Add updates up until 2024-09-05 21:25:49

This commit is contained in:
ggserver 2024-09-05 21:27:55 +00:00
parent 95cf213edf
commit 6031b9e982
35 changed files with 6400 additions and 139 deletions

View file

@ -0,0 +1,846 @@
<document id="348E6588FD317D0D2CBBEBF8A72E25B9" ID-DOI="10.11646/phytotaxa.324.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13697360" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725568056204" checkinUser="felipe" docAuthor="Steyn, Hester M. &amp; Van Wyk, Abraham E." docDate="2017" docId="03F687CD4624FFC6FF2AC9DAFEE753AA" docLanguage="en" docName="phytotaxa.324.2.1.pdf" docOrigin="Phytotaxa 324 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.324.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Acanthopsis glabra H. M. Steyn 2017, comb. nov." docType="treatment" docVersion="2" lastPageNumber="109" masterDocId="FFCFFFB54627FFCEFFA2CD76FF995703" masterDocTitle="Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts" masterLastPageNumber="121" masterPageNumber="101" pageNumber="104" updateTime="1725570993962" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="EC7B7C82A016B49CBAB95FE139BB0DF5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D1DD07CBA30C204BA796A2DFD7589437">
<mods:title id="AC18F1D4C9282C65081C0459F5443675">Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts</mods:title>
</mods:titleInfo>
<mods:name id="8925152E9AFB46940C55CD4795F4D284" type="personal">
<mods:role id="2EC2E6D5EC1040235A216FB77B4BB6EF">
<mods:roleTerm id="D2B6D0CB40A6C6AFC377A1BAA5D77E60">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="53E9208C583D515B2C37A701DCC28C5D">Steyn, Hester M.</mods:namePart>
<mods:affiliation id="E451864361BCBBF74DA3798AFE595165">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:name id="51FB22D36654CCC5E7F3573B257673BB" type="personal">
<mods:role id="E3AE0C0D48600CCDDC9A773F44209963">
<mods:roleTerm id="0BE3FFB24BDFADD49E1A4F7EB034D71C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B2BB7197E5AB4815C9BDF9642EAB769">Van Wyk, Abraham E.</mods:namePart>
<mods:affiliation id="F923EE8D7C4140395890EC1D187B63E9">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="919BBCC8021495B0CF21F216B15CBA89">text</mods:typeOfResource>
<mods:relatedItem id="919648206FF08458062DDFE632AAFF63" type="host">
<mods:titleInfo id="66394036E64CDA0D180A4E34C7D86B49">
<mods:title id="6DDBC459EAF2DBB241700873B3B50E4E">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="11C387D9959924A99E1C321473525CCE">
<mods:date id="8D5816FE1B9CE24373203A92716444D0">2017</mods:date>
<mods:detail id="4DB2187132AC1BA85FF61FD72D2DD8B5" type="pubDate">
<mods:number id="72ED482B2A9599CF11C433B9640A738F">2017-10-10</mods:number>
</mods:detail>
<mods:detail id="229EB255CE530E715E60B67CC11BBAA7" type="volume">
<mods:number id="094665A215234C2FFF0D1D9BD2C906F5">324</mods:number>
</mods:detail>
<mods:detail id="9EBD6E096C4ABD457D00CB5A89FE909C" type="issue">
<mods:number id="7FCD69F0EAB6C982441950C6535C2DB0">2</mods:number>
</mods:detail>
<mods:extent id="7591752B0FC3875B3A72112FEED394D1" unit="page">
<mods:start id="91497664C6B6E760AD9E43568658DB65">101</mods:start>
<mods:end id="19B114BE48BE8460E0C6D9B9289A5678">121</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="E519BCE21F9965F908FAD0C7B9001C95">
<mods:url id="21BA25B8FB1B4E780F62BEEF9A4A2616">http://dx.doi.org/10.11646/phytotaxa.324.2.1</mods:url>
</mods:location>
<mods:classification id="7C16B67C443DA02C032847C9D59670BC">journal article</mods:classification>
<mods:identifier id="095F08F2587981A9BF4540C843BF9A99" type="DOI">10.11646/phytotaxa.324.2.1</mods:identifier>
<mods:identifier id="5A1C83EB2085AB8DEA92EB3B4B2E1F96" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="9FA8B36345D035C322B3A49518B6C41C" type="Zenodo-Dep">13697360</mods:identifier>
</mods:mods>
<treatment id="03F687CD4624FFC6FF2AC9DAFEE753AA" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA" httpUri="http://treatment.plazi.org/id/03F687CD4624FFC6FF2AC9DAFEE753AA" lastPageId="8" lastPageNumber="109" pageId="3" pageNumber="104">
<subSubSection id="C34565504624FFCDFF2AC9DAFC3553C6" box="[136,940,1194,1221]" pageId="3" pageNumber="104" type="nomenclature">
<paragraph id="8BE036DB4624FFCDFF2AC9DAFC3553C6" blockId="3.[136,1452,1194,1400]" box="[136,940,1194,1221]" pageId="3" pageNumber="104">
<heading id="D0A881B74624FFCDFF2AC9DAFC3553C6" box="[136,940,1194,1221]" fontSize="11" level="2" pageId="3" pageNumber="104" reason="4">
<emphasis id="B92BEAC94624FFCDFF2AC9DAFF0453C6" bold="true" box="[136,157,1195,1221]" pageId="3" pageNumber="104">1.</emphasis>
<taxonomicName id="4C5F4D584624FFCDFF01C9DCFDC153C6" ID-CoL="8ZL8" authority="(Nees) H. M. Steyn" authorityName="H. M. Steyn" authorityYear="2017" baseAuthorityName="H. M. Steyn" baseAuthorityYear="2017" box="[163,600,1194,1221]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="104" phylum="Tracheophyta" rank="species" species="glabra" status="comb. nov.">
<emphasis id="B92BEAC94624FFCDFF01C9DCFE1A53C6" bold="true" box="[163,387,1194,1221]" italics="true" pageId="3" pageNumber="104">Acanthopsis glabra</emphasis>
(Nees) H.M.Steyn
</taxonomicName>
,
<taxonomicNameLabel id="A21857B24624FFCDFDC6C9DAFD4753C6" box="[612,734,1196,1221]" pageId="3" pageNumber="104" rank="species">
<emphasis id="B92BEAC94624FFCDFDC6C9DAFD4F53C6" box="[612,726,1196,1221]" italics="true" pageId="3" pageNumber="104">comb. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="13642A5E4624FFCDFD4FC9DAFCA353C6" box="[749,826,1196,1221]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="3" pageNumber="104">Figs. 1</figureCitation>
,
<figureCitation id="13642A5E4624FFCDFCEAC9DAFCF453C6" box="[840,877,1196,1221]" captionStart="FIGURE 3" captionStartId="6.[136,229,1911,1932]" captionTargetBox="[151,1436,190,1886]" captionTargetId="figure-15@6.[151,1436,190,1886]" captionTargetPageId="6" captionText="FIGURE 3. Comparison of floral parts, associated bracts and bracteoles (from left to right): bracts (2), bracteoles (2), calyx (4 sepals: dorsal, ventral and 2 lateral ones) and corolla with anthers.A. A. glabra, B. A. scullyi, C. A. nitida, D. A. glandulopalmata, E. A. insueta, F. A. carduifolia. Note: the bract on the left is from lower down in the same inflorescence. Scale bar = 10 mm. Vouchers:A = Steyn 1893 (PRE), B = Steyn 1911 (PRE), C = Steyn 1884 (PRE), D = Steyn 1890 (PRE), E = Steyn 1805 (PRE), F = Koekemoer 4620 (PRE)." figureDoi="http://doi.org/10.5281/zenodo.13697366" httpUri="https://zenodo.org/record/13697366/files/figure.png" pageId="3" pageNumber="104">3A</figureCitation>
, &amp;
<figureCitation id="13642A5E4624FFCDFC37C9DAFC3D53C6" box="[917,932,1196,1221]" captionStart="FIGURE 4" captionStartId="7.[136,229,1780,1801]" captionTargetBox="[151,1436,175,1755]" captionTargetId="figure-15@7.[151,1436,175,1755]" captionTargetPageId="7" captionText="FIGURE 4. Acanthopsis glabra. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp; F. Bract. G. Calyx. H. Corolla (lobes lilac to cream; throat cream). I. Glandular hairs at base of style. J. Capsule. K. Seed. Vouchers: CF = Acocks 16476 (PRE), GI = Steyn 1893 (PRE), J &amp; K = Steyn 2119 (PRE). Photographed by: H.M. Steyn (A, CK) and M. Koekemoer (B)." figureDoi="http://doi.org/10.5281/zenodo.13697368" httpUri="https://zenodo.org/record/13697368/files/figure.png" pageId="3" pageNumber="104">4</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34565504624FFCDFF2AC9A5FCEB5233" pageId="3" pageNumber="104" type="reference_group">
<paragraph id="8BE036DB4624FFCDFF2AC9A5FCEB5233" blockId="3.[136,1452,1194,1400]" pageId="3" pageNumber="104">
Basionym:—
<taxonomicName id="4C5F4D584624FFCDFEAAC9A5FCBC53EB" ID-CoL="8XGG" authority="Nees von Esenbeck (1847: 278)" authorityName="Nees von Esenbeck" authorityPageNumber="278" authorityYear="1847" box="[264,805,1235,1256]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthodium" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="104" phylum="Tracheophyta" rank="species" species="glabrum">
<emphasis id="B92BEAC94624FFCDFEAAC9A5FE7853EB" box="[264,481,1235,1256]" italics="true" pageId="3" pageNumber="104">Acanthodium glabrum</emphasis>
<bibRefCitation id="EFCE4B2A4624FFCDFE4BC9A5FCBC53EB" author="Nees von Esenbeck, C. G. D." box="[489,805,1235,1256]" pageId="3" pageNumber="104" pagination="46 - 519" refId="ref13072" refString="Nees von Esenbeck, C. G. D. (1847) Acanthaceae. In: Candolle, A. de. (Ed.) Prodromus systematis naturalis regni vegetabilis 11. Treuttel &amp; Wurtz, Paris, pp. 46 - 519." type="book chapter" year="1847">Nees von Esenbeck (1847: 278)</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C5F4D584624FFCDFC92C9A5FDD8520F" ID-CoL="B5LLC" authority="(Nees) Anderson (1864: 35)" authorityName="Anderson" authorityPageNumber="35" authorityYear="1864" baseAuthorityName="Nees" class="Magnoliopsida" family="Acanthaceae" genus="Blepharis" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="104" phylum="Tracheophyta" rank="variety" species="carduifolia" variety="glabra">
<emphasis id="B92BEAC94624FFCDFC92C9A5FB9953EB" box="[816,1024,1235,1256]" italics="true" pageId="3" pageNumber="104">Blepharis carduifolia</emphasis>
(
<bibRefCitation id="EFCE4B2A4624FFCDFBACC9A5FB4B53EB" author="Linnaeus, C." box="[1038,1234,1235,1256]" pageId="3" pageNumber="104" refId="ref12915" refString="Linnaeus, C. (1782) Supplementum plantarum. Impensis Orphanotrophei, Brunsvigae, 502 pp." type="book" year="1782">Linnaeus 1782: 294</bibRefCitation>
)
<bibRefCitation id="EFCE4B2A4624FFCDFB42C9A5FA3253EB" author="Anderson, T." box="[1248,1451,1235,1256]" pageId="3" pageNumber="104" pagination="13 - 54" refId="ref12377" refString="Anderson, T. (1864) An enumeration of the species of Acanthaceae from the continent and the adjacent islands. Journal of the Proceedings of the Linnean Society 7: 13 - 54." type="journal article" year="1864">Anderson (1864: 35)</bibRefCitation>
var.
<emphasis id="B92BEAC94624FFCDFF4AC981FEB1520F" box="[232,296,1271,1292]" italics="true" pageId="3" pageNumber="104">glabra</emphasis>
(Nees)
<bibRefCitation id="EFCE4B2A4624FFCDFED6C981FDD8520F" author="Anderson, T." box="[372,577,1271,1292]" pageId="3" pageNumber="104" pagination="13 - 54" refId="ref12377" refString="Anderson, T. (1864) An enumeration of the species of Acanthaceae from the continent and the adjacent islands. Journal of the Proceedings of the Linnean Society 7: 13 - 54." type="journal article" year="1864">Anderson (1864: 35)</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C5F4D584624FFCDFDE9C981FCAB520F" ID-CoL="8ZKX" authorityName=", Drege" authorityYear="1843" box="[587,818,1271,1292]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="104" phylum="Tracheophyta" rank="species" species="carduifolia">
<emphasis id="B92BEAC94624FFCDFDE9C981FCAB520F" box="[587,818,1271,1292]" italics="true" pageId="3" pageNumber="104">Acanthopsis carduifolia</emphasis>
</taxonomicName>
(L.f.)
<bibRefCitation id="EFCE4B2A4624FFCDFCD0C981FBB5520F" author="Schinz, H." box="[882,1068,1271,1292]" pageId="3" pageNumber="104" pagination="179 - 230" refId="ref13310" refString="Schinz, H. (1890) Beitrage zur Kenntnis der Flora von Deutsch-Sudwest-Afrika und der angrenzenden Gebiete. Verhandlungen des Botanischen Vereins fur die Provinz Brandenbrug und die Angrenzenden Lander 31: 179 - 230." type="journal article" year="1890">Schinz (1890: 200)</bibRefCitation>
<taxonomicName id="4C5F4D584624FFCDFB91C981FA31520F" ID-CoL="8ZL8" authority="(Nees) Schinz (1890: 200)" authorityName="Schinz" authorityPageNumber="200" authorityYear="1890" baseAuthorityName="Nees" box="[1075,1448,1271,1292]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="104" phylum="Tracheophyta" rank="variety" species="carduifolia" variety="glabra">
var.
<emphasis id="B92BEAC94624FFCDFBFFC981FB04520F" box="[1117,1181,1271,1292]" italics="true" pageId="3" pageNumber="104">glabra</emphasis>
(Nees)
<bibRefCitation id="EFCE4B2A4624FFCDFB49C981FA31520F" author="Schinz, H." box="[1259,1448,1271,1292]" pageId="3" pageNumber="104" pagination="179 - 230" refId="ref13310" refString="Schinz, H. (1890) Beitrage zur Kenntnis der Flora von Deutsch-Sudwest-Afrika und der angrenzenden Gebiete. Verhandlungen des Botanischen Vereins fur die Provinz Brandenbrug und die Angrenzenden Lander 31: 179 - 230." type="journal article" year="1890">Schinz (1890: 200)</bibRefCitation>
</taxonomicName>
.
<emphasis id="B92BEAC94624FFCDFF1FC86DFF5C5233" box="[189,197,1307,1328]" italics="true" pageId="3" pageNumber="104"></emphasis>
<taxonomicName id="4C5F4D584624FFCDFF64C86DFE215233" ID-CoL="92SJ" authority="&quot; Meyer" authorityName="Meyer" box="[198,440,1307,1328]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthus" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="104" phylum="Tracheophyta" rank="species" species="glaber">
<emphasis id="B92BEAC94624FFCDFF64C86DFEF05233" box="[198,361,1307,1328]" italics="true" pageId="3" pageNumber="104">Acanthus glaber</emphasis>
” Meyer
</taxonomicName>
in
<bibRefCitation id="EFCE4B2A4624FFCDFE75C86DFDEF5233" author="Drege, J. F." box="[471,630,1307,1328]" pageId="3" pageNumber="104" refId="ref12531" refString="Drege, J. F. (1843) Zwei Pflanzengeographische Documente, nebst einer Einleitung von Dr. E. Meyer. Regensburg, Leipzig, 230 pp. https: // doi. org / 10.5962 / bhl. title. 87612" type="book" year="1843">Drège (1843: 67</bibRefCitation>
, 161),
<emphasis id="B92BEAC94624FFCDFD1AC86DFC885233" box="[696,785,1307,1328]" italics="true" pageId="3" pageNumber="104">nom. nud</emphasis>
(Art. 38).
</paragraph>
</subSubSection>
<subSubSection id="C34565504624FFCDFF2AC849FBAF5F26" pageId="3" pageNumber="104" type="materials_examined">
<paragraph id="8BE036DB4624FFCDFF2AC849FBC9527B" blockId="3.[136,1452,1194,1400]" pageId="3" pageNumber="104">
<emphasis id="B92BEAC94624FFCDFF2AC849FF5B5257" bold="true" box="[136,194,1343,1364]" pageId="3" pageNumber="104">Type:</emphasis>
<collectingCountry id="F348764B4624FFCDFF79C849FE135257" box="[219,394,1343,1364]" name="South Africa" pageId="3" pageNumber="104">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="499BF8394624FFCDFE36C849FDBF5257" box="[404,550,1343,1364]" country="South Africa" name="Northern Cape" pageId="3" pageNumber="104">Northern Cape</collectingRegion>
: Little Namaqualand, Kamies Bergen. [Am Berg bei Kasparskloof, fide
<bibRefCitation id="EFCE4B2A4624FFCDFB42C849FAFA5257" author="Drege, J. F." box="[1248,1379,1343,1364]" pageId="3" pageNumber="104" refId="ref12531" refString="Drege, J. F. (1843) Zwei Pflanzengeographische Documente, nebst einer Einleitung von Dr. E. Meyer. Regensburg, Leipzig, 230 pp. https: // doi. org / 10.5962 / bhl. title. 87612" type="book" year="1843">Drège (1843)</bibRefCitation>
]
<quantity id="4CA79B3E4624FFCDFACDC849FE9B527B" metricMagnitude="3" metricUnit="m" metricValue="1.0668" metricValueMax="1.2192" metricValueMin="0.9144" pageId="3" pageNumber="104" unit="ft" value="3500.0" valueMax="4000.0" valueMin="3000.0">3000 4000 ft</quantity>
[
<quantity id="4CA79B3E4624FFCDFEACC815FE15527B" box="[270,396,1379,1400]" metricMagnitude="3" metricUnit="m" metricValue="1.0675" metricValueMax="1.22" metricValueMin="0.915" pageId="3" pageNumber="104" unit="m" value="1067.5" valueMax="1220.0" valueMin="915.0">9151220 m</quantity>
], [
<date id="FFE1101B4624FFCDFE01C815FDD8527B" box="[419,577,1379,1400]" pageId="3" pageNumber="104" value="1830-08-19">19 August 1830</date>
],
<emphasis id="B92BEAC94624FFCDFDF3C815FD2D527B" box="[593,692,1379,1400]" italics="true" pageId="3" pageNumber="104">Drège s.n.</emphasis>
(
<typeStatus id="54E488794624FFCDFD62C815FC83527B" box="[704,794,1379,1400]" pageId="3" pageNumber="104" type="lectotype">lectotype</typeStatus>
<collectionCode id="ED4EAE1E4624FFCDFC82C815FCAB527B" box="[800,818,1379,1400]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="104" type="Herbarium">K</collectionCode>
000394342! designated here).
</paragraph>
<paragraph id="8BE036DB4624FFCDFF2AC8DEFBAF5F26" blockId="3.[136,1459,1448,2085]" pageId="3" pageNumber="104">
Perennial, acaulescent herb or compact subshrub, up to
<quantity id="4CA79B3E4624FFCDFD59C8DEFCD952C2" box="[763,832,1448,1473]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="104" unit="cm" value="25.0">25 cm</quantity>
tall with very short internodes. Leaves sessile, petiole-like base with spines
<quantity id="4CA79B3E4624FFCDFED5C8BAFE4D52E6" box="[375,468,1484,1509]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="3" pageNumber="104" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
long; lamina oblanceolate, 4570(100) × (6)810(12) mm, often grey to grey-green due to densely packed long, strigose hairs, also with subsessile glands; apex acute, base attenuate, margin undulate, coarsely dentate-spinose, spines fine to rigid, 13(6) mm long, yellow to orange-brown. Inflorescences lax (bract lamina overlap &lt;30%), cylindrical, subsessile with a few infertile bracts at the base, (50)70140(250) mm long, (5)
<quantity id="4CA79B3E4624FFCDFF17CB2AFE8C5176" box="[181,277,1628,1653]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="3" pageNumber="104" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">79 mm</quantity>
in diameter. Bracts oblong to obovate, 1820(23) mm long (including spines), lamina 911(14) mm long, base obtuse; hirsute with deflexed to appressed short hairs, also with isolated, subsessile glandular hairs abaxially, silky-hairy adaxially; middle and upper bracts usually truncate, ending in 5 primary spines; central primary spine usually simple, rarely with 1 pair of short marginal secondary spines; lateral primary spines usually simple; spines spreading (usually 90°) in fruit. Bracteoles linear, (6)
<quantity id="4CA79B3E4624FFCDFD44CB9AFCDA5006" box="[742,835,1772,1797]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="3" pageNumber="104" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
long, pubescent with short, deflexed to spreading hairs and short glandular hairs. Calyx with dorsal sepal ovate, apiculate to acuminate,
<quantity id="4CA79B3E4624FFCDFBAFCA66FB1F502A" box="[1037,1158,1808,1833]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="3" pageNumber="104" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">1314 mm</quantity>
long, hirsute to pubescent with short, deflexed to spreading hairs and short glandular hairs abaxially, silky-hairy adaxially, 79-veined; ventral sepal ovate,
<quantity id="4CA79B3E4624FFCDFEBACA2EFE0B5072" box="[280,402,1880,1905]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="3" pageNumber="104" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">1113 mm</quantity>
long, hirsute to pubescent with short, deflexed to spreading hairs and short glandular hairs abaxially, silky-hairy adaxially, 57-veined; lateral sepals
<quantity id="4CA79B3E4624FFCDFCBCCA0AFCE45096" box="[798,893,1916,1941]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="3" pageNumber="104" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">79 mm</quantity>
long, pubescent with short hairs, also with long, silky hairs along margin. Flowers clearly decussate; corolla cream or lilac with darker veins, cream throat, 2729 × 9(10) mm, tube
<quantity id="4CA79B3E4624FFCDFEEACAB2FE0E50DE" box="[328,407,1988,2013]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="104" unit="mm" value="10.0">10 mm</quantity>
long, pubescent especially on outside, central lobe wider than long or equal, constricted at the base, truncate to emarginate. Filaments
<quantity id="4CA79B3E4624FFCDFDE3CA9EFD045F02" box="[577,669,2024,2049]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="3" pageNumber="104" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
long, glandular; anthers orange-brown,
<quantity id="4CA79B3E4624FFCDFBF8CA9EFB2F5F02" box="[1114,1206,2024,2049]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="104" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long. Style with basal part densely covered with glandular hairs. Capsules 7 × 3(4) mm. Seeds 4 ×
<quantity id="4CA79B3E4624FFCDFC4CC57AFBAB5F26" box="[1006,1074,2060,2085]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="104" unit="mm" value="3.0">3 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF2066534623FFCAFF2AC8F9FB6C52CB" ID-DOI="http://doi.org/10.5281/zenodo.13697362" ID-Zenodo-Dep="13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="4" pageNumber="105" startId="4.[136,229,1423,1444]" targetBox="[151,1436,187,1395]" targetPageId="4" targetType="figure">
<paragraph id="8BE036DB4623FFCAFF2AC8F9FB6C52CB" blockId="4.[136,1452,1423,1480]" pageId="4" pageNumber="105">
<emphasis id="B92BEAC94623FFCAFF2AC8F9FF6752A7" bold="true" box="[136,254,1423,1444]" pageId="4" pageNumber="105">FIGURE 1.</emphasis>
Known distribution of
<taxonomicName id="4C5F4D584623FFCAFE43C8F9FDA552A7" authorityName="H. M. Steyn" authorityYear="2017" baseAuthorityName="H. M. Steyn" baseAuthorityYear="2017" box="[481,572,1423,1444]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="105" phylum="Tracheophyta" rank="species" species="glabra">
<emphasis id="B92BEAC94623FFCAFE43C8F9FDA552A7" box="[481,572,1423,1444]" italics="true" pageId="4" pageNumber="105">A. glabra</emphasis>
</taxonomicName>
(○),
<taxonomicName id="4C5F4D584623FFCAFDCFC8F9FD5E52A7" authorityName="Obermeyer" authorityYear="1937" baseAuthorityName="S. Moore" box="[621,711,1423,1444]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="105" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC94623FFCAFDCFC8F9FD5E52A7" box="[621,711,1423,1444]" italics="true" pageId="4" pageNumber="105">A. scullyi</emphasis>
</taxonomicName>
(▲),
<taxonomicName id="4C5F4D584623FFCAFCA0C8F9FCCC52A7" authorityName="H. M. Steyn" authorityYear="2017" box="[770,853,1423,1444]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="105" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="B92BEAC94623FFCAFCA0C8F9FCCC52A7" box="[770,853,1423,1444]" italics="true" pageId="4" pageNumber="105">A. nitida</emphasis>
</taxonomicName>
(□),
<taxonomicName id="4C5F4D584623FFCAFC24C8F9FBDC52A7" authorityName="H. M. Steyn" authorityYear="2017" box="[902,1093,1423,1444]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="105" phylum="Tracheophyta" rank="species" species="glandulopalmata">
<emphasis id="B92BEAC94623FFCAFC24C8F9FBDC52A7" box="[902,1093,1423,1444]" italics="true" pageId="4" pageNumber="105">A. glandulopalmata</emphasis>
</taxonomicName>
(■)
<taxonomicName id="4C5F4D584623FFCAFBD3C8F9FB4852A7" authorityName="H. M. Steyn" authorityYear="2017" box="[1137,1233,1423,1444]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="105" phylum="Tracheophyta" rank="species" species="insueta">
<emphasis id="B92BEAC94623FFCAFBD3C8F9FB4852A7" box="[1137,1233,1423,1444]" italics="true" pageId="4" pageNumber="105">A. insueta</emphasis>
</taxonomicName>
(Δ) and
<taxonomicName id="4C5F4D584623FFCAFA84C8F9FA3552A7" authorityName=", Drege" authorityYear="1843" box="[1318,1452,1423,1444]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="105" phylum="Tracheophyta" rank="species" species="carduifolia">
<emphasis id="B92BEAC94623FFCAFA84C8F9FA3552A7" box="[1318,1452,1423,1444]" italics="true" pageId="4" pageNumber="105">A. carduifolia</emphasis>
</taxonomicName>
(●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of
<taxonomicName id="4C5F4D584623FFCAFBCBC8C5FB7452CB" authorityName=", Drege" authorityYear="1843" box="[1129,1261,1459,1480]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="105" phylum="Tracheophyta" rank="species" species="carduifolia">
<emphasis id="B92BEAC94623FFCAFBCBC8C5FB7452CB" box="[1129,1261,1459,1480]" italics="true" pageId="4" pageNumber="105">A. carduifolia</emphasis>
</taxonomicName>
)
</paragraph>
</caption>
<subSubSection id="C34565504623FFCAFF1FC88FFDD25135" pageId="4" pageNumber="105" type="etymology">
<paragraph id="8BE036DB4623FFCAFF1FC88FFDD25135" blockId="4.[136,1452,1529,2022]" pageId="4" pageNumber="105">
<emphasis id="B92BEAC94623FFCAFF1FC88FFED15111" bold="true" box="[189,328,1529,1554]" pageId="4" pageNumber="105">Etymology:</emphasis>
—The specific epithet
<taxonomicName id="4C5F4D584623FFCAFDEEC88FFD0E5111" authorityName="H. M. Steyn" authorityYear="2017" baseAuthorityName="H. M. Steyn" baseAuthorityYear="2017" box="[588,663,1529,1554]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="105" phylum="Tracheophyta" rank="species" species="glabra">
<emphasis id="B92BEAC94623FFCAFDEEC88FFD0E5111" box="[588,663,1529,1554]" italics="true" pageId="4" pageNumber="105">glabra</emphasis>
</taxonomicName>
, meaning “smooth/without hairs”, probably refers to the floral bracts which appear glabrous to the naked eye.
</paragraph>
</subSubSection>
<subSubSection id="C34565504623FFC6FF1FCB37FEE753AA" lastPageId="8" lastPageNumber="109" pageId="4" pageNumber="105" type="distribution">
<paragraph id="8BE036DB4623FFCAFF1FCB37FBA95055" blockId="4.[136,1452,1529,2022]" pageId="4" pageNumber="105">
<emphasis id="B92BEAC94623FFCAFF1FCB37FDED5159" bold="true" box="[189,628,1601,1626]" pageId="4" pageNumber="105">Distribution, ecology and phenology:</emphasis>
<taxonomicName id="4C5F4D584623FFCAFD32CB37FCF35159" authorityName="H. M. Steyn" authorityYear="2017" baseAuthorityName="H. M. Steyn" baseAuthorityYear="2017" box="[656,874,1601,1626]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="105" phylum="Tracheophyta" rank="species" species="glabra">
<emphasis id="B92BEAC94623FFCAFD32CB37FCF35159" box="[656,874,1601,1626]" italics="true" pageId="4" pageNumber="105">Acanthopsis glabra</emphasis>
</taxonomicName>
is known from mountain slopes and rocky outcrops on the escarpment from west of Spektakel Pass in the north to just north of Bitterfontein in the south (
<collectingRegion id="499BF8394623FFCAFB58CB13FA3A517D" box="[1274,1443,1637,1662]" country="South Africa" name="Northern Cape" pageId="4" pageNumber="105">Northern Cape</collectingRegion>
) (
<figureCitation id="13642A5E4623FFCAFF32CBFFFF4951A1" box="[144,208,1673,1698]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="4" pageNumber="105">Fig. 1</figureCitation>
) at elevations between 260 and
<quantity id="4CA79B3E4623FFCAFD96CBFFFD1351A1" box="[564,650,1673,1698]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="105" unit="m" value="1500.0">1500 m</quantity>
. This range falls within the Namaqualand Hardeveld Bioregion (
<bibRefCitation id="EFCE4B2A4623FFCAFAF6CBFFFEFF51C5" author="Mucina, L. &amp; Rutherford, M. C." pageId="4" pageNumber="105" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) in the Succulent Karoo Biome (
<bibRefCitation id="EFCE4B2A4623FFCAFD47CBDBFBB251C5" author="Rutherford, M. C. &amp; Westfall, R. H." box="[741,1067,1709,1734]" pageId="4" pageNumber="105" refId="ref13271" refString="Rutherford, M. C. &amp; Westfall, R. H. (1994) Biomes of southern Africa: an objective categorization. [Memoir 63]. National Botanical Institute, Pretoria, 94 pp." type="book" year="1994">Rutherford &amp; Westfall 1994</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A4623FFCAFB9BCBDBFAB051C5" author="Low, A. B. &amp; Rebelo, A. G." box="[1081,1321,1709,1734]" pageId="4" pageNumber="105" refId="ref12933" refString="Low, A. B., Rebelo, A. G. (Eds.) (1996) Vegetation of South Africa, Lesotho and Swaziland. Department of Environmental Affairs &amp; Tourism, Pretoria, 85 pp." type="book" year="1996">Low &amp; Rebelo 1996</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A4623FFCAFA95CBDBFEDF51E9" author="Mucina, L. &amp; Rutherford, M. C." pageId="4" pageNumber="105" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) with the Kamiesberg localities falling in the Namaqualand
<collectingRegion id="499BF8394623FFCAFC5ACBA7FBAB51E9" box="[1016,1074,1745,1770]" country="South Africa" name="Western Cape" pageId="4" pageNumber="105">Cape</collectingRegion>
Shrubland Bioregion (
<bibRefCitation id="EFCE4B2A4623FFCAFA94CBA7FEDD500D" author="Mucina, L. &amp; Rutherford, M. C." pageId="4" pageNumber="105" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) in the Fynbos Biome (
<bibRefCitation id="EFCE4B2A4623FFCAFDEECB83FCAE500D" author="Low, A. B. &amp; Rebelo, A. G." box="[588,823,1781,1806]" pageId="4" pageNumber="105" refId="ref12933" refString="Low, A. B., Rebelo, A. G. (Eds.) (1996) Vegetation of South Africa, Lesotho and Swaziland. Department of Environmental Affairs &amp; Tourism, Pretoria, 85 pp." type="book" year="1996">Low &amp; Rebelo 1996</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A4623FFCAFCE0CB83FBE2500D" author="Mucina, L. &amp; Rutherford, M. C." box="[834,1147,1781,1806]" pageId="4" pageNumber="105" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
). It receives predominantly winter rains with an average of
<quantity id="4CA79B3E4623FFCAFE5BCA6FFD085031" box="[505,657,1817,1842]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="4" pageNumber="105" unit="mm" value="225.0" valueMax="250.0" valueMin="200.0">200250 mm</quantity>
per year. This species prefers deep sandy soils derived from granitic gneisses. Specimens with flowers were collected at the end of August to December.
</paragraph>
<paragraph id="8BE036DB4623FFCAFF1FCA17FCF450E5" blockId="4.[136,1452,1529,2022]" pageId="4" pageNumber="105">
<emphasis id="B92BEAC94623FFCAFF1FCA17FE2C5079" bold="true" box="[189,437,1889,1914]" pageId="4" pageNumber="105">Conservation status:</emphasis>
—This species is widespread and common. Subpopulations are large, and it persists even in heavily overgrazed communal rangelands in the Kamiesberg. As there are no severe threats to this species, the population is not suspected to be declining and it is categorised as Least Concern according to the IUCN Red List Category and Criteria (
<bibRefCitation id="EFCE4B2A4623FFCAFE2BCABBFD8950E5" author="IUCN" box="[393,528,1997,2022]" pageId="4" pageNumber="105" refId="ref12795" refString="IUCN (2012) IUCN Red List categories and criteria version 3.1. IUCN Species Survival Commission, Gland, Switzerland and Cambridge, UK, iv + 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
) (L. von Staden, pers. comm.).
