diff --git a/data/55/71/87/557187D2FFC589329D91FC85FB35F9B6.xml b/data/55/71/87/557187D2FFC589329D91FC85FB35F9B6.xml new file mode 100644 index 00000000000..bb1e21c117e --- /dev/null +++ b/data/55/71/87/557187D2FFC589329D91FC85FB35F9B6.xml @@ -0,0 +1,270 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis visthara + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +0E89DF71-6A9D-4E88-8C49-84281CC84C2A + + + + + +Figs 8M–P +, +9E +, +12E–F +, +14E +, +15E +, +16E +, +20 + + + + + +Differential diagnosis + + + + +Onitis visthara + +sp. nov. +belongs to Group II; it shows affinity to + +Onitis philemon + +, + +O. kethai + +sp. nov +and + +O. singhalensis + +as the metasternum is longitudinally grooved anteriorly and the clypeal margin feebly emarginate. The pronotum of +Onitisvisthara +is more convex and wider and has a smooth median line similar to that in + +O. singhalensis + +but absent in + +O. philemon + +and + +O. kethai + +. +Onitisvisthara +can be differentiated from + +O. singhalensis + +by the pronotal front angle which is sharp in the former while blunt or rather rounded in the latter (compare +Fig. 9E–F +). The clypeus and vertex in + +O. visthara + +are rugose while they are closely granular in + +O. singhalensis + +. The genae of the former is gently rounded while that of the latter is sharply rounded. Profemur in the male of + +O. visthara + +is without tooth while there is a strong tooth towards the end of its anterior edge in the male of + +O. singhalensis + +. The mid femur in the male of + +O. visthara + +is without a tooth near the middle of its posterior edge, but with two teeth at the extremity, a sharp one on the lower edge and a blunt one above; while in the male of + +O. singhalensis + +, the middle femur has a strong tooth near the middle of its posterior edge and another tooth at the extremity. The middle tibia in male + +O. visthara + +is slender at the base and they curve and angularly dilate before the middle; in + +O. singhalensis + +, the middle tibia is slender at the base, less curved, abruptly, angularly dilated before the middle (compare 16E–F). + + + + + +Type material + + + + +Holotype + + + + +INDIA +• +♂ +; +Karnataka +, +Hesaraghatta +, +Bangalore +, hand-picked from cattle dung; + +4 Jan. 2015 + +, +Mohammed Ashraf K. +leg.; specimen deposited at +NBAIR +, +NIM +/ +NBAIR +/COL/ONIT/H-271023C. + + + + +Paratype + + + +COUNTRY • + +♂ +; same collection details as +holotype +; +AIMB +/Co/Sc1000789 + +. + + + + + +Etymology + + + +The pronotal and elytral measurements of this species are almost the same, giving it a broader rectangular appearance while comparing with other species, thus the name + +visthara + +which means ‘expanse’ in Sanskrit. + + + + + +Description + + +MEASUREMENTS (in mm). TL= 17–18; BW =9–10; PL=8–9; PW=9.5–10.5; EL=8–9.5; HL=4–4.5; HW=5–5.5. + +Male +( +Holotype +) + + +Brownish black, not shining, pronotum bronzy black, elytra bronzy red, ventral side blackishred; antennae and mouth reddish; oval, not very convex. Clypeus ( +Fig. 14E +) rugulose, anterior margin parabolic, slightly straight in the middle; clypeal carina not much elevated, bears a few punctures; clypeo-frontal carina not widely interrupted, the tubercle behind blunt, rounded and closer to carina, bears a few punctures; vertex rugulose in the middle and sides more or less smooth with a few scattered granules; genae broad and smooth, slightly reflexed anteriorly with a few scattered punctures. Pronotum strongly and fairly closely punctured, scattered in the middle, more strongly and closely pitted anteriorly, lightly and closely at the base, with a slight median groove starting above the basal foveae and extending to anterior margin as a thin smooth line; front angles moderately sharp, almost right angled, hind angle obsolete, lateral margin straight anteriorly, strongly curved or rounded in the middle and bisinuate behind; base of pronotum bluntly angulate; basal foveae rounded and concave. Elytra moderately shining, lightly and broadly striate, striae smooth, intervals almost flat, strongly and closely punctured; sutural interval convex and slightly elevated than the rest, finely and deeply punctured. Pygidium not shining, lightly and uniformly punctured. Metasternum with deep broad longitudinal groove; metasternal shield smooth in the middle with fine scattered punctures and with strong close aciculate punctures anterio-laterally; sides of metasternum with setigerous granules. Femora smooth with some punctures and hairs on anterior part near the tibial joint. Protibia with four teeth, with the tip extended to a curved process; teeth very short, blunt, the terminal tooth trapezoid. Profemur without teeth. Mid femur has a pair of sharp teeth near the end of the lower edge, and a blunt curved process towards the middle; hind trochanter with a small blunt tooth. + + +AEDEAGUS (in mm) ( +Fig. 8M–P +). LP=3, Lp=2.5; BP=0.75, BpB=0.5, BpT=0.25. Phallobase slightly curved, longer than parameres, the latter curved at the tip, with an opening on the ventral side at the point of articulation with the phallobase. The base of the opening is with a cup-shaped extension which is curved inwards. + + +Female + +Unknown. + + +Habitat + + +The specimens were collected from cattle dung in open grassland near Hesaraghatta lake, the vegetation of which is undergoing rapid changes due to human activities and ecological succession. + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFD0892F9D34FC85FB77FB1B.xml b/data/55/71/87/557187D2FFD0892F9D34FC85FB77FB1B.xml new file mode 100644 index 00000000000..b46b51000e0 --- /dev/null +++ b/data/55/71/87/557187D2FFD0892F9D34FC85FB77FB1B.xml @@ -0,0 +1,495 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis punctatostriatus +Janssens, 1937 + + + + + + +Figs 6M–P +, +9B +, +10F +, +14B +, +15B +, +16B +, +19 + + +Onitispunctatostriatus +Janssens 1937: 19 +(original description). + + + + + + +Onitis punctatostriatus + +– + +Balthasar 1963: 35 + +(monograph). — + +Bezděk & Krell 2006: 159 + +(distribution). — Siddiqui +et. al. +2014: 300 (distribution). — + + +Gupta +et al. +2015: 1037 + + +(list). — + +Bezděk 2016: 179 + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + +Onitispunctatostriatus +is similar to + +O. subopacus + +in having frontal carina broadly interrupted and the metasternum smooth and shining behind with scattered punctures. It differs from the latter in having deep and broad elytral striae and the pronotum being strongly ocellate punctured. This species is reported for the first time from the state of +Assam +in northeastern +India +. + + + + + +Type material examined + + + + +Holotype + + + + +INDIA +• +♂ +; “Darjeeling 1904, +R.P. Verschraeghen +, cf. Mem. Mus. Hist. Nat. Belg., 2: Serie, fasc. 11, 1937, p. 48, Ex. Typis, +A. Janssens +rev., 1936: + +Onitis punctatostriatus +Janss., J. J. Gillet + +det., vend.”; +IRSNB +. + + + + +Paratype + + + + +INDIA +• +♀ +; “ + +Onitisspinipes +Drury R.M.H.N. Belg. + +10. 640, +spinipes +Drury; +J. Gillet +det. 1923, Ex-Typis”; label data same as for male; +IRSNB + +. + + +Additional material studied +( +10 specimens +) + + + +INDIA +• +2 ♂♂ +, +2 ♀♀ +; +Assam +, +Majuli Island +; hand-picked, cow dung; +Rajkamal Goswami +leg.; +AIMB +/ +Co +/ +Sc +1000492 to 1000495 + +• + +♂ +; +Arunachal Pradesh +, +Pasighat +, community forest, +Rani Village +; +27°58ʹ8.8644ʺ N +, +95°19ʹ17.1804ʺ E +; elev. + +127 m +a.s.l. + +; + +30 Sep. 2018 + +; +cow dung baited pitfall trap +; +AIMB +/ +Co +/ +Sc +1000496 + +• + +♀ +; community forest, +Rani Village +; +27°58ʹ10.6068ʺ N +, +95°19ʹ26.1156ʺ E +; elev. + +131 m +a.s.l. + +; + +30 Sep. 2018 + +; cow dung; +baited pitfall trap +; +AIMB +/ +Co +/ +Sc +1000497 + +• + +♂ +, +2 ♀♀ +; community forest, +Rani Village +; +27°58ʹ8.8644ʺ N +, +95°19ʹ17.1804ʺ E +; elev. + +127 m +a.s.l. + +; + +29 Sep. 2018 + +; hand-picked from cow dung, +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000498 to 1000500 + +• + +♀ +; +Tripura +, +Trishna Wildlife Sanctuary +, +South +Tripura +; +23°16ʹ39.9216ʺ N +, +91°24ʹ16.3152ʺ E +; +Forest +, + +18 Feb. 2019 + +; +cow dung baited pitfall trap +; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000501 + +. + + + + +Fig. 15. +Pronotal basal foveae of male specimens. +A +. +Onitisbhomorensis +sp. nov. +, holotype, NIM/ NBAIR/COL/ONIT/H-271023A. +B +. + +O. punctatostriatus +Janssens, 1937 + +, AIMB/Co/Sc1000492. +C +. + +O. kethai + +sp. nov. +, holotype, NIM/NBAIR/COL/ONIT/H-271023B. +D +. + +O. philemon +Fabricius, 1801 + +, AIMB/Co/Sc1000317. +E +. + +O. visthara + +sp. nov. +, holotype, NIM/NBAIR/COL/ONIT/H-271023C. +F. + +O. singhalensis +Lansberge, 1875 + +, AIMB/Co/Sc1000684. (Images: AIMB). + + + + +Fig 16. A +. Protibia of + +Onitis bhomorensis + +sp. nov. +showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. +B +. Anterior tip of protibia of + +O. punctatostriatus +Janssens, 1937 + +, AIMB/Co/Sc1000492. +C +. Mid-leg of + +O. kethai + +sp. nov +, NIM/NBAIR/COL/ONIT/H-271023B. +D +. Mid-leg of + +O. philemon +Fabricius, 1801 + +, AIMB/Co/Sc1000317. +E +. Mid-leg of + +O. visthara + +sp. nov +, NIM/NBAIR/COL/ONIT/H- 271023C. +F +. Mid-leg of + +O. singhalensis +Lansberge, AIMB + +/Co/Sc1000684. (Images: AIMB). + + + + + +Description + + +MEASUREMENTS (in mm). TL=17.5–23; BW= 9–12.5. + +Male + +Black or reddish black, moderately shining; clypeal margin slightly reflexed; frontal carina broadly interrupted in the middle with an anterior short curved clypeal carina and transverse tubercle behind; pronotum strongly punctate anteriorly, basal foveae rounded, closer, lateral borders of pronotum slightly concave before anterior angles; elytral striae broad, shallow, interval imperceptibly punctured; pygidium flat, opaque; metasternum flat in the middle with an anterior weak longitudinal groove, metasternal shield smooth with scattered punctures, sides of the metasternum punctured with long rusty hairs. Clypeus evenly rugulose, pronotum with a broad smooth longitudinal median line, mid legs toothed, trochanters of mid leg long while that of hind legs serrated. + +AEDEAGUS (in mm) ( +Fig. 6M–P +). LP=3, Lp= 2.5; BP=1.25, BpB=0.5, BpT =0.4.Parameres longer than the phallobase, symmetrical, straight above, rounded beneath and curved at the tip. + + +Female + +Clypeus parabolic, closely rugose, pronotal punctures stronger and closer than in male, protibia with four strong teeth. + + + + +Distribution + + + +India +: +Arunachal Pradesh +, +Assam +, +Tripura +, +West Bengal +. Other countries: +Pakistan +( +Bezděk & Krell 2006 +; + +Siddiqui +et al. +2014 + +; + +Gupta +et al. +2015 + +; +Bezděk 2016 +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFDA892B9D65FDD4FC1FFA7B.xml b/data/55/71/87/557187D2FFDA892B9D65FDD4FC1FFA7B.xml new file mode 100644 index 00000000000..81234f2a195 --- /dev/null +++ b/data/55/71/87/557187D2FFDA892B9D65FDD4FC1FFA7B.xml @@ -0,0 +1,706 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis subopacus +Arrow, 1931 + + + + + + +Figs 12C +, +13A–D +, +20 + + +Onitissubopacus +Arrow, 1931: 395 +(original description). + + + + + + +Onitis subopacus + +– + +Balthasar 1935: 94 + +(monograph); 1963: 53 (monograph). — + +Janssens 1937: 51 + +(revision). — + +Biswas & Chatterjee 1986a: 67 + +(distribution). — + +Chandra 2000: 360 + +(distribution); 2005: 150 (distribution). — + +Mittal 2005: 46 + +(list). — + +Chandra & Ahirwar 2005: 149 + +(distribution). — + +Bezděk & Krell 2006: 159 + +(distribution). — + + +Sabu +et al. +2006: 5 + + +(list); 2011: 34 (list). — + +Chandra & Ahirwar 2007: 276 + +(distribution). — + +Sewak 2009b: 60 + +(key, diagnosis, distribution). — + + +Thakare +et al. +2012: 75 + + +(distribution). — + +Chandra & Gupta 2011: 274 + +(distribution); 2012a: 889 (list) Chandra +et al. +2012a: 52 (list); 2012b: 33 (distribution). — + +Chandra & Gupta 2013a: 349 + +(diagnosis, distribution); 2013b: 4667 (distribution). — Chandra +et al. +2011: 67 (distribution). — + + +Gupta +et al. +2014: 230 + + +(distribution); 2015: 1037 (list, distribution). — + + +Siddiqui +et al. +2014: 300 + + +(distribution). — + +Bezděk 2016: 179 + +(distribution). — + + +Ghosh +et al. +2020: 239 + + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + +Onitisphilemon +– Lansberge (nec Fabricius) 1875: 133. — +Boucomont 1914: 336 +(distribution). — +Boucomont & Gillet 1921: 19 +(distribution). + + + + + +Differential diagnosis + + + +Onitissubopacus +belongs to Group III and closely resembles + +O. virens + +of the same group but differs in that the former has a less strongly punctured pronotum and broadly interrupted frontal carina. + + + + + +Type material examined + + + + +Holotype + + + + +SRI LANKA +• +♂ +; “Kalupahani, Haldumulle Ceylon, 1904–171, + +Onitis subopacus + +type +, Arrow (aedeagus point-mounted and attached with specimen)”; +BMNH +(E) 1237174. + + + +Additional material examined +( +39 specimens +) + + + +INDIA +• +♀ +; “ +Kashmir +; +Haran Pltn. +5500, +Sindh Valley +, + +18.v.1928 + +, +C.F.C. Beeson +; ex. coll., Dehradun, B.M. 1928-472; + +Onitissubopacus + +co-type, +Arrow +”; +BMNH +(E) 1237175 + +• + +♂ +; “ +Roorkee +, +Arrow Collection +, +Det. G.J. Arrow +”; +ZSIM + +• + +♀ +; +ZSIM + +• + +4 ♂♂ +, +2 ♀♀ +; +Kerala +, +Shendurney Wildlife Sanctuary +, +Rockwood +grassland; +08°51ʹ41.8ʺ N +, +77°08ʹ00.5ʺ E +; elev. + +839 m +a.s.l. + +; + +8 May 2009 + +; +open cow dung bait traps +; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000711 to 1000716 + +• + +♀ +; +Kerala +, +Shendurney Wildlife Sanctuary +, +Kattilappara +evergreen forest; +08°54ʹ45.6ʺ N +, +77°05ʹ45.3ʺ E +; elev. + +566 ft + +, + +4 Dec. 2009 + +; +AIMB +/ +Co +/ +Sc +1000717 + +• + +2 ♂♂ +, +5 ♀♀ +; +Kerala +, +Shendurney Wildlife Sanctuary +; +Kattilappara +moist deciduous forest; +08°54ʹ54.7ʺ N +, +77°06ʹ28.7ʺ E +; elev. + +185 m +a.s.l. + +; + +3 Dec. 2009 + +; +open cow dung bait traps +; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000718 to 1000724 + +• + +9 ♂♂ +, +6 ♀♀ +; +Kerala +, +Shendurney Wildlife Sanctuary +, +Kattilapara +degraded forest; +08°55ʹ05.6ʺ N +, +77°05ʹ44.7ʺ E +; elev. + +207 m +a.s.l. + +; + +3 Dec. 2009 + +; +open cow dung bait traps +; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000725 to 1000738 + +• + +♂ +; +Kerala +, +Shendurney Wildlife Sanctuary +, +Pandimotta +montane evergreen; +08°49ʹ39.1ʺ N +, +77°13ʹ01.9ʺ E +; elev. + +1234 m +a.s.l. + +; + +29 Nov. 2009 + +; +open cow dung bait traps +; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000739 + +• + +♂ +; +Karnataka +, +Nagarhole National Park +, +Sungathakatte +dry deciduous forest; +11°58ʹ27.12″ N +, +76°11ʹ8.16″ E +; +open cow dung bait traps +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000740 + +• + +♀ +; +Sarathi Nulla +dry deciduous forest; +12°3ʹ48.24″ N +, +76°10ʹ40.8″ E +; +open cattle dung baited trap +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000741 + +• + +♀ +; +Tripura +, +Eden of Bison +, +South +Tripura +, forest; + +18 Feb. 2019 + +; hand-picked from cow dung; +Harsha Malhotra +leg.; +AIMB +/ +Co +/ +Sc +1000742 + +• + +2 ♂♂ +, +♀ +; +Nagaland +, +Singphan Elephant Reserve +, +Forest +; + +24Apr. 2021 + +; hand-picked from cow dung; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000743 to 1000745 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=16–23, BW= 8–12. + +Male + + +Black with a slight metallic lustre; clypeus elliptical, rugulose, vertex granular, more shining; frontal carina widely interrupted in the middle with a conical tubercle behind and a curved clypeal carina in front; pronotum not strongly but rather closely punctured, with an incomplete smooth median longitudinal line, basal foveae deep; pygidium opaque scarcely punctured; elytra finely striate with sutural, 3 +rd +and 5 +th +intervals slightly convex; metasternal shield smooth and shining, the front angles and sides of metasternum closely clothed with reddish hairs. Protibia toothed, mid femur lobed near middle with tooth on posterior edge, trochanter of the hind leg sharply toothed. + + +AEDEAGUS (in mm) ( +Fig. 13A–D +). LP=3, Lp=2.5; BP=1.25, BpB= 0.5, BpT =0.4. Phallobase longer than parameres. Parameres curved outwards at the base and then curved inwards with the tip pointed. + + +Female + +Protibia toothed, middle and hind legs not toothed. + + + + +Distribution + + + +India +: +Arunachal Pradesh +, +Assam +, +Bihar +, +Chhattisgarh +, +Gujarat +, +Haryana +, +Himachal Pradesh +, +Jammu and Kashmir +, +Jharkhand +, +Karnataka +, +Kerala +, +Madhya Pradesh +, +Maharashtra +, +Meghalaya +, +Nagaland +, New +Delhi +, Punjab, +Rajasthan +, +Tamil Nadu +, +Tripura +, +Uttarakhand +, +Uttar Pradesh +, +West Bengal +. Other countries: +Afghanistan +, +Bangladesh +, +Cambodia +, +China +, +Indonesia +, Malay Peninsula, +Myanmar +, +Nepal +, +Pakistan +, +Sri Lanka +, +Thailand +, +Vietnam +. ( +Biswas & Chatterjee 1986a +; +Chandra 2000 +, 2005; +Chandra & Ahirwar 2005 +, +2007 +; +Bezděk & Krell 2006 +; + +Sabu +et al. +2006 + +, +2011 +; +Sewak 2009b +; + +Thakare +et al. +2012 + +; Chandra & Gupta 2012a, 2013a, 2013b; Chandra +et al. +2012a, 2012b; + +Siddiqui +et al. +2014 + +; + +Gupta +et al. +2014 + +, +2015 +; +Bezděk 2016 +; + +Ghosh +et al. +2020 + +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFDB89359D69F987FE3FFD64.xml b/data/55/71/87/557187D2FFDB89359D69F987FE3FFD64.xml new file mode 100644 index 00000000000..7bb2088d38a --- /dev/null +++ b/data/55/71/87/557187D2FFDB89359D69F987FE3FFD64.xml @@ -0,0 +1,835 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis virens +Lansberge, 1875 + + + + + + +Figs 12D +, +13E–H +, +20 + + +Onitisvirens +Lansberge, 1875: 135 +(original description). + + +Onitisamplectens +Lansberge, 1875: 136 +. + + + + + + +Onitis virens + +– + +Boucomont & Gillet 1921: 19 + +(distribution). — + +Arrow 1931: 396 + +(key, description). — + +Balthasar 1935: 52 + +(monograph); 1963: 40 (monograph). — + +Paulian 1945: 144 + +(revision). — + +Biswas & Chatterjee 1991: 260 + +(distribution). — + +Chandra 2000: 360 + +(distribution). — + +Mittal 2005: 46 + +(list). — + +Chandra & Ahirwar 2005: 149 + +(distribution); 2007: 276 (distribution). — + +Bezděk & Krell 2006: 159 + +(distribution). — + +Sewak 2009a: 60 + +(key, diagnosis, distribution), 2009b: 60 (key, diagnosis, distribution). — + + +Sabu +et al. +2011: 34 + + +(list). — Chandra +et al. +2012a: 52 (distribution). — + +Karimbumkara & Rajan 2013: 175 + +(list). — + + +Gupta +et al. +2015: 1037 + + +(list, distribution). — + + +Siddiqui +et al. +2014: 301 + + +(distribution). — + +Bezděk 2016: 180 + +(distribution). — + + +Ghosh +et al. +2020: 239 + + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + +Onitisvirens +belongs to Group III and closely resembles + +O. subopacus + +of the same group but differs from the latter in having a strongly punctured pronotum and clypeo-frontal carina which is narrowly interrupted. + + +Also see remarks under +Onitissubopacus +. + + + + + +Type material examined + + + + +Syntypes + + + +UNKNOWN TYPE LOCALTY • + +♀ +; +“Onitisvirens +Lansb., +G. J. Arrow +det.; +A. Janssens +vid., 1936: + +ONITIS virens +Lansb. + +; +Syntype +; det +Van +Lansberge. +Onitisamplectens +Lansb.; + +Amplectens + +Type Lsb. Mercara; +Onitisvirens +Lansb.Révis.Arrow 1927; Ex-Typis de + +amplectens +Lansb. + +; +Collection E. Candeze +: 109”; +IRSNB + +• + +♀ +; “ +A. Janssens +vid., 1936: + +ONITIS virens +Lansb. + +; cf. Ann. Soc. Ent. Belg. Xviii. 1875, p.135; +Syntype +; Ex-Typis de +amplectens Lansb. +; AmplectensInd br Lansb.; Det. +Van +Lansberge + +Onitis amplectens +Lansb. + +; +Collection E. Candeze +”; +IRSNB + +. + + +Additional material examined +( +47 specimens +) + + + +INDIA +• +♂ +; “ + +Onitis virens +Lansb., Compared + +with +type +, G.J.A.”; +BMNH +(E) 1237153 + +• + +♀ +; “ +Gopaldhara +, +Rungbong Vall. +, +Sikkim +, +H.Stevens. +, 1916-218; +BMNH +(E)1237154” + +• + +♂ +; “ +Nerbuda Survey +, +Harai +, +Bewa State +, + +2700 ft. + +; + +23 Feb. 1927 + +” + +• + +♀ +; +ZSIM + +• + +2 ♂♂ +, +2 ♀♀ +; +Karnataka +, +Biligiri Rangaswamy Tiger Reserve +, +Budipadaga +scrub; +11°50ʹ16.8″ N +, +77°4ʹ39.18″ E +; + +14 May 1999 + +; +open cow dung bait traps +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000746 to 1000749 + +• + +8 ♂♂ +, +5 ♀♀ +; +Kanneri +moist deciduous forest; +11°53ʹ43.62″ N +, +77°8ʹ9.48″ E +; + +22 Dec. 1999 + +; +open cow dung bait traps +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000750 to 1000762 + +• + +2 ♂♂ +, +Purani +dry deciduous forest; +12°1ʹ55.2″ N +, +77°7ʹ52.68″ E +; + +11 May 1999 + +; +open cow dung bait traps +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000763 to 1000764 + +• + +♂ +; +Kerala +, +Shendurney Wildlife Sanctuary +, +Kattilapara +degraded forest; +08°55ʹ05.6ʺ N +, +77°05ʹ44.7ʺ E +; elev. + +207 m +a.s.l. + +; + +3 Dec. 2009 + +; +open cow dung bait traps +; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000765 + +• + +2 ♂♂ +, +2 ♀♀ +; +Arunachal Pradesh +, +Pasighat +, community forest, +Rani Village +; +27°58ʹ8.8644ʺ N +, +95°19ʹ17.1804ʺ E +; elev. + +127 m +a.s.l. + +; + +30 Sep. 2018 + +; +cow dung baited pitfall trap +; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000766 to 1000769 + +• + +6 ♂♂ +, +9 ♀♀ +; agricultural field, +Rani Village +; +27°58ʹ10.6068ʺ N +, +95°19ʹ26.1156ʺ E +, elev. + +131 m +a.s.l. + +; + +30 Sep. 2018 + +; hand-picked from cow dung; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000770 to 1000784 + +• + +♂ +, +2 ♀♀ +; community forest, +Rani Village +; +27°58ʹ9.0156ʺ N +, +95°19ʹ18.9804ʺ E +; elev. + +133 m +a.s.l. + +; + +30 Sep. 2018 + +; +cow dung baited pitfall trap +; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000785 to 1000787 + +• + +♀ +; +Nagaland +, +Bongkolong village +, near +Intanki National Park +, open ground; + +15 Apr. 2021 + +; hand-picked from cow dung; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000788 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=17–23, BW= 9.5–13. + +Male + +Black with a very feeble metallic lustre; clypeus elliptical, margin entire, closely and finely rugulose, vertex rugulose and more shining, frontal carina interrupted by a very short medial gap, with a conical tubercle behind and an anterior short transverse clypeal carina; pronotum fairly closely and strongly punctured, with an anterior narrow smooth median line and slight narrow groove behind; elytra finely striate with intervals flat and imperceptibly punctured, the sutural interval slightly elevated and shining; pygidium flat, smooth and feebly punctured; metasternal shield smooth and shining behind, front angles and sides of metasternum clothed with red hairs. Protibia slender, curved, mid femur and hind trochanter toothed. + + +Fig. 17. +Map showing the distribution of +Onitisassamensis +Biswas, 1980 +; + +O. bhomorensis + +sp. nov. +; + +O. bordati +Cambefort, 1988 + +; + +O. brahma +Lansberge, 1875 + +and + +O. castaneus +Redtenbacher, 1848 + +in the Indian Subcontinent. + + + + +Fig. 18. +Map showing the distribution of + +Onitis crassus +Sharp, 1875 + +; + +O. +excavatus +Arrow, 1931 + +; + +O. falcatus +(Wulfen, 1789) + +; + +O. feae +Felsche, 1907 + +; + +O. humerosus +( +Pallas, 1771 +) + +and + +O. kethai + +sp. nov. +in the Indian Subcontinent. + + + + +Fig. 19. +Map showing the distribution of +Onitislama +Lansberge, 1875 +; + + +O. naviauxi +Cambefort + +, 1988 + +; + +O. philemon +Fabricius, 1801 + +; + +O. punctatostriatus +Janssens, 1937 + +and + +O. singhalensis +Lansberge, 1875 + +in the Indian Subcontinent. + + + + +Fig. 20. +Map showing the distribution of +Onitissiva +Gillet, 1911 +; + +O. subopacus +Arrow, 1931 + +; + +O. virens +Lansberge, 1875 + +and + +O. visthara + +sp. nov. +in the Indian Subcontinent. + + + +AEDEAGUS (in mm) ( +Fig. 13E– H +). LP=3.5, Lp=3; BP=1.6, BpB=0.7, BpT=0.5. Phallobase slightly longer than the paramere, both straight below and sharp at the tip. + + +Female + +Protibia short, broad with four stout teeth, the mid and hind femur without teeth. + + + + +Distribution + + + +India +: +Arunachal Pradesh +, +Assam +, +Bihar +, +Chattisgarh +, +Gujarat +, +Haryana +, +Himachal Pradesh +, +Karnataka +, +Kerala +, +Madhya Pradesh +, +Maharashtra +, +Meghalaya +, +Nagaland +, New +Delhi +, +Odisha +, Punjab, +Rajasthan +, +Sikkim +, +Tamil Nadu +, +Tripura +, +Uttarakhand +, +Uttar Pradesh +, +West Bengal +. Other countries: +China +, +Bangladesh +, +Laos +, +Myanmar +, +Nepal +, +Pakistan +, +Thailand +, +Vietnam +( +Chandra 2000 +; +Chandra & Ahirwar 2005 +, +2007 +; +Bezděk & Krell 2006 +; +Sewak 2009a +, +2009b +; + +Sabu +et al. +2011 + +; Chandra +et al. +2012a; +Karimbumkara & Rajan 2013 +; + +Gupta +et al. +2015 + +; + +Siddiqui +et al. +2014 + +; +Bezděk 2016 +; + +Ghosh +et al. +2020 + +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFDD892A9D90FEB9FDACFD95.xml b/data/55/71/87/557187D2FFDD892A9D90FEB9FDACFD95.xml new file mode 100644 index 00000000000..4e394d88a3c --- /dev/null +++ b/data/55/71/87/557187D2FFDD892A9D90FEB9FDACFD95.xml @@ -0,0 +1,398 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis siva +Gillet, 1911 + + + + + + +( +Figs 4K–N +, +12B +, +20 + + +Onitissiva +Gillet, 1911: 313 +(original description). + + + + + + +Onitis siva + +– + +Arrow 1931: 388 + +(key, description). — + +Janssens 1937: 109 + +(revision). — + +Balthasar 1963: 53 + +(monograph).— + +Rajan 2006:108 + +(list,key, diagnosis, distribution).— + +Chandra2009a:81 + +(distribution). — + +Sewak 2009a: 35 + +(key, diagnosis, distribution); 2009b: 58 (key, diagnosis, distribution). — + + +Sabu +et al. +2011: 34 + + +(list). — + +Karimbumkara & Rajan 2013: 175 + +(list). — + + +Gupta +et al. +2015: 1037 + + +(list, distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + +Onitissiva +belongs to Group XVIII. It is closer to + +O. bordati + +which is the nearest group member in having an uninterrupted frontal carina and the absence of clypeal carina. + +Onitis siva + +is easily distinguishable by the strongly punctured pronotum whereas in the latter it is granular. + + + + + +Type material examined + + + + +Holotype + + + + +INDIA +• +♀ +; “Travancore; A Janssens vid., 1936: +Onitissiva +Gillet, +J.J. Gillet +set., vend. +Onitissiva, +Gillet RMHN. Belg. 10.640, cf. Ann. Soc. Ent. Belg. LV. 1911, 313; +Type +, Wallardi (Travancore) +R.D. Favre +, + +5.9.1903 + +”; +IRSNB +. + + + +Additional material examined +( +10 specimens +) + + + +INDIA +• +♂ +; “Nilgiri Hills, H.L. Andrewes., + +3000 ft. + +, + +Onitis siva + +male, Gillet”; +BMNH +(E) 1237159 + +• + +♂ +; +Karnataka +, +Biligiri Rangaswamy Tiger Reserve +, +Budipadaga +scrub; N 11°50ʹ16.8″, +77°4ʹ39.18″ E +; + +14 May 1999 + +; +open cattle dung baited trap +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000703 + +• + +♂ +; +Karnataka +, +Nagarhole National Park +, core moist deciduous forest; +12°1ʹ13.8″ N +, +76°6ʹ14.76″ E +; +AIMB +/ +Co +/ +Sc +1000704 + +• + +2 ♂♂ +, +2 ♀♀ +; +Hebbala +moist deciduous forest; +12°9ʹ22.68″ N +, +76°6ʹ51.84″ E +; +AIMB +/ +Co +/ +Sc +1000705 to 1000708 + +• + +♂ +, +♀ +; +Sungathakatte Dry Deciduous forest +; +11°58ʹ27.12″ N +, +76°11ʹ8.16″E +; +open cattle dung baited trap +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000709 to 1000710 + +. + + + +NEPAL +• +♀ +; “ +Chitwan +, Tiger Tops. + + +150m + +. + +; + +24 Oct. 1983 + +; +K.K. Gurung +; Rhinoceros dung; Brit. Mus. 1984. 19”; +BMNH +(E) 1237160 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=22.5–31.5; BW=12.5–16. + +Male + +Black or deep reddish-black, not very shining, elongate-oval, moderately convex; clypeus almost semicircular, finely and densely rugose, clypeal carina absent; frontal carina strongly elevated and curved; pronotum with a trace of longitudinal median groove posteriorly, strongly and unevenly punctured, punctures confluent anteriorly, minutely granular near lateral margins, basal foveae elongate and close; pygidium flat, indistinctly punctate; metasternum densely hairy at sides and less densely in middle, base with a small smooth but moderately punctured area. Pronotal punctures not strong and even; front legs elongate, femur with very strong spine in the middle of anterior edge, mid and hind femur also toothed. + +AEDEAGUS (in mm) ( +Fig. 4K–N +). LP=4.5, Lp= 3.5; BP=2, BpB=1, BpT=0.5. Phallobase distinctly longer than parameres. Parameres curved inwards, rounded and blunt at the tip. + + +Female + +Pronotal punctures stronger and closer than in males; median elevation of posterior marginal carina of the head broader and stronger than in male, protibia straight, front femora without teeth. + + + + +Distribution + + + +India +: +Arunachal Pradesh +, +Karnataka +, +Kerala +, +Gujarat +, +Rajasthan +, +Tamil Nadu +, +Uttar Pradesh +( +Rajan 2006 +; +Chandra 2009a +; +Sewak 2009a +, +2009b +; + +Sabu +et al. +2011 + +; +Karimbumkara & Rajan 2013 +; + +Gupta +et al. +2015 + +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFDF892C9D40FB67FBC9F863.xml b/data/55/71/87/557187D2FFDF892C9D40FB67FBC9F863.xml new file mode 100644 index 00000000000..c6a1b7cb9b9 --- /dev/null +++ b/data/55/71/87/557187D2FFDF892C9D40FB67FBC9F863.xml @@ -0,0 +1,658 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis singhalensis +Lansberge, 1875 + + + + + + +Figs 8I–L +, +9F +, +12A +, +14F +, +15F +, +16F +, +19 + + +Onitissinghalensis +Lansberge, 1875: 140 +(original description). + + + + + + +Onitis singhalensis + +– + +Arrow 1931: 394 + +(key, description). — + +Janssens 1937: 53 + +(revision). — + +Balthasar 1963: 40 + +(monograph). — + +Biswas & Chatterjee 1986b: 94 + +(distribution). — + +Chandra 2009a: 81 + +(distribution). — + +Sewak 2009b: 59 + +(key, diagnosis, distribution). — + + +Sabu +et al. +2011: 34 + + +(list). — + +Karimbumkara & Rajan 2013: 175 + +(distribution). — + + +Siddiqui +et al. +2014: 300 + + +(distribution). — + + +Gupta +et al. +2015: 1037 + + +(list, distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + +The clypeus and vertex of + +Onitis singhalensis + +have close granules, giving the clypeus a rugose appearance. + +Onitis singhalensis + +is closer to + +O. philemon + +, + +O. kethai + +sp. nov. +and + +O. visthara + +sp. nov. +but differs in the nature of the granules on the head and also the size. See remarks under + +O. philemon + +, + +O. kethai + +and + +O. visthara + +. + + + + + +Type material examined + + + + +Holotype + + + + +SRI LANKA +• +♂ +; “Ceylon, singhalensis typ. Lansbrg., ex-Musaeo E Harold, +A. Janssens +vid., 1936 +Onitissinghalensis +Lansb., +Type +,” +MNHN +EC4812 +. + + + +Additional material examined +( +204 specimens +) + + + +INDIA +• +♂ +; “ +Nilgiri Hills. +, +H. L. Andrewes. +, Andrewes Bequest., B.M. 1922–221]”; ( +BMNH +(E) 1237145 + +• + +♀ +; +South India +”; ( +BMNH +(E) 1237146) + +• + +20 ♂♂ +, +5 ♀♀ +; +Karnataka +, +Biligiri Rangaswamy Tiger Reserve +, +Budipadaga Scrub +; +11°50ʹ16.8″ N +, +77°4ʹ39.18″ E +; + +14 May 1999 + +; +AIMB +/ +Co +/ +Sc +1000502 to 1000526 + +• + +2♂♂ +, +1♀♀ +; +Gombegallu +evergreen forest; +11°55ʹ54.6″ N +, +77°11ʹ3.48″ E +; + +30 March 2000 + +; +AIMB +/ +Co +/ +Sc +1000527 to 1000558 + +• + +17 ♂♂ +, +10 ♀♀ +; +Kanneri +moist deciduous forest; +11°53ʹ43.62″ N +, +77°8ʹ9.48″ E +; + +22 Dec. 1999 + +; +AIMB +/ +Co +/ +Sc +1000559 to 1000585 + +• + +19 ♂♂ +, +7 ♀♀ +; +Purani +dry deciduous forest; +12°1ʹ55.2″ N +, +77°7ʹ52.68″ E +; + +11 May 1999 + +; +AIMB +/ +Co +/ +Sc +1000586 to 1000611 + +• + +22 ♂♂ +, +15 ♀♀ +; +Honnametti Shola +; +11°53ʹ26.52″ N +, +77°12ʹ4.98″ E +; + +Apr. 1998 + +; +AIMB +/ +Co +/ +Sc +1000612 to 1000648 + +• + +16 ♂♂ +, +9 ♀♀ +; +Gummane Shola +12°1ʹ10.2″ N +, +77°10ʹ30.66″ E +; + +May 1998 + +; +open cow dung baited trap +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000649 to 1000673 + +• + +♂ +, +♀ +; +Karnataka +, +Nagarhole National Park +, +Core +moist deciduous forest; +12°1ʹ13.8″ N +, +76°6ʹ14.76″ E +; +AIMB +/ +Co +/ +Sc +1000674 to 1000675 + +• + +2 ♂♂ +, +♀ +; +Hebbala +moist deciduous forest; +12°9ʹ22.68″ N +, +76°6ʹ51.84″ E +; +AIMB +/ +Co +/ +Sc +1000676 to 1000678 + +• + +♂ +, +2 ♀♀ +; +Sungathakatte +dry deciduous forest; +11°58ʹ27.12″ N +, +76°11ʹ8.16″ E +; +open cow dung baited trap +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000679 to 1000681 + +• + +5 ♂♂ +, +8 ♀♀ +; +Kerala +, +Shendurney Wildlife Sanctuary +, +Rockwood +grassland; +08°51ʹ41.8ʺ N +, +77°08ʹ00.5ʺ E +; elev. + +839 m +a.s.l. + +; + +8 May 2009 + +; +AIMB +/ +Co +/ +Sc +1000682 to 1000694 + +• + +2 ♂♂ +; +Kattilappara +moist deciduous forest; +08°54ʹ54.7ʺ N +, +77°06ʹ28.7ʺ E +; elev. + +185 m +a.s.l. + +; + +3 Dec. 2009 + +; +AIMB +/ +Co +/ +Sc +1000695 to 1000696 + +• + +2 ♂♂ +, +2 ♀♀ +; +Kattilapara +degraded forest; +08°55ʹ05.6ʺ N +, +77°05ʹ44.7ʺ E +; elev. + +207 m +a.s.l. + +; + +3 Dec. 2009 + +; +open cow dung baited trap +; +AIMB +/ +Co +/ +Sc +1000696 to 1000699 + +• + +2 ♂♂ +, +♀ +; +Kattilappara +teak plantation; +08°55ʹ22.9ʺ N +, +77°05ʹ39.2ʺ E +; elev. + +144 m +a.s.l. + +; + +3 Dec. 2009 + +; +open cow dung bait trap +; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000700 to 1000702 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=16–23, BW= 9–12. + +Male + +Coppery or greenish coppery, not very shining, elongate oval; head closely granular, clypeal margin slightly bidentate in the middle, clypeofrontal carina interrupted in the middle with an anterior short clypeal carina and a conical tubercle immediately