</collectingCountry>
, Andalusia, Jaén province, Hornos de Segura, Sistema de la Murcielaguina (38º12′59.35″N, 2º42′37.13″W;
<quantity id="2D579B3CFFE4FF94EF03198CFE1E134F" box="[322,414,1767,1792]" metricMagnitude="3" metricUnit="m" metricValue="1.085" pageId="4" pageNumber="205" unit="m" value="1085.0">1085 m</quantity>
.a.s.l.),
<date id="9E111019FFE4FF94EFA81983FDD6134F" box="[489,598,1768,1792]" pageId="4" pageNumber="205" value="2010-01-31">31.I.2010</date>
,
<specimenCount id="FCA9FD50FFE4FF94EC231983FD0E12B0" box="[610,654,1768,1792]" pageId="4" pageNumber="205" type="generic">1 ♂</specimenCount>
;
<date id="9E111019FFE4FF94ECDB1983FC9C134F" box="[666,796,1768,1792]" pageId="4" pageNumber="205" value="2010-06-27">27.VI.2010</date>
, 1 ♀;
<date id="9E111019FFE4FF94ED1E1983FC73134F" box="[863,1011,1768,1792]" pageId="4" pageNumber="205" value="2011-08-14">14.VIII.2011</date>
,
<specimenCount id="FCA9FD50FFE4FF94EDBF1983FBAA12B0" box="[1022,1066,1768,1792]" pageId="4" pageNumber="205" type="generic">1 ♂</specimenCount>
;
<date id="9E111019FFE4FF94EA771983FB42134F" box="[1078,1218,1768,1792]" pageId="4" pageNumber="205" value="2012-12-24">24.XII.2012</date>
/
<date id="9E111019FFE4FF94EA8A1983FABE134F" box="[1227,1342,1768,1792]" pageId="4" pageNumber="205" value="2013-02-03">03.II.2013</date>
, 2 ♀, all lgt. G.E.V. and deposited in DEUA. Santiago-Pontones, Cueva Covarona (38º11′40.22″N, 2º27′45.18″W;
<quantity id="2D579B3CFFE4FF94EB241867FF341307" metricMagnitude="3" metricUnit="m" metricValue="1.13" pageId="4" pageNumber="205" unit="m" value="1130.0">1130 m</quantity>
.a.s.l.),
<date id="9E111019FFE4FF94EEBE185BFE011307" box="[255,385,1840,1864]" pageId="4" pageNumber="205" value="2002-11-09">09.XI.2002</date>
, lgt. J.L. Lencina &amp; V. Ortuño,
<specimenCount id="FCA9FD50FFE4FF94ECA2185BFC8F1308" box="[739,783,1840,1864]" pageId="4" pageNumber="205" type="generic">2 ♂</specimenCount>
, 5 ♀ (DEUA).
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE4FF95EE86183FFDCE14B8" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205" type="diagnosis">
<paragraph id="EA1036D9FFE4FF95EE86183FFDCE14B8" blockId="4.[151,1437,1732,2009]" lastBlockId="5.[151,1436,151,248]" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">
<emphasis id="D8DBEACBFFE4FF94EE86183FFEC11322" bold="true" box="[199,321,1876,1901]" pageId="4" pageNumber="205">Diagnosis.</emphasis>
Weakly troglomorphic. No eyes or eye-spots. Cheliceral palm with 6 setae and one lateral microseta; without isolated subapical tooth on movable finger; spinneret reduced in males, prominent in females. Carapace without preocular microsetae, two macrosetae on posterior margin. Pedipalp: femur length in males 0.6– 0.67 (6.0–6.1×), females 0.66–0.75 (5.7–6.4×); chela length in males 0.86–0.95 (length/depth 5.3–5.6), females 0.94–1.1 (length/depth 5.2–5.5); fixed finger with 19–21 teeth, movable finger with 10–15 teeth with dental canals reaching halfway between trichobothria
<emphasis id="D8DBEACBFFE5FF95EC1F1FD6FDF1149B" box="[606,625,189,212]" italics="true" pageId="5" pageNumber="206">st</emphasis>
and
<emphasis id="D8DBEACBFFE5FF95ECE81FD7FD42149A" box="[681,706,188,213]" italics="true" pageId="5" pageNumber="206">sb</emphasis>
, raised lamina with 9–11 rounded vestigial teeth without canals; trichobothrium
<emphasis id="D8DBEACBFFE5FF95EF071F8AFEE014B7" box="[326,352,225,248]" italics="true" pageId="5" pageNumber="206">ist</emphasis>
markedly distad
<emphasis id="D8DBEACBFFE5FF95EC631FB4FDC714B7" box="[546,583,223,248]" italics="true" pageId="5" pageNumber="206">esb</emphasis>
.
</paragraph>
</subSubSection>
<caption id="BED06651FFE5FF95EED61A6BFAB6113D" httpUri="https://zenodo.org/record/218270/files/figure.png" pageId="5" pageNumber="206" targetBox="[175,1412,308,1254]" targetPageId="5">
<paragraph id="EA1036D9FFE5FF95EED61A6BFAB6113D" blockId="5.[151,1436,1280,1395]" pageId="5" pageNumber="206">
<emphasis id="D8DBEACBFFE5FF95EED61A6BFEBF115A" bold="true" box="[151,319,1280,1303]" pageId="5" pageNumber="206">FIGURES 2–5.</emphasis>
Habitus of living specimens.
<emphasis id="D8DBEACBFFE5FF95ECC71A6BFD12115A" bold="true" box="[646,658,1280,1301]" pageId="5" pageNumber="206">2</emphasis>
,
<emphasis id="D8DBEACBFFE5FF95ECE41A6BFC071159" box="[677,903,1280,1302]" italics="true" pageId="5" pageNumber="206">
<taxonomicName id="2DAF4D5AFFE5FF95ECE41A6BFC8D1159" box="[677,781,1280,1302]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="206" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
<taxonomicName id="2DAF4D5AFFE5FF95ED0B1A6AFC071159" authority="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez, 2011" authorityName="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez" authorityYear="2011" box="[842,903,1281,1302]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011, Sistema de la Murcielaguina;
<emphasis id="D8DBEACBFFE5FF95EC3C1A74FD09117B" bold="true" box="[637,649,1311,1332]" pageId="5" pageNumber="206">3</emphasis>
,
<emphasis id="D8DBEACBFFE5FF95ECD61A74FC14117A" box="[663,916,1311,1333]" italics="true" pageId="5" pageNumber="206">
<taxonomicName id="2DAF4D5AFFE5FF95ECD61A74FD7F117A" box="[663,767,1311,1333]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="206" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
<taxonomicName id="2DAF4D5AFFE5FF95ED721A74FC14117A" box="[819,916,1311,1333]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="espanyoli" status="sp. nov.">espanyoli</taxonomicName>
</emphasis>
<taxonomicNameLabel id="C3E857B0FFE5FF95EDDA1A74FC6B117B" box="[923,1003,1311,1332]" pageId="5" pageNumber="206" rank="species">
<emphasis id="D8DBEACBFFE5FF95EDDA1A74FC6B117B" bold="true" box="[923,1003,1311,1332]" pageId="5" pageNumber="206">sp. nov.</emphasis>
</taxonomicNameLabel>
, Complejo del Arroyo de la Rambla (PB– 4);
<emphasis id="D8DBEACBFFE5FF95EEFD1A55FF49111C" bold="true" box="[188,201,1342,1363]" pageId="5" pageNumber="206">4</emphasis>
,
<emphasis id="D8DBEACBFFE5FF95EE981A55FE3E111B" box="[217,446,1342,1364]" italics="true" pageId="5" pageNumber="206">
<taxonomicName id="2DAF4D5AFFE5FF95EE981A55FEC5111B" box="[217,325,1342,1364]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="206" phylum="Arthropoda" rank="genus">Neobisium</taxonomicName>
(O.)
<taxonomicName id="2DAF4D5AFFE5FF95EFC01A54FE3E111B" authority="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez, 2011" authorityName="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez" authorityYear="2011" box="[385,446,1343,1364]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011, Sima del Campamento;
<emphasis id="D8DBEACBFFE5FF95EBC81A55FA15111C" bold="true" box="[1417,1429,1342,1363]" pageId="5" pageNumber="206">5</emphasis>
,
<taxonomicName id="2DAF4D5AFFE5FF95EED61A37FE5E113D" authority="Navas, 1923" authorityName="Navas" authorityYear="1923" box="[151,478,1372,1395]" class="Arachnida" family="Chernetidae" genus="Allochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="masi">
<emphasis id="D8DBEACBFFE5FF95EED61A37FEC6113C" box="[151,326,1372,1395]" italics="true" pageId="5" pageNumber="206">Allochernes masi</emphasis>
(Navás, 1923)
</taxonomicName>
, with the woodlouse
<taxonomicName id="2DAF4D5AFFE5FF95ECF41A37FBAF113D" authority="Budde-Lund, 1879" authorityName="Budde-Lund" authorityYear="1879" box="[693,1071,1372,1395]" class="Malacostraca" family="Porcellionidae" genus="Porcellio" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="incanus">
<emphasis id="D8DBEACBFFE5FF95ECF41A37FCE7113C" box="[693,871,1372,1395]" italics="true" pageId="5" pageNumber="206">Porcellio incanus</emphasis>
Budde-Lund, 1879
</taxonomicName>
, Cueva de la Morciguilla.
</paragraph>
</caption>
<subSubSection id="A2B56552FFE5FF96EE861AF4FE2717CF" lastPageId="6" lastPageNumber="207" pageId="5" pageNumber="206" type="description">
<paragraph id="EA1036D9FFE5FF95EE861AF4FBDB11F7" blockId="5.[151,1436,1439,2004]" box="[199,1115,1439,1464]" pageId="5" pageNumber="206">
<emphasis id="D8DBEACBFFE5FF95EE861AF4FED911F7" bold="true" box="[199,345,1439,1464]" pageId="5" pageNumber="206">Description.</emphasis>
Weakly troglomorphic facies; integument depigmented, pale brown.
</paragraph>
<paragraph id="EA1036D9FFE5FF95EE861AAFFED41208" blockId="5.[151,1436,1439,2004]" pageId="5" pageNumber="206">
Carapace slightly longer than broad, distinctly constricted posteriorly, subquadrate; anterior margin medially dentate, without epistome; without eyes or eye-spots; 18 setae, without preocular microsetae, formula: 4:6:4:2:2, anteromedial seta
<quantity id="2D579B3CFFE5FF95EF221967FD85126C" box="[355,517,1548,1572]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.8999999999999999" pageId="5" pageNumber="206" unit="mm" value="0.1" valueMax="0.11" valueMin="0.09">0.09–0.11 mm</quantity>
long, ocular seta
<quantity id="2D579B3CFFE5FF95EC8A1967FCF2126C" box="[715,882,1548,1572]" metricMagnitude="-5" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7" pageId="5" pageNumber="206" unit="mm" value="0.075" valueMax="0.08" valueMin="0.07">0.07–0.08 mm</quantity>
; 4–6 lyrifissures in anterior and ocular areas, two in posterior area.
</paragraph>
<paragraph id="EA1036D9FFE5FF95EE861938FB1F12DF" blockId="5.[151,1436,1439,2004]" pageId="5" pageNumber="206">
Pedipalpal coxa 5 setae (including 2 on manducatory process), coxa
<date id="9E111019FFE5FF95ED93193FFC711223" box="[978,1009,1620,1644]" pageId="5" pageNumber="206">I 3</date>
+ 3 marginal microsetae,
<date id="9E111019FFE5FF95EB5A193FFAC41223" box="[1307,1348,1620,1644]" pageId="5" pageNumber="206">II 4</date>
+ 9–12 bipinnate coxal spines,
<date id="9E111019FFE5FF95EFDF1913FE4F12DF" box="[414,463,1655,1680]" pageId="5" pageNumber="206">III 5</date>
+ 5–7 bipinnate coxal spines,
<date id="9E111019FFE5FF95ED651913FCD912DF" box="[804,857,1655,1680]" pageId="5" pageNumber="206">IV 6</date>
; intercoxal tubercle bisetose.
</paragraph>
<paragraph id="EA1036D9FFE5FF95EE8619F0FDF31350" blockId="5.[151,1436,1439,2004]" pageId="5" pageNumber="206">
Chaetotaxy of tergites I–XI: 4:4:4:4:6:6:6:6:6:4:1T2T1. Sternite II with 8–9 setae in males and
<quantity id="2D579B3CFFE5FF95EB0019F7FA1C12FB" box="[1345,1436,1691,1716]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="5" pageNumber="206" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9–10 in</quantity>
females; sternite III with (3)7–8(3) setae in females; sternite III in males with genital opening flanked by 9–10 setae on each side, 4+4 internal glandular setae and (3)7–8(3) marginal setae. Chaetotaxy of sternites IV–XI: (2)6– 8(2):8:6:6:6:6:6–7:0. Anal cone 0+2 setae.
</paragraph>
<paragraph id="EA1036D9FFE5FF95EE861840FE33139B" blockId="5.[151,1436,1439,2004]" pageId="5" pageNumber="206">
Cheliceral palm with 6 setae and one microseta laterally; seta
<emphasis id="D8DBEACBFFE5FF95ED3C1840FC13130B" box="[893,915,1835,1860]" italics="true" pageId="5" pageNumber="206">gl</emphasis>
0.58–0.62 from base of movable finger. Fixed finger with 10–12 teeth, two distal teeth distinctly larger than others, proximally reduced in size, 2–3 denticles at base. Movable finger without an isolated subapical tooth (
<emphasis id="D8DBEACBFFE5FF95ED711818FCC613C3" box="[816,838,1907,1932]" italics="true" pageId="5" pageNumber="206">di</emphasis>
), with a large distal tooth and 5–6 contiguous teeth, proximally decreasing in size, 2–3 denticles at base; spinneret strongly reduced or small in males, prominent and apically blunt in females.
</paragraph>
<paragraph id="EA1036D9FFE6FF96EE861FFCFE5E1657" blockId="6.[151,1436,151,968]" pageId="6" pageNumber="207">
Pedipalp with femoral chaetotaxy 3:6–7:3:5:1, four lyrifissures: 1 antiaxial and 3 paraxial. Chelal hand markedly depressed at level of
<emphasis id="D8DBEACBFFE6FF96EFB41FD7FDB5149A" box="[501,565,188,213]" italics="true" pageId="6" pageNumber="207">ib-isb</emphasis>
, with a short and prominent hump distad of
<emphasis id="D8DBEACBFFE6FF96EA681FD7FBE9149A" box="[1065,1129,188,213]" italics="true" pageId="6" pageNumber="207">ib-isb</emphasis>
and distinct ventral hollow just before base of movable chelal finger. Fixed finger with 19–20 pointed teeth (last 1–2 can be rounded), all with dental canals except the two small distal ones; third distal tooth from tip modified and little wider than the others; ten microtubercles at base; tip of finger with an accessory tooth (
<emphasis id="D8DBEACBFFE6FF96ED351E4CFC0A150F" box="[884,906,295,320]" italics="true" pageId="6" pageNumber="207">td</emphasis>
) on antiaxial face; four or five teeth at level of
<emphasis id="D8DBEACBFFE6FF96EED61E26FF4D152B" box="[151,205,333,356]" italics="true" pageId="6" pageNumber="207">est/it</emphasis>
occupying
<quantity id="2D579B3CFFE6FF96EF101E27FE29152B" box="[337,425,332,356]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="207" unit="mm" value="0.1">0.1 mm</quantity>
, distance between apices
<quantity id="2D579B3CFFE6FF96EC8B1E27FCF0152B" box="[714,880,332,356]" metricMagnitude="-5" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="6" pageNumber="207" unit="mm" value="0.025" valueMax="0.03" valueMin="0.02">0.02–0.03 mm</quantity>
. Movable finger with 12–15 mostly pointed teeth with dental canals, reaching halfway between trichobothria
<emphasis id="D8DBEACBFFE6FF96ED741E1AFCC815C7" box="[821,840,369,392]" italics="true" pageId="6" pageNumber="207">st</emphasis>
and
<emphasis id="D8DBEACBFFE6FF96ED3F1E04FC1715C7" box="[894,919,367,392]" italics="true" pageId="6" pageNumber="207">sb</emphasis>
, two distal ones tiny and one or two proximal ones reduced in size and blunt; proximally with slightly raised lamina with 9 rounded vestigial teeth, without dental canals, reaching proximad of trichobothrium
<emphasis id="D8DBEACBFFE6FF96EC811EDCFD59159F" box="[704,729,439,464]" italics="true" pageId="6" pageNumber="207">sb</emphasis>
, level with end of dental row of fixed finger; 3–5 tiny microtubercles at base; sensillum
<emphasis id="D8DBEACBFFE6FF96EC581EB6FDB415BB" box="[537,564,477,500]" italics="true" pageId="6" pageNumber="207">pc</emphasis>
between
<emphasis id="D8DBEACBFFE6FF96ECE31EB7FD3B15BA" box="[674,699,476,501]" italics="true" pageId="6" pageNumber="207">sb</emphasis>
and
<emphasis id="D8DBEACBFFE6FF96ECB41EB7FC8315BA" box="[757,771,476,501]" italics="true" pageId="6" pageNumber="207">b</emphasis>
, slightly closer to
<emphasis id="D8DBEACBFFE6FF96ED981EB7FC7215BA" box="[985,1010,476,501]" italics="true" pageId="6" pageNumber="207">sb</emphasis>
. Trichobothria
<emphasis id="D8DBEACBFFE6FF96EAE21EB6FB3D15BB" box="[1187,1213,477,500]" italics="true" pageId="6" pageNumber="207">ist</emphasis>
markedly distad of
<emphasis id="D8DBEACBFFE6FF96EED61E94FF3C1657" box="[151,188,511,536]" italics="true" pageId="6" pageNumber="207">esb</emphasis>
; ratio
<emphasis id="D8DBEACBFFE6FF96EF441D6AFE981657" box="[261,280,513,536]" italics="true" pageId="6" pageNumber="207">st</emphasis>
–
<emphasis id="D8DBEACBFFE6FF96EF671E94FEBF1657" box="[294,319,511,536]" italics="true" pageId="6" pageNumber="207">sb</emphasis>
/
<emphasis id="D8DBEACBFFE6FF96EF071E94FEDF1657" box="[326,351,511,536]" italics="true" pageId="6" pageNumber="207">sb</emphasis>
–
<emphasis id="D8DBEACBFFE6FF96EF2C1E94FEFB1657" box="[365,379,511,536]" italics="true" pageId="6" pageNumber="207">b</emphasis>
1.9–2.1.
</paragraph>
<paragraph id="EA1036D9FFE6FF96EE861D4EFC0C1683" blockId="6.[151,1436,151,968]" pageId="6" pageNumber="207">
<emphasis id="D8DBEACBFFE6FF96EE861D4EFE6E1673" box="[199,494,548,573]" italics="true" pageId="6" pageNumber="207">Measurements and ratios.</emphasis>
Males. Body 1.39–1.48. Carapace 0.42–0.43/0.37–0.42 (1.0–1.1). Chelicera: palm 0.36/0.17–0.19 (2.0–2.1), movable finger 0.18–0.21. Pedipalp: femur 0.60–0.67/0.10–0.11 (6.0–6.1); patella 0.26– 0.29/0.12–0.14 (2.1–2.3); chela 0.86–0.95/0.15–0.17 (5.6–5.7), depth 0.15–0.18 (5.4–5.6); hand 0.33–0.37 (2.2), length/depth (2.1–2.2); movable finger 0.52–0.55; ratio movable finger/hand 1.5–1.6; ratio femur/movable finger 1.1–1.2; ratio femur/carapace 1.4–1.6; ratio chela/carapace 2.1–2.2.
</paragraph>
<paragraph id="EA1036D9FFE6FF96EE861DBCFE2717CF" blockId="6.[151,1436,151,968]" pageId="6" pageNumber="207">Females. Body 1.5–1.56. Carapace 0.48–0.51/0.44–0.46 (1.1). Chelicera: palm 0.43–0.45/0.20–0.21 (2.2), movable finger 0.21–0.22. Pedipalp: femur 0.66–0.71/0.11–0.12 (5.7–6.4); patella 0.29–0.31/0.14–0.15 (2.1); chela 00.94–1.00/0.18 (5.3–5.4), depth 0.18–0.19 (5.2); hand 0.38–0.39(2.1–2.2), length/depth (2.0–2.1); movable finger 0.55–0.60; ratio movable finger/hand 1.4–1.5; ratio femur/movable finger 1.2; ratio femur/carapace 1.4; ratio chela/carapace 2.0.</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE6FF96EE861CE7FDAE17EA" box="[199,558,908,933]" pageId="6" pageNumber="207" type="distribution">
<paragraph id="EA1036D9FFE6FF96EE861CE7FDAE17EA" blockId="6.[151,1436,151,968]" box="[199,558,908,933]" pageId="6" pageNumber="207">
<emphasis id="D8DBEACBFFE6FF96EE861CE7FEE017EA" bold="true" box="[199,352,908,933]" pageId="6" pageNumber="207">Distribution.</emphasis>
Andalusia,
<collectingCountry id="92B87649FFE6FF96EFA71CE7FDAA17EA" box="[486,554,908,933]" name="Spain" pageId="6" pageNumber="207">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE6FF96EE861CC4FD0F1788" box="[199,655,943,968]" pageId="6" pageNumber="207" type="discussion">
<paragraph id="EA1036D9FFE6FF96EE861CC4FD0F1788" blockId="6.[151,1436,151,968]" box="[199,655,943,968]" pageId="6" pageNumber="207">
<emphasis id="D8DBEACBFFE6FF96EE861CC4FEBB1787" bold="true" box="[199,315,943,968]" pageId="6" pageNumber="207">Remarks.</emphasis>
See under
<emphasis id="D8DBEACBFFE6FF96EFF91CDBFD081787" box="[440,648,943,968]" italics="true" pageId="6" pageNumber="207">
C. (E.)
<taxonomicName id="2DAF4D5AFFE6FF96EC4B1CC4FD081787" box="[522,648,943,968]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="cazorlensis">cazorlensis</taxonomicName>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="5A89E160FBE769AA99B759B84A508AA9" type="GBIF-Dataset">3d28eeb6-88a2-4fb2-aebd-796c8b2c2287</mods:identifier>
<mods:identifier id="7B108E7436D1B982A2CC50191BBE856F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="00AD2A2CA39BAD45E91C518B28BC5741" type="Zenodo-Dep">218268</mods:identifier>
<mods:identifier id="7CFB94F1E28AE9F0435E053793BCC2B2" type="ZooBank">8119DAED-D7AF-4858-BC63-F082151A4DAE</mods:identifier>
</mods:mods>
<treatment id="620687CFFFE6FF99EED61B70FA19111B" ID-DOI="http://doi.org/10.5281/zenodo.6149886" ID-GBIF-Taxon="119574440" ID-Zenodo-Dep="6149886" LSID="urn:lsid:plazi:treatment:620687CFFFE6FF99EED61B70FA19111B" httpUri="http://treatment.plazi.org/id/620687CFFFE6FF99EED61B70FA19111B" lastPageId="9" lastPageNumber="210" pageId="6" pageNumber="207">
<subSubSection id="A2B56552FFE6FF96EED61B70FEB41018" pageId="6" pageNumber="207" type="nomenclature">
<paragraph id="EA1036D9FFE6FF96EED61B70FDAA1079" blockId="6.[151,554,1051,1111]" box="[151,554,1051,1078]" pageId="6" pageNumber="207">
<heading id="B15881B5FFE6FF96EED61B70FDAA1079" bold="true" box="[151,554,1051,1078]" fontSize="11" level="1" pageId="6" pageNumber="207" reason="1">
<emphasis id="D8DBEACBFFE6FF96EED61B70FDAA1079" bold="true" box="[151,554,1051,1078]" pageId="6" pageNumber="207">
<emphasis id="D8DBEACBFFE6FF96EED61B70FE45107A" bold="true" box="[151,453,1051,1077]" italics="true" pageId="6" pageNumber="207">
<taxonomicName id="2DAF4D5AFFE6FF96EED61B70FE96107A" ID-CoL="62LNY" box="[151,278,1051,1077]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
<taxonomicName id="2DAF4D5AFFE6FF96EF121B70FE45107A" box="[339,453,1051,1077]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="espanyoli" status="sp. nov.">espanyoli</taxonomicName>
</emphasis>
<taxonomicNameLabel id="C3E857B0FFE6FF96EF8D1B77FDAA1079" box="[460,554,1052,1078]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="EA1036D9FFE6FF96EED61B55FEB41018" blockId="6.[151,554,1051,1111]" box="[151,308,1086,1111]" pageId="6" pageNumber="207">
(
<figureCitation id="72942A5CFFE6FF96EEDE1B55FF671018" box="[159,231,1086,1111]" captionStart="FIGURES 2 – 5" captionStartId="5.[151,264,1280,1303]" captionTargetBox="[175,1412,308,1254]" captionTargetId="figure@5.[165,1421,301,1259]" captionTargetPageId="5" captionText="FIGURES 2 – 5. Habitus of living specimens. 2, Chthonius (E.) perezi Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011, Sistema de la Murcielaguina; 3, Chthonius (E.) espanyoli sp. nov., Complejo del Arroyo de la Rambla (PB – 4); 4, Neobisium (O.) perezi Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011, Sima del Campamento; 5, Allochernes masi (Navás, 1923), with the woodlouse Porcellio incanus Budde-Lund, 1879, Cueva de la Morciguilla." httpUri="https://zenodo.org/record/218270/files/figure.png" pageId="6" pageNumber="207">Figs 3</figureCitation>
,
<figureCitation id="72942A5CFFE6FF96EEB21B55FEAC1018" box="[243,300,1086,1111]" captionStart-0="FIGURES 6 – 9" captionStart-1="FIGURES 10 – 12" captionStartId-0="7.[151,264,895,918]" captionStartId-1="8.[151,264,1952,1975]" captionTargetBox-0="[193,1407,198,862]" captionTargetBox-1="[167,1413,207,1899]" captionTargetId-0="figure@7.[161,1426,193,875]" captionTargetId-1="figure@8.[151,1436,193,1932]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 6 – 9. Chthonius (Ephippiochthonius) espanyoli sp. nov., male holotype. 6, carapace; 7, partial view of anterior margin of carapace; 8, left chelicera; 9, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 7, 9), 0.2 (Fig. 8), 0.3 (Fig. 6)." captionText-1="FIGURES 10 – 12. Chthonius (Ephippiochthonius) espanyoli sp. nov., male holotype. 10, left pedipalp minus chela, dorsal view; 11, left chela, lateral view. Female paratype from Complejo del Arroyo de la Rambla (PB – 4), 12, left chela, lateral view. Scale bar 0.2 mm." httpUri-0="https://zenodo.org/record/218271/files/figure.png" httpUri-1="https://zenodo.org/record/218272/files/figure.png" pageId="6" pageNumber="207">6–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE6FF96EED61BEDFDBD11D4" pageId="6" pageNumber="207" type="materials_examined">
<paragraph id="EA1036D9FFE6FF96EED61BEDFDBD11D4" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="207">
<emphasis id="D8DBEACBFFE6FF96EED61BEDFECC10D0" bold="true" box="[151,332,1158,1183]" pageId="6" pageNumber="207">
<typeStatus id="3514887BFFE6FF96EED61BEDFF5210D0" box="[151,210,1158,1183]" pageId="6" pageNumber="207">Type</typeStatus>
material.
</emphasis>
<typeStatus id="3514887BFFE6FF96EF1B1BEDFE4210D0" box="[346,450,1158,1183]" pageId="6" pageNumber="207" type="holotype">Holotype</typeStatus>
♂,
<collectingCountry id="92B87649FFE6FF96EFBB1BEDFDBD10D0" box="[506,573,1158,1183]" name="Spain" pageId="6" pageNumber="207">Spain</collectingCountry>
, Andalusia, Jaén province, Cazorla, Cueva Secreta del Poyo Manquillo (37°53′18.46″N, 2°52′12.80″W;
<quantity id="2D579B3CFFE6FF96EC491BC1FDE6108C" box="[520,614,1194,1219]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="207" unit="m" value="1500.0">1500 m</quantity>
.a.s.l.),
<date id="9E111019FFE6FF96ECF21BC0FCB5108C" box="[691,821,1195,1219]" pageId="6" pageNumber="207" value="2008-09-21">21.IX.2008</date>
, lgt. G.E.V. (DEUA: 20025).
<typeStatus id="3514887BFFE6FF96EAC61BC0FB78108C" box="[1159,1272,1195,1219]" pageId="6" pageNumber="207" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="FCA9FD50FFE6FF96EB451BC0FAB0108D" box="[1284,1328,1195,1219]" pageId="6" pageNumber="207" type="generic">1 ♂</specimenCount>
(MCNB:
<date id="9E111019FFE6FF96EED61BA5FE9010A8" box="[151,272,1230,1255]" pageId="6" pageNumber="207" value="2013" valueMax="2541">2013-2541</date>
MZB), same locality as
<typeStatus id="3514887BFFE6FF96EC6B1BA5FD0F10A8" box="[554,655,1230,1255]" pageId="6" pageNumber="207" type="holotype">holotype</typeStatus>
,
<date id="9E111019FFE6FF96ECDA1BA4FCA610A8" box="[667,806,1231,1255]" pageId="6" pageNumber="207" value="2011-07-03">03.VII.2011</date>
; 1 ♀ (DEUA: 20009),
<collectingCountry id="92B87649FFE6FF96EA6B1BA5FBED10A8" box="[1066,1133,1230,1255]" name="Spain" pageId="6" pageNumber="207">Spain</collectingCountry>
, Andalusia, Jaén province, Peal de Becerro, Complejo del Arroyo de la Rambla (PB-4) (37º48′1.22″N, 2º53′36.65″W,
<quantity id="2D579B3CFFE6FF96EAB51B98FAD81144" box="[1268,1368,1266,1291]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="207" unit="m" value="1000.0">1000 m</quantity>
.a.s.l.),
<date id="9E111019FFE6FF96EED61A7CFE901160" box="[151,272,1303,1327]" pageId="6" pageNumber="207" value="2007-10-13">13.X.2007</date>
; same locality, 4 ♀, 2 T (2 ♀ DEUA: 20039, 20052, 1 ♀ MCNB:
<date id="9E111019FFE6FF96EDB41A7DFBEF1160" box="[1013,1135,1302,1327]" pageId="6" pageNumber="207" value="2013" valueMax="2542">2013-2542</date>
MZB, 1 ♀ MNCN: 20.02/ 1728, 2 T DEUA: 20053, 20054),
<date id="9E111019FFE6FF96EC621A51FD38111C" box="[547,696,1338,1363]" pageId="6" pageNumber="207" value="2010-08-26">26.VIII.2010</date>
; 1 ♀ (DEUA: 20026),
<date id="9E111019FFE6FF96ED801A50FBCC111C" box="[961,1100,1339,1363]" pageId="6" pageNumber="207" value="2011-12-04">04.XII.2011</date>
; 1 ♀ (DEUA: 20030),
<collectingCountry id="92B87649FFE6FF96EB141A51FA18111C" box="[1365,1432,1338,1363]" name="Spain" pageId="6" pageNumber="207">Spain</collectingCountry>
, Andalusia, Jaén province, Hornos de Segura, Sima Jesusín (38º10′31.90″N, 2º46′58.45″W;
<specimenCount id="FCA9FD50FFE6FF96EABE1A35FADC1138" box="[1279,1372,1374,1399]" pageId="6" pageNumber="207" type="generic">
<quantity id="2D579B3CFFE6FF96EABE1A35FAD81138" box="[1279,1368,1374,1399]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="6" pageNumber="207" unit="m" value="769.0">769 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="9E111019FFE6FF96EED61AE8FEA211D4" box="[151,290,1411,1435]" pageId="6" pageNumber="207" value="2010-07-11">11.VII.2010</date>
. All
<typeStatus id="3514887BFFE6FF96EF161AE8FE4211D4" box="[343,450,1411,1435]" pageId="6" pageNumber="207" type="paratype">paratypes</typeStatus>
lgt. G.E.V.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE6FF96EE861ACDFDFF1200" pageId="6" pageNumber="207" type="etymology">
<paragraph id="EA1036D9FFE6FF96EE861ACDFDFF1200" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="207">
<emphasis id="D8DBEACBFFE6FF96EE861ACDFECE11F0" bold="true" box="[199,334,1446,1471]" pageId="6" pageNumber="207">Etymology.</emphasis>
The new species is dedicated to the memory of Dr Francesc Español Coll (
<date id="9E111019FFE6FF96EADC1ACCFAA011F0" box="[1181,1312,1447,1471]" pageId="6" pageNumber="207" value="1907" valueMax="1999">1907–1999</date>
), specialist in
<taxonomicName id="2DAF4D5AFFE6FF96EEF41AA1FEB311AC" box="[181,307,1482,1507]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
, one of the greatest biospeleologists in the World, invertebrate Curator and later Director of Museo de Zoología de Barcelona (Museu de Ciències Naturals) for more than 50 years. Dr Español collected a pseudoscorpion specimen in the Cueva Secreta del Poyo Manquillo in 1953, identified as
<emphasis id="D8DBEACBFFE6FF96EAE01979FA1C1264" box="[1185,1436,1554,1579]" italics="true" pageId="6" pageNumber="207">
<taxonomicName id="2DAF4D5AFFE6FF96EAE01979FA991264" box="[1185,1305,1554,1579]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">Neobisium</taxonomicName>
(O.)
<taxonomicName id="2DAF4D5AFFE6FF96EB18197FFA1C1264" box="[1369,1436,1556,1579]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
and recorded in this paper for the first time.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE6FF96EE861931FEA51305" pageId="6" pageNumber="207" type="diagnosis">
<paragraph id="EA1036D9FFE6FF96EE861931FEA51305" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="207">
<emphasis id="D8DBEACBFFE6FF96EE861931FEC1123C" bold="true" box="[199,321,1626,1651]" pageId="6" pageNumber="207">Diagnosis.</emphasis>
Strongly troglomorphic facies. No eyes or eye-spots. Chelicera with 6 (rarely 7) setae on palm and one microseta; isolated subapical tooth absent on movable finger; spinneret strongly reduced in males, prominent and apically rounded in females. Carapace with two macrosetae on posterior margin, without preocular microsetae. Pedipalp: femur length 0.99–1.03 (7.6–8.1×) (♂), 0.91–0.94 (6.5–7.2×) (♀); chela length 1.37–1.44 (length/depth 6.6–7.0) (♂), 1.27–1.29 (length/depth 5.8–6.5) (♀); fixed finger with 26–27 (♂), 21–22 (♀) teeth; movable finger with 18–19 (♂), 14–15 (♀) teeth and raised lamina with 8–13 rounded vestigial teeth; trichobothrium
<emphasis id="D8DBEACBFFE6FF96EB531864FAAC1369" box="[1298,1324,1807,1830]" italics="true" pageId="6" pageNumber="207">ist</emphasis>
distinctly distal of
<emphasis id="D8DBEACBFFE6FF96EEB81859FE9E1304" box="[249,286,1842,1867]" italics="true" pageId="6" pageNumber="207">esb</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE6FF99EE86183DFB2F1713" lastPageId="9" lastPageNumber="210" pageId="6" pageNumber="207" type="description">
<paragraph id="EA1036D9FFE6FF96EE86183DFAC61394" blockId="6.[151,1437,1158,2011]" pageId="6" pageNumber="207">
<emphasis id="D8DBEACBFFE6FF96EE86183DFED91320" bold="true" box="[199,345,1878,1903]" pageId="6" pageNumber="207">Description.</emphasis>
Male
<typeStatus id="3514887BFFE6FF96EFE0183DFD861320" box="[417,518,1878,1903]" pageId="6" pageNumber="207" type="holotype">holotype</typeStatus>
, followed (when different) by male
<typeStatus id="3514887BFFE6FF96EDDC183CFC7D1320" box="[925,1021,1879,1903]" pageId="6" pageNumber="207" type="paratype">paratype</typeStatus>
in parentheses and female
<typeStatus id="3514887BFFE6FF96EB71183CFA1C1320" box="[1328,1436,1879,1903]" pageId="6" pageNumber="207" type="paratype">paratypes</typeStatus>
in square brackets. Large species; strongly troglomorphic; integument depigmented, pale yellowish-brown; weak hispid granulation on lateral surfaces of ocular and posterior areas of carapace, on cheliceral palm, on base of paraxial face of pedipalpal femur, on base of movable finger and on dorsodistal surface of pedipalpal hand.
</paragraph>
<caption id="BED06651FFE7FF97EED61C14FE8F179C" httpUri="https://zenodo.org/record/218271/files/figure.png" pageId="7" pageNumber="208" targetBox="[193,1407,198,862]" targetPageId="7">
<paragraph id="EA1036D9FFE7FF97EED61C14FE8F179C" blockId="7.[151,1436,895,979]" pageId="7" pageNumber="208">
<emphasis id="D8DBEACBFFE7FF97EED61C14FEBE17DA" bold="true" box="[151,318,895,918]" pageId="7" pageNumber="208">FIGURES 6–9.</emphasis>
<taxonomicName id="2DAF4D5AFFE7FF97EF061CEBFD7417D9" box="[327,756,896,918]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="espanyoli" status="sp. nov." subGenus="Ephippiochthonius">
<emphasis id="D8DBEACBFFE7FF97EF061CEBFD7417D9" box="[327,756,896,918]" italics="true" pageId="7" pageNumber="208">Chthonius (Ephippiochthonius) espanyoli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E857B0FFE7FF97ECBF1CEBFCD117DA" box="[766,849,896,917]" pageId="7" pageNumber="208" rank="species">
<emphasis id="D8DBEACBFFE7FF97ECBF1CEBFCD117DA" bold="true" box="[766,849,896,917]" pageId="7" pageNumber="208">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype.
<emphasis id="D8DBEACBFFE7FF97EA471CEBFB9317DA" bold="true" box="[1030,1043,896,917]" pageId="7" pageNumber="208">6</emphasis>
, carapace;
<emphasis id="D8DBEACBFFE7FF97EACD1CEBFB1917DA" bold="true" box="[1164,1177,896,917]" pageId="7" pageNumber="208">7</emphasis>
, partial view of anterior margin of carapace;
<emphasis id="D8DBEACBFFE7FF97EF241CF4FEF217FB" bold="true" box="[357,370,927,948]" pageId="7" pageNumber="208">8</emphasis>
, left chelicera;
<emphasis id="D8DBEACBFFE7FF97EC511CF4FD9C17FB" bold="true" box="[528,540,927,948]" pageId="7" pageNumber="208">9</emphasis>
, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 7, 9), 0.2 (Fig. 8), 0.3 (Fig. 6).
</paragraph>
</caption>
<paragraph id="EA1036D9FFE7FF97EE861B6BFBF110CB" blockId="7.[151,1437,1024,2021]" pageId="7" pageNumber="208">
Carapace (
<figureCitation id="72942A5CFFE7FF97EF031B6BFE081056" box="[322,392,1024,1049]" captionStart="FIGURES 6 – 9" captionStartId="7.[151,264,895,918]" captionTargetBox="[193,1407,198,862]" captionTargetId="figure@7.[161,1426,193,875]" captionTargetPageId="7" captionText="FIGURES 6 – 9. Chthonius (Ephippiochthonius) espanyoli sp. nov., male holotype. 6, carapace; 7, partial view of anterior margin of carapace; 8, left chelicera; 9, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 7, 9), 0.2 (Fig. 8), 0.3 (Fig. 6)." httpUri="https://zenodo.org/record/218271/files/figure.png" pageId="7" pageNumber="208">Fig. 6</figureCitation>
) as long as broad or slightly longer than broad, distinctly constricted posteriorly; anterior margin (
<figureCitation id="72942A5CFFE7FF97EEB41B4FFEBA1072" box="[245,314,1060,1085]" captionStart="FIGURES 6 – 9" captionStartId="7.[151,264,895,918]" captionTargetBox="[193,1407,198,862]" captionTargetId="figure@7.[161,1426,193,875]" captionTargetPageId="7" captionText="FIGURES 6 – 9. Chthonius (Ephippiochthonius) espanyoli sp. nov., male holotype. 6, carapace; 7, partial view of anterior margin of carapace; 8, left chelicera; 9, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 7, 9), 0.2 (Fig. 8), 0.3 (Fig. 6)." httpUri="https://zenodo.org/record/218271/files/figure.png" pageId="7" pageNumber="208">Fig. 7</figureCitation>
) weakly prominent, medially dentate, without epistome. Without eyes or eye-spots. Chaetotaxy: 18 (19) setae, without preocular microsetae, formula: 4:6:4:2:2 (4:6:4:3:2), anteromedial seta 0.12 [0.13] mm long, ocular seta
<quantity id="2D579B3CFFE7FF97EF571B07FEFD10CB" box="[278,381,1132,1157]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="208" unit="mm" value="0.06">0.06 mm</quantity>
. Five lyrifissures in anterior and ocular areas, two in posterior area.
</paragraph>
<paragraph id="EA1036D9FFE7FF97EE861BFBFA051082" blockId="7.[151,1437,1024,2021]" pageId="7" pageNumber="208">
Pedipalpal coxa 5 setae (including 2 on manducatory process), coxa
<date id="9E111019FFE7FF97ED931BFAFC7110E7" box="[978,1009,1168,1192]" pageId="7" pageNumber="208">I 3</date>
+ 3 marginal microsetae,
<date id="9E111019FFE7FF97EB5A1BFAFAC410E7" box="[1307,1348,1168,1192]" pageId="7" pageNumber="208">II 4</date>
+ 9–10 (10–11) [10–11] bipinnate coxal spines,
<date id="9E111019FFE7FF97EC1A1BDEFD0C1082" box="[603,652,1204,1229]" pageId="7" pageNumber="208">III 5</date>
+ 6 [5–7] bipinnate coxal spines,
<date id="9E111019FFE7FF97EA481BDEFBBE1082" box="[1033,1086,1204,1229]" pageId="7" pageNumber="208">IV 6</date>
; intercoxal tubercle bisetose.
</paragraph>
<paragraph id="EA1036D9FFE7FF97EE861BB3FCCF1113" blockId="7.[151,1437,1024,2021]" pageId="7" pageNumber="208">
Chaetotaxy of tergites I–XI: 4:4:4:4:6:6:6:6:6:4:1T2T1. Genital area with 9 setae on sternite II in males and females; sternite III with genital opening flanked by 8 setae on each side, 4+4 internal glandular setae and (3)7(3) marginal setae. Chaetotaxy of sternites IV–XI: (2)7(2):7:6:6:6:6:7:0; male
<typeStatus id="3514887BFFE7FF97ED9C1A4AFBBD1176" box="[989,1085,1313,1337]" pageId="7" pageNumber="208" type="paratype">paratype</typeStatus>
with 9 [7–8] marginal setae on sternite III, IV: 6, V: 6 [7–8], X: 6 [6–7]. Anal cone 0+2 setae.
</paragraph>
<paragraph id="EA1036D9FFE7FF97EE861A03FDFA127A" blockId="7.[151,1437,1024,2021]" pageId="7" pageNumber="208">
Chelicera (
<figureCitation id="72942A5CFFE7FF97EF021A03FE2911CF" box="[323,425,1384,1409]" captionStart="FIGURES 6 – 9" captionStartId="7.[151,264,895,918]" captionTargetBox="[193,1407,198,862]" captionTargetId="figure@7.[161,1426,193,875]" captionTargetPageId="7" captionText="FIGURES 6 – 9. Chthonius (Ephippiochthonius) espanyoli sp. nov., male holotype. 6, carapace; 7, partial view of anterior margin of carapace; 8, left chelicera; 9, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 7, 9), 0.2 (Fig. 8), 0.3 (Fig. 6)." httpUri="https://zenodo.org/record/218271/files/figure.png" pageId="7" pageNumber="208">Figs 8–9</figureCitation>
) with 6 [6–7] setae and 1 microseta laterally on palm, seta
<emphasis id="D8DBEACBFFE7FF97EA1B1A03FBF511CE" box="[1114,1141,1384,1409]" italics="true" pageId="7" pageNumber="208">vb</emphasis>
<quantity id="2D579B3CFFE7FF97EA3F1A03FB6411CF" box="[1150,1252,1384,1409]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="208" unit="mm" value="0.05">0.05 mm</quantity>
long, microseta
<quantity id="2D579B3CFFE7FF97EED61AE7FE8D11EB" box="[151,269,1420,1445]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="208" unit="mm" value="0.035">0.035 mm</quantity>
long; seta
<emphasis id="D8DBEACBFFE7FF97EFD51AE7FE2A11EA" box="[404,426,1420,1445]" italics="true" pageId="7" pageNumber="208">gl</emphasis>
0.55 [0.56] from base of movable finger. Fixed finger with 11 teeth, two distal teeth distinctly larger than others, proximally decreasing in size. Movable finger without an isolated subapical tooth (
<emphasis id="D8DBEACBFFE7FF97EB371ADBFA0C1186" box="[1398,1420,1456,1481]" italics="true" pageId="7" pageNumber="208">di</emphasis>
), with a large distal tooth and 10 contiguous teeth reduced in size proximally; spinneret in males strongly reduced, almost absent, prominent and apically rounded in females. Rallum with 11 blades. Serrula exterior with 16 [17–18] blades, serrula interior with 14 [13] blades.
</paragraph>
<paragraph id="EA1036D9FFE7FF97EE86192BFB6713AB" blockId="7.[151,1437,1024,2021]" pageId="7" pageNumber="208">
Pedipalp (
<figureCitation id="72942A5CFFE7FF97EF78192BFE3C1217" box="[313,444,1600,1625]" captionStart="FIGURES 10 – 12" captionStartId="8.[151,264,1952,1975]" captionTargetBox="[167,1413,207,1899]" captionTargetId="figure@8.[151,1436,193,1932]" captionTargetPageId="8" captionText="FIGURES 10 – 12. Chthonius (Ephippiochthonius) espanyoli sp. nov., male holotype. 10, left pedipalp minus chela, dorsal view; 11, left chela, lateral view. Female paratype from Complejo del Arroyo de la Rambla (PB – 4), 12, left chela, lateral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/218272/files/figure.png" pageId="7" pageNumber="208">Figs 10–12</figureCitation>
) with femoral chaetotaxy 3:6:3:5:1 [3:6–7:3:5–6:1], four lyrifissures: 1 antiaxial and 3 paraxial. Chelal hand depressed at level of
<emphasis id="D8DBEACBFFE7FF97EC37190FFD361232" box="[630,694,1636,1661]" italics="true" pageId="7" pageNumber="208">ib-isb</emphasis>
, with a low, rounded hump distad of
<emphasis id="D8DBEACBFFE7FF97EA17190FFB161232" box="[1110,1174,1636,1661]" italics="true" pageId="7" pageNumber="208">ib-isb</emphasis>
in males, slightly more prominent in females, and a weak ventral hollow just before base of movable chelal finger; chaetotaxy 4:5:4. Fixed finger with 26 (27) [21–22] pointed teeth, all with dental canals except the two small distal ones, dental row reaching proximad of trichobothrium
<emphasis id="D8DBEACBFFE7FF97EC1819BBFDF212A6" box="[601,626,1744,1769]" italics="true" pageId="7" pageNumber="208">sb</emphasis>
; third distal tooth from tip modified, little wider than others; four microtubercles at base; tip of finger with an accessory tooth (
<emphasis id="D8DBEACBFFE7FF97ED2A199FFC011342" box="[875,897,1780,1805]" italics="true" pageId="7" pageNumber="208">td</emphasis>
) on antiaxial face; four teeth at level of
<emphasis id="D8DBEACBFFE7FF97EB24199EFA1C1343" box="[1381,1436,1781,1804]" italics="true" pageId="7" pageNumber="208">est/it</emphasis>
occupying
<quantity id="2D579B3CFFE7FF97EF541873FEEF137F" box="[277,367,1816,1840]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="208" unit="mm" value="0.1">0.1 mm</quantity>
, distance between apices 0.025 [0.025–0.03] mm. Distal half of movable finger with 18 (19) [14–15] pointed teeth, all with dental canals except the two tiny distal ones; proximal half of finger with slightly raised lamina with 13 (8) [10–13] rounded vestigial teeth reaching slightly distad of halfway between trichobothria
<emphasis id="D8DBEACBFFE7FF97EED618EFFF2513D2" box="[151,165,1924,1949]" italics="true" pageId="7" pageNumber="208">b</emphasis>
and
<emphasis id="D8DBEACBFFE7FF97EE9F18EFFF7713D2" box="[222,247,1924,1949]" italics="true" pageId="7" pageNumber="208">sb</emphasis>
, level with coupled sensilla
<emphasis id="D8DBEACBFFE7FF97EC7E18EEFDD913D3" box="[575,601,1925,1948]" italics="true" pageId="7" pageNumber="208">pc</emphasis>
, three microtubercles at base; basal apodeme long and apically indented. Trichobothria as in figs 11–12;
<emphasis id="D8DBEACBFFE7FF97EFBA18C2FD96138F" box="[507,534,1961,1984]" italics="true" pageId="7" pageNumber="208">ist</emphasis>
markedly distal of
<emphasis id="D8DBEACBFFE7FF97ECB218C3FC98138E" box="[755,792,1960,1985]" italics="true" pageId="7" pageNumber="208">esb</emphasis>
, distance
<emphasis id="D8DBEACBFFE7FF97EDCB18C2FC25138F" box="[906,933,1961,1984]" italics="true" pageId="7" pageNumber="208">ist</emphasis>
to
<emphasis id="D8DBEACBFFE7FF97ED8B18C3FC6F138E" box="[970,1007,1960,1985]" italics="true" pageId="7" pageNumber="208">esb</emphasis>
0.06 [0.05–0.08] mm, distance
<emphasis id="D8DBEACBFFE7FF97EB1818C3FAFE138E" box="[1369,1406,1960,1985]" italics="true" pageId="7" pageNumber="208">esb</emphasis>
to
<emphasis id="D8DBEACBFFE7FF97EED618A7FF3213AA" box="[151,178,1996,2021]" italics="true" pageId="7" pageNumber="208">eb</emphasis>
0.03 [0.03–0.04] mm; distance between
<emphasis id="D8DBEACBFFE7FF97EC3A18A7FD3513AA" box="[635,693,1996,2021]" italics="true" pageId="7" pageNumber="208">st–sb</emphasis>
2.3 (1.8) [2.1–2.2] longer than that between
<emphasis id="D8DBEACBFFE7FF97EAEA18A7FB6013AA" box="[1195,1248,1996,2021]" italics="true" pageId="7" pageNumber="208">sb–b</emphasis>
.
</paragraph>
<caption id="BED06651FFE8FF98EED618CBFECF13BB" httpUri="https://zenodo.org/record/218272/files/figure.png" pageId="8" pageNumber="209" targetBox="[167,1413,207,1899]" targetPageId="8">
<paragraph id="EA1036D9FFE8FF98EED618CBFECF13BB" blockId="8.[151,1437,1952,2036]" pageId="8" pageNumber="209">
<emphasis id="D8DBEACBFFE8FF98EED618CBFED713F9" bold="true" box="[151,343,1952,1975]" pageId="8" pageNumber="209">FIGURES 10–12.</emphasis>
<taxonomicName id="2DAF4D5AFFE8FF98EF2118CAFC8B13F8" box="[352,779,1953,1975]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="espanyoli" status="sp. nov." subGenus="Ephippiochthonius">
<emphasis id="D8DBEACBFFE8FF98EF2118CAFC8B13F8" box="[352,779,1953,1975]" italics="true" pageId="8" pageNumber="209">Chthonius (Ephippiochthonius) espanyoli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E857B0FFE8FF98ED5418CAFCE913F9" box="[789,873,1953,1974]" pageId="8" pageNumber="209" rank="species">
<emphasis id="D8DBEACBFFE8FF98ED5418CAFCE913F9" bold="true" box="[789,873,1953,1974]" pageId="8" pageNumber="209">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype.
<emphasis id="D8DBEACBFFE8FF98EA5D18CAFBB513F9" bold="true" box="[1052,1077,1953,1974]" pageId="8" pageNumber="209">10</emphasis>
, left pedipalp minus chela, dorsal view;
<emphasis id="D8DBEACBFFE8FF98EE9418ABFF6C139A" bold="true" box="[213,236,1984,2005]" pageId="8" pageNumber="209">11</emphasis>
, left chela, lateral view. Female paratype from Complejo del Arroyo de la Rambla (PB–4),
<emphasis id="D8DBEACBFFE8FF98EACE18ABFB28139A" bold="true" box="[1167,1192,1984,2005]" pageId="8" pageNumber="209">12</emphasis>
, left chela, lateral view. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1036D9FFE9FF99EE861FF3FF58149B" blockId="9.[151,1436,151,1365]" pageId="9" pageNumber="210">Leg IV tactile seta ratios: tibia 0.57 (0.54) [0.54–0.55], basitarsus 0.45 (0.44) [0.40–0.42], tarsus 0.31 [0.27– 0.29].</paragraph>
<paragraph id="EA1036D9FFE9FF99EE861F8AFE661673" blockId="9.[151,1436,151,1365]" pageId="9" pageNumber="210">
<emphasis id="D8DBEACBFFE9FF99EE861F8AFE6A14B7" box="[199,490,223,248]" italics="true" pageId="9" pageNumber="210">Measurements and ratios.</emphasis>
Male
<typeStatus id="3514887BFFE9FF99EC731FB4FD1714B7" box="[562,663,223,248]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
, followed by male
<typeStatus id="3514887BFFE9FF99ED2E1F8BFC4F14B7" box="[879,975,224,248]" pageId="9" pageNumber="210" type="paratype">paratype</typeStatus>
from Cueva Secreta del Poyo Manquillo in square brackets. Body 1.78 [1.88]. Carapace 0.54/0.55 (1.0) [0.59/0.53 (1.1)]. Chelicera palm 0.56/0.25 (2.2) [0.55/0.25 (2.2)], movable finger 0.28 [0.29]. Pedipalp femur 0.99/0.13 (7.6) [1.03/0.13 (8.1)]; patella 0.40/0.16 (2.4) [0.41/0.17 (2.5)]; chela 1.37/0.20 (6.8) [1.44/0.20 (7.0)], depth 0.21 (6.6) [0.21 (7.0)]; hand 0.54 (2.7) [0.55 (2.7)], length/depth (2.6) [(2.6)]; movable finger 0.84 [0.88]; ratio movable finger/hand 1.6; ratio femur/movable finger 1.2; ratio femur/carapace 1.3; ratio chela/carapace 1.8 [1.7]. Leg I: femur 0.52/0.08 (6.2) [0.56/0.08 (6.7)]; patella 0.28/0.08 (3.7) [0.27/0.08 (3.6)]; tibia 0.30/0.06 (5.0) [0.33/0.06 (5.5)]; tarsus 0.60/0.05 (11.8) [0.61/0.05 (12.0)]; ratio femur/patella 1.9 [2.1]. Leg IV: femur+patella 0.81/0.27 (3.0) [0.83/0.27 (3.1)]; tibia 0.56/0.11 (5.2) [0.57/0.11 (5.4)]; basitarsus 0.28/0.08 (3.4) [0.30/0.08 (3.6)]; telotarsus 0.64/0.05 (14.0) [0.66/0.05 (13.3)]; ratio telotarsus/basitarsus 2.3 [2.2].
</paragraph>
<paragraph id="EA1036D9FFE9FF99EE861D2CFB2F1713" blockId="9.[151,1436,151,1365]" pageId="9" pageNumber="210">
Female
<typeStatus id="3514887BFFE9FF99EF611D23FE0B162F" box="[288,395,584,608]" pageId="9" pageNumber="210" type="paratype">paratypes</typeStatus>
from Complejo del Arroyo de la Rambla (PB-4). Body 1.74–1.94. Carapace 0.57–0.59/0.52– 0.55 (1.1). Chelicera: palm 0.53–0.55/0.23–0.25 (2.2–2.3), movable finger 0.27. Pedipalp: femur 0.91–0.94/0.13– 0.14 (6.5–7.2); patella 0.37–0.40/0.15–0.17 (2.4); chela 1.27–1.29/0.20–0.21 (5.9–6.6), depth 0.20–0.22 (5.8–6.5); hand 0.50–0.51 (2.4–2.6), length/depth (2.3–2.5); movable finger 0.76–0.78; ratio movable finger/hand 1.5; ratio femur/movable finger 1.2; ratio femur/carapace 1.5–1.7; ratio chela/carapace 2.2. Leg I: femur 0.48–0.50/0.08 (6.0–6.2); patella 0.25–0.26/0.07 (3.7–3.8); tibia 0.25–0.28/0.06 (4.5–4.9); tarsus 0.54–0.55/0.05 (11.1–11.4); ratio femur/patella 1.9–2.0. Leg IV: femur+patella 0.70–0.72/0.24–0.25 (2.9); tibia 0.49–0.51/0.10–0.11 (4.9); basitarsus 0.25/0.08 (3.1–3.4); telotarsus 0.52–0.56/0.05 (10.7–12.4); ratio telotarsus/basitarsus 2.1–2.2.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE9FF99EE861C0CFDAE17CF" box="[199,558,871,896]" pageId="9" pageNumber="210" type="distribution">
<paragraph id="EA1036D9FFE9FF99EE861C0CFDAE17CF" blockId="9.[151,1436,151,1365]" box="[199,558,871,896]" pageId="9" pageNumber="210">
<emphasis id="D8DBEACBFFE9FF99EE861C0CFEE017CF" bold="true" box="[199,352,871,896]" pageId="9" pageNumber="210">Distribution.</emphasis>
Andalusia,
<collectingCountry id="92B87649FFE9FF99EFA71C0CFDAA17CF" box="[486,554,871,896]" name="Spain" pageId="9" pageNumber="210">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE9FF99EE861CE7FA19111B" pageId="9" pageNumber="210" type="discussion">
<paragraph id="EA1036D9FFE9FF99EE861CE7FD0717A2" blockId="9.[151,1436,151,1365]" pageId="9" pageNumber="210">
<emphasis id="D8DBEACBFFE9FF99EE861CE7FEBB17EA" bold="true" box="[199,315,908,933]" pageId="9" pageNumber="210">Remarks.</emphasis>
Up to now it has not been possible to obtain specimens of both sexes in the same cave. On the basis of the obtained data, we consider the different populations to belong to the same species, but this remains to be verified by the collection of new specimens.
</paragraph>
<paragraph id="EA1036D9FFE9FF99EE861C9CFA19111B" blockId="9.[151,1436,151,1365]" pageId="9" pageNumber="210">
The new species shares with
<emphasis id="D8DBEACBFFE9FF99EC6C1C93FC8A105F" box="[557,778,1015,1040]" italics="true" pageId="9" pageNumber="210">
C. (E.)
<taxonomicName id="2DAF4D5AFFE9FF99ECCD1C9CFC8A105F" box="[652,778,1015,1040]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="cazorlensis">cazorlensis</taxonomicName>
</emphasis>
,
<emphasis id="D8DBEACBFFE9FF99ED5E1C93FC42105F" box="[799,962,1016,1040]" italics="true" pageId="9" pageNumber="210">
C. (E.)
<taxonomicName id="2DAF4D5AFFE9FF99ED3F1C92FC42105F" box="[894,962,1017,1040]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
,
<emphasis id="D8DBEACBFFE9FF99ED971C9CFAA4105F" box="[982,1316,1015,1040]" italics="true" pageId="9" pageNumber="210">
<taxonomicName id="2DAF4D5AFFE9FF99ED971C9CFBC9105F" box="[982,1097,1015,1040]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.) torremarinae
</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2012 (Almería,
<collectingCountry id="92B87649FFE9FF99EDFD1B77FC7D107A" box="[956,1021,1052,1077]" name="Spain" pageId="9" pageNumber="210">Spain</collectingCountry>
) (Carabajal Márquez
<emphasis id="D8DBEACBFFE9FF99EB461B76FAC6107A" box="[1287,1350,1052,1077]" italics="true" pageId="9" pageNumber="210">et al.</emphasis>
2012),
<emphasis id="D8DBEACBFFE9FF99EED61B54FEC21018" box="[151,322,1087,1112]" italics="true" pageId="9" pageNumber="210">
<taxonomicName id="2DAF4D5AFFE9FF99EED61B54FE8A1017" box="[151,266,1087,1112]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
</emphasis>
<taxonomicName id="2DAF4D5AFFE9FF99EF0F1B54FDC41017" authority="Beier, 1939" authorityName="Beier" authorityYear="1939" box="[334,580,1087,1112]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="ventalloi">
<emphasis id="D8DBEACBFFE9FF99EF0F1B54FE301017" box="[334,432,1087,1112]" italics="true" pageId="9" pageNumber="210">ventalloi</emphasis>
Beier, 1939
</taxonomicName>
(Valencia,
<collectingCountry id="92B87649FFE9FF99EC8F1B54FC8F1017" box="[718,783,1087,1112]" name="Spain" pageId="9" pageNumber="210">Spain</collectingCountry>
) (Zaragoza 1985a) and
<emphasis id="D8DBEACBFFE9FF99EA721B2BFB4A1017" box="[1075,1226,1088,1112]" italics="true" pageId="9" pageNumber="210">
C. (E.)
<taxonomicName id="2DAF4D5AFFE9FF99EAD01B2AFB4A1017" box="[1169,1226,1089,1112]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="verai">verai</taxonomicName>
</emphasis>
(Alicante,
<collectingCountry id="92B87649FFE9FF99EB121B54FA141017" box="[1363,1428,1087,1112]" name="Spain" pageId="9" pageNumber="210">Spain</collectingCountry>
) (Zaragoza 1985b) the following characteristics: eyeless, raised lamina present on movable chelal finger, without isolated subapical tooth on movable finger of chelicera, only two macrosetae on posterior margin of carapace and absence of preocular microsetae.
<emphasis id="D8DBEACBFFE9FF99EC4E1BC6FD4E108A" box="[527,718,1196,1221]" italics="true" pageId="9" pageNumber="210">
C. (E.)
<taxonomicName id="2DAF4D5AFFE9FF99EC221BC7FD4E108A" box="[611,718,1196,1221]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="espanyoli" status="sp. nov.">espanyoli</taxonomicName>
</emphasis>
<taxonomicNameLabel id="C3E857B0FFE9FF99EC971BC7FCB0108A" box="[726,816,1196,1221]" pageId="9" pageNumber="210" rank="species">
<emphasis id="D8DBEACBFFE9FF99EC971BC7FCB0108A" bold="true" box="[726,816,1196,1221]" pageId="9" pageNumber="210">sp. nov.</emphasis>
</taxonomicNameLabel>
shares with
<emphasis id="D8DBEACBFFE9FF99EDFE1BC6FBCD108B" box="[959,1101,1196,1220]" italics="true" pageId="9" pageNumber="210">
C. (E.)
<taxonomicName id="2DAF4D5AFFE9FF99EA521BC6FBCD108B" box="[1043,1101,1197,1220]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="verai">verai</taxonomicName>
</emphasis>
a long pedipalp femur (about 0.90–1.00 mm) and chela (about
<quantity id="2D579B3CFFE9FF99EC511BBBFDF610A8" box="[528,630,1232,1256]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="210" unit="mm" value="1.2">1.20 mm</quantity>
or more);
<emphasis id="D8DBEACBFFE9FF99ECB11BBBFC3110A7" box="[752,945,1231,1256]" italics="true" pageId="9" pageNumber="210">
C. (E.)
<taxonomicName id="2DAF4D5AFFE9FF99ED041BA4FC3110A7" box="[837,945,1231,1256]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="espanyoli">espanyoli</taxonomicName>
</emphasis>
has an even slenderer chela (male 6.6–7.0, female 5.8–6.5) than
<emphasis id="D8DBEACBFFE9FF99EFC41B9EFD901143" box="[389,528,1268,1292]" italics="true" pageId="9" pageNumber="210">
C. (E.)
<taxonomicName id="2DAF4D5AFFE9FF99EF961B9EFD901143" box="[471,528,1269,1292]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="verai">verai</taxonomicName>
</emphasis>
(male 6.1, female 5.7–6.0);
<emphasis id="D8DBEACBFFE9FF99ED0C1B9EFB8B1142" box="[845,1035,1268,1293]" italics="true" pageId="9" pageNumber="210">
C. (E.)
<taxonomicName id="2DAF4D5AFFE9FF99EDDE1B9FFB8B1142" box="[927,1035,1268,1293]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="espanyoli">espanyoli</taxonomicName>
</emphasis>
has a higher number of teeth on the fixed and movable fingers, with both dental rows reaching proximad of trichobothrium
<emphasis id="D8DBEACBFFE9FF99EA271A7CFBFF117F" box="[1126,1151,1303,1328]" italics="true" pageId="9" pageNumber="210">sb</emphasis>
, as opposed to that of the fixed finger row being level with
<emphasis id="D8DBEACBFFE9FF99EC4E1A57FDA8111A" box="[527,552,1340,1365]" italics="true" pageId="9" pageNumber="210">sb</emphasis>
, with only that of the movable finger reaching proximad of
<emphasis id="D8DBEACBFFE9FF99EA881A57FB62111A" box="[1225,1250,1340,1365]" italics="true" pageId="9" pageNumber="210">sb</emphasis>
in
<emphasis id="D8DBEACBFFE9FF99EB471A56FA12111B" box="[1286,1426,1340,1364]" italics="true" pageId="9" pageNumber="210">
C. (E.)
<taxonomicName id="2DAF4D5AFFE9FF99EB191A56FA12111B" box="[1368,1426,1341,1364]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="verai">verai</taxonomicName>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="423739D9E06EFDAD16871807F7842877">Hypogean pseudoscorpions (Arachnida) from Jaén province (Andalusia, Spain), with descriptions of four new species and a new synonymy</mods:title>
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<mods:namePart id="80A6CDFC248C16F9A2D378618A15E031">Zaragoza, Juan A.</mods:namePart>
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<treatment id="620687CFFFE9FF9CEED61AF7FDAE17CF" ID-DOI="http://doi.org/10.5281/zenodo.6149888" ID-GBIF-Taxon="119574438" ID-Zenodo-Dep="6149888" LSID="urn:lsid:plazi:treatment:620687CFFFE9FF9CEED61AF7FDAE17CF" httpUri="http://treatment.plazi.org/id/620687CFFFE9FF9CEED61AF7FDAE17CF" lastPageId="12" lastPageNumber="213" pageId="9" pageNumber="210">
<subSubSection id="A2B56552FFE9FF99EED61AF7FEA71197" pageId="9" pageNumber="210" type="nomenclature">
<paragraph id="EA1036D9FFE9FF99EED61AF7FDB811F8" blockId="9.[151,568,1436,1496]" box="[151,568,1436,1463]" pageId="9" pageNumber="210">
<heading id="B15881B5FFE9FF99EED61AF7FDB811F8" bold="true" box="[151,568,1436,1463]" fontSize="11" level="1" pageId="9" pageNumber="210" reason="1">
<emphasis id="D8DBEACBFFE9FF99EED61AF7FDB811F8" bold="true" box="[151,568,1436,1463]" pageId="9" pageNumber="210">
<emphasis id="D8DBEACBFFE9FF99EED61AF7FE5411F9" bold="true" box="[151,468,1436,1462]" italics="true" pageId="9" pageNumber="210">
<taxonomicName id="2DAF4D5AFFE9FF99EED61AF7FE9611F9" ID-CoL="62LNY" box="[151,278,1436,1462]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
<taxonomicName id="2DAF4D5AFFE9FF99EF121AF7FE5411F9" box="[339,468,1436,1462]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="giennensis" status="sp. nov.">giennensis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="C3E857B0FFE9FF99EF9A1AF6FDB811F8" box="[475,568,1437,1463]" pageId="9" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="EA1036D9FFE9FF99EED61AD4FEA71197" blockId="9.[151,568,1436,1496]" box="[151,295,1471,1496]" pageId="9" pageNumber="210">
(
<figureCitation id="72942A5CFFE9FF99EEDE1AD4FE9F1197" box="[159,287,1471,1496]" captionStart-0="FIGURES 13 – 16" captionStart-1="FIGURES 17 – 20" captionStartId-0="10.[151,264,799,822]" captionStartId-1="11.[151,264,1952,1975]" captionTargetBox-0="[214,1364,209,778]" captionTargetBox-1="[192,1430,228,1917]" captionTargetId-0="figure@10.[207,1380,193,779]" captionTargetId-1="figure@11.[151,1436,193,1932]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 13 – 16. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 13, carapace; 14, partial view of anterior margin of carapace; 15, left chelicera; 16, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 14 – 16), 0.2 (Fig. 13)." captionText-1="FIGURES 17 – 20. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 17, right pedipalp femur, dorsal view; 18, right patella, dorsal view; 19, right chela, lateral view; 20, detail of teeth of right chela, lateral view. Scale bars (in mm): 0.05 (Fig. 20), 0.2 (Figs 17 – 19)." httpUri-0="https://zenodo.org/record/218273/files/figure.png" httpUri-1="https://zenodo.org/record/218274/files/figure.png" pageId="9" pageNumber="210">Figs 13–20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE9FF99EED6196CFD271227" pageId="9" pageNumber="210" type="materials_examined">
<paragraph id="EA1036D9FFE9FF99EED6196CFD271227" blockId="9.[151,1436,1543,2037]" pageId="9" pageNumber="210">
<emphasis id="D8DBEACBFFE9FF99EED6196CFECF126F" bold="true" box="[151,335,1543,1568]" pageId="9" pageNumber="210">
<typeStatus id="3514887BFFE9FF99EED6196CFF52126F" box="[151,210,1543,1568]" pageId="9" pageNumber="210">Type</typeStatus>
material.
</emphasis>
<typeStatus id="3514887BFFE9FF99EF1E196CFE47126F" box="[351,455,1543,1568]" pageId="9" pageNumber="210" type="holotype">Holotype</typeStatus>
♂,
<collectingCountry id="92B87649FFE9FF99EC44196CFDC8126F" box="[517,584,1543,1568]" name="Spain" pageId="9" pageNumber="210">Spain</collectingCountry>
, Andalusia, Jaén province, Peal de Becerro, Cueva de los Ladrones (37°51′2.39″N, 2°57′48.69″W;
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.a.s.l.),
<date id="9E111019FFE9FF99ECE11947FCA3120B" box="[672,803,1580,1604]" pageId="9" pageNumber="210" value="2012-06-03">03.VI.2012</date>
, lgt. G.E.V. (DEUA: 20011).
<typeStatus id="3514887BFFE9FF99EA2F1946FB50120A" box="[1134,1232,1581,1605]" pageId="9" pageNumber="210" type="paratype">Paratype</typeStatus>
tritonymph, same details as
<typeStatus id="3514887BFFE9FF99EF441924FEE71227" box="[261,359,1615,1640]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
(MCNB:
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MZB).
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE9FF99EE86191FFDB912FF" pageId="9" pageNumber="210" type="etymology">
<paragraph id="EA1036D9FFE9FF99EE86191FFDB912FF" blockId="9.[151,1436,1543,2037]" pageId="9" pageNumber="210">
<emphasis id="D8DBEACBFFE9FF99EE86191FFECE12C2" bold="true" box="[199,334,1652,1677]" pageId="9" pageNumber="210">Etymology.</emphasis>
Latin adjective derived from
<emphasis id="D8DBEACBFFE9FF99ECDD191EFD7312C3" box="[668,755,1653,1676]" italics="true" pageId="9" pageNumber="210">Giennis</emphasis>
, the Latin name for the Roman city of Jaén, which currently gives its name to the whole province.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE9FF99EE8619D7FC98132B" pageId="9" pageNumber="210" type="diagnosis">
<paragraph id="EA1036D9FFE9FF99EE8619D7FC98132B" blockId="9.[151,1436,1543,2037]" pageId="9" pageNumber="210">
<emphasis id="D8DBEACBFFE9FF99EE8619D7FEC1129A" bold="true" box="[199,321,1724,1749]" pageId="9" pageNumber="210">Diagnosis.</emphasis>
Facies weakly troglomorphic. No eyes or eye-spots. Chelicera with 6 setae on palm and 4–5 microsetae, movable cheliceral finger without isolated subapical tooth, male spinneret strongly reduced. Carapace with two macrosetae on posterior margin, without preocular microsetae. Pedipalp femur length
<quantity id="2D579B3CFFE9FF99EAA1186FFAC61353" box="[1248,1350,1796,1821]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="9" pageNumber="210" unit="mm" value="0.61">0.61 mm</quantity>
(5.8×); chela lateral length 0.87 (5.4×); fixed finger with 20 teeth, movable finger with 12 teeth and raised lamina with 10 rounded vestigial teeth; trichobothrium
<emphasis id="D8DBEACBFFE9FF99EC151826FDEF132B" box="[596,623,1869,1892]" italics="true" pageId="9" pageNumber="210">ist</emphasis>
level with
<emphasis id="D8DBEACBFFE9FF99ECAA1827FC90132A" box="[747,784,1868,1893]" italics="true" pageId="9" pageNumber="210">esb</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFE9FF9CEE861804FBB61683" lastPageId="12" lastPageNumber="213" pageId="9" pageNumber="210" type="description">
<paragraph id="EA1036D9FFE9FF99EE861804FF5513BA" blockId="9.[151,1436,1543,2037]" pageId="9" pageNumber="210">
<emphasis id="D8DBEACBFFE9FF99EE861804FED913C7" bold="true" box="[199,345,1903,1928]" pageId="9" pageNumber="210">Description.</emphasis>
<emphasis id="D8DBEACBFFE9FF99EF231804FD8513C7" box="[354,517,1903,1928]" italics="true" pageId="9" pageNumber="210">
Male
<typeStatus id="3514887BFFE9FF99EFE51804FD8513C7" box="[420,517,1903,1928]" pageId="9" pageNumber="210" type="holotype">holotype</typeStatus>
</emphasis>
. Small, weakly troglomorphic hypogean species; integument depigmented, pale brown; marked hispid granulation on lateral surface of ocular and posterior areas of carapace, on cheliceral palm, on base of paraxial face of pedipalpal femur, on base of movable finger and on dorsodistal surface of pedipalpal hand.
</paragraph>
<caption id="BED06651FFEAFF9AEED61C74FF76173C" httpUri="https://zenodo.org/record/218273/files/figure.png" pageId="10" pageNumber="211" targetBox="[214,1364,209,778]" targetPageId="10">
<paragraph id="EA1036D9FFEAFF9AEED61C74FF76173C" blockId="10.[151,1436,799,883]" pageId="10" pageNumber="211">
<emphasis id="D8DBEACBFFEAFF9AEED61C74FED3177A" bold="true" box="[151,339,799,822]" pageId="10" pageNumber="211">FIGURES 13–16.</emphasis>
<taxonomicName id="2DAF4D5AFFEAFF9AEF191C4BFC871779" box="[344,775,800,822]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="giennensis" status="sp. nov." subGenus="Ephippiochthonius">
<emphasis id="D8DBEACBFFEAFF9AEF191C4BFC871779" box="[344,775,800,822]" italics="true" pageId="10" pageNumber="211">Chthonius (Ephippiochthonius) giennensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E857B0FFEAFF9AED4C1C4BFCDD177A" box="[781,861,800,821]" pageId="10" pageNumber="211" rank="species">
<emphasis id="D8DBEACBFFEAFF9AED4C1C4BFCDD177A" bold="true" box="[781,861,800,821]" pageId="10" pageNumber="211">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype.
<emphasis id="D8DBEACBFFEAFF9AEA441C4BFB9E177A" bold="true" box="[1029,1054,800,821]" pageId="10" pageNumber="211">13</emphasis>
, carapace;
<emphasis id="D8DBEACBFFEAFF9AEACE1C4BFB28177A" bold="true" box="[1167,1192,800,821]" pageId="10" pageNumber="211">14</emphasis>
, partial view of anterior margin of carapace;
<emphasis id="D8DBEACBFFEAFF9AEF2A1C54FE04171B" bold="true" box="[363,388,831,852]" pageId="10" pageNumber="211">15</emphasis>
, left chelicera;
<emphasis id="D8DBEACBFFEAFF9AEC661C54FDC0171B" bold="true" box="[551,576,831,852]" pageId="10" pageNumber="211">16</emphasis>
, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 14–16), 0.2 (Fig. 13).
</paragraph>
</caption>
<paragraph id="EA1036D9FFEAFF9AEE861CCBFD26106B" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">
Carapace (
<figureCitation id="72942A5CFFEAFF9AEF031CCBFE1617F7" box="[322,406,928,953]" captionStart="FIGURES 13 – 16" captionStartId="10.[151,264,799,822]" captionTargetBox="[214,1364,209,778]" captionTargetId="figure@10.[207,1380,193,779]" captionTargetPageId="10" captionText="FIGURES 13 – 16. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 13, carapace; 14, partial view of anterior margin of carapace; 15, left chelicera; 16, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 14 – 16), 0.2 (Fig. 13)." httpUri="https://zenodo.org/record/218273/files/figure.png" pageId="10" pageNumber="211">Fig. 13</figureCitation>
) slightly longer than broad, slightly constricted posteriorly, subquadrate; anterior margin (
<figureCitation id="72942A5CFFEAFF9AEEDE1CAFFF711792" box="[159,241,964,989]" captionStart="FIGURES 13 – 16" captionStartId="10.[151,264,799,822]" captionTargetBox="[214,1364,209,778]" captionTargetId="figure@10.[207,1380,193,779]" captionTargetPageId="10" captionText="FIGURES 13 – 16. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 13, carapace; 14, partial view of anterior margin of carapace; 15, left chelicera; 16, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 14 – 16), 0.2 (Fig. 13)." httpUri="https://zenodo.org/record/218273/files/figure.png" pageId="10" pageNumber="211">Fig. 14</figureCitation>
) prominent, medially dentate and with rudimentary epistome; without eyes or eye-spots; 18 setae, without preocular microsetae, formula: 4:6:4:2:2, anteromedial seta
<quantity id="2D579B3CFFEAFF9AED721C83FC15104F" box="[819,917,1000,1024]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="211" unit="mm" value="0.11">0.11 mm</quantity>
long, ocular seta
<quantity id="2D579B3CFFEAFF9AEA1B1C83FB40104F" box="[1114,1216,1000,1024]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="211" unit="mm" value="0.09">0.09 mm</quantity>
. Five lyrifissures in anterior and ocular areas, one in posterior area.
</paragraph>
<paragraph id="EA1036D9FFEAFF9AEE861B5BFB041022" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">
Pedipalpal coxa 5 setae (including 2 on manducatory process), coxa
<date id="9E111019FFEAFF9AEDA91B5AFB891007" box="[1000,1033,1072,1096]" pageId="10" pageNumber="211">I 3</date>
+ 3 marginal microsetae,
<date id="9E111019FFEAFF9AEB7F1B5AFAE91007" box="[1342,1385,1072,1096]" pageId="10" pageNumber="211">II 4</date>
+ 9 bipinnate coxal spines,
<date id="9E111019FFEAFF9AEFDF1B3EFE4F1022" box="[414,463,1108,1133]" pageId="10" pageNumber="211">III 5</date>
+ 5 bipinnate coxal spines,
<date id="9E111019FFEAFF9AED491B3EFCBD1022" box="[776,829,1108,1133]" pageId="10" pageNumber="211">IV 6</date>
; intercoxal tubercle bisetose.
</paragraph>
<paragraph id="EA1036D9FFEAFF9AEE861B13FC591097" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">Chaetotaxy of tergites I–XI: 4:4:4:4:6:6:6:6:6:4:1T2T1. Genital area with 9 setae on sternite II; sternite III with genital opening flanked by 7 setae on each side, with 4+4 internal glandular setae and (3)8(3) marginal setae. Chaetotaxy of sternites IV–XI: (2)7(2):8:6:6:6:6:6:0. Anal cone 0+2 setae.</paragraph>
<paragraph id="EA1036D9FFEAFF9AEE861B8FFBF111C2" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">
Chelicera (
<figureCitation id="72942A5CFFEAFF9AEF071B8FFE4B10B2" box="[326,459,1252,1277]" captionStart="FIGURES 13 – 16" captionStartId="10.[151,264,799,822]" captionTargetBox="[214,1364,209,778]" captionTargetId="figure@10.[207,1380,193,779]" captionTargetPageId="10" captionText="FIGURES 13 – 16. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 13, carapace; 14, partial view of anterior margin of carapace; 15, left chelicera; 16, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 14 – 16), 0.2 (Fig. 13)." httpUri="https://zenodo.org/record/218273/files/figure.png" pageId="10" pageNumber="211">Figs 15–16</figureCitation>
) with 6 setae on palm and 4–5 microsetae laterally, seta
<emphasis id="D8DBEACBFFEAFF9AEA3D1B8FFB1710B2" box="[1148,1175,1252,1277]" italics="true" pageId="10" pageNumber="211">vb</emphasis>
short,
<quantity id="2D579B3CFFEAFF9AEAAF1B8EFAD810B3" box="[1262,1368,1253,1277]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="211" unit="mm" value="0.04">0.04 mm</quantity>
long, microsetae
<quantity id="2D579B3CFFEAFF9AEF581A63FE3F116F" box="[281,447,1288,1312]" metricMagnitude="-5" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="10" pageNumber="211" unit="mm" value="0.025" valueMax="0.03" valueMin="0.02">0.02–0.03 mm</quantity>
long; seta
<emphasis id="D8DBEACBFFEAFF9AEC7E1A63FDD5116E" box="[575,597,1288,1313]" italics="true" pageId="10" pageNumber="211">gl</emphasis>
0.52 from base of movable finger. Fixed finger with 14 teeth, two distal teeth distinctly larger than others, proximally decreasing in size. Movable finger without an isolated subapical tooth (
<emphasis id="D8DBEACBFFEAFF9AEEA41A3BFF7B1126" box="[229,251,1360,1385]" italics="true" pageId="10" pageNumber="211">di</emphasis>
), a large distal tooth and 9 contiguous teeth reduced in size proximally; spinneret strongly reduced. Rallum with 11 blades, serrulae exterior and interior with 16 and 14 blades respectively.
</paragraph>
<paragraph id="EA1036D9FFEAFF9AEE861AF3FA0A12BB" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">
Pedipalp (
<figureCitation id="72942A5CFFEAFF9AEF7B1AF3FE3E11FF" box="[314,446,1432,1457]" captionStart="FIGURES 17 – 20" captionStartId="11.[151,264,1952,1975]" captionTargetBox="[192,1430,228,1917]" captionTargetId="figure@11.[151,1436,193,1932]" captionTargetPageId="11" captionText="FIGURES 17 – 20. Chthonius (Ephippiochthonius) giennensis sp. nov., male holotype. 17, right pedipalp femur, dorsal view; 18, right patella, dorsal view; 19, right chela, lateral view; 20, detail of teeth of right chela, lateral view. Scale bars (in mm): 0.05 (Fig. 20), 0.2 (Figs 17 – 19)." httpUri="https://zenodo.org/record/218274/files/figure.png" pageId="10" pageNumber="211">Figs 17–20</figureCitation>
) with femoral chaetotaxy 3:6:3:5:1, four lyrifissures: one antiaxial and three paraxial. Chelal hand markedly depressed at level of
<emphasis id="D8DBEACBFFEAFF9AECC01AD7FD41119A" box="[641,705,1468,1493]" italics="true" pageId="10" pageNumber="211">ib-isb</emphasis>
, with a rounded short hump distad of
<emphasis id="D8DBEACBFFEAFF9AEA2A1AD7FB2B119A" box="[1131,1195,1468,1493]" italics="true" pageId="10" pageNumber="211">ib-isb</emphasis>
and a distinct ventral hollow just before base of movable chelal finger; chaetotaxy 4:5:4. Fixed finger with 20 pointed teeth and one rounded proximal tooth, all with dental canals except the two small distal ones and proximal one; third distal tooth from tip modified, a little wider than the others; four microtubercles at base; tip of finger with an accessory tooth (
<emphasis id="D8DBEACBFFEAFF9AEEE11927FF36122A" box="[160,182,1612,1637]" italics="true" pageId="10" pageNumber="211">td</emphasis>
) on antiaxial face; five teeth at level of
<emphasis id="D8DBEACBFFEAFF9AEC341926FD2C122B" box="[629,684,1613,1636]" italics="true" pageId="10" pageNumber="211">est/it</emphasis>
occupying
<quantity id="2D579B3CFFEAFF9AED6E1926FC07122B" box="[815,903,1613,1637]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="211" unit="mm" value="0.1">0.1 mm</quantity>
, distance between apices
<quantity id="2D579B3CFFEAFF9AEAE61926FA9C122B" box="[1191,1308,1613,1637]" metricMagnitude="-5" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="211" unit="mm" value="0.024">0.024 mm</quantity>
. Distal half of movable finger with 12 pointed teeth with dental canals, two distal ones tiny and the proximal one reduced in size; proximal half of finger with slightly raised lamina with 10 rounded vestigial teeth reaching slightly proximad o trichobothrium
<emphasis id="D8DBEACBFFEAFF9AEF1D19D3FEF5129E" box="[348,373,1720,1745]" italics="true" pageId="10" pageNumber="211">sb</emphasis>
, level with coupled sensilla
<emphasis id="D8DBEACBFFEAFF9AECF719D2FD51129F" box="[694,721,1721,1744]" italics="true" pageId="10" pageNumber="211">pc</emphasis>
; basal apodeme long and apically indented. Trichobothria as in figs 18–19;
<emphasis id="D8DBEACBFFEAFF9AEF5D19B6FEB712BB" box="[284,311,1757,1780]" italics="true" pageId="10" pageNumber="211">ist</emphasis>
level with
<emphasis id="D8DBEACBFFEAFF9AEFF519B7FE5912BA" box="[436,473,1756,1781]" italics="true" pageId="10" pageNumber="211">esb</emphasis>
, both forming an obtuse angle with
<emphasis id="D8DBEACBFFEAFF9AED2E19B7FC0A12BA" box="[879,906,1756,1781]" italics="true" pageId="10" pageNumber="211">eb</emphasis>
; distance
<emphasis id="D8DBEACBFFEAFF9AEDBA19B7FBB512BA" box="[1019,1077,1756,1781]" italics="true" pageId="10" pageNumber="211">st–sb</emphasis>
2.1 longer than distance
<emphasis id="D8DBEACBFFEAFF9AEB0F19B7FA0312BA" box="[1358,1411,1756,1781]" italics="true" pageId="10" pageNumber="211">sb–b</emphasis>
.
</paragraph>
<paragraph id="EA1036D9FFEAFF9AEE86186AFC491357" blockId="10.[151,1437,928,2033]" box="[199,969,1792,1817]" pageId="10" pageNumber="211">Leg IV. Tactile seta ratios: tibia 0.57, basitarsus 0.36 and tarsus 0.28.</paragraph>
<paragraph id="EA1036D9FFEAFF9AEE86184EFB0913BF" blockId="10.[151,1437,928,2033]" pageId="10" pageNumber="211">
<emphasis id="D8DBEACBFFEAFF9AEE86184EFE731373" box="[199,499,1828,1853]" italics="true" pageId="10" pageNumber="211">Measurements and ratios.</emphasis>
Body 1.48. Carapace 0.48/0.44 (1.1). Chelicera: palm 0.41/0.19 (2.1), movable finger 0.21. Pedipalp: femur 0.61/0.11 (5.8); patella 0.27/0.13 (2.1); chela 0.87/0.17 (5.3), depth 0.16 (5.4); hand 0.34 (2.1), length/depth (2.1); movable finger 0.52; ratio movable finger/hand 1.5; ratio femur/movable finger 1.2; ratio femur/carapace 1.3; ratio chela/carapace 1.8. Leg I: femur 0.35/0.06 (5.5); patella 0.17/0.06 (2.9); tibia 0.20/ 0.05 (4.3); tarsus 0.39/0.04 (10.4); ratio femur/patella 2.1. Leg IV: femur+patella 0.54/0.22 (2.5); tibia 0.38/0.08 (4.7); basitarsus 0.19/0.06 (3.1); telotarsus 0.39/0.04 (10.1); ratio telotarsus/basitarsus 2.0.
</paragraph>
<caption id="BED06651FFEBFF9BEED618CBFE5E13BB" httpUri="https://zenodo.org/record/218274/files/figure.png" pageId="11" pageNumber="212" targetBox="[192,1430,228,1917]" targetPageId="11">
<paragraph id="EA1036D9FFEBFF9BEED618CBFE5E13BB" blockId="11.[151,1436,1952,2036]" pageId="11" pageNumber="212">
<emphasis id="D8DBEACBFFEBFF9BEED618CBFED513F9" bold="true" box="[151,341,1952,1975]" pageId="11" pageNumber="212">FIGURES 17–20.</emphasis>
<taxonomicName id="2DAF4D5AFFEBFF9BEF1D18CAFC8F13F8" box="[348,783,1953,1975]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="giennensis" status="sp. nov." subGenus="Ephippiochthonius">
<emphasis id="D8DBEACBFFEBFF9BEF1D18CAFC8F13F8" box="[348,783,1953,1975]" italics="true" pageId="11" pageNumber="212">Chthonius (Ephippiochthonius) giennensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E857B0FFEBFF9BED5618CAFCE813F9" box="[791,872,1953,1974]" pageId="11" pageNumber="212" rank="species">
<emphasis id="D8DBEACBFFEBFF9BED5618CAFCE813F9" bold="true" box="[791,872,1953,1974]" pageId="11" pageNumber="212">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype.
<emphasis id="D8DBEACBFFEBFF9BEA5618CAFBB013F9" bold="true" box="[1047,1072,1953,1974]" pageId="11" pageNumber="212">17</emphasis>
, right pedipalp femur, dorsal view;
<emphasis id="D8DBEACBFFEBFF9BEED618ABFF30139A" bold="true" box="[151,176,1984,2005]" pageId="11" pageNumber="212">18</emphasis>
, right patella, dorsal view;
<emphasis id="D8DBEACBFFEBFF9BEF8C18ABFE66139A" bold="true" box="[461,486,1984,2005]" pageId="11" pageNumber="212">19</emphasis>
, right chela, lateral view;
<emphasis id="D8DBEACBFFEBFF9BECB618ABFC90139A" bold="true" box="[759,784,1984,2005]" pageId="11" pageNumber="212">20</emphasis>
, detail of teeth of right chela, lateral view. Scale bars (in mm): 0.05 (Fig. 20), 0.2 (Figs 17–19).
</paragraph>
</caption>
<paragraph id="EA1036D9FFECFF9CEE861FFCFBB61683" blockId="12.[151,1437,151,896]" pageId="12" pageNumber="213">
<emphasis id="D8DBEACBFFECFF9CEE861FFCFED714FF" bold="true" box="[199,343,151,176]" pageId="12" pageNumber="213">Tritonymph</emphasis>
(data coinciding with
<typeStatus id="3514887BFFECFF9CEC121FFCFD3514FF" box="[595,693,151,176]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
omitted). Carapace quadrate. Coxa
<date id="9E111019FFECFF9CEA061FF3FBE614FF" box="[1095,1126,152,176]" pageId="12" pageNumber="213">I 3</date>
+ 2 marginal microsetae,
<date id="9E111019FFECFF9CEBC81FF3FF25149B" pageId="12" pageNumber="213">II 4</date>
+ 7–8 bipinnate coxal spines,
<date id="9E111019FFECFF9CEC101FD6FD11149A" box="[593,657,188,213]" pageId="12" pageNumber="213">III 5</date>
+ 4 bipinnate coxal spines. Chaetotaxy of sternites II–XI: 5:(2)6(2):(1)6(1):6:6:6:6:6:7:0. Cheliceral palm with 5 setae and 4 microsetae laterally, seta
<emphasis id="D8DBEACBFFECFF9CEAF31FB4FB4814B7" box="[1202,1224,223,248]" italics="true" pageId="12" pageNumber="213">gl</emphasis>
0.61 from base of movable finger; fixed finger with 10 teeth, two distal teeth distinctly larger than the others; movable finger without an isolated subapical tooth (
<emphasis id="D8DBEACBFFECFF9CEF961E4CFE6C150F" box="[471,492,295,320]" italics="true" pageId="12" pageNumber="213">di</emphasis>
), a large distal tooth and 6 teeth reduced in size proximally; spinneret blunt toothshaped, more developed than in male
<typeStatus id="3514887BFFECFF9CEC091E27FD2E152A" box="[584,686,332,357]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
. Serrulae exterior and interior with 15 and 14 blades respectively. Chelal hand chaetotaxy 4–4–4; fixed finger with 14 pointed teeth and one rounded proximal tooth, all with dental canals except the two small distal ones and the proximal one; distal half of movable finger with 10 pointed teeth, two tiny distal ones and reduced proximal one without dental canals; proximal half of finger with slightly raised lamina with 9 rounded, vestigial teeth. Leg IV: tactile seta ratios: tibia 0.55, basitarsus 0.39, tarsus 0.28.
<emphasis id="D8DBEACBFFECFF9CEED61D6AFE361657" box="[151,438,511,536]" italics="true" pageId="12" pageNumber="213">Measurements and ratios</emphasis>
: Body 0.96. Carapace 0.33/0.33 (1.0). Chelicera: palm 0.30/0.14 (2.2), movable finger 0.14. Pedipalp: femur 0.38/0.09 (4.4); patella 0.19/0.10 (2.0); chela width and depth 0.62/0.12 (5.1); hand 0.25 (2.0), length/dept (2.0); movable finger 0.36; ratio movable finger/hand 1.5; ratio femur/movable finger 1.0; ratio femur/carapace 1.2, ratio chela/carapace 1.9. Leg I: femur 0.23/0.05 (4.5); patella 0.11/0.05 (2.4); tibia 0.12/0.04 (3.0); tarsus 0.25/0.03 (7.7); ratio femur/patella 2.0. Leg IV: femur+patella 0.36/0.14 (2.5); tibia 0.24/0.06 (4.0); basitarsus 0.12/0.05 (2.5); telotarsus 0.25/0.04 (7.1); ratio telotarsus/basitarsus 2.0.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFECFF9CEE861DBCFC221712" pageId="12" pageNumber="213" type="discussion">
<paragraph id="EA1036D9FFECFF9CEE861DBCFC221712" blockId="12.[151,1437,151,896]" pageId="12" pageNumber="213">
<emphasis id="D8DBEACBFFECFF9CEE861DBCFEBB16BF" bold="true" box="[199,315,727,752]" pageId="12" pageNumber="213">Remarks.</emphasis>
<emphasis id="D8DBEACBFFECFF9CEF031DB3FD8A16BF" box="[322,522,728,752]" italics="true" pageId="12" pageNumber="213">
C. (E.)
<taxonomicName id="2DAF4D5AFFECFF9CEFD51DB2FD8A16BF" box="[404,522,729,752]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="giennensis" status="sp. nov.">giennensis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="C3E857B0FFECFF9CEC501DBCFDEB16BF" box="[529,619,727,752]" pageId="12" pageNumber="213" rank="species">
<emphasis id="D8DBEACBFFECFF9CEC501DBCFDEB16BF" bold="true" box="[529,619,727,752]" pageId="12" pageNumber="213">sp. nov.</emphasis>
</taxonomicNameLabel>
shares with
<emphasis id="D8DBEACBFFECFF9CECB61DB3FC3516BF" box="[759,949,727,752]" italics="true" pageId="12" pageNumber="213">
C. (E.)
<taxonomicName id="2DAF4D5AFFECFF9CED0B1DBCFC3516BF" box="[842,949,727,752]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="espanyoli">espanyoli</taxonomicName>
</emphasis>
and the other five species mentioned in the discussion of the latter the same main characters, but differs from all of them in having 4–5 microsetae on the cheliceral palm and in the position of trichobothrium
<emphasis id="D8DBEACBFFECFF9CECB11C4AFC8B1777" box="[752,779,801,824]" italics="true" pageId="12" pageNumber="213">ist</emphasis>
level with
<emphasis id="D8DBEACBFFECFF9CEDC81C74FC2E1777" box="[905,942,799,824]" italics="true" pageId="12" pageNumber="213">esb</emphasis>
, as opposed to 1 microsetae (rarely 0–2) on the cheliceral palm and
<emphasis id="D8DBEACBFFECFF9CEFE31C2EFE3D1713" box="[418,445,837,860]" italics="true" pageId="12" pageNumber="213">ist</emphasis>
markedly distal of
<emphasis id="D8DBEACBFFECFF9CECD71C2FFD3B1712" box="[662,699,836,861]" italics="true" pageId="12" pageNumber="213">esb</emphasis>
in the other species.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFECFF9CEE861C0CFDAE17CF" box="[199,558,871,896]" pageId="12" pageNumber="213" type="distribution">
<paragraph id="EA1036D9FFECFF9CEE861C0CFDAE17CF" blockId="12.[151,1437,151,896]" box="[199,558,871,896]" pageId="12" pageNumber="213">
<emphasis id="D8DBEACBFFECFF9CEE861C0CFEE017CF" bold="true" box="[199,352,871,896]" pageId="12" pageNumber="213">Distribution.</emphasis>
Andalusia,
<collectingCountry id="92B87649FFECFF9CEFA71C0CFDAA17CF" box="[486,554,871,896]" name="Spain" pageId="12" pageNumber="213">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