</paragraph>
<caption id="DF2066534622FFCBFF2ACABAFE8D5F2A" ID-DOI="http://doi.org/10.5281/zenodo.13697364" ID-Zenodo-Dep="13697364" httpUri="https://zenodo.org/record/13697364/files/figure.png" pageId="5" pageNumber="106" startId="5.[136,229,1996,2017]" targetBox="[230,1356,176,1970]" targetPageId="5" targetType="figure">
<paragraph id="8BE036DB4622FFCBFF2ACABAFE8D5F2A" blockId="5.[136,1452,1996,2089]" pageId="5" pageNumber="106">
<emphasis id="B92BEAC94622FFCBFF2ACABAFF6750E2" bold="true" box="[136,254,1996,2017]" pageId="5" pageNumber="106">FIGURE 2.</emphasis>
Types of inflorescences found in
<taxonomicName id="4C5F4D584622FFCBFDE7CABAFD2350E2" authorityName="Harvey" authorityYear="1842" box="[581,698,1996,2017]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="106" phylum="Tracheophyta" rank="genus">
<emphasis id="B92BEAC94622FFCBFDE7CABAFD2350E2" box="[581,698,1996,2017]" italics="true" pageId="5" pageNumber="106">Acanthopsis</emphasis>
</taxonomicName>
based on density of individual flowers and their bracts. A. dense (&gt;60% bract lamina overlap), B. semi-dense (3060% bract lamina overlap), C. semi-dense (&lt;30% bract lamina overlap). Scale bar = 10 mm. Artist: Daleen Roodt.
</paragraph>
</caption>
<caption id="DF2066534621FFC8FF2ACA01FAFE50FB" ID-DOI="http://doi.org/10.5281/zenodo.13697366" ID-Zenodo-Dep="13697366" httpUri="https://zenodo.org/record/13697366/files/figure.png" pageId="6" pageNumber="107" startId="6.[136,229,1911,1932]" targetBox="[151,1436,190,1886]" targetPageId="6" targetType="figure">
<paragraph id="8BE036DB4621FFC8FF2ACA01FAFE50FB" blockId="6.[136,1452,1911,2040]" pageId="6" pageNumber="107">
<emphasis id="B92BEAC94621FFC8FF2ACA01FF66508F" bold="true" box="[136,255,1911,1932]" pageId="6" pageNumber="107">FIGURE 3.</emphasis>
Comparison of floral parts, associated bracts and bracteoles (from left to right): bracts (2), bracteoles (2), calyx (4 sepals: dorsal, ventral and 2 lateral ones) and corolla with anthers. A.
<taxonomicName id="4C5F4D584621FFC8FD7DCAEDFCA350B3" authorityName="H. M. Steyn" authorityYear="2017" baseAuthorityName="H. M. Steyn" baseAuthorityYear="2017" box="[735,826,1947,1968]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="107" phylum="Tracheophyta" rank="species" species="glabra">
<emphasis id="B92BEAC94621FFC8FD7DCAEDFCA350B3" box="[735,826,1947,1968]" italics="true" pageId="6" pageNumber="107">A. glabra</emphasis>
</taxonomicName>
, B.
<taxonomicName id="4C5F4D584621FFC8FCC0CAEDFC2550B3" authorityName="Obermeyer" authorityYear="1937" baseAuthorityName="S. Moore" box="[866,956,1947,1968]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="107" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC94621FFC8FCC0CAEDFC2550B3" box="[866,956,1947,1968]" italics="true" pageId="6" pageNumber="107">A. scullyi</emphasis>
</taxonomicName>
, C.
<taxonomicName id="4C5F4D584621FFC8FC41CAEDFBAF50B3" authorityName="H. M. Steyn" authorityYear="2017" box="[995,1078,1947,1968]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="107" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="B92BEAC94621FFC8FC41CAEDFBAF50B3" box="[995,1078,1947,1968]" italics="true" pageId="6" pageNumber="107">A. nitida</emphasis>
</taxonomicName>
, D.
<taxonomicName id="4C5F4D584621FFC8FBFDCAEDFA8750B3" authorityName="H. M. Steyn" authorityYear="2017" box="[1119,1310,1947,1968]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="107" phylum="Tracheophyta" rank="species" species="glandulopalmata">
<emphasis id="B92BEAC94621FFC8FBFDCAEDFA8750B3" box="[1119,1310,1947,1968]" italics="true" pageId="6" pageNumber="107">A. glandulopalmata</emphasis>
</taxonomicName>
, E.
<taxonomicName id="4C5F4D584621FFC8FAE7CAEDFA3C50B3" authorityName="H. M. Steyn" authorityYear="2017" box="[1349,1445,1947,1968]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="107" phylum="Tracheophyta" rank="species" species="insueta">
<emphasis id="B92BEAC94621FFC8FAE7CAEDFACD50B3" box="[1349,1364,1947,1968]" italics="true" pageId="6" pageNumber="107">A</emphasis>
.
<emphasis id="B92BEAC94621FFC8FAC2CAEDFA3C50B3" box="[1376,1445,1947,1968]" italics="true" pageId="6" pageNumber="107">insueta</emphasis>
</taxonomicName>
, F.
<taxonomicName id="4C5F4D584621FFC8FF3DCAC9FEBD50D7" authorityName=", Drege" authorityYear="1843" box="[159,292,1983,2004]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="107" phylum="Tracheophyta" rank="species" species="carduifolia">
<emphasis id="B92BEAC94621FFC8FF3DCAC9FEBD50D7" box="[159,292,1983,2004]" italics="true" pageId="6" pageNumber="107">A. carduifolia</emphasis>
</taxonomicName>
. Note: the bract on the left is from lower down in the same inflorescence. Scale bar = 10 mm. Vouchers: A =
<emphasis id="B92BEAC94621FFC8FAE3CAC9FA3250D7" box="[1345,1451,1983,2004]" italics="true" pageId="6" pageNumber="107">Steyn 1893</emphasis>
(PRE), B =
<emphasis id="B92BEAC94621FFC8FF58CA95FEFB50FB" box="[250,354,2019,2040]" italics="true" pageId="6" pageNumber="107">Steyn 1911</emphasis>
(PRE), C =
<emphasis id="B92BEAC94621FFC8FE7BCA95FDDA50FB" box="[473,579,2019,2040]" italics="true" pageId="6" pageNumber="107">Steyn 1884</emphasis>
(PRE), D =
<emphasis id="B92BEAC94621FFC8FD1ECA95FCBF50FB" box="[700,806,2019,2040]" italics="true" pageId="6" pageNumber="107">Steyn 1890</emphasis>
(PRE), E =
<emphasis id="B92BEAC94621FFC8FC3ECA95FB9F50FB" box="[924,1030,2019,2040]" italics="true" pageId="6" pageNumber="107">Steyn 1805</emphasis>
(PRE), F =
<emphasis id="B92BEAC94621FFC8FBD9CA95FA8750FB" box="[1147,1310,2019,2040]" italics="true" pageId="6" pageNumber="107">Koekemoer 4620</emphasis>
(PRE).
</paragraph>
</caption>
<caption id="DF2066534620FFC9FF2ACB82FBDF5052" ID-DOI="http://doi.org/10.5281/zenodo.13697368" ID-Zenodo-Dep="13697368" httpUri="https://zenodo.org/record/13697368/files/figure.png" pageId="7" pageNumber="108" startId="7.[136,229,1780,1801]" targetBox="[151,1436,175,1755]" targetPageId="7" targetType="figure">
<paragraph id="8BE036DB4620FFC9FF2ACB82FBDF5052" blockId="7.[136,1452,1780,1873]" pageId="7" pageNumber="108">
<emphasis id="B92BEAC94620FFC9FF2ACB82FF66500A" bold="true" box="[136,255,1780,1801]" pageId="7" pageNumber="108">FIGURE 4.</emphasis>
<taxonomicName id="4C5F4D584620FFC9FEA4CB82FE5A500A" authorityName="H. M. Steyn" authorityYear="2017" baseAuthorityName="H. M. Steyn" baseAuthorityYear="2017" box="[262,451,1780,1801]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="108" phylum="Tracheophyta" rank="species" species="glabra">
<emphasis id="B92BEAC94620FFC9FEA4CB82FE5A500A" box="[262,451,1780,1801]" italics="true" pageId="7" pageNumber="108">Acanthopsis glabra</emphasis>
</taxonomicName>
. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp; F. Bract. G. Calyx. H. Corolla (lobes lilac to cream; throat cream). I. Glandular hairs at base of style. J. Capsule. K. Seed. Vouchers: CF =
<emphasis id="B92BEAC94620FFC9FB8ACA6EFB36502E" box="[1064,1199,1816,1837]" italics="true" pageId="7" pageNumber="108">Acocks 16476</emphasis>
(PRE), GI =
<emphasis id="B92BEAC94620FFC9FAE2CA6EFA32502E" box="[1344,1451,1816,1837]" italics="true" pageId="7" pageNumber="108">Steyn 1893</emphasis>
(PRE), J &amp; K =
<emphasis id="B92BEAC94620FFC9FE81CA4AFE125052" box="[291,395,1852,1873]" italics="true" pageId="7" pageNumber="108">Steyn 2119</emphasis>
(PRE). Photographed by: H.M. Steyn (A, CK) and M. Koekemoer (B).
</paragraph>
</caption>
<paragraph id="8BE036DB4620FFC6FF1FCA0CFDCD5546" blockId="7.[136,1451,1914,2083]" lastBlockId="8.[136,1452,160,1193]" lastPageId="8" lastPageNumber="109" pageId="7" pageNumber="108">
<emphasis id="B92BEAC94620FFC9FF1FCA0CFDA65090" bold="true" box="[189,575,1914,1939]" pageId="7" pageNumber="108">Additional specimens examined:</emphasis>
<materialsCitation id="3B373C864620FFC9FDF9CA0CFC8750B4" collectingDate="1986-10-09" collectionCode="CD, NBG" collectorName="Le Roux &amp; Lloyd" country="South Africa" elevation="316260" location="Namaqualand. Farm Kourootjie" pageId="7" pageNumber="108" specimenCount="1" stateProvince="Northern Cape">
<collectingCountry id="F348764B4620FFC9FDF9CA0CFCBF5090" box="[603,806,1914,1939]" name="South Africa" pageId="7" pageNumber="108">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="499BF8394620FFC9FC92CA0CFC405090" box="[816,985,1914,1939]" country="South Africa" name="Northern Cape" pageId="7" pageNumber="108">Northern Cape</collectingRegion>
:
<location id="8E8060004620FFC9FC40CA0CFADD5090" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E8060004620FFC9FC40CA0CFADD5090" box="[994,1348,1914,1939]" country="South Africa" name="Namaqualand. Farm Kourootjie" pageId="7" pageNumber="108" stateProvince="Northern Cape">Namaqualand. Farm Kourootjie</location>
<quantity id="4CA79B3E4620FFC9FAE8CA0CFF3950B4" metricMagnitude="5" metricUnit="m" metricValue="3.1626" pageId="7" pageNumber="108" unit="m" value="316260.0">
<elevation id="0072D1E84620FFC9FAE8CA0CFF3950B4" metricMagnitude="5" metricUnit="m" metricValue="3.1626" pageId="7" pageNumber="108" unit="m" value="316260.0">316, 260 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E4620FFC9FF4FCAE8FE8F50B4" box="[237,278,1950,1975]" pageId="7" pageNumber="108">CD</collectionCode>
),
<date id="FFE1101B4620FFC9FE8ACAE8FE4C50B4" box="[296,469,1950,1975]" pageId="7" pageNumber="108" value="1986-10-09">
<collectingDate id="EFA5E9F34620FFC9FE8ACAE8FE4C50B4" box="[296,469,1950,1975]" pageId="7" pageNumber="108" value="1986-10-09">9 October 1986</collectingDate>
</date>
,
<emphasis id="B92BEAC94620FFC9FE7CCAE8FD5150B4" box="[478,712,1950,1975]" italics="true" pageId="7" pageNumber="108">
<collectorName id="26AA530D4620FFC9FE7CCAE8FDA050B4" box="[478,569,1950,1975]" pageId="7" pageNumber="108">Le Roux</collectorName>
&amp;
<collectorName id="26AA530D4620FFC9FDFBCAE8FD0050B4" box="[601,665,1950,1975]" pageId="7" pageNumber="108">Lloyd</collectorName>
608
</emphasis>
(
<collectionCode id="ED4EAE1E4620FFC9FD77CAE8FC8F50B4" box="[725,790,1950,1975]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="7" pageNumber="108" type="Herbarium">NBG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C864620FFC9FC85CAE8FC1550D8" collectingDate="2009-08-12" collectionCode="DD, PRE" collectorName="Steyn" country="South Africa" county="Namaqualand District" elevation="851" location="Messelpadpas" municipality="Springbok-Soebatsfontein" pageId="7" pageNumber="108" specimenCount="1">
<collectingCounty id="62814E574620FFC9FC85CAE8FB8650B4" box="[807,1055,1950,1975]" pageId="7" pageNumber="108">Namaqualand District</collectingCounty>
, on
<collectingMunicipality id="6B84ACA14620FFC9FBEBCAE8FAF450B4" box="[1097,1389,1950,1975]" pageId="7" pageNumber="108">Springbok-Soebatsfontein</collectingMunicipality>
road,
<location id="8E8060004620FFC9FF2ACAB4FEB150D8" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E8060004620FFC9FF2ACAB4FEB150D8" box="[136,296,1986,2011]" country="South Africa" county="Namaqualand District" municipality="Springbok-Soebatsfontein" name="Messelpadpas" pageId="7" pageNumber="108">Messelpadpas</location>
,
<quantity id="4CA79B3E4620FFC9FE90CAB4FEE350D8" box="[306,378,1986,2011]" metricMagnitude="2" metricUnit="m" metricValue="8.51" pageId="7" pageNumber="108" unit="m" value="851.0">
<elevation id="0072D1E84620FFC9FE90CAB4FEE350D8" box="[306,378,1986,2011]" metricMagnitude="2" metricUnit="m" metricValue="8.51" pageId="7" pageNumber="108" unit="m" value="851.0">851 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E4620FFC9FE6ACAB4FE6A50D8" box="[456,499,1986,2011]" country="India" lsid="urn:lsid:biocol.org:col:13033" name="Forest Research Institute, Indian Council of Forestry Research and Education" pageId="7" pageNumber="108" type="Herbarium">DD</collectionCode>
),
<date id="FFE1101B4620FFC9FDA7CAB4FD2150D8" box="[517,696,1986,2011]" pageId="7" pageNumber="108" value="2009-08-12">
<collectingDate id="EFA5E9F34620FFC9FDA7CAB4FD2150D8" box="[517,696,1986,2011]" pageId="7" pageNumber="108" value="2009-08-12">12 August 2009</collectingDate>
</date>
,
<emphasis id="B92BEAC94620FFC9FD60CAB4FCA450D8" box="[706,829,1986,2011]" italics="true" pageId="7" pageNumber="108">
<collectorName id="26AA530D4620FFC9FD60CAB4FD6650D8" box="[706,767,1986,2011]" pageId="7" pageNumber="108">Steyn</collectorName>
1565
</emphasis>
(
<collectionCode id="ED4EAE1E4620FFC9FCE9CAB4FC1D50D8" box="[843,900,1986,2011]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="7" pageNumber="108" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C864620FFC9FC34CAB4FDB450FC" collectingDate="1951-09-03" collectionCode="DA, NBG" country="South Africa" location="Groot Vlei" municipality="Namaqualand District" pageId="7" pageNumber="108" specimenCount="1">
<collectingMunicipality id="6B84ACA14620FFC9FC34CAB4FB1650D8" box="[918,1167,1986,2011]" pageId="7" pageNumber="108">Namaqualand District</collectingMunicipality>
,
<location id="8E8060004620FFC9FB3BCAB4FA8A50D8" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E8060004620FFC9FB3BCAB4FA8A50D8" box="[1177,1299,1986,2011]" country="South Africa" municipality="Namaqualand District" name="Groot Vlei" pageId="7" pageNumber="108">Groot Vlei</location>
, (2918
<collectionCode id="ED4EAE1E4620FFC9FAC2CAB4FA1250D8" box="[1376,1419,1986,2011]" pageId="7" pageNumber="108">DA</collectionCode>
),
<date id="FFE1101B4620FFC9FA3FCAB4FEA650FC" pageId="7" pageNumber="108" value="1951-09-03">
<collectingDate id="EFA5E9F34620FFC9FA3FCAB4FEA650FC" pageId="7" pageNumber="108" value="1951-09-03">3 September 1951</collectingDate>
</date>
,
<emphasis id="B92BEAC94620FFC9FEE8CA90FE4E50FC" box="[330,471,2022,2047]" italics="true" pageId="7" pageNumber="108">
<taxonomicName id="4C5F4D584620FFC9FEE8CA90FE3E50FC" box="[330,423,2022,2047]" genus="Johnson" kingdom="Plantae" pageId="7" pageNumber="108" phylum="Chlorophyta" rank="genus">Johnson</taxonomicName>
204
</emphasis>
(
<collectionCode id="ED4EAE1E4620FFC9FE46CA90FDBC50FC" box="[484,549,2022,2047]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="7" pageNumber="108" type="Herbarium">NBG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C864620FFC9FD95CA90FD4F50FC" box="[567,726,2022,2047]" country="South Africa" location="Namaqualand" pageId="7" pageNumber="108" specimenCount="1">
<location id="8E8060004620FFC9FD95CA90FD4F50FC" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E8060004620FFC9FD95CA90FD4F50FC" box="[567,726,2022,2047]" country="South Africa" name="Namaqualand" pageId="7" pageNumber="108">Namaqualand</location>
</materialsCitation>
;
<materialsCitation id="3B373C864620FFC9FD42CA90FCE250FC" box="[736,891,2022,2047]" country="South Africa" location="Kamieskroon" pageId="7" pageNumber="108" specimenCount="1">
<location id="8E8060004620FFC9FD42CA90FCE250FC" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E8060004620FFC9FD42CA90FCE250FC" box="[736,891,2022,2047]" country="South Africa" name="Kamieskroon" pageId="7" pageNumber="108">Kamieskroon</location>
</materialsCitation>
;
<materialsCitation id="3B373C864620FFC9FC26CA90FCA65F20" collectingDate="1952-09-25" collectionCode="BB, PRE" collectorName="Acocks" country="South Africa" elevation="762" location="Kamieskroon" pageId="7" pageNumber="108" specimenCount="1">
<quantity id="4CA79B3E4620FFC9FC26CA90FC4C50FC" box="[900,981,2022,2047]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="7" pageNumber="108" unit="mi" value="7.0">7 miles</quantity>
[
<quantity id="4CA79B3E4620FFC9FC40CA90FBB250FC" box="[994,1067,2022,2047]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="108" unit="km" value="11.0">11 km</quantity>
] N. by W. of
<location id="8E8060004620FFC9FB1BCA90FACA50FC" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E8060004620FFC9FB1BCA90FACA50FC" box="[1209,1363,2022,2047]" country="South Africa" name="Kamieskroon" pageId="7" pageNumber="108">Kamieskroon</location>
,
<quantity id="4CA79B3E4620FFC9FAFFCA90FA3250FC" box="[1373,1451,2022,2047]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="108" unit="ft" value="2500.0">
<elevation id="0072D1E84620FFC9FAFFCA90FA3250FC" box="[1373,1451,2022,2047]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="108" unit="ft" value="2500.0">2500 ft</elevation>
</quantity>
[
<quantity id="4CA79B3E4620FFC9FF2DC57CFF455F20" box="[143,220,2058,2083]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="108" unit="m" value="760.0">
<elevation id="0072D1E84620FFC9FF2DC57CFF455F20" box="[143,220,2058,2083]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="108" unit="m" value="760.0">760 m</elevation>
</quantity>
], (3017
<collectionCode id="ED4EAE1E4620FFC9FE91C57CFEC05F20" box="[307,345,2058,2083]" country="0" name="Buffalo Bill Museum" pageId="7" pageNumber="108">BB</collectionCode>
),
<date id="FFE1101B4620FFC9FECEC57CFDD15F20" box="[364,584,2058,2083]" pageId="7" pageNumber="108" value="1952-09-25">
<collectingDate id="EFA5E9F34620FFC9FECEC57CFDD15F20" box="[364,584,2058,2083]" pageId="7" pageNumber="108" value="1952-09-25">25 September 1952</collectingDate>
</date>
,
<emphasis id="B92BEAC94620FFC9FDF1C57CFD765F20" box="[595,751,2058,2083]" italics="true" pageId="7" pageNumber="108">
<collectorName id="26AA530D4620FFC9FDF1C57CFD3B5F20" box="[595,674,2058,2083]" pageId="7" pageNumber="108">Acocks</collectorName>
16476
</emphasis>
(
<collectionCode id="ED4EAE1E4620FFC9FD5FC57CFCAF5F20" box="[765,822,2058,2083]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="7" pageNumber="108" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C864620FFC9FCE8C57CFC705F20" box="[842,1001,2058,2083]" country="South Africa" location="Namaqualand" pageId="7" pageNumber="108" specimenCount="1">
<location id="8E8060004620FFC9FCE8C57CFC705F20" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E8060004620FFC9FCE8C57CFC705F20" box="[842,1001,2058,2083]" country="South Africa" name="Namaqualand" pageId="7" pageNumber="108">Namaqualand</location>
</materialsCitation>
;
<materialsCitation id="3B373C864620FFC6FC56C57CFE5157BA" collectingDate="1936-12-15" collectionCode="BB, PRE" collectorName="Adamson" country="South Africa" lastPageId="8" lastPageNumber="109" location="Kamieskroon" pageId="7" pageNumber="108" specimenCount="1">
<location id="8E8060004620FFC9FC56C57CFB175F20" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E8060004620FFC9FC56C57CFB175F20" box="[1012,1166,2058,2083]" country="South Africa" name="Kamieskroon" pageId="7" pageNumber="108">Kamieskroon</location>
, (3017
<collectionCode id="ED4EAE1E4620FFC9FB7EC57CFA9B5F20" box="[1244,1282,2058,2083]" country="0" name="Buffalo Bill Museum" pageId="7" pageNumber="108">BB</collectionCode>
),
<date id="FFE1101B4620FFC6FAB4C57CFF5A57BA" lastPageId="8" lastPageNumber="109" pageId="7" pageNumber="108" value="1936-12-15">
<collectingDate id="EFA5E9F34620FFC6FAB4C57CFF5A57BA" lastPageId="8" lastPageNumber="109" pageId="7" pageNumber="108" value="1936-12-15">15 December 1936</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FF6DCDD6FEEE57BA" box="[207,375,160,185]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FF6DCDD6FEAE57BA" box="[207,311,160,185]" pageId="8" pageNumber="109">Adamson</collectorName>
1422
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FE25CDD6FE5957BA" box="[391,448,160,185]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="8" pageNumber="109" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462FFFC6FE77CDD6FCB257DE" collectingDate="2010-12-29" collectionCode="BB, PRE" collectorName="Koekemoer" country="South Africa" elevation="689" location="Namaqua National Park. On" pageId="8" pageNumber="109" specimenCount="1">
<location id="8E806000462FFFC6FE77CDD6FC8D57BA" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E806000462FFFC6FE77CDD6FC8D57BA" box="[469,788,160,185]" country="South Africa" name="Namaqua National Park. On" pageId="8" pageNumber="109">Namaqua National Park. On</location>
the cement road
<locationDeviation id="9A522FF9462FFFC6FC7BCDD6FAEC57BA" box="[985,1397,160,185]" firstLocation="Skilpad" pageId="8" pageNumber="109" secondLocation="Soebatsfontein">between Skilpad and Soebatsfontein</locationDeviation>
,
<quantity id="4CA79B3E462FFFC6FA23CDD6FF3957DE" metricMagnitude="2" metricUnit="m" metricValue="6.89" pageId="8" pageNumber="109" unit="m" value="689.0">
<elevation id="0072D1E8462FFFC6FA23CDD6FF3957DE" metricMagnitude="2" metricUnit="m" metricValue="6.89" pageId="8" pageNumber="109" unit="m" value="689.0">689 m</elevation>
</quantity>
, (3017
<collectionCode id="ED4EAE1E462FFFC6FF50CDB2FE8157DE" box="[242,280,196,221]" country="0" name="Buffalo Bill Museum" pageId="8" pageNumber="109">BB</collectionCode>
),
<date id="FFE1101B462FFFC6FE8CCDB2FD9357DE" box="[302,522,196,221]" pageId="8" pageNumber="109" value="2010-12-29">
<collectingDate id="EFA5E9F3462FFFC6FE8CCDB2FD9357DE" box="[302,522,196,221]" pageId="8" pageNumber="109" value="2010-12-29">29 December 2010</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FDBACDB2FD4157DE" box="[536,728,196,221]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FDBACDB2FD0157DE" box="[536,664,196,221]" pageId="8" pageNumber="109">Koekemoer</collectorName>
3996
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FD4BCDB2FCBB57DE" box="[745,802,196,221]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="8" pageNumber="109" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462FFFC6FC9ACDB2FBEC5602" collectingDate="2011-08-11" collectionCode="BB, PRE" collectorName="Steyn" country="South Africa" elevation="100" location="Namaqualand District. Namaqua National Park" pageId="8" pageNumber="109" specimenCount="1">
<location id="8E806000462FFFC6FC9ACDB2FACC57DE" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E806000462FFFC6FC9ACDB2FACC57DE" box="[824,1365,196,221]" country="South Africa" name="Namaqualand District. Namaqua National Park" pageId="8" pageNumber="109">Namaqualand District. Namaqua National Park</location>
,
<quantity id="4CA79B3E462FFFC6FAC1CDB2FA3557DE" box="[1379,1452,196,221]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="109" unit="m" value="100.0">
<elevation id="0072D1E8462FFFC6FAC1CDB2FA3557DE" box="[1379,1452,196,221]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="109" unit="m" value="100.0">100 m</elevation>
</quantity>
S of view point on circular route,
<quantity id="4CA79B3E462FFFC6FDAFCD9EFDCE5602" box="[525,599,232,257]" metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" pageId="8" pageNumber="109" unit="m" value="691.0">
<elevation id="0072D1E8462FFFC6FDAFCD9EFDCE5602" box="[525,599,232,257]" metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" pageId="8" pageNumber="109" unit="m" value="691.0">691 m</elevation>
</quantity>
, (3017
<collectionCode id="ED4EAE1E462FFFC6FD0BCD9EFD565602" box="[681,719,232,257]" country="0" name="Buffalo Bill Museum" pageId="8" pageNumber="109">BB</collectionCode>
),
<date id="FFE1101B462FFFC6FD46CD9EFC005602" box="[740,921,232,257]" pageId="8" pageNumber="109" value="2011-08-11">
<collectingDate id="EFA5E9F3462FFFC6FD46CD9EFC005602" box="[740,921,232,257]" pageId="8" pageNumber="109" value="2011-08-11">11 August 2011</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FC04CD9EFBBA5602" box="[934,1059,232,257]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FC04CD9EFC7A5602" box="[934,995,232,257]" pageId="8" pageNumber="109">Steyn</collectorName>
1860
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FB91CD9EFBF55602" box="[1075,1132,232,257]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="8" pageNumber="109" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462FFFC6FB20CD9EFBE45626" collectingDate="2010-09-02" collectionCode="BB, PRE" collectorName="Steyn" country="South Africa" elevation="765" location="Namaqualand District. No Heep" pageId="8" pageNumber="109" specimenCount="1">
<location id="8E806000462FFFC6FB20CD9EFF5A5626" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E806000462FFFC6FB20CD9EFF5A5626" country="South Africa" name="Namaqualand District. No Heep" pageId="8" pageNumber="109">Namaqualand District. No Heep</location>
road
<locationDeviation id="9A522FF9462FFFC6FEAACC7AFE785626" box="[264,481,268,293]" location="Kamieskroon" pageId="8" pageNumber="109">N of Kamieskroon</locationDeviation>
,
<quantity id="4CA79B3E462FFFC6FE4DCC7AFDA25626" box="[495,571,268,293]" metricMagnitude="2" metricUnit="m" metricValue="7.65" pageId="8" pageNumber="109" unit="m" value="765.0">
<elevation id="0072D1E8462FFFC6FE4DCC7AFDA25626" box="[495,571,268,293]" metricMagnitude="2" metricUnit="m" metricValue="7.65" pageId="8" pageNumber="109" unit="m" value="765.0">765 m</elevation>
</quantity>
, (3017
<collectionCode id="ED4EAE1E462FFFC6FD2FCC7AFD2A5626" box="[653,691,268,293]" country="0" name="Buffalo Bill Museum" pageId="8" pageNumber="109">BB</collectionCode>
),
<date id="FFE1101B462FFFC6FD68CC7AFC075626" box="[714,926,268,293]" pageId="8" pageNumber="109" value="2010-09-02">
<collectingDate id="EFA5E9F3462FFFC6FD68CC7AFC075626" box="[714,926,268,293]" pageId="8" pageNumber="109" value="2010-09-02">2 September 2010</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FC0ECC7AFBB35626" box="[940,1066,268,293]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FC0ECC7AFC705626" box="[940,1001,268,293]" pageId="8" pageNumber="109">Steyn</collectorName>
1767
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FB9ECC7AFBEC5626" box="[1084,1141,268,293]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="8" pageNumber="109" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462FFFC6FB29CC7AFC4E564A" collectingDate="2001-09-10" collectionCode="BB, KMG, PRE" collectorName="Venter JV" country="South Africa" elevation="750" location="Kamieskroon. Bloupoort" pageId="8" pageNumber="109" specimenCount="1">
<location id="8E806000462FFFC6FB29CC7AFA3E5626" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E806000462FFFC6FB29CC7AFA3E5626" box="[1163,1447,268,293]" country="South Africa" name="Kamieskroon. Bloupoort" pageId="8" pageNumber="109">Kamieskroon. Bloupoort</location>
,
<quantity id="4CA79B3E462FFFC6FF2ACC46FF48564A" box="[136,209,304,329]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="109" unit="m" value="750.0">
<elevation id="0072D1E8462FFFC6FF2ACC46FF48564A" box="[136,209,304,329]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="109" unit="m" value="750.0">750 m</elevation>
</quantity>
, (3017
<collectionCode id="ED4EAE1E462FFFC6FE82CC46FEDF564A" box="[288,326,304,329]" country="0" name="Buffalo Bill Museum" pageId="8" pageNumber="109">BB</collectionCode>
),
<date id="FFE1101B462FFFC6FEF8CC46FDAC564A" box="[346,565,304,329]" pageId="8" pageNumber="109" value="2001-09-10">
<collectingDate id="EFA5E9F3462FFFC6FEF8CC46FDAC564A" box="[346,565,304,329]" pageId="8" pageNumber="109" value="2001-09-10">10 September 2001</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FDE2CC46FCA1564A" box="[576,824,304,329]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FDE2CC46FD36564A" box="[576,687,304,329]" pageId="8" pageNumber="109">Venter JV</collectorName>
9599(JV-52)
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FCE4CC46FC15564A" box="[838,908,304,329]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13561" lsid="urn:lsid:biocol.org:col:13561" name="McGregor Museum" pageId="8" pageNumber="109" type="Herbarium">KMG</collectionCode>
,
<collectionCode id="ED4EAE1E462FFFC6FC34CC46FC57564A" box="[918,974,304,329]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="8" pageNumber="109" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462FFFC6FC40CC46FC1C566E" collectingDate="1910-12-07" collectionCode="DD, BOL" collectorName="Pillans" country="South Africa" location="Garies" municipality="Namaqualand. In valley" pageId="8" pageNumber="109" specimenCount="1">
<collectingMunicipality id="6B84ACA1462FFFC6FC40CC46FB75564A" box="[994,1260,304,329]" pageId="8" pageNumber="109">Namaqualand. In valley</collectingMunicipality>
<quantity id="4CA79B3E462FFFC6FB51CC46FADC564A" box="[1267,1349,304,329]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="8" pageNumber="109" unit="mi" value="2.0">2 miles</quantity>
[
<quantity id="4CA79B3E462FFFC6FAF0CC46FA3C564A" box="[1362,1445,304,329]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="109" unit="km" value="3.2">3.2 km</quantity>
] west of
<location id="8E806000462FFFC6FF47CC22FEA8566E" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E806000462FFFC6FF47CC22FEA8566E" box="[229,305,340,365]" country="South Africa" municipality="Namaqualand. In valley" name="Garies" pageId="8" pageNumber="109">Garies</location>
, (3017
<collectionCode id="ED4EAE1E462FFFC6FE23CC22FE35566E" box="[385,428,340,365]" country="India" lsid="urn:lsid:biocol.org:col:13033" name="Forest Research Institute, Indian Council of Forestry Research and Education" pageId="8" pageNumber="109" type="Herbarium">DD</collectionCode>
),
<date id="FFE1101B462FFFC6FE60CC22FD09566E" box="[450,656,340,365]" pageId="8" pageNumber="109" value="1910-12-07">
<collectingDate id="EFA5E9F3462FFFC6FE60CC22FD09566E" box="[450,656,340,365]" pageId="8" pageNumber="109" value="1910-12-07">7 December 1910</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FD3CCC22FCB7566E" box="[670,814,340,365]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FD3CCC22FD74566E" box="[670,749,340,365]" pageId="8" pageNumber="109">Pillans</collectorName>
5640
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FCE2CC22FCE7566E" box="[832,894,340,365]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15865" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="8" pageNumber="109" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462FFFC6FC31CC22FD5C5692" collectingDate="2015-10-04" collectionCode="AA, PRE" collectorName="Steyn" country="South Africa" elevation="1043" location="Namaqualand District." pageId="8" pageNumber="109" specimenCount="1">
<location id="8E806000462FFFC6FC31CC22FB0D566E" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E806000462FFFC6FC31CC22FB0D566E" box="[915,1172,340,365]" country="South Africa" name="Namaqualand District." pageId="8" pageNumber="109">Namaqualand District.</location>
<locationDeviation id="9A522FF9462FFFC6FB3ECC22FAFF566E" box="[1180,1382,340,365]" location="Leliefontein" pageId="8" pageNumber="109">N of Leliefontein</locationDeviation>
,
<quantity id="4CA79B3E462FFFC6FAD1CC22FF395692" metricMagnitude="3" metricUnit="m" metricValue="1.043" pageId="8" pageNumber="109" unit="m" value="1043.0">
<elevation id="0072D1E8462FFFC6FAD1CC22FF395692" metricMagnitude="3" metricUnit="m" metricValue="1.043" pageId="8" pageNumber="109" unit="m" value="1043.0">1043 m</elevation>
</quantity>
, (3018
<collectionCode id="ED4EAE1E462FFFC6FF53CC0EFE855692" box="[241,284,376,401]" country="Kazakhstan" lsid="urn:lsid:biocol.org:col:13590" name="Ministry of Science, Academy of Sciences" pageId="8" pageNumber="109" type="Herbarium">AA</collectionCode>
),
<date id="FFE1101B462FFFC6FE90CC0EFE715692" box="[306,488,376,401]" pageId="8" pageNumber="109" value="2015-10-04">
<collectingDate id="EFA5E9F3462FFFC6FE90CC0EFE715692" box="[306,488,376,401]" pageId="8" pageNumber="109" value="2015-10-04">4 October 2015</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FE54CC0EFDEB5692" box="[502,626,376,401]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FE54CC0EFDAA5692" box="[502,563,376,401]" pageId="8" pageNumber="109">Steyn</collectorName>
2119
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FD21CC0EFD255692" box="[643,700,376,401]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="8" pageNumber="109" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462FFFC6FD70CC0EFA3E56B6" collectingDate="1911-01-17" collectionCode="AC, BOL" collectorName="Pearson" country="South Africa" county="Khamiesberg. Sandy" elevation="1524" location="Leliefontein Mission Station" municipality="Beacon" pageId="8" pageNumber="109" specimenCount="1">
<collectingCounty id="62814E57462FFFC6FD70CC0EFC275692" box="[722,958,376,401]" pageId="8" pageNumber="109">Khamiesberg. Sandy</collectingCounty>
ground at foot of
<collectingMunicipality id="6B84ACA1462FFFC6FB37CC0EFB705692" box="[1173,1257,376,401]" pageId="8" pageNumber="109">Beacon</collectingMunicipality>
hill
<quantity id="4CA79B3E462FFFC6FA83CC0EFAEF5692" box="[1313,1398,376,401]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="8" pageNumber="109" unit="mi" value="2.0">2 miles</quantity>
[
<quantity id="4CA79B3E462FFFC6FA24CC0EFF3656B6" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="109" unit="km" value="3.2">3.2 km</quantity>
] south east of
<location id="8E806000462FFFC6FEF0CCEAFD0C56B6" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E806000462FFFC6FEF0CCEAFD0C56B6" box="[338,661,412,437]" country="South Africa" county="Khamiesberg. Sandy" municipality="Beacon" name="Leliefontein Mission Station" pageId="8" pageNumber="109">Leliefontein Mission Station</location>
,
<quantity id="4CA79B3E462FFFC6FD03CCEAFD6856B6" box="[673,753,412,437]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="109" unit="ft" value="5000.0">
<elevation id="0072D1E8462FFFC6FD03CCEAFD6856B6" box="[673,753,412,437]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="109" unit="ft" value="5000.0">5000 ft</elevation>
</quantity>
[
<quantity id="4CA79B3E462FFFC6FD5DCCEAFCC256B6" box="[767,859,412,437]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="109" unit="m" value="1500.0">
<elevation id="0072D1E8462FFFC6FD5DCCEAFCC256B6" box="[767,859,412,437]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="109" unit="m" value="1500.0">1500 m</elevation>
</quantity>
], (3018
<collectionCode id="ED4EAE1E462FFFC6FC13CCEAFC4256B6" box="[945,987,412,437]" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="8" pageNumber="109" type="University or college">AC</collectionCode>
),
<date id="FFE1101B462FFFC6FC4CCCEAFB3056B6" box="[1006,1193,412,437]" pageId="8" pageNumber="109" value="1911-01-17">
<collectingDate id="EFA5E9F3462FFFC6FC4CCCEAFB3056B6" box="[1006,1193,412,437]" pageId="8" pageNumber="109" value="1911-01-17">17 January 1911</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FB17CCEAFACB56B6" box="[1205,1362,412,437]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FB17CCEAFA8B56B6" box="[1205,1298,412,437]" pageId="8" pageNumber="109">Pearson</collectorName>
6378
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FAC3CCEAFA0656B6" box="[1377,1439,412,437]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15865" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="8" pageNumber="109" type="Herbarium">BOL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462FFFC6FF2ACCB6FF4C56DA" box="[136,213,448,473]" country="South Africa" location="Garies" pageId="8" pageNumber="109" specimenCount="1">
<location id="8E806000462FFFC6FF2ACCB6FF4C56DA" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E806000462FFFC6FF2ACCB6FF4C56DA" box="[136,213,448,473]" country="South Africa" name="Garies" pageId="8" pageNumber="109">Garies</location>
</materialsCitation>
;
<materialsCitation id="3B373C86462FFFC6FF41CCB6FF485522" collectingDate="2009-08-22" collectingDateMax="2012-08-27" collectingDateMin="2009-08-22" collectionCode="CA, PRE, CD" collectorName="Steyn" country="South Africa" elevation="491" location="Kwanous" municipality="Namaqualand District" pageId="8" pageNumber="109" specimenCount="1" stateProvince="Western Cape">
on track
<locationDeviation id="9A522FF9462FFFC6FEE9CCB6FC8856DA" box="[331,785,448,473]" firstLocation="Buffelsfontein" pageId="8" pageNumber="109" secondLocation="Doornkraal">between Buffelsfontein and Doornkraal</locationDeviation>
,
<quantity id="4CA79B3E462FFFC6FC82CCB6FCF556DA" box="[800,876,448,473]" metricMagnitude="2" metricUnit="m" metricValue="4.91" pageId="8" pageNumber="109" unit="m" value="491.0">
<elevation id="0072D1E8462FFFC6FC82CCB6FCF556DA" box="[800,876,448,473]" metricMagnitude="2" metricUnit="m" metricValue="4.91" pageId="8" pageNumber="109" unit="m" value="491.0">491 m</elevation>
</quantity>
, (3018
<collectionCode id="ED4EAE1E462FFFC6FC1FCCB6FC7E56DA" box="[957,999,448,473]" country="USA" name="Chicago Academy of Sciences" pageId="8" pageNumber="109" type="Museum">CA</collectionCode>
),
<date id="FFE1101B462FFFC6FC5FCCB6FB2E56DA" box="[1021,1207,448,473]" pageId="8" pageNumber="109" value="2009-08-22">
<collectingDate id="EFA5E9F3462FFFC6FC5FCCB6FB2E56DA" box="[1021,1207,448,473]" pageId="8" pageNumber="109" value="2009-08-22">22 August 2009</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FB67CCB6FACC56DA" box="[1221,1365,448,473]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FB67CCB6FA9B56DA" box="[1221,1282,448,473]" pageId="8" pageNumber="109">Steyn</collectorName>
1681B
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FAC4CCB6FA0656DA" box="[1382,1439,448,473]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="8" pageNumber="109" type="Herbarium">PRE</collectionCode>
).
