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<mods:title id="016960BCDA39A6D880E0BB7D8AA0D700">Four new Dolichopoda species from Greece, one Troglophilus new to Greece and new locality records (Orthoptera, Rhaphidophoridae)</mods:title>
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<mods:namePart id="375BD265261A5F3063C012558A04D451">Alexiou, Sotiris</mods:namePart>
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<mods:affiliation id="CCC6A41606770E34142A51F3B592FFCA">Korinthian Museum of Natural History, Isthmion 201, Korinthos 20100, Greece</mods:affiliation>
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<mods:namePart id="EC8223057121524B5368916974C311C2">Bakolitsas, Kostas</mods:namePart>
<mods:nameIdentifier id="A7EED44E619C18EAF423BB7D192C9ED3" type="ORCID">0000-0003-3137-4024</mods:nameIdentifier>
<mods:affiliation id="99E0E3A6B78B9A93DAFA8F193761DBC1">Laboratory of Natural Sciences 3 rd General Lyceum, 301 31 Agrinion, Greece</mods:affiliation>
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<mods:namePart id="95357E6AB7D4680289C3E48DCEFF37AC">Di Russo, Claudio</mods:namePart>
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<mods:affiliation id="3FF99468E1453BAFD122DF43B5BFA31B">Dipartimento di Biologia e Biotecnologie “ C. Darwin ”, Università di Roma “ La Sapienza ”, Roma, Italy</mods:affiliation>
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<mods:namePart id="A7FEAE1907206EB2E5F1633F92614A2C">Rampini, Mauro</mods:namePart>
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<treatment id="00F51A50235E5E05A13964799158D495" ID-DOI="http://doi.org/10.5281/zenodo.13123941" ID-Zenodo-Dep="13123941" ID-arpha="00F51A50-235E-5E05-A139-64799158D495" LSID="urn:lsid:plazi:treatment:00F51A50235E5E05A13964799158D495" httpUri="http://treatment.plazi.org/id/00F51A50235E5E05A13964799158D495">
<subSubSection id="7DCBA55002499EFBC7497418D27BCF06" type="nomenclature">
<paragraph id="6F99ED599FA70034937892A911498F2A">
<taxonomicName id="8164E5BE35B5B3CED616F38666BDFE8D" ID-CoL="5993T" authority="Chopard, 1934" authorityName="Chopard" authorityYear="1934" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="brevicauda">
<emphasis id="A8B356A5D3528D95DDDC74D0E7F05AAA" italics="true">Troglophilus brevicauda</emphasis>
Chopard, 1934
</taxonomicName>
</paragraph>
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<subSubSection id="5348054582216D66A4815FD3999FF57B" type="description">
<paragraph id="345EDFD8267FB28FE9455D5C0A00E9BB">
<figureCitation id="61E49EFBF3B78FD6D5BDA85B0C2D25E9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Troglophylus brevicauda Chopard, 1934. Greece, Thraki, Cave Lykodromio, 17 Aug. 2020 (photo Kofinas)." figureDoi="10.3897/contrib.entomol.74.e121614.figure8" httpUri="https://binary.pensoft.net/fig/1103855">Fig. 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENWAE" type="material">
<paragraph id="A8E7EAC4EBBDF8D20C8F033C69E33BE2">
<heading id="F1C7EDBD283786B638C0B6B8E4762CC6" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="A5058FCE5AD934CCFBED17D8E1596581">
<materialsCitation id="0734A0877CA3C68F8477DD823A5B489D" collectingDate="2020-08-17" collectorName="G. Kofinas" country="Greece" county="cave Lykodromio" elevation="526" latitude="41.2377" location="Nomos Xanthis" longLatPrecision="1" longitude="24.763968" municipality="Xanthi-Stavroupoli" specimenCount="9" specimenCount-female="3" specimenCount-male="6">
<collectingCountry id="8007ADDEC2F0141DAA0E2D1E8C039284" name="Greece">GREECE</collectingCountry>
•
<specimenCount id="CF354DA16A06C690B1994B2F71FDA5D6" count="6" type="male">6 ♂</specimenCount>
,
<specimenCount id="E4035FCF3B0DE08CCFB9A9E62B3A14EA" count="3" type="female">3 ♀</specimenCount>
; Τhraki,
<location id="B9513FDBEC3FFA84BE693277CB0C47A2" LSID="urn:lsid:plazi:treatment:00F51A50235E5E05A13964799158D495:B9513FDBEC3FFA84BE693277CB0C47A2" country="Greece" county="cave Lykodromio" latitude="41.2377" longLatPrecision="1" longitude="24.763968" municipality="Xanthi-Stavroupoli" name="Nomos Xanthis">Nomos Xanthis</location>
, road
<collectingMunicipality id="0687B8AAC075CCE031F79442C2B9FAA5">Xanthi-Stavroupoli</collectingMunicipality>
,
<collectingCounty id="DA7B27CA159B5A72CF8888881FA644FD">cave Lykodromio</collectingCounty>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWWAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[24.763967,41.237700]}">
<geoCoordinate id="B3D177D7519B51644BE746B6B96B9978" degrees="41" direction="north" minutes="14.262" orientation="latitude" precision="1" value="41.2377">41 ° 14.262 ' N</geoCoordinate>
,
<geoCoordinate id="1D69F8090EE48BB2142385784E336751" degrees="24" direction="east" minutes="45.838" orientation="longitude" precision="1" value="24.763968">24 ° 45.838 ' E</geoCoordinate>
</named-content>
;
<elevation id="A2F7A3EAC23496D1B737E07B38F00F7B" metricMagnitude="2" metricUnit="m" metricValue="5.26" unit="m" value="526.0">
<quantity id="D990CBCEC37645CE1B18C9E6FA3B3221" metricMagnitude="2" metricUnit="m" metricValue="5.26" unit="m" value="526.0">526 m</quantity>
a. s. l.
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;
<date id="79CE5CF2B1038FE34C165224E6D17677" value="2020-08-17">
<collectingDate id="55F9A803CF85A8E0C5DFC7A3B8A46586" value="2020-08-17">17 Aug. 2020</collectingDate>
</date>
;
<collectorName id="4BD1D7B3574723B5A26BFD206EB70351">G. Kofinas</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0E2WAE" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
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.
</paragraph>
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<subSubSection id="SECID0EAXAE" type="comments">
<paragraph id="CCB2C36CD3C22E260079400D8184C810">
<heading id="6D6B1EDDEDE1CCB991F9C44F45A6B67B" reason="title">Comment.</heading>
</paragraph>
<paragraph id="5F36EBA225E6E36FEDBED3ECB0C247E6">
<emphasis id="6E0111AD93DEF646A85F414F446F9645" bold="true">
New for
<collectingCountry id="76341EC15C3FAD73305FE5B79A78CC3D" name="Greece">Greece</collectingCountry>
.
</emphasis>
<taxonomicName id="BFFC37E5ED90401CD0E862110109E42B" authorityName="Chopard" authorityYear="1934" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="brevicauda">
<emphasis id="F6BAB49ECA33C0907E466E4515F72550" italics="true">Troglophilus brevicauda</emphasis>
</taxonomicName>
is a species known from Serbia, Bosnia / Herzegovina, Montenegro, North
<collectingRegion id="1DF8B7ABB664BAE1E29B9BE70A9DB01B" country="Greece" name="Dytiki Makedonia">Macedonia</collectingRegion>
and Bulgaria (
<bibRefCitation id="61020E42613CAB949E49626C0AC61563" DOI="10.1111/j.1096-3642.2011.00738.x" author="Karaman" etAl="et al." firstAuthor="Karaman" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="1035-1063" refId="B10" refString="Karaman I, Hammouti N, Pavićević D, Kiefer A, Horvatović M, Seitz A (2011) The genus Troglophilus Krauss, 1879 (Orthoptera: Rhaphidophoridae) in the west Balkans. Zoological Journal of the Linnean Society 163: 1035–1063. https://doi.org/10.1111/j.1096-3642.2011.00738.x" title="The genus Troglophilus Krauss, 1879 (Orthoptera: Rhaphidophoridae) in the west Balkans." volume="163" year="2011">Karaman et al. 2011</bibRefCitation>
). In Bulgaria, the species is found in the Rhodopi Mountain range in the south. Rhodopi is shared by Greece and Bulgaria, with our location on the Greek side expanding the distribution of the species to cover all of the mountain range. According to
<bibRefCitation id="7E26D382C63DEC5A6535023C90C3F815" DOI="10.1111/j.1096-3642.2011.00738.x" author="Karaman I &amp; Hammouti N &amp; Pavićević D &amp; Kiefer A &amp; Horvatović M &amp; Seitz A" etAl="et al." firstAuthor="Karaman" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="1035-1063" refId="B10" refString="Karaman I, Hammouti N, Pavićević D, Kiefer A, Horvatović M, Seitz A (2011) The genus Troglophilus Krauss, 1879 (Orthoptera: Rhaphidophoridae) in the west Balkans. Zoological Journal of the Linnean Society 163: 1035–1063. https://doi.org/10.1111/j.1096-3642.2011.00738.x" title="The genus Troglophilus Krauss, 1879 (Orthoptera: Rhaphidophoridae) in the west Balkans." volume="163" year="2011">Karaman et al. (2011)</bibRefCitation>
, the population is an isolated local one. All the key morphological characteristics fit those of the species.
</paragraph>
<caption id="1391688F2C7AEAAFA9578ADF7D39B49E" ID-DOI="10.3897/contrib.entomol.74.e121614.figure8" ID-arpha="0B2BECFA-7A5E-5151-99AE-BC85BE82BAA8" httpUri="https://binary.pensoft.net/fig/1103855" startId="F8">
<paragraph id="05B6E4FB3AC47177B4107EF631CA2826">
<label id="264F9544FB26F8FCB1726DC6290E7EDE">Figure 8.</label>
</paragraph>
<paragraph id="1B8EA912CF91489880DA6CDA59AAF8D4">
<taxonomicName id="B5C01F6454C040B914BA2F3CC97F622A" authority="Chopard, 1934" authorityName="Chopard" authorityYear="1934" class="Insecta" family="Rhaphidophoridae" genus="Troglophylus" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="brevicauda">
<emphasis id="2B2ABC928BA3FE78913383E34FA8DF50" italics="true">Troglophylus brevicauda</emphasis>
Chopard, 1934
</taxonomicName>
. Greece, Thraki, Cave Lykodromio, 17 Aug. 2020 (photo Kofinas).
</paragraph>
</caption>
<paragraph id="E15168EEB7BA159592AF7D0F9B89CED7">
Τhe record of an immature individual from Mt Falakro, E.
<collectingCountry id="A8C96DFB3D8086A6A2D867B7A2F39618" name="Macedonia">Macedonia</collectingCountry>
by
<bibRefCitation id="338C2A320ACACBBCD8CE8CC5AA0FF6AF" author="Di Russo C &amp; Rampini M &amp; Cobolli M" etAl="et al." firstAuthor="Di Russo" journalOrPublisher="Biodiversity Journal" pagination="397-420" refId="B7" refString="Di Russo C, Rampini M, Cobolli M (2014) The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss, 1879. Biodiversity Journal 5: 397–420." title="The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss, 1879." volume="5" year="2014">Di Russo et al. (2014)</bibRefCitation>
could very well belong to the same species.
</paragraph>
</subSubSection>
</treatment>
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<mods:classification id="DB627B22CDBFBDD7C252FEB058ABAD0D">journal article</mods:classification>
<mods:identifier id="E3644F139A9D984538650EC93F60363A" type="CLB-Dataset">300338</mods:identifier>
<mods:identifier id="1950C7915E63EA21263DB7B8F14B7697" type="DOI">10.22271/fish.2024.v12.i1a.2893</mods:identifier>
<mods:identifier id="15E8C7864C3E27C79E74028A34BB0A97" type="GBIF-Dataset">5e0ea56f-c06a-4f6d-9a21-b389d8206297</mods:identifier>
<mods:identifier id="E4919239E471262B0BEDA59B71A1A453" type="ISSN">2347-5129</mods:identifier>
<mods:identifier id="6B3848991915524F0A16BB6F7C9066A1" type="Zenodo-Dep">12550280</mods:identifier>
</mods:mods>
<treatment id="0395B06BFFCFFFAD7B53C50AFB3DF84F" ID-DOI="http://doi.org/10.5281/zenodo.13123896" ID-Zenodo-Dep="13123896" LSID="urn:lsid:plazi:treatment:0395B06BFFCFFFAD7B53C50AFB3DF84F" httpUri="http://treatment.plazi.org/id/0395B06BFFCFFFAD7B53C50AFB3DF84F" lastPageId="2" lastPageNumber="65" pageId="1" pageNumber="64">
<subSubSection id="C32652F6FFCFFFAE7B53C50AFECFFB68" box="[96,288,1147,1171]" pageId="1" pageNumber="64" type="nomenclature">
<paragraph id="8B83017DFFCFFFAE7B53C50AFECFFB68" blockId="1.[96,637,963,1171]" box="[96,288,1147,1171]" pageId="1" pageNumber="64">
<heading id="D0CBB611FFCFFFAE7B53C50AFECFFB68" box="[96,288,1147,1171]" fontSize="36" level="2" pageId="1" pageNumber="64" reason="5">
<emphasis id="B948DD6FFFCFFFAE7B53C50AFF53FB68" bold="true" box="[96,188,1147,1171]" pageId="1" pageNumber="64">Species:</emphasis>
<taxonomicName id="4C3C7AFEFFCFFFAE7BF0C50DFECFFB68" ID-CoL="TM48" authority="(Bloch)" baseAuthorityName="Bloch" box="[195,288,1148,1171]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="64" phylum="Chordata" rank="species" species="punctata">
<emphasis id="B948DD6FFFCFFFAE7BF0C50DFECFFB68" box="[195,288,1148,1171]" italics="true" pageId="1" pageNumber="64">punctata</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<caption id="DF4351F5FFCFFFAE7A0FC4C9FDF1FA36" ID-DOI="http://doi.org/10.5281/zenodo.12550283" ID-Zenodo-Dep="12550283" box="[316,542,1464,1486]" httpUri="https://zenodo.org/record/12550283/files/figure.png" pageId="1" pageNumber="64" startId="1.[316,349,1464,1485]" targetBox="[108,751,1202,1442]" targetPageId="1" targetType="figure">
<paragraph id="8B83017DFFCFFFAE7A0FC4C9FDF1FA36" blockId="1.[316,542,1464,1486]" box="[316,542,1464,1486]" pageId="1" pageNumber="64">
<emphasis id="B948DD6FFFCFFFAE7A0FC4C9FE97FA36" bold="true" box="[316,376,1464,1485]" pageId="1" pageNumber="64">Fig 1:</emphasis>
<taxonomicName id="4C3C7AFEFFCFFFAE7A4DC4C9FDF1FA36" box="[382,542,1464,1486]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="64" phylum="Chordata" rank="species" species="punctata">Channa punctata</taxonomicName>
</paragraph>
</caption>
<subSubSection id="C32652F6FFCFFFAE7B53C485FC2DF903" pageId="1" pageNumber="64" type="description">
<paragraph id="8B83017DFFCFFFAE7B53C485FE86F9F7" blockId="1.[96,361,1524,1548]" box="[96,361,1524,1548]" pageId="1" pageNumber="64">
<heading id="D0CBB611FFCFFFAE7B53C485FE86F9F7" bold="true" box="[96,361,1524,1548]" fontSize="36" level="1" pageId="1" pageNumber="64" reason="1">
<emphasis id="B948DD6FFFCFFFAE7B53C485FE86F9F7" bold="true" box="[96,361,1524,1548]" pageId="1" pageNumber="64">Materials and Methods</emphasis>
</heading>
</paragraph>
<paragraph id="8B83017DFFCFFFAE7B53C763FDF6F9D1" blockId="1.[96,763,1554,1792]" box="[96,537,1554,1578]" pageId="1" pageNumber="64">
<heading id="D0CBB611FFCFFFAE7B53C763FDF6F9D1" bold="true" box="[96,537,1554,1578]" fontSize="10" level="2" pageId="1" pageNumber="64" reason="3">
<emphasis id="B948DD6FFFCFFFAE7B53C763FDF6F9D1" bold="true" box="[96,537,1554,1578]" pageId="1" pageNumber="64">Collection and maintenance of test fish</emphasis>
</heading>
</paragraph>
<paragraph id="8B83017DFFCFFFAE7B53C741FEE8F8FB" blockId="1.[96,763,1554,1792]" pageId="1" pageNumber="64">
<taxonomicName id="4C3C7AFEFFCFFFAE7B53C741FECDF9B3" box="[96,290,1584,1608]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="64" phylum="Chordata" rank="species" species="punctata">
<emphasis id="B948DD6FFFCFFFAE7B53C741FECDF9B3" box="[96,290,1584,1608]" italics="true" pageId="1" pageNumber="64">Channa punctata</emphasis>
</taxonomicName>
were collected from culture pond near Serispore T.E., Hailakandi,
<collectingRegion id="49F8CF9FFFCFFFAE7AA0C73EFE31F99C" box="[403,478,1615,1639]" country="India" name="Assam" pageId="1" pageNumber="64">Assam</collectingRegion>
,
<collectingCountry id="F32B41EDFFCFFFAE7ADFC73EFDCCF99C" box="[492,547,1615,1639]" name="India" pageId="1" pageNumber="64">India</collectingCountry>
and brought to the laboratory in aerated containers. In the laboratory, the fishes were maintained in an aquarium with constant aeration. Fishes were fed with commercially available fish meal twice daily and were acclimatized in the aquarium for a week before the experiment.
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC1F6FB30FF64" blockId="1.[824,1247,135,159]" box="[824,1247,135,159]" pageId="1" pageNumber="64">
<heading id="D0CBB611FFCFFFAE780BC1F6FB30FF64" bold="true" box="[824,1247,135,159]" fontSize="10" level="2" pageId="1" pageNumber="64" reason="3">
<emphasis id="B948DD6FFFCFFFAE780BC1F6FB30FF64" bold="true" box="[824,1247,135,159]" pageId="1" pageNumber="64">Experimental design of fish bio-assay</emphasis>
</heading>
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC1D7FA69FEE2" blockId="1.[824,1491,165,281]" pageId="1" pageNumber="64">
For test setup,
<taxonomicName id="4C3C7AFEFFCFFFAE78E4C1D4FB60FF46" box="[983,1167,165,189]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="64" phylum="Chordata" rank="species" species="punctata">
<emphasis id="B948DD6FFFCFFFAE78E4C1D4FB60FF46" box="[983,1167,165,189]" italics="true" pageId="1" pageNumber="64">Channa punctata</emphasis>
</taxonomicName>
were kept in a tank (Group-I) with heavy metal (CdCl
<subScript id="17B80338FFCFFFAE7F72C1BFFBA5FF27" attach="left" box="[1089,1098,206,220]" fontSize="6" pageId="1" pageNumber="64">2</subScript>
) for seven days. A control with no treatment was also run simultaneously in another tank. After seventh day, the protein content of liver was examined.
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC04EFBC8FEAC" blockId="1.[824,1063,319,343]" box="[824,1063,319,343]" pageId="1" pageNumber="64">
<heading id="D0CBB611FFCFFFAE780BC04EFBC8FEAC" bold="true" box="[824,1063,319,343]" fontSize="10" level="2" pageId="1" pageNumber="64" reason="3">
<emphasis id="B948DD6FFFCFFFAE780BC04EFBC8FEAC" bold="true" box="[824,1063,319,343]" pageId="1" pageNumber="64">Experimental Design</emphasis>
</heading>
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC02CFAC4FD72" blockId="1.[824,1491,349,649]" pageId="1" pageNumber="64">
The fishes were randomly divided into three groups
<emphasis id="B948DD6FFFCFFFAE7E9BC02FFA3CFE8E" box="[1448,1491,350,373]" italics="true" pageId="1" pageNumber="64">viz.,</emphasis>
control, low dose, sub-lethal-1 (SL-I, 5ppm) and high dose sub-lethal 2(SL-II, 10ppm). They were kept in three different buckets containing 5 litres of water obtained from a pond and were labelled as control, SL-1(5ppm) and SL-2 (10ppm). The experiment was run for seven days. Fishes were taken out from all the tanks after seven days and were anesthetized and the total length and body weight were recorded. Test fishes were sacrificed and tissues like gills and liver were dissected and stored at refrigerator until further analysis.