340
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<treatment id="620687CFFFECFF9FEED61CA3FDAE162F" ID-DOI="http://doi.org/10.5281/zenodo.6149890" ID-GBIF-Taxon="119574436" ID-Zenodo-Dep="6149890" LSID="urn:lsid:plazi:treatment:620687CFFFECFF9FEED61CA3FDAE162F" httpUri="http://treatment.plazi.org/id/620687CFFFECFF9FEED61CA3FDAE162F" lastPageId="15" lastPageNumber="216" pageId="12" pageNumber="213">
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<paragraph id="EA1036D9FFECFF9CEED61CA3FDC617AC" blockId="12.[151,582,968,1029]" box="[151,582,968,995]" pageId="12" pageNumber="213">
<heading id="B15881B5FFECFF9CEED61CA3FDC617AC" bold="true" box="[151,582,968,995]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<emphasis id="D8DBEACBFFECFF9CEED61CA3FDC617AC" bold="true" box="[151,582,968,995]" pageId="12" pageNumber="213">
<emphasis id="D8DBEACBFFECFF9CEED61CA3FE6017AD" bold="true" box="[151,480,968,994]" italics="true" pageId="12" pageNumber="213">
<taxonomicName id="2DAF4D5AFFECFF9CEED61CA3FE9617AD" ID-CoL="62LNY" box="[151,278,968,994]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
<taxonomicName id="2DAF4D5AFFECFF9CEF121CA3FE6017AD" box="[339,480,968,994]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="villacarrillo" status="sp. nov.">villacarrillo</taxonomicName>
</emphasis>
<taxonomicNameLabel id="C3E857B0FFECFF9CEFA91CA2FDC617AC" box="[488,582,969,995]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="EA1036D9FFECFF9CEED61C87FEA7104A" blockId="12.[151,582,968,1029]" box="[151,295,1004,1029]" pageId="12" pageNumber="213">
(
<figureCitation id="72942A5CFFECFF9CEEDE1C87FE9F104A" box="[159,287,1004,1029]" captionStart-0="FIGURES 21 – 24" captionStart-1="FIGURES 25 – 26" captionStartId-0="13.[151,264,888,911]" captionStartId-1="14.[151,264,1952,1975]" captionTargetBox-0="[182,1415,210,858]" captionTargetBox-1="[178,1385,193,1909]" captionTargetId-0="figure@13.[157,1430,193,867]" captionTargetId-1="figure@14.[174,1413,193,1932]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 21 – 24. Chthonius (Ephippiochthonius) villacarrillo sp. nov., female holotype. 21, carapace; 22, partial view of anterior margin of carapace; 23, left chelicera; 24, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 22 – 24), 0.2 (Fig. 21)." captionText-1="FIGURES 25 – 26. Chthonius (Ephippiochthonius) villacarrillo sp. nov., female holotype. 25, left pedipalp, minus chela and trochanter, dorsal view; 26, left chela, lateral view. Scale bar 0.2 mm." httpUri-0="https://zenodo.org/record/218275/files/figure.png" httpUri-1="https://zenodo.org/record/218276/files/figure.png" pageId="12" pageNumber="213">Figs 21–26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFECFF9CEED61B5FFBAD103F" pageId="12" pageNumber="213" type="materials_examined">
<paragraph id="EA1036D9FFECFF9CEED61B5FFBAD103F" blockId="12.[151,1437,1076,2037]" pageId="12" pageNumber="213">
<emphasis id="D8DBEACBFFECFF9CEED61B5FFED21002" bold="true" box="[151,338,1076,1101]" pageId="12" pageNumber="213">
<typeStatus id="3514887BFFECFF9CEED61B5FFF521002" box="[151,210,1076,1101]" pageId="12" pageNumber="213">Type</typeStatus>
material.
</emphasis>
<typeStatus id="3514887BFFECFF9CEF261B5FFE4F1002" box="[359,463,1076,1101]" pageId="12" pageNumber="213" type="holotype">Holotype</typeStatus>
♀,
<collectingCountry id="92B87649FFECFF9CEC551B5FFDD71002" box="[532,599,1076,1101]" name="Spain" pageId="12" pageNumber="213">Spain</collectingCountry>
, Andalusia, Jaén province,
<taxonomicName id="2DAF4D5AFFECFF9CED801B5FFBCB1002" box="[961,1099,1076,1101]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="215" phylum="Arthropoda" rank="species" species="villacarrillo" status="sp. nov.">Villacarrillo</taxonomicName>
, Cueva de la Morciguilla (38°5′27.87″N, 3°0′5.62″W;
<specimenCount id="FCA9FD50FFECFF9CEF9A1B33FDAD103F" box="[475,557,1111,1136]" pageId="12" pageNumber="213" type="generic">
<quantity id="2D579B3CFFECFF9CEF9A1B33FDA9103F" box="[475,553,1111,1136]" metricMagnitude="2" metricUnit="m" metricValue="7.01" pageId="12" pageNumber="213" unit="m" value="701.0">701 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="9E111019FFECFF9CEC351B33FD6B103F" box="[628,747,1112,1136]" pageId="12" pageNumber="213" value="2013-02-24">24.II.2013</date>
, lgt. G.E.V. (DEUA: 20023).
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFECFF9CEE861B17FDE810F7" pageId="12" pageNumber="213" type="etymology">
<paragraph id="EA1036D9FFECFF9CEE861B17FDE810F7" blockId="12.[151,1437,1076,2037]" pageId="12" pageNumber="213">
<emphasis id="D8DBEACBFFECFF9CEE861B17FECE10DA" bold="true" box="[199,334,1148,1173]" pageId="12" pageNumber="213">Etymology.</emphasis>
This species is dedicated to the municipal area of
<taxonomicName id="2DAF4D5AFFECFF9CED3F1B17FB8510DA" box="[894,1029,1148,1173]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="215" phylum="Arthropoda" rank="species" species="villacarrillo" status="sp. nov.">Villacarrillo</taxonomicName>
where the
<typeStatus id="3514887BFFECFF9CEAC11B16FB3010DA" box="[1152,1200,1149,1173]" pageId="12" pageNumber="213">type</typeStatus>
locality is found and home town of G.E.V.; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFECFF9CEE861BAFFCB31123" pageId="12" pageNumber="213" type="diagnosis">
<paragraph id="EA1036D9FFECFF9CEE861BAFFCB31123" blockId="12.[151,1437,1076,2037]" pageId="12" pageNumber="213">
<emphasis id="D8DBEACBFFECFF9CEE861BAFFEC11092" bold="true" box="[199,321,1220,1245]" pageId="12" pageNumber="213">Diagnosis.</emphasis>
Weakly troglomorphic facies. No eyes or eye-spots. Chelicera with 6 setae on palm and one microseta, movable cheliceral finger with isolated subapical tooth, female with distinct spinneret. Carapace with two macrosetae on posterior margin, without preocular microsetae. Pedipalp: femur length
<quantity id="2D579B3CFFECFF9CEADE1A67FA84116B" box="[1183,1284,1292,1317]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="12" pageNumber="213" unit="mm" value="0.56">0.56 mm</quantity>
(5.5×); chela length 0.78 (5.18×); fixed finger with 19 teeth, movable finger with 12 teeth and raised lamina with 9 rounded vestigial teeth; trichobothrium
<emphasis id="D8DBEACBFFECFF9CEFB01A3EFD8B1123" box="[497,523,1365,1388]" italics="true" pageId="12" pageNumber="213">ist</emphasis>
markedly distal of
<emphasis id="D8DBEACBFFECFF9CECA41A3FFCAC1122" box="[741,812,1364,1389]" italics="true" pageId="12" pageNumber="213">eb/esb</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFECFF9DEE861A1CFB091220" lastPageId="13" lastPageNumber="214" pageId="12" pageNumber="213" type="description">
<paragraph id="EA1036D9FFECFF9CEE861A1CFED111B2" blockId="12.[151,1437,1076,2037]" pageId="12" pageNumber="213">
<emphasis id="D8DBEACBFFECFF9CEE861A1CFED911DF" bold="true" box="[199,345,1399,1424]" pageId="12" pageNumber="213">Description.</emphasis>
Small hypogean species; moderate troglomorphic adaptations; integument depigmented, pale yellowish-brown; weak hispid granulation on lateral surface of ocular and posterior areas of carapace, on cheliceral palm, on base of paraxial face of pedipalpal femur, on base of movable chelal finger and on dorsodistal surface of pedipalpal hand.
</paragraph>
<paragraph id="EA1036D9FFECFF9CEE861963FCC812C3" blockId="12.[151,1437,1076,2037]" pageId="12" pageNumber="213">
Carapace (
<figureCitation id="72942A5CFFECFF9CEF00196CFE16126F" box="[321,406,1543,1568]" captionStart="FIGURES 21 – 24" captionStartId="13.[151,264,888,911]" captionTargetBox="[182,1415,210,858]" captionTargetId="figure@13.[157,1430,193,867]" captionTargetPageId="13" captionText="FIGURES 21 – 24. Chthonius (Ephippiochthonius) villacarrillo sp. nov., female holotype. 21, carapace; 22, partial view of anterior margin of carapace; 23, left chelicera; 24, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 22 – 24), 0.2 (Fig. 21)." httpUri="https://zenodo.org/record/218275/files/figure.png" pageId="12" pageNumber="213">Fig. 21</figureCitation>
) as broad as long, slightly constricted posteriorly, subquadrate; anterior margin (
<figureCitation id="72942A5CFFECFF9CEB7E196CFA13126F" box="[1343,1427,1543,1568]" captionStart="FIGURES 21 – 24" captionStartId="13.[151,264,888,911]" captionTargetBox="[182,1415,210,858]" captionTargetId="figure@13.[157,1430,193,867]" captionTargetPageId="13" captionText="FIGURES 21 – 24. Chthonius (Ephippiochthonius) villacarrillo sp. nov., female holotype. 21, carapace; 22, partial view of anterior margin of carapace; 23, left chelicera; 24, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 22 – 24), 0.2 (Fig. 21)." httpUri="https://zenodo.org/record/218275/files/figure.png" pageId="12" pageNumber="213">Fig. 22</figureCitation>
) prominent, medially dentate and with rudimentary epistome; without eyes or eye-spots; chaetotaxy: 18 setae, without preocular microsetae, formula: 4:6:4:2:2, anteromedial seta
<quantity id="2D579B3CFFECFF9CEDE0193BFB951227" box="[929,1045,1616,1640]" metricMagnitude="-5" metricUnit="m" metricValue="8.3" pageId="12" pageNumber="213" unit="mm" value="0.083">0.083 mm</quantity>
long, ocular seta
<quantity id="2D579B3CFFECFF9CEAA21924FADA1227" box="[1251,1370,1615,1640]" metricMagnitude="-5" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="213" unit="mm" value="0.055">0.055 mm</quantity>
. Four lyrifissures in anterior and ocular areas, two in posterior area.
</paragraph>
<paragraph id="EA1036D9FFECFF9CEE8619FCFEDA12B7" blockId="12.[151,1437,1076,2037]" pageId="12" pageNumber="213">
Pedipalpal coxa 5 setae (including 2 on manducatory process), coxa
<date id="9E111019FFECFF9CED9319F2FC7212FF" box="[978,1010,1688,1712]" pageId="12" pageNumber="213">I 3</date>
+ 3 marginal microsetae,
<date id="9E111019FFECFF9CEB5D19F2FAC512FF" box="[1308,1349,1688,1712]" pageId="12" pageNumber="213">II 4</date>
+ 9–11 bipinnate coxal spines,
<date id="9E111019FFECFF9CEFE619D6FE5B129A" box="[423,475,1724,1749]" pageId="12" pageNumber="213">III 5</date>
+ about 4 (coxae strongly constricted) bipinnate coxal spines and
<date id="9E111019FFECFF9CEAA419D6FA9C129A" box="[1253,1308,1724,1749]" pageId="12" pageNumber="213">IV 6</date>
; intercoxal tubercle bisetose.
</paragraph>
<paragraph id="EA1036D9FFECFF9CEE86186FFC0A130F" blockId="12.[151,1437,1076,2037]" pageId="12" pageNumber="213">Chaetotaxy of tergites I–XI: 4:4:4:4:6:6:6:6:6:4:1T2T1. Genital area with 9 setae on sternite II. Chaetotaxy of sternites III–XI: (3)8(3):(2)7(2):8:6:6:6:6:7:0. Anal cone 0+2 setae.</paragraph>
<paragraph id="EA1036D9FFECFF9CEE861827FBF113BA" blockId="12.[151,1437,1076,2037]" pageId="12" pageNumber="213">
Chelicera (
<figureCitation id="72942A5CFFECFF9CEF051827FE48132B" box="[324,456,1868,1893]" captionStart="FIGURES 21 – 24" captionStartId="13.[151,264,888,911]" captionTargetBox="[182,1415,210,858]" captionTargetId="figure@13.[157,1430,193,867]" captionTargetPageId="13" captionText="FIGURES 21 – 24. Chthonius (Ephippiochthonius) villacarrillo sp. nov., female holotype. 21, carapace; 22, partial view of anterior margin of carapace; 23, left chelicera; 24, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 22 – 24), 0.2 (Fig. 21)." httpUri="https://zenodo.org/record/218275/files/figure.png" pageId="12" pageNumber="213">Figs 23–24</figureCitation>
) with 6 setae on palm and 1 microseta laterally, seta
<emphasis id="D8DBEACBFFECFF9CEA7A1827FBD6132A" box="[1083,1110,1868,1893]" italics="true" pageId="12" pageNumber="213">vb</emphasis>
very short,
<quantity id="2D579B3CFFECFF9CEAA51827FAD9132B" box="[1252,1369,1868,1893]" metricMagnitude="-5" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="213" unit="mm" value="0.025">0.025 mm</quantity>
long, microseta
<quantity id="2D579B3CFFECFF9CEF4F181BFEF513C7" box="[270,373,1904,1928]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="213" unit="mm" value="0.02">0.02 mm</quantity>
long; seta
<emphasis id="D8DBEACBFFECFF9CEFB61804FD8D13C7" box="[503,525,1903,1928]" italics="true" pageId="12" pageNumber="213">gl</emphasis>
0.53 from base of movable finger. Fixed finger with 11 teeth, two distal teeth distinctly larger than others, proximally decreasing in size. Movable finger with an isolated subapical tooth (
<emphasis id="D8DBEACBFFECFF9CEB2318FFFAF813E2" box="[1378,1400,1940,1965]" italics="true" pageId="12" pageNumber="213">di</emphasis>
), a large distal tooth and 7 contiguous teeth reduced in size proximally; spinneret prominent and apically rounded. Rallum with 11 blades, serrulae exterior and interior with 15 and 13 blades respectively.
</paragraph>
<caption id="BED06651FFEDFF9DEED61C13FE9C1784" httpUri="https://zenodo.org/record/218275/files/figure.png" pageId="13" pageNumber="214" targetBox="[182,1415,210,858]" targetPageId="13">
<paragraph id="EA1036D9FFEDFF9DEED61C13FE9C1784" blockId="13.[151,1436,888,971]" pageId="13" pageNumber="214">
<emphasis id="D8DBEACBFFEDFF9DEED61C13FED617C2" bold="true" box="[151,342,888,911]" pageId="13" pageNumber="214">FIGURES 21–24.</emphasis>
<taxonomicName id="2DAF4D5AFFEDFF9DEF1E1C13FCA417C1" box="[351,804,888,910]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="villacarrillo" status="sp. nov." subGenus="Ephippiochthonius">
<emphasis id="D8DBEACBFFEDFF9DEF1E1C13FCA417C1" box="[351,804,888,910]" italics="true" pageId="13" pageNumber="214">Chthonius (Ephippiochthonius) villacarrillo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E857B0FFEDFF9DED6F1C12FC0117C1" box="[814,897,889,910]" pageId="13" pageNumber="214" rank="species">
<emphasis id="D8DBEACBFFEDFF9DED6F1C12FC0117C1" bold="true" box="[814,897,889,910]" pageId="13" pageNumber="214">sp. nov.</emphasis>
</taxonomicNameLabel>
, female holotype.
<emphasis id="D8DBEACBFFEDFF9DEA091C13FBE117C2" bold="true" box="[1096,1121,888,909]" pageId="13" pageNumber="214">21</emphasis>
, carapace;
<emphasis id="D8DBEACBFFEDFF9DEA981C13FB7217C2" bold="true" box="[1241,1266,888,909]" pageId="13" pageNumber="214">22</emphasis>
, partial view of anterior margin of carapace;
<emphasis id="D8DBEACBFFEDFF9DEFF81CFCFE5217E3" bold="true" box="[441,466,919,940]" pageId="13" pageNumber="214">23</emphasis>
, left chelicera;
<emphasis id="D8DBEACBFFEDFF9DEC2F1CFCFD0717E3" bold="true" box="[622,647,919,940]" pageId="13" pageNumber="214">24</emphasis>
, partial view of fingers of left chelicera. Scale bars (in mm): 0.1 (Figs 22–24), 0.2 (Fig. 21).
</paragraph>
</caption>
<paragraph id="EA1036D9FFEDFF9DEE861C98FE4F113C" blockId="13.[151,1437,1011,2008]" pageId="13" pageNumber="214">
Pedipalp (
<figureCitation id="72942A5CFFEDFF9DEF7B1C98FE3E1043" box="[314,446,1011,1036]" captionStart="FIGURES 25 – 26" captionStartId="14.[151,264,1952,1975]" captionTargetBox="[178,1385,193,1909]" captionTargetId="figure@14.[174,1413,193,1932]" captionTargetPageId="14" captionText="FIGURES 25 – 26. Chthonius (Ephippiochthonius) villacarrillo sp. nov., female holotype. 25, left pedipalp, minus chela and trochanter, dorsal view; 26, left chela, lateral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/218276/files/figure.png" pageId="13" pageNumber="214">Figs 25–26</figureCitation>
) with femoral chaetotaxy 3:6:3:5:1, four lyrifissures: one antiaxial and three paraxial. Chelal hand depressed at level of
<emphasis id="D8DBEACBFFEDFF9DEC661B7CFDE7107F" box="[551,615,1047,1072]" italics="true" pageId="13" pageNumber="214">ib-isb</emphasis>
, with a rounded hump, slightly abrupt, distad of
<emphasis id="D8DBEACBFFEDFF9DEAED1B7CFB6C107F" box="[1196,1260,1047,1072]" italics="true" pageId="13" pageNumber="214">ib-isb</emphasis>
, and a marked ventral hollow just before base of movable chelal finger; chaetotaxy 4:5:4. Fixed finger with 18 pointed teeth and one rounded proximal tooth, all with dental canals except the two small distal ones; third distal tooth from tip modified, a little wider than others; four microtubercles at base; tip of finger with an accessory tooth (
<emphasis id="D8DBEACBFFEDFF9DEB141BE8FAEB10D3" box="[1365,1387,1155,1180]" italics="true" pageId="13" pageNumber="214">td</emphasis>
) on antiaxial face; six teeth at level of
<emphasis id="D8DBEACBFFEDFF9DEC5E1BC3FDD510F0" box="[543,597,1192,1215]" italics="true" pageId="13" pageNumber="214">est/it</emphasis>
occupying
<quantity id="2D579B3CFFEDFF9DEC9B1BCCFCB310F0" box="[730,819,1191,1215]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="214" unit="mm" value="0.1">0.1 mm</quantity>
, distance between apices
<quantity id="2D579B3CFFEDFF9DEA161BCCFA9A10F0" box="[1111,1306,1191,1216]" metricMagnitude="-5" metricUnit="m" metricValue="1.65" metricValueMax="1.7999999999999998" metricValueMin="1.5" pageId="13" pageNumber="214" unit="mm" value="0.0165" valueMax="0.018" valueMin="0.015">0.015–0.018 mm</quantity>
. Distal half of movable finger with 12 pointed teeth with dental canals, except distal tiny one and three proximal ones, decreasing in size; proximal half of finger with slightly raised lamina with 9 rounded vestigial teeth reaching halfway between trichobothria
<emphasis id="D8DBEACBFFEDFF9DEFB51A78FD8D1163" box="[500,525,1299,1324]" italics="true" pageId="13" pageNumber="214">sb</emphasis>
and
<emphasis id="D8DBEACBFFEDFF9DEC051A78FDD21163" box="[580,594,1299,1324]" italics="true" pageId="13" pageNumber="214">b</emphasis>
; coupled sensilla
<emphasis id="D8DBEACBFFEDFF9DED5A1A7FFCB61164" box="[795,822,1300,1323]" italics="true" pageId="13" pageNumber="214">pc</emphasis>
between
<emphasis id="D8DBEACBFFEDFF9DEDE01A78FC3A1163" box="[929,954,1299,1324]" italics="true" pageId="13" pageNumber="214">sb</emphasis>
and
<emphasis id="D8DBEACBFFEDFF9DEDB11A78FC7E1163" box="[1008,1022,1299,1324]" italics="true" pageId="13" pageNumber="214">b</emphasis>
, slightly closer to
<emphasis id="D8DBEACBFFEDFF9DEA8F1A78FB671163" box="[1230,1255,1299,1324]" italics="true" pageId="13" pageNumber="214">sb</emphasis>
; basal apodeme long and apically indented. Trichobothria as in fig. 25;
<emphasis id="D8DBEACBFFEDFF9DED571A53FCB11100" box="[790,817,1336,1359]" italics="true" pageId="13" pageNumber="214">ist</emphasis>
markedly distal of
<emphasis id="D8DBEACBFFEDFF9DEA551A5CFBB9111F" box="[1044,1081,1335,1360]" italics="true" pageId="13" pageNumber="214">esb</emphasis>
, distance
<emphasis id="D8DBEACBFFEDFF9DEAEE1A53FB491100" box="[1199,1225,1336,1359]" italics="true" pageId="13" pageNumber="214">ist</emphasis>
to
<emphasis id="D8DBEACBFFEDFF9DEAB31A5CFA97111F" box="[1266,1303,1335,1360]" italics="true" pageId="13" pageNumber="214">esb</emphasis>
<quantity id="2D579B3CFFEDFF9DEB601A5CFA171100" box="[1313,1431,1335,1360]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="214" unit="mm" value="0.035">0.035 mm</quantity>
, distance
<emphasis id="D8DBEACBFFEDFF9DEEBB1A30FE9F113B" box="[250,287,1371,1396]" italics="true" pageId="13" pageNumber="214">esb</emphasis>
to
<emphasis id="D8DBEACBFFEDFF9DEF021A30FEDE113B" box="[323,350,1371,1396]" italics="true" pageId="13" pageNumber="214">eb</emphasis>
<quantity id="2D579B3CFFEDFF9DEF251A37FE4B113C" box="[356,459,1372,1396]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="214" unit="mm" value="0.02">0.02 mm</quantity>
.
</paragraph>
<paragraph id="EA1036D9FFEDFF9DEE861AEBFC2111D8" blockId="13.[151,1437,1011,2008]" box="[199,929,1407,1432]" pageId="13" pageNumber="214">Leg IV. Tactile seta ratios: tibia 0.53, basitarsus 0.43, tarsus 0.33.</paragraph>
<paragraph id="EA1036D9FFEDFF9DEE861ACFFB091220" blockId="13.[151,1437,1011,2008]" pageId="13" pageNumber="214">
<emphasis id="D8DBEACBFFEDFF9DEE861ACFFE7311F4" box="[199,499,1443,1468]" italics="true" pageId="13" pageNumber="214">Measurements and ratios.</emphasis>
Body 1.30. Carapace 0.41/0.41 (1.0). Chelicera: palm 0.36/0.17 (2.1), movable finger 0.19. Pedipalp: femur 0.56/0.10 (5.5); patella 0.23/0.11 (2.0); chela width and depth 0.78/0.15 (5.2); hand 0.32 (2.1), length/depth (2.1); movable finger 0.45; ratio movable finger/hand 1.4; ratio femur/movable finger 1.3; ratio femur/carapace 1.4, ratio chela/carapace 1.9. Leg I: femur 0.29/0.05 (5.4); patella 0.15/0.05 (3.1); tibia 0.17/ 0.04 (4.1); tarsus 0.31/0.03 (9.2); ratio femur/patella 2.0. Leg IV: femur+patella 0.43/0.16 (2.7); tibia 0.28/0.07 (4.1); basitarsus 0.15/0.05 (3.0); telotarsus 0.30/0.03 (10.0); ratio telotarsus/basitarsus 2.0.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFEDFF9FEE861910FCF91672" lastPageId="15" lastPageNumber="216" pageId="13" pageNumber="214" type="discussion">
<paragraph id="EA1036D9FFEDFF9FEE861910FA9415C7" blockId="13.[151,1437,1011,2008]" lastBlockId="15.[151,1436,151,608]" lastPageId="15" lastPageNumber="216" pageId="13" pageNumber="214">
<emphasis id="D8DBEACBFFEDFF9DEE861910FEBB12DB" bold="true" box="[199,315,1659,1684]" pageId="13" pageNumber="214">Remarks.</emphasis>
Among the Iberian species with a raised lamina on movable chelal finger, with an isolated subapical tooth on the movable finger of the chelicera and only two macrosetae on the posterior margin of the carapace,
<emphasis id="D8DBEACBFFEDFF9DEBC319CBFED01293" italics="true" pageId="13" pageNumber="214">
C. (E.)
<taxonomicName id="2DAF4D5AFFEDFF9DEE8819A8FED01293" box="[201,336,1731,1756]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="villacarrillo">villacarrillo</taxonomicName>
</emphasis>
differs from
<emphasis id="D8DBEACBFFEDFF9DEFA519AFFD241293" box="[484,676,1731,1756]" italics="true" pageId="13" pageNumber="214">C. (E.) machadoi</emphasis>
Vachon, 1940 (Algarve,
<collectingCountry id="92B87649FFEDFF9DEDFF19A8FB9F1293" box="[958,1055,1731,1756]" name="Portugal" pageId="13" pageNumber="214">Portugal</collectingCountry>
) (Vachon 1940a) by the markedly distal position of trichobothrium
<emphasis id="D8DBEACBFFEDFF9DEC461983FDA212B0" box="[519,546,1768,1791]" italics="true" pageId="13" pageNumber="214">ist</emphasis>
with respect to
<emphasis id="D8DBEACBFFEDFF9DEC92198CFD78134F" box="[723,760,1767,1792]" italics="true" pageId="13" pageNumber="214">esb</emphasis>
(basal in
<emphasis id="D8DBEACBFFEDFF9DED24198CFC53134F" box="[869,979,1767,1792]" italics="true" pageId="13" pageNumber="214">machadoi</emphasis>
), and in being eyeless. From the epigean species
<emphasis id="D8DBEACBFFEDFF9DEEB81867FE38136C" box="[249,440,1804,1827]" italics="true" pageId="13" pageNumber="214">C. (E.) minutus</emphasis>
Vachon, 1940 (Algarve,
<collectingCountry id="92B87649FFEDFF9DECBB1860FCDD136B" box="[762,861,1803,1828]" name="Portugal" pageId="13" pageNumber="214">Portugal</collectingCountry>
), with similar pedipalp measurements,
<emphasis id="D8DBEACBFFEDFF9DEB071867FE9E1307" italics="true" pageId="13" pageNumber="214">
C. (E.)
<taxonomicName id="2DAF4D5AFFEDFF9DEED61844FE9E1307" box="[151,286,1839,1864]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="villacarrillo">villacarrillo</taxonomicName>
</emphasis>
can be distinguished by the absence of preocular microsetae and the large lamina on the proximal half of the movable chelal finger, with 9 rounded vestigial teeth reaching halfway between trichobothria
<emphasis id="D8DBEACBFFEDFF9DEB6E1838FAC81323" box="[1327,1352,1875,1900]" italics="true" pageId="13" pageNumber="214">sb</emphasis>
and
<emphasis id="D8DBEACBFFEDFF9DEBC61838FA151323" box="[1415,1429,1875,1900]" italics="true" pageId="13" pageNumber="214">b</emphasis>
, versus one preocular microseta on each side and a short proximal lamina with 6 rounded vestigial teeth reaching to level of
<emphasis id="D8DBEACBFFEDFF9DEEB318F0FE8B13FB" box="[242,267,1947,1972]" italics="true" pageId="13" pageNumber="214">sb</emphasis>
in
<emphasis id="D8DBEACBFFEDFF9DEF6F18F7FE5913FC" box="[302,473,1948,1971]" italics="true" pageId="13" pageNumber="214">C. (E.) minutus</emphasis>
.
<emphasis id="D8DBEACBFFEDFF9DEFA618F0FD6F13FC" box="[487,751,1947,1972]" italics="true" pageId="13" pageNumber="214">
<taxonomicName id="2DAF4D5AFFEDFF9DEFA618F0FDDA13FB" box="[487,602,1947,1972]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="214" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.) morenoi
</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011 (Córdoba,
<collectingCountry id="92B87649FFEDFF9DEF0C18D4FE0E1397" box="[333,398,1983,2008]" name="Spain" pageId="13" pageNumber="214">Spain</collectingCountry>
) bears two preocular microsetae on each side and has strong troglomorphic facies with extreme pedipalp ratios (Carabajal Márquez
<emphasis id="D8DBEACBFFEFFF9FECC61FF2FD3F14FF" box="[647,703,151,176]" italics="true" pageId="15" pageNumber="216">et al.</emphasis>
2011), making it very different to
<emphasis id="D8DBEACBFFEFFF9FEA7C1FF3FA9514FF" box="[1085,1301,151,176]" italics="true" pageId="15" pageNumber="216">
C. (E.)
<taxonomicName id="2DAF4D5AFFEFFF9FEACF1FFCFA9514FF" box="[1166,1301,151,176]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="villacarrillo">villacarrillo</taxonomicName>
</emphasis>
. In addition to the three characters mentioned above,
<emphasis id="D8DBEACBFFEFFF9FEC2D1FD6FCC9149A" box="[620,841,188,213]" italics="true" pageId="15" pageNumber="216">
C. (E.)
<taxonomicName id="2DAF4D5AFFEFFF9FEC801FD7FCC9149A" box="[705,841,188,213]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="villacarrillo">villacarrillo</taxonomicName>
</emphasis>
shares the absence of preocular microsetae with the hypogean species
<emphasis id="D8DBEACBFFEFFF9FEF321FB4FD0414B7" box="[371,644,223,248]" italics="true" pageId="15" pageNumber="216">
<taxonomicName id="2DAF4D5AFFEFFF9FEF321FB4FE6614B7" box="[371,486,223,248]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.) sendrai
</emphasis>
Zaragoza, 1985 (Valencia,
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) and
<emphasis id="D8DBEACBFFEFFF9FEA2B1FB4FA1C14B7" box="[1130,1436,223,248]" italics="true" pageId="15" pageNumber="216">
<taxonomicName id="2DAF4D5AFFEFFF9FEA2B1FB4FB5D14B7" box="[1130,1245,223,248]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.) ambrosiae
</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2012 (Cádiz,
<collectingCountry id="92B87649FFEFFF9FED9C1E6FFBA11552" box="[989,1057,260,285]" name="Spain" pageId="15" pageNumber="216">Spain</collectingCountry>
), but it is distinctly different from both of them by its markedly smaller size and robust chelal ratios: e.g. femur 0.56/0.10 (5.5) and chela 0.78/0.15 (5.18) in the new species, femur 0.86/0.12 (7.21) and chela 1.19/0.17 (7.0) in
<emphasis id="D8DBEACBFFEFFF9FEA581E26FB43152A" box="[1049,1219,332,357]" italics="true" pageId="15" pageNumber="216">C. (E.) sendrai</emphasis>
(Zaragoza 1985a), femur 1.19/0.15 (8.16) and chela 1.68/0.24 (6.9) in
<emphasis id="D8DBEACBFFEFFF9FEC991E1BFC1F15C7" box="[728,927,367,392]" italics="true" pageId="15" pageNumber="216">C. (E.) ambrosiae</emphasis>
(Carabajal Márquez
<emphasis id="D8DBEACBFFEFFF9FEACD1E1AFB4515C7" box="[1164,1221,367,392]" italics="true" pageId="15" pageNumber="216">et al.</emphasis>
2012).
</paragraph>
<caption id="BED06651FFEEFF9EEED618CBFCD61395" httpUri="https://zenodo.org/record/218276/files/figure.png" pageId="14" pageNumber="215" targetBox="[178,1385,193,1909]" targetPageId="14">
<paragraph id="EA1036D9FFEEFF9EEED618CBFCD61395" blockId="14.[151,1435,1952,2010]" pageId="14" pageNumber="215">
<emphasis id="D8DBEACBFFEEFF9EEED618CBFED613F9" bold="true" box="[151,342,1952,1975]" pageId="14" pageNumber="215">FIGURES 25–26.</emphasis>
<taxonomicName id="2DAF4D5AFFEEFF9EEF1C18CAFC9F13F8" box="[349,799,1953,1975]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="villacarrillo" status="sp. nov." subGenus="Ephippiochthonius">
<emphasis id="D8DBEACBFFEEFF9EEF1C18CAFC9F13F8" box="[349,799,1953,1975]" italics="true" pageId="14" pageNumber="215">Chthonius (Ephippiochthonius) villacarrillo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E857B0FFEEFF9EED6918CAFCFA13F9" box="[808,890,1953,1974]" pageId="14" pageNumber="215" rank="species">
<emphasis id="D8DBEACBFFEEFF9EED6918CAFCFA13F9" bold="true" box="[808,890,1953,1974]" pageId="14" pageNumber="215">sp. nov.</emphasis>
</taxonomicNameLabel>
, female holotype.
<emphasis id="D8DBEACBFFEEFF9EEA7C18CAFBD613F9" bold="true" box="[1085,1110,1953,1974]" pageId="14" pageNumber="215">25</emphasis>
, left pedipalp, minus chela and trochanter, dorsal view;
<emphasis id="D8DBEACBFFEEFF9EEFCD18AFFE251396" bold="true" box="[396,421,1988,2009]" pageId="14" pageNumber="215">26</emphasis>
, left chela, lateral view. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1036D9FFEFFF9FEE861EFFFCF91672" blockId="15.[151,1436,151,608]" pageId="15" pageNumber="216">
The
<typeStatus id="3514887BFFEFFF9FEEBF1EFEFEAE15E2" box="[254,302,405,429]" pageId="15" pageNumber="216">type</typeStatus>
specimen of
<emphasis id="D8DBEACBFFEFFF9FEF951EFEFD0A15E3" box="[468,650,404,428]" italics="true" pageId="15" pageNumber="216">C. (E.) minutus</emphasis>
, deposited in MNHN, has been examined and used for the above comparison (full data to be given elsewhere). Carabajal Márquez
<emphasis id="D8DBEACBFFEFFF9FEDDA1ED2FC57159F" box="[923,983,439,464]" italics="true" pageId="15" pageNumber="216">et al.</emphasis>
(2011) identified as
<emphasis id="D8DBEACBFFEFFF9FEA901ED3FA05159F" box="[1233,1413,440,464]" italics="true" pageId="15" pageNumber="216">C. (E.) minutus</emphasis>
a sample from Córdoba province and included the species in their key to the Andalusian
<taxonomicName id="2DAF4D5AFFEFFF9FEA351EB7FAC815BA" box="[1140,1352,476,501]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="D8DBEACBFFEFFF9FEA351EB7FAC815BA" box="[1140,1352,476,501]" italics="true" pageId="15" pageNumber="216">Ephippiochthonius</emphasis>
</taxonomicName>
. In our opinion, this record must be discarded, in view of the distance between Algarve and Córdoba and the high endemicity displayed by subterranean
<taxonomicName id="2DAF4D5AFFEFFF9FEC041D4FFC991672" box="[581,793,548,573]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="D8DBEACBFFEFFF9FEC041D4FFC991672" box="[581,793,548,573]" italics="true" pageId="15" pageNumber="216">Ephippiochthonius</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFEFFF9FEE861D2CFDAE162F" box="[199,558,583,608]" pageId="15" pageNumber="216" type="distribution">
<paragraph id="EA1036D9FFEFFF9FEE861D2CFDAE162F" blockId="15.[151,1436,151,608]" box="[199,558,583,608]" pageId="15" pageNumber="216">
<emphasis id="D8DBEACBFFEFFF9FEE861D2CFEE0162F" bold="true" box="[199,352,583,608]" pageId="15" pageNumber="216">Distribution.</emphasis>
Andalusia,
<collectingCountry id="92B87649FFEFFF9FEFA71D2CFDAA162F" box="[486,554,583,608]" name="Spain" pageId="15" pageNumber="216">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

59
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Genus
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<emphasis id="D8DBEACBFFEFFF9FEED61CA3FD2517AD" bold="true" box="[151,677,968,994]" italics="true" pageId="15" pageNumber="216">Neobisium (Neobisium) ischyrum ischyrum</emphasis>
(Navás, 1918)
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<emphasis id="D8DBEACBFFEFFF9FEED61B64FE861067" bold="true" box="[151,262,1039,1064]" pageId="15" pageNumber="216">Material.</emphasis>
<collectingCountry id="92B87649FFEFFF9FEF4F1B64FED11067" box="[270,337,1039,1064]" name="Spain" pageId="15" pageNumber="216">Spain</collectingCountry>
, Andalusia, Jaén province, Siles, Cueva del Nacimiento del Arroyo de San Blas (38º23′15.00″N, 2º32′2.95″W;
<quantity id="2D579B3CFFEFFF9FEF741B5FFE111002" box="[309,401,1076,1101]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="216" unit="m" value="1000.0">1000 m</quantity>
.a.s.l.),
<date id="9E111019FFEFFF9FEF9D1B5FFDE71003" box="[476,615,1076,1100]" pageId="15" pageNumber="216" value="2007-12-23">23.XII.2007</date>
,
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, lgt. G.E.V. (DEUA).
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<emphasis id="D8DBEACBFFEFFF9FEE861B3CFED9103F" bold="true" box="[199,345,1111,1136]" pageId="15" pageNumber="216">Description.</emphasis>
Epigean species, cuticle sclerotized, without troglomorphic adaptations. Carapace as long as broad; two pairs of well developed eyes with lens and reflective tapeta, anterior eyes separated by 0.7 their diameter from anterior margin and 0.5 from posterior eyes; epistome large, triangular and apically acute; 23 setae, formula 4:6:6:7. Tergal chaetotaxy I–VI: 6:8:11:12:12:13. Chelicera with 7 setae on palm; spinneret rounded and prominent; movable finger with teeth regular in size. Pedipalps smooth, femur with one tiny tubercle at middle and other basal of antiaxial face; patella ratio X/Y (diameter of distal opening/length of inner margin of patella) (Gabbutt &amp; Vachon 1965) 0.8; fixed finger with 70 pointed teeth reaching to base; movable finger with 52 teeth, dental row reaching halfway between trichobothria
<emphasis id="D8DBEACBFFEFFF9FEC961A3FFD701122" box="[727,752,1364,1389]" italics="true" pageId="15" pageNumber="216">sb</emphasis>
and
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, 12 pointed distal teeth, 10 rounded teeth up to level of
<emphasis id="D8DBEACBFFEFFF9FEED61A12FF1F11DF" box="[151,159,1401,1424]" italics="true" pageId="15" pageNumber="216">t</emphasis>
, followed by 11 extremely flattened teeth and 21 rudimentary teeth only recognizable by their dental canals. Trichobothrium
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closer to apex of finger than to
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(0.9);
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distad
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; ratio
<emphasis id="D8DBEACBFFEFFF9FEDAC1AF6FB8011FB" box="[1005,1024,1437,1460]" italics="true" pageId="15" pageNumber="216">st</emphasis>
–
<emphasis id="D8DBEACBFFEFFF9FEA4F1AF7FBA711FA" box="[1038,1063,1436,1461]" italics="true" pageId="15" pageNumber="216">sb</emphasis>
/
<emphasis id="D8DBEACBFFEFFF9FEA6F1AF7FBC711FA" box="[1070,1095,1436,1461]" italics="true" pageId="15" pageNumber="216">sb</emphasis>
–
<emphasis id="D8DBEACBFFEFFF9FEA141AF7FBE311FA" box="[1109,1123,1436,1461]" italics="true" pageId="15" pageNumber="216">b</emphasis>
1.5.
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<emphasis id="D8DBEACBFFEFFF9FEE861AAAFE6C1197" box="[199,492,1471,1496]" italics="true" pageId="15" pageNumber="216">Measurements and ratios.</emphasis>
Body 2.44. Carapace 0.66/0.64 (1.0). Pedipalp: femur 0.96/0.21 (4.6); patella 0.63/ 0.25 (2.6); chela+ 1.63/0.4 (4.0); chela - 1.53 (3.8); hand+ 0.72 (1.8); hand - 0.62 (1.5); movable finger 0.93; ratio movable finger/hand+ (1.3); ratio chela+/carapace 2.5; femur/carapace 1.5; femur/movable finger 1.0; femur/patella 1.5; patella/ hand+ 0.9.
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<emphasis id="D8DBEACBFFEFFF9FEE861933FEBB123E" bold="true" box="[199,315,1624,1649]" pageId="15" pageNumber="216">Remarks.</emphasis>
The specimen has a more slender pedipalp femur than indicated in Beier’s redescriptions (1939, 1952, 1963), even allowing for the wide range for the species reported by Mahnert (1985), but it coincides in all other respects, included the strong reduction of the teeth of the movable chelal finger. The species has been recorded under stones, in moss, in grass and in leaf-litter (Zaragoza 2007); this is the first record from a cave and can be considered as accidental.
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<emphasis id="D8DBEACBFFEFFF9FEE861867FEE0136A" bold="true" box="[199,352,1804,1829]" pageId="15" pageNumber="216">Distribution.</emphasis>
Iberian endemic, widespread (Zaragoza 2007), with a subspecies,
<emphasis id="D8DBEACBFFEFFF9FEA161866FA1C136A" box="[1111,1436,1804,1829]" italics="true" pageId="15" pageNumber="216">
N. (N.)
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balearicum
</emphasis>
Beier, 1939, restricted to the Balearic Islands. New for Jaén; previously recorded for several provinces of Andalusia: Cádiz,
<collectingCountry id="92B87649FFEFFF9FEF27183FFE481322" box="[358,456,1876,1901]" name="Grenada" pageId="15" pageNumber="216">Granada</collectingCountry>
, Málaga and Sevilla.
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<emphasis id="D8DBEACBFFF0FF80EED61FB5FDDE14B7" bold="true" box="[151,606,222,248]" italics="true" pageId="16" pageNumber="217">Neobisium (Ommatoblothrus) espinoi</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011
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<emphasis id="D8DBEACBFFF0FF80EED61E2CFE86152F" bold="true" box="[151,262,327,352]" pageId="16" pageNumber="217">Material.</emphasis>
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, Andalusia, Jaén province, Siles, Sima de los 30 años (38º21′20.20″N, 2º35′47.12″W;
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.a.s.l.),
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,
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, lgt. G.E.V. (DEUA).
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<emphasis id="D8DBEACBFFF0FF80EE861EE6FEC115E9" bold="true" box="[199,321,397,422]" pageId="16" pageNumber="217">Diagnosis.</emphasis>
Weakly troglomorphic. Two pairs of well developed eyes. Carapace with 32–37 setae, 7–8 on anterior margin and 11–13 on posterior; epistome triangular and prominent. Tergites I–IV with 8:12:16:18 setae. Spinneret low, prominent. Pedipalp femur ratio 6.3–6.4, femur about same length as movable finger; chela+ ratio 4.9–5.1; ratio hand+ 2.3, widest at the basal half; ratio movable finger/hand+ 1.3; teeth of movable finger hardly recognizable in the basal half; trichobothrium
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distinctly distad of middle of finger.
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<emphasis id="D8DBEACBFFF0FF80EE861D2BFED91616" bold="true" box="[199,345,576,601]" pageId="16" pageNumber="217">Description.</emphasis>
Large
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. Opisthosoma elongate, weak troglomorphic facies. Opisthosoma and legs yellowish. Carapace, chelicerae and pedipalps reddish brown.
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<paragraph id="EA1036D9FFF0FF80EE861DEDFE5216AC" blockId="16.[151,1437,327,1999]" pageId="16" pageNumber="217">Carapace longer than broad. Two pairs of well developed eyes with lens and reflective tapetum. Epistome acute, isosceles triangle shape. 37 setae, formula 7:6:13:11. Two microlyrifissures in ocular zone, four between median and posterior zones.</paragraph>
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Anterior process of coxa I with simple tooth shape, long and apically pointed; medial process slightly elevated and with numerous denticles. Pedipalpal coxa with 11 setae (including 3–4 on process), pedal coxa I with 10–
<date id="9E111019FFF0FF80EB231C7BFAFE1767" box="[1378,1406,784,808]" pageId="16" pageNumber="217" value="2012-02-12">12</date>
, II 12, III
<date id="9E111019FFF0FF80EEA51C58FED61704" box="[228,342,819,843]" pageId="16" pageNumber="217" value="1914-04-20">14, IV 20.</date>
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<paragraph id="EA1036D9FFF0FF80EE861C3EFAD417FC" blockId="16.[151,1437,327,1999]" pageId="16" pageNumber="217">Tergal chaetotaxy I–X: 8:12:16:18:20:19:18:18:16:12. Male genital area with 11 short setae on sternite II; sternite III with 17 setae behind genital opening and (3)24(3) setae on the posterior margin of the sternite. Chaetotaxy of sternites IV–X: (3)22(3):30:28:25:27:24:18. Anal cone with two dorsal and two ventral setae.</paragraph>
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Cheliceral palm with 8–9 setae; seta
<emphasis id="D8DBEACBFFF0FF80EC231CD6FDF81799" box="[610,632,957,982]" italics="true" pageId="16" pageNumber="217">gl</emphasis>
0.71 from base of movable finger. Spinneret a low, prominent, sclerotic dome. Fixed finger with 12 medium- or small-sized teeth; movable finger with 7 worn teeth on a short lamina, two large, blunt, fused medial teeth, others medium-sized or small, dental row ending far proximad of subgaleal seta. Rallum with 8 blades, all smooth or worn, except distal one unilaterally pinnate on anterior face, latter on a raised lamina and distinctly separated from other blades, basal blade very small. Serrula exterior with 32 and serrula interior with 31 blades.
</paragraph>
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Pedipalps completely smooth. Trochanter with a strong tubercle. Femur with one tiny tubercle in middle of antiaxial face, one distal glandular pore present. Patella pedicel with four micropores at base. Chela markedly curved, hand oval, maximum width in basal third; pedicel bearing five dorsal micropores; antiaxial face with three glandular pores close to finger base. Fixed finger with 97 teeth, dental row reaching finger base;
<emphasis id="D8DBEACBFFF0FF80EAB01B9EFA1C1141" box="[1265,1436,1269,1294]" italics="true" pageId="16" pageNumber="217">nodus ramosus</emphasis>
short, at level of 5th distal tooth; trichobothrium
<emphasis id="D8DBEACBFFF0FF80ECEA1A77FD3A117C" box="[683,698,1308,1331]" italics="true" pageId="16" pageNumber="217">it</emphasis>
slightly distad of
<emphasis id="D8DBEACBFFF0FF80EDC31A77FC16117C" box="[898,918,1308,1331]" italics="true" pageId="16" pageNumber="217">et</emphasis>
; trichobothrium
<emphasis id="D8DBEACBFFF0FF80EA121A77FBEE117C" box="[1107,1134,1308,1331]" italics="true" pageId="16" pageNumber="217">ist</emphasis>
markedly closer to apex of finger than to base; trichobothrium
<emphasis id="D8DBEACBFFF0FF80EC671A56FDBC1119" box="[550,572,1341,1366]" italics="true" pageId="16" pageNumber="217">ib</emphasis>
close to
<emphasis id="D8DBEACBFFF0FF80ECE01A56FD421119" box="[673,706,1341,1366]" italics="true" pageId="16" pageNumber="217">isb</emphasis>
; distance between trichobothria
<emphasis id="D8DBEACBFFF0FF80EA711A56FBC61119" box="[1072,1094,1341,1366]" italics="true" pageId="16" pageNumber="217">ib</emphasis>
and
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1.7 times that between
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and apex. Movable finger with 76 teeth, dental row ending distad of trichobothrium
<emphasis id="D8DBEACBFFF0FF80EA251A0BFBF21136" box="[1124,1138,1376,1401]" italics="true" pageId="16" pageNumber="217">b</emphasis>
, 16 pointed teeth distad of
<emphasis id="D8DBEACBFFF0FF80EED61AEFFF1F11D4" box="[151,159,1412,1435]" italics="true" pageId="16" pageNumber="217">t</emphasis>
, followed by 19 rounded teeth reaching, to level of
<emphasis id="D8DBEACBFFF0FF80ECBD1AEFFC8F11D4" box="[764,783,1412,1435]" italics="true" pageId="16" pageNumber="217">st</emphasis>
, and 41 flat teeth, of which the 27 proximal ones are so rudimentary that they can only be recognized by their dental canals; distance between trichobothria
<emphasis id="D8DBEACBFFF0FF80EB4A1ACEFAA411F1" box="[1291,1316,1445,1470]" italics="true" pageId="16" pageNumber="217">sb</emphasis>
and
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1.3 times that between
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and
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.
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Leg IV claws with a very tiny dorsal tooth proximad of middle, subterminal setae with two rami, total length
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, larger ramus
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, shorter one
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, both with tiny spines. Tibia TS 0.45, basitarsus TS 0.11, telotarsus TS 0.41.
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<emphasis id="D8DBEACBFFF0FF80EE86193FFE731224" box="[199,499,1619,1644]" italics="true" pageId="16" pageNumber="217">Measurements and ratios.</emphasis>
Body 4.93. Carapace 1.09/0.90 (1.2). Chelicera: palm 0.69/0.36 (1.9), movable finger 0.43. Pedipalp: trochanter 0.70/0.29 (2.4); femur 1.72/0.27 (6.4); patella 1.10/0.29 (3.8), pedicel 0.34, club 0.76 (2.6), ratio club/pedicel 2.2; chela+ 2.84/0.56 (5.1), chela - 2.72 (4.9); hand+ 1.28 (2.3), hand - 1.16 (2.1); movable finger 1.66; ratio movable finger/hand+ 1.3; ratio chela+/carapace 2.6; femur/carapace 1.6; femur/movable finger 1.0; femur/patella 1.6; patella/ hand+ 0.9. Leg IV: femur+patella 1.48/0.31 (4.8); tibia 1.32/0.16 (8.3); basitarsus 0.69/0.12 (6.0); telotarsus 0.71/0.10 (6.8); ratio telotarsus/basitarsus 1.0.
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<emphasis id="D8DBEACBFFF0FF80EE861840FEE0130B" bold="true" box="[199,352,1835,1860]" pageId="16" pageNumber="217">Distribution.</emphasis>
Andalusia,
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.
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<emphasis id="D8DBEACBFFF0FF80EE861825FEBB1328" bold="true" box="[199,315,1870,1895]" pageId="16" pageNumber="217">Remarks.</emphasis>
This species shares with
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<taxonomicName id="2DAF4D5AFFF0FF80ECC11825FD781328" box="[640,760,1870,1895]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">Neobisium</taxonomicName>
(O.) paucedentatum
</emphasis>
Mahnert, 1982 (
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province) (Mahnert 1982) and
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<taxonomicName id="2DAF4D5AFFF0FF80EFC9181BFD8013C6" box="[392,512,1904,1929]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">Neobisium</taxonomicName>
(O.) baenai
</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011, (Córdoba province) (Carabajal Márquez
<emphasis id="D8DBEACBFFF0FF80EC1C18FFFD0D13E3" box="[605,653,1939,1964]" italics="true" pageId="16" pageNumber="217">et al</emphasis>
. 2011) the increased chaetotaxy of the carapace, tergites and sternites, and the strongly reduced size of teeth in the basal half of the movable chelal finger.
</paragraph>
</subSubSection>
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59
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(
<figureCitation id="72942A5CFFF1FF81EEDE1FD1FF63149C" box="[159,227,186,211]" captionStart="FIGURES 2 – 5" captionStartId="5.[151,264,1280,1303]" captionTargetBox="[175,1412,308,1254]" captionTargetId="figure@5.[165,1421,301,1259]" captionTargetPageId="5" captionText="FIGURES 2 – 5. Habitus of living specimens. 2, Chthonius (E.) perezi Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011, Sistema de la Murcielaguina; 3, Chthonius (E.) espanyoli sp. nov., Complejo del Arroyo de la Rambla (PB – 4); 4, Neobisium (O.) perezi Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011, Sima del Campamento; 5, Allochernes masi (Navás, 1923), with the woodlouse Porcellio incanus Budde-Lund, 1879, Cueva de la Morciguilla." httpUri="https://zenodo.org/record/218270/files/figure.png" pageId="17" pageNumber="218">Fig. 4</figureCitation>
)
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<emphasis id="D8DBEACBFFF1FF81EED61E69FD8A1557" box="[151,522,258,280]" italics="true" pageId="17" pageNumber="218">Neobisium (Ommatoblothrus) perezi</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011: 123
</taxonomicName>
, figs 26–30, table VI.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF1FF81EED61E2BFE1C153B" pageId="17" pageNumber="218" type="description">
<paragraph id="EA1036D9FFF1FF81EED61E2BFE1C153B" blockId="17.[151,1435,258,373]" pageId="17" pageNumber="218">
<taxonomicName id="2DAF4D5AFFF1FF81EED61E2BFB651519" authority="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez, 2011: 125" authorityName="Carabajal Marquez, Garcia Carrillo &amp; Rodriguez Fernandez" authorityPageNumber="125" authorityYear="2011" box="[151,1253,320,342]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="gev" subGenus="Ommatoblothrus">
<emphasis id="D8DBEACBFFF1FF81EED61E2BFE751519" box="[151,501,320,342]" italics="true" pageId="17" pageNumber="218">Neobisium (Ommatoblothrus) gev</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011: 125
</taxonomicName>
–126, figs 31–35, table VII
<emphasis id="D8DBEACBFFF1FF81EF691E35FE16153A" bold="true" box="[296,406,350,373]" pageId="17" pageNumber="218">(syn. nov.)</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF1FF81EED61ECAFBCB176D" pageId="17" pageNumber="218" type="materials_examined">
<paragraph id="EA1036D9FFF1FF81EED61ECAFBCB176D" blockId="17.[151,1436,417,2007]" pageId="17" pageNumber="218">
<emphasis id="D8DBEACBFFF1FF81EED61ECAFE8615F5" bold="true" box="[151,262,417,442]" pageId="17" pageNumber="218">Material.</emphasis>
<collectingCountry id="92B87649FFF1FF81EF561ECAFEDA15F5" box="[279,346,417,442]" name="Spain" pageId="17" pageNumber="218">Spain</collectingCountry>
, Andalucía, Jaén province: Hornos de Segura, Sima de los Alhaurinos (38º10′55.26″N, 2º47′16.79″W;
<specimenCount id="FCA9FD50FFF1FF81EF0B1EADFE231591" box="[330,419,453,478]" pageId="17" pageNumber="218" type="generic">
<quantity id="2D579B3CFFF1FF81EF0B1EADFE1F1591" box="[330,415,453,478]" metricMagnitude="2" metricUnit="m" metricValue="9.28" pageId="17" pageNumber="218" unit="m" value="928.0">928 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="9E111019FFF1FF81EFB31EADFD071591" box="[498,647,454,478]" pageId="17" pageNumber="218" value="2010-08-12">12.VIII.2010</date>
, 1 ♀. Sima del Campamento (38º11′10.10″N, 2º46′48.59″W;
<specimenCount id="FCA9FD50FFF1FF81EB331EADFF38164D" pageId="17" pageNumber="218" type="generic">
<quantity id="2D579B3CFFF1FF81EB331EADFF34164D" metricMagnitude="2" metricUnit="m" metricValue="8.87" pageId="17" pageNumber="218" unit="m" value="887.0">887 m</quantity>
.
</specimenCount>
a.s.l.), 06.I–03.
<date id="9E111019FFF1FF81EF1F1E82FE50164E" box="[350,464,489,514]" pageId="17" pageNumber="218" value="2013-02-01">II.2013, 1</date>
♀;
<date id="9E111019FFF1FF81EC411E82FDF6164E" box="[512,630,489,513]" pageId="17" pageNumber="218" value="2012-02-11">11.II.2012</date>
,
<specimenCount id="FCA9FD50FFF1FF81ECC51E82FD34164E" box="[644,692,489,513]" pageId="17" pageNumber="218" type="generic">3 ♂</specimenCount>
;
<date id="9E111019FFF1FF81EC821E82FCD6164E" box="[707,854,489,513]" pageId="17" pageNumber="218" value="2012-08-11">11.VIII.2012</date>
,
<specimenCount id="FCA9FD50FFF1FF81ED241E82FC13164E" box="[869,915,489,513]" pageId="17" pageNumber="218" type="generic">1 ♂</specimenCount>
, 11 ♀, 2 T;
<date id="9E111019FFF1FF81EA641E82FB27164E" box="[1061,1191,489,513]" pageId="17" pageNumber="218" value="2012-09-23">23.IX.2012</date>
,
<specimenCount id="FCA9FD50FFF1FF81EAF71E82FB64164E" box="[1206,1252,489,513]" pageId="17" pageNumber="218" type="generic">1 ♂</specimenCount>
, 2 ♀, 2 T, 2 D;
<date id="9E111019FFF1FF81EED61D66FE8D1669" box="[151,269,525,550]" pageId="17" pageNumber="218" value="2012-02-08">08.II.2012</date>
–
<date id="9E111019FFF1FF81EF5C1D66FE071669" box="[285,391,525,550]" pageId="17" pageNumber="218" value="2013-01-06">06.I.2013</date>
,
<specimenCount id="FCA9FD50FFF1FF81EFDA1D65FE4E166A" box="[411,462,526,550]" pageId="17" pageNumber="218" type="generic">1 ♂</specimenCount>
, 1 ♀, 1 T. Sima de la Fractura II (38º11′11.58″N, 2º47′17.89″W;
<specimenCount id="FCA9FD50FFF1FF81EB431D66FADC1669" box="[1282,1372,525,550]" pageId="17" pageNumber="218" type="generic">
<quantity id="2D579B3CFFF1FF81EB431D66FAD81669" box="[1282,1368,525,550]" metricMagnitude="2" metricUnit="m" metricValue="9.65" pageId="17" pageNumber="218" unit="m" value="965.0">965 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="9E111019FFF1FF81EED61D5AFE8C1606" box="[151,268,561,585]" pageId="17" pageNumber="218" value="2009-05-31">31.V.2009</date>
, 1 P, 1 T; Sima del Laberinto (38º11′23.79″N, 2º46′47.23″W;
<quantity id="2D579B3CFFF1FF81EDFA1D5AFB971605" box="[955,1047,561,586]" metricMagnitude="3" metricUnit="m" metricValue="1.025" pageId="17" pageNumber="218" unit="m" value="1025.0">1025 m</quantity>
.a.s.l.),
<date id="9E111019FFF1FF81EA201D5AFB641606" box="[1121,1252,561,585]" pageId="17" pageNumber="218" value="2012-09-23">23.IX.2012</date>
,
<specimenCount id="FCA9FD50FFF1FF81EAAE1D5AFA9A1606" box="[1263,1306,561,585]" pageId="17" pageNumber="218" type="generic">2 ♂</specimenCount>
, 2 ♀, 1 T, 2 D,
<specimenCount id="FCA9FD50FFF1FF81EEFC1D3DFF6B1622" box="[189,235,598,622]" pageId="17" pageNumber="218" type="generic">1 P.</specimenCount>
Sima de los Moluscos (38º10′53.97″N, 2º47′18.19″W;
<specimenCount id="FCA9FD50FFF1FF81ED361D3DFC4D1621" box="[887,973,597,622]" pageId="17" pageNumber="218" type="generic">
<quantity id="2D579B3CFFF1FF81ED361D3DFC491621" box="[887,969,597,622]" metricMagnitude="2" metricUnit="m" metricValue="9.04" pageId="17" pageNumber="218" unit="m" value="904.0">904 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="9E111019FFF1FF81EA591D3DFB2E1621" box="[1048,1198,598,622]" pageId="17" pageNumber="218" value="2010-08-10">10.VIII.2010</date>
, 1 ♀. Sistema de la Murcielaguina (38º12′59.35″N, 2º42′37.13″W;
<quantity id="2D579B3CFFF1FF81ECEC1D12FC8A16DD" box="[685,778,633,658]" metricMagnitude="3" metricUnit="m" metricValue="1.085" pageId="17" pageNumber="218" unit="m" value="1085.0">1085 m</quantity>
.a.s.l.),
<date id="9E111019FFF1FF81ED171D11FC6B16DD" box="[854,1003,634,658]" pageId="17" pageNumber="218" value="2007-08-13">13.VIII.2007</date>
, 1 ♀;
<date id="9E111019FFF1FF81EA721D11FB3516DD" box="[1075,1205,634,658]" pageId="17" pageNumber="218" value="2010-06-27">27.VI.2010</date>
,
<specimenCount id="FCA9FD50FFF1FF81EA801D11FB6E16DE" box="[1217,1262,634,658]" pageId="17" pageNumber="218" type="generic">1 ♂</specimenCount>
, 5 ♀, 2 T, 2 D;
<date id="9E111019FFF1FF81EED61DF5FEA216F9" box="[151,290,670,694]" pageId="17" pageNumber="218" value="2012-12-24">24.XII.2012</date>
–
<date id="9E111019FFF1FF81EF731DF5FE2516F9" box="[306,421,670,694]" pageId="17" pageNumber="218" value="2013-02-03">03.II.2013</date>
,
<specimenCount id="FCA9FD50FFF1FF81EFF01DF5FE5E16FA" box="[433,478,670,694]" pageId="17" pageNumber="218" type="generic">1 ♂</specimenCount>
, 1 T;
<date id="9E111019FFF1FF81EC611DF5FD1716F9" box="[544,663,670,694]" pageId="17" pageNumber="218" value="2013-02-03">03.II.2013</date>
, 1 ♀. Sima La Secreta (38º12′12.16″N, 2º45′52.77″W;
<specimenCount id="FCA9FD50FFF1FF81EB481DF5FADC16F9" box="[1289,1372,669,694]" pageId="17" pageNumber="218" type="generic">
<quantity id="2D579B3CFFF1FF81EB481DF5FAD816F9" box="[1289,1368,669,694]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="17" pageNumber="218" unit="m" value="970.0">970 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="9E111019FFF1FF81EED61DA9FEAB1695" box="[151,299,706,730]" pageId="17" pageNumber="218" value="2008-08-11">11.VIII.2008</date>
, 2 T, 1 D. Sima de la Tubería (38º11′1.80″N, 2º47′10.28″W;
<specimenCount id="FCA9FD50FFF1FF81ED8F1DA9FB9F1695" box="[974,1055,705,730]" pageId="17" pageNumber="218" type="generic">
<quantity id="2D579B3CFFF1FF81ED8F1DA9FB9B1695" box="[974,1051,705,730]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="17" pageNumber="218" unit="m" value="930.0">930 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="9E111019FFF1FF81EA271DA9FB791695" box="[1126,1273,706,730]" pageId="17" pageNumber="218" value="2011-08-28">28.VIII.2011</date>
, 2 ♀, 1 T, 1 D. All the specimens lgt. G.E.V. and deposited in DEUA. Cazorla, Cueva Secreta del Poyo Manquillo (37°53′18.46″N, 2°52′12.80″W;
<quantity id="2D579B3CFFF1FF81EF041C62FE21176D" box="[325,417,777,802]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="218" unit="m" value="1500.0">1500 m</quantity>
.a.s.l.),
<date id="9E111019FFF1FF81EFAD1C62FDD8176D" box="[492,600,777,802]" pageId="17" pageNumber="218" value="1953-05-01">V.1953, 1</date>
♀, lgt. F. Español (MCNB: 87–9399 MZB).
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF1FF81EE861C46FEF1106B" pageId="17" pageNumber="218" type="diagnosis">
<paragraph id="EA1036D9FFF1FF81EE861C46FEF1106B" blockId="17.[151,1436,417,2007]" pageId="17" pageNumber="218">
<emphasis id="D8DBEACBFFF1FF81EE861C46FEC11709" bold="true" box="[199,321,813,838]" pageId="17" pageNumber="218">Diagnosis.</emphasis>
Troglomorphic facies. Anterior eyes usually with strongly flattened lens, posterior eyes reduced to eye-spots. Carapace with 21–25 setae, 4 on the anterior and 5–9 on posterior margin; epistome prominent, triangular. Tergites I–IV with 5–6:5–6:6–10:10–12 setae. Spinneret prominent and rounded, slightly smaller in males. Pedipalp femur ratio 6.2–7.0 in males, 6.0–
<quantity id="2D579B3CFFF1FF81EC891CF2FC8717FD" box="[712,775,921,946]" metricMagnitude="-1" metricUnit="m" metricValue="1.6764" pageId="17" pageNumber="218" unit="in" value="6.6">6.6 in</quantity>
females, femur usually slightly longer than movable chelal finger; chela+ ratio 6.0–
<quantity id="2D579B3CFFF1FF81EFDE1CABFE5E1796" box="[415,478,960,985]" metricMagnitude="-1" metricUnit="m" metricValue="1.6764" pageId="17" pageNumber="218" unit="in" value="6.6">6.6 in</quantity>
males, 5.3–6.0 in females; hand+ ratio
<quantity id="2D579B3CFFF1FF81EDD31CABFB831796" box="[914,1027,960,985]" metricMagnitude="-2" metricUnit="m" metricValue="6.985" metricValueMax="7.111999999999999" metricValueMin="6.858" pageId="17" pageNumber="218" unit="in" value="2.75" valueMax="2.8" valueMin="2.7">2.7–2.8 in</quantity>
males,
<quantity id="2D579B3CFFF1FF81EA181CABFB4A1796" box="[1113,1226,960,985]" metricMagnitude="-2" metricUnit="m" metricValue="6.223" metricValueMax="6.604" metricValueMin="5.842" pageId="17" pageNumber="218" unit="in" value="2.45" valueMax="2.6" valueMin="2.3">2.3–2.6 in</quantity>
females, widest in basal half, ratio movable finger/hand+ 1.3–1.5; teeth of movable finger all well developed; trichobothrium
<emphasis id="D8DBEACBFFF1FF81EB791C83FAD317B0" box="[1336,1363,1000,1023]" italics="true" pageId="17" pageNumber="218">ist</emphasis>
distad of middle of finger.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF1FF82EE861B45FD1D15AF" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218" type="description">
<paragraph id="EA1036D9FFF1FF81EE861B45FAE91137" blockId="17.[151,1436,417,2007]" pageId="17" pageNumber="218">
<emphasis id="D8DBEACBFFF1FF81EE861B45FD8A1008" bold="true" box="[199,522,1070,1095]" pageId="17" pageNumber="218">Measurements and ratios.</emphasis>
<emphasis id="D8DBEACBFFF1FF81EC561B44FA141008" box="[535,1428,1070,1095]" italics="true" pageId="17" pageNumber="218">
Specimens from Sima del Laberinto,
<typeStatus id="3514887BFFF1FF81ED941B5BFB841008" box="[981,1028,1072,1095]" pageId="17" pageNumber="218">type</typeStatus>
locality of N. (O.)
<taxonomicName id="2DAF4D5AFFF1FF81EAB91B5BFABF1008" box="[1272,1343,1072,1095]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
. Males
</emphasis>
. Carapace 1.08/0.93 (1.2). Pedipalp: femur 1.76–1.82/0.28–0.29 (6.2–6.3); patella 1.37–1.42/0.32–0.34 (4.2–4.3); chela+ 2.88–3.16/0.48 (6.0–6.6); chela - 2.68–2.94 (5.6–6.1); hand+ 1.28–1.35 (2.7–2.8); hand - 1.08–1.13 (2.3–2.4); movable finger 1.80–1.82; ratio movable finger/hand+ 1.3–1.4; ratio chela+/carapace 2.7; femur/carapace 1.6; femur/movable finger 1.0; femur/patella 1.3; patella/hand+ 1.0–1.1. Distance trichobothria
<emphasis id="D8DBEACBFFF1FF81EACF1BACFB4C10AF" box="[1166,1228,1223,1248]" italics="true" pageId="17" pageNumber="218">ib–ist</emphasis>
1.4–1.6 times that between
<emphasis id="D8DBEACBFFF1FF81EEBF1B87FE98114C" box="[254,280,1260,1283]" italics="true" pageId="17" pageNumber="218">ist</emphasis>
and apex (both sexes).
<emphasis id="D8DBEACBFFF1FF81EC6E1B80FD0E114B" box="[559,654,1259,1284]" italics="true" pageId="17" pageNumber="218">Females</emphasis>
. Carapace 1.0/0.9 (1.1). Pedipalp: femur 1.76–1.87/0.28–0.31 (6.0– 6.3); patella 1.35–1.44/0.32–0.36 (4.0–4.2); chela+ 2.92–3.09/0.48–0.58 (5.3–6.0); chela - 2.74–2.87 (4.9–5.7); hand+ 1.26–1.34 (2.3–2.6); hand - 1.08–1.12 (1.9–2.2); movable finger 1.74–1.88; ratio movable finger/hand+ 1.4; ratio chela+/carapace 2.9; femur/carapace 1.8; femur/movable finger 1.0; femur/patella 1.3; patella/hand+ 1.07.
</paragraph>
<paragraph id="EA1036D9FFF1FF81EE861AEFFF4112BB" blockId="17.[151,1436,417,2007]" pageId="17" pageNumber="218">
<emphasis id="D8DBEACBFFF1FF81EE861AEFFD6B11D3" box="[199,747,1411,1436]" italics="true" pageId="17" pageNumber="218">Specimens from Sima del Campamento. Males</emphasis>
. Carapace 1.00–1.08/0.88–0.92 (1.1–1.2). Pedipalp: femur 1.74–1.93/0.26–0.28 (6.7–7.0); patella 1.30–1.40/0.31–0.32 (4.2–4.3); chela+ 2.83–3.07/0.46–0.48 (6.2–6.4); chela - 2.63–2.87 (5.7–6.0); hand+ 1.22–1.33 (2.7–2.8); hand - 1.02–1.13 (2.2–2.4); movable finger 1.64–1.77; ratio movable finger/hand+ 1.3–1.4; ratio chela+/carapace 2.8; femur/carapace 1.7–1.8; femur/movable finger 1.0–1.1; femur/patella 1.3–1.4; patella/hand+ 1.0–1.1. Distance trichobothria
<emphasis id="D8DBEACBFFF1FF81EDD71975FC551278" box="[918,981,1566,1591]" italics="true" pageId="17" pageNumber="218">ib–ist</emphasis>
1.3–1.6 times that between
<emphasis id="D8DBEACBFFF1FF81EB55194BFAAF1278" box="[1300,1327,1568,1591]" italics="true" pageId="17" pageNumber="218">ist</emphasis>
and apex (both sexes).
<emphasis id="D8DBEACBFFF1FF81EF741928FE141213" box="[309,404,1603,1628]" italics="true" pageId="17" pageNumber="218">Females</emphasis>
. Carapace 1.02–1.16/0.78–0.98 (1.2–1.3). Pedipalp: femur 1.66–1.8/0.26–0.28 (6.4–6.5); patella 1.24–1.4/0.31–0.34 (4.0–4.1); chela+ 2.84–3.04/0.47–0.53 (5.8–6.0); chela - 2.66–2.86 (5.4–5.7); hand+ 1.19–1.28/0.47–0.53 (2.4–2.5); hand - 1.01–1.1 (2.1–2.2); movable finger 1.68–1.82; ratio movable finger/hand+ 1.4; ratio chela+/carapace 2.6–2.8; femur/carapace 1.6; femur/movable finger 1.0; femur/patella 1.3; patella/hand+ 1.0– 1.1.
</paragraph>
<paragraph id="EA1036D9FFF1FF82EE86186BFE3C154E" blockId="17.[151,1436,417,2007]" lastBlockId="18.[151,1437,151,949]" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218">
<emphasis id="D8DBEACBFFF1FF81EE86186BFBE71357" box="[199,1127,1791,1816]" italics="true" pageId="17" pageNumber="218">
Specimens from Sistema de la Murcielaguina,
<typeStatus id="3514887BFFF1FF81ECA2186BFC911358" box="[739,785,1792,1815]" pageId="17" pageNumber="218">type</typeStatus>
locality of N. (O.)
<taxonomicName id="2DAF4D5AFFF1FF81EDB9186BFBA01358" box="[1016,1056,1792,1815]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="gev">gev</taxonomicName>
. Male
</emphasis>
. Carapace 0.96/0.84 (1.1). Pedipalp: femur 1.77/0.26 (6.8); patella 1.29/0.31 (4.1); chela+ 2.87/0.44 (6.5); chela - 2.67 (6.1); hand+ 1.20 (2.7); hand - 1.00 (2.3); movable finger 1.7; ratio movable finger/hand+ 1.4; ratio chela+/carapace 3.0; femur/carapace 1.8; femur/movable finger 1.0; femur/patella 1.4; patella/hand+ 1.1. Distance trichobothria
<emphasis id="D8DBEACBFFF1FF81EAC31818FB4013C3" box="[1154,1216,1907,1932]" italics="true" pageId="17" pageNumber="218">ib–ist</emphasis>
1.2–1.4 times that between
<emphasis id="D8DBEACBFFF1FF81EEBD18F2FE9613FF" box="[252,278,1945,1968]" italics="true" pageId="17" pageNumber="218">ist</emphasis>
and apex (both sexes).
<emphasis id="D8DBEACBFFF1FF81EC6518FCFD0313FF" box="[548,643,1943,1968]" italics="true" pageId="17" pageNumber="218">Females</emphasis>
. Carapace 1.00–1.17/0.87–0.93 (1.1–1.3). Pedipalp: femur 1.78–1.83/ 0.27–0.29 (6.3–6.5); patella 1.30–1.33/0.32–0.34 (3.9–4.0); chela+ 2.87–3.0/0.5–0.52 (5.6–5.8); chela - 2.69–2.81 (5.3–5.4); hand+ 1.22–1.25 (2.4–2.5); hand - 1.04–1.06 (2.0–2.1); movable finger 1.68–1.78; ratio movable finger/ hand+ 1.3–1.4; ratio chela+/carapace 2.6–2.9; femur/carapace 1.5–1.8; femur/movable finger 1.0–1.1; femur/patella 1.3–1.4; patella/hand+ 1.1.
</paragraph>
<paragraph id="EA1036D9FFF2FF82EE861E66FD1D15AF" blockId="18.[151,1437,151,949]" pageId="18" pageNumber="219">
<emphasis id="D8DBEACBFFF2FF82EE861E66FAAB156A" box="[199,1323,268,293]" italics="true" pageId="18" pageNumber="219">Specimens from Sima de los Alhaurinos, Sima de los Moluscos and Sima de la Tubería. Females.</emphasis>
Carapace 0.98–1.10/0.82–0.94 (1.1–1.2). Pedipalp: femur 1.64–1.86/0.26–0.31 (6.0–6.6); patella 1.24–1.42/0.30–0.36 (3.9– 4.2); chela+ 2.82–3.06/0.48–0.55 (5.6–5.9); chela - 2.64–2.86 (5.2–5.5); hand+ 1.16–1.33 (2.4–2.6); hand - 0.98–1.13 (2.0–2.2); movable finger 1.66–1.75; ratio movable finger/hand+ 1.3–1.5; ratio chela+/carapace 2.8–3.0; femur/ carapace 1.6–1.8; femur/movable finger 1.0–1.1; femur/patella 1.3–1.4; patella/hand+ 1.1. Distance trichobothria
<emphasis id="D8DBEACBFFF2FF82EED61EA3FF5515AE" box="[151,213,456,481]" italics="true" pageId="18" pageNumber="219">ib–ist</emphasis>
1.3–1.6 times that between
<emphasis id="D8DBEACBFFF2FF82EC511EA1FDAA15AE" box="[528,554,458,481]" italics="true" pageId="18" pageNumber="219">ist</emphasis>
and apex.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF2FF82EE861E87FDAE164A" box="[199,558,492,517]" pageId="18" pageNumber="219" type="distribution">
<paragraph id="EA1036D9FFF2FF82EE861E87FDAE164A" blockId="18.[151,1437,151,949]" box="[199,558,492,517]" pageId="18" pageNumber="219">
<emphasis id="D8DBEACBFFF2FF82EE861E87FEE0164A" bold="true" box="[199,352,492,517]" pageId="18" pageNumber="219">Distribution.</emphasis>
Andalusia,
<collectingCountry id="92B87649FFF2FF82EFA71E87FDAA164A" box="[486,554,492,517]" name="Spain" pageId="18" pageNumber="219">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF2FF82EE861D7BFDC917FA" pageId="18" pageNumber="219" type="discussion">
<paragraph id="EA1036D9FFF2FF82EE861D7BFACB16DA" blockId="18.[151,1437,151,949]" pageId="18" pageNumber="219">
<emphasis id="D8DBEACBFFF2FF82EE861D7BFEBB1666" bold="true" box="[199,315,528,553]" pageId="18" pageNumber="219">Remarks.</emphasis>
Carabajal Márquez
<emphasis id="D8DBEACBFFF2FF82EC661D79FDE21666" box="[551,610,528,553]" italics="true" pageId="18" pageNumber="219">et al.</emphasis>
(2011) described the closely related species
<emphasis id="D8DBEACBFFF2FF82EA2D1D7BFAE91666" box="[1132,1385,528,553]" italics="true" pageId="18" pageNumber="219">
<taxonomicName id="2DAF4D5AFFF2FF82EA2D1D7BFB641666" box="[1132,1252,528,553]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">Neobisium</taxonomicName>
(O.)
<taxonomicName id="2DAF4D5AFFF2FF82EB671D79FAE91666" box="[1318,1385,530,553]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
and
<emphasis id="D8DBEACBFFF2FF82EED61D5FFEF91602" box="[151,377,564,589]" italics="true" pageId="18" pageNumber="219">
<taxonomicName id="2DAF4D5AFFF2FF82EED61D5FFE8F1602" box="[151,271,564,589]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">Neobisium</taxonomicName>
(O.)
<taxonomicName id="2DAF4D5AFFF2FF82EF131D5DFEF91602" box="[338,377,566,589]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="gev">gev</taxonomicName>
</emphasis>
from small samples. The differences they used to separate them were: chelal hand widest proximad of middle, ratio trichobothria
<emphasis id="D8DBEACBFFF2FF82EC2A1D33FD01163E" box="[619,641,600,625]" italics="true" pageId="18" pageNumber="219">ib</emphasis>
–
<emphasis id="D8DBEACBFFF2FF82ECCE1D31FD29163E" box="[655,681,602,625]" italics="true" pageId="18" pageNumber="219">ist</emphasis>
/
<emphasis id="D8DBEACBFFF2FF82ECF01D31FD59163E" box="[689,729,602,625]" italics="true" pageId="18" pageNumber="219">ist–</emphasis>
apex
<quantity id="2D579B3CFFF2FF82ED5B1D33FC2B163E" box="[794,939,600,625]" metricMagnitude="-2" metricUnit="m" metricValue="3.8608" metricValueMax="3.9878" metricValueMin="3.7338" pageId="18" pageNumber="219" unit="in" value="1.52" valueMax="1.57" valueMin="1.47">1.47–1.57 in</quantity>
<emphasis id="D8DBEACBFFF2FF82EDF71D31FBD8163E" box="[950,1112,601,625]" italics="true" pageId="18" pageNumber="219">
N. (O.)
<taxonomicName id="2DAF4D5AFFF2FF82EA541D31FBD8163E" box="[1045,1112,602,625]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
, as opposed to chelal hand widest at middle, ratio trichobothria
<emphasis id="D8DBEACBFFF2FF82EC711D17FDC616DA" box="[560,582,636,661]" italics="true" pageId="18" pageNumber="219">ib</emphasis>
–
<emphasis id="D8DBEACBFFF2FF82EC151D16FDEF16DB" box="[596,623,637,660]" italics="true" pageId="18" pageNumber="219">ist</emphasis>
/
<emphasis id="D8DBEACBFFF2FF82EC371D16FD1E16DB" box="[630,670,637,660]" italics="true" pageId="18" pageNumber="219">ist–</emphasis>
apex
<quantity id="2D579B3CFFF2FF82EC9B1D16FCA816DA" box="[730,808,636,661]" metricMagnitude="-2" metricUnit="m" metricValue="3.0734" pageId="18" pageNumber="219" unit="in" value="1.21">1.21 in</quantity>
<emphasis id="D8DBEACBFFF2FF82ED6E1D15FC2916DA" box="[815,937,637,661]" italics="true" pageId="18" pageNumber="219">
N. (O.)
<taxonomicName id="2DAF4D5AFFF2FF82EDC51D15FC2916DA" box="[900,937,638,661]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="gev">gev</taxonomicName>
</emphasis>
, as well as the different chelal ratios.
</paragraph>
<paragraph id="EA1036D9FFF2FF82EE861DCBFC141707" blockId="18.[151,1437,151,949]" pageId="18" pageNumber="219">
Analysis of a large sample of specimens from the
<typeStatus id="3514887BFFF2FF82ED451DCAFCB416F6" box="[772,820,673,697]" pageId="18" pageNumber="219">type</typeStatus>
localities of both species and other caves of the same municipality shows a high variability in the chelal ratios, which overlaps the published data of both species. The same applies to the widest part of the chelal hand, which show subtle variations between individuals in the same cave, and to the ratio
<emphasis id="D8DBEACBFFF2FF82EFC71C67FE1C176A" box="[390,412,780,805]" italics="true" pageId="18" pageNumber="219">ib</emphasis>
–
<emphasis id="D8DBEACBFFF2FF82EFEB1C66FE44176B" box="[426,452,781,804]" italics="true" pageId="18" pageNumber="219">ist</emphasis>
/
<emphasis id="D8DBEACBFFF2FF82EF8D1C66FE75176B" box="[460,501,781,804]" italics="true" pageId="18" pageNumber="219">ist–</emphasis>
apex. In conclusion, the two species are here considered to be subjective synonyms. As first revisers, we give
<emphasis id="D8DBEACBFFF2FF82EFF41C59FDCE1706" box="[437,590,817,841]" italics="true" pageId="18" pageNumber="219">
N. (O.)
<taxonomicName id="2DAF4D5AFFF2FF82EC4A1C59FDCE1706" box="[523,590,818,841]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
precedence over
<emphasis id="D8DBEACBFFF2FF82ED521C59FC0D1706" box="[787,909,817,841]" italics="true" pageId="18" pageNumber="219">
N. (O.)
<taxonomicName id="2DAF4D5AFFF2FF82ED291C59FC0D1706" box="[872,909,818,841]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="gev">gev</taxonomicName>
</emphasis>
.
</paragraph>
<paragraph id="EA1036D9FFF2FF82EE861C3FFDC917FA" blockId="18.[151,1437,151,949]" pageId="18" pageNumber="219">
The female specimen from Cueva Secreta del Poyo Manquillo, presents slightly lower pedipalpal ratios than the other populations. We tentatively assign this specimen to
<emphasis id="D8DBEACBFFF2FF82ED141C11FC7317DE" box="[853,1011,889,913]" italics="true" pageId="18" pageNumber="219">
N. (O.)
<taxonomicName id="2DAF4D5AFFF2FF82EDEE1C11FC7317DE" box="[943,1011,890,913]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
until new specimens are collected in that cave to allow a better comparison.
</paragraph>
</subSubSection>
</treatment>
</document>

