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<mods:title id="5D8DCDF99E2258468CB1CF3DDC4F268F">Accumulations of Fossils of the Whale Barnacle Coronula bifida Bronn, 1831 (Thoracica: Coronulidae) Provides Evidence of a Late Pliocene Cetacean Migration Route through the Straits of Taiwan</mods:title>
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<mods:namePart id="9AEA262AF00BF9EAB06450E123398A12">Buckeridge, John Stewart</mods:namePart>
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<taxonomicName id="8189A7F6765FFF9CFC3823F5FAC0FD3F" authority="Bronn, 1831" authorityName="Bronn" authorityYear="1831" box="[935,1291,764,790]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bifida">
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<emphasis id="74FD0067765FFF9CFC3823F5FBA0FD3F" bold="true" box="[935,1131,764,790]" italics="true" pageId="3" pageNumber="4">Coronula bifida</emphasis>
<bibRefCitation id="2218A184765FFF9CFBED23F4FAC0FD3F" author="Bronn" box="[1138,1291,765,790]" firstAuthor="Bronn" pageId="3" pageNumber="4" refId="ref4660" refString="Bronn HG. 1831. Ubersicht der Fossilen Uberreste in den tertiaren subappeninischen Gebirgen. Italiens Tertiar- Gebilde und deren organische Einschlusse. Karl Groos, Heidelberg." type="book" year="1831">Bronn, 1831</bibRefCitation>
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(
<figureCitation id="DEB2C0F0765FFF9CFBA92214FBB7FD1F" box="[1078,1148,797,822]" captionStart="Fig" captionStartId="4.[174,208,1924,1944]" captionTargetBox="[213,1412,249,1898]" captionTargetId="figure-31@4.[213,1412,249,1898]" captionTargetPageId="4" captionText="Fig. 3. Coronula bifida Bronn, 1831 from Niupu, Chiayi, Western Central Taiwan, Liuchungchi Formation (Plio-Pleistocene). Specimens (#0860-0869) held by the Chia-Yi Municipal Museum, Chaiyi City, Taiwan." figureDoi="http://doi.org/10.5281/zenodo.13316508" httpUri="https://zenodo.org/record/13316508/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
)
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<treatmentCitation id="C728FA64765FFF9CFCB2225EFBA2FD45" author="Bronn" box="[813,1129,855,876]" page="126" pageId="3" pageNumber="4" year="1831">
<taxonomicName id="8189A7F6765FFF9CFCB2225EFBA2FD45" authority="Bronn, 1831: 126" authorityName="Bronn" authorityPageNumber="126" authorityYear="1831" box="[813,1129,855,876]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD0067765FFF9CFCB2225EFC77FD45" box="[813,956,855,876]" italics="true" pageId="3" pageNumber="4">Coronula bifida</emphasis>
<bibRefCitation id="2218A184765FFF9CFC5D2251FBA2FD45" author="Bronn" box="[962,1129,856,876]" firstAuthor="Bronn" page="126" pageId="3" pageNumber="4" refId="ref4660" refString="Bronn HG. 1831. Ubersicht der Fossilen Uberreste in den tertiaren subappeninischen Gebirgen. Italiens Tertiar- Gebilde und deren organische Einschlusse. Karl Groos, Heidelberg." type="book" year="1831">Bronn, 1831: 126</bibRefCitation>
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.
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<treatmentCitationGroup id="6699FB5B765FFF9CFCB2227BFB52FDAF" box="[813,1177,882,902]" pageId="3" pageNumber="4">
<treatmentCitation id="C728FA64765FFF9CFCB2227BFB5EFDAF" author="Darwin" box="[813,1173,882,902]" page="421" pageId="3" pageNumber="4" year="1854">
<taxonomicName id="8189A7F6765FFF9CFCB2227BFB5EFDAF" authority="Darwin, 1854 a: 421" authorityName="Darwin" authorityPageNumber="421" authorityYear="1854" box="[813,1173,882,902]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="barbara">
<emphasis id="74FD0067765FFF9CFCB2227BFC18FDAF" box="[813,979,882,902]" italics="true" pageId="3" pageNumber="4">Coronula barbara</emphasis>
<bibRefCitation id="2218A184765FFF9CFC46227BFB5EFDAF" author="Darwin" box="[985,1173,882,902]" firstAuthor="Darwin" page="421" pageId="3" pageNumber="4" refId="ref5006" refString="Darwin C. 1854 a. A Monograph on the fossil Balanidae and Verrucidae of Great Britain. Palaeontographical Society, London." type="book" year="1854" yearSuffix="a">Darwin, 1854a: 421</bibRefCitation>
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.
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<treatmentCitationGroup id="6699FB5B765FFF9CFCB22284FACAFD88" box="[813,1281,909,929]" pageId="3" pageNumber="4">
<treatmentCitation id="C728FA64765FFF9CFCB22284FB36FD88" author="Pilsbry and Olsson" box="[813,1277,909,929]" page="202" pageId="3" pageNumber="4" year="1951">
<taxonomicName id="8189A7F6765FFF9CFCB22284FB36FD88" authority="Pilsbry and Olsson, 1951: 202" authorityName="Pilsbry and Olsson" authorityPageNumber="202" authorityYear="1951" box="[813,1277,909,929]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dormitor">
<emphasis id="74FD0067765FFF9CFCB22284FC1CFD88" box="[813,983,909,929]" italics="true" pageId="3" pageNumber="4">Coronula dormitor</emphasis>
<bibRefCitation id="2218A184765FFF9CFC422284FB36FD88" author="Pilsbry and Olsson" box="[989,1277,909,929]" firstAuthor="Pilsbry" page="202" pageId="3" pageNumber="4" pagination="197 - 210" refId="ref5682" refString="Pilsbry HA, Olsson AA. 1951. Tertiary and Cretaceous Cirripedia from northwestern South America. Acad Nat Sci Phila 103: 197 - 210." type="journal article" year="1951">Pilsbry and Olsson, 1951: 202</bibRefCitation>
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<treatmentCitationGroup id="6699FB5B765FFF9CFCB222AEFB09FD95" box="[813,1218,935,956]" pageId="3" pageNumber="4">
<treatmentCitation id="C728FA64765FFF9CFCB222AEFB75FD95" author="Menesini" box="[813,1214,935,956]" page="387" pageId="3" pageNumber="4" year="1968">
<taxonomicName id="8189A7F6765FFF9CFCB222AEFB75FD95" authority="Menesini, 1968: 387" authorityName="Menesini" authorityPageNumber="387" authorityYear="1968" box="[813,1214,935,956]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="bifida" subSpecies="bifida">
<emphasis id="74FD0067765FFF9CFCB222AEFC3EFD95" box="[813,1013,935,956]" italics="true" pageId="3" pageNumber="4">Coronula bifida bifida</emphasis>
<bibRefCitation id="2218A184765FFF9CFC6422A1FB75FD95" author="Menesini" box="[1019,1214,936,956]" firstAuthor="Menesini" page="387" pageId="3" pageNumber="4" pagination="387 - 398" refId="ref5595" refString="Menesini E. 1968. Osservazioni su Coronula bifida Bronn. Atti della Societa Toscana di Scienze Naturali, Memorie, Serie A 75: 387 - 398." type="journal article" year="1968">Menesini, 1968: 387</bibRefCitation>
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.