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC3DEFB69FD1D" blockId="1.[824,1376,687,742]" pageId="1" pageNumber="64">
<heading id="D0CBB611FFCFFFAE780BC3DEFB69FD1D" bold="true" fontSize="10" level="2" pageId="1" pageNumber="64" reason="3">
<emphasis id="B948DD6FFFCFFFAE780BC3DEFB69FD1D" bold="true" pageId="1" pageNumber="64">Procedure for estimation of total protein content Estimation of protein content</emphasis>
</heading>
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC39DFBDEFCD9" blockId="1.[824,1491,748,803]" pageId="1" pageNumber="64">
Total protein content was estimated by the protocol developed by Lowry
<emphasis id="B948DD6FFFCFFFAE7894C27DFC32FCD9" box="[935,989,778,803]" italics="true" pageId="1" pageNumber="64">et al.</emphasis>
(1951).
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC238FC35FC9A" blockId="1.[824,986,841,865]" box="[824,986,841,865]" pageId="1" pageNumber="64">
<heading id="D0CBB611FFCFFFAE780BC238FC35FC9A" bold="true" box="[824,986,841,865]" fontSize="10" level="2" pageId="1" pageNumber="64" reason="3">
<emphasis id="B948DD6FFFCFFFAE780BC238FC35FC9A" bold="true" box="[824,986,841,865]" pageId="1" pageNumber="64">Reagents used</emphasis>
</heading>
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC216FB67FC84" blockId="1.[824,1491,871,1232]" box="[824,1160,871,895]" pageId="1" pageNumber="64">
<heading id="D0CBB611FFCFFFAE780BC216FB67FC84" bold="true" box="[824,1160,871,895]" fontSize="10" level="2" pageId="1" pageNumber="64" reason="3">
<emphasis id="B948DD6FFFCFFFAE780BC216FB67FC84" bold="true" box="[824,1160,871,895]" pageId="1" pageNumber="64">A. Alkaline copper Reagent.</emphasis>
</heading>
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC2F7FB00FC66" blockId="1.[824,1491,871,1232]" box="[824,1263,901,926]" pageId="1" pageNumber="64">
Reagent A: 2 % Na
<subScript id="17B80338FFCFFFAE7F34C2E1FBFFFC65" attach="both" box="[1031,1040,912,926]" fontSize="6" pageId="1" pageNumber="64">2</subScript>
CO
<quantity id="4CC4AC98FFCFFFAE7F06C2E1FBB6FC66" box="[1077,1113,901,926]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="64" unit="in" value="3.0">
<subScript id="17B80338FFCFFFAE7F06C2E1FBD1FC65" attach="left" box="[1077,1086,912,926]" fontSize="6" pageId="1" pageNumber="64">3</subScript>
in
</quantity>
<geoCoordinate id="EE0867BAFFCFFFAE7F53C2F4FB74FC66" box="[1120,1179,901,925]" degrees="0.1" direction="north" orientation="latitude" pageId="1" pageNumber="64" precision="5555" value="0.1">0.1 N</geoCoordinate>
NaOH.
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC2D4FA2DFC47" blockId="1.[824,1491,871,1232]" box="[824,1474,932,957]" pageId="1" pageNumber="64">
Reagent B: 0.5% CuSO
<subScript id="17B80338FFCFFFAE7F06C2DFFBADFC47" attach="left" box="[1077,1090,932,956]" fontSize="6" pageId="1" pageNumber="64">4.</subScript>
5H
<subScript id="17B80338FFCFFFAE7F56C2DFFB82FC47" attach="both" box="[1125,1133,942,956]" fontSize="6" pageId="1" pageNumber="64">2</subScript>
O in sodium potassium tartarate
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC2B2FB7AFC02" blockId="1.[824,1491,871,1232]" pageId="1" pageNumber="64">The alkaline copper reagent was made by mixing 50 ml of reagent A and 1 ml of reagent B.</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC570FC48FBAE" blockId="1.[824,1491,871,1232]" pageId="1" pageNumber="64">B. Folin Ciocalteu phenol reagent (It is commercially available which is diluted with distilled water in the ratio of 1:2</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC52DFC06FB8F" blockId="1.[824,1491,871,1232]" box="[824,1001,1116,1140]" pageId="1" pageNumber="64">
C.
<geoCoordinate id="EE0867BAFFCFFFAE785BC52DFC73FB8F" box="[872,924,1116,1140]" degrees="0.1" direction="north" orientation="latitude" pageId="1" pageNumber="64" precision="5555" value="0.1">0.1N</geoCoordinate>
NaOH
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC50AFC01FB68" blockId="1.[824,1491,871,1232]" box="[824,1006,1147,1171]" pageId="1" pageNumber="64">D. 70% alcohol</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC5EBFB2BFB2B" blockId="1.[824,1491,871,1232]" pageId="1" pageNumber="64">
E. Protein (stock) Standard solution:
<quantity id="4CC4AC98FFCFFFAE7E31C5E8FAA3FB4A" box="[1282,1356,1177,1201]" metricMagnitude="-4" metricUnit="kg" metricValue="1.0" pageId="1" pageNumber="64" unit="mg" value="100.0">100mg</quantity>
% Bovine Serum Albumin in
<geoCoordinate id="EE0867BAFFCFFFAE7F06C5C9FB9FFB2B" box="[1077,1136,1208,1232]" degrees="0.1" direction="north" orientation="latitude" pageId="1" pageNumber="64" precision="5555" value="0.1">0.1 N</geoCoordinate>
NaOH.
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC587FBE4FAF5" blockId="1.[824,1035,1270,1294]" box="[824,1035,1270,1294]" pageId="1" pageNumber="64">
<heading id="D0CBB611FFCFFFAE780BC587FBE4FAF5" bold="true" box="[824,1035,1270,1294]" fontSize="10" level="2" pageId="1" pageNumber="64" reason="3">
<emphasis id="B948DD6FFFCFFFAE780BC587FBE4FAF5" bold="true" box="[824,1035,1270,1294]" pageId="1" pageNumber="64">Working standard</emphasis>
</heading>
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC465FA53FAD7" blockId="1.[824,1468,1300,1324]" box="[824,1468,1300,1324]" pageId="1" pageNumber="64">10 ml of the stock was diluted to 100ml with distilled water.</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC423FC25FA91" blockId="1.[824,970,1362,1386]" box="[824,970,1362,1386]" pageId="1" pageNumber="64">
<heading id="D0CBB611FFCFFFAE780BC423FC25FA91" bold="true" box="[824,970,1362,1386]" fontSize="10" level="2" pageId="1" pageNumber="64" reason="3">
<emphasis id="B948DD6FFFCFFFAE780BC423FC25FA91" bold="true" box="[824,970,1362,1386]" pageId="1" pageNumber="64">The protocol</emphasis>
</heading>
</paragraph>
<paragraph id="8B83017DFFCFFFAE780BC401FC2DF903" blockId="1.[824,1491,1392,1784]" pageId="1" pageNumber="64">
1 ml of 70% alcohol was added to the test tube after 0.2 ml of tissue homogenate was pipetted out. For ten minutes, the tubes were centrifuged at 5,000 rpm. After discarding the supernatant, 1 ml of
<geoCoordinate id="EE0867BAFFCFFFAE7F1AC4BDFBB1FA1F" box="[1065,1118,1484,1508]" degrees="0.1" direction="north" orientation="latitude" pageId="1" pageNumber="64" precision="5555" value="0.1">0.1N</geoCoordinate>
NaOH was used to dissolve the precipitate. 5 ml of Alkaline Copper Reagent was added, and it was left at room temperature for ten minutes. The Folin-Ciocalteu phenol reagent was added after ten minutes, and it was left in the dark for thirty minutes. The UV-Vis Spectrophotometer was used to measure the absorbance at
<quantity id="4CC4AC98FFCFFFAE780BC7F5FC60F967" box="[824,911,1668,1692]" metricMagnitude="-7" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="64" unit="nm" value="620.0">620 nm</quantity>
against a blank for the reagent. Simultaneously, a series of graded protein standard volumes were run. The data are given as milligrams of protein per gram of tissue weight that is moist.
</paragraph>
</subSubSection>
<caption id="DF4351F5FFCFFFAE7B4EC654FA59F8C1" ID-Table-UUID="DF4351F5FFCFFFAE7B4EC654FA59F8C1" box="[125,1462,1828,1850]" httpUri="http://table.plazi.org/id/DF4351F5FFCFFFAE7B4EC654FA59F8C1" pageId="1" pageNumber="64" startId="1.[125,184,1829,1850]" targetBox="[107,1479,1873,2098]" targetIsTable="true" targetPageId="1" targetType="table">
<paragraph id="8B83017DFFCFFFAE7B4EC654FA59F8C1" blockId="1.[125,1462,1828,1850]" box="[125,1462,1828,1850]" pageId="1" pageNumber="64">
<emphasis id="B948DD6FFFCFFFAE7B4EC654FF3DF8C1" bold="true" box="[125,210,1829,1850]" pageId="1" pageNumber="64">Table 1:</emphasis>
Morphological and behavioural changes in fishes when exposed to doses (5 ppm and 10 ppm) of CdCl
<subScript id="17B80338FFCFFFAE7F83C65AFB57F8C1" attach="left" box="[1200,1208,1835,1850]" fontSize="6" pageId="1" pageNumber="64">2</subScript>
at different exposure time
</paragraph>
</caption>
<paragraph id="8B83017DFFCFFFAE7B58C620FA9EF7C9" pageId="1" pageNumber="64">
<table id="F93CF3DDFFCF00507B58C620FA28F7C9" box="[107,1479,1873,2098]" colsContinueIn="2.[151,1393,135,464]" gridcols="8" gridrows="8" pageId="1" pageNumber="64">
<tr id="350C033FFFCF00507B58C620FA28F89F" box="[107,1479,1873,1892]" gridrow="0" isHeader="true" pageId="1" pageNumber="64">
<th id="76DD6A43FFCF00507B58C620FF19F89F" box="[107,246,1873,1892]" gridcol="0" gridrow="0" pageId="1" pageNumber="64">Exposure Time</th>
<th id="76DD6A43FFCF00507BCDC620FED0F89F" box="[254,319,1873,1892]" gridcol="1" gridrow="0" pageId="1" pageNumber="64">Dose</th>
<th id="76DD6A43FFCF00507A74C620FE34F89F" box="[327,475,1873,1892]" gridcol="2" gridrow="0" pageId="1" pageNumber="64">Colour Changes</th>
<th id="76DD6A43FFCF00507AD5C620FD61F89F" box="[486,654,1873,1892]" gridcol="3" gridrow="0" pageId="1" pageNumber="64">Opercular activity</th>
<th id="76DD6A43FFCF005079AFC620FC4DF89F" box="[668,930,1873,1892]" gridcol="4" gridrow="0" pageId="1" pageNumber="64">Bottom to surface movement</th>
<th id="76DD6A43FFCF00507882C620FBB9F89F" box="[945,1110,1873,1892]" gridcol="5" gridrow="0" pageId="1" pageNumber="64">Resting at Bottom</th>
<th id="76DD6A43FFCF00507F52C620FAECF89F" box="[1121,1283,1873,1892]" gridcol="6" gridrow="0" pageId="1" pageNumber="64">Mucous Secretion</th>
<th id="76DD6A43FFCF00507E23C620FA28F89F" box="[1296,1479,1873,1892]" gridcol="7" gridrow="0" pageId="1" pageNumber="64">
<emphasis id="B948DD6FFFCFFFAE7E23C620FA28F89F" bold="true" box="[1296,1479,1873,1892]" pageId="1" pageNumber="64">Loss of Equilibrium</emphasis>
</th>
</tr>
<tr id="350C033FFFCF00507B58C61BFA28F886" box="[107,1479,1898,1917]" gridrow="1" pageId="1" pageNumber="64">
<th id="76DD6A43FFCF00507B58C61BFA28F886" box="[107,1479,1898,1917]" colspan="8" colspanRight="7" gridcol="0" gridrow="1" pageId="1" pageNumber="64">Control</th>
</tr>
<tr id="350C033FFFCF00507B58C6F5FA28F86C" box="[107,1479,1924,1943]" gridrow="2" pageId="1" pageNumber="64">
<th id="76DD6A43FFCF00507B58C6F5FF19F84A" box="[107,246,1924,1969]" gridcol="0" gridrow="2" pageId="1" pageNumber="64" rowspan="2" rowspanBelow="1">Day 1</th>
<td id="76DD6A43FFCF00507BCDC6F5FED0F86C" box="[254,319,1924,1943]" gridcol="1" gridrow="2" pageId="1" pageNumber="64">SL-I</td>
<td id="76DD6A43FFCF00507A74C6F5FE34F86C" box="[327,475,1924,1943]" gridcol="2" gridrow="2" pageId="1" pageNumber="64">=</td>
<td id="76DD6A43FFCF00507AD5C6F5FD61F86C" box="[486,654,1924,1943]" gridcol="3" gridrow="2" pageId="1" pageNumber="64">*</td>
<td id="76DD6A43FFCF005079AFC6F5FC4DF86C" box="[668,930,1924,1943]" gridcol="4" gridrow="2" pageId="1" pageNumber="64">+</td>
<td id="76DD6A43FFCF00507882C6F5FBB9F86C" box="[945,1110,1924,1943]" gridcol="5" gridrow="2" pageId="1" pageNumber="64">=</td>
<td id="76DD6A43FFCF00507F52C6F5FAECF86C" box="[1121,1283,1924,1943]" gridcol="6" gridrow="2" pageId="1" pageNumber="64">+</td>
<td id="76DD6A43FFCF00507E23C6F5FA28F86C" box="[1296,1479,1924,1943]" gridcol="7" gridrow="2" pageId="1" pageNumber="64">+</td>
</tr>
<tr id="350C033FFFCF00507B58C6EFFA28F84A" box="[107,1479,1950,1969]" gridrow="3" pageId="1" pageNumber="64">
<td id="76DD6A43FFCF00507BCDC6EFFED0F84A" box="[254,319,1950,1969]" gridcol="1" gridrow="3" pageId="1" pageNumber="64">SL-II</td>
<td id="76DD6A43FFCF00507A74C6EFFE34F84A" box="[327,475,1950,1969]" gridcol="2" gridrow="3" pageId="1" pageNumber="64">*</td>
<td id="76DD6A43FFCF00507AD5C6EFFD61F84A" box="[486,654,1950,1969]" gridcol="3" gridrow="3" pageId="1" pageNumber="64">*</td>
<td id="76DD6A43FFCF005079AFC6EFFC4DF84A" box="[668,930,1950,1969]" gridcol="4" gridrow="3" pageId="1" pageNumber="64">+</td>
<td id="76DD6A43FFCF00507882C6EFFBB9F84A" box="[945,1110,1950,1969]" gridcol="5" gridrow="3" pageId="1" pageNumber="64">=</td>
<td id="76DD6A43FFCF00507F52C6EFFAECF84A" box="[1121,1283,1950,1969]" gridcol="6" gridrow="3" pageId="1" pageNumber="64">+</td>
<td id="76DD6A43FFCF00507E23C6EFFA28F84A" box="[1296,1479,1950,1969]" gridcol="7" gridrow="3" pageId="1" pageNumber="64">*</td>
</tr>
<tr id="350C033FFFCF00507B58C6C9FA28F830" box="[107,1479,1976,1995]" gridrow="4" pageId="1" pageNumber="64">
<th id="76DD6A43FFCF00507B58C6C9FA28F830" box="[107,1479,1976,1995]" colspan="8" colspanRight="7" gridcol="0" gridrow="4" pageId="1" pageNumber="64">Control</th>
</tr>
<tr id="350C033FFFCF00507B58C6A3FA28F81E" box="[107,1479,2002,2021]" gridrow="5" pageId="1" pageNumber="64">
<th id="76DD6A43FFCF00507B58C6A3FF19F805" box="[107,246,2002,2046]" gridcol="0" gridrow="5" pageId="1" pageNumber="64" rowspan="2" rowspanBelow="1">Day 2</th>
<td id="76DD6A43FFCF00507BCDC6A3FED0F81E" box="[254,319,2002,2021]" gridcol="1" gridrow="5" pageId="1" pageNumber="64">SL-I</td>
<td id="76DD6A43FFCF00507A74C6A3FE34F81E" box="[327,475,2002,2021]" gridcol="2" gridrow="5" pageId="1" pageNumber="64">+</td>
<td id="76DD6A43FFCF00507AD5C6A3FD61F81E" box="[486,654,2002,2021]" gridcol="3" gridrow="5" pageId="1" pageNumber="64">=</td>
<td id="76DD6A43FFCF005079AFC6A3FC4DF81E" box="[668,930,2002,2021]" gridcol="4" gridrow="5" pageId="1" pageNumber="64">=</td>
<td id="76DD6A43FFCF00507882C6A3FBB9F81E" box="[945,1110,2002,2021]" gridcol="5" gridrow="5" pageId="1" pageNumber="64">*</td>
<td id="76DD6A43FFCF00507F52C6A3FAECF81E" box="[1121,1283,2002,2021]" gridcol="6" gridrow="5" pageId="1" pageNumber="64">+</td>
<td id="76DD6A43FFCF00507E23C6A3FA28F81E" box="[1296,1479,2002,2021]" gridcol="7" gridrow="5" pageId="1" pageNumber="64">=</td>
</tr>
<tr id="350C033FFFCF00507B58C69AFA28F805" box="[107,1479,2027,2046]" gridrow="6" pageId="1" pageNumber="64">
<td id="76DD6A43FFCF00507BCDC69AFED0F805" box="[254,319,2027,2046]" gridcol="1" gridrow="6" pageId="1" pageNumber="64">SL-II</td>
<td id="76DD6A43FFCF00507A74C69AFE34F805" box="[327,475,2027,2046]" gridcol="2" gridrow="6" pageId="1" pageNumber="64">=</td>
<td id="76DD6A43FFCF00507AD5C69AFD61F805" box="[486,654,2027,2046]" gridcol="3" gridrow="6" pageId="1" pageNumber="64">=</td>
<td id="76DD6A43FFCF005079AFC69AFC4DF805" box="[668,930,2027,2046]" gridcol="4" gridrow="6" pageId="1" pageNumber="64">*</td>
<td id="76DD6A43FFCF00507882C69AFBB9F805" box="[945,1110,2027,2046]" gridcol="5" gridrow="6" pageId="1" pageNumber="64">*</td>
<td id="76DD6A43FFCF00507F52C69AFAECF805" box="[1121,1283,2027,2046]" gridcol="6" gridrow="6" pageId="1" pageNumber="64">+</td>
<td id="76DD6A43FFCF00507E23C69AFA28F805" box="[1296,1479,2027,2046]" gridcol="7" gridrow="6" pageId="1" pageNumber="64">*</td>
</tr>
<tr id="350C033FFFCF00507B58C974FA28F7C9" box="[107,1479,2053,2098]" gridrow="7" pageId="1" pageNumber="64">
<th id="76DD6A43FFCF00507B58C974FF19F7C9" box="[107,246,2053,2098]" gridcol="0" gridrow="7" pageId="1" pageNumber="64">Day 3</th>
<td id="76DD6A43FFCF00507BCDC974FED0F7C9" box="[254,319,2053,2098]" gridcol="1" gridrow="7" pageId="1" pageNumber="64">Control SL-I</td>
<td id="76DD6A43FFCF00507A74C974FE34F7C9" box="[327,475,2053,2098]" gridcol="2" gridrow="7" pageId="1" pageNumber="64">+</td>
<td id="76DD6A43FFCF00507AD5C974FD61F7C9" box="[486,654,2053,2098]" gridcol="3" gridrow="7" pageId="1" pageNumber="64">*</td>
<td id="76DD6A43FFCF005079AFC974FC4DF7C9" box="[668,930,2053,2098]" gridcol="4" gridrow="7" pageId="1" pageNumber="64">=</td>
<td id="76DD6A43FFCF00507882C974FBB9F7C9" box="[945,1110,2053,2098]" gridcol="5" gridrow="7" pageId="1" pageNumber="64">=</td>
<td id="76DD6A43FFCF00507F52C974FAECF7C9" box="[1121,1283,2053,2098]" gridcol="6" gridrow="7" pageId="1" pageNumber="64">+</td>
<td id="76DD6A43FFCF00507E23C974FA28F7C9" box="[1296,1479,2053,2098]" gridcol="7" gridrow="7" pageId="1" pageNumber="64">=</td>
</tr>
</table>
</paragraph>
<paragraph id="8B83017DFFCCFFAD7A3BC1F6FA9EFE2B" pageId="2" pageNumber="65">
<table id="F93CF3DDFFCC00507BA4C1F6FA9EFE2B" box="[151,1393,135,464]" colsContinueFrom="1.[107,1479,1873,2098]" gridcols="8" gridrows="13" pageId="2" pageNumber="65">
<tr id="350C033FFFCC00507BA4C1F6FA9EFF61" box="[151,1393,135,154]" gridrow="0" pageId="2" pageNumber="65" rowspan-0="1">
<th id="76DD6A43FFCC00507BCDC1F6FED0FF61" box="[254,319,135,154]" gridcol="1" gridrow="0" pageId="2" pageNumber="65">SL-II</th>
<th id="76DD6A43FFCC00507AB8C1F6FE78FF61" box="[395,407,135,154]" gridcol="2" gridrow="0" pageId="2" pageNumber="65">=</th>
<th id="76DD6A43FFCC00507900C1F6FDD0FF61" box="[563,575,135,154]" gridcol="3" gridrow="0" pageId="2" pageNumber="65">*</th>
<th id="76DD6A43FFCC0050782AC1F6FCCAFF61" box="[793,805,135,154]" gridcol="4" gridrow="0" pageId="2" pageNumber="65">*</th>
<th id="76DD6A43FFCC005078CEC1F6FBE6FF61" box="[1021,1033,135,154]" gridcol="5" gridrow="0" pageId="2" pageNumber="65">=</th>
<th id="76DD6A43FFCC00507F9FC1F6FB57FF61" box="[1196,1208,135,154]" gridcol="6" gridrow="0" pageId="2" pageNumber="65">+</th>
<th id="76DD6A43FFCC00507E56C1F6FA9EFF61" box="[1381,1393,135,154]" gridcol="7" gridrow="0" pageId="2" pageNumber="65">*</th>
</tr>
<tr id="350C033FFFCC00507BA4C1D1FA9EFF48" box="[151,1393,160,179]" gridrow="1" pageId="2" pageNumber="65" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DD6A43FFCC00507BA4C1D1FF24FF1C" box="[151,203,160,231]" gridcol="0" gridrow="1" pageId="2" pageNumber="65" rowspan="3" rowspanBelow="2">Day 4</th>
<td id="76DD6A43FFCC00507BCDC1D1FED0FF48" box="[254,319,160,179]" gridcol="1" gridrow="1" pageId="2" pageNumber="65">Control</td>
</tr>
<tr id="350C033FFFCC00507BA4C1CBFA9EFF36" box="[151,1393,186,205]" gridrow="2" pageId="2" pageNumber="65">
<td id="76DD6A43FFCC00507BCDC1CBFED0FF36" box="[254,319,186,205]" gridcol="1" gridrow="2" pageId="2" pageNumber="65">SL-I</td>
