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<mods:title id="526AE776DC525900666F11A82999FDC9">A taxonomic revision of the ants of the Cardiocondyla wroughtonii group (Hymenoptera: Formicidae) with a checklist of the Cardiocondyla species of the world</mods:title>
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<treatment id="577887E6FF82FFDE6B54591EFE2C37E5" ID-DOI="http://doi.org/10.5281/zenodo.13193217" ID-Zenodo-Dep="13193217" LSID="urn:lsid:plazi:treatment:577887E6FF82FFDE6B54591EFE2C37E5" httpUri="http://treatment.plazi.org/id/577887E6FF82FFDE6B54591EFE2C37E5" lastPageId="20" lastPageNumber="133" pageId="19" pageNumber="132">
<subSubSection id="97CB657BFF82FFD96B54591EFB833704" pageId="19" pageNumber="132" type="nomenclature">
<paragraph id="DF6E36F0FF82FFD96B54591EFAAB3727" blockId="19.[847,1283,833,857]" box="[847,1283,833,857]" pageId="19" pageNumber="132">
<heading id="8426819CFF82FFD96B54591EFAAB3727" box="[847,1283,833,857]" fontSize="10" level="2" pageId="19" pageNumber="132" reason="4">
<taxonomicName id="18D14D73FF82FFD96B54591EFAAB3727" authority="Seifert 2003" authorityName="Seifert" authorityYear="2003" box="[847,1283,833,857]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="shagrinata">
<emphasis id="EDA5EAE2FF82FFD96B54591EFAAB3727" bold="true" box="[847,1283,833,857]" pageId="19" pageNumber="132">
<emphasis id="EDA5EAE2FF82FFD96B54591EFBDA3727" bold="true" box="[847,1138,833,857]" italics="true" pageId="19" pageNumber="132">Cardiocondyla shagrinata</emphasis>
<bibRefCitation id="BB404B01FF82FFD96C62591EFAAB3727" author="Seifert, B." box="[1145,1283,833,857]" pageId="19" pageNumber="132" pagination="203 - 338" refId="ref19624" refString="Seifert, B. (2003): The ant genus Cardiocondyla (Insecta: Hymenoptera: Formicidae) - a taxonomic revision of the C. elegans, C. bulgarica, C. batesii, C. nuda, C. shuckardi, C. stambuloffii, C. wroughtonii, C. emeryi, and C. minutior species groups. - Annalen des Naturhistorischen Museums Wien, Serie B, 104: 203 - 338." type="journal article" year="2003">Seifert 2003</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DF6E36F0FF82FFD96B54593DFB833704" blockId="19.[847,1067,866,890]" box="[847,1067,866,890]" pageId="19" pageNumber="132">
<heading id="8426819CFF82FFD96B54593DFB833704" bold="true" box="[847,1067,866,890]" fontSize="10" level="2" pageId="19" pageNumber="132" reason="3">
<emphasis id="EDA5EAE2FF82FFD96B54593DFB833704" bold="true" box="[847,1067,866,890]" pageId="19" pageNumber="132">
[
<typeStatus id="006A8852FF82FFD96B4E593DFC213704" box="[853,905,866,890]" pageId="19" pageNumber="132">type</typeStatus>
investigation]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF82FFD96B5459DCFC653061" pageId="19" pageNumber="132" type="description">
<paragraph id="DF6E36F0FF82FFD96B5459DCFC653061" blockId="19.[824,1438,899,1975]" pageId="19" pageNumber="132">
<materialsCitation id="6FB93CADFF82FFD96B5459DCFC613061" collectionCode="MZ" country="India" county="South Konkan" location="This" municipality="Forel" pageId="19" pageNumber="132" specimenCount="2" specimenCount-worker="2" typeStatus="holotype">
<location id="DA0E602BFF82FFD96B5459DCFC2837E5" LSID="urn:lsid:plazi:treatment:577887E6FF82FFDE6B54591EFE2C37E5:DA0E602BFF82FFD96B5459DCFC2837E5" box="[847,896,899,923]" country="India" county="South Konkan" municipality="Forel" name="This" pageId="19" pageNumber="132">This</location>
taxon has been described from
<collectingCountry id="A7C67660FF82FFD96CCF59DCFAB837E5" box="[1236,1296,899,923]" name="India" pageId="19" pageNumber="132">India</collectingCountry>
.
<location id="DA0E602BFF82FFD96D0259DCFA3537E5" LSID="urn:lsid:plazi:treatment:577887E6FF82FFDE6B54591EFE2C37E5:DA0E602BFF82FFD96D0259DCFA3537E5" box="[1305,1437,899,923]" country="India" county="South Konkan" municipality="Forel" name="Investigated" pageId="19" pageNumber="132">Investigated</location>
was the
<typeStatus id="006A8852FF82FFD96B9659FBFC4437C2" box="[909,1004,932,956]" pageId="19" pageNumber="132" type="holotype">holotype</typeStatus>
worker labelled by
<collectingMunicipality id="3F0AAC8AFF82FFD96CA159FBFB5937C2" box="[1210,1265,932,956]" pageId="19" pageNumber="132">Forel</collectingMunicipality>
<emphasis id="EDA5EAE2FF82FFD96CE559FBFB1D37A3" italics="true" pageId="19" pageNumber="132">
<taxonomicName id="18D14D73FF82FFD96CE559FBFCDB37A3" authority="Forel" authorityName="Forel" authorityYear="1890" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">C.Wroughtonii Forel</taxonomicName>
<collectingCounty id="360F4E7CFF82FFD96B96599AFB8D37A3" box="[909,1061,965,989]" pageId="19" pageNumber="132">South Konkan</collectingCounty>
(Wroughton)
</emphasis>
I/10,
<collectionCode id="B9C0AE35FF82FFD96CE5599AFA8F37A2" box="[1278,1319,965,988]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="19" pageNumber="132">MZ</collectionCode>
Lausanne; and two
<specimenCount id="C9D7FD79FF82FFD96BBB59B9FBD43780" box="[928,1148,998,1022]" count="2" pageId="19" pageNumber="132" type="worker" typeStatus="paratype">2 paratype workers</specimenCount>
with the same labelling; SMN Görlitz
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF82FFD96B545E78FB0630FF" pageId="19" pageNumber="132" type="materials_examined">
<paragraph id="DF6E36F0FF82FFD96B545E78FA353041" blockId="19.[824,1438,899,1975]" box="[847,1437,1063,1088]" pageId="19" pageNumber="132">
<emphasis id="EDA5EAE2FF82FFD96B545E78FBE63041" bold="true" box="[847,1102,1063,1087]" pageId="19" pageNumber="132">All material examined</emphasis>
. Only the
<typeStatus id="006A8852FF82FFD96CA75E77FB44303E" box="[1212,1260,1064,1088]" pageId="19" pageNumber="132">type</typeStatus>
series is known.
</paragraph>
<paragraph id="DF6E36F0FF82FFD96B545E17FB0630FF" blockId="19.[824,1438,899,1975]" pageId="19" pageNumber="132">
<emphasis id="EDA5EAE2FF82FFD96B545E17FB8C301E" bold="true" box="[847,1060,1096,1120]" pageId="19" pageNumber="132">Geographic range</emphasis>
. The
<typeStatus id="006A8852FF82FFD96C755E16FB36301F" box="[1134,1182,1097,1121]" pageId="19" pageNumber="132">type</typeStatus>
locality is situated at
<geoCoordinate id="BAE55037FF82FFD96B235E36FC3930FF" box="[824,913,1129,1153]" degrees="16.42" direction="north" orientation="latitude" pageId="19" pageNumber="132" precision="555" value="16.42">16.42°N</geoCoordinate>
,
<geoCoordinate id="BAE55037FF82FFD96B875E36FC5730FF" box="[924,1023,1129,1153]" degrees="73.390" direction="east" orientation="longitude" pageId="19" pageNumber="132" precision="55" value="73.39">73.390°E</geoCoordinate>
and
<quantity id="18299B15FF82FFD96C285E36FBDE30FF" box="[1075,1142,1129,1153]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="132" unit="m" value="100.0">100 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF82FFDE6B545ED5FECE3704" lastPageId="20" lastPageNumber="133" pageId="19" pageNumber="132" type="diagnosis">
<paragraph id="DF6E36F0FF82FFDE6B545ED5FE9C36EA" blockId="19.[824,1438,899,1975]" lastBlockId="20.[151,764,241,923]" lastPageId="20" lastPageNumber="133" pageId="19" pageNumber="132">
<emphasis id="EDA5EAE2FF82FFD96B545ED5FC1630DC" bold="true" box="[847,958,1162,1186]" pageId="19" pageNumber="132">Diagnosis</emphasis>
: --Worker (
<tableCitation id="9253034BFF82FFD96C5F5ED5FB2430DC" box="[1092,1164,1162,1186]" captionStart="Table 1" captionStartId="26.[151,204,245,266]" captionText="Table 1. Workers of the C. wroughtonii group of Orientalic-Australasian origin with mean FL/FR &lt;1.040. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="19" pageNumber="132">Tab. 1</tableCitation>
,
<figureCitation id="47EA2A75FF82FFD96C865ED5FB4730DC" box="[1181,1263,1162,1186]" captionStart="Figure 23" captionStartId="10.[151,215,1755,1777]" captionTargetBox="[151,1398,260,1705]" captionTargetPageId="10" captionText="Figure 23. Mesosoma in dorsal view.(A) Cardiocondyla obscurior, neotype, Nago / Okinawa, 3 June 2003; (B) Cardiocondyla shagrinata, paratype, South Konkan / India. Figure 2427. Cardiocondyla shagrinata, paratype, South Konkan / India. (24) head in dorsal view; (25) lateral view; (26) dorsal view; (27) head surface between inner eye margin and paramedian vertex. Figure 28. Cardiocondyla wroughtonii, Lundu / Malaysia, 27 May 2007. (28) head in dorsal view." pageId="19" pageNumber="132">Figs 23</figureCitation>
27, key). Very small, CS 426 µm. Head very short, CL/CW 1.099. Anteromedian clypeal margin notched, median occipital margin in 2 of
<specimenCount id="C9D7FD79FF82FFD96BC05EB2FBC8317B" box="[987,1120,1261,1285]" count="3" pageId="19" pageNumber="132" type="generic">3 specimens</specimenCount>
gently excavated. Postocular distance large, PoOc/CL 0.427. Frons broad (FRS/CS 0.282), frontal carinae immediately behind the FRS level slightly diverging caudad. Eye small, EYE/CS 0.222. Scape short, SL/CS 0.760. Promesonotal plane in dorsal view with a rather straight frontal margin and more concave lateral margins resulting in pronounced pronotal shoulders (
<figureCitation id="47EA2A75FF82FFD96C375F8CFB383195" box="[1068,1168,1491,1515]" captionStart="Figure 23" captionStartId="10.[151,215,1755,1777]" captionTargetBox="[151,1398,260,1705]" captionTargetPageId="10" captionText="Figure 23. Mesosoma in dorsal view.(A) Cardiocondyla obscurior, neotype, Nago / Okinawa, 3 June 2003; (B) Cardiocondyla shagrinata, paratype, South Konkan / India. Figure 2427. Cardiocondyla shagrinata, paratype, South Konkan / India. (24) head in dorsal view; (25) lateral view; (26) dorsal view; (27) head surface between inner eye margin and paramedian vertex. Figure 28. Cardiocondyla wroughtonii, Lundu / Malaysia, 27 May 2007. (28) head in dorsal view." pageId="19" pageNumber="132">Fig. 23B</figureCitation>
) in
<taxonomicName id="18D14D73FF82FFD96CFF5F8CFACF3195" authorityName="Forel" authorityYear="1890" box="[1252,1383,1491,1515]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD96CFF5F8CFACF3195" box="[1252,1383,1491,1515]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
</taxonomicName>
and
<taxonomicName id="18D14D73FF82FFD96B235FABFC083272" authorityName="Wheeler" authorityYear="1929" box="[824,928,1524,1548]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF82FFD96B235FABFC083272" box="[824,928,1524,1548]" italics="true" pageId="19" pageNumber="132">obscurior</emphasis>
</taxonomicName>
, no shoulders are developed due to a strongly convex frontal margin and rather straight lateral margins. Metanotal groove in lateral view deep (MGr/ CS 3.21 %) and with steep anterior and posterior slopes. Propodeal spines moderately long (SP/CS 0.184) and acute, slightly diverging and incurved in dorsal view and with large basal distance (SPBA/CS 0.292). Petiole rather narrow and moderately high (PeW/CS 0.279, PeH/CS 0.333). Axis of petiolar peduncle in lateral aspect deviating by 30° from the longitudinal axis of the petiole node. Postpetiole moderately wide and high (PpW/CS 0.446, PpH/CS 0.308), with a bilateral pair of anteroventral corners (but weaker than usually seen in
<taxonomicName id="18D14D73FF82FFD96B235DC0FC1333C9" authorityName="Forel" authorityYear="1890" box="[824,955,1951,1975]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD96B235DC0FC1333C9" box="[824,955,1951,1975]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
</taxonomicName>
and
<taxonomicName id="18D14D73FF82FFD96BE95DC0FBF233C9" authorityName="Wheeler" authorityYear="1929" box="[1010,1114,1951,1975]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF82FFD96BE95DC0FBF233C9" box="[1010,1114,1951,1975]" italics="true" pageId="19" pageNumber="132">obscurior</emphasis>
</taxonomicName>
); in dorsal view, the strongly convex sides meet with the concave anterior margin in a blunt corner and converge significantly more than in
<taxonomicName id="18D14D73FF85FFDE688C5B6CFEB23535" authorityName="Forel" authorityYear="1890" box="[151,282,307,331]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="133" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF85FFDE688C5B6CFEB23535" box="[151,282,307,331]" italics="true" pageId="20" pageNumber="133">wroughtonii</emphasis>
</taxonomicName>
. Sculpture on head and mesosoma compared to
<taxonomicName id="18D14D73FF85FFDE68A35B0BFE933512" authorityName="Forel" authorityYear="1890" box="[184,315,340,364]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="133" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF85FFDE68A35B0BFE933512" box="[184,315,340,364]" italics="true" pageId="20" pageNumber="133">wroughtonii</emphasis>
</taxonomicName>
and
<taxonomicName id="18D14D73FF85FFDE69605B0BFE4C3512" authorityName="Wheeler" authorityYear="1929" box="[379,484,340,364]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="133" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF85FFDE69605B0BFE4C3512" box="[379,484,340,364]" italics="true" pageId="20" pageNumber="133">obscurior</emphasis>
</taxonomicName>
very irregular and with smaller meshes of reticular sculpture (Fig. 27), without the regularly arranged, large and clearly demarcated foveolae seen in the former species. Lateral mesosoma and waist with dense reticulum the meshes of which have an inner diameter of 45 µm. Pubescence on first gaster tergite moderately long and dilute (PLG/CS 6.2 %, sqPDG 4.87). Head, mesosoma, waist, and appendages more dirty yellowish brown compared to
<taxonomicName id="18D14D73FF85FFDE6A635804FD53360D" authorityName="Forel" authorityYear="1890" box="[632,763,603,627]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="133" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF85FFDE6A635804FD53360D" box="[632,763,603,627]" italics="true" pageId="20" pageNumber="133">wroughtonii</emphasis>
</taxonomicName>
and
<taxonomicName id="18D14D73FF85FFDE68DE5823FE8536EA" authorityName="Wheeler" authorityYear="1929" box="[197,301,636,660]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="133" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF85FFDE68DE5823FE8536EA" box="[197,301,636,660]" italics="true" pageId="20" pageNumber="133">obscurior</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DF6E36F0FF85FFDE68B558C2FECE3704" blockId="20.[151,764,241,923]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE68B558C2FE6836CB" bold="true" box="[174,448,669,693]" pageId="20" pageNumber="133">Taxonomic comments.</emphasis>
Using the 16 standard morphometric characters shown in
<tableCitation id="9253034BFF85FFDE6A0C58E1FDF136A8" box="[535,601,702,726]" captionStart="Table 1" captionStartId="26.[151,204,245,266]" captionText="Table 1. Workers of the C. wroughtonii group of Orientalic-Australasian origin with mean FL/FR &lt;1.040. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="20" pageNumber="133">Tab. 1</tableCitation>
,
<taxonomicName id="18D14D73FF85FFDE6A7D58E1FD5436A8" authorityName="Seifert" authorityYear="2003" box="[614,764,702,726]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="133" phylum="Arthropoda" rank="species" species="shagrinata">
<emphasis id="EDA5EAE2FF85FFDE6A7D58E1FD5436A8" box="[614,764,702,726]" italics="true" pageId="20" pageNumber="133">C. shagrinata</emphasis>
</taxonomicName>
is almost inseparable from
<taxonomicName id="18D14D73FF85FFDE69CB5880FDFB3689" authorityName="Forel" authorityYear="1890" box="[464,595,735,759]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="133" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF85FFDE69CB5880FDFB3689" box="[464,595,735,759]" italics="true" pageId="20" pageNumber="133">wroughtonii</emphasis>
</taxonomicName>
and
<taxonomicName id="18D14D73FF85FFDE6A895880FD533689" authorityName="Wheeler" authorityYear="1929" box="[658,763,735,759]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="133" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF85FFDE6A895880FD533689" box="[658,763,735,759]" italics="true" pageId="20" pageNumber="133">obscurior</emphasis>
</taxonomicName>
and is apparently closely related. An easy separation from these species is given by the strong differences in sculpture of head and mesosoma and the pronounced pronotal shoulders.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF85FFDE68B559DCFE2C37E5" box="[174,388,899,923]" pageId="20" pageNumber="133" type="biology_ecology">
<paragraph id="DF6E36F0FF85FFDE68B559DCFE2C37E5" blockId="20.[151,764,241,923]" box="[174,388,899,923]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE68B559DCFEA337E5" bold="true" box="[174,267,899,923]" pageId="20" pageNumber="133">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,294 @@
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<paragraph id="DF6E36F0FF83FFD86B325804FB5B360D" blockId="18.[809,1267,603,627]" box="[809,1267,603,627]" pageId="18" pageNumber="131">
<heading id="8426819CFF83FFD86B325804FB5B360D" box="[809,1267,603,627]" fontSize="10" level="2" pageId="18" pageNumber="131" reason="4">
<taxonomicName id="18D14D73FF83FFD86B325804FB5B360D" authority="Donisthorpe 1930" authorityName="Donisthorpe" authorityYear="1930" box="[809,1267,603,627]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="EDA5EAE2FF83FFD86B325804FB5B360D" bold="true" box="[809,1267,603,627]" pageId="18" pageNumber="131">
<emphasis id="EDA5EAE2FF83FFD86B325804FB8A360D" bold="true" box="[809,1058,603,627]" italics="true" pageId="18" pageNumber="131">Cardiocondyla bicolor</emphasis>
Donisthorpe 1930
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DF6E36F0FF83FFD86B325823FBAF36EA" blockId="18.[809,1031,636,660]" box="[809,1031,636,660]" pageId="18" pageNumber="131">
<heading id="8426819CFF83FFD86B325823FBAF36EA" bold="true" box="[809,1031,636,660]" fontSize="10" level="2" pageId="18" pageNumber="131" reason="3">
<emphasis id="EDA5EAE2FF83FFD86B325823FBAF36EA" bold="true" box="[809,1031,636,660]" pageId="18" pageNumber="131">
[
<typeStatus id="006A8852FF83FFD86B2B5823FCC736EA" box="[816,879,636,660]" pageId="18" pageNumber="131">types</typeStatus>
investigated]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF83FFD86B3258C2FC6437C2" pageId="18" pageNumber="131" type="description">
<paragraph id="DF6E36F0FF83FFD86B3258C2FC6437C2" blockId="18.[786,1399,669,1153]" pageId="18" pageNumber="131">
This taxon was described from
<collectingCountry id="A7C67660FF83FFD86C6158C2FB1136CB" box="[1146,1209,669,693]" name="Israel" pageId="18" pageNumber="131">Israel</collectingCountry>
. Investigated was the worker
<typeStatus id="006A8852FF83FFD86B9258E1FC4036A8" box="[905,1000,702,726]" pageId="18" pageNumber="131" type="holotype">holotype</typeStatus>
labelled
<collectingCountry id="A7C67660FF83FFD86C5058E1FB1C36A8" box="[1099,1204,702,726]" name="Palestine" pageId="18" pageNumber="131">Palestine</collectingCountry>
: Drs. D.Scheinkin &amp; J.Carmin B.M.1930-163\ On
<taxonomicName id="18D14D73FF83FFD86C6F5880FB053689" box="[1140,1197,735,759]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="18" pageNumber="132" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
sycamore\
<taxonomicName id="18D14D73FF83FFD86D375880FC3F3766" authority="Donisthorpe" authorityName="Donisthorpe" authorityYear="1930" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="bicolor">bicolor Donisthorpe</taxonomicName>
\ Type « and
<specimenCount id="C9D7FD79FF83FFD86C29595FFB063766" box="[1074,1198,768,792]" count="1" pageId="18" pageNumber="131" type="worker">one worker</specimenCount>
<typeStatus id="006A8852FF83FFD86CA3595FFABF3766" box="[1208,1303,768,792]" pageId="18" pageNumber="131" type="paratype">paratype</typeStatus>
labelled
<collectingCountry id="A7C67660FF83FFD86B0D597FFCD73746" box="[790,895,800,824]" name="Palestine" pageId="18" pageNumber="131">Palestine</collectingCountry>
: Drs. D.Scheinkin &amp; J.Carmin B.M.1930-163\ On
<taxonomicName id="18D14D73FF83FFD86B21591EFCDB3727" box="[826,883,833,857]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="18" pageNumber="132" phylum="Tracheophyta" rank="genus">Ficus</taxonomicName>
sycamore\
<taxonomicName id="18D14D73FF83FFD86BF4591EFB2E3727" authority="Donis." authorityName="Donisthorpe" authorityYear="1930" box="[1007,1158,833,857]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="bicolor">bicolor Donis.</taxonomicName>
\ Cotype. Depository: BMNH London. The type sample is allocated to the
<taxonomicName id="18D14D73FF83FFD86D46593CFCD337E5" authorityName="Wheeler" authorityYear="1929" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF83FFD86D46593CFCD337E5" italics="true" pageId="18" pageNumber="131">C. obscurior</emphasis>
</taxonomicName>
cluster with p =0.9992 if run as wild-card in a LDA (see above).
