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<document id="5A87900F7FF5CE840A4A0EFC4CB2A68F" ID-DOI="10.11646/zootaxa.5496.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="13346625" ID-ZooBank="D690291D-7BD2-43D1-95E8-81B3F82D988A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1724132869255" checkinUser="plazi" docAuthor="Takano, Hitoshi" docDate="2024" docId="592E8799FFD9FFCFFF4CF9ACB9865222" docLanguage="en" docName="zootaxa.5496.2.4.pdf" docOrigin="Zootaxa 5496 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5496.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthene sangha Takano, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="220" masterDocId="A517FFE1FFDDFFC9FFDBFF9AB952540F" masterDocTitle="Descriptions of three new species of Lycaenidae (Lepidoptera: Papilionoidea) from Parc National de Nouabalé-Ndoki, Republic of Congo with additional notes on little-known species" masterLastPageNumber="230" masterPageNumber="214" pageNumber="218" updateTime="1724133387488" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="950B49C1444C7A34A73B15C40C3D4AA2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="28754D861648E97D2C823751B69E49D3">
<mods:title id="27551AA18DF69525FFDE1584243DEBE4">Descriptions of three new species of Lycaenidae (Lepidoptera: Papilionoidea) from Parc National de Nouabalé-Ndoki, Republic of Congo with additional notes on little-known species</mods:title>
</mods:titleInfo>
<mods:name id="1B455585C44F47A97B5A4AFBC15C35BD" type="personal">
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<mods:namePart id="46FD9EC8482A719D79EED05D64391353">Takano, Hitoshi</mods:namePart>
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<mods:title id="2DB9186B5A9112166FED2D346592EA0E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A24BC6FF4EE309A3BB589F4F1B4A2D30">
<mods:date id="024914CDC2252659DC7A784EDF01FBEA">2024</mods:date>
<mods:detail id="AC8E5F1B2A26DB1BE7A61DDA9AE3DC33" type="pubDate">
<mods:number id="82F9BC68C3B11C3B46E0ABBA5016F659">2024-08-19</mods:number>
</mods:detail>
<mods:detail id="2F845B468C32FB1C97E73E8428863747" type="volume">
<mods:number id="85729C2C8833183A997F8B027E9CBED9">5496</mods:number>
</mods:detail>
<mods:detail id="47012AAF816D6C394598859DAFDA4A84" type="issue">
<mods:number id="7450A2063D87A1E1961A5816895D59C0">2</mods:number>
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<mods:url id="44EDB83F872CD4EBB3791E692ACD80F9">http://dx.doi.org/10.11646/zootaxa.5496.2.4</mods:url>
</mods:location>
<mods:classification id="07611FA73000129F8727378C6DB92330">journal article</mods:classification>
<mods:identifier id="40AEA8783783E2CF7DBBAA7BC32D3D67" type="DOI">10.11646/zootaxa.5496.2.4</mods:identifier>
<mods:identifier id="9ED092894149E386D3AE781EBCA8F1DF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="713AC5C1B127DACD5C7AE399D758AB4E" type="Zenodo-Dep">13346625</mods:identifier>
<mods:identifier id="952F6270C687526049208362D39D135D" type="ZooBank">D690291D-7BD2-43D1-95E8-81B3F82D988A</mods:identifier>
</mods:mods>
<treatment id="592E8799FFD9FFCFFF4CF9ACB9865222" LSID="urn:lsid:plazi:treatment:592E8799FFD9FFCFFF4CF9ACB9865222" httpUri="http://treatment.plazi.org/id/592E8799FFD9FFCFFF4CF9ACB9865222" lastPageId="6" lastPageNumber="220" pageId="4" pageNumber="218">
<subSubSection id="999D6504FFD9FFCDFF4CF9ACB890525E" box="[151,450,1590,1617]" pageId="4" pageNumber="218" type="nomenclature">
<paragraph id="D138368FFFD9FFCDFF4CF9ACB890525E" blockId="4.[151,450,1590,1653]" box="[151,450,1590,1617]" pageId="4" pageNumber="218">
<heading id="8A7081E3FFD9FFCDFF4CF9ACB890525E" bold="true" box="[151,450,1590,1617]" fontSize="11" level="1" pageId="4" pageNumber="218" reason="1">
<emphasis id="E3F3EA9DFFD9FFCDFF4CF9ACB890525E" bold="true" box="[151,450,1590,1617]" pageId="4" pageNumber="218">
<taxonomicName id="16874D0CFFD9FFCDFF4CF9ACB80F525E" authorityName="Takano" authorityYear="2024" box="[151,349,1590,1617]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="sangha" status="sp. nov.">
<emphasis id="E3F3EA9DFFD9FFCDFF4CF9ACB80F525E" bold="true" box="[151,349,1590,1617]" italics="true" pageId="4" pageNumber="218">Anthene sangha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8C057E6FFD9FFCDFEBEF9ADB890525E" box="[357,450,1591,1617]" pageId="4" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFD9FFCDFF4CF9C1B875527A" box="[151,295,1627,1653]" pageId="4" pageNumber="218" type="description">
<paragraph id="D138368FFFD9FFCDFF4CF9C1B875527A" blockId="4.[151,450,1590,1653]" box="[151,295,1627,1653]" pageId="4" pageNumber="218">
(
<figureCitation id="49BC2A0AFFD9FFCDFF44F9C1B84C527A" box="[159,286,1627,1653]" captionStart="FIGURES 9–14" captionStartId="5.[151,264,1486,1510]" captionTargetBox="[157,1434,184,1449]" captionTargetId="figure-30@5.[151,1435,181,1462]" captionTargetPageId="5" captionText="FIGURES 9–14. Anthene species. Adult ♂♂ (9–11), upperside (above) and underside (below); genitalia (12–14), lateral view (left), ventral view (right) and phallus (below). 9. A. levis (Hewitson), DRC, Mbata-Siala (ANHRTUK 00330135). 10. A. sangha sp. nov., holotype. 11. A. irumu (Stempffer), ROC, PNNN (ANHRTUK 00260783). 12. A. levis. 13. A. sangha. 14. A. irumu." figureDoi="http://doi.org/10.5281/zenodo.13346631" httpUri="https://zenodo.org/record/13346631/files/figure.png" pageId="4" pageNumber="218">Figs 10, 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFD9FFCDFF4CF939BBAC5342" pageId="4" pageNumber="218" type="materials_examined">
<paragraph id="D138368FFFD9FFCDFF4CF939B8F352B2" blockId="4.[151,1436,1699,1941]" box="[151,417,1699,1725]" pageId="4" pageNumber="218">
<materialsCitation id="61EF3CD2FFD9FFCDFF4CF939B8F352B2" box="[151,417,1699,1725]" collectionCode="ANHRT" pageId="4" pageNumber="218" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="E3F3EA9DFFD9FFCDFF4CF939B85652B2" bold="true" box="[151,260,1699,1725]" pageId="4" pageNumber="218">
<typeStatus id="0E3C882DFFD9FFCDFF4CF939B85652B2" box="[151,260,1699,1725]" pageId="4" pageNumber="218" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="C781FD06FFD9FFCDFED0F939B87252B2" box="[267,288,1699,1725]" count="1" pageId="4" pageNumber="218" type="male">♂</specimenCount>
(
<collectionCode id="B796AE4AFFD9FFCDFEF4F939B8C752B2" box="[303,405,1699,1725]" pageId="4" pageNumber="218">ANHRT</collectionCode>
):
</materialsCitation>
</paragraph>
<paragraph id="D138368FFFD9FFCDFF1CF95DBA4E5326" blockId="4.[151,1436,1699,1941]" pageId="4" pageNumber="218">
“
<materialsCitation id="61EF3CD2FFD9FFCDFF0AF95DBA4E5326" collectingDate="2022-10-01" collectingDateMax="2022-10-10" collectingDateMin="2022-10-01" collectionCode="ANHRT" country="Republic of the Congo" county="Mbeli" elevation="372" latitude="2.2399445" location="Sangha Prov." longLatPrecision="1" longitude="16.397806" municipality="National Park" pageId="4" pageNumber="218" specimenCount="1">
<collectingCountry id="A990761FFFD9FFCDFF0AF95DB8BB52EE" box="[209,489,1735,1761]" name="Republic of the Congo" pageId="4" pageNumber="218">REPUBLIC OF CONGO</collectingCountry>
<quantity id="167F9B6AFFD9FFCDFE37F95DBB7E52EE" box="[492,556,1735,1761]" metricMagnitude="2" metricUnit="m" metricValue="3.72" pageId="4" pageNumber="218" unit="m" value="372.0">
<elevation id="5AAAD1BCFFD9FFCDFE37F95DBB7E52EE" box="[492,556,1735,1761]" metricMagnitude="2" metricUnit="m" metricValue="3.72" pageId="4" pageNumber="218" unit="m" value="372.0">372m</elevation>
</quantity>
/
<location id="D4586054FFD9FFCDFDE1F95DBB9F52EE" LSID="urn:lsid:plazi:treatment:592E8799FFD9FFCFFF4CF9ACB9865222:D4586054FFD9FFCDFDE1F95DBB9F52EE" box="[570,717,1735,1761]" country="Republic of the Congo" county="Mbeli" latitude="2.2399445" longLatPrecision="1" longitude="16.397806" municipality="National Park" name="Sangha Prov." pageId="4" pageNumber="218">Sangha Prov.</location>
,
<location id="D4586054FFD9FFCDFD0FF95DBADD52EE" LSID="urn:lsid:plazi:treatment:592E8799FFD9FFCFFF4CF9ACB9865222:D4586054FFD9FFCDFD0FF95DBADD52EE" box="[724,911,1735,1761]" country="Republic of the Congo" county="Mbeli" latitude="2.2399445" longLatPrecision="1" longitude="16.397806" municipality="National Park" name="Nouabale-Ndoki" pageId="4" pageNumber="218">Nouabalé-Ndoki</location>
/
<collectingMunicipality id="315CACF5FFD9FFCDFC46F95DBD6552EE" box="[925,1079,1735,1761]" pageId="4" pageNumber="218">National Park</collectingMunicipality>
,
<collectingCounty id="38594E03FFD9FFCDFBE4F95DBDD052EE" box="[1087,1154,1735,1761]" pageId="4" pageNumber="218">Mbeli</collectingCounty>
camp/(
<taxonomicName id="16874D0CFFD9FFCDFB02F95DBCCE52EE" box="[1241,1436,1735,1761]" class="Magnoliopsida" family="Fabaceae" genus="Gilbertiodendron" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="218" phylum="Tracheophyta" rank="genus">
<emphasis id="E3F3EA9DFFD9FFCDFB02F95DBCCE52EE" box="[1241,1436,1735,1761]" italics="true" pageId="4" pageNumber="218">Gilbertiodendron</emphasis>
</taxonomicName>
forest) /
<geoCoordinate id="B4B35048FFD9FFCDFF22F971B8C8530A" box="[249,410,1771,1797]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="4" pageNumber="218" precision="1" seconds="23.8" value="2.2399445">02°14’23.8”N</geoCoordinate>
,
<geoCoordinate id="B4B35048FFD9FFCDFE7DF970BB13530A" box="[422,577,1770,1797]" degrees="16" direction="east" minutes="23" orientation="longitude" pageId="4" pageNumber="218" precision="1" seconds="52.1" value="16.397806">16°23’52.1”E</geoCoordinate>
/
<date id="A539104FFFD9FFCDFD81F971BBB4530A" box="[602,742,1771,1797]" pageId="4" pageNumber="218" value="2022-10-01" valueMax="2022-10-10" valueMin="2022-10-01">
<collectingDate id="B57DE9A7FFD9FFCDFD81F971BBB4530A" box="[602,742,1771,1797]" pageId="4" pageNumber="218" value="2022-10-01" valueMax="2022-10-10" valueMin="2022-10-01">1–10.x.2022</collectingDate>
</date>
<collectingMethod id="08C64E98FFD9FFCDFD35F971BACE530A" box="[750,924,1771,1797]" pageId="4" pageNumber="218">MV Light Trap</collectingMethod>
/ Dérozier, V., Fouka, B., / Kirk-Spriggs, A., Takano, H. /
<collectionCode id="B796AE4AFFD9FFCDFEFCF895B8DA5326" box="[295,392,1807,1833]" pageId="4" pageNumber="218">ANHRT</collectionCode>
:2022.14 // ANHRTUK / 00260662”
</materialsCitation>
</paragraph>
<paragraph id="D138368FFFD9FFCDFF1CF8A9BBAC5342" blockId="4.[151,1436,1699,1941]" box="[199,766,1843,1869]" pageId="4" pageNumber="218">
<emphasis id="E3F3EA9DFFD9FFCDFF1CF8A9B8665342" bold="true" box="[199,308,1843,1869]" pageId="4" pageNumber="218">
<typeStatus id="0E3C882DFFD9FFCDFF1CF8A9B8665342" box="[199,308,1843,1869]" pageId="4" pageNumber="218" type="paratype">Paratype</typeStatus>
</emphasis>
<specimenCount id="C781FD06FFD9FFCDFEE0F8A9B8015342" box="[315,339,1843,1869]" count="1" pageId="4" pageNumber="218" type="male">♂</specimenCount>
:
<materialsCitation id="61EF3CD2FFD9FFCDFE84F8A9BBA85342" box="[351,762,1843,1869]" collectionCode="ANHRT" pageId="4" pageNumber="218" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
same data as holotype (
<specimenCount id="C781FD06FFD9FFCDFDBDF8A9BBDB5342" box="[614,649,1843,1869]" count="1" pageId="4" pageNumber="218" type="male">1♂</specimenCount>
<collectionCode id="B796AE4AFFD9FFCDFD55F8A9BBA05342" box="[654,754,1843,1869]" pageId="4" pageNumber="218">ANHRT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFD9FFCFFF1CF8CCBD8C56A6" lastPageId="6" lastPageNumber="220" pageId="4" pageNumber="218" type="description">
<paragraph id="D138368FFFD9FFCDFF1CF8CCB80B537F" blockId="4.[151,1436,1699,1941]" box="[199,345,1878,1904]" pageId="4" pageNumber="218">
<emphasis id="E3F3EA9DFFD9FFCDFF1CF8CCB80B537F" bold="true" box="[199,345,1878,1904]" pageId="4" pageNumber="218">Description.</emphasis>
</paragraph>
<paragraph id="D138368FFFD9FFCDFF1CF8E1BACE539A" blockId="4.[151,1436,1699,1941]" box="[199,924,1915,1941]" pageId="4" pageNumber="218">
Forewing length:
<typeStatus id="0E3C882DFFD9FFCDFE55F8E1B8A2539A" box="[398,496,1915,1941]" pageId="4" pageNumber="218" type="holotype">holotype</typeStatus>
male:
<quantity id="167F9B6AFFD9FFCDFDE6F8E1BBDF539A" box="[573,653,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="218" unit="mm" value="11.0">11 mm</quantity>
;
<typeStatus id="0E3C882DFFD9FFCDFD42F8E1BBAB539A" box="[665,761,1915,1941]" pageId="4" pageNumber="218" type="paratype">paratype</typeStatus>
male:
<quantity id="167F9B6AFFD9FFCDFC9EF8E1BAC5539A" box="[837,919,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="218" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
<caption id="85F86607FFD8FFCCFF4CFA54BCD85221" ID-DOI="http://doi.org/10.5281/zenodo.13346631" ID-Zenodo-Dep="13346631" httpUri="https://zenodo.org/record/13346631/files/figure.png" pageId="5" pageNumber="219" startId="5.[151,264,1486,1510]" targetBox="[157,1434,184,1449]" targetPageId="5" targetType="figure">
<paragraph id="D138368FFFD8FFCCFF4CFA54BCD85221" blockId="5.[151,1436,1486,1582]" pageId="5" pageNumber="219">
<emphasis id="E3F3EA9DFFD8FFCCFF4CFA54B81A51E9" bold="true" box="[151,328,1486,1510]" pageId="5" pageNumber="219">FIGURES 9–14.</emphasis>
<taxonomicName id="16874D0CFFD8FFCCFE95FA54B8AB51E9" box="[334,505,1486,1510]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="219" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E3F3EA9DFFD8FFCCFE95FA54B8F351E9" box="[334,417,1486,1510]" italics="true" pageId="5" pageNumber="219">Anthene</emphasis>
species.
</taxonomicName>
Adult ♂♂ (9–11), upperside (above) and underside (below); genitalia (12–14), lateral view (left), ventral view (right) and phallus (below).
<emphasis id="E3F3EA9DFFD8FFCCFDB4FA68BBD35205" bold="true" box="[623,641,1522,1546]" pageId="5" pageNumber="219">9.</emphasis>
<taxonomicName id="16874D0CFFD8FFCCFD5CFA69BA1B5205" authority="(Hewitson)" baseAuthorityName="Hewitson" baseAuthorityYear="1878" box="[647,841,1522,1546]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="219" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="E3F3EA9DFFD8FFCCFD5CFA69BB9D5205" box="[647,719,1522,1546]" italics="true" pageId="5" pageNumber="219">A. levis</emphasis>
(Hewitson)
</taxonomicName>
, DRC, Mbata-Siala (ANHRTUK 00330135).
<emphasis id="E3F3EA9DFFD8FFCCFAC8FA68BC615205" bold="true" box="[1299,1331,1522,1546]" pageId="5" pageNumber="219">10.</emphasis>
<taxonomicName id="16874D0CFFD8FFCCFAE3FA69BCCE5205" authorityName="Takano" authorityYear="2024" box="[1336,1436,1522,1546]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="219" phylum="Arthropoda" rank="species" species="sangha" status="sp. nov.">
<emphasis id="E3F3EA9DFFD8FFCCFAE3FA69BCCE5205" box="[1336,1436,1522,1546]" italics="true" pageId="5" pageNumber="219">A. sangha</emphasis>
</taxonomicName>
<emphasis id="E3F3EA9DFFD8FFCCFF4CF98CB9BA5221" bold="true" box="[151,232,1558,1582]" pageId="5" pageNumber="219">
<taxonomicNameLabel id="F8C057E6FFD8FFCCFF4CF98CB9BA5221" box="[151,232,1558,1582]" pageId="5" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis id="E3F3EA9DFFD8FFCCFE81F98CB82B5221" bold="true" box="[346,377,1558,1582]" pageId="5" pageNumber="219">11.</emphasis>
<taxonomicName id="16874D0CFFD8FFCCFEA4F98DBB045221" authority="(Stempffer)" authorityName="Stempffer" authorityYear="1948" baseAuthorityName="Stempffer" baseAuthorityYear="1948" box="[383,598,1558,1582]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="219" phylum="Arthropoda" rank="species" species="irumu">
<emphasis id="E3F3EA9DFFD8FFCCFEA4F98DB88A5221" box="[383,472,1559,1582]" italics="true" pageId="5" pageNumber="219">A. irumu</emphasis>
(Stempffer)
</taxonomicName>
, ROC, PNNN (ANHRTUK 00260783).
<emphasis id="E3F3EA9DFFD8FFCCFC2BF98CBD5D5221" bold="true" box="[1008,1039,1558,1582]" pageId="5" pageNumber="219">12.</emphasis>
<taxonomicName id="16874D0CFFD8FFCCFBCDF98DBD325221" baseAuthorityName="Hewitson" baseAuthorityYear="1878" box="[1046,1120,1558,1582]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="219" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="E3F3EA9DFFD8FFCCFBCDF98DBD325221" box="[1046,1120,1558,1582]" italics="true" pageId="5" pageNumber="219">A. levis</emphasis>
</taxonomicName>
.
<emphasis id="E3F3EA9DFFD8FFCCFBB6F98CBDDF5221" bold="true" box="[1133,1165,1558,1582]" pageId="5" pageNumber="219">13.</emphasis>
<taxonomicName id="16874D0CFFD8FFCCFB48F98DBDAA5221" authorityName="Takano" authorityYear="2024" box="[1171,1272,1558,1582]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="219" phylum="Arthropoda" rank="species" species="sangha">
<emphasis id="E3F3EA9DFFD8FFCCFB48F98DBDAA5221" box="[1171,1272,1558,1582]" italics="true" pageId="5" pageNumber="219">A. sangha</emphasis>
</taxonomicName>
.
<emphasis id="E3F3EA9DFFD8FFCCFADEF98CBC775221" bold="true" box="[1285,1317,1558,1582]" pageId="5" pageNumber="219">14.</emphasis>
<taxonomicName id="16874D0CFFD8FFCCFAF0F98DBCD65221" authorityName="Stempffer" authorityYear="1948" baseAuthorityName="Stempffer" baseAuthorityYear="1948" box="[1323,1412,1559,1582]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="219" phylum="Arthropoda" rank="species" species="irumu">
<emphasis id="E3F3EA9DFFD8FFCCFAF0F98DBCD65221" box="[1323,1412,1559,1582]" italics="true" pageId="5" pageNumber="219">A. irumu</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="D138368FFFD8FFCFFF1CF91ABD8555FA" blockId="5.[151,1437,1664,2015]" lastBlockId="6.[151,1437,151,1581]" lastPageId="6" lastPageNumber="220" pageId="5" pageNumber="219">
Male. Upperside. Ground colour of head, thorax and abdomen black. Antennae black, with white scaling ventrally at the base of each segment. Ventral surface of club orange. Frons with a narrow row of white scales running adjacent to the eye. Palps longer than diameter of eye with white scaling. Abdominal segments 1–4 with white scaling along latero-posterior margin and along the entirety of the posterior margin of segments 5–8 giving the abdomen a banded appearance posteriorly. Ground colour of both wings dark brown; terminal line black; cilia greyish-brown. Forewing costa gently arcuate, rounded at apex; outer margin gently arcuate. The cell in its entirety, space 1 save for a
<quantity id="167F9B6AFFD8FFCCFEAAF8C3B8E6537C" box="[369,436,1881,1907]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="219" unit="mm" value="2.0">2 mm</quantity>
outer margin and the extreme base of space 2 are covered in violet-blue scales. Discal marking narrow, black. Discal and postdiscal area between veins 2–6 more sparsely covered in scales, forming a patch that differs in appearance from the rest of the wing. Hindwing arcuate, with three
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long, greyish-brown ‘tails’ protruding from the outer margin along veins 1–3. The cell, spaces 1–2 and the inner margin with long light-brown ciliate scales, in the latter turning to white towards the base. The entirety of the cell, the bases of spaces 4 and 5, and the majority of spaces 1a to 3 with the exception of a
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outer margin is covered in violet-blue scales. Discal marking ovoid, well-defined. The outer half of spaces 6 and 7 with sparse scaling similar to the forewing. Marginal spots, black, the one in space 2 larger than that in space 1a. Submarginal line violet-blue, well-defined, sinuate running from vein 1a to 4. Marginal line white, turning violet-blue and terminating in space 3. Underside. Head, thorax and abdomen covered in greyish-white scales. Legs black with a dusting of white scales, with a concentration of white scales at the base of each tarsal segment. Ground colour of both wings grey with indistinct and irregular discal and post-discal bands outlined in white and defined with dark brown on their inner margins but more heavily, almost forming spots, on the outer edge. Forewing submarginal and antemarginal lines white, sinuate, indistinct and interrupted at veins. Hindwing basal spot in space 7 dark brown, edged with white.
