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<document id="06E67BE606B7F36D7CF78D614E07185A" ID-DOI="10.5852/ejt.2024.956.2657" ID-ISSN="2118-9773" ID-ZooBank="9DCDD521-909D-4436-8495-086A11C50DA0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725988271530" checkinUser="plazi" docAuthor="Karimbumkara, Seena Narayanan &amp; Priyadarsanan, Dharma Rajan" docDate="2024" docId="557187D2FFC589329D91FC85FB35F9B6" docLanguage="en" docName="EJT.2024.956.1-61.pdf" docOrigin="European Journal of Taxonomy 956" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onitis visthara Karimbumkara &amp; Priyadarsanan, 2024, sp. nov." docType="treatment" docUuid="0E89DF71-6A9D-4E88-8C49-84281CC84C2A" docUuidSource="ZooBank" docVersion="1" lastPageNumber="53" masterDocId="A948FFAAFFF689069F01FFB1FFF9FF8A" masterDocTitle="Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India" masterLastPageNumber="61" masterPageNumber="1" pageNumber="52" updateTime="1725988442359" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EDF1F21CCF2FE0EB1373BB958621BFF4">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:namePart id="40B2CBA7FBD090612DDF8D98EF77B1D7">Karimbumkara, Seena Narayanan</mods:namePart>
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<mods:affiliation id="C5192386A4CABF0B34A138B575DBF3E6">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
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<mods:namePart id="4DAEB2DC4F7A704969ABE02ACF1C3AD3">Priyadarsanan, Dharma Rajan</mods:namePart>
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<mods:affiliation id="EDC88BA78116A420B0472E4AEED79256">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
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<mods:date id="53FB2A47A74055CE5059275B4A0B2B23">2024</mods:date>
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<paragraph id="DD6736C4FFC589359D91FC85FC5DFCC4" blockId="51.[367,1222,819,917]" box="[656,932,819,846]" pageId="51" pageNumber="52">
<heading id="862F81A8FFC589359D91FC85FC5DFCC4" box="[656,932,819,846]" centered="true" fontSize="11" level="2" pageId="51" pageNumber="52" reason="2">
<taxonomicName id="1AD84D47FFC589359D91FC85FCBBFCC4" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[656,834,819,846]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov.">
<emphasis id="EFACEAD6FFC589359D91FC85FCBBFCC4" bold="true" box="[656,834,819,846]" italics="true" pageId="51" pageNumber="52">Onitis visthara</emphasis>
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<taxonomicNameLabel id="F49F57ADFFC589359C4BFC85FC5DFCC4" box="[842,932,820,846]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="95C2654FFFC589359D22FCCBFBE9FC1E" box="[547,1040,890,917]" pageId="51" pageNumber="52" type="description">
<paragraph id="DD6736C4FFC589359D22FCCBFBE9FC1E" blockId="51.[367,1222,819,917]" box="[547,1040,890,917]" pageId="51" pageNumber="52">
<figureCitation id="45E32A41FFC589359D22FCCBFD5DFC1E" box="[547,676,890,916]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="51" pageNumber="52">Figs 8MP</figureCitation>
,
<figureCitation id="45E32A41FFC589359DB1FCCBFD2AFC1E" box="[688,723,890,916]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="51" pageNumber="52">9E</figureCitation>
,
<figureCitation id="45E32A41FFC589359DDEFCCBFCD7FC1E" box="[735,814,890,916]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="51" pageNumber="52">12EF</figureCitation>
,
<figureCitation id="45E32A41FFC589359C3BFCCBFC92FC1E" box="[826,875,890,916]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="51" pageNumber="52">14E</figureCitation>
,
<figureCitation id="45E32A41FFC589359C76FCCBFC53FC1F" box="[887,938,890,917]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="51" pageNumber="52">15E</figureCitation>
,
<figureCitation id="45E32A41FFC589359CB4FCCBFC1EFC1E" box="[949,999,890,916]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="51" pageNumber="52">16E</figureCitation>
,
<figureCitation id="45E32A41FFC589359CF2FCCBFBE9FC1E" box="[1011,1040,890,916]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." pageId="51" pageNumber="52">20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFC589359FBCFC0AFD90FA7B" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="DD6736C4FFC589359FBCFC0AFE3FFC5F" blockId="51.[189,454,955,981]" box="[189,454,955,981]" pageId="51" pageNumber="52">
<heading id="862F81A8FFC589359FBCFC0AFE3FFC5F" bold="true" box="[189,454,955,981]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="EFACEAD6FFC589359FBCFC0AFE3FFC5F" bold="true" box="[189,454,955,981]" pageId="51" pageNumber="52">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFC589359FBCFC5BFD90FA7B" blockId="51.[189,1399,1002,1521]" pageId="51" pageNumber="52">
<taxonomicName id="1AD84D47FFC589359FBCFC5BFE96FB8E" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[189,367,1002,1028]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov.">
<emphasis id="EFACEAD6FFC589359FBCFC5BFE96FB8E" box="[189,367,1002,1028]" italics="true" pageId="51" pageNumber="52">Onitis visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFC589359E7BFC5AFE21FB8F" box="[378,472,1003,1029]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
belongs to Group II; it shows affinity to
<taxonomicName id="1AD84D47FFC589359CD5FC5BFB6AFB8E" authorityName="Fabricius" authorityYear="1801" box="[980,1171,1002,1028]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFC589359CD5FC5BFB6AFB8E" box="[980,1171,1002,1028]" italics="true" pageId="51" pageNumber="52">Onitis philemon</emphasis>
</taxonomicName>
,
<taxonomicName id="1AD84D47FFC589359BA4FC5BFAEDFB8E" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1189,1300,1002,1028]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov">
<emphasis id="EFACEAD6FFC589359BA4FC5BFAEDFB8E" box="[1189,1300,1002,1028]" italics="true" pageId="51" pageNumber="52">O. kethai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFC589359A1FFC5AFA8FFB8F" box="[1310,1398,1003,1029]" pageId="51" pageNumber="52" rank="species">sp. nov</taxonomicNameLabel>
and
<taxonomicName id="1AD84D47FFC589359FF3FBBFFE52FBAD" authorityName="Lansberge, AIMB" authorityYear="1875" box="[242,427,1037,1064]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFC589359FF3FBBFFE52FBAD" box="[242,427,1037,1064]" italics="true" pageId="51" pageNumber="52">O. singhalensis</emphasis>
</taxonomicName>
as the metasternum is longitudinally grooved anteriorly and the clypeal margin feebly emarginate. The pronotum of
<emphasis id="EFACEAD6FFC589359D69FB81FCEEFBC0" box="[616,791,1072,1098]" italics="true" pageId="51" pageNumber="52">Onitisvisthara</emphasis>
is more convex and wider and has a smooth median line similar to that in
<taxonomicName id="1AD84D47FFC589359EC0FBE5FD8EFBE4" authorityName="Lansberge, AIMB" authorityYear="1875" box="[449,631,1108,1134]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFC589359EC0FBE5FD8EFBE4" box="[449,631,1108,1134]" italics="true" pageId="51" pageNumber="52">O. singhalensis</emphasis>
</taxonomicName>
but absent in
<taxonomicName id="1AD84D47FFC589359C22FBE5FC4CFBE4" authorityName="Fabricius" authorityYear="1801" box="[803,949,1108,1134]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFC589359C22FBE5FC4CFBE4" box="[803,949,1108,1134]" italics="true" pageId="51" pageNumber="52">O. philemon</emphasis>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFC589359CF3FBE5FBA6FBE4" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1010,1119,1108,1134]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="kethai">
<emphasis id="EFACEAD6FFC589359CF3FBE5FBA6FBE4" box="[1010,1119,1108,1134]" italics="true" pageId="51" pageNumber="52">O. kethai</emphasis>
</taxonomicName>
.
<emphasis id="EFACEAD6FFC589359B6EFBE5FAD9FBE4" box="[1135,1312,1108,1134]" italics="true" pageId="51" pageNumber="52">Onitisvisthara</emphasis>
can be differentiated from
<taxonomicName id="1AD84D47FFC589359EA0FBC6FDAFFB1B" authorityName="Lansberge, AIMB" authorityYear="1875" box="[417,598,1143,1169]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFC589359EA0FBC6FDAFFB1B" box="[417,598,1143,1169]" italics="true" pageId="51" pageNumber="52">O. singhalensis</emphasis>
</taxonomicName>
by the pronotal front angle which is sharp in the former while blunt or rather rounded in the latter (compare
<figureCitation id="45E32A41FFC589359D9FFB2BFCE3FB3E" box="[670,794,1178,1204]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="51" pageNumber="52">Fig. 9EF</figureCitation>
). The clypeus and vertex in
<taxonomicName id="1AD84D47FFC589359B6CFB2BFB0BFB3E" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1133,1266,1178,1204]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="visthara">
<emphasis id="EFACEAD6FFC589359B6CFB2BFB0BFB3E" box="[1133,1266,1178,1204]" italics="true" pageId="51" pageNumber="52">O. visthara</emphasis>
</taxonomicName>
are rugose while they are closely granular in
<taxonomicName id="1AD84D47FFC589359D4AFB0FFCF9FB5D" authorityName="Lansberge, AIMB" authorityYear="1875" box="[587,768,1213,1240]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFC589359D4AFB0FFCF9FB5D" box="[587,768,1213,1240]" italics="true" pageId="51" pageNumber="52">O. singhalensis</emphasis>
</taxonomicName>
. The genae of the former is gently rounded while that of the latter is sharply rounded. Profemur in the male of
<taxonomicName id="1AD84D47FFC589359C68FB50FC16FB70" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[873,1007,1248,1275]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="visthara">
<emphasis id="EFACEAD6FFC589359C68FB50FC16FB70" box="[873,1007,1248,1275]" italics="true" pageId="51" pageNumber="52">O. visthara</emphasis>
</taxonomicName>
is without tooth while there is a strong tooth towards the end of its anterior edge in the male of
<taxonomicName id="1AD84D47FFC589359CB3FAB5FB91FA94" authorityName="Lansberge, AIMB" authorityYear="1875" box="[946,1128,1284,1310]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFC589359CB3FAB5FB91FA94" box="[946,1128,1284,1310]" italics="true" pageId="51" pageNumber="52">O. singhalensis</emphasis>
</taxonomicName>
. The mid femur in the male of
<taxonomicName id="1AD84D47FFC589359E20FA96FE5EFACB" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[289,423,1319,1345]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="visthara">
<emphasis id="EFACEAD6FFC589359E20FA96FE5EFACB" box="[289,423,1319,1345]" italics="true" pageId="51" pageNumber="52">O. visthara</emphasis>
</taxonomicName>
is without a tooth near the middle of its posterior edge, but with two teeth at the extremity, a sharp one on the lower edge and a blunt one above; while in the male of
<taxonomicName id="1AD84D47FFC589359B90FAFBFABCFAEE" authorityName="Lansberge, AIMB" authorityYear="1875" box="[1169,1349,1354,1380]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFC589359B90FAFBFABCFAEE" box="[1169,1349,1354,1380]" italics="true" pageId="51" pageNumber="52">O. singhalensis</emphasis>
</taxonomicName>
, the middle femur has a strong tooth near the middle of its posterior edge and another tooth at the extremity. The middle tibia in male
<taxonomicName id="1AD84D47FFC589359EE0FA20FD9DFA20" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[481,612,1424,1451]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="visthara">
<emphasis id="EFACEAD6FFC589359EE0FA20FD9DFA20" box="[481,612,1424,1451]" italics="true" pageId="51" pageNumber="52">O. visthara</emphasis>
</taxonomicName>
is slender at the base and they curve and angularly dilate before the middle; in
<taxonomicName id="1AD84D47FFC589359E3BFA05FE17FA44" authorityName="Lansberge, AIMB" authorityYear="1875" box="[314,494,1460,1486]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFC589359E3BFA05FE17FA44" box="[314,494,1460,1486]" italics="true" pageId="51" pageNumber="52">O. singhalensis</emphasis>
</taxonomicName>
, the middle tibia is slender at the base, less curved, abruptly, angularly dilated before the middle (compare 16EF).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFC589359FBCF9A9FBC1F886" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="DD6736C4FFC589359FBCF9A9FE95F9B8" blockId="51.[189,364,1560,1586]" box="[189,364,1560,1586]" pageId="51" pageNumber="52">
<heading id="862F81A8FFC589359FBCF9A9FE95F9B8" bold="true" box="[189,364,1560,1586]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="EFACEAD6FFC589359FBCF9A9FE95F9B8" bold="true" box="[189,364,1560,1586]" pageId="51" pageNumber="52">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFC589359FEBF9F6FEA5F9EB" blockId="51.[189,1399,1607,1704]" box="[234,348,1607,1633]" pageId="51" pageNumber="52">
<emphasis id="EFACEAD6FFC589359FEBF9F6FEA5F9EB" bold="true" box="[234,348,1607,1633]" pageId="51" pageNumber="52">
<typeStatus id="02638866FFC589359FEBF9F6FEA5F9EB" box="[234,348,1607,1633]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFC589359FBCF9DBFB56F92D" blockId="51.[189,1399,1607,1704]" pageId="51" pageNumber="52">
<materialsCitation id="6DB03C99FFC589359FBCF9DBFB56F92D" collectingDate="2015-01-04" collectionCode="NBAIR, NIM" collectorName="Mohammed Ashraf K." country="India" location="Bangalore" municipality="Hesaraghatta" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka" typeStatus="holotype">
<collectingCountry id="A5CF7654FFC589359FBCF9DBFEF7F90E" box="[189,270,1642,1668]" name="India" pageId="51" pageNumber="52">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFC589359E22F9DAFEC5F90F" box="[291,316,1643,1669]" count="1" pageId="51" pageNumber="52" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFC589359E46F9DBFE3AF90E" box="[327,451,1642,1668]" country="India" name="Karnataka" pageId="51" pageNumber="52">Karnataka</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFC589359ECDF9DBFD91F90E" box="[460,616,1642,1668]" pageId="51" pageNumber="52">Hesaraghatta</collectingMunicipality>
,
<location id="D807601FFFC589359D73F9DBFD17F90E" LSID="urn:lsid:plazi:treatment:557187D2FFC589329D91FC85FB35F9B6:D807601FFFC589359D73F9DBFD17F90E" box="[626,750,1642,1668]" country="India" municipality="Hesaraghatta" name="Bangalore" pageId="51" pageNumber="52" stateProvince="Karnataka">Bangalore</location>
, hand-picked from cattle dung;
<date id="A9661004FFC589359B58F9DBFB26F90F" box="[1113,1247,1642,1669]" pageId="51" pageNumber="52" value="2015-01-04">
<collectingDate id="B922E9ECFFC589359B58F9DBFB26F90F" box="[1113,1247,1642,1669]" pageId="51" pageNumber="52" value="2015-01-04">4 Jan. 2015</collectingDate>
</date>
,
<collectorName id="702D5312FFC589359BE8F9DBFEC8F922" pageId="51" pageNumber="52">Mohammed Ashraf K.</collectorName>
leg.; specimen deposited at
<collectionCode id="BBC9AE01FFC589359D7CF93CFD22F92D" box="[637,731,1677,1703]" pageId="51" pageNumber="52">NBAIR</collectionCode>
,
<collectionCode id="BBC9AE01FFC589359DE6F93CFCD8F92D" box="[743,801,1677,1703]" country="France" name="Museum d'histoire naturelle de Nîmes" pageId="51" pageNumber="52" type="Herbarium">NIM</collectionCode>
/
<collectionCode id="BBC9AE01FFC589359C2BF93CFC7FF92D" box="[810,902,1677,1703]" pageId="51" pageNumber="52">NBAIR</collectionCode>
/COL/ONIT/H-271023C.
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFC589359FEBF97EFEA5F963" blockId="51.[189,1080,1743,1804]" box="[234,348,1743,1769]" pageId="51" pageNumber="52">
<emphasis id="EFACEAD6FFC589359FEBF97EFEA5F963" bold="true" box="[234,348,1743,1769]" pageId="51" pageNumber="52">
<typeStatus id="02638866FFC589359FEBF97EFEA5F963" box="[234,348,1743,1769]" pageId="51" pageNumber="52" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFC589359FBCF943FBC1F886" blockId="51.[189,1080,1743,1804]" box="[189,1080,1778,1804]" pageId="51" pageNumber="52">
COUNTRY •
<materialsCitation id="6DB03C99FFC589359E60F943FBCDF886" box="[353,1076,1778,1804]" collectionCode="AIMB" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="CBDEFD4DFFC589359E60F943FE83F886" box="[353,378,1778,1804]" count="1" pageId="51" pageNumber="52" type="male"></specimenCount>
; same collection details as
<typeStatus id="02638866FFC589359DB8F943FCDAF886" box="[697,803,1778,1804]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
;
<collectionCode id="BBC9AE01FFC589359C2CF943FC82F886" box="[813,891,1778,1804]" pageId="51" pageNumber="52">AIMB</collectionCode>
/Co/Sc1000789
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFC589359FBCF882FEDCF848" pageId="51" pageNumber="52" type="etymology">
<paragraph id="DD6736C4FFC589359FBCF882FEBDF8C7" blockId="51.[189,324,1843,1869]" box="[189,324,1843,1869]" pageId="51" pageNumber="52">
<heading id="862F81A8FFC589359FBCF882FEBDF8C7" bold="true" box="[189,324,1843,1869]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="EFACEAD6FFC589359FBCF882FEBDF8C7" bold="true" box="[189,324,1843,1869]" pageId="51" pageNumber="52">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFC589359FBCF8D3FEDCF848" blockId="51.[189,1399,1890,1986]" pageId="51" pageNumber="52">
The pronotal and elytral measurements of this species are almost the same, giving it a broader rectangular appearance while comparing with other species, thus the name
<taxonomicName id="1AD84D47FFC589359CCCF834FBD4F815" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[973,1069,1925,1951]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="visthara">
<emphasis id="EFACEAD6FFC589359CCCF834FBD4F815" box="[973,1069,1925,1951]" italics="true" pageId="51" pageNumber="52">visthara</emphasis>
</taxonomicName>
which means expanse in Sanskrit.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFC289329FBCFEB9FB35F9B6" pageId="52" pageNumber="53" type="description">
<paragraph id="DD6736C4FFC289329FBCFEB9FEB7FEA8" blockId="52.[189,334,264,290]" box="[189,334,264,290]" pageId="52" pageNumber="53">
<heading id="862F81A8FFC289329FBCFEB9FEB7FEA8" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="EFACEAD6FFC289329FBCFEB9FEB7FEA8" bold="true" box="[189,334,264,290]" pageId="52" pageNumber="53">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFC289329FBCFE86FEB6FEFF" blockId="52.[189,1399,311,373]" pageId="52" pageNumber="53">MEASUREMENTS (in mm). TL= 1718; BW =910; PL=89; PW=9.510.5; EL=89.5; HL=44.5; HW=55.5.</paragraph>
<paragraph id="DD6736C4FFC289329FEBFE2DFE4BFE3C" blockId="52.[189,1399,412,1177]" box="[234,434,412,438]" pageId="52" pageNumber="53">
<emphasis id="EFACEAD6FFC289329FEBFE2DFED3FE3C" bold="true" box="[234,298,412,438]" pageId="52" pageNumber="53">Male</emphasis>
(
<typeStatus id="02638866FFC289329E38FE2DFE53FE3C" box="[313,426,412,438]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
)
</paragraph>
<paragraph id="DD6736C4FFC289329FBCFE0EFD97FB13" blockId="52.[189,1399,412,1177]" pageId="52" pageNumber="53">
Brownish black, not shining, pronotum bronzy black, elytra bronzy red, ventral side blackishred; antennae and mouth reddish; oval, not very convex. Clypeus (
<figureCitation id="45E32A41FFC289329B14FE53FB7DFE76" box="[1045,1156,482,508]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="52" pageNumber="53">Fig. 14E</figureCitation>
) rugulose, anterior margin parabolic, slightly straight in the middle; clypeal carina not much elevated, bears a few punctures; clypeo-frontal carina not widely interrupted, the tubercle behind blunt, rounded and closer to carina, bears a few punctures; vertex rugulose in the middle and sides more or less smooth with a few scattered granules; genae broad and smooth, slightly reflexed anteriorly with a few scattered punctures. Pronotum strongly and fairly closely punctured, scattered in the middle, more strongly and closely pitted anteriorly, lightly and closely at the base, with a slight median groove starting above the basal foveae and extending to anterior margin as a thin smooth line; front angles moderately sharp, almost right angled, hind angle obsolete, lateral margin straight anteriorly, strongly curved or rounded in the middle and bisinuate behind; base of pronotum bluntly angulate; basal foveae rounded and concave. Elytra moderately shining, lightly and broadly striate, striae smooth, intervals almost flat, strongly and closely punctured; sutural interval convex and slightly elevated than the rest, finely and deeply punctured. Pygidium not shining, lightly and uniformly punctured. Metasternum with deep broad longitudinal groove; metasternal shield smooth in the middle with fine scattered punctures and with strong close aciculate punctures anterio-laterally; sides of metasternum with setigerous granules. Femora smooth with some punctures and hairs on anterior part near the tibial joint. Protibia with four teeth, with the tip extended to a curved process; teeth very short, blunt, the terminal tooth trapezoid. Profemur without teeth. Mid femur has a pair of sharp teeth near the end of the lower edge, and a blunt curved process towards the middle; hind trochanter with a small blunt tooth.
</paragraph>
<paragraph id="DD6736C4FFC289329FBCFB71FE6DFACE" blockId="52.[189,1399,1216,1348]" pageId="52" pageNumber="53">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFC289329EB0FB71FDCCFB50" box="[433,565,1216,1242]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="52" pageNumber="53">Fig. 8MP</figureCitation>
). LP=3, Lp=2.5; BP=0.75, BpB=0.5, BpT=0.25. Phallobase slightly curved, longer than parameres, the latter curved at the tip, with an opening on the ventral side at the point of articulation with the phallobase. The base of the opening is with a cup-shaped extension which is curved inwards.
</paragraph>
<paragraph id="DD6736C4FFC289329FEBFADAFEBCFA0F" blockId="52.[189,325,1387,1448]" box="[234,325,1387,1413]" pageId="52" pageNumber="53">
<emphasis id="EFACEAD6FFC289329FEBFADAFEBCFA0F" bold="true" box="[234,325,1387,1413]" pageId="52" pageNumber="53">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFC289329FBCFA3FFEC0FA22" blockId="52.[189,325,1387,1448]" box="[189,313,1422,1448]" pageId="52" pageNumber="53">Unknown.</paragraph>
<paragraph id="DD6736C4FFC289329FBCFA61FEE4FA60" blockId="52.[189,285,1488,1514]" box="[189,285,1488,1514]" pageId="52" pageNumber="53">
<heading id="862F81A8FFC289329FBCFA61FEE4FA60" bold="true" box="[189,285,1488,1514]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="EFACEAD6FFC289329FBCFA61FEE4FA60" bold="true" box="[189,285,1488,1514]" pageId="52" pageNumber="53">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFC289329FBCFA4FFB35F9B6" blockId="52.[189,1398,1534,1596]" pageId="52" pageNumber="53">The specimens were collected from cattle dung in open grassland near Hesaraghatta lake, the vegetation of which is undergoing rapid changes due to human activities and ecological succession.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,495 @@
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<mods:title id="D5251442EDAC9D2D0B9097342E2CE2CB">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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</mods:location>
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<mods:identifier id="BC62B38528A3509182FB9B201B2E8438" type="DOI">10.5852/ejt.2024.956.2657</mods:identifier>
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<treatment id="557187D2FFD0892F9D34FC85FB77FB1B" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B" httpUri="http://treatment.plazi.org/id/557187D2FFD0892F9D34FC85FB77FB1B" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39">
<subSubSection id="95C2654FFFD089209D34FC85FC06FCC4" box="[565,1023,820,846]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="DD6736C4FFD089209D34FC85FC06FCC4" blockId="38.[560,1027,820,882]" box="[565,1023,820,846]" pageId="38" pageNumber="39">
<heading id="862F81A8FFD089209D34FC85FC06FCC4" box="[565,1023,820,846]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="39" reason="2">
<taxonomicName id="1AD84D47FFD089209D34FC85FC06FCC4" ID-CoL="49KKQ" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[565,1023,820,846]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFD089209D34FC85FCB0FCC4" bold="true" box="[565,841,820,846]" italics="true" pageId="38" pageNumber="39">Onitis punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFD089209C51FC85FC06FCC4" author="Janssens A." box="[848,1023,820,846]" pageId="38" pageNumber="39" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD089209D31FCE6FC57FC38" pageId="38" pageNumber="39" type="description">
<paragraph id="DD6736C4FFD089209D31FCE6FBFAFCFB" blockId="38.[560,1027,820,882]" box="[560,1027,855,882]" pageId="38" pageNumber="39">
<figureCitation id="45E32A41FFD089209D31FCE6FD48FCFB" box="[560,689,855,881]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="38" pageNumber="39">Figs 6MP</figureCitation>
,
<figureCitation id="45E32A41FFD089209DBCFCE6FD18FCFB" box="[701,737,855,881]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="38" pageNumber="39">9B</figureCitation>
,
<figureCitation id="45E32A41FFD089209DEFFCE6FCE5FCFB" box="[750,796,855,881]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="38" pageNumber="39">10F</figureCitation>
,
<figureCitation id="45E32A41FFD089209C29FCE6FCA2FCFB" box="[808,859,855,881]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="38" pageNumber="39">14B</figureCitation>
,
<figureCitation id="45E32A41FFD089209C66FCE6FC63FCF8" box="[871,922,855,882]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="38" pageNumber="39">15B</figureCitation>
,
<figureCitation id="45E32A41FFD089209CA6FCE6FC23FCFB" box="[935,986,855,881]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="38" pageNumber="39">16B</figureCitation>
,
<figureCitation id="45E32A41FFD089209CE7FCE6FBFAFCFB" box="[998,1027,855,881]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." pageId="38" pageNumber="39">19</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFD089209FBCFC28FC57FC38" blockId="38.[189,942,920,947]" box="[189,942,920,947]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209FBCFC28FE32FC39" box="[189,459,921,947]" italics="true" pageId="38" pageNumber="39">Onitispunctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFD089209ED3FC28FD5FFC38" author="Janssens A." box="[466,678,920,947]" pageId="38" pageNumber="39" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 19</bibRefCitation>
(original description).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD089209FBCFC6BFC6FFBB0" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="DD6736C4FFD089209FBCFC6BFC6FFBB0" blockId="38.[189,1399,986,1083]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="FDC811EAFFD089209FBCFC6BFC6FFBB0" pageId="38" pageNumber="39">
<taxonomicName id="1AD84D47FFD089209FBCFC6BFE32FC7E" ID-CoL="49KKQ" authorityName="Janssens" authorityYear="1937" box="[189,459,986,1012]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFD089209FBCFC6BFE32FC7E" box="[189,459,986,1012]" italics="true" pageId="38" pageNumber="39">Onitis punctatostriatus</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFD089209EE6FC6BFD3DFC7F" author="Balthasar V." box="[487,708,986,1013]" page="35" pageId="38" pageNumber="39" year="1963">
<bibRefCitation id="B9494B35FFD089209EE6FC6BFD3DFC7F" author="Balthasar V." box="[487,708,986,1013]" pageId="38" pageNumber="39" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar 1963: 35</bibRefCitation>
</treatmentCitation>
(monograph). —
<treatmentCitation id="5C7910D5FFD089209C96FC6BFB34FC7F" author="Bezdek A. &amp; Krell F. - T." box="[919,1229,986,1013]" page="159" pageId="38" pageNumber="39" year="2006">
<bibRefCitation id="B9494B35FFD089209C96FC6BFB34FC7F" author="Bezdek A. &amp; Krell F. - T." box="[919,1229,986,1013]" pageId="38" pageNumber="39" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). — Siddiqui
<emphasis id="EFACEAD6FFD089209E7AFC4FFE47FB9D" box="[379,446,1021,1047]" italics="true" pageId="38" pageNumber="39">et. al.</emphasis>
2014: 300 (distribution). —
<treatmentCitation id="5C7910D5FFD089209C15FC4CFBD5FB9D" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[788,1068,1021,1048]" page="1037" pageId="38" pageNumber="39" year="2015">
<bibRefCitation id="B9494B35FFD089209C15FC4CFBD5FB9D" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[788,1068,1021,1048]" pageId="38" pageNumber="39" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFD089209C62FC4FFC66FB9D" box="[867,927,1021,1047]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFD089209BA1FC4CFA8FFB9D" author="Bezdek A." box="[1184,1398,1021,1047]" page="179" pageId="38" pageNumber="39" year="2016">
<bibRefCitation id="B9494B35FFD089209BA1FC4CFA8FFB9D" author="Bezdek A." box="[1184,1398,1021,1047]" pageId="38" pageNumber="39" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFD089209EB8FB91FD4AFBB0" author="Schoolmeesters P." box="[441,691,1056,1082]" pageId="38" pageNumber="39" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD089209FBCFBD0FC88FA9E" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="DD6736C4FFD089209FBCFBD0FE3FFBF1" blockId="38.[189,454,1121,1147]" box="[189,454,1121,1147]" pageId="38" pageNumber="39">
<heading id="862F81A8FFD089209FBCFBD0FE3FFBF1" bold="true" box="[189,454,1121,1147]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="EFACEAD6FFD089209FBCFBD0FE3FFBF1" bold="true" box="[189,454,1121,1147]" pageId="38" pageNumber="39">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFD089209FBCFB20FC88FA9E" blockId="38.[189,1399,1168,1300]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209FBCFB20FE35FB21" box="[189,460,1169,1195]" italics="true" pageId="38" pageNumber="39">Onitispunctatostriatus</emphasis>
is similar to
<taxonomicName id="1AD84D47FFD089209D6AFB20FCF5FB20" authorityName="Arrow" authorityYear="1931" box="[619,780,1168,1195]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFD089209D6AFB20FCF5FB20" box="[619,780,1168,1195]" italics="true" pageId="38" pageNumber="39">O. subopacus</emphasis>
</taxonomicName>
in having frontal carina broadly interrupted and the metasternum smooth and shining behind with scattered punctures. It differs from the latter in having deep and broad elytral striae and the pronotum being strongly ocellate punctured. This species is reported for the first time from the state of
<collectingRegion id="1F1CF826FFD089209D1EFB4BFD96FA9E" box="[543,623,1274,1300]" country="India" name="Assam" pageId="38" pageNumber="39">Assam</collectingRegion>
in northeastern
<collectingCountry id="A5CF7654FFD089209C2CFB4BFC94FA9E" box="[813,877,1274,1300]" name="India" pageId="38" pageNumber="39">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD0892F9FBCFA8AFDF7FECD" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="DD6736C4FFD089209FBCFA8AFE15FADF" blockId="38.[189,492,1339,1365]" box="[189,492,1339,1365]" pageId="38" pageNumber="39">
<heading id="862F81A8FFD089209FBCFA8AFE15FADF" bold="true" box="[189,492,1339,1365]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="EFACEAD6FFD089209FBCFA8AFE15FADF" bold="true" box="[189,492,1339,1365]" pageId="38" pageNumber="39">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFD089209FEBFADBFEA5FA0E" blockId="38.[189,1399,1386,1517]" box="[234,348,1386,1412]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209FEBFADBFEA5FA0E" bold="true" box="[234,348,1386,1412]" pageId="38" pageNumber="39">
<typeStatus id="02638866FFD089209FEBFADBFEA5FA0E" box="[234,348,1386,1412]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFD089209FBCFA3CFEE2FA67" blockId="38.[189,1399,1386,1517]" pageId="38" pageNumber="39">
<materialsCitation id="6DB03C99FFD089209FBCFA3CFEE2FA67" collectionCode="IRSNB" collectorName="R. P. Verschraeghen &amp; A. Janssens" country="India" location="INDIA" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="A5CF7654FFD089209FBCFA3CFEF7FA2D" box="[189,270,1421,1447]" name="India" pageId="38" pageNumber="39">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFD089209E2AFA3CFEBDFA2D" box="[299,324,1421,1447]" count="1" pageId="38" pageNumber="39" type="male"></specimenCount>
; “Darjeeling 1904,
<collectorName id="702D5312FFD089209D2FFA3CFCEFFA2D" box="[558,790,1421,1447]" pageId="38" pageNumber="39">R.P. Verschraeghen</collectorName>
, cf. Mem. Mus. Hist. Nat. Belg., 2: Serie, fasc. 11, 1937, p. 48, Ex. Typis,
<collectorName id="702D5312FFD089209ED7FA01FD98FA41" box="[470,609,1456,1483]" pageId="38" pageNumber="39">A. Janssens</collectorName>
rev., 1936:
<taxonomicName id="1AD84D47FFD089209DF3FA00FB25FA40" authority="Janss., J. J. Gillet" authorityName="Janss., J. J. Gillet" box="[754,1244,1456,1483]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFD089209DF3FA00FBFBFA40" box="[754,1026,1457,1483]" italics="true" pageId="38" pageNumber="39">Onitis punctatostriatus</emphasis>
Janss., J. J. Gillet
</taxonomicName>
det., vend.”;
<collectionCode id="BBC9AE01FFD089209FBCFA62FEEEFA67" box="[189,279,1491,1517]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="39">IRSNB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFD089209FEBF9A4FEA5F9A5" blockId="38.[189,1398,1557,1654]" box="[234,348,1557,1583]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209FEBF9A4FEA5F9A5" bold="true" box="[234,348,1557,1583]" pageId="38" pageNumber="39">
<typeStatus id="02638866FFD089209FEBF9A4FEA5F9A5" box="[234,348,1557,1583]" pageId="38" pageNumber="39" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFD089209FBCF989FD3FF9FF" blockId="38.[189,1398,1557,1654]" pageId="38" pageNumber="39">
<materialsCitation id="6DB03C99FFD089209FBCF989FD3BF9FF" collectionCode="IRSNB" collectorName="Onitisspinipes Drury R. M. H. N. Belg." country="India" determinerName="J. Gillet" location="INDIA" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="A5CF7654FFD089209FBCF989FEF7F9D8" box="[189,270,1592,1618]" name="India" pageId="38" pageNumber="39">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFD089209E2DF988FEBAF9D9" box="[300,323,1593,1619]" count="1" pageId="38" pageNumber="39" type="female"></specimenCount>
; “
<collectorName id="702D5312FFD089209E60F989FCD2F9D8" box="[353,811,1592,1619]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209E60F989FDEAF9D8" box="[353,531,1592,1618]" italics="true" pageId="38" pageNumber="39">Onitisspinipes</emphasis>
Drury R.M.H.N. Belg.
</collectorName>
10. 640,
<emphasis id="EFACEAD6FFD089209CA1F988FBF9F9D8" box="[928,1024,1593,1618]" italics="true" pageId="38" pageNumber="39">spinipes</emphasis>
Drury;
<determinerName id="28BBDA00FFD089209B63F988FB3BF9D8" box="[1122,1218,1592,1619]" pageId="38" pageNumber="39">J. Gillet</determinerName>
det. 1923, Ex-Typis”; label data same as for male;
<collectionCode id="BBC9AE01FFD089209D69F9EAFD3BF9FF" box="[616,706,1627,1653]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="39">IRSNB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="DD6736C4FFD089209FEBF92DFD07F93D" blockId="38.[189,1398,1692,2001]" box="[234,766,1692,1719]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209FEBF92DFDBDF93C" bold="true" box="[234,580,1692,1718]" pageId="38" pageNumber="39">Additional material studied</emphasis>
(
<specimenCount id="CBDEFD4DFFD089209D55F92CFD0FF93D" box="[596,758,1693,1719]" count="10" pageId="38" pageNumber="39" type="generic">10 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFD0892F9FBCF971FDF7FECD" blockId="38.[189,1398,1692,2001]" lastBlockId="41.[189,1398,263,327]" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39">
<materialsCitation id="6DB03C99FFD089209FBCF971FD93F974" collectionCode="AIMB" collectorName="Rajkamal Goswami &amp; Co &amp; Sc" country="India" location="Majuli Island" pageId="38" pageNumber="39" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Assam">
<collectingCountry id="A5CF7654FFD089209FBCF971FEE8F950" box="[189,273,1728,1754]" name="India" pageId="38" pageNumber="39">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFD089209E33F971FE85F950" box="[306,380,1728,1754]" count="2" pageId="38" pageNumber="39" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD089209E8CF971FE2CF950" box="[397,469,1728,1754]" count="2" pageId="38" pageNumber="39" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFD089209EE6F971FDC4F950" box="[487,573,1728,1754]" country="India" name="Assam" pageId="38" pageNumber="39">Assam</collectingRegion>
,
<location id="D807601FFFD089209D4DF971FD01F950" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B:D807601FFFD089209D4DF971FD01F950" box="[588,760,1728,1754]" country="India" name="Majuli Island" pageId="38" pageNumber="39" stateProvince="Assam">Majuli Island</location>
; hand-picked, cow dung;
<collectorName id="702D5312FFD089209B40F971FACCF950" box="[1089,1333,1728,1754]" pageId="38" pageNumber="39">Rajkamal Goswami</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD089209FBCF952FEF5F977" box="[189,268,1763,1789]" pageId="38" pageNumber="39">AIMB</collectionCode>
/
<collectorName id="702D5312FFD089209E17F952FEC0F977" box="[278,313,1763,1789]" pageId="38" pageNumber="39">Co</collectorName>
/
<collectorName id="702D5312FFD089209E43F952FEA6F977" box="[322,351,1763,1789]" pageId="38" pageNumber="39">Sc</collectorName>
1000492 to 1000495
</materialsCitation>
<materialsCitation id="6DB03C99FFD089209D95F955FD97F8C9" collectingDate="2018-09-30" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" elevation="127" latitude="27.96913" location="Rani Village" longLatPrecision="1" longitude="95.32144" municipality="Pasighat" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFD089209D95F955FD54F974" box="[660,685,1764,1790]" count="1" pageId="38" pageNumber="39" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFD089209DC0F952FC4AF977" box="[705,947,1763,1789]" country="India" name="Arunachal Pradesh" pageId="38" pageNumber="39">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD089209CC6F952FBCBF977" box="[967,1074,1763,1789]" pageId="38" pageNumber="39">Pasighat</collectingMunicipality>
, community forest,
<location id="D807601FFFD089209A3CF952FEE0F8AA" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B:D807601FFFD089209A3CF952FEE0F8AA" country="India" latitude="27.96913" longLatPrecision="1" longitude="95.32144" municipality="Pasighat" name="Rani Village" pageId="38" pageNumber="39" stateProvince="Arunachal Pradesh">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD089209E2BF8B4FE05F8AB" box="[298,508,1797,1825]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="8.8644" value="27.96913">27°58ʹ8.8644ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD089209D0CF8B4FD14F8AB" box="[525,749,1797,1825]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="38" pageNumber="39" precision="1" seconds="17.1804" value="95.32144">95°19ʹ17.1804ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD089209C43F8B7FC20F8AA" box="[834,985,1798,1825]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="38" pageNumber="39" unit="m" value="127.0">
<quantity id="1A209B21FFD089209C43F8B7FC6BF8AB" box="[834,914,1798,1825]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="38" pageNumber="39" unit="m" value="127.0">127 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD089209CEAF8B7FB6CF8AA" box="[1003,1173,1798,1824]" pageId="38" pageNumber="39" value="2018-09-30">
<collectingDate id="B922E9ECFFD089209CEAF8B7FB6CF8AA" box="[1003,1173,1798,1824]" pageId="38" pageNumber="39" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD089209BA7F8B6FEB5F8CE" pageId="38" pageNumber="39">cow dung baited pitfall trap</collectingMethod>
;
<collectionCode id="BBC9AE01FFD089209E63F898FE48F8C9" box="[354,433,1833,1859]" pageId="38" pageNumber="39">AIMB</collectionCode>
/
<collectorName id="702D5312FFD089209EBAF898FE27F8C9" box="[443,478,1833,1859]" pageId="38" pageNumber="39">Co</collectorName>
/
<collectorName id="702D5312FFD089209EE6F898FDFDF8C9" box="[487,516,1833,1859]" pageId="38" pageNumber="39">Sc</collectorName>
1000496
</materialsCitation>
<materialsCitation id="6DB03C99FFD089209D9CF89BFEBCF800" collectingDate="2018-09-30" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" elevation="131" latitude="27.969614" location="Rani Village" longLatPrecision="1" longitude="95.32392" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFD089209D9CF89BFD4DF8CE" box="[669,692,1834,1860]" count="1" pageId="38" pageNumber="39" type="female"></specimenCount>
; community forest,
<location id="D807601FFFD089209CCAF898FB88F8C9" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B:D807601FFFD089209CCAF898FB88F8C9" box="[971,1137,1833,1859]" country="India" latitude="27.969614" longLatPrecision="1" longitude="95.32392" name="Rani Village" pageId="38" pageNumber="39">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD089209B88F899FA88F8CE" box="[1161,1393,1832,1860]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="10.6068" value="27.969614">27°58ʹ10.6068ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD089209FBCF8FAFE67F8ED" box="[189,414,1867,1895]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="38" pageNumber="39" precision="1" seconds="26.1156" value="95.32392">95°19ʹ26.1156ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD089209EF9F8FCFD6DF8ED" box="[504,660,1869,1895]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="38" pageNumber="39" unit="m" value="131.0">
<quantity id="1A209B21FFD089209EF9F8FCFDB3F8ED" box="[504,586,1869,1895]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="38" pageNumber="39" unit="m" value="131.0">131 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD089209DA9F8FCFCA1F8ED" box="[680,856,1869,1895]" pageId="38" pageNumber="39" value="2018-09-30">
<collectingDate id="B922E9ECFFD089209DA9F8FCFCA1F8ED" box="[680,856,1869,1895]" pageId="38" pageNumber="39" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
; cow dung;
<collectingMethod id="04994ED3FFD089209CFFF8FCFB1BF8ED" box="[1022,1250,1869,1895]" pageId="38" pageNumber="39">baited pitfall trap</collectingMethod>
;
<collectionCode id="BBC9AE01FFD089209BF2F8FCFAB8F8ED" box="[1267,1345,1869,1895]" pageId="38" pageNumber="39">AIMB</collectionCode>
/
<collectorName id="702D5312FFD089209A4AF8FCFA94F8ED" box="[1355,1389,1869,1895]" pageId="38" pageNumber="39">Co</collectorName>
/
<collectorName id="702D5312FFD089209FBCF8C1FF23F800" box="[189,218,1904,1930]" pageId="38" pageNumber="39">Sc</collectorName>
1000497
</materialsCitation>
<materialsCitation id="6DB03C99FFD089209E63F8C1FE2EF85B" collectingDate="2018-09-29" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="127" latitude="27.96913" location="Rani Village" longLatPrecision="1" longitude="95.32144" pageId="38" pageNumber="39" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFD089209E63F8C1FE83F800" box="[354,378,1904,1930]" count="1" pageId="38" pageNumber="39" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFD089209E88F8C1FE36F800" box="[393,463,1904,1930]" count="2" pageId="38" pageNumber="39" type="female">2 ♀♀</specimenCount>
; community forest,
<location id="D807601FFFD089209DC8F8C1FC9EF800" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B:D807601FFFD089209DC8F8C1FC9EF800" box="[713,871,1904,1930]" country="India" latitude="27.96913" longLatPrecision="1" longitude="95.32144" name="Rani Village" pageId="38" pageNumber="39">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD089209C75F8DFFBBCF800" box="[884,1093,1902,1930]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="8.8644" value="27.96913">27°58ʹ8.8644ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD089209B52F8DFFAC9F800" box="[1107,1328,1902,1930]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="38" pageNumber="39" precision="1" seconds="17.1804" value="95.32144">95°19ʹ17.1804ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD089209FBCF822FEB2F827" box="[189,331,1939,1966]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="38" pageNumber="39" unit="m" value="127.0">
<quantity id="1A209B21FFD089209FBCF822FEF0F824" box="[189,265,1939,1966]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="38" pageNumber="39" unit="m" value="127.0">127 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD089209E5BF822FE05F827" box="[346,508,1939,1965]" pageId="38" pageNumber="39" value="2018-09-29">
<collectingDate id="B922E9ECFFD089209E5BF822FE05F827" box="[346,508,1939,1965]" pageId="38" pageNumber="39" value="2018-09-29">29 Sep. 2018</collectingDate>
</date>
; hand-picked from cow dung,
<collectorName id="702D5312FFD089209C6FF822FBC2F827" box="[878,1083,1939,1965]" pageId="38" pageNumber="39">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFD089209BF2F822FAB8F827" box="[1267,1345,1939,1965]" pageId="38" pageNumber="39">AIMB</collectionCode>
/
<collectorName id="702D5312FFD089209A4AF822FA94F827" box="[1355,1389,1939,1965]" pageId="38" pageNumber="39">Co</collectorName>
/
<collectorName id="702D5312FFD089209FBCF807FF23F85A" box="[189,218,1974,2000]" pageId="38" pageNumber="39">Sc</collectorName>
1000498 to 1000500
</materialsCitation>
<materialsCitation id="6DB03C99FFD0892F9EF0F806FDF2FECD" collectingDate="2019-02-18" collectionCode="AIMB" collectorName="Forest &amp; Harsha Malhotra &amp; Co &amp; Sc" country="India" lastPageId="41" lastPageNumber="42" latitude="23.277756" location="South" longLatPrecision="1" longitude="91.40453" municipality="Trishna Wildlife Sanctuary" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" stateProvince="Tripura">
<specimenCount id="CBDEFD4DFFD089209EF0F806FDF1F85B" box="[497,520,1975,2001]" count="1" pageId="38" pageNumber="39" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFD089209D19F807FD8FF85A" box="[536,630,1974,2000]" country="India" name="Tripura" pageId="38" pageNumber="39">Tripura</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD089209D80F807FC37F85A" box="[641,974,1974,2000]" pageId="38" pageNumber="39">Trishna Wildlife Sanctuary</collectingMunicipality>
,
<location id="D807601FFFD089209CDBF807FBD8F85A" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B:D807601FFFD089209CDBF807FBD8F85A" box="[986,1057,1974,2000]" country="India" latitude="23.277756" longLatPrecision="1" longitude="91.40453" municipality="Trishna Wildlife Sanctuary" name="South" pageId="38" pageNumber="39" stateProvince="Tripura">South</location>
<collectingRegion id="1F1CF826FFD089209B29F807FB7EF85A" box="[1064,1159,1974,2000]" country="India" name="Tripura" pageId="38" pageNumber="39">Tripura</collectingRegion>
;
<geoCoordinate id="B8EC5003FFD089209B92F804FA88F85B" box="[1171,1393,1973,2001]" degrees="23" direction="north" minutes="16" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="39.9216" value="23.277756">23°16ʹ39.9216ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDF892F9FBCFEB6FE65FEA9" box="[189,412,263,291]" degrees="91" direction="east" minutes="24" orientation="longitude" pageId="41" pageNumber="42" precision="1" seconds="16.3152" value="91.40453">91°24ʹ16.3152ʺ E</geoCoordinate>
;
<collectorName id="702D5312FFDF892F9EABFEB8FE03FEA9" box="[426,506,265,291]" pageId="41" pageNumber="42">Forest</collectorName>
,
<date id="A9661004FFDF892F9D08FEB8FD48FEA9" box="[521,689,265,291]" pageId="41" pageNumber="42" value="2019-02-18">
<collectingDate id="B922E9ECFFDF892F9D08FEB8FD48FEA9" box="[521,689,265,291]" pageId="41" pageNumber="42" value="2019-02-18">18 Feb. 2019</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFDF892F9DBEFEB8FBE4FEA9" box="[703,1053,265,291]" pageId="41" pageNumber="42">cow dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFDF892F9B2DFEB8FB05FEA9" box="[1068,1276,265,291]" pageId="41" pageNumber="42">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFDF892F9FFBFE9DFEB2FECD" box="[250,331,300,327]" pageId="41" pageNumber="42">AIMB</collectionCode>
/
<collectorName id="702D5312FFDF892F9E54FE9DFE80FECD" box="[341,377,300,327]" pageId="41" pageNumber="42">Co</collectorName>
/
<collectorName id="702D5312FFDF892F9E83FE9DFE59FECD" box="[386,416,300,327]" pageId="41" pageNumber="42">Sc</collectorName>
1000501
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="89A7664CFFD189219FBCF8F3FB9DF863" pageId="39" pageNumber="40" startId="39.[189,233,1858,1884]" targetBox="[302,1276,327,1774]" targetPageId="39">
<paragraph id="DD6736C4FFD189219FBCF8F3FB9DF863" blockId="39.[189,1406,1858,2026]" pageId="39" pageNumber="40">
<emphasis id="EFACEAD6FFD189219FBCF8F3FEE7F8D6" bold="true" box="[189,286,1858,1884]" pageId="39" pageNumber="40">Fig. 15.</emphasis>
Pronotal basal foveae of male specimens.
<emphasis id="EFACEAD6FFD189219C2CF8F3FCBBF8D6" bold="true" box="[813,834,1858,1884]" pageId="39" pageNumber="40">A</emphasis>
.
<emphasis id="EFACEAD6FFD189219C52F8F3FBB8F8D6" box="[851,1089,1858,1884]" italics="true" pageId="39" pageNumber="40">Onitisbhomorensis</emphasis>
<taxonomicNameLabel id="F49F57ADFFD189219B4BF8F2FB50F8D7" box="[1098,1193,1859,1885]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, NIM/ NBAIR/COL/ONIT/H-271023A.
<emphasis id="EFACEAD6FFD189219D59F8D4FD95F8F5" bold="true" box="[600,620,1893,1919]" pageId="39" pageNumber="40">B</emphasis>
.
<taxonomicName id="1AD84D47FFD189219D86F8D7FBAFF8F5" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[647,1110,1893,1920]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFD189219D86F8D7FC87F8F5" box="[647,894,1894,1920]" italics="true" pageId="39" pageNumber="40">O. punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFD189219C93F8D7FBAFF8F5" author="Janssens A." box="[914,1110,1893,1920]" pageId="39" pageNumber="40" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000492.
<emphasis id="EFACEAD6FFD189219FBCF839FF2BF828" bold="true" box="[189,210,1928,1954]" pageId="39" pageNumber="40">C</emphasis>
.
<taxonomicName id="1AD84D47FFD189219FE7F838FEA3F828" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[230,346,1928,1955]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov.">
<emphasis id="EFACEAD6FFD189219FE7F838FEA3F828" box="[230,346,1928,1955]" italics="true" pageId="39" pageNumber="40">O. kethai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFD189219E67F838FE31F829" box="[358,456,1929,1955]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, NIM/NBAIR/COL/ONIT/H-271023B.
<emphasis id="EFACEAD6FFD189219B2CF838FBBBF829" bold="true" box="[1069,1090,1929,1955]" pageId="39" pageNumber="40">D</emphasis>
.
<taxonomicName id="1AD84D47FFD189219B57F838FF04F84C" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFD189219B57F838FB09F828" box="[1110,1264,1928,1955]" italics="true" pageId="39" pageNumber="40">O. philemon</emphasis>
<bibRefCitation id="B9494B35FFD189219BFCF839FF04F84C" author="Fabricius J. C." pageId="39" pageNumber="40" refId="ref29058" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Volume 2. Impensis Bibliopolii novi academici, Kiel." type="book" year="1801">Fabricius, 1801</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000317.
<emphasis id="EFACEAD6FFD189219D19F81DFDD5F84C" bold="true" box="[536,556,1964,1990]" pageId="39" pageNumber="40">E</emphasis>
.
<taxonomicName id="1AD84D47FFD189219D3BF81DFD3BF84C" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[570,706,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov.">
<emphasis id="EFACEAD6FFD189219D3BF81DFD3BF84C" box="[570,706,1964,1990]" italics="true" pageId="39" pageNumber="40">O. visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFD189219DCBF81DFCDEF84C" box="[714,807,1964,1990]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, NIM/NBAIR/COL/ONIT/H-271023C.
<emphasis id="EFACEAD6FFD189219FBCF87EFF2FF863" bold="true" box="[189,214,1999,2025]" pageId="39" pageNumber="40">F.</emphasis>
<taxonomicName id="1AD84D47FFD189219FDEF87EFD8DF860" authority="Lansberge, 1875" authorityName="Lansberge, AIMB" authorityYear="1875" box="[223,628,1999,2026]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFD189219FDEF87EFE64F863" box="[223,413,1999,2025]" italics="true" pageId="39" pageNumber="40">O. singhalensis</emphasis>
<bibRefCitation id="B9494B35FFD189219EA4F87EFD8DF860" author="Lansberge G." box="[421,628,1999,2026]" pageId="39" pageNumber="40" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000684. (Images: AIMB).
</paragraph>
</caption>
<caption id="89A7664CFFDE892E9FBCF8F3FB1DF860" pageId="40" pageNumber="41" startId="40.[189,229,1858,1884]" targetBox="[303,1269,309,1806]" targetPageId="40">
<paragraph id="DD6736C4FFDE892E9FBCF8F3FB1DF860" blockId="40.[189,1399,1858,2026]" pageId="40" pageNumber="41">
<emphasis id="EFACEAD6FFDE892E9FBCF8F3FED0F8D6" bold="true" box="[189,297,1858,1885]" pageId="40" pageNumber="41">Fig 16. A</emphasis>
. Protibia of
<taxonomicName id="1AD84D47FFDE892E9EB4F8F2FD6AF8D7" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[437,659,1859,1885]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="bhomorensis" status="sp. nov.">
<emphasis id="EFACEAD6FFDE892E9EB4F8F2FD6AF8D7" box="[437,659,1859,1885]" italics="true" pageId="40" pageNumber="41">Onitis bhomorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFDE892E9D98F8F2FD08F8D7" box="[665,753,1859,1885]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A.
<emphasis id="EFACEAD6FFDE892E9E5BF8D7FE94F80A" bold="true" box="[346,365,1894,1920]" pageId="40" pageNumber="41">B</emphasis>
. Anterior tip of protibia of
<taxonomicName id="1AD84D47FFDE892E9DC3F8D7FB9BF80A" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[706,1122,1894,1920]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFDE892E9DC3F8D7FC5DF80A" box="[706,932,1894,1920]" italics="true" pageId="40" pageNumber="41">O. punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFDE892E9CB1F8D7FB9BF80A" author="Janssens A." box="[944,1122,1894,1920]" pageId="40" pageNumber="41" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000492.
<emphasis id="EFACEAD6FFDE892E9FBCF838FF2BF829" bold="true" box="[189,210,1929,1955]" pageId="40" pageNumber="41">C</emphasis>
. Mid-leg of
<taxonomicName id="1AD84D47FFDE892E9E74F83BFE1CF829" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[373,485,1929,1956]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov">
<emphasis id="EFACEAD6FFDE892E9E74F83BFE1CF829" box="[373,485,1929,1956]" italics="true" pageId="40" pageNumber="41">O. kethai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFDE892E9EF3F83BFDB5F82E" box="[498,588,1930,1956]" pageId="40" pageNumber="41" rank="species">sp. nov</taxonomicNameLabel>
, NIM/NBAIR/COL/ONIT/H-271023B.
<emphasis id="EFACEAD6FFDE892E9B2BF838FBC6F829" bold="true" box="[1066,1087,1929,1955]" pageId="40" pageNumber="41">D</emphasis>
. Mid-leg of
<taxonomicName id="1AD84D47FFDE892E9BE3F83BFE8DF84C" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFDE892E9BE3F83BFA8FF829" box="[1250,1398,1929,1956]" italics="true" pageId="40" pageNumber="41">O. philemon</emphasis>
<bibRefCitation id="B9494B35FFDE892E9FBCF81DFE8DF84C" author="Fabricius J. C." box="[189,372,1964,1990]" pageId="40" pageNumber="41" refId="ref29058" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Volume 2. Impensis Bibliopolii novi academici, Kiel." type="book" year="1801">Fabricius, 1801</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000317.
<emphasis id="EFACEAD6FFDE892E9D8FF81DFD58F84C" bold="true" box="[654,673,1964,1990]" pageId="40" pageNumber="41">E</emphasis>
. Mid-leg of
<taxonomicName id="1AD84D47FFDE892E9C39F81CFC45F84C" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[824,956,1964,1991]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov">
<emphasis id="EFACEAD6FFDE892E9C39F81CFC45F84C" box="[824,956,1964,1991]" italics="true" pageId="40" pageNumber="41">O. visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFDE892E9CC4F81CFBE3F84D" box="[965,1050,1965,1991]" pageId="40" pageNumber="41" rank="species">sp. nov</taxonomicNameLabel>
, NIM/NBAIR/COL/ONIT/H- 271023C.
<emphasis id="EFACEAD6FFDE892E9E34F861FEBEF860" bold="true" box="[309,327,2000,2026]" pageId="40" pageNumber="41">F</emphasis>
. Mid-leg of
<taxonomicName id="1AD84D47FFDE892E9ED6F861FCA7F863" authority="Lansberge, AIMB" authorityName="Lansberge, AIMB" authorityYear="1875" box="[471,862,1999,2026]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFDE892E9ED6F861FD71F863" box="[471,648,1999,2026]" italics="true" pageId="40" pageNumber="41">O. singhalensis</emphasis>
Lansberge, AIMB
</taxonomicName>
/Co/Sc1000684. (Images: AIMB).
</paragraph>
</caption>
<subSubSection id="95C2654FFFDF892F9FBCFEDCFE7EFC74" pageId="41" pageNumber="42" type="description">
<paragraph id="DD6736C4FFDF892F9FBCFEDCFEB7FE0D" blockId="41.[189,334,365,391]" box="[189,334,365,391]" pageId="41" pageNumber="42">
<heading id="862F81A8FFDF892F9FBCFEDCFEB7FE0D" bold="true" box="[189,334,365,391]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="EFACEAD6FFDF892F9FBCFEDCFEB7FE0D" bold="true" box="[189,334,365,391]" pageId="41" pageNumber="42">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFE2DFCCBFE3D" blockId="41.[189,818,412,439]" box="[189,818,412,439]" pageId="41" pageNumber="42">MEASUREMENTS (in mm). TL=17.523; BW= 912.5.</paragraph>
<paragraph id="DD6736C4FFDF892F9FEBFE6FFED3FE72" blockId="41.[189,1399,478,785]" box="[234,298,478,504]" pageId="41" pageNumber="42">
<emphasis id="EFACEAD6FFDF892F9FEBFE6FFED3FE72" bold="true" box="[234,298,478,504]" pageId="41" pageNumber="42">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFDB0FC8CFC9B" blockId="41.[189,1399,478,785]" pageId="41" pageNumber="42">Black or reddish black, moderately shining; clypeal margin slightly reflexed; frontal carina broadly interrupted in the middle with an anterior short curved clypeal carina and transverse tubercle behind; pronotum strongly punctate anteriorly, basal foveae rounded, closer, lateral borders of pronotum slightly concave before anterior angles; elytral striae broad, shallow, interval imperceptibly punctured; pygidium flat, opaque; metasternum flat in the middle with an anterior weak longitudinal groove, metasternal shield smooth with scattered punctures, sides of the metasternum punctured with long rusty hairs. Clypeus evenly rugulose, pronotum with a broad smooth longitudinal median line, mid legs toothed, trochanters of mid leg long while that of hind legs serrated.</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFC88FB70FCFC" blockId="41.[189,1399,824,886]" pageId="41" pageNumber="42">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFDF892F9EADFC88FDD7FCD9" box="[428,558,825,851]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="41" pageNumber="42">Fig. 6MP</figureCitation>
). LP=3, Lp= 2.5; BP=1.25, BpB=0.5, BpT =0.4.Parameres longer than the phallobase, symmetrical, straight above, rounded beneath and curved at the tip.
</paragraph>
<paragraph id="DD6736C4FFDF892F9FEBFC2CFEBCFC3D" blockId="41.[189,1398,925,1022]" box="[234,325,925,951]" pageId="41" pageNumber="42">
<emphasis id="EFACEAD6FFDF892F9FEBFC2CFEBCFC3D" bold="true" box="[234,325,925,951]" pageId="41" pageNumber="42">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFC71FE7EFC74" blockId="41.[189,1398,925,1022]" pageId="41" pageNumber="42">Clypeus parabolic, closely rugose, pronotal punctures stronger and closer than in male, protibia with four strong teeth.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDF892F9FBCFB94FB77FB1B" pageId="41" pageNumber="42" type="distribution">
<paragraph id="DD6736C4FFDF892F9FBCFB94FEAFFBB5" blockId="41.[189,342,1061,1087]" box="[189,342,1061,1087]" pageId="41" pageNumber="42">
<heading id="862F81A8FFDF892F9FBCFB94FEAFFBB5" bold="true" box="[189,342,1061,1087]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="EFACEAD6FFDF892F9FBCFB94FEAFFBB5" bold="true" box="[189,342,1061,1087]" pageId="41" pageNumber="42">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFBE5FB77FB1B" blockId="41.[189,1398,1108,1170]" pageId="41" pageNumber="42">
<collectingCountry id="A5CF7654FFDF892F9FBCFBE5FF07FBE4" box="[189,254,1108,1134]" name="India" pageId="41" pageNumber="42">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFDF892F9E0AFBE5FE09FBE4" box="[267,496,1108,1134]" country="India" name="Arunachal Pradesh" pageId="41" pageNumber="42">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFDF892F9EFCFBE5FDA9FBE4" box="[509,592,1108,1134]" country="India" name="Assam" pageId="41" pageNumber="42">Assam</collectingRegion>
,
<collectingRegion id="1F1CF826FFDF892F9D5EFBE5FD40FBE4" box="[607,697,1108,1134]" country="India" name="Tripura" pageId="41" pageNumber="42">Tripura</collectingRegion>
,
<collectingRegion id="1F1CF826FFDF892F9DC6FBE5FC9AFBE4" box="[711,867,1108,1134]" country="India" name="West Bengal" pageId="41" pageNumber="42">West Bengal</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFDF892F9B3FFBE5FB58FBE4" box="[1086,1185,1108,1134]" name="Pakistan" pageId="41" pageNumber="42">Pakistan</collectingCountry>
(
<bibRefCitation id="B9494B35FFDF892F9BB5FBE5FF02FB1B" author="Bezdek A. &amp; Krell F. - T." pageId="41" pageNumber="42" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDF892F9E06FBC6FE0BFB1B" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[263,498,1143,1169]" pageId="41" pageNumber="42" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFDF892F9E73FBC9FE54FB1B" box="[370,429,1143,1169]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDF892F9EFFFBC6FD34FB18" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[510,717,1143,1170]" pageId="41" pageNumber="42" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFDF892F9D4CFBC9FD71FB1B" box="[589,648,1143,1169]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDF892F9DD8FBC6FC8EFB1B" author="Bezdek A." box="[729,887,1143,1169]" pageId="41" pageNumber="42" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDF892F9C82FBC6FB7BFB1B" author="Schoolmeesters P." box="[899,1154,1143,1169]" pageId="41" pageNumber="42" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,706 @@
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<subSubSection id="95C2654FFFDA892A9D65FDD4FC29FDF5" box="[612,976,612,639]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="DD6736C4FFDA892A9D65FDD4FC29FDF5" blockId="44.[612,976,612,674]" box="[612,976,612,639]" pageId="44" pageNumber="45">
<heading id="862F81A8FFDA892A9D65FDD4FC29FDF5" box="[612,976,612,639]" centered="true" fontSize="11" level="2" pageId="44" pageNumber="45" reason="2">
<taxonomicName id="1AD84D47FFDA892A9D65FDD4FC29FDF5" ID-CoL="49KLJ" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[612,976,612,639]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFDA892A9D65FDD4FCCDFDF5" bold="true" box="[612,820,612,639]" italics="true" pageId="44" pageNumber="45">Onitis subopacus</emphasis>
<bibRefCitation id="B9494B35FFDA892A9C3AFDD4FC29FDF5" author="Arrow G. J." box="[827,976,613,639]" pageId="44" pageNumber="45" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDA892A9D9DFD39FC9CFD6E" pageId="44" pageNumber="45" type="description">
<paragraph id="DD6736C4FFDA892A9D9DFD39FC6EFD28" blockId="44.[612,976,612,674]" box="[668,919,648,674]" pageId="44" pageNumber="45">
<figureCitation id="45E32A41FFDA892A9D9DFD39FCF0FD28" box="[668,777,648,674]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="44" pageNumber="45">Figs 12C</figureCitation>
,
<figureCitation id="45E32A41FFDA892A9C14FD39FC97FD28" box="[789,878,648,674]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. EH. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." pageId="44" pageNumber="45">13AD</figureCitation>
,
<figureCitation id="45E32A41FFDA892A9C7BFD39FC6EFD28" box="[890,919,648,674]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." pageId="44" pageNumber="45">20</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FBCFD7BFC9CFD6E" blockId="44.[189,869,713,740]" box="[189,869,713,740]" pageId="44" pageNumber="45">
<emphasis id="EFACEAD6FFDA892A9FBCFD7BFE71FD6E" box="[189,392,714,740]" italics="true" pageId="44" pageNumber="45">Onitissubopacus</emphasis>
<bibRefCitation id="B9494B35FFDA892A9E8FFD7BFDA4FD6E" author="Arrow G. J." box="[398,605,713,740]" pageId="44" pageNumber="45" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931: 395</bibRefCitation>
(original description).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDA892A9FBCFCBAFD0DFB41" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="DD6736C4FFDA892A9FBCFCBAFDF0FB0F" blockId="44.[189,1406,779,1228]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="FDC811EAFFDA892A9FBCFCBAFDF0FB0F" pageId="44" pageNumber="45">
<taxonomicName id="1AD84D47FFDA892A9FBCFCBAFE75FCAF" ID-CoL="49KLJ" authorityName="Arrow" authorityYear="1931" box="[189,396,779,805]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFDA892A9FBCFCBAFE75FCAF" box="[189,396,779,805]" italics="true" pageId="44" pageNumber="45">Onitis subopacus</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFDA892A9EB0FCBAFD6EFCAF" author="Balthasar V." box="[433,663,779,806]" page="94" pageId="44" pageNumber="45" year="1935">
<bibRefCitation id="B9494B35FFDA892A9EB0FCBAFD6EFCAF" author="Balthasar V." box="[433,663,779,806]" pageId="44" pageNumber="45" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 94</bibRefCitation>
</treatmentCitation>
(monograph); 1963: 53 (monograph). —
<treatmentCitation id="5C7910D5FFDA892A9B9AFCBDFA8FFCAC" author="Janssens A." box="[1179,1398,779,806]" page="51" pageId="44" pageNumber="45" year="1937">
<bibRefCitation id="B9494B35FFDA892A9B9AFCBDFA8FFCAC" author="Janssens A." box="[1179,1398,779,806]" pageId="44" pageNumber="45" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 51</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFDA892A9E95FC9FFCF3FCC2" author="Biswas S. &amp; Chatterjee S. K." box="[404,778,814,840]" page="67" pageId="44" pageNumber="45" year="1986">
<bibRefCitation id="B9494B35FFDA892A9E95FC9FFCF3FCC2" author="Biswas S. &amp; Chatterjee S. K." box="[404,778,814,840]" pageId="44" pageNumber="45" pagination="57 - 67" refId="ref27582" refString="Biswas S. &amp; Chatterjee S. K. 1986 a. Dung beetle fauna (Coleoptera: Scarabaeidae: Scarabaeinae) of Palamou Tiger Reserve, Bihar, with description of a new species. Records of the Zoological Survey of India 83 (3 - 4): 57 - 67. https: // doi. org / 10.26515 / rzsi / v 83 / i 3 - 4 / 1986 / 161311" type="journal article" year="1986">Biswas &amp; Chatterjee 1986a: 67</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDA892A9CE4FC9FFB33FCC2" author="Chandra K." box="[997,1226,814,840]" page="360" pageId="44" pageNumber="45" year="2000">
<bibRefCitation id="B9494B35FFDA892A9CE4FC9FFB33FCC2" author="Chandra K." box="[997,1226,814,840]" pageId="44" pageNumber="45" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000: 360</bibRefCitation>
</treatmentCitation>
(distribution); 2005: 150 (distribution). —
<treatmentCitation id="5C7910D5FFDA892A9D39FCE0FD09FCE1" author="Mittal I. C." box="[568,752,849,876]" page="46" pageId="44" pageNumber="45" year="2005">
<bibRefCitation id="B9494B35FFDA892A9D39FCE0FD09FCE1" author="Mittal I. C." box="[568,752,849,876]" pageId="44" pageNumber="45" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005: 46</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFDA892A9C65FCE0FB35FCE6" author="Chandra K. &amp; Ahirwar S. C." box="[868,1228,849,876]" page="149" pageId="44" pageNumber="45" year="2005">
<bibRefCitation id="B9494B35FFDA892A9C65FCE0FB35FCE6" author="Chandra K. &amp; Ahirwar S. C." box="[868,1228,849,876]" pageId="44" pageNumber="45" pagination="147 - 155" refId="ref28418" refString="Chandra K. &amp; Ahirwar S. C. 2005. Scarabaeid Beetles (Coleoptera) of Kanha Tiger Reserve, Madhya Pradesh. Records Zoological Survey of India 105 (1 - 2): 147 - 155." type="journal article" year="2005">Chandra &amp; Ahirwar 2005: 149</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDA892A9E0FFCC4FDBAFC05" author="Bezdek A. &amp; Krell F. - T." box="[270,579,884,911]" page="159" pageId="44" pageNumber="45" year="2006">
<bibRefCitation id="B9494B35FFDA892A9E0FFCC4FDBAFC05" author="Bezdek A. &amp; Krell F. - T." box="[270,579,884,911]" pageId="44" pageNumber="45" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDA892A9C17FCC4FC08FC05" author="Sabu T. K. &amp; Vinod K. V. &amp; Vineesh P. J." box="[790,1009,884,911]" page="5" pageId="44" pageNumber="45" year="2006">
<bibRefCitation id="B9494B35FFDA892A9C17FCC4FC08FC05" author="Sabu T. K. &amp; Vinod K. V. &amp; Vineesh P. J." box="[790,1009,884,911]" pageId="44" pageNumber="45" pagination="1 - 17" refId="ref30840" refString="Sabu T. K., Vinod K. V. &amp; Vineesh P. J. 2006. Guild structure, diversity and succession of dung beetles associated with Indian elephant dung in South Western Ghats forests. Journal of Insect Science 6 (1): 1 - 17. https: // doi. org / 10.1673 / 2006 _ 06 _ 17.1" type="journal article" year="2006">
Sabu
<emphasis id="EFACEAD6FFDA892A9C59FCC4FC6BFC05" box="[856,914,885,911]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2006: 5
</bibRefCitation>
</treatmentCitation>
(list); 2011: 34 (list). —
<treatmentCitation id="5C7910D5FFDA892A9A12FCC4FE1EFC38" author="Chandra K. &amp; Ahirwar S. C." page="276" pageId="44" pageNumber="45" year="2007">
<bibRefCitation id="B9494B35FFDA892A9A12FCC4FE1EFC38" author="Chandra K. &amp; Ahirwar S. C." pageId="44" pageNumber="45" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">Chandra &amp; Ahirwar 2007: 276</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDA892A9DBFFC29FC73FC38" author="Sewak R." box="[702,906,920,946]" page="60" pageId="44" pageNumber="45" year="2009">
<bibRefCitation id="B9494B35FFDA892A9DBFFC29FC73FC38" author="Sewak R." box="[702,906,920,946]" pageId="44" pageNumber="45" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">Sewak 2009b: 60</bibRefCitation>
</treatmentCitation>
(key, diagnosis, distribution). —
<treatmentCitation id="5C7910D5FFDA892A9A17FC29FE5DFC5C" author="Thakare V. &amp; Zade V. &amp; Chandra K. &amp; Gupta D." page="75" pageId="44" pageNumber="45" year="2012">
<bibRefCitation id="B9494B35FFDA892A9A17FC29FE5DFC5C" author="Thakare V. &amp; Zade V. &amp; Chandra K. &amp; Gupta D." pageId="44" pageNumber="45" pagination="73 - 80" refId="ref31359" refString="Thakare V., Zade V., Chandra K. &amp; Gupta D. 2012. Scarab beetles (Coleoptera: Scarabaeoidea) of Melghat Tiger Reserve, Central India. Academic Journal of Entomology 5 (2): 73 - 80." type="journal article" year="2012">
Thakare
<emphasis id="EFACEAD6FFDA892A9FEBFC0DFED2FC5F" box="[234,299,955,981]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2012: 75
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDA892A9D8BFC0AFC14FC5F" author="Chandra K. &amp; Gupta D." box="[650,1005,955,981]" page="274" pageId="44" pageNumber="45" year="2011">
<bibRefCitation id="B9494B35FFDA892A9D8BFC0AFC14FC5F" author="Chandra K. &amp; Gupta D." box="[650,1005,955,981]" pageId="44" pageNumber="45" pagination="272 - 278" refId="ref28502" refString="Chandra K. &amp; Gupta D. 2011. Study of scarabaeid beetles (Coleoptera) of Veerangana Durgavati Wildlife Sanctuary, Damoh, Madhya Pradesh, India. Deccan Current Science 5: 272 - 278." type="journal article" year="2011">Chandra &amp; Gupta 2011: 274</bibRefCitation>
</treatmentCitation>
(distribution); 2012a: 889 (list) Chandra
<emphasis id="EFACEAD6FFDA892A9E55FC6EFE77FC72" box="[340,398,990,1016]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2012a: 52 (list); 2012b: 33 (distribution). —
<treatmentCitation id="5C7910D5FFDA892A9C9BFC6FFB09FC72" author="Chandra K. &amp; Gupta D." box="[922,1264,990,1016]" page="349" pageId="44" pageNumber="45" year="2013">
<bibRefCitation id="B9494B35FFDA892A9C9BFC6FFB09FC72" author="Chandra K. &amp; Gupta D." box="[922,1264,990,1016]" pageId="44" pageNumber="45" pagination="331 - 360" refId="ref28634" refString="Chandra K. &amp; Gupta D. 2013 a. Taxonomic studies on dung beetles (Coleoptera: Scarabaeidae, Geotrupidae, Hybosoridae) of Chhattisgarh, India. Munis Entomology and Zoology 8 (1): 331 - 360." type="journal article" year="2013">Chandra &amp; Gupta 2013a: 349</bibRefCitation>
</treatmentCitation>
(diagnosis, distribution); 2013b: 4667 (distribution). — Chandra
<emphasis id="EFACEAD6FFDA892A9C67FBB3FC5BFB91" box="[870,930,1025,1051]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2011: 67 (distribution). —
<treatmentCitation id="5C7910D5FFDA892A9BEAFBB3FE98FBB5" author="Gupta D. &amp; Chandra K. &amp; Khan S." page="230" pageId="44" pageNumber="45" year="2014">
<bibRefCitation id="B9494B35FFDA892A9BEAFBB3FE98FBB5" author="Gupta D. &amp; Chandra K. &amp; Khan S." pageId="44" pageNumber="45" pagination="225 - 240" refId="ref29198" refString="Gupta D., Chandra K. &amp; Khan S. 2014. An updated checklist of Scarabaeoid beetles (Coleoptera: Scarabaeoidea) of Pench Tiger Reserve, Madhya Pradesh, India. Journal of Entomology and Zoology Studies 2 (5): 225 - 240. Available from https: // www. entomoljournal. com / vol 2 Issue 5 / 57.1. html [accessed 12 Jun. 2024]" type="journal article" year="2014">
Gupta
<emphasis id="EFACEAD6FFDA892A9A3AFBB3FA8EFB91" box="[1339,1399,1025,1051]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2014: 230
</bibRefCitation>
</treatmentCitation>
(distribution); 2015: 1037 (list, distribution). —
<treatmentCitation id="5C7910D5FFDA892A9CA7FB94FB35FBB5" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[934,1228,1061,1087]" page="300" pageId="44" pageNumber="45" year="2014">
<bibRefCitation id="B9494B35FFDA892A9CA7FB94FB35FBB5" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[934,1228,1061,1087]" pageId="44" pageNumber="45" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFDA892A9B10FB94FBB4FBB5" box="[1041,1101,1061,1087]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2014: 300
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDA892A9E13FBF9FE14FBE8" author="Bezdek A." box="[274,493,1096,1122]" page="179" pageId="44" pageNumber="45" year="2016">
<bibRefCitation id="B9494B35FFDA892A9E13FBF9FE14FBE8" author="Bezdek A." box="[274,493,1096,1122]" pageId="44" pageNumber="45" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDA892A9DCBFBF9FC18FBE8" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." box="[714,993,1096,1122]" page="239" pageId="44" pageNumber="45" year="2020">
<bibRefCitation id="B9494B35FFDA892A9DCBFBF9FC18FBE8" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." box="[714,993,1096,1122]" pageId="44" pageNumber="45" pagination="221 - 275" refId="ref29141" refString="Ghosh J., Das, P., Ghosh S. K., Bhunia D., Kushwaha R. K., Gupta D., &amp; Chandra K. 2020. Insecta: Coleoptera. Zoological Survey of India, Fauna of Haryana, State Fauna Series 24: 221 - 275." type="book chapter" year="2020">
Ghosh
<emphasis id="EFACEAD6FFDA892A9C20FBF8FCA7FBE8" box="[801,862,1096,1122]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2020: 239
</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFDA892A9BBFFBF9FEDCFB0F" author="Schoolmeesters P." pageId="44" pageNumber="45" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FBCFB3FFD0DFB41" blockId="44.[189,1406,779,1228]" pageId="44" pageNumber="45">
<emphasis id="EFACEAD6FFDA892A9FBCFB3FFE85FB22" box="[189,380,1166,1192]" italics="true" pageId="44" pageNumber="45">Onitisphilemon</emphasis>
Lansberge (nec Fabricius) 1875: 133. —
<bibRefCitation id="B9494B35FFDA892A9C94FB3EFB5BFB22" author="Boucomont A." box="[917,1186,1166,1193]" pageId="44" pageNumber="45" pagination="238 - 350" refId="ref27893" refString="Boucomont A. 1914. Les Coprophages de I'Archipel Malais. Annales de la Societe entomologique de France. LXXXIII: 238 - 350." type="book chapter" year="1914">Boucomont 1914: 336</bibRefCitation>
(distribution). —
<bibRefCitation id="B9494B35FFDA892A9FEBFB03FDB3FB46" author="Boucomont A. &amp; Gillet J." box="[234,586,1201,1228]" pageId="44" pageNumber="45" pagination="1 - 76" refId="ref27918" refString="Boucomont A. &amp; Gillet J. 1921. Faune entomologique de l'Indochine francaise. Famille Scarabaeidae Laparosticti (Coleopteres). Portail. Saigon 4: 1 - 76." type="journal article" year="1921">Boucomont &amp; Gillet 1921: 19</bibRefCitation>
(distribution).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDA892A9FBCFB42FAFFFAD5" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="DD6736C4FFDA892A9FBCFB42FE3FFA87" blockId="44.[189,454,1267,1293]" box="[189,454,1267,1293]" pageId="44" pageNumber="45">
<heading id="862F81A8FFDA892A9FBCFB42FE3FFA87" bold="true" box="[189,454,1267,1293]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="EFACEAD6FFDA892A9FBCFB42FE3FFA87" bold="true" box="[189,454,1267,1293]" pageId="44" pageNumber="45">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FBCFA90FAFFFAD5" blockId="44.[189,1399,1313,1375]" pageId="44" pageNumber="45">
<emphasis id="EFACEAD6FFDA892A9FBCFA90FE71FAB1" box="[189,392,1313,1339]" italics="true" pageId="44" pageNumber="45">Onitissubopacus</emphasis>
belongs to Group III and closely resembles
<taxonomicName id="1AD84D47FFDA892A9C93FA93FC04FAB1" authorityName="Lansberge" authorityYear="1875" box="[914,1021,1314,1340]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFDA892A9C93FA93FC04FAB1" box="[914,1021,1314,1340]" italics="true" pageId="44" pageNumber="45">O. virens</emphasis>
</taxonomicName>
of the same group but differs in that the former has a less strongly punctured pronotum and broadly interrupted frontal carina.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDA892B9FBCFA37FA8EFD90" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="DD6736C4FFDA892A9FBCFA37FE15FA2A" blockId="44.[189,492,1414,1440]" box="[189,492,1414,1440]" pageId="44" pageNumber="45">
<heading id="862F81A8FFDA892A9FBCFA37FE15FA2A" bold="true" box="[189,492,1414,1440]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="EFACEAD6FFDA892A9FBCFA37FE15FA2A" bold="true" box="[189,492,1414,1440]" pageId="44" pageNumber="45">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FEBFA04FEA5FA45" blockId="44.[189,1398,1461,1558]" box="[234,348,1461,1487]" pageId="44" pageNumber="45">
<emphasis id="EFACEAD6FFDA892A9FEBFA04FEA5FA45" bold="true" box="[234,348,1461,1487]" pageId="44" pageNumber="45">
<typeStatus id="02638866FFDA892A9FEBFA04FEA5FA45" box="[234,348,1461,1487]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FBCFA69FC2EF99F" blockId="44.[189,1398,1461,1558]" pageId="44" pageNumber="45">
<materialsCitation id="6DB03C99FFDA892A9FBCFA69FC2EF99F" collectionCode="BMNH" country="Sri Lanka" location="SRI LANKA" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="A5CF7654FFDA892A9FBCFA69FEACFA78" box="[189,341,1496,1522]" name="Sri Lanka" pageId="44" pageNumber="45">SRI LANKA</collectingCountry>
<specimenCount id="CBDEFD4DFFDA892A9E6BFA69FE7AFA78" box="[362,387,1496,1522]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
; “Kalupahani, Haldumulle Ceylon, 1904171,
<taxonomicName id="1AD84D47FFDA892A9CA2FA69FB95FA78" authorityName="Arrow" authorityYear="1931" box="[931,1132,1496,1522]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFDA892A9CA2FA69FB95FA78" box="[931,1132,1496,1522]" italics="true" pageId="44" pageNumber="45">Onitis subopacus</emphasis>
</taxonomicName>
<typeStatus id="02638866FFDA892A9B70FA69FB5FFA78" box="[1137,1190,1496,1522]" pageId="44" pageNumber="45">type</typeStatus>
, Arrow (aedeagus point-mounted and attached with specimen)”;
<collectionCode id="BBC9AE01FFDA892A9DDDFA4DFCCDF99C" box="[732,820,1532,1558]" pageId="44" pageNumber="45">BMNH</collectionCode>
(E) 1237174.
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FEBF98DFCE2F9DC" blockId="44.[189,1400,1596,2010]" box="[234,795,1596,1623]" pageId="44" pageNumber="45">
<emphasis id="EFACEAD6FFDA892A9FEBF98DFD9BF9DC" bold="true" box="[234,610,1596,1622]" pageId="44" pageNumber="45">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFDA892A9D70F98CFCEAF9DC" box="[625,787,1596,1623]" count="39" pageId="44" pageNumber="45" type="generic">39 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFDA892B9FBCF9D1FA8EFD90" blockId="44.[189,1400,1596,2010]" lastBlockId="45.[189,1400,263,538]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<materialsCitation id="6DB03C99FFDA892A9FBCF9D1FC4AF914" collectingDate="1928-05-18" collectionCode="BMNH" collectorName="C. F. C. Beeson" country="India" county="Kashmir" location="Sindh Valley" municipality="Haran Pltn." pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1">
<collectingCountry id="A5CF7654FFDA892A9FBCF9D1FEF2F9F0" box="[189,267,1632,1658]" name="India" pageId="44" pageNumber="45">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFDA892A9E25F9D1FEC3F9F0" box="[292,314,1632,1658]" count="1" pageId="44" pageNumber="45" type="female"></specimenCount>
; “
<collectingCounty id="34064E48FFDA892A9E52F9EEFE40F9F3" box="[339,441,1631,1657]" pageId="44" pageNumber="45">Kashmir</collectingCounty>
;
<collectingMunicipality id="3D03ACBEFFDA892A9EC5F9D1FDBCF9F3" box="[452,581,1631,1658]" pageId="44" pageNumber="45">Haran Pltn.</collectingMunicipality>
5500,
<location id="D807601FFFDA892A9D93F9EEFCDBF9F3" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9D93F9EEFCDBF9F3" box="[658,802,1631,1657]" country="India" county="Kashmir" municipality="Haran Pltn." name="Sindh Valley" pageId="44" pageNumber="45">Sindh Valley</location>
,
<date id="A9661004FFDA892A9C2CF9D1FC64F9F0" box="[813,925,1632,1658]" pageId="44" pageNumber="45" value="1928-05-18">
<collectingDate id="B922E9ECFFDA892A9C2CF9D1FC64F9F0" box="[813,925,1632,1658]" pageId="44" pageNumber="45" value="1928-05-18">18.v.1928</collectingDate>
</date>
,
<collectorName id="702D5312FFDA892A9CA8F9D1FBB4F9F0" box="[937,1101,1632,1658]" pageId="44" pageNumber="45">C.F.C. Beeson</collectorName>
; ex. coll., Dehradun, B.M. 1928-472;
<emphasis id="EFACEAD6FFDA892A9E38F932FE05F917" box="[313,508,1667,1693]" italics="true" pageId="44" pageNumber="45">
<location id="D807601FFFDA892A9E38F932FE05F917" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9E38F932FE05F917" box="[313,508,1667,1693]" country="India" county="Kashmir" municipality="Haran Pltn." name="Onitissubopacus" pageId="44" pageNumber="45">Onitissubopacus</location>
</emphasis>
co-type,
<location id="D807601FFFDA892A9D69F932FD4EF917" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9D69F932FD4EF917" box="[616,695,1667,1693]" country="India" county="Kashmir" municipality="Haran Pltn." name="Arrow" pageId="44" pageNumber="45">Arrow</location>
”;
<collectionCode id="BBC9AE01FFDA892A9DCEF932FCDEF917" box="[719,807,1667,1693]" pageId="44" pageNumber="45">BMNH</collectionCode>
(E) 1237175
</materialsCitation>
<materialsCitation id="6DB03C99FFDA892A9CD1F932FE65F94A" collectionCode="ZSIM" country="India" determinerName="Det. G. J. Arrow" location="Arrow Collection" municipality="Roorkee" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFDA892A9CD1F932FC11F917" box="[976,1000,1667,1693]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
; “
<collectingMunicipality id="3D03ACBEFFDA892A9B01F932FB9DF917" box="[1024,1124,1667,1693]" pageId="44" pageNumber="45">Roorkee</collectingMunicipality>
,
<location id="D807601FFFDA892A9B71F932FAC0F917" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9B71F932FAC0F917" box="[1136,1337,1667,1693]" country="India" municipality="Roorkee" name="Arrow Collection" pageId="44" pageNumber="45">Arrow Collection</location>
,
<determinerName id="28BBDA00FFDA892A9A46F932FEC3F94A" pageId="44" pageNumber="45">Det. G.J. Arrow</determinerName>
”;
<collectionCode id="BBC9AE01FFDA892A9E59F917FE65F94A" box="[344,412,1702,1728]" pageId="44" pageNumber="45">ZSIM</collectionCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFDA892A9EB3F916FDE0F94A" box="[434,537,1702,1729]" collectionCode="ZSIM" country="India" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFDA892A9EB3F916FE31F94B" box="[434,456,1703,1729]" count="1" pageId="44" pageNumber="45" type="female"></specimenCount>
;
<collectionCode id="BBC9AE01FFDA892A9ED4F917FDE0F94A" box="[469,537,1702,1728]" pageId="44" pageNumber="45">ZSIM</collectionCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFDA892A9D2EF917FCBFF88C" collectingDate="2009-05-08" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" elevation="839" latitude="8.861611" location="Rockwood" longLatPrecision="1" longitude="77.13347" municipality="Shendurney Wildlife Sanctuary" pageId="44" pageNumber="45" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDA892A9D2EF917FD88F94B" box="[559,625,1702,1729]" count="4" pageId="44" pageNumber="45" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDA892A9D7AF917FD45F94B" box="[635,700,1702,1729]" count="2" pageId="44" pageNumber="45" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFDA892A9DC9F917FCECF94A" box="[712,789,1702,1728]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDA892A9C1EF917FB85F94A" box="[799,1148,1702,1728]" pageId="44" pageNumber="45">Shendurney Wildlife Sanctuary</collectingMunicipality>
,
<location id="D807601FFFDA892A9B87F917FAF9F94A" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9B87F917FAF9F94A" box="[1158,1280,1702,1728]" country="India" latitude="8.861611" longLatPrecision="1" longitude="77.13347" municipality="Shendurney Wildlife Sanctuary" name="Rockwood" pageId="44" pageNumber="45" stateProvince="Kerala">Rockwood</location>
grassland;
<geoCoordinate id="B8EC5003FFDA892A9FBCF979FE99F96E" box="[189,352,1736,1764]" degrees="08" direction="north" minutes="51" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="41.8" value="8.861611">08°51ʹ41.8ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDA892A9E6DF979FDF5F96E" box="[364,524,1736,1764]" degrees="77" direction="east" minutes="08" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="00.5" value="77.13347">77°08ʹ00.5ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFDA892A9D54F978FD25F969" box="[597,732,1737,1764]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="44" pageNumber="45" unit="m" value="839.0">
<quantity id="1A209B21FFDA892A9D54F978FD66F96E" box="[597,671,1737,1764]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="44" pageNumber="45" unit="m" value="839.0">839 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFDA892A9DE8F978FC81F969" box="[745,888,1737,1764]" pageId="44" pageNumber="45" value="2009-05-08">
<collectingDate id="B922E9ECFFDA892A9DE8F978FC81F969" box="[745,888,1737,1764]" pageId="44" pageNumber="45" value="2009-05-08">8 May 2009</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFDA892A9C85F97BFB5CF96E" box="[900,1189,1737,1764]" pageId="44" pageNumber="45">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFDA892A9BB3F978FE9FF88C" pageId="44" pageNumber="45">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDA892A9EB7F95CFE06F88D" box="[438,511,1773,1799]" pageId="44" pageNumber="45">AIMB</collectionCode>
/
<collectorName id="702D5312FFDA892A9D09F95CFDD1F88D" box="[520,552,1773,1799]" pageId="44" pageNumber="45">Co</collectorName>
/
<collectorName id="702D5312FFDA892A9D30F95CFDB5F88D" box="[561,588,1773,1799]" pageId="44" pageNumber="45">Sc</collectorName>
1000711 to 1000716
</materialsCitation>
<materialsCitation id="6DB03C99FFDA892A9C70F95CFA8EF8A0" collectingDate="2009-12-04" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" elevation="173" latitude="8.912666" location="Kattilappara" longLatPrecision="1" longitude="77.09592" municipality="Shendurney Wildlife Sanctuary" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDA892A9C70F95CFC7EF88D" box="[881,903,1773,1799]" count="1" pageId="44" pageNumber="45" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFDA892A9C9EF95DFC15F88C" box="[927,1004,1772,1798]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDA892A9B01F95DFA8AF88D" box="[1024,1395,1772,1799]" pageId="44" pageNumber="45">Shendurney Wildlife Sanctuary</collectingMunicipality>
,
<location id="D807601FFFDA892A9FBCF8BEFEBFF8A3" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9FBCF8BEFEBFF8A3" box="[189,326,1807,1833]" country="India" latitude="8.912666" longLatPrecision="1" longitude="77.09592" municipality="Shendurney Wildlife Sanctuary" name="Kattilappara" pageId="44" pageNumber="45" stateProvince="Kerala">Kattilappara</location>
evergreen forest;
<geoCoordinate id="B8EC5003FFDA892A9D0DF8BFFD55F8A0" box="[524,684,1806,1834]" degrees="08" direction="north" minutes="54" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="45.6" value="8.912666">08°54ʹ45.6ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDA892A9DB7F8BFFCADF8A0" box="[694,852,1806,1834]" degrees="77" direction="east" minutes="05" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="45.3" value="77.09592">77°05ʹ45.3ʺ E</geoCoordinate>
; elev.