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF83FFD96B32599AFE4C362C" lastPageId="19" lastPageNumber="132" pageId="18" pageNumber="131" type="materials_examined">
<paragraph id="DF6E36F0FF83FFD86B32599AFADF30FF" blockId="18.[786,1399,669,1153]" pageId="18" pageNumber="131">
<emphasis id="EDA5EAE2FF83FFD86B32599AFB9D37A3" bold="true" box="[809,1077,965,989]" pageId="18" pageNumber="131">All material examined</emphasis>
.
<materialsCitation id="6FB93CADFF83FFD86C5C599AFAFE30FF" country="Brazil" location="For" municipality="Excluding" pageId="18" pageNumber="131" specimenCount="93" specimenCount-worker="93">
Numeric phenotypical data were available in 43 samples (35 nest samples and 9 single-specimen stray samples) with
<specimenCount id="C9D7FD79FF83FFD86CA45E58FAE83061" box="[1215,1344,1031,1055]" count="93" pageId="18" pageNumber="131" type="worker">93 workers</specimenCount>
.
<location id="DA0E602BFF83FFD86D495E58FADE3060" LSID="urn:lsid:plazi:treatment:577887E6FF83FFD96B325804FB3936CB:DA0E602BFF83FFD86D495E58FADE3060" box="[1362,1398,1031,1054]" country="Brazil" municipality="Excluding" name="For" pageId="18" pageNumber="131">For</location>
details see supplementary information SI1, SI2.
<collectingMunicipality id="3F0AAC8AFF83FFD86D125E78FADF3041" box="[1289,1399,1063,1087]" pageId="18" pageNumber="131">Excluding</collectingMunicipality>
single-specimen samples with unclear separation from
<taxonomicName id="18D14D73FF83FFD86B095E35FC1030FF" authorityName="Forel" authorityYear="1890" box="[786,952,1129,1153]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF83FFD86B095E35FC1030FF" box="[786,952,1129,1153]" italics="true" pageId="18" pageNumber="131">C. wroughtonii</emphasis>
</taxonomicName>
, this material originated from
<collectingCountry id="A7C67660FF83FFD86D0D5E36FAFE30FF" box="[1302,1366,1129,1153]" name="Brazil" pageId="18" pageNumber="131">Brasil</collectingCountry>
</materialsCitation>
(3
</paragraph>
<caption id="8BAE6678FF83FFD8688C5D00FE5433C6" pageId="18" pageNumber="131" startId="18.[151,215,1887,1909]" targetBox="[153,1401,1199,1865]" targetPageId="18" targetType="figure">
<paragraph id="DF6E36F0FF83FFD8688C5D00FE5433C6" blockId="18.[151,1399,1887,1977]" pageId="18" pageNumber="131">
<emphasis id="EDA5EAE2FF83FFD8688C5D00FF5D330A" bold="true" box="[151,245,1887,1909]" pageId="18" pageNumber="131">Figure 53</emphasis>
. Principal component analysis of 84 nest samples of
<taxonomicName id="18D14D73FF83FFD86AFF5D00FC49330A" authorityName="Forel" authorityYear="1890" box="[740,993,1887,1908]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF83FFD86AFF5D00FC49330A" box="[740,993,1887,1908]" italics="true" pageId="18" pageNumber="131">Cardiocondyla wroughtonii</emphasis>
</taxonomicName>
and
<taxonomicName id="18D14D73FF83FFD86C0B5D3FFB2C330A" authorityName="Wheeler" authorityYear="1929" box="[1040,1156,1887,1909]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF83FFD86C0B5D3FFB2C330A" box="[1040,1156,1887,1909]" italics="true" pageId="18" pageNumber="131">C. obscurior</emphasis>
</taxonomicName>
based on 7 morphometric characters taken in 205 worker individuals. The arrow points to the sample determined as
<taxonomicName id="18D14D73FF83FFD86BA05D29FB8433F5" authorityName="Wheeler" authorityYear="1929" box="[955,1068,1910,1931]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF83FFD86BA05D29FB8433F5" box="[955,1068,1910,1931]" italics="true" pageId="18" pageNumber="131">C. obscurior</emphasis>
</taxonomicName>
by a wild-card LDA with p = 0.9456 which, however, showed a pigmentation typical for
<taxonomicName id="18D14D73FF83FFD86A6E5DD2FD5733DC" authorityName="Forel" authorityYear="1890" box="[629,767,1933,1954]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="131" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF83FFD86A6E5DD2FD5733DC" box="[629,767,1933,1954]" italics="true" pageId="18" pageNumber="131">C. wroughtonii</emphasis>
</taxonomicName>
. The mean error of the four exploratory data analyses relative to the controlling LDA (final.sp.hyp) is 1.8%.
</paragraph>
</caption>
<paragraph id="DF6E36F0FF82FFD968A65AAEFC883535" blockId="19.[189,802,241,1975]" pageId="19" pageNumber="132">
samples),
<collectingCountry id="A7C67660FF82FFD9693C5AAEFE233577" box="[295,395,241,265]" name="Bermuda" pageId="19" pageNumber="132">Bermuda</collectingCountry>
(1),
<collectingCountry id="A7C67660FF82FFD969A35AAEFDBA3577" box="[440,530,241,265]" name="Ecuador" pageId="19" pageNumber="132">Ecuador</collectingCountry>
(2),
<collectingCountry id="A7C67660FF82FFD96A5B5AADFD0D3574" box="[576,677,242,266]" name="Germany" pageId="19" pageNumber="132">Germany</collectingCountry>
(3), Hawaii (2),
<collectingCountry id="A7C67660FF82FFD968FC5B4DFEB73554" box="[231,287,274,298]" name="India" pageId="19" pageNumber="132">India</collectingCountry>
(1),
<collectingCountry id="A7C67660FF82FFD969555B4DFE223554" box="[334,394,274,298]" name="Israel" pageId="19" pageNumber="132">Israel</collectingCountry>
(1),
<collectingCountry id="A7C67660FF82FFD969A25B4CFE5E3554" box="[441,502,275,298]" name="Japan" pageId="19" pageNumber="132">Japan</collectingCountry>
(10),
<collectingCountry id="A7C67660FF82FFD96A295B4DFD033554" box="[562,683,274,298]" name="Micronesia" pageId="19" pageNumber="132">Micronesia</collectingCountry>
(1),
<collectingCountry id="A7C67660FF82FFD96AC25B4CFF583535" name="Puerto Rico" pageId="19" pageNumber="132">Puerto Rico</collectingCountry>
(2),
<collectingCountry id="A7C67660FF82FFD9693A5B6CFE3B3535" box="[289,403,307,331]" name="Seychelles" pageId="19" pageNumber="132">Seychelles</collectingCountry>
(5)
<collectingCountry id="A7C67660FF82FFD969A65B6CFD823535" box="[445,554,307,331]" name="Singapore" pageId="19" pageNumber="132">Singapore</collectingCountry>
(1),
<collectingCountry id="A7C67660FF82FFD96A425B6CFD3F3535" box="[601,663,307,331]" name="Spain" pageId="19" pageNumber="132">Spain</collectingCountry>
(5),
<collectingCountry id="A7C67660FF82FFD96AD35B6CFD543535" box="[712,764,307,331]" name="United States of America" pageId="19" pageNumber="132">USA</collectingCountry>
(4)
</paragraph>
<paragraph id="DF6E36F0FF82FFD968CF5B0BFE4C362C" blockId="19.[189,802,241,1975]" pageId="19" pageNumber="132">
<emphasis id="EDA5EAE2FF82FFD968CF5B0BFE0E3512" bold="true" box="[212,422,340,364]" pageId="19" pageNumber="132">Geographic range</emphasis>
. As a tramp species of putatively Southeast Asian origin this species is, like
<taxonomicName id="18D14D73FF82FFD96A8C5B2AFCB235F3" authorityName="Forel" authorityYear="1890" box="[663,794,373,397]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD96A8C5B2AFCB235F3" box="[663,794,373,397]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
</taxonomicName>
, widely distributed over the tropical regions of the Old and New World. However, with exception of a finding from
<collectingCountry id="A7C67660FF82FFD968A65B87FE85358E" box="[189,301,472,496]" name="Singapore" pageId="19" pageNumber="132">Singapore</collectingCountry>
, there are no verified vouchers from the Indo- Malayan and Australasian regions where
<taxonomicName id="18D14D73FF82FFD96A655BA6FC89366E" authorityName="Forel" authorityYear="1890" box="[638,801,504,528]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD96A655BA6FC89366E" box="[638,801,504,528]" italics="true" pageId="19" pageNumber="132">C. wroughtonii</emphasis>
</taxonomicName>
is dominant. It has also been found in buildings in the temperate zone (
<collectingCountry id="A7C67660FF82FFD969745865FE70362C" box="[367,472,570,594]" name="Germany" pageId="19" pageNumber="132">Germany</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF82FFD968CF5804FDD832F1" pageId="19" pageNumber="132" type="diagnosis">
<paragraph id="DF6E36F0FF82FFD968CF5804FDE33233" blockId="19.[189,802,241,1975]" pageId="19" pageNumber="132">
<emphasis id="EDA5EAE2FF82FFD968CF5804FEEA360D" bold="true" box="[212,322,603,627]" pageId="19" pageNumber="132">Diagnosis</emphasis>
: --Worker (
<tableCitation id="9253034BFF82FFD969DC5804FDA7360D" box="[455,527,603,627]" captionStart="Table 1" captionStartId="26.[151,204,245,266]" captionText="Table 1. Workers of the C. wroughtonii group of Orientalic-Australasian origin with mean FL/FR &lt;1.040. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="19" pageNumber="132">Tab. 1</tableCitation>
, Figs 3235, key). Very small and slightly larger than
<taxonomicName id="18D14D73FF82FFD96A125823FD1436EA" authority=", CS" authorityName="CS" box="[521,700,636,660]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD96A125823FD2236EA" box="[521,650,636,660]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
, CS
</taxonomicName>
430 µm. Head short, CL/CW 1.111. Anterocentral clypeal margin straight or slightly notched; central occipital margin usually straight or with a very weak concavity. Postocular distance large, PoOc/CL 0.435. Frons broad (FRS/CS 0.268), frontal carinae caudal of FRS level parallel (FL/FR 1.011). Eye medium-sized, EYE/CS 0.229. Scape short and slightly longer than in
<taxonomicName id="18D14D73FF82FFD969EC593DFD023704" authority=", SL" authorityName="SL" box="[503,682,866,890]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD969EC593DFDD03704" box="[503,632,866,890]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
, SL
</taxonomicName>
/CS 0.772. Metanotal groove in lateral view deep (MGr/CS 3.65 %) and usually with steep anterior and posterior slopes. Promesonotal plane in dorsal view without shoulders due to a strongly convex frontal margin and rather straight lateral margins (Fig. 34). Propodeal spines shorter than in
<taxonomicName id="18D14D73FF82FFD9690E5E78FE3E3041" box="[277,406,1063,1087]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD9690E5E78FE3E3041" box="[277,406,1063,1087]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
</taxonomicName>
(SP/CS 0.185) and their bases less approached (SPBA/CS 0.292). Petiole wider and slightly higher than in
<taxonomicName id="18D14D73FF82FFD969685E36FE5C30FF" box="[371,500,1129,1153]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD969685E36FE5C30FF" box="[371,500,1129,1153]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
</taxonomicName>
(PeW/CS 0.292, PeH/CS 0.338), axis of petiolar peduncle in lateral aspect deviating by 30° from the longitudinal axis of the petiole node. Postpetiole wider and higher than in
<taxonomicName id="18D14D73FF82FFD96A495E93FD7B309A" box="[594,723,1228,1252]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD96A495E93FD7B309A" box="[594,723,1228,1252]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
</taxonomicName>
(PpW/ CS 0.459, PpH/CS 0.314), the sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Sculpture on head, mesosoma and waist similar to
<taxonomicName id="18D14D73FF82FFD969315FCEFE0331D7" box="[298,427,1425,1449]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD969315FCEFE0331D7" box="[298,427,1425,1449]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
</taxonomicName>
. Pubescence on first gaster tergite moderately long and dilute (PLG/CS 6.2 %, sqPDG 5.13). The dark color morph, which resembles the dark color morp of
<taxonomicName id="18D14D73FF82FFD969055FABFE373272" box="[286,415,1524,1548]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD969055FABFE373272" box="[286,415,1524,1548]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
</taxonomicName>
, is found in 98 % of the samples one sample, possibly hybrids, showed a pattern similar to the light color morph of
<taxonomicName id="18D14D73FF82FFD969D95C69FDEB3230" box="[450,579,1590,1614]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD969D95C69FDEB3230" box="[450,579,1590,1614]" italics="true" pageId="19" pageNumber="132">wroughtonii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DF6E36F0FF82FFD968CF5C09FDD832F1" blockId="19.[189,802,241,1975]" pageId="19" pageNumber="132">
<emphasis id="EDA5EAE2FF82FFD968CF5C09FE7D3211" bold="true" box="[212,469,1622,1647]" pageId="19" pageNumber="132">Taxonomic comments.</emphasis>
The morphological separation from
<taxonomicName id="18D14D73FF82FFD968E35C27FE3332F1" box="[248,411,1655,1679]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD968E35C27FE3332F1" box="[248,411,1655,1679]" italics="true" pageId="19" pageNumber="132">C. wroughtonii</emphasis>
</taxonomicName>
is described above.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF82FFD968CF5CC7FB3936CB" pageId="19" pageNumber="132" type="biology_ecology">
<paragraph id="DF6E36F0FF82FFD968CF5CC7FB3936CB" blockId="19.[189,802,241,1975]" lastBlockId="19.[824,1437,241,695]" pageId="19" pageNumber="132">
<emphasis id="EDA5EAE2FF82FFD968CF5CC7FE9832CE" bold="true" box="[212,304,1688,1712]" pageId="19" pageNumber="132">Biology.</emphasis>
In contrast to
<taxonomicName id="18D14D73FF82FFD969C75CC6FD2B32CE" box="[476,643,1688,1712]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF82FFD969C75CC6FD2B32CE" box="[476,643,1688,1712]" italics="true" pageId="19" pageNumber="132">C. wroughtonii</emphasis>
</taxonomicName>
,
<taxonomicName id="18D14D73FF82FFD96A8F5CC6FC8932CE" authorityName="Wheeler" authorityYear="1929" box="[660,801,1688,1712]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF82FFD96A8F5CC6FC8932CE" box="[660,801,1688,1712]" italics="true" pageId="19" pageNumber="132">C. obscurior</emphasis>
</taxonomicName>
was reported to nest in cavities of bushes and trees
<quantity id="18299B15FF82FFD96AE25CE6FF7A328F" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="19" pageNumber="132" unit="m" value="3.5" valueMax="5.0" valueMin="2.0">25 m</quantity>
above the ground level; it was found in dead twigs of trees such as
<taxonomicName id="18D14D73FF82FFD969425CA4FD9D336D" box="[345,565,1787,1811]" class="Magnoliopsida" family="Fabaceae" genus="Erythrina" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="132" phylum="Tracheophyta" rank="species" species="variegata">
<emphasis id="EDA5EAE2FF82FFD969425CA4FD9D336D" box="[345,565,1787,1811]" italics="true" pageId="19" pageNumber="132">Erythrina variegata</emphasis>
</taxonomicName>
(Okinawa), in dwarf coconuts (
<collectingCountry id="A7C67660FF82FFD969305D43FEDA334A" box="[299,370,1820,1844]" name="Brazil" pageId="19" pageNumber="132">Brazil</collectingCountry>
), galls of
<taxonomicName id="18D14D73FF82FFD969C15D43FD8C334D" box="[474,548,1820,1843]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="EDA5EAE2FF82FFD969C15D43FD8C334D" box="[474,548,1820,1843]" italics="true" pageId="19" pageNumber="132">Acacia</emphasis>
</taxonomicName>
trees (
<collectingCountry id="A7C67660FF82FFD96A755D43FD1D334A" box="[622,693,1820,1844]" name="Brazil" pageId="19" pageNumber="132">Brazil</collectingCountry>
), between crippled-curled leaves of a Citrus tree (
<collectingCountry id="A7C67660FF82FFD96A7D5D62FD03332B" box="[614,683,1853,1877]" name="Brazil" pageId="19" pageNumber="132">Brasil</collectingCountry>
), in a dead twig on a tree (Florida), on a
<taxonomicName id="18D14D73FF82FFD96A175D01FDE0330B" box="[524,584,1886,1909]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="19" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="EDA5EAE2FF82FFD96A175D01FDE0330B" box="[524,584,1886,1909]" italics="true" pageId="19" pageNumber="132">Ficus</emphasis>
</taxonomicName>
tree (