behind; pronotum fairly strongly and unevenly punctured and with a smooth median line, lateral margin feebly curved anteriorly, bisinuate behind, front angle blunt, hind angle obsolete, base strongly rounded; elytra moderately strongly striate with intervals very minutely punctured; pygidium scarcely perceptibly punctured; metasternal shield strongly longitudinally grooved in the middle anteriorly, smooth below, granular and hairy at the sides, sides of the metasternum closely granular and hairy. Profemur with a sharp tooth towards the end of its anterior edge, mid femur and hind trochanter also toothed. + +AEDEAGUS (in mm) ( +Fig. 8I–L +). LP=3.5, Lp= 2.75; BP= 1.25, BpB =0.75, BpT= 0.5. Phallobase longer than the parameres. The base of the parameres is straight near the phallobase, forms an angle at one third of its length, is straight anteriorly and curves at the tip. + + +Female + +Clypeus longer, rugulose; protibia broad and armed with four external teeth and an articulated spur. + + + + +Distribution + + + +India +: +Gujarat +, +Karnataka +; +Kerala +, +Rajasthan +, +Tamil Nadu +. Other countries: +Sri Lanka +; +Pakistan +, +Nepal +(Biswas & Chatterjee 1986; +Chandra 2009a +; +Sewak 2009a +, +2009b +; + +Sabu +et al. +2011 + +; +Karimbumkara & Rajan 2013 +; + +Siddiqui +et al. +2014 + +; + +Gupta +et al. +2015 + +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFE089119D82FC59FB66F930.xml b/data/55/71/87/557187D2FFE089119D82FC59FB66F930.xml new file mode 100644 index 00000000000..d214704d498 --- /dev/null +++ b/data/55/71/87/557187D2FFE089119D82FC59FB66F930.xml @@ -0,0 +1,544 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis feae +Felsche, 1907 + + + + + + +Figs 10A +, +11E–H +, +18 + + + + + + + +Onitis feae +Felsche, 1907: 293 + + +(original description). + + + + + +Onitis feae + +– + +Boucomont & Gillet 1921: 17 + +, 18 (distribution). — + +Arrow 1931: 397 + +(key, description). — + +Janssens 1937: 75 + +(revision). — + +Paulian 1945: 141 + +(revision). — Singh +et al. +2010: 1395 (list). — + + +Gupta +et al. +2015: 1037 + + +(checklist). — + +Bezděk 2016: 179 + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + + +Onitis feae + +is distinct in having an elliptical clypeus and a sharply elevated frontal carina which is complete. + + + + + + +Type +material examined + + + + +Cotype + + + +MYANMAR +• +♂ +; “Asciuii Cheba, + +1200-1300 m + +, +L. Fea. +1–88; Co-Typus (red label); +Coll. C. Felsche +, Kauf 20, 1918; Staatl. Museum für Tierkunde Dresden; +SNSD +” + +. + + +Additional material examined +( +111 specimens +) + + + +INDIA +• +♂ +; +Meghalaya +, +Shillong +, village near +Northeast Hill University +( +NEHU +); + +31 Oct. 2012 + +; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000185 + +• + +2 ♂♂ +, +♀ +; +Arunachal Pradesh +, +Mobu +; +27°14ʹ17.7324ʺ N +, +93°32ʹ14.28ʺ E +; elev. + +876 m +a.s.l. + +; + +19 Oct. 2018 + +; hand-picked, mithun dung; +Priyadarsanan Dharma Rajan +and team leg.; +AIMB +/ +Co +/ +Sc +1000186 to 1000188 + +• + +6 ♂♂ +, +5 ♀♀ +; +Arunachal Pradesh +, +Aalong +; +28°8ʹ36.384ʺ N +, +94°49ʹ33.34ʺ E +; elev. + +317 m +a.s.l. + +; + +14 Nov. 2019 + +; hand-picked, mithun dung; +Sneha Haridas +and team leg.; +AIMB +/ +Co +/ +Sc +1000189-1000199 + +• + +13 ♂♂ +, +12 ♀♀ +; +Arunachal Pradesh +, +West Siang +, +Aalong forest +; +28°8ʹ33.882ʺ N +, +94°49ʹ36.1848ʺ E +; elev. + +359 m +a.s.l. + +; + +14 Nov. 2019 + +; +cow dung baited pitfall trap +; +Sneha Haridas +and team leg.; +AIMB +/ +Co +/ +Sc +1000200 to 1000224 + +• + +17 ♂♂ +, +22 ♀♀ +; +Arunachal Pradesh +, +West Siang +, +Kaying forest +; +28°24ʹ35.8668ʺ N +, +94°41ʹ51.4896ʺ E +; elev. + +376 m +a.s.l. + +; + +15 Nov. 2019 + +; +cow dung baited pitfall trap +; +Sneha Haridas +and team leg.; +AIMB +/ +Co +/ +Sc +1000225 to 1000263 + +• + +10 ♂♂ +, +15 ♀♀ +; +Arunachal Pradesh +, +West Siang +, +Darak +/ +Larmuk Pottom forest +; +28°12ʹ17.568ʺ N +, +94°35ʹ49.3008ʺ E +; elev. + +489 m +a.s.l. + +; + +18 Nov. 2019 + +; +mithun dung baited pitfall trap +; +Sneha Haridas +and team leg.; +AIMB +/ +Co +/ +Sc +1000264 to 1000288 + +• + +2 ♂♂ +, +2 ♀♀ +; +Arunachal Pradesh +, +Boru Raksap +/ +Yardi Rabe Supse Wildlife Sanctuary +; +28°15ʹ56.1132ʺ N +, +94°34ʹ58.008ʺ E +; elev. + +529 m +a.s.l. + +; + +24 Nov. 2019 + +; +mithun dung baited pitfall trap +; +Sneha Haridas +and team leg.; +AIMB +/ +Co +/ +Sc +1000289 to 1000292 + +. + + + +LAOS +• +♀ +; “ +Xieng Khouang +, + +8.i.1919 + +; +R.V. de Salvaza +; Indo-China, B.M. 1924-315; +BMNH +(E) 1237162, Southeast Asia” + +. + + + +MYANMAR +• +♂ +; “ex Mus. Calcutta, B.M. 1930–1. Man Hpat, +N. Tawngpeng +, N. +Shan +States, +Upper Burma +, 11-ii-15., + +4600 ft. + +; +J. Coggin Brown +; in dung; Southeast Asia;” +BMNH +(E) 1237161 + +• + +♀ +; “ +Loi +, +Tawng Kyan +, Tawng, N. +Shan +States +, +Upper Burma +• +♂ +; +Tawng Peng +, Pansai, N. +Shan +States +, +Upper Burma +, + +4660 ft. + +” + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=15–20.5; BW= 8.5–11.5. + +Male + +Black, subopaque; head, lower surface and legs dark red; pygidium lower surface and fringe of elytra with reddish hairs; body oval, compact, convex; clypeus elliptical, finely, closely rugulose; forehead rugosely punctured with sharply elevated carina; clypeal carina short, transverse, slightly curved; vertex with a short transverse curved carina before carinate posterior margin of head; pronotum strongly, closely punctured, with feeble median groove, base margined, basal foveae very deep, space between them finely rugose; elytra very convex, opaque, finely striate with intervals carinate along middle; pygidium feebly rugulose, thinly clothed with erect, long reddish hairs; metasternum granular, broadly grooved in the middle; abdomen hairy at sides, smooth in the middle; profemur with sharp tooth near middle of front edge; protibia with finely serrate carina beneath and a tooth near base; middle coxa toothed at its posterior end. + +AEDEAGUS (in mm) ( +Fig. 11E–H +). LP=2.86, Lp =2.68; BP= 1.25, BpB=1.07, BpT=0.36. Phallobase and parameres almost of same size; parameres straight above and slightly curved below with the tips blunt and rounded. + + +Female + +Clypeus extended and reflexed anteriorly, slightly notched; no sharp tooth on underside of protibia and profemur. + + + + +Distribution + + + +India +: +Assam +, +Arunachal Pradesh +, +Meghalaya +, +Mizoram +; Other countries: +China +; +Laos +; +Myanmar +; +Thailand +(Singh +et al. +2010; + +Gupta +et al. +2015 + +; +Bezděk 2016 +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFE1891F9D58F8B1FD76FE31.xml b/data/55/71/87/557187D2FFE1891F9D58F8B1FD76FE31.xml new file mode 100644 index 00000000000..adc5173cf4c --- /dev/null +++ b/data/55/71/87/557187D2FFE1891F9D58F8B1FD76FE31.xml @@ -0,0 +1,375 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis humerosus +( +Pallas, 1771 +) + + + + + + +Figs 10B +, +18 + + +Scarabaeushumerosus +Pallas, 1771: 462 +(original description). + + +Scarabaeusmenalcas +Pallas, 1781: 4 +. + + +Onitischevrolati +Lucas 1846: pl. 23. + + +Onitishumerosusviolaceus +Sahlberg, 1913: 119 +. + + + +Onitis humerosus viridipenni + +sMikŠić, 1950: 165–167. + + +Onitishumerosusviolaceipennis +MikŠić, 1950: 165–167 +. + + +Onitishumerosuscupripennis +Balthasar, 1963: 48 +. + + + + + + +Scarabaeus humerosus + +– Castelnau 1840: 89 (natural history). — + +Lansberge 1875: 116 + +(monograph). — + +Bedel 1892: 268 + +, 271 (revision). — + +Reitter 1892: 97 + +(keys). — + +Arrow 1931: 390 + +(identification key, description). — + +Porta 1932: 414 + +(distribution). — + +Balthasar 1935: 91 + +(monograph); 1963: 47 (monograph).— + +Janssens 1937: 91 + +(revision). — +Hashmi &Tashfeen 1992 +(distribution).— + +Awal 2006: 170 + +(distribution). — + +Bai 2006: 391 + +(identification key). — + +Bezděk & Krell 2006: 159 + +(distribution, synonyms). — + + +Bunalski +et al. +2014: 156 + + +(list of species). — Siddhiqui +et al. +2014: 300 (distribution). — + +Bezděk 2016: 179 + +(distribution, synonyms). — + + +Bunalski +et al. +2016: 36 + + +(distribution). — + +Montreuil 2017: 265 + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + +Onitishumerosus +is a very distinct species having bright yellow elytra with longitudinal green lines upon elevated sutures. + + + + + +Type material examined + + + + +Syntypes + +( +5 specimens +) + + +RUSSIA +• +3 ♂♂ +, +2 ♀♀ +; “ + +Scarabaeus menalcas + +; +MfN +Janssens det. as + +Onitis humerosus + +in 1938; Hist.- Coll. ( +Coleoptera +), Nr. 26753, +Onitismenalcas +Pallas, 1781 +, Russia austr., Bob., Zool. Mus. Berlin; Red Label: +SYNTYPUS +, + +Scarabaeus menalcas +Pallas, 1781 + +, labelled by MNHUB 2007”; +MfN +. + + +Additional material examined +( +7 specimens +) + + + +ISRAEL +• +♂ +; “ + +Onitis humerosus +Pallas, Mid. + +East; +Jerusalem +, +G.R. Crotch. +, Nevinson Coll., 1918- 14”; +BMNH +(E) 1237143 + +• + +♀ +; “ +Onitishumerosus +Pallas, Mid. East; Pascoe, Coll. 93–60;” +BMNH +(E) 1237144 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=12.5–18.5; BW=7–10. + +Male + + +Bright metallic green, coppery, indigo-blue or violet, the elytra bright yellow with four narrow longitudinal green lines on the elevated 1 +st +, 3 +rd +and 5 +th +intervals, pale intervals with light metallic lustre; body oval, convex, not very shining; head closely covered with fine tubercles; clypeus parabolic, rugose with yellow setae, clypeal carina curved, elevated, near to clypeal margin; frontal carina entire with a tubercle behind; pronotum metallic green, unevenly punctured posteriorly, front angles granular, basal foveae deep, close with granules anteriorly; elytra finely striate with sutural, 3 +rd +and 5 +th +intervals elevated, shining, finely punctured, the other intervals opaque, scarcely punctured; pygidium shining, finely, sparsely granular; metasternum with a longitudinal groove, metasternal shield broad, granular and thinly covered with hairs anteriorly, sparingly punctured behind, sides of metasternum closely granular and hairy; protibia elongated, slender with a strong tooth at the middle of the lower surface; hind femur with a very strong hooked tooth near the middle of its upper edge. + + +Female + +Clypeal carina long, curved, space between the carina and front margin covered with long yellow, closelying hairs; legs short, stout, protibia with four strong teeth and an articulated spur. + + + + +Distribution + + + +Afghanistan +; +Albania +; +Armenia +; +China +; +Cyprus +; +Georgia +; +Iran +; +Israel +; +Jordan +; +Kazakhstan +; +Kyrgyzstan +; +Lebanon +; +Pakistan +; +Russia +; +Syria +; +Tajikistan +; +Turkey +; +Turkmenistan +; +Uzbekistan +( +Hashmi & Tashfeen 1992 +; +Awal 2006 +; +Bezděk & Krell 2006 +; Siddhiqui +et al. +2014; +Bezděk 2016 +; + +Bunalski +et al. +2016 + +; +Montreuil 2017 +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFE289109D65FB6AFC27FC2B.xml b/data/55/71/87/557187D2FFE289109D65FB6AFC27FC2B.xml new file mode 100644 index 00000000000..bc73ef12f67 --- /dev/null +++ b/data/55/71/87/557187D2FFE289109D65FB6AFC27FC2B.xml @@ -0,0 +1,898 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis falcatus +( +Wulfen, 1786 +) + + + + + + +Figs 5F +, +6H +, +18 + + + + + + + +Scarabaeus falcatus +Wulfen, 1786: 14 + + +(original description). + + + +Onitishimaleyicus +Redtenbacher, 1844: 518 (synonym). + + + + + +Onitis kiuchii +Masumoto 1995: 88 + + +(synonym). + + + +Onitisfalcatus +– +Lansberge 1875: 126 +(monograph). — Boucomont & Gillet: 1921: 17 (distribution). — +Arrow 1931: 392 +(key and description). — +Balthasar 1935: 93 +(monograph); 1963: 33 (monograph). — +Janssens 1937: 44 +(revision). — +Paulian 1945: 142 +(revision). — +Biswas & Chatterjee 1985: 158 +(distribution); 1991: 260 (distribution). — +Chandra 2000: 360 +(distribution); 2005: 150 (distribution). — +Mittal 2005: 46 +(list). — +Rajan 2006: 108 +(key, distribution). — +Bezděk & Krell 2006: 159 +(distribution). — +Chandra & Ahirwar 2007: 276 +(distributional). — +Sewak 2009a: 36 +(diagnosis, distribution); 2009b: 58 (diagnosis and distribution). — Singh +et al. +2010: 1395 (distribution). — + +Sabu +et al. +2011: 34 + +(list). — Chandra +et al. +2012a: 52 (distribution). — +Karimbumkara & Rajan 2013: 174 +(distribution). — + +Gupta +et al. +2015: 1037 + +(checklist). — +Bezděk 2016: 179 +(distribution). — + +Ghosh +et al. +2020 + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + +Differential diagnosis + + + +Onitisfalcatus +is similar to + +O. excavatus + +in having a minutely punctured pronotum but differs in the nature of rugosity on the clypeus, the presence of longitudinal groove on the pronotum and also in the shape of the body. (Also see remarks under +Onitisexcavatus. +) + + + + + +Type material examined + + + + +Syntypes + +( +2 specimens +) + + +INDIA +• +2 ♀♀ +; “ + +O. himaleyicus + +; +TYPUS +(Red card); Hugel: 170; +Himaleyicus K. Redt. +”; Kashmir; specimen number not assigned; +NHMW +. + + +Additional material examined +( +125 specimens +) + + + +INDIA +• +♂ +, +♀ +; “ +Darjiling +, +Singla +, + +1500 ft + +, +Lord Carmichael’s +collection, + +May 1913 + +”; +ZSIM + +• + +27 ♂♂ +, +16 ♀♀ +; +Karnataka +, BRT +Tiger Reserve +: +Gombegallu +evergreen forest; + +6 Jun. 1999 + +; +11°55ʹ54.6 N +, +77°11ʹ3.48″ E +; +AIMB +/ +Co +/ +Sc +1000064 to 1000106 + +• + +6 ♂♂ +, +5 ♀♀ +; +Karnataka +, BRT +Tiger Reserve +: +Kanneri +moist deciduous forest; + +28 May 1999 + +; +11°53ʹ43.62″ N +, +77°8ʹ9.48″ E +; +open cow dung bait traps +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000107 to 1000117 + +• + +♀ +; +Karnataka +, BRT +Tiger Reserve +: +Purani +dry deciduous forest; + +18 May 1999 + +; +12°1ʹ55.2″ N +, +77°7ʹ52.68″E +; +open cow dung bait traps +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000118 + +• + +9 ♂♂ +, +7 ♀♀ +; +Karnataka +, BRT +Tiger Reserve +: +Honnametti Shola +; + +Apr. 1998 + +; +11°53ʹ26.52″ N +, +77°12ʹ4.98″ E +; +open cow dung bait traps +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000119 to 1000134 + +• + +13 ♂♂ +, +5 ♀♀ +; +Karnataka +, BRT +Tiger Reserve +: +Gummane Shola +; + +May 1998 + +; +12°1ʹ10.2″ N +, +77°10ʹ30.66″ E +; +open cow dung bait traps +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000135 to 1000152 + +• + +2 ♂♂ +, +♀ +; +Kerala +, +Wayanad +, arecanut plantation; + +14 Nov. 2010 + +; +open cow dung bait trap +; +Nisha, P. V. +leg.; +AIMB +/ +Co +/ +Sc +1000153 to 1000155 + +• + +4 ♂♂ +, +3 ♀♀ +; +Kerala +, +Shendurney Wildlife Sanctuary +, grassland; + +8 May 2009 + +; +08°51ʹ41.8ʺ N +, +77°08ʹ00.5ʺ E +; +open cow dung bait traps +; +Kishore A. +leg.; +AIMB +/ +Co +/ +Sc +1000156 to 1000162 + +• + +♂ +, +♀ +; +Kerala +, +Shendurney Wildlife Sanctuary +, degraded forests; + +3 Dec. 2009 + +; +08°55ʹ05.6ʺ N +, +77°05ʹ44.7ʺ E +; +open cow dung bait traps +; +Kishore A. +leg.; +AIMB +/ +Co +/ +Sc +1000163 to 1000164 + +• + +♂ +, +Kerala +, +Shendurney Wildlife Sanctuary +, rubber plantation; + +5 May 2009 + +; +08°51ʹ41.8ʺ N +, +77°08ʹ00.5ʺ E +; +open cow dung bait traps +; +Kishore A. +leg.; +AIMB +/ +Co +/ +Sc +1000165 + +• + +♀ +; +Kerala +, +Shendurney Wildlife Sanctuary +, teak plantations; + +8 Oct. 2008 + +; +open cow dung bait traps +; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000166 + +• + +♀ +; +Mizoram +, +Pasighat +, +Kawlbem +, +Lengteng Wildlife Sanctuary +; +23°52ʹ16.6404ʺ N +, +93°18ʹ22.2408ʺ E +; elev. + +1500 m +a.s.l. + +; + +14 Apr. 2019 + +; +cow dung baited pitfall trap +; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000167 + +• + +5 ♂♂ +, +3 ♀♀ +; +Arunachal Pradesh +, agricultural field, +Rani Village +; +27°58ʹ10.6068ʺ N +, +95°19ʹ26.1156ʺ E +; elev. + +131 m +a.s.l. + +; + +30 Sep. 2018 + +; hand-picked from cow dung; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000168 to 1000175 + +• + +♂ +, +♀ +; +Arunachal Pradesh +, community forest, +Pasighat +, +Rani Village +; +27°58ʹ9.0156ʺ N +, +95°19ʹ18.9804ʺ E +; elev. + +133 m +a.s.l. + +; + +30 Sep. 2018 + +; +cow dung baited pitfall trap +; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000176 to 1000177 + +• + +♂ +; +Arunachal Pradesh +, +Darak +/ +Larmuk Pottom forest +, +West Siang +; +28°12ʹ12.6576ʺ N +, +94°35ʹ45.546ʺ E +; elev. + +434 m +a.s.l. + +; + +18 Nov. 2019 + +, mithun dung baited +pitfall +; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000178 + +• + +3 ♂♂ +, +2 ♀♀ +; +Arunachal Pradesh +, +Kaying forest +, +West Siang +; + +15 Nov. 2019 + +; cow dung, +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000179 to 1000183 + +• + +♀ +; +Nagaland +, +Bongkolong village +, near +Intanki National Park +, open ground; + +15 Apr. 2021 + +; hand-picked from cow dung; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000184 + +. + + + +MALAYSIA +• +♀ +; “ +PERAK +, F.M.S., + +Sep. 1907 + +, Coll.; Ex. F.M.S. Museum., B.M. 1955–354; + +Onitis falcatus +J. Gillet + +det. +Wulf. +1923, +S.E. Asia +”; +BMNH +(E) 1237166 + +. + + + + + +Description + + + +MEASUREMENTS: TL: +16–23 mm +; BW: +8–12 mm +. + + +Male + + +Black, smooth, shining; clypeus elliptical, feebly excised in middle, moderately closely, finely rugulose; frontal carina widely interrupted, a short clypeal carina; pronotum finely and sparsely punctured without a median groove; elytra finely striate, 1 +st +, 3 +rd +and 5 +th +intervals raised, striae very shallow, irregular punctures on sutural intervals; pygidium opaque, smooth, unpunctured; metasternal shield smooth, feebly punctured, hairy anteriorly; protibia elongate with two or three minute teeth before the middle of the lower surface, a sharp incurved terminal process, a minute tooth near the end of the lower edge of the mesofemur. + + +AEDEAGUS (in mm) ( +Fig. 6E–H +). LP= 2, Lp=1.75; BP=1, BpB =0.5, BpT=0.25. Parameres shorter than phallobase, straight above and slightly curved below and hooked at the tip. + + +Female + +Pronotum with stronger punctures; protibia uniformly curved with an articulated spur at the tip. + + + + +Distribution + + + +India +: +Andhra Pradesh +, +Arunachal Pradesh +, +Assam +, +Bihar +, +Gujarat +, +Haryana +, +Himachal Pradesh +, +Jharkhand +, +Karnataka +, +Kerala +, +Madhya Pradesh +, +Maharashtra +, +Manipur +, +Meghalaya +, +Mizoram +, +Nagaland +, +Odisha +, +Puducherry +, Punjab, +Rajasthan +, +Tamil Nadu +, +Tripura +, +Uttarakhand +, +Uttar Pradesh +, New +Delhi +, +West Bengal +. Other countries: +Bangladesh +; +Bhutan +; +Cambodia +; +China +; +Laos +; +Myanmar +; Malay Peninsula; +Nepal +; +Pakistan +; +Philippines +; +Taiwan +; +Thailand +; +Vietnam +( +Biswas & Chatterjee 1985 +; +Chandra 2000 +; +Mittal 2005 +; +Rajan 2006 +; +Bezděk & Krell 2006 +; +Chandra & Ahirwar 2007 +; +Sewak 2009a +, +2009b +; Singh +et al. +2010; + +Sabu +et al. +2011 + +; Chandra +et al. +2012a; +Karimbumkara & Rajan 2013 +; + +Gupta +et al. +2015 + +; +Bezděk 2016 +; + +Ghosh +et al. +2020 + +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFE589129D69FE6CFCB4FB1F.xml b/data/55/71/87/557187D2FFE589129D69FE6CFCB4FB1F.xml new file mode 100644 index 00000000000..7bb8a76b6b6 --- /dev/null +++ b/data/55/71/87/557187D2FFE589129D69FE6CFCB4FB1F.xml @@ -0,0 +1,589 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis excavatus +Arrow, 1931 + + + + + + +Figs 5E +, +6A–D +, +18 + + +Onitisexcavatus +Arrow, 1931: 391 +(original description). + + + + + + +Onitis excavatus + +– + +Balthasar 1935: 93 + +(monograph); 1963: 37 (monograph). — + +Janssens 1937: 50 + +(revision); 1938: 2 (notes). — + +Biswas & Chatterjee 1985: 158 + +(distribution). — + +Mittal 2005: 46 + +(list). — + +Bezdek & Krell 2006: 159 + +(distribution). — Chandra +et al. +2012a: 52 (distribution). — + + +Siddiqui +et al. +2014: 300 + + +(distribution). — + + +Gupta +et al. +2015: 1037 + + +(checklist). — + +Bezděk 2016: 179 + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + +Onitisexcavatus +closely resembles + +O. falcatus + +, both placed in Group I. These two species are similar as the pronotum is smooth or sparingly punctured, and the former differs from the latter in having larger and broader body, a deep transverse excavation behind the middle of the metasternum and presence of strong sharp teeth at the extremity of mesofemur in male. + + + + + +Type material examined + + + + +Lectotype + + + +MYANMAR +• +♂ +; “Tenasserim, Atkinson Coll.92–3, near + +O. Ludekingi, +Lansb. + +, but quite diff. by sculptures, + +Onitis excavates + +male Arr., M.E. Bacchus det. 1975”; +BMNH +(E) 1237173. + + +Additional material examined +( +29 specimens +) + + + +INDIA +• +2 ♂♂ +, +5 ♀♀ +; +Arunachal Pradesh +, +Pakke +; +27°14ʹ17.73″ N +, +93°32ʹ14.28″ E +; elev. + +877 m +a.s.l. + +; hand-picked from elephant dung; + +24 Sep. 2018 + +; +Priyadarsanan Dharma Rajan +and team leg.; +AIMB +/ +Co +/ +Sc +1000037 to 1000044 + +• + +♂ +, +♀ +; +Arunachal Pradesh +, +Sibuk +; +28°21ʹ43.7184ʺ N +, +95°15ʹ57.1968ʺ E +; elev. + +956 m +a.s.l. + +; + +3 Oct. 2018 + +; hand-picked from cow dung; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000045 to 1000046 + +• + +♀ +; +Arunachal Pradesh +, community forest, +Pasighat +, +Rani Village +; +27°31ʹ27.138ʺ N +, +27°31ʹ27.138ʺ E +; elev. + +1557 m +a.s.l. + +; + +9 Oct. 2018 + +; hand-picked from donkey dung; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000047 + +• + +♀ +; +Arunachal Pradesh +, +Ziro +, +Talley Valley +; +27°31ʹ23.4984ʺ N +, +93°50ʹ22.0308ʺ E +; elev. + +1532 m +a.s.l. + +; + +10 Oct. 2018 + +; hand-picked from cow dung; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000048 + +• + +♂ +; +Arunachal Pradesh +, +Sangri +, +Mobu +; +27°14ʹ1.5324ʺ N +, +93°31ʹ58.8576ʺ E +; elev. + +805 m +a.s.l. + +; + +18 Oct. 2018 + +; hand-picked from mithun dung; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000049 + +• + +♀ +; +Arunachal Pradesh +, +Seijosa +, +Pakke Tiger Reserve +; +26°56ʹ56.2596ʺ N +, +92°59ʹ4.3764ʺ E +; elev. + +164 m +a.s.l. + +; + +23 Sep. 2018 + +; hand-picked from elephant dung; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000050 + +• + +6 ♂♂ +, +4 ♀♀ +; +Arunachal Pradesh +, +Seijosa +, +Pakke Tiger Reserve +; +26°56ʹ31.128ʺ N +, +92°58ʹ17.76ʺ E +; elev. + +190 m + +; + +24 Sep. 2018 + +; hand-picked from elephant dung; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000051 to 1000060 + +• + +♂ +; +Arunachal Pradesh +, +Pasighat +, +Rani village +; +27°58ʹ6.0276ʺ N +, +95°19ʹ12.5328ʺ E +; elev. + +135 m +a.s.l. + +; + +29 Sep. 2018 + +; hand-picked from cow dung; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000061 + +• + +♂ +, +♀ +; +Arunachal Pradesh +, +Pasighat +, +Rani village +; +27°58ʹ12.9324ʺ N +, +95°19ʹ20.5284ʺ E +; elev. + +125 m +a.s.l. + +; + +30 Sep. 2018 + +; cow dung; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000061 to 1000062 + +• + +♀ +; +Nagaland +, +Mon District +, +Singphan Elephant Reserve +; + +25 Apr. 2021 + +; hand-picked from cow dung; +Seena Narayanan Karimbumkara +and team leg.; +AIMB +/ +Co +/ +Sc +1000063 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=19–25; BW= 13. + +Male + +Black, shining, elytra and pygidium subopaque; clypeus short, elliptical, moderately closely and finely rugulose; frontal carina widely interrupted in the middle, a short transverse clypeal carina anteriorly, a small tubercle behind, vertex irregularly granular; pronotum very smooth with irregular minute punctures anteriorly, longitudinal groove absent; elytra finely striate with 1st, 3rd and 5th intervals elevated, narrower, rest of the intervals flat; pygidium smooth, unpunctured; metasternum with very deep transverse excavation in the middle, metasternal shield thickly hairy anteriorly, smooth, shining behind; protibia elongate with a serrate carina upon anterior half beneath, four short external teeth and a sharp incurved terminal process; mesofemur with a rounded lobe in the middle of hind edge, two sharp teeth near extremity above and one beneath; trochanters of hind legs acutely produced. + +AEDEAGUS (in mm) ( +Fig. 6A–D +). LP=3.03, Lp=2.93; BP=1.5, BpB=1.14, BpT=0.46. Parameres shorter than phallobase, straight above and slightly curved outwards below and hooked slightly at the tip. + + +Female + +Legs simple, frontal carina slightly diagonal; punctation of the pronotum heavier and denser, elytral striae deeper than in male; metasternum without transverse pit. + + + + +Distribution + + + +India +: +Arunachal Pradesh +, +Assam +, +Himachal Pradesh +, +Meghalaya +, +Nagaland +, +Uttarakhand +, +Uttar Pradesh +. Other countries: +China +; +Myanmar +; +Nepal +; +Pakistan +; +Taiwan +; +Thailand +; +Vietnam +( +Biswas & Chatterjee 1985 +; +Mittal 2005 +; +Bezděk & Krell 2006 +; Chandra +et al. +2012a; + +Siddiqui +et al. +2014 + +; + +Ali +et al. +2015 + +; + +Gupta +et al. +2015 + +; +Bezděk 2016 +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFE789159D7BFEB8FB02FE12.xml b/data/55/71/87/557187D2FFE789159D7BFEB8FB02FE12.xml new file mode 100644 index 00000000000..665c9ccf581 --- /dev/null +++ b/data/55/71/87/557187D2FFE789159D7BFEB8FB02FE12.xml @@ -0,0 +1,388 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis crassus +Sharp, 1875 + + + + + + +Figs 5D +, +18 + + +Onitiscrassus +Sharp, 1875: 48 (original description). + + + + + + +Onitis crassus + +– + +Arrow 1931: 400 + +(keys and description). — + +Janssens 1937: 73 + +(revision). — + +Balthasar 1963: 45 + +(monograph). — + +Chandra 2000: 360 + +(distribution); 2008: 176 (distribution). — + +Bezděk & Krell 2006: 159 + +(distribution). — + +Chandra & Ahirwar 2007: 287 + +(distribution). — Chandra +et al. +2012b: 29 (distribution). — + + +Siddiqui +et al. +2014: 299 + + +(distribution). — + + +Ali +et al. +2015: 4 + + +(distribution). — + + +Gupta +et al. +2015: 1037 + + +(checklist). — + +Bezděk 2016: 179 + +(distribution). + + + +Onitisvischnu +– +Preudhomme de Borre 1881 +: xxxix (synonymised by +Arrow 1931 +). — +Schoolmeesters 2022 +(online catalogue). + + + + + +Differential diagnosis + + + + +Onitis crassus + +is closer to + +O. castaneus + +in having hairy pygidium and uninterrupted clypeo-frontal carina but differs from the latter in having short scanty setae on the pygidium; frontal tubercle being placed behind the carina while in + +O. castaneus + +it is placed on the carina and the anterior margin of clypeus being gently excised in the middle while in the latter it is reflexed and straight. See diagnosis of + +O. brahma + +. + + + + + +Type material examined + + + + +Holotype + + + + +INDIA +• +♀ +; “ +Onitiscrassus +Type D.S.; +Ex. Musaeo D. Sharp +1890, +G. J. Arrow +vid. 1928, +A. Janssens +vid., 1936: +Onitiscrassus +Sharp; +Holotype +(Red Label)”; +MNHN +EC4813 +. + + + +Additional material examined +( +4 specimens +) + + + +INDIA +• +♂ +; “Determined from description G.J.A. + +Onitis Vischnu +, de Borre + +; + +Onitis Vischnu +de Borre, Compared + +with +type +G.J.A., 6745, 7030”; +BMNH +(E) 1237163 + +• + +♀ +; “Dehra Dun Coll. B.M. 1931– 2., +River Suswa +, Dehradun, +U.P. C. Beeson. +XI.’30; Cattle dung on sand”; +BMNH +(E) 1237164 + +• + +♂ +; “A, Janssens vid. 1936: + +ONITIS crassus +Sharp + +; +Onitiscrassus +Shp. Revis. Arrow 1926; +Onitiscrassus +Shp. Compared with +type +G.J.A.; cf. Ann. Soc. Ent. Belg. Xxv, 1881, C.R. p. 39; Ex-Typis de +Vischnu Borre +; det: Predhomme Borre, + +Onitis Vischnu +Borre + +; + +O. Vishnu + +, Reiche, India, 3050, +Coll. J. Thomson +” + +• + +♀ +; same collection data as for preceding; +IRSNB + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=15.5–21; BW=9–10.5. + +Male + +Black; clypeus rugulose, anterior margin strongly elevated; gently excised in the middle; frontal carina entire, curved, with a tubercle behind; pronotum strongly and closely punctured, rugose anteriorly; elytra finely striate, intervals angularly elevated along the middle, finely, fairly closely punctured; pygidium opaque, rugulose with short setae; metasternal shield smooth, shining, feebly, sparsely punctured, sides of metasternum rugosely punctured, with long hairs; front legs elongate with a blunt process on its trochanter; hind femur with a hooked process at the middle of its lower edge. + +Female + +Clypeal margin scarcely excised in the middle; protibia broad with four blunt teeth and an articulated spur. + + + +Fig. 6. +Aedeagus, lateral, dorsal, ventral and apical views. +A–D +. + +Onitis excavatus +Arrow, 1931 + +, AIMB/Co/Sc1000037. +E–H +. + +O. falcatus +( +Wulfen, 1786 +) + +, AIMB/Co/Sc1000070. +I–L +. + +O. bhomorensis + +sp. nov. +, NIM/NBAIR/COL/ONIT/H-271023A. +M–P +. + +O. punctatostriatus +Janssens, 1937 + +, AIMB/ Co/Sc1000492. (Images: AIMB). + + + + + +Distribution + + + +India +: +Madhya Pradesh +, Punjab, +Uttarakhand +, +Uttar Pradesh +. Other countries: +Nepal +; +Pakistan +( +Chandra 2000 +; +Bezděk & Krell 2006 +; +Chandra & Ahirwar 2007 +; +Chandra 2008 +; Chandra +et al. +2012b; + +Siddiqui +et al. +2014 + +; + +Ali +et al. +2015 + +; + +Gupta +et al. +2015 + +; +Bezděk 2016 +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFE889209D58F9C0FB10FD64.xml b/data/55/71/87/557187D2FFE889209D58F9C0FB10FD64.xml new file mode 100644 index 00000000000..5bfdb61f11e --- /dev/null +++ b/data/55/71/87/557187D2FFE889209D58F9C0FB10FD64.xml @@ -0,0 +1,1001 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis philemon +Fabricius, 1801 + + + + + + +Figs 8E–H +, +9D +, +10E +, +14D +, +15D +, +16D +, +19 + + +Onitisphilemon +Fabricius, 1801: 3 +(original description). + + +Onitisdistinctus +Lansberge, 1875: 138 +(synonym). + + +Onitisminor +Lansberge, 1875: 139 +(synonym). + + + + + + +Onitis philemon +– + + +Bates 1891: 10 + +(list). — + +Arrow 1931: 393 + +(key, description). — + +Balthasar 1935: 95 + +(monograph); 1963: 41 (monograph). — + +Janssens 1937: 53 + +(revision). — + +Paulian 1945: 144 + +(revision). — + +Biswas & Chatterjee 1991: 260 + +(distribution). — + +Chandra 2000: 360 + +(distribution); 2005: 150 (distribution); 2009a: 81 (list, distribution). — + +Mittal 2005: 46 + +(list). — + +Chandra & Ahirwar 2005: 149 + +(distribution). — + +Bezděk & Krell 2006: 159 + +(distribution). — + +Sewak 2009a: 37 + +(key, distribution, diagnosis); 2009b:59 (key, distribution, diagnosis).— + + +Sabu +et al. +2011:34 + + +(list).— Chandra +et al. +2011: 66 (distribution). — + +Chandra & Gupta 2011: 274 + +(distribution); 2012a: 889 (distribution); 2013a: 348 (distribution, diagnosis). — Chandra +et al. +2012a: 52 (list and distribution); 2012b: 33 (distribution). — + + +Thakare +et al. +2012: 78 + + +(list and distribution). — + +Karimbumkara & Rajan 2013: 174 + +(list and distribution). — + +Chandra & Gupta 2013a: 348 + +(distribution); 2013b: 4667 (distribution). — + + +Gupta +et al. +2014: 229 + + +(distribution); 2015: 1030 (key, distribution, diagnosis). — + +Bezděk 2016: 179 + +(distribution). — + + +Ghosh +et al. +2020: 238 + + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + +Fig. 9. +Head of male specimens. +A +. +Onitisbhomorensis +sp. nov. +, holotype, NIM/NBAIR/COL/ONIT/H- 271023A. +B +. + +O. punctatostriatus +Janssens, 1937 + +, AIMB/Co/Sc1000492. +C +. + +O. kethai + +sp. nov. +, holotype, NIM/NBAIR/COL/ONIT/H-271023B. +D +. + +O. philemon +Fabricius, 1801 + +, AIMB/Co/Sc1000317. +E +. + +O. visthara + +sp. nov. +, holotype, NIM/NBAIR/COL/ONIT/H-271023C. +F +. + +O. singhalensis +Lansberge, 1875 + +, AIMB/Co/Sc1000684. (Images: AIMB). + + + + +Fig. 10. A +. +Onitisfeae +Felsche, 1907 +, cotype, (image: Olaf Jäger, SNSD). +B +. + +O. humerosus +( +Pallas, 1771 +) + +, BMNH (E) 1237143, (image: BMNH). +C +. + +O. lama +Lansberge, 1875 + +, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). +D +. + +O. naviauxi +Cambefort, 1988 + +, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). +E +. + +O. philemon +Fabricius, 1801 + +, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). +F +. + +O. punctatostriatus +Janssens, 1937 + +, holotype, ♂, (image: Jonathan Brecko, IRSNB). + + + + +Fig. 11. +Aedeagus, lateral, dorsal, ventral and apical views. +A–D +. +Onitiscastaneus +Kollar, 1844 +, AIMB/ Co/Sc1000027. +E–H +. + +O. feae +Felsche, 1907 + +, AIMB/Co/Sc1000188. +I–L +. + +O. lama +Lansberge, 1875 + +, AIMB/Co/Sc1000307. (Images: AIMB). + + + + +Fig. 12.A +. +Onitissinghalensis +Lansberge, 1875 +, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) +B +. +Onitissiva +Gillet, 1911 +, holotype, ♀, (image: Jonathan Brecko, IRSNB). +C +. + +O. subopacus +Arrow, 1931 + +, holotype, ♂, BMNH(E)1237174, (image: BMNH). +D +. + +O.virens +Lansberge, 1875 + +, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). +E–F +Onitisvisthara +sp. nov. +, holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). +E +. Habitus, dorsal view. +F +. Habitus, ventral view. + + + + + +Differential diagnosis + + + +Onitisphilemon +is smaller in size than + +O. singhalensis + +and resembles the latter in the presence of a broad deep longitudinal groove on the metasternum anteriorly and the scarcely punctured pygidium. It differs from the latter in the clypeal sculptures and the absence of a tooth on the profemur of males. In + +O. singhalensis + +the basal pronotal foveae are rounded and closer than in + +O. philemon + +(compare +Fig. 15D, F +) and the pronotum is with a smooth longitudinal median line which extends to the front margin, while the median line is absent in + +O. philemon + +(compare +Fig. 14D, F +). + + + + + +Type material examined + + + + +Lectotype + + + + +UNKNOWN TYPE LOCALITY • + +“ +Philemon + +”; +ZMUK +Fabricius 003706; +Karimbumkara, S.N. +des. 2024; +ZMUK 003706 +. + + + + +Paralectotypes + + + +UNKNOWN TYPE LOCALITY • + +2 specimens +; + +“ +Philemon + +”; +ZMUK +Fabricius 003707, +ZMUK +Fabricius 003708; +Karimbumkara, S.N. +des. 2024; +ZMUK 003707–003708 + +. + + +Remarks + + +In the original description of this species, collection data of the type material are not provided. The Historical Fabricius collections, earlier kept at +ZMUC +and recently transferred to +ZMUK +, has +three specimens +with a simple label + +‘ +Philemon + +’ that matches the original description, which were designated as +syntypes +of +Onitisphilemon +. One of these specimens with code +ZMUK +Fabricius 003706 is now designated as +lectotype +and two other specimens’ +ZMUK +Fabricius 003707 and +ZMUK +Fabricius 003708, are designated as +paralectotypes +. + + +Additional material examined +( +185 specimens +) + + + +INDIA +• +♂ +; +No +label, +ZSIM +; +Karnataka +, BRT +Tiger Reserve + +• + +42 ♂♂ +, +26 ♀♀ +; +Budipadaga Scrub +; +11°50ʹ16.8″N +, +77°4ʹ39.18″E +; + +12Jan.2000 + +; +AIMB +/ +Co +/ +Sc +1000310 to1000377 + +• + +16♂♂ +, +10 ♀♀ +; +Gombegallu Evergreen forest +; N 11°55ʹ54.6″, +77°11ʹ3.48″ E +; + +30 March 2000 + +; +AIMB +/ +Co +/ +Sc +1000378 to 1000403 + +• + +15 ♂♂ +, +12 ♀♀ +; +Kanneri +moist deciduous forest; +11°53ʹ43.62″ N +, +77°8ʹ9.48″ E +; + +22 Dec. 1999 + +; +AIMB +/ +Co +/ +Sc +1000404 to 1000430 + +• + +19 ♂♂ +, +25 ♀♀ +; +Purani +dry deciduous forest; +12°1ʹ55.2″ N +, +77°7ʹ52.68″ E +; + +11 May 1999 + +; +AIMB +/ +Co +/ +Sc +1000431 to 1000474, +cattle dung baited trap +; +Priyadarsanan Dharma Rajan +leg. + +• + +♂ +, +♀ +; +Karnataka +, +Nagarhole National Park +, +Core +moist deciduous forest; +12°1ʹ13.8″ N +, +76°6ʹ14.76″ E +; +cattle dung baited trap +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000475 to 1000476 + +• + +2 ♂♂ +, +2 ♀♀ +; +Kerala +, +Shendurney Wildlife Sanctuary +; +Rockwood +grassland; +08°51ʹ41.8ʺ N +, +77°08ʹ00.5ʺ E +; elev. + +839 m +a.s.l. + +; + +8 May 2009 + +; +open cow dung bait traps +; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000477 to 1000480 + +• + +♂ +; +Tamil Nadu +; +Thirunelveli +pasture land, +Peikulam +; + +13 Jan. 2016 + + +• + +♂ +; +Radhapuram +; + +2Apr. 2016 + +; +AIMB +/ +Co +/ +Sc +1000481 to 1000482 + +• + +3 ♂♂ +, +3 ♀♀ +; +Radhapuram +; + +20 Dec. 2016 + +, +cow dung baited pitfall trap +; +Chian +leg.; +AIMB +/ +Co +/ +Sc +1000483 to 1000488 + +• + +♂ +; +Arunachal Pradesh +, +Pasighat +, community forest, +Rani Village +; +27°58ʹ12.9324ʺ N +, +95°19ʹ20.5284ʺ E +; elev. + +125 m +a.s.l. + +; + +30 Sep. 2018 + +; +cow dung baited pitfall trap +; +AIMB +/ +Co +/ +Sc +1000489 + +• + +♀ +; community forest, +Rani Village +; +27°58ʹ10.6068ʺ N +, +95°19ʹ26.1156ʺ E +; elev. + +131 m +a.s.l. + +; + +30 Sep. 2018 + +; +cow dung baited pitfall trap +; +Harsha Malhotra +and team leg.; +AIMB +/ +Co +/ +Sc +1000490 + +• + +♀ +; +Nagaland +, +Bongkolong village +, near +Intanki National Park +, open ground; + +15 Apr. 2021 + +; hand-picked from cow dung; +Seena Narayanan Karimbumkara +leg.; +AIMB +/ +Co +/ +Sc +1000491 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=14–19, BW= 7.5–10.5. + +Male + + +Green, coppery or bronzy black, moderately shining, oval; clypeus ( +Fig. 9D +) parabolic with a feeble emargination in the middle, fine elongate granules giving it a rugulose appearance; frontal carina interrupted in the middle, a curved clypeal carina anteriorly and a conical tubercle behind, genae rather smooth; pronotum ( +Fig. 14D +) fairly strongly, closely but unevenly punctured without well marked median line, front angles not very sharp, hind angles obsolete; elytra moderately strongly striate, intervals finely and sparsely punctured; pygidium and metasternal shield feebly and sparsely punctured; sides of metasternum densely granulate and hairy. Clypeus with smooth minute granules giving it a rugose appearance; front tibia curved at the anterior tip, mid-femur with a sharp tooth near the end of lower edge, mid-tibia ( +Fig. 16D +) slender at base abruptly dilated near the middle, hind trochanter slightly toothed beneath. + + +AEDEAGUS (in mm) ( +Fig. 8E–H +). LP=1.5, Lp =1.5; BP= 1, BpB= 0.5, BpT=0.25. Phallobase slightly longer than parameres. Parameres almost straight above, curved below and at the tip, with a curved opening on the ventral side near the base. The margin of this opening where the parameres meet is broader in the middle. + + + +Fig. 13. +Aedeagus, lateral, dorsal, ventral and apical views. +A–D +. +Onitissubopacus +Arrow, 1931 +, AIMB/ Co/Sc1000711. +E–H +. + +O. virens +Lansberge, 1875 + +, AIMB/Co/Sc1000765. (Images: AIMB). + + + + +Fig. 14. +Pronotum of male specimens. +A +. + +Onitis bhomorensis + +sp. nov. +, holotype, NIM/NBAIR/ COL/ONIT/H-271023A. +B +. + +O. punctatostriatus +Janssens, 1937 + +, AIMB/Co/Sc1000492. +C +. + +O. kethai + +sp. nov. +, holotype, NIM/NBAIR/COL/ONIT/H-271023B. +D +. + +O. philemon +Fabricius, 1801 + +, AIMB/Co/ Sc1000317. +E +. + +O.visthara + +sp. nov. +, holotype, NIM/NBAIR/COL/ONIT/H-271023C. +F +. + +O. singhalensis +Lansberge, 1875 + +, AIMB/Co/Sc1000684. (Images: AIMB). + + + +Female + +Clypeus closely rugose; protibia broad, narrow with four strong teeth and an articulated spur. + + + + +Distribution + + + +India +: +Andhra Pradesh +, +Arunachal Pradesh +, +Assam +, +Bihar +, +Chhattisgarh +, +Gujarat +, +Haryana +, +Himachal Pradesh +, +Jharkhand +, +Karnataka +, +Kerala +, +Madhya Pradesh +, +Maharashtra +, +Meghalaya +, +Nagaland +, New +Delhi +, +Odisha +, Punjab, +Rajasthan +, +Tamil Nadu +, +Tripura +, +Uttar Pradesh +, +Uttarakhand +and +West Bengal +. Other countries: +China +, +Laos +, +Nepal +, +Pakistan +, +Sri Lanka +, +Taiwan +, +Thailand +and +Vietnam +( +Chandra 2000 +, 2005, 2009a; +Chandra & Ahirwar 2005 +; +Sewak 2009a +, +2009b +; + +Sabu +et al. +2011 + +; Chandra & Gupta 2012a, 2012b; Chandra +et al. +2012a, 2012b; + +Thakare +et al. +2012 + +; +Karimbumkara & Rajan 2013 +; +Chandra & Gupta 2013a +, +2013b +; + +Gupta +et al. +2014 + +, +2015 +; + +Ghosh +et al. +2020 + +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFEB89189D54F9EBFB5AF9A0.xml b/data/55/71/87/557187D2FFEB89189D54F9EBFB5AF9A0.xml new file mode 100644 index 00000000000..575d98a5c76 --- /dev/null +++ b/data/55/71/87/557187D2FFEB89189D54F9EBFB5AF9A0.xml @@ -0,0 +1,204 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis naviauxi +Cambefort, 1988 + + + + + + +Figs 4E–F +, +10D +, +19 + + +Onitisnaviauxi +Cambefort, 1988: 189 +(original description). + + + + + + +Onitis naviauxi + +– + +Bezděk & Krell 2006: 159 + +(distribution). — + +Bezděk 2016: 179 + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + +Cambefort (1988) +classified +Onitisnaviauxi +in Group III of +Janssens (1937) +. It is similar to + +O. assamensis + +in having vertex with punctures and clypeal carina being placed away from vertex. It is distinct from the latter in having shorter clypeal carina and the male protibia having 3–4 teeth on the inferomedial ridge, while in + +O. assamensis + +there are 6–7 teeth. + + +Type material examined +(based on images) + + + +Holotype + + + + +NEPAL +• +♂ +; “ +Onitisnaviauxi +n. sp. +Y. Cambefort +det. 1987, Nepal: Sauraha. Alt. 150, 19.4.82 RN”; +MNHN +EC4807 +. + + + + +Paratype + + + + +NEPAL +• +♀ +; “ +Onitisnaviauxi +n. sp. +Y. Cambefort +det. 1987, Nepal: Sauraha. Alt. 150, 19.4.82 RN”; +MNHN +EC4808 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=20–25. BW= 10–12. + +Male + +Bronzy green, shining; head rounded, slightly flattened anteriorly; clypeus with fine, sparse punctuation, slightly rugose behind; clypeal carina present, short; frontal carina narrowly interrupted in the middle, with a fairly strong sharp tubercle behind; vertex with smooth punctures; pronotum with simple, strong and dense punctures on the disc, finer, more sparse anteriorly, almost imperceptible at the sides; base not bordered, basal foveae round and placed apart; scutellum very small; elytra smooth, striae broad, not very deep, intervals minutely and sparsely punctured; pygidium convex, very minutely punctured; metasternal shield with shallow longitudinal groove, sparse fine long hairs, the sides granulate anteriorly. Clypeus reflexed, straight in the middle. Profemur with a strong tooth on the antero-median ridge, protibia short, tridentate on the infero-internal ridge, median femur unarmed, tibia toothed on the dorsal side, posterior femur with a strong tooth on the ridge below, directed towards the back, tibia bilobed at the apex. + +AEDEAGUS ( +Fig. 4E–F +). Parameres shorter than the phallobase, phallobase cylindrical, slightly curved; parameres straight above and curved below, largely notched on the underside. + + +Female + +Similar to male but more elongated head, clypeus parabolic, not much reflexed, rugose; with confluent transverse rugulosity behind the clypeal carina; vertex as in the male, with simple, very sparse punctuation; metasternum with well-marked narrow groove; legs not toothed. + + + + +Distribution + + + +Nepal +( +Cambefort 1988 +; +Bezděk & Krell 2006 +; +Bezděk 2016 +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFED891B9D6FFB94FDACF99E.xml b/data/55/71/87/557187D2FFED891B9D6FFB94FDACF99E.xml new file mode 100644 index 00000000000..966cd0a6eca --- /dev/null +++ b/data/55/71/87/557187D2FFED891B9D6FFB94FDACF99E.xml @@ -0,0 +1,409 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis lama +Lansberge, 1875 + + + + + + +Figs 10C +, +11I–L +, +19 + + +Onitislama +Lansberge, 1875: 53 +, 123 (original description). + + + + + + +Onitis lama + +– + +Preudhomme De Borre 1881: 40 + +(catalogue). — + +Arrow 1931: 389 + +(key and description). — + +Balthasar 1935: 88 + +(monograph); 1963: 44 (monograph). — + +Janssens 1937: 72 + +(revision). — + +Mittal 2005: 46 + +(distribution). — + +Bezděk & Krell 2006: 159 + +(distribution). — + +Sewak 2009a: 36 + +(diagnosis, distribution); 2009b: 58 (diagnosis, distribution). — + + +Thakare +et al. +2012: 78 + + +(distribution). — + + +Siddiqui +et al. +2014: 300 + + +(distribution). — + + +Gupta +et al. +2015: 1037 + + +(checklist). — + +Bezděk 2016: 179 + +(distribution). — + + +Ghosh +et al. +2020: 238 + + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + +Onitislama +is the sole member of Group V ( +Janssens 1937 +). It has an uninterrupted frontal carina, and a short clypeal carina. + + + + + +Type material examined + + + + +Syntypes + +( +2 specimens +) + + + +INDIA +• 2 “ +♂♂ +; + +“ +Lama +Lansb. + +India bor. Typus, Ex-Musaeo Van Lansberge, Museum Paris ex +Coll. R. Oberthur +, +A. Janssens +vid., 1936: +Onitislama +Lansb. 1875, Type, +Syntype +; Polsky, Himalaya, + +O. lama + +typ. Lansbrg., Ex musaeo +E. Harold +, Museum Paris ex +Coll. R. Oberthur +, +A. Janssens +vid., 1936: + +Onitis lama +Lansb., 1875 + +, +Syntype +”; +MNHN +EC4810 +– +EC4811 + +. + + +Additional material examined +( +9 specimens +) + + + +INDIA +• +♂ +; “Punjab, Nevinson Coll. 1918-14”; +BMNH +(E) 1237147 + +• + +♀ +; “ +Uttarakhand +, Kumaon, +W. Almora +, India. H.G.C., H.G. Champion Coll., B.M. 1953-156., + +Onitis lama, +Lansb., G.J. Arrow + +det.”; +BMNH +(E) 1237148 + +• + +♂ +; “ +Onitislama +, + +Det. J.J.E. Gillet + +; +ZSIM +• +2 ♂♂ +, +2 ♀♀ +; +Tamil Nadu +, Tirunelveli, Radhapuram”; +open cow dung bait trap +, + +20 Dec. 2016 + +, +Chian +leg.; +AIMB +/Co/Sc1000306 to 1000309 + +. + + + + +Fig. 8. +Aedeagus, lateral, dorsal, ventral and apical views. +A–D +. +Onitiskethai +sp. nov. +, NIM/NBAIR/ COL/ONIT/H-271023B. +E–H +. + +O. philemon +Fabricius,1801 + +,AIMB/Co/Sc1000317. +I–L +. + +O. singhalensis +Lansberge, 1875 + +, AIMB/Co/Sc1000684. +M–P +. + +O. visthara + +sp. nov. +, NIM/NBAIR/COL/ONIT/H- 271023C. (Images: AIMB). + + + + + +Description + + +MEASUREMENTS (in mm). TL=19–24, BW =11–13. + +Male + +Black, shining; head with an entire, strongly elevated, slightly arcuate frontal carina, clypeal carina short, curved, closer to clypeal margin; pronotum distinctly, moderately closely punctured anteriorly and extremely minutely behind; front angles blunt, sides strongly rounded, base completely margined and lobed between basal foveae; elytra with fine striae, smooth intervals, sutural intervals with few scattered punctures, posterior edge with short fringe of erect yellow hairs; pygidium smooth, opaque with very minute scanty punctures; metasternum with a narrow longitudinal groove, metasternal shield smooth in the middle and behind, flat and fairly strongly punctured anteriorly and sides, the sides of the metasternum densely punctured and hairy. Clypeus deeply excised in the middle, not very closely rugulose; posterior part of head very finely and sparsely punctured; profemur with a very sharp oblique tooth beyond the middle of its lower edge, protibia elongate, armed with four short external teeth, a strong sharp tooth on the lower face before the middle and three or four minute tubercles and an incurved apical process, hind femur rather long, straight with a tooth near the end of the lower edge, trochanter little produced at the tip and hind tibia finely serrate along its inner edge. + +AEDEAGUS (in mm) ( +Fig. 11I– L +). LP=3.035, Lp= 2.57; BP=1.43, BpB =0.93, BpT=0.36. Phallobase longer than parameres which are blunt, slightly sinuate near the base, curved inwards anteriorly and hooked at the tip. + + +Female + +Clypeus closely rugulose, its front margin entire and slightly truncate, pronotum more strongly punctured anteriorly than male, protibia broad, armed with four blunt external teeth and a small articulated terminal spur. + + + + +Distribution + + + +India +: +Andhra Pradesh +, +Gujarat +, +Haryana +, +Himachal Pradesh +, +Maharashtra +, Punjab, +Rajasthan +, +Tamil Nadu +, +Uttarakhand +, +Uttar Pradesh +. Other countries: +Nepal +, +Pakistan +( +Mittal 2005 +; +Bezděk & Krell 2006 +; +Sewak 2009a +, +2009b +; + +Thakare +et al. +2012 + +; + +Siddiqui +et al. +2014 + +; + +Gupta +et al. +2015 + +; +Bezděk 2016 +; + +Ghosh +et al. +2020 + +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFEF891D9D9AFDB1FE0CFC55.xml b/data/55/71/87/557187D2FFEF891D9D9AFDB1FE0CFC55.xml new file mode 100644 index 00000000000..e9ef6ae9fe7 --- /dev/null +++ b/data/55/71/87/557187D2FFEF891D9D9AFDB1FE0CFC55.xml @@ -0,0 +1,328 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis kethai + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +C3F62D8F-89D9-4034-A209-382401D8B861 + + + + + +Figs 7 +, +8A–D +, +9C +, +14C +, +15C +, +16C +, +18 + + + + + +Differential diagnosis + + + +Onitiskethai +sp. nov. +can be easily distinguished from + +O. philemon + +in having stronger and more closely placed punctures on the pronotum; genae smoothly rounded in + +O. kethai + +, while strongly rounded in + +O. philemon + +, both inconspicuously punctured. The clypeus is rugulose in + +O. kethai + +( +Fig. 9C +); while in + +O. philemon + +it is granular in male ( +Fig. 9D +) and rugose in female. +Onitissinghalensis +differs from both these species in having strong rounded granules on clypeus and head while the genae are sparingly granular ( +Fig. 9F +). In + +O. kethai + +the space between the clypeal carina, frontal carina and the tubercle behind is strongly rugose, while in + +O. philemon + +and + +O. singhalensis + +it is granular. Differences were also observed on comparing the aedeagus of + +O. kethai + +( +Fig. 8A–D +), + +O. philemon + +( +Fig. 8E–H +) and + +O. singhalensis + +( +Fig. 8I–L +). + + + + + +Etymology + + +Named in memory of our field assistant, the late Mr Ketha Gowda who assisted PDR in many of his field trips and helped in sampling dung beetles and other insects. + + + + +Type material examined + + + + +Holotype + + + + +INDIA +• +♂ +; +Karnataka +, +Biligiri Rangaswamy +temple +Tiger Reserve +, +Honnametti +shola; +11°54ʹ7.68″ N +, +77°11ʹ33.84″ E +; +open cow dung bait trap +; + +10 May 1998 + +, +Priyadarsanan Dharma Rajan +leg.; specimen deposited at +NBAIR +, +NIM +/ +NBAIR +/COL/ONIT/H-271023B. + + + + +Paratypes + +( +13 specimens +) + + + +INDIA +• +8 ♂♂ +, +5 ♀♀ +; +Karnataka +, +Biligiri Rangaswamy +temple +Tiger Reserve +, +Honnametti +shola; +11°54ʹ7.68″ N +, +77°11ʹ33.84″ E +; +open cow dung bait trap +; + +10 May 1998 + +; +Priyadarsanan Dharma Rajan +leg.; +AIMB +/ +Co +/ +Sc +1000293 to 1000305 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=11.5–15; BW =6–7.5; PL=4–6; PW= 6–8; EL=6.5–8; HL=3–3.5; HW=3–4.5. + +Male +( +holotype +) + + +Body, elytra, head, legs, antenna and mouthparts reddish, elytra red with a tinge of metallic green; oval, slightly convex. Head ( +Fig. 9C +) closely granular with some rugosity, hind part almost smooth; clypeus parabolic, rugulose, margin slightly reflexed; with a feeble emargination in the middle, separated