<quantity id="1A209B21FFDA892A9C99F8BEFC24F8A3" box="[920,989,1807,1834]" metricMagnitude="2" metricUnit="m" metricValue="1.725168" pageId="44" pageNumber="45" unit="ft" value="566.0">
<elevation id="56F5D1F7FFDA892A9C99F8BEFC24F8A3" box="[920,989,1807,1834]" metricMagnitude="2" metricUnit="m" metricValue="1.725168" pageId="44" pageNumber="45" unit="ft" value="566.0">566 ft</elevation>
</quantity>
,
<date id="A9661004FFDA892A9CECF8A1FB8EF8A0" box="[1005,1143,1808,1834]" pageId="44" pageNumber="45" value="2009-12-04">
<collectingDate id="B922E9ECFFDA892A9CECF8A1FB8EF8A0" box="[1005,1143,1808,1834]" pageId="44" pageNumber="45" value="2009-12-04">4 Dec. 2009</collectingDate>
</date>
;
<collectionCode id="BBC9AE01FFDA892A9B7EF8A1FB31F8A0" box="[1151,1224,1808,1834]" pageId="44" pageNumber="45">AIMB</collectionCode>
/
<collectorName id="702D5312FFDA892A9BD0F8A1FB08F8A0" box="[1233,1265,1808,1834]" pageId="44" pageNumber="45">Co</collectorName>
/
<collectorName id="702D5312FFDA892A9BFBF8A1FAECF8A0" box="[1274,1301,1808,1834]" pageId="44" pageNumber="45">Sc</collectorName>
1000717
</materialsCitation>
<materialsCitation id="6DB03C99FFDA892A9FCFF882FD8CF819" collectingDate="2009-12-03" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" elevation="185" latitude="8.9151945" location="Kattilappara" longLatPrecision="1" longitude="77.10797" municipality="Shendurney Wildlife Sanctuary" pageId="44" pageNumber="45" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDA892A9FCFF882FEE8F8C7" box="[206,273,1843,1869]" count="2" pageId="44" pageNumber="45" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDA892A9E1CF882FEA6F8C7" box="[285,351,1843,1870]" count="5" pageId="44" pageNumber="45" type="female">5 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFDA892A9E6CF882FE43F8C7" box="[365,442,1843,1869]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDA892A9EC7F882FCD1F8C7" box="[454,808,1843,1869]" pageId="44" pageNumber="45">Shendurney Wildlife Sanctuary</collectingMunicipality>
;
<location id="D807601FFFDA892A9C32F882FC45F8C7" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9C32F882FC45F8C7" box="[819,956,1843,1869]" country="India" latitude="8.9151945" longLatPrecision="1" longitude="77.10797" municipality="Shendurney Wildlife Sanctuary" name="Kattilappara" pageId="44" pageNumber="45" stateProvince="Kerala">Kattilappara</location>
moist deciduous forest;
<geoCoordinate id="B8EC5003FFDA892A9BD1F880FA8AF8C7" box="[1232,1395,1841,1869]" degrees="08" direction="north" minutes="54" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="54.7" value="8.9151945">08°54ʹ54.7ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDA892A9FBCF8E5FEA4F8FB" box="[189,349,1876,1905]" degrees="77" direction="east" minutes="06" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="28.7" value="77.10797">77°06ʹ28.7ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFDA892A9EA4F8E7FDD2F8FA" box="[421,555,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="44" pageNumber="45" unit="m" value="185.0">
<quantity id="1A209B21FFDA892A9EA4F8E7FE16F8FB" box="[421,495,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="44" pageNumber="45" unit="m" value="185.0">185 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFDA892A9D38F8E7FD31F8FA" box="[569,712,1878,1905]" pageId="44" pageNumber="45" value="2009-12-03">
<collectingDate id="B922E9ECFFDA892A9D38F8E7FD31F8FA" box="[569,712,1878,1905]" pageId="44" pageNumber="45" value="2009-12-03">3 Dec. 2009</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFDA892A9DD2F8E6FC0DF8FB" box="[723,1012,1878,1905]" pageId="44" pageNumber="45">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFDA892A9B00F8E7FA8EF8FA" box="[1025,1399,1878,1905]" pageId="44" pageNumber="45">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDA892A9FF5F8C8FEC4F819" box="[244,317,1913,1939]" pageId="44" pageNumber="45">AIMB</collectionCode>
/
<collectorName id="702D5312FFDA892A9E47F8C8FE9FF819" box="[326,358,1913,1939]" pageId="44" pageNumber="45">Co</collectorName>
/
<collectorName id="702D5312FFDA892A9E6EF8C8FE73F819" box="[367,394,1913,1939]" pageId="44" pageNumber="45">Sc</collectorName>
1000718 to 1000724
</materialsCitation>
<materialsCitation id="6DB03C99FFDA892A9D91F8C8FC0AF850" collectingDate="2009-12-03" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" elevation="207" latitude="8.918222" location="Kattilapara" longLatPrecision="1" longitude="77.09575" municipality="Shendurney Wildlife Sanctuary" pageId="44" pageNumber="45" specimenCount="15" specimenCount-female="6" specimenCount-male="9" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDA892A9D91F8C8FD2DF81E" box="[656,724,1913,1940]" count="9" pageId="44" pageNumber="45" type="male">9 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDA892A9DE0F8C8FCDDF81E" box="[737,804,1913,1940]" count="6" pageId="44" pageNumber="45" type="female">6 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFDA892A9C32F8C8FC79F819" box="[819,896,1913,1939]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDA892A9C8CF8C8FB17F819" box="[909,1262,1913,1939]" pageId="44" pageNumber="45">Shendurney Wildlife Sanctuary</collectingMunicipality>
,
<location id="D807601FFFDA892A9BFAF8C8FA8EF819" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9BFAF8C8FA8EF819" box="[1275,1399,1913,1939]" country="India" latitude="8.918222" longLatPrecision="1" longitude="77.09575" municipality="Shendurney Wildlife Sanctuary" name="Kattilapara" pageId="44" pageNumber="45" stateProvince="Kerala">Kattilapara</location>
degraded forest;
<geoCoordinate id="B8EC5003FFDA892A9E7DF82AFDD9F83D" box="[380,544,1947,1975]" degrees="08" direction="north" minutes="55" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="05.6" value="8.918222">08°55ʹ05.6ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDA892A9D2CF82AFD37F83D" box="[557,718,1947,1975]" degrees="77" direction="east" minutes="05" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="44.7" value="77.09575">77°05ʹ44.7ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFDA892A9C18F82CFC58F83C" box="[793,929,1948,1975]" metricMagnitude="2" metricUnit="m" metricValue="2.07" pageId="44" pageNumber="45" unit="m" value="207.0">
<quantity id="1A209B21FFDA892A9C18F82CFC9AF83D" box="[793,867,1949,1975]" metricMagnitude="2" metricUnit="m" metricValue="2.07" pageId="44" pageNumber="45" unit="m" value="207.0">207 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFDA892A9CAEF82CFBB9F83D" box="[943,1088,1949,1975]" pageId="44" pageNumber="45" value="2009-12-03">
<collectingDate id="B922E9ECFFDA892A9CAEF82CFBB9F83D" box="[943,1088,1949,1975]" pageId="44" pageNumber="45" value="2009-12-03">3 Dec. 2009</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFDA892A9B4CF82CFA8AF83D" box="[1101,1395,1948,1975]" pageId="44" pageNumber="45">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFDA892A9FBCF871FDCAF853" box="[189,563,1983,2010]" pageId="44" pageNumber="45">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDA892A9D72F80EFD45F850" box="[627,700,1983,2010]" pageId="44" pageNumber="45">AIMB</collectionCode>
/
<collectorName id="702D5312FFDA892A9DC4F80EFD1CF850" box="[709,741,1983,2010]" pageId="44" pageNumber="45">Co</collectorName>
/
<collectorName id="702D5312FFDA892A9DEFF80EFCF0F850" box="[750,777,1983,2010]" pageId="44" pageNumber="45">Sc</collectorName>
1000725 to 1000738
</materialsCitation>
<materialsCitation id="6DB03C99FFDA892B9B0FF871FB4DFECC" collectingDate="2009-11-29" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" elevation="1234" lastPageId="45" lastPageNumber="46" latitude="8.827528" location="Pandimotta" longLatPrecision="1" longitude="77.21719" municipality="Shendurney Wildlife Sanctuary" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDA892A9B0FF871FBDFF850" box="[1038,1062,1984,2010]" count="1" pageId="44" pageNumber="45" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFDA892A9B35F80EFB78F853" box="[1076,1153,1983,2009]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDA892B9B8CF80EFED4FEA9" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">Shendurney Wildlife Sanctuary</collectingMunicipality>
,
<location id="D807601FFFDB892B9E38FEB8FE41FEA9" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDB892B9E38FEB8FE41FEA9" box="[313,440,265,291]" country="India" latitude="8.827528" longLatPrecision="1" longitude="77.21719" municipality="Shendurney Wildlife Sanctuary" name="Pandimotta" pageId="45" pageNumber="46" stateProvince="Kerala">Pandimotta</location>
montane evergreen;
<geoCoordinate id="B8EC5003FFDB892B9DA2FEB6FCBCFEA9" box="[675,837,263,291]" degrees="08" direction="north" minutes="49" orientation="latitude" pageId="45" pageNumber="46" precision="1" seconds="39.1" value="8.827528">08°49ʹ39.1ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDB892B9C51FEB6FC16FEA9" box="[848,1007,263,291]" degrees="77" direction="east" minutes="13" orientation="longitude" pageId="45" pageNumber="46" precision="1" seconds="01.9" value="77.21719">77°13ʹ01.9ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFDB892B9B36FEB8FB30FEA9" box="[1079,1225,265,291]" metricMagnitude="3" metricUnit="m" metricValue="1.234" pageId="45" pageNumber="46" unit="m" value="1234.0">
<quantity id="1A209B21FFDB892B9B36FEB8FB74FEA9" box="[1079,1165,265,291]" metricMagnitude="3" metricUnit="m" metricValue="1.234" pageId="45" pageNumber="46" unit="m" value="1234.0">1234 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFDB892B9BD7FEB8FA8AFEA9" box="[1238,1395,265,291]" pageId="45" pageNumber="46" value="2009-11-29">
<collectingDate id="B922E9ECFFDB892B9BD7FEB8FA8AFEA9" box="[1238,1395,265,291]" pageId="45" pageNumber="46" value="2009-11-29">29 Nov. 2009</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFDB892B9FBCFE9CFE12FECD" box="[189,491,300,327]" pageId="45" pageNumber="46">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFDB892B9EFAFE9DFC8EFECC" box="[507,887,300,327]" pageId="45" pageNumber="46">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDB892B9CBCFE9DFBFFFECC" box="[957,1030,300,326]" pageId="45" pageNumber="46">AIMB</collectionCode>
/
<collectorName id="702D5312FFDB892B9B0EFE9DFBD6FECC" box="[1039,1071,300,326]" pageId="45" pageNumber="46">Co</collectorName>
/
<collectorName id="702D5312FFDB892B9B36FE9DFBABFECC" box="[1079,1106,300,326]" pageId="45" pageNumber="46">Sc</collectorName>
1000739
</materialsCitation>
<materialsCitation id="6DB03C99FFDB892B9BD4FE9CFC48FE07" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="11.974199" location="Sungathakatte" longLatPrecision="1" longitude="76.1856" municipality="Nagarhole National Park" pageId="45" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFDB892B9BD4FE9CFB14FECD" box="[1237,1261,301,327]" count="1" pageId="45" pageNumber="46" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFDB892B9BFCFE9DFA8AFECC" box="[1277,1395,300,326]" country="India" name="Karnataka" pageId="45" pageNumber="46">Karnataka</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDB892B9FBCFEFEFE2BFEE3" box="[189,466,335,361]" pageId="45" pageNumber="46">Nagarhole National Park</collectingMunicipality>
,
<location id="D807601FFFDB892B9EDDFEFEFD83FEE3" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDB892B9EDDFEFEFD83FEE3" box="[476,634,335,361]" country="India" latitude="11.974199" longLatPrecision="1" longitude="76.1856" municipality="Nagarhole National Park" name="Sungathakatte" pageId="45" pageNumber="46" stateProvince="Karnataka">Sungathakatte</location>
dry deciduous forest;
<geoCoordinate id="B8EC5003FFDB892B9C6CFEFFFBE2FEE0" box="[877,1051,334,362]" degrees="11" direction="north" minutes="58" orientation="latitude" pageId="45" pageNumber="46" precision="1" seconds="27.12" value="11.974199">11°58ʹ27.12″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDB892B9B24FEFFFB3AFEE0" box="[1061,1219,334,362]" degrees="76" direction="east" minutes="11" orientation="longitude" pageId="45" pageNumber="46" precision="1" seconds="8.16" value="76.1856">76°11ʹ8.16″ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFDB892B9BCCFEE1FEDEFE07" pageId="45" pageNumber="46">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFDB892B9E32FEC3FD82FE06" box="[307,635,370,396]" pageId="45" pageNumber="46">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDB892B9DBBFEC2FCFAFE07" box="[698,771,371,397]" pageId="45" pageNumber="46">AIMB</collectionCode>
/
<collectorName id="702D5312FFDB892B9C0AFEC2FCD2FE07" box="[779,811,371,397]" pageId="45" pageNumber="46">Co</collectorName>
/
<collectorName id="702D5312FFDB892B9C35FEC2FCB6FE07" box="[820,847,371,397]" pageId="45" pageNumber="46">Sc</collectorName>
1000740
</materialsCitation>
<materialsCitation id="6DB03C99FFDB892B9CCBFEC2FEC2FE59" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="12.0634" location="Sarathi Nulla" longLatPrecision="1" longitude="76.17799" pageId="45" pageNumber="46" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFDB892B9CCBFEC2FC19FE07" box="[970,992,371,397]" count="1" pageId="45" pageNumber="46" type="female"></specimenCount>
;
<location id="D807601FFFDB892B9CEFFEC3FB7AFE06" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDB892B9CEFFEC3FB7AFE06" box="[1006,1155,370,396]" country="India" latitude="12.0634" longLatPrecision="1" longitude="76.17799" name="Sarathi Nulla" pageId="45" pageNumber="46">Sarathi Nulla</location>
dry deciduous forest;
<geoCoordinate id="B8EC5003FFDB892B9FBCFE25FE9BFE3A" box="[189,354,404,432]" degrees="12" direction="north" minutes="3" orientation="latitude" pageId="45" pageNumber="46" precision="1" seconds="48.24" value="12.0634">12°3ʹ48.24″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDB892B9E71FE25FDEBFE3A" box="[368,530,404,432]" degrees="76" direction="east" minutes="10" orientation="longitude" pageId="45" pageNumber="46" precision="1" seconds="40.8" value="76.17799">76°10ʹ40.8″ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFDB892B9D20FE27FC9AFE3A" box="[545,867,406,432]" pageId="45" pageNumber="46">open cattle dung baited trap</collectingMethod>
;
<collectorName id="702D5312FFDB892B9C71FE27FB42FE3A" box="[880,1211,406,432]" pageId="45" pageNumber="46">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDB892B9BFCFE27FABFFE3A" box="[1277,1350,406,432]" pageId="45" pageNumber="46">AIMB</collectionCode>
/
<collectorName id="702D5312FFDB892B9A4FFE27FA97FE3A" box="[1358,1390,406,432]" pageId="45" pageNumber="46">Co</collectorName>
/
<collectorName id="702D5312FFDB892B9FBCFE08FF21FE59" box="[189,216,441,467]" pageId="45" pageNumber="46">Sc</collectorName>
1000741
</materialsCitation>
<materialsCitation id="6DB03C99FFDB892B9E57FE08FD5FFE7C" collectingDate="2019-02-18" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" location="South" municipality="Eden of Bison" pageId="45" pageNumber="46" specimenCount="1" specimenCount-female="1" stateProvince="Tripura">
<specimenCount id="CBDEFD4DFFDB892B9E57FE08FE95FE59" box="[342,364,441,467]" count="1" pageId="45" pageNumber="46" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFDB892B9E7AFE08FE29FE59" box="[379,464,441,467]" country="India" name="Tripura" pageId="45" pageNumber="46">Tripura</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDB892B9EDDFE08FD7BFE59" box="[476,642,441,467]" pageId="45" pageNumber="46">Eden of Bison</collectingMunicipality>
,
<location id="D807601FFFDB892B9D8EFE08FD29FE59" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDB892B9D8EFE08FD29FE59" box="[655,720,441,467]" country="India" municipality="Eden of Bison" name="South" pageId="45" pageNumber="46" stateProvince="Tripura">South</location>
<collectingRegion id="1F1CF826FFDB892B9DD9FE08FCD4FE59" box="[728,813,441,467]" country="India" name="Tripura" pageId="45" pageNumber="46">Tripura</collectingRegion>
, forest;
<date id="A9661004FFDB892B9C89FE08FBDBFE59" box="[904,1058,441,467]" pageId="45" pageNumber="46" value="2019-02-18">
<collectingDate id="B922E9ECFFDB892B9C89FE08FBDBFE59" box="[904,1058,441,467]" pageId="45" pageNumber="46" value="2019-02-18">18 Feb. 2019</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFDB892B9FBCFE6DFE8EFE7C" box="[189,375,476,502]" pageId="45" pageNumber="46">Harsha Malhotra</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDB892B9EAEFE6DFE01FE7C" box="[431,504,476,502]" pageId="45" pageNumber="46">AIMB</collectionCode>
/
<collectorName id="702D5312FFDB892B9D00FE6DFDD8FE7C" box="[513,545,476,502]" pageId="45" pageNumber="46">Co</collectorName>
/
<collectorName id="702D5312FFDB892B9D28FE6DFDBDFE7C" box="[553,580,476,502]" pageId="45" pageNumber="46">Sc</collectorName>
1000742
</materialsCitation>
<materialsCitation id="6DB03C99FFDB892B9DB8FE6DFA8AFD90" collectingDate="2021-04-24" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" location="Forest" municipality="Singphan Elephant Reserve" pageId="45" pageNumber="46" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Nagaland">
<specimenCount id="CBDEFD4DFFDB892B9DB8FE6DFD00FE7D" box="[697,761,476,503]" count="2" pageId="45" pageNumber="46" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDB892B9C03FE6CFCE1FE7D" box="[770,792,477,503]" count="1" pageId="45" pageNumber="46" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFDB892B9C25FE6DFC6BFE7C" box="[804,914,476,502]" country="India" name="Nagaland" pageId="45" pageNumber="46">Nagaland</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDB892B9C9BFE6DFB35FE7D" box="[922,1228,476,503]" pageId="45" pageNumber="46">Singphan Elephant Reserve</collectingMunicipality>
,
<location id="D807601FFFDB892B9BD4FE6CFAE7FE7D" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDB892B9BD4FE6CFAE7FE7D" box="[1237,1310,477,503]" country="India" municipality="Singphan Elephant Reserve" name="Forest" pageId="45" pageNumber="46" stateProvince="Nagaland">Forest</location>
;
<date id="A9661004FFDB892B9A27FE6DFF01FD93" pageId="45" pageNumber="46" value="2021-04-24">
<collectingDate id="B922E9ECFFDB892B9A27FE6DFF01FD93" pageId="45" pageNumber="46" value="2021-04-24">24Apr. 2021</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFDB892B9D48FE4EFC43FD93" box="[585,954,511,538]" pageId="45" pageNumber="46">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDB892B9CF4FE4EFBC7FD93" box="[1013,1086,511,537]" pageId="45" pageNumber="46">AIMB</collectionCode>
/
<collectorName id="702D5312FFDB892B9B46FE4EFB9EFD93" box="[1095,1127,511,537]" pageId="45" pageNumber="46">Co</collectorName>
/
<collectorName id="702D5312FFDB892B9B71FE4EFB72FD93" box="[1136,1163,511,537]" pageId="45" pageNumber="46">Sc</collectorName>
1000743 to 1000745
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDB892B9FBCFDF1FCE8FB00" pageId="45" pageNumber="46" type="description">
<paragraph id="DD6736C4FFDB892B9FBCFDF1FEB7FDD0" blockId="45.[189,334,576,602]" box="[189,334,576,602]" pageId="45" pageNumber="46">
<heading id="862F81A8FFDB892B9FBCFDF1FEB7FDD0" bold="true" box="[189,334,576,602]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="EFACEAD6FFDB892B9FBCFDF1FEB7FDD0" bold="true" box="[189,334,576,602]" pageId="45" pageNumber="46">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCFDDEFCFCFD03" blockId="45.[189,773,623,650]" box="[189,773,623,650]" pageId="45" pageNumber="46">MEASUREMENTS (in mm). TL=1623, BW= 812.</paragraph>
<paragraph id="DD6736C4FFDB892B9FEBFD00FED3FD41" blockId="45.[189,1399,689,962]" box="[234,298,689,715]" pageId="45" pageNumber="46">
<emphasis id="EFACEAD6FFDB892B9FEBFD00FED3FD41" bold="true" box="[234,298,689,715]" pageId="45" pageNumber="46">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCFD65FBFDFC4B" blockId="45.[189,1399,689,962]" pageId="45" pageNumber="46">
Black with a slight metallic lustre; clypeus elliptical, rugulose, vertex granular, more shining; frontal carina widely interrupted in the middle with a conical tubercle behind and a curved clypeal carina in front; pronotum not strongly but rather closely punctured, with an incomplete smooth median longitudinal line, basal foveae deep; pygidium opaque scarcely punctured; elytra finely striate with sutural, 3
<superScript id="2AAD9B8CFFDB892B9E2BFCD1FEC1FCE4" attach="left" box="[298,312,864,878]" fontSize="6" pageId="45" pageNumber="46">rd</superScript>
and 5
<superScript id="2AAD9B8CFFDB892B9E7EFCD1FE75FCE4" attach="left" box="[383,396,864,878]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
intervals slightly convex; metasternal shield smooth and shining, the front angles and sides of metasternum closely clothed with reddish hairs. Protibia toothed, mid femur lobed near middle with tooth on posterior edge, trochanter of the hind leg sharply toothed.
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCFC58FA98FBAC" blockId="45.[189,1399,1000,1062]" pageId="45" pageNumber="46">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFDB892B9EB5FC58FDB1FB89" box="[436,584,1001,1027]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. EH. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." pageId="45" pageNumber="46">Fig. 13AD</figureCitation>
). LP=3, Lp=2.5; BP=1.25, BpB= 0.5, BpT =0.4. Phallobase longer than parameres. Parameres curved outwards at the base and then curved inwards with the tip pointed.
</paragraph>
<paragraph id="DD6736C4FFDB892B9FEBFBFCFEBCFBED" blockId="45.[189,785,1101,1163]" box="[234,325,1101,1127]" pageId="45" pageNumber="46">
<emphasis id="EFACEAD6FFDB892B9FEBFBFCFEBCFBED" bold="true" box="[234,325,1101,1127]" pageId="45" pageNumber="46">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCFBC1FCE8FB00" blockId="45.[189,785,1101,1163]" box="[189,785,1136,1163]" pageId="45" pageNumber="46">Protibia toothed, middle and hind legs not toothed.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDB892B9FBCFB03FC1FFA7B" pageId="45" pageNumber="46" type="distribution">
<paragraph id="DD6736C4FFDB892B9FBCFB03FEAFFB46" blockId="45.[189,342,1202,1228]" box="[189,342,1202,1228]" pageId="45" pageNumber="46">
<heading id="862F81A8FFDB892B9FBCFB03FEAFFB46" bold="true" box="[189,342,1202,1228]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="EFACEAD6FFDB892B9FBCFB03FEAFFB46" bold="true" box="[189,342,1202,1228]" pageId="45" pageNumber="46">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCFB51FC1FFA7B" blockId="45.[189,1399,1248,1522]" pageId="45" pageNumber="46">
<collectingCountry id="A5CF7654FFDB892B9FBCFB51FF07FB70" box="[189,254,1248,1274]" name="India" pageId="45" pageNumber="46">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFDB892B9E0BFB51FE16FB70" box="[266,495,1248,1274]" country="India" name="Arunachal Pradesh" pageId="45" pageNumber="46">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9EFDFB50FDB6FB71" box="[508,591,1249,1275]" country="India" name="Assam" pageId="45" pageNumber="46">Assam</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9D5FFB51FD58FB70" box="[606,673,1248,1274]" country="India" name="Bihar" pageId="45" pageNumber="46">Bihar</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9DB1FB51FCB0FB70" box="[688,841,1248,1274]" country="India" name="Chhattisgarh" pageId="45" pageNumber="46">Chhattisgarh</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9C56FB51FC4BFB70" box="[855,946,1248,1274]" country="India" name="Gujarat" pageId="45" pageNumber="46">Gujarat</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9CC0FB50FBDEFB71" box="[961,1063,1249,1275]" country="India" name="Haryana" pageId="45" pageNumber="46">Haryana</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9B37FB51FAEAFB70" box="[1078,1299,1248,1274]" country="India" name="Himachal Pradesh" pageId="45" pageNumber="46">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9A23FB50FEA2FA94" country="India" name="Jammu and Kashmir" pageId="45" pageNumber="46">Jammu and Kashmir</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9E6AFAB5FE1EFA94" box="[363,487,1284,1310]" country="India" name="Jharkhand" pageId="45" pageNumber="46">Jharkhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9EF6FAB5FD8AFA94" box="[503,627,1284,1310]" country="India" name="Karnataka" pageId="45" pageNumber="46">Karnataka</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9D85FAB5FD2CFA94" box="[644,725,1284,1310]" country="India" name="Kerala" pageId="45" pageNumber="46">Kerala</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9DE7FAB5FC4BFA94" box="[742,946,1284,1310]" country="India" name="Madhya Pradesh" pageId="45" pageNumber="46">Madhya Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9CC2FAB5FBA0FA94" box="[963,1113,1284,1310]" country="India" name="Maharashtra" pageId="45" pageNumber="46">Maharashtra</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9B68FAB5FB17FA94" box="[1129,1262,1284,1310]" country="India" name="Meghalaya" pageId="45" pageNumber="46">Meghalaya</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9BFEFAB5FA8BFA94" box="[1279,1394,1284,1310]" country="India" name="Nagaland" pageId="45" pageNumber="46">Nagaland</collectingRegion>
, New
<collectingRegion id="1F1CF826FFDB892B9FFFFA96FEBBFACB" box="[254,322,1319,1345]" country="India" name="Delhi" pageId="45" pageNumber="46">Delhi</collectingRegion>
, Punjab,
<collectingRegion id="1F1CF826FFDB892B9EB0FA96FDD1FACB" box="[433,552,1319,1345]" country="India" name="Rajasthan" pageId="45" pageNumber="46">Rajasthan</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9D34FA96FD3CFACB" box="[565,709,1319,1345]" country="India" name="Tamil Nadu" pageId="45" pageNumber="46">Tamil Nadu</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9DD3FA96FCD5FACB" box="[722,812,1319,1345]" country="India" name="Tripura" pageId="45" pageNumber="46">Tripura</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9C38FA96FC34FACB" box="[825,973,1319,1345]" country="India" name="Uttarakhand" pageId="45" pageNumber="46">Uttarakhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9CDAFA96FB79FACB" box="[987,1152,1319,1345]" country="India" name="Uttar Pradesh" pageId="45" pageNumber="46">Uttar Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFDB892B9B8CFA96FADEFACB" box="[1165,1319,1319,1345]" country="India" name="West Bengal" pageId="45" pageNumber="46">West Bengal</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFDB892B9E39FAFBFE33FAEE" box="[312,458,1354,1380]" name="Afghanistan" pageId="45" pageNumber="46">Afghanistan</collectingCountry>
,
<collectingCountry id="A5CF7654FFDB892B9ED9FAFBFD9DFAEE" box="[472,612,1354,1380]" name="Bangladesh" pageId="45" pageNumber="46">Bangladesh</collectingCountry>
,
<collectingCountry id="A5CF7654FFDB892B9D70FAFBFD14FAEE" box="[625,749,1354,1380]" name="Cambodia" pageId="45" pageNumber="46">Cambodia</collectingCountry>
,
<collectingCountry id="A5CF7654FFDB892B9DF8FAFBFCBBFAEE" box="[761,834,1354,1380]" name="China" pageId="45" pageNumber="46">China</collectingCountry>
,
<collectingCountry id="A5CF7654FFDB892B9C51FAFBFC3CFAEE" box="[848,965,1354,1380]" name="Indonesia" pageId="45" pageNumber="46">Indonesia</collectingCountry>
, Malay Peninsula,
<collectingCountry id="A5CF7654FFDB892B9BA9FAFAFAE5FAEF" box="[1192,1308,1355,1381]" name="Myanmar" pageId="45" pageNumber="46">Myanmar</collectingCountry>
,
<collectingCountry id="A5CF7654FFDB892B9A28FAFBFA8BFAEE" box="[1321,1394,1354,1380]" name="Nepal" pageId="45" pageNumber="46">Nepal</collectingCountry>
,
<collectingCountry id="A5CF7654FFDB892B9FBCFADCFEDAFA0D" box="[189,291,1389,1415]" name="Pakistan" pageId="45" pageNumber="46">Pakistan</collectingCountry>
,
<collectingCountry id="A5CF7654FFDB892B9E2EFADCFE5CFA0D" box="[303,421,1389,1415]" name="Sri Lanka" pageId="45" pageNumber="46">Sri Lanka</collectingCountry>
,
<collectingCountry id="A5CF7654FFDB892B9EB0FADCFDE5FA0D" box="[433,540,1389,1415]" name="Thailand" pageId="45" pageNumber="46">Thailand</collectingCountry>
,
<collectingCountry id="A5CF7654FFDB892B9D29FADCFD76FA0D" box="[552,655,1389,1415]" name="Vietnam" pageId="45" pageNumber="46">Vietnam</collectingCountry>
. (
<bibRefCitation id="B9494B35FFDB892B9DA3FADCFC10FA0D" author="Biswas S. &amp; Chatterjee S. K." box="[674,1001,1389,1415]" pageId="45" pageNumber="46" pagination="57 - 67" refId="ref27582" refString="Biswas S. &amp; Chatterjee S. K. 1986 a. Dung beetle fauna (Coleoptera: Scarabaeidae: Scarabaeinae) of Palamou Tiger Reserve, Bihar, with description of a new species. Records of the Zoological Survey of India 83 (3 - 4): 57 - 67. https: // doi. org / 10.26515 / rzsi / v 83 / i 3 - 4 / 1986 / 161311" type="journal article" year="1986">Biswas &amp; Chatterjee 1986a</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDB892B9CF7FADCFB66FA0D" author="Chandra K." box="[1014,1183,1389,1415]" pageId="45" pageNumber="46" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000</bibRefCitation>
, 2005;
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,
<bibRefCitation id="B9494B35FFDB892B9E71FA20FE57FA21" author="Chandra K. &amp; Ahirwar S. C." box="[368,430,1425,1451]" pageId="45" pageNumber="46" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">2007</bibRefCitation>
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<bibRefCitation id="B9494B35FFDB892B9EBBFA21FD40FA20" author="Bezdek A. &amp; Krell F. - T." box="[442,697,1424,1451]" pageId="45" pageNumber="46" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
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).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,835 @@
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<taxonomicName id="1AD84D47FFDB892B9D69F987FC35F9DB" ID-CoL="49KMB" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[616,972,1590,1617]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFDB892B9D69F987FCF8F9DA" bold="true" box="[616,769,1590,1616]" italics="true" pageId="45" pageNumber="46">Onitis virens</emphasis>
<bibRefCitation id="B9494B35FFDB892B9C08F987FC35F9DB" author="Lansberge G." box="[777,972,1590,1617]" pageId="45" pageNumber="46" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDB892B9D9CF9EBFD64F952" pageId="45" pageNumber="46" type="description">
<paragraph id="DD6736C4FFDB892B9D9CF9EBFC6EF9FE" blockId="45.[616,972,1590,1652]" box="[669,919,1626,1652]" pageId="45" pageNumber="46">
<figureCitation id="45E32A41FFDB892B9D9CF9EBFCF2F9FE" box="[669,779,1626,1652]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="45" pageNumber="46">Figs 12D</figureCitation>
,
<figureCitation id="45E32A41FFDB892B9C19F9EBFC97F9FE" box="[792,878,1626,1652]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. EH. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." pageId="45" pageNumber="46">13EH</figureCitation>
,
<figureCitation id="45E32A41FFDB892B9C7BF9EBFC6EF9FE" box="[890,919,1626,1652]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." pageId="45" pageNumber="46">20</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCF92DFC99F93F" blockId="45.[189,864,1691,1753]" box="[189,864,1691,1718]" pageId="45" pageNumber="46">
<emphasis id="EFACEAD6FFDB892B9FBCF92DFEAAF93C" box="[189,339,1692,1718]" italics="true" pageId="45" pageNumber="46">Onitisvirens</emphasis>
<bibRefCitation id="B9494B35FFDB892B9E5BF92AFDA0F93C" author="Lansberge G." box="[346,601,1691,1718]" pageId="45" pageNumber="46" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875: 135</bibRefCitation>
(original description).
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCF90FFD64F952" blockId="45.[189,864,1691,1753]" box="[189,669,1726,1753]" pageId="45" pageNumber="46">
<emphasis id="EFACEAD6FFDB892B9FBCF90FFE69F952" box="[189,400,1726,1752]" italics="true" pageId="45" pageNumber="46">Onitisamplectens</emphasis>
<bibRefCitation id="B9494B35FFDB892B9E96F90FFD60F952" author="Lansberge G." box="[407,665,1726,1753]" pageId="45" pageNumber="46" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875: 136</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDB89289FBCF8B1FC33FEE3" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="reference_group">
<paragraph id="DD6736C4FFDB89289FBCF8B1FC33FEE3" blockId="45.[189,1399,1792,1995]" lastBlockId="46.[234,1399,265,362]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<treatmentCitationGroup id="FDC811EAFFDB89289FBCF8B1FC33FEE3" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<taxonomicName id="1AD84D47FFDB892B9FBCF8B1FEAFF890" ID-CoL="49KMB" authorityName="Lansberge" authorityYear="1875" box="[189,342,1792,1818]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFDB892B9FBCF8B1FEAFF890" box="[189,342,1792,1818]" italics="true" pageId="45" pageNumber="46">Onitis virens</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFDB892B9E7BF8B1FD1FF890" author="Boucomont A. &amp; Gillet J." box="[378,742,1792,1818]" page="19" pageId="45" pageNumber="46" year="1921">
<bibRefCitation id="B9494B35FFDB892B9E7BF8B1FD1FF890" author="Boucomont A. &amp; Gillet J." box="[378,742,1792,1818]" pageId="45" pageNumber="46" pagination="1 - 76" refId="ref27918" refString="Boucomont A. &amp; Gillet J. 1921. Faune entomologique de l'Indochine francaise. Famille Scarabaeidae Laparosticti (Coleopteres). Portail. Saigon 4: 1 - 76." type="journal article" year="1921">Boucomont &amp; Gillet 1921: 19</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDB892B9CC5F8B1FB6DF890" author="Arrow G. J." box="[964,1172,1792,1818]" page="396" pageId="45" pageNumber="46" year="1931">
<bibRefCitation id="B9494B35FFDB892B9CC5F8B1FB6DF890" author="Arrow G. J." box="[964,1172,1792,1818]" pageId="45" pageNumber="46" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 396</bibRefCitation>
</treatmentCitation>
(key, description). —
<treatmentCitation id="5C7910D5FFDB892B9E14F892FE06F8B4" author="Balthasar V." box="[277,511,1827,1854]" page="52" pageId="45" pageNumber="46" year="1935">
<bibRefCitation id="B9494B35FFDB892B9E14F892FE06F8B4" author="Balthasar V." box="[277,511,1827,1854]" pageId="45" pageNumber="46" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 52</bibRefCitation>
</treatmentCitation>
(monograph); 1963: 40 (monograph). —
<treatmentCitation id="5C7910D5FFDB892B9B0FF892FB09F8B7" author="Paulian R." box="[1038,1264,1827,1854]" page="144" pageId="45" pageNumber="46" year="1945">
<bibRefCitation id="B9494B35FFDB892B9B0FF892FB09F8B7" author="Paulian R." box="[1038,1264,1827,1854]" pageId="45" pageNumber="46" refId="ref30535" refString="Paulian R. 1945. Faune de l'empire francais: Coleopteres Scarabeides de l'Indochine, Vol. III: 1 - 227. Librairie Larose, Paris." type="book" year="1945">Paulian 1945: 144</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFDB892B9E13F8F7FD74F8EA" author="Biswas S. &amp; Chatterjee S. K." box="[274,653,1862,1888]" page="260" pageId="45" pageNumber="46" year="1991">
<bibRefCitation id="B9494B35FFDB892B9E13F8F7FD74F8EA" author="Biswas S. &amp; Chatterjee S. K." box="[274,653,1862,1888]" pageId="45" pageNumber="46" pagination="243 - 262" refId="ref27741" refString="Biswas S. &amp; Chatterjee S. K. 1991. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, State Fauna Series 1: Fauna of Orissa (Part 3): 243 - 262. Available from: https: // shorturl. at / rZQQv [accessed 15 Oct. 2018]" type="book chapter" year="1991">Biswas &amp; Chatterjee 1991: 260</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDB892B9C6AF8F7FBA8F8EA" author="Chandra K." box="[875,1105,1862,1888]" page="360" pageId="45" pageNumber="46" year="2000">
<bibRefCitation id="B9494B35FFDB892B9C6AF8F7FBA8F8EA" author="Chandra K." box="[875,1105,1862,1888]" pageId="45" pageNumber="46" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000: 360</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDB892B9A2EF8F7FEA8F809" author="Mittal I. C." page="46" pageId="45" pageNumber="46" year="2005">
<bibRefCitation id="B9494B35FFDB892B9A2EF8F7FEA8F809" author="Mittal I. C." pageId="45" pageNumber="46" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005: 46</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFDB892B9EC0F8D8FCDCF80E" author="Chandra K. &amp; Ahirwar S. C." box="[449,805,1897,1924]" page="149" pageId="45" pageNumber="46" year="2005">
<bibRefCitation id="B9494B35FFDB892B9EC0F8D8FCDCF80E" author="Chandra K. &amp; Ahirwar S. C." box="[449,805,1897,1924]" pageId="45" pageNumber="46" pagination="147 - 155" refId="ref28418" refString="Chandra K. &amp; Ahirwar S. C. 2005. Scarabaeid Beetles (Coleoptera) of Kanha Tiger Reserve, Madhya Pradesh. Records Zoological Survey of India 105 (1 - 2): 147 - 155." type="journal article" year="2005">Chandra &amp; Ahirwar 2005: 149</bibRefCitation>
</treatmentCitation>
(distribution); 2007: 276 (distribution). —
<treatmentCitation id="5C7910D5FFDB892B9A1EF8D8FE38F82D" author="Bezdek A. &amp; Krell F. - T." page="159" pageId="45" pageNumber="46" year="2006">
<bibRefCitation id="B9494B35FFDB892B9A1EF8D8FE38F82D" author="Bezdek A. &amp; Krell F. - T." pageId="45" pageNumber="46" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDB892B9D95F83CFCA4F82C" author="Sewak R." box="[660,861,1932,1959]" page="60" pageId="45" pageNumber="46" year="2009">
<bibRefCitation id="B9494B35FFDB892B9D95F83CFCA4F82C" author="Sewak R." box="[660,861,1932,1959]" pageId="45" pageNumber="46" pagination="1 - 48" refId="ref31194" refString="Sewak R. 2009 a. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar desert of Gujarat. Records of the Zoological Survey of India, Occassional Paper No 295: 1 - 48." type="journal article" year="2009">Sewak 2009a: 60</bibRefCitation>
</treatmentCitation>
(key, diagnosis, distribution), 2009b: 60 (key, diagnosis, distribution). —
<treatmentCitation id="5C7910D5FFDB892B9D32F801FCDEF840" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." box="[563,807,1968,1994]" page="34" pageId="45" pageNumber="46" year="2011">
<bibRefCitation id="B9494B35FFDB892B9D32F801FCDEF840" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." box="[563,807,1968,1994]" pageId="45" pageNumber="46" pagination="29 - 38" refId="ref30906" refString="Sabu T. K., Nithya S. &amp; Vinod K. V. 2011. Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa 2830: 29 - 38. https: // doi. org / 10.11646 / ZOOTAXA. 2830.1.3" type="book chapter" year="2011">
Sabu
<emphasis id="EFACEAD6FFDB892B9D79F800FD4CF840" box="[632,693,1968,1994]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
2011: 34
</bibRefCitation>
</treatmentCitation>
(list). — Chandra
<emphasis id="EFACEAD6FFDB892B9B0CF800FBB3F840" box="[1037,1098,1968,1994]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
2012a: 52 (distribution). —
<treatmentCitation id="5C7910D5FFD889289E16FEB8FD37FEAE" author="Karimbumkara S. N. &amp; Rajan P. D." box="[279,718,265,292]" page="175" pageId="46" pageNumber="47" year="2013">
<bibRefCitation id="B9494B35FFD889289E16FEB8FD37FEAE" author="Karimbumkara S. N. &amp; Rajan P. D." box="[279,718,265,292]" pageId="46" pageNumber="47" pagination="173 - 178" refId="ref29842" refString="Karimbumkara S. N. &amp; Rajan P. D. 2013. Insecta: Coleoptera: Scarabaeidae: Scarabaeinae, Dung Beetles. Zoological Survey of India, Fauna of Karnataka, State Fauna Series 21: 173 - 178." type="book chapter" year="2013">Karimbumkara &amp; Rajan 2013: 175</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFD889289C56FEB8FB75FEA9" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[855,1164,265,292]" page="1037" pageId="46" pageNumber="47" year="2015">
<bibRefCitation id="B9494B35FFD889289C56FEB8FB75FEA9" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[855,1164,265,292]" pageId="46" pageNumber="47" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFD889289CAFFEBBFC09FEA9" box="[942,1008,265,291]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(list, distribution). —
<treatmentCitation id="5C7910D5FFD889289E0FFE9DFDD2FECC" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[270,555,300,326]" page="301" pageId="46" pageNumber="47" year="2014">
<bibRefCitation id="B9494B35FFD889289E0FFE9DFDD2FECC" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[270,555,300,326]" pageId="46" pageNumber="47" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFD889289E76FE9CFE48FECC" box="[375,433,300,326]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2014: 301
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFD889289DFCFE9DFC29FECC" author="Bezdek A." box="[765,976,300,326]" page="180" pageId="46" pageNumber="47" year="2016">
<bibRefCitation id="B9494B35FFD889289DFCFE9DFC29FECC" author="Bezdek A." box="[765,976,300,326]" pageId="46" pageNumber="47" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 180</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFD889289BA3FE9DFEEFFEE0" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." page="239" pageId="46" pageNumber="47" year="2020">
<bibRefCitation id="B9494B35FFD889289BA3FE9DFEEFFEE0" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." pageId="46" pageNumber="47" pagination="221 - 275" refId="ref29141" refString="Ghosh J., Das, P., Ghosh S. K., Bhunia D., Kushwaha R. K., Gupta D., &amp; Chandra K. 2020. Insecta: Coleoptera. Zoological Survey of India, Fauna of Haryana, State Fauna Series 24: 221 - 275." type="book chapter" year="2020">
Ghosh
<emphasis id="EFACEAD6FFD889289BF5FE9CFAD7FECC" box="[1268,1326,300,326]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2020: 239
</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFD889289EEDFEFEFD1FFEE0" author="Schoolmeesters P." box="[492,742,335,362]" pageId="46" pageNumber="47" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD889289FBCFE20FD53FDCA" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="DD6736C4FFD889289FBCFE20FE3FFE21" blockId="46.[189,454,401,427]" box="[189,454,401,427]" pageId="46" pageNumber="47">
<heading id="862F81A8FFD889289FBCFE20FE3FFE21" bold="true" box="[189,454,401,427]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="EFACEAD6FFD889289FBCFE20FE3FFE21" bold="true" box="[189,454,401,427]" pageId="46" pageNumber="47">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFD889289FBCFE71FA8EFE77" blockId="46.[189,1399,448,510]" pageId="46" pageNumber="47">
<emphasis id="EFACEAD6FFD889289FBCFE71FEB6FE50" box="[189,335,448,474]" italics="true" pageId="46" pageNumber="47">Onitisvirens</emphasis>
belongs to Group III and closely resembles
<taxonomicName id="1AD84D47FFD889289C4BFE71FC1FFE50" authorityName="Arrow" authorityYear="1931" box="[842,998,448,474]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFD889289C4BFE71FC1FFE50" box="[842,998,448,474]" italics="true" pageId="46" pageNumber="47">O. subopacus</emphasis>
</taxonomicName>
of the same group but differs from the latter in having a strongly punctured pronotum and clypeo-frontal carina which is narrowly interrupted.
</paragraph>
<paragraph id="DD6736C4FFD889289FBCFD94FD53FDCA" blockId="46.[189,682,549,576]" box="[189,682,549,576]" pageId="46" pageNumber="47">
Also see remarks under
<emphasis id="EFACEAD6FFD889289ED9FD94FD5AFDB5" box="[472,675,549,575]" italics="true" pageId="46" pageNumber="47">Onitissubopacus</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD889289FBCFDD6FB88F9B5" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="DD6736C4FFD889289FBCFDD6FE15FD0B" blockId="46.[189,492,615,641]" box="[189,492,615,641]" pageId="46" pageNumber="47">
<heading id="862F81A8FFD889289FBCFDD6FE15FD0B" bold="true" box="[189,492,615,641]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="EFACEAD6FFD889289FBCFDD6FE15FD0B" bold="true" box="[189,492,615,641]" pageId="46" pageNumber="47">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFD889289FEBFD24FEA3FD25" blockId="46.[189,1399,661,900]" box="[234,346,661,687]" pageId="46" pageNumber="47">
<emphasis id="EFACEAD6FFD889289FEBFD24FEA3FD25" bold="true" box="[234,346,661,687]" pageId="46" pageNumber="47">
<typeStatus id="02638866FFD889289FEBFD24FEA3FD25" box="[234,346,661,687]" pageId="46" pageNumber="47" type="syntype">Syntypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFD889289FBCFD08FCD0FC09" blockId="46.[189,1399,661,900]" pageId="46" pageNumber="47">
UNKNOWN TYPE LOCALTY •
<materialsCitation id="6DB03C99FFD889289D57FD0BFEA2FCB7" collectionCode="IRSNB" collectorName="A. Janssens &amp; Collection E. Candeze" determinerName="G. J. Arrow" location="Van" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Van" typeStatus="syntype">
<specimenCount id="CBDEFD4DFFD889289D57FD0BFD94FD5E" box="[598,621,698,724]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
;
<emphasis id="EFACEAD6FFD889289D7FFD0BFCDFFD5E" box="[638,806,698,724]" italics="true" pageId="46" pageNumber="47">“Onitisvirens</emphasis>
Lansb.,
<determinerName id="28BBDA00FFD889289C8EFD08FBE4FD5E" box="[911,1053,697,724]" pageId="46" pageNumber="47">G. J. Arrow</determinerName>
det.;
<collectorName id="702D5312FFD889289B60FD0BFB15FD5E" box="[1121,1260,698,724]" pageId="46" pageNumber="47">A. Janssens</collectorName>
vid., 1936:
<taxonomicName id="1AD84D47FFD889289FBCFD6CFE38FD7C" authority="Lansb." authorityName="Lansberge" authorityYear="1875" box="[189,449,732,759]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFD889289FBCFD6CFE90FD7C" box="[189,361,733,759]" italics="true" pageId="46" pageNumber="47">ONITIS virens</emphasis>
Lansb.
</taxonomicName>
;
<typeStatus id="02638866FFD889289ED3FD6CFDCFFD7D" box="[466,566,733,759]" pageId="46" pageNumber="47" type="syntype">Syntype</typeStatus>
; det
<collectingRegion id="1F1CF826FFD889289D71FD6CFD67FD7D" box="[624,670,733,759]" country="Turkey" name="Van" pageId="46" pageNumber="47">Van</collectingRegion>
Lansberge.
<emphasis id="EFACEAD6FFD889289C30FD6DFBFFFD7C" box="[817,1030,732,758]" italics="true" pageId="46" pageNumber="47">Onitisamplectens</emphasis>
Lansb.;
<taxonomicName id="1AD84D47FFD889289B71FD6DFB0EFD7C" box="[1136,1271,732,758]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplectens">
<emphasis id="EFACEAD6FFD889289B71FD6DFB0EFD7C" box="[1136,1271,732,758]" italics="true" pageId="46" pageNumber="47">Amplectens</emphasis>
</taxonomicName>
Type Lsb. Mercara;
<emphasis id="EFACEAD6FFD889289E2AFCB1FE47FC90" box="[299,446,768,794]" italics="true" pageId="46" pageNumber="47">Onitisvirens</emphasis>
Lansb.Révis.Arrow 1927; Ex-Typis de
<taxonomicName id="1AD84D47FFD889289C8BFCB1FB98FC90" authority="Lansb." authorityName="Lansb." box="[906,1121,768,794]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplectens">
<emphasis id="EFACEAD6FFD889289C8BFCB1FBF7FC90" box="[906,1038,768,794]" italics="true" pageId="46" pageNumber="47">amplectens</emphasis>
Lansb.
</taxonomicName>
;
<collectorName id="702D5312FFD889289B6CFCB1FA8AFC90" box="[1133,1395,768,794]" pageId="46" pageNumber="47">Collection E. Candeze</collectorName>
: 109”;
<collectionCode id="BBC9AE01FFD889289E04FC92FEA2FCB7" box="[261,347,803,829]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="46" pageNumber="47">IRSNB</collectionCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289E74FC92FCDDFC09" collectionCode="IRSNB" collectorName="A. Janssens &amp; Collection E. Candeze" location="Van" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Van" typeStatus="syntype">
<specimenCount id="CBDEFD4DFFD889289E74FC92FE75FCB7" box="[373,396,803,829]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
; “
<collectorName id="702D5312FFD889289EA7FC92FDC9FCB7" box="[422,560,803,829]" pageId="46" pageNumber="47">A. Janssens</collectorName>
vid., 1936:
<taxonomicName id="1AD84D47FFD889289DBCFC92FC44FCB7" authority="Lansb." authorityName="Lansberge" authorityYear="1875" box="[701,957,803,829]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFD889289DBCFC92FC9EFCB7" box="[701,871,803,829]" italics="true" pageId="46" pageNumber="47">ONITIS virens</emphasis>
Lansb.
</taxonomicName>
; cf. Ann. Soc. Ent. Belg. Xviii. 1875, p.135;
<typeStatus id="02638866FFD889289E0EFCF7FE8AFCEA" box="[271,371,838,864]" pageId="46" pageNumber="47" type="syntype">Syntype</typeStatus>
; Ex-Typis de
<taxonomicName id="1AD84D47FFD889289D14FCF7FD09FCEA" authority="Lansb." authorityName="Lansb." box="[533,752,838,864]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplectens">amplectens Lansb.</taxonomicName>
; AmplectensInd br Lansb.; Det.
<collectingRegion id="1F1CF826FFD889289B79FCF6FB5FFCEB" box="[1144,1190,839,865]" country="Turkey" name="Van" pageId="46" pageNumber="47">Van</collectingRegion>
Lansberge
<taxonomicName id="1AD84D47FFD889289A2EFCF6FE61FC09" authority="Lansb." authorityName="Lansb." class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplectens">
<emphasis id="EFACEAD6FFD889289A2EFCF6FEB8FC09" italics="true" pageId="46" pageNumber="47">Onitis amplectens</emphasis>
Lansb.
</taxonomicName>
;
<collectorName id="702D5312FFD889289EA6FCD8FD4DFC09" box="[423,692,873,900]" pageId="46" pageNumber="47">Collection E. Candeze</collectorName>
”;
<collectionCode id="BBC9AE01FFD889289DCBFCD8FCDDFC09" box="[714,804,873,899]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="46" pageNumber="47">IRSNB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="DD6736C4FFD889289FEBFC1AFCE2FC4F" blockId="46.[189,1400,939,1600]" box="[234,795,939,965]" pageId="46" pageNumber="47">
<emphasis id="EFACEAD6FFD889289FEBFC1AFD9BFC4F" bold="true" box="[234,610,939,965]" pageId="46" pageNumber="47">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFD889289D70FC1AFCEAFC4F" box="[625,787,939,965]" count="47" pageId="46" pageNumber="47" type="generic">47 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFD889289FBCFC7EFB88F9B5" blockId="46.[189,1400,939,1600]" pageId="46" pageNumber="47">
<materialsCitation id="6DB03C99FFD889289FBCFC7EFB62FC63" box="[189,1179,974,1001]" collectionCode="BMNH" country="India" location="INDIA" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFD889289FBCFC7EFEF2FC63" box="[189,267,975,1001]" name="India" pageId="46" pageNumber="47">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFD889289E2BFC7EFEBBFC63" box="[298,322,975,1001]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
; “
<taxonomicName id="1AD84D47FFD889289E5CFC7EFD33FC63" authority="Lansb., Compared" authorityName="Lansb., Compared" box="[349,714,975,1001]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFD889289E5CFC7EFE17FC62" box="[349,494,975,1001]" italics="true" pageId="46" pageNumber="47">Onitis virens</emphasis>
Lansb., Compared
</taxonomicName>
with
<typeStatus id="02638866FFD889289C11FC7EFCBAFC63" box="[784,835,975,1001]" pageId="46" pageNumber="47">type</typeStatus>
, G.J.A.”;
<collectionCode id="BBC9AE01FFD889289CB6FC7EFBF6FC63" box="[951,1039,975,1001]" pageId="46" pageNumber="47">BMNH</collectionCode>
(E) 1237153
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289BBBFC7EFC4DFB87" collectionCode="BMNH" collectorName="H. Stevens." country="India" location="Rungbong Vall." municipality="Gopaldhara" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Sikkim">
<specimenCount id="CBDEFD4DFFD889289BBBFC7EFB29FC63" box="[1210,1232,975,1001]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
; “
<collectingMunicipality id="3D03ACBEFFD889289BEAFC7EFA8AFC63" box="[1259,1395,975,1001]" pageId="46" pageNumber="47">Gopaldhara</collectingMunicipality>
,
<location id="D807601FFFD889289FBCFC43FE92FB86" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289FBCFC43FE92FB86" box="[189,363,1010,1036]" country="India" municipality="Gopaldhara" name="Rungbong Vall." pageId="46" pageNumber="47" stateProvince="Sikkim">Rungbong Vall.</location>
,
<collectingRegion id="1F1CF826FFD889289E73FC43FE3FFB86" box="[370,454,1010,1036]" country="India" name="Sikkim" pageId="46" pageNumber="47">Sikkim</collectingRegion>
,
<collectorName id="702D5312FFD889289ECCFC43FDB5FB86" box="[461,588,1010,1036]" pageId="46" pageNumber="47">H.Stevens.</collectorName>
, 1916-218;
<collectionCode id="BBC9AE01FFD889289DC8FC43FCD8FB86" box="[713,801,1010,1036]" pageId="46" pageNumber="47">BMNH</collectionCode>
(E)1237154”
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289CC5FC43FE42FBA5" collectingDate="1927-02-23" country="India" county="Nerbuda Survey" elevation="823" location="Bewa State" municipality="Harai" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFD889289CC5FC43FC25FB86" box="[964,988,1010,1036]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
; “
<collectingCounty id="34064E48FFD889289CEEFC43FB5CFB86" box="[1007,1189,1010,1036]" pageId="46" pageNumber="47">Nerbuda Survey</collectingCounty>
,
<collectingMunicipality id="3D03ACBEFFD889289BADFC43FB14FB86" box="[1196,1261,1010,1036]" pageId="46" pageNumber="47">Harai</collectingMunicipality>
,
<location id="D807601FFFD889289BF5FC43FA8AFB86" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289BF5FC43FA8AFB86" box="[1268,1395,1010,1036]" country="India" county="Nerbuda Survey" municipality="Harai" name="Bewa State" pageId="46" pageNumber="47">Bewa State</location>
,
<quantity id="1A209B21FFD889289FBCFBA4FEECFBA5" box="[189,277,1045,1071]" metricMagnitude="2" metricUnit="m" metricValue="8.2296" pageId="46" pageNumber="47" unit="ft" value="2700.0">
<elevation id="56F5D1F7FFD889289FBCFBA4FEECFBA5" box="[189,277,1045,1071]" metricMagnitude="2" metricUnit="m" metricValue="8.2296" pageId="46" pageNumber="47" unit="ft" value="2700.0">2700 ft.</elevation>
</quantity>
;
<date id="A9661004FFD889289E1FFBA4FE49FBA5" box="[286,432,1045,1071]" pageId="46" pageNumber="47" value="1927-02-23">
<collectingDate id="B922E9ECFFD889289E1FFBA4FE49FBA5" box="[286,432,1045,1071]" pageId="46" pageNumber="47" value="1927-02-23">23 Feb. 1927</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289ECFFBA7FDCDFBA5" box="[462,564,1045,1072]" collectionCode="ZSIM" country="India" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFD889289ECFFBA7FE1DFBBA" box="[462,484,1046,1072]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
;
<collectionCode id="BBC9AE01FFD889289EF1FBA4FDCDFBA5" box="[496,564,1045,1071]" pageId="46" pageNumber="47">ZSIM</collectionCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289D49FBA4FD83FBFC" collectingDate="1999-05-14" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="11.837999" location="Budipadaga" longLatPrecision="1" longitude="77.07755" municipality="Biligiri Rangaswamy Tiger Reserve" pageId="46" pageNumber="47" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFD889289D49FBA4FD71FBBA" box="[584,648,1045,1072]" count="2" pageId="46" pageNumber="47" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD889289D93FBA4FD28FBBA" box="[658,721,1045,1072]" count="2" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFD889289DDCFBA4FCABFBA5" box="[733,850,1045,1071]" country="India" name="Karnataka" pageId="46" pageNumber="47">Karnataka</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD889289C5AFBA4FB11FBBA" box="[859,1256,1045,1072]" pageId="46" pageNumber="47">Biligiri Rangaswamy Tiger Reserve</collectingMunicipality>
,
<location id="D807601FFFD889289BF0FBA4FA8EFBA5" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289BF0FBA4FA8EFBA5" box="[1265,1399,1045,1071]" country="India" latitude="11.837999" longLatPrecision="1" longitude="77.07755" municipality="Biligiri Rangaswamy Tiger Reserve" name="Budipadaga" pageId="46" pageNumber="47" stateProvince="Karnataka">Budipadaga</location>
scrub;
<geoCoordinate id="B8EC5003FFD889289E09FB86FE53FBD9" box="[264,426,1079,1107]" degrees="11" direction="north" minutes="50" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="16.8" value="11.837999">11°50ʹ16.8″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289EB7FB86FDAFFBD9" box="[438,598,1079,1107]" degrees="77" direction="east" minutes="4" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="39.18" value="77.07755">77°4ʹ39.18″ E</geoCoordinate>
;
<date id="A9661004FFD889289D63FB89FD02FBD8" box="[610,763,1080,1107]" pageId="46" pageNumber="47" value="1999-05-14">
<collectingDate id="B922E9ECFFD889289D63FB89FD02FBD8" box="[610,763,1080,1107]" pageId="46" pageNumber="47" value="1999-05-14">14 May 1999</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289C06FB88FBDDFBD9" box="[775,1060,1080,1107]" pageId="46" pageNumber="47">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFD889289B2EFB89FA8EFBD8" box="[1071,1399,1080,1106]" pageId="46" pageNumber="47">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD889289FF7FBEAFEC6FBFF" box="[246,319,1115,1141]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<collectorName id="702D5312FFD889289E49FBEAFE91FBFF" box="[328,360,1115,1141]" pageId="46" pageNumber="47">Co</collectorName>
/
<collectorName id="702D5312FFD889289E70FBEAFE75FBFF" box="[369,396,1115,1141]" pageId="46" pageNumber="47">Sc</collectorName>
1000746 to 1000749
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289D9BFBEDFE3CFB36" collectingDate="1999-12-22" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="11.89545" location="Kanneri" longLatPrecision="1" longitude="77.13596" pageId="46" pageNumber="47" specimenCount="13" specimenCount-female="5" specimenCount-male="8">
<specimenCount id="CBDEFD4DFFD889289D9BFBEDFD19FBFC" box="[666,736,1116,1142]" count="8" pageId="46" pageNumber="47" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD889289DEEFBEAFCCDFBFC" box="[751,820,1115,1142]" count="5" pageId="46" pageNumber="47" type="female">5 ♀♀</specimenCount>
;
<location id="D807601FFFD889289C44FBEAFC66FBFF" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289C44FBEAFC66FBFF" box="[837,927,1115,1141]" country="India" latitude="11.89545" longLatPrecision="1" longitude="77.13596" name="Kanneri" pageId="46" pageNumber="47">Kanneri</location>
moist deciduous forest;
<geoCoordinate id="B8EC5003FFD889289BBFFBEBFA8AFBFC" box="[1214,1395,1114,1142]" degrees="11" direction="north" minutes="53" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="43.62" value="11.89545">11°53ʹ43.62″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289FBCFBCCFEACFB13" box="[189,341,1149,1177]" degrees="77" direction="east" minutes="8" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="9.48" value="77.13596">77°8ʹ9.48″ E</geoCoordinate>
;
<date id="A9661004FFD889289E66FBCEFDF5FB13" box="[359,524,1151,1177]" pageId="46" pageNumber="47" value="1999-12-22">
<collectingDate id="B922E9ECFFD889289E66FBCEFDF5FB13" box="[359,524,1151,1177]" pageId="46" pageNumber="47" value="1999-12-22">22 Dec. 1999</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289D1CFBCEFCA8FB13" box="[541,849,1151,1177]" pageId="46" pageNumber="47">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFD889289C63FBCEFB4DFB13" box="[866,1204,1151,1177]" pageId="46" pageNumber="47">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD889289BFCFBCEFABFFB13" box="[1277,1350,1151,1177]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<collectorName id="702D5312FFD889289A4FFBCEFA97FB13" box="[1358,1390,1151,1177]" pageId="46" pageNumber="47">Co</collectorName>
/
<collectorName id="702D5312FFD889289FBCFB13FF21FB37" box="[189,216,1186,1213]" pageId="46" pageNumber="47">Sc</collectorName>
1000750 to 1000762
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289EE3FB13FB11FB55" collectingDate="1999-05-11" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="12.032" location="Purani" longLatPrecision="1" longitude="77.1313" pageId="46" pageNumber="47" specimenCount="2" specimenCount-male="2">
<specimenCount id="CBDEFD4DFFD889289EE3FB13FDDEFB36" box="[482,551,1186,1212]" count="2" pageId="46" pageNumber="47" type="male">2 ♂♂</specimenCount>
,
<location id="D807601FFFD889289D35FB13FD84FB36" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289D35FB13FD84FB36" box="[564,637,1186,1212]" country="India" latitude="12.032" longLatPrecision="1" longitude="77.1313" name="Purani" pageId="46" pageNumber="47">Purani</location>
dry deciduous forest;
<geoCoordinate id="B8EC5003FFD889289C7EFB11FBEFFB36" box="[895,1046,1184,1212]" degrees="12" direction="north" minutes="1" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="55.2" value="12.032">12°1ʹ55.2″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289B25FB11FB3EFB36" box="[1060,1223,1184,1212]" degrees="77" direction="east" minutes="7" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="52.68" value="77.1313">77°7ʹ52.68″ E</geoCoordinate>
;
<date id="A9661004FFD889289BD4FB13FA8AFB36" box="[1237,1395,1186,1212]" pageId="46" pageNumber="47" value="1999-05-11">
<collectingDate id="B922E9ECFFD889289BD4FB13FA8AFB36" box="[1237,1395,1186,1212]" pageId="46" pageNumber="47" value="1999-05-11">11 May 1999</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289FBCFB77FE23FB6A" box="[189,474,1221,1248]" pageId="46" pageNumber="47">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFD889289EE7FB74FCD7FB55" box="[486,814,1221,1247]" pageId="46" pageNumber="47">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD889289C6AFB74FC4DFB55" box="[875,948,1221,1247]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<collectorName id="702D5312FFD889289CBCFB74FC24FB55" box="[957,989,1221,1247]" pageId="46" pageNumber="47">Co</collectorName>
/
<collectorName id="702D5312FFD889289CE7FB74FBF8FB55" box="[998,1025,1221,1247]" pageId="46" pageNumber="47">Sc</collectorName>
1000763 to 1000764
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289A00FB77FAE3FAAC" collectingDate="2009-12-03" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" elevation="207" latitude="8.918222" location="Kattilapara" longLatPrecision="1" longitude="77.09575" municipality="Shendurney Wildlife Sanctuary" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFD889289A00FB77FAE0FB6A" box="[1281,1305,1222,1248]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFD889289A24FB74FA8AFB55" box="[1317,1395,1221,1247]" country="India" name="Kerala" pageId="46" pageNumber="47">Kerala</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD889289FBCFB59FDE3FA88" box="[189,538,1256,1282]" pageId="46" pageNumber="47">Shendurney Wildlife Sanctuary</collectingMunicipality>
,
<location id="D807601FFFD889289D24FB59FD59FA88" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289D24FB59FD59FA88" box="[549,672,1256,1282]" country="India" latitude="8.918222" longLatPrecision="1" longitude="77.09575" municipality="Shendurney Wildlife Sanctuary" name="Kattilapara" pageId="46" pageNumber="47" stateProvince="Kerala">Kattilapara</location>
degraded forest;
<geoCoordinate id="B8EC5003FFD889289C61FB56FBF9FA89" box="[864,1024,1255,1283]" degrees="08" direction="north" minutes="55" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="05.6" value="8.918222">08°55ʹ05.6ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289B0AFB56FB50FA89" box="[1035,1193,1255,1283]" degrees="77" direction="east" minutes="05" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="44.7" value="77.09575">77°05ʹ44.7ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD889289BEFFB59FA8BFA88" box="[1262,1394,1256,1283]" metricMagnitude="2" metricUnit="m" metricValue="2.07" pageId="46" pageNumber="47" unit="m" value="207.0">
<quantity id="1A209B21FFD889289BEFFB59FACFFA89" box="[1262,1334,1256,1283]" metricMagnitude="2" metricUnit="m" metricValue="2.07" pageId="46" pageNumber="47" unit="m" value="207.0">207 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD889289FBCFABDFEB1FAAC" box="[189,328,1292,1318]" pageId="46" pageNumber="47" value="2009-12-03">
<collectingDate id="B922E9ECFFD889289FBCFABDFEB1FAAC" box="[189,328,1292,1318]" pageId="46" pageNumber="47" value="2009-12-03">3 Dec. 2009</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289E53FABDFD92FAAC" box="[338,619,1291,1318]" pageId="46" pageNumber="47">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFD889289D77FABDFC1EFAAF" box="[630,999,1291,1318]" pageId="46" pageNumber="47">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD889289B22FABAFB95FAAC" box="[1059,1132,1291,1318]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<collectorName id="702D5312FFD889289B74FABAFB6CFAAC" box="[1141,1173,1291,1318]" pageId="46" pageNumber="47">Co</collectorName>
/
<collectorName id="702D5312FFD889289B9FFABAFB40FAAC" box="[1182,1209,1291,1318]" pageId="46" pageNumber="47">Sc</collectorName>
1000765
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289A31FABDFE38FA05" collectingDate="2018-09-30" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="127" latitude="27.96913" location="Rani Village" longLatPrecision="1" longitude="95.32144" municipality="Pasighat" pageId="46" pageNumber="47" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFD889289A31FABDFA8BFAAC" box="[1328,1394,1292,1318]" count="2" pageId="46" pageNumber="47" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD889289FBCFA9EFEF8FAC3" box="[189,257,1327,1353]" count="2" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFD889289E0EFA9EFE1EFAC3" box="[271,487,1327,1353]" country="India" name="Arunachal Pradesh" pageId="46" pageNumber="47">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD889289EF5FA9EFDACFAC3" box="[500,597,1327,1353]" pageId="46" pageNumber="47">Pasighat</collectingMunicipality>
, community forest,
<location id="D807601FFFD889289C3BFA9EFC35FAC3" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289C3BFA9EFC35FAC3" box="[826,972,1327,1353]" country="India" latitude="27.96913" longLatPrecision="1" longitude="95.32144" municipality="Pasighat" name="Rani Village" pageId="46" pageNumber="47" stateProvince="Arunachal Pradesh">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD889289CD8FA9CFB60FAC3" box="[985,1177,1325,1353]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="8.8644" value="27.96913">27°58ʹ8.8644ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289BA6FA9CFA8BFAC3" box="[1191,1394,1325,1353]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="17.1804" value="95.32144">95°19ʹ17.1804ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD889289FFBFAE3FE7AFAE6" box="[250,387,1362,1388]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="46" pageNumber="47" unit="m" value="127.0">
<quantity id="1A209B21FFD889289FFBFAE3FEBDFAE6" box="[250,324,1362,1388]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="46" pageNumber="47" unit="m" value="127.0">127 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD889289E90FAE3FDD7FAE6" box="[401,558,1362,1388]" pageId="46" pageNumber="47" value="2018-09-30">
<collectingDate id="B922E9ECFFD889289E90FAE3FDD7FAE6" box="[401,558,1362,1388]" pageId="46" pageNumber="47" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289D3AFAE3FC84FAE6" box="[571,893,1362,1388]" pageId="46" pageNumber="47">cow dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFD889289C8BFAE3FBB0FAE6" box="[906,1097,1362,1388]" pageId="46" pageNumber="47">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFD889289BFCFAE3FABFFAE6" box="[1277,1350,1362,1388]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<collectorName id="702D5312FFD889289A4FFAE3FA97FAE6" box="[1358,1390,1362,1388]" pageId="46" pageNumber="47">Co</collectorName>
/
<collectorName id="702D5312FFD889289FBCFAC4FF21FA05" box="[189,216,1397,1423]" pageId="46" pageNumber="47">Sc</collectorName>
1000766 to 1000769
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289ED9FAC4FE47FA5C" collectingDate="2018-09-30" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="131" latitude="27.969614" location="Rani Village" longLatPrecision="1" longitude="95.32392" pageId="46" pageNumber="47" specimenCount="15" specimenCount-female="9" specimenCount-male="6">
<specimenCount id="CBDEFD4DFFD889289ED9FAC4FDE2FA1A" box="[472,539,1397,1424]" count="6" pageId="46" pageNumber="47" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD889289D27FAC4FD9EFA1A" box="[550,615,1397,1424]" count="9" pageId="46" pageNumber="47" type="female">9 ♀♀</specimenCount>
; agricultural field,
<location id="D807601FFFD889289C3BFAC4FC33FA05" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289C3BFAC4FC33FA05" box="[826,970,1397,1423]" country="India" latitude="27.969614" longLatPrecision="1" longitude="95.32392" name="Rani Village" pageId="46" pageNumber="47">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD889289CD5FAC2FB66FA1A" box="[980,1183,1395,1424]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="10.6068" value="27.969614">27°58ʹ10.6068ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289BAAFAC2FA8BFA1A" box="[1195,1394,1395,1424]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="26.1156" value="95.32392">95°19ʹ26.1156ʺ E</geoCoordinate>
, elev.
<elevation id="56F5D1F7FFD889289FF8FA29FE7BFA38" box="[249,386,1432,1459]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="46" pageNumber="47" unit="m" value="131.0">
<quantity id="1A209B21FFD889289FF8FA29FEBDFA39" box="[249,324,1432,1459]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="46" pageNumber="47" unit="m" value="131.0">131 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD889289E91FA29FDD2FA38" box="[400,555,1432,1458]" pageId="46" pageNumber="47" value="2018-09-30">
<collectingDate id="B922E9ECFFD889289E91FA29FDD2FA38" box="[400,555,1432,1458]" pageId="46" pageNumber="47" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFD889289C8DFA29FBB3FA38" box="[908,1098,1432,1458]" pageId="46" pageNumber="47">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFD889289BFCFA29FABFFA38" box="[1277,1350,1432,1458]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<collectorName id="702D5312FFD889289A4FFA29FA97FA38" box="[1358,1390,1432,1458]" pageId="46" pageNumber="47">Co</collectorName>
/
<collectorName id="702D5312FFD889289FBCFA0DFF21FA5C" box="[189,216,1468,1494]" pageId="46" pageNumber="47">Sc</collectorName>
1000770 to 1000784
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289ED5FA0DFEC2F996" collectingDate="2018-09-30" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="133" latitude="27.969172" location="Rani Village" longLatPrecision="1" longitude="95.32194" pageId="46" pageNumber="47" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFD889289ED5FA0DFE12FA5C" box="[468,491,1468,1494]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFD889289EF7FA0DFDCFFA5C" box="[502,566,1468,1494]" count="2" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
; community forest,
<location id="D807601FFFD889289C12FA0AFC58FA5F" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289C12FA0AFC58FA5F" box="[787,929,1467,1493]" country="India" latitude="27.969172" longLatPrecision="1" longitude="95.32194" name="Rani Village" pageId="46" pageNumber="47">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD889289CAAFA0BFB9EFA5C" box="[939,1127,1466,1494]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="9.0156" value="27.969172">27°58ʹ9.0156ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289B70FA0BFAC1FA5C" box="[1137,1336,1466,1494]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="18.9804" value="95.32194">95°19ʹ18.9804ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD889289FBCFA6EFEC7FA73" box="[189,318,1503,1529]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="46" pageNumber="47" unit="m" value="133.0">
<quantity id="1A209B21FFD889289FBCFA6EFEFDFA73" box="[189,260,1503,1529]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="46" pageNumber="47" unit="m" value="133.0">133 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD889289E49FA6EFE24FA73" box="[328,477,1503,1529]" pageId="46" pageNumber="47" value="2018-09-30">
<collectingDate id="B922E9ECFFD889289E49FA6EFE24FA73" box="[328,477,1503,1529]" pageId="46" pageNumber="47" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289EE7FA6EFCEEFA73" box="[486,791,1503,1529]" pageId="46" pageNumber="47">cow dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFD889289C20FA6EFC22FA73" box="[801,987,1503,1529]" pageId="46" pageNumber="47">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFD889289B7EFA6FFB30FA73" box="[1151,1225,1502,1529]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<collectorName id="702D5312FFD889289BD3FA6FFB0BFA73" box="[1234,1266,1502,1529]" pageId="46" pageNumber="47">Co</collectorName>
/
<collectorName id="702D5312FFD889289BFAFA6FFAEFFA73" box="[1275,1302,1502,1529]" pageId="46" pageNumber="47">Sc</collectorName>
1000785 to 1000787
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289E51F9B3FB94F9B5" collectingDate="2021-04-15" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" location="Intanki National Park" municipality="Bongkolong village" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Nagaland">
<specimenCount id="CBDEFD4DFFD889289E51F9B3FE9FF996" box="[336,358,1538,1564]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFD889289E72F9B3FE18F996" box="[371,481,1538,1564]" country="India" name="Nagaland" pageId="46" pageNumber="47">Nagaland</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD889289EEBF9B3FD31F996" box="[490,712,1538,1564]" pageId="46" pageNumber="47">Bongkolong village</collectingMunicipality>
, near
<location id="D807601FFFD889289C09F9B3FC01F996" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289C09F9B3FC01F996" box="[776,1016,1538,1564]" country="India" municipality="Bongkolong village" name="Intanki National Park" pageId="46" pageNumber="47" stateProvince="Nagaland">Intanki National Park</location>
, open ground;
<date id="A9661004FFD889289B9BF9B3FAD5F996" box="[1178,1324,1538,1565]" pageId="46" pageNumber="47" value="2021-04-15">
<collectingDate id="B922E9ECFFD889289B9BF9B3FAD5F996" box="[1178,1324,1538,1565]" pageId="46" pageNumber="47" value="2021-04-15">15 Apr. 2021</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFD889289EC7F994FCCEF9B5" box="[454,823,1573,1600]" pageId="46" pageNumber="47">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD889289C73F994FC45F9B5" box="[882,956,1573,1599]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<collectorName id="702D5312FFD889289CC7F994FC1EF9B5" box="[966,999,1573,1599]" pageId="46" pageNumber="47">Co</collectorName>
/
<collectorName id="702D5312FFD889289CEEF994FBF3F9B5" box="[1007,1034,1573,1599]" pageId="46" pageNumber="47">Sc</collectorName>
1000788
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD889359FBCF9D6FB94FE21" lastPageId="51" lastPageNumber="52" pageId="46" pageNumber="47" type="description">
<paragraph id="DD6736C4FFD889289FBCF9D6FEB7F90B" blockId="46.[189,334,1639,1665]" box="[189,334,1639,1665]" pageId="46" pageNumber="47">
<heading id="862F81A8FFD889289FBCF9D6FEB7F90B" bold="true" box="[189,334,1639,1665]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="EFACEAD6FFD889289FBCF9D6FEB7F90B" bold="true" box="[189,334,1639,1665]" pageId="46" pageNumber="47">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFD889289FBCF927FCE2F93B" blockId="46.[189,795,1686,1713]" box="[189,795,1686,1713]" pageId="46" pageNumber="47">MEASUREMENTS (in mm). TL=1723, BW= 9.513.</paragraph>
<paragraph id="DD6736C4FFD889289FEBF969FED3F978" blockId="46.[189,1399,1752,2025]" box="[234,298,1752,1778]" pageId="46" pageNumber="47">
<emphasis id="EFACEAD6FFD889289FEBF969FED3F978" bold="true" box="[234,298,1752,1778]" pageId="46" pageNumber="47">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFD889289FBCF94AFAA7F862" blockId="46.[189,1399,1752,2025]" pageId="46" pageNumber="47">Black with a very feeble metallic lustre; clypeus elliptical, margin entire, closely and finely rugulose, vertex rugulose and more shining, frontal carina interrupted by a very short medial gap, with a conical tubercle behind and an anterior short transverse clypeal carina; pronotum fairly closely and strongly punctured, with an anterior narrow smooth median line and slight narrow groove behind; elytra finely striate with intervals flat and imperceptibly punctured, the sutural interval slightly elevated and shining; pygidium flat, smooth and feebly punctured; metasternal shield smooth and shining behind, front angles and sides of metasternum clothed with red hairs. Protibia slender, curved, mid femur and hind trochanter toothed.</paragraph>
<caption id="89A7664CFFD989299FBCF83BFE4BF860" pageId="47" pageNumber="48" startId="47.[189,232,1930,1956]" targetBox="[223,1365,249,1879]" targetPageId="47">
<paragraph id="DD6736C4FFD989299FBCF83BFE4BF860" blockId="47.[189,1399,1930,2026]" pageId="47" pageNumber="48">
<emphasis id="EFACEAD6FFD989299FBCF83BFEE5F82E" bold="true" box="[189,284,1930,1956]" pageId="47" pageNumber="48">Fig. 17.</emphasis>
Map showing the distribution of
<emphasis id="EFACEAD6FFD989299DB6F83BFC6DF82E" box="[695,916,1930,1956]" italics="true" pageId="47" pageNumber="48">Onitisassamensis</emphasis>
<bibRefCitation id="B9494B35FFD989299C96F83BFBC0F82E" author="Biswas S." box="[919,1081,1930,1956]" pageId="47" pageNumber="48" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980</bibRefCitation>
;
<taxonomicName id="1AD84D47FFD989299B4AF83BFAFEF82E" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1099,1287,1930,1956]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bhomorensis" status="sp. nov.">
<emphasis id="EFACEAD6FFD989299B4AF83BFAFEF82E" box="[1099,1287,1930,1956]" italics="true" pageId="47" pageNumber="48">O. bhomorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFD989299A10F83BFA8BF82E" box="[1297,1394,1930,1956]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="1AD84D47FFD989299FBCF81CFDF1F84D" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[189,520,1965,1991]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bordati">
<emphasis id="EFACEAD6FFD989299FBCF81CFEC6F84D" box="[189,319,1965,1991]" italics="true" pageId="47" pageNumber="48">O. bordati</emphasis>
<bibRefCitation id="B9494B35FFD989299E41F81CFDF1F84D" author="Cambefort Y." box="[320,520,1965,1991]" pageId="47" pageNumber="48" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1AD84D47FFD989299D19F81CFC9CF842" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[536,869,1965,1992]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFD989299D19F81CFD61F84D" box="[536,664,1965,1991]" italics="true" pageId="47" pageNumber="48">O. brahma</emphasis>
<bibRefCitation id="B9494B35FFD989299DA1F81CFC9CF842" author="Lansberge G." box="[672,869,1965,1992]" pageId="47" pageNumber="48" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFD989299CA1F81CFAD2F84D" authority="Redtenbacher, 1848" authorityName="Redtenbacher" authorityYear="1848" box="[928,1323,1965,1991]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFD989299CA1F81CFBC0F84D" box="[928,1081,1965,1991]" italics="true" pageId="47" pageNumber="48">O. castaneus</emphasis>
Redtenbacher, 1848
</taxonomicName>
in the Indian Subcontinent.
</paragraph>
</caption>
<caption id="89A7664CFFC689369FBFF836FE04F862" pageId="48" pageNumber="49" startId="48.[190,233,1927,1953]" targetBox="[224,1364,251,1878]" targetPageId="48">
<paragraph id="DD6736C4FFC689369FBFF836FE04F862" blockId="48.[190,1400,1927,2024]" pageId="48" pageNumber="49">
<emphasis id="EFACEAD6FFC689369FBFF836FED8F82B" bold="true" box="[190,289,1927,1953]" pageId="48" pageNumber="49">Fig. 18.</emphasis>
Map showing the distribution of
<taxonomicName id="1AD84D47FFC689369DCAF839FBE2F828" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[715,1051,1927,1954]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFC689369DCAF839FC8EF82B" box="[715,887,1928,1954]" italics="true" pageId="48" pageNumber="49">Onitis crassus</emphasis>
Sharp, 1875
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC689369B2DF839FA8AF828" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[1068,1395,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="15" phylum="Arthropoda" rank="species" species="excavatus">
O.
<emphasis id="EFACEAD6FFC689369B57F839FB33F82B" box="[1110,1226,1928,1953]" italics="true" pageId="48" pageNumber="49">excavatus</emphasis>
<bibRefCitation id="B9494B35FFC689369BD7F839FA8AF828" author="Arrow G. J." box="[1238,1395,1928,1954]" pageId="48" pageNumber="49" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC689369FBFF81AFE06F84F" authority="(Wulfen, 1789)" baseAuthorityName="Wulfen" baseAuthorityYear="1789" box="[190,511,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFC689369FBFF81AFEBFF84F" box="[190,326,1963,1989]" italics="true" pageId="48" pageNumber="49">O. falcatus</emphasis>
(Wulfen, 1789)
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC689369D0AF81AFCEFF84F" authority="Felsche, 1907" authorityName="Felsche" authorityYear="1907" box="[523,790,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="feae">
<emphasis id="EFACEAD6FFC689369D0AF81AFD99F84F" box="[523,608,1963,1989]" italics="true" pageId="48" pageNumber="49">O. feae</emphasis>
<bibRefCitation id="B9494B35FFC689369D6EF81AFCEFF84F" author="Felsche C." box="[623,790,1963,1989]" pageId="48" pageNumber="49" pagination="273 - 296" refId="ref29092" refString="Felsche C. 1907. Coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift: 273 - 296." type="book chapter" year="1907">Felsche, 1907</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC689369C22F81AFB89F84F" authority="(Pallas, 1771)" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[803,1136,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFC689369C22F81AFC3CF84F" box="[803,965,1963,1989]" italics="true" pageId="48" pageNumber="49">O. humerosus</emphasis>
(
<bibRefCitation id="B9494B35FFC689369CD4F81AFB9EF84F" author="Pallas P. S." box="[981,1127,1963,1989]" pageId="48" pageNumber="49" refId="ref30468" refString="Pallas P. S. 1771. Reise durch verschiedene Provinzen des Russischen Reiches in den Jahren 1768 - 1774 I: 1 - 504 (453 - 504). Petersburg Akademische Buchhandlung, St Petersburg." type="book" year="1771">Pallas, 1771</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="1AD84D47FFC689369BABF81AFAEFF84F" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1194,1302,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov.">
<emphasis id="EFACEAD6FFC689369BABF81AFAEFF84F" box="[1194,1302,1963,1989]" italics="true" pageId="48" pageNumber="49">O. kethai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFC689369A1CF81AFA81F84F" box="[1309,1400,1963,1989]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
in the Indian Subcontinent.
</paragraph>
</caption>
<caption id="89A7664CFFC789379FBCF836FE02F862" pageId="49" pageNumber="50" startId="49.[189,232,1927,1953]" targetBox="[227,1362,257,1876]" targetPageId="49">
<paragraph id="DD6736C4FFC789379FBCF836FE02F862" blockId="49.[189,1399,1927,2024]" pageId="49" pageNumber="50">
<emphasis id="EFACEAD6FFC789379FBCF836FEE2F82B" bold="true" box="[189,283,1927,1953]" pageId="49" pageNumber="50">Fig. 19.</emphasis>
Map showing the distribution of
<emphasis id="EFACEAD6FFC789379DAAF839FCCFF828" box="[683,822,1928,1954]" italics="true" pageId="49" pageNumber="50">Onitislama</emphasis>
<bibRefCitation id="B9494B35FFC789379C3EF839FBFFF828" author="Lansberge G." box="[831,1030,1927,1954]" pageId="49" pageNumber="50" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
;
<taxonomicName id="1AD84D47FFC789379B15F839FA8BF828" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[1044,1394,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFC789379B15F839FADAF828" box="[1044,1315,1928,1954]" italics="true" pageId="49" pageNumber="50">
O. naviauxi
<bibRefCitation id="B9494B35FFC789379BA7F839FADAF828" author="Cambefort Y." box="[1190,1315,1928,1954]" pageId="49" pageNumber="50" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort</bibRefCitation>
</emphasis>
, 1988
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC789379FBCF81AFDEBF84F" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" box="[189,530,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFC789379FBCF81AFEB6F84F" box="[189,335,1963,1989]" italics="true" pageId="49" pageNumber="50">O. philemon</emphasis>
<bibRefCitation id="B9494B35FFC789379E56F81AFDEBF84F" author="Fabricius J. C." box="[343,530,1963,1989]" pageId="49" pageNumber="50" refId="ref29058" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Volume 2. Impensis Bibliopolii novi academici, Kiel." type="book" year="1801">Fabricius, 1801</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC789379D1EF81AFC43F84F" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[543,954,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFC789379D1EF81AFCFAF84F" box="[543,771,1963,1989]" italics="true" pageId="49" pageNumber="50">O. punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFC789379C0BF81AFC43F84F" author="Janssens A." box="[778,954,1963,1989]" pageId="49" pageNumber="50" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFC789379CF4F81AFA8EF84C" authority="Lansberge, 1875" authorityName="Lansberge, AIMB" authorityYear="1875" box="[1013,1399,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFC789379CF4F81AFB53F84F" box="[1013,1194,1963,1989]" italics="true" pageId="49" pageNumber="50">O. singhalensis</emphasis>
<bibRefCitation id="B9494B35FFC789379BB2F81AFA8EF84C" author="Lansberge G." box="[1203,1399,1963,1990]" pageId="49" pageNumber="50" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
in the Indian Subcontinent.
</paragraph>
</caption>
<caption id="89A7664CFFC489349FBCF83BFC1DF84D" pageId="50" pageNumber="51" startId="50.[189,232,1930,1956]" targetBox="[230,1361,265,1880]" targetPageId="50">
<paragraph id="DD6736C4FFC489349FBCF83BFC1DF84D" blockId="50.[189,1399,1930,1992]" pageId="50" pageNumber="51">
<emphasis id="EFACEAD6FFC489349FBCF83BFEE2F82E" bold="true" box="[189,283,1930,1956]" pageId="50" pageNumber="51">Fig. 20.</emphasis>
Map showing the distribution of
<emphasis id="EFACEAD6FFC489349DA2F83BFCD8F82E" box="[675,801,1930,1956]" italics="true" pageId="50" pageNumber="51">Onitissiva</emphasis>
<bibRefCitation id="B9494B35FFC489349C28F83BFC4EF82E" author="Gillet J. J. E." box="[809,951,1930,1956]" pageId="50" pageNumber="51" pagination="313 - 314" refId="ref29108" refString="Gillet J. J. E. 1911. Coprides nouveaux de la region orientale (etc.). Annales de la Societe entomologique de Belgique LV: 313 - 314." type="book chapter" year="1911">Gillet, 1911</bibRefCitation>
;
<taxonomicName id="1AD84D47FFC489349CC2F83BFAF9F82E" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[963,1280,1930,1956]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFC489349CC2F83BFB9AF82E" box="[963,1123,1930,1956]" italics="true" pageId="50" pageNumber="51">O. subopacus</emphasis>
<bibRefCitation id="B9494B35FFC489349B68F83BFAF9F82E" author="Arrow G. J." box="[1129,1280,1930,1956]" pageId="50" pageNumber="51" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC489349A0DF83BFE79F842" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFC489349A0DF83BFA8EF82E" box="[1292,1399,1930,1956]" italics="true" pageId="50" pageNumber="51">O. virens</emphasis>
<bibRefCitation id="B9494B35FFC489349FBCF81CFE79F842" author="Lansberge G." box="[189,384,1965,1992]" pageId="50" pageNumber="51" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFC489349EB8F81CFDC4F84D" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[441,573,1965,1991]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov.">
<emphasis id="EFACEAD6FFC489349EB8F81CFDC4F84D" box="[441,573,1965,1991]" italics="true" pageId="50" pageNumber="51">O. visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFC489349D45F81FFD66F842" box="[580,671,1966,1992]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
in the Indian Subcontinent.
</paragraph>
</caption>
<paragraph id="DD6736C4FFC589359FBCFEB8FC38FECC" blockId="51.[189,1398,265,327]" pageId="51" pageNumber="52">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFC589359EB2FEB8FDB2FEA9" box="[435,587,265,291]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. EH. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." pageId="51" pageNumber="52">Fig. 13E H</figureCitation>
). LP=3.5, Lp=3; BP=1.6, BpB=0.7, BpT=0.5. Phallobase slightly longer than the paramere, both straight below and sharp at the tip.
</paragraph>
<paragraph id="DD6736C4FFC589359FEBFEDFFEBCFE02" blockId="51.[189,1133,366,427]" box="[234,325,366,392]" pageId="51" pageNumber="52">
<emphasis id="EFACEAD6FFC589359FEBFEDFFEBCFE02" bold="true" box="[234,325,366,392]" pageId="51" pageNumber="52">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFC589359FBCFE20FB94FE21" blockId="51.[189,1133,366,427]" box="[189,1133,401,427]" pageId="51" pageNumber="52">Protibia short, broad with four stout teeth, the mid and hind femur without teeth.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFC589359FBCFE63FE3FFD64" pageId="51" pageNumber="52" type="distribution">
<paragraph id="DD6736C4FFC589359FBCFE63FEAFFE66" blockId="51.[189,342,466,492]" box="[189,342,466,492]" pageId="51" pageNumber="52">
<heading id="862F81A8FFC589359FBCFE63FEAFFE66" bold="true" box="[189,342,466,492]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="EFACEAD6FFC589359FBCFE63FEAFFE66" bold="true" box="[189,342,466,492]" pageId="51" pageNumber="52">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFC589359FBCFDB0FE3FFD64" blockId="51.[189,1399,513,750]" pageId="51" pageNumber="52">
<collectingCountry id="A5CF7654FFC589359FBCFDB0FF04FD91" box="[189,253,513,539]" name="India" pageId="51" pageNumber="52">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFC589359E09FDB0FE10FD91" box="[264,489,513,539]" country="India" name="Arunachal Pradesh" pageId="51" pageNumber="52">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359EF5FDB0FDBFFD91" box="[500,582,513,539]" country="India" name="Assam" pageId="51" pageNumber="52">Assam</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359D52FDB0FD6CFD91" box="[595,661,513,539]" country="India" name="Bihar" pageId="51" pageNumber="52">Bihar</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359DA2FDB0FCD5FD91" box="[675,812,513,539]" country="India" name="Chhattisgarh" pageId="51" pageNumber="52">Chattisgarh</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359C38FDB0FC6AFD91" box="[825,915,513,539]" country="India" name="Gujarat" pageId="51" pageNumber="52">Gujarat</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359CA1FDB0FBFCFD91" box="[928,1029,513,539]" country="India" name="Haryana" pageId="51" pageNumber="52">Haryana</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359B13FDB0FB12FD91" box="[1042,1259,513,539]" country="India" name="Himachal Pradesh" pageId="51" pageNumber="52">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359BF9FDB0FA8BFD91" box="[1272,1394,513,539]" country="India" name="Karnataka" pageId="51" pageNumber="52">Karnataka</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359FBCFD95FEF4FDB4" box="[189,269,548,574]" country="India" name="Kerala" pageId="51" pageNumber="52">Kerala</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359E1AFD95FE1AFDB4" box="[283,483,548,574]" country="India" name="Madhya Pradesh" pageId="51" pageNumber="52">Madhya Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359EF0FD95FD7CFDB4" box="[497,645,548,574]" country="India" name="Maharashtra" pageId="51" pageNumber="52">Maharashtra</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359D92FD95FCEEFDB4" box="[659,791,548,574]" country="India" name="Meghalaya" pageId="51" pageNumber="52">Meghalaya</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359C24FD95FC6EFDB4" box="[805,919,548,574]" country="India" name="Nagaland" pageId="51" pageNumber="52">Nagaland</collectingRegion>
, New
<collectingRegion id="1F1CF826FFC589359CE7FD95FBD0FDB4" box="[998,1065,548,574]" country="India" name="Delhi" pageId="51" pageNumber="52">Delhi</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359B39FD95FB74FDB4" box="[1080,1165,548,574]" country="India" name="Odisha" pageId="51" pageNumber="52">Odisha</collectingRegion>
, Punjab,
<collectingRegion id="1F1CF826FFC589359BFCFD95FA8AFDB4" box="[1277,1395,548,574]" country="India" name="Rajasthan" pageId="51" pageNumber="52">Rajasthan</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359FBCFDF6FEEDFDEB" box="[189,276,583,609]" country="India" name="Sikkim" pageId="51" pageNumber="52">Sikkim</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359E29FDF6FE44FDEB" box="[296,445,583,609]" country="India" name="Tamil Nadu" pageId="51" pageNumber="52">Tamil Nadu</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359ED1FDF6FDD0FDEB" box="[464,553,583,609]" country="India" name="Tripura" pageId="51" pageNumber="52">Tripura</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359D3FFDF6FD28FDEB" box="[574,721,583,609]" country="India" name="Uttarakhand" pageId="51" pageNumber="52">Uttarakhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359DE5FDF9FC76FDEB" box="[740,911,583,610]" country="India" name="Uttar Pradesh" pageId="51" pageNumber="52">Uttar Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359CA2FDF9FBBBFDEB" box="[931,1090,583,610]" country="India" name="West Bengal" pageId="51" pageNumber="52">West Bengal</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFC589359A2BFDF6FA8BFDEB" box="[1322,1394,583,609]" name="China" pageId="51" pageNumber="52">China</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359FBCFDDBFEBEFD0E" box="[189,327,618,644]" name="Bangladesh" pageId="51" pageNumber="52">Bangladesh</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359E54FDDAFE69FD0F" box="[341,400,619,645]" name="Laos" pageId="51" pageNumber="52">Laos</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359E9FFDDAFDE8FD0F" box="[414,529,619,645]" name="Myanmar" pageId="51" pageNumber="52">Myanmar</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359D1EFDDBFD9EFD0E" box="[543,615,618,644]" name="Nepal" pageId="51" pageNumber="52">Nepal</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359D74FDDBFD23FD0E" box="[629,730,618,644]" name="Pakistan" pageId="51" pageNumber="52">Pakistan</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359DE6FDDBFCA8FD0E" box="[743,849,618,644]" name="Thailand" pageId="51" pageNumber="52">Thailand</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359C5EFDDBFC38FD0E" box="[863,961,618,644]" name="Vietnam" pageId="51" pageNumber="52">Vietnam</collectingCountry>
(
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<bibRefCitation id="B9494B35FFC589359B8DFDDBFF03FD22" author="Chandra K. &amp; Ahirwar S. C." pageId="51" pageNumber="52" pagination="147 - 155" refId="ref28418" refString="Chandra K. &amp; Ahirwar S. C. 2005. Scarabaeid Beetles (Coleoptera) of Kanha Tiger Reserve, Madhya Pradesh. Records Zoological Survey of India 105 (1 - 2): 147 - 155." type="journal article" year="2005">Chandra &amp; Ahirwar 2005</bibRefCitation>
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<bibRefCitation id="B9494B35FFC589359E0AFD3FFEB0FD22" author="Chandra K. &amp; Ahirwar S. C." box="[267,329,654,680]" pageId="51" pageNumber="52" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">2007</bibRefCitation>
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<bibRefCitation id="B9494B35FFC589359E5BFD3CFD91FD2D" author="Bezdek A. &amp; Krell F. - T." box="[346,616,653,680]" pageId="51" pageNumber="52" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
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<bibRefCitation id="B9494B35FFC589359C2FFD3CFC83FD2D" author="Sewak R." box="[814,890,653,679]" pageId="51" pageNumber="52" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">2009b</bibRefCitation>
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<bibRefCitation id="B9494B35FFC589359C8AFD3CFBA2FD22" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." box="[907,1115,653,680]" pageId="51" pageNumber="52" pagination="29 - 38" refId="ref30906" refString="Sabu T. K., Nithya S. &amp; Vinod K. V. 2011. Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa 2830: 29 - 38. https: // doi. org / 10.11646 / ZOOTAXA. 2830.1.3" type="book chapter" year="2011">
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<bibRefCitation id="B9494B35FFC589359D26FD00FD0FFD46" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[551,758,689,716]" pageId="51" pageNumber="52" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
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).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,398 @@
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<taxonomicName id="1AD84D47FFDD892D9D90FEB9FC5AFEA9" ID-CoL="49KL8" authority="Gillet, 1911" authorityName="Gillet" authorityYear="1911" box="[657,931,264,291]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="siva">
<emphasis id="EFACEAD6FFDD892D9D90FEB9FCE9FEA8" bold="true" box="[657,784,264,290]" italics="true" pageId="43" pageNumber="44">Onitis siva</emphasis>
<bibRefCitation id="B9494B35FFDD892D9C19FEB9FC5AFEA9" author="Gillet J. J. E." box="[792,931,264,291]" pageId="43" pageNumber="44" pagination="313 - 314" refId="ref29108" refString="Gillet J. J. E. 1911. Coprides nouveaux de la region orientale (etc.). Annales de la Societe entomologique de Belgique LV: 313 - 314." type="book chapter" year="1911">Gillet, 1911</bibRefCitation>
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(
<figureCitation id="45E32A41FFDD892D9DA7FE9DFCD5FECC" box="[678,812,300,326]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="43" pageNumber="44">Figs 4KN</figureCitation>
,
<figureCitation id="45E32A41FFDD892D9C39FE9DFC92FECC" box="[824,875,300,326]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="43" pageNumber="44">12B</figureCitation>
,
<figureCitation id="45E32A41FFDD892D9C79FE9DFC6CFECC" box="[888,917,300,326]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." pageId="43" pageNumber="44">20</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFDD892D9FBCFEDFFCE8FE0D" blockId="43.[189,785,365,392]" box="[189,785,365,392]" pageId="43" pageNumber="44">
<emphasis id="EFACEAD6FFDD892D9FBCFEDFFEC2FE02" box="[189,315,366,392]" italics="true" pageId="43" pageNumber="44">Onitissiva</emphasis>
<bibRefCitation id="B9494B35FFDD892D9E42FEDCFDF0FE0D" author="Gillet J. J. E." box="[323,521,365,391]" pageId="43" pageNumber="44" pagination="313 - 314" refId="ref29108" refString="Gillet J. J. E. 1911. Coprides nouveaux de la region orientale (etc.). Annales de la Societe entomologique de Belgique LV: 313 - 314." type="book chapter" year="1911">Gillet, 1911: 313</bibRefCitation>
(original description).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDD892D9FBCFE1EFC85FDDF" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="DD6736C4FFDD892D9FBCFE1EFC85FDDF" blockId="43.[189,1399,430,598]" pageId="43" pageNumber="44">
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<emphasis id="EFACEAD6FFDD892D9FBCFE1EFEC1FE42" box="[189,312,431,457]" italics="true" pageId="43" pageNumber="44">Onitis siva</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFDD892D9E52FE1EFDE1FE43" author="Arrow G. J." box="[339,536,431,457]" page="388" pageId="43" pageNumber="44" year="1931">
<bibRefCitation id="B9494B35FFDD892D9E52FE1EFDE1FE43" author="Arrow G. J." box="[339,536,431,457]" pageId="43" pageNumber="44" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 388</bibRefCitation>
</treatmentCitation>
(key, description). —
<treatmentCitation id="5C7910D5FFDD892D9C18FE1EFC0FFE43" author="Janssens A." box="[793,1014,431,457]" page="109" pageId="43" pageNumber="44" year="1937">
<bibRefCitation id="B9494B35FFDD892D9C18FE1EFC0FFE43" author="Janssens A." box="[793,1014,431,457]" pageId="43" pageNumber="44" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 109</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFDD892D9B9FFE1FFA8FFE43" author="Balthasar V." box="[1182,1398,430,457]" page="53" pageId="43" pageNumber="44" year="1963">
<bibRefCitation id="B9494B35FFDD892D9B9FFE1FFA8FFE43" author="Balthasar V." box="[1182,1398,430,457]" pageId="43" pageNumber="44" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar 1963: 53</bibRefCitation>
</treatmentCitation>
(monograph).—
<treatmentCitation id="5C7910D5FFDD892D9EADFE63FD9AFE66" author="Rajan P. D." box="[428,611,466,492]" page="108" pageId="43" pageNumber="44" year="2006">
<bibRefCitation id="B9494B35FFDD892D9EADFE63FD9AFE66" author="Rajan P. D." box="[428,611,466,492]" pageId="43" pageNumber="44" pagination="91 - 135" refId="ref30757" refString="Rajan P. D. 2006. Insecta: Coleoptera: Scarabaeoidea: Scarabaeidae (dung beetles). Zoological Survey of India. Fauna of Bilgiri Rangaswamy Tiger Wildlife Sanctuary, Conservation Area Series 27: 91 - 135." type="book chapter" year="2006">Rajan 2006:108</bibRefCitation>
</treatmentCitation>
(list,key, diagnosis, distribution).—
<treatmentCitation id="5C7910D5FFDD892D9B00FE63FB2CFE66" author="Chandra K." box="[1025,1237,466,492]" page="81" pageId="43" pageNumber="44" year="2009">
<bibRefCitation id="B9494B35FFDD892D9B00FE63FB2CFE66" author="Chandra K." box="[1025,1237,466,492]" pageId="43" pageNumber="44" pagination="79 - 89" refId="ref28355" refString="Chandra K. 2009 a. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Tamil Nadu, State Fauna Series 1: 79 - 89." type="book chapter" year="2009">Chandra2009a:81</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDD892D9E11FE44FE23FD9A" author="Sewak R." box="[272,474,501,528]" page="35" pageId="43" pageNumber="44" year="2009">
<bibRefCitation id="B9494B35FFDD892D9E11FE44FE23FD9A" author="Sewak R." box="[272,474,501,528]" pageId="43" pageNumber="44" pagination="1 - 48" refId="ref31194" refString="Sewak R. 2009 a. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar desert of Gujarat. Records of the Zoological Survey of India, Occassional Paper No 295: 1 - 48." type="journal article" year="2009">Sewak 2009a: 35</bibRefCitation>
</treatmentCitation>
(key, diagnosis, distribution); 2009b: 58 (key, diagnosis, distribution). —
<treatmentCitation id="5C7910D5FFDD892D9A3CFE44FE6EFDB8" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." page="34" pageId="43" pageNumber="44" year="2011">
<bibRefCitation id="B9494B35FFDD892D9A3CFE44FE6EFDB8" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." pageId="43" pageNumber="44" pagination="29 - 38" refId="ref30906" refString="Sabu T. K., Nithya S. &amp; Vinod K. V. 2011. Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa 2830: 29 - 38. https: // doi. org / 10.11646 / ZOOTAXA. 2830.1.3" type="book chapter" year="2011">
Sabu
<emphasis id="EFACEAD6FFDD892D9FEBFDA8FEDFFDB8" box="[234,294,536,562]" italics="true" pageId="43" pageNumber="44">et al.</emphasis>
2011: 34
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFDD892D9D0CFDA9FC51FDB9" author="Karimbumkara S. N. &amp; Rajan P. D." box="[525,936,536,563]" page="175" pageId="43" pageNumber="44" year="2013">
<bibRefCitation id="B9494B35FFDD892D9D0CFDA9FC51FDB9" author="Karimbumkara S. N. &amp; Rajan P. D." box="[525,936,536,563]" pageId="43" pageNumber="44" pagination="173 - 178" refId="ref29842" refString="Karimbumkara S. N. &amp; Rajan P. D. 2013. Insecta: Coleoptera: Scarabaeidae: Scarabaeinae, Dung Beetles. Zoological Survey of India, Fauna of Karnataka, State Fauna Series 21: 173 - 178." type="book chapter" year="2013">Karimbumkara &amp; Rajan 2013: 175</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFDD892D9B1EFDA9FAC3FDB8" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[1055,1338,536,563]" page="1037" pageId="43" pageNumber="44" year="2015">
<bibRefCitation id="B9494B35FFDD892D9B1EFDA9FAC3FDB8" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[1055,1338,536,563]" pageId="43" pageNumber="44" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFDD892D9B6EFDA8FB52FDB8" box="[1135,1195,536,562]" italics="true" pageId="43" pageNumber="44">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(list, distribution). —
<bibRefCitation id="B9494B35FFDD892D9EABFD8AFD66FDDF" author="Schoolmeesters P." box="[426,671,571,597]" pageId="43" pageNumber="44" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDD892D9FBCFDCCFC2CFC86" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="DD6736C4FFDD892D9FBCFDCCFE3FFD1D" blockId="43.[189,454,637,663]" box="[189,454,637,663]" pageId="43" pageNumber="44">
<heading id="862F81A8FFDD892D9FBCFDCCFE3FFD1D" bold="true" box="[189,454,637,663]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="EFACEAD6FFDD892D9FBCFDCCFE3FFD1D" bold="true" box="[189,454,637,663]" pageId="43" pageNumber="44">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDD892D9FBCFD1DFC2CFC86" blockId="43.[189,1399,683,780]" pageId="43" pageNumber="44">
<emphasis id="EFACEAD6FFDD892D9FBCFD1DFEC0FD4C" box="[189,313,684,710]" italics="true" pageId="43" pageNumber="44">Onitissiva</emphasis>
belongs to Group XVIII. It is closer to
<taxonomicName id="1AD84D47FFDD892D9DF6FD1DFC97FD4F" authorityName="Cambefort" authorityYear="1988" box="[759,878,683,710]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="bordati">
<emphasis id="EFACEAD6FFDD892D9DF6FD1DFC97FD4F" box="[759,878,683,710]" italics="true" pageId="43" pageNumber="44">O. bordati</emphasis>
</taxonomicName>
which is the nearest group member in having an uninterrupted frontal carina and the absence of clypeal carina.