<collectingCountry id="A7C67660FF82FFD96A975D02FD65330B" box="[652,717,1885,1909]" name="Israel" pageId="19" pageNumber="132">Israel</collectingCountry>
), in the gall of a
<taxonomicName id="18D14D73FF82FFD969045D20FED133E8" box="[287,377,1919,1942]" class="Magnoliopsida" family="Tamaricaceae" genus="Tamarix" kingdom="Plantae" order="Caryophyllales" pageId="19" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="EDA5EAE2FF82FFD969045D20FED133E8" box="[287,377,1919,1942]" italics="true" pageId="19" pageNumber="132">Tamarix</emphasis>
</taxonomicName>
bush (
<collectingCountry id="A7C67660FF82FFD969D85D21FDAC33E8" box="[451,516,1918,1942]" name="Israel" pageId="19" pageNumber="132">Israel</collectingCountry>
), below the bark of strong trees (Tenerife) and in the cavity of a coconut high in the tree (
<collectingCountry id="A7C67660FF82FFD96BBA5AAEFBAF3577" box="[929,1031,241,265]" name="Tanzania" pageId="19" pageNumber="132">Zanzibar</collectingCountry>
). Demography and behavior of nest populations as well as male polyphenism and behavior are the same as in
<taxonomicName id="18D14D73FF82FFD96BE65B6BFA913535" authority="(Heinze 2017)" baseAuthorityName="Heinze" baseAuthorityYear="2017" box="[1021,1337,307,331]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="wroughtoni">
<emphasis id="EDA5EAE2FF82FFD96BE65B6BFB313535" box="[1021,1177,307,331]" italics="true" pageId="19" pageNumber="132">C. wroughtoni</emphasis>
(
<bibRefCitation id="BB404B01FF82FFD96CBD5B6CFA9A3535" author="Heinze, J." box="[1190,1330,307,331]" pageId="19" pageNumber="132" refId="ref18842" refString="Heinze, J. (2017): Life-history evolution in ants: the case of Cardiocondyla. - Proceedings of the Royal Society B: Biological Sciences 284 (1): e 20161406 [https: // doi. org / 10.1098 / rspb. 2016.1406]." type="journal volume" year="2017">Heinze 2017</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="BB404B01FF82FFD96D585B6CFC343512" author="Errbii, M. &amp; J. Keilwagen &amp; K. J. Hoff &amp; R. Steffen &amp; J. Altmuller &amp; J. Oettler &amp; L. Schrader" pageId="19" pageNumber="132" pagination="6211 - 6228" refId="ref18634" refString="Errbii, M., J. Keilwagen, K. J. Hoff, R. Steffen, J. Altmuller, J. Oettler &amp; L. Schrader L (2021). Transposable elements and introgression introduce genetic variation in the invasive ant Cardiocondyla obscurior. - Molecular Ecology 30 (23): 6211 - 6228 [https: // doi. org / 10.1111 / mec. 16099] Epub 2021 Aug 17. PMID: 34324751." type="journal article" year="2021">Errbii et al. (2021)</bibRefCitation>
have shown that populations of
<taxonomicName id="18D14D73FF82FFD96CE15B0BFC0935F3" authorityName="Wheeler" authorityYear="1929" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="132" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF82FFD96CE15B0BFC0935F3" italics="true" pageId="19" pageNumber="132">Cardiocondyla obscurior</emphasis>
</taxonomicName>
belong to two distinct lineages, a lineage so far only found in Latin America and a more cosmopolitan Old World lineage. A strong genetic differentiation of these lineages began latest 40,000 generations ago but introgression from the Old World lineage is a dominant source of genetic diversity in the Latin America lineage and is likely to contribute to the adaptive potential of the Latin America lineage. The lineages are infected by different
<taxonomicName id="18D14D73FF82FFD96BB95823FBBD36EA" authorityName="Hertig" authorityYear="1936" box="[930,1045,636,660]" class="Alphaproteobacteria" family="Anaplasmataceae" genus="Wolbachia" kingdom="Bacteria" order="Rickettsiales" pageId="19" pageNumber="132" phylum="Proteobacteria" rank="genus">
<emphasis id="EDA5EAE2FF82FFD96BB95823FBBD36EA" box="[930,1045,636,660]" italics="true" pageId="19" pageNumber="132">Wolbachia</emphasis>
</taxonomicName>
strains, one causing cytoplasmic incompatibility (
<bibRefCitation id="BB404B01FF82FFD96BF758C2FB2E36CB" author="Un, C. &amp; E. Schultner &amp; A. Manzano-Marin &amp; L. V. Florez &amp; B. Seifert &amp; J. Heinze &amp; J. Oettler" box="[1004,1158,669,695]" pageId="19" pageNumber="132" pagination="1775 - 1791" refId="ref19923" refString="Un, C., E. Schultner, A. Manzano-Marin, L. V. Florez, B. Seifert, J. Heinze &amp; J. Oettler (2021): Cytoplasmic incompatibility between Old and New World populations of a tramp ant. - Evolution 75 (7): 1775 - 1791 [https: // doi. org / 10.1111 / evo. 14261]." type="journal article" year="2021">Ün et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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View file

@ -0,0 +1,188 @@
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<paragraph id="DF6E36F0FF84FFDF68CF5D64FDE53329" blockId="21.[212,589,1851,1879]" box="[212,589,1851,1879]" pageId="21" pageNumber="134">
<heading id="8426819CFF84FFDF68CF5D64FDE53329" box="[212,589,1851,1879]" fontSize="11" level="2" pageId="21" pageNumber="134" reason="5">
<emphasis id="EDA5EAE2FF84FFDF68CF5D64FDE53329" bold="true" box="[212,589,1851,1879]" pageId="21" pageNumber="134">
<taxonomicName id="18D14D73FF84FFDF68CF5D64FDA83329" authorityName="Seifert" authorityYear="2024" box="[212,512,1851,1879]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="134" phylum="Arthropoda" rank="species" species="heinzei" status="sp. nov.">
<emphasis id="EDA5EAE2FF84FFDF68CF5D64FDA83329" bold="true" box="[212,512,1851,1879]" italics="true" pageId="21" pageNumber="134">Cardiocondyla heinzei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F6965799FF84FFDF6A115D64FDE53329" box="[522,589,1851,1879]" pageId="21" pageNumber="134" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="97CB657BFF84FFDF68CF5D21FC783554" pageId="21" pageNumber="134" type="etymology">
<paragraph id="DF6E36F0FF84FFDF68CF5D21FC8A33E9" blockId="21.[189,802,1918,1975]" box="[212,802,1918,1943]" pageId="21" pageNumber="134">Etymology: the name is given in honor of Jürgen</paragraph>
<paragraph id="DF6E36F0FF84FFDF68A65DC0FC783554" blockId="21.[189,802,1918,1975]" lastBlockId="21.[824,1437,241,1976]" pageId="21" pageNumber="134">
Heinze wo has done and supervised outstanding research on the fascinating biology of
<taxonomicName id="18D14D73FF84FFDF6C905AAEFA823577" authorityName="Emery" authorityYear="1869" box="[1163,1322,241,265]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="EDA5EAE2FF84FFDF6C905AAEFA823577" box="[1163,1322,241,265]" italics="true" pageId="21" pageNumber="134">Cardiocondyla</emphasis>
</taxonomicName>
ants over three decades.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF84FFDF6B545B6CFBE93747" pageId="21" pageNumber="134" type="materials_examined">
<paragraph id="DF6E36F0FF84FFDF6B545B6CFC503535" blockId="21.[824,1437,241,1976]" box="[847,1016,307,331]" pageId="21" pageNumber="134">
<emphasis id="EDA5EAE2FF84FFDF6B545B6CFC503535" bold="true" box="[847,1016,307,331]" pageId="21" pageNumber="134">Type material:</emphasis>
</paragraph>
<paragraph id="DF6E36F0FF84FFDF6B545B0BFAB636CB" blockId="21.[824,1437,241,1976]" pageId="21" pageNumber="134">
<materialsCitation id="6FB93CADFF84FFDF6B545B0BFAB636CB" collectingDate="2019-04-04" collectingDateMax="2019-04-07" collectingDateMin="2019-04-04" collectionCode="SMNG" county="Res. Station" elevation="191" latitude="8.7703" location="Iringo" longLatPrecision="7" longitude="-3.7897" municipality="Park" pageId="21" pageNumber="134" specimenCode="SMNG20240630" specimenCount="9" specimenCount-worker="8" stateProvince="Comoe" typeStatus="holotype">
<typeStatus id="006A8852FF84FFDF6B545B0BFC1B3512" box="[847,947,340,364]" pageId="21" pageNumber="134" type="holotype">Holotype</typeStatus>
worker labelled IVO:
<geoCoordinate id="BAE55037FF84FFDF6CDF5B0BFA803512" box="[1220,1320,340,364]" degrees="8.7703" direction="north" orientation="latitude" pageId="21" pageNumber="134" precision="5" value="8.7703">8.7703°N</geoCoordinate>
,
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,
<collectingRegion id="1D15F812FF84FFDF6B235B2AFC2D35F3" box="[824,901,373,397]" country="Ivory Coast" name="Comoe" pageId="21" pageNumber="134">Comoé</collectingRegion>
<collectingCounty id="360F4E7CFF84FFDF6B895B29FBB535F3" box="[914,1053,373,397]" pageId="21" pageNumber="134">Res. Station</collectingCounty>
,
<quantity id="18299B15FF84FFDF6C345B2AFBC335F3" box="[1071,1131,373,397]" metricMagnitude="2" metricUnit="m" metricValue="1.91" pageId="21" pageNumber="134" unit="m" value="191.0">
<elevation id="54FCD1C3FF84FFDF6C345B2AFBC335F3" box="[1071,1131,373,397]" metricMagnitude="2" metricUnit="m" metricValue="1.91" pageId="21" pageNumber="134" unit="m" value="191.0">191m</elevation>
</quantity>
, foragers on path halfway between station and river, Heinze
<date id="AB6F1030FF84FFDF6CD95BC9FA9E35D0" box="[1218,1334,406,430]" pageId="21" pageNumber="134" value="2019-04-04">
<collectingDate id="BB2BE9D8FF84FFDF6CD95BC9FA9E35D0" box="[1218,1334,406,430]" pageId="21" pageNumber="134" value="2019-04-04">2019.04.04</collectingDate>
</date>
CI 18,
<specimenCode id="8F779E8BFF84FFDF6B275BE8FC5435B1" box="[828,1020,439,463]" collectionCode="SMNG" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="21" pageNumber="134">SMNG20240630</specimenCode>
[unique specimen identifier];
<specimenCount id="C9D7FD79FF84FFDF6D7E5BE8FC2B358E" count="3" pageId="21" pageNumber="134" type="worker">three worker</specimenCount>
<typeStatus id="006A8852FF84FFDF6B945B87FC51358E" box="[911,1017,472,496]" pageId="21" pageNumber="134" type="paratype">paratypes</typeStatus>
with same labelling on another pin;
<specimenCount id="C9D7FD79FF84FFDF6B235BA7FC67366E" box="[824,975,504,528]" count="3" pageId="21" pageNumber="134" type="worker">three worker</specimenCount>
<typeStatus id="006A8852FF84FFDF6BFE5BA6FBE7366F" box="[997,1103,505,529]" pageId="21" pageNumber="134" type="paratype">paratypes</typeStatus>
labelled IVO:
<geoCoordinate id="BAE55037FF84FFDF6D3A5BA6FA30366F" box="[1313,1432,505,529]" degrees="8.84028" direction="north" orientation="latitude" pageId="21" pageNumber="134" precision="1" value="8.84028">8.84028°N</geoCoordinate>
,
<geoCoordinate id="BAE55037FF84FFDF6B235845FC01364C" box="[824,937,538,562]" degrees="3.77111" direction="west" orientation="longitude" pageId="21" pageNumber="134" precision="1" value="-3.77111">3.77111°W</geoCoordinate>
,
<collectingRegion id="1D15F812FF84FFDF6BA15845FBAF364C" box="[954,1031,538,562]" country="Ivory Coast" name="Comoe" pageId="21" pageNumber="134">Comoé</collectingRegion>
<collectingMunicipality id="3F0AAC8AFF84FFDF6C0F5846FBE0364F" box="[1044,1096,537,561]" pageId="21" pageNumber="134">Park</collectingMunicipality>
,
<location id="DA0E602BFF84FFDF6C415846FB35364F" LSID="urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6C415846FB35364F" box="[1114,1181,537,561]" county="Res. Station" latitude="8.7703" longLatPrecision="7" longitude="-3.7897" municipality="Park" name="Iringo" pageId="21" pageNumber="134" stateProvince="Comoe">Iringo</location>
<collectingRegion id="1D15F812FF84FFDF6CB25846FB4D364F" box="[1193,1253,537,561]" country="Nigeria" name="Rivers" pageId="21" pageNumber="134">River</collectingRegion>
<quantity id="18299B15FF84FFDF6CE95845FA96364F" box="[1266,1342,538,562]" metricMagnitude="2" metricUnit="m" metricValue="2.02" pageId="21" pageNumber="134" unit="m" value="202.0">
<elevation id="54FCD1C3FF84FFDF6CE95845FA96364F" box="[1266,1342,538,562]" metricMagnitude="2" metricUnit="m" metricValue="2.02" pageId="21" pageNumber="134" unit="m" value="202.0">202 m</elevation>
</quantity>
,
<location id="DA0E602BFF84FFDF6D4B5846FA35364F" LSID="urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6D4B5846FA35364F" box="[1360,1437,537,561]" county="Res. Station" latitude="8.7703" longLatPrecision="7" longitude="-3.7897" municipality="Park" name="Heinze" pageId="21" pageNumber="134" stateProvince="Comoe">Heinze</location>
<date id="AB6F1030FF84FFDF6B235865FC04362C" box="[824,940,570,594]" pageId="21" pageNumber="134" value="2019-04-06">
<collectingDate id="BB2BE9D8FF84FFDF6B235865FC04362C" box="[824,940,570,594]" pageId="21" pageNumber="134" value="2019-04-06">2019.04.06</collectingDate>
</date>
CI 45;
<specimenCount id="C9D7FD79FF84FFDF6C0A5864FB23362C" box="[1041,1163,570,594]" count="2" pageId="21" pageNumber="134" type="worker">two worker</specimenCount>
<typeStatus id="006A8852FF84FFDF6C895864FB54362D" box="[1170,1276,571,595]" pageId="21" pageNumber="134" type="paratype">paratypes</typeStatus>
labelled IVO:
<geoCoordinate id="BAE55037FF84FFDF6B235804FC37360D" box="[824,927,603,627]" degrees="8.7703" direction="north" orientation="latitude" pageId="21" pageNumber="134" precision="5" value="8.7703">8.7703°N</geoCoordinate>
,
<geoCoordinate id="BAE55037FF84FFDF6BB35804FBBB360D" box="[936,1043,603,627]" degrees="3.7897" direction="west" orientation="longitude" pageId="21" pageNumber="134" precision="5" value="-3.7897">3.7897°W</geoCoordinate>
,
<collectingRegion id="1D15F812FF84FFDF6C075804FBC1360D" box="[1052,1129,603,627]" country="Ivory Coast" name="Comoe" pageId="21" pageNumber="134">Comoé</collectingRegion>
<location id="DA0E602BFF84FFDF6C6B5803FB5B360D" LSID="urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6C6B5803FB5B360D" box="[1136,1267,603,627]" county="Res. Station" latitude="8.7703" longLatPrecision="7" longitude="-3.7897" municipality="Park" name="Res. Station" pageId="21" pageNumber="134" stateProvince="Comoe">Res. Station</location>
,
<quantity id="18299B15FF84FFDF6CE55804FA92360D" box="[1278,1338,603,627]" metricMagnitude="2" metricUnit="m" metricValue="1.91" pageId="21" pageNumber="134" unit="m" value="191.0">
<elevation id="54FCD1C3FF84FFDF6CE55804FA92360D" box="[1278,1338,603,627]" metricMagnitude="2" metricUnit="m" metricValue="1.91" pageId="21" pageNumber="134" unit="m" value="191.0">191m</elevation>
</quantity>
, foragers on path halfway, between station and river,
<location id="DA0E602BFF84FFDF6D545823FA3436EA" LSID="urn:lsid:plazi:treatment:577887E6FF84FFDC68CF5D64FE2B358E:DA0E602BFF84FFDF6D545823FA3436EA" box="[1359,1436,636,660]" county="Res. Station" latitude="8.7703" longLatPrecision="7" longitude="-3.7897" municipality="Park" name="Heinze" pageId="21" pageNumber="134" stateProvince="Comoe">Heinze</location>
<date id="AB6F1030FF84FFDF6B2358C2FC0336CB" box="[824,939,669,693]" pageId="21" pageNumber="134" value="2019-04-07">
<collectingDate id="BB2BE9D8FF84FFDF6B2358C2FC0336CB" box="[824,939,669,693]" pageId="21" pageNumber="134" value="2019-04-07">2019.04.07</collectingDate>
</date>
CI 55; depository SMN Görlitz.
</materialsCitation>
</paragraph>
<paragraph id="DF6E36F0FF84FFDF6B5458E1FB323689" blockId="21.[824,1437,241,1976]" pageId="21" pageNumber="134">
<emphasis id="EDA5EAE2FF84FFDF6B5458E1FBFD36A8" bold="true" box="[847,1109,702,726]" pageId="21" pageNumber="134">All material examined</emphasis>
.
<materialsCitation id="6FB93CADFF84FFDF6C7F58E1FB3E3689" country="Ivory Coast" location="Ivory Coast" pageId="21" pageNumber="134" specimenCount="3" typeStatus="holotype">
Only the
<specimenCount id="C9D7FD79FF84FFDF6CD658E1FA9536A8" box="[1229,1341,702,726]" count="3" pageId="21" pageNumber="134" type="generic" typeStatus="type">three type</specimenCount>
samples from
<collectingCountry id="A7C67660FF84FFDF6B685880FC593689" box="[883,1009,735,759]" name="Ivory Coast" pageId="21" pageNumber="134">Ivory Coast</collectingCountry>
were available
</materialsCitation>
.
