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<mods:title id="37B68291C81E0AEC4C0D58079A065EAF">Revision of Afrotropical Suragina Walker, 1859 (Diptera, Athericidae)</mods:title>
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<mods:namePart id="EAB43E50C29882183A2DFCF9D0B71450">Muller, Burgert S.</mods:namePart>
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<mods:affiliation id="1277042FA3179ED5FFFFD5E57FFD59C4">Department Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, 9301, South Africa &amp; Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
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<mods:namePart id="96A33D119D5CA006FFFA77346751C8C0">Swart, Vaughn R.</mods:namePart>
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<mods:affiliation id="A1B10529AFBA6FF27D594EBD63419180">Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
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<mods:namePart id="1E04AE11B267E03FF7B107DDD5C294FF">Snyman, Louwrens P.</mods:namePart>
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<mods:affiliation id="E87EB5D6C40087ADFEACEDA1267F7368">Invertebrate Zoology, Royal Alberta Museum, 9810 103 A Avenue NW, Edmonton, Alberta, T 5 J 0 G 2, Canada &amp; Department of Veterinary Microbiology, University of Saskatchewan, 52 Campus Drive, Saskatoon, Saskatchewan, S 7 N 5 B 4, Canada</mods:affiliation>
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<mods:date id="C9951278BC15098C6431A4FBB97524AC">2024</mods:date>
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<subSubSection id="BF28904959F68E8CF6D9AD3DC57EF264" type="nomenclature">
<paragraph id="F1263FBA3322BB8F7854AAF2DB15A3ED">
<taxonomicName id="0D5837E55938A058BD18DC2DBDD59CE6" ID-CoL="7QF2" authority="Walker, 1859" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="F18B891E3C7A4EBDE1EC8AB1C2F5CA3F" italics="true">Suragina</emphasis>
Walker, 1859
</taxonomicName>
</paragraph>
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<subSubSection id="604B20C5540523D45DED0159F9FAECDB" type="reference_group">
<paragraph id="939C57D4505717FB21031B08254EAAB5">
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<treatmentCitation id="45A15F86480E700ADDE36BB89E55C7DD">
<taxonomicName id="E781E2D585E4A69361060DB5DA0956F6" ID-CoL="7QF2" authority="Walker, 1859: 110" authorityName="Walker" authorityPageNumber="110" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="E8F5A56EE563F59166FBE22DE6B25462" italics="true">Suragina</emphasis>
Walker, 1859: 110
</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EWAAE" type="type_taxon">
<paragraph id="9D7EADDB9DC9D7C04EF946F1CCB67314">
<heading id="7F1A5BB323B940F10128DCEE29D3DC59" reason="title">
<typeStatus id="00224633B3D5190A0F716881E5936765">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="B751513A980CCB5ABE8318F58B22BCA0">
<taxonomicName id="DCAC3A3824E3EFAA97841DE3F452A78A" authority="Walker, 1859" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="illucens">
<emphasis id="4513B6F0015BB9B2A10C4EBA2E82DFB5" italics="true">Suragina illucens</emphasis>
Walker, 1859
</taxonomicName>
by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHBAE" type="diagnosis">
<paragraph id="F5B0C1ACEAEEF18E6C6B7AE961D75604">
<heading id="C68F2BD3C13360F8ABB7CF383F008539" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="2409175B46EC2EAE4DAC426587DD90AA">
<taxonomicName id="B4C1FEA39898519FD63581B705E98C62" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="06D2D153AC08CF201BD0609CB9C2B356" italics="true">Suragina</emphasis>
</taxonomicName>
is most similar to the genus
<taxonomicName id="3B056634E8C8872C3FB773118B4FAC37" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="83745B1538BBA05F5630DA4B7D0DFF46" italics="true">Atrichops</emphasis>
</taxonomicName>
with which it shares elongated legs and the hind coxa with a stout apical point or spine-like projection on its anteroventral surface. Similarly,
<taxonomicName id="00B309CD2EBD21A8C4A45C112D10E104" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="36DCD0F41BFF6C5A78397225C4F4E4B0" italics="true">Atrichops</emphasis>
</taxonomicName>
is also haematophagous, with well-developed mandibles. However,
<taxonomicName id="84B1CE96F68C46013F916BE4405F06A5" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="7F40E0F3B284EF4496A33630C124663A" italics="true">Atrichops</emphasis>
</taxonomicName>
has a knoblike proepimeral process (somewhat reduced in Afrotropical species) that is absent in
<taxonomicName id="D7CE2F7B47E584DC883C572C79A17621" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="62FC763992ED06643208B9650D7FD990" italics="true">Suragina</emphasis>
</taxonomicName>
. Additionally,
<taxonomicName id="CC191D37491629012EBA72A2F1ECCD64" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="72366A1E7F8A2DC3D2D4812017D49C3B" italics="true">Suragina</emphasis>
</taxonomicName>
has its frons usually contrasting velvety-black on the upper half and silvery-grey on the lower half (especially evident in
<specimenCount id="476643D000CE5C7CFDDCB7BAF1052BBB" count="1" type="female"></specimenCount>
), whereas the frons in
<taxonomicName id="87E7AE9407F43DB345A569B628B83EFF" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="71D35709E02DDF035C0E59B8B9EBD2E7" italics="true">Atrichops</emphasis>
</taxonomicName>
is more concolorous. The antennal bases are comparatively widely separated in
<taxonomicName id="1C380FED5D0E888CA4129C31BA4B0D01" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="AA86DB972C0B29FC8A036E3331324BCA" italics="true">Suragina</emphasis>
</taxonomicName>
, and close together in
<taxonomicName id="6EA95D13FDC98742E22E1E6C9CBBA977" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="6A03A387C51D216FB2F982E37CB52159" italics="true">Atrichops</emphasis>
</taxonomicName>
.
<taxonomicName id="547DC69A11BC4AC8E13E3B0EF57A8BE0" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="FF5401C1A1416F9C3826D917B39026F7" italics="true">Suragina</emphasis>
</taxonomicName>
typically has a tibial spur ratio of 0: 2: 2 (1: 2:
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some specimens of
<taxonomicName id="53C14E16C1DBFD63C1DC2838E2A77480" authority="(Bequaert, 1921))" authorityName="" baseAuthorityName="Bequaert" baseAuthorityYear="1921" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="binominata">
<emphasis id="CBE2CCDAA34759DB98548E08CA3956B0" italics="true">S. binominata</emphasis>
(
<bibRefCitation id="42C7876425ED269181993E14F0E1F325" DOI="10.1155/1921/95269" author="Bequaert J" issue="1" journalOrPublisher="Psyche (Cambridge, Massachusetts)" pagination="1-7" refId="B2" refString="Bequaert J (1921) Atherix braunsi nov. sp., a South African leptid with gregarious habits (Diptera). Psyche (Cambridge, Massachusetts) 28 (1): 1 - 7. https: // doi. org / 10.1155 / 1921 / 95269" title="Atherix braunsi nov. sp., a South African leptid with gregarious habits (Diptera)." volume="28" year="1921">Bequaert, 1921</bibRefCitation>
))
</taxonomicName>
, while
<taxonomicName id="251E3D11359CB14AC0DDD13ABC2A62C2" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="02C3DED16B3C8D4C792878570BB15E8A" italics="true">Atrichops</emphasis>
</taxonomicName>
has a ratio of 0: 1: 2. There is also a substantial difference in the morphology of the male terminalia, with
<taxonomicName id="C223F7C40DF7D4D1A4D9E74B2F6362DC" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="BA9B525BC1E386D1DDDA7BF559E72261" italics="true">Suragina</emphasis>
</taxonomicName>
having the gonostylus inserted apically and the parameral sheath simple apically, compared to
<taxonomicName id="E13872DDD8167E04BCCA01CF0FAD5F2A" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="89CE5ABC5FC91323781EB0AEEAA4F7A7" italics="true">Atrichops</emphasis>
</taxonomicName>
having the gonostylus inserted subapically, sometimes even medially on the gonocoxite (e. g.
<taxonomicName id="B04400864B4B144142086FA9F37BFA6D" authority="Muller, 2023" authorityName="Muller" authorityYear="2023" class="Insecta" family="Athericidae" genus="Atrichops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="EDE95BACC8C3130B6FB3FE2713F461CB" italics="true">Atrichops intermedius</emphasis>
<bibRefCitation id="9DD576B9861AA44C9D89D2FB2B8685DF" DOI="10.3897/afrinvertebr.64.113133" author="Muller BS &amp; Swart VR &amp; Snyman LP" issue="3" journalOrPublisher="African Invertebrates" pagination="303-322" refId="B14" refString="Muller BS, Swart VR, Snyman LP (2023) Afrotropical Atrichops Verrall (Diptera, Athericidae) with description of a new species. African Invertebrates 64 (3): 303 - 322. https: // doi. org / 10.3897 / afrinvertebr. 64.113133" title="Afrotropical Atrichops Verrall (Diptera, Athericidae) with description of a new species." volume="64" year="2023">
Muller,
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2023
<emphasis id="592360825D65305FF0EE608C6438FDA8" italics="true">in</emphasis>
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</bibRefCitation>
</taxonomicName>
<bibRefCitation id="72DC9D5B3C0ECF35DA682511F9EBD928" DOI="10.3897/afrinvertebr.64.113133" author="Muller" etAl="et al." firstAuthor="Muller" issue="3" journalOrPublisher="African Invertebrates" pagination="303-322" refId="B14" refString="Muller BS, Swart VR, Snyman LP (2023) Afrotropical Atrichops Verrall (Diptera, Athericidae) with description of a new species. African Invertebrates 64 (3): 303322. https://doi.org/10.3897/afrinvertebr.64.113133" title="Afrotropical Atrichops Verrall (Diptera, Athericidae) with description of a new species." volume="64" year="2023">Muller et al. 2023</bibRefCitation>
) and the parameral sheath ending with outward projections.
<bibRefCitation id="C0B8D6619BB49B0655E4EA182C221949" author="Woodley NE" editor="Kirk-Spriggs AH &amp; Sinclair BJ" firstAuthor="Woodley" journalOrPublisher="Volume 2. Nematocerous Diptera and lower Brachycera. Suricata 5. South African National Biodiversity Institute, Pretoria" page="887" pagination="885-891" refId="B28" refString="Woodley NE (2017) 38. Athericidae (Water Snipe Flies). In: Kirk-Spriggs AH, Sinclair BJ (Eds) Manual of Afrotropical Diptera. Volume 2. Nematocerous Diptera and lower Brachycera. Suricata 5. South African National Biodiversity Institute, Pretoria, 885891." title="38. Athericidae (Water Snipe Flies)." volumeTitle="Manual of Afrotropical Diptera." year="2017">Woodley (2017: 887)</bibRefCitation>
provided a key to the known genera of Afrotropical
<taxonomicName id="9FDF52C782AD61561DA765FE54BF80CB" authorityName="Stuckenberg" authorityYear="1973" class="Insecta" family="Athericidae" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="family">Athericidae</taxonomicName>
.
</paragraph>
<paragraph id="05766FDC5935EEE075FF677F080CEE62">
<title id="73209C8DCA86FACBFF4427F1EACDB345">
Key to the Afrotropical species of
<taxonomicName id="462E4A06606C91AE6D969B88BE2D2495" authority="Walker" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="B96EFA85A22D84DE2CFBE2A4489013F7" italics="true">Suragina</emphasis>
Walker
</taxonomicName>
(excluding
<taxonomicName id="7E6E6B7F4F591A8FCE3975631D26C2BA" baseAuthorityName="Speiser" baseAuthorityYear="1914" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="disciclara">
<emphasis id="515D8340FC2A54299E4CD86234E9A36B" italics="true">S. disciclara</emphasis>
</taxonomicName>
and
<taxonomicName id="B5543910CC51A97533BF5F5F3E4BFEEF" baseAuthorityName="Lindner" baseAuthorityYear="1925" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pilitarsis">
<emphasis id="8859DB078DDC2DFF8107D5A4F0C2B68A" italics="true">S. pilitarsis</emphasis>
</taxonomicName>
)
</title>
</paragraph>
<paragraph id="50FB964837058FEA5A427F823D985997">
<table id="TID0EHJAC" rules="all">
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<td id="5AFD6DC3B45F8F1C0F3C382C66BD24D3" colspan="1" rowspan="1">
Scutum and pleura with ground colour mostly orange-yellow (e. g. Fig.
<figureCitation id="C4031CE69121AE8B4015FE8881CFB7C7" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">1</figureCitation>
), if scutum with prominent dark median vitta (e. g. Fig.
<figureCitation id="DB616E38ED0364057E123E34D6852525" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">2</figureCitation>
) then pleura darker in colouration (Figs
<figureCitation id="54E2FB5D3CF137D0363C335E5C5102E4" captionStart="Figures 1417" captionStartId="F3" captionText="Figures 1417. Suragina Walker spp. lateral habitus: S. binominata (Bequaert): 14 ♂ holotype (MLUH) 15 ♀ (BMSA (D) 124083); S. copelandi Muller, sp. nov.: 16 ♂ holotype (BMSA (D) 83426) 17 ♀ paratype (BMSA (D) 84688). 14 Copyright ZNS Halle, January 30, 2024. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures14-17" httpUri="https://binary.pensoft.net/fig/1205865">14</figureCitation>
,
<figureCitation id="C494B32FEBD3534EF09C9A4A8B6928A4" captionStart="Figures 1417" captionStartId="F3" captionText="Figures 1417. Suragina Walker spp. lateral habitus: S. binominata (Bequaert): 14 ♂ holotype (MLUH) 15 ♀ (BMSA (D) 124083); S. copelandi Muller, sp. nov.: 16 ♂ holotype (BMSA (D) 83426) 17 ♀ paratype (BMSA (D) 84688). 14 Copyright ZNS Halle, January 30, 2024. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures14-17" httpUri="https://binary.pensoft.net/fig/1205865">15</figureCitation>
)
</td>
<td id="49A4DFCB9210879A15385349D1C2E58C" colspan="1" rowspan="1">
<emphasis id="308E9E36C78C36C2625F37ACC8C17C9F" bold="true">2</emphasis>
</td>
</tr>
<tr id="097DB0204D6B2FBE78ED276D42D433F6">
<td id="AA721B6D19E31014A1AF200F3C52019C" colspan="1" rowspan="1"></td>
<td id="048645745C83FF2F6C8FF72DE09982C0" colspan="1" rowspan="1">
Scutum and pleura with majority of surface brown to almost black (e. g. Figs
<figureCitation id="5E1F2A3993E4EDC1FE3D7FFC1C1D4B67" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">3</figureCitation>
,
<figureCitation id="5E051CEB359C3B672D9B48A7007C9E29" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">6</figureCitation>
), limited orange-yellow colouration
</td>
<td id="14F060B65167F64D36AC961987A883A9" colspan="1" rowspan="1">
<emphasis id="C9FE853A916F2595BB0AE08D55BF364A" bold="true">8</emphasis>
</td>
</tr>
<tr id="8E70FB9997964B9CB36AD302E64A309E">
<td id="F98BE0B36542BA6ECEAB6E07473C0E98" colspan="1" rowspan="1">2</td>
<td id="9BD224B91469916D29207DAF6F7834CF" colspan="1" rowspan="1">
Notopleural area posterior to postpronotal lobe and anterior to transverse suture orange-yellow (e. g. Fig.
<figureCitation id="82963E06CEFBDF34D353DEC2F645A181" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">2</figureCitation>
), at most with a thin brown band along posterior edge of postpronotal lobe, no pruinosity in area
</td>
<td id="D33FB4E2FFA944902BBFA8E48B9C1A27" colspan="1" rowspan="1">
<emphasis id="D74667CF813726CE4F7D1FD2DC4C574B" bold="true">3</emphasis>
</td>
</tr>
<tr id="6DD7FCF61370A260CE38AB8D68BDAD6A">
<td id="D8A5449B50816C52B6CA6ED5D96BF6EE" colspan="1" rowspan="1"></td>
<td id="D334174D33479BA047845B1716932296" colspan="1" rowspan="1">
Notopleural area posterior to postpronotal lobe and anterior to transverse suture with a brownish marking on surface (e. g. Fig.
<figureCitation id="5E62C0E9346A3F442D4F8E1754BF7CCD" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">4</figureCitation>
), appearing bluish- to silver-grey pruinose in dorsal view, brown in lateral view (dependent on viewing angle)
</td>
<td id="4C1386BC48B4C94315480A30FDBD89F6" colspan="1" rowspan="1">
<emphasis id="25BB0D8E9B40EFF7D109324132878403" bold="true">5</emphasis>
</td>
</tr>
<tr id="D8666AE0951AB36DEB12938EE5D34317">
<td id="2490E36A0B3FA8D0AF6C52F6B9271A00" colspan="1" rowspan="1">3</td>
<td id="A8F0CE155AF6EF80CCF8464AF254A994" colspan="1" rowspan="1">
Scutum with a prominent dark median vitta (Fig.
<figureCitation id="2C0BDDA5B365EF14AF12897D542BD924" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">2</figureCitation>
)
</td>
<td id="2919DE1C26BECCA053984E4843321EA8" colspan="1" rowspan="1">
<taxonomicName id="C4DD110E27BC93E67C1F5D943B77B480" authority="(Bequaert)" authorityName="" baseAuthorityName="Bequaert" baseAuthorityYear="1921" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="binominata">
<emphasis id="F4AACE6D9C897ECA0F29BC51A43BB887" bold="true">
<emphasis id="4EA5DD07960D1D4953E69A687FD5AA0C" bold="true" italics="true">S. binominata</emphasis>
(Bequaert)
</emphasis>
</taxonomicName>
(in part, ♀)
</td>
</tr>
<tr id="07E156F5B17177D6E7AA91E0A933A3B3">
<td id="048D2FB75B4D272D82459077787C406A" colspan="1" rowspan="1"></td>
<td id="89698E16278F258E1BEFE6FC1EC86609" colspan="1" rowspan="1">
Scutum without a median vitta (Fig.
<figureCitation id="19989246D2B1BBDF63442CF7BF1FF139" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">1</figureCitation>
)
</td>
<td id="98696C01D5224F1427D438AB6B4D7822" colspan="1" rowspan="1">
<emphasis id="248DC3689CE072E603483EA447C58580" bold="true">4</emphasis>
</td>
</tr>
<tr id="FEA505D64548682CFB00A7EF2029B268">
<td id="05EF5A066D2EB51FC0D8F6EB916EC57E" colspan="1" rowspan="1">4</td>
<td id="40CB1F0E8CF2D2CA40CB43E52A7FAEBD" colspan="1" rowspan="1">
Fore and hind tibiae dark brown to black; lateral margins of tergites 24 dark (Figs
<figureCitation id="4614C8A6A0F4C447E645FF2A21952059" captionStart="Figures 1417" captionStartId="F3" captionText="Figures 1417. Suragina Walker spp. lateral habitus: S. binominata (Bequaert): 14 ♂ holotype (MLUH) 15 ♀ (BMSA (D) 124083); S. copelandi Muller, sp. nov.: 16 ♂ holotype (BMSA (D) 83426) 17 ♀ paratype (BMSA (D) 84688). 14 Copyright ZNS Halle, January 30, 2024. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures14-17" httpUri="https://binary.pensoft.net/fig/1205865">16</figureCitation>
,
<figureCitation id="3B0BC5ED2974D3352920E91945A83F4F" captionStart="Figures 1417" captionStartId="F3" captionText="Figures 1417. Suragina Walker spp. lateral habitus: S. binominata (Bequaert): 14 ♂ holotype (MLUH) 15 ♀ (BMSA (D) 124083); S. copelandi Muller, sp. nov.: 16 ♂ holotype (BMSA (D) 83426) 17 ♀ paratype (BMSA (D) 84688). 14 Copyright ZNS Halle, January 30, 2024. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures14-17" httpUri="https://binary.pensoft.net/fig/1205865">17</figureCitation>
)
</td>
<td id="9999E5A2DB41A00AEC23C341CF1F8AE4" colspan="1" rowspan="1">
<emphasis id="5E5BE41EC883CEC792138D017257E3D8" bold="true">
<taxonomicName id="C26A128762F26192B73481905B6BC471" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="copelandi" status="sp. nov.">
<emphasis id="0D075CBEB50A2A3B0B1901E0BBA1072C" bold="true" italics="true">S. copelandi</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="113D461CF698BC7DB67397934BFC2F12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
</tr>
<tr id="1A0061D12A8AF2AAC86FBCED6A895B27">
<td id="33B8ABA7CA02B2EC1F52843196C99D4B" colspan="1" rowspan="1"></td>
<td id="42CD5139AF2C2BD06B67047DD1FAB933" colspan="1" rowspan="1">
Fore and hind tibiae yellow to orange-yellow, some specimens with apical half of front tibia darker; lateral margins of tergites 24 concolorous with rest of abdomen or only lateral margin of tergite 2 a slightly darker colour (Figs
<figureCitation id="A4B1BE33CA3A2747A1FBDD143DC5F09C" captionStart="Figures 913" captionStartId="F2" captionText="Figures 913. Suragina Walker spp. lateral habitus: S. agramma (Bezzi): 9 ♂ (NMSA - DIP 028196) 10 ♀ holotype (NMSA - DIP 028199); S. bezzii (Curran): 11 ♂ (SAM-DIP-A 018382) 12 ♀ (SAM-DIP-A 018410); S. bilobata Muller, sp. nov.: 13 a ♀ paratype 1 st flagellomere (CSCA) 13 b ♀ holotype (CSCA). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures9-13" httpUri="https://binary.pensoft.net/fig/1205864">9</figureCitation>
,
<figureCitation id="E65618244AD491D7D47003D50E75CAC0" captionStart="Figures 913" captionStartId="F2" captionText="Figures 913. Suragina Walker spp. lateral habitus: S. agramma (Bezzi): 9 ♂ (NMSA - DIP 028196) 10 ♀ holotype (NMSA - DIP 028199); S. bezzii (Curran): 11 ♂ (SAM-DIP-A 018382) 12 ♀ (SAM-DIP-A 018410); S. bilobata Muller, sp. nov.: 13 a ♀ paratype 1 st flagellomere (CSCA) 13 b ♀ holotype (CSCA). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures9-13" httpUri="https://binary.pensoft.net/fig/1205864">10</figureCitation>
)
</td>
<td id="26F08BFB32BEE0965A2529E7DCCA6EB0" colspan="1" rowspan="1">
<taxonomicName id="D65478EB331AEE9FD5B98306CA8215A8" authority="(Bezzi)" authorityName="ICIPE" baseAuthorityName="Bezzi" baseAuthorityYear="1926" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="agramma">
<emphasis id="3E8E1DBF19BA748CD5EC19FAEF54838C" bold="true">
<emphasis id="0D19E85F9F01AEE1349A2B376E075F4B" bold="true" italics="true">S. agramma</emphasis>
(Bezzi)
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="645CD15F959CC8DF656CB1FD19967457">
<td id="701FA90A48559C0462F51312566C6668" colspan="1" rowspan="1">5</td>
<td id="30CF48AEE9D5F695E453A8B421798347" colspan="1" rowspan="1">Scutum with a prominent dark median vitta, sometimes with additional dark markings on pre- and postsutural lateral areas</td>
<td id="02EDBA10161AA08D55B76B3728918B3F" colspan="1" rowspan="1">
<emphasis id="EACD8E9EDCC6AFB91894765D6BCDE3BC" bold="true">6</emphasis>
</td>
</tr>
<tr id="7E01EC3666D0C989E27880B2000A2CC7">
<td id="495CA63D76C0F1E2D220A6F6E5C2FDEE" colspan="1" rowspan="1"></td>
<td id="346D009C3FEE46EDC22AB6FC7CC85D3A" colspan="1" rowspan="1">
Scutum at most with two feint dorsocentral bluish-grey pruinose vittae (e. g. Fig.
<figureCitation id="F29DEE85BF9BA002C2CA99A7A5179680" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">5</figureCitation>
)
</td>
<td id="6B1A702AA72F2CBE7864A6AA4F240C17" colspan="1" rowspan="1">
<emphasis id="2E8C107C23889C42DCFD15332EF16B8B" bold="true">7</emphasis>
</td>
</tr>
<tr id="D6DA3411BE8116F2B4845E2309358421">
<td id="55DC482891DEA8C3899BB7F81CF2C81A" colspan="1" rowspan="1">6</td>
<td id="5DF3EAAE7F6640E0B5A38F31C96639DB" colspan="1" rowspan="1">
Scutum with additional dark markings on pre- and postsutural lateral areas in addition to notopleural marking (Fig.
<figureCitation id="C6F0D9C2346EDFF44D96D976F6F9167B" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">4</figureCitation>
); fore tibia dark brown to almost black
</td>
<td id="7E76501BCC92B489B3E706B337A62779" colspan="1" rowspan="1">
<taxonomicName id="92E4B92CB9C84057260652691B03BAA7" authority="(Oldroyd)" baseAuthorityName="Oldroyd" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="falsa">
<emphasis id="946F3A2CB1DA4C24BF328EAC5535DC6A" bold="true">
<emphasis id="96EBD50E0DACE9AB31779C93B2B66A87" bold="true" italics="true">S. falsa</emphasis>
(Oldroyd)
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="64CD8688A5E3A7FD8E778ADCB347DAFC">
<td id="EC2EFE3E8A01AFF747AA6C4B08FEC936" colspan="1" rowspan="1"></td>
<td id="72177B8A260F3AB08254C45FD82E0F95" colspan="1" rowspan="1">
Scutum without any additional pre- or postsutural dark markings apart from central vitta; fore and hind tibiae light brownish-yellow to orange-yellow (Figs
<figureCitation id="C146C5C12D74C5C001FAA8626521AEF8" captionStart="Figures 2730" captionStartId="F7" captionText="Figures 2730. Suragina Walker spp. lateral habitus: S. monogramma (Bezzi): 27 ♂ (NMSA - DIP 028180) 28 ♀ (NMSA - DIP 158385); S. mulanjeensis Muller, sp. nov.: 29 ♂ holotype (NMSA - DIP 158398) 30 ♀ paratype (NMSA - DIP 158394). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures27-30" httpUri="https://binary.pensoft.net/fig/1205869">27</figureCitation>
,
<figureCitation id="2BAB1ACCAEBFB5B87BEFC7E880173794" captionStart="Figures 2730" captionStartId="F7" captionText="Figures 2730. Suragina Walker spp. lateral habitus: S. monogramma (Bezzi): 27 ♂ (NMSA - DIP 028180) 28 ♀ (NMSA - DIP 158385); S. mulanjeensis Muller, sp. nov.: 29 ♂ holotype (NMSA - DIP 158398) 30 ♀ paratype (NMSA - DIP 158394). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures27-30" httpUri="https://binary.pensoft.net/fig/1205869">28</figureCitation>
)
</td>
<td id="F923D98E2AF61C6E73848026A2DA37E8" colspan="1" rowspan="1">
<taxonomicName id="4EBF3A1E049C40B27FF549EB881D58E9" authority="(Bezzi)" baseAuthorityName="Bezzi" baseAuthorityYear="1926" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="monogramma">
<emphasis id="ADD8E9B96E8A222AA7FAFAD8B23C40A9" bold="true">
<emphasis id="D8B34D59C83BD6BCC397F2966F788414" bold="true" italics="true">S. monogramma</emphasis>
(Bezzi)
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="3A66AA85AFBC342CBCD486396F113C1F">
<td id="A953D64C63A471B66E000C8E8D901404" colspan="1" rowspan="1">7</td>
<td id="9AC41674439BC607A3C613B1A43C5F70" colspan="1" rowspan="1">
Wing stigma dark brown with similarly dark substigmal mark running down to base of cell
<emphasis id="BC7C1F29F2B96A7D675583D5138C55A3" italics="true">
m
<subScript id="A64B041F8A4B4684FB13954FF9ECE9B1">3</subScript>
</emphasis>
(Fig.
<figureCitation id="7E7ADF5965CFAB45DABEFA11841C44ED" captionStart="Figures 4348" captionStartId="F11" captionText="Figures 4348. Suragina Walker spp. wing dorsal view: 43 S. dimidiatipennis (Brunetti) ♀ holotype (NHMUK 014064158) 44 S. falsa Oldroyd ♀ (NHMUK 014064160) 45 S. freidbergi Muller, sp. nov. ♀ paratype (BMSA (D) 92158) 46 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158445) 47 S. malavaensis Muller, sp. nov. ♂ (ICIPE) 48 S. milloti (Séguy) ♀ (CSCA). 43, 44 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures43-48" httpUri="https://binary.pensoft.net/fig/1205873">48</figureCitation>
); prescutellar area orange-yellow without any pruinosity; Madagascan endemic
</td>
<td id="0ECA843356FF78E4128B996BF004826B" colspan="1" rowspan="1">
<taxonomicName id="EFD8F7731838349F7145002295D103DC" authority="(Seguy)" authorityName="CSCA" baseAuthorityName="Seguy" baseAuthorityYear="1951" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="milloti">
<emphasis id="D6744EEEBB1C3263646B0ED1634BDD89" bold="true">
<emphasis id="69BF4811A7F2246E7F38354701F3EC67" bold="true" italics="true">S. milloti</emphasis>
(Séguy)
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="00149235073050411F19E2FAD22E8748">
<td id="DC40EF4CC8E10A803ED49D97EB738F84" colspan="1" rowspan="1"></td>
<td id="349B5CA55E74A188E9338E36220FBD8F" colspan="1" rowspan="1">
Wing stigma light brown, with only some darker suffusions over posterior cells (Fig.
<figureCitation id="DBF84442EF3F7F684B1AC0519889781F" captionStart="Figures 4954" captionStartId="F12" captionText="Figures 4954. Suragina Walker spp. wing dorsal view: 49 S. monogramma (Bezzi) ♀ (BMSA (D) 129988) 50 S. mulanjeensis Muller, sp. nov. ♀ (BMSA (D) 92374) 51 S. nigromaculata Muller, sp. nov. ♂ holotype (NHMUK 014064156) 52 S. pauliani ♂ paratype (NMSA - DIP 028172) 53 S. semiobscura Muller, sp. nov. ♀ paratype (ICIPE) 54 S. zombaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158451). 51 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures49-54" httpUri="https://binary.pensoft.net/fig/1205874">50</figureCitation>
); prescutellar area with bluish-grey pruinosity similar to that of notopleuron (Fig.