<taxonomicName id="1AD84D47FFDD892D9CB6FD7EFBCDFD62" authorityName="Gillet" authorityYear="1911" box="[951,1076,719,745]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="siva">
<emphasis id="EFACEAD6FFDD892D9CB6FD7EFBCDFD62" box="[951,1076,719,745]" italics="true" pageId="43" pageNumber="44">Onitis siva</emphasis>
</taxonomicName>
is easily distinguishable by the strongly punctured pronotum whereas in the latter it is granular.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDD892D9FBCFC82FDCAFA2F" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="DD6736C4FFDD892D9FBCFC82FE15FCC7" blockId="43.[189,492,819,845]" box="[189,492,819,845]" pageId="43" pageNumber="44">
<heading id="862F81A8FFDD892D9FBCFC82FE15FCC7" bold="true" box="[189,492,819,845]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="EFACEAD6FFDD892D9FBCFC82FE15FCC7" bold="true" box="[189,492,819,845]" pageId="43" pageNumber="44">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDD892D9FEBFCD3FEA5FCF6" blockId="43.[189,1399,866,998]" box="[234,348,866,892]" pageId="43" pageNumber="44">
<emphasis id="EFACEAD6FFDD892D9FEBFCD3FEA5FCF6" bold="true" box="[234,348,866,892]" pageId="43" pageNumber="44">
<typeStatus id="02638866FFDD892D9FEBFCD3FEA5FCF6" box="[234,348,866,892]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDD892D9FBCFC34FE67FC6F" blockId="43.[189,1399,866,998]" pageId="43" pageNumber="44">
<materialsCitation id="6DB03C99FFDD892D9FBCFC34FE67FC6F" collectingDate="1903-09-05" collectionCode="IRSNB" collectorName="J. J. Gillet &amp; R. D. Favre" country="India" location="INDIA" pageId="43" pageNumber="44" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingCountry id="A5CF7654FFDD892D9FBCFC34FEF7FC15" box="[189,270,901,927]" name="India" pageId="43" pageNumber="44">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFDD892D9E25FC34FEC2FC15" box="[292,315,901,927]" count="1" pageId="43" pageNumber="44" type="female"></specimenCount>
; “Travancore; A Janssens vid., 1936:
<emphasis id="EFACEAD6FFDD892D9DEAFC34FC9EFC15" box="[747,871,901,927]" italics="true" pageId="43" pageNumber="44">Onitissiva</emphasis>
Gillet,
<collectorName id="702D5312FFDD892D9CBDFC34FBD3FC15" box="[956,1066,901,927]" pageId="43" pageNumber="44">J.J. Gillet</collectorName>
set., vend.
<emphasis id="EFACEAD6FFDD892D9BABFC34FACDFC15" box="[1194,1332,901,927]" italics="true" pageId="43" pageNumber="44">Onitissiva,</emphasis>
Gillet RMHN. Belg. 10.640, cf. Ann. Soc. Ent. Belg. LV. 1911, 313;
<typeStatus id="02638866FFDD892D9C9BFC18FC2EFC49" box="[922,983,937,963]" pageId="43" pageNumber="44">Type</typeStatus>
, Wallardi (Travancore)
<collectorName id="702D5312FFDD892D9BECFC18FA8BFC49" box="[1261,1394,937,963]" pageId="43" pageNumber="44">R.D. Favre</collectorName>
,
<date id="A9661004FFDD892D9FBCFC7AFED3FC6C" box="[189,298,971,998]" pageId="43" pageNumber="44" value="1903-09-05">
<collectingDate id="B922E9ECFFDD892D9FBCFC7AFED3FC6C" box="[189,298,971,998]" pageId="43" pageNumber="44" value="1903-09-05">5.9.1903</collectingDate>
</date>
”;
<collectionCode id="BBC9AE01FFDD892D9E41FC7AFE63FC6F" box="[320,410,971,997]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="43" pageNumber="44">IRSNB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFDD892D9FEBFBBCFCE2FBAD" blockId="43.[189,1398,1037,1345]" box="[234,795,1037,1063]" pageId="43" pageNumber="44">
<emphasis id="EFACEAD6FFDD892D9FEBFBBCFD98FBAD" bold="true" box="[234,609,1037,1063]" pageId="43" pageNumber="44">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFDD892D9D70FBBCFCEAFBAD" box="[625,787,1037,1063]" count="10" pageId="43" pageNumber="44" type="generic">10 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFDD892D9FBCFB81FEB5FACB" blockId="43.[189,1398,1037,1345]" pageId="43" pageNumber="44">
<materialsCitation id="6DB03C99FFDD892D9FBCFB81FAB8FBC1" box="[189,1345,1072,1099]" collectionCode="BMNH" country="India" elevation="914" location="INDIA" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFDD892D9FBCFB81FEE9FBC0" box="[189,272,1072,1098]" name="India" pageId="43" pageNumber="44">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFDD892D9E27FB80FEC6FBC1" box="[294,319,1073,1099]" count="1" pageId="43" pageNumber="44" type="male"></specimenCount>
; “Nilgiri Hills, H.L. Andrewes.,
<quantity id="1A209B21FFDD892D9DBFFB81FCE5FBC0" box="[702,796,1072,1098]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="43" pageNumber="44" unit="ft" value="3000.0">
<elevation id="56F5D1F7FFDD892D9DBFFB81FCE5FBC0" box="[702,796,1072,1098]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="43" pageNumber="44" unit="ft" value="3000.0">3000 ft.</elevation>
</quantity>
,
<taxonomicName id="1AD84D47FFDD892D9C26FB81FC5FFBC0" authorityName="Gillet" authorityYear="1911" box="[807,934,1072,1098]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="siva">
<emphasis id="EFACEAD6FFDD892D9C26FB81FC5FFBC0" box="[807,934,1072,1098]" italics="true" pageId="43" pageNumber="44">Onitis siva</emphasis>
</taxonomicName>
male, Gillet”;
<collectionCode id="BBC9AE01FFDD892D9B55FB81FB48FBC0" box="[1108,1201,1072,1098]" pageId="43" pageNumber="44">BMNH</collectionCode>
(E) 1237159
</materialsCitation>
<materialsCitation id="6DB03C99FFDD892D9A59FB80FA8FFB1B" collectingDate="1999-05-14" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" location="Budipadaga" longitude="77.07755" municipality="Biligiri Rangaswamy Tiger Reserve" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFDD892D9A59FB80FA88FBC1" box="[1368,1393,1073,1099]" count="1" pageId="43" pageNumber="44" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFDD892D9FBCFBE2FEC2FBE7" box="[189,315,1107,1133]" country="India" name="Karnataka" pageId="43" pageNumber="44">Karnataka</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDD892D9E47FBE2FD0FFBE4" box="[326,758,1107,1134]" pageId="43" pageNumber="44">Biligiri Rangaswamy Tiger Reserve</collectingMunicipality>
,
<location id="D807601FFFDD892D9C00FBE2FC69FBE7" LSID="urn:lsid:plazi:treatment:557187D2FFDD892A9D90FEB9FDACFD95:D807601FFFDD892D9C00FBE2FC69FBE7" box="[769,912,1107,1133]" country="India" longitude="77.07755" municipality="Biligiri Rangaswamy Tiger Reserve" name="Budipadaga" pageId="43" pageNumber="44" stateProvince="Karnataka">Budipadaga</location>
scrub; N 11°50ʹ16.8″,
<geoCoordinate id="B8EC5003FFDD892D9BA0FBE3FAB5FBE4" box="[1185,1356,1106,1134]" degrees="77" direction="east" minutes="4" orientation="longitude" pageId="43" pageNumber="44" precision="1" seconds="39.18" value="77.07755">77°4ʹ39.18″ E</geoCoordinate>
;
<date id="A9661004FFDD892D9A58FBE2FEC7FB1B" pageId="43" pageNumber="44" value="1999-05-14">
<collectingDate id="B922E9ECFFDD892D9A58FBE2FEC7FB1B" pageId="43" pageNumber="44" value="1999-05-14">14 May 1999</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFDD892D9E4FFBC6FD57FB1B" box="[334,686,1142,1169]" pageId="43" pageNumber="44">open cattle dung baited trap</collectingMethod>
;
<collectorName id="702D5312FFDD892D9DBFFBC7FBDCFB1A" box="[702,1061,1142,1168]" pageId="43" pageNumber="44">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDD892D9B6EFBC6FB44FB1B" box="[1135,1213,1143,1169]" pageId="43" pageNumber="44">AIMB</collectionCode>
/
<collectorName id="702D5312FFDD892D9BC7FBC6FB11FB1B" box="[1222,1256,1143,1169]" pageId="43" pageNumber="44">Co</collectorName>
/
<collectorName id="702D5312FFDD892D9BF3FBC6FAF7FB1B" box="[1266,1294,1143,1169]" pageId="43" pageNumber="44">Sc</collectorName>
1000703
</materialsCitation>
<materialsCitation id="6DB03C99FFDD892D9FCEFB2BFE3CFB5D" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" latitude="12.0205" location="Nagarhole National Park" longLatPrecision="1" longitude="76.104095" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFDD892D9FCEFB2BFF11FB3E" box="[207,232,1178,1204]" count="1" pageId="43" pageNumber="44" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFDD892D9FF7FB2BFE8DFB3E" box="[246,372,1178,1204]" country="India" name="Karnataka" pageId="43" pageNumber="44">Karnataka</collectingRegion>
,
<location id="D807601FFFDD892D9E80FB2BFD57FB3E" LSID="urn:lsid:plazi:treatment:557187D2FFDD892A9D90FEB9FDACFD95:D807601FFFDD892D9E80FB2BFD57FB3E" box="[385,686,1178,1204]" country="India" latitude="12.0205" longLatPrecision="1" longitude="76.104095" name="Nagarhole National Park" pageId="43" pageNumber="44" stateProvince="Karnataka">Nagarhole National Park</location>
, core moist deciduous forest;
<geoCoordinate id="B8EC5003FFDD892D9B16FB29FB41FB3E" box="[1047,1208,1176,1204]" degrees="12" direction="north" minutes="1" orientation="latitude" pageId="43" pageNumber="44" precision="1" seconds="13.8" value="12.0205">12°1ʹ13.8″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDD892D9BC4FB29FA88FB3E" box="[1221,1393,1176,1204]" degrees="76" direction="east" minutes="6" orientation="longitude" pageId="43" pageNumber="44" precision="1" seconds="14.76" value="76.104095">76°6ʹ14.76″ E</geoCoordinate>
;
<collectionCode id="BBC9AE01FFDD892D9FBCFB0CFEF2FB5D" box="[189,267,1213,1239]" pageId="43" pageNumber="44">AIMB</collectionCode>
/
<collectorName id="702D5312FFDD892D9E15FB0CFECFFB5D" box="[276,310,1213,1239]" pageId="43" pageNumber="44">Co</collectorName>
/
<collectorName id="702D5312FFDD892D9E41FB0CFEA5FB5D" box="[320,348,1213,1239]" pageId="43" pageNumber="44">Sc</collectorName>
1000704
</materialsCitation>
<materialsCitation id="6DB03C99FFDD892D9EDFFB0CFDA4FB70" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" latitude="12.1563" location="Hebbala" longLatPrecision="1" longitude="76.114395" pageId="43" pageNumber="44" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCount id="CBDEFD4DFFDD892D9EDFFB0CFDDAFB5D" box="[478,547,1213,1239]" count="2" pageId="43" pageNumber="44" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDD892D9D2EFB0CFD8AFB5D" box="[559,627,1213,1239]" count="2" pageId="43" pageNumber="44" type="female">2 ♀♀</specimenCount>
;
<location id="D807601FFFDD892D9D80FB0CFD1DFB5D" LSID="urn:lsid:plazi:treatment:557187D2FFDD892A9D90FEB9FDACFD95:D807601FFFDD892D9D80FB0CFD1DFB5D" box="[641,740,1213,1239]" country="India" latitude="12.1563" longLatPrecision="1" longitude="76.114395" name="Hebbala" pageId="43" pageNumber="44">Hebbala</location>
moist deciduous forest;
<geoCoordinate id="B8EC5003FFDD892D9B0BFB0AFB40FB5D" box="[1034,1209,1211,1239]" degrees="12" direction="north" minutes="9" orientation="latitude" pageId="43" pageNumber="44" precision="1" seconds="22.68" value="12.1563">12°9ʹ22.68″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDD892D9BC4FB0AFA88FB5D" box="[1221,1393,1211,1239]" degrees="76" direction="east" minutes="6" orientation="longitude" pageId="43" pageNumber="44" precision="1" seconds="51.84" value="76.114395">76°6ʹ51.84″ E</geoCoordinate>
;
<collectionCode id="BBC9AE01FFDD892D9FBCFB51FEF2FB71" box="[189,267,1248,1275]" pageId="43" pageNumber="44">AIMB</collectionCode>
/
<collectorName id="702D5312FFDD892D9E15FB51FECFFB71" box="[276,310,1248,1275]" pageId="43" pageNumber="44">Co</collectorName>
/
<collectorName id="702D5312FFDD892D9E41FB51FEA4FB71" box="[320,349,1248,1275]" pageId="43" pageNumber="44">Sc</collectorName>
1000705 to 1000708
</materialsCitation>
<materialsCitation id="6DB03C99FFDD892D9D83FB50FEB1FACB" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="11.974199" location="Sungathakatte Dry Deciduous forest" longLatPrecision="1" longitude="76.1856" pageId="43" pageNumber="44" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFDD892D9D83FB50FD63FB71" box="[642,666,1249,1275]" count="1" pageId="43" pageNumber="44" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFDD892D9DADFB50FD3AFB71" box="[684,707,1249,1275]" count="1" pageId="43" pageNumber="44" type="female"></specimenCount>
;
<location id="D807601FFFDD892D9DD9FB51FB67FB70" LSID="urn:lsid:plazi:treatment:557187D2FFDD892A9D90FEB9FDACFD95:D807601FFFDD892D9DD9FB51FB67FB70" box="[728,1182,1248,1275]" country="India" latitude="11.974199" longLatPrecision="1" longitude="76.1856" name="Sungathakatte Dry Deciduous forest" pageId="43" pageNumber="44">Sungathakatte Dry Deciduous forest</location>
;
<geoCoordinate id="B8EC5003FFDD892D9BAEFB6FFA88FB71" box="[1199,1393,1246,1275]" degrees="11" direction="north" minutes="58" orientation="latitude" pageId="43" pageNumber="44" precision="1" seconds="27.12" value="11.974199">11°58ʹ27.12″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDD892D9FBCFAB3FE98FA94" box="[189,353,1282,1310]" degrees="76" direction="east" minutes="11" orientation="longitude" pageId="43" pageNumber="44" precision="1" seconds="8.16" value="76.1856">76°11ʹ8.16″E</geoCoordinate>
;
<collectingMethod id="04994ED3FFDD892D9E6FFAB5FD46FA94" box="[366,703,1283,1310]" pageId="43" pageNumber="44">open cattle dung baited trap</collectingMethod>
;
<collectorName id="702D5312FFDD892D9DCDFAB2FBD5FA97" box="[716,1068,1283,1309]" pageId="43" pageNumber="44">Priyadarsanan Dharma Rajan</collectorName>
leg.;
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/
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/
<collectorName id="702D5312FFDD892D9BF0FAB2FAF4FA97" box="[1265,1293,1283,1309]" pageId="43" pageNumber="44">Sc</collectorName>
1000709 to 1000710
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.
</paragraph>
<paragraph id="DD6736C4FFDD892D9FBCFAD9FDCAFA2F" blockId="43.[189,1399,1384,1445]" pageId="43" pageNumber="44">
<materialsCitation id="6DB03C99FFDD892D9FBCFAD9FDD6FA2F" collectingDate="1983-10-24" collectionCode="BMNH" collectorName="K. K. Gurung" country="Nepal" elevation="150" location="Chitwan" pageId="43" pageNumber="44" specimenCount="151" specimenCount-female="1">
<collectingCountry id="A5CF7654FFDD892D9FBCFAD9FEE1FA08" box="[189,280,1384,1410]" name="Nepal" pageId="43" pageNumber="44">NEPAL</collectingCountry>
<specimenCount id="CBDEFD4DFFDD892D9E35FAD9FEB2FA08" box="[308,331,1384,1410]" count="1" pageId="43" pageNumber="44" type="female"></specimenCount>
; “
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, Tiger Tops.
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<quantity id="1A209B21FFDD892D9D6DFAD9FD4CFA09" box="[620,693,1384,1411]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="44" unit="m" value="150.0">
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</quantity>
.
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;
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<collectingDate id="B922E9ECFFDD892D9DC6FAD9FC9EFA08" box="[711,871,1384,1410]" pageId="43" pageNumber="44" value="1983-10-24">24 Oct. 1983</collectingDate>
</date>
;
<collectorName id="702D5312FFDD892D9C74FAD9FBEDFA08" box="[885,1044,1384,1410]" pageId="43" pageNumber="44">K.K. Gurung</collectorName>
; Rhinoceros dung; Brit. Mus. 1984. 19”;
<collectionCode id="BBC9AE01FFDD892D9E41FA3AFE62FA2F" box="[320,411,1419,1445]" pageId="43" pageNumber="44">BMNH</collectionCode>
(E) 1237160
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDD892A9FBCFA7DFB3FFEE3" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
<paragraph id="DD6736C4FFDD892D9FBCFA7DFEB7FA6C" blockId="43.[189,334,1484,1510]" box="[189,334,1484,1510]" pageId="43" pageNumber="44">
<heading id="862F81A8FFDD892D9FBCFA7DFEB7FA6C" bold="true" box="[189,334,1484,1510]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="EFACEAD6FFDD892D9FBCFA7DFEB7FA6C" bold="true" box="[189,334,1484,1510]" pageId="43" pageNumber="44">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDD892D9FBCFA4AFCAFF99C" blockId="43.[189,854,1531,1558]" box="[189,854,1531,1558]" pageId="43" pageNumber="44">MEASUREMENTS (in mm). TL=22.531.5; BW=12.516.</paragraph>
<paragraph id="DD6736C4FFDD892D9FEBF98CFED3F9DD" blockId="43.[189,1399,1597,1869]" box="[234,298,1597,1623]" pageId="43" pageNumber="44">
<emphasis id="EFACEAD6FFDD892D9FEBF98CFED3F9DD" bold="true" box="[234,298,1597,1623]" pageId="43" pageNumber="44">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDD892D9FBCF9D1FA8EF8C7" blockId="43.[189,1399,1597,1869]" pageId="43" pageNumber="44">Black or deep reddish-black, not very shining, elongate-oval, moderately convex; clypeus almost semicircular, finely and densely rugose, clypeal carina absent; frontal carina strongly elevated and curved; pronotum with a trace of longitudinal median groove posteriorly, strongly and unevenly punctured, punctures confluent anteriorly, minutely granular near lateral margins, basal foveae elongate and close; pygidium flat, indistinctly punctate; metasternum densely hairy at sides and less densely in middle, base with a small smooth but moderately punctured area. Pronotal punctures not strong and even; front legs elongate, femur with very strong spine in the middle of anterior edge, mid and hind femur also toothed.</paragraph>
<paragraph id="DD6736C4FFDD892D9FBCF8C5FBA6F83B" blockId="43.[189,1398,1908,1970]" pageId="43" pageNumber="44">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFDD892D9EB8F8C5FDB9F804" box="[441,576,1908,1934]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="43" pageNumber="44">Fig. 4KN</figureCitation>
). LP=4.5, Lp= 3.5; BP=2, BpB=1, BpT=0.5. Phallobase distinctly longer than parameres. Parameres curved inwards, rounded and blunt at the tip.
</paragraph>
<paragraph id="DD6736C4FFDA892A9FEBFEB8FEBCFEA9" blockId="44.[189,1398,265,361]" box="[234,325,265,291]" pageId="44" pageNumber="45">
<emphasis id="EFACEAD6FFDA892A9FEBFEB8FEBCFEA9" bold="true" box="[234,325,265,291]" pageId="44" pageNumber="45">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FBCFE9DFB3FFEE3" blockId="44.[189,1398,265,361]" pageId="44" pageNumber="45">Pronotal punctures stronger and closer than in males; median elevation of posterior marginal carina of the head broader and stronger than in male, protibia straight, front femora without teeth.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDA892A9FBCFE21FDACFD95" pageId="44" pageNumber="45" type="distribution">
<paragraph id="DD6736C4FFDA892A9FBCFE21FEAFFE20" blockId="44.[189,342,400,426]" box="[189,342,400,426]" pageId="44" pageNumber="45">
<heading id="862F81A8FFDA892A9FBCFE21FEAFFE20" bold="true" box="[189,342,400,426]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="EFACEAD6FFDA892A9FBCFE21FEAFFE20" bold="true" box="[189,342,400,426]" pageId="44" pageNumber="45">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FBCFE0EFDACFD95" blockId="44.[189,1399,447,544]" pageId="44" pageNumber="45">
<collectingCountry id="A5CF7654FFDA892A9FBCFE0EFF07FE53" box="[189,254,447,473]" name="India" pageId="44" pageNumber="45">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFDA892A9E0AFE0EFE09FE53" box="[267,496,447,473]" country="India" name="Arunachal Pradesh" pageId="44" pageNumber="45">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFDA892A9EFEFE0EFD82FE53" box="[511,635,447,473]" country="India" name="Karnataka" pageId="44" pageNumber="45">Karnataka</collectingRegion>
,
<collectingRegion id="1F1CF826FFDA892A9D88FE0EFD23FE53" box="[649,730,447,473]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
,
<collectingRegion id="1F1CF826FFDA892A9DEBFE0EFCBCFE53" box="[746,837,447,473]" country="India" name="Gujarat" pageId="44" pageNumber="45">Gujarat</collectingRegion>
,
<collectingRegion id="1F1CF826FFDA892A9C55FE0EFC32FE53" box="[852,971,447,473]" country="India" name="Rajasthan" pageId="44" pageNumber="45">Rajasthan</collectingRegion>
,
<collectingRegion id="1F1CF826FFDA892A9CD8FE0EFB92FE53" box="[985,1131,447,473]" country="India" name="Tamil Nadu" pageId="44" pageNumber="45">Tamil Nadu</collectingRegion>
,
<collectingRegion id="1F1CF826FFDA892A9B78FE0EFAE7FE53" box="[1145,1310,447,473]" country="India" name="Uttar Pradesh" pageId="44" pageNumber="45">Uttar Pradesh</collectingRegion>
(
<bibRefCitation id="B9494B35FFDA892A9A31FE0EFF02FE76" author="Rajan P. D." pageId="44" pageNumber="45" pagination="91 - 135" refId="ref30757" refString="Rajan P. D. 2006. Insecta: Coleoptera: Scarabaeoidea: Scarabaeidae (dung beetles). Zoological Survey of India. Fauna of Bilgiri Rangaswamy Tiger Wildlife Sanctuary, Conservation Area Series 27: 91 - 135." type="book chapter" year="2006">Rajan 2006</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDA892A9E0BFE53FE3AFE76" author="Chandra K." box="[266,451,482,508]" pageId="44" pageNumber="45" pagination="79 - 89" refId="ref28355" refString="Chandra K. 2009 a. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Tamil Nadu, State Fauna Series 1: 79 - 89." type="book chapter" year="2009">Chandra 2009a</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDA892A9ED0FE53FD8DFE76" author="Sewak R." box="[465,628,482,508]" pageId="44" pageNumber="45" pagination="1 - 48" refId="ref31194" refString="Sewak R. 2009 a. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar desert of Gujarat. Records of the Zoological Survey of India, Occassional Paper No 295: 1 - 48." type="journal article" year="2009">Sewak 2009a</bibRefCitation>
,
<bibRefCitation id="B9494B35FFDA892A9D82FE53FD29FE76" author="Sewak R." box="[643,720,482,508]" pageId="44" pageNumber="45" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">2009b</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDA892A9DDFFE53FC51FE76" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." box="[734,936,482,508]" pageId="44" pageNumber="45" pagination="29 - 38" refId="ref30906" refString="Sabu T. K., Nithya S. &amp; Vinod K. V. 2011. Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa 2830: 29 - 38. https: // doi. org / 10.11646 / ZOOTAXA. 2830.1.3" type="book chapter" year="2011">
Sabu
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2011
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDA892A9CB6FE53FAD9FE76" author="Karimbumkara S. N. &amp; Rajan P. D." box="[951,1312,482,508]" pageId="44" pageNumber="45" pagination="173 - 178" refId="ref29842" refString="Karimbumkara S. N. &amp; Rajan P. D. 2013. Insecta: Coleoptera: Scarabaeidae: Scarabaeinae, Dung Beetles. Zoological Survey of India, Fauna of Karnataka, State Fauna Series 21: 173 - 178." type="book chapter" year="2013">Karimbumkara &amp; Rajan 2013</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDA892A9A2EFE53FEC4FDAA" author="Gupta D. &amp; Chandra K. &amp; Khan S." pageId="44" pageNumber="45" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
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).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,658 @@
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<heading id="862F81A8FFDF892F9D40FB67FC0AFB7B" box="[577,1011,1238,1265]" centered="true" fontSize="11" level="2" pageId="41" pageNumber="42" reason="2">
<taxonomicName id="1AD84D47FFDF892F9D40FB67FC0AFB7B" ID-CoL="49KL7" authority="Lansberge, 1875" authorityName="Lansberge, AIMB" authorityYear="1875" box="[577,1011,1238,1265]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFDF892F9D40FB67FCD1FB7B" bold="true" box="[577,808,1238,1265]" italics="true" pageId="41" pageNumber="42">Onitis singhalensis</emphasis>
<bibRefCitation id="B9494B35FFDF892F9C2EFB67FC0AFB7B" author="Lansberge G." box="[815,1011,1238,1265]" pageId="41" pageNumber="42" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
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<subSubSection id="95C2654FFFDF892F9D3CFB4BFC52FADF" pageId="41" pageNumber="42" type="description">
<paragraph id="DD6736C4FFDF892F9D3CFB4BFC0FFA9E" blockId="41.[573,1014,1238,1301]" box="[573,1014,1274,1301]" pageId="41" pageNumber="42">
<figureCitation id="45E32A41FFDF892F9D3CFB4BFD4AFA9E" box="[573,691,1274,1300]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="41" pageNumber="42">Figs 8IL</figureCitation>
,
<figureCitation id="45E32A41FFDF892F9DBEFB4BFD27FA9E" box="[703,734,1274,1300]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="41" pageNumber="42">9F</figureCitation>
,
<figureCitation id="45E32A41FFDF892F9DEBFB4BFCE6FA9E" box="[746,799,1274,1300]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="41" pageNumber="42">12A</figureCitation>
,
<figureCitation id="45E32A41FFDF892F9C2AFB4BFCA3FA9E" box="[811,858,1274,1300]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="41" pageNumber="42">14F</figureCitation>
,
<figureCitation id="45E32A41FFDF892F9C64FB4BFC6AFA9F" box="[869,915,1274,1301]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="41" pageNumber="42">15F</figureCitation>
,
<figureCitation id="45E32A41FFDF892F9C9EFB4BFC34FA9E" box="[927,973,1274,1300]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="41" pageNumber="42">16F</figureCitation>
,
<figureCitation id="45E32A41FFDF892F9CD8FB4BFC0FFA9E" box="[985,1014,1274,1300]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." pageId="41" pageNumber="42">19</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFA8AFC52FADF" blockId="41.[189,939,1339,1366]" box="[189,939,1339,1366]" pageId="41" pageNumber="42">
<emphasis id="EFACEAD6FFDF892F9FBCFA8AFE64FADF" box="[189,413,1339,1365]" italics="true" pageId="41" pageNumber="42">Onitissinghalensis</emphasis>
<bibRefCitation id="B9494B35FFDF892F9EA5FA8AFD5AFADF" author="Lansberge G." box="[420,675,1339,1366]" pageId="41" pageNumber="42" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875: 140</bibRefCitation>
(original description).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDF892F9FBCFACCFAF2F9A9" pageId="41" pageNumber="42" type="reference_group">
<paragraph id="DD6736C4FFDF892F9FBCFACCFAF2F9A9" blockId="41.[189,1399,1404,1572]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="FDC811EAFFDF892F9FBCFACCFAF2F9A9" pageId="41" pageNumber="42">
<taxonomicName id="1AD84D47FFDF892F9FBCFACCFE64FA1D" ID-CoL="49KL7" authorityName="Lansberge, AIMB" authorityYear="1875" box="[189,413,1405,1431]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFDF892F9FBCFACCFE64FA1D" box="[189,413,1405,1431]" italics="true" pageId="41" pageNumber="42">Onitis singhalensis</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFDF892F9EB9FACCFD7BFA1D" author="Arrow G. J." box="[440,642,1405,1431]" page="394" pageId="41" pageNumber="42" year="1931">
<bibRefCitation id="B9494B35FFDF892F9EB9FACCFD7BFA1D" author="Arrow G. J." box="[440,642,1405,1431]" pageId="41" pageNumber="42" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 394</bibRefCitation>
</treatmentCitation>
(key, description). —
<treatmentCitation id="5C7910D5FFDF892F9C89FACCFBA2FA1D" author="Janssens A." box="[904,1115,1404,1431]" page="53" pageId="41" pageNumber="42" year="1937">
<bibRefCitation id="B9494B35FFDF892F9C89FACCFBA2FA1D" author="Janssens A." box="[904,1115,1404,1431]" pageId="41" pageNumber="42" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 53</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFDF892F9A09FACCFEACFA30" author="Balthasar V." page="40" pageId="41" pageNumber="42" year="1963">
<bibRefCitation id="B9494B35FFDF892F9A09FACCFEACFA30" author="Balthasar V." pageId="41" pageNumber="42" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar 1963: 40</bibRefCitation>
</treatmentCitation>
(monograph). —
<treatmentCitation id="5C7910D5FFDF892F9D32FA11FC48FA30" author="Biswas S. &amp; Chatterjee S. K." box="[563,945,1440,1466]" page="94" pageId="41" pageNumber="42" year="1986">
<bibRefCitation id="B9494B35FFDF892F9D32FA11FC48FA30" author="Biswas S. &amp; Chatterjee S. K." box="[563,945,1440,1466]" pageId="41" pageNumber="42" pagination="79 - 96" refId="ref27658" refString="Biswas S. &amp; Chatterjee S. K. 1986 b. Scarabaeidae (India Coleoptera) of Silent Valley, Kerala, India with descriptions of three new species. Records of the Zoological Survey of India 84 (1 - 4): 79 - 96. Available from: https: // faunaofindia. nic. in / PDFVolumes / records / 084 / 01 - 04 / 0079 - 0096. pdf [accessed 15 Oct. 2018]" type="book chapter" year="1986">Biswas &amp; Chatterjee 1986b: 94</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDF892F9B91FA11FA8FFA30" author="Chandra K." box="[1168,1398,1440,1466]" page="81" pageId="41" pageNumber="42" year="2009">
<bibRefCitation id="B9494B35FFDF892F9B91FA11FA8FFA30" author="Chandra K." box="[1168,1398,1440,1466]" pageId="41" pageNumber="42" pagination="79 - 89" refId="ref28355" refString="Chandra K. 2009 a. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Tamil Nadu, State Fauna Series 1: 79 - 89." type="book chapter" year="2009">Chandra 2009a: 81</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDF892F9EC1FA72FD6DFA54" author="Sewak R." box="[448,660,1475,1502]" page="59" pageId="41" pageNumber="42" year="2009">
<bibRefCitation id="B9494B35FFDF892F9EC1FA72FD6DFA54" author="Sewak R." box="[448,660,1475,1502]" pageId="41" pageNumber="42" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">Sewak 2009b: 59</bibRefCitation>
</treatmentCitation>
(key, diagnosis, distribution). —
<treatmentCitation id="5C7910D5FFDF892F9B33FA72FAD5FA57" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." box="[1074,1324,1475,1501]" page="34" pageId="41" pageNumber="42" year="2011">
<bibRefCitation id="B9494B35FFDF892F9B33FA72FAD5FA57" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." box="[1074,1324,1475,1501]" pageId="41" pageNumber="42" pagination="29 - 38" refId="ref30906" refString="Sabu T. K., Nithya S. &amp; Vinod K. V. 2011. Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa 2830: 29 - 38. https: // doi. org / 10.11646 / ZOOTAXA. 2830.1.3" type="book chapter" year="2011">
Sabu
<emphasis id="EFACEAD6FFDF892F9B79FA75FB4EFA57" box="[1144,1207,1475,1501]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2011: 34
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFDF892F9E13FA57FD40F98B" author="Karimbumkara S. N. &amp; Rajan P. D." box="[274,697,1510,1537]" page="175" pageId="41" pageNumber="42" year="2013">
<bibRefCitation id="B9494B35FFDF892F9E13FA57FD40F98B" author="Karimbumkara S. N. &amp; Rajan P. D." box="[274,697,1510,1537]" pageId="41" pageNumber="42" pagination="173 - 178" refId="ref29842" refString="Karimbumkara S. N. &amp; Rajan P. D. 2013. Insecta: Coleoptera: Scarabaeidae: Scarabaeinae, Dung Beetles. Zoological Survey of India, Fauna of Karnataka, State Fauna Series 21: 173 - 178." type="book chapter" year="2013">Karimbumkara &amp; Rajan 2013: 175</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDF892F9C98FA57FB33F98A" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[921,1226,1510,1536]" page="300" pageId="41" pageNumber="42" year="2014">
<bibRefCitation id="B9494B35FFDF892F9C98FA57FB33F98A" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[921,1226,1510,1536]" pageId="41" pageNumber="42" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFDF892F9B06FA56FBBCF98A" box="[1031,1093,1510,1536]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2014: 300
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDF892F9E0EF9BBFDDCF9AE" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[271,549,1545,1572]" page="1037" pageId="41" pageNumber="42" year="2015">
<bibRefCitation id="B9494B35FFDF892F9E0EF9BBFDDCF9AE" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[271,549,1545,1572]" pageId="41" pageNumber="42" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFDF892F9E5FF9BBFE60F9A9" box="[350,409,1545,1571]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(list, distribution). —
<bibRefCitation id="B9494B35FFDF892F9C2CF9B8FBDEF9AE" author="Schoolmeesters P." box="[813,1063,1545,1572]" pageId="41" pageNumber="42" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDF892F9FBCF9FAFE14F974" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="DD6736C4FFDF892F9FBCF9FAFE3FF9EF" blockId="41.[189,454,1611,1637]" box="[189,454,1611,1637]" pageId="41" pageNumber="42">
<heading id="862F81A8FFDF892F9FBCF9FAFE3FF9EF" bold="true" box="[189,454,1611,1637]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="EFACEAD6FFDF892F9FBCF9FAFE3FF9EF" bold="true" box="[189,454,1611,1637]" pageId="41" pageNumber="42">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCF9C8FE14F974" blockId="41.[189,1399,1657,1790]" pageId="41" pageNumber="42">
The clypeus and vertex of
<taxonomicName id="1AD84D47FFDF892F9D18F9CBFCF9F919" authorityName="Lansberge, AIMB" authorityYear="1875" box="[537,768,1657,1684]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFDF892F9D18F9CBFCF9F919" box="[537,768,1657,1684]" italics="true" pageId="41" pageNumber="42">Onitis singhalensis</emphasis>
</taxonomicName>
have close granules, giving the clypeus a rugose appearance.
<taxonomicName id="1AD84D47FFDF892F9E53F92CFDCCF93D" authorityName="Lansberge, AIMB" authorityYear="1875" box="[338,565,1693,1719]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFDF892F9E53F92CFDCCF93D" box="[338,565,1693,1719]" italics="true" pageId="41" pageNumber="42">Onitis singhalensis</emphasis>
</taxonomicName>
is closer to
<taxonomicName id="1AD84D47FFDF892F9DCEF92CFC9BF93D" authorityName="Fabricius" authorityYear="1801" box="[719,866,1693,1719]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFDF892F9DCEF92CFC9BF93D" box="[719,866,1693,1719]" italics="true" pageId="41" pageNumber="42">O. philemon</emphasis>
</taxonomicName>
,
<taxonomicName id="1AD84D47FFDF892F9C75F92CFC13F93D" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[884,1002,1693,1719]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov.">
<emphasis id="EFACEAD6FFDF892F9C75F92CFC13F93D" box="[884,1002,1693,1719]" italics="true" pageId="41" pageNumber="42">O. kethai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFDF892F9CECF92CFBB3F93D" box="[1005,1098,1693,1719]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="1AD84D47FFDF892F9B89F92CFAF6F93D" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1160,1295,1693,1719]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov.">
<emphasis id="EFACEAD6FFDF892F9B89F92CFAF6F93D" box="[1160,1295,1693,1719]" italics="true" pageId="41" pageNumber="42">O. visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFDF892F9A18F92CFA8EF93D" box="[1305,1399,1693,1719]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
but differs in the nature of the granules on the head and also the size. See remarks under
<taxonomicName id="1AD84D47FFDF892F9BDCF971FA96F950" authorityName="Fabricius" authorityYear="1801" box="[1245,1391,1728,1754]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFDF892F9BDCF971FA96F950" box="[1245,1391,1728,1754]" italics="true" pageId="41" pageNumber="42">O. philemon</emphasis>
</taxonomicName>
,
<taxonomicName id="1AD84D47FFDF892F9FBCF952FED0F977" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[189,297,1763,1789]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="kethai">
<emphasis id="EFACEAD6FFDF892F9FBCF952FED0F977" box="[189,297,1763,1789]" italics="true" pageId="41" pageNumber="42">O. kethai</emphasis>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFDF892F9E63F952FE1CF977" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[354,485,1763,1789]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="visthara">
<emphasis id="EFACEAD6FFDF892F9E63F952FE1CF977" box="[354,485,1763,1789]" italics="true" pageId="41" pageNumber="42">O. visthara</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDF892C9FBCF894FBE3FBA5" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="DD6736C4FFDF892F9FBCF894FE15F8B4" blockId="41.[189,492,1828,1855]" box="[189,492,1828,1855]" pageId="41" pageNumber="42">
<heading id="862F81A8FFDF892F9FBCF894FE15F8B4" bold="true" box="[189,492,1828,1855]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="EFACEAD6FFDF892F9FBCF894FE15F8B4" bold="true" box="[189,492,1828,1855]" pageId="41" pageNumber="42">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FEBF8E2FEA5F8E7" blockId="41.[189,1399,1875,1972]" box="[234,348,1875,1901]" pageId="41" pageNumber="42">
<emphasis id="EFACEAD6FFDF892F9FEBF8E2FEA5F8E7" bold="true" box="[234,348,1875,1901]" pageId="41" pageNumber="42">
<typeStatus id="02638866FFDF892F9FEBF8E2FEA5F8E7" box="[234,348,1875,1901]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCF8C7FCD8F83E" blockId="41.[189,1399,1875,1972]" pageId="41" pageNumber="42">
<materialsCitation id="6DB03C99FFDF892F9FBCF8C7FCD8F83E" collectionCode="MNHN" collectorName="A. Janssens" country="Sri Lanka" location="SRI LANKA" pageId="41" pageNumber="42" specimenCode="EC4812" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="A5CF7654FFDF892F9FBCF8C7FEA3F81A" box="[189,346,1910,1936]" name="Sri Lanka" pageId="41" pageNumber="42">SRI LANKA</collectingCountry>
<specimenCount id="CBDEFD4DFFDF892F9E78F8C6FE6BF81B" box="[377,402,1911,1937]" count="1" pageId="41" pageNumber="42" type="male"></specimenCount>
; “Ceylon, singhalensis typ. Lansbrg., ex-Musaeo E Harold,
<collectorName id="702D5312FFDF892F9B66F8C7FB0AF81B" box="[1127,1267,1910,1937]" pageId="41" pageNumber="42">A. Janssens</collectorName>
vid., 1936
<emphasis id="EFACEAD6FFDF892F9FBCF828FE64F839" box="[189,413,1945,1971]" italics="true" pageId="41" pageNumber="42">Onitissinghalensis</emphasis>
Lansb.,
<typeStatus id="02638866FFDF892F9D03F82BFDB8F83E" box="[514,577,1946,1972]" pageId="41" pageNumber="42">Type</typeStatus>
,”
<collectionCode id="BBC9AE01FFDF892F9D59F82BFD4BF83E" box="[600,690,1946,1972]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="41" pageNumber="42">MNHN</collectionCode>
<specimenCode id="8D7E9EBFFFDF892F9DB8F82BFCE5F83E" box="[697,796,1946,1972]" collectionCode="EC" pageId="41" pageNumber="42">EC4812</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFDC892C9FEBFEB8FCD3FEA9" blockId="42.[189,1399,265,1071]" box="[234,810,265,291]" pageId="42" pageNumber="43">
<emphasis id="EFACEAD6FFDC892C9FEBFEB8FD9BFEA9" bold="true" box="[234,610,265,291]" pageId="42" pageNumber="43">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFDC892C9D70FEB8FCDBFEA9" box="[625,802,265,291]" count="204" pageId="42" pageNumber="43" type="generic">204 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFDC892C9FBCFE9FFBE3FBA5" blockId="42.[189,1399,265,1071]" pageId="42" pageNumber="43">
<materialsCitation id="6DB03C99FFDC892C9FBCFE9FFEDDFEE4" collectionCode="BMNH" collectorName="H. L. Andrewes." country="India" location="Nilgiri Hills." pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFDC892C9FBCFE9FFEF7FEC2" box="[189,270,302,328]" name="India" pageId="42" pageNumber="43">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFDC892C9E32FE9EFEB5FEC3" box="[307,332,303,329]" count="1" pageId="42" pageNumber="43" type="male"></specimenCount>
; “
<location id="D807601FFFDC892C9E69FE9FFDF0FEC2" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9E69FE9FFDF0FEC2" box="[360,521,302,328]" country="India" name="Nilgiri Hills." pageId="42" pageNumber="43">Nilgiri Hills.</location>
,
<collectorName id="702D5312FFDC892C9D1AFE9EFD13FEC2" box="[539,746,302,329]" pageId="42" pageNumber="43">H. L. Andrewes.</collectorName>
, Andrewes Bequest., B.M. 1922221]”; (
<collectionCode id="BBC9AE01FFDC892C9BEBFE9FFABDFEC2" box="[1258,1348,302,328]" pageId="42" pageNumber="43">BMNH</collectionCode>
(E) 1237145
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9E40FEE5FCE2FEE7" box="[321,795,339,366]" collectionCode="BMNH" country="India" location="South India" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFDC892C9E40FEE5FEA1FEE4" box="[321,344,340,366]" count="1" pageId="42" pageNumber="43" type="female"></specimenCount>
;
<location id="D807601FFFDC892C9E69FEE2FE01FEE7" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9E69FEE2FE01FEE7" box="[360,504,339,365]" country="India" name="South India" pageId="42" pageNumber="43">South India</location>
”; (
<collectionCode id="BBC9AE01FFDC892C9D18FEE2FD8AFEE7" box="[537,627,339,365]" pageId="42" pageNumber="43">BMNH</collectionCode>
(E) 1237146)
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9C39FEE2FEBBFE32" collectingDate="1999-05-14" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" latitude="11.837999" location="Budipadaga Scrub" longLatPrecision="1" longitude="77.07755" municipality="Biligiri Rangaswamy Tiger Reserve" pageId="42" pageNumber="43" specimenCount="25" specimenCount-female="5" specimenCount-male="20" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFDC892C9C39FEE2FC74FEE4" box="[824,909,339,366]" count="20" pageId="42" pageNumber="43" type="male">20 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9C9AFEE2FC18FEE4" box="[923,993,339,366]" count="5" pageId="42" pageNumber="43" type="female">5 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFDC892C9CF0FEE2FB94FEE7" box="[1009,1133,339,365]" country="India" name="Karnataka" pageId="42" pageNumber="43">Karnataka</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDC892C9B7BFEE2FE9DFE19" pageId="42" pageNumber="43">Biligiri Rangaswamy Tiger Reserve</collectingMunicipality>
,
<location id="D807601FFFDC892C9E70FEC9FDB4FE18" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9E70FEC9FDB4FE18" box="[369,589,376,402]" country="India" latitude="11.837999" longLatPrecision="1" longitude="77.07755" municipality="Biligiri Rangaswamy Tiger Reserve" name="Budipadaga Scrub" pageId="42" pageNumber="43" stateProvince="Karnataka">Budipadaga Scrub</location>
;
<geoCoordinate id="B8EC5003FFDC892C9D58FEC6FCFCFE19" box="[601,773,375,403]" degrees="11" direction="north" minutes="50" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="16.8" value="11.837999">11°50ʹ16.8″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9C10FEC6FC43FE19" box="[785,954,375,403]" degrees="77" direction="east" minutes="4" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="39.18" value="77.07755">77°4ʹ39.18″ E</geoCoordinate>
;
<date id="A9661004FFDC892C9CC6FEC9FB91FE18" box="[967,1128,376,403]" pageId="42" pageNumber="43" value="1999-05-14">
<collectingDate id="B922E9ECFFDC892C9CC6FEC9FB91FE18" box="[967,1128,376,403]" pageId="42" pageNumber="43" value="1999-05-14">14 May 1999</collectingDate>
</date>
;
<collectionCode id="BBC9AE01FFDC892C9B72FEC9FB39FE19" box="[1139,1216,376,403]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9BC8FEC9FB12FE19" box="[1225,1259,376,403]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9BF5FEC9FAE9FE19" box="[1268,1296,376,403]" pageId="42" pageNumber="43">Sc</collectorName>
1000502 to 1000526
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9E5BFE2FFDA9FE57" collectingDate="2000-03-30" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" latitude="11.931833" location="Gombegallu" longLatPrecision="1" longitude="77.1843" pageId="42" pageNumber="43" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="CBDEFD4DFFDC892C9E5BFE2FFE61FE32" box="[346,408,414,440]" count="2" pageId="42" pageNumber="43" type="male">2♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9EA2FE2FFE26FE32" box="[419,479,414,440]" count="1" pageId="42" pageNumber="43" type="female">1♀♀</specimenCount>
;
<location id="D807601FFFDC892C9EECFE2CFD87FE3D" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9EECFE2CFD87FE3D" box="[493,638,413,439]" country="India" latitude="11.931833" longLatPrecision="1" longitude="77.1843" name="Gombegallu" pageId="42" pageNumber="43">Gombegallu</location>
evergreen forest;
<geoCoordinate id="B8EC5003FFDC892C9C53FE2DFC05FE32" box="[850,1020,412,440]" degrees="11" direction="north" minutes="55" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="54.6" value="11.931833">11°55ʹ54.6″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9B09FE2DFB56FE32" box="[1032,1199,412,440]" degrees="77" direction="east" minutes="11" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="3.48" value="77.1843">77°11ʹ3.48″ E</geoCoordinate>
;
<date id="A9661004FFDC892C9BBAFE2FFA8BFE32" box="[1211,1394,413,440]" pageId="42" pageNumber="43" value="2000-03-30">
<collectingDate id="B922E9ECFFDC892C9BBAFE2FFA8BFE32" box="[1211,1394,413,440]" pageId="42" pageNumber="43" value="2000-03-30">30 March 2000</collectingDate>
</date>
;
<collectionCode id="BBC9AE01FFDC892C9FBCFE73FEF3FE57" box="[189,266,450,477]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9E12FE73FECCFE57" box="[275,309,450,477]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9E3FFE73FEA3FE57" box="[318,346,450,477]" pageId="42" pageNumber="43">Sc</collectorName>
1000527 to 1000558
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9D6DFE72FC51FD89" collectingDate="1999-12-22" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" latitude="11.89545" location="Kanneri" longLatPrecision="1" longitude="77.13596" pageId="42" pageNumber="43" specimenCount="27" specimenCount-female="10" specimenCount-male="17">
<specimenCount id="CBDEFD4DFFDC892C9D6DFE72FD39FE57" box="[620,704,451,477]" count="17" pageId="42" pageNumber="43" type="male">17 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9DCFFE72FCD8FE57" box="[718,801,451,477]" count="10" pageId="42" pageNumber="43" type="female">10 ♀♀</specimenCount>
;
<location id="D807601FFFDC892C9C33FE72FC69FE57" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9C33FE72FC69FE57" box="[818,912,451,477]" country="India" latitude="11.89545" longLatPrecision="1" longitude="77.13596" name="Kanneri" pageId="42" pageNumber="43">Kanneri</location>
moist deciduous forest;
<geoCoordinate id="B8EC5003FFDC892C9BB6FE70FA8BFE57" box="[1207,1394,449,477]" degrees="11" direction="north" minutes="53" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="43.62" value="11.89545">11°53ʹ43.62″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9FBCFE57FEA1FD88" box="[189,344,486,514]" degrees="77" direction="east" minutes="8" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="9.48" value="77.13596">77°8ʹ9.48″ E</geoCoordinate>
;
<date id="A9661004FFDC892C9E67FE59FDF3FD88" box="[358,522,488,514]" pageId="42" pageNumber="43" value="1999-12-22">
<collectingDate id="B922E9ECFFDC892C9E67FE59FDF3FD88" box="[358,522,488,514]" pageId="42" pageNumber="43" value="1999-12-22">22 Dec. 1999</collectingDate>
</date>
;
<collectionCode id="BBC9AE01FFDC892C9D17FE59FD9AFD89" box="[534,611,488,515]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9D6AFE59FD74FD89" box="[619,653,488,515]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9D97FE59FD4BFD89" box="[662,690,488,515]" pageId="42" pageNumber="43">Sc</collectorName>
1000559 to 1000585
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9CC5FE59FB5CFDAD" collectingDate="1999-05-11" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" latitude="12.032" location="Purani" longLatPrecision="1" longitude="77.1313" pageId="42" pageNumber="43" specimenCount="26" specimenCount-female="7" specimenCount-male="19">
<specimenCount id="CBDEFD4DFFDC892C9CC5FE59FBE1FD88" box="[964,1048,488,514]" count="19" pageId="42" pageNumber="43" type="male">19 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9B27FE59FB93FD88" box="[1062,1130,488,514]" count="7" pageId="42" pageNumber="43" type="female">7 ♀♀</specimenCount>
;
<location id="D807601FFFDC892C9B7BFE59FB3EFD88" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9B7BFE59FB3EFD88" box="[1146,1223,488,514]" country="India" latitude="12.032" longLatPrecision="1" longitude="77.1313" name="Purani" pageId="42" pageNumber="43">Purani</location>
dry deciduous forest;
<geoCoordinate id="B8EC5003FFDC892C9E0EFDBAFE55FDAD" box="[271,428,523,551]" degrees="12" direction="north" minutes="1" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="55.2" value="12.032">12°1ʹ55.2″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9EB9FDBAFD98FDAD" box="[440,609,523,551]" degrees="77" direction="east" minutes="7" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="52.68" value="77.1313">77°7ʹ52.68″ E</geoCoordinate>
;
<date id="A9661004FFDC892C9D6CFDBCFCF5FDAD" box="[621,780,525,551]" pageId="42" pageNumber="43" value="1999-05-11">
<collectingDate id="B922E9ECFFDC892C9D6CFDBCFCF5FDAD" box="[621,780,525,551]" pageId="42" pageNumber="43" value="1999-05-11">11 May 1999</collectingDate>
</date>
;
<collectionCode id="BBC9AE01FFDC892C9C16FDBCFC9DFDA2" box="[791,868,525,552]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9C6DFDBCFC77FDA2" box="[876,910,525,552]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9C96FDBCFC4AFDA2" box="[919,947,525,552]" pageId="42" pageNumber="43">Sc</collectorName>
1000586 to 1000611
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9BBFFDBCFA8EFDC6" collectingDate="1998-04" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" latitude="11.8907" location="Honnametti Shola" longLatPrecision="1" longitude="77.20138" pageId="42" pageNumber="43" specimenCount="37" specimenCount-female="15" specimenCount-male="22">
<specimenCount id="CBDEFD4DFFDC892C9BBFFDBCFAE8FDAD" box="[1214,1297,525,551]" count="22" pageId="42" pageNumber="43" type="male">22 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9A1CFDBCFA96FDAD" box="[1309,1391,525,552]" count="15" pageId="42" pageNumber="43" type="female">15 ♀♀</specimenCount>
;
<location id="D807601FFFDC892C9FBCFD83FE65FDC6" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9FBCFD83FE65FDC6" box="[189,412,562,588]" country="India" latitude="11.8907" longLatPrecision="1" longitude="77.20138" name="Honnametti Shola" pageId="42" pageNumber="43">Honnametti Shola</location>
;
<geoCoordinate id="B8EC5003FFDC892C9EB1FD81FD88FDC7" box="[432,625,560,589]" degrees="11" direction="north" minutes="53" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="26.52" value="11.8907">11°53ʹ26.52″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9D85FD81FCCDFDC7" box="[644,820,560,589]" degrees="77" direction="east" minutes="12" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="4.98" value="77.20138">77°12ʹ4.98″ E</geoCoordinate>
;
<date id="A9661004FFDC892C9C47FD83FC3FFDC6" box="[838,966,562,588]" pageId="42" pageNumber="43" value="1998-04">
<collectingDate id="B922E9ECFFDC892C9C47FD83FC3FFDC6" box="[838,966,562,588]" pageId="42" pageNumber="43" value="1998-04">Apr. 1998</collectingDate>
</date>
;
<collectionCode id="BBC9AE01FFDC892C9CD9FD83FBDCFDC6" box="[984,1061,562,588]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9B2FFD83FBA9FDC6" box="[1070,1104,562,588]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9B58FD83FB8CFDC6" box="[1113,1141,562,588]" pageId="42" pageNumber="43">Sc</collectorName>
1000612 to 1000648
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9FCFFDE6FC04FD1C" collectingDate="1998-05" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="12.0195" location="Gummane Shola" longLatPrecision="1" longitude="77.17518" pageId="42" pageNumber="43" specimenCount="25" specimenCount-female="9" specimenCount-male="16">
<specimenCount id="CBDEFD4DFFDC892C9FCFFDE6FED9FDF8" box="[206,288,599,626]" count="16" pageId="42" pageNumber="43" type="male">16 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9E2DFDE6FE96FDF8" box="[300,367,599,626]" count="9" pageId="42" pageNumber="43" type="female">9 ♀♀</specimenCount>
;
<location id="D807601FFFDC892C9E7CFDE6FDB8FDFB" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9E7CFDE6FDB8FDFB" box="[381,577,599,625]" country="India" latitude="12.0195" longLatPrecision="1" longitude="77.17518" name="Gummane Shola" pageId="42" pageNumber="43">Gummane Shola</location>
<geoCoordinate id="B8EC5003FFDC892C9D49FDE7FD1DFDF8" box="[584,740,598,626]" degrees="12" direction="north" minutes="1" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="10.2" value="12.0195">12°1ʹ10.2″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9DF1FDE7FC5EFDF8" box="[752,935,598,626]" degrees="77" direction="east" minutes="10" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="30.66" value="77.17518">77°10ʹ30.66″ E</geoCoordinate>
;
<date id="A9661004FFDC892C9CB2FDE9FBD4FDFB" box="[947,1069,599,626]" pageId="42" pageNumber="43" value="1998-05">
<collectingDate id="B922E9ECFFDC892C9CB2FDE9FBD4FDFB" box="[947,1069,599,626]" pageId="42" pageNumber="43" value="1998-05">May 1998</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFDC892C9B38FDE9FA8BFDF8" box="[1081,1394,599,626]" pageId="42" pageNumber="43">open cow dung baited trap</collectingMethod>
;
<collectorName id="702D5312FFDC892C9FBCFDCDFDE5FD1C" box="[189,540,636,662]" pageId="42" pageNumber="43">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDC892C9D67FDCDFD4AFD1C" box="[614,691,636,662]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9DBAFDCDFD24FD1C" box="[699,733,636,662]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9DE7FDCDFCFBFD1C" box="[742,770,636,662]" pageId="42" pageNumber="43">Sc</collectorName>
1000649 to 1000673
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9B1FFDCCFEBFFD6B" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" latitude="12.0205" location="Core" longLatPrecision="1" longitude="76.104095" municipality="Nagarhole National Park" pageId="42" pageNumber="43" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFDC892C9B1FFDCCFBCFFD1D" box="[1054,1078,637,663]" count="1" pageId="42" pageNumber="43" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9B46FDCCFBA7FD1D" box="[1095,1118,637,663]" count="1" pageId="42" pageNumber="43" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFDC892C9B70FDCDFB14FD1C" box="[1137,1261,636,662]" country="India" name="Karnataka" pageId="42" pageNumber="43">Karnataka</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDC892C9BFDFDCDFE93FD31" pageId="42" pageNumber="43">Nagarhole National Park</collectingMunicipality>
,
<location id="D807601FFFDC892C9E7FFD13FE4EFD36" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9E7FFD13FE4EFD36" box="[382,439,674,700]" country="India" latitude="12.0205" longLatPrecision="1" longitude="76.104095" municipality="Nagarhole National Park" name="Core" pageId="42" pageNumber="43" stateProvince="Karnataka">Core</location>
moist deciduous forest;
<geoCoordinate id="B8EC5003FFDC892C9DF6FD11FC65FD36" box="[759,924,672,700]" degrees="12" direction="north" minutes="1" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="13.8" value="12.0205">12°1ʹ13.8″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9CB1FD11FB99FD36" box="[944,1120,672,700]" degrees="76" direction="east" minutes="6" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="14.76" value="76.104095">76°6ʹ14.76″ E</geoCoordinate>
;
<collectionCode id="BBC9AE01FFDC892C9B72FD10FB39FD31" box="[1139,1216,673,699]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9BC8FD10FB12FD31" box="[1225,1259,673,699]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9BF5FD10FAE9FD31" box="[1268,1296,673,699]" pageId="42" pageNumber="43">Sc</collectorName>
1000674 to 1000675
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9E66FD76FE34FC8C" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" latitude="12.1563" location="Hebbala" longLatPrecision="1" longitude="76.114395" pageId="42" pageNumber="43" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="CBDEFD4DFFDC892C9E66FD76FE56FD6B" box="[359,431,711,737]" count="2" pageId="42" pageNumber="43" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9EC1FD76FE2EFD6B" box="[448,471,711,737]" count="1" pageId="42" pageNumber="43" type="female"></specimenCount>
;
<location id="D807601FFFDC892C9EE8FD77FDB2FD6A" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9EE8FD77FDB2FD6A" box="[489,587,710,736]" country="India" latitude="12.1563" longLatPrecision="1" longitude="76.114395" name="Hebbala" pageId="42" pageNumber="43">Hebbala</location>
moist deciduous forest;
<geoCoordinate id="B8EC5003FFDC892C9C7DFD74FBD5FD6B" box="[892,1068,709,737]" degrees="12" direction="north" minutes="9" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="22.68" value="12.1563">12°9ʹ22.68″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9B3DFD74FB11FD6B" box="[1084,1256,709,737]" degrees="76" direction="east" minutes="6" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="51.84" value="76.114395">76°6ʹ51.84″ E</geoCoordinate>
;
<collectionCode id="BBC9AE01FFDC892C9BF6FD76FABAFD6B" box="[1271,1347,711,737]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9A4DFD76FA94FD6B" box="[1356,1389,711,737]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9FBCFD5DFF20FC8C" box="[189,217,748,774]" pageId="42" pageNumber="43">Sc</collectorName>
1000676 to 1000678
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9EE4FD5DFAE0FCA1" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="11.974199" location="Sungathakatte" longLatPrecision="1" longitude="76.1856" pageId="42" pageNumber="43" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFDC892C9EE4FD5DFE04FC8C" box="[485,509,748,774]" count="1" pageId="42" pageNumber="43" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9D08FD5DFDB5FC8C" box="[521,588,748,774]" count="2" pageId="42" pageNumber="43" type="female">2 ♀♀</specimenCount>
;
<location id="D807601FFFDC892C9D5BFD5DFCF9FC8C" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9D5BFD5DFCF9FC8C" box="[602,768,748,774]" country="India" latitude="11.974199" longLatPrecision="1" longitude="76.1856" name="Sungathakatte" pageId="42" pageNumber="43">Sungathakatte</location>
dry deciduous forest;
<geoCoordinate id="B8EC5003FFDC892C9B04FD5BFB47FC8C" box="[1029,1214,746,774]" degrees="11" direction="north" minutes="58" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="27.12" value="11.974199">11°58ʹ27.12″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9BCBFD5BFA88FC8C" box="[1226,1393,746,774]" degrees="76" direction="east" minutes="11" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="8.16" value="76.1856">76°11ʹ8.16″ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFDC892C9FBCFCA0FE08FCA1" box="[189,497,785,811]" pageId="42" pageNumber="43">open cow dung baited trap</collectingMethod>
;
<collectorName id="702D5312FFDC892C9EFAFCA0FCB6FCA1" box="[507,847,785,811]" pageId="42" pageNumber="43">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDC892C9C8CFCA0FC23FCA1" box="[909,986,785,811]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9CE2FCA0FBFCFCA1" box="[995,1029,785,811]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9B0CFCA0FBD0FCA1" box="[1037,1065,785,811]" pageId="42" pageNumber="43">Sc</collectorName>
1000679 to 1000681
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9A2EFCA0FC41FCFF" collectingDate="2009-05-08" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" elevation="839" latitude="8.861611" location="Rockwood" longLatPrecision="1" longitude="77.13347" municipality="Shendurney Wildlife Sanctuary" pageId="42" pageNumber="43" specimenCount="13" specimenCount-female="8" specimenCount-male="5" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDC892C9A2EFCA0FA88FCA1" box="[1327,1393,785,812]" count="5" pageId="42" pageNumber="43" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9FBCFC87FEFAFCDB" box="[189,259,822,849]" count="8" pageId="42" pageNumber="43" type="female">8 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFDC892C9E14FC87FE9FFCDA" box="[277,358,822,848]" country="India" name="Kerala" pageId="42" pageNumber="43">Kerala</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFDC892C9E77FC87FD17FCDA" box="[374,750,822,848]" pageId="42" pageNumber="43">Shendurney Wildlife Sanctuary</collectingMunicipality>
,
<location id="D807601FFFDC892C9DFCFC87FC85FCDA" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9DFCFC87FC85FCDA" box="[765,892,822,848]" country="India" latitude="8.861611" longLatPrecision="1" longitude="77.13347" municipality="Shendurney Wildlife Sanctuary" name="Rockwood" pageId="42" pageNumber="43" stateProvince="Kerala">Rockwood</location>
grassland;
<geoCoordinate id="B8EC5003FFDC892C9B08FC85FB4EFCDB" box="[1033,1207,820,849]" degrees="08" direction="north" minutes="51" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="41.8" value="8.861611">08°51ʹ41.8ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9BC7FC85FA88FCDB" box="[1222,1393,820,849]" degrees="77" direction="east" minutes="08" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="00.5" value="77.13347">77°08ʹ00.5ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFDC892C9FFBFCEAFE7AFCFF" box="[250,387,859,886]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="42" pageNumber="43" unit="m" value="839.0">
<quantity id="1A209B21FFDC892C9FFBFCEAFEBDFCFC" box="[250,324,859,886]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="42" pageNumber="43" unit="m" value="839.0">839 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFDC892C9E91FCEAFDD9FCFF" box="[400,544,859,886]" pageId="42" pageNumber="43" value="2009-05-08">
<collectingDate id="B922E9ECFFDC892C9E91FCEAFDD9FCFF" box="[400,544,859,886]" pageId="42" pageNumber="43" value="2009-05-08">8 May 2009</collectingDate>
</date>
;
<collectionCode id="BBC9AE01FFDC892C9D2BFCEAFD8EFCFF" box="[554,631,859,885]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9D7EFCEAFD58FCFF" box="[639,673,859,885]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9DABFCEAFD3FFCFF" box="[682,710,859,885]" pageId="42" pageNumber="43">Sc</collectorName>
1000682 to 1000694
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9CCEFCEAFA8EFC10" collectingDate="2009-12-03" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" elevation="185" latitude="8.9151945" location="Kattilappara" longLatPrecision="1" longitude="77.10797" pageId="42" pageNumber="43" specimenCount="2" specimenCount-male="2">
<specimenCount id="CBDEFD4DFFDC892C9CCEFCEAFBEDFCFC" box="[975,1044,859,886]" count="2" pageId="42" pageNumber="43" type="male">2 ♂♂</specimenCount>
;
<location id="D807601FFFDC892C9B21FCEAFB48FCFF" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9B21FCEAFB48FCFF" box="[1056,1201,859,885]" country="India" latitude="8.9151945" longLatPrecision="1" longitude="77.10797" name="Kattilappara" pageId="42" pageNumber="43">Kattilappara</location>
moist deciduous forest;
<geoCoordinate id="B8EC5003FFDC892C9E0EFCCEFE40FC11" box="[271,441,895,923]" degrees="08" direction="north" minutes="54" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="54.7" value="8.9151945">08°54ʹ54.7ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9EC5FCCEFD92FC11" box="[452,619,895,923]" degrees="77" direction="east" minutes="06" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="28.7" value="77.10797">77°06ʹ28.7ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFDC892C9DB4FC31FCC6FC10" box="[693,831,896,923]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="42" pageNumber="43" unit="m" value="185.0">
<quantity id="1A209B21FFDC892C9DB4FC31FD06FC11" box="[693,767,896,923]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="42" pageNumber="43" unit="m" value="185.0">185 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFDC892C9C4DFC31FC24FC10" box="[844,989,896,923]" pageId="42" pageNumber="43" value="2009-12-03">
<collectingDate id="B922E9ECFFDC892C9C4DFC31FC24FC10" box="[844,989,896,923]" pageId="42" pageNumber="43" value="2009-12-03">3 Dec. 2009</collectingDate>
</date>
;
<collectionCode id="BBC9AE01FFDC892C9CE6FC31FBCDFC11" box="[999,1076,896,923]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9B3FFC31FB99FC11" box="[1086,1120,896,923]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9B69FC31FB7DFC11" box="[1128,1156,896,923]" pageId="42" pageNumber="43">Sc</collectorName>
1000695 to 1000696
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9FD0FC17FBFBFC6E" collectingDate="2009-12-03" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" elevation="207" latitude="8.918222" location="Kattilapara" longLatPrecision="1" longitude="77.09575" pageId="42" pageNumber="43" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCount id="CBDEFD4DFFDC892C9FD0FC17FEE1FC4A" box="[209,280,934,960]" count="2" pageId="42" pageNumber="43" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9E26FC17FE95FC4A" box="[295,364,934,960]" count="2" pageId="42" pageNumber="43" type="female">2 ♀♀</specimenCount>
;
<location id="D807601FFFDC892C9E7CFC14FE06FC35" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9E7CFC14FE06FC35" box="[381,511,933,959]" country="India" latitude="8.918222" longLatPrecision="1" longitude="77.09575" name="Kattilapara" pageId="42" pageNumber="43">Kattilapara</location>
degraded forest;
<geoCoordinate id="B8EC5003FFDC892C9DD5FC15FC78FC4A" box="[724,897,932,960]" degrees="08" direction="north" minutes="55" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="05.6" value="8.918222">08°55ʹ05.6ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9C8EFC15FBC0FC4A" box="[911,1081,932,960]" degrees="77" direction="east" minutes="05" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="44.7" value="77.09575">77°05ʹ44.7ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFDC892C9B89FC17FAEEFC35" box="[1160,1303,933,960]" metricMagnitude="2" metricUnit="m" metricValue="2.07" pageId="42" pageNumber="43" unit="m" value="207.0">
<quantity id="1A209B21FFDC892C9B89FC17FB2CFC4A" box="[1160,1237,934,960]" metricMagnitude="2" metricUnit="m" metricValue="2.07" pageId="42" pageNumber="43" unit="m" value="207.0">207 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFDC892C9A26FC17FF02FC6F" pageId="42" pageNumber="43" value="2009-12-03">
<collectingDate id="B922E9ECFFDC892C9A26FC17FF02FC6F" pageId="42" pageNumber="43" value="2009-12-03">3 Dec. 2009</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFDC892C9E0DFC7AFDA0FC6F" box="[268,601,970,997]" pageId="42" pageNumber="43">open cow dung baited trap</collectingMethod>
;
<collectionCode id="BBC9AE01FFDC892C9D68FC7BFD4FFC6E" box="[617,694,970,996]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9DBFFC7BFD19FC6E" box="[702,736,970,996]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9DE8FC7BFCFCFC6E" box="[745,773,970,996]" pageId="42" pageNumber="43">Sc</collectorName>
1000696 to 1000699
</materialsCitation>
<materialsCitation id="6DB03C99FFDC892C9B25FC7AFBECFBA5" collectingDate="2009-12-03" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" elevation="144" latitude="8.923028" location="Kattilappara" longLatPrecision="1" longitude="77.09422" pageId="42" pageNumber="43" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="CBDEFD4DFFDC892C9B25FC7AFB94FC6F" box="[1060,1133,971,997]" count="2" pageId="42" pageNumber="43" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFDC892C9B7FFC7AFB6CFC6F" box="[1150,1173,971,997]" count="1" pageId="42" pageNumber="43" type="female"></specimenCount>
;
<location id="D807601FFFDC892C9BA9FC7BFAC0FC6E" LSID="urn:lsid:plazi:treatment:557187D2FFDF892C9D40FB67FBC9F863:D807601FFFDC892C9BA9FC7BFAC0FC6E" box="[1192,1337,970,996]" country="India" latitude="8.923028" longLatPrecision="1" longitude="77.09422" name="Kattilappara" pageId="42" pageNumber="43">Kattilappara</location>
teak plantation;
<geoCoordinate id="B8EC5003FFDC892C9E44FC5FFE0BFB80" box="[325,498,1006,1034]" degrees="08" direction="north" minutes="55" orientation="latitude" pageId="42" pageNumber="43" precision="1" seconds="22.9" value="8.923028">08°55ʹ22.9ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDC892C9D00FC5FFD55FB80" box="[513,684,1006,1034]" degrees="77" direction="east" minutes="05" orientation="longitude" pageId="42" pageNumber="43" precision="1" seconds="39.2" value="77.09422">77°05ʹ39.2ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFDC892C9DFDFC41FC74FB80" box="[764,909,1008,1034]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="42" pageNumber="43" unit="m" value="144.0">
<quantity id="1A209B21FFDC892C9DFDFC41FCB3FB80" box="[764,842,1008,1034]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="42" pageNumber="43" unit="m" value="144.0">144 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFDC892C9C9FFC41FBCFFB80" box="[926,1078,1008,1034]" pageId="42" pageNumber="43" value="2009-12-03">
<collectingDate id="B922E9ECFFDC892C9C9FFC41FBCFFB80" box="[926,1078,1008,1034]" pageId="42" pageNumber="43" value="2009-12-03">3 Dec. 2009</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFDC892C9B47FC41FA8BFB80" box="[1094,1394,1008,1034]" pageId="42" pageNumber="43">open cow dung bait trap</collectingMethod>
;
<collectorName id="702D5312FFDC892C9FBCFBA4FDBAFBA5" box="[189,579,1045,1071]" pageId="42" pageNumber="43">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFDC892C9D82FBA4FD29FBA5" box="[643,720,1045,1071]" pageId="42" pageNumber="43">AIMB</collectionCode>
/
<collectorName id="702D5312FFDC892C9DD9FBA4FD03FBA5" box="[728,762,1045,1071]" pageId="42" pageNumber="43">Co</collectorName>
/
<collectorName id="702D5312FFDC892C9C02FBA4FCE6FBA5" box="[771,799,1045,1071]" pageId="42" pageNumber="43">Sc</collectorName>
1000700 to 1000702
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDC892C9FBCFBE9FAB2F8A1" pageId="42" pageNumber="43" type="description">
<paragraph id="DD6736C4FFDC892C9FBCFBE9FEB7FBF8" blockId="42.[189,334,1112,1138]" box="[189,334,1112,1138]" pageId="42" pageNumber="43">
<heading id="862F81A8FFDC892C9FBCFBE9FEB7FBF8" bold="true" box="[189,334,1112,1138]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="EFACEAD6FFDC892C9FBCFBE9FEB7FBF8" bold="true" box="[189,334,1112,1138]" pageId="42" pageNumber="43">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDC892C9FBCFB38FCFCFB29" blockId="42.[189,773,1161,1187]" box="[189,773,1161,1187]" pageId="42" pageNumber="43">MEASUREMENTS (in mm). TL=1623, BW= 912.</paragraph>
<paragraph id="DD6736C4FFDC892C9FEBFB7DFED3FB6C" blockId="42.[189,1399,1228,1588]" box="[234,298,1228,1254]" pageId="42" pageNumber="43">
<emphasis id="EFACEAD6FFDC892C9FEBFB7DFED3FB6C" bold="true" box="[234,298,1228,1254]" pageId="42" pageNumber="43">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDC892C9FBCFB40FCFEF9BE" blockId="42.[189,1399,1228,1588]" pageId="42" pageNumber="43">Coppery or greenish coppery, not very shining, elongate oval; head closely granular, clypeal margin slightly bidentate in the middle, clypeofrontal carina interrupted in the middle with an anterior short clypeal carina and a conical tubercle immediately behind; pronotum fairly strongly and unevenly punctured and with a smooth median line, lateral margin feebly curved anteriorly, bisinuate behind, front angle blunt, hind angle obsolete, base strongly rounded; elytra moderately strongly striate with intervals very minutely punctured; pygidium scarcely perceptibly punctured; metasternal shield strongly longitudinally grooved in the middle anteriorly, smooth below, granular and hairy at the sides, sides of the metasternum closely granular and hairy. Profemur with a sharp tooth towards the end of its anterior edge, mid femur and hind trochanter also toothed.</paragraph>
<paragraph id="DD6736C4FFDC892C9FBCF9EFFCC6F948" blockId="42.[189,1398,1629,1730]" pageId="42" pageNumber="43">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFDC892C9EAFF9EFFDDBF9F2" box="[430,546,1630,1656]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="42" pageNumber="43">Fig. 8IL</figureCitation>
). LP=3.5, Lp= 2.75; BP= 1.25, BpB =0.75, BpT= 0.5. Phallobase longer than the parameres. The base of the parameres is straight near the phallobase, forms an angle at one third of its length, is straight anteriorly and curves at the tip.
</paragraph>
<paragraph id="DD6736C4FFDC892C9FEBF95AFEBCF88F" blockId="42.[189,1355,1771,1835]" box="[234,325,1771,1797]" pageId="42" pageNumber="43">
<emphasis id="EFACEAD6FFDC892C9FEBF95AFEBCF88F" bold="true" box="[234,325,1771,1797]" pageId="42" pageNumber="43">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDC892C9FBCF8A1FAB2F8A1" blockId="42.[189,1355,1771,1835]" box="[189,1355,1808,1835]" pageId="42" pageNumber="43">Clypeus longer, rugulose; protibia broad and armed with four external teeth and an articulated spur.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDC892C9FBCF8E2FBC9F863" pageId="42" pageNumber="43" type="distribution">
<paragraph id="DD6736C4FFDC892C9FBCF8E2FEAFF8E7" blockId="42.[189,342,1875,1901]" box="[189,342,1875,1901]" pageId="42" pageNumber="43">
<heading id="862F81A8FFDC892C9FBCF8E2FEAFF8E7" bold="true" box="[189,342,1875,1901]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="EFACEAD6FFDC892C9FBCF8E2FEAFF8E7" bold="true" box="[189,342,1875,1901]" pageId="42" pageNumber="43">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDC892C9FBCF835FBC9F863" blockId="42.[189,1398,1924,2025]" pageId="42" pageNumber="43">
<collectingCountry id="A5CF7654FFDC892C9FBCF835FF07F814" box="[189,254,1924,1950]" name="India" pageId="42" pageNumber="43">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFDC892C9E08F835FE9DF814" box="[265,356,1924,1950]" country="India" name="Gujarat" pageId="42" pageNumber="43">Gujarat</collectingRegion>
,
<collectingRegion id="1F1CF826FFDC892C9E71F835FE15F814" box="[368,492,1924,1950]" country="India" name="Karnataka" pageId="42" pageNumber="43">Karnataka</collectingRegion>
;
<collectingRegion id="1F1CF826FFDC892C9EF8F835FDB3F814" box="[505,586,1924,1950]" country="India" name="Kerala" pageId="42" pageNumber="43">Kerala</collectingRegion>
,
<collectingRegion id="1F1CF826FFDC892C9D56F835FD37F814" box="[599,718,1924,1950]" country="India" name="Rajasthan" pageId="42" pageNumber="43">Rajasthan</collectingRegion>
,
<collectingRegion id="1F1CF826FFDC892C9DD8F835FC91F814" box="[729,872,1924,1950]" country="India" name="Tamil Nadu" pageId="42" pageNumber="43">Tamil Nadu</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFDC892C9B3BF835FB48F814" box="[1082,1201,1924,1950]" name="Sri Lanka" pageId="42" pageNumber="43">Sri Lanka</collectingCountry>
;
<collectingCountry id="A5CF7654FFDC892C9BBFF835FADDF814" box="[1214,1316,1924,1950]" name="Pakistan" pageId="42" pageNumber="43">Pakistan</collectingCountry>
,
<collectingCountry id="A5CF7654FFDC892C9A31F835FA8FF814" box="[1328,1398,1924,1950]" name="Nepal" pageId="42" pageNumber="43">Nepal</collectingCountry>
(Biswas &amp; Chatterjee 1986;
<bibRefCitation id="B9494B35FFDC892C9D0CF818FD3DF84E" author="Chandra K." box="[525,708,1961,1988]" pageId="42" pageNumber="43" pagination="79 - 89" refId="ref28355" refString="Chandra K. 2009 a. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Tamil Nadu, State Fauna Series 1: 79 - 89." type="book chapter" year="2009">Chandra 2009a</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDC892C9DD1F818FC88F84E" author="Sewak R." box="[720,881,1961,1988]" pageId="42" pageNumber="43" pagination="1 - 48" refId="ref31194" refString="Sewak R. 2009 a. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar desert of Gujarat. Records of the Zoological Survey of India, Occassional Paper No 295: 1 - 48." type="journal article" year="2009">Sewak 2009a</bibRefCitation>
,
<bibRefCitation id="B9494B35FFDC892C9C7CF818FC33F849" author="Sewak R." box="[893,970,1961,1987]" pageId="42" pageNumber="43" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">2009b</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDC892C9CD6F818FB60F84E" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." box="[983,1177,1961,1988]" pageId="42" pageNumber="43" pagination="29 - 38" refId="ref30906" refString="Sabu T. K., Nithya S. &amp; Vinod K. V. 2011. Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa 2830: 29 - 38. https: // doi. org / 10.11646 / ZOOTAXA. 2830.1.3" type="book chapter" year="2011">
Sabu
<emphasis id="EFACEAD6FFDC892C9B18F81BFBACF849" box="[1049,1109,1961,1987]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
2011
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDC892C9BA7F818FEBEF863" author="Karimbumkara S. N. &amp; Rajan P. D." pageId="42" pageNumber="43" pagination="173 - 178" refId="ref29842" refString="Karimbumkara S. N. &amp; Rajan P. D. 2013. Insecta: Coleoptera: Scarabaeidae: Scarabaeinae, Dung Beetles. Zoological Survey of India, Fauna of Karnataka, State Fauna Series 21: 173 - 178." type="book chapter" year="2013">Karimbumkara &amp; Rajan 2013</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDC892C9E52F87FFDC7F863" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[339,574,1998,2025]" pageId="42" pageNumber="43" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
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2014
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<bibRefCitation id="B9494B35FFDC892C9D4BF87EFCE0F863" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[586,793,1998,2025]" pageId="42" pageNumber="43" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
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<bibRefCitation id="B9494B35FFDC892C9C24F87FFBDDF863" author="Schoolmeesters P." box="[805,1060,1998,2025]" pageId="42" pageNumber="43" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,544 @@
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<subSubSection id="95C2654FFFE089109D82FC59FC48FB88" box="[643,945,999,1026]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="DD6736C4FFE089109D82FC59FC48FB88" blockId="22.[643,945,999,1063]" box="[643,945,999,1026]" pageId="22" pageNumber="23">
<heading id="862F81A8FFE089109D82FC59FC48FB88" box="[643,945,999,1026]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="23" reason="2">
<taxonomicName id="1AD84D47FFE089109D82FC59FC48FB88" ID-CoL="49KG7" authority="Felsche, 1907" authorityName="Felsche" authorityYear="1907" box="[643,945,999,1026]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="feae">
<emphasis id="EFACEAD6FFE089109D82FC59FCFFFB8B" bold="true" box="[643,774,999,1026]" italics="true" pageId="22" pageNumber="23">Onitis feae</emphasis>
<bibRefCitation id="B9494B35FFE089109C0FFC59FC48FB88" author="Felsche C." box="[782,945,1000,1026]" pageId="22" pageNumber="23" pagination="273 - 296" refId="ref29092" refString="Felsche C. 1907. Coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift: 273 - 296." type="book chapter" year="1907">Felsche, 1907</bibRefCitation>
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<subSubSection id="95C2654FFFE089109D9CFBBDFC6FFBAD" box="[669,918,1036,1063]" pageId="22" pageNumber="23" type="description">
<paragraph id="DD6736C4FFE089109D9CFBBDFC6FFBAD" blockId="22.[643,945,999,1063]" box="[669,918,1036,1063]" pageId="22" pageNumber="23">
<figureCitation id="45E32A41FFE089109D9CFBBDFCF4FBAD" box="[669,781,1036,1063]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="22" pageNumber="23">Figs 10A</figureCitation>
,
<figureCitation id="45E32A41FFE089109C19FBBCFC94FBAD" box="[792,877,1037,1063]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. EH. O. feae Felsche, 1907, AIMB/Co/Sc1000188. IL. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." pageId="22" pageNumber="23">11EH</figureCitation>
,
<figureCitation id="45E32A41FFE089109C78FBBCFC6FFBAD" box="[889,918,1037,1063]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="22" pageNumber="23">18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE089109FBCFBE1FE3EFA9C" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="DD6736C4FFE089109FBCFBE1FCD3FBE3" blockId="22.[189,810,1103,1130]" box="[189,810,1103,1130]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="FDC811EAFFE089109FBCFBE1FCD3FBE3" box="[189,810,1103,1130]" pageId="22" pageNumber="23">
<treatmentCitation id="5C7910D5FFE089109FBCFBE1FDDAFBE0" author="Felsche C." box="[189,547,1103,1130]" page="293" pageId="22" pageNumber="23" year="1907">
<taxonomicName id="1AD84D47FFE089109FBCFBE1FDDAFBE0" ID-CoL="49KG7" authority="Felsche, 1907: 293" authorityName="Felsche" authorityPageNumber="293" authorityYear="1907" box="[189,547,1103,1130]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="feae">
<emphasis id="EFACEAD6FFE089109FBCFBE1FEC4FBE3" box="[189,317,1103,1130]" italics="true" pageId="22" pageNumber="23">Onitis feae</emphasis>
<bibRefCitation id="B9494B35FFE089109E45FBFEFDDAFBE0" author="Felsche C." box="[324,547,1103,1130]" pageId="22" pageNumber="23" pagination="273 - 296" refId="ref29092" refString="Felsche C. 1907. Coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift: 273 - 296." type="book chapter" year="1907">Felsche, 1907: 293</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD6736C4FFE089109FBCFB22FE3EFA9C" blockId="22.[189,1399,1170,1302]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="FDC811EAFFE089109FBCFB22FE3EFA9C" pageId="22" pageNumber="23">
<taxonomicName id="1AD84D47FFE089109FBCFB22FEC6FB26" ID-CoL="49KG7" authorityName="Felsche" authorityYear="1907" box="[189,319,1170,1197]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="feae">
<emphasis id="EFACEAD6FFE089109FBCFB22FEC6FB26" box="[189,319,1170,1197]" italics="true" pageId="22" pageNumber="23">Onitis feae</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFE089109E5EFB22FD3EFB26" author="Boucomont A. &amp; Gillet J." box="[351,711,1170,1197]" page="17" pageId="22" pageNumber="23" year="1921">
<bibRefCitation id="B9494B35FFE089109E5EFB22FD3EFB26" author="Boucomont A. &amp; Gillet J." box="[351,711,1170,1197]" pageId="22" pageNumber="23" pagination="1 - 76" refId="ref27918" refString="Boucomont A. &amp; Gillet J. 1921. Faune entomologique de l'Indochine francaise. Famille Scarabaeidae Laparosticti (Coleopteres). Portail. Saigon 4: 1 - 76." type="journal article" year="1921">Boucomont &amp; Gillet 1921: 17</bibRefCitation>
</treatmentCitation>
, 18 (distribution). —
<treatmentCitation id="5C7910D5FFE089109CCAFB22FB61FB26" author="Arrow G. J." box="[971,1176,1170,1197]" page="397" pageId="22" pageNumber="23" year="1931">
<bibRefCitation id="B9494B35FFE089109CCAFB22FB61FB26" author="Arrow G. J." box="[971,1176,1170,1197]" pageId="22" pageNumber="23" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 397</bibRefCitation>
</treatmentCitation>
(key, description). —
<treatmentCitation id="5C7910D5FFE089109E11FB07FE1FFB5A" author="Janssens A." box="[272,486,1205,1232]" page="75" pageId="22" pageNumber="23" year="1937">
<bibRefCitation id="B9494B35FFE089109E11FB07FE1FFB5A" author="Janssens A." box="[272,486,1205,1232]" pageId="22" pageNumber="23" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 75</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFE089109D96FB04FC94FB45" author="Paulian R." box="[663,877,1205,1232]" page="141" pageId="22" pageNumber="23" year="1945">
<bibRefCitation id="B9494B35FFE089109D96FB04FC94FB45" author="Paulian R." box="[663,877,1205,1232]" pageId="22" pageNumber="23" refId="ref30535" refString="Paulian R. 1945. Faune de l'empire francais: Coleopteres Scarabeides de l'Indochine, Vol. III: 1 - 227. Librairie Larose, Paris." type="book" year="1945">Paulian 1945: 141</bibRefCitation>
</treatmentCitation>
(revision). — Singh
<emphasis id="EFACEAD6FFE089109B65FB07FB58FB45" box="[1124,1185,1205,1231]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2010: 1395 (list). —
<treatmentCitation id="5C7910D5FFE089109E0FFB68FDDBFB79" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[270,546,1241,1268]" page="1037" pageId="22" pageNumber="23" year="2015">
<bibRefCitation id="B9494B35FFE089109E0FFB68FDDBFB79" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[270,546,1241,1268]" pageId="22" pageNumber="23" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFE089109E5CFB6BFE6EFB79" box="[349,407,1241,1267]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(checklist). —
<treatmentCitation id="5C7910D5FFE089109DD7FB68FC53FB79" author="Bezdek A." box="[726,938,1241,1267]" page="179" pageId="22" pageNumber="23" year="2016">
<bibRefCitation id="B9494B35FFE089109DD7FB68FC53FB79" author="Bezdek A." box="[726,938,1241,1267]" pageId="22" pageNumber="23" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFE089109B7CFB68FA8EFB79" author="Schoolmeesters P." box="[1149,1399,1241,1267]" pageId="22" pageNumber="23" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE089109FBCFA8EFEC9FA26" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="DD6736C4FFE089109FBCFA8EFE3FFAD3" blockId="22.[189,454,1343,1369]" box="[189,454,1343,1369]" pageId="22" pageNumber="23">
<heading id="862F81A8FFE089109FBCFA8EFE3FFAD3" bold="true" box="[189,454,1343,1369]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="EFACEAD6FFE089109FBCFA8EFE3FFAD3" bold="true" box="[189,454,1343,1369]" pageId="22" pageNumber="23">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE089109FBCFADEFEC9FA26" blockId="22.[189,1399,1391,1452]" pageId="22" pageNumber="23">
<taxonomicName id="1AD84D47FFE089109FBCFADEFEB8FA03" authorityName="Felsche" authorityYear="1907" box="[189,321,1391,1417]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="feae">
<emphasis id="EFACEAD6FFE089109FBCFADEFEB8FA03" box="[189,321,1391,1417]" italics="true" pageId="22" pageNumber="23">Onitis feae</emphasis>
</taxonomicName>
is distinct in having an elliptical clypeus and a sharply elevated frontal carina which is complete.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE089119FBCFA64FE72FC8C" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="DD6736C4FFE089109FBCFA64FE15FA65" blockId="22.[189,492,1493,1519]" box="[189,492,1493,1519]" pageId="22" pageNumber="23">
<heading id="862F81A8FFE089109FBCFA64FE15FA65" bold="true" box="[189,492,1493,1519]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="EFACEAD6FFE089109FBCFA64FE15FA65" bold="true" box="[189,492,1493,1519]" pageId="22" pageNumber="23">
<typeStatus id="02638866FFE089109FBCFA64FF03FA65" box="[189,250,1493,1519]" pageId="22" pageNumber="23">Type</typeStatus>
material examined
</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE089109FEBF9B4FEBDF995" blockId="22.[189,1398,1541,1640]" box="[234,324,1541,1567]" pageId="22" pageNumber="23">
<emphasis id="EFACEAD6FFE089109FEBF9B4FEBDF995" bold="true" box="[234,324,1541,1567]" pageId="22" pageNumber="23">Cotype</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE089109FBCF99BFC54F9ED" blockId="22.[189,1398,1541,1640]" pageId="22" pageNumber="23">
<materialsCitation id="6DB03C99FFE089109FBCF99BFC50F9ED" collectionCode="SNSD" collectorName="L. Fea. &amp; Coll. C. Felsche" country="Myanmar" elevation="1250" location="MYANMAR" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFE089109FBCF99BFEAFF9CE" box="[189,342,1578,1604]" name="Myanmar" pageId="22" pageNumber="23">MYANMAR</collectingCountry>
<specimenCount id="CBDEFD4DFFE089109E71F99BFE70F9CE" box="[368,393,1578,1604]" count="1" pageId="22" pageNumber="23" type="male"></specimenCount>
; “Asciuii Cheba,
<quantity id="1A209B21FFE089109D5BF99BFD03F9CE" box="[602,762,1578,1604]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="22" pageNumber="23" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">
<elevation id="56F5D1F7FFE089109D5BF99BFD03F9CE" box="[602,762,1578,1604]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="22" pageNumber="23" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200-1300 m</elevation>
</quantity>
,
<collectorName id="702D5312FFE089109C06F99BFCA3F9CE" box="[775,858,1578,1604]" pageId="22" pageNumber="23">L. Fea.</collectorName>
188; Co-Typus (red label);
<collectorName id="702D5312FFE089109BB0F99BFA8AF9CE" box="[1201,1395,1578,1604]" pageId="22" pageNumber="23">Coll. C. Felsche</collectorName>
, Kauf 20, 1918; Staatl. Museum für Tierkunde Dresden;
<collectionCode id="BBC9AE01FFE089109C4FF9FCFC66F9ED" box="[846,927,1613,1639]" pageId="22" pageNumber="23">SNSD</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="DD6736C4FFE089109FEBF921FCD1F920" blockId="22.[189,1399,1680,2025]" box="[234,808,1680,1706]" pageId="22" pageNumber="23">
<emphasis id="EFACEAD6FFE089109FEBF921FD9BF920" bold="true" box="[234,610,1680,1706]" pageId="22" pageNumber="23">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFE089109D70F921FCE6F920" box="[625,799,1680,1706]" count="111" pageId="22" pageNumber="23" type="generic">111 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFE089119FBCF904FED2FE7C" blockId="22.[189,1399,1680,2025]" lastBlockId="23.[189,1399,265,502]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<materialsCitation id="6DB03C99FFE089109FBCF904FC6DF979" collectingDate="2012-10-31" collectionCode="NEHU, AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" location="Northeast Hill University" municipality="Shillong" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Meghalaya">
<collectingCountry id="A5CF7654FFE089109FBCF904FEF7F945" box="[189,270,1717,1743]" name="India" pageId="22" pageNumber="23">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFE089109E35F904FEB4F945" box="[308,333,1717,1743]" count="1" pageId="22" pageNumber="23" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFE089109E5EF904FE1DF945" box="[351,484,1717,1743]" country="India" name="Meghalaya" pageId="22" pageNumber="23">Meghalaya</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE089109EF7F904FDA5F945" box="[502,604,1717,1743]" pageId="22" pageNumber="23">Shillong</collectingMunicipality>
, village near
<location id="D807601FFFE089109C0AF904FBBBF945" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE089109C0AF904FBBBF945" box="[779,1090,1717,1743]" country="India" municipality="Shillong" name="Northeast Hill University" pageId="22" pageNumber="23" stateProvince="Meghalaya">Northeast Hill University</location>
(
<collectionCode id="BBC9AE01FFE089109B56F904FB56F945" box="[1111,1199,1717,1743]" pageId="22" pageNumber="23">NEHU</collectionCode>
);
<date id="A9661004FFE089109BCBF904FA8BF945" box="[1226,1394,1717,1743]" pageId="22" pageNumber="23" value="2012-10-31">
<collectingDate id="B922E9ECFFE089109BCBF904FA8BF945" box="[1226,1394,1717,1743]" pageId="22" pageNumber="23" value="2012-10-31">31 Oct. 2012</collectingDate>
</date>
;
<collectorName id="702D5312FFE089109FBCF969FDB3F978" box="[189,586,1752,1778]" pageId="22" pageNumber="23">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE089109D90F969FD27F979" box="[657,734,1752,1779]" pageId="22" pageNumber="23">AIMB</collectionCode>
/
<collectorName id="702D5312FFE089109DE6F969FCF0F979" box="[743,777,1752,1779]" pageId="22" pageNumber="23">Co</collectorName>
/
<collectorName id="702D5312FFE089109C13F969FCD7F979" box="[786,814,1752,1779]" pageId="22" pageNumber="23">Sc</collectorName>
1000185
</materialsCitation>
<materialsCitation id="6DB03C99FFE089109CB4F969FBB5F8B2" collectingDate="2018-10-19" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" elevation="876" latitude="27.238258" location="Mobu" longLatPrecision="1" longitude="93.5373" pageId="22" pageNumber="23" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE089109CB4F969FC05F978" box="[949,1020,1752,1778]" count="2" pageId="22" pageNumber="23" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE089109B0DF969FBDAF978" box="[1036,1059,1752,1778]" count="1" pageId="22" pageNumber="23" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE089109B35F969FAE0F978" box="[1076,1305,1752,1778]" country="India" name="Arunachal Pradesh" pageId="22" pageNumber="23">Arunachal Pradesh</collectingRegion>
,
<location id="D807601FFFE089109A29F969FA8BF978" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE089109A29F969FA8BF978" box="[1320,1394,1752,1778]" country="India" latitude="27.238258" longLatPrecision="1" longitude="93.5373" name="Mobu" pageId="22" pageNumber="23" stateProvince="Arunachal Pradesh">Mobu</location>
;
<geoCoordinate id="B8EC5003FFE089109FBCF94BFE63F89C" box="[189,410,1786,1814]" degrees="27" direction="north" minutes="14" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="17.7324" value="27.238258">27°14ʹ17.7324ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE089109EACF94BFD93F89C" box="[429,618,1786,1814]" degrees="93" direction="east" minutes="32" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="14.28" value="93.5373">93°32ʹ14.28ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE089109DC3F94AFCA3F89F" box="[706,858,1787,1814]" metricMagnitude="2" metricUnit="m" metricValue="8.76" pageId="22" pageNumber="23" unit="m" value="876.0">
<quantity id="1A209B21FFE089109DC3F94AFCEAF89C" box="[706,787,1787,1814]" metricMagnitude="2" metricUnit="m" metricValue="8.76" pageId="22" pageNumber="23" unit="m" value="876.0">876 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE089109C6EF94AFBE1F89F" box="[879,1048,1787,1813]" pageId="22" pageNumber="23" value="2018-10-19">
<collectingDate id="B922E9ECFFE089109C6EF94AFBE1F89F" box="[879,1048,1787,1813]" pageId="22" pageNumber="23" value="2018-10-19">19 Oct. 2018</collectingDate>
</date>
; hand-picked, mithun dung;
<collectorName id="702D5312FFE089109FBCF8AFFDEBF8B2" box="[189,530,1822,1848]" pageId="22" pageNumber="23">Priyadarsanan Dharma Rajan</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE089109DBEF8AFFCF5F8B2" box="[703,780,1822,1848]" pageId="22" pageNumber="23">AIMB</collectionCode>
/
<collectorName id="702D5312FFE089109C15F8AFFCCFF8B2" box="[788,822,1822,1848]" pageId="22" pageNumber="23">Co</collectorName>
/
<collectorName id="702D5312FFE089109C3EF8AFFCA2F8B2" box="[831,859,1822,1848]" pageId="22" pageNumber="23">Sc</collectorName>
1000186 to 1000188
</materialsCitation>
<materialsCitation id="6DB03C99FFE089109B60F8AFFBD2F8F5" collectingDate="2019-11-14" collectionCode="AIMB" collectorName="Sneha Haridas &amp; Co &amp; Sc" country="India" elevation="317" latitude="28.14344" location="Aalong" longLatPrecision="1" longitude="94.82593" pageId="22" pageNumber="23" specimenCount="11" specimenCount-female="5" specimenCount-male="6" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE089109B60F8AFFB5DF8B3" box="[1121,1188,1822,1849]" count="6" pageId="22" pageNumber="23" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE089109BAEF8AFFB08F8B3" box="[1199,1265,1822,1849]" count="5" pageId="22" pageNumber="23" type="female">5 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE089109BFDF8AFFEE4F8D1" country="India" name="Arunachal Pradesh" pageId="22" pageNumber="23">Arunachal Pradesh</collectingRegion>
,
<location id="D807601FFFE089109E2DF8F0FE7FF8D1" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE089109E2DF8F0FE7FF8D1" box="[300,390,1857,1883]" country="India" latitude="28.14344" longLatPrecision="1" longitude="94.82593" name="Aalong" pageId="22" pageNumber="23" stateProvince="Arunachal Pradesh">Aalong</location>
;
<geoCoordinate id="B8EC5003FFE089109E99F8F1FDAFF8D6" box="[408,598,1856,1884]" degrees="28" direction="north" minutes="8" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="36.384" value="28.14344">28°8ʹ36.384ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE089109D66F8F1FCDAF8D6" box="[615,803,1856,1884]" degrees="94" direction="east" minutes="49" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="33.34" value="94.82593">94°49ʹ33.34ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE089109C79F8F3FBF4F8D1" box="[888,1037,1857,1884]" metricMagnitude="2" metricUnit="m" metricValue="3.17" pageId="22" pageNumber="23" unit="m" value="317.0">
<quantity id="1A209B21FFE089109C79F8F3FC3EF8D6" box="[888,967,1858,1884]" metricMagnitude="2" metricUnit="m" metricValue="3.17" pageId="22" pageNumber="23" unit="m" value="317.0">317 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE089109B21F8F3FB35F8D6" box="[1056,1228,1858,1884]" pageId="22" pageNumber="23" value="2019-11-14">
<collectingDate id="B922E9ECFFE089109B21F8F3FB35F8D6" box="[1056,1228,1858,1884]" pageId="22" pageNumber="23" value="2019-11-14">14 Nov. 2019</collectingDate>
</date>
; hand-picked, mithun dung;
<collectorName id="702D5312FFE089109E61F8D4FDF0F8F5" box="[352,521,1893,1919]" pageId="22" pageNumber="23">Sneha Haridas</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE089109DB9F8D4FCFCF8F5" box="[696,773,1893,1919]" pageId="22" pageNumber="23">AIMB</collectionCode>
/
<collectorName id="702D5312FFE089109C0FF8D4FCC9F8F5" box="[782,816,1893,1919]" pageId="22" pageNumber="23">Co</collectorName>
/
<collectorName id="702D5312FFE089109C38F8D4FCACF8F5" box="[825,853,1893,1919]" pageId="22" pageNumber="23">Sc</collectorName>
1000189-1000199
</materialsCitation>
<materialsCitation id="6DB03C99FFE089109B43F8D4FA8EF84F" collectingDate="2019-11-14" collectionCode="AIMB" collectorName="Sneha Haridas &amp; Co &amp; Sc" country="India" elevation="359" latitude="28.142744" location="Aalong forest" longLatPrecision="1" longitude="94.82671" municipality="West Siang" pageId="22" pageNumber="23" specimenCount="25" specimenCount-female="12" specimenCount-male="13" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE089109B43F8D4FB6DF8F5" box="[1090,1172,1893,1919]" count="13" pageId="22" pageNumber="23" type="male">13 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE089109BA1F8D4FB09F8F5" box="[1184,1264,1893,1919]" count="12" pageId="22" pageNumber="23" type="female">12 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE089109BFDF8D4FEE4F828" country="India" name="Arunachal Pradesh" pageId="22" pageNumber="23">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE089109E2DF839FE4EF828" box="[300,439,1928,1954]" pageId="22" pageNumber="23">West Siang</collectingMunicipality>
,
<location id="D807601FFFE089109EC4F839FD97F828" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE089109EC4F839FD97F828" box="[453,622,1928,1954]" country="India" latitude="28.142744" longLatPrecision="1" longitude="94.82671" municipality="West Siang" name="Aalong forest" pageId="22" pageNumber="23" stateProvince="Arunachal Pradesh">Aalong forest</location>
;
<geoCoordinate id="B8EC5003FFE089109D7CF837FCC3F828" box="[637,826,1926,1954]" degrees="28" direction="north" minutes="8" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="33.882" value="28.142744">28°8ʹ33.882ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE089109C48F837FBD8F828" box="[841,1057,1926,1954]" degrees="94" direction="east" minutes="49" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="36.1848" value="94.82671">94°49ʹ36.1848ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE089109B73F839FAFDF828" box="[1138,1284,1928,1955]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="22" pageNumber="23" unit="m" value="359.0">
<quantity id="1A209B21FFE089109B73F839FB38F828" box="[1138,1217,1928,1955]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="22" pageNumber="23" unit="m" value="359.0">359 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE089109A17F839FF02F84F" pageId="22" pageNumber="23" value="2019-11-14">
<collectingDate id="B922E9ECFFE089109A17F839FF02F84F" pageId="22" pageNumber="23" value="2019-11-14">14 Nov. 2019</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE089109E0AF81DFD98F84C" box="[267,609,1963,1990]" pageId="22" pageNumber="23">cow dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFE089109D70F81AFCE6F84F" box="[625,799,1963,1989]" pageId="22" pageNumber="23">Sneha Haridas</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE089109CE1F81AFBD4F84F" box="[992,1069,1963,1989]" pageId="22" pageNumber="23">AIMB</collectionCode>
/
<collectorName id="702D5312FFE089109B34F81AFBAEF84F" box="[1077,1111,1963,1989]" pageId="22" pageNumber="23">Co</collectorName>
/
<collectorName id="702D5312FFE089109B61F81AFB85F84F" box="[1120,1148,1963,1989]" pageId="22" pageNumber="23">Sc</collectorName>
1000200 to 1000224
</materialsCitation>
<materialsCitation id="6DB03C99FFE089119FCEF87FFE35FECC" collectingDate="2019-11-15" collectionCode="AIMB" collectorName="Sneha Haridas &amp; Co &amp; Sc" country="India" elevation="376" lastPageId="23" lastPageNumber="24" latitude="28.409962" location="Kaying forest" longLatPrecision="1" longitude="94.69764" municipality="West Siang" pageId="22" pageNumber="23" specimenCount="39" specimenCount-female="22" specimenCount-male="17" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE089109FCEF87FFED8F863" box="[207,289,1998,2025]" count="17" pageId="22" pageNumber="23" type="male">17 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE089109E2FF87FFE79F863" box="[302,384,1998,2025]" count="22" pageId="22" pageNumber="23" type="female">22 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE089109E8CF87FFD97F862" box="[397,622,1998,2024]" country="India" name="Arunachal Pradesh" pageId="22" pageNumber="23">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE089109D78F87EFCF9F862" box="[633,768,1998,2025]" pageId="22" pageNumber="23">West Siang</collectingMunicipality>
,
<location id="D807601FFFE089109C0DF87FFC48F862" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE089109C0DF87FFC48F862" box="[780,945,1998,2024]" country="India" latitude="28.409962" longLatPrecision="1" longitude="94.69764" municipality="West Siang" name="Kaying forest" pageId="22" pageNumber="23" stateProvince="Arunachal Pradesh">Kaying forest</location>
;
<geoCoordinate id="B8EC5003FFE089109CBDF87CFB6BF863" box="[956,1170,1997,2025]" degrees="28" direction="north" minutes="24" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="35.8668" value="28.409962">28°24ʹ35.8668ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE089109B9FF87CFA88F863" box="[1182,1393,1997,2025]" degrees="94" direction="east" minutes="41" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="51.4896" value="94.69764">94°41ʹ51.4896ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE189119FFAFEB8FE7FFEA9" box="[251,390,265,291]" metricMagnitude="2" metricUnit="m" metricValue="3.76" pageId="23" pageNumber="24" unit="m" value="376.0">
<quantity id="1A209B21FFE189119FFAFEB8FEBFFEA9" box="[251,326,265,291]" metricMagnitude="2" metricUnit="m" metricValue="3.76" pageId="23" pageNumber="24" unit="m" value="376.0">376 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE189119E95FEB8FDCFFEA9" box="[404,566,265,292]" pageId="23" pageNumber="24" value="2019-11-15">
<collectingDate id="B922E9ECFFE189119E95FEB8FDCFFEA9" box="[404,566,265,292]" pageId="23" pageNumber="24" value="2019-11-15">15 Nov. 2019</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE189119D42FEB8FC75FEA9" box="[579,908,265,291]" pageId="23" pageNumber="24">cow dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFE189119C98FEB8FBBAFEA9" box="[921,1091,265,291]" pageId="23" pageNumber="24">Sneha Haridas</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE189119BF6FEB8FABAFEA9" box="[1271,1347,265,291]" pageId="23" pageNumber="24">AIMB</collectionCode>
/
<collectorName id="702D5312FFE189119A4DFEB8FA94FEA9" box="[1356,1389,265,291]" pageId="23" pageNumber="24">Co</collectorName>
/
<collectorName id="702D5312FFE189119FBCFE9DFF20FECD" box="[189,217,300,327]" pageId="23" pageNumber="24">Sc</collectorName>
1000225 to 1000263
</materialsCitation>
<materialsCitation id="6DB03C99FFE189119EE5FE9DFC59FE07" collectingDate="2019-11-18" collectionCode="AIMB" collectorName="Sneha Haridas &amp; Co &amp; Sc" country="India" county="West Siang" elevation="489" latitude="28.204882" location="Larmuk Pottom forest" longLatPrecision="1" longitude="94.59703" municipality="Darak" pageId="23" pageNumber="24" specimenCount="25" specimenCount-female="15" specimenCount-male="10" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE189119EE5FE9DFDCFFECD" box="[484,566,300,327]" count="10" pageId="23" pageNumber="24" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE189119D43FE9DFD6AFECD" box="[578,659,300,327]" count="15" pageId="23" pageNumber="24" type="female">15 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE189119D9EFE9DFC79FECC" box="[671,896,300,326]" country="India" name="Arunachal Pradesh" pageId="23" pageNumber="24">Arunachal Pradesh</collectingRegion>
,
<collectingCounty id="34064E48FFE189119C8AFE9CFBEBFECC" box="[907,1042,300,327]" pageId="23" pageNumber="24">West Siang</collectingCounty>
,
<collectingMunicipality id="3D03ACBEFFE189119B1DFE9DFB9AFECC" box="[1052,1123,300,326]" pageId="23" pageNumber="24">Darak</collectingMunicipality>
/
<location id="D807601FFFE189119B6CFE9DFA8BFECC" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE189119B6CFE9DFA8BFECC" box="[1133,1394,300,327]" country="India" county="West Siang" latitude="28.204882" longLatPrecision="1" longitude="94.59703" municipality="Darak" name="Larmuk Pottom forest" pageId="23" pageNumber="24" stateProvince="Arunachal Pradesh">Larmuk Pottom forest</location>
;
<geoCoordinate id="B8EC5003FFE189119FBCFEFFFE7FFEE0" box="[189,390,334,362]" degrees="28" direction="north" minutes="12" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="17.568" value="28.204882">28°12ʹ17.568ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE189119E92FEFFFD91FEE0" box="[403,616,334,362]" degrees="94" direction="east" minutes="35" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="49.3008" value="94.59703">94°35ʹ49.3008ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE189119DB5FEFEFCB8FEE3" box="[692,833,335,362]" metricMagnitude="2" metricUnit="m" metricValue="4.89" pageId="23" pageNumber="24" unit="m" value="489.0">
<quantity id="1A209B21FFE189119DB5FEFEFCF9FEE0" box="[692,768,335,362]" metricMagnitude="2" metricUnit="m" metricValue="4.89" pageId="23" pageNumber="24" unit="m" value="489.0">489 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE189119C51FEFEFC0CFEE3" box="[848,1013,335,362]" pageId="23" pageNumber="24" value="2019-11-18">
<collectingDate id="B922E9ECFFE189119C51FEFEFC0CFEE3" box="[848,1013,335,362]" pageId="23" pageNumber="24" value="2019-11-18">18 Nov. 2019</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE189119B03FEFEFA8BFEE0" box="[1026,1394,335,362]" pageId="23" pageNumber="24">mithun dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFE189119FBCFEC3FE9FFE06" box="[189,358,370,396]" pageId="23" pageNumber="24">Sneha Haridas</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE189119D12FEC3FD99FE06" box="[531,608,370,396]" pageId="23" pageNumber="24">AIMB</collectionCode>
/
<collectorName id="702D5312FFE189119D68FEC3FD72FE06" box="[617,651,370,396]" pageId="23" pageNumber="24">Co</collectorName>
/
<collectorName id="702D5312FFE189119D92FEC3FD56FE06" box="[659,687,370,396]" pageId="23" pageNumber="24">Sc</collectorName>
1000264 to 1000288
</materialsCitation>
<materialsCitation id="6DB03C99FFE189119CB7FEC2FEDEFE7C" collectingDate="2019-11-24" collectionCode="AIMB" collectorName="Sneha Haridas &amp; Co &amp; Sc" country="India" elevation="529" latitude="28.265587" location="Yardi Rabe Supse Wildlife Sanctuary" longLatPrecision="1" longitude="94.58278" municipality="Boru Raksap" pageId="23" pageNumber="24" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE189119CB7FEC2FC00FE07" box="[950,1017,371,397]" count="2" pageId="23" pageNumber="24" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE189119B05FEC2FBBFFE07" box="[1028,1094,371,397]" count="2" pageId="23" pageNumber="24" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE189119B50FEC3FACBFE06" box="[1105,1330,370,396]" country="India" name="Arunachal Pradesh" pageId="23" pageNumber="24">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE189119A3DFEC2FEE8FE3A" pageId="23" pageNumber="24">Boru Raksap</collectingMunicipality>
/
<location id="D807601FFFE189119E1BFE27FD25FE3A" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE189119E1BFE27FD25FE3A" box="[282,732,406,432]" country="India" latitude="28.265587" longLatPrecision="1" longitude="94.58278" municipality="Boru Raksap" name="Yardi Rabe Supse Wildlife Sanctuary" pageId="23" pageNumber="24" stateProvince="Arunachal Pradesh">Yardi Rabe Supse Wildlife Sanctuary</location>
;
<geoCoordinate id="B8EC5003FFE189119DE8FE25FC39FE3A" box="[745,960,404,432]" degrees="28" direction="north" minutes="15" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="56.1132" value="28.265587">28°15ʹ56.1132ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE189119CCFFE25FB6CFE3A" box="[974,1173,404,432]" degrees="94" direction="east" minutes="34" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="58.008" value="94.58278">94°34ʹ58.008ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE189119BE3FE27FA89FE3A" box="[1250,1392,406,433]" metricMagnitude="2" metricUnit="m" metricValue="5.29" pageId="23" pageNumber="24" unit="m" value="529.0">
<quantity id="1A209B21FFE189119BE3FE27FAD6FE3A" box="[1250,1327,406,433]" metricMagnitude="2" metricUnit="m" metricValue="5.29" pageId="23" pageNumber="24" unit="m" value="529.0">529 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE189119FBCFE08FE9BFE59" box="[189,354,441,467]" pageId="23" pageNumber="24" value="2019-11-24">
<collectingDate id="B922E9ECFFE189119FBCFE08FE9BFE59" box="[189,354,441,467]" pageId="23" pageNumber="24" value="2019-11-24">24 Nov. 2019</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE189119E71FE08FD18FE59" box="[368,737,441,467]" pageId="23" pageNumber="24">mithun dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFE189119DEFFE08FC63FE59" box="[750,922,441,467]" pageId="23" pageNumber="24">Sneha Haridas</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE189119B52FE08FB59FE59" box="[1107,1184,441,467]" pageId="23" pageNumber="24">AIMB</collectionCode>
/
<collectorName id="702D5312FFE189119BA8FE08FB32FE59" box="[1193,1227,441,467]" pageId="23" pageNumber="24">Co</collectorName>
/
<collectorName id="702D5312FFE189119BD5FE08FB09FE59" box="[1236,1264,441,467]" pageId="23" pageNumber="24">Sc</collectorName>
1000289 to 1000292
</materialsCitation>
.