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<treatmentCitationGroup id="6699FB5B765FFF9CFCB222CBFB12FDFF" box="[813,1241,962,982]" pageId="3" pageNumber="4">
<treatmentCitation id="C728FA64765FFF9CFCB222CBFB1EFDFF" author="Menesini" box="[813,1237,962,982]" page="395" pageId="3" pageNumber="4" year="1968">
<taxonomicName id="8189A7F6765FFF9CFCB222CBFB1EFDFF" authority="Menesini, 1968: 395" authorityName="Menesini" authorityPageNumber="395" authorityYear="1968" box="[813,1237,962,982]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="bifida" subSpecies="barbara">
<emphasis id="74FD0067765FFF9CFCB222CBFBC7FDFF" box="[813,1036,962,982]" italics="true" pageId="3" pageNumber="4">Coronula bifida barbara</emphasis>
<bibRefCitation id="2218A184765FFF9CFB8E22CBFB1EFDFF" author="Menesini" box="[1041,1237,962,982]" firstAuthor="Menesini" page="395" pageId="3" pageNumber="4" pagination="387 - 398" refId="ref5595" refString="Menesini E. 1968. Osservazioni su Coronula bifida Bronn. Atti della Societa Toscana di Scienze Naturali, Memorie, Serie A 75: 387 - 398." type="journal article" year="1968">Menesini, 1968: 395</bibRefCitation>
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<treatmentCitationGroup id="6699FB5B765FFF9CFCB222D4FB76FDD8" box="[813,1213,989,1009]" pageId="3" pageNumber="4">
<treatmentCitation id="C728FA64765FFF9CFCB222D4FB71FDD8" author="Collareta" box="[813,1210,989,1009]" page="15" pageId="3" pageNumber="4" year="2018">
<taxonomicName id="8189A7F6765FFF9CFCB222D4FB71FDD8" authority="Collareta et al., 2018 a: 15" authorityName="Collareta" authorityPageNumber="15" authorityYear="2018" box="[813,1210,989,1009]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD0067765FFF9CFCB222D4FC77FDD8" box="[813,956,989,1009]" italics="true" pageId="3" pageNumber="4">Coronula bifida</emphasis>
<bibRefCitation id="2218A184765FFF9CFC5D22D4FB71FDD8" author="Collareta" box="[962,1210,989,1009]" etAl="et al." firstAuthor="Collareta" page="15" pageId="3" pageNumber="4" pagination="9 - 22" refId="ref4875" refString="Collareta A, Insacco G, Reitano A, Catanzariti R, Bosselaers M, Montes M, Bianucci G. 2018 a. Fossil whale barnacles from the lower Pleistocene of Sicily shed light on the coeval Mediterranean cetacean fauna. Carnets de geologie 18: 9 - 22. doi: 10.1016 / j. crpv. 2015.10.006." type="journal article" year="2018" yearSuffix="a">Collareta et al., 2018a: 15</bibRefCitation>
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<paragraph id="4636DC75765FFF9CFCB222FEFB08FA25" blockId="3.[813,1281,855,1036]" box="[813,1219,1015,1036]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="6699FB5B765FFF9CFCB222FEFB08FA25" box="[813,1219,1015,1036]" pageId="3" pageNumber="4">
<treatmentCitation id="C728FA64765FFF9CFCB222FEFB08FA25" author="Collareta" box="[813,1219,1015,1036]" page="147" pageId="3" pageNumber="4" year="2018">
<taxonomicName id="8189A7F6765FFF9CFCB222FEFB08FA25" authority="Collareta et al., 2018 b: 147" authorityName="Collareta" authorityPageNumber="147" authorityYear="2018" box="[813,1219,1015,1036]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD0067765FFF9CFCB222FEFC77FA25" box="[813,956,1015,1036]" italics="true" pageId="3" pageNumber="4">Coronula bifida</emphasis>
<bibRefCitation id="2218A184765FFF9CFC5D22FEFB08FA25" author="Collareta" box="[962,1219,1015,1036]" etAl="et al." firstAuthor="Collareta" page="147" pageId="3" pageNumber="4" pagination="143 - 159" refId="ref4936" refString="Collareta A, Regattieri E, Zanchetta G, Lambert O, Catanzariti R, Bosselaers M, Covelo P, Varola A, Bianucci G. 2018 b. New insights on ancient cetacean movement patterns from oxygenisotope analyses of a Mediterranean Pleistocene whale barnacle. Neues Jahrbuch fur Geologie und Palaontologie-Abhandlungen 288 (2): 143 - 159. doi: 10.4267 / 2042 / 65747." type="journal article" year="2018" yearSuffix="b">Collareta et al., 2018b: 147</bibRefCitation>
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<emphasis id="74FD0067765FFF9CFCF6253AFBDDFA65" box="[873,1046,1075,1100]" italics="true" pageId="3" pageNumber="4">Distribution</emphasis>
: Late Pliocene to early Pleistocene (Piacenzian to Gelasian,
<collectingCountry id="3E9E9CE5765FFF9CFB71255AFA8AFA45" box="[1262,1345,1107,1132]" name="Taiwan" pageId="3" pageNumber="4">Taiwan</collectingCountry>
- this work); Middle Pliocene to mid-lower Pleistocene (Calabrian, Western Europe; California;
<collectingCountry id="3E9E9CE5765FFF9CFA8E259AFAB0FA85" box="[1297,1403,1171,1196]" name="Ecuador" pageId="3" pageNumber="4">Ecuador</collectingCountry>
). As
<taxonomicName id="8189A7F6765FFF9CFCC825BAFC17FAE5" authorityName="Darwin" authorityYear="1854" box="[855,988,1203,1228]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="barbara">
<emphasis id="74FD0067765FFF9CFCC825BAFC17FAE5" box="[855,988,1203,1228]" italics="true" pageId="3" pageNumber="4">C. barbara</emphasis>
</taxonomicName>
:
<bibRefCitation id="2218A184765FFF9CFC6F25BAFB27FAE5" author="Marquet" box="[1008,1260,1203,1228]" etAl="et al." firstAuthor="Marquet" pageId="3" pageNumber="4" pagination="41 - 61" refId="ref5555" refString="Marquet R, Collins J, Jagt J, van Bakel B. 2009. The invertebrate fauna of the Pliocene in Belgium. Part I. Annellida, Anthozoa and Crustacea. Palaeofocus 2: 41 - 61." type="journal article" year="2009">Marquet et al. 2009</bibRefCitation>
;
<bibRefCitation id="2218A184765FFF9CFB6525BAFA49FAE5" author="Zullo VA" box="[1274,1410,1203,1228]" pageId="3" pageNumber="4" pagination="1 - 25" refId="ref5994" refString="Zullo VA. 1969. Thoracic Cirripedia of the San Diego Formation, San Diego County, California. Contrib Sci, Natural History Museum of Los Angeles County 159: 1 - 25." type="journal article" year="1969">Zullo 1969</bibRefCitation>
;
<bibRefCitation id="2218A184765FFF9CFCB225DAFC64FAC5" author="Beu AG" box="[813,943,1235,1260]" journalOrPublisher="New Zeal J Geol Geop" pageId="3" pageNumber="4" pagination="897 - 904" part="14" refId="ref4527" refString="Beu AG. 1971. Further fossil whale barnacles from New Zealand. New Zeal J Geol Geop 14: 897 - 904." title="Further fossil whale barnacles from New Zealand" type="journal article" year="1971">Beu 1971</bibRefCitation>