<figureCitation id="1033D990B659A77952D40092E27639D5" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">5</figureCitation>
); mainland Africa
</td>
<td id="201623D5F062078B03425959DCF13199" colspan="1" rowspan="1">
<emphasis id="E1447B0099EF3B8DB0B6528E542DE9DE" bold="true">
<taxonomicName id="430883953189BF73B010AE28CA0473E3" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="mulanjeensis" status="sp. nov.">
<emphasis id="0161EED329232950C54DF2CFE4A35839" bold="true" italics="true">S. mulanjeensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="8C68552F61F31A3C51CE6A4F88FF3ECE" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
</tr>
<tr id="4ED69662A2E1FA2BD7C7C8FEEDBCD153">
<td id="624A54AF1B2DC5E5D4CF3D35E15163EF" colspan="1" rowspan="1">8</td>
<td id="7457D5A130CFBE2BE20C3740F176DE4E" colspan="1" rowspan="1">
Wing stigma dark brown with similarly dark, but irregularly shaped substigmal mark running down to base of cell
<emphasis id="599E77ADCBF630F54A8EDDA18F4629B1" italics="true">
m
<subScript id="0916642D72E63106E9CDDCF6833124FF">3</subScript>
</emphasis>
; at least apical third of wing with brown suffused appearance, area between substigmal mark and suffusion appearing to form a hyaline band (e. g. Figs
<figureCitation id="2AE171E094523B25B10D93CF50A7F66C" captionStart="Figures 3742" captionStartId="F10" captionText="Figures 3742. Suragina Walker spp. wing dorsal view: 37 S. agramma (Bezzi) ♀ (BMSA (D) 124627) 38 S. bezzii (Curran) ♂ (SAM-DIP-A 018410) 39 S. bilobata Muller, sp. nov. ♀ paratype (CSCA) 40 S. copelandi Muller, sp. nov. ♀ paratype (BMSA (D) 83423); S. binominata (Bequaert): 41 holotype ♂ (MLUH) 42 ♀ (NMSA - DIP 158446). 41 Copyright ZNS Halle, January 30, 2024. Abbreviations: br basal radial cell; bm basal medial cell; cua anterior cubital cell; CuA + CuP anterior branch of cubital vein + posterior branch of cubital vein; d discal cell; M 1 first branch of media; m 1 first medial cell; M 2 second branch of media; m 2 second medial cell; M 3 third branch of media; m 3 third medial cell; R 1 anterior branch of radius; r 1 first radial cell; R 2 + 3 second branch of radius; r 2 + 3 second radial cell; R 4 upper branch of third branch of radius; R 5 lower branch of third branch of radius; Sc subcostal vein; sc subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures37-42" httpUri="https://binary.pensoft.net/fig/1205872">39</figureCitation>
,
<figureCitation id="730C01E2C7FC7250BE9E6270731032D8" captionStart="Figures 4348" captionStartId="F11" captionText="Figures 4348. Suragina Walker spp. wing dorsal view: 43 S. dimidiatipennis (Brunetti) ♀ holotype (NHMUK 014064158) 44 S. falsa Oldroyd ♀ (NHMUK 014064160) 45 S. freidbergi Muller, sp. nov. ♀ paratype (BMSA (D) 92158) 46 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158445) 47 S. malavaensis Muller, sp. nov. ♂ (ICIPE) 48 S. milloti (Séguy) ♀ (CSCA). 43, 44 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures43-48" httpUri="https://binary.pensoft.net/fig/1205873">4547</figureCitation>
)
</td>
<td id="D5E5D9C8B012D64609BF317EF15D681E" colspan="1" rowspan="1">
<emphasis id="1718BC96FCAC05813BC4E784EA6945FC" bold="true">9</emphasis>
</td>
</tr>
<tr id="106AE2FD392998029CFFE347DDDFE602">
<td id="4255C66704BE01B4514341952A18F324" colspan="1" rowspan="1"></td>
<td id="D281B9751072A90C04CE7B46914601F0" colspan="1" rowspan="1">
Wing stigma variable, darker substigmal mark present or absent, if present, then brown suffusion runs up to or near it, not visually forming a clear hyaline band (e. g. Figs
<figureCitation id="A239F1ED230067EC6932807E4C85BA1C" captionStart="Figures 3742" captionStartId="F10" captionText="Figures 3742. Suragina Walker spp. wing dorsal view: 37 S. agramma (Bezzi) ♀ (BMSA (D) 124627) 38 S. bezzii (Curran) ♂ (SAM-DIP-A 018410) 39 S. bilobata Muller, sp. nov. ♀ paratype (CSCA) 40 S. copelandi Muller, sp. nov. ♀ paratype (BMSA (D) 83423); S. binominata (Bequaert): 41 holotype ♂ (MLUH) 42 ♀ (NMSA - DIP 158446). 41 Copyright ZNS Halle, January 30, 2024. Abbreviations: br basal radial cell; bm basal medial cell; cua anterior cubital cell; CuA + CuP anterior branch of cubital vein + posterior branch of cubital vein; d discal cell; M 1 first branch of media; m 1 first medial cell; M 2 second branch of media; m 2 second medial cell; M 3 third branch of media; m 3 third medial cell; R 1 anterior branch of radius; r 1 first radial cell; R 2 + 3 second branch of radius; r 2 + 3 second radial cell; R 4 upper branch of third branch of radius; R 5 lower branch of third branch of radius; Sc subcostal vein; sc subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures37-42" httpUri="https://binary.pensoft.net/fig/1205872">38</figureCitation>
,
<figureCitation id="A9902A711637756763E2BC2C07C87E98" captionStart="Figures 4954" captionStartId="F12" captionText="Figures 4954. Suragina Walker spp. wing dorsal view: 49 S. monogramma (Bezzi) ♀ (BMSA (D) 129988) 50 S. mulanjeensis Muller, sp. nov. ♀ (BMSA (D) 92374) 51 S. nigromaculata Muller, sp. nov. ♂ holotype (NHMUK 014064156) 52 S. pauliani ♂ paratype (NMSA - DIP 028172) 53 S. semiobscura Muller, sp. nov. ♀ paratype (ICIPE) 54 S. zombaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158451). 51 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures49-54" httpUri="https://binary.pensoft.net/fig/1205874">54</figureCitation>
)
</td>
<td id="9A94FAB9EE83FBBF7549B36D4E188C3E" colspan="1" rowspan="1">
<emphasis id="74E79D82F5C536C967619B23994B582A" bold="true">12</emphasis>
</td>
</tr>
<tr id="516D71F6772D7294ED4B08A60CC83BCA">
<td id="36EB3CDE4460425F14BE6F52AC57C1DB" colspan="1" rowspan="1">9</td>
<td id="72C110F85EB73C0D63D40A974D2D7A99" colspan="1" rowspan="1">
Antennal 1
<superScript id="8A71AE5E79DA609BC34C2E5E5551B2BE">st</superScript>
flagellomere bilobate, c-shaped (Fig.
<figureCitation id="208F9358BEF4A8B252E578C810749A40" captionStart="Figures 913" captionStartId="F2" captionText="Figures 913. Suragina Walker spp. lateral habitus: S. agramma (Bezzi): 9 ♂ (NMSA - DIP 028196) 10 ♀ holotype (NMSA - DIP 028199); S. bezzii (Curran): 11 ♂ (SAM-DIP-A 018382) 12 ♀ (SAM-DIP-A 018410); S. bilobata Muller, sp. nov.: 13 a ♀ paratype 1 st flagellomere (CSCA) 13 b ♀ holotype (CSCA). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures9-13" httpUri="https://binary.pensoft.net/fig/1205864">13 a</figureCitation>
); Madagascan endemic
</td>
<td id="B5C0089681266384351FCE854A4CA8D9" colspan="1" rowspan="1">
<emphasis id="D0FA1DD8D17C55D8AB3F590147303C61" bold="true">
<taxonomicName id="8743A40EBEC2B3942DD841265AFF3424" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bilobata" status="sp. nov.">
<emphasis id="66669D50BEC9E4401D36608BF00345D8" bold="true" italics="true">S. bilobata</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="4E52D3DD6616D2A1B263353760759DC4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
</tr>
<tr id="ED139EAC98A6D50CD18EADE808937771">
<td id="F407EEC82B916C1A0FFCB7F17B347410" colspan="1" rowspan="1"></td>
<td id="FF185C19AD885FCCCC7A721DD61702AE" colspan="1" rowspan="1">
Antennal 1
<superScript id="F832E3976092FE0567DCEFFA7623388C">st</superScript>
flagellomere typical subreniform shape; mainland Africa
</td>
<td id="A414490716A45C499909B926801474F2" colspan="1" rowspan="1">
<emphasis id="9C4DD4ABE022638BAA74284FD435D797" bold="true">10</emphasis>
</td>
</tr>
<tr id="1ECF2809371E335ECAB98EB1669CDE3D">
<td id="2F63A28229FF4438D0EB4641868579A2" colspan="1" rowspan="1">10</td>
<td id="932F47D810E3D82C9DA6E8D215754439" colspan="1" rowspan="1">
Upper occiput with a pair of well-developed dark markings that run down to occipital foramen (Fig.
<figureCitation id="F072BF1A59C46EA4D7A3BC930C0472A4" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">7</figureCitation>
); mid femur dark brown, only yellow at extreme base and apex (Fig.
<figureCitation id="F9DB8264C7744B4BCFE20909EE421082" captionStart="Figures 2022" captionStartId="F5" captionText="Figures 2022. Suragina Walker spp. lateral habitus: S. freidbergi Muller, sp. nov.: 20 ♂ holotype (NMSA - DIP 158423) 21 ♀ paratype (NMSA - DIP 158424); S. liberiaensis Muller, sp. nov.: 22 ♂ holotype (NMSA - DIP 158443). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures20-22" httpUri="https://binary.pensoft.net/fig/1205867">22</figureCitation>
); all abdominal tergites dark brown, without any posterior grey pruinose bands
</td>
<td id="FCBCC31A1B1EF6CB6F44241FBB7116A3" colspan="1" rowspan="1">
<emphasis id="AE19EBDC9E532E94FDCE5A016796E918" bold="true">
<taxonomicName id="6740F7D23D37E9F87FB1F6BD8F2AEAB9" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="liberiaensis" status="sp. nov.">
<emphasis id="CB081D2A09DE4C34F1ED22E62DE32D62" bold="true" italics="true">S. liberiaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="B0E3C84A7FFCC1330C2DD00A450BAE2B" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
</tr>
<tr id="238AFCDCE1586A6609D9EEDED3ABD95F">
<td id="BC05A7F59C331A99FCF802D9D6E11DED" colspan="1" rowspan="1"></td>
<td id="BCFB8E18A08E04D1A710C21A0C7C3A3F" colspan="1" rowspan="1">Upper occiput with narrower dark markings, either subrectangular or subtriangular; mid femur yellow, at most brownish on extreme base; tergite 1 with subtriangular dark marking, appearance of other tergites variable</td>
<td id="E4DEB3A8539E4F919F016C013C241C39" colspan="1" rowspan="1">
<emphasis id="21E527E928FEB50314CA427D95C6676B" bold="true">11</emphasis>
</td>
</tr>
<tr id="0035E48A5E0CFC9C667B445C15B06560">
<td id="3820247068472DBFCAF8674ADCF449BC" colspan="1" rowspan="1">11</td>
<td id="DDE76517331A63DF70430D464D20E843" colspan="1" rowspan="1">
Upper occiput with a pair of narrow subrectangular dark markings (Fig.
<figureCitation id="35831D979D5819F46FCD97DF5309EADF" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">8</figureCitation>
); mid femur yellow (Figs
<figureCitation id="59875321843E63F9DFCB65D81841FB5B" captionStart="Figures 2022" captionStartId="F5" captionText="Figures 2022. Suragina Walker spp. lateral habitus: S. freidbergi Muller, sp. nov.: 20 ♂ holotype (NMSA - DIP 158423) 21 ♀ paratype (NMSA - DIP 158424); S. liberiaensis Muller, sp. nov.: 22 ♂ holotype (NMSA - DIP 158443). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures20-22" httpUri="https://binary.pensoft.net/fig/1205867">20</figureCitation>
,
<figureCitation id="08A124230B91DA10EE283BD6DFAA4911" captionStart="Figures 2022" captionStartId="F5" captionText="Figures 2022. Suragina Walker spp. lateral habitus: S. freidbergi Muller, sp. nov.: 20 ♂ holotype (NMSA - DIP 158423) 21 ♀ paratype (NMSA - DIP 158424); S. liberiaensis Muller, sp. nov.: 22 ♂ holotype (NMSA - DIP 158443). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures20-22" httpUri="https://binary.pensoft.net/fig/1205867">21</figureCitation>
); abdominal tergites yellowish-brown to dark brown, tergite 1 with a subtriangular marking, tergite 2 in some specimens with a fenestrated appearance, at least tergites 35 with posterior grey pruinose bands
</td>
<td id="9F8FD893FE652027FC9F2955EF773F55" colspan="1" rowspan="1">
<emphasis id="98038125CDA2F6E00641761323B24944" bold="true">
<taxonomicName id="D1F55C2908E81ED3971E8CB8790D8081" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="freidbergi" status="sp. nov.">
<emphasis id="93A9D4E680CC52803DCDE593B052663D" bold="true" italics="true">S. freidbergi</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="F670B95ED1BAB51D1340ECBCB673B7C4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
</tr>
<tr id="851F812EA9CE7D192432A693D4DEC0D7">
<td id="5D0051C65693736B8918B45067958779" colspan="1" rowspan="1"></td>
<td id="792A5AEA6399E8A076F2D051A6CBC589" colspan="1" rowspan="1">
Upper occiput with a subtriangular pair of well-developed dark markings (Fig.
<figureCitation id="AE0F9E8D877288C13FC0A03E81F3CBCF" captionStart="Figures 18" captionStartId="F1" captionText="Figures 18. Dorsal view of thorax (14) and head and thorax (58) of Suragina Walker spp. 1 S. agramma (Bezzi) ♀ (NMSA - DIP 028177) 2 S. binominata ♀ (NMSA - DIP 158408) 3 S. bilobata Muller, sp. nov. ♀ (CSCA) 4 S. falsa ♀ holotype (NHMUK 014064160) 5 S. mulanjeensis Muller, sp. nov. ♂ paratype (NMSA - DIP 162049) 6 S. malavaensis Muller, sp. nov. ♀ paratype (ICIPE) 7 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158442) 8 S. freidbergi Muller, sp. nov. ♀ paratype (NMSA - DIP 158425). 4 Copyright NHMUK under CC BY 4.0. Not to scale." figureDoi="10.3897/afrinvertebr.65.140524.figures1-8" httpUri="https://binary.pensoft.net/fig/1205863">6</figureCitation>
); mid femur yellow, at most with extreme base dark; abdomen with orange-yellow appearance; elongated dark median and lateral markings on tergites 2 and 3, remaining tergites orange-yellow without external markings
</td>
<td id="C3BE17D06413AEAE77409E758827D7DE" colspan="1" rowspan="1">
<emphasis id="0AA9142C398928146C64739110DFA0B7" bold="true">
<taxonomicName id="FA726893775138AE0508A2A865495AD8" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="malavaensis" status="sp. nov.">
<emphasis id="478C1E6CBB489D45E2BE645671149E3A" bold="true" italics="true">S. malavaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="B30A8CD3B17134FD53B2BEA3D56D6643" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
</tr>
<tr id="FB44DAEF06C650ABDB596C2120D3CB94">
<td id="4328A3B7A646DC49C015BC0BB84CE625" colspan="1" rowspan="1">12</td>
<td id="B5814C968B04E8259D62F26CB90A04EB" colspan="1" rowspan="1">Mid femur brown on at least two-thirds of surface, fore femur yellow on over two-thirds of surface</td>
<td id="242DBE1C6D957E8CDD5E62EB05C92EDE" colspan="1" rowspan="1">
<emphasis id="3C8A9C863B0C55396081557423FA3BBA" bold="true">13</emphasis>
</td>
</tr>
<tr id="BAB08FED822FCA69FF05A667055CD391">
<td id="EB097ACC7285C6E9232620C4F27FFC55" colspan="1" rowspan="1"></td>
<td id="2A1F87D613BA223DBD3D7BE90C86C34C" colspan="1" rowspan="1">Fore and mid femora yellow, at most brown on extreme apex and base</td>
<td id="D08F29CB89070760944AF2AC93731678" colspan="1" rowspan="1">
<emphasis id="7D3745E06C7576472253DB7A4714DA01" bold="true">14</emphasis>
</td>
</tr>
<tr id="767AD3F0CB5353BFDDC245A914FB172D">
<td id="B7A5B4C214B6F15EED02DCD236E3658D" colspan="1" rowspan="1">13</td>
<td id="2011CC8B34EF7B41B7B5BE222792590D" colspan="1" rowspan="1">
Wing with dark brown suffusion more pronounced on anterior half, lighter on posterior half (Fig.
<figureCitation id="FB1C60DFBE028FB8693D90550BE9AB9F" captionStart="Figures 4954" captionStartId="F12" captionText="Figures 4954. Suragina Walker spp. wing dorsal view: 49 S. monogramma (Bezzi) ♀ (BMSA (D) 129988) 50 S. mulanjeensis Muller, sp. nov. ♀ (BMSA (D) 92374) 51 S. nigromaculata Muller, sp. nov. ♂ holotype (NHMUK 014064156) 52 S. pauliani ♂ paratype (NMSA - DIP 028172) 53 S. semiobscura Muller, sp. nov. ♀ paratype (ICIPE) 54 S. zombaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158451). 51 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures49-54" httpUri="https://binary.pensoft.net/fig/1205874">53</figureCitation>
); postpronotal lobe pale yellow; fore femur yellow on at least basal half to two-thirds, remaining apical third to half dark brown
</td>
<td id="61A658C8EBB8D300AC8F7BA6E816BC9F" colspan="1" rowspan="1">
<emphasis id="F99DFAFC417BEEABACA20005982A1C2C" bold="true">
<taxonomicName id="B2C1E39B46F3C7A19483ACA450D5A442" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="semiobscura" status="sp. nov.">
<emphasis id="CF06EDD24A48C7B2B9E02F916D1121CD" bold="true" italics="true">S. semiobscura</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="CBB65E96E35B7F0A1E9F8612BB79DD85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
</tr>
<tr id="390348A1044D9A051383AE70F040D896">
<td id="45FDB7D43AA800C5121F0E753782AB81" colspan="1" rowspan="1"></td>
<td id="266E10B200B93156E80279CD9071C12F" colspan="1" rowspan="1">
Wing with brownish suffusion uniformly distributed over anterior and posterior half (Fig.
<figureCitation id="EF88BE801728FEE170A4B51E0F51F43B" captionStart="Figures 3742" captionStartId="F10" captionText="Figures 3742. Suragina Walker spp. wing dorsal view: 37 S. agramma (Bezzi) ♀ (BMSA (D) 124627) 38 S. bezzii (Curran) ♂ (SAM-DIP-A 018410) 39 S. bilobata Muller, sp. nov. ♀ paratype (CSCA) 40 S. copelandi Muller, sp. nov. ♀ paratype (BMSA (D) 83423); S. binominata (Bequaert): 41 holotype ♂ (MLUH) 42 ♀ (NMSA - DIP 158446). 41 Copyright ZNS Halle, January 30, 2024. Abbreviations: br basal radial cell; bm basal medial cell; cua anterior cubital cell; CuA + CuP anterior branch of cubital vein + posterior branch of cubital vein; d discal cell; M 1 first branch of media; m 1 first medial cell; M 2 second branch of media; m 2 second medial cell; M 3 third branch of media; m 3 third medial cell; R 1 anterior branch of radius; r 1 first radial cell; R 2 + 3 second branch of radius; r 2 + 3 second radial cell; R 4 upper branch of third branch of radius; R 5 lower branch of third branch of radius; Sc subcostal vein; sc subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures37-42" httpUri="https://binary.pensoft.net/fig/1205872">38</figureCitation>
); postpronotal lobe dark brown, at most with a slight orange-yellow colouration on lateral margin; fore femur dark brown on basal third or almost entirely yellow with only extreme apex and base brown
</td>
<td id="29F854B43940A427D864E24874F1792D" colspan="1" rowspan="1">
<taxonomicName id="DAB570CB68084365FC9A3CBB67BEE47F" authority="(Curran)" baseAuthorityName="Curran" baseAuthorityYear="1928" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="5B274A222354627C58B08C217872437A" bold="true">
<emphasis id="B353152114E302E055AA41F37A89EEF6" bold="true" italics="true">S. bezzii</emphasis>
(Curran)
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="DF9EF571C91712521DDEAA1454FC3BEE">
<td id="64BE7E14D042BC3D8C229DE113E70105" colspan="1" rowspan="1">14</td>
<td id="18DE1C28C4E0DFB73957B546A809372B" colspan="1" rowspan="1">All coxae orange-yellow; hind femur orange-yellow on majority of surface, with small anterior apical dark mark, hind tibia and tarsi orange-yellow; occiput with pair of narrow dark subtriangular markings; ♂ cercus orange-yellow (♀ unknown)</td>
<td id="481C0D718419604C982C5CB2CEA2937D" colspan="1" rowspan="1">
<taxonomicName id="02E9ADE24874DCCAACAF2913CEC22880" authority="(Brunetti)" baseAuthorityName="Brunetti" baseAuthorityYear="1929" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="nigromaculata">
<emphasis id="D4A33F2E011BCF3E4A9F868F24987988" bold="true">
<emphasis id="206AE00531E58A14458A61633DE327C9" bold="true" italics="true">S. nigromaculata</emphasis>
(Brunetti)
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="BDF600E11702940A12C29DF112E8CCEA">
<td id="BF71FAC39CCF63FA655170C871B0E6D4" colspan="1" rowspan="1"></td>
<td id="F68F029AFE0929F84843C7FB1642C591" colspan="1" rowspan="1">Fore coxa variable, mid and hind coxae brown with some greyish pruinosity on at least anterior surface; hind femur with middle third with some degree of light infuscation to being blackish-brown, hind tibia and tarsi with some degree of overall infuscation; ♂ ♀ cercus darkened</td>
<td id="4FEA0B6747C456F70B398F4FA934DC93" colspan="1" rowspan="1">
<emphasis id="68DAFDB3E53906B4104F584820444FB4" bold="true">15</emphasis>
</td>
</tr>
<tr id="B5909F3AC0C3A7AABD4D1AD2A0029F90">
<td id="2A36BC0BD8EEE541BA8555B44056BE0B" colspan="1" rowspan="1">15</td>
<td id="9B83F3E7A9065318021000B6BDE2A6FF" colspan="1" rowspan="1">
Antenna with 1
<superScript id="E0889B0DD4C8F94842FD4EEC56B87DF7">st</superScript>
flagellomere brown (Figs
<figureCitation id="C9273A923AF419305AF3338A40A9318D" captionStart="Figures 3133" captionStartId="F8" captionText="Figures 3133. Suragina Walker spp. dorsal habitus: 31 S. nigromaculata ♂ holotype (NHMUK 014064156); lateral habitus, S. pauliani Stuckenberg: 32 ♂ paratype (NMSA - DIP 28172) 33 ♀ paratype (NMSA - DIP 028174). 31 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures31-33" httpUri="https://binary.pensoft.net/fig/1205870">32</figureCitation>
,
<figureCitation id="B3DE1717BAB913096D21529FC9F240C7" captionStart="Figures 3133" captionStartId="F8" captionText="Figures 3133. Suragina Walker spp. dorsal habitus: 31 S. nigromaculata ♂ holotype (NHMUK 014064156); lateral habitus, S. pauliani Stuckenberg: 32 ♂ paratype (NMSA - DIP 28172) 33 ♀ paratype (NMSA - DIP 028174). 31 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures31-33" httpUri="https://binary.pensoft.net/fig/1205870">33</figureCitation>
); postpronotal lobe and postalar callus pale orange-yellow, without pruinosity; at least tergites 15 with dark subtriangular medial markings; wing overall light brown suffused (Fig.
<figureCitation id="6A63619F233CCBA634159126A6A07641" captionStart="Figures 4954" captionStartId="F12" captionText="Figures 4954. Suragina Walker spp. wing dorsal view: 49 S. monogramma (Bezzi) ♀ (BMSA (D) 129988) 50 S. mulanjeensis Muller, sp. nov. ♀ (BMSA (D) 92374) 51 S. nigromaculata Muller, sp. nov. ♂ holotype (NHMUK 014064156) 52 S. pauliani ♂ paratype (NMSA - DIP 028172) 53 S. semiobscura Muller, sp. nov. ♀ paratype (ICIPE) 54 S. zombaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158451). 51 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures49-54" httpUri="https://binary.pensoft.net/fig/1205874">52</figureCitation>
) apparent darker suffused substigmal mark in ♂; Madagascan endemic
</td>
<td id="130CAACB7F64BA7B6FC4F80C4DCCCCEA" colspan="1" rowspan="1">
<taxonomicName id="61BFDB43E5C4D1EDE954D7BDA0118D21" authority="(Stuckenberg)" baseAuthorityName="Stuckenberg" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="9BD0BDDF5A71D00E5779DE43302B8798" bold="true">
<emphasis id="D15E51962DAEEF4106D0F8D3E05591C2" bold="true" italics="true">S. pauliani</emphasis>
(Stuckenberg)
</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="D9CC2A9B60C8E32619E3734DED570EDE">
<td id="8FD8822A1B8DD8911D84289FF50FEEEC" colspan="1" rowspan="1"></td>
<td id="5A5F457F106AAE3EC056483C1237ECA8" colspan="1" rowspan="1">
Antenna with 1
<superScript id="8C53B943B49168320801D02B6CE664B4">st</superScript>
flagellomere orange-yellow; postpronotal lobe orange-brown to brown with greyish pruinosity, or dark brown without pruinosity, postalar callus colour variable; abdominal tergite markings variable, if dark, subtriangular then wing without apparent dark substigmal mark in ♂ ♀; mainland Africa
</td>
<td id="B2E54A43EE85FC7AB1D199441748848D" colspan="1" rowspan="1">
<emphasis id="AFED37AF3034DB8A708A9A144175BEA0" bold="true">16</emphasis>
</td>
</tr>
<tr id="CC4B235BA10DDB4165667FC55839111C">
<td id="61BCB550029F4671EF9F7D9B9C79E0F0" colspan="1" rowspan="1">16</td>
<td id="7CE298C8B7F7771D85C1265254917808" colspan="1" rowspan="1">
Hind femur with median third only slightly infuscated, if at all; wing with light brown stigma, wing membrane lightly and evenly brown suffused (Figs
<figureCitation id="D760D5EAFD3EAFD17075BA0B0128B660" captionStart="Figures 3742" captionStartId="F10" captionText="Figures 3742. Suragina Walker spp. wing dorsal view: 37 S. agramma (Bezzi) ♀ (BMSA (D) 124627) 38 S. bezzii (Curran) ♂ (SAM-DIP-A 018410) 39 S. bilobata Muller, sp. nov. ♀ paratype (CSCA) 40 S. copelandi Muller, sp. nov. ♀ paratype (BMSA (D) 83423); S. binominata (Bequaert): 41 holotype ♂ (MLUH) 42 ♀ (NMSA - DIP 158446). 41 Copyright ZNS Halle, January 30, 2024. Abbreviations: br basal radial cell; bm basal medial cell; cua anterior cubital cell; CuA + CuP anterior branch of cubital vein + posterior branch of cubital vein; d discal cell; M 1 first branch of media; m 1 first medial cell; M 2 second branch of media; m 2 second medial cell; M 3 third branch of media; m 3 third medial cell; R 1 anterior branch of radius; r 1 first radial cell; R 2 + 3 second branch of radius; r 2 + 3 second radial cell; R 4 upper branch of third branch of radius; R 5 lower branch of third branch of radius; Sc subcostal vein; sc subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures37-42" httpUri="https://binary.pensoft.net/fig/1205872">41</figureCitation>
,
<figureCitation id="2BFB3B25C58ED1C4C040EF00AA579522" captionStart="Figures 3742" captionStartId="F10" captionText="Figures 3742. Suragina Walker spp. wing dorsal view: 37 S. agramma (Bezzi) ♀ (BMSA (D) 124627) 38 S. bezzii (Curran) ♂ (SAM-DIP-A 018410) 39 S. bilobata Muller, sp. nov. ♀ paratype (CSCA) 40 S. copelandi Muller, sp. nov. ♀ paratype (BMSA (D) 83423); S. binominata (Bequaert): 41 holotype ♂ (MLUH) 42 ♀ (NMSA - DIP 158446). 41 Copyright ZNS Halle, January 30, 2024. Abbreviations: br basal radial cell; bm basal medial cell; cua anterior cubital cell; CuA + CuP anterior branch of cubital vein + posterior branch of cubital vein; d discal cell; M 1 first branch of media; m 1 first medial cell; M 2 second branch of media; m 2 second medial cell; M 3 third branch of media; m 3 third medial cell; R 1 anterior branch of radius; r 1 first radial cell; R 2 + 3 second branch of radius; r 2 + 3 second radial cell; R 4 upper branch of third branch of radius; R 5 lower branch of third branch of radius; Sc subcostal vein; sc subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures37-42" httpUri="https://binary.pensoft.net/fig/1205872">42</figureCitation>
); abdominal tergites orange-yellow with blackish subtriangular median markings and similarly coloured lateral markings
</td>
<td id="77BF30DB50EA6DE77D4D8F37C2D5A151" colspan="1" rowspan="1">
<taxonomicName id="E20FBF8FCA33B3FE76EF5150AFBA45E9" authority="(Bequaert)" authorityName="" baseAuthorityName="Bequaert" baseAuthorityYear="1921" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="binominata">
<emphasis id="6559D64992D4F94E81A39AFB0EAD2D30" bold="true">
<emphasis id="008D3A5942A8B8F74BF299FB75AE0233" bold="true" italics="true">S. binominata</emphasis>
(Bequaert)
</emphasis>
</taxonomicName>
(in part, ♂)
</td>
</tr>
<tr id="C97A481C432D5FEBAB20AD6094B7F46D">
<td id="41E81476B38E13797C6F123E38D06BBE" colspan="1" rowspan="1"></td>
<td id="C8B65036A69A7C65BC65A0E5852F304F" colspan="1" rowspan="1">Hind femur with median third blackish; wing with dark brown stigma, wing membrane darker brown suffused on apical half, with some cells having a small hyaline patch at their centres; abdominal tergite colour variable, but never with subtriangular markings</td>
<td id="BF86BCF5C4CE8018147FE8D33220435A" colspan="1" rowspan="1">
<emphasis id="372A5B81AB732C996B5A7A3FD3ACF957" bold="true">17</emphasis>
</td>
</tr>
<tr id="BAE7BF8379C9E78D33BCA718086A3C06">
<td id="9B6629A85D6ED9A4BFB190879B51614D" colspan="1" rowspan="1">17</td>
<td id="9453A360F4E515EABBEF3295B78F799F" colspan="1" rowspan="1">
Upper occiput with narrow subrectangular markings not reaching down past upper third of occiput (Fig.
<figureCitation id="543B9CE7BEE773D709A79BCA130C356E" captionStart="Figures 3436" captionStartId="F9" captionText="Figures 3436. Suragina Walker spp. lateral habitus: 34 S. semiobscura Muller, sp. nov. ♀ holotype (SAM-DIP-A 018385); S. zombaensis Muller, sp. nov.: 35 ♂ paratype (NMSA - DIP 158421) 36 ♀ paratype (NMSA - DIP 158415). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures34-36" httpUri="https://binary.pensoft.net/fig/1205871">36</figureCitation>
); tergites 15 each with a median longitudinal blackish marking flanked for most part by bluish-grey pruinosity, additionally tergites 35 each with a grey pruinose posterior band
</td>
<td id="2D5231456F710064EF0CB45A5613E46F" colspan="1" rowspan="1">
<emphasis id="B05767C4F5A51EA150283D55E47D0BBE" bold="true">
<taxonomicName id="BEB3A1EC612D347A3D70E7FE815EEF7E" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="zombaensis" status="sp. nov.">
<emphasis id="6AC223E0459E711C60BA35185547C8CD" bold="true" italics="true">S. zombaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="1B9B0DA02B3672965A3CA2EE05D9E1AD" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
</tr>
<tr id="0890E5BD4F39612D70834F577A17627E">
<td id="F3A54BAE6C2B8472DAFDF9138E980C4C" colspan="1" rowspan="1"></td>
<td id="D1D29C51C6AA81C75C868BA1AE791238" colspan="1" rowspan="1">
Upper occiput with wide subrectangular markings running down to occipital foramen; tergite 1 appearing orange-yellow without any pruinosity, tergites 2 and 3 with a light brown dorsal marking appearing to cover most of dorsal surfaces; tergites 4 and 5 orange-yellow, with tergites 6 and 7 appearing to have brownish posterior margins, all tergites without apparent pruinosity (Fig.