</paragraph>
<paragraph id="DF6E36F0FF84FFDF6B5458A0FBE93747" blockId="21.[824,1437,241,1976]" pageId="21" pageNumber="134">
<emphasis id="EDA5EAE2FF84FFDF6B5458A0FB8A3766" bold="true" box="[847,1058,767,792]" pageId="21" pageNumber="134">Geographic range</emphasis>
.
<materialsCitation id="6FB93CADFF84FFDF6C2E595FFB953747" country="Ivory Coast" location="Ivory Coast" pageId="21" pageNumber="134" specimenCount="2" typeStatus="holotype">
Only known from the
<specimenCount id="C9D7FD79FF84FFDF6D23595FFA343766" box="[1336,1436,768,792]" count="2" pageId="21" pageNumber="134" type="generic" typeStatus="type">two type</specimenCount>
localities in
<collectingCountry id="A7C67660FF84FFDF6BA7597EFB953747" box="[956,1085,801,825]" name="Ivory Coast" pageId="21" pageNumber="134">Ivory Coast</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF84FFDC6B54591EFDFB35D0" lastPageId="22" lastPageNumber="135" pageId="21" pageNumber="134" type="diagnosis">
<paragraph id="DF6E36F0FF84FFDC6B54591EFD953532" blockId="21.[824,1437,241,1976]" lastBlockId="22.[151,764,241,496]" lastPageId="22" lastPageNumber="135" pageId="21" pageNumber="134">
<emphasis id="EDA5EAE2FF84FFDF6B54591EFC163727" bold="true" box="[847,958,833,857]" pageId="21" pageNumber="134">Diagnosis</emphasis>
: --Worker (
<tableCitation id="9253034BFF84FFDF6C4C591EFB033727" box="[1111,1195,833,857]" captionStart="Table 2" captionStartId="26.[150,203,1476,1497]" captionText="Table 2. Workers of the C. wroughtonii group with mean FL/FR&gt; 1.040 and originating from Africa and theArabian Peninsula. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="21" pageNumber="134">Tab. 2</tableCitation>
, Figs 4448, key). Extremely small, CS 377 µm. Head elongated, CL/CW 1.189. Median third of anterior clypeal margin slightly concave, median third of occipital margin straight. Postocular distance large, PoOc/CL 0.457. Frons very narrow (FRS/CS 0.219), widening frontad (FL/FR 1.063), caudal of FRS level parallel. Eye without any microsetae and rather smalll, EYE/CS 0.232. Scape moderately long, SL/CS 0.786. Metanotal groove in profile in absolute terms rather deep (MGr/CS 3.25 %) but with shallow slopes to mesonotum and propodeum. Prododeal spines long (SP/CS 0.201), thin and acute; in profile deviating from longitudinal mesosomal axis by only 25°, in dorsal view slightly diverging and and slightly incurved; their bases narrow (SPBA/CS 0.240). Petiole very narrow and moderately high (PeW/CS 0.234, PeH/CS 0.333); in lateral view with a concave anterior face and a long weakly convex dorsal profile; petiole in dorsal view very slender, its node 1.5 fold as long as wide and narrowing frontad. Postpetiole rather wide and low (PpW/CS 0.454, PpH/CS 0.276); the sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Postpetiole in dorsal view with a concave anterior margin and convex sides. Anterior clypeus with fine transverse rugulae. Vertex with densely-arranged, flat-bottomed and moderately large foveolae (dFov 16.7). The largest foveolae show a flat tubercle around the hair base of 68 µm diameter which is sometimes connected with the outer ring through 12 fine microcarinulae (
<figureCitation id="47EA2A75FF84FFDF6C405D62FB18332B" box="[1115,1200,1853,1877]" captionStart="Figure 4748" captionStartId="14.[151,215,1787,1809]" captionTargetBox="[151,1398,261,1741]" captionTargetPageId="14" captionText="Figure 4748. Cardiocondyla heinzei n.sp., holotype, Comoe / Ivory Coast, 4 April 2019. (47) head surface between inner eye margin and paramedian vertex. (48) anterior dorsum of 1st gaster tergite. Figure 4951: Cardiocondyla neferka, holotype, Mampong / Ghana, 10 February 1970; from www.antweb.org CASENT0901747, images by Ryan Perry. (49) head in dorsal view; (50) lateral view; (51) dorsal view." pageId="21" pageNumber="134">Fig. 47</figureCitation>
). Whole lateral area of mesosoma microreticulate, promesontum foveolate-reticulate. Petiole microreticulate, postpetiole more shiny. First gaster tergite with a rather long and dense pubescence (PLG/CS 7.12%, sqPDG 4.14) and a weak microreticulum (
<figureCitation id="47EA2A75FF87FFDC69545B4DFE373554" box="[335,415,274,298]" captionStart="Figure 4748" captionStartId="14.[151,215,1787,1809]" captionTargetBox="[151,1398,261,1741]" captionTargetPageId="14" captionText="Figure 4748. Cardiocondyla heinzei n.sp., holotype, Comoe / Ivory Coast, 4 April 2019. (47) head surface between inner eye margin and paramedian vertex. (48) anterior dorsum of 1st gaster tergite. Figure 4951: Cardiocondyla neferka, holotype, Mampong / Ghana, 10 February 1970; from www.antweb.org CASENT0901747, images by Ryan Perry. (49) head in dorsal view; (50) lateral view; (51) dorsal view." pageId="22" pageNumber="135">Fig. 48</figureCitation>
). All body parts yellowish with exception of the blackish brown gaster.
</paragraph>
<paragraph id="DF6E36F0FF87FFDC68B55B0BFDFB35D0" blockId="22.[151,764,241,496]" pageId="22" pageNumber="135">
<emphasis id="EDA5EAE2FF87FFDC68B55B0BFE063512" bold="true" box="[174,430,340,364]" pageId="22" pageNumber="135">Taxonomic comments.</emphasis>
As a combination of large CL/ CW and small PeW/CS and FRS/CS not confuse with other members of the
<taxonomicName id="18D14D73FF87FFDC69985BC9FDAE35D0" authorityName="Forel" authorityYear="1890" box="[387,518,406,430]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="135" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF87FFDC69985BC9FDAE35D0" box="[387,518,406,430]" italics="true" pageId="22" pageNumber="135">wroughtonii</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF87FFDC68B55BE8FE2B358E" pageId="22" pageNumber="135" type="biology_ecology">
<paragraph id="DF6E36F0FF87FFDC68B55BE8FE2B358E" blockId="22.[151,764,241,496]" pageId="22" pageNumber="135">
<emphasis id="EDA5EAE2FF87FFDC68B55BE8FEA335B1" bold="true" box="[174,267,439,463]" pageId="22" pageNumber="135">Biology.</emphasis>
The workers foraged on ground in both open and woody situations.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,120 @@
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<taxonomicName id="18D14D73FF85FFDE68B55E78FDB43041" authority="Seifert 2003" authorityName="Seifert" authorityYear="2003" box="[174,540,1063,1087]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="133" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="EDA5EAE2FF85FFDE68B55E78FE383041" bold="true" box="[174,400,1063,1087]" italics="true" pageId="20" pageNumber="133">Cardiocondyla nana</emphasis>
<bibRefCitation id="BB404B01FF85FFDE698F5E78FDB43041" author="Seifert, B." box="[404,540,1063,1087]" pageId="20" pageNumber="133" pagination="203 - 338" refId="ref19624" refString="Seifert, B. (2003): The ant genus Cardiocondyla (Insecta: Hymenoptera: Formicidae) - a taxonomic revision of the C. elegans, C. bulgarica, C. batesii, C. nuda, C. shuckardi, C. stambuloffii, C. wroughtonii, C. emeryi, and C. minutior species groups. - Annalen des Naturhistorischen Museums Wien, Serie B, 104: 203 - 338." type="journal article" year="2003">Seifert 2003</bibRefCitation>
</taxonomicName>
[
<typeStatus id="006A8852FF85FFDE6A3C5E77FDF3303E" box="[551,603,1064,1088]" pageId="20" pageNumber="133">type</typeStatus>
investigation]
</emphasis>
</heading>
</paragraph>
<paragraph id="DF6E36F0FF85FFDE68B55E17FEE230BD" blockId="20.[151,764,1063,1975]" pageId="20" pageNumber="133">
<materialsCitation id="6FB93CADFF85FFDE68B55E17FEEE30BD" collectionCode="T, MC, BMNH" country="Brunei" location="Day'" municipality="Ulu" pageId="20" pageNumber="133" specimenCount="1" stateProvince="Temburong" typeStatus="holotype">
This taxon has been described from Brunei. Investigated was the
<typeStatus id="006A8852FF85FFDE69175E36FEC330FF" box="[268,363,1129,1153]" pageId="20" pageNumber="133" type="holotype">holotype</typeStatus>
worker labelled
<collectingCountry id="A7C67660FF85FFDE6A545E36FD1330FF" box="[591,699,1129,1153]" name="Brunei" pageId="20" pageNumber="133">BRUNEI</collectingCountry>
:
<collectingMunicipality id="3F0AAC8AFF85FFDE6AC85E36FD5330FF" box="[723,763,1129,1153]" pageId="20" pageNumber="133">Ulu</collectingMunicipality>
<collectingRegion id="1D15F812FF85FFDE688C5ED5FEBA30DC" box="[151,274,1162,1186]" country="Brunei" name="Temburong" pageId="20" pageNumber="133">Temburong</collectingRegion>
L.P.-283.m.
<collectionCode id="B9C0AE35FF85FFDE69835ED5FE0230DC" box="[408,426,1162,1186]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="133">T</collectionCode>
. 22.ii.82.
<collectionCode id="B9C0AE35FF85FFDE6A155ED5FD9430DC" box="[526,572,1162,1186]" country="0" name="Museo de Cipolleti" pageId="20" pageNumber="133">MC</collectionCode>
.
<location id="DA0E602BFF85FFDE6A5B5ED5FDDF30DC" LSID="urn:lsid:plazi:treatment:577887E6FF85FFDE68B55E78FC5730DC:DA0E602BFF85FFDE6A5B5ED5FDDF30DC" box="[576,631,1162,1186]" country="Brunei" municipality="Ulu" name="Day'" pageId="20" pageNumber="133" stateProvince="Temburong">Day</location>
, depository
<collectionCode id="B9C0AE35FF85FFDE688C5EF3FF4030BD" box="[151,232,1196,1219]" pageId="20" pageNumber="133">BMNH</collectionCode>
London
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF85FFDE68B55E93FE523138" pageId="20" pageNumber="133" type="materials_examined">
<paragraph id="DF6E36F0FF85FFDE68B55E93FF41317B" blockId="20.[151,764,1063,1975]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE68B55E93FE12309A" bold="true" box="[174,442,1228,1252]" pageId="20" pageNumber="133">All material examined</emphasis>
. Only the
<typeStatus id="006A8852FF85FFDE6A205E93FDC3309A" box="[571,619,1228,1252]" pageId="20" pageNumber="133">type</typeStatus>
specimen is known.
</paragraph>
<paragraph id="DF6E36F0FF85FFDE68B55F52FE523138" blockId="20.[151,764,1063,1975]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE68B55F52FE2B3158" bold="true" box="[174,387,1293,1318]" pageId="20" pageNumber="133">Geographic range</emphasis>
. The
<typeStatus id="006A8852FF85FFDE69D65F51FE553158" box="[461,509,1294,1318]" pageId="20" pageNumber="133">type</typeStatus>
locality is situated at
<geoCoordinate id="BAE55037FF85FFDE688C5F70FF773139" box="[151,223,1327,1351]" degrees="1.47" direction="south" orientation="latitude" pageId="20" pageNumber="133" precision="555" value="-1.47">1.47°S</geoCoordinate>
,
<geoCoordinate id="BAE55037FF85FFDE68F15F71FEF03138" box="[234,344,1326,1350]" degrees="123.559" direction="east" orientation="longitude" pageId="20" pageNumber="133" precision="55" value="123.559">123.559°E</geoCoordinate>
and
<quantity id="18299B15FF85FFDE69965F70FE6B3138" box="[397,451,1327,1351]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="133" unit="m" value="12.0">12 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF85FFDE68B55F10FAA230FF" pageId="20" pageNumber="133" type="diagnosis">
<paragraph id="DF6E36F0FF85FFDE68B55F10FB313061" blockId="20.[151,764,1063,1975]" lastBlockId="20.[786,1399,241,1186]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE68B55F10FEB23119" bold="true" box="[174,282,1359,1383]" pageId="20" pageNumber="133">Diagnosis</emphasis>
: --Worker (
<tableCitation id="9253034BFF85FFDE69945F10FE673116" box="[399,463,1359,1384]" captionStart="Table 1" captionStartId="26.[151,204,245,266]" captionText="Table 1. Workers of the C. wroughtonii group of Orientalic-Australasian origin with mean FL/FR &lt;1.040. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="20" pageNumber="133">Tab. 1</tableCitation>
, Figs1618, key).Extremely small, CS 366 µm. Head extremely short, CL/CW 1.069. Median third of anterior clypeal margin deeply, median third of occipital margin slightly concave. Postocular distance large, PoOc/CL 0.464. Frons moderately broad (FRS/CS 0.266), frontal carinae diverging caudad. Eye without any microsetae and small, EYE/CS 0.220. Scape rather long, SL/CS 0.818. Metanotal groove in lateral view relatively deep (MGr/CS 2.50 %). Propodeal spines long and thin, rather steep, in profile deviating from longitudinal mesosomal axis by 43° (SP/CS 0.192), in dorsal view strongly diverging and their bases moderately wide (SPBA/ CS 0.277). Petiole rather narrow and moderately high (PeW/CS 0.264, PeH/CS 0.336); in profile with almost linear (only slightly concave) anterior face and semicircular dorsum; in dorsal view with rather slender peduncle and almost globular node, which is slightly longer than wide. Postpetiole moderately wide and low (PpW/CS 0.426, PpH/CS 0.289); the sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Postpetiole in dorsal view with a concave anterior margin and convex sides. Anterior clypeus smooth, not shining. Frontal laminae and anteromedian vertex with weak microsculpture, consisting of an irregular mixture of corrugated, foveolate, and carinulate elements. Paramedian and lateral areas of vertex with a unique sculpture: densely-arranged, deeply impressed, flat-bottomed and very large foveae of 1923 µm diameter. The largest foveolae show a well-demarcated central ring of 89 µm diameter which is connected with the outer ring through 24 very fine microcarinulae in 90° cross-wire arrangement, such suggesting a four-leaf clover; in the smaller foveolae the number of microcarinulae may be reduced. Whole lateral area of mesosoma, anterior area of pronotum, and dorsal area of propodeum strongly microreticulate; dorsal area of promesonotum similarly foveate as paramedian vertex. Longitudinal sculpture on whole mesosoma, including metapleural gland bulla, completely absent. Petiole strongly microreticulate, postpetiole shining, very finely microreticulate. Pubescence on first gaster tergite moderately long and very dilute (PLG/CS 6.29 %, sqPDG 5.60). All body parts light-yellowish.
</paragraph>
<paragraph id="DF6E36F0FF85FFDE6B325E78FAA230FF" blockId="20.[786,1399,241,1186]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE6B325E78FB82303E" bold="true" box="[809,1066,1063,1088]" pageId="20" pageNumber="133">Taxonomic comments.</emphasis>
This species is not to confuse due to extremely small size, the extremely low CL/CW and the large characteristic foveolae on vertex.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF85FFDE6B325ED5FC5730DC" box="[809,1023,1162,1186]" pageId="20" pageNumber="133" type="biology_ecology">
<paragraph id="DF6E36F0FF85FFDE6B325ED5FC5730DC" blockId="20.[786,1399,241,1186]" box="[809,1023,1162,1186]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE6B325ED5FC2D30DC" bold="true" box="[809,901,1162,1186]" pageId="20" pageNumber="133">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,194 @@
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<treatment id="577887E6FF85FFDF6B325F71FE12328C" LSID="urn:lsid:plazi:treatment:577887E6FF85FFDF6B325F71FE12328C" httpUri="http://treatment.plazi.org/id/577887E6FF85FFDF6B325F71FE12328C" lastPageId="21" lastPageNumber="134" pageId="20" pageNumber="133">
<subSubSection id="97CB657BFF85FFDE6B325F71FBAD3119" pageId="20" pageNumber="133" type="nomenclature">
<paragraph id="DF6E36F0FF85FFDE6B325F71FB713138" blockId="20.[809,1241,1326,1350]" box="[809,1241,1326,1350]" pageId="20" pageNumber="133">
<heading id="8426819CFF85FFDE6B325F71FB713138" box="[809,1241,1326,1350]" fontSize="10" level="2" pageId="20" pageNumber="133" reason="4">
<taxonomicName id="18D14D73FF85FFDE6B325F71FB713138" authority="Seifert 2023" authorityName="Seifert" authorityYear="2023" box="[809,1241,1326,1350]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="133" phylum="Arthropoda" rank="species" species="allonivalis">
<emphasis id="EDA5EAE2FF85FFDE6B325F71FB713138" bold="true" box="[809,1241,1326,1350]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE6B325F71FBE03138" bold="true" box="[809,1096,1326,1350]" italics="true" pageId="20" pageNumber="133">Cardiocondyla allonivalis</emphasis>
Seifert 2023
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DF6E36F0FF85FFDE6B325F10FBAD3119" blockId="20.[809,1029,1359,1383]" box="[809,1029,1359,1383]" pageId="20" pageNumber="133">
<heading id="8426819CFF85FFDE6B325F10FBAD3119" bold="true" box="[809,1029,1359,1383]" fontSize="10" level="2" pageId="20" pageNumber="133" reason="3">
<emphasis id="EDA5EAE2FF85FFDE6B325F10FBAD3119" bold="true" box="[809,1029,1359,1383]" pageId="20" pageNumber="133">
[
<typeStatus id="006A8852FF85FFDE6B2B5F10FCCC3119" box="[816,868,1359,1383]" pageId="20" pageNumber="133">type</typeStatus>
investigation]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF85FFDE6B325F2FFA893272" pageId="20" pageNumber="133" type="description">
<paragraph id="DF6E36F0FF85FFDE6B325F2FFA893272" blockId="20.[786,1399,1392,1976]" pageId="20" pageNumber="133">
<materialsCitation id="6FB93CADFF85FFDE6B325F2FFA893272" collectingDate="2007-10-23" collectorName="Wanang &amp; Sogeram &amp; Janda &amp; Gorlitz." country="Papua New Guinea" elevation="95" latitude="145.267" location="This" municipality="Investigated" pageId="20" pageNumber="133" specimenCount="1" specimenCount-worker="1" typeStatus="holotype">
<location id="DA0E602BFF85FFDE6B325F2FFCF231F6" LSID="urn:lsid:plazi:treatment:577887E6FF85FFDF6B325F71FE12328C:DA0E602BFF85FFDE6B325F2FFCF231F6" box="[809,858,1392,1416]" country="Papua New Guinea" latitude="145.267" municipality="Investigated" name="This" pageId="20" pageNumber="133">This</location>
taxon has been described from
<collectingCountry id="A7C67660FF85FFDE6CBF5F2EFADB31F6" box="[1188,1395,1392,1416]" name="Papua New Guinea" pageId="20" pageNumber="133">Papua New Guinea</collectingCountry>
.