<bibRefCitation id="2218A184765FFF9CFC2225DAFADCFAC5" author="Pilsbry HA &amp; Olsson AA" box="[957,1303,1235,1261]" pageId="3" pageNumber="4" pagination="197 - 210" refId="ref5682" refString="Pilsbry HA, Olsson AA. 1951. Tertiary and Cretaceous Cirripedia from northwestern South America. Acad Nat Sci Phila 103: 197 - 210." type="journal article" year="1951">Pilsbry and Olsson 1951</bibRefCitation>
;
<bibRefCitation id="2218A184765FFF9CFAB625DAFCB1FB25" author="Darwin C." pageId="3" pageNumber="4" refId="ref5029" refString="Darwin C. 1854 b. A Monograph on the Sub-Class Cirripedia. The Balanidae and Verrucidae. Ray Society, London." type="book" year="1854">Darwin 1854b</bibRefCitation>
; De Alessandri 1960
<bibRefCitation id="2218A184765FFF9CFBE025FAFA49FB25" author="Bianucci G &amp; Landini W &amp; Buckeridge JS" box="[1151,1410,1267,1292]" pageId="3" pageNumber="4" pagination="115 - 120" refId="ref4603" refString="Bianucci G, Landini W, Buckeridge JS. 2006 b. Whale barnacles and Neogene cetacean migration routes. New Zeal J Geol Geop 49: 115 - 120. doi: 10.1080 / 00288306.2006.9515152." type="journal article" year="2006">Bianucci et al. 2006b</bibRefCitation>
;
<bibRefCitation id="2218A184765FFF9CFCB2241AFBEEFB05" author="Collareta A &amp; Margiotta S &amp; Varola A &amp; Catanzariti R &amp; Bosselaers M &amp; Bianucci G." box="[813,1061,1299,1325]" pageId="3" pageNumber="4" pagination="473 - 481" refId="ref4809" refString="Collareta A, Margiotta S, Varola A, Catanzariti R, Bosselaers M, Bianucci G. 2016. A new whale barnacle from the early Pleistocene of Italy suggests an ancient right whale breeding ground in the Mediterranean. Comptes Rendus Palevol 15: 473 - 481. doi: 10.1016 / j. crpv. 2015.10.006." type="journal article" year="2016">Collareta et al. 2016</bibRefCitation>
<bibRefCitation id="2218A184765FFF9CFBB0241AFB59FB05" author="Collareta A &amp; Insacco G &amp; Reitano A &amp; Catanzariti R &amp; Bosselaers M &amp; Montes M &amp; Bianucci G." box="[1071,1170,1299,1324]" pageId="3" pageNumber="4" pagination="9 - 22" refId="ref4875" refString="Collareta A, Insacco G, Reitano A, Catanzariti R, Bosselaers M, Montes M, Bianucci G. 2018 a. Fossil whale barnacles from the lower Pleistocene of Sicily shed light on the coeval Mediterranean cetacean fauna. Carnets de geologie 18: 9 - 22. doi: 10.1016 / j. crpv. 2015.10.006." type="journal article" year="2018">2018a b</bibRefCitation>
(see full references in
<bibRefCitation id="2218A184765FFF9CFCD6243AFB86FB65" author="Collareta A &amp; Insacco G &amp; Reitano A &amp; Catanzariti R &amp; Bosselaers M &amp; Montes M &amp; Bianucci G." box="[841,1101,1331,1357]" pageId="3" pageNumber="4" pagination="9 - 22" refId="ref4875" refString="Collareta A, Insacco G, Reitano A, Catanzariti R, Bosselaers M, Montes M, Bianucci G. 2018 a. Fossil whale barnacles from the lower Pleistocene of Sicily shed light on the coeval Mediterranean cetacean fauna. Carnets de geologie 18: 9 - 22. doi: 10.1016 / j. crpv. 2015.10.006." type="journal article" year="2018">Collareta et al. 2018a</bibRefCitation>
).
</paragraph>
<paragraph id="4636DC75765FFF9CFCF6245AFB43F925" blockId="3.[813,1414,1075,1964]" pageId="3" pageNumber="4">
<emphasis id="74FD0067765FFF9CFCF6245AFB98FB45" box="[873,1107,1363,1388]" italics="true" pageId="3" pageNumber="4">Material Examined</emphasis>
:
<collectingCountry id="3E9E9CE5765FFF9CFBF9245AFB0AFB45" box="[1126,1217,1363,1388]" name="Taiwan" pageId="3" pageNumber="4">Taiwan</collectingCountry>
. 0860-0869: 10 complete shells (without opercula), 17.4-37.0 mm C-R diameter, 13.6-31.2 height; CIMFO0047- CIMFO0058 and NMMSTM000125: 13 complete shells (without opercula),
<quantity id="81717190765FFF9CFB1224DAFA8DFBC5" box="[1165,1350,1491,1516]" metricMagnitude="-2" metricUnit="m" metricValue="5.985" metricValueMax="7.85" metricValueMin="4.12" pageId="3" pageNumber="4" unit="mm" value="59.85" valueMax="78.5" valueMin="41.2">41.2-78.5 mm</quantity>
C-R diameter, 25.0-
<quantity id="81717190765FFF9CFC7824FAFB9EF825" box="[999,1109,1523,1548]" metricMagnitude="-2" metricUnit="m" metricValue="5.4799999999999995" pageId="3" pageNumber="4" unit="mm" value="54.8">54.8 mm</quantity>
height; ASIZCR000400- ASIZCR000409: 10 complete shells (without opercula) 21.0-
<quantity id="81717190765FFF9CFC67273AFBA5F865" box="[1016,1134,1587,1612]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" pageId="3" pageNumber="4" unit="mm" value="56.5">56.5 mm</quantity>
C-R diameter,
<quantity id="81717190765FFF9CFADD273AFC50F845" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="4.47" metricValueMin="1.83" pageId="3" pageNumber="4" unit="mm" value="31.5" valueMax="44.7" valueMin="18.3">18.3- 44.7 mm</quantity>
height; G.340:
<specimenCount id="508F17FC765FFF9CFBF9275AFAD3F845" box="[1126,1304,1619,1644]" count="1" pageId="3" pageNumber="4" type="generic">one specimen</specimenCount>
(without opercula), C-R diameter
<quantity id="81717190765FFF9CFBF4277AFB0CF8A5" box="[1131,1223,1651,1676]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="4" unit="mm" value="47.0">47 mm</quantity>
, height
<quantity id="81717190765FFF9CFAB1277AFA4FF8A5" box="[1326,1412,1651,1676]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="4" unit="mm" value="38.0">38 mm</quantity>
(
<tableCitation id="0B0BE9CE765FFF9CFCAC279AFC62F884" box="[819,937,1683,1709]" captionStart="Table 1" captionStartId="1.[136,205,1424,1449]" captionText="Table 1. Accumulations of Coronula shells. This table only includes fossil localities where two or more specimens have been recovered. As such, they can likely be assessed as whale congregation sites. (For a list of all localities from which fossil Coronula has been recorded, see Bianucci et al. 2006a)" pageId="3" pageNumber="4">Table S1</tableCitation>
). All specimens were collected by Mr Wen-Ji Xue between 1980 and 2000 and from an actively eroding escarpment at Chaiyi Niupu -
<geoCoordinate id="23BDBAB2765FFF9CFCB227FAFC06F925" box="[813,973,1779,1804]" degrees="23" direction="north" minutes="25" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="40.8" value="23.428">23°25'40.8&quot;N</geoCoordinate>
,
<geoCoordinate id="23BDBAB2765FFF9CFC4827FAFB4FF925" box="[983,1156,1779,1804]" degrees="120" direction="east" minutes="33" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="25.7" value="120.557144">120°33'25.7&quot;E</geoCoordinate>
.