264
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<mods:title id="65F88BBD60F278A2770C747479D8A2D4">Hypogean pseudoscorpions (Arachnida) from Jaén province (Andalusia, Spain), with descriptions of four new species and a new synonymy</mods:title>
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<treatment id="620687CFFFF2FF83EED61B63FD3213E1" ID-DOI="http://doi.org/10.5281/zenodo.6149908" ID-GBIF-Taxon="119574426" ID-Zenodo-Dep="6149908" LSID="urn:lsid:zoobank.org:act:07686D99-7B18-4B58-BF05-1B982188DC47" httpUri="http://treatment.plazi.org/id/620687CFFFF2FF83EED61B63FD3213E1" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219">
<subSubSection id="A2B56552FFF2FF82EED61B63FEA7100B" pageId="18" pageNumber="219" type="nomenclature">
<paragraph id="EA1036D9FFF2FF82EED61B63FD64106C" blockId="18.[151,740,1032,1092]" box="[151,740,1032,1059]" pageId="18" pageNumber="219">
<heading id="B15881B5FFF2FF82EED61B63FD64106C" bold="true" box="[151,740,1032,1059]" fontSize="11" level="1" pageId="18" pageNumber="219" reason="1">
<emphasis id="D8DBEACBFFF2FF82EED61B63FD64106C" bold="true" box="[151,740,1032,1059]" pageId="18" pageNumber="219">
<taxonomicName id="2DAF4D5AFFF2FF82EED61B63FDFF106D" ID-CoL="7VZBJ" box="[151,639,1032,1058]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="perezruizi" status="sp. nov." subGenus="Ommatoblothrus">
<emphasis id="D8DBEACBFFF2FF82EED61B63FDFF106D" bold="true" box="[151,639,1032,1058]" italics="true" pageId="18" pageNumber="219">Neobisium (Ommatoblothrus) perezruizi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3E857B0FFF2FF82ECC71B62FD64106C" box="[646,740,1033,1059]" pageId="18" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="EA1036D9FFF2FF82EED61B40FEA7100B" blockId="18.[151,740,1032,1092]" box="[151,295,1067,1092]" pageId="18" pageNumber="219">
(
<figureCitation id="72942A5CFFF2FF82EEDE1B40FE9E100B" box="[159,286,1067,1092]" captionStart="FIGURES 27 – 33" captionStartId="20.[151,264,1909,1932]" captionTargetBox="[168,1399,186,1885]" captionTargetId="figure@20.[151,1436,179,1918]" captionTargetPageId="20" captionText="FIGURES 27 – 33. Neobisium (Ommatoblothrus) perezruizi sp. nov., male holotype. 27, carapace; 28, partial view of anterior margin of carapace, showing epistome; 29, anterior and medial processes of right coxa I; 30, left chelicera; 31, partial view of movable finger of left chelicera; 32, right pedipalp, minus chela, dorsal view; 33, right chela, dorsal view. Scale bars (in mm): 0.1 (Figs 28, 29, 31), 0.3 (Fig. 30), 0.5 (Figs 27, 32, 33)." httpUri="https://zenodo.org/record/218278/files/figure.png" pageId="18" pageNumber="219">Figs 27–33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF2FF82EED61B18FB18109B" pageId="18" pageNumber="219" type="materials_examined">
<paragraph id="EA1036D9FFF2FF82EED61B18FB18109B" blockId="18.[151,1436,1139,2034]" pageId="18" pageNumber="219">
<emphasis id="D8DBEACBFFF2FF82EED61B18FEC810C3" bold="true" box="[151,328,1139,1164]" pageId="18" pageNumber="219">
<typeStatus id="3514887BFFF2FF82EED61B18FF5210C3" box="[151,210,1139,1164]" pageId="18" pageNumber="219">Type</typeStatus>
material.
</emphasis>
<typeStatus id="3514887BFFF2FF82EF121B18FE3B10C3" box="[339,443,1139,1164]" pageId="18" pageNumber="219" type="holotype">Holotype</typeStatus>
♂ (DEUA: 20046),
<collectingCountry id="92B87649FFF2FF82ECF11B18FD7310C3" box="[688,755,1139,1164]" name="Spain" pageId="18" pageNumber="219">Spain</collectingCountry>
, Andalusia, Jaén province, Peal de Becerro, Complejo del Arroyo de la Rambla (PB-4) (37º48′1.22″N, 2º53′36.65″W;
<quantity id="2D579B3CFFF2FF82ED071BF3FC2410FF" box="[838,932,1175,1200]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="219" unit="m" value="1000.0">1000 m</quantity>
.a.s.l.),
<date id="9E111019FFF2FF82EDB01BF3FBFC10FF" box="[1009,1148,1176,1200]" pageId="18" pageNumber="219" value="2011-12-04">04.XII.2011</date>
.
<typeStatus id="3514887BFFF2FF82EAC81BF3FB7A10FF" box="[1161,1274,1176,1200]" pageId="18" pageNumber="219" type="paratype">Paratypes</typeStatus>
: 1 T (MCNB:
<date id="9E111019FFF2FF82EED61BD0FE90109B" box="[151,272,1211,1236]" pageId="18" pageNumber="219" value="2013" valueMax="2544">2013-2544</date>
MZB), 1 D (DEUA: 20055), same details as
<typeStatus id="3514887BFFF2FF82ED4C1BD0FCF3109B" box="[781,883,1211,1236]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
. All specimens lgt. G.E.V.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF2FF82EE861BB4FF50110F" pageId="18" pageNumber="219" type="etymology">
<paragraph id="EA1036D9FFF2FF82EE861BB4FF50110F" blockId="18.[151,1436,1139,2034]" pageId="18" pageNumber="219">
<emphasis id="D8DBEACBFFF2FF82EE861BB4FECE10B7" bold="true" box="[199,334,1247,1272]" pageId="18" pageNumber="219">Etymology.</emphasis>
The species is dedicated to Antonio Pérez Ruiz, ex-president and founding member of G.E.V., to honour his great contribution to our knowledge of the speleology and hypogean fauna of Andalusia, particularly of Jaén.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF2FF82EE861A20FE4611B5" pageId="18" pageNumber="219" type="diagnosis">
<paragraph id="EA1036D9FFF2FF82EE861A20FE4611B5" blockId="18.[151,1436,1139,2034]" pageId="18" pageNumber="219">
<emphasis id="D8DBEACBFFF2FF82EE861A20FEC1112B" bold="true" box="[199,321,1355,1380]" pageId="18" pageNumber="219">Diagnosis.</emphasis>
Troglomorphic facies. No eyes or eye-spots. Carapace with about 20 setae, posterior margin probably with 4–6 setae (
<quantity id="2D579B3CFFF2FF82EFFA1A04FE6811C7" box="[443,488,1391,1416]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="18" pageNumber="219" unit="in" value="5.0">5 in</quantity>
<typeStatus id="3514887BFFF2FF82EFB01A04FDD811C7" box="[497,600,1391,1416]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
). Tergites I–IV with 6:6:6:9 setae. Male cheliceral spinneret a low, broad dome. Pedipalp femur ratio about 7.0, femur as long as movable finger; chela+ ratio 7.5; hand+ ratio 3.4, widest at middle, ratio movable finger/hand+ 1.4; movable finger teeth recognizable in whole dental row; trichobothrium
<emphasis id="D8DBEACBFFF2FF82EBC01AD5FA1C119A" box="[1409,1436,1470,1493]" italics="true" pageId="18" pageNumber="219">ist</emphasis>
distad middle of the finger.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF2FF83EE86196EFE2A122B" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219" type="description">
<paragraph id="EA1036D9FFF2FF82EE86196EFB30120D" blockId="18.[151,1436,1139,2034]" pageId="18" pageNumber="219">
<emphasis id="D8DBEACBFFF2FF82EE86196EFE4B1251" bold="true" box="[199,459,1541,1566]" pageId="18" pageNumber="219">Description of male.</emphasis>
Medium-sized
<taxonomicName id="2DAF4D5AFFF2FF82ECCF196EFCCC1251" box="[654,844,1541,1566]" class="Arachnida" family="Neobisiidae" genus="Ommatoblothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="D8DBEACBFFF2FF82ECCF196EFCCC1251" box="[654,844,1541,1566]" italics="true" pageId="18" pageNumber="219">Ommatoblothrus</emphasis>
</taxonomicName>
of moderate troglomorphic facies. Opisthosoma elongate. Opisthosoma and legs yellowish, carapace, chelicerae and pedipalps reddish brown.
</paragraph>
<paragraph id="EA1036D9FFF2FF82EE861926FF5E12E2" blockId="18.[151,1436,1139,2034]" pageId="18" pageNumber="219">
Carapace (
<figureCitation id="72942A5CFFF2FF82EF071926FE1F122A" box="[326,415,1613,1638]" captionStart="FIGURES 27 – 33" captionStartId="20.[151,264,1909,1932]" captionTargetBox="[168,1399,186,1885]" captionTargetId="figure@20.[151,1436,179,1918]" captionTargetPageId="20" captionText="FIGURES 27 – 33. Neobisium (Ommatoblothrus) perezruizi sp. nov., male holotype. 27, carapace; 28, partial view of anterior margin of carapace, showing epistome; 29, anterior and medial processes of right coxa I; 30, left chelicera; 31, partial view of movable finger of left chelicera; 32, right pedipalp, minus chela, dorsal view; 33, right chela, dorsal view. Scale bars (in mm): 0.1 (Figs 28, 29, 31), 0.3 (Fig. 30), 0.5 (Figs 27, 32, 33)." httpUri="https://zenodo.org/record/218278/files/figure.png" pageId="18" pageNumber="219">Fig. 27</figureCitation>
) longer than broad. Without eyes or eye-spots. Epistome long and acute (
<figureCitation id="72942A5CFFF2FF82EB721926FA10122A" box="[1331,1424,1613,1638]" captionStart="FIGURES 27 – 33" captionStartId="20.[151,264,1909,1932]" captionTargetBox="[168,1399,186,1885]" captionTargetId="figure@20.[151,1436,179,1918]" captionTargetPageId="20" captionText="FIGURES 27 – 33. Neobisium (Ommatoblothrus) perezruizi sp. nov., male holotype. 27, carapace; 28, partial view of anterior margin of carapace, showing epistome; 29, anterior and medial processes of right coxa I; 30, left chelicera; 31, partial view of movable finger of left chelicera; 32, right pedipalp, minus chela, dorsal view; 33, right chela, dorsal view. Scale bars (in mm): 0.1 (Figs 28, 29, 31), 0.3 (Fig. 30), 0.5 (Figs 27, 32, 33)." httpUri="https://zenodo.org/record/218278/files/figure.png" pageId="18" pageNumber="219">Fig. 28</figureCitation>
). Chaetotaxy 21 setae, formula 4:6:6:5. Four microlyrifissures on ocular zone, four between median and posterior zones.
</paragraph>
<paragraph id="EA1036D9FFF2FF82EE8619D2FE1B1356" blockId="18.[151,1436,1139,2034]" pageId="18" pageNumber="219">
Manducatory process with 4 setae. Anterolateral process of coxa I with simple tooth shape, long and apically pointed; medial process straight with 2–3 denticles (
<figureCitation id="72942A5CFFF2FF82EC9A19B6FCAF12BA" box="[731,815,1757,1782]" captionStart="FIGURES 27 – 33" captionStartId="20.[151,264,1909,1932]" captionTargetBox="[168,1399,186,1885]" captionTargetId="figure@20.[151,1436,179,1918]" captionTargetPageId="20" captionText="FIGURES 27 – 33. Neobisium (Ommatoblothrus) perezruizi sp. nov., male holotype. 27, carapace; 28, partial view of anterior margin of carapace, showing epistome; 29, anterior and medial processes of right coxa I; 30, left chelicera; 31, partial view of movable finger of left chelicera; 32, right pedipalp, minus chela, dorsal view; 33, right chela, dorsal view. Scale bars (in mm): 0.1 (Figs 28, 29, 31), 0.3 (Fig. 30), 0.5 (Figs 27, 32, 33)." httpUri="https://zenodo.org/record/218278/files/figure.png" pageId="18" pageNumber="219">Fig. 29</figureCitation>
). Pedipalpal coxa with 8 setae, pedal coxa I with 6–7,
<date id="9E111019FFF2FF82EBC819B5FF431355" pageId="18" pageNumber="219">II 5–6</date>
,
<date id="9E111019FFF2FF82EE8E1869FE9E1356" box="[207,286,1793,1817]" pageId="18" pageNumber="219">III 7–8</date>
,
<date id="9E111019FFF2FF82EF6B1869FE1B1356" box="[298,411,1793,1817]" pageId="18" pageNumber="219" value="2010-04-11">IV 10–11.</date>
</paragraph>
<paragraph id="EA1036D9FFF2FF82EE86184EFB6413CA" blockId="18.[151,1436,1139,2034]" pageId="18" pageNumber="219">
Tergal chaetotaxy I–X: 6:6:6:9:9:9:9:9:8:7. Male genital area with 11 short setae on sternite II; sternite III with 15 setae behind genital opening and (3)11(3) setae on the posterior margin of the sternite. Sternal chaetotaxy IV–X: (3)13(3):13:13:13:13:12:10. Segment
<date id="9E111019FFF2FF82EC031805FDF513CA" box="[578,629,1901,1925]" pageId="18" pageNumber="219">XI 8</date>
setae. Anal cone with two dorsal and two ventral setae.
</paragraph>
<paragraph id="EA1036D9FFF2FF83EE8618FAFF6614B7" blockId="18.[151,1436,1139,2034]" lastBlockId="19.[151,1437,151,1966]" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219">
Chelicerae (
<figureCitation id="72942A5CFFF2FF82EF1018FAFE5513E6" box="[337,469,1937,1962]" captionStart="FIGURES 27 – 33" captionStartId="20.[151,264,1909,1932]" captionTargetBox="[168,1399,186,1885]" captionTargetId="figure@20.[151,1436,179,1918]" captionTargetPageId="20" captionText="FIGURES 27 – 33. Neobisium (Ommatoblothrus) perezruizi sp. nov., male holotype. 27, carapace; 28, partial view of anterior margin of carapace, showing epistome; 29, anterior and medial processes of right coxa I; 30, left chelicera; 31, partial view of movable finger of left chelicera; 32, right pedipalp, minus chela, dorsal view; 33, right chela, dorsal view. Scale bars (in mm): 0.1 (Figs 28, 29, 31), 0.3 (Fig. 30), 0.5 (Figs 27, 32, 33)." httpUri="https://zenodo.org/record/218278/files/figure.png" pageId="18" pageNumber="219">Figs 30–31</figureCitation>
) with 7 setae on palm; seta
<emphasis id="D8DBEACBFFF2FF82ED6718FAFCBC13E5" box="[806,828,1937,1962]" italics="true" pageId="18" pageNumber="219">gl</emphasis>
0.62 from base of movable finger. Spinneret a low, prominent sclerotic dome. Fixed finger with 9 teeth; movable finger with 6 teeth, one large and blunt (probably worn) medial tooth, others medium or small, dental row ending just proximad subgaleal seta. Rallum with 8 blades, all smooth except distal one, which is unilaterally pinnate on anterior face and situated on a raised lamina, distinctly separated from other blades, basal blade very small. Serrula exterior with 32 blades, serrula interior 27 blades.
</paragraph>
<paragraph id="EA1036D9FFF3FF83EE861E6FFD47162D" blockId="19.[151,1437,151,1966]" pageId="19" pageNumber="220">
Pedipalps (
<figureCitation id="72942A5CFFF3FF83EF021E6FFE441553" box="[323,452,260,285]" captionStart="FIGURES 27 – 33" captionStartId="20.[151,264,1909,1932]" captionTargetBox="[168,1399,186,1885]" captionTargetId="figure@20.[151,1436,179,1918]" captionTargetPageId="20" captionText="FIGURES 27 – 33. Neobisium (Ommatoblothrus) perezruizi sp. nov., male holotype. 27, carapace; 28, partial view of anterior margin of carapace, showing epistome; 29, anterior and medial processes of right coxa I; 30, left chelicera; 31, partial view of movable finger of left chelicera; 32, right pedipalp, minus chela, dorsal view; 33, right chela, dorsal view. Scale bars (in mm): 0.1 (Figs 28, 29, 31), 0.3 (Fig. 30), 0.5 (Figs 27, 32, 33)." httpUri="https://zenodo.org/record/218278/files/figure.png" pageId="19" pageNumber="220">Figs 32–33</figureCitation>
) completely smooth. Lyrifissures as illustrated. Femur with one tiny tubercle at middle of antiaxial face, one distal glandular pore present. Patella pedicel with two micropores at base and one in distal portion. Chela markedly curved, hand nearly parallel sides at middle area in dorsal view, maximum width at middle; pedicel bears four dorsal micropores; two glandular pores on antiaxial face, close to finger base. Fixed finger with 119 teeth, dental row reaching base of finger;
<emphasis id="D8DBEACBFFF3FF83ED181EFDFB8815E0" box="[857,1032,406,431]" italics="true" pageId="19" pageNumber="220">nodus ramosus</emphasis>
short, at level of 4th distal tooth; trichobothrium
<emphasis id="D8DBEACBFFF3FF83EF0B1ED7FEDA159C" box="[330,346,444,467]" italics="true" pageId="19" pageNumber="220">it</emphasis>
slightly distad
<emphasis id="D8DBEACBFFF3FF83EC521ED7FDA7159C" box="[531,551,444,467]" italics="true" pageId="19" pageNumber="220">et</emphasis>
; trichobothrium
<emphasis id="D8DBEACBFFF3FF83ECAF1ED7FC89159C" box="[750,777,444,467]" italics="true" pageId="19" pageNumber="220">ist</emphasis>
closer to apex of finger than to base; distance between trichobothria
<emphasis id="D8DBEACBFFF3FF83EF6F1EB5FEC415B8" box="[302,324,478,503]" italics="true" pageId="19" pageNumber="220">ib</emphasis>
and
<emphasis id="D8DBEACBFFF3FF83EF381E8BFE1415B8" box="[377,404,480,503]" italics="true" pageId="19" pageNumber="220">ist</emphasis>
1.2 times than that between
<emphasis id="D8DBEACBFFF3FF83EC911E8BFD6A15B8" box="[720,746,480,503]" italics="true" pageId="19" pageNumber="220">ist</emphasis>
and apex; distance between trichobothria
<emphasis id="D8DBEACBFFF3FF83EAFE1EB5FB6015B8" box="[1215,1248,478,503]" italics="true" pageId="19" pageNumber="220">isb</emphasis>
and
<emphasis id="D8DBEACBFFF3FF83EB541E8BFAB015B8" box="[1301,1328,480,503]" italics="true" pageId="19" pageNumber="220">ist</emphasis>
5.6 times than that between
<emphasis id="D8DBEACBFFF3FF83EF2B1D69FE0B1654" box="[362,395,514,539]" italics="true" pageId="19" pageNumber="220">isb</emphasis>
and
<emphasis id="D8DBEACBFFF3FF83EF841D69FE5B1654" box="[453,475,514,539]" italics="true" pageId="19" pageNumber="220">ib</emphasis>
. Movable finger with 112 teeth, dental row nearly reaching base of finger; distance between trichobothria
<emphasis id="D8DBEACBFFF3FF83EFDA1D4DFE341670" box="[411,436,550,575]" italics="true" pageId="19" pageNumber="220">sb</emphasis>
and
<emphasis id="D8DBEACBFFF3FF83EFB21D43FD861670" box="[499,518,552,575]" italics="true" pageId="19" pageNumber="220">st</emphasis>
2.0 times that between
<emphasis id="D8DBEACBFFF3FF83ED641D4DFCBE1670" box="[805,830,550,575]" italics="true" pageId="19" pageNumber="220">sb</emphasis>
and
<emphasis id="D8DBEACBFFF3FF83ED3D1D4DFC0A1670" box="[892,906,550,575]" italics="true" pageId="19" pageNumber="220">b</emphasis>
. One sensillum near tip of both fingers; two separated sensilla between trichobothria
<emphasis id="D8DBEACBFFF3FF83EC1F1D21FDF7162C" box="[606,631,586,611]" italics="true" pageId="19" pageNumber="220">sb</emphasis>
and
<emphasis id="D8DBEACBFFF3FF83ECEF1D27FD41162C" box="[686,705,588,611]" italics="true" pageId="19" pageNumber="220">st</emphasis>
.
</paragraph>
<paragraph id="EA1036D9FFF3FF83EE861D05FD041680" blockId="19.[151,1437,151,1966]" pageId="19" pageNumber="220">
Claws of legs I and IV with a tiny dorsal tooth proximad of middle, subterminal setae with two rami, that of leg IV total length
<quantity id="2D579B3CFFF3FF83EF001DF8FE3516E5" box="[321,437,659,683]" metricMagnitude="-5" metricUnit="m" metricValue="8.4" pageId="19" pageNumber="220" unit="mm" value="0.084">0.084 mm</quantity>
, larger ramus
<quantity id="2D579B3CFFF3FF83EC151DF8FD4816E5" box="[596,712,659,683]" metricMagnitude="-5" metricUnit="m" metricValue="4.299999999999999" pageId="19" pageNumber="220" unit="mm" value="0.043">0.043 mm</quantity>
, shorter ramus
<quantity id="2D579B3CFFF3FF83ED351DF8FC6816E5" box="[884,1000,659,683]" metricMagnitude="-5" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="220" unit="mm" value="0.024">0.024 mm</quantity>
, both with tiny spines. Leg IV tibia TS 0.43, basitarsus TS 0.07, telotarsus TS 0.38.
</paragraph>
<paragraph id="EA1036D9FFF3FF83EE861DB7FE2A17B8" blockId="19.[151,1437,151,1966]" pageId="19" pageNumber="220">
<emphasis id="D8DBEACBFFF3FF83EE861DB7FE7316BC" box="[199,499,730,755]" italics="true" pageId="19" pageNumber="220">Measurements and ratios.</emphasis>
Body 2.65. Carapace 0.87/0.76 (1.2). Chelicera: palm 0.65/0.33 (2.0), movable finger 0.41. Pedipalp: trochanter 0.70/0.25 (2.8); femur 1.66/0.24 (6.9); patella 1.32/0.27 (4.9), pedicel 0.38, club 0.94 (3.5), ratio club/pedicel 2.5; chela+ 2.74/0.37 (7.5), chela - 2.58 (7.1); hand+ 1.23 (3.4), hand - 1.07 (2.9); movable finger 1.66; ratio movable finger/hand+ 1.4; ratio chela+/carapace 3.1; femur/carapace 1.9; femur/movable finger 1.0; femur/patella 1.3; patella/hand+ 1.1. Leg I: femur 0.96/0.14 (6.7); patella 0.60/0.12 (4.9); tibia 0.65/0.10 (6.5); basitarsus 0.45/0.08 (5.6); telotarsus 0.65/0.10 (6.7); ratio femur/patella 1.6; telotarsus/basitarsus 1.4. Leg IV: femur+patella 1.46/0.29 (5.1); tibia 1.28/0.17 (7.5); basitarsus 0.66/0.13 (4.9); telotarsus 0.84/0.12 (7.1); ratio telotarsus/basitarsus 1.3.
</paragraph>
<paragraph id="EA1036D9FFF3FF83EE861B68FD041174" blockId="19.[151,1437,151,1966]" pageId="19" pageNumber="220">
<emphasis id="D8DBEACBFFF3FF83EE861B68FED71053" bold="true" box="[199,343,1027,1052]" pageId="19" pageNumber="220">Tritonymph</emphasis>
(data coinciding with
<typeStatus id="3514887BFFF3FF83EC121B68FD351053" box="[595,693,1027,1052]" pageId="19" pageNumber="220" type="holotype">holotype</typeStatus>
omitted). Carapace with 22 setae, formula 4:6:6:6; epistome short and acute. Pedipalpal coxa with 7 setae, pedal coxa
<date id="9E111019FFF3FF83EC9A1B43FD7B1070" box="[731,763,1063,1087]" pageId="19" pageNumber="220">I 4</date>
,
<date id="9E111019FFF3FF83ED461B43FCB0100F" box="[775,816,1063,1088]" pageId="19" pageNumber="220">II 5</date>
,
<date id="9E111019FFF3FF83ED7D1B43FCEF100F" box="[828,879,1063,1088]" pageId="19" pageNumber="220">III 5</date>
,
<date id="9E111019FFF3FF83ED3A1B43FC34100F" box="[891,948,1063,1088]" pageId="19" pageNumber="220">IV 6.</date>
Tergal chaetotaxy I–X: 6:6:7:6:6:8:7:8:7:7. Sternite II with 4 microsetae; sternal chaetotaxy III–X: (2)7(2):(2)9(2):11:12:11:10:10:9. Cheliceral palm with 6 setae; spinneret slightly more prominent than in male; fixed finger with 14 teeth; movable finger with 8 teeth, three teeth large and close together, the medial one even longer than the others. Fixed chelal finger with 91 teeth; movable finger with 88 teeth, decreasing in size in basal half but still well developed and much more easily recognized than in male; trichobothrium
<emphasis id="D8DBEACBFFF3FF83EC251BB7FDF310BC" box="[612,627,1244,1267]" italics="true" pageId="19" pageNumber="220">it</emphasis>
distinctly distad of
<emphasis id="D8DBEACBFFF3FF83ED141BB7FCE910BC" box="[853,873,1244,1267]" italics="true" pageId="19" pageNumber="220">et</emphasis>
; trichobothrium
<emphasis id="D8DBEACBFFF3FF83EA681BB7FBC310BC" box="[1065,1091,1244,1267]" italics="true" pageId="19" pageNumber="220">ist</emphasis>
only slightly closer to apex of finger than to base; distance between trichobothria
<emphasis id="D8DBEACBFFF3FF83EC9A1B94FD711157" box="[731,753,1279,1304]" italics="true" pageId="19" pageNumber="220">ib</emphasis>
and
<emphasis id="D8DBEACBFFF3FF83ED681A6BFCC41158" box="[809,836,1280,1303]" italics="true" pageId="19" pageNumber="220">ist</emphasis>
0.9 times that between
<emphasis id="D8DBEACBFFF3FF83EA131A6BFBEC1158" box="[1106,1132,1280,1303]" italics="true" pageId="19" pageNumber="220">ist</emphasis>
and apex. Leg IV tibia TS 0.36, basitarsus TS 0.02, telotarsus TS 0.38.
</paragraph>
<paragraph id="EA1036D9FFF3FF83EE861A23FE2A122B" blockId="19.[151,1437,151,1966]" pageId="19" pageNumber="220">
<emphasis id="D8DBEACBFFF3FF83EE861A23FE6B1110" box="[199,491,1351,1376]" italics="true" pageId="19" pageNumber="220">Measurements and ratios</emphasis>
: Body 2.80. Carapace 0.71/0.60 (1.2). Chelicera: palm 0.51/0.27 (1.9), movable finger 0.32. Pedipalp: trochanter 0.48/0.21 (2.3); femur 1.16/0.20 (5.8); patella 0.84/0.23 (3.7), pedicel 0.25, club 0.59 (2.6), ratio club/pedicel 2.4; chela+ 1.98/0.32 (6.3), chela - 1.89 (6.0); hand+ 0.82 (2.6), hand - 0.73 (2.3); movable finger 1.19; ratio movable finger/hand+ 1.5; ratio chela+/carapace 2.8; femur/carapace 1.6; femur/movable finger 1.0; femur/patella 1.4; patella/ hand+ 1.0. Leg I: femur 0.63/0.12 (5.3); patella 0.39/0.10 (3.8); tibia 0.44/0.9 (4.9); basitarsus 0.30/0.07 (4.3); telotarsus 0.46/0.07 (6.7); ratio femur/patella 1.6; telotarsus/basitarsus 1.5. Leg IV: femur+patella 1.04/0.21 (5.0); tibia 0.85/0.11 (7.5); basitarsus 0.41/0.09 (4.6); telotarsus 0.60/0.08 (7.5); ratio telotarsus/basitarsus 1.5.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF3FF83EE861904FDAE12C7" box="[199,558,1647,1672]" pageId="19" pageNumber="220" type="distribution">
<paragraph id="EA1036D9FFF3FF83EE861904FDAE12C7" blockId="19.[151,1437,151,1966]" box="[199,558,1647,1672]" pageId="19" pageNumber="220">
<emphasis id="D8DBEACBFFF3FF83EE861904FEE012C7" bold="true" box="[199,352,1647,1672]" pageId="19" pageNumber="220">Distribution.</emphasis>
Andalusia,
<collectingCountry id="92B87649FFF3FF83EFA71904FDAA12C7" box="[486,554,1647,1672]" name="Spain" pageId="19" pageNumber="220">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF3FF83EE8619F8FD3213E1" pageId="19" pageNumber="220" type="discussion">
<paragraph id="EA1036D9FFF3FF83EE8619F8FD3213E1" blockId="19.[151,1437,151,1966]" pageId="19" pageNumber="220">
<emphasis id="D8DBEACBFFF3FF83EE8619F8FEBB12E3" bold="true" box="[199,315,1683,1708]" pageId="19" pageNumber="220">Remarks.</emphasis>
<emphasis id="D8DBEACBFFF3FF83EF0319F8FDE612E3" box="[322,614,1683,1708]" italics="true" pageId="19" pageNumber="220">
<taxonomicName id="2DAF4D5AFFF3FF83EF0319F8FE3A12E3" box="[322,442,1683,1708]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="220" phylum="Arthropoda" rank="genus">Neobisium</taxonomicName>
(O.)
<taxonomicName id="2DAF4D5AFFF3FF83EFB619FEFDE612E3" box="[503,614,1685,1708]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="perezruizi">perezruizi</taxonomicName>
</emphasis>
belongs to the Spanish group of species with well developed teeth in the basal half of the dental row on the movable chelal finger, usually 6 or less setae on posterior margin of carapace and a large tooth on the movable cheliceral finger, but it differs of all them in the chela+ ratio (
<quantity id="2D579B3CFFF3FF83EA8A19B4FA9C12B8" box="[1227,1308,1758,1783]" metricMagnitude="-1" metricUnit="m" metricValue="1.2623799999999998" pageId="19" pageNumber="220" unit="in" value="4.97">4.97 in</quantity>
<emphasis id="D8DBEACBFFF3FF83EB6519B5FEA81354" italics="true" pageId="19" pageNumber="220">
<taxonomicName id="2DAF4D5AFFF3FF83EB6519B5FA1C12B8" box="[1316,1436,1758,1783]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="220" phylum="Arthropoda" rank="genus">Neobisium</taxonomicName>
(O.) rodrigoi
</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2000, 6.0 in
<emphasis id="D8DBEACBFFF3FF83EA221868FA941354" box="[1123,1300,1794,1819]" italics="true" pageId="19" pageNumber="220">N. (O.) hiberum</emphasis>
Beier, 1931 and
<quantity id="2D579B3CFFF3FF83EE86184CFE951370" box="[199,277,1830,1855]" metricMagnitude="-1" metricUnit="m" metricValue="2.49682" pageId="19" pageNumber="220" unit="in" value="9.83">9.83 in</quantity>
<emphasis id="D8DBEACBFFF3FF83EF5D184CFE3B1371" box="[284,443,1831,1854]" italics="true" pageId="19" pageNumber="220">N. (O.) tinauti</emphasis>
Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández, 2011). The geographically closest species,
<emphasis id="D8DBEACBFFF3FF83EF081820FE61132D" box="[329,481,1867,1890]" italics="true" pageId="19" pageNumber="220">
N. (O.)
<taxonomicName id="2DAF4D5AFFF3FF83EFDF1820FE61132D" box="[414,481,1867,1890]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="perezi">perezi</taxonomicName>
</emphasis>
, shares a regular dental row on cheliceral movable finger, but is more robust than the new species (ratios male chela hand+ 2.7–2.8 and chela+ 6.0–6.6, versus hand+ 3.4 and chela+
<quantity id="2D579B3CFFF3FF83EAB7181AFABB13C5" box="[1270,1339,1905,1930]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" pageId="19" pageNumber="220" unit="in" value="7.5">7.5 in</quantity>
<emphasis id="D8DBEACBFFF3FF83EB091819FE8613E2" italics="true" pageId="19" pageNumber="220">
N. (O.)
<taxonomicName id="2DAF4D5AFFF3FF83EED618FDFE8613E2" box="[151,262,1942,1965]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="perezruizi">perezruizi</taxonomicName>
</emphasis>
and has a different chelal hand shape.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="596D95F47017F92846067224F494D1A4" ID-DOI="10.11646/zootaxa.3700.2.1" ID-GBIF-Dataset="3d28eeb6-88a2-4fb2-aebd-796c8b2c2287" ID-ISSN="1175-5326" ID-Zenodo-Dep="218268" ID-ZooBank="8119DAED-D7AF-4858-BC63-F082151A4DAE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460662093739" checkinUser="plazi" docAuthor="Zaragoza, Juan A. &amp; Pérez, Toni" docDate="2013" docId="620687CFFFF5FF85EED61E0CFAD113D3" docLanguage="en" docName="zt03700p225.pdf" docOrigin="Zootaxa 3700 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microcreagrella caeca subsp. caeca Simon 1883" docType="treatment" docVersion="8" lastPageNumber="222" masterDocId="9E3FFFB7FFE0FF90EE411F6BFF80144F" masterDocTitle="Hypogean pseudoscorpions (Arachnida) from Jaén province (Andalusia, Spain), with descriptions of four new species and a new synonymy" masterLastPageNumber="225" masterPageNumber="201" pageNumber="222" updateTime="1698347634734" updateUser="plazi">
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<mods:title id="4B78A3B47317BF2147A8F2CBD5287875">Hypogean pseudoscorpions (Arachnida) from Jaén province (Andalusia, Spain), with descriptions of four new species and a new synonymy</mods:title>
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<mods:namePart id="EEEB03DF635C136CB9F191DA5F4D407D">Zaragoza, Juan A.</mods:namePart>
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<mods:namePart id="AC848DDE183645C3BCB8FE10A7ABCAB8">Pérez, Toni</mods:namePart>
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<mods:date id="B61176F3E63E08B65EBE657CFE4E1B74">2013</mods:date>
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<mods:number id="A8F5DBF3178BAE4FEA6E72D742AF9C82">3700</mods:number>
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<treatment id="620687CFFFF5FF85EED61E0CFAD113D3" ID-DOI="http://doi.org/10.5281/zenodo.6149916" ID-GBIF-Taxon="119574422" ID-Zenodo-Dep="6149916" LSID="urn:lsid:plazi:treatment:620687CFFFF5FF85EED61E0CFAD113D3" httpUri="http://treatment.plazi.org/id/620687CFFFF5FF85EED61E0CFAD113D3" lastPageNumber="222" pageId="21" pageNumber="222">
<subSubSection id="A2B56552FFF5FF85EED61E0CFD2515CD" box="[151,677,359,386]" pageId="21" pageNumber="222" type="nomenclature">
<paragraph id="EA1036D9FFF5FF85EED61E0CFD2515CD" blockId="21.[151,677,359,386]" box="[151,677,359,386]" pageId="21" pageNumber="222">
<heading id="B15881B5FFF5FF85EED61E0CFD2515CD" bold="true" box="[151,677,359,386]" fontSize="11" level="1" pageId="21" pageNumber="222" reason="1">
<taxonomicName id="2DAF4D5AFFF5FF85EED61E0CFD2515CD" ID-CoL="5JP2W" authority="Simon, 1883" authorityName="Simon" authorityYear="1883" box="[151,677,359,386]" class="Arachnida" family="Syarinidae" genus="Microcreagrella" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="222" phylum="Arthropoda" rank="subSpecies" species="caeca" subSpecies="caeca">
<emphasis id="D8DBEACBFFF5FF85EED61E0CFD2515CD" bold="true" box="[151,677,359,386]" pageId="21" pageNumber="222">
<emphasis id="D8DBEACBFFF5FF85EED61E0CFE7115CE" bold="true" box="[151,497,359,385]" italics="true" pageId="21" pageNumber="222">Microcreagrella caeca caeca</emphasis>
(Simon, 1883)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF5FF85EED61EC5FD5415A4" pageId="21" pageNumber="222" type="materials_examined">
<paragraph id="EA1036D9FFF5FF85EED61EC5FD5415A4" blockId="21.[151,1436,430,1948]" pageId="21" pageNumber="222">
<emphasis id="D8DBEACBFFF5FF85EED61EC5FE861588" bold="true" box="[151,262,430,455]" pageId="21" pageNumber="222">Material.</emphasis>
<collectingCountry id="92B87649FFF5FF85EF4C1EC5FED01588" box="[269,336,430,455]" name="Spain" pageId="21" pageNumber="222">Spain</collectingCountry>
, Andalusia, Jaén province,
<taxonomicName id="2DAF4D5AFFF5FF85ECC21EC5FC8D1588" box="[643,781,430,455]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="215" phylum="Arthropoda" rank="species" species="villacarrillo" status="sp. nov.">Villacarrillo</taxonomicName>
, Cueva de la Morciguilla (38°5′27.87″N, 3°0′5.62″W;
<specimenCount id="FCA9FD50FFF5FF85EB331EC4FF3815A4" pageId="21" pageNumber="222" type="generic">
<quantity id="2D579B3CFFF5FF85EB331EC4FF3415A4" metricMagnitude="2" metricUnit="m" metricValue="7.01" pageId="21" pageNumber="222" unit="m" value="701.0">701 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="9E111019FFF5FF85EEBE1EB8FEF615A4" box="[255,374,467,491]" pageId="21" pageNumber="222" value="2013-02-24">24.II.2013</date>
, lgt. G.E.V.,
<specimenCount id="FCA9FD50FFF5FF85EC421EB8FDAF15A5" box="[515,559,467,491]" pageId="21" pageNumber="222" type="generic">1 ♂</specimenCount>
, 1 ♀ (DEUA).
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF5FF85EE861E9DFAA81113" pageId="21" pageNumber="222" type="description">
<paragraph id="EA1036D9FFF5FF85EE861E9DFDCD167C" blockId="21.[151,1436,430,1948]" pageId="21" pageNumber="222">
<emphasis id="D8DBEACBFFF5FF85EE861E9DFED91640" bold="true" box="[199,345,502,527]" pageId="21" pageNumber="222">Description.</emphasis>
Carapace, chelicerae and pedipalps brownish-red, legs and opisthosoma pale. Opisthosoma elongate, partially destroyed in female.
</paragraph>
<paragraph id="EA1036D9FFF5FF85EE861D54FCB51634" blockId="21.[151,1436,430,1948]" pageId="21" pageNumber="222">Carapace elongate, distinctly longer than broad, anterior margin slightly prominent, no recognizable epistome; no eyes or eye-spots; chaetotaxy: 24 setae, formula 4-6-8-6.</paragraph>
<paragraph id="EA1036D9FFF5FF85EE861DEDFD4E1760" blockId="21.[151,1436,430,1948]" pageId="21" pageNumber="222">Tergal chaetotaxy I–XI: 9:11:11:11:12:11:11:11:11:9:6 (at least 2 TS). Manducatory process with 2 setae on each side. Male genital area with 9 setae on sternite II; sternite III with 2 medial setae behind genital opening and (3)6(3) setae on posterior margin of the sternite; genitalia with 3+3 internal setae. Sternal chaetotaxy IV–XI: (3)9(3):14:14:15:15:13:11:6(2 TS); anal cone with two dorsal and two ventral setae; pleural membrane dorsally and ventrally striate, weakly granulate in-between.</paragraph>
<paragraph id="EA1036D9FFF5FF85EE861C51FCB517D4" blockId="21.[151,1436,430,1948]" pageId="21" pageNumber="222">
Chelicera with five setae on palm, seta
<emphasis id="D8DBEACBFFF5FF85ECC51C51FD1A171C" box="[644,666,826,851]" italics="true" pageId="21" pageNumber="222">gl</emphasis>
0.62 from base of movable finger. Galea simple, length
<quantity id="2D579B3CFFF5FF85EB5B1C50FAFF171D" box="[1306,1407,827,851]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="222" unit="mm" value="0.04">0.04 mm</quantity>
in male,
<quantity id="2D579B3CFFF5FF85EE9F1C35FEAF1738" box="[222,303,862,887]" metricMagnitude="-3" metricUnit="m" metricValue="1.524" pageId="21" pageNumber="222" unit="in" value="0.06">0.06 in</quantity>
female. Fixed finger with 5 tiny distal denticles, 10 medium or small teeth and 3 basal denticles; movable finger with 8 large teeth and 2 tiny basal denticles.
</paragraph>
<paragraph id="EA1036D9FFF5FF85EE861CCDFDEF1000" blockId="21.[151,1436,430,1948]" pageId="21" pageNumber="222">
Pedipalp femur shorter than the carapace. Movable chelal finger longer than hand with pedicel. Trichobothriotaxy as illustrated by Vachon (1940b: fig. 3). Trichobothrium
<emphasis id="D8DBEACBFFF5FF85EDB91CA7FB8017AC" box="[1016,1024,972,995]" italics="true" pageId="21" pageNumber="222">t</emphasis>
bent backwards, acuminate, shorter and thicker than others on movable finger, not flattened or lanceolate;
<emphasis id="D8DBEACBFFF5FF85EDE51C84FC371049" box="[932,951,1007,1030]" italics="true" pageId="21" pageNumber="222">st</emphasis>
close to
<emphasis id="D8DBEACBFFF5FF85EA581C84FBA11049" box="[1049,1057,1007,1030]" italics="true" pageId="21" pageNumber="222">t</emphasis>
, distance
<emphasis id="D8DBEACBFFF5FF85EAD11C85FB1E1048" box="[1168,1182,1006,1031]" italics="true" pageId="21" pageNumber="222">b</emphasis>
–
<emphasis id="D8DBEACBFFF5FF85EAED1C85FB451048" box="[1196,1221,1006,1031]" italics="true" pageId="21" pageNumber="222">sb</emphasis>
1.24× distance
<emphasis id="D8DBEACBFFF5FF85EB341C85FA0E1048" box="[1397,1422,1006,1031]" italics="true" pageId="21" pageNumber="222">sb</emphasis>
–
<emphasis id="D8DBEACBFFF5FF85EED61B7FFF2A1064" box="[151,170,1044,1067]" italics="true" pageId="21" pageNumber="222">st</emphasis>
,
<emphasis id="D8DBEACBFFF5FF85EEF91B79FF511064" box="[184,209,1042,1067]" italics="true" pageId="21" pageNumber="222">sb</emphasis>
not forming group with
<emphasis id="D8DBEACBFFF5FF85EFA61B7FFE7A1064" box="[487,506,1044,1067]" italics="true" pageId="21" pageNumber="222">st</emphasis>
-
<emphasis id="D8DBEACBFFF5FF85EC421B7FFD8B1064" box="[515,523,1044,1067]" italics="true" pageId="21" pageNumber="222">t</emphasis>
. A row of 4 antiaxial feather-shaped setae on distal half of hand and another row of 4 on antiaxial basal half of fixed finger.
</paragraph>
<paragraph id="EA1036D9FFF5FF85EE861B37FAA81113" blockId="21.[151,1436,430,1948]" pageId="21" pageNumber="222">
<emphasis id="D8DBEACBFFF5FF85EE861B37FDC7103C" box="[199,583,1114,1139]" italics="true" pageId="21" pageNumber="222">Measurements and ratios. Male.</emphasis>
Body 1.94. Carapace 0.59/0.43 (1.4). Chelicera: palm 0.30/0.16 (1.9), movable finger 0.20. Pedipalp: femur 0.51/0.17 (3.1); patella 0.45/0.22 (2.0), pedicel 0.15; chela+ 0.86/0.29 (3.0), chela - 0.79 (2.7); hand+ 0.41 (1.4), hand - 0.34 (1.2); movable finger 0.45, ratio movable finger/hand (1.1); ratio chela+/carapace 1.5; femur/carapace 0.9; femur/movable finger 1.1; femur/patella 1.1; patella/ hand+ 1.1.
<emphasis id="D8DBEACBFFF5FF85EB001BA5FA1C10A8" box="[1345,1436,1230,1255]" italics="true" pageId="21" pageNumber="222">Female.</emphasis>
Carapace 0.61/0.46 (1.34). Pedipalp: femur 0.57/0.18 (3.1); 0.49/0.24 (2.0), pedicel 0.17; chela+ 0.93/0.32 (2.9), chela - 0.85/0.32 (2.6); hand+ 0.40 (1.3), hand - 0.34 1.0; movable finger 0.53, ratio movable finger/hand 1.3; ratio chela+/carapace 1.5; femur/carapace 0.9; femur/movable finger 1.1; femur/patella 1.2; patella/ hand+ 1.2.
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF5FF85EE861A0CFBDA11CF" box="[199,1114,1383,1408]" pageId="21" pageNumber="222" type="distribution">
<paragraph id="EA1036D9FFF5FF85EE861A0CFBDA11CF" blockId="21.[151,1436,430,1948]" box="[199,1114,1383,1408]" pageId="21" pageNumber="222">
<emphasis id="D8DBEACBFFF5FF85EE861A0CFEE011CF" bold="true" box="[199,352,1383,1408]" pageId="21" pageNumber="222">Distribution.</emphasis>
<collectingCountry id="92B87649FFF5FF85EF261A0CFE4611CF" box="[359,454,1383,1408]" name="Portugal" pageId="21" pageNumber="222">Portugal</collectingCountry>
(Azores; Madeira; mainland),
<collectingCountry id="92B87649FFF5FF85ED611A0CFCE011CF" box="[800,864,1383,1408]" name="Spain" pageId="21" pageNumber="222">Spain</collectingCountry>
(Canarias; mainland).
</paragraph>
</subSubSection>
<subSubSection id="A2B56552FFF5FF85EE861AE0FAD113D3" pageId="21" pageNumber="222" type="discussion">
<paragraph id="EA1036D9FFF5FF85EE861AE0FC79127B" blockId="21.[151,1436,430,1948]" pageId="21" pageNumber="222">
<emphasis id="D8DBEACBFFF5FF85EE861AE0FEBB11EB" bold="true" box="[199,315,1419,1444]" pageId="21" pageNumber="222">Remarks.</emphasis>
The assignment of the above studied specimens to the subspecies
<taxonomicName id="2DAF4D5AFFF5FF85EA7B1AE7FAFF11EC" box="[1082,1407,1420,1443]" class="Arachnida" family="Syarinidae" genus="Microcreagrina" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="222" phylum="Arthropoda" rank="subSpecies" species="caeca" subSpecies="caeca">
<emphasis id="D8DBEACBFFF5FF85EA7B1AE7FAFF11EC" box="[1082,1407,1420,1443]" italics="true" pageId="21" pageNumber="222">Microcreagrina caeca caeca</emphasis>
</taxonomicName>
is based in the length of the movable chelal finger relative to the hand with pedicel, which is equal in
<taxonomicName id="2DAF4D5AFFF5FF85EAAC1ADBFA1C1188" box="[1261,1436,1456,1479]" class="Arachnida" family="Syarinidae" genus="Microcreagrina" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="222" phylum="Arthropoda" rank="subSpecies" species="caeca" subSpecies="caeca">
<emphasis id="D8DBEACBFFF5FF85EAAC1ADBFA1C1188" box="[1261,1436,1456,1479]" italics="true" pageId="21" pageNumber="222">M. caeca caeca</emphasis>
</taxonomicName>
versus shorter in the Madeiran subspecies
<taxonomicName id="2DAF4D5AFFF5FF85EC2F1ABFFC7711A3" authority="Beier 1963" authorityName="Beier" authorityYear="1963" box="[622,1015,1491,1516]" class="Arachnida" family="Syarinidae" genus="Microcreagrina" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="222" phylum="Arthropoda" rank="subSpecies" species="caeca" subSpecies="madeirensis">
<emphasis id="D8DBEACBFFF5FF85EC2F1ABFFCE011A3" box="[622,864,1491,1516]" italics="true" pageId="21" pageNumber="222">M. caeca madeirensis</emphasis>
(Beier 1963)
</taxonomicName>
. However, the lengths of the pedipalp femur and chela, and the chelal movable finger/hand are higher than in the nominate subspecies, as was also the case for the specimen described by Zaragoza (2010) from mainland
<collectingCountry id="92B87649FFF5FF85EDD31970FC74127B" box="[914,1012,1563,1588]" name="Portugal" pageId="21" pageNumber="222">Portugal</collectingCountry>
.
</paragraph>
<paragraph id="EA1036D9FFF5FF85EE861954FBE61343" blockId="21.[151,1436,430,1948]" pageId="21" pageNumber="222">
The presence of feather-shaped setae on the pedipalp chela was argued by Zaragoza (2010) to be a characteristic for the subfamily
<taxonomicName id="2DAF4D5AFFF5FF85EFBE1908FD081233" box="[511,648,1635,1660]" class="Arachnida" family="Syarinidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="222" phylum="Arthropoda" rank="subFamily" subFamily="Ideobisiinae">Ideobisiinae</taxonomicName>
within the family
<taxonomicName id="2DAF4D5AFFF5FF85ED1A1908FC561233" box="[859,982,1635,1660]" class="Arachnida" family="Syarinidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="222" phylum="Arthropoda" rank="family">Syarinidae</taxonomicName>
. A group of 4–5 such setae on the basal half of the chelal fixed finger has been mentioned by Mahnert (1976, 1993) and Zaragoza (1990, 2010) for the syarinid genera
<taxonomicName id="2DAF4D5AFFF5FF85EF0E19C7FD0E128B" authority="Beier, 1961" authorityName="Beier" authorityYear="1961" box="[335,654,1707,1732]" class="Arachnida" family="Syarinidae" genus="Microcreagrina" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="D8DBEACBFFF5FF85EF0E19C7FD80128C" box="[335,512,1708,1731]" italics="true" pageId="21" pageNumber="222">Microcreagrina</emphasis>
Beier, 1961
</taxonomicName>
and
<taxonomicName id="2DAF4D5AFFF5FF85EC8B19C0FC00128B" box="[714,896,1707,1732]" class="Arachnida" family="Syarinidae" genus="Microcreagrella" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="D8DBEACBFFF5FF85EC8B19C0FC00128B" box="[714,896,1707,1732]" italics="true" pageId="21" pageNumber="222">Microcreagrella</emphasis>
</taxonomicName>
, both inhabiting Macaronesian Islands and the Iberian Peninsula. An additional group 4 setae feather-shaped is also present on antiaxial distal half of chelal hand in
<taxonomicName id="2DAF4D5AFFF5FF85EEF51998FEEA1343" box="[180,362,1779,1804]" class="Arachnida" family="Syarinidae" genus="Microcreagrella" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="D8DBEACBFFF5FF85EEF51998FEEA1343" box="[180,362,1779,1804]" italics="true" pageId="21" pageNumber="222">Microcreagrella</emphasis>
</taxonomicName>
. The taxonomic importance of this characteristic remains unknown.
</paragraph>
<paragraph id="EA1036D9FFF5FF85EE86187CFAD113D3" blockId="21.[151,1436,430,1948]" pageId="21" pageNumber="222">
The genus
<taxonomicName id="2DAF4D5AFFF5FF85EF04187CFE7B137F" box="[325,507,1815,1840]" class="Arachnida" family="Syarinidae" genus="Microcreagrella" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="D8DBEACBFFF5FF85EF04187CFE7B137F" box="[325,507,1815,1840]" italics="true" pageId="21" pageNumber="222">Microcreagrella</emphasis>
</taxonomicName>
was first described from the Azores and subsequently found on Madeira and the Canary Islands. Mahnert (1980) and Zaragoza (2010) recorded its presence in mainland
<collectingCountry id="92B87649FFF5FF85EAFC1850FA9C131B" box="[1213,1308,1851,1876]" name="Portugal" pageId="21" pageNumber="222">Portugal</collectingCountry>
(Coimbra, Setúbal). The new location in Southern
<collectingCountry id="92B87649FFF5FF85EC241834FD281337" box="[613,680,1887,1912]" name="Spain" pageId="21" pageNumber="222">Spain</collectingCountry>
, reported here, and unpublished data of J.A. Zaragoza extend the distribution range of the genus to a great part of mainland
<collectingCountry id="92B87649FFF5FF85ED6218E8FC0213D3" box="[803,898,1923,1948]" name="Portugal" pageId="21" pageNumber="222">Portugal</collectingCountry>
and to other southern Spanish provinces.
</paragraph>
</subSubSection>
</treatment>
</document>

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data/62/06/87/620687CFFFF5FF85EED61E49FDC21573.xml Normal file
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(
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)
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<emphasis id="D8DBEACBFFF6FF86EED61EB9FE8615A4" bold="true" box="[151,262,466,491]" pageId="22" pageNumber="223">Material.</emphasis>
<collectingCountry id="92B87649FFF6FF86EF4C1EB9FED015A4" box="[269,336,466,491]" name="Spain" pageId="22" pageNumber="223">Spain</collectingCountry>
, Andalusia, Jaén province,
<taxonomicName id="2DAF4D5AFFF6FF86ECC21EB9FC8D15A4" box="[643,781,466,491]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="22" pageNumber="215" phylum="Arthropoda" rank="species" species="villacarrillo" status="sp. nov.">Villacarrillo</taxonomicName>
, Cueva de la Morciguilla (38°5′27.87″N, 3°0′5.62″W;
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<quantity id="2D579B3CFFF6FF86EB331EB8FF341640" metricMagnitude="2" metricUnit="m" metricValue="7.01" pageId="22" pageNumber="223" unit="m" value="701.0">701 m</quantity>
.
</specimenCount>
a.s.l.),
<date id="9E111019FFF6FF86EEBE1E9DFEFF1640" box="[255,383,502,527]" pageId="22" pageNumber="223" value="2008-03-16">16.III.2008</date>
, ♂, 4 ♀, 3 T;
<date id="9E111019FFF6FF86EC5B1E9CFD1B1640" box="[538,667,503,527]" pageId="22" pageNumber="223" value="2008-03-23">23.III.2008</date>
,
<specimenCount id="FCA9FD50FFF6FF86ECE71E9CFD521641" box="[678,722,503,527]" pageId="22" pageNumber="223" type="generic">1 ♂</specimenCount>
, 4 ♀;
<date id="9E111019FFF6FF86ED561E9CFC181640" box="[791,920,503,527]" pageId="22" pageNumber="223" value="2013-03-09">09.III.2013</date>
,
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, 11 ♀, 1 T. All specimens lgt. G.E.V. and deposited in DEUA.
</paragraph>
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<emphasis id="D8DBEACBFFF6FF86EE861D55FEE01618" bold="true" box="[199,352,574,599]" pageId="22" pageNumber="223">Distribution.</emphasis>
<collectingCountry id="92B87649FFF6FF86EF291D55FE291618" box="[360,425,574,599]" name="Israel" pageId="22" pageNumber="223">Israel</collectingCountry>
,
<collectingCountry id="92B87649FFF6FF86EFF41D55FE691618" box="[437,489,574,599]" name="Italy" pageId="22" pageNumber="223">Italy</collectingCountry>
(Sardinia, Sicily),
<collectingCountry id="92B87649FFF6FF86EC801D55FCA31618" box="[705,803,574,599]" name="Portugal" pageId="22" pageNumber="223">Portugal</collectingCountry>
,
<collectingCountry id="92B87649FFF6FF86ED6E1D55FCEF1618" box="[815,879,574,599]" name="Spain" pageId="22" pageNumber="223">Spain</collectingCountry>
(Harvey 2011; Gardini 2000). Iberian Peninsula: Algarve (
<collectingCountry id="92B87649FFF6FF86EF431D09FEE31634" box="[258,355,610,635]" name="Portugal" pageId="22" pageNumber="223">Portugal</collectingCountry>
), Alicante, Barcelona, Castellón, Huesca, Sevilla, Valencia (
<collectingCountry id="92B87649FFF6FF86EA661D09FBEA1634" box="[1063,1130,610,635]" name="Spain" pageId="22" pageNumber="223">Spain</collectingCountry>
) (Zaragoza 2007). Second record for Andalusia, after Sevilla. In the Iberian Peninsula, this species seems to be restricted to southern and eastern areas.
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<mods:classification id="6A991D91EADC8EFB8183634ACE8F1648">journal article</mods:classification>
<mods:identifier id="3D6E1AD73B246CE4C77D3AE7F532D80E" type="DOI">http://dx.doi.org/10.3897/dez.62.5285</mods:identifier>
<mods:identifier id="C0189BDD9FE49862BAE0973794CAF660" type="Pensoft-Pub">1860-1324-2-211</mods:identifier>
<mods:identifier id="D90EDB37DB3383AAF1766ACC4EBBA268" type="ZooBank">075E22007B72481B8D1587708FD1A8BE</mods:identifier>
</mods:mods>
<treatment id="62072679C9DE25408D3460819DD249ED" ID-GBIF-Taxon="127858928" LSID="urn:lsid:zoobank.org:act:1CC0B157-15A9-4F86-85AA-29FB14857F92" httpUri="http://treatment.plazi.org/id/62072679C9DE25408D3460819DD249ED" lastPageId="4" lastPageNumber="215" pageId="2" pageNumber="213">
<subSubSection id="62E9151C3A04E29AFE73656CF8C53596" pageId="2" pageNumber="213" type="multiple">
<paragraph id="D51D1243F532E3DFCC0C2DC07C3FA87F" pageId="2" pageNumber="213">Taxon classification Animalia Coleoptera Coccinellidae</paragraph>
</subSubSection>
<subSubSection id="DF8A0DBA86F954295EF51BF9114B3F37" pageId="2" pageNumber="213" type="nomenclature">
<paragraph id="429B0D7086C2DC58AB3A3DDC6D1E4E77" pageId="2" pageNumber="213">
<taxonomicName id="2CF7698BEA9DE4B421FDC32EC45C2783" LSID="http://zoobank.org/1CC0B157-15A9-4F86-85AA-29FB14857F92" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) yanzigouensis" order="Coleoptera" pageId="2" pageNumber="213" phylum="Arthropoda" rank="species" species="yanzigouensis" subGenus="Parapullus">Scymnus (Parapullus) yanzigouensis</taxonomicName>
<taxonomicNameLabel id="F435B51FDF271204291D28A891D55C8E" pageId="2" pageNumber="213">sp. n.</taxonomicNameLabel>
Figs 8, 12
</paragraph>
</subSubSection>
<subSubSection id="99DF1A5224FEB0A94C77D49F04025755" pageId="3" pageNumber="214" type="diagnosis">
<paragraph id="BADF6FE82B320691CE878BCD0DB1FE99" pageId="3" pageNumber="214">
<pageBreakToken id="FDD804B4FB6F0EF37F7EE6E3531029D7" pageId="3" pageNumber="214" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph id="994DDCD6C1B9ACCD2ADF9AC050FFCD8B" pageId="3" pageNumber="214">
This species is similar to
<taxonomicName id="B65D557A0988AA67540627437F1655F3" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) nanlingicus" order="Coleoptera" pageId="3" pageNumber="214" phylum="Arthropoda" rank="species" species="nanlingicus" subGenus="Parapullus">Scymnus (Parapullus) nanlingicus</taxonomicName>
Chen, Ren &amp; Wang, 2012 in general appearance and male genitalia, but can be separated from it by having broad area enclosed by abdominal postcoxal lines (Fig. 8d) and the parameres narrowest at base, expanding toward apex in lateral view (Fig. 8h).
</paragraph>
<caption id="BEE7CF42EEBA62769B56772EBC8502ED" pageId="3" pageNumber="214">
<paragraph id="840C5461E4CF70ADB96879642C5A7DB0" pageId="3" pageNumber="214">
Figure 8.
<taxonomicName id="785F05385670AF3FC3905FE1E027A0DF" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) yanzigouensis" order="Coleoptera" pageId="3" pageNumber="214" phylum="Arthropoda" rank="species" species="yanzigouensis" subGenus="Parapullus">Scymnus (Parapullus) yanzigouensis</taxonomicName>
sp. n.: a dorsal view; b anterior view; c lateral view; d abdomen; e penis; f apex of penis; g tegmen, ventral view; h tegmen, lateral view. Scale bars:
<normalizedToken id="7865870101FD68FAEAB34164E3EB57FC" originalValue="a–c">a-c</normalizedToken>
0.5 mm, d 0.2 mm,
<normalizedToken id="1A769D621A85F193DE8F4325F2A353E9" originalValue="e–h">e-h</normalizedToken>
0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4193A6A02C796665CAB7521DFD57D3D8" pageId="3" pageNumber="214" type="description">
<paragraph id="88140966F222227BD19D46BF9B32A82A" pageId="3" pageNumber="214">Description.</paragraph>
<paragraph id="B667BBAACE743D36CA5BF94C92FBACC4" pageId="3" pageNumber="214">TL: 2.20-2.42 mm, TW: 1.52-1.67 mm, TH: 0.99-1.05 mm, TL/TW: 1.45, PL/PW: 0.51, EL/EW: 1.13-1.14, HW/PW: 0.56-0.58, PW/EW: 0.73-0.74.</paragraph>
<paragraph id="85C64114BC2E5ECEE5DBA68773D5C1EA" pageId="3" pageNumber="214">
Body oval, moderately convex, dorsum covered with white pubescence (Fig. 8
<normalizedToken id="33F89100F014B2FB99E6F93328282B98" originalValue="a–c">a-c</normalizedToken>
). Head, antennae and mouthparts brown. Pronotum yellowish brown. Scutellum dark brown. Elytra black with apical 1/4 yellowish brown. Prothoracic hypomeron and prosternum yellowish brown. Mesoventrite, metaventrite and elytral epipleura black. Legs brown.
</paragraph>
<paragraph id="FF8D1764D891EB11406893A4446C9A0F" pageId="3" pageNumber="214">Head with fine frontal punctures, as large as eye facets, 0.5-1.0 diameters apart. Eyes densely faceted, interocular distance 0.47 times head width. Pronotal punctures similar to those on frons, 1.0-2.0 diameters apart. Surface of elytra with punctures larger than those on pronotum, separated by 2.0-3.0 diameters. Prosternal process rectangular, 2 times as long as its width at base; with lateral carinae distinct, parallel, extending to anterior margin of prosternum. Abdominal postcoxal lines strongly recurved and distinctly incomplete laterally (Fig. 8d), reaching 7/8 length of abdominal ventrite 1, area enclosed by lines irregularly punctate, broadly smooth along line. Abdominal ventrite 5 in male with apex truncate.</paragraph>
<paragraph id="2B49DAE5C59B515E790E195DD1A0788B" pageId="3" pageNumber="214">
Male genitalia: penis stout (Fig. 8e). Penis capsule with outer arm stout and inner arm strongly curved, hook-shaped. Apex of penis simple (Fig. 8f). Tegmen stout (Fig. 8
<normalizedToken id="ED9FF8D27D92937CC655235BEF5128D8" originalValue="g–h">g-h</normalizedToken>
) with penis guide parallel sided from base to 4/5 length, then tapering abruptly to blunt apex in ventral view (Fig. 8g). Parameres narrowest at base, expanding toward apex, distinctly shorter than penis guide, densely covered with two groups of long setae at apices (Fig. 8h).
</paragraph>
<paragraph id="A7B2AA61415B90662C64DB0832C958A1" pageId="3" pageNumber="214">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="AF87B39B836D66A9F3BA32B234AC15C0" pageId="4" pageNumber="215" type="type material">
<paragraph id="4B4A114D319CE8C2C9AB1B3E9C086E0C" pageId="4" pageNumber="215">
<pageBreakToken id="B0E663ABFCA6766FF50F218C70CEF82D" pageId="4" pageNumber="215" start="start">Type</pageBreakToken>
materials.
</paragraph>
<paragraph id="1234CC6536224761DB58259DC2FB655C" pageId="4" pageNumber="215">
Holotype: male, No. SCAU (E) 11531, China: Sichuan: Yanzigou, Gonggashan National Nature Reserve, Luding County,
<geoCoordinate id="EC18CFD9E02E0CC7CA5251326A72DF5C" direction="north" orientation="latitude" precision="9" value="29.698668">29°41.92'N</geoCoordinate>
,
<geoCoordinate id="FB7E9F55B466C3A825994751E8E211E2" direction="east" orientation="longitude" precision="9" value="102.00367">102°0.22'E</geoCoordinate>
, ca 2500 m, 28. IX. 2007, Chen XS leg. Paratypes (1): 1♂ with same data as holotype.
</paragraph>
</subSubSection>
<subSubSection id="A26017E57A7E44BF912405C10051DCE9" pageId="4" pageNumber="215" type="distribution">
<paragraph id="47D206ED45F08C41251AE01E5A5000E6" pageId="4" pageNumber="215">Distribution.</paragraph>
<paragraph id="0B6AE57D8CC59D2B8548F73D7C95E52C" pageId="4" pageNumber="215">China (Sichuan).</paragraph>
</subSubSection>
<subSubSection id="93C2721A435EFCCEABFA02A82F89E218" pageId="4" pageNumber="215" type="etymology">
<paragraph id="2B4276BE2A57C058535F1C4D1335308F" pageId="4" pageNumber="215">Etymology.</paragraph>
<paragraph id="B2DFF97810737BFFB79B2DC1138FFDEF" pageId="4" pageNumber="215">The specific epithet refers to its type locality, Yanzigou Scenic Spots, Gonggashan National Nature Reserve, Sichuan Province.</paragraph>
<caption id="0BCA1892959C91EF48477588DE7F943C" pageId="4" pageNumber="215">
<paragraph id="78F27B38001559FFEA442D600C732F98" pageId="4" pageNumber="215">
Figure 9. Distribution map.
<taxonomicName id="B97B7FABF4C8DB24AB059A90ECA759F6" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) hastatus" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="hastatus" subGenus="Parapullus">Scymnus (Parapullus) hastatus</taxonomicName>
sp. n. (★);
<taxonomicName id="11F10904122DF5449D2635042AE21B68" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) tsugae" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="tsugae" subGenus="Parapullus">Scymnus (Parapullus) tsugae</taxonomicName>
Yu &amp; Yao (▲);
<taxonomicName id="6F242BE8C7E4B19F65705F2E42A772C0" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) alishanensis" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="alishanensis" subGenus="Parapullus">Scymnus (Parapullus) alishanensis</taxonomicName>
Pang &amp; Yu (●);
<taxonomicName id="45704276308598278277D1E00F7B43A6" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) baxianshanensis" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="baxianshanensis" subGenus="Parapullus">Scymnus (Parapullus) baxianshanensis</taxonomicName>
sp. n. (■).
</paragraph>
</caption>
<caption id="47482614442BF81401EF517AA3EB41D6" pageId="4" pageNumber="215">
<paragraph id="445E6539488C49A52F842D7C474ED40E" pageId="4" pageNumber="215">
Figure 10. Distribution map.
<taxonomicName id="D944CB48B82FB4ED93B48ADC5B06010D" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) laojielingensis" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="laojielingensis" subGenus="Parapullus">Scymnus (Parapullus) laojielingensis</taxonomicName>
sp. n. (●);
<taxonomicName id="E49A2AF3699A2373AC889E6521C4F305" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) parallelicus" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="parallelicus" subGenus="Parapullus">Scymnus (Parapullus) parallelicus</taxonomicName>
Chen, Ren &amp; Wang (▲);
<taxonomicName id="7AB66DFBFEF2FF8CF62A42CB035EACA3" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) annuliformis" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="annuliformis" subGenus="Parapullus">Scymnus (Parapullus) annuliformis</taxonomicName>
sp. n. (★);
<taxonomicName id="44E3F16CF483C3A647BE3A0E76EEAA93" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) malleatus" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="malleatus" subGenus="Parapullus">Scymnus (Parapullus) malleatus</taxonomicName>
Chen, Ren &amp; Wang (■).
</paragraph>
</caption>
<caption id="95A2C1B5A133C395B6AB65DCABB66FAF" pageId="4" pageNumber="215">
<paragraph id="9237049A1499F96C479DF526DBFFB91A" pageId="4" pageNumber="215">
Figure 11. Distribution map.
<taxonomicName id="D6644525B70022A14E6DA5C0D0BD2279" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) papillatus" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="papillatus" subGenus="Parapullus">Scymnus (Parapullus) papillatus</taxonomicName>
sp. n. (★);
<taxonomicName id="2E5A46E3A9DDCDE11357A8AF2143BDB6" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) solus" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="solus" subGenus="Parapullus">Scymnus (Parapullus) solus</taxonomicName>
Chen, Ren &amp; Wang (▲);
<taxonomicName id="42EDDFEE7F39A5FAC761E77094EE8F1C" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) aduncatus" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="aduncatus" subGenus="Parapullus">Scymnus (Parapullus) aduncatus</taxonomicName>
Chen, Ren &amp; Wang (●);
<taxonomicName id="5064A4CD441D0567AA5063C43F7DBBE6" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) dichotomus" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="dichotomus" subGenus="Parapullus">Scymnus (Parapullus) dichotomus</taxonomicName>
sp. n. (■).
</paragraph>
</caption>
<caption id="3760198C6D53422B794E6338FDF4B5A1" pageId="4" pageNumber="215">
<paragraph id="9EA19A565E4107121B139967C93AA9FF" pageId="4" pageNumber="215">
Figure 12. Distribution map.
<taxonomicName id="F14EE023B422C913542CAC5D95B21588" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) shenmuensis" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="shenmuensis" subGenus="Parapullus">Scymnus (Parapullus) shenmuensis</taxonomicName>
sp. n. (●);
<taxonomicName id="0A7A44296EC9BAE21A9AD7D9B9F7A7BD" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) secula" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="secula" subGenus="Parapullus">Scymnus (Parapullus) secula</taxonomicName>
Yang (▲);
<taxonomicName id="D32BE813DF3C37C8686EE77463691F66" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) nanlingicus" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="nanlingicus" subGenus="Parapullus">Scymnus (Parapullus) nanlingicus</taxonomicName>
Chen, Ren &amp; Wang (★);
<taxonomicName id="00B89473DA55AFDC7E211B414B7BA1DF" class="Insecta" family="Coccinellidae" genus="Scymnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scymnus (Parapullus) yanzigouensis" order="Coleoptera" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="yanzigouensis" subGenus="Parapullus">Scymnus (Parapullus) yanzigouensis</taxonomicName>
sp. n. (■).
</paragraph>
</caption>
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28.
<taxonomicName id="90932F0EA82B6B28B29E77A8E3D395A8" authority="A.Berlese, 1916" authorityName="A.Berlese" authorityYear="1916" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="marinus">Lasioseius marinus</taxonomicName>
<taxonomicNameLabel id="4A942AF4F55A8518A1C388D3C223118C" pageId="7" pageNumber="146" rank="species">nov. spec.</taxonomicNameLabel>
(Abb. 4a, b).1)
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<paragraph id="167583636F5DC6092B7FAFC7F6CC1DED" pageId="7" pageNumber="146">
Das hinten geradlinig abgeschnittene Genitale und das
<normalizedToken id="F50EBA13716C16D97AFAAFF304D8ED2C" originalValue="große" pageId="7" pageNumber="146">grosse</normalizedToken>
Ventrianale weisen die Species in die Subfam.
<taxonomicName id="E564A0F0DE43E2CD4340EBDF781CAFF5" pageId="7" pageNumber="146" rank="subFamily" subFamily="Podocininae">Podocininae</taxonomicName>
, und sie
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da wegen der deutlichen
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und der fast glatten Struktur in die Gattung
<taxonomicName id="CB5B05C0A361292D7F6FB1BE6341D116" authority="A.Berlese, 1916" authorityName="A.Berlese" authorityYear="1916" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Lasioseius</taxonomicName>
.
</paragraph>
<paragraph id="BD691D3A1E5DBDFA1AC183EE41BD7F30" pageId="7" pageNumber="146">
1) Es ist
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,
<normalizedToken id="FF3DD6D36F9113596001176BF0842F52" originalValue="daß" pageId="7" pageNumber="146">dass</normalizedToken>
<taxonomicName id="2183FA7B1F7EFDE614C1B6755AE7B349" authority="A.Berlese, 1916" authorityName="A.Berlese" authorityYear="1916" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="marinus">Lasioseius marinus</taxonomicName>
in das Genus
<taxonomicName id="977B99C551CDE539F8AA8A1B3581D23B" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Typhlodromus</taxonomicName>
eingereiht werden
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, ebenso der als Vergleichsart herangezogene
<taxonomicName id="38C3E5D6F84F7CF083287397A00CDB4E" authority="A.Berlese, 1916" authorityName="A.Berlese" authorityYear="1916" class="Arachnida" family="Ascidae" genus="Lasioseius" genus.outerRound="1" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="polonicus">L. polonicus</taxonomicName>
. Dann
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die Art zu den
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<normalizedToken id="6A237B8B1A0076F8246BD927CC08190E" originalValue="gehören" pageId="7" pageNumber="146">gehoeren</normalizedToken>
. Um die systematische Stellung dieser Species sicher entscheiden zu
<normalizedToken id="ABD0C2AB575A69B8978DA50FEFA5B29E" originalValue="können" pageId="7" pageNumber="146">koennen</normalizedToken>
,
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noch weitere Untersuchungen an
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Material beiderlei Geschlechts
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werden.
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Weibchen (Abb. 4 a, b):
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480-495, Breite 225
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</paragraph>
<paragraph id="91EBA727C31FB4D400C7A2B307C12F24" pageId="8" pageNumber="147">
<pageBreakToken id="AEC8DAD9CBA3235BB33A918AE520A562" pageId="8" pageNumber="147" start="start">Als</pageBreakToken>
besonders charakteristisches Merkmal nenne ich zwei kurze, aber stark verbreiterte Borsten auf dem hinteren Teile des
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, die bei anderen Arten der Gattung fehlen. Die Borsten am Hinterrande sind nicht besonders
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. Das Sternale ist kurz und breit, Vorderrand undeutlich, man hat den Eindruck, als ob das erste Borstenpaar vor dem Schilde stehe. Die Hinterecken des Sternale sind etwas ausgezogen. Die Metasternalschilder sind
<normalizedToken id="82598AB644F4D67CEE570758424A82B8" originalValue="auffällig" pageId="8" pageNumber="147">auffaellig</normalizedToken>
spitzeckig.
</paragraph>
</subSubSection>
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<paragraph id="99E5C657CAFF120F67A75C7E6EBF1E30" pageId="8" pageNumber="147">
Differenzialdiagnose: In der
<normalizedToken id="7BA2038DAD7B688183867CF07B81B832" originalValue="Größe" pageId="8" pageNumber="147">Groesse</normalizedToken>
steht die Art zwischen
<taxonomicName id="E38EC4680BCB5A7E4B18D2BB957046DC" authority="A.Berlese, 1916" authorityName="A.Berlese" authorityYear="1916" class="Arachnida" family="Ascidae" genus="Lasioseius" genus.outerRound="1" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="polonicus">L. polonicus</taxonomicName>
Willmann 1949 (360/180
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) und
<taxonomicName id="B354937A37BCC6489169DA8029642AB0" authority="A.Berlese, 1916" authorityName="A.Berlese" authorityYear="1916" class="Arachnida" family="Ascidae" genus="Lasioseius" genus.bestMatchDistance="359" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="berlesei">L. berlesei Oudemans</taxonomicName>
(540/340
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), in Gestalt und auch in der Form des Ventrianale
<normalizedToken id="4004C28E0C05A8F721E7E547DDFC5649" originalValue="ähnelt" pageId="8" pageNumber="147">aehnelt</normalizedToken>
die Species
<taxonomicName id="47A71BAE2CA8F1B24DB877813FD46463" authority="A.Berlese, 1916" authorityName="A.Berlese" authorityYear="1916" class="Arachnida" family="Ascidae" genus="Lasioseius" genus.outerRound="1" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="polonicus">L. polonicus</taxonomicName>
. Sie unterscheidet sich aber von dieser Art durch das Paar verbreiterter, fast
<normalizedToken id="F2A86AD467353701DE2625917BC73312" originalValue="blattförmiger" pageId="8" pageNumber="147">blattfoermiger</normalizedToken>
Haare auf dem
<normalizedToken id="2C15517314BE023680B467C9520A38AF" originalValue="Rücken" pageId="8" pageNumber="147">Ruecken</normalizedToken>
und durch das hinten in zwei
<normalizedToken id="56FB42748ECCC2AB6EAC77EE864C3E62" originalValue="Fortsätze" pageId="8" pageNumber="147">Fortsaetze</normalizedToken>
ausgezogene Sternalschild.
</paragraph>
</subSubSection>
<subSubSection id="BBBB8852763A6706FD14DFB7915656F9" pageId="8" pageNumber="147" type="materials_examined">
<paragraph id="2CDA384AA5CF11E750A5F4FC32E679A4" pageId="8" pageNumber="147">
<materialsCitation id="A63694048F178917A4BD2DB430F62C0B" ID-GBIF-Occurrence="923906361" collectingDate="1949-10-09" collectingMethod="Boden- und Bewuchsprobe, Berlese-Extraktion" collectorName="Carl Willmann" country="Germany" elevation="0 m" latitude="53.7820" location="Vogelschutzgebiet-Ost" longLatPrecision="600" longitude="7.9380" stateProvince="Wangerooge" typeStatus="Holotypus">
Fundort:
<location id="37CE4B198D2FDEF59FAFA3E47B7628E1" LSID="urn:lsid:plazi:treatment:62076A7153684B06E014F856920B43C2:37CE4B198D2FDEF59FAFA3E47B7628E1" country="Germany" elevation="0 m" latitude="53.7820" longLatPrecision="600" longitude="7.9380" name="Vogelschutzgebiet-Ost" stateProvince="Wangerooge">Vogelschutzgebiet-Ost</location>
,
<collectingMethod id="FEE8042CDCF7B12E22A58848455CAFD0">Bodenprobe</collectingMethod>
mit
<taxonomicName id="CC61A3BC49A78AF13ECAF4817CD78191" class="Magnoliopsida" family="Amaranthaceae" genus="Salicornia" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="147" phylum="Tracheophyta" rank="genus">Salicornia</taxonomicName>
und anderen Wattpflanzen,
<date id="B96FF7191AE61A17A98D23274BAB4200" value="1949-10-09">
<collectingDate id="3BD01067AD3D877EFA95D98ABB438C46" value="1949-10-09">9. X. 49</collectingDate>
</date>
.
<paragraph id="EB33A658FF19974C032B016FCB4BC0FA" pageId="8" pageNumber="147">
<typeStatus id="CC1DE0B20736605AD92447D9B9BDB40C">Holotypus</typeStatus>
:
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mit einem Weibchen in meiner Sammlung.
</paragraph>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title>Checklist of British and Irish Hymenoptera - Braconidae</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Broad, Gavin R.</mods:namePart>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaw, Mark R.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Godfray, H. Charles J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
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<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>8151</mods:start>
<mods:end>8151</mods:end>
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<mods:location>
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<treatment LSID="urn:lsid:plazi:treatment:620787AF74D8A7A22430CBA61FE4837D" httpUri="http://treatment.plazi.org/id/620787AF74D8A7A22430CBA61FE4837D" lastPageNumber="8151" pageId="0" pageNumber="8151">
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<paragraph pageId="0" pageNumber="8151">
<taxonomicName authority="Nees, 1834" authorityName="Nees" authorityYear="1834" genus="Ichneumonoidea" lsidName="Ichneumonoidea (Euphorus)" pageId="0" pageNumber="8151" rank="subGenus" subGenus="Euphorus">Euphorus Nees, 1834</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="8151" type="notes">
<paragraph pageId="0" pageNumber="8151">Notes</paragraph>
<paragraph pageId="0" pageNumber="8151">
Treated as a subgenus of
<taxonomicName class="Insecta" family="Braconidae" genus="Leiophron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leiophron" order="Hymenoptera" pageId="0" pageNumber="8151" phylum="Arthropoda" rank="genus">Leiophron</taxonomicName>
by
<bibRefCitation author="Belokobylskij, S. A." editor="Ler, P. A." journalOrPublisher="Dalnauka, Vladivostok" pageId="0" pageNumber="8151" title="[Key to the insects of Russian Far East. Vol. IV. Neuropteroidea, Mecoptera, Hymenoptera. Pt 4.] Opredelitel nasekomykh Dalnego Vostoka Rossii. T. IV" year="2000">Belokobylskij (2000b)</bibRefCitation>
.
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</subSubSection>
</treatment>
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<mods:dateIssued>1758</mods:dateIssued>
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</mods:place>
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<mods:location>
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</mods:location>
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<taxonomicName LSID="urn:lsid:zoobank.org:act:7D3655DE-AD40-4554-80DF-F119AC05CBB9" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Oestridae" genus="Oestrus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bovis" status="spec. nov." zbkClass="Insecta" zbkKingdom="Animalia" zbkOrder="Diptera">Oestrus bovis</taxonomicName>
[
<taxonomicNameLabel rank="species">spec. nov.</taxonomicNameLabel>
]
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<paragraph pageNumber="584">
OE. alis maculatis, thorace flavo fascia fusca, abdomine flavo apice nigro.
<emphasis italics="true">Fn. svec.</emphasis>
1024.
<emphasis italics="true">It. Wgot.</emphasis>
202.
</paragraph>
<paragraph pageNumber="584">
<emphasis italics="true">Raj. ins</emphasis>
. 271
</paragraph>
<paragraph pageNumber="584">
<emphasis italics="true">Derham theolog. l.</emphasis>
8.
<emphasis italics="true">c.</emphasis>
6.
<emphasis italics="true">n.</emphasis>
11.
</paragraph>
<paragraph pageNumber="584">
<emphasis italics="true">Vallisn. nat.</emphasis>
1.
<emphasis italics="true">t.</emphasis>
28.
</paragraph>
<paragraph pageNumber="584">
<emphasis italics="true">Reaum. ins.</emphasis>
4.
<emphasis italics="true">t.</emphasis>
36-38.
</paragraph>
</subSubSection>
<subSubSection type="distribution">
<paragraph pageNumber="584">
<emphasis italics="true">Habitat intra</emphasis>
Boum
<emphasis italics="true">dorsum.</emphasis>
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageNumber="584">
<emphasis italics="true">Hoc illud insectum-cui nomen Asilo</emphasis>
</paragraph>
<paragraph pageNumber="584">
<emphasis italics="true">Romanum est, Oestrum graeci vertere vocantes,</emphasis>
</paragraph>
<paragraph pageNumber="584">
<emphasis italics="true">Asper, acerba sonans, quo toto exterrita silvis</emphasis>
</paragraph>
<paragraph pageNumber="584">
<emphasis italics="true">Diffugiunt armenta.</emphasis>
</paragraph>
</subSubSection>
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<mods:namePart id="96BDF8AE01498BB9B04BA76FB1C9F571">Kezo, Kezhocuyi</mods:namePart>
<mods:nameIdentifier id="3DDE6E90D26F026E9247D79A973A8197" type="ORCID">https://orcid.org/0000-0002-3723-0462</mods:nameIdentifier>
<mods:affiliation id="E60226D6C1718EA64426969429C5CECE">Centre for Advanced Studies in Botany, University of Madras, Guindy Campus, Chennai 600025, Tamil Nadu, India</mods:affiliation>
</mods:name>
<mods:name id="C400306059E90099C6BFD2705FCBF21F" type="personal">
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<mods:roleTerm id="87898DC1CD249D449F8B30EC86477076">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7311CFCBFB39C28F1AE0097A96984B35">Karunarathna, Samantha C.</mods:namePart>
<mods:nameIdentifier id="D2894B1C81DDB29D102CF3418C3B5CE0" type="ORCID">https://orcid.org/0000-0001-7080-0781</mods:nameIdentifier>
<mods:affiliation id="14CFFF7D322631295A46F26A0355653B">Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, China &amp; National Institute of Fundamental Studies (NIFS), Kandy, Sri Lanka</mods:affiliation>
</mods:name>
<mods:name id="25FC7C21A0B6D89D6D43C638862D6CA7" type="personal">
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</mods:role>
<mods:namePart id="DF2D897D143110C0AD21F03DB37531E0">Yang, Erfu</mods:namePart>
<mods:nameIdentifier id="72F7AA4D845926050B2BC346027946DE" type="ORCID">https://orcid.org/0000-0003-2385-6402</mods:nameIdentifier>
<mods:affiliation id="78361E3BB18E461DEB42109C5B0ED877">Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, China &amp; Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
</mods:name>
<mods:name id="4AD076EE03BD480BA59014D94AF3B487" type="personal">
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<mods:roleTerm id="B265793BF37563351B69075C0626CF98">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B577756B411E3537FBAAB12CBEEFE34">Zhao, Changlin</mods:namePart>
<mods:nameIdentifier id="D8FA60275A664DE8B72D038D9F1F8ACC" type="ORCID">https://orcid.org/0000-0002-1395-591X</mods:nameIdentifier>
<mods:affiliation id="4EF618DFF76D6B19880EF37B32130302">College of Biodiversity Conservation, Southwest Forestry University, Kunming 650224, China</mods:affiliation>
</mods:name>
<mods:name id="4CB1BB1DCD4DDE121FFA8AC87359E0B3" type="personal">
<mods:role id="79B4135C8BA02DF37C142A2935EF3626">
<mods:roleTerm id="8445625E74555C8EB4C985EA7CF905A5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="49EDF20822646A8E7008751D5421E1B0">Elgorban, Abdallah M.</mods:namePart>
<mods:nameIdentifier id="9A13F152ED9CDDA1692B5167442B3C0A" type="ORCID">https://orcid.org/0000-0003-3664-7853</mods:nameIdentifier>
<mods:affiliation id="02EC1637314B7AB6F1FE2AFE21807B53">Department of Botany and Microbiology, College of Science, King Saud University, Riyadh 11451, Saudi Arabia</mods:affiliation>
</mods:name>
<mods:name id="1161B7CD0D8EEB838C903B9828AA87F4" type="personal">
<mods:role id="2CB16DF79570F3C641CCD2601B100288">
<mods:roleTerm id="BEE9C74BBE37300525056A345465BF0E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB805127CE9014BC88E9BAB69F9B7188">Tibpromma, Saowaluck</mods:namePart>
<mods:nameIdentifier id="2DAA01187E630A44C6865C03E84DB1DE" type="ORCID">https://orcid.org/0000-0002-4706-6547</mods:nameIdentifier>
<mods:affiliation id="183609F872890ED61B600406C3F09E1A">Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, China</mods:affiliation>
</mods:name>
<mods:name id="19014A7741B43D770FC27BCA34446B5F" type="personal">
<mods:role id="125ECDEEBA39F5C20ABA2A43F2C62730">
<mods:roleTerm id="D4F94BB3B29325C63576795F0B31C3E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="850B2C4B48C5646E93F6696689404C62">Kaliyaperumal, Malarvizhi</mods:namePart>
<mods:nameIdentifier id="BDF15F2C17062042453D5DF9843AC468" type="ORCID">https://orcid.org/0000-0002-1218-3778</mods:nameIdentifier>
<mods:affiliation id="177836F78E42235C317E8F32E117CC2F">Centre for Advanced Studies in Botany, University of Madras, Guindy Campus, Chennai 600025, Tamil Nadu, India</mods:affiliation>
<mods:nameIdentifier id="08759552596E12E708EA90B3BA1E9706" type="email">malar.kaliyaperumal@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="426AEAD166BE5D945A220FB3796A2621">text</mods:typeOfResource>
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<mods:title id="56664C4CCEF8DDF2C017E6E71092A9C7">MycoKeys</mods:title>
</mods:titleInfo>
<mods:part id="664B6EB305EDA85CD307664CB64F9B39">
<mods:date id="427B699225A3FC3CADB7DD772CD597DC">2024</mods:date>
<mods:detail id="5FA416CBD99B68279FFD261EC44E4734" type="pubDate">
<mods:number id="B3084D0D3E08DF107DD032C7FF09C261">2024-02-05</mods:number>
</mods:detail>
<mods:detail id="AE75750DCF13A1A13A8DDFFF9DBF7E04" type="volume">
<mods:number id="4300CCB16967C21E219B3C25CF3E927A">102</mods:number>
</mods:detail>
<mods:extent id="40C8C539F7C412C0D5D5574DF738F807" unit="page">
<mods:start id="225CDC787B6E93C83D7B94147639A5CE">29</mods:start>
<mods:end id="D2916C99379823C2C493015A5B00C3B1">54</mods:end>
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<mods:location id="43D9310AEA3349B3EBED7C3CEBCD4868">
<mods:url id="C072AA8426C26E2CEF3D38D684536E7B">http://dx.doi.org/10.3897/mycokeys.102.117067</mods:url>
</mods:location>
<mods:classification id="B2F91F571A0FBD746902D78A5B5963E8">journal article</mods:classification>
<mods:identifier id="473AB05837EB3A583F1E5BED6152799B" type="DOI">http://dx.doi.org/10.3897/mycokeys.102.117067</mods:identifier>
<mods:identifier id="AB47DE2F96593E40CF61318A8F62E10A" type="Pensoft-Pub">1314-4049-102-29</mods:identifier>
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</mods:mods>
<treatment id="62082AE9516E542FB4B61BFE5B345743" LSID="urn:lsid:plazi:treatment:62082AE9516E542FB4B61BFE5B345743" httpUri="http://treatment.plazi.org/id/62082AE9516E542FB4B61BFE5B345743" lastPageNumber="29" pageId="0" pageNumber="29">
<subSubSection id="5AF2E0B5F77AC7BD11A8CD98C4C50A05" pageId="0" pageNumber="29" type="nomenclature">
<paragraph id="386DD79B1E686B57019C41C410DED53D" pageId="0" pageNumber="29">
<taxonomicName id="2BF937C4BFA1BCA84418456136C6565A" LSID="62082AE9-516E-542F-B4B6-1BFE5B345743" authority="S. Gunaseelan &amp; M. Kaliyaperumal" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus" status="sp. nov.">Tropicoporus indicus S. Gunaseelan &amp; M. Kaliyaperumal</taxonomicName>
<taxonomicNameLabel id="0BF4AFA386FBB85951355A4EDEEA2573" pageId="0" pageNumber="29">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="46251EDD8D25EBE3E279290BBF0DC49A" pageId="0" pageNumber="29" type="description">
<paragraph id="5B97403B05901000C06D035BE6EFF4FB" pageId="0" pageNumber="29">
<figureCitation id="2E5081CFE91A99CFEC099B4C696B618F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tropicoporus indicus (MUBL 1083 holotype) A basidiomata (Holotype) B basidiome (SMK-MK 2 a- Isotype) C pore surface D cross-section of basidiome (arrows indicating stratified tube layers) E hymenial setae F cystidioles G-J basidiospores: G basidiospores in water H basidiospores in KOH I basidiospores in cotton blue J basidiospores in Melzer's reagent. Scale bars: 1 cm (A-D); 5 µm (E-J)." figureDoi="10.3897/mycokeys.102.117067.figure5" httpUri="https://binary.pensoft.net/fig/979926" pageId="0" pageNumber="29">Figs 5</figureCitation>
<figureCitation id="F671952AD4917BD5CC18DA591B58C2D5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tropicoporus indicus (MUBL 1083 holotype) A contextual hyphae B tramal hyphae C hymenial setae D cystidioles E basidioles F basidia G basidiospores. Scale bars: 5 µm." figureDoi="10.3897/mycokeys.102.117067.figure6" httpUri="https://binary.pensoft.net/fig/979927" pageId="0" pageNumber="29">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BAF800DC178929EBB9A16C954EE6E41C" pageId="0" pageNumber="29" type="etymology">
<paragraph id="0B02D8C13FD011D749F2A58C7573D613" pageId="0" pageNumber="29">Etymology.</paragraph>
<paragraph id="C800399C1B4C0FA04A3167D43CA96491" pageId="0" pageNumber="29">
The species epithet &quot;
<emphasis id="044F20B73D669DB94A5EA2B61957BEAC" italics="true" pageId="0" pageNumber="29">indicus</emphasis>
&quot; (Lat.): referring to the species being collected from India.
</paragraph>
</subSubSection>
<subSubSection id="368C4DC75023C71E89CC7ADD0FDF838F" pageId="0" pageNumber="29" type="diagnosis">
<paragraph id="4D9597E8B1BF60BB4F27FA8D361F6607" pageId="0" pageNumber="29">Diagnosis.</paragraph>
<paragraph id="EF899FD667D892D618687B0837171020" pageId="0" pageNumber="29">
<taxonomicName id="FA0501C519CADD63F1CB94AAC0505925" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="744132A9C18066647965B63534AE0F56" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
is characterised by applanate to meagrely triquetrous basidiome with concentrically zonate, sulcate, glabrous, deeply cracked to rimose pilear surface, homogenous context, acute margin, mono-dimitic hyphal system, presence of cystidioles, subglobose to broadly ellipsoid basidiospores measuring 5-6
<normalizedToken id="FEB817D4525411C29B3E2585ADE237C0" originalValue="×">x</normalizedToken>
4.2-4.9
<normalizedToken id="7E918504528DB828C6EE214A14020E0E" originalValue="μm">μm</normalizedToken>
.
</paragraph>
<caption id="F7F7BEC721272C72A5CFC1E0E1B1016E" doi="10.3897/mycokeys.102.117067.figure5" httpUri="https://binary.pensoft.net/fig/979926" pageId="0" pageNumber="29" start="Figure 5" startId="F5">
<paragraph id="BEE5D3E91315B53A61FC158B7BA185C6" pageId="0" pageNumber="29">
<emphasis id="8F4AE627BCC5A69425279DEED6D03C90" bold="true" pageId="0" pageNumber="29">Figure 5.</emphasis>
<taxonomicName id="04BE451B3C8F09BCB2D7F23246C83DFB" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="D455BB614D1F11AA5CC819489486E994" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
(MUBL1083 holotype)
<emphasis id="1BFC7E586A241C25522011B93504BA69" bold="true" pageId="0" pageNumber="29">A</emphasis>
basidiomata (Holotype)
<emphasis id="0AF7E936E01BC0AD876868A59FC844C6" bold="true" pageId="0" pageNumber="29">B</emphasis>
basidiome (SMK-MK2a- Isotype)
<emphasis id="66CBFA7998079088C115A3F048A39DC5" bold="true" pageId="0" pageNumber="29">C</emphasis>
pore surface
<emphasis id="9DA68684183AE8E546AC05B172031183" bold="true" pageId="0" pageNumber="29">D</emphasis>
cross-section of basidiome (arrows indicating stratified tube layers)
<emphasis id="4C4572BF89BDE299D826DA962D96C3BC" bold="true" pageId="0" pageNumber="29">E</emphasis>
hymenial setae
<emphasis id="C1258520A83F2E3E47C8DF9CE8FDEA97" bold="true" pageId="0" pageNumber="29">F</emphasis>
cystidioles
<emphasis id="98117C7B4DA3A802CB87F1D21E80F871" bold="true" pageId="0" pageNumber="29">G-J</emphasis>
basidiospores:
<emphasis id="C71223762EED1995AD2EBDA60AAAE7B7" bold="true" pageId="0" pageNumber="29">G</emphasis>
basidiospores in water
<emphasis id="D3F7B13D88B643D602939871B3FB730A" bold="true" pageId="0" pageNumber="29">H</emphasis>
basidiospores in KOH
<emphasis id="D00A54486967153821948C4346314343" bold="true" pageId="0" pageNumber="29">I</emphasis>
basidiospores in cotton blue
<emphasis id="25A034D9FFD0E4FF95A1E322BEC90426" bold="true" pageId="0" pageNumber="29">J</emphasis>
basidiospores in
<normalizedToken id="395FCE2E8AA71DD46801BD7C9977FD0B" originalValue="Melzer’s">Melzer's</normalizedToken>
reagent. Scale bars: 1 cm (
<emphasis id="399895FDD4F5A6B8771A2C3E2CBBDFB7" bold="true" pageId="0" pageNumber="29">A-D</emphasis>
); 5
<normalizedToken id="573B4995A57AD8D6034AF5B2F45DD600" originalValue="µm">µm</normalizedToken>
(
<emphasis id="60FDFD3C201F4CAE26B9CAC8518D1794" bold="true" pageId="0" pageNumber="29">E-J</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B153723D747BFB2999D29B76464C52D4" pageId="0" pageNumber="29" type="materials_examined">
<paragraph id="C054F912DFEA41B6962D22E0E300BE2D" pageId="0" pageNumber="29">
<typeStatus id="DE05FB7F564E07A637D0135F674E73A8">Type</typeStatus>
.
</paragraph>
<paragraph id="7E1762E532BF4ADB76AE04320BA14B7E" pageId="0" pageNumber="29">
<materialsCitation id="65D5A8F4A463E2F68D1FC8D2E53391B5" collectingDate="2022-09-29" country="India" location="Kalvarayan Hills" municipality="Kallakurichi District" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="Holotype">
<collectingCountry id="838D83A17E568A2EAFFABB656510106E" name="India">India</collectingCountry>
,
<collectingRegion id="E03789AA88E377A64B85919D5C46A908" country="India" name="Tamil Nadu">Tamil Nadu</collectingRegion>
,
<collectingMunicipality id="C64899D78194C957A1374D7F56367A04">Kallakurichi District</collectingMunicipality>
,
<location id="B92D70A2586D84DCE5CA4F1DDCAF65D6" LSID="urn:lsid:plazi:treatment:62082AE9516E542FB4B61BFE5B345743:B92D70A2586D84DCE5CA4F1DDCAF65D6" country="India" municipality="Kallakurichi District" name="Kalvarayan Hills" stateProvince="Tamil Nadu">Kalvarayan Hills</location>
; 11°91'30&quot;N, 78°57'86&quot;E;
<collectingDate id="A3E21C5CB676E09428D79FFE3028B91E" value="2022-09-29">29 Sep 2022</collectingDate>
; Sugantha Gunaseelan; on living angiosperm tree of
<taxonomicName id="7356494E7AE3E2204FE06E49D24E9E46" class="Dicotyledoneae" family="Fabaceae" genus="Albizia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Albizia amara" order="Rosales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="amara">
<emphasis id="D061194C9A6FD987B7D27FFC4E0A674B" italics="true" pageId="0" pageNumber="29">Albizia amara</emphasis>
</taxonomicName>
; SMK- MK2 (MUBL1083,
<typeStatus id="EF522058354CCD8E6F1D6F37889759D1" type="holotype">Holotype</typeStatus>
); GenBank: OR272293 (ITS); OR272338 (nLSU)
</materialsCitation>
.
</paragraph>
<caption id="0417FD3D2E1C7D5D6F79F5571170ECB1" doi="10.3897/mycokeys.102.117067.figure6" httpUri="https://binary.pensoft.net/fig/979927" pageId="0" pageNumber="29" start="Figure 6" startId="F6">
<paragraph id="3404361760D4ABE24912E5BB43A51F73" pageId="0" pageNumber="29">
<emphasis id="81EE7AA8516163E33DB81F4D737DA260" bold="true" pageId="0" pageNumber="29">Figure 6.</emphasis>
<taxonomicName id="3140D02FE7705B07BE1AC9FAFB909EA3" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="8A1381499FD1A636CECD2AB74F42B836" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
(MUBL1083
<typeStatus id="43CB44B3B279BC9EC3F70F74F354C36E" type="holotype">holotype</typeStatus>
)
<emphasis id="8B4B091459877DA2B9FA07399DB6C7B8" bold="true" pageId="0" pageNumber="29">A</emphasis>
contextual hyphae
<emphasis id="A710C7B2FEC6853120673896E6216D8D" bold="true" pageId="0" pageNumber="29">B</emphasis>
tramal hyphae
<emphasis id="571A649858CB441BA96CD9BF8CCD2735" bold="true" pageId="0" pageNumber="29">C</emphasis>
hymenial setae
<emphasis id="C603982024A17BA0804568D4C6DEA17E" bold="true" pageId="0" pageNumber="29">D</emphasis>
cystidioles
<emphasis id="0D67633B6FC55CBA2A7B3278D4BC234B" bold="true" pageId="0" pageNumber="29">E</emphasis>
basidioles
<emphasis id="432667E0EFB1A32AA1442E3E7E4A8806" bold="true" pageId="0" pageNumber="29">F</emphasis>
basidia
<emphasis id="E5C6F6A2267748611CB9FEF583FDF2A8" bold="true" pageId="0" pageNumber="29">G</emphasis>
basidiospores. Scale bars: 5
<normalizedToken id="84CD5828A7E50191D668CCFE9945EBC1" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1FDDB638A1442F8A46F22F5B0CE00762" pageId="0" pageNumber="29" type="description">
<paragraph id="98DBCB21A5C82F7F6F7998E865A6712C" pageId="0" pageNumber="29">Description.</paragraph>
<paragraph id="4FAF4EB773F10A8041B31FABF94617D1" pageId="0" pageNumber="29">
<emphasis id="4A07B64BD97536DC7316937B47CEB234" bold="true" italics="true" pageId="0" pageNumber="29">Basidiome</emphasis>
perennial, pileate, woody and without distinctive odour or taste when fresh, hard when dry.
<emphasis id="93D13D632F46E25A5324C9BBEF7E0C27" bold="true" italics="true" pageId="0" pageNumber="29">Pilei</emphasis>
applanate to meagrely triquetrous, projecting up to 5.5 cm, 7 cm wide and 4 cm thick at the base; pileal surface concentrically zonate, sulcate, glabrous, deeply cracked to rimose near attachment, dark brown (7E4), greyish-brown (7F3).
<emphasis id="A7E0D09CCC88EAACBA811B122321522D" bold="true" italics="true" pageId="0" pageNumber="29">Margin</emphasis>
velutinate, 2 mm thick, acute, brown (6E7).
<emphasis id="B3A7983052C52B00E24F82DC7479311E" bold="true" italics="true" pageId="0" pageNumber="29">Pore surface</emphasis>
light brown (5D5) to yellowish-brown (5E7); pores circular, 4-6 per mm.
<emphasis id="FAABF0438CCB8B9A8CCF780DB31BE4A5" bold="true" italics="true" pageId="0" pageNumber="29">Context</emphasis>
homogenous, up to 0.8 cm thick, brown (6E8) to dark brown (6F8).
<emphasis id="5773F0A59CEB0E64AD1DC1EA3068B995" bold="true" italics="true" pageId="0" pageNumber="29">Tubes</emphasis>
woody hard, up to 2.5 cm long, brown (6E7); tube layers stratified, each stratum up to 0.5 cm long.
</paragraph>
</subSubSection>
<subSubSection id="141A9D6385836279BCD32ED43E141274" pageId="0" pageNumber="29" type="hyphal structures">
<paragraph id="1C2E9AD4E03BA6AD7ACE1130AD4A0549" pageId="0" pageNumber="29">Hyphal structures.</paragraph>
<paragraph id="4B849D2824CEA3990096DA2F9029E339" pageId="0" pageNumber="29">Hyphal system monomitic in the context and dimitic in the trama, tissue darkening with KOH without swelling</paragraph>
</subSubSection>
<subSubSection id="0CD9E240ACCB5F7C663FFBD92544170A" pageId="0" pageNumber="29" type="context">
<paragraph id="443A9278FCEE2A993C7AEE546B8FD941" pageId="0" pageNumber="29">Context.</paragraph>
<paragraph id="62D8E7A78673314CE750C523DF96C78E" pageId="0" pageNumber="29">
Generative hyphae, thin to thick-walled, hyaline to golden yellow, simple septate, rarely branched, 2-5
<normalizedToken id="2567CF407934D51785B603BADD37E539" originalValue="μm">μm</normalizedToken>
diam.
</paragraph>
</subSubSection>
<subSubSection id="75E574BF614993C299550F9CCA1DA46F" pageId="0" pageNumber="29" type="trama">
<paragraph id="B9ECE6697971C4404F00486FADC0E489" pageId="0" pageNumber="29">Trama.</paragraph>
<paragraph id="2A73862BE2196029BBF1DD80DD26168D" pageId="0" pageNumber="29">
Generative hyphae dominant, thin to thick-walled, hyaline to pale yellow, septate, occasionally branched, 2-4.5
<normalizedToken id="1CDEC11197425FE8CA29F48EEFAA561F" originalValue="μm">μm</normalizedToken>
diam. Skeletal hyphae thick-walled with narrow to wide lumen, yellowish-brown, aseptate, unbranched, 2-4
<normalizedToken id="C9FCFA4431D5BA6841EA7F02CF2BCF93" originalValue="μm">μm</normalizedToken>
diam.
</paragraph>
</subSubSection>
<subSubSection id="FBDDA3000150531EB66506A207CC41A8" pageId="0" pageNumber="29" type="hymenium">
<paragraph id="EBDDC36B0F4FD6F84F2236C697940DD7" pageId="0" pageNumber="29">Hymenium.</paragraph>
<paragraph id="DAEB2E79F4C8CE6E6E8EF0C20D86E894" pageId="0" pageNumber="29">
Hymenial setae dark brown, thick-walled, ventricose to subulate with sharp to blunt tips, 7-28
<normalizedToken id="607304B009249B4C40DFB2FC3F93C9B7" originalValue="×">x</normalizedToken>
3-5
<normalizedToken id="EFAEE034537567CF407C25F7F5131E8B" originalValue="μm">μm</normalizedToken>
. Cystidia absent. Cystidioles hyaline, thin-walled, ventricose to fusoid with elongated tapering apical portion, 5-21
<normalizedToken id="09EBEC289E4A747B587B77E89B7ACD98" originalValue="×">x</normalizedToken>
3-5
<normalizedToken id="2D6FACE1178FDCC2FC6793021D69672B" originalValue="μm">μm</normalizedToken>
. Basidia clavate to broadly clavate, 7-17
<normalizedToken id="C7CE3238EBB532C414159EE1936ECD38" originalValue="×">x</normalizedToken>
3-6
<normalizedToken id="E989E5236DB34BFE676E74CB8F2CEF26" originalValue="μm">μm</normalizedToken>
, with four sterigmata and a simple septum at the base. Basidioles clavate, 5-14
<normalizedToken id="868F6BC28E599AB4402CA0090B14C50B" originalValue="×">x</normalizedToken>
3-5
<normalizedToken id="6E522470C071D757166C9A8D234A162E" originalValue="μm">μm</normalizedToken>
. Basidiospores broadly ellipsoid to subglobose, pale yellow in water, turning golden yellow to brown in KOH, thick-walled, smooth, CB ̄, IKI ̄, (5-) 5.3-5.8 (- 6)
<normalizedToken id="FC7D71B7C78D3840173D035BE0BB85CD" originalValue="×">x</normalizedToken>
(4.2-) 4.7-4.9
<normalizedToken id="E18EB24817901976567F0180CB8DF878" originalValue="μm">μm</normalizedToken>
(n = 50/2), Q = 1.16 (Q range 1.05-1.3).
</paragraph>
</subSubSection>
<subSubSection id="2F8EEA2EEBAF0BA881C42DCA287E5FBE" pageId="0" pageNumber="29" type="habitat">
<paragraph id="1A6CC74B66624F147CD6512684B29BAF" pageId="0" pageNumber="29">Habitat and distribution.</paragraph>
<paragraph id="F0747385A9392E373997D9D88F4538DD" pageId="0" pageNumber="29">
Basidiomes were found on living trees of
<taxonomicName id="768FD47B32D89D5C0490A0351D54C084" authorityName="L.W.Zhou, Y.C.Dai &amp; Sheng H.Wu" authorityYear="2015" family="Fabaceae" kingdom="Fungi" lsidName="" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="family">Fabaceae</taxonomicName>
members (
<taxonomicName id="5BF3EC14EF19DA47CDC26296A0BC433F" class="Dicotyledoneae" family="Fabaceae" genus="Albizia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Albizia amara" order="Rosales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="amara">
<emphasis id="3C3B38163648D176BF1F51C684ED06EF" italics="true" pageId="0" pageNumber="29">Albizia amara</emphasis>
</taxonomicName>
and
<taxonomicName id="EC65242800100FA474C5C95030CDD797" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Prosopis cineraria" order="Fabales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="cineraria">
<emphasis id="33CA6C2825C2B88E1DD2D8F40F362739" italics="true" pageId="0" pageNumber="29">Prosopis cineraria</emphasis>
</taxonomicName>
), distributed in Kalvarayan Hills, Kallakurichi District, Tamil Nadu, India.
</paragraph>
</subSubSection>
<subSubSection id="7BA82C9FD7C07A575AF531310B8DE92C" pageId="0" pageNumber="29" type="materials_examined">
<paragraph id="FF1D6B35849BA76C1B7E48E05707C2E8" pageId="0" pageNumber="29">Additional material examined.</paragraph>
<paragraph id="483A64701DF586D4F190A14F5B631C80" pageId="0" pageNumber="29">
<materialsCitation id="88EC66C3741B1BA40BE3BB40DC7AB17A" collectingDate="2022-09-29" country="India" location="Kalvarayan Hills" municipality="Kallakurichi District" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="Paratype">
<collectingCountry id="84B6BA94B8495944C0BEB96621C5CF14" name="India">India</collectingCountry>
,
<collectingRegion id="68AC01C23429F3484422E1175201FEE0" country="India" name="Tamil Nadu">Tamil Nadu</collectingRegion>
,
<collectingMunicipality id="367EDD920F008D3F2E3DC595057A54AA">Kallakurichi District</collectingMunicipality>
,
<location id="113318AC5CEE30896BCBCF5E5FEC6653" LSID="urn:lsid:plazi:treatment:62082AE9516E542FB4B61BFE5B345743:113318AC5CEE30896BCBCF5E5FEC6653" country="India" municipality="Kallakurichi District" name="Kalvarayan Hills" stateProvince="Tamil Nadu">Kalvarayan Hills</location>
; 11°90'39&quot;N, 78°55'69&quot;E; on a living angiosperm tree (
<taxonomicName id="3332666F69E88D433B1F97A2BC251725" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Prosopis cineraria" order="Fabales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="cineraria">
<emphasis id="52B2E9050875A720B86A5ACBCD69457A" italics="true" pageId="0" pageNumber="29">Prosopis cineraria</emphasis>
</taxonomicName>
);
<collectingDate id="7F1186D5D9DA111D96C70E3B5A30F47A" value="2022-09-29">29 Sep 2022</collectingDate>
; Kezhocuyi Kezo; SMK-MK2a (MUBL1084,
<typeStatus id="124DBAB3BE02C44BEFA1DFC7191BAC66" type="paratype">Paratype</typeStatus>
); GenBank: OR272294 (ITS); OR272339 (nLSU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="21542072A96A093A2F4DAC979EEB7137" pageId="0" pageNumber="29" type="notes">
<paragraph id="7508C31DAE256A46DAD9257199805ABA" pageId="0" pageNumber="29">Notes.</paragraph>
<paragraph id="CBF2696AB924C521661F3942FFB366C1" pageId="0" pageNumber="29">
Phylogenetically,
<taxonomicName id="A7D15F9C6E91E1A2005759ABCD11FB23" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="F6B16A66DC6178BFFC34B79ABB5AE0D4" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
was recovered in the
<taxonomicName id="757C53F94C54C1AA1D425C16B4E455F0" lsidName="T. linteus" pageId="0" pageNumber="29" rank="species" species="linteus">
<emphasis id="706063A4D60CC79DE7983763834A5959" italics="true" pageId="0" pageNumber="29">T. linteus</emphasis>
</taxonomicName>
clade.
<taxonomicName id="290473D566A981A2EB27449C51AF4F8F" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="AB61F023984706CE3D47B8B0572DEBAC" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="F9ADBB6FF122D484DDE26F70CC2DF57B" lsidName="T. linteus" pageId="0" pageNumber="29" rank="species" species="linteus">
<emphasis id="04270A33FE3F0025C83083D15598BEF6" italics="true" pageId="0" pageNumber="29">T. linteus</emphasis>
</taxonomicName>
by sharing, pileate, dimidiate basidiomes, concentrically sulcate pilear surface, zonate context, smaller pores (5-7/mm), a mono-dimitic hyphal system and presence of cystidioles. While varying in the nature of cracks,
<taxonomicName id="0997FF0E8004F89E6C761DEB8ABF6600" lsidName="T. linteus" pageId="0" pageNumber="29" rank="species" species="linteus">
<emphasis id="B3977EB245F3D6E499F377572B9FADAB" italics="true" pageId="0" pageNumber="29">T. linteus</emphasis>
</taxonomicName>
has more or less cracked basidiomes,
<taxonomicName id="010052204A51053ADE6CF54A429B0E86" lsidName="T. indicus" pageId="0" pageNumber="29" rank="species" species="indicus">
<emphasis id="8C0C32E4AA0732285609918F39C76C90" italics="true" pageId="0" pageNumber="29">T. indicus</emphasis>
</taxonomicName>
has irregular deep cracks in basidiomes, with larger basidiospores (
<taxonomicName id="72830352DCE2554A7235626789FCD6DD" lsidName="T. linteus" pageId="0" pageNumber="29" rank="species" species="linteus">
<emphasis id="5F555F4C4C97E454B5E3BEFC19F9FEC5" italics="true" pageId="0" pageNumber="29">T. linteus</emphasis>
</taxonomicName>
4.8-5.7
<normalizedToken id="48D049455B43C16187C8E379AE62A386" originalValue="×">x</normalizedToken>
3.8-4.8
<normalizedToken id="0F748C7A7088E2942C1E5059F00FFF76" originalValue="μm">μm</normalizedToken>
and
<taxonomicName id="885DADA9BE758F105C9062726B5CB986" lsidName="T. indicus" pageId="0" pageNumber="29" rank="species" species="indicus">
<emphasis id="8DBB3F8BF3078F77C60B6DCF0673F6D1" italics="true" pageId="0" pageNumber="29">T. indicus</emphasis>
</taxonomicName>
5-6
<normalizedToken id="B6DDC768A90196223CB40D1A2DD4653C" originalValue="×">x</normalizedToken>
4.2-4.9
<normalizedToken id="3A4BC333663BCD1B7D0C6DD94C2C55C8" originalValue="μm">μm</normalizedToken>
) (
<bibRefCitation id="3E01B73412D6C6BC5C4B6666A1BB04AB" DOI="https://doi.org/10.1007/s13225-012-0202-9" author="Tian, XM" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="159 - 169" refId="B25" refString="Tian, XM, Yu, HY, Zhou, LW, Decock, C, Vlasak, J, Dai, YC, 2013. Phylogeny and taxonomy of the Inonotus linteus complex. Fungal Diversity 58 (1): 159 - 169, DOI: https://doi.org/10.1007/s13225-012-0202-9" title="Phylogeny and taxonomy of the Inonotus linteus complex." url="https://doi.org/10.1007/s13225-012-0202-9" volume="58" year="2013">Tian et al. 2013</bibRefCitation>
;
<bibRefCitation id="7AAEDE9F84BFC8CB20E630A9799C48F9" DOI="https://doi.org/10.1007/s13225-021-00496-4" author="Wu, F" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="1 - 192" refId="B30" refString="Wu, F, Zhou, LW, Vlasak, J, Dai, YC, 2022. Global diversity and systematics of Hymenochaetaceae with poroid hymenophore. Fungal Diversity 113 (1): 1 - 192, DOI: https://doi.org/10.1007/s13225-021-00496-4" title="Global diversity and systematics of Hymenochaetaceae with poroid hymenophore." url="https://doi.org/10.1007/s13225-021-00496-4" volume="113" year="2022">Wu et al. 2022</bibRefCitation>
).
<taxonomicName id="B85B63A409C56D98CF489E0C0BE724E7" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="0A32D642975E3FD1C106B145EE1FA9F2" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="C87F1423FBB08087E5E903B345951218" lsidName="T. rudis" pageId="0" pageNumber="29" rank="species" species="rudis">
<emphasis id="CFD24BEFE5F8CE1742D9D7CF1D7332B1" italics="true" pageId="0" pageNumber="29">T. rudis</emphasis>
</taxonomicName>
share a mono-dimitic hyphal system, but
<taxonomicName id="D7701BAFAF7D08BC69DFF9216C40D5F2" lsidName="T. indicus" pageId="0" pageNumber="29" rank="species" species="indicus">
<emphasis id="A49328DA1EAE65E722B428386F04EA60" italics="true" pageId="0" pageNumber="29">T. indicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="62228A9067A01ECC79E5741838BED530" lsidName="T. rudis" pageId="0" pageNumber="29" rank="species" species="rudis">
<emphasis id="61E5AED4F8C4E1141AF242602451453C" italics="true" pageId="0" pageNumber="29">T. rudis</emphasis>
</taxonomicName>
in having zonate, sulcate, deeply cracked to rimose basidiomes and larger pores (4-6/mm). In these regards,
<taxonomicName id="06F0CAD7D2B8FF2AA67F100D4B920138" lsidName="T. rudis" pageId="0" pageNumber="29" rank="species" species="rudis">
<emphasis id="97190D5527A9B3C78C1EEC564343B500" italics="true" pageId="0" pageNumber="29">T. rudis</emphasis>
</taxonomicName>
is characterised by fulvous, velvety, concentrically zonate, matted, rimose basidiome and smaller pores (6-7/mm) (
<bibRefCitation id="D2E879DC954C6D669C9F1650E39E0E6B" DOI="https://doi.org/10.1007/s13225-021-00496-4" author="Wu, F" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="1 - 192" refId="B30" refString="Wu, F, Zhou, LW, Vlasak, J, Dai, YC, 2022. Global diversity and systematics of Hymenochaetaceae with poroid hymenophore. Fungal Diversity 113 (1): 1 - 192, DOI: https://doi.org/10.1007/s13225-021-00496-4" title="Global diversity and systematics of Hymenochaetaceae with poroid hymenophore." url="https://doi.org/10.1007/s13225-021-00496-4" volume="113" year="2022">Wu et al. 2022</bibRefCitation>
).
<taxonomicName id="51CFD021F464E69FD8BD19D25CB2E48B" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="BD68A9F67E1CFE47A3D7D769F69EA4E2" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="DC4ECCEBBADE26624948B0A6D011ECEF" lsidName="T. angustisulcatus" pageId="0" pageNumber="29" rank="species" species="angustisulcatus">
<emphasis id="1704375BF970D9C294A131BAC7D25E3D" italics="true" pageId="0" pageNumber="29">T. angustisulcatus</emphasis>
</taxonomicName>
,
<taxonomicName id="22EB3450D6EF3EB914A2ED3173DC8544" lsidName="T. excentrodendri" pageId="0" pageNumber="29" rank="species" species="excentrodendri">
<emphasis id="08F1F80D2B38E6906DFF76F454E4C563" italics="true" pageId="0" pageNumber="29">T. excentrodendri</emphasis>
</taxonomicName>
,
<taxonomicName id="8C0E34D1E4F32D5024ADC9BBBFA57F31" lsidName="T. lineatus" pageId="0" pageNumber="29" rank="species" species="lineatus">
<emphasis id="BED1EF3E56689C26F45FA7A7C7FE4039" italics="true" pageId="0" pageNumber="29">T. lineatus</emphasis>
</taxonomicName>
and
<taxonomicName id="C3B1B86418960A0C08E0E27739B7E135" lsidName="T. substratificans" pageId="0" pageNumber="29" rank="species" species="substratificans">
<emphasis id="F67AFBEE50F5FCA3D161CA3AAD708340" italics="true" pageId="0" pageNumber="29">T. substratificans</emphasis>
</taxonomicName>
in having mono-dimitic hyphal system and a cracked basidiome (
<bibRefCitation id="64BB0B816B6959FB42B66D2782633F5B" DOI="https://doi.org/10.1007/s13225-015-0335-8" author="Zhou, LW" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="335 - 347" refId="B31" refString="Zhou, LW, Vlasak, J, Decock, C, Assefa, A, Stenlid, J, Abate, D, Wu, SH, Dai, YC, 2015. Global diversity and taxonomy of the Inonotus linteus complex (Hymenochaetales, Basidiomycota): Sanghuangporus gen. nov., Tropicoporus excentrodendri and T. guanacastensis gen. et spp. nov., and 17 new combinations. Fungal Diversity 77 (1): 335 - 347, DOI: https://doi.org/10.1007/s13225-015-0335-8" title="Global diversity and taxonomy of the Inonotus linteus complex (Hymenochaetales, Basidiomycota): Sanghuangporus gen. nov., Tropicoporus excentrodendri and T. guanacastensis gen. et spp. nov., and 17 new combinations." url="https://doi.org/10.1007/s13225-015-0335-8" volume="77" year="2015">Zhou et al. 2015</bibRefCitation>
;
<bibRefCitation id="365C9053C4A4877ADF6045036A0BC3A8" DOI="https://doi.org/10.1007/s13225-021-00496-4" author="Wu, F" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="1 - 192" refId="B30" refString="Wu, F, Zhou, LW, Vlasak, J, Dai, YC, 2022. Global diversity and systematics of Hymenochaetaceae with poroid hymenophore. Fungal Diversity 113 (1): 1 - 192, DOI: https://doi.org/10.1007/s13225-021-00496-4" title="Global diversity and systematics of Hymenochaetaceae with poroid hymenophore." url="https://doi.org/10.1007/s13225-021-00496-4" volume="113" year="2022">Wu et al. 2022</bibRefCitation>
).
<taxonomicName id="8A7D735461597D07595ACECA099D033E" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="7AC01A0EB90535B9052160AF5D9D1917" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="2887B4C182C3FD5C33E5DD64BE4944DE" lsidName="T. flabellatus" pageId="0" pageNumber="29" rank="species" species="flabellatus">
<emphasis id="B8DB698885E5BD77BC72C66FFF58FEF6" italics="true" pageId="0" pageNumber="29">T. flabellatus</emphasis>
</taxonomicName>
are similar in having a mono-dimitic hyphal system, but differ significantly by having concentrical zones, sulcate, glabrous, deeply-cracked to rimose pilear surface with larger pores (4-6/mm) and broadly ellipsoid to subglobose spores (5-6
<normalizedToken id="F6716CC838A4A323C91607FE08BB4A59" originalValue="×">x</normalizedToken>
4.2-4.9
<normalizedToken id="5FBD43EA80F88F4BE6381F7433FF7359" originalValue="μm">μm</normalizedToken>
).
<taxonomicName id="FC1B44D11C81C48381332AAA0C965DDA" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus flabellatus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="flabellatus">
<emphasis id="C7FE21E2F2A58AC3C3E3DDFEC2A6DA61" italics="true" pageId="0" pageNumber="29">Tropicoporus flabellatus</emphasis>
</taxonomicName>
, in contrast, has a velutinate pilear surface, uncracked basidiomes with smaller pores (7-9/mm) and smaller basidiospores (4.5-5
<normalizedToken id="5A4936E733DB2B94986096871DBDFACE" originalValue="×">x</normalizedToken>
3.5-4
<normalizedToken id="A4C1701FC7C12EC8FE79DD0C6B5C36BB" originalValue="μm">μm</normalizedToken>
) (
<bibRefCitation id="D3C5238A3AE64D51DC8678CA28D1A8DF" DOI="https://doi.org/10.5252/cryptogamie-mycologie2022v43a1" author="Lima, VX" journalOrPublisher="Mycologie" pageId="0" pageNumber="29" pagination="1 - 21" refId="B19" refString="Lima, VX, de Oliveira, VRT, de Lima-Junior, NC, Oliveira-Filho, JRC, Santos, C, Lima, N, Gibertoni, TB, 2022. Taxonomy and phylogenetic analysis reveal one new genus and three new species in Inonotus s.l. (Hymenochaetaceae) from Brazil. Cryptogamie. Mycologie 43 (1): 1 - 21, DOI: https://doi.org/10.5252/cryptogamie-mycologie2022v43a1" title="Taxonomy and phylogenetic analysis reveal one new genus and three new species in Inonotus s. l. (Hymenochaetaceae) from Brazil. Cryptogamie." url="https://doi.org/10.5252/cryptogamie-mycologie2022v43a1" volume="43" year="2022">Lima et al. 2022</bibRefCitation>
).
<taxonomicName id="2230DFB9DD5EA61B590758E07106062A" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="A75CFFF4B858D143E8B78E340EFE5873" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
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and
<taxonomicName id="F87A8A4D81B35A7BABA342DF3938A9D8" lsidName="T. guanacastensis" pageId="0" pageNumber="29" rank="species" species="guanacastensis">
<emphasis id="39E69F2204BF61C8931144860F496456" italics="true" pageId="0" pageNumber="29">T. guanacastensis</emphasis>
</taxonomicName>
are similar in having sulcate, cracked basidiome, stratified tube, mono-dimitic hyphal system and ventricose setae. However, the former differs in larger pores (4-6/mm) and larger basidiospores (
<taxonomicName id="FCFBDDFAFD4F6B02B9A6E02AD699CA4A" lsidName="T. indicus" pageId="0" pageNumber="29" rank="species" species="indicus">
<emphasis id="FD4F9733CA341389FCAE2A0F3EFECB24" italics="true" pageId="0" pageNumber="29">T. indicus</emphasis>
</taxonomicName>
5-6
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4.2-4.9
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vs.
<taxonomicName id="9E8199B862C36D0A21F8DA0C4BBAC532" lsidName="T. guanacastensis" pageId="0" pageNumber="29" rank="species" species="guanacastensis">
<emphasis id="AAACAD3CF1864039C16CFB1990D8F1FE" italics="true" pageId="0" pageNumber="29">T. guanacastensis</emphasis>
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4.1-5
<normalizedToken id="725D64489769E4312F118087D3519BA1" originalValue="×">x</normalizedToken>
3.1-4
<normalizedToken id="43EDD852A765743E4BD7E9BA933C86E4" originalValue="μm">μm</normalizedToken>
) (
<bibRefCitation id="8911BFE71B3217A7E13918CE3335E80E" DOI="https://doi.org/10.1007/s13225-015-0335-8" author="Zhou, LW" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="29" pagination="335 - 347" refId="B31" refString="Zhou, LW, Vlasak, J, Decock, C, Assefa, A, Stenlid, J, Abate, D, Wu, SH, Dai, YC, 2015. Global diversity and taxonomy of the Inonotus linteus complex (Hymenochaetales, Basidiomycota): Sanghuangporus gen. nov., Tropicoporus excentrodendri and T. guanacastensis gen. et spp. nov., and 17 new combinations. Fungal Diversity 77 (1): 335 - 347, DOI: https://doi.org/10.1007/s13225-015-0335-8" title="Global diversity and taxonomy of the Inonotus linteus complex (Hymenochaetales, Basidiomycota): Sanghuangporus gen. nov., Tropicoporus excentrodendri and T. guanacastensis gen. et spp. nov., and 17 new combinations." url="https://doi.org/10.1007/s13225-015-0335-8" volume="77" year="2015">Zhou et al. 2015</bibRefCitation>
).
<taxonomicName id="9593665F2A4A44F9101CB12344E55438" authorityName="S. Gunaseelan &amp; M. Kaliyaperumal" authorityYear="2024" class="Agaricomycetes" family="Hymenochaetaceae" genus="Tropicoporus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Tropicoporus indicus" order="Hymenochaetales" pageId="0" pageNumber="29" phylum="Basidiomycota" rank="species" species="indicus">
<emphasis id="ACE313B25FC7830F12A4A7AE2591B6F9" italics="true" pageId="0" pageNumber="29">Tropicoporus indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="C4989550866730E10919384A41F807D3" lsidName="T. drechsleri" pageId="0" pageNumber="29" rank="species" species="drechsleri">
<emphasis id="8EC068E2DDC6E6DDD8B9DD5A532F3D83" italics="true" pageId="0" pageNumber="29">T. drechsleri</emphasis>
</taxonomicName>
share concentrically sulcate deeply-cracked pilei with mono-dimitic hyphal system, larger pores (&lt;6/mm) and presence of cystidioles, but the South American species differs in basidiospore size (4-5.5
<normalizedToken id="97247C47E8A131858A3A41F682495144" originalValue="×">x</normalizedToken>
3-4.5
<normalizedToken id="28DD91A828D881E6B15E4154B23252E6" originalValue="μm">μm</normalizedToken>
) (
<bibRefCitation id="804381AA5378D0B8092AA0ABBE7567FD" DOI="https://doi.org/10.11646/phytotaxa.338.1.6" author="Salvador-Montoya, CA" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="29" pagination="075 - 089" refId="B22" refString="Salvador-Montoya, CA, Costa-Rezende, DH, Ferreira-Lopes, V, Borba-Silva, MA, Popoff, OF, 2018. Tropicoporus drechsleri (Hymenochaetales, Basidiomycota), a new species in the &quot; Inonotus linteus &quot; complex from Northern Argentina. Phytotaxa 338: 075 - 089, DOI: https://doi.org/10.11646/phytotaxa.338.1.6" title="Tropicoporus drechsleri (Hymenochaetales, Basidiomycota), a new species in the &quot; Inonotus linteus &quot; complex from Northern Argentina." url="https://doi.org/10.11646/phytotaxa.338.1.6" volume="338" year="2018">Salvador-Montoya et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

