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Key to species of
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1. Elytra uniformly coloured (
<figureCitation id="1300F98AFFF5FFF11290F8CDFDADAD6D" box="[494,608,1833,1855]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="9" pageNumber="393">Figs 14, 17</figureCitation>
,
<figureCitation id="1300F98AFFF5FFF11115F8CEFD67AD6D" box="[619,682,1834,1855]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="9" pageNumber="393">21, 24</figureCitation>
) or at most with pale-brown humeral spot (
<figureCitation id="1300F98AFFF5FFF11729F8CDFB6CAD6D" box="[1111,1185,1833,1855]" captionStart="FIGURE 2832" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 2832. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="9" pageNumber="393">Fig. 28</figureCitation>
). Pronotum parallel-sided (
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). Space between eyes relatively broad, HW / FW &lt;4................................................. 2
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- Elytra bicoloured with anterior half black and posterior half pale brown. (
<figureCitation id="1300F98AFFF5FFF110E7F880FC2DAD2B" box="[921,992,1892,1914]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="9" pageNumber="393">Fig. 34</figureCitation>
). Sides of pronotum not parallel, widest before middle (
<figureCitation id="1300F98AFFF5FFF1124CF865FEA0ADC5" box="[306,365,1921,1943]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="9" pageNumber="393">Fig. 9</figureCitation>
). Space between eyes very narrow, HW / FW&gt; 4. Bituberculate foveae lacking in apical third of elytra.........................................................................................
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2. Elytra pale brown, pronotum and head dark brown. Larger species with body length greater than
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. Inner interstriae completely smooth, without any tubercles. Bituberculate foveae lacking in apical fourth of elytra. Space between foveae larger than their diameter..................................................................
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B.
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- Whole body black. Smaller species with body length less than
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. All interstriae slightly but distinctly ungulated and with tubercles. Bituberculate foveae present on almost whole elytral surface. Space between foveae smaller than their diameter.................................................................................
<taxonomicName id="4C3B9E8CFFF6FFF217EAFF14FA51AB56" authority="Menier, 1979" authorityName="Menier" authorityYear="1979" box="[1172,1436,239,261]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="394" phylum="Arthropoda" rank="species" species="peyrierasi">
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<emphasis id="B94F391DFFF6FFF217EAFF14FADCAB56" bold="true" box="[1172,1297,239,261]" italics="true" pageId="10" pageNumber="394">B. peyrierasi</emphasis>
<bibRefCitation id="EFAA98FEFFF6FFF21666FF0BFA51AB56" author="Menier, J. J." box="[1304,1436,239,261]" pageId="10" pageNumber="394" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier, 1979</bibRefCitation>
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<mods:identifier id="59BEFCB32A49395CDD1ED65C372171FB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4A0F4AA587506D0151727D874E64137D" type="Zenodo-Dep">13915935</mods:identifier>
<mods:identifier id="B61EC32DA6F00AB513DC7B66302F3FCE" type="ZooBank">988A65E0-4B68-4E8F-B52C-112FEAE8F06C</mods:identifier>
</mods:mods>
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<subSubSection id="C321B684FFF9FFFD13E9FABDFDD2AF26" box="[151,543,1369,1396]" pageId="5" pageNumber="389" type="nomenclature">
<paragraph id="8B84E50FFFF9FFFD13E9FABDFDD2AF26" blockId="5.[151,543,1369,1432]" box="[151,543,1369,1396]" pageId="5" pageNumber="389">
<heading id="D0CC5263FFF9FFFD13E9FABDFDD2AF26" bold="true" box="[151,543,1369,1396]" fontSize="11" level="1" pageId="5" pageNumber="389" reason="1">
<emphasis id="B94F391DFFF9FFFD13E9FABDFDD2AF26" bold="true" box="[151,543,1369,1396]" pageId="5" pageNumber="389">
<taxonomicName id="4C3B9E8CFFF9FFFD13E9FABDFE77AF26" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[151,442,1369,1396]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF9FFFD13E9FABDFE77AF26" bold="true" box="[151,442,1369,1396]" italics="true" pageId="5" pageNumber="389">Biflabelotillus bicolorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C8466FFF9FFFD12BFFABEFDD2AF26" box="[449,543,1370,1396]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFFD13E9FA9AFE5EAFCA" box="[151,403,1406,1432]" pageId="5" pageNumber="389" type="description">
<paragraph id="8B84E50FFFF9FFFD13E9FA9AFE5EAFCA" blockId="5.[151,543,1369,1432]" box="[151,403,1406,1432]" pageId="5" pageNumber="389">
<figureCitation id="1300F98AFFF9FFFD13E9FA9AFE8DAFCA" box="[151,320,1406,1432]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="5" pageNumber="389">Figs 3, 6, 9, 12</figureCitation>
, 3337
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFFD13E9FA22FB8BACEA" pageId="5" pageNumber="389" type="materials_examined">
<paragraph id="8B84E50FFFF9FFFD13E9FA22FF20AC56" blockId="5.[151,1437,1477,2008]" pageId="5" pageNumber="389">
<materialsCitation id="3B53EF52FFF9FFFD13E9FA22FF24AC56" collectionCode="RN" country="Madagascar" location="Moramanga" pageId="5" pageNumber="389" specimenCode="RN 2" specimenCount="1" stateProvince="Antananarivo" typeStatus="holotype">
<emphasis id="B94F391DFFF9FFFD13E9FA22FEFAAF8D" bold="true" box="[151,311,1477,1504]" pageId="5" pageNumber="389">
<typeStatus id="54805BADFFF9FFFD13E9FA22FF1FAFB2" box="[151,210,1478,1504]" pageId="5" pageNumber="389">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32CA59FFFF9FFFD1241FA22FE05AFB2" box="[319,456,1478,1504]" name="Madagascar" pageId="5" pageNumber="389">Madagascar</collectingCountry>
,
<quantity id="4CC348EAFFF9FFFD12AAFA22FDD1AFB2" box="[468,540,1478,1505]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="389" unit="km" value="50.0">50 km</quantity>
W of
<collectingRegion id="49FF2BEDFFF9FFFD111AFA22FD32AFB2" box="[612,767,1478,1504]" country="Madagascar" name="Antananarivo" pageId="5" pageNumber="389">Antananarivo</collectingRegion>
,
<quantity id="4CC348EAFFF9FFFD1072FA22FC99AFB2" box="[780,852,1478,1504]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="389" unit="km" value="20.0">20 km</quantity>
E-NE of
<location id="8EE4B3D4FFF9FFFD10BAFA22FB83AFB2" LSID="urn:lsid:plazi:treatment:03925419FFF9FFF113E9FABDFAAFACC1:8EE4B3D4FFF9FFFD10BAFA22FB83AFB2" box="[964,1102,1478,1504]" country="Madagascar" name="Moramanga" pageId="5" pageNumber="389" stateProvince="Antananarivo">Moramanga</location>
,
<locationDeviation id="9A36FC2DFFF9FFFD1724FA22FADAAFB2" box="[1114,1303,1478,1505]" location="Route" pageId="5" pageNumber="389">
<quantity id="4CC348EAFFF9FFFD1724FA22FB58AFB2" box="[1114,1173,1478,1505]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="389" unit="km" value="5.0">5 km</quantity>
S of Route
</locationDeviation>
nationale 2 (
<specimenCode id="DB9D4D74FFF9FFFD13E1FA0EFF12AC56" box="[159,223,1514,1540]" collectionCode="RN" pageId="5" pageNumber="389">RN 2</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B84E50FFFF9FFFD13B9F9EAFE62ACC6" blockId="5.[151,1437,1477,2008]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9F9EAFEB0AC75" bold="true" box="[199,381,1549,1576]" pageId="5" pageNumber="389">Type material.</emphasis>
<materialsCitation id="3B53EF52FFF9FFFD12F5F9EAFDF3AC22" collectionCode="DATE" country="Madagascar" location="Route" pageId="5" pageNumber="389" specimenCode="DATE 1-70" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54805BADFFF9FFFD12F5F9EAFE3AAC7A" box="[395,503,1550,1576]" pageId="5" pageNumber="389" type="holotype">Holotype</typeStatus>
. (
<specimenCount id="9D3D2E86FFF9FFFD116AF9EAFD89AC7A" box="[532,580,1550,1576]" count="1" pageId="5" pageNumber="389" type="male">1 ♂</specimenCount>
FPC): [white, printed label]
<collectingCountry id="F32CA59FFFF9FFFD10D3F9EAFBB9AC7A" box="[941,1140,1550,1576]" name="Madagascar" pageId="5" pageNumber="389">MADAGASCAR</collectingCountry>
EST // [white, printed + handwritten label]
<location id="8EE4B3D4FFF9FFFD1214F9D6FE60AC1E" LSID="urn:lsid:plazi:treatment:03925419FFF9FFF113E9FABDFAAFACC1:8EE4B3D4FFF9FFFD1214F9D6FE60AC1E" box="[362,429,1586,1612]" country="Madagascar" name="Route" pageId="5" pageNumber="389">Route</location>
ANOSIBE / P.K. 40 /
<specimenCode id="DB9D4D74FFF9FFFD11D6F9D6FCE3AC1E" box="[680,814,1586,1612]" collectionCode="DATE" pageId="5" pageNumber="389">DATE 170</specimenCode>
// [white, printed label] DUJARDIN-DELACOUR G. / ENTOMOLOGIE/ TANANARIVE //
</materialsCitation>
<typeStatus id="54805BADFFF9FFFD1138F9B2FD21AC22" box="[582,748,1622,1648]" pageId="5" pageNumber="389" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="4C3B9E8CFFF9FFFD118AF9B2FBC9AC22" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[756,1028,1622,1648]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF9FFFD118AF9B2FBC9AC22" box="[756,1028,1622,1648]" italics="true" pageId="5" pageNumber="389">Biflabelotillus bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFF9FFFD1775F9B2FBABAC22" bold="true" box="[1035,1126,1622,1648]" pageId="5" pageNumber="389">
<taxonomicNameLabel id="A27C8466FFF9FFFD1775F9B2FBABAC22" box="[1035,1126,1622,1648]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
det. F. Pavel &amp; L. Purchart 2024 [red, printed label].
</paragraph>
<paragraph id="8B84E50FFFF9FFFD13B9F979FB8BACEA" blockId="5.[151,1437,1477,2008]" box="[199,1094,1693,1720]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9F979FE52ACEA" bold="true" box="[199,415,1693,1720]" pageId="5" pageNumber="389">
Condition of
<typeStatus id="54805BADFFF9FFFD121AF97AFE54ACEA" box="[356,409,1694,1720]" pageId="5" pageNumber="389">type</typeStatus>
.
</emphasis>
Right front leg missing. Right antennomeres IV-XI missing.
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFF013B9F925FD21AA87" lastPageId="8" lastPageNumber="392" pageId="5" pageNumber="389" type="diagnosis">
<paragraph id="8B84E50FFFF9FFF013B9F925FD21AA87" blockId="5.[151,1437,1477,2008]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="392" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9F925FE1CAC89" bold="true" box="[199,465,1729,1755]" pageId="5" pageNumber="389">Differential diagnosis.</emphasis>
Both
<taxonomicName id="4C3B9E8CFFF9FFFD1162F927FD64AC8E" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[540,681,1730,1756]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF9FFFD1162F927FD64AC8E" box="[540,681,1730,1756]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFF9FFFD11CDF926FCDDAC8E" bold="true" box="[691,784,1730,1756]" pageId="5" pageNumber="389">
<taxonomicNameLabel id="A27C8466FFF9FFFD11CDF926FCDDAC8E" box="[691,784,1730,1756]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3B9E8CFFF9FFFD1033F927FC2EAC8E" authorityName="Pic" authorityYear="1949" box="[845,995,1730,1756]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFF9FFFD1033F927FC2EAC8E" box="[845,995,1730,1756]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
</taxonomicName>
can be distinguished reliably from
<taxonomicName id="4C3B9E8CFFF9FFFD16FAF927FEC4AD52" authorityName="Menier" authorityYear="1979" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF9FFFD16FAF927FEC4AD52" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
</taxonomicName>
mainly by elytral surface, as they have smooth elytral interstriae (tubercles present in elytral interstriae in
<taxonomicName id="4C3B9E8CFFF9FFFD13C9F8EFFE87AD76" authorityName="Menier" authorityYear="1979" box="[183,330,1803,1828]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF9FFFD13C9F8EFFE87AD76" box="[183,330,1803,1828]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
</taxonomicName>
), by bituberculate foveae which are lacking in apical fourth in
<taxonomicName id="4C3B9E8CFFF9FFFD1757F8EFFB73AD76" authorityName="Pic" authorityYear="1949" box="[1065,1214,1802,1828]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFF9FFFD1757F8EFFB73AD76" box="[1065,1214,1802,1828]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
</taxonomicName>
and apical third of elytra in
<taxonomicName id="4C3B9E8CFFF9FFFD1385F8CBFE48AD1A" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[251,389,1838,1864]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus">
<emphasis id="B94F391DFFF9FFFD1385F8CBFE48AD1A" box="[251,389,1838,1864]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
</taxonomicName>
, respectively (foveae cover whole surface in
<taxonomicName id="4C3B9E8CFFF9FFFD10FDF8CBFBD8AD1A" authorityName="Menier" authorityYear="1979" box="[899,1045,1839,1864]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF9FFFD10FDF8CBFBD8AD1A" box="[899,1045,1839,1864]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
</taxonomicName>
), and space between foveae which is equal or larger than their diameter (smaller in
<taxonomicName id="4C3B9E8CFFF9FFFD11C9F8B7FC84AD3E" authorityName="Menier" authorityYear="1979" box="[695,841,1875,1900]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF9FFFD11C9F8B7FC84AD3E" box="[695,841,1875,1900]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
</taxonomicName>
). Last but not least, ultimate antennomere is more or less securiform in
<taxonomicName id="4C3B9E8CFFF9FFFD121FF893FE24ADC2" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[353,489,1910,1936]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF9FFFD121FF893FE24ADC2" box="[353,489,1910,1936]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFF9FFFD1291F892FD8AADC2" bold="true" box="[495,583,1910,1936]" pageId="5" pageNumber="389">
<taxonomicNameLabel id="A27C8466FFF9FFFD1291F892FD8AADC2" box="[495,583,1910,1936]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3B9E8CFFF9FFFD1105F893FCC1ADC2" authorityName="Pic" authorityYear="1949" box="[635,780,1910,1936]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFF9FFFD1105F893FCC1ADC2" box="[635,780,1910,1936]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
</taxonomicName>
(
<figureCitation id="1300F98AFFF9FFFD1067F892FCB6ADC2" box="[793,891,1910,1936]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="5" pageNumber="389">Figs 4, 6</figureCitation>
) whereas it is elongate-oval in
<taxonomicName id="4C3B9E8CFFF9FFFD17AFF893FAACADC2" authorityName="Menier" authorityYear="1979" box="[1233,1377,1911,1936]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF9FFFD17AFF893FAACADC2" box="[1233,1377,1911,1936]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
</taxonomicName>
(
<figureCitation id="1300F98AFFF9FFFD1611F892FF6AADE7" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="5" pageNumber="389">Fig. 5</figureCitation>
). The new species most resembles
<taxonomicName id="4C3B9E8CFFF9FFFD114FF87FFD0EADE6" authorityName="Pic" authorityYear="1949" box="[561,707,1946,1972]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFF9FFFD114FF87FFD0EADE6" box="[561,707,1946,1972]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
</taxonomicName>
but differs in following characteristics:
<taxonomicName id="4C3B9E8CFFF9FFFD17FBF87FFAC3ADE6" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1157,1294,1946,1972]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF9FFFD17FBF87FFAC3ADE6" box="[1157,1294,1946,1972]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFF9FFFD166BF87EFAA2ADE6" bold="true" box="[1301,1391,1946,1972]" pageId="5" pageNumber="389">
<taxonomicNameLabel id="A27C8466FFF9FFFD166BF87EFAA2ADE6" box="[1301,1391,1946,1972]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a smaller species with bicoloured elytra and non-parallel sided pronotum, the latter widest before middle (
<figureCitation id="1300F98AFFF9FFFD165BF85AFAA0AD8A" box="[1317,1389,1982,2008]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="5" pageNumber="389">Figs 9</figureCitation>
, 33, 34) whereas
<taxonomicName id="4C3B9E8CFFFBFFFF1257F925FE70AC88" authorityName="Pic" authorityYear="1949" box="[297,445,1728,1754]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFBFFFF1257F925FE70AC88" box="[297,445,1728,1754]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