<td id="76DD6A43FFCC00507AB8C1CBFE78FF36" box="[395,407,186,205]" gridcol="2" gridrow="2" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507900C1CBFDD0FF36" box="[563,575,186,205]" gridcol="3" gridrow="2" pageId="2" pageNumber="65">*</td>
<td id="76DD6A43FFCC0050782AC1CBFCCAFF36" box="[793,805,186,205]" gridcol="4" gridrow="2" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC005078CEC1CBFBE6FF36" box="[1021,1033,186,205]" gridcol="5" gridrow="2" pageId="2" pageNumber="65">*</td>
<td id="76DD6A43FFCC00507F9FC1CBFB57FF36" box="[1196,1208,186,205]" gridcol="6" gridrow="2" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507E56C1CBFA9EFF36" box="[1381,1393,186,205]" gridcol="7" gridrow="2" pageId="2" pageNumber="65">*</td>
</tr>
<tr id="350C033FFFCC00507BA4C1A5FA9EFF1C" box="[151,1393,212,231]" gridrow="3" pageId="2" pageNumber="65">
<td id="76DD6A43FFCC00507BCDC1A5FED0FF1C" box="[254,319,212,231]" gridcol="1" gridrow="3" pageId="2" pageNumber="65">SL-II</td>
<td id="76DD6A43FFCC00507AB8C1A5FE78FF1C" box="[395,407,212,231]" gridcol="2" gridrow="3" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC00507900C1A5FDD0FF1C" box="[563,575,212,231]" gridcol="3" gridrow="3" pageId="2" pageNumber="65">*</td>
<td id="76DD6A43FFCC0050782AC1A5FCCAFF1C" box="[793,805,212,231]" gridcol="4" gridrow="3" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC005078CEC1A5FBE6FF1C" box="[1021,1033,212,231]" gridcol="5" gridrow="3" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC00507F9FC1A5FB57FF1C" box="[1196,1208,212,231]" gridcol="6" gridrow="3" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507E56C1A5FA9EFF1C" box="[1381,1393,212,231]" gridcol="7" gridrow="3" pageId="2" pageNumber="65">*</td>
</tr>
<tr id="350C033FFFCC00507BA4C19FFA9EFEFA" box="[151,1393,238,257]" gridrow="4" pageId="2" pageNumber="65" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DD6A43FFCC00507BA4C19FFF24FECE" box="[151,203,238,309]" gridcol="0" gridrow="4" pageId="2" pageNumber="65" rowspan="3" rowspanBelow="2">Day 5</th>
<td id="76DD6A43FFCC00507BCDC19FFED0FEFA" box="[254,319,238,257]" gridcol="1" gridrow="4" pageId="2" pageNumber="65">Control</td>
</tr>
<tr id="350C033FFFCC00507BA4C079FA9EFEE0" box="[151,1393,264,283]" gridrow="5" pageId="2" pageNumber="65">
<td id="76DD6A43FFCC00507BCDC079FED0FEE0" box="[254,319,264,283]" gridcol="1" gridrow="5" pageId="2" pageNumber="65">SL-I</td>
<td id="76DD6A43FFCC00507AB8C079FE78FEE0" box="[395,407,264,283]" gridcol="2" gridrow="5" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507900C079FDD0FEE0" box="[563,575,264,283]" gridcol="3" gridrow="5" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC0050782AC079FCCAFEE0" box="[793,805,264,283]" gridcol="4" gridrow="5" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC005078CEC079FBE6FEE0" box="[1021,1033,264,283]" gridcol="5" gridrow="5" pageId="2" pageNumber="65">*</td>
<td id="76DD6A43FFCC00507F9FC079FB57FEE0" box="[1196,1208,264,283]" gridcol="6" gridrow="5" pageId="2" pageNumber="65">*</td>
<td id="76DD6A43FFCC00507E56C079FA9EFEE0" box="[1381,1393,264,283]" gridcol="7" gridrow="5" pageId="2" pageNumber="65">=</td>
</tr>
<tr id="350C033FFFCC00507BA4C053FA9EFECE" box="[151,1393,290,309]" gridrow="6" pageId="2" pageNumber="65">
<td id="76DD6A43FFCC00507BCDC053FED0FECE" box="[254,319,290,309]" gridcol="1" gridrow="6" pageId="2" pageNumber="65">SL-II</td>
<td id="76DD6A43FFCC00507AB8C053FE78FECE" box="[395,407,290,309]" gridcol="2" gridrow="6" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC00507900C053FDD0FECE" box="[563,575,290,309]" gridcol="3" gridrow="6" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC0050782AC053FCCAFECE" box="[793,805,290,309]" gridcol="4" gridrow="6" pageId="2" pageNumber="65">*</td>
<td id="76DD6A43FFCC005078CEC053FBE6FECE" box="[1021,1033,290,309]" gridcol="5" gridrow="6" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507F9FC053FB57FECE" box="[1196,1208,290,309]" gridcol="6" gridrow="6" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC00507E56C053FA9EFECE" box="[1381,1393,290,309]" gridcol="7" gridrow="6" pageId="2" pageNumber="65">*</td>
</tr>
<tr id="350C033FFFCC00507BA4C04DFA9EFEB4" box="[151,1393,316,335]" gridrow="7" pageId="2" pageNumber="65" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DD6A43FFCC00507BA4C04DFF24FE79" box="[151,203,316,386]" gridcol="0" gridrow="7" pageId="2" pageNumber="65" rowspan="3" rowspanBelow="2">Day 6</th>
<td id="76DD6A43FFCC00507BCDC04DFED0FEB4" box="[254,319,316,335]" gridcol="1" gridrow="7" pageId="2" pageNumber="65">Control</td>
</tr>
<tr id="350C033FFFCC00507BA4C024FA9EFE93" box="[151,1393,341,360]" gridrow="8" pageId="2" pageNumber="65">
<td id="76DD6A43FFCC00507BCDC024FED0FE93" box="[254,319,341,360]" gridcol="1" gridrow="8" pageId="2" pageNumber="65">SL-I</td>
<td id="76DD6A43FFCC00507AB8C024FE78FE93" box="[395,407,341,360]" gridcol="2" gridrow="8" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507900C024FDD0FE93" box="[563,575,341,360]" gridcol="3" gridrow="8" pageId="2" pageNumber="65">*</td>
<td id="76DD6A43FFCC0050782AC024FCCAFE93" box="[793,805,341,360]" gridcol="4" gridrow="8" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC005078CEC024FBE6FE93" box="[1021,1033,341,360]" gridcol="5" gridrow="8" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507F9FC024FB57FE93" box="[1196,1208,341,360]" gridcol="6" gridrow="8" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC00507E56C024FA9EFE93" box="[1381,1393,341,360]" gridcol="7" gridrow="8" pageId="2" pageNumber="65">*</td>
</tr>
<tr id="350C033FFFCC00507BA4C01EFA9EFE79" box="[151,1393,367,386]" gridrow="9" pageId="2" pageNumber="65">
<td id="76DD6A43FFCC00507BCDC01EFED0FE79" box="[254,319,367,386]" gridcol="1" gridrow="9" pageId="2" pageNumber="65">SL-II</td>
<td id="76DD6A43FFCC00507AB8C01EFE78FE79" box="[395,407,367,386]" gridcol="2" gridrow="9" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507900C01EFDD0FE79" box="[563,575,367,386]" gridcol="3" gridrow="9" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC0050782AC01EFCCAFE79" box="[793,805,367,386]" gridcol="4" gridrow="9" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC005078CEC01EFBE6FE79" box="[1021,1033,367,386]" gridcol="5" gridrow="9" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC00507F9FC01EFB57FE79" box="[1196,1208,367,386]" gridcol="6" gridrow="9" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC00507E56C01EFA9EFE79" box="[1381,1393,367,386]" gridcol="7" gridrow="9" pageId="2" pageNumber="65">+</td>
</tr>
<tr id="350C033FFFCC00507BA4C0F8FA9EFE67" box="[151,1393,393,412]" gridrow="10" pageId="2" pageNumber="65" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76DD6A43FFCC00507BA4C0F8FF24FE2B" box="[151,203,393,464]" gridcol="0" gridrow="10" pageId="2" pageNumber="65" rowspan="3" rowspanBelow="2">Day 7</th>
<td id="76DD6A43FFCC00507BCDC0F8FED0FE67" box="[254,319,393,412]" gridcol="1" gridrow="10" pageId="2" pageNumber="65">Control</td>
</tr>
<tr id="350C033FFFCC00507BA4C0D2FA9EFE4D" box="[151,1393,419,438]" gridrow="11" pageId="2" pageNumber="65">
<td id="76DD6A43FFCC00507BCDC0D2FED0FE4D" box="[254,319,419,438]" gridcol="1" gridrow="11" pageId="2" pageNumber="65">SL-I</td>
<td id="76DD6A43FFCC00507AB8C0D2FE78FE4D" box="[395,407,419,438]" gridcol="2" gridrow="11" pageId="2" pageNumber="65">*</td>
<td id="76DD6A43FFCC00507900C0D2FDD0FE4D" box="[563,575,419,438]" gridcol="3" gridrow="11" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC0050782AC0D2FCCAFE4D" box="[793,805,419,438]" gridcol="4" gridrow="11" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC005078CEC0D2FBE6FE4D" box="[1021,1033,419,438]" gridcol="5" gridrow="11" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507F9FC0D2FB57FE4D" box="[1196,1208,419,438]" gridcol="6" gridrow="11" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC00507E56C0D2FA9EFE4D" box="[1381,1393,419,438]" gridcol="7" gridrow="11" pageId="2" pageNumber="65">=</td>
</tr>
<tr id="350C033FFFCC00507BA4C0CCFA9EFE2B" box="[151,1393,445,464]" gridrow="12" pageId="2" pageNumber="65">
<td id="76DD6A43FFCC00507BCDC0CCFED0FE2B" box="[254,319,445,464]" gridcol="1" gridrow="12" pageId="2" pageNumber="65">SL-II</td>
<td id="76DD6A43FFCC00507AB8C0CCFE78FE2B" box="[395,407,445,464]" gridcol="2" gridrow="12" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507900C0CCFDD0FE2B" box="[563,575,445,464]" gridcol="3" gridrow="12" pageId="2" pageNumber="65">=</td>
<td id="76DD6A43FFCC0050782AC0CCFCCAFE2B" box="[793,805,445,464]" gridcol="4" gridrow="12" pageId="2" pageNumber="65">*</td>
<td id="76DD6A43FFCC005078CEC0CCFBE6FE2B" box="[1021,1033,445,464]" gridcol="5" gridrow="12" pageId="2" pageNumber="65">+</td>
<td id="76DD6A43FFCC00507F9FC0CCFB57FE2B" box="[1196,1208,445,464]" gridcol="6" gridrow="12" pageId="2" pageNumber="65">*</td>
<td id="76DD6A43FFCC00507E56C0CCFA9EFE2B" box="[1381,1393,445,464]" gridcol="7" gridrow="12" pageId="2" pageNumber="65">+</td>
</tr>
</table>
</paragraph>
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<paragraph id="8B83017DFFCCFFAD7B53C0A6FE6AFE16" blockId="2.[96,654,471,576]" box="[96,389,471,493]" pageId="2" pageNumber="65">SL-I = 5 ppm; SL-II = 10 ppm</paragraph>
<paragraph id="8B83017DFFCCFFAD7B53C082FDC4FDF2" blockId="2.[96,654,471,576]" box="[96,555,499,521]" pageId="2" pageNumber="65">“=” indicates no change or no increase/decrease</paragraph>
<paragraph id="8B83017DFFCCFFAD7B53C37FFD63FDDF" blockId="2.[96,654,471,576]" box="[96,652,526,548]" pageId="2" pageNumber="65">“+” indicates occurrence of changes or increase in change</paragraph>
<paragraph id="8B83017DFFCCFFAD7B53C35BFD61FDBB" blockId="2.[96,654,471,576]" box="[96,654,554,576]" pageId="2" pageNumber="65">“*” indicates occurrence of changes or decrease in change</paragraph>
</tableNote>
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<emphasis id="B948DD6FFFCCFFAD7B99C314FF11FD81" bold="true" box="[170,254,613,634]" pageId="2" pageNumber="65">Table 2:</emphasis>
Protein Content of the tissues (gill and liver) of
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at different concentrations of CdCl
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after 7 days of exposure
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<emphasis id="B948DD6FFFCCFFAD7BE5C3E3FEA2FD5C" bold="true" box="[214,333,658,679]" pageId="2" pageNumber="65">Treatments</emphasis>
</th>
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<emphasis id="B948DD6FFFCCFFAD790EC3E3FD75FD5C" bold="true" box="[573,666,658,679]" pageId="2" pageNumber="65">Duration</emphasis>
</th>
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<emphasis id="B948DD6FFFCCFFAD7887C3E3FC34FD5C" bold="true" box="[948,987,658,679]" pageId="2" pageNumber="65">Gill</emphasis>
</th>
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<emphasis id="B948DD6FFFCCFFAD7E35C3E3FAD1FD5C" bold="true" box="[1286,1342,658,679]" pageId="2" pageNumber="65">Liver</emphasis>
</th>
</tr>
<tr id="350C033FFFCC00507B9BC3DFFABDFD3F" box="[168,1362,686,708]" gridrow="1" pageId="2" pageNumber="65">
<th id="76DD6A43FFCC00507B9BC3DFFE95FD3F" box="[168,378,686,708]" gridcol="0" gridrow="1" pageId="2" pageNumber="65">5 ppm CdCl2 (mg/L)</th>
<td id="76DD6A43FFCC0050790EC3DFFD75FD3F" box="[573,666,686,708]" gridcol="1" gridrow="1" pageId="2" pageNumber="65">7 Days</td>
<td id="76DD6A43FFCC005078A5C3DFFC17FD3F" box="[918,1016,686,708]" gridcol="2" gridrow="1" pageId="2" pageNumber="65">70.3±0.25</td>
<td id="76DD6A43FFCC00507FC2C3DFFABDFD3F" box="[1265,1362,686,708]" gridcol="3" gridrow="1" pageId="2" pageNumber="65">79.3±0.20</td>
</tr>
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<th id="76DD6A43FFCC00507B9BC3BBFE95FD1B" box="[168,378,714,736]" gridcol="0" gridrow="2" pageId="2" pageNumber="65">10 ppm CdCl2 (mg/L)</th>
<td id="76DD6A43FFCC0050790EC3BBFD75FD1B" box="[573,666,714,736]" gridcol="1" gridrow="2" pageId="2" pageNumber="65">7 Days</td>
<td id="76DD6A43FFCC005078A5C3BBFC17FD1B" box="[918,1016,714,736]" gridcol="2" gridrow="2" pageId="2" pageNumber="65">65.3±0.30</td>
<td id="76DD6A43FFCC00507FC2C3BBFABDFD1B" box="[1265,1362,714,736]" gridcol="3" gridrow="2" pageId="2" pageNumber="65">71.3±0.25</td>
</tr>
<tr id="350C033FFFCC00507B9BC396FABDFD06" box="[168,1362,743,765]" gridrow="3" pageId="2" pageNumber="65">
<th id="76DD6A43FFCC00507B9BC396FE95FD06" box="[168,378,743,765]" gridcol="0" gridrow="3" pageId="2" pageNumber="65">Control</th>
<td id="76DD6A43FFCC0050790EC396FD75FD06" box="[573,666,743,765]" gridcol="1" gridrow="3" pageId="2" pageNumber="65">7 Days</td>
<td id="76DD6A43FFCC005078A5C396FC17FD06" box="[918,1016,743,765]" gridcol="2" gridrow="3" pageId="2" pageNumber="65">82.5±0.20</td>
<td id="76DD6A43FFCC00507FC2C396FABDFD06" box="[1265,1362,743,765]" gridcol="3" gridrow="3" pageId="2" pageNumber="65">89.7±0.15</td>
</tr>
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</paragraph>
<caption id="DF4351F5FFCCFFAD7BFCC75EFA8BF9BF" ID-DOI="http://doi.org/10.5281/zenodo.12550285" ID-Zenodo-Dep="12550285" box="[207,1380,1583,1605]" httpUri="https://zenodo.org/record/12550285/files/figure.png" pageId="2" pageNumber="65" startId="2.[207,240,1583,1604]" targetBox="[271,1336,816,1551]" targetPageId="2" targetType="figure">
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<emphasis id="B948DD6FFFCCFFAD7BFCC75EFEE5F9BF" bold="true" box="[207,266,1583,1604]" pageId="2" pageNumber="65">Fig 1:</emphasis>
Protein Content of the tissues (gill and liver) of
<taxonomicName id="4C3C7AFEFFCCFFAD79EFC75EFCA3F9BE" box="[732,844,1583,1605]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="65" phylum="Chordata" rank="species" species="punctata">
<emphasis id="B948DD6FFFCCFFAD79EFC75EFCA3F9BE" box="[732,844,1583,1605]" italics="true" pageId="2" pageNumber="65">C. punctata</emphasis>
</taxonomicName>
at 5 ppm and 10 ppm of CdCl
<subScript id="17B80338FFCCFFAD7F43C747FB97F9BE" attach="left" box="[1136,1144,1590,1605]" fontSize="6" pageId="2" pageNumber="65">2</subScript>
after 7 days of exposure
</paragraph>
</caption>
<subSubSection id="C32652F6FFCCFFAD7B53C71AFB3DF84F" pageId="2" pageNumber="65" type="discussion">
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<emphasis id="B948DD6FFFCCFFAD7B53C71AFE8CF978" bold="true" box="[96,355,1643,1667]" pageId="2" pageNumber="65">Results and Discussion</emphasis>
</heading>
</paragraph>
<paragraph id="8B83017DFFCCFFAD7B53C7F8FC5BF944" blockId="2.[96,763,1673,2095]" lastBlockId="2.[824,1491,1642,1972]" pageId="2" pageNumber="65">
The present study indicated that the heavy metal Cadmium Chloride (CdCl
<subScript id="17B80338FFCCFFAD7A3BC7C0FEFEF944" attach="left" box="[264,273,1713,1727]" fontSize="6" pageId="2" pageNumber="65">2</subScript>
) has a dreadful effect on the freshwater fish
<taxonomicName id="4C3C7AFEFFCCFFAD7B53C7B7FE98F925" authority="(Bloch)" baseAuthorityName="Bloch" box="[96,375,1734,1758]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="65" phylum="Chordata" rank="species" species="punctata">
<emphasis id="B948DD6FFFCCFFAD7B53C7B7FEF4F925" box="[96,283,1734,1758]" italics="true" pageId="2" pageNumber="65">Channa punctata</emphasis>
(Bloch)
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(Qureshi et al, 2011)
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[
<bibRefCitation id="EFAD7C8CFFCCFFAD7944C7B5FD6EF929" author="Ahmad B &amp; Qureshi TA &amp; Manohar S &amp; Kaur P &amp; Khaliq R." box="[631,641,1732,1746]" pageId="2" pageNumber="65" pagination="531 - 536" refId="ref2247" refString="3. Ahmad B, Qureshi TA, Manohar S, Kaur P, Khaliq R. Effect of cadmium chloride on the histoarchitecture of liver and kidney of a freshwater Catfish, Clariasbatrachus. International Journal of Environmental Sciences. 2011; 2 (2): 531 - 536." type="journal article" year="2011">3</bibRefCitation>
]
</superScript>
. From the behavioural responses of fish exposed to sub-lethal concentration of the heavy metal showed that they were under stress. The increase in opercular movement and the bottom to upward movement of the fishes to overcome hypoxic condition was seen in the 5
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and 7
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day of exposure. Mucus secretion, starting of depletion of scales, skin lesions on dorsal and lateral side of the body, copious mucus, clumping of gills increased with the increasing concentration of the toxicant (Rastogi,
<emphasis id="B948DD6FFFCCFFAD7A15C6AAFEBAF809" box="[294,341,2010,2034]" italics="true" pageId="2" pageNumber="65">et al</emphasis>
., 2018)
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[
<bibRefCitation id="EFAD7C8CFFCCFFAD7A87C6A9FE51F81D" author="Rastogi S &amp; Yadav P &amp; Singh M." box="[436,446,2008,2022]" pageId="2" pageNumber="65" pagination="66 - 68" refId="ref2344" refString="5. Rastogi S, Yadav P, Singh M. Toxicity of cadmium and its effect on the morphology of the fresh water Teleost, Heteropneustes fossilis (Bloch). International Journal on Biological Sciences. 2018; 9 (1): 66 - 68." type="journal article" year="2018">5</bibRefCitation>
]
</superScript>
. Colour changes of the body from blackish to pale yellowish and restlessness again confirmed the anxiety level of the fishes in the experiment (
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). Also many visible lesions were observed near operculum, mouth, pectoral and pelvic fins and also near the caudal fins.