</paragraph>
<paragraph id="DD6736C4FFE189119FBCFDAFFE0CFDD1" blockId="23.[189,1399,541,603]" pageId="23" pageNumber="24">
<materialsCitation id="6DB03C99FFE189119FBCFDAFFE09FDD1" collectingDate="1919-01-08" collectionCode="BMNH" collectorName="R. V. de Salvaza" country="Laos" location="Xieng Khouang" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="Xiangkhouang">
<collectingCountry id="A5CF7654FFE189119FBCFDAFFEF3FDB2" box="[189,266,542,568]" name="Laos" pageId="23" pageNumber="24">LAOS</collectingCountry>
<specimenCount id="CBDEFD4DFFE189119E2CFDAFFEBDFDB2" box="[301,324,542,568]" count="1" pageId="23" pageNumber="24" type="female"></specimenCount>
; “
<collectingRegion id="1F1CF826FFE189119E63FDAFFDD1FDB2" box="[354,552,542,568]" country="Laos" name="Xiangkhouang" pageId="23" pageNumber="24">Xieng Khouang</collectingRegion>
,
<date id="A9661004FFE189119D39FDAFFD62FDB2" box="[568,667,542,568]" pageId="23" pageNumber="24" value="1919-01-08">
<collectingDate id="B922E9ECFFE189119D39FDAFFD62FDB2" box="[568,667,542,568]" pageId="23" pageNumber="24" value="1919-01-08">8.i.1919</collectingDate>
</date>
;
<collectorName id="702D5312FFE189119DACFDAFFC8DFDB2" box="[685,884,542,568]" pageId="23" pageNumber="24">R.V. de Salvaza</collectorName>
; Indo-China, B.M. 1924-315;
<collectionCode id="BBC9AE01FFE189119BEDFDAFFABDFDB2" box="[1260,1348,542,568]" pageId="23" pageNumber="24">BMNH</collectionCode>
(E) 1237162, Southeast Asia”
</materialsCitation>
.
</paragraph>
<paragraph id="DD6736C4FFE189119FBCFD33FE72FC8C" blockId="23.[189,1399,642,774]" pageId="23" pageNumber="24">
<materialsCitation id="6DB03C99FFE189119FBCFD33FB03FD35" collectionCode="BMNH" collectorName="N. Tawngpeng &amp; J. Coggin Brown" country="Myanmar" elevation="1402" location="Upper Burma" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Shan">
<collectingCountry id="A5CF7654FFE189119FBCFD33FEAFFD16" box="[189,342,642,668]" name="Myanmar" pageId="23" pageNumber="24">MYANMAR</collectingCountry>
<specimenCount id="CBDEFD4DFFE189119E70FD32FE73FD17" box="[369,394,643,669]" count="1" pageId="23" pageNumber="24" type="male"></specimenCount>
; “ex Mus. Calcutta, B.M. 19301. Man Hpat,
<collectorName id="702D5312FFE189119CB3FD32FB9AFD17" box="[946,1123,643,669]" pageId="23" pageNumber="24">N. Tawngpeng</collectorName>
, N.
<collectingRegion id="1F1CF826FFE189119B95FD33FB36FD16" box="[1172,1231,642,668]" country="Myanmar" name="Shan" pageId="23" pageNumber="24">Shan</collectingRegion>
States,
<location id="D807601FFFE189119A2CFD32FEE9FD4A" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE189119A2CFD32FEE9FD4A" country="Myanmar" name="Upper Burma" pageId="23" pageNumber="24" stateProvince="Shan">Upper Burma</location>
, 11-ii-15.,
<quantity id="1A209B21FFE189119E91FD14FE17FD35" box="[400,494,677,703]" metricMagnitude="3" metricUnit="m" metricValue="1.40208" pageId="23" pageNumber="24" unit="ft" value="4600.0">
<elevation id="56F5D1F7FFE189119E91FD14FE17FD35" box="[400,494,677,703]" metricMagnitude="3" metricUnit="m" metricValue="1.40208" pageId="23" pageNumber="24" unit="ft" value="4600.0">4600 ft.</elevation>
</quantity>
;
<collectorName id="702D5312FFE189119EFAFD17FD3EFD4A" box="[507,711,677,704]" pageId="23" pageNumber="24">J. Coggin Brown</collectorName>
; in dung; Southeast Asia;”
<collectionCode id="BBC9AE01FFE189119B06FD17FBA6FD4A" box="[1031,1119,678,704]" pageId="23" pageNumber="24">BMNH</collectionCode>
(E) 1237161
</materialsCitation>
<materialsCitation id="6DB03C99FFE189119A15FD17FE7EFC8C" country="Myanmar" county="Loi" elevation="1420" location="States" municipality="Tawng Kyan" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Shan">
<specimenCount id="CBDEFD4DFFE189119A15FD17FAD2FD4A" box="[1300,1323,678,704]" count="1" pageId="23" pageNumber="24" type="female"></specimenCount>
; “
<collectingCounty id="34064E48FFE189119A44FD14FA8BFD35" box="[1349,1394,677,703]" pageId="23" pageNumber="24">Loi</collectingCounty>
,
<collectingMunicipality id="3D03ACBEFFE189119FBCFD78FEA1FD69" box="[189,344,713,739]" pageId="23" pageNumber="24">Tawng Kyan</collectingMunicipality>
, Tawng, N.
<collectingRegion id="1F1CF826FFE189119EECFD78FDD1FD69" box="[493,552,713,739]" country="Myanmar" name="Shan" pageId="23" pageNumber="24">Shan</collectingRegion>
<location id="D807601FFFE189119D33FD78FD82FD69" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE189119D33FD78FD82FD69" box="[562,635,713,739]" country="Myanmar" county="Loi" municipality="Tawng Kyan" name="States" pageId="23" pageNumber="24" stateProvince="Shan">States</location>
,
<location id="D807601FFFE189119D88FD78FCD5FD69" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE189119D88FD78FCD5FD69" box="[649,812,713,739]" country="Myanmar" county="Loi" municipality="Tawng Kyan" name="Upper Burma" pageId="23" pageNumber="24" stateProvince="Shan">Upper Burma</location>
<specimenCount id="CBDEFD4DFFE189119C4BFD78FC9AFD69" box="[842,867,713,739]" count="1" pageId="23" pageNumber="24" type="male"></specimenCount>
;
<location id="D807601FFFE189119C73FD78FBF0FD69" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE189119C73FD78FBF0FD69" box="[882,1033,713,739]" country="Myanmar" county="Loi" municipality="Tawng Kyan" name="Tawng Peng" pageId="23" pageNumber="24" stateProvince="Shan">Tawng Peng</location>
, Pansai, N.
<collectingRegion id="1F1CF826FFE189119B8EFD78FB35FD69" box="[1167,1228,713,739]" country="Myanmar" name="Shan" pageId="23" pageNumber="24">Shan</collectingRegion>
<location id="D807601FFFE189119BD7FD78FAE6FD69" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE189119BD7FD78FAE6FD69" box="[1238,1311,713,739]" country="Myanmar" county="Loi" municipality="Tawng Kyan" name="States" pageId="23" pageNumber="24" stateProvince="Shan">States</location>
,
<location id="D807601FFFE189119A2CFD78FEE9FC8C" LSID="urn:lsid:plazi:treatment:557187D2FFE089119D82FC59FB66F930:D807601FFFE189119A2CFD78FEE9FC8C" country="Myanmar" county="Loi" municipality="Tawng Kyan" name="Upper Burma" pageId="23" pageNumber="24" stateProvince="Shan">Upper Burma</location>
,
<quantity id="1A209B21FFE189119E1AFD5DFE83FC8C" box="[283,378,748,774]" metricMagnitude="3" metricUnit="m" metricValue="1.420368" pageId="23" pageNumber="24" unit="ft" value="4660.0">
<elevation id="56F5D1F7FFE189119E1AFD5DFE83FC8C" box="[283,378,748,774]" metricMagnitude="3" metricUnit="m" metricValue="1.420368" pageId="23" pageNumber="24" unit="ft" value="4660.0">4660 ft.</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE189119FBCFC9CFEC8F9AD" pageId="23" pageNumber="24" type="description">
<paragraph id="DD6736C4FFE189119FBCFC9CFEB7FCCD" blockId="23.[189,334,813,839]" box="[189,334,813,839]" pageId="23" pageNumber="24">
<heading id="862F81A8FFE189119FBCFC9CFEB7FCCD" bold="true" box="[189,334,813,839]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="EFACEAD6FFE189119FBCFC9CFEB7FCCD" bold="true" box="[189,334,813,839]" pageId="23" pageNumber="24">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE189119FBCFCEDFCBEFCFD" blockId="23.[189,839,860,887]" box="[189,839,860,887]" pageId="23" pageNumber="24">MEASUREMENTS (in mm). TL=1520.5; BW= 8.511.5.</paragraph>
<paragraph id="DD6736C4FFE189119FEBFC2CFED3FC3D" blockId="23.[189,1399,925,1303]" box="[234,298,925,951]" pageId="23" pageNumber="24">
<emphasis id="EFACEAD6FFE189119FEBFC2CFED3FC3D" bold="true" box="[234,298,925,951]" pageId="23" pageNumber="24">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE189119FBCFC71FEA6FA9D" blockId="23.[189,1399,925,1303]" pageId="23" pageNumber="24">Black, subopaque; head, lower surface and legs dark red; pygidium lower surface and fringe of elytra with reddish hairs; body oval, compact, convex; clypeus elliptical, finely, closely rugulose; forehead rugosely punctured with sharply elevated carina; clypeal carina short, transverse, slightly curved; vertex with a short transverse curved carina before carinate posterior margin of head; pronotum strongly, closely punctured, with feeble median groove, base margined, basal foveae very deep, space between them finely rugose; elytra very convex, opaque, finely striate with intervals carinate along middle; pygidium feebly rugulose, thinly clothed with erect, long reddish hairs; metasternum granular, broadly grooved in the middle; abdomen hairy at sides, smooth in the middle; profemur with sharp tooth near middle of front edge; protibia with finely serrate carina beneath and a tooth near base; middle coxa toothed at its posterior end.</paragraph>
<paragraph id="DD6736C4FFE189119FBCFA8EFE63FA15" blockId="23.[189,1398,1343,1439]" pageId="23" pageNumber="24">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFE189119EB6FA8EFDBFFAD3" box="[439,582,1343,1369]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. EH. O. feae Felsche, 1907, AIMB/Co/Sc1000188. IL. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." pageId="23" pageNumber="24">Fig. 11EH</figureCitation>
). LP=2.86, Lp =2.68; BP= 1.25, BpB=1.07, BpT=0.36. Phallobase and parameres almost of same size; parameres straight above and slightly curved below with the tips blunt and rounded.
</paragraph>
<paragraph id="DD6736C4FFE189119FEBFA76FEBCFA6B" blockId="23.[189,1398,1479,1575]" box="[234,325,1479,1505]" pageId="23" pageNumber="24">
<emphasis id="EFACEAD6FFE189119FEBFA76FEBCFA6B" bold="true" box="[234,325,1479,1505]" pageId="23" pageNumber="24">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE189119FBCFA5BFEC8F9AD" blockId="23.[189,1398,1479,1575]" pageId="23" pageNumber="24">Clypeus extended and reflexed anteriorly, slightly notched; no sharp tooth on underside of protibia and profemur.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE189119FBCF9FFFB66F930" pageId="23" pageNumber="24" type="distribution">
<paragraph id="DD6736C4FFE189119FBCF9FFFEAFF9E2" blockId="23.[189,342,1614,1640]" box="[189,342,1614,1640]" pageId="23" pageNumber="24">
<heading id="862F81A8FFE189119FBCF9FFFEAFF9E2" bold="true" box="[189,342,1614,1640]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="EFACEAD6FFE189119FBCF9FFFEAFF9E2" bold="true" box="[189,342,1614,1640]" pageId="23" pageNumber="24">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE189119FBCF9CCFB66F930" blockId="23.[189,1398,1661,1723]" pageId="23" pageNumber="24">
<collectingCountry id="A5CF7654FFE189119FBCF9CCFF07F91D" box="[189,254,1661,1687]" name="India" pageId="23" pageNumber="24">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFE189119E0CF9CCFE99F91D" box="[269,352,1661,1687]" country="India" name="Assam" pageId="23" pageNumber="24">Assam</collectingRegion>
,
<collectingRegion id="1F1CF826FFE189119E71F9CCFDA1F91D" box="[368,600,1661,1687]" country="India" name="Arunachal Pradesh" pageId="23" pageNumber="24">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE189119D68F9CCFD17F91D" box="[617,750,1661,1687]" country="India" name="Meghalaya" pageId="23" pageNumber="24">Meghalaya</collectingRegion>
,
<collectingRegion id="1F1CF826FFE189119C01F9CCFC96F91D" box="[768,879,1661,1687]" country="India" name="Mizoram" pageId="23" pageNumber="24">Mizoram</collectingRegion>
; Other countries:
<collectingCountry id="A5CF7654FFE189119B53F9CCFB65F91D" box="[1106,1180,1661,1687]" name="China" pageId="23" pageNumber="24">China</collectingCountry>
;
<collectingCountry id="A5CF7654FFE189119BAFF9CCFB12F91D" box="[1198,1259,1661,1687]" name="Laos" pageId="23" pageNumber="24">Laos</collectingCountry>
;
<collectingCountry id="A5CF7654FFE189119BFDF9CCFA8BF91D" box="[1276,1394,1661,1687]" name="Myanmar" pageId="23" pageNumber="24">Myanmar</collectingCountry>
;
<collectingCountry id="A5CF7654FFE189119FBCF911FEDCF930" box="[189,293,1696,1722]" name="Thailand" pageId="23" pageNumber="24">Thailand</collectingCountry>
(Singh
<emphasis id="EFACEAD6FFE189119E83F910FE44F930" box="[386,445,1696,1722]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2010;
<bibRefCitation id="B9494B35FFE189119D0FF911FD24F931" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[526,733,1696,1723]" pageId="23" pageNumber="24" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFE189119D5CF910FD61F930" box="[605,664,1696,1722]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE189119DE8F911FC7EF930" author="Bezdek A." box="[745,903,1696,1722]" pageId="23" pageNumber="24" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE189119C92F911FB6BF930" author="Schoolmeesters P." box="[915,1170,1696,1722]" pageId="23" pageNumber="24" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,375 @@
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<mods:title id="09572AA176841DD5E1737D83E972B637">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<subSubSection id="95C2654FFFE189119D58F8B1FC23F890" box="[601,986,1791,1818]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="DD6736C4FFE189119D58F8B1FC23F890" blockId="23.[601,986,1791,1853]" box="[601,986,1791,1818]" pageId="23" pageNumber="24">
<heading id="862F81A8FFE189119D58F8B1FC23F890" box="[601,986,1791,1818]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<taxonomicName id="1AD84D47FFE189119D58F8B1FC23F890" ID-CoL="49KGS" authority="(Pallas, 1771)" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[601,986,1791,1818]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFE189119D58F8B1FCC9F890" bold="true" box="[601,816,1791,1818]" italics="true" pageId="23" pageNumber="24">Onitis humerosus</emphasis>
(
<bibRefCitation id="B9494B35FFE189119C3EF8B1FC2BF890" author="Pallas P. S." box="[831,978,1792,1818]" pageId="23" pageNumber="24" refId="ref30468" refString="Pallas P. S. 1771. Reise durch verschiedene Provinzen des Russischen Reiches in den Jahren 1768 - 1774 I: 1 - 504 (453 - 504). Petersburg Akademische Buchhandlung, St Petersburg." type="book" year="1771">Pallas, 1771</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE1891E9DCFF892FCF7FEE4" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
<paragraph id="DD6736C4FFE189119DCFF892FC9CF8B7" blockId="23.[601,986,1791,1853]" box="[718,869,1827,1853]" pageId="23" pageNumber="24">
<figureCitation id="45E32A41FFE189119DCFF892FCC2F8B7" box="[718,827,1827,1853]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="23" pageNumber="24">Figs 10B</figureCitation>
,
<figureCitation id="45E32A41FFE189119C49F892FC9CF8B7" box="[840,869,1827,1853]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="23" pageNumber="24">18</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFE189119FBCF8D5FC5CF8F4" blockId="23.[189,933,1892,2025]" box="[189,933,1892,1919]" pageId="23" pageNumber="24">
<emphasis id="EFACEAD6FFE189119FBCF8D5FE32F8F4" box="[189,459,1892,1918]" italics="true" pageId="23" pageNumber="24">Scarabaeushumerosus</emphasis>
<bibRefCitation id="B9494B35FFE189119ED2F8D5FD64F8F4" author="Pallas P. S." box="[467,669,1892,1919]" pageId="23" pageNumber="24" refId="ref30468" refString="Pallas P. S. 1771. Reise durch verschiedene Provinzen des Russischen Reiches in den Jahren 1768 - 1774 I: 1 - 504 (453 - 504). Petersburg Akademische Buchhandlung, St Petersburg." type="book" year="1771">Pallas, 1771: 462</bibRefCitation>
(original description).
</paragraph>
<paragraph id="DD6736C4FFE189119FBCF839FD81F828" blockId="23.[189,933,1892,2025]" box="[189,632,1928,1954]" pageId="23" pageNumber="24">
<emphasis id="EFACEAD6FFE189119FBCF839FE45F828" box="[189,444,1928,1954]" italics="true" pageId="23" pageNumber="24">Scarabaeusmenalcas</emphasis>
<bibRefCitation id="B9494B35FFE189119EC2F839FD88F828" author="Pallas P. S." box="[451,625,1928,1954]" pageId="23" pageNumber="24" refId="ref30506" refString="Pallas P. S. 1781. Icones Insectorum praesertim Rossiae Siberiaeque pecularium quae collegit et Descriptionibus illustravit: 1 - 96. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1781">Pallas, 1781: 4</bibRefCitation>
.
</paragraph>
<paragraph id="DD6736C4FFE189119FBCF81AFD92F84F" blockId="23.[189,933,1892,2025]" box="[189,619,1963,1989]" pageId="23" pageNumber="24">
<emphasis id="EFACEAD6FFE189119FBCF81AFE7BF84F" box="[189,386,1963,1989]" italics="true" pageId="23" pageNumber="24">Onitischevrolati</emphasis>
Lucas 1846: pl. 23.
</paragraph>
<paragraph id="DD6736C4FFE189119FBCF87FFD03F863" blockId="23.[189,933,1892,2025]" box="[189,762,1998,2025]" pageId="23" pageNumber="24">
<emphasis id="EFACEAD6FFE189119FBCF87FFDF9F862" box="[189,512,1998,2024]" italics="true" pageId="23" pageNumber="24">Onitishumerosusviolaceus</emphasis>
<bibRefCitation id="B9494B35FFE189119D09F87FFD0AF862" author="Sahlberg J." box="[520,755,1998,2024]" pageId="23" pageNumber="24" pagination="113 - 128" refId="ref30971" refString="Sahlberg J. 1913. Coleoptera mediterranea orientalia, quae Aegypto, Palaestina, Syria, Caramania atque in Anatolia occidentali anno 1904 (Scarabaeidae). Ofversight af Finska Vetenkaps-Societatens Forhandlingar. Helsingfors 55 (19): 113 - 128." type="journal article" year="1913">Sahlberg, 1913: 119</bibRefCitation>
.
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFEB8FCC4FEAE" blockId="24.[189,858,265,366]" box="[189,829,265,294]" pageId="24" pageNumber="25">
<taxonomicName id="1AD84D47FFEE891E9FBCFEB8FDEBFEA9" authorityName="Mikšić" authorityYear="1950" box="[189,530,265,291]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="humerosus" subSpecies="viridipenni">
<emphasis id="EFACEAD6FFEE891E9FBCFEB8FDEBFEA9" box="[189,530,265,291]" italics="true" pageId="24" pageNumber="25">Onitis humerosus viridipenni</emphasis>
</taxonomicName>
sMikŠić, 1950: 165167.
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFE9FFCA3FEC3" blockId="24.[189,858,265,366]" box="[189,858,302,331]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FBCFE9FFDC2FEC2" box="[189,571,302,328]" italics="true" pageId="24" pageNumber="25">Onitishumerosusviolaceipennis</emphasis>
<bibRefCitation id="B9494B35FFEE891E9D43FE9FFCAFFEC3" author="Miksic, R." box="[578,854,302,331]" pageId="24" pageNumber="25" pagination="165 - 167" refId="ref30176" refString="Miksic, R. (1950). Biljeske o Scarabaeidama palearkticke faune. Glasnik Bioloski Sekcije Hrvatkog Privredno Drustvo Zagreb (2 - B) 2 - 3: 165 - 167." type="book chapter" year="1950">MikŠić, 1950: 165167</bibRefCitation>
.
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFEE5FCF7FEE4" blockId="24.[189,858,265,366]" box="[189,782,340,366]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FBCFEE5FDE5FEE4" box="[189,540,340,366]" italics="true" pageId="24" pageNumber="25">Onitishumerosuscupripennis</emphasis>
<bibRefCitation id="B9494B35FFEE891E9D22FEE5FCF2FEE4" author="Balthasar V." box="[547,779,340,366]" pageId="24" pageNumber="25" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar, 1963: 48</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEE891E9FBCFE29FC0EFD32" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="DD6736C4FFEE891E9FBCFE29FC0EFD32" blockId="24.[189,1399,407,697]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="FDC811EAFFEE891E9FBCFE29FC0EFD32" pageId="24" pageNumber="25">
<taxonomicName id="1AD84D47FFEE891E9FBCFE29FE31FE38" ID-CoL="4V269" authorityName="Pallas" authorityYear="1771" box="[189,456,408,434]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFEE891E9FBCFE29FE31FE38" box="[189,456,408,434]" italics="true" pageId="24" pageNumber="25">Scarabaeus humerosus</emphasis>
</taxonomicName>
Castelnau 1840: 89 (natural history). —
<treatmentCitation id="5C7910D5FFEE891E9CD6FE29FB36FE38" author="Lansberge G." box="[983,1231,407,434]" page="116" pageId="24" pageNumber="25" year="1875">
<bibRefCitation id="B9494B35FFEE891E9CD6FE29FB36FE38" author="Lansberge G." box="[983,1231,407,434]" pageId="24" pageNumber="25" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge 1875: 116</bibRefCitation>
</treatmentCitation>
(monograph). —
<treatmentCitation id="5C7910D5FFEE891E9E10FE0CFE2FFE5D" author="Bedel L." box="[273,470,445,471]" page="268" pageId="24" pageNumber="25" year="1892">
<bibRefCitation id="B9494B35FFEE891E9E10FE0CFE2FFE5D" author="Bedel L." box="[273,470,445,471]" pageId="24" pageNumber="25" pagination="282 - 288" refId="ref27264" refString="Bedel L. 1892. Revision des Scarabaeus palearctiques. L'Abeille, Journal d'Entomologie 27: 282 - 288." type="journal article" year="1892">Bedel 1892: 268</bibRefCitation>
</treatmentCitation>
, 271 (revision). —
<treatmentCitation id="5C7910D5FFEE891E9DBFFE0CFC84FE5D" author="Reitter E." box="[702,893,445,471]" page="97" pageId="24" pageNumber="25" year="1892">
<bibRefCitation id="B9494B35FFEE891E9DBFFE0CFC84FE5D" author="Reitter E." box="[702,893,445,471]" pageId="24" pageNumber="25" refId="ref30798" refString="Reitter E. 1892. Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des Naturforschenden Vereines in Brunn. https: // www. zobodat. at / pdf / Verh-naturf-Ver-Bruenn _ 31 _ 0003 - 0109. pdf" type="book" year="1892">Reitter 1892: 97</bibRefCitation>
</treatmentCitation>
(keys). —
<treatmentCitation id="5C7910D5FFEE891E9B05FE0CFB37FE5D" author="Arrow G. J." box="[1028,1230,445,471]" page="390" pageId="24" pageNumber="25" year="1931">
<bibRefCitation id="B9494B35FFEE891E9B05FE0CFB37FE5D" author="Arrow G. J." box="[1028,1230,445,471]" pageId="24" pageNumber="25" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 390</bibRefCitation>
</treatmentCitation>
(identification key, description). —
<treatmentCitation id="5C7910D5FFEE891E9EE4FE52FD5AFE77" author="Porta A." box="[485,675,483,509]" page="414" pageId="24" pageNumber="25" year="1932">
<bibRefCitation id="B9494B35FFEE891E9EE4FE52FD5AFE77" author="Porta A." box="[485,675,483,509]" pageId="24" pageNumber="25" refId="ref30657" refString="Porta A. 1932. Fauna Coleopterorum Italica, Vol. II: Scarabaeidae: 373 - 454. Stabilimento Tipografico piancentino, Piacenza." type="book" year="1932">Porta 1932: 414</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFEE891E9C7CFE52FBA5FE77" author="Balthasar V." box="[893,1116,482,509]" page="91" pageId="24" pageNumber="25" year="1935">
<bibRefCitation id="B9494B35FFEE891E9C7CFE52FBA5FE77" author="Balthasar V." box="[893,1116,482,509]" pageId="24" pageNumber="25" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 91</bibRefCitation>
</treatmentCitation>
(monograph); 1963: 47 (monograph).—
<treatmentCitation id="5C7910D5FFEE891E9EADFDB8FD8CFDA8" author="Janssens A." box="[428,629,520,547]" page="91" pageId="24" pageNumber="25" year="1937">
<bibRefCitation id="B9494B35FFEE891E9EADFDB8FD8CFDA8" author="Janssens A." box="[428,629,520,547]" pageId="24" pageNumber="25" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 91</bibRefCitation>
</treatmentCitation>
(revision). —
<bibRefCitation id="B9494B35FFEE891E9C15FDB9FBC9FDA8" author="Hashmi A. &amp; Tashfeen A." box="[788,1072,520,546]" pageId="24" pageNumber="25" pagination="133 - 170" refId="ref29508" refString="Hashmi A. &amp; Tashfeen A. 1992. Coleoptera of Pakistan. Proceedings of Pakistan Congress of Zoology 12: 133 - 170." type="journal article" year="1992">Hashmi &amp;Tashfeen 1992</bibRefCitation>
(distribution).—
<treatmentCitation id="5C7910D5FFEE891E9BF4FDB9FEECFDC2" author="Awal M. M." page="170" pageId="24" pageNumber="25" year="2006">
<bibRefCitation id="B9494B35FFEE891E9BF4FDB9FEECFDC2" author="Awal M. M." pageId="24" pageNumber="25" pagination="163 - 172" refId="ref27017" refString="Awal M. M. 2006. Preliminary studies on Scarabaeoidea (Coleoptera) fauna of Razavi Khorasan province of Iran. Turkiye Entomoloji Dergisi 30 (3): 163 - 172. Available from: https: // dergipark. org. tr / tr / download / article-file / 64989 [accessed 27 Nov. 2019]" type="journal article" year="2006">Awal 2006: 170</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFEE891E9EE9FD9FFD74FDC2" author="Bai M. &amp; Yang X. &amp; Youwei Z." box="[488,653,557,584]" page="391" pageId="24" pageNumber="25" year="2006">
<bibRefCitation id="B9494B35FFEE891E9EE9FD9FFD74FDC2" author="Bai M. &amp; Yang X. &amp; Youwei Z." box="[488,653,557,584]" pageId="24" pageNumber="25" pagination="389 - 395" refId="ref27077" refString="Bai M., Yang X. &amp; Youwei Z. 2006. A key to species of the genus Onitis Fabricius (Coleoptera: Scarabaeidae: Scarabaeinae) from China, with the description of a new species and a new record for China. Proceedings of the Entomological Society of Washington 108: 389 - 395. Available from: https: // www. biodiversitylibrary. org / part / 69133 [accessed 27 nov. 2019]" type="journal article" year="2006">Bai 2006: 391</bibRefCitation>
</treatmentCitation>
(identification key). —
<treatmentCitation id="5C7910D5FFEE891E9CA7FD9FFB22FDC2" author="Bezdek A. &amp; Krell F. - T." box="[934,1243,557,584]" page="159" pageId="24" pageNumber="25" year="2006">
<bibRefCitation id="B9494B35FFEE891E9CA7FD9FFB22FDC2" author="Bezdek A. &amp; Krell F. - T." box="[934,1243,557,584]" pageId="24" pageNumber="25" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution, synonyms). —
<treatmentCitation id="5C7910D5FFEE891E9E9AFDE2FD44FDE4" author="Bunalski M. &amp; Samin N. &amp; Ghahari H. &amp; Hawkeswood T. J." box="[411,701,595,622]" page="156" pageId="24" pageNumber="25" year="2014">
<bibRefCitation id="B9494B35FFEE891E9E9AFDE2FD44FDE4" author="Bunalski M. &amp; Samin N. &amp; Ghahari H. &amp; Hawkeswood T. J." box="[411,701,595,622]" pageId="24" pageNumber="25" pagination="141 - 170" refId="ref28015" refString="Bunalski M., Samin N., Ghahari H. &amp; Hawkeswood T. J. 2014. Contributions to the knowledge the scarab beetles of Golestan province, Northern Iran with checklist of Iranian Scarabaeoidea (Coleoptera). Polish Journal of Entomology 83: 141 - 170. https: // doi. org / 10.2478 / pjen- 2014 - 0011" type="journal article" year="2014">
Bunalski
<emphasis id="EFACEAD6FFEE891E9D09FDE5FDBBFDE7" box="[520,578,595,621]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2014: 156
</bibRefCitation>
</treatmentCitation>
(list of species). — Siddhiqui
<emphasis id="EFACEAD6FFEE891E9B1CFDE5FBAEFDE7" box="[1053,1111,595,621]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2014: 300 (distribution). —
<treatmentCitation id="5C7910D5FFEE891E9E0FFDC9FE25FD19" author="Bezdek A." box="[270,476,632,659]" page="179" pageId="24" pageNumber="25" year="2016">
<bibRefCitation id="B9494B35FFEE891E9E0FFDC9FE25FD19" author="Bezdek A." box="[270,476,632,659]" pageId="24" pageNumber="25" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution, synonyms). —
<treatmentCitation id="5C7910D5FFEE891E9C2BFDC9FBC1FD18" author="Bunalski M. &amp; Samin N. &amp; Ghahari H." box="[810,1080,632,658]" page="36" pageId="24" pageNumber="25" year="2016">
<bibRefCitation id="B9494B35FFEE891E9C2BFDC9FBC1FD18" author="Bunalski M. &amp; Samin N. &amp; Ghahari H." box="[810,1080,632,658]" pageId="24" pageNumber="25" pagination="31 - 40" refId="ref28079" refString="Bunalski M., Samin N. &amp; Ghahari H. 2016. A contribution to the faunistic study of Scarabaeoidea (Coleoptera) from Mazandaran Province, northern Iran. Wiadomosci Entomologiczne 35 (1): 31 - 40. Available from: https: // sparrow. up. poznan. pl / pte / we / 2016 / 35 _ 3 _ Bunalski _ i _ in. pdf [Accessed 15 Nov. 2018]" type="journal article" year="2016">
Bunalski
<emphasis id="EFACEAD6FFEE891E9C94FDC8FC37FD18" box="[917,974,632,658]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2016: 36
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFEE891E9A04FDC9FEA4FD33" author="Montreuil O." page="265" pageId="24" pageNumber="25" year="2017">
<bibRefCitation id="B9494B35FFEE891E9A04FDC9FEA4FD33" author="Montreuil O." pageId="24" pageNumber="25" pagination="259 - 277" refId="ref30308" refString="Montreuil O. 2017. New records of Geotrupidae and Scarabeidae dung beetles (Insecta, Coleoptera) from Iran. Journal of Entomological Society of Iran 36 (4): 259 - 277." type="journal article" year="2017">Montreuil 2017: 265</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFEE891E9D2BFD2FFCE2FD32" author="Schoolmeesters P." box="[554,795,670,696]" pageId="24" pageNumber="25" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEE891E9FBCFD50FE3AFCCD" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="DD6736C4FFEE891E9FBCFD50FE3FFD71" blockId="24.[189,1398,737,839]" box="[189,454,737,763]" pageId="24" pageNumber="25">
<heading id="862F81A8FFEE891E9FBCFD50FE3FFD71" bold="true" box="[189,454,737,763]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="EFACEAD6FFEE891E9FBCFD50FE3FFD71" bold="true" box="[189,454,737,763]" pageId="24" pageNumber="25">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFCB6FE3AFCCD" blockId="24.[189,1398,737,839]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FBCFCB6FE74FCAB" box="[189,397,775,801]" italics="true" pageId="24" pageNumber="25">Onitishumerosus</emphasis>
is a very distinct species having bright yellow elytra with longitudinal green lines upon elevated sutures.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEE891E9FBCFCC0FED2FB6A" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="DD6736C4FFEE891E9FBCFCC0FE15FC00" blockId="24.[189,492,880,907]" box="[189,492,880,907]" pageId="24" pageNumber="25">
<heading id="862F81A8FFEE891E9FBCFCC0FE15FC00" bold="true" box="[189,492,880,907]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="EFACEAD6FFEE891E9FBCFCC0FE15FC00" bold="true" box="[189,492,880,907]" pageId="24" pageNumber="25">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEE891E9FEBFC10FDFFFC31" blockId="24.[189,1398,929,1068]" box="[234,518,929,956]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FEBFC10FEA3FC31" bold="true" box="[234,346,929,955]" pageId="24" pageNumber="25">
<typeStatus id="02638866FFEE891E9FEBFC10FEA3FC31" box="[234,346,929,955]" pageId="24" pageNumber="25" type="syntype">Syntypes</typeStatus>
</emphasis>
(
<specimenCount id="CBDEFD4DFFEE891E9E6AFC10FE07FC31" box="[363,510,929,956]" count="5" pageId="24" pageNumber="25" type="generic">5 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFC76FB16FBA6" blockId="24.[189,1398,929,1068]" pageId="24" pageNumber="25">
<collectingCountry id="A5CF7654FFEE891E9FBCFC76FEDDFC6B" box="[189,292,967,993]" name="Russia" pageId="24" pageNumber="25">RUSSIA</collectingCountry>
<specimenCount id="CBDEFD4DFFEE891E9E3CFC76FE7BFC6B" box="[317,386,967,993]" count="3" pageId="24" pageNumber="25" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFEE891E9E8EFC76FE2BFC6B" box="[399,466,967,993]" count="2" pageId="24" pageNumber="25" type="female">2 ♀♀</specimenCount>
; “
<taxonomicName id="1AD84D47FFEE891E9EEFFC76FD17FC6B" authorityName="Pallas" authorityYear="1781" box="[494,750,967,993]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="menalcas">
<emphasis id="EFACEAD6FFEE891E9EEFFC76FD17FC6B" box="[494,750,967,993]" italics="true" pageId="24" pageNumber="25">Scarabaeus menalcas</emphasis>
</taxonomicName>
;
<collectionCode id="BBC9AE01FFEE891E9DFCFC76FCCFFC6B" box="[765,822,967,993]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="24" pageNumber="25" type="Museum">MfN</collectionCode>
Janssens det. as
<taxonomicName id="1AD84D47FFEE891E9CFFFC76FB32FC6B" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[1022,1227,967,993]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFEE891E9CFFFC76FB32FC6B" box="[1022,1227,967,993]" italics="true" pageId="24" pageNumber="25">Onitis humerosus</emphasis>
</taxonomicName>
in 1938; Hist.- Coll. (
<taxonomicName id="1AD84D47FFEE891E9E04FC5DFE71FB8C" authorityName="Linnaeus" authorityYear="1758" box="[261,392,1004,1030]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
), Nr. 26753,
<emphasis id="EFACEAD6FFEE891E9D20FC5DFD27FB8C" box="[545,734,1004,1030]" italics="true" pageId="24" pageNumber="25">Onitismenalcas</emphasis>
<bibRefCitation id="B9494B35FFEE891E9DE4FC5DFC8DFB8C" author="Pallas P. S." box="[741,884,1004,1030]" pageId="24" pageNumber="25" refId="ref30506" refString="Pallas P. S. 1781. Icones Insectorum praesertim Rossiae Siberiaeque pecularium quae collegit et Descriptionibus illustravit: 1 - 96. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1781">Pallas, 1781</bibRefCitation>
, Russia austr., Bob., Zool. Mus. Berlin; Red Label:
<typeStatus id="02638866FFEE891E9E0EFBA3FE53FBA6" box="[271,426,1042,1068]" pageId="24" pageNumber="25" type="syntype">SYNTYPUS</typeStatus>
,
<taxonomicName id="1AD84D47FFEE891E9EB4FBA3FCB2FBA6" authority="Pallas, 1781" authorityName="Pallas" authorityYear="1781" box="[437,843,1042,1068]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="menalcas">
<emphasis id="EFACEAD6FFEE891E9EB4FBA3FD4CFBA6" box="[437,693,1042,1068]" italics="true" pageId="24" pageNumber="25">Scarabaeus menalcas</emphasis>
<bibRefCitation id="B9494B35FFEE891E9DBDFBA3FCB2FBA6" author="Pallas P. S." box="[700,843,1042,1068]" pageId="24" pageNumber="25" refId="ref30506" refString="Pallas P. S. 1781. Icones Insectorum praesertim Rossiae Siberiaeque pecularium quae collegit et Descriptionibus illustravit: 1 - 96. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1781">Pallas, 1781</bibRefCitation>
</taxonomicName>
, labelled by MNHUB 2007”;
<collectionCode id="BBC9AE01FFEE891E9BAFFBA3FB12FBA6" box="[1198,1259,1042,1068]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="24" pageNumber="25" type="Museum">MfN</collectionCode>
.
</paragraph>
<paragraph id="DD6736C4FFEE891E9FEBFBE4FCF5FBE5" blockId="24.[189,1399,1109,1248]" box="[234,780,1109,1136]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FEBFBE4FD98FBE5" bold="true" box="[234,609,1109,1135]" pageId="24" pageNumber="25">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFEE891E9D70FBE7FCFDFBE5" box="[625,772,1109,1136]" count="7" pageId="24" pageNumber="25" type="generic">7 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFBCAFED2FB6A" blockId="24.[189,1399,1109,1248]" pageId="24" pageNumber="25">
<materialsCitation id="6DB03C99FFEE891E9FBCFBCAFE15FB31" collectionCode="BMNH" collectorName="G. R. Crotch." country="Israel" location="Jerusalem" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Jerusalem">
<collectingCountry id="A5CF7654FFEE891E9FBCFBCAFEDAFB1F" box="[189,291,1147,1173]" name="Israel" pageId="24" pageNumber="25">ISRAEL</collectingCountry>
<specimenCount id="CBDEFD4DFFEE891E9E42FBCAFEA5FB1F" box="[323,348,1147,1173]" count="1" pageId="24" pageNumber="25" type="male"></specimenCount>
; “
<taxonomicName id="1AD84D47FFEE891E9E78FBCAFD1FFB1F" authority="Pallas, Mid." authorityName="Pallas, Mid." box="[377,742,1147,1173]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFEE891E9E78FBCAFDB3FB1F" box="[377,586,1147,1173]" italics="true" pageId="24" pageNumber="25">Onitis humerosus</emphasis>
Pallas, Mid.
</taxonomicName>
East;
<collectingRegion id="1F1CF826FFEE891E9C37FBCAFC57FB1F" box="[822,942,1147,1173]" country="Israel" name="Jerusalem" pageId="24" pageNumber="25">Jerusalem</collectingRegion>
,
<collectorName id="702D5312FFEE891E9CBFFBCAFBA2FB1F" box="[958,1115,1147,1173]" pageId="24" pageNumber="25">G.R. Crotch.</collectorName>
, Nevinson Coll., 1918- 14”;
<collectionCode id="BBC9AE01FFEE891E9FF6FB10FEB6FB31" box="[247,335,1185,1211]" pageId="24" pageNumber="25">BMNH</collectionCode>
(E) 1237143
</materialsCitation>
<materialsCitation id="6DB03C99FFEE891E9D07FB10FEDEFB6A" collectionCode="BMNH" country="Israel" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFEE891E9D07FB10FDE4FB31" box="[518,541,1185,1211]" count="1" pageId="24" pageNumber="25" type="female"></specimenCount>
; “
<emphasis id="EFACEAD6FFEE891E9D38FB11FCFEFB30" box="[569,775,1184,1210]" italics="true" pageId="24" pageNumber="25">Onitishumerosus</emphasis>
Pallas, Mid. East; Pascoe, Coll. 9360;”
<collectionCode id="BBC9AE01FFEE891E9BF0FB10FAB0FB31" box="[1265,1353,1185,1211]" pageId="24" pageNumber="25">BMNH</collectionCode>
(E) 1237144
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEE891E9FBCFAB8FB70F863" pageId="24" pageNumber="25" type="description">
<paragraph id="DD6736C4FFEE891E9FBCFAB8FEB7FAA9" blockId="24.[189,334,1289,1315]" box="[189,334,1289,1315]" pageId="24" pageNumber="25">
<heading id="862F81A8FFEE891E9FBCFAB8FEB7FAA9" bold="true" box="[189,334,1289,1315]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="EFACEAD6FFEE891E9FBCFAB8FEB7FAA9" bold="true" box="[189,334,1289,1315]" pageId="24" pageNumber="25">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFA8AFCCBFADF" blockId="24.[189,818,1338,1365]" box="[189,818,1338,1365]" pageId="24" pageNumber="25">MEASUREMENTS (in mm). TL=12.518.5; BW=710.</paragraph>
<paragraph id="DD6736C4FFEE891E9FEBFACFFED3FA12" blockId="24.[189,1398,1406,1882]" box="[234,298,1406,1432]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FEBFACFFED3FA12" bold="true" box="[234,298,1406,1432]" pageId="24" pageNumber="25">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCFA15FE6AF8D0" blockId="24.[189,1398,1406,1882]" pageId="24" pageNumber="25">
Bright metallic green, coppery, indigo-blue or violet, the elytra bright yellow with four narrow longitudinal green lines on the elevated 1
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, 3
<superScript id="2AAD9B8CFFEE891E9DF1FA79FD06FA5C" attach="left" box="[752,767,1480,1494]" fontSize="6" pageId="24" pageNumber="25">rd</superScript>
and 5
<superScript id="2AAD9B8CFFEE891E9C4FFA79FCA5FA5C" attach="left" box="[846,860,1480,1494]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
intervals, pale intervals with light metallic lustre; body oval, convex, not very shining; head closely covered with fine tubercles; clypeus parabolic, rugose with yellow setae, clypeal carina curved, elevated, near to clypeal margin; frontal carina entire with a tubercle behind; pronotum metallic green, unevenly punctured posteriorly, front angles granular, basal foveae deep, close with granules anteriorly; elytra finely striate with sutural, 3
<superScript id="2AAD9B8CFFEE891E9E2EF935FEC7F918" attach="left" box="[303,318,1668,1682]" fontSize="6" pageId="24" pageNumber="25">rd</superScript>
and 5
<superScript id="2AAD9B8CFFEE891E9E89F935FE6FF918" attach="left" box="[392,406,1668,1682]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
intervals elevated, shining, finely punctured, the other intervals opaque, scarcely punctured; pygidium shining, finely, sparsely granular; metasternum with a longitudinal groove, metasternal shield broad, granular and thinly covered with hairs anteriorly, sparingly punctured behind, sides of metasternum closely granular and hairy; protibia elongated, slender with a strong tooth at the middle of the lower surface; hind femur with a very strong hooked tooth near the middle of its upper edge.
</paragraph>
<paragraph id="DD6736C4FFEE891E9FEBF835FEBCF814" blockId="24.[189,1398,1924,2025]" box="[234,325,1924,1950]" pageId="24" pageNumber="25">
<emphasis id="EFACEAD6FFEE891E9FEBF835FEBCF814" bold="true" box="[234,325,1924,1950]" pageId="24" pageNumber="25">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFEE891E9FBCF818FB70F863" blockId="24.[189,1398,1924,2025]" pageId="24" pageNumber="25">Clypeal carina long, curved, space between the carina and front margin covered with long yellow, closelying hairs; legs short, stout, protibia with four strong teeth and an articulated spur.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEF891F9FBCFEB9FD76FE31" pageId="25" pageNumber="26" type="distribution">
<paragraph id="DD6736C4FFEF891F9FBCFEB9FEAFFEA8" blockId="25.[189,342,264,290]" box="[189,342,264,290]" pageId="25" pageNumber="26">
<heading id="862F81A8FFEF891F9FBCFEB9FEAFFEA8" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="EFACEAD6FFEF891F9FBCFEB9FEAFFEA8" bold="true" box="[189,342,264,290]" pageId="25" pageNumber="26">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEF891F9FBCFE86FD76FE31" blockId="25.[189,1398,311,443]" pageId="25" pageNumber="26">
<collectingCountry id="A5CF7654FFEF891F9FBCFE86FEA9FEDB" box="[189,336,311,337]" name="Afghanistan" pageId="25" pageNumber="26">Afghanistan</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9E5BFE86FE42FEDB" box="[346,443,311,337]" name="Albania" pageId="25" pageNumber="26">Albania</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9EC4FE86FDC9FEDB" box="[453,560,311,337]" name="Armenia" pageId="25" pageNumber="26">Armenia</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9D3AFE86FD7CFEDB" box="[571,645,311,337]" name="China" pageId="25" pageNumber="26">China</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9D90FE86FD10FEDB" box="[657,745,311,337]" name="Cyprus" pageId="25" pageNumber="26">Cyprus</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9DF4FE86FCAEFEDB" box="[757,855,311,337]" name="Georgia" pageId="25" pageNumber="26">Georgia</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9C63FE89FC6CFED8" box="[866,917,312,338]" name="Iran" pageId="25" pageNumber="26">Iran</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9CA0FE86FC1FFEDB" box="[929,998,311,337]" name="Israel" pageId="25" pageNumber="26">Israel</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9CF3FE86FBBDFEDB" box="[1010,1092,311,337]" name="Jordan" pageId="25" pageNumber="26">Jordan</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9B4EFE86FB25FEDB" box="[1103,1244,311,337]" name="Kazakhstan" pageId="25" pageNumber="26">Kazakhstan</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9BE9FE89FA8AFED8" box="[1256,1395,312,338]" name="Kyrgyzstan" pageId="25" pageNumber="26">Kyrgyzstan</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9FBCFEEBFEDEFEFE" box="[189,295,346,372]" name="Lebanon" pageId="25" pageNumber="26">Lebanon</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9E34FEEBFE65FEFE" box="[309,412,346,372]" name="Pakistan" pageId="25" pageNumber="26">Pakistan</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9EA8FEEBFE02FEFE" box="[425,507,346,372]" name="Russia" pageId="25" pageNumber="26">Russia</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9D08FEEBFDB2FEFE" box="[521,587,346,372]" name="Syria" pageId="25" pageNumber="26">Syria</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9D56FEEBFD37FEFE" box="[599,718,346,372]" name="Tajikistan" pageId="25" pageNumber="26">Tajikistan</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9DDAFEEBFCCAFEFE" box="[731,819,346,372]" name="Turkey" pageId="25" pageNumber="26">Turkey</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9C41FEEBFC1FFEFE" box="[832,998,346,372]" name="Turkmenistan" pageId="25" pageNumber="26">Turkmenistan</collectingCountry>
;
<collectingCountry id="A5CF7654FFEF891F9CF2FEEBFB8EFEFE" box="[1011,1143,346,372]" name="Uzbekistan" pageId="25" pageNumber="26">Uzbekistan</collectingCountry>
(
<bibRefCitation id="B9494B35FFEF891F9B86FEEBFF02FE12" author="Hashmi A. &amp; Tashfeen A." pageId="25" pageNumber="26" pagination="133 - 170" refId="ref29508" refString="Hashmi A. &amp; Tashfeen A. 1992. Coleoptera of Pakistan. Proceedings of Pakistan Congress of Zoology 12: 133 - 170." type="journal article" year="1992">Hashmi &amp; Tashfeen 1992</bibRefCitation>
;
<bibRefCitation id="B9494B35FFEF891F9E08FECCFE77FE1D" author="Awal M. M." box="[265,398,381,407]" pageId="25" pageNumber="26" pagination="163 - 172" refId="ref27017" refString="Awal M. M. 2006. Preliminary studies on Scarabaeoidea (Coleoptera) fauna of Razavi Khorasan province of Iran. Turkiye Entomoloji Dergisi 30 (3): 163 - 172. Available from: https: // dergipark. org. tr / tr / download / article-file / 64989 [accessed 27 Nov. 2019]" type="journal article" year="2006">Awal 2006</bibRefCitation>
;
<bibRefCitation id="B9494B35FFEF891F9E9FFECCFD5FFE1D" author="Bezdek A. &amp; Krell F. - T." box="[414,678,381,408]" pageId="25" pageNumber="26" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
; Siddhiqui
<emphasis id="EFACEAD6FFEF891F9C33FECFFC89FE1D" box="[818,880,381,407]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2014;
<bibRefCitation id="B9494B35FFEF891F9CC9FECCFB90FE1D" author="Bezdek A." box="[968,1129,381,407]" pageId="25" pageNumber="26" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
;
<bibRefCitation id="B9494B35FFEF891F9B79FECCFA8BFE1D" author="Bunalski M. &amp; Samin N. &amp; Ghahari H." box="[1144,1394,381,407]" pageId="25" pageNumber="26" pagination="31 - 40" refId="ref28079" refString="Bunalski M., Samin N. &amp; Ghahari H. 2016. A contribution to the faunistic study of Scarabaeoidea (Coleoptera) from Mazandaran Province, northern Iran. Wiadomosci Entomologiczne 35 (1): 31 - 40. Available from: https: // sparrow. up. poznan. pl / pte / we / 2016 / 35 _ 3 _ Bunalski _ i _ in. pdf [Accessed 15 Nov. 2018]" type="journal article" year="2016">
Bunalski
<emphasis id="EFACEAD6FFEF891F9BEAFECFFAD0FE1D" box="[1259,1321,381,407]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2016
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;
<bibRefCitation id="B9494B35FFEF891F9FBCFE10FE81FE31" author="Montreuil O." box="[189,376,417,443]" pageId="25" pageNumber="26" pagination="259 - 277" refId="ref30308" refString="Montreuil O. 2017. New records of Geotrupidae and Scarabeidae dung beetles (Insecta, Coleoptera) from Iran. Journal of Entomological Society of Iran 36 (4): 259 - 277." type="journal article" year="2017">Montreuil 2017</bibRefCitation>
;
<bibRefCitation id="B9494B35FFEF891F9E85FE10FD7AFE31" author="Schoolmeesters P." box="[388,643,417,443]" pageId="25" pageNumber="26" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,898 @@
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<subSubSection id="95C2654FFFE289129D65FB6AFC36FB7F" box="[612,975,1242,1269]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="DD6736C4FFE289129D65FB6AFC36FB7F" blockId="20.[612,975,1242,1305]" box="[612,975,1242,1269]" pageId="20" pageNumber="21">
<heading id="862F81A8FFE289129D65FB6AFC36FB7F" box="[612,975,1242,1269]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
<taxonomicName id="1AD84D47FFE289129D65FB6AFC36FB7F" ID-CoL="49KG5" authority="(Wulfen, 1786)" baseAuthorityName="Wulfen" baseAuthorityYear="1786" box="[612,975,1242,1269]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFE289129D65FB6AFCECFB7F" bold="true" box="[612,789,1242,1269]" italics="true" pageId="20" pageNumber="21">Onitis falcatus</emphasis>
(
<bibRefCitation id="B9494B35FFE289129C25FB6AFC3EFB7F" author="Wulfen X." box="[804,967,1243,1269]" pageId="20" pageNumber="21" refId="ref31404" refString="Wulfen X. 1786. Descriptiones quorumdam Capensium insectorum. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1786">Wulfen, 1786</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE289129DBEFB4FFC8DFA92" box="[703,884,1278,1305]" pageId="20" pageNumber="21" type="description">
<paragraph id="DD6736C4FFE289129DBEFB4FFC8DFA92" blockId="20.[612,975,1242,1305]" box="[703,884,1278,1305]" pageId="20" pageNumber="21">
<figureCitation id="45E32A41FFE289129DBEFB4FFCE0FA93" box="[703,793,1278,1305]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="20" pageNumber="21">Figs 5F</figureCitation>
,
<figureCitation id="45E32A41FFE289129C25FB4FFCB3FA92" box="[804,842,1278,1304]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="20" pageNumber="21">6H</figureCitation>
,
<figureCitation id="45E32A41FFE289129C56FB4FFC8DFA92" box="[855,884,1278,1304]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="20" pageNumber="21">18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE289129FBCFAF1FB6AF894" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="DD6736C4FFE289129FBCFAF1FC7FFAD0" blockId="20.[189,902,1344,1441]" box="[189,902,1344,1370]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="FDC811EAFFE289129FBCFAF1FC7FFAD0" box="[189,902,1344,1370]" pageId="20" pageNumber="21">
<treatmentCitation id="5C7910D5FFE289129FBCFAF1FD87FAD0" author="Wulfen X." box="[189,638,1344,1370]" page="14" pageId="20" pageNumber="21" year="1786">
<taxonomicName id="1AD84D47FFE289129FBCFAF1FD87FAD0" ID-CoL="79W7R" authority="Wulfen, 1786: 14" authorityName="Wulfen" authorityPageNumber="14" authorityYear="1786" box="[189,638,1344,1370]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFE289129FBCFAF1FE53FAD0" box="[189,426,1344,1370]" italics="true" pageId="20" pageNumber="21">Scarabaeus falcatus</emphasis>
<bibRefCitation id="B9494B35FFE289129EB0FAF1FD87FAD0" author="Wulfen X." box="[433,638,1344,1370]" pageId="20" pageNumber="21" refId="ref31404" refString="Wulfen X. 1786. Descriptiones quorumdam Capensium insectorum. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1786">Wulfen, 1786: 14</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFAD2FCAAFAF7" blockId="20.[189,902,1344,1441]" box="[189,851,1379,1406]" pageId="20" pageNumber="21">
<emphasis id="EFACEAD6FFE289129FBCFAD2FE61FAF7" box="[189,408,1379,1405]" italics="true" pageId="20" pageNumber="21">Onitishimaleyicus</emphasis>
Redtenbacher, 1844: 518 (synonym).
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFA37FD27FA2A" blockId="20.[189,902,1344,1441]" box="[189,734,1414,1441]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="FDC811EAFFE289129FBCFA37FD27FA2A" box="[189,734,1414,1441]" pageId="20" pageNumber="21">
<treatmentCitation id="5C7910D5FFE289129FBCFA37FDB6FA2A" author="Masumoto, K." box="[189,591,1414,1441]" page="88" pageId="20" pageNumber="21" year="1995">
<taxonomicName id="1AD84D47FFE289129FBCFA37FDB6FA2A" ID-CoL="49KHB" authority="Masumoto 1995: 88" authorityName="Masumoto" authorityPageNumber="88" authorityYear="1995" box="[189,591,1414,1441]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="kiuchii">
<emphasis id="EFACEAD6FFE289129FBCFA37FEA5FA2A" box="[189,348,1414,1440]" italics="true" pageId="20" pageNumber="21">Onitis kiuchii</emphasis>
<bibRefCitation id="B9494B35FFE289129E62FA36FDB6FA2A" author="Masumoto, K." box="[355,591,1414,1441]" pageId="20" pageNumber="21" pagination="87 - 94" refId="ref30072" refString="Masumoto, K. 1995. Coprophagid-beetles from Northwest Thailand (X) (Coleoptera, Scarabaeidae). The Entomological Review of Japan 50 (2): 87 - 94. https: // archive. org / details / entomologicalreviewofjapan 19960050002 / page / n 16" type="journal article" year="1995">Masumoto 1995: 88</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(synonym).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFA76FB6AF894" blockId="20.[189,1399,1479,1823]" pageId="20" pageNumber="21">
<emphasis id="EFACEAD6FFE289129FBCFA76FE91FA6B" box="[189,360,1479,1505]" italics="true" pageId="20" pageNumber="21">Onitisfalcatus</emphasis>
<bibRefCitation id="B9494B35FFE289129E82FA76FD8EFA6B" author="Lansberge G." box="[387,631,1479,1506]" pageId="20" pageNumber="21" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge 1875: 126</bibRefCitation>
(monograph). — Boucomont &amp; Gillet: 1921: 17 (distribution). —
<bibRefCitation id="B9494B35FFE289129FEBFA5AFE4BF98F" author="Arrow G. J." box="[234,434,1515,1541]" pageId="20" pageNumber="21" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 392</bibRefCitation>
(key and description). —
<bibRefCitation id="B9494B35FFE289129DE1FA5AFC42F98F" author="Balthasar V." box="[736,955,1515,1542]" pageId="20" pageNumber="21" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 93</bibRefCitation>
(monograph); 1963: 33 (monograph). —
<bibRefCitation id="B9494B35FFE289129E0FF9BFFE19F9A2" author="Janssens A." box="[270,480,1550,1576]" pageId="20" pageNumber="21" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 44</bibRefCitation>
(revision). —
<bibRefCitation id="B9494B35FFE289129D8AF9BFFCA6F9A2" author="Paulian R." box="[651,863,1550,1577]" pageId="20" pageNumber="21" refId="ref30535" refString="Paulian R. 1945. Faune de l'empire francais: Coleopteres Scarabeides de l'Indochine, Vol. III: 1 - 227. Librairie Larose, Paris." type="book" year="1945">Paulian 1945: 142</bibRefCitation>
(revision). —
<bibRefCitation id="B9494B35FFE289129B08F9BFFA8FF9A3" author="Biswas S. &amp; Chatterjee S. K." box="[1033,1398,1550,1577]" pageId="20" pageNumber="21" pagination="1 - 4" refId="ref27511" refString="Biswas S. &amp; Chatterjee S. K. 1985. Insecta: Coleoptera: Scarabaeidae: Coprinae. Records of the Zoological Survey of India, Fauna of Namdapha, Arunachal Pradesh (Special Issue), 82 (1 - 4): 147 - 177. https: // doi. org / 10.26515 / rzsi / v 82 / i 1 - 4 / 1984 / 161291" type="journal article" year="1985">Biswas &amp; Chatterjee 1985: 158</bibRefCitation>
(distribution); 1991: 260 (distribution). —
<bibRefCitation id="B9494B35FFE289129DD5F980FC56F9C1" author="Chandra K." box="[724,943,1585,1611]" pageId="20" pageNumber="21" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000: 360</bibRefCitation>
(distribution); 2005: 150 (distribution). —
<bibRefCitation id="B9494B35FFE289129E12F9E5FE29F9E4" author="Mittal I. C." box="[275,464,1620,1647]" pageId="20" pageNumber="21" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005: 46</bibRefCitation>
(list). —
<bibRefCitation id="B9494B35FFE289129D4CF9E5FCEFF9E4" author="Rajan P. D." box="[589,790,1620,1646]" pageId="20" pageNumber="21" pagination="91 - 135" refId="ref30757" refString="Rajan P. D. 2006. Insecta: Coleoptera: Scarabaeoidea: Scarabaeidae (dung beetles). Zoological Survey of India. Fauna of Bilgiri Rangaswamy Tiger Wildlife Sanctuary, Conservation Area Series 27: 91 - 135." type="book chapter" year="2006">Rajan 2006: 108</bibRefCitation>
(key, distribution). —
<bibRefCitation id="B9494B35FFE289129B30F9E5FA8FF9E5" author="Bezdek A. &amp; Krell F. - T." box="[1073,1398,1620,1647]" pageId="20" pageNumber="21" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
(distribution). —
<bibRefCitation id="B9494B35FFE289129EBCF9C6FCD4F91B" author="Chandra K. &amp; Ahirwar S. C." box="[445,813,1655,1682]" pageId="20" pageNumber="21" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">Chandra &amp; Ahirwar 2007: 276</bibRefCitation>
(distributional). —
<bibRefCitation id="B9494B35FFE289129B1FF9C6FB14F91B" author="Sewak R." box="[1054,1261,1655,1682]" pageId="20" pageNumber="21" pagination="1 - 48" refId="ref31194" refString="Sewak R. 2009 a. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar desert of Gujarat. Records of the Zoological Survey of India, Occassional Paper No 295: 1 - 48." type="journal article" year="2009">Sewak 2009a: 36</bibRefCitation>
(diagnosis, distribution); 2009b: 58 (diagnosis and distribution). — Singh
<emphasis id="EFACEAD6FFE289129CD9F92DFBECF93F" box="[984,1045,1691,1717]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2010: 1395 (distribution). —
<bibRefCitation id="B9494B35FFE289129FEBF90FFE25F952" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." box="[234,476,1726,1752]" pageId="20" pageNumber="21" pagination="29 - 38" refId="ref30906" refString="Sabu T. K., Nithya S. &amp; Vinod K. V. 2011. Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa 2830: 29 - 38. https: // doi. org / 10.11646 / ZOOTAXA. 2830.1.3" type="book chapter" year="2011">
Sabu
<emphasis id="EFACEAD6FFE289129E2FF90EFE92F952" box="[302,363,1726,1752]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2011: 34
</bibRefCitation>
(list). — Chandra
<emphasis id="EFACEAD6FFE289129DC1F90EFD04F952" box="[704,765,1726,1752]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2012a: 52 (distribution). —
<bibRefCitation id="B9494B35FFE289129B56F90FFE99F971" author="Karimbumkara S. N. &amp; Rajan P. D." pageId="20" pageNumber="21" pagination="173 - 178" refId="ref29842" refString="Karimbumkara S. N. &amp; Rajan P. D. 2013. Insecta: Coleoptera: Scarabaeidae: Scarabaeinae, Dung Beetles. Zoological Survey of India, Fauna of Karnataka, State Fauna Series 21: 173 - 178." type="book chapter" year="2013">Karimbumkara &amp; Rajan 2013: 174</bibRefCitation>
(distribution). —
<bibRefCitation id="B9494B35FFE289129D32F950FCBFF971" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[563,838,1761,1788]" pageId="20" pageNumber="21" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFE289129D80F953FD42F971" box="[641,699,1761,1787]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2015: 1037
</bibRefCitation>
(checklist). —
<bibRefCitation id="B9494B35FFE289129CFBF950FB37F971" author="Bezdek A." box="[1018,1230,1761,1787]" pageId="20" pageNumber="21" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
(distribution). —
<bibRefCitation id="B9494B35FFE289129E0EF8B5FE19F894" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." box="[271,480,1796,1822]" pageId="20" pageNumber="21" pagination="221 - 275" refId="ref29141" refString="Ghosh J., Das, P., Ghosh S. K., Bhunia D., Kushwaha R. K., Gupta D., &amp; Chandra K. 2020. Insecta: Coleoptera. Zoological Survey of India, Fauna of Haryana, State Fauna Series 24: 221 - 275." type="book chapter" year="2020">
Ghosh
<emphasis id="EFACEAD6FFE289129E62F8B4FE67F894" box="[355,414,1796,1822]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2020
</bibRefCitation>
(distribution). —
<bibRefCitation id="B9494B35FFE289129DB4F8B5FC49F894" author="Schoolmeesters P." box="[693,944,1796,1822]" pageId="20" pageNumber="21" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE289129FBCF8F4FC6AF85F" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="DD6736C4FFE289129FBCF8F4FE3FF8D5" blockId="20.[189,454,1861,1887]" box="[189,454,1861,1887]" pageId="20" pageNumber="21">
<heading id="862F81A8FFE289129FBCF8F4FE3FF8D5" bold="true" box="[189,454,1861,1887]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="EFACEAD6FFE289129FBCF8F4FE3FF8D5" bold="true" box="[189,454,1861,1887]" pageId="20" pageNumber="21">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCF8C5FC6AF85F" blockId="20.[189,1399,1908,2005]" pageId="20" pageNumber="21">
<emphasis id="EFACEAD6FFE289129FBCF8C5FE92F804" box="[189,363,1908,1934]" italics="true" pageId="20" pageNumber="21">Onitisfalcatus</emphasis>
is similar to
<taxonomicName id="1AD84D47FFE289129D0FF8C4FD51F804" authorityName="Arrow" authorityYear="1931" box="[526,680,1909,1935]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="EFACEAD6FFE289129D0FF8C4FD51F804" box="[526,680,1909,1935]" italics="true" pageId="20" pageNumber="21">O. excavatus</emphasis>
</taxonomicName>
in having a minutely punctured pronotum but differs in the nature of rugosity on the clypeus, the presence of longitudinal groove on the pronotum and also in the shape of the body. (Also see remarks under
<emphasis id="EFACEAD6FFE289129DBEF80AFC70F85F" box="[703,905,1979,2005]" italics="true" pageId="20" pageNumber="21">Onitisexcavatus.</emphasis>
)
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE389139FBCFEB8FC37F92E" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="DD6736C4FFE389139FBCFEB8FE15FEA8" blockId="21.[189,492,264,291]" box="[189,492,264,291]" pageId="21" pageNumber="22">
<heading id="862F81A8FFE389139FBCFEB8FE15FEA8" bold="true" box="[189,492,264,291]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="EFACEAD6FFE389139FBCFEB8FE15FEA8" bold="true" box="[189,492,264,291]" pageId="21" pageNumber="22">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE389139FEBFE86FDFFFEDB" blockId="21.[189,1398,311,408]" box="[234,518,311,337]" pageId="21" pageNumber="22">
<emphasis id="EFACEAD6FFE389139FEBFE86FEA3FEDB" bold="true" box="[234,346,311,337]" pageId="21" pageNumber="22">
<typeStatus id="02638866FFE389139FEBFE86FEA3FEDB" box="[234,346,311,337]" pageId="21" pageNumber="22" type="syntype">Syntypes</typeStatus>
</emphasis>
(
<specimenCount id="CBDEFD4DFFE389139E6AFE86FE07FEDB" box="[363,510,311,337]" count="2" pageId="21" pageNumber="22" type="generic">2 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFE389139FBCFEEBFD62FE12" blockId="21.[189,1398,311,408]" pageId="21" pageNumber="22">
<collectingCountry id="A5CF7654FFE389139FBCFEEBFEF7FEFE" box="[189,270,346,372]" name="India" pageId="21" pageNumber="22">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFE389139E2CFEEBFE8AFEFF" box="[301,371,346,373]" count="2" pageId="21" pageNumber="22" type="female">2 ♀♀</specimenCount>
; “
<taxonomicName id="1AD84D47FFE389139E90FEEAFDBDFEFE" box="[401,580,346,373]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="himaleyicus">
<emphasis id="EFACEAD6FFE389139E90FEEAFDBDFEFE" box="[401,580,346,373]" italics="true" pageId="21" pageNumber="22">O. himaleyicus</emphasis>
</taxonomicName>
;
<typeStatus id="02638866FFE389139D54FEEBFD4BFEFE" box="[597,690,346,372]" pageId="21" pageNumber="22">TYPUS</typeStatus>
(Red card); Hugel: 170;
<taxonomicName id="1AD84D47FFE389139CE7FEEBFB15FEFE" authority="K. Redt." authorityName="K. Redt." box="[998,1260,346,373]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="himaleyicus">Himaleyicus K. Redt.</taxonomicName>
”; Kashmir; specimen number not assigned;
<collectionCode id="BBC9AE01FFE389139D34FECFFD6FFE12" box="[565,662,382,408]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="22" type="Museum">NHMW</collectionCode>
.