</taxonomicName>
is a larger species with uniformly coloured elytra and parallel sided pronotum (
<figureCitation id="1300F98AFFFBFFFF1633F924FA5AAC89" box="[1357,1431,1728,1755]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="7" pageNumber="391">Figs 7</figureCitation>
, 1417). Additionally, these species can be distinguished as follows: in
<taxonomicName id="4C3B9E8CFFFBFFFF10B2F901FB95ACAC" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[972,1112,1764,1790]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFBFFFF10B2F901FB95ACAC" box="[972,1112,1764,1790]" italics="true" pageId="7" pageNumber="391">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFBFFFF171FF900FB73ACAC" bold="true" box="[1121,1214,1764,1790]" pageId="7" pageNumber="391">
<taxonomicNameLabel id="A27C8466FFFBFFFF171FF900FB73ACAC" box="[1121,1214,1764,1790]" pageId="7" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the eyes are larger and less emarginated anteriorly, the space between eyes (vertex) very narrow with HW / FW = 4.26 (
<figureCitation id="1300F98AFFFBFFFF1666F8ECFA43AD70" box="[1304,1422,1800,1826]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="7" pageNumber="391">Figs 9, 12</figureCitation>
), whereas in
<taxonomicName id="4C3B9E8CFFFBFFFF1262F8C9FE7DAD14" authorityName="Pic" authorityYear="1949" box="[284,432,1836,1862]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFBFFFF1262F8C9FE7DAD14" box="[284,432,1836,1862]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
</taxonomicName>
the eyes are smaller, more emarginated anteriorly and the space between eyes is much wider with HW / FW = 2.993.31 (
<figureCitation id="1300F98AFFFBFFFF115DF8B4FD5AAD38" box="[547,663,1872,1899]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="7" pageNumber="391">Figs 7, 10</figureCitation>
). The terminal antennomere of
<taxonomicName id="4C3B9E8CFFFBFFFF1087F8B5FB4FAD38" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1017,1154,1872,1898]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFBFFFF1087F8B5FB4FAD38" box="[1017,1154,1872,1898]" italics="true" pageId="7" pageNumber="391">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFBFFFF17F7F8B4FB2EAD38" bold="true" box="[1161,1251,1872,1898]" pageId="7" pageNumber="391">
<taxonomicNameLabel id="A27C8466FFFBFFFF17F7F8B4FB2EAD38" box="[1161,1251,1872,1898]" pageId="7" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is broadened on one side (2.75x longer than broad) and somewhat resembling the shape of a securiform shape of a palpus (
<figureCitation id="1300F98AFFFBFFFF1637F890FA43ADDC" box="[1353,1422,1908,1934]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="7" pageNumber="391">Fig. 6</figureCitation>
), whereas in
<taxonomicName id="4C3B9E8CFFFBFFFF126BF87DFE6BADE0" authorityName="Pic" authorityYear="1949" box="[277,422,1944,1970]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFBFFFF126BF87DFE6BADE0" box="[277,422,1944,1970]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
</taxonomicName>
it is more elongate (3.25x longer than broad) and not securiform in shape (
<figureCitation id="1300F98AFFFBFFFF179CF87CFAEAADE0" box="[1250,1319,1944,1970]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="7" pageNumber="391">Fig. 4</figureCitation>
). Shape of antennomeres III-V is different, more tooth-like widened in
<taxonomicName id="4C3B9E8CFFFBFFFF104EF859FC0CAD84" authorityName="Pic" authorityYear="1949" box="[816,961,1980,2006]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFBFFFF104EF859FC0CAD84" box="[816,961,1980,2006]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
</taxonomicName>
(
<figureCitation id="1300F98AFFFBFFFF10B1F858FBFCAD84" box="[975,1073,1980,2006]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="7" pageNumber="391">Figs 1, 3</figureCitation>
). Scutellum almost squared with slightly rounded sides in
<taxonomicName id="4C3B9E8CFFFBFFFF12CDF805FDF0ADA8" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[435,573,2016,2042]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFBFFFF12CDF805FDF0ADA8" box="[435,573,2016,2042]" italics="true" pageId="7" pageNumber="391">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFBFFFF113BF804FD52ADA8" bold="true" box="[581,671,2016,2042]" pageId="7" pageNumber="391">
<taxonomicNameLabel id="A27C8466FFFBFFFF113BF804FD52ADA8" box="[581,671,2016,2042]" pageId="7" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
whereas it is almost rounded in
<taxonomicName id="4C3B9E8CFFFBFFFF1770F805FB6CADA8" authorityName="Pic" authorityYear="1949" box="[1038,1185,2016,2042]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFBFFFF1770F805FB6CADA8" box="[1038,1185,2016,2042]" italics="true" pageId="7" pageNumber="391">B. foveolatus</emphasis>
</taxonomicName>
with the basal margin distinctly cut-out in middle (
<figureCitation id="1300F98AFFF4FFF012A4FF73FDFCAAE3" box="[474,561,151,177]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="8" pageNumber="392">Figs 19</figureCitation>
,
<figureCitation id="1300F98AFFF4FFF01140FF73FD93AAE3" box="[574,606,151,177]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="8" pageNumber="392">36</figureCitation>
). Prosternal inter-coxal process narrower in
<taxonomicName id="4C3B9E8CFFF4FFF0172DFF73FB10AAE3" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1107,1245,151,177]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="392" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF4FFF0172DFF73FB10AAE3" box="[1107,1245,151,177]" italics="true" pageId="8" pageNumber="392">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFF4FFF0179AFF73FAF2AAE3" bold="true" box="[1252,1343,151,177]" pageId="8" pageNumber="392">
<taxonomicNameLabel id="A27C8466FFF4FFF0179AFF73FAF2AAE3" box="[1252,1343,151,177]" pageId="8" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(WPC / WPP = 3.25) than in
<taxonomicName id="4C3B9E8CFFF4FFF012FDFF5FFDD8AA87" authorityName="Pic" authorityYear="1949" box="[387,533,187,213]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="392" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFF4FFF012FDFF5FFDD8AA87" box="[387,533,187,213]" italics="true" pageId="8" pageNumber="392">B. foveolatus</emphasis>
</taxonomicName>
(WPC / WPP = 5).
</paragraph>
</subSubSection>
<caption id="DF44B587FFFAFFFE13E9F873FD58ADA5" ID-DOI="http://doi.org/10.5281/zenodo.13915941" ID-Zenodo-Dep="13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="6" pageNumber="390" startId="6.[151,250,1943,1967]" targetBox="[225,1371,187,1919]" targetPageId="6" targetType="figure">
<paragraph id="8B84E50FFFFAFFFE13E9F873FD58ADA5" blockId="6.[151,1436,1943,2039]" pageId="6" pageNumber="390">
<emphasis id="B94F391DFFFAFFFE13E9F873FE85ADFD" bold="true" box="[151,328,1943,1967]" pageId="6" pageNumber="390">FIGURE 2127.</emphasis>
<taxonomicName id="4C3B9E8CFFFAFFFE122EF873FD12ADFD" authority="Menier, 1979" authorityName="Menier" authorityYear="1979" box="[336,735,1943,1967]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="390" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFAFFFE122EF873FD82ADFD" box="[336,591,1943,1967]" italics="true" pageId="6" pageNumber="390">Biflabelotillus peyrierasi</emphasis>
<bibRefCitation id="EFAA98FEFFFAFFFE1129F873FD12ADFD" author="Menier, J. J." box="[599,735,1943,1967]" pageId="6" pageNumber="390" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier, 1979</bibRefCitation>
</taxonomicName>
—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea).
</paragraph>
</caption>
<caption id="DF44B587FFFBFFFF13E9F9D6FE20ACC0" ID-DOI="http://doi.org/10.5281/zenodo.13915943" ID-Zenodo-Dep="13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="7" pageNumber="391" startId="7.[151,250,1586,1610]" targetBox="[170,1418,184,1562]" targetPageId="7" targetType="figure">
<paragraph id="8B84E50FFFFBFFFF13E9F9D6FE20ACC0" blockId="7.[151,1436,1586,1682]" pageId="7" pageNumber="391">
<emphasis id="B94F391DFFFBFFFF13E9F9D6FE84AC18" bold="true" box="[151,329,1586,1610]" pageId="7" pageNumber="391">FIGURE 2832.</emphasis>
<taxonomicName id="4C3B9E8CFFFBFFFF122FF9D6FD28AC18" authority="Menier, 1979" authorityName="Menier" authorityYear="1979" box="[337,741,1586,1610]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="391" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFBFFFF122FF9D6FD9DAC18" box="[337,592,1586,1610]" italics="true" pageId="7" pageNumber="391">Biflabelotillus peyrierasi</emphasis>
<bibRefCitation id="EFAA98FEFFFBFFFF1127F9D6FD28AC18" author="Menier, J. J." box="[601,741,1586,1610]" pageId="7" pageNumber="391" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier, 1979</bibRefCitation>
</taxonomicName>
, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea).
</paragraph>
</caption>
<caption id="DF44B587FFF4FFF013E9F96BFF12ACBD" ID-DOI="http://doi.org/10.5281/zenodo.13915945" ID-Zenodo-Dep="13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="8" pageNumber="392" startId="8.[151,250,1679,1703]" targetBox="[173,1409,286,1655]" targetPageId="8" targetType="figure">
<paragraph id="8B84E50FFFF4FFF013E9F96BFF12ACBD" blockId="8.[151,1436,1679,1775]" pageId="8" pageNumber="392">
<emphasis id="B94F391DFFF4FFF013E9F96BFE85ACF5" bold="true" box="[151,328,1679,1703]" pageId="8" pageNumber="392">FIGURE 3337.</emphasis>
<taxonomicName id="4C3B9E8CFFF4FFF01231F96BFD8AACF5" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[335,583,1679,1703]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="392" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF4FFF01231F96BFD8AACF5" box="[335,583,1679,1703]" italics="true" pageId="8" pageNumber="392">Biflabelotillus bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFF4FFF01131F96BFD6CACF5" bold="true" box="[591,673,1679,1703]" pageId="8" pageNumber="392">
<taxonomicNameLabel id="A27C8466FFF4FFF01131F96BFD6CACF5" box="[591,673,1679,1703]" pageId="8" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea).
</paragraph>
</caption>
<subSubSection id="C321B684FFF4FFF113B9F8F9FBB3AC19" lastPageId="9" lastPageNumber="393" pageId="8" pageNumber="392" type="description">
<paragraph id="8B84E50FFFF4FFF013B9F8F9FE41AD09" blockId="8.[151,1437,1821,1991]" pageId="8" pageNumber="392">
<emphasis id="B94F391DFFF4FFF013B9F8F9FE94AD65" bold="true" box="[199,345,1821,1847]" pageId="8" pageNumber="392">Description.</emphasis>
Size of
<typeStatus id="54805BADFFF4FFF012CBF8F9FDDAAD65" box="[437,535,1821,1847]" pageId="8" pageNumber="392" type="holotype">holotype</typeStatus>
9.0 x
<quantity id="4CC348EAFFF4FFF01123F8F9FD7BAD6A" box="[605,694,1821,1848]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="392" unit="mm" value="2.5">2.5 mm</quantity>
. Winged, black, with apical half of elytra and tarsi rust-brown and abdomen pale-brown.
</paragraph>
<paragraph id="8B84E50FFFF4FFF013B9F881FE0DAD95" blockId="8.[151,1437,1821,1991]" pageId="8" pageNumber="392">
<emphasis id="B94F391DFFF4FFF013B9F881FEC3AD2D" bold="true" box="[199,270,1893,1919]" pageId="8" pageNumber="392">Head.</emphasis>
Head including eyes broader than pronotum (DV) in ratio 1:0.7 (HW =
<quantity id="4CC348EAFFF4FFF01736F881FB7CAD2D" box="[1096,1201,1893,1919]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" pageId="8" pageNumber="392" unit="mm" value="2.13">2.13 mm</quantity>
). The space between eyes (frons) very narrow (HW / FW = 4.26) (
<figureCitation id="1300F98AFFF4FFF011F4F86DFD11ADF1" box="[650,732,1929,1955]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="8" pageNumber="392">Fig. 12</figureCitation>
). Antennae 1.47x longer than pronotum. Ultimate antennomere axe-like, widened (
<figureCitation id="1300F98AFFF4FFF01210F849FE7FAD95" box="[366,434,1965,1991]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="8" pageNumber="392">Fig. 6</figureCitation>
).
</paragraph>
<caption id="DF44B587FFF5FFF113E9FBBAFE60AEC8" ID-DOI="http://doi.org/10.5281/zenodo.13915947" ID-Zenodo-Dep="13915947" httpUri="https://zenodo.org/record/13915947/files/figure.png" pageId="9" pageNumber="393" startId="9.[151,250,1118,1142]" targetBox="[489,1099,181,1094]" targetPageId="9" targetType="figure">
<paragraph id="8B84E50FFFF5FFF113E9FBBAFE60AEC8" blockId="9.[151,1437,1118,1178]" pageId="9" pageNumber="393">
<emphasis id="B94F391DFFF5FFF113E9FBBAFED3AE24" bold="true" box="[151,286,1118,1142]" pageId="9" pageNumber="393">FIGURE 38.</emphasis>
Distribution map of the genus
<taxonomicName id="4C3B9E8CFFF5FFF1112EFBBAFD2DAE24" authorityName="Pic" authorityYear="1949" box="[592,736,1118,1142]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFF5FFF1112EFBBAFD2DAE24" box="[592,736,1118,1142]" italics="true" pageId="9" pageNumber="393">Biflabelotillus</emphasis>
</taxonomicName>
.
<emphasis id="B94F391DFFF5FFF1118EFBBBFC44AE24" box="[752,905,1118,1142]" italics="true" pageId="9" pageNumber="393">
<taxonomicName id="4C3B9E8CFFF5FFF1118EFBBBFCBDAE24" authorityName="Pic" authorityYear="1949" box="[752,880,1118,1142]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="species" species="foveolatus">B. foveolatus</taxonomicName>
</emphasis>
red triangle,
<taxonomicName id="4C3B9E8CFFF5FFF11779FBBBFB45AE24" authorityName="Menier" authorityYear="1979" box="[1031,1160,1119,1142]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF5FFF11779FBBBFB45AE24" box="[1031,1160,1119,1142]" italics="true" pageId="9" pageNumber="393">B. peyrierasi</emphasis>
</taxonomicName>
—white circle,
<taxonomicName id="4C3B9E8CFFF5FFF1165CFBBBFA50AE24" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1314,1437,1118,1142]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF5FFF1165CFBBBFA50AE24" box="[1314,1437,1118,1142]" italics="true" pageId="9" pageNumber="393">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFF5FFF113E9FB66FF25AEC8" bold="true" box="[151,232,1154,1178]" pageId="9" pageNumber="393">
<taxonomicNameLabel id="A27C8466FFF5FFF113E9FB66FF25AEC8" box="[151,232,1154,1178]" pageId="9" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
—yellow rhombus.
</paragraph>
</caption>
<paragraph id="8B84E50FFFF5FFF113B9FB2DFA4DAF1D" blockId="9.[151,1437,1224,1683]" pageId="9" pageNumber="393">
<emphasis id="B94F391DFFF5FFF113B9FB2DFE8AAEB1" bold="true" box="[199,327,1225,1251]" pageId="9" pageNumber="393">Pronotum.</emphasis>
Longer than wide in ratio 1:1.5 (PL =
<quantity id="4CC348EAFFF5FFF1106EFB2CFCB5AEB1" box="[784,888,1224,1251]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="9" pageNumber="393" unit="mm" value="2.25">2.25 mm</quantity>
, PW =
<quantity id="4CC348EAFFF5FFF110ADFB2CFBFDAEB1" box="[979,1072,1224,1251]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="393" unit="mm" value="1.5">1.5 mm</quantity>
), subcylindrical but not parallel sided, widest before middle, sides of pronotum narrowing in basal third from pronotal base toward the middle part of pronotum, where it is distinctly widening and then slightly narrowing toward anterior edge of pronotum, surface roughly wrinkled, irregularly in dorsal part, more or less longitudinally coarsely wrinkled in lateral part (
<figureCitation id="1300F98AFFF5FFF11653FAD0FABFAF1D" box="[1325,1394,1332,1359]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="9" pageNumber="393">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFF5FFF113B9FABDFE3DAC50" blockId="9.[151,1437,1224,1683]" pageId="9" pageNumber="393">
<emphasis id="B94F391DFFF5FFF113B9FABDFED7AF21" bold="true" box="[199,282,1369,1395]" pageId="9" pageNumber="393">Elytra.</emphasis>
Subparallel, 2.7x longer than their width (EL =
<quantity id="4CC348EAFFF5FFF1103FFABCFC65AF21" box="[833,936,1368,1395]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" pageId="9" pageNumber="393" unit="mm" value="6.75">6.75 mm</quantity>
, EW =
<quantity id="4CC348EAFFF5FFF1177EFABCFB96AF21" box="[1024,1115,1368,1395]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="393" unit="mm" value="2.5">2.5 mm</quantity>
), with posterior half slightly widening posteriad. Anterior half of elytra black, posterior half rust-brown (
<figureCitation id="1300F98AFFF5FFF11773FA98FBA8AFC5" box="[1037,1125,1404,1431]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="9" pageNumber="393">Fig. 34</figureCitation>
). Black part of elytra with bituberculate foveae arranged in regular rows and ending at interface of black and rust-brown area, whereas the latter is smooth with only several bituberculate foveae close to black area. Space between foveae approx. 13x larger than diameter of foveae.