113
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<mods:title id="C8F5B13F63A1D74419331569B5A8F369">Review of the genus Tersilochus Holmgren (Hymenoptera, Ichneumonidae, Tersilochinae) from South Korea</mods:title>
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<paragraph id="026A6773F5E6846DB249836E733909C0" pageId="7" pageNumber="34">
<taxonomicName id="BB67401F0190FD8E24AD4FB6E08354B5" LSID="http://zoobank.org/83AB92C8-1729-4ADA-9F15-286F4861DAE5" authority="Khalaim &amp; Lee" class="Insecta" family="Ichneumonidae" genus="Tersilochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tersilochus (Tersilochus) iracundus" order="Hymenoptera" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="iracundus" subGenus="Tersilochus">Tersilochus (Tersilochus) iracundus Khalaim &amp; Lee</taxonomicName>
<taxonomicNameLabel id="8B73E562334A4BBF820EDAD6FAF83068" pageId="7" pageNumber="34">sp. n.</taxonomicNameLabel>
Figs 18-27
</paragraph>
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<paragraph id="39FC2E7081BD520A779D93626E996E10" pageId="7" pageNumber="34">Description.</paragraph>
<paragraph id="D99655AA16AFD9EDC795500A7F5BA942" pageId="7" pageNumber="34">Female (holotype). Body length 3.7 mm. Fore wing length 2.8 mm.</paragraph>
<paragraph id="AA80A4344D532171C255E2A9B552BECD" lastPageId="8" lastPageNumber="35" pageId="7" pageNumber="34">
Head strongly rounded behind eyes in dorsal view (Fig. 19); temple 0.74 times as long as eye width. Inner eye orbits parallel (Fig. 23). Mandible with upper tooth much longer than lower tooth. Clypeus lenticular, almost 3.0 times as broad as long, in profile convex, with lower 0.4 bent backwards (Fig. 23); sparsely punctate, finely granulate, and dull in upper 0.7. Malar space 0.4 times as long as basal width of mandible. Flagellum of antenna weakly tapered towards apex, with 19 segments (Fig. 20); subbasal flagellomeres 1.3-1.4 times, and subapical flagellomeres about 1.2 times
<pageBreakToken id="938592AEE4C935A308C51A30A43234B2" pageId="8" pageNumber="35" start="start">as</pageBreakToken>
long as broad; flagellomeres 4-6 with distinct and flagellomere 7 with rudimental subapical finger-shaped structures on outer surface (Fig. 21). Face, frons, vertex, and temple distinctly granulate, dull, and impunctate (Figs 19, 23).
</paragraph>
<caption id="59784AE2DBCC07ECE01C871032C77BA3" pageId="8" pageNumber="35">
<paragraph id="4F6385D43824896D1FE73BB763E56969" pageId="8" pageNumber="35">
Figures 23-30.
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sp. n., female, holotype: 23 head, frontal view 24 mesosoma, ventrolateral view 25 fore wings 26 metasoma with ovipositor, lateral view 27 tergites 1-3 of metasoma, dorsal view.
<taxonomicName id="6EE7EE21461DD3BBDFEBFDE61A7CB851" class="Insecta" family="Ichneumonidae" genus="Tersilochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tersilochus nigellus" order="Hymenoptera" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="nigellus">Tersilochus nigellus</taxonomicName>
sp. n., female, holotype: 28 head with antenna and anterior part of mesosoma, lateral view 29 mesosoma, lateral view 30 propodeum and base of metasoma, dorsolateral view.
</paragraph>
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<pageBreakToken id="718AE4E7CDDC04C382C5A99838B6A748" pageId="9" pageNumber="36" start="start">Mesosoma</pageBreakToken>
entirely granulate, dull, impunctate; mesopleuron centrally with fine oblique striae on granulate background (Fig. 24). Notaulus absent. Foveate groove weak and short, oblique, situated in anterior half of mesopleuron (Fig. 24). Propodeum with narrow basal area, which is 0.4 times as long as apical area (Fig. 22). Propodeal spiracle separated from pleural carina by 1.5 times diameter of spiracle. Apical area slightly impressed, anteriorly rounded (Fig. 22). Apical longitudinal carinae developed in posterior half, anteriorly absent. Fore wing (Fig. 25) with intercubitus thickened, as long as abscissa of cubitus between intercubitus and second recurrent vein. First abscissa of radius slightly longer than width of pterostigma. Metacarpus ending far from apex of fore wing. Postnervulus intercepted below middle. Hind wing with nervellus vertical. First tergite almost 3.0 times as long as broad posteriorly, mostly smooth, with petiole more or less round in cross-section, well separated from postpetiole in dorsal view, finely striate laterally before glymma (Fig. 27). Glymma deep, situated somewhat behind center of first tergite, joining by distinct furrow to ventral part of postpetiole (Fig. 26). Second tergite as long as anteriorly broad (Fig. 27). Thyridial depression short, transverse. Ovipositor short, slender, almost straight basally and upcurved in apical 0.3, with fine teeth dorsally and ventrally at apex (Fig. 26); sheath 0.7 times as long as first tergite.
</paragraph>
<paragraph id="353BD09C879988D1537A21942D9FBF3F" pageId="9" pageNumber="36">Head, mesosoma, and first tergite black; palpi, mandible (except reddish black teeth), and lower 0.3 of clypeus yellow-brown; tegula yellow. Antenna dark brown. Pterostigma brown. Legs brownish yellow; fore and mid coxae weakly brown, and hind coxa strongly brown. Metasoma behind first tergite yellow-brown.</paragraph>
<paragraph id="C47448BFBCDCDF1BCC9C97742FCF6AF9" pageId="9" pageNumber="36">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="D03B0777FBD4FABDB66D879F58B2CEEB" pageId="9" pageNumber="36" type="comparison">
<paragraph id="CF642D83E07A2D530BEEE52A0A4E75B9" pageId="9" pageNumber="36">Comparison.</paragraph>
<paragraph id="67509D0009D7564D9FF8B97F4C78E912" pageId="9" pageNumber="36">
Differs from other Palaearctic species of
<taxonomicName id="BE9CD147247ACC9076C14FA931CD7B60" class="Insecta" family="Ichneumonidae" genus="Tersilochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tersilochus" order="Hymenoptera" pageId="9" pageNumber="36" phylum="Arthropoda" rank="genus">Tersilochus</taxonomicName>
by the combination short malar space, reddish brown behind first tergite of metasoma (Figs 26, 27) and short ovipositor (Fig. 26).
</paragraph>
</subSubSection>
<subSubSection id="1EC5A0D6BF1F3F79D8E1F829994F1F4F" pageId="9" pageNumber="36" type="type material">
<paragraph id="89C69654DE920C2464C4B25EE1C36A23" pageId="9" pageNumber="36">Type material.</paragraph>
<paragraph id="28A3CDAA85BFC415A391488C63BA1DE6" pageId="9" pageNumber="36">
Holotype female, South Korea, Chungbuk-do (CB), Jecheon-si, Deoksan-myeon, Worak-ri, Deoksanmaepyoso,
<geoCoordinate id="F4CE89986FF4B4032F83582DC3E9FA65" direction="north" orientation="latitude" precision="925" value="36.866665">36°52'N</geoCoordinate>
,
<geoCoordinate id="76CC461690A7A6D940AE4E93AE843352" direction="east" orientation="longitude" precision="925" value="128.21666">128°13'E</geoCoordinate>
, Malaise trap, 6-20.V.2006, coll. J.W. Lee (YUG).
</paragraph>
</subSubSection>
<subSubSection id="571EBABA336EEAE971BF9A4B8AA6C41D" pageId="9" pageNumber="36" type="distribution">
<paragraph id="603FC1573DE4751E398DE4DD4E0567B2" pageId="9" pageNumber="36">Distribution.</paragraph>
<paragraph id="2C025249658ED7CDB718D374BABAF19C" pageId="9" pageNumber="36">South Korea.</paragraph>
</subSubSection>
<subSubSection id="E83AB0B2FE257A79999682F119AB9C60" pageId="9" pageNumber="36" type="etymology">
<paragraph id="E5150D1BAA3804C058E94139EE9908BD" pageId="9" pageNumber="36">Etymology.</paragraph>
<paragraph id="FE58EB145CA2EA49EEAF37422D17AA9F" pageId="9" pageNumber="36">Named after the Latin iracundus (angry, hot-tempered, furious).</paragraph>
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215
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<mods:title id="4AFBC13B21A0B3B590FD8F02E181F684">Zootaxa</mods:title>
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<mods:part id="C1B039E79CF023CC0821499D1C920927">
<mods:date id="C9382594B534BB91B80FAC37F8DFDE4C">2003</mods:date>
<mods:detail id="8F3774DF506C73048BD39DC892BD173D" type="volume">
<mods:number id="2E33000FB9FD63285C294C4A9A4D7958">142</mods:number>
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<mods:identifier id="2833733FE34351E499560DF3FE4A73A2" type="CLB-Dataset">51371</mods:identifier>
<mods:identifier id="856D6A594588CB7E7679C96EA6AA6B01" type="DOI">10.5281/zenodo.156420</mods:identifier>
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<treatment id="62088784C301C741E12E75F2FE9CF9C6" ID-DOI="http://doi.org/10.5281/zenodo.6274664" ID-Zenodo-Dep="6274664" LSID="urn:lsid:plazi:treatment:62088784C301C741E12E75F2FE9CF9C6" httpUri="http://treatment.plazi.org/id/62088784C301C741E12E75F2FE9CF9C6" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48">
<subSubSection id="A2BB6519C301C75FE12E75F2FC12FE35" box="[264,936,483,510]" pageId="47" pageNumber="48" type="key">
<paragraph id="EA1E3692C301C75FE12E75F2FC12FE35" blockId="47.[264,936,483,510]" box="[264,936,483,510]" pageId="47" pageNumber="48">
<heading id="B15681FEC301C75FE12E75F2FC12FE35" bold="true" box="[264,936,483,510]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<emphasis id="D8D5EA80C301C75FE12E75F2FC12FE35" bold="true" box="[264,936,483,510]" pageId="47" pageNumber="48">
<key id="9E3061A3C301C75FE12E75F2FC12FE35" box="[264,936,483,510]" pageId="47" pageNumber="48">
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13.
<taxonomicName id="2DA14D11C301C75FE11275F5FD20FE35" ID-CoL="6YBD9" authority="Gusarov" authorityName="Gusarov" box="[308,666,483,510]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="D8D5EA80C301C75FE11275F5FD92FE36" bold="true" box="[308,552,484,510]" italics="true" pageId="47" pageNumber="48">Seeversiella minima</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC301C75FE28175F2FD53FE35" box="[679,745,483,509]" pageId="47" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C301C75FE2DF75F2FC1AFE35" box="[761,928,483,509]" captionStart-0="FIGURES 175 ­ 178" captionStart-1="FIGURES 179 ­ 189" captionStartId-0="47.[264,383,1629,1653]" captionStartId-1="48.[264,383,1595,1619]" captionTargetBox-0="[300,1285,812,1597]" captionTargetBox-1="[298,1302,460,1563]" captionTargetId-0="figure@47.[294,1294,804,1603]" captionTargetId-1="figure@48.[278,1310,449,1569]" captionTargetPageId-0="47" captionTargetPageId-1="48" captionText-0="FIGURES 175 ­ 178. Abdominal segment 8 of Seeversiella minima Gusarov, sp. n. (175 ­ 176, holotype; 177 ­ 178, paratype, 21.7 km NE Metapan, El Salvador). 175 – male tergum 8; 176 – male sternum 8; 177 – female tergum 8; 178 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri-0="https://zenodo.org/record/156451/files/figure.png" httpUri-1="https://zenodo.org/record/156452/files/figure.png" pageId="47" pageNumber="48">Figs. 175­189</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C301C75FE12E763FFECFFD09" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="EA1E3692C301C75FE12E763FFB8CFDB9" blockId="47.[264,1323,558,705]" pageId="47" pageNumber="48">
<emphasis id="D8D5EA80C301C75FE12E763FFE7AFD80" bold="true" box="[264,448,558,584]" pageId="47" pageNumber="48">
<typeStatus id="351A8830C301C75FE12E763FFEFFFD80" box="[264,325,558,584]" pageId="47" pageNumber="48">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C301C75FE1F7763EFDFBFD81" box="[465,577,559,585]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C301C75FE254763FFC53FD80" bold="true" box="[626,1001,558,584]" pageId="47" pageNumber="48">
<collectingCountry id="92B67602C301C75FE254763FFCFFFD80" box="[626,837,558,584]" name="El Salvador" pageId="47" pageNumber="48">EL SALVADOR</collectingCountry>
: Santa Ana:
</emphasis>
Montecristo,
<quantity id="2D599B77C301C75FE4BF763EFB41FD81" box="[1177,1275,559,585]" metricMagnitude="4" metricUnit="m" metricValue="2.17" pageId="47" pageNumber="48" unit="km" value="21.7">21.7 km</quantity>
NE Metapan,
<quantity id="2D599B77C301C75FE15B7646FE63FDB9" box="[381,473,599,625]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="47" pageNumber="48" unit="m" value="2100.0">2100 m</quantity>
, forest litter (R.S.Anderson),
<date id="9E1F1052C301C75FE3127646FC78FDB9" box="[820,962,599,625]" pageId="47" pageNumber="48" value="1994-08-29">29.viii.1994</date>
(
<collectionCode id="8CB0AE57C301C75FE3F77646FB93FDB9" box="[977,1065,599,625]" pageId="47" pageNumber="48">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C301C75FE11E766EFECFFD09" blockId="47.[264,1323,558,705]" pageId="47" pageNumber="48">
<typeStatus id="351A8830C301C75FE11E766EFE15FD51" box="[312,431,639,665]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C301C75FE1EB766FFCECFD50" bold="true" box="[461,854,638,664]" pageId="47" pageNumber="48">
<collectingCountry id="92B67602C301C75FE1EB766FFD1EFD50" box="[461,676,638,664]" name="El Salvador" pageId="47" pageNumber="48">EL SALVADOR</collectingCountry>
: Santa Ana:
</emphasis>
, 4, same data as the
<typeStatus id="351A8830C301C75FE4E2766EFA91FD51" box="[1220,1323,639,665]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C301C75FE13676B6FED2FD09" box="[272,360,679,705]" pageId="47" pageNumber="48">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC301C75FE12E724CFBEAF972" httpUri="https://zenodo.org/record/156451/files/figure.png" pageId="47" pageNumber="48" targetBox="[300,1285,812,1597]" targetPageId="47">
<paragraph id="EA1E3692C301C75FE12E724CFBEAF972" blockId="47.[264,1322,1629,1722]" pageId="47" pageNumber="48">
<emphasis id="D8D5EA80C301C75FE12E724CFE65F9BD" bold="true" box="[264,479,1629,1653]" pageId="47" pageNumber="48">FIGURES 175­178</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C301C75FE2DA724CFB8FF9BC" authority="Gusarov" authorityName="Gusarov" box="[764,1077,1629,1652]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="D8D5EA80C301C75FE2DA724CFC69F9BC" box="[764,979,1629,1652]" italics="true" pageId="47" pageNumber="48">Seeversiella minima</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC301C75FE466724CFBC6F9BD" box="[1088,1148,1629,1653]" pageId="47" pageNumber="48" rank="species">
<emphasis id="D8D5EA80C301C75FE466724CFBC6F9BD" bold="true" box="[1088,1148,1629,1653]" pageId="47" pageNumber="48">sp. n.</emphasis>
</taxonomicNameLabel>
(175­176, holotype; 177­178, paratype, 21.7 km NE Metapan, El Salvador). 175 – male tergum 8; 176 – male sternum 8; 177 – female tergum 8; 178 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C301C741E11E731BFDBAFDEE" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="EA1E3692C301C740E11E731BFC4BFE4E" blockId="47.[264,1323,1802,1869]" lastBlockId="48.[264,1324,284,390]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="D8D5EA80C301C75FE11E731BFE02F8EC" bold="true" box="[312,440,1802,1828]" pageId="47" pageNumber="48">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C301C75FE1EA731AFD07F8EC" box="[460,701,1803,1828]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="D8D5EA80C301C75FE1EA731AFD07F8EC" box="[460,701,1803,1828]" italics="true" pageId="47" pageNumber="48">Seeversiella minima</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C301C75FE4F5731AFEFDF884" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C301C75FE4F5731AFEFDF884" italics="true" pageId="47" pageNumber="48">Seeversiella</emphasis>
</taxonomicName>
by having brownish yellow; small eyes (temple length to eye length ratio 4.0­7.0); glossy pronotum with fine and weak microsculpture; elytra much shorter than pronotum; reduced wings; tergum 8 with four pairs of macrosetae; the shape of aedeagus (
<figureCitation id="729A2A17C31EC740E4947555FE1CFE4E" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="48" pageNumber="49">Figs. 179­ 183, 185­189</figureCitation>
) and the distinct shape of spermatheca (
<figureCitation id="729A2A17C31EC740E35A757DFC59FE4E" box="[892,995,364,390]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="48" pageNumber="49">Fig. 184</figureCitation>
).
</paragraph>
<caption id="BEDE661AC31EC740E12E722AFECBF892" httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="48" pageNumber="49" targetBox="[298,1302,460,1563]" targetPageId="48">
<paragraph id="EA1E3692C31EC740E12E722AFECBF892" blockId="48.[264,1323,1595,1882]" pageId="48" pageNumber="49">
<emphasis id="D8D5EA80C31EC740E12E722AFE5BF99B" bold="true" box="[264,481,1595,1619]" pageId="48" pageNumber="49">FIGURES 179­189</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C31EC740E2A2722DFC78F99B" authority="Gusarov" authorityName="Gusarov" box="[644,962,1596,1619]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="D8D5EA80C31EC740E2A2722DFCE5F99B" box="[644,863,1596,1619]" italics="true" pageId="48" pageNumber="49">Seeversiella minima</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC31EC740E3F6722AFBB4F99B" box="[976,1038,1595,1619]" pageId="48" pageNumber="49" rank="species">
<emphasis id="D8D5EA80C31EC740E3F6722AFBB4F99B" bold="true" box="[976,1038,1595,1619]" pageId="48" pageNumber="49">sp. n.</emphasis>
</taxonomicNameLabel>
(179­183, 185­186, holotype; 184, 187­189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185­186), 0.1 mm (180, 182­184, 187­189).
</paragraph>
</caption>
<paragraph id="EA1E3692C31FC741E11E750DFDBAFDEE" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
<taxonomicName id="2DA14D11C31FC741E11E750DFD9EFEFD" box="[312,548,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="D8D5EA80C31FC741E11E750DFD9EFEFD" box="[312,548,284,309]" italics="true" pageId="49" pageNumber="50">Seeversiella minima</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C31FC741E378750DFC76FEFD" box="[862,972,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="D8D5EA80C31FC741E378750DFC76FEFD" box="[862,972,284,309]" italics="true" pageId="49" pageNumber="50">S. flavida</emphasis>
</taxonomicName>
and other similar species (24­ 28) with small body and very short elytra by the combination of the following characters: pubescence in lateral portions of pronotum directed towards midline and obliquely posteriorly; tergum 8 with four pairs of macrosetae; proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C31FC741E2F875ADFCFCFE1E" box="[734,838,444,470]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Fig. 183</figureCitation>
); apex of median lobe in parameral view wide (
<figureCitation id="729A2A17C31FC741E17275F5FE45FE36" box="[340,511,484,510]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Figs. 179­180</figureCitation>
); spermatheca J­shaped with bulbous proximal portion and without umbilicus (
<figureCitation id="729A2A17C31FC741E1AA761DFE48FDEE" box="[396,498,524,550]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Fig. 184</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C31FC741E11E7622FDC7FA5E" pageId="49" pageNumber="50" type="description">
<paragraph id="EA1E3692C31FC741E11E7622FE7CFDBE" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
<emphasis id="D8D5EA80C31FC741E11E7622FE6AFD85" bold="true" box="[312,464,563,589]" pageId="49" pageNumber="50">Description.</emphasis>
Length 1.6­2.0 mm. Body brownish­yellow with yellow legs, antennae and mouthparts.
</paragraph>
<paragraph id="EA1E3692C31FC741E11E7695FD35FCDE" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">Head surface glossy, with fine and weak isodiametric microsculpture, fine and sparse punctation, distance between punctures equals 3­4 times their diameter. Temple length to eye length ratio 4.0­7.0. Antennal article 2 as long as 3, article 4 transverse (ratio 1.5), 5­ 10 strongly transverse (ratio 2.0).</paragraph>
<paragraph id="EA1E3692C31FC741E11E7735FE7EFBE6" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C31FC741E40D7735FB6CFCF6" box="[1067,1238,804,830]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="4.0" metricValueMin="3.4" pageId="49" pageNumber="50" unit="mm" value="0.37" valueMax="0.4" valueMin="0.34">0.34­0.40 mm</quantity>
, length
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, width to length ratio 1.1, surface glossy, microsculpture and punctation as on head. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.9 times wider than long, glossy, with fine and poorly visible isodiametric microsculpture; punctation asperate and stronger than on pronotum, distance between punctures equals 2­3 times their diameter. Wings shorter than elytra or absent.
</paragraph>
<paragraph id="EA1E3692C31FC741E11E702DFC1BFB06" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 2­3 times their diameter on terga 3­5 and 3­6 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C31FC741E2C170A5FC2EFB06" box="[743,916,1204,1230]" captionStart="FIGURES 175 ­ 178" captionStartId="47.[264,383,1629,1653]" captionTargetBox="[300,1285,812,1597]" captionTargetId="figure@47.[294,1294,804,1603]" captionTargetPageId="47" captionText="FIGURES 175 ­ 178. Abdominal segment 8 of Seeversiella minima Gusarov, sp. n. (175 ­ 176, holotype; 177 ­ 178, paratype, 21.7 km NE Metapan, El Salvador). 175 – male tergum 8; 176 – male sternum 8; 177 – female tergum 8; 178 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156451/files/figure.png" pageId="49" pageNumber="50">Figs. 175, 177</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C31FC741E11E70CDFBD7FAD6" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
In males posterior angles of tergum 3 not projecting, tergum 7 without medial carina or tubercle, posterior margin of tergum 8 without emargination (
<figureCitation id="729A2A17C31FC741E3DF7115FBDAFAD6" box="[1017,1120,1284,1310]" captionStart="FIGURES 175 ­ 178" captionStartId="47.[264,383,1629,1653]" captionTargetBox="[300,1285,812,1597]" captionTargetId="figure@47.[294,1294,804,1603]" captionTargetPageId="47" captionText="FIGURES 175 ­ 178. Abdominal segment 8 of Seeversiella minima Gusarov, sp. n. (175 ­ 176, holotype; 177 ­ 178, paratype, 21.7 km NE Metapan, El Salvador). 175 – male tergum 8; 176 – male sternum 8; 177 – female tergum 8; 178 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156451/files/figure.png" pageId="49" pageNumber="50">Fig. 175</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C31FC741E11E713DFCF6FAA6" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
Aedeagus as in
<figureCitation id="729A2A17C31FC741E1DB713DFCA6FA8E" box="[509,796,1324,1350]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Figs. 179­183, 185­189</figureCitation>
. Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C31FC741E2FE7145FC85FAA6" box="[728,831,1364,1390]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Fig. 183</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C31FC741E11E716DFDC7FA5E" blockId="49.[264,1324,284,1550]" box="[312,637,1404,1430]" pageId="49" pageNumber="50">
Spermatheca as in
<figureCitation id="729A2A17C31FC741E232716DFDC3FA5E" box="[532,633,1404,1430]" captionStart="FIGURES 179 ­ 189" captionStartId="48.[264,383,1595,1619]" captionTargetBox="[298,1302,460,1563]" captionTargetId="figure@48.[278,1310,449,1569]" captionTargetPageId="48" captionText="FIGURES 179 ­ 189. Genitalia of Seeversiella minima Gusarov, sp. n. (179 ­ 183, 185 ­ 186, holotype; 184, 187 ­ 189, paratypes, 21.7 km NE Metapan, El Salvador). 179 – median lobe, parameral view; 180 – apex of median lobe, parameral view; 181 – median lobe, lateral view; 182 – apex of median lobe, lateral view; 183 – apex of left paramere; 184 – spermatheca; 185 – details of internal sac retracted into median lobe, lateral view; 186 – details of internal sac retracted into median lobe, abparameral view; 187 – medial lamellae, parameral view; 188 – medial lamellae, lateral view; 189 – copulatory piece, lateral view. Scale bar 0.2 mm (179, 181, 185 ­ 186), 0.1 mm (180, 182 ­ 184, 187 ­ 189)." httpUri="https://zenodo.org/record/156452/files/figure.png" pageId="49" pageNumber="50">Fig. 184</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C31FC741E11E71B2FE9CF9C6" pageId="49" pageNumber="50" type="distribution">
<paragraph id="EA1E3692C31FC741E11E71B2FBC2FA76" blockId="49.[264,1324,284,1550]" box="[312,1144,1443,1470]" pageId="49" pageNumber="50">
<emphasis id="D8D5EA80C31FC741E11E71B2FE62FA75" bold="true" box="[312,472,1443,1469]" pageId="49" pageNumber="50">Distribution.</emphasis>
Known from Cerro Montecristo,
<collectingCountry id="92B67602C31FC741E34D71B5FC4EFA76" box="[875,1012,1444,1470]" name="El Salvador" pageId="49" pageNumber="50">El Salvador</collectingCountry>
(
<figureCitation id="729A2A17C31FC741E42271B5FBD1FA76" box="[1028,1131,1444,1470]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="49" pageNumber="50">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C31FC741E11E71DAFE9CF9C6" blockId="49.[264,1324,284,1550]" pageId="49" pageNumber="50">
<emphasis id="D8D5EA80C31FC741E11E71DAFDBFFA2D" bold="true" box="[312,517,1483,1509]" pageId="49" pageNumber="50">Natural History.</emphasis>
<taxonomicName id="2DA14D11C31FC741E23571DDFCBAFA2D" box="[531,768,1484,1509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="D8D5EA80C31FC741E23571DDFCBAFA2D" box="[531,768,1484,1509]" italics="true" pageId="49" pageNumber="50">Seeversiella minima</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C31FC741E4C971DDFE98F9C6" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="49" pageNumber="50" unit="m" value="2100.0">2100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

247
data/62/08/87/62088784C302C75FE12E775AFED3FE4E.xml Normal file
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@ -0,0 +1,247 @@
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12.
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<emphasis id="D8D5EA80C302C75CE112775DFDA3FCAE" bold="true" box="[308,537,844,870]" italics="true" pageId="44" pageNumber="45">Seeversiella similis</emphasis>
Gusarov
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,
<taxonomicNameLabel id="C3E657FBC302C75CE2BE775AFD60FCAD" box="[664,730,843,869]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C302C75CE2CC775AFC2BFCAD" box="[746,913,843,869]" captionStart-0="FIGURES 161 ­ 164" captionStart-1="FIGURES 165 ­ 168" captionStart-2="FIGURES 169 ­ 174" captionStartId-0="44.[264,383,1760,1784]" captionStartId-1="45.[264,383,1583,1607]" captionStartId-2="46.[264,383,1661,1685]" captionTargetBox-0="[317,1270,975,1727]" captionTargetBox-1="[302,1289,779,1549]" captionTargetBox-2="[288,1298,900,1628]" captionTargetId-0="figure@44.[309,1279,964,1734]" captionTargetId-1="figure@45.[292,1295,769,1556]" captionTargetId-2="figure@46.[282,1304,889,1634]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionText-0="FIGURES 161 ­ 164. Abdominal segment 8 of Seeversiella similis Gusarov, sp. n. (161 ­ 162, holotype; 163 ­ 164, paratype from La Tigra, Honduras). 161 – male tergum 8; 162 – male sternum 8; 163 – female tergum 8; 164 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 – median lobe, parameral view; 166 – apex of median lobe, parameral view; 167 – median lobe, lateral view; 168 – apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." captionText-2="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri-0="https://zenodo.org/record/156448/files/figure.png" httpUri-1="https://zenodo.org/record/156449/files/figure.png" httpUri-2="https://zenodo.org/record/156450/files/figure.png" pageId="44" pageNumber="45">Figs. 161­174</figureCitation>
)
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<emphasis id="D8D5EA80C302C75CE12E72F1FE65F930" bold="true" box="[264,479,1760,1784]" pageId="44" pageNumber="45">FIGURES 161­164</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C302C75CE32772F0FB89F930" authority="Gusarov" authorityName="Gusarov" box="[769,1075,1761,1784]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="D8D5EA80C302C75CE32772F0FC75F930" box="[769,975,1761,1784]" italics="true" pageId="44" pageNumber="45">Seeversiella similis</emphasis>
Gusarov
</taxonomicName>
,
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</emphasis>
(161­162, holotype; 163­164, paratype from La Tigra, Honduras). 161 – male tergum 8; 162 – male sternum 8; 163 – female tergum 8; 164 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C303C75DE12E750AFD85FE36" pageId="45" pageNumber="46" type="materials_examined">
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<typeStatus id="351A8830C303C75DE12E750AFEF9FEFD" box="[264,323,283,309]" pageId="45" pageNumber="46">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C303C75DE1FC750DFDF0FEFE" box="[474,586,284,310]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C303C75DE2A7750AFBF6FEFD" bold="true" box="[641,1100,283,309]" pageId="45" pageNumber="46">
<collectingCountry id="92B67602C303C75DE2A7750AFC97FEFD" box="[641,813,283,309]" name="Honduras" pageId="45" pageNumber="46">HONDURAS</collectingCountry>
: Francisco Morazán:
</emphasis>
<quantity id="2D599B77C303C75DE442750DFB72FEFE" box="[1124,1224,284,310]" metricMagnitude="4" metricUnit="m" metricValue="2.13" pageId="45" pageNumber="46" unit="km" value="21.3">21.3 km</quantity>
N Teguicigalpa, La Tigra,
<quantity id="2D599B77C303C75DE2237555FDDCFE96" box="[517,614,324,350]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="45" pageNumber="46" unit="m" value="2100.0">2100 m</quantity>
,
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<geoCoordinate id="8F955055C303C75DE2C37555FCF6FE96" box="[741,844,324,350]" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-86.1">86°06'W</geoCoordinate>
, forest litter (R.S.Anderson),
<date id="9E1F1052C303C75DE49B7555FA96FE96" box="[1213,1324,324,350]" pageId="45" pageNumber="46" value="1994-06-07">7.vi.1994</date>
(
<collectionCode id="8CB0AE57C303C75DE136757DFED2FE4E" box="[272,360,364,390]" pageId="45" pageNumber="46">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C303C75DE11E7585FD85FE36" blockId="45.[264,1324,283,710]" pageId="45" pageNumber="46">
<typeStatus id="351A8830C303C75DE11E7585FE15FE66" box="[312,431,404,430]" pageId="45" pageNumber="46" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C303C75DE1E17582FC37FE65" bold="true" box="[455,909,403,429]" pageId="45" pageNumber="46">
<collectingCountry id="92B67602C303C75DE1E17582FDCEFE65" box="[455,628,403,429]" name="Honduras" pageId="45" pageNumber="46">HONDURAS</collectingCountry>
: Francisco Morazán:
</emphasis>
4, same data as the
<typeStatus id="351A8830C303C75DE49A7585FA9CFE66" box="[1212,1318,404,430]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
; 2, ditto but
<date id="9E1F1052C303C75DE1EA75ADFDE7FE1E" box="[460,605,444,470]" pageId="45" pageNumber="46" value="1994-08-15">15.viii.1994</date>
; 2, 4, ditto but
<quantity id="2D599B77C303C75DE35875ADFC5CFE1E" box="[894,998,444,470]" metricMagnitude="4" metricUnit="m" metricValue="2.2199999999999998" pageId="45" pageNumber="46" unit="km" value="22.2">22.2 km</quantity>
N Teguicigalpa,
<quantity id="2D599B77C303C75DE4E275ADFA9DFE1E" box="[1220,1319,444,470]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="45" pageNumber="46" unit="m" value="2030.0">2030 m</quantity>
,
<date id="9E1F1052C303C75DE12E75F5FE2CFE36" box="[264,406,484,510]" pageId="45" pageNumber="46" value="1994-08-15">15.viii.1994</date>
(all ­
<collectionCode id="8CB0AE57C303C75DE1FB75F5FD8EFE36" box="[477,564,484,510]" pageId="45" pageNumber="46">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C303C75DE11E761AFB7FF8E9" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="EA1E3692C303C75DE11E761AFB6CFD0E" blockId="45.[264,1324,283,710]" pageId="45" pageNumber="46">
<emphasis id="D8D5EA80C303C75DE11E761AFE02FDED" bold="true" box="[312,440,523,549]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C303C75DE1E3761DFD1CFDED" box="[453,678,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C303C75DE1E3761DFD1CFDED" box="[453,678,524,549]" italics="true" pageId="45" pageNumber="46">Seeversiella similis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C303C75DE4BB761DFA91FDED" box="[1181,1323,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C303C75DE4BB761DFA91FDED" box="[1181,1323,524,549]" italics="true" pageId="45" pageNumber="46">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.4­1.8 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra as long or slightly shorter than pronotum (elytral length to pronotal length ratio 1.0­0.9); reduced wings; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C303C75DE2FE76BDFCC7FD0E" box="[728,893,684,710]" captionStart-0="FIGURES 165 ­ 168" captionStart-1="FIGURES 169 ­ 174" captionStartId-0="45.[264,383,1583,1607]" captionStartId-1="46.[264,383,1661,1685]" captionTargetBox-0="[302,1289,779,1549]" captionTargetBox-1="[288,1298,900,1628]" captionTargetId-0="figure@45.[292,1295,769,1556]" captionTargetId-1="figure@46.[282,1304,889,1634]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 – median lobe, parameral view; 166 – apex of median lobe, parameral view; 167 – median lobe, lateral view; 168 – apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." captionText-1="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri-0="https://zenodo.org/record/156449/files/figure.png" httpUri-1="https://zenodo.org/record/156450/files/figure.png" pageId="45" pageNumber="46">Figs. 165­173</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C303C75DE44476BDFB72FD0E" box="[1122,1224,684,710]" captionStart="FIGURES 169 ­ 174" captionStartId="46.[264,383,1661,1685]" captionTargetBox="[288,1298,900,1628]" captionTargetId="figure@46.[282,1304,889,1634]" captionTargetPageId="46" captionText="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="45" pageNumber="46">Fig. 174</figureCitation>
).
</paragraph>
<caption id="BEDE661AC303C75DE12E723EFB71F943" httpUri="https://zenodo.org/record/156449/files/figure.png" pageId="45" pageNumber="46" targetBox="[302,1289,779,1549]" targetPageId="45">
<paragraph id="EA1E3692C303C75DE12E723EFB71F943" blockId="45.[264,1322,1583,1675]" pageId="45" pageNumber="46">
<emphasis id="D8D5EA80C303C75DE12E723EFE5AF98F" bold="true" box="[264,480,1583,1607]" pageId="45" pageNumber="46">FIGURES 165­168</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C303C75DE2AE723EFC01F98E" authority="Gusarov" authorityName="Gusarov" box="[648,955,1583,1606]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="D8D5EA80C303C75DE2AE723EFCE2F98E" box="[648,856,1583,1606]" italics="true" pageId="45" pageNumber="46">Seeversiella similis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C303C75DE3EE723EFBBCF98F" bold="true" box="[968,1030,1583,1607]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="C3E657FBC303C75DE3EE723EFBBCF98F" box="[968,1030,1583,1607]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 165 – median lobe, parameral view; 166 – apex of median lobe, parameral view; 167 – median lobe, lateral view; 168 – apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168).
</paragraph>
</caption>
<paragraph id="EA1E3692C303C75DE11E72CEFB7FF8E9" blockId="45.[264,1323,1759,1905]" pageId="45" pageNumber="46">
<taxonomicName id="2DA14D11C303C75DE11E72CEFDA0F930" box="[312,538,1759,1784]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C303C75DE11E72CEFDA0F930" box="[312,538,1759,1784]" italics="true" pageId="45" pageNumber="46">Seeversiella similis</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C303C75DE34372CEFC7CF930" box="[869,966,1759,1784]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C303C75DE34372CEFC7CF930" box="[869,966,1759,1784]" italics="true" pageId="45" pageNumber="46">S. badia</emphasis>
</taxonomicName>
in that apex of median lobe in lateral view lacks clear emargination on parameral side (
<figureCitation id="729A2A17C303C75DE3B97316FBFCF8E9" box="[927,1094,1799,1825]" captionStart="FIGURES 165 ­ 168" captionStartId="45.[264,383,1583,1607]" captionTargetBox="[302,1289,779,1549]" captionTargetId="figure@45.[292,1295,769,1556]" captionTargetPageId="45" captionText="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 – median lobe, parameral view; 166 – apex of median lobe, parameral view; 167 – median lobe, lateral view; 168 – apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." httpUri="https://zenodo.org/record/156449/files/figure.png" pageId="45" pageNumber="46">Figs. 167­168</figureCitation>
; 156­157).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C303C75EE11E733EFC0CF889" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph id="EA1E3692C303C75EE11E733EFCDCFE66" blockId="45.[264,1323,1759,1905]" lastBlockId="46.[264,1324,284,790]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<taxonomicName id="2DA14D11C303C75DE11E733EFDA0F880" box="[312,538,1839,1864]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C303C75DE11E733EFDA0F880" box="[312,538,1839,1864]" italics="true" pageId="45" pageNumber="46">Seeversiella similis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C303C75DE37A733EFC4CF880" box="[860,1014,1839,1864]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C303C75DE37A733EFC4CF880" box="[860,1014,1839,1864]" italics="true" pageId="45" pageNumber="46">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 5­8, 10) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have small spicules (
<figureCitation id="729A2A17C300C75EE2DD7555FCDFFE96" box="[763,869,324,350]" captionStart="FIGURES 169 ­ 174" captionStartId="46.[264,383,1661,1685]" captionTargetBox="[288,1298,900,1628]" captionTargetId="figure@46.[282,1304,889,1634]" captionTargetPageId="46" captionText="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="46" pageNumber="47">Fig. 173</figureCitation>
); apex of median lobe in lateral view widened subapically and not emarginate on parameral side (
<figureCitation id="729A2A17C300C75EE3F1757DFB3BFE4E" box="[983,1153,364,390]" captionStart="FIGURES 165 ­ 168" captionStartId="45.[264,383,1583,1607]" captionTargetBox="[302,1289,779,1549]" captionTargetId="figure@45.[292,1295,769,1556]" captionTargetPageId="45" captionText="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 – median lobe, parameral view; 166 – apex of median lobe, parameral view; 167 – median lobe, lateral view; 168 – apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." httpUri="https://zenodo.org/record/156449/files/figure.png" pageId="46" pageNumber="47">Figs. 167­168</figureCitation>
); spermatheca S­shaped with umbilicus facing laterally (
<figureCitation id="729A2A17C300C75EE2D47585FCE3FE66" box="[754,857,404,430]" captionStart="FIGURES 169 ­ 174" captionStartId="46.[264,383,1661,1685]" captionTargetBox="[288,1298,900,1628]" captionTargetId="figure@46.[282,1304,889,1634]" captionTargetPageId="46" captionText="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="46" pageNumber="47">Fig. 174</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C300C75EE11E75AAFC17FE36" blockId="46.[264,1324,284,790]" pageId="46" pageNumber="47">
<emphasis id="D8D5EA80C300C75EE11E75AAFE6AFE1D" bold="true" box="[312,464,443,469]" pageId="46" pageNumber="47">Description.</emphasis>
Length
<quantity id="2D599B77C300C75EE21F75ADFD7FFE1E" box="[569,709,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="46" pageNumber="47" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C300C75EE3A575ADFB96FE1E" box="[899,1068,444,470]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="46" pageNumber="47" unit="mm" value="0.51" valueMax="0.54" valueMin="0.48">0.48­0.54 mm</quantity>
. Body brown to dark brown with lighter elytra, legs brown to yellowish brown.
</paragraph>
<paragraph id="EA1E3692C300C75EE11E761DFC52FD86" blockId="46.[264,1324,284,790]" pageId="46" pageNumber="47">
In all character states
<taxonomicName id="2DA14D11C300C75EE211761DFD1BFDED" box="[567,673,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C300C75EE211761DFD1BFDED" box="[567,673,524,549]" italics="true" pageId="46" pageNumber="47">S. similis</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C300C75EE350761DFBB5FDED" box="[886,1039,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C300C75EE350761DFBB5FDED" box="[886,1039,524,549]" italics="true" pageId="46" pageNumber="47">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C300C75EE11E764DFDE1FD0E" blockId="46.[264,1324,284,790]" pageId="46" pageNumber="47">In males posterior angles of tergum 3 projecting as spines, up to 1 time as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to 2/3 as long as tergum length.</paragraph>
<paragraph id="EA1E3692C300C75EE11E76C5FD21FD26" blockId="46.[264,1324,284,790]" box="[312,667,724,750]" pageId="46" pageNumber="47">
Aedeagus as in
<figureCitation id="729A2A17C300C75EE1D776C5FD2DFD26" box="[497,663,724,750]" captionStart-0="FIGURES 165 ­ 168" captionStart-1="FIGURES 169 ­ 174" captionStartId-0="45.[264,383,1583,1607]" captionStartId-1="46.[264,383,1661,1685]" captionTargetBox-0="[302,1289,779,1549]" captionTargetBox-1="[288,1298,900,1628]" captionTargetId-0="figure@45.[292,1295,769,1556]" captionTargetId-1="figure@46.[282,1304,889,1634]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 – median lobe, parameral view; 166 – apex of median lobe, parameral view; 167 – median lobe, lateral view; 168 – apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." captionText-1="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri-0="https://zenodo.org/record/156449/files/figure.png" httpUri-1="https://zenodo.org/record/156450/files/figure.png" pageId="46" pageNumber="47">Figs. 165­173</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C300C75EE11E76EDFDC4FCDE" blockId="46.[264,1324,284,790]" box="[312,638,764,790]" pageId="46" pageNumber="47">
Spermatheca as in
<figureCitation id="729A2A17C300C75EE23376EDFDC0FCDE" box="[533,634,764,790]" captionStart="FIGURES 169 ­ 174" captionStartId="46.[264,383,1661,1685]" captionTargetBox="[288,1298,900,1628]" captionTargetId="figure@46.[282,1304,889,1634]" captionTargetPageId="46" captionText="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="46" pageNumber="47">Fig. 174</figureCitation>
.
</paragraph>
<caption id="BEDE661AC300C75EE12E726CFCCCF8D7" httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="46" pageNumber="47" targetBox="[288,1298,900,1628]" targetPageId="46">
<paragraph id="EA1E3692C300C75EE12E726CFCCCF8D7" blockId="46.[264,1323,1661,1857]" pageId="46" pageNumber="47">
<emphasis id="D8D5EA80C300C75EE12E726CFE64F95D" bold="true" box="[264,478,1661,1685]" pageId="46" pageNumber="47">FIGURES 169­174</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C300C75EE253726FFC19F95C" authority="Gusarov" authorityName="Gusarov" box="[629,931,1661,1685]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="D8D5EA80C300C75EE253726FFCFBF95D" box="[629,833,1662,1685]" italics="true" pageId="46" pageNumber="47">Seeversiella similis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C300C75EE38B726CFC52F95D" bold="true" box="[941,1000,1661,1685]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="C3E657FBC300C75EE38B726CFC52F95D" box="[941,1000,1661,1685]" pageId="46" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(171, 173, holotype; 169­170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172­173), 0.1 mm (170­171, 174).
</paragraph>
</caption>
<paragraph id="EA1E3692C300C75EE12E733BFC0CF889" blockId="46.[264,1323,1661,1857]" box="[264,950,1834,1857]" pageId="46" pageNumber="47">CP – copulatory piece; LD – lateral diverticulum of internal sac.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C301C75FE11E750AFED3FE4E" pageId="47" pageNumber="48" type="distribution">
<paragraph id="EA1E3692C301C75FE11E750AFED3FE4E" blockId="47.[264,1323,283,390]" pageId="47" pageNumber="48">
<emphasis id="D8D5EA80C301C75FE11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="47" pageNumber="48">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C301C75FE2A3750DFD4DFEFE" box="[645,759,284,310]" name="Honduras" pageId="47" pageNumber="48">Honduras</collectingCountry>
(
<figureCitation id="729A2A17C301C75FE321750DFCD5FEFE" box="[775,879,284,310]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="47" pageNumber="48">Fig. 387</figureCitation>
).
<emphasis id="D8D5EA80C301C75FE11E7552FDBFFE95" bold="true" box="[312,517,323,349]" pageId="47" pageNumber="48">Natural History.</emphasis>
<taxonomicName id="2DA14D11C301C75FE2327555FD4CFE95" box="[532,758,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C301C75FE2327555FD4CFE95" box="[532,758,324,349]" italics="true" pageId="47" pageNumber="48">Seeversiella similis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C301C75FE4C07555FEDEFE4E" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="47" pageNumber="48" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">2000­ 2100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

223
data/62/08/87/62088784C304C75CE12E750AFED2FD26.xml Normal file
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@ -0,0 +1,223 @@
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<mods:namePart id="9049AD10A8FF8B62AFD09E71DA11E528">Gusarov, Vladimir I.</mods:namePart>
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<mods:date id="240B8EA845699741D84305E6755252D7">2003</mods:date>
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<mods:identifier id="78E08ACE7070B3986359D6BE42AF3908" type="DOI">10.5281/zenodo.156420</mods:identifier>
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<subSubSection id="A2BB6519C304C75AE12E750AFC36FEFD" box="[264,908,283,310]" pageId="42" pageNumber="43" type="key">
<paragraph id="EA1E3692C304C75AE12E750AFC36FEFD" blockId="42.[264,908,283,310]" box="[264,908,283,310]" pageId="42" pageNumber="43">
<heading id="B15681FEC304C75AE12E750AFC36FEFD" bold="true" box="[264,908,283,310]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<emphasis id="D8D5EA80C304C75AE12E750AFC36FEFD" bold="true" box="[264,908,283,310]" pageId="42" pageNumber="43">
<key id="9E3061A3C304C75AE12E750AFC36FEFD" box="[264,908,283,310]" pageId="42" pageNumber="43">
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<keyLead id="515095A7C304C75AE12E750AFC36FEFD" box="[264,908,283,310]" pageId="42" pageNumber="43">
11.
<taxonomicName id="2DA14D11C304C75AE115750DFDC4FEFD" ID-CoL="6YBCN" authority="Gusarov" authorityName="Gusarov" box="[307,638,283,310]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="badia" status="sp. nov.">
<emphasis id="D8D5EA80C304C75AE115750DFDB6FEFE" bold="true" box="[307,524,284,310]" italics="true" pageId="42" pageNumber="43">Seeversiella badia</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC304C75AE2AD750AFD74FEFD" box="[651,718,283,309]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C304C75AE2FB750AFC3EFEFD" box="[733,900,283,309]" captionStart-0="FIGURES 150 ­ 153" captionStart-1="FIGURES 154 ­ 160" captionStartId-0="42.[264,382,1453,1477]" captionStartId-1="43.[264,383,1715,1739]" captionTargetBox-0="[286,1300,613,1419]" captionTargetBox-1="[292,1295,611,1684]" captionTargetId-0="figure@42.[279,1308,604,1426]" captionTargetId-1="figure@43.[281,1307,601,1691]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionText-0="FIGURES 150 ­ 153. Abdominal segment 8 of Seeversiella badia Gusarov, sp. n. (150 ­ 151, holotype; 152 ­ 153, paratype, 21.7 km NE Metapan, El Salvador). 150 – male tergum 8; 151 – male sternum 8; 152 – female tergum 8; 153 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri-0="https://zenodo.org/record/156446/files/figure.png" httpUri-1="https://zenodo.org/record/156447/files/figure.png" pageId="42" pageNumber="43">Figs. 150­160</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C304C75AE12E7577FED1FE31" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="EA1E3692C304C75AE12E7577FBC7FE61" blockId="42.[264,1323,358,505]" pageId="42" pageNumber="43">
<emphasis id="D8D5EA80C304C75AE12E7577FE05FE48" bold="true" box="[264,447,358,384]" pageId="42" pageNumber="43">
<typeStatus id="351A8830C304C75AE12E7577FEFFFE48" box="[264,325,358,384]" pageId="42" pageNumber="43">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C304C75AE1F77576FDFBFE49" box="[465,577,359,385]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C304C75AE2547577FC53FE48" bold="true" box="[626,1001,358,384]" pageId="42" pageNumber="43">
<collectingCountry id="92B67602C304C75AE2547577FCFFFE48" box="[626,837,358,384]" name="El Salvador" pageId="42" pageNumber="43">EL SALVADOR</collectingCountry>
: Santa Ana:
</emphasis>
Montecristo,
<quantity id="2D599B77C304C75AE4BF7576FB41FE49" box="[1177,1275,359,385]" metricMagnitude="4" metricUnit="m" metricValue="2.17" pageId="42" pageNumber="43" unit="km" value="21.7">21.7 km</quantity>
NE Metapan,
<quantity id="2D599B77C304C75AE15B759EFE63FE61" box="[381,473,399,425]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="42" pageNumber="43" unit="m" value="2100.0">2100 m</quantity>
, cloud forest litter (R.S.Anderson),
<date id="9E1F1052C304C75AE35D759EFBB2FE61" box="[891,1032,399,425]" pageId="42" pageNumber="43" value="1994-08-29">29.viii.1994</date>
(
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).
</paragraph>
<paragraph id="EA1E3692C304C75AE11E75A6FED1FE31" blockId="42.[264,1323,358,505]" pageId="42" pageNumber="43">
<typeStatus id="351A8830C304C75AE11E75A6FE15FE19" box="[312,431,439,465]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C304C75AE1EE75A7FCF2FE18" bold="true" box="[456,840,438,464]" pageId="42" pageNumber="43">
<collectingCountry id="92B67602C304C75AE1EE75A7FD27FE18" box="[456,669,438,464]" name="El Salvador" pageId="42" pageNumber="43">EL SALVADOR</collectingCountry>
: Santa Ana:
</emphasis>
2, same data as the
<typeStatus id="351A8830C304C75AE45B75A6FB5EFE19" box="[1149,1252,439,465]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
(all ­
<collectionCode id="8CB0AE57C304C75AE12E75CEFEE4FE31" box="[264,350,479,505]" pageId="42" pageNumber="43">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC304C75AE12E71BCFBEAF9C1" httpUri="https://zenodo.org/record/156446/files/figure.png" pageId="42" pageNumber="43" targetBox="[286,1300,613,1419]" targetPageId="42">
<paragraph id="EA1E3692C304C75AE12E71BCFBEAF9C1" blockId="42.[264,1322,1453,1545]" pageId="42" pageNumber="43">
<emphasis id="D8D5EA80C304C75AE12E71BCFE65FA0D" bold="true" box="[264,479,1453,1477]" pageId="42" pageNumber="43">FIGURES 150­153</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C304C75AE32F71BCFB88FA0C" authority="Gusarov" authorityName="Gusarov" box="[777,1074,1453,1476]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="badia" status="sp. nov.">
<emphasis id="D8D5EA80C304C75AE32F71BCFC75FA0C" box="[777,975,1453,1476]" italics="true" pageId="42" pageNumber="43">Seeversiella badia</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C304C75AE41B71BCFBC0FA0D" bold="true" box="[1085,1146,1453,1477]" pageId="42" pageNumber="43">
<taxonomicNameLabel id="C3E657FBC304C75AE41B71BCFBC0FA0D" box="[1085,1146,1453,1477]" pageId="42" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(150­151, holotype; 152­153, paratype, 21.7 km NE Metapan, El Salvador). 150 – male tergum 8; 151 – male sternum 8; 152 – female tergum 8; 153 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C304C75AE11E724EFB26F88A" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="EA1E3692C304C75AE11E724EFB6CF93A" blockId="42.[264,1324,1631,1858]" pageId="42" pageNumber="43">
<emphasis id="D8D5EA80C304C75AE11E724EFE02F9B1" bold="true" box="[312,440,1631,1657]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C304C75AE1E17271FD25F9B1" box="[455,671,1632,1657]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C304C75AE1E17271FD25F9B1" box="[455,671,1632,1657]" italics="true" pageId="42" pageNumber="43">Seeversiella badia</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C304C75AE4BB7271FA91F9B1" box="[1181,1323,1632,1657]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C304C75AE4BB7271FA91F9B1" box="[1181,1323,1632,1657]" italics="true" pageId="42" pageNumber="43">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.7­2.0 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra as long as pronotum; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C304C75AE2FE72C9FCC7F93A" box="[728,893,1752,1778]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="42" pageNumber="43">Figs. 154­159</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C304C75AE44472C9FB72F93A" box="[1122,1224,1752,1778]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="42" pageNumber="43">Fig. 160</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C304C75AE11E7311FB26F88A" blockId="42.[264,1324,1631,1858]" pageId="42" pageNumber="43">
<taxonomicName id="2DA14D11C304C75AE11E7311FDA9F8D1" box="[312,531,1792,1817]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C304C75AE11E7311FDA9F8D1" box="[312,531,1792,1817]" italics="true" pageId="42" pageNumber="43">Seeversiella badia</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C304C75AE3567311FC67F8D1" box="[880,989,1792,1817]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="D8D5EA80C304C75AE3567311FC67F8D1" box="[880,989,1792,1817]" italics="true" pageId="42" pageNumber="43">S. similis</emphasis>
</taxonomicName>
in that the apex of median lobe in lateral view is emarginate on parameral side (
<figureCitation id="729A2A17C304C75AE3537339FBA7F88A" box="[885,1053,1832,1858]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="42" pageNumber="43">Figs. 156­157</figureCitation>
; 167­168).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C305C75CE11E750DFDC4FDBE" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
<paragraph id="EA1E3692C305C75BE11E750DFCF6FE36" blockId="43.[264,1324,284,510]" pageId="43" pageNumber="44">
<taxonomicName id="2DA14D11C305C75BE11E750DFDABFEFD" box="[312,529,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C305C75BE11E750DFDABFEFD" box="[312,529,284,309]" italics="true" pageId="43" pageNumber="44">Seeversiella badia</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C305C75BE371750DFC48FEFD" box="[855,1010,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C305C75BE371750DFC48FEFD" box="[855,1010,284,309]" italics="true" pageId="43" pageNumber="44">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 5­8, 10) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have small spicules (
<figureCitation id="729A2A17C305C75BE2DD7585FCDFFE66" box="[763,869,404,430]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="43" pageNumber="44">Fig. 158</figureCitation>
); apex of median lobe in lateral view widened subapically and emarginate on parameral side (
<figureCitation id="729A2A17C305C75BE39775ADFBE1FE1E" box="[945,1115,444,470]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="43" pageNumber="44">Figs. 156­157</figureCitation>
); spermatheca Sshaped with umbilicus facing laterally (
<figureCitation id="729A2A17C305C75BE2FE75F5FC84FE36" box="[728,830,484,510]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="43" pageNumber="44">Fig. 160</figureCitation>
).
</paragraph>
<caption id="BEDE661AC305C75BE12E72A2FB46F89D" httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="43" pageNumber="44" targetBox="[292,1295,611,1684]" targetPageId="43">
<paragraph id="EA1E3692C305C75BE12E72A2FB46F89D" blockId="43.[264,1324,1715,1877]" pageId="43" pageNumber="44">
<emphasis id="D8D5EA80C305C75BE12E72A2FE59F903" bold="true" box="[264,483,1715,1739]" pageId="43" pageNumber="44">FIGURES 154­160</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C305C75BE2A872A5FC07F902" authority="Gusarov" authorityName="Gusarov" box="[654,957,1715,1739]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="badia" status="sp. nov.">
<emphasis id="D8D5EA80C305C75BE2A872A5FCE2F903" box="[654,856,1716,1739]" italics="true" pageId="43" pageNumber="44">Seeversiella badia</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C305C75BE3EB72A2FBB7F903" bold="true" box="[973,1037,1715,1739]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="C3E657FBC305C75BE3EB72A2FBB7F903" box="[973,1037,1715,1739]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(154­159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159­160).
</paragraph>
</caption>
<paragraph id="EA1E3692C302C75CE11E750AFC17FE96" blockId="44.[264,1323,283,750]" pageId="44" pageNumber="45">
<emphasis id="D8D5EA80C302C75CE11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="44" pageNumber="45">Description.</emphasis>
Length
<quantity id="2D599B77C302C75CE262750DFD1FFEFE" box="[580,677,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="44" pageNumber="45" unit="mm" value="2.5">2.5 mm</quantity>
, pronotal width
<quantity id="2D599B77C302C75CE348750DFBA1FEFE" box="[878,1051,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="44" pageNumber="45" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50­0.53 mm</quantity>
. Body brown to dark brown with lighter elytra, legs brown to yellowish brown.
</paragraph>
<paragraph id="EA1E3692C302C75CE11E757DFC52FE66" blockId="44.[264,1323,283,750]" pageId="44" pageNumber="45">
In all character states
<taxonomicName id="2DA14D11C302C75CE21C757DFD21FE4D" box="[570,667,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C302C75CE21C757DFD21FE4D" box="[570,667,364,389]" italics="true" pageId="44" pageNumber="45">S. badia</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C302C75CE355757DFBB6FE4D" box="[883,1036,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C302C75CE355757DFBB6FE4D" box="[883,1036,364,389]" italics="true" pageId="44" pageNumber="45">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C302C75CE11E75ADFCE8FDEE" blockId="44.[264,1323,283,750]" pageId="44" pageNumber="45">In the only known male specimen the posterior angles of tergum 3 projecting as spines, as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to 2/3 as long as tergum length.</paragraph>
<paragraph id="EA1E3692C302C75CE11E7625FD26FD86" blockId="44.[264,1323,283,750]" box="[312,668,564,590]" pageId="44" pageNumber="45">
Aedeagus as in
<figureCitation id="729A2A17C302C75CE1D77625FD2DFD86" box="[497,663,564,590]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="44" pageNumber="45">Figs. 154­159</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C302C75CE11E764DFDC4FDBE" blockId="44.[264,1323,283,750]" box="[312,638,604,630]" pageId="44" pageNumber="45">
Spermatheca as in
<figureCitation id="729A2A17C302C75CE233764DFDC1FDBE" box="[533,635,604,630]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="44" pageNumber="45">Fig. 160</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C302C75CE11E7692FED2FD26" pageId="44" pageNumber="45" type="distribution">
<paragraph id="EA1E3692C302C75CE11E7692FC29FD56" blockId="44.[264,1323,283,750]" box="[312,915,643,670]" pageId="44" pageNumber="45">
<emphasis id="D8D5EA80C302C75CE11E7692FE62FD55" bold="true" box="[312,472,643,669]" pageId="44" pageNumber="45">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C302C75CE2A37695FCB4FD56" box="[645,782,644,670]" name="El Salvador" pageId="44" pageNumber="45">El Salvador</collectingCountry>
(
<figureCitation id="729A2A17C302C75CE3397695FC3DFD56" box="[799,903,644,670]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="44" pageNumber="45">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C302C75CE11E76BAFED2FD26" blockId="44.[264,1323,283,750]" pageId="44" pageNumber="45">
<emphasis id="D8D5EA80C302C75CE11E76BAFDBCFD0D" bold="true" box="[312,518,683,709]" pageId="44" pageNumber="45">Natural History.</emphasis>
<taxonomicName id="2DA14D11C302C75CE23176BDFD4AFD0D" box="[535,752,684,709]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="badia">
<emphasis id="D8D5EA80C302C75CE23176BDFD4AFD0D" box="[535,752,684,709]" italics="true" pageId="44" pageNumber="45">Seeversiella badia</emphasis>
</taxonomicName>
was collected in cloud forest litter at altitude of
<quantity id="2D599B77C302C75CE12E76C5FEDEFD26" box="[264,356,724,750]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="44" pageNumber="45" unit="m" value="2100.0">2100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

246
data/62/08/87/62088784C309C759E12E76C2FE17F89D.xml Normal file
View file

@ -0,0 +1,246 @@
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<paragraph id="EA1E3692C309C757E12E76C2FC7BFD25" blockId="39.[264,961,723,750]" box="[264,961,723,750]" pageId="39" pageNumber="40">
<heading id="B15681FEC309C757E12E76C2FC7BFD25" bold="true" box="[264,961,723,750]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
<emphasis id="D8D5EA80C309C757E12E76C2FC7BFD25" bold="true" box="[264,961,723,750]" pageId="39" pageNumber="40">
<keyStep id="51552E37C309C757E12E76C2FC7BFD25" box="[264,961,723,750]" pageId="39" pageNumber="40">
<key id="9E3061A3C309C757E12E76C2FC7BFD25" box="[264,961,723,750]" pageId="39" pageNumber="40">
<keyLead id="515095A7C309C757E12E76C2FC7BFD25" box="[264,961,723,750]" pageId="39" pageNumber="40">
10.
<taxonomicName id="2DA14D11C309C757E11276C5FD08FD25" ID-CoL="6YBD6" authority="Gusarov" authorityName="Gusarov" box="[308,690,723,750]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C309C757E11276C5FDFBFD26" bold="true" box="[308,577,724,750]" italics="true" pageId="39" pageNumber="40">Seeversiella mexicana</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC309C757E29976C2FCB8FD25" box="[703,770,723,749]" pageId="39" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C309C757E33776C2FC03FD25" box="[785,953,723,749]" captionStart-0="FIGURES 135 ­ 138" captionStart-1="FIGURES 139 ­ 142" captionStart-2="FIGURES 143 ­ 149" captionStartId-0="39.[264,383,1739,1763]" captionStartId-1="40.[264,383,1421,1445]" captionStartId-2="41.[264,383,1483,1507]" captionTargetBox-0="[313,1272,932,1705]" captionTargetBox-1="[306,1279,781,1384]" captionTargetBox-2="[298,1288,744,1449]" captionTargetId-0="figure@39.[304,1283,924,1712]" captionTargetId-1="figure@40.[299,1288,769,1395]" captionTargetId-2="figure@41.[289,1297,729,1456]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="41" captionText-0="FIGURES 135 ­ 138. Abdominal segment 8 of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (135 ­ 136) and Suchixtepec (137 ­ 138), Mexico). 135 – male tergum 8; 136 – male sternum 8; 137 – female tergum 8; 138 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." captionText-2="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri-0="https://zenodo.org/record/156443/files/figure.png" httpUri-1="https://zenodo.org/record/156444/files/figure.png" httpUri-2="https://zenodo.org/record/156445/files/figure.png" pageId="39" pageNumber="40">Figs. 135­149</figureCitation>
)
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C309C758E12E770FFD44FE1E" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="EA1E3692C309C757E12E770FFBA9FCA9" blockId="39.[264,1323,798,865]" pageId="39" pageNumber="40">
<emphasis id="D8D5EA80C309C757E12E770FFE07FCF0" bold="true" box="[264,445,798,824]" pageId="39" pageNumber="40">
<typeStatus id="351A8830C309C757E12E770FFEF9FCF0" box="[264,323,798,824]" pageId="39" pageNumber="40">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C309C757E1F6770EFDFAFCF1" box="[464,576,799,825]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C309C757E257770FFCCAFCF0" bold="true" box="[625,880,798,824]" pageId="39" pageNumber="40">
<collectingCountry id="92B67602C309C757E257770FFD4BFCF0" box="[625,753,798,824]" name="Mexico" pageId="39" pageNumber="40">MEXICO</collectingCountry>
: Oaxaca:
</emphasis>
<quantity id="2D599B77C309C757E3A7770EFC6EFCF1" box="[897,980,799,825]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="39" pageNumber="40" unit="km" value="4.6">4.6 km</quantity>
S Suchixtepec,
<quantity id="2D599B77C309C757E4B3770EFB48FCF1" box="[1173,1266,799,825]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="39" pageNumber="40" unit="m" value="2150.0">2150 m</quantity>
, leaf litter in riparian alder forest (R.S.Anderson),
<date id="9E1F1052C309C757E33C7756FC1AFCA9" box="[794,928,839,865]" pageId="39" pageNumber="40" value="1992-07-23">23.vii.1992</date>
(
<collectionCode id="8CB0AE57C309C757E3897756FBBDFCA9" box="[943,1031,839,865]" pageId="39" pageNumber="40">KSEM</collectionCode>
).
</paragraph>
<caption id="BEDE661AC309C757E12E72DAFB31F8EF" httpUri="https://zenodo.org/record/156443/files/figure.png" pageId="39" pageNumber="40" targetBox="[313,1272,932,1705]" targetPageId="39">
<paragraph id="EA1E3692C309C757E12E72DAFB31F8EF" blockId="39.[264,1323,1739,1831]" pageId="39" pageNumber="40">
<emphasis id="D8D5EA80C309C757E12E72DAFE5BF92B" bold="true" box="[264,481,1739,1763]" pageId="39" pageNumber="40">FIGURES 135­138</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C309C757E33472DAFBDDF92A" authority="Gusarov" authorityName="Gusarov" box="[786,1127,1739,1762]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C309C757E33472DAFBB8F92A" box="[786,1026,1739,1762]" italics="true" pageId="39" pageNumber="40">Seeversiella mexicana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C309C757E45272DAFB08F92B" bold="true" box="[1140,1202,1739,1763]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="C3E657FBC309C757E45272DAFB08F92B" box="[1140,1202,1739,1763]" pageId="39" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Filo de Caballo (135­136) and Suchixtepec (137­138), Mexico). 135 – male tergum 8; 136 – male sternum 8; 137 – female tergum 8; 138 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C306C758E11E750DFD44FE1E" blockId="40.[264,1324,283,710]" pageId="40" pageNumber="41">
<typeStatus id="351A8830C306C758E11E750DFE14FEFE" box="[312,430,284,310]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C306C758E1E6750AFD06FEFD" bold="true" box="[448,700,283,309]" pageId="40" pageNumber="41">
<collectingCountry id="92B67602C306C758E1E6750AFDFAFEFD" box="[448,576,283,309]" name="Mexico" pageId="40" pageNumber="41">MEXICO</collectingCountry>
: Oaxaca:
</emphasis>
2, 5, same data as the
<typeStatus id="351A8830C306C758E43F750DFB39FEFE" box="[1049,1155,284,310]" pageId="40" pageNumber="41" type="holotype">holotype</typeStatus>
;, 2, ditto but
<quantity id="2D599B77C306C758E1107555FE33FE96" box="[310,393,324,350]" metricMagnitude="3" metricUnit="m" metricValue="5.1" pageId="40" pageNumber="41" unit="km" value="5.1">5.1 km</quantity>
S Suchixtepec, oak, pine and alder forest litter,
<date id="9E1F1052C306C758E3EE7555FBEBFE96" box="[968,1105,324,350]" pageId="40" pageNumber="41" value="1992-07-25">25.vii.1992</date>
;, 4, ditto but
<date id="9E1F1052C306C758E12E757DFE2BFE4E" box="[264,401,364,390]" pageId="40" pageNumber="41" value="1992-07-24">24.vii.1992</date>
;
<emphasis id="D8D5EA80C306C758E186757AFDA5FE4D" bold="true" box="[416,543,363,389]" pageId="40" pageNumber="41">Guerrero:</emphasis>
3, 4,
<quantity id="2D599B77C306C758E2E7757DFCB4FE4E" box="[705,782,364,390]" metricMagnitude="4" metricUnit="m" metricValue="7.1" pageId="40" pageNumber="41" unit="km" value="71.0">71 km</quantity>
NE Atoyac de Alvarez,
<quantity id="2D599B77C306C758E41C757DFB23FE4E" box="[1082,1177,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="40" pageNumber="41" unit="m" value="1700.0">1700 m</quantity>
, forest litter (J.S.Ashe),
<date id="9E1F1052C306C758E1B57585FDA6FE66" box="[403,540,404,430]" pageId="40" pageNumber="41" value="1992-07-25">25.vii.1992</date>
; 3,,
<quantity id="2D599B77C306C758E2B17585FD51FE66" box="[663,747,404,430]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="40" pageNumber="41" unit="km" value="5.6">5.6 km</quantity>
SW Filo de Caballo,
<quantity id="2D599B77C306C758E3D07585FBE9FE66" box="[1014,1107,404,430]" metricMagnitude="3" metricUnit="m" metricValue="2.31" pageId="40" pageNumber="41" unit="m" value="2310.0">2310 m</quantity>
, alder forest litter (R.S.Anderson),
<date id="9E1F1052C306C758E1E975ADFDEEFE1E" box="[463,596,444,470]" pageId="40" pageNumber="41" value="1992-07-13">13.vii.1992</date>
(all ­
<collectionCode id="8CB0AE57C306C758E2BD75ADFD48FE1E" box="[667,754,444,470]" pageId="40" pageNumber="41">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C306C758E11E75F2FCD6F8A2" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="EA1E3692C306C758E11E75F2FE3FFD0E" blockId="40.[264,1324,283,710]" pageId="40" pageNumber="41">
<emphasis id="D8D5EA80C306C758E11E75F2FE0DFE35" bold="true" box="[312,439,483,509]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C306C758E1EE75F5FD75FE35" box="[456,719,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="D8D5EA80C306C758E1EE75F5FD75FE35" box="[456,719,484,509]" italics="true" pageId="40" pageNumber="41">Seeversiella mexicana</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C306C758E4F575F5FEFDFDED" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C306C758E4F575F5FEFDFDED" italics="true" pageId="40" pageNumber="41">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.2­2.0 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra longer or a little shorter than pronotum (elytral length to pronotal length ratio 1.2­0.9); wings fully developed or reduced; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C306C758E3887695FBEFFD56" box="[942,1109,644,670]" captionStart-0="FIGURES 139 ­ 142" captionStart-1="FIGURES 143 ­ 149" captionStartId-0="40.[264,383,1421,1445]" captionStartId-1="41.[264,383,1483,1507]" captionTargetBox-0="[306,1279,781,1384]" captionTargetBox-1="[298,1288,744,1449]" captionTargetId-0="figure@40.[299,1288,769,1395]" captionTargetId-1="figure@41.[289,1297,729,1456]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." captionText-1="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri-0="https://zenodo.org/record/156444/files/figure.png" httpUri-1="https://zenodo.org/record/156445/files/figure.png" pageId="40" pageNumber="41">Figs. 139­148</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C306C758E13776BDFEC2FD0E" box="[273,376,684,710]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="40" pageNumber="41">Fig. 149</figureCitation>
).
</paragraph>
<caption id="BEDE661AC306C758E12E719CFDAEF9C5" httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="40" pageNumber="41" targetBox="[306,1279,781,1384]" targetPageId="40">
<paragraph id="EA1E3692C306C758E12E719CFDAEF9C5" blockId="40.[264,1323,1421,1549]" pageId="40" pageNumber="41">
<emphasis id="D8D5EA80C306C758E12E719CFE58FA6D" bold="true" box="[264,482,1421,1445]" pageId="40" pageNumber="41">FIGURES 139­142</emphasis>
. Aedeagus
<taxonomicName id="2DA14D11C306C758E254719FFC70FA6D" authority="Gusarov" authorityName="Gusarov" box="[626,970,1422,1445]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C306C758E254719FFCDEFA6D" box="[626,868,1422,1445]" italics="true" pageId="40" pageNumber="41">Seeversiella mexicana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C306C758E3FF719CFBA2FA6D" bold="true" box="[985,1048,1421,1445]" pageId="40" pageNumber="41">
<taxonomicNameLabel id="C3E657FBC306C758E3FF719CFBA2FA6D" box="[985,1048,1421,1445]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142).
</paragraph>
</caption>
<paragraph id="EA1E3692C306C758E11E7271FCD6F8A2" blockId="40.[264,1323,1632,1898]" pageId="40" pageNumber="41">
<taxonomicName id="2DA14D11C306C758E11E7271FDFAF9B1" box="[312,576,1632,1657]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="D8D5EA80C306C758E11E7271FDFAF9B1" box="[312,576,1632,1657]" italics="true" pageId="40" pageNumber="41">Seeversiella mexicana</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C306C758E3AD7271FB9DF9B1" box="[907,1063,1632,1657]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C306C758E3AD7271FB9DF9B1" box="[907,1063,1632,1657]" italics="true" pageId="40" pageNumber="41">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 5­8, 11­12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have strong spicules (
<figureCitation id="729A2A17C306C758E36D72C9FC09F93A" box="[843,947,1752,1778]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="40" pageNumber="41">Fig. 146</figureCitation>
); apex of median lobe in lateral view widened subapically and not emarginate on parameral side (
<figureCitation id="729A2A17C306C758E46F7311FB42F8D2" box="[1097,1272,1792,1818]" captionStart="FIGURES 139 ­ 142" captionStartId="40.[264,383,1421,1445]" captionTargetBox="[306,1279,781,1384]" captionTargetId="figure@40.[299,1288,769,1395]" captionTargetPageId="40" captionText="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="40" pageNumber="41">Figs. 141­142</figureCitation>
), in parameral view narrow and with weak basal constriction (
<figureCitation id="729A2A17C306C758E3917339FBDAF88A" box="[951,1120,1832,1858]" captionStart="FIGURES 139 ­ 142" captionStartId="40.[264,383,1421,1445]" captionTargetBox="[306,1279,781,1384]" captionTargetId="figure@40.[299,1288,769,1395]" captionTargetPageId="40" captionText="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="40" pageNumber="41">Figs. 139­140</figureCitation>
); spermatheca Sshaped with umbilicus facing proximally (
<figureCitation id="729A2A17C306C758E2DE7341FCE5F8A2" box="[760,863,1872,1898]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="40" pageNumber="41">Fig. 149</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C307C759E11E750AFDCDF947" pageId="41" pageNumber="42" type="description">
<paragraph id="EA1E3692C307C759E11E750AFC17FE96" blockId="41.[264,1323,283,630]" pageId="41" pageNumber="42">
<emphasis id="D8D5EA80C307C759E11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="41" pageNumber="42">Description.</emphasis>
Length
<quantity id="2D599B77C307C759E21F750DFD7FFEFE" box="[569,709,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="41" pageNumber="42" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C307C759E3A5750DFB96FEFE" box="[899,1068,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.3" metricValueMin="4.1" pageId="41" pageNumber="42" unit="mm" value="0.47" valueMax="0.53" valueMin="0.41">0.41­0.53 mm</quantity>
. Body brown to dark brown with lighter elytra, legs brown to yellowish brown.
</paragraph>
<paragraph id="EA1E3692C307C759E11E757DFB89FE66" blockId="41.[264,1323,283,630]" pageId="41" pageNumber="42">
In all character states
<taxonomicName id="2DA14D11C307C759E264757DFD68FE4D" box="[578,722,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="D8D5EA80C307C759E264757DFD68FE4D" box="[578,722,364,389]" italics="true" pageId="41" pageNumber="42">S. mexicana</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C307C759E392757DFBEAFE4D" box="[948,1104,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C307C759E392757DFBEAFE4D" box="[948,1104,364,389]" italics="true" pageId="41" pageNumber="42">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C307C759E11E75ADFD73FDEE" blockId="41.[264,1323,283,630]" pageId="41" pageNumber="42">In large males posterior angles of male tergum 3 projecting as spines up to twice as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to 2/3 as long as tergum length.</paragraph>
<paragraph id="EA1E3692C307C759E11E7625FD26FD86" blockId="41.[264,1323,283,630]" box="[312,668,564,590]" pageId="41" pageNumber="42">
Aedeagus as in
<figureCitation id="729A2A17C307C759E1D77625FD2DFD86" box="[497,663,564,590]" captionStart-0="FIGURES 139 ­ 142" captionStart-1="FIGURES 143 ­ 149" captionStartId-0="40.[264,383,1421,1445]" captionStartId-1="41.[264,383,1483,1507]" captionTargetBox-0="[306,1279,781,1384]" captionTargetBox-1="[298,1288,744,1449]" captionTargetId-0="figure@40.[299,1288,769,1395]" captionTargetId-1="figure@41.[289,1297,729,1456]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." captionText-1="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri-0="https://zenodo.org/record/156444/files/figure.png" httpUri-1="https://zenodo.org/record/156445/files/figure.png" pageId="41" pageNumber="42">Figs. 139­148</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C307C759E11E764DFDC4FDBE" blockId="41.[264,1323,283,630]" box="[312,638,604,630]" pageId="41" pageNumber="42">
Spermatheca as in
<figureCitation id="729A2A17C307C759E233764DFDC0FDBE" box="[533,634,604,630]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="41" pageNumber="42">Fig. 149</figureCitation>
.
</paragraph>
<caption id="BEDE661AC307C759E12E71DAFBB5F9A5" httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="41" pageNumber="42" targetBox="[298,1288,744,1449]" targetPageId="41">
<paragraph id="EA1E3692C307C759E12E71DAFBB5F9A5" blockId="41.[264,1325,1483,1679]" pageId="41" pageNumber="42">
<emphasis id="D8D5EA80C307C759E12E71DAFE5AFA2B" bold="true" box="[264,480,1483,1507]" pageId="41" pageNumber="42">FIGURES 143­149</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C307C759E2A471DDFC6FFA2A" authority="Gusarov" authorityName="Gusarov" box="[642,981,1483,1507]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C307C759E2A471DDFCC8FA2B" box="[642,882,1484,1507]" italics="true" pageId="41" pageNumber="42">Seeversiella mexicana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C307C759E3C471DAFB9BFA2B" bold="true" box="[994,1057,1483,1507]" pageId="41" pageNumber="42">
<taxonomicNameLabel id="C3E657FBC307C759E3C471DAFB9BFA2B" box="[994,1057,1483,1507]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Filo de Caballo (143, 145­148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145­146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145­146), 0.1 mm (144, 147­149).
</paragraph>
</caption>
<paragraph id="EA1E3692C307C759E12E7269FDCDF947" blockId="41.[264,1325,1483,1679]" box="[264,631,1656,1679]" pageId="41" pageNumber="42">DS – distal sclerite of internal sac.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C307C759E11E72FAFE17F89D" pageId="41" pageNumber="42" type="distribution">
<paragraph id="EA1E3692C307C759E11E72FAFE17F89D" blockId="41.[264,1323,1771,1877]" pageId="41" pageNumber="42">
<emphasis id="D8D5EA80C307C759E11E72FAFE62F8CD" bold="true" box="[312,472,1771,1797]" pageId="41" pageNumber="42">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C307C759E2A372FAFD65F8CD" box="[645,735,1771,1797]" name="Mexico" pageId="41" pageNumber="42">Mexico</collectingCountry>
(
<figureCitation id="729A2A17C307C759E2C972FAFCEDF8CD" box="[751,855,1771,1797]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="41" pageNumber="42">Fig. 387</figureCitation>
).
<emphasis id="D8D5EA80C307C759E11E7302FDB3F8E5" bold="true" box="[312,521,1811,1837]" pageId="41" pageNumber="42">Natural History.</emphasis>
<taxonomicName id="2DA14D11C307C759E2387302FC9DF8E4" box="[542,807,1811,1836]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="D8D5EA80C307C759E2387302FC9DF8E4" box="[542,807,1811,1836]" italics="true" pageId="41" pageNumber="42">Seeversiella mexicana</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C307C759E12E732AFE12F89D" box="[264,424,1851,1877]" metricMagnitude="3" metricUnit="m" metricValue="2.005" metricValueMax="2.31" metricValueMin="1.7" pageId="41" pageNumber="42" unit="m" value="2005.0" valueMax="2310.0" valueMin="1700.0">1700­2310 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

168
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<treatment id="62088784C30BC757E12E77D2FD1DFDBE" ID-DOI="http://doi.org/10.5281/zenodo.6274652" ID-Zenodo-Dep="6274652" LSID="urn:lsid:plazi:treatment:62088784C30BC757E12E77D2FD1DFDBE" httpUri="http://treatment.plazi.org/id/62088784C30BC757E12E77D2FD1DFDBE" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
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<paragraph id="EA1E3692C30BC755E12E77D2FC22FC15" blockId="37.[264,920,963,990]" box="[264,920,963,990]" pageId="37" pageNumber="38">
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<emphasis id="D8D5EA80C30BC755E12E77D2FC22FC15" bold="true" box="[264,920,963,990]" pageId="37" pageNumber="38">
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9.
<taxonomicName id="2DA14D11C30BC755E10077D5FD30FC15" ID-CoL="6YBD7" authority="Gusarov" authorityName="Gusarov" box="[294,650,963,990]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="D8D5EA80C30BC755E10077D5FDADFC16" bold="true" box="[294,535,964,990]" italics="true" pageId="37" pageNumber="38">Seeversiella grandis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC30BC755E2B177D2FD63FC15" box="[663,729,963,989]" pageId="37" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C30BC755E2CF77D2FC2BFC15" box="[745,913,963,989]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 – male tergum 8; 127 – male sternum 8; 128 – median lobe, parameral view; 129 – apex of median lobe, parameral view; 130 – median lobe, lateral view; 131 – apex of median lobe, lateral view; 132 – details of internal sac retracted into median lobe, abparameral view; 133 – details of internal sac retracted into median lobe, lateral view; 134 – apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="37" pageNumber="38">Figs. 126­134</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30BC755E12E701FFBF7FB99" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="EA1E3692C30BC755E12E701FFBF7FB99" blockId="37.[264,1324,1038,1904]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E12E701FFE07FBE0" bold="true" box="[264,445,1038,1064]" pageId="37" pageNumber="38">
<typeStatus id="351A8830C30BC755E12E701FFEF9FBE0" box="[264,323,1038,1064]" pageId="37" pageNumber="38">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C30BC755E1F4701EFDF8FBE1" box="[466,578,1039,1065]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C30BC755E255701FFBA7FBE0" bold="true" box="[627,1053,1038,1064]" pageId="37" pageNumber="38">
<collectingCountry id="92B67602C30BC755E255701FFC82FBE0" box="[627,824,1038,1064]" name="Guatemala" pageId="37" pageNumber="38">GUATEMALA</collectingCountry>
: Quetzaltenango:
</emphasis>
<quantity id="2D599B77C30BC755E409701EFBD7FBE1" box="[1071,1133,1039,1065]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="38" unit="km" value="8.0">8 km</quantity>
SE Zunil,
<quantity id="2D599B77C30BC755E4D6701EFE98FB99" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="37" pageNumber="38" unit="m" value="2620.0">2620 m</quantity>
, flight intercept trap (J.S.Ashe &amp; R.Brooks),
<date id="9E1F1052C30BC755E3137026FC60FB99" box="[821,986,1079,1105]" pageId="37" pageNumber="38" value="1993-06-19" valueMax="1993-06-21" valueMin="1993-06-19">19­21.vi.1993</date>
(
<collectionCode id="8CB0AE57C30BC755E3CF7026FBFBFB99" box="[1001,1089,1079,1105]" pageId="37" pageNumber="38">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30BC755E11E704FFBEAFAD1" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="EA1E3692C30BC755E11E704FFBEAFAD1" blockId="37.[264,1324,1038,1904]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E11E704FFE0DFBB0" bold="true" box="[312,439,1118,1144]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C30BC755E1EA704EFD04FBB0" box="[460,702,1119,1144]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="D8D5EA80C30BC755E1EA704EFD04FBB0" box="[460,702,1119,1144]" italics="true" pageId="37" pageNumber="38">Seeversiella grandis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C30BC755E4F5704EFEFDFB68" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C30BC755E4F5704EFEFDFB68" italics="true" pageId="37" pageNumber="38">Seeversiella</emphasis>
</taxonomicName>
by having large body (
<quantity id="2D599B77C30BC755E2457096FD7FFB69" box="[611,709,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="37" pageNumber="38" unit="mm" value="3.7">3.7 mm</quantity>
), brownish black body with brown elytra and legs; large eyes (temple length to eye length ratio 0.9); matte pronotum with strong isodiametric microsculpture; elytra longer than pronotum; wings fully developed; tergum 8 with four pairs of macrosetae; and the distinct shape of aedeagus (
<figureCitation id="729A2A17C30BC755E3BD70EEFBF9FAD1" box="[923,1091,1279,1305]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 – male tergum 8; 127 – male sternum 8; 128 – median lobe, parameral view; 129 – apex of median lobe, parameral view; 130 – median lobe, lateral view; 131 – apex of median lobe, lateral view; 132 – details of internal sac retracted into median lobe, abparameral view; 133 – details of internal sac retracted into median lobe, lateral view; 134 – apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="37" pageNumber="38">Figs. 129­134</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30BC757E11E7137FDB1FE36" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
<paragraph id="EA1E3692C30BC755E11E7137FB0CFA89" blockId="37.[264,1324,1038,1904]" box="[312,1206,1318,1345]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E11E7137FE6AFA88" bold="true" box="[312,464,1318,1344]" pageId="37" pageNumber="38">Description.</emphasis>
Length
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. Body brownish black, elytra and legs brown.
</paragraph>
<paragraph id="EA1E3692C30BC755E11E715EFBF7FA28" blockId="37.[264,1324,1038,1904]" pageId="37" pageNumber="38">Head surface matte, with fine isodiametric microsculpture, fine punctation, distance between punctures equals 1­2 times their diameter, anteromedial portion of the disc without punctation. Temple length to eye length ratio 0.9. Antennal article 2 as long as 3, article 4 elongate (ratio 1.2), 5 quadrate, 6­10 transverse (ratio 1.2­1.3).</paragraph>
<paragraph id="EA1E3692C30BC755E11E71FFFC99F930" blockId="37.[264,1324,1038,1904]" pageId="37" pageNumber="38">
Pronotum slightly transverse, 1.3 times as wide as head, width
<quantity id="2D599B77C30BC755E40D71FFFB23F9C0" box="[1067,1177,1518,1544]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="37" pageNumber="38" unit="mm" value="0.74">0.74 mm</quantity>
, length
<quantity id="2D599B77C30BC755E4DF71FFFE83F9F8" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="37" pageNumber="38" unit="mm" value="0.66">0.66 mm</quantity>
, width to length ratio 1.1, surface matte, with fine isodiametric microsculpture and fine punctation, distance between punctures equals ½­1 times their diameter. Elytra wider and longer (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 0.8), 1.3 times wider than long, glossy, with fine and weak isodiametric microsculpture; with fine and asperate punctation, distance between punctures equal to ½­ 1 time their diameter. Wings fully developed.
</paragraph>
<paragraph id="EA1E3692C30BC756E11E7317FDCAFE4E" blockId="37.[264,1324,1038,1904]" lastBlockId="38.[264,1325,284,390]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
Abdominal terga glossy, terga 3­6 with fine microsculpture consisting of transverse waves and strongly transverse meshes, with fine punctation, distance between punctures equals 3­6 times their diameter. Tergum 7 with microsculpture consisting of transverse meshes and with fine and sparse punctation, distance between punctures equal to 4­7 times their diameter. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C308C756E1DA757DFDD9FE4E" box="[508,611,364,390]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 – male tergum 8; 127 – male sternum 8; 128 – median lobe, parameral view; 129 – apex of median lobe, parameral view; 130 – median lobe, lateral view; 131 – apex of median lobe, lateral view; 132 – details of internal sac retracted into median lobe, abparameral view; 133 – details of internal sac retracted into median lobe, lateral view; 134 – apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="38" pageNumber="39">Fig. 126</figureCitation>
).
</paragraph>
<caption id="BEDE661AC308C756E12E7264FB85F880" httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="38" pageNumber="39" targetBox="[283,1299,500,1615]" targetPageId="38">
<paragraph id="EA1E3692C308C756E12E7264FB85F880" blockId="38.[264,1323,1653,1864]" pageId="38" pageNumber="39">
<emphasis id="D8D5EA80C308C756E12E7264FE5AF945" bold="true" box="[264,480,1653,1677]" pageId="38" pageNumber="39">FIGURES 126­134</emphasis>
. Abdominal segment 8 and aedeagus of
<taxonomicName id="2DA14D11C308C756E3857267FB58F944" authority="Gusarov" authorityName="Gusarov" box="[931,1250,1653,1677]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="D8D5EA80C308C756E3857267FBC5F945" box="[931,1151,1654,1677]" italics="true" pageId="38" pageNumber="39">Seeversiella grandis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C308C756E4CB7264FA90F945" bold="true" box="[1261,1322,1653,1677]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="C3E657FBC308C756E4CB7264FA90F945" box="[1261,1322,1653,1677]" pageId="38" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 126 – male tergum 8; 127 – male sternum 8; 128 – median lobe, parameral view; 129 – apex of median lobe, parameral view; 130 – median lobe, lateral view; 131 – apex of median lobe, lateral view; 132 – details of internal sac retracted into median lobe, abparameral view; 133 – details of internal sac retracted into median lobe, lateral view; 134 – apex of left paramere. Scale bar 0.4 mm (126­127), 0.3 mm (128, 130, 132­133), 0.1 mm (129, 131, 134).
</paragraph>
</caption>
<paragraph id="EA1E3692C309C757E11E750DFC61FE1E" blockId="39.[264,1323,284,630]" pageId="39" pageNumber="40">
In the only known male specimen pronotum with wide medial impression, posterior margin of male tergum 3 with two short and obtuse projections (as in
<figureCitation id="729A2A17C309C757E4777555FB13FE96" box="[1105,1193,324,350]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="39" pageNumber="40">Fig. 19</figureCitation>
), tergum 7 with medial tubercle in front of posterior margin. Posterior margin of male tergum 8 without emargination (
<figureCitation id="729A2A17C309C757E1C77585FDF2FE66" box="[481,584,404,430]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 – male tergum 8; 127 – male sternum 8; 128 – median lobe, parameral view; 129 – apex of median lobe, parameral view; 130 – median lobe, lateral view; 131 – apex of median lobe, lateral view; 132 – details of internal sac retracted into median lobe, abparameral view; 133 – details of internal sac retracted into median lobe, lateral view; 134 – apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="39" pageNumber="40">Fig. 126</figureCitation>
). Aedeagus as in
<figureCitation id="729A2A17C309C757E3047585FC73FE66" box="[802,969,404,430]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 – male tergum 8; 127 – male sternum 8; 128 – median lobe, parameral view; 129 – apex of median lobe, parameral view; 130 – median lobe, lateral view; 131 – apex of median lobe, lateral view; 132 – details of internal sac retracted into median lobe, abparameral view; 133 – details of internal sac retracted into median lobe, lateral view; 134 – apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="39" pageNumber="40">Figs. 129­134</figureCitation>
. Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C309C757E34175ADFC74FE1E" box="[871,974,444,470]" captionStart="FIGURES 126 ­ 134" captionStartId="38.[264,383,1653,1677]" captionTargetBox="[283,1299,500,1615]" captionTargetId="figure@38.[273,1314,489,1626]" captionTargetPageId="38" captionText="FIGURES 126 ­ 134. Abdominal segment 8 and aedeagus of Seeversiella grandis Gusarov, sp. n. (holotype). 126 – male tergum 8; 127 – male sternum 8; 128 – median lobe, parameral view; 129 – apex of median lobe, parameral view; 130 – median lobe, lateral view; 131 – apex of median lobe, lateral view; 132 – details of internal sac retracted into median lobe, abparameral view; 133 – details of internal sac retracted into median lobe, lateral view; 134 – apex of left paramere. Scale bar 0.4 mm (126 ­ 127), 0.3 mm (128, 130, 132 ­ 133), 0.1 mm (129, 131, 134)." httpUri="https://zenodo.org/record/156442/files/figure.png" pageId="39" pageNumber="40">Fig. 134</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C309C757E11E75F5FDB1FE36" blockId="39.[264,1323,284,630]" box="[312,523,484,510]" pageId="39" pageNumber="40">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C309C757E11E761AFD1DFDBE" pageId="39" pageNumber="40" type="distribution">
<paragraph id="EA1E3692C309C757E11E761AFC33FDEE" blockId="39.[264,1323,284,630]" box="[312,905,523,550]" pageId="39" pageNumber="40">
<emphasis id="D8D5EA80C309C757E11E761AFE62FDED" bold="true" box="[312,472,523,549]" pageId="39" pageNumber="40">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C309C757E2A3761DFCBEFDEE" box="[645,772,524,550]" name="Guatemala" pageId="39" pageNumber="40">Guatemala</collectingCountry>
(
<figureCitation id="729A2A17C309C757E332761DFCC6FDEE" box="[788,892,524,550]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="39" pageNumber="40">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C309C757E11E7622FD1DFDBE" blockId="39.[264,1323,284,630]" pageId="39" pageNumber="40">
<emphasis id="D8D5EA80C309C757E11E7622FDBFFD85" bold="true" box="[312,517,563,589]" pageId="39" pageNumber="40">Natural History.</emphasis>
The only known specimen of
<taxonomicName id="2DA14D11C309C757E3527625FC56FD85" box="[884,1004,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="D8D5EA80C309C757E3527625FC56FD85" box="[884,1004,564,589]" italics="true" pageId="39" pageNumber="40">S. grandis</emphasis>
</taxonomicName>
was collected with a flight intercept trap at altitude of
<quantity id="2D599B77C309C757E261764DFD19FDBE" box="[583,675,604,630]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="39" pageNumber="40" unit="m" value="2620.0">2620 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