from the vertex by an interrupted frontal carina with a short curved clypeal carina and a blunt tubercle behind; genae with scattered imperceptible punctures. Pronotum ( +Fig. 14C +) strongly and closely punctured in the middle, less closely at the sides, more strongly and closely pitted anteriorly, with a very slight median line starting above the basal impression, extending almost up to the anterior margin as thin smooth area; anterior sides of pronotum slightly curved, strongly rounded in the middle, bisinuate behind; front angles sharp, base bluntly angulate in the middle; basal fovea deep near the base with imperceptible punctures in circular rows at the periphery. Elytra strongly striate, striae without punctures, elytral intervals finely and sparsely punctured with the 1 +st +, 3 +rd +and the 5 +th +intervals convex, sutural interval shining while the rest opaque. Pygidium not shining, imperceptibly punctured. Metasternum with deep broad longitudinal groove in the front, metasternal shield imperceptibly punctured, with scattered aciculate punctures at the sides anteriorly; sides of the metasternum granular and thinly covered with long hairs. Protibia slightly curved with four strong external teeth and a sharp process at the tip, profemur not toothed; mesofemur ( +Fig. 16C +) plain and curved near the tibial joint where it forms a sharp tooth-like process on both upper and lower ridges; hind trochanter slightly toothed. + + + +Fig. 7. + +Onitis kethai + +sp. nov. +A–E +. Holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023B. +A +. Habitus, dorsal view. +B +. Habitus, ventral view. +C +. Head, dorsal view. +D +. Pygidium. +E +. Elytra. +F +. Paratype, ♀, habitus, dorsal view. (Images: AIMB). + + + +AEDEAGUS (in mm) ( +Fig. 8A–D +). LP= 2.5, Lp=2.2; BP=1.3, BpB=1, BpT=0.5. Phallobase longer than parameres. Parameres slightly sinuate near the base, curved inwards anteriorly and hooked at the tip. + + +Female + +Clypeus closely rugose; protibia with four strong teeth and an articulated spur; no protrusions on the legs. + + +Habitat + + + +The collections were made in the cloud forests of Honnametti in Biligiri Rangaswamy Temple Tiger Reserve, +Karnataka +, +India +. + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFF389019D5AFB71FABDF814.xml b/data/55/71/87/557187D2FFF389019D5AFB71FABDF814.xml new file mode 100644 index 00000000000..52849d356a8 --- /dev/null +++ b/data/55/71/87/557187D2FFF389019D5AFB71FABDF814.xml @@ -0,0 +1,284 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis assamensis +Biswas, 1980 + + + + + + +Figs 1 +, +4A–D +, +17 + + +Onitisassamensis +Biswas, 1980: 339 +(original description). + + + + + + +Onitis assamensis + +– + + +Gupta +et al. +2015: 1037 + + +(checklist). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + + +Onitis assamensis + +and + +O. naviauxi + +differ from each other in the number of teeth on the anteriomedian ridge of the protibia in the male. Clypeal carina short in male of + +O. assamensis + +and + +O. naviauxi + +while it is longer in female; clypeus in female pointed in both species; metasternal shield hairy in males and smooth in females. In the original description of + +O. assamensis + +the frontal carina is considered entire, but on observing the specimen in detail, it is confirmed that it is interrupted due to the placement of tubercle on it. The teeth on the infero-internal ridge in the profemur is a very prominent character of the species and the arrangement of teeth, their number and size may vary with individuals. In a specimen we collected, an extra small tooth was present near the femoro-tibial joint on one of the fore legs. + + + + + +Material examined + + + + +Holotype + + + + +INDIA +• +♀ +; +Assam +, +Kaziranga National Park +; + +21 Dec. 1972 + +; +A.K. Ghosh +leg.; from rhinoceros dung; +ZSIM +Regd. No. A +1/4397. + + + + +Fig. 1. + +Onitis assamensis +Biswas, 1980 + +, ♂ (A–D), AIMB/Co/Sc1000001; ♀ (E–F), AIMB/Co/ Sc1000009. +A +. Habitus, dorsal view. +B +. Habitus, ventral view. +C +. Protibia. +D +. Pygidium. +E +. Habitus, dorsal view. +F +. Habitus, ventral view. (Images: AIMB) + + + +Additional material +( +15 specimens +) + + + +INDIA +• +8 ♂♂ +, +7 ♀♀ +; +Assam +, +Kaziranga National Park +; + +21 May 2019 + +; +Priyadarsanan Dharma Rajan +& +Karimbumkara Seena Narayanan +leg.; hand–picked, rhinoceros dung; +AIMB AIMB +/ +Co +/ +Sc +1000001 to 1000015 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=21.5–24.8; BW =10.7–12.5; PL=8.5–11; PW= 10.5–13.5; EL=10.5– 12.5; HL= 5–6; HW=5.5–6.8. + +Male + +Body shiny green or bronzy, broadly oval. Head rounded, clypeus straight in the middle anteriorly, finely rugose with scattered punctures; clypeal carina short, rugose behind; frontal carina almost straight, narrowly interrupted in the middle by the placement of a strong conical blunt tubercle; vertex sparsely punctured, elevated towards the eyes with a depression in the middle devoid of any punctures. Pronotum with small, close dense punctures between the basal foveae, punctures larger and less close anterior to the foveae, finer and sparse anteriorly and at the sides; basal foveae oval, deep, connected by a bordered base; scutellum triangular visibly large. Elytra broadly curved at the sides; striae deep, broader at the base, narrow and smooth towards the tip; elytral intervals smooth without punctures, sutural interval slightly convex. Pygidium margined, lesser sheen, imperceptibly punctured. Metasternal shield with minute punctures, front angles with minute granules and fine yellow hairs; sides of metasternum with reddish hairs, densely granular. Fore legs with rows of reddish long hairs or setae on the ridges; a strong curved tooth on the anteromedian ridge of profemur with a minute oblique denticle near the base; protibia with four distantly placed external teeth, the infero-medial ridge with 6–7 teeth, small proximally and stouter distally; mid femur unarmed; hind femur with a strong curved lateral tooth towards the apex. + +AEDEAGUS (in mm) ( +Fig. 4A–D +). LP=3.6, Lp =2.86; BP=1.86, BpB=1.43, BpT= 0.71. Phallobase longer than parameres, broader at base and further tapers, becomes blunt and slightly curved at the tip. Phallobase slightly broader at the base. + + +Female + +Body greenish, broadly oval, not very convex; head sub triangular, frontal carina entire with a tubercle. Clypeal carina long and curved placed almost midway between the clypeal margin and the frontal carina; clypeus parabolic, rugulose. Pronotum strongly and closely punctured; front angles sharp; basal foveae rounded. Elytra: dull, striae shallow and broad, intervals flat, not perceptibly punctate. Ventral surface: metasternum flat, smooth with two deep pits behind and a shallow longitudinal groove between; sides of metasternum closely granular and hairy. Pygidium without hair, slightly shining and coarse; legs unarmed. + + + + +Distribution + + + +India +: +Assam +( +Biswas 1980 +; + +Gupta +et al. +2015 + +; +Schoolmeesters 2022 +). + + + + + +Remarks + + + +A single female specimen of + +O. assamensis + +was collected by +Biswas (1980) +from rhinoceros dung and described as the +holotype +. During our visit to Kaziranga National Park, we have collected +15 specimens +of the species of which eight were males and +seven females +. The rarity of the species mentioned by Biswas can be attributed to the winter in December when he had made the collection. The species was common and active in May (pre-monsoon), when we collected them at the same location. We have described the male of +Onitisassamensis +which had never previously been discovered or described. + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFF589039D6CFA62FA8FFBF0.xml b/data/55/71/87/557187D2FFF589039D6CFA62FA8FFBF0.xml new file mode 100644 index 00000000000..a0228346641 --- /dev/null +++ b/data/55/71/87/557187D2FFF589039D6CFA62FA8FFBF0.xml @@ -0,0 +1,627 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + +Genus + +Onitis +Fabricius,1798 + + + + + + + +Key to species of + +Onitis +Fabricius, 1798 + +from the Indian subcontinent + + + +The following key is based on external morphological characters and can be used to identify both male and female adults of + +Onitis + +. For the convenience of identification, we have retained the groups suggested by +Janssens (1937) +. The keys have been modified from the identification keys to the species provided by +Arrow (1931) +and +Balthasar (1963) +. + + + + +1. Head with frontal carina interrupted medially, with tubercle placed adjoining the carina; clypeus always with a short or long transverse carina.................................................................................... 2 + + +– Head with frontal carina uninterrupted or entire, with tubercle behind or absent; clypeal carina feebly expressed.......................................................................................................................................... 13 + + + + + +2. Body completely black or dark brown ( +GROUP I +) .......................................................................... 3 + + + + + +– Body metallic, with coppery or green tinge....................................................................................... 6 + +3. Pronotum minutely punctured ( +Fig. 5E–F +)........................................................................................ 4 + + + + + +– Pronotum strongly punctured ( +Fig. 14A–B +)...................................................................................... 5 + + + + + +4. Metasternal shield in male with deep transverse excavation in the middle; without an anterior longitudinal groove ................................................... + +O. excavatus +Arrow, 1931 + +( +Figs 5E +, +6A–D +, +18 +) + + + + +– Metasternal shield without transverse excavation in the middle, with an anterior longitudinal groove ...................................................................... + +O. falcatus +( +Wulfen, 1786 +) + +( +Figs 5F +, +6E–H +, +18 +) + + + + + + +5. Pygidium with a median longitudinal groove; protibia with a rudimentary spur before the extremity ( +Fig. 16A +) ..................................... + +O. bhomorensis + +sp. nov. +( +Figs 2 +, +6I–L +, +9A +, +14A +, +15A +, +16A +, +17 +) + + + + +– Pygidium without a median longitudinal groove; protibia without a rudimentary spur ( +Fig. 16B +).... ................................. + +O. punctatostriatus +Janssens, 1937 + +( +Figs 6M–P +, +9B +, +10F +, +14B +, +15B +, +16B +, +19 +) + + + + + + +6. Vertex covered with granules or highly raspy punctuation; clypeal carina closer to the frontal carina than to the clypeal margin ( +Fig. 9A–F +) ( +GROUP II +)........................................................................ 7 + + + + +– Vertex covered with punctures and never rough or granular; clypeal carina midway between clypeal margin and frontal carina ( +Figs 1A +, +10D +) ( +GROUP III +)................................................................ 12 + + + + + + +7. Metasternum longitudinally grooved anteriorly ( +Fig. 12F +)............................................................... 8 + + + + +– Metasternum not grooved anteriorly ( +Fig. 1B +).................................................................................11 + + + + + + +8. Pronotum without a smooth median longitudinal line; genae smooth, inconspicuously punctured ( +Fig. 14C–D +); male profemur not toothed......................................................................................... 9 + + + + +– Pronotum with a smooth median longitudinal line; genae sparingly granular ( +Fig. 14E–F +); male profemur toothed ............................................................................................................................. 10 + + + + + + +9. Clypeus granular in male and rugose in female ( +Figs 9D +, +14D +), pronotum strongly and unevenly punctured .......................... + +O. philemon +Fabricius, 1801 + +( +Figs 8E–H +, +9D +, +10E +, +14D +, +15D +, +16D +, +19 +) + + + + +– Clypeus rugulose in both sexes; pronotum strongly and closely punctured ( +Figs 9C +, +14C +)............... ................................................................. + +O. kethai + +sp. nov. +( +Figs 7 +, +8A–D +, +9C +, +14C +, +15C +, +16C +, +18 +) + + + + + + +10. Lateral margin of pronotum feebly curved; front angles of pronotum blunt ( +Figs 9F +, +14F +); mid-femur in male with a tooth near the middle of its posterior edge, another at the extremity ( +Fig. 16F +) ......... ......................................... + +O. singhalensis +Lansberge, 1875 + +( +Figs 8I–L +, +9F +, +12A +, +14F +, +15F +, +16F +, +19 +) + + + + +– Lateral margin of pronotum strongly curved; front angles of pronotum sharp ( +Figs 9E +, +14E +); mid femur in male without a tooth near the middle of the posterior edge, the extremity with a sharp tooth on the lower edge and a blunt one above ( +Fig. 16E +) ........................................................................... ....................................................... + +O. visthara + +sp. nov. +( +Figs 8M–P +, +9E +, +12E–F +, +14E +, +15E +, +16E +, +20 +) + + + + + + +11. Frontal carina narrowly interrupted; pygidium smooth with minute punctures.................................. ............................................................................. + +O. virens +Lansberge, 1875 + +( +Figs 12D +, +13E–H +, +20 +) + + + + +– Frontal carina widely interrupted; pygidium opaque with imperceptible punctures........................... ............................................................................ + +O. subopacus +Arrow, 1931 + +( +Figs 12C +, +13A–D +, +20 +) + + + + + + +12. Pronotal anterior angles not extended, blunt; male protibia with 3–4 teeth on the inferomedial ridge ........................................................................... + +O. naviauxi +Cambefort, 1988 + +( +Figs 4E–F +, +10D +, +19 +) + + + + +– Pronotal anterior angles slightly extended, sharp; male protibia with 6–7 teeth on the inferomedial ridge ( +Fig. 1C +) ........................................................ + +O. assamensis +Biswas, 1980 + +( +Figs 1 +, +4A–D +, +17 +) + + + + + + +13. Pygidium covered with short or long hair ( +GROUP VII +)............................................................... 14 + + + + + +– Pygidium not covered with hair or setae ......................................................................................... 17 + +14. Clypeus elliptical; head not tuberculate; elytral intervals slightly carinate in the middle................... ..................................................................................... + +O. feae +Felsche, 1907 + +( +Figs 10A +, +11E–H +, +18 +) + + + + +– Clypeus truncate or excised; head tuberculate; elytral intervals convex......................................... 15 + + + + +15. Clypeal margin deeply excised with an obtuse angle; frontal carina with a small gap ....................... .......................................................................................... + +O. brahma +Lansberge, 1875 + +( +Figs 5B +, +17 +) + + + +– Clypeal margin not deeply excised; frontal carina entire ................................................................ 16 + + + + + +16. Hair on the pygidium short; elytral interval angularly elevated along middle with fine close punctures................................................................................... + +O. crassus +Sharp, 1875 + +( +Figs 5D +, +18 +) + + + + +– Hair on the pygidium long; elytral interval flat with feeble punctures................................................ .................................................................... + +O. castaneus +Redtenbacher, 1848 + +( +Figs 5C +, +11A–D +, +17 +) + + + + + +17. Clypeal carina present ..................................................................................................................... 18 + + +– Clypeal carina absent....................................................................................................................... 19 + + + + + +18. First, third and fifth inter-striae of the elytra not much convex; elytra dark, metasternal shield smooth behind, with a median groove ( +GROUP V +)............ + +O. lama +Lansberge, 1875 + +( +Figs 10C +, +11I–L +, +19 +) + + + + +– First, third, and fifth inter-striae distinctly convex; elytra yellow with some intervals green; metasternal shield punctured behind, without a median groove ( +GROUP XIV +) ............................... ....................................................................................... + +O. humerosus +( +Pallas, 1771 +) + +( +Figs 10B +, +18 +) + + + + + + +19. Pronotum covered with close granules ( +GROUP IX +)......................................................................... ............................................................................ + +O. bordati +Cambefort, 1988 + +( +Figs 3 +, +4G–J +, +5A +, +17 +) + + + + +– Pronotum covered with strong punctures ( +GROUP XVIII +)............................................................... .......................................................................................... + +O. siva +Gillet, 1911 + +( +Figs 4K–N +, +12B +, +20 +) + + + + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFF989169D68F994FE37F82E.xml b/data/55/71/87/557187D2FFF989169D68F994FE37F82E.xml new file mode 100644 index 00000000000..94a60d36b4f --- /dev/null +++ b/data/55/71/87/557187D2FFF989169D68F994FE37F82E.xml @@ -0,0 +1,379 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis castaneus +Kollar, 1844 + + + + + + +Figs 5C +, +11A–D +, +17 + + +Onitiscastaneus +Kollar, 1844: 517 +(original description). + + + + + + +Onitis castaneus + +– + +Lansberge 1875: 144 + +(revision). — + +Arrow 1931: 398 + +(keys, description). — + +Balthasar 1935: 92 + +(monograph); 1963: 46 (monograph). — + +Janssens 1937: 74 + +(revision). — + +Chandra 2000: 360 + +(distribution). — + +Mittal 2005: 46 + +(list). — + +Bezděk & Krell 2006: 159 + +(describing author, year correction). — + +Chandra & Ahirwar 2007: 287 + +(distribution). — + +Sewak 2009b: 61 + +(distribution, diagnosis). — Singh +et al. +2010: 1394 (distribution). — + + +Gupta +et al. +2015: 1037 + + +(checklist). — + +Bezděk 2016: 179 + +(describing author, year correction, distribution). — + + +Ghosh +et al. +2020: 238 + + +(distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + +Onitiscastaneus +is similar to + +O. brahma + +and + +O. crassus + +in having a hairy pygidium and it differs from them in having tubercle on the clypeo-frontal carina while the tubercle is placed behind the clypeo-frontal carina in both the latter species. It differs from + +O. brahma + +in the frontal carina being entire; the clypeal margin being straight while the carina is interrupted and the clypeal margin is slightly excised in + +O. brahma + +. +Onitiscastaneus +differs from + +O. crassus + +in having longer hairs on the pygidium while the latter has short setae. + + + + + +Material examined + + + + +Syntype + +( +3 specimens +) + + + +INDIA +• +♂ +, +2 ♀♀ +; “Kaschmir, Hügel”; +NHMW + +. + + +Additional material +( +17 specimens +) + + + +INDIA +• +♂ +; “ +Kheri Forest +, +U.P. India. +, Jan’ 16 HGC; +H.G. Champion Coll. +, B.M. 1953-156.; + +Onitis castaneus +, Redt. G. J. Arrow + +det.”; +BMNH +(E) 1237151 + +• + +♀ +; “ +Gazighat +, +Multan Div. Punjab +, 15. 2. 28., +C.F.C. Beeson +; ex coll. +Dehradun, B.M. +1928-472., ex Cattle dung;” +BMNH +(E) 1237152 + +• + +♀ +; “ +Pilani +, +Rajasthan +; 1957; Coll: +H.L. Khandu +, Det: +T. G. Vazirani +”; +ZSIM + +• + +10 ♂♂ +, +4 ♀♀ +; +Assam +, +Tezpur +, under +Kolia +bhomora bridge; hand-picked, cattle dung; + +19 Dec. 2018 + +; +Rajkamal Goswami +leg.; +AIMB +/ +Co +/ +Sc +1000023 to 1000036 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=16–19; BW=9.5–12; PL=6.5–7.5; PW=9–11; EL= 8.5–10; HL=4. 5–5.5; HW =5–6. + +Male + +Dark chestnut brown; head with strongly elevated frontal carina acuminate in the middle; clypeal margin strongly reflexed, straight in the middle, with long transverse arcuate carina, frontal carina straight, rugulose in front, with tubercle behind; vertex strongly excavated; pronotum unevenly punctured, strongly rugulose anteriorly, tuberculate between the basal foveae; front angles rather blunt, deep basal foveae closely placed; elytra opaque, finely striate, intervals flat, feebly punctured; pygidium sparsely hairy; metasternum closely tuberculate, hairy with a longitudinal median groove; front legs long, slender, strongly curved, armed with four short teeth externally, profemur with a strong oblique spine near the middle of anterior edge, middle tibia with two strong processes at its outer edge. + +AEDEAGUS (in mm) ( +Fig. 11A–D +). LP=3.03, Lp =2.78; BP=1.32, BpB=1.07, BpT=0.43. Parameres shorter than phallobase, dorsal side of parameres straight at the base, tip broader and curved and tip blunt. + + +Female + +Protibia short, broad bearing four strong external teeth; clypeus more rounded in the middle than male; frontal tubercle more prominent. + + + + +Distribution + + + +India +: +Arunachal Pradesh +, +Assam +, +Bihar +, +Gujarat +, +Haryana +, +Himachal Pradesh +, +Jammu and Kashmir +, +Madhya Pradesh +, +Meghalaya +, Punjab, +Rajasthan +, +Uttarakhand +, +Uttar Pradesh +, +West Bengal +. Other countries: +Nepal +; +Pakistan +( +Chandra 2000 +; +Mittal 2005 +; +Bezděk & Krell 2006 +; +Chandra & Ahirwar 2007 +; +Sewak 2009b +; Singh +et al. +2010; + +Gupta +et al. +2015 + +; +Bezděk 2016 +; + +Ghosh +et al. +2020 + +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFFA89099D5CF9AEFBA3FA55.xml b/data/55/71/87/557187D2FFFA89099D5CF9AEFBA3FA55.xml new file mode 100644 index 00000000000..8c68157cef0 --- /dev/null +++ b/data/55/71/87/557187D2FFFA89099D5CF9AEFBA3FA55.xml @@ -0,0 +1,346 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis brahma +Lansberge, 1875 + + + + + + +Figs 5B +, +17 + + +Onitisbrahma +Lansberge, 1875: 142 +(original description). + + +Onitisbrahma +– +Preudhomme De Borre 1881: 41 +(catalogue). — +Arrow 1931: 399 +(keys, description). — +Janssens 1937: 74 +(revision). — +Balthasar 1963: 45 +(monograph). — +Chandra 2000: 360 +(distribution). — +Chandra & Ahirwar 2005: 149 +(distribution); 2007: 286 (distribution). — +Sewak 2009b: 61 +(distribution, diagnosis). — Chandra +et al. +2011: 66 (distribution). — +Chandra & Gupta 2012b: 817 +(distribution). — +Karimbumkara & Rajan 2013: 174 +(distribution). — + +Gupta +et al. +2014: 230 + +(distribution); 2015: 1034 (key, distribution, description). — +Schoolmeesters 2022 +(online catalogue). + + + + +Fig. 4. A–D +. + +Onitis assamensis +Biswas, 1980 + +, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). +E–F +. + +O. naviauxi +Cambefort, 1988 + +, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). +G–J +. + +O. bordati +Cambefort, 1988 + +, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). +K–N +. + +O. siva +Gillet, 1911 + +, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB). + + + + +Fig. 5. A +. + +Onitis bordati +Cambefort, 1988 + +, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). +B +. + +O. brahma +Lansberge, 1875 + +, ♂, BMNH (E) 1237157, (image: BMNH). +C +. + +O.castaneus +Redtenbacher, 1848 + +, syntype, (image: Harald Schillhammer, NHMW). +D +. + +O. crassus +Sharp, 1875 + +, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). +E +. + +O. excavatus +Arrow, 1931 + +, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). +F +. + +O. falcatus +( +Wulfen, 1786 +) + +, ♂, AIMB/Co/ Sc1000070, (image: AIMB). + + + + + +Differential diagnosis + + + +Onitisbrahma +belongs to Group VII and is closer to + +O. crassus + +and + +O. castaneus + +in having a hairy pygidium and punctured pronotum but differs from both in having an interrupted frontal carina and strongly excised clypeus. + + + + + +Material examined + + + + +Holotype + + + + +INDIA +• +♂ +; “Mniszech, Ex-Musaeo Van Lansberge, Muséum Paris ex +Coll. R. Oberthür +1952, +A. Janssens +vid., 1936: +Onitisbrahma +Lansb.”, +MNHN +EC4809 +. + + + +Additional material +( +4 specimens +) + + + +INDIA +• +♂ +, +♀ +; Dharwar, Bombay, “ +H. Swale. +1913–117”, +BMNH +(E) 1237157, +BMNH +(E) 1237158 + +• + +♂ +, +♀ +; +Nerbuda Survey +, +Barra +, +Bewa State +, + +8 Mar. 1927 + +; + +2700 ft +a.s.l. + +; +ZSIM + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=21–26; BW=11–13.5. + +Male + +Black or dark brown; clypeus shining, not closely granulate; clypeal margin strongly reflexed, excised in the middle; head finely granulate; frontal carina interrupted in the middle; pronotum shining, pronotum rugose anteriorly and punctured behind; elytra deeply striate, intervals convex, unpunctured, not shining; pygidium sparingly punctured; scanty hairs on pygidium long; metasternum with close reddish hairs, a raised longitudinal ridge in the middle; fore legs elongate, femur bears a blunt anterior process near the extremity, tibia slender, strongly curved towards the tip, armed with four external teeth, a very sharp tooth beneath, before the middle and a long incurved terminal process; mesofemur with a sharp tooth near the middle of its lower edge; hind femur with a sharp outwardly directed tooth at its upper edge near the base and a blunt lobe before middle of the lower edge. + +Female + +Clypeus more densely granular, not shining; pronotum closely rugose on major part. + + + + +Distribution + + + +India +: +Andhra Pradesh +, +Chhattisgarh +, +Gujarat +, +Karnataka +, +Madhya Pradesh +, +Maharashtra +, +Rajasthan +, +Tamil Nadu +( +Chandra 2000 +; +Chandra & Ahirwar 2005 +, +2007 +; +Sewak 2009b +; +Chandra & Gupta 2012b +; +Karimbumkara & Rajan 2013 +; + +Gupta +et al. +2014 + +, +2015 +; +Schoolmeesters 2022 +). + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFFC890A9D5EFC0AFEA2FA53.xml b/data/55/71/87/557187D2FFFC890A9D5EFC0AFEA2FA53.xml new file mode 100644 index 00000000000..96f450b10e5 --- /dev/null +++ b/data/55/71/87/557187D2FFFC890A9D5EFC0AFEA2FA53.xml @@ -0,0 +1,327 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis bordati +Cambefort, 1988 + + + + + + +Figs 3 +, +4G–J +, +5A +, +17 + + +Onitisbordati +Cambefort, 1988: 190 +(original description). + + + + + + +Onitis bordati +– + + +Kabakov&Napolov1999:70 + +(distribution).— + + +Hanboonsong +et al. +1999:466 + + +(distribution). — +Hanboonsong & Masumoto 2000 +. — 103, 108 (key, distribution). — +Schoolmeesters 2022 +(online catalogue). + + + + + + +Differential diagnosis + + + +Onitisbordati +belongs to Group IX and it resembles species of Group XVIII in having an uninterrupted frontal carina. It resembles + +O. siva + +, in the absence of clypeal carina and differs from it in the pronotum being densely granular. + + + + + +Material examined + + + + +Holotype + + + + +VIETNAM +• +♂ +; Djiring, Annam; +11°35ʹ N +, +108°05ʹ E +; +Coll. H. Perrot +; +Coll. P. Bordat +leg.; +Y. Cambefort +det. 1987; +MNHN +, +EC1187 +. + + + +Additional material +( +8 specimens +) + + + +THAILAND +• +♀ +; “Prov. +TAK Region +de UMPHANG VI; + +1 Jun. 1991 + +– + +6 Jun. 1991 + +; Y. CAMBEFORT det. 1992”; +MNHN +, +EC1185 + +. + + + +INDIA +• +2 ♂♂ +, +5 ♀♀ +; +Meghalaya +, +Nongkhyllem +; +25°54ʹ56.51″ N +, +91°46ʹ4.55″ E +; elev. + +269 m +a.s.l. + +; + +20 May 2012 + +; +Karimbumkara Seena Narayanan +& +Rajkamal Goswami +leg.; hand-picked, elephant dung; +AIMB +/ +Co +/ +Sc +1000016 to 1000022 + +. + + + + + +Description + + +MEASUREMENTS (in mm). TL=23.5–32; BW =11.5–15.5; PL=9–12; PW=11.5–15.5; EL=11.5–15; HL=6–9; HW= 7–8.5. + +Male +( +Fig. 5A +) + +Body including legs and antennae reddish black, except the antennal club which is reddish yellow; moderately shining, elongate oval, slightly convex, devoid of any setae on the dorsal surface except at the lower margin of the elytra where a few scattered setae are present. Head with clypeus elongate, strongly rounded, slightly reflexed anteriorly, clypeal carina absent; genae strongly curved, extended outwards and upwards anteriorly, with fine granules; vertex significantly depressed, bordered behind, finely, densely granulate. Pronotum smoothly margined on all sides, the sides of pronotum straight anteriorly, slightly sinuate behind; front angles sharp, obtuse, hind angles obsolete; base angulate in the middle; basal foveae shallow, deeper towards the base, with small ovate granules both within and between them; rows of spindle shaped granules present around the foveae which are not very close, closer and longer in the middle of the pronotum, squamiform and closer anteriorly, the granules smaller, ovate, not very close in the lateral margins; a trace of longitudinal groove extends as a slight depression from the front of the foveae to the front margin of the pronotum; lateral foveae with fine close ovate granules; scutellum very small. Elytra finely striate, striae not perceptibly punctured; intervals flat, opaque, with scattered imperceptible punctures. The sutural interval slightly convex, shining with visible fine scattered punctures; the 5th interval with minute, close, slightly rugose punctures. The lateral carina of the elytra sharp. Pygidium without any hair or setae, opaque, rugose with minute scattered granules. Metasternal shield closely granular and densely hairy at the sides, less granular and hairy in the middle, smooth with imperceptible granules anteriorly and at the base. Sides of the metasternum finely, closely and evenly granular with dense hairs; the hairs shorter than those on the shield. Front legs elongate, tibia slender, strongly curved towards the end with the tip produced into a finger-like process, terminal spur absent, teeth short and triangular; lower side of protibia bears a tooth almost halfway from the base, a shorter one near the base and another short sharper one between these two, the inner side serrate. A tuft of short setae is present in an oval groove near the base of profemur, the upper margins of the legs fringed. The mesofemur is with a sharp tooth just beyond the middle of the lower edge, the hind femur without any teeth. The femoral teeth may not be prominent in minor males. + + +Fig. 3. + +Onitis bordati +Cambefort, 1988 + +, ♀, AIMB/Co/Sc1000018. +A +. Habitus, dorsal view. +B +. Habitus, ventral view. +C +. Head, dorsal view. +D +. Pygidium. (Images: AIMB). + + + +AEDEAGUS (in mm) ( +Fig. 4G–J +). LP= 3.5–4.5, Lp=4.5–5; BP=2, BpB =1.5, BpT=0.5. Phallobase straight. Parameres longer than the phallobase, broader at the base and further tapers and bluntly curved above at the tip. + + +Female +( +Fig. 3A–D +) + +Body oval, not as convex as male; clypeus elongate, strongly elliptical, slightly reflexed anteriorly, closely rugose. The posterior carina curved inwards, with a laterally compressed conical tubercle in the middle behind. Pygidium more shining than in male, rugose with imperceptible scattered granules. Profemur not toothed, tuft of short setae near the base absent. Protibia with an articulated terminal spur; teeth blunt, closer to each other than in male. + + + + +Distribution + + + +India +: +Meghalaya +, other countries: +Vietnam +, +Thailand +( + +Hanboonsong +et al. +1999 + +; +Kabakov & Napolov 1999 +; +Hanboonsong & Masumoto 2000 +; +Schoolmeesters 2022 +). + + + +Habitat + + + +In +India +, this species was collected from bamboo dominated secondary forest near Nonkhyllem Wildlife Sanctuary, +Meghalaya +. + + + + + +Remarks + + + +Cambefort (1988) +did not describe the female of +Onitisbordati +, but a female of the species deposited at MNHN was determined by Cambefort (1992) (https://shorturl.at/v4Dlr). In this paper, we provide a description for the female of + +O. bordati + +as +Hanboonsong & Masumoto (2000) +mentioned only some morphological characters. This is a new distribution record of the species from +India +and the Indian subcontinent. + + + + \ No newline at end of file diff --git a/data/55/71/87/557187D2FFFE890C9D72FEB8FABFFCFC.xml b/data/55/71/87/557187D2FFFE890C9D72FEB8FABFFCFC.xml new file mode 100644 index 00000000000..746c9c148ce --- /dev/null +++ b/data/55/71/87/557187D2FFFE890C9D72FEB8FABFFCFC.xml @@ -0,0 +1,275 @@ + + + +Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India + + + +Author + +Karimbumkara, Seena Narayanan +A6B50103-8D6D-4977-9111-EF8BA4AA0477 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +seena.narayanan@atree.org + + + +Author + +Priyadarsanan, Dharma Rajan +03907E93-F783-4720-9F7B-CB662E7FC745 +Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India. +priyan@atree.org + +text + + +European Journal of Taxonomy + + +2024 + +2024-09-10 + + +956 + + +1 +61 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287 + +journal article +10.5852/ejt.2024.956.2657 +2118-9773 +9DCDD521-909D-4436-8495-086A11C50DA0 + + + + + + +Onitis bhomorensis + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +7D70D754-D82E-4BCA-A3C9-D42645B4F708 + + + + + +Figs 2 +, +6I–L +, +9A +, +14A +, +15A +, +16A +, +17 + + + + + +Differential diagnosis + + + +Onitisbhomorensis +sp. nov. +is assigned to Group I as its body is completely black and the frontal carina is interrupted. It is similar to + +O. punctatostriatus + +in having broad elytral striae. It differs from the latter in the nature of pronotal punctures which are stronger in + +O. bhomorensis + +than in + +O. punctatostriatus + +; basal foveae are more spaced in the former than the