</paragraph>
<paragraph id="8B84E50FFFF5FFF113B9F9E8FC49AC74" blockId="9.[151,1437,1224,1683]" box="[199,900,1548,1575]" pageId="9" pageNumber="393">
<emphasis id="B94F391DFFF5FFF113B9F9E8FE8BAC74" bold="true" box="[199,326,1548,1574]" pageId="9" pageNumber="393">Scutellum.</emphasis>
Almost square with slightly curved sides (
<figureCitation id="1300F98AFFF5FFF1105DF9E8FCB5AC74" box="[803,888,1548,1574]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="9" pageNumber="393">Fig. 36</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFF5FFF113B9F9D5FBB3AC19" blockId="9.[151,1437,1224,1683]" box="[199,1150,1584,1611]" pageId="9" pageNumber="393">
<emphasis id="B94F391DFFF5FFF113B9F9D5FEB7AC18" bold="true" box="[199,378,1584,1611]" pageId="9" pageNumber="393">Lower surface.</emphasis>
Ventrite I brown to pale-brown, ventrites II-VI pale brown (
<figureCitation id="1300F98AFFF5FFF11763F9D4FBBCAC19" box="[1053,1137,1584,1611]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="9" pageNumber="393">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF5FFF113B9F9B0FAEDAC3D" box="[199,1312,1620,1647]" pageId="9" pageNumber="393" type="distribution">
<paragraph id="8B84E50FFFF5FFF113B9F9B0FAEDAC3D" blockId="9.[151,1437,1224,1683]" box="[199,1312,1620,1647]" pageId="9" pageNumber="393">
<emphasis id="B94F391DFFF5FFF113B9F9B0FEACAC3C" bold="true" box="[199,353,1620,1646]" pageId="9" pageNumber="393">Distribution.</emphasis>
<collectingCountry id="F32CA59FFFF5FFF11219F9B0FE3DAC3C" box="[359,496,1620,1646]" name="Madagascar" pageId="9" pageNumber="393">Madagascar</collectingCountry>
, Moramanga district. Known only from the
<typeStatus id="54805BADFFF5FFF110A0F9B1FBC3AC3D" box="[990,1038,1621,1647]" pageId="9" pageNumber="393">type</typeStatus>
locality so far (
<figureCitation id="1300F98AFFF5FFF117C1F9B0FADEAC3D" box="[1215,1299,1620,1647]" captionStart="FIGURE 38" captionStartId="9.[151,250,1118,1142]" captionTargetBox="[489,1099,181,1094]" captionTargetId="figure-26@9.[489,1099,181,1094]" captionTargetPageId="9" captionText="FIGURE 38. Distribution map of the genus Biflabelotillus. B. foveolatus—red triangle, B. peyrierasi—white circle, B. bicolorus sp. nov.—yellow rhombus." figureDoi="http://doi.org/10.5281/zenodo.13915947" httpUri="https://zenodo.org/record/13915947/files/figure.png" pageId="9" pageNumber="393">Fig. 38</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF5FFF113B9F99DFAAFACC1" box="[199,1378,1656,1683]" pageId="9" pageNumber="393" type="etymology">
<paragraph id="8B84E50FFFF5FFF113B9F99DFAAFACC1" blockId="9.[151,1437,1224,1683]" box="[199,1378,1656,1683]" pageId="9" pageNumber="393">
<emphasis id="B94F391DFFF5FFF113B9F99DFE83ACC1" bold="true" box="[199,334,1657,1683]" pageId="9" pageNumber="393">Etymology.</emphasis>
Latin expression for having two colors or to be of two colors. Adjective
<taxonomicName id="4C3B9E8CFFF5FFF117ECF99CFB31ACC0" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1170,1276,1656,1682]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="393" phylum="Arthropoda" rank="species" species="bicolorus">
<emphasis id="B94F391DFFF5FFF117ECF99CFB31ACC0" box="[1170,1276,1656,1682]" italics="true" pageId="9" pageNumber="393">bicolorus</emphasis>
</taxonomicName>
, -
<emphasis id="B94F391DFFF5FFF1166AF99DFAEFACC0" box="[1300,1314,1657,1682]" italics="true" pageId="9" pageNumber="393">a</emphasis>
, -
<emphasis id="B94F391DFFF5FFF11647F99DFA96ACC0" box="[1337,1371,1657,1682]" italics="true" pageId="9" pageNumber="393">um</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,287 @@
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<mods:title id="F44B23F31F9437055AC4C3D1400378BD">Revision of Madagascan Cleridae (Coleoptera: Cleroidea). Part I. - The genus Biflabelotillus Pic, 1949 (Tillinae)</mods:title>
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<subSubSection id="C321B684FFF9FFFD13E9FF73FDA7AAE3" box="[151,618,151,178]" pageId="5" pageNumber="389" type="nomenclature">
<paragraph id="8B84E50FFFF9FFFD13E9FF73FDA7AAE3" blockId="5.[151,618,151,214]" box="[151,618,151,178]" pageId="5" pageNumber="389">
<heading id="D0CC5263FFF9FFFD13E9FF73FDA7AAE3" bold="true" box="[151,618,151,178]" fontSize="11" level="1" pageId="5" pageNumber="389" reason="1">
<taxonomicName id="4C3B9E8CFFF9FFFD13E9FF73FDA7AAE3" authority="Menier, 1979" authorityName="Menier" authorityYear="1979" box="[151,618,151,178]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF9FFFD13E9FF73FDA7AAE3" bold="true" box="[151,618,151,178]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13E9FF73FE0CAAE3" bold="true" box="[151,449,151,178]" italics="true" pageId="5" pageNumber="389">Biflabelotillus peyrierasi</emphasis>
<bibRefCitation id="EFAA98FEFFF9FFFD12B6FF73FDA7AAE3" author="Menier, J. J." box="[456,618,151,177]" pageId="5" pageNumber="389" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier, 1979</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFFD13E9FF5FFE5CAA87" box="[151,401,187,214]" pageId="5" pageNumber="389" type="description">
<paragraph id="8B84E50FFFF9FFFD13E9FF5FFE5CAA87" blockId="5.[151,618,151,214]" box="[151,401,187,214]" pageId="5" pageNumber="389">
<figureCitation id="1300F98AFFF9FFFD13E9FF5FFEF2AA87" box="[151,319,187,214]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="5" pageNumber="389">Figs 2, 5, 8, 11</figureCitation>
, 2132
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFFD13E9FEE4FCA2AB4A" box="[151,879,255,280]" pageId="5" pageNumber="389" type="reference_group">
<paragraph id="8B84E50FFFF9FFFD13E9FEE4FCA2AB4A" blockId="5.[151,879,255,280]" box="[151,879,255,280]" pageId="5" pageNumber="389">
<treatmentCitationGroup id="AB2BC221FFF9FFFD13E9FEE4FCA2AB4A" box="[151,879,255,280]" pageId="5" pageNumber="389">
<treatmentCitation id="0A9AC31EFFF9FFFD13E9FEE4FD46AB45" author="Menier, J. J." box="[151,651,255,280]" page="75" pageId="5" pageNumber="389" year="1979">
<taxonomicName id="4C3B9E8CFFF9FFFD13E9FEE4FD46AB45" authority="Menier, 1979: 75" authorityName="Menier" authorityPageNumber="75" authorityYear="1979" box="[151,651,255,280]" class="Insecta" family="Cleridae" genus="Biflabellotillus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF9FFFD13E9FEE4FEE3AB4A" box="[151,302,256,280]" italics="true" pageId="5" pageNumber="389">Biflabellotillus</emphasis>
[sic!]
<emphasis id="B94F391DFFF9FFFD120EFEE4FE1AAB45" box="[368,471,256,279]" italics="true" pageId="5" pageNumber="389">peyrierasi</emphasis>
<bibRefCitation id="EFAA98FEFFF9FFFD12A3FEE4FD46AB45" author="Menier, J. J." box="[477,651,255,280]" pageId="5" pageNumber="389" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier, 1979: 75</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFFD13E9FEA2FCD9A846" pageId="5" pageNumber="389" type="materials_examined">
<paragraph id="8B84E50FFFF9FFFD13E9FEA2FC1DAB32" blockId="5.[151,1436,325,1288]" box="[151,976,325,352]" pageId="5" pageNumber="389">
<materialsCitation id="3B53EF52FFF9FFFD13E9FEA2FC00AB32" box="[151,973,325,352]" county="Mananara Nord District" location="Antanambe" pageId="5" pageNumber="389" specimenCount="1" stateProvince="Analanjirofo Region">
<emphasis id="B94F391DFFF9FFFD13E9FEA2FEFBAB0D" bold="true" box="[151,310,325,352]" pageId="5" pageNumber="389">Type locality.</emphasis>
<location id="8EE4B3D4FFF9FFFD1245FEA2FE72AB32" LSID="urn:lsid:plazi:treatment:03925419FFF9FFFD13E9FF73FA40AF5A:8EE4B3D4FFF9FFFD1245FEA2FE72AB32" box="[315,447,326,352]" county="Mananara Nord District" municipality="Mananara Nord District" name="Antanambe" pageId="5" pageNumber="389" stateProvince="Analanjirofo Region">Antanambe</location>
,
<collectingCounty id="62E59D83FFF9FFFD12B4FEA2FD15AB32" box="[458,728,326,352]" pageId="5" pageNumber="389">Mananara Nord District</collectingCounty>
,
<collectingRegion id="49FF2BEDFFF9FFFD119CFEA2FC00AB32" box="[738,973,326,352]" pageId="5" pageNumber="389">Analanjirofo Region</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="8B84E50FFFF9FFFD13B9FE8EFE69AB9E" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FE8EFE16ABD1" bold="true" box="[199,475,361,388]" pageId="5" pageNumber="389">Type material studied.</emphasis>
<materialsCitation id="3B53EF52FFF9FFFD1298FE8EFE21ABFA" collectionCode="MNHN" country="Madagascar" county="Mananara Nord District" location="Madagascar" pageId="5" pageNumber="389" specimenCount="1" specimenCount-male="1" stateProvince="Analanjirofo Region" typeStatus="holotype">
<typeStatus id="54805BADFFF9FFFD1298FE8EFD91ABD6" box="[486,604,362,388]" pageId="5" pageNumber="389" type="holotype">HOLOTYPE</typeStatus>
,
<specimenCount id="9D3D2E86FFF9FFFD1110FE8EFD4EABD6" box="[622,643,362,388]" count="1" pageId="5" pageNumber="389" type="male"></specimenCount>
(
<collectionCode id="ED2A7DCAFFF9FFFD11EBFE8EFD3FABD6" box="[661,754,362,388]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="389">MNHN</collectionCode>
): [white, printed]
<collectingCountry id="F32CA59FFFF9FFFD10BBFE8EFB81ABD6" box="[965,1100,362,388]" name="Madagascar" pageId="5" pageNumber="389">Madagascar</collectingCountry>
Est / distr. Mananara N. / Antanambe // [red, printed] ♂
</materialsCitation>
<typeStatus id="54805BADFFF9FFFD128BFE6AFD44ABFA" box="[501,649,398,424]" pageId="5" pageNumber="389" type="holotype">HOLOTYPE</typeStatus>
// [white, handwritten]
<taxonomicName id="4C3B9E8CFFF9FFFD10E6FE6AFB62ABFA" authorityName="Menier" authorityYear="1979" box="[920,1199,398,424]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">Biflabelotillus peyrierasi</taxonomicName>
<specimenCount id="9D3D2E86FFF9FFFD17C6FE6AFB00ABFA" box="[1208,1229,398,424]" count="1" pageId="5" pageNumber="389" type="male"></specimenCount>
/ n.sp.
<typeStatus id="54805BADFFF9FFFD165AFE6AFA41ABFA" box="[1316,1420,398,424]" pageId="5" pageNumber="389" type="holotype">Holotype</typeStatus>
/ J.J. MENIER det. 1979.
</paragraph>
<paragraph id="8B84E50FFFF9FFFD13B9FE31FCD9A846" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FE31FDF0ABBD" bold="true" box="[199,573,469,495]" pageId="5" pageNumber="389">Additional material examined.</emphasis>
(
<specimenCount id="9D3D2E86FFF9FFFD112DFE32FD4FABA2" box="[595,642,470,496]" count="1" pageId="5" pageNumber="389" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D3D2E86FFF9FFFD11ECFE32FD0CABA2" box="[658,705,470,496]" count="1" pageId="5" pageNumber="389" type="female">1 ♀</specimenCount>
IJC):
<materialsCitation id="3B53EF52FFF9FFFD106EFE32FCD9A846" collectingDate="1995-11-27" collectorName="Ivo Jenis" country="Madagascar" county="Moramanga" location="East" municipality="Moramanga District" pageId="5" pageNumber="389" specimenCount="1">
<location id="8EE4B3D4FFF9FFFD106EFE32FC8DABA2" LSID="urn:lsid:plazi:treatment:03925419FFF9FFFD13E9FF73FA40AF5A:8EE4B3D4FFF9FFFD106EFE32FC8DABA2" box="[784,832,470,496]" country="Madagascar" county="Moramanga" municipality="Moramanga District" name="East" pageId="5" pageNumber="389">East</location>
<collectingCountry id="F32CA59FFFF9FFFD1035FE32FC19ABA2" box="[843,980,470,496]" name="Madagascar" pageId="5" pageNumber="389">Madagascar</collectingCountry>
,
<location id="8EE4B3D4FFF9FFFD109CFE32FACBABA2" LSID="urn:lsid:plazi:treatment:03925419FFF9FFFD13E9FF73FA40AF5A:8EE4B3D4FFF9FFFD109CFE32FACBABA2" box="[994,1286,470,496]" country="Madagascar" county="Moramanga" municipality="Moramanga District" name="Alaotra-Mangoro Region" pageId="5" pageNumber="389">Alaotra-Mangoro Region</location>
,
<collectingMunicipality id="6BE07F75FFF9FFFD166BFE32FF23A846" pageId="5" pageNumber="389">Moramanga District</collectingMunicipality>
,
<collectingCounty id="62E59D83FFF9FFFD1387FE1EFE4DA846" box="[249,384,506,532]" pageId="5" pageNumber="389">Moramanga</collectingCounty>
env., 25.