<collectingRegion id="499BF839462FFFC6FF2ACC92FEB156FE" box="[136,296,484,509]" country="South Africa" name="Western Cape" pageId="8" pageNumber="109">Western Cape</collectingRegion>
:
<collectingMunicipality id="6B84ACA1462FFFC6FE93CC92FDB256FE" box="[305,555,484,509]" pageId="8" pageNumber="109">Namaqualand District</collectingMunicipality>
. Farm Eenkokerboom/
<location id="8E806000462FFFC6FC8CCC92FC0F56FE" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E806000462FFFC6FC8CCC92FC0F56FE" box="[814,918,484,509]" country="South Africa" municipality="Namaqualand District" name="Kwanous" pageId="8" pageNumber="109" stateProvince="Western Cape">Kwanous</location>
,
<quantity id="4CA79B3E462FFFC6FC02CC92FC7056FE" box="[928,1001,484,509]" metricMagnitude="2" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="109" unit="m" value="320.0">
<elevation id="0072D1E8462FFFC6FC02CC92FC7056FE" box="[928,1001,484,509]" metricMagnitude="2" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="109" unit="m" value="320.0">320 m</elevation>
</quantity>
, (3018
<collectionCode id="ED4EAE1E462FFFC6FB95CC92FBF956FE" box="[1079,1120,484,509]" pageId="8" pageNumber="109">CD</collectionCode>
),
<date id="FFE1101B462FFFC6FBD1CC92FABF56FE" box="[1139,1318,484,509]" pageId="8" pageNumber="109" value="2012-08-27">
<collectingDate id="EFA5E9F3462FFFC6FBD1CC92FABF56FE" box="[1139,1318,484,509]" pageId="8" pageNumber="109" value="2012-08-27">27 August 2012</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FA92CC92FA3256FE" box="[1328,1451,484,509]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FA92CC92FAF456FE" box="[1328,1389,484,509]" pageId="8" pageNumber="109">Steyn</collectorName>
1893
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FF32CF7EFF505522" box="[144,201,520,545]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="8" pageNumber="109" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462FFFC6FF7CCF7EFDC95546" collectingDate="1908-12-21" collectionCode="DA, BOL, P, S" collectorName="Pearson &amp; Without" country="South Africa" county="Bushmanland" location="Drege" municipality="Upper. Near Nieuwfontein" pageId="8" pageNumber="109" specimenCount="1">
<collectingCounty id="62814E57462FFFC6FF7CCF7EFEE15522" box="[222,376,520,545]" pageId="8" pageNumber="109">Bushmanland</collectingCounty>
and
<collectingMunicipality id="6B84ACA1462FFFC6FE12CF7EFD465522" box="[432,735,520,545]" pageId="8" pageNumber="109">Upper. Near Nieuwfontein</collectingMunicipality>
, (3018
<collectionCode id="ED4EAE1E462FFFC6FC8FCF7EFCC15522" box="[813,856,520,545]" pageId="8" pageNumber="109">DA</collectionCode>
),
<date id="FFE1101B462FFFC6FCCFCF7EFBDE5522" box="[877,1095,520,545]" pageId="8" pageNumber="109" value="1908-12-21">
<collectingDate id="EFA5E9F3462FFFC6FCCFCF7EFBDE5522" box="[877,1095,520,545]" pageId="8" pageNumber="109" value="1908-12-21">21 December 1908</collectingDate>
</date>
,
<emphasis id="B92BEAC9462FFFC6FBF0CF7EFB695522" box="[1106,1264,520,545]" italics="true" pageId="8" pageNumber="109">
<collectorName id="26AA530D462FFFC6FBF0CF7EFB365522" box="[1106,1199,520,545]" pageId="8" pageNumber="109">Pearson</collectorName>
3317
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FAA2CF7EFAA75522" box="[1280,1342,520,545]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15865" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="8" pageNumber="109" type="Herbarium">BOL</collectionCode>
).
<collectorName id="26AA530D462FFFC6FAF3CF7EFA355522" box="[1361,1452,520,545]" pageId="8" pageNumber="109">Without</collectorName>
precise locality:
<emphasis id="B92BEAC9462FFFC6FEE3CF5AFE5F5546" box="[321,454,556,581]" italics="true" pageId="8" pageNumber="109">
<location id="8E806000462FFFC6FEE3CF5AFE1E5546" LSID="urn:lsid:plazi:treatment:03F687CD4624FFC6FF2AC9DAFEE753AA:8E806000462FFFC6FEE3CF5AFE1E5546" box="[321,391,556,581]" country="South Africa" county="Bushmanland" municipality="Upper. Near Nieuwfontein" name="Drege" pageId="8" pageNumber="109">Drège</location>
2436
</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FE74CF5AFE715546" box="[470,488,556,581]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="8" pageNumber="109" type="Herbarium">P</collectionCode>
),
<emphasis id="B92BEAC9462FFFC6FE5FCF5AFDBD5546" box="[509,548,556,581]" italics="true" pageId="8" pageNumber="109">s.n.</emphasis>
(
<collectionCode id="ED4EAE1E462FFFC6FD97CF5AFDDF5546" box="[565,582,556,581]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="8" pageNumber="109" type="Herbarium">S</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE036DB462FFFC6FF1FCF26FB28558E" blockId="8.[136,1452,160,1193]" pageId="8" pageNumber="109">
<emphasis id="B92BEAC9462FFFC6FF1FCF26FE90556A" bold="true" box="[189,265,592,617]" pageId="8" pageNumber="109">Notes:</emphasis>
<emphasis id="B92BEAC9462FFFC6FE84CF26FE7F556A" box="[294,486,592,617]" italics="true" pageId="8" pageNumber="109">Koekemoer 3996</emphasis>
has villose leaves and antrorse hairs on the bracts but is here included in
<taxonomicName id="4C5F4D58462FFFC6FA9DCF26FA35556A" authorityName="H. M. Steyn" authorityYear="2017" baseAuthorityName="H. M. Steyn" baseAuthorityYear="2017" box="[1343,1452,592,617]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="glabra">
<emphasis id="B92BEAC9462FFFC6FA9DCF26FA35556A" box="[1343,1452,592,617]" italics="true" pageId="8" pageNumber="109">A. glabra</emphasis>
</taxonomicName>
rather than in
<taxonomicName id="4C5F4D58462FFFC6FE86CF02FE15558E" authorityName="Obermeyer" authorityYear="1937" baseAuthorityName="S. Moore" box="[292,396,628,653]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC9462FFFC6FE86CF02FE15558E" box="[292,396,628,653]" italics="true" pageId="8" pageNumber="109">A. scullyi</emphasis>
</taxonomicName>
due to the oblong, truncate bracts ending in five simple primary spines.
</paragraph>
<paragraph id="8BE036DB462FFFC6FF1FCFEEFBD554F6" blockId="8.[136,1452,160,1193]" pageId="8" pageNumber="109">
<taxonomicName id="4C5F4D58462FFFC6FF68CFEEFE1855B2" box="[202,385,664,689]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthus" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="glaber">
<emphasis id="B92BEAC9462FFFC6FF68CFEEFE1855B2" box="[202,385,664,689]" italics="true" pageId="8" pageNumber="109">Acanthus glaber</emphasis>
</taxonomicName>
” was proposed by Meyer in
<bibRefCitation id="EFCE4B2A462FFFC6FD1FCFEEFCC855B2" author="Drege, J. F." box="[701,849,664,689]" pageId="8" pageNumber="109" refId="ref12531" refString="Drege, J. F. (1843) Zwei Pflanzengeographische Documente, nebst einer Einleitung von Dr. E. Meyer. Regensburg, Leipzig, 230 pp. https: // doi. org / 10.5962 / bhl. title. 87612" type="book" year="1843">Drège (1843)</bibRefCitation>
based on a herbarium specimen in Drèges herbarium, but it was not validly published because no diagnosis or description was provided.
<bibRefCitation id="EFCE4B2A462FFFC6FB90CFCAFAF855D6" author="Nees von Esenbeck, C. G. D." box="[1074,1377,700,725]" pageId="8" pageNumber="109" pagination="46 - 519" refId="ref13072" refString="Nees von Esenbeck, C. G. D. (1847) Acanthaceae. In: Candolle, A. de. (Ed.) Prodromus systematis naturalis regni vegetabilis 11. Treuttel &amp; Wurtz, Paris, pp. 46 - 519." type="book chapter" year="1847">Nees von Esenbeck (1847)</bibRefCitation>
firstly validly published the name
<taxonomicName id="4C5F4D58462FFFC6FE65CF96FD5D55FA" authorityName="Nees von Esenbeck" authorityYear="1847" box="[455,708,736,761]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthodium" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="glabrum">
<emphasis id="B92BEAC9462FFFC6FE65CF96FD5D55FA" box="[455,708,736,761]" italics="true" pageId="8" pageNumber="109">Acanthodium glabrum</emphasis>
</taxonomicName>
for the taxon with citing
<taxonomicName id="4C5F4D58462FFFC6FC4CCF96FB3055FA" box="[1006,1193,736,761]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthus" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="glaber">
<emphasis id="B92BEAC9462FFFC6FC4CCF96FB3055FA" box="[1006,1193,736,761]" italics="true" pageId="8" pageNumber="109">Acanthus glaber</emphasis>
</taxonomicName>
as a synonym.
<bibRefCitation id="EFCE4B2A462FFFC6FAFDCF96FF4A541E" author="Schinz, H." pageId="8" pageNumber="109" pagination="179 - 230" refId="ref13310" refString="Schinz, H. (1890) Beitrage zur Kenntnis der Flora von Deutsch-Sudwest-Afrika und der angrenzenden Gebiete. Verhandlungen des Botanischen Vereins fur die Provinz Brandenbrug und die Angrenzenden Lander 31: 179 - 230." type="journal article" year="1890">Schinz (1890)</bibRefCitation>
treated it as a variety of
<taxonomicName id="4C5F4D58462FFFC6FE55CE72FC9D541E" authorityName=", Drege" authorityYear="1843" box="[503,772,772,797]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="carduifolia">
<emphasis id="B92BEAC9462FFFC6FE55CE72FC9D541E" box="[503,772,772,797]" italics="true" pageId="8" pageNumber="109">Acanthopsis carduifolia</emphasis>
</taxonomicName>
(L.f.)
<bibRefCitation id="EFCE4B2A462FFFC6FCF0CE72FBB4541E" author="Schinz, H." box="[850,1069,772,797]" pageId="8" pageNumber="109" pagination="179 - 230" refId="ref13310" refString="Schinz, H. (1890) Beitrage zur Kenntnis der Flora von Deutsch-Sudwest-Afrika und der angrenzenden Gebiete. Verhandlungen des Botanischen Vereins fur die Provinz Brandenbrug und die Angrenzenden Lander 31: 179 - 230." type="journal article" year="1890">Schinz (1890: 200)</bibRefCitation>
and the authorship of the
<taxonomicName id="4C5F4D58462FFFC6FAFCCE72FF5C5442" pageId="8" pageNumber="109" rank="variety" variety="name">variety name</taxonomicName>
was incorrectly attributed to “(E.Mey.) Schinz”.The correct author citation of the variety
<taxonomicName id="4C5F4D58462FFFC6FB03CE5EFE9E5466" authorityName="Schinz" authorityYear="1890" baseAuthorityName="Nees" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="variety" species="carduifolia" variety="glabra">
<emphasis id="B92BEAC9462FFFC6FB03CE5EFA325442" box="[1185,1451,808,833]" italics="true" pageId="8" pageNumber="109">Acanthopsis carduifolia</emphasis>
var.
<emphasis id="B92BEAC9462FFFC6FF1ECE3AFE9E5466" box="[188,263,844,869]" italics="true" pageId="8" pageNumber="109">glabra</emphasis>
</taxonomicName>
is therefore (Nees) Schinz. Simultaneously, Schinz published an additional variety,
<taxonomicName id="4C5F4D58462FFFC6FB78CE3AFE6F548A" authority="Schinz (1890: 200)" authorityName="Schinz" authorityPageNumber="200" authorityYear="1890" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="variety" species="carduifolia" variety="longearistata">
<emphasis id="B92BEAC9462FFFC6FB78CE3AFAEE5466" box="[1242,1399,844,869]" italics="true" pageId="8" pageNumber="109">A. carduifolia</emphasis>
var.
<emphasis id="B92BEAC9462FFFC6FF2ACE06FE84548A" box="[136,285,880,905]" italics="true" pageId="8" pageNumber="109">longearistata</emphasis>
<bibRefCitation id="EFCE4B2A462FFFC6FE81CE06FE6F548A" author="Schinz, H." box="[291,502,880,905]" pageId="8" pageNumber="109" pagination="179 - 230" refId="ref13310" refString="Schinz, H. (1890) Beitrage zur Kenntnis der Flora von Deutsch-Sudwest-Afrika und der angrenzenden Gebiete. Verhandlungen des Botanischen Vereins fur die Provinz Brandenbrug und die Angrenzenden Lander 31: 179 - 230." type="journal article" year="1890">Schinz (1890: 200)</bibRefCitation>
</taxonomicName>
based on a specimen (
<emphasis id="B92BEAC9462FFFC6FD49CE06FCEF548A" box="[747,886,880,905]" italics="true" pageId="8" pageNumber="109">Schenck 318</emphasis>
) collected at “Aus in Gross-Namaland” [
<collectingCountry id="F348764B462FFFC6FA9ACE06FA3B548A" box="[1336,1442,880,905]" name="Namibia" pageId="8" pageNumber="109">Namibia</collectingCountry>
]. This specimen (
<emphasis id="B92BEAC9462FFFC6FE9CCEE2FE5454AE" box="[318,461,916,941]" italics="true" pageId="8" pageNumber="109">Schenck 318</emphasis>
) could not be traced and verified and therefore this name is excluded from the current treatment. In his treatment of the genus for Flora Capensis,
<bibRefCitation id="EFCE4B2A462FFFC6FC87CECEFC5854D2" author="Clarke, C. B." box="[805,961,952,977]" pageId="8" pageNumber="109" pagination="1 - 92" refId="ref12444" refString="Clarke, C. B. (1901) Acanthaceae. In: Thiselton-Dyer, W. T. (Ed.) Flora capensis 5, 1. Lovell Reeve &amp; Co., London, pp. 1 - 92. https: // doi. org / 10.5962 / bhl. title. 821" type="book chapter" year="1901">Clarke (1901)</bibRefCitation>
upheld
<taxonomicName id="4C5F4D58462FFFC6FBBECECEFB0E54D2" authorityName="Schinz" authorityYear="1890" baseAuthorityName="Nees" box="[1052,1175,952,977]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="variety" species="carduifolia" variety="glabra">
var.
<emphasis id="B92BEAC9462FFFC6FBEECECEFB0E54D2" box="[1100,1175,952,977]" italics="true" pageId="8" pageNumber="109">glabra</emphasis>
</taxonomicName>
but did not mention
<taxonomicName id="4C5F4D58462FFFC6FA20CECEFE8454F6" authorityName="Schinz" authorityYear="1890" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="variety" species="carduifolia" variety="longearistata">
var.
<emphasis id="B92BEAC9462FFFC6FF2ACEAAFE8454F6" box="[136,285,988,1013]" italics="true" pageId="8" pageNumber="109">longearistata</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFCE4B2A462FFFC6FE89CEAAFE5F54F6" author="Meyer, P. G." box="[299,454,988,1013]" pageId="8" pageNumber="109" refId="ref12974" refString="Meyer, P. G. (1961) Die Acanthaceae und Euphorbiaceae Sudwestafrikas. Publisher unknown, Munchen, 151 pp." type="book" year="1961">Meyer (1961)</bibRefCitation>
sunk
<taxonomicName id="4C5F4D58462FFFC6FDABCEAAFCE954F6" authorityName="Schinz" authorityYear="1890" box="[521,880,988,1013]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="variety" species="carduifolia" variety="longearistata">
<emphasis id="B92BEAC9462FFFC6FDABCEAAFD3A54F6" box="[521,675,988,1013]" italics="true" pageId="8" pageNumber="109">A. carduifolia</emphasis>
var.
<emphasis id="B92BEAC9462FFFC6FD79CEAAFCE954F6" box="[731,880,988,1013]" italics="true" pageId="8" pageNumber="109">longearistata</emphasis>
</taxonomicName>
under
<taxonomicName id="4C5F4D58462FFFC6FC1CCEAAFBDC54F6" authorityName="Nees von Esenbeck" authorityYear="1847" box="[958,1093,988,1013]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="disperma">
<emphasis id="B92BEAC9462FFFC6FC1CCEAAFBDC54F6" box="[958,1093,988,1013]" italics="true" pageId="8" pageNumber="109">A. disperma</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE036DB462FFFC6FF1FC976FEE753AA" blockId="8.[136,1452,160,1193]" pageId="8" pageNumber="109">
Although
<bibRefCitation id="EFCE4B2A462FFFC6FE88C976FE71531A" author="Anderson, T." box="[298,488,1024,1049]" pageId="8" pageNumber="109" pagination="13 - 54" refId="ref12377" refString="Anderson, T. (1864) An enumeration of the species of Acanthaceae from the continent and the adjacent islands. Journal of the Proceedings of the Linnean Society 7: 13 - 54." type="journal article" year="1864">Anderson (1864)</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A462FFFC6FE53C976FD15531A" author="Schinz, H." box="[497,652,1024,1049]" pageId="8" pageNumber="109" pagination="179 - 230" refId="ref13310" refString="Schinz, H. (1890) Beitrage zur Kenntnis der Flora von Deutsch-Sudwest-Afrika und der angrenzenden Gebiete. Verhandlungen des Botanischen Vereins fur die Provinz Brandenbrug und die Angrenzenden Lander 31: 179 - 230." type="journal article" year="1890">Schinz (1890)</bibRefCitation>
and
<bibRefCitation id="EFCE4B2A462FFFC6FD1FC976FCCE531A" author="Clarke, C. B." box="[701,855,1024,1049]" pageId="8" pageNumber="109" pagination="1 - 92" refId="ref12444" refString="Clarke, C. B. (1901) Acanthaceae. In: Thiselton-Dyer, W. T. (Ed.) Flora capensis 5, 1. Lovell Reeve &amp; Co., London, pp. 1 - 92. https: // doi. org / 10.5962 / bhl. title. 821" type="book chapter" year="1901">Clarke (1901)</bibRefCitation>
treated
<taxonomicName id="4C5F4D58462FFFC6FC0EC976FB1D531A" authorityName="H. M. Steyn" authorityYear="2017" baseAuthorityName="H. M. Steyn" baseAuthorityYear="2017" box="[940,1156,1024,1049]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="glabra">
<emphasis id="B92BEAC9462FFFC6FC0EC976FB1D531A" box="[940,1156,1024,1049]" italics="true" pageId="8" pageNumber="109">Acanthopsis glabra</emphasis>
</taxonomicName>
as a variety of
<taxonomicName id="4C5F4D58462FFFC6FA80C976FE31533E" authority=", Drege (1843)" authorityName=", Drege" authorityYear="1843" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="carduifolia">
<emphasis id="B92BEAC9462FFFC6FA80C976FE93533E" italics="true" pageId="8" pageNumber="109">Acanthopsis carduifolia,</emphasis>
<bibRefCitation id="EFCE4B2A462FFFC6FEB0C952FE31533E" author="Drege, J. F." box="[274,424,1060,1085]" pageId="8" pageNumber="109" refId="ref12531" refString="Drege, J. F. (1843) Zwei Pflanzengeographische Documente, nebst einer Einleitung von Dr. E. Meyer. Regensburg, Leipzig, 230 pp. https: // doi. org / 10.5962 / bhl. title. 87612" type="book" year="1843">Drège (1843)</bibRefCitation>
</taxonomicName>
and
<bibRefCitation id="EFCE4B2A462FFFC6FE7CC952FC95533E" author="Nees von Esenbeck, C. G. D." box="[478,780,1060,1085]" pageId="8" pageNumber="109" pagination="46 - 519" refId="ref13072" refString="Nees von Esenbeck, C. G. D. (1847) Acanthaceae. In: Candolle, A. de. (Ed.) Prodromus systematis naturalis regni vegetabilis 11. Treuttel &amp; Wurtz, Paris, pp. 46 - 519." type="book chapter" year="1847">Nees von Esenbeck (1847)</bibRefCitation>
recognised it at species level in
<taxonomicName id="4C5F4D58462FFFC6FBD5C952FB46533E" box="[1143,1247,1060,1085]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthus" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="genus">
<emphasis id="B92BEAC9462FFFC6FBD5C952FB46533E" box="[1143,1247,1060,1085]" italics="true" pageId="8" pageNumber="109">Acanthus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D58462FFFC6FAB4C952FA32533E" box="[1302,1451,1060,1085]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthodium" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="genus">
<emphasis id="B92BEAC9462FFFC6FAB4C952FA32533E" box="[1302,1451,1060,1085]" italics="true" pageId="8" pageNumber="109">Acanthodium</emphasis>
</taxonomicName>
respectively. Until more information becomes available, we have decided to follow Drège and Nees in recognizing
<taxonomicName id="4C5F4D58462FFFC6FA31C93EFF4A5386" authorityName="H. M. Steyn" authorityYear="2017" baseAuthorityName="H. M. Steyn" baseAuthorityYear="2017" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="glabra">
<emphasis id="B92BEAC9462FFFC6FA31C93EFF4A5386" italics="true" pageId="8" pageNumber="109">A. glabra</emphasis>
</taxonomicName>
as a distinct species as the latter is probably more closely related to
<taxonomicName id="4C5F4D58462FFFC6FC1DC91AFA325386" authority="(S. Moore) Obermeyer (1937: 138)" authorityName="Obermeyer" authorityPageNumber="138" authorityYear="1937" baseAuthorityName="S. Moore" box="[959,1451,1132,1157]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC9462FFFC6FC1DC91AFBBF5386" box="[959,1062,1132,1157]" italics="true" pageId="8" pageNumber="109">A. scullyi</emphasis>
(S.Moore)
<bibRefCitation id="EFCE4B2A462FFFC6FB07C91AFA325386" author="Obermeyer, A. A." box="[1189,1451,1132,1157]" pageId="8" pageNumber="109" pagination="105 - 139" refId="ref13166" refString="Obermeyer, A. A. (1937) The South African species of Blepharis. Annals of the Transvaal Museum 14: 105 - 139." type="journal article" year="1937">Obermeyer (1937: 138)</bibRefCitation>
</taxonomicName>
than to
<taxonomicName id="4C5F4D58462FFFC6FF7EC9E6FEEF53AA" authorityName=", Drege" authorityYear="1843" box="[220,374,1168,1193]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="carduifolia">
<emphasis id="B92BEAC9462FFFC6FF7EC9E6FEEF53AA" box="[220,374,1168,1193]" italics="true" pageId="8" pageNumber="109">A. carduifolia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,291 @@
<document id="42BC6C265041C1131AE1133B6B50F23D" ID-DOI="10.11646/phytotaxa.324.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13697360" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725568056204" checkinUser="felipe" docAuthor="Steyn, Hester M. &amp; Van Wyk, Abraham E." docDate="2017" docId="03F687CD4629FFDEFF2ACE3AFD665693" docLanguage="en" docName="phytotaxa.324.2.1.pdf" docOrigin="Phytotaxa 324 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.324.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Acanthopsis insueta H. M. Steyn 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="117" masterDocId="FFCFFFB54627FFCEFFA2CD76FF995703" masterDocTitle="Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts" masterLastPageNumber="121" masterPageNumber="101" pageNumber="115" updateTime="1725570993962" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="60C0A506E873C928D71356A68BFABFEA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="92583700E141D0AE96B7AFDCC80CB2BB">
<mods:title id="B2E6A0BE82EABFEFC42FA079D8575818">Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts</mods:title>
</mods:titleInfo>
<mods:name id="D594C687D596F1DB9AB4F6EC72A3CF6F" type="personal">
<mods:role id="047F14272AB3CAF6A45E1020C86CA23B">
<mods:roleTerm id="3080877C60D80796538A28894009EC29">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="798978845F1292D21D9EC509A038115E">Steyn, Hester M.</mods:namePart>
<mods:affiliation id="35DAF4F54C9CBAC95FAD118232413824">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:name id="E48E6ED1DA65C23C3F884FE6F31B5A8C" type="personal">
<mods:role id="864A4E788BE42291AE23182BC8490BD8">
<mods:roleTerm id="AD07B2274D1A4CB7AA743675D166FC9D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F23E45EACBD05A8AE62E6A4B5B85E006">Van Wyk, Abraham E.</mods:namePart>
<mods:affiliation id="68F4A872FA61EDACF7F226CBAFE893B2">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0F6F424EDA9D6BF858E2B910F3B052EE">text</mods:typeOfResource>
<mods:relatedItem id="3A8DCE7B2FABE4F685B4E74692834D6F" type="host">
<mods:titleInfo id="73051457FDDC2EFBBD9E302E3A5430BC">
<mods:title id="43177C4ACFE635E903D5500142C212E1">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="016139D890F508A63A6294F976C2B42F">
<mods:date id="BBE7BED56B81EDC96788DAE69B783D18">2017</mods:date>
<mods:detail id="048B0CF884EAAB137911BAA5BB8F746A" type="pubDate">
<mods:number id="867B44079E3136B323279754325BE97A">2017-10-10</mods:number>
</mods:detail>
<mods:detail id="FA477C4C9A633C68CA96B895AE99B503" type="volume">
<mods:number id="44C29620AB7E3BFFC4A78E52E8AA42C8">324</mods:number>
</mods:detail>
<mods:detail id="4AC728B26EF689CD98E58FB794870D40" type="issue">
<mods:number id="4A85235B930F25105B76E4622BD553D4">2</mods:number>
</mods:detail>
<mods:extent id="59060614B6BB2B5FA8ED2C0088D89B85" unit="page">
<mods:start id="F8F0CEFB15BCAB9F46B345D8EF2838EE">101</mods:start>
<mods:end id="9DC9DBEC3B1FBE0FED64EEF10B6C3A02">121</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="55420898646912754CD26AD7A779C504">
<mods:url id="BE3E42AA77FE9875EB7930394C996986">http://dx.doi.org/10.11646/phytotaxa.324.2.1</mods:url>
</mods:location>
<mods:classification id="EBA5A6D3719D81981D9FBFD2D1DFEF79">journal article</mods:classification>
<mods:identifier id="CF41FAE7EFFD9F9AF26B6162E0BFE794" type="DOI">10.11646/phytotaxa.324.2.1</mods:identifier>
<mods:identifier id="4F21024C1EE00CF29FA87FA9CE659FA9" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="364BBB6E912D20BAA708C19CC463249F" type="Zenodo-Dep">13697360</mods:identifier>
</mods:mods>
<treatment id="03F687CD4629FFDEFF2ACE3AFD665693" ID-DOI="http://doi.org/10.5281/zenodo.13697386" ID-Zenodo-Dep="13697386" LSID="urn:lsid:plazi:treatment:03F687CD4629FFDEFF2ACE3AFD665693" httpUri="http://treatment.plazi.org/id/03F687CD4629FFDEFF2ACE3AFD665693" lastPageId="16" lastPageNumber="117" pageId="14" pageNumber="115">
<subSubSection id="C34565504629FFC0FF2ACE3AFCAE5466" box="[136,823,842,869]" pageId="14" pageNumber="115" type="nomenclature">
<paragraph id="8BE036DB4629FFC0FF2ACE3AFCAE5466" blockId="14.[136,1452,842,1119]" box="[136,823,842,869]" pageId="14" pageNumber="115">
<heading id="D0A881B74629FFC0FF2ACE3AFCAE5466" box="[136,823,842,869]" fontSize="11" level="2" pageId="14" pageNumber="115" reason="4">
<emphasis id="B92BEAC94629FFC0FF2ACE3AFF045466" bold="true" box="[136,157,844,869]" pageId="14" pageNumber="115">5.</emphasis>
<taxonomicName id="4C5F4D584629FFC0FF06CE3CFD885466" ID-CoL="8ZLD" authority="H.M.Steyn" authorityName="H. M. Steyn" authorityYear="2017" box="[164,529,842,869]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="species" species="insueta" status="sp. nov.">
<emphasis id="B92BEAC94629FFC0FF06CE3CFE145467" bold="true" box="[164,397,842,869]" italics="true" pageId="14" pageNumber="115">Acanthopsis insueta</emphasis>
H.M.Steyn
</taxonomicName>
,
<taxonomicNameLabel id="A21857B24629FFC0FDBFCE3AFDEA5466" box="[541,627,844,869]" pageId="14" pageNumber="115" rank="species">
<emphasis id="B92BEAC94629FFC0FDBFCE3AFDF25466" box="[541,619,844,869]" italics="true" pageId="14" pageNumber="115">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="13642A5E4629FFC0FD20CE3AFD565466" box="[642,719,844,869]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="14" pageNumber="115">
Figs.