191
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@ -0,0 +1,191 @@
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<mods:detail id="42E92DF706DB184E3EB015139C3E5FA8" type="volume">
<mods:number id="E44F8070E19C561582A51425C4F2AD2F">142</mods:number>
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<mods:classification id="02280353E69306FD47E0C79B8C17C387">journal article</mods:classification>
<mods:identifier id="DCE8F0DDE0D7A5AC92EF0C55ABF55C65" type="CLB-Dataset">51371</mods:identifier>
<mods:identifier id="7779120051DD2D44A0769B5D52CD1EFF" type="DOI">10.5281/zenodo.156420</mods:identifier>
<mods:identifier id="CCE6C61C29AA0DE868192A971918444E" type="GBIF-Dataset">76c096e3-d27c-4abc-a504-a62ae8c7ba92</mods:identifier>
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<treatment id="62088784C30DC755E12E7252FD90FCAE" ID-DOI="http://doi.org/10.5281/zenodo.6274650" ID-Zenodo-Dep="6274650" LSID="urn:lsid:plazi:treatment:62088784C30DC755E12E7252FD90FCAE" httpUri="http://treatment.plazi.org/id/62088784C30DC755E12E7252FD90FCAE" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36">
<subSubSection id="A2BB6519C30DC753E12E7252FC13F995" box="[264,937,1603,1629]" pageId="35" pageNumber="36" type="key">
<paragraph id="EA1E3692C30DC753E12E7252FC13F995" blockId="35.[264,937,1603,1629]" box="[264,937,1603,1629]" pageId="35" pageNumber="36">
<heading id="B15681FEC30DC753E12E7252FC13F995" bold="true" box="[264,937,1603,1629]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<emphasis id="D8D5EA80C30DC753E12E7252FC13F995" bold="true" box="[264,937,1603,1629]" pageId="35" pageNumber="36">
<key id="9E3061A3C30DC753E12E7252FC13F995" box="[264,937,1603,1629]" pageId="35" pageNumber="36">
<keyStep id="51552E37C30DC753E12E7252FC13F995" box="[264,937,1603,1629]" pageId="35" pageNumber="36">
<keyLead id="515095A7C30DC753E12E7252FC13F995" box="[264,937,1603,1629]" pageId="35" pageNumber="36">
8.
<taxonomicName id="2DA14D11C30DC753E1007252FD27F995" ID-CoL="6YBCY" authority="Gusarov" authorityName="Gusarov" box="[294,669,1603,1629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="nigriceps" status="sp. nov.">
<emphasis id="D8D5EA80C30DC753E1007252FD91F995" bold="true" box="[294,555,1603,1629]" italics="true" pageId="35" pageNumber="36">Seeversiella nigriceps</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC30DC753E28C7252FD56F995" box="[682,748,1603,1629]" pageId="35" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C30DC753E2DA7252FC18F995" box="[764,930,1603,1629]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 – male tergum 8; 118 – male sternum 8; 119 – median lobe, parameral view; 120 – apex of median lobe, parameral view; 121 – median lobe, lateral view; 122 – apex of median lobe, lateral view; 123 ­ 124 – details of internal sac retracted into median lobe, lateral view; 125 – apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS – distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="35" pageNumber="36">Figs. 117­125</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30DC753E12E729CFD4BF918" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="EA1E3692C30DC753E12E729CFD4BF918" blockId="35.[264,1324,1677,1904]" pageId="35" pageNumber="36">
<emphasis id="D8D5EA80C30DC753E12E729CFE04F96F" bold="true" box="[264,446,1677,1703]" pageId="35" pageNumber="36">
<typeStatus id="351A8830C30DC753E12E729CFEFFF96F" box="[264,325,1677,1703]" pageId="35" pageNumber="36">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C30DC753E1EB729FFD87F960" box="[461,573,1678,1704]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C30DC753E24D729CFCE4F96F" bold="true" box="[619,862,1677,1703]" pageId="35" pageNumber="36">
<collectingCountry id="92B67602C30DC753E24D729CFD51F96F" box="[619,747,1677,1703]" name="Mexico" pageId="35" pageNumber="36">MEXICO</collectingCountry>
: Puebla:
</emphasis>
<quantity id="2D599B77C30DC753E348729FFC05F960" box="[878,959,1678,1704]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="35" pageNumber="36" unit="km" value="1.6">1.6 km</quantity>
E Nicolas Bravo,
<quantity id="2D599B77C30DC753E4B1729FFB48F960" box="[1175,1266,1678,1704]" metricMagnitude="3" metricUnit="m" metricValue="2.41" pageId="35" pageNumber="36" unit="m" value="2410.0">2410 m</quantity>
, forest litter (J.S.Ashe),
<date id="9E1F1052C30DC753E1D172A7FDC7F918" box="[503,637,1718,1744]" pageId="35" pageNumber="36" value="1992-07-17">17.vii.1992</date>
(
<collectionCode id="8CB0AE57C30DC753E2AA72A7FD5EF918" box="[652,740,1718,1744]" pageId="35" pageNumber="36">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30DC754E11E72CCFC6DFE96" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="EA1E3692C30DC753E11E72CCFA91F8B8" blockId="35.[264,1324,1677,1904]" pageId="35" pageNumber="36">
<emphasis id="D8D5EA80C30DC753E11E72CCFE0DF93F" bold="true" box="[312,439,1757,1783]" pageId="35" pageNumber="36">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C30DC753E1EE72CFFD76F93F" box="[456,716,1758,1783]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C30DC753E1EE72CFFD76F93F" box="[456,716,1758,1783]" italics="true" pageId="35" pageNumber="36">Seeversiella nigriceps</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C30DC753E4F572CFFEFDF8D7" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C30DC753E4F572CFFEFDF8D7" italics="true" pageId="35" pageNumber="36">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples twice as long as eyes; glossy pronotum with weak and partially missing microsculpture; elytra longer than pronotum; wings fully developed; tergum 8 with four pairs of macrosetae; and the distinct shape of aedeagus (
<figureCitation id="729A2A17C30DC753E4517347FAA5F8B8" box="[1143,1311,1878,1904]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 – male tergum 8; 118 – male sternum 8; 119 – median lobe, parameral view; 120 – apex of median lobe, parameral view; 121 – median lobe, lateral view; 122 – apex of median lobe, lateral view; 123 ­ 124 – details of internal sac retracted into median lobe, lateral view; 125 – apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS – distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="35" pageNumber="36">Figs. 119­125</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30AC754E11E750DFC6DFE96" blockId="36.[264,1323,284,350]" pageId="36" pageNumber="37">
<taxonomicName id="2DA14D11C30AC754E11E750DFD80FEFD" box="[312,570,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C30AC754E11E750DFD80FEFD" box="[312,570,284,309]" italics="true" pageId="36" pageNumber="37">Seeversiella nigriceps</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C30AC754E3A0750DFBFFFEFD" box="[902,1093,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C30AC754E3A0750DFBFFFEFD" box="[902,1093,284,309]" italics="true" pageId="36" pageNumber="37">S. sonomotoides</emphasis>
</taxonomicName>
in having narrower distal sclerites of internal sac of aedeagus (
<figureCitation id="729A2A17C30AC754E2D97555FCCAFE96" box="[767,880,324,350]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 – male tergum 8; 118 – male sternum 8; 119 – median lobe, parameral view; 120 – apex of median lobe, parameral view; 121 – median lobe, lateral view; 122 – apex of median lobe, lateral view; 123 ­ 124 – details of internal sac retracted into median lobe, lateral view; 125 – apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS – distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="36" pageNumber="37">Figs. 124</figureCitation>
; 63, 65).
</paragraph>
</subSubSection>
<caption id="BEDE661AC30AC754E12E727DFDCDF8F8" httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="36" pageNumber="37" targetBox="[297,1299,423,1607]" targetPageId="36">
<paragraph id="EA1E3692C30AC754E12E727DFDCDF8F8" blockId="36.[264,1322,1644,1840]" pageId="36" pageNumber="37">
<emphasis id="D8D5EA80C30AC754E12E727DFE67F94C" bold="true" box="[264,477,1644,1668]" pageId="36" pageNumber="37">FIGURES 117­125</emphasis>
. Abdominal segment 8 and aedeagus of
<taxonomicName id="2DA14D11C30AC754E3B1727DFB59F94B" authority="Gusarov" authorityName="Gusarov" box="[919,1251,1644,1667]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nigriceps" status="sp. nov.">
<emphasis id="D8D5EA80C30AC754E3B1727DFB3BF94B" box="[919,1153,1644,1667]" italics="true" pageId="36" pageNumber="37">Seeversiella nigriceps</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C30AC754E4C9727DFA90F94C" bold="true" box="[1263,1322,1644,1668]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="C3E657FBC30AC754E4C9727DFA90F94C" box="[1263,1322,1644,1668]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 117 – male tergum 8; 118 – male sternum 8; 119 – median lobe, parameral view; 120 – apex of median lobe, parameral view; 121 – median lobe, lateral view; 122 – apex of median lobe, lateral view; 123­124 – details of internal sac retracted into median lobe, lateral view; 125 – apex of left paramere. Scale bar 0.3 mm (117­118), 0.2 mm (119, 121, 123­124), 0.1 mm (120, 122, 125). DS – distal sclerite of internal sac.
</paragraph>
</caption>
<subSubSection id="A2BB6519C30BC755E11E750DFDB0FD26" pageId="37" pageNumber="38" type="description">
<paragraph id="EA1E3692C30BC755E11E750DFB6EFE66" blockId="37.[264,1324,284,870]" pageId="37" pageNumber="38">
<taxonomicName id="2DA14D11C30BC755E11E750DFD84FEFD" box="[312,574,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C30BC755E11E750DFD84FEFD" box="[312,574,284,309]" italics="true" pageId="37" pageNumber="38">Seeversiella nigriceps</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C30BC755E3AC750DFB9CFEFD" box="[906,1062,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C30BC755E3AC750DFB9CFEFD" box="[906,1062,284,309]" italics="true" pageId="37" pageNumber="38">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (5­7, 10­12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac sclerotized and dentiform (
<figureCitation id="729A2A17C30BC755E4DA757DFE82FE66" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 – male tergum 8; 118 – male sternum 8; 119 – median lobe, parameral view; 120 – apex of median lobe, parameral view; 121 – median lobe, lateral view; 122 – apex of median lobe, lateral view; 123 ­ 124 – details of internal sac retracted into median lobe, lateral view; 125 – apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS – distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="37" pageNumber="38">Fig. 124</figureCitation>
); apex of median lobe in lateral view not widened subapically (
<figureCitation id="729A2A17C30BC755E4397585FB7DFE66" box="[1055,1223,404,430]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 – male tergum 8; 118 – male sternum 8; 119 – median lobe, parameral view; 120 – apex of median lobe, parameral view; 121 – median lobe, lateral view; 122 – apex of median lobe, lateral view; 123 ­ 124 – details of internal sac retracted into median lobe, lateral view; 125 – apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS – distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="37" pageNumber="38">Figs. 121­122</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30BC755E11E75AAFE2BFE36" blockId="37.[264,1324,284,870]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E11E75AAFE6AFE1D" bold="true" box="[312,464,443,469]" pageId="37" pageNumber="38">Description.</emphasis>
Length
<quantity id="2D599B77C30BC755E21D75ADFD2DFE1E" box="[571,663,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="37" pageNumber="38" unit="mm" value="2.4">2.4 mm</quantity>
, pronotal width
<quantity id="2D599B77C30BC755E37E75ADFC7FFE1E" box="[856,965,444,470]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="37" pageNumber="38" unit="mm" value="0.47">0.47 mm</quantity>
. Body dark brown, elytra and legs brown.
</paragraph>
<paragraph id="EA1E3692C30BC755E11E761DFBF5FD86" blockId="37.[264,1324,284,870]" pageId="37" pageNumber="38">
In all character states
<taxonomicName id="2DA14D11C30BC755E21F761DFD79FDED" box="[569,707,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C30BC755E21F761DFD79FDED" box="[569,707,524,549]" italics="true" pageId="37" pageNumber="38">S. nigriceps</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C30BC755E3BC761DFBE3FDED" box="[922,1113,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C30BC755E3BC761DFBE3FDED" box="[922,1113,524,549]" italics="true" pageId="37" pageNumber="38">S. sonomotoides</emphasis>
</taxonomicName>
, but differs in the shape of the distal sclerites of internal sac as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C30BC755E11E764DFBCDFD56" blockId="37.[264,1324,284,870]" pageId="37" pageNumber="38">
In the only known male specimen the posterior angles of male tergum 3 not projecting as spines, tergum
<quantity id="2D599B77C30BC755E1FC7695FDBDFD56" box="[474,519,644,670]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="37" pageNumber="38" unit="in" value="7.0">7 in</quantity>
front of posterior margin with small medial tubercle.
</paragraph>
<paragraph id="EA1E3692C30BC755E11E76BDFD20FD0E" blockId="37.[264,1324,284,870]" box="[312,666,684,710]" pageId="37" pageNumber="38">
Aedeagus as in
<figureCitation id="729A2A17C30BC755E1D676BDFD2CFD0E" box="[496,662,684,710]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 – male tergum 8; 118 – male sternum 8; 119 – median lobe, parameral view; 120 – apex of median lobe, parameral view; 121 – median lobe, lateral view; 122 – apex of median lobe, lateral view; 123 ­ 124 – details of internal sac retracted into median lobe, lateral view; 125 – apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS – distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="37" pageNumber="38">Figs. 119­125</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C30BC755E11E76C5FDB0FD26" blockId="37.[264,1324,284,870]" box="[312,522,724,750]" pageId="37" pageNumber="38">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30BC755E11E76EAFD90FCAE" pageId="37" pageNumber="38" type="distribution">
<paragraph id="EA1E3692C30BC755E11E76EAFCD8FCDE" blockId="37.[264,1324,284,870]" box="[312,866,763,790]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E11E76EAFE62FCDD" bold="true" box="[312,472,763,789]" pageId="37" pageNumber="38">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C30BC755E2A376EDFD65FCDE" box="[645,735,764,790]" name="Mexico" pageId="37" pageNumber="38">Mexico</collectingCountry>
(
<figureCitation id="729A2A17C30BC755E2C876EDFCEFFCDE" box="[750,853,764,790]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="37" pageNumber="38">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30BC755E11E7732FD90FCAE" blockId="37.[264,1324,284,870]" pageId="37" pageNumber="38">
<emphasis id="D8D5EA80C30BC755E11E7732FDBEFCF5" bold="true" box="[312,516,803,829]" pageId="37" pageNumber="38">Natural History.</emphasis>
The only known specimen of
<taxonomicName id="2DA14D11C30BC755E34A7735FC4CFCF5" box="[876,1014,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C30BC755E34A7735FC4CFCF5" box="[876,1014,804,829]" italics="true" pageId="37" pageNumber="38">S. nigriceps</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C30BC755E1EC775DFD9FFCAE" box="[458,549,844,870]" metricMagnitude="3" metricUnit="m" metricValue="2.41" pageId="37" pageNumber="38" unit="m" value="2410.0">2410 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

201
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<document id="18FB544ABE09F23031CA926E62D28BDD" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C30FC753E12E7162FE5DFE4E" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella tuberculicauda Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="36" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="34" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="0E57A0CCF8A554F7241A28DA324D52AC">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C30FC751E12E7162FC48FA45" box="[264,1010,1395,1422]" pageId="33" pageNumber="34" type="key">
<paragraph id="EA1E3692C30FC751E12E7162FC48FA45" blockId="33.[264,1010,1395,1422]" box="[264,1010,1395,1422]" pageId="33" pageNumber="34">
<heading id="B15681FEC30FC751E12E7162FC48FA45" bold="true" box="[264,1010,1395,1422]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="D8D5EA80C30FC751E12E7162FC48FA45" bold="true" box="[264,1010,1395,1422]" pageId="33" pageNumber="34">
<keyStep id="51552E37C30FC751E12E7162FC48FA45" box="[264,1010,1395,1422]" pageId="33" pageNumber="34">
<key id="9E3061A3C30FC751E12E7162FC48FA45" box="[264,1010,1395,1422]" pageId="33" pageNumber="34">
<keyLead id="515095A7C30FC751E12E7162FC48FA45" box="[264,1010,1395,1422]" pageId="33" pageNumber="34">
7.
<taxonomicName id="2DA14D11C30FC751E1007165FD5FFA45" ID-CoL="6YBBJ" authority="Gusarov" authorityName="Gusarov" box="[294,741,1395,1422]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tuberculicauda" status="sp. nov.">
<emphasis id="D8D5EA80C30FC751E1007165FDC8FA46" bold="true" box="[294,626,1396,1422]" italics="true" pageId="33" pageNumber="34">Seeversiella tuberculicauda</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC30FC751E2D47162FC8EFA45" box="[754,820,1395,1421]" pageId="33" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C30FC751E3627162FC50FA45" box="[836,1002,1395,1421]" captionStart-0="FIGURES 105 ­ 108" captionStart-1="FIGURES 109 ­ 116" captionStartId-0="34.[264,383,1348,1372]" captionStartId-1="35.[264,383,1334,1358]" captionTargetBox-0="[296,1289,499,1312]" captionTargetBox-1="[306,1282,441,1310]" captionTargetId-0="figure@34.[285,1302,489,1322]" captionTargetId-1="figure@35.[293,1295,433,1320]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionText-0="FIGURES 105 ­ 108. Abdominal segment 8 of Seeversiella tuberculicauda Gusarov, sp. n. (105 ­ 106, holotype; 107 ­ 108, paratype from Jalapa, Mexico). 105 – male tergum 8; 106 – male sternum 8; 107 – female tergum 8; 108 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri-0="https://zenodo.org/record/156439/files/figure.png" httpUri-1="https://zenodo.org/record/156440/files/figure.png" pageId="33" pageNumber="34">Figs. 105­116</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30FC751E12E71AFFB19F9E0" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="EA1E3692C30FC751E12E71AFFD45F9C8" blockId="33.[264,1323,1470,1896]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E12E71AFFE07FA10" bold="true" box="[264,445,1470,1496]" pageId="33" pageNumber="34">
<typeStatus id="351A8830C30FC751E12E71AFFEF9FA10" box="[264,323,1470,1496]" pageId="33" pageNumber="34">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C30FC751E1F471AFFDF8FA10" box="[466,578,1470,1496]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C30FC751E25571AFFC3DFA10" bold="true" box="[627,903,1470,1496]" pageId="33" pageNumber="34">
<collectingCountry id="92B67602C30FC751E25571AFFD49FA10" box="[627,755,1470,1496]" name="Mexico" pageId="33" pageNumber="34">MEXICO</collectingCountry>
: Veracruz:
</emphasis>
<quantity id="2D599B77C30FC751E3BF71AFFC57FA10" box="[921,1005,1470,1496]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="33" pageNumber="34" unit="km" value="2.3">2.3 km</quantity>
S Jalapa,
<quantity id="2D599B77C30FC751E44E71AFFB7CFA10" box="[1128,1222,1470,1496]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="33" pageNumber="34" unit="m" value="1320.0">1320 m</quantity>
, streamside litter (J.S.Ashe),
<date id="9E1F1052C30FC751E22071F7FD36F9C8" box="[518,652,1510,1536]" pageId="33" pageNumber="34" value="1992-07-13">13.vii.1992</date>
(
<collectionCode id="8CB0AE57C30FC751E2BD71F7FD49F9C8" box="[667,755,1510,1536]" pageId="33" pageNumber="34">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C30FC751E11E721FFB19F9E0" blockId="33.[264,1323,1470,1896]" box="[312,1187,1550,1576]" pageId="33" pageNumber="34">
<typeStatus id="351A8830C30FC751E11E721FFE14F9E0" box="[312,430,1550,1576]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C30FC751E1E6721FFD6AF9E0" bold="true" box="[448,720,1550,1576]" pageId="33" pageNumber="34">
<collectingCountry id="92B67602C30FC751E1E6721FFDFAF9E0" box="[448,576,1550,1576]" name="Mexico" pageId="33" pageNumber="34">MEXICO</collectingCountry>
: Veracruz:
</emphasis>
, same data as the
<typeStatus id="351A8830C30FC751E3EE721FFB94F9E0" box="[968,1070,1550,1576]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C30FC751E418721FFB2CF9E0" box="[1086,1174,1550,1576]" pageId="33" pageNumber="34">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30FC752E11E7227FD0AFE66" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="EA1E3692C30FC751E11E7227FD31F938" blockId="33.[264,1323,1470,1896]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E11E7227FE02F998" bold="true" box="[312,440,1590,1616]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C30FC751E1E87226FCA2F998" box="[462,792,1591,1616]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tuberculicauda">
<emphasis id="D8D5EA80C30FC751E1E87226FCA2F998" box="[462,792,1591,1616]" italics="true" pageId="33" pageNumber="34">Seeversiella tuberculicauda</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C30FC751E12E724EFE2CF9B0" box="[264,406,1631,1656]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C30FC751E12E724EFE2CF9B0" box="[264,406,1631,1656]" italics="true" pageId="33" pageNumber="34">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.2 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra longer than pronotum; wings fully developed; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C30FC751E49472BFFE8EF938" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="33" pageNumber="34">Figs. 110­ 115</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C30FC751E23E72C7FDC4F938" box="[536,638,1750,1776]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="33" pageNumber="34">Fig. 116</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30FC752E11E72EEFD0AFE66" blockId="33.[264,1323,1470,1896]" lastBlockId="34.[264,1323,284,430]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<taxonomicName id="2DA14D11C30FC751E11E72EEFDC7F8D0" box="[312,637,1791,1816]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tuberculicauda">
<emphasis id="D8D5EA80C30FC751E11E72EEFDC7F8D0" box="[312,637,1791,1816]" italics="true" pageId="33" pageNumber="34">Seeversiella tuberculicauda</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C30FC751E39E72EEFBEBF8D0" box="[952,1105,1791,1816]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C30FC751E39E72EEFBEBF8D0" box="[952,1105,1791,1816]" italics="true" pageId="33" pageNumber="34">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 5­6, 8, 10­12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have weak spicules (
<figureCitation id="729A2A17C30CC752E3BD750DFBB8FEFE" box="[923,1026,284,310]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Fig. 114</figureCitation>
); apex of median lobe in lateral view widened subapically and not emarginate on parameral side (
<figureCitation id="729A2A17C30CC752E47F7555FB45FE96" box="[1113,1279,324,350]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Figs. 112­113</figureCitation>
), in parameral view wide (
<figureCitation id="729A2A17C30CC752E23D757DFD7FFE4E" box="[539,709,364,390]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Figs. 110­111</figureCitation>
); spermatheca L­shaped with small umbilicus and straight proximal portion (
<figureCitation id="729A2A17C30CC752E21B7585FD19FE66" box="[573,675,404,430]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Fig. 116</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC30CC752E12E7155FC4EFA69" httpUri="https://zenodo.org/record/156439/files/figure.png" pageId="34" pageNumber="35" targetBox="[296,1289,499,1312]" targetPageId="34">
<paragraph id="EA1E3692C30CC752E12E7155FC4EFA69" blockId="34.[264,1323,1348,1441]" pageId="34" pageNumber="35">
<emphasis id="D8D5EA80C30CC752E12E7155FE5AFA94" bold="true" box="[264,480,1348,1372]" pageId="34" pageNumber="35">FIGURES 105­108</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C30CC752E3297154FB27FA94" authority="Gusarov" authorityName="Gusarov" box="[783,1181,1349,1372]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="tuberculicauda" status="sp. nov.">
<emphasis id="D8D5EA80C30CC752E3297154FB83FA94" box="[783,1081,1349,1372]" italics="true" pageId="34" pageNumber="35">Seeversiella tuberculicauda</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC30CC752E48C7155FB52FA94" box="[1194,1256,1348,1372]" pageId="34" pageNumber="35" rank="species">
<emphasis id="D8D5EA80C30CC752E48C7155FB52FA94" bold="true" box="[1194,1256,1348,1372]" pageId="34" pageNumber="35">sp. n.</emphasis>
</taxonomicNameLabel>
(105­ 106, holotype; 107­108, paratype from Jalapa, Mexico). 105 – male tergum 8; 106 – male sternum 8; 107 – female tergum 8; 108 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C30CC752E11E71E7FDC7F899" pageId="34" pageNumber="35" type="description">
<paragraph id="EA1E3692C30CC752E11E71E7FEE0F9F1" blockId="34.[264,1324,1526,1873]" pageId="34" pageNumber="35">
<emphasis id="D8D5EA80C30CC752E11E71E7FE6AF9D8" bold="true" box="[312,464,1526,1552]" pageId="34" pageNumber="35">Description.</emphasis>
Length
<quantity id="2D599B77C30CC752E26D71E6FD14F9D9" box="[587,686,1527,1553]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="34" pageNumber="35" unit="mm" value="2.4">2.4 mm</quantity>
, pronotal width
<quantity id="2D599B77C30CC752E35871E6FC4AF9D9" box="[894,1008,1527,1553]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="34" pageNumber="35" unit="mm" value="0.49">0.49 mm</quantity>
. Body dark brown, legs brown.
</paragraph>
<paragraph id="EA1E3692C30CC752E11E7256FB75F941" blockId="34.[264,1324,1526,1873]" pageId="34" pageNumber="35">
In all external character states
<taxonomicName id="2DA14D11C30CC752E28E7256FCC2F9A8" box="[680,888,1607,1632]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="tuberculicauda">
<emphasis id="D8D5EA80C30CC752E28E7256FCC2F9A8" box="[680,888,1607,1632]" italics="true" pageId="34" pageNumber="35">S. tuberculicauda</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C30CC752E47F7256FB4EF9A8" box="[1113,1268,1607,1632]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C30CC752E47F7256FB4EF9A8" box="[1113,1268,1607,1632]" italics="true" pageId="34" pageNumber="35">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C30CC752E11E7286FED4F8C9" blockId="34.[264,1324,1526,1873]" pageId="34" pageNumber="35">
In the only known male specimen the posterior angles of male tergum 3 projecting as spines twice as long as tergum 3 (measured medially), tergum
<quantity id="2D599B77C30CC752E3CE72AEFBAFF911" box="[1000,1045,1727,1753]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="34" pageNumber="35" unit="in" value="7.0">7 in</quantity>
basal third with medial tubercle.
</paragraph>
<paragraph id="EA1E3692C30CC752E11E731EFD23F8E1" blockId="34.[264,1324,1526,1873]" box="[312,665,1807,1833]" pageId="34" pageNumber="35">
Aedeagus as in
<figureCitation id="729A2A17C30CC752E1D6731EFD2FF8E1" box="[496,661,1807,1833]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Figs. 110­115</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C30CC752E11E7326FDC7F899" blockId="34.[264,1324,1526,1873]" box="[312,637,1847,1873]" pageId="34" pageNumber="35">
Spermatheca as in
<figureCitation id="729A2A17C30CC752E2327326FDC2F899" box="[532,632,1847,1873]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="34" pageNumber="35">Fig. 116</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30DC753E11E750AFE5DFE4E" pageId="35" pageNumber="36" type="distribution">
<paragraph id="EA1E3692C30DC753E11E750AFCD9FEFE" blockId="35.[264,1323,283,390]" box="[312,867,283,310]" pageId="35" pageNumber="36">
<emphasis id="D8D5EA80C30DC753E11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="35" pageNumber="36">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C30DC753E2A3750DFD65FEFE" box="[645,735,284,310]" name="Mexico" pageId="35" pageNumber="36">Mexico</collectingCountry>
(
<figureCitation id="729A2A17C30DC753E2C9750DFCEDFEFE" box="[751,855,284,310]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="35" pageNumber="36">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30DC753E11E7552FE5DFE4E" blockId="35.[264,1323,283,390]" pageId="35" pageNumber="36">
<emphasis id="D8D5EA80C30DC753E11E7552FDB0FE95" bold="true" box="[312,522,323,349]" pageId="35" pageNumber="36">Natural History.</emphasis>
<taxonomicName id="2DA14D11C30DC753E2077555FCD1FE95" box="[545,875,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="tuberculicauda">
<emphasis id="D8D5EA80C30DC753E2077555FCD1FE95" box="[545,875,324,349]" italics="true" pageId="35" pageNumber="36">Seeversiella tuberculicauda</emphasis>
</taxonomicName>
was collected in streamside litter at altitude of
<quantity id="2D599B77C30DC753E1A1757DFE5EFE4E" box="[391,484,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="35" pageNumber="36" unit="m" value="1320.0">1320 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

301
data/62/08/87/62088784C311C733E12E7780FE17FE4E.xml Normal file
View file

@ -0,0 +1,301 @@
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<paragraph id="EA1E3692C311C74FE12E7780FC6FFC63" blockId="63.[264,981,913,939]" box="[264,981,913,939]" pageId="63" pageNumber="64">
<heading id="B15681FEC311C74FE12E7780FC6FFC63" bold="true" box="[264,981,913,939]" fontSize="11" level="1" pageId="63" pageNumber="64" reason="1">
<key id="9E3061A3C311C74FE12E7780FC6FFC63" box="[264,981,913,939]" pageId="63" pageNumber="64">
<keyStep id="51552E37C311C74FE12E7780FC6FFC63" box="[264,981,913,939]" pageId="63" pageNumber="64">
<keyLead id="515095A7C311C74FE12E7780FC6FFC63" box="[264,981,913,939]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE12E7780FC6FFC63" bold="true" box="[264,981,913,939]" pageId="63" pageNumber="64">
18.
<taxonomicName id="2DA14D11C311C74FE1127780FD7DFC63" ID-CoL="6YBCS" authority="Gusarov" authorityName="Gusarov" box="[308,711,913,939]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="curtipennis" status="sp. nov.">
<emphasis id="D8D5EA80C311C74FE1127780FDEEFC63" bold="true" box="[308,596,913,939]" italics="true" pageId="63" pageNumber="64">Seeversiella curtipennis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC311C74FE2F57780FCAFFC63" box="[723,789,913,939]" pageId="63" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C311C74FE3037780FC77FC63" box="[805,973,913,939]" captionStart-0="FIGURES 247 ­ 250" captionStart-1="FIGURES 251 ­ 255" captionStart-2="FIGURES 256 ­ 264" captionStartId-0="63.[264,383,1778,1802]" captionStartId-1="64.[264,383,1694,1718]" captionStartId-2="66.[264,383,1579,1603]" captionTargetBox-0="[283,1297,1044,1745]" captionTargetBox-1="[284,1301,785,1656]" captionTargetBox-2="[278,1308,342,1490]" captionTargetId-0="figure@63.[271,1315,1033,1752]" captionTargetId-1="figure@64.[270,1316,769,1667]" captionTargetId-2="figure@66.[270,1318,331,1497]" captionTargetPageId-0="63" captionTargetPageId-1="64" captionTargetPageId-2="66" captionText-0="FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 – male tergum 8; 248 – male sternum 8; 249 – female tergum 8; 250 – female sternum 8. Scale bar 0.4 mm." captionText-1="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." captionText-2="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 – details of internal sac retracted into median lobe, abparameral view; 257 – details of internal sac retracted into median lobe, parameral view; 258 – medial lamellae, parameral view; 259 – everted internal sac, parameral view; 260 – apex of left paramere; 261 ­ 262 – details of internal sac retracted into median lobe, lateral view; 263 – right distal sclerite of internal sac; 264 – everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri-0="https://zenodo.org/record/156463/files/figure.png" httpUri-1="https://zenodo.org/record/156464/files/figure.png" httpUri-2="https://zenodo.org/record/156465/files/figure.png" pageId="63" pageNumber="64">Figs. 247­264</figureCitation>
)
</emphasis>
</keyLead>
</keyStep>
</key>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="EA1E3692C311C74FE12E72E3FD11F887" blockId="63.[264,1323,1778,1871]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE12E72E3FE65F8C2" bold="true" box="[264,479,1778,1802]" pageId="63" pageNumber="64">FIGURES 247­250</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C311C74FE32272E2FBDDF8C2" authority="Gusarov" authorityName="Gusarov" box="[772,1127,1779,1802]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="curtipennis" status="sp. nov.">
<emphasis id="D8D5EA80C311C74FE32272E2FBBEF8C2" box="[772,1028,1779,1802]" italics="true" pageId="63" pageNumber="64">Seeversiella curtipennis</emphasis>
Gusarov
</taxonomicName>
,
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<emphasis id="D8D5EA80C311C74FE45772E3FB17F8C2" bold="true" box="[1137,1197,1778,1802]" pageId="63" pageNumber="64">sp. n.</emphasis>
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(paratypes, Volcán Irazú, Costa Rica). 247 – male tergum 8; 248 – male sternum 8; 249 – female tergum 8; 250 – female sternum 8. Scale bar 0.4 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C36EC730E12E750AFC57FE1E" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="EA1E3692C36EC730E12E750AFB35FE96" blockId="64.[264,1324,283,670]" pageId="64" pageNumber="65">
<emphasis id="D8D5EA80C36EC730E12E750AFE07FEFD" bold="true" box="[264,445,283,309]" pageId="64" pageNumber="65">
<typeStatus id="351A8830C36EC730E12E750AFEF9FEFD" box="[264,323,283,309]" pageId="64" pageNumber="65">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C36EC730E1F7750DFDFBFEFE" box="[465,577,284,310]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C36EC730E254750AFC0EFEFD" bold="true" box="[626,948,283,309]" pageId="64" pageNumber="65">
<collectingCountry id="92B67602C36EC730E254750AFC97FEFD" box="[626,813,283,309]" name="Costa Rica" pageId="64" pageNumber="65">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
P. N. Volcán Irazú,
<geoCoordinate id="8F955055C36EC730E489750DFA96FEFE" box="[1199,1324,284,310]" direction="north" orientation="latitude" pageId="64" pageNumber="65" precision="15" value="9.974999">9°58'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C36EC730E12E7555FE23FE96" box="[264,409,324,350]" direction="west" orientation="longitude" pageId="64" pageNumber="65" precision="15" value="-83.85833">83°51'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C36EC730E1827555FDBAFE96" box="[420,512,324,350]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="64" pageNumber="65" unit="m" value="3300.0">3300 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C36EC730E3B87555FBA1FE96" box="[926,1051,324,350]" pageId="64" pageNumber="65" value="1997-06-18">18.vi.1997</date>
(
<collectionCode id="8CB0AE57C36EC730E40D7555FB39FE96" box="[1067,1155,324,350]" pageId="64" pageNumber="65">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C36EC730E11E757DFC57FE1E" blockId="64.[264,1324,283,670]" pageId="64" pageNumber="65">
<typeStatus id="351A8830C36EC730E11E757DFE14FE4E" box="[312,430,364,390]" pageId="64" pageNumber="65" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C36EC730E1E6757AFD47FE4D" bold="true" box="[448,765,363,389]" pageId="64" pageNumber="65">
<collectingCountry id="92B67602C36EC730E1E6757AFDC3FE4D" box="[448,633,363,389]" name="Costa Rica" pageId="64" pageNumber="65">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
<specimenCount id="FCA7FD1BC36EC730E32A757DFC17FE4E" box="[780,941,364,390]" pageId="64" pageNumber="65" type="generic">28 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C36EC730E459757DFB53FE4E" box="[1151,1257,364,390]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
;
<emphasis id="D8D5EA80C36EC730E4DD757AFEFDFE65" bold="true" pageId="64" pageNumber="65">San Jose:</emphasis>
<specimenCount id="FCA7FD1BC36EC730E17E7585FE56FE66" box="[344,492,404,430]" pageId="64" pageNumber="65" type="generic">4 specimens</specimenCount>
, Estación Cuerici,
<quantity id="2D599B77C36EC730E2EA7585FCA4FE66" box="[716,798,404,430]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="64" pageNumber="65" unit="km" value="4.6">4.6 km</quantity>
E Villa Mills,
<geoCoordinate id="8F955055C36EC730E3F67585FBF6FE66" box="[976,1100,404,430]" direction="north" orientation="latitude" pageId="64" pageNumber="65" precision="15" value="9.566667">9°34'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C36EC730E4727585FB5EFE66" box="[1108,1252,404,430]" direction="west" orientation="longitude" pageId="64" pageNumber="65" precision="15" value="-83.666664">83°40'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C36EC730E4D77585FE98FE1E" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="64" pageNumber="65" unit="m" value="2600.0">2600 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C36EC730E2E775ADFC84FE1E" box="[705,830,444,470]" pageId="64" pageNumber="65" value="1997-06-26">26.vi.1997</date>
(all –
<collectionCode id="8CB0AE57C36EC730E3AC75ADFC5BFE1E" box="[906,993,444,470]" pageId="64" pageNumber="65">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36EC731E11E75F2FC95FDEE" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="EA1E3692C36EC730E11E75F2FAB3FD56" blockId="64.[264,1324,283,670]" pageId="64" pageNumber="65">
<emphasis id="D8D5EA80C36EC730E11E75F2FE02FE35" bold="true" box="[312,440,483,509]" pageId="64" pageNumber="65">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C36EC730E1ED75F5FD5DFE35" box="[459,743,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36EC730E1ED75F5FD5DFE35" box="[459,743,484,509]" italics="true" pageId="64" pageNumber="65">Seeversiella curtipennis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C36EC730E4DE75F5FED7FDED" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C36EC730E4DE75F5FED7FDED" italics="true" pageId="64" pageNumber="65">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.9­2.0 times as long as eyes; elytra shorter than pronotum; reduced wings (shorter than elytra); by lacking the medial emargination at the posterior margin of the male tergum 7 (
<figureCitation id="729A2A17C36EC730E2F3764DFC86FDBE" box="[725,828,604,630]" captionStart="FIGURES 247 ­ 250" captionStartId="63.[264,383,1778,1802]" captionTargetBox="[283,1297,1044,1745]" captionTargetId="figure@63.[271,1315,1033,1752]" captionTargetPageId="63" captionText="FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 – male tergum 8; 248 – male sternum 8; 249 – female tergum 8; 250 – female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156463/files/figure.png" pageId="64" pageNumber="65">Fig. 247</figureCitation>
); tergum 8 with three pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C36EC730E2BD7695FCFBFD56" box="[667,833,644,670]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="64" pageNumber="65">Figs. 251­254</figureCitation>
,
<figureCitation id="729A2A17C36EC730E36B7695FC0BFD56" box="[845,945,644,670]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 – details of internal sac retracted into median lobe, abparameral view; 257 – details of internal sac retracted into median lobe, parameral view; 258 – medial lamellae, parameral view; 259 – everted internal sac, parameral view; 260 – apex of left paramere; 261 ­ 262 – details of internal sac retracted into median lobe, lateral view; 263 – right distal sclerite of internal sac; 264 – everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="64" pageNumber="65">256­264</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C36EC730E4B27695FB46FD56" box="[1172,1276,644,670]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="64" pageNumber="65">Fig. 255</figureCitation>
).
</paragraph>
<caption id="BEDE661AC36EC730E12E728FFD66F8D5" httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="64" pageNumber="65" targetBox="[284,1301,785,1656]" targetPageId="64">
<paragraph id="EA1E3692C36EC730E12E728FFD66F8D5" blockId="64.[264,1323,1694,1821]" pageId="64" pageNumber="65">
<emphasis id="D8D5EA80C36EC730E12E728FFE67F97E" bold="true" box="[264,477,1694,1718]" pageId="64" pageNumber="65">FIGURES 251­255</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C36EC730E253728FFC6CF97D" authority="Gusarov" authorityName="Gusarov" box="[629,982,1694,1717]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="curtipennis" status="sp. nov.">
<emphasis id="D8D5EA80C36EC730E253728FFCCEF97D" box="[629,884,1694,1717]" italics="true" pageId="64" pageNumber="65">Seeversiella curtipennis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC36EC730E3C6728FFBA1F97E" box="[992,1051,1694,1718]" pageId="64" pageNumber="65" rank="species">
<emphasis id="D8D5EA80C36EC730E3C6728FFBA1F97E" bold="true" box="[992,1051,1694,1718]" pageId="64" pageNumber="65">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254­255).
</paragraph>
</caption>
<paragraph id="EA1E3692C36FC731E11E750DFC95FDEE" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
<taxonomicName id="2DA14D11C36FC731E11E750DFDEAFEFD" box="[312,592,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36FC731E11E750DFDEAFEFD" box="[312,592,284,309]" italics="true" pageId="65" pageNumber="66">Seeversiella curtipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C36FC731E2CF750DFC35FEFD" box="[745,911,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C36FC731E2CF750DFC35FEFD" box="[745,911,284,309]" italics="true" pageId="65" pageNumber="66">S. scabricollis</emphasis>
</taxonomicName>
,
<taxonomicName id="2DA14D11C36FC731E3BB750DFBEDFEFD" box="[925,1111,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C36FC731E3BB750DFBEDFEFD" box="[925,1111,284,309]" italics="true" pageId="65" pageNumber="66">S. furcativentris</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C36FC731E4A9750DFAB7FEFD" box="[1167,1293,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C36FC731E4A9750DFAB7FEFD" box="[1167,1293,284,309]" italics="true" pageId="65" pageNumber="66">S. brunnea</emphasis>
</taxonomicName>
in having shorter elytra (shorter than pronotum); reduced wings (shorter than elytra); posterior margin of male tergum 8 without medial emargination (
<figureCitation id="729A2A17C36FC731E3FF757DFBFBFE4E" box="[985,1089,364,390]" captionStart="FIGURES 247 ­ 250" captionStartId="63.[264,383,1778,1802]" captionTargetBox="[283,1297,1044,1745]" captionTargetId="figure@63.[271,1315,1033,1752]" captionTargetPageId="63" captionText="FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 – male tergum 8; 248 – male sternum 8; 249 – female tergum 8; 250 – female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156463/files/figure.png" pageId="65" pageNumber="66">Fig. 247</figureCitation>
); broad apex of the median lobe (in parameral view) (
<figureCitation id="729A2A17C36FC731E2B57585FC80FE66" box="[659,826,404,430]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="65" pageNumber="66">Figs. 251­252</figureCitation>
; 194­196; 212­213; 239­240); long medial lamellae of the internal sac (
<figureCitation id="729A2A17C36FC731E27175ADFD45FE1E" box="[599,767,444,470]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 – details of internal sac retracted into median lobe, abparameral view; 257 – details of internal sac retracted into median lobe, parameral view; 258 – medial lamellae, parameral view; 259 – everted internal sac, parameral view; 260 – apex of left paramere; 261 ­ 262 – details of internal sac retracted into median lobe, lateral view; 263 – right distal sclerite of internal sac; 264 – everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="65" pageNumber="66">Figs. 258­259</figureCitation>
; 201) protruding apically when the internal sac is retracted (
<figureCitation id="729A2A17C36FC731E18475F5FDF5FE36" box="[418,591,484,510]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 – details of internal sac retracted into median lobe, abparameral view; 257 – details of internal sac retracted into median lobe, parameral view; 258 – medial lamellae, parameral view; 259 – everted internal sac, parameral view; 260 – apex of left paramere; 261 ­ 262 – details of internal sac retracted into median lobe, lateral view; 263 – right distal sclerite of internal sac; 264 – everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="65" pageNumber="66">Figs. 256­257</figureCitation>
); lateral diverticula without sclerotized denticles (
<figureCitation id="729A2A17C36FC731E49775F5FA9DFE36" box="[1201,1319,484,510]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 – details of internal sac retracted into median lobe, abparameral view; 257 – details of internal sac retracted into median lobe, parameral view; 258 – medial lamellae, parameral view; 259 – everted internal sac, parameral view; 260 – apex of left paramere; 261 ­ 262 – details of internal sac retracted into median lobe, lateral view; 263 – right distal sclerite of internal sac; 264 – everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="65" pageNumber="66">Figs. 259</figureCitation>
; 200); and longer spermatheca (
<figureCitation id="729A2A17C36FC731E252761DFD5FFDEE" box="[628,741,524,550]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="65" pageNumber="66">Figs. 255</figureCitation>
; 205).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36FC731E11E7625FDC4F9FE" pageId="65" pageNumber="66" type="description">
<paragraph id="EA1E3692C36FC731E11E7625FBDCFDBE" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
<taxonomicName id="2DA14D11C36FC731E11E7625FDEBFD85" box="[312,593,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36FC731E11E7625FDEBFD85" box="[312,593,564,589]" italics="true" pageId="65" pageNumber="66">Seeversiella curtipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C36FC731E2CB7625FC38FD85" box="[749,898,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="D8D5EA80C36FC731E2CB7625FC38FD85" box="[749,898,564,589]" italics="true" pageId="65" pageNumber="66">S. lativentris</emphasis>
</taxonomicName>
in having weaker basal constriction of the apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C36FC731E366764DFC5DFDBE" box="[832,999,604,630]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="65" pageNumber="66">Figs. 251­252</figureCitation>
; 269­270).
</paragraph>
<paragraph id="EA1E3692C36FC731E11E7692FCCBFD56" blockId="65.[264,1324,284,1870]" box="[312,881,643,670]" pageId="65" pageNumber="66">
<emphasis id="D8D5EA80C36FC731E11E7692FE6AFD55" bold="true" box="[312,464,643,669]" pageId="65" pageNumber="66">Description.</emphasis>
Length
<quantity id="2D599B77C36FC731E21F7695FD7EFD56" box="[569,708,644,670]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="65" pageNumber="66" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6­3.1 mm</quantity>
. Body brown.
</paragraph>
<paragraph id="EA1E3692C36FC731E11E76BDFD8BFCF6" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">Head surface matte, with fine isodiametric microsculpture, fine and slightly asperate punctation, distance between punctures equals 1­2 times their diameter. Temples 1.9­2.0 times as long as eyes. Antennal article 2 as long as 3, article 4 slightly transverse, 5­10 transverse (ratio 1.5­2.0).</paragraph>
<paragraph id="EA1E3692C36FC731E11E775DFEE8FBB6" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C36FC731E40D775DFB6CFCAE" box="[1067,1238,844,870]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="5.6" metricValueMin="5.0" pageId="65" pageNumber="66" unit="mm" value="0.53" valueMax="0.56" valueMin="0.5">0.50­0.56 mm</quantity>
, length
<quantity id="2D599B77C36FC731E12E7765FE0BFC46" box="[264,433,884,910]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="65" pageNumber="66" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
, width to length ratio 1.1, surface matte, with fine isodiametric microsculpture; punctation fine and weak, distance between punctures equals 1­2 times their diameter. Elytra wider and shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.2), 1.7 times wider than long, glossy, with fine and weak isodiametric microsculpture, and fine and slightly asperate punctation, distance between punctures equals 1­2 times their diameter. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C36FC731E11E709DFC16FAD6" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 2­4 times their diameter on terga 3­5 and 4­7 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C36FC731E2D47115FC1AFAD6" box="[754,928,1284,1310]" captionStart="FIGURES 247 ­ 250" captionStartId="63.[264,383,1778,1802]" captionTargetBox="[283,1297,1044,1745]" captionTargetId="figure@63.[271,1315,1033,1752]" captionTargetPageId="63" captionText="FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 – male tergum 8; 248 – male sternum 8; 249 – female tergum 8; 250 – female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156463/files/figure.png" pageId="65" pageNumber="66">Figs. 247, 249</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36FC731E11E713DFEFFFA76" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
In large males posterior angles of tergum 3 projecting as spines up to ½ as long as tergum 3 (measured medially); tergum 7 with medial carina along midline, the carina up to 2/3 as long as tergum length; posterior margin of tergum 8 without emargination (
<figureCitation id="729A2A17C36FC731E4DD716DFE82FA76" captionStart="FIGURES 247 ­ 250" captionStartId="63.[264,383,1778,1802]" captionTargetBox="[283,1297,1044,1745]" captionTargetId="figure@63.[271,1315,1033,1752]" captionTargetPageId="63" captionText="FIGURES 247 ­ 250. Abdominal segment 8 of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 247 – male tergum 8; 248 – male sternum 8; 249 – female tergum 8; 250 – female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156463/files/figure.png" pageId="65" pageNumber="66">Fig. 247</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36FC731E11E71DDFC19F9C6" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
Aedeagus as in
<figureCitation id="729A2A17C36FC731E1D971DDFD13FA2E" box="[511,681,1484,1510]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="65" pageNumber="66">Figs. 251­254</figureCitation>
,
<figureCitation id="729A2A17C36FC731E29F71DDFCA5FA2E" box="[697,799,1484,1510]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 – details of internal sac retracted into median lobe, abparameral view; 257 – details of internal sac retracted into median lobe, parameral view; 258 – medial lamellae, parameral view; 259 – everted internal sac, parameral view; 260 – apex of left paramere; 261 ­ 262 – details of internal sac retracted into median lobe, lateral view; 263 – right distal sclerite of internal sac; 264 – everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="65" pageNumber="66">256­264</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C36FC731E30971E5FC2CF9C6" box="[815,918,1524,1550]" captionStart="FIGURES 256 ­ 264" captionStartId="66.[264,383,1579,1603]" captionTargetBox="[278,1308,342,1490]" captionTargetId="figure@66.[270,1318,331,1497]" captionTargetPageId="66" captionText="FIGURES 256 ­ 264. Aedeagus of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 256 – details of internal sac retracted into median lobe, abparameral view; 257 – details of internal sac retracted into median lobe, parameral view; 258 – medial lamellae, parameral view; 259 – everted internal sac, parameral view; 260 – apex of left paramere; 261 ­ 262 – details of internal sac retracted into median lobe, lateral view; 263 – right distal sclerite of internal sac; 264 – everted internal sac, lateral view. Scale bar 0.2 mm (256 ­ 257, 259, 261 ­ 262, 264), 0.1 mm (258, 260, 263)." httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="65" pageNumber="66">Fig. 260</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C36FC731E11E720DFDC4F9FE" blockId="65.[264,1324,284,1870]" box="[312,638,1564,1590]" pageId="65" pageNumber="66">
Spermatheca as in
<figureCitation id="729A2A17C36FC731E233720DFDC1F9FE" box="[533,635,1564,1590]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="65" pageNumber="66">Fig. 255</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36FC732E11E7252FC0DF8FF" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="discussion">
<paragraph id="EA1E3692C36FC731E11E7252FB57F886" blockId="65.[264,1324,284,1870]" pageId="65" pageNumber="66">
<emphasis id="D8D5EA80C36FC731E11E7252FE79F995" bold="true" box="[312,451,1603,1629]" pageId="65" pageNumber="66">Discussion.</emphasis>
Volcán Irazú and Cerro Cuerici, the localities where
<taxonomicName id="2DA14D11C36FC731E4687255FB49F995" box="[1102,1267,1604,1629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36FC731E4687255FB49F995" box="[1102,1267,1604,1629]" italics="true" pageId="65" pageNumber="66">S. curtipennis</emphasis>
</taxonomicName>
was found, are separated by the distance of
<quantity id="2D599B77C36FC731E2FF727DFC9FF94E" box="[729,805,1644,1670]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="65" pageNumber="66" unit="km" value="50.0">50 km</quantity>
and the gap as low as
<quantity id="2D599B77C36FC731E415727DFB2AF94E" box="[1075,1168,1644,1670]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="65" pageNumber="66" unit="m" value="1400.0">1400 m</quantity>
. Considering the distribution of the other species of
<taxonomicName id="2DA14D11C36FC731E2FB7285FCD1F965" box="[733,875,1684,1709]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C36FC731E2FB7285FCD1F965" box="[733,875,1684,1709]" italics="true" pageId="65" pageNumber="66">Seeversiella</emphasis>
</taxonomicName>
in
<collectingCountry id="92B67602C36FC731E3B27285FBA3F966" box="[916,1049,1684,1710]" name="Costa Rica" pageId="65" pageNumber="66">Costa Rica</collectingCountry>
, and the fact that in
<taxonomicName id="2DA14D11C36FC731E5337285FE36F91D" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36FC731E5337285FE36F91D" italics="true" pageId="65" pageNumber="66">S. curtipennis</emphasis>
</taxonomicName>
the wings are reduced to vestiges that are shorter than elytra, it is clear that the two known populations of this species are isolated from each other. Despite this, both are identical in the structure of genitalia, including the internal sac. Apparently, the isolation of the two population is recent and has not allowed enough time for their divergence.
</paragraph>
<caption id="BEDE661AC36CC732E12E723AFE5EF8DB" httpUri="https://zenodo.org/record/156465/files/figure.png" pageId="66" pageNumber="67" targetBox="[278,1308,342,1490]" targetPageId="66">
<paragraph id="EA1E3692C36CC732E12E723AFE5EF8DB" blockId="66.[264,1323,1579,1847]" pageId="66" pageNumber="67">
<emphasis id="D8D5EA80C36CC732E12E723AFE59F98B" bold="true" box="[264,483,1579,1603]" pageId="66" pageNumber="67">FIGURES 256­264</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C36CC732E2BE723DFBB9F98B" authority="Gusarov" authorityName="Gusarov" box="[664,1027,1580,1603]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="curtipennis" status="sp. nov.">
<emphasis id="D8D5EA80C36CC732E2BE723DFC26F98B" box="[664,924,1580,1603]" italics="true" pageId="66" pageNumber="67">Seeversiella curtipennis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C36CC732E435723AFBEEF98B" bold="true" box="[1043,1108,1579,1603]" pageId="66" pageNumber="67">
<taxonomicNameLabel id="C3E657FBC36CC732E435723AFBEEF98B" box="[1043,1108,1579,1603]" pageId="66" pageNumber="67" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes, Volcán Irazú, Costa Rica). 256 – details of internal sac retracted into median lobe, abparameral view; 257 – details of internal sac retracted into median lobe, parameral view; 258 – medial lamellae, parameral view; 259 – everted internal sac, parameral view; 260 – apex of left paramere; 261­262 – details of internal sac retracted into median lobe, lateral view; 263 – right distal sclerite of internal sac; 264 – everted internal sac, lateral view. Scale bar 0.2 mm (256­257, 259, 261­262, 264), 0.1 mm (258, 260, 263).
</paragraph>
</caption>
<paragraph id="EA1E3692C36CC732E12E7331FC0DF8FF" blockId="66.[264,1323,1579,1847]" box="[264,951,1824,1847]" pageId="66" pageNumber="67">LD – lateral diverticulum of internal sac; ML – medial lamellae.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C36DC733E11E750AFE17FE4E" pageId="67" pageNumber="68" type="distribution">
<paragraph id="EA1E3692C36DC733E11E750AFE17FE4E" blockId="67.[264,1323,283,390]" pageId="67" pageNumber="68">
<emphasis id="D8D5EA80C36DC733E11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="67" pageNumber="68">Distribution.</emphasis>
Known from Volcán Irazú and Cerro Cuerici,
<collectingCountry id="92B67602C36DC733E427750DFB3BFEFE" box="[1025,1153,284,310]" name="Costa Rica" pageId="67" pageNumber="68">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C36DC733E4B7750DFB43FEFE" box="[1169,1273,284,310]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="67" pageNumber="68">Fig. 388</figureCitation>
).
<emphasis id="D8D5EA80C36DC733E11E7552FDBDFE95" bold="true" box="[312,519,323,349]" pageId="67" pageNumber="68">Natural History.</emphasis>
<taxonomicName id="2DA14D11C36DC733E23F7555FC8EFE95" box="[537,820,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C36DC733E23F7555FC8EFE95" box="[537,820,324,349]" italics="true" pageId="67" pageNumber="68">Seeversiella curtipennis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C36DC733E12E757DFE12FE4E" box="[264,424,364,390]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.3" metricValueMin="2.6" pageId="67" pageNumber="68" unit="m" value="2950.0" valueMax="3300.0" valueMin="2600.0">2600­3300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