<figureCitation id="48DA9D0E88AB33EEE00BC15CB3B7012D" captionStart="Figures 18, 19" captionStartId="F4" captionText="Figures 18, 19. Suragina Walker spp. dorsal habitus: S. dimidiatipennis (Brunetti): 18 ♀ holotype (NHMUK 014064158); S. falsa Oldroyd: 19 ♀ holotype (NHMUK 014064160). 18, 19 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures18-19" httpUri="https://binary.pensoft.net/fig/1205866">18</figureCitation>
)
</td>
<td id="362667C059880DF6A6820BF73BC12CE3" colspan="1" rowspan="1">
<taxonomicName id="F564505B1598C6EAD4FDDDB4FC22F541" authority="(Brunetti)" baseAuthorityName="Brunetti" baseAuthorityYear="1929" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dimidiatipennis">
<emphasis id="6D6E0AD762BCFF313E7D4402DEAF004F" bold="true">
<emphasis id="439F64232364D405067DB980EF31EC73" bold="true" italics="true">S. dimidiatipennis</emphasis>
(Brunetti)
</emphasis>
</taxonomicName>
</td>
</tr>
</table-wrap>
</tbody>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,782 @@
<document id="C3D65165E610742BDCBF67D364C9B06F" ID-DOI="10.3897/afrinvertebr.65.140524" ID-ZooBank="BCB49D2E-F772-49EB-A17A-47EB21194212" ID-publisher-id="140524" URI-arpha="8BBD979A-A43B-550B-853F-36895E8AFD3A" URI-zoobank="http://zoobank.org/BCB49D2E-F772-49EB-A17A-47EB21194212" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1734967240283" checkinUser="pensoft" docAuthor="Muller, Burgert S., Swart, Vaughn R. &amp; Snyman, Louwrens P." docDate="2024" docId="46AA75DDC8A55CE38FAEE183DFDC2814" docLanguage="en" docName="African Invertebrates 65(2): 247-327" docOrigin="African Invertebrates 65 (2)" docSource="https://africaninvertebrates.pensoft.net/article/140524/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Suragina zombaensis Muller 2024, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="327" masterDocId="8BBD979AA43B550B853F36895E8AFD3A" masterDocTitle="Revision of Afrotropical Suragina Walker, 1859 (Diptera, Athericidae)" masterLastPageNumber="327" masterPageNumber="247" pageNumber="247" updateTime="1735007760347" updateUser="admin">
<mods:mods id="C2CB217EB2003C38F9E58C0201FE664A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7A9EECF8F03508DF75B7C268CDE546F9">
<mods:title id="11E22BABFECD47DDCBF7C500B99636BC">Revision of Afrotropical Suragina Walker, 1859 (Diptera, Athericidae)</mods:title>
</mods:titleInfo>
<mods:name id="0FE7F602F8AFB0D20A2BD836B04A1503" type="personal">
<mods:role id="4ABF6E94E5C8463553F5DC191DEBAAC5">
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<mods:namePart id="AF66B0D5A8FED65C95D8CC32EF356375">Muller, Burgert S.</mods:namePart>
<mods:nameIdentifier id="CE3CA7AECBAABB5B6831D4C812C2675C" type="ORCID">0000-0002-7304-4050</mods:nameIdentifier>
<mods:affiliation id="175C5698817F84BD597C53732F6FC676">Department Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, 9301, South Africa &amp; Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
</mods:name>
<mods:name id="D928FEFDC75797431059440D3C8943BE" type="personal">
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</mods:role>
<mods:namePart id="EC58F4D658C783F19D99A5F7F99FD9DB">Swart, Vaughn R.</mods:namePart>
<mods:nameIdentifier id="AE1A1727B5C9DC55397D5D903CABFFB8" type="ORCID">0000-0001-7905-5298</mods:nameIdentifier>
<mods:affiliation id="464A5103FD1EE76BE340F9B28A49200D">Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
</mods:name>
<mods:name id="8D77CB431C4EFD31C3E5B7F02C3F1A5E" type="personal">
<mods:role id="1B4ED3F3F6907A2B727D0E8AB2DDC223">
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</mods:role>
<mods:namePart id="2C8EDAC5B1AEFFA5F31903E20CCD9C27">Snyman, Louwrens P.</mods:namePart>
<mods:nameIdentifier id="64293A11F16B6703D6FC34F3CD55D53B" type="ORCID">0000-0002-5768-7216</mods:nameIdentifier>
<mods:affiliation id="F273FBBAF81CD3D451372F75CE0148A4">Invertebrate Zoology, Royal Alberta Museum, 9810 103 A Avenue NW, Edmonton, Alberta, T 5 J 0 G 2, Canada &amp; Department of Veterinary Microbiology, University of Saskatchewan, 52 Campus Drive, Saskatoon, Saskatchewan, S 7 N 5 B 4, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource id="35F2BDEE658D9A6B23F51C6980FB6B23">text</mods:typeOfResource>
<mods:relatedItem id="FFB1221EB0A646FD62FD71277A3F01E8" type="host">
<mods:titleInfo id="1E99C9CF48537DF3059F925590449BB8">
<mods:title id="6C47ABE010FDB44692896FD5FC769757">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="20206E12DFAF19BC03377808EAAC94BE">
<mods:date id="518D5DD9F212B0252525E41AE8EFA3D7">2024</mods:date>
<mods:detail id="FE963ABB48D013EDAF0C2A3DB34760A9" type="pubDate">
<mods:number id="A04543D3381BEBA0F8BD55FFE95D724D">2024-12-23</mods:number>
</mods:detail>
<mods:detail id="528FE13278574D3BC472222356238F01" type="volume">
<mods:number id="D9ADD13A7F35A960D09CF2A893D83D32">65</mods:number>
</mods:detail>
<mods:detail id="5B17F92DEFAEF5CCF98938E501120661" type="issue">
<mods:number id="7499D7847DB3E19F8A715EEE2DB2B632">2</mods:number>
</mods:detail>
<mods:extent id="782A1DD4B886D766FDBA129D20D68D04" unit="page">
<mods:start id="F788274ECD0871E0D48CABF51B3D02EC">247</mods:start>
<mods:end id="76B3D81A7776D1F95DFC75C3C3B2D4EC">327</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="10CDF12C0A5CF57F3E6DD2ABFD699BBD">journal article</mods:classification>
<mods:identifier id="170D0B43A6BFC77647E7940902631895" type="DOI">10.3897/afrinvertebr.65.140524</mods:identifier>
<mods:identifier id="35AC3B307FBF241CBC8E5B49BE4EB9A3" type="ZooBank">BCB49D2E-F772-49EB-A17A-47EB21194212</mods:identifier>
</mods:mods>
<treatment id="46AA75DDC8A55CE38FAEE183DFDC2814" ID-arpha="46AA75DD-C8A5-5CE3-8FAE-E183DFDC2814" ID-zoobank="https://zoobank.org/80FD6CC3-1332-469B-8FB6-0C45CD0E06A5" LSID="urn:lsid:plazi:treatment:46AA75DDC8A55CE38FAEE183DFDC2814" LSID-ZBK="urn:lsid:zoobank.org:act:80FD6CC3-1332-469B-8FB6-0C45CD0E06A5" httpUri="http://treatment.plazi.org/id/46AA75DDC8A55CE38FAEE183DFDC2814">
<subSubSection id="A3A38C630A6D2C4D6A68E3C8B66ED33E" type="nomenclature">
<paragraph id="F24E262147C78E8AE46597C8E19C757B">
<taxonomicName id="D045582B999A756D33BFF36E29B71B34" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="zombaensis" status="sp. nov.">
<emphasis id="C0298756B0E53AA96A72A014E9FCCCB1" italics="true">Suragina zombaensis</emphasis>
Muller
</taxonomicName>
<taxonomicNameLabel id="95C405A1EA5CC6A95E054A2CA1CEDC9D" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="567E397B51E40AA8693031E7546FA769" type="description">
<paragraph id="5BFA590049D5F5FADB6B3FC627F14E97">
<figureCitation id="DF0146A520BDEAA47BE4A2F812C00E03" captionStart="Figures 3436" captionStartId="F9" captionText="Figures 3436. Suragina Walker spp. lateral habitus: 34 S. semiobscura Muller, sp. nov. ♀ holotype (SAM-DIP-A 018385); S. zombaensis Muller, sp. nov.: 35 ♂ paratype (NMSA - DIP 158421) 36 ♀ paratype (NMSA - DIP 158415). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures34-36" httpUri="https://binary.pensoft.net/fig/1205871">Figs 35</figureCitation>
,
<figureCitation id="334A2AFE660B7142BC52727F524CD7ED" captionStart="Figures 3436" captionStartId="F9" captionText="Figures 3436. Suragina Walker spp. lateral habitus: 34 S. semiobscura Muller, sp. nov. ♀ holotype (SAM-DIP-A 018385); S. zombaensis Muller, sp. nov.: 35 ♂ paratype (NMSA - DIP 158421) 36 ♀ paratype (NMSA - DIP 158415). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures34-36" httpUri="https://binary.pensoft.net/fig/1205871">36</figureCitation>
,
<figureCitation id="791E00F4DB6CC75B82888F4A6DA58C7B" captionStart="Figures 4954" captionStartId="F12" captionText="Figures 4954. Suragina Walker spp. wing dorsal view: 49 S. monogramma (Bezzi) ♀ (BMSA (D) 129988) 50 S. mulanjeensis Muller, sp. nov. ♀ (BMSA (D) 92374) 51 S. nigromaculata Muller, sp. nov. ♂ holotype (NHMUK 014064156) 52 S. pauliani ♂ paratype (NMSA - DIP 028172) 53 S. semiobscura Muller, sp. nov. ♀ paratype (ICIPE) 54 S. zombaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158451). 51 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures49-54" httpUri="https://binary.pensoft.net/fig/1205874">54</figureCitation>
,
<figureCitation id="AAEC4C1A50CDE8D79C87657BA7423B5F" captionStart="Figures 7174" captionStartId="F15" captionText="Figures 7174. Suragina Walker spp. male terminalia, ventral view (71, 72) and epandrium with cerci, dorsal view (73, 74): 71, 73 S. mulanjeensis Muller, sp. nov. paratype (NMSA - DIP 162049) 72, 74 S. zombaensis Muller, sp. nov. paratype (NMSA - DIP 158450). Scale bar (applicable to all illustrations): 0.6 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures71-74" httpUri="https://binary.pensoft.net/fig/1205877">72</figureCitation>
,
<figureCitation id="4704D6C01F5542D40D5CB1C83432DDC5" captionStart="Figures 7174" captionStartId="F15" captionText="Figures 7174. Suragina Walker spp. male terminalia, ventral view (71, 72) and epandrium with cerci, dorsal view (73, 74): 71, 73 S. mulanjeensis Muller, sp. nov. paratype (NMSA - DIP 162049) 72, 74 S. zombaensis Muller, sp. nov. paratype (NMSA - DIP 158450). Scale bar (applicable to all illustrations): 0.6 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures71-74" httpUri="https://binary.pensoft.net/fig/1205877">74</figureCitation>
,
<figureCitation id="622B5C43EAEEE46BD4ACAFC80B76179A" captionStart="Figures 7587" captionStartId="F16" captionText="Figures 7587. Suragina Walker spp. female genital fork and spermathecae, ventral view: 75 S. agramma (Bezzi) (ICIPE) 76 S. bezzii (Curran) (SAM-DIP- 018403) 77 S. bilobata Muller, sp. nov. (CSCA) 78 S. binominata (Bequaert) (NMSA - DIP 158411) 79 S. copelandi Muller, sp. nov. paratype (BMSA (D) 84686) 80 S. freidbergi Muller, sp. nov. paratype (BMSA (D) 92158) 81 S. liberiaensis Muller, sp. nov. paratype (NMSA - DIP 158442) 82 S. malavaensis Muller, sp. nov. paratype (ICIPE) 83 S. milloti (Séguy) (CSCA) 84 S. monogramma (Bezzi) (BMSA (D) 129988) 85 S. mulanjeensis Muller, sp. nov. paratype (NMSA - DIP 158392) 86 S. semiobscura Muller, sp. nov. paratype (ICIPE) 87 S. zombaensis Muller, sp. nov. paratype (NMSA - DIP 1584719). Scale bar (applicable to all illustrations): 0.1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures75-87" httpUri="https://binary.pensoft.net/fig/1205878">87</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E65CK" type="type material">
<paragraph id="FF121AAE195E63E7ECCEC1BB83E9189F">
<heading id="C0FF12160C63FDAE5821469A577E510F" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="B2A495DA9EC9094A831680EE0E16A02B">
<materialsCitation id="C2D2E868B0244EEE2E9C59185914894D" collectingDate="1980-11-24" collectingDateMax="1980-11-27" collectingDateMin="1980-11-24" collectionCode="NMSA" collectorName="J. G. H. Londt &amp; B. R. Stuckenberg" country="Malawi" latitude="-15.383333" location="Ad" longLatPrecision="13" longitude="35.333332" specimenCode="DIP 158422" specimenCount="1" specimenCount-male="1" stateProvince="Zomba" typeStatus="holotype">
<emphasis id="BCEEF777D5318C246568EC2D53CA1234" bold="true" italics="true">
<typeStatus id="7CB2D98A01B94F155385398EF6CCF38E" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="C23EA556FD7088982B02B6932ACC6B2F" name="Malawi">Malawi</collectingCountry>
<specimenCount id="DBB0126EE354DF9EFA22FD6D6FA115BF" count="1" type="male">1 ♂</specimenCount>
;
<collectingRegion id="A3B91D6D8C1B80E01C78A4FD8CA40A64" country="Malawi" name="Zomba">Zomba</collectingRegion>
; 1535
<location id="CA23F19ADB5F8EB88704AA352BA83200" LSID="urn:lsid:plazi:treatment:46AA75DDC8A55CE38FAEE183DFDC2814:CA23F19ADB5F8EB88704AA352BA83200" country="Malawi" latitude="-15.383333" longLatPrecision="13" longitude="35.333332" name="Ad" stateProvince="Zomba">Ad</location>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EN6CK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[35.333333,-15.383333]}">
<geoCoordinate id="648B6771324AA355A77C0D1FF27BD4DE" degrees="15" direction="south" minutes="23.00" orientation="latitude" precision="9" value="-15.383333">15°23.00'S</geoCoordinate>
,
<geoCoordinate id="D66F2E28B107AEEF4400C1713C30115F" degrees="35" direction="east" minutes="20.00" orientation="longitude" precision="9" value="35.333332">35°20.00'E</geoCoordinate>
</named-content>
];
<date id="7DA9F3B446702417B7E3B47EB86E31FE" value="1980-11-24" valueMax="1980-11-27" valueMin="1980-11-24">
<collectingDate id="A99FFFDF9476695860B889711CDB3606" value="1980-11-24" valueMax="1980-11-27" valueMin="1980-11-24">2427 Nov. 1980</collectingDate>
</date>
;
<collectorName id="4F9F8C538507618EC0C15E78D12AE384">J. G. H. Londt</collectorName>
&amp;
<collectorName id="D7A9E44B39B1C907BA18B414B6578432">B. R. Stuckenberg</collectorName>
leg.; (
<named-content id="C8A7AB63D41BBF7DD9FBFB6335456D26" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="AD31AF76F6ABBAE0F6DFE596A5495D17" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
-
<specimenCode id="2CEDABFE3F0F8E100AE2DB24FD1686DC" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">DIP 158422</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="7CFE88E996FEAE342CF6B82F77E8A0CA">
<materialsCitation id="C19B9FC8E20ED4C0682D06C01DB7B583" collectingDate="1980-11-24" collectingDateMax="1980-11-27" collectingDateMin="1980-11-24" collectionCode="NMSA" collectorName="J. G. H. Londt &amp; B. R. Stuckenberg" country="Malawi" latitude="-15.383333" location="Ad" longLatPrecision="13" longitude="35.333332" specimenCode="DIP 158420, 158421, 158450, DIP 158417, 158418, 158419, 158451" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Zomba" typeStatus="paratype">
<emphasis id="22884D8E99AE1BC11EFAE3C2D42CCCBB" bold="true" italics="true">
<typeStatus id="AD8EDAD6AA6A05B87E9A8EB3E25589A5" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="8F275F14A607D946E8441012399467A5" count="3" type="male">3 ♂</specimenCount>
<specimenCount id="617D38B4B45705C990CF413CCAC221F6" count="4" type="female">4 ♀</specimenCount>
; Same data as holotype; (
<specimenCount id="1B87ADE712F79C2FE9B4350B8EE0E33F" count="1" type="male"></specimenCount>
:
<named-content id="6211D922A57A3FF477D7DA6CC2B717E5" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="F3FB896CBB5007B09E61AF63C51E6FFF" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
-
<specimenCode id="2369ACB8E5845B6A14DAD13E4C64A097" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">DIP 158420</specimenCode>
,
<specimenCode id="48863ACB0294C14039CC64C669151FF3" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">158421</specimenCode>
,
<specimenCode id="4E4B6EF685542982C15C1B365F388627" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">158450</specimenCode>
;
<specimenCount id="C1C0DB499A432BCCF3F7BD0DB8CBE2D5" count="1" type="female"></specimenCount>
:
<named-content id="AE65C5B13BE9B5BB3484B63BFC3CDF7A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="63132D56FF3FA3EE6BCE649BB57DA28E" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
-
<specimenCode id="E83E80E5ACE7EBCCC26A968051953B71" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">DIP 158417</specimenCode>
,
<specimenCode id="2032B319BE82A1C585076D98D7BFB341" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">158418</specimenCode>
,
<specimenCode id="BF2200E8527EDF41057D1B20DA3F5FEC" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">158419</specimenCode>
,
<specimenCode id="171322EE803BD8487B21DC01F7B5B001" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">158451</specimenCode>
)
</materialsCitation>
<materialsCitation id="20771BD514503797F6A766FC3BE79945" collectingDate="1980-12-03" collectingDateMax="1980-12-04" collectingDateMin="1980-12-03" collectionCode="NMSA" collectorName="J. G. H. Londt &amp; B. R. Stuckenberg" country="Malawi" county="Forest reserve" elevation="1500" latitude="-13.366667" location="Montane forest" longLatPrecision="13" longitude="34.0" municipality="Ac" specimenCode="DIP 158415" specimenCount="1" specimenCount-female="1" stateProvince="Ntchisi" typeStatus="paratype">
<specimenCount id="398AB4F8FA528027A3379EFE67BD59A9" count="1" type="female">1 ♀</specimenCount>
;
<collectingRegion id="1C8DA1B02CB58F3F56C6E2B43DB63D47" country="Malawi" name="Ntchisi">Ntchisi</collectingRegion>
<collectingCounty id="06BA1B51061C8199DE14DDA91FC5A2E1">Forest reserve</collectingCounty>
; 1334
<collectingMunicipality id="A4870902007E04C346BF879397E5A710">Ac</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIADK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[34.000000,-13.366667]}">
<geoCoordinate id="25A186DB12C9043B96C50D96C89C766F" degrees="13" direction="south" minutes="22.00" orientation="latitude" precision="9" value="-13.366667">13°22.00'S</geoCoordinate>
,
<geoCoordinate id="652FDFEF3C5D0765EC4C0D4FCE0638F1" degrees="34" direction="east" minutes="00.00" orientation="longitude" precision="9" value="34.0">34°00.00'E</geoCoordinate>
</named-content>
];
<quantity id="52DBBBA91B1242E175473C0F632AA3DD" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
<elevation id="88D65B999404104D746B4C5CBC5F5E89" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 masl</elevation>
</quantity>
;
<date id="79B2FB7589C39CE67E641145EF40820D" value="1980-12-03" valueMax="1980-12-04" valueMin="1980-12-03">
<collectingDate id="F0DAB204330C7389B4035D33E206B78A" value="1980-12-03" valueMax="1980-12-04" valueMin="1980-12-03">34 Dec. 1980</collectingDate>
</date>
;
<collectorName id="B5705C868B2E8951FCA2DE2486F51F58">J. G. H. Londt</collectorName>
&amp;
<collectorName id="DCD46B4B490ADE659E283D9FF5F6A38A">B. R. Stuckenberg</collectorName>
leg.;
<location id="528AFA121493BE28A994409B0F8937B9" LSID="urn:lsid:plazi:treatment:46AA75DDC8A55CE38FAEE183DFDC2814:528AFA121493BE28A994409B0F8937B9" country="Malawi" county="Forest reserve" latitude="-13.366667" longLatPrecision="13" longitude="34.0" municipality="Ac" name="Montane forest" stateProvince="Ntchisi">Montane forest</location>
, woodland; (
<named-content id="FE4475230A4EAC14CABBB2EDE93C2C57" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="01C3772C8A7F920A0F038253548D68AC" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
-
<specimenCode id="8AE5E94AA09925E045DAAE420DCE82AD" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">DIP 158415</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8948F879D9A433DB793D81EFCD713E02">
<typeStatus id="6F0F2A5CD669E8AEF23C4DFBB6C5AD5A" type="holotype">Holotype</typeStatus>
and
<typeStatus id="8838624382CAAB85625D531FF773C4CB" type="paratype">paratypes</typeStatus>
deposited in
<named-content id="30B80ACE4E57A37627AEEC321651E259" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">
<collectionCode id="4F9DA7139C714EB4A29C779E388FC822" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">NMSA</collectionCode>
</named-content>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXADK" type="diagnosis">
<paragraph id="407C4D0AF125C32D5E452E546EEFF4A0">
<heading id="7D3E5E46EFE5B4E243779FFB0B0EDF4F" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="7070DB6BB91EE454A0CC6009B1039620">
<taxonomicName id="EA69E7A521AA609073D8CC822E7003AE" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="zombaensis" status="sp. nov.">
<emphasis id="3AC1164635CB9E9F3B8DEC110794D0C2" italics="true">Suragina zombaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="16270EA3721142AA3DCAAA24087B3A83" rank="species">sp. nov.</taxonomicNameLabel>
has its thorax almost entirely brown with bluish-grey pruionosity, the scutellum with apical margin yellow. The males and females are similarly coloured (Figs
<figureCitation id="0A439626E89FBE76ED0A4A8D122D7AB8" captionStart="Figures 3436" captionStartId="F9" captionText="Figures 3436. Suragina Walker spp. lateral habitus: 34 S. semiobscura Muller, sp. nov. ♀ holotype (SAM-DIP-A 018385); S. zombaensis Muller, sp. nov.: 35 ♂ paratype (NMSA - DIP 158421) 36 ♀ paratype (NMSA - DIP 158415). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures34-36" httpUri="https://binary.pensoft.net/fig/1205871">35</figureCitation>
,
<figureCitation id="DE1640E1BA6BEEA3F7E1AF3D96979A8C" captionStart="Figures 3436" captionStartId="F9" captionText="Figures 3436. Suragina Walker spp. lateral habitus: 34 S. semiobscura Muller, sp. nov. ♀ holotype (SAM-DIP-A 018385); S. zombaensis Muller, sp. nov.: 35 ♂ paratype (NMSA - DIP 158421) 36 ♀ paratype (NMSA - DIP 158415). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures34-36" httpUri="https://binary.pensoft.net/fig/1205871">36</figureCitation>
), except for the males having the lateral margins of tergites 3 and 4 yellow compared to dark brown in the female. Tergite
<quantity id="0CDBF03A1F2C9F3143BA59CC94398856" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
both sexes has dark vitta medially, flanked by brownish-yellow colouration, giving it a fenestrated appearance. The wing of
<taxonomicName id="F4D55E8735C66C09B53B1BCD381D0FB4" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="zombaensis" status="sp. nov.">
<emphasis id="39CD32F88B34D5B364D00A3EF75AFE17" italics="true">S. zombaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="011BC7865CB35F5F01F59410127895CE" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to that of
<taxonomicName id="CF0FF7F9910E68C58E64D799EFFD46A1" baseAuthorityName="Brunetti" baseAuthorityYear="1929" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dimidiatipennis">
<emphasis id="0A663B061DE85BD334F799F48D6DBCFB" italics="true">S. dimidiatipennis</emphasis>
</taxonomicName>
, being brown suffused on apical half (Fig.
<figureCitation id="5F26C1F6FDFB725AC9D21EAA11789754" captionStart="Figures 4954" captionStartId="F12" captionText="Figures 4954. Suragina Walker spp. wing dorsal view: 49 S. monogramma (Bezzi) ♀ (BMSA (D) 129988) 50 S. mulanjeensis Muller, sp. nov. ♀ (BMSA (D) 92374) 51 S. nigromaculata Muller, sp. nov. ♂ holotype (NHMUK 014064156) 52 S. pauliani ♂ paratype (NMSA - DIP 028172) 53 S. semiobscura Muller, sp. nov. ♀ paratype (ICIPE) 54 S. zombaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158451). 51 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures49-54" httpUri="https://binary.pensoft.net/fig/1205874">54</figureCitation>
vs Fig.
<figureCitation id="CBF3FE8F46FD628DA3781A669DC75B8A" captionStart="Figures 4348" captionStartId="F11" captionText="Figures 4348. Suragina Walker spp. wing dorsal view: 43 S. dimidiatipennis (Brunetti) ♀ holotype (NHMUK 014064158) 44 S. falsa Oldroyd ♀ (NHMUK 014064160) 45 S. freidbergi Muller, sp. nov. ♀ paratype (BMSA (D) 92158) 46 S. liberiaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158445) 47 S. malavaensis Muller, sp. nov. ♂ (ICIPE) 48 S. milloti (Séguy) ♀ (CSCA). 43, 44 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures43-48" httpUri="https://binary.pensoft.net/fig/1205873">43</figureCitation>
).
<taxonomicName id="5F33C4098262A68C916B6A8F6A104494" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="zombaensis" status="sp. nov.">
<emphasis id="2E77BC5ACCCAF8DB25BAF0F1D95DFAC7" italics="true">Suragina zombaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="0F61C7BA97113A3327010BB22C15F2D2" rank="species">sp. nov.</taxonomicNameLabel>
(Figs
<figureCitation id="A614148F9F6778F92D87C5B7FC0C5479" captionStart="Figures 3436" captionStartId="F9" captionText="Figures 3436. Suragina Walker spp. lateral habitus: 34 S. semiobscura Muller, sp. nov. ♀ holotype (SAM-DIP-A 018385); S. zombaensis Muller, sp. nov.: 35 ♂ paratype (NMSA - DIP 158421) 36 ♀ paratype (NMSA - DIP 158415). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures34-36" httpUri="https://binary.pensoft.net/fig/1205871">35</figureCitation>
,
<figureCitation id="2A6F63C07E750BB998125FE35DCFB0B7" captionStart="Figures 3436" captionStartId="F9" captionText="Figures 3436. Suragina Walker spp. lateral habitus: 34 S. semiobscura Muller, sp. nov. ♀ holotype (SAM-DIP-A 018385); S. zombaensis Muller, sp. nov.: 35 ♂ paratype (NMSA - DIP 158421) 36 ♀ paratype (NMSA - DIP 158415). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures34-36" httpUri="https://binary.pensoft.net/fig/1205871">36</figureCitation>
), however, differs from
<taxonomicName id="A957EA1D6DEFC3D6E85DAFEBF89E457A" baseAuthorityName="Brunetti" baseAuthorityYear="1929" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dimidiatipennis">
<emphasis id="3EF6DF093C620E6AD1F3EB812A39DAAF" italics="true">S. dimidiatipennis</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="A6FFE59471FDFF26E316490846666A30" captionStart="Figures 18, 19" captionStartId="F4" captionText="Figures 18, 19. Suragina Walker spp. dorsal habitus: S. dimidiatipennis (Brunetti): 18 ♀ holotype (NHMUK 014064158); S. falsa Oldroyd: 19 ♀ holotype (NHMUK 014064160). 18, 19 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures18-19" httpUri="https://binary.pensoft.net/fig/1205866">18</figureCitation>
) in being a much darker species, with mostly dark abdominal tergites with grey dusted apical margins, compared to
<taxonomicName id="A208B5F862DFC6991C3866EC35BBACD3" baseAuthorityName="Brunetti" baseAuthorityYear="1929" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dimidiatipennis">
<emphasis id="DB4859DE9C2D402AC3E6C8F48D7EB61A" italics="true">S. dimidiatipennis</emphasis>
</taxonomicName>
having a mostly orange-yellow to brown abdomen without any dark markings.
<taxonomicName id="872AB5C20654B43DA7EC9BC52403EBDA" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="freidbergi" status="sp. nov.">
<emphasis id="2EEF49A010B9B931296909DFCD20A5D7" italics="true">Suragina freidbergi</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="82DFC25A4636F54C5F24B3A9D9127CFE" rank="species">sp. nov.</taxonomicNameLabel>
is also similar to
<taxonomicName id="F3019F26874994966015E101C87CC28A" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="zombaensis" status="sp. nov.">
<emphasis id="3FEEF27DADACA9050A4A8FCFD5A7BA9F" italics="true">S. zombaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="C4E230A32EBDF080674EDC3C90DD1C45" rank="species">sp. nov.</taxonomicNameLabel>
(see
<taxonomicName id="022BF531532984006719FECE04DF1F9C" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="freidbergi" status="sp. nov.">
<emphasis id="2BAC07772C41965841CCD740E68C2419" italics="true">S. freidbergi</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="FCF6C4360DDA77317CE73986B405E295" rank="species">sp. nov.</taxonomicNameLabel>
diagnosis).
</paragraph>
<caption id="8D7F87F9FA2A5BA0C740214B1EA9F43D" ID-DOI="10.3897/afrinvertebr.65.140524.figures55-62" ID-arpha="C2697A46-D67B-5892-AFEA-369E3231393E" httpUri="https://binary.pensoft.net/fig/1205875" startId="F13">
<paragraph id="A9B1D6ADB27A6B490E8BC286277241B1">
<label id="331E7F9D303579E03845DB21FA4FB815">Figures 5562.</label>
</paragraph>
<paragraph id="775F018533670944B4909871626114BE">
<taxonomicName id="D379092926DECD750F719AF7B9F3409B" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C84565E822B6D05CF92C85137CCC934E" italics="true">Suragina</emphasis>
Walker spp.
</taxonomicName>
male terminalia, ventral view (
<emphasis id="1A1EDDAC79354674D220EA5F27BD2963" bold="true">5558</emphasis>
) and epandrium with cerci, dorsal view (
<emphasis id="BD3568C4F15A9C862172072AD8C40B83" bold="true">5962</emphasis>
):
<emphasis id="4BF7635555B0116DCA99158145DCE690" bold="true">55, 59</emphasis>
<taxonomicName id="1401840A1B9F8CBE206E19C84A989FAE" authority="(Bezzi) (ICIPE)" authorityName="ICIPE" baseAuthorityName="Bezzi" baseAuthorityYear="1926" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="agramma">
<emphasis id="F8D12B84BF518A3076255146A8B5DC05" italics="true">S. agramma</emphasis>
(Bezzi) (
<abbrev id="ABBRID0E3FDK" xlink_title="International Centre of Insect Physiology and Ecology">ICIPE</abbrev>
)
</taxonomicName>
<emphasis id="77F80632416D807C963382ECEB57F5AF" bold="true">56, 60</emphasis>
<taxonomicName id="86AF32D739E1AE6E2067EA98AFC88EFD" authority="(Curran)" baseAuthorityName="Curran" baseAuthorityYear="1928" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="59197AA5F500AFF4335B53F612BD957D" italics="true">S. bezzii</emphasis>
(Curran)
</taxonomicName>
(SAM-DIP A 018382)
<emphasis id="60110E9F11EDD795AD81A8F3845BB3C4" bold="true">57, 61</emphasis>
<taxonomicName id="FE5E9326B307E93202D0D89E799254E6" authority="(Bequaert)" authorityName="" baseAuthorityName="Bequaert" baseAuthorityYear="1921" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="binominata">
<emphasis id="EBCF6423D96EC445D528229F0E4EDCDF" italics="true">S. binominata</emphasis>
(Bequaert)
</taxonomicName>
(
<named-content id="D20C7BFA390B5BDB96184ABD9BEA22BD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
- DIP 028165)
<emphasis id="C7C887C75A378D69D72779B8A73E8F38" bold="true">58, 62</emphasis>
<taxonomicName id="6B1CA56095622168E1E6867815BB8410" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="copelandi" status="sp. nov.">
<emphasis id="B91FE9617958A88559802D979D5F271B" italics="true">S. copelandi</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="1C3FB38E55C2E1B5A60ED2E8126CA5B8" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<named-content id="9758995FC8A1774B00D3C13473F6CBA2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-bloemfontein" xlink_title="National Museum, Bloemfontein">BMSA</named-content>
(D) 83425). Abbreviations: aed tn aedeagal tine; cerc cercus; ej apod ejaculatory apodeme; epand epandrium; goncx gonocoxite; goncx apod gonocoxal apodeme; gonst gonostylus; pm paramere; pm apod parameral apodeme; pm sh parameral sheath. Scale bar (applicable to all illustrations): 0.6 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EUHDK" type="description">
<paragraph id="22E77278F0F197A1D171FA22AF0021A4">
<heading id="898D2480948ACAA942C9BE0EC753638B" reason="title">Description.</heading>
</paragraph>
<paragraph id="4F37E9D6F2FF37521A0EF5E8B0F715B6">
<emphasis id="282C405103570E93BF7615B879E69BAF" bold="true" italics="true">Measurements</emphasis>
(
<specimenCount id="E2F357D6CCAA80E943BBC2061C94F9FD" count="1" type="male"></specimenCount>
n = 2,
<specimenCount id="A53D97C60706C4770C34D6E637DF62A9" count="1" type="female"></specimenCount>
n = 2): Wing span:
<specimenCount id="67E746B4673F2763B9D773380E83CC73" count="1" type="male"></specimenCount>
<quantity id="94ACC4C871527D5F09C9104E584EDBAF" metricMagnitude="-3" metricUnit="m" metricValue="8.2" metricValueMax="8.7" metricValueMin="7.7" unit="mm" value="8.2" valueMax="8.7" valueMin="7.7">7.78.7 mm</quantity>
(avg.
<quantity id="D8A1233B15AEC1EB7A6687FB21764AEA" metricMagnitude="-3" metricUnit="m" metricValue="8.2" unit="mm" value="8.2">8.2 mm</quantity>
);
<specimenCount id="39C84170A4E54167483306BC326846CF" count="1" type="female"></specimenCount>
<quantity id="89A54E1E959B601506DC18FF3D8DDEB7" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="9.8" metricValueMin="8.7" unit="mm" value="9.25" valueMax="9.8" valueMin="8.7">8.79.8 mm</quantity>
(avg.
<quantity id="EF429A4CC3C1AE4FF858529D613446F9" metricMagnitude="-3" metricUnit="m" metricValue="9.3" unit="mm" value="9.3">9.3 mm</quantity>
); body length:
<specimenCount id="234401AA5B748EC5E894D7C7987F7A72" count="1" type="male"></specimenCount>
<quantity id="A34A557AF0516268C7AC6EBA270C6A79" metricMagnitude="-3" metricUnit="m" metricValue="9.4" metricValueMax="10.5" metricValueMin="8.3" unit="mm" value="9.4" valueMax="10.5" valueMin="8.3">8.310.5 mm</quantity>
(avg.
<quantity id="78AC24FD0A05C1E7FA7A6C0F621CF0DC" metricMagnitude="-3" metricUnit="m" metricValue="9.4" unit="mm" value="9.4">9.4 mm</quantity>
);
<specimenCount id="5A715CD38087E18B40ABA5B1F1E7B3A8" count="1" type="female"></specimenCount>
<quantity id="36632877F2D03F2C11FE71F41B707219" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="9.6" metricValueMin="8.9" unit="mm" value="9.25" valueMax="9.6" valueMin="8.9">8.99.6 mm</quantity>
(avg.
<quantity id="33A479261B97DA408A1DA24D7987A115" metricMagnitude="-3" metricUnit="m" metricValue="9.2" unit="mm" value="9.2">9.2 mm</quantity>
); wing span to body length ratio (avg.):
<specimenCount id="2F82295897E9BCC48EA0A687960533A8" count="1" type="male"></specimenCount>
0.97;
<specimenCount id="6EA7BFAD6FE04388D767130EB2B87DC0" count="1" type="female"></specimenCount>
0.99.
</paragraph>
<paragraph id="2D30EABEDABDC49D6CB6DA0E28C5475B">
<emphasis id="906FBBC0788F4E3CC6EAB736A9AAE410" bold="true">Male</emphasis>
(Fig.
<figureCitation id="209E63B44D9BFDAB3FB6490CED2D1362" captionStart="Figures 3436" captionStartId="F9" captionText="Figures 3436. Suragina Walker spp. lateral habitus: 34 S. semiobscura Muller, sp. nov. ♀ holotype (SAM-DIP-A 018385); S. zombaensis Muller, sp. nov.: 35 ♂ paratype (NMSA - DIP 158421) 36 ♀ paratype (NMSA - DIP 158415). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures34-36" httpUri="https://binary.pensoft.net/fig/1205871">35</figureCitation>
).