</paragraph>
<paragraph id="4636DC75765FFF9AFCF6261AFE9AFF1F" blockId="3.[813,1414,1075,1964]" lastBlockId="5.[136,738,253,1782]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
<emphasis id="74FD0067765FFF9CFCF6261AFC27F905" box="[873,1004,1811,1836]" italics="true" pageId="3" pageNumber="4">Diagnosis</emphasis>
:
<taxonomicName id="8189A7F6765FFF9CFB9D261AFBB1F905" authorityName="Lamarck" authorityYear="1802" box="[1026,1146,1811,1836]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="74FD0067765FFF9CFB9D261AFBB1F905" box="[1026,1146,1811,1836]" italics="true" pageId="3" pageNumber="4">Coronula</emphasis>
</taxonomicName>
with globose shell; compartments with convex longitudinal parietal ribs that often bifurcate, and with strong transverse ridges; radii broad, flat, becoming narrower in their lowermost third; body chamber relatively shallow, sub-cylindrical; sheath less than half that of the total shell height.
</paragraph>
<footnote id="2592C07B765FFF9CFF1726D4FE53F9C6" box="[136,408,2013,2031]" pageId="3" pageNumber="4">
<paragraph id="4636DC75765FFF9CFF1726D4FE53F9C6" blockId="3.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="3" pageNumber="4">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="12F68CFD7658FF9BFF31268DFCAEF99D" ID-DOI="http://doi.org/10.5281/zenodo.13316508" ID-Zenodo-Dep="13316508" httpUri="https://zenodo.org/record/13316508/files/figure.png" pageId="4" pageNumber="5" startId="4.[174,208,1924,1944]" targetBox="[213,1412,249,1898]" targetPageId="4" targetType="figure">
<paragraph id="4636DC757658FF9BFF31268DFCAEF99D" blockId="4.[174,1452,1924,1972]" pageId="4" pageNumber="5">
<emphasis id="74FD00677658FF9BFF31268DFF20F9B1" bold="true" box="[174,235,1924,1944]" pageId="4" pageNumber="5">Fig. 3.</emphasis>
<taxonomicName id="8189A7F67658FF9BFF67268DFDCAF9B1" authority="Bronn, 1831" authorityName="Bronn" authorityYear="1831" box="[248,513,1924,1944]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD00677658FF9BFF67268DFE4CF9B1" box="[248,391,1924,1944]" italics="true" pageId="4" pageNumber="5">Coronula bifida</emphasis>
<bibRefCitation id="2218A1847658FF9BFE12268DFDCAF9B1" author="Bronn HG" box="[397,513,1924,1944]" pageId="4" pageNumber="5" refId="ref4660" refString="Bronn HG. 1831. Ubersicht der Fossilen Uberreste in den tertiaren subappeninischen Gebirgen. Italiens Tertiar- Gebilde und deren organische Einschlusse. Karl Groos, Heidelberg." type="book" year="1831">Bronn, 1831</bibRefCitation>
</taxonomicName>
from Niupu, Chiayi, Western Central Taiwan, Liuchungchi Formation (Plio-Pleistocene). Specimens (#0860-0869) held by the Chia-Yi Municipal Museum, Chaiyi City, Taiwan.
</paragraph>
</caption>
<footnote id="2592C07B7658FF9BFB0326D4FA60F9C6" box="[1180,1451,2013,2031]" pageId="4" pageNumber="5">
<paragraph id="4636DC757658FF9BFB0326D4FA60F9C6" blockId="4.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="4" pageNumber="5">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="4636DC757659FF9AFF5A2034FEA4FCFF" blockId="5.[136,738,253,1782]" pageId="5" pageNumber="6">
<emphasis id="74FD00677659FF9AFF5A2034FE9AFF7F" box="[197,337,317,342]" italics="true" pageId="5" pageNumber="6">Description</emphasis>
: Shell globose; in adults, lower half of compartments developing five to six convex longitudinal parietal ribs; parietal ribs with strong transverse ridges in which bifurcation is rare; ridges spaced by at least their own height; basal edges of ribs crenated; orifice narrower than basal opening; compartments (
<emphasis id="74FD00677659FF9AFE2820F4FE10FC3F" box="[439,475,509,534]" italics="true" pageId="5" pageNumber="6">i.e.</emphasis>
ribs) noticeably worn in upper third of shell; radii broad, flat, summits horizontal; centrally radii are slightly wider than paries, becoming narrower in their lowermost third; body chamber relatively shallow, sub-cylindrical; sheath less than half of the total height of the shell. Opercula unknown.
</paragraph>
<paragraph id="4636DC757659FF9AFF5A23D4FF28FB5F" blockId="5.[136,738,253,1782]" pageId="5" pageNumber="6">
<emphasis id="74FD00677659FF9AFF5A23D4FEE4FCDF" box="[197,303,733,758]" italics="true" pageId="5" pageNumber="6">Remarks</emphasis>
:
<taxonomicName id="8189A7F67659FF9AFEA123D4FE3AFCDF" authorityName="Bronn" authorityYear="1831" box="[318,497,733,758]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD00677659FF9AFEA123D4FE3AFCDF" box="[318,497,733,758]" italics="true" pageId="5" pageNumber="6">Coronula bifida</emphasis>
</taxonomicName>
is close to
<taxonomicName id="8189A7F67659FF9AFDEA23D4FF39FD3F" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD00677659FF9AFDEA23D4FF39FD3F" italics="true" pageId="5" pageNumber="6">Coronula diadema</emphasis>
</taxonomicName>
; a phyletic lineage from
<taxonomicName id="8189A7F67659FF9AFDAF23F4FD52FD3F" authorityName="Bronn" authorityYear="1831" box="[560,665,765,790]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD00677659FF9AFDAF23F4FD52FD3F" box="[560,665,765,790]" italics="true" pageId="5" pageNumber="6">C. bifida</emphasis>
</taxonomicName>
to
<taxonomicName id="8189A7F67659FF9AFD5923F4FF38FD1F" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD00677659FF9AFD5923F4FF38FD1F" italics="true" pageId="5" pageNumber="6">C. diadema</emphasis>
</taxonomicName>
has been proposed by
<bibRefCitation id="2218A1847659FF9AFDB62214FECFFD7F" author="Dominici S &amp; Bartalini M &amp; Benvenuti M &amp; Balestra B." pageId="5" pageNumber="6" pagination="95 - 101" refId="ref5109" refString="Dominici S, Bartalini M, Benvenuti M, Balestra B. 2011. Large kings with small crowns: A Mediterranean Pleistocene whale barnacle. Bollettino della Societa Paleontologica Italiana, Modena 50: 95 - 101. doi: 10.4435 / BSPI. 2011.10." type="journal article" year="2011">Dominici et al. (2011: 99)</bibRefCitation>
, and as the ranges of these species are not known to overlap, this seems a reasonable conclusion. It differs from
<taxonomicName id="8189A7F67659FF9AFE572274FD9EFDBF" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[456,597,893,918]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD00677659FF9AFE572274FD9EFDBF" box="[456,597,893,918]" italics="true" pageId="5" pageNumber="6">C. diadema</emphasis>