</paragraph>
<paragraph id="DD6736C4FFE389139FEBFE0EFCD3FE53" blockId="21.[189,1399,447,1599]" box="[234,810,447,474]" pageId="21" pageNumber="22">
<emphasis id="EFACEAD6FFE389139FEBFE0EFD9BFE53" bold="true" box="[234,610,447,473]" pageId="21" pageNumber="22">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFE389139D70FE0EFCDBFE53" box="[625,802,447,474]" count="125" pageId="21" pageNumber="22" type="generic">125 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFE389139FBCFE53FD43F9B5" blockId="21.[189,1399,447,1599]" pageId="21" pageNumber="22">
<materialsCitation id="6DB03C99FFE389139FBCFE53FEFAFD95" collectingDate="1913-05" collectionCode="ZSIM" country="India" county="Darjiling" elevation="457" location="Lord Carmichael's" municipality="Singla" pageId="21" pageNumber="22" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFE389139FBCFE53FEF7FE76" box="[189,270,482,508]" name="India" pageId="21" pageNumber="22">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFE389139E4BFE52FE9BFE77" box="[330,354,483,509]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139E81FE52FE6EFE77" box="[384,407,483,509]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
; “
<collectingCounty id="34064E48FFE389139EC3FE53FDCBFE76" box="[450,562,482,508]" pageId="21" pageNumber="22">Darjiling</collectingCounty>
,
<collectingMunicipality id="3D03ACBEFFE389139D4FFE53FD65FE76" box="[590,668,482,508]" pageId="21" pageNumber="22">Singla</collectingMunicipality>
,
<quantity id="1A209B21FFE389139DB8FE53FCDBFE76" box="[697,802,482,509]" metricMagnitude="2" metricUnit="m" metricValue="4.572" pageId="21" pageNumber="22" unit="ft" value="1500.0">
<elevation id="56F5D1F7FFE389139DB8FE53FCDBFE76" box="[697,802,482,509]" metricMagnitude="2" metricUnit="m" metricValue="4.572" pageId="21" pageNumber="22" unit="ft" value="1500.0">1500 ft</elevation>
</quantity>
,
<location id="D807601FFFE389139C3EFE53FBD5FE76" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139C3EFE53FBD5FE76" box="[831,1068,482,508]" country="India" county="Darjiling" municipality="Singla" name="Lord Carmichael's" pageId="21" pageNumber="22">Lord Carmichaels</location>
collection,
<date id="A9661004FFE389139BD9FE52FA9FFE76" box="[1240,1382,482,509]" pageId="21" pageNumber="22" value="1913-05">
<collectingDate id="B922E9ECFFE389139BD9FE52FA9FFE76" box="[1240,1382,482,509]" pageId="21" pageNumber="22" value="1913-05">May 1913</collectingDate>
</date>
”;
<collectionCode id="BBC9AE01FFE389139FBCFDB4FEFAFD95" box="[189,259,517,543]" pageId="21" pageNumber="22">ZSIM</collectionCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139E4EFDB4FB4AFDC8" collectingDate="1999-06-06" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" latitude="11.931833" location="Gombegallu" longLatPrecision="1" longitude="77.1843" municipality="Tiger Reserve" pageId="21" pageNumber="22" specimenCount="43" specimenCount-female="16" specimenCount-male="27" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFE389139E4EFDB4FE42FDAA" box="[335,443,517,544]" count="27" pageId="21" pageNumber="22" type="male">27 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139EE0FDB4FDB5FDAA" box="[481,588,517,544]" count="16" pageId="21" pageNumber="22" type="female">16 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE389139D75FDB4FD09FD95" box="[628,752,517,543]" country="India" name="Karnataka" pageId="21" pageNumber="22">Karnataka</collectingRegion>
, BRT
<collectingMunicipality id="3D03ACBEFFE389139C6AFDB4FBD4FDAA" box="[875,1069,517,544]" pageId="21" pageNumber="22">Tiger Reserve</collectingMunicipality>
:
<location id="D807601FFFE389139B53FDB4FB1AFD95" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139B53FDB4FB1AFD95" box="[1106,1251,517,543]" country="India" latitude="11.931833" longLatPrecision="1" longitude="77.1843" municipality="Tiger Reserve" name="Gombegallu" pageId="21" pageNumber="22" stateProvince="Karnataka">Gombegallu</location>
evergreen forest;
<date id="A9661004FFE389139E13FD99FE5DFDC9" box="[274,420,552,579]" pageId="21" pageNumber="22" value="1999-06-06">
<collectingDate id="B922E9ECFFE389139E13FD99FE5DFDC9" box="[274,420,552,579]" pageId="21" pageNumber="22" value="1999-06-06">6 Jun. 1999</collectingDate>
</date>
;
<geoCoordinate id="B8EC5003FFE389139EB5FD96FDACFDC9" box="[436,597,551,579]" degrees="11" direction="north" minutes="55" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="54.6" value="11.931833">11°55ʹ54.6 N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139D65FD96FCF6FDC9" box="[612,783,551,579]" degrees="77" direction="east" minutes="11" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="3.48" value="77.1843">77°11ʹ3.48″ E</geoCoordinate>
;
<collectionCode id="BBC9AE01FFE389139C1CFD99FC93FDC8" box="[797,874,552,578]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139C72FD99FC6CFDC8" box="[883,917,552,578]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139C9CFD99FC40FDC8" box="[925,953,552,578]" pageId="21" pageNumber="22">Sc</collectorName>
1000064 to 1000106
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139BD3FD99FEDAFD26" collectingDate="1999-05-28" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="11.89545" location="Kanneri" longLatPrecision="1" longitude="77.13596" municipality="Tiger Reserve" pageId="21" pageNumber="22" specimenCount="11" specimenCount-female="5" specimenCount-male="6" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFE389139BD3FD99FAE0FDC9" box="[1234,1305,552,579]" count="6" pageId="21" pageNumber="22" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139A28FD99FA96FDC9" box="[1321,1391,552,579]" count="5" pageId="21" pageNumber="22" type="female">5 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE389139FBCFDFDFEC0FDEC" box="[189,313,588,614]" country="India" name="Karnataka" pageId="21" pageNumber="22">Karnataka</collectingRegion>
, BRT
<collectingMunicipality id="3D03ACBEFFE389139E91FDFDFDB8FDEC" box="[400,577,588,614]" pageId="21" pageNumber="22">Tiger Reserve</collectingMunicipality>
:
<location id="D807601FFFE389139D52FDFDFD48FDEC" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139D52FDFDFD48FDEC" box="[595,689,588,614]" country="India" latitude="11.89545" longLatPrecision="1" longitude="77.13596" municipality="Tiger Reserve" name="Kanneri" pageId="21" pageNumber="22" stateProvince="Karnataka">Kanneri</location>
moist deciduous forest;
<date id="A9661004FFE389139CEEFDFDFB67FDEC" box="[1007,1182,588,614]" pageId="21" pageNumber="22" value="1999-05-28">
<collectingDate id="B922E9ECFFE389139CEEFDFDFB67FDEC" box="[1007,1182,588,614]" pageId="21" pageNumber="22" value="1999-05-28">28 May 1999</collectingDate>
</date>
;
<geoCoordinate id="B8EC5003FFE389139BB0FDFBFA8BFDEC" box="[1201,1394,586,614]" degrees="11" direction="north" minutes="53" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="43.62" value="11.89545">11°53ʹ43.62″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139FBCFDDCFEA3FD03" box="[189,346,621,649]" degrees="77" direction="east" minutes="8" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="9.48" value="77.13596">77°8ʹ9.48″ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFE389139E68FDDEFD59FD03" box="[361,672,623,649]" pageId="21" pageNumber="22">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFE389139DAFFDDEFBF2FD03" box="[686,1035,623,649]" pageId="21" pageNumber="22">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE389139B53FDDEFB66FD03" box="[1106,1183,623,649]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139BA9FDDEFB33FD03" box="[1192,1226,623,649]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139BD2FDDEFB16FD03" box="[1235,1263,623,649]" pageId="21" pageNumber="22">Sc</collectorName>
1000107 to 1000117
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139E3AFD22FAE7FD45" collectingDate="1999-05-18" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="12.032" location="Purani" longLatPrecision="1" longitude="77.1313" municipality="Tiger Reserve" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFE389139E3AFD22FEABFD27" box="[315,338,659,685]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE389139E61FD23FE25FD26" box="[352,476,658,684]" country="India" name="Karnataka" pageId="21" pageNumber="22">Karnataka</collectingRegion>
, BRT
<collectingMunicipality id="3D03ACBEFFE389139D24FD23FD34FD27" box="[549,717,658,685]" pageId="21" pageNumber="22">Tiger Reserve</collectingMunicipality>
:
<location id="D807601FFFE389139DD9FD23FCDCFD26" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139DD9FD23FCDCFD26" box="[728,805,658,684]" country="India" latitude="12.032" longLatPrecision="1" longitude="77.1313" municipality="Tiger Reserve" name="Purani" pageId="21" pageNumber="22" stateProvince="Karnataka">Purani</location>
dry deciduous forest;
<date id="A9661004FFE389139B2BFD23FB30FD26" box="[1066,1225,658,685]" pageId="21" pageNumber="22" value="1999-05-18">
<collectingDate id="B922E9ECFFE389139B2BFD23FB30FD26" box="[1066,1225,658,685]" pageId="21" pageNumber="22" value="1999-05-18">18 May 1999</collectingDate>
</date>
;
<geoCoordinate id="B8EC5003FFE389139BD4FD21FA8BFD27" box="[1237,1394,656,685]" degrees="12" direction="north" minutes="1" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="55.2" value="12.032">12°1ʹ55.2″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139FBCFD05FE9BFD5A" box="[189,354,692,720]" degrees="77" direction="east" minutes="7" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="52.68" value="77.1313">77°7ʹ52.68″E</geoCoordinate>
;
<collectingMethod id="04994ED3FFE389139E6AFD07FD70FD5A" box="[363,649,693,720]" pageId="21" pageNumber="22">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFE389139D90FD04FC1BFD45" box="[657,994,693,719]" pageId="21" pageNumber="22">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE389139B1DFD04FB90FD45" box="[1052,1129,693,719]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139B73FD04FB6DFD45" box="[1138,1172,693,719]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139B9DFD04FB41FD45" box="[1180,1208,693,719]" pageId="21" pageNumber="22">Sc</collectorName>
1000118
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139A31FD04FAF3FC9C" collectingDate="1998-04" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="11.8907" location="Honnametti Shola" longLatPrecision="1" longitude="77.20138" municipality="Tiger Reserve" pageId="21" pageNumber="22" specimenCount="16" specimenCount-female="7" specimenCount-male="9" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFE389139A31FD04FA88FD5A" box="[1328,1393,693,720]" count="9" pageId="21" pageNumber="22" type="male">9 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139FBCFD68FEFBFD79" box="[189,258,729,755]" count="7" pageId="21" pageNumber="22" type="female">7 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE389139E13FD69FE77FD78" box="[274,398,728,754]" country="India" name="Karnataka" pageId="21" pageNumber="22">Karnataka</collectingRegion>
, BRT
<collectingMunicipality id="3D03ACBEFFE389139EDBFD69FD7DFD79" box="[474,644,728,755]" pageId="21" pageNumber="22">Tiger Reserve</collectingMunicipality>
:
<location id="D807601FFFE389139D90FD69FC93FD78" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139D90FD69FC93FD78" box="[657,874,728,754]" country="India" latitude="11.8907" longLatPrecision="1" longitude="77.20138" municipality="Tiger Reserve" name="Honnametti Shola" pageId="21" pageNumber="22" stateProvince="Karnataka">Honnametti Shola</location>
;
<date id="A9661004FFE389139C76FD68FC08FD79" box="[887,1009,729,755]" pageId="21" pageNumber="22" value="1998-04">
<collectingDate id="B922E9ECFFE389139C76FD68FC08FD79" box="[887,1009,729,755]" pageId="21" pageNumber="22" value="1998-04">Apr. 1998</collectingDate>
</date>
;
<geoCoordinate id="B8EC5003FFE389139CFEFD66FB43FD79" box="[1023,1210,727,755]" degrees="11" direction="north" minutes="53" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="26.52" value="11.8907">11°53ʹ26.52″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139BC6FD66FA88FD79" box="[1223,1393,727,755]" degrees="77" direction="east" minutes="12" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="4.98" value="77.20138">77°12ʹ4.98″ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFE389139FBCFD4DFE1BFC9C" box="[189,482,764,790]" pageId="21" pageNumber="22">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFE389139EEDFD4DFCB8FC9C" box="[492,833,764,790]" pageId="21" pageNumber="22">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE389139C7FFD4DFC33FC9C" box="[894,970,764,790]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139CD5FD4DFC0FFC9C" box="[980,1014,764,790]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139CFEFD4DFBE2FC9C" box="[1023,1051,764,790]" pageId="21" pageNumber="22">Sc</collectorName>
1000119 to 1000134
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139A21FD4DFB10FCD7" collectingDate="1998-05" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" latitude="12.0195" location="Gummane Shola" longLatPrecision="1" longitude="77.17518" municipality="Tiger Reserve" pageId="21" pageNumber="22" specimenCount="18" specimenCount-female="5" specimenCount-male="13" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFE389139A21FD4DFA88FC9C" box="[1312,1393,764,790]" count="13" pageId="21" pageNumber="22" type="male">13 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139FBCFCAEFF06FCB3" box="[189,255,799,826]" count="5" pageId="21" pageNumber="22" type="female">5 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE389139E0DFCAEFE71FCB3" box="[268,392,799,825]" country="India" name="Karnataka" pageId="21" pageNumber="22">Karnataka</collectingRegion>
, BRT
<collectingMunicipality id="3D03ACBEFFE389139ECFFCAEFD8FFCB3" box="[462,630,799,825]" pageId="21" pageNumber="22">Tiger Reserve</collectingMunicipality>
:
<location id="D807601FFFE389139D81FCAEFCBFFCB3" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139D81FCAEFCBFFCB3" box="[640,838,799,825]" country="India" latitude="12.0195" longLatPrecision="1" longitude="77.17518" municipality="Tiger Reserve" name="Gummane Shola" pageId="21" pageNumber="22" stateProvince="Karnataka">Gummane Shola</location>
;
<date id="A9661004FFE389139C50FCAEFC32FCB3" box="[849,971,799,825]" pageId="21" pageNumber="22" value="1998-05">
<collectingDate id="B922E9ECFFE389139C50FCAEFC32FCB3" box="[849,971,799,825]" pageId="21" pageNumber="22" value="1998-05">May 1998</collectingDate>
</date>
;
<geoCoordinate id="B8EC5003FFE389139CD7FCACFB88FCB3" box="[982,1137,797,825]" degrees="12" direction="north" minutes="1" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="10.2" value="12.0195">12°1ʹ10.2″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139B7DFCACFACBFCB3" box="[1148,1330,797,825]" degrees="77" direction="east" minutes="10" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="30.66" value="77.17518">77°10ʹ30.66″ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFE389139A3CFCAEFE55FCD7" pageId="21" pageNumber="22">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFE389139EB8FCF3FCEAFCD6" box="[441,787,834,860]" pageId="21" pageNumber="22">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE389139C56FCF3FC5DFCD7" box="[855,932,834,861]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139CACFCF3FC36FCD7" box="[941,975,834,861]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139CD9FCF3FC0DFCD7" box="[984,1012,834,861]" pageId="21" pageNumber="22">Sc</collectorName>
1000135 to 1000152
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139A04FCF3FE05FC29" collectingDate="2010-11-14" collectionCode="AIMB" collectorName="Nisha, P. V. &amp; Co &amp; Sc" country="India" location="Wayanad" pageId="21" pageNumber="22" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFE389139A04FCF3FAB3FCD7" box="[1285,1354,834,861]" count="2" pageId="21" pageNumber="22" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139A59FCF2FA96FCD7" box="[1368,1391,835,861]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE389139FBCFCD4FEF7FCF5" box="[189,270,869,895]" country="India" name="Kerala" pageId="21" pageNumber="22">Kerala</collectingRegion>
,
<location id="D807601FFFE389139E18FCD4FE70FCF5" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139E18FCD4FE70FCF5" box="[281,393,869,895]" country="India" name="Wayanad" pageId="21" pageNumber="22" stateProvince="Kerala">Wayanad</location>
, arecanut plantation;
<date id="A9661004FFE389139D82FCD4FCDCFCF5" box="[643,805,869,896]" pageId="21" pageNumber="22" value="2010-11-14">
<collectingDate id="B922E9ECFFE389139D82FCD4FCDCFCF5" box="[643,805,869,896]" pageId="21" pageNumber="22" value="2010-11-14">14 Nov. 2010</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE389139C30FCD7FBB6FC0A" box="[817,1103,869,896]" pageId="21" pageNumber="22">open cow dung bait trap</collectingMethod>
;
<collectorName id="702D5312FFE389139B5AFCD4FB1BFC0A" box="[1115,1250,869,896]" pageId="21" pageNumber="22">Nisha, P. V.</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE389139A23FCD4FA97FCF5" box="[1314,1390,869,895]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139FBCFC39FF26FC29" box="[189,223,904,931]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139FE9FC39FEFDFC29" box="[232,260,904,931]" pageId="21" pageNumber="22">Sc</collectorName>
1000153 to 1000155
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139D1BFC38FEDDFC63" collectingDate="2009-05-08" collectionCode="AIMB" collectorName="Kishore A. &amp; Co &amp; Sc" country="India" latitude="8.861611" location="Shendurney Wildlife Sanctuary" longLatPrecision="1" longitude="77.13347" pageId="21" pageNumber="22" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFE389139D1BFC38FD99FC29" box="[538,608,905,931]" count="4" pageId="21" pageNumber="22" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139D6EFC38FD4DFC29" box="[623,692,905,931]" count="3" pageId="21" pageNumber="22" type="female">3 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE389139DC4FC39FCEFFC28" box="[709,790,904,930]" country="India" name="Kerala" pageId="21" pageNumber="22">Kerala</collectingRegion>
,
<location id="D807601FFFE389139C25FC39FB60FC29" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139C25FC39FB60FC29" box="[804,1177,904,931]" country="India" latitude="8.861611" longLatPrecision="1" longitude="77.13347" name="Shendurney Wildlife Sanctuary" pageId="21" pageNumber="22" stateProvince="Kerala">Shendurney Wildlife Sanctuary</location>
, grassland;
<date id="A9661004FFE389139A29FC38FF02FC4C" pageId="21" pageNumber="22" value="2009-05-08">
<collectingDate id="B922E9ECFFE389139A29FC38FF02FC4C" pageId="21" pageNumber="22" value="2009-05-08">8 May 2009</collectingDate>
</date>
;
<geoCoordinate id="B8EC5003FFE389139E07FC1BFE56FC4C" box="[262,431,938,966]" degrees="08" direction="north" minutes="51" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="41.8" value="8.861611">08°51ʹ41.8ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139EB8FC1BFDA6FC4C" box="[441,607,938,966]" degrees="77" direction="east" minutes="08" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="00.5" value="77.13347">77°08ʹ00.5ʺ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFE389139D6BFC1DFC69FC4C" box="[618,912,940,966]" pageId="21" pageNumber="22">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFE389139C9AFC1DFBE0FC4C" box="[923,1049,940,966]" pageId="21" pageNumber="22">Kishore A.</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE389139B57FC1AFB5AFC4C" box="[1110,1187,939,966]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139BADFC1AFB37FC4C" box="[1196,1230,939,966]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139BD6FC1AFB0AFC4C" box="[1239,1267,939,966]" pageId="21" pageNumber="22">Sc</collectorName>
1000156 to 1000162
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139E3DFC7EFB1EFB86" collectingDate="2009-12-03" collectionCode="AIMB" collectorName="Kishore A. &amp; Co &amp; Sc" country="India" latitude="8.918222" location="Shendurney Wildlife Sanctuary" longLatPrecision="1" longitude="77.09575" pageId="21" pageNumber="22" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFE389139E3DFC7EFEADFC63" box="[316,340,975,1001]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139E61FC7EFE8EFC63" box="[352,375,975,1001]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE389139E84FC7EFE2FFC63" box="[389,470,975,1001]" country="India" name="Kerala" pageId="21" pageNumber="22">Kerala</collectingRegion>
,
<location id="D807601FFFE389139EE0FC7EFCA8FC63" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139EE0FC7EFCA8FC63" box="[481,849,975,1001]" country="India" latitude="8.918222" longLatPrecision="1" longitude="77.09575" name="Shendurney Wildlife Sanctuary" pageId="21" pageNumber="22" stateProvince="Kerala">Shendurney Wildlife Sanctuary</location>
, degraded forests;
<date id="A9661004FFE389139B2DFC7EFB44FC63" box="[1068,1213,975,1001]" pageId="21" pageNumber="22" value="2009-12-03">
<collectingDate id="B922E9ECFFE389139B2DFC7EFB44FC63" box="[1068,1213,975,1001]" pageId="21" pageNumber="22" value="2009-12-03">3 Dec. 2009</collectingDate>
</date>
;
<geoCoordinate id="B8EC5003FFE389139BC9FC7CFA8BFC63" box="[1224,1394,973,1001]" degrees="08" direction="north" minutes="55" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="05.6" value="8.918222">08°55ʹ05.6ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139FBCFC41FE9AFB87" box="[189,355,1008,1037]" degrees="77" direction="east" minutes="05" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="44.7" value="77.09575">77°05ʹ44.7ʺ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFE389139E6FFC42FD6DFB87" box="[366,660,1010,1037]" pageId="21" pageNumber="22">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFE389139D9FFC43FCE5FB86" box="[670,796,1010,1036]" pageId="21" pageNumber="22">Kishore A.</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE389139C5BFC43FC5EFB86" box="[858,935,1010,1036]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139CB1FC43FC2BFB86" box="[944,978,1010,1036]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139CDAFC43FC0EFB86" box="[987,1015,1010,1036]" pageId="21" pageNumber="22">Sc</collectorName>
1000163 to 1000164
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139BFCFC42FC94FBD9" collectingDate="2009-05-05" collectionCode="AIMB" collectorName="Kishore A. &amp; Co &amp; Sc" country="India" latitude="8.861611" location="Shendurney Wildlife Sanctuary" longLatPrecision="1" longitude="77.13347" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFE389139BFCFC42FAECFB87" box="[1277,1301,1011,1037]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
,
<collectingRegion id="1F1CF826FFE389139A20FC43FA8BFB86" box="[1313,1394,1010,1036]" country="India" name="Kerala" pageId="21" pageNumber="22">Kerala</collectingRegion>
,
<location id="D807601FFFE389139FBCFBA4FDCDFBA5" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139FBCFBA4FDCDFBA5" box="[189,564,1045,1071]" country="India" latitude="8.861611" longLatPrecision="1" longitude="77.13347" name="Shendurney Wildlife Sanctuary" pageId="21" pageNumber="22" stateProvince="Kerala">Shendurney Wildlife Sanctuary</location>
, rubber plantation;
<date id="A9661004FFE389139C20FBA4FC40FBA5" box="[801,953,1045,1072]" pageId="21" pageNumber="22" value="2009-05-05">
<collectingDate id="B922E9ECFFE389139C20FBA4FC40FBA5" box="[801,953,1045,1072]" pageId="21" pageNumber="22" value="2009-05-05">5 May 2009</collectingDate>
</date>
;
<geoCoordinate id="B8EC5003FFE389139CC9FBA5FB8CFBBA" box="[968,1141,1044,1072]" degrees="08" direction="north" minutes="51" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="41.8" value="8.861611">08°51ʹ41.8ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139B85FBA5FAD7FBBA" box="[1156,1326,1044,1072]" degrees="77" direction="east" minutes="08" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="00.5" value="77.13347">77°08ʹ00.5ʺ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFE389139A3CFBA7FE5DFBD9" pageId="21" pageNumber="22">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFE389139EAFFB89FDD5FBD9" box="[430,556,1080,1107]" pageId="21" pageNumber="22">Kishore A.</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE389139D6BFB89FD4EFBD9" box="[618,695,1080,1107]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139DC1FB89FD1BFBD9" box="[704,738,1080,1107]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139DEAFB89FCFEFBD9" box="[747,775,1080,1107]" pageId="21" pageNumber="22">Sc</collectorName>
1000165
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139C82FB88FE92FB13" collectingDate="2008-10-08" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" location="Shendurney Wildlife Sanctuary" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFE389139C82FB88FC63FBD9" box="[899,922,1081,1107]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE389139CA9FB89FC00FBD8" box="[936,1017,1080,1106]" country="India" name="Kerala" pageId="21" pageNumber="22">Kerala</collectingRegion>
,
<location id="D807601FFFE389139B02FB89FA8BFBD9" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139B02FB89FA8BFBD9" box="[1027,1394,1080,1107]" country="India" name="Shendurney Wildlife Sanctuary" pageId="21" pageNumber="22" stateProvince="Kerala">Shendurney Wildlife Sanctuary</location>
, teak plantations;
<date id="A9661004FFE389139E87FBEDFDEDFBFC" box="[390,532,1116,1142]" pageId="21" pageNumber="22" value="2008-10-08">
<collectingDate id="B922E9ECFFE389139E87FBEDFDEDFBFC" box="[390,532,1116,1142]" pageId="21" pageNumber="22" value="2008-10-08">8 Oct. 2008</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE389139D20FBEDFCB7FBFC" box="[545,846,1116,1142]" pageId="21" pageNumber="22">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFE389139C5BFBEDFB18FBFC" box="[858,1249,1116,1142]" pageId="21" pageNumber="22">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE389139A23FBEDFA97FBFC" box="[1314,1390,1116,1142]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139FBCFBCEFF26FB13" box="[189,223,1151,1177]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139FE9FBCEFEFDFB13" box="[232,260,1151,1177]" pageId="21" pageNumber="22">Sc</collectorName>
1000166
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139E8AFBCEFD84FB55" collectingDate="2019-04-14" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" county="Pasighat" elevation="1500" latitude="23.871288" location="Lengteng Wildlife Sanctuary" longLatPrecision="1" longitude="93.30618" municipality="Kawlbem" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="Mizoram">
<specimenCount id="CBDEFD4DFFE389139E8AFBCEFE5BFB13" box="[395,418,1151,1177]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE389139EB5FBCEFDDAFB13" box="[436,547,1151,1177]" country="India" name="Mizoram" pageId="21" pageNumber="22">Mizoram</collectingRegion>
,
<collectingCounty id="34064E48FFE389139D33FBCEFD61FB13" box="[562,664,1151,1177]" pageId="21" pageNumber="22">Pasighat</collectingCounty>
,
<collectingMunicipality id="3D03ACBEFFE389139DA6FBCEFCE5FB13" box="[679,796,1151,1177]" pageId="21" pageNumber="22">Kawlbem</collectingMunicipality>
,
<location id="D807601FFFE389139C2DFBCEFB70FB13" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139C2DFBCEFB70FB13" box="[812,1161,1151,1177]" country="India" county="Pasighat" latitude="23.871288" longLatPrecision="1" longitude="93.30618" municipality="Kawlbem" name="Lengteng Wildlife Sanctuary" pageId="21" pageNumber="22" stateProvince="Mizoram">Lengteng Wildlife Sanctuary</location>
;
<geoCoordinate id="B8EC5003FFE389139B99FBCCFA8BFB13" box="[1176,1394,1149,1177]" degrees="23" direction="north" minutes="52" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="16.6404" value="23.871288">23°52ʹ16.6404ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139FBCFB11FE6AFB37" box="[189,403,1184,1213]" degrees="93" direction="east" minutes="18" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="22.2408" value="93.30618">93°18ʹ22.2408ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE389139EE5FB13FD7EFB36" box="[484,647,1186,1213]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="22" unit="m" value="1500.0">
<quantity id="1A209B21FFE389139EE5FB13FDBDFB37" box="[484,580,1186,1213]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="22" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE389139D99FB13FCC3FB36" box="[664,826,1186,1212]" pageId="21" pageNumber="22" value="2019-04-14">
<collectingDate id="B922E9ECFFE389139D99FB13FCC3FB36" box="[664,826,1186,1212]" pageId="21" pageNumber="22" value="2019-04-14">14 Apr. 2019</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE389139C48FB12FB67FB37" box="[841,1182,1186,1213]" pageId="21" pageNumber="22">cow dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFE389139BACFB13FA8EFB36" box="[1197,1399,1186,1212]" pageId="21" pageNumber="22">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE389139E7BFB74FE3EFB55" box="[378,455,1221,1247]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139ED1FB74FE0BFB55" box="[464,498,1221,1247]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139EFAFB74FDEEFB55" box="[507,535,1221,1247]" pageId="21" pageNumber="22">Sc</collectorName>
1000167
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139DB1FB74FBDAFAAC" collectingDate="2018-09-30" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="131" latitude="27.969614" location="Rani Village" longLatPrecision="1" longitude="95.32392" pageId="21" pageNumber="22" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE389139DB1FB74FD03FB6A" box="[688,762,1221,1248]" count="5" pageId="21" pageNumber="22" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139C0CFB74FCAFFB6A" box="[781,854,1221,1248]" count="3" pageId="21" pageNumber="22" type="female">3 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE389139C69FB74FBA9FB55" box="[872,1104,1221,1247]" country="India" name="Arunachal Pradesh" pageId="21" pageNumber="22">Arunachal Pradesh</collectingRegion>
, agricultural field,
<location id="D807601FFFE389139A3EFB74FEECFA88" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139A3EFB74FEECFA88" country="India" latitude="27.969614" longLatPrecision="1" longitude="95.32392" name="Rani Village" pageId="21" pageNumber="22" stateProvince="Arunachal Pradesh">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFE389139E23FB56FE00FA89" box="[290,505,1255,1283]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="10.6068" value="27.969614">27°58ʹ10.6068ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139D07FB56FD20FA89" box="[518,729,1255,1283]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="26.1156" value="95.32392">95°19ʹ26.1156ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE389139C24FB58FC48FA88" box="[805,945,1256,1283]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="21" pageNumber="22" unit="m" value="131.0">
<quantity id="1A209B21FFE389139C24FB58FC89FA89" box="[805,880,1257,1283]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="21" pageNumber="22" unit="m" value="131.0">131 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE389139CBEFB58FBA7FA89" box="[959,1118,1257,1283]" pageId="21" pageNumber="22" value="2018-09-30">
<collectingDate id="B922E9ECFFE389139CBEFB58FBA7FA89" box="[959,1118,1257,1283]" pageId="21" pageNumber="22" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFE389139E0BFABDFE2AFAAC" box="[266,467,1292,1318]" pageId="21" pageNumber="22">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE389139D8FFABDFD22FAAC" box="[654,731,1292,1318]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139DE5FABDFCFFFAAC" box="[740,774,1292,1318]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139C0EFABDFCD2FAAC" box="[783,811,1292,1318]" pageId="21" pageNumber="22">Sc</collectorName>
1000168 to 1000175
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139B40FABDFA8EFAE6" collectingDate="2018-09-30" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="133" latitude="27.969172" location="Rani Village" longLatPrecision="1" longitude="95.32194" municipality="Pasighat" pageId="21" pageNumber="22" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE389139B40FABDFBA0FAAC" box="[1089,1113,1292,1318]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139B69FABDFB86FAAC" box="[1128,1151,1292,1318]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE389139B8FFABDFA8BFAAC" box="[1166,1394,1292,1318]" country="India" name="Arunachal Pradesh" pageId="21" pageNumber="22">Arunachal Pradesh</collectingRegion>
, community forest,
<collectingMunicipality id="3D03ACBEFFE389139E99FA9EFE07FAC3" box="[408,510,1327,1353]" pageId="21" pageNumber="22">Pasighat</collectingMunicipality>
,
<location id="D807601FFFE389139D09FA9EFD65FAC3" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139D09FA9EFD65FAC3" box="[520,668,1327,1353]" country="India" latitude="27.969172" longLatPrecision="1" longitude="95.32194" municipality="Pasighat" name="Rani Village" pageId="21" pageNumber="22" stateProvince="Arunachal Pradesh">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFE389139DA6FA9CFC95FAC3" box="[679,876,1325,1353]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="9.0156" value="27.969172">27°58ʹ9.0156ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139C77FA9CFBB1FAC3" box="[886,1096,1325,1353]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="18.9804" value="95.32194">95°19ʹ18.9804ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE389139B8FFA9EFAECFAC3" box="[1166,1301,1327,1353]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="21" pageNumber="22" unit="m" value="133.0">
<quantity id="1A209B21FFE389139B8FFA9EFB2EFAC3" box="[1166,1239,1327,1353]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="21" pageNumber="22" unit="m" value="133.0">133 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE389139A21FA9EFF02FAE6" pageId="21" pageNumber="22" value="2018-09-30">
<collectingDate id="B922E9ECFFE389139A21FA9EFF02FAE6" pageId="21" pageNumber="22" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE389139E08FAE2FDAFFAE7" box="[265,598,1362,1389]" pageId="21" pageNumber="22">cow dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFE389139D65FAE3FCD5FAE6" box="[612,812,1362,1388]" pageId="21" pageNumber="22">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE389139CE5FAE3FBC8FAE6" box="[996,1073,1362,1388]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139B3BFAE3FBA5FAE6" box="[1082,1116,1362,1388]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139B64FAE3FB78FAE6" box="[1125,1153,1362,1388]" pageId="21" pageNumber="22">Sc</collectorName>
1000176 to 1000177
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139FCFFAC7FEB8FA5C" collectingDate="2019-11-18" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" county="Darak" elevation="434" latitude="28.203516" location="West Siang" longLatPrecision="1" longitude="94.595985" municipality="Larmuk Pottom forest" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE389139FCFFAC7FF1EFA1A" box="[206,231,1398,1424]" count="1" pageId="21" pageNumber="22" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFE389139FF0FAC4FE2BFA05" box="[241,466,1397,1423]" country="India" name="Arunachal Pradesh" pageId="21" pageNumber="22">Arunachal Pradesh</collectingRegion>
,
<collectingCounty id="34064E48FFE389139EDDFAC4FDDAFA05" box="[476,547,1397,1423]" pageId="21" pageNumber="22">Darak</collectingCounty>
/
<collectingMunicipality id="3D03ACBEFFE389139D2DFAC4FCC8FA05" box="[556,817,1397,1424]" pageId="21" pageNumber="22">Larmuk Pottom forest</collectingMunicipality>
,
<location id="D807601FFFE389139C3AFAC7FC3BFA05" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139C3AFAC7FC3BFA05" box="[827,962,1397,1424]" country="India" county="Darak" latitude="28.203516" longLatPrecision="1" longitude="94.595985" municipality="Larmuk Pottom forest" name="West Siang" pageId="21" pageNumber="22" stateProvince="Arunachal Pradesh">West Siang</location>
;
<geoCoordinate id="B8EC5003FFE389139CCCFAC5FB5AFA1A" box="[973,1187,1396,1424]" degrees="28" direction="north" minutes="12" orientation="latitude" pageId="21" pageNumber="22" precision="1" seconds="12.6576" value="28.203516">28°12ʹ12.6576ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE389139BAFFAC5FA88FA1A" box="[1198,1393,1396,1424]" degrees="94" direction="east" minutes="35" orientation="longitude" pageId="21" pageNumber="22" precision="1" seconds="45.546" value="94.595985">94°35ʹ45.546ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE389139FFAFA28FE7DFA38" box="[251,388,1432,1459]" metricMagnitude="2" metricUnit="m" metricValue="4.34" pageId="21" pageNumber="22" unit="m" value="434.0">
<quantity id="1A209B21FFE389139FFAFA28FEBCFA39" box="[251,325,1433,1459]" metricMagnitude="2" metricUnit="m" metricValue="4.34" pageId="21" pageNumber="22" unit="m" value="434.0">434 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE389139E93FA28FDCDFA39" box="[402,564,1433,1459]" pageId="21" pageNumber="22" value="2019-11-18">
<collectingDate id="B922E9ECFFE389139E93FA28FDCDFA39" box="[402,564,1433,1459]" pageId="21" pageNumber="22" value="2019-11-18">18 Nov. 2019</collectingDate>
</date>
, mithun dung baited
<collectingMethod id="04994ED3FFE389139C2BFA29FC8DFA38" box="[810,884,1432,1458]" pageId="21" pageNumber="22">pitfall</collectingMethod>
;
<collectorName id="702D5312FFE389139C7EFA29FBBFFA38" box="[895,1094,1432,1458]" pageId="21" pageNumber="22">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE389139BF6FA28FABAFA39" box="[1271,1347,1433,1459]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139A4DFA28FA94FA39" box="[1356,1389,1433,1459]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139FBCFA0DFF20FA5C" box="[189,217,1468,1494]" pageId="21" pageNumber="22">Sc</collectorName>
1000178
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139E5FFA0DFC31FA73" collectingDate="2019-11-15" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" location="West Siang" municipality="Kaying forest" pageId="21" pageNumber="22" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE389139E5FFA0DFE5CFA5C" box="[350,421,1468,1494]" count="3" pageId="21" pageNumber="22" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE389139EB5FA0DFE00FA5C" box="[436,505,1468,1494]" count="2" pageId="21" pageNumber="22" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE389139D09FA0DFD15FA5C" box="[520,748,1468,1494]" country="India" name="Arunachal Pradesh" pageId="21" pageNumber="22">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE389139DFBFA0DFC59FA5C" box="[762,928,1468,1494]" pageId="21" pageNumber="22">Kaying forest</collectingMunicipality>
,
<location id="D807601FFFE389139CACFA0DFBCFFA5C" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139CACFA0DFBCFFA5C" box="[941,1078,1468,1494]" country="India" municipality="Kaying forest" name="West Siang" pageId="21" pageNumber="22" stateProvince="Arunachal Pradesh">West Siang</location>
;
<date id="A9661004FFE389139B44FA0AFB15FA5C" box="[1093,1260,1467,1494]" pageId="21" pageNumber="22" value="2019-11-15">
<collectingDate id="B922E9ECFFE389139B44FA0AFB15FA5C" box="[1093,1260,1467,1494]" pageId="21" pageNumber="22" value="2019-11-15">15 Nov. 2019</collectingDate>
</date>
; cow dung,
<collectorName id="702D5312FFE389139FBCFA6EFE7DFA73" box="[189,388,1503,1529]" pageId="21" pageNumber="22">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE389139D36FA6EFD7DFA73" box="[567,644,1503,1529]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139D8CFA6EFD56FA73" box="[653,687,1503,1529]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139DB9FA6EFD2DFA73" box="[696,724,1503,1529]" pageId="21" pageNumber="22">Sc</collectorName>
1000179 to 1000183
</materialsCitation>
<materialsCitation id="6DB03C99FFE389139CE0FA6EFD4FF9B5" collectingDate="2021-04-15" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" location="Intanki National Park" municipality="Bongkolong village" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="Nagaland">
<specimenCount id="CBDEFD4DFFE389139CE0FA6EFC01FA73" box="[993,1016,1503,1529]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE389139B06FA6EFB83FA73" box="[1031,1146,1503,1529]" country="India" name="Nagaland" pageId="21" pageNumber="22">Nagaland</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE389139B87FA6EFA8BFA73" box="[1158,1394,1503,1529]" pageId="21" pageNumber="22">Bongkolong village</collectingMunicipality>
, near
<location id="D807601FFFE389139FF6F9B3FE0EF996" LSID="urn:lsid:plazi:treatment:557187D2FFE289109D65FB6AFC27FC2B:D807601FFFE389139FF6F9B3FE0EF996" box="[247,503,1538,1564]" country="India" municipality="Bongkolong village" name="Intanki National Park" pageId="21" pageNumber="22" stateProvince="Nagaland">Intanki National Park</location>
, open ground;
<date id="A9661004FFE389139DA7F9B3FCBBF996" box="[678,834,1538,1565]" pageId="21" pageNumber="22" value="2021-04-15">
<collectingDate id="B922E9ECFFE389139DA7F9B3FCBBF996" box="[678,834,1538,1565]" pageId="21" pageNumber="22" value="2021-04-15">15 Apr. 2021</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFE389139BABF9B3FE96F9B5" pageId="21" pageNumber="22">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE389139EAEF994FE04F9B5" box="[431,509,1573,1599]" pageId="21" pageNumber="22">AIMB</collectionCode>
/
<collectorName id="702D5312FFE389139D07F994FDD1F9B5" box="[518,552,1573,1599]" pageId="21" pageNumber="22">Co</collectorName>
/
<collectorName id="702D5312FFE389139D30F994FDB4F9B5" box="[561,589,1573,1599]" pageId="21" pageNumber="22">Sc</collectorName>
1000184
</materialsCitation>
.
</paragraph>
<paragraph id="DD6736C4FFE389139FBCF9D6FC37F92E" blockId="21.[189,1399,1639,1700]" pageId="21" pageNumber="22">
<materialsCitation id="6DB03C99FFE389139FBCF9D6FC33F92E" collectingDate="1907-09" collectionCode="BMNH" collectorName="S. E. Asia" country="Malaysia" determinerName="Wulf." location="PERAK" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" stateProvince="Perak">
<collectingCountry id="A5CF7654FFE389139FBCF9D6FEADF90B" box="[189,340,1639,1665]" name="Malaysia" pageId="21" pageNumber="22">MALAYSIA</collectingCountry>
<specimenCount id="CBDEFD4DFFE389139E72F9D6FE73F90B" box="[371,394,1639,1665]" count="1" pageId="21" pageNumber="22" type="female"></specimenCount>
; “
<collectingRegion id="1F1CF826FFE389139EA6F9D6FDF4F90B" box="[423,525,1639,1665]" country="Malaysia" name="Perak" pageId="21" pageNumber="22">PERAK</collectingRegion>
, F.M.S.,
<date id="A9661004FFE389139D7DF9D6FD0EF90B" box="[636,759,1639,1665]" pageId="21" pageNumber="22" value="1907-09">
<collectingDate id="B922E9ECFFE389139D7DF9D6FD0EF90B" box="[636,759,1639,1665]" pageId="21" pageNumber="22" value="1907-09">Sep. 1907</collectingDate>
</date>
, Coll.; Ex. F.M.S. Museum., B.M. 1955354;
<taxonomicName id="1AD84D47FFE389139A2EF9D6FE87F92E" authority="J. Gillet" authorityName="J. Gillet" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFE389139A2EF9D6FEE3F92E" italics="true" pageId="21" pageNumber="22">Onitis falcatus</emphasis>
J. Gillet
</taxonomicName>
det.
<determinerName id="28BBDA00FFE389139EB6F93BFE03F92E" box="[439,506,1674,1700]" pageId="21" pageNumber="22">Wulf.</determinerName>
1923,
<collectorName id="702D5312FFE389139D4AF93BFD45F92E" box="[587,700,1674,1700]" pageId="21" pageNumber="22">S.E. Asia</collectorName>
”;
<collectionCode id="BBC9AE01FFE389139DD2F93BFCD2F92E" box="[723,811,1674,1700]" pageId="21" pageNumber="22">BMNH</collectionCode>
(E) 1237166
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE389109FBCF97AFAE4FDBD" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="DD6736C4FFE389139FBCF97AFEB7F96F" blockId="21.[189,334,1739,1765]" box="[189,334,1739,1765]" pageId="21" pageNumber="22">
<heading id="862F81A8FFE389139FBCF97AFEB7F96F" bold="true" box="[189,334,1739,1765]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="EFACEAD6FFE389139FBCF97AFEB7F96F" bold="true" box="[189,334,1739,1765]" pageId="21" pageNumber="22">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE389139FBCF94BFD03F89E" blockId="21.[189,762,1786,1812]" box="[189,762,1786,1812]" pageId="21" pageNumber="22">
MEASUREMENTS: TL:
<quantity id="1A209B21FFE389139EB6F94BFDC0F89E" box="[439,569,1786,1812]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="21" pageNumber="22" unit="mm" value="19.5" valueMax="23.0" valueMin="16.0">1623 mm</quantity>
; BW:
<quantity id="1A209B21FFE389139D83F94BFD0CF89E" box="[642,757,1786,1812]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="21" pageNumber="22" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">812 mm</quantity>
.
</paragraph>
<paragraph id="DD6736C4FFE389139FEBF88AFED3F8DF" blockId="21.[189,1399,1851,2019]" box="[234,298,1851,1877]" pageId="21" pageNumber="22">
<emphasis id="EFACEAD6FFE389139FEBF88AFED3F8DF" bold="true" box="[234,298,1851,1877]" pageId="21" pageNumber="22">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE389109FBCF8EFFE8AFEE3" blockId="21.[189,1399,1851,2019]" lastBlockId="22.[189,1398,265,361]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
Black, smooth, shining; clypeus elliptical, feebly excised in middle, moderately closely, finely rugulose; frontal carina widely interrupted, a short clypeal carina; pronotum finely and sparsely punctured without a median groove; elytra finely striate, 1
<superScript id="2AAD9B8CFFE389139DA6F814FD4BF839" attach="none" box="[679,690,1957,1971]" fontSize="6" pageId="21" pageNumber="22">st</superScript>
, 3
<superScript id="2AAD9B8CFFE389139DD5F815FD1BF838" attach="left" box="[724,738,1956,1970]" fontSize="6" pageId="21" pageNumber="22">rd</superScript>
and 5
<superScript id="2AAD9B8CFFE389139C35F815FCB8F838" attach="left" box="[820,833,1956,1970]" fontSize="6" pageId="21" pageNumber="22">th</superScript>
intervals raised, striae very shallow, irregular punctures on sutural intervals; pygidium opaque, smooth, unpunctured; metasternal shield smooth, feebly punctured, hairy anteriorly; protibia elongate with two or three minute teeth before the middle of the lower surface, a sharp incurved terminal process, a minute tooth near the end of the lower edge of the mesofemur.
</paragraph>
<paragraph id="DD6736C4FFE089109FBCFE23FB9CFE45" blockId="22.[189,1398,402,464]" pageId="22" pageNumber="23">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFE089109EBAFE23FDC6FE26" box="[443,575,402,428]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="22" pageNumber="23">Fig. 6EH</figureCitation>
). LP= 2, Lp=1.75; BP=1, BpB =0.5, BpT=0.25. Parameres shorter than phallobase, straight above and slightly curved below and hooked at the tip.
</paragraph>
<paragraph id="DD6736C4FFE089109FEBFE48FEBCFD99" blockId="22.[189,1309,505,568]" box="[234,325,505,531]" pageId="22" pageNumber="23">
<emphasis id="EFACEAD6FFE089109FEBFE48FEBCFD99" bold="true" box="[234,325,505,531]" pageId="22" pageNumber="23">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE089109FBCFDAFFAE4FDBD" blockId="22.[189,1309,505,568]" box="[189,1309,541,568]" pageId="22" pageNumber="23">Pronotum with stronger punctures; protibia uniformly curved with an articulated spur at the tip.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE089109FBCFDD1FC27FC2B" pageId="22" pageNumber="23" type="distribution">
<paragraph id="DD6736C4FFE089109FBCFDD1FEAFFDF0" blockId="22.[189,342,608,634]" box="[189,342,608,634]" pageId="22" pageNumber="23">
<heading id="862F81A8FFE089109FBCFDD1FEAFFDF0" bold="true" box="[189,342,608,634]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="EFACEAD6FFE089109FBCFDD1FEAFFDF0" bold="true" box="[189,342,608,634]" pageId="22" pageNumber="23">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE089109FBCFD21FC27FC2B" blockId="22.[189,1399,656,929]" pageId="22" pageNumber="23">
<collectingCountry id="A5CF7654FFE089109FBCFD21FF07FD20" box="[189,254,656,682]" name="India" pageId="22" pageNumber="23">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFE089109E11FD21FE21FD20" box="[272,472,656,682]" country="India" name="Andhra Pradesh" pageId="22" pageNumber="23">Andhra Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109EEAFD21FD2CFD20" box="[491,725,656,682]" country="India" name="Arunachal Pradesh" pageId="22" pageNumber="23">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109DE6FD21FCC3FD20" box="[743,826,656,682]" country="India" name="Assam" pageId="22" pageNumber="23">Assam</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109C4EFD21FC6BFD20" box="[847,914,656,682]" country="India" name="Bihar" pageId="22" pageNumber="23">Bihar</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109CA7FD21FBF8FD20" box="[934,1025,656,682]" country="India" name="Gujarat" pageId="22" pageNumber="23">Gujarat</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109B14FD20FB82FD21" box="[1045,1147,657,683]" country="India" name="Haryana" pageId="22" pageNumber="23">Haryana</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109B91FD21FA8BFD20" box="[1168,1394,656,682]" country="India" name="Himachal Pradesh" pageId="22" pageNumber="23">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109FBCFD02FEC0FD47" box="[189,313,691,717]" country="India" name="Jharkhand" pageId="22" pageNumber="23">Jharkhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109E50FD02FE34FD47" box="[337,461,691,717]" country="India" name="Karnataka" pageId="22" pageNumber="23">Karnataka</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109EE5FD02FDCCFD47" box="[484,565,691,717]" country="India" name="Kerala" pageId="22" pageNumber="23">Kerala</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109D4FFD02FCD8FD47" box="[590,801,691,717]" country="India" name="Madhya Pradesh" pageId="22" pageNumber="23">Madhya Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109C38FD02FC36FD47" box="[825,975,691,717]" country="India" name="Maharashtra" pageId="22" pageNumber="23">Maharashtra</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109CE6FD02FBB7FD47" box="[999,1102,691,717]" country="India" name="Manipur" pageId="22" pageNumber="23">Manipur</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109B67FD02FB12FD47" box="[1126,1259,691,717]" country="India" name="Meghalaya" pageId="22" pageNumber="23">Meghalaya</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109A05FD02FA8AFD47" box="[1284,1395,691,717]" country="India" name="Mizoram" pageId="22" pageNumber="23">Mizoram</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109FBCFD67FEC9FD7A" box="[189,304,726,752]" country="India" name="Nagaland" pageId="22" pageNumber="23">Nagaland</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109E3EFD67FE6CFD7A" box="[319,405,726,752]" country="India" name="Odisha" pageId="22" pageNumber="23">Odisha</collectingRegion>
,
<collectingCountry id="A5CF7654FFE089109EA5FD67FDD4FD7A" box="[420,557,726,752]" name="India" pageId="22" pageNumber="23">Puducherry</collectingCountry>
, Punjab,
<collectingRegion id="1F1CF826FFE089109D9CFD67FCEDFD7A" box="[669,788,726,752]" country="India" name="Rajasthan" pageId="22" pageNumber="23">Rajasthan</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109C23FD67FC4AFD7A" box="[802,947,726,752]" country="India" name="Tamil Nadu" pageId="22" pageNumber="23">Tamil Nadu</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109CC0FD67FBE2FD7A" box="[961,1051,726,752]" country="India" name="Tripura" pageId="22" pageNumber="23">Tripura</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109B28FD67FB44FD7A" box="[1065,1213,726,752]" country="India" name="Uttarakhand" pageId="22" pageNumber="23">Uttarakhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109BCDFD66FA8BFD7A" box="[1228,1394,726,753]" country="India" name="Uttar Pradesh" pageId="22" pageNumber="23">Uttar Pradesh</collectingRegion>
, New
<collectingRegion id="1F1CF826FFE089109FFEFD4BFEBAFC9E" box="[255,323,762,788]" country="India" name="Delhi" pageId="22" pageNumber="23">Delhi</collectingRegion>
,
<collectingRegion id="1F1CF826FFE089109E53FD4BFE17FC9E" box="[338,494,762,788]" country="India" name="West Bengal" pageId="22" pageNumber="23">West Bengal</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFE089109DC9FD4BFCACFC9E" box="[712,853,762,788]" name="Bangladesh" pageId="22" pageNumber="23">Bangladesh</collectingCountry>
;
<collectingCountry id="A5CF7654FFE089109C65FD4BFC45FC9E" box="[868,956,762,788]" name="Bhutan" pageId="22" pageNumber="23">Bhutan</collectingCountry>
;
<collectingCountry id="A5CF7654FFE089109CCDFD4BFBB1FC9E" box="[972,1096,762,788]" name="Cambodia" pageId="22" pageNumber="23">Cambodia</collectingCountry>
;
<collectingCountry id="A5CF7654FFE089109B56FD4BFB58FC9E" box="[1111,1185,762,788]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
;
<collectingCountry id="A5CF7654FFE089109BB1FD4BFB14FC9E" box="[1200,1261,762,788]" name="Laos" pageId="22" pageNumber="23">Laos</collectingCountry>
;
<collectingCountry id="A5CF7654FFE089109BFDFD4BFA8BFC9E" box="[1276,1394,762,788]" name="Myanmar" pageId="22" pageNumber="23">Myanmar</collectingCountry>
; Malay Peninsula;
<collectingCountry id="A5CF7654FFE089109E91FCACFE23FCBD" box="[400,474,797,823]" name="Nepal" pageId="22" pageNumber="23">Nepal</collectingCountry>
;
<collectingCountry id="A5CF7654FFE089109EE4FCACFDB5FCBD" box="[485,588,797,823]" name="Pakistan" pageId="22" pageNumber="23">Pakistan</collectingCountry>
;
<collectingCountry id="A5CF7654FFE089109D59FCACFD19FCBD" box="[600,736,797,823]" name="Philippines" pageId="22" pageNumber="23">Philippines</collectingCountry>
;
<collectingCountry id="A5CF7654FFE089109DEAFCACFCBCFCBD" box="[747,837,797,823]" name="Taiwan" pageId="22" pageNumber="23">Taiwan</collectingCountry>
;
<collectingCountry id="A5CF7654FFE089109C4EFCACFC42FCBD" box="[847,955,797,823]" name="Thailand" pageId="22" pageNumber="23">Thailand</collectingCountry>
;
<collectingCountry id="A5CF7654FFE089109CC7FCACFBD0FCBD" box="[966,1065,797,823]" name="Vietnam" pageId="22" pageNumber="23">Vietnam</collectingCountry>
(
<bibRefCitation id="B9494B35FFE089109B38FCACFA8BFCB2" author="Biswas S. &amp; Chatterjee S. K." box="[1081,1394,797,824]" pageId="22" pageNumber="23" pagination="1 - 4" refId="ref27511" refString="Biswas S. &amp; Chatterjee S. K. 1985. Insecta: Coleoptera: Scarabaeidae: Coprinae. Records of the Zoological Survey of India, Fauna of Namdapha, Arunachal Pradesh (Special Issue), 82 (1 - 4): 147 - 177. https: // doi. org / 10.26515 / rzsi / v 82 / i 1 - 4 / 1984 / 161291" type="journal article" year="1985">Biswas &amp; Chatterjee 1985</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE089109FBCFCF1FE9DFCD0" author="Chandra K." box="[189,356,832,858]" pageId="22" pageNumber="23" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000</bibRefCitation>
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<bibRefCitation id="B9494B35FFE089109C9FFCF1FB31FCD0" author="Chandra K. &amp; Ahirwar S. C." box="[926,1224,832,858]" pageId="22" pageNumber="23" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">Chandra &amp; Ahirwar 2007</bibRefCitation>
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<bibRefCitation id="B9494B35FFE089109FBCFCD2FEF3FCF7" author="Sewak R." box="[189,266,867,893]" pageId="22" pageNumber="23" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">2009b</bibRefCitation>
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).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,589 @@
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<paragraph id="DD6736C4FFE589159D69FE6CFC35FE7D" blockId="19.[616,972,477,539]" box="[616,972,477,503]" pageId="19" pageNumber="20">
<heading id="862F81A8FFE589159D69FE6CFC35FE7D" box="[616,972,477,503]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<taxonomicName id="1AD84D47FFE589159D69FE6CFC35FE7D" ID-CoL="49KG2" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[616,972,477,503]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="EFACEAD6FFE589159D69FE6CFCC9FE7D" bold="true" box="[616,816,477,503]" italics="true" pageId="19" pageNumber="20">Onitis excavatus</emphasis>
<bibRefCitation id="B9494B35FFE589159C36FE6CFC35FE7D" author="Arrow G. J." box="[823,972,477,503]" pageId="19" pageNumber="20" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
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<paragraph id="DD6736C4FFE589159DAAFDB1FC71FD90" blockId="19.[616,972,477,539]" box="[683,904,512,539]" pageId="19" pageNumber="20">
<figureCitation id="45E32A41FFE589159DAAFDB1FCF0FD91" box="[683,777,512,539]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="19" pageNumber="20">Figs 5E</figureCitation>
,
<figureCitation id="45E32A41FFE589159C15FDB1FCA6FD90" box="[788,863,512,538]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="19" pageNumber="20">6AD</figureCitation>
,
<figureCitation id="45E32A41FFE589159C6AFDB1FC71FD90" box="[875,904,512,538]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="19" pageNumber="20">18</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFE589159FBCFDF3FCA4FDD6" blockId="19.[189,861,578,604]" box="[189,861,578,604]" pageId="19" pageNumber="20">
<emphasis id="EFACEAD6FFE589159FBCFDF3FE79FDD6" box="[189,384,578,604]" italics="true" pageId="19" pageNumber="20">Onitisexcavatus</emphasis>
<bibRefCitation id="B9494B35FFE589159E84FDF3FDACFDD6" author="Arrow G. J." box="[389,597,578,604]" pageId="19" pageNumber="20" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931: 391</bibRefCitation>
(original description).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE589159FBCFD35FC33FCA0" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="DD6736C4FFE589159FBCFD35FC33FCA0" blockId="19.[189,1399,643,811]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="FDC811EAFFE589159FBCFD35FC33FCA0" pageId="19" pageNumber="20">
<taxonomicName id="1AD84D47FFE589159FBCFD35FE7CFD17" ID-CoL="49KG2" authorityName="Arrow" authorityYear="1931" box="[189,389,644,670]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="EFACEAD6FFE589159FBCFD35FE7CFD17" box="[189,389,644,670]" italics="true" pageId="19" pageNumber="20">Onitis excavatus</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFE589159EAAFD32FD6BFD17" author="Balthasar V." box="[427,658,643,670]" page="93" pageId="19" pageNumber="20" year="1935">
<bibRefCitation id="B9494B35FFE589159EAAFD32FD6BFD17" author="Balthasar V." box="[427,658,643,670]" pageId="19" pageNumber="20" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 93</bibRefCitation>
</treatmentCitation>
(monograph); 1963: 37 (monograph). —
<treatmentCitation id="5C7910D5FFE589159B98FD35FA8FFD14" author="Janssens A." box="[1177,1398,643,670]" page="50" pageId="19" pageNumber="20" year="1937">
<bibRefCitation id="B9494B35FFE589159B98FD35FA8FFD14" author="Janssens A." box="[1177,1398,643,670]" pageId="19" pageNumber="20" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 50</bibRefCitation>
</treatmentCitation>
(revision); 1938: 2 (notes). —
<treatmentCitation id="5C7910D5FFE589159D61FD17FC25FD4B" author="Biswas S. &amp; Chatterjee S. K." box="[608,988,678,705]" page="158" pageId="19" pageNumber="20" year="1985">
<bibRefCitation id="B9494B35FFE589159D61FD17FC25FD4B" author="Biswas S. &amp; Chatterjee S. K." box="[608,988,678,705]" pageId="19" pageNumber="20" pagination="1 - 4" refId="ref27511" refString="Biswas S. &amp; Chatterjee S. K. 1985. Insecta: Coleoptera: Scarabaeidae: Coprinae. Records of the Zoological Survey of India, Fauna of Namdapha, Arunachal Pradesh (Special Issue), 82 (1 - 4): 147 - 177. https: // doi. org / 10.26515 / rzsi / v 82 / i 1 - 4 / 1984 / 161291" type="journal article" year="1985">Biswas &amp; Chatterjee 1985: 158</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFE589159BBBFD17FA8FFD4A" author="Mittal I. C." box="[1210,1398,678,705]" page="46" pageId="19" pageNumber="20" year="2005">
<bibRefCitation id="B9494B35FFE589159BBBFD17FA8FFD4A" author="Mittal I. C." box="[1210,1398,678,705]" pageId="19" pageNumber="20" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005: 46</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFE589159E61FD7BFD55FD6E" author="Bezdek A. &amp; Krell F. - T." box="[352,684,713,740]" page="159" pageId="19" pageNumber="20" year="2006">
<bibRefCitation id="B9494B35FFE589159E61FD7BFD55FD6E" author="Bezdek A. &amp; Krell F. - T." box="[352,684,713,740]" pageId="19" pageNumber="20" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezdek &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). — Chandra
<emphasis id="EFACEAD6FFE589159B00FD7BFBB8FD6E" box="[1025,1089,714,740]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2012a: 52 (distribution). —
<treatmentCitation id="5C7910D5FFE589159E0EFD5CFDCBFC8D" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[271,562,749,775]" page="300" pageId="19" pageNumber="20" year="2014">
<bibRefCitation id="B9494B35FFE589159E0EFD5CFDCBFC8D" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[271,562,749,775]" pageId="19" pageNumber="20" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFE589159E7BFD5FFE4CFC8D" box="[378,437,749,775]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2014: 300
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFE589159C09FD5CFBE7FC8D" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[776,1054,749,776]" page="1037" pageId="19" pageNumber="20" year="2015">
<bibRefCitation id="B9494B35FFE589159C09FD5CFBE7FC8D" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[776,1054,749,776]" pageId="19" pageNumber="20" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFE589159C56FD5FFC6BFC8D" box="[855,914,749,775]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(checklist). —
<treatmentCitation id="5C7910D5FFE589159BD5FD5CFEEFFCA0" author="Bezdek A." page="179" pageId="19" pageNumber="20" year="2016">
<bibRefCitation id="B9494B35FFE589159BD5FD5CFEEFFCA0" author="Bezdek A." pageId="19" pageNumber="20" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFE589159EEDFCA1FD1FFCA0" author="Schoolmeesters P." box="[492,742,784,810]" pageId="19" pageNumber="20" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE589159FBCFCE0FCA7FB8E" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="DD6736C4FFE589159FBCFCE0FE3EFCE1" blockId="19.[189,455,849,875]" box="[189,455,849,875]" pageId="19" pageNumber="20">
<heading id="862F81A8FFE589159FBCFCE0FE3EFCE1" bold="true" box="[189,455,849,875]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="EFACEAD6FFE589159FBCFCE0FE3EFCE1" bold="true" box="[189,455,849,875]" pageId="19" pageNumber="20">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE589159FBCFC31FCA7FB8E" blockId="19.[189,1399,896,1028]" pageId="19" pageNumber="20">
<emphasis id="EFACEAD6FFE589159FBCFC31FE86FC10" box="[189,383,896,922]" italics="true" pageId="19" pageNumber="20">Onitisexcavatus</emphasis>
closely resembles
<taxonomicName id="1AD84D47FFE589159D5DFC31FD24FC10" baseAuthorityName="Wulfen" baseAuthorityYear="1786" box="[604,733,896,922]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFE589159D5DFC31FD24FC10" box="[604,733,896,922]" italics="true" pageId="19" pageNumber="20">O. falcatus</emphasis>
</taxonomicName>
, both placed in Group I. These two species are similar as the pronotum is smooth or sparingly punctured, and the former differs from the latter in having larger and broader body, a deep transverse excavation behind the middle of the metasternum and presence of strong sharp teeth at the extremity of mesofemur in male.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE589159FBCFB9AFCD5F855" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="DD6736C4FFE589159FBCFB9AFE15FBCF" blockId="19.[189,492,1067,1093]" box="[189,492,1067,1093]" pageId="19" pageNumber="20">
<heading id="862F81A8FFE589159FBCFB9AFE15FBCF" bold="true" box="[189,492,1067,1093]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="EFACEAD6FFE589159FBCFB9AFE15FBCF" bold="true" box="[189,492,1067,1093]" pageId="19" pageNumber="20">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE589159FEBFBEBFE9FFBFE" blockId="19.[189,1398,1114,1211]" box="[234,358,1114,1140]" pageId="19" pageNumber="20">
<emphasis id="EFACEAD6FFE589159FEBFBEBFE9FFBFE" bold="true" box="[234,358,1114,1140]" pageId="19" pageNumber="20">
<typeStatus id="02638866FFE589159FEBFBEBFE9FFBFE" box="[234,358,1114,1140]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE589159FBCFBCCFB41FB30" blockId="19.[189,1398,1114,1211]" pageId="19" pageNumber="20">
<collectingCountry id="A5CF7654FFE589159FBCFBCCFEAFFB1D" box="[189,342,1149,1175]" name="Myanmar" pageId="19" pageNumber="20">MYANMAR</collectingCountry>
<specimenCount id="CBDEFD4DFFE589159E7AFBCCFE6DFB1D" box="[379,404,1149,1175]" count="1" pageId="19" pageNumber="20" type="male"></specimenCount>
; “Tenasserim, Atkinson Coll.923, near
<taxonomicName id="1AD84D47FFE589159C8DFBCCFB77FB1D" authority=", Lansb." authorityName="Lansb." box="[908,1166,1149,1175]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ludekingi">
<emphasis id="EFACEAD6FFE589159C8DFBCCFBC8FB1D" box="[908,1073,1149,1175]" italics="true" pageId="19" pageNumber="20">O. Ludekingi,</emphasis>
Lansb.
</taxonomicName>
, but quite diff. by sculptures,
<taxonomicName id="1AD84D47FFE589159E42FB11FDFDFB30" authorityName="Arrow" authorityYear="1931" box="[323,516,1184,1210]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="excavates">
<emphasis id="EFACEAD6FFE589159E42FB11FDFDFB30" box="[323,516,1184,1210]" italics="true" pageId="19" pageNumber="20">Onitis excavates</emphasis>
</taxonomicName>
male Arr., M.E. Bacchus det. 1975”;
<collectionCode id="BBC9AE01FFE589159CBCFB10FBECFB31" box="[957,1045,1185,1211]" pageId="19" pageNumber="20">BMNH</collectionCode>
(E) 1237173.
</paragraph>
<paragraph id="DD6736C4FFE589159FEBFB50FCE2FB71" blockId="19.[189,1399,1249,2015]" box="[234,795,1249,1276]" pageId="19" pageNumber="20">
<emphasis id="EFACEAD6FFE589159FEBFB50FD9BFB71" bold="true" box="[234,610,1249,1275]" pageId="19" pageNumber="20">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFE589159D70FB53FCEAFB71" box="[625,787,1249,1276]" count="29" pageId="19" pageNumber="20" type="generic">29 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFE589159FBCFAB4FCD5F855" blockId="19.[189,1399,1249,2015]" pageId="19" pageNumber="20">
<materialsCitation id="6DB03C99FFE589159FBCFAB4FE07FAEF" collectingDate="2018-09-24" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" elevation="877" latitude="27.238258" location="Pakke" longLatPrecision="1" longitude="93.5373" pageId="19" pageNumber="20" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Arunachal Pradesh">
<collectingCountry id="A5CF7654FFE589159FBCFAB4FEF7FA95" box="[189,270,1285,1311]" name="India" pageId="19" pageNumber="20">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFE589159E2BFAB4FE96FA95" box="[298,367,1285,1311]" count="2" pageId="19" pageNumber="20" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE589159E7CFAB5FE38FA95" box="[381,449,1284,1311]" count="5" pageId="19" pageNumber="20" type="female">5 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE589159ECEFAB4FD4BFA95" box="[463,690,1285,1311]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<location id="D807601FFFE589159DBFFAB4FCF3FA95" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159DBFFAB4FCF3FA95" box="[702,778,1285,1311]" country="India" latitude="27.238258" longLatPrecision="1" longitude="93.5373" name="Pakke" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Pakke</location>
;
<geoCoordinate id="B8EC5003FFE589159C16FAB2FC2BFA95" box="[791,978,1283,1311]" degrees="27" direction="north" minutes="14" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="17.73" value="27.238258">27°14ʹ17.73″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159CDEFAB2FB61FA95" box="[991,1176,1283,1311]" degrees="93" direction="east" minutes="32" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="14.28" value="93.5373">93°32ʹ14.28″ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159BE5FAB4FA89FA95" box="[1252,1392,1285,1311]" metricMagnitude="2" metricUnit="m" metricValue="8.77" pageId="19" pageNumber="20" unit="m" value="877.0">
<quantity id="1A209B21FFE589159BE5FAB4FAD6FA95" box="[1252,1327,1285,1311]" metricMagnitude="2" metricUnit="m" metricValue="8.77" pageId="19" pageNumber="20" unit="m" value="877.0">877 m</quantity>
a.s.l.
</elevation>
; hand-picked from elephant dung;
<date id="A9661004FFE589159D54FA99FD01FAC8" box="[597,760,1320,1346]" pageId="19" pageNumber="20" value="2018-09-24">
<collectingDate id="B922E9ECFFE589159D54FA99FD01FAC8" box="[597,760,1320,1346]" pageId="19" pageNumber="20" value="2018-09-24">24 Sep. 2018</collectingDate>
</date>
;
<collectorName id="702D5312FFE589159C06FA99FB9DFAC8" box="[775,1124,1320,1346]" pageId="19" pageNumber="20">Priyadarsanan Dharma Rajan</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159A23FA99FA97FAC8" box="[1314,1390,1320,1346]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<collectorName id="702D5312FFE589159FBCFAFAFF26FAEF" box="[189,223,1355,1381]" pageId="19" pageNumber="20">Co</collectorName>
/
<collectorName id="702D5312FFE589159FE9FAFAFEFDFAEF" box="[232,260,1355,1381]" pageId="19" pageNumber="20">Sc</collectorName>
1000037 to 1000044
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159D1DFAFDFDF2FA21" collectingDate="2018-10-03" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="956" latitude="28.362144" location="Sibuk" longLatPrecision="1" longitude="95.265884" pageId="19" pageNumber="20" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159D1DFAFDFDCDFAEC" box="[540,564,1356,1382]" count="1" pageId="19" pageNumber="20" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFE589159D42FAFDFDA3FAEC" box="[579,602,1356,1382]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159D6BFAFAFCB7FAEF" box="[618,846,1355,1381]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<location id="D807601FFFE589159C5CFAFAFC5CFAEF" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159C5CFAFAFC5CFAEF" box="[861,933,1355,1381]" country="India" latitude="28.362144" longLatPrecision="1" longitude="95.265884" name="Sibuk" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Sibuk</location>
;
<geoCoordinate id="B8EC5003FFE589159CB5FAF8FB74FAEC" box="[948,1165,1353,1382]" degrees="28" direction="north" minutes="21" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="43.7184" value="28.362144">28°21ʹ43.7184ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159B9AFAF8FA88FAEC" box="[1179,1393,1353,1382]" degrees="95" direction="east" minutes="15" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="57.1968" value="95.265884">95°15ʹ57.1968ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159FFCFADFFE72FA02" box="[253,395,1390,1417]" metricMagnitude="2" metricUnit="m" metricValue="9.559999999999999" pageId="19" pageNumber="20" unit="m" value="956.0">
<quantity id="1A209B21FFE589159FFCFADFFEB0FA03" box="[253,329,1390,1417]" metricMagnitude="2" metricUnit="m" metricValue="9.559999999999999" pageId="19" pageNumber="20" unit="m" value="956.0">956 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159E9AFADFFDD5FA02" box="[411,556,1390,1416]" pageId="19" pageNumber="20" value="2018-10-03">
<collectingDate id="B922E9ECFFE589159E9AFADFFDD5FA02" box="[411,556,1390,1416]" pageId="19" pageNumber="20" value="2018-10-03">3 Oct. 2018</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFE589159C9FFADFFB9EFA02" box="[926,1127,1390,1416]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159A23FADFFA97FA02" box="[1314,1390,1390,1416]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<collectorName id="702D5312FFE589159FBCFA20FF26FA26" box="[189,223,1425,1452]" pageId="19" pageNumber="20">Co</collectorName>
/
<collectorName id="702D5312FFE589159FE9FA20FEFDFA26" box="[232,260,1425,1452]" pageId="19" pageNumber="20">Sc</collectorName>
1000045 to 1000046
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159D36FA23FCB5FA78" collectingDate="2018-10-09" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="1557" latitude="27.524204" location="Rani Village" longLatPrecision="1" longitude="27.524204" municipality="Pasighat" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159D36FA23FDB7FA26" box="[567,590,1426,1452]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159D65FA20FCB6FA21" box="[612,847,1425,1451]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
, community forest,
<collectingMunicipality id="3D03ACBEFFE589159B57FA20FB45FA21" box="[1110,1212,1425,1451]" pageId="19" pageNumber="20">Pasighat</collectingMunicipality>
,
<location id="D807601FFFE589159BD3FA20FA8BFA21" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159BD3FA20FA8BFA21" box="[1234,1394,1425,1451]" country="India" latitude="27.524204" longLatPrecision="1" longitude="27.524204" municipality="Pasighat" name="Rani Village" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFE589159FBCFA02FE7EFA45" box="[189,391,1459,1487]" degrees="27" direction="north" minutes="31" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="27.138" value="27.524204">27°31ʹ27.138ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159E94FA02FDA2FA45" box="[405,603,1459,1487]" degrees="27" direction="east" minutes="31" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="27.138" value="27.524204">27°31ʹ27.138ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159DABFA05FCBEFA45" box="[682,839,1460,1487]" metricMagnitude="3" metricUnit="m" metricValue="1.557" pageId="19" pageNumber="20" unit="m" value="1557.0">
<quantity id="1A209B21FFE589159DABFA05FCFCFA45" box="[682,773,1460,1487]" metricMagnitude="3" metricUnit="m" metricValue="1.557" pageId="19" pageNumber="20" unit="m" value="1557.0">1557 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159C56FA04FC11FA45" box="[855,1000,1461,1487]" pageId="19" pageNumber="20" value="2018-10-09">
<collectingDate id="B922E9ECFFE589159C56FA04FC11FA45" box="[855,1000,1461,1487]" pageId="19" pageNumber="20" value="2018-10-09">9 Oct. 2018</collectingDate>
</date>
; hand-picked from donkey dung;
<collectorName id="702D5312FFE589159FBCFA69FE7EFA78" box="[189,391,1496,1522]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159D4BFA69FD6EFA78" box="[586,663,1496,1522]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<collectorName id="702D5312FFE589159D9EFA69FD38FA78" box="[671,705,1496,1522]" pageId="19" pageNumber="20">Co</collectorName>
/
<collectorName id="702D5312FFE589159DCBFA69FD1FFA78" box="[714,742,1496,1522]" pageId="19" pageNumber="20">Sc</collectorName>
1000047
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159C6FFA69FC6BF9B2" collectingDate="2018-10-10" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="1532" latitude="27.523193" location="Talley Valley" longLatPrecision="1" longitude="93.839455" municipality="Ziro" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159C6FFA69FC7CFA78" box="[878,901,1496,1522]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159C97FA69FB82FA78" box="[918,1147,1496,1522]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159B8AFA69FB39FA78" box="[1163,1216,1496,1522]" pageId="19" pageNumber="20">Ziro</collectingMunicipality>
,
<location id="D807601FFFE589159BD1FA69FA8BFA78" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159BD1FA69FA8BFA78" box="[1232,1394,1496,1522]" country="India" latitude="27.523193" longLatPrecision="1" longitude="93.839455" municipality="Ziro" name="Talley Valley" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Talley Valley</location>
;
<geoCoordinate id="B8EC5003FFE589159FBCFA48FE62F99C" box="[189,411,1529,1558]" degrees="27" direction="north" minutes="31" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="23.4984" value="27.523193">27°31ʹ23.4984ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159EAFFA48FD70F99C" box="[430,649,1529,1558]" degrees="93" direction="east" minutes="50" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="22.0308" value="93.839455">93°50ʹ22.0308ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159DE3FA4AFC73F99F" box="[738,906,1531,1558]" metricMagnitude="3" metricUnit="m" metricValue="1.5319999999999998" pageId="19" pageNumber="20" unit="m" value="1532.0">
<quantity id="1A209B21FFE589159DE3FA4AFCBAF99C" box="[738,835,1531,1558]" metricMagnitude="3" metricUnit="m" metricValue="1.5319999999999998" pageId="19" pageNumber="20" unit="m" value="1532.0">1532 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159C9EFA4AFBB3F99F" box="[927,1098,1531,1557]" pageId="19" pageNumber="20" value="2018-10-10">
<collectingDate id="B922E9ECFFE589159C9EFA4AFBB3F99F" box="[927,1098,1531,1557]" pageId="19" pageNumber="20" value="2018-10-10">10 Oct. 2018</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFE589159E0BF9AFFE2AF9B2" box="[266,467,1566,1592]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159D91F9AFFD24F9B2" box="[656,733,1566,1592]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<collectorName id="702D5312FFE589159DE7F9AFFCF1F9B2" box="[742,776,1566,1592]" pageId="19" pageNumber="20">Co</collectorName>
/
<collectorName id="702D5312FFE589159C11F9AFFCD5F9B2" box="[784,812,1566,1592]" pageId="19" pageNumber="20">Sc</collectorName>
1000048
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159CB0F9AEFCB1F9F5" collectingDate="2018-10-18" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="805" latitude="27.233759" location="Mobu" longLatPrecision="1" longitude="93.53302" municipality="Sangri" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159CB0F9AEFC33F9B3" box="[945,970,1567,1593]" count="1" pageId="19" pageNumber="20" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159CD9F9AFFB45F9B2" box="[984,1212,1566,1592]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159BCBF9AFFAE0F9B2" box="[1226,1305,1566,1592]" pageId="19" pageNumber="20">Sangri</collectingMunicipality>
,
<location id="D807601FFFE589159A29F9AFFA8BF9B2" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159A29F9AFFA8BF9B2" box="[1320,1394,1566,1592]" country="India" latitude="27.233759" longLatPrecision="1" longitude="93.53302" municipality="Sangri" name="Mobu" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Mobu</location>
;
<geoCoordinate id="B8EC5003FFE589159FBCF9F1FE7FF9D6" box="[189,390,1600,1628]" degrees="27" direction="north" minutes="14" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="1.5324" value="27.233759">27°14ʹ1.5324ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159E92F9F1FD91F9D6" box="[403,616,1600,1628]" degrees="93" direction="east" minutes="31" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="58.8576" value="93.53302">93°31ʹ58.8576ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159DB5F9F0FCB8F9D1" box="[692,833,1601,1628]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="19" pageNumber="20" unit="m" value="805.0">
<quantity id="1A209B21FFE589159DB5F9F0FCF9F9D6" box="[692,768,1601,1628]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="19" pageNumber="20" unit="m" value="805.0">805 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159C51F9F3FC16F9D6" box="[848,1007,1602,1628]" pageId="19" pageNumber="20" value="2018-10-18">
<collectingDate id="B922E9ECFFE589159C51F9F3FC16F9D6" box="[848,1007,1602,1628]" pageId="19" pageNumber="20" value="2018-10-18">18 Oct. 2018</collectingDate>
</date>
; hand-picked from mithun dung;
<collectorName id="702D5312FFE589159FBCF9D4FE7FF9F5" box="[189,390,1637,1663]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159D45F9D4FD68F9F5" box="[580,657,1637,1663]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<collectorName id="702D5312FFE589159D9BF9D4FD45F9F5" box="[666,700,1637,1663]" pageId="19" pageNumber="20">Co</collectorName>
/
<collectorName id="702D5312FFE589159DC4F9D4FD18F9F5" box="[709,737,1637,1663]" pageId="19" pageNumber="20">Sc</collectorName>
1000049
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159C67F9D4FC1BF94C" collectingDate="2018-09-23" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="164" latitude="26.94896" location="Pakke Tiger Reserve" longLatPrecision="1" longitude="92.98454" municipality="Seijosa" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159C67F9D4FC84F9F5" box="[870,893,1637,1663]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159C8CF9D4FB88F9F5" box="[909,1137,1637,1663]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159B81F9D4FB21F9F5" box="[1152,1240,1637,1663]" pageId="19" pageNumber="20">Seijosa</collectingMunicipality>
,
<location id="D807601FFFE589159BE7F9D4FEE6F928" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159BE7F9D4FEE6F928" country="India" latitude="26.94896" longLatPrecision="1" longitude="92.98454" municipality="Seijosa" name="Pakke Tiger Reserve" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Pakke Tiger Reserve</location>
;
<geoCoordinate id="B8EC5003FFE589159E31F937FDF5F928" box="[304,524,1670,1698]" degrees="26" direction="north" minutes="56" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="56.2596" value="26.94896">26°56ʹ56.2596ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159D1FF937FD11F928" box="[542,744,1670,1698]" degrees="92" direction="east" minutes="59" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="4.3764" value="92.98454">92°59ʹ4.3764ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159C3CF939FC2BF928" box="[829,978,1672,1698]" metricMagnitude="2" metricUnit="m" metricValue="1.64" pageId="19" pageNumber="20" unit="m" value="164.0">
<quantity id="1A209B21FFE589159C3CF939FC74F928" box="[829,909,1672,1698]" metricMagnitude="2" metricUnit="m" metricValue="1.64" pageId="19" pageNumber="20" unit="m" value="164.0">164 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159CE4F939FB77F928" box="[997,1166,1672,1698]" pageId="19" pageNumber="20" value="2018-09-23">
<collectingDate id="B922E9ECFFE589159CE4F939FB77F928" box="[997,1166,1672,1698]" pageId="19" pageNumber="20" value="2018-09-23">23 Sep. 2018</collectingDate>
</date>
; hand-picked from elephant dung;
<collectorName id="702D5312FFE589159E6EF91AFDCAF94F" box="[367,563,1707,1733]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159DDEF91AFCD5F94C" box="[735,812,1707,1734]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<collectorName id="702D5312FFE589159C34F91AFCAEF94C" box="[821,855,1707,1734]" pageId="19" pageNumber="20">Co</collectorName>
/
<collectorName id="702D5312FFE589159C5EF91AFC82F94C" box="[863,891,1707,1734]" pageId="19" pageNumber="20">Sc</collectorName>
1000050
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159CF9F91AFB47F881" collectingDate="2018-09-24" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="190" latitude="26.941978" location="Pakke Tiger Reserve" longLatPrecision="1" longitude="92.9716" municipality="Seijosa" pageId="19" pageNumber="20" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159CF9F91AFBC3F94C" box="[1016,1082,1707,1734]" count="6" pageId="19" pageNumber="20" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE589159B44F91AFB7EF94C" box="[1093,1159,1707,1734]" count="4" pageId="19" pageNumber="20" type="female">4 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE589159B93F91AFA8BF94F" box="[1170,1394,1707,1733]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159FBCF97FFEECF962" box="[189,277,1742,1768]" pageId="19" pageNumber="20">Seijosa</collectingMunicipality>
,
<location id="D807601FFFE589159E1EF97FFDEBF963" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159E1EF97FFDEBF963" box="[287,530,1742,1769]" country="India" latitude="26.941978" longLatPrecision="1" longitude="92.9716" municipality="Seijosa" name="Pakke Tiger Reserve" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Pakke Tiger Reserve</location>
;
<geoCoordinate id="B8EC5003FFE589159D1CF97CFD1AF963" box="[541,739,1741,1769]" degrees="26" direction="north" minutes="56" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="31.128" value="26.941978">26°56ʹ31.128ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159DEFF97CFC5BF963" box="[750,930,1741,1769]" degrees="92" direction="east" minutes="58" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="17.76" value="92.9716">92°58ʹ17.76ʺ E</geoCoordinate>
; elev.
<quantity id="1A209B21FFE589159CE8F97FFBCCF963" box="[1001,1077,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="19" pageNumber="20" unit="m" value="190.0">
<elevation id="56F5D1F7FFE589159CE8F97FFBCCF963" box="[1001,1077,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="19" pageNumber="20" unit="m" value="190.0">190 m</elevation>
</quantity>
;
<date id="A9661004FFE589159B41F97FFB22F962" box="[1088,1243,1742,1768]" pageId="19" pageNumber="20" value="2018-09-24">
<collectingDate id="B922E9ECFFE589159B41F97FFB22F962" box="[1088,1243,1742,1768]" pageId="19" pageNumber="20" value="2018-09-24">24 Sep. 2018</collectingDate>
</date>
; hand-picked from elephant dung;
<collectorName id="702D5312FFE589159EB2F940FD83F881" box="[435,634,1777,1803]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159C2FF940FC82F886" box="[814,891,1777,1804]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<collectorName id="702D5312FFE589159C85F940FC5FF886" box="[900,934,1777,1804]" pageId="19" pageNumber="20">Co</collectorName>
/
<collectorName id="702D5312FFE589159CAEF940FC32F886" box="[943,971,1777,1804]" pageId="19" pageNumber="20">Sc</collectorName>
1000051 to 1000060
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159BD9F943FB61F8D8" collectingDate="2018-09-29" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="135" latitude="27.968342" location="Rani village" longLatPrecision="1" longitude="95.320145" municipality="Pasighat" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159BD9F943FB08F886" box="[1240,1265,1778,1804]" count="1" pageId="19" pageNumber="20" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159BFDF940FEE4F8A5" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159E2BF8A4FE69F8A5" box="[298,400,1813,1839]" pageId="19" pageNumber="20">Pasighat</collectingMunicipality>
,
<location id="D807601FFFE589159E9CF8A4FDC9F8A5" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159E9CF8A4FDC9F8A5" box="[413,560,1813,1839]" country="India" latitude="27.968342" longLatPrecision="1" longitude="95.320145" municipality="Pasighat" name="Rani village" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Rani village</location>
;
<geoCoordinate id="B8EC5003FFE589159D3FF8A2FCFEF8A5" box="[574,775,1811,1839]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="6.0276" value="27.968342">27°58ʹ6.0276ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159C15F8A2FC10F8A5" box="[788,1001,1811,1839]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="12.5328" value="95.320145">95°19ʹ12.5328ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159B37F8A5FB3AF8A5" box="[1078,1219,1812,1839]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="19" pageNumber="20" unit="m" value="135.0">
<quantity id="1A209B21FFE589159B37F8A5FB78F8A5" box="[1078,1153,1812,1839]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="19" pageNumber="20" unit="m" value="135.0">135 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159BD0F8A4FA88F8A5" box="[1233,1393,1813,1839]" pageId="19" pageNumber="20" value="2018-09-29">
<collectingDate id="B922E9ECFFE589159BD0F8A4FA88F8A5" box="[1233,1393,1813,1839]" pageId="19" pageNumber="20" value="2018-09-29">29 Sep. 2018</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFE589159D1BF889FD18F8D8" box="[538,737,1848,1874]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159C94F889FC1BF8D8" box="[917,994,1848,1874]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<collectorName id="702D5312FFE589159CEDF889FBF7F8D8" box="[1004,1038,1848,1874]" pageId="19" pageNumber="20">Co</collectorName>
/
<collectorName id="702D5312FFE589159B16F889FBCAF8D8" box="[1047,1075,1848,1874]" pageId="19" pageNumber="20">Sc</collectorName>
1000061
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159BB3F889FB89F812" collectingDate="2018-09-30" collectionCode="AIMB" collectorName="Harsha Malhotra &amp; Co &amp; Sc" country="India" elevation="125" latitude="27.970259" location="Rani village" longLatPrecision="1" longitude="95.322365" municipality="Pasighat" pageId="19" pageNumber="20" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159BB3F889FB33F8D8" box="[1202,1226,1848,1874]" count="1" pageId="19" pageNumber="20" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFE589159BD9F889FB16F8D8" box="[1240,1263,1848,1874]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159BFDF889FEE4F8FF" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159E28F8EAFE76F8FF" box="[297,399,1883,1909]" pageId="19" pageNumber="20">Pasighat</collectingMunicipality>
,
<location id="D807601FFFE589159E9AF8EAFDD4F8FF" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159E9AF8EAFDD4F8FF" box="[411,557,1883,1909]" country="India" latitude="27.970259" longLatPrecision="1" longitude="95.322365" municipality="Pasighat" name="Rani village" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Rani village</location>
;
<geoCoordinate id="B8EC5003FFE589159D3BF8E8FCE9F8FC" box="[570,784,1881,1910]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="12.9324" value="27.970259">27°58ʹ12.9324ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159C1CF8E8FC09F8FC" box="[797,1008,1881,1910]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="20.5284" value="95.322365">95°19ʹ20.5284ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159B3AF8EAFB3FF8FF" box="[1083,1222,1883,1910]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="19" pageNumber="20" unit="m" value="125.0">
<quantity id="1A209B21FFE589159B3AF8EAFB7FF8FC" box="[1083,1158,1883,1910]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="19" pageNumber="20" unit="m" value="125.0">125 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159BD2F8EAFA8BF8FF" box="[1235,1394,1883,1909]" pageId="19" pageNumber="20" value="2018-09-30">
<collectingDate id="B922E9ECFFE589159BD2F8EAFA8BF8FF" box="[1235,1394,1883,1909]" pageId="19" pageNumber="20" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
; cow dung;
<collectorName id="702D5312FFE589159E49F8CFFDEAF812" box="[328,531,1918,1944]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159DD6F8CFFCDDF812" box="[727,804,1918,1944]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<collectorName id="702D5312FFE589159C2FF8CFFCA9F812" box="[814,848,1918,1944]" pageId="19" pageNumber="20">Co</collectorName>
/
<collectorName id="702D5312FFE589159C58F8CFFC8CF812" box="[857,885,1918,1944]" pageId="19" pageNumber="20">Sc</collectorName>
1000061 to 1000062
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159B90F8CEFCD1F855" collectingDate="2021-04-25" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara &amp; Co &amp; Sc" country="India" location="Singphan Elephant Reserve" municipality="Mon District" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Nagaland">
<specimenCount id="CBDEFD4DFFE589159B90F8CEFB51F813" box="[1169,1192,1919,1945]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159BBAF8CFFAD7F812" box="[1211,1326,1918,1944]" country="India" name="Nagaland" pageId="19" pageNumber="20">Nagaland</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159A3EF8CEFEE1F831" pageId="19" pageNumber="20">Mon District</collectingMunicipality>
,
<location id="D807601FFFE589159E29F810FD8EF836" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159E29F810FD8EF836" box="[296,631,1953,1980]" country="India" municipality="Mon District" name="Singphan Elephant Reserve" pageId="19" pageNumber="20" stateProvince="Nagaland">Singphan Elephant Reserve</location>
;
<date id="A9661004FFE589159D86F810FCD2F836" box="[647,811,1953,1980]" pageId="19" pageNumber="20" value="2021-04-25">
<collectingDate id="B922E9ECFFE589159D86F810FCD2F836" box="[647,811,1953,1980]" pageId="19" pageNumber="20" value="2021-04-25">25 Apr. 2021</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFE589159BA7F813FE96F855" pageId="19" pageNumber="20">Seena Narayanan Karimbumkara</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159D20F874FD96F855" box="[545,623,1989,2015]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<collectorName id="702D5312FFE589159D79F874FD63F855" box="[632,666,1989,2015]" pageId="19" pageNumber="20">Co</collectorName>
/
<collectorName id="702D5312FFE589159DA5F874FD39F855" box="[676,704,1989,2015]" pageId="19" pageNumber="20">Sc</collectorName>
1000063
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE289129FBCFEB9FC66FC36" pageId="20" pageNumber="21" type="description">
<paragraph id="DD6736C4FFE289129FBCFEB9FEB7FEA8" blockId="20.[189,334,264,290]" box="[189,334,264,290]" pageId="20" pageNumber="21">
<heading id="862F81A8FFE289129FBCFEB9FEB7FEA8" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="EFACEAD6FFE289129FBCFEB9FEB7FEA8" bold="true" box="[189,334,264,290]" pageId="20" pageNumber="21">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFE86FD11FEDB" blockId="20.[189,744,311,338]" box="[189,744,311,338]" pageId="20" pageNumber="21">MEASUREMENTS (in mm). TL=1925; BW= 13.</paragraph>
<paragraph id="DD6736C4FFE289129FEBFEC8FED3FE19" blockId="20.[189,1399,377,720]" box="[234,298,377,403]" pageId="20" pageNumber="21">
<emphasis id="EFACEAD6FFE289129FEBFEC8FED3FE19" bold="true" box="[234,298,377,403]" pageId="20" pageNumber="21">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFE2DFB4EFD45" blockId="20.[189,1399,377,720]" pageId="20" pageNumber="21">Black, shining, elytra and pygidium subopaque; clypeus short, elliptical, moderately closely and finely rugulose; frontal carina widely interrupted in the middle, a short transverse clypeal carina anteriorly, a small tubercle behind, vertex irregularly granular; pronotum very smooth with irregular minute punctures anteriorly, longitudinal groove absent; elytra finely striate with 1st, 3rd and 5th intervals elevated, narrower, rest of the intervals flat; pygidium smooth, unpunctured; metasternum with very deep transverse excavation in the middle, metasternal shield thickly hairy anteriorly, smooth, shining behind; protibia elongate with a serrate carina upon anterior half beneath, four short external teeth and a sharp incurved terminal process; mesofemur with a rounded lobe in the middle of hind edge, two sharp teeth near extremity above and one beneath; trochanters of hind legs acutely produced.</paragraph>
<paragraph id="DD6736C4FFE289129FBCFD46FAE3FCBE" blockId="20.[189,1399,759,820]" pageId="20" pageNumber="21">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFE289129EA8FD46FDD0FC9B" box="[425,553,759,785]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="20" pageNumber="21">Fig. 6AD</figureCitation>
). LP=3.03, Lp=2.93; BP=1.5, BpB=1.14, BpT=0.46. Parameres shorter than phallobase, straight above and slightly curved outwards below and hooked slightly at the tip.
</paragraph>
<paragraph id="DD6736C4FFE289129FEBFCEAFEBCFCFF" blockId="20.[189,1399,859,956]" box="[234,325,859,885]" pageId="20" pageNumber="21">
<emphasis id="EFACEAD6FFE289129FEBFCEAFEBCFCFF" bold="true" box="[234,325,859,885]" pageId="20" pageNumber="21">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFCCEFC66FC36" blockId="20.[189,1399,859,956]" pageId="20" pageNumber="21">Legs simple, frontal carina slightly diagonal; punctation of the pronotum heavier and denser, elytral striae deeper than in male; metasternum without transverse pit.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE289129FBCFC52FCB4FB1F" pageId="20" pageNumber="21" type="distribution">
<paragraph id="DD6736C4FFE289129FBCFC52FEAFFC77" blockId="20.[189,342,995,1021]" box="[189,342,995,1021]" pageId="20" pageNumber="21">
<heading id="862F81A8FFE289129FBCFC52FEAFFC77" bold="true" box="[189,342,995,1021]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="EFACEAD6FFE289129FBCFC52FEAFFC77" bold="true" box="[189,342,995,1021]" pageId="20" pageNumber="21">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFBA3FCB4FB1F" blockId="20.[189,1399,1042,1174]" pageId="20" pageNumber="21">
<collectingCountry id="A5CF7654FFE289129FBCFBA3FF07FBA6" box="[189,254,1042,1068]" name="India" pageId="20" pageNumber="21">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFE289129E04FBA3FE1DFBA6" box="[261,484,1042,1068]" country="India" name="Arunachal Pradesh" pageId="20" pageNumber="21">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129EEDFBA3FDC6FBA6" box="[492,575,1042,1068]" country="India" name="Assam" pageId="20" pageNumber="21">Assam</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129D49FBA3FCE6FBA6" box="[584,799,1042,1068]" country="India" name="Himachal Pradesh" pageId="20" pageNumber="21">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129C29FBA3FC54FBA6" box="[808,941,1042,1068]" country="India" name="Meghalaya" pageId="20" pageNumber="21">Meghalaya</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129CB6FBA3FBD3FBA6" box="[951,1066,1042,1068]" country="India" name="Nagaland" pageId="20" pageNumber="21">Nagaland</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129B35FBA3FB31FBA6" box="[1076,1224,1042,1068]" country="India" name="Uttarakhand" pageId="20" pageNumber="21">Uttarakhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129BD0FBA3FA8AFBA6" box="[1233,1395,1042,1068]" country="India" name="Uttar Pradesh" pageId="20" pageNumber="21">Uttar Pradesh</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFE289129E84FB84FE36FBC5" box="[389,463,1077,1103]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129EDCFB84FDAAFBC5" box="[477,595,1077,1103]" name="Myanmar" pageId="20" pageNumber="21">Myanmar</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129D61FB84FD53FBC5" box="[608,682,1077,1103]" name="Nepal" pageId="20" pageNumber="21">Nepal</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129DB6FB84FCE7FBC5" box="[695,798,1077,1103]" name="Pakistan" pageId="20" pageNumber="21">Pakistan</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129C2AFB84FC7CFBC5" box="[811,901,1077,1103]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129C93FB84FC07FBC5" box="[914,1022,1077,1103]" name="Thailand" pageId="20" pageNumber="21">Thailand</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129B0BFB84FB94FBC5" box="[1034,1133,1077,1103]" name="Vietnam" pageId="20" pageNumber="21">Vietnam</collectingCountry>
(
<bibRefCitation id="B9494B35FFE289129B7EFB84FF02FBF9" author="Biswas S. &amp; Chatterjee S. K." pageId="20" pageNumber="21" pagination="1 - 4" refId="ref27511" refString="Biswas S. &amp; Chatterjee S. K. 1985. Insecta: Coleoptera: Scarabaeidae: Coprinae. Records of the Zoological Survey of India, Fauna of Namdapha, Arunachal Pradesh (Special Issue), 82 (1 - 4): 147 - 177. https: // doi. org / 10.26515 / rzsi / v 82 / i 1 - 4 / 1984 / 161291" type="journal article" year="1985">Biswas &amp; Chatterjee 1985</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE289129E08FBE9FE61FBF9" author="Mittal I. C." box="[265,408,1112,1139]" pageId="20" pageNumber="21" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE289129EA7FBE9FD53FBF8" author="Bezdek A. &amp; Krell F. - T." box="[422,682,1112,1138]" pageId="20" pageNumber="21" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
; Chandra
<emphasis id="EFACEAD6FFE289129C24FBE8FC98FBF8" box="[805,865,1112,1138]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2012a;
<bibRefCitation id="B9494B35FFE289129CC2FBE9FB4AFBF8" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[963,1203,1112,1138]" pageId="20" pageNumber="21" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFE289129B31FBE8FB95FBF8" box="[1072,1132,1112,1138]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE289129BBEFBE9FA8BFBF9" author="Ali S. M. S. &amp; Naeem M. &amp; Baig F. &amp; Shazad A. &amp; Zia A." box="[1215,1394,1112,1139]" pageId="20" pageNumber="21" pagination="1 - 6" refId="ref26882" refString="Ali S. M. S., Naeem M., Baig F., Shazad A. &amp; Zia A. 2015. New records, distributional notes and species diversity of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Pothohar Plateau of Punjab. Pakistan. Journal of Entomology and Zoology Studies 3 (3): 1 - 6. Available from: https: // www. entomoljournal. com / archives / 2015 / vol 3 issue 3 / PartA / 3 - 3 - 23. pdf [accessed 27 Nov. 2019]" type="journal article" year="2015">
Ali
<emphasis id="EFACEAD6FFE289129BEFFBE8FAD2FBF8" box="[1262,1323,1112,1138]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE289129FBCFBCAFE75FB1C" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[189,396,1147,1174]" pageId="20" pageNumber="21" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFE289129E0DFBCDFEBEFB1F" box="[268,327,1147,1173]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE289129E99FBCAFDCCFB1F" author="Bezdek A." box="[408,565,1147,1173]" pageId="20" pageNumber="21" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE289129D43FBCAFCB8FB1F" author="Schoolmeesters P." box="[578,833,1147,1173]" pageId="20" pageNumber="21" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,388 @@
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<mods:title id="47C2032495E3705A5487BC54F042C6C1">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<subSubSection id="95C2654FFFE789179D7BFEB8FC43FEAE" box="[634,954,265,292]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="DD6736C4FFE789179D7BFEB8FC43FEAE" blockId="17.[634,954,265,334]" box="[634,954,265,292]" pageId="17" pageNumber="18">
<heading id="862F81A8FFE789179D7BFEB8FC43FEAE" box="[634,954,265,292]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
<taxonomicName id="1AD84D47FFE789179D7BFEB8FC43FEAE" ID-CoL="49KFD" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[634,954,265,292]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFE789179D7BFEB8FCDCFEA9" bold="true" box="[634,805,265,291]" italics="true" pageId="17" pageNumber="18">Onitis crassus</emphasis>
Sharp, 1875
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<subSubSection id="95C2654FFFE789179DD4FE82FCD7FE1C" pageId="17" pageNumber="18" type="description">
<paragraph id="DD6736C4FFE789179DD4FE82FCA7FEC7" blockId="17.[634,954,265,334]" box="[725,862,307,334]" pageId="17" pageNumber="18">
<figureCitation id="45E32A41FFE789179DD4FE82FCCCFEC4" box="[725,821,307,334]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="17" pageNumber="18">Figs 5D</figureCitation>
,
<figureCitation id="45E32A41FFE789179C40FE82FCA7FEC7" box="[833,862,307,333]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="17" pageNumber="18">18</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCFECDFCD7FE1C" blockId="17.[189,814,379,406]" box="[189,814,379,406]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FBCFECDFE9DFE1C" box="[189,356,380,406]" italics="true" pageId="17" pageNumber="18">Onitiscrassus</emphasis>
Sharp, 1875: 48 (original description).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE789179FBCFE74FDF0FD32" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="DD6736C4FFE789179FBCFE74FAFDFDE1" blockId="17.[189,1399,452,697]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="FDC811EAFFE789179FBCFE74FAFDFDE1" pageId="17" pageNumber="18">
<taxonomicName id="1AD84D47FFE789179FBCFE74FE9DFE54" ID-CoL="49KFD" authorityName="Sharp" authorityYear="1875" box="[189,356,453,479]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFE789179FBCFE74FE9DFE54" box="[189,356,453,479]" italics="true" pageId="17" pageNumber="18">Onitis crassus</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFE789179E7EFE75FDB0FE54" author="Arrow G. J." box="[383,585,452,478]" page="400" pageId="17" pageNumber="18" year="1931">
<bibRefCitation id="B9494B35FFE789179E7EFE75FDB0FE54" author="Arrow G. J." box="[383,585,452,478]" pageId="17" pageNumber="18" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 400</bibRefCitation>
</treatmentCitation>
(keys and description). —
<treatmentCitation id="5C7910D5FFE789179C89FE74FBA2FE54" author="Janssens A." box="[904,1115,452,479]" page="73" pageId="17" pageNumber="18" year="1937">
<bibRefCitation id="B9494B35FFE789179C89FE74FBA2FE54" author="Janssens A." box="[904,1115,452,479]" pageId="17" pageNumber="18" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 73</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFE789179A09FE75FEADFD88" author="Balthasar V." page="45" pageId="17" pageNumber="18" year="1963">
<bibRefCitation id="B9494B35FFE789179A09FE75FEADFD88" author="Balthasar V." pageId="17" pageNumber="18" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar 1963: 45</bibRefCitation>
</treatmentCitation>
(monograph). —
<treatmentCitation id="5C7910D5FFE789179D2EFE56FCEDFD8B" author="Chandra K." box="[559,788,487,513]" page="360" pageId="17" pageNumber="18" year="2000">
<bibRefCitation id="B9494B35FFE789179D2EFE56FCEDFD8B" author="Chandra K." box="[559,788,487,513]" pageId="17" pageNumber="18" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000: 360</bibRefCitation>
</treatmentCitation>
(distribution); 2008: 176 (distribution). —
<treatmentCitation id="5C7910D5FFE789179A1EFE56FE28FDAF" author="Bezdek A. &amp; Krell F. - T." page="159" pageId="17" pageNumber="18" year="2006">
<bibRefCitation id="B9494B35FFE789179A1EFE56FE28FDAF" author="Bezdek A. &amp; Krell F. - T." pageId="17" pageNumber="18" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFE789179DB4FDBBFBD6FDAF" author="Chandra K. &amp; Ahirwar S. C." box="[693,1071,522,549]" page="287" pageId="17" pageNumber="18" year="2007">
<bibRefCitation id="B9494B35FFE789179DB4FDBBFBD6FDAF" author="Chandra K. &amp; Ahirwar S. C." box="[693,1071,522,549]" pageId="17" pageNumber="18" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">Chandra &amp; Ahirwar 2007: 287</bibRefCitation>
</treatmentCitation>
(distribution). — Chandra
<emphasis id="EFACEAD6FFE789179FEBFD9EFED0FDC2" box="[234,297,558,584]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2012b: 29 (distribution). —
<treatmentCitation id="5C7910D5FFE789179D8EFD9FFC38FDC2" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[655,961,558,584]" page="299" pageId="17" pageNumber="18" year="2014">
<bibRefCitation id="B9494B35FFE789179D8EFD9FFC38FDC2" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[655,961,558,584]" pageId="17" pageNumber="18" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFE789179DFCFD9EFCC5FDC2" box="[765,828,558,584]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2014: 299
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFE789179BA1FD9FFA8EFDC2" author="Ali S. M. S. &amp; Naeem M. &amp; Baig F. &amp; Shazad A. &amp; Zia A." box="[1184,1399,558,585]" page="4" pageId="17" pageNumber="18" year="2015">
<bibRefCitation id="B9494B35FFE789179BA1FD9FFA8EFDC2" author="Ali S. M. S. &amp; Naeem M. &amp; Baig F. &amp; Shazad A. &amp; Zia A." box="[1184,1399,558,585]" pageId="17" pageNumber="18" pagination="1 - 6" refId="ref26882" refString="Ali S. M. S., Naeem M., Baig F., Shazad A. &amp; Zia A. 2015. New records, distributional notes and species diversity of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Pothohar Plateau of Punjab. Pakistan. Journal of Entomology and Zoology Studies 3 (3): 1 - 6. Available from: https: // www. entomoljournal. com / archives / 2015 / vol 3 issue 3 / PartA / 3 - 3 - 23. pdf [accessed 27 Nov. 2019]" type="journal article" year="2015">
Ali
<emphasis id="EFACEAD6FFE789179BD1FD9EFAF6FDC2" box="[1232,1295,558,584]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2015: 4
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFE789179EB9FDE0FD36FDE1" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[440,719,593,620]" page="1037" pageId="17" pageNumber="18" year="2015">
<bibRefCitation id="B9494B35FFE789179EB9FDE0FD36FDE1" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[440,719,593,620]" pageId="17" pageNumber="18" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFE789179D06FDE3FDBBFDE1" box="[519,578,593,619]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(checklist). —
<treatmentCitation id="5C7910D5FFE789179C84FDE0FBA3FDE1" author="Bezdek A." box="[901,1114,593,619]" page="179" pageId="17" pageNumber="18" year="2016">
<bibRefCitation id="B9494B35FFE789179C84FDE0FBA3FDE1" author="Bezdek A." box="[901,1114,593,619]" pageId="17" pageNumber="18" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCFDCAFDF0FD32" blockId="17.[189,1399,452,697]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FBCFDCAFE9CFD1F" box="[189,357,635,661]" italics="true" pageId="17" pageNumber="18">Onitisvischnu</emphasis>
<bibRefCitation id="B9494B35FFE789179E83FDCAFD28FD1F" author="Preudhomme de Borre C. F. P. A." box="[386,721,635,662]" pageId="17" pageNumber="18" refId="ref30731" refString="Preudhomme de Borre C. F. P. A. 1881. Bulletin ou Comptes-Rendus des Seances de la Societe entomologique de Belgique XXXIX." type="book" year="1881">Preudhomme de Borre 1881</bibRefCitation>
: xxxix (synonymised by
<bibRefCitation id="B9494B35FFE789179CF5FDCAFB7FFD1F" author="Arrow G. J." box="[1012,1158,635,661]" pageId="17" pageNumber="18" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931</bibRefCitation>
). —
<bibRefCitation id="B9494B35FFE789179BBFFDCAFEDCFD33" author="Schoolmeesters P." pageId="17" pageNumber="18" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE789179FBCFD56FE9AFC4E" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="DD6736C4FFE789179FBCFD56FE3FFC8B" blockId="17.[189,454,743,769]" box="[189,454,743,769]" pageId="17" pageNumber="18">
<heading id="862F81A8FFE789179FBCFD56FE3FFC8B" bold="true" box="[189,454,743,769]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="EFACEAD6FFE789179FBCFD56FE3FFC8B" bold="true" box="[189,454,743,769]" pageId="17" pageNumber="18">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCFCACFE9AFC4E" blockId="17.[189,1399,796,964]" pageId="17" pageNumber="18">
<taxonomicName id="1AD84D47FFE789179FBCFCACFE91FCBC" authorityName="Sharp" authorityYear="1875" box="[189,360,797,823]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFE789179FBCFCACFE91FCBC" box="[189,360,797,823]" italics="true" pageId="17" pageNumber="18">Onitis crassus</emphasis>
</taxonomicName>
is closer to
<taxonomicName id="1AD84D47FFE789179D06FCACFD5AFCBC" box="[519,675,797,823]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFE789179D06FCACFD5AFCBC" box="[519,675,797,823]" italics="true" pageId="17" pageNumber="18">O. castaneus</emphasis>
</taxonomicName>
in having hairy pygidium and uninterrupted clypeo-frontal carina but differs from the latter in having short scanty setae on the pygidium; frontal tubercle being placed behind the carina while in
<taxonomicName id="1AD84D47FFE789179D5AFCD2FD0FFCF7" box="[603,758,867,893]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFE789179D5AFCD2FD0FFCF7" box="[603,758,867,893]" italics="true" pageId="17" pageNumber="18">O. castaneus</emphasis>
</taxonomicName>
it is placed on the carina and the anterior margin of clypeus being gently excised in the middle while in the latter it is reflexed and straight. See diagnosis of
<taxonomicName id="1AD84D47FFE789179FDCFC1BFEA5FC49" authorityName="Lansberge" authorityYear="1875" box="[221,348,937,964]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFE789179FDCFC1BFEA5FC49" box="[221,348,937,964]" italics="true" pageId="17" pageNumber="18">O. brahma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE789179FBCFC43FCECFA5F" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="DD6736C4FFE789179FBCFC43FE15FB81" blockId="17.[189,492,1009,1036]" box="[189,492,1009,1036]" pageId="17" pageNumber="18">
<heading id="862F81A8FFE789179FBCFC43FE15FB81" bold="true" box="[189,492,1009,1036]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="EFACEAD6FFE789179FBCFC43FE15FB81" bold="true" box="[189,492,1009,1036]" pageId="17" pageNumber="18">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE789179FEBFB96FEA5FBCB" blockId="17.[234,348,1063,1089]" box="[234,348,1063,1089]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FEBFB96FEA5FBCB" bold="true" box="[234,348,1063,1089]" pageId="17" pageNumber="18">
<typeStatus id="02638866FFE789179FEBFB96FEA5FBCB" box="[234,348,1063,1089]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCFBE3FBD5FB05" blockId="17.[189,1398,1105,1167]" pageId="17" pageNumber="18">
<materialsCitation id="6DB03C99FFE789179FBCFBE3FBD5FB05" collectionCode="MNHN" collectorName="Ex. Musaeo D. Sharp &amp; G. J. Arrow &amp; A. Janssens" country="India" location="INDIA" pageId="17" pageNumber="18" specimenCode="EC4813" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingCountry id="A5CF7654FFE789179FBCFBE3FEF7FBE6" box="[189,270,1106,1132]" name="India" pageId="17" pageNumber="18">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFE789179E29FBE3FEC6FBE6" box="[296,319,1106,1132]" count="1" pageId="17" pageNumber="18" type="female"></specimenCount>
; “
<emphasis id="EFACEAD6FFE789179E5AFBE3FDFBFBE6" box="[347,514,1106,1132]" italics="true" pageId="17" pageNumber="18">Onitiscrassus</emphasis>
Type D.S.;
<collectorName id="702D5312FFE789179D91FBE3FC74FBE1" box="[656,909,1105,1132]" pageId="17" pageNumber="18">Ex. Musaeo D. Sharp</collectorName>
1890,
<collectorName id="702D5312FFE789179CDEFBE3FB90FBE6" box="[991,1129,1106,1132]" pageId="17" pageNumber="18">G. J. Arrow</collectorName>
vid. 1928,
<collectorName id="702D5312FFE789179BECFBE3FA8FFBE6" box="[1261,1398,1106,1132]" pageId="17" pageNumber="18">A. Janssens</collectorName>
vid., 1936:
<emphasis id="EFACEAD6FFE789179E42FBC4FE13FB05" box="[323,490,1141,1167]" italics="true" pageId="17" pageNumber="18">Onitiscrassus</emphasis>
Sharp;
<typeStatus id="02638866FFE789179D44FBC4FD4BFB05" box="[581,690,1141,1167]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
(Red Label)”;
<collectionCode id="BBC9AE01FFE789179C62FBC4FC44FB05" box="[867,957,1141,1167]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
<specimenCode id="8D7E9EBFFFE789179CC5FBC4FBD1FB05" box="[964,1064,1141,1167]" collectionCode="EC" pageId="17" pageNumber="18">EC4813</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFE789179FEBFB0CFCF5FB5D" blockId="17.[234,780,1213,1239]" box="[234,780,1213,1239]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FEBFB0CFD9BFB5D" bold="true" box="[234,610,1213,1239]" pageId="17" pageNumber="18">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFE789179D70FB0CFCFDFB5D" box="[625,772,1213,1239]" count="4" pageId="17" pageNumber="18" type="generic">4 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFE789179FBCFB59FCECFA5F" blockId="17.[189,1399,1255,1493]" pageId="17" pageNumber="18">
<materialsCitation id="6DB03C99FFE789179FBCFB59FC44FAAF" collectionCode="BMNH" country="India" location="INDIA" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFE789179FBCFB59FEF7FA88" box="[189,270,1256,1282]" name="India" pageId="17" pageNumber="18">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFE789179E2DFB59FEBCFA88" box="[300,325,1256,1282]" count="1" pageId="17" pageNumber="18" type="male"></specimenCount>
; “Determined from description G.J.A.