269
data/62/08/87/62088784C313C74FE12E761AFE17FCF6.xml Normal file
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<heading id="B15681FEC313C74DE12E761AFC08FDED" bold="true" box="[264,946,523,550]" fontSize="11" level="1" pageId="61" pageNumber="62" reason="1">
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17.
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<emphasis id="D8D5EA80C313C74DE112761DFD88FDEE" bold="true" box="[308,562,524,550]" italics="true" pageId="61" pageNumber="62">Seeversiella brunnea</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC313C74DE297761AFD49FDED" box="[689,755,523,549]" pageId="61" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C313C74DE324761AFC10FDED" box="[770,938,523,549]" captionStart-0="FIGURES 235 ­ 238" captionStart-1="FIGURES 239 ­ 246" captionStartId-0="61.[264,382,1659,1683]" captionStartId-1="62.[264,383,1630,1654]" captionTargetBox-0="[313,1272,852,1625]" captionTargetBox-1="[313,1277,660,1592]" captionTargetId-0="figure@61.[304,1283,844,1632]" captionTargetId-1="figure@62.[302,1285,649,1603]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionText-0="FIGURES 235 ­ 238. Abdominal segment 8 of Seeversiella brunnea Gusarov, sp. n. (235 ­ 236, holotype; 237 ­ 238, paratype, P. N. Tapantí, Costa Rica). 235 – male tergum 8; 236 – male sternum 8; 237 – female tergum 8; 238 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 – median lobe, parameral view; 240 – apex of median lobe, parameral view; 241 – median lobe, lateral view; 242 – apex of median lobe, lateral view; 243 – details of internal sac retracted into median lobe, lateral view; 244 – apex of left paramere; 245 – everted internal sac, parameral view; 246 – spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri-0="https://zenodo.org/record/156461/files/figure.png" httpUri-1="https://zenodo.org/record/156462/files/figure.png" pageId="61" pageNumber="62">Figs. 235­246</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C313C74DE12E7647FE29FD21" pageId="61" pageNumber="62" type="materials_examined">
<paragraph id="EA1E3692C313C74DE12E7647FB3BFD51" blockId="61.[264,1323,598,745]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE12E7647FE78FDB8" bold="true" box="[264,450,598,624]" pageId="61" pageNumber="62">
<typeStatus id="351A8830C313C74DE12E7647FEF9FDB8" box="[264,323,598,624]" pageId="61" pageNumber="62">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C313C74DE1C67646FDEAFDB9" box="[480,592,599,625]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C313C74DE2AC7647FC61FDB8" bold="true" box="[650,987,598,624]" pageId="61" pageNumber="62">
<collectingCountry id="92B67602C313C74DE2AC7647FCF0FDB8" box="[650,842,598,624]" name="Costa Rica" pageId="61" pageNumber="62">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
P. N. Tapantí,
<geoCoordinate id="8F955055C313C74DE4897646FA91FDB9" box="[1199,1323,599,625]" direction="north" orientation="latitude" pageId="61" pageNumber="62" precision="15" value="9.75">9°45'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C313C74DE12E766EFE23FD51" box="[264,409,639,665]" direction="west" orientation="longitude" pageId="61" pageNumber="62" precision="15" value="-83.816666">83°49'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C313C74DE182766EFDBAFD51" box="[420,512,639,665]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="61" pageNumber="62" unit="m" value="1500.0">1500 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C313C74DE3B8766EFBB7FD51" box="[926,1037,639,665]" pageId="61" pageNumber="62" value="1997-06-04">4.vi.1997</date>
(
<collectionCode id="8CB0AE57C313C74DE43A766EFBCEFD51" box="[1052,1140,639,665]" pageId="61" pageNumber="62">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C313C74DE11E76B6FE29FD21" blockId="61.[264,1323,598,745]" pageId="61" pageNumber="62">
<typeStatus id="351A8830C313C74DE11E76B6FE15FD09" box="[312,431,679,705]" pageId="61" pageNumber="62" type="paratype">Paratypes</typeStatus>
:
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<collectingCountry id="92B67602C313C74DE1E576B7FDC7FD08" box="[451,637,678,704]" name="Costa Rica" pageId="61" pageNumber="62">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
,, ditto but,
<geoCoordinate id="8F955055C313C74DE3E976B6FBF0FD09" box="[975,1098,679,705]" direction="north" orientation="latitude" pageId="61" pageNumber="62" precision="15" value="9.741667">9°44'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C313C74DE47476B6FB59FD09" box="[1106,1251,679,705]" direction="west" orientation="longitude" pageId="61" pageNumber="62" precision="15" value="-83.808334">83°48'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C313C74DE4D676B6FEA5FD21" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="61" pageNumber="62" unit="m" value="1740.0">1740 m</quantity>
(
<collectionCode id="8CB0AE57C313C74DE10876DEFE3CFD21" box="[302,390,719,745]" pageId="61" pageNumber="62">KSEM</collectionCode>
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</paragraph>
</subSubSection>
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<paragraph id="EA1E3692C313C74DE12E726AFC4EF91F" blockId="61.[264,1323,1659,1751]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE12E726AFE53F95B" bold="true" box="[264,489,1659,1683]" pageId="61" pageNumber="62">FIGURES 235­238.</emphasis>
Abdominal segment 8 of
<taxonomicName id="2DA14D11C313C74DE338726AFBD2F95A" authority="Gusarov" authorityName="Gusarov" box="[798,1128,1659,1682]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="D8D5EA80C313C74DE338726AFBB8F95A" box="[798,1026,1659,1682]" italics="true" pageId="61" pageNumber="62">Seeversiella brunnea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C313C74DE451726AFB0DF95B" bold="true" box="[1143,1207,1659,1683]" pageId="61" pageNumber="62">
<taxonomicNameLabel id="C3E657FBC313C74DE451726AFB0DF95B" box="[1143,1207,1659,1683]" pageId="61" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(235­236, holotype; 237­238, paratype, P. N. Tapantí, Costa Rica). 235 – male tergum 8; 236 – male sternum 8; 237 – female tergum 8; 238 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C313C74EE11E733CFB24FE1E" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="diagnosis">
<paragraph id="EA1E3692C313C74EE11E733CFC51FE4E" blockId="61.[264,1323,1837,1904]" lastBlockId="62.[264,1323,284,590]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE11E733CFE0DF88F" bold="true" box="[312,439,1837,1863]" pageId="61" pageNumber="62">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C313C74DE1ED733FFD7EF88F" box="[459,708,1838,1863]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C313C74DE1ED733FFD7EF88F" box="[459,708,1838,1863]" italics="true" pageId="61" pageNumber="62">Seeversiella brunnea</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C313C74DE4F5733FFEFDF8A7" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C313C74DE4F5733FFEFDF8A7" italics="true" pageId="61" pageNumber="62">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 0.8­1.2 times as long as eyes; matte pronotum with strong microsculpture; elytra longer than pronotum; wings fully developed or partially reduced, 2­4 times as long as elytra; tergum 8 with three pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C310C74EE1CB757DFD29FE4E" box="[493,659,364,390]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 – median lobe, parameral view; 240 – apex of median lobe, parameral view; 241 – median lobe, lateral view; 242 – apex of median lobe, lateral view; 243 – details of internal sac retracted into median lobe, lateral view; 244 – apex of left paramere; 245 – everted internal sac, parameral view; 246 – spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Figs. 239­245</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C310C74EE351757DFC64FE4E" box="[887,990,364,390]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 – median lobe, parameral view; 240 – apex of median lobe, parameral view; 241 – median lobe, lateral view; 242 – apex of median lobe, lateral view; 243 – details of internal sac retracted into median lobe, lateral view; 244 – apex of left paramere; 245 – everted internal sac, parameral view; 246 – spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Fig. 246</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C310C74EE11E7585FB24FE1E" blockId="62.[264,1323,284,590]" pageId="62" pageNumber="63">
<taxonomicName id="2DA14D11C310C74EE11E7585FD8BFE65" box="[312,561,404,429]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C310C74EE11E7585FD8BFE65" box="[312,561,404,429]" italics="true" pageId="62" pageNumber="63">Seeversiella brunnea</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C310C74EE2F47585FCC0FE65" box="[722,890,404,429]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C310C74EE2F47585FCC0FE65" box="[722,890,404,429]" italics="true" pageId="62" pageNumber="63">S. scabricollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C310C74EE3917585FBCEFE65" box="[951,1140,404,429]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C310C74EE3917585FBCEFE65" box="[951,1140,404,429]" italics="true" pageId="62" pageNumber="63">S. furcativentris</emphasis>
</taxonomicName>
in having concave apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C310C74EE32075ADFC14FE1E" box="[774,942,444,470]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 – median lobe, parameral view; 240 – apex of median lobe, parameral view; 241 – median lobe, lateral view; 242 – apex of median lobe, lateral view; 243 – details of internal sac retracted into median lobe, lateral view; 244 – apex of left paramere; 245 – everted internal sac, parameral view; 246 – spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Figs. 241­242</figureCitation>
; 197­198; 214­215).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C310C74FE11E75F5FDC4FE1E" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="description">
<paragraph id="EA1E3692C310C74EE11E75F5FD68FD86" blockId="62.[264,1323,284,590]" pageId="62" pageNumber="63">
<taxonomicName id="2DA14D11C310C74EE11E75F5FD8BFE35" box="[312,561,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C310C74EE11E75F5FD8BFE35" box="[312,561,484,509]" italics="true" pageId="62" pageNumber="63">Seeversiella brunnea</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C310C74EE2F775F5FCC3FE35" box="[721,889,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C310C74EE2F775F5FCC3FE35" box="[721,889,484,509]" italics="true" pageId="62" pageNumber="63">S. brevipennis</emphasis>
</taxonomicName>
in having matte pronotum; pronotal punctation not asperate; elytra longer than pronotum; and concave apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C310C74EE18D7625FDE9FD86" box="[427,595,564,590]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 – median lobe, parameral view; 240 – apex of median lobe, parameral view; 241 – median lobe, lateral view; 242 – apex of median lobe, lateral view; 243 – details of internal sac retracted into median lobe, lateral view; 244 – apex of left paramere; 245 – everted internal sac, parameral view; 246 – spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63">Figs. 241­242</figureCitation>
; 228­229).
</paragraph>
<caption id="BEDE661AC310C74EE12E724FFD50F8EA" httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="62" pageNumber="63" targetBox="[313,1277,660,1592]" targetPageId="62">
<paragraph id="EA1E3692C310C74EE12E724FFD50F8EA" blockId="62.[264,1324,1630,1863]" pageId="62" pageNumber="63">
<emphasis id="D8D5EA80C310C74EE12E724FFE5AF9BE" bold="true" box="[264,480,1630,1654]" pageId="62" pageNumber="63">FIGURES 239­246</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C310C74EE2A4724FFC72F9BD" authority="Gusarov" authorityName="Gusarov" box="[642,968,1630,1653]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="D8D5EA80C310C74EE2A4724FFCDEF9BD" box="[642,868,1630,1653]" italics="true" pageId="62" pageNumber="63">Seeversiella brunnea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C310C74EE3F3724FFBA8F9BE" bold="true" box="[981,1042,1630,1654]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="C3E657FBC310C74EE3F3724FFBA8F9BE" box="[981,1042,1630,1654]" pageId="62" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(239­244, holotype; 245­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 – median lobe, parameral view; 240 – apex of median lobe, parameral view; 241 – median lobe, lateral view; 242 – apex of median lobe, lateral view; 243 – details of internal sac retracted into median lobe, lateral view; 244 – apex of left paramere; 245 – everted internal sac, parameral view; 246 – spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246).
</paragraph>
</caption>
<paragraph id="EA1E3692C310C74EE12E7321FE47F88F" blockId="62.[264,1324,1630,1863]" box="[264,509,1840,1863]" pageId="62" pageNumber="63">CB – connecting band.</paragraph>
<paragraph id="EA1E3692C311C74FE11E750AFB5BFEFE" blockId="63.[264,1323,283,830]" box="[312,1249,283,310]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="63" pageNumber="64">Description.</emphasis>
Length
<quantity id="2D599B77C311C74FE21F750DFD7EFEFE" box="[569,708,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="63" pageNumber="64" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5­2.8 mm</quantity>
. Pronotal width
<quantity id="2D599B77C311C74FE3AA750DFB8FFEFE" box="[908,1077,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.0" metricValueMin="4.4" pageId="63" pageNumber="64" unit="mm" value="0.47" valueMax="0.5" valueMin="0.44">0.44­0.50 mm</quantity>
. Body brown.
</paragraph>
<paragraph id="EA1E3692C311C74FE11E7555FBEFFE4E" blockId="63.[264,1323,283,830]" pageId="63" pageNumber="64">
In all character states
<taxonomicName id="2DA14D11C311C74FE2117555FD0CFE95" box="[567,694,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE2117555FD0CFE95" box="[567,694,324,349]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C311C74FE3AC7555FB8AFE95" box="[906,1072,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C311C74FE3AC7555FB8AFE95" box="[906,1072,324,349]" italics="true" pageId="63" pageNumber="64">S. scabricollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C311C74FE44F7555FA9EFE95" box="[1129,1316,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C311C74FE44F7555FA9EFE95" box="[1129,1316,324,349]" italics="true" pageId="63" pageNumber="64">S. furcativentris</emphasis>
</taxonomicName>
, but differs in the shape of the median lobe as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C311C74FE11E7585FD26FE66" blockId="63.[264,1323,283,830]" box="[312,668,404,430]" pageId="63" pageNumber="64">
Aedeagus as in
<figureCitation id="729A2A17C311C74FE1D77585FD2DFE66" box="[497,663,404,430]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 – median lobe, parameral view; 240 – apex of median lobe, parameral view; 241 – median lobe, lateral view; 242 – apex of median lobe, lateral view; 243 – details of internal sac retracted into median lobe, lateral view; 244 – apex of left paramere; 245 – everted internal sac, parameral view; 246 – spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="63" pageNumber="64">Figs. 239­245</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C311C74FE11E75ADFDC4FE1E" blockId="63.[264,1323,283,830]" box="[312,638,444,470]" pageId="63" pageNumber="64">
Spermatheca as in
<figureCitation id="729A2A17C311C74FE23375ADFDC1FE1E" box="[533,635,444,470]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 – median lobe, parameral view; 240 – apex of median lobe, parameral view; 241 – median lobe, lateral view; 242 – apex of median lobe, lateral view; 243 – details of internal sac retracted into median lobe, lateral view; 244 – apex of left paramere; 245 – everted internal sac, parameral view; 246 – spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="63" pageNumber="64">Fig. 246</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C311C74FE11E75F2FBBFFD0E" pageId="63" pageNumber="64" type="discussion">
<paragraph id="EA1E3692C311C74FE11E75F2FBBFFD0E" blockId="63.[264,1323,283,830]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E75F2FE79FE35" bold="true" box="[312,451,483,509]" pageId="63" pageNumber="64">Discussion.</emphasis>
Despite the similarity of
<taxonomicName id="2DA14D11C311C74FE2D775F5FCCAFE35" box="[753,880,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE2D775F5FCCAFE35" box="[753,880,484,509]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
to
<taxonomicName id="2DA14D11C311C74FE3B275F5FBF4FE35" box="[916,1102,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C311C74FE3B275F5FBF4FE35" box="[916,1102,484,509]" italics="true" pageId="63" pageNumber="64">S. furcativentris</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C311C74FE4A375F5FA91FE35" box="[1157,1323,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C311C74FE4A375F5FA91FE35" box="[1157,1323,484,509]" italics="true" pageId="63" pageNumber="64">S. scabricollis</emphasis>
</taxonomicName>
in external characters and the shape of genitalia, the difference between
<taxonomicName id="2DA14D11C311C74FE476761DFB75FDED" box="[1104,1231,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE476761DFB75FDED" box="[1104,1231,524,549]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
and the two species in the shape of the apex of median lobe (
<figureCitation id="729A2A17C311C74FE35B7625FB9FFD86" box="[893,1061,564,590]" captionStart="FIGURES 239 ­ 246" captionStartId="62.[264,383,1630,1654]" captionTargetBox="[313,1277,660,1592]" captionTargetId="figure@62.[302,1285,649,1603]" captionTargetPageId="62" captionText="FIGURES 239 ­ 246. Genitalia of Seeversiella brunnea Gusarov, sp. n. (239 ­ 244, holotype; 245 ­ 246, paratypes, P. N. Tapantí, Costa Rica). 239 – median lobe, parameral view; 240 – apex of median lobe, parameral view; 241 – median lobe, lateral view; 242 – apex of median lobe, lateral view; 243 – details of internal sac retracted into median lobe, lateral view; 244 – apex of left paramere; 245 – everted internal sac, parameral view; 246 – spermatheca. Scale bar 0.2 mm (239, 241, 243, 245), 0.1 mm (240, 242, 244, 246)." httpUri="https://zenodo.org/record/156462/files/figure.png" pageId="63" pageNumber="64">Figs. 239­242</figureCitation>
; 194­198; 212­215) is of the same scale as the difference between
<taxonomicName id="2DA14D11C311C74FE336764DFC0DFDBD" box="[784,951,604,629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C311C74FE336764DFC0DFDBD" box="[784,951,604,629]" italics="true" pageId="63" pageNumber="64">S. scabricollis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C311C74FE3D4764DFB17FDBD" box="[1010,1197,604,629]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C311C74FE3D4764DFB17FDBD" box="[1010,1197,604,629]" italics="true" pageId="63" pageNumber="64">S. furcativentris</emphasis>
</taxonomicName>
. Since no intermediate forms have been found,
<taxonomicName id="2DA14D11C311C74FE2E77695FCFBFD55" box="[705,833,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE2E77695FCFBFD55" box="[705,833,644,669]" italics="true" pageId="63" pageNumber="64">S. brunnea</emphasis>
</taxonomicName>
is considered to represent a separate species, possibly a local endemic of the area near Tapantí (
<figureCitation id="729A2A17C311C74FE3B776BDFC42FD0E" box="[913,1016,684,710]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="63" pageNumber="64">Fig. 388</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C311C74FE11E76C2FE17FCF6" pageId="63" pageNumber="64" type="distribution">
<paragraph id="EA1E3692C311C74FE11E76C2FB29FD26" blockId="63.[264,1323,283,830]" box="[312,1171,723,750]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E76C2FE62FD25" bold="true" box="[312,472,723,749]" pageId="63" pageNumber="64">Distribution.</emphasis>
Known from the area near Tapantí,
<collectingCountry id="92B67602C311C74FE3A976C5FBB5FD26" box="[911,1039,724,750]" name="Costa Rica" pageId="63" pageNumber="64">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C311C74FE43976C5FB3DFD26" box="[1055,1159,724,750]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="63" pageNumber="64">Fig. 388</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C311C74FE11E76EAFE17FCF6" blockId="63.[264,1323,283,830]" pageId="63" pageNumber="64">
<emphasis id="D8D5EA80C311C74FE11E76EAFDB0FCDD" bold="true" box="[312,522,763,789]" pageId="63" pageNumber="64">Natural History.</emphasis>
<taxonomicName id="2DA14D11C311C74FE20776EDFCA7FCDD" box="[545,797,764,789]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C311C74FE20776EDFCA7FCDD" box="[545,797,764,789]" italics="true" pageId="63" pageNumber="64">Seeversiella brunnea</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C311C74FE12E7735FE12FCF6" box="[264,424,804,830]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" metricValueMax="1.7399999999999998" metricValueMin="1.5" pageId="63" pageNumber="64" unit="m" value="1620.0" valueMax="1740.0" valueMin="1500.0">1500­1740 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="6690E386CD041993E6B5C8C93D5AE505" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C317C74DE12E72B7FED2FE66" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella brevipennis Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="62" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="58" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="2F04BCC81938C116CA5C4FDB180A847D">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58" type="key">
<paragraph id="EA1E3692C317C749E12E72B7FC62F908" blockId="57.[264,984,1702,1729]" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<heading id="B15681FEC317C749E12E72B7FC62F908" bold="true" box="[264,984,1702,1729]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<keyLead id="515095A7C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<emphasis id="D8D5EA80C317C749E12E72B7FC62F908" bold="true" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<keyStep id="51552E37C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
<key id="9E3061A3C317C749E12E72B7FC62F908" box="[264,984,1702,1729]" pageId="57" pageNumber="58">
16.
<taxonomicName id="2DA14D11C317C749E11272B6FD70F908" ID-CoL="6YBCM" authority="Gusarov" authorityName="Gusarov" box="[308,714,1702,1729]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C317C749E11272B6FDEDF909" bold="true" box="[308,599,1703,1729]" italics="true" pageId="57" pageNumber="58">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC317C749E2F072B7FCA3F908" box="[726,793,1702,1728]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C317C749E30E72B7FC6AF908" box="[808,976,1702,1728]" captionStart-0="FIGURES 222 ­ 225" captionStart-1="FIGURES 226 ­ 229" captionStart-2="FIGURES 230 ­ 234" captionStartId-0="58.[264,382,1219,1243]" captionStartId-1="59.[264,383,1174,1198]" captionStartId-2="60.[264,382,1729,1753]" captionTargetBox-0="[284,1303,333,1185]" captionTargetBox-1="[307,1283,459,1141]" captionTargetBox-2="[294,1292,824,1693]" captionTargetId-0="figure@58.[273,1314,320,1192]" captionTargetId-1="figure@59.[296,1291,449,1147]" captionTargetId-2="figure@60.[281,1305,809,1702]" captionTargetPageId-0="58" captionTargetPageId-1="59" captionTargetPageId-2="60" captionText-0="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 – male tergum 8; 223 – male sternum 8; 224 – female tergum 8; 225 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-2="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156458/files/figure.png" httpUri-1="https://zenodo.org/record/156459/files/figure.png" httpUri-2="https://zenodo.org/record/156460/files/figure.png" pageId="57" pageNumber="58">Figs. 222­234</figureCitation>
)
</key>
</keyStep>
</emphasis>
</keyLead>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C317C74AE12E72E0FB6CFA58" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="EA1E3692C317C749E12E72E0FCC8F894" blockId="57.[264,1323,1777,1884]" pageId="57" pageNumber="58">
<emphasis id="D8D5EA80C317C749E12E72E0FE04F8C3" bold="true" box="[264,446,1777,1803]" pageId="57" pageNumber="58">
<typeStatus id="351A8830C317C749E12E72E0FEF9F8C3" box="[264,323,1777,1803]" pageId="57" pageNumber="58">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C317C749E1F372E3FDFFF8C4" box="[469,581,1778,1804]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C317C749E25E72E0FC31F8C3" bold="true" box="[632,907,1777,1803]" pageId="57" pageNumber="58">
<collectingCountry id="92B67602C317C749E25E72E0FD47F8C3" box="[632,765,1777,1803]" name="Panama" pageId="57" pageNumber="58">PANAMA</collectingCountry>
: Chiriquí:
</emphasis>
<quantity id="2D599B77C317C749E3B972E3FC4EF8C4" box="[927,1012,1778,1804]" metricMagnitude="3" metricUnit="m" metricValue="5.9" pageId="57" pageNumber="58" unit="km" value="5.9">5.9 km</quantity>
N Cerro Punta, Par. Nac. Volcán Barú,
<geoCoordinate id="8F955055C317C749E181730BFE43F8FC" box="[423,505,1818,1844]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="925" value="8.866667">8°52'N</geoCoordinate>
[the latitude on the label (
<geoCoordinate id="8F955055C317C749E30D730BFC22F8FC" box="[811,920,1818,1844]" direction="north" orientation="latitude" pageId="57" pageNumber="58" precision="15" value="8.366667">8°22'0&quot;N</geoCoordinate>
) is incorrect]
<geoCoordinate id="8F955055C317C749E41A730BFB07F8FC" box="[1084,1213,1818,1844]" direction="west" orientation="longitude" pageId="57" pageNumber="58" precision="15" value="-82.566666">82°34'0&quot;W</geoCoordinate>
,
<quantity id="2D599B77C317C749E4EF730BFA9EF8FC" box="[1225,1316,1818,1844]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="57" pageNumber="58" unit="m" value="2150.0">2150 m</quantity>
, alder forest litter (R.Anderson),
<date id="9E1F1052C317C749E2A77353FD44F894" box="[641,766,1858,1884]" pageId="57" pageNumber="58" value="1995-06-14">14.vi.1995</date>
(
<collectionCode id="8CB0AE57C317C749E3287353FCDCF894" box="[782,870,1858,1884]" pageId="57" pageNumber="58">KSEM</collectionCode>
).
</paragraph>
<caption id="BEDE661AC314C74AE12E70D2FBEAFAE8" httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="58" pageNumber="59" targetBox="[284,1303,333,1185]" targetPageId="58">
<paragraph id="EA1E3692C314C74AE12E70D2FBEAFAE8" blockId="58.[264,1323,1219,1312]" pageId="58" pageNumber="59">
<emphasis id="D8D5EA80C314C74AE12E70D2FE5CFB13" bold="true" box="[264,486,1219,1243]" pageId="58" pageNumber="59">FIGURES 222­225.</emphasis>
Abdominal segment 8 of
<taxonomicName id="2DA14D11C314C74AE32C70D5FBC8FB12" authority="Gusarov" authorityName="Gusarov" box="[778,1138,1219,1243]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C314C74AE32C70D5FBB4FB13" box="[778,1038,1220,1243]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C314C74AE45B70D2FB00FB13" bold="true" box="[1149,1210,1219,1243]" pageId="58" pageNumber="59">
<taxonomicNameLabel id="C3E657FBC314C74AE45B70D2FB00FB13" box="[1149,1210,1219,1243]" pageId="58" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(222­223, holotype; 224­225, paratype, 5.9 km N Cerro Punta, Panama). 222 – male tergum 8; 223 – male sternum 8; 224 – female tergum 8; 225 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C314C74AE11E7167FB6CFA58" blockId="58.[264,1324,1397,1904]" box="[312,1238,1397,1424]" pageId="58" pageNumber="59">
<typeStatus id="351A8830C314C74AE11E7167FE14FA58" box="[312,430,1398,1424]" pageId="58" pageNumber="59" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C314C74AE1E67164FD74FA47" bold="true" box="[448,718,1397,1423]" pageId="58" pageNumber="59">
<collectingCountry id="92B67602C314C74AE1E67164FDFFFA47" box="[448,581,1397,1423]" name="Panama" pageId="58" pageNumber="59">PANAMA</collectingCountry>
: Chiriquí:
</emphasis>
,, same data as the
<typeStatus id="351A8830C314C74AE3DD7167FBD8FA58" box="[1019,1122,1398,1424]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C314C74AE4547167FB70FA58" box="[1138,1226,1398,1424]" pageId="58" pageNumber="59">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C314C74AE11E718CFCA0F930" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="EA1E3692C314C74AE11E718CFBF2F948" blockId="58.[264,1324,1397,1904]" pageId="58" pageNumber="59">
<emphasis id="D8D5EA80C314C74AE11E718CFE02FA7F" bold="true" box="[312,440,1437,1463]" pageId="58" pageNumber="59">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C314C74AE1EC718FFD50FA7F" box="[458,746,1438,1463]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C314C74AE1EC718FFD50FA7F" box="[458,746,1438,1463]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C314C74AE4DE718FFED7FA17" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C314C74AE4DE718FFED7FA17" italics="true" pageId="58" pageNumber="59">Seeversiella</emphasis>
</taxonomicName>
by having brown or reddish brown body with yellowish brown elytra, abdominal segments 3­4, legs and mouthparts; large eyes (temple length to eye length ratio 1.6­2.0); glossy pronotum with weak isodiametric microsculpture and asperate punctation; elytra as long as pronotum; wings partially reduced; tergum 8 with three pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C314C74AE26D7277FD4AF948" box="[587,752,1638,1664]" captionStart-0="FIGURES 226 ­ 229" captionStart-1="FIGURES 230 ­ 234" captionStartId-0="59.[264,383,1174,1198]" captionStartId-1="60.[264,382,1729,1753]" captionTargetBox-0="[307,1283,459,1141]" captionTargetBox-1="[294,1292,824,1693]" captionTargetId-0="figure@59.[296,1291,449,1147]" captionTargetId-1="figure@60.[281,1305,809,1702]" captionTargetPageId-0="59" captionTargetPageId-1="60" captionText-0="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-1="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156459/files/figure.png" httpUri-1="https://zenodo.org/record/156460/files/figure.png" pageId="58" pageNumber="59">Figs. 226­233</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C314C74AE3F27277FB86F948" box="[980,1084,1638,1664]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="58" pageNumber="59">Fig. 234</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C314C74AE11E729FFCA0F930" blockId="58.[264,1324,1397,1904]" pageId="58" pageNumber="59">
<taxonomicName id="2DA14D11C314C74AE11E729FFDEDF96F" box="[312,599,1678,1703]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C314C74AE11E729FFDEDF96F" box="[312,599,1678,1703]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C314C74AE2DF729FFC18F96F" box="[761,930,1678,1703]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C314C74AE2DF729FFC18F96F" box="[761,930,1678,1703]" italics="true" pageId="58" pageNumber="59">S. scabricollis</emphasis>
</taxonomicName>
in having glossy pronotum with asperate punctation; elytra as long as pronotum; apex of median lobe in parameral view constricted basally (
<figureCitation id="729A2A17C314C74AE1D572CFFD20F930" box="[499,666,1758,1784]" captionStart="FIGURES 226 ­ 229" captionStartId="59.[264,383,1174,1198]" captionTargetBox="[307,1283,459,1141]" captionTargetId="figure@59.[296,1291,449,1147]" captionTargetPageId="59" captionText="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="58" pageNumber="59">Figs. 226­227</figureCitation>
; 194­196).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C314C74CE11E7317FDC4FD0E" lastPageId="60" lastPageNumber="61" pageId="58" pageNumber="59" type="description">
<paragraph id="EA1E3692C314C74AE11E7317FE56F8B8" blockId="58.[264,1324,1397,1904]" pageId="58" pageNumber="59">
<taxonomicName id="2DA14D11C314C74AE11E7317FDEFF8D7" box="[312,597,1798,1823]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C314C74AE11E7317FDEFF8D7" box="[312,597,1798,1823]" italics="true" pageId="58" pageNumber="59">Seeversiella brevipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C314C74AE2D77317FC16F8D7" box="[753,940,1798,1823]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C314C74AE2D77317FC16F8D7" box="[753,940,1798,1823]" italics="true" pageId="58" pageNumber="59">S. furcativentris</emphasis>
</taxonomicName>
in having glossy pronotum with asperate punctation; elytra as long as pronotum; larger median lobe with longer apex (
<figureCitation id="729A2A17C314C74AE4D7733FFED7F8B8" captionStart="FIGURES 226 ­ 229" captionStartId="59.[264,383,1174,1198]" captionTargetBox="[307,1283,459,1141]" captionTargetId="figure@59.[296,1291,449,1147]" captionTargetPageId="59" captionText="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="58" pageNumber="59">Figs. 226­229</figureCitation>
; 212­215).
</paragraph>
<paragraph id="EA1E3692C315C74BE11E750DFD3BFE4E" blockId="59.[264,1323,284,390]" pageId="59" pageNumber="60">
<taxonomicName id="2DA14D11C315C74BE11E750DFDE1FEFD" box="[312,603,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C315C74BE11E750DFDE1FEFD" box="[312,603,284,309]" italics="true" pageId="59" pageNumber="60">Seeversiella brevipennis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C315C74BE32F750DFC35FEFD" box="[777,911,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C315C74BE32F750DFC35FEFD" box="[777,911,284,309]" italics="true" pageId="59" pageNumber="60">S. brunnea</emphasis>
</taxonomicName>
in having glossy pronotum with asperate punctation; elytra as long as pronotum; straight apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C315C74BE17D757DFDB8FE4E" box="[347,514,364,390]" captionStart="FIGURES 226 ­ 229" captionStartId="59.[264,383,1174,1198]" captionTargetBox="[307,1283,459,1141]" captionTargetId="figure@59.[296,1291,449,1147]" captionTargetPageId="59" captionText="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="59" pageNumber="60">Figs. 228­229</figureCitation>
; 241­242).
</paragraph>
<caption id="BEDE661AC315C74BE12E7087FEFAFADD" httpUri="https://zenodo.org/record/156459/files/figure.png" pageId="59" pageNumber="60" targetBox="[307,1283,459,1141]" targetPageId="59">
<paragraph id="EA1E3692C315C74BE12E7087FEFAFADD" blockId="59.[264,1323,1174,1301]" pageId="59" pageNumber="60">
<emphasis id="D8D5EA80C315C74BE12E7087FE59FB66" bold="true" box="[264,483,1174,1198]" pageId="59" pageNumber="60">FIGURES 226­229</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C315C74BE2B37087FBB9FB65" authority="Gusarov" authorityName="Gusarov" box="[661,1027,1174,1197]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C315C74BE2B37087FC26FB65" box="[661,924,1174,1197]" italics="true" pageId="59" pageNumber="60">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
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<emphasis id="D8D5EA80C315C74BE4347087FBE9FB66" bold="true" box="[1042,1107,1174,1198]" pageId="59" pageNumber="60">sp. n.</emphasis>
</taxonomicNameLabel>
(holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229).
</paragraph>
</caption>
<paragraph id="EA1E3692C315C74BE11E717AFC1BFA66" blockId="59.[264,1324,1387,1894]" pageId="59" pageNumber="60">
<emphasis id="D8D5EA80C315C74BE11E717AFE75FA4D" bold="true" box="[312,463,1387,1413]" pageId="59" pageNumber="60">Description.</emphasis>
Length 2.6­3.0 mm. Body brown or reddish brown, elytra, abdominal segments 3 and 4, legs and mouthparts yellowish brown.
</paragraph>
<paragraph id="EA1E3692C315C74BE11E71ADFE87F986" blockId="59.[264,1324,1387,1894]" pageId="59" pageNumber="60">Head surface glossy, with fine isodiametric microsculpture, fine and slightly asperate punctation, distance between punctures equals 2­3 times their diameter. Temple length to eye length ratio 1.6­2.0. Antennal article 2 as long as 3, articles 4­10 transverse (ratio 1.3­ 1.6).</paragraph>
<paragraph id="EA1E3692C315C74BE11E724DFB27F8AE" blockId="59.[264,1324,1387,1894]" pageId="59" pageNumber="60">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C315C74BE40D724DFB6CF9BE" box="[1067,1238,1628,1654]" metricMagnitude="-4" metricUnit="m" metricValue="5.45" metricValueMax="5.6" metricValueMin="5.3" pageId="59" pageNumber="60" unit="mm" value="0.545" valueMax="0.56" valueMin="0.53">0.53­0.56 mm</quantity>
, length
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, width to length ratio 1.1, surface glossy, with fine and weak isodiametric microsculpture; punctation fine and asperate, distance between punctures equal to 1­1.5 times their diameter. Elytra wider and as long (measured from humeral angle) as pronotum (pronotal length to elytral length ratio 1.0), 1.4 times wider than long, glossy, with fine and weak isodiametric microsculpture; with fine and asperate punctation, distance between punctures equal to 1­2 times their diameter. Wings partially reduced.
</paragraph>
<paragraph id="EA1E3692C312C74CE11E750DFC3CFE66" blockId="60.[264,1324,284,710]" pageId="60" pageNumber="61">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 3­6 times their diameter on terga 3­5 and 4­7 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C312C74CE2ED7585FCC3FE66" box="[715,889,404,430]" captionStart="FIGURES 222 ­ 225" captionStartId="58.[264,382,1219,1243]" captionTargetBox="[284,1303,333,1185]" captionTargetId="figure@58.[273,1314,320,1192]" captionTargetPageId="58" captionText="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 – male tergum 8; 223 – male sternum 8; 224 – female tergum 8; 225 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="60" pageNumber="61">Figs. 222, 224</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C312C74CE11E75ADFEFFFD86" blockId="60.[264,1324,284,710]" pageId="60" pageNumber="61">
In males posterior angles of tergum 3 projecting as spines up to 2 times as long as tergum 3 (measured medially); tergum
<quantity id="2D599B77C312C74CE2E775F5FD4AFE36" box="[705,752,484,510]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="60" pageNumber="61" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial tubercle; posterior margin of male tergum 8 with emargination delimited by short projections (
<figureCitation id="729A2A17C312C74CE4DD761DFE83FD86" captionStart="FIGURES 222 ­ 225" captionStartId="58.[264,382,1219,1243]" captionTargetBox="[284,1303,333,1185]" captionTargetId="figure@58.[273,1314,320,1192]" captionTargetPageId="58" captionText="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 – male tergum 8; 223 – male sternum 8; 224 – female tergum 8; 225 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="60" pageNumber="61">Fig. 222</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C312C74CE11E764DFD48FD56" blockId="60.[264,1324,284,710]" pageId="60" pageNumber="61">
Aedeagus as in
<figureCitation id="729A2A17C312C74CE1D4764DFD23FDBE" box="[498,665,604,630]" captionStart-0="FIGURES 226 ­ 229" captionStart-1="FIGURES 230 ­ 234" captionStartId-0="59.[264,383,1174,1198]" captionStartId-1="60.[264,382,1729,1753]" captionTargetBox-0="[307,1283,459,1141]" captionTargetBox-1="[294,1292,824,1693]" captionTargetId-0="figure@59.[296,1291,449,1147]" captionTargetId-1="figure@60.[281,1305,809,1702]" captionTargetPageId-0="59" captionTargetPageId-1="60" captionText-0="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-1="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156459/files/figure.png" httpUri-1="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61">Figs. 226­233</figureCitation>
. Apex of paramere broad, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C312C74CE2587695FD5FFD56" box="[638,741,644,670]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61">Fig. 233</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C312C74CE11E76BDFDC4FD0E" blockId="60.[264,1324,284,710]" box="[312,638,684,710]" pageId="60" pageNumber="61">
Spermatheca as in
<figureCitation id="729A2A17C312C74CE23276BDFDC0FD0E" box="[532,634,684,710]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61">Fig. 234</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="BEDE661AC312C74CE12E72D0FB63F888" httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="60" pageNumber="61" targetBox="[294,1292,824,1693]" targetPageId="60">
<paragraph id="EA1E3692C312C74CE12E72D0FB63F888" blockId="60.[264,1323,1729,1856]" pageId="60" pageNumber="61">
<emphasis id="D8D5EA80C312C74CE12E72D0FE67F911" bold="true" box="[264,477,1729,1753]" pageId="60" pageNumber="61">FIGURES 230­234</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C312C74CE25272D3FC6DF910" authority="Gusarov" authorityName="Gusarov" box="[628,983,1729,1753]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C312C74CE25272D3FCCCF911" box="[628,886,1730,1753]" italics="true" pageId="60" pageNumber="61">Seeversiella brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C312C74CE3C772D0FBA6F911" bold="true" box="[993,1052,1729,1753]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="C3E657FBC312C74CE3C772D0FBA6F911" box="[993,1052,1729,1753]" pageId="60" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(230­233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230­231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230­232), 0.1 mm (233­234).
</paragraph>
</caption>
<subSubSection id="A2BB6519C313C74DE11E750AFED2FE66" pageId="61" pageNumber="62" type="distribution">
<paragraph id="EA1E3692C313C74DE11E750AFD25FE96" blockId="61.[264,1323,283,430]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="61" pageNumber="62">Distribution.</emphasis>
Known from Cerro Punta,
<collectingCountry id="92B67602C313C74DE302750DFC3BFEFE" box="[804,897,284,310]" name="Panama" pageId="61" pageNumber="62">Panama</collectingCountry>
(
<figureCitation id="729A2A17C313C74DE3B4750DFC40FEFE" box="[914,1018,284,310]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="61" pageNumber="62">Fig. 388</figureCitation>
). In this locality
<taxonomicName id="2DA14D11C313C74DE4EE750DFEEFFE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C313C74DE4EE750DFEEFFE95" italics="true" pageId="61" pageNumber="62">S. brevipennis</emphasis>
</taxonomicName>
is sympatric
<taxonomicName id="2DA14D11C313C74DE1D47555FD22FE95" box="[498,664,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C313C74DE1D47555FD22FE95" box="[498,664,324,349]" italics="true" pageId="61" pageNumber="62">S. scabricollis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EA1E3692C313C74DE11E757AFED2FE66" blockId="61.[264,1323,283,430]" pageId="61" pageNumber="62">
<emphasis id="D8D5EA80C313C74DE11E757AFDBCFE4D" bold="true" box="[312,518,363,389]" pageId="61" pageNumber="62">Natural History.</emphasis>
<taxonomicName id="2DA14D11C313C74DE23E757DFC8DFE4D" box="[536,823,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C313C74DE23E757DFC8DFE4D" box="[536,823,364,389]" italics="true" pageId="61" pageNumber="62">Seeversiella brevipennis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
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.
</paragraph>
</subSubSection>
</treatment>
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489
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15.
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<emphasis id="D8D5EA80C318C746E11271B5FDD4FA76" bold="true" box="[308,622,1444,1470]" italics="true" pageId="54" pageNumber="55">Seeversiella furcativentris</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC318C746E2CB71B2FC8AFA75" box="[749,816,1443,1469]" pageId="54" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C318C746E31971B2FC5CFA75" box="[831,998,1443,1469]" captionStart-0="FIGURES 208 ­ 211" captionStart-1="FIGURES 212 ­ 221" captionStartId-0="56.[264,383,1366,1390]" captionStartId-1="57.[264,383,1351,1375]" captionTargetBox-0="[299,1287,536,1332]" captionTargetBox-1="[279,1264,462,1310]" captionTargetId-0="figure@56.[292,1294,529,1339]" captionTargetId-1="figure@57.[252,1336,433,1340]" captionTargetPageId-0="56" captionTargetPageId-1="57" captionText-0="FIGURES 208 ­ 211. Abdominal segment 8 of Seeversiella furcativentris Gusarov, sp. n. (208 ­ 209, holotype; 210 ­ 211, paratype, Zurquí de Moravia, Costa Rica). 208 – male tergum 8; 209 – male sternum 8; 210 – female tergum 8; 211 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 – median lobe, parameral view; 213 – apex of median lobe, parameral view; 214 – apex of median lobe, lateral view; 215 – median lobe, lateral view; 216 – apex of left paramere, 217 – details of internal sac retracted into median lobe, abparameral view; 218 – details of internal sac retracted into median lobe, lateral view; 219 – right distal sclerite of internal sac; 220 – left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 – spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri-0="https://zenodo.org/record/156456/files/figure.png" httpUri-1="https://zenodo.org/record/156457/files/figure.png" pageId="54" pageNumber="55">Figs. 208­221</figureCitation>
)
</keyLead>
</key>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C318C747E12E71FFFBD4FA8E" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="EA1E3692C318C746E12E71FFFCC8F9F8" blockId="54.[264,1325,1518,1904]" pageId="54" pageNumber="55">
<emphasis id="D8D5EA80C318C746E12E71FFFE7BF9C0" bold="true" box="[264,449,1518,1544]" pageId="54" pageNumber="55">
<typeStatus id="351A8830C318C746E12E71FFFEF9F9C0" box="[264,323,1518,1544]" pageId="54" pageNumber="55">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C318C746E1F871FEFDF4F9C1" box="[478,590,1519,1545]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C318C746E2AE71FFFC45F9C0" bold="true" box="[648,1023,1518,1544]" pageId="54" pageNumber="55">
<collectingCountry id="92B67602C318C746E2AE71FFFCFDF9C0" box="[648,839,1518,1544]" name="Costa Rica" pageId="54" pageNumber="55">COSTA RICA</collectingCountry>
: Puntarenas:
</emphasis>
Monte Verde,
<quantity id="2D599B77C318C746E4EA71FEFA91F9C1" box="[1228,1323,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="54" pageNumber="55" unit="m" value="1800.0">1800 m</quantity>
(J.Ashe, R.Brooks &amp; R.Leschen),
<date id="9E1F1052C318C746E2BF7207FD45F9F8" box="[665,767,1558,1584]" pageId="54" pageNumber="55" value="1989-05-05">5.v.1989</date>
(
<collectionCode id="8CB0AE57C318C746E3287207FCDCF9F8" box="[782,870,1558,1584]" pageId="54" pageNumber="55">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C318C747E11E722FFBD4FA8E" blockId="54.[264,1325,1518,1904]" lastBlockId="55.[264,1324,284,1830]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<typeStatus id="351A8830C318C746E11E722FFE14F990" box="[312,430,1598,1624]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C318C746E1E6722FFC9CF990" bold="true" box="[448,806,1598,1624]" pageId="54" pageNumber="55">
<collectingCountry id="92B67602C318C746E1E6722FFDC0F990" box="[448,634,1598,1624]" name="Costa Rica" pageId="54" pageNumber="55">COSTA RICA</collectingCountry>
: Puntarenas:
</emphasis>
<specimenCount id="FCA7FD1BC318C746E312722FFC7DF990" box="[820,967,1598,1624]" pageId="54" pageNumber="55" type="generic">5 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C318C746E4BF722FFAB9F990" box="[1177,1283,1598,1624]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
;, ditto but
<quantity id="2D599B77C318C746E15E7277FE6DF948" box="[376,471,1638,1664]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="54" pageNumber="55" unit="m" value="1580.0">1580 m</quantity>
, ex
<taxonomicName id="2DA14D11C318C746E22B7276FD2CF948" box="[525,662,1639,1664]" class="Agaricomycetes" family="Auriculariaceae" genus="Auricularia" kingdom="Animalia" order="Auriculariales" pageId="54" pageNumber="55" phylum="Basidiomycota" rank="genus">
<emphasis id="D8D5EA80C318C746E22B7276FD2CF948" box="[525,662,1639,1664]" italics="true" pageId="54" pageNumber="55">Auricularia</emphasis>
</taxonomicName>
sp.;, ditto but
<quantity id="2D599B77C318C746E34C7277FC73F948" box="[874,969,1638,1664]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="54" pageNumber="55" unit="m" value="1520.0">1520 m</quantity>
, flight intercept trap (J.Ashe, R.Brooks &amp; R.Leschen),
<date id="9E1F1052C318C746E21A729FFD08F960" box="[572,690,1678,1704]" pageId="54" pageNumber="55" value="1989-05-11">11.v.1989</date>
;
<specimenCount id="FCA7FD1BC318C746E2E4729FFCE2F960" box="[706,856,1678,1704]" pageId="54" pageNumber="55" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C318C746E3F1729FFBF4F960" box="[983,1102,1678,1704]" pageId="54" pageNumber="55" value="1989-05-14">14.v.1989</date>
;
<specimenCount id="FCA7FD1BC318C746E47B729FFB52F960" box="[1117,1256,1678,1704]" pageId="54" pageNumber="55" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C318C746E11072A7FE24F918" box="[310,414,1718,1744]" pageId="54" pageNumber="55" value="1989-05-09">9.v.1989</date>
;
<specimenCount id="FCA7FD1BC318C746E18D72A7FD8EF918" box="[427,564,1718,1744]" pageId="54" pageNumber="55" type="generic">1 specimen</specimenCount>
, ditto but pitfall trap,
<date id="9E1F1052C318C746E31F72A7FC15F918" box="[825,943,1718,1744]" pageId="54" pageNumber="55" value="1989-05-24">24.v.1989</date>
;
<specimenCount id="FCA7FD1BC318C746E39B72A7FBEBF918" box="[957,1105,1718,1744]" pageId="54" pageNumber="55" type="generic">2 specimens</specimenCount>
, ditto but
<quantity id="2D599B77C318C746E4EC72A7FA9DF918" box="[1226,1319,1718,1744]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="54" pageNumber="55" unit="m" value="1550.0">1550 m</quantity>
, flight intercept trap,
<date id="9E1F1052C318C746E1DC72CFFDCBF930" box="[506,625,1758,1784]" pageId="54" pageNumber="55" value="1989-05-23">23.v.1989</date>
;, ditto but
<quantity id="2D599B77C318C746E32F72CFFCDCF930" box="[777,870,1758,1784]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="54" pageNumber="55" unit="m" value="1570.0">1570 m</quantity>
, flight intercept trap,
<date id="9E1F1052C318C746E44572CFFB71F930" box="[1123,1227,1758,1784]" pageId="54" pageNumber="55" value="1989-05-09">9.v.1989</date>
;
<specimenCount id="FCA7FD1BC318C746E4F172CFFEFCF8E8" pageId="54" pageNumber="55" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C318C746E1EF7317FD85F8E8" box="[457,575,1798,1824]" pageId="54" pageNumber="55" value="1989-05-11">11.v.1989</date>
;, ditto but
<quantity id="2D599B77C318C746E2CC7317FCF1F8E8" box="[746,843,1798,1824]" metricMagnitude="3" metricUnit="m" metricValue="1.77" pageId="54" pageNumber="55" unit="m" value="1770.0">1770 m</quantity>
, ex
<taxonomicName id="2DA14D11C318C746E3A47316FC5DF8E8" box="[898,999,1799,1824]" class="Agaricomycetes" family="Tricholomataceae" genus="Collybia" kingdom="Fungi" order="Agaricales" pageId="54" pageNumber="55" phylum="Basidiomycota" rank="genus">
<emphasis id="D8D5EA80C318C746E3A47316FC5DF8E8" box="[898,999,1799,1824]" italics="true" pageId="54" pageNumber="55">Collybia</emphasis>
</taxonomicName>
sp.,
<date id="9E1F1052C318C746E40E7317FB25F8E8" box="[1064,1183,1798,1824]" pageId="54" pageNumber="55" value="1989-05-21">21.v.1989</date>
;, Monteverde reserve, trail near lab, flight intercept trap (C.Michalski),
<date id="9E1F1052C318C746E43A733FFB34F880" box="[1052,1166,1838,1864]" pageId="54" pageNumber="55" value="1993-06-03">3.vi.1993</date>
;
<specimenCount id="FCA7FD1BC318C746E4BB733FFA9DF880" box="[1181,1319,1838,1864]" pageId="54" pageNumber="55" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C318C746E15F7347FE51F8B8" box="[377,491,1878,1904]" pageId="54" pageNumber="55" value="1993-06-01">1.vi.1993</date>
;
<specimenCount id="FCA7FD1BC318C746E1DD7347FD28F8B8" box="[507,658,1878,1904]" pageId="54" pageNumber="55" type="generic">9 specimens</specimenCount>
, Monte Verde, Cerro Amigos,
<quantity id="2D599B77C318C746E42A7347FBD6F8B8" box="[1036,1132,1878,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="54" pageNumber="55" unit="m" value="1780.0">1780 m</quantity>
, flight intercept trap (J.Ashe, R.Brooks &amp; R.Leschen),
<date id="9E1F1052C319C747E2C3750DFCE6FEFE" box="[741,860,284,310]" pageId="55" pageNumber="56" value="1989-05-14">14.v.1989</date>
;
<specimenCount id="FCA7FD1BC319C747E34A750DFC43FEFE" box="[876,1017,284,310]" pageId="55" pageNumber="56" type="generic">1 specimen</specimenCount>
, ditto but pitfall trap;
<specimenCount id="FCA7FD1BC319C747E529750DFE3FFE96" pageId="55" pageNumber="56" type="generic">11 specimens</specimenCount>
, ditto but flight intercept trap,
<date id="9E1F1052C319C747E2D27555FCD1FE96" box="[756,875,324,350]" pageId="55" pageNumber="56" value="1989-05-21">21.v.1989</date>
;
<specimenCount id="FCA7FD1BC319C747E35E7555FBA6FE96" box="[888,1052,324,350]" pageId="55" pageNumber="56" type="generic">12 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E4B07555FAB6FE96" box="[1174,1292,324,350]" pageId="55" pageNumber="56" value="1989-05-24">24.v.1989</date>
;
<specimenCount id="FCA7FD1BC319C747E53C7555FE3FFE4E" pageId="55" pageNumber="56" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E1DB757DFDC9FE4E" box="[509,627,364,390]" pageId="55" pageNumber="56" value="1989-05-17">17.v.1989</date>
;
<emphasis id="D8D5EA80C319C747E2A7757AFC7FFE4D" bold="true" box="[641,965,363,389]" pageId="55" pageNumber="56">Puntarenas / Guanacaste:</emphasis>
, Monte Verde, Cerro Amigos,
<quantity id="2D599B77C319C747E1647585FE1AFE66" box="[322,416,404,430]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="55" pageNumber="56" unit="m" value="1780.0">1780 m</quantity>
, sifted leaf litter (J.Ashe, R.Brooks &amp; R.Leschen),
<date id="9E1F1052C319C747E42D7585FBC9FE66" box="[1035,1139,404,430]" pageId="55" pageNumber="56" value="1989-05-09">9.v.1989</date>
;
<specimenCount id="FCA7FD1BC319C747E4A47585FA9FFE66" box="[1154,1317,404,430]" pageId="55" pageNumber="56" type="generic">11 specimens</specimenCount>
, ditto but
<quantity id="2D599B77C319C747E15675ADFE71FE1E" box="[368,459,444,470]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="55" pageNumber="56" unit="m" value="1760.0">1760 m</quantity>
, but flight intercept trap,
<date id="9E1F1052C319C747E2D675ADFCDDFE1E" box="[752,871,444,470]" pageId="55" pageNumber="56" value="1989-05-12">12.v.1989</date>
;
<specimenCount id="FCA7FD1BC319C747E35475ADFBBFFE1E" box="[882,1029,444,470]" pageId="55" pageNumber="56" type="generic">6 specimens</specimenCount>
, ditto but pitfall traps;
<specimenCount id="FCA7FD1BC319C747E52D75ADFE3FFE36" pageId="55" pageNumber="56" type="generic">29 specimens</specimenCount>
, ditto but flight intercept trap,
<date id="9E1F1052C319C747E2CB75F5FCDEFE36" box="[749,868,484,510]" pageId="55" pageNumber="56" value="1989-05-10">10.v.1989</date>
;
<emphasis id="D8D5EA80C319C747E35775F2FBB7FE35" bold="true" box="[881,1037,483,509]" pageId="55" pageNumber="56">Guanacaste:</emphasis>
, Guanacaste Conservacion Area, Cacao Field Station,
<quantity id="2D599B77C319C747E2EF761DFC95FDEE" box="[713,815,524,550]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="55" pageNumber="56" unit="m" value="1400.0">1400 m</quantity>
, wet cloud forest litter (R.S.Anderson),
<date id="9E1F1052C319C747E12E7625FE39FD86" box="[264,387,564,590]" pageId="55" pageNumber="56" value="1996-02-12">12.ii.1996</date>
;
<emphasis id="D8D5EA80C319C747E1B57622FDB8FD85" bold="true" box="[403,514,563,589]" pageId="55" pageNumber="56">Heredia:</emphasis>
,, Cerro Chompipe,
<quantity id="2D599B77C319C747E3637625FC3FFD86" box="[837,901,564,590]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="56" unit="km" value="2.0">2 km</quantity>
N Monte de la Cruz,
<geoCoordinate id="8F955055C319C747E4847625FA96FD86" box="[1186,1324,564,590]" direction="north" orientation="latitude" pageId="55" pageNumber="56" precision="15" value="10.088888">10°05'20&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C319C747E12E764DFE23FDBE" box="[264,409,604,630]" direction="west" orientation="longitude" pageId="55" pageNumber="56" precision="15" value="-84.075">84°04'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C319C747E181764DFDBCFDBE" box="[423,518,604,630]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="56" unit="m" value="2000.0">2000 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C319C747E394764DFB89FDBE" box="[946,1075,604,630]" pageId="55" pageNumber="56" value="1997-06-12">12.vi.1997</date>
;
<specimenCount id="FCA7FD1BC319C747E464764DFB5DFDBE" box="[1090,1255,604,630]" pageId="55" pageNumber="56" type="generic">13 specimens</specimenCount>
, ditto but
<quantity id="2D599B77C319C747E11E7695FE37FD56" box="[312,397,644,670]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="55" pageNumber="56" unit="m" value="2100.0">2100m</quantity>
, flight intercept trap (S. &amp; J.Peck),
<date id="9E1F1052C319C747E36E7695FC55FD56" box="[840,1007,644,670]" pageId="55" pageNumber="56" value="1997-06-12" valueMax="1997-06-27" valueMin="1997-06-12">12­27.vi.1997</date>
;
<emphasis id="D8D5EA80C319C747E3D97692FBC3FD55" bold="true" box="[1023,1145,643,669]" pageId="55" pageNumber="56">San Jose:</emphasis>
<specimenCount id="FCA7FD1BC319C747E4A97695FA9CFD56" box="[1167,1318,644,670]" pageId="55" pageNumber="56" type="generic">5 specimens</specimenCount>
, Zurquí de Moravia [
<geoCoordinate id="8F955055C319C747E1D576BDFDECFD0E" box="[499,598,684,710]" direction="north" orientation="latitude" pageId="55" pageNumber="56" precision="925" value="10.05">10°03'N</geoCoordinate>
<geoCoordinate id="8F955055C319C747E27B76BDFD73FD0E" box="[605,713,684,710]" direction="west" orientation="longitude" pageId="55" pageNumber="56" precision="925" value="-84.01667">84°01'W</geoCoordinate>
],
<quantity id="2D599B77C319C747E2FA76BDFC8DFD0E" box="[732,823,684,710]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="55" pageNumber="56" unit="m" value="1600.0">1600 m</quantity>
, malaise traps (P.Hanson),
<date id="9E1F1052C319C747E45476BDFB76FD0E" box="[1138,1228,684,710]" pageId="55" pageNumber="56" value="1992-04">iv.1992</date>
;
<specimenCount id="FCA7FD1BC319C747E4FE76BDFEEBFD26" pageId="55" pageNumber="56" type="generic">7 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E1E076C5FDA3FD26" box="[454,537,724,750]" pageId="55" pageNumber="56" value="1992-05">v.1992</date>
;
<specimenCount id="FCA7FD1BC319C747E20276C5FD0DFD26" box="[548,695,724,750]" pageId="55" pageNumber="56" type="generic">5 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E30B76C5FC33FD26" box="[813,905,724,750]" pageId="55" pageNumber="56" value="1992-06">vi.1992</date>
;
<specimenCount id="FCA7FD1BC319C747E3B376C5FB92FD26" box="[917,1064,724,750]" pageId="55" pageNumber="56" type="generic">5 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E4BB76C5FAB8FD26" box="[1181,1282,724,750]" pageId="55" pageNumber="56" value="1992-07">vii.1992</date>
;
<specimenCount id="FCA7FD1BC319C747E52876C5FE3FFCDE" pageId="55" pageNumber="56" type="generic">10 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E1DA76EDFDE8FCDE" box="[508,594,764,790]" pageId="55" pageNumber="56" value="1993-02">ii.1993</date>
;
<specimenCount id="FCA7FD1BC319C747E27876EDFD48FCDE" box="[606,754,764,790]" pageId="55" pageNumber="56" type="generic">6 specimens</specimenCount>
, ditto but iii­iv.1993;
<specimenCount id="FCA7FD1BC319C747E3D776EDFB2EFCDE" box="[1009,1172,764,790]" pageId="55" pageNumber="56" type="generic">36 specimens</specimenCount>
, ditto but ivv.1993;
<specimenCount id="FCA7FD1BC319C747E1417735FE41FCF6" box="[359,507,804,830]" pageId="55" pageNumber="56" type="generic">7 specimens</specimenCount>
, ditto but ix­x.1993;
<specimenCount id="FCA7FD1BC319C747E2D37735FC33FCF6" box="[757,905,804,830]" pageId="55" pageNumber="56" type="generic">2 specimens</specimenCount>
, ditto but xi­xii.1993;
<specimenCount id="FCA7FD1BC319C747E4B57735FA9DFCF6" box="[1171,1319,804,830]" pageId="55" pageNumber="56" type="generic">5 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E150775DFE71FCAE" box="[374,459,844,870]" pageId="55" pageNumber="56" value="1994-02">ii.1994</date>
;
<specimenCount id="FCA7FD1BC319C747E1FF775DFDD4FCAE" box="[473,622,844,870]" pageId="55" pageNumber="56" type="generic">8 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E2CF775DFCFDFCAE" box="[745,839,844,870]" pageId="55" pageNumber="56" value="1994-03">iii.1994</date>
;
<specimenCount id="FCA7FD1BC319C747E373775DFC50FCAE" box="[853,1002,844,870]" pageId="55" pageNumber="56" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E443775DFB05FCAE" box="[1125,1215,844,870]" pageId="55" pageNumber="56" value="1994-04">iv.1994</date>
;
<specimenCount id="FCA7FD1BC319C747E4EB775DFEF3FC46" pageId="55" pageNumber="56" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E1F67765FD99FC46" box="[464,547,884,910]" pageId="55" pageNumber="56" value="1994-05">v.1994</date>
;
<specimenCount id="FCA7FD1BC319C747E2137765FD74FC46" box="[565,718,884,910]" pageId="55" pageNumber="56" type="generic">9 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E3737765FC5AFC46" box="[853,992,884,910]" pageId="55" pageNumber="56" value="1995-01-01" valueMax="1995-01-31" valueMin="1995-01-01">1­31.i.1995</date>
;
<specimenCount id="FCA7FD1BC319C747E3D47765FB31FC46" box="[1010,1163,884,910]" pageId="55" pageNumber="56" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E5357765FE3DFC7E" pageId="55" pageNumber="56" value="1995-04-01" valueMax="1995-04-30" valueMin="1995-04-01">1­ 30.iv.1995</date>
;
<specimenCount id="FCA7FD1BC319C747E1BE778DFD8AFC7E" box="[408,560,924,950]" pageId="55" pageNumber="56" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E292778DFCF7FC7E" box="[692,845,924,950]" pageId="55" pageNumber="56" value="1995-09-01" valueMax="1995-09-30" valueMin="1995-09-01">1­30.ix.1995</date>
;
<specimenCount id="FCA7FD1BC319C747E379778DFC4CFC7E" box="[863,1014,924,950]" pageId="55" pageNumber="56" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C319C747E45D778DFAB6FC7E" box="[1147,1292,924,950]" pageId="55" pageNumber="56" value="1995-10-01" valueMax="1995-10-30" valueMin="1995-10-01">1­30.x.1995</date>
;
<specimenCount id="FCA7FD1BC319C747E53B778DFEC0FC16" pageId="55" pageNumber="56" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C319C747E1DC77D5FD29FC16" box="[506,659,964,990]" pageId="55" pageNumber="56" value="1996-06-01" valueMax="1996-06-30" valueMin="1996-06-01">1­30.vi.1996</date>
;,
<quantity id="2D599B77C319C747E2EE77D5FCACFC16" box="[712,790,964,990]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="55" pageNumber="56" unit="km" value="26.0">26 km</quantity>
N San Isidro,
<quantity id="2D599B77C319C747E3EB77D5FB90FC16" box="[973,1066,964,990]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="55" pageNumber="56" unit="m" value="2100.0">2100 m</quantity>
(P.Hanson),
<date id="9E1F1052C319C747E4EE77D5FA98FC16" box="[1224,1314,964,990]" pageId="55" pageNumber="56" value="1992-11">xi.1992</date>
­
<date id="9E1F1052C319C747E12E77FDFEEFFBCE" box="[264,341,1004,1030]" pageId="55" pageNumber="56" value="1993-01">i.1993</date>
;,, San Antonio de Escazú,
<quantity id="2D599B77C319C747E2E177FDFC99FBCE" box="[711,803,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="55" pageNumber="56" unit="m" value="2000.0">2000 m</quantity>
, flight intercept trap (C.
<collectingCountry id="92B67602C319C747E41E77FDFB3CFBCE" box="[1080,1158,1004,1030]" name="Indonesia" pageId="55" pageNumber="56">Flores</collectingCountry>
&amp; P.Hanson),
<date id="9E1F1052C319C747E12E7005FE0DFBE6" box="[264,439,1044,1070]" pageId="55" pageNumber="56" value="1996-09-01" valueMax="1996-10-30" valueMin="1996-09-01">1.ix­30.x.1996</date>
;
<emphasis id="D8D5EA80C319C747E1E37002FD06FBE5" bold="true" box="[453,700,1043,1069]" pageId="55" pageNumber="56">San Jose / Cartago:</emphasis>
, km 45, Int. Amer. Hwy.,
<quantity id="2D599B77C319C747E43D7005FBE2FBE6" box="[1051,1112,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="55" pageNumber="56" unit="km" value="6.0">6 km</quantity>
NE El Empalme,
<geoCoordinate id="8F955055C319C747E12E702DFE3EFB9E" box="[264,388,1084,1110]" direction="north" orientation="latitude" pageId="55" pageNumber="56" precision="15" value="9.75">9°45'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C319C747E1AB702DFDA4FB9E" box="[397,542,1084,1110]" direction="west" orientation="longitude" pageId="55" pageNumber="56" precision="15" value="-83.975">83°58'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C319C747E20B702DFD31FB9E" box="[557,651,1084,1110]" metricMagnitude="3" metricUnit="m" metricValue="1.975" pageId="55" pageNumber="56" unit="m" value="1975.0">1975 m</quantity>
, flight intercept trap (S. &amp; J.Peck),
<date id="9E1F1052C319C747E419702DFB62FB9E" box="[1087,1240,1084,1110]" pageId="55" pageNumber="56" value="1997-06-08" valueMax="1997-06-26" valueMin="1997-06-08">8­26.vi.1997</date>
;
<emphasis id="D8D5EA80C319C747E4C0702AFE84FBB5" bold="true" pageId="55" pageNumber="56">Cartago:</emphasis>
10,
<specimenCount id="FCA7FD1BC319C747E1897075FDE8FBB6" box="[431,594,1124,1150]" pageId="55" pageNumber="56" type="generic">26 specimens</specimenCount>
, La Cangreja,
<quantity id="2D599B77C319C747E3247075FCDAFBB6" box="[770,864,1124,1150]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="55" pageNumber="56" unit="m" value="1950.0">1950 m</quantity>
, (P.Hanson), ix­xii.1992;
<specimenCount id="FCA7FD1BC319C747E4B57075FA9DFBB6" box="[1171,1319,1124,1150]" pageId="55" pageNumber="56" type="generic">2 specimens</specimenCount>
, ditto but iii­v.1992;
<specimenCount id="FCA7FD1BC319C747E1DC709DFD24FB6E" box="[506,670,1164,1190]" pageId="55" pageNumber="56" type="generic">17 specimens</specimenCount>
, ditto but vi­vii.1992; 2, ditto but
<quantity id="2D599B77C319C747E443709DFB79FB6E" box="[1125,1219,1164,1190]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="55" pageNumber="56" unit="m" value="1450.0">1450 m</quantity>
, malaise trap,
<date id="9E1F1052C319C747E16270A5FE4BFB06" box="[324,497,1204,1230]" pageId="55" pageNumber="56" value="1995-04-01" valueMax="1995-05-30" valueMin="1995-04-01">1.iv­30.v.1995</date>
;, Tapantí, (M.L.Jameson),
<date id="9E1F1052C319C747E36070A5FC7DFB06" box="[838,967,1204,1230]" pageId="55" pageNumber="56" value="1992-06-23">23.vi.1992</date>
;
<emphasis id="D8D5EA80C319C747E3F570A2FB5AFB05" bold="true" box="[979,1248,1203,1229]" pageId="55" pageNumber="56">
<collectingCountry id="92B67602C319C747E3F570A2FBE2FB05" box="[979,1112,1203,1229]" name="Panama" pageId="55" pageNumber="56">PANAMA</collectingCountry>
: Chiriquí:
</emphasis>
,,
<quantity id="2D599B77C319C747E12E70CDFEE1FB3E" box="[264,347,1244,1270]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="55" pageNumber="56" unit="km" value="5.6">5.6 km</quantity>
N Boquete, La Culebra Trail,
<geoCoordinate id="8F955055C319C747E2E870CDFCF3FB3E" box="[718,841,1244,1270]" direction="north" orientation="latitude" pageId="55" pageNumber="56" precision="15" value="8.823055">8°49'23&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C319C747E37570CDFC59FB3E" box="[851,995,1244,1270]" direction="west" orientation="longitude" pageId="55" pageNumber="56" precision="15" value="-82.42166">82°25'18&quot;W</geoCoordinate>
,
<quantity id="2D599B77C319C747E3D770CDFBF5FB3E" box="[1009,1103,1244,1270]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="55" pageNumber="56" unit="m" value="1800.0">1800 m</quantity>
, cloud forest litter (R.S.Anderson),
<date id="9E1F1052C319C747E1E97115FDEAFAD6" box="[463,592,1284,1310]" pageId="55" pageNumber="56" value="1996-06-15">15.vi.1996</date>
; 3, ditto but 6.0 km NE Boquete,
<geoCoordinate id="8F955055C319C747E3D97115FBEBFAD6" box="[1023,1105,1284,1310]" direction="north" orientation="latitude" pageId="55" pageNumber="56" precision="925" value="8.8">8°48'N</geoCoordinate>
<geoCoordinate id="8F955055C319C747E47E7115FB05FAD6" box="[1112,1215,1284,1310]" direction="west" orientation="longitude" pageId="55" pageNumber="56" precision="925" value="-82.433334">82°26'W</geoCoordinate>
,
<quantity id="2D599B77C319C747E4ED7115FA9DFAD6" box="[1227,1319,1284,1310]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="55" pageNumber="56" unit="m" value="1650.0">1650 m</quantity>
, flight intercept traps (J.S.Ashe &amp; R.Brooks),
<date id="9E1F1052C319C747E33D713DFC05FA8E" box="[795,959,1324,1350]" pageId="55" pageNumber="56" value="1996-06-14" valueMax="1996-06-19" valueMin="1996-06-14">14­19.vi.1996</date>
(all –
<collectionCode id="8CB0AE57C319C747E42D713DFBD8FA8E" box="[1035,1122,1324,1350]" pageId="55" pageNumber="56">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C319C747E11E7142FBF8F996" pageId="55" pageNumber="56" type="diagnosis">
<paragraph id="EA1E3692C319C747E11E7142FEFFF9C6" blockId="55.[264,1324,284,1830]" pageId="55" pageNumber="56">
<emphasis id="D8D5EA80C319C747E11E7142FE0DFAA5" bold="true" box="[312,439,1363,1389]" pageId="55" pageNumber="56">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C319C747E1E07145FD43FAA5" box="[454,761,1364,1389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C319C747E1E07145FD43FAA5" box="[454,761,1364,1389]" italics="true" pageId="55" pageNumber="56">Seeversiella furcativentris</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C319C747E4DE7145FED7FA5D" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C319C747E4DE7145FED7FA5D" italics="true" pageId="55" pageNumber="56">Seeversiella</emphasis>
</taxonomicName>
by brown body; large eyes (temples as long as eyes); matte pronotum with strong microsculpture; elytra longer than pronotum; wings fully developed or partially reduced, longer than elytra; the distinct shape of aedeagus (
<figureCitation id="729A2A17C319C747E34F71DDFBABFA2E" box="[873,1041,1484,1510]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 – median lobe, parameral view; 213 – apex of median lobe, parameral view; 214 – apex of median lobe, lateral view; 215 – median lobe, lateral view; 216 – apex of left paramere, 217 – details of internal sac retracted into median lobe, abparameral view; 218 – details of internal sac retracted into median lobe, lateral view; 219 – right distal sclerite of internal sac; 220 – left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 – spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="55" pageNumber="56">Figs. 212­220</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C319C747E4DA71DDFE82F9C6" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 – median lobe, parameral view; 213 – apex of median lobe, parameral view; 214 – apex of median lobe, lateral view; 215 – median lobe, lateral view; 216 – apex of left paramere, 217 – details of internal sac retracted into median lobe, abparameral view; 218 – details of internal sac retracted into median lobe, lateral view; 219 – right distal sclerite of internal sac; 220 – left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 – spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="55" pageNumber="56">Fig. 221</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C319C747E11E720DFBF8F996" blockId="55.[264,1324,284,1830]" pageId="55" pageNumber="56">
<taxonomicName id="2DA14D11C319C747E11E720DFDD7F9FD" box="[312,621,1564,1589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C319C747E11E720DFDD7F9FD" box="[312,621,1564,1589]" italics="true" pageId="55" pageNumber="56">Seeversiella furcativentris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C319C747E337720DFC01F9FD" box="[785,955,1564,1589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C319C747E337720DFC01F9FD" box="[785,955,1564,1589]" italics="true" pageId="55" pageNumber="56">S. scabricollis</emphasis>
</taxonomicName>
in having the apex of median lobe constricted basally (in parameral view) (
<figureCitation id="729A2A17C319C747E33D7255FC79F996" box="[795,963,1604,1630]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 – median lobe, parameral view; 213 – apex of median lobe, parameral view; 214 – apex of median lobe, lateral view; 215 – median lobe, lateral view; 216 – apex of left paramere, 217 – details of internal sac retracted into median lobe, abparameral view; 218 – details of internal sac retracted into median lobe, lateral view; 219 – right distal sclerite of internal sac; 220 – left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 – spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="55" pageNumber="56">Figs. 212­213</figureCitation>
; 194­196).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C319C748E11E727DFDC7F983" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="description">
<paragraph id="EA1E3692C319C747E11E727DFC89F966" blockId="55.[264,1324,284,1830]" pageId="55" pageNumber="56">
<taxonomicName id="2DA14D11C319C747E11E727DFDD6F94D" box="[312,620,1644,1669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C319C747E11E727DFDD6F94D" box="[312,620,1644,1669]" italics="true" pageId="55" pageNumber="56">Seeversiella furcativentris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C319C747E32B727DFC34F94D" box="[781,910,1644,1669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C319C747E32B727DFC34F94D" box="[781,910,1644,1669]" italics="true" pageId="55" pageNumber="56">S. brunnea</emphasis>
</taxonomicName>
in having straight apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C319C747E22B7285FD0EF966" box="[525,692,1684,1710]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 – median lobe, parameral view; 213 – apex of median lobe, parameral view; 214 – apex of median lobe, lateral view; 215 – median lobe, lateral view; 216 – apex of left paramere, 217 – details of internal sac retracted into median lobe, abparameral view; 218 – details of internal sac retracted into median lobe, lateral view; 219 – right distal sclerite of internal sac; 220 – left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 – spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="55" pageNumber="56">Figs. 214­215</figureCitation>
; 241­242).
</paragraph>
<paragraph id="EA1E3692C319C747E11E72ADFCB0F8EE" blockId="55.[264,1324,284,1830]" pageId="55" pageNumber="56">
<taxonomicName id="2DA14D11C319C747E11E72ADFDCBF91D" box="[312,625,1724,1749]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C319C747E11E72ADFDCBF91D" box="[312,625,1724,1749]" italics="true" pageId="55" pageNumber="56">Seeversiella furcativentris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C319C747E30472ADFC75F91D" box="[802,975,1724,1749]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C319C747E30472ADFC75F91D" box="[802,975,1724,1749]" italics="true" pageId="55" pageNumber="56">S. brevipennis</emphasis>
</taxonomicName>
in having matte pronotum; pronotal punctation not asperate; elytra longer than pronotum; and smaller median lobe with shorter apex (
<figureCitation id="729A2A17C319C747E1C5731DFD31F8EE" box="[483,651,1804,1830]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 – median lobe, parameral view; 213 – apex of median lobe, parameral view; 214 – apex of median lobe, lateral view; 215 – median lobe, lateral view; 216 – apex of left paramere, 217 – details of internal sac retracted into median lobe, abparameral view; 218 – details of internal sac retracted into median lobe, lateral view; 219 – right distal sclerite of internal sac; 220 – left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 – spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="55" pageNumber="56">Figs. 212­215</figureCitation>
; 226­229).
</paragraph>
<paragraph id="EA1E3692C316C748E11E750AFCABFE96" blockId="56.[264,1323,283,430]" pageId="56" pageNumber="57">
<emphasis id="D8D5EA80C316C748E11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="56" pageNumber="57">Description.</emphasis>
Length
<quantity id="2D599B77C316C748E219750DFD76FEFE" box="[575,716,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="56" pageNumber="57" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C316C748E3B7750DFB86FEFE" box="[913,1084,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="56" pageNumber="57" unit="mm" value="0.42" valueMax="0.44" valueMin="0.4">0.40­0.44 mm</quantity>
. Body brown, with darker head and lighter legs and mouthparts.
</paragraph>
<paragraph id="EA1E3692C316C748E11E757DFC0DFE66" blockId="56.[264,1323,283,430]" pageId="56" pageNumber="57">
In all character states
<taxonomicName id="2DA14D11C316C748E219757DFD41FE4D" box="[575,763,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C316C748E219757DFD41FE4D" box="[575,763,364,389]" italics="true" pageId="56" pageNumber="57">S. furcativentris</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C316C748E3FE757DFB3AFE4D" box="[984,1152,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C316C748E3FE757DFB3AFE4D" box="[984,1152,364,389]" italics="true" pageId="56" pageNumber="57">S. scabricollis</emphasis>
</taxonomicName>
, but differs in the shape of the median lobe as described in the diagnosis.
</paragraph>
<caption id="BEDE661AC316C748E12E7147FBF5FA7A" httpUri="https://zenodo.org/record/156456/files/figure.png" pageId="56" pageNumber="57" targetBox="[299,1287,536,1332]" targetPageId="56">
<paragraph id="EA1E3692C316C748E12E7147FBF5FA7A" blockId="56.[264,1325,1366,1458]" pageId="56" pageNumber="57">
<emphasis id="D8D5EA80C316C748E12E7147FE5EFAA6" bold="true" box="[264,484,1366,1390]" pageId="56" pageNumber="57">FIGURES 208­211.</emphasis>
Abdominal segment 8 of
<taxonomicName id="2DA14D11C316C748E2D87147FBCCFAA5" authority="Gusarov" authorityName="Gusarov" box="[766,1142,1366,1389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="furcativentris" status="sp. nov.">
<emphasis id="D8D5EA80C316C748E2D87147FBAEFAA5" box="[766,1044,1366,1389]" italics="true" pageId="56" pageNumber="57">Seeversiella furcativentris</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C316C748E4A67147FB01FAA6" bold="true" box="[1152,1211,1366,1390]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="C3E657FBC316C748E4A67147FB01FAA6" box="[1152,1211,1366,1390]" pageId="56" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(208­209, holotype; 210­211, paratype, Zurquí de Moravia, Costa Rica). 208 – male tergum 8; 209 – male sternum 8; 210 – female tergum 8; 211 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C316C748E11E7218FD21F9EB" blockId="56.[264,1324,1545,1891]" box="[312,667,1545,1571]" pageId="56" pageNumber="57">
Aedeagus as in
<figureCitation id="729A2A17C316C748E1D67218FD2DF9EB" box="[496,663,1545,1571]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 – median lobe, parameral view; 213 – apex of median lobe, parameral view; 214 – apex of median lobe, lateral view; 215 – median lobe, lateral view; 216 – apex of left paramere, 217 – details of internal sac retracted into median lobe, abparameral view; 218 – details of internal sac retracted into median lobe, lateral view; 219 – right distal sclerite of internal sac; 220 – left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 – spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="56" pageNumber="57">Figs. 212­220</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C316C748E11E7220FDC7F983" blockId="56.[264,1324,1545,1891]" box="[312,637,1585,1611]" pageId="56" pageNumber="57">
Spermatheca as in
<figureCitation id="729A2A17C316C748E2327220FDC3F983" box="[532,633,1585,1611]" captionStart="FIGURES 212 ­ 221" captionStartId="57.[264,383,1351,1375]" captionTargetBox="[279,1264,462,1310]" captionTargetId="figure@57.[252,1336,433,1340]" captionTargetPageId="57" captionText="FIGURES 212 ­ 221. Genitalia of Seeversiella furcativentris Gusarov, sp. n. (212 ­ 218, 221, holotype; 219 ­ 220, paratype from Monte Verde, Costa Rica). 212 – median lobe, parameral view; 213 – apex of median lobe, parameral view; 214 – apex of median lobe, lateral view; 215 – median lobe, lateral view; 216 – apex of left paramere, 217 – details of internal sac retracted into median lobe, abparameral view; 218 – details of internal sac retracted into median lobe, lateral view; 219 – right distal sclerite of internal sac; 220 – left lateral diverticulum of internal sac, with denticulate sclerotized portion; 221 – spermatheca. Scale bar 0.2 mm (212, 215, 217 ­ 218), 0.1 mm (213 ­ 214, 216, 219 ­ 221)." httpUri="https://zenodo.org/record/156457/files/figure.png" pageId="56" pageNumber="57">Fig. 221</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C316C748E11E7249FD88F8AB" pageId="56" pageNumber="57" type="discussion">
<paragraph id="EA1E3692C316C748E11E7249FD88F8AB" blockId="56.[264,1324,1545,1891]" pageId="56" pageNumber="57">
<emphasis id="D8D5EA80C316C748E11E7249FE79F9BA" bold="true" box="[312,451,1624,1650]" pageId="56" pageNumber="57">Discussion.</emphasis>
Despite the fact that
<taxonomicName id="2DA14D11C316C748E2ED7248FC3CF9BA" box="[715,902,1625,1650]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C316C748E2ED7248FC3CF9BA" box="[715,902,1625,1650]" italics="true" pageId="56" pageNumber="57">S. furcativentris</emphasis>
</taxonomicName>
and
<taxonomicName id="2DA14D11C316C748E3E77248FBD2F9BA" box="[961,1128,1625,1650]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C316C748E3E77248FBD2F9BA" box="[961,1128,1625,1650]" italics="true" pageId="56" pageNumber="57">S. scabricollis</emphasis>
</taxonomicName>
are very similar in their genitalia, the difference in the shape of the apex of median lobe is stable and no intermediate forms have been observed. In one locality both species have been collected in the same sample, which also confirms their status of separate species. The two species appear to be parapatric:
<taxonomicName id="2DA14D11C316C748E20E72E8FD5BF8DA" box="[552,737,1785,1810]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C316C748E20E72E8FD5BF8DA" box="[552,737,1785,1810]" italics="true" pageId="56" pageNumber="57">S. furcativentris</emphasis>
</taxonomicName>
occurs mostly below
<quantity id="2D599B77C316C748E3C672E8FB83F8DB" box="[992,1081,1785,1811]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="56" pageNumber="57" unit="m" value="2000.0">2000 m</quantity>
(has never been collected above
<quantity id="2D599B77C316C748E1877330FE47F8F3" box="[417,509,1825,1851]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="56" pageNumber="57" unit="m" value="2100.0">2100 m</quantity>
), while
<taxonomicName id="2DA14D11C316C748E27F7330FD45F8F2" box="[601,767,1825,1850]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C316C748E27F7330FD45F8F2" box="[601,767,1825,1850]" italics="true" pageId="56" pageNumber="57">S. scabricollis</emphasis>
</taxonomicName>
occurs mostly above
<quantity id="2D599B77C316C748E3DA7330FBEEF8F3" box="[1020,1108,1825,1851]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="56" pageNumber="57" unit="m" value="2000.0">2000 m</quantity>
(occasionally collected as low as
<quantity id="2D599B77C316C748E1EE7358FD9FF8AB" box="[456,549,1865,1891]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="56" pageNumber="57" unit="m" value="1450.0">1450 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C317C749E11E750AFC59FE4E" pageId="57" pageNumber="58" type="distribution">
<paragraph id="EA1E3692C317C749E11E750AFBA5FEFE" blockId="57.[264,1323,283,390]" box="[312,1055,283,310]" pageId="57" pageNumber="58">
<emphasis id="D8D5EA80C317C749E11E750AFE62FEFD" bold="true" box="[312,472,283,309]" pageId="57" pageNumber="58">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C317C749E2A3750DFCBFFEFE" box="[645,773,284,310]" name="Costa Rica" pageId="57" pageNumber="58">Costa Rica</collectingCountry>
and
<collectingCountry id="92B67602C317C749E318750DFC21FEFE" box="[830,923,284,310]" name="Panama" pageId="57" pageNumber="58">Panama</collectingCountry>
(
<figureCitation id="729A2A17C317C749E38D750DFBA9FEFE" box="[939,1043,284,310]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="57" pageNumber="58">Fig. 388</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C317C749E11E7552FC59FE4E" blockId="57.[264,1323,283,390]" pageId="57" pageNumber="58">
<emphasis id="D8D5EA80C317C749E11E7552FDB3FE95" bold="true" box="[312,521,323,349]" pageId="57" pageNumber="58">Natural History.</emphasis>
<taxonomicName id="2DA14D11C317C749E2397555FCEFFE95" box="[543,853,324,349]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C317C749E2397555FCEFFE95" box="[543,853,324,349]" italics="true" pageId="57" pageNumber="58">Seeversiella furcativentris</emphasis>
</taxonomicName>
was collected in forest litter and with flight intercept and malaise traps, at altitudes of
<quantity id="2D599B77C317C749E318757DFC65FE4E" box="[830,991,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.1" metricValueMin="1.4" pageId="57" pageNumber="58" unit="m" value="1750.0" valueMax="2100.0" valueMin="1400.0">1400­2100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