<date id="FF85C3CFFFF9FFFD128EFE1EFDB9A847" box="[496,628,506,533]" pageId="5" pageNumber="389" value="1995-11-27">
<collectingDate id="EFC13A27FFF9FFFD128EFE1EFDB9A847" box="[496,628,506,533]" pageId="5" pageNumber="389" value="1995-11-27">27.11.1995</collectingDate>
</date>
,
<collectorName id="26CE80D9FFF9FFFD1101FE1EFD2EA846" box="[639,739,506,532]" pageId="5" pageNumber="389">Ivo Jeniš</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFFD13B9FDFAFBEBA86A" box="[199,1062,542,568]" pageId="5" pageNumber="389" type="discussion">
<paragraph id="8B84E50FFFF9FFFD13B9FDFAFBEBA86A" blockId="5.[151,1436,325,1288]" box="[199,1062,542,568]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FDFAFEF6A86A" bold="true" box="[199,315,542,568]" pageId="5" pageNumber="389">Remarks.</emphasis>
The
<typeStatus id="54805BADFFF9FFFD120AFDFAFE1BA86A" box="[372,470,542,568]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
was fixed by original designation by
<bibRefCitation id="EFAA98FEFFF9FFFD1002FDFAFBEFA86A" author="Menier, J. J." box="[892,1058,542,568]" pageId="5" pageNumber="389" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier (1979)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFFD13B9FDA5FDE3A89A" pageId="5" pageNumber="389" type="diagnosis">
<paragraph id="8B84E50FFFF9FFFD13B9FDA5FDE3A89A" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FDA5FE1DA809" bold="true" box="[199,464,577,603]" pageId="5" pageNumber="389">Differential diagnosis.</emphasis>
<taxonomicName id="4C3B9E8CFFF9FFFD12A6FDA7FDA6A80E" authorityName="Menier" authorityYear="1979" box="[472,619,579,604]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF9FFFD12A6FDA7FDA6A80E" box="[472,619,579,604]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
</taxonomicName>
can be distinguished from both
<taxonomicName id="4C3B9E8CFFF9FFFD10A1FDA7FBBEA80E" authorityName="Pic" authorityYear="1949" box="[991,1139,578,604]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFF9FFFD10A1FDA7FBBEA80E" box="[991,1139,578,604]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3B9E8CFFF9FFFD17D3FDA7FAF5A80E" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1197,1336,578,604]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF9FFFD17D3FDA7FAF5A80E" box="[1197,1336,578,604]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFF9FFFD163FFDA6FA51A80E" bold="true" box="[1345,1436,578,604]" pageId="5" pageNumber="389">
<taxonomicNameLabel id="A27C8466FFF9FFFD163FFDA6FA51A80E" box="[1345,1436,578,604]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
mainly by black parallel sided elytra and tuberculate elytral interstriae, whereas the elytra are brown in
<taxonomicName id="4C3B9E8CFFF9FFFD1675FD83FA51A8D2" authorityName="Pic" authorityYear="1949" box="[1291,1436,614,640]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFF9FFFD1675FD83FA51A8D2" box="[1291,1436,614,640]" italics="true" pageId="5" pageNumber="389">B. foveolatus</emphasis>
</taxonomicName>
and bicoloured in
<taxonomicName id="4C3B9E8CFFF9FFFD1224FD6FFE2CA8F6" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[346,481,650,676]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFF9FFFD1224FD6FFE2CA8F6" box="[346,481,650,676]" italics="true" pageId="5" pageNumber="389">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFF9FFFD129BFD6EFDF1A8F6" bold="true" box="[485,572,650,676]" pageId="5" pageNumber="389">
<taxonomicNameLabel id="A27C8466FFF9FFFD129BFD6EFDF1A8F6" box="[485,572,650,676]" pageId="5" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, subparallel, slightly widened with smooth elytral interstriae in both mentioned species (
<figureCitation id="1300F98AFFF9FFFD1389FD4AFEB5A89A" box="[247,376,686,712]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="5" pageNumber="389">Figs 14, 17</figureCitation>
,
<figureCitation id="1300F98AFFF9FFFD12FAFD4AFE01A89A" box="[388,460,686,712]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="5" pageNumber="389">22, 24</figureCitation>
,
<figureCitation id="1300F98AFFF9FFFD12A6FD4AFE3BA89A" box="[472,502,686,712]" captionStart="FIGURE 2832" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 2832. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="5" pageNumber="389">28</figureCitation>
,
<figureCitation id="1300F98AFFF9FFFD117CFD4AFDEFA89A" box="[514,546,686,712]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="5" pageNumber="389">34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFFD13B9FD35FD9DA9BA" pageId="5" pageNumber="389" type="description">
<paragraph id="8B84E50FFFF9FFFD13B9FD35FAD5A8BF" blockId="5.[151,1436,325,1288]" box="[199,1304,721,749]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FD35FEB8A8B9" bold="true" box="[199,373,721,747]" pageId="5" pageNumber="389">Redescription.</emphasis>
Size of
<typeStatus id="54805BADFFF9FFFD12AFFD36FDFEA8BE" box="[465,563,722,748]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
8.1 x
<quantity id="4CC348EAFFF9FFFD1107FD36FD1FA8BE" box="[633,722,722,748]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="389" unit="mm" value="2.2">2.2 mm</quantity>
. Winged, black, with tarsi pale brown (
<figureCitation id="1300F98AFFF9FFFD17F6FD36FAC1A8BF" box="[1160,1292,722,749]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="5" pageNumber="389">Figs 2125</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFF9FFFD13B9FD12FADBA967" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FD12FEC3A942" bold="true" box="[199,270,758,784]" pageId="5" pageNumber="389">Head.</emphasis>
Head including eyes broader than pronotum (DV) in ratio 1:0.8 (HW =
<quantity id="4CC348EAFFF9FFFD1756FD12FB4CA942" box="[1064,1153,758,784]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="389" unit="mm" value="1.7">1.7 mm</quantity>
). The space between eyes (frons) narrow (HW / FW = 2.8). Ultimate antennomere elongate-oval, 8.8x longer then broad (
<figureCitation id="1300F98AFFF9FFFD17BDFCFEFAC5A967" box="[1219,1288,794,821]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="5" pageNumber="389">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFF9FFFD13B9FCDAFB16A92E" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FCDAFE8AA90A" bold="true" box="[199,327,830,856]" pageId="5" pageNumber="389">Pronotum.</emphasis>
Longer than wide in ratio 1:1.3 (PL =
<quantity id="4CC348EAFFF9FFFD1070FCDAFCA5A90A" box="[782,872,830,856]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="389" unit="mm" value="1.8">1.8 mm</quantity>
, PW =
<quantity id="4CC348EAFFF9FFFD10BCFCDAFBD2A90A" box="[962,1055,830,856]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="389" unit="mm" value="1.4">1.4 mm</quantity>
), cylindrical, apical two thirds of pronotum parallel sided and slightly trangled in apical third, basal third strongly trangled (
<figureCitation id="1300F98AFFF9FFFD17F7FC86FB03A92E" box="[1161,1230,866,892]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="5" pageNumber="389">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFF9FFFD13B9FC62FB39A996" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FC62FED7A9F2" bold="true" box="[199,282,902,928]" pageId="5" pageNumber="389">Elytra.</emphasis>
Parallel, 2.3x longer than their width (EL =
<quantity id="4CC348EAFFF9FFFD1058FC62FC4CA9F2" box="[806,897,902,929]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="5" pageNumber="389" unit="mm" value="5.1">5.1 mm</quantity>
, EW =
<quantity id="4CC348EAFFF9FFFD10A0FC62FBF6A9F2" box="[990,1083,902,928]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="389" unit="mm" value="2.2">2.2 mm</quantity>
). Almost whole elytral surface covered with bituberculate foveae (
<figureCitation id="1300F98AFFF9FFFD115DFC4EFDBBA996" box="[547,630,938,964]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="5" pageNumber="389">Fig. 21</figureCitation>
). Space between foveae smaller than diameter of foveae.
</paragraph>
<paragraph id="8B84E50FFFF9FFFD13B9FC29FD9DA9BA" blockId="5.[151,1436,325,1288]" box="[199,592,973,1000]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FC29FE8BA9B5" bold="true" box="[199,326,973,999]" pageId="5" pageNumber="389">Scutellum.</emphasis>
Heart shaped (
<figureCitation id="1300F98AFFF9FFFD1291FC2AFD89A9BA" box="[495,580,974,1000]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="5" pageNumber="389">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFFD13B9FC16FCE6AEB6" pageId="5" pageNumber="389" type="discussion">
<paragraph id="8B84E50FFFF9FFFD13B9FC16FBC8AE2A" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FC16FEFDAE5E" bold="true" box="[199,304,1010,1036]" pageId="5" pageNumber="389">Remark.</emphasis>
The species was described based on a single male. The female is unknown so far. Fortunately, we had the opportunity to examine the female of
<taxonomicName id="4C3B9E8CFFF9FFFD111BFBF3FD38AE62" authorityName="Menier" authorityYear="1979" box="[613,757,1047,1072]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFF9FFFD111BFBF3FD38AE62" box="[613,757,1047,1072]" italics="true" pageId="5" pageNumber="389">B. peyrierasi</emphasis>
</taxonomicName>
and found out that it fully agrees with the male
<typeStatus id="54805BADFFF9FFFD1675FBF2FAA0AE62" box="[1291,1389,1046,1072]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
and Menier´s description except the following characters: antennae serrate (
<figureCitation id="1300F98AFFF9FFFD10CBFBDEFBCBAE06" box="[949,1030,1082,1108]" captionStart="FIGURE 2832" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 2832. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="5" pageNumber="389">Fig. 30</figureCitation>
) (biflabeloid in males), each elytron with pale-brown humeral spot (whole elytral surface black in males) (
<figureCitation id="1300F98AFFF9FFFD10DAFBBAFC34AE2A" box="[932,1017,1118,1144]" captionStart="FIGURE 2832" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 2832. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="5" pageNumber="389">Fig. 28</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFF9FFFD13B9FB65FCE6AEB6" blockId="5.[151,1436,325,1288]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FB65FE6FAEC9" bold="true" box="[199,418,1153,1179]" pageId="5" pageNumber="389">Variability of size.</emphasis>
Examined male: body length
<quantity id="4CC348EAFFF9FFFD1186FB66FC9CAECE" box="[760,849,1154,1181]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="389" unit="mm" value="7.5">7.5 mm</quantity>
, HW =
<quantity id="4CC348EAFFF9FFFD10D3FB66FBD9AECE" box="[941,1044,1154,1181]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="5" pageNumber="389" unit="mm" value="1.65">1.65 mm</quantity>
, HW / FW = 2.75, PL =
<quantity id="4CC348EAFFF9FFFD164FFB66FA55AECE" box="[1329,1432,1154,1181]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="5" pageNumber="389" unit="mm" value="1.75">1.75 mm</quantity>
, PW =
<quantity id="4CC348EAFFF9FFFD13A1FB42FEFAAE92" box="[223,311,1190,1216]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="389" unit="mm" value="1.2">1.2 mm</quantity>
, EL =
<quantity id="4CC348EAFFF9FFFD12FFFB42FE17AE92" box="[385,474,1190,1217]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="389" unit="mm" value="4.5">4.5 mm</quantity>
, EW =
<quantity id="4CC348EAFFF9FFFD1150FB42FD4AAE92" box="[558,647,1190,1216]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="389" unit="mm" value="2.1">2.1 mm</quantity>
. Examined female body length 8.0 mm, HW = 1.65, HW / FW = 1.94, PL =
<quantity id="4CC348EAFFF9FFFD13ABFB2EFEE0AEB6" box="[213,301,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="389" unit="mm" value="1.9">1.9 mm</quantity>
, PW =
<quantity id="4CC348EAFFF9FFFD1201FB2EFE1AAEB6" box="[383,471,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="389" unit="mm" value="1.4">1.4 mm</quantity>
, EL =
<quantity id="4CC348EAFFF9FFFD115CFB2EFDB7AEB6" box="[546,634,1226,1253]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="5" pageNumber="389" unit="mm" value="5.4">5.4 mm</quantity>
, EW =
<quantity id="4CC348EAFFF9FFFD11B0FB2EFCEAAEB6" box="[718,807,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="389" unit="mm" value="2.4">2.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFF9FFFD13B9FB09FA40AF5A" box="[199,1421,1261,1288]" pageId="5" pageNumber="389" type="distribution">
<paragraph id="8B84E50FFFF9FFFD13B9FB09FA40AF5A" blockId="5.[151,1436,325,1288]" box="[199,1421,1261,1288]" pageId="5" pageNumber="389">
<emphasis id="B94F391DFFF9FFFD13B9FB09FEACAF55" bold="true" box="[199,353,1261,1287]" pageId="5" pageNumber="389">Distribution.</emphasis>
Eastern
<collectingCountry id="F32CA59FFFF9FFFD12BCFB0AFD86AF5A" box="[450,587,1262,1288]" name="Madagascar" pageId="5" pageNumber="389">Madagascar</collectingCountry>
, known only from Moramanga and Maroantsetra Districts so far (
<figureCitation id="1300F98AFFF9FFFD1653FB0AFA4CAF5A" box="[1325,1409,1262,1288]" captionStart="FIGURE 38" captionStartId="9.[151,250,1118,1142]" captionTargetBox="[489,1099,181,1094]" captionTargetId="figure-26@9.[489,1099,181,1094]" captionTargetPageId="9" captionText="FIGURE 38. Distribution map of the genus Biflabelotillus. B. foveolatus—red triangle, B. peyrierasi—white circle, B. bicolorus sp. nov.—yellow rhombus." figureDoi="http://doi.org/10.5281/zenodo.13915947" httpUri="https://zenodo.org/record/13915947/files/figure.png" pageId="5" pageNumber="389">Fig. 38</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,261 @@
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<subSubSection id="C321B684FFFDFFF913E9FE3EFDDFABA6" box="[151,530,474,501]" pageId="1" pageNumber="385" type="nomenclature">
<paragraph id="8B84E50FFFFDFFF913E9FE3EFDDFABA6" blockId="1.[151,530,474,501]" box="[151,530,474,501]" pageId="1" pageNumber="385">
<heading id="D0CC5263FFFDFFF913E9FE3EFDDFABA6" bold="true" box="[151,530,474,501]" fontSize="11" level="1" pageId="1" pageNumber="385" reason="1">
<emphasis id="B94F391DFFFDFFF913E9FE3EFDDFABA6" bold="true" box="[151,530,474,501]" pageId="1" pageNumber="385">
Genus
<taxonomicName id="4C3B9E8CFFFDFFF91390FE3EFDDFABA6" authority="Pic, 1949" authorityName="Pic" authorityYear="1949" box="[238,530,474,501]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFFDFFF91390FE3EFE54ABA7" bold="true" box="[238,409,474,501]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
<bibRefCitation id="EFAA98FEFFFDFFF912DFFE3EFDDFABA6" author="Pic, M." box="[417,530,474,500]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFDFFF913E9FDFEFBC5A860" box="[151,1032,538,562]" pageId="1" pageNumber="385" type="reference_group">
<paragraph id="8B84E50FFFFDFFF913E9FDFEFBC5A860" blockId="1.[151,1436,538,685]" box="[151,1032,538,562]" pageId="1" pageNumber="385">
<treatmentCitationGroup id="AB2BC221FFFDFFF913E9FDFEFBC5A860" box="[151,1032,538,562]" pageId="1" pageNumber="385">
<treatmentCitation id="0A9AC31EFFFDFFF913E9FDFEFE65A860" author="Pic, M." box="[151,424,538,562]" page="3" pageId="1" pageNumber="385" year="1949">
<taxonomicName id="4C3B9E8CFFFDFFF913E9FDFEFE65A860" authority="Pic, 1949: 3" authorityName="Pic" authorityPageNumber="3" authorityYear="1949" box="[151,424,538,562]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFFDFFF913E9FDFEFEEAA860" box="[151,295,538,562]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
<bibRefCitation id="EFAA98FEFFFDFFF91250FDFEFE65A860" author="Pic, M." box="[302,424,538,562]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949: 3</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
<treatmentCitation id="0A9AC31EFFFDFFF911D8FDFEFBC9A860" author="Edwards, M. A. &amp; Hopwood, A. T." box="[678,1028,538,562]" page="36" pageId="1" pageNumber="385" year="1966">
<bibRefCitation id="EFAA98FEFFFDFFF911D8FDFEFC13A860" author="Edwards, M. A. &amp; Hopwood, A. T." box="[678,990,538,562]" pageId="1" pageNumber="385" refId="ref5094" refString="Edwards, M. A. &amp; Hopwood, A. T. (1966) Nomenclator Zoologicus. Vol. VI. 1946 - 1955. Zoological Society of London, London, viii + 329 pp." type="book" year="1966">EDWARDS &amp; HOPWOOD (1966)</bibRefCitation>
: 36
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFDFFF91770FDFEFDE8A802" pageId="1" pageNumber="385" type="type_taxon">
<paragraph id="8B84E50FFFFDFFF91770FDFEFDE8A802" blockId="1.[151,1436,538,685]" pageId="1" pageNumber="385">
<typeStatus id="54805BADFFFDFFF91770FDFEFB8DA860" box="[1038,1088,538,562]" pageId="1" pageNumber="385">Type</typeStatus>
species:
<taxonomicName id="4C3B9E8CFFFDFFF917E0FDFEFA51A860" authorityYear="1949" box="[1182,1436,538,562]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF917E0FDFEFA51A860" box="[1182,1436,538,562]" italics="true" pageId="1" pageNumber="385">Biflabelotillus foveolatus</emphasis>
</taxonomicName>
<bibRefCitation id="EFAA98FEFFFDFFF913B9FDDCFE8CA803" author="Pic, M." box="[199,321,568,593]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949: 3</bibRefCitation>
(original description).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFDFFF913E9FDB3FB90A8FF" pageId="1" pageNumber="385" type="reference_group">
<paragraph id="8B84E50FFFFDFFF913E9FDB3FC1EA83D" blockId="1.[151,1436,538,685]" box="[151,979,599,623]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913E9FDB3FEEAA83D" box="[151,295,599,623]" italics="true" pageId="1" pageNumber="385">Biflabellotilus</emphasis>
[sic!]