</paragraph>
<paragraph id="8B83017DFFCCFFAD780BC7B7FB3DF84F" blockId="2.[824,1491,1642,1972]" pageId="2" pageNumber="65">
Also from the total protein content assay, it was observed that the gills and liver tissues when exposed to sub lethal heavy metal concentration of CdCl
<subScript id="17B80338FFCCFFAD7F4CC67CFB67F8E0" attach="left" box="[1151,1160,1805,1819]" fontSize="6" pageId="2" pageNumber="65">2</subScript>
(5ppm and 10ppm) showed gradual decline in the protein content with the increase in exposure time, which further confirms the toxicity of the heavy metal (
<tableCitation id="C6BE34C6FFCCFFAD78E6C62EFBDDF88C" box="[981,1074,1887,1911]" captionStart="Table 2" captionStartId="2.[170,229,613,634]" captionTargetBox="[168,1362,658,765]" captionTargetId="graphics-818@2.[99,1488,650,767]" captionTargetPageId="2" captionText="Table 2: Protein Content of the tissues (gill and liver) of C. punctata at different concentrations of CdCl2 after 7 days of exposure" httpUri="http://table.plazi.org/id/DF4351F5FFCCFFAD7B99C314FA66FD81" pageId="2" pageNumber="65" tableUuid="DF4351F5FFCCFFAD7B99C314FA66FD81">Table 2</tableCitation>
). This result indicates that the vital organs like gills and liver if affected proves lethal in survival of the fishes (Agrahari
<emphasis id="B948DD6FFFCCFFAD7F1CC6ECFBB1F84F" box="[1071,1118,1948,1972]" italics="true" pageId="2" pageNumber="65">et al</emphasis>
., 2007)
<superScript id="7C49AC35FFCCFFAD7F84C6EAFB24F852" attach="right" box="[1207,1227,1947,1961]" fontSize="6" pageId="2" pageNumber="65">
[
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]
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.
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</treatment>
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<mods:identifier id="6ABC96BD5CD8AB7F853E8F8F325BED30" type="ZooBank">154CE236-EFA8-4411-834B-234A9B45A63F</mods:identifier>
</mods:mods>
<treatment id="0987463F2E2D509F88594C782A30D269" ID-DOI="http://doi.org/10.5281/zenodo.13124023" ID-Zenodo-Dep="13124023" ID-arpha="0987463F-2E2D-509F-8859-4C782A30D269" ID-zoobank="https://zoobank.org/6196FFDC-ED56-484E-A845-0C0A7D542BF9" LSID="urn:lsid:zoobank.org:act:6196FFDC-ED56-484E-A845-0C0A7D542BF9" LSID-ZBK="urn:lsid:zoobank.org:act:6196FFDC-ED56-484E-A845-0C0A7D542BF9" httpUri="http://treatment.plazi.org/id/0987463F2E2D509F88594C782A30D269">
<subSubSection id="C88DABE113EA3CC2AD41B83EF80D7B31" type="nomenclature">
<paragraph id="4C9B295292CD4C6BA9E2ADD878D79724">
<taxonomicName id="B66B065DF31F1C5AD1F836200E528CA0" authority="Boruah, Narayanan, Deepak &amp; Das" authorityName="Boruah, Narayanan, Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="barailensis" status="sp. nov.">
<emphasis id="936847D169F3067E74E68A4D60A774A8" italics="true">Cyrtodactylus barailensis</emphasis>
Boruah, Narayanan, Deepak &amp; Das
</taxonomicName>
<taxonomicNameLabel id="7AA93D7D71C9F7B8E5778A5EFD21E1E3" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="29F61530D277961A3A190D661BEFB00D" type="description">
<paragraph id="88F2C05BBD53DDD121879BB3C62119CE">
<figureCitation id="432F9A987D9C301E25AD05B0051EF3FE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Cyrtodactylus barailensis sp. nov. (WII-ADR 971). A dorsal and B ventral view of the whole body; C dorsal, D ventral, and E lateral view of head; F dorsal tubercles on trunk; G ventral view of left manus, and H left pes. I C. barailensis sp. nov. in life. Scale bar: 10 mm." figureDoi="10.3897/vz.74.e124752.figure5" httpUri="https://binary.pensoft.net/fig/1102271">Figure 5</figureCitation>
;
<tableCitation id="D5D90750E8A8736F85586E003DB7354F" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">Tables 2, S 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZSAK" type="Holotype">
<paragraph id="F4A7584E3530587ECDBEB5BECE7B13A8">
<heading id="35D500D43A570723DBC2257228B0832E" reason="title">
<typeStatus id="9BE515FC895673A5805C4E93B88B0926" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="B61575AA4DECB8C95540F059045F4859">
<materialsCitation id="29B43AB5640DF5D78509BD6FE93C504F" collectingDate="2021-08-14" collectorName="Abhijit Das" elevation="740" latitude="25.54199" location="Peren District" longLatPrecision="5" longitude="93.6307" specimenCount="1" stateProvince="Nagaland" typeStatus="holotype">
Adult female (WII-ADR 971), from Athibung (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECTAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[93.630700,25.541990]}">
<geoCoordinate id="C48E58CB783A9E1E667ECE6E75C9B6AA" degrees="25.54199" direction="north" orientation="latitude" precision="1" value="25.54199">25.54199 ° N</geoCoordinate>
;
<geoCoordinate id="6406EAD8A7EB729D56D027145513E607" degrees="93.6307" direction="east" orientation="longitude" precision="5" value="93.6307">93.6307 ° E</geoCoordinate>
</named-content>
;
<elevation id="9E3695E80E00DEAF39214F6766017FE2" metricMagnitude="2" metricUnit="m" metricValue="7.4" unit="m" value="740.0">
elevation
<quantity id="5109F9736321BBF4445998FEDC08D315" metricMagnitude="2" metricUnit="m" metricValue="7.4" unit="m" value="740.0">740 m</quantity>
a. s. l.
</elevation>
) (Fig.
<figureCitation id="E533ABC182595ED938F3C30C6D134557" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 A</figureCitation>
),
<location id="FF6612D812061757FC57E42A61FB3A60" LSID="urn:lsid:plazi:treatment:0987463F2E2D509F88594C782A30D269:FF6612D812061757FC57E42A61FB3A60" latitude="25.54199" longLatPrecision="5" longitude="93.6307" name="Peren District" stateProvince="Nagaland">Peren District</location>
,
<collectingRegion id="9E413FE8D03EB5A5C3AAB2BF5D1D4092" country="India" name="Nagaland">Nagaland</collectingRegion>
, collected by
<collectorName id="AAC1735A433D724883938601D5790EAC">Abhijit Das</collectorName>
and Bitupan Boruah on
<date id="227153EA81FE12A333F3BD32F70642D7" value="2021-08-14">
<collectingDate id="3EC9E7BCF7086D4E4C945D513629488F" value="2021-08-14">14 August 2021</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELTAK" type="diagnosis">
<paragraph id="2810FE24D332ABC3C92E314F09ABD6EE">
<heading id="E1F47E85CFB09BBFA39F24BE34E989D9" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="2487AE75720DF93160718256D2B9F065">
Medium-sized gecko,
<abbrev id="ABBRID0ERTAK" xlink_title="snout to vent length">SVL</abbrev>
at least
<quantity id="DA1B3E0F319595E5F9D8DAA872F5D038" metricMagnitude="-2" metricUnit="m" metricValue="6.88" unit="mm" value="68.8">68.8 mm</quantity>
in adult female; supralabials 9–12 and infralabials nine; 12 or 13 lamellae beneath the digit IV of manus; 17 lamellae beneath digit IV of pes; 17 feebly keeled and bluntly conical tubercles across mid dorsum and 32 paravertebral tubercles; 36 smooth mid-ventral scales between ventrolateral folds; at least 10 small precloacal pores in female; head on top pale-brown with purplish tinge; dark-brown postorbital stripe continuing to above the ear opening; a dark-brown cross bar with irregular edges on nape; indistinct pale-yellow patch on occipital region; dorsally and laterally neck and back pale yellowish-brown with dark-brown irregular reticulation.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVTAK" type="description">
<paragraph id="A4DE332E4532EC63C76699E5D878821B">
<heading id="234491D36D0351A88814438D6E40E50D" reason="title">
Description of
<typeStatus id="1D25E81C4191BF7B66FCF9C80884D4DB" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="D0907AC1EE32905AA0AE1C20F67927FF">
<typeStatus id="1D5D72EBE15CBD5BC748324204C0C92F" type="holotype">Holotype</typeStatus>
well preserved except an incision below left axilla ventrolaterally. Snout-vent length
<quantity id="011E1D7F4FD86F11717030FF601778F1" metricMagnitude="-2" metricUnit="m" metricValue="6.88" unit="mm" value="68.8">68.8 mm</quantity>
. Head moderately large (
<abbrev id="ABBRID0E2TAK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
/
<abbrev id="ABBRID0E6TAK" xlink_title="snout to vent length">SVL</abbrev>
= 0.26), dorsoventrally depressed, longer than width (
<abbrev id="ABBRID0EDUAK" xlink_title="maximum head width">HW</abbrev>
/
<abbrev id="ABBRID0EHUAK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.65), oval, distinct from neck, broader at occipital region; snout tip rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat; a longitudinal furrow on dorsal surface of the snout; snout short (
<abbrev id="ABBRID0ELUAK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
/
<abbrev id="ABBRID0EPUAK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.39), longer than orbit (
<abbrev id="ABBRID0ETUAK" xlink_title="orbit diameter, taken between the anterior and posterior bony orbital borders">OD</abbrev>
/
<abbrev id="ABBRID0EXUAK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
= 0.59); nostril rounded, opening directed posterolaterally; ear opening oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous, supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at the anterodorsal region; rostral wide, a short groove at the middle on top; rostral connected with nasals, supranasals and first supralabials; a single scale between the supranasals, larger than the rest of the granular snout scales; granular scales at parietal, occipital and temporal region intermixed with slightly larger rounded and bluntly conical granular tubercles starting from the level of posterior margin of the upper eyelids, smaller in parietal region, size increases towards nape; supralabial 12 on right side and nine on left side, supralabials count up to midorbit is eight on right side and six on left side, size decreases towards angle of jaw; a series of narrow, enlarged scales above the supralabials between nostril and anterior orbital border; mental triangular, connected with first infralabials and inner postmentals; nine infralabials on both side, size decreases towards angle of jaw; inner pair of postmentals are larger than the outer postmentals; posterior margin of the inner postmentals bordered by eight granular scales of different size; two rows of slightly enlarged, narrow scales, larger than gular scales present along the infralabials starting below the outer postmentals, posteriorly size decreases; gular scales granular, juxtaposed, homogeneous, size increases towards the throat where they become imbricate.
</paragraph>
<paragraph id="00085B6F36BB370F40D2BB495C87E464">
Habitus slender (
<abbrev id="ABBRID0E4UAK" xlink_title="maximum body width">BW</abbrev>
/
<abbrev id="ABBRID0EBVAK" xlink_title="snout to vent length">SVL</abbrev>
= 0.16,
<abbrev id="ABBRID0EFVAK" xlink_title="trunk length, between the axilla and groin">TRL</abbrev>
/
<abbrev id="ABBRID0EJVAK" xlink_title="snout to vent length">SVL</abbrev>
= 0.47), dorsoventrally depressed; dorsal scales granular, rounded, heterogeneous, intermixed with densely placed rounded, weakly keeled and bluntly conical tubercles, irregularly arranged, continues to seventh segment of the tail, size increases towards posterior body and pronounced at the base of tail; 17 dorsal tubercles across mid dorsum; 32 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid, subimbricate to imbricate, largest on posterior part of the belly, 36 mid-ventral scales between ventrolateral folds; 10 small precloacal pores arranged in an inverted “
<collectionCode id="88E2C3EEA1CCC2F19570764A8CA4961E" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
” shaped continuous series, followed by a series eight unpored, enlarged scales below it.
</paragraph>
<paragraph id="60E9221D69FCB4372EBA5009B150A43A">
Forelimbs and hindlimbs slender (
<abbrev id="ABBRID0EPVAK" xlink_title="forearm length, taken from the elbow to the wrist">FL</abbrev>
/
<abbrev id="ABBRID0ETVAK" xlink_title="snout to vent length">SVL</abbrev>
= 0.14,
<abbrev id="ABBRID0EXVAK" xlink_title="crus length, taken from the knee to the base of the foot">CL</abbrev>
/
<abbrev id="ABBRID0E2VAK" xlink_title="snout to vent length">SVL</abbrev>
= 0.18); digits strongly inflected at the joints, all bearing large recurved claw and enlarged subdigital lamellae; lamellae count beneath digit IV of right and left manus (given as basal + distal) is 5 + 7 and 5 + 8 respectively; lamellae count beneath digit IV of both right and left pes (given as basal + distal) is 8 + 9; dorsal scales on forelimbs heterogeneous in size; hind arm scales smooth and subimbricate; forearm scales small and granular towards proximal and towards distal end it is smooth, cycloid and imbricate; forearm scales intermixed with enlarged feebly keeled tubercles; dorsal scales of hindlimbs heterogeneous, intermixed with densely placed large, rounded and bluntly conical tubercles; scales on inner lateral and dorsolateral side of the thighs smooth, large and imbricate, rest of the scales are granular; scales on tibia are small, granular juxtaposed; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous in shape and size, granular juxtaposed; ventral scales of hind limbs smaller than those of belly, smooth, cycloid and subimbricate; scales on the knee, above cloaca and on thigh below the level of precloacal pores smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate.
</paragraph>
<paragraph id="8998594114F3A6A38B6DBB856AB6723A">
Tail complete, posterior
<quantity id="72351A90DF2F44911AE5003B50F80985" metricMagnitude="-3" metricUnit="m" metricValue="6.0" unit="mm" value="6.0">6 mm</quantity>
regenerated (
<abbrev id="ABBRID0EBWAK" xlink_title="tail length, from the cloaca to the tail tip">TL</abbrev>
=
<quantity id="81A8AB68CFAE1FFDD2F15338ECEA2F5C" metricMagnitude="-2" metricUnit="m" metricValue="6.4" unit="mm" value="64.0">64 mm</quantity>
), slender, gradually tapering towards tip; tail segments indistinct; dorsal scales small, granular, juxtaposed at the base, posteriorly size of the scales increases, flat, smooth, subimbricate, heterogeneous in shape and size; enlarged feebly keeled scales up to seventh segment of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; four bluntly conical spurs on both sides of the tail base.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFWAK" type="Colouration in life">
<paragraph id="3658C5A7A092351A948A9698B687D6E0">
<heading id="41507745A821A4E9FDE6F9127A58486E" reason="title">Colouration in life.</heading>
</paragraph>
<paragraph id="F92E119312CEC9DBA4E7F925EF27A2D7">
Head on top pale-brown with purplish tinge, indistinct brown irregular spots on occipital region; a short brown streak behind the posterior corner of the upper eyelid; a broad, dark-brown postorbital stripe continuing to above the ear opening; area between these two postorbital streaks is paler than dorsal head colour; an indistinct brown loreal stripe covering the nasal and an indistinct pale stripe above it; indistinct pale-yellow patch on occipital region; lips and mandibular region paler with indistinct yellow spots; dorsally and laterally neck and back pale yellowish-brown with dark-brown irregular reticulation starting from neck to sacrum; a dark-brown cross bar with irregular edges on nape; limbs pale-yellowish-brown with dark-brown reticulation; digits with alternative dark-brown and pale-yellow bars; tail with alternative 11 dark and 10 light bars in the original part, dark bars posteriorly more darker and broader; light bars posteriorly more whitish, first bar at the base of tail broken into two spots; fourth and fifth bar connected on left side; ventrally head, trunk and limbs whitish; tail with irregular brown and pale-yellow patches (Fig.
<figureCitation id="40CBA6E483DE3CAF85E163A315DE0080" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Cyrtodactylus barailensis sp. nov. (WII-ADR 971). A dorsal and B ventral view of the whole body; C dorsal, D ventral, and E lateral view of head; F dorsal tubercles on trunk; G ventral view of left manus, and H left pes. I C. barailensis sp. nov. in life. Scale bar: 10 mm." figureDoi="10.3897/vz.74.e124752.figure5" httpUri="https://binary.pensoft.net/fig/1102271">5 I</figureCitation>
).
</paragraph>
<caption id="F4C76D73A30E05F51F87882F035989EB" ID-DOI="10.3897/vz.74.e124752.figure5" ID-arpha="62A9A230-D983-5987-9B27-B3A850DA09D9" httpUri="https://binary.pensoft.net/fig/1102271" startId="F5">
<paragraph id="9CA3B835DB6C4640EC56F9FBBB999EA2">
<label id="672710045696006C8A88EF24942C434F">Figure 5.</label>
</paragraph>
<paragraph id="C4019411650398C3C7928BF670CD24A1">
Holotype of
<taxonomicName id="1C5138CA777B39D59A7883C1032233F0" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="barailensis" status="sp. nov.">
<emphasis id="9353AEB52A0839B2A0FA4459751CE773" italics="true">Cyrtodactylus barailensis</emphasis>
</taxonomicName>
<emphasis id="4F17B258138F82E2088D92DA6A928304" bold="true">
<taxonomicNameLabel id="7261F079EED76924C624281285129106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(WII-ADR 971).
<emphasis id="5430F8A719136F5BDFAB24384D985926" bold="true">A</emphasis>
dorsal and
<emphasis id="4B94EFE71FB7DEF1CF0A3C6F47C87106" bold="true">B</emphasis>
ventral view of the whole body;
<emphasis id="56B3B23B51680F5C289281812B4B3907" bold="true">C</emphasis>
dorsal,
<emphasis id="E43637DE914A1780CB05F5CC1C286120" bold="true">D</emphasis>
ventral, and
<emphasis id="3EC8EEBE4DC0EBE3F723F68DB665EF54" bold="true">E</emphasis>
lateral view of head;
<emphasis id="299093C2C6968F12C578BDC487F9A5D9" bold="true">F</emphasis>
dorsal tubercles on trunk;
<emphasis id="51F8263A50CE7B2F4C27C94E1B3E867E" bold="true">G</emphasis>
ventral view of left manus, and
<emphasis id="D1978F82733FE0703855A647871D1C2B" bold="true">H</emphasis>
left pes.