<geoCoordinate id="EE6B501C4629FFC0FD63CE3AFD565466" box="[705,719,844,869]" degrees="1, 3" direction="east" orientation="longitude" pageId="14" pageNumber="115" precision="5555" value="1.3">1</geoCoordinate>
</figureCitation>
,
<figureCitation id="13642A5E4629FFC0FD7FCE3AFD645466" box="[733,765,844,869]" captionStart="FIGURE 3" captionStartId="6.[136,229,1911,1932]" captionTargetBox="[151,1436,190,1886]" captionTargetId="figure-15@6.[151,1436,190,1886]" captionTargetPageId="6" captionText="FIGURE 3. Comparison of floral parts, associated bracts and bracteoles (from left to right): bracts (2), bracteoles (2), calyx (4 sepals: dorsal, ventral and 2 lateral ones) and corolla with anthers.A. A. glabra, B. A. scullyi, C. A. nitida, D. A. glandulopalmata, E. A. insueta, F. A. carduifolia. Note: the bract on the left is from lower down in the same inflorescence. Scale bar = 10 mm. Vouchers:A = Steyn 1893 (PRE), B = Steyn 1911 (PRE), C = Steyn 1884 (PRE), D = Steyn 1890 (PRE), E = Steyn 1805 (PRE), F = Koekemoer 4620 (PRE)." figureDoi="http://doi.org/10.5281/zenodo.13697366" httpUri="https://zenodo.org/record/13697366/files/figure.png" pageId="14" pageNumber="115">3E</figureCitation>
&amp;
<figureCitation id="13642A5E4629FFC0FC82CE3AFCB75466" box="[800,814,844,869]" captionStart="FIGURE 8" captionStartId="15.[136,229,1958,1979]" captionTargetBox="[151,1436,353,1933]" captionTargetId="figure-96@15.[151,1436,353,1933]" captionTargetPageId="15" captionText="FIGURE 8. Acanthopsis insuenta. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp; F. E &amp;F. Bract. G. Calyx. H. Corolla (lobes lilac; throat lemon-yellow). I. Glandular hairs at base of style. J. Capsule. K. Capsule with seed. Vouchers: CK = Steyn 1805 (PRE). Photographed by: H.M. Steyn (AK)." figureDoi="http://doi.org/10.5281/zenodo.13697378" httpUri="https://zenodo.org/record/13697378/files/figure.png" pageId="14" pageNumber="115">8</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34565504629FFC0FF2ACE04FB0E54A8" pageId="14" pageNumber="115" type="materials_examined">
<paragraph id="8BE036DB4629FFC0FF2ACE04FB0E54A8" blockId="14.[136,1452,842,1119]" pageId="14" pageNumber="115">
<emphasis id="B92BEAC94629FFC0FF2ACE04FF5B5484" bold="true" box="[136,194,882,903]" pageId="14" pageNumber="115">Type:</emphasis>
<materialsCitation id="3B373C864629FFC0FF79CE04FB7A5484" box="[219,1251,882,903]" country="South Africa" location="Namaqualand District. Richtersveld National Park" pageId="14" pageNumber="115" specimenCount="1" stateProvince="Northern Cape">
<collectingCountry id="F348764B4629FFC0FF79CE04FE165484" box="[219,399,882,903]" name="South Africa" pageId="14" pageNumber="115">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="499BF8394629FFC0FE3FCE04FDAD5484" box="[413,564,882,903]" country="South Africa" name="Northern Cape" pageId="14" pageNumber="115">Northern Cape</collectingRegion>
:
<location id="8E8060004629FFC0FDE0CE04FBA55484" LSID="urn:lsid:plazi:treatment:03F687CD4629FFDEFF2ACE3AFD665693:8E8060004629FFC0FDE0CE04FBA55484" box="[578,1084,882,903]" country="South Africa" name="Namaqualand District. Richtersveld National Park" pageId="14" pageNumber="115" stateProvince="Northern Cape">Namaqualand District. Richtersveld National Park</location>
. Vyftienmylberg
</materialsCitation>
;
<materialsCitation id="3B373C864629FFC0FB50CE04FB0A54A8" collectingDate="2010-09-07" collectionCode="AA, NBG" country="South Africa" elevation="292" pageId="14" pageNumber="115" specimenCode="PRE0861485" specimenCount="2" typeStatus="holotype">
on track to top of mountain,
<quantity id="4CA79B3E4629FFC0FE86CEE0FEFA54A8" box="[292,355,918,939]" metricMagnitude="2" metricUnit="m" metricValue="2.92" pageId="14" pageNumber="115" unit="m" value="292.0">
<elevation id="0072D1E84629FFC0FE86CEE0FEFA54A8" box="[292,355,918,939]" metricMagnitude="2" metricUnit="m" metricValue="2.92" pageId="14" pageNumber="115" unit="m" value="292.0">292 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E4629FFC0FE04CEE0FE5254A8" box="[422,459,918,939]" country="Kazakhstan" lsid="urn:lsid:biocol.org:col:13590" name="Ministry of Science, Academy of Sciences" pageId="14" pageNumber="115" type="Herbarium">AA</collectionCode>
),
<date id="FFE1101B4629FFC0FE79CEE0FD1554A8" box="[475,652,918,939]" pageId="14" pageNumber="115" value="2010-09-07">
<collectingDate id="EFA5E9F34629FFC0FE79CEE0FD1554A8" box="[475,652,918,939]" pageId="14" pageNumber="115" value="2010-09-07">7 September 2010</collectingDate>
</date>
,
<emphasis id="B92BEAC94629FFC0FD34CEE0FC9954A8" box="[662,768,918,939]" italics="true" pageId="14" pageNumber="115">Steyn 1805</emphasis>
, (
<typeStatus id="54E488794629FFC0FCB0CEE0FCF154A8" box="[786,872,918,939]" pageId="14" pageNumber="115" type="holotype">holotype</typeStatus>
<specimenCode id="DBF99EA04629FFC0FCCCCEE0FC6F54A8" box="[878,1014,918,939]" collectionCode="PRE" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="14" pageNumber="115" type="Herbarium">PRE0861485</specimenCode>
!;
<typeStatus id="54E488794629FFC0FBA1CEE0FBD154A8" box="[1027,1096,918,939]" pageId="14" pageNumber="115" type="isotype">isotype</typeStatus>
<collectionCode id="ED4EAE1E4629FFC0FBECCEE0FB1054A8" box="[1102,1161,918,939]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="14" pageNumber="115" type="Herbarium">NBG</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34565504629FFC0FF2ACECCFBD351A2" pageId="14" pageNumber="115" type="diagnosis">
<paragraph id="8BE036DB4629FFC0FF2ACECCFC71535C" blockId="14.[136,1452,842,1119]" pageId="14" pageNumber="115">
<emphasis id="B92BEAC94629FFC0FF2ACECCFF6A54CC" bold="true" box="[136,243,954,975]" pageId="14" pageNumber="115">Diagnosis:</emphasis>
<taxonomicName id="4C5F4D584629FFC0FEA9CECCFE5254CC" authorityName="H. M. Steyn" authorityYear="2017" box="[267,459,954,975]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="species" species="insueta">
<emphasis id="B92BEAC94629FFC0FEA9CECCFE5254CC" box="[267,459,954,975]" italics="true" pageId="14" pageNumber="115">Acanthopsis insueta</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5F4D584629FFC0FDEBCECCFC9E54CC" authorityName="H. M. Steyn" authorityYear="2017" box="[585,775,954,975]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="species" species="glandulopalmata">
<emphasis id="B92BEAC94629FFC0FDEBCECCFC9E54CC" box="[585,775,954,975]" italics="true" pageId="14" pageNumber="115">A. glandulopalmata</emphasis>
</taxonomicName>
in having middle to upper bracts ending in (3)5 ensiform or narrowly winged primary spines (
<emphasis id="B92BEAC94629FFC0FE05CEA8FE5F54F0" box="[423,454,990,1011]" italics="true" pageId="14" pageNumber="115">not</emphasis>
5 winged primary spines) and the lateral spines usually with a basal secondary spine on the side facing the central primary spine (
<emphasis id="B92BEAC94629FFC0FE1EC974FE425314" box="[444,475,1026,1047]" italics="true" pageId="14" pageNumber="115">not</emphasis>
lateral spines always simple) and from
<taxonomicName id="4C5F4D584629FFC0FCF9C974FC795314" authorityName=", Drege" authorityYear="1843" box="[859,992,1026,1047]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="species" species="carduifolia">
<emphasis id="B92BEAC94629FFC0FCF9C974FC795314" box="[859,992,1026,1047]" italics="true" pageId="14" pageNumber="115">A. carduifolia</emphasis>
</taxonomicName>
in having grey leaves with appressed hairs (
<emphasis id="B92BEAC94629FFC0FA2FC974FA355314" box="[1421,1452,1026,1047]" italics="true" pageId="14" pageNumber="115">not</emphasis>
green leaves with strigose hairs) and bracts with secondary spines often longer than primary spines (
<emphasis id="B92BEAC94629FFC0FB34C950FB2C5338" box="[1174,1205,1062,1083]" italics="true" pageId="14" pageNumber="115">not</emphasis>
secondary spines always shorter than primary spines) and a lemon-yellow (
<emphasis id="B92BEAC94629FFC0FD3CC93CFD24535C" box="[670,701,1098,1119]" italics="true" pageId="14" pageNumber="115">not</emphasis>
cream or white) corolla throat.
</paragraph>
<paragraph id="8BE036DB4629FFC0FF2AC9E6FBD351A2" blockId="14.[136,1453,1168,2057]" pageId="14" pageNumber="115">
Perennial, compact subshrub,
<quantity id="4CA79B3E4629FFC0FE7EC9E6FDBA53AA" box="[476,547,1168,1193]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="115" unit="cm" value="15.0">15 cm</quantity>
tall with very short internodes. Leaves sessile, decurrent, base petiole-like with spines
<quantity id="4CA79B3E4629FFC0FF75C9C2FEAC53CE" box="[215,309,1204,1229]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="14" pageNumber="115" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">36 mm</quantity>
long; lamina oblanceolate, (25)3550(70) × 810(12) mm, grey with appressed, densely packed short hairs; apex acute, base attenuate, margin undulate, dentate-spinose to coarsely dentate-spinose, spines rigid, 47(8) mm long, yellow to dark brown. Inflorescences semi-dense, cylindrical, subsessile, ca.
<quantity id="4CA79B3E4629FFC0FB69C98AFADF5216" box="[1227,1350,1276,1301]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="14" pageNumber="115" unit="mm" value="40.0" valueMax="45.0" valueMin="35.0">3545 mm</quantity>
long, 8 10(12) mm in diameter. Bracts obovate to wedge-shaped,
<quantity id="4CA79B3E4629FFC0FC84C856FC07523A" box="[806,926,1312,1337]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="14" pageNumber="115" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">1113 mm</quantity>
long (including spines), lamina
<quantity id="4CA79B3E4629FFC0FAAFC856FAF2523A" box="[1293,1387,1312,1337]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="14" pageNumber="115" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">58 mm</quantity>
long; spines and lamina pubescent to hirsute with deflexed to spreading short hairs, lamina often with isolated long white hairs abaxially, silky-hairy adaxially; lower bracts ending in 5 primary spines; middle and upper bracts ending in (3)5 primary spines, upper bracts very often 3-fid (see
<figureCitation id="13642A5E4629FFC0FCB2C8FAFCC752A6" box="[784,862,1420,1445]" captionStart="FIGURE 3" captionStartId="6.[136,229,1911,1932]" captionTargetBox="[151,1436,190,1886]" captionTargetId="figure-15@6.[151,1436,190,1886]" captionTargetPageId="6" captionText="FIGURE 3. Comparison of floral parts, associated bracts and bracteoles (from left to right): bracts (2), bracteoles (2), calyx (4 sepals: dorsal, ventral and 2 lateral ones) and corolla with anthers.A. A. glabra, B. A. scullyi, C. A. nitida, D. A. glandulopalmata, E. A. insueta, F. A. carduifolia. Note: the bract on the left is from lower down in the same inflorescence. Scale bar = 10 mm. Vouchers:A = Steyn 1893 (PRE), B = Steyn 1911 (PRE), C = Steyn 1884 (PRE), D = Steyn 1890 (PRE), E = Steyn 1805 (PRE), F = Koekemoer 4620 (PRE)." figureDoi="http://doi.org/10.5281/zenodo.13697366" httpUri="https://zenodo.org/record/13697366/files/figure.png" pageId="14" pageNumber="115">Figs. 3</figureCitation>
&amp;
<figureCitation id="13642A5E4629FFC0FC25C8FAFC0F52A6" box="[903,918,1420,1445]" captionStart="FIGURE 8" captionStartId="15.[136,229,1958,1979]" captionTargetBox="[151,1436,353,1933]" captionTargetId="figure-96@15.[151,1436,353,1933]" captionTargetPageId="15" captionText="FIGURE 8. Acanthopsis insuenta. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp; F. E &amp;F. Bract. G. Calyx. H. Corolla (lobes lilac; throat lemon-yellow). I. Glandular hairs at base of style. J. Capsule. K. Capsule with seed. Vouchers: CK = Steyn 1805 (PRE). Photographed by: H.M. Steyn (AK)." figureDoi="http://doi.org/10.5281/zenodo.13697378" httpUri="https://zenodo.org/record/13697378/files/figure.png" pageId="14" pageNumber="115">8</figureCitation>
); central primary spine usually simple or with short marginal secondary spines; lateral primary spines usually with long basal secondary spine on side facing central primary spine, often longer than primary spines; spines spreading to recurved in fruit;. Bracteoles linear, 45(6) mm long, silky-hairy. Calyx with dorsal sepal ovate, acuminate,
<quantity id="4CA79B3E4629FFC0FC9DC88EFC205112" box="[831,953,1528,1553]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="14" pageNumber="115" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">1112 mm</quantity>
long, silky-hairy especially abaxially, 79- veined; ventral sepal ovate,
<quantity id="4CA79B3E4629FFC0FE6FCB6AFDA05136" box="[461,569,1564,1589]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="14" pageNumber="115" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">911 mm</quantity>
long, silky-hairy especially abaxially, 57-veined; lateral sepals
<quantity id="4CA79B3E4629FFC0FA84CB6AFAF05136" box="[1318,1385,1564,1589]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="115" unit="mm" value="9.0">9 mm</quantity>
long, silky-hairy. Corolla lilac to purple with lemon-yellow throat, 25 ×
<quantity id="4CA79B3E4629FFC0FC22CB36FC4A515A" box="[896,979,1600,1625]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="115" unit="mm" value="10.0">10 mm</quantity>
, tube
<quantity id="4CA79B3E4629FFC0FBBBCB36FBC5515A" box="[1049,1116,1600,1625]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="115" unit="mm" value="9.0">9 mm</quantity>
long, central lobe wider than long, constricted at base, emarginate. Filaments
<quantity id="4CA79B3E4629FFC0FD10CB12FD6C517E" box="[690,757,1636,1661]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="115" unit="mm" value="6.0">6 mm</quantity>
long, glandular; anthers beige to orange-brown,
<quantity id="4CA79B3E4629FFC0FA85CB12FAF3517E" box="[1319,1386,1636,1661]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="115" unit="mm" value="3.0">3 mm</quantity>
long. Style with patch of glandular hairs at the base. Capsules 8 ×
<quantity id="4CA79B3E4629FFC0FC8CCBFEFC1751A2" box="[814,910,1672,1697]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="14" pageNumber="115" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
. Seed 5 ×
<quantity id="4CA79B3E4629FFC0FBA0CBFEFBDF51A2" box="[1026,1094,1672,1697]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="115" unit="mm" value="3.0">3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34565504629FFC0FF1FCBDAFD5B51EA" pageId="14" pageNumber="115" type="etymology">
<paragraph id="8BE036DB4629FFC0FF1FCBDAFD5B51EA" blockId="14.[136,1453,1168,2057]" pageId="14" pageNumber="115">
<emphasis id="B92BEAC94629FFC0FF1FCBDAFED151C6" bold="true" box="[189,328,1708,1733]" pageId="14" pageNumber="115">Etymology:</emphasis>
—The specific epithet
<taxonomicName id="4C5F4D584629FFC0FDEFCBDAFD0751C6" authorityName="H. M. Steyn" authorityYear="2017" box="[589,670,1708,1733]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="species" species="insueta">
<emphasis id="B92BEAC94629FFC0FDEFCBDAFD0751C6" box="[589,670,1708,1733]" italics="true" pageId="14" pageNumber="115">insueta</emphasis>
</taxonomicName>
, meaning “unusual”, refers to the unusual combination of middle to upper floral bracts ending in 3 or 5 primary spines.
</paragraph>
</subSubSection>
<subSubSection id="C34565504629FFDEFF1FCB82FD665693" lastPageId="16" lastPageNumber="117" pageId="14" pageNumber="115" type="distribution">
<paragraph id="8BE036DB4629FFC0FF1FCB82FD075F0A" blockId="14.[136,1453,1168,2057]" pageId="14" pageNumber="115">
<emphasis id="B92BEAC94629FFC0FF1FCB82FDEC500E" bold="true" box="[189,629,1780,1805]" pageId="14" pageNumber="115">Distribution, ecology and phenology:</emphasis>
<taxonomicName id="4C5F4D584629FFC0FD33CB82FCEB500E" authorityName="H. M. Steyn" authorityYear="2017" box="[657,882,1780,1805]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="species" species="insueta">
<emphasis id="B92BEAC94629FFC0FD33CB82FCEB500E" box="[657,882,1780,1805]" italics="true" pageId="14" pageNumber="115">Acanthopsis insueta</emphasis>
</taxonomicName>
is endemic to the Vyftienmylberg (
<collectingRegion id="499BF8394629FFC0FB59CB82FA3D500E" box="[1275,1444,1780,1805]" country="South Africa" name="Northern Cape" pageId="14" pageNumber="115">Northern Cape</collectingRegion>
) and is known from two localities (
<figureCitation id="13642A5E4629FFC0FDA4CA6EFDDE5032" box="[518,583,1816,1841]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="14" pageNumber="115">Fig. 1</figureCitation>
). It is centred in the core part of the Gariep Centre of Endemism (
<bibRefCitation id="EFCE4B2A4629FFC0FA84CA6EFE965056" author="Van Wyk, A. E. &amp; Smith, G. F." pageId="14" pageNumber="115" refId="ref13657" refString="Van Wyk, A. E. &amp; Smith, G. F. (2001) Regions of floristic endemism in southern Africa: a review with emphasis on succulents. Umdaus Press, Pretoria, 160 pp." type="book" year="2001">Van Wyk &amp; Smith 2001</bibRefCitation>
). This distribution falls within the Succulent Karoo Biome (
<bibRefCitation id="EFCE4B2A4629FFC0FC14CA4AFB6E5056" author="Rutherford, M. C. &amp; Westfall, R. H." box="[950,1271,1852,1877]" pageId="14" pageNumber="115" refId="ref13271" refString="Rutherford, M. C. &amp; Westfall, R. H. (1994) Biomes of southern Africa: an objective categorization. [Memoir 63]. National Botanical Institute, Pretoria, 94 pp." type="book" year="1994">Rutherford &amp; Westfall 1994</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A4629FFC0FAA1CA4AFF5A507A" author="Low, A. B. &amp; Rebelo, A. G." pageId="14" pageNumber="115" refId="ref12933" refString="Low, A. B., Rebelo, A. G. (Eds.) (1996) Vegetation of South Africa, Lesotho and Swaziland. Department of Environmental Affairs &amp; Tourism, Pretoria, 85 pp." type="book" year="1996">Low &amp; Rebelo 1996</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A4629FFC0FF6FCA16FD98507A" author="Mucina, L. &amp; Rutherford, M. C." box="[205,513,1888,1913]" pageId="14" pageNumber="115" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
). It is found in the Richtersveld Bioregion, Vyftienmyl se Berge Succulent Shrubland (
<bibRefCitation id="EFCE4B2A4629FFC0FF32CAF2FE55509E" author="Mucina, L. &amp; Rutherford, M. C." box="[144,460,1924,1949]" pageId="14" pageNumber="115" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) where it receives predominantly winter rains with an average of
<quantity id="4CA79B3E4629FFC0FB1CCAF2FADE509E" box="[1214,1351,1924,1949]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="14" pageNumber="115" unit="mm" value="90.0" valueMax="120.0" valueMin="60.0">60120 mm</quantity>
per year (including considerable amounts of fog). The specimens were collected on mountain slopes associated with quartzite stones in well-drained, sandy soils at elevations between 230 and
<specimenCount id="9D59FD524629FFC0FCDACABAFC5150E6" box="[888,968,1996,2021]" count="445" pageId="14" pageNumber="115" type="generic">
<quantity id="4CA79B3E4629FFC0FCDACABAFC5D50E6" box="[888,964,1996,2021]" metricMagnitude="2" metricUnit="m" metricValue="4.45" pageId="14" pageNumber="115" unit="m" value="445.0">445 m</quantity>
.
</specimenCount>
Specimens with flowers were collected in August (probably also flowers into September).
</paragraph>
<paragraph id="8BE036DB4628FFC1FF1FCDD6FEBA564A" blockId="15.[136,1452,160,329]" pageId="15" pageNumber="116">
<emphasis id="B92BEAC94628FFC1FF1FCDD6FE2E57BA" bold="true" box="[189,439,160,185]" pageId="15" pageNumber="116">Conservation status:</emphasis>
<taxonomicName id="4C5F4D584628FFC1FE71CDD6FD2F57BA" authorityName="H. M. Steyn" authorityYear="2017" box="[467,694,160,185]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="15" pageNumber="116" phylum="Tracheophyta" rank="species" species="insueta">
<emphasis id="B92BEAC94628FFC1FE71CDD6FD2F57BA" box="[467,694,160,185]" italics="true" pageId="15" pageNumber="116">Acanthopsis insueta</emphasis>
</taxonomicName>
is a highly localized Richtersveld endemic (EOO &lt;
<quantity id="4CA79B3E4628FFC1FAADCDD6FAC557BA" box="[1295,1372,160,185]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="116" unit="km" value="10.0">10 km</quantity>
²). This species is known from a small area on the northern section of the summit of Vyftienmyl-se-Berg, an inselberg that is home to many local endemics. As there are no severe threats, the population is not likely to be declining. As its habitat is well-protected and the species is not in danger of extinction, it is given a status of Critically Rare (L. von Staden, pers. comm.).
</paragraph>
<caption id="DF2066534628FFC1FF2ACAD0FE6A5F00" ID-DOI="http://doi.org/10.5281/zenodo.13697378" ID-Zenodo-Dep="13697378" httpUri="https://zenodo.org/record/13697378/files/figure.png" pageId="15" pageNumber="116" startId="15.[136,229,1958,1979]" targetBox="[151,1436,353,1933]" targetPageId="15" targetType="figure">
<paragraph id="8BE036DB4628FFC1FF2ACAD0FE6A5F00" blockId="15.[136,1452,1958,2051]" pageId="15" pageNumber="116">
<emphasis id="B92BEAC94628FFC1FF2ACAD0FF6650B8" bold="true" box="[136,255,1958,1979]" pageId="15" pageNumber="116">FIGURE 8.</emphasis>
<taxonomicName id="4C5F4D584628FFC1FEA4CAD0FE4A50B8" box="[262,467,1958,1979]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="15" pageNumber="116" phylum="Tracheophyta" rank="species" species="insuenta">
<emphasis id="B92BEAC94628FFC1FEA4CAD0FE4A50B8" box="[262,467,1958,1979]" italics="true" pageId="15" pageNumber="116">Acanthopsis insuenta</emphasis>
</taxonomicName>
. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp; F. E &amp;F. Bract. G. Calyx. H. Corolla (lobes lilac; throat lemon-yellow). I. Glandular hairs at base of style. J. Capsule. K. Capsule with seed. Vouchers: CK =
<emphasis id="B92BEAC94628FFC1FB54CABCFAF850DC" box="[1270,1377,1994,2015]" italics="true" pageId="15" pageNumber="116">Steyn 1805</emphasis>
(PRE). Photographed by: H.M. Steyn (AK).
</paragraph>
</caption>
<paragraph id="8BE036DB4637FFDEFF1FCDE9FF465603" blockId="16.[136,1452,159,400]" pageId="16" pageNumber="117">
<emphasis id="B92BEAC94637FFDEFF1FCDE9FD4157BB" bold="true" box="[189,728,159,184]" pageId="16" pageNumber="117">
Additional specimen examined (
<typeStatus id="54E488794637FFDEFDF5CDE9FD5257BB" box="[599,715,159,184]" pageId="16" pageNumber="117" type="paratype">Paratype</typeStatus>
):
</emphasis>
<materialsCitation id="3B373C864637FFDEFD57CDE9FD1C57DF" country="South Africa" location="Namaqualand District. Richtersveld National Park" pageId="16" pageNumber="117" specimenCount="1" stateProvince="Northern Cape">
<collectingCountry id="F348764B4637FFDEFD57CDE9FC5357BB" box="[757,970,159,184]" name="South Africa" pageId="16" pageNumber="117">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="499BF8394637FFDEFC7CCDE9FB0857BB" box="[990,1169,159,184]" country="South Africa" name="Northern Cape" pageId="16" pageNumber="117">Northern Cape</collectingRegion>
:
<location id="8E8060004637FFDEFB07CDE9FE5D57DF" LSID="urn:lsid:plazi:treatment:03F687CD4629FFDEFF2ACE3AFD665693:8E8060004637FFDEFB07CDE9FE5D57DF" country="South Africa" name="Namaqualand District. Richtersveld National Park" pageId="16" pageNumber="117" stateProvince="Northern Cape">Namaqualand District. Richtersveld National Park</location>
. Vyftienmylberg
</materialsCitation>
;
<materialsCitation id="3B373C864637FFDEFD36CDB5FF425603" collectingDate="2010-09-09" collectionCode="AA, PRE" country="South Africa" elevation="444" pageId="16" pageNumber="117" specimenCount="1">
top of mountain,
<quantity id="4CA79B3E4637FFDEFCC0CDB5FC3657DF" box="[866,943,195,220]" metricMagnitude="2" metricUnit="m" metricValue="4.4399999999999995" pageId="16" pageNumber="117" unit="m" value="444.0">
<elevation id="0072D1E84637FFDEFCC0CDB5FC3657DF" box="[866,943,195,220]" metricMagnitude="2" metricUnit="m" metricValue="4.4399999999999995" pageId="16" pageNumber="117" unit="m" value="444.0">444 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E4637FFDEFBA0CDB5FBB457DF" box="[1026,1069,195,220]" country="Kazakhstan" lsid="urn:lsid:biocol.org:col:13590" name="Ministry of Science, Academy of Sciences" pageId="16" pageNumber="117" type="Herbarium">AA</collectionCode>
),
<date id="FFE1101B4637FFDEFBE7CDB5FA8557DF" box="[1093,1308,195,220]" pageId="16" pageNumber="117" value="2010-09-09">
<collectingDate id="EFA5E9F34637FFDEFBE7CDB5FA8557DF" box="[1093,1308,195,220]" pageId="16" pageNumber="117" value="2010-09-09">9 September 2010</collectingDate>
</date>
,
<emphasis id="B92BEAC94637FFDEFA89CDB5FA3257DF" box="[1323,1451,195,220]" italics="true" pageId="16" pageNumber="117">Steyn 1806</emphasis>
(
<collectionCode id="ED4EAE1E4637FFDEFF33CD91FF565603" box="[145,207,231,256]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="16" pageNumber="117" type="Herbarium">PRE</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE036DB4637FFDEFF1FCC7DFD665693" blockId="16.[136,1452,159,400]" pageId="16" pageNumber="117">
<emphasis id="B92BEAC94637FFDEFF1FCC7DFE905627" bold="true" box="[189,265,267,292]" pageId="16" pageNumber="117">Notes:</emphasis>
—In the case where
<taxonomicName id="4C5F4D584637FFDEFDA0CC7DFD7E5627" authorityName="H. M. Steyn" authorityYear="2017" box="[514,743,267,292]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="species" species="insueta">
<emphasis id="B92BEAC94637FFDEFDA0CC7DFD7E5627" box="[514,743,267,292]" italics="true" pageId="16" pageNumber="117">Acanthopsis insueta</emphasis>
</taxonomicName>
has trifid bracts, it differs from
<taxonomicName id="4C5F4D584637FFDEFBCCCC7DFB445627" authorityName="Schinz" authorityYear="1890" baseAuthorityName="Nees" box="[1134,1245,267,292]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="species" species="glauca">
<emphasis id="B92BEAC94637FFDEFBCCCC7DFB445627" box="[1134,1245,267,292]" italics="true" pageId="16" pageNumber="117">A. glauca</emphasis>
</taxonomicName>
in having sessile inflorescences and bracts with primary spines of approximately the same length (
<emphasis id="B92BEAC94637FFDEFB82CC59FBDD564B" box="[1056,1092,303,328]" italics="true" pageId="16" pageNumber="117">not</emphasis>
pedunculate inflorescences and bracts with central primary spine much longer than lateral spines) and from all other species with trifid bracts in having ensiform spines (
<emphasis id="B92BEAC94637FFDEFEEBCC01FEF45693" box="[329,365,375,400]" italics="true" pageId="16" pageNumber="117">not</emphasis>
ovate, mucronate to spinose lobes).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,379 @@
<document id="384D7B834061D75C7D2DEA13FF241D21" ID-DOI="10.11646/phytotaxa.324.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13697360" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725568056204" checkinUser="felipe" docAuthor="Steyn, Hester M. &amp; Van Wyk, Abraham E." docDate="2017" docId="03F687CD462BFFC0FF2ACE89FB27541E" docLanguage="en" docName="phytotaxa.324.2.1.pdf" docOrigin="Phytotaxa 324 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.324.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Acanthopsis glandulopalmata H. M. Steyn 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="115" masterDocId="FFCFFFB54627FFCEFFA2CD76FF995703" masterDocTitle="Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts" masterLastPageNumber="121" masterPageNumber="101" pageNumber="113" updateTime="1725570993962" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F976DBAC9EBA1003F4FD1047F337D1FB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8F3B236CBE987B1664D1BDE22BD7AB54">
<mods:title id="B35FDB77DDCC990BA31E2C86CD121C21">Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts</mods:title>
</mods:titleInfo>
<mods:name id="A97D9616FC9C83507B8CD75113C434FC" type="personal">
<mods:role id="1868A1220DE933448D5C822A0FFA02C2">
<mods:roleTerm id="155E0F2ABB7F4FF19E289BBDF1251779">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F492B22952E3555F6A83A0272B841502">Steyn, Hester M.</mods:namePart>
<mods:affiliation id="A0BDD354E72D9BB4D06827847A87B85B">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:name id="F1D88CE7F4C69F8209A0B8E0C80B338E" type="personal">
<mods:role id="B920AC0ADC17EFFF8E6D4F444CB29B80">
<mods:roleTerm id="FEF9EA20588152F52336BBD1D1BACE37">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D711E24B12601B385FFB533D9117D67">Van Wyk, Abraham E.</mods:namePart>
<mods:affiliation id="385EBFE7C15F5D4D2ADD71D29492F839">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="725CB84B1EEC1BEB78C11F8C019EE6B6">text</mods:typeOfResource>
<mods:relatedItem id="243102D59435BC04024268E83280683F" type="host">
<mods:titleInfo id="0A511635DD20FDB348407D04CB57EDF7">
<mods:title id="528C629315643D8EAE8DBCACFC6BB917">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="722675C18F58CFB16CEE4464D0301C82">
<mods:date id="B2D47EB6DA95E4D7712A084D7751801A">2017</mods:date>
<mods:detail id="D369FAD89A0A16C058F81C2E5991ABA3" type="pubDate">
<mods:number id="2F3350F9693A743AE4F50F16165D0E45">2017-10-10</mods:number>
</mods:detail>
<mods:detail id="7D47FF2E78D3C8858808F7504C4E8EEC" type="volume">
<mods:number id="5F3CC9BCBAD6E4F200CCE89CF309F111">324</mods:number>
</mods:detail>
<mods:detail id="A5210AD46EFBE6F13CEA13FB4F2D00C5" type="issue">
<mods:number id="62A7DBE3A009CFF32084BD6F0C4487F0">2</mods:number>
</mods:detail>
<mods:extent id="AFC2529AAB4847BF2E0F897ED4EA8625" unit="page">
<mods:start id="354475053B5B0D4175416166D5974387">101</mods:start>
<mods:end id="7439D94938086CDBA1F9523EF2C19E88">121</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="0CFD1EEC0CD20EB87E3658841F4E639D">
<mods:url id="295690B0506892C7BC8EFEC32C799515">http://dx.doi.org/10.11646/phytotaxa.324.2.1</mods:url>
</mods:location>
<mods:classification id="930C47704747BB426AC11366A0AD48BA">journal article</mods:classification>
<mods:identifier id="EECD1784F41059F0FE457ECF9EFA27F3" type="DOI">10.11646/phytotaxa.324.2.1</mods:identifier>
<mods:identifier id="094F6B9D687FCF8D86F6704EAD20AE79" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="22FEF116D1F8AFF93643AD96325FFAE1" type="Zenodo-Dep">13697360</mods:identifier>
</mods:mods>
<treatment id="03F687CD462BFFC0FF2ACE89FB27541E" ID-DOI="http://doi.org/10.5281/zenodo.13697384" ID-Zenodo-Dep="13697384" LSID="urn:lsid:plazi:treatment:03F687CD462BFFC0FF2ACE89FB27541E" httpUri="http://treatment.plazi.org/id/03F687CD462BFFC0FF2ACE89FB27541E" lastPageId="14" lastPageNumber="115" pageId="12" pageNumber="113">
<subSubSection id="C3456550462BFFC2FF2ACE89FC33531B" box="[136,938,1021,1048]" pageId="12" pageNumber="113" type="nomenclature">
<paragraph id="8BE036DB462BFFC2FF2ACE89FC33531B" blockId="12.[136,1452,1021,1227]" box="[136,938,1021,1048]" pageId="12" pageNumber="113">
<heading id="D0A881B7462BFFC2FF2ACE89FC33531B" box="[136,938,1021,1048]" fontSize="11" level="2" pageId="12" pageNumber="113" reason="4">
<emphasis id="B92BEAC9462BFFC2FF2ACE89FF04531B" bold="true" box="[136,157,1022,1048]" pageId="12" pageNumber="113">4.</emphasis>
<taxonomicName id="4C5F4D58462BFFC2FF01CE8BFD19531B" ID-CoL="8ZL9" authority="H.M.Steyn" authorityName="H. M. Steyn" authorityYear="2017" box="[163,640,1021,1048]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="glandulopalmata" status="sp. nov.">
<emphasis id="B92BEAC9462BFFC2FF01CE8BFE65531B" bold="true" box="[163,508,1021,1048]" italics="true" pageId="12" pageNumber="113">Acanthopsis glandulopalmata</emphasis>
H.M.Steyn
</taxonomicName>
,
<taxonomicNameLabel id="A21857B2462BFFC2FD2ECE89FD7B531B" box="[652,738,1023,1048]" pageId="12" pageNumber="113" rank="species">
<emphasis id="B92BEAC9462BFFC2FD2ECE89FD43531B" box="[652,730,1023,1048]" italics="true" pageId="12" pageNumber="113">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="13642A5E462BFFC2FD53CE89FCA7531B" box="[753,830,1023,1048]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="12" pageNumber="113">Figs. 1</figureCitation>
,
<figureCitation id="13642A5E462BFFC2FCEECE89FCF6531B" box="[844,879,1023,1048]" captionStart="FIGURE 3" captionStartId="6.[136,229,1911,1932]" captionTargetBox="[151,1436,190,1886]" captionTargetId="figure-15@6.[151,1436,190,1886]" captionTargetPageId="6" captionText="FIGURE 3. Comparison of floral parts, associated bracts and bracteoles (from left to right): bracts (2), bracteoles (2), calyx (4 sepals: dorsal, ventral and 2 lateral ones) and corolla with anthers.A. A. glabra, B. A. scullyi, C. A. nitida, D. A. glandulopalmata, E. A. insueta, F. A. carduifolia. Note: the bract on the left is from lower down in the same inflorescence. Scale bar = 10 mm. Vouchers:A = Steyn 1893 (PRE), B = Steyn 1911 (PRE), C = Steyn 1884 (PRE), D = Steyn 1890 (PRE), E = Steyn 1805 (PRE), F = Koekemoer 4620 (PRE)." figureDoi="http://doi.org/10.5281/zenodo.13697366" httpUri="https://zenodo.org/record/13697366/files/figure.png" pageId="12" pageNumber="113">3D</figureCitation>
&amp;
<figureCitation id="13642A5E462BFFC2FC30CE89FC38531B" box="[914,929,1023,1048]" captionStart="FIGURE 7" captionStartId="13.[136,229,1958,1979]" captionTargetBox="[151,1436,353,1933]" captionTargetId="figure-96@13.[151,1436,353,1933]" captionTargetPageId="13" captionText="FIGURE 7. Acanthopsis glandulopalmata. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp;F. Bract. G. Calyx. H. Corolla (lobes lilac to purple; throat lemon-yellow). I. Glandular hairs at base of style. J. Capsule with seed. K. Seed. Vouchers: CI = Steyn 2125 (PRE), J &amp; K = Steyn 1889 (PRE). Photographed by: M. Koekemoer (A) and H.M. Steyn (BK)." figureDoi="http://doi.org/10.5281/zenodo.13697374" httpUri="https://zenodo.org/record/13697374/files/figure.png" pageId="12" pageNumber="113">7</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3456550462BFFC2FF2AC950FB23535C" pageId="12" pageNumber="113" type="materials_examined">
<paragraph id="8BE036DB462BFFC2FF2AC950FB23535C" blockId="12.[136,1452,1021,1227]" pageId="12" pageNumber="113">
<emphasis id="B92BEAC9462BFFC2FF2AC950FF5B5338" bold="true" box="[136,194,1062,1083]" pageId="12" pageNumber="113">Type:</emphasis>
<collectingCountry id="F348764B462BFFC2FF79C950FE145338" box="[219,397,1062,1083]" name="South Africa" pageId="12" pageNumber="113">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="499BF839462BFFC2FE3BC950FDB75338" box="[409,558,1062,1083]" country="South Africa" name="Northern Cape" pageId="12" pageNumber="113">Northern Cape</collectingRegion>
: Namaqualand District. On track between Anenous/Kosies to Umdaus in Wyepoort River Valley,
<quantity id="4CA79B3E462BFFC2FEA7C93CFEDD535C" box="[261,324,1098,1119]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="12" pageNumber="113" unit="m" value="690.0">690 m</quantity>
, (2917
<collectionCode id="ED4EAE1E462BFFC2FE2AC93CFE32535C" box="[392,427,1098,1119]" country="Argentina" lsid="urn:lsid:biocol.org:col:15002" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="12" pageNumber="113" type="Herbarium">BA</collectionCode>
),
<date id="FFE1101B462BFFC2FE19C93CFDCC535C" box="[443,597,1098,1119]" pageId="12" pageNumber="113" value="2012-08-25">25 August 2012</date>
,
<emphasis id="B92BEAC9462BFFC2FDFDC93CFD50535C" box="[607,713,1098,1119]" italics="true" pageId="12" pageNumber="113">Steyn 1890</emphasis>
(
<typeStatus id="54E48879462BFFC2FD74C93CFCB5535C" box="[726,812,1098,1119]" pageId="12" pageNumber="113" type="holotype">holotype</typeStatus>
<collectionCode id="ED4EAE1E462BFFC2FC93C93CFCF9535C" box="[817,864,1098,1119]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="12" pageNumber="113" type="Herbarium">PRE</collectionCode>
0861467!,
<typeStatus id="54E48879462BFFC2FC67C93CFB93535C" box="[965,1034,1098,1119]" pageId="12" pageNumber="113" type="isotype">isotype</typeStatus>
<collectionCode id="ED4EAE1E462BFFC2FBB2C93CFBDF535C" box="[1040,1094,1098,1119]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="12" pageNumber="113" type="Herbarium">NBG</collectionCode>
01461963!).