</taxonomicName>
in its more widely spaced transverse ridges on the parietal ribs and a shallower body chamber. In general
<taxonomicName id="8189A7F67659FF9AFD5922B4FF03FDDF" authorityName="Bronn" authorityYear="1831" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD00677659FF9AFD5922B4FF03FDDF" italics="true" pageId="5" pageNumber="6">C. bifida</emphasis>
</taxonomicName>
has more parietal ribs on each compartment than observed in
<taxonomicName id="8189A7F67659FF9AFEC122F4FE22FA3F" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[350,489,1021,1046]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD00677659FF9AFEC122F4FE22FA3F" box="[350,489,1021,1046]" italics="true" pageId="5" pageNumber="6">C. diadema</emphasis>
</taxonomicName>
(
<emphasis id="74FD00677659FF9AFE6322F4FDEAFA3F" box="[508,545,1021,1046]" italics="true" pageId="5" pageNumber="6">i.e.</emphasis>
the ribs branch less frequently in the latter species). The worn (= exposed) portion of the compartments suggests that the shell was embedded into the cetacean skin by slightly more than half of its height. Specimens of
<taxonomicName id="8189A7F67659FF9AFEAD2594FE71FA9F" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[306,442,1181,1206]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD00677659FF9AFEAD2594FE71FA9F" box="[306,442,1181,1206]" italics="true" pageId="5" pageNumber="6">C. diadema</emphasis>
</taxonomicName>
in the collection at RMIT tend to show a proportionately greater worn zone, permitting us to conclude that
<taxonomicName id="8189A7F67659FF9AFE6C25D4FD9DFADF" authorityName="Bronn" authorityYear="1831" box="[499,598,1245,1270]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD00677659FF9AFE6C25D4FD9DFADF" box="[499,598,1245,1270]" italics="true" pageId="5" pageNumber="6">C. bifida</emphasis>
</taxonomicName>
was buried deeper into the whale’s skin. These observations are consistent with specimens of
<taxonomicName id="8189A7F67659FF9AFDF52414FD2AFB1F" authorityName="Bronn" authorityYear="1831" box="[618,737,1309,1334]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD00677659FF9AFDF52414FD2AFB1F" box="[618,737,1309,1334]" italics="true" pageId="5" pageNumber="6">C. bifida</emphasis>
</taxonomicName>
recovered from the Mediterranean (
<bibRefCitation id="2218A1847659FF9AFDB02434FF12FB5F" author="Collareta A &amp; Insacco G &amp; Reitano A &amp; Catanzariti R &amp; Bosselaers M &amp; Montes M &amp; Bianucci G." pageId="5" pageNumber="6" pagination="9 - 22" refId="ref4875" refString="Collareta A, Insacco G, Reitano A, Catanzariti R, Bosselaers M, Montes M, Bianucci G. 2018 a. Fossil whale barnacles from the lower Pleistocene of Sicily shed light on the coeval Mediterranean cetacean fauna. Carnets de geologie 18: 9 - 22. doi: 10.1016 / j. crpv. 2015.10.006." type="journal article" year="2018">Collareta et al. 2018a</bibRefCitation>
).
</paragraph>
<paragraph id="4636DC757659FF9AFF5A2474FD73F8DF" blockId="5.[136,738,253,1782]" pageId="5" pageNumber="6">
It is noteworthy that both
<bibRefCitation id="2218A1847659FF9AFE702474FD2AFBBF" author="Darwin C." box="[495,737,1405,1430]" pageId="5" pageNumber="6" refId="ref5006" refString="Darwin C. 1854 a. A Monograph on the fossil Balanidae and Verrucidae of Great Britain. Palaeontographical Society, London." type="book" year="1854">Darwin (1854a: 420)</bibRefCitation>
and later
<bibRefCitation id="2218A1847659FF9AFF682494FDD7FB9F" author="Dominici S &amp; Bartalini M &amp; Benvenuti M &amp; Balestra B." box="[247,540,1437,1462]" pageId="5" pageNumber="6" pagination="95 - 101" refId="ref5109" refString="Dominici S, Bartalini M, Benvenuti M, Balestra B. 2011. Large kings with small crowns: A Mediterranean Pleistocene whale barnacle. Bollettino della Societa Paleontologica Italiana, Modena 50: 95 - 101. doi: 10.4435 / BSPI. 2011.10." type="journal article" year="2011">Dominici et al. (2011: 97)</bibRefCitation>
commented that the differences between
<taxonomicName id="8189A7F67659FF9AFE3124B4FDFCFBFF" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[430,567,1469,1494]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD00677659FF9AFE3124B4FDFCFBFF" box="[430,567,1469,1494]" italics="true" pageId="5" pageNumber="6">C. diadema</emphasis>
</taxonomicName>
and
<taxonomicName id="8189A7F67659FF9AFDEA24B4FF2AFBDF" authorityName="Darwin" authorityYear="1854" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="reginae">
<emphasis id="74FD00677659FF9AFDEA24B4FF2AFBDF" italics="true" pageId="5" pageNumber="6">Coronula reginae</emphasis>
</taxonomicName>
are minimal. Darwin considered
<taxonomicName id="8189A7F67659FF9AFDF924D4FD2AFBDF" authorityName="Darwin" authorityYear="1854" box="[614,737,1501,1526]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="reginae">
<emphasis id="74FD00677659FF9AFDF924D4FD2AFBDF" box="[614,737,1501,1526]" italics="true" pageId="5" pageNumber="6">C. reginae</emphasis>
</taxonomicName>
more closely related to
<taxonomicName id="8189A7F67659FF9AFE5A24F4FDBFF83F" baseAuthorityName="Gmelin" baseAuthorityYear="1791" box="[453,628,1533,1558]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="balaenaris">
<emphasis id="74FD00677659FF9AFE5A24F4FDBFF83F" box="[453,628,1533,1558]" italics="true" pageId="5" pageNumber="6">C. balaenaris</emphasis>
</taxonomicName>
than
<taxonomicName id="8189A7F67659FF9AFD5A24F4FF24F81F" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD00677659FF9AFD5A24F4FF24F81F" italics="true" pageId="5" pageNumber="6">C. diadema</emphasis>
</taxonomicName>
, which is interesting, as
<taxonomicName id="8189A7F67659FF9AFDBD2714FD08F81F" baseAuthorityName="Gmelin" baseAuthorityYear="1791" box="[546,707,1565,1590]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="balaenaris">
<emphasis id="74FD00677659FF9AFDBD2714FD08F81F" box="[546,707,1565,1590]" italics="true" pageId="5" pageNumber="6">C. balaenaris</emphasis>
</taxonomicName>
is now recognized as
<taxonomicName id="8189A7F67659FF9AFEE32734FE3AF87F" box="[380,497,1597,1622]" class="Maxillopoda" family="Coronulidae" genus="Cetopirus" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="74FD00677659FF9AFEE32734FE3AF87F" box="[380,497,1597,1622]" italics="true" pageId="5" pageNumber="6">Cetopirus</emphasis>