647
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@ -0,0 +1,647 @@
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<paragraph id="EA1E3692C31FC741E12E727AFB90F94D" blockId="49.[264,1066,1643,1670]" box="[264,1066,1643,1670]" pageId="49" pageNumber="50">
<heading id="B15681FEC31FC741E12E727AFB90F94D" bold="true" box="[264,1066,1643,1670]" fontSize="11" level="1" pageId="49" pageNumber="50" reason="1">
<emphasis id="D8D5EA80C31FC741E12E727AFB90F94D" bold="true" box="[264,1066,1643,1670]" pageId="49" pageNumber="50">
<key id="9E3061A3C31FC741E12E727AFB90F94D" box="[264,1066,1643,1670]" pageId="49" pageNumber="50">
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14.
<taxonomicName id="2DA14D11C31FC741E112727DFD7CF94D" ID-CoL="6YBDD" authority="Gusarov" authorityName="Gusarov" box="[308,710,1643,1670]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="scabricollis" status="sp. nov.">
<emphasis id="D8D5EA80C31FC741E112727DFDEEF94E" bold="true" box="[308,596,1644,1670]" italics="true" pageId="49" pageNumber="50">Seeversiella scabricollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC31FC741E2F5727AFCAFF94D" box="[723,789,1643,1669]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C31FC741E302727AFC0BF94D" box="[804,945,1643,1669]" captionStart="FIGURES 10 ­ 16" captionStartId="6.[264,383,1647,1671]" captionTargetBox="[319,1271,331,1612]" captionTargetId="figure@6.[304,1283,320,1620]" captionTargetPageId="6" captionText="FIGURES 10 ­ 16. Details of Seeversiella globicollis (Bernhauer) (10, Anticosti, Quebec; 13, Waterton Lakes National Park, Alberta), S. scabricollis Gusarov, sp. n. (11, 15, paratype from Volcán Barva, Costa Rica) and S. micralymma Gusarov, sp. n. (12, 14, 16, paratype from Cerro Buenavista, Costa Rica). 10 ­ 12 – pronotum; 13 ­ 14 – mesometathorax; 15 – male tergum 3 and paratergites; 16 – male tergum 3. Scale bar 0.4 mm (10 ­ 11, 15), 0.3 mm (12 ­ 13, 16), 0.2 mm (14)." httpUri="https://zenodo.org/record/156423/files/figure.png" pageId="49" pageNumber="50">Figs. 11, 15</figureCitation>
,
<figureCitation id="729A2A17C31FC741E399727AFB98F94D" box="[959,1058,1643,1669]" captionStart-0="FIGURES 190 ­ 193" captionStart-1="FIGURES 194 ­ 198" captionStart-2="FIGURES 199 ­ 207" captionStartId-0="51.[264,383,1774,1798]" captionStartId-1="52.[264,383,1328,1352]" captionStartId-2="53.[264,383,1638,1662]" captionTargetBox-0="[282,1304,939,1740]" captionTargetBox-1="[289,1303,584,1286]" captionTargetBox-2="[306,1280,457,1604]" captionTargetId-0="figure@51.[275,1312,929,1748]" captionTargetId-1="figure@52.[272,1315,569,1301]" captionTargetId-2="figure@53.[299,1288,449,1611]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="53" captionText-0="FIGURES 190 ­ 193. Abdominal segment 8 of Seeversiella scabricollis Gusarov, sp. n. (190 ­ 191, holotype; 192 ­ 193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 – male tergum 8; 191 – male sternum 8; 192 – female tergum 8; 193 – female sternum 8. Scale bar 0.4 mm." captionText-1="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 – median lobe, parameral view; 196 – apex of median lobe, parameral view; 197 – median lobe, lateral view; 198 – apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." captionText-2="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri-0="https://zenodo.org/record/156453/files/figure.png" httpUri-1="https://zenodo.org/record/156454/files/figure.png" httpUri-2="https://zenodo.org/record/156455/files/figure.png" pageId="49" pageNumber="50">190­207</figureCitation>
)
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</subSubSection>
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<emphasis id="D8D5EA80C31FC741E12E72A7FE7BF918" bold="true" box="[264,449,1718,1744]" pageId="49" pageNumber="50">
<typeStatus id="351A8830C31FC741E12E72A7FEF9F918" box="[264,323,1718,1744]" pageId="49" pageNumber="50">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C31FC741E1FA72A7FDF6F918" box="[476,588,1718,1744]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C31FC741E2A372A7FC68F918" bold="true" box="[645,978,1718,1744]" pageId="49" pageNumber="50">
<collectingCountry id="92B67602C31FC741E2A372A7FCF9F918" box="[645,835,1718,1744]" name="Costa Rica" pageId="49" pageNumber="50">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
Sendero a Cerro Chirripó,
<geoCoordinate id="8F955055C31FC741E12E72CFFE3EF930" box="[264,388,1758,1784]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="15" value="9.446389">9°26'47&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31FC741E1B072CFFD9DF930" box="[406,551,1758,1784]" direction="west" orientation="longitude" pageId="49" pageNumber="50" precision="15" value="-83.53667">83°32'12&quot;W</geoCoordinate>
,
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, mixed oak forest litter (R.Anderson),
<date id="9E1F1052C31FC741E48B72CFFA91F930" box="[1197,1323,1758,1784]" pageId="49" pageNumber="50" value="1999-06-27">27.vi.1999</date>
(
<collectionCode id="8CB0AE57C31FC741E1367317FED2F8E8" box="[272,360,1798,1824]" pageId="49" pageNumber="50">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C31FC742E11E733FFA91FA8E" blockId="49.[264,1323,1718,1904]" lastBlockId="50.[264,1325,284,1870]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<typeStatus id="351A8830C31FC741E11E733FFE15F880" box="[312,431,1838,1864]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C31FC741E1E3733FFCBAF880" bold="true" box="[453,768,1838,1864]" pageId="49" pageNumber="50">
<collectingCountry id="92B67602C31FC741E1E3733FFD3AF880" box="[453,640,1838,1864]" name="Costa Rica" pageId="49" pageNumber="50">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
<specimenCount id="FCA7FD1BC31FC741E335733FFC0DF880" box="[787,951,1838,1864]" pageId="49" pageNumber="50" type="generic">42 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C31FC741E4B2733FFB44F880" box="[1172,1278,1838,1864]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
;
<specimenCount id="FCA7FD1BC31FC741E52A733FFE3FF8B8" pageId="49" pageNumber="50" type="generic">50 specimens</specimenCount>
, ditto but
<geoCoordinate id="8F955055C31FC741E2227347FD3AF8B8" box="[516,640,1878,1904]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="15" value="9.443611">9°26'37&quot;N</geoCoordinate>
83°32'14&quot;,
<quantity id="2D599B77C31FC741E3377347FCCAF8B8" box="[785,880,1878,1904]" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="49" pageNumber="50" unit="m" value="2950.0">2950 m</quantity>
, oak forest litter;, Cerro Chirripó, Base Crestones,
<geoCoordinate id="8F955055C31CC742E1C5750DFDE4FEFE" box="[483,606,284,310]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.454722">9°27'17&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E257750DFCB8FEFE" box="[625,770,284,310]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.50445">83°30'16&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E33C750DFC38FEFE" box="[794,898,284,310]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="50" pageNumber="51" unit="m" value="3350.0">3350 m</quantity>
, elfin forest litter (R.Anderson),
<date id="9E1F1052C31CC742E12E7555FE32FE96" box="[264,392,324,350]" pageId="50" pageNumber="51" value="1999-06-25">25.vi.1999</date>
;
<specimenCount id="FCA7FD1BC31CC742E1B97555FDF0FE96" box="[415,586,324,350]" pageId="50" pageNumber="51" type="generic">17 specimens</specimenCount>
,
<quantity id="2D599B77C31CC742E2797555FD1FFE96" box="[607,677,324,350]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="km" value="2.0">2 km</quantity>
S Villa Mills,
<quantity id="2D599B77C31CC742E3517555FC66FE96" box="[887,988,324,350]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="m" value="3000.0">3000 m</quantity>
, ridgetop oak forest litter (R.S.Anderson),
<date id="9E1F1052C31CC742E1E9757DFDF0FE4E" box="[463,586,364,390]" pageId="50" pageNumber="51" value="1996-02-10">10.ii.1996</date>
;, 2.0 km E Villa Mills,
<geoCoordinate id="8F955055C31CC742E35F757DFC4FFE4E" box="[889,1013,364,390]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.566667">9°34'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E3DA757DFB36FE4E" box="[1020,1164,364,390]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.69722">83°41'50&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E4BF757DFB4FFE4E" box="[1177,1269,364,390]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="50" pageNumber="51" unit="m" value="2750.0">2750 m</quantity>
, oak forest litter (R.S.Anderson),
<date id="9E1F1052C31CC742E24A7585FD5DFE66" box="[620,743,404,430]" pageId="50" pageNumber="51" value="1998-02-15">15.ii.1998</date>
;, Cerro Buenavista,
<geoCoordinate id="8F955055C31CC742E42A7585FB32FE66" box="[1036,1160,404,430]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.717222">9°43'02&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E4B07585FA9CFE66" box="[1174,1318,404,430]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.75389">83°45'14&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E12E75ADFED3FE1E" box="[264,361,444,470]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="50" pageNumber="51" unit="m" value="3300.0">3300 m</quantity>
, subparamo litter (R.Anderson);
<specimenCount id="FCA7FD1BC31CC742E2D875ADFC1FFE1E" box="[766,933,444,470]" pageId="50" pageNumber="51" type="generic">10 specimens</specimenCount>
, Cerro de la Muerte,
<quantity id="2D599B77C31CC742E49375ADFB4CFE1E" box="[1205,1270,444,470]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="50" pageNumber="51" unit="km" value="4.0">4 km</quantity>
NE Cañon,
<quantity id="2D599B77C31CC742E14375F5FE7BFE36" box="[357,449,484,510]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="50" pageNumber="51" unit="m" value="2350.0">2350 m</quantity>
, malaise trap (P.Hanson),
<date id="9E1F1052C31CC742E2DF75F5FCFCFE36" box="[761,838,484,510]" pageId="50" pageNumber="51" value="1995-01">i.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E37275F5FC52FE36" box="[852,1000,484,510]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E44475F5FAB6FE36" box="[1122,1292,484,510]" pageId="50" pageNumber="51" value="1995-01-01" valueMax="1995-03-30" valueMin="1995-01-01">1.i­30.iii.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E53C75F5FE3FFDEE" pageId="50" pageNumber="51" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E225761DFD20FDEE" box="[515,666,524,550]" pageId="50" pageNumber="51" value="1995-04-01" valueMax="1995-04-30" valueMin="1995-04-01">1­30.iv.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E28F761DFC89FDEE" box="[681,819,524,550]" pageId="50" pageNumber="51" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E397761DFBFAFDEE" box="[945,1088,524,550]" pageId="50" pageNumber="51" value="1995-05-01" valueMax="1995-05-30" valueMin="1995-05-01">1­30.v.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E469761DFB5FFDEE" box="[1103,1253,524,550]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E1667625FE62FD86" box="[320,472,564,590]" pageId="50" pageNumber="51" value="1995-06-01" valueMax="1995-06-30" valueMin="1995-06-01">1­30.vi.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E1D67625FD34FD86" box="[496,654,564,590]" pageId="50" pageNumber="51" type="generic">6 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E3027625FC7FFD86" box="[804,965,564,590]" pageId="50" pageNumber="51" value="1995-07-01" valueMax="1995-07-30" valueMin="1995-07-01">1­30.vii.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E3FA7625FBC0FD86" box="[988,1146,564,590]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E5377625FE23FDBE" pageId="50" pageNumber="51" value="1995-08-01" valueMax="1995-08-30" valueMin="1995-08-01">1­ 30.viii.1995</date>
;
<specimenCount id="FCA7FD1BC31CC742E18F764DFD8FFDBE" box="[425,565,604,630]" pageId="50" pageNumber="51" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E290764DFCFAFDBE" box="[694,832,604,630]" pageId="50" pageNumber="51" value="1996-01-01" valueMax="1996-01-30" valueMin="1996-01-01">1­30.i.1996</date>
;
<specimenCount id="FCA7FD1BC31CC742E376764DFC66FDBE" box="[848,988,604,630]" pageId="50" pageNumber="51" type="generic">1 specimen</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E47B764DFAB0FDBE" box="[1117,1290,604,630]" pageId="50" pageNumber="51" value="1996-04-01" valueMax="1996-05-30" valueMin="1996-04-01">1.iv­30.v.1996</date>
;
<specimenCount id="FCA7FD1BC31CC742E53C764DFE3FFD56" pageId="50" pageNumber="51" type="generic">7 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E1DF7695FD20FD56" box="[505,666,644,670]" pageId="50" pageNumber="51" value="1996-07-01" valueMax="1996-07-30" valueMin="1996-07-01">1­30.vii.1996</date>
;
<specimenCount id="FCA7FD1BC31CC742E2807695FC82FD56" box="[678,824,644,670]" pageId="50" pageNumber="51" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E38A7695FBFEFD56" box="[940,1092,644,670]" pageId="50" pageNumber="51" value="1996-09-01" valueMax="1996-09-30" valueMin="1996-09-01">1­30.ix.1996</date>
;
<specimenCount id="FCA7FD1BC31CC742E4767695FB59FD56" box="[1104,1251,644,670]" pageId="50" pageNumber="51" type="generic">4 specimens</specimenCount>
,
<quantity id="2D599B77C31CC742E4C87695FA93FD56" box="[1262,1321,644,670]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="km" value="2.0">2 km</quantity>
N P. N. Volcán Irazú,
<geoCoordinate id="8F955055C31CC742E22E76BDFD28FD0E" box="[520,658,684,710]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="10.0">10°00'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E2BC76BDFC93FD0E" box="[666,809,684,710]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.85">83°51'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E31076BDFC28FD0E" box="[822,914,684,710]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="50" pageNumber="51" unit="m" value="2900.0">2900 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E12E76C5FE32FD26" box="[264,392,724,750]" pageId="50" pageNumber="51" value="1997-06-18">18.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC742E1B376C5FD92FD26" box="[405,552,724,750]" pageId="50" pageNumber="51" type="generic">6 specimens</specimenCount>
,
<quantity id="2D599B77C31CC742E21576C5FDD4FD26" box="[563,622,724,750]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="km" value="3.0">3 km</quantity>
N P. N. Volcán Irazú,
<geoCoordinate id="8F955055C31CC742E35376C5FC4BFD26" box="[885,1009,724,750]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.995833">9°59'45&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E3DE76C5FB32FD26" box="[1016,1160,724,750]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.85">83°51'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E4B276C5FB4AFD26" box="[1172,1264,724,750]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="50" pageNumber="51" unit="m" value="2750.0">2750 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E25F76EDFD43FCDE" box="[633,761,764,790]" pageId="50" pageNumber="51" value="1997-06-18">18.vi.1997</date>
;, Madreselva, Finca Los Lagos, A. C. Amistad,
<quantity id="2D599B77C31CC742E11C7735FE63FCF6" box="[314,473,804,830]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="50" pageNumber="51" unit="m" value="2300.0" valueMax="2600.0" valueMin="2000.0">2000­2600 m</quantity>
,
<date id="9E1F1052C31CC742E1C27735FD36FCF6" box="[484,652,804,830]" pageId="50" pageNumber="51" value="1993-06-12" valueMax="1993-06-19" valueMin="1993-06-12">12­19.vi.1993</date>
; 2, ditto but
<date id="9E1F1052C31CC742E3627735FBA9FCF6" box="[836,1043,804,830]" pageId="50" pageNumber="51" value="1993-06-28" valueMax="1993-07-10" valueMin="1993-06-28">28.vi­10.vii.1993</date>
;, ditto but
<date id="9E1F1052C31CC742E48E7735FABEFCF6" box="[1192,1284,804,830]" pageId="50" pageNumber="51" value="1993-09">ix.1993</date>
;, ditto but
<date id="9E1F1052C31CC742E155775DFE7DFCAE" box="[371,455,844,870]" pageId="50" pageNumber="51" value="1993-10">x.1993</date>
;
<emphasis id="D8D5EA80C31CC742E1F2775AFD73FCAD" bold="true" box="[468,713,843,869]" pageId="50" pageNumber="51">San Jose / Cartago:</emphasis>
<specimenCount id="FCA7FD1BC31CC742E2FF775DFCC1FCAE" box="[729,891,844,870]" pageId="50" pageNumber="51" type="generic">11 specimens</specimenCount>
, km 71, Int. Amer. Hwy., nr. Tres de Junio,
<geoCoordinate id="8F955055C31CC742E17F7765FE6EFC46" box="[345,468,884,910]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.628889">9°37'44&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E1F87765FDD4FC46" box="[478,622,884,910]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.836945">83°50'13&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E25A7765FD61FC46" box="[636,731,884,910]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="50" pageNumber="51" unit="m" value="2900.0">2900 m</quantity>
, wet cloud forest litter (R.Anderson),
<date id="9E1F1052C31CC742E4807765FA9DFC46" box="[1190,1319,884,910]" pageId="50" pageNumber="51" value="1999-06-23">23.vi.1999</date>
; 3, km 69, Int. Amer. Hwy., nr. Tres de Junio,
<geoCoordinate id="8F955055C31CC742E37D778DFC6DFC7E" box="[859,983,924,950]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.658333">9°39'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E3C6778DFBCBFC7E" box="[992,1137,924,950]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.85833">83°51'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E459778DFB64FC7E" box="[1151,1246,924,950]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="50" pageNumber="51" unit="m" value="2600.0">2600 m</quantity>
, flight intercept trap (S. &amp; J.Peck),
<date id="9E1F1052C31CC742E27C77D5FD49FC16" box="[602,755,964,990]" pageId="50" pageNumber="51" value="1997-06-07" valueMax="1997-06-19" valueMin="1997-06-07">7­19.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC742E32777D5FC18FC16" box="[769,930,964,990]" pageId="50" pageNumber="51" type="generic">11 specimens</specimenCount>
, km 78, Int. Amer. Hwy., nr. Ojo de Agua,
<geoCoordinate id="8F955055C31CC742E15B77FDFE43FBCE" box="[381,505,1004,1030]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.608334">9°36'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E22477FDFD28FBCE" box="[514,658,1004,1030]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.83889">83°50'20&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E28677FDFD47FBCE" box="[672,765,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="50" pageNumber="51" unit="m" value="2800.0">2800 m</quantity>
, cloud forest litter (R.Anderson),
<date id="9E1F1052C31CC742E4B477FDFAB6FBCE" box="[1170,1292,1004,1030]" pageId="50" pageNumber="51" value="1999-07-08">8.vii.1999</date>
;
<specimenCount id="FCA7FD1BC31CC742E53C77FDFE3FFBE6" pageId="50" pageNumber="51" type="generic">9 specimens</specimenCount>
, km 72, Int. Amer. Hwy.,
<quantity id="2D599B77C31CC742E29B7005FD40FBE6" box="[701,762,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="km" value="3.0">3 km</quantity>
W Ojo de Agua,
<geoCoordinate id="8F955055C31CC742E3E97005FBF0FBE6" box="[975,1098,1044,1070]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.625">9°37'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E4757005FB59FBE6" box="[1107,1251,1044,1070]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.84167">83°50'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E4D77005FE98FB9E" metricMagnitude="3" metricUnit="m" metricValue="2.95" pageId="50" pageNumber="51" unit="m" value="2950.0">2950 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E2F1702DFCF2FB9E" box="[727,840,1084,1110]" pageId="50" pageNumber="51" value="1997-06-07">7.vi.1997</date>
;, km 87, Int. Amer. Hwy., nr. Cerro Buenavista,
<geoCoordinate id="8F955055C31CC742E1BF7075FDAFFBB6" box="[409,533,1124,1150]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.608334">9°36'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E23A7075FD17FBB6" box="[540,685,1124,1150]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.76667">83°46'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E29F7075FCACFBB6" box="[697,790,1124,1150]" metricMagnitude="3" metricUnit="m" metricValue="3.15" pageId="50" pageNumber="51" unit="m" value="3150.0">3150 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E4927075FA9CFBB6" box="[1204,1318,1124,1150]" pageId="50" pageNumber="51" value="1997-06-08">8.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC742E12E709DFE21FB6E" box="[264,411,1164,1190]" pageId="50" pageNumber="51" type="generic">8 specimens</specimenCount>
, Cerro Buenavista, km 89, Int. Amer. Hwy.,
<geoCoordinate id="8F955055C31CC742E38A709DFB92FB6E" box="[940,1064,1164,1190]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.55">9°33'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E409709DFB7AFB6E" box="[1071,1216,1164,1190]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.75833">83°45'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E4EA709DFA93FB6E" box="[1228,1321,1164,1190]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="50" pageNumber="51" unit="m" value="3200.0">3200 m</quantity>
, berlese leaf litter (R.Anderson),
<date id="9E1F1052C31CC742E2AD70A5FCB1FB06" box="[651,779,1204,1230]" pageId="50" pageNumber="51" value="1998-06-18">18.vi.1998</date>
;, 2, km 55, Int. Amer. Hwy.,
<quantity id="2D599B77C31CC742E49670A5FB57FB06" box="[1200,1261,1204,1230]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="km" value="3.0">3 km</quantity>
S El Empalme,
<geoCoordinate id="8F955055C31CC742E1AF70CDFDBEFB3E" box="[393,516,1244,1270]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.708333">9°42'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E22870CDFD25FB3E" box="[526,671,1244,1270]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.95">83°57'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E28B70CDFCB1FB3E" box="[685,779,1244,1270]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="50" pageNumber="51" unit="m" value="2350.0">2350 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E49270CDFA9CFB3E" box="[1204,1318,1244,1270]" pageId="50" pageNumber="51" value="1997-06-08">8.vi.1997</date>
;
<emphasis id="D8D5EA80C31CC742E12E7112FEC4FAD5" bold="true" box="[264,382,1283,1309]" pageId="50" pageNumber="51">San Jose:</emphasis>
<specimenCount id="FCA7FD1BC31CC742E1A97115FD8BFAD6" box="[399,561,1284,1310]" pageId="50" pageNumber="51" type="generic">15 specimens</specimenCount>
, Estación Cuerici,
<quantity id="2D599B77C31CC742E3367115FCD9FAD6" box="[784,867,1284,1310]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="50" pageNumber="51" unit="km" value="4.6">4.6 km</quantity>
E Villa Mills,
<geoCoordinate id="8F955055C31CC742E4357115FB35FAD6" box="[1043,1167,1284,1310]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.566667">9°34'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E4B07115FA9DFAD6" box="[1174,1319,1284,1310]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.666664">83°40'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E12E713DFEDFFA8E" box="[264,357,1324,1350]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="50" pageNumber="51" unit="m" value="2600.0">2600 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E32C713DFC31FA8E" box="[778,907,1324,1350]" pageId="50" pageNumber="51" value="1997-06-26">26.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC742E3BE713DFB96FA8E" box="[920,1068,1324,1350]" pageId="50" pageNumber="51" type="generic">3 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E480713DFA9CFA8E" box="[1190,1318,1324,1350]" pageId="50" pageNumber="51" value="1997-06-19">19.vi.1997</date>
;
</paragraph>
<paragraph id="EA1E3692C31CC743E13A7145FA97FD86" blockId="50.[264,1325,284,1870]" lastBlockId="51.[264,1325,284,870]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
, 2.0 km E Villa Mills,
<geoCoordinate id="8F955055C31CC742E21E7145FD0EFAA6" box="[568,692,1364,1390]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.558333">9°33'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E2987145FCF5FAA6" box="[702,847,1364,1390]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-83.7">83°42'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E37B7145FC01FAA6" box="[861,955,1364,1390]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="50" pageNumber="51" unit="m" value="2800.0">2800 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E16A716DFE77FA5E" box="[332,461,1404,1430]" pageId="50" pageNumber="51" value="1997-06-26">26.vi.1997</date>
;, 2, km 97, Int. Amer. Hwy., Villa Mills,
<quantity id="2D599B77C31CC742E428716DFBD6FA5E" box="[1038,1132,1404,1430]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="m" value="3000.0">3000 m</quantity>
, flight intercept traps (S. &amp; J.Peck),
<date id="9E1F1052C31CC742E23371B5FD2CFA76" box="[533,662,1444,1470]" pageId="50" pageNumber="51" value="1997-06-26">26.vi.1997</date>
; 3,, Cerros de Escazú,
<quantity id="2D599B77C31CC742E43271B5FBEDFA76" box="[1044,1111,1444,1470]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="km" value="2.0">2 km</quantity>
S San Antonio,
<geoCoordinate id="8F955055C31CC742E12E71DDFE3EFA2E" box="[264,388,1484,1510]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="9.891666">9°53'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E1A871DDFDA5FA2E" box="[398,543,1484,1510]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-84.15">84°09'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E20871DDFD37FA2E" box="[558,653,1484,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="50" pageNumber="51" unit="m" value="1650.0">1650 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E41A71DDFB07FA2E" box="[1084,1213,1484,1510]" pageId="50" pageNumber="51" value="1997-06-13">13.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC742E4EA71DDFEF3F9C6" pageId="50" pageNumber="51" type="generic">8 specimens</specimenCount>
, Cerro de la Muerte,
<quantity id="2D599B77C31CC742E26571E5FDC5F9C6" box="[579,639,1524,1550]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="50" pageNumber="51" unit="km" value="6.0">6 km</quantity>
N San Gerardo,
<quantity id="2D599B77C31CC742E36071E5FC25F9C6" box="[838,927,1524,1550]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="50" pageNumber="51" unit="m" value="2800.0">2800 m</quantity>
(P.Hanson), viii­ix.1992;
<specimenCount id="FCA7FD1BC31CC742E4F771E5FEF3F9FE" pageId="50" pageNumber="51" type="generic">4 specimens</specimenCount>
, ditto but x­xii.1992;
<specimenCount id="FCA7FD1BC31CC742E270720DFD57F9FE" box="[598,749,1564,1590]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
, ditto but
<date id="9E1F1052C31CC742E348720DFC72F9FE" box="[878,968,1564,1590]" pageId="50" pageNumber="51" value="1992-04">iv.1992</date>
; 2, malaise trap, ditto but
<date id="9E1F1052C31CC742E12E7255FED9F996" box="[264,355,1604,1630]" pageId="50" pageNumber="51" value="1993-04">iv.1993</date>
;, Cerro de la Muerte,
<quantity id="2D599B77C31CC742E2A07255FD68F996" box="[646,722,1604,1630]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="50" pageNumber="51" unit="km" value="19.0">19 km</quantity>
S and
<quantity id="2D599B77C31CC742E3007255FCDEF996" box="[806,868,1604,1630]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="50" pageNumber="51" unit="km" value="3.0">3 km</quantity>
W of Empalme,
<quantity id="2D599B77C31CC742E4147255FB34F996" box="[1074,1166,1604,1630]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="50" pageNumber="51" unit="m" value="2600.0">2600 m</quantity>
, malaise trap (P.Hanson), ix­x.1992; 7, 26 km N San Isidro,
<quantity id="2D599B77C31CC742E347727DFC00F94E" box="[865,954,1644,1670]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="50" pageNumber="51" unit="m" value="2100.0">2100 m</quantity>
(P.Hanson),
<date id="9E1F1052C31CC742E472727DFB0AF94E" box="[1108,1200,1644,1670]" pageId="50" pageNumber="51" value="1992-11">xi.1992</date>
­
<date id="9E1F1052C31CC742E49F727DFABEF94E" box="[1209,1284,1644,1670]" pageId="50" pageNumber="51" value="1993-01">i.1993</date>
;, ditto but vi­vii.1990;, ditto but ix­x.1992;
<specimenCount id="FCA7FD1BC31CC742E33C7285FC14F966" box="[794,942,1684,1710]" pageId="50" pageNumber="51" type="generic">2 specimens</specimenCount>
,
<quantity id="2D599B77C31CC742E39D7285FC42F966" box="[955,1016,1684,1710]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="km" value="2.0">2 km</quantity>
W Empalme, Cerro de la Muerte,
<quantity id="2D599B77C31CC742E14872ADFE70F91E" box="[366,458,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="50" pageNumber="51" unit="m" value="2300.0">2300 m</quantity>
(P.Hanson),
<date id="9E1F1052C31CC742E24172ADFCB2F91E" box="[615,776,1724,1750]" pageId="50" pageNumber="51" value="1995-07-01" valueMax="1995-07-30" valueMin="1995-07-01">1­30.vii.1995</date>
, malaise trap;
<emphasis id="D8D5EA80C31CC742E39C72AAFB93F91D" bold="true" box="[954,1065,1723,1749]" pageId="50" pageNumber="51">Heredia:</emphasis>
2, 2,
<specimenCount id="FCA7FD1BC31CC742E4EA72ADFEF3F936" pageId="50" pageNumber="51" type="generic">3 specimens</specimenCount>
, Porrosati,
<quantity id="2D599B77C31CC742E1F672F5FDB4F936" box="[464,526,1764,1790]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="50" pageNumber="51" unit="km" value="6.0">6 km</quantity>
N San José de la Montana,
<geoCoordinate id="8F955055C31CC742E37972F5FC53F936" box="[863,1001,1764,1790]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="10.091666">10°05'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E3D472F5FB38F936" box="[1010,1154,1764,1790]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-84.11667">84°07'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E4A972F5FB56F936" box="[1167,1260,1764,1790]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="50" pageNumber="51" unit="m" value="1900.0">1900 m</quantity>
, berlese forest litter (R.Anderson);
<specimenCount id="FCA7FD1BC31CC742E2BF731DFCF9F8EE" box="[665,835,1804,1830]" pageId="50" pageNumber="51" type="generic">44 specimens</specimenCount>
, P. N. Braulio Carrillo Volcán Barva,
<geoCoordinate id="8F955055C31CC742E12E7325FE28F886" box="[264,402,1844,1870]" direction="north" orientation="latitude" pageId="50" pageNumber="51" precision="15" value="10.125">10°07'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31CC742E1BC7325FD90F886" box="[410,554,1844,1870]" direction="west" orientation="longitude" pageId="50" pageNumber="51" precision="15" value="-84.125">84°07'30&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31CC742E2117325FD29F886" box="[567,659,1844,1870]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="50" pageNumber="51" unit="m" value="2600.0">2600 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31CC742E4127325FB09F886" box="[1076,1203,1844,1870]" pageId="50" pageNumber="51" value="1997-06-11">11.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31CC743E4E67325FEF3FEFE" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="generic">21 specimens</specimenCount>
, ditto but, flight intercept trap (S. &amp; J.Peck),
<date id="9E1F1052C31DC743E3B7750DFB82FEFE" box="[913,1080,284,310]" pageId="51" pageNumber="52" value="1997-06-11" valueMax="1997-06-27" valueMin="1997-06-11">11­27.vi.1997</date>
;
<specimenCount id="FCA7FD1BC31DC743E46D750DFB5EFEFE" box="[1099,1252,284,310]" pageId="51" pageNumber="52" type="generic">4 specimens</specimenCount>
, Vara Blanca, Finca Georgina,
<quantity id="2D599B77C31DC743E2147555FD37FE96" box="[562,653,324,350]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="51" pageNumber="52" unit="m" value="2100.0">2100 m</quantity>
(P.Hanson), vi­vii.1990;
<emphasis id="D8D5EA80C31DC743E39C7552FB97FE95" bold="true" box="[954,1069,323,349]" pageId="51" pageNumber="52">Alajuela:</emphasis>
<specimenCount id="FCA7FD1BC31DC743E4197555FB59FE96" box="[1087,1251,324,350]" pageId="51" pageNumber="52" type="generic">28 specimens</specimenCount>
, P. N. Volcán Poás,
<geoCoordinate id="8F955055C31DC743E19B757DFDFCFE4E" box="[445,582,364,390]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="10.191667">10°11'30&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31DC743E27E757DFD53FE4E" box="[600,745,364,390]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-84.23333">84°14'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31DC743E326757DFCD2FE4E" box="[768,872,364,390]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="51" pageNumber="52" unit="m" value="2500.0">2500 m</quantity>
, berlese forest litter (R.Anderson),
<date id="9E1F1052C31DC743E12E7585FEC0FE66" box="[264,378,404,430]" pageId="51" pageNumber="52" value="1997-06-06">6.vi.1997</date>
;,, ditto but flight intercept trap (S. &amp; J.Peck),
<date id="9E1F1052C31DC743E3DF7585FB28FE66" box="[1017,1170,404,430]" pageId="51" pageNumber="52" value="1997-06-06" valueMax="1997-06-28" valueMin="1997-06-06">6­28.vi.1997</date>
;
<emphasis id="D8D5EA80C31DC743E4877582FEC7FE1D" bold="true" pageId="51" pageNumber="52">
<collectingCountry id="92B67602C31DC743E4877582FA9CFE65" box="[1185,1318,403,429]" name="Panama" pageId="51" pageNumber="52">PANAMA</collectingCountry>
: Chiriquí:
</emphasis>
<specimenCount id="FCA7FD1BC31DC743E1AA75ADFD94FE1E" box="[396,558,444,470]" pageId="51" pageNumber="52" type="generic">31 specimens</specimenCount>
,
<quantity id="2D599B77C31DC743E21F75ADFD31FE1E" box="[569,651,444,470]" metricMagnitude="3" metricUnit="m" metricValue="5.9" pageId="51" pageNumber="52" unit="km" value="5.9">5.9 km</quantity>
NE Cerro Punta, Par. Nac. Volcán Barú,
<geoCoordinate id="8F955055C31DC743E44B75ADFB7AFE1E" box="[1133,1216,444,470]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="925" value="8.866667">8°52'N</geoCoordinate>
[the latitude on the label (
<geoCoordinate id="8F955055C31DC743E1FF75F5FDF2FE36" box="[473,584,484,510]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="8.366667">8°22'0&quot;N</geoCoordinate>
) is incorrect]
<geoCoordinate id="8F955055C31DC743E2CD75F5FCE8FE36" box="[747,850,484,510]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="925" value="-82.566666">82°34'W</geoCoordinate>
,
<quantity id="2D599B77C31DC743E37875F5FC03FE36" box="[862,953,484,510]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="51" pageNumber="52" unit="m" value="2100.0">2100 m</quantity>
, flight intercept traps (J.S.Ashe &amp; R.Brooks),
<date id="9E1F1052C31DC743E193761DFDE7FDEE" box="[437,605,524,550]" pageId="51" pageNumber="52" value="1995-06-14" valueMax="1995-06-16" valueMin="1995-06-14">14­16.vi.1995</date>
;, ditto but
<quantity id="2D599B77C31DC743E322761DFCD8FDEE" box="[772,866,524,550]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="51" pageNumber="52" unit="m" value="2400.0">2400 m</quantity>
, bamboo forest litter (R.S.Anderson),
<date id="9E1F1052C31DC743E12E7625FE33FD86" box="[264,393,564,590]" pageId="51" pageNumber="52" value="1995-06-14">14.vi.1995</date>
;
<specimenCount id="FCA7FD1BC31DC743E1B37625FD82FD86" box="[405,568,564,590]" pageId="51" pageNumber="52" type="generic">12 specimens</specimenCount>
, ditto but
<quantity id="2D599B77C31DC743E28B7625FCB3FD86" box="[685,777,564,590]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="51" pageNumber="52" unit="m" value="2150.0">2150 m</quantity>
, alder forest litter (R.S.Anderson),
<date id="9E1F1052C31DC743E4817625FA92FD86" box="[1191,1320,564,590]" pageId="51" pageNumber="52" value="1995-06-14">14.vi.1995</date>
;
</paragraph>
<paragraph id="EA1E3692C31DC743E13A764DFB0CFCAE" blockId="51.[264,1325,284,870]" pageId="51" pageNumber="52">
,
<quantity id="2D599B77C31DC743E10D764DFE37FDBE" box="[299,397,604,630]" metricMagnitude="4" metricUnit="m" metricValue="2.77" pageId="51" pageNumber="52" unit="km" value="27.7">27.7 km</quantity>
W Volcán Hartmann's Finca,
<quantity id="2D599B77C31DC743E2D5764DFCEAFDBE" box="[755,848,604,630]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="51" pageNumber="52" unit="m" value="1450.0">1450 m</quantity>
,
<geoCoordinate id="8F955055C31DC743E37B764DFC6DFDBE" box="[861,983,604,630]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="8.863334">8°51'48&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31DC743E3C6764DFBCBFDBE" box="[992,1137,604,630]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-82.74333">82°44'36&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31DC743E458764DFB61FDBE" box="[1150,1243,604,630]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="51" pageNumber="52" unit="m" value="1450.0">1450 m</quantity>
, forest litter (R.S.Anderson),
<date id="9E1F1052C31DC743E22B7695FD34FD56" box="[525,654,644,670]" pageId="51" pageNumber="52" value="1995-06-14">14.vi.1995</date>
;
<specimenCount id="FCA7FD1BC31DC743E2BF7695FC81FD56" box="[665,827,644,670]" pageId="51" pageNumber="52" type="generic">22 specimens</specimenCount>
, NNE of Las Nubes @ continental divide,
<quantity id="2D599B77C31DC743E12E76BDFEDBFD0E" box="[264,353,684,710]" metricMagnitude="3" metricUnit="m" metricValue="2.4899999999999998" pageId="51" pageNumber="52" unit="m" value="2490.0">2490 m</quantity>
(A.R.Gillogly),
<date id="9E1F1052C31DC743E20576BDFD70FD0E" box="[547,714,684,710]" pageId="51" pageNumber="52" value="1995-06-14" valueMax="1995-06-15" valueMin="1995-06-14">14­15.vi.1995</date>
;
<specimenCount id="FCA7FD1BC31DC743E2F076BDFCD3FD0E" box="[726,873,684,710]" pageId="51" pageNumber="52" type="generic">5 specimens</specimenCount>
, N of Las Nubes,
<quantity id="2D599B77C31DC743E41D76BDFB61FD0E" box="[1083,1243,684,710]" metricMagnitude="3" metricUnit="m" metricValue="2.4" metricValueMax="2.45" metricValueMin="2.35" pageId="51" pageNumber="52" unit="m" value="2400.0" valueMax="2450.0" valueMin="2350.0">2350­2450 m</quantity>
, continental divide (A.R.Gillogly),
<date id="9E1F1052C31DC743E2AE76C5FCB2FD26" box="[648,776,724,750]" pageId="51" pageNumber="52" value="1995-06-15">15.vi.1995</date>
; 2, Cerro Pando,
<quantity id="2D599B77C31DC743E40976C5FB2DFD26" box="[1071,1175,724,750]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="51" pageNumber="52" unit="m" value="1875.0">1875 m</quantity>
,
<geoCoordinate id="8F955055C31DC743E48976C5FA91FD26" box="[1199,1323,724,750]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="8.911666">8°54'42&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31DC743E12E76EDFE23FCDE" box="[264,409,764,790]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-82.721664">82°43'18&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31DC743E18E76EDFDBDFCDE" box="[424,519,764,790]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="51" pageNumber="52" unit="m" value="1875.0">1875 m</quantity>
, flight intercept trap (J.S.Ashe, R.Brooks),
<date id="9E1F1052C31DC743E43176EDFB04FCDE" box="[1047,1214,764,790]" pageId="51" pageNumber="52" value="1996-06-17" valueMax="1996-06-18" valueMin="1996-06-17">17­18.vi.1996</date>
;, ditto but
<quantity id="2D599B77C31DC743E1187735FE18FCF6" box="[318,418,804,830]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="51" pageNumber="52" unit="m" value="1850.0">1850 m</quantity>
;
<specimenCount id="FCA7FD1BC31DC743E19E7735FDEEFCF6" box="[440,596,804,830]" pageId="51" pageNumber="52" type="generic">6 specimens</specimenCount>
,
<quantity id="2D599B77C31DC743E24E7735FD00FCF6" box="[616,698,804,830]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="51" pageNumber="52" unit="km" value="11.0">11 km</quantity>
NW Boquete, Volcán Barú,
<quantity id="2D599B77C31DC743E4107735FB21FCF6" box="[1078,1179,804,830]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="51" pageNumber="52" unit="m" value="2150.0">2150 m</quantity>
,
<geoCoordinate id="8F955055C31DC743E4967735FA91FCF6" box="[1200,1323,804,830]" direction="north" orientation="latitude" pageId="51" pageNumber="52" precision="15" value="8.8">8°48'00&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C31DC743E12E775DFE23FCAE" box="[264,409,844,870]" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="15" value="-82.48333">82°29'00&quot;W</geoCoordinate>
,
<quantity id="2D599B77C31DC743E183775DFDBBFCAE" box="[421,513,844,870]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="51" pageNumber="52" unit="m" value="2150.0">2150 m</quantity>
, oak forest litter (R.S.Anderson),
<date id="9E1F1052C31DC743E3A8775DFBB6FCAE" box="[910,1036,844,870]" pageId="51" pageNumber="52" value="1995-06-18">18.vi.1995</date>
(all ­
<collectionCode id="8CB0AE57C31DC743E475775DFB10FCAE" box="[1107,1194,844,870]" pageId="51" pageNumber="52">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC31DC743E12E72FFFB34F883" httpUri="https://zenodo.org/record/156453/files/figure.png" pageId="51" pageNumber="52" targetBox="[282,1304,939,1740]" targetPageId="51">
<paragraph id="EA1E3692C31DC743E12E72FFFB34F883" blockId="51.[264,1325,1774,1867]" pageId="51" pageNumber="52">
<emphasis id="D8D5EA80C31DC743E12E72FFFE5CF8CE" bold="true" box="[264,486,1774,1798]" pageId="51" pageNumber="52">FIGURES 190­193.</emphasis>
Abdominal segment 8 of
<taxonomicName id="2DA14D11C31DC743E32F72FEFBC8F8CE" authority="Gusarov" authorityName="Gusarov" box="[777,1138,1775,1798]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="scabricollis" status="sp. nov.">
<emphasis id="D8D5EA80C31DC743E32F72FEFBB5F8CE" box="[777,1039,1775,1798]" italics="true" pageId="51" pageNumber="52">Seeversiella scabricollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC31DC743E45B72FFFB00F8CE" box="[1149,1210,1774,1798]" pageId="51" pageNumber="52" rank="species">
<emphasis id="D8D5EA80C31DC743E45B72FFFB00F8CE" bold="true" box="[1149,1210,1774,1798]" pageId="51" pageNumber="52">sp. n.</emphasis>
</taxonomicNameLabel>
(190­191, holotype; 192­193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 – male tergum 8; 191 – male sternum 8; 192 – female tergum 8; 193 – female sternum 8. Scale bar 0.4 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C31AC744E11E750AFC04F98A" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="EA1E3692C31AC744E11E750AFDA5FE36" blockId="52.[264,1323,283,510]" pageId="52" pageNumber="53">
<emphasis id="D8D5EA80C31AC744E11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C31AC744E1EC750DFD51FEFD" box="[458,747,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C31AC744E1EC750DFD51FEFD" box="[458,747,284,309]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C31AC744E4DE750DFED7FE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C31AC744E4DE750DFED7FE95" italics="true" pageId="52" pageNumber="53">Seeversiella</emphasis>
</taxonomicName>
by having dark brown body with brown elytra, legs and mouthparts; large eyes (temple length to eye length ratio 0.9­1.1); matte pronotum with strong microsculpture; elytra longer than pronotum; wings fully developed or reduced; tergum 8 with three pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C31AC744E31F75ADFBE9FE1E" box="[825,1107,444,470]" captionStart-0="FIGURES 194 ­ 198" captionStart-1="FIGURES 199 ­ 207" captionStartId-0="52.[264,383,1328,1352]" captionStartId-1="53.[264,383,1638,1662]" captionTargetBox-0="[289,1303,584,1286]" captionTargetBox-1="[306,1280,457,1604]" captionTargetId-0="figure@52.[272,1315,569,1301]" captionTargetId-1="figure@53.[299,1288,449,1611]" captionTargetPageId-0="52" captionTargetPageId-1="53" captionText-0="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 – median lobe, parameral view; 196 – apex of median lobe, parameral view; 197 – median lobe, lateral view; 198 – apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." captionText-1="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri-0="https://zenodo.org/record/156454/files/figure.png" httpUri-1="https://zenodo.org/record/156455/files/figure.png" pageId="52" pageNumber="53">Figs. 194­204, 206­207</figureCitation>
) and the shape of spermatheca (
<figureCitation id="729A2A17C31AC744E18D75F5FDA8FE36" box="[427,530,484,510]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="52" pageNumber="53">Fig. 205</figureCitation>
).
</paragraph>
<caption id="BEDE661AC31AC744E12E7121FBB5FA67" httpUri="https://zenodo.org/record/156454/files/figure.png" pageId="52" pageNumber="53" targetBox="[289,1303,584,1286]" targetPageId="52">
<paragraph id="EA1E3692C31AC744E12E7121FBB5FA67" blockId="52.[264,1322,1328,1455]" pageId="52" pageNumber="53">
<emphasis id="D8D5EA80C31AC744E12E7121FE5BFA80" bold="true" box="[264,481,1328,1352]" pageId="52" pageNumber="53">FIGURES 194­198</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C31AC744E2AC7121FC4CFA8F" authority="Gusarov" authorityName="Gusarov" box="[650,1014,1328,1351]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis" status="sp. nov.">
<emphasis id="D8D5EA80C31AC744E2AC7121FC2BFA8F" box="[650,913,1328,1351]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C31AC744E4247121FBFAFA80" bold="true" box="[1026,1088,1328,1352]" pageId="52" pageNumber="53">
<taxonomicNameLabel id="C3E657FBC31AC744E4247121FBFAFA80" box="[1026,1088,1328,1352]" pageId="52" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(194, 196­198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194­195 – median lobe, parameral view; 196 – apex of median lobe, parameral view; 197 – median lobe, lateral view; 198 – apex of median lobe, lateral view. Scale bar 0.2 mm (194­195, 197), 0.1 mm (196, 198).
</paragraph>
</caption>
<paragraph id="EA1E3692C31AC744E11E7211FC04F98A" blockId="52.[264,1324,1536,1882]" pageId="52" pageNumber="53">
<taxonomicName id="2DA14D11C31AC744E11E7211FDECF9D1" box="[312,598,1536,1561]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C31AC744E11E7211FDECF9D1" box="[312,598,1536,1561]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C31AC744E2D57211FC17F9D1" box="[755,941,1536,1561]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="furcativentris">
<emphasis id="D8D5EA80C31AC744E2D57211FC17F9D1" box="[755,941,1536,1561]" italics="true" pageId="52" pageNumber="53">S. furcativentris</emphasis>
</taxonomicName>
in having parallel­sided apex of median lobe (in parameral view) (
<figureCitation id="729A2A17C31AC744E2B17239FC84F98A" box="[663,830,1576,1602]" captionStart="FIGURES 194 ­ 198" captionStartId="52.[264,383,1328,1352]" captionTargetBox="[289,1303,584,1286]" captionTargetId="figure@52.[272,1315,569,1301]" captionTargetPageId="52" captionText="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 – median lobe, parameral view; 196 – apex of median lobe, parameral view; 197 – median lobe, lateral view; 198 – apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." httpUri="https://zenodo.org/record/156454/files/figure.png" pageId="52" pageNumber="53">Figs. 194­196</figureCitation>
; 212­213).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C31AC746E11E7241FDC4FB06" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53" type="description">
<paragraph id="EA1E3692C31AC744E11E7241FC89F95A" blockId="52.[264,1324,1536,1882]" pageId="52" pageNumber="53">
<taxonomicName id="2DA14D11C31AC744E11E7241FDE0F9A1" box="[312,602,1616,1641]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C31AC744E11E7241FDE0F9A1" box="[312,602,1616,1641]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C31AC744E3267241FC39F9A1" box="[768,899,1616,1641]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="D8D5EA80C31AC744E3267241FC39F9A1" box="[768,899,1616,1641]" italics="true" pageId="52" pageNumber="53">S. brunnea</emphasis>
</taxonomicName>
in having straight apex of median lobe (in lateral view) (
<figureCitation id="729A2A17C31AC744E22B7269FD0EF95A" box="[525,692,1656,1682]" captionStart="FIGURES 194 ­ 198" captionStartId="52.[264,383,1328,1352]" captionTargetBox="[289,1303,584,1286]" captionTargetId="figure@52.[272,1315,569,1301]" captionTargetPageId="52" captionText="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 – median lobe, parameral view; 196 – apex of median lobe, parameral view; 197 – median lobe, lateral view; 198 – apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." httpUri="https://zenodo.org/record/156454/files/figure.png" pageId="52" pageNumber="53">Figs. 197­198</figureCitation>
; 241­242).
</paragraph>
<paragraph id="EA1E3692C31AC744E11E72B1FCD1F8C2" blockId="52.[264,1324,1536,1882]" pageId="52" pageNumber="53">
<taxonomicName id="2DA14D11C31AC744E11E72B1FDECF971" box="[312,598,1696,1721]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C31AC744E11E72B1FDECF971" box="[312,598,1696,1721]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C31AC744E2D672B1FC2CF971" box="[752,918,1696,1721]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="D8D5EA80C31AC744E2D672B1FC2CF971" box="[752,918,1696,1721]" italics="true" pageId="52" pageNumber="53">S. brevipennis</emphasis>
</taxonomicName>
in having matte pronotum; pronotal punctation not asperate; elytra longer than pronotum; apex of median lobe parallelsided (in parameral view) (
<figureCitation id="729A2A17C31AC744E26372E1FD56F8C2" box="[581,748,1776,1802]" captionStart="FIGURES 194 ­ 198" captionStartId="52.[264,383,1328,1352]" captionTargetBox="[289,1303,584,1286]" captionTargetId="figure@52.[272,1315,569,1301]" captionTargetPageId="52" captionText="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 – median lobe, parameral view; 196 – apex of median lobe, parameral view; 197 – median lobe, lateral view; 198 – apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." httpUri="https://zenodo.org/record/156454/files/figure.png" pageId="52" pageNumber="53">Figs. 194­196</figureCitation>
; 226­227).
</paragraph>
<paragraph id="EA1E3692C31AC745E11E7309FB51FE4E" blockId="52.[264,1324,1536,1882]" lastBlockId="53.[264,1324,284,390]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<taxonomicName id="2DA14D11C31AC744E11E7309FDE6F8F9" box="[312,604,1816,1841]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C31AC744E11E7309FDE6F8F9" box="[312,604,1816,1841]" italics="true" pageId="52" pageNumber="53">Seeversiella scabricollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C31AC744E32D7309FC09F8F9" box="[779,947,1816,1841]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="D8D5EA80C31AC744E32D7309FC09F8F9" box="[779,947,1816,1841]" italics="true" pageId="52" pageNumber="53">S. curtipennis</emphasis>
</taxonomicName>
in having larger eyes; longer elytra; wings longer than elytra; posterior margin of male tergum 8 with medial emargination (
<figureCitation id="729A2A17C31BC745E175750DFE00FEFE" box="[339,442,284,310]" captionStart="FIGURES 190 ­ 193" captionStartId="51.[264,383,1774,1798]" captionTargetBox="[282,1304,939,1740]" captionTargetId="figure@51.[275,1312,929,1748]" captionTargetPageId="51" captionText="FIGURES 190 ­ 193. Abdominal segment 8 of Seeversiella scabricollis Gusarov, sp. n. (190 ­ 191, holotype; 192 ­ 193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 – male tergum 8; 191 – male sternum 8; 192 – female tergum 8; 193 – female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156453/files/figure.png" pageId="53" pageNumber="54">Fig. 190</figureCitation>
); narrow apex of the median lobe (in parameral view) (
<figureCitation id="729A2A17C31BC745E464750DFB53FEFE" box="[1090,1257,284,310]" captionStart="FIGURES 194 ­ 198" captionStartId="52.[264,383,1328,1352]" captionTargetBox="[289,1303,584,1286]" captionTargetId="figure@52.[272,1315,569,1301]" captionTargetPageId="52" captionText="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 – median lobe, parameral view; 196 – apex of median lobe, parameral view; 197 – median lobe, lateral view; 198 – apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." httpUri="https://zenodo.org/record/156454/files/figure.png" pageId="53" pageNumber="54">Figs. 194­196</figureCitation>
; 251­ 252); short medial lamellae of the internal sac (
<figureCitation id="729A2A17C31BC745E3687555FC79FE96" box="[846,963,324,350]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="53" pageNumber="54">Figs. 201</figureCitation>
; 258); lateral diverticula with sclerotized denticles (
<figureCitation id="729A2A17C31BC745E220757DFD09FE4E" box="[518,691,364,390]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="53" pageNumber="54">Figs. 200, 207</figureCitation>
; 259); and shorter spermatheca (
<figureCitation id="729A2A17C31BC745E417757DFB18FE4E" box="[1073,1186,364,390]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="53" pageNumber="54">Figs. 205</figureCitation>
,
<figureCitation id="729A2A17C31BC745E488757DFB65FE4E" box="[1198,1247,364,390]" captionStart="FIGURES 251 ­ 255" captionStartId="64.[264,383,1694,1718]" captionTargetBox="[284,1301,785,1656]" captionTargetId="figure@64.[270,1316,769,1667]" captionTargetPageId="64" captionText="FIGURES 251 ­ 255. Genitalia of Seeversiella curtipennis Gusarov, sp. n. (paratypes, Volcán Irazú, Costa Rica). 251 – median lobe, parameral view; 252 – apex of median lobe, parameral view; 253 – median lobe, lateral view; 254 – apex of median lobe, lateral view; 255 ­ spermatheca. Scale bar 0.2 mm (251, 253), 0.1 mm (252, 254 ­ 255)." httpUri="https://zenodo.org/record/156464/files/figure.png" pageId="53" pageNumber="54">255</figureCitation>
).
</paragraph>
<caption id="BEDE661AC31BC745E12E7277FB09F885" httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="53" pageNumber="54" targetBox="[306,1280,457,1604]" targetPageId="53">
<paragraph id="EA1E3692C31BC745E12E7277FB09F885" blockId="53.[264,1323,1638,1904]" pageId="53" pageNumber="54">
<emphasis id="D8D5EA80C31BC745E12E7277FE58F9B6" bold="true" box="[264,482,1638,1662]" pageId="53" pageNumber="54">FIGURES 199­207</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C31BC745E2AE7277FC4EF9B5" authority="Gusarov" authorityName="Gusarov" box="[648,1012,1638,1661]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="scabricollis" status="sp. nov.">
<emphasis id="D8D5EA80C31BC745E2AE7277FC35F9B5" box="[648,911,1638,1661]" italics="true" pageId="53" pageNumber="54">Seeversiella scabricollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C31BC745E4247277FBF8F9B6" bold="true" box="[1026,1090,1638,1662]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="C3E657FBC31BC745E4247277FBF8F9B6" box="[1026,1090,1638,1662]" pageId="53" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(199, 204, holotype; 200­203, 205­207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199­200, 203­204), 0.1 mm (201­202, 205­207).
</paragraph>
</caption>
<paragraph id="EA1E3692C31BC745E12E7348FC0CF8B8" blockId="53.[264,1323,1638,1904]" box="[264,950,1881,1904]" pageId="53" pageNumber="54">CP – copulatory piece; LD – lateral diverticulum of internal sac.</paragraph>
<paragraph id="EA1E3692C318C746E11E750AFE2EFE96" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
<emphasis id="D8D5EA80C318C746E11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="54" pageNumber="55">Description.</emphasis>
Length
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. Body dark brown, with brown elytra, legs and mouthparts.
</paragraph>
<paragraph id="EA1E3692C318C746E11E757DFCE9FE36" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">Head surface matte, with fine isodiametric microsculpture, fine and slightly asperate punctation, distance between punctures equals 2 times their diameter. Temple length to eye length ratio 0.9­1.1. Antennal article 2 longer than article 3, article 4 subquadrate, 5 slightly transverse, 6­10 transverse (ratio 1.5­2.0).</paragraph>
<paragraph id="EA1E3692C318C746E11E761DFBE5FCDE" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C318C746E40D761DFB6CFDEE" box="[1067,1238,524,550]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.4" metricValueMin="4.9" pageId="54" pageNumber="55" unit="mm" value="0.515" valueMax="0.54" valueMin="0.49">0.49­0.54 mm</quantity>
, length
<quantity id="2D599B77C318C746E12E7625FE0BFD86" box="[264,433,564,590]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.6" metricValueMin="4.1" pageId="54" pageNumber="55" unit="mm" value="0.435" valueMax="0.46" valueMin="0.41">0.41­0.46 mm</quantity>
, width to length ratio 1.2, surface matte, with fine isodiametric microsculpture; punctation fine and weak, distance between punctures equal to their diameter. Elytra wider and longer (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 0.8), 1.3 times as wide as long, glossy, with fine and weak isodiametric microsculpture; with fine punctation, distance between punctures equal to ½­1 time their diameter. Wings fully developed or partially reduced, longer than elytra.
</paragraph>
<paragraph id="EA1E3692C318C746E11E7735FBBDFC7E" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
Abdominal terga glossy, with fine microsculpture consisting of strongly transverse meshes, with fine punctation, distance between punctures equals 2­4 times their diameter on terga 3­5 and 4­7 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C318C746E36B778DFC40FC7E" box="[845,1018,924,950]" captionStart="FIGURES 190 ­ 193" captionStartId="51.[264,383,1774,1798]" captionTargetBox="[282,1304,939,1740]" captionTargetId="figure@51.[275,1312,929,1748]" captionTargetPageId="51" captionText="FIGURES 190 ­ 193. Abdominal segment 8 of Seeversiella scabricollis Gusarov, sp. n. (190 ­ 191, holotype; 192 ­ 193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 – male tergum 8; 191 – male sternum 8; 192 – female tergum 8; 193 – female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156453/files/figure.png" pageId="54" pageNumber="55">Figs. 190, 192</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C318C746E11E77D5FC99FB9E" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
In males posterior angles of tergum 3 projecting as spines up to 3 times as long as tergum 3 (measured medially); tergum 7 with medial carina along midline, the carina up to 2/ 3 as long as tergum length; posterior margin of tergum 8 with medial emargination delimited by short dentiform projections (
<figureCitation id="729A2A17C318C746E289702DFCACFB9E" box="[687,790,1084,1110]" captionStart="FIGURES 190 ­ 193" captionStartId="51.[264,383,1774,1798]" captionTargetBox="[282,1304,939,1740]" captionTargetId="figure@51.[275,1312,929,1748]" captionTargetPageId="51" captionText="FIGURES 190 ­ 193. Abdominal segment 8 of Seeversiella scabricollis Gusarov, sp. n. (190 ­ 191, holotype; 192 ­ 193, paratype, Sendero a Cerro Chirripó, Costa Rica). 190 – male tergum 8; 191 – male sternum 8; 192 – female tergum 8; 193 – female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156453/files/figure.png" pageId="54" pageNumber="55">Fig. 190</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C318C746E11E7075FC19FB6E" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
Aedeagus as in
<figureCitation id="729A2A17C318C746E2247075FC9FFBB6" box="[514,805,1124,1150]" captionStart-0="FIGURES 194 ­ 198" captionStart-1="FIGURES 199 ­ 207" captionStartId-0="52.[264,383,1328,1352]" captionStartId-1="53.[264,383,1638,1662]" captionTargetBox-0="[289,1303,584,1286]" captionTargetBox-1="[306,1280,457,1604]" captionTargetId-0="figure@52.[272,1315,569,1301]" captionTargetId-1="figure@53.[299,1288,449,1611]" captionTargetPageId-0="52" captionTargetPageId-1="53" captionText-0="FIGURES 194 ­ 198. Aedeagus of Seeversiella scabricollis Gusarov, sp. n. (194, 196 ­ 198, holotype; 195, paratype, San José de la Montana, Costa Rica). 194 ­ 195 – median lobe, parameral view; 196 – apex of median lobe, parameral view; 197 – median lobe, lateral view; 198 – apex of median lobe, lateral view. Scale bar 0.2 mm (194 ­ 195, 197), 0.1 mm (196, 198)." captionText-1="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri-0="https://zenodo.org/record/156454/files/figure.png" httpUri-1="https://zenodo.org/record/156455/files/figure.png" pageId="54" pageNumber="55">Figs. 194­204, 206­207</figureCitation>
. Apex of paramere broad, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C318C746E309709DFC2CFB6E" box="[815,918,1164,1190]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="54" pageNumber="55">Fig. 202</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C318C746E11E70A5FDC4FB06" blockId="54.[264,1324,283,1350]" box="[312,638,1204,1230]" pageId="54" pageNumber="55">
Spermatheca as in
<figureCitation id="729A2A17C318C746E23370A5FDC1FB06" box="[533,635,1204,1230]" captionStart="FIGURES 199 ­ 207" captionStartId="53.[264,383,1638,1662]" captionTargetBox="[306,1280,457,1604]" captionTargetId="figure@53.[299,1288,449,1611]" captionTargetPageId="53" captionText="FIGURES 199 ­ 207. Genitalia of Seeversiella scabricollis Gusarov, sp. n. (199, 204, holotype; 200 ­ 203, 205 ­ 207, paratypes, sendero a Cerro Chirripó, Costa Rica). 199 – details of internal sac retracted into median lobe, abparameral view; 200 – everted internal sac, parameral view; 201 – medial lamellae, parameral view; 202 – apex of left paramere; 203 – everted internal sac, lateral view; 204 – details of internal sac retracted into median lobe, lateral view; 205 – spermatheca; 206 – right distal sclerite of internal sac; 207 – left lateral diverticulum of internal sac, with denticulate sclerotized portion. Scale bar 0.2 mm (199 ­ 200, 203 ­ 204), 0.1 mm (201 ­ 202, 205 ­ 207)." httpUri="https://zenodo.org/record/156455/files/figure.png" pageId="54" pageNumber="55">Fig. 205</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C318C746E11E70CAFC21FA8E" pageId="54" pageNumber="55" type="distribution">
<paragraph id="EA1E3692C318C746E11E70CAFBA5FB3E" blockId="54.[264,1324,283,1350]" box="[312,1055,1243,1270]" pageId="54" pageNumber="55">
<emphasis id="D8D5EA80C318C746E11E70CAFE62FB3D" bold="true" box="[312,472,1243,1269]" pageId="54" pageNumber="55">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C318C746E2A370CDFCBFFB3E" box="[645,773,1244,1270]" name="Costa Rica" pageId="54" pageNumber="55">Costa Rica</collectingCountry>
and
<collectingCountry id="92B67602C318C746E31870CDFC21FB3E" box="[830,923,1244,1270]" name="Panama" pageId="54" pageNumber="55">Panama</collectingCountry>
(
<figureCitation id="729A2A17C318C746E38D70CDFBA9FB3E" box="[939,1043,1244,1270]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="54" pageNumber="55">Fig. 388</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C318C746E11E7112FC21FA8E" blockId="54.[264,1324,283,1350]" pageId="54" pageNumber="55">
<emphasis id="D8D5EA80C318C746E11E7112FDBCFAD5" bold="true" box="[312,518,1283,1309]" pageId="54" pageNumber="55">Natural History.</emphasis>
<taxonomicName id="2DA14D11C318C746E2317115FC8DFAD5" box="[535,823,1284,1309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="scabricollis">
<emphasis id="D8D5EA80C318C746E2317115FC8DFAD5" box="[535,823,1284,1309]" italics="true" pageId="54" pageNumber="55">Seeversiella scabricollis</emphasis>
</taxonomicName>
was collected in leaf litter and with flight intercept and malaise traps, at altitudes of
<quantity id="2D599B77C318C746E2D1713DFC2DFA8E" box="[759,919,1324,1350]" metricMagnitude="3" metricUnit="m" metricValue="2.375" metricValueMax="3.3" metricValueMin="1.45" pageId="54" pageNumber="55" unit="m" value="2375.0" valueMax="3300.0" valueMin="1450.0">1450­3300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C195F2F9F8657ED8F0160998A3987E3D">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C327C779E12E750AFCF3FEFE" box="[264,841,283,310]" pageId="9" pageNumber="10" type="nomenclature">
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Key for identification of species of
<taxonomicName id="2DA14D11C327C779E29F750DFCF3FEFE" ID-CoL="8VS3L" box="[697,841,284,310]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C327C779E29F750DFCF3FEFE" bold="true" box="[697,841,284,310]" italics="true" pageId="9" pageNumber="10">Seeversiella</emphasis>
</taxonomicName>
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</heading>
</paragraph>
</subSubSection>
<keyStep id="51552E37C327C779E12E7576FA91FE31" pageId="9" pageNumber="10">
<subSubSection id="A2BB6519C327C779E12E7576FA96FE61" pageId="9" pageNumber="10" type="key">
<paragraph id="EA1E3692C327C779E12E7576FA96FE61" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7576FA96FE61" pageId="9" pageNumber="10">1 Elytra longer or only a little shorter than pronotum. Pronotal length to elytral length (measured from humeral angle) ratio 1.1 or less ......................................................... 2</keyLead>
</paragraph>
</subSubSection>
<paragraph id="EA1E3692C327C779E12E75A6FA91FE31" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E75A6FA91FE31" pageId="9" pageNumber="10">
– Elytra much shorter than pronotum (
<figureCitation id="729A2A17C327C779E2D075A6FC3EFE19" box="[758,900,439,465]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="9" pageNumber="10">Figs. 18­19</figureCitation>
). Pronotal length to elytral length (measured from humeral angle) ratio more than 1.1 ................................................. 28
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E7616FA91FD81" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E7616FA96FDE9" blockId="9.[264,1327,359,1904]" box="[264,1324,519,545]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7616FA96FDE9" box="[264,1324,519,545]" pageId="9" pageNumber="10">2 Pronotum glossy, with weak isodiametric microsculpture or without microsculpture 3</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E763EFA91FD81" blockId="9.[264,1327,359,1904]" box="[264,1323,559,585]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E763EFA91FD81" box="[264,1323,559,585]" pageId="9" pageNumber="10">– Pronotum matte, with strong isodiametric microsculpture ....................................... 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E7646FA91FC41" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E7646FA91FCD8" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7646FA91FCD8" pageId="9" pageNumber="10">
3 Punctation of pronotum asperate. Tergum 8 with three pairs of macrosetae (
<figureCitation id="729A2A17C327C779E4907646FED3FD51" captionStart="FIGURES 222 ­ 225" captionStartId="58.[264,382,1219,1243]" captionTargetBox="[284,1303,333,1185]" captionTargetId="figure@58.[273,1314,320,1192]" captionTargetPageId="58" captionText="FIGURES 222 ­ 225. Abdominal segment 8 of Seeversiella brevipennis Gusarov, sp. n. (222 ­ 223, holotype; 224 ­ 225, paratype, 5.9 km N Cerro Punta, Panama). 222 – male tergum 8; 223 – male sternum 8; 224 – female tergum 8; 225 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156458/files/figure.png" pageId="9" pageNumber="10">Figs. 222, 224</figureCitation>
). Proximal seta of the apex of paramere is approximately as long as the other three setae (
<figureCitation id="729A2A17C327C779E1EA76B6FD8DFD09" box="[460,567,679,705]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="9" pageNumber="10">Fig. 233</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C327C779E2CB76B6FC2DFD09" box="[749,919,679,705]" captionStart-0="FIGURES 226 ­ 229" captionStart-1="FIGURES 230 ­ 234" captionStartId-0="59.[264,383,1174,1198]" captionStartId-1="60.[264,382,1729,1753]" captionTargetBox-0="[307,1283,459,1141]" captionTargetBox-1="[294,1292,824,1693]" captionTargetId-0="figure@59.[296,1291,449,1147]" captionTargetId-1="figure@60.[281,1305,809,1702]" captionTargetPageId-0="59" captionTargetPageId-1="60" captionText-0="FIGURES 226 ­ 229. Aedeagus of Seeversiella brevipennis Gusarov, sp. n. (holotype). 226 – median lobe, parameral view; 227 – apex of median lobe, parameral view; 228 – median lobe, lateral view; 229 – apex of median lobe, lateral view. Scale bar 0.2 mm (226, 228), 0.1 mm (227, 229)." captionText-1="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri-0="https://zenodo.org/record/156459/files/figure.png" httpUri-1="https://zenodo.org/record/156460/files/figure.png" pageId="9" pageNumber="10">Figs. 226­233</figureCitation>
. Spermatheca:
<figureCitation id="729A2A17C327C779E44E76B6FB68FD09" box="[1128,1234,679,705]" captionStart="FIGURES 230 ­ 234" captionStartId="60.[264,382,1729,1753]" captionTargetBox="[294,1292,824,1693]" captionTargetId="figure@60.[281,1305,809,1702]" captionTargetPageId="60" captionText="FIGURES 230 ­ 234. Genitalia of Seeversiella brevipennis Gusarov, sp. n. (230 ­ 233, holotype; 234, paratype, 5.9 km N Cerro Punta, Panama). 230 ­ 231 – details of internal sac retracted into median lobe, lateral view; 232 – details of internal sac retracted into median lobe, abparameral view; 233 – apex of left paramere; 234 – spermatheca. Scale bar 0.2 mm (230 ­ 232), 0.1 mm (233 ­ 234)." httpUri="https://zenodo.org/record/156460/files/figure.png" pageId="9" pageNumber="10">Fig. 234</figureCitation>
. Body length 2.6­3.0 mm, pronotal width
<quantity id="2D599B77C327C779E2F576DEFCC7FD21" box="[723,893,719,745]" metricMagnitude="-4" metricUnit="m" metricValue="5.45" metricValueMax="5.6" metricValueMin="5.3" pageId="9" pageNumber="10" unit="mm" value="0.545" valueMax="0.56" valueMin="0.53">0.53­0.56 mm</quantity>
. Known from Cerro Punta,
<collectingCountry id="92B67602C327C779E4EF76DEFA93FD21" box="[1225,1321,719,745]" name="Panama" pageId="9" pageNumber="10">Panama</collectingCountry>
, at altitude of
<quantity id="2D599B77C327C779E1F576E6FD96FCD9" box="[467,556,759,785]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="10" unit="m" value="2100.0">2100 m</quantity>
(
<figureCitation id="729A2A17C327C779E21A76E6FD1AFCD9" box="[572,672,759,785]" captionStart="FIGURE 388" captionStartId="98.[264,368,1798,1822]" captionTargetBox="[420,1168,317,1782]" captionTargetId="figure@98.[413,1175,313,1788]" captionTargetPageId="98" captionText="FIGURE 388. Geographical distribution of Scabricollis group in Costa Rica and Panama. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156490/files/figure.png" pageId="9" pageNumber="10">Fig. 388</figureCitation>
) .............................. 16.
<taxonomicName id="2DA14D11C327C779E3EA76E6FB67FCD9" authority="Gusarov" authorityName="Gusarov" box="[972,1245,759,785]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brevipennis" status="sp. nov.">
<emphasis id="D8D5EA80C327C779E3EA76E6FBC8FCD8" box="[972,1138,759,784]" italics="true" pageId="9" pageNumber="10">S. brevipennis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C327C779E4CF76E7FA91FCD8" bold="true" box="[1257,1323,758,784]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="C3E657FBC327C779E4CF76E7FA91FCD8" box="[1257,1323,758,784]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E770EFA91FC41" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E770EFA91FC41" pageId="9" pageNumber="10">
– Punctation of pronotum not asperate. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C327C779E4D7770EFE32FCA9" captionStart="FIGURES 20 ­ 23" captionStartId="15.[264,383,1258,1282]" captionTargetBox="[344,1240,531,1222]" captionTargetId="figure@15.[337,1250,524,1230]" captionTargetPageId="15" captionText="FIGURES 20 ­ 23. Abdominal segment 8 of Seeversiella globicollis (Bernhauer) (20 ­ 21, Cloudcroft, New Mexico; 22 ­ 23, Salmo, British Columbia). 20 – male tergum 8; 21 – male sternum 8; 22 – female tergum 8; 23 – female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156425/files/figure.png" pageId="9" pageNumber="10">Figs. 20, 22</figureCitation>
). Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C327C779E1A4777EFE6DFC41" box="[386,471,879,905]" captionStart="FIGURES 28 ­ 36" captionStartId="19.[264,383,1598,1622]" captionTargetBox="[297,1289,603,1577]" captionTargetId="figure@19.[285,1302,592,1587]" captionTargetPageId="19" captionText="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 – details of internal sac retracted into median lobe, abparameral view; 29 – everted internal sac, parameral view; 30 – medial lamellae, parameral view; 31 – right hook of internal sac; 32 – spermatheca; 33 – apex of left paramere; 34 ­ 35 – details of internal sac retracted into median lobe, lateral view; 36 – copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri="https://zenodo.org/record/156427/files/figure.png" pageId="9" pageNumber="10">Fig. 33</figureCitation>
)............................................................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E7786FA96FC11" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E7786FA96FC79" blockId="9.[264,1327,359,1904]" box="[264,1324,919,945]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7786FA96FC79" box="[264,1324,919,945]" pageId="9" pageNumber="10">4 Pronotum with weak isodiametric microsculpture ...................................................... 5</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E77AEFA96FC11" blockId="9.[264,1327,359,1904]" box="[264,1324,959,985]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E77AEFA96FC11" box="[264,1324,959,985]" pageId="9" pageNumber="10">– Pronotum without microsculpture ............................................................................. 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E77F6FA96FBE1" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E77F6FA91FBC9" blockId="9.[264,1327,359,1904]" box="[264,1323,999,1025]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E77F6FA91FBC9" box="[264,1323,999,1025]" pageId="9" pageNumber="10">
<specimenCount id="FCA7FD1BC327C779E12E77F6FEC5FBC9" box="[264,383,999,1025]" pageId="9" pageNumber="10" type="male">5 Males</specimenCount>
............................................................................................................................ 6
</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E701EFA96FBE1" blockId="9.[264,1327,359,1904]" box="[264,1324,1039,1065]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E701EFA96FBE1" box="[264,1324,1039,1065]" pageId="9" pageNumber="10">
– Females (Females of
<taxonomicName id="2DA14D11C327C779E215701EFD07FBE0" box="[563,701,1039,1064]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C327C779E215701EFD07FBE0" box="[563,701,1039,1064]" italics="true" pageId="9" pageNumber="10">S. nigriceps</emphasis>
</taxonomicName>
are unknown) ........................................................ 15
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E7027FA94FB00" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E7027FA96FBB0" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7027FA96FBB0" pageId="9" pageNumber="10">
6 Distal sclerites of internal sac strongly sclerotized, hook­shaped (
<figureCitation id="729A2A17C327C779E41D7027FB71FB98" box="[1083,1227,1078,1104]" captionStart="FIGURES 28 ­ 36" captionStartId="19.[264,383,1598,1622]" captionTargetBox="[297,1289,603,1577]" captionTargetId="figure@19.[285,1302,592,1587]" captionTargetPageId="19" captionText="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 – details of internal sac retracted into median lobe, abparameral view; 29 – everted internal sac, parameral view; 30 – medial lamellae, parameral view; 31 – right hook of internal sac; 32 – spermatheca; 33 – apex of left paramere; 34 ­ 35 – details of internal sac retracted into median lobe, lateral view; 36 – copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri="https://zenodo.org/record/156427/files/figure.png" pageId="9" pageNumber="10">Figs. 31, 35</figureCitation>
; 48, 51; DS) or dentiform (
<figureCitation id="729A2A17C327C779E234704FFD1BFBB0" box="[530,673,1118,1144]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="9" pageNumber="10">Figs. 63, 65</figureCitation>
; DS) 7
</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E7097FA94FB00" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7097FA94FB00" pageId="9" pageNumber="10">
– Distal sclerites of internal sac poorly sclerotized, not hook­shaped or dentiform and may have numerous spicules (
<figureCitation id="729A2A17C327C779E2BC70BFFD47FB00" box="[666,765,1198,1224]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="9" pageNumber="10">Figs. 87</figureCitation>
; 99, 101; 124; 146; DS) ............................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E70C7FA95FAD0" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E70C7FA95FB38" blockId="9.[264,1327,359,1904]" box="[264,1327,1238,1264]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E70C7FA95FB38" box="[264,1327,1238,1264]" pageId="9" pageNumber="10">
7 Distal sclerites of internal sac hook­shaped (
<figureCitation id="729A2A17C327C779E36670C7FC75FB38" box="[832,975,1238,1264]" captionStart="FIGURES 28 ­ 36" captionStartId="19.[264,383,1598,1622]" captionTargetBox="[297,1289,603,1577]" captionTargetId="figure@19.[285,1302,592,1587]" captionTargetPageId="19" captionText="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 – details of internal sac retracted into median lobe, abparameral view; 29 – everted internal sac, parameral view; 30 – medial lamellae, parameral view; 31 – right hook of internal sac; 32 – spermatheca; 33 – apex of left paramere; 34 ­ 35 – details of internal sac retracted into median lobe, lateral view; 36 – copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri="https://zenodo.org/record/156427/files/figure.png" pageId="9" pageNumber="10">Figs. 31, 35</figureCitation>
; 48, 51) ............................. 8
</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E70EFFA95FAD0" blockId="9.[264,1327,359,1904]" box="[264,1327,1278,1304]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E70EFFA95FAD0" box="[264,1327,1278,1304]" pageId="9" pageNumber="10">
– Distal sclerites of internal sac dentiform (
<figureCitation id="729A2A17C327C779E33870EFFC16FAD0" box="[798,940,1278,1304]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="9" pageNumber="10">Figs. 63, 65</figureCitation>
; 124) ...................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C779E12E7137FA90F8E8" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E7137FA91F9F8" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E7137FA91F9F8" pageId="9" pageNumber="10">
8 Body dark brown. Microsculpture on head and pronotum less distinct. Apex of median lobe in lateral view straight (
<figureCitation id="729A2A17C327C779E2DF715FFC3DFAA0" box="[761,903,1358,1384]" captionStart="FIGURES 24 ­ 27" captionStartId="18.[264,383,1768,1792]" captionTargetBox="[301,1288,1258,1750]" captionTargetId="figure@18.[287,1300,1249,1757]" captionTargetPageId="18" captionText="FIGURES 24 ­ 27. Aedeagus of Seeversiella globicollis (Bernhauer) (Cypress Hills Provincial Park, Alberta). 24 – median lobe, parameral view; 25 – apex of median lobe, parameral view; 26 – median lobe, lateral view; 27 – apex of median lobe, lateral view. Scale bar 0.2 mm (24, 26), 0.1 mm (25, 27)." httpUri="https://zenodo.org/record/156426/files/figure.png" pageId="9" pageNumber="10">Figs. 26­27</figureCitation>
). Copulatory piece of internal sac long (
<figureCitation id="729A2A17C327C779E1A77167FDA3FA58" box="[385,537,1398,1424]" captionStart="FIGURES 28 ­ 36" captionStartId="19.[264,383,1598,1622]" captionTargetBox="[297,1289,603,1577]" captionTargetId="figure@19.[285,1302,592,1587]" captionTargetPageId="19" captionText="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 – details of internal sac retracted into median lobe, abparameral view; 29 – everted internal sac, parameral view; 30 – medial lamellae, parameral view; 31 – right hook of internal sac; 32 – spermatheca; 33 – apex of left paramere; 34 ­ 35 – details of internal sac retracted into median lobe, lateral view; 36 – copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri="https://zenodo.org/record/156427/files/figure.png" pageId="9" pageNumber="10">Figs. 29, 34</figureCitation>
). Elytra (measured from humeral angle) longer than pronotum. Wings fully developed, 4 times as long as elytra. Aedeagus:
<figureCitation id="729A2A17C327C779E464718FFA9DFA70" box="[1090,1319,1438,1464]" captionStart-0="FIGURES 24 ­ 27" captionStart-1="FIGURES 28 ­ 36" captionStartId-0="18.[264,383,1768,1792]" captionStartId-1="19.[264,383,1598,1622]" captionTargetBox-0="[301,1288,1258,1750]" captionTargetBox-1="[297,1289,603,1577]" captionTargetId-0="figure@18.[287,1300,1249,1757]" captionTargetId-1="figure@19.[285,1302,592,1587]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURES 24 ­ 27. Aedeagus of Seeversiella globicollis (Bernhauer) (Cypress Hills Provincial Park, Alberta). 24 – median lobe, parameral view; 25 – apex of median lobe, parameral view; 26 – median lobe, lateral view; 27 – apex of median lobe, lateral view. Scale bar 0.2 mm (24, 26), 0.1 mm (25, 27)." captionText-1="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 – details of internal sac retracted into median lobe, abparameral view; 29 – everted internal sac, parameral view; 30 – medial lamellae, parameral view; 31 – right hook of internal sac; 32 – spermatheca; 33 – apex of left paramere; 34 ­ 35 – details of internal sac retracted into median lobe, lateral view; 36 – copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri-0="https://zenodo.org/record/156426/files/figure.png" httpUri-1="https://zenodo.org/record/156427/files/figure.png" pageId="9" pageNumber="10">Figs. 24­31, 33­36</figureCitation>
. Body length
<quantity id="2D599B77C327C779E1F471D7FDE5FA28" box="[466,607,1478,1504]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="3.3" metricValueMin="2.4" pageId="9" pageNumber="10" unit="mm" value="2.8499999999999996" valueMax="3.3" valueMin="2.4">2.4­3.3 mm</quantity>
, pronotal width
<quantity id="2D599B77C327C779E30771D7FC76FA28" box="[801,972,1478,1504]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.4" metricValueMin="4.4" pageId="9" pageNumber="10" unit="mm" value="0.49" valueMax="0.54" valueMin="0.44">0.44­0.54 mm</quantity>
. Widespread in North
<collectingCountry id="92B67602C327C779E4C671D7FEE7F9C0" name="United States of America" pageId="9" pageNumber="10">America</collectingCountry>
, known from
<collectingCountry id="92B67602C327C779E22671FFFDE2F9C0" box="[512,600,1518,1544]" name="Canada" pageId="9" pageNumber="10">Canada</collectingCountry>
to
<collectingCountry id="92B67602C327C779E25B71FFFD55F9C0" box="[637,751,1518,1544]" name="Honduras" pageId="9" pageNumber="10">Honduras</collectingCountry>
(
<figureCitation id="729A2A17C327C779E2D971FFFC1EF9C0" box="[767,932,1518,1544]" captionStart-0="FIGURE 385" captionStart-1="FIGURE 386" captionStart-2="FIGURE 387" captionStartId-0="95.[264,368,1458,1482]" captionStartId-1="96.[264,368,1354,1378]" captionStartId-2="97.[264,368,1861,1885]" captionTargetBox-0="[309,1276,654,1420]" captionTargetBox-1="[315,1274,412,1319]" captionTargetBox-2="[313,1275,771,1826]" captionTargetId-0="figure@95.[301,1286,649,1431]" captionTargetId-1="figure@96.[309,1279,409,1327]" captionTargetId-2="figure@97.[304,1282,766,1834]" captionTargetPageId-0="95" captionTargetPageId-1="96" captionTargetPageId-2="97" captionText-0="FIGURE 385. Geographical distribution of Seeversiella globicollis (Bernhauer)." captionText-1="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." captionText-2="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri-0="https://zenodo.org/record/156487/files/figure.png" httpUri-1="https://zenodo.org/record/156488/files/figure.png" httpUri-2="https://zenodo.org/record/156489/files/figure.png" pageId="9" pageNumber="10">Figs. 385­387</figureCitation>
) .................................................. ...............................................................................1.
<taxonomicName id="2DA14D11C327C779E3B87206FB76F9F8" authority="Bernhauer" authorityName="Bernhauer" box="[926,1228,1558,1584]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C327C779E3B87206FB8DF9F8" box="[926,1079,1559,1584]" italics="true" pageId="9" pageNumber="10">S. globicollis</emphasis>
(Bernhauer)
</taxonomicName>
(males)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C327C779E12E722FFA90F8E8" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E722FFA90F8E8" pageId="9" pageNumber="10">
– Body brown. Microsculpture on head and pronotum more distinct. Apex of median lobe in lateral view bent paramerally (
<figureCitation id="729A2A17C327C779E3277277FC35F948" box="[769,911,1638,1664]" captionStart="FIGURES 41 ­ 44" captionStartId="22.[264,383,1298,1322]" captionTargetBox="[293,1296,576,1260]" captionTargetId="figure@22.[281,1306,569,1271]" captionTargetPageId="22" captionText="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 – median lobe, parameral view; 42 – apex of median lobe, parameral view; 43 – median lobe, lateral view; 44 – apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." httpUri="https://zenodo.org/record/156429/files/figure.png" pageId="9" pageNumber="10">Figs. 43­44</figureCitation>
). Copulatory piece of internal sac short (
<figureCitation id="729A2A17C327C779E1A5729FFD80F960" box="[387,570,1678,1704]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="9" pageNumber="10">Figs. 46, 49­50</figureCitation>
). Elytra (measured from humeral angle) shorter than pronotum. Wings short, 2 times as long as elytra. Aedeagus:
<figureCitation id="729A2A17C327C779E3B872A7FB93F918" box="[926,1065,1718,1744]" captionStart-0="FIGURES 41 ­ 44" captionStart-1="FIGURES 45 ­ 53" captionStartId-0="22.[264,383,1298,1322]" captionStartId-1="23.[264,383,1314,1338]" captionTargetBox-0="[293,1296,576,1260]" captionTargetBox-1="[313,1274,323,1277]" captionTargetId-0="figure@22.[281,1306,569,1271]" captionTargetId-1="figure@23.[306,1280,313,1287]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 – median lobe, parameral view; 42 – apex of median lobe, parameral view; 43 – median lobe, lateral view; 44 – apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-1="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156429/files/figure.png" httpUri-1="https://zenodo.org/record/156430/files/figure.png" pageId="9" pageNumber="10">Figs. 41­52</figureCitation>
. Body length
<quantity id="2D599B77C327C779E4FE72A7FED3F930" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="9" pageNumber="10" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C327C779E20F72CFFD69F930" box="[553,723,1758,1784]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.9" metricValueMin="4.4" pageId="9" pageNumber="10" unit="mm" value="0.46499999999999997" valueMax="0.49" valueMin="0.44">0.44­0.49 mm</quantity>
. Known from the
<collectingCountry id="92B67602C327C779E39772CFFB8AF930" box="[945,1072,1758,1784]" name="Guadeloupe" pageId="9" pageNumber="10">Guadalupe</collectingCountry>
Mountains, Texas, at altitude of
<quantity id="2D599B77C327C779E1917317FDAAF8E8" box="[439,528,1798,1824]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="10" unit="m" value="2400.0">2400 m</quantity>
(
<figureCitation id="729A2A17C327C779E2397317FD39F8E8" box="[543,643,1798,1824]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="9" pageNumber="10">Fig. 386</figureCitation>
) ............................... 2.
<taxonomicName id="2DA14D11C327C779E3817316FBC7F8E8" authority="Gusarov" authorityName="Gusarov" box="[935,1149,1798,1824]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C327C779E3817316FBA8F8E8" box="[935,1042,1799,1824]" italics="true" pageId="9" pageNumber="10">S. texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C327C779E4AF7317FB76F8E8" bold="true" box="[1161,1228,1798,1824]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="C3E657FBC327C779E4AF7317FB76F8E8" box="[1161,1228,1798,1824]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C327C77AE12E733FFA91FE1E" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<paragraph id="EA1E3692C327C779E12E733FFA97F8B8" blockId="9.[264,1327,359,1904]" pageId="9" pageNumber="10">
<keyLead id="515095A7C327C779E12E733FFA97F8B8" pageId="9" pageNumber="10">
9 Distal sclerites of internal sac wider (
<figureCitation id="729A2A17C327C779E2D8733FFC29F880" box="[766,915,1838,1864]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="9" pageNumber="10">Figs. 63, 65</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C327C779E463733FFA92F880" box="[1093,1320,1838,1864]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="9" pageNumber="10">Figs. 58­61, 63­67</figureCitation>
. Body length
<quantity id="2D599B77C327C779E1F27347FDDBF8B8" box="[468,609,1878,1904]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="9" pageNumber="10" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C327C779E3007347FC68F8B8" box="[806,978,1878,1904]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="9" pageNumber="10" unit="mm" value="0.42" valueMax="0.44" valueMin="0.4">0.40­0.44 mm</quantity>
. Known from the Huachuca
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE11E750DFA91FE1E" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
Mountains, Arizona, at altitude of
<quantity id="2D599B77C324C77AE2EA750DFC9FFEFE" box="[716,805,284,310]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="10" pageNumber="11" unit="m" value="2150.0">2150 m</quantity>
(
<figureCitation id="729A2A17C324C77AE313750DFC22FEFE" box="[821,920,284,310]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="10" pageNumber="11">Fig. 386</figureCitation>
) ................................................... ........................................................ 3.
<taxonomicName id="2DA14D11C324C77AE2DF7555FB9EFE96" authority="Gusarov" authorityName="Gusarov" box="[761,1060,324,350]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sonomotoides" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE2DF7555FC02FE95" box="[761,952,324,349]" italics="true" pageId="10" pageNumber="11">S. sonomotoides</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4097552FBCBFE95" bold="true" box="[1071,1137,323,349]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4097552FBCBFE95" box="[1071,1137,323,349]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males, in part)
<keyLead id="515095A7C324C77AE12E757DFA91FE1E" pageId="10" pageNumber="11">
– Distal sclerites of internal sac narrower (
<figureCitation id="729A2A17C324C77AE33A757DFC3EFE4E" box="[796,900,364,390]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 – male tergum 8; 118 – male sternum 8; 119 – median lobe, parameral view; 120 – apex of median lobe, parameral view; 121 – median lobe, lateral view; 122 – apex of median lobe, lateral view; 123 ­ 124 – details of internal sac retracted into median lobe, lateral view; 125 – apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS – distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="10" pageNumber="11">Fig. 124</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE416757DFB6DFE4E" box="[1072,1239,364,390]" captionStart="FIGURES 117 ­ 125" captionStartId="36.[264,383,1644,1668]" captionTargetBox="[297,1299,423,1607]" captionTargetId="figure@36.[279,1307,409,1617]" captionTargetPageId="36" captionText="FIGURES 117 ­ 125. Abdominal segment 8 and aedeagus of Seeversiella nigriceps Gusarov, sp. n. (holotype). 117 – male tergum 8; 118 – male sternum 8; 119 – median lobe, parameral view; 120 – apex of median lobe, parameral view; 121 – median lobe, lateral view; 122 – apex of median lobe, lateral view; 123 ­ 124 – details of internal sac retracted into median lobe, lateral view; 125 – apex of left paramere. Scale bar 0.3 mm (117 ­ 118), 0.2 mm (119, 121, 123 ­ 124), 0.1 mm (120, 122, 125). DS – distal sclerite of internal sac." httpUri="https://zenodo.org/record/156441/files/figure.png" pageId="10" pageNumber="11">Figs. 119­125</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE1AF7585FE52FE66" box="[393,488,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="11" unit="mm" value="2.4">2.4 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE28F7585FCADFE66" box="[681,791,404,430]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="10" pageNumber="11" unit="mm" value="0.47">0.47 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE3EC7585FB9DFE66" box="[970,1063,404,430]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE4F47585FA96FE66" box="[1234,1324,404,430]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="11" unit="m" value="2400.0">2400 m</quantity>
(
<figureCitation id="729A2A17C324C77AE16775ADFE1FFE1E" box="[321,421,444,470]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
)...............................................8.
<taxonomicName id="2DA14D11C324C77AE30875ADFB99FE1E" authority="Gusarov" authorityName="Gusarov" box="[814,1059,444,470]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nigriceps" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE30875ADFC02FE1D" box="[814,952,444,469]" italics="true" pageId="10" pageNumber="11">S. nigriceps</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE40975AAFBCBFE1D" bold="true" box="[1071,1137,443,469]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE40975AAFBCBFE1D" box="[1071,1137,443,469]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males, in part)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E75F5FA97FDEE" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E75F5FA91FE36" blockId="10.[264,1326,284,1870]" box="[264,1323,484,510]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E75F5FA91FE36" box="[264,1323,484,510]" pageId="10" pageNumber="11">
10 Apex of median lobe in lateral view widened subapically (
<figureCitation id="729A2A17C324C77AE3C375F5FB31FE36" box="[997,1163,484,510]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="10" pageNumber="11">Figs. 112­113</figureCitation>
; 156­157) 11
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E761DFA97FDEE" blockId="10.[264,1326,284,1870]" box="[264,1325,524,550]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E761DFA97FDEE" box="[264,1325,524,550]" pageId="10" pageNumber="11">
– Apex of median lobe in lateral view not widened subapically (
<figureCitation id="729A2A17C324C77AE435761DFB20FDEE" box="[1043,1178,524,550]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="10" pageNumber="11">Figs. 85­86</figureCitation>
) ............. 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E7625FA96FCDE" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E7625FA90FD0E" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7625FA90FD0E" pageId="10" pageNumber="11">
11 Apex of median lobe in lateral view emarginate on parameral side (
<figureCitation id="729A2A17C324C77AE4527625FA9AFD86" box="[1140,1312,564,590]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="10" pageNumber="11">Figs. 156­157</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE1F6764DFDC7FDBE" box="[464,637,604,630]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="10" pageNumber="11">Figs. 154­159</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE364764DFC1FFDBE" box="[834,933,604,630]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="11" unit="mm" value="2.5">2.5 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE45E764DFA92FDBE" box="[1144,1320,604,630]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="10" pageNumber="11" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50­0.53 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE1F07695FDDBFD56" box="[470,609,644,670]" name="El Salvador" pageId="10" pageNumber="11">El Salvador</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE32E7695FCDBFD56" box="[776,865,644,670]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="11" unit="m" value="2100.0">2100 m</quantity>
(
<figureCitation id="729A2A17C324C77AE3577695FC6EFD56" box="[881,980,644,670]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ........................................... ................................................................................11.
<taxonomicName id="2DA14D11C324C77AE39476BDFBC7FD0E" authority="Gusarov" authorityName="Gusarov" box="[946,1149,684,710]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="badia" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE39476BDFBA8FD0D" box="[946,1042,684,709]" italics="true" pageId="10" pageNumber="11">S. badia</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4AF76BAFB76FD0D" bold="true" box="[1161,1228,683,709]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4AF76BAFB76FD0D" box="[1161,1228,683,709]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E76C5FA96FCDE" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E76C5FA96FCDE" pageId="10" pageNumber="11">
– Apex of median lobe in lateral view may be slightly bent paramerally (
<figureCitation id="729A2A17C324C77AE45B76C5FA99FD26" box="[1149,1315,724,750]" captionStart="FIGURES 139 ­ 142" captionStartId="40.[264,383,1421,1445]" captionTargetBox="[306,1279,781,1384]" captionTargetId="figure@40.[299,1288,769,1395]" captionTargetPageId="40" captionText="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="10" pageNumber="11">Figs. 141­142</figureCitation>
) but never clearly emarginate (
<figureCitation id="729A2A17C324C77AE2B076EDFC81FCDE" box="[662,827,764,790]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="10" pageNumber="11">Figs. 112­113</figureCitation>
; 141­142) ............................................ 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E7735FA97FC7E" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E7735FA90FC46" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7735FA90FC46" pageId="10" pageNumber="11">
12 Apex of median lobe in parameral view wide (
<figureCitation id="729A2A17C324C77AE3447735FBB3FCF6" box="[866,1033,804,830]" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="10" pageNumber="11">Figs. 110­111</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE4947735FEDCFCAE" captionStart="FIGURES 109 ­ 116" captionStartId="35.[264,383,1334,1358]" captionTargetBox="[306,1282,441,1310]" captionTargetId="figure@35.[293,1295,433,1320]" captionTargetPageId="35" captionText="FIGURES 109 ­ 116. Genitalia of Seeversiella tuberculicauda Gusarov, sp. n. (109 ­ 115, holotype; 116, paratype from Jalapa, Mexico). 109 – apex of left paramere; 110 – median lobe, parameral view; 111 – apex of median lobe, parameral view; 112 – median lobe, lateral view; 113 – apex of median lobe, lateral view; 114 ­ 115 – details of internal sac retracted into median lobe, lateral view; 116 – spermatheca. Scale bar 0.1 mm (109, 111, 113, 116), 0.2 mm (110, 112, 114 ­ 115)." httpUri="https://zenodo.org/record/156440/files/figure.png" pageId="10" pageNumber="11">Figs. 110­ 115</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE234775DFDD5FCAE" box="[530,623,844,870]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="11" unit="mm" value="2.4">2.4 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE308775DFC21FCAE" box="[814,923,844,870]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="10" pageNumber="11" unit="mm" value="0.49">0.49 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE46A775DFB13FCAE" box="[1100,1193,844,870]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE17E7765FE0BFC46" box="[344,433,884,910]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="11" unit="m" value="1300.0">1300 m</quantity>
(
<figureCitation id="729A2A17C324C77AE1E67765FD9EFC46" box="[448,548,884,910]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ............................... 7.
<taxonomicName id="2DA14D11C324C77AE3637765FBC4FC46" authority="Gusarov" authorityName="Gusarov" box="[837,1150,884,910]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tuberculicauda" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE3637765FBA8FC45" box="[837,1042,884,909]" italics="true" pageId="10" pageNumber="11">S. tuberculicauda</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4AF7762FB76FC45" bold="true" box="[1161,1228,883,909]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4AF7762FB76FC45" box="[1161,1228,883,909]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E778DFA97FC7E" blockId="10.[264,1326,284,1870]" box="[264,1325,924,950]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E778DFA97FC7E" box="[264,1325,924,950]" pageId="10" pageNumber="11">
– Apex of median lobe in parameral view narrow (
<figureCitation id="729A2A17C324C77AE353778DFBA6FC7E" box="[885,1052,924,950]" captionStart="FIGURES 139 ­ 142" captionStartId="40.[264,383,1421,1445]" captionTargetBox="[306,1279,781,1384]" captionTargetId="figure@40.[299,1288,769,1395]" captionTargetPageId="40" captionText="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="10" pageNumber="11">Figs. 139­140</figureCitation>
; 165­166) .............. 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E77D5FA90FB3E" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E77D5FA90FB9E" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E77D5FA90FB9E" pageId="10" pageNumber="11">
13 Apex of median lobe in parameral view with weak basal constriction (
<figureCitation id="729A2A17C324C77AE45077D5FAA5FC16" box="[1142,1311,964,990]" captionStart="FIGURES 139 ­ 142" captionStartId="40.[264,383,1421,1445]" captionTargetBox="[306,1279,781,1384]" captionTargetId="figure@40.[299,1288,769,1395]" captionTargetPageId="40" captionText="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." httpUri="https://zenodo.org/record/156444/files/figure.png" pageId="10" pageNumber="11">Figs. 139­140</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE1EE77FDFDCBFBCE" box="[456,625,1004,1030]" captionStart-0="FIGURES 139 ­ 142" captionStart-1="FIGURES 143 ­ 149" captionStartId-0="40.[264,383,1421,1445]" captionStartId-1="41.[264,383,1483,1507]" captionTargetBox-0="[306,1279,781,1384]" captionTargetBox-1="[298,1288,744,1449]" captionTargetId-0="figure@40.[299,1288,769,1395]" captionTargetId-1="figure@41.[289,1297,729,1456]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 139 ­ 142. Aedeagus Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo, Mexico). 139 – median lobe, parameral view; 140 – apex of median lobe, parameral view; 141 – median lobe, lateral view; 142 – apex of median lobe, lateral view. Scale bar 0.2 mm (139, 141), 0.1 mm (140, 142)." captionText-1="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri-0="https://zenodo.org/record/156444/files/figure.png" httpUri-1="https://zenodo.org/record/156445/files/figure.png" pageId="10" pageNumber="11">Figs. 139­148</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE30177FDFC0FFBCE" box="[807,949,1004,1030]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="10" pageNumber="11" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE45A77FDFA92FBCE" box="[1148,1320,1004,1030]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.3" metricValueMin="4.1" pageId="10" pageNumber="11" unit="mm" value="0.47" valueMax="0.53" valueMin="0.41">0.41­0.53 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE1F07005FD88FBE6" box="[470,562,1044,1070]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
, at altitudes of
<quantity id="2D599B77C324C77AE2C27005FC3BFBE6" box="[740,897,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.3" metricValueMin="1.7" pageId="10" pageNumber="11" unit="m" value="2000.0" valueMax="2300.0" valueMin="1700.0">1700­2300 m</quantity>
(
<figureCitation id="729A2A17C324C77AE3B77005FC4FFBE6" box="[913,1013,1044,1070]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ....................................... ........................................................................ 10.
<taxonomicName id="2DA14D11C324C77AE3A3702DFBC4FB9E" authority="Gusarov" authorityName="Gusarov" box="[901,1150,1084,1110]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE3A3702DFBA8FB9D" box="[901,1042,1084,1109]" italics="true" pageId="10" pageNumber="11">S. mexicana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4AF702AFB76FB9D" bold="true" box="[1161,1228,1083,1109]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4AF702AFB76FB9D" box="[1161,1228,1083,1109]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E7075FA90FB3E" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7075FA90FB3E" pageId="10" pageNumber="11">
– Apex of median lobe in parameral view gradually widened basally, not constricted (
<figureCitation id="729A2A17C324C77AE167709DFE50FB6E" box="[321,490,1164,1190]" captionStart="FIGURES 165 ­ 168" captionStartId="45.[264,383,1583,1607]" captionTargetBox="[302,1289,779,1549]" captionTargetId="figure@45.[292,1295,769,1556]" captionTargetPageId="45" captionText="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 – median lobe, parameral view; 166 – apex of median lobe, parameral view; 167 – median lobe, lateral view; 168 – apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." httpUri="https://zenodo.org/record/156449/files/figure.png" pageId="10" pageNumber="11">Figs. 165­166</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE2B5709DFC80FB6E" box="[659,826,1164,1190]" captionStart-0="FIGURES 165 ­ 168" captionStart-1="FIGURES 169 ­ 174" captionStartId-0="45.[264,383,1583,1607]" captionStartId-1="46.[264,383,1661,1685]" captionTargetBox-0="[302,1289,779,1549]" captionTargetBox-1="[288,1298,900,1628]" captionTargetId-0="figure@45.[292,1295,769,1556]" captionTargetId-1="figure@46.[282,1304,889,1634]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="FIGURES 165 ­ 168. Aedeagus of Seeversiella similis Gusarov, sp. n. (holotype). 165 – median lobe, parameral view; 166 – apex of median lobe, parameral view; 167 – median lobe, lateral view; 168 – apex of median lobe, lateral view. Scale bar 0.2 mm (165, 167), 0.1 mm (166, 168)." captionText-1="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri-0="https://zenodo.org/record/156449/files/figure.png" httpUri-1="https://zenodo.org/record/156450/files/figure.png" pageId="10" pageNumber="11">Figs. 165­173</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE3CE709DFBCEFB6E" box="[1000,1140,1164,1190]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="10" pageNumber="11" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE11E70A5FE58FB06" box="[312,482,1204,1230]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="10" pageNumber="11" unit="mm" value="0.51" valueMax="0.54" valueMin="0.48">0.48­0.54 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE2B470A5FCBDFB06" box="[658,775,1204,1230]" name="Honduras" pageId="10" pageNumber="11">Honduras</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE38970A5FBF6FB06" box="[943,1100,1204,1230]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="10" pageNumber="11" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">2000­2100 m</quantity>
(
<figureCitation id="729A2A17C324C77AE47A70A5FB7BFB06" box="[1116,1217,1204,1230]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ............ ............................................................................. 12.
<taxonomicName id="2DA14D11C324C77AE38E70CDFBC4FB3E" authority="Gusarov" authorityName="Gusarov" box="[936,1150,1244,1270]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE38E70CDFBA8FB3D" box="[936,1042,1244,1269]" italics="true" pageId="10" pageNumber="11">S. similis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4AF70CAFB76FB3D" bold="true" box="[1161,1228,1243,1269]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4AF70CAFB76FB3D" box="[1161,1228,1243,1269]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E7115FA91F9C6" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E7115FA90FAA6" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7115FA90FAA6" pageId="10" pageNumber="11">
14 Distal sclerites of internal sac have weak spicules (
<figureCitation id="729A2A17C324C77AE3B17115FC4BFAD6" box="[919,1009,1284,1310]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="10" pageNumber="11">Fig. 87</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE4BB7115FA9DFAD6" box="[1181,1319,1284,1310]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="10" pageNumber="11">Figs. 83­88</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE1F7713DFDE6FA8E" box="[465,604,1324,1350]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="10" pageNumber="11" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3­2.5 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE33A713DFC7CFA8E" box="[796,966,1324,1350]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="10" pageNumber="11" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE45E713DFB6FFA8E" box="[1144,1237,1324,1350]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE1B77145FD94FAA6" box="[401,558,1364,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="10" pageNumber="11" unit="m" value="1800.0" valueMax="1900.0" valueMin="1700.0">1700­1900 m</quantity>
(
<figureCitation id="729A2A17C324C77AE2187145FD18FAA6" box="[574,674,1364,1390]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ............................. 5.
<taxonomicName id="2DA14D11C324C77AE3917145FBC4FAA6" authority="Gusarov" authorityName="Gusarov" box="[951,1150,1364,1390]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fusca" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE3917145FBA8FAA5" box="[951,1042,1364,1389]" italics="true" pageId="10" pageNumber="11">S. fusca</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC324C77AE4AF7142FB76FAA5" box="[1161,1228,1363,1389]" pageId="10" pageNumber="11" rank="species">
<emphasis id="D8D5EA80C324C77AE4AF7142FB76FAA5" bold="true" box="[1161,1228,1363,1389]" pageId="10" pageNumber="11">sp. n.</emphasis>
</taxonomicNameLabel>
(males)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E716DFA91F9C6" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E716DFA91F9C6" pageId="10" pageNumber="11">
– Distal sclerites of internal sac have strong spicules (
<figureCitation id="729A2A17C324C77AE385716DFBFFFA5E" box="[931,1093,1404,1430]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="10" pageNumber="11">Figs. 99, 101</figureCitation>
). Aedeagus:
<figureCitation id="729A2A17C324C77AE4D7716DFE35FA76" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="10" pageNumber="11">Figs. 94­103</figureCitation>
. Body length
<quantity id="2D599B77C324C77AE21871B5FD71FA76" box="[574,715,1444,1470]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="10" pageNumber="11" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE3AB71B5FB83FA76" box="[909,1081,1444,1470]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="10" pageNumber="11" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE4C871B5FEE6FA2E" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
and
<collectingCountry id="92B67602C324C77AE1B271DDFDB3FA2E" box="[404,521,1484,1510]" name="Honduras" pageId="10" pageNumber="11">Honduras</collectingCountry>
, at altitude of
<quantity id="2D599B77C324C77AE29671DDFCF7FA2E" box="[688,845,1484,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.4" metricValueMin="1.5" pageId="10" pageNumber="11" unit="m" value="1950.0" valueMax="2400.0" valueMin="1500.0">1500­2400 m</quantity>
(
<figureCitation id="729A2A17C324C77AE37B71DDFC7BFA2E" box="[861,961,1484,1510]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ............................................................................................................................6.
<taxonomicName id="2DA14D11C324C77AE3AB71E5FBC4F9C6" authority="Gusarov" authorityName="Gusarov" box="[909,1150,1524,1550]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="castanea" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE3AB71E5FBA8F9C5" box="[909,1042,1524,1549]" italics="true" pageId="10" pageNumber="11">S. castanea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE4AF71E2FB76F9C5" bold="true" box="[1161,1228,1523,1549]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE4AF71E2FB76F9C5" box="[1161,1228,1523,1549]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(males)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E720DFA91F996" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E720DFA96F9FE" blockId="10.[264,1326,284,1870]" box="[264,1324,1564,1590]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E720DFA96F9FE" box="[264,1324,1564,1590]" pageId="10" pageNumber="11">
15 Spermatheca S­shaped (
<figureCitation id="729A2A17C324C77AE277720DFD7BF9FE" box="[593,705,1564,1590]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="10" pageNumber="11">Figs. 149</figureCitation>
,
<figureCitation id="729A2A17C324C77AE2E8720DFD41F9FE" box="[718,763,1564,1590]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="10" pageNumber="11">160</figureCitation>
) .................................................................... 16
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E7255FA91F996" blockId="10.[264,1326,284,1870]" box="[264,1323,1604,1630]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7255FA91F996" box="[264,1323,1604,1630]" pageId="10" pageNumber="11">
– Spermatheca L­, J­ or C­shaped (
<figureCitation id="729A2A17C324C77AE29D7255FCA7F996" box="[699,797,1604,1630]" captionStart="FIGURES 28 ­ 36" captionStartId="19.[264,383,1598,1622]" captionTargetBox="[297,1289,603,1577]" captionTargetId="figure@19.[285,1302,592,1587]" captionTargetPageId="19" captionText="FIGURES 28 ­ 36. Genitalia of Seeversiella globicollis (Bernhauer) (28, 34 ­ 35, Cypress Hills Provincial Park, Alberta; 29 ­ 31, 33, 36, Cloudcroft, New Mexico; 32, Salmo, British Columbia). 28 – details of internal sac retracted into median lobe, abparameral view; 29 – everted internal sac, parameral view; 30 – medial lamellae, parameral view; 31 – right hook of internal sac; 32 – spermatheca; 33 – apex of left paramere; 34 ­ 35 – details of internal sac retracted into median lobe, lateral view; 36 – copulatory piece. Scale bar 0.2 mm (28, 34 ­ 35), 0.1 mm (29 ­ 33, 36)." httpUri="https://zenodo.org/record/156427/files/figure.png" pageId="10" pageNumber="11">Figs. 32</figureCitation>
,
<figureCitation id="729A2A17C324C77AE30F7255FCF3F996" box="[809,841,1604,1630]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="10" pageNumber="11">53</figureCitation>
,
<figureCitation id="729A2A17C324C77AE3737255FCCFF996" box="[853,885,1604,1630]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="10" pageNumber="11">62</figureCitation>
,
<figureCitation id="729A2A17C324C77AE3A77255FC1BF996" box="[897,929,1604,1630]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="10" pageNumber="11">89</figureCitation>
,
<figureCitation id="729A2A17C324C77AE38B7255FC60F996" box="[941,986,1604,1630]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="10" pageNumber="11">104</figureCitation>
) ....................................... 18
</keyLead>
</paragraph>
</keyStep>
<keyStep id="51552E37C324C77AE12E727DFA97F8EE" pageId="10" pageNumber="11">
<paragraph id="EA1E3692C324C77AE12E727DFA91F91E" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E727DFA91F91E" pageId="10" pageNumber="11">
16 Umbilicus facing proximally (
<figureCitation id="729A2A17C324C77AE28F727DFCAEF94E" box="[681,788,1644,1670]" captionStart="FIGURES 143 ­ 149" captionStartId="41.[264,383,1483,1507]" captionTargetBox="[298,1288,744,1449]" captionTargetId="figure@41.[289,1297,729,1456]" captionTargetPageId="41" captionText="FIGURES 143 ­ 149. Genitalia of Seeversiella mexicana Gusarov, sp. n. (paratypes from Filo de Caballo (143, 145 ­ 148) Suchixtepec (144, 149), Mexico). 143 – everted internal sac, parameral view; 144 – medial lamellae, parameral view; 145 ­ 146 – details of internal sac retracted into median lobe, lateral view; 147 – copulatory piece, lateral view; 148 – apex of left paramere; 149 – spermatheca. Scale bar 0.2 mm (143, 145 ­ 146), 0.1 mm (144, 147 ­ 149)." httpUri="https://zenodo.org/record/156445/files/figure.png" pageId="10" pageNumber="11">Fig. 149</figureCitation>
). Body length
<quantity id="2D599B77C324C77AE3FD727DFBD1F94E" box="[987,1131,1644,1670]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="10" pageNumber="11" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE11E7285FE58F966" box="[312,482,1684,1710]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.3" metricValueMin="4.1" pageId="10" pageNumber="11" unit="mm" value="0.47" valueMax="0.53" valueMin="0.41">0.41­0.53 mm</quantity>
. Known from
<collectingCountry id="92B67602C324C77AE2B47285FD55F966" box="[658,751,1684,1710]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
, at altitudes of
<quantity id="2D599B77C324C77AE3847285FB85F966" box="[930,1087,1684,1710]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.3" metricValueMin="1.7" pageId="10" pageNumber="11" unit="m" value="2000.0" valueMax="2300.0" valueMin="1700.0">1700­2300 m</quantity>
(
<figureCitation id="729A2A17C324C77AE4697285FB0EF966" box="[1103,1204,1684,1710]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="10" pageNumber="11">Fig. 387</figureCitation>
) ............. ..................................................................... 10.
<taxonomicName id="2DA14D11C324C77AE34872ADFBD2F91E" authority="Gusarov" authorityName="Gusarov" box="[878,1128,1724,1750]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mexicana" status="sp. nov.">
<emphasis id="D8D5EA80C324C77AE34872ADFC41F91D" box="[878,1019,1724,1749]" italics="true" pageId="10" pageNumber="11">S. mexicana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C324C77AE45572AAFB0FF91D" bold="true" box="[1139,1205,1723,1749]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="C3E657FBC324C77AE45572AAFB0FF91D" box="[1139,1205,1723,1749]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(females)
</keyLead>
</paragraph>
<paragraph id="EA1E3692C324C77AE12E72F5FA97F8EE" blockId="10.[264,1326,284,1870]" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E72F5FA97F8EE" pageId="10" pageNumber="11">
– Umbilicus facing laterally (
<figureCitation id="729A2A17C324C77AE25D72F5FD56F936" box="[635,748,1764,1790]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="10" pageNumber="11">Figs. 160</figureCitation>
,
<figureCitation id="729A2A17C324C77AE2DE72F5FC92F936" box="[760,808,1764,1790]" captionStart="FIGURES 169 ­ 174" captionStartId="46.[264,383,1661,1685]" captionTargetBox="[288,1298,900,1628]" captionTargetId="figure@46.[282,1304,889,1634]" captionTargetPageId="46" captionText="FIGURES 169 ­ 174. Genitalia of Seeversiella similis Gusarov, sp. n. (171, 173, holotype; 169 ­ 170, 172, 174, paratypes from La Tigra, Honduras). 169 – everted internal sac, parameral view; 170 – medial lamellae, parameral view; 171 – apex of left paramere; 172 – everted internal sac, lateral view; 173 – details of internal sac retracted into median lobe, lateral view; 174 – spermatheca. Scale bar 0.2 mm (169, 172 ­ 173), 0.1 mm (170 ­ 171, 174)." httpUri="https://zenodo.org/record/156450/files/figure.png" pageId="10" pageNumber="11">174</figureCitation>
). Two closely related species which cannot be distinguished by females ....................................................................................... 17
</keyLead>
</paragraph>
</keyStep>
<paragraph id="EA1E3692C324C77BE12E7325FA91FE96" blockId="10.[264,1326,284,1870]" lastBlockId="11.[264,1325,284,1870]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<keyLead id="515095A7C324C77AE12E7325FA91F886" box="[264,1323,1844,1870]" pageId="10" pageNumber="11">
<keyStep id="51552E37C324C77AE12E7325FA91F886" box="[264,1323,1844,1870]" pageId="10" pageNumber="11">
17 Body length
<quantity id="2D599B77C324C77AE1F67325FD94F886" box="[464,558,1844,1870]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="11" unit="mm" value="2.5">2.5 mm</quantity>
, pronotal width
<quantity id="2D599B77C324C77AE2C87325FC22F886" box="[750,920,1844,1870]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="10" pageNumber="11" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50­0.53 mm</quantity>
. Spermatheca:
<figureCitation id="729A2A17C324C77AE47D7325FB7AF886" box="[1115,1216,1844,1870]" captionStart="FIGURES 154 ­ 160" captionStartId="43.[264,383,1715,1739]" captionTargetBox="[292,1295,611,1684]" captionTargetId="figure@43.[281,1307,601,1691]" captionTargetPageId="43" captionText="FIGURES 154 ­ 160. Genitalia of Seeversiella badia Gusarov, sp. n. (154 ­ 159, holotype; 160, paratype, 21.7 km NE Metapan, El Salvador). 154 – median lobe, parameral view; 155 – apex of median lobe, parameral view; 156 – median lobe, lateral view; 157 – apex of median lobe, lateral view; 158 – details of internal sac retracted into median lobe, lateral view; 159 – apex of left paramere; 160 – spermatheca. Scale bar 0.2 mm (154, 156, 158), 0.1 mm (155, 157, 159 ­ 160)." httpUri="https://zenodo.org/record/156447/files/figure.png" pageId="10" pageNumber="11">Fig. 160</figureCitation>
. Known
</keyStep>
</keyLead>
<subSection id="832E2F79C325C77BE11E750DFA91FE96" pageId="11" pageNumber="12" type="multiple">
from
<collectingCountry id="92B67602C325C77BE15E750DFDBEFEFE" box="[376,516,284,310]" name="El Salvador" pageId="11" pageNumber="12">El Salvador</collectingCountry>
, at altitude of
<quantity id="2D599B77C325C77BE28C750DFCB9FEFE" box="[682,771,284,310]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="12" unit="m" value="2100.0">2100 m</quantity>
(
<figureCitation id="729A2A17C325C77BE335750DFCCDFEFE" box="[787,887,284,310]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="11" pageNumber="12">Fig. 387</figureCitation>
) .....................................................................................................................................11.
<taxonomicName id="2DA14D11C325C77BE3BD7555FBD2FE96" authority="Gusarov" authorityName="Gusarov" box="[923,1128,324,350]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="badia" status="sp. nov.">
<emphasis id="D8D5EA80C325C77BE3BD7555FC46FE95" box="[923,1020,324,349]" italics="true" pageId="11" pageNumber="12">S. badia</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C325C77BE4557552FB0FFE95" bold="true" box="[1139,1205,323,349]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="C3E657FBC325C77BE4557552FB0FFE95" box="[1139,1205,323,349]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(females)
</subSection>
</paragraph>
</treatment>
</document>

407
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<document id="F91FBA367D17F0CB3F3FC9F01E09BF67" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="1175­5326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C331C751E12E76BAFE17FADE" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella castanea Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="34" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="32" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:title id="DB14E98F9B5B764BF4DF829AA1BD855E">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C331C76FE12E76BAFC22FD0D" box="[264,920,683,710]" pageId="31" pageNumber="32" type="key">
<paragraph id="EA1E3692C331C76FE12E76BAFC22FD0D" blockId="31.[264,920,683,710]" box="[264,920,683,710]" pageId="31" pageNumber="32">
<heading id="B15681FEC331C76FE12E76BAFC22FD0D" bold="true" box="[264,920,683,710]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="D8D5EA80C331C76FE12E76BAFC22FD0D" bold="true" box="[264,920,683,710]" pageId="31" pageNumber="32">
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<key id="9E3061A3C331C76FE12E76BAFC22FD0D" box="[264,920,683,710]" pageId="31" pageNumber="32">
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6.
<taxonomicName id="2DA14D11C331C76FE10076BDFD22FD0D" ID-CoL="6YBD3" authority="Gusarov" authorityName="Gusarov" box="[294,664,683,710]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="castanea" status="sp. nov.">
<emphasis id="D8D5EA80C331C76FE10076BDFD9CFD0E" bold="true" box="[294,550,684,710]" italics="true" pageId="31" pageNumber="32">Seeversiella castanea</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC331C76FE28076BAFD52FD0D" box="[678,744,683,709]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C331C76FE2D176BAFC2AFD0D" box="[759,912,683,709]" captionStart-0="FIGURES 90 ­ 93" captionStart-1="FIGURES 94 ­ 104" captionStartId-0="31.[264,383,1803,1827]" captionStartId-1="32.[264,383,1439,1463]" captionTargetBox-0="[290,1295,933,1772]" captionTargetBox-1="[296,1295,339,1401]" captionTargetId-0="figure@31.[281,1306,921,1779]" captionTargetId-1="figure@32.[282,1305,329,1412]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="FIGURES 90 ­ 93. Abdominal segment 8 of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (90 ­ 91) and Las Piedrecitas (92 ­ 93), Mexico). 90 – male tergum 8; 91 – male sternum 8; 92 – female tergum 8; 93 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri-0="https://zenodo.org/record/156437/files/figure.png" httpUri-1="https://zenodo.org/record/156438/files/figure.png" pageId="31" pageNumber="32">Figs. 90­104</figureCitation>
)
</keyLead>
</key>
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</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C331C751E12E76E7FD55FE49" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="EA1E3692C331C76FE12E76E7FE40FCA9" blockId="31.[264,1324,758,865]" pageId="31" pageNumber="32">
<emphasis id="D8D5EA80C331C76FE12E76E7FE7AFCD8" bold="true" box="[264,448,758,784]" pageId="31" pageNumber="32">
<typeStatus id="351A8830C331C76FE12E76E7FEFFFCD8" box="[264,325,758,784]" pageId="31" pageNumber="32">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C331C76FE1F576E6FDF9FCD9" box="[467,579,759,785]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C331C76FE25376E7FCC4FCD8" bold="true" box="[629,894,758,784]" pageId="31" pageNumber="32">
<collectingCountry id="92B67602C331C76FE25376E7FD4FFCD8" box="[629,757,758,784]" name="Mexico" pageId="31" pageNumber="32">MEXICO</collectingCountry>
: Chiapas:
</emphasis>
<quantity id="2D599B77C331C76FE3B776E6FC5FFCD9" box="[913,997,759,785]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="31" pageNumber="32" unit="km" value="2.8">2.8 km</quantity>
NW Las Piedrecitas,
<quantity id="2D599B77C331C76FE4C876E6FA96FCD9" box="[1262,1324,759,785]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="km" value="5.0">5 km</quantity>
NW
<collectingCountry id="92B67602C331C76FE16F770EFE4BFCF1" box="[329,497,799,825]" name="Saint Kitts and Nevis" pageId="31" pageNumber="32">San Cristobal</collectingCountry>
de las Casas,
<quantity id="2D599B77C331C76FE29F770EFCA7FCF1" box="[697,797,799,825]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="32" unit="m" value="2400.0">2400 m</quantity>
, oak and pine forest litter (R.Anderson),
<date id="9E1F1052C331C76FE12E7756FE3CFCA9" box="[264,390,839,865]" pageId="31" pageNumber="32" value="1991-09-20">20.ix.1991</date>
(
<collectionCode id="8CB0AE57C331C76FE1B37756FE57FCA9" box="[405,493,839,865]" pageId="31" pageNumber="32">KSEM</collectionCode>
).
</paragraph>
<caption id="BEDE661AC331C76FE12E731AFC26F8A0" httpUri="https://zenodo.org/record/156437/files/figure.png" pageId="31" pageNumber="32" targetBox="[290,1295,933,1772]" targetPageId="31">
<paragraph id="EA1E3692C331C76FE12E731AFC26F8A0" blockId="31.[264,1323,1803,1896]" pageId="31" pageNumber="32">
<emphasis id="D8D5EA80C331C76FE12E731AFE79F8EB" bold="true" box="[264,451,1803,1827]" pageId="31" pageNumber="32">FIGURES 90­93</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C331C76FE2CF731DFB88F8EA" authority="Gusarov" authorityName="Gusarov" box="[745,1074,1803,1827]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="castanea" status="sp. nov.">
<emphasis id="D8D5EA80C331C76FE2CF731DFC6AF8EB" box="[745,976,1804,1827]" italics="true" pageId="31" pageNumber="32">Seeversiella castanea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C331C76FE41B731AFBC0F8EB" bold="true" box="[1085,1146,1803,1827]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="C3E657FBC331C76FE41B731AFBC0F8EB" box="[1085,1146,1803,1827]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Rayon (90­91) and Las Piedrecitas (92­93), Mexico). 90 – male tergum 8; 91 – male sternum 8; 92 – female tergum 8; 93 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<caption id="BEDE661AC30EC750E12E718EFE4BF94E" httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="32" pageNumber="33" targetBox="[296,1295,339,1401]" targetPageId="32">
<paragraph id="EA1E3692C30EC750E12E718EFE4BF94E" blockId="32.[264,1323,1439,1704]" pageId="32" pageNumber="33">
<emphasis id="D8D5EA80C30EC750E12E718EFE6BFA7F" bold="true" box="[264,465,1439,1463]" pageId="32" pageNumber="33">FIGURES 94­104</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C30EC750E24D718EFC09FA7E" authority="Gusarov" authorityName="Gusarov" box="[619,947,1439,1462]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="castanea" status="sp. nov.">
<emphasis id="D8D5EA80C30EC750E24D718EFCE8FA7E" box="[619,850,1439,1462]" italics="true" pageId="32" pageNumber="33">Seeversiella castanea</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C30EC750E398718EFC40FA7F" bold="true" box="[958,1018,1439,1463]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="C3E657FBC30EC750E398718EFC40FA7F" box="[958,1018,1439,1463]" pageId="32" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Rayon (94­ 97, 99­102) and Las Piedrecitas (98, 103­104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98– everted internal sac, parameral view; 99 – right hook of internal sac; 100­101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100­101), 0.1 mm (95, 97, 99, 102­104).
</paragraph>
</caption>
<paragraph id="EA1E3692C30EC750E12E7280FDCDF960" blockId="32.[264,1323,1439,1704]" box="[264,631,1681,1704]" pageId="32" pageNumber="33">DS – distal sclerite of internal sac.</paragraph>
<paragraph id="EA1E3692C30EC751E11E72E8FD55FE49" blockId="32.[264,1324,1785,1886]" lastBlockId="33.[264,1324,283,1302]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<typeStatus id="351A8830C30EC750E11E72E8FE15F8DB" box="[312,431,1785,1811]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C30EC750E1E572E8FD70F8DB" bold="true" box="[451,714,1785,1811]" pageId="32" pageNumber="33">
<collectingCountry id="92B67602C30EC750E1E572E8FDF9F8DB" box="[451,579,1785,1811]" name="Mexico" pageId="32" pageNumber="33">MEXICO</collectingCountry>
: Chiapas:
</emphasis>
<specimenCount id="FCA7FD1BC30EC750E2FD72E8FCC4F8DB" box="[731,894,1785,1811]" pageId="32" pageNumber="33" type="generic">10 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C30EC750E47372E8FB05F8DB" box="[1109,1215,1785,1811]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
;
<specimenCount id="FCA7FD1BC30EC750E4EB72E8FEF3F8F1" pageId="32" pageNumber="33" type="generic">6 specimens</specimenCount>
, Yerbabuena Reserve,
<quantity id="2D599B77C30EC750E271730EFD13F8F1" box="[599,681,1823,1849]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="32" pageNumber="33" unit="km" value="2.1">2.1 km</quantity>
NW Pueblo Nuevo Solistahuacan,
<quantity id="2D599B77C30EC750E469730EFB16F8F1" box="[1103,1196,1823,1849]" metricMagnitude="3" metricUnit="m" metricValue="2.07" pageId="32" pageNumber="33" unit="m" value="2070.0">2070 m</quantity>
, liquidambar forest litter (R.Anderson),
<date id="9E1F1052C30EC750E2507355FD4CF896" box="[630,758,1860,1886]" pageId="32" pageNumber="33" value="1992-09-23">23.ix.1992</date>
;
<specimenCount id="FCA7FD1BC30EC750E3237355FC20F896" box="[773,922,1860,1886]" pageId="32" pageNumber="33" type="generic">6 specimens</specimenCount>
,
<quantity id="2D599B77C30EC750E38F7355FC47F896" box="[937,1021,1860,1886]" metricMagnitude="3" metricUnit="m" metricValue="8.9" pageId="32" pageNumber="33" unit="km" value="8.9">8.9 km</quantity>
E Rayon,
<quantity id="2D599B77C30EC750E4597355FB66F896" box="[1151,1244,1860,1886]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="m" value="1500.0">1500 m</quantity>
, cloud forest litter (R.Anderson),
<date id="9E1F1052C30FC751E261750DFD72FEFE" box="[583,712,284,310]" pageId="33" pageNumber="34" value="1991-09-19">19.ix.1991</date>
;
<emphasis id="D8D5EA80C30FC751E2F0750AFC47FEFD" bold="true" box="[726,1021,283,309]" pageId="33" pageNumber="34">
<collectingCountry id="92B67602C30FC751E2F0750AFC38FEFD" box="[726,898,283,309]" name="Honduras" pageId="33" pageNumber="34">HONDURAS</collectingCountry>
: La Paz:
</emphasis>
2, 4, Cord. Montecillos,
<quantity id="2D599B77C30FC751E1487550FE06FE93" box="[366,444,321,347]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="33" pageNumber="34" unit="km" value="18.0">18 km</quantity>
NW La Paz,
<geoCoordinate id="8F955055C30FC751E2407550FD7DFE93" box="[614,711,321,347]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="14.35">14°21'N</geoCoordinate>
<geoCoordinate id="8F955055C30FC751E2F77550FC83FE93" box="[721,825,321,347]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-87.78333">87°47'W</geoCoordinate>
,
<quantity id="2D599B77C30FC751E3617550FC1CFE93" box="[839,934,321,347]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="34" unit="m" value="2000.0">2000 m</quantity>
, pine and hardwood forest litter (R.Anderson),
<date id="9E1F1052C30FC751E1917576FDFFFE49" box="[439,581,359,385]" pageId="33" pageNumber="34" value="1994-08-21">21.viii.1994</date>
(all ­
<collectionCode id="8CB0AE57C30FC751E2AA7576FD59FE49" box="[652,739,359,385]" pageId="33" pageNumber="34">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30FC751E11E759AFDBAFCFE" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="EA1E3692C30FC751E11E759AFDEAFD8E" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E11E759AFE0DFE6D" bold="true" box="[312,439,395,421]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C30FC751E1EF759DFD72FE6D" box="[457,712,396,421]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="D8D5EA80C30FC751E1EF759DFD72FE6D" box="[457,712,396,421]" italics="true" pageId="33" pageNumber="34">Seeversiella castanea</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C30FC751E4F5759DFEFDFE05" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C30FC751E4F5759DFEFDFE05" italics="true" pageId="33" pageNumber="34">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 1.5­1.7 times as long as eyes; glossy pronotum with weak isodiametric microsculpture; elytra longer or as long as pronotum; wings fully developed; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C30FC751E4AB7615FA99FDD6" box="[1165,1315,516,542]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Figs. 94­103</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C30FC751E1FA763DFDF9FD8E" box="[476,579,556,582]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Fig. 104</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30FC751E11E7645FDBAFCFE" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">
<taxonomicName id="2DA14D11C30FC751E11E7645FD82FDA5" box="[312,568,596,621]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="D8D5EA80C30FC751E11E7645FD82FDA5" box="[312,568,596,621]" italics="true" pageId="33" pageNumber="34">Seeversiella castanea</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C30FC751E3A17645FB9EFDA5" box="[903,1060,596,621]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C30FC751E3A17645FB9EFDA5" box="[903,1060,596,621]" italics="true" pageId="33" pageNumber="34">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 5, 7­8, 10­12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have strong spicules (
<figureCitation id="729A2A17C30FC751E34B76DDFBB3FD2E" box="[877,1033,716,742]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Figs. 99, 101</figureCitation>
); apex of median lobe in lateral view not widened subapically (
<figureCitation id="729A2A17C30FC751E2C076E5FCCDFCC6" box="[742,887,756,782]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Figs. 96­97</figureCitation>
); spermatheca C­shaped with large umbilicus (
<figureCitation id="729A2A17C30FC751E1AA770DFE48FCFE" box="[396,498,796,822]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Fig. 104</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30FC751E11E7752FDC4FB56" pageId="33" pageNumber="34" type="description">
<paragraph id="EA1E3692C30FC751E11E7752FCD4FC4E" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E11E7752FE6AFC95" bold="true" box="[312,464,835,861]" pageId="33" pageNumber="34">Description.</emphasis>
Length
<quantity id="2D599B77C30FC751E21F7755FD7FFC96" box="[569,709,836,862]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="33" pageNumber="34" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="2D599B77C30FC751E3A57755FB96FC96" box="[899,1068,836,862]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="33" pageNumber="34" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
. Body brown to dark brown with lighter pronotum and elytra, legs brown.
</paragraph>
<paragraph id="EA1E3692C30FC751E11E7785FB89FC1E" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">
In all external character states
<taxonomicName id="2DA14D11C30FC751E2B97785FC9EFC65" box="[671,804,916,941]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="D8D5EA80C30FC751E2B97785FC9EFC65" box="[671,804,916,941]" italics="true" pageId="33" pageNumber="34">S. castanea</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="2DA14D11C30FC751E3997785FBE2FC65" box="[959,1112,916,941]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C30FC751E3997785FBE2FC65" box="[959,1112,916,941]" italics="true" pageId="33" pageNumber="34">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C30FC751E11E77F5FD73FB86" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">In large males posterior angles of male tergum 3 projecting as spines up to 1 time as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to 2/3 as long as tergum length.</paragraph>
<paragraph id="EA1E3692C30FC751E11E704DFD37FBBE" blockId="33.[264,1324,283,1302]" box="[312,653,1116,1142]" pageId="33" pageNumber="34">
Aedeagus as in
<figureCitation id="729A2A17C30FC751E1D7704DFD33FBBE" box="[497,649,1116,1142]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Figs. 94­103</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C30FC751E11E7095FDC4FB56" blockId="33.[264,1324,283,1302]" box="[312,638,1156,1182]" pageId="33" pageNumber="34">
Spermatheca as in
<figureCitation id="729A2A17C30FC751E2337095FDC0FB56" box="[533,634,1156,1182]" captionStart="FIGURES 94 ­ 104" captionStartId="32.[264,383,1439,1463]" captionTargetBox="[296,1295,339,1401]" captionTargetId="figure@32.[282,1305,329,1412]" captionTargetPageId="32" captionText="FIGURES 94 ­ 104. Genitalia of Seeversiella castanea Gusarov, sp. n. (paratypes from Rayon (94 ­ 97, 99 ­ 102) and Las Piedrecitas (98, 103 ­ 104), Mexico). 94 – median lobe, parameral view; 95 – apex of median lobe, parameral view; 96 – median lobe, lateral view; 97 – apex of median lobe, lateral view; 98 – everted internal sac, parameral view; 99 – right hook of internal sac; 100 ­ 101 – details of internal sac retracted into median lobe, lateral view; 102 – apex of left paramere; 103 – copulatory piece, lateral view; 104 – spermatheca. Scale bar 0.2 mm (94, 96, 98, 100 ­ 101), 0.1 mm (95, 97, 99, 102 ­ 104)." httpUri="https://zenodo.org/record/156438/files/figure.png" pageId="33" pageNumber="34">Fig. 104</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C30FC751E11E70BAFE17FADE" pageId="33" pageNumber="34" type="distribution">
<paragraph id="EA1E3692C30FC751E11E70BAFBB4FB0E" blockId="33.[264,1324,283,1302]" box="[312,1038,1195,1222]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E11E70BAFE62FB0D" bold="true" box="[312,472,1195,1221]" pageId="33" pageNumber="34">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C30FC751E2A370BDFD65FB0E" box="[645,735,1196,1222]" name="Mexico" pageId="33" pageNumber="34">Mexico</collectingCountry>
and
<collectingCountry id="92B67602C30FC751E33E70BDFC30FB0E" box="[792,906,1196,1222]" name="Honduras" pageId="33" pageNumber="34">Honduras</collectingCountry>
(
<figureCitation id="729A2A17C30FC751E3BC70BDFBB8FB0E" box="[922,1026,1196,1222]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="33" pageNumber="34">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C30FC751E11E70C2FE17FADE" blockId="33.[264,1324,283,1302]" pageId="33" pageNumber="34">
<emphasis id="D8D5EA80C30FC751E11E70C2FDB3FB25" bold="true" box="[312,521,1235,1261]" pageId="33" pageNumber="34">Natural History.</emphasis>
<taxonomicName id="2DA14D11C30FC751E20670C5FC9BFB25" box="[544,801,1236,1261]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="D8D5EA80C30FC751E20670C5FC9BFB25" box="[544,801,1236,1261]" italics="true" pageId="33" pageNumber="34">Seeversiella castanea</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C30FC751E12E70EDFE12FADE" box="[264,424,1276,1302]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="33" pageNumber="34" unit="m" value="2200.0" valueMax="2400.0" valueMin="2000.0">2000­2400 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

219
data/62/08/87/62088784C333C76FE12E750AFD7BFD86.xml Normal file
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5.
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<emphasis id="D8D5EA80C333C76DE100750DFE47FEFE" bold="true" box="[294,509,284,310]" italics="true" pageId="29" pageNumber="30">Seeversiella fusca</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC333C76DE25B750AFD05FEFD" box="[637,703,283,309]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C333C76DE2E9750AFCE3FEFD" box="[719,857,283,309]" captionStart-0="FIGURES 79 ­ 82" captionStart-1="FIGURES 83 ­ 89" captionStartId-0="29.[264,382,1486,1510]" captionStartId-1="30.[264,383,1432,1456]" captionTargetBox-0="[291,1296,652,1452]" captionTargetBox-1="[285,1297,538,1400]" captionTargetId-0="figure@29.[277,1311,641,1462]" captionTargetId-1="figure@30.[274,1314,526,1408]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="FIGURES 79 ­ 82. Abdominal segment 8 of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 79 – male tergum 8; 80 – male sternum 8; 81 – female tergum 8; 82 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri-0="https://zenodo.org/record/156435/files/figure.png" httpUri-1="https://zenodo.org/record/156436/files/figure.png" pageId="29" pageNumber="30">Figs. 79­89</figureCitation>
)
</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C333C76DE12E7577FCF3FDE9" pageId="29" pageNumber="30" type="materials_examined">
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<emphasis id="D8D5EA80C333C76DE12E7577FE01FE48" bold="true" box="[264,443,358,384]" pageId="29" pageNumber="30">
<typeStatus id="351A8830C333C76DE12E7577FEF9FE48" box="[264,323,358,384]" pageId="29" pageNumber="30">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C333C76DE1ED7576FD81FE49" box="[459,571,359,385]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C333C76DE24E7577FCC0FE48" bold="true" box="[616,890,358,384]" pageId="29" pageNumber="30">
<collectingCountry id="92B67602C333C76DE24E7577FD52FE48" box="[616,744,358,384]" name="Mexico" pageId="29" pageNumber="30">MEXICO</collectingCountry>
: Guerrero:
</emphasis>
<quantity id="2D599B77C333C76DE3AF7576FC69FE49" box="[905,979,359,385]" metricMagnitude="4" metricUnit="m" metricValue="7.1" pageId="29" pageNumber="30" unit="km" value="71.0">71 km</quantity>
NE Atoyac de Alvarez,
<quantity id="2D599B77C333C76DE4D67576FE98FE61" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="29" pageNumber="30" unit="m" value="1700.0">1700 m</quantity>
, flight intercept trap (J.S.Ashe),
<date id="9E1F1052C333C76DE286759EFC9FFE61" box="[672,805,399,425]" pageId="29" pageNumber="30" value="1992-07-27">27.vii.1992</date>
(
<collectionCode id="8CB0AE57C333C76DE313759EFC37FE61" box="[821,909,399,425]" pageId="29" pageNumber="30">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C333C76DE11E75A6FCF3FDE9" blockId="29.[264,1323,358,545]" pageId="29" pageNumber="30">
<typeStatus id="351A8830C333C76DE11E75A6FE15FE19" box="[312,431,439,465]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
:
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<collectingCountry id="92B67602C333C76DE1E375A7FDFFFE18" box="[453,581,438,464]" name="Mexico" pageId="29" pageNumber="30">MEXICO</collectingCountry>
: Guerrero:
</emphasis>
, 2, same data as the
<typeStatus id="351A8830C333C76DE40F75A6FB2EFE19" box="[1065,1172,439,465]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
;, ditto but
<date id="9E1F1052C333C76DE12E75CEFE2BFE31" box="[264,401,479,505]" pageId="29" pageNumber="30" value="1992-07-25">25.vii.1992</date>
, forest litter;
<emphasis id="D8D5EA80C333C76DE21675CFFD23FE30" bold="true" box="[560,665,478,504]" pageId="29" pageNumber="30">Oaxaca:</emphasis>
2,,
<quantity id="2D599B77C333C76DE33475CEFCCEFE31" box="[786,884,479,505]" metricMagnitude="4" metricUnit="m" metricValue="3.84" pageId="29" pageNumber="30" unit="km" value="38.4">38.4 km</quantity>
N Telixtlahuaca, Hwy. 131, 1880 m, forest litter (J.S.Ashe),
<date id="9E1F1052C333C76DE23C7616FD25FDE9" box="[538,671,519,545]" pageId="29" pageNumber="30" value="1992-07-19">19.vii.1992</date>
(all ­
<collectionCode id="8CB0AE57C333C76DE2C07616FC87FDE9" box="[742,829,519,545]" pageId="29" pageNumber="30">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC333C76DE12E71DFFD76F9E3" httpUri="https://zenodo.org/record/156435/files/figure.png" pageId="29" pageNumber="30" targetBox="[291,1296,652,1452]" targetPageId="29">
<paragraph id="EA1E3692C333C76DE12E71DFFD76F9E3" blockId="29.[264,1322,1486,1579]" pageId="29" pageNumber="30">
<emphasis id="D8D5EA80C333C76DE12E71DFFE7FFA2E" bold="true" box="[264,453,1486,1510]" pageId="29" pageNumber="30">FIGURES 79­82</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C333C76DE2D771DEFBADFA2E" authority="Gusarov" authorityName="Gusarov" box="[753,1047,1487,1510]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="fusca" status="sp. nov.">
<emphasis id="D8D5EA80C333C76DE2D771DEFC08FA2E" box="[753,946,1487,1510]" italics="true" pageId="29" pageNumber="30">Seeversiella fusca</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C333C76DE40471DFFBDAFA2E" bold="true" box="[1058,1120,1486,1510]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="C3E657FBC333C76DE40471DFFBDAFA2E" box="[1058,1120,1486,1510]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes, 71 km NE Atoyac de Alvarez, Mexico). 79 – male tergum 8; 80 – male sternum 8; 81 – female tergum 8; 82 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C333C76EE11E7297FDCDF997" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="EA1E3692C333C76DE11E7297FB60F8D1" blockId="29.[264,1323,1670,1897]" pageId="29" pageNumber="30">
<emphasis id="D8D5EA80C333C76DE11E7297FE02F968" bold="true" box="[312,440,1670,1696]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C333C76DE1E17296FD21F968" box="[455,667,1671,1696]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="D8D5EA80C333C76DE1E17296FD21F968" box="[455,667,1671,1696]" italics="true" pageId="29" pageNumber="30">Seeversiella fusca</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C333C76DE4BB7296FA91F968" box="[1181,1323,1671,1696]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C333C76DE4BB7296FA91F968" box="[1181,1323,1671,1696]" italics="true" pageId="29" pageNumber="30">Seeversiella</emphasis>
</taxonomicName>
by having dark brown body; temples as long as eyes; glossy pronotum with weak microsculpture; elytra longer than pronotum; wings fully developed, tergum 8 with four pairs of macrosetae, the distinct shape of aedeagus (
<figureCitation id="729A2A17C333C76DE32E72EEFC2AF8D1" box="[776,912,1791,1817]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="29" pageNumber="30">Figs. 83­88</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C333C76DE45272EEFB76F8D1" box="[1140,1228,1791,1817]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="29" pageNumber="30">Fig. 89</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C333C76EE11E7336FCF9FE66" blockId="29.[264,1323,1670,1897]" lastBlockId="30.[264,1324,284,430]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<taxonomicName id="2DA14D11C333C76DE11E7336FDB7F888" box="[312,525,1831,1856]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="D8D5EA80C333C76DE11E7336FDB7F888" box="[312,525,1831,1856]" italics="true" pageId="29" pageNumber="30">Seeversiella fusca</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="2DA14D11C333C76DE3737336FC4AF888" box="[853,1008,1831,1856]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C333C76DE3737336FC4AF888" box="[853,1008,1831,1856]" italics="true" pageId="29" pageNumber="30">S. globicollis</emphasis>
</taxonomicName>
and the other similar species (3, 6­8, 10­12) with long elytra and weak pronotal microsculpture by the combination of the following characters: distal sclerites of internal sac poorly sclerotized, not dentiform or hook­shaped and have weak spicules (
<figureCitation id="729A2A17C330C76EE3037555FCC6FE96" box="[805,892,324,350]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="30" pageNumber="31">Fig. 87</figureCitation>
); apex of median lobe in lateral view not widened subapically (
<figureCitation id="729A2A17C330C76EE219757DFD76FE4E" box="[575,716,364,390]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="30" pageNumber="31">Figs. 85­86</figureCitation>
); spermatheca L­shaped with small umbilicus and slightly curved in the proximal portion (
<figureCitation id="729A2A17C330C76EE2F87585FC8FFE66" box="[734,821,404,430]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="30" pageNumber="31">Fig. 89</figureCitation>
).
</paragraph>
<caption id="BEDE661AC330C76EE12E7189FD53F9F2" httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="30" pageNumber="31" targetBox="[285,1297,538,1400]" targetPageId="30">
<paragraph id="EA1E3692C330C76EE12E7189FD53F9F2" blockId="30.[264,1323,1432,1631]" pageId="30" pageNumber="31">
<emphasis id="D8D5EA80C330C76EE12E7189FE7EFA78" bold="true" box="[264,452,1432,1456]" pageId="30" pageNumber="31">FIGURES 83­89</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C330C76EE27B7188FCC5FA78" authority="Gusarov" authorityName="Gusarov" box="[605,895,1433,1456]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="fusca" status="sp. nov.">
<emphasis id="D8D5EA80C330C76EE27B7188FCA7FA78" box="[605,797,1433,1456]" italics="true" pageId="30" pageNumber="31">Seeversiella fusca</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C330C76EE3AC7189FC7FFA78" bold="true" box="[906,965,1432,1456]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="C3E657FBC330C76EE3AC7189FC7FFA78" box="[906,965,1432,1456]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88­89).
</paragraph>
</caption>
<paragraph id="EA1E3692C330C76EE12E7259FDCDF997" blockId="30.[264,1323,1432,1631]" box="[264,631,1608,1631]" pageId="30" pageNumber="31">DS – distal sclerite of internal sac.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C330C76FE11E72ACFDD5FE1E" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
<paragraph id="EA1E3692C330C76EE11E72ACFDCBF8C8" blockId="30.[264,1323,1725,1872]" pageId="30" pageNumber="31">
<emphasis id="D8D5EA80C330C76EE11E72ACFE6AF91F" bold="true" box="[312,464,1725,1751]" pageId="30" pageNumber="31">Description.</emphasis>
Length
<quantity id="2D599B77C330C76EE26772AFFD75F910" box="[577,719,1726,1752]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="30" pageNumber="31" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3­2.5 mm</quantity>
, pronotal width
<quantity id="2D599B77C330C76EE3B372AFFBFBF910" box="[917,1089,1726,1752]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="30" pageNumber="31" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.46­0.50 mm</quantity>
. Body dark brown with lighter elytra, legs brown.
</paragraph>
<paragraph id="EA1E3692C330C76EE11E731FFB89F898" blockId="30.[264,1323,1725,1872]" pageId="30" pageNumber="31">
In all external character states
<taxonomicName id="2DA14D11C330C76EE2BF731FFD4EF8EF" box="[665,756,1806,1831]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="D8D5EA80C330C76EE2BF731FFD4EF8EF" box="[665,756,1806,1831]" italics="true" pageId="30" pageNumber="31">S. fusca</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C330C76EE3E3731FFBE7F8EF" box="[965,1117,1806,1831]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C330C76EE3E3731FFBE7F8EF" box="[965,1117,1806,1831]" italics="true" pageId="30" pageNumber="31">S. globicollis</emphasis>
</taxonomicName>
, but differs in the shape of the aedeagus and spermatheca as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C331C76FE11E750DFE03FE4E" blockId="31.[264,1324,284,590]" pageId="31" pageNumber="32">In four examined males posterior angles of tergum 3 not projecting at all or projecting as spines up to 1/3 as long as tergum 3 (measured medially), tergum 7 without medial carina or tubercle.</paragraph>
<paragraph id="EA1E3692C331C76FE11E7585FDC7FE66" blockId="31.[264,1324,284,590]" box="[312,637,404,430]" pageId="31" pageNumber="32">
Aedeagus as in
<figureCitation id="729A2A17C331C76FE1D67585FDC3FE66" box="[496,633,404,430]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="31" pageNumber="32">Figs. 83­88</figureCitation>
.
</paragraph>
<paragraph id="EA1E3692C331C76FE11E75ADFDD5FE1E" blockId="31.[264,1324,284,590]" box="[312,623,444,470]" pageId="31" pageNumber="32">
Spermatheca as in
<figureCitation id="729A2A17C331C76FE23275ADFDD1FE1E" box="[532,619,444,470]" captionStart="FIGURES 83 ­ 89" captionStartId="30.[264,383,1432,1456]" captionTargetBox="[285,1297,538,1400]" captionTargetId="figure@30.[274,1314,526,1408]" captionTargetPageId="30" captionText="FIGURES 83 ­ 89. Genitalia of Seeversiella fusca Gusarov, sp. n. (paratypes, 71 km NE Atoyac de Alvarez, Mexico). 83 – median lobe, parameral view; 84 – apex of median lobe, parameral view; 85 – median lobe, lateral view; 86 – apex of median lobe, lateral view; 87 – details of internal sac retracted into median lobe, lateral view; 88 – apex of left paramere; 89 – spermatheca. Scale bar 0.2 mm (83, 85, 87), 0.1 mm (84, 86, 88 ­ 89)." httpUri="https://zenodo.org/record/156436/files/figure.png" pageId="31" pageNumber="32">Fig. 89</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C331C76FE11E75F2FD7BFD86" pageId="31" pageNumber="32" type="distribution">
<paragraph id="EA1E3692C331C76FE11E75F2FCD8FE36" blockId="31.[264,1324,284,590]" box="[312,866,483,510]" pageId="31" pageNumber="32">
<emphasis id="D8D5EA80C331C76FE11E75F2FE62FE35" bold="true" box="[312,472,483,509]" pageId="31" pageNumber="32">Distribution.</emphasis>
Known from
<collectingCountry id="92B67602C331C76FE2A375F5FD65FE36" box="[645,735,484,510]" name="Mexico" pageId="31" pageNumber="32">Mexico</collectingCountry>
(
<figureCitation id="729A2A17C331C76FE2C875F5FCEFFE36" box="[750,853,484,510]" captionStart="FIGURE 387" captionStartId="97.[264,368,1861,1885]" captionTargetBox="[313,1275,771,1826]" captionTargetId="figure@97.[304,1282,766,1834]" captionTargetPageId="97" captionText="FIGURE 387. Geographical distribution of Seeversiella from Mexico to Honduras." httpUri="https://zenodo.org/record/156489/files/figure.png" pageId="31" pageNumber="32">Fig. 387</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C331C76FE11E761AFD7BFD86" blockId="31.[264,1324,284,590]" pageId="31" pageNumber="32">
<emphasis id="D8D5EA80C331C76FE11E761AFDBEFDED" bold="true" box="[312,516,523,549]" pageId="31" pageNumber="32">Natural History.</emphasis>
<taxonomicName id="2DA14D11C331C76FE235761DFD5CFDED" box="[531,742,524,549]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="fusca">
<emphasis id="D8D5EA80C331C76FE235761DFD5CFDED" box="[531,742,524,549]" italics="true" pageId="31" pageNumber="32">Seeversiella fusca</emphasis>
</taxonomicName>
was collected in forest litter and with flight intercept traps at altitude of
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.
</paragraph>
</subSubSection>
</treatment>
</document>