<emphasis id="275AAB25A558833059CEF36085039A2E" bold="true">I</emphasis>
<taxonomicName id="AB842A9AB90F6E171BC2053A254FF3B3" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="barailensis" status="sp. nov.">
<emphasis id="35FA803203DBD601DA4C4BBC5FEBD1B7" italics="true">C. barailensis</emphasis>
</taxonomicName>
<emphasis id="A5889D4FFDF22294E1104248B1AD27CD" bold="true">
<taxonomicNameLabel id="4D25415B502ED1D2AC954F2E68E9E0AF" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life. Scale bar: 10 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EIYAK" type="Colouration in preservative">
<paragraph id="3F3A07ABDF435B848856571ED2D7FEDF">
<heading id="268576E696C85CBEF6ACE07BB0146944" reason="title">Colouration in preservative.</heading>
</paragraph>
<paragraph id="3AEAA1985598988CD6F860301CD877A8">Top of head pale-greyish-brown; neck, back, limbs and tail dorsally light-grey with dark-brown markings; marking pattern visible as those in life condition; ventrally whitish with brown specks on tail.</paragraph>
</subSubSection>
<subSubSection id="SECID0ENYAK" type="comparisons">
<paragraph id="14D0FB9A232B3A6F26692F37E3A3349D">
<heading id="958D27993DBCA48BB94B2211A462F4DC" reason="title">Comparison.</heading>
</paragraph>
<paragraph id="9951D85C2E0F2B3BA7CD30CD9094DC17">
<taxonomicName id="EE9053E966AEE3837D898BCDE0285856" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="barailensis" status="sp. nov.">
<emphasis id="679DC03514D31445B628B499848A7215" italics="true">Cyrtodactylus barailensis</emphasis>
</taxonomicName>
<emphasis id="00B39F5DAB9E997297BF2E5796FA40A8" bold="true">
<taxonomicNameLabel id="17C8B2A5246C677A8B931A8EE7227C92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="BABBD9856947506023A006376C72444F" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aaronbaueri">
<emphasis id="BFEDA8D28AF318A0D8BFF17E65A50BE6" italics="true">C. aaronbaueri</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0ELZAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs.
<abbrev id="ABBRID0EPZAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="793A9D712CB9D66E2803E6C815C53029">DTR</collectionCode>
</abbrev>
22–28), 10 precloacal pores (vs. 6–8 pitted precloacal scales in female); differs from
<taxonomicName id="C7E78467229496C57ACA8B3BFD6D1224" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="agarwali">
<emphasis id="2F813B33F39020ACEC3C58955B05680C" italics="true">C. agarwali</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0E5ZAK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs. 21–25); differs from
<taxonomicName id="7D0BEFD37E7BBBE6D8E6D17B15BF20CC" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aunglini">
<emphasis id="E14EC5CA9F1196BC799529AE1F1C9154" italics="true">C. aunglini</emphasis>
</taxonomicName>
by fewer mid-ventral scale rows,
<abbrev id="ABBRID0EN1AK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="B6138E6F640E421D9FA0365D6E2646E0">MVSR</collectionCode>
</abbrev>
36 (vs.
<abbrev id="ABBRID0ER1AK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="982440F435D53428C14F6124C06316C6">MVSR</collectionCode>
</abbrev>
47–49), 32 paravertebral tubercle (vs.
<abbrev id="ABBRID0EV1AK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="01516156B9CBB4D5E7E756321431CD77">PVT</collectionCode>
</abbrev>
36–45), 17 dorsal tubercle rows (vs.
<abbrev id="ABBRID0EZ1AK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="4177BF69C88E3369FC38A8D4C8F4A1D8">DTR</collectionCode>
</abbrev>
21–26); differs from
<taxonomicName id="58382408F0DF7EBE182F74BCFAA5B5F6" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="bengkhuaiai">
<emphasis id="362F6A219783889A99D576C8CD14204A" italics="true">C. bengkhuaiai</emphasis>
</taxonomicName>
by having fewer dorsal tubercle rows,
<abbrev id="ABBRID0EI2AK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs.
<abbrev id="ABBRID0EM2AK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="CDB8C76150397B8E86C41F6C10B7F349">DTR</collectionCode>
</abbrev>
22–26), 10 precloacal pores (vs. 6–8 pitted precloacal scales in females); differs from
<taxonomicName id="9E5BD89C0E7609F10876F38756DD0D1A" authorityName="Bauer" authorityYear="2002" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="brevidactylus">
<emphasis id="1FBB304649C780BC1729670DDDC26DFC" italics="true">C. brevidactylus</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0E22AK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs.
<abbrev id="ABBRID0E62AK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="06A81101737E1A5DCA2C3EB0738995EB">DTR</collectionCode>
</abbrev>
27–30), fewer paravertebral tubercles,
<abbrev id="ABBRID0ED3AK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
32 (vs.
<abbrev id="ABBRID0EH3AK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="8E95BF5EECABA669338DBE8F08E7B2D0">PVT</collectionCode>
</abbrev>
38–42), enlarged dark blotches on head and dorsum absent (vs. large dark blotches on dorsum between nape and sacrum); differs from
<taxonomicName id="EABEFCE03889017E0353ED14C92A1044" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="chrysopylos">
<emphasis id="8E9F43C07D9DB079351D88DB4FEF18B3" italics="true">C. chrysopylos</emphasis>
</taxonomicName>
by presence of precloacal pores (vs.
<abbrev id="ABBRID0EW3AK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
absent in female), fewer mid-ventral scales,
<abbrev id="ABBRID0E13AK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="2A112C9C533A41ECF7DC7ED44D78A4AB">MVSR</collectionCode>
</abbrev>
36 (vs.
<abbrev id="ABBRID0E53AK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="B190EFFB926D5651D71582040C6BBF6A">MVSR</collectionCode>
</abbrev>
37–55); differs from
<taxonomicName id="7A8782B88B14C81D3E62796A2C88E829" authorityName="Liu &amp; Rao" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="dianxiensis">
<emphasis id="D9BC78A650EDB17A9A3D42BA44C9B5DD" italics="true">C. dianxiensis</emphasis>
</taxonomicName>
by smaller body size,
<abbrev id="ABBRID0EN4AK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="2B3476EA79E8E71789834C08B534E4A2" metricMagnitude="-2" metricUnit="m" metricValue="6.88" unit="mm" value="68.8">68.8 mm</quantity>
(vs.
<abbrev id="ABBRID0ER4AK" xlink_title="snout to vent length">
<collectionCode id="C6EB39E54B56737F178CB707E68F6B1A">SVL</collectionCode>
</abbrev>
<quantity id="A922C2501EAEA10CEB052DE653E65E07" metricMagnitude="-2" metricUnit="m" metricValue="7.5" unit="mm" value="75.0">75 mm</quantity>
), presence of precloacal pores (vs.
<abbrev id="ABBRID0EV4AK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
absent in female), fewer subdigital lamellae, 12 or 13 under fourth finger and 16 under fourth toe (vs. 16 or 17 under fourth finger and 19 or 20 under fourth toe); differs from
<taxonomicName id="866247A3DB9B0B840613F169C7467F2E" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="gansi">
<emphasis id="BE1FDC54360815987660B7DC34AB24CA" italics="true">C. gansi</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EE5AK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs.
<abbrev id="ABBRID0EI5AK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="ECD0FE937E1E31411058D4C9C388D7A0">DTR</collectionCode>
</abbrev>
20–25), ventrolateral fold present on trunk (vs. absent), fewer precloacal pores,
<abbrev id="ABBRID0EM5AK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
10 (vs.
<abbrev id="ABBRID0EQ5AK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
<quantity id="5254E19837A91353CAB6E40738A063FF" metricMagnitude="-1" metricUnit="m" metricValue="3.302" unit="in" value="13.0">13 in</quantity>
female); differs from
<taxonomicName id="0B8E4E440985209DDB9854A209A222EA" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="jaintiaensis">
<emphasis id="C3C138ABB31F713F8603DEDE66755FFD" italics="true">C. jaintiaensis</emphasis>
</taxonomicName>
by much smaller body size,
<abbrev id="ABBRID0E65AK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="1D592E8B4D7C2EDC8DB8F72D9C54085D" metricMagnitude="-2" metricUnit="m" metricValue="6.88" unit="mm" value="68.8">68.8 mm</quantity>
(vs.
<abbrev id="ABBRID0ED6AK" xlink_title="snout to vent length">
<collectionCode id="2E9C60AD1AF35954D6B79746F172DE6E">SVL</collectionCode>
</abbrev>
<quantity id="0D1E1953363D2C699653A2FAD26EC10B" metricMagnitude="-2" metricUnit="m" metricValue="9.62" unit="mm" value="96.2">96.2 mm</quantity>
in female), fewer mid-ventral scale rows,
<abbrev id="ABBRID0EH6AK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="BA8E9F14E19443E3716D7B504DC893A4">MVSR</collectionCode>
</abbrev>
36 (vs.
<abbrev id="ABBRID0EL6AK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="9814F8DB647EDA912F47CFF73236DA69">MVSR</collectionCode>
</abbrev>
40–42), fewer precloacal pores,
<abbrev id="ABBRID0EP6AK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
10 (vs. 12
<abbrev id="ABBRID0ET6AK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
in female); differs from
<taxonomicName id="91436873DF8F720F83B59DDF8657A584" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="A5DD8A1FCE6E1EB945ED5884A670A71A" italics="true">C. kiphire</emphasis>
</taxonomicName>
<emphasis id="88F321955D1EB8E7B46C499A465AACBD" bold="true">
<taxonomicNameLabel id="4B458379BAB7EA4661950F54594514FD" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the number of paravertebral tubercles 32 (vs.
<abbrev id="ABBRID0EEABK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="936C0E80A7A02F6EB4C744B71D461EDE">PVT</collectionCode>
</abbrev>
26 or 29); differs from
<taxonomicName id="C3BD4AF1361A74F84D8B502F2F4AA4C7" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="lungleiensis">
<emphasis id="841202DB56314B22E6EAEC284EF53C99" italics="true">C. lungleiensis</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0ETABK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs. 24–28), 10 precloacal pores (vs. 5–7 pitted precloacal scales in females); differs from
<taxonomicName id="00F9EDBD795FF7DD115C17BE566AAD7C" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="montanus">
<emphasis id="A97A4440B04BF2E9E5C7A8EAABD8A4A5" italics="true">C. montanus</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0ECBBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs.
<abbrev id="ABBRID0EGBBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="E6CD0C7098B6214073271942B27D3F17">DTR</collectionCode>
</abbrev>
21–23), fewer paravertebral tubercles,
<abbrev id="ABBRID0EKBBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
32 (vs.
<abbrev id="ABBRID0EOBBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="5F1907FD429C9B97D43D138131200782">PVT</collectionCode>
</abbrev>
37–43), precloacal pores present (vs.
<abbrev id="ABBRID0ESBBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
absent in female); differs from
<taxonomicName id="F4F34404CD97D1F504BC4ED576996219" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="myaleiktaung">
<emphasis id="5711F8E23A539651E827093D2E71BFC4" italics="true">C. myaleiktaung</emphasis>
</taxonomicName>
by fewer mid-ventral scale rows,
<abbrev id="ABBRID0EBCBK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="6D365CD5F51714D9EAD32C699365CBBB">MVSR</collectionCode>
</abbrev>
36 (vs.
<abbrev id="ABBRID0EFCBK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="FA5B15BC7424DFB085311A09888DE958">MVSR</collectionCode>
</abbrev>
57), precloacal pores present (vs.
<abbrev id="ABBRID0EJCBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
absent), broad regular dark bands absent on dorsum (vs. present); differs from
<taxonomicName id="E8120A72AF813AA30A2175A32BFA8B68" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="nagalandensis">
<emphasis id="8CA81DDA91EC20F124A5CEAD26D55786" italics="true">C. nagalandensis</emphasis>
</taxonomicName>
by presence of 10 precloacal pores (vs. six pitted precloacal scales in female), 32 paravertebral tubercles (vs.
<abbrev id="ABBRID0EYCBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="E9624C7EB910317EDF387D03F8DF6AD4">PVT</collectionCode>
</abbrev>
35–37); differs from
<taxonomicName id="678BC77F913A210926657F4FD7F7FDF2" authorityName="Mahony &amp; Kamei" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namtiram">
<emphasis id="52BD93A198F17F2506D6B943B3C6DFD4" italics="true">C. namtiram</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EHDBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs.
<abbrev id="ABBRID0ELDBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="0006DDEE9906B1FE0C8EAA8C039F0347">DTR</collectionCode>
</abbrev>
21); differs from
<taxonomicName id="695AE568F97D980E109C24AE89977AD0" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngopensis">
<emphasis id="04092CBBF046442267BBF75E6460F1FA" italics="true">C. ngopensis</emphasis>
</taxonomicName>
by presence of 10 precloacal pores in female, (vs. 0–6 pitted precloacal scales in female); differs from
<taxonomicName id="E868560F1E9734A0226E2836A77EEB5A" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="E1BD0A1DA0514CBD7B632CF9F501AC28" italics="true">C. septentrionalis</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EFEBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs.
<abbrev id="ABBRID0EJEBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="77599ED7F6DDEB8F73FD8350641133A8">DTR</collectionCode>
</abbrev>
23 or 24), less paravertebral tubercles,
<abbrev id="ABBRID0ENEBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
32 (vs.
<abbrev id="ABBRID0EREBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="EAE9AFD62826CDA5FA1DB48324D27F3C">PVT</collectionCode>
</abbrev>
38–42), 10 precloacal pores present (vs. 14 precloacal scales with indistinct depression in female); differs from
<taxonomicName id="4BCC36B3B09C2CE33C60C609D8F8B4E2" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siahaensis">
<emphasis id="FB22DBF73970E26C98B20D52E0182083" italics="true">C. siahaensis</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EAFBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs.
<abbrev id="ABBRID0EEFBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="3EFBBD5AFF10C122FA20C80C2396C627">DTR</collectionCode>
</abbrev>
22–24), presence of precloacal pores in female (vs.
<abbrev id="ABBRID0EIFBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
absent in female); differs from
<taxonomicName id="A0327654DBAFD19BA398E3251EEFB0AE" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="vairengtensis">
<emphasis id="A770F9303CA5C7D55A437D7C6310291F" italics="true">C. vairengtensis</emphasis>
</taxonomicName>
by the number of dorsal tubercle rows,
<abbrev id="ABBRID0EXFBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17 (vs.
<abbrev id="ABBRID0E2FBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="4B0A2386437575E6A13AD20F18DA17FE">DTR</collectionCode>
</abbrev>
22–23), 32 paravertebral tubercles (vs.
<abbrev id="ABBRID0E6FBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="D8243F29DE49E88102660917B106BA22">PVT</collectionCode>
</abbrev>
34–39), 10 precloacal pores present (vs. 5–9 precloacal pits in female). Morphological differences with other members of
<taxonomicName id="17AB4FD109A9F0EFB824A8E39C9C940E" baseAuthorityName="Jerdon" baseAuthorityYear="1870" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="7926E0E60C2B2AB331C5F162D3A81F10" italics="true">khasiensis</emphasis>
</taxonomicName>
group is presented in Table
<tableCitation id="5BF013D3B9495F3524E24F96F9D92834" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERGBK" type="Sequence divergence">
<paragraph id="7C0049B1924B22FD5DA10B230A2AA47E">
<heading id="B0B17004120025DDCD70277B7B3ACCBC" reason="title">Sequence divergence.</heading>
</paragraph>
<paragraph id="7887D709337977C5137BC7F001D6DBDE">
<taxonomicName id="11A019310ED45945A9534FE490B79BE4" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="barailensis" status="sp. nov.">
<emphasis id="25E6FF380934A07C60A84E395ABDD412" italics="true">Cyrtodactylus barailensis</emphasis>
</taxonomicName>
<emphasis id="E3EFB0D585C3FF3FCA8B05024BDF6032" bold="true">
<taxonomicNameLabel id="309B9B82E4F4CE83F9B34B93797A8C57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a high genetic divergence of 10.9 % from its closely related
<taxonomicName id="1D5B4540B60B3C2455FB841364384BC1" authorityName="Mahony &amp; Kamei" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namtiram">
<emphasis id="4F7BA7F8B111C1EE6A463BA96CB6B25C" italics="true">C. namtiram</emphasis>
</taxonomicName>
. With other members of the clade,
<taxonomicName id="09EFAD8242AA1FC34F7BCB941FF79E56" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="66EC246FF543FDE18DC15674F71E8F23" italics="true">C. kiphire</emphasis>
</taxonomicName>
<emphasis id="06E294F42E3F92A1BD732A4FB850CBB3" bold="true">
<taxonomicNameLabel id="DE4C13F95B363F66576F71C010579A28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a genetic divergence of 10.7 % and 22.3 % in the ND 2 gene.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3HBK" type="etymology">
<paragraph id="C2A055C2F6F4AEA250E6D8B9A45B8DA5">
<heading id="545AAB5F77882AB9FF882F5C4B59E367" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="C1C74D550F578A7714F3003B736CF333">
The specific epithet is a toponym derived from the name of the hill range “ Barail ” where the
<typeStatus id="8403B414FEEA81ACF2858CDD9587830F">type</typeStatus>
locality of the species lies.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBIBK" type="Suggested common name">
<paragraph id="B867BA96A8DB7210668A22F50726F75C">
<heading id="3BB16F3C6B4BCEA97CE4CA202E889A9A" reason="title">Suggested common name.</heading>
</paragraph>
<paragraph id="207A5EA24CA7978086896D13C2D635AE">Barail Hills bent-toed gecko.</paragraph>
</subSubSection>
<subSubSection id="SECID0EGIBK" type="distribution">
<paragraph id="750F79DA3D770EC3FB4586B481C015FA">
<heading id="21E9BC883F8FA3AA01577E1C27A45E43" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="52480E70FB1130A6E26EB4A6DCE59551">
During our two-day survey we only located a single individual of this species. Thus, the new species is currently known only from the
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locality, in Peren District,
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,
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. We recorded this species on the trunk of a small tree at a height of approximately
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from the ground in the Athibung Reserve Forest at approximately 20: 00 hrs on
<date id="08E9E3E3025C754C49F50712CD4C17D7" value="2021-08-14">14 August 2021</date>
. The forest
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is semi-evergreen with relatively little anthropogenic pressure.
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156
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<emphasis id="90AFCB7FB79F58FF3F83EE8DAE060509" italics="true">Dolichopoda gasparoi</emphasis>
<bibRefCitation id="6CDFCC604BF892D4491C071756C3BAEA" DOI="10.11646/zootaxa.1923.1.1" author="Rampini M &amp; Di Russo C &amp; Pavesi F &amp; Cobolli M" journalOrPublisher="Zootaxa" pagination="1-17" refId="B12" refString="Rampini M, Di Russo C, Pavesi F, Cobolli M (2008) The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae). Zootaxa 1923: 1 - 17. https: // doi. org / 10.11646 / zootaxa. 1923.1. 1" title="The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae)." volume="1923" year="2008">Rampini &amp; Di Russo, 2008</bibRefCitation>
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<paragraph id="2DBE8087BF94701A071E69C9F08CDDF8">
<heading id="75DF715284A5CA1EEE09D0D21FDF827E" reason="title">Material examined.</heading>
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<paragraph id="AC70E57F5FE779E16F0FEB473FAE0752">
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-
<emphasis id="303755BA0786387BE12557DECF1F180C" bold="true">
<collectingRegion id="74B0D9471BA9426A75094C5B38707FC1" country="Greece" name="Ionia Nisia">Ionian Islands</collectingRegion>
</emphasis>
•
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; Νomos
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,
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,
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;
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</date>
;
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leg. (
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)
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•
<materialsCitation id="83B0251570547B83175579AE3D96FE3E" collectingDate="2016-08-23" collectorName="K. Bakolitsas" country="Greece" county="Lefkadas" location="cave Pouliezou" municipality="Lefkada Isl." specimenCount="2" specimenCount-male="2" stateProvince="Ionia Nisia">
<specimenCount id="8D9185E9DC03B9E844C7B7582BC4BE4A" count="2" type="male">2 ♂</specimenCount>
; Νomos
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,
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;
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;
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leg. (
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•
<materialsCitation id="6D93CF1BEABA026EFA2413ADBFC9D70B" collectingDate="2016-09-25" collectorName="K. Bakolitsas" country="Greece" county="Lefkadas" location="cave Tsotsonia" municipality="Lefkada Isl." specimenCount="4" specimenCount-male="4" stateProvince="Ionia Nisia">
<specimenCount id="1E948171847F7553372C83E90DF69C65" count="4" type="male">4 ♂</specimenCount>
; Νomos
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,
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<location id="C70FA4EFFABFF184CB8F6B6CFE125C00" LSID="urn:lsid:plazi:treatment:0C80CCAAF19651FE8C2A886279550539:C70FA4EFFABFF184CB8F6B6CFE125C00" country="Greece" county="Lefkadas" municipality="Lefkada Isl." name="cave Tsotsonia" stateProvince="Ionia Nisia">cave Tsotsonia</location>
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<collectingDate id="4065666A4788185570A4AE8A4FA4C3A8" value="2016-09-25">25 Sept. 2016</collectingDate>
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;
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leg. (
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.