<bibRefCitation id="EFAA98FEFFFDFFF91202FDB3FE27A83D" author="Pic, M." box="[380,490,599,623]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic (1949)</bibRefCitation>
: 3 (used as
<emphasis id="B94F391DFFFDFFF91121FDB3FD22A83D" box="[607,751,599,623]" italics="true" pageId="1" pageNumber="385">Biflabellotilus</emphasis>
<emphasis id="B94F391DFFFDFFF9118BFDB3FC90A83D" box="[757,861,599,623]" italics="true" pageId="1" pageNumber="385">foveolatus</emphasis>
<bibRefCitation id="EFAA98FEFFFDFFF9101DFDB3FC0AA83D" author="Pic, M." box="[867,967,599,623]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949</bibRefCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFDFFF913E9FD92FB90A8FF" blockId="1.[151,1436,538,685]" pageId="1" pageNumber="385">
<taxonomicName id="4C3B9E8CFFFDFFF913E9FD92FEE3A8DC" authorityName="Pic" authorityYear="1949" box="[151,302,630,654]" class="Insecta" family="Cleridae" genus="Biflabellotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFFDFFF913E9FD92FEE3A8DC" box="[151,302,630,654]" italics="true" pageId="1" pageNumber="385">Biflabellotillus</emphasis>
</taxonomicName>
[sic!]
<bibRefCitation id="EFAA98FEFFFDFFF912F7FD92FD9EA8DC" author="Corporaal, J. B." box="[393,595,630,654]" pageId="1" pageNumber="385" refId="ref5062" refString="Corporaal, J. B. (1950) Cleridae. Coleopterorum Catalogus. Supplementa Pars 23. Uitgeverij. Dr. W. Junk, ' s-Gravenhage, 373 pp." type="book" year="1950">CORPORAAL (1950)</bibRefCitation>
: 17 (catalogue);
<bibRefCitation id="EFAA98FEFFFDFFF9107FFD92FC7CA8DC" author="Winkler, J. R." box="[769,945,630,654]" pageId="1" pageNumber="385" refId="ref5415" refString="Winkler, J. R. (1961) Die Buntkafer (Cleridae). A. Ziemsen Verlag, Wittenberg, Lutherstadt, 108 pp." type="book" year="1961">WINKLER (1961)</bibRefCitation>
: 20, 32 (list of genera);
<bibRefCitation id="EFAA98FEFFFDFFF917CEFD92FA82A8DC" author="Menier, J. J." box="[1200,1359,630,654]" pageId="1" pageNumber="385" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">MENIER (1979)</bibRefCitation>
: 75, 76 (redefinition, key to species);
<bibRefCitation id="EFAA98FEFFFDFFF91288FD71FD7BA8FF" author="Mawdsley, J." box="[502,694,661,685]" pageId="1" pageNumber="385" pagination="19 - 29" refId="ref5261" refString="Mawdsley, J. (1993) A key to the species of the genus Myrmecomaea Fairmaire (Coleoptera: Cleridae) with checklist of species, notes on mimicry, and discussion of endemism in Madagascan Cleridae. Y. E. S. Quarterly, 10 (4), 19 - 29" type="journal article" year="1993">MAWDSLEY (1993)</bibRefCitation>
: 26 (list of endemic Madagascan genera).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFDFFF913E9FD3FFE67A96F" pageId="1" pageNumber="385" type="multiple">
<paragraph id="8B84E50FFFFDFFF913E9FD3FFE67A96F" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913E9FD3FFF1BA8A7" bold="true" box="[151,214,731,757]" pageId="1" pageNumber="385">Note.</emphasis>
<bibRefCitation id="EFAA98FEFFFDFFF913A3FD3FFE9EA8A7" author="Pic, M." box="[221,339,731,757]" pageId="1" pageNumber="385" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic (1949)</bibRefCitation>
used two different spellings in his original paper,
<taxonomicName id="4C3B9E8CFFFDFFF910FBFD3FFBE9A8A7" authorityName="Pic" authorityYear="1949" box="[901,1060,731,757]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFFDFFF910FBFD3FFBE9A8A7" box="[901,1060,731,757]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
</taxonomicName>
and
<emphasis id="B94F391DFFFDFFF91724FD3FFB34A8A7" box="[1114,1273,731,757]" italics="true" pageId="1" pageNumber="385">Biflabellotilus</emphasis>
(pp. 3). As far as we know, no one has acted as First Reviser (
<bibRefCitation id="EFAA98FEFFFDFFF911DDFD1BFCE5A94B" author="ICZN" box="[675,808,767,793]" pageId="1" pageNumber="385" refId="ref5225" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. Adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
, Article 24.2.3) and
<emphasis id="B94F391DFFFDFFF91772FD1BFB62A94B" bold="true" box="[1036,1199,767,793]" pageId="1" pageNumber="385">we here select</emphasis>
<taxonomicName id="4C3B9E8CFFFDFFF917C8FD1BFA98A94B" authorityName="Pic" authorityYear="1949" box="[1206,1365,767,793]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFFDFFF917C8FD1BFA98A94B" box="[1206,1365,767,793]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
</taxonomicName>
as the correct original spelling.
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFDFFF913B9FCA2FC10AE9B" pageId="1" pageNumber="385" type="diagnosis">
<paragraph id="8B84E50FFFFDFFF913B9FCA2FC10AE9B" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913B9FCA2FE8FA932" bold="true" box="[199,322,838,864]" pageId="1" pageNumber="385">Diagnosis.</emphasis>
Although a review of Madagascan tilline genera is needed to determine the generic status for most of the genera (
<bibRefCitation id="EFAA98FEFFFDFFF91247FC8FFE02A9D7" author="Menier, J. J." box="[313,463,875,901]" pageId="1" pageNumber="385" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier 1979</bibRefCitation>
, Gerstmeier &amp; Weis 2009),
<taxonomicName id="4C3B9E8CFFFDFFF91074FC8FFC64A9D7" authorityName="Pic" authorityYear="1949" box="[778,937,875,901]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFFDFFF91074FC8FFC64A9D7" box="[778,937,875,901]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
</taxonomicName>
seems to be a relatively well defined genus. According to
<bibRefCitation id="EFAA98FEFFFDFFF91251FC6BFE1FA9FB" author="Menier, J. J." box="[303,466,911,937]" pageId="1" pageNumber="385" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier (1979)</bibRefCitation>
, this genus belongs to the group of several genera (besides
<taxonomicName id="4C3B9E8CFFFDFFF91720FC6BFB30A9FB" authorityName="Pic" authorityYear="1949" box="[1118,1277,911,937]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFFDFFF91720FC6BFB30A9FB" box="[1118,1277,911,937]" italics="true" pageId="1" pageNumber="385">Biflabelotillus</emphasis>
</taxonomicName>
e.g.
<taxonomicName id="4C3B9E8CFFFDFFF9164EFC6BFE81A99F" authority="Fairmaire, 1903" authorityName="Fairmaire" authorityYear="1903" class="Insecta" family="Cleridae" genus="Archalius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFFDFFF9164EFC6BFA51A9FB" box="[1328,1436,911,937]" italics="true" pageId="1" pageNumber="385">Archalius</emphasis>
Fairmaire, 1903
</taxonomicName>
) within Madagascan
<taxonomicName id="4C3B9E8CFFFDFFF9113FFC57FD6EA99F" authorityName="Latreille" authorityYear="1802" box="[577,675,947,973]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="family">Cleridae</taxonomicName>
, whose elytral surface is covered with bituberculate foveae - a pair of tubercles situated opposite each other on top of a large round depression (see also
<figureCitation id="1300F98AFFFDFFF91719FC33FB72A9A3" box="[1127,1215,983,1009]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="1" pageNumber="385">Figs 20</figureCitation>
,
<figureCitation id="1300F98AFFFDFFF917B2FC33FB27A9A3" box="[1228,1258,983,1009]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="1" pageNumber="385">27</figureCitation>
,
<figureCitation id="1300F98AFFFDFFF91786FC33FADBA9A3" box="[1272,1302,983,1009]" captionStart="FIGURE 2832" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 2832. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="1" pageNumber="385">32</figureCitation>
,
<figureCitation id="1300F98AFFFDFFF9165DFC33FA8CA9A3" box="[1315,1345,983,1009]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="1" pageNumber="385">37</figureCitation>
, 41). It, however, can reliably be distinguished by biflabeloid antennae, with two flabelae situated at the beginning of each antennomere IIIX. Moreover, the flabelae have the same diameter along their whole length (see also
<figureCitation id="1300F98AFFFDFFF91673FBFBFABDAE6B" box="[1293,1392,1055,1081]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="1" pageNumber="385">Figs 13</figureCitation>
). In other Madagascan genera which possess flabeloid or biflabeloid antennae, the flabelae are situated in the middle or at the end of antennomeres IIIX and their diameter is not equal along their length but usually gradually widened toward their apex. Moreover, in some genera (e.g. in
<taxonomicName id="4C3B9E8CFFFDFFF91186FB6FFB8CAEF7" authority="Fairmaire, 1885" authorityName="Fairmaire" authorityYear="1885" box="[760,1089,1162,1189]" class="Insecta" family="Cleridae" genus="Diplocladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B94F391DFFFDFFF91186FB6FFC4CAEF7" box="[760,897,1163,1189]" italics="true" pageId="1" pageNumber="385">Diplocladus</emphasis>
Fairmaire, 1885
</taxonomicName>
; Gerstmeier &amp; Weis 2009) the base of flabelae is so close that it seems to be in fact one bifurcate flabella.
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFDFFFA13B9FB36FB0EAAAB" lastPageId="2" lastPageNumber="386" pageId="1" pageNumber="385" type="description">
<paragraph id="8B84E50FFFFDFFF913B9FB36FF07AF43" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913B9FB36FEB8AEBE" bold="true" box="[199,373,1234,1260]" pageId="1" pageNumber="385">Redescription.</emphasis>
Body cylindrical, shiny, sparsely pubescent. Body length
<quantity id="4CC348EAFFFDFFF9176DFB36FB61AEBF" box="[1043,1196,1234,1261]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" metricValueMax="11.9" metricValueMin="7.5" pageId="1" pageNumber="385" unit="mm" value="9.7" valueMax="11.9" valueMin="7.5">7.511.9 mm</quantity>
. Body width
<quantity id="4CC348EAFFFDFFF91636FB37FF0BAF43" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.6" metricValueMin="2.1" pageId="1" pageNumber="385" unit="mm" value="2.85" valueMax="3.6" valueMin="2.1">2.13.6 mm</quantity>
.
</paragraph>
<paragraph id="8B84E50FFFFDFFF913B9FAFFFDEAAC2B" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913B9FAFFFEC3AF67" bold="true" box="[199,270,1307,1333]" pageId="1" pageNumber="385">Head.</emphasis>
Head including eyes broader than pronotum. Eyes large, with sparse erect setae, protruding laterally, anteriorly kidney-shaped (
<figureCitation id="1300F98AFFFDFFF912BDFADBFDF6AF0B" box="[451,571,1343,1369]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="1" pageNumber="385">Figs 712</figureCitation>
). Frons and vertex wrinkled. Labrum bilobed, anterior margin with yellow setae and emarginate in middle (
<figureCitation id="1300F98AFFFDFFF9117EFA87FDB8AF2F" box="[512,629,1379,1405]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="1" pageNumber="385">Figs 712</figureCitation>
). Terminal maxillary palpomeres cylindrical; terminal labial palpomeres securiform. Clypeus shiny, straight anteriorly, slightly dilated posteriad, sparsely pubescent.Antennae 11-segmented, biflabeloid, with flabelae situated at base of each antennomere III-X (
<figureCitation id="1300F98AFFFDFFF910C4FA4FFBEFAF97" box="[954,1058,1451,1477]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="1" pageNumber="385">Figs 13</figureCitation>
). Flabelae 23 times longer than length of respective antennomere. Flabelae equal in width along their whole length, divided, unjointed.Antennomere I thickened, approx. 2.5 longer than antennomere II, which is strongly shortened. Antennomere II with a small pronounced bump on inner side. Antennomeres III-X approx. equal in length. Dorsal side of antennomeres sparsely covered with long yellow recumbent hairs whereas inner side of antennomeres and flabelae is densely covered with shorter white erect hairs (
<figureCitation id="1300F98AFFFDFFF912CAF9BBFDD4AC2B" box="[436,537,1631,1657]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="1" pageNumber="385">Figs 13</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFDFFF913B9F967FCC4AC93" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913B9F967FE8AACCF" bold="true" box="[199,327,1667,1693]" pageId="1" pageNumber="385">Pronotum.</emphasis>
Shiny, longer than wide, cylindrical, surface roughly and irregularly wrinkled, whole surface covered with long erect and semi-erect hairs (
<figureCitation id="1300F98AFFFDFFF911E8F943FD30AC93" box="[662,765,1703,1729]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="1" pageNumber="385">Figs 79</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFDFFF913B9F92FFD09AD27" blockId="1.[151,1437,731,1981]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913B9F92FFED7ACB7" bold="true" box="[199,282,1739,1765]" pageId="1" pageNumber="385">Elytra.</emphasis>
Elytra shiny, parallel or subparallel, distinctly longer than their width, with posterior half parallel or slightly widening posteriad, whole surface sparsely covered with semi-erect yellow hairs, more densely in lateral part of elytra. Humeri well developed, distinct, almost rectangular. Elytral surface with bituberculate foveae - a pair of tubercles situated opposite each other on the top of the large round depressions (
<figureCitation id="1300F98AFFFDFFF91759F8D3FBB0AD03" box="[1063,1149,1847,1873]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="1" pageNumber="385">Figs 20</figureCitation>
,
<figureCitation id="1300F98AFFFDFFF917F9F8D3FB68AD03" box="[1159,1189,1847,1873]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="1" pageNumber="385">27</figureCitation>
,
<figureCitation id="1300F98AFFFDFFF917CEF8D3FB01AD03" box="[1200,1228,1847,1873]" captionStart="FIGURE 2832" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 2832. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="1" pageNumber="385">32</figureCitation>
and
<figureCitation id="1300F98AFFFDFFF9167EF8D3FAEDAD03" box="[1280,1312,1847,1873]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="1" pageNumber="385">37</figureCitation>
). The latter arranged in regular rows (
<figureCitation id="1300F98AFFFDFFF912C6F8BFFDC2AD27" box="[440,527,1883,1909]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="1" pageNumber="385">Figs 17</figureCitation>
,
<figureCitation id="1300F98AFFFDFFF91165F8BFFDF4AD27" box="[539,569,1883,1909]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="1" pageNumber="385">24</figureCitation>
,
<figureCitation id="1300F98AFFFDFFF9113BF8BFFDACAD27" box="[581,609,1883,1909]" captionStart="FIGURE 2832" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 2832. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="1" pageNumber="385">28</figureCitation>
and
<figureCitation id="1300F98AFFFDFFF911E6F8BFFD75AD27" box="[664,696,1883,1909]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="1" pageNumber="385">34</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFDFFF913B9F89AFC0FADCB" blockId="1.[151,1437,731,1981]" box="[199,962,1918,1945]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913B9F89AFE8BADCA" bold="true" box="[199,326,1918,1944]" pageId="1" pageNumber="385">Scutellum.</emphasis>
Almost square or subtriangular (
<figureCitation id="1300F98AFFFDFFF911C8F89BFCC0ADCB" box="[694,781,1919,1945]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="1" pageNumber="385">Figs 19</figureCitation>
,
<figureCitation id="1300F98AFFFDFFF91067F89BFCFAADCB" box="[793,823,1919,1945]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="1" pageNumber="385">26</figureCitation>
,
<figureCitation id="1300F98AFFFDFFF9103DF89BFC92ADCB" box="[835,863,1919,1945]" captionStart="FIGURE 2832" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 2832. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="1" pageNumber="385">31</figureCitation>
and
<figureCitation id="1300F98AFFFDFFF910EBF89BFC78ADCB" box="[917,949,1919,1945]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="1" pageNumber="385">36</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFDFFF913B9F847FCFDADEF" blockId="1.[151,1437,731,1981]" box="[199,816,1955,1981]" pageId="1" pageNumber="385">
<emphasis id="B94F391DFFFDFFF913B9F847FECBADEF" bold="true" box="[199,262,1955,1981]" pageId="1" pageNumber="385">Legs.</emphasis>
Relatively densely covered with long erect hairs.