</taxonomicName>
(see above). What this does is demonstrate the significant variation in whale barnacle morphology due to crowding and location on the whale. This variation is also demonstrated by the rather broad spectrum of shell morphologies in this collection of
<taxonomicName id="8189A7F67659FF9AFDD027D4FD7AF8DF" authorityName="Bronn" authorityYear="1831" box="[591,689,1757,1782]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD00677659FF9AFDD027D4FD7AF8DF" box="[591,689,1757,1782]" italics="true" pageId="5" pageNumber="6">C. bifida</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

224
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<mods:title id="CC4DB31156498C585BA2DF615BA959DE">Accumulations of Fossils of the Whale Barnacle Coronula bifida Bronn, 1831 (Thoracica: Coronulidae) Provides Evidence of a Late Pliocene Cetacean Migration Route through the Straits of Taiwan</mods:title>
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<treatment id="CE206D63765FFF9CFE852555FBDEFCFF" ID-DOI="http://doi.org/10.5281/zenodo.12859695" ID-Zenodo-Dep="12859695" LSID="urn:lsid:plazi:treatment:CE206D63765FFF9CFE852555FBDEFCFF" httpUri="http://treatment.plazi.org/id/CE206D63765FFF9CFE852555FBDEFCFF" lastPageNumber="4" pageId="3" pageNumber="4">
<subSubSection id="0E938FFE765FFF9CFE852555FD85FA5C" box="[282,590,1116,1142]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="4636DC75765FFF9CFE852555FD85FA5C" blockId="3.[191,682,1052,1142]" box="[282,590,1116,1142]" pageId="3" pageNumber="4">
<heading id="1D7E6B19765FFF9CFE852555FD85FA5C" bold="true" box="[282,590,1116,1142]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
<taxonomicName id="8189A7F6765FFF9CFE852555FD85FA5C" ID-CoL="62MGB" authority="Lamarck, 1802" authorityName="Lamarck" authorityYear="1802" box="[282,590,1116,1142]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="74FD0067765FFF9CFE852555FD85FA5C" bold="true" box="[282,590,1116,1142]" pageId="3" pageNumber="4">
<emphasis id="74FD0067765FFF9CFE852555FE44FA5F" bold="true" box="[282,399,1116,1142]" italics="true" pageId="3" pageNumber="4">Coronula</emphasis>
<bibRefCitation id="2218A184765FFF9CFE082555FD85FA5C" author="Lamarck" box="[407,590,1116,1141]" firstAuthor="Lamarck" pageId="3" pageNumber="4" pagination="461 - 464" refId="ref5361" refString="Lamarck JB. 1802. Memoire sur la Tubicinelle. Annales du Museum National d'Histoire Naturelle 1: 461 - 464." type="journal article" year="1802">Lamarck, 1802</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0E938FFE765FFF9CFF5A2595FBDEFCFF" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="4636DC75765FFF9CFF5A2595FD05FBBC" blockId="3.[136,738,1180,1973]" pageId="3" pageNumber="4">
<emphasis id="74FD0067765FFF9CFF5A2595FEF2FA9C" box="[197,313,1180,1205]" italics="true" pageId="3" pageNumber="4">Diagnosis</emphasis>
: Shell comprising six compartments of equal size, walls thin, deeply folded, with folds forming cavities, open basally; radii wide; body chamber relatively small, cup-shaped; opercular valves, paired terga and scuta, diminished in size, not articulated, together much smaller than the orifice; orifice of body chamber wider than basal opening; basis membranous. Attached to whales.
</paragraph>
<paragraph id="4636DC75765FFF9CFF5A2495FB6FFCBF" blockId="3.[136,738,1180,1973]" lastBlockId="3.[813,1414,253,726]" pageId="3" pageNumber="4">
<emphasis id="74FD0067765FFF9CFF5A2495FEFBFB9C" box="[197,304,1436,1461]" italics="true" pageId="3" pageNumber="4">Remarks</emphasis>
: The number of species in
<taxonomicName id="8189A7F6765FFF9CFDEA2495FD2AFB9C" authorityName="Lamarck" authorityYear="1802" box="[629,737,1436,1461]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="74FD0067765FFF9CFDEA2495FD2AFB9C" box="[629,737,1436,1461]" italics="true" pageId="3" pageNumber="4">Coronula</emphasis>
</taxonomicName>
is now somewhat reduced from the seven listed in
<bibRefCitation id="2218A184765FFF9CFF3424D5FDC7FBDC" author="Newman and Ross" box="[171,524,1500,1525]" firstAuthor="Newman" pageId="3" pageNumber="4" pagination="1 - 108" refId="ref5648" refString="Newman WA, Ross A. 1976. Revision of the balanomorph barnacles; including a catalog of the species. San Diego Society of Natural History Memoir 9: 1 - 108." type="journal article" year="1976">Newman and Ross (1976)</bibRefCitation>
. Of the three in
<bibRefCitation id="2218A184765FFF9CFF1724F5FE91F83C" author="Darwin's" box="[136,346,1532,1557]" firstAuthor="Darwin's" pageId="3" pageNumber="4" refId="ref5006" refString="Darwin C. 1854 a. A Monograph on the fossil Balanidae and Verrucidae of Great Britain. Palaeontographical Society, London." type="book" year="1854" yearSuffix="a">Darwin’s 1854a</bibRefCitation>
monographs, only
<taxonomicName id="8189A7F6765FFF9CFDF724F5FE00F81C" authority="(Linnaeus, 1767)" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD0067765FFF9CFDF724F5FF3BF81C" italics="true" pageId="3" pageNumber="4">Coronula diadema</emphasis>
(
<bibRefCitation id="2218A184765FFF9CFE9D2715FE0FF81C" author="Linnaeus" box="[258,452,1564,1589]" firstAuthor="Linnaeus" pageId="3" pageNumber="4" refId="ref5486" refString="Linnaeus C. 1767. Systema Naturae sive Regna Tria Naturae, secundum Classes, Ordines, Genera, Species, cum characteribus, differentiis, synonymis, locis, Tomus I. Editio duodecima, Reformata. L. Salvius, Stockholm." type="book" year="1767">Linnaeus, 1767</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="8189A7F6765FFF9CFD922715FE85F87C" authority="Darwin, 1854 a" authorityName="Darwin" authorityYear="1854" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="reginae">
<emphasis id="74FD0067765FFF9CFD922715FD2AF81C" box="[525,737,1564,1589]" italics="true" pageId="3" pageNumber="4">Coronula reginae</emphasis>
<bibRefCitation id="2218A184765FFF9CFF172735FE85F87C" author="Darwin" box="[136,334,1596,1621]" firstAuthor="Darwin" pageId="3" pageNumber="4" refId="ref5006" refString="Darwin C. 1854 a. A Monograph on the fossil Balanidae and Verrucidae of Great Britain. Palaeontographical Society, London." type="book" year="1854" yearSuffix="a">Darwin, 1854a</bibRefCitation>
</taxonomicName>
remain.