<collectingMunicipality id="3F0AAC8AFF85FFDE6B095FCEFC3E31D7" box="[786,918,1425,1449]" pageId="20" pageNumber="133">Investigated</collectingMunicipality>
was the
<typeStatus id="006A8852FF85FFDE6BE25FCEFBF531D7" box="[1017,1117,1425,1449]" pageId="20" pageNumber="133" type="holotype">Holotype</typeStatus>
and
<specimenCount id="C9D7FD79FF85FFDE6C8E5FCDFADE31D7" box="[1173,1398,1425,1450]" count="1" pageId="20" pageNumber="133" type="worker" typeStatus="paratype">one paratype worker</specimenCount>
on separate pins, both labelled PNG:
<geoCoordinate id="BAE55037FF85FFDE6CB05FEDFB5D31B4" box="[1195,1269,1458,1482]" degrees="5.25" direction="south" orientation="latitude" pageId="20" pageNumber="133" precision="555" value="-5.25">5.25°S</geoCoordinate>
,
<geoCoordinate id="BAE55037FF85FFDE6CE45FEDFADB31B4" box="[1279,1395,1458,1482]" degrees="145.267" direction="north" orientation="latitude" pageId="20" pageNumber="133" precision="55" value="145.267">145.267°N</geoCoordinate>
,
<collectorName id="72245326FF85FFDE6B095F8CFCC53195" box="[786,877,1491,1515]" pageId="20" pageNumber="133">Wanang</collectorName>
,
<collectorName id="72245326FF85FFDE6B605F8CFC733195" box="[891,987,1491,1515]" pageId="20" pageNumber="133">Sogeram</collectorName>
riv.
<quantity id="18299B15FF85FFDE6C095F8CFBE63194" box="[1042,1102,1491,1515]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="20" pageNumber="133" unit="m" value="95.0">
<elevation id="54FCD1C3FF85FFDE6C095F8CFBE63194" box="[1042,1102,1491,1515]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="20" pageNumber="133" unit="m" value="95.0">95 m</elevation>
</quantity>
,
<collectingMethod id="06904EE7FF85FFDE6C465F8CFB173194" box="[1117,1215,1491,1515]" pageId="20" pageNumber="133">Bait trap</collectingMethod>
ground D4, coll.
<collectorName id="72245326FF85FFDE6B5C5FABFC2C3272" box="[839,900,1524,1548]" pageId="20" pageNumber="133">Janda</collectorName>
<date id="AB6F1030FF85FFDE6B975FABFBAD3272" box="[908,1029,1524,1548]" pageId="20" pageNumber="133" value="2007-10-23">
<collectingDate id="BB2BE9D8FF85FFDE6B975FABFBAD3272" box="[908,1029,1524,1548]" pageId="20" pageNumber="133" value="2007-10-23">2007.10.23</collectingDate>
</date>
; depository SMN
<collectorName id="72245326FF85FFDE6CD65FABFA893272" box="[1229,1313,1524,1548]" pageId="20" pageNumber="133">Görlitz.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF85FFDE6B325C4BFB253308" pageId="20" pageNumber="133" type="materials_examined">
<paragraph id="DF6E36F0FF85FFDE6B325C4BFB3432AF" blockId="20.[786,1399,1392,1976]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE6B325C4BFB843252" bold="true" box="[809,1068,1556,1580]" pageId="20" pageNumber="133">All material examined</emphasis>
.
<materialsCitation id="6FB93CADFF85FFDE6C215C4AFB3F32AF" collectionCode="CASENT" country="Papua New Guinea" location="Papua New Guinea" pageId="20" pageNumber="133" specimenCode="CASENT0914964" specimenCount="5" specimenCount-worker="5">
Examined were four samples with
<specimenCount id="C9D7FD79FF85FFDE6B495C69FC763230" box="[850,990,1590,1614]" count="5" pageId="20" pageNumber="133" type="worker">five workers</specimenCount>
from
<collectingCountry id="A7C67660FF85FFDE6C295C69FABB3230" box="[1074,1299,1590,1614]" name="Papua New Guinea" pageId="20" pageNumber="133">Papua New Guinea</collectingCountry>
three samples through direct stereomicroscopic evaluation and one sample through photo evaluation of the specimen
<specimenCode id="8F779E8BFF85FFDE6B095CC7FC7C32CE" box="[786,980,1688,1712]" collectionCode="CASENT" pageId="20" pageNumber="133">CASENT0914964</specimenCode>
in www.antweb.org. For details see supplementary information SI1, SI2
</materialsCitation>
.
</paragraph>
<paragraph id="DF6E36F0FF85FFDE6B325C85FB253308" blockId="20.[786,1399,1392,1976]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE6B325C85FC52328C" bold="true" box="[809,1018,1754,1778]" pageId="20" pageNumber="133">Geographic range</emphasis>
.
<materialsCitation id="6FB93CADFF85FFDE6C125C85FAAD334A" collectorName="Cape Wom" country="Papua New Guinea" elevation="350" latitude="-3.533" location="The" longLatPrecision="78" longitude="143.583" pageId="20" pageNumber="133" specimenCount="1">
<location id="DA0E602BFF85FFDE6C125C85FB9B328C" LSID="urn:lsid:plazi:treatment:577887E6FF85FFDF6B325F71FE12328C:DA0E602BFF85FFDE6C125C85FB9B328C" box="[1033,1075,1754,1778]" country="Papua New Guinea" latitude="-3.533" longLatPrecision="78" longitude="143.583" name="The" pageId="20" pageNumber="133">The</location>
species is so far only known from sea level up to
<quantity id="18299B15FF85FFDE6BE65CA4FBEA336C" box="[1021,1090,1787,1811]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="20" pageNumber="133" unit="m" value="350.0">
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</quantity>
in four sites in
<collectingCountry id="A7C67660FF85FFDE6CE25CA4FCCD334A" name="Papua New Guinea" pageId="20" pageNumber="133">Papua New Guinea</collectingCountry>
:
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(
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,
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)
</materialsCitation>
;
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<location id="DA0E602BFF85FFDE6D055D43FADE334A" LSID="urn:lsid:plazi:treatment:577887E6FF85FFDF6B325F71FE12328C:DA0E602BFF85FFDE6D055D43FADE334A" box="[1310,1398,1820,1844]" country="Papua New Guinea" latitude="-5.25" longLatPrecision="78" longitude="145.267" name="Wanang" pageId="20" pageNumber="133">Wanang</location>
(
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,
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)
</materialsCitation>
,
<materialsCitation id="6FB93CADFF85FFDE6C175D62FADF332B" box="[1036,1399,1853,1877]" country="Papua New Guinea" latitude="-9.3" location="Goldie River" longLatPrecision="780" longitude="147.42" pageId="20" pageNumber="133" specimenCount="1">
<location id="DA0E602BFF85FFDE6C175D62FB37332B" LSID="urn:lsid:plazi:treatment:577887E6FF85FFDF6B325F71FE12328C:DA0E602BFF85FFDE6C175D62FB37332B" box="[1036,1183,1853,1877]" country="Papua New Guinea" latitude="-9.3" longLatPrecision="780" longitude="147.42" name="Goldie River" pageId="20" pageNumber="133">Goldie River</location>
(
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,
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)
</materialsCitation>
<materialsCitation id="6FB93CADFF85FFDE6B095D02FB203308" box="[786,1160,1885,1910]" country="Papua New Guinea" latitude="-8.77" location="Popondetta" longLatPrecision="781" longitude="148.24" pageId="20" pageNumber="133" specimenCount="1">
and
<location id="DA0E602BFF85FFDE6B5B5D02FC11330B" LSID="urn:lsid:plazi:treatment:577887E6FF85FFDF6B325F71FE12328C:DA0E602BFF85FFDE6B5B5D02FC11330B" box="[832,953,1885,1909]" country="Papua New Guinea" latitude="-8.77" longLatPrecision="781" longitude="148.24" name="Popondetta" pageId="20" pageNumber="133">Popondetta</location>
(
<geoCoordinate id="BAE55037FF85FFDE6BD35D01FBB93308" box="[968,1041,1886,1910]" degrees="8.77" direction="south" orientation="latitude" pageId="20" pageNumber="133" precision="555" value="-8.77">8.77°S</geoCoordinate>
,
<geoCoordinate id="BAE55037FF85FFDE6C005D01FB293308" box="[1051,1153,1886,1910]" degrees="148.24" direction="east" orientation="longitude" pageId="20" pageNumber="133" precision="555" value="148.24">148.24°E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF85FFDF6B325D21FD2032CE" lastPageId="21" lastPageNumber="134" pageId="20" pageNumber="133" type="diagnosis">
<paragraph id="DF6E36F0FF85FFDE6B325D21FADE33C9" blockId="20.[786,1399,1392,1976]" pageId="20" pageNumber="133">
<emphasis id="EDA5EAE2FF85FFDE6B325D21FC3D33E8" bold="true" box="[809,917,1918,1942]" pageId="20" pageNumber="133">Diagnosis</emphasis>
: --Worker (Figs 1922,
<tableCitation id="9253034BFF85FFDE6CA45D21FAA333E8" box="[1215,1291,1918,1942]" captionStart="Table 1" captionStartId="26.[151,204,245,266]" captionText="Table 1. Workers of the C. wroughtonii group of Orientalic-Australasian origin with mean FL/FR &lt;1.040. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="20" pageNumber="133">Tab. 1</tableCitation>
, pictures CASENT0914964 in ww.antweb.org): Very small size,
</paragraph>
<paragraph id="DF6E36F0FF84FFDF68A65AADFD003230" blockId="21.[189,802,241,1778]" pageId="21" pageNumber="134">
CS 380. Head moderately long (CL/CW 1.137); with maximum CL and CW in visual plane, its posterior margin straight or very slightly concave and anterior clypeal margin slightly concave. Postocular distance rather large, PoOc/CL 0.440. Scape rather long, SL/CS 0.808. Eye relatively large, EYE/CS 0.242. Frons rather narrow with short and almost parallel frontal carinae, FRS/CS 0.299, FL/FR 1.017. Mesosoma slender; dorsal profile of promesonotum and propodeum slightly convex but with a notable metanotal depression. Anterior pronotum in dorsal view rounded, without pronounced corners. Propodeal spines short (SP/CS 0.185), in dorsal view slightly diverging, in lateral view nearly straight and their axis deviating 2628° from longitudinal axis of mesosoma. Petiole wide and high (PeW/CS 0.319, PeH/CS 0.363), in lateral view with a short peduncle, a concave anterior margin, a weakly convex dorsum of node that more or less linearly slopes down to caudal cylinder; petiole in dorsal view with a semiglobular node which is slightly wider than long. Postpetiole moderately wide and very high (PpW/CS 0.470, PpH/CS 0.356), in dorsal view much wider than long and with a concave anterior and slightly convex posterior margin when these margins are adjusted to the same focal level; the sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Whole surface of dorsal head with circular foveolae that show a flat central tubercle, mean dFov 15.5 µm; the interspaces between foveolae on vertex smaller than foveolar diameter and delicately longitudinally striate (Fig. 22). Microsculpture on dorsum of mesosoma weaker than on vertex, microreticulate-corrugated and without foveolae; lateral mesosoma microreticulate. Petiole laterally finely microreticulate, dorsally more microcorrugated. Postpetiole finely microcorrugated and in overall appearance matt. Basal part of 1
<superScript id="28A49BB8FF84FFDF6A9E5FCFFD2731E0" attach="left" box="[645,655,1424,1438]" fontSize="6" pageId="21" pageNumber="134">st</superScript>
gaster tergite very finely microreticulate and appearing rather shiny at lower magnifications. Pubescence on 1st gaster tergite of medium length and rather dilute (PLG/CS 6.44 %, sqPDG 4.36]. Gaster jet black and all remaining body parts excluding the eyes very pale yellowish brown.
</paragraph>
<paragraph id="DF6E36F0FF84FFDF68CF5C09FD2032CE" blockId="21.[189,802,241,1778]" pageId="21" pageNumber="134">
<emphasis id="EDA5EAE2FF84FFDF68CF5C09FE7D3211" bold="true" box="[212,469,1622,1647]" pageId="21" pageNumber="134">Taxonomic comments.</emphasis>
As a combination of high and wide postpetiole, long scape and (probably) diagnostic coloration,
<taxonomicName id="18D14D73FF84FFDF692C5CC6FE6032CE" authorityName="Seifert" authorityYear="2023" box="[311,456,1688,1712]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="134" phylum="Arthropoda" rank="species" species="allonivalis">
<emphasis id="EDA5EAE2FF84FFDF692C5CC6FE6032CE" box="[311,456,1688,1712]" italics="true" pageId="21" pageNumber="134">C. allonivalis</emphasis>
</taxonomicName>
is not to confuse.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF84FFDF68CF5CE6FE12328C" pageId="21" pageNumber="134" type="biology_ecology">
<paragraph id="DF6E36F0FF84FFDF68CF5CE6FE12328C" blockId="21.[189,802,241,1778]" pageId="21" pageNumber="134">
<emphasis id="EDA5EAE2FF84FFDF68CF5CE6FE8132AF" bold="true" box="[212,297,1721,1745]" pageId="21" pageNumber="134">Biology</emphasis>
. One nest was found by P.S. Ward in soil of a semi-dry littoral forest.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,201 @@
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<treatment id="577887E6FF86FFD268CF5F2FFEBF366E" LSID="urn:lsid:plazi:treatment:577887E6FF86FFD268CF5F2FFEBF366E" httpUri="http://treatment.plazi.org/id/577887E6FF86FFD268CF5F2FFEBF366E" lastPageId="24" lastPageNumber="137" pageId="23" pageNumber="136">
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<heading id="8426819CFF86FFDD68CF5F2FFDF531F6" box="[212,605,1392,1416]" fontSize="10" level="2" pageId="23" pageNumber="136" reason="4">
<taxonomicName id="18D14D73FF86FFDD68CF5F2FFDF531F6" authority="Rigato 2002" authorityName="Rigato" authorityYear="2002" box="[212,605,1392,1416]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD68CF5F2FFDF531F6" bold="true" box="[212,605,1392,1416]" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD68CF5F2FFE6431F6" bold="true" box="[212,460,1392,1416]" italics="true" pageId="23" pageNumber="136">Cardiocondyla yoruba</emphasis>
<bibRefCitation id="BB404B01FF86FFDD69C85F2FFDF531F6" author="Rigato, F." box="[467,605,1392,1416]" pageId="23" pageNumber="136" pagination="167 - 173" refId="ref19389" refString="Rigato, F. (2002): Three new Afrotropical Cardiocondyla Emery, with a revised key to the workers (Hymenoptera Formicidae). - Bollettino della Societa Entomologica Italiana 134: 167 - 173." type="journal article" year="2002">Rigato 2002</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<heading id="8426819CFF86FFDD68CF5FCEFE1831D7" bold="true" box="[212,432,1425,1449]" fontSize="10" level="2" pageId="23" pageNumber="136" reason="3">
<emphasis id="EDA5EAE2FF86FFDD68CF5FCEFE1831D7" bold="true" box="[212,432,1425,1449]" pageId="23" pageNumber="136">
[
<typeStatus id="006A8852FF86FFDD68C05FCEFEA731D7" box="[219,271,1425,1449]" pageId="23" pageNumber="136">type</typeStatus>
investigation]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF86FFDD68CF5FEDFE1A330B" pageId="23" pageNumber="136" type="description">
<paragraph id="DF6E36F0FF86FFDD68CF5FEDFEB032F1" blockId="23.[189,802,1458,1976]" pageId="23" pageNumber="136">
<materialsCitation id="6FB93CADFF86FFDD68CF5FEDFEBC32F1" collectionCode="IITA, T, BMNH" country="Nigeria" county="Ibadan" location="This" municipality="Investigated" pageId="23" pageNumber="136" specimenCode="CASENT0901752" specimenCount="3" specimenCount-worker="3" stateProvince="Niger" typeStatus="holotype">
<location id="DA0E602BFF86FFDD68CF5FEDFEAD31B4" LSID="urn:lsid:plazi:treatment:577887E6FF86FFD268CF5F2FFEBF366E:DA0E602BFF86FFDD68CF5FEDFEAD31B4" box="[212,261,1458,1482]" country="Nigeria" county="Ibadan" municipality="Investigated" name="This" pageId="23" pageNumber="136" stateProvince="Niger">This</location>
taxon has been described from
<collectingRegion id="1D15F812FF86FFDD6A4D5FEDFD3D31B4" box="[598,661,1458,1482]" country="Nigeria" name="Niger" pageId="23" pageNumber="136">Niger</collectingRegion>
.
<collectingMunicipality id="3F0AAC8AFF86FFDD6A855FEDFC8A31B4" box="[670,802,1458,1482]" pageId="23" pageNumber="136">Investigated</collectingMunicipality>
were
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on the same pin with the site equal to the
<typeStatus id="006A8852FF86FFDD69725FABFE6F3272" box="[361,455,1524,1548]" pageId="23" pageNumber="136" type="holotype">holotype</typeStatus>
(see
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in www. antweb.org), labelled
<collectingCountry id="A7C67660FF86FFDD69D85C4AFD9D3253" box="[451,565,1557,1581]" name="Nigeria" pageId="23" pageNumber="136">NIGERIA</collectingCountry>
<collectingCounty id="360F4E7CFF86FFDD6A5C5C4AFD3B3253" box="[583,659,1557,1581]" pageId="23" pageNumber="136">Ibadan</collectingCounty>
,
<collectionCode id="B9C0AE35FF86FFDD6AB15C4AFD4D3253" box="[682,741,1557,1581]" country="Nigeria" httpUri="http://grbio.org/cool/8h49-s777" name="International Institute of Tropical Agriculture" pageId="23" pageNumber="136">IITA</collectionCode>
.xi.87,
<collectionCode id="B9C0AE35FF86FFDD68A65C69FF663230" box="[189,206,1590,1614]" name="Tavera, Department of Geology and Geophysics" pageId="23" pageNumber="136">T</collectionCode>
.Noyes (handwritten),
<taxonomicName id="18D14D73FF86FFDD69C45C69FD763230" authorityName="Seifert" authorityYear="2024" box="[479,734,1590,1614]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba" status="sp. nov.">Cardiocondyla yoruba</taxonomicName>
<taxonomicNameLabel id="F6965799FF86FFDD6AF45C69FC893233" box="[751,801,1590,1613]" pageId="23" pageNumber="136" rank="species">n.sp.</taxonomicNameLabel>
<typeStatus id="006A8852FF86FFDD68A65C08FEEA3211" box="[189,322,1623,1647]" pageId="23" pageNumber="136">PARATYPI</typeStatus>
, det. F.