</paragraph>
</subSubSection>
<subSubSection id="C3456550462BFFC2FF2AC918FC5C5033" pageId="12" pageNumber="113" type="diagnosis">
<paragraph id="8BE036DB462BFFC2FF2AC918FBA453C8" blockId="12.[136,1452,1021,1227]" pageId="12" pageNumber="113">
<emphasis id="B92BEAC9462BFFC2FF2AC918FF6A5380" bold="true" box="[136,243,1134,1155]" pageId="12" pageNumber="113">Diagnosis:</emphasis>
<taxonomicName id="4C5F4D58462BFFC2FEA9C918FDB75380" authorityName="H. M. Steyn" authorityYear="2017" box="[267,558,1134,1155]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="glandulopalmata">
<emphasis id="B92BEAC9462BFFC2FEA9C918FDB75380" box="[267,558,1134,1155]" italics="true" pageId="12" pageNumber="113">Acanthopsis glandulopalmata</emphasis>
</taxonomicName>
differs from other species with semi-dense spikes and 5-fid bracts in having bracts with winged primary spines (nearly always simple) with long, glandular hairs (
<emphasis id="B92BEAC9462BFFC2FC23C9E4FC3953A4" box="[897,928,1170,1191]" italics="true" pageId="12" pageNumber="113">not</emphasis>
narrow to narrowly winged primary spines with short, eglandular hairs) and a relatively large corolla of
<quantity id="4CA79B3E462BFFC2FD3AC9C0FD6653C8" box="[664,767,1206,1227]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="12" pageNumber="113" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">2832 mm</quantity>
long (
<emphasis id="B92BEAC9462BFFC2FC9FC9C0FCC553C8" box="[829,860,1206,1227]" italics="true" pageId="12" pageNumber="113">not</emphasis>
2025(27) mm long).
</paragraph>
<paragraph id="8BE036DB462BFFC2FF2AC98DFC5C5033" blockId="12.[136,1452,1275,2092]" pageId="12" pageNumber="113">
Perennial, compact subshrub,
<quantity id="4CA79B3E462BFFC2FE74C98DFDAF5217" box="[470,566,1275,1300]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="12" pageNumber="113" unit="cm" value="8.5" valueMax="10.0" valueMin="7.0">710 cm</quantity>
tall with very short internodes. Leaves sessile, base petiole-like with spines
<quantity id="4CA79B3E462BFFC2FA20C98DFF2D523B" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="12" pageNumber="113" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">37 mm</quantity>
long; lamina oblanceolate, (25)3550(70) × 510(12) mm, strigose with short, white hairs, also with subsessile glandular hairs; apex acute, base attenuate, margin undulate, dentate-spinose, spines rigid,
<quantity id="4CA79B3E462BFFC2FB35C835FB6F525F" box="[1175,1270,1347,1372]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="12" pageNumber="113" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">36 mm</quantity>
long, yellow to brown. Inflorescences semi-dense, cylindrical, subsessile, 4550(70) mm long, 810(12) mm in diameter. Bracts obovate to broadly obovate, (1519)
<quantity id="4CA79B3E462BFFC2FD85C8FDFDEE52A7" box="[551,631,1419,1444]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="113" unit="mm" value="24.0">24 mm</quantity>
long (including spines), lamina
<quantity id="4CA79B3E462BFFC2FC79C8FDFBC852A7" box="[987,1105,1419,1444]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="12" pageNumber="113" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">1011 mm</quantity>
long, base obtuse; lower bracts hirsute with deflexed to spreading short, white hairs, middle to upper bracts hirsute with spreading short, white hairs, together with glandular hairs abaxially, silky-hairy, also with short, white hairs and glandular hairs adaxially; middle and upper bracts ending in 5 primary spines; central primary spine usually to nearly always simple, if compound, then with 1 pair of short marginal spines (prickles); lateral primary spines nearly always simple; spines spreading to recurved in fruit. Bracteoles linear, 78(11) mm long, silky-hairy with some isolated glandular hairs. Calyx with dorsal sepal ovate, acuminate, 1314(16) mm long, silky-hairy with some isolated glandular hairs abaxially, silky-hairy adaxially, 79-veined; ventral sepal ovate,
<quantity id="4CA79B3E462BFFC2FE54CBF1FDF651A3" box="[502,623,1671,1696]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="12" pageNumber="113" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">1214 mm</quantity>
long, silky-hairy with some isolated glandular hairs abaxially, silky-hairy adaxially, 57-veined; lateral sepals
<quantity id="4CA79B3E462BFFC2FD8BCBDDFDF551C7" box="[553,620,1707,1732]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="113" unit="mm" value="9.0">9 mm</quantity>
long, with some isolated glandular hairs abaxially, silky-hairy adaxially. Corolla lilac to purple with lemon-yellow throat, 2832 ×
<quantity id="4CA79B3E462BFFC2FC8BCBB9FC3C51EB" box="[809,933,1743,1768]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="12" pageNumber="113" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">1011 mm</quantity>
, tube
<quantity id="4CA79B3E462BFFC2FC4FCBB9FBC351EB" box="[1005,1114,1743,1768]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="12" pageNumber="113" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">910 mm</quantity>
long, central lobe as wide as long, truncate to rounded. Filaments
<quantity id="4CA79B3E462BFFC2FD88CB85FD1E500F" box="[554,647,1779,1804]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="12" pageNumber="113" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
long, glandular; anthers orange-brown,
<quantity id="4CA79B3E462BFFC2FBECCB85FB35500F" box="[1102,1196,1779,1804]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="12" pageNumber="113" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
long. Style with patch of glandular hairs at the base. Capsules 68 ×
<quantity id="4CA79B3E462BFFC2FD2FCA61FD745033" box="[653,749,1815,1840]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="12" pageNumber="113" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
. Seed 5 ×
<quantity id="4CA79B3E462BFFC2FCC2CA61FC595033" box="[864,960,1815,1840]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="12" pageNumber="113" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3456550462BFFC2FF1FCA4DFD9D507B" pageId="12" pageNumber="113" type="etymology">
<paragraph id="8BE036DB462BFFC2FF1FCA4DFD9D507B" blockId="12.[136,1452,1275,2092]" pageId="12" pageNumber="113">
<emphasis id="B92BEAC9462BFFC2FF1FCA4DFED15057" bold="true" box="[189,328,1851,1876]" pageId="12" pageNumber="113">Etymology:</emphasis>
—The specific epithet
<taxonomicName id="4C5F4D58462BFFC2FDF1CA4DFC8B5057" authorityName="H. M. Steyn" authorityYear="2017" box="[595,786,1851,1876]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="glandulopalmata">
<emphasis id="B92BEAC9462BFFC2FDF1CA4DFC8B5057" box="[595,786,1851,1876]" italics="true" pageId="12" pageNumber="113">glandulopalmata</emphasis>
</taxonomicName>
, meaning “glandular-palmate”, refers to the hand-shaped bracts, covered in glandular hairs.
</paragraph>
</subSubSection>
<subSubSection id="C3456550462BFFC0FF1FCAF5FB27541E" lastPageId="14" lastPageNumber="115" pageId="12" pageNumber="113" type="distribution">
<paragraph id="8BE036DB462BFFC3FF1FCAF5FC09564A" blockId="12.[136,1452,1275,2092]" lastBlockId="13.[136,1452,160,329]" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113">
<emphasis id="B92BEAC9462BFFC2FF1FCAF5FD04509F" bold="true" box="[189,669,1923,1948]" pageId="12" pageNumber="113">Distribution, ecology and phenology:—</emphasis>
<taxonomicName id="4C5F4D58462BFFC2FD3FCAF5FC76509F" authorityName="H. M. Steyn" authorityYear="2017" box="[669,1007,1923,1948]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="glandulopalmata">
<emphasis id="B92BEAC9462BFFC2FD3FCAF5FC76509F" box="[669,1007,1923,1948]" italics="true" pageId="12" pageNumber="113">Acanthopsis glandulopalmata</emphasis>
</taxonomicName>
is endemic to the
<collectingRegion id="499BF839462BFFC2FB6DCAF5FAE0509F" box="[1231,1401,1923,1948]" country="South Africa" name="Northern Cape" pageId="12" pageNumber="113">Northern Cape</collectingRegion>
and is known from three localities in an area to the north-west of Steinkopf with one outlier population north-west of Springbok (
<figureCitation id="13642A5E462BFFC2FEACCABDFEC850E7" box="[270,337,1995,2020]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="12" pageNumber="113">Fig. 1</figureCitation>
). It is centred in the core part of the Gariep Centre of Endemism (
<bibRefCitation id="EFCE4B2A462BFFC2FBEECABDFAFC50E7" author="Van Wyk, A. E. &amp; Smith, G. F." box="[1100,1381,1995,2020]" pageId="12" pageNumber="113" refId="ref13657" refString="Van Wyk, A. E. &amp; Smith, G. F. (2001) Regions of floristic endemism in southern Africa: a review with emphasis on succulents. Umdaus Press, Pretoria, 160 pp." type="book" year="2001">Van Wyk &amp; Smith 2001</bibRefCitation>
). This distribution falls within the Succulent Karoo Biome (
<bibRefCitation id="EFCE4B2A462BFFC2FD4BCA99FBB75F0B" author="Rutherford, M. C. &amp; Westfall, R. H." box="[745,1070,2031,2056]" pageId="12" pageNumber="113" refId="ref13271" refString="Rutherford, M. C. &amp; Westfall, R. H. (1994) Biomes of southern Africa: an objective categorization. [Memoir 63]. National Botanical Institute, Pretoria, 94 pp." type="book" year="1994">Rutherford &amp; Westfall 1994</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A462BFFC2FB99CA99FAB35F0B" author="Low, A. B. &amp; Rebelo, A. G." box="[1083,1322,2031,2056]" pageId="12" pageNumber="113" refId="ref12933" refString="Low, A. B., Rebelo, A. G. (Eds.) (1996) Vegetation of South Africa, Lesotho and Swaziland. Department of Environmental Affairs &amp; Tourism, Pretoria, 85 pp." type="book" year="1996">Low &amp; Rebelo 1996</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A462BFFC2FA95CA99FEDD5F2F" author="Mucina, L. &amp; Rutherford, M. C." pageId="12" pageNumber="113" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) in the Richtersveld Bioregion: Umdaus Mountain Succulent Shrubland, Anenous Plateau Shrubland and Lekkersing Succulent Shrubland (
<bibRefCitation id="EFCE4B2A462AFFC3FD8ECDD6FCC757BA" author="Mucina, L. &amp; Rutherford, M. C." box="[556,862,160,185]" pageId="13" pageNumber="114" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) where it receives predominantly winter rains with an average of
<quantity id="4CA79B3E462AFFC3FEA0CDB2FE0C57DE" box="[258,405,196,221]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="114" unit="mm" value="150.0" valueMax="200.0" valueMin="100.0">100200 mm</quantity>
per year. The specimens were collected on plains with quartzite pebbles in well-drained, sandy soils at elevations between
<quantity id="4CA79B3E462AFFC3FE63CD9EFDDD5602" box="[449,580,232,257]" metricMagnitude="2" metricUnit="m" metricValue="6.85" metricValueMax="7.0" metricValueMin="6.7" pageId="13" pageNumber="114" unit="m" value="685.0" valueMax="700.0" valueMin="670.0">
670
<specimenCount id="9D59FD52462AFFC3FE5BCD9EFDDD5602" box="[505,580,232,257]" count="700" pageId="13" pageNumber="114" type="generic">700 m</specimenCount>
</quantity>
.
<emphasis id="B92BEAC9462AFFC3FDF3CD9EFCAB5602" box="[593,818,232,257]" italics="true" pageId="13" pageNumber="114">Desmet &amp; Ellis 656</emphasis>
was collected at
<quantity id="4CA79B3E462AFFC3FC5BCD9EFBD85602" box="[1017,1089,232,257]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="114" unit="m" value="120.0">120 m</quantity>
in the Namaqualand Hardeveld Bioregion where it receives &lt;
<quantity id="4CA79B3E462AFFC3FE77CC7AFDAA5626" box="[469,563,268,293]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="114" unit="mm" value="100.0">100 mm</quantity>
of rain per annum, but probably more frequent fog that the inland populations. Specimens with flowers were collected between August and October.
</paragraph>
<caption id="DF206653462AFFC3FF2ACAD0FC7D5F00" ID-DOI="http://doi.org/10.5281/zenodo.13697374" ID-Zenodo-Dep="13697374" httpUri="https://zenodo.org/record/13697374/files/figure.png" pageId="13" pageNumber="114" startId="13.[136,229,1958,1979]" targetBox="[151,1436,353,1933]" targetPageId="13" targetType="figure">
<paragraph id="8BE036DB462AFFC3FF2ACAD0FC7D5F00" blockId="13.[136,1452,1958,2051]" pageId="13" pageNumber="114">
<emphasis id="B92BEAC9462AFFC3FF2ACAD0FF6750B8" bold="true" box="[136,254,1958,1979]" pageId="13" pageNumber="114">FIGURE 7.</emphasis>
<taxonomicName id="4C5F4D58462AFFC3FEA7CAD0FDBD50B8" authorityName="H. M. Steyn" authorityYear="2017" box="[261,548,1958,1979]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="13" pageNumber="114" phylum="Tracheophyta" rank="species" species="glandulopalmata">
<emphasis id="B92BEAC9462AFFC3FEA7CAD0FDBD50B8" box="[261,548,1958,1979]" italics="true" pageId="13" pageNumber="114">Acanthopsis glandulopalmata</emphasis>
</taxonomicName>
. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp;F. Bract. G. Calyx. H. Corolla (lobes lilac to purple; throat lemon-yellow). I. Glandular hairs at base of style. J. Capsule with seed. K. Seed. Vouchers: CI =
<emphasis id="B92BEAC9462AFFC3FB58CABCFAFD50DC" box="[1274,1380,1994,2015]" italics="true" pageId="13" pageNumber="114">Steyn 2125</emphasis>
(PRE), J &amp; K =
<emphasis id="B92BEAC9462AFFC3FF79CA98FEDC5F00" box="[219,325,2030,2051]" italics="true" pageId="13" pageNumber="114">Steyn 1889</emphasis>
(PRE). Photographed by: M. Koekemoer (A) and H.M. Steyn (BK).
</paragraph>
</caption>
<paragraph id="8BE036DB4629FFC0FF1FCDD6FC615692" blockId="14.[136,1452,160,797]" pageId="14" pageNumber="115">
<emphasis id="B92BEAC94629FFC0FF1FCDD6FE2A57BA" bold="true" box="[189,435,160,185]" pageId="14" pageNumber="115">Conservation status:</emphasis>
<taxonomicName id="4C5F4D584629FFC0FE6DCDD6FC8557BA" authorityName="H. M. Steyn" authorityYear="2017" box="[463,796,160,185]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="species" species="glandulopalmata">
<emphasis id="B92BEAC94629FFC0FE6DCDD6FC8557BA" box="[463,796,160,185]" italics="true" pageId="14" pageNumber="115">Acanthopsis glandulopalmata</emphasis>
</taxonomicName>
is known from four widely scattered subpopulations (EOO
<quantity id="4CA79B3E4629FFC0FF2ACDB2FF6A57DE" box="[136,243,196,221]" metricMagnitude="6" metricUnit="m" metricValue="1.954" pageId="14" pageNumber="115" unit="km" value="1954.0">1954 km</quantity>
²). The threats affecting this species, particularly overgrazing, is likely to affect most areas with suitable habitat, and therefore it is threatened across its range. The population is likely to be declining at all known locations. It is therefore assessed as Endangered based on precautionary principles (Endangered B1ab(iii)) according to the IUCN Red List Category and Criteria (
<bibRefCitation id="EFCE4B2A4629FFC0FE4DCC46FDEF564A" author="IUCN" box="[495,630,304,329]" pageId="14" pageNumber="115" refId="ref12795" refString="IUCN (2012) IUCN Red List categories and criteria version 3.1. IUCN Species Survival Commission, Gland, Switzerland and Cambridge, UK, iv + 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
) (L. von Staden, pers. comm.), but may need to be downlisted in future if further field surveys indicate that there are more than four subpopulations. Further field surveys are needed to gain a better understanding of the population structure and abundance of this species.
</paragraph>
<paragraph id="8BE036DB4629FFC0FF1FCCEAFBAF558E" blockId="14.[136,1452,160,797]" pageId="14" pageNumber="115">
<materialsCitation id="3B373C864629FFC0FF1FCCEAFB6656DA" collectingDate="1985-09-15" collectionCode="CA, NMB, PRE, PRU" collectorName="Zietsman" country="South Africa" elevation="550" location="Tussen Ploegberg en Stinkfonteinberge" pageId="14" pageNumber="115" specimenCount="2" stateProvince="Northern Cape">
<emphasis id="B92BEAC94629FFC0FF1FCCEAFD7256B6" bold="true" box="[189,747,412,437]" pageId="14" pageNumber="115">
<typeStatus id="54E488794629FFC0FF1FCCEAFDD156B6" box="[189,584,412,437]" isOtherTypeStatus="true" pageId="14" pageNumber="115">Additional specimens examined</typeStatus>
(
<typeStatus id="54E488794629FFC0FDFDCCEAFD4756B6" box="[607,734,412,437]" pageId="14" pageNumber="115" type="paratype">Paratypes</typeStatus>
):
</emphasis>
<collectingCountry id="F348764B4629FFC0FCA5CCEAFC4556B6" box="[775,988,412,437]" name="South Africa" pageId="14" pageNumber="115">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="499BF8394629FFC0FC4DCCEAFB3856B6" box="[1007,1185,412,437]" country="South Africa" name="Northern Cape" pageId="14" pageNumber="115">Northern Cape</collectingRegion>
:
<location id="8E8060004629FFC0FB11CCEAFECA56DA" LSID="urn:lsid:plazi:treatment:03F687CD462BFFC0FF2ACE89FB27541E:8E8060004629FFC0FB11CCEAFECA56DA" country="South Africa" name="Tussen Ploegberg en Stinkfonteinberge" pageId="14" pageNumber="115" stateProvince="Northern Cape">Tussen Ploegberg en Stinkfonteinberge</location>
,
<quantity id="4CA79B3E4629FFC0FEFECCB6FE3B56DA" box="[348,418,448,473]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="115" unit="m" value="550.0">
<elevation id="0072D1E84629FFC0FEFECCB6FE3B56DA" box="[348,418,448,473]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="115" unit="m" value="550.0">550 m</elevation>
</quantity>
, (2817
<collectionCode id="ED4EAE1E4629FFC0FE4DCCB6FD8156DA" box="[495,536,448,473]" country="USA" name="Chicago Academy of Sciences" pageId="14" pageNumber="115" type="Museum">CA</collectionCode>
),
<date id="FFE1101B4629FFC0FD8BCCB6FC9956DA" box="[553,768,448,473]" pageId="14" pageNumber="115" value="1985-09-15">
<collectingDate id="EFA5E9F34629FFC0FD8BCCB6FC9956DA" box="[553,768,448,473]" pageId="14" pageNumber="115" value="1985-09-15">15 September 1985</collectingDate>
</date>
,
<emphasis id="B92BEAC94629FFC0FCABCCB6FBBD56DA" box="[777,1060,448,473]" italics="true" pageId="14" pageNumber="115">
<collectorName id="26AA530D4629FFC0FCABCCB6FCE956DA" box="[777,880,448,473]" pageId="14" pageNumber="115">Zietsman</collectorName>
&amp;
<collectorName id="26AA530D4629FFC0FC2DCCB6FC6F56DA" box="[911,1014,448,473]" pageId="14" pageNumber="115">Zietsman</collectorName>
773
</emphasis>
(
<collectionCode id="ED4EAE1E4629FFC0FB92CCB6FBED56DA" box="[1072,1140,448,473]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="14" pageNumber="115">NMB</collectionCode>
,
<collectionCode id="ED4EAE1E4629FFC0FBDFCCB6FB2A56DA" box="[1149,1203,448,473]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="14" pageNumber="115" type="Herbarium">PRE</collectionCode>
,
<collectionCode id="ED4EAE1E4629FFC0FB1ECCB6FB6E56DA" box="[1212,1271,448,473]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14594" lsid="urn:lsid:biocol.org:col:14594" name="University of Pretoria" pageId="14" pageNumber="115" type="Herbarium">PRU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C864629FFC0FAAACCB6FA3E56DA" box="[1288,1447,448,473]" country="South Africa" location="Namaqualand" pageId="14" pageNumber="115" specimenCount="1">
<location id="8E8060004629FFC0FAAACCB6FA3E56DA" LSID="urn:lsid:plazi:treatment:03F687CD462BFFC0FF2ACE89FB27541E:8E8060004629FFC0FAAACCB6FA3E56DA" box="[1288,1447,448,473]" country="South Africa" name="Namaqualand" pageId="14" pageNumber="115">Namaqualand</location>
</materialsCitation>
; ±
<materialsCitation id="3B373C864629FFC0FF01CC92FA3E56FE" box="[163,1447,484,509]" collectingDate="1956-10-08" collectionCode="BA, PRE" collectorName="Leistner" country="South Africa" elevation="671" location="Hangbal River" pageId="14" pageNumber="115" specimenCount="1">
<locationDeviation id="9A522FF94629FFC0FF01CC92FE5C56FE" box="[163,453,484,509]" location="Steinkopf" pageId="14" pageNumber="115">
<quantity id="4CA79B3E4629FFC0FF01CC92FF6956FE" box="[163,240,484,509]" metricMagnitude="4" metricUnit="m" metricValue="2.5749504" pageId="14" pageNumber="115" unit="mi" value="16.0">16 mi.</quantity>
NW of Steinkopf
</locationDeviation>
(
<location id="8E8060004629FFC0FE7BCC92FD1E56FE" LSID="urn:lsid:plazi:treatment:03F687CD462BFFC0FF2ACE89FB27541E:8E8060004629FFC0FE7BCC92FD1E56FE" box="[473,647,484,509]" country="South Africa" name="Hangbal River" pageId="14" pageNumber="115">Hangbal River</location>
),
<quantity id="4CA79B3E4629FFC0FD3DCC92FD6D56FE" box="[671,756,484,509]" metricMagnitude="2" metricUnit="m" metricValue="6.7056" pageId="14" pageNumber="115" unit="ft" value="2200.0">
<elevation id="0072D1E84629FFC0FD3DCC92FD6D56FE" box="[671,756,484,509]" metricMagnitude="2" metricUnit="m" metricValue="6.7056" pageId="14" pageNumber="115" unit="ft" value="2200.0">2200 ft</elevation>
</quantity>
[
<quantity id="4CA79B3E4629FFC0FCA5CC92FCC056FE" box="[775,857,484,509]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="14" pageNumber="115" unit="m" value="670.0">
<elevation id="0072D1E84629FFC0FCA5CC92FCC056FE" box="[775,857,484,509]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="14" pageNumber="115" unit="m" value="670.0">670 m</elevation>
</quantity>
], (2917
<collectionCode id="ED4EAE1E4629FFC0FC17CC92FC4756FE" box="[949,990,484,509]" country="Argentina" lsid="urn:lsid:biocol.org:col:15002" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="14" pageNumber="115" type="Herbarium">BA</collectionCode>
),
<date id="FFE1101B4629FFC0FC54CC92FB2956FE" box="[1014,1200,484,509]" pageId="14" pageNumber="115" value="1956-10-08">
<collectingDate id="EFA5E9F34629FFC0FC54CC92FB2956FE" box="[1014,1200,484,509]" pageId="14" pageNumber="115" value="1956-10-08">8 October 1956</collectingDate>
</date>
,
<emphasis id="B92BEAC94629FFC0FB62CC92FACB56FE" box="[1216,1362,484,509]" italics="true" pageId="14" pageNumber="115">
<collectorName id="26AA530D4629FFC0FB62CC92FA8556FE" box="[1216,1308,484,509]" pageId="14" pageNumber="115">Leistner</collectorName>
797
</emphasis>
(
<collectionCode id="ED4EAE1E4629FFC0FAC4CC92FA0656FE" box="[1382,1439,484,509]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="14" pageNumber="115" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C864629FFC0FF2ACF7EFE745546" collectingDate="2012-08-25" collectionCode="BA, PRE" collectorName="Steyn" country="South Africa" county="Namaqualand District" elevation="697" location="Kosies" municipality="Anenous" pageId="14" pageNumber="115" specimenCount="1">
<collectingCounty id="62814E574629FFC0FF2ACF7EFE1B5522" box="[136,386,520,545]" pageId="14" pageNumber="115">Namaqualand District</collectingCounty>
. On track between
<collectingMunicipality id="6B84ACA14629FFC0FDF7CF7EFD205522" box="[597,697,520,545]" pageId="14" pageNumber="115">Anenous</collectingMunicipality>
/
<location id="8E8060004629FFC0FD60CF7EFC945522" LSID="urn:lsid:plazi:treatment:03F687CD462BFFC0FF2ACE89FB27541E:8E8060004629FFC0FD60CF7EFC945522" box="[706,781,520,545]" country="South Africa" county="Namaqualand District" municipality="Anenous" name="Kosies" pageId="14" pageNumber="115">Kosies</location>
to
<location id="8E8060004629FFC0FC92CF7EFB2B5522" LSID="urn:lsid:plazi:treatment:03F687CD462BFFC0FF2ACE89FB27541E:8E8060004629FFC0FC92CF7EFB2B5522" box="[816,1202,520,545]" country="South Africa" county="Namaqualand District" municipality="Anenous" name="Umdaus in Wyepoort River Valley" pageId="14" pageNumber="115">Umdaus in Wyepoort River Valley</location>
,
<quantity id="4CA79B3E4629FFC0FB1ECF7EFA9C5522" box="[1212,1285,520,545]" metricMagnitude="2" metricUnit="m" metricValue="6.97" pageId="14" pageNumber="115" unit="m" value="697.0">
<elevation id="0072D1E84629FFC0FB1ECF7EFA9C5522" box="[1212,1285,520,545]" metricMagnitude="2" metricUnit="m" metricValue="6.97" pageId="14" pageNumber="115" unit="m" value="697.0">697 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E4629FFC0FAF6CF7EFAE45522" box="[1364,1405,520,545]" country="Argentina" lsid="urn:lsid:biocol.org:col:15002" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="14" pageNumber="115" type="Herbarium">BA</collectionCode>
),
<date id="FFE1101B4629FFC0FA2DCF7EFE835546" pageId="14" pageNumber="115" value="2012-08-25">
<collectingDate id="EFA5E9F34629FFC0FA2DCF7EFE835546" pageId="14" pageNumber="115" value="2012-08-25">25 August 2012</collectingDate>
</date>
,
<emphasis id="B92BEAC94629FFC0FE81CF5AFE075546" box="[291,414,556,581]" italics="true" pageId="14" pageNumber="115">
<collectorName id="26AA530D4629FFC0FE81CF5AFEF95546" box="[291,352,556,581]" pageId="14" pageNumber="115">Steyn</collectorName>
1889
</emphasis>
(
<collectionCode id="ED4EAE1E4629FFC0FE09CF5AFE7D5546" box="[427,484,556,581]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="14" pageNumber="115" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C864629FFC0FE55CF5AFE88556A" collectingDate="2015-10-09" collectionCode="BA, PRE" collectorName="Steyn" country="South Africa" location="Namaqualand District. Wyepoort River Valley" pageId="14" pageNumber="115" specimenCount="1">
<location id="8E8060004629FFC0FE55CF5AFC615546" LSID="urn:lsid:plazi:treatment:03F687CD462BFFC0FF2ACE89FB27541E:8E8060004629FFC0FE55CF5AFC615546" box="[503,1016,556,581]" country="South Africa" name="Namaqualand District. Wyepoort River Valley" pageId="14" pageNumber="115">Namaqualand District. Wyepoort River Valley</location>
, 695, (2917
<collectionCode id="ED4EAE1E4629FFC0FBDECF5AFB3C5546" box="[1148,1189,556,581]" country="Argentina" lsid="urn:lsid:biocol.org:col:15002" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="14" pageNumber="115" type="Herbarium">BA</collectionCode>
),
<date id="FFE1101B4629FFC0FB15CF5AFAFC5546" box="[1207,1381,556,581]" pageId="14" pageNumber="115" value="2015-10-09">
<collectingDate id="EFA5E9F34629FFC0FB15CF5AFAFC5546" box="[1207,1381,556,581]" pageId="14" pageNumber="115" value="2015-10-09">9 October 2015</collectingDate>
</date>
,
<emphasis id="B92BEAC94629FFC0FACDCF5AFF59556A" italics="true" pageId="14" pageNumber="115">
<collectorName id="26AA530D4629FFC0FACDCF5AFA355546" box="[1391,1452,556,581]" pageId="14" pageNumber="115">Steyn</collectorName>
2125
</emphasis>
(
<collectionCode id="ED4EAE1E4629FFC0FF6DCF26FE91556A" box="[207,264,592,617]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="14" pageNumber="115" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C864629FFC0FEBFCF26FBAB558E" collectingDate="1997-06-21" collectionCode="CB, NBG" collectorName="Desmet &amp; Ellis" country="South Africa" county="Nuttabooi Kleinzee" elevation="120" location="Kleinzee" municipality="Nuttabooi" pageId="14" pageNumber="115" specimenCount="1">
<collectingCounty id="62814E574629FFC0FEBFCF26FE63556A" box="[285,506,592,617]" pageId="14" pageNumber="115">Nuttabooi Kleinzee</collectingCounty>
,
<collectingMunicipality id="6B84ACA14629FFC0FDA4CF26FDEF556A" box="[518,630,592,617]" pageId="14" pageNumber="115">Nuttabooi</collectingMunicipality>
mining area approximately