</paragraph>
<paragraph id="9C5FADBFFA8810BEDF5B470C18B3167B">
<emphasis id="F4816C36BBE7C4D05CF3F388C8069EF2" bold="true" italics="true">Head</emphasis>
: Blackish-brown ground colour, with bluish-grey pruinosity on majority of head; eye bare; holoptic; ommatidia on lower and upper quarter of eye smaller than those on rest of eye; lateral edge of eye with slight indentation, and an apparent tubercle next to indentation; ocellar tubercle barely visible in profile, blackish-brown in colour with slight brownish- to bluish-grey pruinosity and short dark setulae; vertex blackish-brown with bluish-grey pruinosity and long dark setulose; anterior ocellus larger than posterior pair; ocellar tubercle in front of dorsal margin of eye, not placed as deeply towards middle of head as in
<specimenCount id="A8D731AAD7111C3D1D33751FE8DF71BA" count="1" type="female"></specimenCount>
; dorsal inner edge of eye abutting ocellar tubercle; occiput with same bluish-grey pruinosity as rest of head; paired narrow subrectangular black markings with short dark setulae on upper occiput, widening only slightly towards lateral margin of head in some specimens, abutting posterior margin of eyes, flanking vertex; upper occiput with pale setulae; lower occiput with long pale setulae, these continue ventrally on head to mouthparts that have similar long ventral setulae; frons bluish-grey pruinose, velvety-black from ocellar tubercle to before lower half of eye when viewed anteriorly, when viewed anteroventrally entire frons appears blackish-brown with a slight velvety appearance; frons widening from where eyes touch down to antennal base; frons with long dark setulae (setulae can appear pale depending on viewing angle); face and gena bluish-grey, lateral margins of face with long pale setulae, gena with long dark setulae; clypeus dark brown with bluish-grey pruinosity, bare; face separated anteriorly from clypeus by a deep transverse suture, similar to lateral sutures; face not appearing to bulge laterally when viewed in profile; clypeus visible in profile, face not; antennal bases separated ca 0.7 × width of scape, with slight longitudinal groove running between; scape dark brown and pedicel orange-brown, both with silvery pruinosity; 1
<superScript id="2C519DBAA30D9F2696CC7F46D83424B8">st</superScript>
flagellomere orange-yellow, with sparse silvery pruinosity; 2
<superScript id="CDDCB580C92A3570725135D17E365C1A">nd</superScript>
flagellomere dark brown; scape 1.5 × length of pedicel; 1
<superScript id="0244DB0721E6422CD80B2B080973C8BD">st</superScript>
flagellomere reniform, ca 1.5 × size of pedicel; 2
<superScript id="96DFA14230C363308D7341E3264BAE3F">nd</superScript>
flagellomere arista-like; pedicel with dark dorsal and ventral setulae, similar in size, scape with only dark dorsal setulae; palpus orange-yellow with dense bluish-grey pruinosity, with dark setulae throughout; palpus ca 0.5 × length of proboscis; proboscis slightly shorter than head height; proboscis brown to dark brown in colour, prementum brown, proboscis with long pale setulae basoventrally and dark setulae on rest of ventral and dorsal surfaces, except for short pale setulae apically.
</paragraph>
<paragraph id="901FEE7F954D374C82541209F3B48E79">
<emphasis id="F470A4043BF1CA80D81B985BD0116100" bold="true" italics="true">Thorax</emphasis>
: Scutum brown, mostly with short dark setulae, with two dorsocentral whitish-brown pruinose vittae running from posterior half of postpronotal lobes to transverse suture; prescutellar area bluish-grey pruinose; pronotum brown with bluish-grey pruinosity and short pale setulae; postpronotal lobe orange-brown, appearing slightly browner dorsally, with sparse bluish-grey pruinosity, setulae pale; notopleuron bluish-grey pruinose with long pale setulae anteriorly and long dark setulae posteriorly; postalar wall dark brown and postalar callus orange-brown, both with slight bluish-grey pruinosity, supra-alar area with dark setulae, postalar callus with some short pale setulae interspersed between dark setulae; scutellum dark brown with a brownish- to orange-yellow apical margin with long dark setulae; majority of pleura brown with bluish-grey pruinosity, except for anepimeron, anatergite, katatergite and katepimeron orange-brown with bluish-grey pruinosity; proepisternum and proepimeron, katatergite and katepisternum with long pale setulae; anatergite and meron bare; anterior and posterior spiracles whitish-yellow, bare; postspiracular scale orange-brown; postscutellum orange-brown.
</paragraph>
<paragraph id="02BA6A5C18DF85A7D044CC61E3B612FB">
<emphasis id="68D3097A9BE2C5B00193820A9A007351" bold="true" italics="true">Legs</emphasis>
: Fore coxa yellowish-brown, mid and hind coxae brown, all three coxae with bluish-grey pruinosity, hind coxa much less pronounced; fore coxal setulae long pale on anterior surface, with dark setulae apically; mid coxal setulae long pale on anterior surface, dark apically; hind coxal setulae dark on anterior edge surrounding well-developed anterior apical point with some scattered short setulae, lateral apical edges with a mix of long pale and dark setulae; all trochanters yellowish-brown with short pale setulae; fore femur almost entirely yellow, except for brown apex; mid femur entirely yellow; hind femur yellow with brown to dark brown median band; fore and hind tibiae brown to dark brown, mid tibia yellow; fore and hind tarsi brown to dark brown, mid tarsi yellow except for terminal 3 segments that are brown; fore tarsal claws asymmetrical, outer claw much larger than inner claw, foreleg empodium ca 2 × size of inner pulvillus, outer pulvillus ca 2 × length of inner, approaching size of outer claw; fore femur overall with short pale setulae, with long pale setulae on posteroventral surface; mid femur with long pale setulae on ventral surface, otherwise with short pale setulae; hind femur with a mix of pale and dark setulae on dorsal and ventral surfaces, base of femur with pale setulae; fore tarsi covered with long sensory setulae along antero- and posteroventral surfaces, sensory setulae ca 2 × as long as tarsal segment is wide; hind leg overall stouter than remaining legs; hind tarsal segments 0.91.0 × as long as hind tibia; tibial spur formula 0: 2: 2.
</paragraph>
<paragraph id="FFC588AEC81A581175594F640CBCDBDF">
<emphasis id="A8E48FD1B89F0D892F2B2EF6B662903C" bold="true" italics="true">Wing</emphasis>
: Brown suffused on majority of surface; dark brown stigma over area of veins
<emphasis id="E065119A965488C73707650A101B28A5" italics="true">
R
<subScript id="77898C7D4FC1F2F6251D546679FA5A88">1</subScript>
</emphasis>
and
<emphasis id="325B99BC2B53C37CA229994BFC6DAABB" italics="true">
R
<subScript id="AF2E5DE60E8A36C0C1F4154948C957F1">2 + 3</subScript>
</emphasis>
and cell
<emphasis id="651E338605638A66E48AC8AAA11EA2BC" italics="true">
r
<subScript id="B33CED215BB28E7722D46F7769E215A5">1</subScript>
</emphasis>
, difficult to discern due to surrounding brown suffusion on apical half of wing; preapical parts of cells
<emphasis id="8B76EA4B78489D3B4BC14B2D2667F250" italics="true">br</emphasis>
,
<emphasis id="6E238A3C6A65672817FCDEA563A27E9A" italics="true">bm</emphasis>
hyaline, most of cell
<emphasis id="51071DE66788C18681190CBC4B8C5B6F" italics="true">cua</emphasis>
and anal lobe, and apical two-thirds of discal cell only lightly brown suffused; some specimens with cells
<emphasis id="D401EC893E929459208D38C123473A0B" italics="true">
r
<subScript id="D7A819DFA6E9D4918D445A7636CB23C3">2 + 3</subScript>
</emphasis>
and
<emphasis id="4FD9795B28E9D91860A576A0DE81188A" italics="true">
m
<subScript id="CEF3959062E35E32B1C96E9B2094A6B0">4</subScript>
</emphasis>
having a small hyaline marking in each of their centres; veins dark brown; costa without distinct downward flexure over stigma; cell
<emphasis id="0FAE851898D065D837FEC06C75F32DA8" italics="true">cua</emphasis>
closed at wing margin, cell
<emphasis id="7F1B772710D13BC623879FBF4136EC65" italics="true">
m
<subScript id="F1E564D8BBF08940B9074C7E46D8615D">3</subScript>
</emphasis>
open, veins
<emphasis id="4ED876CDAC46F3E13C94E6DFAEE1AB69" italics="true">
M
<subScript id="5CE4B3BAAE6DE0B192BB761556BBC01F">1</subScript>
</emphasis>
,
<emphasis id="BB728B5A0E0FC85CC52A26D4C99E785B" italics="true">
M
<subScript id="2061A6E83707E05C7C3C90ED53871CC9">2</subScript>
</emphasis>
,
<emphasis id="8DAFB7EC838C2C2CC42B415D50B6164F" italics="true">
M
<subScript id="E6D4CDF7B882E1DFE01C1EA0A7B0DCDD">3</subScript>
</emphasis>
present; haltere stalk (except for yellow basal half) and knob dark brown, with a few short dark setulae.
</paragraph>
<caption id="75EDE9B7FC21854DD6D9890BD7BE33D1" ID-DOI="10.3897/afrinvertebr.65.140524.figures63-70" ID-arpha="32C54E94-766C-50E3-A12C-AD3E4874CD88" httpUri="https://binary.pensoft.net/fig/1205876" startId="F14">
<paragraph id="19C560E4F1EB6A3EA8484975F6575C01">
<label id="A279B7C3CE2A6C041F4BD84D7DEC235C">Figures 6370.</label>
</paragraph>
<paragraph id="350FFCD6D3A901B35B62A4BF5F5F62E2">
<taxonomicName id="0F63D1F7670F40805F93816DAF7124A2" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2CD4A4EBE2DFD2B91DC308897AA85AF4" italics="true">Suragina</emphasis>
Walker spp.
</taxonomicName>
male terminalia, ventral view (
<emphasis id="562AEEF2DDD4024AF5ABDF635F5D522A" bold="true">6366</emphasis>
) and epandrium with cerci, dorsal view (
<emphasis id="6DC09C7A0EAF0290EDDCE173752AACDD" bold="true">6770</emphasis>
):
<emphasis id="7372935A77BB5ACE6B45805AB4C9FCE1" bold="true">63, 67</emphasis>
<taxonomicName id="9A13241EC6F97F390FAE5A7DAA44E61D" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="freidbergi" status="sp. nov.">
<emphasis id="A29642963C7DDD8283F8613A7657241A" italics="true">S. freidbergi</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="7B17FAB149221B4AE9BD1859720229D4" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<named-content id="4F2F8A171EA1C7D7D392499BE211F0E7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-bloemfontein" xlink_title="National Museum, Bloemfontein">BMSA</named-content>
(D) 92376)
<emphasis id="068132E7348F36FD54A4A204A491A0AF" bold="true">64, 68</emphasis>
<taxonomicName id="E6A94F47BFF5962D3C41B08C73B13EFE" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="malavaensis" status="sp. nov.">
<emphasis id="006F97EE5AC4125C4062F0B319FB5049" italics="true">S. malavaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="F26CD51D2D749758B5E78EA36E868A43" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<abbrev id="ABBRID0EIMDK" xlink_title="International Centre of Insect Physiology and Ecology">ICIPE</abbrev>
)
<emphasis id="1C7F063CA628FC0D9B203CF91967C13E" bold="true">65, 69</emphasis>
<taxonomicName id="BF400186F313D0658E72644AF5B6E8CD" authority="(Seguy) (CSCA)" authorityName="CSCA" baseAuthorityName="Seguy" baseAuthorityYear="1951" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="milloti">
<emphasis id="EE14437B7A503CE7079E661965505A89" italics="true">S. milloti</emphasis>
(Séguy) (
<named-content id="8D26A9C5E415A282627235809573CC2D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-state-collection-arthropods" xlink_title="California State Collection of Arthropods">CSCA</named-content>
)
</taxonomicName>
<emphasis id="5FCE5DA57B31EAEE6431377E0BC2F871" bold="true">66, 70</emphasis>
<taxonomicName id="CA465178D0B0199A51252EC73C008305" authority="(Bezzi)" baseAuthorityName="Bezzi" baseAuthorityYear="1926" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="monogramma">
<emphasis id="70F849BA3A93D7074470425B212AC642" italics="true">S. monogramma</emphasis>
(Bezzi)
</taxonomicName>
(
<named-content id="9DFA2DC22C46E9447093EB6FF5DE23F2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
- DIP 028180). Scale bar (applicable to all illustrations): 0.6 mm.
</paragraph>
</caption>
<paragraph id="03CEAF2026A158EA8F6983B7DEFF7455">
<emphasis id="6EC57FD84C2229469A1FC0331A544FDD" bold="true" italics="true">Abdomen</emphasis>
: Overall brownish-yellow colour; tergite 1 light brownish-yellow with a dark brown subtriangular dorsal marking surrounded by bluish-grey pruinosity up to lateral margins; tergites 2 and 3 with a dark brown longitudinal rectangular marking, otherwise brownish-yellow, tergite 3 also with a grey pruinose posterior band; tergite 4 with a dark brown subtriangular marking running towards a dark posterior band with greyish pruinosity; tergite 5 entirely dark brown with posterior grey pruinose band; remaining tergites dark brown; tergite 1 medially without a longitudinal suture; tergites with dark setulae dorsally, and long pale setulae laterally up to margins; sternites mostly yellowish, except sternites 4 and 5 dark brown; sternites with short and long pale setulae.
</paragraph>
<paragraph id="B05428A860DDCC63C313F48527A8AEA0">
<emphasis id="FB9F22F0C6913F0FCFA01FC60E9A3DC2" bold="true" italics="true">Terminalia</emphasis>
(Figs
<figureCitation id="06EC9D20515A927002F4172308E3C9FA" captionStart="Figures 7174" captionStartId="F15" captionText="Figures 7174. Suragina Walker spp. male terminalia, ventral view (71, 72) and epandrium with cerci, dorsal view (73, 74): 71, 73 S. mulanjeensis Muller, sp. nov. paratype (NMSA - DIP 162049) 72, 74 S. zombaensis Muller, sp. nov. paratype (NMSA - DIP 158450). Scale bar (applicable to all illustrations): 0.6 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures71-74" httpUri="https://binary.pensoft.net/fig/1205877">72</figureCitation>
,
<figureCitation id="5AA00F5CF18D10944754DF822E0ADD1F" captionStart="Figures 7174" captionStartId="F15" captionText="Figures 7174. Suragina Walker spp. male terminalia, ventral view (71, 72) and epandrium with cerci, dorsal view (73, 74): 71, 73 S. mulanjeensis Muller, sp. nov. paratype (NMSA - DIP 162049) 72, 74 S. zombaensis Muller, sp. nov. paratype (NMSA - DIP 158450). Scale bar (applicable to all illustrations): 0.6 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures71-74" httpUri="https://binary.pensoft.net/fig/1205877">74</figureCitation>
): Epandrium and cercus dark brown with dark setulae; gonocoxite, hypoproct and hypandrium with pale setulae; gonostylus almost parallel shaped with truncated apex, outer edge of gonostylus with short setulae, inner edge with protrusion with 4 setulae, apex of gonostylus sparsely covered in microtrichia; gonocoxite widening and appearing more rounded on apical half, apex somewhat flattened, gonocoxite outer and ventral medial surface with long setulae, inner and median surface of upper half with short setulae, lower ventral surface comparatively less setulose; gonocoxite with microtrichia between setulae; parameral apodeme with truncated apex, not reaching base of gonocoxite in ventral view, parameral sheath including parameral apodeme ca 0.7 × length of gonocoxite; gonocoxal apodeme similar in length to gonocoxite and slightly shorter than ejaculatory apodeme; aedeagal tine curvature barely extending down past gonocoxites, apex of tines extending out slightly past parameral sheath; endoaedeagal process apically truncated and widened.
</paragraph>
<caption id="3836FD21B7D8C2ACD3D5C7434BE882DE" ID-DOI="10.3897/afrinvertebr.65.140524.figures71-74" ID-arpha="55CED672-3716-51FB-B728-FD7DE3EC2B8A" httpUri="https://binary.pensoft.net/fig/1205877" startId="F15">
<paragraph id="237DA520DD6C91D0308D9B07D8D2B602">
<label id="7DF29C97AA11914366142DBF1EE43E53">Figures 7174.</label>
</paragraph>
<paragraph id="0F3A1EE0998AC9D7F3BCDE7E9C65B342">
<taxonomicName id="3C80C88C8369CE927EC2C56FE0EB1269" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="83325F3EF7900F2F61C65A5F98A38339" italics="true">Suragina</emphasis>
Walker spp.
</taxonomicName>
male terminalia, ventral view (
<emphasis id="FEC62C58E3A37DFEB2AFCC2AAA2B95CC" bold="true">71, 72</emphasis>
) and epandrium with cerci, dorsal view (
<emphasis id="5597A18E80535264A9BC2AB8E8C68A5D" bold="true">73, 74</emphasis>
):
<emphasis id="6FF9F2D0A58D75FE9E571E6EC3795998" bold="true">71, 73</emphasis>
<taxonomicName id="AA4D4F0BAF6147FAD342399B16AC698A" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="mulanjeensis" status="sp. nov.">
<emphasis id="9EF537C203B4A0C01AE3639F515BCD21" italics="true">S. mulanjeensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="28B700801EDF4D8F6F763CD2CC3F9989" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<named-content id="C8019E34404733F7F203048A72524DDB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
- DIP 162049)
<emphasis id="5B0C93C77D4A58E25058E61A3993F4D6" bold="true">72, 74</emphasis>
<taxonomicName id="B9CA84AC9CA67D5C6F1A78A0B0F44EFD" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="zombaensis" status="sp. nov.">
<emphasis id="79D77B1A3E5FAEC08BBEA2227E10C260" italics="true">S. zombaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="699987D181D31B005357C646580A75B0" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<named-content id="59A25268BBDC4FCAAE90B78D4637EF5A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
- DIP 158450). Scale bar (applicable to all illustrations): 0.6 mm.
</paragraph>
</caption>
<paragraph id="464A4A39616A01C0099F531B30E030EF">
<emphasis id="77C30472FD29936A4784014C1EAF6287" bold="true">Female</emphasis>
(Fig.
<figureCitation id="4AA161BE888A272DAD8B4686E8938E9A" captionStart="Figures 3436" captionStartId="F9" captionText="Figures 3436. Suragina Walker spp. lateral habitus: 34 S. semiobscura Muller, sp. nov. ♀ holotype (SAM-DIP-A 018385); S. zombaensis Muller, sp. nov.: 35 ♂ paratype (NMSA - DIP 158421) 36 ♀ paratype (NMSA - DIP 158415). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures34-36" httpUri="https://binary.pensoft.net/fig/1205871">36</figureCitation>
): Similar to
<specimenCount id="0440101DA096E8E7B5C6345093DCCF04" count="1" type="male"></specimenCount>
except for the following:
</paragraph>
<paragraph id="F55C11E76317F906CBCD2D8E70B8E8F4">
<emphasis id="94748CD95B4CC35C12E768C4EA8FA24C" bold="true" italics="true">Head</emphasis>
: Dichoptic; ommatidia of similar size except for those on upper quarter that are somewhat smaller; lateral edge of eye without any indentation, but also with apparent tubercle as in
<specimenCount id="B6067B2EC31F6C26C54CD6A713DA2598" count="1" type="male"></specimenCount>
; anterior ocellus similar in size to posterior pair; ocellar tubercle placed deeper in front of dorsal margin of eye compared to
<specimenCount id="EB9F1BF6363A834BD23A321C676A18CF" count="1" type="male"></specimenCount>
; dorsal inner edge of eye separated from ocellar tubercle by paired silver-grey markings, appearing to extend down from vertex; upper occiput with some short dark setulae below black rectangular markings (
<specimenCount id="819494C71407E2FD34219BA6EF430885" count="1" type="male"></specimenCount>
without); frons bluish-grey pruinose between lower half of eye down to antennal base, velvety-black from ocellar tubercle to lower half of eye when viewed anteriorly; frons running almost parallel down to antennal base; frons densely covered with dark setulae on velvety-black upper half, with dark setulae on lateral margins of lower half; face separated anteriorly from clypeus by transverse suture, (less prominent than in
<specimenCount id="C08A5AA79596AA29DFBE08C2E9E34B65" count="1" type="male"></specimenCount>
); antennal bases separated ca 0.60.8 × width of scape, with slight longitudinal groove running between; scape 1.7 × length of pedicel; 1
<superScript id="8D9ED96A42F3B43436A9137B4BB342D9">st</superScript>
flagellomere ca 2 × size of pedicel; proboscis ca same length as head height.
</paragraph>
<paragraph id="3E747ECE2C9EFD6D7E3A65A307869E45">
<emphasis id="84CD7A8000116C066045413CA8C5CA57" bold="true" italics="true">Thorax</emphasis>
: Overall similar to
<specimenCount id="3379F3C0D2FC20A2C653C6EC9720E184" count="1" type="male"></specimenCount>
; scutum similar to that of
<specimenCount id="4F79DC62809859B70F8CC5CC996CE5BE" count="1" type="male"></specimenCount>
, except two dorsocentral vittae are bluish-grey pruinose compared to whitish-brown in
<specimenCount id="2DCC97A5671B4357EA4C3C2FC2893186" count="1" type="male"></specimenCount>
.
</paragraph>
<paragraph id="00991524BA185766ECB64FBCC25B0A56">
<emphasis id="725A221E3D25B6C95E1ACD4BA9454687" bold="true" italics="true">Legs</emphasis>
: Similar to
<specimenCount id="326B8AA06C7B0CE45077178191254BCE" count="1" type="male"></specimenCount>
in colour; fore tarsal claws symmetrical; overall leg setation similar to
<specimenCount id="1C361590825FD29842CC302DC764D845" count="1" type="male"></specimenCount>
except generally shorter and coxal setulae all pale except for dark setulae surrounding anterior apical point of hind coxa; hind tarsal segments 0.981.01 × as long as hind tibia
</paragraph>
<paragraph id="CACD95BDD02A080788E29E57D4AC8654">
<emphasis id="957364D8F9891262B1209DADB8296208" bold="true" italics="true">Wing</emphasis>
(Fig.
<figureCitation id="169EF53FFB8228475D5B767F71081ED6" captionStart="Figures 4954" captionStartId="F12" captionText="Figures 4954. Suragina Walker spp. wing dorsal view: 49 S. monogramma (Bezzi) ♀ (BMSA (D) 129988) 50 S. mulanjeensis Muller, sp. nov. ♀ (BMSA (D) 92374) 51 S. nigromaculata Muller, sp. nov. ♂ holotype (NHMUK 014064156) 52 S. pauliani ♂ paratype (NMSA - DIP 028172) 53 S. semiobscura Muller, sp. nov. ♀ paratype (ICIPE) 54 S. zombaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158451). 51 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures49-54" httpUri="https://binary.pensoft.net/fig/1205874">54</figureCitation>
): Similar to
<specimenCount id="CE002EA5588257DFA4146B5FF4C8B59A" count="1" type="male"></specimenCount>
.
</paragraph>
<paragraph id="9B8570ED03519D78A99BA1269175F2F9">
<emphasis id="4AEC4B681D19D31D69489C095A4C51E1" bold="true" italics="true">Abdomen</emphasis>
: Tergite 1 with a narrower, rectangular (
<specimenCount id="AE4B79C71B168379950081C803E3F9D1" count="1" type="male"></specimenCount>
subtriangular) dark brown marking, more widely bluish-grey pruinose than in
<specimenCount id="0D79F6506AA14CBE18DCC7BD3F773BF3" count="1" type="male"></specimenCount>
, medially with a longitudinal suture; tergite 2 with similar dark longitudinal marking, but surrounded by bluish-grey pruinosity instead of brownish-yellow ground colour; tergites 35 dark brown, posterior margins with similar grey pruinose bands as in
<specimenCount id="D98E268117A4910CE293DF2041B23ACF" count="1" type="male"></specimenCount>
; tergite 1 with long pale setulae on pruinose surface, and short dark setulae on dark brown surfaces, tergites 25 with short pale setulae on posterior pruinose bands and short dark setulae on rest of surface; tergites 6 and 7 with short dark setulae; tergites 15 with long pale setulae on lateral margins, tergites 6 and 7 with dark setulae laterally; sternites all yellow, compared to sternites 5 and 6 dark brown in
<specimenCount id="1B8B79B1846646F67F3A4077E308BB9F" count="1" type="male"></specimenCount>
.
</paragraph>
<paragraph id="DCD061B8489C0B80D735FB507909CF88">
<emphasis id="755039375DBB212E2FB517A942799CA3" bold="true" italics="true">Terminalia</emphasis>
(Fig.
<figureCitation id="496C2DAF964CC92DDE2A6093EBC18C88" captionStart="Figures 7587" captionStartId="F16" captionText="Figures 7587. Suragina Walker spp. female genital fork and spermathecae, ventral view: 75 S. agramma (Bezzi) (ICIPE) 76 S. bezzii (Curran) (SAM-DIP- 018403) 77 S. bilobata Muller, sp. nov. (CSCA) 78 S. binominata (Bequaert) (NMSA - DIP 158411) 79 S. copelandi Muller, sp. nov. paratype (BMSA (D) 84686) 80 S. freidbergi Muller, sp. nov. paratype (BMSA (D) 92158) 81 S. liberiaensis Muller, sp. nov. paratype (NMSA - DIP 158442) 82 S. malavaensis Muller, sp. nov. paratype (ICIPE) 83 S. milloti (Séguy) (CSCA) 84 S. monogramma (Bezzi) (BMSA (D) 129988) 85 S. mulanjeensis Muller, sp. nov. paratype (NMSA - DIP 158392) 86 S. semiobscura Muller, sp. nov. paratype (ICIPE) 87 S. zombaensis Muller, sp. nov. paratype (NMSA - DIP 1584719). Scale bar (applicable to all illustrations): 0.1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures75-87" httpUri="https://binary.pensoft.net/fig/1205878">87</figureCitation>
): Cercus yellowish-brown with some greyish pruinosity, with pale setulae; genital fork with distal apodeme ending broadly truncated; median lobe with narrow emargination; paired apical lobes with somewhat slender appearance, inner surface with clustered microtrichia at apex; arms gradually rounded; three ovate and sclerotised spermathecae.
</paragraph>
<caption id="82B2365461EFB559CBF87A61FC0BB252" ID-DOI="10.3897/afrinvertebr.65.140524.figures75-87" ID-arpha="A46B73BF-0DF3-5EFD-9F88-8430B3953721" httpUri="https://binary.pensoft.net/fig/1205878" startId="F16">
<paragraph id="D97A5042BD95AF9B474BD2217493CA72">
<label id="16E8CE23D7482D6CB9EC9410C0D80999">Figures 7587.</label>
</paragraph>
<paragraph id="34C56E23E15B1601D539E93D4B77BB9E">
<taxonomicName id="11B5CB71D4D41F2B130BA514CA5B5F34" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="12AD038BECE9D6631A1956557D178A7B" italics="true">Suragina</emphasis>
Walker spp.
</taxonomicName>
female genital fork and spermathecae, ventral view:
<emphasis id="468FB02F027AB28ADC614C09C2DA7CE5" bold="true">75</emphasis>
<taxonomicName id="6AA7A22BB6F0F02B948A8054CF6D7AD8" authority="(Bezzi) (ICIPE)" authorityName="ICIPE" baseAuthorityName="Bezzi" baseAuthorityYear="1926" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="agramma">
<emphasis id="0372D6D60036A64A641B087FEB882923" italics="true">S. agramma</emphasis>
(Bezzi) (
<abbrev id="ABBRID0E2SDK" xlink_title="International Centre of Insect Physiology and Ecology">ICIPE</abbrev>
)
</taxonomicName>
<emphasis id="0A2AF41F06A6155E52C8260FDCE3603F" bold="true">76</emphasis>
<taxonomicName id="31DE183E7775EF273A98C62932D2743A" authority="(Curran)" baseAuthorityName="Curran" baseAuthorityYear="1928" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="A66ACBA01A2D8945E91F93F93B049FE4" italics="true">S. bezzii</emphasis>
(Curran)
</taxonomicName>
(SAM-DIP- 018403)
<emphasis id="6E8DE8F13537C866AF0EE0F517628CD5" bold="true">77</emphasis>
<taxonomicName id="AEB58E4A5C3F7F0977831A750E0783DE" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bilobata" status="sp. nov.">
<emphasis id="6D7BE02CADF2FC86ADD08BE808730A5F" italics="true">S. bilobata</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="867CD57FEE83A06D9A9A8FF76CFCC189" rank="species">sp. nov.</taxonomicNameLabel>
(
<named-content id="53ED852CE2F2B3DA41C5A03361B35A1B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-state-collection-arthropods" xlink_title="California State Collection of Arthropods">CSCA</named-content>
)
<emphasis id="8E53D89447E9E91BF0429B824267237E" bold="true">78</emphasis>
<taxonomicName id="E83A03CEC7B1BDB5CA32A57376E3460D" authority="(Bequaert)" authorityName="" baseAuthorityName="Bequaert" baseAuthorityYear="1921" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="binominata">
<emphasis id="FC8FF0DC193C2135DE55BFBB1BEFFB70" italics="true">S. binominata</emphasis>
(Bequaert)
</taxonomicName>
(
<named-content id="B19077E140D5DCC3B51FCD7D3CEC1BBB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
- DIP 158411)
<emphasis id="29735F12D25F6B72210A9AE6CBFB0FB5" bold="true">79</emphasis>
<taxonomicName id="86FD68977D98C1416EBBE8368488DA27" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="copelandi" status="sp. nov.">
<emphasis id="6E8ED4261BB4D7824EDFB5FD776C2924" italics="true">S. copelandi</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="4859A6B567131D94C5D98B839BDC8B88" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<named-content id="E9BD66BA20A21FBB3519F14DC345902D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-bloemfontein" xlink_title="National Museum, Bloemfontein">BMSA</named-content>
(D) 84686)
<emphasis id="3A43EB061994350F78F2E35CFB136853" bold="true">80</emphasis>
<taxonomicName id="3AA268CFC01D649A8C1284DA9DF7B2A2" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="freidbergi" status="sp. nov.">
<emphasis id="32088F3058B6931ABA421A5B6B3C20D7" italics="true">S. freidbergi</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="FFBF26AEF872E7466D004E8E509E0660" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<named-content id="86D830B011B73405F5F34A716BF6D47B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-bloemfontein" xlink_title="National Museum, Bloemfontein">BMSA</named-content>
(D) 92158)
<emphasis id="9D800606D4259019B2C09D005D8B93D8" bold="true">81</emphasis>
<taxonomicName id="EEAFA955E30DB4E68A39C588DFDF0952" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="liberiaensis" status="sp. nov.">
<emphasis id="F803B8878BB2DE34DD499E89B36F3B8D" italics="true">S. liberiaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="F01545B39E7B7C8B627E5E0058E3ED92" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<named-content id="914F0E0E6EAF93A33FD2D67A0C6AED45" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
- DIP 158442)
<emphasis id="3ACB3B95FF0EAA43E2DCF320643ACEBA" bold="true">82</emphasis>
<taxonomicName id="E5D67F7474FF8EC3837A48D3A202DDA4" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="malavaensis" status="sp. nov.">
<emphasis id="CA4F469EEF3319CEBAD25BD00F5CC499" italics="true">S. malavaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="26A1A70172A947DF9627B96AD941AE97" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<abbrev id="ABBRID0EOWDK" xlink_title="International Centre of Insect Physiology and Ecology">ICIPE</abbrev>
)
<emphasis id="D233F0DE32EAA9CF583E0B618967B9A5" bold="true">83</emphasis>
<taxonomicName id="E275CB0CDE78CB18E123939060185EBE" authority="(Seguy) (CSCA)" authorityName="CSCA" baseAuthorityName="Seguy" baseAuthorityYear="1951" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="milloti">
<emphasis id="FB5D629C89D3C114665C9215673B8771" italics="true">S. milloti</emphasis>
(Séguy) (
<named-content id="6DBE10AC6D796B690403FCE17429602B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-state-collection-arthropods" xlink_title="California State Collection of Arthropods">CSCA</named-content>
)
</taxonomicName>
<emphasis id="92DD2D3751DFE6AB64EA947A8F8E86B3" bold="true">84</emphasis>
<taxonomicName id="9EE5A09CD6F08EF13605CA3056B3AEE3" authority="(Bezzi)" baseAuthorityName="Bezzi" baseAuthorityYear="1926" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="monogramma">
<emphasis id="A105A61227592810AC8F6F455FB7B7AD" italics="true">S. monogramma</emphasis>
(Bezzi)
</taxonomicName>
(
<named-content id="27A2E0096B5C5E00604CEEE593D44477" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-bloemfontein" xlink_title="National Museum, Bloemfontein">BMSA</named-content>
(D) 129988)
<emphasis id="787B36F5D78A4DF62732072ED8A80586" bold="true">85</emphasis>
<taxonomicName id="48640C29107D307A0B2A8E7B21CA4374" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="mulanjeensis" status="sp. nov.">
<emphasis id="DB5BF1D3F4A0E68612F301CE975B7934" italics="true">S. mulanjeensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="B1E228DDB538E0C8670DAD153F92BB6D" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<named-content id="65B9ED64E50EA012ABFAAD2A47801D9D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
- DIP 158392)
<emphasis id="EC0EB71A78EC979EEC61321CCAB8D0C7" bold="true">86</emphasis>
<taxonomicName id="A5D80B3F75960BFC52425C429827E00C" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="semiobscura" status="sp. nov.">
<emphasis id="A0B17316FA383B56E74232F209CF73BD" italics="true">S. semiobscura</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="54494ACE3416E4868EC8E22F264910A7" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<abbrev id="ABBRID0ESYDK" xlink_title="International Centre of Insect Physiology and Ecology">ICIPE</abbrev>
)
<emphasis id="AA4C7BA744CDF85BE3529B5C977C0F9F" bold="true">87</emphasis>
<taxonomicName id="839F251E3594B0A83E58857587612EDD" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="zombaensis" status="sp. nov.">
<emphasis id="5B57438EB3416FFE1796871249CB4BB1" italics="true">S. zombaensis</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="4F1CF9D6B98291C3798BCE0F47FA0644" rank="species">sp. nov.</taxonomicNameLabel>
paratype (
<named-content id="2ACE39EB09C1ACC86044559F2AC954E8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/kwazulu-natal-museum" xlink_title="KwaZulu-Natal Museum">NMSA</named-content>
- DIP 1584719). Scale bar (applicable to all illustrations): 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EMZDK" type="etymology">
<paragraph id="1C6293FEC8F320D1F9A1941BCBDC1939">
<heading id="86FC8B5439FB6786BC11E55A9EDF6A85" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="3DE2D2CC2326E3282069C789456A764D">
Named after the
<typeStatus id="ADDB1F7123AED50668B3CDB289549DD3">type</typeStatus>
locality, the
<collectingRegion id="AD60C1AFC52FEA97B733AC298C7FFB6C" country="Malawi" name="Zomba">Zomba</collectingRegion>
Plateau,
<collectingCountry id="C0468C1B382151AC91B86BEE01D5C61E" name="Malawi">Malawi</collectingCountry>
. Feminine adjective in the nominative singular case.