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9FF73FB0EAAAB" blockId="2.[151,1436,150,285]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FF73FEB1AAE2" bold="true" box="[199,380,150,177]" pageId="2" pageNumber="386">Lower surface.</emphasis>
Shiny, covered with long semi-erect hairs; pro-intercoxal process narrow, slightly expanded distally; anterior margin of mesoventrite straight, lateral margins divergent posteriad, meso-intercoxal process narrow, pointed; metaventrite subquadrate; abdomen with six visible ventrites (
<figureCitation id="1300F98AFFFEFFFA1770FF3BFBADAAAB" box="[1038,1120,223,249]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Fig. 18</figureCitation>
,
<figureCitation id="1300F98AFFFEFFFA1712FF3BFB47AAAB" box="[1132,1162,223,249]" captionStart="FIGURE 2832" captionStartId="7.[151,250,1586,1610]" captionTargetBox="[170,1418,184,1562]" captionTargetId="figure-26@7.[170,1418,181,1562]" captionTargetPageId="7" captionText="FIGURE 2832. Biflabelotillus peyrierasi Menier, 1979, female. Fig. 28—habitus, dorsal view; 29—habitus, ventral view; 30—antenna (red arrow indicates last antennomere); 31—scutellum; 32—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915943" httpUri="https://zenodo.org/record/13915943/files/figure.png" pageId="2" pageNumber="386">29</figureCitation>
,
<figureCitation id="1300F98AFFFEFFFA17E8FF3AFB78AAAB" box="[1174,1205,222,249]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="2" pageNumber="386">35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13B9FEE6FD14AB4F" box="[199,729,258,285]" pageId="2" pageNumber="386" type="distribution">
<paragraph id="8B84E50FFFFEFFFA13B9FEE6FD14AB4F" blockId="2.[151,1436,150,285]" box="[199,729,258,285]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FEE6FEACAB4E" bold="true" box="[199,353,258,284]" pageId="2" pageNumber="386">Distribution.</emphasis>
Central and Eastern
<collectingCountry id="F32CA59FFFFEFFFA1135FEE7FD19AB4F" box="[587,724,259,285]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,417 @@
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<paragraph id="8B84E50FFFFEFFFA13E9FE8AFDF3ABDA" blockId="2.[151,574,366,429]" box="[151,574,366,393]" pageId="2" pageNumber="386">
<heading id="D0CC5263FFFEFFFA13E9FE8AFDF3ABDA" bold="true" box="[151,574,366,393]" fontSize="11" level="1" pageId="2" pageNumber="386" reason="1">
<taxonomicName id="4C3B9E8CFFFEFFFA13E9FE8AFDF3ABDA" authority="Pic, 1949" authorityName="Pic" authorityYear="1949" box="[151,574,366,393]" class="Insecta" family="Cleridae" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA13E9FE8AFDF3ABDA" bold="true" box="[151,574,366,393]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13E9FE8AFE0BABDB" bold="true" box="[151,454,366,393]" italics="true" pageId="2" pageNumber="386">Biflabelotillus foveolatus</emphasis>
<bibRefCitation id="EFAA98FEFFFEFFFA12B3FE8AFDF3ABDA" author="Pic, M." box="[461,574,366,392]" pageId="2" pageNumber="386" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C321B684FFFEFFFA13E9FE77FE5FABFF" box="[151,402,403,429]" pageId="2" pageNumber="386" type="description">
<paragraph id="8B84E50FFFFEFFFA13E9FE77FE5FABFF" blockId="2.[151,574,366,429]" box="[151,402,403,429]" pageId="2" pageNumber="386">
<figureCitation id="1300F98AFFFEFFFA13E9FE77FE8DABFF" box="[151,320,403,429]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="2" pageNumber="386">Figs 1, 4, 7, 10</figureCitation>
, 1320
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13E9FE33FEF9A87E" pageId="2" pageNumber="386" type="reference_group">
<paragraph id="8B84E50FFFFEFFFA13E9FE33FCF8ABBD" blockId="2.[151,1436,471,556]" box="[151,821,471,495]" pageId="2" pageNumber="386">
<treatmentCitationGroup id="AB2BC221FFFEFFFA13E9FE33FCF8ABBD" box="[151,821,471,495]" pageId="2" pageNumber="386">
<taxonomicName id="4C3B9E8CFFFEFFFA13E9FE33FD9CABBD" authority="Pic, 1949: 3" authorityName="Pic" authorityPageNumber="3" authorityYear="1949" box="[151,593,471,495]" class="Insecta" family="Cleridae" genus="Biflabellotilus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA13E9FE33FEEAABBD" box="[151,295,471,495]" italics="true" pageId="2" pageNumber="386">Biflabellotilus</emphasis>
<treatmentCitation id="0A9AC31EFFFEFFFA1253FE33FD9CABBD" author="Pic, M." box="[301,593,471,495]" page="3" pageId="2" pageNumber="386" year="1949">
[sic!]
<emphasis id="B94F391DFFFEFFFA1217FE33FE1CABBD" box="[361,465,471,495]" italics="true" pageId="2" pageNumber="386">foveolatus</emphasis>
<bibRefCitation id="EFAA98FEFFFEFFFA12A9FE33FD9CABBD" author="Pic, M." box="[471,593,471,495]" pageId="2" pageNumber="386" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic, 1949: 3</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(original description).
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<paragraph id="8B84E50FFFFEFFFA13E9FE12FEF9A87E" blockId="2.[151,1436,471,556]" pageId="2" pageNumber="386">
<treatmentCitationGroup id="AB2BC221FFFEFFFA13E9FE12FEF9A87E" pageId="2" pageNumber="386">
<taxonomicName id="4C3B9E8CFFFEFFFA13E9FE12FE18A85C" authority="Pic, 1949" authorityName="Pic" authorityYear="1949" box="[151,469,502,526]" class="Insecta" family="Cleridae" genus="Biflabellotillus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA13E9FE12FEE3A85C" box="[151,302,502,526]" italics="true" pageId="2" pageNumber="386">Biflabellotillus</emphasis>
[sic!]
<emphasis id="B94F391DFFFEFFFA1213FE12FE18A85C" box="[365,469,502,526]" italics="true" pageId="2" pageNumber="386">foveolatus</emphasis>
</taxonomicName>
<treatmentCitation id="0A9AC31EFFFEFFFA1295FE12FD1CA85C" author="Corporaal, J. B." box="[491,721,502,526]" page="17" pageId="2" pageNumber="386" year="1950">
<bibRefCitation id="EFAA98FEFFFEFFFA1295FE12FD7DA85C" author="Corporaal, J. B." box="[491,688,502,526]" pageId="2" pageNumber="386" refId="ref5062" refString="Corporaal, J. B. (1950) Cleridae. Coleopterorum Catalogus. Supplementa Pars 23. Uitgeverij. Dr. W. Junk, ' s-Gravenhage, 373 pp." type="book" year="1950">CORPORAAL (1950)</bibRefCitation>
: 17
</treatmentCitation>
(catalogue);
<treatmentCitation id="0A9AC31EFFFEFFFA102DFE12FBEEA85C" author="Winkler, J. R." box="[851,1059,502,526]" page="32" pageId="2" pageNumber="386" year="1961">
<bibRefCitation id="EFAA98FEFFFEFFFA102DFE12FC32A85C" author="Winkler, J. R." box="[851,1023,502,526]" pageId="2" pageNumber="386" refId="ref5415" refString="Winkler, J. R. (1961) Die Buntkafer (Cleridae). A. Ziemsen Verlag, Wittenberg, Lutherstadt, 108 pp." type="book" year="1961">WINKLER (1961)</bibRefCitation>
: 32
</treatmentCitation>
;
<treatmentCitation id="0A9AC31EFFFEFFFA1752FE12FB25A85C" author="Menier, J. J." box="[1068,1256,502,526]" page="75" pageId="2" pageNumber="386" year="1979">
<bibRefCitation id="EFAA98FEFFFEFFFA1752FE12FB05A85C" author="Menier, J. J." box="[1068,1224,502,526]" pageId="2" pageNumber="386" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">MENIER (1979)</bibRefCitation>
: 75
</treatmentCitation>
(habitat character drawings).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<materialsCitation id="3B53EF52FFFEFFFA13E9FDBFFB4AA827" box="[151,1159,602,629]" country="Madagascar" county="Maroantsetra District" location="Antongonivitsika" municipality="Antongonivitsika" pageId="2" pageNumber="386" specimenCount="1" stateProvince="Analanjirofo Region">
<emphasis id="B94F391DFFFEFFFA13E9FDBFFEFAA827" bold="true" box="[151,311,602,629]" pageId="2" pageNumber="386">Type locality.</emphasis>
<collectingCountry id="F32CA59FFFFEFFFA1240FDBFFE0AA827" box="[318,455,603,629]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
,
<collectingRegion id="49FF2BEDFFFEFFFA12AEFDBFFD76A827" box="[464,699,603,629]" pageId="2" pageNumber="386">Analanjirofo Region</collectingRegion>
,
<collectingCounty id="62E59D83FFFEFFFA11BBFDBFFC7AA827" box="[709,951,603,629]" pageId="2" pageNumber="386">Maroantsetra District</collectingCounty>
,
<collectingMunicipality id="6BE07F75FFFEFFFA10BFFDBFFB4AA827" box="[961,1159,603,629]" pageId="2" pageNumber="386">Antongonivitsika</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9FD9BFC7EA97B" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FD9BFE2EA8CA" bold="true" box="[199,483,638,665]" pageId="2" pageNumber="386">Type material studied.</emphasis>
<materialsCitation id="3B53EF52FFFEFFFA128CFD9BFC27A8B3" collectingDate="1947-11" collectionCode="MNHN" collectorName="Michel" country="Madagascar" county="Maroantsetra District" location="Antongonivitsika" municipality="Antongonivitsika" pageId="2" pageNumber="386" specimenCount="1" specimenCount-male="1" stateProvince="Analanjirofo Region" typeStatus="lectotype">
<typeStatus id="54805BADFFFEFFFA128CFD9BFDBDA8CB" box="[498,624,639,665]" pageId="2" pageNumber="386" type="lectotype">LECTOTYPE</typeStatus>
(designated here),
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(
<collectionCode id="ED2A7DCAFFFEFFFA10F5FD9BFC25A8CB" box="[907,1000,639,665]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="386">MNHN</collectionCode>
): [white, handwritten]
<collectingCountry id="F32CA59FFFFEFFFA1780FD9BFA48A8CB" box="[1278,1413,639,665]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
/
<collectingMunicipality id="6BE07F75FFFEFFFA13E9FD47FE94A8EF" box="[151,345,675,701]" pageId="2" pageNumber="386">Antongonivitsika</collectingMunicipality>
/
<date id="FF85C3CFFFFEFFFA1211FD47FE74A8EF" box="[367,441,675,701]" pageId="2" pageNumber="386" value="1947-11">
<collectingDate id="EFC13A27FFFEFFFA1211FD47FE74A8EF" box="[367,441,675,701]" pageId="2" pageNumber="386" value="1947-11">XI. 47</collectingDate>
</date>
,
<collectorName id="26CE80D9FFFEFFFA12BAFD47FDD4A8EF" box="[452,537,675,701]" pageId="2" pageNumber="386">Michel</collectorName>
! [white, handwritten] Type // [white, handwritten]
<taxonomicName id="4C3B9E8CFFFEFFFA1731FD47FB22A8EF" authorityName="Pic" authorityYear="1949" box="[1103,1263,675,701]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="genus">Biflabelotillus</taxonomicName>
ng / foveolatus nsp // [white, handwritten] genre pres Paradoxocerus Kr. // [red, printed] ♂
</materialsCitation>
<materialsCitation id="3B53EF52FFFEFFFA108CFD23FB8AA957" collectorName="J. J. MENIER" country="Madagascar" county="Maroantsetra District" location="Antongonivitsika" municipality="Antongonivitsika" pageId="2" pageNumber="386" specimenCount="1" stateProvince="Analanjirofo Region" typeStatus="holotype">
<typeStatus id="54805BADFFFEFFFA108CFD23FB4BA8B3" box="[1010,1158,711,737]" pageId="2" pageNumber="386" type="holotype">HOLOTYPE</typeStatus>
// [blue] // [red, printed] SPECIMEN FIGURE / FAUNE DE MADAG. /
<collectorName id="26CE80D9FFFEFFFA11A5FD0FFCB8A957" box="[731,885,747,773]" pageId="2" pageNumber="386">J.J. MENIER</collectorName>
/ 1933 1991 //
</materialsCitation>
<typeStatus id="54805BADFFFEFFFA172CFD0FFB3FA957" box="[1106,1266,747,773]" pageId="2" pageNumber="386" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="4C3B9E8CFFFEFFFA1780FD0FFECCA97B" authorityName="Pavel &amp; Purchart" authorityYear="2024" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA1780FD0FFECCA97B" italics="true" pageId="2" pageNumber="386">Biflabelotillus bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA1276FCEBFEAFA97B" bold="true" box="[264,354,783,809]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA1276FCEBFEAFA97B" box="[264,354,783,809]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
det. F. Pavel &amp; L. Purchart 2024 [red, printed label].
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9FCD6FC85A98F" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FCD6FDF8A91E" bold="true" box="[199,565,818,844]" pageId="2" pageNumber="386">Additional material examined.</emphasis>
(
<specimenCount id="9D3D2E86FFFEFFFA1136FCD7FDBCA91F" box="[584,625,819,845]" count="1" pageId="2" pageNumber="386" type="male">1 ♂</specimenCount>
IJC):
<materialsCitation id="3B53EF52FFFEFFFA11C6FCD7FB6CA923" collectingDate="2001-11-28" collectorName="Z. Mracek" country="Madagascar" county="Central" location="Ankazobe District" municipality="Analamanga Region" pageId="2" pageNumber="386" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<collectingCounty id="62E59D83FFFEFFFA11C6FCD7FCC7A91F" box="[696,778,819,845]" pageId="2" pageNumber="386">Central</collectingCounty>
<collectingCountry id="F32CA59FFFFEFFFA106CFCD7FC56A91F" box="[786,923,819,845]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
,
<collectingMunicipality id="6BE07F75FFFEFFFA10DBFCD7FB46A91F" box="[933,1163,819,845]" pageId="2" pageNumber="386">Analamanga Region</collectingMunicipality>
[formerly
<collectingRegion id="49FF2BEDFFFEFFFA167AFCD7FF32A923" country="Madagascar" name="Antananarivo" pageId="2" pageNumber="386">Antananarivo province</collectingRegion>
],
<location id="8EE4B3D4FFFEFFFA126AFCB3FE24A923" LSID="urn:lsid:plazi:treatment:03925419FFFEFFFA13E9FE8AFE8CAD9F:8EE4B3D4FFFEFFFA126AFCB3FE24A923" box="[276,489,855,881]" country="Madagascar" county="Central" municipality="Analamanga Region" name="Ankazobe District" pageId="2" pageNumber="386" stateProvince="Antananarivo">Ankazobe District</location>
,
<location id="8EE4B3D4FFFEFFFA1287FCB3FDB1A923" LSID="urn:lsid:plazi:treatment:03925419FFFEFFFA13E9FE8AFE8CAD9F:8EE4B3D4FFFEFFFA1287FCB3FDB1A923" box="[505,636,855,881]" country="Madagascar" county="Central" municipality="Analamanga Region" name="Manankazo" pageId="2" pageNumber="386" stateProvince="Antananarivo">Manankazo</location>
env.,
<date id="FF85C3CFFFFEFFFA11B7FCB3FC85A923" box="[713,840,855,881]" pageId="2" pageNumber="386" value="2001-11-28">
<collectingDate id="EFC13A27FFFEFFFA11B7FCB3FC85A923" box="[713,840,855,881]" pageId="2" pageNumber="386" value="2001-11-28">28.11.2001</collectingDate>
</date>
,
<collectorName id="26CE80D9FFFEFFFA1027FCB3FC1EA923" box="[857,979,855,881]" pageId="2" pageNumber="386">Z. Mráček</collectorName>
leg.; (
<specimenCount id="9D3D2E86FFFEFFFA1759FCB3FB98A923" box="[1063,1109,855,881]" count="1" pageId="2" pageNumber="386" type="male">1 ♂</specimenCount>
FPC)
</materialsCitation>
:
<materialsCitation id="3B53EF52FFFEFFFA17CEFCB3FB00A9C7" collectorName="G. Dujardin-Delacour" country="Madagascar" county="Alaotra Mangora Region" location="Lakato" municipality="Moramanga District" pageId="2" pageNumber="386" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F32CA59FFFFEFFFA17CEFCB3FAF4A923" box="[1200,1337,855,881]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
,
<collectingCounty id="62E59D83FFFEFFFA1636FCB3FE94A9C7" pageId="2" pageNumber="386">Alaotra Mangora Region</collectingCounty>
,
<collectingMunicipality id="6BE07F75FFFEFFFA1218FC9FFD80A9C7" box="[358,589,891,917]" pageId="2" pageNumber="386">Moramanga District</collectingMunicipality>
,
<location id="8EE4B3D4FFFEFFFA1124FC9FFD66A9C7" LSID="urn:lsid:plazi:treatment:03925419FFFEFFFA13E9FE8AFE8CAD9F:8EE4B3D4FFFEFFFA1124FC9FFD66A9C7" box="[602,683,891,917]" country="Madagascar" county="Alaotra Mangora Region" municipality="Moramanga District" name="Lakato" pageId="2" pageNumber="386">Lakato</location>
, 1.1969,
<collectorName id="26CE80D9FFFEFFFA106BFC9FFBC1A9C7" box="[789,1036,891,917]" pageId="2" pageNumber="386">G. Dujardin-Delacour</collectorName>
leg.; (
<specimenCount id="9D3D2E86FFFEFFFA1724FC9FFB49A9C7" box="[1114,1156,891,917]" count="1" pageId="2" pageNumber="386" type="male">1 ♂</specimenCount>
FPC)
</materialsCitation>
:
<materialsCitation id="3B53EF52FFFEFFFA17A7FC9FFC85A98F" collectingDate="2011-11-29" collectorName="M. Tryzna" country="Madagascar" county="Central" elevation="1530" latitude="-18.197498" location="Ambohitantely Special Reserve" longLatPrecision="21" longitude="47.284164" municipality="Analamanga Region" pageId="2" pageNumber="386" specimenCount="1" specimenCount-male="1">
same as previous but 2. 1969; (
<specimenCount id="9D3D2E86FFFEFFFA1244FC7BFEA8A9EB" box="[314,357,927,953]" count="1" pageId="2" pageNumber="386" type="male">1 ♂</specimenCount>
IJC):
<collectingCounty id="62E59D83FFFEFFFA12CCFC7BFDC9A9EB" box="[434,516,927,953]" pageId="2" pageNumber="386">Central</collectingCounty>
<collectingCountry id="F32CA59FFFFEFFFA1170FC7BFD5AA9EB" box="[526,663,927,953]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
,
<collectingMunicipality id="6BE07F75FFFEFFFA11DAFC7BFC5DA9EB" box="[676,912,927,953]" pageId="2" pageNumber="386">Analamanga Region</collectingMunicipality>
,
<location id="8EE4B3D4FFFEFFFA10E2FC7BFAC5A9EB" LSID="urn:lsid:plazi:treatment:03925419FFFEFFFA13E9FE8AFE8CAD9F:8EE4B3D4FFFEFFFA10E2FC7BFAC5A9EB" box="[924,1288,927,953]" country="Madagascar" county="Central" latitude="-18.197498" longLatPrecision="21" longitude="47.284164" municipality="Analamanga Region" name="Ambohitantely Special Reserve" pageId="2" pageNumber="386">Ambohitantely Special Reserve</location>
,
<geoCoordinate id="EE0F83C8FFFEFFFA1669FC7AFA51A9EB" box="[1303,1436,926,953]" degrees="18" direction="south" minutes="11" orientation="latitude" pageId="2" pageNumber="386" precision="15" seconds="51" value="-18.197498">S18°1151</geoCoordinate>
<geoCoordinate id="EE0F83C8FFFEFFFA13E9FC27FEE9A98F" box="[151,292,963,989]" degrees="47" direction="east" minutes="17" orientation="longitude" pageId="2" pageNumber="386" precision="15" seconds="03" value="47.284164">E47°1703</geoCoordinate>
,
<quantity id="4CC348EAFFFEFFFA1251FC26FE4DA98F" box="[303,384,962,989]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="2" pageNumber="386" unit="m" value="1530.0">
<elevation id="0016023CFFFEFFFA1251FC26FE4DA98F" box="[303,384,962,989]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="2" pageNumber="386" unit="m" value="1530.0">1530m</elevation>
</quantity>
, at light, 24.