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(handwritten),
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London
</materialsCitation>
.
</paragraph>
<paragraph id="DF6E36F0FF86FFDD68CF5CC7FEAA336D" blockId="23.[189,802,1458,1976]" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD68CF5CC7FE6632CE" bold="true" box="[212,462,1688,1712]" pageId="23" pageNumber="136">Allmaterialexamined</emphasis>
.
<materialsCitation id="6FB93CADFF86FFDD69CC5CC7FF56336D" country="Ivory Coast" location="Bondoukou" pageId="23" pageNumber="136" specimenCount="11" specimenCount-worker="11">
Morphometricallyinvestigated were two samples with
<specimenCount id="C9D7FD79FF86FFDD69DE5CE5FDCD32AF" box="[453,613,1721,1745]" count="7" pageId="23" pageNumber="136" type="worker">seven workers</specimenCount>
: the
<typeStatus id="006A8852FF86FFDD6A875CE5FD6432AC" box="[668,716,1722,1746]" pageId="23" pageNumber="136">type</typeStatus>
sample plus a sample of
<specimenCount id="C9D7FD79FF86FFDD699A5C85FDA6328C" box="[385,526,1754,1778]" count="4" pageId="23" pageNumber="136" type="worker">four workers</specimenCount>
from
<location id="DA0E602BFF86FFDD6A435C85FD7D328C" LSID="urn:lsid:plazi:treatment:577887E6FF86FFD268CF5F2FFEBF366E:DA0E602BFF86FFDD6A435C85FD7D328C" box="[600,725,1754,1778]" country="Ivory Coast" name="Bondoukou" pageId="23" pageNumber="136">Bondoukou</location>
/
<collectingCountry id="A7C67660FF86FFDD6AFC5C85FF56336D" name="Ivory Coast" pageId="23" pageNumber="136">Ivory Coast</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="DF6E36F0FF86FFDD68CF5D44FE1A330B" blockId="23.[189,802,1458,1976]" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD68CF5D44FE01334A" bold="true" box="[212,425,1819,1844]" pageId="23" pageNumber="136">Geographic range</emphasis>
. Known so far only from West Africa:
<collectingCountry id="A7C67660FF86FFDD69155D62FEFD332B" box="[270,341,1853,1877]" name="Ghana" pageId="23" pageNumber="136">Ghana</collectingCountry>
(by image evaluation of CASENT0901752),
<collectingCountry id="A7C67660FF86FFDD68A65D01FE923308" box="[189,314,1886,1910]" name="Ivory Coast" pageId="23" pageNumber="136">Ivory Coast</collectingCountry>
and
<collectingCountry id="A7C67660FF86FFDD69745D02FE06330B" box="[367,430,1885,1909]" name="Niger" pageId="23" pageNumber="136">Niger</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF86FFD268CF5D21FDC83535" lastPageId="24" lastPageNumber="137" pageId="23" pageNumber="136" type="diagnosis">
<paragraph id="DF6E36F0FF86FFDD68CF5D21FC8A33C9" blockId="23.[189,802,1458,1976]" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD68CF5D21FEEB33E8" bold="true" box="[212,323,1918,1942]" pageId="23" pageNumber="136">Diagnosis</emphasis>
: --Worker (
<tableCitation id="9253034BFF86FFDD69D75D21FDB033E8" box="[460,536,1918,1942]" captionStart="Table 2" captionStartId="26.[150,203,1476,1497]" captionText="Table 2. Workers of the C. wroughtonii group with mean FL/FR&gt; 1.040 and originating from Africa and theArabian Peninsula. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="23" pageNumber="136">Tab. 2</tableCitation>
, Figs 610, key). Very small, CS 380 µm. Head very long, CL/CW 1.247.
</paragraph>
<paragraph id="DF6E36F0FF86FFDD6B235AAEFB143210" blockId="23.[824,1437,241,1976]" pageId="23" pageNumber="136">Median third of anterior clypeal margin straight, median third of occipital margin slightly concave. Postocular distance large, PoOc/CL 0.459. Frons relatively narrow (FRS/CS 0.245); frontal carinae slightly diverging frontal of FRS level (FL/FR 1.063), caudal of FRS level parallel or slightly converging. Eye rather small, EYE/CS 0.240 and with scattered microsetae of 67 µm length. Scape very short, SL/CS 0.724. Metanotal depression shallow (MGr/CS 1.27 %). Propodeal spines short (SP/CS 0.120) and almost triangular in lateral view, their axis in profile deviating from longitudinal mesosomal axis by 25°, in dorsal view not diverging and their bases moderately wide (SPBA/CS 0.275). Petiole rather narrow and moderately high (PeW/CS 0.274, PeH/CS 0.334); in profile with a concave anterior face and semicircular dorsum; in dorsal view with an almost globular node, which is only slightly longer than wide. Postpetiole narrow and low (PpW/CS 0.427, PpH/CS 0.288); postpetiolar sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Postpetiole in dorsal view with a straight to slightly concave anterior margin and convex sides. Lateral parts of clypeus each with 23 rugulae, central part slightly carinulate-reticulate. Frontal laminae with weak microsculpture, consisting of an irregular mixture of microrugulate, foveolate and microreticulate elements. Whole vertex foveolate, mediad of the eyes with densely-arranged, flat-bottomed foveolae of 1416 µm diameter. The largest foveolae show a well-demarcated but flat central tubercule of 68 µm diameter around the hair bases (Fig. 9). Whole surface of mesosoma foveolate-microreticulate; petiole strongly and postpetiolar tergite less strongly microreticulate. First gaster tergite with moderately long and dense pubescence (PLG/CS 6.89 %, sqPDG 3.86) and a more strongly developed microreticum, with the hair bases placed in microfoveolae of 46 µm diameter (Fig. 10). The microreticulum is an optical illusion in perpendicular view on the surface that is produced by the margins of roof-tile-like surface structures. Color: whole dorsum of gaster blackish brown and all remaining body parts yellowish or yellowish-brown.</paragraph>
<paragraph id="DF6E36F0FF86FFD26B545C28FDC83535" blockId="23.[824,1437,241,1976]" lastBlockId="24.[151,764,241,529]" lastPageId="24" lastPageNumber="137" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD6B545C28FBFC32F1" bold="true" box="[847,1108,1655,1679]" pageId="23" pageNumber="136">Taxonomic comments.</emphasis>
<taxonomicName id="18D14D73FF86FFDD6C7A5C27FB7B32F1" authorityName="Seifert" authorityYear="2024" box="[1121,1235,1655,1679]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD6C7A5C27FB7B32F1" box="[1121,1235,1655,1679]" italics="true" pageId="23" pageNumber="136">C. yoruba</emphasis>
</taxonomicName>
is inseparable in a PCA from
<taxonomicName id="18D14D73FF86FFDD6BD05CC6FB9232CE" authorityName="Seifert" authorityYear="2024" box="[971,1082,1689,1712]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yemeni">
<emphasis id="EDA5EAE2FF86FFDD6BD05CC6FB9232CE" box="[971,1082,1689,1712]" italics="true" pageId="23" pageNumber="136">C. yemeni</emphasis>
</taxonomicName>
considering all 16 NUMOBAT characters: the
<typeStatus id="006A8852FF86FFDD6BFA5CE5FBB932AC" box="[993,1041,1722,1746]" pageId="23" pageNumber="136">type</typeStatus>
samples of
<taxonomicName id="18D14D73FF86FFDD6C825CE5FB4D32AF" authorityName="Seifert" authorityYear="2024" box="[1177,1253,1722,1745]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yemeni">
<emphasis id="EDA5EAE2FF86FFDD6C825CE5FB4D32AF" box="[1177,1253,1722,1745]" italics="true" pageId="23" pageNumber="136">yemeni</emphasis>
</taxonomicName>
and
<taxonomicName id="18D14D73FF86FFDD6D045CE6FAC432AF" authorityName="Seifert" authorityYear="2024" box="[1311,1388,1721,1745]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD6D045CE6FAC432AF" box="[1311,1388,1721,1745]" italics="true" pageId="23" pageNumber="136">yoruba</emphasis>
</taxonomicName>
and the non-type sample of
<taxonomicName id="18D14D73FF86FFDD6C535C85FB3D328C" authorityName="Seifert" authorityYear="2024" box="[1096,1173,1754,1778]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD6C535C85FB3D328C" box="[1096,1173,1754,1778]" italics="true" pageId="23" pageNumber="136">yoruba</emphasis>
</taxonomicName>
from
<collectingCountry id="A7C67660FF86FFDD6CFB5C85FAF5328F" box="[1248,1373,1754,1778]" name="Ivory Coast" pageId="23" pageNumber="136">Ivory coast</collectingCountry>
form a coherent cluster in a PCA (
<figureCitation id="47EA2A75FF86FFDD6C615CA4FB62336D" box="[1146,1226,1787,1811]" captionStart="Figure 54" captionStartId="24.[151,215,1886,1908]" captionTargetBox="[151,1399,1160,1864]" captionTargetId="figure-477@24.[169,1205,1180,1850]" captionTargetPageId="24" captionText="Figure 54. Principal component analysis of worker individuals considering all 16 morphometric characters. Triangles: Cardiocondyla weserka (grey: type of weserka, black: type of neferka, white: no type). Grey discs: syntypes of Cardiocondyla yemeni (Black rhombs: types of Cardiocondyla yoruba. White rhombs: C. yoruba sample from Comoe / Ivory coast). Black squares: Holotype and eight paratypes of Cardiocondyla heinzei n.sp. from Comoe / Ivory coast." pageId="23" pageNumber="136">Fig. 54</figureCitation>
). However, I stood back from synonymization of
<taxonomicName id="18D14D73FF86FFDD6C605D43FB60334A" authorityName="Seifert" authorityYear="2024" box="[1147,1224,1820,1844]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD6C605D43FB60334A" box="[1147,1224,1820,1844]" italics="true" pageId="23" pageNumber="136">yoruba</emphasis>
</taxonomicName>
because of the very remote allopatric ranges, the blackish brown gaster not observed so far in the
<taxonomicName id="18D14D73FF86FFDD6C2A5D01FBD5330B" authorityName="Seifert" authorityYear="2024" box="[1073,1149,1886,1909]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yemeni">
<emphasis id="EDA5EAE2FF86FFDD6C2A5D01FBD5330B" box="[1073,1149,1886,1909]" italics="true" pageId="23" pageNumber="136">yemeni</emphasis>
</taxonomicName>
samples from
<collectingCountry id="A7C67660FF86FFDD6D395D01FAC5330B" box="[1314,1389,1886,1909]" name="Yemen" pageId="23" pageNumber="136">Yemen</collectingCountry>
and
<collectingCountry id="A7C67660FF86FFDD6B235D21FCD233E8" box="[824,890,1918,1942]" name="Oman" pageId="23" pageNumber="136">Oman</collectingCountry>
and because of the stronger microsculpture of
<taxonomicName id="18D14D73FF86FFDD6B235DC0FC2D33C9" authorityName="Seifert" authorityYear="2024" box="[824,901,1951,1975]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD6B235DC0FC2D33C9" box="[824,901,1951,1975]" italics="true" pageId="23" pageNumber="136">yoruba</emphasis>
</taxonomicName>
in particular on first gaster tergite. We have to wait if
<taxonomicName id="18D14D73FF89FFD268F85AAEFE983577" authorityName="Seifert" authorityYear="2024" box="[227,304,241,265]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="137" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF89FFD268F85AAEFE983577" box="[227,304,241,265]" italics="true" pageId="24" pageNumber="137">yoruba</emphasis>
</taxonomicName>
might be more convingly demonstrated as a separate cryptic species when plenty of samples were analyzed morphometrically or genetically.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF89FFD268B55B0BFEBF366E" pageId="24" pageNumber="137" type="biology_ecology">
<paragraph id="DF6E36F0FF89FFD268B55B0BFEBF366E" blockId="24.[151,764,241,529]" pageId="24" pageNumber="137">
<emphasis id="EDA5EAE2FF89FFD268B55B0BFEA33512" bold="true" box="[174,267,340,364]" pageId="24" pageNumber="137">Biology.</emphasis>
The species is seems to follow other
<taxonomicName id="18D14D73FF89FFD2688C5B2AFE3D35F3" box="[151,405,373,397]" pageId="24" pageNumber="137">
<emphasis id="EDA5EAE2FF89FFD2688C5B2AFE9135F3" box="[151,313,373,397]" italics="true" pageId="24" pageNumber="137">Cardiocondyla</emphasis>
species
</taxonomicName>
by living in anthropogenously disturbed areas. As habitats were reported a grassy patch adjacent to a paved road in the urban center of Accra /
<collectingCountry id="A7C67660FF89FFD2688C5B87FF76358E" box="[151,222,472,496]" name="Ghana" pageId="24" pageNumber="137">Ghana</collectingCountry>
and a path in the suburban area of Bondoukou /
<collectingCountry id="A7C67660FF89FFD2688C5BA6FEBB366E" box="[151,275,505,529]" name="Ivory Coast" pageId="24" pageNumber="137">Ivory coast</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,137 @@
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<subSubSection id="97CB657BFF87FFDC6B325E59FBAD3041" pageId="22" pageNumber="135" type="nomenclature">
<paragraph id="DF6E36F0FF87FFDC6B325E59FACE3060" blockId="22.[809,1382,1030,1055]" box="[809,1382,1030,1055]" pageId="22" pageNumber="135">
<heading id="8426819CFF87FFDC6B325E59FACE3060" box="[809,1382,1030,1055]" centered="true" fontSize="10" level="2" pageId="22" pageNumber="135" reason="4">
<taxonomicName id="18D14D73FF87FFDC6B325E59FACE3060" authority="Collingwood &amp; Agosti 1996" authorityName="Collingwood &amp; Agosti" authorityYear="1996" box="[809,1382,1030,1055]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="135" phylum="Arthropoda" rank="species" species="yemeni">
<emphasis id="EDA5EAE2FF87FFDC6B325E59FACE3060" bold="true" box="[809,1382,1030,1055]" pageId="22" pageNumber="135">
<emphasis id="EDA5EAE2FF87FFDC6B325E59FB8B3061" bold="true" box="[809,1059,1030,1055]" italics="true" pageId="22" pageNumber="135">Cardiocondyla yemeni</emphasis>
Collingwood &amp; Agosti 1996
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DF6E36F0FF87FFDC6B325E77FBAD3041" blockId="22.[809,1029,1063,1088]" box="[809,1029,1063,1088]" pageId="22" pageNumber="135">
<heading id="8426819CFF87FFDC6B325E77FBAD3041" bold="true" box="[809,1029,1063,1088]" fontSize="10" level="2" pageId="22" pageNumber="135" reason="3">
<emphasis id="EDA5EAE2FF87FFDC6B325E77FBAD3041" bold="true" box="[809,1029,1063,1088]" pageId="22" pageNumber="135">
[
<typeStatus id="006A8852FF87FFDC6B2B5E77FCCC303E" box="[816,868,1064,1088]" pageId="22" pageNumber="135">type</typeStatus>
investigation]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF87FFDC6B325E17FAF13195" pageId="22" pageNumber="135" type="description">
<paragraph id="DF6E36F0FF87FFDC6B325E17FBC13138" blockId="22.[786,1399,1096,1975]" pageId="22" pageNumber="135">
This taxon has been described from
<collectingCountry id="A7C67660FF87FFDC6C855E16FB45301E" box="[1182,1261,1097,1120]" name="Yemen" pageId="22" pageNumber="135">Yemen</collectingCountry>
. Investigated wasonetypeworkerlabelledinCollingwoodshandwriting
<collectingCountry id="A7C67660FF87FFDC6B0D5ED5FCD730DC" box="[790,895,1162,1186]" name="Yemen" pageId="22" pageNumber="135">YEMEN</collectingCountry>
:
<collectingRegion id="1D15F812FF87FFDC6B9C5ED5FC4D30DC" box="[903,997,1162,1186]" country="Yemen" name="San'a'" pageId="22" pageNumber="135">SANAA</collectingRegion>
(sandy path)
<date id="AB6F1030FF87FFDC6C695ED5FB4230DC" box="[1138,1258,1162,1186]" pageId="22" pageNumber="135" value="1993-03-05">05 03 1993</date>
, Typus and
<taxonomicName id="18D14D73FF87FFDC6B0D5EF4FC1030BD" authorityName="Emery" authorityYear="1869" box="[790,952,1195,1219]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="135" phylum="Arthropoda" rank="genus">Cardiocondyla</taxonomicName>
yemeni; one topotypical worker, without postpetiole and gaster, labelled
<emphasis id="EDA5EAE2FF87FFDC6C735E92FAC6309A" box="[1128,1390,1228,1252]" italics="true" pageId="22" pageNumber="135">
<collectingCountry id="A7C67660FF87FFDC6C735E92FB68309A" box="[1128,1216,1229,1252]" name="Yemen" pageId="22" pageNumber="135">YEMEN</collectingCountry>
<collectingRegion id="1D15F812FF87FFDC6CDD5E93FAA7309D" box="[1222,1295,1228,1251]" country="Yemen" name="San'a'" pageId="22" pageNumber="135">Sanaa</collectingRegion>
17 III 93
</emphasis>
(day not clearly legible, other possible reading
<emphasis id="EDA5EAE2FF87FFDC6D195EB2FAC4317B" box="[1282,1388,1261,1285]" italics="true" pageId="22" pageNumber="135">27 III 93</emphasis>
) and
<taxonomicName id="18D14D73FF87FFDC6B5F5F51FB953158" authorityName="Seifert" authorityYear="2024" box="[836,1085,1294,1318]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="135" phylum="Arthropoda" rank="species" species="yemeni" status="sp. nov.">Cardiocondyla yemeni</taxonomicName>
<taxonomicNameLabel id="F6965799FF87FFDC6C5E5F51FBDF315B" box="[1093,1143,1294,1317]" pageId="22" pageNumber="135" rank="species">n.sp.</taxonomicNameLabel>
Collingwood &amp; Agosti 1996; depository SMN Görlitz.
</paragraph>
<paragraph id="DF6E36F0FF87FFDC6B325F10FB9731F6" blockId="22.[786,1399,1096,1975]" pageId="22" pageNumber="135">
All material examined. Morphometrically investigated were only the
<typeStatus id="006A8852FF87FFDC6BB05F2EFC7331F7" box="[939,987,1393,1417]" pageId="22" pageNumber="135">type</typeStatus>
samples.