</paragraph>
<caption id="42999F94929C35C894D90BFB4C1FE74A" ID-DOI="10.3897/afrinvertebr.65.140524.figures88-89" ID-arpha="B878678C-9BFC-59BC-BA3B-77A7A8113828" httpUri="https://binary.pensoft.net/fig/1205879" startId="F17">
<paragraph id="976DF6851ADD4F51015C99B7DFF0E7CA">
<label id="5462CE1236D02A18244BB04110E88FAD">Figures 88, 89.</label>
</paragraph>
<paragraph id="C772946F3B0A8E31BBAF2B7C4E1FB8AD">
Distribution of
<taxonomicName id="2A2D3021F127C0651633A3FE54E97260" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8723FE0043301513F32B90E574E7524B" italics="true">Suragina</emphasis>
Walker spp.
</taxonomicName>
within the Afrotropical Region.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EF1DK" type="distribution">
<paragraph id="8B2072D6B27E7C7764548B5591A48254">
<heading id="7913DEC4245F5377127E93F7FE6B3143" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="71B63281C08B99473FCEAF05B13B664A">
<collectingCountry id="961A55660D5A9211F148824112C07B96" name="Malawi">Malawi</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,751 @@
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<mods:title id="00E15960FFDCB824DD4EA67947CACE59">Revision of Afrotropical Suragina Walker, 1859 (Diptera, Athericidae)</mods:title>
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<mods:role id="A750B639ECBBB0D6455745BD6F4EC782">
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<mods:namePart id="4AAF426B0D7B7F83428ED86EADEA3668">Muller, Burgert S.</mods:namePart>
<mods:nameIdentifier id="07ED1A646E174DE8A6763EA99A888680" type="ORCID">0000-0002-7304-4050</mods:nameIdentifier>
<mods:affiliation id="FF46EA19AD986C3D97E00BAC6C68B37A">Department Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, 9301, South Africa &amp; Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
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<mods:namePart id="08118479DE4BDE10513E980DB78F1E30">Swart, Vaughn R.</mods:namePart>
<mods:nameIdentifier id="7F1515BC291D796081AEA193E9F65814" type="ORCID">0000-0001-7905-5298</mods:nameIdentifier>
<mods:affiliation id="671F9A44A8044D7701F01606107D4AD6">Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
</mods:name>
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<mods:namePart id="9597EFDE7AEAF210142B4BD819CF02C0">Snyman, Louwrens P.</mods:namePart>
<mods:nameIdentifier id="304F60447FE6F09AD6B23ED87F6E093A" type="ORCID">0000-0002-5768-7216</mods:nameIdentifier>
<mods:affiliation id="A736FDDB8ABC64DA2A61FDB832B98A7C">Invertebrate Zoology, Royal Alberta Museum, 9810 103 A Avenue NW, Edmonton, Alberta, T 5 J 0 G 2, Canada &amp; Department of Veterinary Microbiology, University of Saskatchewan, 52 Campus Drive, Saskatoon, Saskatchewan, S 7 N 5 B 4, Canada</mods:affiliation>
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<mods:part id="94E4C01A22991A3B8DF88F5D76DBBD23">
<mods:date id="CDB940C637AFAA38121FA1A011C64447">2024</mods:date>
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<treatment id="50AEAD430D13552497579AA544512AEB" ID-arpha="50AEAD43-0D13-5524-9757-9AA544512AEB" LSID="urn:lsid:plazi:treatment:50AEAD430D13552497579AA544512AEB" httpUri="http://treatment.plazi.org/id/50AEAD430D13552497579AA544512AEB">
<subSubSection id="50FF54109AE92C950F3A95604E34A678" type="nomenclature">
<paragraph id="D968B7876541AC450436E150F9D36134">
<taxonomicName id="7650CE66442CD362A66A86934454466C" ID-CoL="53H66" authority="(Curran, 1928)" baseAuthorityName="Curran" baseAuthorityYear="1928" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="F916486C002EB654FCD117204B71ECC8" italics="true">Suragina bezzii</emphasis>
(
<bibRefCitation id="9ECAFA75BFC70978CE2E1AD4F71183D4" author="Curran CH" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="167-187" refId="B6" refString="Curran CH (1928) Diptera of the American Museum Congo Expedition. Part III. - Stratiomyidae, Rhagionidae, Therevidae, Scenopinidae, Ortalidae, Micropezidae, Piophilidae, Sepsidae and Diopsidae. Bulletin of the American Museum of Natural History 58: 167 - 187." title="Diptera of the American Museum Congo Expedition. Part III. - Stratiomyidae, Rhagionidae, Therevidae, Scenopinidae, Ortalidae, Micropezidae, Piophilidae, Sepsidae and Diopsidae." volume="58" year="1928">Curran, 1928</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="8C51A4F7F5F124DF29637AB0CD48A42E" type="description">
<paragraph id="27F77205DD8425F1319618BF1C9586C2">
<figureCitation id="7A7DD3811630A031917A81CE235310BD" captionStart="Figures 913" captionStartId="F2" captionText="Figures 913. Suragina Walker spp. lateral habitus: S. agramma (Bezzi): 9 ♂ (NMSA - DIP 028196) 10 ♀ holotype (NMSA - DIP 028199); S. bezzii (Curran): 11 ♂ (SAM-DIP-A 018382) 12 ♀ (SAM-DIP-A 018410); S. bilobata Muller, sp. nov.: 13 a ♀ paratype 1 st flagellomere (CSCA) 13 b ♀ holotype (CSCA). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures9-13" httpUri="https://binary.pensoft.net/fig/1205864">Figs 11</figureCitation>
,
<figureCitation id="7B7387127F6EBAA2BAF3F20858A56B36" captionStart="Figures 913" captionStartId="F2" captionText="Figures 913. Suragina Walker spp. lateral habitus: S. agramma (Bezzi): 9 ♂ (NMSA - DIP 028196) 10 ♀ holotype (NMSA - DIP 028199); S. bezzii (Curran): 11 ♂ (SAM-DIP-A 018382) 12 ♀ (SAM-DIP-A 018410); S. bilobata Muller, sp. nov.: 13 a ♀ paratype 1 st flagellomere (CSCA) 13 b ♀ holotype (CSCA). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures9-13" httpUri="https://binary.pensoft.net/fig/1205864">12</figureCitation>
,
<figureCitation id="F057A9771A2230ED8326DA09EAF09502" captionStart="Figures 3742" captionStartId="F10" captionText="Figures 3742. Suragina Walker spp. wing dorsal view: 37 S. agramma (Bezzi) ♀ (BMSA (D) 124627) 38 S. bezzii (Curran) ♂ (SAM-DIP-A 018410) 39 S. bilobata Muller, sp. nov. ♀ paratype (CSCA) 40 S. copelandi Muller, sp. nov. ♀ paratype (BMSA (D) 83423); S. binominata (Bequaert): 41 holotype ♂ (MLUH) 42 ♀ (NMSA - DIP 158446). 41 Copyright ZNS Halle, January 30, 2024. Abbreviations: br basal radial cell; bm basal medial cell; cua anterior cubital cell; CuA + CuP anterior branch of cubital vein + posterior branch of cubital vein; d discal cell; M 1 first branch of media; m 1 first medial cell; M 2 second branch of media; m 2 second medial cell; M 3 third branch of media; m 3 third medial cell; R 1 anterior branch of radius; r 1 first radial cell; R 2 + 3 second branch of radius; r 2 + 3 second radial cell; R 4 upper branch of third branch of radius; R 5 lower branch of third branch of radius; Sc subcostal vein; sc subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures37-42" httpUri="https://binary.pensoft.net/fig/1205872">38</figureCitation>
,
<figureCitation id="C750B6F38BA5C452603968D87863589D" captionStart="Figures 5562" captionStartId="F13" captionText="Figures 5562. Suragina Walker spp. male terminalia, ventral view (5558) and epandrium with cerci, dorsal view (5962): 55, 59 S. agramma (Bezzi) (ICIPE) 56, 60 S. bezzii (Curran) (SAM-DIP A 018382) 57, 61 S. binominata (Bequaert) (NMSA - DIP 028165) 58, 62 S. copelandi Muller, sp. nov. paratype (BMSA (D) 83425). Abbreviations: aed tn aedeagal tine; cerc cercus; ej apod ejaculatory apodeme; epand epandrium; goncx gonocoxite; goncx apod gonocoxal apodeme; gonst gonostylus; pm paramere; pm apod parameral apodeme; pm sh parameral sheath. Scale bar (applicable to all illustrations): 0.6 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures55-62" httpUri="https://binary.pensoft.net/fig/1205875">57</figureCitation>
,
<figureCitation id="F47E23CF8AE2A439E0F12610F9A6A7A5" captionStart="Figures 5562" captionStartId="F13" captionText="Figures 5562. Suragina Walker spp. male terminalia, ventral view (5558) and epandrium with cerci, dorsal view (5962): 55, 59 S. agramma (Bezzi) (ICIPE) 56, 60 S. bezzii (Curran) (SAM-DIP A 018382) 57, 61 S. binominata (Bequaert) (NMSA - DIP 028165) 58, 62 S. copelandi Muller, sp. nov. paratype (BMSA (D) 83425). Abbreviations: aed tn aedeagal tine; cerc cercus; ej apod ejaculatory apodeme; epand epandrium; goncx gonocoxite; goncx apod gonocoxal apodeme; gonst gonostylus; pm paramere; pm apod parameral apodeme; pm sh parameral sheath. Scale bar (applicable to all illustrations): 0.6 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures55-62" httpUri="https://binary.pensoft.net/fig/1205875">56</figureCitation>
,
<figureCitation id="BBCC130BCB572FCDFD75914DDCDCC97F" captionStart="Figures 5562" captionStartId="F13" captionText="Figures 5562. Suragina Walker spp. male terminalia, ventral view (5558) and epandrium with cerci, dorsal view (5962): 55, 59 S. agramma (Bezzi) (ICIPE) 56, 60 S. bezzii (Curran) (SAM-DIP A 018382) 57, 61 S. binominata (Bequaert) (NMSA - DIP 028165) 58, 62 S. copelandi Muller, sp. nov. paratype (BMSA (D) 83425). Abbreviations: aed tn aedeagal tine; cerc cercus; ej apod ejaculatory apodeme; epand epandrium; goncx gonocoxite; goncx apod gonocoxal apodeme; gonst gonostylus; pm paramere; pm apod parameral apodeme; pm sh parameral sheath. Scale bar (applicable to all illustrations): 0.6 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures55-62" httpUri="https://binary.pensoft.net/fig/1205875">60</figureCitation>
,
<figureCitation id="36E42A569435E9AE689AADAE9D1DB6C7" captionStart="Figures 7587" captionStartId="F16" captionText="Figures 7587. Suragina Walker spp. female genital fork and spermathecae, ventral view: 75 S. agramma (Bezzi) (ICIPE) 76 S. bezzii (Curran) (SAM-DIP- 018403) 77 S. bilobata Muller, sp. nov. (CSCA) 78 S. binominata (Bequaert) (NMSA - DIP 158411) 79 S. copelandi Muller, sp. nov. paratype (BMSA (D) 84686) 80 S. freidbergi Muller, sp. nov. paratype (BMSA (D) 92158) 81 S. liberiaensis Muller, sp. nov. paratype (NMSA - DIP 158442) 82 S. malavaensis Muller, sp. nov. paratype (ICIPE) 83 S. milloti (Séguy) (CSCA) 84 S. monogramma (Bezzi) (BMSA (D) 129988) 85 S. mulanjeensis Muller, sp. nov. paratype (NMSA - DIP 158392) 86 S. semiobscura Muller, sp. nov. paratype (ICIPE) 87 S. zombaensis Muller, sp. nov. paratype (NMSA - DIP 1584719). Scale bar (applicable to all illustrations): 0.1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures75-87" httpUri="https://binary.pensoft.net/fig/1205878">76</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="228DA8F19B27BF6A79E31BFDCAF3BF1A" type="reference_group">
<paragraph id="2F616449CBDC22D3B1F036CB03CE0E5A">
<treatmentCitationGroup id="A5FE5D817CA383A4C5D99981468C45FC">
<treatmentCitation id="10CAB9C974E32D8DBB89764C950A3373" author="Curran CH" page="172" year="1928">
<taxonomicName id="A2E07C1DC94CFADF25A11A0BC8EB1DE5" ID-CoL="8B5JV" authority="Curran, 1928: 172" authorityName="Curran" authorityPageNumber="172" authorityYear="1928" class="Insecta" family="Athericidae" genus="Atrichops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="B550581A5262C48EE718ED809A9E2405" italics="true">Atrichops bezzii</emphasis>
<comment id="01387639D21118C4CBE27B8C02D5D84D">
<bibRefCitation id="6F99B3C4AA393E212A6EAD2C21022EF0" author="Curran CH" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="167-187" refId="B6" refString="Curran CH (1928) Diptera of the American Museum Congo Expedition. Part III. - Stratiomyidae, Rhagionidae, Therevidae, Scenopinidae, Ortalidae, Micropezidae, Piophilidae, Sepsidae and Diopsidae. Bulletin of the American Museum of Natural History 58: 167 - 187." title="Diptera of the American Museum Congo Expedition. Part III. - Stratiomyidae, Rhagionidae, Therevidae, Scenopinidae, Ortalidae, Micropezidae, Piophilidae, Sepsidae and Diopsidae." volume="58" year="1928">Curran, 1928: 172</bibRefCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="44358BDC1F61044F3C14C34249E966F2">
<treatmentCitationGroup id="5349C69D17AAAD85BED9B53C8AE1541B">
<taxonomicName id="42F3C5F9D228BB347A4F6DA6A0992540" ID-CoL="53H66" baseAuthorityName="Curran" baseAuthorityYear="1928" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="FD69F67C199B205B8C91805E29F0E080" italics="true">Suragina bezzii</emphasis>
</taxonomicName>
<comment id="1CF68FF8C6914F53E513D6FE5E7686D1">
:
<treatmentCitation id="197C9DAD1702281912104B02F8BE7577" author="Stuckenberg BR" page="312" year="1980">
<bibRefCitation id="4E372055DDB1F78F5768AB23726CEE83" author="Stuckenberg BR" editor="Crosskey RW" firstAuthor="Stuckenberg" journalOrPublisher="British Museum (Natural History), London" page="312" pagination="312-313" refId="B23" refString="Stuckenberg BR (1980) 23. Family Athericidae. In: Crosskey RW (Ed.) Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, 312313." title="23. Family Athericidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Stuckenberg 1980: 312</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="6DDE0EDC7FF8F17E7DD26DEB88635F6B">
<treatmentCitationGroup id="753B377EC7558A539940AECE1AA6CDBA">
<treatmentCitation id="B6CCAC65DEC2E3927BCF037831354301" author="Brunetti E" page="2" year="1929">
<taxonomicName id="A5443270EA55C9F27F3F84232FA2F57A" ID-CoL="8B5K5" authority="Brunetti, 1929: 2" authorityName="Brunetti" authorityPageNumber="2" authorityYear="1929" class="Insecta" family="Athericidae" genus="Atrichops" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="C34AEE48A341CB9DFCB10D753D19D456" italics="true">Atrichops varicolor</emphasis>
<comment id="615E955465F627F29C6FF0A8F42BA407">
<bibRefCitation id="64C20D6C8CB29E40341C455A217864C1" DOI="10.1080/00222932908673024" author="Brunetti E" issue="19" journalOrPublisher="Annals &amp; Magazine of Natural History" pagination="1-35" refId="B4" refString="Brunetti E (1929) I. - New African Diptera. Annals &amp; Magazine of Natural History 4 (19): 1 - 35. https: // doi. org / 10.1080 / 00222932908673024" title="I. - New African Diptera." volume="4" year="1929">Brunetti, 1929: 2</bibRefCitation>
. syn. nov.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="606D0EC05A5C77348C9178E82DCFD4E6">
<treatmentCitationGroup id="F021C42E6A3C2F549192303235B73A20">
<taxonomicName id="1409D656A64E4A6B9E7397CD6644184E" ID-CoL="53H7G" baseAuthorityName="Brunetti" baseAuthorityYear="1929" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="9C1D6ADD464F43702DBD1E916DF90EE4" italics="true">Suragina varicolor</emphasis>
</taxonomicName>
<comment id="2F97BE7713A531297E31FC665E13D067">
:
<treatmentCitation id="A89C6F6384297B350A825B4CDDBB2FC9" author="Stuckenberg BR" page="313" year="1980">
<bibRefCitation id="F1A06E173E894A3A748EB356480AC04F" author="Stuckenberg BR" editor="Crosskey RW" firstAuthor="Stuckenberg" journalOrPublisher="British Museum (Natural History), London" page="313" pagination="312-313" refId="B23" refString="Stuckenberg BR (1980) 23. Family Athericidae. In: Crosskey RW (Ed.) Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, 312313." title="23. Family Athericidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Stuckenberg 1980: 313</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2EBG" type="Synonymy of Suragina varicolor (Brunetti, 1929)">
<paragraph id="EB1CA06D5901C5736BC2AB5D2157D382">
<heading id="BCEE76026ACAC68E04A456538150143E" reason="title">
Synonymy of
<taxonomicName id="0CE2A1453663585C0D2EA04A58A32DB7" authority="(Brunetti, 1929)" baseAuthorityName="Brunetti" baseAuthorityYear="1929" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="76B9AF60BE144570E8D1ED66E381BB95" italics="true">Suragina varicolor</emphasis>
(
<bibRefCitation id="98E193DEA9C35C1495D24A57F780C6FE" DOI="10.1080/00222932908673024" author="Brunetti E" issue="19" journalOrPublisher="Annals &amp; Magazine of Natural History" pagination="1-35" refId="B4" refString="Brunetti E (1929) I. - New African Diptera. Annals &amp; Magazine of Natural History 4 (19): 1 - 35. https: // doi. org / 10.1080 / 00222932908673024" title="I. - New African Diptera." volume="4" year="1929">Brunetti, 1929</bibRefCitation>
)
</taxonomicName>
.
</heading>
</paragraph>
<paragraph id="C33463206B32884F2C104056239200EE">
Images of both the
<typeStatus id="58FF219582E4AD2EDA1529E59D8A2601" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="0BDB09CBD7F4D57215F0B706B3F29AB9" baseAuthorityName="Curran" baseAuthorityYear="1928" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="38B1F488331BBBF50C27FB83602C238A" italics="true">S. bezzii</emphasis>
</taxonomicName>
and
<taxonomicName id="38AB4FD8DE11B4F028E17FFD6DC5A17D" baseAuthorityName="Brunetti" baseAuthorityYear="1929" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="07F623D0996662AFFDCC25970FFF152F" italics="true">S. varicolor</emphasis>
</taxonomicName>
, as well as identified material from
<collectingCountry id="E4EA955CB03734CD58037E5D6EF0353C" name="Uganda">Uganda</collectingCountry>
,
<collectingCountry id="30C9B3B2CE9CB48A270DD9A8873220AB" name="Kenya">Kenya</collectingCountry>
and
<collectingCountry id="92ED6F04AF178AEA21B1D281453E1037" name="Burundi">Burundi</collectingCountry>
were examined and compared. There is no marked difference between the two species, apart from slight variation in colour intensity due to different preservation methods. Thus,
<taxonomicName id="B6017A0FE3B539499BCB549DED30F201" baseAuthorityName="Brunetti" baseAuthorityYear="1929" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="AEBA457555591EEB44BFA3C98434F527" italics="true">S. varicolor</emphasis>
</taxonomicName>
is hereby designated as junior synonym of
<taxonomicName id="1F5F866BC2832415A9F5FDF5B7437FE5" baseAuthorityName="Curran" baseAuthorityYear="1928" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="6EC778C2B74C91A78B7D90863308ECC9" italics="true">S. bezzii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZGBG" type="type material">
<paragraph id="193E5F830A1AB95DF53B9B1CA8ECBDB6">
<heading id="8DA527D457EDA38E4246F4FB11E5A9AA" reason="title">
<typeStatus id="5D20E43D43AEC4F5C05EEC6A4BC768D0">Type</typeStatus>
material examined.
</heading>
</paragraph>
<paragraph id="F7C8F7CDD7DDBEB5DA7D2AD599F11B2C">
[
<materialsCitation id="55C151E29924E3AC49C0115B998AF43F" country="Democratic Republic of the Congo" location="Democratic Republic of Congo" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
from digital photographs]
<emphasis id="8D370171D7394139D3F09163E1BB2703" bold="true" italics="true">
<typeStatus id="FB142C02251B9E78BFFD47957D54BC65" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="044FB5815E59E15FD3FBD4CEC1649A10" count="1" type="female"></specimenCount>
; [
<collectingCountry id="CAB71E4B747E922C97094DBE01B5FF9E" name="Democratic Republic of the Congo">Democratic Republic of Congo</collectingCountry>
]
</materialsCitation>
<materialsCitation id="59D417783F09FE56788669E86F939C1C" collectingDate="1914-09-29" collectionCode="AMNH" country="Democratic Republic of the Congo" latitude="0.9701666" location="Bengamisa" longLatPrecision="13" longitude="25.210667" specimenCount="1" typeStatus="holotype">
<location id="224A2E23E08B9984A4E059B076E5CDAA" LSID="urn:lsid:plazi:treatment:50AEAD430D13552497579AA544512AEB:224A2E23E08B9984A4E059B076E5CDAA" country="Democratic Republic of the Congo" latitude="0.9701666" longLatPrecision="13" longitude="25.210667" name="Bengamisa">Bengamisa</location>
,
<collectingCountry id="043E54AC8F8237E49442BD85296D47AA" name="Democratic Republic of the Congo">Belgian Congo</collectingCountry>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFHBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[25.210667,0.970167]}">
<geoCoordinate id="480437E2F4E12E68B417385C9E43F97A" degrees="0" direction="north" minutes="58.21" orientation="latitude" precision="9" value="0.9701666">0°58.21'N</geoCoordinate>
,
<geoCoordinate id="FDBD73A8EF953B4E32758EBE41531D6E" degrees="25" direction="east" minutes="12.64" orientation="longitude" precision="9" value="25.210667">25°12.64'E</geoCoordinate>
</named-content>
];
<date id="9438AA0C8A9165007E266835FAD8351E" value="1914-09-29">
<collectingDate id="11F071AC015303790724939974B916B2" value="1914-09-29">29 Sep. 1914</collectingDate>
</date>
; (
<named-content id="B82D8E425AE219FEFC151129E3366953" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/american-museum-natural-history" xlink_title="American Museum of Natural History">
<collectionCode id="A7DE07CEA1147E782C439DF26F690B69" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" type="">AMNH</collectionCode>
</named-content>
). Type photos accessible at:
<ext-link id="C0D523FFDBE9098A7B6E9440FF2DA554" ext-link-type="uri" xlink_href="https://emu-prod.amnh.org/imulive/iz/iz.html#details=ecatalogue.10021491" xlink_type="simple">https://emu-prod.amnh.org/imulive/iz/iz.html#details=ecatalogue.10021491</ext-link>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETHBG" type="type material">
<paragraph id="1A51C0BFD4E744DE3748F73927E89315">
<heading id="12939A67339272FD80B9FE691DB15AD9" reason="title">
Other
<typeStatus id="B5FDA013A004AD23B5DFD9486264000C">type</typeStatus>
material examined.
</heading>
</paragraph>
<paragraph id="2BA47E48E19B077B225BD36B0CD01D0A">
<taxonomicName id="AAA873C99CAFF104C7D60E003085D754" baseAuthorityName="Brunetti" baseAuthorityYear="1929" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="varicolor" status="syn. nov.">
<emphasis id="C089D05D24966206B685D0264533D885" italics="true">S. varicolor</emphasis>
</taxonomicName>
<materialsCitation id="61DC40B9427085E938B669AABDCB0421" collectingDate="1911-08-13" collectionCode="NHMUK" country="Uganda" county="Southern Region" latitude="0.31361166" location="Imperial Bureau of Entomology" longLatPrecision="1" longitude="32.581112" municipality="Presented" specimenCode="NHMUK 014064157" specimenCount="1" specimenCount-female="1" stateProvince="Kampala" typeStatus="holotype">
<taxonomicNameLabel id="9020AEE3031D806A45A1271598ECC0F4">syn. nov.</taxonomicNameLabel>
type [from digital photographs]:
<emphasis id="C4940A0560E145AE6C1B4CD5FF6A853D" bold="true" italics="true">
<typeStatus id="EAB2E07C4FBB6BD533DF197BDD0CB129" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="306751D71A66900ED0F13E5012F5912E" name="Uganda">Uganda</collectingCountry>
<specimenCount id="1385FF254D96F2DCAE493305D117EF59" count="1" type="female">1 ♀</specimenCount>
;
<collectingCounty id="6AA6E77FC1AEDA14541AD52280CBA332">Southern Region</collectingCounty>
;
<collectingRegion id="77E7FCF6F4700C61CFC73E412BF1CCEF" country="Uganda" name="Kampala">Kampala</collectingRegion>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKIBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[32.581112,0.313612]}">
<geoCoordinate id="E8C510A25D28F55393D9950B47BDE859" degrees="0" direction="north" minutes="18.8167" orientation="latitude" precision="1" value="0.31361166">0°18.8167'N</geoCoordinate>
,
<geoCoordinate id="0BFE3002FD9F9D5CED3CD4EC375A92DA" degrees="32" direction="east" minutes="34.8667" orientation="longitude" precision="1" value="32.581112">32°34.8667'E</geoCoordinate>
</named-content>
];
<date id="0660CF09AA1C99082A65E975A3483CE2" value="1911-08-13">
<collectingDate id="8A4F578D8D5AAD91FCA3647C51E7392B" value="1911-08-13">13 Aug. 1911</collectingDate>
</date>
;
<collectingMunicipality id="981043066B6A510FCAE2E8B5EE534DAB">Presented</collectingMunicipality>
by the
<location id="E4CC11C39E06733A7A237FC66DC72573" LSID="urn:lsid:plazi:treatment:50AEAD430D13552497579AA544512AEB:E4CC11C39E06733A7A237FC66DC72573" country="Uganda" county="Southern Region" latitude="0.31361166" longLatPrecision="1" longitude="32.581112" municipality="Presented" name="Imperial Bureau of Entomology" stateProvince="Kampala">Imperial Bureau of Entomology</location>
,
<location id="EF1A5747B21EA2FB15D8501424371E5F" LSID="urn:lsid:plazi:treatment:50AEAD430D13552497579AA544512AEB:EF1A5747B21EA2FB15D8501424371E5F" country="Uganda" county="Southern Region" latitude="0.31361166" longLatPrecision="1" longitude="32.581112" municipality="Presented" name="British Museum" stateProvince="Kampala">British Museum</location>
192948; (
<specimenCode id="518D987935C1B6E854BAEA2A1944709D" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">
<named-content id="CE96B96738F119577DA2F3870716ABE8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">NHMUK</named-content>
014064157
</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETIBG" type="material">
<paragraph id="407A427C31BC051DB829FE42A3089B6D">
<heading id="38F11B5E5E5C7711F18534989E989F58" reason="title">Other material examined.</heading>
</paragraph>
<paragraph id="18B961916301496ED322BA549DFDD226">
<collectingCountry id="339D7B6B9148E13D0576463CAFC0894D" name="Burundi">Burundi</collectingCountry>
<materialsCitation id="BDB4CEEABD34988FAEE805FB1BA88B9C" collectingDate="2010-02-21" collectingDateMax="2010-03-08" collectingDateMin="2010-02-21" collectorName="R. Copeland" country="Kenya" elevation="1382" latitude="-3.1666667" location="Ruvubu River" longLatPrecision="13" longitude="30.333334" municipality="Ruvubu National Park" specimenCount="1" specimenCount-female="1">
<specimenCount id="80DEF9A741C7EE669A14FB5FE80BC395" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="25EC9602A9DA3527CE3B71AD31E0B187">Ruvubu National Park</collectingMunicipality>
, nr.
<location id="2EEC1522E7B59FC48864C1AA79D32514" LSID="urn:lsid:plazi:treatment:50AEAD430D13552497579AA544512AEB:2EEC1522E7B59FC48864C1AA79D32514" country="Kenya" latitude="-3.1666667" longLatPrecision="13" longitude="30.333334" municipality="Ruvubu National Park" name="Ruvubu River">Ruvubu River</location>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3IBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[30.333333,-3.166667]}">
<geoCoordinate id="0874C143A4705BB3C4A82E8346C88797" degrees="3" direction="south" minutes="10.00" orientation="latitude" precision="9" value="-3.1666667">3°10.00'S</geoCoordinate>
,
<geoCoordinate id="2446AB22A1CE4C637A03F6591AA59A5F" degrees="30" direction="east" minutes="20.00" orientation="longitude" precision="9" value="30.333334">30°20.00'E</geoCoordinate>
</named-content>
];
<quantity id="DCA796E46B4994FF3DC71CAF5B4329DE" metricMagnitude="3" metricUnit="m" metricValue="1.382" unit="m" value="1382.0">
<elevation id="6DB1A7774BC557067887216F72B96568" metricMagnitude="3" metricUnit="m" metricValue="1.382" unit="m" value="1382.0">1382 masl</elevation>
</quantity>
;
<date id="EDE8ED1E8503F4E442161459C9ABA117" value="2010-02-21" valueMax="2010-03-08" valueMin="2010-02-21">
<collectingDate id="A699A3030C6F5548605E418B7CEBB167" value="2010-02-21" valueMax="2010-03-08" valueMin="2010-02-21">21 Feb. 8 Mar. 2010</collectingDate>
</date>
;
<collectorName id="BFE68BDC1774B8F548327420003B051E">R. Copeland</collectorName>
leg.; edge of forest;
<collectingMethod id="5B78F0CF3164876FEE676A99C19B4982">Malaise trap</collectingMethod>
.
<collectingCountry id="F5FADF5094B4E9D8F520E7CC15871189" name="Kenya">Kenya</collectingCountry>
</materialsCitation>
<materialsCitation id="9DF7B7C45E9491F7D0167735E072FB33" collectingDate="2006-10-22" collectingDateMax="2006-11-05" collectingDateMin="2006-10-22" collectionCode="ICIPE" collectorName="R. Copeland" country="Kenya" county="Western Province" elevation="1630" latitude="0.22767" location="Rondo Guest House" longLatPrecision="1" longitude="34.88533" municipality="Forest" specimenCount="1" specimenCount-female="1" stateProvince="Kakamega">
<specimenCount id="C433A3647DD26CE97A181BA9AB09577B" count="1" type="female">1 ♀</specimenCount>
;
<collectingCounty id="332253387A0937371D8A245A325E6211">Western Province</collectingCounty>
;
<collectingRegion id="285CE7842949C055E65D4850FE386A3A" country="Kenya" name="Kakamega">Kakamega</collectingRegion>
<collectingMunicipality id="05A1BCA4F78778CA225C888B7DFA42B0">Forest</collectingMunicipality>
, nr.