</paragraph>
<paragraph id="D138368FFFDBFFCFFF1CFE65BD02568A" blockId="6.[151,1437,151,1581]" pageId="6" pageNumber="220">Male genitalia. Uncus robust, broadly indented medially to form two lobes. Subuncus long and fine, mediallyangled, with hooked apex. Valve elongate, without ornamentation, strongly bifurcate at its apex forming two long, sharply-pointed processes, the dorsal process slightly longer than the other. Saccus long with evenly arcuate margin. Aedeagus almost three times as long as valve, distally narrower and tapering at apex.</paragraph>
<paragraph id="D138368FFFDBFFCFFF1CFD15BD8C56A6" blockId="6.[151,1437,151,1581]" box="[199,1246,655,681]" pageId="6" pageNumber="220">Female. No specimens examined (except for a photograph of a potential female—see below).</paragraph>
</subSubSection>
<subSubSection id="999D6504FFDBFFCFFF1CFD28B82650CA" pageId="6" pageNumber="220" type="diagnosis">
<paragraph id="D138368FFFDBFFCFFF1CFD28B82650CA" blockId="6.[151,1437,151,1581]" pageId="6" pageNumber="220">
<emphasis id="E3F3EA9DFFDBFFCFFF1CFD28B81056C3" bold="true" box="[199,322,690,716]" pageId="6" pageNumber="220">Diagnosis.</emphasis>
<taxonomicName id="16874D0CFFDBFFCFFE93FD29B8A956C2" authorityName="Takano" authorityYear="2024" box="[328,507,691,717]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="sangha" status="sp. nov.">
<emphasis id="E3F3EA9DFFDBFFCFFE93FD29B8A956C2" box="[328,507,691,717]" italics="true" pageId="6" pageNumber="220">Anthene sangha</emphasis>
</taxonomicName>
<emphasis id="E3F3EA9DFFDBFFCFFDD9FD29BB0956C2" bold="true" box="[514,603,691,717]" pageId="6" pageNumber="220">
<taxonomicNameLabel id="F8C057E6FFDBFFCFFDD9FD29BB0956C2" box="[514,603,691,717]" pageId="6" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
clearly belongs to the
<taxonomicName id="16874D0CFFDBFFCFFC83FD29BAF856C2" baseAuthorityName="Hewitson" baseAuthorityYear="1878" box="[856,938,691,717]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="E3F3EA9DFFDBFFCFFC83FD29BAF856C2" box="[856,938,691,717]" italics="true" pageId="6" pageNumber="220">A. levis</emphasis>
</taxonomicName>
group of
<bibRefCitation id="B5164B7EFFDBFFCFFBCCFD29BDFF56C2" author="Libert, M." box="[1047,1197,691,717]" pageId="6" pageNumber="220" refId="ref10860" refString="Libert, M. (2010) Revision des Anthene africains (Lepidoptera, Lycaenidae). ABRI, Nairobi, Kenya and Lambillionea, Tervuren, 420 pp." type="book" year="2010">Libert (2010)</bibRefCitation>
and is reminiscent of both
<taxonomicName id="16874D0CFFDBFFCFFF0AFD4DBB5656FE" authority="(Hewitson, [1878])" baseAuthorityName="Hewitson" baseAuthorityYear="1878" box="[209,516,727,753]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="E3F3EA9DFFDBFFCFFF0AFD4DB87656FE" box="[209,292,727,753]" italics="true" pageId="6" pageNumber="220">A. levis</emphasis>
(Hewitson, [1878])
</taxonomicName>
(
<figureCitation id="49BC2A0AFFDBFFCFFDC8FD4DBB0A56FE" box="[531,600,727,753]" captionStart="FIGURES 9–14" captionStartId="5.[151,264,1486,1510]" captionTargetBox="[157,1434,184,1449]" captionTargetId="figure-30@5.[151,1435,181,1462]" captionTargetPageId="5" captionText="FIGURES 9–14. Anthene species. Adult ♂♂ (9–11), upperside (above) and underside (below); genitalia (12–14), lateral view (left), ventral view (right) and phallus (below). 9. A. levis (Hewitson), DRC, Mbata-Siala (ANHRTUK 00330135). 10. A. sangha sp. nov., holotype. 11. A. irumu (Stempffer), ROC, PNNN (ANHRTUK 00260783). 12. A. levis. 13. A. sangha. 14. A. irumu." figureDoi="http://doi.org/10.5281/zenodo.13346631" httpUri="https://zenodo.org/record/13346631/files/figure.png" pageId="6" pageNumber="220">Fig. 9</figureCitation>
) and
<taxonomicName id="16874D0CFFDBFFCFFD43FD4DBA9856FE" authority="(Stempffer, 1948)" authorityName="Stempffer" authorityYear="1948" baseAuthorityName="Stempffer" baseAuthorityYear="1948" box="[664,970,727,753]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="irumu">
<emphasis id="E3F3EA9DFFDBFFCFFD43FD4DBBA956FF" box="[664,763,727,753]" italics="true" pageId="6" pageNumber="220">A. irumu</emphasis>
(
<bibRefCitation id="B5164B7EFFDBFFCFFCD1FD4DBA9056FE" author="Stempffer, H." box="[778,962,727,753]" pageId="6" pageNumber="220" pagination="185 - 196" refId="ref11419" refString="Stempffer, H. (1948) Contribution a l'etude des Lycaenidae de la faune ethiopienne. Revue Francaise d'Entomologie, 15, 185 - 196." type="journal article" year="1948">Stempffer, 1948</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="49BC2A0AFFDBFFCFFC01FD4DBD7956FE" box="[986,1067,727,753]" captionStart="FIGURES 9–14" captionStartId="5.[151,264,1486,1510]" captionTargetBox="[157,1434,184,1449]" captionTargetId="figure-30@5.[151,1435,181,1462]" captionTargetPageId="5" captionText="FIGURES 9–14. Anthene species. Adult ♂♂ (9–11), upperside (above) and underside (below); genitalia (12–14), lateral view (left), ventral view (right) and phallus (below). 9. A. levis (Hewitson), DRC, Mbata-Siala (ANHRTUK 00330135). 10. A. sangha sp. nov., holotype. 11. A. irumu (Stempffer), ROC, PNNN (ANHRTUK 00260783). 12. A. levis. 13. A. sangha. 14. A. irumu." figureDoi="http://doi.org/10.5281/zenodo.13346631" httpUri="https://zenodo.org/record/13346631/files/figure.png" pageId="6" pageNumber="220">Fig. 11</figureCitation>
) with which it is sympatric. It is closest to the former sharing the paler discal patch of the forewing, but the extent of the blue patch on the hindwing upperside in
<taxonomicName id="16874D0CFFDBFFCFFEF7FC85B8D25736" baseAuthorityName="Hewitson" baseAuthorityYear="1878" box="[300,384,799,825]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="E3F3EA9DFFDBFFCFFEF7FC85B8D25736" box="[300,384,799,825]" italics="true" pageId="6" pageNumber="220">A. levis</emphasis>
</taxonomicName>
is much greater whilst the discoidal marking is heavily reduced. The blue colouration of
<taxonomicName id="16874D0CFFDBFFCFFA5FFC85B9BA5752" authorityName="Takano" authorityYear="2024" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="sangha" status="sp. nov.">
<emphasis id="E3F3EA9DFFDBFFCFFA5FFC85B9BA5752" italics="true" pageId="6" pageNumber="220">A. sangha</emphasis>
</taxonomicName>
<emphasis id="E3F3EA9DFFDBFFCFFF2BFCD9B8195752" bold="true" box="[240,331,835,861]" pageId="6" pageNumber="220">
<taxonomicNameLabel id="F8C057E6FFDBFFCFFF2BFCD9B8195752" box="[240,331,835,861]" pageId="6" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is however less lustrous, more akin to the markings of
<taxonomicName id="16874D0CFFDBFFCFFC1AFCD9BD765753" authorityName="Stempffer" authorityYear="1948" baseAuthorityName="Stempffer" baseAuthorityYear="1948" box="[961,1060,835,861]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="irumu">
<emphasis id="E3F3EA9DFFDBFFCFFC1AFCD9BD765753" box="[961,1060,835,861]" italics="true" pageId="6" pageNumber="220">A. irumu</emphasis>
</taxonomicName>
. Male genitalia share similarities with both the aforementioned congeners in the elongate valve (as in
<taxonomicName id="16874D0CFFDBFFCFFC43FCFDBAA9578F" authorityName="Stempffer" authorityYear="1948" baseAuthorityName="Stempffer" baseAuthorityYear="1948" box="[920,1019,871,897]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="irumu">
<emphasis id="E3F3EA9DFFDBFFCFFC43FCFDBAA9578F" box="[920,1019,871,897]" italics="true" pageId="6" pageNumber="220">A. irumu</emphasis>
</taxonomicName>
(
<figureCitation id="49BC2A0AFFDBFFCFFBD1FCFDBD0F578E" box="[1034,1117,871,897]" captionStart="FIGURES 9–14" captionStartId="5.[151,264,1486,1510]" captionTargetBox="[157,1434,184,1449]" captionTargetId="figure-30@5.[151,1435,181,1462]" captionTargetPageId="5" captionText="FIGURES 9–14. Anthene species. Adult ♂♂ (9–11), upperside (above) and underside (below); genitalia (12–14), lateral view (left), ventral view (right) and phallus (below). 9. A. levis (Hewitson), DRC, Mbata-Siala (ANHRTUK 00330135). 10. A. sangha sp. nov., holotype. 11. A. irumu (Stempffer), ROC, PNNN (ANHRTUK 00260783). 12. A. levis. 13. A. sangha. 14. A. irumu." figureDoi="http://doi.org/10.5281/zenodo.13346631" httpUri="https://zenodo.org/record/13346631/files/figure.png" pageId="6" pageNumber="220">Fig. 14</figureCitation>
), although the configuration is fundamentally different) and its bifurcate apex (as in
<taxonomicName id="16874D0CFFDBFFCFFCD1FC11BA0E57AA" baseAuthorityName="Hewitson" baseAuthorityYear="1878" box="[778,860,907,933]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="E3F3EA9DFFDBFFCFFCD1FC11BA0E57AA" box="[778,860,907,933]" italics="true" pageId="6" pageNumber="220">A. levis</emphasis>
</taxonomicName>
). The genital capsule is however closest to
<taxonomicName id="16874D0CFFDBFFCFFA92FC11BCCE57AA" baseAuthorityName="Hewitson" baseAuthorityYear="1878" box="[1353,1436,907,933]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="E3F3EA9DFFDBFFCFFA92FC11BCCE57AA" box="[1353,1436,907,933]" italics="true" pageId="6" pageNumber="220">A. levis</emphasis>
</taxonomicName>
(
<figureCitation id="49BC2A0AFFDBFFCFFF44FC35B9A657C6" box="[159,244,943,969]" captionStart="FIGURES 9–14" captionStartId="5.[151,264,1486,1510]" captionTargetBox="[157,1434,184,1449]" captionTargetId="figure-30@5.[151,1435,181,1462]" captionTargetPageId="5" captionText="FIGURES 9–14. Anthene species. Adult ♂♂ (9–11), upperside (above) and underside (below); genitalia (12–14), lateral view (left), ventral view (right) and phallus (below). 9. A. levis (Hewitson), DRC, Mbata-Siala (ANHRTUK 00330135). 10. A. sangha sp. nov., holotype. 11. A. irumu (Stempffer), ROC, PNNN (ANHRTUK 00260783). 12. A. levis. 13. A. sangha. 14. A. irumu." figureDoi="http://doi.org/10.5281/zenodo.13346631" httpUri="https://zenodo.org/record/13346631/files/figure.png" pageId="6" pageNumber="220">Fig. 12</figureCitation>
) which differs from the new species in the smaller size (ca. two-thirds the size), the shallower medial emargination of the uncus, the shorter (ca. half the length) and broader (about twice the width) valves, the broader, shorter saccus and the phallus which is almost half the length. Photographs have been examined of another male specimen and a potential female from Dzanga-Sangha National Park,
<collectingCountry id="A990761FFFDBFFCFFC60FB81BAA7503A" box="[955,1013,1051,1077]" name="Central African Republic" pageId="6" pageNumber="220">CAR</collectingCountry>
in the collection of the late Philippe Annoyer (M. Libert, pers. comm.). This female specimen is similar in appearance to the female of
<taxonomicName id="16874D0CFFDBFFCFFB34FBA5BC105056" baseAuthorityName="Hewitson" baseAuthorityYear="1878" box="[1263,1346,1087,1113]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="levis">
<emphasis id="E3F3EA9DFFDBFFCFFB34FBA5BC105056" box="[1263,1346,1087,1113]" italics="true" pageId="6" pageNumber="220">A. levis</emphasis>
</taxonomicName>
sharing an all brown-upperside and a grey (rather than brown as in
<taxonomicName id="16874D0CFFDBFFCFFCF7FBF9BADC5073" authorityName="Stempffer" authorityYear="1948" baseAuthorityName="Stempffer" baseAuthorityYear="1948" box="[812,910,1123,1149]" class="Insecta" family="Lycaenidae" genus="Anthene" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="irumu">
<emphasis id="E3F3EA9DFFDBFFCFFCF7FBF9BADC5073" box="[812,910,1123,1149]" italics="true" pageId="6" pageNumber="220">A. irumu</emphasis>
</taxonomicName>
) underside but the dark brown discal markings are well-defined on both wings (like the male of the new species) and the antemarginal white band on the hindwing upperside is absent.
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFDBFFCFFF1CFB55BDAB5176" pageId="6" pageNumber="220" type="discussion">
<paragraph id="D138368FFFDBFFCFFF1CFB55BDAB5176" blockId="6.[151,1437,151,1581]" pageId="6" pageNumber="220">
<emphasis id="E3F3EA9DFFDBFFCFFF1CFB55B86950E6" bold="true" box="[199,315,1231,1257]" pageId="6" pageNumber="220">Remarks.</emphasis>
The
<typeStatus id="0E3C882DFFDBFFCFFEADFB55B88A50E6" box="[374,472,1231,1257]" pageId="6" pageNumber="220" type="holotype">holotype</typeStatus>
and
<typeStatus id="0E3C882DFFDBFFCFFDCBFB55BB2250E6" box="[528,624,1231,1257]" pageId="6" pageNumber="220" type="paratype">paratype</typeStatus>
of the new species were both captured on the same night at a MV light trap at Mbeli camp (
<figureCitation id="49BC2A0AFFDBFFCFFEA3FB69B8985102" box="[376,458,1267,1293]" captionStart="FIGURES 38–40" captionStartId="12.[151,264,1214,1238]" captionTargetBox="[151,1436,181,1190]" captionTargetId="figure-15@12.[151,1436,181,1190]" captionTargetPageId="12" captionText="FIGURES 38–40.Parc National de Nouabalé-Ndoki, Republic of Congo.38.Mbeli camp, typelocality of Falcuna nouabaleensis sp. nov. and Anthene sangha sp. nov. 39. Ndoki formation, type locality of Neurellipes smithi sp. nov. 40. F. nouabaleensis sp. nov., paratype ♂, taken in natura at Mondika camp. (All photographs V. Dérozier)." figureDoi="http://doi.org/10.5281/zenodo.13346643" httpUri="https://zenodo.org/record/13346643/files/figure.png" pageId="6" pageNumber="220">Fig. 38</figureCitation>
), within a small clearing in the forest. The specimens were attracted to the light during a very heavy downpour together with numerous other butterfly species belonging to various families, presumably all of the individuals having been disturbed by the strong winds and rain. Numerous other butterflies were collected at light traps throughout the time in the field, of which several were unique to this sampling technique.
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFDBFFCFFF1CFA19BBD551CE" pageId="6" pageNumber="220" type="etymology">
<paragraph id="D138368FFFDBFFCFFF1CFA19BBD551CE" blockId="6.[151,1437,151,1581]" pageId="6" pageNumber="220">
<emphasis id="E3F3EA9DFFDBFFCFFF1CFA19B8CF5192" bold="true" box="[199,413,1411,1437]" pageId="6" pageNumber="220">
<emphasis id="E3F3EA9DFFDBFFCFFF1CFA19B8C45192" bold="true" box="[199,406,1411,1437]" italics="true" pageId="6" pageNumber="220">Derivatio nominis</emphasis>
.
</emphasis>
The new species is named after the Sangha River which flows through the region where the only known records of the species originate.
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFDBFFCFFF1CFA50B9865222" pageId="6" pageNumber="220" type="distribution">
<paragraph id="D138368FFFDBFFCFFF1CFA50B9865222" blockId="6.[151,1437,151,1581]" pageId="6" pageNumber="220">
<emphasis id="E3F3EA9DFFDBFFCFFF1CFA50B83351EB" bold="true" box="[199,353,1482,1508]" pageId="6" pageNumber="220">Distribution.</emphasis>
The new species is so far only known with certainty from
<specimenCount id="C781FD06FFDBFFCFFBC1FA51BDC051EA" box="[1050,1170,1483,1509]" count="3" pageId="6" pageNumber="220" type="male">three male</specimenCount>
specimens captured in PNNN and Dzanga-Sangha National Park,
<collectingCountry id="A990761FFFDBFFCFFD53FA75BB965206" box="[648,708,1519,1545]" name="Central African Republic" pageId="6" pageNumber="220">CAR</collectingCountry>
, and it is potentially endemic to the Sangha-Congo interfluvial zone.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:roleTerm id="CB96EF59714998F5AE6FD182E8A8FE75">Author</mods:roleTerm>
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<mods:namePart id="CB0CB0BF8737DEA8F760FAFBEAF86190">Takano, Hitoshi</mods:namePart>
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<mods:title id="4DB6B9764F7F7B5EA2E938C4C99D472E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E943D30F6B65D4C98472FCCBD449DD38">
<mods:date id="05FFC73567BBCD6C8AC887784A7BAB8E">2024</mods:date>
<mods:detail id="6BD53743291DF994F068D844720DEEA3" type="pubDate">
<mods:number id="658576531FBA8F1383C13CB07E7A6C84">2024-08-19</mods:number>
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<mods:number id="FA1AFDF94F60D3F247D2CECA5711F0AC">5496</mods:number>
</mods:detail>
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<mods:number id="E4F855E50D9248F5DE810E1CBE539146">2</mods:number>
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<mods:start id="38FD329B6BD372737ED0D3BD39F5E970">214</mods:start>
<mods:end id="3373C253B998BAC536BC79D281594EB5">230</mods:end>
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<mods:location id="915EA9C787FCEFA32E0827B9A2C2ABF6">
<mods:url id="654C126B404C68B3ACC9866EEAC2C968">http://dx.doi.org/10.11646/zootaxa.5496.2.4</mods:url>
</mods:location>
<mods:classification id="09ECF3D1C51CC060D439A3BED163CCB7">journal article</mods:classification>
<mods:identifier id="8E358A92C74C943DDF771210ADB3EA18" type="DOI">10.11646/zootaxa.5496.2.4</mods:identifier>
<mods:identifier id="01CEFFAA3C80CA44BF89173B619D0A9A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B5CB2A599956BD0233A82B234277916D" type="Zenodo-Dep">13346625</mods:identifier>
<mods:identifier id="51CCCDCAB7D51A1CCF40B1F1ECE4271F" type="ZooBank">D690291D-7BD2-43D1-95E8-81B3F82D988A</mods:identifier>
</mods:mods>
<treatment id="592E8799FFDBFFC0FF4CF9E4B8E95206" LSID="urn:lsid:plazi:treatment:592E8799FFDBFFC0FF4CF9E4B8E95206" httpUri="http://treatment.plazi.org/id/592E8799FFDBFFC0FF4CF9E4B8E95206" lastPageId="9" lastPageNumber="223" pageId="6" pageNumber="220">
<subSubSection id="999D6504FFDBFFCFFF4CF9E4B88B5296" box="[151,473,1662,1689]" pageId="6" pageNumber="220" type="nomenclature">
<paragraph id="D138368FFFDBFFCFFF4CF9E4B88B5296" blockId="6.[151,473,1662,1725]" box="[151,473,1662,1689]" pageId="6" pageNumber="220">
<heading id="8A7081E3FFDBFFCFFF4CF9E4B88B5296" bold="true" box="[151,473,1662,1689]" fontSize="11" level="1" pageId="6" pageNumber="220" reason="1">
<emphasis id="E3F3EA9DFFDBFFCFFF4CF9E4B88B5296" bold="true" box="[151,473,1662,1689]" pageId="6" pageNumber="220">
<taxonomicName id="16874D0CFFDBFFCFFF4CF9E4B8265296" authorityName="Takano" authorityYear="2024" box="[151,372,1662,1689]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="220" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="E3F3EA9DFFDBFFCFFF4CF9E4B8265296" bold="true" box="[151,372,1662,1689]" italics="true" pageId="6" pageNumber="220">Neurellipes smithi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8C057E6FFDBFFCFFEA0F9E5B88B5296" box="[379,473,1663,1689]" pageId="6" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFDBFFCFFF4CF939B80352B2" box="[151,337,1699,1725]" pageId="6" pageNumber="220" type="description">
<paragraph id="D138368FFFDBFFCFFF4CF939B80352B2" blockId="6.[151,473,1662,1725]" box="[151,337,1699,1725]" pageId="6" pageNumber="220">
(
<figureCitation id="49BC2A0AFFDBFFCFFF44F939B87252B2" box="[159,288,1699,1725]" captionStart="FIGURES 15–24" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[158,1432,382,1805]" captionTargetId="figure-109@7.[151,1435,381,1818]" captionTargetPageId="7" captionText="FIGURES 15–24. Neurellipes species. Adults, upperside (above) and underside (below). 15. N. ngoko ngoko (Stempffer), ♂, ROC, PNNN (ANHRTUK 00321840). 16. Id. (ANHRTUK 00260807). 17. Id. (ANHRTUK 00260771). 18. Id., ♀, ROC, PNNN (ANHRTUK 00260682). 19. N. smithi sp. nov., paratype ♂ (ANHRTUK 00289864). 20. Id., holotype ♂. 21. Id., paratype ♂ (ANHRTUK 00260696). 22. Id., paratype ♀ (ANHRTUK 00321906). 23. N. cf. ngoko, ♂, DRC, Kwokoro (ANHRTUK 00289939). 24. N. rufomarginata (Bethune-Baker), ♂, DRC, Mamove (ANHRTUK 00363939)." figureDoi="http://doi.org/10.5281/zenodo.13346635" httpUri="https://zenodo.org/record/13346635/files/figure.png" pageId="6" pageNumber="220">Figs 19–22</figureCitation>
,
<figureCitation id="49BC2A0AFFDBFFCFFEF7F939B81B52B2" box="[300,329,1699,1725]" captionStart="FIGURES 25–27" captionStartId="8.[151,264,854,878]" captionTargetBox="[152,1435,184,825]" captionTargetId="figure-15@8.[151,1436,181,830]" captionTargetPageId="8" captionText="FIGURES 25–27. Neurellipes species ♂ genitalia. Capsule in lateral view (top), phallus (middle) and valve in three-quarter view (bottom). 25. N. ngoko ngoko (Stempffer), ROC, PNNN (ANHRTUK 00260768). 26. N. smithi sp. nov., holotype. 27. N. rufomarginata (Bethune-Baker), DRC, Mamove (ANHRTUK 00363939)." figureDoi="http://doi.org/10.5281/zenodo.13346637" httpUri="https://zenodo.org/record/13346637/files/figure.png" pageId="6" pageNumber="220">26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFDBFFCEFF4CF971BA3E556A" lastPageId="7" lastPageNumber="221" pageId="6" pageNumber="220" type="materials_examined">
<paragraph id="D138368FFFDBFFCFFF4CF971B8F3530A" blockId="6.[151,1437,1771,2013]" box="[151,417,1771,1797]" pageId="6" pageNumber="220">
<materialsCitation id="61EF3CD2FFDBFFCFFF4CF971B8F3530A" box="[151,417,1771,1797]" collectionCode="ANHRT" pageId="6" pageNumber="220" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="E3F3EA9DFFDBFFCFFF4CF971B856530A" bold="true" box="[151,260,1771,1797]" pageId="6" pageNumber="220">
<typeStatus id="0E3C882DFFDBFFCFFF4CF971B856530A" box="[151,260,1771,1797]" pageId="6" pageNumber="220" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="C781FD06FFDBFFCFFED0F971B872530A" box="[267,288,1771,1797]" count="1" pageId="6" pageNumber="220" type="male">♂</specimenCount>
(
<collectionCode id="B796AE4AFFDBFFCFFEF4F971B8C7530A" box="[303,405,1771,1797]" pageId="6" pageNumber="220">ANHRT</collectionCode>
):
</materialsCitation>
</paragraph>
<paragraph id="D138368FFFDBFFCFFF1CF895BA4E537E" blockId="6.[151,1437,1771,2013]" pageId="6" pageNumber="220">
“
<collectingCountry id="A990761FFFDBFFCFFF0AF895B8BF5326" box="[209,493,1807,1833]" name="Republic of the Congo" pageId="6" pageNumber="220">REPUBLIC OF CONGO</collectingCountry>
<quantity id="167F9B6AFFDBFFCFFE29F894BB605326" box="[498,562,1806,1833]" metricMagnitude="2" metricUnit="m" metricValue="3.52" pageId="6" pageNumber="220" unit="m" value="352.0">352m</quantity>
/ Sangha Prov., Nouabalé-Ndoki / National Park, Ndoki formation / (secondary forest) /
<geoCoordinate id="B4B35048FFDBFFCFFF2DF8A9B8C55342" box="[246,407,1843,1869]" degrees="02" direction="north" minutes="12" orientation="latitude" pageId="6" pageNumber="220" precision="1" seconds="47.7" value="2.2132502">02°12’47.7”N</geoCoordinate>
,
<geoCoordinate id="B4B35048FFDBFFCFFE79F8A8BB6F5342" box="[418,573,1842,1869]" degrees="16" direction="east" minutes="23" orientation="longitude" pageId="6" pageNumber="220" precision="1" seconds="45.8" value="16.396055">16°23’45.8”E</geoCoordinate>
/
<date id="A539104FFFDBFFCFFD88F8A9BA585342" box="[595,778,1843,1869]" pageId="6" pageNumber="220" value="2022-09-29" valueMax="2022-10-01" valueMin="2022-09-29">29.ix–01.x.2022</date>
General Coll. / Dérozier, V., Fouka, B., / Kirk-Spriggs, A., Takano, H. /
<collectionCode id="B796AE4AFFDBFFCFFEFCF8CDB8DA537E" box="[295,392,1879,1905]" pageId="6" pageNumber="220">ANHRT</collectionCode>
:2022.14 // ANHRTUK / 00260669”
</paragraph>
<paragraph id="D138368FFFDBFFCFFF1CF8E1B8B4539A" blockId="6.[151,1437,1771,2013]" box="[199,486,1915,1941]" pageId="6" pageNumber="220">
<emphasis id="E3F3EA9DFFDBFFCFFF1CF8E1B86D539A" bold="true" box="[199,319,1915,1941]" pageId="6" pageNumber="220">
<typeStatus id="0E3C882DFFDBFFCFFF1CF8E1B86D539A" box="[199,319,1915,1941]" pageId="6" pageNumber="220" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="C781FD06FFDBFFCFFE95F8E1B8C7539A" box="[334,405,1915,1941]" count="13" pageId="6" pageNumber="220" type="male">13♂♂</specimenCount>
<specimenCount id="C781FD06FFDBFFCFFE47F8E1B884539A" box="[412,470,1915,1941]" count="2" pageId="6" pageNumber="220" type="female">2♀♀</specimenCount>
):
</paragraph>
<paragraph id="D138368FFFDBFFCEFF1CF804BA3E556A" blockId="6.[151,1437,1771,2013]" lastBlockId="7.[151,1437,151,357]" lastPageId="7" lastPageNumber="221" pageId="6" pageNumber="220">
<materialsCitation id="61EF3CD2FFDBFFCFFF1CF804BAF653B6" box="[199,932,1950,1977]" collectionCode="ANHRT" country="Republic of the Congo" pageId="6" pageNumber="220" specimenCount="2" specimenCount-male="2">
<emphasis id="E3F3EA9DFFDBFFCFFF1CF804B8A053B7" bold="true" box="[199,498,1950,1976]" pageId="6" pageNumber="220">
<collectingCountry id="A990761FFFDBFFCFFF1CF804B8A053B7" box="[199,498,1950,1976]" name="Republic of the Congo" pageId="6" pageNumber="220">REPUBLIC OF CONGO</collectingCountry>
</emphasis>
: same data as holotype (
<specimenCount id="C781FD06FFDBFFCFFD27F805BA6753B6" box="[764,821,1951,1977]" count="2" pageId="6" pageNumber="220" type="male">2♂♂</specimenCount>
<collectionCode id="B796AE4AFFDBFFCFFCE3F805BACE53B6" box="[824,924,1951,1977]" pageId="6" pageNumber="220">ANHRT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="61EF3CD2FFDBFFCEFC76F805B85154BE" collectingDate="2022-10-01" collectingDateMax="2022-10-10" collectingDateMin="2022-10-01" collectionCode="ANHRT" collectorName="V. Derozier &amp; B. Fouka &amp; A. Kirk-Spriggs &amp; H. Takano" country="Republic of the Congo" elevation="372" lastPageId="7" lastPageNumber="221" latitude="2.2399445" location="Mbeli" longLatPrecision="1" longitude="16.397806" municipality="Nouabale-Ndoki National Park" pageId="6" pageNumber="220" specimenCount="2" specimenCount-male="2">
<location id="D4586054FFDBFFCFFC76F805BAA253B6" LSID="urn:lsid:plazi:treatment:592E8799FFDBFFC0FF4CF9E4B8E95206:D4586054FFDBFFCFFC76F805BAA253B6" box="[941,1008,1951,1977]" country="Republic of the Congo" latitude="2.2399445" longLatPrecision="1" longitude="16.397806" municipality="Nouabale-Ndoki National Park" name="Mbeli" pageId="6" pageNumber="220">Mbeli</location>
camp,
<collectingMunicipality id="315CACF5FFDBFFCFFBE6F805BCCA53B6" box="[1085,1432,1951,1977]" pageId="6" pageNumber="220">Nouabalé-Ndoki National Park</collectingMunicipality>
,
<quantity id="167F9B6AFFDBFFCFFF4CF859B98853D2" box="[151,218,1987,2013]" metricMagnitude="2" metricUnit="m" metricValue="3.72" pageId="6" pageNumber="220" unit="m" value="372.0">
<elevation id="5AAAD1BCFFDBFFCFFF4CF859B98853D2" box="[151,218,1987,2013]" metricMagnitude="2" metricUnit="m" metricValue="3.72" pageId="6" pageNumber="220" unit="m" value="372.0">372m</elevation>
</quantity>
,
<geoCoordinate id="B4B35048FFDBFFCFFF3EF859B8D453D2" box="[229,390,1987,2013]" degrees="02" direction="north" minutes="14" orientation="latitude" pageId="6" pageNumber="220" precision="1" seconds="23.8" value="2.2399445">02°14’23.8”N</geoCoordinate>
,
<geoCoordinate id="B4B35048FFDBFFCFFE49F858BB6253D2" box="[402,560,1986,2013]" degrees="16" direction="east" minutes="23" orientation="longitude" pageId="6" pageNumber="220" precision="1" seconds="52.1" value="16.397806">16°23’52.1”E</geoCoordinate>
,
<date id="A539104FFFDBFFCFFDE0F859BB9853D2" box="[571,714,1987,2013]" pageId="6" pageNumber="220" value="2022-10-01" valueMax="2022-10-10" valueMin="2022-10-01">
<collectingDate id="B57DE9A7FFDBFFCFFDE0F859BB9853D2" box="[571,714,1987,2013]" pageId="6" pageNumber="220" value="2022-10-01" valueMax="2022-10-10" valueMin="2022-10-01">1–10.x.2022</collectingDate>
</date>
, leg.