<date id="FF85C3CFFFFEFFFA1168FC27FD55A98F" box="[534,664,963,989]" pageId="2" pageNumber="386" value="2011-11-29">
<collectingDate id="EFC13A27FFFEFFFA1168FC27FD55A98F" box="[534,664,963,989]" pageId="2" pageNumber="386" value="2011-11-29">29.11.2011</collectingDate>
</date>
,
<collectorName id="26CE80D9FFFEFFFA11DDFC27FCDAA98F" box="[675,791,963,989]" pageId="2" pageNumber="386">M. Trýzna</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13B9FC03FD77AEC3" pageId="2" pageNumber="386" type="discussion">
<paragraph id="8B84E50FFFFEFFFA13B9FC03FD77AEC3" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FC03FEF6AE53" bold="true" box="[199,315,999,1025]" pageId="2" pageNumber="386">Remarks.</emphasis>
As
<bibRefCitation id="EFAA98FEFFFEFFFA1210FC03FE2AAE53" author="Pic, M." box="[366,487,999,1025]" pageId="2" pageNumber="386" pagination="1 - 16" refId="ref5365" refString="Pic, M. (1949) Clerides de Madagascar. Diversites Entomologiques V., 1949, 1 - 16." type="journal article" year="1949">Pic (1949)</bibRefCitation>
made no reference to specimens in his description it cannot be assumed that the nominal species-group name,
<taxonomicName id="4C3B9E8CFFFEFFFA1291FBEFFCC7AE77" authorityName="Pic" authorityYear="1949" box="[495,778,1035,1061]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA1291FBEFFCC7AE77" box="[495,778,1035,1061]" italics="true" pageId="2" pageNumber="386">Biflabelotillus foveolatus</emphasis>
</taxonomicName>
, was based on a single specimen. Despite being labelled holotype (a recent, not original, label) most likely by Menier the type located in the Muséum National dHistoire Naturelle, Paris (MNHN) must be considered a
<typeStatus id="54805BADFFFEFFFA11C3FBB7FCD9AE3F" box="[701,788,1107,1133]" pageId="2" pageNumber="386" type="syntype">syntype</typeStatus>
(
<bibRefCitation id="EFAA98FEFFFEFFFA105AFBB7FC60AE3F" author="ICZN" box="[804,941,1107,1133]" pageId="2" pageNumber="386" refId="ref5225" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. Adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Article 73.2) and is therefore available for
<typeStatus id="54805BADFFFEFFFA13E9FB93FF33AEC3" box="[151,254,1143,1169]" pageId="2" pageNumber="386" type="lectotype">lectotype</typeStatus>
designation (
<bibRefCitation id="EFAA98FEFFFEFFFA12EAFB93FDD6AEC3" author="ICZN" box="[404,539,1143,1169]" pageId="2" pageNumber="386" refId="ref5225" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. Adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Article 74.1).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13B9FB7EFEE8AF87" pageId="2" pageNumber="386" type="diagnosis">
<paragraph id="8B84E50FFFFEFFFA13B9FB7EFEE8AF87" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FB7EFE1CAEE6" bold="true" box="[199,465,1178,1204]" pageId="2" pageNumber="386">Differential diagnosis.</emphasis>
<taxonomicName id="4C3B9E8CFFFEFFFA12A2FB7FFDBFAEE7" authorityName="Pic" authorityYear="1949" box="[476,626,1179,1205]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA12A2FB7FFDBFAEE7" box="[476,626,1179,1205]" italics="true" pageId="2" pageNumber="386">B. foveolatus</emphasis>
</taxonomicName>
can easily be distinguished from both
<taxonomicName id="4C3B9E8CFFFEFFFA1743FB7FFB1FAEE6" authorityName="Menier" authorityYear="1979" box="[1085,1234,1179,1204]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA1743FB7FFB1FAEE6" box="[1085,1234,1179,1204]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3B9E8CFFFEFFFA166EFB7FFA51AEE7" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1296,1436,1179,1205]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA166EFB7FFA51AEE7" box="[1296,1436,1179,1205]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA13E9FB5BFF3CAE8B" bold="true" box="[151,241,1215,1241]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA13E9FB5BFF3CAE8B" box="[151,241,1215,1241]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by brown elytra (black in
<taxonomicName id="4C3B9E8CFFFEFFFA1160FB5BFD7DAE8A" authorityName="Menier" authorityYear="1979" box="[542,688,1215,1240]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA1160FB5BFD7DAE8A" box="[542,688,1215,1240]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
and bicoloured in
<taxonomicName id="4C3B9E8CFFFEFFFA10FAFB5BFBC3AE8B" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[900,1038,1215,1241]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA10FAFB5BFBC3AE8B" box="[900,1038,1215,1241]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA176BFB5BFBA2AE8B" bold="true" box="[1045,1135,1215,1241]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA176BFB5BFBA2AE8B" box="[1045,1135,1215,1241]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="1300F98AFFFEFFFA1701FB5BFB18AE8B" box="[1151,1237,1215,1241]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Figs 14</figureCitation>
,
<figureCitation id="1300F98AFFFEFFFA179FFB5BFACDAE8B" box="[1249,1280,1215,1241]" captionStart="FIGURE 2127" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[225,1371,187,1919]" captionTargetId="figure-17@6.[215,1371,181,1919]" captionTargetPageId="6" captionText="FIGURE 2127. Biflabelotillus peyrierasi Menier, 1979—2123 (holotype), 2427 (non-type, male). Fig. 21—labels of the holotype; 21, 24—habitus, dorsal view; 23, 25—habitus, lateral view; 26—scutellum; 27—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915941" httpUri="https://zenodo.org/record/13915941/files/figure.png" pageId="2" pageNumber="386">22</figureCitation>
,
<figureCitation id="1300F98AFFFEFFFA1672FB5BFAE1AE8B" box="[1292,1324,1215,1241]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="2" pageNumber="386">34</figureCitation>
). From
<taxonomicName id="4C3B9E8CFFFEFFFA16FAFB5BFEC4AEAE" authorityName="Menier" authorityYear="1979" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA16FAFB5BFEC4AEAE" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
it further differs by body length which is equal or larger than
<quantity id="4CC348EAFFFEFFFA10B7FB07FBD4AEAF" box="[969,1049,1251,1277]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="386" unit="mm" value="10.0">10 mm</quantity>
(less than
<quantity id="4CC348EAFFFEFFFA17EAFB07FB18AEAF" box="[1172,1237,1251,1277]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="386" unit="mm" value="9.0">9 mm</quantity>
in
<taxonomicName id="4C3B9E8CFFFEFFFA1784FB07FA41AEAE" authorityName="Menier" authorityYear="1979" box="[1274,1420,1251,1276]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA1784FB07FA41AEAE" box="[1274,1420,1251,1276]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
), subparallel, slightly widened elytra (parallel in
<taxonomicName id="4C3B9E8CFFFEFFFA11CBFAE3FC85AF72" authorityName="Menier" authorityYear="1979" box="[693,840,1287,1312]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA11CBFAE3FC85AF72" box="[693,840,1287,1312]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
), smooth elytral interstriae (the latter with tubercles in
<taxonomicName id="4C3B9E8CFFFEFFFA13C8FACFFE84AF16" authorityName="Menier" authorityYear="1979" box="[182,329,1323,1348]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA13C8FACFFE84AF16" box="[182,329,1323,1348]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
) and last but not least the shape of ultimate antennomere more or less securiform (elongate-oval in
<taxonomicName id="4C3B9E8CFFFEFFFA13C8FAABFE84AF3A" authorityName="Menier" authorityYear="1979" box="[182,329,1359,1384]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="peyrierasi">
<emphasis id="B94F391DFFFEFFFA13C8FAABFE84AF3A" box="[182,329,1359,1384]" italics="true" pageId="2" pageNumber="386">B. peyrierasi</emphasis>
</taxonomicName>
;
<figureCitation id="1300F98AFFFEFFFA1227FAABFE0DAF3B" box="[345,448,1358,1385]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="2" pageNumber="386">Figs 4, 5</figureCitation>
). From the new species
<taxonomicName id="4C3B9E8CFFFEFFFA11A6FAABFCAEAF3B" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[728,867,1359,1385]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA11A6FAABFCAEAF3B" box="[728,867,1359,1385]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA1012FAABFC0AAF3B" bold="true" box="[876,967,1359,1385]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA1012FAABFC0AAF3B" box="[876,967,1359,1385]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
it can also be distinguished by distinctly securiform ultimate antennomere (more or less securiform in
<taxonomicName id="4C3B9E8CFFFEFFFA1037FA97FC16AFDF" authorityName="Pic" authorityYear="1949" box="[841,987,1395,1421]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis id="B94F391DFFFEFFFA1037FA97FC16AFDF" box="[841,987,1395,1421]" italics="true" pageId="2" pageNumber="386">B. foveolatus</emphasis>
</taxonomicName>
; Figs 4,6), parallel-sided pronotum (not parallel in
<taxonomicName id="4C3B9E8CFFFEFFFA1273FA73FE5BAFE3" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[269,406,1431,1457]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA1273FA73FE5BAFE3" box="[269,406,1431,1457]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA12E2FA73FE38AFE3" bold="true" box="[412,501,1431,1457]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA12E2FA73FE38AFE3" box="[412,501,1431,1457]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="1300F98AFFFEFFFA117DFA73FDA9AFE3" box="[515,612,1431,1457]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="2" pageNumber="386">Figs 7, 9</figureCitation>
) and dark brown abdomen (distinctly pale-brown in
<taxonomicName id="4C3B9E8CFFFEFFFA17D3FA73FAFBAFE3" authorityName="Pavel &amp; Purchart" authorityYear="2024" box="[1197,1334,1431,1457]" class="Insecta" genus="Biflabelotillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="bicolorus" status="sp. nov.">
<emphasis id="B94F391DFFFEFFFA17D3FA73FAFBAFE3" box="[1197,1334,1431,1457]" italics="true" pageId="2" pageNumber="386">B. bicolorus</emphasis>
</taxonomicName>
<emphasis id="B94F391DFFFEFFFA1642FA73FA59AFE3" bold="true" box="[1340,1428,1431,1457]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="A27C8466FFFEFFFA1642FA73FA59AFE3" box="[1340,1428,1431,1457]" pageId="2" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="1300F98AFFFEFFFA13E9FA5FFF21AF87" box="[151,236,1467,1493]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Figs 18</figureCitation>
,
<figureCitation id="1300F98AFFFEFFFA1386FA5EFEDAAF87" box="[248,279,1466,1493]" captionStart="FIGURE 3337" captionStartId="8.[151,250,1679,1703]" captionTargetBox="[173,1409,286,1655]" captionTargetId="figure-47@8.[165,1422,273,1655]" captionTargetPageId="8" captionText="FIGURE 3337. Biflabelotillus bicolorus sp. nov., holotype. Fig. 33—habitus, lateral view; 34—habitus, dorsal view; 35— habitus, ventral view; 36—scutellum; 37—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915945" httpUri="https://zenodo.org/record/13915945/files/figure.png" pageId="2" pageNumber="386">35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13B9FA3AFAADADD7" pageId="2" pageNumber="386" type="description">
<paragraph id="8B84E50FFFFEFFFA13B9FA3AFF23AC4F" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9FA3AFEB8AFAA" bold="true" box="[199,373,1502,1528]" pageId="2" pageNumber="386">Redescription.</emphasis>
Size of
<typeStatus id="54805BADFFFEFFFA12A8FA3BFDF5AFAB" box="[470,568,1503,1529]" pageId="2" pageNumber="386" type="holotype">holotype</typeStatus>
10.0 x
<quantity id="4CC348EAFFFEFFFA11EFFA3BFD26AFAB" box="[657,747,1503,1529]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="386" unit="mm" value="2.4">2.4 mm</quantity>
. Winged, dark brown, with elytra and abdomen brown (
<figureCitation id="1300F98AFFFEFFFA1614FA3BFF2CAC4F" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Figs 1418</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F9C3FAF2AC37" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F9C3FEC3AC13" bold="true" box="[199,270,1575,1601]" pageId="2" pageNumber="386">Head.</emphasis>
Head including eyes broader than pronotum (DV) in ratio 1:0.8 (HW =
<quantity id="4CC348EAFFFEFFFA1756F9C3FB4CAC13" box="[1064,1153,1575,1601]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="386" unit="mm" value="2.1">2.1 mm</quantity>
). The space between eyes (frons) narrow (HW / FW = 3.5). Ultimate antennomere almost securifom, 3.4x longer then broad (
<figureCitation id="1300F98AFFFEFFFA1793F9AFFAFFAC37" box="[1261,1330,1611,1637]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="2" pageNumber="386">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F98BFC81ACFF" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F98BFE8AACDB" bold="true" box="[199,327,1647,1673]" pageId="2" pageNumber="386">Pronotum.</emphasis>
Longer than wide in ratio 1:1.7 (PL =
<quantity id="4CC348EAFFFEFFFA1070F98AFCA5ACDB" box="[782,872,1646,1673]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="386" unit="mm" value="2.5">2.5 mm</quantity>
, PW =
<quantity id="4CC348EAFFFEFFFA10BCF98AFBD2ACDB" box="[962,1055,1646,1673]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="386" unit="mm" value="1.5">1.5 mm</quantity>
), cylindrical, apical two thirds of pronotum parallel sided, basal third strongly trangled (
<figureCitation id="1300F98AFFFEFFFA1187F977FCF3ACFF" box="[761,830,1683,1709]" captionStart="FIGURE 112" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[202,1385,187,1917]" captionTargetId="figure-26@3.[202,1385,179,1917]" captionTargetPageId="3" captionText="FIGURE 112. Biflabelotillus foveolatus—1, 4, 7, 10; B. peyrierasi—2, 5, 8, 11; B. bicolorus sp. nov., holotype—3, 6, 9, 12. Fig. 13—right antenna, antennomers II-VI; 46—right antenna, antennomers IXXI (red arrows indicate last antennomere); 79—head and pronotum, dorsal view; 1012—head, front view." pageId="2" pageNumber="386">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F953FC25AD4B" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F953FED7AC83" bold="true" box="[199,282,1719,1745]" pageId="2" pageNumber="386">Elytra.</emphasis>
Subparallel, 2.6x longer than their width (EL =
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, EW =
<quantity id="4CC348EAFFFEFFFA1082F953FB95AC83" box="[1020,1112,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="386" unit="mm" value="2.4">2.4 mm</quantity>
), with posterior half slightly widening posteriad. Three fourths of elytral surface with bitubecrulate foveae, the latter absent in apical fourth (
<figureCitation id="1300F98AFFFEFFFA1611F93FFF2DAD4B" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Fig. 14, 17</figureCitation>
). Space between foveae approx. 13x larger than diameter of foveae.