<taxonomicName id="1AD84D47FFE789179C19FB59FBC6FA88" authority=", de Borre" authorityName="de Borre" box="[792,1087,1255,1282]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vischnu">
<emphasis id="EFACEAD6FFE789179C19FB59FC3CFA8B" box="[792,965,1255,1282]" italics="true" pageId="17" pageNumber="18">Onitis Vischnu</emphasis>
, de Borre
</taxonomicName>
;
<taxonomicName id="1AD84D47FFE789179B51FB59FECEFAAF" authority="de Borre, Compared" authorityName="de Borre, Compared" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vischnu">
<emphasis id="EFACEAD6FFE789179B51FB59FB04FA8B" box="[1104,1277,1255,1282]" italics="true" pageId="17" pageNumber="18">Onitis Vischnu</emphasis>
de Borre, Compared
</taxonomicName>
with
<typeStatus id="02638866FFE789179E80FABAFE4DFAAF" box="[385,436,1291,1317]" pageId="17" pageNumber="18">type</typeStatus>
G.J.A., 6745, 7030”;
<collectionCode id="BBC9AE01FFE789179DC2FABAFCE2FAAF" box="[707,795,1291,1317]" pageId="17" pageNumber="18">BMNH</collectionCode>
(E) 1237163
</materialsCitation>
<materialsCitation id="6DB03C99FFE789179CDDFABAFAC2FAC2" collectionCode="BMNH" collectorName="U. P. C. Beeson." country="India" location="River Suswa" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFE789179CDDFABAFC0AFAAF" box="[988,1011,1291,1317]" count="1" pageId="17" pageNumber="18" type="female"></specimenCount>
; “Dehra Dun Coll. B.M. 1931 2.,
<location id="D807601FFFE789179FE5FA9FFE84FAC2" LSID="urn:lsid:plazi:treatment:557187D2FFE789159D7BFEB8FB02FE12:D807601FFFE789179FE5FA9FFE84FAC2" box="[228,381,1326,1352]" country="India" name="River Suswa" pageId="17" pageNumber="18">River Suswa</location>
, Dehradun,
<collectorName id="702D5312FFE789179D11FA9FFD34FAC2" box="[528,717,1326,1352]" pageId="17" pageNumber="18">U.P. C. Beeson.</collectorName>
XI.30; Cattle dung on sand”;
<collectionCode id="BBC9AE01FFE789179B45FA9FFB65FAC2" box="[1092,1180,1326,1352]" pageId="17" pageNumber="18">BMNH</collectionCode>
(E) 1237164
</materialsCitation>
<materialsCitation id="6DB03C99FFE789179A59FA9FFA8FFA3B" collectorName="O. Vishnu &amp; Coll. J. Thomson" country="India" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFE789179A59FA9FFA88FAC2" box="[1368,1393,1326,1352]" count="1" pageId="17" pageNumber="18" type="male"></specimenCount>
; “A, Janssens vid. 1936:
<taxonomicName id="1AD84D47FFE789179EDAFAE0FD1BFAE1" authority="Sharp" authorityName="Sharp" authorityYear="1875" box="[475,738,1361,1387]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFE789179EDAFAE0FD6FFAE1" box="[475,662,1361,1387]" italics="true" pageId="17" pageNumber="18">ONITIS crassus</emphasis>
Sharp
</taxonomicName>
;
<emphasis id="EFACEAD6FFE789179DF2FAE3FC62FAE6" box="[755,923,1362,1388]" italics="true" pageId="17" pageNumber="18">Onitiscrassus</emphasis>
Shp. Revis. Arrow 1926;
<emphasis id="EFACEAD6FFE789179BCEFAE3FA8EFAE6" box="[1231,1399,1362,1388]" italics="true" pageId="17" pageNumber="18">Onitiscrassus</emphasis>
Shp. Compared with
<typeStatus id="02638866FFE789179EB9FAC4FE12FA05" box="[440,491,1397,1423]" pageId="17" pageNumber="18">type</typeStatus>
G.J.A.; cf. Ann. Soc. Ent. Belg. Xxv, 1881, C.R. p. 39; Ex-Typis de
<taxonomicName id="1AD84D47FFE789179A17FAC5FEF9FA38" authority="Borre" authorityName="Borre" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vischnu">Vischnu Borre</taxonomicName>
; det: Predhomme Borre,
<taxonomicName id="1AD84D47FFE789179D29FA29FCE4FA38" authority="Borre" authorityName="Borre" box="[552,797,1431,1458]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vischnu">
<emphasis id="EFACEAD6FFE789179D29FA29FD2AFA3B" box="[552,723,1431,1458]" italics="true" pageId="17" pageNumber="18">Onitis Vischnu</emphasis>
Borre
</taxonomicName>
;
<emphasis id="EFACEAD6FFE789179C2DFA29FC67FA3B" box="[812,926,1431,1458]" italics="true" pageId="17" pageNumber="18">
<collectorName id="702D5312FFE789179C2DFA29FC67FA3B" box="[812,926,1431,1458]" pageId="17" pageNumber="18">O. Vishnu</collectorName>
</emphasis>
, Reiche, India, 3050,
<collectorName id="702D5312FFE789179BA1FA26FA94FA3B" box="[1184,1389,1431,1458]" pageId="17" pageNumber="18">Coll. J. Thomson</collectorName>
</materialsCitation>
<materialsCitation id="6DB03C99FFE789179FCFFA0AFCE8FA5F" box="[206,785,1467,1493]" collectionCode="IRSNB" collectorName="O. Vishnu &amp; Coll. J. Thomson" country="India" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFE789179FCFFA0AFF1CFA5F" box="[206,229,1467,1493]" count="1" pageId="17" pageNumber="18" type="female"></specimenCount>
; same collection data as for preceding;
<collectionCode id="BBC9AE01FFE789179DB6FA0AFCE8FA5F" box="[695,785,1467,1493]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="17" pageNumber="18">IRSNB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE789179FBCF9B2FA8EF863" pageId="17" pageNumber="18" type="description">
<paragraph id="DD6736C4FFE789179FBCF9B2FEB7F997" blockId="17.[189,334,1539,1565]" box="[189,334,1539,1565]" pageId="17" pageNumber="18">
<heading id="862F81A8FFE789179FBCF9B2FEB7F997" bold="true" box="[189,334,1539,1565]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="EFACEAD6FFE789179FBCF9B2FEB7F997" bold="true" box="[189,334,1539,1565]" pageId="17" pageNumber="18">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCF988FCCBF9DE" blockId="17.[189,818,1593,1620]" box="[189,818,1593,1620]" pageId="17" pageNumber="18">MEASUREMENTS (in mm). TL=15.521; BW=910.5.</paragraph>
<paragraph id="DD6736C4FFE789179FEBF930FED3F911" blockId="17.[234,298,1665,1691]" box="[234,298,1665,1691]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FEBF930FED3F911" bold="true" box="[234,298,1665,1691]" pageId="17" pageNumber="18">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCF91AFBBFF8FF" blockId="17.[189,1399,1707,1910]" pageId="17" pageNumber="18">Black; clypeus rugulose, anterior margin strongly elevated; gently excised in the middle; frontal carina entire, curved, with a tubercle behind; pronotum strongly and closely punctured, rugose anteriorly; elytra finely striate, intervals angularly elevated along the middle, finely, fairly closely punctured; pygidium opaque, rugulose with short setae; metasternal shield smooth, shining, feebly, sparsely punctured, sides of metasternum rugosely punctured, with long hairs; front legs elongate with a blunt process on its trochanter; hind femur with a hooked process at the middle of its lower edge.</paragraph>
<paragraph id="DD6736C4FFE789179FEBF815FEBCF834" blockId="17.[234,325,1956,1982]" box="[234,325,1956,1982]" pageId="17" pageNumber="18">
<emphasis id="EFACEAD6FFE789179FEBF815FEBCF834" bold="true" box="[234,325,1956,1982]" pageId="17" pageNumber="18">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE789179FBCF87FFA8EF863" blockId="17.[189,1399,1998,2025]" box="[189,1399,1998,2025]" pageId="17" pageNumber="18">Clypeal margin scarcely excised in the middle; protibia broad with four blunt teeth and an articulated spur.</paragraph>
</subSubSection>
<caption id="89A7664CFFE489149FBFF8D4FDB7F863" pageId="18" pageNumber="19" startId="18.[190,234,1893,1919]" targetBox="[295,1246,340,1832]" targetPageId="18">
<paragraph id="DD6736C4FFE489149FBFF8D4FDB7F863" blockId="18.[190,1407,1893,2025]" pageId="18" pageNumber="19">
<emphasis id="EFACEAD6FFE489149FBFF8D4FEEBF8F5" bold="true" box="[190,274,1893,1919]" pageId="18" pageNumber="19">Fig. 6.</emphasis>
Aedeagus, lateral, dorsal, ventral and apical views.
<emphasis id="EFACEAD6FFE489149CACF8D4FC1EF8F5" bold="true" box="[941,999,1893,1920]" pageId="18" pageNumber="19">AD</emphasis>
.
<taxonomicName id="1AD84D47FFE489149CFAF8D7FA80F8F5" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[1019,1401,1893,1920]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="EFACEAD6FFE489149CFAF8D7FB30F8F5" box="[1019,1225,1894,1920]" italics="true" pageId="18" pageNumber="19">Onitis excavatus</emphasis>
<bibRefCitation id="B9494B35FFE489149BD2F8D7FA80F8F5" author="Arrow G. J." box="[1235,1401,1893,1920]" pageId="18" pageNumber="19" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000037.
<emphasis id="EFACEAD6FFE489149ECEF838FDF3F829" bold="true" box="[463,522,1929,1955]" pageId="18" pageNumber="19">EH</emphasis>
.
<taxonomicName id="1AD84D47FFE489149D19F838FCA7F828" authority="(Wulfen, 1786)" baseAuthorityName="Wulfen" baseAuthorityYear="1786" box="[536,862,1928,1955]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFE489149D19F838FD64F828" box="[536,669,1928,1955]" italics="true" pageId="18" pageNumber="19">O. falcatus</emphasis>
(
<bibRefCitation id="B9494B35FFE489149DADF839FCACF828" author="Wulfen X." box="[684,853,1928,1954]" pageId="18" pageNumber="19" refId="ref31404" refString="Wulfen X. 1786. Descriptiones quorumdam Capensium insectorum. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1786">Wulfen, 1786</bibRefCitation>
)
</taxonomicName>
, AIMB/Co/Sc1000070.
<emphasis id="EFACEAD6FFE489149B7DF838FB52F829" bold="true" box="[1148,1195,1929,1955]" pageId="18" pageNumber="19">IL</emphasis>
.
<taxonomicName id="1AD84D47FFE489149BB8F838FA81F828" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1209,1400,1928,1955]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bhomorensis" status="sp. nov.">
<emphasis id="EFACEAD6FFE489149BB8F838FA81F828" box="[1209,1400,1928,1955]" italics="true" pageId="18" pageNumber="19">O. bhomorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFE489149FBFF81DFED8F84C" box="[190,289,1964,1990]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, NIM/NBAIR/COL/ONIT/H-271023A.
<emphasis id="EFACEAD6FFE489149C08F81DFCBEF84C" bold="true" box="[777,839,1964,1990]" pageId="18" pageNumber="19">MP</emphasis>
.
<taxonomicName id="1AD84D47FFE489149C5AF81DFAE8F84C" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[859,1297,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFE489149C5AF81DFBB2F84C" box="[859,1099,1964,1990]" italics="true" pageId="18" pageNumber="19">O. punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFE489149B59F81DFAE8F84C" author="Janssens A." box="[1112,1297,1964,1990]" pageId="18" pageNumber="19" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
, AIMB/ Co/Sc1000492. (Images: AIMB).
</paragraph>
</caption>
<subSubSection id="95C2654FFFE589159FBCFEB9FB02FE12" pageId="19" pageNumber="20" type="distribution">
<paragraph id="DD6736C4FFE589159FBCFEB9FEAFFEA8" blockId="19.[189,342,264,290]" box="[189,342,264,290]" pageId="19" pageNumber="20">
<heading id="862F81A8FFE589159FBCFEB9FEAFFEA8" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="EFACEAD6FFE589159FBCFEB9FEAFFEA8" bold="true" box="[189,342,264,290]" pageId="19" pageNumber="20">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE589159FBCFE86FB02FE12" blockId="19.[189,1398,311,408]" pageId="19" pageNumber="20">
<collectingCountry id="A5CF7654FFE589159FBCFE86FF07FEDB" box="[189,254,311,337]" name="India" pageId="19" pageNumber="20">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFE589159E15FE86FE1FFEDB" box="[276,486,311,337]" country="India" name="Madhya Pradesh" pageId="19" pageNumber="20">Madhya Pradesh</collectingRegion>
, Punjab,
<collectingRegion id="1F1CF826FFE589159D69FE86FD05FEDB" box="[616,764,311,337]" country="India" name="Uttarakhand" pageId="19" pageNumber="20">Uttarakhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFE589159C13FE89FC3BFEDB" box="[786,962,311,338]" country="India" name="Uttar Pradesh" pageId="19" pageNumber="20">Uttar Pradesh</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFE589159BB2FE86FB04FEDB" box="[1203,1277,311,337]" name="Nepal" pageId="19" pageNumber="20">Nepal</collectingCountry>
;
<collectingCountry id="A5CF7654FFE589159A12FE86FA8FFEDB" box="[1299,1398,311,337]" name="Pakistan" pageId="19" pageNumber="20">Pakistan</collectingCountry>
(
<bibRefCitation id="B9494B35FFE589159FC4FEEBFE89FEFE" author="Chandra K." box="[197,368,346,372]" pageId="19" pageNumber="20" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE589159E7DFEEBFD82FEFE" author="Bezdek A. &amp; Krell F. - T." box="[380,635,346,372]" pageId="19" pageNumber="20" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE589159D89FEEBFC4FFEFE" author="Chandra K. &amp; Ahirwar S. C." box="[648,950,346,372]" pageId="19" pageNumber="20" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">Chandra &amp; Ahirwar 2007</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE589159CC2FEEBFB95FEFE" author="Chandra K." box="[963,1132,346,372]" pageId="19" pageNumber="20" pagination="159 - 186" refId="ref28328" refString="Chandra K. 2008. Insecta: Coleoptera. Zoological Survey of India, Faunal Diversity of Jabalpur District, Madhya Pradesh: 159 - 186." type="book chapter" year="2008">Chandra 2008</bibRefCitation>
; Chandra
<emphasis id="EFACEAD6FFE589159BE2FEEAFAE7FEFE" box="[1251,1310,346,372]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2012b;
<bibRefCitation id="B9494B35FFE589159FBCFECCFE51FE12" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[189,424,381,408]" pageId="19" pageNumber="20" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFE589159E29FECFFE9AFE1D" box="[296,355,381,407]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE589159EB3FECCFDA6FE12" author="Ali S. M. S. &amp; Naeem M. &amp; Baig F. &amp; Shazad A. &amp; Zia A." box="[434,607,381,408]" pageId="19" pageNumber="20" pagination="1 - 6" refId="ref26882" refString="Ali S. M. S., Naeem M., Baig F., Shazad A. &amp; Zia A. 2015. New records, distributional notes and species diversity of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Pothohar Plateau of Punjab. Pakistan. Journal of Entomology and Zoology Studies 3 (3): 1 - 6. Available from: https: // www. entomoljournal. com / archives / 2015 / vol 3 issue 3 / PartA / 3 - 3 - 23. pdf [accessed 27 Nov. 2019]" type="journal article" year="2015">
Ali
<emphasis id="EFACEAD6FFE589159EDEFECFFDE3FE1D" box="[479,538,381,407]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE589159D6AFECFFCC3FE12" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[619,826,381,408]" pageId="19" pageNumber="20" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFE589159DBBFECFFD0CFE1D" box="[698,757,381,407]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE589159C47FECCFC1DFE1D" author="Bezdek A." box="[838,996,381,407]" pageId="19" pageNumber="20" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE589159CF1FECCFB16FE12" author="Schoolmeesters P." box="[1008,1263,381,408]" pageId="19" pageNumber="20" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="1F476A55C1A001343856BC075915C0D9" ID-DOI="10.5852/ejt.2024.956.2657" ID-ISSN="2118-9773" ID-ZooBank="9DCDD521-909D-4436-8495-086A11C50DA0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725988271530" checkinUser="plazi" docAuthor="Karimbumkara, Seena Narayanan &amp; Priyadarsanan, Dharma Rajan" docDate="2024" docId="557187D2FFEB89189D54F9EBFB5AF9A0" docLanguage="en" docName="EJT.2024.956.1-61.pdf" docOrigin="European Journal of Taxonomy 956" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onitis naviauxi Cambefort 1988" docType="treatment" docVersion="1" lastPageNumber="31" masterDocId="A948FFAAFFF689069F01FFB1FFF9FF8A" masterDocTitle="Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India" masterLastPageNumber="61" masterPageNumber="1" pageNumber="30" updateTime="1725988442359" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F57E9D66268369C611A22208FCD3A47D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="36248BDBDFB12E1217D813CDC0FCC06C">
<mods:title id="16F2D57B908DFA8EC692005F1025D1CB">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
</mods:titleInfo>
<mods:name id="76685A060078D066007B5C3BFDFE3772" type="personal">
<mods:role id="70EB9CA6A89DB0E8BBC4128B60D2FE90">
<mods:roleTerm id="5FBC70652B925200326A6B31DC8C44C8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D6E0D9AF04CBD559A2D4B4FFDE51FDB">Karimbumkara, Seena Narayanan</mods:namePart>
<mods:nameIdentifier id="FDB8F5941612A2CC5E280F0B04775663" type="LSID">A6B50103-8D6D-4977-9111-EF8BA4AA0477</mods:nameIdentifier>
<mods:affiliation id="40C2B6EE3B62B67852527CC442E07831">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="6C5EBA4F1DFAA9485F6A7B020C2F504C" type="email">seena.narayanan@atree.org</mods:nameIdentifier>
</mods:name>
<mods:name id="3FB7BC95EE762AFBB41143BDFE3D3DAB" type="personal">
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<mods:namePart id="6CC097E69411E304E6428CFEC3A6BEEA">Priyadarsanan, Dharma Rajan</mods:namePart>
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<mods:affiliation id="1D323B8E8F42B8BCC8BFB9B43F43EFA2">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
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<mods:title id="D0F65CEA6112AFD8EA8EF301F5A97309">European Journal of Taxonomy</mods:title>
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<mods:part id="BABC1D0F6C78A8FCA745D336A56E48D0">
<mods:date id="C53E0B41D0FA92061F903DB8415208AF">2024</mods:date>
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<mods:number id="6BE53622620ACB1497DC528CC395323F">2024-09-10</mods:number>
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<mods:detail id="79C41CE4F002DCC881B4E679CD52D604" type="volume">
<mods:number id="E12E214988566CFD4566944BE694E2FA">956</mods:number>
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<mods:start id="080CE32FE5C75C94B26A0373B2E008A1">1</mods:start>
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<mods:url id="71088CF967484033D4F32ACCEC7430C2">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287</mods:url>
</mods:location>
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<mods:identifier id="FDFFF0DB44EEF20E019AE42E04AE0B45" type="DOI">10.5852/ejt.2024.956.2657</mods:identifier>
<mods:identifier id="8C9EA5D9361D6AF0558331C440F28E70" type="ISSN">2118-9773</mods:identifier>
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<treatment id="557187D2FFEB89189D54F9EBFB5AF9A0" LSID="urn:lsid:plazi:treatment:557187D2FFEB89189D54F9EBFB5AF9A0" httpUri="http://treatment.plazi.org/id/557187D2FFEB89189D54F9EBFB5AF9A0" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="95C2654FFFEB891B9D54F9EBFC26F9FE" box="[597,991,1626,1652]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="DD6736C4FFEB891B9D54F9EBFC26F9FE" blockId="29.[597,991,1626,1687]" box="[597,991,1626,1652]" pageId="29" pageNumber="30">
<heading id="862F81A8FFEB891B9D54F9EBFC26F9FE" box="[597,991,1626,1652]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
<taxonomicName id="1AD84D47FFEB891B9D54F9EBFC26F9FE" ID-CoL="49KJD" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[597,991,1626,1652]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFEB891B9D54F9EBFCE9F9FE" bold="true" box="[597,784,1626,1652]" italics="true" pageId="29" pageNumber="30">Onitis naviauxi</emphasis>
<bibRefCitation id="B9494B35FFEB891B9C16F9EBFC26F9FE" author="Cambefort Y." box="[791,991,1626,1652]" pageId="29" pageNumber="30" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEB891B9DA9F9CCFC7BF952" pageId="29" pageNumber="30" type="description">
<paragraph id="DD6736C4FFEB891B9DA9F9CCFC75F91D" blockId="29.[597,991,1626,1687]" box="[680,908,1661,1687]" pageId="29" pageNumber="30">
<figureCitation id="45E32A41FFEB891B9DA9F9CCFCD8F91D" box="[680,801,1661,1687]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="29" pageNumber="30">Figs 4EF</figureCitation>
,
<figureCitation id="45E32A41FFEB891B9C2CF9CCFC9BF91D" box="[813,866,1661,1687]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="29" pageNumber="30">10D</figureCitation>
,
<figureCitation id="45E32A41FFEB891B9C6EF9CCFC75F91D" box="[879,908,1661,1687]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." pageId="29" pageNumber="30">19</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFEB891B9FBCF90EFC7BF952" blockId="29.[189,898,1726,1753]" box="[189,898,1726,1753]" pageId="29" pageNumber="30">
<emphasis id="EFACEAD6FFEB891B9FBCF90EFE88F953" box="[189,369,1727,1753]" italics="true" pageId="29" pageNumber="30">Onitisnaviauxi</emphasis>
<bibRefCitation id="B9494B35FFEB891B9E78F90FFD82F953" author="Cambefort Y." box="[377,635,1726,1753]" pageId="29" pageNumber="30" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988: 189</bibRefCitation>
(original description).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEB891B9FBCF8B1FD14F8B7" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="DD6736C4FFEB891B9FBCF8B1FD14F8B7" blockId="29.[189,1399,1792,1854]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="FDC811EAFFEB891B9FBCF8B1FD14F8B7" pageId="29" pageNumber="30">
<taxonomicName id="1AD84D47FFEB891B9FBCF8B1FE81F890" ID-CoL="49KJD" authorityName="Cambefort" authorityYear="1988" box="[189,376,1792,1818]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFEB891B9FBCF8B1FE81F890" box="[189,376,1792,1818]" italics="true" pageId="29" pageNumber="30">Onitis naviauxi</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFEB891B9EA5F8B1FD01F891" author="Bezdek A. &amp; Krell F. - T." box="[420,760,1792,1819]" page="159" pageId="29" pageNumber="30" year="2006">
<bibRefCitation id="B9494B35FFEB891B9EA5F8B1FD01F891" author="Bezdek A. &amp; Krell F. - T." box="[420,760,1792,1819]" pageId="29" pageNumber="30" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFEB891B9CE3F8B1FB3FF890" author="Bezdek A." box="[994,1222,1792,1818]" page="179" pageId="29" pageNumber="30" year="2016">
<bibRefCitation id="B9494B35FFEB891B9CE3F8B1FB3FF890" author="Bezdek A." box="[994,1222,1792,1818]" pageId="29" pageNumber="30" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFEB891B9E0EF892FDF3F8B7" author="Schoolmeesters P." box="[271,522,1827,1853]" pageId="29" pageNumber="30" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEB89189FBCF8D5FE7CFD06" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="DD6736C4FFEB891B9FBCF8D5FE3EF8F4" blockId="29.[189,455,1892,1918]" box="[189,455,1892,1918]" pageId="29" pageNumber="30">
<heading id="862F81A8FFEB891B9FBCF8D5FE3EF8F4" bold="true" box="[189,455,1892,1918]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="EFACEAD6FFEB891B9FBCF8D5FE3EF8F4" bold="true" box="[189,455,1892,1918]" pageId="29" pageNumber="30">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEB89189FBCF822FD4DFECC" blockId="29.[189,1399,1939,2001]" lastBlockId="30.[189,1398,265,327]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<bibRefCitation id="B9494B35FFEB891B9FBCF822FE77F827" author="Cambefort Y." box="[189,398,1939,1965]" pageId="29" pageNumber="30" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort (1988)</bibRefCitation>
classified
<emphasis id="EFACEAD6FFEB891B9D07F822FD41F827" box="[518,696,1939,1965]" italics="true" pageId="29" pageNumber="30">Onitisnaviauxi</emphasis>
in Group III of
<bibRefCitation id="B9494B35FFEB891B9C68F825FBDCF827" author="Janssens A." box="[873,1061,1939,1966]" pageId="29" pageNumber="30" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens (1937)</bibRefCitation>
. It is similar to
<taxonomicName id="1AD84D47FFEB891B9BD0F822FA8EF827" authorityName="Biswas" authorityYear="1980" box="[1233,1399,1939,1965]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFEB891B9BD0F822FA8EF827" box="[1233,1399,1939,1965]" italics="true" pageId="29" pageNumber="30">O. assamensis</emphasis>
</taxonomicName>
in having vertex with punctures and clypeal carina being placed away from vertex. It is distinct from the latter in having shorter clypeal carina and the male protibia having 34 teeth on the inferomedial ridge, while in
<taxonomicName id="1AD84D47FFE889189E25FE9CFE32FECC" authorityName="Biswas" authorityYear="1980" box="[292,459,301,327]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFE889189E25FE9CFE32FECC" box="[292,459,301,327]" italics="true" pageId="30" pageNumber="31">O. assamensis</emphasis>
</taxonomicName>
there are 67 teeth.
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFEDEFD31FE03" blockId="30.[189,712,367,394]" box="[189,712,367,394]" pageId="30" pageNumber="31">
<emphasis id="EFACEAD6FFE889189FBCFEDEFE15FE03" bold="true" box="[189,492,367,393]" pageId="30" pageNumber="31">Type material examined</emphasis>
(based on images)
</paragraph>
<paragraph id="DD6736C4FFE889189FEBFE2EFEA5FE33" blockId="30.[189,1399,415,513]" box="[234,348,415,441]" pageId="30" pageNumber="31">
<emphasis id="EFACEAD6FFE889189FEBFE2EFEA5FE33" bold="true" box="[234,348,415,441]" pageId="30" pageNumber="31">
<typeStatus id="02638866FFE889189FEBFE2EFEA5FE33" box="[234,348,415,441]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFE75FE7FFD8B" blockId="30.[189,1399,415,513]" pageId="30" pageNumber="31">
<materialsCitation id="6DB03C99FFE889189FBCFE75FE7FFD8B" collectionCode="MNHN" country="Nepal" determinerName="Y. Cambefort" location="NEPAL" pageId="30" pageNumber="31" specimenCode="EC4807" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="A5CF7654FFE889189FBCFE75FEE1FE54" box="[189,280,452,478]" name="Nepal" pageId="30" pageNumber="31">NEPAL</collectingCountry>
<specimenCount id="CBDEFD4DFFE889189E34FE75FEB7FE54" box="[309,334,452,478]" count="1" pageId="30" pageNumber="31" type="male"></specimenCount>
; “
<emphasis id="EFACEAD6FFE889189E6BFE75FDD9FE54" box="[362,544,452,478]" italics="true" pageId="30" pageNumber="31">Onitisnaviauxi</emphasis>
<taxonomicNameLabel id="F49F57ADFFE889189D2BFE75FD92FE54" box="[554,619,452,478]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
<determinerName id="28BBDA00FFE889189D75FE75FCEAFE54" box="[628,787,452,478]" pageId="30" pageNumber="31">Y. Cambefort</determinerName>
det. 1987, Nepal: Sauraha. Alt. 150, 19.4.82 RN”;
<collectionCode id="BBC9AE01FFE889189FBCFE56FEEEFD8B" box="[189,279,487,513]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
<specimenCode id="8D7E9EBFFFE889189E1FFE56FE7BFD8B" box="[286,386,487,513]" collectionCode="EC" pageId="30" pageNumber="31">EC4807</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFE889189FEBFD9BFEA5FDCE" blockId="30.[189,1399,554,652]" box="[234,348,554,580]" pageId="30" pageNumber="31">
<emphasis id="EFACEAD6FFE889189FEBFD9BFEA5FDCE" bold="true" box="[234,348,554,580]" pageId="30" pageNumber="31">
<typeStatus id="02638866FFE889189FEBFD9BFEA5FDCE" box="[234,348,554,580]" pageId="30" pageNumber="31" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFDFEFE7CFD06" blockId="30.[189,1399,554,652]" pageId="30" pageNumber="31">
<materialsCitation id="6DB03C99FFE889189FBCFDFEFE87FD06" collectionCode="MNHN" country="Nepal" determinerName="Y. Cambefort" location="NEPAL" pageId="30" pageNumber="31" specimenCode="EC4808" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="A5CF7654FFE889189FBCFDFEFEE1FDE3" box="[189,280,591,617]" name="Nepal" pageId="30" pageNumber="31">NEPAL</collectingCountry>
<specimenCount id="CBDEFD4DFFE889189E37FDFEFEB4FDE3" box="[310,333,591,617]" count="1" pageId="30" pageNumber="31" type="female"></specimenCount>
; “
<emphasis id="EFACEAD6FFE889189E6DFDFEFDDAFDE3" box="[364,547,591,617]" italics="true" pageId="30" pageNumber="31">Onitisnaviauxi</emphasis>
<taxonomicNameLabel id="F49F57ADFFE889189D2CFDFEFD96FDE3" box="[557,623,591,617]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
<determinerName id="28BBDA00FFE889189D6EFDFEFCF6FDE2" box="[623,783,590,617]" pageId="30" pageNumber="31">Y. Cambefort</determinerName>
det. 1987, Nepal: Sauraha. Alt. 150, 19.4.82 RN”;
<collectionCode id="BBC9AE01FFE889189FBCFDC3FEEEFD06" box="[189,279,626,652]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="30" pageNumber="31">MNHN</collectionCode>
<specimenCode id="8D7E9EBFFFE889189E1FFDC3FE87FD06" box="[286,382,626,652]" collectionCode="EC" pageId="30" pageNumber="31">EC4808</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE889189FBCFD05FBB5FA3D" pageId="30" pageNumber="31" type="description">
<paragraph id="DD6736C4FFE889189FBCFD05FEB7FD44" blockId="30.[189,334,692,718]" box="[189,334,692,718]" pageId="30" pageNumber="31">
<heading id="862F81A8FFE889189FBCFD05FEB7FD44" bold="true" box="[189,334,692,718]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="EFACEAD6FFE889189FBCFD05FEB7FD44" bold="true" box="[189,334,692,718]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFD54FCEAFD75" blockId="30.[189,787,740,767]" box="[189,787,740,767]" pageId="30" pageNumber="31">MEASUREMENTS (in mm). TL=2025. BW= 1012.</paragraph>
<paragraph id="DD6736C4FFE889189FEBFC99FED3FCC8" blockId="30.[189,1399,808,1187]" box="[234,298,808,834]" pageId="30" pageNumber="31">
<emphasis id="EFACEAD6FFE889189FEBFC99FED3FCC8" bold="true" box="[234,298,808,834]" pageId="30" pageNumber="31">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFCFCFAEEFB29" blockId="30.[189,1399,808,1187]" pageId="30" pageNumber="31">Bronzy green, shining; head rounded, slightly flattened anteriorly; clypeus with fine, sparse punctuation, slightly rugose behind; clypeal carina present, short; frontal carina narrowly interrupted in the middle, with a fairly strong sharp tubercle behind; vertex with smooth punctures; pronotum with simple, strong and dense punctures on the disc, finer, more sparse anteriorly, almost imperceptible at the sides; base not bordered, basal foveae round and placed apart; scutellum very small; elytra smooth, striae broad, not very deep, intervals minutely and sparsely punctured; pygidium convex, very minutely punctured; metasternal shield with shallow longitudinal groove, sparse fine long hairs, the sides granulate anteriorly. Clypeus reflexed, straight in the middle. Profemur with a strong tooth on the antero-median ridge, protibia short, tridentate on the infero-internal ridge, median femur unarmed, tibia toothed on the dorsal side, posterior femur with a strong tooth on the ridge below, directed towards the back, tibia bilobed at the apex.</paragraph>
<paragraph id="DD6736C4FFE889189FBCFB7DFBB4FA83" blockId="30.[189,1398,1228,1289]" pageId="30" pageNumber="31">
AEDEAGUS (
<figureCitation id="45E32A41FFE889189E4BFB7DFE3EFB6C" box="[330,455,1228,1254]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="30" pageNumber="31">Fig. 4EF</figureCitation>
). Parameres shorter than the phallobase, phallobase cylindrical, slightly curved; parameres straight above and curved below, largely notched on the underside.
</paragraph>
<paragraph id="DD6736C4FFE889189FEBFA83FEBCFAC6" blockId="30.[189,1398,1330,1463]" box="[234,325,1330,1356]" pageId="30" pageNumber="31">
<emphasis id="EFACEAD6FFE889189FEBFA83FEBCFAC6" bold="true" box="[234,325,1330,1356]" pageId="30" pageNumber="31">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCFAE7FBB5FA3D" blockId="30.[189,1398,1330,1463]" pageId="30" pageNumber="31">Similar to male but more elongated head, clypeus parabolic, not much reflexed, rugose; with confluent transverse rugulosity behind the clypeal carina; vertex as in the male, with simple, very sparse punctuation; metasternum with well-marked narrow groove; legs not toothed.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE889189FBCFA6EFB5AF9A0" box="[189,1187,1503,1578]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="DD6736C4FFE889189FBCFA6EFEAFFA73" blockId="30.[189,342,1503,1529]" box="[189,342,1503,1529]" pageId="30" pageNumber="31">
<heading id="862F81A8FFE889189FBCFA6EFEAFFA73" bold="true" box="[189,342,1503,1529]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="EFACEAD6FFE889189FBCFA6EFEAFFA73" bold="true" box="[189,342,1503,1529]" pageId="30" pageNumber="31">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE889189FBCF9A1FB5AF9A0" blockId="30.[189,1187,1552,1578]" box="[189,1187,1552,1578]" pageId="30" pageNumber="31">
<collectingCountry id="A5CF7654FFE889189FBCF9A1FEFAF9A0" box="[189,259,1552,1578]" name="Nepal" pageId="30" pageNumber="31">Nepal</collectingCountry>
(
<bibRefCitation id="B9494B35FFE889189E13F9A1FE2EF9A0" author="Cambefort Y." box="[274,471,1552,1578]" pageId="30" pageNumber="31" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort 1988</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE889189EE2F9A1FD1BF9A0" author="Bezdek A. &amp; Krell F. - T." box="[483,738,1552,1578]" pageId="30" pageNumber="31" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE889189DEFF9A1FC75F9A0" author="Bezdek A." box="[750,908,1552,1578]" pageId="30" pageNumber="31" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE889189C99F9A1FB6EF9A0" author="Schoolmeesters P." box="[920,1175,1552,1578]" pageId="30" pageNumber="31" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,409 @@
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<paragraph id="DD6736C4FFED891D9D6FFB94FC3FFBCA" blockId="27.[622,966,1061,1122]" box="[622,966,1061,1088]" pageId="27" pageNumber="28">
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<taxonomicName id="1AD84D47FFED891D9D6FFB94FC3FFBCA" ID-CoL="74N7T" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[622,966,1061,1088]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lama">
<emphasis id="EFACEAD6FFED891D9D6FFB94FD02FBCA" bold="true" box="[622,763,1061,1088]" italics="true" pageId="27" pageNumber="28">Onitis lama</emphasis>
<bibRefCitation id="B9494B35FFED891D9C03FB94FC3FFBCA" author="Lansberge G." box="[770,966,1061,1088]" pageId="27" pageNumber="28" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
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<paragraph id="DD6736C4FFED891D9DA5FBF9FC76FBE8" blockId="27.[622,966,1061,1122]" box="[676,911,1096,1122]" pageId="27" pageNumber="28">
<figureCitation id="45E32A41FFED891D9DA5FBF9FCE8FBE8" box="[676,785,1096,1122]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="27" pageNumber="28">Figs 10C</figureCitation>
,
<figureCitation id="45E32A41FFED891D9C1CFBF9FC9EFBE8" box="[797,871,1096,1122]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. EH. O. feae Felsche, 1907, AIMB/Co/Sc1000188. IL. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." pageId="27" pageNumber="28">11IL</figureCitation>
,
<figureCitation id="45E32A41FFED891D9C73FBF9FC76FBE8" box="[882,911,1096,1122]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." pageId="27" pageNumber="28">19</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFED891D9FBCFB3BFC79FB2E" blockId="27.[189,896,1162,1189]" box="[189,896,1162,1189]" pageId="27" pageNumber="28">
<emphasis id="EFACEAD6FFED891D9FBCFB3BFEBEFB2E" box="[189,327,1162,1188]" italics="true" pageId="27" pageNumber="28">Onitislama</emphasis>
<bibRefCitation id="B9494B35FFED891D9E4FFB3BFDC4FB2F" author="Lansberge G." box="[334,573,1162,1189]" pageId="27" pageNumber="28" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875: 53</bibRefCitation>
, 123 (original description).
</paragraph>
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<subSubSection id="95C2654FFFED891D9FBCFB7DFE95FA32" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="DD6736C4FFED891D9FBCFB7DFE95FA32" blockId="27.[189,1399,1227,1464]" pageId="27" pageNumber="28">
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<emphasis id="EFACEAD6FFED891D9FBCFB7DFEBEFB6F" box="[189,327,1227,1254]" italics="true" pageId="27" pageNumber="28">Onitis lama</emphasis>
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<treatmentCitation id="5C7910D5FFED891D9E64FB7AFD1CFB6F" author="Preudhomme de Borre C. F. P. A." box="[357,741,1227,1254]" page="40" pageId="27" pageNumber="28" year="1881">
<bibRefCitation id="B9494B35FFED891D9E64FB7AFD1CFB6F" author="Preudhomme de Borre C. F. P. A." box="[357,741,1227,1254]" pageId="27" pageNumber="28" refId="ref30731" refString="Preudhomme de Borre C. F. P. A. 1881. Bulletin ou Comptes-Rendus des Seances de la Societe entomologique de Belgique XXXIX." type="book" year="1881">Preudhomme De Borre 1881: 40</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="5C7910D5FFED891D9CA2FB7AFB97FB6F" author="Arrow G. J." box="[931,1134,1227,1253]" page="389" pageId="27" pageNumber="28" year="1931">
<bibRefCitation id="B9494B35FFED891D9CA2FB7AFB97FB6F" author="Arrow G. J." box="[931,1134,1227,1253]" pageId="27" pageNumber="28" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 389</bibRefCitation>
</treatmentCitation>
(key and description). —
<treatmentCitation id="5C7910D5FFED891D9E13FB5FFE0CFA83" author="Balthasar V." box="[274,501,1262,1289]" page="88" pageId="27" pageNumber="28" year="1935">
<bibRefCitation id="B9494B35FFED891D9E13FB5FFE0CFA83" author="Balthasar V." box="[274,501,1262,1289]" pageId="27" pageNumber="28" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 88</bibRefCitation>
</treatmentCitation>
(monograph); 1963: 44 (monograph). —
<treatmentCitation id="5C7910D5FFED891D9CF3FB5EFB32FA83" author="Janssens A." box="[1010,1227,1263,1289]" page="72" pageId="27" pageNumber="28" year="1937">
<bibRefCitation id="B9494B35FFED891D9CF3FB5EFB32FA83" author="Janssens A." box="[1010,1227,1263,1289]" pageId="27" pageNumber="28" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 72</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFED891D9FEBFAA0FE5FFAA1" author="Mittal I. C." box="[234,422,1297,1324]" page="46" pageId="27" pageNumber="28" year="2005">
<bibRefCitation id="B9494B35FFED891D9FEBFAA0FE5FFAA1" author="Mittal I. C." box="[234,422,1297,1324]" pageId="27" pageNumber="28" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005: 46</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFED891D9D84FAA0FC31FAA6" author="Bezdek A. &amp; Krell F. - T." box="[645,968,1297,1324]" page="159" pageId="27" pageNumber="28" year="2006">
<bibRefCitation id="B9494B35FFED891D9D84FAA0FC31FAA6" author="Bezdek A. &amp; Krell F. - T." box="[645,968,1297,1324]" pageId="27" pageNumber="28" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFED891D9BA7FAA0FA8FFAA1" author="Sewak R." box="[1190,1398,1297,1324]" page="36" pageId="27" pageNumber="28" year="2009">
<bibRefCitation id="B9494B35FFED891D9BA7FAA0FA8FFAA1" author="Sewak R." box="[1190,1398,1297,1324]" pageId="27" pageNumber="28" pagination="1 - 48" refId="ref31194" refString="Sewak R. 2009 a. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar desert of Gujarat. Records of the Zoological Survey of India, Occassional Paper No 295: 1 - 48." type="journal article" year="2009">Sewak 2009a: 36</bibRefCitation>
</treatmentCitation>
(diagnosis, distribution); 2009b: 58 (diagnosis, distribution). —
<treatmentCitation id="5C7910D5FFED891D9CC8FA84FB36FAC5" author="Thakare V. &amp; Zade V. &amp; Chandra K. &amp; Gupta D." box="[969,1231,1333,1359]" page="78" pageId="27" pageNumber="28" year="2012">
<bibRefCitation id="B9494B35FFED891D9CC8FA84FB36FAC5" author="Thakare V. &amp; Zade V. &amp; Chandra K. &amp; Gupta D." box="[969,1231,1333,1359]" pageId="27" pageNumber="28" pagination="73 - 80" refId="ref31359" refString="Thakare V., Zade V., Chandra K. &amp; Gupta D. 2012. Scarab beetles (Coleoptera: Scarabaeoidea) of Melghat Tiger Reserve, Central India. Academic Journal of Entomology 5 (2): 73 - 80." type="journal article" year="2012">
Thakare
<emphasis id="EFACEAD6FFED891D9B2FFA87FB9FFAC5" box="[1070,1126,1333,1359]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2012: 78
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFED891D9E15FAE9FDB2FAF8" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[276,587,1368,1394]" page="300" pageId="27" pageNumber="28" year="2014">
<bibRefCitation id="B9494B35FFED891D9E15FAE9FDB2FAF8" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[276,587,1368,1394]" pageId="27" pageNumber="28" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFED891D9E85FAE8FE3DFAF8" box="[388,452,1368,1394]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2014: 300
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFED891D9C2EFAE9FBA0FAF8" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[815,1113,1368,1395]" page="1037" pageId="27" pageNumber="28" year="2015">
<bibRefCitation id="B9494B35FFED891D9C2EFAE9FBA0FAF8" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[815,1113,1368,1395]" pageId="27" pageNumber="28" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFED891D9C82FAE8FC3AFAF8" box="[899,963,1368,1394]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(checklist). —
<treatmentCitation id="5C7910D5FFED891D9A1FFAE9FE98FA1F" author="Bezdek A." page="179" pageId="27" pageNumber="28" year="2016">
<bibRefCitation id="B9494B35FFED891D9A1FFAE9FE98FA1F" author="Bezdek A." pageId="27" pageNumber="28" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFED891D9D38FACAFCB1FA1F" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." box="[569,840,1403,1429]" page="238" pageId="27" pageNumber="28" year="2020">
<bibRefCitation id="B9494B35FFED891D9D38FACAFCB1FA1F" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." box="[569,840,1403,1429]" pageId="27" pageNumber="28" pagination="221 - 275" refId="ref29141" refString="Ghosh J., Das, P., Ghosh S. K., Bhunia D., Kushwaha R. K., Gupta D., &amp; Chandra K. 2020. Insecta: Coleoptera. Zoological Survey of India, Fauna of Haryana, State Fauna Series 24: 221 - 275." type="book chapter" year="2020">
Ghosh
<emphasis id="EFACEAD6FFED891D9D8FFACDFD30FA1F" box="[654,713,1403,1429]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2020: 238
</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFED891D9B21FACAFAE2FA1F" author="Schoolmeesters P." box="[1056,1307,1403,1429]" pageId="27" pageNumber="28" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFED891D9FBCFA51FE46F9C1" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="DD6736C4FFED891D9FBCFA51FE3FFA70" blockId="27.[189,454,1504,1530]" box="[189,454,1504,1530]" pageId="27" pageNumber="28">
<heading id="862F81A8FFED891D9FBCFA51FE3FFA70" bold="true" box="[189,454,1504,1530]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="EFACEAD6FFED891D9FBCFA51FE3FFA70" bold="true" box="[189,454,1504,1530]" pageId="27" pageNumber="28">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFED891D9FBCF9BFFE46F9C1" blockId="27.[189,1398,1550,1612]" pageId="27" pageNumber="28">
<emphasis id="EFACEAD6FFED891D9FBCF9BFFEBEF9A2" box="[189,327,1550,1576]" italics="true" pageId="27" pageNumber="28">Onitislama</emphasis>
is the sole member of Group V (
<bibRefCitation id="B9494B35FFED891D9DCAF9BFFC80F9A2" author="Janssens A." box="[715,889,1550,1576]" pageId="27" pageNumber="28" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937</bibRefCitation>
). It has an uninterrupted frontal carina, and a short clypeal carina.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFED891B9FBCF9C2FEB3FEE0" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="DD6736C4FFED891D9FBCF9C2FE15F907" blockId="27.[189,492,1651,1677]" box="[189,492,1651,1677]" pageId="27" pageNumber="28">
<heading id="862F81A8FFED891D9FBCF9C2FE15F907" bold="true" box="[189,492,1651,1677]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="EFACEAD6FFED891D9FBCF9C2FE15F907" bold="true" box="[189,492,1651,1677]" pageId="27" pageNumber="28">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFED891D9FEBF910FDFFF931" blockId="27.[189,1399,1697,1865]" box="[234,518,1697,1724]" pageId="27" pageNumber="28">
<emphasis id="EFACEAD6FFED891D9FEBF910FEA3F931" bold="true" box="[234,346,1697,1723]" pageId="27" pageNumber="28">
<typeStatus id="02638866FFED891D9FEBF910FEA3F931" box="[234,346,1697,1723]" pageId="27" pageNumber="28" type="syntype">Syntypes</typeStatus>
</emphasis>
(
<specimenCount id="CBDEFD4DFFED891D9E6AF913FE07F931" box="[363,510,1697,1724]" count="2" pageId="27" pageNumber="28" type="generic">2 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFED891D9FBCF974FCA0F8C2" blockId="27.[189,1399,1697,1865]" pageId="27" pageNumber="28">
<materialsCitation id="6DB03C99FFED891D9FBCF974FCACF8C2" collectionCode="MNHN" collectorName="Coll. R. Oberthur &amp; A. Janssens &amp; E. Harold" country="India" location="INDIA" pageId="27" pageNumber="28" specimenCode="EC4810, EC4811" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<collectingCountry id="A5CF7654FFED891D9FBCF974FEF7F955" box="[189,270,1733,1759]" name="India" pageId="27" pageNumber="28">INDIA</collectingCountry>
• 2 “
<specimenCount id="CBDEFD4DFFED891D9E51F974FE79F955" box="[336,384,1733,1759]" count="1" pageId="27" pageNumber="28" type="male">♂♂</specimenCount>
;
<taxonomicName id="1AD84D47FFED891D9E8FF974FDCFF955" authority="Lansb." authorityName="Lansberge" authorityYear="1875" box="[398,566,1733,1759]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lama">
<emphasis id="EFACEAD6FFED891D9E9AF977FE27F955" box="[411,478,1734,1759]" italics="true" pageId="27" pageNumber="28">Lama</emphasis>
Lansb.
</taxonomicName>
India bor. Typus, Ex-Musaeo Van Lansberge, Museum Paris ex
<collectorName id="702D5312FFED891D9A3DF974FEB3F888" pageId="27" pageNumber="28">Coll. R. Oberthur</collectorName>
,
<collectorName id="702D5312FFED891D9E53F959FE20F889" box="[338,473,1768,1795]" pageId="27" pageNumber="28">A. Janssens</collectorName>
vid., 1936:
<emphasis id="EFACEAD6FFED891D9D60F959FD10F888" box="[609,745,1768,1794]" italics="true" pageId="27" pageNumber="28">Onitislama</emphasis>
Lansb. 1875, Type,
<typeStatus id="02638866FFED891D9CD3F959FBCFF888" box="[978,1078,1768,1794]" pageId="27" pageNumber="28" type="syntype">Syntype</typeStatus>
; Polsky, Himalaya,
<taxonomicName id="1AD84D47FFED891D9A1BF959FA8EF888" authorityName="Lansberge" authorityYear="1875" box="[1306,1399,1768,1794]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lama">
<emphasis id="EFACEAD6FFED891D9A1BF959FA8EF888" box="[1306,1399,1768,1794]" italics="true" pageId="27" pageNumber="28">O. lama</emphasis>
</taxonomicName>
typ. Lansbrg., Ex musaeo
<collectorName id="702D5312FFED891D9EEEF8BDFD9AF8AF" box="[495,611,1803,1830]" pageId="27" pageNumber="28">E. Harold</collectorName>
, Museum Paris ex
<collectorName id="702D5312FFED891D9C40F8BAFBE9F8AF" box="[833,1040,1803,1830]" pageId="27" pageNumber="28">Coll. R. Oberthur</collectorName>
,
<collectorName id="702D5312FFED891D9B1BF8BAFB5BF8AC" box="[1050,1186,1803,1830]" pageId="27" pageNumber="28">A. Janssens</collectorName>
vid., 1936:
<taxonomicName id="1AD84D47FFED891D9A2EF8BDFE61F8C3" authority="Lansb., 1875" authorityName="Lansberge" authorityYear="1875" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lama">
<emphasis id="EFACEAD6FFED891D9A2EF8BDFF01F8C2" italics="true" pageId="27" pageNumber="28">Onitis lama</emphasis>
Lansb., 1875
</taxonomicName>
,
<typeStatus id="02638866FFED891D9EA7F89FFDF5F8C2" box="[422,524,1838,1864]" pageId="27" pageNumber="28" type="syntype">Syntype</typeStatus>
”;
<collectionCode id="BBC9AE01FFED891D9D22F89EFD84F8C3" box="[547,637,1839,1865]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="28">MNHN</collectionCode>
<specimenCode id="8D7E9EBFFFED891D9D85F89FFD1FF8C2" box="[644,742,1838,1864]" collectionCode="EC" pageId="27" pageNumber="28">EC4810</specimenCode>
<specimenCode id="8D7E9EBFFFED891D9DF4F89FFCACF8C2" box="[757,853,1838,1864]" collectionCode="EC" pageId="27" pageNumber="28">EC4811</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="DD6736C4FFED891D9FEBF8C1FCF5F800" blockId="27.[189,1399,1904,2001]" box="[234,780,1904,1930]" pageId="27" pageNumber="28">
<emphasis id="EFACEAD6FFED891D9FEBF8C1FD98F800" bold="true" box="[234,609,1904,1930]" pageId="27" pageNumber="28">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFED891D9D73F8C1FCFDF800" box="[626,772,1904,1930]" count="9" pageId="27" pageNumber="28" type="generic">9 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFED891B9FBCF822FEB3FEE0" blockId="27.[189,1399,1904,2001]" lastBlockId="29.[189,1399,265,362]" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
<materialsCitation id="6DB03C99FFED891D9FBCF822FBF4F827" box="[189,1037,1939,1966]" collectionCode="BMNH" country="India" location="INDIA" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFED891D9FBCF822FEF7F827" box="[189,270,1939,1965]" name="India" pageId="27" pageNumber="28">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFED891D9E31F825FEB0F824" box="[304,329,1940,1966]" count="1" pageId="27" pageNumber="28" type="male"></specimenCount>
; “Punjab, Nevinson Coll. 1918-14”;
<collectionCode id="BBC9AE01FFED891D9C10F825FC90F824" box="[785,873,1940,1966]" pageId="27" pageNumber="28">BMNH</collectionCode>
(E) 1237147
</materialsCitation>
<materialsCitation id="6DB03C99FFED891B9B2EF825FDF8FEA9" collectionCode="BMNH" collectorName="W. Almora" country="India" lastPageId="29" lastPageNumber="30" location="Uttarakhand" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1" stateProvince="Uttarakhand">
<specimenCount id="CBDEFD4DFFED891D9B2EF825FBBFF824" box="[1071,1094,1940,1966]" count="1" pageId="27" pageNumber="28" type="female"></specimenCount>
; “
<collectingRegion id="1F1CF826FFED891D9B62F822FB03F827" box="[1123,1274,1939,1965]" country="India" name="Uttarakhand" pageId="27" pageNumber="28">Uttarakhand</collectingRegion>
, Kumaon,
<collectorName id="702D5312FFED891D9FBCF806FEBDF85A" box="[189,324,1974,2001]" pageId="27" pageNumber="28">W. Almora</collectorName>
, India. H.G.C., H.G. Champion Coll., B.M. 1953-156.,
<taxonomicName id="1AD84D47FFED891D9CF1F806FA8EF85A" authority=", Lansb., G. J. Arrow" authorityName="Lansb., G. J. Arrow" box="[1008,1399,1974,2001]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lama">
<emphasis id="EFACEAD6FFED891D9CF1F806FB75F85A" box="[1008,1164,1974,2001]" italics="true" pageId="27" pageNumber="28">Onitis lama,</emphasis>
Lansb., G.J. Arrow
</taxonomicName>
det.”;
<collectionCode id="BBC9AE01FFEB891B9E09FEBBFE99FEAE" box="[264,352,266,292]" pageId="29" pageNumber="30">BMNH</collectionCode>
(E) 1237148
</materialsCitation>
<materialsCitation id="6DB03C99FFEB891B9D21FEBBFEBCFEE0" collectingDate="2016-12-20" collectionCode="ZSIM, AIMB" collectorName="Det. J. J. E. Gillet &amp; Chian" country="India" determinerName="Det. J. J. E. Gillet" location="Tamil Nadu" pageId="29" pageNumber="30" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Tamil Nadu">
<specimenCount id="CBDEFD4DFFEB891B9D21FEBBFDC0FEAE" box="[544,569,266,292]" count="1" pageId="29" pageNumber="30" type="male"></specimenCount>
; “
<emphasis id="EFACEAD6FFEB891B9D54FEB8FD1BFEA9" box="[597,738,265,291]" italics="true" pageId="29" pageNumber="30">Onitislama</emphasis>
,
<determinerName id="28BBDA00FFEB891B9DF5FEBBFC39FEA9" box="[756,960,265,292]" pageId="29" pageNumber="30">
<collectorName id="702D5312FFEB891B9DF5FEBBFC39FEA9" box="[756,960,265,292]" pageId="29" pageNumber="30">Det. J.J.E. Gillet</collectorName>
</determinerName>
;
<collectionCode id="BBC9AE01FFEB891B9CCFFEB8FBEDFEA9" box="[974,1044,265,291]" pageId="29" pageNumber="30">ZSIM</collectionCode>
<specimenCount id="CBDEFD4DFFEB891B9B32FEB8FB83FEAE" box="[1075,1146,265,292]" count="2" pageId="29" pageNumber="30" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFEB891B9B8BFEB8FB29FEAE" box="[1162,1232,265,292]" count="2" pageId="29" pageNumber="30" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFEB891B9BE0FEB8FA8AFEA9" box="[1249,1395,265,291]" country="India" name="Tamil Nadu" pageId="29" pageNumber="30">Tamil Nadu</collectingRegion>
, Tirunelveli, Radhapuram”;
<collectingMethod id="04994ED3FFEB891B9D02FE9CFCD3FECD" box="[515,810,300,327]" pageId="29" pageNumber="30">open cow dung bait trap</collectingMethod>
,
<date id="A9661004FFEB891B9C36FE9DFC25FECC" box="[823,988,300,327]" pageId="29" pageNumber="30" value="2016-12-20">
<collectingDate id="B922E9ECFFEB891B9C36FE9DFC25FECC" box="[823,988,300,327]" pageId="29" pageNumber="30" value="2016-12-20">20 Dec. 2016</collectingDate>
</date>
,
<collectorName id="702D5312FFEB891B9CE8FE9DFBD6FECC" box="[1001,1071,300,326]" pageId="29" pageNumber="30">Chian</collectorName>
leg.;
<collectionCode id="BBC9AE01FFEB891B9B72FE9DFB39FECC" box="[1139,1216,300,326]" pageId="29" pageNumber="30">AIMB</collectionCode>
/Co/Sc1000306 to 1000309
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="89A7664CFFEA891A9FBCF8D5FDFBF862" pageId="28" pageNumber="29" startId="28.[189,232,1892,1918]" targetBox="[295,1242,313,1806]" targetPageId="28">
<paragraph id="DD6736C4FFEA891A9FBCF8D5FDFBF862" blockId="28.[189,1399,1892,2024]" pageId="28" pageNumber="29">
<emphasis id="EFACEAD6FFEA891A9FBCF8D5FEF5F8F5" bold="true" box="[189,268,1892,1919]" pageId="28" pageNumber="29">Fig. 8.</emphasis>
Aedeagus, lateral, dorsal, ventral and apical views.
<emphasis id="EFACEAD6FFEA891A9C7AF8D5FC4DF8F5" bold="true" box="[891,948,1892,1919]" pageId="28" pageNumber="29">AD</emphasis>
.
<emphasis id="EFACEAD6FFEA891A9CC4F8D4FBA4F8F5" box="[965,1117,1893,1919]" italics="true" pageId="28" pageNumber="29">Onitiskethai</emphasis>
<taxonomicNameLabel id="F49F57ADFFEA891A9B67F8D4FB3CF8F5" box="[1126,1221,1893,1919]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
, NIM/NBAIR/ COL/ONIT/H-271023B.
<emphasis id="EFACEAD6FFEA891A9EE4F839FDE7F828" bold="true" box="[485,542,1928,1954]" pageId="28" pageNumber="29">EH</emphasis>
.
<taxonomicName id="1AD84D47FFEA891A9D2BF839FC8AF828" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" box="[554,883,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFEA891A9D2BF839FD41F828" box="[554,696,1928,1954]" italics="true" pageId="28" pageNumber="29">O. philemon</emphasis>
<bibRefCitation id="B9494B35FFEA891A9DBCF839FC8AF828" author="Fabricius J. C." box="[701,883,1928,1954]" pageId="28" pageNumber="29" refId="ref29058" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Volume 2. Impensis Bibliopolii novi academici, Kiel." type="book" year="1801">Fabricius,1801</bibRefCitation>
</taxonomicName>
,AIMB/Co/Sc1000317.
<emphasis id="EFACEAD6FFEA891A9B8BF839FB41F828" bold="true" box="[1162,1208,1928,1954]" pageId="28" pageNumber="29">IL</emphasis>
.
<taxonomicName id="1AD84D47FFEA891A9BC5F839FE70F84C" authority="Lansberge, 1875" authorityName="Lansberge, AIMB" authorityYear="1875" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFEA891A9BC5F839FA8FF828" box="[1220,1398,1928,1954]" italics="true" pageId="28" pageNumber="29">O. singhalensis</emphasis>
<bibRefCitation id="B9494B35FFEA891A9FBCF81AFE70F84C" author="Lansberge G." box="[189,393,1963,1990]" pageId="28" pageNumber="29" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000684.
<emphasis id="EFACEAD6FFEA891A9DB9F81AFD0CF84F" bold="true" box="[696,757,1963,1989]" pageId="28" pageNumber="29">MP</emphasis>
.
<taxonomicName id="1AD84D47FFEA891A9C0DF81DFC61F84F" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[780,920,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov.">
<emphasis id="EFACEAD6FFEA891A9C0DF81DFC61F84F" box="[780,920,1963,1990]" italics="true" pageId="28" pageNumber="29">O. visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFEA891A9CA9F81DFBF2F84C" box="[936,1035,1964,1990]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
, NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB).
</paragraph>
</caption>
<subSubSection id="95C2654FFFEB891B9FBCFE20FE10FAB1" pageId="29" pageNumber="30" type="description">
<paragraph id="DD6736C4FFEB891B9FBCFE20FEB7FE21" blockId="29.[189,334,401,427]" box="[189,334,401,427]" pageId="29" pageNumber="30">
<heading id="862F81A8FFEB891B9FBCFE20FEB7FE21" bold="true" box="[189,334,401,427]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="EFACEAD6FFEB891B9FBCFE20FEB7FE21" bold="true" box="[189,334,401,427]" pageId="29" pageNumber="30">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEB891B9FBCFE0EFCEBFE53" blockId="29.[189,786,447,474]" box="[189,786,447,474]" pageId="29" pageNumber="30">MEASUREMENTS (in mm). TL=1924, BW =1113.</paragraph>
<paragraph id="DD6736C4FFEB891B9FEBFDB0FED3FD91" blockId="29.[189,1399,513,1032]" box="[234,298,513,539]" pageId="29" pageNumber="30">
<emphasis id="EFACEAD6FFEB891B9FEBFDB0FED3FD91" bold="true" box="[234,298,513,539]" pageId="29" pageNumber="30">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFEB891B9FBCFD95FD25FB82" blockId="29.[189,1399,513,1032]" pageId="29" pageNumber="30">Black, shining; head with an entire, strongly elevated, slightly arcuate frontal carina, clypeal carina short, curved, closer to clypeal margin; pronotum distinctly, moderately closely punctured anteriorly and extremely minutely behind; front angles blunt, sides strongly rounded, base completely margined and lobed between basal foveae; elytra with fine striae, smooth intervals, sutural intervals with few scattered punctures, posterior edge with short fringe of erect yellow hairs; pygidium smooth, opaque with very minute scanty punctures; metasternum with a narrow longitudinal groove, metasternal shield smooth in the middle and behind, flat and fairly strongly punctured anteriorly and sides, the sides of the metasternum densely punctured and hairy. Clypeus deeply excised in the middle, not very closely rugulose; posterior part of head very finely and sparsely punctured; profemur with a very sharp oblique tooth beyond the middle of its lower edge, protibia elongate, armed with four short external teeth, a strong sharp tooth on the lower face before the middle and three or four minute tubercles and an incurved apical process, hind femur rather long, straight with a tooth near the end of the lower edge, trochanter little produced at the tip and hind tibia finely serrate along its inner edge.</paragraph>
<paragraph id="DD6736C4FFEB891B9FBCFB9EFE70FB05" blockId="29.[189,1398,1071,1168]" pageId="29" pageNumber="30">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFEB891B9EB2FB9EFDC7FBC3" box="[435,574,1071,1097]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. EH. O. feae Felsche, 1907, AIMB/Co/Sc1000188. IL. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." pageId="29" pageNumber="30">Fig. 11I L</figureCitation>
). LP=3.035, Lp= 2.57; BP=1.43, BpB =0.93, BpT=0.36. Phallobase longer than parameres which are blunt, slightly sinuate near the base, curved inwards anteriorly and hooked at the tip.
</paragraph>
<paragraph id="DD6736C4FFEB891B9FEBFB06FEBCFB5B" blockId="29.[189,1399,1207,1339]" box="[234,325,1207,1233]" pageId="29" pageNumber="30">
<emphasis id="EFACEAD6FFEB891B9FEBFB06FEBCFB5B" bold="true" box="[234,325,1207,1233]" pageId="29" pageNumber="30">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFEB891B9FBCFB6BFE10FAB1" blockId="29.[189,1399,1207,1339]" pageId="29" pageNumber="30">Clypeus closely rugulose, its front margin entire and slightly truncate, pronotum more strongly punctured anteriorly than male, protibia broad, armed with four blunt external teeth and a small articulated terminal spur.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEB891B9FBCFAD3FDACF99E" pageId="29" pageNumber="30" type="distribution">
<paragraph id="DD6736C4FFEB891B9FBCFAD3FEAFFAF6" blockId="29.[189,342,1378,1404]" box="[189,342,1378,1404]" pageId="29" pageNumber="30">
<heading id="862F81A8FFEB891B9FBCFAD3FEAFFAF6" bold="true" box="[189,342,1378,1404]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="EFACEAD6FFEB891B9FBCFAD3FEAFFAF6" bold="true" box="[189,342,1378,1404]" pageId="29" pageNumber="30">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEB891B9FBCFA21FDACF99E" blockId="29.[189,1399,1424,1556]" pageId="29" pageNumber="30">
<collectingCountry id="A5CF7654FFEB891B9FBCFA21FF07FA20" box="[189,254,1424,1450]" name="India" pageId="29" pageNumber="30">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFEB891B9E0BFA21FE34FA20" box="[266,461,1424,1450]" country="India" name="Andhra Pradesh" pageId="29" pageNumber="30">Andhra Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFEB891B9EDDFA21FDCEFA20" box="[476,567,1424,1450]" country="India" name="Gujarat" pageId="29" pageNumber="30">Gujarat</collectingRegion>
,
<collectingRegion id="1F1CF826FFEB891B9D44FA20FD52FA21" box="[581,683,1425,1451]" country="India" name="Haryana" pageId="29" pageNumber="30">Haryana</collectingRegion>
,
<collectingRegion id="1F1CF826FFEB891B9DBAFA21FC6EFA20" box="[699,919,1424,1450]" country="India" name="Himachal Pradesh" pageId="29" pageNumber="30">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFEB891B9CA7FA21FBC5FA20" box="[934,1084,1424,1450]" country="India" name="Maharashtra" pageId="29" pageNumber="30">Maharashtra</collectingRegion>
, Punjab,
<collectingRegion id="1F1CF826FFEB891B9BAFFA21FADCFA20" box="[1198,1317,1424,1450]" country="India" name="Rajasthan" pageId="29" pageNumber="30">Rajasthan</collectingRegion>
,
<collectingRegion id="1F1CF826FFEB891B9A32FA21FEF9FA44" country="India" name="Tamil Nadu" pageId="29" pageNumber="30">Tamil Nadu</collectingRegion>
,
<collectingRegion id="1F1CF826FFEB891B9E0BFA05FE67FA44" box="[266,414,1460,1486]" country="India" name="Uttarakhand" pageId="29" pageNumber="30">Uttarakhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFEB891B9EA8FA05FDB5FA44" box="[425,588,1460,1486]" country="India" name="Uttar Pradesh" pageId="29" pageNumber="30">Uttar Pradesh</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFEB891B9C19FA05FC98FA44" box="[792,865,1460,1486]" name="Nepal" pageId="29" pageNumber="30">Nepal</collectingCountry>
,
<collectingCountry id="A5CF7654FFEB891B9C6DFA05FC36FA44" box="[876,975,1460,1486]" name="Pakistan" pageId="29" pageNumber="30">Pakistan</collectingCountry>
(
<bibRefCitation id="B9494B35FFEB891B9CDCFA05FB92FA44" author="Mittal I. C." box="[989,1131,1459,1486]" pageId="29" pageNumber="30" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005</bibRefCitation>
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<bibRefCitation id="B9494B35FFEB891B9B77FA05FA8BFA44" author="Bezdek A. &amp; Krell F. - T." box="[1142,1394,1460,1486]" pageId="29" pageNumber="30" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
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<bibRefCitation id="B9494B35FFEB891B9FBCFA66FEA4FA7B" author="Sewak R." box="[189,349,1495,1521]" pageId="29" pageNumber="30" pagination="1 - 48" refId="ref31194" refString="Sewak R. 2009 a. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar desert of Gujarat. Records of the Zoological Survey of India, Occassional Paper No 295: 1 - 48." type="journal article" year="2009">Sewak 2009a</bibRefCitation>
,
<bibRefCitation id="B9494B35FFEB891B9E69FA66FE4CFA7B" author="Sewak R." box="[360,437,1495,1521]" pageId="29" pageNumber="30" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">2009b</bibRefCitation>
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).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,328 @@
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<mods:title id="48B93249546CB17D7FED2BD77C595C74">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<emphasis id="EFACEAD6FFEF891F9D9AFDB1FCCEFD91" bold="true" box="[667,823,512,539]" italics="true" pageId="25" pageNumber="26">Onitis kethai</emphasis>
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</subSubSection>
<subSubSection id="95C2654FFFEF891F9D3DFDF6FC0EFDEB" box="[572,1015,583,610]" pageId="25" pageNumber="26" type="description">
<paragraph id="DD6736C4FFEF891F9D3DFDF6FC0EFDEB" blockId="25.[373,1214,512,610]" box="[572,1015,583,610]" pageId="25" pageNumber="26">
<figureCitation id="45E32A41FFEF891F9D3DFDF6FD7EFDEB" box="[572,647,583,609]" captionStart="Fig" captionStartId="26.[189,232,1923,1949]" captionTargetBox="[299,1295,368,1851]" captionTargetId="figure-42@26.[191,1400,269,1864]" captionTargetPageId="26" captionText="Fig. 7. Onitis kethai sp. nov. AE. Holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023B. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head, dorsal view. D. Pygidium. E. Elytra. F. Paratype, ♀, habitus, dorsal view. (Images: AIMB)." pageId="25" pageNumber="26">Figs 7</figureCitation>
,
<figureCitation id="45E32A41FFEF891F9D95FDF6FD27FDEB" box="[660,734,583,609]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="25" pageNumber="26">8AD</figureCitation>
,
<figureCitation id="45E32A41FFEF891F9DEBFDF6FCF7FDEB" box="[746,782,583,609]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="25" pageNumber="26">9C</figureCitation>
,
<figureCitation id="45E32A41FFEF891F9C1AFDF6FCB7FDEB" box="[795,846,583,609]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="25" pageNumber="26">14C</figureCitation>
,
<figureCitation id="45E32A41FFEF891F9C5BFDF6FC77FDE8" box="[858,910,583,610]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="25" pageNumber="26">15C</figureCitation>
,
<figureCitation id="45E32A41FFEF891F9C9BFDF6FC34FDEB" box="[922,973,583,609]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="25" pageNumber="26">16C</figureCitation>
,
<figureCitation id="45E32A41FFEF891F9CDBFDF6FC0EFDEB" box="[986,1015,583,609]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="25" pageNumber="26">18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEF891F9FBCFD39FDFAFC60" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="DD6736C4FFEF891F9FBCFD39FE3FFD28" blockId="25.[189,454,648,674]" box="[189,454,648,674]" pageId="25" pageNumber="26">
<heading id="862F81A8FFEF891F9FBCFD39FE3FFD28" bold="true" box="[189,454,648,674]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="EFACEAD6FFEF891F9FBCFD39FE3FFD28" bold="true" box="[189,454,648,674]" pageId="25" pageNumber="26">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEF891F9FBCFD06FDFAFC60" blockId="25.[189,1399,695,1003]" pageId="25" pageNumber="26">
<emphasis id="EFACEAD6FFEF891F9FBCFD06FEAAFD5B" box="[189,339,695,721]" italics="true" pageId="25" pageNumber="26">Onitiskethai</emphasis>
<taxonomicNameLabel id="F49F57ADFFEF891F9E5BFD06FE4DFD5B" box="[346,436,695,721]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
can be easily distinguished from
<taxonomicName id="1AD84D47FFEF891F9C3DFD06FC35FD5B" authorityName="Fabricius" authorityYear="1801" box="[828,972,695,721]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFEF891F9C3DFD06FC35FD5B" box="[828,972,695,721]" italics="true" pageId="25" pageNumber="26">O. philemon</emphasis>
</taxonomicName>
in having stronger and more closely placed punctures on the pronotum; genae smoothly rounded in
<taxonomicName id="1AD84D47FFEF891F9CBFFD6BFBD4FD7E" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[958,1069,730,756]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kethai">
<emphasis id="EFACEAD6FFEF891F9CBFFD6BFBD4FD7E" box="[958,1069,730,756]" italics="true" pageId="25" pageNumber="26">O. kethai</emphasis>
</taxonomicName>
, while strongly rounded in
<taxonomicName id="1AD84D47FFEF891F9FBCFD4FFEA8FC9D" authorityName="Fabricius" authorityYear="1801" box="[189,337,765,792]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFEF891F9FBCFD4FFEA8FC9D" box="[189,337,765,792]" italics="true" pageId="25" pageNumber="26">O. philemon</emphasis>
</taxonomicName>
, both inconspicuously punctured. The clypeus is rugulose in
<taxonomicName id="1AD84D47FFEF891F9B3AFD4FFB48FC9D" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1083,1201,765,792]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kethai">
<emphasis id="EFACEAD6FFEF891F9B3AFD4FFB48FC9D" box="[1083,1201,765,792]" italics="true" pageId="25" pageNumber="26">O. kethai</emphasis>
</taxonomicName>
(
<figureCitation id="45E32A41FFEF891F9BBCFD4CFAE3FC9D" box="[1213,1306,765,791]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="25" pageNumber="26">Fig. 9C</figureCitation>
); while in
<taxonomicName id="1AD84D47FFEF891F9FDCFC90FE96FCB0" authorityName="Fabricius" authorityYear="1801" box="[221,367,800,827]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFEF891F9FDCFC90FE96FCB0" box="[221,367,800,827]" italics="true" pageId="25" pageNumber="26">O. philemon</emphasis>
</taxonomicName>
it is granular in male (
<figureCitation id="45E32A41FFEF891F9D85FC91FD1BFCB0" box="[644,738,800,826]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="25" pageNumber="26">Fig. 9D</figureCitation>
) and rugose in female.
<emphasis id="EFACEAD6FFEF891F9CFEFC91FB18FCB0" box="[1023,1249,800,826]" italics="true" pageId="25" pageNumber="26">Onitissinghalensis</emphasis>
differs from both these species in having strong rounded granules on clypeus and head while the genae are sparingly granular (
<figureCitation id="45E32A41FFEF891F9E33FCD6FE77FC0B" box="[306,398,871,897]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="25" pageNumber="26">Fig. 9F</figureCitation>
). In
<taxonomicName id="1AD84D47FFEF891F9EC9FCD6FDCEFC0B" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[456,567,871,897]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kethai">
<emphasis id="EFACEAD6FFEF891F9EC9FCD6FDCEFC0B" box="[456,567,871,897]" italics="true" pageId="25" pageNumber="26">O. kethai</emphasis>
</taxonomicName>
the space between the clypeal carina, frontal carina and the tubercle behind is strongly rugose, while in
<taxonomicName id="1AD84D47FFEF891F9D6DFC3BFD06FC2E" authorityName="Fabricius" authorityYear="1801" box="[620,767,906,932]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFEF891F9D6DFC3BFD06FC2E" box="[620,767,906,932]" italics="true" pageId="25" pageNumber="26">O. philemon</emphasis>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFEF891F9C3FFC3BFC0FFC2E" authorityName="Lansberge, AIMB" authorityYear="1875" box="[830,1014,906,932]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFEF891F9C3FFC3BFC0FFC2E" box="[830,1014,906,932]" italics="true" pageId="25" pageNumber="26">O. singhalensis</emphasis>
</taxonomicName>
it is granular. Differences were also observed on comparing the aedeagus of
<taxonomicName id="1AD84D47FFEF891F9DEBFC1FFCA3FC4D" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[746,858,941,968]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kethai">
<emphasis id="EFACEAD6FFEF891F9DEBFC1FFCA3FC4D" box="[746,858,941,968]" italics="true" pageId="25" pageNumber="26">O. kethai</emphasis>
</taxonomicName>
(
<figureCitation id="45E32A41FFEF891F9C6FFC1CFC0DFC4D" box="[878,1012,941,967]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="25" pageNumber="26">Fig. 8AD</figureCitation>
),
<taxonomicName id="1AD84D47FFEF891F9B0FFC1FFB5AFC4D" authorityName="Fabricius" authorityYear="1801" box="[1038,1187,941,968]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFEF891F9B0FFC1FFB5AFC4D" box="[1038,1187,941,968]" italics="true" pageId="25" pageNumber="26">O. philemon</emphasis>
</taxonomicName>
(
<figureCitation id="45E32A41FFEF891F9BB6FC1CFAC1FC4D" box="[1207,1336,941,967]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="25" pageNumber="26">Fig. 8EH</figureCitation>
) and
<taxonomicName id="1AD84D47FFEF891F9FBCFC60FE8BFC60" authorityName="Lansberge, AIMB" authorityYear="1875" box="[189,370,976,1003]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFEF891F9FBCFC60FE8BFC60" box="[189,370,976,1003]" italics="true" pageId="25" pageNumber="26">O. singhalensis</emphasis>
</taxonomicName>
(
<figureCitation id="45E32A41FFEF891F9E83FC61FE0FFC60" box="[386,502,976,1002]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="25" pageNumber="26">Fig. 8IL</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEF891F9FBCFBA3FC81FBF7" pageId="25" pageNumber="26" type="etymology">
<paragraph id="DD6736C4FFEF891F9FBCFBA3FEBDFBA6" blockId="25.[189,324,1042,1068]" box="[189,324,1042,1068]" pageId="25" pageNumber="26">
<heading id="862F81A8FFEF891F9FBCFBA3FEBDFBA6" bold="true" box="[189,324,1042,1068]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="EFACEAD6FFEF891F9FBCFBA3FEBDFBA6" bold="true" box="[189,324,1042,1068]" pageId="25" pageNumber="26">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEF891F9FBCFBF1FC81FBF7" blockId="25.[189,1398,1088,1149]" pageId="25" pageNumber="26">Named in memory of our field assistant, the late Mr Ketha Gowda who assisted PDR in many of his field trips and helped in sampling dung beetles and other insects.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEF891F9FBCFB14FD74F989" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="DD6736C4FFEF891F9FBCFB14FE15FB35" blockId="25.[189,492,1189,1215]" box="[189,492,1189,1215]" pageId="25" pageNumber="26">
<heading id="862F81A8FFEF891F9FBCFB14FE15FB35" bold="true" box="[189,492,1189,1215]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="EFACEAD6FFEF891F9FBCFB14FE15FB35" bold="true" box="[189,492,1189,1215]" pageId="25" pageNumber="26">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEF891F9FEBFB65FEA5FB64" blockId="25.[189,1399,1236,1368]" box="[234,348,1236,1262]" pageId="25" pageNumber="26">
<emphasis id="EFACEAD6FFEF891F9FEBFB65FEA5FB64" bold="true" box="[234,348,1236,1262]" pageId="25" pageNumber="26">
<typeStatus id="02638866FFEF891F9FEBFB65FEA5FB64" box="[234,348,1236,1262]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFEF891F9FBCFB46FC7AFADD" blockId="25.[189,1399,1236,1368]" pageId="25" pageNumber="26">
<materialsCitation id="6DB03C99FFEF891F9FBCFB46FC7AFADD" collectingDate="1998-05-10" collectionCode="NBAIR, NIM" collectorName="Priyadarsanan Dharma Rajan" country="India" county="Biligiri Rangaswamy" latitude="11.902133" location="Honnametti" longLatPrecision="1" longitude="77.19273" municipality="Tiger Reserve" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka" typeStatus="holotype">
<collectingCountry id="A5CF7654FFEF891F9FBCFB46FEF7FA9B" box="[189,270,1271,1297]" name="India" pageId="25" pageNumber="26">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFEF891F9E28FB46FEBBFA9B" box="[297,322,1271,1297]" count="1" pageId="25" pageNumber="26" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFEF891F9E4EFB46FE32FA9B" box="[335,459,1271,1297]" country="India" name="Karnataka" pageId="25" pageNumber="26">Karnataka</collectingRegion>
,
<collectingCounty id="34064E48FFEF891F9ED6FB46FD2BFA9B" box="[471,722,1271,1297]" pageId="25" pageNumber="26">Biligiri Rangaswamy</collectingCounty>
temple
<collectingMunicipality id="3D03ACBEFFEF891F9C30FB46FC20FA9B" box="[817,985,1271,1297]" pageId="25" pageNumber="26">Tiger Reserve</collectingMunicipality>
,
<location id="D807601FFFEF891F9CE7FB46FB88FA9B" LSID="urn:lsid:plazi:treatment:557187D2FFEF891D9D9AFDB1FE0CFC55:D807601FFFEF891F9CE7FB46FB88FA9B" box="[998,1137,1271,1297]" country="India" county="Biligiri Rangaswamy" latitude="11.902133" longLatPrecision="1" longitude="77.19273" municipality="Tiger Reserve" name="Honnametti" pageId="25" pageNumber="26" stateProvince="Karnataka">Honnametti</location>
shola;
<geoCoordinate id="B8EC5003FFEF891F9BC6FB44FA8BFA9B" box="[1223,1394,1269,1297]" degrees="11" direction="north" minutes="54" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="7.68" value="11.902133">11°54ʹ7.68″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFEF891F9FBCFAA9FE8CFABE" box="[189,373,1304,1332]" degrees="77" direction="east" minutes="11" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="33.84" value="77.19273">77°11ʹ33.84″ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFEF891F9E82FAABFD50FABE" box="[387,681,1306,1332]" pageId="25" pageNumber="26">open cow dung bait trap</collectingMethod>
;
<date id="A9661004FFEF891F9DB6FAABFCA0FABE" box="[695,857,1306,1332]" pageId="25" pageNumber="26" value="1998-05-10">
<collectingDate id="B922E9ECFFEF891F9DB6FAABFCA0FABE" box="[695,857,1306,1332]" pageId="25" pageNumber="26" value="1998-05-10">10 May 1998</collectingDate>
</date>
,
<collectorName id="702D5312FFEF891F9C66FAABFB3BFABE" box="[871,1218,1306,1332]" pageId="25" pageNumber="26">Priyadarsanan Dharma Rajan</collectorName>
leg.; specimen deposited at
<collectionCode id="BBC9AE01FFEF891F9E50FA8CFE56FADD" box="[337,431,1341,1367]" pageId="25" pageNumber="26">NBAIR</collectionCode>
,
<collectionCode id="BBC9AE01FFEF891F9EBAFA8CFE0CFADD" box="[443,501,1341,1367]" country="France" name="Museum d'histoire naturelle de Nîmes" pageId="25" pageNumber="26" type="Herbarium">NIM</collectionCode>
/
<collectionCode id="BBC9AE01FFEF891F9EFFFA8CFDA2FADD" box="[510,603,1341,1367]" pageId="25" pageNumber="26">NBAIR</collectionCode>
/COL/ONIT/H-271023B.
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFEF891F9FEBFACEFDDBFA12" blockId="25.[189,1399,1406,1539]" box="[234,546,1406,1433]" pageId="25" pageNumber="26">
<emphasis id="EFACEAD6FFEF891F9FEBFACEFE9EFA13" bold="true" box="[234,359,1407,1433]" pageId="25" pageNumber="26">
<typeStatus id="02638866FFEF891F9FEBFACEFE9EFA13" box="[234,359,1407,1433]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="CBDEFD4DFFEF891F9E79FACEFDE3FA12" box="[376,538,1406,1433]" count="13" pageId="25" pageNumber="26" type="generic">13 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFEF891F9FBCFA13FD74F989" blockId="25.[189,1399,1406,1539]" pageId="25" pageNumber="26">
<materialsCitation id="6DB03C99FFEF891F9FBCFA13FD70F989" collectingDate="1998-05-10" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Co &amp; Sc" country="India" county="Biligiri Rangaswamy" latitude="11.902133" location="Honnametti" longLatPrecision="1" longitude="77.19273" municipality="Tiger Reserve" pageId="25" pageNumber="26" specimenCount="13" specimenCount-female="5" specimenCount-male="8" stateProvince="Karnataka">
<collectingCountry id="A5CF7654FFEF891F9FBCFA13FEF7FA36" box="[189,270,1442,1468]" name="India" pageId="25" pageNumber="26">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFEF891F9E33FA13FE82FA36" box="[306,379,1442,1468]" count="8" pageId="25" pageNumber="26" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFEF891F9E8CFA13FE2CFA36" box="[397,469,1442,1469]" count="5" pageId="25" pageNumber="26" type="female">5 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFEF891F9EE9FA13FD9DFA36" box="[488,612,1442,1468]" country="India" name="Karnataka" pageId="25" pageNumber="26">Karnataka</collectingRegion>
,
<collectingCounty id="34064E48FFEF891F9D74FA13FC8DFA36" box="[629,884,1442,1468]" pageId="25" pageNumber="26">Biligiri Rangaswamy</collectingCounty>
temple
<collectingMunicipality id="3D03ACBEFFEF891F9CDDFA13FB71FA36" box="[988,1160,1442,1468]" pageId="25" pageNumber="26">Tiger Reserve</collectingMunicipality>
,
<location id="D807601FFFEF891F9B98FA13FADDFA36" LSID="urn:lsid:plazi:treatment:557187D2FFEF891D9D9AFDB1FE0CFC55:D807601FFFEF891F9B98FA13FADDFA36" box="[1177,1316,1442,1468]" country="India" county="Biligiri Rangaswamy" latitude="11.902133" longLatPrecision="1" longitude="77.19273" municipality="Tiger Reserve" name="Honnametti" pageId="25" pageNumber="26" stateProvince="Karnataka">Honnametti</location>
shola;
<geoCoordinate id="B8EC5003FFEF891F9FBCFA72FE90FA55" box="[189,361,1475,1503]" degrees="11" direction="north" minutes="54" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="7.68" value="11.902133">11°54ʹ7.68″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFEF891F9E77FA72FDD4FA55" box="[374,557,1475,1503]" degrees="77" direction="east" minutes="11" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="33.84" value="77.19273">77°11ʹ33.84″ E</geoCoordinate>
;
<collectingMethod id="04994ED3FFEF891F9D3AFA74FCA6FA55" box="[571,863,1477,1503]" pageId="25" pageNumber="26">open cow dung bait trap</collectingMethod>
;
<date id="A9661004FFEF891F9C6DFA74FBF6FA55" box="[876,1039,1477,1503]" pageId="25" pageNumber="26" value="1998-05-10">
<collectingDate id="B922E9ECFFEF891F9C6DFA74FBF6FA55" box="[876,1039,1477,1503]" pageId="25" pageNumber="26" value="1998-05-10">10 May 1998</collectingDate>
</date>
;
<collectorName id="702D5312FFEF891F9B1DFA74FA8FFA55" box="[1052,1398,1477,1503]" pageId="25" pageNumber="26">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFEF891F9FF7FA59FEBAF988" box="[246,323,1512,1538]" pageId="25" pageNumber="26">AIMB</collectionCode>
/
<collectorName id="702D5312FFEF891F9E4DFA59FE97F988" box="[332,366,1512,1538]" pageId="25" pageNumber="26">Co</collectorName>
/
<collectorName id="702D5312FFEF891F9E76FA59FE6AF988" box="[375,403,1512,1538]" pageId="25" pageNumber="26">Sc</collectorName>
1000293 to 1000305
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFEF891D9FBCF998FE0CFC55" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
<paragraph id="DD6736C4FFEF891F9FBCF998FEB7F9C9" blockId="25.[189,334,1577,1603]" box="[189,334,1577,1603]" pageId="25" pageNumber="26">
<heading id="862F81A8FFEF891F9FBCF998FEB7F9C9" bold="true" box="[189,334,1577,1603]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="EFACEAD6FFEF891F9FBCF998FEB7F9C9" bold="true" box="[189,334,1577,1603]" pageId="25" pageNumber="26">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFEF891F9FBCF9E9FEB6F91C" blockId="25.[189,1399,1624,1686]" pageId="25" pageNumber="26">MEASUREMENTS (in mm). TL=11.515; BW =67.5; PL=46; PW= 68; EL=6.58; HL=33.5; HW=34.5.</paragraph>
<paragraph id="DD6736C4FFEF891F9FEBF90CFE52F95D" blockId="25.[189,1399,1725,1998]" box="[234,427,1725,1751]" pageId="25" pageNumber="26">
<emphasis id="EFACEAD6FFEF891F9FEBF90CFED3F95D" bold="true" box="[234,298,1725,1751]" pageId="25" pageNumber="26">Male</emphasis>
(
<typeStatus id="02638866FFEF891F9E38F90CFE5AF95D" box="[313,419,1725,1751]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="DD6736C4FFEF891D9FBCF951FDC6FD09" blockId="25.[189,1399,1725,1998]" lastBlockId="27.[189,1399,265,643]" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
Body, elytra, head, legs, antenna and mouthparts reddish, elytra red with a tinge of metallic green; oval, slightly convex. Head (
<figureCitation id="45E32A41FFEF891F9D1BF8B2FD8EF897" box="[538,631,1795,1821]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="25" pageNumber="26">Fig. 9C</figureCitation>
) closely granular with some rugosity, hind part almost smooth; clypeus parabolic, rugulose, margin slightly reflexed; with a feeble emargination in the middle, separated from the vertex by an interrupted frontal carina with a short curved clypeal carina and a blunt tubercle behind; genae with scattered imperceptible punctures. Pronotum (
<figureCitation id="45E32A41FFEF891F9B95F8DCFAF8F80D" box="[1172,1281,1901,1927]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="25" pageNumber="26">Fig. 14C</figureCitation>
) strongly and closely punctured in the middle, less closely at the sides, more strongly and closely pitted anteriorly, with a very slight median line starting above the basal impression, extending almost up to the anterior margin as thin smooth area; anterior sides of pronotum slightly curved, strongly rounded in the middle, bisinuate behind; front angles sharp, base bluntly angulate in the middle; basal fovea deep near the base with imperceptible punctures in circular rows at the periphery. Elytra strongly striate, striae without punctures, elytral intervals finely and sparsely punctured with the 1
<superScript id="2AAD9B8CFFED891D9A62FEC3FA96FE0A" attach="none" box="[1379,1391,370,384]" fontSize="6" pageId="27" pageNumber="28">st</superScript>
, 3
<superScript id="2AAD9B8CFFED891D9FCDFE24FF23FE29" attach="left" box="[204,218,405,419]" fontSize="6" pageId="27" pageNumber="28">rd</superScript>
and the 5
<superScript id="2AAD9B8CFFED891D9E50FE24FEA7FE29" attach="left" box="[337,350,405,419]" fontSize="6" pageId="27" pageNumber="28">th</superScript>
intervals convex, sutural interval shining while the rest opaque. Pygidium not shining, imperceptibly punctured. Metasternum with deep broad longitudinal groove in the front, metasternal shield imperceptibly punctured, with scattered aciculate punctures at the sides anteriorly; sides of the metasternum granular and thinly covered with long hairs. Protibia slightly curved with four strong external teeth and a sharp process at the tip, profemur not toothed; mesofemur (
<figureCitation id="45E32A41FFED891D9B89FD93FB0CFDB6" box="[1160,1269,546,572]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="27" pageNumber="28">Fig. 16C</figureCitation>
) plain and curved near the tibial joint where it forms a sharp tooth-like process on both upper and lower ridges; hind trochanter slightly toothed.
</paragraph>
<caption id="89A7664CFFEC891C9FBCF832FD79F86E" pageId="26" pageNumber="27" startId="26.[189,232,1923,1949]" targetBox="[299,1295,368,1851]" targetPageId="26">
<paragraph id="DD6736C4FFEC891C9FBCF832FD79F86E" blockId="26.[189,1398,1923,2020]" pageId="26" pageNumber="27">
<emphasis id="EFACEAD6FFEC891C9FBCF832FEF6F814" bold="true" box="[189,271,1923,1950]" pageId="26" pageNumber="27">Fig. 7.</emphasis>
<taxonomicName id="1AD84D47FFEC891C9E1BF835FE4CF817" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[282,437,1923,1950]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov.">
<emphasis id="EFACEAD6FFEC891C9E1BF835FE4CF817" box="[282,437,1923,1950]" italics="true" pageId="26" pageNumber="27">Onitis kethai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFEC891C9EBEF835FDE7F814" box="[447,542,1924,1950]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="EFACEAD6FFEC891C9D29F832FDA6F817" bold="true" box="[552,607,1923,1949]" pageId="26" pageNumber="27">AE</emphasis>
. Holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023B.
<emphasis id="EFACEAD6FFEC891C9BEDF832FAF8F817" bold="true" box="[1260,1281,1923,1949]" pageId="26" pageNumber="27">A</emphasis>
. Habitus, dorsal view.
<emphasis id="EFACEAD6FFEC891C9E57F816FE93F84B" bold="true" box="[342,362,1959,1985]" pageId="26" pageNumber="27">B</emphasis>
. Habitus, ventral view.
<emphasis id="EFACEAD6FFEC891C9D8BF817FD66F84A" bold="true" box="[650,671,1958,1984]" pageId="26" pageNumber="27">C</emphasis>
. Head, dorsal view.
<emphasis id="EFACEAD6FFEC891C9C99F816FC54F84B" bold="true" box="[920,941,1959,1985]" pageId="26" pageNumber="27">D</emphasis>
. Pygidium.
<emphasis id="EFACEAD6FFEC891C9B40F816FBACF84B" bold="true" box="[1089,1109,1959,1985]" pageId="26" pageNumber="27">E</emphasis>
. Elytra.
<emphasis id="EFACEAD6FFEC891C9BBEF816FB28F84B" bold="true" box="[1215,1233,1959,1985]" pageId="26" pageNumber="27">F</emphasis>
. Paratype, ♀, habitus, dorsal view. (Images: AIMB).
</paragraph>
</caption>
<paragraph id="DD6736C4FFED891D9FBCFD1AFAA3FD62" blockId="27.[189,1399,682,744]" pageId="27" pageNumber="28">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFED891D9EACFD1BFDD6FD4F" box="[429,559,682,709]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="27" pageNumber="28">Fig. 8AD</figureCitation>
). LP= 2.5, Lp=2.2; BP=1.3, BpB=1, BpT=0.5. Phallobase longer than parameres. Parameres slightly sinuate near the base, curved inwards anteriorly and hooked at the tip.