</paragraph>
<paragraph id="DF6E36F0FF87FFDC6B325FCEFAF13195" blockId="22.[786,1399,1096,1975]" pageId="22" pageNumber="135">
<emphasis id="EDA5EAE2FF87FFDC6B325FCEFBAA31D7" bold="true" box="[809,1026,1425,1449]" pageId="22" pageNumber="135">Geographic range</emphasis>
. Known so far only from the Arabian Peninsula:
<collectingCountry id="A7C67660FF87FFDC6C1D5FECFBF931B4" box="[1030,1105,1459,1482]" name="Yemen" pageId="22" pageNumber="135">Yemen</collectingCountry>
and
<collectingCountry id="A7C67660FF87FFDC6CB85FEDFB4D31B4" box="[1187,1253,1458,1482]" name="Oman" pageId="22" pageNumber="135">Oman</collectingCountry>
(by image evaluation of CASENT0922296 in www.antweb.org).
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF87FFDD6B325FABFDC730DC" lastPageId="23" lastPageNumber="136" pageId="22" pageNumber="135" type="diagnosis">
<paragraph id="DF6E36F0FF87FFDC6B325FABFADF33C9" blockId="22.[786,1399,1096,1975]" pageId="22" pageNumber="135">
<emphasis id="EDA5EAE2FF87FFDC6B325FABFC323272" bold="true" box="[809,922,1524,1548]" pageId="22" pageNumber="135">Diagnosis</emphasis>
: --Worker (
<tableCitation id="9253034BFF87FFDC6C3B5FABFBC33272" box="[1056,1131,1524,1548]" captionStart="Table 2" captionStartId="26.[150,203,1476,1497]" captionText="Table 2. Workers of the C. wroughtonii group with mean FL/FR&gt; 1.040 and originating from Africa and theArabian Peninsula. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="22" pageNumber="135">Tab. 2</tableCitation>
,
<figureCitation id="47EA2A75FF87FFDC6C615FABFB5A3272" box="[1146,1266,1524,1548]" captionStart="Figure 1115" captionStartId="8.[151,215,1755,1777]" captionTargetBox="[151,1399,259,1706]" captionTargetPageId="8" captionText="Figure 1115. Cardiocondyla yemeni, type, Sanaa / Yemen, 5 March 1993. (11) head in dorsal view; (12) lateral view; (13) dorsal view; (14) head surface between inner eye margin and paramedian vertex. (15) anterior dorsum of 1st gaster tergite. Figure 16. Cardiocondyla nana, holotype, Ulu Temburong / Brunei 22 February 1982; from www.antweb.org CASENT0901757, images by Ryan Perry." pageId="22" pageNumber="135">Figs 1115</figureCitation>
, key). Very small, CS 377 µm. Head very long, CL/CW 1.260. Median third of anterior clypeal margin straight, median third of occipital margin slightly concave. Postocular distance large, PoOc/CL 0.453. Frons relatively narrow (FRS/CS 0.242); frontal carinae slightly diverging frontal of FRS level (FL/FR 1.063), caudal of FRS level parallel or slightly converging. Eye rather small, EYE/ CS 0.238. Scape very short, SL/CS 0.724. Metanotal depression nearly absent (MGr/CS 0.55 %). Propodeal spines short (SP/CS 0.103) and almost triangular in lateral view, their axis in profile deviating from longitudinal mesosomal axis by 25°, in dorsal view not diverging and their bases relatively narrow (SPBA/CS 0.266).
</paragraph>
<paragraph id="DF6E36F0FF86FFDD68A65AAEFDA4301E" blockId="23.[189,802,241,1251]" pageId="23" pageNumber="136">
Petiole rather narrow and moderately high (PeW/CS 0.276, PeH/CS 0.336); in profile with a concave anterior face and a semicircular dorsum; in dorsal view with an almost globular node, which is only slightly longer than wide. Postpetiole narrow and low (PpW/CS 0.430, PpH/CS 0.268); postpetiolar sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Postpetiole in dorsal view with a straight to slightly concave anterior margin and convex sides. Lateral parts of clypeus each with 23 rugulae, central part slightly carinulate-reticulate. Frontal laminae with weak microsculpture, consisting of an irregular mixture of microrugulate, foveolate and microreticulate elements. Whole vertex foveolate, mediad of the eyes with densely-arranged, flat-bottomed foveolae of 1416 µm diameter. The largest foveolae show a well-demarcated but flat central tubercule of 68 µm diameter around the hair bases (
<figureCitation id="47EA2A75FF86FFDD6AD9593DFCBD3704" box="[706,789,866,890]" captionStart="Figure 1115" captionStartId="8.[151,215,1755,1777]" captionTargetBox="[151,1399,259,1706]" captionTargetPageId="8" captionText="Figure 1115. Cardiocondyla yemeni, type, Sanaa / Yemen, 5 March 1993. (11) head in dorsal view; (12) lateral view; (13) dorsal view; (14) head surface between inner eye margin and paramedian vertex. (15) anterior dorsum of 1st gaster tergite. Figure 16. Cardiocondyla nana, holotype, Ulu Temburong / Brunei 22 February 1982; from www.antweb.org CASENT0901757, images by Ryan Perry." pageId="23" pageNumber="136">Fig. 14</figureCitation>
). Whole surface of mesosoma foveolate-microreticulate; petiole and postpetiolar tergite microreticulate. First gaster tergite with moderately long and dense pubescence (PLG/CS 6.78 %, sqPDG 3.76) and a weakly developed microreticulum, with the hair bases placed in microfoveolae of 46 µm diameter. All body parts yellowish or yellowish-brown.
</paragraph>
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<emphasis id="EDA5EAE2FF86FFDD68CF5E36FE7230FF" bold="true" box="[212,474,1129,1153]" pageId="23" pageNumber="136">Taxonomic comments.</emphasis>
For separation from closely related
<taxonomicName id="18D14D73FF86FFDD69165ED5FDAA30DC" box="[269,514,1162,1186]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD69165ED5FDAA30DC" box="[269,514,1162,1186]" italics="true" pageId="23" pageNumber="136">Cardiocondyla yoruba</emphasis>
</taxonomicName>
see there.
</paragraph>
</subSubSection>
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<emphasis id="EDA5EAE2FF86FFDD68CF5EF4FE9830BD" bold="true" box="[212,304,1195,1219]" pageId="23" pageNumber="136">Biology.</emphasis>
As habitat was reported a sandy path in Saana /
<collectingCountry id="A7C67660FF86FFDD68DF5E93FEBB309D" box="[196,275,1228,1251]" name="Yemen" pageId="23" pageNumber="136">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="EDA5EAE2FF9EFFC56B545881FBC73688" bold="true" box="[847,1135,734,758]" italics="true" pageId="15" pageNumber="128">Cardiocondyla longispina</emphasis>
Karavajev 1935
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[
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investigated]
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This taxon has been described from
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. Investigated were
<specimenCount id="C9D7FD79FF9EFFC56C1B591EFBC93727" box="[1024,1121,833,857]" count="5" pageId="15" pageNumber="128" type="worker">5 worker</specimenCount>
<typeStatus id="006A8852FF9EFFC56C72591DFB613724" box="[1129,1225,834,858]" pageId="15" pageNumber="128" type="syntype">syntypes</typeStatus>
labelled Tjibodas,
<collectingRegion id="1D15F812FF9EFFC56B23593CFCCF3704" box="[824,871,867,890]" country="Indonesia" name="Jawa Timur" pageId="15" pageNumber="128">Java</collectingRegion>
Karavaiev\ 5377. Coll. Karavaievi \
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Typus; IZ
<collectingRegion id="1D15F812FF9EFFC56C6E59DCFB0337E5" box="[1141,1195,899,923]" country="Ukraine" name="Kyiv" pageId="15" pageNumber="128">Kiev</collectingRegion>
. The type workers are allocated to the
<taxonomicName id="18D14D73FF9EFFC56BFF59FBFB2E37C2" authorityName="Forel" authorityYear="1890" box="[996,1158,932,956]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF9EFFC56BFF59FBFB2E37C2" box="[996,1158,932,956]" italics="true" pageId="15" pageNumber="128">C. wroughtonii</emphasis>
</taxonomicName>
cluster with p = 0.9997 if run as wild-card in a LDA (for details see below).
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<emphasis id="EDA5EAE2FF9EFFC56B545E59FBC63060" bold="true" box="[847,1134,1030,1054]" italics="true" pageId="15" pageNumber="128">Cardiocondyla yamauchii</emphasis>
Terayama 1999
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[
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</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF9EFFC56B545E17FC563158" pageId="15" pageNumber="128" type="description">
<paragraph id="DF6E36F0FF9EFFC56B545E17FC563158" blockId="15.[824,1438,1096,1975]" pageId="15" pageNumber="128">
<collectingRegion id="1D15F812FF9EFFC56B545E17FC18301E" box="[847,944,1096,1120]" country="Japan" name="Okinawa" pageId="15" pageNumber="128">Okinawa</collectingRegion>
/
<collectingCountry id="A7C67660FF9EFFC56BDE5E17FBA9301E" box="[965,1025,1096,1120]" name="Japan" pageId="15" pageNumber="128">Japan</collectingCountry>
<specimenCount id="C9D7FD79FF9EFFC56C155E17FBDB301E" box="[1038,1139,1096,1120]" count="3" pageId="15" pageNumber="128" type="worker">3 worker</specimenCount>
<typeStatus id="006A8852FF9EFFC56C645E16FB41301F" box="[1151,1257,1097,1121]" pageId="15" pageNumber="128" type="paratype">paratypes</typeStatus>
from the same sample as
<typeStatus id="006A8852FF9EFFC56BA65E36FBB730FF" box="[957,1055,1129,1153]" pageId="15" pageNumber="128" type="holotype">holotype</typeStatus>
: Ada, Okinawa-jima
<collectingRegion id="1D15F812FF9EFFC56D205E36FCC230DC" country="Japan" name="Okinawa" pageId="15" pageNumber="128">Okinawa Pref.</collectingRegion>
<date id="AB6F1030FF9EFFC56B665ED5FC4630DC" box="[893,1006,1162,1186]" pageId="15" pageNumber="128" value="1991-06-12">12.VI.1991</date>
K.Yamauchi leg. and
<taxonomicName id="18D14D73FF9EFFC56CE15ED5FBC230BD" authority="Terayama, 1999" authorityName="Terayama" authorityYear="1999" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="yamauchii">Cardiocondyla yamauchii Terayama, 1999</taxonomicName>
, Paratype, SMN Goerlitz. The type workers are allocated to the
<taxonomicName id="18D14D73FF9EFFC56CEF5E93FA34309A" authorityName="Forel" authorityYear="1890" box="[1268,1436,1228,1252]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF9EFFC56CEF5E93FA34309A" box="[1268,1436,1228,1252]" italics="true" pageId="15" pageNumber="128">C. wroughtonii</emphasis>
</taxonomicName>
cluster with p = 0.9958 if run as wild-card in a LDA (for details see below).
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF9EFFC56B545F71FC18330B" pageId="15" pageNumber="128" type="materials_examined">
<paragraph id="DF6E36F0FF9EFFC56B545F71FB0232F1" blockId="15.[824,1438,1096,1975]" pageId="15" pageNumber="128">
<emphasis id="EDA5EAE2FF9EFFC56B545F71FBF23138" bold="true" box="[847,1114,1326,1350]" pageId="15" pageNumber="128">All material examined</emphasis>
. Numeric phenotypical data were available in 64 samples (51 nest samples and 17 single-specimen stray samples) with
<specimenCount id="C9D7FD79FF9EFFC56CC75F2FFAC031F6" box="[1244,1384,1392,1416]" count="140" pageId="15" pageNumber="128" type="worker">140 workers</specimenCount>
. For details see supplementary information SI1, SI2. Excluding single-specimen samples with unclear separation from
<taxonomicName id="18D14D73FF9EFFC56B235F8CFC683195" authorityName="Wheeler" authorityYear="1929" box="[824,960,1491,1515]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF9EFFC56B235F8CFC683195" box="[824,960,1491,1515]" italics="true" pageId="15" pageNumber="128">C. obscurior</emphasis>
</taxonomicName>
, this material originated from
<collectingCountry id="A7C67660FF9EFFC56D0B5F8CFADC3195" box="[1296,1396,1491,1515]" name="Australia" pageId="15" pageNumber="128">Australia</collectingCountry>
(10 samples),
<collectingCountry id="A7C67660FF9EFFC56BBF5FABFC463272" box="[932,1006,1524,1548]" name="Brunei" pageId="15" pageNumber="128">Brunei</collectingCountry>
(1), the Comores (1),
<collectingCountry id="A7C67660FF9EFFC56CC75FABFAB53272" box="[1244,1309,1524,1548]" name="Egypt" pageId="15" pageNumber="128">Egypt</collectingCountry>
(1), Hawaii (4),
<collectingCountry id="A7C67660FF9EFFC56B7A5C4AFC313253" box="[865,921,1557,1581]" name="India" pageId="15" pageNumber="128">India</collectingCountry>
(1),
<collectingCountry id="A7C67660FF9EFFC56BDC5C4AFB983253" box="[967,1072,1557,1581]" name="Indonesia" pageId="15" pageNumber="128">Indonesia</collectingCountry>
(7),
<collectingCountry id="A7C67660FF9EFFC56C7B5C4AFB353252" box="[1120,1181,1557,1580]" name="Japan" pageId="15" pageNumber="128">Japan</collectingCountry>
(3), Kenia (1),
<collectingCountry id="A7C67660FF9EFFC56D215C4AFA343253" box="[1338,1436,1557,1581]" name="Malaysia" pageId="15" pageNumber="128">Malaysia</collectingCountry>
(13),
<collectingCountry id="A7C67660FF9EFFC56B765C69FC7F3230" box="[877,983,1590,1614]" name="Mauritius" pageId="15" pageNumber="128">Mauritius</collectingCountry>
(3),
<collectingCountry id="A7C67660FF9EFFC56C135C69FBE03230" box="[1032,1096,1590,1614]" name="Nepal" pageId="15" pageNumber="128">Nepal</collectingCountry>
(1),
<collectingCountry id="A7C67660FF9EFFC56C6C5C69FB593230" box="[1143,1265,1590,1614]" name="Philippines" pageId="15" pageNumber="128">Philippines</collectingCountry>
(5),
<collectingCountry id="A7C67660FF9EFFC56D395C69FC2F3210" name="Papua New Guinea" pageId="15" pageNumber="128">Papua New Guinea</collectingCountry>
(2),
<collectingCountry id="A7C67660FF9EFFC56BDE5C09FB9A3210" box="[965,1074,1622,1646]" name="Singapore" pageId="15" pageNumber="128">Singapore</collectingCountry>
(1), Sri Lanka (1),
<collectingCountry id="A7C67660FF9EFFC56D055C09FAC53210" box="[1310,1389,1622,1646]" name="Taiwan" pageId="15" pageNumber="128">Taiwan</collectingCountry>
(1), Tanzania (1), Tailand (4),
<collectingCountry id="A7C67660FF9EFFC56C505C28FBD732F1" box="[1099,1151,1655,1679]" name="United States of America" pageId="15" pageNumber="128">USA</collectingCountry>
(5).
</paragraph>
<paragraph id="DF6E36F0FF9EFFC56B545CC7FC18330B" blockId="15.[824,1438,1096,1975]" pageId="15" pageNumber="128">
<emphasis id="EDA5EAE2FF9EFFC56B545CC7FB8932CE" bold="true" box="[847,1057,1688,1712]" pageId="15" pageNumber="128">Geographic range</emphasis>
. As a tramp species of putatively Southeast Asian origin this species is widely distributed over the tropical regions of the world. However, it has not been confirmed so far for South and Central America where
<taxonomicName id="18D14D73FF9EFFC56B9E5D43FBBB334A" authorityName="Wheeler" authorityYear="1929" box="[901,1043,1820,1844]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF9EFFC56B9E5D43FBBB334A" box="[901,1043,1820,1844]" italics="true" pageId="15" pageNumber="128">C. obscurior</emphasis>
</taxonomicName>
is found. Occurrence in buildings in the temperate zone is so far not verified by voucher specimens.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF9EFFDA6B545D21FB143253" lastPageId="16" lastPageNumber="129" pageId="15" pageNumber="128" type="diagnosis">
<paragraph id="DF6E36F0FF9EFFC56B545D21FA3433C9" blockId="15.[824,1438,1096,1975]" pageId="15" pageNumber="128">
<emphasis id="EDA5EAE2FF9EFFC56B545D21FC1733E8" bold="true" box="[847,959,1918,1942]" pageId="15" pageNumber="128">Diagnosis</emphasis>
: --Worker (
<tableCitation id="9253034BFF9EFFC56C5E5D21FB2533E8" box="[1093,1165,1918,1942]" captionStart="Table 1" captionStartId="26.[151,204,245,266]" captionText="Table 1. Workers of the C. wroughtonii group of Orientalic-Australasian origin with mean FL/FR &lt;1.040. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="15" pageNumber="128">Tab. 1</tableCitation>
, Figs 2831, key). Very small and slightly smaller than
<taxonomicName id="18D14D73FF9EFFC56CD85DC0FACE33C9" authority=", CS" authorityName="CS" box="[1219,1382,1951,1975]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="128" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF9EFFC56CD85DC0FA8433C9" box="[1219,1324,1951,1975]" italics="true" pageId="15" pageNumber="128">obscurior</emphasis>
, CS
</taxonomicName>
410
</paragraph>
<paragraph id="DF6E36F0FF81FFDA688C5AADFD8932CE" blockId="16.[151,765,241,1975]" pageId="16" pageNumber="129">
µm. Head short, CL/CW 1.120. Anterocentral clypeal margin straight or slightly notched; central occipital margin usually straight or with a very weak concavity. Postocular distance large, PoOc/CL 0.440. Frons broad (FRS/CS 0.272), frontal carinae caudal of FRS level parallel (FL/FR 1.008). Eye medium-sized, EYE/CS 0.233. Scape slightly shorter than in
<taxonomicName id="18D14D73FF81FFDA6A4D5BE8FD5B35B1" authority=", SL" authorityName="SL" box="[598,755,439,463]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA6A4D5BE8FD1735B1" box="[598,703,439,463]" italics="true" pageId="16" pageNumber="129">obscurior</emphasis>
, SL
</taxonomicName>
/ CS 0.772. Promesonotal plane in dorsal view without shoulders due to a strongly convex frontal margin and rather straight lateral margins (
<figureCitation id="47EA2A75FF81FFDA6A125846FDDB364C" box="[521,627,537,562]" captionStart="Figure 23" captionStartId="10.[151,215,1755,1777]" captionTargetBox="[151,1398,260,1705]" captionTargetPageId="10" captionText="Figure 23. Mesosoma in dorsal view.(A) Cardiocondyla obscurior, neotype, Nago / Okinawa, 3 June 2003; (B) Cardiocondyla shagrinata, paratype, South Konkan / India. Figure 2427. Cardiocondyla shagrinata, paratype, South Konkan / India. (24) head in dorsal view; (25) lateral view; (26) dorsal view; (27) head surface between inner eye margin and paramedian vertex. Figure 28. Cardiocondyla wroughtonii, Lundu / Malaysia, 27 May 2007. (28) head in dorsal view." pageId="16" pageNumber="129">Fig. 23A</figureCitation>
). Metanotal groove in lateral view deep (MGr/CS 3.65 %) and usually with steep anterior and posterior slopes. Propodeal spines longer than in
<taxonomicName id="18D14D73FF81FFDA69B25823FDBB36EA" box="[425,531,636,660]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA69B25823FDBB36EA" box="[425,531,636,660]" italics="true" pageId="16" pageNumber="129">obscurior</emphasis>
</taxonomicName>
(SP/CS 0.193) and their bases more approached (SPBA/CS 0.278). Petiole narrower and slightly lower than in
<taxonomicName id="18D14D73FF81FFDA6A5958E1FD0536A8" box="[578,685,702,726]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA6A5958E1FD0536A8" box="[578,685,702,726]" italics="true" pageId="16" pageNumber="129">obscurior</emphasis>
</taxonomicName>
(PeW/ CS 0.277, PeH/CS 0.328), Axis of petiolar peduncle in lateral aspect deviating by 30° from the longitudinal axis of the petiole node. Postpetiole narrower and lower than in
<taxonomicName id="18D14D73FF81FFDA68A8591EFEB53727" box="[179,285,833,857]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA68A8591EFEB53727" box="[179,285,833,857]" italics="true" pageId="16" pageNumber="129">obscurior</emphasis>
</taxonomicName>
(PpW/CS 0.437, PpH/CS 0.303), the sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Whole head and mesosoma without any notable rugosity. Paramedian and lateral vertex with deep (sometimes shallower) foveolae of 1721 µm diameter which are frequently arranged in dense honey-comb arrangement and show an inner elevation around the hair bases of 69 µm diameter (
<figureCitation id="47EA2A75FF81FFDA69735ED5FE1D30DC" box="[360,437,1162,1186]" captionStart="Figure 2931" captionStartId="11.[189,253,1757,1779]" captionTargetBox="[189,1437,260,1707]" captionTargetPageId="11" captionText="Figure 2931. Cardiocondyla wroughtonii, Lundu / Malaysia, 27 May 2007.(29) lateral view; (30) dorsal view; (31) head surface between inner eye margin and paramedian vertex. Figure 3234. Cardiocondyla obscurior, neotype, Nago / Okinawa, 3 June 2003. (32) head in dorsal view; (33) lateral view; (34) dorsal view." pageId="16" pageNumber="129">Fig. 31</figureCitation>
); median vertex with slightly smaller and weaker foveolae, occasionally weakly foveolate-carinulate. Whole surface of mesosoma and waist densely foveolate-reticulate, the meshes with lower diameters than on vertex (
<figureCitation id="47EA2A75FF81FFDA69E95F51FDC23158" box="[498,618,1294,1318]" captionStart="Figure 2931" captionStartId="11.[189,253,1757,1779]" captionTargetBox="[189,1437,260,1707]" captionTargetPageId="11" captionText="Figure 2931. Cardiocondyla wroughtonii, Lundu / Malaysia, 27 May 2007.(29) lateral view; (30) dorsal view; (31) head surface between inner eye margin and paramedian vertex. Figure 3234. Cardiocondyla obscurior, neotype, Nago / Okinawa, 3 June 2003. (32) head in dorsal view; (33) lateral view; (34) dorsal view." pageId="16" pageNumber="129">Figs 29, 30</figureCitation>
). Pubescence on first gaster tergite moderately long and dilute (PLG/ CS 6.3 %, sqPDG 5.05). Color dimorphism. Light morph (76 % of samples): whole ant entirely light yellowish except for a diffuse brown band in the posterior half of 1
<superScript id="28A49BB8FF81FFDA68BF5FEEFF0731C1" attach="left" box="[164,175,1457,1471]" fontSize="6" pageId="16" pageNumber="129">st</superScript>
gaster tergite; this band may be interrupted in the centre to form 2 separated lateral patches (as seen in Wheeler´s
<taxonomicName id="18D14D73FF81FFDA69115FABFE1B3272" authorityName="Wheeler" authorityYear="1929" box="[266,435,1524,1548]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="variety" species="wroughtonii" variety="bimaculata">
var.