220
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<mods:identifier id="CA3215A00096105A0E882D50C296BA1A" type="Zenodo-Dep">156420</mods:identifier>
</mods:mods>
<treatment id="62088784C335C76CE12E750AFB79F90D" ID-DOI="http://doi.org/10.5281/zenodo.6274642" ID-Zenodo-Dep="6274642" LSID="urn:lsid:plazi:treatment:62088784C335C76CE12E750AFB79F90D" httpUri="http://treatment.plazi.org/id/62088784C335C76CE12E750AFB79F90D" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<subSubSection id="A2BB6519C335C76BE12E750AFC2FFEFD" box="[264,917,283,310]" pageId="27" pageNumber="28" type="key">
<paragraph id="EA1E3692C335C76BE12E750AFC2FFEFD" blockId="27.[264,917,283,310]" box="[264,917,283,310]" pageId="27" pageNumber="28">
<heading id="B15681FEC335C76BE12E750AFC2FFEFD" bold="true" box="[264,917,283,310]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="D8D5EA80C335C76BE12E750AFC2FFEFD" bold="true" box="[264,917,283,310]" pageId="27" pageNumber="28">
<keyLead id="515095A7C335C76BE12E750AFC2FFEFD" box="[264,917,283,310]" pageId="27" pageNumber="28">
<key id="9E3061A3C335C76BE12E750AFC2FFEFD" box="[264,917,283,310]" pageId="27" pageNumber="28">
<keyStep id="51552E37C335C76BE12E750AFC2FFEFD" box="[264,917,283,310]" pageId="27" pageNumber="28">
4.
<taxonomicName id="2DA14D11C335C76BE100750DFD1EFEFD" ID-CoL="6YBCT" authority="Gusarov" authorityName="Gusarov" box="[294,676,283,310]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="liliputana" status="sp. nov.">
<emphasis id="D8D5EA80C335C76BE100750DFD8BFEFE" bold="true" box="[294,561,284,310]" italics="true" pageId="27" pageNumber="28">Seeversiella liliputana</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC335C76BE297750AFD49FEFD" box="[689,755,283,309]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C335C76BE325750AFC37FEFD" box="[771,909,283,309]" captionStart-0="FIGURES 68 ­ 71" captionStart-1="FIGURES 72 ­ 78" captionStartId-0="27.[264,383,1474,1498]" captionStartId-1="28.[264,383,1298,1322]" captionTargetBox-0="[282,1307,629,1441]" captionTargetBox-1="[300,1285,615,1266]" captionTargetId-0="figure@27.[268,1319,604,1448]" captionTargetId-1="figure@28.[296,1291,609,1272]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionText-0="FIGURES 68 ­ 71. Abdominal segment 8 of Seeversiella liliputana Gusarov, sp. n. (68 ­ 69, holotype; 70 ­ 71, possible female of S. liliputana from Bear Saddle, Arizona). 68 – male tergum 8; 69 – male sternum 8; 70 – female tergum 8; 71 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 – median lobe, parameral view; 73 – apex of median lobe, parameral view; 74 – median lobe, lateral view; 75 – apex of median lobe, lateral view; 76 – details of internal sac retracted into median lobe, lateral view; 77 – apex of left paramere; 78 – spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78)." httpUri-0="https://zenodo.org/record/156433/files/figure.png" httpUri-1="https://zenodo.org/record/156434/files/figure.png" pageId="27" pageNumber="28">Figs. 68­78</figureCitation>
)
</keyStep>
</key>
</keyLead>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C335C76BE12E7577FDB8FE31" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="EA1E3692C335C76BE12E7577FC01FE61" blockId="27.[264,1323,358,505]" pageId="27" pageNumber="28">
<emphasis id="D8D5EA80C335C76BE12E7577FE05FE48" bold="true" box="[264,447,358,384]" pageId="27" pageNumber="28">
<typeStatus id="351A8830C335C76BE12E7577FEF9FE48" box="[264,323,358,384]" pageId="27" pageNumber="28">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C335C76BE1F07576FDFCFE49" box="[470,582,359,385]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C335C76BE25D7577FC4AFE48" bold="true" box="[635,1008,358,384]" pageId="27" pageNumber="28">
<collectingCountry id="92B67602C335C76BE25D7577FCD2FE48" box="[635,872,358,384]" name="United States of America" pageId="27" pageNumber="28">UNITED STATES</collectingCountry>
: Arizona:
</emphasis>
Cochise Co.: Huachuca Mts., Bear Saddle, 8100' (A.Smetana),
<date id="9E1F1052C335C76BE2F5759EFCE9FE61" box="[723,851,399,425]" pageId="27" pageNumber="28" value="1979-08-05">5.viii.1979</date>
(
<collectionCode id="8CB0AE57C335C76BE344759EFC14FE61" box="[866,942,399,425]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="28">CNCI</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C335C76BE11E75A7FDB8FE31" blockId="27.[264,1323,358,505]" pageId="27" pageNumber="28">
<emphasis id="D8D5EA80C335C76BE11E75A7FC0DFE18" bold="true" box="[312,951,438,464]" pageId="27" pageNumber="28">
Additional material.
<collectingCountry id="92B67602C335C76BE26175A7FC89FE18" box="[583,819,438,464]" name="United States of America" pageId="27" pageNumber="28">UNITED STATES</collectingCountry>
: Arizona:
</emphasis>
Cochise Co.:, same data as the
<typeStatus id="351A8830C335C76BE11575CEFE20FE31" box="[307,410,479,505]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C335C76BE18C75CEFE4DFE31" box="[426,503,479,505]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="28">CNCI</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC335C76BE12E71D3FBCAF9D7" httpUri="https://zenodo.org/record/156433/files/figure.png" pageId="27" pageNumber="28" targetBox="[282,1307,629,1441]" targetPageId="27">
<paragraph id="EA1E3692C335C76BE12E71D3FBCAF9D7" blockId="27.[264,1322,1474,1567]" pageId="27" pageNumber="28">
<emphasis id="D8D5EA80C335C76BE12E71D3FE7FFA12" bold="true" box="[264,453,1474,1498]" pageId="27" pageNumber="28">FIGURES 68­71</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C335C76BE2D271D2FBF3FA12" authority="Gusarov" authorityName="Gusarov" box="[756,1097,1475,1498]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="liliputana" status="sp. nov.">
<emphasis id="D8D5EA80C335C76BE2D271D2FC5FFA12" box="[756,997,1475,1498]" italics="true" pageId="27" pageNumber="28">Seeversiella liliputana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C335C76BE47071D3FB29FA12" bold="true" box="[1110,1171,1474,1498]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="C3E657FBC335C76BE47071D3FB29FA12" box="[1110,1171,1474,1498]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(68­69, holotype; 70­71, possible female of
<taxonomicName id="2DA14D11C335C76BE27C71F7FD66FA35" box="[602,732,1510,1533]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C335C76BE27C71F7FD66FA35" box="[602,732,1510,1533]" italics="true" pageId="27" pageNumber="28">S. liliputana</emphasis>
</taxonomicName>
from Bear Saddle, Arizona). 68 – male tergum 8; 69 – male sternum 8; 70 – female tergum 8; 71 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C335C76BE11E726BFCF8F895" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="EA1E3692C335C76BE11E726BFD3FF8C5" blockId="27.[264,1323,1658,1885]" pageId="27" pageNumber="28">
<emphasis id="D8D5EA80C335C76BE11E726BFE0DF95C" bold="true" box="[312,439,1658,1684]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C335C76BE1EE726AFD75F95C" box="[456,719,1659,1684]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C335C76BE1EE726AFD75F95C" box="[456,719,1659,1684]" italics="true" pageId="27" pageNumber="28">Seeversiella liliputana</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C335C76BE4F5726AFEFDF974" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C335C76BE4F5726AFEFDF974" italics="true" pageId="27" pageNumber="28">Seeversiella</emphasis>
</taxonomicName>
by having brown body; temples 2 times as long as eyes; pronotum without microsculpture; elytra as long as pronotum; tergum 8 with four pairs of macrosetae, and the distinct shape of aedeagus (
<figureCitation id="729A2A17C335C76BE1CB72E2FDC3F8C5" box="[493,633,1779,1805]" captionStart="FIGURES 72 ­ 78" captionStartId="28.[264,383,1298,1322]" captionTargetBox="[300,1285,615,1266]" captionTargetId="figure@28.[296,1291,609,1272]" captionTargetPageId="28" captionText="FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 – median lobe, parameral view; 73 – apex of median lobe, parameral view; 74 – median lobe, lateral view; 75 – apex of median lobe, lateral view; 76 – details of internal sac retracted into median lobe, lateral view; 77 – apex of left paramere; 78 – spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78)." httpUri="https://zenodo.org/record/156434/files/figure.png" pageId="27" pageNumber="28">Figs. 72­77</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C335C76BE11E730AFCF8F895" blockId="27.[264,1323,1658,1885]" pageId="27" pageNumber="28">
<taxonomicName id="2DA14D11C335C76BE11E730AFDFAF8FC" box="[312,576,1819,1844]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C335C76BE11E730AFDFAF8FC" box="[312,576,1819,1844]" italics="true" pageId="27" pageNumber="28">Seeversiella liliputana</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C335C76BE2FB730AFC26F8FC" box="[733,924,1819,1844]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C335C76BE2FB730AFC26F8FC" box="[733,924,1819,1844]" italics="true" pageId="27" pageNumber="28">S. sonomotoides</emphasis>
</taxonomicName>
in having smaller median lobe of aedeagus with shorter apex (
<figureCitation id="729A2A17C335C76BE2707352FD5AF895" box="[598,736,1859,1885]" captionStart="FIGURES 72 ­ 78" captionStartId="28.[264,383,1298,1322]" captionTargetBox="[300,1285,615,1266]" captionTargetId="figure@28.[296,1291,609,1272]" captionTargetPageId="28" captionText="FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 – median lobe, parameral view; 73 – apex of median lobe, parameral view; 74 – median lobe, lateral view; 75 – apex of median lobe, lateral view; 76 – details of internal sac retracted into median lobe, lateral view; 77 – apex of left paramere; 78 – spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78)." httpUri="https://zenodo.org/record/156434/files/figure.png" pageId="27" pageNumber="28">Figs. 72­75</figureCitation>
; 58­61).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C332C76CE11E750AFBB8F9BD" pageId="28" pageNumber="29" type="description">
<paragraph id="EA1E3692C332C76CE11E750AFC36FE96" blockId="28.[264,1323,283,550]" pageId="28" pageNumber="29">
<emphasis id="D8D5EA80C332C76CE11E750AFE6AFEFD" bold="true" box="[312,464,283,309]" pageId="28" pageNumber="29">Description.</emphasis>
Length
<quantity id="2D599B77C332C76CE21E750DFD2EFEFE" box="[568,660,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="28" pageNumber="29" unit="mm" value="2.4">2.4 mm</quantity>
, pronotal width
<quantity id="2D599B77C332C76CE377750DFC0CFEFE" box="[849,950,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="28" pageNumber="29" unit="mm" value="0.41">0.41mm</quantity>
. Body brown, elytra and antennae light brown, legs and mouthparts brownish yellow.
</paragraph>
<paragraph id="EA1E3692C332C76CE11E757DFC30FE66" blockId="28.[264,1323,283,550]" pageId="28" pageNumber="29">
In all character states
<taxonomicName id="2DA14D11C332C76CE211757DFD7CFE4D" box="[567,710,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C332C76CE211757DFD7CFE4D" box="[567,710,364,389]" italics="true" pageId="28" pageNumber="29">S. liliputana</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="2DA14D11C332C76CE3BD757DFBE0FE4D" box="[923,1114,364,389]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C332C76CE3BD757DFBE0FE4D" box="[923,1114,364,389]" italics="true" pageId="28" pageNumber="29">S. sonomotoides</emphasis>
</taxonomicName>
, but differs in the shape of the median lobe as described in the diagnosis.
</paragraph>
<paragraph id="EA1E3692C332C76CE11E75ADFD34FE36" blockId="28.[264,1323,283,550]" pageId="28" pageNumber="29">In the only known male specimen the posterior angles of tergum 3 not projecting, tergum 7 without carina or tubercle.</paragraph>
<paragraph id="EA1E3692C332C76CE11E761DFDC7FDEE" blockId="28.[264,1323,283,550]" box="[312,637,524,550]" pageId="28" pageNumber="29">
Aedeagus as in
<figureCitation id="729A2A17C332C76CE1D6761DFDC3FDEE" box="[496,633,524,550]" captionStart="FIGURES 72 ­ 78" captionStartId="28.[264,383,1298,1322]" captionTargetBox="[300,1285,615,1266]" captionTargetId="figure@28.[296,1291,609,1272]" captionTargetPageId="28" captionText="FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 – median lobe, parameral view; 73 – apex of median lobe, parameral view; 74 – median lobe, lateral view; 75 – apex of median lobe, lateral view; 76 – details of internal sac retracted into median lobe, lateral view; 77 – apex of left paramere; 78 – spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78)." httpUri="https://zenodo.org/record/156434/files/figure.png" pageId="28" pageNumber="29">Figs. 72­77</figureCitation>
.
</paragraph>
<caption id="BEDE661AC332C76CE12E7103FB28FA7C" httpUri="https://zenodo.org/record/156434/files/figure.png" pageId="28" pageNumber="29" targetBox="[300,1285,615,1266]" targetPageId="28">
<paragraph id="EA1E3692C332C76CE12E7103FB28FA7C" blockId="28.[264,1323,1298,1460]" pageId="28" pageNumber="29">
<emphasis id="D8D5EA80C332C76CE12E7103FE7FFAE2" bold="true" box="[264,453,1298,1322]" pageId="28" pageNumber="29">FIGURES 72­78</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C332C76CE2417102FC01FAE2" authority="Gusarov" authorityName="Gusarov" box="[615,955,1299,1322]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liliputana" status="sp. nov.">
<emphasis id="D8D5EA80C332C76CE2417102FCE2FAE2" box="[615,856,1299,1322]" italics="true" pageId="28" pageNumber="29">Seeversiella liliputana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C332C76CE3EE7103FBBCFAE2" bold="true" box="[968,1030,1298,1322]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="C3E657FBC332C76CE3EE7103FBBCFAE2" box="[968,1030,1298,1322]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(72­77, holotype; 78, possible female of
<taxonomicName id="2DA14D11C332C76CE1967127FD89FA85" box="[432,563,1334,1357]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C332C76CE1967127FD89FA85" box="[432,563,1334,1357]" italics="true" pageId="28" pageNumber="29">S. liliputana</emphasis>
</taxonomicName>
from Bear Saddle, Arizona). 72 – median lobe, parameral view; 73 – apex of median lobe, parameral view; 74 – median lobe, lateral view; 75 – apex of median lobe, lateral view; 76 – details of internal sac retracted into median lobe, lateral view; 77 – apex of left paramere; 78 – spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77­78).
</paragraph>
</caption>
<paragraph id="EA1E3692C332C76CE11E721AFBB8F9BD" blockId="28.[264,1323,1547,1733]" pageId="28" pageNumber="29">
One female specimen, externally similar to the
<typeStatus id="351A8830C332C76CE347721AFC72F9ED" box="[865,968,1547,1573]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
of
<taxonomicName id="2DA14D11C332C76CE3C9721AFBC4F9EC" box="[1007,1150,1547,1572]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C332C76CE3C9721AFBC4F9EC" box="[1007,1150,1547,1572]" italics="true" pageId="28" pageNumber="29">S. liliputana</emphasis>
</taxonomicName>
, was collected together with this
<typeStatus id="351A8830C332C76CE1FB7222FDFDF985" box="[477,583,1587,1613]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
. However that female specimen has the spermatheca like in
<taxonomicName id="2DA14D11C332C76CE5337222FE13F9BC" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C332C76CE5337222FE13F9BC" italics="true" pageId="28" pageNumber="29">S. sonomotoides</emphasis>
</taxonomicName>
(
<figureCitation id="729A2A17C332C76CE19F724AFDA6F9BD" box="[441,540,1627,1653]" captionStart="FIGURES 72 ­ 78" captionStartId="28.[264,383,1298,1322]" captionTargetBox="[300,1285,615,1266]" captionTargetId="figure@28.[296,1291,609,1272]" captionTargetPageId="28" captionText="FIGURES 72 ­ 78. Genitalia of Seeversiella liliputana Gusarov, sp. n. (72 ­ 77, holotype; 78, possible female of S. liliputana from Bear Saddle, Arizona). 72 – median lobe, parameral view; 73 – apex of median lobe, parameral view; 74 – median lobe, lateral view; 75 – apex of median lobe, lateral view; 76 – details of internal sac retracted into median lobe, lateral view; 77 – apex of left paramere; 78 – spermatheca. Scale bar 0.2 mm (72, 74, 76), 0.1 mm (73, 75, 77 ­ 78)." httpUri="https://zenodo.org/record/156434/files/figure.png" pageId="28" pageNumber="29">Figs. 78</figureCitation>
,
<figureCitation id="729A2A17C332C76CE20E724AFDFDF9BD" box="[552,583,1627,1653]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="28" pageNumber="29">62</figureCitation>
) and may belong to the latter species.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C332C76CE11E7293FB79F90D" pageId="28" pageNumber="29" type="distribution">
<paragraph id="EA1E3692C332C76CE11E7293FB21F955" blockId="28.[264,1323,1547,1733]" box="[312,1179,1666,1693]" pageId="28" pageNumber="29">
<emphasis id="D8D5EA80C332C76CE11E7293FE62F954" bold="true" box="[312,472,1666,1692]" pageId="28" pageNumber="29">Distribution.</emphasis>
Known from the Huachuca Mountains, Arizona (
<figureCitation id="729A2A17C332C76CE4017292FB35F955" box="[1063,1167,1667,1693]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="28" pageNumber="29">Fig. 386</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C332C76CE11E72BBFB79F90D" blockId="28.[264,1323,1547,1733]" box="[312,1219,1706,1733]" pageId="28" pageNumber="29">
<emphasis id="D8D5EA80C332C76CE11E72BBFDBFF90C" bold="true" box="[312,517,1706,1732]" pageId="28" pageNumber="29">Natural History.</emphasis>
<taxonomicName id="2DA14D11C332C76CE23572BAFCA1F90C" box="[531,795,1707,1732]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C332C76CE23572BAFCA1F90C" box="[531,795,1707,1732]" italics="true" pageId="28" pageNumber="29">Seeversiella liliputana</emphasis>
</taxonomicName>
was collected at altitude of
<quantity id="2D599B77C332C76CE44572BAFB05F90D" box="[1123,1215,1707,1733]" metricMagnitude="3" metricUnit="m" metricValue="2.4699999999999998" pageId="28" pageNumber="29" unit="m" value="2470.0">2470 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

242
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@ -0,0 +1,242 @@
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<mods:title id="393DCAC902FA4C05EBCF86FA451CDF50">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<paragraph id="EA1E3692C336C768E12E77F7FC7EFBC8" blockId="24.[264,964,998,1024]" box="[264,964,998,1024]" pageId="24" pageNumber="25">
<heading id="B15681FEC336C768E12E77F7FC7EFBC8" bold="true" box="[264,964,998,1024]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<key id="9E3061A3C336C768E12E77F7FC7EFBC8" box="[264,964,998,1024]" pageId="24" pageNumber="25">
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<emphasis id="D8D5EA80C336C768E12E77F7FC7EFBC8" bold="true" box="[264,964,998,1024]" pageId="24" pageNumber="25">
<keyLead id="515095A7C336C768E12E77F7FC7EFBC8" box="[264,964,998,1024]" pageId="24" pageNumber="25">
3.
<taxonomicName id="2DA14D11C336C768E10077F7FD69FBC8" ID-CoL="6YBBF" authority="Gusarov" authorityName="Gusarov" box="[294,723,998,1024]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sonomotoides" status="sp. nov.">
<emphasis id="D8D5EA80C336C768E10077F7FDDAFBC8" bold="true" box="[294,608,998,1024]" italics="true" pageId="24" pageNumber="25">Seeversiella sonomotoides</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC336C768E2C677F7FC98FBC8" box="[736,802,998,1024]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C336C768E31477F7FC06FBC8" box="[818,956,998,1024]" captionStart-0="FIGURES 54 ­ 57" captionStart-1="FIGURES 58 ­ 67" captionStartId-0="25.[264,383,1800,1824]" captionStartId-1="26.[264,382,1663,1687]" captionTargetBox-0="[322,1265,1015,1769]" captionTargetBox-1="[289,1302,621,1628]" captionTargetId-0="figure@25.[313,1274,1006,1776]" captionTargetId-1="figure@26.[279,1307,609,1637]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="FIGURES 54 ­ 57. Abdominal segment 8 of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 54 – male tergum 8; 55 – male sternum 8; 56 – female tergum 8; 57 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri-0="https://zenodo.org/record/156431/files/figure.png" httpUri-1="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 54­67</figureCitation>
)
</keyLead>
</emphasis>
</keyStep>
</key>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C336C768E12E7020FBFBFB0B" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="EA1E3692C336C768E12E7020FB16FBBB" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E12E7020FE05FB83" bold="true" box="[264,447,1073,1099]" pageId="24" pageNumber="25">
<typeStatus id="351A8830C336C768E12E7020FEF9FB83" box="[264,323,1073,1099]" pageId="24" pageNumber="25">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C336C768E1F07020FDFCFB83" box="[470,582,1073,1099]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C336C768E25D7020FC4AFB83" bold="true" box="[635,1008,1073,1099]" pageId="24" pageNumber="25">
<collectingCountry id="92B67602C336C768E25D7020FCD2FB83" box="[635,872,1073,1099]" name="United States of America" pageId="24" pageNumber="25">UNITED STATES</collectingCountry>
: Arizona:
</emphasis>
Cochise Co.: Huachuca Mts., Carr Cny.,
<quantity id="2D599B77C336C768E1EA7048FD92FBBB" box="[460,552,1113,1139]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="24" pageNumber="25" unit="m" value="2150.0">2150 m</quantity>
, leaf oak litter (P.M.Hammond),
<date id="9E1F1052C336C768E38D7048FB8BFBBB" box="[939,1073,1113,1139]" pageId="24" pageNumber="25" value="1976-07-23">23.vii.1976</date>
(
<collectionCode id="8CB0AE57C336C768E4677048FB1AFBBB" box="[1089,1184,1113,1139]" pageId="24" pageNumber="25">BMNH</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E7090FBFBFB0B" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<typeStatus id="351A8830C336C768E11E7090FE15FB53" box="[312,431,1153,1179]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C336C768E1E77090FC8AFB53" bold="true" box="[449,816,1153,1179]" pageId="24" pageNumber="25">
<collectingCountry id="92B67602C336C768E1E77090FD16FB53" box="[449,684,1153,1179]" name="United States of America" pageId="24" pageNumber="25">UNITED STATES</collectingCountry>
: Arizona:
</emphasis>
Cochise Co.:
<specimenCount id="FCA7FD1BC336C768E3C17090FB33FB53" box="[999,1161,1153,1179]" pageId="24" pageNumber="25" type="generic">44 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C336C768E11570B8FE20FB0B" box="[307,410,1193,1219]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C336C768E18F70B8FDBDFB0B" box="[425,519,1193,1219]" pageId="24" pageNumber="25">BMNH</collectionCode>
); 2, ditto but (J.M.Campbell) (
<collectionCode id="8CB0AE57C336C768E3B570B8FC64FB0B" box="[915,990,1193,1219]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="24" pageNumber="25">CNCI</collectionCode>
,
<collectionCode id="8CB0AE57C336C768E3CF70B8FB8EFB0B" box="[1001,1076,1193,1219]" pageId="24" pageNumber="25">SPSU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C336C768E11E70C0FC6DF9CB" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="EA1E3692C336C768E11E70C0FE65FA7B" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E11E70C0FE0DFB23" bold="true" box="[312,439,1233,1259]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C336C768E1E070C3FD47FB23" box="[454,765,1234,1259]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C336C768E1E070C3FD47FB23" box="[454,765,1234,1259]" italics="true" pageId="24" pageNumber="25">Seeversiella sonomotoides</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C336C768E4DE70C3FED7FADB" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C336C768E4DE70C3FED7FADB" italics="true" pageId="24" pageNumber="25">Seeversiella</emphasis>
</taxonomicName>
by having brown body; large eyes (temples 1.7­2.1 times as long as eyes); glossy pronotum without microsculpture or with weak isodiametric microsculpture; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C336C768E3D37158FB6DFAAB" box="[1013,1239,1353,1379]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 58­61, 63­67</figureCitation>
), especially the dentiform distal sclerites of internal sac (
<figureCitation id="729A2A17C336C768E3547160FBBDFA43" box="[882,1031,1393,1419]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 63, 65</figureCitation>
); and the shape of spermatheca (
<figureCitation id="729A2A17C336C768E15C7188FE6BFA7B" box="[378,465,1433,1459]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E71D3FC6DF9CB" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<taxonomicName id="2DA14D11C336C768E11E71D3FDC9FA13" box="[312,627,1474,1499]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C336C768E11E71D3FDC9FA13" box="[312,627,1474,1499]" italics="true" pageId="24" pageNumber="25">Seeversiella sonomotoides</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C336C768E3F771D3FBDAFA13" box="[977,1120,1474,1499]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="D8D5EA80C336C768E3F771D3FBDAFA13" box="[977,1120,1474,1499]" italics="true" pageId="24" pageNumber="25">S. nigriceps</emphasis>
</taxonomicName>
in having wider distal sclerites of internal sac of aedeagus (
<figureCitation id="729A2A17C336C768E2D971F8FC37F9CB" box="[767,909,1513,1539]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 63, 65</figureCitation>
; 124).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C336C76AE11E7203FDD5FE66" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="description">
<paragraph id="EA1E3692C336C768E11E7203FC86F99B" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<taxonomicName id="2DA14D11C336C768E11E7203FDCAF9E3" box="[312,624,1554,1579]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C336C768E11E7203FDCAF9E3" box="[312,624,1554,1579]" italics="true" pageId="24" pageNumber="25">Seeversiella sonomotoides</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C336C768E3347203FC1EF9E3" box="[786,932,1554,1579]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="liliputana">
<emphasis id="D8D5EA80C336C768E3347203FC1EF9E3" box="[786,932,1554,1579]" italics="true" pageId="24" pageNumber="25">S. liliputana</emphasis>
</taxonomicName>
in having larger median lobe of aedeagus with longer apex (
<figureCitation id="729A2A17C336C768E2767228FD63F99B" box="[592,729,1593,1619]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 58­61</figureCitation>
; 72­75).
</paragraph>
<paragraph id="EA1E3692C336C768E11E7273FB5EF903" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<taxonomicName id="2DA14D11C336C768E11E7273FDCBF9B3" box="[312,625,1634,1659]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C336C768E11E7273FDCBF9B3" box="[312,625,1634,1659]" italics="true" pageId="24" pageNumber="25">Seeversiella sonomotoides</emphasis>
</taxonomicName>
differs from the similar species of
<taxonomicName id="2DA14D11C336C768E4067273FB14F9B3" box="[1056,1198,1634,1659]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C336C768E4067273FB14F9B3" box="[1056,1198,1634,1659]" italics="true" pageId="24" pageNumber="25">Seeversiella</emphasis>
</taxonomicName>
with long elytra and weak pronotal microsculpture in having strongly dentiform distal sclerites of internal sac (
<figureCitation id="729A2A17C336C768E1B872A0FD90F903" box="[414,554,1713,1739]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Figs. 63, 65</figureCitation>
) and L­shaped spermatheca with small umbilicus (
<figureCitation id="729A2A17C336C768E45872A0FB6CF903" box="[1150,1238,1713,1739]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="24" pageNumber="25">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E72C8FDB1F8D3" blockId="24.[264,1324,1073,1899]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E11E72C8FE6AF93B" bold="true" box="[312,464,1753,1779]" pageId="24" pageNumber="25">Description.</emphasis>
Length
<quantity id="2D599B77C336C768E21D72C8FD72F93B" box="[571,712,1753,1779]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="24" pageNumber="25" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
. Body brown, in some elytra and pronotum lighter, legs yellowish brown.
</paragraph>
<paragraph id="EA1E3692C336C769E11E7338FB84FE96" blockId="24.[264,1324,1073,1899]" lastBlockId="25.[264,1324,284,910]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">Head surface glossy, without microsculpture or with weak isodiametric microsculpture, with fine, weak and poorly visible punctation, distance between punctures equals 2­3 times their diameter. Temples 1.7­2.1 times as long as eyes. Antennal article 2 longer than article 3, articles 4­10 transverse or strongly transverse (ratio 1.5­2.0).</paragraph>
<paragraph id="EA1E3692C337C769E11E757DFDAEFDBE" blockId="25.[264,1324,284,910]" pageId="25" pageNumber="26">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C337C769E40D757DFB6CFE4E" box="[1067,1238,364,390]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="25" pageNumber="26" unit="mm" value="0.42" valueMax="0.44" valueMin="0.4">0.40­0.44 mm</quantity>
, length
<quantity id="2D599B77C337C769E12E7585FE09FE66" box="[264,435,404,430]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.5999999999999996" pageId="25" pageNumber="26" unit="mm" value="0.395" valueMax="0.43" valueMin="0.36">0.36­0.43 mm</quantity>
, width to length ratio 1.1, surface glossy, without microsculpture; punctation as on head. Elytra wider than pronotum and as long (measured from humeral angle) as pronotum (pronotal length to elytral length ratio 1.0), 1.3 times wider than long, glossy, with weak isodiametric microsculpture, with fine and slightly asperate punctation, distance between punctures equal to 1­2 times their diameter. Wings fully developed, 3­4 times as long as elytra.
</paragraph>
<paragraph id="EA1E3692C337C769E11E7695FCA7FCDE" blockId="25.[264,1324,284,910]" pageId="25" pageNumber="26">Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 2­3 times their diameter on terga 3­5 and 3­6 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with four pairs of macrosetae (54, 56).</paragraph>
<paragraph id="EA1E3692C337C769E11E7735FECCFC46" blockId="25.[264,1324,284,910]" pageId="25" pageNumber="26">
In males posterior angles of tergum 3 not projecting, tergum 7 without medial carina. Posterior margin of male tergum 8 slightly concave medially, but without emargination (
<figureCitation id="729A2A17C337C769E1377765FED3FC46" box="[273,361,884,910]" captionStart="FIGURES 54 ­ 57" captionStartId="25.[264,383,1800,1824]" captionTargetBox="[322,1265,1015,1769]" captionTargetId="figure@25.[313,1274,1006,1776]" captionTargetPageId="25" captionText="FIGURES 54 ­ 57. Abdominal segment 8 of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 54 – male tergum 8; 55 – male sternum 8; 56 – female tergum 8; 57 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156431/files/figure.png" pageId="25" pageNumber="26">Fig. 54</figureCitation>
).
</paragraph>
<caption id="BEDE661AC337C769E12E7319FD2DF8AD" httpUri="https://zenodo.org/record/156431/files/figure.png" pageId="25" pageNumber="26" targetBox="[322,1265,1015,1769]" targetPageId="25">
<paragraph id="EA1E3692C337C769E12E7319FD2DF8AD" blockId="25.[264,1323,1800,1893]" pageId="25" pageNumber="26">
<emphasis id="D8D5EA80C337C769E12E7319FE7EF8E8" bold="true" box="[264,452,1800,1824]" pageId="25" pageNumber="26">FIGURES 54­57</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C337C769E2CB7318FBD1F8D7" authority="Gusarov" authorityName="Gusarov" box="[749,1131,1800,1824]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sonomotoides" status="sp. nov.">
<emphasis id="D8D5EA80C337C769E2CB7318FBB3F8E8" box="[749,1033,1801,1824]" italics="true" pageId="25" pageNumber="26">Seeversiella sonomotoides</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C337C769E4517319FB0EF8E8" bold="true" box="[1143,1204,1800,1824]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="C3E657FBC337C769E4517319FB0EF8E8" box="[1143,1204,1800,1824]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Carr Canyon, Arizona). 54 – male tergum 8; 55 – male sternum 8; 56 – female tergum 8; 57 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C334C76AE11E750DFE8CFE4E" blockId="26.[264,1324,284,550]" pageId="26" pageNumber="27">
Aedeagus as in
<figureCitation id="729A2A17C334C76AE1D7750DFD77FEFE" box="[497,717,284,310]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="26" pageNumber="27">Figs. 58­61, 63­67</figureCitation>
. Distal sclerites of internal sac dentiform (
<figureCitation id="729A2A17C334C76AE4E2750DFE93FE96" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="26" pageNumber="27">Figs. 63, 65</figureCitation>
). Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C334C76AE4DA7555FE93FE4E" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="26" pageNumber="27">Fig. 64</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C334C76AE11E7585FDD5FE66" blockId="26.[264,1324,284,550]" box="[312,623,404,430]" pageId="26" pageNumber="27">
Spermatheca as in
<figureCitation id="729A2A17C334C76AE2327585FDD1FE66" box="[532,619,404,430]" captionStart="FIGURES 58 ­ 67" captionStartId="26.[264,382,1663,1687]" captionTargetBox="[289,1302,621,1628]" captionTargetId="figure@26.[279,1307,609,1637]" captionTargetPageId="26" captionText="FIGURES 58 ­ 67. Genitalia of Seeversiella sonomotoides Gusarov, sp. n. (paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61 ­ 62, 64 ­ 66)." httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="26" pageNumber="27">Fig. 62</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C334C76AE11E75AAFDCDF8A0" pageId="26" pageNumber="27" type="distribution">
<paragraph id="EA1E3692C334C76AE11E75AAFB20FE1E" blockId="26.[264,1324,284,550]" box="[312,1178,443,470]" pageId="26" pageNumber="27">
<emphasis id="D8D5EA80C334C76AE11E75AAFE62FE1D" bold="true" box="[312,472,443,469]" pageId="26" pageNumber="27">Distribution.</emphasis>
Known from the Huachuca Mountains, Arizona (
<figureCitation id="729A2A17C334C76AE40075ADFB34FE1E" box="[1062,1166,444,470]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="26" pageNumber="27">Fig. 386</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C334C76AE11E75F2FE32FDEE" blockId="26.[264,1324,284,550]" pageId="26" pageNumber="27">
<emphasis id="D8D5EA80C334C76AE11E75F2FDBEFE35" bold="true" box="[312,516,483,509]" pageId="26" pageNumber="27">Natural History.</emphasis>
<taxonomicName id="2DA14D11C334C76AE23575F5FCF0FE35" box="[531,842,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sonomotoides">
<emphasis id="D8D5EA80C334C76AE23575F5FCF0FE35" box="[531,842,484,509]" italics="true" pageId="26" pageNumber="27">Seeversiella sonomotoides</emphasis>
</taxonomicName>
was collected in oak leaf litter at altitude of
<quantity id="2D599B77C334C76AE10E761DFE3EFDEE" box="[296,388,524,550]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="26" pageNumber="27" unit="m" value="2150.0">2150 m</quantity>
.
</paragraph>
<caption id="BEDE661AC334C76AE12E726EFC27F88C" httpUri="https://zenodo.org/record/156432/files/figure.png" pageId="26" pageNumber="27" targetBox="[289,1302,621,1628]" targetPageId="26">
<paragraph id="EA1E3692C334C76AE12E726EFC27F88C" blockId="26.[264,1324,1663,1896]" pageId="26" pageNumber="27">
<emphasis id="D8D5EA80C334C76AE12E726EFE7DF95F" bold="true" box="[264,455,1663,1687]" pageId="26" pageNumber="27">FIGURES 58­67</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C334C76AE24A7291FC54F95F" authority="Gusarov" authorityName="Gusarov" box="[620,1006,1664,1687]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sonomotoides" status="sp. nov.">
<emphasis id="D8D5EA80C334C76AE24A7291FC30F95F" box="[620,906,1664,1687]" italics="true" pageId="26" pageNumber="27">Seeversiella sonomotoides</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C334C76AE3DA726EFB81F95F" bold="true" box="[1020,1083,1663,1687]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="C3E657FBC334C76AE3DA726EFB81F95F" box="[1020,1083,1663,1687]" pageId="26" pageNumber="27" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Carr Canyon, Arizona). 58 – median lobe, parameral view; 59 – apex of median lobe, parameral view; 60 – median lobe, lateral view; 61 – apex of median lobe, lateral view; 62 – spermatheca; 63 – details of internal sac retracted into median lobe, lateral view; 64 – apex of left paramere; 65 – right hook of internal sac; 66 – copulatory piece, lateral view; 67 – everted internal sac, parameral view. Scale bar 0.2 mm (58, 60, 63, 67), 0.1 mm (59, 61­62, 64­66).
</paragraph>
</caption>
<paragraph id="EA1E3692C334C76AE12E7340FDCDF8A0" blockId="26.[264,1324,1663,1896]" box="[264,631,1873,1896]" pageId="26" pageNumber="27">DS – distal sclerite of internal sac.</paragraph>
</subSubSection>
</treatment>
</document>

255
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2.
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<emphasis id="D8D5EA80C33BC765E10075F5FDB7FE36" bold="true" box="[294,525,484,510]" italics="true" pageId="21" pageNumber="22">Seeversiella texana</emphasis>
Gusarov
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,
<taxonomicNameLabel id="C3E657FBC33BC765E2AB75F2FD6AFE35" box="[653,720,483,509]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C33BC765E2F975F2FCD0FE35" box="[735,874,483,509]" captionStart-0="FIGURES 37 ­ 40" captionStart-1="FIGURES 41 ­ 44" captionStart-2="FIGURES 45 ­ 53" captionStartId-0="21.[264,383,1773,1797]" captionStartId-1="22.[264,383,1298,1322]" captionStartId-2="23.[264,383,1314,1338]" captionTargetBox-0="[274,1312,896,1738]" captionTargetBox-1="[293,1296,576,1260]" captionTargetBox-2="[313,1274,323,1277]" captionTargetId-0="figure@21.[264,1323,884,1747]" captionTargetId-1="figure@22.[281,1306,569,1271]" captionTargetId-2="figure@23.[306,1280,313,1287]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="FIGURES 37 ­ 40. Abdominal segment 8 of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 37 – male tergum 8; 38 – male sternum 8; 39 – female tergum 8; 40 – female sternum 8. Scale bar 0.4 mm." captionText-1="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 – median lobe, parameral view; 42 – apex of median lobe, parameral view; 43 – median lobe, lateral view; 44 – apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-2="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156428/files/figure.png" httpUri-1="https://zenodo.org/record/156429/files/figure.png" httpUri-2="https://zenodo.org/record/156430/files/figure.png" pageId="21" pageNumber="22">Figs. 37­53</figureCitation>
)
</keyLead>
</emphasis>
</keyStep>
</key>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C33BC765E12E763FFD6EFCD9" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="EA1E3692C33BC765E12E763FFC83FD51" blockId="21.[264,1325,558,785]" pageId="21" pageNumber="22">
<emphasis id="D8D5EA80C33BC765E12E763FFE7BFD80" bold="true" box="[264,449,558,584]" pageId="21" pageNumber="22">
<typeStatus id="351A8830C33BC765E12E763FFEFFFD80" box="[264,325,558,584]" pageId="21" pageNumber="22">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C33BC765E1F3763EFDFFFD81" box="[469,581,559,585]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C33BC765E25E763FFC75FD80" bold="true" box="[632,975,558,584]" pageId="21" pageNumber="22">
<collectingCountry id="92B67602C33BC765E25E763FFCDCFD80" box="[632,870,558,584]" name="United States of America" pageId="21" pageNumber="22">UNITED STATES</collectingCountry>
: Texas:
</emphasis>
Culberson Co.:
<collectingCountry id="92B67602C33BC765E48A763EFA91FD81" box="[1196,1323,559,585]" name="Guadeloupe" pageId="21" pageNumber="22">Guadalupe</collectingCountry>
Mts. N. P., The Bowl, jct. Tejas Tr. and Juniper Tr.,
<quantity id="2D599B77C33BC765E3457646FC05FDB9" box="[867,959,599,625]" metricMagnitude="3" metricUnit="m" metricValue="2.407" pageId="21" pageNumber="22" unit="m" value="2407.0">2407 m</quantity>
, berlese maple, oak, pine, ironwood litter (R.Anderson),
<date id="9E1F1052C33BC765E219766EFD7FFD51" box="[575,709,639,665]" pageId="21" pageNumber="22" value="1989-07-31">31.vii.1989</date>
(
<collectionCode id="8CB0AE57C33BC765E2F2766EFC96FD51" box="[724,812,639,665]" pageId="21" pageNumber="22">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C33BC765E11E76B6FD6EFCD9" blockId="21.[264,1325,558,785]" pageId="21" pageNumber="22">
<typeStatus id="351A8830C33BC765E11E76B6FE15FD09" box="[312,431,679,705]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C33BC765E1E476B7FCA9FD08" bold="true" box="[450,787,678,704]" pageId="21" pageNumber="22">
<collectingCountry id="92B67602C33BC765E1E476B7FD17FD08" box="[450,685,678,704]" name="United States of America" pageId="21" pageNumber="22">UNITED STATES</collectingCountry>
: Texas:
</emphasis>
Culberson Co.:
<specimenCount id="FCA7FD1BC33BC765E3C076B6FB33FD09" box="[998,1161,679,705]" pageId="21" pageNumber="22" type="generic">14 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C33BC765E11176DEFE1BFD21" box="[311,417,719,745]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
;
<specimenCount id="FCA7FD1BC33BC765E19676DEFDFCFD21" box="[432,582,719,745]" pageId="21" pageNumber="22" type="generic">4 specimens</specimenCount>
, ditto but mixed hardwood litter;
<specimenCount id="FCA7FD1BC33BC765E3C276DEFBC0FD21" box="[996,1146,719,745]" pageId="21" pageNumber="22" type="generic">8 specimens</specimenCount>
, ditto but oak, conifer litter,
<date id="9E1F1052C33BC765E18176E6FD9FFCD9" box="[423,549,759,785]" pageId="21" pageNumber="22" value="1988-09-13">13.ix.1988</date>
(all –
<collectionCode id="8CB0AE57C33BC765E25676E6FD7DFCD9" box="[624,711,759,785]" pageId="21" pageNumber="22">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="BEDE661AC33BC765E12E72FCFC32F882" httpUri="https://zenodo.org/record/156428/files/figure.png" pageId="21" pageNumber="22" targetBox="[274,1312,896,1738]" targetPageId="21">
<paragraph id="EA1E3692C33BC765E12E72FCFC32F882" blockId="21.[264,1323,1773,1866]" pageId="21" pageNumber="22">
<emphasis id="D8D5EA80C33BC765E12E72FCFE7FF8CD" bold="true" box="[264,453,1773,1797]" pageId="21" pageNumber="22">FIGURES 37­40</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C33BC765E2D072FFFB90F8CD" authority="Gusarov" authorityName="Gusarov" box="[758,1066,1774,1797]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C33BC765E2D072FFFC7DF8CD" box="[758,967,1774,1797]" italics="true" pageId="21" pageNumber="22">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C33BC765E41E72FCFBCCF8CD" bold="true" box="[1080,1142,1773,1797]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="C3E657FBC33BC765E41E72FCFBCCF8CD" box="[1080,1142,1773,1797]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Guadalupe Mountains National Park, Texas). 37 – male tergum 8; 38 – male sternum 8; 39 – female tergum 8; 40 – female sternum 8. Scale bar 0.4 mm.
</paragraph>
</caption>
<subSubSection id="A2BB6519C338C766E11E750AFC9CF969" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="EA1E3692C338C766E11E750AFD8AFE36" blockId="22.[264,1323,283,510]" pageId="22" pageNumber="23">
<emphasis id="D8D5EA80C338C766E11E750AFE02FEFD" bold="true" box="[312,440,283,309]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C338C766E1E8750DFD0FFEFD" box="[462,693,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C338C766E1E8750DFD0FFEFD" box="[462,693,284,309]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C338C766E4F5750DFEFDFE95" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C338C766E4F5750DFEFDFE95" italics="true" pageId="22" pageNumber="23">Seeversiella</emphasis>
</taxonomicName>
by having brown or brownish yellow body; small eyes (temples 2.1­2.7 times as long as eyes); glossy pronotum with weak microsculpture; wings reduced, 2 times as long as elytra; tergum 8 with four pairs of macrosetae; the distinct shape of aedeagus (
<figureCitation id="729A2A17C338C766E4E77585FE92FE1E" captionStart-0="FIGURES 41 ­ 44" captionStart-1="FIGURES 45 ­ 53" captionStartId-0="22.[264,383,1298,1322]" captionStartId-1="23.[264,383,1314,1338]" captionTargetBox-0="[293,1296,576,1260]" captionTargetBox-1="[313,1274,323,1277]" captionTargetId-0="figure@22.[281,1306,569,1271]" captionTargetId-1="figure@23.[306,1280,313,1287]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 – median lobe, parameral view; 42 – apex of median lobe, parameral view; 43 – median lobe, lateral view; 44 – apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-1="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156429/files/figure.png" httpUri-1="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Figs. 41­ 52</figureCitation>
), especially the hook­shaped distal sclerites of internal sac (
<figureCitation id="729A2A17C338C766E3C175ADFBCCFE1E" box="[999,1142,444,470]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Figs. 48, 51</figureCitation>
); and the shape of spermatheca (
<figureCitation id="729A2A17C338C766E1EC75F5FD98FE36" box="[458,546,484,510]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Fig. 53</figureCitation>
).
</paragraph>
<caption id="BEDE661AC338C766E12E7103FDCFFA59" httpUri="https://zenodo.org/record/156429/files/figure.png" pageId="22" pageNumber="23" targetBox="[293,1296,576,1260]" targetPageId="22">
<paragraph id="EA1E3692C338C766E12E7103FDCFFA59" blockId="22.[264,1323,1298,1425]" pageId="22" pageNumber="23">
<emphasis id="D8D5EA80C338C766E12E7103FE7DFAE2" bold="true" box="[264,455,1298,1322]" pageId="22" pageNumber="23">FIGURES 41­44</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C338C766E2527103FC10FAE1" authority="Gusarov" authorityName="Gusarov" box="[628,938,1298,1321]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C338C766E2527103FCFFFAE1" box="[628,837,1298,1321]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C338C766E39E7103FC4CFAE2" bold="true" box="[952,1014,1298,1322]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="C3E657FBC338C766E39E7103FC4CFAE2" box="[952,1014,1298,1322]" pageId="22" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Guadalupe Mountains National Park, Texas). 41 – median lobe, parameral view; 42 – apex of median lobe, parameral view; 43 – median lobe, lateral view; 44 – apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44).
</paragraph>
</caption>
<paragraph id="EA1E3692C338C766E11E71F9FC9CF969" blockId="22.[264,1323,1511,1897]" pageId="22" pageNumber="23">
<taxonomicName id="2DA14D11C338C766E11E71F9FDA4F9C9" box="[312,542,1512,1537]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C338C766E11E71F9FDA4F9C9" box="[312,542,1512,1537]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
</taxonomicName>
differs from closely related
<taxonomicName id="2DA14D11C338C766E35E71F9FBAFF9C9" box="[888,1045,1512,1537]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="globicollis">
<emphasis id="D8D5EA80C338C766E35E71F9FBAFF9C9" box="[888,1045,1512,1537]" italics="true" pageId="22" pageNumber="23">S. globicollis</emphasis>
</taxonomicName>
in having lighter body colour; more distinct microsculpture of head and pronotum; elytra shorter than pronotum; reduced wings, 2 times as long as elytra; apex of median lobe bent paramerally (in lateral view) (
<figureCitation id="729A2A17C338C766E17B724EFE53F9B1" box="[349,489,1631,1657]" captionStart="FIGURES 41 ­ 44" captionStartId="22.[264,383,1298,1322]" captionTargetBox="[293,1296,576,1260]" captionTargetId="figure@22.[281,1306,569,1271]" captionTargetPageId="22" captionText="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 – median lobe, parameral view; 42 – apex of median lobe, parameral view; 43 – median lobe, lateral view; 44 – apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." httpUri="https://zenodo.org/record/156429/files/figure.png" pageId="22" pageNumber="23">Figs. 43­44</figureCitation>
; 26­27), shorter copulatory piece of internal sac (
<figureCitation id="729A2A17C338C766E41B724EFB42F9B1" box="[1085,1272,1631,1657]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Figs. 46, 49­50</figureCitation>
; 29, 34) and spermatheca with umbilicus (
<figureCitation id="729A2A17C338C766E2E77296FCA3F969" box="[705,793,1671,1697]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="22" pageNumber="23">Fig. 53</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C338C768E11E72A1FDD5FCF6" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
<paragraph id="EA1E3692C338C766E11E72A1FBF5F8D1" blockId="22.[264,1323,1511,1897]" pageId="22" pageNumber="23">
<taxonomicName id="2DA14D11C338C766E11E72A1FDA5F901" box="[312,543,1712,1737]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C338C766E11E72A1FDA5F901" box="[312,543,1712,1737]" italics="true" pageId="22" pageNumber="23">Seeversiella texana</emphasis>
</taxonomicName>
differs from the other similar species of
<taxonomicName id="2DA14D11C338C766E43872A1FB16F901" box="[1054,1196,1712,1737]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C338C766E43872A1FB16F901" box="[1054,1196,1712,1737]" italics="true" pageId="22" pageNumber="23">Seeversiella</emphasis>
</taxonomicName>
with long elytra and weak pronotal microsculpture in having strongly sclerotized hook­shaped distal sclerites of internal sac and J­shaped spermatheca with large umbilicus.
</paragraph>
<paragraph id="EA1E3692C338C766E11E7336FB32F8A1" blockId="22.[264,1323,1511,1897]" pageId="22" pageNumber="23">
<emphasis id="D8D5EA80C338C766E11E7336FE6AF889" bold="true" box="[312,464,1831,1857]" pageId="22" pageNumber="23">Description.</emphasis>
Length
<quantity id="2D599B77C338C766E2677336FD75F889" box="[577,719,1831,1857]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="22" pageNumber="23" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5­2.8 mm</quantity>
. Body from brownish yellow to brown, head and abdominal segments 6­7 often darker, legs and mouthparts brownish yellow.
</paragraph>
<caption id="BEDE661AC339C767E12E7133FDC5FA2E" httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="23" pageNumber="24" targetBox="[313,1274,323,1277]" targetPageId="23">
<paragraph id="EA1E3692C339C767E12E7133FDC5FA2E" blockId="23.[264,1323,1314,1545]" pageId="23" pageNumber="24">
<emphasis id="D8D5EA80C339C767E12E7133FE7DFAF2" bold="true" box="[264,455,1314,1338]" pageId="23" pageNumber="24">FIGURES 45­53</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C339C767E2497133FC1DFAF1" authority="Gusarov" authorityName="Gusarov" box="[623,935,1314,1337]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="texana" status="sp. nov.">
<emphasis id="D8D5EA80C339C767E2497133FCFBFAF1" box="[623,833,1314,1337]" italics="true" pageId="23" pageNumber="24">Seeversiella texana</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C339C767E3907133FC4FFAF2" bold="true" box="[950,1013,1314,1338]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="C3E657FBC339C767E3907133FC4FFAF2" box="[950,1013,1314,1338]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48­49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48­49, 53), 0.1 mm (46­47, 50­52).
</paragraph>
</caption>
<paragraph id="EA1E3692C339C767E12E71E3FDCDF9C1" blockId="23.[264,1323,1314,1545]" box="[264,631,1522,1545]" pageId="23" pageNumber="24">DS – distal sclerite of internal sac.</paragraph>
<paragraph id="EA1E3692C339C767E11E7274FBD5F93F" blockId="23.[264,1324,1637,1903]" pageId="23" pageNumber="24">
Head surface glossy, with weak isodiametric microsculpture, with fine, weak and poorly visible (at
<date id="9E1F1052C339C767E1F0729CFDBEF96F" box="[470,516,1677,1703]" pageId="23" pageNumber="24" value="1970-10">70x</date>
) punctation, distance between punctures equals 2­3 times their diameter. Temples 2.1­2.7 times as long as eyes. Antennal article 2 longer than article 3, article 4 slightly transverse, 5­10 transverse or strongly transverse (ratio 1.5­2.0).
</paragraph>
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Pronotum slightly transverse, 1.2 times as wide as head, width
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, length
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, width to length ratio 1.1, surface glossy, with weak isodiametric microsculpture; punctation as on head. Elytra wider and shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.1), 1.4 times wider than long, glossy, with weak isodiametric microsculpture, with fine and slightly asperate punctation, distance between punctures equal to 1­2 times their diameter. Wings reduced, 2 times as long as elytra.
</paragraph>
<paragraph id="EA1E3692C336C768E11E75ADFCA7FD86" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 2­3 times their diameter on terga 3­5 and 3­6 times on tergum 7. Apical margin of tergum 7 with white palisade fringe. Tergum 8 with four pairs of macrosetae (37, 39).</paragraph>
<paragraph id="EA1E3692C336C768E11E764DFA92FD0E" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">
In males posterior angles of tergum 3 projecting as spines up to half as long as tergum 3 (measured medially), tergum 7 with medial carina along midline, the carina up to half as long as tergum length. Posterior margin of male tergum 8 without emargination (
<figureCitation id="729A2A17C336C768E4E576BDFAA0FD0E" box="[1219,1306,684,710]" captionStart="FIGURES 37 ­ 40" captionStartId="21.[264,383,1773,1797]" captionTargetBox="[274,1312,896,1738]" captionTargetId="figure@21.[264,1323,884,1747]" captionTargetPageId="21" captionText="FIGURES 37 ­ 40. Abdominal segment 8 of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 37 – male tergum 8; 38 – male sternum 8; 39 – female tergum 8; 40 – female sternum 8. Scale bar 0.4 mm." httpUri="https://zenodo.org/record/156428/files/figure.png" pageId="24" pageNumber="25">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E76C5FAA7FCDE" blockId="24.[264,1324,284,910]" pageId="24" pageNumber="25">
Aedeagus as in
<figureCitation id="729A2A17C336C768E1C976C5FDC3FD26" box="[495,633,724,750]" captionStart-0="FIGURES 41 ­ 44" captionStart-1="FIGURES 45 ­ 53" captionStartId-0="22.[264,383,1298,1322]" captionStartId-1="23.[264,383,1314,1338]" captionTargetBox-0="[293,1296,576,1260]" captionTargetBox-1="[313,1274,323,1277]" captionTargetId-0="figure@22.[281,1306,569,1271]" captionTargetId-1="figure@23.[306,1280,313,1287]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURES 41 ­ 44. Aedeagus of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 41 – median lobe, parameral view; 42 – apex of median lobe, parameral view; 43 – median lobe, lateral view; 44 – apex of median lobe, lateral view. Scale bar 0.2 mm (41, 43), 0.1 mm (42, 44)." captionText-1="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri-0="https://zenodo.org/record/156429/files/figure.png" httpUri-1="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Figs. 41­52</figureCitation>
. Distal sclerites of internal sac hook­shaped (
<figureCitation id="729A2A17C336C768E4B676C5FAA5FD26" box="[1168,1311,724,750]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Figs. 48, 51</figureCitation>
). Proximal seta of the apex of paramere is much longer than the other three setae (
<figureCitation id="729A2A17C336C768E49E76EDFAAAFCDE" box="[1208,1296,764,790]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Fig. 52</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E7735FDD5FCF6" blockId="24.[264,1324,284,910]" box="[312,623,804,830]" pageId="24" pageNumber="25">
Spermatheca as in
<figureCitation id="729A2A17C336C768E2337735FDD6FCF6" box="[533,620,804,830]" captionStart="FIGURES 45 ­ 53" captionStartId="23.[264,383,1314,1338]" captionTargetBox="[313,1274,323,1277]" captionTargetId="figure@23.[306,1280,313,1287]" captionTargetPageId="23" captionText="FIGURES 45 ­ 53. Genitalia of Seeversiella texana Gusarov, sp. n. (paratypes from Guadalupe Mountains National Park, Texas). 45 – details of internal sac retracted into median lobe, abparameral view; 46 – copulatory piece, parameral view; 47 – medial lamellae, parameral view; 48 ­ 49 – details of internal sac retracted into median lobe, lateral view; 50 – copulatory piece, lateral view; 51 – right hook of internal sac; 52 – apex of left paramere; 53 – spermatheca. Scale bar 0.2 mm (45, 48 ­ 49, 53), 0.1 mm (46 ­ 47, 50 ­ 52)." httpUri="https://zenodo.org/record/156430/files/figure.png" pageId="24" pageNumber="25">Fig. 53</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C336C768E11E775AFA96FC46" pageId="24" pageNumber="25" type="distribution">
<paragraph id="EA1E3692C336C768E11E775AFB31FCAE" blockId="24.[264,1324,284,910]" box="[312,1163,843,870]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E11E775AFE62FCAD" bold="true" box="[312,472,843,869]" pageId="24" pageNumber="25">Distribution.</emphasis>
Known from the
<collectingCountry id="92B67602C336C768E296775DFC95FCAE" box="[688,815,844,870]" name="Guadeloupe" pageId="24" pageNumber="25">Guadalupe</collectingCountry>
Mountains, Texas (
<figureCitation id="729A2A17C336C768E430775DFBC4FCAE" box="[1046,1150,844,870]" captionStart="FIGURE 386" captionStartId="96.[264,368,1354,1378]" captionTargetBox="[315,1274,412,1319]" captionTargetId="figure@96.[309,1279,409,1327]" captionTargetPageId="96" captionText="FIGURE 386. Geographical distribution of Seeversiella in America north of Mexico." httpUri="https://zenodo.org/record/156488/files/figure.png" pageId="24" pageNumber="25">Fig. 386</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C336C768E11E7762FA96FC46" blockId="24.[264,1324,284,910]" box="[312,1324,883,910]" pageId="24" pageNumber="25">
<emphasis id="D8D5EA80C336C768E11E7762FDBFFC45" bold="true" box="[312,517,883,909]" pageId="24" pageNumber="25">Natural History.</emphasis>
<taxonomicName id="2DA14D11C336C768E2357765FD4CFC45" box="[531,758,884,909]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="D8D5EA80C336C768E2357765FD4CFC45" box="[531,758,884,909]" italics="true" pageId="24" pageNumber="25">Seeversiella texana</emphasis>
</taxonomicName>
was collected at altitude of
<quantity id="2D599B77C336C768E41B7765FB2CFC46" box="[1085,1174,884,910]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="24" pageNumber="25" unit="m" value="2400.0">2400 m</quantity>
in leaf litter.
</paragraph>
</subSubSection>
</treatment>
</document>

231
data/62/08/87/62088784C360C722E12E727AFED2FE96.xml Normal file
View file

@ -0,0 +1,231 @@
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<paragraph id="EA1E3692C360C73EE12E727AFC7FF94D" blockId="78.[264,965,1643,1670]" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
<heading id="B15681FEC360C73EE12E727AFC7FF94D" bold="true" box="[264,965,1643,1670]" fontSize="11" level="1" pageId="78" pageNumber="79" reason="1">
<emphasis id="D8D5EA80C360C73EE12E727AFC7FF94D" bold="true" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
<key id="9E3061A3C360C73EE12E727AFC7FF94D" box="[264,965,1643,1670]" pageId="78" pageNumber="79">
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23.
<taxonomicName id="2DA14D11C360C73EE112727DFD0DF94D" ID-CoL="6YBBC" authority="Gusarov" authorityName="Gusarov" box="[308,695,1643,1670]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sulcicollis" status="sp. nov.">
<emphasis id="D8D5EA80C360C73EE112727DFDFEF94E" bold="true" box="[308,580,1644,1670]" italics="true" pageId="78" pageNumber="79">Seeversiella sulcicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC360C73EE2E5727AFCBCF94D" box="[707,774,1643,1669]" pageId="78" pageNumber="79" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C360C73EE333727AFC06F94D" box="[789,956,1643,1669]" captionStart-0="FIGURES 321 ­ 324" captionStart-1="FIGURES 325 ­ 330" captionStartId-0="79.[264,382,1679,1703]" captionStartId-1="80.[264,382,1592,1616]" captionTargetBox-0="[283,1308,537,1644]" captionTargetBox-1="[280,1306,336,1561]" captionTargetId-0="figure@79.[272,1316,529,1652]" captionTargetId-1="figure@80.[274,1313,329,1565]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 – median lobe, parameral view; 322 – apex of median lobe, parameral view; 323 – median lobe, lateral view; 324 – apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." captionText-1="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri-0="https://zenodo.org/record/156476/files/figure.png" httpUri-1="https://zenodo.org/record/156477/files/figure.png" pageId="78" pageNumber="79">Figs. 321­330</figureCitation>
)
</keyLead>
</keyStep>
</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C360C720E12E72A7FC86F8D7" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79" type="materials_examined">
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<emphasis id="D8D5EA80C360C73EE12E72A7FE04F918" bold="true" box="[264,446,1718,1744]" pageId="78" pageNumber="79">
<typeStatus id="351A8830C360C73EE12E72A7FEFFF918" box="[264,325,1718,1744]" pageId="78" pageNumber="79">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C360C73EE1EA72A7FD86F918" box="[460,572,1718,1744]" pageId="78" pageNumber="79" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C360C73EE24C72A7FBA6F918" bold="true" box="[618,1052,1718,1744]" pageId="78" pageNumber="79">
<collectingCountry id="92B67602C360C73EE24C72A7FC99F918" box="[618,803,1718,1744]" name="Costa Rica" pageId="78" pageNumber="79">COSTA RICA</collectingCountry>
: San Jose/Cartago:
</emphasis>
Cerro Buenavista, km 89, Int. Amer. Hwy.,
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<geoCoordinate id="8F955055C360C73EE2A472CFFCA8F930" box="[642,786,1758,1784]" direction="west" orientation="longitude" pageId="78" pageNumber="79" precision="15" value="-83.75833">83°45'30&quot;W</geoCoordinate>
,
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, forest litter (R.Anderson),
<date id="9E1F1052C360C73EE49A72CFFA90F930" box="[1212,1322,1758,1784]" pageId="78" pageNumber="79" value="1997-06-08">8.vi.1997</date>
(
<collectionCode id="8CB0AE57C360C73EE1367317FED2F8E8" box="[272,360,1798,1824]" pageId="78" pageNumber="79">KSEM</collectionCode>
).
</paragraph>
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<typeStatus id="351A8830C360C73EE11E733FFE14F880" box="[312,430,1838,1864]" pageId="78" pageNumber="79" type="paratype">Paratypes</typeStatus>
:
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<collectingCountry id="92B67602C360C73EE1E6733FFDC0F880" box="[448,634,1838,1864]" name="Costa Rica" pageId="78" pageNumber="79">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
, same data as the
<typeStatus id="351A8830C360C73EE3D0733FFBE7F880" box="[1014,1117,1838,1864]" pageId="78" pageNumber="79" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C360C73EE44A733FFB7EF880" box="[1132,1220,1838,1864]" pageId="78" pageNumber="79">KSEM</collectionCode>
).
<emphasis id="D8D5EA80C360C73EE11E7347FE02F8B8" bold="true" box="[312,440,1878,1904]" pageId="78" pageNumber="79">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C360C73EE1E17346FD69F8B8" box="[455,723,1879,1904]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C360C73EE1E17346FD69F8B8" box="[455,723,1879,1904]" italics="true" pageId="78" pageNumber="79">Seeversiella sulcicollis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C360C73FE4F57346FEFDFEFD" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" lastPageId="79" lastPageNumber="80" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C360C73FE4F57346FEFDFEFD" italics="true" lastPageId="79" lastPageNumber="80" pageId="78" pageNumber="79">Seeversiella</emphasis>
</taxonomicName>
by having reddish brown body; temples 1.8­2.3 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by having small medial emargination delimited by two short and blunt projections at the posterior margin of the male tergum 8 (
<figureCitation id="729A2A17C361C73FE1AE7585FE49FE66" box="[392,499,404,430]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="79" pageNumber="80">Fig. 330</figureCitation>
); tergum 8 with four pairs of macrosetae; and the distinct shape of aedeagus (
<figureCitation id="729A2A17C361C73FE1A575ADFD96FE1E" box="[387,556,444,470]" captionStart-0="FIGURES 321 ­ 324" captionStart-1="FIGURES 325 ­ 330" captionStartId-0="79.[264,382,1679,1703]" captionStartId-1="80.[264,382,1592,1616]" captionTargetBox-0="[283,1308,537,1644]" captionTargetBox-1="[280,1306,336,1561]" captionTargetId-0="figure@79.[272,1316,529,1652]" captionTargetId-1="figure@80.[274,1313,329,1565]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 – median lobe, parameral view; 322 – apex of median lobe, parameral view; 323 – median lobe, lateral view; 324 – apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." captionText-1="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri-0="https://zenodo.org/record/156476/files/figure.png" httpUri-1="https://zenodo.org/record/156477/files/figure.png" pageId="79" pageNumber="80">Figs. 321­328</figureCitation>
).
</paragraph>
<caption id="BEDE661AC361C73FE12E729EFB76F923" httpUri="https://zenodo.org/record/156476/files/figure.png" pageId="79" pageNumber="80" targetBox="[283,1308,537,1644]" targetPageId="79">
<paragraph id="EA1E3692C361C73FE12E729EFB76F923" blockId="79.[264,1322,1679,1771]" pageId="79" pageNumber="80">
<emphasis id="D8D5EA80C361C73FE12E729EFE67F96F" bold="true" box="[264,477,1679,1703]" pageId="79" pageNumber="80">FIGURES 321­324</emphasis>
. Aedeagus of
<taxonomicName id="2DA14D11C361C73FE25B729EFC68F96E" authority="Gusarov" authorityName="Gusarov" box="[637,978,1679,1702]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="sulcicollis" status="sp. nov.">
<emphasis id="D8D5EA80C361C73FE25B729EFCCAF96E" box="[637,880,1679,1702]" italics="true" pageId="79" pageNumber="80">Seeversiella sulcicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C361C73FE3FA729EFBADF96F" bold="true" box="[988,1047,1679,1703]" pageId="79" pageNumber="80">
<taxonomicNameLabel id="C3E657FBC361C73FE3FA729EFBADF96F" box="[988,1047,1679,1703]" pageId="79" pageNumber="80" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). 321 – median lobe, parameral view; 322 – apex of median lobe, parameral view; 323 – median lobe, lateral view; 324 – apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324).
</paragraph>
</caption>
<caption id="BEDE661AC37EC720E12E7229FE10F934" httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="80" pageNumber="81" targetBox="[280,1306,336,1561]" targetPageId="80">
<paragraph id="EA1E3692C37EC720E12E7229FE10F934" blockId="80.[264,1323,1592,1823]" pageId="80" pageNumber="81">
<emphasis id="D8D5EA80C37EC720E12E7229FE67F998" bold="true" box="[264,477,1592,1616]" pageId="80" pageNumber="81">FIGURES 325­330</emphasis>
. Abdominal segment 8 and aedeagus of
<taxonomicName id="2DA14D11C37EC720E3B77229FB5FF987" authority="Gusarov" authorityName="Gusarov" box="[913,1253,1592,1615]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="sulcicollis" status="sp. nov.">
<emphasis id="D8D5EA80C37EC720E3B77229FB3EF987" box="[913,1156,1592,1615]" italics="true" pageId="80" pageNumber="81">Seeversiella sulcicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC37EC720E4C97229FA91F998" box="[1263,1323,1592,1616]" pageId="80" pageNumber="81" rank="species">
<emphasis id="D8D5EA80C37EC720E4C97229FA91F998" bold="true" box="[1263,1323,1592,1616]" pageId="80" pageNumber="81">sp. n.</emphasis>
</taxonomicNameLabel>
(325, 327­330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326­328), 0.4 mm (329­330).
</paragraph>
</caption>
<paragraph id="EA1E3692C37EC720E12E7319FC86F8D7" blockId="80.[264,1323,1592,1823]" box="[264,828,1800,1823]" pageId="80" pageNumber="81">SLD – sclerite of lateral diverticulum of internal sac.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C37FC721E11E750DFDB1F8EE" pageId="81" pageNumber="82" type="description">
<paragraph id="EA1E3692C37FC721E11E750DFE76FDEE" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
<taxonomicName id="2DA14D11C37FC721E11E750DFDFCFEFD" box="[312,582,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C37FC721E11E750DFDFCFEFD" box="[312,582,284,309]" italics="true" pageId="81" pageNumber="82">Seeversiella sulcicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37FC721E2C1750DFC12FEFD" box="[743,936,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C37FC721E2C1750DFC12FEFD" box="[743,936,284,309]" italics="true" pageId="81" pageNumber="82">S. impressicollis</emphasis>
</taxonomicName>
in having larger body; in males pronotum with medial furrow, lateral portions of posterior margin of tergum 3 extending as short and obtuse projections (as in
<figureCitation id="729A2A17C37FC721E298757DFCA9FE4E" box="[702,787,364,390]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="81" pageNumber="82">Fig. 19</figureCitation>
), tergum 7 with short medial carina; posterior margin of tergum 8 with two medial denticles (
<figureCitation id="729A2A17C37FC721E3647585FC11FE66" box="[834,939,404,430]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
); apex of median lobe with two divergent carinae (in parameral view) (
<figureCitation id="729A2A17C37FC721E2C375ADFC28FE1E" box="[741,914,444,470]" captionStart="FIGURES 321 ­ 324" captionStartId="79.[264,382,1679,1703]" captionTargetBox="[283,1308,537,1644]" captionTargetId="figure@79.[272,1316,529,1652]" captionTargetPageId="79" captionText="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 – median lobe, parameral view; 322 – apex of median lobe, parameral view; 323 – median lobe, lateral view; 324 – apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." httpUri="https://zenodo.org/record/156476/files/figure.png" pageId="81" pageNumber="82">Figs. 321­322</figureCitation>
); copulatory piece of internal sac longer (
<figureCitation id="729A2A17C37FC721E14175F5FDA6FE36" box="[359,540,484,510]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Figs. 326, 328</figureCitation>
; 317); sclerites of lateral diverticula of internal sac more robust (
<figureCitation id="729A2A17C37FC721E137761DFE38FDEE" box="[273,386,524,550]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Figs. 326</figureCitation>
; 318).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E7625FEFFFD26" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
<taxonomicName id="2DA14D11C37FC721E11E7625FDFEFD85" box="[312,580,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C37FC721E11E7625FDFEFD85" box="[312,580,564,589]" italics="true" pageId="81" pageNumber="82">Seeversiella sulcicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C37FC721E2C77625FC08FD85" box="[737,946,564,589]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C37FC721E2C77625FC08FD85" box="[737,946,564,589]" italics="true" pageId="81" pageNumber="82">S. microphthalma</emphasis>
</taxonomicName>
in having more asperate pronotal punctation; in males pronotum with medial furrow, posterior angles of tergum 3 not projecting as spines, posterior margin of tergum 8 with two medial denticles (
<figureCitation id="729A2A17C37FC721E4947695FAA6FD56" box="[1202,1308,644,670]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
); and the apex of median lobe with two divergent carinae (in parameral view) (
<figureCitation id="729A2A17C37FC721E48976BDFE82FD26" captionStart="FIGURES 321 ­ 324" captionStartId="79.[264,382,1679,1703]" captionTargetBox="[283,1308,537,1644]" captionTargetId="figure@79.[272,1316,529,1652]" captionTargetPageId="79" captionText="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 – median lobe, parameral view; 322 – apex of median lobe, parameral view; 323 – median lobe, lateral view; 324 – apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." httpUri="https://zenodo.org/record/156476/files/figure.png" pageId="81" pageNumber="82">Figs. 321­ 322</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E76EAFC1AFCDE" blockId="81.[264,1324,284,1870]" box="[312,928,763,790]" pageId="81" pageNumber="82">
<emphasis id="D8D5EA80C37FC721E11E76EAFE6AFCDD" bold="true" box="[312,464,763,789]" pageId="81" pageNumber="82">Description.</emphasis>
Length 3.0 mm. Body reddish brown.
</paragraph>
<paragraph id="EA1E3692C37FC721E11E7735FDB1FC7E" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">Head surface glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter. Temples 1.8­2.3 times as long as eyes. Antennal article 2 as long as 3, articles 4­6 subquadrate, 7­ 10 slightly transverse.</paragraph>
<paragraph id="EA1E3692C37FC721E11E77D5FCD3FB3E" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
Pronotum slightly transverse, 1.3 times as wide as head, width
<quantity id="2D599B77C37FC721E40D77D5FB6CFC16" box="[1067,1238,964,990]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="81" pageNumber="82" unit="mm" value="0.665" valueMax="0.67" valueMin="0.66">0.66­0.67 mm</quantity>
, length
<quantity id="2D599B77C37FC721E12E77FDFE0BFBCE" box="[264,433,1004,1030]" metricMagnitude="-4" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.0" metricValueMin="5.8999999999999995" pageId="81" pageNumber="82" unit="mm" value="0.595" valueMax="0.6" valueMin="0.59">0.59­0.60 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­1.5 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.8 times wider than long, glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter, punctures stronger than on pronotum. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C37FC721E11E7115FDD5FA2E" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
Abdominal terga glossy, terga 3­5 with microsculpture consisting of transverse meshes, and with fine punctation, distance between punctures equals 1­4 times their diameter; tergum 7 with microsculpture consisting of isodiametric or slightly transverse meshes and with fine and sparse punctation, distance between punctures equals 3­6 times their diameter. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C37FC721E1DD71DDFDD9FA2E" box="[507,611,1484,1510]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E71E5FD43F966" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
In males pronotum with medial furrow; lateral portions of posterior margin of tergum 3 extending as short and blunt projections up to 1/8 as long as tergum (measured medially) (as in
<figureCitation id="729A2A17C37FC721E1777255FE1DF996" box="[337,423,1604,1630]" captionStart="FIGURES 17 ­ 19" captionStartId="7.[264,383,1739,1763]" captionTargetBox="[407,1184,936,1705]" captionTargetId="figure@7.[399,1188,929,1712]" captionTargetPageId="7" captionText="FIGURES 17 ­ 19. Body outline of Seeversiella globicollis (Bernhauer) (17, Volcán Tacana, Mexico), S. micralymma Gusarov, sp. n. (18, paratype from Cerro de la Muerte, Costa Rica) and S. flavida Gusarov, sp. n. (19, paratype from Cerro de la Muerte, Costa Rica). Scale bar 1 mm." httpUri="https://zenodo.org/record/156424/files/figure.png" pageId="81" pageNumber="82">Fig. 19</figureCitation>
); tergum
<quantity id="2D599B77C37FC721E23F7255FDFDF996" box="[537,583,1604,1630]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="81" pageNumber="82" unit="in" value="7.0">7 in</quantity>
front of posterior margin with medial carina, up to 1/4 as long as the tergum; posterior margin of tergum 8 with small medial emargination delimited by two short and blunt projections (
<figureCitation id="729A2A17C37FC721E2A37285FD57F966" box="[645,749,1684,1710]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 330</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E72ADFD48F936" blockId="81.[264,1324,284,1870]" pageId="81" pageNumber="82">
Aedeagus as in
<figureCitation id="729A2A17C37FC721E1C872ADFD2EF91E" box="[494,660,1724,1750]" captionStart-0="FIGURES 321 ­ 324" captionStart-1="FIGURES 325 ­ 330" captionStartId-0="79.[264,382,1679,1703]" captionStartId-1="80.[264,382,1592,1616]" captionTargetBox-0="[283,1308,537,1644]" captionTargetBox-1="[280,1306,336,1561]" captionTargetId-0="figure@79.[272,1316,529,1652]" captionTargetId-1="figure@80.[274,1313,329,1565]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURES 321 ­ 324. Aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (holotype). 321 – median lobe, parameral view; 322 – apex of median lobe, parameral view; 323 – median lobe, lateral view; 324 – apex of median lobe, lateral view. Scale bar 0.2 mm (321, 323), 0.1 mm (322, 324)." captionText-1="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri-0="https://zenodo.org/record/156476/files/figure.png" httpUri-1="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Figs. 321­328</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C37FC721E25872F5FD5FF936" box="[638,741,1764,1790]" captionStart="FIGURES 325 ­ 330" captionStartId="80.[264,382,1592,1616]" captionTargetBox="[280,1306,336,1561]" captionTargetId="figure@80.[274,1313,329,1565]" captionTargetPageId="80" captionText="FIGURES 325 ­ 330. Abdominal segment 8 and aedeagus of Seeversiella sulcicollis Gusarov, sp. n. (325, 327 ­ 330, holotype; 326, paratype from Cerro Buenavista, Costa Rica). 325 – apex of left paramere; 326 – everted internal sac, parameral view; 327 – details of internal sac retracted into median lobe, abparameral view; 328 – details of internal sac retracted into median lobe, lateral view; 329 – male sternum 8; 330 – male tergum 8. Scale bar 0.1 mm (325), 0.2 mm (326 ­ 328), 0.4 mm (329 ­ 330)." httpUri="https://zenodo.org/record/156477/files/figure.png" pageId="81" pageNumber="82">Fig. 325</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37FC721E11E731DFDB1F8EE" blockId="81.[264,1324,284,1870]" box="[312,523,1804,1830]" pageId="81" pageNumber="82">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C37FC722E11E7322FED2FE96" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="distribution">
<paragraph id="EA1E3692C37FC721E11E7322FBCDF886" blockId="81.[264,1324,284,1870]" box="[312,1143,1843,1870]" pageId="81" pageNumber="82">
<emphasis id="D8D5EA80C37FC721E11E7322FE62F885" bold="true" box="[312,472,1843,1869]" pageId="81" pageNumber="82">Distribution.</emphasis>
Known from Cerro de la Muerte,
<collectingCountry id="92B67602C37FC721E3547325FC49F886" box="[882,1011,1844,1870]" name="Costa Rica" pageId="81" pageNumber="82">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C37FC721E4257325FBD0F886" box="[1027,1130,1844,1870]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="81" pageNumber="82">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C37CC722E11E750AFED2FE96" blockId="82.[264,1323,283,350]" pageId="82" pageNumber="83">
<emphasis id="D8D5EA80C37CC722E11E750AFDB2FEFD" bold="true" box="[312,520,283,309]" pageId="82" pageNumber="83">Natural History.</emphasis>
<taxonomicName id="2DA14D11C37CC722E23A750DFC91FEFD" box="[540,811,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C37CC722E23A750DFC91FEFD" box="[540,811,284,309]" italics="true" pageId="82" pageNumber="83">Seeversiella sulcicollis</emphasis>
</taxonomicName>
was collected in forest litter at altitude of
<quantity id="2D599B77C37CC722E12E7555FEDEFE96" box="[264,356,324,350]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="82" pageNumber="83" unit="m" value="3300.0">3300 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

238
data/62/08/87/62088784C362C73EE12E750AFE5DF9C6.xml Normal file
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@ -0,0 +1,238 @@
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<mods:title id="43CB2D7B546D52CF95DD091CA37CFE25">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="A2BB6519C362C73CE12E750AFC4AFEFD" box="[264,1008,283,310]" pageId="76" pageNumber="77" type="key">
<paragraph id="EA1E3692C362C73CE12E750AFC4AFEFD" blockId="76.[264,1008,283,310]" box="[264,1008,283,310]" pageId="76" pageNumber="77">
<heading id="B15681FEC362C73CE12E750AFC4AFEFD" bold="true" box="[264,1008,283,310]" fontSize="11" level="1" pageId="76" pageNumber="77" reason="1">
<emphasis id="D8D5EA80C362C73CE12E750AFC4AFEFD" bold="true" box="[264,1008,283,310]" pageId="76" pageNumber="77">
<key id="9E3061A3C362C73CE12E750AFC4AFEFD" box="[264,1008,283,310]" pageId="76" pageNumber="77">
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22.
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<emphasis id="D8D5EA80C362C73CE112750DFDCAFEFE" bold="true" box="[308,624,284,310]" italics="true" pageId="76" pageNumber="77">Seeversiella impressicollis</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="C3E657FBC362C73CE2C9750AFC8BFEFD" box="[751,817,283,309]" pageId="76" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="729A2A17C362C73CE366750AFC52FEFD" box="[832,1000,283,309]" captionStart-0="FIGURES 307 ­ 310" captionStart-1="FIGURES 311 ­ 320" captionStartId-0="76.[264,383,1792,1816]" captionStartId-1="77.[264,383,1623,1647]" captionTargetBox-0="[290,1292,893,1759]" captionTargetBox-1="[286,1302,571,1605]" captionTargetId-0="figure@76.[282,1305,884,1766]" captionTargetId-1="figure@77.[276,1310,569,1613]" captionTargetPageId-0="76" captionTargetPageId-1="77" captionText-0="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 – male tergum 8; 308 – male sternum 8; 309 – female tergum 8; 310 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri-0="https://zenodo.org/record/156474/files/figure.png" httpUri-1="https://zenodo.org/record/156475/files/figure.png" pageId="76" pageNumber="77">Figs. 307­320</figureCitation>
)
</keyLead>
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</key>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C362C73CE12E7577FECFFDE9" pageId="76" pageNumber="77" type="materials_examined">
<paragraph id="EA1E3692C362C73CE12E7577FECFFE19" blockId="76.[264,1323,358,785]" pageId="76" pageNumber="77">
<emphasis id="D8D5EA80C362C73CE12E7577FE05FE48" bold="true" box="[264,447,358,384]" pageId="76" pageNumber="77">
<typeStatus id="351A8830C362C73CE12E7577FEF9FE48" box="[264,323,358,384]" pageId="76" pageNumber="77">Type</typeStatus>
material.
</emphasis>
<typeStatus id="351A8830C362C73CE1F17576FDFDFE49" box="[471,583,359,385]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
,,
<emphasis id="D8D5EA80C362C73CE25D7577FC79FE48" bold="true" box="[635,963,358,384]" pageId="76" pageNumber="77">
<collectingCountry id="92B67602C362C73CE25D7577FC82FE48" box="[635,824,358,384]" name="Costa Rica" pageId="76" pageNumber="77">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
Cerro Chirripó, Valle de los Conelos,
<geoCoordinate id="8F955055C362C73CE15C759EFE4FFE61" box="[378,501,399,425]" direction="north" orientation="latitude" pageId="76" pageNumber="77" precision="15" value="9.476944">9°28'37&quot;N</geoCoordinate>
<geoCoordinate id="8F955055C362C73CE1D9759EFD35FE61" box="[511,655,399,425]" direction="west" orientation="longitude" pageId="76" pageNumber="77" precision="15" value="-83.489716">83°29'23&quot;W</geoCoordinate>
,
<quantity id="2D599B77C362C73CE2B9759EFD44FE61" box="[671,766,399,425]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="76" pageNumber="77" unit="m" value="3600.0">3600 m</quantity>
, paramo shrub litter (R.Anderson),
<date id="9E1F1052C362C73CE48B759EFA90FE61" box="[1197,1322,399,425]" pageId="76" pageNumber="77" value="1999-06-26">26.vi.1999</date>
(
<collectionCode id="8CB0AE57C362C73CE13675A6FED2FE19" box="[272,360,439,465]" pageId="76" pageNumber="77">KSEM</collectionCode>
).
</paragraph>
<paragraph id="EA1E3692C362C73CE11E75CEFECFFDE9" blockId="76.[264,1323,358,785]" pageId="76" pageNumber="77">
<typeStatus id="351A8830C362C73CE11E75CEFE15FE31" box="[312,431,479,505]" pageId="76" pageNumber="77" type="paratype">Paratypes</typeStatus>
:
<emphasis id="D8D5EA80C362C73CE1ED75CFFCA2FE30" bold="true" box="[459,792,478,504]" pageId="76" pageNumber="77">
<collectingCountry id="92B67602C362C73CE1ED75CFFD33FE30" box="[459,649,478,504]" name="Costa Rica" pageId="76" pageNumber="77">COSTA RICA</collectingCountry>
: Cartago:
</emphasis>
<specimenCount id="FCA7FD1BC362C73CE31775CEFC62FE31" box="[817,984,479,505]" pageId="76" pageNumber="77" type="generic">14 specimens</specimenCount>
, same data as the
<typeStatus id="351A8830C362C73CE4E575CEFA90FE31" box="[1219,1322,479,505]" pageId="76" pageNumber="77" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CB0AE57C362C73CE1367616FED2FDE9" box="[272,360,519,545]" pageId="76" pageNumber="77">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C362C73EE11E763FFE3BFE1E" lastPageId="78" lastPageNumber="79" pageId="76" pageNumber="77" type="diagnosis">
<paragraph id="EA1E3692C362C73CE11E763FFD78FCD9" blockId="76.[264,1323,358,785]" pageId="76" pageNumber="77">
<emphasis id="D8D5EA80C362C73CE11E763FFE0DFD80" bold="true" box="[312,439,558,584]" pageId="76" pageNumber="77">Diagnosis.</emphasis>
<taxonomicName id="2DA14D11C362C73CE1E0763EFD47FD80" box="[454,765,559,584]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C362C73CE1E0763EFD47FD80" box="[454,765,559,584]" italics="true" pageId="76" pageNumber="77">Seeversiella impressicollis</emphasis>
</taxonomicName>
can be distinguished from other species of
<taxonomicName id="2DA14D11C362C73CE4DE763EFED7FDB8" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="D8D5EA80C362C73CE4DE763EFED7FDB8" italics="true" pageId="76" pageNumber="77">Seeversiella</emphasis>
</taxonomicName>
by having brown or dark brown body with lighter pronotum and elytra; temples 1.8­2.2 times as long as eyes; elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking the medial emargination at the posterior margin of male tergum 7 (
<figureCitation id="729A2A17C362C73CE13776DEFECDFD21" box="[273,375,719,745]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 – male tergum 8; 308 – male sternum 8; 309 – female tergum 8; 310 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="76" pageNumber="77">Fig. 307</figureCitation>
); tergum 8 with four pairs of macrosetae; by the distinct shape of aedeagus (
<figureCitation id="729A2A17C362C73CE4D676DEFED0FCD9" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="76" pageNumber="77">Figs. 311­319</figureCitation>
) and spermatheca (
<figureCitation id="729A2A17C362C73CE26876E6FD0CFCD9" box="[590,694,759,785]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="76" pageNumber="77">Fig. 320</figureCitation>
).
</paragraph>
<caption id="BEDE661AC362C73CE12E7311FC2CF895" httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="76" pageNumber="77" targetBox="[290,1292,893,1759]" targetPageId="76">
<paragraph id="EA1E3692C362C73CE12E7311FC2CF895" blockId="76.[264,1323,1792,1885]" pageId="76" pageNumber="77">
<emphasis id="D8D5EA80C362C73CE12E7311FE52F8D0" bold="true" box="[264,488,1792,1816]" pageId="76" pageNumber="77">FIGURES 307­310</emphasis>
. Abdominal segment 8 of
<taxonomicName id="2DA14D11C362C73CE3647310FB6BF8D0" authority="Gusarov" authorityName="Gusarov" box="[834,1233,1793,1816]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="impressicollis" status="sp. nov.">
<emphasis id="D8D5EA80C362C73CE3647310FBDCF8D0" box="[834,1126,1793,1816]" italics="true" pageId="76" pageNumber="77">Seeversiella impressicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C362C73CE4C37311FA91F8D0" bold="true" box="[1253,1323,1792,1816]" pageId="76" pageNumber="77">
<taxonomicNameLabel id="C3E657FBC362C73CE4C37311FA91F8D0" box="[1253,1323,1792,1816]" pageId="76" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Cerro Chirripó, Costa Rica). 307 – male tergum 8; 308 – male sternum 8; 309 – female tergum 8; 310 – female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="EA1E3692C363C73DE11E750DFD43FE36" blockId="77.[264,1323,284,510]" pageId="77" pageNumber="78">
<taxonomicName id="2DA14D11C363C73DE11E750DFDD5FEFD" box="[312,623,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C363C73DE11E750DFDD5FEFD" box="[312,623,284,309]" italics="true" pageId="77" pageNumber="78">Seeversiella impressicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C363C73DE32A750DFC1AFEFD" box="[780,928,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="D8D5EA80C363C73DE32A750DFC1AFEFD" box="[780,928,284,309]" italics="true" pageId="77" pageNumber="78">S. sulcicollis</emphasis>
</taxonomicName>
in having smaller body; in males pronotum with broad medial impression, posterior margin of tergum 3 not projecting, tergum 7 without medial carina and posterior margin of tergum 8 without medial denticles (
<figureCitation id="729A2A17C363C73DE1377585FEC2FE66" box="[273,376,404,430]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 – male tergum 8; 308 – male sternum 8; 309 – female tergum 8; 310 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="77" pageNumber="78">Fig. 307</figureCitation>
); apex of median lobe in parameral view without carinae (
<figureCitation id="729A2A17C363C73DE40D7585FB69FE66" box="[1067,1235,404,430]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78">Figs. 311­312</figureCitation>
); copulatory piece of internal sac shorter (
<figureCitation id="729A2A17C363C73DE28975ADFC99FE1E" box="[687,803,444,470]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78">Figs. 317</figureCitation>
; 326, 328); sclerites of lateral diverticula of internal sac more slender (
<figureCitation id="729A2A17C363C73DE21875F5FD15FE36" box="[574,687,484,510]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78">Figs. 318</figureCitation>
; 326).
</paragraph>
<caption id="BEDE661AC363C73DE12E7246FEFAF8F7" httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="77" pageNumber="78" targetBox="[286,1302,571,1605]" targetPageId="77">
<paragraph id="EA1E3692C363C73DE12E7246FEFAF8F7" blockId="77.[264,1323,1623,1891]" pageId="77" pageNumber="78">
<emphasis id="D8D5EA80C363C73DE12E7246FE67F9A7" bold="true" box="[264,477,1623,1647]" pageId="77" pageNumber="78">FIGURES 311­320</emphasis>
. Genitalia of
<taxonomicName id="2DA14D11C363C73DE2537249FC4BF9A7" authority="Gusarov" authorityName="Gusarov" box="[629,1009,1624,1647]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="impressicollis" status="sp. nov.">
<emphasis id="D8D5EA80C363C73DE2537249FC2AF9A7" box="[629,912,1624,1647]" italics="true" pageId="77" pageNumber="78">Seeversiella impressicollis</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="D8D5EA80C363C73DE3DA7246FB82F9A7" bold="true" box="[1020,1080,1623,1647]" pageId="77" pageNumber="78">
<taxonomicNameLabel id="C3E657FBC363C73DE3DA7246FB82F9A7" box="[1020,1080,1623,1647]" pageId="77" pageNumber="78" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312­313, 315, 317, 319­ 320).
</paragraph>
</caption>
<paragraph id="EA1E3692C363C73DE12E735DFA91F8AB" blockId="77.[264,1323,1623,1891]" box="[264,1323,1868,1891]" pageId="77" pageNumber="78">CP – copulatory piece; ML – medial lamellae; SLD – sclerite of lateral diverticulum of internal sac.</paragraph>
<paragraph id="EA1E3692C360C73EE11E750DFE3BFE1E" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
<taxonomicName id="2DA14D11C360C73EE11E750DFDC9FEFD" box="[312,627,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C360C73EE11E750DFDC9FEFD" box="[312,627,284,309]" italics="true" pageId="78" pageNumber="79">Seeversiella impressicollis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2DA14D11C360C73EE33D750DFC55FEFD" box="[795,1007,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="D8D5EA80C360C73EE33D750DFC55FEFD" box="[795,1007,284,309]" italics="true" pageId="78" pageNumber="79">S. microphthalma</emphasis>
</taxonomicName>
in having pronotum with more asperate punctation; in males pronotum with broad medial impression, posterior angles of tergum 3 not projecting as spines; lateral diverticula of internal sac with long spiniform sclerites (
<figureCitation id="729A2A17C360C73EE1DE7585FD11FE66" box="[504,683,404,430]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Figs. 316, 318</figureCitation>
; 343); spermatheca with smaller distal portion (
<figureCitation id="729A2A17C360C73EE4D77585FE8DFE1E" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Figs. 320</figureCitation>
; 341).
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C360C73EE11E75F2FDC4FA5E" pageId="78" pageNumber="79" type="description">
<paragraph id="EA1E3692C360C73EE11E75F2FD37FDEE" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE11E75F2FE6AFE35" bold="true" box="[312,464,483,509]" pageId="78" pageNumber="79">Description.</emphasis>
Length
<quantity id="2D599B77C360C73EE21A75F5FD73FE36" box="[572,713,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="78" pageNumber="79" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2­2.7 mm</quantity>
. Body brown or dark brown with lighter pronotum and elytra; legs brownish yellow.
</paragraph>
<paragraph id="EA1E3692C360C73EE11E7625FCB1FD0E" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">Head surface glossy, with fine and weak isodiametric microsculpture, with fine and poorly visible punctation, distance between punctures equals 2 times their diameter. Temples 1.8­2.2 times as long as eyes. Antennal article 2 longer than 3, article 4 subquadrate, 5­10 transverse (ratio 1.5­1.8).</paragraph>
<paragraph id="EA1E3692C360C73EE11E76C5FE5FFBCE" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
Pronotum slightly transverse, 1.2 times as wide as head, width
<quantity id="2D599B77C360C73EE40D76C5FB6CFD26" box="[1067,1238,724,750]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.3" metricValueMin="4.699999999999999" pageId="78" pageNumber="79" unit="mm" value="0.5" valueMax="0.53" valueMin="0.47">0.47­0.53 mm</quantity>
, length
<quantity id="2D599B77C360C73EE12E76EDFE0BFCDE" box="[264,433,764,790]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="4.9" metricValueMin="4.3" pageId="78" pageNumber="79" unit="mm" value="0.45999999999999996" valueMax="0.49" valueMin="0.43">0.43­0.49 mm</quantity>
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine and slightly asperate punctation, distance between punctures equals 1­2 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.5), 1.8 times wider than long, glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 1­2 times their diameter. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="EA1E3692C360C73EE11E7005FC18FB6E" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
Abdominal terga glossy, with fine microsculpture consisting of transverse meshes, with fine punctation, distance between punctures equals 1­4 times their diameter on terga 3­5 and 3­5 times on tergum 7. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
<figureCitation id="729A2A17C360C73EE2C1709DFC2FFB6E" box="[743,917,1164,1190]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 – male tergum 8; 308 – male sternum 8; 309 – female tergum 8; 310 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="78" pageNumber="79">Figs. 307, 309</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E70A5FD9CFAD6" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
In males pronotum with wide medial impression; posterior angles of tergum 3 not projecting; tergum 7 without medial carina or tubercle; posterior margin of tergum 8 without emargination (
<figureCitation id="729A2A17C360C73EE1947115FDA0FAD6" box="[434,538,1284,1310]" captionStart="FIGURES 307 ­ 310" captionStartId="76.[264,383,1792,1816]" captionTargetBox="[290,1292,893,1759]" captionTargetId="figure@76.[282,1305,884,1766]" captionTargetPageId="76" captionText="FIGURES 307 ­ 310. Abdominal segment 8 of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 307 – male tergum 8; 308 – male sternum 8; 309 – female tergum 8; 310 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156474/files/figure.png" pageId="78" pageNumber="79">Fig. 307</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E713DFD48FAA6" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
Aedeagus as in
<figureCitation id="729A2A17C360C73EE1C9713DFD2EFA8E" box="[495,660,1324,1350]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Figs. 311­319</figureCitation>
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
<figureCitation id="729A2A17C360C73EE2587145FD5FFAA6" box="[638,741,1364,1390]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Fig. 313</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E716DFDC4FA5E" blockId="78.[264,1324,284,1550]" box="[312,638,1404,1430]" pageId="78" pageNumber="79">
Spermatheca as in
<figureCitation id="729A2A17C360C73EE232716DFDC0FA5E" box="[532,634,1404,1430]" captionStart="FIGURES 311 ­ 320" captionStartId="77.[264,383,1623,1647]" captionTargetBox="[286,1302,571,1605]" captionTargetId="figure@77.[276,1310,569,1613]" captionTargetPageId="77" captionText="FIGURES 311 ­ 320. Genitalia of Seeversiella impressicollis Gusarov, sp. n. (paratypes from Cerro Chirripó, Costa Rica). 311 – median lobe, parameral view; 312 – apex of median lobe, parameral view; 313 – apex of left paramere; 314 – median lobe, lateral view; 315 – apex of median lobe, lateral view; 316 – details of internal sac retracted into median lobe, lateral view; 317 – copulatory piece, lateral view; 318 – everted internal sac, parameral view; 319 – medial lamellae, parameral view; 320 ­ spermatheca. Scale bar 0.2 mm (311, 314, 316, 318), 0.1 mm (312 ­ 313, 315, 317, 319 ­ 320)." httpUri="https://zenodo.org/record/156475/files/figure.png" pageId="78" pageNumber="79">Fig. 320</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2BB6519C360C73EE11E71B2FE5DF9C6" pageId="78" pageNumber="79" type="distribution">
<paragraph id="EA1E3692C360C73EE11E71B2FBFEFA76" blockId="78.[264,1324,284,1550]" box="[312,1092,1443,1470]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE11E71B2FE62FA75" bold="true" box="[312,472,1443,1469]" pageId="78" pageNumber="79">Distribution.</emphasis>
Known from Cerro Chirripó,
<collectingCountry id="92B67602C360C73EE36671B5FC7AFA76" box="[832,960,1444,1470]" name="Costa Rica" pageId="78" pageNumber="79">Costa Rica</collectingCountry>
(
<figureCitation id="729A2A17C360C73EE3F671B5FB82FA76" box="[976,1080,1444,1470]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="78" pageNumber="79">Fig. 389</figureCitation>
).
</paragraph>
<paragraph id="EA1E3692C360C73EE11E71DAFE5DF9C6" blockId="78.[264,1324,284,1550]" pageId="78" pageNumber="79">
<emphasis id="D8D5EA80C360C73EE11E71DAFDBDFA2D" bold="true" box="[312,519,1483,1509]" pageId="78" pageNumber="79">Natural History.</emphasis>
<taxonomicName id="2DA14D11C360C73EE23C71DDFCEEFA2D" box="[538,852,1484,1509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="impressicollis">
<emphasis id="D8D5EA80C360C73EE23C71DDFCEEFA2D" box="[538,852,1484,1509]" italics="true" pageId="78" pageNumber="79">Seeversiella impressicollis</emphasis>
</taxonomicName>
was collected in paramo shrub litter at altitude of
<quantity id="2D599B77C360C73EE1A171E5FE5EF9C6" box="[391,484,1524,1550]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="78" pageNumber="79" unit="m" value="3600.0">3600 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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