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<heading id="5846F1E49A8062B74B71197EB66F6452" reason="title">Comment.</heading>
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<taxonomicName id="F92BF6B00F90FB7688D7426C3D99C265" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="gasparoi">
<emphasis id="1EF5A2B9C758E7A956AD674FE915785B" italics="true">Dolichopoda gasparoi</emphasis>
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is endemic to Lefkada Isl. (
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). Our records indicate that the species is widespread on the island.
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<mods:namePart id="7E26282E6BC1DADF1B8773EADCED6487">Deepak, V.</mods:namePart>
<mods:nameIdentifier id="684F3C20F98FDB815755275C0A18BB5F" type="ORCID">0000-0002-8826-9367</mods:nameIdentifier>
<mods:affiliation id="9130AC36B4A7CC3D03EBDD0909E0A8E8">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK &amp; School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne NE 1 7 RU, UK</mods:affiliation>
</mods:name>
<mods:name id="C5EF85AAEB8B6AEFBE351F8CA88F1720" type="personal">
<mods:role id="CF7280CB449E33F5B9E4A1C77A23B457">
<mods:roleTerm id="67D25E2B9595A1A788042111B05FD0DA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DFBA1B4D10E7652F87170E75F3EEF8F2">Das, Abhijit</mods:namePart>
<mods:nameIdentifier id="C00029AE4AF356E57C32AE78FA5AAEB2" type="ORCID">0000-0002-5851-8457</mods:nameIdentifier>
<mods:affiliation id="2C11865E6AAEE13D90CB063648DDC091">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
</mods:name>
<mods:typeOfResource id="09F86B83B40016E0F5243BDFD242BAD0">text</mods:typeOfResource>
<mods:relatedItem id="FD2258827C35799DB39007E0E471DB12" type="host">
<mods:titleInfo id="111C58AF8323B530D4FB73442842BEBD">
<mods:title id="89F0C3A99DE03613D368670C4CC3BDFD">Vertebrate Zoology</mods:title>
</mods:titleInfo>
<mods:part id="AFE62DD89A3291D87B607F99550E8689">
<mods:date id="94C5E2B0441F2BCCC9742E5E664E1279">2024</mods:date>
<mods:detail id="C0DC0D9E812D5DCE0555737E61A388D1" type="pubDate">
<mods:number id="6E6FA9D88749B2A1582760008BE9EE98">2024-07-29</mods:number>
</mods:detail>
<mods:detail id="7A4F6C6A4F676C2D600720BD03B1D091" type="volume">
<mods:number id="78BB1F30864BA568CEF9894CCBDA36F6">74</mods:number>
</mods:detail>
<mods:extent id="6DD44EB1680DDCEBF7A6F569BFA045CD" unit="page">
<mods:start id="8136BFEF25489A5846A6DEB4DB14E5AD">453</mods:start>
<mods:end id="A48CDEADAB122E2A0B42B3C1E144B211">486</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8D3037C8E91840849EE454D48F983398">journal article</mods:classification>
<mods:identifier id="91261E89FB86011DEEA619C03C271DBD" type="DOI">10.3897/vz.74.e124752</mods:identifier>
<mods:identifier id="F006CF2EF0A64F67A6AB6BC1F2F8E0F2" type="ZooBank">154CE236-EFA8-4411-834B-234A9B45A63F</mods:identifier>
</mods:mods>
<treatment id="0F6F61B9C7765FC09DEBF74295D25A34" ID-DOI="http://doi.org/10.5281/zenodo.13124031" ID-Zenodo-Dep="13124031" ID-arpha="0F6F61B9-C776-5FC0-9DEB-F74295D25A34" ID-zoobank="https://zoobank.org/D04AF555-89A2-467F-B5CE-419328F6934F" LSID="urn:lsid:zoobank.org:act:D04AF555-89A2-467F-B5CE-419328F6934F" LSID-ZBK="urn:lsid:zoobank.org:act:D04AF555-89A2-467F-B5CE-419328F6934F" httpUri="http://treatment.plazi.org/id/0F6F61B9C7765FC09DEBF74295D25A34">
<subSubSection id="10200B5998F80FD13D200FC30F5A6B79" type="nomenclature">
<paragraph id="814F80E9B375D3E010EE6775E59D6AA8">
<taxonomicName id="A4E3A8BEB97759E4286C0B6C4F1F44DD" authority="Boruah, Narayanan, Lalronunga, Deepak &amp; Das" authorityName="Boruah, Narayanan, Lalronunga, Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngengpuiensis" status="sp. nov.">
<emphasis id="265C304D67170542C8B657412DD89A3E" italics="true">Cyrtodactylus ngengpuiensis</emphasis>
Boruah, Narayanan, Lalronunga, Deepak &amp; Das
</taxonomicName>
<taxonomicNameLabel id="49ACD2263E35921568EF1AD12DB228AF" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9A60711384BFF5675C0252B6242B4F75" type="description">
<paragraph id="C03CE186CF8707804F626824A8B7EE80">
<figureCitation id="6F59AE6383E3AC01AB99D8705DDB983B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyrtodactylus ngengpuiensis sp. nov. A – J holotype (ADR-WII 1057), A dorsal and B ventral view of the whole body; C dorsal, D ventral, and E lateral view of head; F dorsal tubercles on trunk; G ventral view of right manus, and H right pes, I precloacofemoral pores; J holotype (ADR-WII 1057), and K paratype (WII-ADR 991) in life. Scale bar: 10 mm. J, K not to scale." figureDoi="10.3897/vz.74.e124752.figure7" httpUri="https://binary.pensoft.net/fig/1102273">Figure 7</figureCitation>
;
<tableCitation id="347830FDAAC71F101154C147085F744C" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">Tables 2, S 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3ECK" type="Holotype">
<paragraph id="F4C4DBF561738995EEE9DD38DDF85F1B">
<heading id="2C2B771922F8569B4FC2B0E346E03373" reason="title">
<typeStatus id="2F6BDF7D330A577A14A219DE3A10865A" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="D15C57331985C22210458AA9E9E7BFB4">
<materialsCitation id="A8F3783BDB43A7D62D7278323050A5EC" collectingDate="2021-09-08" collectorName="Abhijit Das" country="India" elevation="160" latitude="22.4906" location="Lawngtlai District" longLatPrecision="7" longitude="92.7575" specimenCount="1" stateProvince="Mizoram" typeStatus="holotype">
Adult male (WII-ADR 1057; Fig.
<figureCitation id="4A916E0A813E2ECDEE3E557088B3D685" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyrtodactylus ngengpuiensis sp. nov. A – J holotype (ADR-WII 1057), A dorsal and B ventral view of the whole body; C dorsal, D ventral, and E lateral view of head; F dorsal tubercles on trunk; G ventral view of right manus, and H right pes, I precloacofemoral pores; J holotype (ADR-WII 1057), and K paratype (WII-ADR 991) in life. Scale bar: 10 mm. J, K not to scale." figureDoi="10.3897/vz.74.e124752.figure7" httpUri="https://binary.pensoft.net/fig/1102273">7 A – J</figureCitation>
), from Ngengpui Wildlife Sanctuary (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJFCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[92.757500,22.490600]}">
<geoCoordinate id="7893EF7CF7BF8E0AFB7074458717647B" degrees="22.4906" direction="north" orientation="latitude" precision="5" value="22.4906">22.4906 ° N</geoCoordinate>
;
<geoCoordinate id="B99AA9B8F8D830BEFB37855ED3892783" degrees="92.7575" direction="east" orientation="longitude" precision="5" value="92.7575">92.7575 ° E</geoCoordinate>
</named-content>
;
<elevation id="DACE750CE532BCC2892D09FADEBA5208" metricMagnitude="2" metricUnit="m" metricValue="1.6" unit="m" value="160.0">
elevation
<quantity id="B72B4D2F6FDC9A80340AE080C8D37817" metricMagnitude="2" metricUnit="m" metricValue="1.6" unit="m" value="160.0">160 m</quantity>
a. s. l.
</elevation>
) (Fig.
<figureCitation id="3880332778AFA36BFF0C2D1599353545" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 A</figureCitation>
),
<location id="0A8B47636EBAE2F646C2471D0BF2A076" LSID="urn:lsid:plazi:treatment:0F6F61B9C7765FC09DEBF74295D25A34:0A8B47636EBAE2F646C2471D0BF2A076" country="India" latitude="22.4906" longLatPrecision="7" longitude="92.7575" name="Lawngtlai District" stateProvince="Mizoram">Lawngtlai District</location>
,
<collectingRegion id="6DE0706B502BEB3612B2264D3EAB2D49" country="India" name="Mizoram">Mizoram</collectingRegion>
,
<collectingCountry id="B27BE8C1B29F9CE635CAFD7E1CD6FD7D" name="India">India</collectingCountry>
collected by
<collectorName id="705AC7EB4C2D6AB19BF6AE5D15D43CAD">Abhijit Das</collectorName>
, Bitupan Boruah and Samuel Lalronunga on
<date id="3D00617E7CC5BC18416A3E86A059A354" value="2021-09-08">
<collectingDate id="57133D6E2E9AA415808F49A46080C750" value="2021-09-08">8 September 2021</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption id="5303538D07AD17608EC264402B8EC947" ID-DOI="10.3897/vz.74.e124752.figure7" ID-arpha="9669FBEA-04B2-5ECE-84FE-8B33822D2368" httpUri="https://binary.pensoft.net/fig/1102273" startId="F7">
<paragraph id="147C36DFB02262EA6FD4915211A8A994">
<label id="5693DE070D64C94DC88B1E40BD440A57">Figure 7.</label>
</paragraph>
<paragraph id="E8AA3229F517BCD3FEB031AA46B700CA">
<taxonomicName id="6651003DF428FA2BE83846FE5633F419" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngengpuiensis" status="sp. nov.">
<emphasis id="D17AF8DA9229721634129656A75FF77D" italics="true">Cyrtodactylus ngengpuiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60914E3BC8B73C3998CFE0054784515F" rank="species">
<emphasis id="48984A7A441878052D4818B8B6D9D536" bold="true">sp. nov</emphasis>
.
</taxonomicNameLabel>
<emphasis id="98DAC7B51925693A33E4D59C2CBB7F64" bold="true">A</emphasis>
–
<emphasis id="F3D90BE3D97EEBA3C8DEFF2E057C0839" bold="true">J</emphasis>
holotype (ADR-WII 1057),
<emphasis id="EFA8FE720FD4B406B075D5FEAC244BB2" bold="true">A</emphasis>
dorsal and
<emphasis id="D7864A54964BCD14E4267B0005ECE3E1" bold="true">B</emphasis>
ventral view of the whole body;
<emphasis id="A92822EFC8E1786B1A2AAC73C5059B79" bold="true">C</emphasis>
dorsal,
<emphasis id="239D1BD81A24F6C0B9ED0A95AAA3018D" bold="true">D</emphasis>
ventral, and
<emphasis id="75F292161425A8574A37D6AECDF3AF83" bold="true">E</emphasis>
lateral view of head;
<emphasis id="0CC62A22112EEE2C24C4AD3F355B2F94" bold="true">F</emphasis>
dorsal tubercles on trunk;
<emphasis id="FB2B451271D41322223BB18861C3BB9C" bold="true">G</emphasis>
ventral view of right manus, and
<emphasis id="A287893FB115FE7FBF4902E11CE0B5D9" bold="true">H</emphasis>
right pes,
<emphasis id="DA0479C623A637DA3314FA8CE62398BA" bold="true">I</emphasis>
precloacofemoral pores;
<emphasis id="FBD5865EA2AE63B124C970E6D89E4F8F" bold="true">J</emphasis>
holotype (ADR-WII 1057), and
<emphasis id="84D05F05D82233A864CA7A4AAD4D2046" bold="true">K</emphasis>
paratype (WII-ADR 991) in life. Scale bar: 10 mm.
<emphasis id="F0CC3B570F73174CE5E012A84384D1BD" bold="true">J, K</emphasis>
not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EIHCK" type="Paratypes">
<paragraph id="B0BBD46E9AB8ADE694452243FA2D6F7F">
<heading id="86C242D3991DD180F6C04984926AAB34" reason="title">
<typeStatus id="ED108A0D59FF6DCD82571193318A5EF6" type="paratype">Paratypes</typeStatus>
.
</heading>
</paragraph>
<paragraph id="D0CFD4E8796C3198E29114BFF63A9EB4">
<specimenCount id="8CAE3C4A05313DFD2FD3F763D9B9B4AB" count="2" type="adult">Two adult</specimenCount>
females (WII-ADR 991 and WII-ADR 1058) and
<specimenCount id="DC1BA10AF099CF6AC67745612ABB086D" count="1" type="subadult">one subadult</specimenCount>
female (WII-ADR 1059) collected from the same locality in Ngengpui Wildlife Sanctuary by the same team on
<date id="2F13415877074AE32BB76A4E4CE4F96F" value="2021-09-07" valueMax="2021-09-08" valueMin="2021-09-07">7 and 8 September 2021</date>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENHCK" type="diagnosis">
<paragraph id="64392709E39D43C2AD45AF6E46EB2B47">
<heading id="8687EF43ABFCD37B89FF41F6928CB6DF" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D7F9B0EB31487A70E7D010A2AEE6D0F3">
Medium-sized gecko (
<abbrev id="ABBRID0ETHCK" xlink_title="snout to vent length">SVL</abbrev>
at least
<quantity id="7E142AB1EA6EBFD970C539D87AC72FA1" metricMagnitude="-2" metricUnit="m" metricValue="6.12" unit="mm" value="61.2">61.2 mm</quantity>
in adult male and
<quantity id="8804F53AF7A4CB78F15DD840572082CD" metricMagnitude="-2" metricUnit="m" metricValue="7.305" metricValueMax="7.409999999999999" metricValueMin="7.2" unit="mm" value="73.05" valueMax="74.1" valueMin="72.0">72–74.1 mm</quantity>
in adult females); 9–12 supralabials; 8–11 infralabials; 18–20 bluntly conical and feebly keeled dorsal tubercles; 29–34 paravertebral tubercles; 38 or 39 midventral scale rows between the weak ventrolateral folds; no precloacal groove; at least 27 precloacofemoral pores in continuous series in male and 10–16 small precloacal pores in females; short dark-brown bars between a pair of dorsolateral stripes or enlarged irregular dark-brown spots present on dorsum.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXHCK" type="description">
<paragraph id="B09E451E770A929737C034433A210444">
<heading id="42107C89E15888DDB87D220637BF408D" reason="title">
Description of
<typeStatus id="A01CC01AB598B2AB52C51EE08BE88682" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="9E68392E6CB3DCA2E02DB49AAB00819D">
<typeStatus id="7BE14BD8D0836391338F3B2CC44214D5" type="holotype">Holotype</typeStatus>
well preserved except an incision below left axilla ventrolaterally. Snout-vent length
<quantity id="DE742026CB23345767DEEE5AD94D3237" metricMagnitude="-2" metricUnit="m" metricValue="6.12" unit="mm" value="61.2">61.2 mm</quantity>
. Head moderately large (
<abbrev id="ABBRID0E4HCK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
/
<abbrev id="ABBRID0EBICK" xlink_title="snout to vent length">SVL</abbrev>
= 0.27), dorsoventrally depressed, longer than width (
<abbrev id="ABBRID0EFICK" xlink_title="maximum head width">HW</abbrev>
/
<abbrev id="ABBRID0EJICK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.68), distinct from neck, broader at occipital region, snout tip rounded in both dorsal and lateral view, loreal region convex, canthus rostralis rounded, indistinct, interorbital space flat, a longitudinal furrow on dorsal surface of the snout, snout short (
<abbrev id="ABBRID0ENICK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
/
<abbrev id="ABBRID0ERICK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.4), longer than orbit (
<abbrev id="ABBRID0EVICK" xlink_title="orbit diameter, taken between the anterior and posterior bony orbital borders">OD</abbrev>
/
<abbrev id="ABBRID0EZICK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
= 0.66), nostril nearly rounded, opening directed posterolaterally, ear opening oval and oblique, scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed, scales on upper eyelids heterogeneous, supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anterior and posterior end decreases, rostral wide, a short groove at the middle on top, rostral connected with nasals, supranasals, an internasal and first supralabials, two scales between the supranasals, larger than the rest of the granular snout scales, granular scales at parietal region and occipital region intermixed with slightly large rounded granular tubercles, dense in occipital and temporal region and size increases towards nape, nine supralabials on right and 10 on left side, size decreases towards angle of jaw, supralabials up to midorbit seven on right and eight on left side, a series of narrow and slightly elongated scales above the supralabials between nostril and anterior border of the orbit, mental as wide as rostral, triangular, connected with first infralabials, inner postmentals, nine infralabials on both side, size decreases towards angle of jaw, first infralabials connected with mental, second infralabial, inner and outer postmentals, inner pair of postmentals are larger than the outer postmentals, posterior margin of the inner postmentals bordered by seven granular scales of different size, two or three rows of enlarged scales along the infralabials starting below the outer postmentals, posteriorly size of those decreases, elongated and narrow, rest of the gular scales are small, granular juxtaposed, homogeneous, size increases towards the throat where they become imbricate.
</paragraph>
<paragraph id="C72998EF1F101931AED19135662BB2D7">
Habitus slender (
<abbrev id="ABBRID0E6ICK" xlink_title="maximum body width">BW</abbrev>
/
<abbrev id="ABBRID0EDJCK" xlink_title="snout to vent length">SVL</abbrev>
= 0.17,
<abbrev id="ABBRID0EHJCK" xlink_title="trunk length, between the axilla and groin">TRL</abbrev>
/
<abbrev id="ABBRID0ELJCK" xlink_title="snout to vent length">SVL</abbrev>
= 0.41), dorsoventrally depressed, dorsal scales granular, rounded, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, starting from occipital region to seventh segment of the tail, size increases towards posterior body, pronounced at sacrum and base of the tail, 18 dorsal tubercles across mid dorsum, 29 paravertebral tubercles, ventrolateral fold weak, ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate, largest on belly, 39 mid-ventral scales between ventrolateral fold, 27 precloacal femoral (
<abbrev id="ABBRID0EPJCK" xlink_title="precloacofemoral pores, a continuous series of pore-bearing scales that extends from the precloacal region onto the thighs">PcFP</abbrev>
) pores in a continuous series (Fig.
<figureCitation id="BC567E89397E6A1887B5F0FAC366C14B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyrtodactylus ngengpuiensis sp. nov. A – J holotype (ADR-WII 1057), A dorsal and B ventral view of the whole body; C dorsal, D ventral, and E lateral view of head; F dorsal tubercles on trunk; G ventral view of right manus, and H right pes, I precloacofemoral pores; J holotype (ADR-WII 1057), and K paratype (WII-ADR 991) in life. Scale bar: 10 mm. J, K not to scale." figureDoi="10.3897/vz.74.e124752.figure7" httpUri="https://binary.pensoft.net/fig/1102273">7 I</figureCitation>
), followed by eight unpored, large scales below the
<abbrev id="ABBRID0EXJCK" xlink_title="precloacofemoral pores, a continuous series of pore-bearing scales that extends from the precloacal region onto the thighs">PcFP</abbrev>
at the middle.
</paragraph>
<paragraph id="B86D497D2FC0D8F95721F4CE06B371FB">
Forelimbs and hindlimbs slender (
<abbrev id="ABBRID0E4JCK" xlink_title="forearm length, taken from the elbow to the wrist">FL</abbrev>
/
<abbrev id="ABBRID0EBKCK" xlink_title="snout to vent length">SVL</abbrev>
= 0.14,
<abbrev id="ABBRID0EFKCK" xlink_title="crus length, taken from the knee to the base of the foot">CL</abbrev>
/
<abbrev id="ABBRID0EJKCK" xlink_title="snout to vent length">SVL</abbrev>
= 0.19); digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae; lamellae beneath digit IV of right and left manus (given as basal + distal) is 5 + 8 and 5 + 9 respectively; lamellae beneath digit IV of right and left pes (given as basal + distal) is 4 + 10 and 5 + 10 respectively; dorsal scales on forelimbs heterogeneous, granular juxtaposed; dorsal scales of hindlimbs heterogeneous, granular, intermixed with densely placed large, rounded and bluntly conical tubercles, scales on inner lateral side of the thighs near knee are subimbricate; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous in shape and size, granular; scales on ventral side of hindlimbs nearly equal to those of belly, smooth, cycloid and subimbricate, but on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate.