<collectorName id="7C725359FFDBFFCFFCDEF859BADA53D2" box="[773,904,1987,2013]" pageId="6" pageNumber="220">V. Dérozier</collectorName>
,
<collectorName id="7C725359FFDBFFCFFC48F859BAAF53D2" box="[915,1021,1987,2013]" pageId="6" pageNumber="220">B. Fouka</collectorName>
,
<collectorName id="7C725359FFDBFFCFFBDCF859BDE953D2" box="[1031,1211,1987,2013]" pageId="6" pageNumber="220">A. Kirk-Spriggs</collectorName>
&amp;
<collectorName id="7C725359FFDBFFCFFB3BF859BC0653D2" box="[1248,1364,1987,2013]" pageId="6" pageNumber="220">H. Takano</collectorName>
(
<specimenCount id="C781FD06FFDBFFCFFAB8F859BCCF53D2" box="[1379,1437,1987,2013]" count="2" pageId="6" pageNumber="220" type="male">2♂♂</specimenCount>
<collectionCode id="B796AE4AFFDAFFCEFF4CFF0DB9A954BE" box="[151,251,151,177]" pageId="7" pageNumber="221">ANHRT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="61EF3CD2FFDAFFCEFED6FF0DBD4154DA" collectingDate="2023-05-06" collectingDateMax="2023-05-14" collectingDateMin="2023-05-06" collectionCode="ANHRT" collectorName="V. Derozier &amp; A. Kirk-Spriggs &amp; G. Laszlo &amp; S. Mvouende" country="Republic of the Congo" elevation="341" latitude="2.2102501" location="Bomassa" longLatPrecision="1" longitude="16.191721" municipality="Nouabale-Ndoki National Park" pageId="7" pageNumber="221" specimenCount="5" specimenCount-female="2" specimenCount-male="3">
<location id="D4586054FFDAFFCEFED6FF0DB82054BE" LSID="urn:lsid:plazi:treatment:592E8799FFDBFFC0FF4CF9E4B8E95206:D4586054FFDAFFCEFED6FF0DB82054BE" box="[269,370,151,177]" country="Republic of the Congo" latitude="2.2102501" longLatPrecision="1" longitude="16.191721" municipality="Nouabale-Ndoki National Park" name="Bomassa" pageId="7" pageNumber="221">Bomassa</location>
camp,
<collectingMunicipality id="315CACF5FFDAFFCEFE19FF0DBA4D54BE" box="[450,799,151,177]" pageId="7" pageNumber="221">Nouabalé-Ndoki National Park</collectingMunicipality>
,
<quantity id="167F9B6AFFDAFFCEFCF2FF0DBA3954BE" box="[809,875,151,177]" metricMagnitude="2" metricUnit="m" metricValue="3.41" pageId="7" pageNumber="221" unit="m" value="341.0">
<elevation id="5AAAD1BCFFDAFFCEFCF2FF0DBA3954BE" box="[809,875,151,177]" metricMagnitude="2" metricUnit="m" metricValue="3.41" pageId="7" pageNumber="221" unit="m" value="341.0">341m</elevation>
</quantity>
,
<geoCoordinate id="B4B35048FFDAFFCEFCADFF0DBD4554BE" box="[886,1047,151,177]" degrees="02" direction="north" minutes="12" orientation="latitude" pageId="7" pageNumber="221" precision="1" seconds="36.9" value="2.2102501">02°12’36.9”N</geoCoordinate>
,
<geoCoordinate id="B4B35048FFDAFFCEFBFAFF0DBDEC54BE" box="[1057,1214,151,177]" degrees="16" direction="east" minutes="11" orientation="longitude" pageId="7" pageNumber="221" precision="1" seconds="30.2" value="16.191721">16°11’30.2”E</geoCoordinate>
,
<date id="A539104FFFDAFFCEFB13FF0DBC3B54BE" box="[1224,1385,151,177]" pageId="7" pageNumber="221" value="2023-05-06" valueMax="2023-05-14" valueMin="2023-05-06">
<collectingDate id="B57DE9A7FFDAFFCEFB13FF0DBC3B54BE" box="[1224,1385,151,177]" pageId="7" pageNumber="221" value="2023-05-06" valueMax="2023-05-14" valueMin="2023-05-06">06–14. v.2023</collectingDate>
</date>
, leg.
<collectorName id="7C725359FFDAFFCEFF4CFF21B84954DA" box="[151,283,187,213]" pageId="7" pageNumber="221">V. Dérozier</collectorName>
,
<collectorName id="7C725359FFDAFFCEFEFDFF21B88C54DA" box="[294,478,187,213]" pageId="7" pageNumber="221">A. Kirk-Spriggs</collectorName>
,
<collectorName id="7C725359FFDAFFCEFE30FF21BB0B54DA" box="[491,601,187,213]" pageId="7" pageNumber="221">G. László</collectorName>
&amp;
<collectorName id="7C725359FFDAFFCEFDA4FF21BA4454DA" box="[639,790,187,213]" pageId="7" pageNumber="221">S. Mvouende</collectorName>
(
<specimenCount id="C781FD06FFDAFFCEFCFCFF21BA3254DA" box="[807,864,187,213]" count="3" pageId="7" pageNumber="221" type="male">3♂♂</specimenCount>
<specimenCount id="C781FD06FFDAFFCEFCB3FF21BAF254DA" box="[872,928,187,213]" count="2" pageId="7" pageNumber="221" type="female">2♀♀</specimenCount>
<collectionCode id="B796AE4AFFDAFFCEFC7CFF21BD5954DA" box="[935,1035,187,213]" pageId="7" pageNumber="221">ANHRT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="61EF3CD2FFDAFFCEFBC4FF21BB165512" collectingDate="2022-10-17" collectingDateMax="2022-10-21" collectingDateMin="2022-10-17" collectionCode="ANHRT" collectorName="V. Derozier &amp; B. Fouka &amp; A. Kirk-Spriggs &amp; H. Takano" country="Republic of the Congo" elevation="358" latitude="2.1994722" location="Bomassa Forest" longLatPrecision="1" longitude="16.19947" municipality="Nouabale-Ndoki National Park" pageId="7" pageNumber="221" specimenCount="4" specimenCount-male="4">
<location id="D4586054FFDAFFCEFBC4FF21BD8754DA" LSID="urn:lsid:plazi:treatment:592E8799FFDBFFC0FF4CF9E4B8E95206:D4586054FFDAFFCEFBC4FF21BD8754DA" box="[1055,1237,187,213]" country="Republic of the Congo" latitude="2.1994722" longLatPrecision="1" longitude="16.19947" municipality="Nouabale-Ndoki National Park" name="Bomassa Forest" pageId="7" pageNumber="221">Bomassa Forest</location>
,
<collectingMunicipality id="315CACF5FFDAFFCEFB39FF21B86154F6" pageId="7" pageNumber="221">Nouabalé-Ndoki National Park</collectingMunicipality>
,
<quantity id="167F9B6AFFDAFFCEFEE6FF44B82D54F6" box="[317,383,222,249]" metricMagnitude="2" metricUnit="m" metricValue="3.5799999999999996" pageId="7" pageNumber="221" unit="m" value="358.0">
<elevation id="5AAAD1BCFFDAFFCEFEE6FF44B82D54F6" box="[317,383,222,249]" metricMagnitude="2" metricUnit="m" metricValue="3.5799999999999996" pageId="7" pageNumber="221" unit="m" value="358.0">358m</elevation>
</quantity>
,
<geoCoordinate id="B4B35048FFDAFFCEFE52FF44BB7B54F6" box="[393,553,222,249]" degrees="02" direction="north" minutes="11" orientation="latitude" pageId="7" pageNumber="221" precision="1" seconds="58.1" value="2.1994722">02°11’58.1”N</geoCoordinate>
,
<geoCoordinate id="B4B35048FFDAFFCEFDE8FF44BB9D54F6" box="[563,719,222,249]" degrees="16" direction="east" minutes="11" orientation="longitude" pageId="7" pageNumber="221" precision="1" seconds="58.1" value="16.19947">16°11’58.1”E</geoCoordinate>
,
<date id="A539104FFFDAFFCEFD02FF45BA2454F6" box="[729,886,223,249]" pageId="7" pageNumber="221" value="2022-10-17" valueMax="2022-10-21" valueMin="2022-10-17">
<collectingDate id="B57DE9A7FFDAFFCEFD02FF45BA2454F6" box="[729,886,223,249]" pageId="7" pageNumber="221" value="2022-10-17" valueMax="2022-10-21" valueMin="2022-10-17">17–21.x.2022</collectingDate>
</date>
,
<collectingMethod id="08C64E98FFDAFFCEFC5BFF45BD5E54F6" box="[896,1036,223,249]" pageId="7" pageNumber="221">Malaise trap</collectingMethod>
, leg.
<collectorName id="7C725359FFDAFFCEFB9FFF45BD9754F6" box="[1092,1221,223,249]" pageId="7" pageNumber="221">V. Dérozier</collectorName>
,
<collectorName id="7C725359FFDAFFCEFB14FF45BC6554F6" box="[1231,1335,223,249]" pageId="7" pageNumber="221">B. Fouka</collectorName>
,
<collectorName id="7C725359FFDAFFCEFAE4FF45B9BF5512" pageId="7" pageNumber="221">A. Kirk-Spriggs</collectorName>
&amp;
<collectorName id="7C725359FFDAFFCEFEC8FE99B8DA5512" box="[275,392,259,285]" pageId="7" pageNumber="221">H. Takano</collectorName>
(
<specimenCount id="C781FD06FFDAFFCEFE43FE99B8835512" box="[408,465,259,285]" count="4" pageId="7" pageNumber="221" type="male">4♂♂</specimenCount>
<collectionCode id="B796AE4AFFDAFFCEFE03FE99BB6E5512" box="[472,572,259,285]" pageId="7" pageNumber="221">ANHRT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="61EF3CD2FFDAFFCEFD8BFE99BC5C5512" box="[592,1294,259,285]" collectingDate="2022-10-17" collectingDateMax="2022-10-23" collectingDateMin="2022-10-17" collectionCode="ANHRT" collectorName="V. Derozier &amp; B. Fouka &amp; A. Kirk-Spriggs &amp; H. Takano" country="Republic of the Congo" elevation="358" latitude="2.1994722" location="Bomassa Forest" longLatPrecision="1" longitude="16.19947" municipality="Nouabale-Ndoki National Park" pageId="7" pageNumber="221" specimenCount="1" specimenCount-male="1">
same data but
<date id="A539104FFFDAFFCEFD2FFE99BAC35512" box="[756,913,259,285]" pageId="7" pageNumber="221" value="2022-10-17" valueMax="2022-10-23" valueMin="2022-10-17">
<collectingDate id="B57DE9A7FFDAFFCEFD2FFE99BAC35512" box="[756,913,259,285]" pageId="7" pageNumber="221" value="2022-10-17" valueMax="2022-10-23" valueMin="2022-10-17">17–23.x.2022</collectingDate>
</date>
, general collection (
<specimenCount id="C781FD06FFDAFFCEFBA3FE99BDC95512" box="[1144,1179,259,285]" count="1" pageId="7" pageNumber="221" type="male">1♂</specimenCount>
<collectionCode id="B796AE4AFFDAFFCEFB7AFE99BC575512" box="[1185,1285,259,285]" pageId="7" pageNumber="221">ANHRT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="61EF3CD2FFDAFFCEFAC2FE99BA3A556A" collectingDate="2023-02-02" collectingDateMax="2023-02-06" collectingDateMin="2023-02-02" collectionCode="ANHRT" collectorName="N. Bakala &amp; V. Derozier &amp; A. Kirk-Spriggs &amp; G. Laszlo" country="Republic of the Congo" elevation="352" latitude="2.1751945" location="Mombongo" longLatPrecision="1" longitude="16.143805" municipality="Nouabale-Ndoki National Park" pageId="7" pageNumber="221" specimenCount="1" specimenCount-male="1">
<location id="D4586054FFDAFFCEFAC2FE99BCCE5512" LSID="urn:lsid:plazi:treatment:592E8799FFDBFFC0FF4CF9E4B8E95206:D4586054FFDAFFCEFAC2FE99BCCE5512" box="[1305,1436,259,285]" country="Republic of the Congo" latitude="2.1751945" longLatPrecision="1" longitude="16.143805" municipality="Nouabale-Ndoki National Park" name="Mombongo" pageId="7" pageNumber="221">Mombongo</location>
camp,
<collectingMunicipality id="315CACF5FFDAFFCEFF3EFEBDBB1B554E" box="[229,585,295,321]" pageId="7" pageNumber="221">Nouabalé-Ndoki National Park</collectingMunicipality>
,
<quantity id="167F9B6AFFDAFFCEFD8CFEBCBBC8554E" box="[599,666,294,321]" metricMagnitude="2" metricUnit="m" metricValue="3.52" pageId="7" pageNumber="221" unit="m" value="352.0">
<elevation id="5AAAD1BCFFDAFFCEFD8CFEBCBBC8554E" box="[599,666,294,321]" metricMagnitude="2" metricUnit="m" metricValue="3.52" pageId="7" pageNumber="221" unit="m" value="352.0">352m</elevation>
</quantity>
,
<geoCoordinate id="B4B35048FFDAFFCEFD73FEBDBA1B554E" box="[680,841,295,321]" degrees="02" direction="north" minutes="10" orientation="latitude" pageId="7" pageNumber="221" precision="1" seconds="30.7" value="2.1751945">02°10’30.7”N</geoCoordinate>
,
<geoCoordinate id="B4B35048FFDAFFCEFC8CFEBDBAA6554E" box="[855,1012,295,321]" degrees="16" direction="east" minutes="08" orientation="longitude" pageId="7" pageNumber="221" precision="1" seconds="37.7" value="16.143805">16°08’37.7”E</geoCoordinate>
,
<date id="A539104FFFDAFFCEFBD8FEBDBDF3554E" box="[1027,1185,295,321]" pageId="7" pageNumber="221" value="2023-02-02" valueMax="2023-02-06" valueMin="2023-02-02">
<collectingDate id="B57DE9A7FFDAFFCEFBD8FEBDBDF3554E" box="[1027,1185,295,321]" pageId="7" pageNumber="221" value="2023-02-02" valueMax="2023-02-06" valueMin="2023-02-02">02–06.ii.2023</collectingDate>
</date>
,
<collectingMethod id="08C64E98FFDAFFCEFB74FEBDBC12554E" box="[1199,1344,295,321]" pageId="7" pageNumber="221">Malaise trap</collectingMethod>
, leg.
<collectorName id="7C725359FFDAFFCEFA5AFEBDB9BA556A" pageId="7" pageNumber="221">N. Bakala</collectorName>
,
<collectorName id="7C725359FFDAFFCEFF29FED1B827556A" box="[242,373,331,357]" pageId="7" pageNumber="221">V. Dérozier</collectorName>
,
<collectorName id="7C725359FFDAFFCEFEA5FED1BB60556A" box="[382,562,331,357]" pageId="7" pageNumber="221">A. Kirk-Spriggs</collectorName>
&amp;
<collectorName id="7C725359FFDAFFCEFD8DFED1BB96556A" box="[598,708,331,357]" pageId="7" pageNumber="221">G. László</collectorName>
(
<specimenCount id="C781FD06FFDAFFCEFD0FFED1BBA5556A" box="[724,759,331,357]" count="1" pageId="7" pageNumber="221" type="male">1♂</specimenCount>
<collectionCode id="B796AE4AFFDAFFCEFD27FED1BA32556A" box="[764,864,331,357]" pageId="7" pageNumber="221">ANHRT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="85F86607FFDAFFCEFF4CF8ABBD3653D6" ID-DOI="http://doi.org/10.5281/zenodo.13346635" ID-Zenodo-Dep="13346635" httpUri="https://zenodo.org/record/13346635/files/figure.png" pageId="7" pageNumber="221" startId="7.[151,264,1841,1865]" targetBox="[158,1432,382,1805]" targetPageId="7" targetType="figure">
<paragraph id="D138368FFFDAFFCEFF4CF8ABBD3653D6" blockId="7.[151,1437,1841,2009]" pageId="7" pageNumber="221">
<emphasis id="E3F3EA9DFFDAFFCEFF4CF8ABB8075346" bold="true" box="[151,341,1841,1865]" pageId="7" pageNumber="221">FIGURES 15–24.</emphasis>
<taxonomicName id="16874D0CFFDAFFCEFE87F8ABBB7A5346" box="[348,552,1841,1865]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="221" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E3F3EA9DFFDAFFCEFE87F8ABB8825346" box="[348,464,1841,1865]" italics="true" pageId="7" pageNumber="221">Neurellipes</emphasis>
species.
</taxonomicName>
Adults, upperside (above) and underside (below).
<emphasis id="E3F3EA9DFFDAFFCEFBF4F8ABBD1D5346" bold="true" box="[1071,1103,1841,1865]" pageId="7" pageNumber="221">15.</emphasis>
<taxonomicName id="16874D0CFFDAFFCEFB8DF8A8BC2A5346" authority="(Stempffer)" baseAuthorityName="Stempffer" box="[1110,1400,1841,1865]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="221" phylum="Arthropoda" rank="subSpecies" species="ngoko" subSpecies="ngoko">
<emphasis id="E3F3EA9DFFDAFFCEFB8DF8A8BDAA5346" box="[1110,1272,1841,1865]" italics="true" pageId="7" pageNumber="221">N. ngoko ngoko</emphasis>
(Stempffer)
</taxonomicName>
, ♂, ROC, PNNN (ANHRTUK 00321840).
<emphasis id="E3F3EA9DFFDAFFCEFDF9F8CFBB135362" bold="true" box="[546,577,1877,1901]" pageId="7" pageNumber="221">16.</emphasis>
<emphasis id="E3F3EA9DFFDAFFCEFD9DF8CFBB095362" box="[582,603,1877,1901]" italics="true" pageId="7" pageNumber="221">Id</emphasis>
. (ANHRTUK 00260807).
<emphasis id="E3F3EA9DFFDAFFCEFCB3F8CFBAD55362" bold="true" box="[872,903,1877,1901]" pageId="7" pageNumber="221">17.</emphasis>
<emphasis id="E3F3EA9DFFDAFFCEFC57F8CFBAF35362" box="[908,929,1877,1901]" italics="true" pageId="7" pageNumber="221">Id</emphasis>
. (ANHRTUK 00260771).
<emphasis id="E3F3EA9DFFDAFFCEFB75F8CFBD9C5362" bold="true" box="[1198,1230,1877,1901]" pageId="7" pageNumber="221">18.</emphasis>
<emphasis id="E3F3EA9DFFDAFFCEFB08F8CFBDBA5362" box="[1235,1256,1877,1901]" italics="true" pageId="7" pageNumber="221">Id</emphasis>
., ♀, ROC, PNNN (ANHRTUK 00260682).
<emphasis id="E3F3EA9DFFDAFFCEFE7BF8E3B892539E" bold="true" box="[416,448,1913,1937]" pageId="7" pageNumber="221">19.</emphasis>
<taxonomicName id="16874D0CFFDAFFCEFE12F8E0BB75539E" authorityName="Takano" authorityYear="2024" box="[457,551,1913,1937]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="221" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="E3F3EA9DFFDAFFCEFE12F8E0BB75539E" box="[457,551,1913,1937]" italics="true" pageId="7" pageNumber="221">N. smithi</emphasis>
</taxonomicName>
<emphasis id="E3F3EA9DFFDAFFCEFDEAF8E3BBD6539E" bold="true" box="[561,644,1913,1937]" pageId="7" pageNumber="221">
<taxonomicNameLabel id="F8C057E6FFDAFFCEFDEAF8E3BBD6539E" box="[561,644,1913,1937]" pageId="7" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype ♂ (ANHRTUK 00289864).
<emphasis id="E3F3EA9DFFDAFFCEFBC2F8E3BD6B539E" bold="true" box="[1049,1081,1913,1937]" pageId="7" pageNumber="221">20.</emphasis>
<emphasis id="E3F3EA9DFFDAFFCEFB99F8E3BD05539E" box="[1090,1111,1913,1937]" italics="true" pageId="7" pageNumber="221">Id</emphasis>
., holotype ♂.
<emphasis id="E3F3EA9DFFDAFFCEFB2AF8E3BC43539E" bold="true" box="[1265,1297,1913,1937]" pageId="7" pageNumber="221">21.</emphasis>
<emphasis id="E3F3EA9DFFDAFFCEFAC1F8E3BC7D539E" box="[1306,1327,1913,1937]" italics="true" pageId="7" pageNumber="221">Id</emphasis>
., paratype ♂ (ANHRTUK 00260696).
<emphasis id="E3F3EA9DFFDAFFCEFE67F807B88953BA" bold="true" box="[444,475,1949,1973]" pageId="7" pageNumber="221">22.</emphasis>
<emphasis id="E3F3EA9DFFDAFFCEFE3FF807B8AB53BA" box="[484,505,1949,1973]" italics="true" pageId="7" pageNumber="221">Id</emphasis>
., paratype ♀ (ANHRTUK 00321906).
<emphasis id="E3F3EA9DFFDAFFCEFC48F807BAE153BA" bold="true" box="[915,947,1949,1973]" pageId="7" pageNumber="221">23.</emphasis>
<taxonomicName id="16874D0CFFDAFFCEFC67F804BD6E53BA" box="[956,1084,1949,1973]" class="Insecta" family="Lycaenidae" genus="Neurellipes" isUncertain="true" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="221" phylum="Arthropoda" rank="species" species="ngoko">
<emphasis id="E3F3EA9DFFDAFFCEFC67F804BA8653BA" box="[956,980,1950,1973]" italics="true" pageId="7" pageNumber="221">N.</emphasis>
cf.
<emphasis id="E3F3EA9DFFDAFFCEFC25F807BD6E53BA" box="[1022,1084,1949,1973]" italics="true" pageId="7" pageNumber="221">ngoko</emphasis>
</taxonomicName>
, ♂, DRC, Kwokoro (ANHRTUK 00289939).
<emphasis id="E3F3EA9DFFDAFFCEFEC9F85BB86053D6" bold="true" box="[274,306,1985,2009]" pageId="7" pageNumber="221">24.</emphasis>
<taxonomicName id="16874D0CFFDAFFCEFEE3F858BBC953D6" authority="(Bethune-Baker)" baseAuthorityName="Bethune-Baker" box="[312,667,1985,2009]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="221" phylum="Arthropoda" rank="species" species="rufomarginata">
<emphasis id="E3F3EA9DFFDAFFCEFEE3F858B8BA53D6" box="[312,488,1985,2009]" italics="true" pageId="7" pageNumber="221">N. rufomarginata</emphasis>
(Bethune-Baker)
</taxonomicName>
, ♂, DRC, Mamove (ANHRTUK 00363939).
</paragraph>
</caption>
<caption id="85F86607FFD5FFC1FF4CFCCCBAD557B9" ID-DOI="http://doi.org/10.5281/zenodo.13346637" ID-Zenodo-Dep="13346637" httpUri="https://zenodo.org/record/13346637/files/figure.png" pageId="8" pageNumber="222" startId="8.[151,264,854,878]" targetBox="[152,1435,184,825]" targetPageId="8" targetType="figure">
<paragraph id="D138368FFFD5FFC1FF4CFCCCBAD557B9" blockId="8.[151,1437,854,950]" pageId="8" pageNumber="222">
<emphasis id="E3F3EA9DFFD5FFC1FF4CFCCCB8045761" bold="true" box="[151,342,854,878]" pageId="8" pageNumber="222">FIGURES 25–27.</emphasis>
<taxonomicName id="16874D0CFFD5FFC1FE85FCCCBB715761" box="[350,547,854,878]" pageId="8" pageNumber="222">
<emphasis id="E3F3EA9DFFD5FFC1FE85FCCCB8805761" box="[350,466,854,878]" italics="true" pageId="8" pageNumber="222">Neurellipes</emphasis>
species
</taxonomicName>
♂ genitalia. Capsule in lateral view (top), phallus (middle) and valve in three-quarter view (bottom).
<emphasis id="E3F3EA9DFFD5FFC1FEEFFCE0B806579D" bold="true" box="[308,340,890,914]" pageId="8" pageNumber="222">25.</emphasis>
<taxonomicName id="16874D0CFFD5FFC1FE81FCE1BB2B579D" authority="(Stempffer)" baseAuthorityName="Stempffer" box="[346,633,890,914]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="222" phylum="Arthropoda" rank="subSpecies" species="ngoko" subSpecies="ngoko">
<emphasis id="E3F3EA9DFFD5FFC1FE81FCE1B8A8579D" box="[346,506,890,914]" italics="true" pageId="8" pageNumber="222">N. ngoko ngoko</emphasis>
(Stempffer)
</taxonomicName>
, ROC, PNNN (ANHRTUK 00260768).
<emphasis id="E3F3EA9DFFD5FFC1FBC8FCE0BD61579D" bold="true" box="[1043,1075,890,914]" pageId="8" pageNumber="222">26.</emphasis>
<taxonomicName id="16874D0CFFD5FFC1FBE2FCE1BDC7579D" authorityName="Takano" authorityYear="2024" box="[1081,1173,890,914]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="222" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="E3F3EA9DFFD5FFC1FBE2FCE1BDC7579D" box="[1081,1173,890,914]" italics="true" pageId="8" pageNumber="222">N. smithi</emphasis>
</taxonomicName>
<emphasis id="E3F3EA9DFFD5FFC1FB47FCE0BDBE579D" bold="true" box="[1180,1260,890,914]" pageId="8" pageNumber="222">
<taxonomicNameLabel id="F8C057E6FFD5FFC1FB47FCE0BDBE579D" box="[1180,1260,890,914]" pageId="8" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis id="E3F3EA9DFFD5FFC1FA84FCE0BC2D579D" bold="true" box="[1375,1407,890,914]" pageId="8" pageNumber="222">27.</emphasis>
<taxonomicName id="16874D0CFFD5FFC1FA5EFCE1B88F57B9" authority="(Bethune-Baker)" baseAuthorityName="Bethune-Baker" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="222" phylum="Arthropoda" rank="species" species="rufomarginata">
<emphasis id="E3F3EA9DFFD5FFC1FA5EFCE1B87857B9" italics="true" pageId="8" pageNumber="222">N. rufomarginata</emphasis>
(Bethune-Baker)
</taxonomicName>
, DRC, Mamove (ANHRTUK 00363939).
</paragraph>
</caption>
<subSubSection id="999D6504FFD5FFC0FF1CFB92BDDB556A" lastPageId="9" lastPageNumber="223" pageId="8" pageNumber="222" type="description">
<paragraph id="D138368FFFD5FFC1FF1CFB92B80B502D" blockId="8.[151,1437,1032,2031]" box="[199,345,1032,1058]" pageId="8" pageNumber="222">
<emphasis id="E3F3EA9DFFD5FFC1FF1CFB92B80B502D" bold="true" box="[199,345,1032,1058]" pageId="8" pageNumber="222">Description.</emphasis>
</paragraph>
<paragraph id="D138368FFFD5FFC1FF1CFBB6BC775048" blockId="8.[151,1437,1032,2031]" box="[199,1317,1068,1095]" pageId="8" pageNumber="222">
Forewing length:
<typeStatus id="0E3C882DFFD5FFC1FE55FBB6B8A25049" box="[398,496,1068,1094]" pageId="8" pageNumber="222" type="holotype">holotype</typeStatus>
male:
<quantity id="167F9B6AFFD5FFC1FDE6FBB6BBDD5048" box="[573,655,1068,1095]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="222" unit="mm" value="13.0">13 mm</quantity>
;
<typeStatus id="0E3C882DFFD5FFC1FD41FBB7BBA85048" box="[666,762,1069,1095]" pageId="8" pageNumber="222" type="paratype">paratype</typeStatus>
males:
<quantity id="167F9B6AFFD5FFC1FC8AFBB6BA9E5048" box="[849,972,1068,1095]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="8" pageNumber="222" unit="mm" value="13.5" valueMax="16.0" valueMin="11.0">11–16 mm</quantity>
;
<typeStatus id="0E3C882DFFD5FFC1FC03FBB7BD6A5048" box="[984,1080,1069,1095]" pageId="8" pageNumber="222" type="paratype">paratype</typeStatus>
females:
<quantity id="167F9B6AFFD5FFC1FB7EFBB6BC735048" box="[1189,1313,1068,1095]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="8" pageNumber="222" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">14–15 mm</quantity>
.
</paragraph>
<paragraph id="D138368FFFD5FFC1FF1CFBCAB89753E1" blockId="8.[151,1437,1032,2031]" pageId="8" pageNumber="222">
Male. Upperside: ground colour of head, thorax and abdomen black.Antennae black, with white scaling ventrally at the base of each segment. Frons with a narrow row of white scales running adjacent to the eye. Palps longer than diameter of eye with white scaling. Abdominal segments with dark greyish-brown scaling along posterior margin of segments giving the abdomen a slightly banded appearance. Ground colour of both wings dark brown; terminal line black; cilia greyish-brown. Forewing costa gently arcuate, rounded at apex; outer margin gently arcuate. Discal marking ovoid, longitudinally elongate, pale-orange in colour, occupying the dorsal half of space 1 and distally to within
<quantity id="167F9B6AFFD5FFC1FF3CFAB2B87B514C" box="[231,297,1320,1347]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="222" unit="mm" value="3.0">3 mm</quantity>
of the outer margin, space
<quantity id="167F9B6AFFD5FFC1FDBBFAB2BBDE514D" box="[608,652,1320,1346]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="222" unit="in" value="2.0">2 in</quantity>
its entirety except for a
<quantity id="167F9B6AFFD5FFC1FC78FAB2BAB7514C" box="[931,997,1320,1347]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="222" unit="mm" value="2.0">2 mm</quantity>
outer margin, and the base of space 3. Hindwing arcuate, with three
<quantity id="167F9B6AFFD5FFC1FE37FAD6BB7D5168" box="[492,559,1356,1383]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="222" unit="mm" value="1.0">1 mm</quantity>
long, greyish-brown ‘tails’ protruding from the outer margin along veins 1a, 2 and 3. Ground colour slightly paler dorsally, from space 7 to 8. The cell, spaces 1–2 and the inner margin with long light-brown ciliate scales. Discal marking pale orange, in the form of a submarginal band running broadly parallel with the marginal band from the dorsal half of space 1a to the ventral half of space 4. Marginal band formed of black spots boldly surrounded proximally and laterally with pale orange and a thin white fascia parallel to the outer margin distally; spots in space 1 conjoined, the proximal margin strongly emarginate medially. Both discal and marginal bands strongly bisected with heavy black scaling along the veins. Underside. Head and thorax covered in greyish-white scales. Legs dark brown with a dusting of white scales, with a concentration of white scales at the base of each tarsal segment. Abdomen with dense cream ciliate scales. Ground colour of both wings pale brownish-grey with white bilineate bands and fasciae; discal triangle with pale-orange marking from upperside weakly showing through; terminal line dark brown. Forewing basal band with two lines converging dorsally, filled with darker brown scales and in space 1 deep charcoal. Discal band defined with dark brown on the inner edges. Postdiscal band indented, defined with dark brown forming spots on its inner edge. Antemarginal and Submarginal lines indistinct, broadening towards ventral margin; the space in between filled with dark brown scaling from space 3 to ventral margin. Marginal line fine, indistinct and interrupted at veins. Hindwing with basal and discal bands irrorated internally with darker scales. Postmedial band indented, defined with dark brown forming spots on its inner edge. Antemarginal fascia arcuate, diffuse. Marginal fascia mirroring upperside, but the black spots replaced with the ground colour and the orange replaced with white except for in spaces 1 and 2; these two spots with a ring of iridescent green scaling.