<location id="208A860E07112B08F326D41B642459C6" LSID="urn:lsid:plazi:treatment:50AEAD430D13552497579AA544512AEB:208A860E07112B08F326D41B642459C6" country="Kenya" county="Western Province" latitude="0.22767" longLatPrecision="1" longitude="34.88533" municipality="Forest" name="Rondo Guest House" stateProvince="Kakamega">Rondo Guest House</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EEJBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[34.885330,0.227670]}">
<geoCoordinate id="65A34846E4B1276D3BA7BDCC830952C7" degrees="00" direction="north" minutes="13.6602" orientation="latitude" precision="1" value="0.22767">00°13.6602'N</geoCoordinate>
,
<geoCoordinate id="A37F7FAF0BDCDE72AB4EE55BBE68DBCC" degrees="34" direction="east" minutes="53.1198" orientation="longitude" precision="1" value="34.88533">34°53.1198'E</geoCoordinate>
</named-content>
;
<quantity id="66547F508860DBF15123C3BE13752977" metricMagnitude="3" metricUnit="m" metricValue="1.63" unit="m" value="1630.0">
<elevation id="CAEF0044D64007D08B2009D4155A11C5" metricMagnitude="3" metricUnit="m" metricValue="1.63" unit="m" value="1630.0">1630 masl</elevation>
</quantity>
;
<date id="B093A18A163C72F577804814CD583BFE" value="2006-10-22" valueMax="2006-11-05" valueMin="2006-10-22">
<collectingDate id="A0F3198ABE25EA2F7A9CE6E9A57590D4" value="2006-10-22" valueMax="2006-11-05" valueMin="2006-10-22">22 Oct. 5 Nov. 2006</collectingDate>
</date>
;
<collectorName id="2314A8E76215EE34365AE380A5CFA364">R. Copeland</collectorName>
leg.; across small permanent stream;
<collectingMethod id="F71AFFB56116DA379C27E20271921C23">Malaise trap</collectingMethod>
; (
<abbrev id="ABBRID0EJJBG" xlink_title="International Centre of Insect Physiology and Ecology">
<collectionCode id="446EDB297C4EE2EFDCE5CF212768D557">ICIPE</collectionCode>
</abbrev>
)
</materialsCitation>
<materialsCitation id="9F1368D6810E82AB503517031DD59F49" collectingDate="2006-09-24" collectingDateMax="2006-10-08" collectingDateMin="2006-09-24" collectionCode="ICIPE" collectorName="R. Copeland" country="Uganda" county="Western Province" elevation="1630" latitude="0.22767" location="Rondo Guest House" longLatPrecision="1" longitude="34.88533" municipality="Forest" specimenCount="1" specimenCount-female="1" stateProvince="Kakamega">
<specimenCount id="AEBF1C50933B57D995BDF446BE86048A" count="1" type="female">1 ♀</specimenCount>
;
<collectingCounty id="D87D46E6F0F428437C207F8D0D3BF951">Western Province</collectingCounty>
;
<collectingRegion id="C543B3B1AEA4D37E24867D43578C7979" country="Kenya" name="Kakamega">Kakamega</collectingRegion>
<collectingMunicipality id="2E9EBF15F45663DD0756FFB4D72A00FD">Forest</collectingMunicipality>
, nr.
<location id="5D965F7A98035DA94712F42A8732E025" LSID="urn:lsid:plazi:treatment:50AEAD430D13552497579AA544512AEB:5D965F7A98035DA94712F42A8732E025" country="Uganda" county="Western Province" latitude="0.22767" longLatPrecision="1" longitude="34.88533" municipality="Forest" name="Rondo Guest House" stateProvince="Kakamega">Rondo Guest House</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQJBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[34.885330,0.227670]}">
<geoCoordinate id="F9221193C2561FC920B624A5F9AE588B" degrees="00" direction="north" minutes="13.6602" orientation="latitude" precision="1" value="0.22767">00°13.6602'N</geoCoordinate>
,
<geoCoordinate id="9520163916B6DDF587018D624A454A5E" degrees="34" direction="east" minutes="53.1198" orientation="longitude" precision="1" value="34.88533">34°53.1198'E</geoCoordinate>
</named-content>
;
<quantity id="8431EF06C2F8333A492194A1F1B32E92" metricMagnitude="3" metricUnit="m" metricValue="1.63" unit="m" value="1630.0">
<elevation id="D8D066C827CE2CA9FC31C433CAB8AE3A" metricMagnitude="3" metricUnit="m" metricValue="1.63" unit="m" value="1630.0">1630 masl</elevation>
</quantity>
;
<date id="CB8FD82E48C9B3CF8988BD4D14819DCF" value="2006-09-24" valueMax="2006-10-08" valueMin="2006-09-24">
<collectingDate id="542E55F04DF1950D50B27E0B78C76035" value="2006-09-24" valueMax="2006-10-08" valueMin="2006-09-24">24 Sep. 8 Oct. 2006</collectingDate>
</date>
;
<collectorName id="65A91A3591EC71214E8F5B551E97DC0A">R. Copeland</collectorName>
leg.; across small permanent stream;
<collectingMethod id="A85DFC63053CAF54C1CF8B432694CAEF">Malaise trap</collectingMethod>
; (
<abbrev id="ABBRID0EVJBG" xlink_title="International Centre of Insect Physiology and Ecology">
<collectionCode id="F7FC0B3D2C9733EE536D9AC68F6D84D2">ICIPE</collectionCode>
</abbrev>
).
<collectingCountry id="B8A8837BF4F935A2921B81B416855375" name="Uganda">Uganda</collectingCountry>
</materialsCitation>
<materialsCitation id="7167ADE6554139F597F4E3C3E40FE40B" collectingDate="2008-10-12" collectingDateMax="2008-10-26" collectingDateMin="2008-10-12" collectorName="S van Noort &amp; Rainforest" country="Uganda" county="National Park" elevation="1505" latitude="0.5637167" location="Makerere University Biological Field Station" longLatPrecision="1" longitude="30.358166" municipality="Kanyawara" specimenCount="51" specimenCount-female="49" specimenCount-male="2" stateProvince="Kibale">
<specimenCount id="6571BEA6A1D549685B8D99EC7018C575" count="1" type="male">1 ♂</specimenCount>
<specimenCount id="EF5BA96DEA00764940954D5D9C0B3613" count="48" type="female">48 ♀</specimenCount>
;
<collectingRegion id="B993A6114399F2210983922D6D645473" country="Uganda" name="Kibale">Kibale</collectingRegion>
<collectingCounty id="D85C2EB4C7538C26C568762CC2C1C759">National Park</collectingCounty>
,
<collectingMunicipality id="42CFFA45AC882312F47ED2478A8570BE">Kanyawara</collectingMunicipality>
,
<location id="FCF428C4F301EE96E6C12E6732049673" LSID="urn:lsid:plazi:treatment:50AEAD430D13552497579AA544512AEB:FCF428C4F301EE96E6C12E6732049673" country="Uganda" county="National Park" latitude="0.5637167" longLatPrecision="1" longitude="30.358166" municipality="Kanyawara" name="Makerere University Biological Field Station" stateProvince="Kibale">Makerere University Biological Field Station</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3JBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[30.358167,0.563717]}">
<geoCoordinate id="390A9EC276E1EA248A84F2B320119206" degrees="00" direction="north" minutes="33.823" orientation="latitude" precision="1" value="0.5637167">00°33.823'N</geoCoordinate>
,
<geoCoordinate id="888A9638CF1B7E730523D384E1E02A46" degrees="30" direction="east" minutes="21.490" orientation="longitude" precision="1" value="30.358166">30°21.490'E</geoCoordinate>
</named-content>
;
<quantity id="8B3E295042526546C92E69F09C325805" metricMagnitude="3" metricUnit="m" metricValue="1.505" unit="m" value="1505.0">
<elevation id="CCECB47577DC73435DADE4C008023309" metricMagnitude="3" metricUnit="m" metricValue="1.505" unit="m" value="1505.0">1505 masl</elevation>
</quantity>
;
<date id="50A48AF56D240258D60242F79D2DA926" value="2008-10-12" valueMax="2008-10-26" valueMin="2008-10-12">
<collectingDate id="C09437E612C5520E1910D608BFC69FC5" value="2008-10-12" valueMax="2008-10-26" valueMin="2008-10-12">1226 Oct. 2008</collectingDate>
</date>
;
<collectorName id="8C0C514FA26F7A5AE7D8468E0353416F">S van Noort</collectorName>
leg.; UG 08 - KF 3 - M 13;
<collectingMethod id="4270B98325299765292CA1CFD8E90A10">Malaise trap</collectingMethod>
; primary mid-altitude
<collectorName id="4DFF482EBFE0472F203B627C14834E57">Rainforest</collectorName>
(
<specimenCount id="3B320DBE44A97003E1BC8FCB28549300" count="1" type="male"></specimenCount>
: SAM-DIP-A 018410;
<specimenCount id="8BD778DE2ED2845337F8E360A7A163D3" count="1" type="female"></specimenCount>
: SAM-DIP-A 018378, A 018379, A 018380, A 018381, A 018383, A 018384, A 018386, A 018387, A 018388, A 018389, A 018391, A 018392, A 018393, A 018394, A 018395, A 018396, A 018397, A 018398, A 018399, A 018400, A 018401, A 018402, A 018403, A 018404, A 018405, A 018406, A 018407, A 018408, A 018409, A 018411, A 018412, A 018413, A 018414, A 018416, A 018417, A 018418, A 018419, A 018420, A 018421, A 018422, A 018423, A 018424, A 018425, A 018426, A 018427, A 018428, A 018429, A 018432)
</materialsCitation>
.
</paragraph>
<paragraph id="46ED593BBBADD6FF1A086E76AA5F02DB">
The female material from
<collectingRegion id="80ADB46B470AA426A4BF9D955E418512" country="Uganda" name="Kibale">Kibale</collectingRegion>
National Park agrees entirely with that of the original type description as well as with the publically available photos of the
<typeStatus id="772E2D24D8F0A6432CD65CF4907DBCCC" type="holotype">holotype</typeStatus>
female.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECKBG" type="diagnosis">
<paragraph id="265277A0D4B18C0AEBD3424577DF29D7">
<heading id="F323623CFC3A00D8A7C4508BADD12F54" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="90D552C205AB7B7EEE00AFAD49EC44E8">
<taxonomicName id="7CA4367D5752130FD9AE4A49EBA0D778" baseAuthorityName="Curran" baseAuthorityYear="1928" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="38A4BAF549C8A5F119CCBCD59E2B5A76" italics="true">Suragina bezzii</emphasis>
</taxonomicName>
has an overall dark appearance (Figs
<figureCitation id="DEACD3A20EE9C3B2500ED3B6B76E9BD1" captionStart="Figures 913" captionStartId="F2" captionText="Figures 913. Suragina Walker spp. lateral habitus: S. agramma (Bezzi): 9 ♂ (NMSA - DIP 028196) 10 ♀ holotype (NMSA - DIP 028199); S. bezzii (Curran): 11 ♂ (SAM-DIP-A 018382) 12 ♀ (SAM-DIP-A 018410); S. bilobata Muller, sp. nov.: 13 a ♀ paratype 1 st flagellomere (CSCA) 13 b ♀ holotype (CSCA). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures9-13" httpUri="https://binary.pensoft.net/fig/1205864">11</figureCitation>
,
<figureCitation id="9DF9FC4199F25703A65F151C23302CE9" captionStart="Figures 913" captionStartId="F2" captionText="Figures 913. Suragina Walker spp. lateral habitus: S. agramma (Bezzi): 9 ♂ (NMSA - DIP 028196) 10 ♀ holotype (NMSA - DIP 028199); S. bezzii (Curran): 11 ♂ (SAM-DIP-A 018382) 12 ♀ (SAM-DIP-A 018410); S. bilobata Muller, sp. nov.: 13 a ♀ paratype 1 st flagellomere (CSCA) 13 b ♀ holotype (CSCA). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures9-13" httpUri="https://binary.pensoft.net/fig/1205864">12</figureCitation>
), with its entire thorax dark brown to blackish with varying levels of bluish-grey pruinosity throughout. The wings are light brown suffused with a dark brown stigma and a brown suffused substigmal marking (Fig.
<figureCitation id="818053AEE5E2AD7267B171C42D838E60" captionStart="Figures 3742" captionStartId="F10" captionText="Figures 3742. Suragina Walker spp. wing dorsal view: 37 S. agramma (Bezzi) ♀ (BMSA (D) 124627) 38 S. bezzii (Curran) ♂ (SAM-DIP-A 018410) 39 S. bilobata Muller, sp. nov. ♀ paratype (CSCA) 40 S. copelandi Muller, sp. nov. ♀ paratype (BMSA (D) 83423); S. binominata (Bequaert): 41 holotype ♂ (MLUH) 42 ♀ (NMSA - DIP 158446). 41 Copyright ZNS Halle, January 30, 2024. Abbreviations: br basal radial cell; bm basal medial cell; cua anterior cubital cell; CuA + CuP anterior branch of cubital vein + posterior branch of cubital vein; d discal cell; M 1 first branch of media; m 1 first medial cell; M 2 second branch of media; m 2 second medial cell; M 3 third branch of media; m 3 third medial cell; R 1 anterior branch of radius; r 1 first radial cell; R 2 + 3 second branch of radius; r 2 + 3 second radial cell; R 4 upper branch of third branch of radius; R 5 lower branch of third branch of radius; Sc subcostal vein; sc subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures37-42" httpUri="https://binary.pensoft.net/fig/1205872">38</figureCitation>
). The abdomen has tergite 2 with a fenestrated appearance and the tergites 46 orange, contrasting with the otherwise dark brown abdomen. The species is most similar to
<taxonomicName id="7400D5D4B7C28CD26A64BEF0C8DCEEEE" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="semiobscura" status="sp. nov.">
<emphasis id="0506E61F96B17FDA830619CD34C64EEE" italics="true">S. semiobscura</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="82B2A04B30921C6072DED278F75391F9" rank="species">sp. nov.</taxonomicNameLabel>
but differs from it by having the postpronotal lobes dark brown as opposed to pale yellow.
<taxonomicName id="3E14F67259C50D9306986903863D05D0" authority="Muller" authorityName="Muller" authorityYear="2024" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="semiobscura" status="sp. nov.">
<emphasis id="E54BBAE0B395B530301951E9A848DE2C" italics="true">Suragina semiobscura</emphasis>
Muller
</taxonomicName>
,
<taxonomicNameLabel id="980E8B04E2BB97852AFDFAF1E407A9D6" rank="species">sp. nov.</taxonomicNameLabel>
, most strikingly, has the upper half of the wing dark brown suffused, the lower half appearing lighter, creating a two-toned appearance (Fig.
<figureCitation id="9B7EAABFB3E08112AB89BF930625F9E0" captionStart="Figures 4954" captionStartId="F12" captionText="Figures 4954. Suragina Walker spp. wing dorsal view: 49 S. monogramma (Bezzi) ♀ (BMSA (D) 129988) 50 S. mulanjeensis Muller, sp. nov. ♀ (BMSA (D) 92374) 51 S. nigromaculata Muller, sp. nov. ♂ holotype (NHMUK 014064156) 52 S. pauliani ♂ paratype (NMSA - DIP 028172) 53 S. semiobscura Muller, sp. nov. ♀ paratype (ICIPE) 54 S. zombaensis Muller, sp. nov. ♀ paratype (NMSA - DIP 158451). 51 Copyright NHMUK under CC BY 4.0. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures49-54" httpUri="https://binary.pensoft.net/fig/1205874">53</figureCitation>
) compared to that of
<taxonomicName id="C7E73D5220D7FD214E67F5BE13B34E60" baseAuthorityName="Curran" baseAuthorityYear="1928" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="F52F6983B20179F1E16F2803BBBFB687" italics="true">S. bezzii</emphasis>
</taxonomicName>
that has the wing uniformly brown suffused.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEMBG" type="remarks">
<paragraph id="5BB1A6FAF5871358EB27BBE233FEBB93">
<heading id="EFE9B039D97F7A3C1F6F7ED6CF470C1B" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="3C8C967698AD8FF72B4DC4A6B9562FD1">
<bibRefCitation id="EE2F48F63C6F65E9E54633A0F734215E" author="Curran CH" firstAuthor="Curran" journalOrPublisher="Bulletin of the American Museum of Natural History" page="172" pagination="167-187" refId="B6" refString="Curran CH (1928) Diptera of the American Museum Congo Expedition. Part III. Stratiomyidae, Rhagionidae, Therevidae, Scenopinidae, Ortalidae, Micropezidae, Piophilidae, Sepsidae and Diopsidae. Bulletin of the American Museum of Natural History 58: 167187." title="Diptera of the American Museum Congo Expedition. Part III. - Stratiomyidae, Rhagionidae, Therevidae, Scenopinidae, Ortalidae, Micropezidae, Piophilidae, Sepsidae and Diopsidae." volume="58" year="1928">Curran (1928: 172)</bibRefCitation>
remarked that it was possible that the “ black undescribed ” specimen mentioned by
<bibRefCitation id="9E47584D48290092859A1726EDBE1EFF" author="Bezzi M" firstAuthor="Bezzi" journalOrPublisher="Annals of the South African Museum" page="304" pagination="297-324" refId="B3" refString="Bezzi M (1926) South African Rhagionidae (Diptera) in the South African Museum. Annals of the South African Museum 23: 297324. https://biostor.org/reference/128969 [Last accessed: 2024 / 10 / 29]" title="South African Rhagionidae (Diptera) in the South African Museum." url="https://biostor.org/reference/128969" volume="23" year="1926">Bezzi (1926: 304)</bibRefCitation>
is in fact
<taxonomicName id="EB5F2B4143E667C5D1CA20F8645A1ADA" baseAuthorityName="Curran" baseAuthorityYear="1928" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="bezzii">
<emphasis id="FCF94D867A0ADC897F55D7B9D81476A2" italics="true">S. bezzii</emphasis>
</taxonomicName>
. That is however just conjecture at this point in time as no further information is available regarding the whereabouts of the specimen mentioned by Bezzi, nor was it ever described. There are several dark species from west and central African countries. Bezzis comment: “ This last species, like
<taxonomicName id="3F01E36629339061863D8B8BFCEE9A6F" authority="Speiser, 1914" authorityName="Speiser" authorityYear="1914" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="disciclara">
<emphasis id="5016475247B513B26E86D0EB6AB16E44" italics="true">A. disciclara</emphasis>
<bibRefCitation id="12FC410C760FF99AD6426653ACCBD68E" DOI="10.1002/mmnd.191419140102" author="Speiser P" issue="1" journalOrPublisher="Deutsche Entomologische Zeitschrift" pagination="1-16" refId="B20" refString="Speiser P (1914) Beitrage zur Dipterenfauna von Kamerun II. Deutsche Entomologische Zeitschrift 1914 (1): 1 - 16. https: // doi. org / 10.1002 / mmnd. 191419140102" title="Beiträge zur Dipterenfauna von Kamerun II." volume="1914" year="1914">Speiser, 1914</bibRefCitation>
</taxonomicName>
, shows a deep longitudinal median furrow on the fore part of the frons. ” is not diagnostic, as all known Afrotropical
<taxonomicName id="EF6095F491B0FFA940E57601AC5EFFD1" authorityName="Walker" authorityYear="1859" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="3EA909DA044DAE6D9022D0294582595B" italics="true">Suragina</emphasis>
</taxonomicName>
have a longitudinal median groove or “ furrow ” running up from between the antennal bases, with varying degrees of depth.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPNBG" type="description">
<paragraph id="B7EBC3A8FCA24B1B556FFD6D156E5D33">
<heading id="6CC21A44AA84D21289E887415DB964CF" reason="title">Description.</heading>
</paragraph>
<paragraph id="4D418AAE8048545D93E809CA32632486">
<emphasis id="166C09C06D19E02312AAFD4A490E5093" bold="true" italics="true">Measurements</emphasis>
(
<specimenCount id="CBC97ABA549207F575CE066DDF0E54A6" count="1" type="male"></specimenCount>
n = 1,
<specimenCount id="029FAFBB2D1D9A19A2D9DB79E9211DB7" count="1" type="female"></specimenCount>
n = 5): Wing span:
<specimenCount id="B407CBDD0FCC0E021FC9DEC201D962F0" count="1" type="male"></specimenCount>
<quantity id="5E47823916B492EDB66A0F2BEC6592A9" metricMagnitude="-3" metricUnit="m" metricValue="7.4" unit="mm" value="7.4">7.4 mm</quantity>
;
<specimenCount id="EF40DECCE6285FBD032D1D4A90555A71" count="1" type="female"></specimenCount>
<quantity id="8102E3B3DA41AA5B7040B5517C2D44F2" metricMagnitude="-3" metricUnit="m" metricValue="8.85" metricValueMax="9.1" metricValueMin="8.6" unit="mm" value="8.85" valueMax="9.1" valueMin="8.6">8.69.1 mm</quantity>
(avg.
<quantity id="12E8666986544BE3E4E4EFED2CD676A1" metricMagnitude="-3" metricUnit="m" metricValue="8.4" unit="mm" value="8.4">8.4 mm</quantity>
); body length:
<specimenCount id="7D0E164F4F60DE99618BF125387BB190" count="1" type="male"></specimenCount>
<quantity id="07F0A28E29674DF5CABD9B2B7003593F" metricMagnitude="-3" metricUnit="m" metricValue="9.3" unit="mm" value="9.3">9.3 mm</quantity>
;
<specimenCount id="B5448D3BD5701396A3E1B87E4025D6A7" count="1" type="female"></specimenCount>
9.0
<quantity id="61BE7CCB2D9146CE0002B339F73A63BA" metricMagnitude="-3" metricUnit="m" metricValue="9.8" unit="mm" value="9.8">9.8 mm</quantity>
(avg.
<quantity id="EC12F223BFEC5827DDCC9ABA6324AFEB" metricMagnitude="-3" metricUnit="m" metricValue="9.5" unit="mm" value="9.5">9.5 mm</quantity>
); wing span to body length ratio (avg.):
<specimenCount id="1187524A71F8E3D4367FA7401EA9AB12" count="1" type="male"></specimenCount>
0.80;
<specimenCount id="F9500042C6618C7F6B65DA2D637FB34A" count="1" type="female"></specimenCount>
0.94.
</paragraph>
<paragraph id="3D1881CFF219C5B0EC97E59FB7382AE8">
<emphasis id="CECF61F45D5B42CF55D10CD76B332D79" bold="true">Male</emphasis>
(Fig.
<figureCitation id="B52CEC31877855973B6234207B40B6F7" captionStart="Figures 913" captionStartId="F2" captionText="Figures 913. Suragina Walker spp. lateral habitus: S. agramma (Bezzi): 9 ♂ (NMSA - DIP 028196) 10 ♀ holotype (NMSA - DIP 028199); S. bezzii (Curran): 11 ♂ (SAM-DIP-A 018382) 12 ♀ (SAM-DIP-A 018410); S. bilobata Muller, sp. nov.: 13 a ♀ paratype 1 st flagellomere (CSCA) 13 b ♀ holotype (CSCA). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures9-13" httpUri="https://binary.pensoft.net/fig/1205864">11</figureCitation>
).
</paragraph>
<paragraph id="6CB5EB8E24617ADFBF776E4A245B014D">
<emphasis id="78859BF854ACB09F02C5053D5D251879" bold="true" italics="true">Head</emphasis>
: Black ground colour, with bluish-grey pruinosity on majority of head; eye bare; holoptic; ommatidia of similar size; lateral edge of eye with slight indentation; ocellar tubercle barely visible in profile, black in colour (appears to be rubbed bare in both sexes due to degradation from preservation method); vertex with slight grey pruinosity, otherwise appearing black in colour and dark setulose; anterior ocellus similar in size to posterior pair; ocellar tubercle in front of dorsal margin of eye, not placed as deeply towards middle of head as in
<specimenCount id="84AC490C0B1B8A908087BE479FD5531D" count="1" type="female"></specimenCount>
; dorsal inner edge of eye abutting ocellar tubercle; occiput with same bluish-grey pruinosity as rest of head; paired large rectangular black markings with dark setulae on upper occiput widening towards lateral margin of head, abutting posterior margin of eyes, flanking vertex; upper occiput with similar dark setulae; lower occiput with long pale setulae medially, dark setulae laterally; genal area bluish-grey with dark setulae, these dark setulae continue ventrally on head to mouthparts that have similar dark ventral setulae; frons bluish-grey pruinose, velvety-black from ocellar tubercle to lower half of eye; frons widening from velvety-black patch towards antennal base; frons bare; face with pale setulae; clypeus black with bluish-grey pruinosity, bare; face separated anteriorly from clypeus by a deep transverse suture, similar to lateral sutures; face not appearing to bulge laterally when viewed in profile; clypeus visible in profile, face not; antennal bases separated ca 0.5 × width of scape, with slight longitudinal groove running between; scape, pedicel dark brown with whitish pruinosity; 1
<superScript id="50F8FFA48502C48DBE1D8196E6620BC6">st</superScript>
flagellomere orange, with some slight darker markings and with similar pruinosity as other segments, 2
<superScript id="00EB6A58EC6FC59F75FAC71D41D9CF42">nd</superScript>
flagellomere dark brown; scape and pedicel of similar size; 1
<superScript id="72AB0138E65200E89B9326FBC82138EF">st</superScript>
flagellomere reniform, ca 1.5 × size of pedicel; 2
<superScript id="8A4E4C4A39EBD3E21B58810E19192581">nd</superScript>
flagellomere arista-like; pedicel with dark dorsal and ventral setulae, similar in size, scape with only dark dorsal setulae; palpus black on majority of surface with scattered white pruinosity, orange-yellow on basal third, with dark setulae throughout; palpus ca 0.5 × length of proboscis; proboscis ca same length as head height; proboscis mostly brown, basally more orange-yellow, entire structure interspersed with some dark setulae and some shorter pale setulae.
</paragraph>
<paragraph id="1E0E51C269D6066F076DED726B9F8573">
<emphasis id="9316F8398E123E5377285488C80ED51C" bold="true" italics="true">Thorax</emphasis>
: Scutum rubbed bare due to degradation from preservation method (
<specimenCount id="5876EE83249CAD5BD0C469688332D5DB" count="1" type="female"></specimenCount>
with two feint dorsocentral bluish-grey pruinose vittae running from pronotum and joining before scutellum), otherwise dark brown with bluish-grey pruinosity; pronotum bluish-grey pruinose with long pale setulae; postpronotal lobe same dark brown as rest of scutum, slight bluish-grey pruinose, setation unknown, anterolateral margin of lobe lighter yellowish- to orange-brown (
<specimenCount id="30AF038D76B08F84F345E8303957874C" count="1" type="female"></specimenCount>
colouring more apparent); notopleuron appearing slight bluish-grey pruinose when viewed in profile, however, when viewed dorsally pruinosity shifts into a silver-grey pruinose appearance, running up towards edge of mesonotum (notopleuron mostly rubbed bare except for anterior edge with long dark setulae); postalar wall and postalar callus dark brown with slight bluish-grey pruinosity, anterior of postalar callus orange-yellow; scutellum dark brown with bluish-grey pruinosity, entire margin orange-yellow from base to apex; supra-alar area, postalar wall and postalar callus with patches of dark setulae (less apparent and numerous than in
<specimenCount id="9E101EBC30FC359328DB8BFD5F6890CB" count="1" type="female"></specimenCount>
); majority of pleura bluish-grey pruinose, except for anterior of anepimeron and meron shiny blackish-brown; all pleura that are bluish-grey pruinose have pale setulae (only some remnant setulae remain due to damage from preservation method), except for katatergite that has long dark setulae; anatergite and meron bare; proepisternum and proepimeron with long pale setulae; anterior and posterior spiracles whitish-yellow and surroundings yellowish-brown, bare; postspiracular scale dark brown.
</paragraph>
<paragraph id="B93966367F15A9BE15C2E29B499E8BD8">
<emphasis id="F1E9355CFC8876BEA2935CB41D7CCF3A" bold="true" italics="true">Legs</emphasis>
: All coxae entirely blackish-brown with only some scattered white pruinosity; fore coxa with mostly pale setulae except for some dark setulae apically; mid coxa with long dark setulae on anterior surface, posterior margin appearing bare; hind coxa with a mix of long pale and dark setulae on anterior edge surrounding well-developed anterior apical point, lateral apical edges with long dark setulae; all trochanters reddish dark brown, all trochanters with short pale setulae; fore femur almost entirely yellow, except for blackish-brown apex and base; mid femur missing; hind femur dark brown except for yellow basal and apical quarters; fore tibia and 1
<superScript id="D6153BB41A85E1A70D99A3DEECB82465">st</superScript>
and 2
<superScript id="5DEB8AC1F7CCAFD2C79A4B56052000D2">nd</superScript>
tarsal segments dark brown almost black, rest of for tarsi missing; mid tibia and tarsi missing; hind tibia and basitarsus blackish-brown, with base and apex of hind tibia yellowish-brown, rest of hind tarsi missing; fore and hind femora rubbed clean, setation unknown; fore tibia with short dark setulae; hind tibia with dark setulae that are at least as long as segment is wide; hind leg overall stouter than remaining legs; combined length of hind tarsal segments subequal to hind tibia; tibial spur formula 0: 2: 2; fore tibial spur weaker than that of hind tibia.
</paragraph>
<paragraph id="119C5A0B426CF2C161D46478085E8C73">
<emphasis id="B65E23922755D80ED59A39D66E3BD6E9" bold="true" italics="true">Wing</emphasis>
(Fig.
<figureCitation id="871FC6A019BA10A0B3F62C0FC6542125" captionStart="Figures 3742" captionStartId="F10" captionText="Figures 3742. Suragina Walker spp. wing dorsal view: 37 S. agramma (Bezzi) ♀ (BMSA (D) 124627) 38 S. bezzii (Curran) ♂ (SAM-DIP-A 018410) 39 S. bilobata Muller, sp. nov. ♀ paratype (CSCA) 40 S. copelandi Muller, sp. nov. ♀ paratype (BMSA (D) 83423); S. binominata (Bequaert): 41 holotype ♂ (MLUH) 42 ♀ (NMSA - DIP 158446). 41 Copyright ZNS Halle, January 30, 2024. Abbreviations: br basal radial cell; bm basal medial cell; cua anterior cubital cell; CuA + CuP anterior branch of cubital vein + posterior branch of cubital vein; d discal cell; M 1 first branch of media; m 1 first medial cell; M 2 second branch of media; m 2 second medial cell; M 3 third branch of media; m 3 third medial cell; R 1 anterior branch of radius; r 1 first radial cell; R 2 + 3 second branch of radius; r 2 + 3 second radial cell; R 4 upper branch of third branch of radius; R 5 lower branch of third branch of radius; Sc subcostal vein; sc subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures37-42" httpUri="https://binary.pensoft.net/fig/1205872">38</figureCitation>
): Light brown suffused on majority of surface except for cell
<emphasis id="CDFEDD0F717A58CE9EEBE3B26A55062B" italics="true">cua</emphasis>
and anal lobe; dark brown stigma over cell
<emphasis id="D9976D23D2AF5B828E36A5B67501F239" italics="true">
r
<subScript id="44A3163BCDB8AC91ACB8E42C7EAEE9B9">1</subScript>
</emphasis>
; darker suffused substigmal marking running down from stigma over crossvein
<emphasis id="94DFBAFDF7E139C23DE63FBA1406AD3D" italics="true">r m</emphasis>
, bases of discal cell, cell
<emphasis id="9A8EE1EF95FF79368DEC215A8ABF5E5E" italics="true">
m
<subScript id="ABF830226FB82EB3071B23A1A9E7F756">3</subScript>
</emphasis>
and apex of cell
<emphasis id="EC4DD90B028990388F4A9C91C6235461" italics="true">br</emphasis>
; veins dark brown, with additional darker brown suffusion around vein
<emphasis id="B3D57A3B2B578EF9D905DFD7FDFB95E1" italics="true">CuA</emphasis>
; costa without distinct downward flexure over stigma; cell
<emphasis id="E3907CE7E4DC4A252C1A785DD351E89B" italics="true">cua</emphasis>
closed at wing margin, cell
<emphasis id="BBE2E3E21FD98128A5D646011B1C7E3E" italics="true">
m
<subScript id="97AE0BE5F0F6EA8AECD7D12DF29ED250">3</subScript>
</emphasis>
open, veins
<emphasis id="2D6529779FCBC74E4553BDBE6B961D0A" italics="true">
M
<subScript id="2175E0FF96AB615C6DF5EA58A219BD0A">1</subScript>
</emphasis>
,
<emphasis id="E694693AE9290164EBC0ECA0FE63F45E" italics="true">
M
<subScript id="E221E42E7D2F32DDC1AA0FDCD1193919">2</subScript>
</emphasis>
,
<emphasis id="45F262A70A36D41406C7239952E9904B" italics="true">
M
<subScript id="367CFE96215970323C8C617056F9B45A">3</subScript>
</emphasis>
present; haltere stalk dirty yellow, knob darker yellowish-brown, with a few short and dark setulae.
</paragraph>
<paragraph id="DD079BF2CE520B37061981E6143E51FF">
<emphasis id="A9720137C4AAB57AC70C122EB5AD4A2D" bold="true" italics="true">Abdomen</emphasis>
: Tergite 1 entirely blackish-brown, with dark setulae (pale setulae in
<specimenCount id="04C79FCCC311002F1BF7436CE31C4F4A" count="1" type="female"></specimenCount>
); tergite 2 pale, with dark brown median marking, giving it a fenestrated appearance; tergite 3 dark brown, tergites 4 and 5 orange, with tergite 5 with dark brown posterior band; tergite 6 with orange markings anteriorly, otherwise dark brown; remaining tergites dark brown; all tergites rubbed bare except for tergite 1; tergite 1 medially without a longitudinal suture; sternites 13 cream coloured, sternite 3 with posterior margin brown; rest of sternites orange-yellow; all sternites with pale setulae.