<quantity id="4CA79B3E4629FFC0FC11CF26FC63556A" box="[947,1018,592,617]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="115" unit="km" value="40.0">40 km</quantity>
from
<location id="8E8060004629FFC0FB9DCF26FB3A556A" LSID="urn:lsid:plazi:treatment:03F687CD462BFFC0FF2ACE89FB27541E:8E8060004629FFC0FB9DCF26FB3A556A" box="[1087,1187,592,617]" country="South Africa" county="Nuttabooi Kleinzee" municipality="Nuttabooi" name="Kleinzee" pageId="14" pageNumber="115">Kleinzee</location>
on the
<location id="8E8060004629FFC0FB55CF26FAF5556A" LSID="urn:lsid:plazi:treatment:03F687CD462BFFC0FF2ACE89FB27541E:8E8060004629FFC0FB55CF26FAF5556A" box="[1271,1388,592,617]" country="South Africa" county="Nuttabooi Kleinzee" municipality="Nuttabooi" name="Springbok" pageId="14" pageNumber="115">Springbok</location>
road, south bank of
<location id="8E8060004629FFC0FE85CF02FEE5558E" LSID="urn:lsid:plazi:treatment:03F687CD462BFFC0FF2ACE89FB27541E:8E8060004629FFC0FE85CF02FEE5558E" box="[295,380,628,653]" country="South Africa" county="Nuttabooi Kleinzee" municipality="Nuttabooi" name="Buffels" pageId="14" pageNumber="115">Buffels</location>
,
<quantity id="4CA79B3E4629FFC0FE25CF02FE49558E" box="[391,464,628,653]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="115" unit="m" value="120.0">
<elevation id="0072D1E84629FFC0FE25CF02FE49558E" box="[391,464,628,653]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="115" unit="m" value="120.0">120 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E4629FFC0FD82CF02FDDF558E" box="[544,582,628,653]" country="Australia" name="The CB Rhizobium Collection" pageId="14" pageNumber="115">CB</collectionCode>
),
<date id="FFE1101B4629FFC0FDF8CF02FD6A558E" box="[602,755,628,653]" pageId="14" pageNumber="115" value="1997-06-21">
<collectingDate id="EFA5E9F34629FFC0FDF8CF02FD6A558E" box="[602,755,628,653]" pageId="14" pageNumber="115" value="1997-06-21">21 June 1997</collectingDate>
</date>
,
<emphasis id="B92BEAC94629FFC0FD5CCF02FC43558E" box="[766,986,628,653]" italics="true" pageId="14" pageNumber="115">
<collectorName id="26AA530D4629FFC0FD5CCF02FCCB558E" box="[766,850,628,653]" pageId="14" pageNumber="115">Desmet</collectorName>
&amp;
<collectorName id="26AA530D4629FFC0FCD7CF02FC31558E" box="[885,936,628,653]" pageId="14" pageNumber="115">Ellis</collectorName>
956
</emphasis>
(
<collectionCode id="ED4EAE1E4629FFC0FC4BCF02FBB3558E" box="[1001,1066,628,653]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="14" pageNumber="115" type="Herbarium">NBG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE036DB4629FFC0FF1FCFEEFB27541E" blockId="14.[136,1452,160,797]" pageId="14" pageNumber="115">
<emphasis id="B92BEAC94629FFC0FF1FCFEEFE0555B2" box="[189,412,664,689]" italics="true" pageId="14" pageNumber="115">Desmet &amp; Ellis 956</emphasis>
has greyish leaves covered with densely packed appressed hairs. It was collected at a much lower elevation and in a different bioregion than the rest of the specimens, and is here included in
<taxonomicName id="4C5F4D584629FFC0FB6FCFCAFA3255D6" authorityName="H. M. Steyn" authorityYear="2017" box="[1229,1451,700,725]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="species" species="glandulopalmata">
<emphasis id="B92BEAC94629FFC0FB6FCFCAFA3255D6" box="[1229,1451,700,725]" italics="true" pageId="14" pageNumber="115">A. glandulopalmata</emphasis>
</taxonomicName>
based on the semi-dense inflorescences with palmate bracts. However, the inflorescence is very narrow and no glandular hairs are visible on the bracts, as the inflorescences are old (probably from the previous season).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,306 @@
<document id="59C4807E644B93C6A4653C8E49826221" ID-DOI="10.11646/phytotaxa.324.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13697360" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725568056204" checkinUser="felipe" docAuthor="Steyn, Hester M. &amp; Van Wyk, Abraham E." docDate="2017" docId="03F687CD462DFFC2FF2AC8F3FBF954D3" docLanguage="en" docName="phytotaxa.324.2.1.pdf" docOrigin="Phytotaxa 324 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.324.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Acanthopsis nitida H. M. Steyn 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="113" masterDocId="FFCFFFB54627FFCEFFA2CD76FF995703" masterDocTitle="Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts" masterLastPageNumber="121" masterPageNumber="101" pageNumber="111" updateTime="1725570993962" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1C598788F9FCFB32B15149923A543881" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="478B2E1E53B7D8DCFD21D6D1E9DAFF3B">
<mods:title id="B2F94A77B83BF338418AB975CC6E139C">Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts</mods:title>
</mods:titleInfo>
<mods:name id="7E26C6F6B837A4DD84A9642EA8211D38" type="personal">
<mods:role id="A18419F15B0C3077E365DA14787106FF">
<mods:roleTerm id="583D2F248E8D504E79C5C27A3362C930">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="67BBB0A3FB8513CB7C574F299CD89339">Steyn, Hester M.</mods:namePart>
<mods:affiliation id="2831BE680915409F181A9BBE4171FBBF">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:name id="97A52CDF4947B87B387A8A22E64D9C72" type="personal">
<mods:role id="AF4053FEC157D783E556E55C3776E65A">
<mods:roleTerm id="A8E351C780975AE58514C12D42DA0ABE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E1486456C0EA223BF14BA23D5A078BE1">Van Wyk, Abraham E.</mods:namePart>
<mods:affiliation id="8929A8C75C4109A6F67FF7CC49216826">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="66351D9CBA922DD089EBD5186DBFD962">text</mods:typeOfResource>
<mods:relatedItem id="7AD8406F7C35262DBA74A60A1C9B5166" type="host">
<mods:titleInfo id="8A0D00160CA325452422CFCAF15CB40F">
<mods:title id="12C30591F9F379C877C3CB4017D0CE5C">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="8CA0875265AD9EBF7AA04CC55A62B164">
<mods:date id="660DF3C623B2E423BC5F7968C5A4E729">2017</mods:date>
<mods:detail id="81E4C12F2465BC9B2378FF94FD3E6E8E" type="pubDate">
<mods:number id="B9349A33974EE83B5ADC260BF2960B0F">2017-10-10</mods:number>
</mods:detail>
<mods:detail id="33E7EFB0CBDA077D1DB3896C89F4BBF2" type="volume">
<mods:number id="413DB1B335FF9CC8584E4DC4DF121DFC">324</mods:number>
</mods:detail>
<mods:detail id="B77392AF24E4265117BAFEDC46461200" type="issue">
<mods:number id="C804C14AA9CAC5144B86374DDBBC41C7">2</mods:number>
</mods:detail>
<mods:extent id="A451765BE5E0D9406FAB6B836E6C0421" unit="page">
<mods:start id="44B10DF375C0174E4EB7E32FD8C8B025">101</mods:start>
<mods:end id="2229A38CF9AFDC9A2E9000EB6546A6EE">121</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="B94D52B8C9F9F549E48374F7B14B3A57">
<mods:url id="3B104EEBBEEF98B37F77DD659D141E8F">http://dx.doi.org/10.11646/phytotaxa.324.2.1</mods:url>
</mods:location>
<mods:classification id="E681FE30945B1CBA7164A3093B9F437E">journal article</mods:classification>
<mods:identifier id="24B723F4D887C3B46378980C61BACA52" type="DOI">10.11646/phytotaxa.324.2.1</mods:identifier>
<mods:identifier id="A675F239871B13DC2A81FA0267BB4AB2" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="713C88C1686C5A61A390FECCED7AC273" type="Zenodo-Dep">13697360</mods:identifier>
</mods:mods>
<treatment id="03F687CD462DFFC2FF2AC8F3FBF954D3" ID-DOI="http://doi.org/10.5281/zenodo.13697382" ID-Zenodo-Dep="13697382" LSID="urn:lsid:plazi:treatment:03F687CD462DFFC2FF2AC8F3FBF954D3" httpUri="http://treatment.plazi.org/id/03F687CD462DFFC2FF2AC8F3FBF954D3" lastPageId="12" lastPageNumber="113" pageId="10" pageNumber="111">
<subSubSection id="C3456550462DFFC4FF2AC8F3FCBE529D" box="[136,807,1411,1438]" pageId="10" pageNumber="111" type="nomenclature">
<paragraph id="8BE036DB462DFFC4FF2AC8F3FCBE529D" blockId="10.[136,1452,1411,1617]" box="[136,807,1411,1438]" pageId="10" pageNumber="111">
<heading id="D0A881B7462DFFC4FF2AC8F3FCBE529D" box="[136,807,1411,1438]" fontSize="11" level="2" pageId="10" pageNumber="111" reason="4">
<emphasis id="B92BEAC9462DFFC4FF2AC8F3FF04529D" bold="true" box="[136,157,1412,1438]" pageId="10" pageNumber="111">3.</emphasis>
<taxonomicName id="4C5F4D58462DFFC4FF01C8F5FE66529D" ID-CoL="8ZLG" authority="H.M.Steyn" authorityName="H. M. Steyn" authorityYear="2017" box="[163,511,1411,1438]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="111" phylum="Tracheophyta" rank="species" species="nitida" status="sp. nov.">
<emphasis id="B92BEAC9462DFFC4FF01C8F5FEE2529D" bold="true" box="[163,379,1411,1438]" italics="true" pageId="10" pageNumber="111">Acanthopsis nitida</emphasis>
H.M.Steyn
</taxonomicName>
,
<taxonomicNameLabel id="A21857B2462DFFC4FDA8C8F3FDF8529D" box="[522,609,1413,1438]" pageId="10" pageNumber="111" rank="species">
<emphasis id="B92BEAC9462DFFC4FDA8C8F3FDC0529D" box="[522,601,1413,1438]" italics="true" pageId="10" pageNumber="111">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="13642A5E462DFFC4FDD2C8F3FD24529D" box="[624,701,1413,1438]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="10" pageNumber="111">Figs. 1</figureCitation>
,
<figureCitation id="13642A5E462DFFC4FD69C8F3FD75529D" box="[715,748,1413,1438]" captionStart="FIGURE 3" captionStartId="6.[136,229,1911,1932]" captionTargetBox="[151,1436,190,1886]" captionTargetId="figure-15@6.[151,1436,190,1886]" captionTargetPageId="6" captionText="FIGURE 3. Comparison of floral parts, associated bracts and bracteoles (from left to right): bracts (2), bracteoles (2), calyx (4 sepals: dorsal, ventral and 2 lateral ones) and corolla with anthers.A. A. glabra, B. A. scullyi, C. A. nitida, D. A. glandulopalmata, E. A. insueta, F. A. carduifolia. Note: the bract on the left is from lower down in the same inflorescence. Scale bar = 10 mm. Vouchers:A = Steyn 1893 (PRE), B = Steyn 1911 (PRE), C = Steyn 1884 (PRE), D = Steyn 1890 (PRE), E = Steyn 1805 (PRE), F = Koekemoer 4620 (PRE)." figureDoi="http://doi.org/10.5281/zenodo.13697366" httpUri="https://zenodo.org/record/13697366/files/figure.png" pageId="10" pageNumber="111">3C</figureCitation>
&amp;
<figureCitation id="13642A5E462DFFC4FCB2C8F3FC86529D" box="[784,799,1413,1438]" captionStart="FIGURE 6" captionStartId="11.[136,229,1958,1979]" captionTargetBox="[151,1436,353,1933]" captionTargetId="figure-89@11.[151,1436,353,1933]" captionTargetPageId="11" captionText="FIGURE 6. Acanthopsis nitida. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp; F. Bract. G. Calyx. H. Corolla (lobes lilac to purple; throat lemon-yellow). I. Glandular hairs at base of style. J. Capsule. K. Capsule with seed. Vouchers: CI = Steyn 1884 (PRE), J = Burgoyne 1514 (PRE), K = Steyn 2129 (PRE). Photographed by: M. Koekemoer (A &amp; B) and H.M. Steyn (CK)." figureDoi="http://doi.org/10.5281/zenodo.13697372" httpUri="https://zenodo.org/record/13697372/files/figure.png" pageId="10" pageNumber="111">6</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3456550462DFFC4FF2AC8DAFD1D52E6" pageId="10" pageNumber="111" type="materials_examined">
<paragraph id="8BE036DB462DFFC4FF2AC8DAFD1D52E6" blockId="10.[136,1452,1411,1617]" pageId="10" pageNumber="111">
<emphasis id="B92BEAC9462DFFC4FF2AC8DAFF5B52C2" bold="true" box="[136,194,1452,1473]" pageId="10" pageNumber="111">Type:</emphasis>
<collectingCountry id="F348764B462DFFC4FF79C8DAFE1552C2" box="[219,396,1452,1473]" name="South Africa" pageId="10" pageNumber="111">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="499BF839462DFFC4FE35C8DAFDB352C2" box="[407,554,1452,1473]" country="South Africa" name="Northern Cape" pageId="10" pageNumber="111">Northern Cape</collectingRegion>
: Namaqualand District. Anenous Pass, S of road,
<quantity id="4CA79B3E462DFFC4FBB7C8DAFBCF52C2" box="[1045,1110,1452,1473]" metricMagnitude="2" metricUnit="m" metricValue="7.09" pageId="10" pageNumber="111" unit="m" value="709.0">709 m</quantity>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FB39C8DAFB2752C2" box="[1179,1214,1452,1473]" country="Argentina" lsid="urn:lsid:biocol.org:col:15002" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="10" pageNumber="111" type="Herbarium">BA</collectionCode>
),
<date id="FFE1101B462DFFC4FB73C8DAFAF552C2" box="[1233,1388,1452,1473]" pageId="10" pageNumber="111" value="2015-10-09">9 October 2015</date>
,
<emphasis id="B92BEAC9462DFFC4FAD5C8DAFF6D52E6" italics="true" pageId="10" pageNumber="111">Steyn 2129,</emphasis>
(
<typeStatus id="54E48879462DFFC4FEA2C8A6FECF52E6" box="[256,342,1488,1509]" pageId="10" pageNumber="111" type="holotype">holotype</typeStatus>
<collectionCode id="ED4EAE1E462DFFC4FEF9C8A6FE1352E6" box="[347,394,1488,1509]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
0866187!,
<typeStatus id="54E48879462DFFC4FE4DC8A6FDAD52E6" box="[495,564,1488,1509]" pageId="10" pageNumber="111" type="isotype">isotype</typeStatus>
<collectionCode id="ED4EAE1E462DFFC4FD99C8A6FDEF52E6" box="[571,630,1488,1509]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="10" pageNumber="111" type="Herbarium">NBG</collectionCode>
!).
</paragraph>
</subSubSection>
<subSubSection id="C3456550462DFFC5FF2AC882FF65564A" lastPageId="11" lastPageNumber="112" pageId="10" pageNumber="111" type="diagnosis">
<paragraph id="8BE036DB462DFFC4FF2AC882FD855152" blockId="10.[136,1452,1411,1617]" pageId="10" pageNumber="111">
<emphasis id="B92BEAC9462DFFC4FF2AC882FF6A510A" bold="true" box="[136,243,1524,1545]" pageId="10" pageNumber="111">Diagnosis:</emphasis>
<taxonomicName id="4C5F4D58462DFFC4FEA9C882FE5B510A" authorityName="H. M. Steyn" authorityYear="2017" box="[267,450,1524,1545]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="111" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="B92BEAC9462DFFC4FEA9C882FE5B510A" box="[267,450,1524,1545]" italics="true" pageId="10" pageNumber="111">Acanthopsis nitida</emphasis>
</taxonomicName>
differs from all other members of the group with semi-dense spikes and 5-fid bracts in its glossy appearance (
<emphasis id="B92BEAC9462DFFC4FE95CB6EFECF512E" box="[311,342,1560,1581]" italics="true" pageId="10" pageNumber="111">not</emphasis>
matt), yellow-green puberulent to sub-glabrous leaves (
<emphasis id="B92BEAC9462DFFC4FCD0CB6EFC08512E" box="[882,913,1560,1581]" italics="true" pageId="10" pageNumber="111">not</emphasis>
grey or green strigose to villose leaves) and in having a glabrous (
<emphasis id="B92BEAC9462DFFC4FEBCCB4AFEA45152" box="[286,317,1596,1617]" italics="true" pageId="10" pageNumber="111">not</emphasis>
variously hairy) calyx.
</paragraph>
<paragraph id="8BE036DB462DFFC5FF2ACB0CFF65564A" blockId="10.[136,1452,1658,2079]" lastBlockId="11.[136,1452,160,329]" lastPageId="11" lastPageNumber="112" pageId="10" pageNumber="111">
Perennial, compact subshrub,
<quantity id="4CA79B3E462DFFC4FE42CB0CFDDA5190" box="[480,579,1658,1683]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="10" pageNumber="111" unit="cm" value="8.5" valueMax="10.0" valueMin="7.0">710 cm</quantity>
tall with very short internodes.
<emphasis id="B92BEAC9462DFFC4FC14CB0CFB9D5190" box="[950,1028,1658,1683]" italics="true" pageId="10" pageNumber="111">Leaves</emphasis>
sessile, base petiole-like with spines
<quantity id="4CA79B3E462DFFC4FF2ACBE8FF7151B4" box="[136,232,1694,1719]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="10" pageNumber="111" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">37 mm</quantity>
long; lamina oblanceolate, 4060(90) ×
<quantity id="4CA79B3E462DFFC4FD6DCBE8FCA651B4" box="[719,831,1694,1719]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="10" pageNumber="111" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">812 mm</quantity>
, often with yellow-green appearance, puberulent with strigose to spreading short hairs, also with subsessile glandular hairs; apex acute, base attenuate, margin undulate, dentate-spinose, spines rigid, 35(8) mm long, yellow to brown. Inflorescences semi-dense, cylindrical, subsessile,
<quantity id="4CA79B3E462DFFC4FF2ACA7CFE995020" box="[136,256,1802,1827]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="10" pageNumber="111" unit="mm" value="50.0" valueMax="55.0" valueMin="45.0">4555 mm</quantity>
long, (8)
<quantity id="4CA79B3E462DFFC4FED3CA7CFE735020" box="[369,490,1802,1827]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="10" pageNumber="111" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">1013 mm</quantity>
in diameter. Bracts broadly obovate to wedge-shaped, (15)
<quantity id="4CA79B3E462DFFC4FB22CA7CFB655020" box="[1152,1276,1802,1827]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="10" pageNumber="111" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">2030 mm</quantity>
long (including spines), lamina
<quantity id="4CA79B3E462DFFC4FE9ECA58FE3E5044" box="[316,423,1838,1863]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="10" pageNumber="111" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">811 mm</quantity>
long, base obtuse to attenuate; lamina and spines glabrous to puberulent with short white, spreading, hairs and isolated subsessile glandular hairs abaxially, silky-hairy adaxially; middle and upper bracts ending in 5(7) primary spines; central primary spine often compound with one pair of marginal secondary spines, often simple; lateral primary spines very often with 1 long basal secondary spine (shorter than primary spine) on side facing central primary spine and a short marginal secondary spine on opposite side; spines divergent in flower, spreading to recurved in fruit. Bracteoles linear, 8(12) mm long, silky-hairy adaxially. Calyx brown-tipped with dorsal sepal ovate, acuminate, 1315(16) mm long, glabrous abaxially, silky-hairy adaxially, 79(11)-veined; ventral sepal ovate, 12 13(15) mm long, glabrous abaxially, silky-hairy adaxially, 57(9)-veined; lateral sepals 89(10) mm long, glabrous abaxially, silky-hairy adaxially, especially towards base. Corolla lilac to purple with lemon-yellow throat, 2527 ×
<quantity id="4CA79B3E462CFFC5FF2ACD9EFE9D5602" box="[136,260,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="11" pageNumber="112" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">1012 mm</quantity>
, tube 8(9) mm long, central lobe as wide as long, constricted at the base, emarginate. Filaments
<quantity id="4CA79B3E462CFFC5FAEDCD9EFA355602" box="[1359,1452,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="11" pageNumber="112" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
long, glandular; anthers beige,
<quantity id="4CA79B3E462CFFC5FE40CC7AFDBA5626" box="[482,547,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="112" unit="mm" value="3.0">3 mm</quantity>
long. Style covered with glandular hairs at the base. Capsules 89 ×
<quantity id="4CA79B3E462CFFC5FA85CC7AFAF35626" box="[1319,1386,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="112" unit="mm" value="3.0">3 mm</quantity>
. Seed 3 ×
<quantity id="4CA79B3E462CFFC5FF16CC46FF61564A" box="[180,248,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="112" unit="mm" value="3.0">3 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF206653462CFFC5FF2ACAD0FB505F00" ID-DOI="http://doi.org/10.5281/zenodo.13697372" ID-Zenodo-Dep="13697372" httpUri="https://zenodo.org/record/13697372/files/figure.png" pageId="11" pageNumber="112" startId="11.[136,229,1958,1979]" targetBox="[151,1436,353,1933]" targetPageId="11" targetType="figure">
<paragraph id="8BE036DB462CFFC5FF2ACAD0FB505F00" blockId="11.[136,1452,1958,2051]" pageId="11" pageNumber="112">
<emphasis id="B92BEAC9462CFFC5FF2ACAD0FF6650B8" bold="true" box="[136,255,1958,1979]" pageId="11" pageNumber="112">FIGURE 6.</emphasis>
<taxonomicName id="4C5F4D58462CFFC5FEA5CAD0FE2550B8" authorityName="H. M. Steyn" authorityYear="2017" box="[263,444,1958,1979]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="112" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="B92BEAC9462CFFC5FEA5CAD0FE2550B8" box="[263,444,1958,1979]" italics="true" pageId="11" pageNumber="112">Acanthopsis nitida</emphasis>
</taxonomicName>
. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp; F. Bract. G. Calyx. H. Corolla (lobes lilac to purple; throat lemon-yellow). I. Glandular hairs at base of style. J. Capsule. K. Capsule with seed. Vouchers: CI =
<emphasis id="B92BEAC9462CFFC5FB7ACABCFAD850DC" box="[1240,1345,1994,2015]" italics="true" pageId="11" pageNumber="112">Steyn 1884</emphasis>
(PRE), J =
<emphasis id="B92BEAC9462CFFC5FF2ACA98FE835F00" box="[136,282,2030,2051]" italics="true" pageId="11" pageNumber="112">Burgoyne 1514</emphasis>
(PRE), K =
<emphasis id="B92BEAC9462CFFC5FE31CA98FE645F00" box="[403,509,2030,2051]" italics="true" pageId="11" pageNumber="112">Steyn 2129</emphasis>
(PRE). Photographed by: M. Koekemoer (A &amp; B) and H.M. Steyn (CK).
</paragraph>
</caption>
<subSubSection id="C3456550462BFFC2FF1FCDE9FEEB57DF" pageId="12" pageNumber="113" type="etymology">
<paragraph id="8BE036DB462BFFC2FF1FCDE9FEEB57DF" blockId="12.[136,1452,159,976]" pageId="12" pageNumber="113">
<emphasis id="B92BEAC9462BFFC2FF1FCDE9FED157BB" bold="true" box="[189,328,159,184]" pageId="12" pageNumber="113">Etymology:</emphasis>
—The specific epithet
<taxonomicName id="4C5F4D58462BFFC2FDEECDE9FD1457BB" authorityName="H. M. Steyn" authorityYear="2017" box="[588,653,159,184]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="B92BEAC9462BFFC2FDEECDE9FD1457BB" box="[588,653,159,184]" italics="true" pageId="12" pageNumber="113">nitida</emphasis>
</taxonomicName>
, meaning “shiny/glossy”, refers to the glossy leaves, floral bracts and calyx of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3456550462BFFC2FF1FCD91FBF954D3" pageId="12" pageNumber="113" type="distribution">
<paragraph id="8BE036DB462BFFC2FF1FCD91FE4A56FF" blockId="12.[136,1452,159,976]" pageId="12" pageNumber="113">
<emphasis id="B92BEAC9462BFFC2FF1FCD91FD1A5603" bold="true" box="[189,643,231,256]" pageId="12" pageNumber="113">Distribution, ecology and phenology:</emphasis>
<taxonomicName id="4C5F4D58462BFFC2FD3DCD91FCED5603" authorityName="H. M. Steyn" authorityYear="2017" box="[671,884,231,256]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="B92BEAC9462BFFC2FD3DCD91FCED5603" box="[671,884,231,256]" italics="true" pageId="12" pageNumber="113">Acanthopsis nitida</emphasis>
</taxonomicName>
is endemic to the
<collectingRegion id="499BF839462BFFC2FBFACD91FA9B5603" box="[1112,1282,231,256]" country="South Africa" name="Northern Cape" pageId="12" pageNumber="113">Northern Cape</collectingRegion>
and is known from three localities in an area to the west of Steinkopf (
<figureCitation id="13642A5E462BFFC2FCB5CC7DFCC25627" box="[791,859,267,292]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="12" pageNumber="113">Fig. 1</figureCitation>
). It is centred in the core part of the Gariep Centre of Endemism (
<bibRefCitation id="EFCE4B2A462BFFC2FE93CC59FDD4564B" author="Van Wyk, A. E. &amp; Smith, G. F." box="[305,589,303,328]" pageId="12" pageNumber="113" refId="ref13657" refString="Van Wyk, A. E. &amp; Smith, G. F. (2001) Regions of floristic endemism in southern Africa: a review with emphasis on succulents. Umdaus Press, Pretoria, 160 pp." type="book" year="2001">Van Wyk &amp; Smith 2001</bibRefCitation>
), a biogeographical region rich in restricted-range plants and animals. This distribution falls within the Succulent Karoo Biome (
<bibRefCitation id="EFCE4B2A462BFFC2FD4BCC25FBB7566F" author="Rutherford, M. C. &amp; Westfall, R. H." box="[745,1070,339,364]" pageId="12" pageNumber="113" refId="ref13271" refString="Rutherford, M. C. &amp; Westfall, R. H. (1994) Biomes of southern Africa: an objective categorization. [Memoir 63]. National Botanical Institute, Pretoria, 94 pp." type="book" year="1994">Rutherford &amp; Westfall 1994</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A462BFFC2FB99CC25FAB3566F" author="Low, A. B. &amp; Rebelo, A. G." box="[1083,1322,339,364]" pageId="12" pageNumber="113" refId="ref12933" refString="Low, A. B., Rebelo, A. G. (Eds.) (1996) Vegetation of South Africa, Lesotho and Swaziland. Department of Environmental Affairs &amp; Tourism, Pretoria, 85 pp." type="book" year="1996">Low &amp; Rebelo 1996</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A462BFFC2FA95CC25FEDF5693" author="Mucina, L. &amp; Rutherford, M. C." pageId="12" pageNumber="113" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) in the Richtersveld Bioregion, Anenous Plateau Shrubland (
<bibRefCitation id="EFCE4B2A462BFFC2FBA3CC01FAA75693" author="Mucina, L. &amp; Rutherford, M. C." box="[1025,1342,375,400]" pageId="12" pageNumber="113" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) where it receives predominantly winter rains with an average of ±
<quantity id="4CA79B3E462BFFC2FCBECCEDFCE356B7" box="[796,890,411,436]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="113" unit="mm" value="150.0">150 mm</quantity>
per year. The specimens were collected on stony mountain slopes in well-drained sandy soils at elevations of
<quantity id="4CA79B3E462BFFC2FCEACCC9FC4356DB" box="[840,986,447,472]" metricMagnitude="2" metricUnit="m" metricValue="8.95" metricValueMax="11.0" metricValueMin="6.9" pageId="12" pageNumber="113" unit="m" value="895.0" valueMax="1100.0" valueMin="690.0">6901100 m</quantity>
. Specimens with flowers were collected between August and October.
</paragraph>
<paragraph id="8BE036DB462BFFC2FF1FCF71FEE4558F" blockId="12.[136,1452,159,976]" pageId="12" pageNumber="113">
<emphasis id="B92BEAC9462BFFC2FF1FCF71FE2B5523" bold="true" box="[189,434,519,544]" pageId="12" pageNumber="113">Conservation status:</emphasis>
<taxonomicName id="4C5F4D58462BFFC2FE6CCF71FD045523" authorityName="H. M. Steyn" authorityYear="2017" box="[462,669,519,544]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="B92BEAC9462BFFC2FE6CCF71FD045523" box="[462,669,519,544]" italics="true" pageId="12" pageNumber="113">Acanthopsis nitida</emphasis>
</taxonomicName>
is a range-restricted species (EOO
<quantity id="4CA79B3E462BFFC2FB82CF71FBFF5523" box="[1056,1126,519,544]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="113" unit="km" value="42.0">42 km</quantity>
²) known from three locations within a small area. It is potentially threatened by habitat loss and degradation due to mining and overgrazing and is categorised as Endangered B1ab(iii) according to the IUCN Red List Category and Criteria (
<bibRefCitation id="EFCE4B2A462BFFC2FB18CF39FADC556B" author="IUCN" box="[1210,1349,591,616]" pageId="12" pageNumber="113" refId="ref12795" refString="IUCN (2012) IUCN Red List categories and criteria version 3.1. IUCN Species Survival Commission, Gland, Switzerland and Cambridge, UK, iv + 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
) (L. von Staden, pers. comm.).