</paragraph>
<paragraph id="D138368FFFD4FFC0FF1CFF0DBDDA5512" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="223">Male genitalia. Uncus robust, indented with a broad concavity medially. Subuncus long and fine, almost rightangled medially with hooked apices. Valve rectangular, the distal margin finely serrate with two sharp projections ventro-distally, the ventral projection longer than the dorsal one. Saccus long, its margins evenly arcuate in ventral view. Aedeagus almost three times as long as valve, distally narrower and tapered at apex.</paragraph>
<paragraph id="D138368FFFD4FFC0FF1CFEBDBDDB556A" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="223">Female. Similar to male but paler and fringe longer. Outer margin of forewing more arcuate. Hindwing upperside marginal band extending to space 6 becoming paler and more indistinct towards the costa.</paragraph>
</subSubSection>
<subSubSection id="999D6504FFD4FFC0FF1CFEF4BCDB571A" pageId="9" pageNumber="223" type="diagnosis">
<paragraph id="D138368FFFD4FFC0FF1CFEF4BCDB571A" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="223">
<emphasis id="E3F3EA9DFFD4FFC0FF1CFEF4B8105587" bold="true" box="[199,322,366,392]" pageId="9" pageNumber="223">Diagnosis.</emphasis>
The new species is very similar in appearance to the sympatric
<taxonomicName id="16874D0CFFD4FFC0FBD9FEF5B98B55A2" authority="(Stempffer, 1962)" baseAuthorityName="Stempffer" baseAuthorityYear="1962" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="subSpecies" species="ngoko" subSpecies="ngoko">
<emphasis id="E3F3EA9DFFD4FFC0FBD9FEF5BC455586" box="[1026,1303,367,393]" italics="true" pageId="9" pageNumber="223">Neurellipes ngoko ngoko</emphasis>
(
<bibRefCitation id="B5164B7EFFD4FFC0FAFEFEF5B98255A2" author="Stempffer, H." pageId="9" pageNumber="223" pagination="1135 - 1181" refId="ref11446" refString="Stempffer, H. (1962) Contribution a l'etude des Lepidopteres Lycaenidae de l'Afrique tropicale et equatoriale. Bulletin de l'Institut Francais d'Afrique Noire, Series A, 24, 1135 - 1181." type="journal article" year="1962">Stempffer, 1962</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="49BC2A0AFFD4FFC0FF31FE09B83C55A2" box="[234,366,402,429]" captionStart="FIGURES 15–24" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[158,1432,382,1805]" captionTargetId="figure-109@7.[151,1435,381,1818]" captionTargetPageId="7" captionText="FIGURES 15–24. Neurellipes species. Adults, upperside (above) and underside (below). 15. N. ngoko ngoko (Stempffer), ♂, ROC, PNNN (ANHRTUK 00321840). 16. Id. (ANHRTUK 00260807). 17. Id. (ANHRTUK 00260771). 18. Id., ♀, ROC, PNNN (ANHRTUK 00260682). 19. N. smithi sp. nov., paratype ♂ (ANHRTUK 00289864). 20. Id., holotype ♂. 21. Id., paratype ♂ (ANHRTUK 00260696). 22. Id., paratype ♀ (ANHRTUK 00321906). 23. N. cf. ngoko, ♂, DRC, Kwokoro (ANHRTUK 00289939). 24. N. rufomarginata (Bethune-Baker), ♂, DRC, Mamove (ANHRTUK 00363939)." figureDoi="http://doi.org/10.5281/zenodo.13346635" httpUri="https://zenodo.org/record/13346635/files/figure.png" pageId="9" pageNumber="223">Figs 15–18</figureCitation>
), but in both sexes of the latter, the upperside markings are a darker reddish-orange, and the hindwing discal marking is larger often extending into space 5 and reaching the cell in spaces 2 and 3 combined with limited fine black scaling along the veins giving the appearance of a continuous patch. The underside ground colour of
<taxonomicName id="16874D0CFFD4FFC0FF6CFE65B84F5616" box="[183,285,511,537]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="ngoko">
<emphasis id="E3F3EA9DFFD4FFC0FF6CFE65B84F5616" box="[183,285,511,537]" italics="true" pageId="9" pageNumber="223">N. ngoko</emphasis>
</taxonomicName>
is a darker brownish-grey, and the dark brown spots in the postdiscal bands are much better defined. Markings are however variable in both species and the development of the upperside orange markings can be heavily reduced (
<figureCitation id="49BC2A0AFFD4FFC0FE8CFDDDB885566E" box="[343,471,582,609]" captionStart="FIGURES 15–24" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[158,1432,382,1805]" captionTargetId="figure-109@7.[151,1435,381,1818]" captionTargetPageId="7" captionText="FIGURES 15–24. Neurellipes species. Adults, upperside (above) and underside (below). 15. N. ngoko ngoko (Stempffer), ♂, ROC, PNNN (ANHRTUK 00321840). 16. Id. (ANHRTUK 00260807). 17. Id. (ANHRTUK 00260771). 18. Id., ♀, ROC, PNNN (ANHRTUK 00260682). 19. N. smithi sp. nov., paratype ♂ (ANHRTUK 00289864). 20. Id., holotype ♂. 21. Id., paratype ♂ (ANHRTUK 00260696). 22. Id., paratype ♀ (ANHRTUK 00321906). 23. N. cf. ngoko, ♂, DRC, Kwokoro (ANHRTUK 00289939). 24. N. rufomarginata (Bethune-Baker), ♂, DRC, Mamove (ANHRTUK 00363939)." figureDoi="http://doi.org/10.5281/zenodo.13346635" httpUri="https://zenodo.org/record/13346635/files/figure.png" pageId="9" pageNumber="223">Figs 15, 19</figureCitation>
) or strongly expressed (
<figureCitation id="49BC2A0AFFD4FFC0FD38FDDDBA36566E" box="[739,868,583,609]" captionStart="FIGURES 15–24" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[158,1432,382,1805]" captionTargetId="figure-109@7.[151,1435,381,1818]" captionTargetPageId="7" captionText="FIGURES 15–24. Neurellipes species. Adults, upperside (above) and underside (below). 15. N. ngoko ngoko (Stempffer), ♂, ROC, PNNN (ANHRTUK 00321840). 16. Id. (ANHRTUK 00260807). 17. Id. (ANHRTUK 00260771). 18. Id., ♀, ROC, PNNN (ANHRTUK 00260682). 19. N. smithi sp. nov., paratype ♂ (ANHRTUK 00289864). 20. Id., holotype ♂. 21. Id., paratype ♂ (ANHRTUK 00260696). 22. Id., paratype ♀ (ANHRTUK 00321906). 23. N. cf. ngoko, ♂, DRC, Kwokoro (ANHRTUK 00289939). 24. N. rufomarginata (Bethune-Baker), ♂, DRC, Mamove (ANHRTUK 00363939)." figureDoi="http://doi.org/10.5281/zenodo.13346635" httpUri="https://zenodo.org/record/13346635/files/figure.png" pageId="9" pageNumber="223">Figs 17, 21</figureCitation>
). In the male genitalia, the general configuration is very similar but in
<taxonomicName id="16874D0CFFD4FFC0FEB3FDF1B899568A" box="[360,459,619,645]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="ngoko">
<emphasis id="E3F3EA9DFFD4FFC0FEB3FDF1B899568A" box="[360,459,619,645]" italics="true" pageId="9" pageNumber="223">N. ngoko</emphasis>
</taxonomicName>
(
<figureCitation id="49BC2A0AFFD4FFC0FE01FDF1BB7E568A" box="[474,556,618,645]" captionStart="FIGURES 25–27" captionStartId="8.[151,264,854,878]" captionTargetBox="[152,1435,184,825]" captionTargetId="figure-15@8.[151,1436,181,830]" captionTargetPageId="8" captionText="FIGURES 25–27. Neurellipes species ♂ genitalia. Capsule in lateral view (top), phallus (middle) and valve in three-quarter view (bottom). 25. N. ngoko ngoko (Stempffer), ROC, PNNN (ANHRTUK 00260768). 26. N. smithi sp. nov., holotype. 27. N. rufomarginata (Bethune-Baker), DRC, Mamove (ANHRTUK 00363939)." figureDoi="http://doi.org/10.5281/zenodo.13346637" httpUri="https://zenodo.org/record/13346637/files/figure.png" pageId="9" pageNumber="223">Fig. 25</figureCitation>
), the capsule is larger, the ventro-distal projections of the valve are nearly twice as long, and the concavity between them is broader. Although the upperside markings and colouration are different,
<taxonomicName id="16874D0CFFD4FFC0FF4CFD29BB3B56C2" authority="(Bethune-Baker, 1910)" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1910" box="[151,617,691,717]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="rufomarginata">
<emphasis id="E3F3EA9DFFD4FFC0FF4CFD29B80E56C2" box="[151,348,691,717]" italics="true" pageId="9" pageNumber="223">N. rufomarginata</emphasis>
(
<bibRefCitation id="B5164B7EFFD4FFC0FEB6FD29BB3356C2" author="Bethune-Baker, G. T." box="[365,609,691,717]" pageId="9" pageNumber="223" pagination="1 - 84" refId="ref9569" refString="Bethune-Baker, G. T. (1910) A revision of the African species of the Lycaenesthes group of the Lycaenidae. Transactions of the Entomological Society of London, 1910, 1 - 84. https: // doi. org / 10.1111 / j. 1365 - 2311.1910. tb 01165. x" type="journal article" year="1910">Bethune-Baker, 1910</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="49BC2A0AFFD4FFC0FDA0FD29BB9D56C2" box="[635,719,691,717]" captionStart="FIGURES 15–24" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[158,1432,382,1805]" captionTargetId="figure-109@7.[151,1435,381,1818]" captionTargetPageId="7" captionText="FIGURES 15–24. Neurellipes species. Adults, upperside (above) and underside (below). 15. N. ngoko ngoko (Stempffer), ♂, ROC, PNNN (ANHRTUK 00321840). 16. Id. (ANHRTUK 00260807). 17. Id. (ANHRTUK 00260771). 18. Id., ♀, ROC, PNNN (ANHRTUK 00260682). 19. N. smithi sp. nov., paratype ♂ (ANHRTUK 00289864). 20. Id., holotype ♂. 21. Id., paratype ♂ (ANHRTUK 00260696). 22. Id., paratype ♀ (ANHRTUK 00321906). 23. N. cf. ngoko, ♂, DRC, Kwokoro (ANHRTUK 00289939). 24. N. rufomarginata (Bethune-Baker), ♂, DRC, Mamove (ANHRTUK 00363939)." figureDoi="http://doi.org/10.5281/zenodo.13346635" httpUri="https://zenodo.org/record/13346635/files/figure.png" pageId="9" pageNumber="223">Fig. 24</figureCitation>
) shares a similar genital ground plan but differs in the shorter valves, finer ventro-distal projections of the valve and a narrower concavity between them (
<figureCitation id="49BC2A0AFFD4FFC0FB7CFD4DBDAE56FE" box="[1191,1276,727,753]" captionStart="FIGURES 25–27" captionStartId="8.[151,264,854,878]" captionTargetBox="[152,1435,184,825]" captionTargetId="figure-15@8.[151,1436,181,830]" captionTargetPageId="8" captionText="FIGURES 25–27. Neurellipes species ♂ genitalia. Capsule in lateral view (top), phallus (middle) and valve in three-quarter view (bottom). 25. N. ngoko ngoko (Stempffer), ROC, PNNN (ANHRTUK 00260768). 26. N. smithi sp. nov., holotype. 27. N. rufomarginata (Bethune-Baker), DRC, Mamove (ANHRTUK 00363939)." figureDoi="http://doi.org/10.5281/zenodo.13346637" httpUri="https://zenodo.org/record/13346637/files/figure.png" pageId="9" pageNumber="223">Fig. 27</figureCitation>
). Intraspecific PWDs of the new species were 0.0–0.2% (n=5) and diverged interspecifically from
<taxonomicName id="16874D0CFFD4FFC0FB99FD61BDF5571A" box="[1090,1191,763,789]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="ngoko">
<emphasis id="E3F3EA9DFFD4FFC0FB99FD61BDF5571A" box="[1090,1191,763,789]" italics="true" pageId="9" pageNumber="223">N. ngoko</emphasis>
</taxonomicName>
by 2.5–3.5% (n=6).
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFD4FFC0FF1CFC85BD75515A" pageId="9" pageNumber="223" type="discussion">
<paragraph id="D138368FFFD4FFC0FF1CFC85BD75515A" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="223">
<emphasis id="E3F3EA9DFFD4FFC0FF1CFC85B8695736" bold="true" box="[199,315,799,825]" pageId="9" pageNumber="223">Remarks.</emphasis>
A long series of
<taxonomicName id="16874D0CFFD4FFC0FE21FC85BB285736" authorityName="Bethune-Baker" authorityYear="1910" box="[506,634,799,825]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="E3F3EA9DFFD4FFC0FE21FC85BB285736" box="[506,634,799,825]" italics="true" pageId="9" pageNumber="223">Neurellipes</emphasis>
</taxonomicName>
specimens with the elongate orange oval marking on the forewing was captured, in part due to the deployment of Malaise traps in suitable habitats (Takano
<emphasis id="E3F3EA9DFFD4FFC0FBB5FCD9BDFB5752" box="[1134,1193,835,861]" italics="true" pageId="9" pageNumber="223">et al.</emphasis>
2024). Malaise traps accounted for 32 of the
<specimenCount id="C781FD06FFD4FFC0FE75FCFCBB1A578E" box="[430,584,870,897]" count="54" pageId="9" pageNumber="223" type="generic">54 specimens</specimenCount>
of
<taxonomicName id="16874D0CFFD4FFC0FDA9FCFDBB8B578E" box="[626,729,871,897]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="ngoko">
<emphasis id="E3F3EA9DFFD4FFC0FDA9FCFDBB8B578E" box="[626,729,871,897]" italics="true" pageId="9" pageNumber="223">N. ngoko</emphasis>
</taxonomicName>
, and five of the
<specimenCount id="C781FD06FFD4FFC0FC46FCFDBD65578E" box="[925,1079,871,897]" count="16" pageId="9" pageNumber="223" type="generic">16 specimens</specimenCount>
of
<taxonomicName id="16874D0CFFD4FFC0FBBAFCFDBD9A578E" authorityName="Takano" authorityYear="2024" box="[1121,1224,871,897]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov.">
<emphasis id="E3F3EA9DFFD4FFC0FBBAFCFDBD9A578E" box="[1121,1224,871,897]" italics="true" pageId="9" pageNumber="223">N. smithi</emphasis>
</taxonomicName>
<emphasis id="E3F3EA9DFFD4FFC0FB09FCFDBC7C578E" bold="true" box="[1234,1326,871,897]" pageId="9" pageNumber="223">
<taxonomicNameLabel id="F8C057E6FFD4FFC0FB09FCFDBC7C578E" box="[1234,1326,871,897]" pageId="9" pageNumber="223" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
collected at PNNN.
<taxonomicName id="16874D0CFFD4FFC0FECDFC11B8B457AA" box="[278,486,907,933]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="ngoko">
<emphasis id="E3F3EA9DFFD4FFC0FECDFC11B8B457AA" box="[278,486,907,933]" italics="true" pageId="9" pageNumber="223">Neurellipes ngoko</emphasis>
</taxonomicName>
was described from Ouesso,
<collectingCountry id="A990761FFFD4FFC0FC9CFC11BAD357AA" box="[839,897,907,933]" name="Taiwan" pageId="9" pageNumber="223">ROC</collectingCountry>
and is not known from anywhere east of the
<collectingCountry id="A990761FFFD4FFC0FF4CFC35B9B057C6" box="[151,226,943,969]" name="Democratic Republic of the Congo" pageId="9" pageNumber="223">Congo</collectingCountry>
River (
<bibRefCitation id="B5164B7EFFD4FFC0FEE6FC35B89957C6" author="Libert, M." box="[317,459,943,969]" pageId="9" pageNumber="223" refId="ref10860" refString="Libert, M. (2010) Revision des Anthene africains (Lepidoptera, Lycaenidae). ABRI, Nairobi, Kenya and Lambillionea, Tervuren, 420 pp." type="book" year="2010">Libert 2010</bibRefCitation>
), whilst its allopatric sibling
<taxonomicName id="16874D0CFFD4FFC0FCF8FC35BABB57C6" box="[803,1001,943,969]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="rufomarginata">
<emphasis id="E3F3EA9DFFD4FFC0FCF8FC35BABB57C6" box="[803,1001,943,969]" italics="true" pageId="9" pageNumber="223">N. rufomarginata</emphasis>
</taxonomicName>
, distributed in north-eastern
<collectingCountry id="A990761FFFD4FFC0FAE5FC35BC2857C6" box="[1342,1402,943,969]" name="Democratic Republic of the Congo" pageId="9" pageNumber="223">DRC</collectingCountry>
, is separated by some
<quantity id="167F9B6AFFD4FFC0FEB0FC49B8ED57E2" box="[363,447,979,1005]" metricMagnitude="5" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="223" unit="km" value="800.0">800 km</quantity>
(its most westerly record being Kisangani). There is however a short series of specimens (n=11) from Kwokoro,
<collectingCountry id="A990761FFFD4FFC0FE7DFC6DB8B2501E" box="[422,480,1015,1041]" name="Democratic Republic of the Congo" pageId="9" pageNumber="223">DRC</collectingCountry>
(
<figureCitation id="49BC2A0AFFD4FFC0FE29FC6DBB14501E" box="[498,582,1015,1041]" captionStart="FIGURES 15–24" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[158,1432,382,1805]" captionTargetId="figure-109@7.[151,1435,381,1818]" captionTargetPageId="7" captionText="FIGURES 15–24. Neurellipes species. Adults, upperside (above) and underside (below). 15. N. ngoko ngoko (Stempffer), ♂, ROC, PNNN (ANHRTUK 00321840). 16. Id. (ANHRTUK 00260807). 17. Id. (ANHRTUK 00260771). 18. Id., ♀, ROC, PNNN (ANHRTUK 00260682). 19. N. smithi sp. nov., paratype ♂ (ANHRTUK 00289864). 20. Id., holotype ♂. 21. Id., paratype ♂ (ANHRTUK 00260696). 22. Id., paratype ♀ (ANHRTUK 00321906). 23. N. cf. ngoko, ♂, DRC, Kwokoro (ANHRTUK 00289939). 24. N. rufomarginata (Bethune-Baker), ♂, DRC, Mamove (ANHRTUK 00363939)." figureDoi="http://doi.org/10.5281/zenodo.13346635" httpUri="https://zenodo.org/record/13346635/files/figure.png" pageId="9" pageNumber="223">Fig. 23</figureCitation>
) in the ANHRT collections somewhat intermediate between
<taxonomicName id="16874D0CFFD4FFC0FAD8FC6DBC38501E" box="[1283,1386,1015,1041]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="ngoko">
<emphasis id="E3F3EA9DFFD4FFC0FAD8FC6DBC38501E" box="[1283,1386,1015,1041]" italics="true" pageId="9" pageNumber="223">N. ngoko</emphasis>
</taxonomicName>
and a
<taxonomicName id="16874D0CFFD4FFC0FF77FB81B823503A" box="[172,369,1051,1077]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="rufomarginata">
<emphasis id="E3F3EA9DFFD4FFC0FF77FB81B823503A" box="[172,369,1051,1077]" italics="true" pageId="9" pageNumber="223">N. rufomarginata</emphasis>
</taxonomicName>
specimen from Epulu,
<collectingCountry id="A990761FFFD4FFC0FDA6FB81BBE5503A" box="[637,695,1051,1077]" name="Democratic Republic of the Congo" pageId="9" pageNumber="223">DRC</collectingCountry>
figured on Plate XXIV of
<bibRefCitation id="B5164B7EFFD4FFC0FC37FB81BDC5503A" author="Libert, M." box="[1004,1175,1051,1077]" pageId="9" pageNumber="223" refId="ref10860" refString="Libert, M. (2010) Revision des Anthene africains (Lepidoptera, Lycaenidae). ABRI, Nairobi, Kenya and Lambillionea, Tervuren, 420 pp." type="book" year="2010">Libert’s (2010)</bibRefCitation>
revision. Although the genitalia and the deeper orange colouration of the wing markings are similar to true
<taxonomicName id="16874D0CFFD4FFC0FBBAFBA5BD955056" box="[1121,1223,1087,1113]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="ngoko">
<emphasis id="E3F3EA9DFFD4FFC0FBBAFBA5BD955056" box="[1121,1223,1087,1113]" italics="true" pageId="9" pageNumber="223">N. ngoko</emphasis>
</taxonomicName>
, the wing patterns are consistently more reduced and less well-defined; this population may yet constitute an undescribed taxon but without barcode data, its identity is not dealt with any further herein and merely highlighted as being a group that requires further investigations. The male specimens of
<taxonomicName id="16874D0CFFD4FFC0FD26FB31BA3350CA" box="[765,865,1195,1221]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="ngoko">
<emphasis id="E3F3EA9DFFD4FFC0FD26FB31BA3350CA" box="[765,865,1195,1221]" italics="true" pageId="9" pageNumber="223">N. ngoko</emphasis>
</taxonomicName>
from PNNN also show a high degree of variability with some males (
<figureCitation id="49BC2A0AFFD4FFC0FEB2FB55B8EF50E6" box="[361,445,1231,1257]" captionStart="FIGURES 15–24" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[158,1432,382,1805]" captionTargetId="figure-109@7.[151,1435,381,1818]" captionTargetPageId="7" captionText="FIGURES 15–24. Neurellipes species. Adults, upperside (above) and underside (below). 15. N. ngoko ngoko (Stempffer), ♂, ROC, PNNN (ANHRTUK 00321840). 16. Id. (ANHRTUK 00260807). 17. Id. (ANHRTUK 00260771). 18. Id., ♀, ROC, PNNN (ANHRTUK 00260682). 19. N. smithi sp. nov., paratype ♂ (ANHRTUK 00289864). 20. Id., holotype ♂. 21. Id., paratype ♂ (ANHRTUK 00260696). 22. Id., paratype ♀ (ANHRTUK 00321906). 23. N. cf. ngoko, ♂, DRC, Kwokoro (ANHRTUK 00289939). 24. N. rufomarginata (Bethune-Baker), ♂, DRC, Mamove (ANHRTUK 00363939)." figureDoi="http://doi.org/10.5281/zenodo.13346635" httpUri="https://zenodo.org/record/13346635/files/figure.png" pageId="9" pageNumber="223">Fig. 17</figureCitation>
) displaying phenotypes very similar to
<taxonomicName id="16874D0CFFD4FFC0FC5EFB55BDBD50E6" authority="Libert, 2010" authorityName="Libert" authorityYear="2010" box="[901,1263,1231,1257]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="subSpecies" species="ngoko" subSpecies="fontainei">
<emphasis id="E3F3EA9DFFD4FFC0FC5EFB55BD0850E6" box="[901,1114,1231,1257]" italics="true" pageId="9" pageNumber="223">N. ngoko fontainei</emphasis>
<bibRefCitation id="B5164B7EFFD4FFC0FBB8FB55BDBD50E6" author="Libert, M." box="[1123,1263,1231,1257]" pageId="9" pageNumber="223" refId="ref10860" refString="Libert, M. (2010) Revision des Anthene africains (Lepidoptera, Lycaenidae). ABRI, Nairobi, Kenya and Lambillionea, Tervuren, 420 pp." type="book" year="2010">Libert, 2010</bibRefCitation>
</taxonomicName>
from Sankuru,
<collectingCountry id="A990761FFFD4FFC0FF4CFB69B9865102" box="[151,212,1267,1293]" name="Democratic Republic of the Congo" pageId="9" pageNumber="223">DRC</collectingCountry>
. Based on the series collected at PNNN, it is possible that
<taxonomicName id="16874D0CFFD4FFC0FCBAFB69BAA55102" authorityName="Libert" authorityYear="2010" box="[865,1015,1267,1293]" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="subSpecies" species="ngoko" subSpecies="fontainei">
ssp.
<emphasis id="E3F3EA9DFFD4FFC0FC48FB69BAA55102" box="[915,1015,1267,1293]" italics="true" pageId="9" pageNumber="223">fontainei</emphasis>
</taxonomicName>
is merely a form of nominotypical
<taxonomicName id="16874D0CFFD4FFC0FA58FB69B989513E" class="Insecta" family="Lycaenidae" genus="Neurellipes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="223" phylum="Arthropoda" rank="species" species="ngoko">
<emphasis id="E3F3EA9DFFD4FFC0FA58FB69B989513E" italics="true" pageId="9" pageNumber="223">N. ngoko</emphasis>
</taxonomicName>
, but since the distribution is disjunct (ca.
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apart), the two taxa are maintained as being separate for the time being until further material becomes available in between these populations.
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFD4FFC0FF1CFAC5BB4E51CE" pageId="9" pageNumber="223" type="etymology">
<paragraph id="D138368FFFD4FFC0FF1CFAC5BB4E51CE" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="223">
<emphasis id="E3F3EA9DFFD4FFC0FF1CFAC5B8CC5176" bold="true" box="[199,414,1375,1401]" pageId="9" pageNumber="223">
<emphasis id="E3F3EA9DFFD4FFC0FF1CFAC5B8C55176" bold="true" box="[199,407,1375,1401]" italics="true" pageId="9" pageNumber="223">Derivatio nominis</emphasis>
.
</emphasis>
It is with great pleasure that I dedicate this new species to Richard Smith, Chairman of the Board of Trustees, ANHRT, for his outstanding and ongoing contribution to African entomology and biogeography through the institution he founded.
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFD4FFC0FF1CFA50B8E95206" pageId="9" pageNumber="223" type="distribution">
<paragraph id="D138368FFFD4FFC0FF1CFA50B8E95206" blockId="9.[151,1437,151,1545]" pageId="9" pageNumber="223">
<emphasis id="E3F3EA9DFFD4FFC0FF1CFA50B83351EB" bold="true" box="[199,353,1482,1508]" pageId="9" pageNumber="223">Distribution.</emphasis>
The new species is known only from PNNN but may be more widely distributed in the Sangha-Congo interfluvial region.