latter; metasternal shield slightly convex, granular anteriorly with reddish hairs in + +O. bhomorensis + +while in + +O. punctatostriatus + +it is flat in the middle, with a weak longitudinal groove and with smooth scattered punctures. Clypeal margin truncated, straight in the middle in the former, while it is slightly reflexed in the latter. The presence of a median groove on the pygidium and the rudimentary spur on the protibia make it different from all other Oriental + +Onitis +species. + +Onitisbhomorensis +, is compared with + +O. assamensis + +, a species described from a nearby locality as that of the former by +Biswas (1980) +based on a single specimen. + +O. bhomorensis + +is smaller and the body is reddish black, while + +O. assamensis + +is greenish. The frontal carina is entire with a tubercle in the former while the carina is interrupted in the latter. The protibia of + +O. bhomorensis + +is with a rudimentary spur and the pygidium is with a vertical groove, both these characters are absent in + +O. assamensis + +. + + + + + +Etymology + + + +Named after the bridge “Kolia Bhomora” across Brahmaputra – the widest river in +India +, near which the +type +specimen was collected. + + + + + +Material examined + + + + +Holotype + + + + +INDIA +• +♀ +; +Assam +, +Tezpur +, +Kolia +bhomora; + +10 Oct. 2012 + +; +26°34ʹ56.66ʺ N +, +92°51ʹ52.65ʺ E +; elev. + +74 m +a.s.l. + +; +Karimbumkara Seena Narayanan +leg.; hand-picked, cattle dung; +NBAIR +, +Bangalore +; +NIM +/ +NBAIR +/COL/ONIT/H-271023A. + + + + + + +Description + + +MEASUREMENTS (in mm). TL=18; BW= 10; PL=7; PW=11; EL=10; HL=5; HW=5.1. + +Male +( +holotype +) + + +Body narrow, elongate oval, reddish black, not very shining, slightly convex. Antennae red with the club reddish yellow, legs reddish black; body devoid of setae on the dorsal side, ventral side covered with reddish hairs except on the middle of metasternum and the abdominal sclerites. Head shining with the clypeus truncate ( +Fig. 9A +), almost straight anteriorly and at the sides, strongly rugose; clypeal carina present, slightly elevated and curved. The frontal carina is broadly interrupted in the middle with a blunt conical tubercle behind. The vertex punctuation granular, closer in the middle and scattered at the sides appears rugose behind the tubercle. Genae smooth, curved, extended slightly outwards and upwards anteriorly; inner margin slightly elevated and continues behind as a posterior carina. Pronotum moderately convex, sides smoothly margined, except the base which is slightly angulate in the middle; with strong close punctures, closer between the basal foveae; uneven in the middle; deeper and closer anteriorly. Pronotal front angles not very sharp, almost right-angled, the sides straight anteriorly, rounded in the middle, bisinuate behind; hind angles obsolete. A pair of deep basal foveae present at the base on either side of the middle separated by twice its length. Lateral foveae smooth with small punctures in the periphery. Elytra finely and broadly striate, striae devoid of punctures; intervals flat, opaque; sutural interval convex, shining with visible fine scattered punctures. The lateral carinae straight, with minute visible punctures. Pygidium ( +Fig. 2F +) without hair or setae, slightly shining with imperceptible scattered punctures. A smooth median longitudinal groove extending from base to almost the middle is present. Metasternal shield shining, ( +Fig. 2E +) with minute punctures in the middle; anterior part of the metasternal shield slightly convex, with uneven granules and reddish hairs. Sides of the metasternum closely and evenly granular, covered with smooth short reddish hairs. Protibia slender, bears four external teeth, the tip not very long or curved, with a rudimentary spur near the tip ( +Figs 2D +, +16A +); teeth short, triangular, placed almost equidistantly. The inner margins of femur and tibia fringed. Femur without protrusion on any of the legs. + + + +Fig. 2. +Onitisbhomorensis +sp. nov. +, holotype, ♂, NIM/NBAIR/COL/ONIT/H-271023A. +A +. Habitus, dorsal view. +B +. Habitus, ventral view. +C +. Head and prontum. +D +. Protibia, arrow showing rudimentary spur. +E +. Metasternum. +F +. Pygidium, arrow showing longitudinal groove. (Images: AIMB). + + + +AEDEAGUS (in mm) ( +Fig. 6I–L +). LP=3, Lp=2.5; BP=1.4, BpB =0.7, BpT=0.4. Parameres almost same length as phallobase, broader at the base and further tapers become slender and slightly curved at the tip. Phallobase slightly broader at the base. + + +Female + +Unknown. + + +Habitat + + + +The specimen was collected from cattle dung on the river bed of the Brahmaputra, where the vegetation is dry seasonal grasslands, typical of Brahmaputra flood plains dominated by +Ziziphusjujuba +trees. + + + + \ No newline at end of file diff --git a/data/64/E7/C1/64E7C11C564251B3B50CBA1B151C5937.xml b/data/64/E7/C1/64E7C11C564251B3B50CBA1B151C5937.xml new file mode 100644 index 00000000000..4f265b4bf7e --- /dev/null +++ b/data/64/E7/C1/64E7C11C564251B3B50CBA1B151C5937.xml @@ -0,0 +1,837 @@ + + + +Description of a new species of the genus Oreonectes (Teleostei, Cypriniformes, Nemacheilidae) from Guangxi, China + + + +Author + +Zhong, Jia-Hong +Key Laboratory of Wildlife Evolution and Conservation in Mountain Ecosystem of Guangxi, Nanning Normal University, Nanning 530001, Guangxi, China + + + +Author + +Yang, Jian +0000-0001-7115-6713 +Key Laboratory of Wildlife Evolution and Conservation in Mountain Ecosystem of Guangxi, Nanning Normal University, Nanning 530001, Guangxi, China + + + +Author + +Mo, Hao-Lin +Key Laboratory of Wildlife Evolution and Conservation in Mountain Ecosystem of Guangxi, Nanning Normal University, Nanning 530001, Guangxi, China + + + +Author + +Chen, Wei-Cai +0000-0002-2398-4079 +Key Laboratory of Wildlife Evolution and Conservation in Mountain Ecosystem of Guangxi, Nanning Normal University, Nanning 530001, Guangxi, China + +text + + +Zoosystematics and Evolution + + +2024 + +2024-09-10 + + +100 + + +4 + + +1259 +1267 + + + +journal article +10.3897/zse.100.125866 +29F191C5-30BF-491B-8B59-5EC344BD2584 + + + + + +Oreonectes zhangi +Zhong, Yang & Chen + +sp. nov. + + + + +Figs 4 +, +5 +, +6 +, +7 +, +8 +, +10 +, +Tables 3 +, +4 + + + + +Materials. + + + + + +Holotype + +. + +• +NNNU 2023100203 +(Fig. +4 +); +71.1 mm +TL +, +63.1 mm +SL +; collected by +Jia-Hong Zhong +on + +2 October 2023 + +; in +Mt. Hanshan +, +Xinye County +, +Yulin City +, +Guangxi +, +China +; ( + +22.7364 ° N +, +110.0605 ° E + +; elevation + +199 m + +; Fig. +1 +) + +. + + + +Paratypes + +. + +• +14 specimens +; +NNU 2023100201 +–202, 2023100204–215; were collected by +Jia-Hong Zhong +from the same locality and at the same time as the holotype + +. + + + + + + +Dorsal, lateral, ventral, and living views of + +Oreonectes zhangi + +sp. nov. +NNNU 2023100203, holotype, 71.1 mm +TL +, 63.1 mm +SL +. + + + + + +Etymology. + + +The species was named in honor of the three Zhang brothers, who assisted local residents in locating water during an ancient drought. Following their passing, they were revered as rain deities by the community ( +Li 2014 +). We suggest the common Chinese name “ 张氏岭鳅 ” (Zhang Shi Ling Qiu). + + + + +Comments. + + + +Oreonectes zhangi + +sp. nov. +is classified within the genus + +Oreonectes + +based on molecular phylogenetic analyses and genus-specific characteristics, including narrowly separated anterior and posterior nostrils, elongated barbel-like anterior nostrils longer than the depth of the nostril tube (Fig. +5 +), having an epural in the caudal skeleton (Fig. +7 +), and a rounded caudal fin with 7 branched dorsal-fin rays. ( +Du et al. 2023 +; +Ito 2024 +) + + + + + + + +Oreonectes zhangi + +sp. nov. +, dorsal view of head., NNNU 2023100203, holotype, 63.1 mm +SL +. + + + + + +Diagnosis. + + +Comparative data between + +Oreonectes zhangi + +sp. nov. +and all seven known species within the genus + +Oreonectes + +are provided in Table +3 +. + +Oreonectes zhangi + +sp. nov. +can be distinguished from + +O. guilinensis + +, + +O. luochengensis + +, + +O. guananensis + +, + +O. andongensis + +, and + +O. polystigmus + +by the reduced posterior chamber of air-bladder (vs. developed), from + +O. platycephalus + +by six branched plevic-fin rays (vs. 7), seven branched dorsal-fin rays (vs. 6), from + +O. damingshanensis + +by six branched pelvic-fin rays (vs. 7), the tip of the male pelvic fin not reaching the anus (vs. exceeding) (Fig. +8 +), vertebrae 4 + 32 (vs. 4 + 34), head depth at nape (59.6–68.1 % +HL +vs. 46.7–54.6 %), body depth at dorsal-fin origin (14.0–16.1 % +SL +vs. 11.1–13.3 %). + + + + + + + +Oreonectes +species + +and comparison of identifying characteristics between the new species and its congeners. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
-Body pigmentationDorsal-fin raysPectoral-fin raysPelvic-fin raysAnal-fin raysCaudal-fin raysPosterior chamber of air-bladder
+ +Oreonectes zhangi + +sp. nov. +Presentiii, 7i, 8–9i, 6iii, 515–16Reduced
+ +O. damingshanensis + +Presentiii, 7i, 9i, 7iii, 514–15Reduced
+ +O. platycephalus + +Presentiii, 6i, 9–11i, 7iii, 513–15Reduced
+ +O. guilinensis + +Presentii, 7i, 10–11i, 6iii, 513–14Developed
+ +O. luochengensis + +Absentiii, 7i, 11–12i, 7ii, 514–16Developed
+ +O. guananensis + +Presentiii, 7i, 10–11i, 7–8iii, 513–17Developed
+ +O. andongensis + +Presentiii, 7i, 9–10i, 6iii, 513–14Developed
+ +O. polystigmus + +Presentiii, 6–7i, 9i, 6ii, 514–15Developed
+ + + + +Description. + + +Morphometric data are given in Tables +3 +, +4 +. Body elongated and cylindrical, with insignificant depth decreasing from dorsal-fin origin to caudal-fin base. Head short, head length 18.6–20.2 % of standard length, slightly depressed and flattened, width greater than depth (head width / head depth = 1.1–1.5), head depth 59.6–68.1 % of head length. Snout round, oblique, and flat, length 38.2–48.8 % of head length ( +HL +). Mouth inferior, curved, upper, and lower lips have shallow wrinkles and small spinous processes; lower lip with V-shaped median notch (Fig. +6 +). Three pairs of barbels, long: inner rostral barbel 35.4–44.1 % of +HL +, extending backward, not reaching anterior margin of eye; outer rostral barbel 49.0–67.6 % of +HL +, extending backward beyond posterior margin of eye. Maxillary barbel 38.4–52.2 % of +HL +, tip not reaching posterior margin of gill cover. Anterior and posterior nostrils are separated by a short distance, 16.8 % – 36.8 % of eye diameter. Anterior nostril tube short, with elongated, short barbel-like tip. Eyes normal, diameter 11.9–17.1 % of +HL +. Gill opening small; gill rakers not developed; nine inner gill rakers on first gill arch (n = 2). + + + + + + +Comparison of measurable traits between + +Oreonectes zhangi + +sp. nov. +and similar species, + +O. damingshanensis + +. The numbers in parentheses are the mean ± standard deviation (SD). + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
- + +Oreonectes zhangi + +sp. nov. + +O. damingshanensis +
Number of specimens1510
+TL +(mm) +59.8–81.8 (71.0 ± 8.3)69.8–85.7 (78.9 ± 4.5)
+SL +(mm) +49.8–67.6 (58.8 ± 7.1)56–70.6 (65.4 ± 4.6)
+ +Percentage of +SL +(%) + +
Body depth at dorsal-fin origin14.0–16.1 (14.7 ± 0.7)11.1–13.3 (12.2 ± 0.7)
Body width at dorsal-fin origin9.3–12.8 (11.3 ± 1.1)8.1–10.1 (9.1 ± 0.7)
Lateral head length18.6–20.2 (19.3 ± 0.7)18.5–21.2 (19.7 ± 1.1)
Predorsal length56.2–59.4 (58.0 ± 1.2)56.8–60.5 (58.3 ± 1.1)
Dorsal fin length16.7–21.1 (17.7 ± 1.4)16.9–20.7 (18.9 ± 1.3)
Dorsal fin base length7.2–9.7 (8.7 ± 0.8)8.2–11.2 (9.5 ± 1.0)
Prepelvic length48.1–50.8 (49.1 ± 1.0)48.7–52.6 (50.7 ± 1.1)
Pelvic fin length16.1–17.7 (16.7 ± 0.7)14.4–18.3 (15.8 ± 1.2)
Pelvic fin base length3.9–4.9 (4.3 ± 0.3)2.8–4.4 (3.5 ± 0.5)
Preanus length67.4–72.1 (69.3 ± 1.5)67.0–70.8 (69.2 ± 1.0)
Preanal length67.7–78.7 (73.8 ± 0.4)73.8–78.3 (76.0 ± 1.8)
Anal fin length15.1–19.5 (16.7 ± 1.5)14.8–20.9 (17.4 ± 1.6)
Anal fin base length6.6–9.0 (7.4 ± 0.8)6.9–9.5 (7.8 ± 0.8)
Prepectoral length18.3–21.2 (19.7 ± 1.1)17.6–27.7 (21.6 ± 2.6)
Pectoral fin length15.9–18.9 (17.3 ± 0.9)15.6–19.1 (17.1 ± 1.1)
Pectoral fin base length4.4–5.3 (4.8 ± 0.3)3.3–4.6 (3.9 ± 0.4)
Caudal peduncle length13.3–19.9 (16.1 ± 1.9)14.3–18.3 (16.3 ± 1.0)
Caudal peduncle depth11.5–12.5 (11.9 ± 0.3)9.2–11.8 (10.4 ± 0.8)
+ +Percentage of +HL +(%) + +
Head depth at nape59.6–68.1 (63.2 ± 2.9)46.7–54.6 (51.2 ± 2.6)
Maximum head width70.4–89.3 (82.2 ± 5.3)71.9–83.2 (76.3 ± 3.7)
Snout length38.2–48.8 (41.8 ± 3.1)37.0–44.7 (39.5 ± 2.6)
Outer rostral barber length49.0–67.6 (57.9 ± 5.2)50.0–68.2 (56.3 ± 5.6)
Maxillary barber length38.4–52.2 (46.3 ± 4.7)37.1–46.4 (41.9 ± 3.6)
Inner barber length35.4–44.1 (38.8 ± 2.8)37.1–46.4 (41.9 ± 3.1)
Eye diameter11.9–17.1 (14.6 ± 1.8)10.2–17.7 (14.5 ± 2.4)
Interorbital width29.2–52.5 (44.2 ± 6.7)39.9–47.5 (43.5 ± 2.2)
Distance between anterior nostrils25.4–34.4 (31.9 ± 2.8)25.5–30.6 (28.6 ± 1.8)
Distance between posterior nostril29.6–41.9 (37.2 ± 3.8)30.5–34.6 (33.3 ± 1.4)
+ + + + + + + +Oreonectes zhangi + +sp. nov. +, NNNU 2023100203, holotype, 71.1 mm +TL +, 63.1 mm +SL +. The ventral view of the mouth ( +A +) and close-ups of the upper ( +B +) and lower ( +C +) lips are shown; the black arrow indicates the position of the small spinous processes. + + + +Dorsal-fin rays iii, 7, pectoral-fin rays i, 8–9, pelvic-fin rays i, 6, anal-fin rays iii, 5, branched caudal fin rays 15–16. Dorsal fin short, length 16.7 % – 21.1 % of +SL +, distal margin round, origin posterior to pelvic-fin insertion, situated slightly posterior to two-thirds distance between snout tip and caudal-fin base. Pectoral fin short, length 15.9 % – 18.9 % of +SL +. Pelvic fin length 16.1–17.7 % of +SL +; tips of pelvic fin not reaching anus; distance between tips of pelvic fin and anus 0.7 times eye diameter. Anal fin short, length 15.1 % – 19.5 %, origin short distance from anus. Caudal fin rounded, length 13.3 % – 19.9 % of +SL +, length greater than depth (caudal peduncle length / caudal peduncle depth = 1.1–1.7). vertebrae 4 + 32 (n = 4), having an epural in the caudal skeleton (Fig. +7 +). + + + + + + +The caudal fin skeleton of + +Oreonectes zhangi + +sp. nov. +, NNNU 2023100202, paratype, 64.2 mm +SL +, pc 1, first preural centrum; pc 2, second preural centrum; h, hypural; hs, haemal spine and arch; ph, parhypural; pls, pleurostylar. + + + + + + + +Gonadal structure of + +O. damingshanensis + +( +A +), NNNU 2024052007, 56 mm +SL +, and + +O. zhangi + +sp. nov. +( +B +), NNNU 2023100215, paratype, 43.2 mm +SL +; the black arrow refers to the position of the anus. + + +Body completely covered by scales, except head; lateral line incomplete, with 7–13 pores, last one not reaching above tip of pectoral fin; cephalic lateral-line system, with 8 supraorbital pores, 4 + 9–10 infraorbital pores, three supratemporal canal pores, and 5–8 preoperculo-mandibular canal pores. + +U-shaped stomach with straight intestines (Fig. +9 +). Anterior chamber covered by dumbbell-shaped bony capsule, and posterior chamber reduced, reaching the vertical of the pectoral-fin base (Fig. +10 +). + + + + + + + +Oreonectes zhangi + +sp. nov. +, NNNU 2023100202, paratype, 64.18 mm +SL +; digestive tract. + + + + + + + +Air-bladder of + +Oreonectes zhangi + +sp. nov. +: a, anterior chamber; b, tube connecting anterior and posterior chambers of the air-bladder; c, posterior chamber. + + + + + +Coloration. + + +The body is mainly light yellowish brown in color, with dark brown extending from the front of the eyes to the outer rostral barbel. The base of the caudal fin is black, with each fin showing a light yellow hue. When preserved in 10 % formalin, the body color fades to a dark brown shade (Fig. +4 B +). + + + + +Sexual dimorphism. + + +Males have an oval genital papilla located immediately posterior to the anus (Fig. +8 B +), which is unclear in females (Fig. +4 C +). + + + + +Distribution, habitat, and populations. + + +Based on field investigations, the species is abundant and commonly found in the streams of Mt. Hanshan in Xingye County, Yulin City, and +Guangxi +, +China +(Fig. +11 +). During the day, the species typically seeks shelter in stone crevices and substrates, emerging at night to feed. Exposure to intense light can cause their body color to change from light yellow to dark brown or black. Their diet consists of sediment and tadpoles, including those of Quasipaa spinosa. Interestingly, many pregnant females were collected, even in December. The stream habitats also harbored other species, such as + +Schistura fasciolatus + +, + +Liniparhomaloptera disparis + +, and + +Rhinogobius similis + +. + + + + + + +Habitat of + +Oreonectes zhangi + +sp. nov. + + + + + \ No newline at end of file