</paragraph>
<paragraph id="CB462813635B5A117B7BE25C48B57121">
Tail complete (
<abbrev id="ABBRID0EPKCK" xlink_title="tail length, from the cloaca to the tail tip">TL</abbrev>
=
<quantity id="7E555ECB2F78270717E3CB6555F79152" metricMagnitude="-2" metricUnit="m" metricValue="7.0" unit="mm" value="70.0">70 mm</quantity>
), slender, gradually tapering towards tip, segments indistinct, dorsal scales granular, juxtaposed, flat, smooth, heterogeneous in shape and size; enlarged feebly keeled scales up to eight segments of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETKCK" type="Colouration in life">
<paragraph id="FB8969DF48545DD09B8EEEE1859351DC">
<heading id="E60BAD34429CF930F972FBEB16E9A22F" reason="title">Colouration in life.</heading>
</paragraph>
<paragraph id="F8B1A10FC3C17A9D866822E99F8591F2">
Fig.
<figureCitation id="5A9A19FBEA8CFC79EEE68E2621E548D1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyrtodactylus ngengpuiensis sp. nov. A – J holotype (ADR-WII 1057), A dorsal and B ventral view of the whole body; C dorsal, D ventral, and E lateral view of head; F dorsal tubercles on trunk; G ventral view of right manus, and H right pes, I precloacofemoral pores; J holotype (ADR-WII 1057), and K paratype (WII-ADR 991) in life. Scale bar: 10 mm. J, K not to scale." figureDoi="10.3897/vz.74.e124752.figure7" httpUri="https://binary.pensoft.net/fig/1102273">7 J</figureCitation>
. Top of head pale-brown with indistinct pale-cream coloured spots on anterior part, a few slightly dark-brown patches on upper eyelids, temporal and occipital region; supraciliary yellowish-brown; dark-brown postocular streak on each side which continues dorsolaterally along neck and trunk to tail base, on right side this stripe is disjunct; a pale-cream coloured steak on each side of the dark-brown post orbital streak; an indistinct brown stripe along loreal region and a pale-cream coloured streak above it; upper lip with irregular brown and pale-yellow spots; a pair of dark-brown stripe with irregular edge on neck interspaced by an elongated pale-cream coloured patch, posterior ends of these stripes are connected; an enlarged dark-brown spot at the middle of nape; a pair of narrow pale-cream coloured stripe on neck above the lateral dark-brown stripe running up to slightly behind the level of axilla; dorsum pale-brown with seven irregular shaped and sized, broad dark-brown cross bars, these bars posteriorly edged with narrow cream coloured patch, first two cross bars are shorter than the rests; a few enlarged whitish spots along lateral side of the trunk; dorsally limbs pale-brown intermixed with slightly darker spots and light patches; dorsally tail with alternating slightly dark-brown broad bands and narrow pale-cream coloured bands of irregular shape and size; spurs cream coloured; ventrally head and trunk whitish with pinkish tinge and scales with brown marbling; some scales above and below the cloaca are pale-yellow; tail with slightly dense brown marbling.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4KCK" type="Colouration in preservative">
<paragraph id="38DE13BF4B4CC56F529BFDDA8B03FCB0">
<heading id="F571B9B07FF75F62BC479A2904668BA2" reason="title">Colouration in preservative.</heading>
</paragraph>
<paragraph id="979401C451FA1F5B01AF98CF624EB51D">Dorsal and ventral colour nearly the same as that of in life; dorsal markings visible as that of life condition.</paragraph>
</subSubSection>
<subSubSection id="SECID0ECLCK" type="Morphological variation">
<paragraph id="C343237C74443B02ABFE52015DAB2A80">
<heading id="85E32DA20D27329194DCCB423A924B25" reason="title">Morphological variation.</heading>
</paragraph>
<paragraph id="2D32A4FDA796BA98BFC1E5A6BB0C320C">
Details of the variations in morphometric and meristic characters of the type series are provided in Table S 2. Apart from these dorsal markings among the
<typeStatus id="7BB7CC918C817442284EAB77A85A7A87" type="paratype">paratypes</typeStatus>
slightly varied. In the
<specimenCount id="3CF2D2A17A0896245BA9A60942906D33" count="2" type="generic" typeStatus="paratype">two paratypes</specimenCount>
(WII-ADR 1058 and WII-ADR 1059), dorsal bands are broken mid dorsally giving the appearance of enlarged spots (Fig. S 2). Precloacal pores in female are smaller than that of male.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHLCK" type="comparisons">
<paragraph id="A20BBF2F3AB53BFBBC4334595342C133">
<heading id="79CB560E423B6431A0A41F676F9F7AA4" reason="title">Comparison.</heading>
</paragraph>
<paragraph id="A26ABB6C8AE3E218CB3315F346D3BC41">
<taxonomicName id="80D5D41447342ECAD88A1B1E8A10ACAF" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngengpuiensis" status="sp. nov.">
<emphasis id="D7F613C4CC10D565765F4125B9D0BAF8" italics="true">Cyrtodactylus ngengpuiensis</emphasis>
</taxonomicName>
<emphasis id="C14A7CCF713F6F3876CE3F1E973916D2" bold="true">
<taxonomicNameLabel id="5F25D99D76E655FF4D67D16A49C15345" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from the members of the
<taxonomicName id="A12D66F4C9732D3E16BF25225094FB15" baseAuthorityName="Jerdon" baseAuthorityYear="1870" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="46F8E6D1419F55CD578F773F0F40C3EE" italics="true">C. khasiensis</emphasis>
</taxonomicName>
clade (except
<taxonomicName id="073645CD4EA3DFF99ACDE3084203E39A" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ayeyarwadyensis">
<emphasis id="11250F175CF6EB52E454E6C31C4B0FEF" italics="true">C. ayeyarwadyensis</emphasis>
</taxonomicName>
,
<taxonomicName id="80CB9B67CDD7ED69FD81F7349C4FA1FD" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="guwahatiensis">
<emphasis id="12FB1A9DE2E930F1BD001D26F1DA7E38" italics="true">C. guwahatiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="38C735E730E94413BB5E1861E9B54AAD" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="karsticola">
<emphasis id="E6CC0B1D807EBE855355A7A546D53186" italics="true">C. karsticola</emphasis>
</taxonomicName>
and
<taxonomicName id="64BD10A922AA0372057C7678FE34A890" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="tripuraensis">
<emphasis id="118D989B31522D32C20B38D7CC160CE0" italics="true">C. tripuraensis</emphasis>
</taxonomicName>
) by the presence of precloacofemoral pores in male (vs. no femoral pores in male of
<taxonomicName id="917BF8DE02BC6CE670BB251A8C60A6E1" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="agarwali">
<emphasis id="C21F8C547E522A98D35D11C04016CE73" italics="true">C. agarwali</emphasis>
</taxonomicName>
,
<taxonomicName id="4385251B4FB4DF3060173788380EF053" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="exercitus">
<emphasis id="6766B4FA46272712D63F7C6644F4AC98" italics="true">C. exercitus</emphasis>
</taxonomicName>
,
<taxonomicName id="CA6678CA6D6AD76909EF3B71AB94CA15" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kazirangaensis">
<emphasis id="BA4D2AD72DF21420C54C28E91FA88450" italics="true">C. kazirangaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="CF7E9186561B30557349E49C67932D8D" baseAuthorityName="Jerdon" baseAuthorityYear="1870" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="8001330B92D8B1B545B3424AF2C99397" italics="true">C. khasiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="DAC349B2C339B3AF436741F43DA55907" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="CC05CA36CF1BD5405D549C09FD976182" italics="true">C. septentrionalis</emphasis>
</taxonomicName>
,
<taxonomicName id="8F4B1B854F837250FA5B25156271CAB9" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="urbanus">
<emphasis id="2C13696549A2989FE9D90932F441AD13" italics="true">C. urbanus</emphasis>
</taxonomicName>
).
<taxonomicName id="E7916E908AD8FDBA6596BD02BA4694E7" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngengpuiensis" status="sp. nov.">
<emphasis id="049D5CC7DA141A2AD52BF95832A9D7FF" italics="true">Cyrtodactylus ngengpuiensis</emphasis>
</taxonomicName>
<emphasis id="B26E01F61E0F757BC5416000AEC91B5F" bold="true">
<taxonomicNameLabel id="015BBA36964F70B1A9A4E606644DFEA5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="BD62B43DBA83B53E4831F583387F9524" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ayeyarwadyensis">
<emphasis id="3AE1CB5E1E0039FBD863F0EC34E1C340" italics="true">C. ayeyarwadyensis</emphasis>
</taxonomicName>
by presence of tiny precloacal pores in female,
<abbrev id="ABBRID0ELQCK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
10–16 (vs.
<abbrev id="ABBRID0EPQCK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
absent in female); differs from
<taxonomicName id="2004AFCCE9BCD2E003BB138AB649E051" authorityName="Kamei &amp; Mahony" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="bapme">
<emphasis id="6127D64EC6A6B05A475D25749EDEC109" italics="true">C. bapme</emphasis>
</taxonomicName>
by presence of 10–16 precloacal pores in female (vs. 0–13 pitted precloacal scales), dorsal tubercle rows,
<abbrev id="ABBRID0E5QCK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
18 or 20 (vs. 21–24); differs from
<taxonomicName id="ECD4F31B3D3639C0B79712995FA48348" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="guwahatiensis">
<emphasis id="36E751EC4D9FB211DC37C65B1A2F030F" italics="true">C. guwahatiensis</emphasis>
</taxonomicName>
by fewer preclocaofemoral pores,
<abbrev id="ABBRID0ENRCK" xlink_title="precloacofemoral pores, a continuous series of pore-bearing scales that extends from the precloacal region onto the thighs">PcFP</abbrev>
27 (vs.
<abbrev id="ABBRID0ERRCK" xlink_title="precloacofemoral pores, a continuous series of pore-bearing scales that extends from the precloacal region onto the thighs">PcFP</abbrev>
35–39),
<abbrev id="ABBRID0EVRCK" xlink_title="precloacofemoral pores, a continuous series of pore-bearing scales that extends from the precloacal region onto the thighs">PcFP</abbrev>
in a continuous series (vs. 26
<abbrev id="ABBRID0EZRCK" xlink_title="precloacofemoral pores, a continuous series of pore-bearing scales that extends from the precloacal region onto the thighs">PcFP</abbrev>
interrupted by 11 unpored scales in
<typeStatus id="154F197CE7D18A7AE92C549A4CCBE819" type="holotype">holotype</typeStatus>
<taxonomicName id="A1D2A1F3F772AFE309DBE821900D7316" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="guwahatiensis">
<emphasis id="692A5248F91AD32676B9ADA9134B543C" italics="true">C. guwahatiensis</emphasis>
</taxonomicName>
),
<abbrev id="ABBRID0EISCK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
present in females (vs.
<abbrev id="ABBRID0EMSCK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
absent in females), higher mid-ventral scale rows,
<abbrev id="ABBRID0EQSCK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="E47B9967422C8BDF2E812FD37B4CB645">MVSR</collectionCode>
</abbrev>
38–39 (
<abbrev id="ABBRID0EUSCK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="2BD8C99971E7E004AC1746A931A1D288">MVSR</collectionCode>
</abbrev>
vs. 30–35), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EYSCK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
18–20 (vs.
<abbrev id="ABBRID0E3SCK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="0CEA1A913B073DDCDDEA102BEFCE5A10">DTR</collectionCode>
</abbrev>
21–24); differs from
<taxonomicName id="0842F1EECE030F1E4DB6507EA631164B" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="karsticola">
<emphasis id="95C85EF6873C9C63124B5B1321378E4E" italics="true">C. karsticola</emphasis>
</taxonomicName>
by fewer precloacal femoral pores in male,
<abbrev id="ABBRID0ELTCK" xlink_title="precloacofemoral pores, a continuous series of pore-bearing scales that extends from the precloacal region onto the thighs">PcFP</abbrev>
27 (vs.
<abbrev id="ABBRID0EPTCK" xlink_title="precloacofemoral pores, a continuous series of pore-bearing scales that extends from the precloacal region onto the thighs">PcFP</abbrev>
34–38), less number of dorsal tubercle rows,
<abbrev id="ABBRID0ETTCK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
18–20 (vs.
<abbrev id="ABBRID0EXTCK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="8A69815D7CCE192AC5D6453A1DCC50B0">DTR</collectionCode>
</abbrev>
21–24), fewer paravertebral tubercles,
<abbrev id="ABBRID0E2TCK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
29–34 (vs.
<abbrev id="ABBRID0E6TCK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="77EDEF4AAE0339F1EF55BDF462EDDF0F">PVT</collectionCode>
</abbrev>
34–39); differs from
<taxonomicName id="D0EF4DD82B7EC57B25D622D0A2C3B6FA" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="tripuraensis">
<emphasis id="1F536E97C0DC425243C939D0C8B395CF" italics="true">C. tripuraensis</emphasis>
</taxonomicName>
by fewer precloacal femoral pores in male,
<abbrev id="ABBRID0EOUCK" xlink_title="precloacofemoral pores, a continuous series of pore-bearing scales that extends from the precloacal region onto the thighs">PcFP</abbrev>
27 (vs.
<abbrev id="ABBRID0ESUCK" xlink_title="precloacofemoral pores, a continuous series of pore-bearing scales that extends from the precloacal region onto the thighs">PcFP</abbrev>
29–37) and less number of precloacal pores,
<abbrev id="ABBRID0EWUCK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
<quantity id="13453BC85AEDD6459DAE8C965313E55C" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="4.064" metricValueMin="2.54" unit="in" value="13.0" valueMax="16.0" valueMin="10.0">10–16 in</quantity>
female (vs.
<abbrev id="ABBRID0E1UCK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
<quantity id="46AA94C7EB19BCD34EBBF929F7FD69C3" metricMagnitude="-1" metricUnit="m" metricValue="6.096" metricValueMax="7.366" metricValueMin="4.826" unit="in" value="24.0" valueMax="29.0" valueMin="19.0">19–29 in</quantity>
female). Morphological differences with other members of
<taxonomicName id="E22F2987EB4952BCAEF30EFE0A4BFC03" baseAuthorityName="Jerdon" baseAuthorityYear="1870" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="EECAEF70563AC3B1CAA27628223DB9BF" italics="true">khasiensis</emphasis>
</taxonomicName>
group is presented in Table
<tableCitation id="6886D9D1178DD98C4E3C7F1B57ABDFCA" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMVCK" type="Sequence divergence">
<paragraph id="269FA1035014FC7DE9EE9C5E99549EF1">
<heading id="819CD88D3A4BCF7D32DC0A39F4009B70" reason="title">Sequence divergence.</heading>
</paragraph>
<paragraph id="97E848580648B450CC1A6A02CDD94636">
<taxonomicName id="8F82CB78DAAF850D280FDA97AFD867D3" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngengpuiensis" status="sp. nov.">
<emphasis id="5C2C3A5CE77474E546D619A332B69D0F" italics="true">Cyrtodactylus ngengpuiensis</emphasis>
</taxonomicName>
<emphasis id="88B7787029928A081A0927B6C4D20CA9" bold="true">
<taxonomicNameLabel id="A421DC0185E325731225C13D6AFEF2BD" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a moderate genetic divergence of 4.1–6.6 % from its closely related
<taxonomicName id="99B7B08DA5D8E4C973606F7832C57B62" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ayeyarwadyensis">
<emphasis id="C3DDB86C9FA7BF2B409A7C460D8C61D6" italics="true">C. ayeyarwadyensis</emphasis>
</taxonomicName>
, 4.2–6.0 % from
<taxonomicName id="5B06D4332125F94004096CE66925E4C7" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="tripuraensis">
<emphasis id="8A7BC45B8F92E1CB4176990C3679F706" italics="true">C. tripuraensis</emphasis>
</taxonomicName>
. With other members of the clade,
<taxonomicName id="ABAE30325E0BB28CC79D3C3B4CEBD451" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngengpuiensis" status="sp. nov.">
<emphasis id="B04A9C338C376960AEC0F0B0DBB2B55D" italics="true">C. ngengpuiensis</emphasis>
</taxonomicName>
<emphasis id="CEF56898FF9ED8BF1E4580CA7B91F9BB" bold="true">
<taxonomicNameLabel id="6876A000F76FCB58B40A75376C69BE20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a genetic divergence between 8.6 % and 15.7 % in the ND 2 gene. The intraspecific divergence is between the two samples of
<taxonomicName id="51A2581721240646E796C45EF5D3CE67" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngengpuiensis" status="sp. nov.">
<emphasis id="0F3339F098920B91986C39D5BAF75F43" italics="true">C. ngengpuiensis</emphasis>
</taxonomicName>
<emphasis id="03C810AA205F5DDFECCA404DE5580180" bold="true">
<taxonomicNameLabel id="84D7FCF3E6977757E0235C6160F780E1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is 0.8 %.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPXCK" type="etymology">
<paragraph id="CA7625AADB35E08BF8DFD36D6ED5AFE9">
<heading id="3EBE8F49203E1937719322D38674448C" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="469226927F875902E1CCFB888B17437C">
The specific epithet is a toponym derived from the name “ Ngengpui Wildlife Sanctuary ” of
<collectingRegion id="BA1398980158B38BB5F47273A9604F98" country="India" name="Mizoram">Mizoram state</collectingRegion>
from where the
<typeStatus id="66C9BF3FF107A35FD1C445E1EFFEB11B">type</typeStatus>
series of the species were collected.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUXCK" type="Suggested common name">
<paragraph id="AF9B56A81204C81B740FAA9FA9682843">
<heading id="2664FC17998A712DB493999BF65A0B93" reason="title">Suggested common name.</heading>
</paragraph>
<paragraph id="EC83F1445768E96861221F49152643E7">Ngengpui bent-toed gecko.</paragraph>
</subSubSection>
<subSubSection id="SECID0EZXCK" type="distribution">
<paragraph id="50654F29F817DA44B31EC759EA6933E4">
<heading id="EEC0AC18ACD9E3BF4F5A2185646CD453" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="D587B71BBC38020762B6B4D1C84C9EA6">
<taxonomicName id="B7BF4225B08E7F72D6A89188202B7529" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngengpuiensis" status="sp. nov.">
<emphasis id="13E89BBE6EEA33CA9703C239D472DCA4" italics="true">Cyrtodactylus ngengpuiensis</emphasis>
</taxonomicName>
<emphasis id="CBFFA2613E31F760F8DA5DB26A850958" bold="true">
<taxonomicNameLabel id="A29B6B0A61186B394694229C7218225D" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
new species is currently only known from the
<typeStatus id="27077758B0BE48388362EC39323B1E03">type</typeStatus>
locality, the Ngengpui Wildlife Sanctuary, Lawngtlai District,
<collectingRegion id="5B9C8A266B21E10253524F2EBC70E953" country="India" name="Mizoram">Mizoram</collectingRegion>
,
<collectingCountry id="6E7C31C7FDABCC656C3D104C1AD2DBA6" name="India">India</collectingCountry>
. The forest is characterised as tropical semi-evergreen to moist evergreen forest. The forest is dominated by
<taxonomicName id="5B518A3A479AFD8F5F3541A0B43C3072" class="Magnoliopsida" family="Dipterocarpaceae" genus="Dipterocarpus" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="F65918F94996C65D837790E7C1E7F3A9" italics="true">Dipterocarpus</emphasis>
spp.
</taxonomicName>
, palms, canes and rattans. Individuals of
<taxonomicName id="4425261B421BFDB040E3957CD059C67E" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngengpuiensis" status="sp. nov.">
<emphasis id="F6F0705F586489D0438FBA0773B75A78" italics="true">C. ngengpuiensis</emphasis>
</taxonomicName>
<emphasis id="D9C3DE5E0E9082650DB3316E5B32124E" bold="true">
<taxonomicNameLabel id="AE059AA90F05A7864924E56BFA530EBC" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected during
<date id="29E0619D53978CFF627C8E3D4BCC3ED4" value="2021-09-06" valueMax="2021-09-09" valueMin="2021-09-06">6–9 September 2021</date>
at 21: 00–22: 00 hrs. Individuals were recorded in bamboo thickets, on tree buttresses and trunks, and amongst ferns and rocks along the banks of evergreen forest streams.