</paragraph>
</subSubSection>
</treatment>
</document>

440
data/59/2E/87/592E8799FFDFFFCDFF4CFB56BA1E51EA.xml Normal file
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@ -0,0 +1,440 @@
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<mods:title id="D164967B62863A40BA90341786A5997E">Descriptions of three new species of Lycaenidae (Lepidoptera: Papilionoidea) from Parc National de Nouabalé-Ndoki, Republic of Congo with additional notes on little-known species</mods:title>
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<emphasis id="E3F3EA9DFFDFFFCBFF4CFB56B8E050E8" bold="true" box="[151,434,1228,1255]" italics="true" pageId="2" pageNumber="216">Falcuna nouabaleensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8C057E6FFDFFFCBFE62FB56BB4550E9" box="[441,535,1228,1254]" pageId="2" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="999D6504FFDFFFCBFF4CFB6AB8035105" box="[151,337,1264,1290]" pageId="2" pageNumber="216" type="description">
<paragraph id="D138368FFFDFFFCBFF4CFB6AB8035105" blockId="2.[151,535,1228,1290]" box="[151,337,1264,1290]" pageId="2" pageNumber="216">
(
<figureCitation id="49BC2A0AFFDFFFCBFF44FB6AB84D5105" box="[159,287,1264,1290]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1907,1931]" captionTargetBox="[152,1434,184,1870]" captionTargetId="figure-30@3.[151,1435,181,1883]" captionTargetPageId="3" captionText="FIGURES 1–8. Falcuna species. Adults (1–6), upperside (above) and underside (below); ♂ genitalia (7–8) in lateral view. 1. F. lybia (Staudinger), paralectotype ♂ of Larinopoda libya (MfN) with labels. 2. Id., ♂, Cameroon, Nkol Nlong (RCJS). 3. Id., ♀, Cameroon, Zoétélé (RCJS). 4. F. nouabaleensis sp. nov., holotype ♂. 5. Id., paratype ♂ (ANHRTUK 00260604). 6. Id., paratype ♀ (ANHRTUK 00321798). 7. F. lybia. 8. F. nouabaleensis." figureDoi="http://doi.org/10.5281/zenodo.13346629" httpUri="https://zenodo.org/record/13346629/files/figure.png" pageId="2" pageNumber="216">Figs 4–6, 8</figureCitation>
,
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)
</paragraph>
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(
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):
</materialsCitation>
</paragraph>
<paragraph id="D138368FFFDFFFCBFF1CFAC2BA4E51B5" blockId="2.[151,1437,1332,2043]" pageId="2" pageNumber="216">
“
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<collectingCountry id="A990761FFFDFFFCBFF0AFAC2B8BB517D" box="[209,489,1368,1394]" name="Republic of the Congo" pageId="2" pageNumber="216">REPUBLIC OF CONGO</collectingCountry>
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</quantity>
/
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,
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/
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,
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camp/(
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<emphasis id="E3F3EA9DFFDFFFCBFB02FAC2BCCE517D" box="[1241,1436,1368,1394]" italics="true" pageId="2" pageNumber="216">Gilbertiodendron</emphasis>
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forest) /
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,
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General Coll. / Dérozier, V., Fouka, B., / Kirk-Spriggs, A., Takano, H. /
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:2022.14 // ANHRTUK / 00260603”
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</emphasis>
(
<specimenCount id="C781FD06FFDFFFCBFE95FA5EB8D551D0" box="[334,391,1476,1503]" count="5" pageId="2" pageNumber="216" type="male">5♂♂</specimenCount>
<specimenCount id="C781FD06FFDFFFCBFE55FA5EB89551D1" box="[398,455,1476,1502]" count="3" pageId="2" pageNumber="216" type="female">3♀♀</specimenCount>
):
</paragraph>
<paragraph id="D138368FFFDFFFCBFF1CFA72BAC752B9" blockId="2.[151,1437,1332,2043]" pageId="2" pageNumber="216">
<materialsCitation id="61EF3CD2FFDFFFCBFF1CFA72BAA2520D" box="[199,1008,1512,1539]" collectionCode="ANHRT" country="Republic of the Congo" pageId="2" pageNumber="216" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
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</emphasis>
: same data as holotype (
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)
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;
<materialsCitation id="61EF3CD2FFDFFFCBFC26FA73BAFC5229" collectingDate="2023-02-14" collectingDateMax="2023-02-20" collectingDateMin="2023-02-14" collectionCode="ANHRT" collectorName="N. Bakala &amp; V. Derozier &amp; A. Kirk-Spriggs &amp; G. Laszlo" country="Republic of the Congo" pageId="2" pageNumber="216" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
same data but
<date id="A539104FFFDFFFCBFB78FA72BC10520D" box="[1187,1346,1512,1538]" pageId="2" pageNumber="216" value="2023-02-14" valueMax="2023-02-20" valueMin="2023-02-14">
<collectingDate id="B57DE9A7FFDFFFCBFB78FA72BC10520D" box="[1187,1346,1512,1538]" pageId="2" pageNumber="216" value="2023-02-14" valueMax="2023-02-20" valueMin="2023-02-14">14–20.ii.2023</collectingDate>
</date>
, leg.
<collectorName id="7C725359FFDFFFCBFA5AFA73B9BA5229" pageId="2" pageNumber="216">N. Bakala</collectorName>
,
<collectorName id="7C725359FFDFFFCBFF2EF997B8285229" box="[245,378,1548,1575]" pageId="2" pageNumber="216">V. Dérozier</collectorName>
,
<collectorName id="7C725359FFDFFFCBFE5CF997BB6F5229" box="[391,573,1548,1575]" pageId="2" pageNumber="216">A. Kirk-Spriggs</collectorName>
&amp;
<collectorName id="7C725359FFDFFFCBFDBCF996BB855229" box="[615,727,1548,1574]" pageId="2" pageNumber="216">G. László</collectorName>
(
<specimenCount id="C781FD06FFDFFFCBFD31F996BA5F5229" box="[746,781,1548,1574]" count="1" pageId="2" pageNumber="216" type="male">1♂</specimenCount>
<specimenCount id="C781FD06FFDFFFCBFCCDF996BA6B5229" box="[790,825,1548,1574]" count="1" pageId="2" pageNumber="216" type="female">1♀</specimenCount>
<collectionCode id="B796AE4AFFDFFFCBFC99F996BAF45229" box="[834,934,1548,1574]" pageId="2" pageNumber="216">ANHRT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="61EF3CD2FFDFFFCBFC60F997B8665261" collectingDate="2022-10-10" collectingDateMax="2022-10-16" collectingDateMin="2022-10-10" collectionCode="ANHRT" collectorName="V. Derozier &amp; B. Fouka &amp; A. Kirk-Spriggs &amp; H. Takano" country="Republic of the Congo" elevation="341" latitude="2.2102501" location="Bomassa" longLatPrecision="1" longitude="16.191721" municipality="Nouabale-Ndoki National Park" pageId="2" pageNumber="216" specimenCount="1" specimenCount-female="1">
<location id="D4586054FFDFFFCBFC60F997BD725228" LSID="urn:lsid:plazi:treatment:592E8799FFDFFFCDFF4CFB56BA1E51EA:D4586054FFDFFFCBFC60F997BD725228" box="[955,1056,1549,1575]" country="Republic of the Congo" latitude="2.2102501" longLatPrecision="1" longitude="16.191721" municipality="Nouabale-Ndoki National Park" name="Bomassa" pageId="2" pageNumber="216">Bomassa</location>
camp,
<collectingMunicipality id="315CACF5FFDFFFCBFBACF996B99F5245" pageId="2" pageNumber="216">Nouabalé-Ndoki National Park</collectingMunicipality>
,
<quantity id="167F9B6AFFDFFFCBFF03F9AAB8495245" box="[216,283,1584,1610]" metricMagnitude="2" metricUnit="m" metricValue="3.41" pageId="2" pageNumber="216" unit="m" value="341.0">
<elevation id="5AAAD1BCFFDFFFCBFF03F9AAB8495245" box="[216,283,1584,1610]" metricMagnitude="2" metricUnit="m" metricValue="3.41" pageId="2" pageNumber="216" unit="m" value="341.0">341m</elevation>
</quantity>
,
<geoCoordinate id="B4B35048FFDFFFCBFEFDF9AAB8955245" box="[294,455,1584,1610]" degrees="02" direction="north" minutes="12" orientation="latitude" pageId="2" pageNumber="216" precision="1" seconds="36.9" value="2.2102501">02°12’36.9”N</geoCoordinate>
,
<geoCoordinate id="B4B35048FFDFFFCBFE09F9AABB3D5245" box="[466,623,1584,1610]" degrees="16" direction="east" minutes="11" orientation="longitude" pageId="2" pageNumber="216" precision="1" seconds="30.2" value="16.191721">16°11’30.2”E</geoCoordinate>
,
<date id="A539104FFFDFFFCBFDA1F9AABA455245" box="[634,791,1584,1610]" pageId="2" pageNumber="216" value="2022-10-10" valueMax="2022-10-16" valueMin="2022-10-10">
<collectingDate id="B57DE9A7FFDFFFCBFDA1F9AABA455245" box="[634,791,1584,1610]" pageId="2" pageNumber="216" value="2022-10-10" valueMax="2022-10-16" valueMin="2022-10-10">10–16.x.2022</collectingDate>
</date>
, leg.
<collectorName id="7C725359FFDFFFCBFC8AF9ABBA865245" box="[849,980,1584,1611]" pageId="2" pageNumber="216">V. Dérozier</collectorName>
,
<collectorName id="7C725359FFDFFFCBFC05F9ABBD1A5245" box="[990,1096,1584,1611]" pageId="2" pageNumber="216">B. Fouka</collectorName>
,
<collectorName id="7C725359FFDFFFCBFB8AF9ABBC575245" box="[1105,1285,1584,1611]" pageId="2" pageNumber="216">A. Kirk-Spriggs</collectorName>
&amp;
<collectorName id="7C725359FFDFFFCBFAF2F9ABBCCE5245" box="[1321,1436,1584,1611]" pageId="2" pageNumber="216">H. Takano</collectorName>
(
<specimenCount id="C781FD06FFDFFFCBFF78F9CEB9965261" box="[163,196,1620,1646]" count="1" pageId="2" pageNumber="216" type="female">1♀</specimenCount>
<collectionCode id="B796AE4AFFDFFFCBFF13F9CEB87E5261" box="[200,300,1620,1646]" pageId="2" pageNumber="216">ANHRT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="61EF3CD2FFDFFFCBFEE5F9CEBA85529D" collectingDate="2023-02-07" collectingDateMax="2023-02-14" collectingDateMin="2023-02-07" collectionCode="ANHRT" collectorName="N. Bakala &amp; V. Derozier &amp; A. Kirk-Spriggs &amp; G. Laszlo" country="Republic of the Congo" elevation="365" latitude="2.3640554" location="Mondika" longLatPrecision="1" longitude="16.273832" municipality="Nouabale-Ndoki National Park" pageId="2" pageNumber="216" specimenCount="1" specimenCount-male="1">
<location id="D4586054FFDFFFCBFEE5F9CEB8F15261" LSID="urn:lsid:plazi:treatment:592E8799FFDFFFCDFF4CFB56BA1E51EA:D4586054FFDFFFCBFEE5F9CEB8F15261" box="[318,419,1620,1646]" country="Republic of the Congo" latitude="2.3640554" longLatPrecision="1" longitude="16.273832" municipality="Nouabale-Ndoki National Park" name="Mondika" pageId="2" pageNumber="216">Mondika</location>
camp,
<collectingMunicipality id="315CACF5FFDFFFCBFE29F9CEBA1D5261" box="[498,847,1620,1646]" pageId="2" pageNumber="216">Nouabalé-Ndoki National Park</collectingMunicipality>
,
<quantity id="167F9B6AFFDFFFCBFC81F9CEBACE5260" box="[858,924,1620,1647]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="2" pageNumber="216" unit="m" value="365.0">
<elevation id="5AAAD1BCFFDFFFCBFC81F9CEBACE5260" box="[858,924,1620,1647]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="2" pageNumber="216" unit="m" value="365.0">365m</elevation>
</quantity>
,
<geoCoordinate id="B4B35048FFDFFFCBFC7DF9CEBD1A5260" box="[934,1096,1620,1647]" degrees="02" direction="north" minutes="21" orientation="latitude" pageId="2" pageNumber="216" precision="1" seconds="50.6" value="2.3640554">02°21’50.6”N</geoCoordinate>
,
<geoCoordinate id="B4B35048FFDFFFCBFB89F9CEBDBD5260" box="[1106,1263,1620,1647]" degrees="16" direction="east" minutes="16" orientation="longitude" pageId="2" pageNumber="216" precision="1" seconds="25.8" value="16.273832">16°16’25.8”E</geoCoordinate>
,
<date id="A539104FFFDFFFCBFB21F9CEBCCA5261" box="[1274,1432,1620,1646]" pageId="2" pageNumber="216" value="2023-02-07" valueMax="2023-02-14" valueMin="2023-02-07">
<collectingDate id="B57DE9A7FFDFFFCBFB21F9CEBCCA5261" box="[1274,1432,1620,1646]" pageId="2" pageNumber="216" value="2023-02-07" valueMax="2023-02-14" valueMin="2023-02-07">07–14.ii.2023</collectingDate>
</date>
, leg.
<collectorName id="7C725359FFDFFFCBFF11F9E3B812529D" box="[202,320,1656,1683]" pageId="2" pageNumber="216">N. Bakala</collectorName>
,
<collectorName id="7C725359FFDFFFCBFE96F9E3B880529D" box="[333,466,1656,1683]" pageId="2" pageNumber="216">V. Dérozier</collectorName>
,
<collectorName id="7C725359FFDFFFCBFE06F9E3BBC6529D" box="[477,660,1656,1683]" pageId="2" pageNumber="216">A. Kirk-Spriggs</collectorName>
&amp;
<collectorName id="7C725359FFDFFFCBFD66F9E2BA7F529D" box="[701,813,1656,1682]" pageId="2" pageNumber="216">G. László</collectorName>
(
<specimenCount id="C781FD06FFDFFFCBFC9BF9E2BA31529D" box="[832,867,1656,1682]" count="1" pageId="2" pageNumber="216" type="male">1♂</specimenCount>
<collectionCode id="B796AE4AFFDFFFCBFCB0F9E2BA9D529D" box="[875,975,1656,1682]" pageId="2" pageNumber="216">ANHRT</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="61EF3CD2FFDFFFCBFC3FF9E3BAC052B9" collectingDate="2023-04-27" collectingDateMax="2023-05-06" collectingDateMin="2023-04-27" collectionCode="ANHRT" collectorName="V. Derozier &amp; A. Kirk-Spriggs &amp; G. Laszlo &amp; S. Mvouende" country="Republic of the Congo" elevation="365" latitude="2.3640554" location="Mondika" longLatPrecision="1" longitude="16.273832" municipality="Nouabale-Ndoki National Park" pageId="2" pageNumber="216" specimenCount="1" specimenCount-male="1">
same data but
<date id="A539104FFFDFFFCBFB56F9E2BC16529D" box="[1165,1348,1656,1682]" pageId="2" pageNumber="216" value="2023-04-27" valueMax="2023-05-06" valueMin="2023-04-27">
<collectingDate id="B57DE9A7FFDFFFCBFB56F9E2BC16529D" box="[1165,1348,1656,1682]" pageId="2" pageNumber="216" value="2023-04-27" valueMax="2023-05-06" valueMin="2023-04-27">27.iv–06.v.2023</collectingDate>
</date>
, leg.
<collectorName id="7C725359FFDFFFCBFA5EF9E3B9A952B9" pageId="2" pageNumber="216">V. Dérozier</collectorName>
,
<collectorName id="7C725359FFDFFFCBFEDFF907B8EE52B9" box="[260,444,1692,1719]" pageId="2" pageNumber="216">A. Kirk-Spriggs</collectorName>
,
<collectorName id="7C725359FFDFFFCBFE1CF906BB6652B9" box="[455,564,1692,1718]" pageId="2" pageNumber="216">G. László</collectorName>
&amp;
<collectorName id="7C725359FFDFFFCBFD83F906BBBF52B9" box="[600,749,1692,1718]" pageId="2" pageNumber="216">S. Mvouende</collectorName>
(
<specimenCount id="C781FD06FFDFFFCBFD25F906BA7252B9" box="[766,800,1692,1718]" count="1" pageId="2" pageNumber="216" type="male">1♂</specimenCount>
<collectionCode id="B796AE4AFFDFFFCBFCFDF906BAD852B9" box="[806,906,1692,1718]" pageId="2" pageNumber="216">ANHRT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFDFFFCDFF1CF95ABB1C566E" lastPageId="4" lastPageNumber="218" pageId="2" pageNumber="216" type="description">
<paragraph id="D138368FFFDFFFCBFF1CF95AB80B52D5" blockId="2.[151,1437,1332,2043]" box="[199,345,1728,1754]" pageId="2" pageNumber="216">
<emphasis id="E3F3EA9DFFDFFFCBFF1CF95AB80B52D5" bold="true" box="[199,345,1728,1754]" pageId="2" pageNumber="216">Description.</emphasis>
</paragraph>
<paragraph id="D138368FFFDFFFCBFF1CF97EBDAE52F0" blockId="2.[151,1437,1332,2043]" box="[199,1276,1764,1791]" pageId="2" pageNumber="216">
Forewing length:
<typeStatus id="0E3C882DFFDFFFCBFE55F97EB8A252F1" box="[398,496,1764,1790]" pageId="2" pageNumber="216" type="holotype">holotype</typeStatus>
male:
<quantity id="167F9B6AFFDFFFCBFDE6F97EBBDD52F0" box="[573,655,1764,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="216" unit="mm" value="12.0">12 mm</quantity>
;
<typeStatus id="0E3C882DFFDFFFCBFD41F97FBBA852F0" box="[666,762,1765,1791]" pageId="2" pageNumber="216" type="paratype">paratype</typeStatus>
males:
<quantity id="167F9B6AFFDFFFCBFC8AF97EBA9F52F0" box="[849,973,1764,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="2" pageNumber="216" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
;
<typeStatus id="0E3C882DFFDFFFCBFC02F97FBD6B52F0" box="[985,1081,1765,1791]" pageId="2" pageNumber="216" type="paratype">paratype</typeStatus>
females:
<quantity id="167F9B6AFFDFFFCBFB7DF97EBDAA52F0" box="[1190,1272,1764,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="216" unit="mm" value="15.0">15 mm</quantity>
.
</paragraph>
<paragraph id="D138368FFFDFFFCDFF1CF892BA2F5586" blockId="2.[151,1437,1332,2043]" lastBlockId="4.[151,1437,151,1509]" lastPageId="4" lastPageNumber="218" pageId="2" pageNumber="216">
Male. Upperside. Ground colour of head and thorax black, abdomen dark grey. Antennae black, with white scaling ventrally at the base of each segment. Frons with a narrow row of white scales running adjacent to the eye. Palps longer than diameter of eye, black, with creamy-white scaling at base. Thorax with long, ciliate, dark greyish-brown scales anteriorly, turning paler posteriorly. Ground colour of wings creamy-white, bordered with fuscous charcoal-brown. Forewing costa evenly arcuate, rounded at apex; outer margin convex. Dark border ca.
<quantity id="167F9B6AFFDFFFCBFAC4F802BC3253BC" box="[1311,1376,1944,1971]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="216" unit="mm" value="2.0">2 mm</quantity>
wide along costa, almost
<quantity id="167F9B6AFFDFFFCBFEACF826B8EB53D8" box="[375,441,1980,2007]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="216" unit="mm" value="5.0">5 mm</quantity>
at the base of vein 5, and
<quantity id="167F9B6AFFDFFFCBFD3AF826BA7153D8" box="[737,803,1980,2007]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="216" unit="mm" value="3.0">3 mm</quantity>
along vein 2. Inner edge of border clearly-defined, with incipient tooth at the end of cell and a slight protrusion in space 2. Discal spot small and indistinct, fused with border. Hindwing rounded, with
<quantity id="167F9B6AFFD9FFCDFE61FF0DBB4854BE" box="[442,538,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="4" pageNumber="218" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
wide border terminating at tornus, suffused with paler, slightly-greyer scales. Discoidal spot of underside showing through faintly. Underside. Ground colour of head and thorax black, abdomen creamy-white. Legs black with yellow scaling at the base of each tarsal segment. Ground colour of wings the same as the upperside. Forewing border narrower along the outer margin, evenly rounded and tapering towards tornal angle. Elongate subapical marking, creamy-white, and very indistinct. Hindwing border the same width as the upperside, continuing along the length of the inner margin. Small discoidal spot the same colour as border, longitudinally ovoid, centred around the juncture of veins 6 and 7.
</paragraph>
<caption id="85F86607FFDEFFCAFF4CF8E9BBBD53F8" ID-DOI="http://doi.org/10.5281/zenodo.13346629" ID-Zenodo-Dep="13346629" httpUri="https://zenodo.org/record/13346629/files/figure.png" pageId="3" pageNumber="217" startId="3.[151,264,1907,1931]" targetBox="[152,1434,184,1870]" targetPageId="3" targetType="figure">
<paragraph id="D138368FFFDEFFCAFF4CF8E9BBBD53F8" blockId="3.[151,1437,1907,2039]" pageId="3" pageNumber="217">
<emphasis id="E3F3EA9DFFDEFFCAFF4CF8E9B8685384" bold="true" box="[151,314,1907,1931]" pageId="3" pageNumber="217">FIGURES 1–8.</emphasis>
<taxonomicName id="16874D0CFFDEFFCAFE9BF8E9B8B85384" box="[320,490,1907,1931]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="217" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E3F3EA9DFFDEFFCAFE9BF8E9B8C65384" box="[320,404,1907,1931]" italics="true" pageId="3" pageNumber="217">Falcuna</emphasis>
species.
</taxonomicName>
Adults (1–6), upperside (above) and underside (below); ♂ genitalia (7–8) in lateral view.
<emphasis id="E3F3EA9DFFDEFFCAFAAAF8E9BCD65384" bold="true" box="[1393,1412,1907,1931]" pageId="3" pageNumber="217">1.</emphasis>
<taxonomicName id="16874D0CFFDEFFCAFA51F8EEB80353A0" authority="(Staudinger)" baseAuthorityName="Staudinger" baseAuthorityYear="1892" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="217" phylum="Arthropoda" rank="species" species="lybia">
<emphasis id="E3F3EA9DFFDEFFCAFA51F8EEB99853A0" italics="true" pageId="3" pageNumber="217">F. lybia</emphasis>
(Staudinger)
</taxonomicName>
, paralectotype ♂ of
<taxonomicName id="16874D0CFFDEFFCAFDC5F80DBB9C53A0" authorityName="Staudinger" authorityYear="1892" box="[542,718,1943,1967]" class="Insecta" family="Lycaenidae" genus="Larinopoda" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="217" phylum="Arthropoda" rank="species" species="libya">
<emphasis id="E3F3EA9DFFDEFFCAFDC5F80DBB9C53A0" box="[542,718,1943,1967]" italics="true" pageId="3" pageNumber="217">Larinopoda libya</emphasis>
</taxonomicName>
(MfN) with labels.
<emphasis id="E3F3EA9DFFDEFFCAFC4CF80DBAF853A0" bold="true" box="[919,938,1943,1967]" pageId="3" pageNumber="217">2.</emphasis>
<emphasis id="E3F3EA9DFFDEFFCAFC6BF80DBA9753A0" box="[944,965,1943,1967]" italics="true" pageId="3" pageNumber="217">Id</emphasis>
., ♂, Cameroon, Nkol Nlong (RCJS).
<emphasis id="E3F3EA9DFFDEFFCAFA99F80DBC0753A0" bold="true" box="[1346,1365,1943,1967]" pageId="3" pageNumber="217">3.</emphasis>
<emphasis id="E3F3EA9DFFDEFFCAFA80F80DBC2253A0" box="[1371,1392,1943,1967]" italics="true" pageId="3" pageNumber="217">Id</emphasis>
., ♀, Cameroon, Zoétélé (RCJS).
<emphasis id="E3F3EA9DFFDEFFCAFE68F821B89453DC" bold="true" box="[435,454,1979,2003]" pageId="3" pageNumber="217">4.</emphasis>
<taxonomicName id="16874D0CFFDEFFCAFE17F826BB2653DC" authorityName="Takano" authorityYear="2024" box="[460,628,1979,2003]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="217" phylum="Arthropoda" rank="species" species="nouabaleensis" status="sp. nov.">
<emphasis id="E3F3EA9DFFDEFFCAFE17F826BB2653DC" box="[460,628,1979,2003]" italics="true" pageId="3" pageNumber="217">F. nouabaleensis</emphasis>
</taxonomicName>
<emphasis id="E3F3EA9DFFDEFFCAFDA2F821BB9B53DC" bold="true" box="[633,713,1979,2003]" pageId="3" pageNumber="217">
<taxonomicNameLabel id="F8C057E6FFDEFFCAFDA2F821BB9B53DC" box="[633,713,1979,2003]" pageId="3" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂.
<emphasis id="E3F3EA9DFFDEFFCAFC8AF821BA3653DC" bold="true" box="[849,868,1979,2003]" pageId="3" pageNumber="217">5.</emphasis>
<emphasis id="E3F3EA9DFFDEFFCAFCB2F821BA2C53DC" box="[873,894,1979,2003]" italics="true" pageId="3" pageNumber="217">Id</emphasis>
., paratype ♂ (ANHRTUK 00260604).
<emphasis id="E3F3EA9DFFDEFFCAFADDF821BC4B53DC" bold="true" box="[1286,1305,1979,2003]" pageId="3" pageNumber="217">6.</emphasis>
<emphasis id="E3F3EA9DFFDEFFCAFAC5F821BC6153DC" box="[1310,1331,1979,2003]" italics="true" pageId="3" pageNumber="217">Id</emphasis>
., paratype ♀ (ANHRTUK 00321798).
<emphasis id="E3F3EA9DFFDEFFCAFE6FF845B89553F8" bold="true" box="[436,455,2015,2039]" pageId="3" pageNumber="217">7.</emphasis>
<taxonomicName id="16874D0CFFDEFFCAFE16F87ABB4B53F8" baseAuthorityName="Staudinger" baseAuthorityYear="1892" box="[461,537,2015,2039]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="217" phylum="Arthropoda" rank="species" species="lybia">
<emphasis id="E3F3EA9DFFDEFFCAFE16F87ABB4B53F8" box="[461,537,2015,2039]" italics="true" pageId="3" pageNumber="217">F. lybia</emphasis>
</taxonomicName>
.
<emphasis id="E3F3EA9DFFDEFFCAFDFDF845BB6A53F8" bold="true" box="[550,568,2015,2039]" pageId="3" pageNumber="217">8.</emphasis>
<taxonomicName id="16874D0CFFDEFFCAFDE4F87ABBBB53F8" authorityName="Takano" authorityYear="2024" box="[575,745,2015,2039]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="217" phylum="Arthropoda" rank="species" species="nouabaleensis">
<emphasis id="E3F3EA9DFFDEFFCAFDE4F87ABBBB53F8" box="[575,745,2015,2039]" italics="true" pageId="3" pageNumber="217">F. nouabaleensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="D138368FFFD9FFCDFF1CFE09BC1A5616" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="218">Male genitalia. Uncus triangular, with weakly-depressed, pointed apex. Tegumen broad, half the width of valve. Subuncal lobe, slender, with a pair of long, slightly curved arms on the outer margin; the free end drawn out into a long, gently excurved, pointed apex. Valve oblong, apex recurved towards dorsum, with a small digitate process on the dorsal margin. Aedeagus long, slightly longer than valve, sinuate, tapering gradually from base to apex.</paragraph>
<paragraph id="D138368FFFD9FFCDFF1CFDB9BB1C566E" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="218">Female. Similar to male but larger, the forewing outer margin is more convex and the creamy-white basal area is much more extensive on both wings.</paragraph>
</subSubSection>
<subSubSection id="999D6504FFD9FFCDFF1CFDF0BD28501E" pageId="4" pageNumber="218" type="diagnosis">
<paragraph id="D138368FFFD9FFCDFF1CFDF0BD28501E" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="218">
<emphasis id="E3F3EA9DFFD9FFCDFF1CFDF0B810568B" bold="true" box="[199,322,618,644]" pageId="4" pageNumber="218">Diagnosis.</emphasis>
The new species is similar in appearance to
<taxonomicName id="16874D0CFFD9FFCDFCFEFDF1BD1E568A" authority="(Staudinger, 1892)" baseAuthorityName="Staudinger" baseAuthorityYear="1892" box="[805,1100,619,645]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="lybia">
<emphasis id="E3F3EA9DFFD9FFCDFCFEFDF1BA25568A" box="[805,887,619,645]" italics="true" pageId="4" pageNumber="218">F. lybia</emphasis>
(
<bibRefCitation id="B5164B7EFFD9FFCDFC5FFDF1BD11568A" author="Staudinger, O." box="[900,1091,619,645]" pageId="4" pageNumber="218" pagination="215 - 223" refId="ref11393" refString="Staudinger, O. (1892) Neue afrikanische Lycaeniden. Deutsche entomologische Zeitschrift, Iris, 4 (2), 215 - 223." type="journal article" year="1892">Staudinger, 1892</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="49BC2A0AFFD9FFCDFB82FDF1BDE9568A" box="[1113,1211,619,645]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1907,1931]" captionTargetBox="[152,1434,184,1870]" captionTargetId="figure-30@3.[151,1435,181,1883]" captionTargetPageId="3" captionText="FIGURES 1–8. Falcuna species. Adults (1–6), upperside (above) and underside (below); ♂ genitalia (7–8) in lateral view. 1. F. lybia (Staudinger), paralectotype ♂ of Larinopoda libya (MfN) with labels. 2. Id., ♂, Cameroon, Nkol Nlong (RCJS). 3. Id., ♀, Cameroon, Zoétélé (RCJS). 4. F. nouabaleensis sp. nov., holotype ♂. 5. Id., paratype ♂ (ANHRTUK 00260604). 6. Id., paratype ♀ (ANHRTUK 00321798). 7. F. lybia. 8. F. nouabaleensis." figureDoi="http://doi.org/10.5281/zenodo.13346629" httpUri="https://zenodo.org/record/13346629/files/figure.png" pageId="4" pageNumber="218">Figs 1–3</figureCitation>
), a peculiar member of the genus distinguished by a single discoidal spot on the hindwing underside, somewhat reminiscent of the white
<taxonomicName id="16874D0CFFD9FFCDFF4CFD29B89456C2" authority="Westwood, [1851]" authorityName="Westwood" authorityYear="1851" box="[151,454,690,717]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="E3F3EA9DFFD9FFCDFF4CFD29B9BF56C3" box="[151,237,691,716]" italics="true" pageId="4" pageNumber="218">Liptena</emphasis>
Westwood, [1851]
</taxonomicName>
in the
<taxonomicName id="16874D0CFFD9FFCDFDC1FD29BB2C56C3" baseAuthorityName="Kirby" baseAuthorityYear="1890" box="[538,638,691,716]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="E3F3EA9DFFD9FFCDFDC1FD29BB2C56C3" box="[538,638,691,716]" italics="true" pageId="4" pageNumber="218">L. opaca</emphasis>
</taxonomicName>
group of
<bibRefCitation id="B5164B7EFFD9FFCDFD29FD29BADC56C2" author="Libert, M." box="[754,910,691,717]" pageId="4" pageNumber="218" pagination="367 - 396" refId="ref10890" refString="Libert, M. (2021) Nouveaux taxons et synonymies dans le groupe de Liptena opaca (Kirby, 1890) (Lepidoptera, Lycaenidae, Poritiinae). Bulletin de la Societe entomologique de France, 126 (3), 367 - 396. https: // doi. org / 10.32475 / bsef _ 2154" type="journal article" year="2021">Libert (2021)</bibRefCitation>
. The genitalia are also distinctive, the slender construction of the subuncal lobes being strongly excurved anterio-ventrally and lacking projections from the inner surface (
<bibRefCitation id="B5164B7EFFD9FFCDFF23FD61BB79571A" author="Stempffer, H. &amp; Bennett, N. H." box="[248,555,763,789]" pageId="4" pageNumber="218" pagination="171 - 194" refId="ref11481" refString="Stempffer, H. &amp; Bennett, N. H. (1963) A new genus of Lipteninae (Lepidoptera: Lycaenidae). Bulletin of the British Museum (Natural History): Entomology, 13 (6), 171 - 194." type="journal article" year="1963">Stempffer &amp; Bennett 1963</bibRefCitation>
).