<taxonomicName id="8189A7F6765FFF9CFE522735FE88F85C" authority="(Gmelin, 1791)" baseAuthorityName="Gmelin" baseAuthorityYear="1791" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="balaenaris">
<emphasis id="74FD0067765FFF9CFE522735FD14F87C" box="[461,735,1596,1621]" italics="true" pageId="3" pageNumber="4">Coronula balaenaris</emphasis>
(
<bibRefCitation id="2218A184765FFF9CFF102755FEF0F85C" author="Gmelin" box="[143,315,1628,1654]" firstAuthor="Gmelin" pageId="3" pageNumber="4" pagination="3021 - 3910" refId="ref5228" refString="Gmelin JF. 1791. Vermes. In: Gmelin JF (ed) Caroli a Linnaei Systema Naturae per Regna Tria Naturae, Ed. 13. Tome 1 (6). GE. Beer, Lipsiae [Leipzig], pp. 3021 - 3910." type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
)
</taxonomicName>
has been assigned to
<taxonomicName id="8189A7F6765FFF9CFDF52755FE19F8BC" authority="(Morch, 1853)" baseAuthorityName="Morch" baseAuthorityYear="1853" class="Maxillopoda" family="Coronulidae" genus="Cetopirus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="complanatus">
<emphasis id="74FD0067765FFF9CFDF52755FED4F8BC" italics="true" pageId="3" pageNumber="4">Cetopirus complanatus</emphasis>
(
<bibRefCitation id="2218A184765FFF9CFEB12775FE01F8BC" author="Morch" box="[302,458,1660,1685]" firstAuthor="Morch" pageId="3" pageNumber="4" pagination="65 - 68" refId="ref5624" refString="Morch OAL. 1853. Cephalophora. In: Catalogus Conchyliorum. Typis Ludovici Kleini, Copenhagen, pp. 65 - 68." type="book chapter" year="1853">Mörch, 1853</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="8189A7F6765FFF9CFD8E2775FEF2F89C" authority="Darwin, 1854 a" authorityName="Darwin" authorityYear="1854" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="barbara">
<emphasis id="74FD0067765FFF9CFD8E2775FD2AF8BC" box="[529,737,1660,1685]" italics="true" pageId="3" pageNumber="4">Coronula barbara</emphasis>
<bibRefCitation id="2218A184765FFF9CFF172795FEF2F89C" author="Darwin" box="[136,313,1692,1717]" firstAuthor="Darwin" pageId="3" pageNumber="4" refId="ref5006" refString="Darwin C. 1854 a. A Monograph on the fossil Balanidae and Verrucidae of Great Britain. Palaeontographical Society, London." type="book" year="1854" yearSuffix="a">Darwin, 1854a</bibRefCitation>
</taxonomicName>
, is now regarded as
<taxonomicName id="8189A7F6765FFF9CFDB22795FD2AF89C" box="[557,737,1692,1717]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD0067765FFF9CFDB22795FD2AF89C" box="[557,737,1692,1717]" italics="true" pageId="3" pageNumber="4">Coronula bifida</emphasis>
</taxonomicName>
- a possibility that was contemplated by Darwin himself - see
<bibRefCitation id="2218A184765FFF9CFEAE27D5FDC1F8DC" author="Darwin" box="[305,522,1756,1781]" firstAuthor="Darwin" page="40" pageId="3" pageNumber="4" refId="ref5006" refString="Darwin C. 1854 a. A Monograph on the fossil Balanidae and Verrucidae of Great Britain. Palaeontographical Society, London." type="book" year="1854" yearSuffix="a">Darwin 1854a: 40</bibRefCitation>
(
<bibRefCitation id="2218A184765FFF9CFD8427D5FD1DF8DC" author="Menesini" box="[539,726,1756,1781]" firstAuthor="Menesini" pageId="3" pageNumber="4" pagination="387 - 398" refId="ref5595" refString="Menesini E. 1968. Osservazioni su Coronula bifida Bronn. Atti della Societa Toscana di Scienze Naturali, Memorie, Serie A 75: 387 - 398." type="journal article" year="1968">Menesini 1968</bibRefCitation>
). Species proposed since Darwin include
<taxonomicName id="8189A7F6765FFF9CFDEB27F5FDC1F91C" authority="De Gregorio, 1895" authorityName="De Gregorio" authorityYear="1895" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ficarazzensis">
<emphasis id="74FD0067765FFF9CFDEB27F5FEE9F91C" italics="true" pageId="3" pageNumber="4">Coronula ficarazzensis</emphasis>
<bibRefCitation id="2218A184765FFF9CFEB42615FDC1F91C" author="De Gregorio" box="[299,522,1820,1846]" firstAuthor="De Gregorio" pageId="3" pageNumber="4" pagination="1 - 7" refId="ref5073" refString="De Gregorio A. 1895. Note sur un asteride et un cirripede du post-Pliocene de Sicile des genres Astrogonium et Coronula. Annali di Geologia e Paleontologia di Palermo 19: 1 - 7." type="journal article" year="1895">De Gregorio, 1895</bibRefCitation>
</taxonomicName>
- now likely to be
<taxonomicName id="8189A7F6765FFF9CFF172635FDA9F97C" authority="(Collareta et al. 2018 a: 13)" baseAuthorityName="Collareta" baseAuthorityPageNumber="13" baseAuthorityYear="2018" box="[136,610,1852,1878]" class="Maxillopoda" family="Coronulidae" genus="Cetopirus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="74FD0067765FFF9CFF172635FF34F97C" box="[136,255,1852,1877]" italics="true" pageId="3" pageNumber="4">Cetopirus</emphasis>
(
<bibRefCitation id="2218A184765FFF9CFE8F2635FD91F97C" author="Collareta" box="[272,602,1852,1878]" etAl="et al." firstAuthor="Collareta" page="13" pageId="3" pageNumber="4" pagination="9 - 22" refId="ref4875" refString="Collareta A, Insacco G, Reitano A, Catanzariti R, Bosselaers M, Montes M, Bianucci G. 2018 a. Fossil whale barnacles from the lower Pleistocene of Sicily shed light on the coeval Mediterranean cetacean fauna. Carnets de geologie 18: 9 - 22. doi: 10.1016 / j. crpv. 2015.10.006." type="journal article" year="2018" yearSuffix="a">Collareta et al. 2018a: 13</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="8189A7F6765FFF9CFDEE2635FDF2F95C" authority="Pilsbry &amp; Olsson, 1951" authorityName="Pilsbry &amp; Olsson" authorityYear="1951" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dormitor">
<emphasis id="74FD0067765FFF9CFDEE2635FF3FF95C" italics="true" pageId="3" pageNumber="4">Coronula dormitor</emphasis>
<bibRefCitation id="2218A184765FFF9CFE9E2655FDF2F95C" author="Pilsbry &amp; Olsson" box="[257,569,1884,1910]" firstAuthor="Pilsbry" pageId="3" pageNumber="4" pagination="197 - 210" refId="ref5682" refString="Pilsbry HA, Olsson AA. 1951. Tertiary and Cretaceous Cirripedia from northwestern South America. Acad Nat Sci Phila 103: 197 - 210." type="journal article" year="1951">Pilsbry &amp; Olsson, 1951</bibRefCitation>
</taxonomicName>
, is
<taxonomicName id="8189A7F6765FFF9CFDF42655FF01F9BC" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD0067765FFF9CFDF42655FF01F9BC" italics="true" pageId="3" pageNumber="4">Coronula bifida</emphasis>
</taxonomicName>
, and
<taxonomicName id="8189A7F6765FFF9CFE8A2675FD16F9BC" authority="Weisbord, 1971" authorityName="Weisbord" authorityYear="1971" box="[277,733,1916,1941]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="macsotayi">
<emphasis id="74FD0067765FFF9CFE8A2675FDC7F9BC" box="[277,524,1916,1941]" italics="true" pageId="3" pageNumber="4">Coronula macsotayi</emphasis>
<bibRefCitation id="2218A184765FFF9CFD892675FD16F9BC" author="Weisbord" box="[534,733,1916,1941]" firstAuthor="Weisbord" pageId="3" pageNumber="4" pagination="87 - 96" refId="ref5942" refString="Weisbord NE. 1971. A new species of Coronula (Cirripedia) from the lower Pliocene of Venezuela. Bull Amer Paleo 60: 87 - 96." type="journal article" year="1971">Weisbord, 1971</bibRefCitation>
</taxonomicName>
, is now considered