</paragraph>
<paragraph id="29EC858C745BF9F3565AE1FC05C5F5AE">
<emphasis id="79E2672503B34D6788DF9DB7C221A1AB" bold="true" italics="true">Terminalia</emphasis>
(Figs
<figureCitation id="845D0F565A22183F13A1655563DF5771" captionStart="Figures 5562" captionStartId="F13" captionText="Figures 5562. Suragina Walker spp. male terminalia, ventral view (5558) and epandrium with cerci, dorsal view (5962): 55, 59 S. agramma (Bezzi) (ICIPE) 56, 60 S. bezzii (Curran) (SAM-DIP A 018382) 57, 61 S. binominata (Bequaert) (NMSA - DIP 028165) 58, 62 S. copelandi Muller, sp. nov. paratype (BMSA (D) 83425). Abbreviations: aed tn aedeagal tine; cerc cercus; ej apod ejaculatory apodeme; epand epandrium; goncx gonocoxite; goncx apod gonocoxal apodeme; gonst gonostylus; pm paramere; pm apod parameral apodeme; pm sh parameral sheath. Scale bar (applicable to all illustrations): 0.6 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures55-62" httpUri="https://binary.pensoft.net/fig/1205875">56</figureCitation>
,
<figureCitation id="85EB8D585B1FF77D5D8B2ED3A8E4F6A8" captionStart="Figures 5562" captionStartId="F13" captionText="Figures 5562. Suragina Walker spp. male terminalia, ventral view (5558) and epandrium with cerci, dorsal view (5962): 55, 59 S. agramma (Bezzi) (ICIPE) 56, 60 S. bezzii (Curran) (SAM-DIP A 018382) 57, 61 S. binominata (Bequaert) (NMSA - DIP 028165) 58, 62 S. copelandi Muller, sp. nov. paratype (BMSA (D) 83425). Abbreviations: aed tn aedeagal tine; cerc cercus; ej apod ejaculatory apodeme; epand epandrium; goncx gonocoxite; goncx apod gonocoxal apodeme; gonst gonostylus; pm paramere; pm apod parameral apodeme; pm sh parameral sheath. Scale bar (applicable to all illustrations): 0.6 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures55-62" httpUri="https://binary.pensoft.net/fig/1205875">60</figureCitation>
): Epandrium and cercus dark brown with dark setulae; gonocoxite, hypoproct and hypandrium with pale setulae; gonostylus tapering with truncated apex, outer edge of gonostylus with short setulae, inner edge with a protrusion with 4 setulae, apical third of gonostylus with microtrichia; gonocoxite widening and appearing more rounded on apical half, apex somewhat flattened, gonocoxite outer and ventral surface down to parameral apodeme with long setulae, inner surface of upper half with short setulae, lower half with long inward-facing setulae; parameral apodeme well-developed, not reaching base of gonocoxite in ventral view, parameral sheath including parameral apodeme ca 0.7 × length of gonocoxite; gonocoxal apodeme about same length as gonocoxite and slightly shorter than ejaculatory apodeme; aedeagal tine curvature extending down past gonocoxites; endoaedeagal process apically truncated and widened.
</paragraph>
<paragraph id="8847B38104834564E4DEBB13754066A9">
<emphasis id="762E160AC00CA22624408C957A5E8BA2" bold="true">Female</emphasis>
(Fig.
<figureCitation id="9FEA65DA2C2B4F01D625A5764E68B52A" captionStart="Figures 913" captionStartId="F2" captionText="Figures 913. Suragina Walker spp. lateral habitus: S. agramma (Bezzi): 9 ♂ (NMSA - DIP 028196) 10 ♀ holotype (NMSA - DIP 028199); S. bezzii (Curran): 11 ♂ (SAM-DIP-A 018382) 12 ♀ (SAM-DIP-A 018410); S. bilobata Muller, sp. nov.: 13 a ♀ paratype 1 st flagellomere (CSCA) 13 b ♀ holotype (CSCA). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures9-13" httpUri="https://binary.pensoft.net/fig/1205864">12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EORBG" type="redescription">
<paragraph id="5CAD0E772A478E1344DEE175094E93DD">
<heading id="C3C87FE40175D8741085D638A2B868DC" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="0D5F95CF720DEC6EAC7B244707166333">
(Based on
<specimenCount id="E4426B870BBDEA8D8F85CC23CB12835F" count="1" type="female"></specimenCount>
<typeStatus id="1BDA8CA75B97CF67721D1E9A1D7E1C76" type="holotype">holotype</typeStatus>
photographs and additional
<specimenCount id="CF9272F6FAE46CAB5E7DDF165C355ABF" count="1" type="female"></specimenCount>
material from
<collectingCountry id="144DBBA34ECE50CC7975A393F1DAFC30" name="Uganda">Uganda</collectingCountry>
.)
<emphasis id="DAAA8E8B1E44FB3E755A203241871A0D" bold="true" italics="true">Head</emphasis>
: Black ground colour, with bluish-grey pruinosity on majority of head; eye bare; dichoptic; ommatidia of similar size; lateral edge of eye without any indentation; ocellar tubercle barely visible in profile, dark setulose (rubbed bare in
<specimenCount id="D609F6D9E317FE3FEEB0873F8A83D3D7" count="1" type="male"></specimenCount>
due to damage from preservation method), bluish-grey pruinose medially when viewed dorsally, otherwise appearing black; vertex grey pruinose and dark setulose; anterior ocellus similar in size to posterior pair; ocellar tubercle placed deeper in front of dorsal margin of eye compared to
<specimenCount id="ABBA96CFEF99EA0DC5D29DAD45FBD273" count="1" type="male"></specimenCount>
; dorsal inner edge of eye separated from ocellar tubercle by paired silver-grey markings; occiput with same bluish-grey pruinosity as rest of head; paired large rectangular black markings on upper occiput widening towards lateral margin of head with dark setulae, abutting posterior margin of eyes, flanking vertex; rest of upper occiput with pale setulae; lower occiput with long pale setulae medially and laterally; genal area with dark setulae, rest of ventral head setulae pale, these continue ventrally on head to mouthparts that have long pale ventral setulae and some shorter dark setulae; frons bluish-grey pruinose between lower half of eye down to antennal base, velvety-black from ocellar tubercle to lower half of eye; frons widening only slightly from velvety-black patch towards antennal bases; frons with dark setulae on velvety-black area; face, gena and clypeus with bluish-grey pruinosity; face sparsley populated with long pale setulae; clypeus black with bluish-grey pruinosity, bare; face separated anteriorly from clypeus by transverse suture, (less prominent than in
<specimenCount id="7D346CE5C3C0B781F932FFAFCA55CD79" count="1" type="male"></specimenCount>
) similar to lateral sutures; antennal bases separated ca 0.5 × 0.75 × width of scape, with slight longitudinal groove running between; scape dark brown with whitish pruinosity and with yellowish lateral margins; pedicel dark brown with whitish pruinosity; 1
<superScript id="2415D92A49E1889F8BA699F879E0CD65">st</superScript>
flagellomere orange to orange-brown with similar pruinosity as other segments, scape ca 1.52 × length of pedicel; 1
<superScript id="A5A95F6FD38D27C0D93D186D639B6032">st</superScript>
flagellomere ca 2 × size of pedicel; 2
<superScript id="E8305EC7791DD20D6E1D07D90ACB1D39">nd</superScript>
flagellomere, brown, arista-like; pedicel with dark dorsal and ventral setulae, similar in size, scape with only dark dorsal setulae; palpus black on majority of surface with scattered white pruinosity, orange-yellow on at most basal half, with dark setulae throughout; palpus ca 0.5 × length of proboscis; proboscis ca same length as head height; proboscis mostly brown, basally more orange-yellow, entire structure interspersed with some pale and dark setulae of varying length.
</paragraph>
<paragraph id="BFFA46B91E19E7EB4798781FAFCC56B3">
<emphasis id="A8B235134B9DF7CDAA0BA873EDAF008E" bold="true" italics="true">Thorax</emphasis>
: Scutum shining black with two feint dorsocentral bluish-grey pruinose vittae running from pronotum to before scutellum; pronotum bluish-grey pruinose with long pale setulae; postpronotal lobe dark brown, slight bluish-grey pruinose with long pale setulae, anterolateral margin of lobe lighter yellowish- to orange-brown (more apparent than in
<specimenCount id="879FE99F54FB574BBBA666AF901C1211" count="1" type="male"></specimenCount>
); scutum with paired, black almost velvety, rectangular marking behind postpronotal lobe (not visible in
<specimenCount id="77BE5F11692F6B76267AF3513D2747E9" count="1" type="male"></specimenCount>
due to damage); notopleuron with same colouration as in
<specimenCount id="1755BF1479704309BF3F6C3DB6B82405" count="1" type="male"></specimenCount>
, however,
<specimenCount id="CD71BEF6AD7816DD97612021EEBF67A8" count="1" type="female"></specimenCount>
notopleuron with scattered pale setulae interspersed with some dark setulae, anteriorly with pale setulae compared to dark group of setulae in
<specimenCount id="6E5BEDE8601A393A20D1C5412A43455A" count="1" type="male"></specimenCount>
; postalar wall and postalar callus dark brown with slight bluish-grey pruinosity, anterior of postalar callus orange-yellow; supra-alar area, postalar wall and postalar callus with patches of dark setulae (more apparent and numerous than in
<specimenCount id="776CF22C8CCDA2C31C8DD063D3E2A36B" count="1" type="male"></specimenCount>
); scutellum dark brown with bluish-grey pruinosity, apical margin orange-yellow; scutum generally with short dark setulae with postsutural setulae longer than presutural setulae, especially prescutellar setulae; majority of pleura bluish-grey pruinose, except for anterior of anepimeron and meron shiny blackish-brown; all pleura that are bluish-grey pruinose have some pale setulae; katatergite with long pale setulae, in contrast to
<specimenCount id="0A29CAEA4F9CD97E37216D50736D3115" count="1" type="male"></specimenCount>
that has long dark setulae; anatergite and meron bare; proepisternum and proepimeron with long pale setulae; anterior and posterior spiracles yellowish-brown to whitish-yellow (due to damage from preservation method) and surroundings yellowish-brown, bare; postspiracular scale dark brown.
</paragraph>
<paragraph id="2805A2BB238BB8D1ED8E3CF2B7D958AD">
<emphasis id="5217FAA626626A71DAD36F135FFD7288" bold="true" italics="true">Legs</emphasis>
: All coxae brown to blackish-brown with some bluish-grey pruinosity; fore coxa with mostly pale setulae except for some dark setulae apically; mid coxa with long dark setulae on anterior apical surface, otherwise with pale setulae, and sparsley setulose along posterior margin (
<specimenCount id="BC5EA21F965350C71DB9331B5E9598AE" count="1" type="male"></specimenCount>
with only dark setulae); hind coxa with dark setulae on anterior edge surrounding well-developed anterior apical point, lateral apical edges with long pale setulae; all trochanters reddish dark brown, some more yellowish apically, all trochanters with short pale setulae; fore femur in most specimens yellow with extreme base and apex dark brown, some individuals with fore femur dark brown on basal half and apical quarter to third with remainder of segment yellow; mid femur dark brown except for ca apical third that is yellow; hind femur dark brown except for yellow basal and apical quarters; fore tibia and tarsi dark brown to blackish; mid tibia and tarsi yellow, tarsi appearing somewhat darker yellow; hind tibia and basitarsus blackish-brown, with apex of tibia yellowish-brown in some specimens; fore tarsal claws and pulvilli symmetrical, pulvilus and empodium of similar size; fore tarsi with long, somewhat curved along antero- and posteroventral surfaces, sensory setulae ca 2 × as long as tarsal segment is wide; fore femur dorsally with short dark setulae, except for several scattered short pale setulae, apically and medioventrally with long pale setulae, and with a group of 13 dark setulae located in middle of femur; mid femur covered in short pale setulae, but with longer pale setulae ventrally and short dark setulae apically; hind femur with mix of short pale and dark setulae, with apex having somewhat longer pale and dark setulae ventrally; hind leg overall stouter than remaining legs; all tibiae with short dark setulae; hind tarsal segments 0.91.0 × as long as hind tibia; tibial spur formula 0: 2: 2.
</paragraph>
<paragraph id="67342C8708C489E0C4FEAAB6E58C4E58">
<emphasis id="97C335533D47A25F38FD0A68A7D76ABD" bold="true" italics="true">Wing</emphasis>
: Light brown suffused on majority of surface, except for cells
<emphasis id="3BF9486F261499CCEFD98FAE1A74C855" italics="true">bm</emphasis>
and
<emphasis id="6C94F73F49927D21B8551BBC692685DA" italics="true">cua</emphasis>
, and anal lobe, overall appearing darker on apical half; dark brown stigma over cell
<emphasis id="03D7EA30EC463B3809E89104985976CA" italics="true">
r
<subScript id="BD2641864BC140E7F13B4F50A3984D40">1</subScript>
</emphasis>
; darker suffused substigmal marking running down from stigma over crossvein
<emphasis id="951FBA8CF5CF5282B61E486E29DA8BAA" italics="true">r m</emphasis>
, bases of discal cell, cell
<emphasis id="6B5B0D69F0056B7EB4A23B7D5201CA4A" italics="true">
m
<subScript id="C5474BE324A0A30482B5582AB191AF42">3</subScript>
</emphasis>
and apex of cell
<emphasis id="4F99A1FC86DA6E48BA7BE2384ED273AA" italics="true">br</emphasis>
; veins dark brown, with additional darker brown suffusion around vein
<emphasis id="00A6893BB79B212D458D2FDFF34F69F0" italics="true">CuA</emphasis>
; costa without distinct downward flexure over stigma; cell
<emphasis id="902D726CEC13026CA8FBDB8845DA43FF" italics="true">cua</emphasis>
closed at wing margin or at a short distance from wing margin; cell
<emphasis id="CD8BAACDFE495589D3F6D0A40B831436" italics="true">
m
<subScript id="0F4B528B9436B98A3A4411373AE63B5F">3</subScript>
</emphasis>
open, veins
<emphasis id="24E9A3CA0F10ED0C48270D2070543FC7" italics="true">
M
<subScript id="57C23D751E13B5A74532A3FA105187BA">1</subScript>
</emphasis>
,
<emphasis id="C74E460F2DC93BEFF6F28FCE6A0731C5" italics="true">
M
<subScript id="98C39B4822477A4A5CD39D1C8B9F9C0C">2</subScript>
</emphasis>
,
<emphasis id="6D01228179EC7A1D33726942570C6E38" italics="true">
M
<subScript id="0D62A60BFA9FEBF64E33D8F02E5716E7">3</subScript>
</emphasis>
present; haltere stalk dirty yellow, knob darker yellowish-brown, with a few short and dark setulae.
</paragraph>
<paragraph id="D0FBC155C0E1421250EDCEC3AF71494F">
<emphasis id="F1C7C1EFAA9A0B4715FFE49834CC30E6" bold="true" italics="true">Abdomen</emphasis>
: Tergite 1 dark to blackish-brown, with basal margin of some specimens more orange-brown, tergite 1 with pale setulae; tergite 2 mostly pale to orange-yellow, with brown to dark brown median marking running down to brown to dark brown band along posterior margin of tergite, giving it a fenestrated appearance, lateral margins of tergite similar brown colour to rest of tergite; tergites 3 and 4 dark brown with tergite 4 with some specimens also having orange markings posteriorly, tergites 57 orange, each with a dark brown posterior band; lateral margins of tergites 47 with blackish markings; some specimens have tergite 7 entirely dark brown; dark brown parts of tergites with pale setulae, and orange parts with dark setulae; tergite 1 medially with a longitudinal suture; sternites 1 and 2 cream coloured, sternites 3 and 4 dark brown with posterior margins darker; rest of sternites orange-yellow, ending in dark brown terminalia; all sternites with pale setulae.
</paragraph>
<paragraph id="E7F18FDB69BCE8C77A7ED7C22BD344AC">
<emphasis id="2763A917B98B7805A6A128891316E2AC" bold="true" italics="true">Terminalia</emphasis>
(Fig.
<figureCitation id="90C45A33F7026F94C90837620AF6FDD0" captionStart="Figures 7587" captionStartId="F16" captionText="Figures 7587. Suragina Walker spp. female genital fork and spermathecae, ventral view: 75 S. agramma (Bezzi) (ICIPE) 76 S. bezzii (Curran) (SAM-DIP- 018403) 77 S. bilobata Muller, sp. nov. (CSCA) 78 S. binominata (Bequaert) (NMSA - DIP 158411) 79 S. copelandi Muller, sp. nov. paratype (BMSA (D) 84686) 80 S. freidbergi Muller, sp. nov. paratype (BMSA (D) 92158) 81 S. liberiaensis Muller, sp. nov. paratype (NMSA - DIP 158442) 82 S. malavaensis Muller, sp. nov. paratype (ICIPE) 83 S. milloti (Séguy) (CSCA) 84 S. monogramma (Bezzi) (BMSA (D) 129988) 85 S. mulanjeensis Muller, sp. nov. paratype (NMSA - DIP 158392) 86 S. semiobscura Muller, sp. nov. paratype (ICIPE) 87 S. zombaensis Muller, sp. nov. paratype (NMSA - DIP 1584719). Scale bar (applicable to all illustrations): 0.1 mm." figureDoi="10.3897/afrinvertebr.65.140524.figures75-87" httpUri="https://binary.pensoft.net/fig/1205878">76</figureCitation>
): Cercus dark brown with pale setulae; genital fork has distal apodeme slender, forked; median lobe with deep apical emargination, paired apical lobes with a slender appearance, inner surface inward projecting with clustered microtrichia at apex, arms gradually rounded; three ovate-shaped and sclerotised spermathecae.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIUBG" type="distribution">
<paragraph id="AC400E8CAC0D20F59ABF577B69E62C45">
<heading id="3167698ABB1896064D7808371C84FFC8" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="F327E12099234668B1FDA7D8BF6E64AD">
<collectingCountry id="5D0B5B757E20FD98A97520656B8B5636" name="Burundi">Burundi</collectingCountry>
(new record),
<collectingCountry id="95E867F59E8864E8E7B3896D7B7EC932" name="Democratic Republic of the Congo">Democratic Republic of Congo</collectingCountry>
,
<collectingCountry id="8721AF18739FFEB3C3B417EF56A609AF" name="Kenya">Kenya</collectingCountry>
(new record),
<collectingCountry id="B8C89C1F59E97B3E707B8A0B9C394BB9" name="Uganda">Uganda</collectingCountry>
(new record).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,59 +1,59 @@
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<mods:affiliation id="BDD341CE5CF5F1C463C0335E99DA4B53">Department Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, 9301, South Africa &amp; Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
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<treatment id="8F5BAEF55B745DEB816C739831546EB2" ID-arpha="8F5BAEF5-5B74-5DEB-816C-739831546EB2" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2" httpUri="http://treatment.plazi.org/id/8F5BAEF55B745DEB816C739831546EB2">
<treatment id="8F5BAEF55B745DEB816C739831546EB2" ID-DOI="http://doi.org/10.5281/zenodo.14548345" ID-Zenodo-Dep="14548345" ID-arpha="8F5BAEF5-5B74-5DEB-816C-739831546EB2" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2" httpUri="http://treatment.plazi.org/id/8F5BAEF55B745DEB816C739831546EB2">
<subSubSection id="548997F287440549C72A8FA1D926CAD9" type="nomenclature">
<paragraph id="EB3C3399955BD0D1A0D45BF4901F96F1">
<taxonomicName id="46EDBCFF875B0C2DB0622C9CB9E9CF45" ID-CoL="53H65" authority="(Bezzi, 1926)" authorityName="ICIPE" baseAuthorityName="Bezzi" baseAuthorityYear="1926" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="agramma">
@ -97,7 +97,6 @@
</paragraph>
<paragraph id="903F0C235B6766DAB3551500C67F7DB7">
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<taxonomicName id="D41B5012A0FDEB68CC6974CEEFB66EC9" ID-CoL="53H65" authorityName="ICIPE" baseAuthorityName="Bezzi" baseAuthorityYear="1926" class="Insecta" family="Athericidae" genus="Suragina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="agramma">
<emphasis id="4D32325C78CEE26EB820AFB477765F67" italics="true">Suragina agramma</emphasis>
</taxonomicName>
@ -112,7 +111,6 @@
</treatmentCitation>
.
</comment>
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</paragraph>
</subSubSection>
@ -145,7 +143,7 @@ of
<heading id="3334B31144034D22D96D8B0C543705CD" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="F526E0075D4B7E04CF416DC66DB4B95E">
<materialsCitation id="A730AA770912F3A96394567CADE18994" collectingDate="1919-10-05" collectionCode="NMSA" collectorName="De Kaap &amp; H. K. Munro" country="South Africa" latitude="-25.587555" location="De Kaap" longLatPrecision="1" longitude="30.973667" municipality="Baberton" specimenCode="DIP 028199" specimenCount="1" specimenCount-female="1" stateProvince="Mpumalanga" typeStatus="holotype">
<materialsCitation id="A730AA770912F3A96394567CADE18994" collectingDate="1919-10-05" collectionCode="NMSA" collectorName="H. K. Munro" country="South Africa" latitude="-25.587555" location="De Kaap" longLatPrecision="1" longitude="30.973667" municipality="Baberton" specimenCode="DIP 028199" specimenCount="1" specimenCount-female="1" stateProvince="Mpumalanga" typeStatus="holotype">
<emphasis id="5B65D9C083E20F5AD4062C1632CA9ED8" bold="true" italics="true">
<typeStatus id="FA62F3C3F08C6F24205B3E715029A8A9" type="holotype">Holotype</typeStatus>
</emphasis>
@ -158,14 +156,12 @@ of
];
<collectingMunicipality id="775D9C15E9317EB608497C10B22B683B">Baberton</collectingMunicipality>
,
<collectorName id="36A3E537A037F352AF74F419EE005A1F">
<location id="E082285B330C6EA9031B1C99577C1941" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:E082285B330C6EA9031B1C99577C1941" country="South Africa" latitude="-25.587555" longLatPrecision="1" longitude="30.973667" municipality="Baberton" name="De Kaap" stateProvince="Mpumalanga">De Kaap</location>
</collectorName>
[Valley], B. 50; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPGAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[30.973667,-25.587555]}">
<geoCoordinate id="B8BB2E2F2B38771F7E7DDB369FCD2856" degrees="25" direction="south" minutes="35.2533" orientation="latitude" precision="1" value="-25.587555">25 ° 35.2533 ' S</geoCoordinate>
<geoCoordinate id="B8BB2E2F2B38771F7E7DDB369FCD2856" degrees="25" direction="south" minutes="35.2533" orientation="latitude" precision="1" value="-25.587555">25°35.2533'S</geoCoordinate>
,
<geoCoordinate id="68A9120AEFB16FA2CE5CE17423C1AB47" degrees="30" direction="east" minutes="58.4200" orientation="longitude" precision="1" value="30.973667">30 ° 58.4200 ' E</geoCoordinate>
<geoCoordinate id="68A9120AEFB16FA2CE5CE17423C1AB47" degrees="30" direction="east" minutes="58.4200" orientation="longitude" precision="1" value="30.973667">30°58.4200'E</geoCoordinate>
</named-content>
];
<date id="E4A1C839E071918CC8664264C024AB77" value="1919-10-05">
@ -200,9 +196,9 @@ leg.; (
<location id="3D54E501E143D1067E57E1E347F49319" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:3D54E501E143D1067E57E1E347F49319" country="Kenya" latitude="-4.23456" longLatPrecision="1" longitude="39.401688" municipality="Shimba Hills Nat. Pk." name="Longomwagandi Forest" stateProvince="Coast Province">Longomwagandi Forest</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBHAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[39.401687,-4.234560]}">
<geoCoordinate id="0DA0F549FB02F8AD1F873F57B09D6369" degrees="04" direction="south" minutes="14.0736" orientation="latitude" precision="1" value="-4.23456">04 ° 14.0736 ' S</geoCoordinate>
<geoCoordinate id="0DA0F549FB02F8AD1F873F57B09D6369" degrees="04" direction="south" minutes="14.0736" orientation="latitude" precision="1" value="-4.23456">04°14.0736'S</geoCoordinate>
,
<geoCoordinate id="15D27515CFA4766A8B06FE93855D228B" degrees="39" direction="east" minutes="24.1012" orientation="longitude" precision="1" value="39.401688">39 ° 24.1012 ' E</geoCoordinate>
<geoCoordinate id="15D27515CFA4766A8B06FE93855D228B" degrees="39" direction="east" minutes="24.1012" orientation="longitude" precision="1" value="39.401688">39°24.1012'E</geoCoordinate>
</named-content>
;
<quantity id="50CE8778E7CD256AF80C183FD4CD96F3" metricMagnitude="2" metricUnit="m" metricValue="3.8899999999999997" unit="m" value="389.0">
@ -231,9 +227,9 @@ leg.;
<location id="FCA604E22C471E203393BE31ECE5AB4F" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:FCA604E22C471E203393BE31ECE5AB4F" country="Kenya" latitude="-4.23456" longLatPrecision="1" longitude="39.401688" municipality="Shimba Hills Nat. Pk." name="Longomwagandi Forest" stateProvince="Coast Province">Longomwagandi Forest</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENHAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[39.401687,-4.234560]}">
<geoCoordinate id="2DB4DC0E30F8CE0B9326F4BC4304C4F6" degrees="04" direction="south" minutes="14.0736" orientation="latitude" precision="1" value="-4.23456">04 ° 14.0736 ' S</geoCoordinate>
<geoCoordinate id="2DB4DC0E30F8CE0B9326F4BC4304C4F6" degrees="04" direction="south" minutes="14.0736" orientation="latitude" precision="1" value="-4.23456">04°14.0736'S</geoCoordinate>
,
<geoCoordinate id="2E010DED8FA5726348E1C6FDA1AF3D8D" degrees="39" direction="east" minutes="24.1012" orientation="longitude" precision="1" value="39.401688">39 ° 24.1012 ' E</geoCoordinate>
<geoCoordinate id="2E010DED8FA5726348E1C6FDA1AF3D8D" degrees="39" direction="east" minutes="24.1012" orientation="longitude" precision="1" value="39.401688">39°24.1012'E</geoCoordinate>
</named-content>
;
<quantity id="35BECD6E95CF932653AF3E7DDAD7F934" metricMagnitude="2" metricUnit="m" metricValue="3.8899999999999997" unit="m" value="389.0">
@ -262,9 +258,9 @@ leg.;
<location id="9216376CAC13097BC433117F90405142" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:9216376CAC13097BC433117F90405142" country="Kenya" latitude="-4.23456" longLatPrecision="1" longitude="39.401688" municipality="Shimba Hills Nat. Pk." name="Longomwagandi Forest" stateProvince="Coast Province">Longomwagandi Forest</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZHAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[39.401687,-4.234560]}">
<geoCoordinate id="1A2813994211F037DF19DA053FD83EF9" degrees="04" direction="south" minutes="14.0736" orientation="latitude" precision="1" value="-4.23456">04 ° 14.0736 ' S</geoCoordinate>
<geoCoordinate id="1A2813994211F037DF19DA053FD83EF9" degrees="04" direction="south" minutes="14.0736" orientation="latitude" precision="1" value="-4.23456">04°14.0736'S</geoCoordinate>
,
<geoCoordinate id="F491072A6612C00EE20ACB468371B0CB" degrees="39" direction="east" minutes="24.1012" orientation="longitude" precision="1" value="39.401688">39 ° 24.1012 ' E</geoCoordinate>
<geoCoordinate id="F491072A6612C00EE20ACB468371B0CB" degrees="39" direction="east" minutes="24.1012" orientation="longitude" precision="1" value="39.401688">39°24.1012'E</geoCoordinate>
</named-content>
;
<quantity id="02F892A84013D289FBF75B0F767C6826" metricMagnitude="2" metricUnit="m" metricValue="3.8899999999999997" unit="m" value="389.0">
@ -299,9 +295,9 @@ leg.;
<location id="27F09EB102110BE6A1EBDDFD78CF8B67" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:27F09EB102110BE6A1EBDDFD78CF8B67" country="Kenya" county="Coast Province" latitude="-3.4844334" longLatPrecision="1" longitude="38.332516" municipality="Hills" name="Bura Bluff" stateProvince="Taita/Taveta">Bura Bluff</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFIAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[38.332517,-3.484433]}">
<geoCoordinate id="4368AFF524580EAE4D0FA481013AC832" degrees="03" direction="south" minutes="29.066" orientation="latitude" precision="1" value="-3.4844334">03 ° 29.066 ' S</geoCoordinate>
<geoCoordinate id="4368AFF524580EAE4D0FA481013AC832" degrees="03" direction="south" minutes="29.066" orientation="latitude" precision="1" value="-3.4844334">03°29.066'S</geoCoordinate>
,
<geoCoordinate id="336A10E281024D38628A6983E3813CF7" degrees="38" direction="east" minutes="19.951" orientation="longitude" precision="1" value="38.332516">38 ° 19.951 ' E</geoCoordinate>
<geoCoordinate id="336A10E281024D38628A6983E3813CF7" degrees="38" direction="east" minutes="19.951" orientation="longitude" precision="1" value="38.332516">38°19.951'E</geoCoordinate>
</named-content>
;
<quantity id="D25F1069DFDAC407DE7264A7540D3C49" metricMagnitude="3" metricUnit="m" metricValue="1.011" unit="m" value="1011.0">
@ -333,9 +329,9 @@ leg.; riverine forest;
<location id="DAB8EC33FDF470C67DB33BFAF13D32CA" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:DAB8EC33FDF470C67DB33BFAF13D32CA" country="Kenya" county="Coast Province" latitude="-3.4790833" longLatPrecision="1" longitude="38.341618" municipality="Hills" name="Chawia" stateProvince="Taita/Taveta">Chawia</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERIAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[38.341617,-3.479083]}">
<geoCoordinate id="2CA0AAB1F1B7BF90416A7D9A40561A00" degrees="03" direction="south" minutes="28.745" orientation="latitude" precision="1" value="-3.4790833">03 ° 28.745 ' S</geoCoordinate>
<geoCoordinate id="2CA0AAB1F1B7BF90416A7D9A40561A00" degrees="03" direction="south" minutes="28.745" orientation="latitude" precision="1" value="-3.4790833">03°28.745'S</geoCoordinate>
,
<geoCoordinate id="3ED6AD39F5EA329A4CB24B2D0E689B6C" degrees="38" direction="east" minutes="20.497" orientation="longitude" precision="1" value="38.341618">38 ° 20.497 ' E</geoCoordinate>
<geoCoordinate id="3ED6AD39F5EA329A4CB24B2D0E689B6C" degrees="38" direction="east" minutes="20.497" orientation="longitude" precision="1" value="38.341618">38°20.497'E</geoCoordinate>
</named-content>
;
<quantity id="984FEE2D00D198E9614EDCB1575E37D1" metricMagnitude="3" metricUnit="m" metricValue="1.614" unit="m" value="1614.0">
@ -367,9 +363,9 @@ leg.; next to small forest pond;
<location id="54A9209B1171B3318664E295B8937599" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:54A9209B1171B3318664E295B8937599" country="Kenya" county="Coast Province" latitude="-3.4790833" longLatPrecision="1" longitude="38.341618" municipality="Hills" name="Chawia" stateProvince="Taita/Taveta">Chawia</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4IAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[38.341617,-3.479083]}">
<geoCoordinate id="BB1803A172C9BDE1342AD6E1B4E61937" degrees="03" direction="south" minutes="28.745" orientation="latitude" precision="1" value="-3.4790833">03 ° 28.745 ' S</geoCoordinate>
<geoCoordinate id="BB1803A172C9BDE1342AD6E1B4E61937" degrees="03" direction="south" minutes="28.745" orientation="latitude" precision="1" value="-3.4790833">03°28.745'S</geoCoordinate>
,
<geoCoordinate id="F6C954C928FF83E58B7C3A3AA8B45E51" degrees="38" direction="east" minutes="20.497" orientation="longitude" precision="1" value="38.341618">38 ° 20.497 ' E</geoCoordinate>
<geoCoordinate id="F6C954C928FF83E58B7C3A3AA8B45E51" degrees="38" direction="east" minutes="20.497" orientation="longitude" precision="1" value="38.341618">38°20.497'E</geoCoordinate>
</named-content>
;
<quantity id="89F155F56FD1ADE624DE2612AAB4727C" metricMagnitude="3" metricUnit="m" metricValue="1.614" unit="m" value="1614.0">
@ -403,9 +399,9 @@ leg.; next to small forest pond;