</paragraph>
<paragraph id="8BE036DB462BFFC2FF1FCFE1FE665443" blockId="12.[136,1452,159,976]" pageId="12" pageNumber="113">
<materialsCitation id="3B373C86462BFFC2FF1FCFE1FBF555D7" collectingDate="2012-08-22" collectionCode="BA, PRE" collectorName="Namaqualand District. Anenous Pass, S" country="South Africa" elevation="691" location="Northern Cape" pageId="12" pageNumber="113" specimenCount="2" stateProvince="Northern Cape">
<emphasis id="B92BEAC9462BFFC2FF1FCFE1FD7F55B3" bold="true" box="[189,742,663,688]" pageId="12" pageNumber="113">
<typeStatus id="54E48879462BFFC2FF1FCFE1FDDD55B3" box="[189,580,663,688]" isOtherTypeStatus="true" pageId="12" pageNumber="113">Additional specimens examined</typeStatus>
(
<typeStatus id="54E48879462BFFC2FDF8CFE1FD4055B3" box="[602,729,663,688]" pageId="12" pageNumber="113" type="paratype">Paratypes</typeStatus>
):
</emphasis>
<collectingCountry id="F348764B462BFFC2FCA0CFE1FC4C55B3" box="[770,981,663,688]" name="South Africa" pageId="12" pageNumber="113">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="499BF839462BFFC2FC44CFE1FB0E55B3" box="[998,1175,663,688]" country="South Africa" name="Northern Cape" pageId="12" pageNumber="113">Northern Cape</collectingRegion>
:
<collectorName id="26AA530D462BFFC2FB05CFE1FEA655D7" pageId="12" pageNumber="113">Namaqualand District. Anenous Pass, S</collectorName>
slope of mountain,
<quantity id="4CA79B3E462BFFC2FDBACFCDFDF955D7" box="[536,608,699,724]" metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" pageId="12" pageNumber="113" unit="m" value="691.0">
<elevation id="0072D1E8462BFFC2FDBACFCDFDF955D7" box="[536,608,699,724]" metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" pageId="12" pageNumber="113" unit="m" value="691.0">691 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462BFFC2FD0FCFCDFD4F55D7" box="[685,726,699,724]" country="Argentina" lsid="urn:lsid:biocol.org:col:15002" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="12" pageNumber="113" type="Herbarium">BA</collectionCode>
),
<date id="FFE1101B462BFFC2FD4ACFCDFC0055D7" box="[744,921,699,724]" pageId="12" pageNumber="113" value="2012-08-22">
<collectingDate id="EFA5E9F3462BFFC2FD4ACFCDFC0055D7" box="[744,921,699,724]" pageId="12" pageNumber="113" value="2012-08-22">22 August 2012</collectingDate>
</date>
,
<emphasis id="B92BEAC9462BFFC2FC01CFCDFB8455D7" box="[931,1053,699,724]" italics="true" pageId="12" pageNumber="113">Steyn 1884</emphasis>
(
<collectionCode id="ED4EAE1E462BFFC2FB88CFCDFBFA55D7" box="[1066,1123,699,724]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="12" pageNumber="113" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462BFFC2FBD4CFCDFAD955D7" box="[1142,1344,699,724]" country="South Africa" location="Steinkopf District" pageId="12" pageNumber="113" specimenCount="1">
<location id="8E806000462BFFC2FBD4CFCDFAD955D7" LSID="urn:lsid:plazi:treatment:03F687CD462DFFC2FF2AC8F3FBF954D3:8E806000462BFFC2FBD4CFCDFAD955D7" box="[1142,1344,699,724]" country="South Africa" name="Steinkopf District" pageId="12" pageNumber="113">Steinkopf District</location>
</materialsCitation>
; Anenous Pass;
<materialsCitation id="3B373C86462BFFC2FF6ACFA9FD0C55FB" box="[200,661,735,760]" country="South Africa" location="17 km West of Steinkopf" pageId="12" pageNumber="113" specimenCount="1">
Approximately
<location id="8E806000462BFFC2FEDACFA9FD0C55FB" LSID="urn:lsid:plazi:treatment:03F687CD462DFFC2FF2AC8F3FBF954D3:8E806000462BFFC2FEDACFA9FD0C55FB" box="[376,661,735,760]" country="South Africa" name="17 km West of Steinkopf" pageId="12" pageNumber="113">
<quantity id="4CA79B3E462BFFC2FEDACFA9FE5955FB" box="[376,448,735,760]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="113" unit="km" value="17.0">17 km</quantity>
West of Steinkopf
</location>
</materialsCitation>
;
<materialsCitation id="3B373C86462BFFC2FD02CFA9FDF0541F" collectingDate="1992-12-20" collectionCode="BC, PRE" collectorName="Pass" country="South Africa" elevation="1100" pageId="12" pageNumber="113" specimenCount="1">
hills on southern side of the
<collectorName id="26AA530D462BFFC2FC42CFA9FB8B55FB" box="[992,1042,735,760]" pageId="12" pageNumber="113">Pass</collectorName>
on the old road,
<quantity id="4CA79B3E462BFFC2FB70CFA9FAB155FB" box="[1234,1320,735,760]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="113" unit="m" value="1100.0">
<elevation id="0072D1E8462BFFC2FB70CFA9FAB155FB" box="[1234,1320,735,760]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="113" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462BFFC2FADACFA9FA0655FB" box="[1400,1439,735,760]" country="Spain" lsid="urn:lsid:biocol.org:col:13287" name="Institut Botànic de Barcelona" pageId="12" pageNumber="113" type="Herbarium">BC</collectionCode>
),
<date id="FFE1101B462BFFC2FF2ACE75FEF8541F" box="[136,353,771,796]" pageId="12" pageNumber="113" value="1992-12-20">
<collectingDate id="EFA5E9F3462BFFC2FF2ACE75FEF8541F" box="[136,353,771,796]" pageId="12" pageNumber="113" value="1992-12-20">20 December 1992</collectingDate>
</date>
,
<emphasis id="B92BEAC9462BFFC2FECFCE75FD81541F" box="[365,536,771,796]" italics="true" pageId="12" pageNumber="113">Burgoyne 1514</emphasis>
(
<collectionCode id="ED4EAE1E462BFFC2FD8ACE75FDF8541F" box="[552,609,771,796]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="12" pageNumber="113" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462BFFC2FDD7CE75FE625443" collectingDate="2015-10-10" collectionCode="BC, PRE" country="South Africa" elevation="838" location="Namaqualand District" pageId="12" pageNumber="113" specimenCount="1">
<location id="8E806000462BFFC2FDD7CE75FCE9541F" LSID="urn:lsid:plazi:treatment:03F687CD462DFFC2FF2AC8F3FBF954D3:8E806000462BFFC2FDD7CE75FCE9541F" box="[629,880,771,796]" country="South Africa" name="Namaqualand District" pageId="12" pageNumber="113">Namaqualand District</location>
. On dirt road
<locationDeviation id="9A522FF9462BFFC2FBAECE75FB36541F" box="[1036,1199,771,796]" location="Eyams" pageId="12" pageNumber="113">NW of Eyams</locationDeviation>
,
<quantity id="4CA79B3E462BFFC2FB19CE75FA9C541F" box="[1211,1285,771,796]" metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" pageId="12" pageNumber="113" unit="m" value="838.0">
<elevation id="0072D1E8462BFFC2FB19CE75FA9C541F" box="[1211,1285,771,796]" metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" pageId="12" pageNumber="113" unit="m" value="838.0">838 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462BFFC2FAF6CE75FAE2541F" box="[1364,1403,771,796]" country="Spain" lsid="urn:lsid:biocol.org:col:13287" name="Institut Botànic de Barcelona" pageId="12" pageNumber="113" type="Herbarium">BC</collectionCode>
),
<date id="FFE1101B462BFFC2FA2DCE75FEBD5443" pageId="12" pageNumber="113" value="2015-10-10">
<collectingDate id="EFA5E9F3462BFFC2FA2DCE75FEBD5443" pageId="12" pageNumber="113" value="2015-10-10">10 October 2015</collectingDate>
</date>
,
<emphasis id="B92BEAC9462BFFC2FE8DCE51FE325443" box="[303,427,807,832]" italics="true" pageId="12" pageNumber="113">Steyn 2138</emphasis>
(
<collectionCode id="ED4EAE1E462BFFC2FE18CE51FE6A5443" box="[442,499,807,832]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="12" pageNumber="113" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE036DB462BFFC2FF1FCE3DFBF954D3" blockId="12.[136,1452,159,976]" pageId="12" pageNumber="113">
<emphasis id="B92BEAC9462BFFC2FF1FCE3DFE905467" bold="true" box="[189,265,843,868]" pageId="12" pageNumber="113">Notes:</emphasis>
<taxonomicName id="4C5F4D58462BFFC2FE84CE3DFE6F5467" authorityName="H. M. Steyn" authorityYear="2017" box="[294,502,843,868]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="113" phylum="Tracheophyta" rank="species" species="nitida">
<emphasis id="B92BEAC9462BFFC2FE84CE3DFE6F5467" box="[294,502,843,868]" italics="true" pageId="12" pageNumber="113">Acanthopsis nitida</emphasis>
</taxonomicName>
can be easily distinguished from the other subshrubs with semi-dense inflorescences in the genus by its glossy appearance (
<emphasis id="B92BEAC9462BFFC2FDECCE19FDEB548B" box="[590,626,879,904]" italics="true" pageId="12" pageNumber="113">not</emphasis>
matt) and yellow-green (
<emphasis id="B92BEAC9462BFFC2FC39CE19FC26548B" box="[923,959,879,904]" italics="true" pageId="12" pageNumber="113">not</emphasis>
green or grey-green) leaves. Although the central primary spine could be simple or compound (one pair of short, marginal secondary spines), the lateral primary spines always have a basal secondary spine on the side facing the central primary spine.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,581 @@
<document id="F8DF87704EC9B040ED5F82BB64E2E160" ID-DOI="10.11646/phytotaxa.324.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13697360" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725568056204" checkinUser="felipe" docAuthor="Steyn, Hester M. &amp; Van Wyk, Abraham E." docDate="2017" docId="03F687CD462FFFC4FF2AC9A7FD3B525E" docLanguage="en" docName="phytotaxa.324.2.1.pdf" docOrigin="Phytotaxa 324 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.324.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Acanthopsis scullyi Obermeyer 1937" docType="treatment" docVersion="2" lastPageNumber="111" masterDocId="FFCFFFB54627FFCEFFA2CD76FF995703" masterDocTitle="Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts" masterLastPageNumber="121" masterPageNumber="101" pageNumber="109" updateTime="1725570993962" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="A77DF413A755FD0D7240D8FCC372DAD4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E262F8172C14DBEB09FA53A42CA5F9E9">
<mods:title id="824E2A26DF471388A85E3862E02110BE">Taxonomic notes on Acanthopsis (Acanthaceae, tribe Acantheae): the group with semi-dense spikes and 5 - fid bracts</mods:title>
</mods:titleInfo>
<mods:name id="A4ABE88621AB0B102C67561BBBE8EBED" type="personal">
<mods:role id="CFC265CEF2FC83A9B8A6D143C4E00815">
<mods:roleTerm id="F553B2B0DA70F4CE6CFA674CC96E285A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EFFA3436BC75BB6C2440F4322697D252">Steyn, Hester M.</mods:namePart>
<mods:affiliation id="663CC7007B429EF5CAD4AA4C945EFB33">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:name id="D42B896FA015AACFDD0E79EE0247214F" type="personal">
<mods:role id="DAEDE7C4732CFDC39F159B86F937A417">
<mods:roleTerm id="DBAA0E9FD8CCFC69A9AD0C39AC236C48">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5194596C01E3469A79E91237DD3DDCF8">Van Wyk, Abraham E.</mods:namePart>
<mods:affiliation id="5523AB947F1DEB7CCBD58B3E019A9591">National Herbarium, South African National Biodiversity Institute, Private Bag X 101, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="30ACE4A38D7E9AA9AA86DD059970C40F">text</mods:typeOfResource>
<mods:relatedItem id="F4EFC89D83D1FC794A29DC908775A5FB" type="host">
<mods:titleInfo id="8A48C736207AEE8BF300E69E4E035DFA">
<mods:title id="D19305BD17841CF67E5EB0B37CF172F2">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="0601F0CFA8FC53D629100F74475F2514">
<mods:date id="B75C643637E4F017C8D20C492CDD4450">2017</mods:date>
<mods:detail id="62872888D25B09A3C4B0298EDDA15B00" type="pubDate">
<mods:number id="E4281F605BC04C728A7AA1C105C1CEDA">2017-10-10</mods:number>
</mods:detail>
<mods:detail id="BFD8F84F91ECF9069DA4C367779F1164" type="volume">
<mods:number id="7C47B1D876D4305FC64148AA41FF1881">324</mods:number>
</mods:detail>
<mods:detail id="231A7B1EC015827858EAAEE39A711EDF" type="issue">
<mods:number id="498EC407CE5DB1C10AC41F8D29071B49">2</mods:number>
</mods:detail>
<mods:extent id="E5186B0B569DC1404700715A2E85752F" unit="page">
<mods:start id="958600F15771AE41856F256532813BEA">101</mods:start>
<mods:end id="FC660C242D46A9C1B41F09B863268D88">121</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="2FBA0E69981440FB600957A69482E926">
<mods:url id="5426D1284DAEA7C6EB4FB9ABD99CE026">http://dx.doi.org/10.11646/phytotaxa.324.2.1</mods:url>
</mods:location>
<mods:classification id="F1372588441D42A6C995A1D605C3F8CE">journal article</mods:classification>
<mods:identifier id="A070D70D376AA1C16D61BEEB00BE9C0E" type="DOI">10.11646/phytotaxa.324.2.1</mods:identifier>
<mods:identifier id="4E2408850ED559264BA52BC1BCDE9784" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="61659460AD6B85FA1CD12992CA141403" type="Zenodo-Dep">13697360</mods:identifier>
</mods:mods>
<treatment id="03F687CD462FFFC4FF2AC9A7FD3B525E" ID-DOI="http://doi.org/10.5281/zenodo.13697376" ID-Zenodo-Dep="13697376" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E" httpUri="http://treatment.plazi.org/id/03F687CD462FFFC4FF2AC9A7FD3B525E" lastPageId="10" lastPageNumber="111" pageId="8" pageNumber="109">
<subSubSection id="C3456550462FFFC6FF2AC9A7FB5053E9" box="[136,1225,1231,1258]" pageId="8" pageNumber="109" type="nomenclature">
<paragraph id="8BE036DB462FFFC6FF2AC9A7FB5053E9" blockId="8.[136,1452,1231,1401]" box="[136,1225,1231,1258]" pageId="8" pageNumber="109">
<heading id="D0A881B7462FFFC6FF2AC9A7FB5053E9" box="[136,1225,1231,1258]" fontSize="11" level="2" pageId="8" pageNumber="109" reason="4">
<emphasis id="B92BEAC9462FFFC6FF2AC9A7FF0453E9" bold="true" box="[136,157,1232,1258]" pageId="8" pageNumber="109">2.</emphasis>
<taxonomicName id="4C5F4D58462FFFC6FF01C9B9FC8953E9" ID-CoL="8ZLH" authority="(S. Moore) Obermeyer (1937: 138)" authorityName="Obermeyer" authorityPageNumber="138" authorityYear="1937" baseAuthorityName="S. Moore" box="[163,784,1231,1258]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC9462FFFC6FF01C9B9FE1A53E9" bold="true" box="[163,387,1231,1258]" italics="true" pageId="8" pageNumber="109">Acanthopsis scullyi</emphasis>
(S.Moore)
<bibRefCitation id="EFCE4B2A462FFFC6FDA6C9A7FC8953E9" author="Obermeyer, A. A." box="[516,784,1233,1258]" pageId="8" pageNumber="109" pagination="105 - 139" refId="ref13166" refString="Obermeyer, A. A. (1937) The South African species of Blepharis. Annals of the Transvaal Museum 14: 105 - 139." type="journal article" year="1937">Obermeyer (1937: 138)</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EFCE4B2A462FFFC6FCB9C9A7FB9A53E9" author="Snijman, D. A." box="[795,1027,1233,1258]" pageId="8" pageNumber="109" refId="ref13383" refString="Snijman, D. A. (2013) Acanthopis. In: Snijman, D. A. (Ed.) Plants of the Greater Cape Floristic Region, Vol. 2: the Extra Cape Flora. Strelitzia 30. South African National Biodiversity Institute, Pretoria, 163 pp." type="book" year="2013">Snijman (2013: 163)</bibRefCitation>
(
<figureCitation id="13642A5E462FFFC6FBB0C9A7FBF953E9" box="[1042,1120,1233,1258]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="8" pageNumber="109">Figs. 1</figureCitation>
,
<figureCitation id="13642A5E462FFFC6FBCCC9A7FB1753E9" box="[1134,1166,1233,1258]" captionStart="FIGURE 3" captionStartId="6.[136,229,1911,1932]" captionTargetBox="[151,1436,190,1886]" captionTargetId="figure-15@6.[151,1436,190,1886]" captionTargetPageId="6" captionText="FIGURE 3. Comparison of floral parts, associated bracts and bracteoles (from left to right): bracts (2), bracteoles (2), calyx (4 sepals: dorsal, ventral and 2 lateral ones) and corolla with anthers.A. A. glabra, B. A. scullyi, C. A. nitida, D. A. glandulopalmata, E. A. insueta, F. A. carduifolia. Note: the bract on the left is from lower down in the same inflorescence. Scale bar = 10 mm. Vouchers:A = Steyn 1893 (PRE), B = Steyn 1911 (PRE), C = Steyn 1884 (PRE), D = Steyn 1890 (PRE), E = Steyn 1805 (PRE), F = Koekemoer 4620 (PRE)." figureDoi="http://doi.org/10.5281/zenodo.13697366" httpUri="https://zenodo.org/record/13697366/files/figure.png" pageId="8" pageNumber="109">3B</figureCitation>
&amp;
<figureCitation id="13642A5E462FFFC6FB10C9A7FB5953E9" box="[1202,1216,1233,1258]" captionStart="FIGURE 5" captionStartId="9.[136,229,1780,1801]" captionTargetBox="[151,1436,175,1755]" captionTargetId="figure-15@9.[151,1436,175,1755]" captionTargetPageId="9" captionText="FIGURE 5. Acanthopsis scullyi. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp; F. Bract. G. Calyx. H. Corolla (lobes cream to pale purple; throat cream). I. Glandular hairs at base of style. J. Capsule. K. Capsule with seed. Vouchers: C = Steyn 2134 (PRE), DK = Steyn 1911 (PRE). Photographed by: H.M. Steyn (A, CK) and M. Koekemoer (B)." figureDoi="http://doi.org/10.5281/zenodo.13697370" httpUri="https://zenodo.org/record/13697370/files/figure.png" pageId="8" pageNumber="109">5</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3456550462FFFC6FF2AC98EFEEB5232" pageId="8" pageNumber="109" type="reference_group">
<paragraph id="8BE036DB462FFFC6FF2AC98EFEEB5232" blockId="8.[136,1452,1231,1401]" pageId="8" pageNumber="109">
Basionym:—
<taxonomicName id="4C5F4D58462FFFC6FEAAC98EFDE9520E" ID-CoL="M5Z4" authority="Moore (1901: 301)" authorityName="Moore" authorityPageNumber="301" authorityYear="1901" box="[264,624,1272,1293]" class="Magnoliopsida" family="Acanthaceae" genus="Blepharis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC9462FFFC6FEAAC98EFE35520E" box="[264,428,1272,1293]" italics="true" pageId="8" pageNumber="109">Blepharis scullyi</emphasis>
<bibRefCitation id="EFCE4B2A462FFFC6FE11C98EFDE9520E" author="Moore, S. le" box="[435,624,1272,1293]" pageId="8" pageNumber="109" pagination="300 - 305" refId="ref12997" refString="Moore, S. le M. (1901) Some recent additions to the British Museum: Acanthaceae. Journal of Botany, British and Foreign 39: 300 - 305." type="journal article" year="1901">Moore (1901: 301)</bibRefCitation>
</taxonomicName>
.
<emphasis id="B92BEAC9462FFFC6FDD9C98EFD1A520E" box="[635,643,1272,1293]" italics="true" pageId="8" pageNumber="109"></emphasis>
<taxonomicName id="4C5F4D58462FFFC6FD21C98EFC33520E" ID-CoL="92S2" authority="Thunb." authorityName="Thunb." box="[643,938,1272,1293]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthus" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="carduifolius">
<emphasis id="B92BEAC9462FFFC6FD21C98EFCC4520E" box="[643,861,1272,1293]" italics="true" pageId="8" pageNumber="109">Acanthus carduifolius</emphasis>
Thunb.
</taxonomicName>
<emphasis id="B92BEAC9462FFFC6FC12C981FB1F520E" box="[944,1158,1271,1293]" italics="true" pageId="8" pageNumber="109">ά spica glabriuscula”</emphasis>
, ms. [name written on sheet],
<emphasis id="B92BEAC9462FFFC6FF1FC86AFEEB5232" box="[189,370,1308,1329]" italics="true" pageId="8" pageNumber="109">sensu Drège 2431.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3456550462FFFC6FF2AC836FD525F1C" pageId="8" pageNumber="109" type="materials_examined">
<paragraph id="8BE036DB462FFFC6FF2AC836FD1F527A" blockId="8.[136,1452,1231,1401]" pageId="8" pageNumber="109">
<emphasis id="B92BEAC9462FFFC6FF2AC836FF5B5256" bold="true" box="[136,194,1344,1365]" pageId="8" pageNumber="109">Type:</emphasis>
<materialsCitation id="3B373C86462FFFC6FF79C836FD1B527A" collectionCode="BM, PRE" country="South Africa" location="Namaqualand" pageId="8" pageNumber="109" specimenCode="BM000931147, PRE0129526" specimenCount="2" stateProvince="Northern Cape" typeStatus="holotype">
<collectingCountry id="F348764B462FFFC6FF79C836FE0B5256" box="[219,402,1344,1365]" name="South Africa" pageId="8" pageNumber="109">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="499BF839462FFFC6FE01C836FDA55256" box="[419,572,1344,1365]" country="South Africa" name="Northern Cape" pageId="8" pageNumber="109">Northern Cape</collectingRegion>
:
<location id="8E806000462FFFC6FDEFC836FD4B5256" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462FFFC6FDEFC836FD4B5256" box="[589,722,1344,1365]" country="South Africa" name="Namaqualand" pageId="8" pageNumber="109" stateProvince="Northern Cape">Namaqualand</location>
[no precise locality],
<emphasis id="B92BEAC9462FFFC6FC66C836FBA15256" box="[964,1080,1344,1365]" italics="true" pageId="8" pageNumber="109">
<taxonomicName id="4C5F4D58462FFFC6FC66C836FB9C5256" authorityName="Moore" authorityYear="1901" box="[964,1029,1344,1365]" class="Magnoliopsida" family="Acanthaceae" genus="Blepharis" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="109" phylum="Tracheophyta" rank="species" species="scullyi">Scullyi</taxonomicName>
249
</emphasis>
(
<typeStatus id="54E48879462FFFC6FBEEC836FB3B5256" box="[1100,1186,1344,1365]" pageId="8" pageNumber="109" type="holotype">holotype</typeStatus>
<specimenCode id="DBF99EA0462FFFC6FB12C836FAD15256" box="[1200,1352,1344,1365]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="8" pageNumber="109">BM000931147</specimenCode>
!;
<typeStatus id="54E48879462FFFC6FAFFC836FA355256" box="[1373,1452,1344,1365]" pageId="8" pageNumber="109" type="isotype">isotypes</typeStatus>
BOL138596!, K000394339!,
<specimenCode id="DBF99EA0462FFFC6FE79C812FDFF527A" box="[475,614,1380,1401]" collectionCode="PRE" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="8" pageNumber="109" type="Herbarium">PRE0129526</specimenCode>
-0!)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE036DB462FFFC6FF2AC8D4FD525F1C" blockId="8.[136,1452,1442,2079]" pageId="8" pageNumber="109">
Perennial, acaulescent herb or compact subshrub,
<quantity id="4CA79B3E462FFFC6FD61C8D4FCAD52B8" box="[707,820,1442,1467]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="8" pageNumber="109" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">1015 cm</quantity>
tall with very short internodes. Leaves sessile, petiole-like base with spines
<quantity id="4CA79B3E462FFFC6FEDFC8B0FE4252DC" box="[381,475,1478,1503]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="8" pageNumber="109" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long; lamina oblanceolate, 55105 × 1015(18) mm, pubescent to villose with long, silky hairs, occasionally interspersed with long glandular hairs; apex acute, base attenuate, margin undulate, coarsely dentate-spinose, spines fine to rigid,
<quantity id="4CA79B3E462FFFC6FD82CB78FDE45124" box="[544,637,1550,1575]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="8" pageNumber="109" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
long, yellowish. Inflorescences usually lax (bract lamina overlap &lt;30%), cylindrical, subsessile with a number of infertile bracts at the base; 80120(150) mm long, 810(12) mm in diameter. Bracts obovate to broadly obovate, (16)
<quantity id="4CA79B3E462FFFC6FDEECB20FD5F516C" box="[588,710,1622,1647]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="8" pageNumber="109" unit="mm" value="20.0" valueMax="22.0" valueMin="18.0">1822 mm</quantity>
long (including spines), lamina 1012(14) mm long, base obtuse; villose with antrorse (often spreading) long, silky hairs interspersed with short hairs and occasionally also with glandular hairs; middle and upper bracts ending in 5 primary spines, 3 central primary spines usually broadly triangular; central primary spine nearly always compound with 1 or 2 pairs of short, marginal secondary spines (prickles); lateral primary spines usually with 1 short marginal secondary spine on side facing central primary spine, occasionally also with 1 short, marginal secondary spine on opposite side; primary spines spreading in flower and fruit. Bracteoles linear, 68( 9) mm long, silky-hairy, also with long, glandular hairs. Calyx with dorsal sepal lanceolate to ovate, acuminate,
<quantity id="4CA79B3E462FFFC6FAC4CA58FF2D5068" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="8" pageNumber="109" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">1214 mm</quantity>
long, silky-hairy, often also with long, glandular hairs, 57-veined; ventral sepal lanceolate to narrowly ovate,
<quantity id="4CA79B3E462FFFC6FF2ACA00FF66508C" box="[136,255,1910,1935]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="8" pageNumber="109" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">1113 mm</quantity>
long, silky-hairy, 5(7)-veined; lateral sepals
<quantity id="4CA79B3E462FFFC6FD59CA00FCFC508C" box="[763,869,1910,1935]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="8" pageNumber="109" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
long, densely silky-hairy especially at base. Flowers clearly decussate; corolla cream to light mauve with lilac veins, cream throat, (21)2531 × (8)
<quantity id="4CA79B3E462FFFC6FB61CAECFAA950B0" box="[1219,1328,1946,1971]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="8" pageNumber="109" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">911 mm</quantity>
, tube
<quantity id="4CA79B3E462FFFC6FAD6CAECFF2D50D4" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="8" pageNumber="109" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
long, pubescent, central lobe often square (as wide as long), not or only slightly constricted at the base, truncate to emarginate. Filaments 78(10) mm long, glandular; anthers beige,
<quantity id="4CA79B3E462FFFC6FC03CA94FC7B50F8" box="[929,994,2018,2043]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="109" unit="mm" value="3.0">3 mm</quantity>
long. Style with patch of glandular hairs at the base. Capsules 89 ×
<quantity id="4CA79B3E462FFFC6FE62C570FD9D5F1C" box="[448,516,2054,2079]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="109" unit="mm" value="4.0">4 mm</quantity>
. Seeds 4 ×
<quantity id="4CA79B3E462FFFC6FD21C570FD5E5F1C" box="[643,711,2054,2079]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="109" unit="mm" value="2.0">2 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF206653462EFFC7FF2ACB82FC325052" ID-DOI="http://doi.org/10.5281/zenodo.13697370" ID-Zenodo-Dep="13697370" httpUri="https://zenodo.org/record/13697370/files/figure.png" pageId="9" pageNumber="110" startId="9.[136,229,1780,1801]" targetBox="[151,1436,175,1755]" targetPageId="9" targetType="figure">
<paragraph id="8BE036DB462EFFC7FF2ACB82FC325052" blockId="9.[136,1452,1780,1873]" pageId="9" pageNumber="110">
<emphasis id="B92BEAC9462EFFC7FF2ACB82FF67500A" bold="true" box="[136,254,1780,1801]" pageId="9" pageNumber="110">FIGURE 5.</emphasis>
<taxonomicName id="4C5F4D58462EFFC7FEA7CB82FE59500A" authorityName="Obermeyer" authorityYear="1937" baseAuthorityName="S. Moore" box="[261,448,1780,1801]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC9462EFFC7FEA7CB82FE59500A" box="[261,448,1780,1801]" italics="true" pageId="9" pageNumber="110">Acanthopsis scullyi</emphasis>
</taxonomicName>
. A &amp; B. Habit. C &amp; D. Leaf indumentum (abaxial). E &amp; F. Bract. G. Calyx. H. Corolla (lobes cream to pale purple; throat cream). I. Glandular hairs at base of style. J. Capsule. K. Capsule with seed. Vouchers: C =
<emphasis id="B92BEAC9462EFFC7FB12CA6EFA83502E" box="[1200,1306,1816,1837]" italics="true" pageId="9" pageNumber="110">Steyn 2134</emphasis>
(PRE), DK =
<emphasis id="B92BEAC9462EFFC7FF2ACA4AFF695052" box="[136,240,1852,1873]" italics="true" pageId="9" pageNumber="110">Steyn 1911</emphasis>
(PRE). Photographed by: H.M. Steyn (A, CK) and M. Koekemoer (B).
</paragraph>
</caption>
<subSubSection id="C3456550462EFFC7FF1FCAF7FB7050BD" pageId="9" pageNumber="110" type="etymology">
<paragraph id="8BE036DB462EFFC7FF1FCAF7FB7050BD" blockId="9.[136,1452,1921,2090]" pageId="9" pageNumber="110">
<emphasis id="B92BEAC9462EFFC7FF1FCAF7FED15099" bold="true" box="[189,328,1921,1946]" pageId="9" pageNumber="110">Etymology:</emphasis>
—The specific epithet commemorates William Charles Scully [18551943], a magistrate, author and collector of natural history specimens, who collected the
<typeStatus id="54E48879462EFFC7FCAACAD3FCA150BD" box="[776,824,1957,1982]" pageId="9" pageNumber="110">type</typeStatus>
material somewhere in Namaqualand.
</paragraph>
</subSubSection>
<subSubSection id="C3456550462EFFC4FF1FCABFFD3B525E" lastPageId="10" lastPageNumber="111" pageId="9" pageNumber="110" type="distribution">
<paragraph id="8BE036DB462EFFC4FF1FCABFFD03564A" blockId="9.[136,1452,1921,2090]" lastBlockId="10.[136,1452,160,1373]" lastPageId="10" lastPageNumber="111" pageId="9" pageNumber="110">
<emphasis id="B92BEAC9462EFFC7FF1FCABFFDEF50E1" bold="true" box="[189,630,1993,2018]" pageId="9" pageNumber="110">Distribution, ecology and phenology:</emphasis>
—Known from a number of localities between Steinkopf in the north and the farm Eenkoker (Bitterfontein area) in the south (
<figureCitation id="13642A5E462EFFC7FD73CA9BFC8A5F05" box="[721,787,2029,2054]" captionStart="FIGURE 1" captionStartId="4.[136,229,1423,1444]" captionTargetBox="[151,1436,187,1395]" captionTargetId="figure-15@4.[151,1436,187,1395]" captionTargetPageId="4" captionText="FIGURE 1. Known distribution of A. glabra (○), A. scullyi (▲), A. nitida (□), A. glandulopalmata (■) A. insueta (Δ) and A. carduifolia (●). Thunbergs route (1774) indicated in red. (for explanation see Notes under taxonomic treatment of A. carduifolia)" figureDoi="http://doi.org/10.5281/zenodo.13697362" httpUri="https://zenodo.org/record/13697362/files/figure.png" pageId="9" pageNumber="110">Fig. 1</figureCitation>
).
<taxonomicName id="4C5F4D58462EFFC7FC89CA9BFB9D5F05" authorityName="Obermeyer" authorityYear="1937" baseAuthorityName="S. Moore" box="[811,1028,2029,2054]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="9" pageNumber="110" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC9462EFFC7FC89CA9BFB9D5F05" box="[811,1028,2029,2054]" italics="true" pageId="9" pageNumber="110">Acanthopsis scullyi</emphasis>
</taxonomicName>
is mainly found in sandy soil derived from granitic gneisses on hill slopes and sandy plains (elevation between
<quantity id="4CA79B3E462EFFC7FC13C567FBA75F29" box="[945,1086,2065,2090]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.4" metricValueMin="0.7" pageId="9" pageNumber="110" unit="m" value="1050.0" valueMax="1400.0" valueMin="700.0">7001400 m</quantity>
) in a region receiving an average of
<quantity id="4CA79B3E462DFFC4FF04CDD6FEA557BA" box="[166,316,160,185]" metricMagnitude="-1" metricUnit="m" metricValue="2.65" metricValueMax="3.7" metricValueMin="1.6" pageId="10" pageNumber="111" unit="mm" value="265.0" valueMax="370.0" valueMin="160.0">160370 mm</quantity>
of rain per year. The known distribution falls within the Namaqualand Hardeveld Bioregion (
<bibRefCitation id="EFCE4B2A462DFFC4FAF6CDD6FEFF57DE" author="Mucina, L. &amp; Rutherford, M. C." pageId="10" pageNumber="111" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) in the Succulent Karoo Biome (
<bibRefCitation id="EFCE4B2A462DFFC4FD47CDB2FBB257DE" author="Rutherford, M. C. &amp; Westfall, R. H." box="[741,1067,196,221]" pageId="10" pageNumber="111" refId="ref13271" refString="Rutherford, M. C. &amp; Westfall, R. H. (1994) Biomes of southern Africa: an objective categorization. [Memoir 63]. National Botanical Institute, Pretoria, 94 pp." type="book" year="1994">Rutherford &amp; Westfall 1994</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A462DFFC4FB9BCDB2FAB057DE" author="Low, A. B. &amp; Rebelo, A. G." box="[1081,1321,196,221]" pageId="10" pageNumber="111" refId="ref12933" refString="Low, A. B., Rebelo, A. G. (Eds.) (1996) Vegetation of South Africa, Lesotho and Swaziland. Department of Environmental Affairs &amp; Tourism, Pretoria, 85 pp." type="book" year="1996">Low &amp; Rebelo 1996</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A462DFFC4FA95CDB2FED15602" author="Mucina, L. &amp; Rutherford, M. C." pageId="10" pageNumber="111" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
), with the Kamiesberg locality falling in the Namaqualand
<collectingRegion id="499BF839462DFFC4FC57CD9EFBB65602" box="[1013,1071,232,257]" country="South Africa" name="Western Cape" pageId="10" pageNumber="111">Cape</collectingRegion>
Shrubland Bioregion (
<bibRefCitation id="EFCE4B2A462DFFC4FA97CD9EFEDC5626" author="Mucina, L. &amp; Rutherford, M. C." pageId="10" pageNumber="111" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) in the Fynbos Biome (
<bibRefCitation id="EFCE4B2A462DFFC4FDF0CC7AFCD95626" author="Low, A. B. &amp; Rebelo, A. G." box="[594,832,268,293]" pageId="10" pageNumber="111" refId="ref12933" refString="Low, A. B., Rebelo, A. G. (Eds.) (1996) Vegetation of South Africa, Lesotho and Swaziland. Department of Environmental Affairs &amp; Tourism, Pretoria, 85 pp." type="book" year="1996">Low &amp; Rebelo 1996</bibRefCitation>
,
<bibRefCitation id="EFCE4B2A462DFFC4FCEECC7AFB115626" author="Mucina, L. &amp; Rutherford, M. C." box="[844,1160,268,293]" pageId="10" pageNumber="111" refId="ref13030" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
). Specimens with flowers were collected between August and September.
</paragraph>
<paragraph id="8BE036DB462DFFC4FF1FCC22FA1D56FE" blockId="10.[136,1452,160,1373]" pageId="10" pageNumber="111">
<emphasis id="B92BEAC9462DFFC4FF1FCC22FE21566E" bold="true" box="[189,440,340,365]" pageId="10" pageNumber="111">Conservation status:</emphasis>
<taxonomicName id="4C5F4D58462DFFC4FE76CC22FD28566E" authorityName="Obermeyer" authorityYear="1937" baseAuthorityName="S. Moore" box="[468,689,340,365]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="111" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC9462DFFC4FE76CC22FD28566E" box="[468,689,340,365]" italics="true" pageId="10" pageNumber="111">Acanthopsis scullyi</emphasis>
</taxonomicName>
is known from many herbarium records and is locally common. There are at least 10 known subpopulations, and it is likely that many more exist as suitable habitat is widespread across its range. Subpopulations are large and appear to be resilient to disturbance, and therefore the population is not suspected to be declining. Although it is range-restricted (EOO
<quantity id="4CA79B3E462DFFC4FCFCCCB6FC5C56DA" box="[862,965,448,473]" metricMagnitude="6" metricUnit="m" metricValue="1.272" pageId="10" pageNumber="111" unit="km" value="1272.0">1272 km</quantity>
²), it is not threatened and is categorised as Least Concern according to the IUCN Red List Category and Criteria (
<bibRefCitation id="EFCE4B2A462DFFC4FC02CC92FBB156FE" author="IUCN" box="[928,1064,484,509]" pageId="10" pageNumber="111" refId="ref12795" refString="IUCN (2012) IUCN Red List categories and criteria version 3.1. IUCN Species Survival Commission, Gland, Switzerland and Cambridge, UK, iv + 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
) (L. von Staden, pers. comm.).