<taxonomicName id="8189A7F6765FFF9CFEE52695FDA9F99C" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[378,610,1948,1973]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD0067765FFF9CFEE52695FDA9F99C" box="[378,610,1948,1973]" italics="true" pageId="3" pageNumber="4">Coronula diadema</emphasis>
</taxonomicName>
(
<bibRefCitation id="2218A184765FFF9CFDEB2695FC23FF3F" author="Bianucci" box="[628,1000,253,1973]" etAl="et al." firstAuthor="Bianucci" pageId="3" pageNumber="4" pagination="326 - 342" refId="ref4551" refString="Bianucci G, Di Celma C, Landini W, Buckeridge JS. 2006 a. Palaeoecology and taphonomy of an extraordinary whale barnacle accumulation from the Plio-Pleistocene of Ecuador. Palaeogeogr, Palaeoclimatol, Palaeoecol 242: 326 - 342. doi: 10.1016 / j. palaeo. 2006.07.004." type="journal article" year="2006" yearSuffix="a">Bianucci et al. 2006a b</bibRefCitation>
). This leaves two taxa from the antipodes, both fossil, and both defined by isolated plates:
<taxonomicName id="8189A7F6765FFF9CFC772034FA9AFF7F" authority="Fleming, 1959" authorityName="Fleming" authorityYear="1959" box="[1000,1361,317,342]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="aotea">
<emphasis id="74FD0067765FFF9CFC772034FB54FF7F" box="[1000,1183,317,342]" italics="true" pageId="3" pageNumber="4">Coronula aotea</emphasis>
<bibRefCitation id="2218A184765FFF9CFB382034FA9AFF7F" author="Fleming" box="[1191,1361,317,342]" firstAuthor="Fleming" pageId="3" pageNumber="4" pagination="242 - 247" refId="ref5199" refString="Fleming CA. 1959. A Pliocene whale-barnacle from Hawke's bay, New Zealand. New Zeal J Geol Geoph 2 (1): 242 - 247." type="journal article" year="1959">Fleming, 1959</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="8189A7F6765FFF9CFCB22054FAD0FF5F" authority="Buckeridge, 1983" authorityName="Buckeridge" authorityYear="1983" box="[813,1307,349,374]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="74FD0067765FFF9CFCB22054FBE6FF5F" box="[813,1069,349,374]" italics="true" pageId="3" pageNumber="4">Coronula intermedia</emphasis>
<bibRefCitation id="2218A184765FFF9CFBA62054FAD0FF5F" author="Buckeridge" box="[1081,1307,349,374]" firstAuthor="Buckeridge" pageId="3" pageNumber="4" pagination="1 - 151" refId="ref4688" refString="Buckeridge JS. 1983. The fossil barnacles (Cirripedia: Thoracica) of New Zealand and Australia. New Zealand Geological Survey Paleontological B 50: 1 - 151." type="journal article" year="1983">Buckeridge, 1983</bibRefCitation>
</taxonomicName>
. Both of these species are likely to have had a lower profile shell than
<taxonomicName id="8189A7F6765FFF9CFC2F2094FBA6FF9F" box="[944,1133,413,438]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="74FD0067765FFF9CFC2F2094FBA6FF9F" box="[944,1133,413,438]" italics="true" pageId="3" pageNumber="4">Coronula bifida</emphasis>
</taxonomicName>
or
<taxonomicName id="8189A7F6765FFF9CFB052094FAB5FF9F" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1178,1406,413,438]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD0067765FFF9CFB052094FAB5FF9F" box="[1178,1406,413,438]" italics="true" pageId="3" pageNumber="4">Coronula diadema</emphasis>
</taxonomicName>
, to lie somewhat intermediate between these and
<taxonomicName id="8189A7F6765FFF9CFCB220D4FC62FFDF" authorityName="Darwin" authorityYear="1854" box="[813,937,477,502]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="reginae">
<emphasis id="74FD0067765FFF9CFCB220D4FC62FFDF" box="[813,937,477,502]" italics="true" pageId="3" pageNumber="4">C. reginae</emphasis>
</taxonomicName>
, or perhaps
<taxonomicName id="8189A7F6765FFF9CFBDB20D4FB7DFFDF" box="[1092,1206,477,502]" class="Maxillopoda" family="Coronulidae" genus="Cetopirus" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="74FD0067765FFF9CFBDB20D4FB7DFFDF" box="[1092,1206,477,502]" italics="true" pageId="3" pageNumber="4">Cetopirus</emphasis>
</taxonomicName>
. Even with these later additions, only five species (
<emphasis id="74FD0067765FFF9CFB4A20F4FB31FC3F" box="[1237,1274,509,534]" italics="true" pageId="3" pageNumber="4">i.e.</emphasis>
<emphasis id="74FD0067765FFF9CFA9A20F4FAD9FC3C" bold="true" box="[1285,1298,509,533]" pageId="3" pageNumber="4">†</emphasis>
<taxonomicName id="8189A7F6765FFF9CFA8C20F4FAB5FC3F" authorityName="Fleming" authorityYear="1959" box="[1299,1406,509,534]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="aotea">
<emphasis id="74FD0067765FFF9CFA8C20F4FAB5FC3F" box="[1299,1406,509,534]" italics="true" pageId="3" pageNumber="4">C. aotea</emphasis>
</taxonomicName>
,
<emphasis id="74FD0067765FFF9CFCB22314FCF1FC1C" bold="true" box="[813,826,541,565]" pageId="3" pageNumber="4">†</emphasis>
<emphasis id="74FD0067765FFF9CFCA42314FB85FC1F" box="[827,1102,541,566]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="8189A7F6765FFF9CFCA42314FC61FC1F" box="[827,938,541,566]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bifida">C. bifida</taxonomicName>
,
<taxonomicName id="8189A7F6765FFF9CFC242314FB85FC1F" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[955,1102,541,566]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="diadema">C. diadema</taxonomicName>
</emphasis>
,
<emphasis id="74FD0067765FFF9CFBFD2314FBA4FC1C" bold="true" box="[1122,1135,541,565]" pageId="3" pageNumber="4">†</emphasis>
<taxonomicName id="8189A7F6765FFF9CFBEF2314FAD7FC1F" authorityName="Buckeridge" authorityYear="1983" box="[1136,1308,541,566]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="74FD0067765FFF9CFBEF2314FAD7FC1F" box="[1136,1308,541,566]" italics="true" pageId="3" pageNumber="4">C. intermedia</emphasis>
</taxonomicName>
, and
<taxonomicName id="8189A7F6765FFF9CFAF52314FC4DFC7F" authorityName="Darwin" authorityYear="1854" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="reginae">
<emphasis id="74FD0067765FFF9CFAF52314FC4DFC7F" italics="true" pageId="3" pageNumber="4">C. reginae</emphasis>
</taxonomicName>
) are currently assigned to
<taxonomicName id="8189A7F6765FFF9CFB5E2334FAE6FC7F" authorityName="Lamarck" authorityYear="1802" box="[1217,1325,573,598]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="74FD0067765FFF9CFB5E2334FAE6FC7F" box="[1217,1325,573,598]" italics="true" pageId="3" pageNumber="4">Coronula</emphasis>
</taxonomicName>
. Those with
<emphasis id="74FD0067765FFF9CFCFA2354FCB9FC5C" bold="true" box="[869,882,605,629]" pageId="3" pageNumber="4">†</emphasis>
are known only as fossil,
<taxonomicName id="8189A7F6765FFF9CFB302354FAF2FC5F" authorityName="Plio-Pleistocene" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1199,1337,605,630]" class="Maxillopoda" family="Coronulidae" genus="Coronula" kingdom="Animalia" order="Sessilia" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="74FD0067765FFF9CFB302354FAF2FC5F" box="[1199,1337,605,630]" italics="true" pageId="3" pageNumber="4">C. diadema</emphasis>
</taxonomicName>
has a record from Pliocene to Recent.
</paragraph>
<paragraph id="4636DC75765FFF9CFCF62394FBDEFCFF" blockId="3.[813,1414,253,726]" pageId="3" pageNumber="4">
<emphasis id="74FD0067765FFF9CFCF62394FBCDFC9F" box="[873,1030,669,694]" italics="true" pageId="3" pageNumber="4">Distribution</emphasis>
: Cosmopolitan, on whales. Pliocene to Recent.
</paragraph>
</subSubSection>
</treatment>
</document>