<location id="3BB7638A4AA1F0158EA4C25B903C4220" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:3BB7638A4AA1F0158EA4C25B903C4220" country="Kenya" county="Coast Province" latitude="-3.4844334" longLatPrecision="1" longitude="38.332516" municipality="Hills" name="Bura Bluff" stateProvince="Taita/Taveta">Bura Bluff</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJJAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[38.332517,-3.484433]}">
<geoCoordinate id="808155A929450B7B395822D6A33BC068" degrees="03" direction="south" minutes="29.066" orientation="latitude" precision="1" value="-3.4844334">03 ° 29.066 ' S</geoCoordinate>
<geoCoordinate id="808155A929450B7B395822D6A33BC068" degrees="03" direction="south" minutes="29.066" orientation="latitude" precision="1" value="-3.4844334">03°29.066'S</geoCoordinate>
,
<geoCoordinate id="8B97B7F0B2FBC8185927745D3DE455B2" degrees="38" direction="east" minutes="19.951" orientation="longitude" precision="1" value="38.332516">38 ° 19.951 ' E</geoCoordinate>
<geoCoordinate id="8B97B7F0B2FBC8185927745D3DE455B2" degrees="38" direction="east" minutes="19.951" orientation="longitude" precision="1" value="38.332516">38°19.951'E</geoCoordinate>
</named-content>
;
<quantity id="9BD4FB406772B9628F02D93CD7E83C0E" metricMagnitude="3" metricUnit="m" metricValue="1.011" unit="m" value="1011.0">
@ -437,9 +433,9 @@ leg.; riverine forest;
<location id="29696B2CFB7AEDF5143973A5FE871DC9" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:29696B2CFB7AEDF5143973A5FE871DC9" country="Kenya" county="Coast Province" latitude="-3.4790833" longLatPrecision="1" longitude="38.341618" municipality="Hills" name="Chawia" stateProvince="Taita/Taveta">Chawia</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVJAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[38.341617,-3.479083]}">
<geoCoordinate id="B0B8AA8F15980FD7616182AC72DA463C" degrees="03" direction="south" minutes="28.745" orientation="latitude" precision="1" value="-3.4790833">03 ° 28.745 ' S</geoCoordinate>
<geoCoordinate id="B0B8AA8F15980FD7616182AC72DA463C" degrees="03" direction="south" minutes="28.745" orientation="latitude" precision="1" value="-3.4790833">03°28.745'S</geoCoordinate>
,
<geoCoordinate id="4504FC4334E9B9C73AA6D03D9536BC2C" degrees="38" direction="east" minutes="20.497" orientation="longitude" precision="1" value="38.341618">38 ° 20.497 ' E</geoCoordinate>
<geoCoordinate id="4504FC4334E9B9C73AA6D03D9536BC2C" degrees="38" direction="east" minutes="20.497" orientation="longitude" precision="1" value="38.341618">38°20.497'E</geoCoordinate>
</named-content>
;
<quantity id="0622B9D92105E843362F2591E036C490" metricMagnitude="3" metricUnit="m" metricValue="1.614" unit="m" value="1614.0">
@ -473,9 +469,9 @@ leg.; next to small forest pond;
<location id="7C2169E89DC95C09C5718C060736C7A1" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:7C2169E89DC95C09C5718C060736C7A1" country="Kenya" county="Coast Province" latitude="-3.4844334" longLatPrecision="1" longitude="38.332516" municipality="Hills" name="Bura Bluff" stateProvince="Taita/Taveta">Bura Bluff</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBKAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[38.332517,-3.484433]}">
<geoCoordinate id="F59AD0E6BC6D3B6D58B2E6251E1D9F0E" degrees="03" direction="south" minutes="29.066" orientation="latitude" precision="1" value="-3.4844334">03 ° 29.066 ' S</geoCoordinate>
<geoCoordinate id="F59AD0E6BC6D3B6D58B2E6251E1D9F0E" degrees="03" direction="south" minutes="29.066" orientation="latitude" precision="1" value="-3.4844334">03°29.066'S</geoCoordinate>
,
<geoCoordinate id="A66FE7A402869A84A73AA49C963B4B6C" degrees="38" direction="east" minutes="19.951" orientation="longitude" precision="1" value="38.332516">38 ° 19.951 ' E</geoCoordinate>
<geoCoordinate id="A66FE7A402869A84A73AA49C963B4B6C" degrees="38" direction="east" minutes="19.951" orientation="longitude" precision="1" value="38.332516">38°19.951'E</geoCoordinate>
</named-content>
;
<quantity id="0E1BE5CF54564713C412B0E88F97BF6C" metricMagnitude="3" metricUnit="m" metricValue="1.011" unit="m" value="1011.0">
@ -510,9 +506,9 @@ leg.; riverine forest;
<location id="42B096575607C3EE2A9804655CC8EF37" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:42B096575607C3EE2A9804655CC8EF37" country="Kenya" county="Coast Province" latitude="-3.4844334" longLatPrecision="1" longitude="38.332516" municipality="Hills" name="Bura Bluff" stateProvince="Taita/Taveta">Bura Bluff</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENKAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[38.332517,-3.484433]}">
<geoCoordinate id="0EEB5832F4D8717D560272C40A50BFF1" degrees="03" direction="south" minutes="29.066" orientation="latitude" precision="1" value="-3.4844334">03 ° 29.066 ' S</geoCoordinate>
<geoCoordinate id="0EEB5832F4D8717D560272C40A50BFF1" degrees="03" direction="south" minutes="29.066" orientation="latitude" precision="1" value="-3.4844334">03°29.066'S</geoCoordinate>
,
<geoCoordinate id="476CECFACB5916B5DB1CA44ACA25A346" degrees="38" direction="east" minutes="19.951" orientation="longitude" precision="1" value="38.332516">38 ° 19.951 ' E</geoCoordinate>
<geoCoordinate id="476CECFACB5916B5DB1CA44ACA25A346" degrees="38" direction="east" minutes="19.951" orientation="longitude" precision="1" value="38.332516">38°19.951'E</geoCoordinate>
</named-content>
;
<quantity id="2EBE7764BA16BA98ED9DF2FA3D30D068" metricMagnitude="3" metricUnit="m" metricValue="1.011" unit="m" value="1011.0">
@ -546,9 +542,9 @@ leg.; riverine forest;
<location id="5D18C93746E5CAEC73DAB544F09E7050" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:5D18C93746E5CAEC73DAB544F09E7050" country="Kenya" county="Coast Province" latitude="-3.4844334" longLatPrecision="1" longitude="38.332516" municipality="Hills" name="Bura Bluff" stateProvince="Taita/Taveta">Bura Bluff</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZKAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[38.332517,-3.484433]}">
<geoCoordinate id="56ECB040FED304C1923868920652505C" degrees="03" direction="south" minutes="29.066" orientation="latitude" precision="1" value="-3.4844334">03 ° 29.066 ' S</geoCoordinate>
<geoCoordinate id="56ECB040FED304C1923868920652505C" degrees="03" direction="south" minutes="29.066" orientation="latitude" precision="1" value="-3.4844334">03°29.066'S</geoCoordinate>
,
<geoCoordinate id="F2CAEAFAD097219410AA4EA82F82B89A" degrees="38" direction="east" minutes="19.951" orientation="longitude" precision="1" value="38.332516">38 ° 19.951 ' E</geoCoordinate>
<geoCoordinate id="F2CAEAFAD097219410AA4EA82F82B89A" degrees="38" direction="east" minutes="19.951" orientation="longitude" precision="1" value="38.332516">38°19.951'E</geoCoordinate>
</named-content>
;
<quantity id="6499009C65B9D8446E502062D5E7B41A" metricMagnitude="3" metricUnit="m" metricValue="1.011" unit="m" value="1011.0">
@ -583,9 +579,9 @@ leg.; riverine forest;
<location id="653DCFEA135343626CFE78705D742213" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:653DCFEA135343626CFE78705D742213" country="Kenya" latitude="-0.51663" longLatPrecision="1" longitude="37.41852" municipality="Njuki-ini Forest" name="Forest station" stateProvince="Eastern Province">Forest station</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFLAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[37.418520,-0.516630]}">
<geoCoordinate id="792A1FAB3DD744419265112E10D07D4E" degrees="00" direction="south" minutes="30.9978" orientation="latitude" precision="1" value="-0.51663">00 ° 30.9978 ' S</geoCoordinate>
<geoCoordinate id="792A1FAB3DD744419265112E10D07D4E" degrees="00" direction="south" minutes="30.9978" orientation="latitude" precision="1" value="-0.51663">00°30.9978'S</geoCoordinate>
,
<geoCoordinate id="723685BCCAB0E79A1B8A7586C578C832" degrees="37" direction="east" minutes="25.1112" orientation="longitude" precision="1" value="37.41852">37 ° 25.1112 ' E</geoCoordinate>
<geoCoordinate id="723685BCCAB0E79A1B8A7586C578C832" degrees="37" direction="east" minutes="25.1112" orientation="longitude" precision="1" value="37.41852">37°25.1112'E</geoCoordinate>
</named-content>
;
<quantity id="EF8B5C4EC3CDF20B3710E5DE18854EE8" metricMagnitude="3" metricUnit="m" metricValue="1.4709999999999999" unit="m" value="1471.0">
@ -606,7 +602,7 @@ leg.; inside indigenous forest;
)
</materialsCitation>
<materialsCitation id="BBB3C32AF3767F9555D7BD578E9DD35A" collectingDate="2006-02-12" collectingDateMax="2006-02-26" collectingDateMin="2006-02-12" collectionCode="ICIPE" collectorName="R. Copeland" country="Malawi" elevation="1455" latitude="-0.5166" location="Forest station" longLatPrecision="1" longitude="37.41843" municipality="Njuki-ini Forest" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Province">
<materialsCitation id="70423BEFA9971EC1B9733620C5046D19" collectingDate="2006-02-12" collectingDateMax="2006-02-26" collectingDateMin="2006-02-12" collectionCode="ICIPE" collectorName="R. Copeland" country="Malawi" elevation="1455" latitude="-0.5166" location="Forest station" longLatPrecision="1" longitude="37.41843" municipality="Njuki-ini Forest" specimenCount="1" specimenCount-female="1" stateProvince="Eastern Province">
<specimenCount id="CFF18E1CB59C7237F5B4FC4261A833DE" count="1" type="female">1 ♀</specimenCount>
;
<collectingRegion id="C65FC8767310F351404A1B7BF4F33B49" inferredBySuffix="true">Eastern Province</collectingRegion>
@ -616,9 +612,9 @@ leg.; inside indigenous forest;
<location id="774DE859781B7BB918093250521A32F3" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:774DE859781B7BB918093250521A32F3" country="Malawi" latitude="-0.5166" longLatPrecision="1" longitude="37.41843" municipality="Njuki-ini Forest" name="Forest station" stateProvince="Eastern Province">Forest station</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERLAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[37.418430,-0.516600]}">
<geoCoordinate id="D7318EE989C48ECCC6E81B5943AE8D82" degrees="00" direction="south" minutes="30.9960" orientation="latitude" precision="1" value="-0.5166">00 ° 30.9960 ' S</geoCoordinate>
<geoCoordinate id="D7318EE989C48ECCC6E81B5943AE8D82" degrees="00" direction="south" minutes="30.9960" orientation="latitude" precision="1" value="-0.5166">00°30.9960'S</geoCoordinate>
,
<geoCoordinate id="FA36CE855AC858363199DC91D17F2471" degrees="37" direction="east" minutes="25.1058" orientation="longitude" precision="1" value="37.41843">37 ° 25.1058 ' E</geoCoordinate>
<geoCoordinate id="FA36CE855AC858363199DC91D17F2471" degrees="37" direction="east" minutes="25.1058" orientation="longitude" precision="1" value="37.41843">37°25.1058'E</geoCoordinate>
</named-content>
;
<quantity id="B0ED17EF1B6432DCA55CCA0F0B28759C" metricMagnitude="3" metricUnit="m" metricValue="1.455" unit="m" value="1455.0">
@ -636,11 +632,12 @@ leg.; inside indigenous forest;
<abbrev id="ABBRID0EWLAG" xlink_title="International Centre of Insect Physiology and Ecology">
<collectionCode id="503410CE61713B1319A0E60E59CB94AD">ICIPE</collectionCode>
</abbrev>
).
<collectingCountry id="C5BA292BBA9AB2832EC720E7BEFD14F7" name="Malawi">Malawi</collectingCountry>
)
</materialsCitation>
.
<materialsCitation id="D5CE65277F53B1F64B407B677DB419AB" collectingDate="1980-09-24" collectingDateMax="1980-09-27" collectingDateMin="1980-09-24" collectionCode="NMSA" collectorName="J. G. H. Londt &amp; B. R. Stuckenberg" country="South Africa" elevation="1100" latitude="-15.383333" location="Ad" longLatPrecision="13" longitude="35.333332" specimenCode="DIP 028177" specimenCount="1" specimenCount-female="1" stateProvince="Zomba">
<collectingCountry id="C5BA292BBA9AB2832EC720E7BEFD14F7" name="Malawi">Malawi</collectingCountry>
<materialsCitation id="333F5E36D51A5B174FFB4C9D6032D7F9" collectingDate="1980-09-24" collectingDateMax="1980-09-27" collectingDateMin="1980-09-24" collectionCode="NMSA" collectorName="J. G. H. Londt &amp; B. R. Stuckenberg" country="South Africa" elevation="1100" latitude="-15.383333" location="Ad" longLatPrecision="13" longitude="35.333332" specimenCode="DIP 028177" specimenCount="1" specimenCount-female="1" stateProvince="Zomba">
<specimenCount id="AA91BE949F999E82BA142E15322A0BBB" count="1" type="female">1 ♀</specimenCount>
; [Southern Region];
<collectingRegion id="4D759D2A423184C76413411F30CDAD89" country="Malawi" name="Zomba">Zomba</collectingRegion>
@ -648,9 +645,9 @@ leg.; inside indigenous forest;
<location id="8795431C6B7BF9889BEFB8EF81143CD3" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:8795431C6B7BF9889BEFB8EF81143CD3" country="South Africa" latitude="-15.383333" longLatPrecision="13" longitude="35.333332" name="Ad" stateProvince="Zomba">Ad</location>
[
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4LAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[35.333333,-15.383333]}">
<geoCoordinate id="F5F38752C376D342E35E945C87A8A567" degrees="15" direction="south" minutes="23.00" orientation="latitude" precision="9" value="-15.383333">15 ° 23.00 ' S</geoCoordinate>
<geoCoordinate id="F5F38752C376D342E35E945C87A8A567" degrees="15" direction="south" minutes="23.00" orientation="latitude" precision="9" value="-15.383333">15°23.00'S</geoCoordinate>
,
<geoCoordinate id="B1066EF99FDF53E41F3EB0043E699194" degrees="35" direction="east" minutes="20.00" orientation="longitude" precision="9" value="35.333332">35 ° 20.00 ' E</geoCoordinate>
<geoCoordinate id="B1066EF99FDF53E41F3EB0043E699194" degrees="35" direction="east" minutes="20.00" orientation="longitude" precision="9" value="35.333332">35°20.00'E</geoCoordinate>
</named-content>
];
<quantity id="9DCDA44B017BCD361B003DEDE005D660" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
@ -670,11 +667,12 @@ leg.; (
</named-content>
-
<specimenCode id="F1147FB802125FA6692D50AAB5F54406" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" type="Museum">DIP 028177</specimenCode>
).
<collectingCountry id="079B376B0DCC29B69F26535A73E2AFA4" name="South Africa">South Africa</collectingCountry>
)
</materialsCitation>
.
<materialsCitation id="25B102BC0FD401439FB42FFE47F3F481" collectingDate="1965-01-16" collectingDateMax="1965-01-21" collectingDateMin="1965-01-16" collectionCode="NMSA" collectorName="Transvaal Museum Expedition" country="South Africa" county="Blouberg" latitude="-23.8206" location="Glenferness" longLatPrecision="1" longitude="29.789566" municipality="Northern" specimenCode="DIP 028189" specimenCount="1" specimenCount-male="1" stateProvince="Limpopo">
<collectingCountry id="079B376B0DCC29B69F26535A73E2AFA4" name="South Africa">South Africa</collectingCountry>
<materialsCitation id="CE6F3CD91BD995E07DC33943E8EC5C13" collectingDate="1965-01-16" collectingDateMax="1965-01-21" collectingDateMin="1965-01-16" collectionCode="NMSA" collectorName="Transvaal Museum Expedition" country="South Africa" county="Blouberg" latitude="-23.8206" location="Glenferness" longLatPrecision="1" longitude="29.789566" municipality="Northern" specimenCode="DIP 028189" specimenCount="1" specimenCount-male="1" stateProvince="Limpopo">
<specimenCount id="30ED9ABC30A76BBFAB6C44323E22E822" count="1" type="male">1 ♂</specimenCount>
;
<collectingRegion id="84C5E84626CFC512E170FB19B1F38119" country="South Africa" name="Limpopo">Limpopo</collectingRegion>
@ -686,9 +684,9 @@ side,
<location id="164B5EB4BD799970A2A7F509D571E248" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:164B5EB4BD799970A2A7F509D571E248" country="South Africa" county="Blouberg" latitude="-23.8206" longLatPrecision="1" longitude="29.789566" municipality="Northern" name="Glenferness" stateProvince="Limpopo">Glenferness</location>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJMAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[29.789567,-23.820600]}">
<geoCoordinate id="3B94539499CF4F26F594BD7ADCDA4A2D" degrees="23" direction="south" minutes="49.236" orientation="latitude" precision="1" value="-23.8206">23 ° 49.236 ' S</geoCoordinate>
<geoCoordinate id="3B94539499CF4F26F594BD7ADCDA4A2D" degrees="23" direction="south" minutes="49.236" orientation="latitude" precision="1" value="-23.8206">23°49.236'S</geoCoordinate>
,
<geoCoordinate id="4FAA1DAFE452CB70E7DA689AA860452C" degrees="29" direction="east" minutes="47.374" orientation="longitude" precision="1" value="29.789566">29 ° 47.374 ' E</geoCoordinate>
<geoCoordinate id="4FAA1DAFE452CB70E7DA689AA860452C" degrees="29" direction="east" minutes="47.374" orientation="longitude" precision="1" value="29.789566">29°47.374'E</geoCoordinate>
</named-content>
]
<date id="EBAFF17A1251C147AC3C3D644AD696B4" value="1965-01-16" valueMax="1965-01-21" valueMin="1965-01-16">
@ -705,7 +703,7 @@ side,
)
</materialsCitation>
<materialsCitation id="52984EDA5248A325A6ABCDEBB3633EE2" collectingDate="2017-01-20" collectingDateMax="2017-01-24" collectingDateMin="2017-01-20" collectionCode="BMSA" collectorName="B. S. Muller &amp; A. H. Kirk-Spriggs &amp; Legogote Sour Bushveld" country="Zimbabwe" elevation="850" latitude="-24.595554" location="Motlasedi River" longLatPrecision="1" longitude="30.888056" municipality="Mariepskop State Forest" specimenCount="3" specimenCount-female="3" stateProvince="Mpumalanga">
<materialsCitation id="C0EF71E56A4281557FA23DEE88843345" collectingDate="2017-01-20" collectingDateMax="2017-01-24" collectingDateMin="2017-01-20" collectionCode="BMSA" collectorName="B. S. Muller &amp; A. H. Kirk-Spriggs &amp; Legogote Sour Bushveld" country="Zimbabwe" elevation="850" latitude="-24.595554" location="Motlasedi River" longLatPrecision="1" longitude="30.888056" municipality="Mariepskop State Forest" specimenCount="3" specimenCount-female="3" stateProvince="Mpumalanga">
<specimenCount id="ED9227CCE0A458ACA1CCA7DB92AC65DB" count="3" type="female">3 ♀</specimenCount>
;
<collectingRegion id="D18BAA7C4FB3CA30AFE0C812F7911DA5" country="South Africa" name="Mpumalanga">Mpumalanga</collectingRegion>
@ -715,9 +713,9 @@ side,
<location id="28D8F2B6373D92256C18B7C3540595D8" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:28D8F2B6373D92256C18B7C3540595D8" country="Zimbabwe" latitude="-24.595554" longLatPrecision="1" longitude="30.888056" municipality="Mariepskop State Forest" name="Motlasedi River" stateProvince="Mpumalanga">Motlasedi River</location>
at:;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVMAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[30.888055,-24.595555]}">
<geoCoordinate id="42FB0B7BD13F7708B24D3D22455E0C7B" degrees="24" direction="south" minutes="35.7333" orientation="latitude" precision="1" value="-24.595554">24 ° 35.7333 ' S</geoCoordinate>
<geoCoordinate id="42FB0B7BD13F7708B24D3D22455E0C7B" degrees="24" direction="south" minutes="35.7333" orientation="latitude" precision="1" value="-24.595554">24°35.7333'S</geoCoordinate>
,
<geoCoordinate id="C51A2346522BA660A3E25796C0A8EE4B" degrees="30" direction="east" minutes="53.2833" orientation="longitude" precision="1" value="30.888056">30 ° 53.2833 ' E</geoCoordinate>
<geoCoordinate id="C51A2346522BA660A3E25796C0A8EE4B" degrees="30" direction="east" minutes="53.2833" orientation="longitude" precision="1" value="30.888056">30°53.2833'E</geoCoordinate>
</named-content>
;
<quantity id="DDA9530C3FF03DA911EAE05C2073AC94" metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">
@ -747,11 +745,12 @@ leg.; streambed and grassy floodplain,
<named-content id="3157836DF5BD7E921737B5B105F1037F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-bloemfontein" xlink_title="National Museum, Bloemfontein">
<collectionCode id="09D2764B2785BCD9F77E6AF17E447743" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" type="">BMSA</collectionCode>
</named-content>
(D) 124627).
<collectingCountry id="9FA79ABE23DA8CF6AA2961D228A6CA1F" name="Zimbabwe">Zimbabwe</collectingCountry>
(D) 124627)
</materialsCitation>
.
<materialsCitation id="09C7ED998322E21EFBA7B18361A091CB" collectingDate="1931-04-12" collectionCode="NMSA" collectorName="P. A. Sheppard" country="Zimbabwe" latitude="-18.966667" location="Umtali District" longLatPrecision="13" longitude="32.633335" specimenCode="DIP 028202, DIP 158390, DIP 158389, DIP 209061" specimenCount="5" specimenCount-female="5" stateProvince="Manicaland">
<collectingCountry id="9FA79ABE23DA8CF6AA2961D228A6CA1F" name="Zimbabwe">Zimbabwe</collectingCountry>
<materialsCitation id="8DA67CE1B2D74D6EF3DD3A6EBF3AAA72" collectingDate="1931-04-12" collectionCode="NMSA" collectorName="P. A. Sheppard" country="Zimbabwe" latitude="-18.966667" location="Umtali District" longLatPrecision="13" longitude="32.633335" specimenCode="DIP 028202, DIP 158390, DIP 158389, DIP 209061" specimenCount="5" specimenCount-female="5" stateProvince="Manicaland">
<specimenCount id="30B55056E087C5F44E395BD39D0E325B" count="4" type="female">4 ♀</specimenCount>
; [
<collectingRegion id="5B8029A45C07C9241B8830B1B3FDC53E" country="Zimbabwe" name="Manicaland">Manicaland Province</collectingRegion>
@ -761,9 +760,9 @@ leg.; streambed and grassy floodplain,
<collectingCountry id="750E1929D3230BD2FA3FEE3D9BF90305" name="Zimbabwe">Rhodesia</collectingCountry>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJNAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[32.633333,-18.966667]}">
<geoCoordinate id="A903F1C9D96B3ABB7E961FC2DD972636" degrees="18" direction="south" minutes="58.00" orientation="latitude" precision="9" value="-18.966667">18 ° 58.00 ' S</geoCoordinate>
<geoCoordinate id="A903F1C9D96B3ABB7E961FC2DD972636" degrees="18" direction="south" minutes="58.00" orientation="latitude" precision="9" value="-18.966667">18°58.00'S</geoCoordinate>
,
<geoCoordinate id="3A54588FED73493DB89CFF7CE625C379" degrees="32" direction="east" minutes="38.00" orientation="longitude" precision="9" value="32.633335">32 ° 38.00 ' E</geoCoordinate>
<geoCoordinate id="3A54588FED73493DB89CFF7CE625C379" degrees="32" direction="east" minutes="38.00" orientation="longitude" precision="9" value="32.633335">32°38.00'E</geoCoordinate>
</named-content>
];
<date id="76894CAA14473FADDB325C6B130FE07F" value="1931-04-12">
@ -810,9 +809,9 @@ leg.; (
<collectingCountry id="11A5F845C644868D6A470ACD059E2679" name="Zimbabwe">Rhodesia</collectingCountry>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBOAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[32.633333,-18.966667]}">
<geoCoordinate id="58654B7C223A767812CC4518D5DD36BB" degrees="18" direction="south" minutes="58.00" orientation="latitude" precision="9" value="-18.966667">18 ° 58.00 ' S</geoCoordinate>
<geoCoordinate id="58654B7C223A767812CC4518D5DD36BB" degrees="18" direction="south" minutes="58.00" orientation="latitude" precision="9" value="-18.966667">18°58.00'S</geoCoordinate>
,
<geoCoordinate id="B3658B09ABFDFBEF92258428478498D5" degrees="32" direction="east" minutes="38.00" orientation="longitude" precision="9" value="32.633335">32 ° 38.00 ' E</geoCoordinate>
<geoCoordinate id="B3658B09ABFDFBEF92258428478498D5" degrees="32" direction="east" minutes="38.00" orientation="longitude" precision="9" value="32.633335">32°38.00'E</geoCoordinate>
</named-content>
];
<date id="7BC4F02A6AD24606B93568D5656507FF" value="1931-02-24">
@ -839,9 +838,9 @@ leg.; (
<collectingCountry id="CD8BE2E426C9544BE1796673A93A70ED" name="Zimbabwe">Rhodesia</collectingCountry>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENOAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[32.633333,-18.966667]}">
<geoCoordinate id="12738C207FAC4D36CE485B0395299B6A" degrees="18" direction="south" minutes="58.00" orientation="latitude" precision="9" value="-18.966667">18 ° 58.00 ' S</geoCoordinate>
<geoCoordinate id="12738C207FAC4D36CE485B0395299B6A" degrees="18" direction="south" minutes="58.00" orientation="latitude" precision="9" value="-18.966667">18°58.00'S</geoCoordinate>
,
<geoCoordinate id="FB5E4851BB15FAF9B23EFFBF2C269837" degrees="32" direction="east" minutes="38.00" orientation="longitude" precision="9" value="32.633335">32 ° 38.00 ' E</geoCoordinate>
<geoCoordinate id="FB5E4851BB15FAF9B23EFFBF2C269837" degrees="32" direction="east" minutes="38.00" orientation="longitude" precision="9" value="32.633335">32°38.00'E</geoCoordinate>
</named-content>
];
<date id="F27DA8C3D99AC843ED3CE936993B93BA" value="1930-02-16">
@ -868,9 +867,9 @@ leg.; (
<location id="3299B5564374E0FBB83BBBC5018075ED" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:3299B5564374E0FBB83BBBC5018075ED" country="Zimbabwe" latitude="-19.1" longLatPrecision="13" longitude="32.783333" name="Vumba" stateProvince="Manicaland">Vumba</location>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZOAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[32.783333,-19.100000]}">
<geoCoordinate id="C1F1904A1A04FD7DA5CA2ADEC0EAA50D" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19 ° 06.00 ' S</geoCoordinate>
<geoCoordinate id="C1F1904A1A04FD7DA5CA2ADEC0EAA50D" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19°06.00'S</geoCoordinate>
,
<geoCoordinate id="834C5B5184EBF0B06EA94DDE8F29811D" degrees="32" direction="east" minutes="47.00" orientation="longitude" precision="9" value="32.783333">32 ° 47.00 ' E</geoCoordinate>
<geoCoordinate id="834C5B5184EBF0B06EA94DDE8F29811D" degrees="32" direction="east" minutes="47.00" orientation="longitude" precision="9" value="32.783333">32°47.00'E</geoCoordinate>
</named-content>
];
<date id="363B3F9E4B9B4C07E4FDCD5801540078" value="1965-03-07">
@ -898,9 +897,9 @@ leg.; (
[ain]., near
<location id="BC44E613ADFD96702F06CA8D223D68E5" LSID="urn:lsid:plazi:treatment:8F5BAEF55B745DEB816C739831546EB2:BC44E613ADFD96702F06CA8D223D68E5" country="Zimbabwe" latitude="-19.1" longLatPrecision="874" longitude="32.783333" municipality="Vumba Mount" name="Umtali" stateProvince="Manicaland">Umtali</location>
; [
<geoCoordinate id="AA16B04F80B8DB810246E2D5D95E0957" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19 ° 06.00 ' S</geoCoordinate>
<geoCoordinate id="AA16B04F80B8DB810246E2D5D95E0957" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19°06.00'S</geoCoordinate>
,
<geoCoordinate id="557D331C646531B2FD679A94AEF098C6" degrees="32" direction="east" minutes="47" orientation="longitude" precision="925" value="32.783333">32 ° 47 ' E</geoCoordinate>
<geoCoordinate id="557D331C646531B2FD679A94AEF098C6" degrees="32" direction="east" minutes="47" orientation="longitude" precision="925" value="32.783333">32°47'E</geoCoordinate>
];
<date id="905A559AFDE30645EBE461C61E8D02E4" value="1955-01-18">
<collectingDate id="56483AD97DB704DDD1AB45D55ADBE3CD" value="1955-01-18">18 Jan. 1955</collectingDate>
@ -926,9 +925,9 @@ leg.; (
<collectingCountry id="831D7E3C701F8E5E63CFB48A9DB1723C" name="Zimbabwe">Rhodesia</collectingCountry>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJPAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[32.783333,-19.100000]}">
<geoCoordinate id="3E9CF2557CFAEF09F32D9A99C46070BB" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19 ° 06.00 ' S</geoCoordinate>
<geoCoordinate id="3E9CF2557CFAEF09F32D9A99C46070BB" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19°06.00'S</geoCoordinate>
,
<geoCoordinate id="7241568BB710864ADDD13D13D359C3B5" degrees="32" direction="east" minutes="47.00" orientation="longitude" precision="9" value="32.783333">32 ° 47.00 ' E</geoCoordinate>
<geoCoordinate id="7241568BB710864ADDD13D13D359C3B5" degrees="32" direction="east" minutes="47.00" orientation="longitude" precision="9" value="32.783333">32°47.00'E</geoCoordinate>
</named-content>
];
<date id="C67C5E128B28BC58617B8DF060B34BC6" value="1965-03-11">
@ -953,9 +952,9 @@ leg. (
<collectingCountry id="987D8522BD141ABCAC63ED1E95A2AB66" name="Zimbabwe">Rhodesia</collectingCountry>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVPAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[32.783333,-19.100000]}">
<geoCoordinate id="8BB2D0B4D8D8C96AE647B62F887ABBBD" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19 ° 06.00 ' S</geoCoordinate>
<geoCoordinate id="8BB2D0B4D8D8C96AE647B62F887ABBBD" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19°06.00'S</geoCoordinate>
,
<geoCoordinate id="4EF321A3E77F29056AA303281CF979AE" degrees="32" direction="east" minutes="47.00" orientation="longitude" precision="9" value="32.783333">32 ° 47.00 ' E</geoCoordinate>
<geoCoordinate id="4EF321A3E77F29056AA303281CF979AE" degrees="32" direction="east" minutes="47.00" orientation="longitude" precision="9" value="32.783333">32°47.00'E</geoCoordinate>
</named-content>
];
<date id="A65060AB4292187272EDE76D8FF59DD6" value="1965-10-14">
@ -980,9 +979,9 @@ leg. (
<collectingCountry id="39709C3C84A92D45E3C404365951DAA0" name="Zimbabwe">Rhodesia</collectingCountry>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBQAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[32.783333,-19.100000]}">
<geoCoordinate id="3DC89FF6F1E16181D45BEEC0E7538E2D" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19 ° 06.00 ' S</geoCoordinate>
<geoCoordinate id="3DC89FF6F1E16181D45BEEC0E7538E2D" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19°06.00'S</geoCoordinate>
,
<geoCoordinate id="123D52A9714724763499D5BA5D8B5435" degrees="32" direction="east" minutes="47.00" orientation="longitude" precision="9" value="32.783333">32 ° 47.00 ' E</geoCoordinate>
<geoCoordinate id="123D52A9714724763499D5BA5D8B5435" degrees="32" direction="east" minutes="47.00" orientation="longitude" precision="9" value="32.783333">32°47.00'E</geoCoordinate>
</named-content>
];
<date id="E37CE3163071BFF17C97E5AE26EB02A7" value="1965-03-13">
@ -1007,9 +1006,9 @@ leg.; (
<collectingCountry id="90B04FD428F113E05BFA4E388A308DBE" name="Zimbabwe">Rhodesia</collectingCountry>
; [
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENQAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[32.783333,-19.100000]}">
<geoCoordinate id="A85552ADA147A0276B081978A255AF8C" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19 ° 06.00 ' S</geoCoordinate>
<geoCoordinate id="A85552ADA147A0276B081978A255AF8C" degrees="19" direction="south" minutes="06.00" orientation="latitude" precision="9" value="-19.1">19°06.00'S</geoCoordinate>
,
<geoCoordinate id="4C8B2F4655E7C0BF9C5C8F2B1D901F7A" degrees="32" direction="east" minutes="47.00" orientation="longitude" precision="9" value="32.783333">32 ° 47.00 ' E</geoCoordinate>
<geoCoordinate id="4C8B2F4655E7C0BF9C5C8F2B1D901F7A" degrees="32" direction="east" minutes="47.00" orientation="longitude" precision="9" value="32.783333">32°47.00'E</geoCoordinate>
</named-content>
];
<date id="6ECAC7643B6C6B2813F1D10DE3E39D17" value="1965-03-29">