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F8C6FB6DAD6F" blockId="2.[151,1437,602,1997]" box="[199,1184,1826,1853]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F8C6FE8BAD6E" bold="true" box="[199,326,1826,1852]" pageId="2" pageNumber="386">Scutellum.</emphasis>
Almost rounded with the basal margin distinctly cut-out in middle (
<figureCitation id="1300F98AFFFEFFFA1741F8C7FB5EAD6F" box="[1087,1171,1827,1853]" captionStart="FIGURE 1320" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[222,1365,181,1919]" captionTargetId="figure-17@4.[222,1365,181,1919]" captionTargetPageId="4" captionText="FIGURE 1320. Biflabelotillus foveolatus Pic, 1949—1315 (lectotype), 1620 (non-type). Fig. 13—labels of the lectotype; 14, 17—habitus, dorsal view; 15,16—habitus, lateral view; 18—habitus, ventral view; 19—scutellum; 20—bituberculate foveae (white arrows indicate the two tubercles on the top of the fovea)." figureDoi="http://doi.org/10.5281/zenodo.13915937" httpUri="https://zenodo.org/record/13915937/files/figure.png" pageId="2" pageNumber="386">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8B84E50FFFFEFFFA13B9F8A2FAADADD7" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F8A2FE6EAD32" bold="true" box="[199,419,1862,1888]" pageId="2" pageNumber="386">Variability of size.</emphasis>
Examined males (n=2): body length
<quantity id="4CC348EAFFFEFFFA1031F8A2FBCCAD33" box="[847,1025,1862,1889]" metricMagnitude="-2" metricUnit="m" metricValue="1.1824999999999999" metricValueMax="1.19" metricValueMin="1.175" pageId="2" pageNumber="386" unit="mm" value="11.825" valueMax="11.9" valueMin="11.75">11.7511.9 mm</quantity>
, HW =
<quantity id="4CC348EAFFFEFFFA1720F8A3FAC8AD33" box="[1118,1285,1863,1889]" metricMagnitude="-3" metricUnit="m" metricValue="2.525" metricValueMax="2.63" metricValueMin="2.42" pageId="2" pageNumber="386" unit="mm" value="2.525" valueMax="2.63" valueMin="2.42">2.422.63 mm</quantity>
, HW / FW = 2.993.31, PL =
<quantity id="4CC348EAFFFEFFFA122DF88EFE27ADD7" box="[339,490,1898,1925]" metricMagnitude="-3" metricUnit="m" metricValue="2.54" metricValueMax="2.58" metricValueMin="2.5" pageId="2" pageNumber="386" unit="mm" value="2.54" valueMax="2.58" valueMin="2.5">2.52.58 mm</quantity>
, PW =
<quantity id="4CC348EAFFFEFFFA1142F88FFD7DADD7" box="[572,688,1899,1925]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="2" pageNumber="386" unit="mm" value="1.9" valueMax="2.0" valueMin="1.8">1.82 mm</quantity>
, EL =
<quantity id="4CC348EAFFFEFFFA1185F88EFC5FADD7" box="[763,914,1898,1925]" metricMagnitude="-3" metricUnit="m" metricValue="8.475" metricValueMax="8.7" metricValueMin="8.25" pageId="2" pageNumber="386" unit="mm" value="8.475" valueMax="8.7" valueMin="8.25">8.258.7 mm</quantity>
, EW =
<quantity id="4CC348EAFFFEFFFA1098F88EFB41ADD7" box="[998,1164,1898,1925]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.55" metricValueMin="3.25" pageId="2" pageNumber="386" unit="mm" value="3.4" valueMax="3.55" valueMin="3.25">3.253.55 mm</quantity>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C321B684FFFEFFFA13B9F86AFE8CAD9F" pageId="2" pageNumber="386" type="distribution">
<paragraph id="8B84E50FFFFEFFFA13B9F86AFE8CAD9F" blockId="2.[151,1437,602,1997]" pageId="2" pageNumber="386">
<emphasis id="B94F391DFFFEFFFA13B9F86AFEACADFA" bold="true" box="[199,353,1934,1960]" pageId="2" pageNumber="386">Distribution.</emphasis>
Central and Eastern
<collectingCountry id="F32CA59FFFFEFFFA112AF86BFD16ADFB" box="[596,731,1935,1961]" name="Madagascar" pageId="2" pageNumber="386">Madagascar</collectingCountry>
Analamanga, Alaotra Mangora and Sava Regions (
<figureCitation id="1300F98AFFFEFFFA163EF86BFA59ADFB" box="[1344,1428,1935,1961]" captionStart="FIGURE 38" captionStartId="9.[151,250,1118,1142]" captionTargetBox="[489,1099,181,1094]" captionTargetId="figure-26@9.[489,1099,181,1094]" captionTargetPageId="9" captionText="FIGURE 38. Distribution map of the genus Biflabelotillus. B. foveolatus—red triangle, B. peyrierasi—white circle, B. bicolorus sp. nov.—yellow rhombus." figureDoi="http://doi.org/10.5281/zenodo.13915947" httpUri="https://zenodo.org/record/13915947/files/figure.png" pageId="2" pageNumber="386">Fig. 38</figureCitation>
) (
<bibRefCitation id="EFAA98FEFFFEFFFA13E1F857FEF9AD9F" author="Menier, J. J." box="[159,308,1971,1997]" pageId="2" pageNumber="386" pagination="75 - 77" refId="ref5316" refString="Menier, J. J. (1979) Une espece malgache inedite du genere Biflabellotillus: B. peyrierasi, n. sp. (Col. Cleridae). Revue franc ¸ aise d'Entomologie, New Series, 1 (8), 75 - 77." type="journal article" year="1979">Menier 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,59 +1,62 @@
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<treatment id="03FE887FFFE53008AB99F92B5D8F7300" LSID="urn:lsid:plazi:treatment:03FE887FFFE53008AB99F92B5D8F7300" httpUri="http://treatment.plazi.org/id/03FE887FFFE53008AB99F92B5D8F7300" lastPageId="4" lastPageNumber="300" pageId="0" pageNumber="296">
<subSubSection id="C34D6AE2FFE5300CAB99F92B580B7634" pageId="0" pageNumber="296" type="nomenclature">
<treatment id="03FE887FFFE53008AB99F92B5D8F7300" ID-DOI="http://doi.org/10.5281/zenodo.13916594" ID-GBIF-Taxon="243278326" ID-Zenodo-Dep="13916594" LSID="urn:lsid:plazi:treatment:03FE887FFFE53008AB99F92B5D8F7300" httpUri="http://treatment.plazi.org/id/03FE887FFFE53008AB99F92B5D8F7300" lastPageId="4" lastPageNumber="300" pageId="0" pageNumber="296">
<subSubSection id="C34D6AE2FFE5300CAB99F92B5B0A7611" box="[151,574,1773,1800]" pageId="0" pageNumber="296" type="nomenclature">
<paragraph id="8BE83969FFE5300CAB99F92B5B0A7611" blockId="0.[151,574,1773,1872]" box="[151,574,1773,1800]" pageId="0" pageNumber="296">
<heading id="D0A08E05FFE5300CAB99F92B5B0A7611" bold="true" box="[151,574,1773,1800]" fontSize="11" level="1" pageId="0" pageNumber="296" reason="1">
<emphasis id="B923E57BFFE5300CAB99F92B5B0A7611" bold="true" box="[151,574,1773,1800]" pageId="0" pageNumber="296">
<taxonomicName id="4C5742EAFFE5300CAB99F92B58ED7611" authorityName="Chen" authorityYear="2024" box="[151,473,1773,1800]" class="Insecta" family="Gryllacrididae" genus="Marthogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="xiangjini" status="sp. nov.">
<taxonomicName id="4C5742EAFFE5300CAB99F92B58ED7611" authority="Chen, 2024" authorityName="Chen" authorityYear="2024" box="[151,473,1773,1800]" class="Insecta" family="Gryllacrididae" genus="Marthogryllacris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="0" pageNumber="296" phylum="Arthropoda" rank="species" species="xiangjini" status="sp. nov.">
<emphasis id="B923E57BFFE5300CAB99F92B58ED7611" bold="true" box="[151,473,1773,1800]" italics="true" pageId="0" pageNumber="296">Marthogryllacris xiangjini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2105800FFE5300CAAEEF9285B0A7611" box="[480,574,1774,1800]" pageId="0" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D6AE2FFE5300CAB99F8D5580B7634" box="[151,319,1811,1837]" pageId="0" pageNumber="296" type="vernacular_names">
<paragraph id="8BE83969FFE5300CAB99F8D5580B7634" blockId="0.[151,574,1773,1872]" box="[151,319,1811,1837]" pageId="0" pageNumber="296">
<heading id="D0A08E05FFE5300CAB99F8D5580B7634" bold="true" box="[151,319,1811,1837]" fontSize="11" level="1" pageId="0" pageNumber="296" reason="1">
<emphasis id="B923E57BFFE5300CAB99F8D5580B7634" bold="true" box="[151,319,1811,1837]" pageId="0" pageNumber="296">ḂaeLjṖṞǞ</emphasis>
@ -209,9 +212,11 @@ BL:
<subSubSection id="C34D6AE2FFE13008ABC9FF1859FD7058" pageId="4" pageNumber="300" type="materials_examined">
<paragraph id="8BE83969FFE13008ABC9FF1859FD7058" blockId="4.[151,1437,150,537]" pageId="4" pageNumber="300">
<emphasis id="B923E57BFFE13008ABC9FF18588471E1" bold="true" box="[199,432,222,248]" pageId="4" pageNumber="300">Material examined.</emphasis>
<materialsCitation id="3B3F3334FFE13008AAB8FF195CAC71E0" box="[438,1432,223,249]" collectingDate="2024-08-20" location="Gongshan" municipality="Dulongjiang" pageId="4" pageNumber="300" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<materialsCitation id="3B3F3334FFE13008AAB8FF195CAC71E0" ID-GBIF-Occurrence="4951993301" box="[438,1432,223,249]" collectingDate="2024-08-20" collectorName="Xiangjin Liu" location="Gongshan" municipality="Dulongjiang" pageId="4" pageNumber="300" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54EC87CBFFE13008AAB8FF195B1771E0" box="[438,547,223,249]" pageId="4" pageNumber="300" type="holotype">Holotype</typeStatus>
: male,
:
<specimenCount id="9D51F2E0FFE13008A922FF195B5171E0" box="[556,613,223,249]" pageId="4" pageNumber="300" type="male">male</specimenCount>
,
<collectingMunicipality id="6B8CA313FFE13008A961FF195BCA71E0" box="[623,766,223,249]" pageId="4" pageNumber="300">Dulongjiang</collectingMunicipality>
,
<location id="8E886FB2FFE13008A807FF195A4971E0" LSID="urn:lsid:plazi:treatment:03FE887FFFE53008AB99F92B5D8F7300:8E886FB2FFE13008A807FF195A4971E0" box="[777,893,223,249]" municipality="Dulongjiang" name="Gongshan" pageId="4" pageNumber="300" stateProvince="Yunnan">Gongshan</location>
@ -221,17 +226,18 @@ BL:
<date id="FFE91FA9FFE13008A8E9FF195D9771E0" box="[999,1187,223,249]" pageId="4" pageNumber="300" value="2024-08-20">
<collectingDate id="EFADE641FFE13008A8E9FF195D9771E0" box="[999,1187,223,249]" pageId="4" pageNumber="300" value="2024-08-20">August 20, 2024</collectingDate>
</date>
, coll. by Xiangjin Liu
, coll. by
<collectorName id="26A25CBFFFE13008AE08FF195CAC71E0" box="[1286,1432,223,249]" pageId="4" pageNumber="300">Xiangjin Liu</collectorName>
</materialsCitation>
.
<materialsCitation id="3B3F3334FFE13008AB99FEC559F17058" collectingDate="2024-08-21" county="Dulongjiang" location="Ting Luo" municipality="Gongshan" pageId="4" pageNumber="300" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<materialsCitation id="3B3F3334FFE13008AB99FEC559F17058" ID-GBIF-Occurrence="4951993302" collectingDate="2024-08-21" collectorName="Ting Luo &amp; Yanting Qin" location="Dulongjiang" municipality="Gongshan" pageId="4" pageNumber="300" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54EC87CBFFE13008AB99FEC5583C7004" box="[151,264,259,285]" pageId="4" pageNumber="300" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D51F2E0FFE13008AA1DFEC5586B7004" box="[275,351,259,285]" count="1" pageId="4" pageNumber="300" type="male">1 male</specimenCount>
and
<specimenCount id="9D51F2E0FFE13008AA98FEC558CF7004" box="[406,507,259,285]" count="1" pageId="4" pageNumber="300" type="female">1 female</specimenCount>
,
<collectingCounty id="628941E5FFE13008A908FEC55BA17004" box="[518,661,259,285]" pageId="4" pageNumber="300">Dulongjiang</collectingCounty>
<location id="8E886FB2FFE13008A908FEC55BA17004" LSID="urn:lsid:plazi:treatment:03FE887FFFE53008AB99F92B5D8F7300:8E886FB2FFE13008A908FEC55BA17004" box="[518,661,259,285]" county="Dulongjiang" municipality="Gongshan" name="Dulongjiang" pageId="4" pageNumber="300" stateProvince="Yunnan">Dulongjiang</location>
,
<collectingMunicipality id="6B8CA313FFE13008A9AFFEC55A217004" box="[673,789,259,285]" pageId="4" pageNumber="300">Gongshan</collectingMunicipality>
,
@ -241,9 +247,9 @@ and
<collectingDate id="EFADE641FFE13008A88CFEC55D747004" box="[898,1088,259,285]" pageId="4" pageNumber="300" value="2024-08-21">August 21, 2024</collectingDate>
</date>
, coll. by
<location id="8E886FB2FFE13008AFA8FEC55C3B7004" LSID="urn:lsid:plazi:treatment:03FE887FFFE53008AB99F92B5D8F7300:8E886FB2FFE13008AFA8FEC55C3B7004" box="[1190,1295,259,285]" county="Dulongjiang" municipality="Gongshan" name="Ting Luo" pageId="4" pageNumber="300" stateProvince="Yunnan">Ting Luo</location>
<collectorName id="26A25CBFFFE13008AFA8FEC55C3B7004" box="[1190,1295,259,285]" pageId="4" pageNumber="300">Ting Luo</collectorName>
and
<location id="8E886FB2FFE13008AE4BFEC559F17058" LSID="urn:lsid:plazi:treatment:03FE887FFFE53008AB99F92B5D8F7300:8E886FB2FFE13008AE4BFEC559F17058" county="Dulongjiang" municipality="Gongshan" name="Yanting Qin" pageId="4" pageNumber="300" stateProvince="Yunnan">Yanting Qin</location>
<collectorName id="26A25CBFFFE13008AE4BFEC559F17058" pageId="4" pageNumber="300">Yanting Qin</collectorName>
</materialsCitation>
.
</paragraph>