<emphasis id="EDA5EAE2FF81FFDA69225FABFE1B3272" box="[313,435,1524,1548]" italics="true" pageId="16" pageNumber="129">bimaculata</emphasis>
</taxonomicName>
), PigG1 9.5 ± 8.2 [0, 45] % n= 108. Dark morph (24 % of samples): Head, mesosoma, waist, and appendages light yellowish brown; funicular club blackish brown; 1
<superScript id="28A49BB8FF81FFDA69885C0AFE36321D" attach="left" box="[403,414,1621,1635]" fontSize="6" pageId="16" pageNumber="129">st</superScript>
gaster tergite and sternite dark brown, the following segments substantially lighter, PigG1 86.8 ± 19.1 [55, 100] %, n=34.
</paragraph>
<paragraph id="DF6E36F0FF81FFDA68B55CE6FB143253" blockId="16.[151,765,241,1975]" lastBlockId="16.[786,1399,242,1975]" pageId="16" pageNumber="129">
<emphasis id="EDA5EAE2FF81FFDA68B55CE6FD6332AF" bold="true" box="[174,715,1721,1745]" pageId="16" pageNumber="129">Taxonomic comments and clustering results</emphasis>
.
<taxonomicName id="18D14D73FF81FFDA6AF85CE5FEB0328C" authorityName="Forel" authorityYear="1890" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF81FFDA6AF85CE5FEB0328C" italics="true" pageId="16" pageNumber="129">C. wroughtonii</emphasis>
</taxonomicName>
is extremely similar to
<taxonomicName id="18D14D73FF81FFDA6A225C85FD6F328C" box="[569,711,1754,1778]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA6A225C85FD6F328C" box="[569,711,1754,1778]" italics="true" pageId="16" pageNumber="129">C. obscurior</emphasis>
</taxonomicName>
and the occurrence of a dark morph in
<taxonomicName id="18D14D73FF81FFDA6A075CA4FD35336D" authorityName="Forel" authorityYear="1890" box="[540,669,1787,1811]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF81FFDA6A075CA4FD35336D" box="[540,669,1787,1811]" italics="true" pageId="16" pageNumber="129">wroughtonii</emphasis>
</taxonomicName>
restricts the use of pigmentation for species separation. Species hypotheses were formed in 84 samples of both species with
<specimenCount id="C9D7FD79FF81FFDA68CA5D02FEE7330B" box="[209,335,1885,1909]" count="205" pageId="16" pageNumber="129" type="worker">205 worker</specimenCount>
individuals under exclusion of single-specimen samples. In the first step of analysis, the 15 characters CS, CL/CW, PoOc/CL, SL/CS, EYE/CS, FRS/ CS, SPBA/CS, SP/CS, PeW/CS, PeH/CS, PPW/CS, PpH/CS, MGr/CS, sqPDG and PLG/CS were run in the exploratory data analyses NC-Ward, NC-NMDS.kmeans and PCA. Hypotheses were fixed in the controlling LDA when the classifications in the three exploratory data analyses coincided whereas samples with controversial classification were run as wild-cards. In the next run of the LDA the corrected classifications were accepted but now the type samples were run as wild-cards. The posterior probabilities for allocation to the
<taxonomicName id="18D14D73FF81FFDA6CCC5845FADE364F" authorityName="Forel" authorityYear="1890" box="[1239,1398,537,561]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF81FFDA6CCC5845FADE364F" box="[1239,1398,537,561]" italics="true" pageId="16" pageNumber="129">C. wroughtonii</emphasis>
</taxonomicName>
cluster were 1.0000 in the types of
<taxonomicName id="18D14D73FF81FFDA6C9F5865FAAD362C" authorityName="Forel" authorityYear="1890" box="[1156,1285,570,594]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF81FFDA6C9F5865FAAD362C" box="[1156,1285,570,594]" italics="true" pageId="16" pageNumber="129">wroughtonii</emphasis>
</taxonomicName>
,
<quantity id="18299B15FF81FFDA6D095865FADF362C" box="[1298,1399,570,594]" metricMagnitude="-2" metricUnit="m" metricValue="2.539746" pageId="16" pageNumber="129" unit="in" value="0.9999">0.9999 in</quantity>
the type of
<emphasis id="EDA5EAE2FF81FFDA6B975804FBAB360D" box="[908,1027,603,627]" italics="true" pageId="16" pageNumber="129">hawaiensis</emphasis>
,
<quantity id="18299B15FF81FFDA6C0B5804FBDD360D" box="[1040,1141,603,627]" metricMagnitude="-2" metricUnit="m" metricValue="2.539238" pageId="16" pageNumber="129" unit="in" value="0.9997">0.9997 in</quantity>
the types of
<taxonomicName id="18D14D73FF81FFDA6CE45804FAD8360D" authorityName="Karavajev" authorityYear="1935" box="[1279,1392,603,627]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="longispina">
<emphasis id="EDA5EAE2FF81FFDA6CE45804FAD8360D" box="[1279,1392,603,627]" italics="true" pageId="16" pageNumber="129">longispina</emphasis>
</taxonomicName>
,
<quantity id="18299B15FF81FFDA6B095823FCD136EA" box="[786,889,636,660]" metricMagnitude="-2" metricUnit="m" metricValue="2.53873" pageId="16" pageNumber="129" unit="in" value="0.9995">0.9995 in</quantity>
the types of
<emphasis id="EDA5EAE2FF81FFDA6C105823FB3E36EA" box="[1035,1174,636,660]" italics="true" pageId="16" pageNumber="129">quadraticeps</emphasis>
,
<quantity id="18299B15FF81FFDA6CBE5823FAA436EA" box="[1189,1292,636,660]" metricMagnitude="-2" metricUnit="m" metricValue="2.535936" pageId="16" pageNumber="129" unit="in" value="0.9984">0.9984 in</quantity>
the types of
<emphasis id="EDA5EAE2FF81FFDA6B2B58C2FC0036CB" box="[816,936,669,693]" italics="true" pageId="16" pageNumber="129">bimaculata</emphasis>
and
<quantity id="18299B15FF81FFDA6BC658C2FBEB36CB" box="[989,1091,669,693]" metricMagnitude="-2" metricUnit="m" metricValue="2.529332" pageId="16" pageNumber="129" unit="in" value="0.9958">0.9958 in</quantity>
the types of
<taxonomicName id="18D14D73FF81FFDA6CCB58C2FA9736CB" authorityName="Terayama" authorityYear="1999" box="[1232,1343,669,693]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="yamauchii">
<emphasis id="EDA5EAE2FF81FFDA6CCB58C2FA9736CB" box="[1232,1343,669,693]" italics="true" pageId="16" pageNumber="129">yamauchii</emphasis>
</taxonomicName>
. The posterior probabilities for allocation to the
<taxonomicName id="18D14D73FF81FFDA6CF658E1FADF36A8" box="[1261,1399,702,726]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA6CF658E1FADF36A8" box="[1261,1399,702,726]" italics="true" pageId="16" pageNumber="129">C. obscurior</emphasis>
</taxonomicName>
cluster were
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the
<typeStatus id="006A8852FF81FFDA6C325880FBD53689" box="[1065,1149,735,759]" pageId="16" pageNumber="129" type="neotype">neotype</typeStatus>
series of
<taxonomicName id="18D14D73FF81FFDA6CF95880FAE23689" box="[1250,1354,735,759]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA6CF95880FAE23689" box="[1250,1354,735,759]" italics="true" pageId="16" pageNumber="129">obscurior</emphasis>
</taxonomicName>
and
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the types of
<taxonomicName id="18D14D73FF81FFDA6C1F595FFBF93766" authorityName="Donisthorpe" authorityYear="1930" box="[1028,1105,768,792]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="EDA5EAE2FF81FFDA6C1F595FFBF93766" box="[1028,1105,768,792]" italics="true" pageId="16" pageNumber="129">bicolor</emphasis>
</taxonomicName>
. As a third step, a stepwise character reduction to CS, CL/CW, SL/CS, EYE/CS, FRS/ CS, SP/CS and PPW/CS was performed.This increased the agreement of the exploratory data analyses with the final species hypothesis determined by the controlling LDA to 96.4 % in NC-Ward, 98.8 % in NC-NMDS.kmeans, 98.8 % in NC-part.kmeans (
<figureCitation id="47EA2A75FF81FFDA6C26599AFB2337A3" box="[1085,1163,965,989]" captionStart="Figure 52" captionStartId="17.[189,253,1910,1932]" captionText="Figure 52. Classification of 84 nest samples of Cardiocondyla wroughtonii and C. obscurior by the exploratory data analyses NC-Ward (hierarchical), NC-part.kmeans, NC-NMDS.kmeans and PCA based on 7 morphometric characters taken in 205 worker individuals. The mean error of the four exploratory data analyses relative to the controlling LDA (final.sp.hyp) is 1.8%." pageId="16" pageNumber="129">Fig. 52</figureCitation>
) and to 98.8 % and in the PCA (
<figureCitation id="47EA2A75FF81FFDA6B9B59B9FC613780" box="[896,969,998,1022]" captionStart="Figure 53" captionStartId="18.[151,215,1887,1909]" captionTargetBox="[153,1401,1199,1865]" captionTargetId="graphics-518@18.[155,1400,1199,1865]" captionTargetPageId="18" captionText="Figure 53. Principal component analysis of 84 nest samples of Cardiocondyla wroughtonii and C. obscurior based on 7 morphometric characters taken in 205 worker individuals.The arrow points to the sample determined as C. obscurior by a wild-card LDA with p = 0.9456 which, however, showed a pigmentation typical for C. wroughtonii. The mean error of the four exploratory data analyses relative to the controlling LDA (final.sp.hyp) is 1.8%." pageId="16" pageNumber="129">Fig.53</figureCitation>
). This is a mean error in four forms of analysis of only 1.8 %. In laboratory experiments, males of
<taxonomicName id="18D14D73FF81FFDA6B285E77FC163041" box="[819,958,1063,1087]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA6B285E77FC163041" box="[819,958,1063,1087]" italics="true" pageId="16" pageNumber="129">C. obscurior</emphasis>
</taxonomicName>
and gynes of
<taxonomicName id="18D14D73FF81FFDA6C7F5E77FAA03041" authorityName="Forel" authorityYear="1890" box="[1124,1288,1063,1087]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF81FFDA6C7F5E77FAA03041" box="[1124,1288,1063,1087]" italics="true" pageId="16" pageNumber="129">C. wroughtonii</emphasis>
</taxonomicName>
produced hybrids but the opposite mating combination so far not (K. Yamauchi pers. comm. 2000). Hybridisation is supposed to be rare or absent under natural conditions as concluded from the strong separation of the clusters. However, the available sample size, in particular the low number of 2.4 workers per sample, makes assessment of hybridization frequency difficult. There is so far only one suggestion on a possible hybridization: the only sample of
<taxonomicName id="18D14D73FF81FFDA6D145F71FADF3138" box="[1295,1399,1326,1350]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA6D145F71FADF3138" box="[1295,1399,1326,1350]" italics="true" pageId="16" pageNumber="129">obscurior</emphasis>
</taxonomicName>
with a gaster pigmentation corresponding to light morph of
<taxonomicName id="18D14D73FF81FFDA6B295F2FFC1B31F6" authorityName="Forel" authorityYear="1890" box="[818,947,1392,1416]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="wroughtonii">
<emphasis id="EDA5EAE2FF81FFDA6B295F2FFC1B31F6" box="[818,947,1392,1416]" italics="true" pageId="16" pageNumber="129">wroughtonii</emphasis>
</taxonomicName>
(SaNo 1072
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) was allocated to the
<taxonomicName id="18D14D73FF81FFDA6B215FCEFC0A31D7" box="[826,930,1425,1449]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA6B215FCEFC0A31D7" box="[826,930,1425,1449]" italics="true" pageId="16" pageNumber="129">obscurior</emphasis>
</taxonomicName>
cluster with p=0.9456 if run as a wild-card in a LDA of the reduced 7-characters data set and was placed by the PCA in a marginal position (
<figureCitation id="47EA2A75FF81FFDA6CCE5F8CFA8C3195" box="[1237,1316,1491,1515]" captionStart="Figure 53" captionStartId="18.[151,215,1887,1909]" captionTargetBox="[153,1401,1199,1865]" captionTargetId="graphics-518@18.[155,1400,1199,1865]" captionTargetPageId="18" captionText="Figure 53. Principal component analysis of 84 nest samples of Cardiocondyla wroughtonii and C. obscurior based on 7 morphometric characters taken in 205 worker individuals.The arrow points to the sample determined as C. obscurior by a wild-card LDA with p = 0.9456 which, however, showed a pigmentation typical for C. wroughtonii. The mean error of the four exploratory data analyses relative to the controlling LDA (final.sp.hyp) is 1.8%." pageId="16" pageNumber="129">Fig. 53</figureCitation>
). It was found under the bark of a tree in
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height which supports the determination by the LDA.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF81FFDA6B325C6AFBFD33C9" pageId="16" pageNumber="129" type="biology_ecology">
<paragraph id="DF6E36F0FF81FFDA6B325C6AFBFD33C9" blockId="16.[786,1399,242,1975]" pageId="16" pageNumber="129">
<emphasis id="EDA5EAE2FF81FFDA6B325C6AFCD63233" bold="true" box="[809,894,1589,1613]" pageId="16" pageNumber="129">Biology</emphasis>
. In contrast to its sibling species
<taxonomicName id="18D14D73FF81FFDA6CF35C69FAD83230" box="[1256,1392,1590,1614]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="obscurior">
<emphasis id="EDA5EAE2FF81FFDA6CF35C69FAD83230" box="[1256,1392,1590,1614]" italics="true" pageId="16" pageNumber="129">C. obscurior</emphasis>
</taxonomicName>
, it was reported to nest near to or on the ground; it was found in hollow stems of dead
<taxonomicName id="18D14D73FF81FFDA6C4D5C28FB0D32F1" box="[1110,1189,1655,1679]" class="Insecta" family="Stratiomyidae" genus="Eulalia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="EDA5EAE2FF81FFDA6C4D5C28FB0D32F1" box="[1110,1189,1655,1679]" italics="true" pageId="16" pageNumber="129">Eulalia</emphasis>
</taxonomicName>
grasses (Okinawa), in a dead twig on the ground (New Orleans/
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), between layers of
<taxonomicName id="18D14D73FF81FFDA6BF65CE5FB6032AF" box="[1005,1224,1721,1745]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="129" phylum="Arthropoda" rank="species" species="jambolana">
<emphasis id="EDA5EAE2FF81FFDA6BF65CE5FB6032AF" box="[1005,1224,1721,1745]" italics="true" pageId="16" pageNumber="129">Eugenia jambolana</emphasis>
</taxonomicName>
leaves (
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), in litter (Sulawesi), and under leaves in a silk patch (
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). Nest populations are polygynous and adopt alien queens. There is polyphenism showing winged and wingless (ergatoid) males. The wingless males have sickle-shaped mandibles used to kill male callows or pupae whereas they besmear adult rivals with a secretion that elicits worker aggression.
</paragraph>
</subSubSection>
</treatment>
</document>