<taxonomicName id="16874D0CFFD9FFCDFD9BFD61BA17571A" authorityName="Takano" authorityYear="2024" box="[576,837,763,789]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="nouabaleensis" status="sp. nov.">
<emphasis id="E3F3EA9DFFD9FFCDFD9BFD61BA17571A" box="[576,837,763,789]" italics="true" pageId="4" pageNumber="218">Falcuna nouabaleensis</emphasis>
</taxonomicName>
<emphasis id="E3F3EA9DFFD9FFCDFC95FD61BAFB571A" bold="true" box="[846,937,763,789]" pageId="4" pageNumber="218">
<taxonomicNameLabel id="F8C057E6FFD9FFCDFC95FD61BAFB571A" box="[846,937,763,789]" pageId="4" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can most readily be separated from
<taxonomicName id="16874D0CFFD9FFCDFA92FD61BCCE571A" baseAuthorityName="Staudinger" baseAuthorityYear="1892" box="[1353,1436,763,789]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="lybia">
<emphasis id="E3F3EA9DFFD9FFCDFA92FD61BCCE571A" box="[1353,1436,763,789]" italics="true" pageId="4" pageNumber="218">F. lybia</emphasis>
</taxonomicName>
based on the presence of subapical and apical white spots on the forewing and hindwing undersides respectively, the broader hindwing border, and the larger discoidal spot of the hindwing in the latter. The ground plan of the genital capsule is similar but in
<taxonomicName id="16874D0CFFD9FFCDFE6AFCFDBB55578E" baseAuthorityName="Staudinger" baseAuthorityYear="1892" box="[433,519,871,897]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="lybia">
<emphasis id="E3F3EA9DFFD9FFCDFE6AFCFDBB55578E" box="[433,519,871,897]" italics="true" pageId="4" pageNumber="218">F. lybia</emphasis>
</taxonomicName>
(
<figureCitation id="49BC2A0AFFD9FFCDFDC3FCFDBB0C578E" box="[536,606,871,897]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1907,1931]" captionTargetBox="[152,1434,184,1870]" captionTargetId="figure-30@3.[151,1435,181,1883]" captionTargetPageId="3" captionText="FIGURES 1–8. Falcuna species. Adults (1–6), upperside (above) and underside (below); ♂ genitalia (7–8) in lateral view. 1. F. lybia (Staudinger), paralectotype ♂ of Larinopoda libya (MfN) with labels. 2. Id., ♂, Cameroon, Nkol Nlong (RCJS). 3. Id., ♀, Cameroon, Zoétélé (RCJS). 4. F. nouabaleensis sp. nov., holotype ♂. 5. Id., paratype ♂ (ANHRTUK 00260604). 6. Id., paratype ♀ (ANHRTUK 00321798). 7. F. lybia. 8. F. nouabaleensis." figureDoi="http://doi.org/10.5281/zenodo.13346629" httpUri="https://zenodo.org/record/13346629/files/figure.png" pageId="4" pageNumber="218">Fig. 7</figureCitation>
), it is generally larger, the subuncal lobes are cranially shorter and more robust, and caudally more excurved, the valve is more elongate, its dorsal margin straighter and the recurved apex broader. Intraspecific barcode PWDs of the new species were 0.0% (n=3) and diverged from a
<taxonomicName id="16874D0CFFD9FFCDFB17FC35BC7357C6" baseAuthorityName="Staudinger" baseAuthorityYear="1892" box="[1228,1313,943,969]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="lybia">
<emphasis id="E3F3EA9DFFD9FFCDFB17FC35BC7357C6" box="[1228,1313,943,969]" italics="true" pageId="4" pageNumber="218">F. lybia</emphasis>
</taxonomicName>
sample on GenBank (MCZ:RE08J104) by 1.6%. This is a greater divergence than that observed between GenBank sequences of
<taxonomicName id="16874D0CFFD9FFCDFF6DFC6DB8BD501E" authority="(Suffert, 1904)" baseAuthorityName="Suffert" baseAuthorityYear="1904" box="[182,495,1015,1041]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="margarita">
<emphasis id="E3F3EA9DFFD9FFCDFF6DFC6DB810501F" box="[182,322,1015,1040]" italics="true" pageId="4" pageNumber="218">F. margarita</emphasis>
(
<bibRefCitation id="B5164B7EFFD9FFCDFE8BFC6DB8B4501E" author="Suffert, E." box="[336,486,1015,1041]" pageId="4" pageNumber="218" pagination="12 - 107" refId="ref11527" refString="Suffert, E. (1904) Neue afrikanische Tagfalter aus dem kon. zool. Museum, Berlin, und meiner Sammlung. Deutsche entomologische Zeitschrift, Iris, 17 (1), 12 - 107." type="journal article" year="1904">Suffert, 1904</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="16874D0CFFD9FFCDFDFEFC6DBAE6501E" authority="(Bethune-Baker, 1906)" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1906" box="[549,948,1015,1041]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="E3F3EA9DFFD9FFCDFDFEFC6DBBF9501E" box="[549,683,1015,1041]" italics="true" pageId="4" pageNumber="218">F. orientalis</emphasis>
(
<bibRefCitation id="B5164B7EFFD9FFCDFD61FC6DBAF9501E" author="Bethune-Baker, G. T." box="[698,939,1015,1041]" pageId="4" pageNumber="218" pagination="339 - 346" refId="ref9526" refString="Bethune-Baker, G. T. (1906) Descriptions of African Lepidoptera. Annals and Magazine of Natural History, Series 7, 18 (107), 339 - 346. https: // doi. org / 10.1080 / 00222930608562623" type="journal article" year="1906">Bethune-Baker, 1906</bibRefCitation>
)
</taxonomicName>
which was 1.1%.
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFD9FFCDFF1CFB81B8885056" pageId="4" pageNumber="218" type="etymology">
<paragraph id="D138368FFFD9FFCDFF1CFB81B8885056" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="218">
<emphasis id="E3F3EA9DFFD9FFCDFF1CFB81B8F0503A" bold="true" box="[199,418,1051,1077]" pageId="4" pageNumber="218">
<emphasis id="E3F3EA9DFFD9FFCDFF1CFB81B8C9503A" bold="true" box="[199,411,1051,1077]" italics="true" pageId="4" pageNumber="218">Derivatio nominis</emphasis>
.
</emphasis>
The new species is named after its
<typeStatus id="0E3C882DFFD9FFCDFC89FB81BAD0503A" box="[850,898,1051,1077]" pageId="4" pageNumber="218">type</typeStatus>
locality, Parc National de Nouabalé-Ndoki in northern
<collectingCountry id="A990761FFFD9FFCDFF26FBA5B8875056" box="[253,469,1087,1113]" name="Republic of the Congo" pageId="4" pageNumber="218">Republic of Congo</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="999D6504FFD9FFCDFF1CFBF8BA1E51EA" pageId="4" pageNumber="218" type="distribution">
<paragraph id="D138368FFFD9FFCDFF1CFBF8BA1E51EA" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="218">
<emphasis id="E3F3EA9DFFD9FFCDFF1CFBF8B8335073" bold="true" box="[199,353,1122,1148]" pageId="4" pageNumber="218">Distribution.</emphasis>
The new species is only known from the short series collected at the type locality. Its sister species,
<taxonomicName id="16874D0CFFD9FFCDFF2DFB1DB81B50AE" baseAuthorityName="Staudinger" baseAuthorityYear="1892" box="[246,329,1159,1185]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="lybia">
<emphasis id="E3F3EA9DFFD9FFCDFF2DFB1DB81B50AE" box="[246,329,1159,1185]" italics="true" pageId="4" pageNumber="218">F. lybia</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="B5164B7EFFD9FFCDFDCDFB1DBBB250AE" author="Staudinger, O." box="[534,736,1159,1185]" pageId="4" pageNumber="218" pagination="215 - 223" refId="ref11393" refString="Staudinger, O. (1892) Neue afrikanische Lycaeniden. Deutsche entomologische Zeitschrift, Iris, 4 (2), 215 - 223." type="journal article" year="1892">Staudinger (1892)</bibRefCitation>
from a small series of both sexes collected in
<collectingCountry id="A990761FFFD9FFCDFB39FB1DBC7F50AE" box="[1250,1325,1159,1185]" name="Gabon" pageId="4" pageNumber="218">Gabon</collectingCountry>
by Albert Mocquerys (see
<figureCitation id="49BC2A0AFFD9FFCDFE8BFB31B8C050CA" box="[336,402,1195,1221]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1907,1931]" captionTargetBox="[152,1434,184,1870]" captionTargetId="figure-30@3.[151,1435,181,1883]" captionTargetPageId="3" captionText="FIGURES 1–8. Falcuna species. Adults (1–6), upperside (above) and underside (below); ♂ genitalia (7–8) in lateral view. 1. F. lybia (Staudinger), paralectotype ♂ of Larinopoda libya (MfN) with labels. 2. Id., ♂, Cameroon, Nkol Nlong (RCJS). 3. Id., ♀, Cameroon, Zoétélé (RCJS). 4. F. nouabaleensis sp. nov., holotype ♂. 5. Id., paratype ♂ (ANHRTUK 00260604). 6. Id., paratype ♀ (ANHRTUK 00321798). 7. F. lybia. 8. F. nouabaleensis." figureDoi="http://doi.org/10.5281/zenodo.13346629" httpUri="https://zenodo.org/record/13346629/files/figure.png" pageId="4" pageNumber="218">Fig. 1</figureCitation>
for
<typeStatus id="0E3C882DFFD9FFCDFE1BFB31BB0550CA" box="[448,599,1195,1221]" pageId="4" pageNumber="218" type="paralectotype">paralectotype</typeStatus>
in MfN), and is known from
<collectingCountry id="A990761FFFD9FFCDFC79FB31BD4550CA" box="[930,1047,1195,1221]" name="Cameroon" pageId="4" pageNumber="218">Cameroon</collectingCountry>
(a long series in ABRI), as well as from Etoumbi, Kellé and Ketta in
<collectingCountry id="A990761FFFD9FFCDFDD4FB55BB1B50E6" box="[527,585,1231,1257]" name="Taiwan" pageId="4" pageNumber="218">ROC</collectingCountry>
(
<bibRefCitation id="B5164B7EFFD9FFCDFD8DFB55BA6E50E6" author="Bampton, I. &amp; Collins, S. C. &amp; Dowsett, R. J." box="[598,828,1231,1257]" pageId="4" pageNumber="218" pagination="57 - 108" refId="ref9368" refString="Bampton, I., Collins, S. C. &amp; Dowsett, R. J. (1991) Butterflies (Lepidoptera) of the Mayombe and lower Kouilu (with a checklist for Congo). Tauraco Research Report, 4, 57 - 108." type="journal article" year="1991">
Bampton
<emphasis id="E3F3EA9DFFD9FFCDFD1FFB55BBA650E6" box="[708,756,1231,1257]" italics="true" pageId="4" pageNumber="218">et al</emphasis>
. 1991
</bibRefCitation>
). These
<collectingCountry id="A990761FFFD9FFCDFC4DFB55BA8250E6" box="[918,976,1231,1257]" name="Taiwan" pageId="4" pageNumber="218">ROC</collectingCountry>
records are based on specimens collected by T.H.E. Jackson in the National Museum of
<collectingCountry id="A990761FFFD9FFCDFD49FB69BB895102" box="[658,731,1267,1293]" name="Kenya" pageId="4" pageNumber="218">Kenya</collectingCountry>
but since they have not been examined, it is not known to which species they belong. Despite the close proximity of Ketta to PNNN, it may be possible to speculate that all these specimens belong to
<taxonomicName id="16874D0CFFD9FFCDFE5EFAA1B88B515A" baseAuthorityName="Staudinger" baseAuthorityYear="1892" box="[389,473,1339,1365]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="lybia">
<emphasis id="E3F3EA9DFFD9FFCDFE5EFAA1B88B515A" box="[389,473,1339,1365]" italics="true" pageId="4" pageNumber="218">F. lybia</emphasis>
</taxonomicName>
;
<taxonomicName id="16874D0CFFD9FFCDFE32FAA1BB3B515A" authorityName="Leach" authorityYear="1815" box="[489,617,1339,1365]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="family">Lycaenidae</taxonomicName>
and in particular the
<taxonomicName id="16874D0CFFD9FFCDFC81FAA1BA93515A" box="[858,961,1339,1365]" class="Insecta" family="Lycaenidae" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="tribe" tribe="Liptenini">Liptenini</taxonomicName>
were a speciality of Jackson’s (
<bibRefCitation id="B5164B7EFFD9FFCDFAFCFAA1B8115176" author="Carcasson, R. H. &amp; Clench, H. K." pageId="4" pageNumber="218" pagination="131 - 134" refId="ref9759" refString="Carcasson, R. H. &amp; Clench, H. K. (1969) Obituary. Thomas Herbert Elliot Jackson (1903 - 1968). Journal of the Lepidopterists' Society, 23 (2), 131 - 134." type="journal article" year="1969">Carcasson &amp; Clench 1969</bibRefCitation>
) and had the underside lacked the white subapical and apical spots on his specimens, he most likely would have noticed such differences in these distinctive insects. Taking into consideration the evolutionary history and biogeography of this region (discussed further below), it is believed these two species are allopatric vicariants and
<taxonomicName id="16874D0CFFD9FFCDFF1CFA51B8D051EA" authorityName="Takano" authorityYear="2024" box="[199,386,1483,1509]" class="Insecta" family="Lycaenidae" genus="Falcuna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="218" phylum="Arthropoda" rank="species" species="nouabaleensis" status="sp. nov.">
<emphasis id="E3F3EA9DFFD9FFCDFF1CFA51B8D051EA" box="[199,386,1483,1509]" italics="true" pageId="4" pageNumber="218">F. nouabaleensis</emphasis>
</taxonomicName>
<emphasis id="E3F3EA9DFFD9FFCDFE52FA51B8B151EA" bold="true" box="[393,483,1483,1509]" pageId="4" pageNumber="218">
<taxonomicNameLabel id="F8C057E6FFD9FFCDFE52FA51B8B151EA" box="[393,483,1483,1509]" pageId="4" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is likely endemic to this region.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart id="0708D519342760247BBF6D3B013B0BFA">Gonçalves, Clayton Corrêa</mods:namePart>
<mods:nameIdentifier id="9CB717B52A646B9B17A4425F75783A99" type="ORCID">0000-0003-4588-4236</mods:nameIdentifier>
<mods:nameIdentifier id="4B25E1F78CEF4FEE8EE6D48AF6D8C434" type="email">clayton.correa.goncalves@gmail.com</mods:nameIdentifier>
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<mods:namePart id="913088D59BD009CAFF936B5BBE53EE42">Takiya, Daniela Maeda</mods:namePart>
<mods:nameIdentifier id="6BC90F477DB02B35C48232E9E80B6E36" type="ORCID">0000-0003-4588-4236</mods:nameIdentifier>
<mods:nameIdentifier id="886618E4F8A889816ACBDD2782E559DE" type="email">takiya@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="1D83AB52242112BA6667C8EB9B13B0D2">text</mods:typeOfResource>
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<mods:title id="F23D7DF6725CD9372044178892F4394F">Zootaxa</mods:title>
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<mods:part id="1527A07E06A971FA74FB3FD87573A8E2">
<mods:date id="ABDAC6D350862D2010C042E9567464F1">2024</mods:date>
<mods:detail id="124E9BFBB694E0EADBC8BD9FEBEC3828" type="pubDate">
<mods:number id="31F61CEF0DEED295F1F33D61A5F447C9">2024-08-20</mods:number>
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<mods:detail id="AB77DE019A192363D56DA0E71AB8B0C2" type="volume">
<mods:number id="6A69A736C5FF7823AB98D87516B5BC8E">5496</mods:number>
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<mods:number id="04284D19B9F5C9CF427610A2C1040647">3</mods:number>
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<mods:start id="C1CF31128164B2D8D25C8D8B7CCC7B3F">441</mods:start>
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<mods:url id="61BCEA64124210979BB4C6ABBACCA3DB">http://dx.doi.org/10.11646/zootaxa.5496.3.9</mods:url>
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<mods:classification id="B324FD49ED2C386FBF3E39EBFF2F6D14">journal article</mods:classification>
<mods:identifier id="621ACF728BC3E6F163B7BE5D9A8A11BA" type="DOI">10.11646/zootaxa.5496.3.9</mods:identifier>
<mods:identifier id="8DA9A7FF8BE1DEFE7C663FB7335682FF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="242A4044145028E0FBE97F6B3FE36790" type="Zenodo-Dep">13347001</mods:identifier>
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<treatment id="594B87B1FF98FF9EFF0FFE73D849F83F" ID-DOI="http://doi.org/10.5281/zenodo.13346997" ID-Zenodo-Dep="13346997" LSID="urn:lsid:plazi:treatment:594B87B1FF98FF9EFF0FFE73D849F83F" httpUri="http://treatment.plazi.org/id/594B87B1FF98FF9EFF0FFE73D849F83F" lastPageId="6" lastPageNumber="447" pageId="5" pageNumber="446">
<subSubSection id="99F8652CFF98FF9DFF0FFE73DB0BFE3A" box="[151,523,402,429]" pageId="5" pageNumber="446" type="nomenclature">
<paragraph id="D15D36A7FF98FF9DFF0FFE73DB0BFE3A" blockId="5.[151,523,402,465]" box="[151,523,402,429]" pageId="5" pageNumber="446">
<heading id="8A1581CBFF98FF9DFF0FFE73DB0BFE3A" bold="true" box="[151,523,402,429]" fontSize="11" level="1" pageId="5" pageNumber="446" reason="1">
<emphasis id="E396EAB5FF98FF9DFF0FFE73DB0BFE3A" bold="true" box="[151,523,402,429]" pageId="5" pageNumber="446">
<taxonomicName id="16E24D24FF98FF9DFF0FFE73D8A7FE3B" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[151,423,402,429]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="tetrarama" status="sp. nov.">
<emphasis id="E396EAB5FF98FF9DFF0FFE73D8A7FE3B" bold="true" box="[151,423,402,429]" italics="true" pageId="5" pageNumber="446">Gracilipona tetrarama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A557CEFF98FF9DFE36FE73DB0BFE3A" box="[430,523,402,428]" pageId="5" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="99F8652CFF98FF9DFF0FFE57D85CFE47" box="[151,348,438,465]" pageId="5" pageNumber="446" type="description">
<paragraph id="D15D36A7FF98FF9DFF0FFE57D85CFE47" blockId="5.[151,523,402,465]" box="[151,348,438,465]" pageId="5" pageNumber="446">
(
<figureCitation id="49D92A22FF98FF9DFF07FE57D9E7FE46" box="[159,231,438,464]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Figs 3</figureCitation>
,
<figureCitation id="49D92A22FF98FF9DFF6CFE57D804FE46" box="[244,260,438,464]" captionStart="FIGURE 4" captionStartId="7.[151,250,1464,1488]" captionTargetBox="[151,1434,181,1431]" captionTargetId="figure-20@7.[151,1435,181,1440]" captionTargetPageId="7" captionText="FIGURE 4. Gracilipona tetrarama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347009" httpUri="https://zenodo.org/record/13347009/files/figure.png" pageId="5" pageNumber="446">4</figureCitation>
,
<figureCitation id="49D92A22FF98FF9DFE88FE57D854FE47" box="[272,340,438,465]" captionStart="FIGURE 5" captionStartId="8.[151,250,1189,1213]" captionTargetBox="[151,1436,491,1165]" captionTargetId="figure-654@8.[151,1436,491,1165]" captionTargetPageId="8" captionText="FIGURE 5. Habitus, dorsal view (left) and lateral view (right). A, B, Gracilipona birama sp. nov., holotype male. C, D, Gracilipona tetrarama sp. nov., holotype male. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347011" httpUri="https://zenodo.org/record/13347011/files/figure.png" pageId="5" pageNumber="446">5C–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="99F8652CFF98FF9DFF0FFE1FDA5BFD15" pageId="5" pageNumber="446" type="diagnosis">
<paragraph id="D15D36A7FF98FF9DFF0FFE1FDA5BFD15" blockId="5.[151,1437,510,1930]" pageId="5" pageNumber="446">
<emphasis id="E396EAB5FF98FF9DFF0FFE1FD812FD8E" bold="true" box="[151,274,510,536]" pageId="5" pageNumber="446">Diagnosis.</emphasis>
Style (
<figureCitation id="49D92A22FF98FF9DFEFDFE1FD8BFFD8E" box="[357,447,510,536]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Fig. 3K</figureCitation>
) blade with ventral margin expanded at apical two-thirds; dorsal margin rounded near base; apex thin, weakly directed dorsally; aedeagus (
<figureCitation id="49D92A22FF98FF9DFD6EFDC3DA79FDAA" box="[758,889,546,572]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Figs 3L–M</figureCitation>
) shaft short with two pairs of processes arising ventrally: one pair as long as shaft and the other pair two times longer than shaft length; female sternite VII (
<figureCitation id="49D92A22FF98FF9DFAF7FDA7D9BAFD12" captionStart="FIGURE 4" captionStartId="7.[151,250,1464,1488]" captionTargetBox="[151,1434,181,1431]" captionTargetId="figure-20@7.[151,1435,181,1440]" captionTargetPageId="7" captionText="FIGURE 4. Gracilipona tetrarama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347009" httpUri="https://zenodo.org/record/13347009/files/figure.png" pageId="5" pageNumber="446">Fig. 4A</figureCitation>
) with posterolateral angles rounded, not produced posterad.
</paragraph>
</subSubSection>
<subSubSection id="99F8652CFF98FF9DFF5FFD6CDDF2FB75" pageId="5" pageNumber="446" type="description">
<paragraph id="D15D36A7FF98FF9DFF5FFD6CD9F3FD5D" blockId="5.[151,1437,510,1930]" pageId="5" pageNumber="446">
<emphasis id="E396EAB5FF98FF9DFF5FFD6CD8C5FD31" bold="true" box="[199,453,653,679]" pageId="5" pageNumber="446">Measurements (mm).</emphasis>
<typeStatus id="0E598805FF98FF9DFE53FD6CDB33FD31" box="[459,563,653,679]" pageId="5" pageNumber="446" type="holotype">Holotype</typeStatus>
male: total length 8.0.
<typeStatus id="0E598805FF98FF9DFCAAFD6FDAA3FD3E" box="[818,931,654,680]" pageId="5" pageNumber="446" type="paratype">Paratypes</typeStatus>
: males (n = 6),
<quantity id="161A9B42FF98FF9DFBD4FD6CDDD6FD3E" box="[1100,1238,653,680]" metricMagnitude="-3" metricUnit="m" metricValue="7.55" metricValueMax="7.8" metricValueMin="7.3" pageId="5" pageNumber="446" unit="mm" value="7.55" valueMax="7.8" valueMin="7.3">7.3–7.8 mm</quantity>
; females (n = 11), 8.0–9.0.
</paragraph>
<paragraph id="D15D36A7FF98FF9DFF5FFD34D875FBC2" blockId="5.[151,1437,510,1930]" pageId="5" pageNumber="446">
<emphasis id="E396EAB5FF98FF9DFF5FFD34D88FFD78" bold="true" box="[199,399,724,751]" pageId="5" pageNumber="446">Male terminalia.</emphasis>
Sternite VIII (
<figureCitation id="49D92A22FF98FF9DFDADFD34DB8AFD79" box="[565,650,725,751]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Fig. 3E</figureCitation>
) in ventral view, as wide as long; lateral margins straight and slightly tapering to apex; posterior margin rounded. Valve (
<figureCitation id="49D92A22FF98FF9DFD45FD19DA32FC84" box="[733,818,760,786]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Fig. 3F</figureCitation>
) in ventral view, 1.7x wider than long; ventral margin rounded. Pygofer (
<figureCitation id="49D92A22FF98FF9DFEF2FCFDD8C2FCA0" box="[362,450,796,822]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Fig. 3G</figureCitation>
) in lateral view, 1.8x longer than high, ventral margin broadly rounded; apex subacute. Subgenital plate (
<figureCitation id="49D92A22FF98FF9DFEC7FCA1D8BBFCCC" box="[351,443,832,858]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Fig. 3H</figureCitation>
), in ventral view, approximately 5.5x longer than wide; lateral margins nearly parallel; apex rounded. Connective (
<figureCitation id="49D92A22FF98FF9DFE4CFC82DB22FCE8" box="[468,546,867,894]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Fig. 3I</figureCitation>
) in dorsal view, with arms slightly longer than stem; stem narrow, with well developed dorsal keel. Style (
<figureCitation id="49D92A22FF98FF9DFE46FC66DB2BFC37" box="[478,555,903,929]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Fig. 3J</figureCitation>
) in dorsal view, with outer lobe developed; in lateral view (
<figureCitation id="49D92A22FF98FF9DFB24FC66DC15FC37" box="[1212,1301,903,929]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Fig. 3K</figureCitation>
), blade with ventral margin expanded at apical two-thirds; dorsal margin rounded near base; apex thin, acute weakly directed dorsally. Aedeagus (
<figureCitation id="49D92A22FF98FF9DFEE2FC2ED8FBFC7F" box="[378,507,975,1001]" captionStart="FIGURE 3" captionStartId="6.[151,250,1649,1673]" captionTargetBox="[154,1435,186,1620]" captionTargetId="figure-28@6.[151,1435,181,1625]" captionTargetPageId="6" captionText="FIGURE 3. Gracilipona tetrarama sp. nov., holotype male.A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="5" pageNumber="446">Figs 3L–M</figureCitation>
) shaft short, with two pairs of processes arising ventrally: one pair thin, as long as shaft, parallel and adjacent to shaft, and the other pair wider, two times longer than shaft length, adjacent to shaft at basal third and then abruptly directed posterodorsally; apex of shaft without processes. Other characteristics as in generic description.