</paragraph>
<paragraph id="DD6736C4FFED891D9FEBFCBEFEBCFCA3" blockId="27.[189,1399,783,845]" box="[234,325,783,809]" pageId="27" pageNumber="28">
<emphasis id="EFACEAD6FFED891D9FEBFCBEFEBCFCA3" bold="true" box="[234,325,783,809]" pageId="27" pageNumber="28">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFED891D9FBCFC83FA8EFCC6" blockId="27.[189,1399,783,845]" box="[189,1399,818,845]" pageId="27" pageNumber="28">Clypeus closely rugose; protibia with four strong teeth and an articulated spur; no protrusions on the legs.</paragraph>
<paragraph id="DD6736C4FFED891D9FBCFCC2FEE4FC07" blockId="27.[189,285,883,909]" box="[189,285,883,909]" pageId="27" pageNumber="28">
<heading id="862F81A8FFED891D9FBCFCC2FEE4FC07" bold="true" box="[189,285,883,909]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="EFACEAD6FFED891D9FBCFCC2FEE4FC07" bold="true" box="[189,285,883,909]" pageId="27" pageNumber="28">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFED891D9FBCFC13FE0CFC55" blockId="27.[189,1399,930,992]" pageId="27" pageNumber="28">
The collections were made in the cloud forests of Honnametti in Biligiri Rangaswamy Temple Tiger Reserve,
<collectingRegion id="1F1CF826FFED891D9E2BFC74FE5FFC55" box="[298,422,965,991]" country="India" name="Karnataka" pageId="27" pageNumber="28">Karnataka</collectingRegion>
,
<collectingCountry id="A5CF7654FFED891D9EB0FC74FE08FC55" box="[433,497,965,991]" name="India" pageId="27" pageNumber="28">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,284 @@
<document id="F6F209940A3D4C7961C44525EB1468C6" ID-DOI="10.5852/ejt.2024.956.2657" ID-ISSN="2118-9773" ID-ZooBank="9DCDD521-909D-4436-8495-086A11C50DA0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725988271530" checkinUser="plazi" docAuthor="Karimbumkara, Seena Narayanan &amp; Priyadarsanan, Dharma Rajan" docDate="2024" docId="557187D2FFF389019D5AFB71FABDF814" docLanguage="en" docName="EJT.2024.956.1-61.pdf" docOrigin="European Journal of Taxonomy 956" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onitis assamensis Biswas 1980" docType="treatment" docVersion="1" lastPageNumber="8" masterDocId="A948FFAAFFF689069F01FFB1FFF9FF8A" masterDocTitle="Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India" masterLastPageNumber="61" masterPageNumber="1" pageNumber="6" updateTime="1725988442359" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E64CA1F6D06D753C018156FF4569E9FA">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:role id="3A9E8178005AA965A5414034FF1FA2A7">
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<mods:namePart id="BC0FAFB902BD7043C391A3BD46E4331D">Karimbumkara, Seena Narayanan</mods:namePart>
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<mods:affiliation id="900909F59261F260E081636F92E11F7B">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="78C5B48822F40DF3E7C8133F8F3E0027" type="email">seena.narayanan@atree.org</mods:nameIdentifier>
</mods:name>
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<mods:role id="D10F44ACC19C060B3D0476BC5431211F">
<mods:roleTerm id="3C4DCE9E57C8C3F1ABB1CB99A8C2AEE5">Author</mods:roleTerm>
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<mods:namePart id="45F35481498C2671CBA49C3AC0AA316D">Priyadarsanan, Dharma Rajan</mods:namePart>
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<mods:affiliation id="393BB918A55E534C977DB38B36F269E9">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="F8685AE158DB2E2800501520FC3BA576" type="email">priyan@atree.org</mods:nameIdentifier>
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<mods:title id="A93578F7DE434A201F1B6FD67142636E">European Journal of Taxonomy</mods:title>
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<mods:part id="A83BB59C2A87970058FCFA448D238A1F">
<mods:date id="72A2278D65A2E4A52A8DA7C06A27E4D6">2024</mods:date>
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<mods:number id="2656EC15D7ACDFC5A30F82D2F4204D20">2024-09-10</mods:number>
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<mods:number id="158E909DB0948B84BC9251467CA688DD">956</mods:number>
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<mods:start id="1A0ECFB08381B8A33D6C454AAA3E4063">1</mods:start>
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<mods:url id="DA6801B2F81FE6DC2F14F5F033040A52">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287</mods:url>
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<mods:identifier id="99DD5821F1051140C95701F06707A4D3" type="DOI">10.5852/ejt.2024.956.2657</mods:identifier>
<mods:identifier id="BA20EE0A3F1417B9E5FF655CA56014D8" type="ISSN">2118-9773</mods:identifier>
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<treatment id="557187D2FFF389019D5AFB71FABDF814" LSID="urn:lsid:plazi:treatment:557187D2FFF389019D5AFB71FABDF814" httpUri="http://treatment.plazi.org/id/557187D2FFF389019D5AFB71FABDF814" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="95C2654FFFF389039D5AFB71FC20FB50" box="[603,985,1216,1242]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="DD6736C4FFF389039D5AFB71FC20FB50" blockId="5.[603,985,1216,1277]" box="[603,985,1216,1242]" pageId="5" pageNumber="6">
<heading id="862F81A8FFF389039D5AFB71FC20FB50" box="[603,985,1216,1242]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
<taxonomicName id="1AD84D47FFF389039D5AFB71FC20FB50" ID-CoL="6SMW9" authority="Biswas, 1980" authorityName="Biswas" authorityYear="1980" box="[603,985,1216,1242]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF389039D5AFB71FCCAFB50" bold="true" box="[603,819,1216,1242]" italics="true" pageId="5" pageNumber="6">Onitis assamensis</emphasis>
<bibRefCitation id="B9494B35FFF389039C3BFB71FC20FB50" author="Biswas S." box="[826,985,1216,1242]" pageId="5" pageNumber="6" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF389039DB5FB52FC81FAB4" pageId="5" pageNumber="6" type="description">
<paragraph id="DD6736C4FFF389039DB5FB52FC86FB77" blockId="5.[603,985,1216,1277]" box="[692,895,1251,1277]" pageId="5" pageNumber="6">
<figureCitation id="45E32A41FFF389039DB5FB52FD04FB77" box="[692,765,1251,1277]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[262,1340,366,1854]" captionTargetId="figure-1@6.[189,1398,315,1882]" captionTargetPageId="6" captionText="Fig. 1. Onitis assamensis Biswas, 1980, ♂ (AD), AIMB/Co/Sc1000001; ♀ (EF), AIMB/Co/ Sc1000009. A. Habitus, dorsal view. B. Habitus, ventral view. C. Protibia. D. Pygidium. E. Habitus, dorsal view. F. Habitus, ventral view. (Images: AIMB)" pageId="5" pageNumber="6">Figs 1</figureCitation>
,
<figureCitation id="45E32A41FFF389039C0AFB52FCAFFB77" box="[779,854,1251,1277]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="5" pageNumber="6">4AD</figureCitation>
,
<figureCitation id="45E32A41FFF389039C63FB52FC86FB77" box="[866,895,1251,1277]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." pageId="5" pageNumber="6">17</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFA94FC81FAB4" blockId="5.[189,888,1316,1343]" box="[189,888,1316,1343]" pageId="5" pageNumber="6">
<emphasis id="EFACEAD6FFF389039FBCFA94FE69FAB5" box="[189,400,1317,1343]" italics="true" pageId="5" pageNumber="6">Onitisassamensis</emphasis>
<bibRefCitation id="B9494B35FFF389039E96FA95FD88FAB5" author="Biswas S." box="[407,625,1316,1343]" pageId="5" pageNumber="6" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980: 339</bibRefCitation>
(original description).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF389039FBCFAD7FAAEFA0A" box="[189,1367,1382,1409]" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="DD6736C4FFF389039FBCFAD7FAAEFA0A" blockId="5.[189,1367,1382,1409]" box="[189,1367,1382,1409]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="FDC811EAFFF389039FBCFAD7FAAEFA0A" box="[189,1367,1382,1409]" pageId="5" pageNumber="6">
<taxonomicName id="1AD84D47FFF389039FBCFAD7FE69FA0A" ID-CoL="6SMW9" authorityName="Biswas" authorityYear="1980" box="[189,400,1382,1408]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF389039FBCFAD7FE69FA0A" box="[189,400,1382,1408]" italics="true" pageId="5" pageNumber="6">Onitis assamensis</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFF389039EACFAD7FD3AFA0A" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[429,707,1382,1409]" page="1037" pageId="5" pageNumber="6" year="2015">
<bibRefCitation id="B9494B35FFF389039EACFAD7FD3AFA0A" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[429,707,1382,1409]" pageId="5" pageNumber="6" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFF389039EFDFAD6FDCEFA0A" box="[508,567,1382,1408]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(checklist). —
<bibRefCitation id="B9494B35FFF389039C7BFAD7FB8DFA0A" author="Schoolmeesters P." box="[890,1140,1382,1408]" pageId="5" pageNumber="6" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF389039FBCFA16FDECF883" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="DD6736C4FFF389039FBCFA16FE3FFA4B" blockId="5.[189,454,1447,1473]" box="[189,454,1447,1473]" pageId="5" pageNumber="6">
<heading id="862F81A8FFF389039FBCFA16FE3FFA4B" bold="true" box="[189,454,1447,1473]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="EFACEAD6FFF389039FBCFA16FE3FFA4B" bold="true" box="[189,454,1447,1473]" pageId="5" pageNumber="6">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFA67FDECF883" blockId="5.[189,1399,1494,1802]" pageId="5" pageNumber="6">
<taxonomicName id="1AD84D47FFF389039FBCFA67FE6CFA7A" authorityName="Biswas" authorityYear="1980" box="[189,405,1494,1520]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF389039FBCFA67FE6CFA7A" box="[189,405,1494,1520]" italics="true" pageId="5" pageNumber="6">Onitis assamensis</emphasis>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFF389039ECFFA67FDA3FA7A" authorityName="Cambefort" authorityYear="1988" box="[462,602,1494,1520]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFF389039ECFFA67FDA3FA7A" box="[462,602,1494,1520]" italics="true" pageId="5" pageNumber="6">O. naviauxi</emphasis>
</taxonomicName>
differ from each other in the number of teeth on the anteriomedian ridge of the protibia in the male. Clypeal carina short in male of
<taxonomicName id="1AD84D47FFF389039CF5FA4BFB5CF999" authorityName="Biswas" authorityYear="1980" box="[1012,1189,1530,1556]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF389039CF5FA4BFB5CF999" box="[1012,1189,1530,1556]" italics="true" pageId="5" pageNumber="6">O. assamensis</emphasis>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFF389039BE7FA4BFA8FF999" authorityName="Cambefort" authorityYear="1988" box="[1254,1398,1530,1556]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFF389039BE7FA4BFA8FF999" box="[1254,1398,1530,1556]" italics="true" pageId="5" pageNumber="6">O. naviauxi</emphasis>
</taxonomicName>
while it is longer in female; clypeus in female pointed in both species; metasternal shield hairy in males and smooth in females. In the original description of
<taxonomicName id="1AD84D47FFF389039CCBF9F1FB80F9D3" authorityName="Biswas" authorityYear="1980" box="[970,1145,1600,1626]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF389039CCBF9F1FB80F9D3" box="[970,1145,1600,1626]" italics="true" pageId="5" pageNumber="6">O. assamensis</emphasis>
</taxonomicName>
the frontal carina is considered entire, but on observing the specimen in detail, it is confirmed that it is interrupted due to the placement of tubercle on it. The teeth on the infero-internal ridge in the profemur is a very prominent character of the species and the arrangement of teeth, their number and size may vary with individuals. In a specimen we collected, an extra small tooth was present near the femoro-tibial joint on one of the fore legs.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF389019FBCF880FEB3FE07" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="DD6736C4FFF389039FBCF880FE53F8C1" blockId="5.[189,426,1841,1867]" box="[189,426,1841,1867]" pageId="5" pageNumber="6">
<heading id="862F81A8FFF389039FBCF880FE53F8C1" bold="true" box="[189,426,1841,1867]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="EFACEAD6FFF389039FBCF880FE53F8C1" bold="true" box="[189,426,1841,1867]" pageId="5" pageNumber="6">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF389039FEBF8D1FEA5F8F0" blockId="5.[189,1399,1888,1984]" box="[234,348,1888,1914]" pageId="5" pageNumber="6">
<emphasis id="EFACEAD6FFF389039FEBF8D1FEA5F8F0" bold="true" box="[234,348,1888,1914]" pageId="5" pageNumber="6">
<typeStatus id="02638866FFF389039FEBF8D1FEA5F8F0" box="[234,348,1888,1914]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCF832FE0FF84A" blockId="5.[189,1399,1888,1984]" pageId="5" pageNumber="6">
<materialsCitation id="6DB03C99FFF389039FBCF832FE0FF84A" collectingDate="1972-12-21" collectionCode="ZSIM" collectorName="A. K. Ghosh &amp; Regd. No. A" country="India" location="Kaziranga National Park" pageId="5" pageNumber="6" specimenCount="1" specimenCount-female="1" stateProvince="Assam" typeStatus="holotype">
<collectingCountry id="A5CF7654FFF389039FBCF832FEF7F817" box="[189,270,1923,1949]" name="India" pageId="5" pageNumber="6">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFF389039E2AF832FEBBF817" box="[299,322,1923,1949]" count="1" pageId="5" pageNumber="6" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFF389039E51F832FE5AF817" box="[336,419,1923,1949]" country="India" name="Assam" pageId="5" pageNumber="6">Assam</collectingRegion>
,
<location id="D807601FFFF389039EB0F832FD22F817" LSID="urn:lsid:plazi:treatment:557187D2FFF389019D5AFB71FABDF814:D807601FFFF389039EB0F832FD22F817" box="[433,731,1923,1949]" country="India" name="Kaziranga National Park" pageId="5" pageNumber="6" stateProvince="Assam">Kaziranga National Park</location>
;
<date id="A9661004FFF389039DE9F832FC75F817" box="[744,908,1923,1949]" pageId="5" pageNumber="6" value="1972-12-21">
<collectingDate id="B922E9ECFFF389039DE9F832FC75F817" box="[744,908,1923,1949]" pageId="5" pageNumber="6" value="1972-12-21">21 Dec. 1972</collectingDate>
</date>
;
<collectorName id="702D5312FFF389039C99F832FBDEF817" box="[920,1063,1923,1949]" pageId="5" pageNumber="6">A.K. Ghosh</collectorName>
leg.; from rhinoceros dung;
<collectionCode id="BBC9AE01FFF389039FBCF817FEFAF84A" box="[189,259,1958,1984]" pageId="5" pageNumber="6">ZSIM</collectionCode>
<collectorName id="702D5312FFF389039E0BF817FE66F84A" box="[266,415,1958,1984]" pageId="5" pageNumber="6">Regd. No. A</collectorName>
1/4397.
</materialsCitation>
</paragraph>
<caption id="89A7664CFFF089009FBCF839FCC7F862" pageId="6" pageNumber="7" startId="6.[189,232,1928,1954]" targetBox="[262,1340,366,1854]" targetPageId="6">
<paragraph id="DD6736C4FFF089009FBCF839FCC7F862" blockId="6.[189,1398,1928,2025]" pageId="6" pageNumber="7">
<emphasis id="EFACEAD6FFF089009FBCF839FEECF828" bold="true" box="[189,277,1928,1954]" pageId="6" pageNumber="7">Fig. 1.</emphasis>
<taxonomicName id="1AD84D47FFF089009E24F839FD40F828" authority="Biswas, 1980" authorityName="Biswas" authorityYear="1980" box="[293,697,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF089009E24F839FDF8F828" box="[293,513,1928,1954]" italics="true" pageId="6" pageNumber="7">Onitis assamensis</emphasis>
<bibRefCitation id="B9494B35FFF089009D13F839FD40F828" author="Biswas S." box="[530,697,1928,1954]" pageId="6" pageNumber="7" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980</bibRefCitation>
</taxonomicName>
, ♂ (AD), AIMB/Co/Sc1000001; ♀ (EF), AIMB/Co/ Sc1000009.
<emphasis id="EFACEAD6FFF089009E53F81AFE9EF84F" bold="true" box="[338,359,1963,1989]" pageId="6" pageNumber="7">A</emphasis>
. Habitus, dorsal view.
<emphasis id="EFACEAD6FFF089009D7FF81AFD6BF84F" bold="true" box="[638,658,1963,1989]" pageId="6" pageNumber="7">B</emphasis>
. Habitus, ventral view.
<emphasis id="EFACEAD6FFF089009CB3F81AFC3EF84F" bold="true" box="[946,967,1963,1989]" pageId="6" pageNumber="7">C</emphasis>
. Protibia.
<emphasis id="EFACEAD6FFF089009B47F81AFBA2F84F" bold="true" box="[1094,1115,1963,1989]" pageId="6" pageNumber="7">D</emphasis>
. Pygidium.
<emphasis id="EFACEAD6FFF089009BEEF81AFAFAF84F" bold="true" box="[1263,1283,1963,1989]" pageId="6" pageNumber="7">E</emphasis>
. Habitus, dorsal view.
<emphasis id="EFACEAD6FFF089009E53F87FFE9DF862" bold="true" box="[338,356,1998,2024]" pageId="6" pageNumber="7">F</emphasis>
. Habitus, ventral view. (Images: AIMB)
</paragraph>
</caption>
<paragraph id="DD6736C4FFF189019FEBFEB8FD62FEA9" blockId="7.[189,1398,265,397]" box="[234,667,265,292]" pageId="7" pageNumber="8">
<emphasis id="EFACEAD6FFF189019FEBFEB8FE18FEA9" bold="true" box="[234,481,265,291]" pageId="7" pageNumber="8">Additional material</emphasis>
(
<specimenCount id="CBDEFD4DFFF189019EF3FEB8FD6AFEA9" box="[498,659,265,292]" count="15" pageId="7" pageNumber="8" type="generic">15 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFF189019FBCFE9DFEB3FE07" blockId="7.[189,1398,265,397]" pageId="7" pageNumber="8">
<materialsCitation id="6DB03C99FFF189019FBCFE9DFEBCFE07" collectingDate="2019-05-21" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Karimbumkara Seena Narayanan &amp; Co &amp; Sc" country="India" location="Kaziranga National Park" pageId="7" pageNumber="8" specimenCount="15" specimenCount-female="7" specimenCount-male="8" stateProvince="Assam">
<collectingCountry id="A5CF7654FFF189019FBCFE9DFEF7FECC" box="[189,270,300,326]" name="India" pageId="7" pageNumber="8">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFF189019E2AFE9DFE88FECD" box="[299,369,300,327]" count="8" pageId="7" pageNumber="8" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFF189019E7EFE9CFE3DFECD" box="[383,452,301,327]" count="7" pageId="7" pageNumber="8" type="female">7 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFF189019ED2FE9DFDDFFECC" box="[467,550,300,326]" country="India" name="Assam" pageId="7" pageNumber="8">Assam</collectingRegion>
,
<location id="D807601FFFF189019D32FE9DFCA4FECC" LSID="urn:lsid:plazi:treatment:557187D2FFF389019D5AFB71FABDF814:D807601FFFF189019D32FE9DFCA4FECC" box="[563,861,300,326]" country="India" name="Kaziranga National Park" pageId="7" pageNumber="8" stateProvince="Assam">Kaziranga National Park</location>
;
<date id="A9661004FFF189019C6BFE9DFBF4FECC" box="[874,1037,300,327]" pageId="7" pageNumber="8" value="2019-05-21">
<collectingDate id="B922E9ECFFF189019C6BFE9DFBF4FECC" box="[874,1037,300,327]" pageId="7" pageNumber="8" value="2019-05-21">21 May 2019</collectingDate>
</date>
;
<collectorName id="702D5312FFF189019B1AFE9DFA8FFECC" box="[1051,1398,300,326]" pageId="7" pageNumber="8">Priyadarsanan Dharma Rajan</collectorName>
&amp;
<collectorName id="702D5312FFF189019FDCFEFEFD9CFEE0" box="[221,613,335,362]" pageId="7" pageNumber="8">Karimbumkara Seena Narayanan</collectorName>
leg.; handpicked, rhinoceros dung;
<collectionCode id="BBC9AE01FFF189019B1EFEE1FB39FEE3" box="[1055,1216,335,362]" pageId="7" pageNumber="8">AIMB AIMB</collectionCode>
/
<collectorName id="702D5312FFF189019BCBFEFEFB15FEE3" box="[1226,1260,335,361]" pageId="7" pageNumber="8">Co</collectorName>
/
<collectorName id="702D5312FFF189019BF4FEFEFAE8FEE3" box="[1269,1297,335,361]" pageId="7" pageNumber="8">Sc</collectorName>
1000001 to 1000015
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF189019FBCFE05FED5F984" pageId="7" pageNumber="8" type="description">
<paragraph id="DD6736C4FFF189019FBCFE05FEB7FE44" blockId="7.[189,334,436,462]" box="[189,334,436,462]" pageId="7" pageNumber="8">
<heading id="862F81A8FFF189019FBCFE05FEB7FE44" bold="true" box="[189,334,436,462]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="EFACEAD6FFF189019FBCFE05FEB7FE44" bold="true" box="[189,334,436,462]" pageId="7" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF189019FBCFE52FDD8FDAB" blockId="7.[189,1399,482,545]" pageId="7" pageNumber="8">MEASUREMENTS (in mm). TL=21.524.8; BW =10.712.5; PL=8.511; PW= 10.513.5; EL=10.5 12.5; HL= 56; HW=5.56.8.</paragraph>
<paragraph id="DD6736C4FFF189019FEBFDF6FED3FDEB" blockId="7.[189,1399,583,1103]" box="[234,298,583,609]" pageId="7" pageNumber="8">
<emphasis id="EFACEAD6FFF189019FEBFDF6FED3FDEB" bold="true" box="[234,298,583,609]" pageId="7" pageNumber="8">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFF189019FBCFDDBFAAFFBC5" blockId="7.[189,1399,583,1103]" pageId="7" pageNumber="8">Body shiny green or bronzy, broadly oval. Head rounded, clypeus straight in the middle anteriorly, finely rugose with scattered punctures; clypeal carina short, rugose behind; frontal carina almost straight, narrowly interrupted in the middle by the placement of a strong conical blunt tubercle; vertex sparsely punctured, elevated towards the eyes with a depression in the middle devoid of any punctures. Pronotum with small, close dense punctures between the basal foveae, punctures larger and less close anterior to the foveae, finer and sparse anteriorly and at the sides; basal foveae oval, deep, connected by a bordered base; scutellum triangular visibly large. Elytra broadly curved at the sides; striae deep, broader at the base, narrow and smooth towards the tip; elytral intervals smooth without punctures, sutural interval slightly convex. Pygidium margined, lesser sheen, imperceptibly punctured. Metasternal shield with minute punctures, front angles with minute granules and fine yellow hairs; sides of metasternum with reddish hairs, densely granular. Fore legs with rows of reddish long hairs or setae on the ridges; a strong curved tooth on the anteromedian ridge of profemur with a minute oblique denticle near the base; protibia with four distantly placed external teeth, the infero-medial ridge with 67 teeth, small proximally and stouter distally; mid femur unarmed; hind femur with a strong curved lateral tooth towards the apex.</paragraph>
<paragraph id="DD6736C4FFF189019FBCFBC7FD7EFB5C" blockId="7.[189,1399,1141,1238]" pageId="7" pageNumber="8">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFF189019EBFFBC4FDBFFB1A" box="[446,582,1141,1168]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="7" pageNumber="8">Fig. 4AD</figureCitation>
). LP=3.6, Lp =2.86; BP=1.86, BpB=1.43, BpT= 0.71. Phallobase longer than parameres, broader at base and further tapers, becomes blunt and slightly curved at the tip. Phallobase slightly broader at the base.
</paragraph>
<paragraph id="DD6736C4FFF189019FEBFB4CFEBCFA9D" blockId="7.[189,1399,1277,1550]" box="[234,325,1277,1303]" pageId="7" pageNumber="8">
<emphasis id="EFACEAD6FFF189019FEBFB4CFEBCFA9D" bold="true" box="[234,325,1277,1303]" pageId="7" pageNumber="8">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFF189019FBCFA91FED5F984" blockId="7.[189,1399,1277,1550]" pageId="7" pageNumber="8">Body greenish, broadly oval, not very convex; head sub triangular, frontal carina entire with a tubercle. Clypeal carina long and curved placed almost midway between the clypeal margin and the frontal carina; clypeus parabolic, rugulose. Pronotum strongly and closely punctured; front angles sharp; basal foveae rounded. Elytra: dull, striae shallow and broad, intervals flat, not perceptibly punctate. Ventral surface: metasternum flat, smooth with two deep pits behind and a shallow longitudinal groove between; sides of metasternum closely granular and hairy. Pygidium without hair, slightly shining and coarse; legs unarmed.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF189019FBCF984FC0CF9F4" box="[189,1013,1589,1662]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="DD6736C4FFF189019FBCF984FEAFF9C5" blockId="7.[189,342,1589,1615]" box="[189,342,1589,1615]" pageId="7" pageNumber="8">
<heading id="862F81A8FFF189019FBCF984FEAFF9C5" bold="true" box="[189,342,1589,1615]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="EFACEAD6FFF189019FBCF984FEAFF9C5" bold="true" box="[189,342,1589,1615]" pageId="7" pageNumber="8">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF189019FBCF9D5FC0CF9F4" blockId="7.[189,1013,1635,1662]" box="[189,1013,1635,1662]" pageId="7" pageNumber="8">
<collectingCountry id="A5CF7654FFF189019FBCF9D5FF07F9F4" box="[189,254,1636,1662]" name="India" pageId="7" pageNumber="8">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFF189019E06F9D5FEAEF9F4" box="[263,343,1636,1662]" country="India" name="Assam" pageId="7" pageNumber="8">Assam</collectingRegion>
(
<bibRefCitation id="B9494B35FFF189019E67F9D5FDFBF9F4" author="Biswas S." box="[358,514,1636,1662]" pageId="7" pageNumber="8" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas 1980</bibRefCitation>
;
<bibRefCitation id="B9494B35FFF189019D0FF9D5FD24F9F4" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[526,733,1635,1662]" pageId="7" pageNumber="8" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFF189019D5CF9D5FD61F9F4" box="[605,664,1636,1662]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFF189019DE8F9D5FC10F9F4" author="Schoolmeesters P." box="[745,1001,1636,1662]" pageId="7" pageNumber="8" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF189019FBCF914FABDF814" pageId="7" pageNumber="8" type="discussion">
<paragraph id="DD6736C4FFF189019FBCF914FED6F935" blockId="7.[189,303,1701,1727]" box="[189,303,1701,1727]" pageId="7" pageNumber="8">
<heading id="862F81A8FFF189019FBCF914FED6F935" bold="true" box="[189,303,1701,1727]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="EFACEAD6FFF189019FBCF914FED6F935" bold="true" box="[189,303,1701,1727]" pageId="7" pageNumber="8">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF189019FBCF965FABDF814" blockId="7.[189,1399,1748,1950]" pageId="7" pageNumber="8">
A single female specimen of
<taxonomicName id="1AD84D47FFF189019D14F965FD47F967" authorityName="Biswas" authorityYear="1980" box="[533,702,1748,1774]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF189019D14F965FD47F967" box="[533,702,1748,1774]" italics="true" pageId="7" pageNumber="8">O. assamensis</emphasis>
</taxonomicName>
was collected by
<bibRefCitation id="B9494B35FFF189019C93F965FBC4F964" author="Biswas S." box="[914,1085,1748,1774]" pageId="7" pageNumber="8" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas (1980)</bibRefCitation>
from rhinoceros dung and described as the
<typeStatus id="02638866FFF189019E7FF946FE10F89B" box="[382,489,1783,1809]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
. During our visit to Kaziranga National Park, we have collected
<specimenCount id="CBDEFD4DFFF189019BD9F946FA8FF89B" box="[1240,1398,1783,1810]" count="15" pageId="7" pageNumber="8" type="generic">15 specimens</specimenCount>
of the species of which eight were males and
<specimenCount id="CBDEFD4DFFF189019DEBF8AAFC6FF8BE" box="[746,918,1818,1845]" count="7" pageId="7" pageNumber="8" type="female">seven females</specimenCount>
. The rarity of the species mentioned by Biswas can be attributed to the winter in December when he had made the collection. The species was common and active in May (pre-monsoon), when we collected them at the same location. We have described the male of
<emphasis id="EFACEAD6FFF189019EC1F835FD6AF814" box="[448,659,1924,1950]" italics="true" pageId="7" pageNumber="8">Onitisassamensis</emphasis>
which had never previously been discovered or described.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,627 @@
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<mods:title id="E714754C24BC6E0C85BF1C0988B72AE3">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<subSubSection id="95C2654FFFF589059D6CFA62FC3EFA67" box="[621,967,1491,1517]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="DD6736C4FFF589059D6CFA62FC3EFA67" blockId="3.[621,967,1491,1517]" box="[621,967,1491,1517]" pageId="3" pageNumber="4">
<heading id="862F81A8FFF589059D6CFA62FC3EFA67" box="[621,967,1491,1517]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
Genus
<taxonomicName id="1AD84D47FFF589059DBEFA62FC3EFA67" ID-CoL="68C6" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[703,967,1491,1517]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EFACEAD6FFF589059DBEFA62FCF1FA67" bold="true" box="[703,776,1491,1517]" italics="true" pageId="3" pageNumber="4">Onitis</emphasis>
<bibRefCitation id="B9494B35FFF589059C11FA62FC3EFA67" author="Fabricius J. C." box="[784,967,1491,1517]" pageId="3" pageNumber="4" refId="ref29024" refString="Fabricius J. C. 1798. Supplementum Entomologiae Systematicae. Proft et Storch, Copenhagen [Hafniae]. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1798">Fabricius,1798</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF589039FBCF9A9FA8FFBF0" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="key">
<paragraph id="DD6736C4FFF589059FBCF9A9FB93F9BE" blockId="3.[189,1130,1559,1588]" box="[189,1130,1559,1588]" pageId="3" pageNumber="4">
<emphasis id="EFACEAD6FFF589059FBCF9A9FB93F9BE" bold="true" box="[189,1130,1559,1588]" pageId="3" pageNumber="4">
Key to species of
<taxonomicName id="1AD84D47FFF589059EA9F9A6FD28F9BE" ID-CoL="68C6" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[424,721,1559,1588]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EFACEAD6FFF589059EA9F9A6FE01F9B9" bold="true" box="[424,504,1559,1587]" italics="true" pageId="3" pageNumber="4">Onitis</emphasis>
<bibRefCitation id="B9494B35FFF589059D01F9A6FD28F9BE" author="Fabricius J. C." box="[512,721,1559,1588]" pageId="3" pageNumber="4" refId="ref29024" refString="Fabricius J. C. 1798. Supplementum Entomologiae Systematicae. Proft et Storch, Copenhagen [Hafniae]. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1798">Fabricius, 1798</bibRefCitation>
</taxonomicName>
from the Indian subcontinent
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFF589059FBCF9FCFD9DF95B" blockId="3.[189,1399,1613,1745]" pageId="3" pageNumber="4">
The following key is based on external morphological characters and can be used to identify both male and female adults of
<taxonomicName id="1AD84D47FFF589059EACF9C0FE0CF901" ID-CoL="68C6" authorityName="Fabricius" authorityYear="1798" box="[429,501,1649,1675]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="EFACEAD6FFF589059EACF9C0FE0CF901" box="[429,501,1649,1675]" italics="true" pageId="3" pageNumber="4">Onitis</emphasis>
</taxonomicName>
. For the convenience of identification, we have retained the groups suggested by
<bibRefCitation id="B9494B35FFF589059FE0F925FE67F924" author="Janssens A." box="[225,414,1684,1710]" pageId="3" pageNumber="4" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens (1937)</bibRefCitation>
. The keys have been modified from the identification keys to the species provided by
<bibRefCitation id="B9494B35FFF589059FBCF906FEA6F95B" author="Arrow G. J." box="[189,351,1719,1745]" pageId="3" pageNumber="4" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow (1931)</bibRefCitation>
and
<bibRefCitation id="B9494B35FFF589059E99F906FD99F95B" author="Balthasar V." box="[408,608,1719,1745]" pageId="3" pageNumber="4" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar (1963)</bibRefCitation>
.
</paragraph>
<key id="A94961F5FFF589039FBCF94BFA8FFBF0" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
<keyStep id="662C2E61FFF589059FBCF94BFA8FF80A" pageId="3" pageNumber="4">
<paragraph id="DD6736C4FFF589059FBCF94BFA8EF8B2" blockId="3.[189,1399,1786,1920]" pageId="3" pageNumber="4">
<keyLead id="662995F1FFF589059FBCF94BFA8EF8B2" pageId="3" pageNumber="4">1. Head with frontal carina interrupted medially, with tubercle placed adjoining the carina; clypeus always with a short or long transverse carina.................................................................................... 2</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF589059FBCF8F2FA8FF80A" blockId="3.[189,1399,1786,1920]" pageId="3" pageNumber="4">
<keyLead id="662995F1FFF589059FBCF8F2FA8FF80A" pageId="3" pageNumber="4"> Head with frontal carina uninterrupted or entire, with tubercle behind or absent; clypeal carina feebly expressed.......................................................................................................................................... 13</keyLead>
</paragraph>
</keyStep>
<paragraph id="DD6736C4FFF589059FBCF818FA8EF849" blockId="3.[189,1399,1961,2025]" box="[189,1399,1961,1988]" pageId="3" pageNumber="4">
<keyLead id="662995F1FFF589059FBCF818FA8EF849" box="[189,1399,1961,1988]" pageId="3" pageNumber="4">
<keyStep id="662C2E61FFF589059FBCF818FA8EF849" box="[189,1399,1961,1988]" pageId="3" pageNumber="4">
2. Body completely black or dark brown (
<emphasis id="EFACEAD6FFF589059DB8F818FCCDF849" bold="true" box="[697,820,1961,1987]" pageId="3" pageNumber="4">GROUP I</emphasis>
) .......................................................................... 3
</keyStep>
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF589029FBCF87EFA8EFEA9" blockId="3.[189,1399,1961,2025]" lastBlockId="4.[189,1399,265,328]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<keyLead id="662995F1FFF589029FBCF87EFA8EFEA9" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Body metallic, with coppery or green tinge....................................................................................... 6
<keyStep id="662C2E61FFF289029FBCFEB8FA8EFEA9" box="[189,1399,265,292]" pageId="4" pageNumber="5">
3. Pronotum minutely punctured (
<figureCitation id="45E32A41FFF289029D5BFEB8FD2BFEAE" box="[602,722,265,292]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="4" pageNumber="5">Fig. 5EF</figureCitation>
)........................................................................................ 4
</keyStep>
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289029FBCFE9FFA8EFEC2" blockId="4.[189,1399,265,328]" box="[189,1399,301,328]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFE9FFA8EFEC2" box="[189,1399,301,328]" pageId="4" pageNumber="5">
Pronotum strongly punctured (
<figureCitation id="45E32A41FFF289029D53FE9CFD26FECD" box="[594,735,301,327]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Fig. 14AB</figureCitation>
)...................................................................................... 5
</keyLead>
</paragraph>
<keyStep id="662C2E61FFF289029FBCFEC1FA8FFE7F" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFEC1FA8FFE27" blockId="4.[189,1399,368,501]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFEC1FA8FFE27" pageId="4" pageNumber="5">
4. Metasternal shield in male with deep transverse excavation in the middle; without an anterior longitudinal groove ...................................................
<taxonomicName id="1AD84D47FFF289029C4BFE22FB86FE27" ID-CoL="49KG2" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[842,1151,403,429]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="EFACEAD6FFF289029C4BFE22FC1CFE27" bold="true" box="[842,997,403,429]" italics="true" pageId="4" pageNumber="5">O. excavatus</emphasis>
<bibRefCitation id="B9494B35FFF289029CEAFE22FB86FE27" author="Arrow G. J." box="[1003,1151,403,429]" pageId="4" pageNumber="5" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029B8FFE22FB14FE24" box="[1166,1261,403,430]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="4" pageNumber="5">Figs 5E</figureCitation>
,
<figureCitation id="45E32A41FFF289029BF8FE22FABDFE27" box="[1273,1348,403,429]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="4" pageNumber="5">6AD</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EFE22FA94FE27" box="[1359,1389,403,429]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="4" pageNumber="5">18</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289029FBCFE09FA8FFE7F" blockId="4.[189,1399,368,501]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFE09FA8FFE7F" pageId="4" pageNumber="5">
Metasternal shield without transverse excavation in the middle, with an anterior longitudinal groove ......................................................................
<taxonomicName id="1AD84D47FFF289029C46FE6BFB7FFE7E" ID-CoL="49KG5" authority="(Wulfen, 1786)" baseAuthorityName="Wulfen" baseAuthorityYear="1786" box="[839,1158,474,501]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFF289029C46FE6BFC32FE7F" bold="true" box="[839,971,474,501]" italics="true" pageId="4" pageNumber="5">O. falcatus</emphasis>
(
<bibRefCitation id="B9494B35FFF289029CDBFE6BFB87FE7E" author="Wulfen X." box="[986,1150,474,500]" pageId="4" pageNumber="5" refId="ref31404" refString="Wulfen X. 1786. Descriptiones quorumdam Capensium insectorum. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1786">Wulfen, 1786</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029B94FE6BFB09FE7F" box="[1173,1264,474,501]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="4" pageNumber="5">Figs 5F</figureCitation>
,
<figureCitation id="45E32A41FFF289029BFDFE6BFABAFE7E" box="[1276,1347,474,500]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="4" pageNumber="5">6EH</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EFE6AFA94FE7F" box="[1359,1389,475,501]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="4" pageNumber="5">18</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCFDACFA8FFD28" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFDACFA8FFDD0" blockId="4.[189,1398,541,674]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFDACFA8FFDD0" pageId="4" pageNumber="5">
5. Pygidium with a median longitudinal groove; protibia with a rudimentary spur before the extremity (
<figureCitation id="45E32A41FFF289029FF2FDF1FEA7FDD0" box="[243,350,576,602]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Fig. 16A</figureCitation>
) .....................................
<taxonomicName id="1AD84D47FFF289029D88FDF1FCB1FDD1" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[649,840,576,603]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bhomorensis" status="sp. nov.">
<emphasis id="EFACEAD6FFF289029D88FDF1FCB1FDD1" bold="true" box="[649,840,576,603]" italics="true" pageId="4" pageNumber="5">O. bhomorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFF289029C4EFDF0FC53FDD1" box="[847,938,577,603]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="45E32A41FFF289029CB8FDF1FBFCFDD0" box="[953,1029,576,602]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[245,1340,356,1841]" captionTargetId="figure-1@9.[189,1398,265,1856]" captionTargetPageId="9" captionText="Fig. 2. Onitisbhomorensis sp. nov., holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head and prontum. D. Protibia, arrow showing rudimentary spur. E. Metasternum. F. Pygidium, arrow showing longitudinal groove. (Images: AIMB)." pageId="4" pageNumber="5">Figs 2</figureCitation>
,
<figureCitation id="45E32A41FFF289029B13FDF1FBB4FDD0" box="[1042,1101,576,602]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="4" pageNumber="5">6IL</figureCitation>
,
<figureCitation id="45E32A41FFF289029B58FDF1FB79FDD0" box="[1113,1152,576,602]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">9A</figureCitation>
,
<figureCitation id="45E32A41FFF289029B8DFDF1FB38FDD0" box="[1164,1217,576,602]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">14A</figureCitation>
,
<figureCitation id="45E32A41FFF289029BCCFDF1FAFBFDD1" box="[1229,1282,576,603]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">15A</figureCitation>
,
<figureCitation id="45E32A41FFF289029A0FFDF1FABDFDD0" box="[1294,1348,576,602]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">16A</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EFDF1FA94FDD0" box="[1359,1389,576,602]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." pageId="4" pageNumber="5">17</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289029FBCFDD4FA8FFD28" blockId="4.[189,1398,541,674]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFDD4FA8FFD28" pageId="4" pageNumber="5">
Pygidium without a median longitudinal groove; protibia without a rudimentary spur (
<figureCitation id="45E32A41FFF289029BDFFDD5FABEFDF4" box="[1246,1351,612,638]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Fig. 16B</figureCitation>
).... .................................
<taxonomicName id="1AD84D47FFF289029EEBFD39FC71FD28" ID-CoL="49KKQ" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[490,904,648,674]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFF289029EEBFD39FD2BFD28" bold="true" box="[490,722,648,674]" italics="true" pageId="4" pageNumber="5">O. punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFF289029DD8FD39FC71FD28" author="Janssens A." box="[729,904,648,674]" pageId="4" pageNumber="5" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029C96FD39FBE3FD28" box="[919,1050,648,674]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="4" pageNumber="5">Figs 6MP</figureCitation>
,
<figureCitation id="45E32A41FFF289029B27FD39FBB3FD28" box="[1062,1098,648,674]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">9B</figureCitation>
,
<figureCitation id="45E32A41FFF289029B56FD39FB7CFD28" box="[1111,1157,648,674]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="4" pageNumber="5">10F</figureCitation>
,
<figureCitation id="45E32A41FFF289029B90FD39FB3DFD28" box="[1169,1220,648,674]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">14B</figureCitation>
,
<figureCitation id="45E32A41FFF289029BD1FD36FAFAFD28" box="[1232,1283,647,674]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">15B</figureCitation>
,
<figureCitation id="45E32A41FFF289029A11FD39FABAFD28" box="[1296,1347,648,674]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">16B</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EFD39FA94FD28" box="[1359,1389,648,674]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." pageId="4" pageNumber="5">19</figureCitation>
)
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</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCFD7BFA8FFCC5" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFD7BFA8EFC82" blockId="4.[189,1399,714,847]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFD7BFA8EFC82" pageId="4" pageNumber="5">
6. Vertex covered with granules or highly raspy punctuation; clypeal carina closer to the frontal carina than to the clypeal margin (
<figureCitation id="45E32A41FFF289029D2DFD5CFD51FC82" box="[556,680,749,776]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Fig. 9AF</figureCitation>
) (
<emphasis id="EFACEAD6FFF289029DC0FD5CFCBEFC82" bold="true" box="[705,839,749,776]" pageId="4" pageNumber="5">GROUP II</emphasis>
)........................................................................ 7
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289029FBCFCA3FA8FFCC5" blockId="4.[189,1399,714,847]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFCA3FA8FFCC5" pageId="4" pageNumber="5">
Vertex covered with punctures and never rough or granular; clypeal carina midway between clypeal margin and frontal carina (
<figureCitation id="45E32A41FFF289029D22FC84FD7DFCC5" box="[547,644,821,847]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[262,1340,366,1854]" captionTargetId="figure-1@6.[189,1398,315,1882]" captionTargetPageId="6" captionText="Fig. 1. Onitis assamensis Biswas, 1980, ♂ (AD), AIMB/Co/Sc1000001; ♀ (EF), AIMB/Co/ Sc1000009. A. Habitus, dorsal view. B. Habitus, ventral view. C. Protibia. D. Pygidium. E. Habitus, dorsal view. F. Habitus, ventral view. (Images: AIMB)" pageId="4" pageNumber="5">Figs 1A</figureCitation>
,
<figureCitation id="45E32A41FFF289029D90FC84FD3CFCC5" box="[657,709,821,847]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="4" pageNumber="5">10D</figureCitation>
) (
<emphasis id="EFACEAD6FFF289029DDFFC84FC89FCC5" bold="true" box="[734,880,821,847]" pageId="4" pageNumber="5">GROUP III</emphasis>
)................................................................ 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCFCC9FA8FFC3C" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFCC9FA8EFC18" blockId="4.[189,1399,887,950]" box="[189,1399,887,914]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFCC9FA8EFC18" box="[189,1399,887,914]" pageId="4" pageNumber="5">
7. Metasternum longitudinally grooved anteriorly (
<figureCitation id="45E32A41FFF289029C23FCC6FC71FC18" box="[802,904,887,914]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="4" pageNumber="5">Fig. 12F</figureCitation>
)............................................................... 8
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289029FBCFC2DFA8FFC3C" blockId="4.[189,1399,887,950]" box="[189,1398,924,950]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFC2DFA8FFC3C" box="[189,1398,924,950]" pageId="4" pageNumber="5">
Metasternum not grooved anteriorly (
<figureCitation id="45E32A41FFF289029DA2FC2DFD05FC3C" box="[675,764,924,950]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[262,1340,366,1854]" captionTargetId="figure-1@6.[189,1398,315,1882]" captionTargetPageId="6" captionText="Fig. 1. Onitis assamensis Biswas, 1980, ♂ (AD), AIMB/Co/Sc1000001; ♀ (EF), AIMB/Co/ Sc1000009. A. Habitus, dorsal view. B. Habitus, ventral view. C. Protibia. D. Pygidium. E. Habitus, dorsal view. F. Habitus, ventral view. (Images: AIMB)" pageId="4" pageNumber="5">Fig. 1B</figureCitation>
).................................................................................11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCFC6FFA8FFBE9" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFC6FFA8EFB96" blockId="4.[189,1399,990,1123]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFC6FFA8EFB96" pageId="4" pageNumber="5">
8. Pronotum without a smooth median longitudinal line; genae smooth, inconspicuously punctured (
<figureCitation id="45E32A41FFF289029FF2FBB0FE7DFB96" box="[243,388,1025,1052]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Fig. 14CD</figureCitation>
); male profemur not toothed......................................................................................... 9
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF289029FBCFB97FA8FFBE9" blockId="4.[189,1399,990,1123]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFB97FA8FFBE9" pageId="4" pageNumber="5">
Pronotum with a smooth median longitudinal line; genae sparingly granular (
<figureCitation id="45E32A41FFF289029B99FB97FADCFBCA" box="[1176,1317,1062,1088]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Fig. 14EF</figureCitation>
); male profemur toothed ............................................................................................................................. 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCFB3DFA8FFA9A" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCFB3DFA8FFB43" blockId="4.[189,1399,1163,1297]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCFB3DFA8FFB43" pageId="4" pageNumber="5">
9. Clypeus granular in male and rugose in female (
<figureCitation id="45E32A41FFF289029C30FB3AFC6CFB2C" box="[817,917,1163,1190]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Figs 9D</figureCitation>
,
<figureCitation id="45E32A41FFF289029CA5FB3DFC23FB2C" box="[932,986,1164,1190]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">14D</figureCitation>
), pronotum strongly and unevenly punctured ..........................
<taxonomicName id="1AD84D47FFF289029D2BFB1EFC87FB43" ID-CoL="49KK8" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" box="[554,894,1198,1225]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFF289029D2BFB1EFD46FB43" bold="true" box="[554,703,1198,1225]" italics="true" pageId="4" pageNumber="5">O. philemon</emphasis>
<bibRefCitation id="B9494B35FFF289029DC6FB1EFC87FB43" author="Fabricius J. C." box="[711,894,1199,1225]" pageId="4" pageNumber="5" refId="ref29058" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Volume 2. Impensis Bibliopolii novi academici, Kiel." type="book" year="1801">Fabricius, 1801</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029C8CFB1EFBE9FB43" box="[909,1040,1199,1225]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="4" pageNumber="5">Figs 8EH</figureCitation>
,
<figureCitation id="45E32A41FFF289029B1AFB1EFBB8FB43" box="[1051,1089,1199,1225]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">9D</figureCitation>
,
<figureCitation id="45E32A41FFF289029B4FFB1EFB79FB43" box="[1102,1152,1199,1225]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="4" pageNumber="5">10E</figureCitation>
,
<figureCitation id="45E32A41FFF289029B8DFB1EFB38FB43" box="[1164,1217,1199,1225]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">14D</figureCitation>
,
<figureCitation id="45E32A41FFF289029BCCFB1FFAFBFB43" box="[1229,1282,1198,1225]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">15D</figureCitation>
,
<figureCitation id="45E32A41FFF289029A0FFB1EFABAFB43" box="[1294,1347,1199,1225]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">16D</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EFB1EFA94FB43" box="[1359,1389,1199,1225]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." pageId="4" pageNumber="5">19</figureCitation>
)
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<paragraph id="DD6736C4FFF289029FBCFB62FA8FFA9A" blockId="4.[189,1399,1163,1297]" pageId="4" pageNumber="5">
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Clypeus rugulose in both sexes; pronotum strongly and closely punctured (
<figureCitation id="45E32A41FFF289029B58FB62FB41FB67" box="[1113,1208,1235,1261]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Figs 9C</figureCitation>
,
<figureCitation id="45E32A41FFF289029BC4FB62FB0EFB67" box="[1221,1271,1235,1261]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">14C</figureCitation>
)............... .................................................................
<taxonomicName id="1AD84D47FFF289029DD0FB47FCB9FA9B" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[721,832,1270,1297]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov.">
<emphasis id="EFACEAD6FFF289029DD0FB47FCB9FA9B" bold="true" box="[721,832,1270,1297]" italics="true" pageId="4" pageNumber="5">O. kethai</emphasis>
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<taxonomicNameLabel id="F49F57ADFFF289029C46FB46FC5BFA9B" box="[839,930,1271,1297]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="45E32A41FFF289029CB0FB47FC04FA9B" box="[945,1021,1270,1297]" captionStart="Fig" captionStartId="26.[189,232,1923,1949]" captionTargetBox="[299,1295,368,1851]" captionTargetId="figure-42@26.[191,1400,269,1864]" captionTargetPageId="26" captionText="Fig. 7. Onitis kethai sp. nov. AE. Holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023B. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head, dorsal view. D. Pygidium. E. Elytra. F. Paratype, ♀, habitus, dorsal view. (Images: AIMB)." pageId="4" pageNumber="5">Figs 7</figureCitation>
,
<figureCitation id="45E32A41FFF289029B08FB47FBADFA9A" box="[1033,1108,1270,1296]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="4" pageNumber="5">8AD</figureCitation>
,
<figureCitation id="45E32A41FFF289029B61FB47FB7DFA9A" box="[1120,1156,1270,1296]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">9C</figureCitation>
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<figureCitation id="45E32A41FFF289029B90FB47FB3DFA9A" box="[1169,1220,1270,1296]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">14C</figureCitation>
,
<figureCitation id="45E32A41FFF289029BD1FB47FAFDFA9B" box="[1232,1284,1270,1297]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">15C</figureCitation>
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<figureCitation id="45E32A41FFF289029A11FB47FABAFA9A" box="[1296,1347,1270,1296]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">16C</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EFB47FA94FA9A" box="[1359,1389,1270,1296]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="4" pageNumber="5">18</figureCitation>
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10. Lateral margin of pronotum feebly curved; front angles of pronotum blunt (
<figureCitation id="45E32A41FFF289029B51FA88FB50FAD9" box="[1104,1193,1337,1363]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Figs 9F</figureCitation>
,
<figureCitation id="45E32A41FFF289029BB2FA88FB1CFAD9" box="[1203,1253,1337,1363]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">14F</figureCitation>
); mid-femur in male with a tooth near the middle of its posterior edge, another at the extremity (
<figureCitation id="45E32A41FFF289029BBBFAEDFAD9FAFC" box="[1210,1312,1372,1398]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Fig. 16F</figureCitation>
) ......... .........................................
<taxonomicName id="1AD84D47FFF289029D1FFACEFC5AFA10" ID-CoL="49KL7" authority="Lansberge, 1875" authorityName="Lansberge, AIMB" authorityYear="1875" box="[542,931,1406,1434]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFF289029D1FFACEFD21FA13" bold="true" box="[542,728,1406,1433]" italics="true" pageId="4" pageNumber="5">O. singhalensis</emphasis>
<bibRefCitation id="B9494B35FFF289029DDEFACEFC5AFA10" author="Lansberge G." box="[735,931,1407,1434]" pageId="4" pageNumber="5" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
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(
<figureCitation id="45E32A41FFF289029CB3FACEFBD0FA13" box="[946,1065,1407,1433]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="4" pageNumber="5">Figs 8IL</figureCitation>
,
<figureCitation id="45E32A41FFF289029B34FACEFBADFA13" box="[1077,1108,1407,1433]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">9F</figureCitation>
,
<figureCitation id="45E32A41FFF289029B61FACEFB6FFA13" box="[1120,1174,1407,1433]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="4" pageNumber="5">12A</figureCitation>
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<figureCitation id="45E32A41FFF289029BA0FACEFB29FA13" box="[1185,1232,1407,1433]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">14F</figureCitation>
,
<figureCitation id="45E32A41FFF289029BDAFACEFAF0FA10" box="[1243,1289,1407,1434]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">15F</figureCitation>
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<figureCitation id="45E32A41FFF289029A14FACEFABDFA13" box="[1301,1348,1407,1433]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">16F</figureCitation>
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<figureCitation id="45E32A41FFF289029A4EFACEFA94FA13" box="[1359,1389,1407,1433]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." pageId="4" pageNumber="5">19</figureCitation>
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<paragraph id="DD6736C4FFF289029FBCFA15FA8FF9AD" blockId="4.[189,1399,1337,1576]" pageId="4" pageNumber="5">
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Lateral margin of pronotum strongly curved; front angles of pronotum sharp (
<figureCitation id="45E32A41FFF289029B91FA12FB09FA37" box="[1168,1264,1443,1469]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Figs 9E</figureCitation>
,
<figureCitation id="45E32A41FFF289029BFFFA12FACBFA37" box="[1278,1330,1443,1469]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">14E</figureCitation>
); mid femur in male without a tooth near the middle of the posterior edge, the extremity with a sharp tooth on the lower edge and a blunt one above (
<figureCitation id="45E32A41FFF289029DD4FA5BFCC5F98E" box="[725,828,1514,1540]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">Fig. 16E</figureCitation>
) ........................................................................... .......................................................
<taxonomicName id="1AD84D47FFF289029D86F9BCFCF5F9AD" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[647,780,1548,1575]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov.">
<emphasis id="EFACEAD6FFF289029D86F9BCFCF5F9AD" bold="true" box="[647,780,1548,1575]" italics="true" pageId="4" pageNumber="5">O. visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFF289029C12F9BCFC97F9AD" box="[787,878,1549,1575]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="45E32A41FFF289029C7CF9BCFBF8F9AD" box="[893,1025,1549,1575]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[295,1242,313,1806]" captionTargetId="figure-50@28.[189,1398,262,1824]" captionTargetPageId="28" captionText="Fig. 8. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiskethai sp. nov., NIM/NBAIR/ COL/ONIT/H-271023B. EH. O. philemon Fabricius,1801,AIMB/Co/Sc1000317. IL. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. MP. O. visthara sp. nov., NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB)." pageId="4" pageNumber="5">Figs 8MP</figureCitation>
,
<figureCitation id="45E32A41FFF289029B0DF9BCFBD6F9AD" box="[1036,1071,1549,1575]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">9E</figureCitation>
,
<figureCitation id="45E32A41FFF289029B3AF9BCFB73F9AD" box="[1083,1162,1549,1575]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="4" pageNumber="5">12EF</figureCitation>
,
<figureCitation id="45E32A41FFF289029B97F9BCFB3EF9AD" box="[1174,1223,1549,1575]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">14E</figureCitation>
,
<figureCitation id="45E32A41FFF289029BD5F9BCFAFFF9A2" box="[1236,1286,1549,1576]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">15E</figureCitation>
,
<figureCitation id="45E32A41FFF289029A10F9BCFABAF9AD" box="[1297,1347,1549,1575]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="4" pageNumber="5">16E</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EF9BCFA97F9AD" box="[1359,1390,1549,1575]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." pageId="4" pageNumber="5">20</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCF9E1FA8FF95E" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCF9E1FA8FF907" blockId="4.[189,1398,1615,1749]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCF9E1FA8FF907" pageId="4" pageNumber="5">
11. Frontal carina narrowly interrupted; pygidium smooth with minute punctures.................................. .............................................................................
<taxonomicName id="1AD84D47FFF289029C2BF9C2FB9BF907" ID-CoL="49KMB" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[810,1122,1650,1677]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFF289029C2BF9C2FC66F907" bold="true" box="[810,927,1651,1677]" italics="true" pageId="4" pageNumber="5">O. virens</emphasis>
<bibRefCitation id="B9494B35FFF289029C9FF9C2FB9BF907" author="Lansberge G." box="[926,1122,1650,1677]" pageId="4" pageNumber="5" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029B70F9C2FB18F907" box="[1137,1249,1651,1677]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="4" pageNumber="5">Figs 12D</figureCitation>
,
<figureCitation id="45E32A41FFF289029BEFF9C2FABDF907" box="[1262,1348,1651,1677]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. EH. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." pageId="4" pageNumber="5">13EH</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EF9C2FA97F907" box="[1359,1390,1651,1677]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." pageId="4" pageNumber="5">20</figureCitation>
)
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</paragraph>
<paragraph id="DD6736C4FFF289029FBCF926FA8FF95E" blockId="4.[189,1398,1615,1749]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCF926FA8FF95E" pageId="4" pageNumber="5">
Frontal carina widely interrupted; pygidium opaque with imperceptible punctures........................... ............................................................................
<taxonomicName id="1AD84D47FFF289029C22F90BFB99F95E" ID-CoL="49KLJ" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[803,1120,1722,1749]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFF289029C22F90BFC3FF95F" bold="true" box="[803,966,1722,1749]" italics="true" pageId="4" pageNumber="5">O. subopacus</emphasis>
<bibRefCitation id="B9494B35FFF289029CCDF90BFB99F95E" author="Arrow G. J." box="[972,1120,1722,1748]" pageId="4" pageNumber="5" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029B6EF90BFB27F95E" box="[1135,1246,1722,1748]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="4" pageNumber="5">Figs 12C</figureCitation>
,
<figureCitation id="45E32A41FFF289029BEBF90BFABAF95E" box="[1258,1347,1722,1748]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. EH. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." pageId="4" pageNumber="5">13AD</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EF90BFA97F95E" box="[1359,1390,1722,1748]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." pageId="4" pageNumber="5">20</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF289029FBCF94CFA8FF80B" pageId="4" pageNumber="5">
<paragraph id="DD6736C4FFF289029FBCF94CFA8FF8B0" blockId="4.[189,1399,1789,1922]" pageId="4" pageNumber="5">
<keyLead id="662995F1FFF289029FBCF94CFA8FF8B0" pageId="4" pageNumber="5">
12. Pronotal anterior angles not extended, blunt; male protibia with 34 teeth on the inferomedial ridge ...........................................................................
<taxonomicName id="1AD84D47FFF289029C1AF891FB81F8B0" ID-CoL="49KJD" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[795,1144,1824,1850]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFF289029C1AF891FC50F8B0" bold="true" box="[795,937,1824,1850]" italics="true" pageId="4" pageNumber="5">O. naviauxi</emphasis>
<bibRefCitation id="B9494B35FFF289029CB0F891FB81F8B0" author="Cambefort Y." box="[945,1144,1824,1850]" pageId="4" pageNumber="5" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029B86F891FAFBF8B0" box="[1159,1282,1824,1850]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="4" pageNumber="5">Figs 4EF</figureCitation>
,
<figureCitation id="45E32A41FFF289029A0FF891FABAF8B0" box="[1294,1347,1824,1850]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="4" pageNumber="5">10D</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EF891FA94F8B0" box="[1359,1389,1824,1850]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." pageId="4" pageNumber="5">19</figureCitation>
)
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</paragraph>
<paragraph id="DD6736C4FFF289029FBCF8F4FA8FF80B" blockId="4.[189,1399,1789,1922]" pageId="4" pageNumber="5">
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Pronotal anterior angles slightly extended, sharp; male protibia with 67 teeth on the inferomedial ridge (
<figureCitation id="45E32A41FFF289029E36F8D6FE76F80B" box="[311,399,1895,1921]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[262,1340,366,1854]" captionTargetId="figure-1@6.[189,1398,315,1882]" captionTargetPageId="6" captionText="Fig. 1. Onitis assamensis Biswas, 1980, ♂ (AD), AIMB/Co/Sc1000001; ♀ (EF), AIMB/Co/ Sc1000009. A. Habitus, dorsal view. B. Habitus, ventral view. C. Protibia. D. Pygidium. E. Habitus, dorsal view. F. Habitus, ventral view. (Images: AIMB)" pageId="4" pageNumber="5">Fig. 1C</figureCitation>
) ........................................................
<taxonomicName id="1AD84D47FFF289029C41F8D6FB68F80B" ID-CoL="6SMW9" authority="Biswas, 1980" authorityName="Biswas" authorityYear="1980" box="[832,1169,1895,1921]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF289029C41F8D6FC12F80B" bold="true" box="[832,1003,1895,1921]" italics="true" pageId="4" pageNumber="5">O. assamensis</emphasis>
<bibRefCitation id="B9494B35FFF289029CF2F8D6FB68F80B" author="Biswas S." box="[1011,1169,1895,1921]" pageId="4" pageNumber="5" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF289029BA1F8D6FB12F80B" box="[1184,1259,1895,1921]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[262,1340,366,1854]" captionTargetId="figure-1@6.[189,1398,315,1882]" captionTargetPageId="6" captionText="Fig. 1. Onitis assamensis Biswas, 1980, ♂ (AD), AIMB/Co/Sc1000001; ♀ (EF), AIMB/Co/ Sc1000009. A. Habitus, dorsal view. B. Habitus, ventral view. C. Protibia. D. Pygidium. E. Habitus, dorsal view. F. Habitus, ventral view. (Images: AIMB)" pageId="4" pageNumber="5">Figs 1</figureCitation>
,
<figureCitation id="45E32A41FFF289029BF8F8D6FABDF80B" box="[1273,1348,1895,1921]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="4" pageNumber="5">4AD</figureCitation>
,
<figureCitation id="45E32A41FFF289029A4EF8D6FA94F80B" box="[1359,1389,1895,1921]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." pageId="4" pageNumber="5">17</figureCitation>
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<paragraph id="DD6736C4FFF289029FBCF81BFA8FF84E" blockId="4.[189,1398,1962,2025]" box="[189,1398,1962,1988]" pageId="4" pageNumber="5">
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13. Pygidium covered with short or long hair (
<emphasis id="EFACEAD6FFF289029DDEF81BFC83F84E" bold="true" box="[735,890,1962,1988]" pageId="4" pageNumber="5">GROUP VII</emphasis>
)............................................................... 14
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</paragraph>
<paragraph id="DD6736C4FFF289039FBCF87EFA8FFECC" blockId="4.[189,1398,1962,2025]" lastBlockId="5.[189,1398,265,362]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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Pygidium not covered with hair or setae ......................................................................................... 17
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14. Clypeus elliptical; head not tuberculate; elytral intervals slightly carinate in the middle................... .....................................................................................
<taxonomicName id="1AD84D47FFF389039C63FE9DFB9AFECC" ID-CoL="49KG7" authority="Felsche, 1907" authorityName="Felsche" authorityYear="1907" box="[866,1123,300,326]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="feae">
<emphasis id="EFACEAD6FFF389039C63FE9DFC41FECC" bold="true" box="[866,952,300,326]" italics="true" pageId="5" pageNumber="6">O. feae</emphasis>
<bibRefCitation id="B9494B35FFF389039CC1FE9DFB9AFECC" author="Felsche C." box="[960,1123,300,326]" pageId="5" pageNumber="6" pagination="273 - 296" refId="ref29092" refString="Felsche C. 1907. Coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift: 273 - 296." type="book chapter" year="1907">Felsche, 1907</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039B73FE9DFB1AFECC" box="[1138,1251,300,326]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="5" pageNumber="6">Figs 10A</figureCitation>
,
<figureCitation id="45E32A41FFF389039BEEFE9DFABDFECC" box="[1263,1348,300,326]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. EH. O. feae Felsche, 1907, AIMB/Co/Sc1000188. IL. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." pageId="5" pageNumber="6">11EH</figureCitation>
,
<figureCitation id="45E32A41FFF389039A4EFE9DFA94FECC" box="[1359,1389,300,326]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="5" pageNumber="6">18</figureCitation>
)
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</paragraph>
<paragraph id="DD6736C4FFF389039FBCFEE1FA8FFEE0" blockId="5.[189,1398,265,362]" box="[189,1398,335,362]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFEE1FA8FFEE0" box="[189,1398,335,362]" pageId="5" pageNumber="6"> Clypeus truncate or excised; head tuberculate; elytral intervals convex......................................... 15</keyLead>
</paragraph>
<keyStep id="662C2E61FFF389039FBCFE20FA8FFE7B" pageId="5" pageNumber="6">
<paragraph id="DD6736C4FFF389039FBCFE20FA8FFE44" blockId="5.[189,1398,401,498]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFE20FA8FFE44" pageId="5" pageNumber="6">
15. Clypeal margin deeply excised with an obtuse angle; frontal carina with a small gap ....................... ..........................................................................................
<taxonomicName id="1AD84D47FFF389039C86FE05FB2DFE45" ID-CoL="74MW8" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[903,1236,435,463]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFF389039C86FE05FBF3FE44" bold="true" box="[903,1034,435,462]" italics="true" pageId="5" pageNumber="6">O. brahma</emphasis>
<bibRefCitation id="B9494B35FFF389039B11FE05FB2DFE45" author="Lansberge G." box="[1040,1236,436,463]" pageId="5" pageNumber="6" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
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(
<figureCitation id="45E32A41FFF389039BE2FE05FABBFE45" box="[1251,1346,436,463]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="5" pageNumber="6">Figs 5B</figureCitation>
,
<figureCitation id="45E32A41FFF389039A4EFE05FA94FE44" box="[1359,1389,436,462]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." pageId="5" pageNumber="6">17</figureCitation>
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</paragraph>
<paragraph id="DD6736C4FFF389039FBCFE69FA8FFE7B" blockId="5.[189,1398,401,498]" box="[189,1398,471,498]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFE69FA8FFE7B" box="[189,1398,471,498]" pageId="5" pageNumber="6"> Clypeal margin not deeply excised; frontal carina entire ................................................................ 16</keyLead>
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<keyStep id="662C2E61FFF389039FBCFDA8FA8FFD16" pageId="5" pageNumber="6">
<paragraph id="DD6736C4FFF389039FBCFDA8FA8FFDDC" blockId="5.[189,1398,537,669]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFDA8FA8FFDDC" pageId="5" pageNumber="6">
16. Hair on the pygidium short; elytral interval angularly elevated along middle with fine close punctures...................................................................................
<taxonomicName id="1AD84D47FFF389039CBEFD8DFB2BFDDD" ID-CoL="49KFD" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[959,1234,572,599]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFF389039CBEFD8DFBBDFDDC" bold="true" box="[959,1092,572,598]" italics="true" pageId="5" pageNumber="6">O. crassus</emphasis>
Sharp, 1875
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039BE0FD8DFABAFDDD" box="[1249,1347,572,599]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="5" pageNumber="6">Figs 5D</figureCitation>
,
<figureCitation id="45E32A41FFF389039A4EFD8DFA94FDDC" box="[1359,1389,572,598]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="5" pageNumber="6">18</figureCitation>
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</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFDD1FA8FFD16" blockId="5.[189,1398,537,669]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFDD1FA8FFD16" pageId="5" pageNumber="6">
Hair on the pygidium long; elytral interval flat with feeble punctures................................................ ....................................................................
<taxonomicName id="1AD84D47FFF389039DE5FD33FB89FD16" ID-CoL="6SN86" authority="Redtenbacher, 1848" authorityName="Redtenbacher" authorityYear="1848" box="[740,1136,642,669]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFF389039DE5FD33FC86FD17" bold="true" box="[740,895,642,669]" italics="true" pageId="5" pageNumber="6">O. castaneus</emphasis>
Redtenbacher, 1848
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039B7EFD33FB26FD17" box="[1151,1247,642,669]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="5" pageNumber="6">Figs 5C</figureCitation>
,
<figureCitation id="45E32A41FFF389039BEAFD33FABAFD16" box="[1259,1347,642,668]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. EH. O. feae Felsche, 1907, AIMB/Co/Sc1000188. IL. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." pageId="5" pageNumber="6">11AD</figureCitation>
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<figureCitation id="45E32A41FFF389039A4EFD33FA94FD16" box="[1359,1389,642,668]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." pageId="5" pageNumber="6">17</figureCitation>
)
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</paragraph>
</keyStep>
<keyStep id="662C2E61FFF389039FBCFD75FA8FFC8B" pageId="5" pageNumber="6">
<paragraph id="DD6736C4FFF389039FBCFD75FA8FFD54" blockId="5.[189,1398,708,769]" box="[189,1398,708,734]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFD75FA8FFD54" box="[189,1398,708,734]" pageId="5" pageNumber="6">17. Clypeal carina present ..................................................................................................................... 18</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFD56FA8FFC8B" blockId="5.[189,1398,708,769]" box="[189,1398,743,769]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFD56FA8FFC8B" box="[189,1398,743,769]" pageId="5" pageNumber="6"> Clypeal carina absent....................................................................................................................... 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="662C2E61FFF389039FBCFC99FA8FFC45" pageId="5" pageNumber="6">
<paragraph id="DD6736C4FFF389039FBCFC99FA8FFCEC" blockId="5.[189,1398,808,976]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFC99FA8FFCEC" pageId="5" pageNumber="6">
18. First, third and fifth inter-striae of the elytra not much convex; elytra dark, metasternal shield smooth behind, with a median groove (
<emphasis id="EFACEAD6FFF389039D58FCFAFD24FCEC" bold="true" box="[601,733,843,870]" pageId="5" pageNumber="6">GROUP V</emphasis>
)............
<taxonomicName id="1AD84D47FFF389039C44FCFAFB89FCEC" ID-CoL="74N7T" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[837,1136,843,870]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lama">
<emphasis id="EFACEAD6FFF389039C44FCFAFC5CFCEC" bold="true" box="[837,933,843,870]" italics="true" pageId="5" pageNumber="6">O. lama</emphasis>
<bibRefCitation id="B9494B35FFF389039CADFCFAFB89FCEC" author="Lansberge G." box="[940,1136,843,870]" pageId="5" pageNumber="6" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
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(
<figureCitation id="45E32A41FFF389039B7EFCFAFB17FCEC" box="[1151,1262,843,870]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="5" pageNumber="6">Figs 10C</figureCitation>
,
<figureCitation id="45E32A41FFF389039BFBFCFDFABAFCEC" box="[1274,1347,844,870]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. EH. O. feae Felsche, 1907, AIMB/Co/Sc1000188. IL. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." pageId="5" pageNumber="6">11IL</figureCitation>
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<figureCitation id="45E32A41FFF389039A4EFCFDFA94FCEC" box="[1359,1389,844,870]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." pageId="5" pageNumber="6">19</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFCDEFA8FFC45" blockId="5.[189,1398,808,976]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFCDEFA8FFC45" pageId="5" pageNumber="6">
First, third, and fifth inter-striae distinctly convex; elytra yellow with some intervals green; metasternal shield punctured behind, without a median groove (
<emphasis id="EFACEAD6FFF389039CD6FC23FB85FC26" bold="true" box="[983,1148,914,940]" pageId="5" pageNumber="6">GROUP XIV</emphasis>
) ............................... .......................................................................................
<taxonomicName id="1AD84D47FFF389039C70FC04FB3CFC45" ID-CoL="49KGS" authority="(Pallas, 1771)" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[881,1221,949,976]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFF389039C70FC04FBE2FC5A" bold="true" box="[881,1051,949,976]" italics="true" pageId="5" pageNumber="6">O. humerosus</emphasis>
(
<bibRefCitation id="B9494B35FFF389039B2BFC04FB45FC45" author="Pallas P. S." box="[1066,1212,949,975]" pageId="5" pageNumber="6" refId="ref30468" refString="Pallas P. S. 1771. Reise durch verschiedene Provinzen des Russischen Reiches in den Jahren 1768 - 1774 I: 1 - 504 (453 - 504). Petersburg Akademische Buchhandlung, St Petersburg." type="book" year="1771">Pallas, 1771</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039BD5FC04FABBFC45" box="[1236,1346,949,975]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." pageId="5" pageNumber="6">Figs 10B</figureCitation>
,
<figureCitation id="45E32A41FFF389039A4EFC04FA94FC45" box="[1359,1389,949,975]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." pageId="5" pageNumber="6">18</figureCitation>
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</paragraph>
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<keyStep id="662C2E61FFF389039FBCFC46FA8FFBF0" pageId="5" pageNumber="6">
<paragraph id="DD6736C4FFF389039FBCFC46FA8FFBBE" blockId="5.[189,1398,1014,1147]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFC46FA8FFBBE" pageId="5" pageNumber="6">
19. Pronotum covered with close granules (
<emphasis id="EFACEAD6FFF389039DBCFC47FCB7FB9B" bold="true" box="[701,846,1014,1041]" pageId="5" pageNumber="6">GROUP IX</emphasis>
)......................................................................... ............................................................................
<taxonomicName id="1AD84D47FFF389039C1EFBABFB91FBBE" ID-CoL="74MVW" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[799,1128,1049,1076]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bordati">
<emphasis id="EFACEAD6FFF389039C1EFBABFC58FBBE" bold="true" box="[799,929,1049,1076]" italics="true" pageId="5" pageNumber="6">O. bordati</emphasis>
<bibRefCitation id="B9494B35FFF389039CA0FBABFB91FBBE" author="Cambefort Y." box="[929,1128,1050,1076]" pageId="5" pageNumber="6" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039B76FBABFB3AFBBE" box="[1143,1219,1050,1076]" captionStart="Fig" captionStartId="11.[189,232,1953,1979]" captionTargetBox="[280,1294,921,1858]" captionTargetId="figure-246@11.[189,1398,900,1917]" captionTargetPageId="11" captionText="Fig. 3. Onitis bordati Cambefort, 1988, ♀, AIMB/Co/Sc1000018. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head, dorsal view. D. Pygidium. (Images: AIMB)." pageId="5" pageNumber="6">Figs 3</figureCitation>
,
<figureCitation id="45E32A41FFF389039BD1FBABFAE8FBBE" box="[1232,1297,1050,1076]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="5" pageNumber="6">4GJ</figureCitation>
,
<figureCitation id="45E32A41FFF389039A1CFBABFABAFBBF" box="[1309,1347,1050,1077]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="5" pageNumber="6">5A</figureCitation>
,
<figureCitation id="45E32A41FFF389039A4EFBABFA94FBBE" box="[1359,1389,1050,1076]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." pageId="5" pageNumber="6">17</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFB8FFA8FFBF0" blockId="5.[189,1398,1014,1147]" pageId="5" pageNumber="6">
<keyLead id="662995F1FFF389039FBCFB8FFA8FFBF0" pageId="5" pageNumber="6">
Pronotum covered with strong punctures (
<emphasis id="EFACEAD6FFF389039DD8FB8CFC6CFBDD" bold="true" box="[729,917,1085,1111]" pageId="5" pageNumber="6">GROUP XVIII</emphasis>
)............................................................... ..........................................................................................
<taxonomicName id="1AD84D47FFF389039C88FBD1FB96FBF0" ID-CoL="49KL8" authority="Gillet, 1911" authorityName="Gillet" authorityYear="1911" box="[905,1135,1120,1146]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="siva">
<emphasis id="EFACEAD6FFF389039C88FBD1FC25FBF0" bold="true" box="[905,988,1120,1146]" italics="true" pageId="5" pageNumber="6">O. siva</emphasis>
<bibRefCitation id="B9494B35FFF389039CE2FBD1FB96FBF0" author="Gillet J. J. E." box="[995,1135,1120,1146]" pageId="5" pageNumber="6" pagination="313 - 314" refId="ref29108" refString="Gillet J. J. E. 1911. Coprides nouveaux de la region orientale (etc.). Annales de la Societe entomologique de Belgique LV: 313 - 314." type="book chapter" year="1911">Gillet, 1911</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45E32A41FFF389039B7FFBD1FAFDFBF0" box="[1150,1284,1120,1146]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="5" pageNumber="6">Figs 4KN</figureCitation>
,
<figureCitation id="45E32A41FFF389039A11FBD1FABBFBF0" box="[1296,1346,1120,1146]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." pageId="5" pageNumber="6">12B</figureCitation>
,
<figureCitation id="45E32A41FFF389039A4EFBD1FA97FBF0" box="[1359,1390,1120,1146]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." pageId="5" pageNumber="6">20</figureCitation>
)
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@ -0,0 +1,379 @@
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<paragraph id="DD6736C4FFF989099D68F994FC32F9B5" blockId="15.[617,971,1573,1635]" box="[617,971,1573,1599]" pageId="15" pageNumber="16">
<heading id="862F81A8FFF989099D68F994FC32F9B5" box="[617,971,1573,1599]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
<taxonomicName id="1AD84D47FFF989099D68F994FC32F9B5" ID-CoL="6SN86" authority="Kollar, 1844" authorityName="Kollar" authorityYear="1844" box="[617,971,1573,1599]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFF989099D68F994FCC9F9B5" bold="true" box="[617,816,1573,1599]" italics="true" pageId="15" pageNumber="16">Onitis castaneus</emphasis>
<bibRefCitation id="B9494B35FFF989099C39F994FC32F9B5" author="Kollar V. &amp; Redtenbacher L." box="[824,971,1573,1599]" pageId="15" pageNumber="16" pagination="395 - 564" refId="ref29778" refString="Kollar V. &amp; Redtenbacher L. 1844. Aufzahlung und Beschreibung der von Freiherrn Carl v. Hugel auf seiner Reise durch Kaschmir und das Himaleyagebirge gesammelten Insecten. Mit 28 Steindruck- Tafeln. In Hugel C. A. A. von (ed.), Kaschmir und das Reich der Siek: 395 - 564. Hallberger, Stuttgart." type="book chapter" year="1844">Kollar, 1844</bibRefCitation>
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<paragraph id="DD6736C4FFF989099DA5F9F9FC76F9E8" blockId="15.[617,971,1573,1635]" box="[676,911,1608,1635]" pageId="15" pageNumber="16">
<figureCitation id="45E32A41FFF989099DA5F9F9FCFBF9E9" box="[676,770,1608,1635]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="15" pageNumber="16">Figs 5C</figureCitation>
,
<figureCitation id="45E32A41FFF989099C0FF9F9FC9FF9E8" box="[782,870,1608,1634]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. EH. O. feae Felsche, 1907, AIMB/Co/Sc1000188. IL. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." pageId="15" pageNumber="16">11AD</figureCitation>
,
<figureCitation id="45E32A41FFF989099C73F9F9FC76F9E8" box="[882,911,1608,1634]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." pageId="15" pageNumber="16">17</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFF989099FBCF93BFCA4F929" blockId="15.[189,861,1673,1700]" box="[189,861,1673,1700]" pageId="15" pageNumber="16">
<emphasis id="EFACEAD6FFF989099FBCF93BFE79F92E" box="[189,384,1674,1700]" italics="true" pageId="15" pageNumber="16">Onitiscastaneus</emphasis>
<bibRefCitation id="B9494B35FFF989099E86F938FDAFF92E" author="Kollar V. &amp; Redtenbacher L." box="[391,598,1673,1700]" pageId="15" pageNumber="16" pagination="395 - 564" refId="ref29778" refString="Kollar V. &amp; Redtenbacher L. 1844. Aufzahlung und Beschreibung der von Freiherrn Carl v. Hugel auf seiner Reise durch Kaschmir und das Himaleyagebirge gesammelten Insecten. Mit 28 Steindruck- Tafeln. In Hugel C. A. A. von (ed.), Kaschmir und das Reich der Siek: 395 - 564. Hallberger, Stuttgart." type="book chapter" year="1844">Kollar, 1844: 517</bibRefCitation>
(original description).
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<taxonomicName id="1AD84D47FFF989099FBCF97AFE87F96F" ID-CoL="6SN86" authorityName="Kollar" authorityYear="1844" box="[189,382,1739,1765]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFF989099FBCF97AFE87F96F" box="[189,382,1739,1765]" italics="true" pageId="15" pageNumber="16">Onitis castaneus</emphasis>
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<bibRefCitation id="B9494B35FFF989099E99F97AFD75F96F" author="Lansberge G." box="[408,652,1739,1766]" pageId="15" pageNumber="16" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge 1875: 144</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFF989099C33F97AFC00F96F" author="Arrow G. J." box="[818,1017,1739,1765]" page="398" pageId="15" pageNumber="16" year="1931">
<bibRefCitation id="B9494B35FFF989099C33F97AFC00F96F" author="Arrow G. J." box="[818,1017,1739,1765]" pageId="15" pageNumber="16" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 398</bibRefCitation>
</treatmentCitation>
(keys, description). —
<treatmentCitation id="5C7910D5FFF989099A09F97AFEB6F882" author="Balthasar V." page="92" pageId="15" pageNumber="16" year="1935">
<bibRefCitation id="B9494B35FFF989099A09F97AFEB6F882" author="Balthasar V." pageId="15" pageNumber="16" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 92</bibRefCitation>
</treatmentCitation>
(monograph); 1963: 46 (monograph). —
<treatmentCitation id="5C7910D5FFF989099C28F95FFC0EF882" author="Janssens A." box="[809,1015,1774,1800]" page="74" pageId="15" pageNumber="16" year="1937">
<bibRefCitation id="B9494B35FFF989099C28F95FFC0EF882" author="Janssens A." box="[809,1015,1774,1800]" pageId="15" pageNumber="16" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 74</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFF989099B9AF95FFA8FF882" author="Chandra K." box="[1179,1398,1774,1800]" page="360" pageId="15" pageNumber="16" year="2000">
<bibRefCitation id="B9494B35FFF989099B9AF95FFA8FF882" author="Chandra K." box="[1179,1398,1774,1800]" pageId="15" pageNumber="16" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000: 360</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFF989099EC3F8A0FD78F8A1" author="Mittal I. C." box="[450,641,1809,1836]" page="46" pageId="15" pageNumber="16" year="2005">
<bibRefCitation id="B9494B35FFF989099EC3F8A0FD78F8A1" author="Mittal I. C." box="[450,641,1809,1836]" pageId="15" pageNumber="16" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005: 46</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFF989099C00F8A0FBB2F8A6" author="Bezdek A. &amp; Krell F. - T." box="[769,1099,1809,1836]" page="159" pageId="15" pageNumber="16" year="2006">
<bibRefCitation id="B9494B35FFF989099C00F8A0FBB2F8A6" author="Bezdek A. &amp; Krell F. - T." box="[769,1099,1809,1836]" pageId="15" pageNumber="16" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(describing author, year correction). —
<treatmentCitation id="5C7910D5FFF989099EA7F885FCE4F8C4" author="Chandra K. &amp; Ahirwar S. C." box="[422,797,1844,1870]" page="287" pageId="15" pageNumber="16" year="2007">
<bibRefCitation id="B9494B35FFF989099EA7F885FCE4F8C4" author="Chandra K. &amp; Ahirwar S. C." box="[422,797,1844,1870]" pageId="15" pageNumber="16" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">Chandra &amp; Ahirwar 2007: 287</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFF989099CFEF885FB2BF8C4" author="Sewak R." box="[1023,1234,1844,1870]" page="61" pageId="15" pageNumber="16" year="2009">
<bibRefCitation id="B9494B35FFF989099CFEF885FB2BF8C4" author="Sewak R." box="[1023,1234,1844,1870]" pageId="15" pageNumber="16" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">Sewak 2009b: 61</bibRefCitation>
</treatmentCitation>
(distribution, diagnosis). — Singh
<emphasis id="EFACEAD6FFF989099EF4F8E9FDCFF8FB" box="[501,566,1879,1905]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2010: 1394 (distribution). —
<treatmentCitation id="5C7910D5FFF989099CB6F8E9FB1FF8F8" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[951,1254,1879,1906]" page="1037" pageId="15" pageNumber="16" year="2015">
<bibRefCitation id="B9494B35FFF989099CB6F8E9FB1FF8F8" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[951,1254,1879,1906]" pageId="15" pageNumber="16" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFF989099B0DF8E9FBB4F8FB" box="[1036,1101,1879,1905]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(checklist). —
<treatmentCitation id="5C7910D5FFF989099E10F8CAFE10F81F" author="Bezdek A." box="[273,489,1915,1941]" page="179" pageId="15" pageNumber="16" year="2016">
<bibRefCitation id="B9494B35FFF989099E10F8CAFE10F81F" author="Bezdek A." box="[273,489,1915,1941]" pageId="15" pageNumber="16" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(describing author, year correction, distribution). —
<treatmentCitation id="5C7910D5FFF989099B65F8CAFA8FF81F" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." box="[1124,1398,1915,1941]" page="238" pageId="15" pageNumber="16" year="2020">
<bibRefCitation id="B9494B35FFF989099B65F8CAFA8FF81F" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." box="[1124,1398,1915,1941]" pageId="15" pageNumber="16" pagination="221 - 275" refId="ref29141" refString="Ghosh J., Das, P., Ghosh S. K., Bhunia D., Kushwaha R. K., Gupta D., &amp; Chandra K. 2020. Insecta: Coleoptera. Zoological Survey of India, Fauna of Haryana, State Fauna Series 24: 221 - 275." type="book chapter" year="2020">
Ghosh
<emphasis id="EFACEAD6FFF989099BB8F8CDFB0FF81F" box="[1209,1270,1915,1941]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2020: 238
</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFF989099EB9F82FFD4AF832" author="Schoolmeesters P." box="[440,691,1950,1976]" pageId="15" pageNumber="16" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE689169FBCFEB9FE4AFD88" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="DD6736C4FFE689169FBCFEB9FE3FFEA8" blockId="16.[189,454,264,290]" box="[189,454,264,290]" pageId="16" pageNumber="17">
<heading id="862F81A8FFE689169FBCFEB9FE3FFEA8" bold="true" box="[189,454,264,290]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="EFACEAD6FFE689169FBCFEB9FE3FFEA8" bold="true" box="[189,454,264,290]" pageId="16" pageNumber="17">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE689169FBCFE86FE4AFD88" blockId="16.[189,1399,311,514]" pageId="16" pageNumber="17">
<emphasis id="EFACEAD6FFE689169FBCFE86FE79FEDB" box="[189,384,311,337]" italics="true" pageId="16" pageNumber="17">Onitiscastaneus</emphasis>
is similar to
<taxonomicName id="1AD84D47FFE689169D1AFE86FD62FEDB" authorityName="Lansberge" authorityYear="1875" box="[539,667,311,337]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFE689169D1AFE86FD62FEDB" box="[539,667,311,337]" italics="true" pageId="16" pageNumber="17">O. brahma</emphasis>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFE689169DD4FE86FCA8FEDB" authorityName="Sharp" authorityYear="1875" box="[725,849,311,337]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFE689169DD4FE86FCA8FEDB" box="[725,849,311,337]" italics="true" pageId="16" pageNumber="17">O. crassus</emphasis>
</taxonomicName>
in having a hairy pygidium and it differs from them in having tubercle on the clypeo-frontal carina while the tubercle is placed behind the clypeo-frontal carina in both the latter species. It differs from
<taxonomicName id="1AD84D47FFE689169C3EFECFFC47FE1D" authorityName="Lansberge" authorityYear="1875" box="[831,958,381,408]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFE689169C3EFECFFC47FE1D" box="[831,958,381,408]" italics="true" pageId="16" pageNumber="17">O. brahma</emphasis>
</taxonomicName>
in the frontal carina being entire; the clypeal margin being straight while the carina is interrupted and the clypeal margin is slightly excised in
<taxonomicName id="1AD84D47FFE689169FBCFE75FEC4FE54" authorityName="Lansberge" authorityYear="1875" box="[189,317,452,478]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFE689169FBCFE75FEC4FE54" box="[189,317,452,478]" italics="true" pageId="16" pageNumber="17">O. brahma</emphasis>
</taxonomicName>
.
<emphasis id="EFACEAD6FFE689169E4CFE75FDE8FE54" box="[333,529,452,478]" italics="true" pageId="16" pageNumber="17">Onitiscastaneus</emphasis>
differs from
<taxonomicName id="1AD84D47FFE689169DACFE75FCD3FE54" authorityName="Sharp" authorityYear="1875" box="[685,810,452,478]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFE689169DACFE75FCD3FE54" box="[685,810,452,478]" italics="true" pageId="16" pageNumber="17">O. crassus</emphasis>
</taxonomicName>
in having longer hairs on the pygidium while the latter has short setae.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE689169FBCFD99FE2CFC23" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="DD6736C4FFE689169FBCFD99FE53FDC8" blockId="16.[189,426,552,578]" box="[189,426,552,578]" pageId="16" pageNumber="17">
<heading id="862F81A8FFE689169FBCFD99FE53FDC8" bold="true" box="[189,426,552,578]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="EFACEAD6FFE689169FBCFD99FE53FDC8" bold="true" box="[189,426,552,578]" pageId="16" pageNumber="17">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE689169FEBFDE6FE02FDFB" blockId="16.[189,756,599,661]" box="[234,507,599,625]" pageId="16" pageNumber="17">
<emphasis id="EFACEAD6FFE689169FEBFDE6FEB6FDFB" bold="true" box="[234,335,599,625]" pageId="16" pageNumber="17">
<typeStatus id="02638866FFE689169FEBFDE6FEB6FDFB" box="[234,335,599,625]" pageId="16" pageNumber="17" type="syntype">Syntype</typeStatus>
</emphasis>
(
<specimenCount id="CBDEFD4DFFE689169E61FDE6FE0BFDFB" box="[352,498,599,625]" count="3" pageId="16" pageNumber="17" type="generic">3 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFE689169FBCFDCBFD0DFD1F" blockId="16.[189,756,599,661]" box="[189,756,634,661]" pageId="16" pageNumber="17">
<materialsCitation id="6DB03C99FFE689169FBCFDCBFD16FD1F" box="[189,751,634,661]" collectionCode="NHMW" country="India" location="INDIA" pageId="16" pageNumber="17" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<collectingCountry id="A5CF7654FFE689169FBCFDCBFEF7FD1E" box="[189,270,634,660]" name="India" pageId="16" pageNumber="17">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFE689169E26FDCAFEC6FD1F" box="[295,319,635,661]" count="1" pageId="16" pageNumber="17" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFE689169E4DFDCBFE76FD1F" box="[332,399,634,661]" count="2" pageId="16" pageNumber="17" type="female">2 ♀♀</specimenCount>
; “Kaschmir, Hügel”;
<collectionCode id="BBC9AE01FFE689169D8FFDCAFD16FD1F" box="[654,751,635,661]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="17" type="Museum">NHMW</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="DD6736C4FFE689169FEBFD0AFD62FD5C" blockId="16.[189,1399,699,937]" box="[234,667,699,726]" pageId="16" pageNumber="17">
<emphasis id="EFACEAD6FFE689169FEBFD0AFE18FD5F" bold="true" box="[234,481,699,725]" pageId="16" pageNumber="17">Additional material</emphasis>
(
<specimenCount id="CBDEFD4DFFE689169EF3FD0DFD6AFD5C" box="[498,659,700,726]" count="17" pageId="16" pageNumber="17" type="generic">17 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFE689169FBCFD6EFE2CFC23" blockId="16.[189,1399,699,937]" pageId="16" pageNumber="17">
<materialsCitation id="6DB03C99FFE689169FBCFD6EFCB4FC97" collectionCode="BMNH" collectorName="U. P. India. &amp; H. G. Champion Coll." country="India" location="Kheri Forest" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFE689169FBCFD6EFEF7FD73" box="[189,270,735,761]" name="India" pageId="16" pageNumber="17">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFE689169E2CFD6EFEBFFD73" box="[301,326,735,761]" count="1" pageId="16" pageNumber="17" type="male"></specimenCount>
; “
<location id="D807601FFFE689169E61FD6EFE02FD73" LSID="urn:lsid:plazi:treatment:557187D2FFF989169D68F994FE37F82E:D807601FFFE689169E61FD6EFE02FD73" box="[352,507,735,761]" country="India" name="Kheri Forest" pageId="16" pageNumber="17">Kheri Forest</location>
,
<collectorName id="702D5312FFE689169D0BFD6EFD75FD73" box="[522,652,735,761]" pageId="16" pageNumber="17">U.P. India.</collectorName>
, Jan 16 HGC;
<collectorName id="702D5312FFE689169C4AFD6EFBB6FD73" box="[843,1103,735,761]" pageId="16" pageNumber="17">H.G. Champion Coll.</collectorName>
, B.M. 1953-156.;
<taxonomicName id="1AD84D47FFE689169A2EFD6EFDF4FC96" authority=", Redt. G. J. Arrow" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFE689169A2EFD6EFEC8FC96" italics="true" pageId="16" pageNumber="17">Onitis castaneus</emphasis>
, Redt. G. J. Arrow
</taxonomicName>
det.”;
<collectionCode id="BBC9AE01FFE689169D5AFCB2FD4AFC97" box="[603,691,771,797]" pageId="16" pageNumber="17">BMNH</collectionCode>
(E) 1237151
</materialsCitation>
<materialsCitation id="6DB03C99FFE689169C65FCB2FB27FCCA" collectionCode="BMNH" collectorName="C. F. C. Beeson &amp; Dehradun, B. M." country="India" location="Multan Div. Punjab" municipality="Gazighat" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFE689169C65FCB2FC82FC97" box="[868,891,771,797]" count="1" pageId="16" pageNumber="17" type="female"></specimenCount>
; “
<collectingMunicipality id="3D03ACBEFFE689169C95FCB3FBFAFC96" box="[916,1027,770,796]" pageId="16" pageNumber="17">Gazighat</collectingMunicipality>
,
<location id="D807601FFFE689169B0FFCB3FB0EFC96" LSID="urn:lsid:plazi:treatment:557187D2FFF989169D68F994FE37F82E:D807601FFFE689169B0FFCB3FB0EFC96" box="[1038,1271,770,796]" country="India" municipality="Gazighat" name="Multan Div. Punjab" pageId="16" pageNumber="17">Multan Div. Punjab</location>
, 15. 2. 28.,
<collectorName id="702D5312FFE689169FBCFC94FE93FCCA" box="[189,362,805,832]" pageId="16" pageNumber="17">C.F.C. Beeson</collectorName>
; ex coll.
<collectorName id="702D5312FFE689169ED5FC94FD6AFCCA" box="[468,659,805,832]" pageId="16" pageNumber="17">Dehradun, B.M.</collectorName>
1928-472., ex Cattle dung;”
<collectionCode id="BBC9AE01FFE689169CEAFC97FBBAFCCA" box="[1003,1091,806,832]" pageId="16" pageNumber="17">BMNH</collectionCode>
(E) 1237152
</materialsCitation>
<materialsCitation id="6DB03C99FFE689169BF9FC97FC48FCE9" collectionCode="ZSIM" collectorName="H. L. Khandu &amp; T. G. Vazirani" country="India" location="Pilani" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1" stateProvince="Rajasthan">
<specimenCount id="CBDEFD4DFFE689169BF9FC97FAF6FCCA" box="[1272,1295,806,832]" count="1" pageId="16" pageNumber="17" type="female"></specimenCount>
; “
<location id="D807601FFFE689169A29FC94FA8BFCB5" LSID="urn:lsid:plazi:treatment:557187D2FFF989169D68F994FE37F82E:D807601FFFE689169A29FC94FA8BFCB5" box="[1320,1394,805,831]" country="India" name="Pilani" pageId="16" pageNumber="17" stateProvince="Rajasthan">Pilani</location>
,
<collectingRegion id="1F1CF826FFE689169FBCFCF9FECCFCE8" box="[189,309,840,866]" country="India" name="Rajasthan" pageId="16" pageNumber="17">Rajasthan</collectingRegion>
; 1957; Coll:
<collectorName id="702D5312FFE689169ECAFCF8FD91FCE8" box="[459,616,840,867]" pageId="16" pageNumber="17">H.L. Khandu</collectorName>
, Det:
<collectorName id="702D5312FFE689169DADFCF8FCAFFCE8" box="[684,854,840,867]" pageId="16" pageNumber="17">T. G. Vazirani</collectorName>
”;
<collectionCode id="BBC9AE01FFE689169C6AFCF8FC48FCE9" box="[875,945,841,867]" pageId="16" pageNumber="17">ZSIM</collectionCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE689169CCBFCF8FE29FC23" collectingDate="2018-12-19" collectionCode="AIMB" collectorName="Rajkamal Goswami &amp; Co &amp; Sc" country="India" location="Kolia" municipality="Tezpur" pageId="16" pageNumber="17" specimenCount="14" specimenCount-female="4" specimenCount-male="10" stateProvince="Assam">
<specimenCount id="CBDEFD4DFFE689169CCBFCF8FBE5FCE9" box="[970,1052,841,867]" count="10" pageId="16" pageNumber="17" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE689169B28FCF8FB92FCE9" box="[1065,1131,841,867]" count="4" pageId="16" pageNumber="17" type="female">4 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE689169B79FCF8FB32FCE9" box="[1144,1227,841,867]" country="India" name="Assam" pageId="16" pageNumber="17">Assam</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE689169BD4FCF8FAD1FCE9" box="[1237,1320,841,867]" pageId="16" pageNumber="17">Tezpur</collectingMunicipality>
, under
<location id="D807601FFFE689169FBCFCDDFF07FC0C" LSID="urn:lsid:plazi:treatment:557187D2FFF989169D68F994FE37F82E:D807601FFFE689169FBCFCDDFF07FC0C" box="[189,254,876,902]" country="India" municipality="Tezpur" name="Kolia" pageId="16" pageNumber="17" stateProvince="Assam">Kolia</location>
bhomora bridge; hand-picked, cattle dung;
<date id="A9661004FFE689169C10FCDDFC4EFC0C" box="[785,951,876,902]" pageId="16" pageNumber="17" value="2018-12-19">
<collectingDate id="B922E9ECFFE689169C10FCDDFC4EFC0C" box="[785,951,876,902]" pageId="16" pageNumber="17" value="2018-12-19">19 Dec. 2018</collectingDate>
</date>
;
<collectorName id="702D5312FFE689169CC7FCDDFB48FC0C" box="[966,1201,876,902]" pageId="16" pageNumber="17">Rajkamal Goswami</collectorName>
leg.;
<collectionCode id="BBC9AE01FFE689169BF6FCDDFABAFC0C" box="[1271,1347,876,902]" pageId="16" pageNumber="17">AIMB</collectionCode>
/
<collectorName id="702D5312FFE689169A4DFCDDFA94FC0C" box="[1356,1389,876,902]" pageId="16" pageNumber="17">Co</collectorName>
/
<collectorName id="702D5312FFE689169FBCFC3EFF20FC23" box="[189,217,911,937]" pageId="16" pageNumber="17">Sc</collectorName>
1000023 to 1000036
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE689169FBCFC61FDC4F92D" pageId="16" pageNumber="17" type="description">
<paragraph id="DD6736C4FFE689169FBCFC61FEB7FC60" blockId="16.[189,334,976,1002]" box="[189,334,976,1002]" pageId="16" pageNumber="17">
<heading id="862F81A8FFE689169FBCFC61FEB7FC60" bold="true" box="[189,334,976,1002]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="EFACEAD6FFE689169FBCFC61FEB7FC60" bold="true" box="[189,334,976,1002]" pageId="16" pageNumber="17">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE689169FBCFC4EFE73FBB7" blockId="16.[189,1398,1023,1085]" pageId="16" pageNumber="17">MEASUREMENTS (in mm). TL=1619; BW=9.512; PL=6.57.5; PW=911; EL= 8.510; HL=4. 55.5; HW =56.</paragraph>
<paragraph id="DD6736C4FFE689169FEBFBD5FED3FBF4" blockId="16.[189,1399,1124,1467]" box="[234,298,1124,1150]" pageId="16" pageNumber="17">
<emphasis id="EFACEAD6FFE689169FEBFBD5FED3FBF4" bold="true" box="[234,298,1124,1150]" pageId="16" pageNumber="17">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE689169FBCFB36FEBAFA30" blockId="16.[189,1399,1124,1467]" pageId="16" pageNumber="17">Dark chestnut brown; head with strongly elevated frontal carina acuminate in the middle; clypeal margin strongly reflexed, straight in the middle, with long transverse arcuate carina, frontal carina straight, rugulose in front, with tubercle behind; vertex strongly excavated; pronotum unevenly punctured, strongly rugulose anteriorly, tuberculate between the basal foveae; front angles rather blunt, deep basal foveae closely placed; elytra opaque, finely striate, intervals flat, feebly punctured; pygidium sparsely hairy; metasternum closely tuberculate, hairy with a longitudinal median groove; front legs long, slender, strongly curved, armed with four short teeth externally, profemur with a strong oblique spine near the middle of anterior edge, middle tibia with two strong processes at its outer edge.</paragraph>
<paragraph id="DD6736C4FFE689169FBCFA53FA8EF995" blockId="16.[189,1399,1506,1567]" pageId="16" pageNumber="17">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFE689169EBBFA53FDB4FA76" box="[442,589,1506,1532]" captionStart="Fig" captionStartId="33.[189,232,1913,1939]" captionTargetBox="[295,1282,334,1816]" captionTargetId="figure-42@33.[189,1398,262,1833]" captionTargetPageId="33" captionText="Fig. 11. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitiscastaneus Kollar, 1844, AIMB/ Co/Sc1000027. EH. O. feae Felsche, 1907, AIMB/Co/Sc1000188. IL. O. lama Lansberge, 1875, AIMB/Co/Sc1000307. (Images: AIMB)." pageId="16" pageNumber="17">Fig. 11AD</figureCitation>
). LP=3.03, Lp =2.78; BP=1.32, BpB=1.07, BpT=0.43. Parameres shorter than phallobase, dorsal side of parameres straight at the base, tip broader and curved and tip blunt.