</paragraph>
<paragraph id="8BE036DB462DFFC4FF1FCF7EFABC531A" blockId="10.[136,1452,160,1373]" pageId="10" pageNumber="111">
<emphasis id="B92BEAC9462DFFC4FF1FCF7EFDDC5522" bold="true" box="[189,581,520,545]" pageId="10" pageNumber="111">Additional specimens examined:</emphasis>
<materialsCitation id="3B373C86462DFFC4FDC3CF7EFCA85546" collectingDate="1985-08-21" collectionCode="BC, NBG" collectorName="Le Roux" country="South Africa" location="Josfontein" municipality="Namaqualand Klipkoppe. Hills" pageId="10" pageNumber="111" specimenCount="1" stateProvince="Northern Cape">
<collectingCountry id="F348764B462DFFC4FDC3CF7EFCB65522" box="[609,815,520,545]" name="South Africa" pageId="10" pageNumber="111">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="499BF839462DFFC4FC9ECF7EFC715522" box="[828,1000,520,545]" country="South Africa" name="Northern Cape" pageId="10" pageNumber="111">Northern Cape</collectingRegion>
:
<collectingMunicipality id="6B84ACA1462DFFC4FC56CF7EFACD5522" box="[1012,1364,520,545]" pageId="10" pageNumber="111">Namaqualand Klipkoppe. Hills</collectingMunicipality>
around
<location id="8E806000462DFFC4FF2ACF5AFF675546" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FF2ACF5AFF675546" box="[136,254,556,581]" country="South Africa" municipality="Namaqualand Klipkoppe. Hills" name="Josfontein" pageId="10" pageNumber="111" stateProvince="Northern Cape">Josfontein</location>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FEEECF5AFEEA5546" box="[332,371,556,581]" country="Spain" lsid="urn:lsid:biocol.org:col:13287" name="Institut Botànic de Barcelona" pageId="10" pageNumber="111" type="Herbarium">BC</collectionCode>
),
<date id="FFE1101B462DFFC4FE27CF5AFDAE5546" box="[389,567,556,581]" pageId="10" pageNumber="111" value="1985-08-21">
<collectingDate id="EFA5E9F3462DFFC4FE27CF5AFDAE5546" box="[389,567,556,581]" pageId="10" pageNumber="111" value="1985-08-21">21 August 1985</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FDE3CF5AFD435546" box="[577,730,556,581]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FDE3CF5AFD045546" box="[577,669,556,581]" pageId="10" pageNumber="111">Le Roux</collectorName>
3256
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FD4ACF5AFCB05546" box="[744,809,556,581]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="10" pageNumber="111" type="Herbarium">NBG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462DFFC4FC9ECF5AFA9F556A" collectingDate="2015-10-10" collectionCode="BC, PRE" collectorName="Steyn" country="South Africa" county="Namaqualand District" elevation="694" location="Eyams" municipality="Perdewater" pageId="10" pageNumber="111" specimenCount="1">
<collectingCounty id="62814E57462DFFC4FC9ECF5AFBAC5546" box="[828,1077,556,581]" pageId="10" pageNumber="111">Namaqualand District</collectingCounty>
. On dirt road between
<collectingMunicipality id="6B84ACA1462DFFC4FA8CCF5AFA355546" box="[1326,1452,556,581]" pageId="10" pageNumber="111">Perdewater</collectingMunicipality>
and
<location id="8E806000462DFFC4FF14CF26FE9C556A" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FF14CF26FE9C556A" box="[182,261,592,617]" country="South Africa" county="Namaqualand District" municipality="Perdewater" name="Eyams" pageId="10" pageNumber="111">Eyams</location>
,
<locationDeviation id="9A522FF9462DFFC4FEADCF26FE73556A" box="[271,490,592,617]" location="Springbok" pageId="10" pageNumber="111">NNW of Springbok</locationDeviation>
(river crossing),
<quantity id="4CA79B3E462DFFC4FD04CF26FD77556A" box="[678,750,592,617]" metricMagnitude="2" metricUnit="m" metricValue="6.94" pageId="10" pageNumber="111" unit="m" value="694.0">
<elevation id="0072D1E8462DFFC4FD04CF26FD77556A" box="[678,750,592,617]" metricMagnitude="2" metricUnit="m" metricValue="6.94" pageId="10" pageNumber="111" unit="m" value="694.0">694 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FC9ECF26FCFA556A" box="[828,867,592,617]" country="Spain" lsid="urn:lsid:biocol.org:col:13287" name="Institut Botànic de Barcelona" pageId="10" pageNumber="111" type="Herbarium">BC</collectionCode>
),
<date id="FFE1101B462DFFC4FCD4CF26FBAB556A" box="[886,1074,592,617]" pageId="10" pageNumber="111" value="2015-10-10">
<collectingDate id="EFA5E9F3462DFFC4FCD4CF26FBAB556A" box="[886,1074,592,617]" pageId="10" pageNumber="111" value="2015-10-10">10 October 2015</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FB9ECF26FB2E556A" box="[1084,1207,592,617]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FB9ECF26FBE0556A" box="[1084,1145,592,617]" pageId="10" pageNumber="111">Steyn</collectorName>
2134
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FB66CF26FB64556A" box="[1220,1277,592,617]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462DFFC4FAB2CF26FA3E558E" collectingDate="2015-10-09" collectionCode="BD, PRE" collectorName="Steyn" country="South Africa" elevation="1061" location="Namaqualand District" pageId="10" pageNumber="111" specimenCount="1">
<location id="8E806000462DFFC4FAB2CF26FF79558E" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FAB2CF26FF79558E" country="South Africa" name="Namaqualand District" pageId="10" pageNumber="111">Namaqualand District</location>
. On dirt road
<locationDeviation id="9A522FF9462DFFC4FE2ACF02FC8B558E" box="[392,786,628,653]" firstLocation="Steinkopf" pageId="10" pageNumber="111" secondLocation="Concordia">between Steinkopf and Concordia</locationDeviation>
,
<quantity id="4CA79B3E462DFFC4FC82CF02FCE2558E" box="[800,891,628,653]" metricMagnitude="3" metricUnit="m" metricValue="1.061" pageId="10" pageNumber="111" unit="m" value="1061.0">
<elevation id="0072D1E8462DFFC4FC82CF02FCE2558E" box="[800,891,628,653]" metricMagnitude="3" metricUnit="m" metricValue="1.061" pageId="10" pageNumber="111" unit="m" value="1061.0">1061 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FC6CCF02FC6F558E" box="[974,1014,628,653]" pageId="10" pageNumber="111">BD</collectionCode>
),
<date id="FFE1101B462DFFC4FBAFCF02FB5C558E" box="[1037,1221,628,653]" pageId="10" pageNumber="111" value="2015-10-09">
<collectingDate id="EFA5E9F3462DFFC4FBAFCF02FB5C558E" box="[1037,1221,628,653]" pageId="10" pageNumber="111" value="2015-10-09">9 October 2015</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FB76CF02FACA558E" box="[1236,1363,628,653]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FB76CF02FA88558E" box="[1236,1297,628,653]" pageId="10" pageNumber="111">Steyn</collectorName>
2130
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FAC4CF02FA06558E" box="[1382,1439,628,653]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462DFFC4FF2ACFEEFE3955D6" collectingDate="2015-10-10" collectionCode="BD, PRE" collectorName="Steyn" country="South Africa" county="Namaqualand District" elevation="800" location="Gladkop" municipality="Bulletrap" pageId="10" pageNumber="111" specimenCount="1">
<collectingCounty id="62814E57462DFFC4FF2ACFEEFE1855B2" box="[136,385,664,689]" pageId="10" pageNumber="111">Namaqualand District</collectingCounty>
. On dirt road between
<collectingMunicipality id="6B84ACA1462DFFC4FDD7CFEEFD4455B2" box="[629,733,664,689]" pageId="10" pageNumber="111">Bulletrap</collectingMunicipality>
and
<location id="8E806000462DFFC4FCB2CFEEFCEA55B2" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FCB2CFEEFCEA55B2" box="[784,883,664,689]" country="South Africa" county="Namaqualand District" municipality="Bulletrap" name="Gladkop" pageId="10" pageNumber="111">Gladkop</location>
,
<locationDeviation id="9A522FF9462DFFC4FCDFCFEEFBC055B2" box="[893,1113,664,689]" location="Springbok" pageId="10" pageNumber="111">NNW of Springbok</locationDeviation>
,
<quantity id="4CA79B3E462DFFC4FBC3CFEEFB3055B2" box="[1121,1193,664,689]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="111" unit="m" value="800.0">
<elevation id="0072D1E8462DFFC4FBC3CFEEFB3055B2" box="[1121,1193,664,689]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="111" unit="m" value="800.0">800 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FB54CFEEFA8755B2" box="[1270,1310,664,689]" pageId="10" pageNumber="111">BD</collectionCode>
),
<date id="FFE1101B462DFFC4FA92CFEEFF5A55D6" pageId="10" pageNumber="111" value="2015-10-10">
<collectingDate id="EFA5E9F3462DFFC4FA92CFEEFF5A55D6" pageId="10" pageNumber="111" value="2015-10-10">10 October 2015</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FF72CFCAFED755D6" box="[208,334,700,725]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FF72CFCAFE9455D6" box="[208,269,700,725]" pageId="10" pageNumber="111">Steyn</collectorName>
2131
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FEFCCFCAFE0E55D6" box="[350,407,700,725]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462DFFC4FE0FCFCAFC9855FA" collectingDate="2015-10-10" collectionCode="BD, PRE" collectorName="Steyn" country="South Africa" county="Namaqualand District" elevation="856" location="Gladkop" municipality="Bulletrap" pageId="10" pageNumber="111" specimenCount="1">
<collectingCounty id="62814E57462DFFC4FE0FCFCAFD3355D6" box="[429,682,700,725]" pageId="10" pageNumber="111">Namaqualand District</collectingCounty>
. On dirt road between
<collectingMunicipality id="6B84ACA1462DFFC4FC13CFCAFB8055D6" box="[945,1049,700,725]" pageId="10" pageNumber="111">Bulletrap</collectingMunicipality>
and
<location id="8E806000462DFFC4FBF6CFCAFB2E55D6" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FBF6CFCAFB2E55D6" box="[1108,1207,700,725]" country="South Africa" county="Namaqualand District" municipality="Bulletrap" name="Gladkop" pageId="10" pageNumber="111">Gladkop</location>
,
<locationDeviation id="9A522FF9462DFFC4FB66CFCAFA3155D6" box="[1220,1448,700,725]" location="Springbok" pageId="10" pageNumber="111">NNW of Springbok</locationDeviation>
,
<quantity id="4CA79B3E462DFFC4FF2ACF96FF4A55FA" box="[136,211,736,761]" metricMagnitude="2" metricUnit="m" metricValue="8.559999999999999" pageId="10" pageNumber="111" unit="m" value="856.0">
<elevation id="0072D1E8462DFFC4FF2ACF96FF4A55FA" box="[136,211,736,761]" metricMagnitude="2" metricUnit="m" metricValue="8.559999999999999" pageId="10" pageNumber="111" unit="m" value="856.0">856 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FE86CF96FED555FA" box="[292,332,736,761]" pageId="10" pageNumber="111">BD</collectionCode>
),
<date id="FFE1101B462DFFC4FEC0CF96FDBC55FA" box="[354,549,736,761]" pageId="10" pageNumber="111" value="2015-10-10">
<collectingDate id="EFA5E9F3462DFFC4FEC0CF96FDBC55FA" box="[354,549,736,761]" pageId="10" pageNumber="111" value="2015-10-10">10 October 2015</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FD93CF96FD3655FA" box="[561,687,736,761]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FD93CF96FDF755FA" box="[561,622,736,761]" pageId="10" pageNumber="111">Steyn</collectorName>
2132
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FD62CF96FD6055FA" box="[704,761,736,761]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462DFFC4FCACCF96FBD9541E" collectingDate="2015-10-10" collectionCode="BD, PRE" collectorName="Steyn" country="South Africa" county="Namaqualand District" elevation="835" location="Perdewater" municipality="Bulletrap" pageId="10" pageNumber="111" specimenCount="1">
<collectingCounty id="62814E57462DFFC4FCACCF96FB9255FA" box="[782,1035,736,761]" pageId="10" pageNumber="111">Namaqualand District</collectingCounty>
. On dirt road between
<collectingMunicipality id="6B84ACA1462DFFC4FAB0CF96FAE355FA" box="[1298,1402,736,761]" pageId="10" pageNumber="111">Bulletrap</collectingMunicipality>
and
<location id="8E806000462DFFC4FF2ACE72FE91541E" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FF2ACE72FE91541E" box="[136,264,772,797]" country="South Africa" county="Namaqualand District" municipality="Bulletrap" name="Perdewater" pageId="10" pageNumber="111">Perdewater</location>
,
<locationDeviation id="9A522FF9462DFFC4FEB2CE72FE70541E" box="[272,489,772,797]" location="Springbok" pageId="10" pageNumber="111">NNW of Springbok</locationDeviation>
,
<quantity id="4CA79B3E462DFFC4FE52CE72FDAF541E" box="[496,566,772,797]" metricMagnitude="2" metricUnit="m" metricValue="8.35" pageId="10" pageNumber="111" unit="m" value="835.0">
<elevation id="0072D1E8462DFFC4FE52CE72FDAF541E" box="[496,566,772,797]" metricMagnitude="2" metricUnit="m" metricValue="8.35" pageId="10" pageNumber="111" unit="m" value="835.0">835 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FD20CE72FD33541E" box="[642,682,772,797]" pageId="10" pageNumber="111">BD</collectionCode>
),
<date id="FFE1101B462DFFC4FD19CE72FCEA541E" box="[699,883,772,797]" pageId="10" pageNumber="111" value="2015-10-10">
<collectingDate id="EFA5E9F3462DFFC4FD19CE72FCEA541E" box="[699,883,772,797]" pageId="10" pageNumber="111" value="2015-10-10">10October 2015</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FCD9CE72FC6A541E" box="[891,1011,772,797]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FCD9CE72FC21541E" box="[891,952,772,797]" pageId="10" pageNumber="111">Steyn</collectorName>
2133
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FC5DCE72FBA1541E" box="[1023,1080,772,797]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462DFFC4FBEBCE72FD7F5442" collectingDate="1988-07-13" collectionCode="DB, J" collectorName="Dean" country="South Africa" elevation="800" location="Motel" municipality="Springbok" pageId="10" pageNumber="111" specimenCount="1">
<collectingMunicipality id="6B84ACA1462DFFC4FBEBCE72FB58541E" box="[1097,1217,772,797]" pageId="10" pageNumber="111">Springbok</collectingMunicipality>
,
<location id="8E806000462DFFC4FB6ACE72FA92541E" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FB6ACE72FA92541E" box="[1224,1291,772,797]" country="South Africa" municipality="Springbok" name="Motel" pageId="10" pageNumber="111">Motel</location>
<quantity id="4CA79B3E462DFFC4FAADCE72FADD541E" box="[1295,1348,772,797]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="111" unit="km" value="5.0">5km</quantity>
downhill of town,
<quantity id="4CA79B3E462DFFC4FF48CE5EFEA85442" box="[234,305,808,833]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="111" unit="m" value="800.0">
<elevation id="0072D1E8462DFFC4FF48CE5EFEA85442" box="[234,305,808,833]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="111" unit="m" value="800.0">800 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FE22CE5EFE315442" box="[384,424,808,833]" pageId="10" pageNumber="111">DB</collectionCode>
),
<date id="FFE1101B462DFFC4FE18CE5EFDD25442" box="[442,587,808,833]" pageId="10" pageNumber="111" value="1988-07-13">
<collectingDate id="EFA5E9F3462DFFC4FE18CE5EFDD25442" box="[442,587,808,833]" pageId="10" pageNumber="111" value="1988-07-13">13 July 1988</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FDF6CE5EFD585442" box="[596,705,808,833]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FDF6CE5EFD085442" box="[596,657,808,833]" pageId="10" pageNumber="111">Dean</collectorName>
547
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FD72CE5EFD445442" box="[720,733,808,833]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="10" pageNumber="111" type="Herbarium">J</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462DFFC4FD53CE5EFD315466" collectingDate="2013-08-09" collectionCode="DB, PRE" collectorName="Steyn" country="South Africa" elevation="812" location="Kokerboom Motel" municipality="Namaqualand District" pageId="10" pageNumber="111" specimenCount="1">
<collectingMunicipality id="6B84ACA1462DFFC4FD53CE5EFC735442" box="[753,1002,808,833]" pageId="10" pageNumber="111">Namaqualand District</collectingMunicipality>
.
<location id="8E806000462DFFC4FC51CE5EFB5D5442" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FC51CE5EFB5D5442" box="[1011,1220,808,833]" country="South Africa" municipality="Namaqualand District" name="Kokerboom Motel" pageId="10" pageNumber="111">Kokerboom Motel</location>
,
<locationDeviation id="9A522FF9462DFFC4FB6CCE5EFAE15442" box="[1230,1400,808,833]" location="Springbok" pageId="10" pageNumber="111">S of Springbok</locationDeviation>
,
<quantity id="4CA79B3E462DFFC4FA23CE5EFF395466" metricMagnitude="2" metricUnit="m" metricValue="8.12" pageId="10" pageNumber="111" unit="m" value="812.0">
<elevation id="0072D1E8462DFFC4FA23CE5EFF395466" metricMagnitude="2" metricUnit="m" metricValue="8.12" pageId="10" pageNumber="111" unit="m" value="812.0">812 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FF52CE3AFE815466" box="[240,280,844,869]" pageId="10" pageNumber="111">DB</collectionCode>
),
<date id="FFE1101B462DFFC4FE8ECE3AFE4A5466" box="[300,467,844,869]" pageId="10" pageNumber="111" value="2013-08-09">
<collectingDate id="EFA5E9F3462DFFC4FE8ECE3AFE4A5466" box="[300,467,844,869]" pageId="10" pageNumber="111" value="2013-08-09">9 August 2013</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FE7CCE3AFDC15466" box="[478,600,844,869]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FE7CCE3AFD825466" box="[478,539,844,869]" pageId="10" pageNumber="111">Steyn</collectorName>
1911
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FDC5CE3AFD395466" box="[615,672,844,869]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462DFFC4FD16CE3AFD8E548A" collectingDate="2015-10-08" collectionCode="DB, PRE" collectorName="Steyn" country="South Africa" elevation="813" location="Namaqualand District" pageId="10" pageNumber="111" specimenCount="1">
<location id="8E806000462DFFC4FD16CE3AFC365466" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FD16CE3AFC365466" box="[692,943,844,869]" country="South Africa" name="Namaqualand District" pageId="10" pageNumber="111">Namaqualand District</location>
.
<locationDeviation id="9A522FF9462DFFC4FC1BCE3AFB4B5466" box="[953,1234,844,869]" location="Kokerboom Motel" pageId="10" pageNumber="111">SE of Kokerboom Motel</locationDeviation>
,
<quantity id="4CA79B3E462DFFC4FB7FCE3AFABE5466" box="[1245,1319,844,869]" metricMagnitude="2" metricUnit="m" metricValue="8.129999999999999" pageId="10" pageNumber="111" unit="m" value="813.0">
<elevation id="0072D1E8462DFFC4FB7FCE3AFABE5466" box="[1245,1319,844,869]" metricMagnitude="2" metricUnit="m" metricValue="8.129999999999999" pageId="10" pageNumber="111" unit="m" value="813.0">813 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FAD5CE3AFA065466" box="[1399,1439,844,869]" pageId="10" pageNumber="111">DB</collectionCode>
),
<date id="FFE1101B462DFFC4FF2ACE06FEA5548A" box="[136,316,880,905]" pageId="10" pageNumber="111" value="2015-10-08">
<collectingDate id="EFA5E9F3462DFFC4FF2ACE06FEA5548A" box="[136,316,880,905]" pageId="10" pageNumber="111" value="2015-10-08">8 October 2015</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FEEBCE06FE5F548A" box="[329,454,880,905]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FEEBCE06FE1F548A" box="[329,390,880,905]" pageId="10" pageNumber="111">Steyn</collectorName>
2124
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FE74CE06FD96548A" box="[470,527,880,905]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462DFFC4FD86CE06FD0C54AE" collectingDate="2015-10-11" collectionCode="DB, PRE" collectorName="Steyn" country="South Africa" elevation="813" location="Namaqualand District" pageId="10" pageNumber="111" specimenCount="1">
<location id="8E806000462DFFC4FD86CE06FCB9548A" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FD86CE06FCB9548A" box="[548,800,880,905]" country="South Africa" name="Namaqualand District" pageId="10" pageNumber="111">Namaqualand District</location>
. On road
<locationDeviation id="9A522FF9462DFFC4FC32CE06FAD6548A" box="[912,1359,880,905]" firstLocation="Springbok" pageId="10" pageNumber="111" secondLocation="Spektakel Pass">between Springbok and Spektakel Pass</locationDeviation>
,
<quantity id="4CA79B3E462DFFC4FAFECE06FA3E548A" box="[1372,1447,880,905]" metricMagnitude="2" metricUnit="m" metricValue="8.129999999999999" pageId="10" pageNumber="111" unit="m" value="813.0">
<elevation id="0072D1E8462DFFC4FAFECE06FA3E548A" box="[1372,1447,880,905]" metricMagnitude="2" metricUnit="m" metricValue="8.129999999999999" pageId="10" pageNumber="111" unit="m" value="813.0">813 m</elevation>
</quantity>
, (2917
<collectionCode id="ED4EAE1E462DFFC4FF6ECEE2FF6D54AE" box="[204,244,916,941]" pageId="10" pageNumber="111">DB</collectionCode>
),
<date id="FFE1101B462DFFC4FEA4CEE2FE5B54AE" box="[262,450,916,941]" pageId="10" pageNumber="111" value="2015-10-11">
<collectingDate id="EFA5E9F3462DFFC4FEA4CEE2FE5B54AE" box="[262,450,916,941]" pageId="10" pageNumber="111" value="2015-10-11">11 October 2015</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FE69CEE2FDDF54AE" box="[459,582,916,941]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FE69CEE2FD9154AE" box="[459,520,916,941]" pageId="10" pageNumber="111">Steyn</collectorName>
2139
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FDF1CEE2FD1554AE" box="[595,652,916,941]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B373C86462DFFC4FD3DCEE2FAB8531A" collectingDate="1830-11-12" collectingDateMax="2001-09-22" collectingDateMin="1830-11-12" collectionCode="AC, NBG, CC, NMB, PRE, Z, P" collectorName="Mucina &amp; Santos &amp; Eenkokerboom &amp; Schlechter &amp; Without &amp; Ezelsfontein &amp; Roodeberg &amp; Drege" country="South Africa" elevation="1420" latitude="-30.312471" location="Leliefontein" longLatPrecision="1" longitude="18.078474" pageId="10" pageNumber="111" specimenCount="1" stateProvince="Western Cape">
<location id="8E806000462DFFC4FD3DCEE2FCB254AE" LSID="urn:lsid:plazi:treatment:03F687CD462FFFC4FF2AC9A7FD3B525E:8E806000462DFFC4FD3DCEE2FCB254AE" box="[671,811,916,941]" country="South Africa" latitude="-30.312471" longLatPrecision="1" longitude="18.078474" name="Leliefontein" pageId="10" pageNumber="111" stateProvince="Western Cape">Leliefontein</location>
, pass W of settlement.
<geoCoordinate id="EE6B501C462DFFC4FB84CEE2FB5854AE" box="[1062,1217,916,941]" degrees="30" direction="south" minutes="18" orientation="latitude" pageId="10" pageNumber="111" precision="1" seconds="44.9" value="-30.312471">30°1844.9”S</geoCoordinate>
,
<geoCoordinate id="EE6B501C462DFFC4FB6ECEE2FAF054AE" box="[1228,1385,916,941]" degrees="18" direction="east" minutes="04" orientation="longitude" pageId="10" pageNumber="111" precision="1" seconds="42.5" value="18.078474">18°0442.5”E</geoCoordinate>
,
<quantity id="4CA79B3E462DFFC4FAD1CEE2FF3954D2" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="10" pageNumber="111" unit="m" value="1420.0">
<elevation id="0072D1E8462DFFC4FAD1CEE2FF3954D2" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="10" pageNumber="111" unit="m" value="1420.0">1420 m</elevation>
</quantity>
, (3018
<collectionCode id="ED4EAE1E462DFFC4FF4DCECEFE8054D2" box="[239,281,952,977]" country="USA" lsid="urn:lsid:biocol.org:col:12582" name="Amherst College, Beneski Museum of Natural History" pageId="10" pageNumber="111" type="University or college">AC</collectionCode>
),
<date id="FFE1101B462DFFC4FE8ECECEFD9054D2" box="[300,521,952,977]" pageId="10" pageNumber="111" value="2001-09-22">
<collectingDate id="EFA5E9F3462DFFC4FE8ECECEFD9054D2" box="[300,521,952,977]" pageId="10" pageNumber="111" value="2001-09-22">22 September 2001</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FDB6CECEFCA254D2" box="[532,827,952,977]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FDB6CECEFDF354D2" box="[532,618,952,977]" pageId="10" pageNumber="111">Mucina</collectorName>
&amp;
<collectorName id="26AA530D462DFFC4FD2FCECEFD4154D2" box="[653,728,952,977]" pageId="10" pageNumber="111">Santos</collectorName>
7257/22
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FCE8CECEFC1254D2" box="[842,907,952,977]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="10" pageNumber="111" type="Herbarium">NBG</collectionCode>
).
<collectingRegion id="499BF839462DFFC4FC3CCECEFBA754D2" box="[926,1086,952,977]" country="South Africa" name="Western Cape" pageId="10" pageNumber="111">Western Cape</collectingRegion>
:
<collectorName id="26AA530D462DFFC4FBEACECEFB6F54D2" box="[1096,1270,952,977]" pageId="10" pageNumber="111">Eenkokerboom</collectorName>
,
<quantity id="4CA79B3E462DFFC4FAA0CECEFADD54D2" box="[1282,1348,952,977]" metricMagnitude="2" metricUnit="m" metricValue="2.7432" pageId="10" pageNumber="111" unit="ft" value="900.0">
<elevation id="0072D1E8462DFFC4FAA0CECEFADD54D2" box="[1282,1348,952,977]" metricMagnitude="2" metricUnit="m" metricValue="2.7432" pageId="10" pageNumber="111" unit="ft" value="900.0">900 ft</elevation>
</quantity>
[
<quantity id="4CA79B3E462DFFC4FAF0CECEFA0654D2" box="[1362,1439,952,977]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="10" pageNumber="111" unit="m" value="275.0">
<elevation id="0072D1E8462DFFC4FAF0CECEFA0654D2" box="[1362,1439,952,977]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="10" pageNumber="111" unit="m" value="275.0">275 m</elevation>
</quantity>
], (3018
<collectionCode id="ED4EAE1E462DFFC4FF69CEAAFF6A54F6" box="[203,243,988,1013]" country="Australia" name="CSIRO Canberra Rhizobium Collection" pageId="10" pageNumber="111">CC</collectionCode>
),
<date id="FFE1101B462DFFC4FEA5CEAAFE4154F6" box="[263,472,988,1013]" pageId="10" pageNumber="111" value="1897-09-03">
<collectingDate id="EFA5E9F3462DFFC4FEA5CEAAFE4154F6" box="[263,472,988,1013]" pageId="10" pageNumber="111" value="1897-09-03">3 September 1897</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FE46CEAAFD3F54F6" box="[484,678,988,1013]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FE46CEAAFDC354F6" box="[484,602,988,1013]" pageId="10" pageNumber="111">Schlechter</collectorName>
11061
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FD14CEAAFD6354F6" box="[694,762,988,1013]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="10" pageNumber="111">NMB</collectionCode>
,
<collectionCode id="ED4EAE1E462DFFC4FCA4CEAAFCA554F6" box="[774,828,988,1013]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="10" pageNumber="111" type="Herbarium">PRE</collectionCode>
,
<collectionCode id="ED4EAE1E462DFFC4FCEBCEAAFCC754F6" box="[841,862,988,1013]" country="Switzerland" name="Universität Zürich" pageId="10" pageNumber="111" type="Herbarium">Z</collectionCode>
).
<collectorName id="26AA530D462DFFC4FCD0CEAAFC5454F6" box="[882,973,988,1013]" pageId="10" pageNumber="111">Without</collectorName>
precise locality: [Illegible] zw [zwischen]
<collectorName id="26AA530D462DFFC4FF2AC976FE8D531A" box="[136,276,1024,1049]" pageId="10" pageNumber="111">Ezelsfontein</collectorName>
u [und]
<collectorName id="26AA530D462DFFC4FED6C976FE68531A" box="[372,497,1024,1049]" pageId="10" pageNumber="111">Roodeberg</collectorName>
,
<quantity id="4CA79B3E462DFFC4FE5EC976FD0B531A" box="[508,658,1024,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.143" metricValueMax="1.2192" metricValueMin="1.0668" pageId="10" pageNumber="111" unit="ft" value="3750.0" valueMax="4000.0" valueMin="3500.0">
<elevation id="0072D1E8462DFFC4FE5EC976FD0B531A" box="[508,658,1024,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.143" metricValueMax="1.2192" metricValueMin="1.0668" pageId="10" pageNumber="111" unit="ft" value="3750.0" valueMax="4000.0" valueMin="3500.0">35004000 ft</elevation>
</quantity>
[
<quantity id="4CA79B3E462DFFC4FD02C976FCD8531A" box="[672,833,1024,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.14" metricValueMax="1.22" metricValueMin="1.06" pageId="10" pageNumber="111" unit="m" value="1140.0" valueMax="1220.0" valueMin="1060.0">
<elevation id="0072D1E8462DFFC4FD02C976FCD8531A" box="[672,833,1024,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.14" metricValueMax="1.22" metricValueMin="1.06" pageId="10" pageNumber="111" unit="m" value="1140.0" valueMax="1220.0" valueMin="1060.0">10601220 m</elevation>
</quantity>
],
<date id="FFE1101B462DFFC4FCF7C976FBA9531A" box="[853,1072,1024,1049]" pageId="10" pageNumber="111" value="1830-11-12">
<collectingDate id="EFA5E9F3462DFFC4FCF7C976FBA9531A" box="[853,1072,1024,1049]" pageId="10" pageNumber="111" value="1830-11-12">12 November 1830</collectingDate>
</date>
,
<emphasis id="B92BEAC9462DFFC4FB99C976FB6D531A" box="[1083,1268,1024,1049]" italics="true" pageId="10" pageNumber="111">
<collectorName id="26AA530D462DFFC4FB99C976FB18531A" box="[1083,1153,1024,1049]" pageId="10" pageNumber="111">Drège</collectorName>
2431, s.n.
</emphasis>
(
<collectionCode id="ED4EAE1E462DFFC4FAA7C976FA81531A" box="[1285,1304,1024,1049]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="10" pageNumber="111" type="Herbarium">P</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE036DB462DFFC4FF1FC952FE9753F2" blockId="10.[136,1452,160,1373]" pageId="10" pageNumber="111">
<emphasis id="B92BEAC9462DFFC4FF1FC952FE90533E" bold="true" box="[189,265,1060,1085]" pageId="10" pageNumber="111">Notes:</emphasis>
—The
<typeStatus id="54E48879462DFFC4FEFBC952FE10533E" box="[345,393,1060,1085]" pageId="10" pageNumber="111">type</typeStatus>
specimen of
<taxonomicName id="4C5F4D58462DFFC4FD81C952FD15533E" authorityName="Obermeyer" authorityYear="1937" baseAuthorityName="S. Moore" box="[547,652,1060,1085]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="111" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC9462DFFC4FD81C952FD15533E" box="[547,652,1060,1085]" italics="true" pageId="10" pageNumber="111">A. scullyi</emphasis>
</taxonomicName>
is very distinct and has lax inflorescences with bracts ending in short, broadly triangular primary spines with a compound central primary spine (usually two pairs of short prickles on margin), and villose leaf and bract indumentum. However, in recent collections of the species it is clear that these characters are variable as specimens occasionally have simple or compound central primary spines (
<emphasis id="B92BEAC9462DFFC4FB43C9E6FAC553AA" box="[1249,1372,1168,1193]" italics="true" pageId="10" pageNumber="111">Steyn 2139</emphasis>
) or the spines could vary between shortly triangular to almost needle-shaped. However, compound, broadly triangular spines is the norm.
</paragraph>
<paragraph id="8BE036DB462DFFC4FF1FC98AFD3B525E" blockId="10.[136,1452,160,1373]" pageId="10" pageNumber="111">
<emphasis id="B92BEAC9462DFFC4FF1FC98AFE785216" box="[189,481,1276,1301]" italics="true" pageId="10" pageNumber="111">Mucina &amp; Santos 7257/22</emphasis>
and
<emphasis id="B92BEAC9462DFFC4FDB7C98AFD015216" box="[533,664,1276,1301]" italics="true" pageId="10" pageNumber="111">Drège 2431</emphasis>
are included in
<taxonomicName id="4C5F4D58462DFFC4FCEAC98AFC365216" authorityName="Obermeyer" authorityYear="1937" baseAuthorityName="S. Moore" box="[840,943,1276,1301]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="111" phylum="Tracheophyta" rank="species" species="scullyi">
<emphasis id="B92BEAC9462DFFC4FCEAC98AFC365216" box="[840,943,1276,1301]" italics="true" pageId="10" pageNumber="111">A. scullyi</emphasis>
</taxonomicName>
as they have the typical bract shape (obovate) and the latter also has antrorse, villose hairs on the bracts, but both specimens have leaves with a strigose indumentum and a distinct petiole-like base (as in
<taxonomicName id="4C5F4D58462DFFC4FD8AC832FD0B525E" authorityName="H. M. Steyn" authorityYear="2017" baseAuthorityName="H. M. Steyn" baseAuthorityYear="2017" box="[552,658,1348,1373]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthopsis" kingdom="Plantae" order="Lamiales" pageId="10" pageNumber="111" phylum="Tracheophyta" rank="species" species="glabra">
<emphasis id="B92BEAC9462DFFC4FD8AC832FD0B525E" box="[552,658,1348,1373]" italics="true" pageId="10" pageNumber="111">A. glabra</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>