</paragraph>
<paragraph id="D15D36A7FF98FF9DFF5FFBBFDDF2FB75" blockId="5.[151,1437,510,1930]" pageId="5" pageNumber="446">
<emphasis id="E396EAB5FF98FF9DFF5FFBBFD8AEFBE1" bold="true" box="[199,430,1117,1144]" pageId="5" pageNumber="446">Female terminalia.</emphasis>
Sternite VII (
<figureCitation id="49D92A22FF98FF9DFDC0FBBFDBB7FBEE" box="[600,695,1118,1144]" captionStart="FIGURE 4" captionStartId="7.[151,250,1464,1488]" captionTargetBox="[151,1434,181,1431]" captionTargetId="figure-20@7.[151,1435,181,1440]" captionTargetPageId="7" captionText="FIGURE 4. Gracilipona tetrarama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347009" httpUri="https://zenodo.org/record/13347009/files/figure.png" pageId="5" pageNumber="446">Fig. 4A</figureCitation>
), in ventral view, 1.4x wider than long; posterolateral margins rounded, not produced posterad; posterior margin weakly rounded. First valvifer (
<figureCitation id="49D92A22FF98FF9DFBAEFB60DD8DFB0A" box="[1078,1165,1153,1180]" captionStart="FIGURE 4" captionStartId="7.[151,250,1464,1488]" captionTargetBox="[151,1434,181,1431]" captionTargetId="figure-20@7.[151,1435,181,1440]" captionTargetPageId="7" captionText="FIGURE 4. Gracilipona tetrarama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347009" httpUri="https://zenodo.org/record/13347009/files/figure.png" pageId="5" pageNumber="446">Fig. 4C</figureCitation>
) 1.5x higher than wide. First valvula (
<figureCitation id="49D92A22FF98FF9DFEAAFB44D888FB29" box="[306,392,1189,1215]" captionStart="FIGURE 4" captionStartId="7.[151,250,1464,1488]" captionTargetBox="[151,1434,181,1431]" captionTargetId="figure-20@7.[151,1435,181,1440]" captionTargetPageId="7" captionText="FIGURE 4. Gracilipona tetrarama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347009" httpUri="https://zenodo.org/record/13347009/files/figure.png" pageId="5" pageNumber="446">Fig. 4C</figureCitation>
) 4x longer than high. Second valvula (
<figureCitation id="49D92A22FF98FF9DFCAFFB44DA8AFB29" box="[823,906,1189,1215]" captionStart="FIGURE 4" captionStartId="7.[151,250,1464,1488]" captionTargetBox="[151,1434,181,1431]" captionTargetId="figure-20@7.[151,1435,181,1440]" captionTargetPageId="7" captionText="FIGURE 4. Gracilipona tetrarama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347009" httpUri="https://zenodo.org/record/13347009/files/figure.png" pageId="5" pageNumber="446">Fig. 4F</figureCitation>
) 3.5x longer than high; apical portion (
<figureCitation id="49D92A22FF98FF9DFAA5FB44DC94FB29" box="[1341,1428,1189,1215]" captionStart="FIGURE 4" captionStartId="7.[151,250,1464,1488]" captionTargetBox="[151,1434,181,1431]" captionTargetId="figure-20@7.[151,1435,181,1440]" captionTargetPageId="7" captionText="FIGURE 4. Gracilipona tetrarama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347009" httpUri="https://zenodo.org/record/13347009/files/figure.png" pageId="5" pageNumber="446">Fig. 4G</figureCitation>
) dorsal margin with approximately ten rounded denticles. Gonoplac (
<figureCitation id="49D92A22FF98FF9DFC0CFB28DAEBFB75" box="[916,1003,1225,1251]" captionStart="FIGURE 4" captionStartId="7.[151,250,1464,1488]" captionTargetBox="[151,1434,181,1431]" captionTargetId="figure-20@7.[151,1435,181,1440]" captionTargetPageId="7" captionText="FIGURE 4. Gracilipona tetrarama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347009" httpUri="https://zenodo.org/record/13347009/files/figure.png" pageId="5" pageNumber="446">Fig. 4H</figureCitation>
) 3.3x longer than high.
</paragraph>
</subSubSection>
<subSubSection id="99F8652CFF98FF9DFF5FFB0CD805FABC" pageId="5" pageNumber="446" type="etymology">
<paragraph id="D15D36A7FF98FF9DFF5FFB0CD805FABC" blockId="5.[151,1437,510,1930]" pageId="5" pageNumber="446">
<emphasis id="E396EAB5FF98FF9DFF5FFB0CD84EFA91" bold="true" box="[199,334,1261,1287]" pageId="5" pageNumber="446">Etymology.</emphasis>
The specific epithet
<taxonomicName id="16E24D24FF98FF9DFDDCFB0CDBB4FA90" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[580,692,1261,1286]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="tetrarama">
<emphasis id="E396EAB5FF98FF9DFDDCFB0CDBB4FA90" box="[580,692,1261,1286]" italics="true" pageId="5" pageNumber="446">tetrarama</emphasis>
</taxonomicName>
(noun in apposition), refers to the two pairs of processes in the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="99F8652CFF98FF9DFF5FFAD5D837F81C" pageId="5" pageNumber="446" type="materials_examined">
<paragraph id="D15D36A7FF98FF9DFF5FFAD5D837F81C" blockId="5.[151,1437,510,1930]" pageId="5" pageNumber="446">
<emphasis id="E396EAB5FF98FF9DFF5FFAD5D8B3FAD8" bold="true" box="[199,435,1332,1358]" pageId="5" pageNumber="446">Material examined.</emphasis>
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<typeStatus id="0E598805FF98FF9DFE23FAD5DB22FAD8" box="[443,546,1332,1358]" pageId="5" pageNumber="446" type="holotype">holotype</typeStatus>
</emphasis>
male:
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<collectingRegion id="1326F845FF98FF9DFDEAFAD5DA28FAD8" box="[626,808,1332,1358]" country="Brazil" name="Rio de Janeiro" pageId="5" pageNumber="446">Rio de Janeiro</collectingRegion>
:
</emphasis>
“P. N.
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, RJ,
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\
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luz \
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.
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.
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leg.” (
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:
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:
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</emphasis>
♀, “
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: RJ,
<collectorName id="7C175371FF98FF9DFAD0FAB9DC99FAE4" box="[1352,1433,1368,1394]" pageId="5" pageNumber="446">Itatiaia</collectorName>
,
<collectorName id="7C175371FF98FF9DFF0FFA9DD9E5FA00" box="[151,229,1404,1430]" pageId="5" pageNumber="446">Parque</collectorName>
\
<collectingCounty id="383C4E2BFF98FF9DFF63FA9AD8DCFA03" box="[251,476,1403,1429]" pageId="5" pageNumber="446">Nacional do Itatiaia</collectingCounty>
,
<collectingMunicipality id="3139ACDDFF98FF9DFE7FFA9DDB1DFA00" box="[487,541,1404,1430]" pageId="5" pageNumber="446">Casa</collectingMunicipality>
do \
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2, -22.452500, \ -44.610681,
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, light, 12- \
<date id="A55C1067FF98FF9DFA99FA9DDC6FFA00" box="[1281,1391,1404,1430]" pageId="5" pageNumber="446" value="2023-01-14">
<collectingDate id="B518E98FFF98FF9DFA99FA9DDC6FFA00" box="[1281,1391,1404,1430]" pageId="5" pageNumber="446" value="2023-01-14">14.I.2023</collectingDate>
</date>
,
<collectorName id="7C175371FF98FF9DFAE1FA9DD9D5FA2F" pageId="5" pageNumber="446">AP Pinto</collectorName>
,
<collectionCode id="B7F3AE62FF98FF9DFF47FA7ED806FA2F" box="[223,262,1439,1465]" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="5" pageNumber="446" type="University or college">AC</collectionCode>
</materialsCitation>
<materialsCitation id="618A3CFAFF98FF9DFE95FA41DB65FA2C" box="[269,613,1439,1466]" collectionCode="LP" collectorName="Ehlert" country="Brazil" pageId="5" pageNumber="446" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
Doma- \ hovski, J
<collectorName id="7C175371FF98FF9DFE44FA7EDB20FA2F" box="[476,544,1439,1465]" pageId="5" pageNumber="446">Ehlert</collectorName>
&amp;
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<materialsCitation id="618A3CFAFF98FF9DFDF2FA7ED9C7FA4B" collectionCode="DZRJ, MG" collectorName="Aguiar &amp; S. Goncalo Rio Abaixo" country="Brazil" location="Minas Gerais" pageId="5" pageNumber="446" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
<collectorName id="7C175371FF98FF9DFDF2FA7EDBB8FA2F" box="[618,696,1439,1465]" pageId="5" pageNumber="446">Aguiar</collectorName>
leg.” (
<collectionCode id="B7F3AE62FF98FF9DFC9CFA7EDA4DFA2F" box="[772,845,1439,1465]" pageId="5" pageNumber="446">DZRJ</collectionCode>
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:
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♂,
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<emphasis id="E396EAB5FF98FF9DFBDCFA7EDD52FA2F" bold="true" box="[1092,1106,1439,1465]" pageId="5" pageNumber="446">2</emphasis>
♀
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, “
<collectorName id="7C175371FF98FF9DFB1AFA7EDC89FA2F" box="[1154,1417,1439,1465]" pageId="5" pageNumber="446">S. Gonçalo Rio Abaixo</collectorName>
, \
<collectionCode id="B7F3AE62FF98FF9DFF0FFA22D9C7FA4B" box="[151,199,1475,1501]" country="China" name="Museum of Zoology" pageId="5" pageNumber="446">MG</collectionCode>
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,
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<collectingCounty id="383C4E2BFF98FF9DFF48FA22D816FA4B" box="[208,278,1475,1501]" pageId="5" pageNumber="446">Brasil</collectingCounty>
, (
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)
<date id="A55C1067FF98FF9DFDA3FA22DBB0FA4B" box="[571,688,1475,1501]" pageId="5" pageNumber="446" value="2013-04-12">
<collectingDate id="B518E98FFF98FF9DFDA3FA22DBB0FA4B" box="[571,688,1475,1501]" pageId="5" pageNumber="446" value="2013-04-12">12.iv.2013</collectingDate>
</date>
\
<location id="D43D607CFF98FF9DFD5AFA22DBFFFA4B" LSID="urn:lsid:plazi:treatment:594B87B1FF98FF9EFF0FFE73D849F83F:D43D607CFF98FF9DFD5AFA22DBFFFA4B" box="[706,767,1475,1501]" county="Brasil" latitude="-19.883888" municipality="Est. Amb" name="Light" pageId="5" pageNumber="446">Light</location>
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43º22’ \ 21”
<collectorName id="7C175371FF98FF9DFB95FA25DD89FA4B" box="[1037,1161,1475,1502]" pageId="5" pageNumber="446">W A. Lima</collectorName>
;
<collectorName id="7C175371FF98FF9DFB09FA22DC31FA4B" box="[1169,1329,1475,1502]" pageId="5" pageNumber="446">A. F. Kumagai</collectorName>
&amp;
<collectorName id="7C175371FF98FF9DFAC9FA25DC9CFA4B" box="[1361,1436,1475,1502]" pageId="5" pageNumber="446">P. Dias</collectorName>
legs.” (
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,
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,
<collectionCode id="B7F3AE62FF98FF9DFED4FA06D892F997" box="[332,402,1511,1537]" pageId="5" pageNumber="446">DZRJ</collectionCode>
,
<specimenCount id="C7E4FD2EFF98FF9DFE06FA06D8C0F997" box="[414,448,1511,1537]" count="1" pageId="5" pageNumber="446" type="female">1♀</specimenCount>
<collectionCode id="B7F3AE62FF98FF9DFE50FA06DB16F997" box="[456,534,1511,1537]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
);
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<emphasis id="E396EAB5FF98FF9DFDB2FA06DB38F997" bold="true" box="[554,568,1511,1537]" pageId="5" pageNumber="446">1</emphasis>
♀
</specimenCount>
, same data as preceding, except “
<date id="A55C1067FF98FF9DFC53FA06DD38F997" box="[971,1080,1511,1537]" pageId="5" pageNumber="446" value="2004-01-02">
<collectingDate id="B518E98FFF98FF9DFC53FA06DD38F997" box="[971,1080,1511,1537]" pageId="5" pageNumber="446" value="2004-01-02">02.I.2004</collectingDate>
</date>
\
<collectorName id="7C175371FF98FF9DFBD5FA06DDFBF997" box="[1101,1275,1511,1537]" pageId="5" pageNumber="446">A. F. Kumagai.</collectorName>
col.” (
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);
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<emphasis id="E396EAB5FF98FF9DFF0FF9EBD9A5F9B2" bold="true" box="[151,165,1546,1572]" pageId="5" pageNumber="446">1</emphasis>
♂
</specimenCount>
, “Brasil,
<collectionCode id="B7F3AE62FF98FF9DFEB1F9EAD859F9B3" box="[297,345,1547,1573]" country="China" name="Museum of Zoology" pageId="5" pageNumber="446">MG</collectionCode>
</materialsCitation>
,
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<collectingCounty id="383C4E2BFF98FF9DFEFEF9EAD8A1F9B3" box="[358,417,1547,1573]" pageId="5" pageNumber="446">Serra</collectingCounty>
do \
<collectingMunicipality id="3139ACDDFF98FF9DFE47F9EADB30F9B3" box="[479,560,1547,1573]" pageId="5" pageNumber="446">Caraça</collectingMunicipality>
, arm. lum.
<location id="D43D607CFF98FF9DFD2BF9EADBDFF9B3" LSID="urn:lsid:plazi:treatment:594B87B1FF98FF9EFF0FFE73D849F83F:D43D607CFF98FF9DFD2BF9EADBDFF9B3" box="[691,735,1547,1573]" country="Brazil" county="Serra" municipality="Caraca" name="Luz" pageId="5" pageNumber="446" stateProvince="Parana">Luz</location>
\
<collectorName id="7C175371FF98FF9DFD60F9EBDA75F9B2" box="[760,885,1546,1572]" pageId="5" pageNumber="446">de Queiróz</collectorName>
<date id="A55C1067FF98FF9DFCE6F9EBDAFDF9B2" box="[894,1021,1546,1572]" pageId="5" pageNumber="446" value="2014-12-11">
<collectingDate id="B518E98FFF98FF9DFCE6F9EBDAFDF9B2" box="[894,1021,1546,1572]" pageId="5" pageNumber="446" value="2014-12-11">11.xii.2014</collectingDate>
</date>
\
<collectorName id="7C175371FF98FF9DFB8DF9EADD8DF9B2" box="[1045,1165,1546,1573]" pageId="5" pageNumber="446">A. Capelli</collectorName>
;
<collectorName id="7C175371FF98FF9DFB02F9EADC06F9B3" box="[1178,1286,1547,1573]" pageId="5" pageNumber="446">D. Souza</collectorName>
;
<collectorName id="7C175371FF98FF9DFA8BF9EADC98F9B2" box="[1299,1432,1546,1573]" pageId="5" pageNumber="446">M. Cupello</collectorName>
;
<collectorName id="7C175371FF98FF9DFF0FF9CED812F9DF" box="[151,274,1583,1609]" pageId="5" pageNumber="446">J.P. Botero</collectorName>
leg.” (
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);
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<collectingRegion id="1326F845FF98FF9DFE5AF9CCDB1DF9DE" box="[450,541,1581,1608]" country="Brazil" name="Parana" pageId="5" pageNumber="446">Paraná</collectingRegion>
:
<specimenCount id="C7E4FD2EFF98FF9DFDB4F9CFDB3AF9DE" box="[556,570,1582,1608]" count="1" pageId="5" pageNumber="446" type="male">1</specimenCount>
</emphasis>
♂, “Brasil
</materialsCitation>
,
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<collectingRegion id="1326F845FF98FF9DFD27F9CFDA0EF9DE" box="[703,782,1582,1608]" country="Brazil" name="Parana" pageId="5" pageNumber="446">Paraná</collectingRegion>
,
<collectingMunicipality id="3139ACDDFF98FF9DFC84F9CFDA80F9DE" box="[796,896,1582,1608]" pageId="5" pageNumber="446">São José</collectingMunicipality>
\ dos
<location id="D43D607CFF98FF9DFC53F9CFDD20F9DE" LSID="urn:lsid:plazi:treatment:594B87B1FF98FF9EFF0FFE73D849F83F:D43D607CFF98FF9DFC53F9CFDD20F9DE" box="[971,1056,1582,1608]" country="Brazil" latitude="-25.603514" longLatPrecision="1" longitude="-49.19266" municipality="Sao Jose" name="Pinhais" pageId="5" pageNumber="446" stateProvince="Parana">Pinhais</location>
,
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\
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<quantity id="161A9B42FF98FF9DFF0FF9B3D9D7F9FA" box="[151,215,1618,1644]" metricMagnitude="2" metricUnit="m" metricValue="8.83" pageId="5" pageNumber="446" unit="m" value="883.0">
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\
<date id="A55C1067FF98FF9DFF6AF9B3D88DF9FB" box="[242,397,1618,1645]" pageId="5" pageNumber="446" value="2015-10-10" valueMax="2015-10-17" valueMin="2015-10-10">
<collectingDate id="B518E98FFF98FF9DFF6AF9B3D88DF9FB" box="[242,397,1618,1645]" pageId="5" pageNumber="446" value="2015-10-10" valueMax="2015-10-17" valueMin="2015-10-10">10-17.X.2015</collectingDate>
</date>
Malaise-2 \
<collectorName id="7C175371FF98FF9DFDB8F9B3DBEEF9FA" box="[544,750,1618,1644]" pageId="5" pageNumber="446">A.C. Domahovski</collectorName>
leg.” (
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);
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<emphasis id="E396EAB5FF98FF9DFC3CF9B3DAB2F9FA" bold="true" box="[932,946,1618,1644]" pageId="5" pageNumber="446">1</emphasis>
♂
</specimenCount>
, “
<collectorName id="7C175371FF98FF9DFC7AF9B3DD2AF9FA" box="[994,1066,1618,1644]" pageId="5" pageNumber="446">Brasil</collectorName>
</materialsCitation>
,
<materialsCitation id="618A3CFAFF98FF9DFBAFF9B3DC66F8D4" collectingDate="2015-09-05" collectingDateMax="2016-07-16" collectingDateMin="2015-09-05" collectionCode="DZUP" collectorName="dos Pinhais &amp; A. C. Domahovski" country="Brazil" elevation="880" latitude="-25.60039" location="Sao Jose" longLatPrecision="1" longitude="-49.190186" municipality="Sao Jose" pageId="5" pageNumber="446" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Parana" typeStatus="paratype">
<collectingRegion id="1326F845FF98FF9DFBAFF9B3DD86F9FA" box="[1079,1158,1618,1644]" country="Brazil" name="Parana" pageId="5" pageNumber="446">Paraná</collectingRegion>
,
<location id="D43D607CFF98FF9DFB0CF9B3DDF7F9FA" LSID="urn:lsid:plazi:treatment:594B87B1FF98FF9EFF0FFE73D849F83F:D43D607CFF98FF9DFB0CF9B3DDF7F9FA" box="[1172,1271,1618,1644]" latitude="-25.60039" longLatPrecision="1" longitude="-49.190186" name="Sao Jose" pageId="5" pageNumber="446" stateProvince="Parana">São José</location>
\
<collectorName id="7C175371FF98FF9DFA8AF9B3DC98F9FA" box="[1298,1432,1618,1644]" pageId="5" pageNumber="446">dos Pinhais</collectorName>
,
<geoCoordinate id="B4D65060FF98FF9DFF0FF994D83CF906" box="[151,316,1653,1680]" degrees="25" direction="south" minutes="36" orientation="latitude" pageId="5" pageNumber="446" precision="1" seconds="01.40" value="-25.60039">25º36’01.40”S</geoCoordinate>
\
<geoCoordinate id="B4D65060FF98FF9DFEC9F997D8FFF906" box="[337,511,1654,1680]" degrees="49" direction="west" minutes="11" orientation="longitude" pageId="5" pageNumber="446" precision="1" seconds="24.66" value="-49.190186">49º11’24.66”W</geoCoordinate>
<quantity id="161A9B42FF98FF9DFD9DF997DB45F906" box="[517,581,1654,1680]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="5" pageNumber="446" unit="m" value="880.0">
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</quantity>
\
<date id="A55C1067FF98FF9DFDC1F994DBFEF906" box="[601,766,1653,1680]" pageId="5" pageNumber="446" value="2015-09-05" valueMax="2015-09-12" valueMin="2015-09-05">
<collectingDate id="B518E98FFF98FF9DFDC1F994DBFEF906" box="[601,766,1653,1680]" pageId="5" pageNumber="446" value="2015-09-05" valueMax="2015-09-12" valueMin="2015-09-05">05-12.IX.2015</collectingDate>
</date>
Malaise-3 \
<collectorName id="7C175371FF98FF9DFC1FF997DD51F906" box="[903,1105,1654,1680]" pageId="5" pageNumber="446">A.C. Domahovski</collectorName>
leg.” (
<collectionCode id="B7F3AE62FF98FF9DFB04F997DDEBF906" box="[1180,1259,1654,1680]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
);
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<emphasis id="E396EAB5FF98FF9DFB65F994DC0BF919" bold="true" box="[1277,1291,1653,1679]" pageId="5" pageNumber="446">1</emphasis>
♀
</specimenCount>
, same data as preceding, except
<date id="A55C1067FF98FF9DFE19F978DB26F922" box="[385,550,1689,1716]" pageId="5" pageNumber="446" value="2015-09-19" valueMax="2015-09-26" valueMin="2015-09-19">
<collectingDate id="B518E98FFF98FF9DFE19F978DB26F922" box="[385,550,1689,1716]" pageId="5" pageNumber="446" value="2015-09-19" valueMax="2015-09-26" valueMin="2015-09-19">19-26.IX.2015</collectingDate>
</date>
(
<collectionCode id="B7F3AE62FF98FF9DFDACF97BDB83F922" box="[564,643,1690,1716]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
);
<specimenCount id="C7E4FD2EFF98FF9DFD0EF978DBB9F922" box="[662,697,1689,1716]" count="1" pageId="5" pageNumber="446" type="female">
<emphasis id="E396EAB5FF98FF9DFD0EF978DBA4F925" bold="true" box="[662,676,1689,1715]" pageId="5" pageNumber="446">1</emphasis>
♀
</specimenCount>
, same data as preceding, except
<date id="A55C1067FF98FF9DFBBEF978DDC1F922" box="[1062,1217,1689,1716]" pageId="5" pageNumber="446" value="2015-10-03" valueMax="2015-10-10" valueMin="2015-10-03">
<collectingDate id="B518E98FFF98FF9DFBBEF978DDC1F922" box="[1062,1217,1689,1716]" pageId="5" pageNumber="446" value="2015-10-03" valueMax="2015-10-10" valueMin="2015-10-03">03-10.X.2015</collectingDate>
</date>
(
<collectionCode id="B7F3AE62FF98FF9DFB48F97BDC1FF922" box="[1232,1311,1690,1716]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
);
<specimenCount id="C7E4FD2EFF98FF9DFAAAF978DC55F922" box="[1330,1365,1689,1716]" count="1" pageId="5" pageNumber="446" type="female">
<emphasis id="E396EAB5FF98FF9DFAAAF978DC40F925" bold="true" box="[1330,1344,1689,1715]" pageId="5" pageNumber="446">1</emphasis>
♀
</specimenCount>
, same data as preceding, except
<date id="A55C1067FF98FF9DFE5DF95CDB61F94E" box="[453,609,1725,1752]" pageId="5" pageNumber="446" value="2015-10-17" valueMax="2015-10-24" valueMin="2015-10-17">
<collectingDate id="B518E98FFF98FF9DFE5DF95CDB61F94E" box="[453,609,1725,1752]" pageId="5" pageNumber="446" value="2015-10-17" valueMax="2015-10-24" valueMin="2015-10-17">17-24.X.2015</collectingDate>
</date>
(
<collectionCode id="B7F3AE62FF98FF9DFDEBF95CDBC2F941" box="[627,706,1725,1751]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
);
<specimenCount id="C7E4FD2EFF98FF9DFD41F95CDBFCF94E" box="[729,764,1725,1752]" count="1" pageId="5" pageNumber="446" type="female">
<emphasis id="E396EAB5FF98FF9DFD41F95CDBE7F941" bold="true" box="[729,743,1725,1751]" pageId="5" pageNumber="446">1</emphasis>
♀
</specimenCount>
, same data as preceding, except
<date id="A55C1067FF98FF9DFB18F95CDC2EF941" box="[1152,1326,1725,1751]" pageId="5" pageNumber="446" value="2016-07-09" valueMax="2016-07-16" valueMin="2016-07-09">
<collectingDate id="B518E98FFF98FF9DFB18F95CDC2EF941" box="[1152,1326,1725,1751]" pageId="5" pageNumber="446" value="2016-07-09" valueMax="2016-07-16" valueMin="2016-07-09">09-16.VII.2016</collectingDate>
</date>
(
<collectionCode id="B7F3AE62FF98FF9DFAD9F95CDC90F941" box="[1345,1424,1725,1751]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
);
<specimenCount id="C7E4FD2EFF98FF9DFF0FF900D9BAF96D" box="[151,186,1761,1787]" count="1" pageId="5" pageNumber="446" type="female">
<emphasis id="E396EAB5FF98FF9DFF0FF900D9A5F96D" bold="true" box="[151,165,1761,1787]" pageId="5" pageNumber="446">1</emphasis>
♀
</specimenCount>
, “Brasil,
<collectingRegion id="1326F845FF98FF9DFEB5F900D87CF96D" box="[301,380,1761,1787]" country="Brazil" name="Parana" pageId="5" pageNumber="446">Paraná</collectingRegion>
, São José \ dos Pinhais,
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<geoCoordinate id="B4D65060FF98FF9DFCCCF900DD02F96D" box="[852,1026,1761,1787]" degrees="49" direction="west" minutes="11" orientation="longitude" pageId="5" pageNumber="446" precision="1" seconds="22.80" value="-49.189667">49º11’22.80”W</geoCoordinate>
<quantity id="161A9B42FF98FF9DFB95F900DD4CF96D" box="[1037,1100,1761,1787]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="5" pageNumber="446" unit="m" value="880.0">
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</quantity>
\
<date id="A55C1067FF98FF9DFBF2F900DC0FF96A" box="[1130,1295,1761,1788]" pageId="5" pageNumber="446" value="2015-09-19" valueMax="2015-09-26" valueMin="2015-09-19">
<collectingDate id="B518E98FFF98FF9DFBF2F900DC0FF96A" box="[1130,1295,1761,1788]" pageId="5" pageNumber="446" value="2015-09-19" valueMax="2015-09-26" valueMin="2015-09-19">19-26.IX.2015</collectingDate>
</date>
Malaise-4 \
<collectorName id="7C175371FF98FF9DFF0FF8E4D865F889" box="[151,357,1797,1823]" pageId="5" pageNumber="446">A.C. Domahovski</collectorName>
leg.” (
<collectionCode id="B7F3AE62FF98FF9DFE2EF8E4DB05F889" box="[438,517,1797,1823]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
);
<specimenCount id="C7E4FD2EFF98FF9DFD84F8E5DB41F889" box="[540,577,1796,1823]" count="1" pageId="5" pageNumber="446" type="male">
<emphasis id="E396EAB5FF98FF9DFD84F8E5DB2AF888" bold="true" box="[540,554,1796,1822]" pageId="5" pageNumber="446">1</emphasis>
♂
</specimenCount>
, same data as preceding, except
<date id="A55C1067FF98FF9DFC27F8E5DD7FF889" box="[959,1151,1796,1823]" pageId="5" pageNumber="446" value="2015-10-31" valueMax="2015-11-07" valueMin="2015-10-31">
<collectingDate id="B518E98FFF98FF9DFC27F8E5DD7FF889" box="[959,1151,1796,1823]" pageId="5" pageNumber="446" value="2015-10-31" valueMax="2015-11-07" valueMin="2015-10-31">31.X-07.XI.2015</collectingDate>
</date>
(
<collectionCode id="B7F3AE62FF98FF9DFB09F8E4DDE0F889" box="[1169,1248,1797,1823]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
);
<specimenCount id="C7E4FD2EFF98FF9DFB6EF8E5DC1BF889" box="[1270,1307,1796,1823]" count="1" pageId="5" pageNumber="446" type="male">
<emphasis id="E396EAB5FF98FF9DFB6EF8E5DC04F888" bold="true" box="[1270,1284,1796,1822]" pageId="5" pageNumber="446">1</emphasis>
♂
</specimenCount>
, same data as preceding, except
<date id="A55C1067FF98FF9DFE1DF8C9DB58F8D5" box="[389,600,1832,1859]" pageId="5" pageNumber="446" value="2015-11-28" valueMax="2015-12-05" valueMin="2015-11-28">
<collectingDate id="B518E98FFF98FF9DFE1DF8C9DB58F8D5" box="[389,600,1832,1859]" pageId="5" pageNumber="446" value="2015-11-28" valueMax="2015-12-05" valueMin="2015-11-28">28.XI-05.XII.2015</collectingDate>
</date>
(
<collectionCode id="B7F3AE62FF98FF9DFDF0F8C9DBB7F8D4" box="[616,695,1832,1858]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
);
<specimenCount id="C7E4FD2EFF98FF9DFD54F8C9DBEFF8D5" box="[716,751,1832,1859]" count="1" pageId="5" pageNumber="446" type="female">
<emphasis id="E396EAB5FF98FF9DFD54F8C9DBDAF8D4" bold="true" box="[716,730,1832,1858]" pageId="5" pageNumber="446">1</emphasis>
♀
</specimenCount>
, same data as preceding, except
<date id="A55C1067FF98FF9DFBFCF8C9DDFEF8D4" box="[1124,1278,1832,1858]" pageId="5" pageNumber="446" value="2016-02-20" valueMax="2016-02-27" valueMin="2016-02-20">
<collectingDate id="B518E98FFF98FF9DFBFCF8C9DDFEF8D4" box="[1124,1278,1832,1858]" pageId="5" pageNumber="446" value="2016-02-20" valueMax="2016-02-27" valueMin="2016-02-20">20-27.II.2016</collectingDate>
</date>
(
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♀, “Brasil, RS,
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</paragraph>
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<caption id="859D662FFF9BFF9EFF0FF990DADAF963" ID-DOI="http://doi.org/10.5281/zenodo.13347007" ID-Zenodo-Dep="13347007" httpUri="https://zenodo.org/record/13347007/files/figure.png" pageId="6" pageNumber="447" startId="6.[151,250,1649,1673]" targetBox="[154,1435,186,1620]" targetPageId="6" targetType="figure">
<paragraph id="D15D36A7FF9BFF9EFF0FF990DADAF963" blockId="6.[151,1437,1649,1781]" pageId="6" pageNumber="447">
<emphasis id="E396EAB5FF9BFF9EFF0FF990D812F91F" bold="true" box="[151,274,1649,1673]" pageId="6" pageNumber="447">FIGURE 3.</emphasis>
<taxonomicName id="16E24D24FF9BFF9EFE80F990D8FFF91F" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[280,511,1649,1673]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="447" phylum="Arthropoda" rank="species" species="tetrarama" status="sp. nov.">
<emphasis id="E396EAB5FF9BFF9EFE80F990D8FFF91F" box="[280,511,1649,1673]" italics="true" pageId="6" pageNumber="447">Gracilipona tetrarama</emphasis>
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<taxonomicNameLabel id="F8A557CEFF9BFF9EFD9DF990DB54F91F" box="[517,596,1649,1673]" pageId="6" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D, forewing. E–M, male terminalia: E, sternite VIII, ventral view. F, valve, ventral view. G, pygofer, valve, and subgenital plate, lateral view. H, subgenital plate, ventral view. I, connective, dorsal view. J, style, dorsal view. K, style, lateral view. L, aedeagus, lateral view. M, aedeagus, posterior view. Scales in mm.