</paragraph>
<paragraph id="DD6736C4FFE689169FEBF9F6FEBCF9EB" blockId="16.[189,1398,1607,1703]" box="[234,325,1607,1633]" pageId="16" pageNumber="17">
<emphasis id="EFACEAD6FFE689169FEBF9F6FEBCF9EB" bold="true" box="[234,325,1607,1633]" pageId="16" pageNumber="17">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE689169FBCF9DBFDC4F92D" blockId="16.[189,1398,1607,1703]" pageId="16" pageNumber="17">Protibia short, broad bearing four strong external teeth; clypeus more rounded in the middle than male; frontal tubercle more prominent.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE689169FBCF97FFE37F82E" pageId="16" pageNumber="17" type="distribution">
<paragraph id="DD6736C4FFE689169FBCF97FFEAFF962" blockId="16.[189,342,1742,1768]" box="[189,342,1742,1768]" pageId="16" pageNumber="17">
<heading id="862F81A8FFE689169FBCF97FFEAFF962" bold="true" box="[189,342,1742,1768]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="EFACEAD6FFE689169FBCF97FFEAFF962" bold="true" box="[189,342,1742,1768]" pageId="16" pageNumber="17">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE689169FBCF94CFE37F82E" blockId="16.[189,1399,1789,1956]" pageId="16" pageNumber="17">
<collectingCountry id="A5CF7654FFE689169FBCF94CFF06F89D" box="[189,255,1789,1815]" name="India" pageId="16" pageNumber="17">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFE689169E0BF94CFE08F89D" box="[266,497,1789,1815]" country="India" name="Arunachal Pradesh" pageId="16" pageNumber="17">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169EFDF94CFDA9F89D" box="[508,592,1789,1815]" country="India" name="Assam" pageId="16" pageNumber="17">Assam</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169D5CF94CFD58F89D" box="[605,673,1789,1815]" country="India" name="Bihar" pageId="16" pageNumber="17">Bihar</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169DAFF94CFCF2F89D" box="[686,779,1789,1815]" country="India" name="Gujarat" pageId="16" pageNumber="17">Gujarat</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169C16F94CFC86F89D" box="[791,895,1789,1815]" country="India" name="Haryana" pageId="16" pageNumber="17">Haryana</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169C8CF94CFB92F89D" box="[909,1131,1789,1815]" country="India" name="Himachal Pradesh" pageId="16" pageNumber="17">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169B79F94CFA8BF89D" box="[1144,1394,1789,1815]" country="India" name="Jammu and Kashmir" pageId="16" pageNumber="17">Jammu and Kashmir</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169FBCF891FE6CF8B0" box="[189,405,1824,1850]" country="India" name="Madhya Pradesh" pageId="16" pageNumber="17">Madhya Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169EAFF891FDCFF8B0" box="[430,566,1824,1850]" country="India" name="Meghalaya" pageId="16" pageNumber="17">Meghalaya</collectingRegion>
, Punjab,
<collectingRegion id="1F1CF826FFE689169DBCF891FCCFF8B0" box="[701,822,1824,1850]" country="India" name="Rajasthan" pageId="16" pageNumber="17">Rajasthan</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169C4EF891FC1FF8B0" box="[847,998,1824,1850]" country="India" name="Uttarakhand" pageId="16" pageNumber="17">Uttarakhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169CFEF890FB4AF8B0" box="[1023,1203,1824,1851]" country="India" name="Uttar Pradesh" pageId="16" pageNumber="17">Uttar Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE689169BCAF890FA8BF8B0" box="[1227,1394,1824,1851]" country="India" name="West Bengal" pageId="16" pageNumber="17">West Bengal</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFE689169E8EF8F2FE23F8D7" box="[399,474,1859,1885]" name="Nepal" pageId="16" pageNumber="17">Nepal</collectingCountry>
;
<collectingCountry id="A5CF7654FFE689169EEAF8F2FDA9F8D7" box="[491,592,1859,1885]" name="Pakistan" pageId="16" pageNumber="17">Pakistan</collectingCountry>
(
<bibRefCitation id="B9494B35FFE689169D65F8F2FCEEF8D7" author="Chandra K." box="[612,791,1859,1885]" pageId="16" pageNumber="17" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE689169C29F8F2FC45F8D4" author="Mittal I. C." box="[808,956,1859,1886]" pageId="16" pageNumber="17" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE689169CCCF8F2FB27F8D7" author="Bezdek A. &amp; Krell F. - T." box="[973,1246,1859,1885]" pageId="16" pageNumber="17" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE689169BEEF8F2FE91F80B" author="Chandra K. &amp; Ahirwar S. C." pageId="16" pageNumber="17" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">Chandra &amp; Ahirwar 2007</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE689169E74F8D7FDE2F80A" author="Sewak R." box="[373,539,1894,1920]" pageId="16" pageNumber="17" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">Sewak 2009b</bibRefCitation>
; Singh
<emphasis id="EFACEAD6FFE689169D77F8D6FD4AF80A" box="[630,691,1894,1920]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2010;
<bibRefCitation id="B9494B35FFE689169C06F8D6FC25F80B" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[775,988,1894,1921]" pageId="16" pageNumber="17" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFE689169C59F8D6FC6CF80A" box="[856,917,1894,1920]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE689169CEBF8D7FB73F80A" author="Bezdek A." box="[1002,1162,1894,1920]" pageId="16" pageNumber="17" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE689169B99F8D7FA88F80B" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." box="[1176,1393,1894,1921]" pageId="16" pageNumber="17" pagination="221 - 275" refId="ref29141" refString="Ghosh J., Das, P., Ghosh S. K., Bhunia D., Kushwaha R. K., Gupta D., &amp; Chandra K. 2020. Insecta: Coleoptera. Zoological Survey of India, Fauna of Haryana, State Fauna Series 24: 221 - 275." type="book chapter" year="2020">
Ghosh
<emphasis id="EFACEAD6FFE689169BEFF8D6FAD3F80A" box="[1262,1322,1894,1920]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2020
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE689169FBCF838FE3BF82E" author="Schoolmeesters P." box="[189,450,1929,1956]" pageId="16" pageNumber="17" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,346 @@
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<paragraph id="DD6736C4FFFA890A9D5CF9AEFC2EF9B3" blockId="12.[605,983,1566,1629]" box="[605,983,1566,1593]" pageId="12" pageNumber="13">
<heading id="862F81A8FFFA890A9D5CF9AEFC2EF9B3" box="[605,983,1566,1593]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="1AD84D47FFFA890A9D5CF9AEFC2EF9B3" ID-CoL="74MW8" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[605,983,1566,1593]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFFA890A9D5CF9AEFCF4F9B3" bold="true" box="[605,781,1566,1593]" italics="true" pageId="12" pageNumber="13">Onitis brahma</emphasis>
<bibRefCitation id="B9494B35FFFA890A9C12F9AEFC2EF9B3" author="Lansberge G." box="[787,983,1566,1593]" pageId="12" pageNumber="13" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
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<subSubSection id="95C2654FFFFA890A9DD7F9F3FA95F805" pageId="12" pageNumber="13" type="description">
<paragraph id="DD6736C4FFFA890A9DD7F9F3FCA4F9D6" blockId="12.[605,983,1566,1629]" box="[726,861,1602,1629]" pageId="12" pageNumber="13">
<figureCitation id="45E32A41FFFA890A9DD7F9F3FCCDF9D7" box="[726,820,1602,1629]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="12" pageNumber="13">Figs 5B</figureCitation>
,
<figureCitation id="45E32A41FFFA890A9C41F9F3FCA4F9D6" box="[832,861,1602,1628]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." pageId="12" pageNumber="13">17</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFFA890A9FBCF932FC8CF917" blockId="12.[189,885,1667,1694]" box="[189,885,1667,1694]" pageId="12" pageNumber="13">
<emphasis id="EFACEAD6FFFA890A9FBCF932FE9EF917" box="[189,359,1667,1693]" italics="true" pageId="12" pageNumber="13">Onitisbrahma</emphasis>
<bibRefCitation id="B9494B35FFFA890A9E6EF932FD94F917" author="Lansberge G." box="[367,621,1667,1694]" pageId="12" pageNumber="13" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875: 142</bibRefCitation>
(original description).
</paragraph>
<paragraph id="DD6736C4FFFA890A9FBCF974FA95F805" blockId="12.[189,1399,1733,1935]" pageId="12" pageNumber="13">
<emphasis id="EFACEAD6FFFA890A9FBCF974FE9CF955" box="[189,357,1733,1759]" italics="true" pageId="12" pageNumber="13">Onitisbrahma</emphasis>
<bibRefCitation id="B9494B35FFFA890A9E86F974FCFEF955" author="Preudhomme de Borre C. F. P. A." box="[391,775,1733,1759]" pageId="12" pageNumber="13" refId="ref30731" refString="Preudhomme de Borre C. F. P. A. 1881. Bulletin ou Comptes-Rendus des Seances de la Societe entomologique de Belgique XXXIX." type="book" year="1881">Preudhomme De Borre 1881: 41</bibRefCitation>
(catalogue). —
<bibRefCitation id="B9494B35FFFA890A9CC7F974FB69F955" author="Arrow G. J." box="[966,1168,1733,1759]" pageId="12" pageNumber="13" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 399</bibRefCitation>
(keys, description). —
<bibRefCitation id="B9494B35FFFA890A9E16F959FE0EF888" author="Janssens A." box="[279,503,1768,1794]" pageId="12" pageNumber="13" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 74</bibRefCitation>
(revision). —
<bibRefCitation id="B9494B35FFFA890A9DB8F959FC5BF889" author="Balthasar V." box="[697,930,1768,1795]" pageId="12" pageNumber="13" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar 1963: 45</bibRefCitation>
(monograph). —
<bibRefCitation id="B9494B35FFFA890A9B8BF959FA8FF888" author="Chandra K." box="[1162,1398,1768,1794]" pageId="12" pageNumber="13" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000: 360</bibRefCitation>
(distribution). —
<bibRefCitation id="B9494B35FFFA890A9EBBF8BAFCDFF8AF" author="Chandra K. &amp; Ahirwar S. C." box="[442,806,1803,1830]" pageId="12" pageNumber="13" pagination="147 - 155" refId="ref28418" refString="Chandra K. &amp; Ahirwar S. C. 2005. Scarabaeid Beetles (Coleoptera) of Kanha Tiger Reserve, Madhya Pradesh. Records Zoological Survey of India 105 (1 - 2): 147 - 155." type="journal article" year="2005">Chandra &amp; Ahirwar 2005: 149</bibRefCitation>
(distribution); 2007: 286 (distribution). —
<bibRefCitation id="B9494B35FFFA890A9A2BF8BAFE98F8C2" author="Sewak R." pageId="12" pageNumber="13" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">Sewak 2009b: 61</bibRefCitation>
(distribution, diagnosis). — Chandra
<emphasis id="EFACEAD6FFFA890A9C21F89EFCA5F8C2" box="[800,860,1838,1864]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2011: 66 (distribution). —
<bibRefCitation id="B9494B35FFFA890A9BA4F89FFE90F8E6" author="Chandra K. &amp; Gupta D." pageId="12" pageNumber="13" pagination="812 - 826" refId="ref28584" refString="Chandra K. &amp; Gupta D. 2012 b. Diversity and composition of dung beetles (Scarabaeidae: Scarabaeinae and Aphodiinae) assemblages in Singhori Wildlife Sanctuary, Raisen, Madhya Pradesh (India). Munis Entomology and Zoology 7 (2): 812 - 826." type="journal article" year="2012">Chandra &amp; Gupta 2012b: 817</bibRefCitation>
(distribution). —
<bibRefCitation id="B9494B35FFFA890A9D33F8E0FC4EF8E6" author="Karimbumkara S. N. &amp; Rajan P. D." box="[562,951,1873,1900]" pageId="12" pageNumber="13" pagination="173 - 178" refId="ref29842" refString="Karimbumkara S. N. &amp; Rajan P. D. 2013. Insecta: Coleoptera: Scarabaeidae: Scarabaeinae, Dung Beetles. Zoological Survey of India, Fauna of Karnataka, State Fauna Series 21: 173 - 178." type="book chapter" year="2013">Karimbumkara &amp; Rajan 2013: 174</bibRefCitation>
(distribution). —
<bibRefCitation id="B9494B35FFFA890A9B7EF8E3FA8EF8E6" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[1151,1399,1873,1900]" pageId="12" pageNumber="13" pagination="225 - 240" refId="ref29198" refString="Gupta D., Chandra K. &amp; Khan S. 2014. An updated checklist of Scarabaeoid beetles (Coleoptera: Scarabaeoidea) of Pench Tiger Reserve, Madhya Pradesh, India. Journal of Entomology and Zoology Studies 2 (5): 225 - 240. Available from https: // www. entomoljournal. com / vol 2 Issue 5 / 57.1. html [accessed 12 Jun. 2024]" type="journal article" year="2014">
Gupta
<emphasis id="EFACEAD6FFFA890A9BCBF8E3FAF8F8E1" box="[1226,1281,1873,1899]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2014: 230
</bibRefCitation>
(distribution); 2015: 1034 (key, distribution, description). —
<bibRefCitation id="B9494B35FFFA890A9C9FF8C4FB69F805" author="Schoolmeesters P." box="[926,1168,1909,1935]" pageId="12" pageNumber="13" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</paragraph>
</subSubSection>
<caption id="89A7664CFFFB890B9FBCF8F1FB25F86D" pageId="13" pageNumber="14" startId="13.[189,232,1856,1882]" targetBox="[300,1241,290,1779]" targetPageId="13">
<paragraph id="DD6736C4FFFB890B9FBCF8F1FB25F86D" blockId="13.[189,1399,1856,2023]" pageId="13" pageNumber="14">
<emphasis id="EFACEAD6FFFB890B9FBCF8F1FEB1F8D0" bold="true" box="[189,328,1856,1882]" pageId="13" pageNumber="14">Fig. 4. AD</emphasis>
.
<taxonomicName id="1AD84D47FFFB890B9E55F8F0FD31F8D0" authority="Biswas, 1980" authorityName="Biswas" authorityYear="1980" box="[340,712,1856,1883]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFFB890B9E55F8F0FDDFF8D0" box="[340,550,1857,1883]" italics="true" pageId="13" pageNumber="14">Onitis assamensis</emphasis>
<bibRefCitation id="B9494B35FFFB890B9D2DF8F1FD31F8D0" author="Biswas S." box="[556,712,1856,1882]" pageId="13" pageNumber="14" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980</bibRefCitation>
</taxonomicName>
, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB).
<emphasis id="EFACEAD6FFFB890B9D47F8D2FD83F8F7" bold="true" box="[582,634,1891,1918]" pageId="13" pageNumber="14">EF</emphasis>
.
<taxonomicName id="1AD84D47FFFB890B9D8BF8D5FC1FF8F7" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[650,998,1891,1918]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFFB890B9D8BF8D5FCEDF8F7" box="[650,788,1892,1918]" italics="true" pageId="13" pageNumber="14">O. naviauxi</emphasis>
<bibRefCitation id="B9494B35FFFB890B9C1DF8D2FC1FF8F7" author="Cambefort Y." box="[796,998,1891,1917]" pageId="13" pageNumber="14" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN).
<emphasis id="EFACEAD6FFFB890B9C00F837FCCCF82B" bold="true" box="[769,821,1926,1953]" pageId="13" pageNumber="14">GJ</emphasis>
.
<taxonomicName id="1AD84D47FFFB890B9C47F836FB60F82B" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[838,1177,1926,1953]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bordati">
<emphasis id="EFACEAD6FFFB890B9C47F836FC3BF82A" box="[838,962,1926,1953]" italics="true" pageId="13" pageNumber="14">O. bordati</emphasis>
<bibRefCitation id="B9494B35FFFB890B9CCCF837FB60F82B" author="Cambefort Y." box="[973,1177,1926,1953]" pageId="13" pageNumber="14" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB).
<emphasis id="EFACEAD6FFFB890B9B33F81BFB94F84E" bold="true" box="[1074,1133,1962,1988]" pageId="13" pageNumber="14">KN</emphasis>
.
<taxonomicName id="1AD84D47FFFB890B9B7EF81BFA88F84E" authority="Gillet, 1911" authorityName="Gillet" authorityYear="1911" box="[1151,1393,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="siva">
<emphasis id="EFACEAD6FFFB890B9B7EF81BFB2CF84E" box="[1151,1237,1962,1988]" italics="true" pageId="13" pageNumber="14">O. siva</emphasis>
<bibRefCitation id="B9494B35FFFB890B9BE1F81BFA88F84E" author="Gillet J. J. E." box="[1248,1393,1962,1988]" pageId="13" pageNumber="14" pagination="313 - 314" refId="ref29108" refString="Gillet J. J. E. 1911. Coprides nouveaux de la region orientale (etc.). Annales de la Societe entomologique de Belgique LV: 313 - 314." type="book chapter" year="1911">Gillet, 1911</bibRefCitation>
</taxonomicName>
, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB).
</paragraph>
</caption>
<caption id="89A7664CFFF889089FBCF8A2FDE8F854" pageId="14" pageNumber="15" startId="14.[189,233,1811,1837]" targetBox="[327,1251,267,1773]" targetPageId="14">
<paragraph id="DD6736C4FFF889089FBCF8A2FDE8F854" blockId="14.[189,1399,1811,2014]" pageId="14" pageNumber="15">
<emphasis id="EFACEAD6FFF889089FBCF8A2FED0F8A7" bold="true" box="[189,297,1811,1838]" pageId="14" pageNumber="15">Fig. 5. A</emphasis>
.
<taxonomicName id="1AD84D47FFF889089E39F8A5FD4AF8A4" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[312,691,1811,1838]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bordati">
<emphasis id="EFACEAD6FFF889089E39F8A5FE19F8A7" box="[312,480,1811,1838]" italics="true" pageId="14" pageNumber="15">Onitis bordati</emphasis>
<bibRefCitation id="B9494B35FFF889089EE9F8A2FD4AF8A4" author="Cambefort Y." box="[488,691,1811,1838]" pageId="14" pageNumber="15" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN).
<emphasis id="EFACEAD6FFF889089E33F886FEBFF8DB" bold="true" box="[306,326,1847,1873]" pageId="14" pageNumber="15">B</emphasis>
.
<taxonomicName id="1AD84D47FFF889089E57F886FD53F8D8" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[342,682,1847,1874]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFF889089E57F886FE20F8DB" box="[342,473,1847,1873]" italics="true" pageId="14" pageNumber="15">O. brahma</emphasis>
<bibRefCitation id="B9494B35FFF889089EE3F886FD53F8D8" author="Lansberge G." box="[482,682,1847,1874]" pageId="14" pageNumber="15" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
, ♂, BMNH (E) 1237157, (image: BMNH).
<emphasis id="EFACEAD6FFF889089BBFF887FB2AF8DA" bold="true" box="[1214,1235,1846,1872]" pageId="14" pageNumber="15">C</emphasis>
.
<taxonomicName id="1AD84D47FFF889089BE2F886FE56F8FE" authority="Redtenbacher, 1848" authorityName="Redtenbacher" authorityYear="1848" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFF889089BE2F886FA8FF8DB" box="[1251,1398,1847,1873]" italics="true" pageId="14" pageNumber="15">O.castaneus</emphasis>
Redtenbacher, 1848
</taxonomicName>
, syntype, (image: Harald Schillhammer, NHMW).
<emphasis id="EFACEAD6FFF889089B29F8EBFBC4F8FE" bold="true" box="[1064,1085,1882,1908]" pageId="14" pageNumber="15">D</emphasis>
.
<taxonomicName id="1AD84D47FFF889089B4EF8EBFA96F8FF" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[1103,1391,1882,1909]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFF889089B4EF8EBFB28F8FE" box="[1103,1233,1882,1908]" italics="true" pageId="14" pageNumber="15">O. crassus</emphasis>
Sharp, 1875
</taxonomicName>
, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN).
<emphasis id="EFACEAD6FFF889089B09F8CCFBE5F81D" bold="true" box="[1032,1052,1917,1943]" pageId="14" pageNumber="15">E</emphasis>
.
<taxonomicName id="1AD84D47FFF889089B2FF8CFFA96F81D" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[1070,1391,1917,1944]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="EFACEAD6FFF889089B2FF8CFFB35F81D" box="[1070,1228,1918,1944]" italics="true" pageId="14" pageNumber="15">O. excavatus</emphasis>
<bibRefCitation id="B9494B35FFF889089BD4F8CCFA96F81D" author="Arrow G. J." box="[1237,1391,1917,1943]" pageId="14" pageNumber="15" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
, lectotype, ♂, BMNH (E) 1237173, (image: BMNH).
<emphasis id="EFACEAD6FFF889089C4CF811FCA6F830" bold="true" box="[845,863,1952,1978]" pageId="14" pageNumber="15">F</emphasis>
.
<taxonomicName id="1AD84D47FFF889089C73F810FB47F830" authority="(Wulfen, 1786)" baseAuthorityName="Wulfen" baseAuthorityYear="1786" box="[882,1214,1952,1979]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFF889089C73F810FC01F830" box="[882,1016,1952,1979]" italics="true" pageId="14" pageNumber="15">O. falcatus</emphasis>
(
<bibRefCitation id="B9494B35FFF889089B0AF811FB4CF830" author="Wulfen X." box="[1035,1205,1952,1978]" pageId="14" pageNumber="15" refId="ref31404" refString="Wulfen X. 1786. Descriptiones quorumdam Capensium insectorum. Sumptu Wolfgangi Waltheri, Erlangen." type="book" year="1786">Wulfen, 1786</bibRefCitation>
)
</taxonomicName>
, ♂, AIMB/Co/ Sc1000070, (image: AIMB).
</paragraph>
</caption>
<subSubSection id="95C2654FFFF989099FBCFEB9FE1CFE1D" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="DD6736C4FFF989099FBCFEB9FE3FFEA8" blockId="15.[189,454,264,290]" box="[189,454,264,290]" pageId="15" pageNumber="16">
<heading id="862F81A8FFF989099FBCFEB9FE3FFEA8" bold="true" box="[189,454,264,290]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="EFACEAD6FFF989099FBCFEB9FE3FFEA8" bold="true" box="[189,454,264,290]" pageId="15" pageNumber="16">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF989099FBCFE86FE1CFE1D" blockId="15.[189,1398,311,407]" pageId="15" pageNumber="16">
<emphasis id="EFACEAD6FFF989099FBCFE86FE93FEDB" box="[189,362,311,337]" italics="true" pageId="15" pageNumber="16">Onitisbrahma</emphasis>
belongs to Group VII and is closer to
<taxonomicName id="1AD84D47FFF989099C47FE86FC3DFEDB" authorityName="Sharp" authorityYear="1875" box="[838,964,311,337]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFF989099C47FE86FC3DFEDB" box="[838,964,311,337]" italics="true" pageId="15" pageNumber="16">O. crassus</emphasis>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFF989099B02FE86FB67FEDB" authorityName="Redtenbacher" authorityYear="1848" box="[1027,1182,311,337]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFF989099B02FE86FB67FEDB" box="[1027,1182,311,337]" italics="true" pageId="15" pageNumber="16">O. castaneus</emphasis>
</taxonomicName>
in having a hairy pygidium and punctured pronotum but differs from both in having an interrupted frontal carina and strongly excised clypeus.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF989099FBCFE0EFBACFD5C" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="DD6736C4FFF989099FBCFE0EFE53FE53" blockId="15.[189,426,447,473]" box="[189,426,447,473]" pageId="15" pageNumber="16">
<heading id="862F81A8FFF989099FBCFE0EFE53FE53" bold="true" box="[189,426,447,473]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="EFACEAD6FFF989099FBCFE0EFE53FE53" bold="true" box="[189,426,447,473]" pageId="15" pageNumber="16">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF989099FEBFE5FFEA5FD82" blockId="15.[189,1398,494,590]" box="[234,348,494,520]" pageId="15" pageNumber="16">
<emphasis id="EFACEAD6FFF989099FEBFE5FFEA5FD82" bold="true" box="[234,348,494,520]" pageId="15" pageNumber="16">
<typeStatus id="02638866FFF989099FEBFE5FFEA5FD82" box="[234,348,494,520]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFF989099FBCFDA0FCD1FDC4" blockId="15.[189,1398,494,590]" pageId="15" pageNumber="16">
<materialsCitation id="6DB03C99FFF989099FBCFDA0FCD1FDC4" collectionCode="MNHN" collectorName="Coll. R. Oberthur &amp; A. Janssens" country="India" location="INDIA" pageId="15" pageNumber="16" specimenCode="EC4809" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="A5CF7654FFF989099FBCFDA0FEF7FDA1" box="[189,270,529,555]" name="India" pageId="15" pageNumber="16">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFF989099E23FDA0FEC2FDA1" box="[290,315,529,555]" count="1" pageId="15" pageNumber="16" type="male"></specimenCount>
; “Mniszech, Ex-Musaeo Van Lansberge, Muséum Paris ex
<collectorName id="702D5312FFF989099CDFFDA0FB5EFDA1" box="[990,1191,529,555]" pageId="15" pageNumber="16">Coll. R. Oberthür</collectorName>
1952,
<collectorName id="702D5312FFF989099BF1FDA0FA8FFDA1" box="[1264,1398,529,555]" pageId="15" pageNumber="16">A. Janssens</collectorName>
vid., 1936:
<emphasis id="EFACEAD6FFF989099E42FD85FE14FDC4" box="[323,493,564,590]" italics="true" pageId="15" pageNumber="16">Onitisbrahma</emphasis>
Lansb.”,
<collectionCode id="BBC9AE01FFF989099D5EFD85FD40FDC4" box="[607,697,564,590]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="16">MNHN</collectionCode>
<specimenCode id="8D7E9EBFFFF989099DC1FD85FCDDFDC4" box="[704,804,564,590]" collectionCode="EC" pageId="15" pageNumber="16">EC4809</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFF989099FEBFDC4FD75FD05" blockId="15.[189,1399,629,726]" box="[234,652,629,655]" pageId="15" pageNumber="16">
<emphasis id="EFACEAD6FFF989099FEBFDC4FE18FD05" bold="true" box="[234,481,629,655]" pageId="15" pageNumber="16">Additional material</emphasis>
(
<specimenCount id="CBDEFD4DFFF989099EF0FDC4FD7DFD05" box="[497,644,629,655]" count="4" pageId="15" pageNumber="16" type="generic">4 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFF989099FBCFD28FBACFD5C" blockId="15.[189,1399,629,726]" pageId="15" pageNumber="16">
<materialsCitation id="6DB03C99FFF989099FBCFD28FA8EFD39" box="[189,1399,664,691]" collectionCode="BMNH" collectorName="H. Swale." country="India" location="INDIA" pageId="15" pageNumber="16" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFF989099FBCFD28FEF7FD39" box="[189,270,665,691]" name="India" pageId="15" pageNumber="16">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFF989099E2BFD28FEBBFD39" box="[298,322,665,691]" count="1" pageId="15" pageNumber="16" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFF989099E50FD28FE91FD39" box="[337,360,665,691]" count="1" pageId="15" pageNumber="16" type="female"></specimenCount>
; Dharwar, Bombay, “
<collectorName id="702D5312FFF989099D6AFD28FD18FD38" box="[619,737,664,691]" pageId="15" pageNumber="16">H. Swale.</collectorName>
1913117”,
<collectionCode id="BBC9AE01FFF989099C7AFD28FC2AFD39" box="[891,979,665,691]" pageId="15" pageNumber="16">BMNH</collectionCode>
(E) 1237157,
<collectionCode id="BBC9AE01FFF989099B80FD28FB20FD39" box="[1153,1241,665,691]" pageId="15" pageNumber="16">BMNH</collectionCode>
(E) 1237158
</materialsCitation>
<materialsCitation id="6DB03C99FFF989099FCEFD0DFBABFD5C" box="[207,1106,700,726]" collectingDate="1927-03-08" collectionCode="ZSIM" collectorName="Nerbuda Survey &amp; Barra &amp; Bewa State" country="India" elevation="823" pageId="15" pageNumber="16" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFF989099FCEFD0DFF1EFD5C" box="[207,231,700,726]" count="1" pageId="15" pageNumber="16" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFF989099FF2FD0DFEF3FD5C" box="[243,266,700,726]" count="1" pageId="15" pageNumber="16" type="female"></specimenCount>
;
<collectorName id="702D5312FFF989099E18FD0DFE20FD5C" box="[281,473,700,726]" pageId="15" pageNumber="16">Nerbuda Survey</collectorName>
,
<collectorName id="702D5312FFF989099EE4FD0DFDD0FD5C" box="[485,553,700,726]" pageId="15" pageNumber="16">Barra</collectorName>
,
<collectorName id="702D5312FFF989099D34FD0DFD44FD5C" box="[565,701,700,726]" pageId="15" pageNumber="16">Bewa State</collectorName>
,
<date id="A9661004FFF989099DC9FD0DFCA3FD5C" box="[712,858,700,726]" pageId="15" pageNumber="16" value="1927-03-08">
<collectingDate id="B922E9ECFFF989099DC9FD0DFCA3FD5C" box="[712,858,700,726]" pageId="15" pageNumber="16" value="1927-03-08">8 Mar. 1927</collectingDate>
</date>
;
<elevation id="56F5D1F7FFF989099C67FD0DFC03FD5C" box="[870,1018,700,726]" metricMagnitude="2" metricUnit="m" metricValue="8.2296" pageId="15" pageNumber="16" unit="ft" value="2700.0">
<quantity id="1A209B21FFF989099C67FD0DFC43FD5C" box="[870,954,700,726]" metricMagnitude="2" metricUnit="m" metricValue="8.2296" pageId="15" pageNumber="16" unit="ft" value="2700.0">2700 ft</quantity>
a.s.l.
</elevation>
;
<collectionCode id="BBC9AE01FFF989099B09FD0DFBABFD5C" box="[1032,1106,700,726]" pageId="15" pageNumber="16">ZSIM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF989099FBCFD4CFB63FAA3" pageId="15" pageNumber="16" type="description">
<paragraph id="DD6736C4FFF989099FBCFD4CFEB7FC9D" blockId="15.[189,334,765,791]" box="[189,334,765,791]" pageId="15" pageNumber="16">
<heading id="862F81A8FFF989099FBCFD4CFEB7FC9D" bold="true" box="[189,334,765,791]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="EFACEAD6FFF989099FBCFD4CFEB7FC9D" bold="true" box="[189,334,765,791]" pageId="15" pageNumber="16">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF989099FBCFC9DFCD0FCCC" blockId="15.[189,809,811,838]" box="[189,809,811,838]" pageId="15" pageNumber="16">MEASUREMENTS (in mm). TL=2126; BW=1113.5.</paragraph>
<paragraph id="DD6736C4FFF989099FEBFCDCFED3FC0D" blockId="15.[189,1399,877,1220]" box="[234,298,877,903]" pageId="15" pageNumber="16">
<emphasis id="EFACEAD6FFF989099FEBFCDCFED3FC0D" bold="true" box="[234,298,877,903]" pageId="15" pageNumber="16">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFF989099FBCFC21FC9FFB4E" blockId="15.[189,1399,877,1220]" pageId="15" pageNumber="16">Black or dark brown; clypeus shining, not closely granulate; clypeal margin strongly reflexed, excised in the middle; head finely granulate; frontal carina interrupted in the middle; pronotum shining, pronotum rugose anteriorly and punctured behind; elytra deeply striate, intervals convex, unpunctured, not shining; pygidium sparingly punctured; scanty hairs on pygidium long; metasternum with close reddish hairs, a raised longitudinal ridge in the middle; fore legs elongate, femur bears a blunt anterior process near the extremity, tibia slender, strongly curved towards the tip, armed with four external teeth, a very sharp tooth beneath, before the middle and a long incurved terminal process; mesofemur with a sharp tooth near the middle of its lower edge; hind femur with a sharp outwardly directed tooth at its upper edge near the base and a blunt lobe before middle of the lower edge.</paragraph>
<paragraph id="DD6736C4FFF989099FEBFB5AFEBCFA8F" blockId="15.[189,1178,1259,1321]" box="[234,325,1259,1285]" pageId="15" pageNumber="16">
<emphasis id="EFACEAD6FFF989099FEBFB5AFEBCFA8F" bold="true" box="[234,325,1259,1285]" pageId="15" pageNumber="16">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFF989099FBCFABFFB63FAA3" blockId="15.[189,1178,1259,1321]" box="[189,1178,1294,1321]" pageId="15" pageNumber="16">Clypeus more densely granular, not shining; pronotum closely rugose on major part.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF989099FBCFAE1FBA3FA55" pageId="15" pageNumber="16" type="distribution">
<paragraph id="DD6736C4FFF989099FBCFAE1FEAFFAE0" blockId="15.[189,342,1360,1386]" box="[189,342,1360,1386]" pageId="15" pageNumber="16">
<heading id="862F81A8FFF989099FBCFAE1FEAFFAE0" bold="true" box="[189,342,1360,1386]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="EFACEAD6FFF989099FBCFAE1FEAFFAE0" bold="true" box="[189,342,1360,1386]" pageId="15" pageNumber="16">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF989099FBCFACFFBA3FA55" blockId="15.[189,1398,1406,1504]" pageId="15" pageNumber="16">
<collectingCountry id="A5CF7654FFF989099FBCFACFFF07FA12" box="[189,254,1406,1432]" name="India" pageId="15" pageNumber="16">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFF989099E0AFACFFE37FA12" box="[267,462,1406,1432]" country="India" name="Andhra Pradesh" pageId="15" pageNumber="16">Andhra Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFF989099EDFFACFFD8EFA12" box="[478,631,1406,1432]" country="India" name="Chhattisgarh" pageId="15" pageNumber="16">Chhattisgarh</collectingRegion>
,
<collectingRegion id="1F1CF826FFF989099D87FACFFD18FA12" box="[646,737,1406,1432]" country="India" name="Gujarat" pageId="15" pageNumber="16">Gujarat</collectingRegion>
,
<collectingRegion id="1F1CF826FFF989099DF1FACFFC95FA12" box="[752,876,1406,1432]" country="India" name="Karnataka" pageId="15" pageNumber="16">Karnataka</collectingRegion>
,
<collectingRegion id="1F1CF826FFF989099C7DFACFFBBEFA12" box="[892,1095,1406,1432]" country="India" name="Madhya Pradesh" pageId="15" pageNumber="16">Madhya Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFF989099B57FACFFB15FA12" box="[1110,1260,1406,1432]" country="India" name="Maharashtra" pageId="15" pageNumber="16">Maharashtra</collectingRegion>
,
<collectingRegion id="1F1CF826FFF989099BFAFACFFA8BFA12" box="[1275,1394,1406,1432]" country="India" name="Rajasthan" pageId="15" pageNumber="16">Rajasthan</collectingRegion>
,
<collectingRegion id="1F1CF826FFF989099FBCFA13FEB0FA36" box="[189,329,1442,1468]" country="India" name="Tamil Nadu" pageId="15" pageNumber="16">Tamil Nadu</collectingRegion>
(
<bibRefCitation id="B9494B35FFF989099E58FA13FDFDFA36" author="Chandra K." box="[345,516,1442,1468]" pageId="15" pageNumber="16" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000</bibRefCitation>
;
<bibRefCitation id="B9494B35FFF989099D13FA13FCBAFA37" author="Chandra K. &amp; Ahirwar S. C." box="[530,835,1442,1469]" pageId="15" pageNumber="16" pagination="147 - 155" refId="ref28418" refString="Chandra K. &amp; Ahirwar S. C. 2005. Scarabaeid Beetles (Coleoptera) of Kanha Tiger Reserve, Madhya Pradesh. Records Zoological Survey of India 105 (1 - 2): 147 - 155." type="journal article" year="2005">Chandra &amp; Ahirwar 2005</bibRefCitation>
,
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</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:start id="DC39EB1439AC4A0D1A43122CEEB01E7F">1</mods:start>
<mods:end id="E554FCBA558A3078205DC8F8A60D1712">61</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:location id="2AFA2E5AB8E3CE8AC2AF13C940FE3C30">
<mods:url id="86C630853F9F45E1A1BA079851B99491">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287</mods:url>
</mods:location>
<mods:classification id="C598FA81C85377243363C8328D3494E2">journal article</mods:classification>
<mods:identifier id="8A1EBFB7657643F3973DFA33FCE92CA0" type="DOI">10.5852/ejt.2024.956.2657</mods:identifier>
<mods:identifier id="F9E1AC7453E51C02A2ECB5B33D1E73F3" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="BF93FD04E731E1F0796B823A4D4105CB" type="ZooBank">9DCDD521-909D-4436-8495-086A11C50DA0</mods:identifier>
</mods:mods>
<treatment id="557187D2FFFC890A9D5EFC0AFEA2FA53" LSID="urn:lsid:plazi:treatment:557187D2FFFC890A9D5EFC0AFEA2FA53" httpUri="http://treatment.plazi.org/id/557187D2FFFC890A9D5EFC0AFEA2FA53" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="95C2654FFFFC890C9D5EFC0AFC2CFC5C" box="[607,981,955,982]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="DD6736C4FFFC890C9D5EFC0AFC2CFC5C" blockId="10.[607,981,955,1018]" box="[607,981,955,982]" pageId="10" pageNumber="11">
<heading id="862F81A8FFFC890C9D5EFC0AFC2CFC5C" box="[607,981,955,982]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="1AD84D47FFFC890C9D5EFC0AFC2CFC5C" ID-CoL="74MVW" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[607,981,955,982]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bordati">
<emphasis id="EFACEAD6FFFC890C9D5EFC0AFCFFFC5C" bold="true" box="[607,774,955,982]" italics="true" pageId="10" pageNumber="11">Onitis bordati</emphasis>
<bibRefCitation id="B9494B35FFFC890C9C0CFC0AFC2CFC5C" author="Cambefort Y." box="[781,981,955,982]" pageId="10" pageNumber="11" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFC890C9DA1FC6EFC8AFBB0" pageId="10" pageNumber="11" type="description">
<paragraph id="DD6736C4FFFC890C9DA1FC6EFC6AFC73" blockId="10.[607,981,955,1018]" box="[672,915,991,1018]" pageId="10" pageNumber="11">
<figureCitation id="45E32A41FFFC890C9DA1FC6EFD13FC73" box="[672,746,991,1017]" captionStart="Fig" captionStartId="11.[189,232,1953,1979]" captionTargetBox="[280,1294,921,1858]" captionTargetId="figure-246@11.[189,1398,900,1917]" captionTargetPageId="11" captionText="Fig. 3. Onitis bordati Cambefort, 1988, ♀, AIMB/Co/Sc1000018. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head, dorsal view. D. Pygidium. (Images: AIMB)." pageId="10" pageNumber="11">Figs 3</figureCitation>
,
<figureCitation id="45E32A41FFFC890C9DF6FC6EFCC1FC73" box="[759,824,991,1017]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="10" pageNumber="11">4GJ</figureCitation>
,
<figureCitation id="45E32A41FFFC890C9C45FC6EFC93FC70" box="[836,874,991,1018]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="10" pageNumber="11">5A</figureCitation>
,
<figureCitation id="45E32A41FFFC890C9C77FC6EFC6AFC73" box="[886,915,991,1017]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." pageId="10" pageNumber="11">17</figureCitation>
</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCFB91FC8AFBB0" blockId="10.[189,883,1056,1082]" box="[189,883,1056,1082]" pageId="10" pageNumber="11">
<emphasis id="EFACEAD6FFFC890C9FBCFB91FE98FBB0" box="[189,353,1056,1082]" italics="true" pageId="10" pageNumber="11">Onitisbordati</emphasis>
<bibRefCitation id="B9494B35FFFC890C9E68FB91FD92FBB0" author="Cambefort Y." box="[361,619,1056,1082]" pageId="10" pageNumber="11" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988: 190</bibRefCitation>
(original description).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFC890C9FBCFBD3FE92FB48" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="DD6736C4FFFC890C9FBCFBD3FE92FB48" blockId="10.[189,1399,1121,1218]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="FDC811EAFFFC890C9FBCFBD3FE92FB48" pageId="10" pageNumber="11">
<emphasis id="EFACEAD6FFFC890C9FBCFBD3FE94FBF1" box="[189,365,1121,1148]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="1AD84D47FFFC890C9FBCFBD3FEA2FBF1" ID-CoL="74MVW" authorityName="Cambefort" authorityYear="1988" box="[189,347,1121,1148]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bordati">Onitis bordati</taxonomicName>
</emphasis>
<treatmentCitation id="5C7910D5FFFC890C9E71FBD0FD47FBF6" author="Kabakov O. N. &amp; Napolov A." box="[368,702,1121,1148]" page="70" pageId="10" pageNumber="11" year="1999">
<bibRefCitation id="B9494B35FFFC890C9E71FBD0FD47FBF6" author="Kabakov O. N. &amp; Napolov A." box="[368,702,1121,1148]" pageId="10" pageNumber="11" pagination="58 - 96" refId="ref29730" refString="Kabakov O. N. &amp; Napolov A. 1999. Fauna and ecology of Lamellicornia of subfamily Scarabaeinae (Scarabaeidae, Coleoptera) of Vietnam and some parts of adjacent countries: South China, Laos and Thailand. Latvijas Entomologs 37: 58 - 96." type="journal article" year="1999">Kabakov&amp;Napolov1999:70</bibRefCitation>
</treatmentCitation>
(distribution).—
<treatmentCitation id="5C7910D5FFFC890C9C84FBD0FB2AFBF1" author="Hanboonsong Y. &amp; Chunram S. &amp; Pimpasalee S. &amp; Emberso R. W. &amp; Masumoto K." box="[901,1235,1121,1148]" page="466" pageId="10" pageNumber="11" year="1999">
<bibRefCitation id="B9494B35FFFC890C9C84FBD0FB2AFBF1" author="Hanboonsong Y. &amp; Chunram S. &amp; Pimpasalee S. &amp; Emberso R. W. &amp; Masumoto K." box="[901,1235,1121,1148]" pageId="10" pageNumber="11" pagination="463 - 469" refId="ref29406" refString="Hanboonsong Y., Chunram S., Pimpasalee S., Emberso R. W. &amp; Masumoto K. 1999. The dung beetle fauna (Coleoptera: Scarabaeidae) of Northeast Thailand. Elytra, Tokyo 27 (2): 463 - 469." type="journal article" year="1999">
Hanboonsong
<emphasis id="EFACEAD6FFFC890C9B28FBD3FBA6FBF1" box="[1065,1119,1121,1147]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
1999:466
</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFFC890C9E0CFB34FD6BFB15" author="Hanboonsong Y. &amp; Masumoto K." box="[269,658,1157,1183]" pageId="10" pageNumber="11" pagination="101 - 114" refId="ref29369" refString="Hanboonsong Y. &amp; Masumoto K. 2000. Dung beetles (Coleoptera, Scarabaeidae) of Thailand, Part 2, Genus Onitis. Elytra, Tokyo 28 (1): 101 - 114." type="journal article" year="2000">Hanboonsong &amp; Masumoto 2000</bibRefCitation>
. — 103, 108 (key, distribution). —
<bibRefCitation id="B9494B35FFFC890C9B29FB34FAE7FB15" author="Schoolmeesters P." box="[1064,1310,1157,1183]" pageId="10" pageNumber="11" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFC890C9FBCFB58FE37FAF2" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="DD6736C4FFFC890C9FBCFB58FE3FFA89" blockId="10.[189,454,1257,1283]" box="[189,454,1257,1283]" pageId="10" pageNumber="11">
<heading id="862F81A8FFFC890C9FBCFB58FE3FFA89" bold="true" box="[189,454,1257,1283]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="EFACEAD6FFFC890C9FBCFB58FE3FFA89" bold="true" box="[189,454,1257,1283]" pageId="10" pageNumber="11">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCFAA9FE37FAF2" blockId="10.[189,1398,1304,1400]" pageId="10" pageNumber="11">
<emphasis id="EFACEAD6FFFC890C9FBCFAA9FE98FAB8" box="[189,353,1304,1330]" italics="true" pageId="10" pageNumber="11">Onitisbordati</emphasis>
belongs to Group IX and it resembles species of Group XVIII in having an uninterrupted frontal carina. It resembles
<taxonomicName id="1AD84D47FFFC890C9D01FA8DFDAAFADF" authorityName="Gillet" authorityYear="1911" box="[512,595,1340,1366]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="siva">
<emphasis id="EFACEAD6FFFC890C9D01FA8DFDAAFADF" box="[512,595,1340,1366]" italics="true" pageId="10" pageNumber="11">O. siva</emphasis>
</taxonomicName>
, in the absence of clypeal carina and differs from it in the pronotum being densely granular.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFC890C9FBCFA2EFD64F8B5" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="DD6736C4FFFC890C9FBCFA2EFE53FA33" blockId="10.[189,426,1439,1465]" box="[189,426,1439,1465]" pageId="10" pageNumber="11">
<heading id="862F81A8FFFC890C9FBCFA2EFE53FA33" bold="true" box="[189,426,1439,1465]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="EFACEAD6FFFC890C9FBCFA2EFE53FA33" bold="true" box="[189,426,1439,1465]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFC890C9FEBFA7FFEA5FA62" blockId="10.[189,1398,1486,1583]" box="[234,348,1486,1512]" pageId="10" pageNumber="11">
<emphasis id="EFACEAD6FFFC890C9FEBFA7FFEA5FA62" bold="true" box="[234,348,1486,1512]" pageId="10" pageNumber="11">
<typeStatus id="02638866FFFC890C9FEBFA7FFEA5FA62" box="[234,348,1486,1512]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCFA43FDF0F9A5" blockId="10.[189,1398,1486,1583]" pageId="10" pageNumber="11">
<materialsCitation id="6DB03C99FFFC890C9FBCFA43FDF0F9A5" collectionCode="MNHN" collectorName="Coll. H. Perrot &amp; Coll. P. Bordat" country="Vietnam" determinerName="Y. Cambefort" latitude="11.583333" location="VIETNAM" longLatPrecision="1295" longitude="108.083336" pageId="10" pageNumber="11" specimenCode="EC1187" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="A5CF7654FFFC890C9FBCFA43FEBDF986" box="[189,324,1522,1548]" name="Vietnam" pageId="10" pageNumber="11">VIETNAM</collectingCountry>
<specimenCount id="CBDEFD4DFFFC890C9E5AFA43FE8DF986" box="[347,372,1522,1548]" count="1" pageId="10" pageNumber="11" type="male"></specimenCount>
; Djiring, Annam;
<geoCoordinate id="B8EC5003FFFC890C9D45FA41FD54F986" box="[580,685,1520,1548]" degrees="11" direction="north" minutes="35" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="11.583333">11°35ʹ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFFC890C9DB9FA41FCD7F986" box="[696,814,1520,1548]" degrees="108" direction="east" minutes="05" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="108.083336">108°05ʹ E</geoCoordinate>
;
<collectorName id="702D5312FFFC890C9C3BFA40FC11F986" box="[826,1000,1521,1548]" pageId="10" pageNumber="11">Coll. H. Perrot</collectorName>
;
<collectorName id="702D5312FFFC890C9CF3FA40FB65F981" box="[1010,1180,1521,1548]" pageId="10" pageNumber="11">Coll. P. Bordat</collectorName>
leg.;
<determinerName id="28BBDA00FFFC890C9BDBFA43FA8FF981" box="[1242,1398,1521,1548]" pageId="10" pageNumber="11">Y. Cambefort</determinerName>
det. 1987;
<collectionCode id="BBC9AE01FFFC890C9E3BF9A4FE6EF9A5" box="[314,407,1557,1583]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
,
<specimenCode id="8D7E9EBFFFFC890C9EA3F9A4FDFCF9A5" box="[418,517,1557,1583]" collectionCode="EC" pageId="10" pageNumber="11">EC1187</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="DD6736C4FFFC890C9FEBF9E7FD75F9FA" blockId="10.[189,1398,1622,1719]" box="[234,652,1622,1648]" pageId="10" pageNumber="11">
<emphasis id="EFACEAD6FFFC890C9FEBF9E7FE18F9FA" bold="true" box="[234,481,1622,1648]" pageId="10" pageNumber="11">Additional material</emphasis>
(
<specimenCount id="CBDEFD4DFFFC890C9EF3F9E7FD7DF9FA" box="[498,644,1622,1648]" count="8" pageId="10" pageNumber="11" type="generic">8 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCF9CBFDEFF93D" blockId="10.[189,1398,1622,1719]" pageId="10" pageNumber="11">
<materialsCitation id="6DB03C99FFFC890C9FBCF9CBFDEBF93D" collectingDate="1991-06-01" collectingDateMax="1991-06-06" collectingDateMin="1991-06-01" collectionCode="MNHN" country="Thailand" location="TAK Region" pageId="10" pageNumber="11" specimenCode="EC1185" specimenCount="1" specimenCount-female="1" stateProvince="Tak">
<collectingCountry id="A5CF7654FFFC890C9FBCF9CBFEACF91E" box="[189,341,1658,1684]" name="Thailand" pageId="10" pageNumber="11">THAILAND</collectingCountry>
<specimenCount id="CBDEFD4DFFFC890C9E6EF9CBFE7FF91E" box="[367,390,1658,1684]" count="1" pageId="10" pageNumber="11" type="female"></specimenCount>
; “Prov.
<collectingRegion id="1F1CF826FFFC890C9EE7F9CBFD84F919" box="[486,637,1657,1684]" country="Thailand" name="Tak" pageId="10" pageNumber="11">TAK Region</collectingRegion>
de UMPHANG VI;
<date id="A9661004FFFC890C9C75F9C8FC04F919" box="[884,1021,1657,1684]" pageId="10" pageNumber="11" value="1991-06-01">
<collectingDate id="B922E9ECFFFC890C9C75F9C8FC04F919" box="[884,1021,1657,1684]" pageId="10" pageNumber="11" value="1991-06-01">1 Jun. 1991</collectingDate>
</date>
<date id="A9661004FFFC890C9B0CF9C8FB60F919" box="[1037,1177,1657,1684]" pageId="10" pageNumber="11" value="1991-06-06">
<collectingDate id="B922E9ECFFFC890C9B0CF9C8FB60F919" box="[1037,1177,1657,1684]" pageId="10" pageNumber="11" value="1991-06-06">6 Jun. 1991</collectingDate>
</date>
; Y. CAMBEFORT det. 1992”;
<collectionCode id="BBC9AE01FFFC890C9E46F92CFE5DF93D" box="[327,420,1693,1719]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
,
<specimenCode id="8D7E9EBFFFFC890C9EAEF92DFDEBF93D" box="[431,530,1692,1719]" collectionCode="EC" pageId="10" pageNumber="11">EC1185</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCF96FFD64F8B5" blockId="10.[189,1398,1756,1855]" pageId="10" pageNumber="11">
<materialsCitation id="6DB03C99FFFC890C9FBCF96FFD61F8B5" collectingDate="2012-05-20" collectionCode="AIMB" collectorName="Karimbumkara Seena Narayanan &amp; Rajkamal Goswami &amp; Co &amp; Sc" country="India" elevation="269" latitude="25.915697" location="Nongkhyllem" longLatPrecision="1" longitude="91.76794" pageId="10" pageNumber="11" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Meghalaya">
<collectingCountry id="A5CF7654FFFC890C9FBCF96FFEF7F972" box="[189,270,1758,1784]" name="India" pageId="10" pageNumber="11">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFFC890C9E2DF96FFE8BF973" box="[300,370,1758,1785]" count="2" pageId="10" pageNumber="11" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFFC890C9E80F96FFE3FF973" box="[385,454,1758,1785]" count="5" pageId="10" pageNumber="11" type="female">5 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFFC890C9ED6F96FFDA5F972" box="[471,604,1758,1784]" country="India" name="Meghalaya" pageId="10" pageNumber="11">Meghalaya</collectingRegion>
,
<location id="D807601FFFFC890C9D6BF96FFCF6F972" LSID="urn:lsid:plazi:treatment:557187D2FFFC890A9D5EFC0AFEA2FA53:D807601FFFFC890C9D6BF96FFCF6F972" box="[618,783,1758,1784]" country="India" latitude="25.915697" longLatPrecision="1" longitude="91.76794" name="Nongkhyllem" pageId="10" pageNumber="11" stateProvince="Meghalaya">Nongkhyllem</location>
;
<geoCoordinate id="B8EC5003FFFC890C9C1CF96DFC23F973" box="[797,986,1756,1785]" degrees="25" direction="north" minutes="54" orientation="latitude" pageId="10" pageNumber="11" precision="1" seconds="56.51" value="25.915697">25°54ʹ56.51″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFFC890C9CE9F96DFB6AF973" box="[1000,1171,1756,1785]" degrees="91" direction="east" minutes="46" orientation="longitude" pageId="10" pageNumber="11" precision="1" seconds="4.55" value="91.76794">91°46ʹ4.55″ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFFC890C9BE3F96FFA89F972" box="[1250,1392,1758,1785]" metricMagnitude="2" metricUnit="m" metricValue="2.69" pageId="10" pageNumber="11" unit="m" value="269.0">
<quantity id="1A209B21FFFC890C9BE3F96FFAD7F973" box="[1250,1326,1758,1785]" metricMagnitude="2" metricUnit="m" metricValue="2.69" pageId="10" pageNumber="11" unit="m" value="269.0">269 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFFC890C9FBCF8B0FE9FF891" box="[189,358,1793,1820]" pageId="10" pageNumber="11" value="2012-05-20">
<collectingDate id="B922E9ECFFFC890C9FBCF8B0FE9FF891" box="[189,358,1793,1820]" pageId="10" pageNumber="11" value="2012-05-20">20 May 2012</collectingDate>
</date>
;
<collectorName id="702D5312FFFC890C9E76F8B0FCFFF896" box="[375,774,1793,1820]" pageId="10" pageNumber="11">Karimbumkara Seena Narayanan</collectorName>
&amp;
<collectorName id="702D5312FFFC890C9C37F8B0FBDBF891" box="[822,1058,1793,1819]" pageId="10" pageNumber="11">Rajkamal Goswami</collectorName>
leg.; hand-picked, elephant dung;
<collectionCode id="BBC9AE01FFFC890C9E04F895FEABF8B4" box="[261,338,1828,1854]" pageId="10" pageNumber="11">AIMB</collectionCode>
/
<collectorName id="702D5312FFFC890C9E5AF895FE84F8B4" box="[347,381,1828,1854]" pageId="10" pageNumber="11">Co</collectorName>
/
<collectorName id="702D5312FFFC890C9E87F895FE5BF8B4" box="[390,418,1828,1854]" pageId="10" pageNumber="11">Sc</collectorName>
1000016 to 1000022
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFC890A9FBCF8D7FD34FC3C" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
<paragraph id="DD6736C4FFFC890C9FBCF8D7FEB7F80A" blockId="10.[189,334,1894,1920]" box="[189,334,1894,1920]" pageId="10" pageNumber="11">
<heading id="862F81A8FFFC890C9FBCF8D7FEB7F80A" bold="true" box="[189,334,1894,1920]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="EFACEAD6FFFC890C9FBCF8D7FEB7F80A" bold="true" box="[189,334,1894,1920]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCF824FE31F858" blockId="10.[189,1399,1940,2002]" pageId="10" pageNumber="11">MEASUREMENTS (in mm). TL=23.532; BW =11.515.5; PL=912; PW=11.515.5; EL=11.515; HL=69; HW= 78.5.</paragraph>
<paragraph id="DD6736C4FFFD890D9FEBFEB8FE66FEAE" blockId="11.[189,1399,265,855]" box="[234,415,265,292]" pageId="11" pageNumber="12">
<emphasis id="EFACEAD6FFFD890D9FEBFEB8FED3FEA9" bold="true" box="[234,298,265,291]" pageId="11" pageNumber="12">Male</emphasis>
(
<figureCitation id="45E32A41FFFD890D9E3BFEB8FE6CFEAE" box="[314,405,265,292]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." pageId="11" pageNumber="12">Fig. 5A</figureCitation>
)
</paragraph>
<paragraph id="DD6736C4FFFD890A9FBCFE9DFBC9FDB6" blockId="11.[189,1399,265,855]" lastBlockId="12.[189,1399,265,573]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">Body including legs and antennae reddish black, except the antennal club which is reddish yellow; moderately shining, elongate oval, slightly convex, devoid of any setae on the dorsal surface except at the lower margin of the elytra where a few scattered setae are present. Head with clypeus elongate, strongly rounded, slightly reflexed anteriorly, clypeal carina absent; genae strongly curved, extended outwards and upwards anteriorly, with fine granules; vertex significantly depressed, bordered behind, finely, densely granulate. Pronotum smoothly margined on all sides, the sides of pronotum straight anteriorly, slightly sinuate behind; front angles sharp, obtuse, hind angles obsolete; base angulate in the middle; basal foveae shallow, deeper towards the base, with small ovate granules both within and between them; rows of spindle shaped granules present around the foveae which are not very close, closer and longer in the middle of the pronotum, squamiform and closer anteriorly, the granules smaller, ovate, not very close in the lateral margins; a trace of longitudinal groove extends as a slight depression from the front of the foveae to the front margin of the pronotum; lateral foveae with fine close ovate granules; scutellum very small. Elytra finely striate, striae not perceptibly punctured; intervals flat, opaque, with scattered imperceptible punctures. The sutural interval slightly convex, shining with visible fine scattered punctures; the 5th interval with minute, close, slightly rugose punctures. The lateral carina of the elytra sharp. Pygidium without any hair or setae, opaque, rugose with minute scattered granules. Metasternal shield closely granular and densely hairy at the sides, less granular and hairy in the middle, smooth with imperceptible granules anteriorly and at the base. Sides of the metasternum finely, closely and evenly granular with dense hairs; the hairs shorter than those on the shield. Front legs elongate, tibia slender, strongly curved towards the end with the tip produced into a finger-like process, terminal spur absent, teeth short and triangular; lower side of protibia bears a tooth almost halfway from the base, a shorter one near the base and another short sharper one between these two, the inner side serrate. A tuft of short setae is present in an oval groove near the base of profemur, the upper margins of the legs fringed. The mesofemur is with a sharp tooth just beyond the middle of the lower edge, the hind femur without any teeth. The femoral teeth may not be prominent in minor males.</paragraph>
<caption id="89A7664CFFFD890D9FBCF810FC29F855" pageId="11" pageNumber="12" startId="11.[189,232,1953,1979]" targetBox="[280,1294,921,1858]" targetPageId="11">
<paragraph id="DD6736C4FFFD890D9FBCF810FC29F855" blockId="11.[189,1398,1953,2015]" pageId="11" pageNumber="12">
<emphasis id="EFACEAD6FFFD890D9FBCF810FEF5F831" bold="true" box="[189,268,1953,1979]" pageId="11" pageNumber="12">Fig. 3.</emphasis>
<taxonomicName id="1AD84D47FFFD890D9E15F813FD71F836" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[276,648,1954,1980]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bordati">
<emphasis id="EFACEAD6FFFD890D9E15F813FE41F836" box="[276,440,1954,1980]" italics="true" pageId="11" pageNumber="12">Onitis bordati</emphasis>
<bibRefCitation id="B9494B35FFFD890D9EC1F813FD71F836" author="Cambefort Y." box="[448,648,1954,1980]" pageId="11" pageNumber="12" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
, ♀, AIMB/Co/Sc1000018.
<emphasis id="EFACEAD6FFFD890D9CCCF810FC1BF831" bold="true" box="[973,994,1953,1979]" pageId="11" pageNumber="12">A</emphasis>
. Habitus, dorsal view.
<emphasis id="EFACEAD6FFFD890D9BF0F813FAFCF836" bold="true" box="[1265,1285,1954,1980]" pageId="11" pageNumber="12">B</emphasis>
. Habitus, ventral view.
<emphasis id="EFACEAD6FFFD890D9E5DF874FE88F855" bold="true" box="[348,369,1989,2015]" pageId="11" pageNumber="12">C</emphasis>
. Head, dorsal view.
<emphasis id="EFACEAD6FFFD890D9D60F874FD8FF855" bold="true" box="[609,630,1989,2015]" pageId="11" pageNumber="12">D</emphasis>
. Pygidium. (Images: AIMB).
</paragraph>
</caption>
<paragraph id="DD6736C4FFFA890A9FBCFDD5FE80FD4E" blockId="12.[189,1399,612,709]" pageId="12" pageNumber="13">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFFA890A9EC0FDD5FDB9FDF4" box="[449,576,612,638]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. AD. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). EF. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). GJ. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). KN. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." pageId="12" pageNumber="13">Fig. 4GJ</figureCitation>
). LP= 3.54.5, Lp=4.55; BP=2, BpB =1.5, BpT=0.5. Phallobase straight. Parameres longer than the phallobase, broader at the base and further tapers and bluntly curved above at the tip.
</paragraph>
<paragraph id="DD6736C4FFFA890A9FEBFD5DFE27FC8C" blockId="12.[189,1399,748,950]" box="[234,478,748,774]" pageId="12" pageNumber="13">
<emphasis id="EFACEAD6FFFA890A9FEBFD5DFEBCFC8C" bold="true" box="[234,325,748,774]" pageId="12" pageNumber="13">Female</emphasis>
(
<figureCitation id="45E32A41FFFA890A9E57FD5DFE2FFC8C" box="[342,470,748,774]" captionStart="Fig" captionStartId="11.[189,232,1953,1979]" captionTargetBox="[280,1294,921,1858]" captionTargetId="figure-246@11.[189,1398,900,1917]" captionTargetPageId="11" captionText="Fig. 3. Onitis bordati Cambefort, 1988, ♀, AIMB/Co/Sc1000018. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head, dorsal view. D. Pygidium. (Images: AIMB)." pageId="12" pageNumber="13">Fig. 3AD</figureCitation>
)
</paragraph>
<paragraph id="DD6736C4FFFA890A9FBCFCBEFD34FC3C" blockId="12.[189,1399,748,950]" pageId="12" pageNumber="13">Body oval, not as convex as male; clypeus elongate, strongly elliptical, slightly reflexed anteriorly, closely rugose. The posterior carina curved inwards, with a laterally compressed conical tubercle in the middle behind. Pygidium more shining than in male, rugose with imperceptible scattered granules. Profemur not toothed, tuft of short setae near the base absent. Protibia with an articulated terminal spur; teeth blunt, closer to each other than in male.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFA890A9FBCFC6CFE30FB56" pageId="12" pageNumber="13" type="distribution">
<paragraph id="DD6736C4FFFA890A9FBCFC6CFEAFFC7D" blockId="12.[189,342,989,1015]" box="[189,342,989,1015]" pageId="12" pageNumber="13">
<heading id="862F81A8FFFA890A9FBCFC6CFEAFFC7D" bold="true" box="[189,342,989,1015]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="EFACEAD6FFFA890A9FBCFC6CFEAFFC7D" bold="true" box="[189,342,989,1015]" pageId="12" pageNumber="13">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFA890A9FBCFBBDFC53FBC3" blockId="12.[189,1398,1036,1098]" pageId="12" pageNumber="13">
<collectingCountry id="A5CF7654FFFA890A9FBCFBBDFF07FBAC" box="[189,254,1036,1062]" name="India" pageId="12" pageNumber="13">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFFA890A9E08FBBDFE77FBAC" box="[265,398,1036,1062]" country="India" name="Meghalaya" pageId="12" pageNumber="13">Meghalaya</collectingRegion>
, other countries:
<collectingCountry id="A5CF7654FFFA890A9D5AFBBDFD38FBAC" box="[603,705,1036,1062]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
,
<collectingCountry id="A5CF7654FFFA890A9DCCFBBDFCCCFBAC" box="[717,821,1036,1062]" name="Thailand" pageId="12" pageNumber="13">Thailand</collectingCountry>
(
<bibRefCitation id="B9494B35FFFA890A9C44FBBDFB8DFBAC" author="Hanboonsong Y. &amp; Chunram S. &amp; Pimpasalee S. &amp; Emberso R. W. &amp; Masumoto K." box="[837,1140,1036,1062]" pageId="12" pageNumber="13" pagination="463 - 469" refId="ref29406" refString="Hanboonsong Y., Chunram S., Pimpasalee S., Emberso R. W. &amp; Masumoto K. 1999. The dung beetle fauna (Coleoptera: Scarabaeidae) of Northeast Thailand. Elytra, Tokyo 27 (2): 463 - 469." type="journal article" year="1999">
Hanboonsong
<emphasis id="EFACEAD6FFFA890A9CF3FBBCFBD7FBAC" box="[1010,1070,1036,1062]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
1999
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFFA890A9B80FBBDFF02FBC3" author="Kabakov O. N. &amp; Napolov A." pageId="12" pageNumber="13" pagination="58 - 96" refId="ref29730" refString="Kabakov O. N. &amp; Napolov A. 1999. Fauna and ecology of Lamellicornia of subfamily Scarabaeinae (Scarabaeidae, Coleoptera) of Vietnam and some parts of adjacent countries: South China, Laos and Thailand. Latvijas Entomologs 37: 58 - 96." type="journal article" year="1999">Kabakov &amp; Napolov 1999</bibRefCitation>
;
<bibRefCitation id="B9494B35FFFA890A9E06FB9EFD6BFBC3" author="Hanboonsong Y. &amp; Masumoto K." box="[263,658,1071,1098]" pageId="12" pageNumber="13" pagination="101 - 114" refId="ref29369" refString="Hanboonsong Y. &amp; Masumoto K. 2000. Dung beetles (Coleoptera, Scarabaeidae) of Thailand, Part 2, Genus Onitis. Elytra, Tokyo 28 (1): 101 - 114." type="journal article" year="2000">Hanboonsong &amp; Masumoto 2000</bibRefCitation>
;
<bibRefCitation id="B9494B35FFFA890A9D9EFB9EFC67FBC3" author="Schoolmeesters P." box="[671,926,1071,1097]" pageId="12" pageNumber="13" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
<paragraph id="DD6736C4FFFA890A9FBCFBC1FEE4FB00" blockId="12.[189,285,1136,1162]" box="[189,285,1136,1162]" pageId="12" pageNumber="13">
<heading id="862F81A8FFFA890A9FBCFBC1FEE4FB00" bold="true" box="[189,285,1136,1162]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="EFACEAD6FFFA890A9FBCFBC1FEE4FB00" bold="true" box="[189,285,1136,1162]" pageId="12" pageNumber="13">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFA890A9FBCFB11FE30FB56" blockId="12.[189,1399,1183,1244]" pageId="12" pageNumber="13">
In
<collectingCountry id="A5CF7654FFFA890A9FDDFB2EFEE2FB33" box="[220,283,1183,1209]" name="India" pageId="12" pageNumber="13">India</collectingCountry>
, this species was collected from bamboo dominated secondary forest near Nonkhyllem Wildlife Sanctuary,
<collectingRegion id="1F1CF826FFFA890A9E3EFB73FE3CFB56" box="[319,453,1218,1244]" country="India" name="Meghalaya" pageId="12" pageNumber="13">Meghalaya</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFA890A9FBCFAB5FEA2FA53" pageId="12" pageNumber="13" type="discussion">
<paragraph id="DD6736C4FFFA890A9FBCFAB5FED6FA94" blockId="12.[189,303,1284,1310]" box="[189,303,1284,1310]" pageId="12" pageNumber="13">
<heading id="862F81A8FFFA890A9FBCFAB5FED6FA94" bold="true" box="[189,303,1284,1310]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="EFACEAD6FFFA890A9FBCFAB5FED6FA94" bold="true" box="[189,303,1284,1310]" pageId="12" pageNumber="13">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFA890A9FBCFA83FEA2FA53" blockId="12.[189,1399,1330,1497]" pageId="12" pageNumber="13">
<bibRefCitation id="B9494B35FFFA890A9FBCFA83FE6BFAC6" author="Cambefort Y." box="[189,402,1330,1356]" pageId="12" pageNumber="13" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort (1988)</bibRefCitation>
did not describe the female of
<emphasis id="EFACEAD6FFFA890A9C09FA83FC57FAC6" box="[776,942,1330,1356]" italics="true" pageId="12" pageNumber="13">Onitisbordati</emphasis>
, but a female of the species deposited at MNHN was determined by Cambefort (1992) (https://shorturl.at/v4Dlr). In this paper, we provide a description for the female of
<taxonomicName id="1AD84D47FFFA890A9D1EFAC8FD62FA19" authorityName="Cambefort" authorityYear="1988" box="[543,667,1401,1427]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bordati">
<emphasis id="EFACEAD6FFFA890A9D1EFAC8FD62FA19" box="[543,667,1401,1427]" italics="true" pageId="12" pageNumber="13">O. bordati</emphasis>
</taxonomicName>
as
<bibRefCitation id="B9494B35FFFA890A9DC6FAC8FB92FA19" author="Hanboonsong Y. &amp; Masumoto K." box="[711,1131,1401,1427]" pageId="12" pageNumber="13" pagination="101 - 114" refId="ref29369" refString="Hanboonsong Y. &amp; Masumoto K. 2000. Dung beetles (Coleoptera, Scarabaeidae) of Thailand, Part 2, Genus Onitis. Elytra, Tokyo 28 (1): 101 - 114." type="journal article" year="2000">Hanboonsong &amp; Masumoto (2000)</bibRefCitation>
mentioned only some morphological characters. This is a new distribution record of the species from
<collectingCountry id="A5CF7654FFFA890A9B83FA2DFB47FA3C" box="[1154,1214,1436,1462]" name="India" pageId="12" pageNumber="13">India</collectingCountry>
and the Indian subcontinent.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,275 @@
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<mods:title id="3A08BCFC7EE06A4F5D8799AB19D8E9BB">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:role id="DABC12CED2BA82BE9B20575A02A9F3FC">
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<mods:namePart id="FA2ED42900FE8BC2A5CBD12B94F4072C">Karimbumkara, Seena Narayanan</mods:namePart>
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</mods:name>
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<mods:namePart id="5A76D85D5CC687E84C8D5F0C8AD4EB0E">Priyadarsanan, Dharma Rajan</mods:namePart>
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<mods:affiliation id="1E7A201E20419C99B5000D23E8ECB919">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="BA26A1746D76E803E7D5F2D101A38A34" type="email">priyan@atree.org</mods:nameIdentifier>
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<mods:typeOfResource id="8C00BEEB8B4F5F209EC1DF0AD55CE0AF">text</mods:typeOfResource>
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<mods:titleInfo id="6E27F1230477B737D48C1AE7A4AB2210">
<mods:title id="76CCBF53004CABAC09F5207696969726">European Journal of Taxonomy</mods:title>
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<mods:part id="4F7D613AA62E92E4B8364E4EE2103010">
<mods:date id="D8A2C994131587FF4AFE4433A4757860">2024</mods:date>
<mods:detail id="B90B7CAC9DCE1AF60B78B05AB8960DFF" type="pubDate">
<mods:number id="A96B8F7BFD31DEE8AE1191BE1606BC64">2024-09-10</mods:number>
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<mods:detail id="8EC8247F80993F34C862849F2E860C9A" type="volume">
<mods:number id="5FB4B1717E4ABA24C71F9DF1B6444B7F">956</mods:number>
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<mods:start id="746CE3D5FFEBE692151E1B0746F04D39">1</mods:start>
<mods:end id="15B52B3BA44F953DC94C4FEF864BDFC7">61</mods:end>
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<mods:location id="A5325953B4E4EB8603F4807B830CE078">
<mods:url id="84E688A1FC73A6FE261C968D0749123A">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287</mods:url>
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<mods:identifier id="F2EAA174E24E6324CC910A37ACB0438D" type="DOI">10.5852/ejt.2024.956.2657</mods:identifier>
<mods:identifier id="15BCC2501697D1F43702DD5D1BC88D14" type="ISSN">2118-9773</mods:identifier>
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<treatment id="557187D2FFFE890C9D72FEB8FABFFCFC" LSID="urn:lsid:zoobank.org:act:7D70D754-D82E-4BCA-A3C9-D42645B4F708" httpUri="http://treatment.plazi.org/id/557187D2FFFE890C9D72FEB8FABFFCFC" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="95C2654FFFFE890E9D72FEB8FB33FECD" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="DD6736C4FFFE890E9D72FEB8FC38FEA9" blockId="8.[361,1226,264,362]" box="[627,961,264,291]" pageId="8" pageNumber="9">
<heading id="862F81A8FFFE890E9D72FEB8FC38FEA9" box="[627,961,264,291]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
<taxonomicName id="1AD84D47FFFE890E9D72FEB8FCA6FEA9" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[627,863,264,291]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bhomorensis" status="sp. nov.">
<emphasis id="EFACEAD6FFFE890E9D72FEB8FCA6FEA9" bold="true" box="[627,863,264,291]" italics="true" pageId="8" pageNumber="9">Onitis bhomorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFFE890E9C67FEB8FC38FEA9" box="[870,961,265,291]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFE890E9E68FE9DFB33FECD" blockId="8.[361,1226,264,362]" box="[361,1226,300,327]" pageId="8" pageNumber="9">
<uri id="A9493AC6FFFE890E9E68FE9DFB33FECD" box="[361,1226,300,327]" pageId="8" pageNumber="9">
urn:lsid:zoobank.org:act:
<uuid id="A97E0C11FFFE890E9D86FE9DFB33FECD" box="[647,1226,300,327]" pageId="8" pageNumber="9">7D70D754-D82E-4BCA-A3C9-D42645B4F708</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFE890E9D40FEFEFC0BFEE3" box="[577,1010,335,362]" pageId="8" pageNumber="9" type="description">
<paragraph id="DD6736C4FFFE890E9D40FEFEFC0BFEE3" blockId="8.[361,1226,264,362]" box="[577,1010,335,362]" pageId="8" pageNumber="9">
<figureCitation id="45E32A41FFFE890E9D40FEFEFD72FEE3" box="[577,651,335,361]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[245,1340,356,1841]" captionTargetId="figure-1@9.[189,1398,265,1856]" captionTargetPageId="9" captionText="Fig. 2. Onitisbhomorensis sp. nov., holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head and prontum. D. Protibia, arrow showing rudimentary spur. E. Metasternum. F. Pygidium, arrow showing longitudinal groove. (Images: AIMB)." pageId="8" pageNumber="9">Figs 2</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9D99FEFEFD2AFEE3" box="[664,723,335,361]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="8" pageNumber="9">6IL</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9DDEFEFEFCFFFEE3" box="[735,774,335,361]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="8" pageNumber="9">9A</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9C13FEFEFCBEFEE3" box="[786,839,335,361]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="8" pageNumber="9">14A</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9C52FEFEFC71FEE0" box="[851,904,335,362]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="8" pageNumber="9">15A</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9C95FEFEFC33FEE3" box="[916,970,335,361]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="8" pageNumber="9">16A</figureCitation>
,
<figureCitation id="45E32A41FFFE890E9CD4FEFEFC0BFEE3" box="[981,1010,335,361]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." pageId="8" pageNumber="9">17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFE890E9FBCFE21FAC4FC0A" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="DD6736C4FFFE890E9FBCFE21FE3FFE20" blockId="8.[189,454,400,426]" box="[189,454,400,426]" pageId="8" pageNumber="9">
<heading id="862F81A8FFFE890E9FBCFE21FE3FFE20" bold="true" box="[189,454,400,426]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EFACEAD6FFFE890E9FBCFE21FE3FFE20" bold="true" box="[189,454,400,426]" pageId="8" pageNumber="9">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFE890E9FBCFE0EFAC4FC0A" blockId="8.[189,1399,447,896]" pageId="8" pageNumber="9">
<emphasis id="EFACEAD6FFFE890E9FBCFE0EFE58FE53" box="[189,417,447,473]" italics="true" pageId="8" pageNumber="9">Onitisbhomorensis</emphasis>
<taxonomicNameLabel id="F49F57ADFFFE890E9EA6FE71FDFBFE50" box="[423,514,448,474]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
is assigned to Group I as its body is completely black and the frontal carina is interrupted. It is similar to
<taxonomicName id="1AD84D47FFFE890E9D14FE52FD01FE76" authorityName="Janssens" authorityYear="1937" box="[533,760,483,509]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFFE890E9D14FE52FD01FE76" box="[533,760,483,509]" italics="true" pageId="8" pageNumber="9">O. punctatostriatus</emphasis>
</taxonomicName>
in having broad elytral striae. It differs from the latter in the nature of pronotal punctures which are stronger in
<taxonomicName id="1AD84D47FFFE890E9C6AFDB7FBDCFDAA" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[875,1061,518,544]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bhomorensis">
<emphasis id="EFACEAD6FFFE890E9C6AFDB7FBDCFDAA" box="[875,1061,518,544]" italics="true" pageId="8" pageNumber="9">O. bhomorensis</emphasis>
</taxonomicName>
than in
<taxonomicName id="1AD84D47FFFE890E9B8BFDB7FA96FDAA" authorityName="Janssens" authorityYear="1937" box="[1162,1391,518,544]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFFE890E9B8BFDB7FA96FDAA" box="[1162,1391,518,544]" italics="true" pageId="8" pageNumber="9">O. punctatostriatus</emphasis>
</taxonomicName>
; basal foveae are more spaced in the former than the latter; metasternal shield slightly convex, granular anteriorly with reddish hairs in
<taxonomicName id="1AD84D47FFFE890E9D2CFDFCFD1CFDEC" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[557,741,588,615]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bhomorensis">
<emphasis id="EFACEAD6FFFE890E9D2CFDFCFD1CFDEC" box="[557,741,588,615]" italics="true" pageId="8" pageNumber="9">O. bhomorensis</emphasis>
</taxonomicName>
while in
<taxonomicName id="1AD84D47FFFE890E9C53FDFCFBCCFDEC" authorityName="Janssens" authorityYear="1937" box="[850,1077,589,615]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFFE890E9C53FDFCFBCCFDEC" box="[850,1077,589,615]" italics="true" pageId="8" pageNumber="9">O. punctatostriatus</emphasis>
</taxonomicName>
it is flat in the middle, with a weak longitudinal groove and with smooth scattered punctures. Clypeal margin truncated, straight in the middle in the former, while it is slightly reflexed in the latter. The presence of a median groove on the pygidium and the rudimentary spur on the protibia make it different from all other Oriental
<taxonomicName id="1AD84D47FFFE890E9A2EFD07FEE0FD79" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EFACEAD6FFFE890E9A2EFD07FA8EFD5A" box="[1327,1399,694,720]" italics="true" pageId="8" pageNumber="9">Onitis</emphasis>
species.
</taxonomicName>
<emphasis id="EFACEAD6FFFE890E9E1EFD68FDFBFD79" box="[287,514,729,755]" italics="true" pageId="8" pageNumber="9">Onitisbhomorensis</emphasis>
, is compared with
<taxonomicName id="1AD84D47FFFE890E9DDCFD68FC7DFD79" authorityName="Biswas" authorityYear="1980" box="[733,900,729,755]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFFE890E9DDCFD68FC7DFD79" box="[733,900,729,755]" italics="true" pageId="8" pageNumber="9">O. assamensis</emphasis>
</taxonomicName>
, a species described from a nearby locality as that of the former by
<bibRefCitation id="B9494B35FFFE890E9EDCFD4DFD71FC9C" author="Biswas S." box="[477,648,764,790]" pageId="8" pageNumber="9" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas (1980)</bibRefCitation>
based on a single specimen.
<taxonomicName id="1AD84D47FFFE890E9CE2FD4CFB64FC9C" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[995,1181,764,791]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bhomorensis">
<emphasis id="EFACEAD6FFFE890E9CE2FD4CFB64FC9C" box="[995,1181,764,791]" italics="true" pageId="8" pageNumber="9">O. bhomorensis</emphasis>
</taxonomicName>
is smaller and the body is reddish black, while
<taxonomicName id="1AD84D47FFFE890E9D0DFC91FD4AFCB3" authorityName="Biswas" authorityYear="1980" box="[524,691,800,826]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFFE890E9D0DFC91FD4AFCB3" box="[524,691,800,826]" italics="true" pageId="8" pageNumber="9">O. assamensis</emphasis>
</taxonomicName>
is greenish. The frontal carina is entire with a tubercle in the former while the carina is interrupted in the latter. The protibia of
<taxonomicName id="1AD84D47FFFE890E9CBFFCF2FB8EFCD6" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[958,1143,834,861]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bhomorensis">
<emphasis id="EFACEAD6FFFE890E9CBFFCF2FB8EFCD6" box="[958,1143,834,861]" italics="true" pageId="8" pageNumber="9">O. bhomorensis</emphasis>
</taxonomicName>
is with a rudimentary spur and the pygidium is with a vertical groove, both these characters are absent in
<taxonomicName id="1AD84D47FFFE890E9B8FFCD7FACFFC0A" authorityName="Biswas" authorityYear="1980" box="[1166,1334,870,896]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFFE890E9B8FFCD7FACFFC0A" box="[1166,1334,870,896]" italics="true" pageId="8" pageNumber="9">O. assamensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFE890E9FBCFC16FDEDFB99" pageId="8" pageNumber="9" type="etymology">
<paragraph id="DD6736C4FFFE890E9FBCFC16FEBDFC4B" blockId="8.[189,324,935,961]" box="[189,324,935,961]" pageId="8" pageNumber="9">
<heading id="862F81A8FFFE890E9FBCFC16FEBDFC4B" bold="true" box="[189,324,935,961]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EFACEAD6FFFE890E9FBCFC16FEBDFC4B" bold="true" box="[189,324,935,961]" pageId="8" pageNumber="9">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFE890E9FBCFC67FDEDFB99" blockId="8.[189,1399,982,1043]" pageId="8" pageNumber="9">
Named after the bridge “Kolia Bhomora” across Brahmaputra the widest river in
<collectingCountry id="A5CF7654FFFE890E9B83FC67FB38FC7A" box="[1154,1217,982,1008]" name="India" pageId="8" pageNumber="9">India</collectingCountry>
, near which the
<typeStatus id="02638866FFFE890E9FBCFC48FF09FB99" box="[189,240,1017,1043]" pageId="8" pageNumber="9">type</typeStatus>
specimen was collected.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFE890E9FBCFB8BFDBCFB67" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="DD6736C4FFFE890E9FBCFB8BFE53FBDE" blockId="8.[189,426,1082,1108]" box="[189,426,1082,1108]" pageId="8" pageNumber="9">
<heading id="862F81A8FFFE890E9FBCFB8BFE53FBDE" bold="true" box="[189,426,1082,1108]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EFACEAD6FFFE890E9FBCFB8BFE53FBDE" bold="true" box="[189,426,1082,1108]" pageId="8" pageNumber="9">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFE890E9FEBFBD8FEA5FB09" blockId="8.[189,1399,1129,1261]" box="[234,348,1129,1155]" pageId="8" pageNumber="9">
<emphasis id="EFACEAD6FFFE890E9FEBFBD8FEA5FB09" bold="true" box="[234,348,1129,1155]" pageId="8" pageNumber="9">
<typeStatus id="02638866FFFE890E9FEBFBD8FEA5FB09" box="[234,348,1129,1155]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFFE890E9FBCFB3DFDBCFB67" blockId="8.[189,1399,1129,1261]" pageId="8" pageNumber="9">
<materialsCitation id="6DB03C99FFFE890E9FBCFB3DFDBCFB67" collectingDate="2012-10-10" collectionCode="NBAIR, NIM" collectorName="Karimbumkara Seena Narayanan &amp; Bangalore" country="India" elevation="74" latitude="26.582407" location="Kolia" longLatPrecision="1" longitude="92.864624" municipality="Tezpur" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Assam" typeStatus="holotype">
<collectingCountry id="A5CF7654FFFE890E9FBCFB3DFEF7FB2C" box="[189,270,1164,1190]" name="India" pageId="8" pageNumber="9">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFFE890E9E33FB3CFEB0FB2D" box="[306,329,1165,1191]" count="1" pageId="8" pageNumber="9" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFFE890E9E5AFB3DFE57FB2C" box="[347,430,1164,1190]" country="India" name="Assam" pageId="8" pageNumber="9">Assam</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFFE890E9EBFFB3CFDE8FB2D" box="[446,529,1165,1191]" pageId="8" pageNumber="9">Tezpur</collectingMunicipality>
,
<location id="D807601FFFFE890E9D23FB3DFD9AFB2C" LSID="urn:lsid:plazi:treatment:557187D2FFFE890C9D72FEB8FABFFCFC:D807601FFFFE890E9D23FB3DFD9AFB2C" box="[546,611,1164,1190]" country="India" latitude="26.582407" longLatPrecision="1" longitude="92.864624" municipality="Tezpur" name="Kolia" pageId="8" pageNumber="9" stateProvince="Assam">Kolia</location>
bhomora;
<date id="A9661004FFFE890E9DECFB3DFC6AFB2C" box="[749,915,1164,1190]" pageId="8" pageNumber="9" value="2012-10-10">
<collectingDate id="B922E9ECFFFE890E9DECFB3DFC6AFB2C" box="[749,915,1164,1190]" pageId="8" pageNumber="9" value="2012-10-10">10 Oct. 2012</collectingDate>
</date>
;
<geoCoordinate id="B8EC5003FFFE890E9CA5FB3AFB9BFB2D" box="[932,1122,1163,1191]" degrees="26" direction="north" minutes="34" orientation="latitude" pageId="8" pageNumber="9" precision="1" seconds="56.66" value="26.582407">26°34ʹ56.66ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFFE890E9B72FB3AFAD7FB2D" box="[1139,1326,1163,1191]" degrees="92" direction="east" minutes="51" orientation="longitude" pageId="8" pageNumber="9" precision="1" seconds="52.65" value="92.864624">92°51ʹ52.65ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFFE890E9FBCFB1EFEC0FB43" box="[189,313,1199,1226]" metricMagnitude="1" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="9" unit="m" value="74.0">
<quantity id="1A209B21FFFE890E9FBCFB1EFF00FB40" box="[189,249,1199,1226]" metricMagnitude="1" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="9" unit="m" value="74.0">74 m</quantity>
a.s.l.
</elevation>
;
<collectorName id="702D5312FFFE890E9E46FB1EFD34FB40" box="[327,717,1199,1226]" pageId="8" pageNumber="9">Karimbumkara Seena Narayanan</collectorName>
leg.; hand-picked, cattle dung;
<collectionCode id="BBC9AE01FFFE890E9B43FB01FB59FB40" box="[1090,1184,1200,1226]" pageId="8" pageNumber="9">NBAIR</collectionCode>
,
<collectorName id="702D5312FFFE890E9BACFB1EFAD0FB43" box="[1197,1321,1199,1225]" pageId="8" pageNumber="9">Bangalore</collectorName>
;
<collectionCode id="BBC9AE01FFFE890E9A34FB1EFA94FB43" box="[1333,1389,1199,1225]" country="France" name="Museum d'histoire naturelle de Nîmes" pageId="8" pageNumber="9" type="Herbarium">NIM</collectionCode>
/
<collectionCode id="BBC9AE01FFFE890E9FBCFB62FEE0FB67" box="[189,281,1235,1261]" pageId="8" pageNumber="9">NBAIR</collectionCode>
/COL/ONIT/H-271023A.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFFE890C9FBCFAA5FABFFCFC" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="DD6736C4FFFE890E9FBCFAA5FEB7FAA4" blockId="8.[189,334,1300,1326]" box="[189,334,1300,1326]" pageId="8" pageNumber="9">
<heading id="862F81A8FFFE890E9FBCFAA5FEB7FAA4" bold="true" box="[189,334,1300,1326]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="EFACEAD6FFFE890E9FBCFAA5FEB7FAA4" bold="true" box="[189,334,1300,1326]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFE890E9FBCFAF2FB3BFAD7" blockId="8.[189,1218,1346,1373]" box="[189,1218,1346,1373]" pageId="8" pageNumber="9">MEASUREMENTS (in mm). TL=18; BW= 10; PL=7; PW=11; EL=10; HL=5; HW=5.1.</paragraph>
<paragraph id="DD6736C4FFFE890E9FEBFA35FE52FA14" blockId="8.[189,1399,1412,2002]" box="[234,427,1412,1438]" pageId="8" pageNumber="9">
<emphasis id="EFACEAD6FFFE890E9FEBFA35FED3FA14" bold="true" box="[234,298,1412,1438]" pageId="8" pageNumber="9">Male</emphasis>
(
<typeStatus id="02638866FFFE890E9E38FA35FE5AFA14" box="[313,419,1412,1438]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="DD6736C4FFFE890C9FBCFA16FE89FE7C" blockId="8.[189,1399,1412,2002]" lastBlockId="10.[189,1399,265,503]" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
Body narrow, elongate oval, reddish black, not very shining, slightly convex. Antennae red with the club reddish yellow, legs reddish black; body devoid of setae on the dorsal side, ventral side covered with reddish hairs except on the middle of metasternum and the abdominal sclerites. Head shining with the clypeus truncate (
<figureCitation id="45E32A41FFFE890E9EF4F9A0FDAAF9A1" box="[501,595,1553,1579]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="8" pageNumber="9">Fig. 9A</figureCitation>
), almost straight anteriorly and at the sides, strongly rugose; clypeal carina present, slightly elevated and curved. The frontal carina is broadly interrupted in the middle with a blunt conical tubercle behind. The vertex punctuation granular, closer in the middle and scattered at the sides appears rugose behind the tubercle. Genae smooth, curved, extended slightly outwards and upwards anteriorly; inner margin slightly elevated and continues behind as a posterior carina. Pronotum moderately convex, sides smoothly margined, except the base which is slightly angulate in the middle; with strong close punctures, closer between the basal foveae; uneven in the middle; deeper and closer anteriorly. Pronotal front angles not very sharp, almost right-angled, the sides straight anteriorly, rounded in the middle, bisinuate behind; hind angles obsolete. A pair of deep basal foveae present at the base on either side of the middle separated by twice its length. Lateral foveae smooth with small punctures in the periphery. Elytra finely and broadly striate, striae devoid of punctures; intervals flat, opaque; sutural interval convex, shining with visible fine scattered punctures. The lateral carinae straight, with minute visible punctures. Pygidium (
<figureCitation id="45E32A41FFFE890E9D13F806FD90F85B" box="[530,617,1975,2001]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[245,1340,356,1841]" captionTargetId="figure-1@9.[189,1398,265,1856]" captionTargetPageId="9" captionText="Fig. 2. Onitisbhomorensis sp. nov., holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head and prontum. D. Protibia, arrow showing rudimentary spur. E. Metasternum. F. Pygidium, arrow showing longitudinal groove. (Images: AIMB)." pageId="8" pageNumber="9">Fig. 2F</figureCitation>
) without hair or setae, slightly shining with imperceptible scattered punctures. A smooth median longitudinal groove extending from base to almost the middle is present. Metasternal shield shining, (
<figureCitation id="45E32A41FFFC890C9D05FE9DFDA2FECC" box="[516,603,300,326]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[245,1340,356,1841]" captionTargetId="figure-1@9.[189,1398,265,1856]" captionTargetPageId="9" captionText="Fig. 2. Onitisbhomorensis sp. nov., holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head and prontum. D. Protibia, arrow showing rudimentary spur. E. Metasternum. F. Pygidium, arrow showing longitudinal groove. (Images: AIMB)." pageId="10" pageNumber="11">Fig. 2E</figureCitation>
) with minute punctures in the middle; anterior part of the metasternal shield slightly convex, with uneven granules and reddish hairs. Sides of the metasternum closely and evenly granular, covered with smooth short reddish hairs. Protibia slender, bears four external teeth, the tip not very long or curved, with a rudimentary spur near the tip (
<figureCitation id="45E32A41FFFC890C9CBBFE27FBE5FE3A" box="[954,1052,406,432]" captionStart="Fig" captionStartId="9.[189,232,1928,1954]" captionTargetBox="[245,1340,356,1841]" captionTargetId="figure-1@9.[189,1398,265,1856]" captionTargetPageId="9" captionText="Fig. 2. Onitisbhomorensis sp. nov., holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. A. Habitus, dorsal view. B. Habitus, ventral view. C. Head and prontum. D. Protibia, arrow showing rudimentary spur. E. Metasternum. F. Pygidium, arrow showing longitudinal groove. (Images: AIMB)." pageId="10" pageNumber="11">Figs 2D</figureCitation>
,
<figureCitation id="45E32A41FFFC890C9B29FE27FBA6FE3A" box="[1064,1119,406,432]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." pageId="10" pageNumber="11">16A</figureCitation>
); teeth short, triangular, placed almost equidistantly. The inner margins of femur and tibia fringed. Femur without protrusion on any of the legs.
</paragraph>
<caption id="89A7664CFFFF890F9FBCF839FB19F862" pageId="9" pageNumber="10" startId="9.[189,232,1928,1954]" targetBox="[245,1340,356,1841]" targetPageId="9">
<paragraph id="DD6736C4FFFF890F9FBCF839FB19F862" blockId="9.[189,1398,1928,2025]" pageId="9" pageNumber="10">
<emphasis id="EFACEAD6FFFF890F9FBCF839FEF7F828" bold="true" box="[189,270,1928,1954]" pageId="9" pageNumber="10">Fig. 2.</emphasis>
<emphasis id="EFACEAD6FFFF890F9E19F839FE06F828" box="[280,511,1928,1954]" italics="true" pageId="9" pageNumber="10">Onitisbhomorensis</emphasis>
<taxonomicNameLabel id="F49F57ADFFFF890F9D0BF839FD9EF828" box="[522,615,1928,1954]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A.
<emphasis id="EFACEAD6FFFF890F9BECF839FAFBF828" bold="true" box="[1261,1282,1928,1954]" pageId="9" pageNumber="10">A</emphasis>
. Habitus, dorsal view.
<emphasis id="EFACEAD6FFFF890F9E54F81AFE90F84F" bold="true" box="[341,361,1963,1989]" pageId="9" pageNumber="10">B</emphasis>
. Habitus, ventral view.
<emphasis id="EFACEAD6FFFF890F9D86F81AFD65F84F" bold="true" box="[647,668,1963,1989]" pageId="9" pageNumber="10">C</emphasis>
. Head and prontum.
<emphasis id="EFACEAD6FFFF890F9C9BF81AFC56F84F" bold="true" box="[922,943,1963,1989]" pageId="9" pageNumber="10">D</emphasis>
. Protibia, arrow showing rudimentary spur.
<emphasis id="EFACEAD6FFFF890F9FFCF87FFEE8F862" bold="true" box="[253,273,1998,2024]" pageId="9" pageNumber="10">E</emphasis>
. Metasternum.
<emphasis id="EFACEAD6FFFF890F9EC9F87FFE23F862" bold="true" box="[456,474,1998,2024]" pageId="9" pageNumber="10">F</emphasis>
. Pygidium, arrow showing longitudinal groove. (Images: AIMB).
</paragraph>
</caption>
<paragraph id="DD6736C4FFFC890C9FBCFDAFFD7EFDF4" blockId="10.[189,1399,541,639]" pageId="10" pageNumber="11">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFFC890C9EAFFDAFFDDAFDBD" box="[430,547,541,568]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." pageId="10" pageNumber="11">Fig. 6IL</figureCitation>
). LP=3, Lp=2.5; BP=1.4, BpB =0.7, BpT=0.4. Parameres almost same length as phallobase, broader at the base and further tapers become slender and slightly curved at the tip. Phallobase slightly broader at the base.
</paragraph>
<paragraph id="DD6736C4FFFC890C9FEBFD17FEBCFD4A" blockId="10.[189,325,678,739]" box="[234,325,678,704]" pageId="10" pageNumber="11">
<emphasis id="EFACEAD6FFFC890C9FEBFD17FEBCFD4A" bold="true" box="[234,325,678,704]" pageId="10" pageNumber="11">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCFD78FEC0FD69" blockId="10.[189,325,678,739]" box="[189,313,713,739]" pageId="10" pageNumber="11">Unknown.</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCFCBBFEE4FCAE" blockId="10.[189,285,778,804]" box="[189,285,778,804]" pageId="10" pageNumber="11">
<heading id="862F81A8FFFC890C9FBCFCBBFEE4FCAE" bold="true" box="[189,285,778,804]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="EFACEAD6FFFC890C9FBCFCBBFEE4FCAE" bold="true" box="[189,285,778,804]" pageId="10" pageNumber="11">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFFC890C9FBCFC88FABFFCFC" blockId="10.[189,1398,825,886]" pageId="10" pageNumber="11">
The specimen was collected from cattle dung on the river bed of the Brahmaputra, where the vegetation is dry seasonal grasslands, typical of Brahmaputra flood plains dominated by
<emphasis id="EFACEAD6FFFC890C9B4BFCEDFAF9FCFC" box="[1098,1280,860,886]" italics="true" pageId="10" pageNumber="11">Ziziphusjujuba</emphasis>
trees.
</paragraph>
</subSubSection>
</treatment>
</document>