</paragraph>
</caption>
<subSubSection id="99F8652CFF9BFF9EFF5FF8A6D849F83F" pageId="6" pageNumber="447" type="discussion">
<paragraph id="D15D36A7FF9BFF9EFF5FF8A6D849F83F" blockId="6.[151,1436,1863,1961]" pageId="6" pageNumber="447">
<emphasis id="E396EAB5FF9BFF9EFF5FF8A6D83BF8F7" bold="true" box="[199,315,1863,1889]" pageId="6" pageNumber="447">Remarks.</emphasis>
<taxonomicName id="16E24D24FF9BFF9EFED8F8A9D8D1F8F7" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[320,465,1864,1890]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="447" phylum="Arthropoda" rank="species" species="tetrarama" status="sp. nov.">
<emphasis id="E396EAB5FF9BFF9EFED8F8A9D8D1F8F7" box="[320,465,1864,1890]" italics="true" pageId="6" pageNumber="447">G. tetrarama</emphasis>
</taxonomicName>
<emphasis id="E396EAB5FF9BFF9EFE4EF8A6DB2EF8F7" bold="true" box="[470,558,1863,1889]" pageId="6" pageNumber="447">
<taxonomicNameLabel id="F8A557CEFF9BFF9EFE4EF8A6DB2EF8F7" box="[470,558,1863,1889]" pageId="6" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is easily distinguished from
<taxonomicName id="16E24D24FF9BFF9EFCF0F8A9DAD9F8F7" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[872,985,1863,1890]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="447" phylum="Arthropoda" rank="species" species="birama" status="sp. nov.">
<emphasis id="E396EAB5FF9BFF9EFCF0F8A9DAD9F8F7" box="[872,985,1863,1890]" italics="true" pageId="6" pageNumber="447">G. birama</emphasis>
</taxonomicName>
<emphasis id="E396EAB5FF9BFF9EFC46F8A6DD36F8F7" bold="true" box="[990,1078,1863,1889]" pageId="6" pageNumber="447">
<taxonomicNameLabel id="F8A557CEFF9BFF9EFC46F8A6DD36F8F7" box="[990,1078,1863,1889]" pageId="6" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having the aedeagus with two pairs of processes, being the ventral pair 2x longer than the shaft length and female sternite VII with posterolateral angles rounded.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="0961AE6CA3DF5ACC2D3FC64F07B3363F">A new genus of Gyponini (Hemiptera, Cicadellidae, Iassinae) with description of two new species from Brazil and key to genera of the Hecalapona complex</mods:title>
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<emphasis id="E396EAB5FF9FFF9AFF0FF8C6D8EBF8D4" bold="true" box="[151,491,1831,1858]" pageId="2" pageNumber="443">
<taxonomicName id="16E24D24FF9FFF9AFF0FF8C6D886F8D4" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[151,390,1831,1858]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="birama" status="sp. nov.">
<emphasis id="E396EAB5FF9FFF9AFF0FF8C6D886F8D4" bold="true" box="[151,390,1831,1858]" italics="true" pageId="2" pageNumber="443">Gracilipona birama</emphasis>
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<taxonomicNameLabel id="F8A557CEFF9FFF9AFE15F8C9D8EBF8D4" box="[397,491,1832,1858]" pageId="2" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="99F8652CFF9FFF9AFF0FF8ADD85CF8F1" box="[151,348,1868,1895]" pageId="2" pageNumber="443" type="description">
<paragraph id="D15D36A7FF9FFF9AFF0FF8ADD85CF8F1" blockId="2.[151,491,1831,1895]" box="[151,348,1868,1895]" pageId="2" pageNumber="443">
(
<figureCitation id="49D92A22FF9FFF9AFF07F8ADD9E6F8F0" box="[159,230,1868,1894]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="2" pageNumber="443">Figs 1</figureCitation>
,
<figureCitation id="49D92A22FF9FFF9AFF6CF8ADD804F8F0" box="[244,260,1868,1894]" captionStart="FIGURE 2" captionStartId="4.[151,250,1763,1787]" captionTargetBox="[154,1431,381,1735]" captionTargetId="figure-116@4.[151,1435,381,1740]" captionTargetPageId="4" captionText="FIGURE 2. Gracilipona birama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347005" httpUri="https://zenodo.org/record/13347005/files/figure.png" pageId="2" pageNumber="443">2</figureCitation>
,
<figureCitation id="49D92A22FF9FFF9AFE88F8ADD854F8F1" box="[272,340,1868,1895]" captionStart="FIGURE 5" captionStartId="8.[151,250,1189,1213]" captionTargetBox="[151,1436,491,1165]" captionTargetId="figure-654@8.[151,1436,491,1165]" captionTargetPageId="8" captionText="FIGURE 5. Habitus, dorsal view (left) and lateral view (right). A, B, Gracilipona birama sp. nov., holotype male. C, D, Gracilipona tetrarama sp. nov., holotype male. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347011" httpUri="https://zenodo.org/record/13347011/files/figure.png" pageId="2" pageNumber="443">5A–B</figureCitation>
)
</paragraph>
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<subSubSection id="99F8652CFF9FFF9AFF0FF875DC9DF860" pageId="2" pageNumber="443" type="diagnosis">
<paragraph id="D15D36A7FF9FFF9AFF0FF875DC9DF860" blockId="2.[151,1437,1940,2039]" pageId="2" pageNumber="443">
<emphasis id="E396EAB5FF9FFF9AFF0FF875D812F838" bold="true" box="[151,274,1940,1966]" pageId="2" pageNumber="443">Diagnosis.</emphasis>
Style (
<figureCitation id="49D92A22FF9FFF9AFEF8F875D8ACF838" box="[352,428,1940,1966]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="2" pageNumber="443">Fig. 1J</figureCitation>
) blade with ventral margin expanded at apical half; dorsal margin somewhat straight; apex directed dorsally; aedeagus (
<figureCitation id="49D92A22FF9FFF9AFE47F859DB5DF844" box="[479,605,1976,2002]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="2" pageNumber="443">Figs 1K–L</figureCitation>
) shaft long with pair of processes, each elongate and extending slightly beyond apex of shaft; female sternite VII (
<figureCitation id="49D92A22FF9FFF9AFDEAF83DDBCAF860" box="[626,714,2012,2038]" captionStart="FIGURE 2" captionStartId="4.[151,250,1763,1787]" captionTargetBox="[154,1431,381,1735]" captionTargetId="figure-116@4.[151,1435,381,1740]" captionTargetPageId="4" captionText="FIGURE 2. Gracilipona birama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347005" httpUri="https://zenodo.org/record/13347005/files/figure.png" pageId="2" pageNumber="443">Fig. 2A</figureCitation>
) with posterolateral angles subacute, weakly produced posterad.
</paragraph>
</subSubSection>
<caption id="859D662FFF9EFF9BFF0FF9EADA35F918" ID-DOI="http://doi.org/10.5281/zenodo.13347003" ID-Zenodo-Dep="13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="3" pageNumber="444" startId="3.[151,250,1547,1571]" targetBox="[151,1436,204,1524]" targetPageId="3" targetType="figure">
<paragraph id="D15D36A7FF9EFF9BFF0FF9EADA35F918" blockId="3.[151,1437,1547,1679]" pageId="3" pageNumber="444">
<emphasis id="E396EAB5FF9EFF9BFF0FF9EAD814F9B5" bold="true" box="[151,276,1547,1571]" pageId="3" pageNumber="444">FIGURE 1.</emphasis>
<taxonomicName id="16E24D24FF9EFF9BFE83F9EAD8E7F9B5" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[283,487,1547,1571]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="birama" status="sp. nov.">
<emphasis id="E396EAB5FF9EFF9BFE83F9EAD8E7F9B5" box="[283,487,1547,1571]" italics="true" pageId="3" pageNumber="444">Gracilipona birama</emphasis>
</taxonomicName>
<emphasis id="E396EAB5FF9EFF9BFE76F9EADB3FF9B5" bold="true" box="[494,575,1547,1571]" pageId="3" pageNumber="444">
<taxonomicNameLabel id="F8A557CEFF9EFF9BFE76F9EADB3FF9B5" box="[494,575,1547,1571]" pageId="3" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm.
</paragraph>
</caption>
<subSubSection id="99F8652CFF9EFF9CFF5FF95CDDBCFEF3" lastPageId="4" lastPageNumber="445" pageId="3" pageNumber="444" type="description">
<paragraph id="D15D36A7FF9EFF9BFF5FF95CDDC8F941" blockId="3.[151,1436,1725,2040]" box="[199,1224,1725,1752]" pageId="3" pageNumber="444">
<emphasis id="E396EAB5FF9EFF9BFF5FF95CD8C5F941" bold="true" box="[199,453,1725,1751]" pageId="3" pageNumber="444">Measurements (mm).</emphasis>
<typeStatus id="0E598805FF9EFF9BFE55F95CDB35F941" box="[461,565,1725,1751]" pageId="3" pageNumber="444" type="holotype">Holotype</typeStatus>
male: total length 7.1.
<typeStatus id="0E598805FF9EFF9BFCA0F95FDAA9F94E" box="[824,937,1726,1752]" pageId="3" pageNumber="444" type="paratype">Paratypes</typeStatus>
: females (n = 2), 7.9–8.1.
</paragraph>
<paragraph id="D15D36A7FF9EFF9CFF5FF900DA74FF43" blockId="3.[151,1436,1725,2040]" lastBlockId="4.[151,1436,151,357]" lastPageId="4" lastPageNumber="445" pageId="3" pageNumber="444">
<emphasis id="E396EAB5FF9EFF9BFF5FF900D88FF96D" bold="true" box="[199,399,1761,1787]" pageId="3" pageNumber="444">Male terminalia.</emphasis>
Sternite VIII (
<figureCitation id="49D92A22FF9EFF9BFDACF900DB8BF96D" box="[564,651,1761,1787]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="3" pageNumber="444">Fig. 1D</figureCitation>
) in ventral view, as wide as long; lateral margins straight and slightly tapering to apex; posterior margin rounded. Valve (
<figureCitation id="49D92A22FF9EFF9BFD44F8E4DA31F889" box="[732,817,1797,1823]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="3" pageNumber="444">Fig. 1E</figureCitation>
) in ventral view, 2x wider than long; posterior margin acute. Pygofer (
<figureCitation id="49D92A22FF9EFF9BFEDBF8C8D893F8D5" box="[323,403,1833,1859]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="3" pageNumber="444">Fig. 1F</figureCitation>
) in lateral view, 2x longer than high, ventral margin broadly rounded; apical portion triangular, tapering to acute apex. Subgenital plate (
<figureCitation id="49D92A22FF9EFF9BFDC5F8ACDBB4F8F1" box="[605,692,1869,1895]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="3" pageNumber="444">Fig. 1G</figureCitation>
), in ventral view, approximately 5.8x longer than wide; wider near apex than near base; lateral margins straight; apex rounded. Connective (
<figureCitation id="49D92A22FF9EFF9BFC7FF890DD40F81D" box="[999,1088,1905,1931]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="3" pageNumber="444">Fig. 1H</figureCitation>
) in dorsal view, with arms as long as stem; stem expanded apically; dorsal keel developed. Style (
<figureCitation id="49D92A22FF9EFF9BFC0BF874DADFF839" box="[915,991,1941,1967]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="3" pageNumber="444">Fig. 1I</figureCitation>
) in dorsal view, with outer lobe weakly developed, rounded; in lateral view (
<figureCitation id="49D92A22FF9EFF9BFDDAF858DB93F845" box="[578,659,1977,2003]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="3" pageNumber="444">Fig. 1J</figureCitation>
), blade with ventral margin expanded at apical half; dorsal margin somewhat straight; apex thin, acute, directed dorsally. Aedeagus (
<figureCitation id="49D92A22FF9EFF9BFC13F83CDD0AF861" box="[907,1034,2013,2039]" captionStart="FIGURE 1" captionStartId="3.[151,250,1547,1571]" captionTargetBox="[151,1436,204,1524]" captionTargetId="figure-20@3.[151,1436,201,1524]" captionTargetPageId="3" captionText="FIGURE 1. Gracilipona birama sp. nov., holotype male. A, head and thorax, dorsal view. B, head, ventral view. C, head and thorax, lateral view. D–L, male terminalia: D, sternite VIII, ventral view. E, valve, ventral view. F, pygofer, valve, and subgenital plate, lateral view. G, subgenital plate, ventral view. H, connective, dorsal view. I, style, dorsal view. J, style, lateral view. K, aedeagus, lateral view. L, aedeagus, posterior view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347003" httpUri="https://zenodo.org/record/13347003/files/figure.png" pageId="3" pageNumber="444">Figs 1K–L</figureCitation>
) shaft long, with pair of processes arising lateroventrally and extending slightly beyond apex of shaft, curved anterad, weakly sclerotized; apex of shaft without processes. Other characteristics as in generic description.
</paragraph>
<paragraph id="D15D36A7FF99FF9CFF5FFF3EDDBCFEF3" blockId="4.[151,1436,151,357]" pageId="4" pageNumber="445">
<emphasis id="E396EAB5FF99FF9CFF5FFF3ED8A7FF6E" bold="true" box="[199,423,222,249]" pageId="4" pageNumber="445">Female terminalia.</emphasis>
Sternite VII (
<figureCitation id="49D92A22FF99FF9CFDA5FF3EDB96FF6F" box="[573,662,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1763,1787]" captionTargetBox="[154,1431,381,1735]" captionTargetId="figure-116@4.[151,1435,381,1740]" captionTargetPageId="4" captionText="FIGURE 2. Gracilipona birama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347005" httpUri="https://zenodo.org/record/13347005/files/figure.png" pageId="4" pageNumber="445">Fig. 2A</figureCitation>
), in ventral view, 1.5x wider than long; posterolateral angles subacute, weakly produced posterad; posterior margin straight. First valvifer (
<figureCitation id="49D92A22FF99FF9CFC04FEE2DAF4FE8B" box="[924,1012,259,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1763,1787]" captionTargetBox="[154,1431,381,1735]" captionTargetId="figure-116@4.[151,1435,381,1740]" captionTargetPageId="4" captionText="FIGURE 2. Gracilipona birama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347005" httpUri="https://zenodo.org/record/13347005/files/figure.png" pageId="4" pageNumber="445">Fig. 2C</figureCitation>
) 1.5x higher than wide. First valvula (
<figureCitation id="49D92A22FF99FF9CFF07FEC6D9F7FED7" box="[159,247,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1763,1787]" captionTargetBox="[154,1431,381,1735]" captionTargetId="figure-116@4.[151,1435,381,1740]" captionTargetPageId="4" captionText="FIGURE 2. Gracilipona birama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347005" httpUri="https://zenodo.org/record/13347005/files/figure.png" pageId="4" pageNumber="445">Fig. 2C</figureCitation>
) 4.2x longer than high. Second valvula (
<figureCitation id="49D92A22FF99FF9CFD48FEC6DA25FED7" box="[720,805,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1763,1787]" captionTargetBox="[154,1431,381,1735]" captionTargetId="figure-116@4.[151,1435,381,1740]" captionTargetPageId="4" captionText="FIGURE 2. Gracilipona birama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347005" httpUri="https://zenodo.org/record/13347005/files/figure.png" pageId="4" pageNumber="445">Fig. 2F</figureCitation>
) 3.4x longer than high; apical portion (
<figureCitation id="49D92A22FF99FF9CFB74FEC6DC46FED7" box="[1260,1350,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1763,1787]" captionTargetBox="[154,1431,381,1735]" captionTargetId="figure-116@4.[151,1435,381,1740]" captionTargetPageId="4" captionText="FIGURE 2. Gracilipona birama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347005" httpUri="https://zenodo.org/record/13347005/files/figure.png" pageId="4" pageNumber="445">Fig. 2G</figureCitation>
) dorsal margin with approximately eight rounded denticles. Gonoplac (
<figureCitation id="49D92A22FF99FF9CFCC6FEAADAB5FEF3" box="[862,949,331,357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1763,1787]" captionTargetBox="[154,1431,381,1735]" captionTargetId="figure-116@4.[151,1435,381,1740]" captionTargetPageId="4" captionText="FIGURE 2. Gracilipona birama sp. nov., paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.13347005" httpUri="https://zenodo.org/record/13347005/files/figure.png" pageId="4" pageNumber="445">Fig. 2H</figureCitation>
) 2.8x longer than high.
</paragraph>
</subSubSection>
<caption id="859D662FFF99FF9CFF0FF902DDBCF8F1" ID-DOI="http://doi.org/10.5281/zenodo.13347005" ID-Zenodo-Dep="13347005" httpUri="https://zenodo.org/record/13347005/files/figure.png" pageId="4" pageNumber="445" startId="4.[151,250,1763,1787]" targetBox="[154,1431,381,1735]" targetPageId="4" targetType="figure">
<paragraph id="D15D36A7FF99FF9CFF0FF902DDBCF8F1" blockId="4.[151,1436,1763,1895]" pageId="4" pageNumber="445">
<emphasis id="E396EAB5FF99FF9CFF0FF902D815F96D" bold="true" box="[151,277,1763,1787]" pageId="4" pageNumber="445">FIGURE 2.</emphasis>
<taxonomicName id="16E24D24FF99FF9CFE85F902D8EAF96D" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[285,490,1763,1787]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="birama" status="sp. nov.">
<emphasis id="E396EAB5FF99FF9CFE85F902D8EAF96D" box="[285,490,1763,1787]" italics="true" pageId="4" pageNumber="445">Gracilipona birama</emphasis>
</taxonomicName>
<emphasis id="E396EAB5FF99FF9CFE6BF902DB45F96D" bold="true" box="[499,581,1763,1787]" pageId="4" pageNumber="445">
<taxonomicNameLabel id="F8A557CEFF99FF9CFE6BF902DB45F96D" box="[499,581,1763,1787]" pageId="4" pageNumber="445" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype female. A, apical portion of abdomen, ventral view. B, apical portion of abdomen, lateral view. C, first valvifer and first valvula, lateral view. D, enlarged view of dorsal sculptured area. E, enlarged view of apical portion of first valvula. F, second valvula, lateral view. G, enlarged view of apical portion of second valvula. G, second valvifer and gonoplac, lateral view. I, enlarged view of apical portion of gonoplac. Scales in mm.
</paragraph>
</caption>
<subSubSection id="99F8652CFF99FF9CFF5FF874D805F845" pageId="4" pageNumber="445" type="etymology">
<paragraph id="D15D36A7FF99FF9CFF5FF874D805F845" blockId="4.[151,1436,1941,2003]" pageId="4" pageNumber="445">
<emphasis id="E396EAB5FF99FF9CFF5FF874D84EF839" bold="true" box="[199,334,1941,1967]" pageId="4" pageNumber="445">Etymology.</emphasis>
The specific epithet
<taxonomicName id="16E24D24FF99FF9CFDDFF874DB98F839" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[583,664,1941,1967]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="birama">
<emphasis id="E396EAB5FF99FF9CFDDFF874DB98F839" box="[583,664,1941,1967]" italics="true" pageId="4" pageNumber="445">birama</emphasis>
</taxonomicName>
(noun in apposition), refers to the single pair of processes on the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="99F8652CFF98FF9DFF5FFF77DA1BFF42" pageId="5" pageNumber="446" type="materials_examined">
<paragraph id="D15D36A7FF98FF9DFF5FFF77DA1BFF42" blockId="5.[151,1436,150,320]" pageId="5" pageNumber="446">
<emphasis id="E396EAB5FF98FF9DFF5FFF77D8B2FF26" bold="true" box="[199,434,150,176]" pageId="5" pageNumber="446">Material examined.</emphasis>
<materialsCitation id="618A3CFAFF98FF9DFE21FF77DB47FF42" collectionCode="DZUP" collectorName="M. Alvarenga" country="Brazil" elevation="980" location="Encruzilhada - Bahia" pageId="5" pageNumber="446" specimenCount="2" specimenCount-female="2" stateProvince="Bahia" typeStatus="holotype">
<emphasis id="E396EAB5FF98FF9DFE21FF77DB20FF26" bold="true" box="[441,544,150,176]" pageId="5" pageNumber="446">
<typeStatus id="0E598805FF98FF9DFE21FF77DB20FF26" box="[441,544,150,176]" pageId="5" pageNumber="446" type="holotype">holotype</typeStatus>
</emphasis>
male:
<emphasis id="E396EAB5FF98FF9DFDF6FF77DBBEFF26" bold="true" box="[622,702,150,176]" pageId="5" pageNumber="446">
<collectingRegion id="1326F845FF98FF9DFDF6FF77DBB8FF26" box="[622,696,150,176]" country="Brazil" name="Bahia" pageId="5" pageNumber="446">Bahia</collectingRegion>
:
</emphasis>
“
<location id="D43D607CFF98FF9DFD57FF77DAC0FF26" LSID="urn:lsid:plazi:treatment:594B87B1FF9FFF9DFF0FF8C6DB64FED6:D43D607CFF98FF9DFD57FF77DAC0FF26" box="[719,960,150,177]" country="Brazil" name="Encruzilhada - Bahia" pageId="5" pageNumber="446" stateProvince="Bahia">Encruzilhada - Bahia</location>
\
<collectingCountry id="A9F57637FF98FF9DFC40FF77DD1BFF26" box="[984,1051,150,176]" name="Brazil" pageId="5" pageNumber="446">Brasil</collectingCountry>
<quantity id="161A9B42FF98FF9DFBBBFF77DD63FF26" box="[1059,1123,150,176]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="5" pageNumber="446" unit="m" value="980.0">
<elevation id="5ACFD194FF98FF9DFBBBFF77DD63FF26" box="[1059,1123,150,176]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="5" pageNumber="446" unit="m" value="980.0">980m</elevation>
</quantity>
XI/74 \
<collectorName id="7C175371FF98FF9DFB5CFF76DC5FFF26" box="[1220,1375,150,177]" pageId="5" pageNumber="446">M. Alvarenga</collectorName>
leg.” (
<collectionCode id="B7F3AE62FF98FF9DFF07FF5BD9EFFF42" box="[159,239,186,212]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
).
<emphasis id="E396EAB5FF98FF9DFE99FF5BD897FF42" bold="true" box="[257,407,186,212]" pageId="5" pageNumber="446">
<typeStatus id="0E598805FF98FF9DFE99FF5BD87DFF42" box="[257,381,186,212]" pageId="5" pageNumber="446" type="paratype">Paratypes</typeStatus>
:
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</emphasis>
♀, same data of
</materialsCitation>
<materialsCitation id="618A3CFAFF98FF9DFDD7FF5BDA17FF42" box="[591,791,186,212]" collectionCode="DZUP" country="Brazil" pageId="5" pageNumber="446" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<typeStatus id="0E598805FF98FF9DFDD7FF5BDBB1FF42" box="[591,689,186,212]" pageId="5" pageNumber="446" type="holotype">holotype</typeStatus>
(
<collectionCode id="B7F3AE62FF98FF9DFD27FF5BDA0FFF42" box="[703,783,186,212]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="5" pageNumber="446" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="99F8652CFF98FF9DFF5FFF3FDB64FED6" pageId="5" pageNumber="446" type="discussion">
<paragraph id="D15D36A7FF98FF9DFF5FFF3FDB64FED6" blockId="5.[151,1436,150,320]" pageId="5" pageNumber="446">
<emphasis id="E396EAB5FF98FF9DFF5FFF3FD83BFF6E" bold="true" box="[199,315,222,248]" pageId="5" pageNumber="446">Remarks.</emphasis>
<taxonomicName id="16E24D24FF98FF9DFEDCFF3FDB26FF6E" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[324,550,222,248]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="birama" status="sp. nov.">
<emphasis id="E396EAB5FF98FF9DFEDCFF3FDB26FF6E" box="[324,550,222,248]" italics="true" pageId="5" pageNumber="446">Gracilipona birama</emphasis>
</taxonomicName>
<emphasis id="E396EAB5FF98FF9DFDB6FF3FDB89FF6E" bold="true" box="[558,649,222,248]" pageId="5" pageNumber="446">
<taxonomicNameLabel id="F8A557CEFF98FF9DFDB6FF3FDB89FF6E" box="[558,649,222,248]" pageId="5" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="16E24D24FF98FF9DFCBAFF3EDAB5FF6E" authorityName="Domahovski &amp; Gonçalves &amp; Takiya" authorityYear="2024" box="[802,949,223,249]" class="Insecta" family="Cicadellidae" genus="Gracilipona" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="tetrarama" status="sp. nov.">
<emphasis id="E396EAB5FF98FF9DFCBAFF3EDAB5FF6E" box="[802,949,223,249]" italics="true" pageId="5" pageNumber="446">G. tetrarama</emphasis>
</taxonomicName>
<emphasis id="E396EAB5FF98FF9DFC26FF3FDD19FF6E" bold="true" box="[958,1049,222,248]" pageId="5" pageNumber="446">
<taxonomicNameLabel id="F8A557CEFF98FF9DFC26FF3FDD19FF6E" box="[958,1049,222,248]" pageId="5" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in color and morphology, but can be easily distinguished from the latter by the aedeagus with a single pair of processes and female sternite VII with lateral angles slightly produced posterad.
</paragraph>
</subSubSection>
</treatment>
</document>