</paragraph>
</subSubSection>
</treatment>
</document>

259
data/2C/C6/1F/2CC61FD327F05CD5990514CCCF1D0F2B.xml Normal file
View file

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<mods:title id="2D045DB69D69935F88BB26C21088C5B9">Four new Dolichopoda species from Greece, one Troglophilus new to Greece and new locality records (Orthoptera, Rhaphidophoridae)</mods:title>
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<mods:namePart id="580700A475998E652A6701B8D38A2AC6">Alexiou, Sotiris</mods:namePart>
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<taxonomicName id="7DB5AF5D1EA605F39A15D2F85B3F46CC" ID-CoL="8PYGK" authority="Rampini &amp; Di Russo, 2008" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="lustriae">
<emphasis id="01283741802BA61776802FF6D7EF3B1A" italics="true">Dolichopoda lustriae</emphasis>
<bibRefCitation id="0907814FCE8FA1AEE2BAF08972B68DA9" DOI="10.11646/zootaxa.1923.1.1" author="Rampini M &amp; Di Russo C &amp; Pavesi F &amp; Cobolli M" journalOrPublisher="Zootaxa" pagination="1-17" refId="B12" refString="Rampini M, Di Russo C, Pavesi F, Cobolli M (2008) The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae). Zootaxa 1923: 1 - 17. https: // doi. org / 10.11646 / zootaxa. 1923.1. 1" title="The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae)." volume="1923" year="2008">Rampini &amp; Di Russo, 2008</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="2070BA7DF23BD745A4A780824706045F">
<figureCitation id="CBC83317072255C6D9ED73EE78B78BE4" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Dolichopoda lustriae Rampini &amp; Di Russo, 2008. Greece, Thessalia, Anthousa, Monastery of Galaktotrophousa, 1 Aug. 2020 (photo Kofinas)." figureDoi="10.3897/contrib.entomol.74.e121614.figure10" httpUri="https://binary.pensoft.net/fig/1103857">Fig. 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EY6AE" type="material">
<paragraph id="6DA8CCCE7E1D6AA05728A0E3E39EAD32">
<heading id="5B637C35E0EF1D75FFC1C57900F03815" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="EE477E29CD2FC27D1417E52E3641E38E">
<materialsCitation id="BE735C920FE583B7551151BB05367A70" collectingDate="2018-12-09" collectorName="S. Alexiou" country="Greece" county="Nomos Karditsas" location="cave Gaki" municipality="Mt. Agrapha" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Thessalia">
<collectingCountry id="0298B51A49AEE7577707FFB6DC351CF1" name="Greece">GREECE</collectingCountry>
–
<emphasis id="D871E755239358143805170134FA64C9" bold="true">
<collectingRegion id="2B7C898BEEED507E762E05A1D8245E5F" country="Greece" name="Thessalia">Thessalia</collectingRegion>
</emphasis>
•
<specimenCount id="60572326B51B55B46A15667187C9788F" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="6074AE5544CE102D9F300030C7B2201C" count="2" type="female">2 ♀</specimenCount>
;
<collectingCounty id="7585D33B1089106B146FF0186FD59A0E">Nomos Karditsas</collectingCounty>
,
<collectingMunicipality id="70678252E5EE58AA794719197807E033">Mt. Agrapha</collectingMunicipality>
,
<location id="5B86654077289C62BBB0351C5206D278" LSID="urn:lsid:plazi:treatment:2CC61FD327F05CD5990514CCCF1D0F2B:5B86654077289C62BBB0351C5206D278" country="Greece" county="Nomos Karditsas" municipality="Mt. Agrapha" name="cave Gaki" stateProvince="Thessalia">cave Gaki</location>
;
<date id="827B7AD8240B790E85EA38AEB29C7138" value="2018-12-09">
<collectingDate id="83FA41D45EF27D114F3D6B6DBC6BC24B" value="2018-12-09">9 Dec. 2018</collectingDate>
</date>
;
<collectorName id="04F694269C0F64DD93408B75DAAFDB7C">S. Alexiou</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0EBAAG" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
•
<materialsCitation id="2B222EBB2A3BECE447DE69B675A4CEA6" collectingDate="2018-10-08" collectorName="L. Makrostergios" country="Greece" location="Agrapha" municipality="Nomos Karditsas" specimenCount="1" specimenCount-female="1" stateProvince="Thessalia">
<specimenCount id="AE9155245018D017B5BFA19541657BBD" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="60E28EAF45EDC5C1D07FDF9D53E6F1D1">Nomos Karditsas</collectingMunicipality>
, Μt.
<location id="14F39E2F3BB913E4C920DE4051582A79" LSID="urn:lsid:plazi:treatment:2CC61FD327F05CD5990514CCCF1D0F2B:14F39E2F3BB913E4C920DE4051582A79" country="Greece" municipality="Nomos Karditsas" name="Agrapha" stateProvince="Thessalia">Agrapha</location>
, refuge ;
<date id="D9B09EE6C2EB1BBA663F05C978C8502B" value="2018-10-08">
<collectingDate id="33E943D6E0337D25C2764146E30629BA" value="2018-10-08">8 Oct. 2018</collectingDate>
</date>
;
<collectorName id="8F5A3A12BA4E093778AA580131DB2835">L. Makrostergios</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0EGAAG" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
•
<materialsCitation id="A3DCF8A12CD1E42796267E63CFD950D3" collectingDate="2020-08-01" collectorName="G. Kofinas &amp; Sterea Ellas" country="Greece" county="Nomos Trikalon" elevation="1150" location="Monastery of Galaktotrophousa" municipality="Anthousa" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Thessalia">
<specimenCount id="A082769239C10183A596194AB7648B03" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="488894D4FD2CE273633B25CA94F818D1" count="3" type="female">3 ♀</specimenCount>
;
<collectingCounty id="8BA4CBA237595608EE583080D82DBE65">Nomos Trikalon</collectingCounty>
,
<collectingMunicipality id="FBC9B8B699ED7C2C537DFA1AF9FF3B01">Anthousa</collectingMunicipality>
,
<location id="93B996AE909C29203D79FF807F29F30C" LSID="urn:lsid:plazi:treatment:2CC61FD327F05CD5990514CCCF1D0F2B:93B996AE909C29203D79FF807F29F30C" country="Greece" county="Nomos Trikalon" municipality="Anthousa" name="Monastery of Galaktotrophousa" stateProvince="Thessalia">Monastery of Galaktotrophousa</location>
;
<date id="0DE297798BE5698681D57A1FE6A3B444" value="2020-08-01">
<collectingDate id="C25512618C0E4DCE883AF442A6E9D6E8" value="2020-08-01">1 Aug. 2020</collectingDate>
</date>
; c.
<elevation id="03A04FF18D6578345C922DCB98BB51CA" metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">
<quantity id="292E1416EBC53B0DD7ED309C6B9EA514" metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">1150 m</quantity>
a. s. l.
</elevation>
;
<collectorName id="E16C07D8C68256BF170C540CEFF0151C">G. Kofinas</collectorName>
leg. (ΚΜΝΗ). -
<emphasis id="507F792D25FAB621C7DC62C6D5ADE05E" bold="true">
<collectorName id="3CC1C162B7FB1DC13BD6182C0E7C9B41">Sterea Ellas</collectorName>
</emphasis>
</materialsCitation>
•
<materialsCitation id="9829D23498D330F363486541DFC9A3A1" collectingDate="2021-08-13" collectorName="K. Bakolitsas" country="Greece" county="Nomos" location="cave Patriarchis" municipality="Neraida" specimenCount="1" specimenCount-female="1" stateProvince="Thessalia">
<specimenCount id="B5CE75D98D24FE446A5B3F9F51FCBBF7" count="1" type="female">1 ♀</specimenCount>
;
<collectingCounty id="D786B79B291FCADC6F2CB60745636798">Nomos</collectingCounty>
Εvritanias,
<collectingMunicipality id="AA6D02FC9F125133CF3C50AC30F5337B">Neraida</collectingMunicipality>
,
<location id="5864E004E6219AECEA653AB006DCA9BA" LSID="urn:lsid:plazi:treatment:2CC61FD327F05CD5990514CCCF1D0F2B:5864E004E6219AECEA653AB006DCA9BA" country="Greece" county="Nomos" municipality="Neraida" name="cave Patriarchis" stateProvince="Thessalia">cave Patriarchis</location>
;
<date id="19EDEC70B3B8B345D95C4B9728C778B4" value="2021-08-13">
<collectingDate id="638E4CC886DF27AABB2E32E3DE3ABD56" value="2021-08-13">13 Aug. 2021</collectingDate>
</date>
;
<collectorName id="C0BE24481B02790A8A19CBA3F745D287">K. Bakolitsas</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0ENAAG" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
•
<materialsCitation id="DA72B2C0880233B60673BFFECB0B28D5" collectingDate="2021-08-31" collectorName="K. Bakolitsas" country="Greece" location="cave Mesokomis" municipality="Nomos" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Thessalia">
<specimenCount id="9D70AB991BF430E26B1799D3D8BAB92A" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="E880BA73D51FA99D174B793D47D1B04F" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="C77F6A003CDD3ED6C216FE50E836AD7D">Nomos</collectingMunicipality>
Εvritanias,
<location id="E49B73B520E73D05CFBB249833ABCFDE" LSID="urn:lsid:plazi:treatment:2CC61FD327F05CD5990514CCCF1D0F2B:E49B73B520E73D05CFBB249833ABCFDE" country="Greece" municipality="Nomos" name="cave Mesokomis" stateProvince="Thessalia">cave Mesokomis</location>
;
<date id="7E08564919A7D3238EF171A48D8A5DBA" value="2021-08-31">
<collectingDate id="2E754BD7A095E36B43344D992ACD948D" value="2021-08-31">31 Aug. 2021</collectingDate>
</date>
; leg.
<collectorName id="2F0B750AD3BEFFC88501567CC5A79A10">K. Bakolitsas</collectorName>
(
<abbrev content-type="institution" id="ABBRID0ESAAG" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
•
<materialsCitation id="62C3EFA4F1DA0FC6F441F2ADB596B264" collectorName="K. Bakolitsas" country="Greece" county="Nomos Etolias-Akarnanias" elevation="854" latitude="38.81925" location="lake Kastrakiou" longLatPrecision="1" longitude="21.421333" municipality="Stoa Acheloou" specimenCount="857" specimenCount-female="2" specimenCount-male="1" stateProvince="Thessalia">
<specimenCount id="5F03618E147E1CF286F2FE65A2560894" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="14D8AE9924E10F895464F423FD167E62" count="2" type="female">2 ♀</specimenCount>
;
<collectingCounty id="F8AC33CC8C79B3A2256CB85984F7D622">Nomos Etolias-Akarnanias</collectingCounty>
,
<collectingMunicipality id="7D5E50A7E7289C667947306950235AEE">Stoa Acheloou</collectingMunicipality>
, close to
<location id="9CF21D2F351AD0056AAD101FE31159AF" LSID="urn:lsid:plazi:treatment:2CC61FD327F05CD5990514CCCF1D0F2B:9CF21D2F351AD0056AAD101FE31159AF" country="Greece" county="Nomos Etolias-Akarnanias" latitude="38.81925" longLatPrecision="1" longitude="21.421333" municipality="Stoa Acheloou" name="lake Kastrakiou" stateProvince="Thessalia">lake Kastrakiou</location>
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a. s. l ;
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leg. (
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.
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<label id="2E3A55F3C10145DDC80F33D4A9BC8B67">Figure 10.</label>
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<paragraph id="E7EBC2AE797E15A3214BEA84CC88328D">
<taxonomicName id="A120F26EE47852171ED3D4D2EE3E41C1" authority="Rampini &amp; Di Russo, 2008" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="lustriae">
<emphasis id="9ADB2FC8A8351CAAC52E2E3D3681B91F" italics="true">Dolichopoda lustriae</emphasis>
<bibRefCitation id="4458354D5D61AFC98AA56C80B73DDEE6" DOI="10.11646/zootaxa.1923.1.1" author="Rampini M &amp; Di Russo C &amp; Pavesi F &amp; Cobolli M" journalOrPublisher="Zootaxa" pagination="1-17" refId="B12" refString="Rampini M, Di Russo C, Pavesi F, Cobolli M (2008) The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae). Zootaxa 1923: 1 - 17. https: // doi. org / 10.11646 / zootaxa. 1923.1. 1" title="The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae)." volume="1923" year="2008">Rampini &amp; Di Russo, 2008</bibRefCitation>
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. Greece, Thessalia, Anthousa, Monastery of Galaktotrophousa, 1 Aug. 2020 (photo Kofinas).
</paragraph>
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<heading id="087C0F890D661E39847C13D3423BA71A" reason="title">Comment.</heading>
</paragraph>
<paragraph id="1659FBD30E4B30B7E62B8E1C26E59DBE">
<taxonomicName id="411D6A738F7F6E2E0BF684C68EEBBFA0" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="lustriae">
<emphasis id="C43249992DCFF24F19DB360F4EA9E48F" italics="true">Dolichopoda lustriae</emphasis>
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was originally described from Μt. Pselovuni of Etolia-Akarnania. Two more populations were added in the following years: Mt. Vardousia Fokidos (
<bibRefCitation id="CA6AAF1FBC11DEAFEFA1D61FF801E1B0" author="Alexiou" etAl="et al." firstAuthor="Alexiou" journalOrPublisher="Parnassiana Archives" pagination="71-81" refId="B2" refString="Alexiou S, Bakolitsas K, Di Russo C, Latella L, Rampini M, Zacharias S (2014) Biospeleological notes on the caves of the Parnassos, Giona and Vardousia Mountains (Central Greece). Parnassiana Archives 2: 71–81." title="Biospeleological notes on the caves of the Parnassos, Giona and Vardousia Mountains (Central Greece)." volume="2" year="2014">Alexiou et al. 2014</bibRefCitation>
) and Pades
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(Davranoglou et al. 2018). The new populations added here demonstrate the wide distribution of the species along the high altitudes of the Pindos mountain range (Fig.
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), which may imply the more epigean biological circle of
<taxonomicName id="57C8B40D5AFB32A2C2CC358204BF6946" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="lustriae">
<emphasis id="190CE03D813D29671A634F9EFF4C5F96" italics="true">D. lustriae</emphasis>
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, as discussed at
<bibRefCitation id="B3871977AC49034D8040EAC4A4F278D0" DOI="10.1080/00222933.2021.1891316" author="Davranoglou L-R &amp; Matsumoto K &amp; Sohc Z &amp; Kempton J" etAl="et al." firstAuthor="Davranoglou" journalOrPublisher="Journal of Natural History" pagination="1-13" refId="B6" refString="Davranoglou L-R, Matsumoto K, Sohc Z, Kempton J (2021) New records and ecological observations on Greek cave crickets (Orthoptera: Rhaphidophoridae). Journal of Natural History 55: 1–13. https://doi.org/10.1080/00222933.2021.1891316" title="New records and ecological observations on Greek cave crickets (Orthoptera: Rhaphidophoridae)." volume="55" year="2021">Davranoglou et al. (2021)</bibRefCitation>
.
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</treatment>
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120
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<mods:title id="B772D5767A529BFD22ED2B1A9304D060">Four new Dolichopoda species from Greece, one Troglophilus new to Greece and new locality records (Orthoptera, Rhaphidophoridae)</mods:title>
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<emphasis id="F07558BECFB9773137925E6151A81EA5" italics="true">Dolichopoda giachinoi</emphasis>
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<paragraph id="AFBBD290C0E17A1AB111C918CAE4840B">
<heading id="BED0CDBA11014AB2E320C909274FF792" reason="title">Material examined.</heading>
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<paragraph id="94C8CC84755D5FE80186AB236BCCF9D3">
<materialsCitation id="B426B99CBE3B8C506BC39C14012C54B0" collectingDate="2016-06-06" collectorName="K. Bakolitsas" country="Greece" county="cave Likonikos" latitude="38.839233" location="Sterea Ellas" longLatPrecision="1" longitude="20.92775" municipality="Vonitsa" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
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•
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<geoCoordinate id="F58B192A523C8951FF0BA5304D3561F3" degrees="38" direction="north" minutes="50.354" orientation="latitude" precision="1" value="38.839233">38 ° 50.354 ' N</geoCoordinate>
,
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;
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<collectingDate id="F6B6BC3762EF0177358302063FB966A9" value="2016-06-06">6 Jun. 2016</collectingDate>
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leg.
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The species has been described recently from Megalo Spilio cave, at the northern side of Mt. Serekas (part of Mt. Akarnanika) and from an altitude of
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. a. s. l. (
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). The new locality extends the distribution of the species to the north, close to the Ionian Sea and away from the mountainous area of Akarnanika.
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<emphasis id="82715410094675577E6A2E53AF2B178C" italics="true">Dolichopoda vandeli</emphasis>
Boundou-Saltet, 1970
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<heading id="FF8F8CA43B0B84223DB29F595CE7814B" reason="title">Material examined.</heading>
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•
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,
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, 2 nymphs ;
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leg. (
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.
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<heading id="BC2A6A1AE823DC1C9353DD53E280E79C" reason="title">Comment.</heading>
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<paragraph id="8CC570782CBDD74D68863D544556FB98">
<taxonomicName id="B207A428040C21124E1A2908A2060270" authorityName="Boundou-Saltet" authorityYear="1970" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="vandeli">
<emphasis id="BF295D79BC106523A067833E926449E9" italics="true">Dolichopoda vandeli</emphasis>
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was described from Hermes cave, Nomos Biotias, Orchomenos, near the village of Dionysos, at the north of the plane of Kopais. Subsequently it has been collected from two more caves of Kopais plane, near Akontio and Katavothra Aliartos, in addition to Mt. Elikon, Agia Triada cave and the north foothills of Mt. Parnassos (Ano Polydroso, cave Kontilospilia) (
<bibRefCitation id="A4FB5FF1B93243FC1F61042FBACBF556" author="Alexiou" etAl="et al." firstAuthor="Alexiou" journalOrPublisher="Parnassiana Archives" pagination="71-81" refId="B2" refString="Alexiou S, Bakolitsas K, Di Russo C, Latella L, Rampini M, Zacharias S (2014) Biospeleological notes on the caves of the Parnassos, Giona and Vardousia Mountains (Central Greece). Parnassiana Archives 2: 71–81." title="Biospeleological notes on the caves of the Parnassos, Giona and Vardousia Mountains (Central Greece)." volume="2" year="2014">Alexiou et al. 2014</bibRefCitation>
;
<bibRefCitation id="D4B556013F7A3D1C4E87FD7267CA1BC5" author="Di Russo" etAl="et al." firstAuthor="Di Russo" journalOrPublisher="Biodiversity Journal" pagination="397-420" refId="B7" refString="Di Russo C, Rampini M, Cobolli M (2014) The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss, 1879. Biodiversity Journal 5: 397–420." title="The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss, 1879." volume="5" year="2014">Di Russo et al. 2014</bibRefCitation>
). Our record is a significant expansion of the species range to the south (Fig.
<figureCitation id="1A833D41CF56E801B70D59EE988978EA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Geographic distribution of D. lustriae Rampini &amp; Di Russo, 2008 and D. vandeli Boundou-Saltet, 1970." figureDoi="10.3897/contrib.entomol.74.e121614.figure9" httpUri="https://binary.pensoft.net/fig/1103856">9</figureCitation>
).
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<caption id="64BA837F77F1D2EAB23C87915EFE8339" ID-DOI="10.3897/contrib.entomol.74.e121614.figure9" ID-arpha="3D3D9579-2082-5AE9-AA8D-262D5278A0E8" httpUri="https://binary.pensoft.net/fig/1103856" startId="F9">
<paragraph id="915447DE59A77E823B0E9926462A2C4F">
<label id="4E7639035A1F6ACB2068826A9F1E2E93">Figure 9.</label>
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<paragraph id="CC863614563147713E5A056431A8D102">
Geographic distribution of
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<emphasis id="0E2C9D052FF95C7E08DAF222DE88562F" italics="true">D. lustriae</emphasis>
<bibRefCitation id="77CDD977025E3A642DBC1AE1614BCAEF" DOI="10.11646/zootaxa.1923.1.1" author="Rampini M &amp; Di Russo C &amp; Pavesi F &amp; Cobolli M" journalOrPublisher="Zootaxa" pagination="1-17" refId="B12" refString="Rampini M, Di Russo C, Pavesi F, Cobolli M (2008) The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae). Zootaxa 1923: 1 - 17. https: // doi. org / 10.11646 / zootaxa. 1923.1. 1" title="The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae)." volume="1923" year="2008">Rampini &amp; Di Russo, 2008</bibRefCitation>
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and
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<emphasis id="91B6B7AE4CCF67E80D5D129B838B6BA5" italics="true">D. vandeli</emphasis>
Boundou-Saltet, 1970
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.
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,
<geoCoordinate id="3C2DE4978D7CC0E4F401A2EE10E3E316" degrees="65" direction="west" minutes="17" orientation="longitude" precision="15" seconds="22" value="-65.289444">65 ° 17 ′ 22 ″ W</geoCoordinate>
</named-content>
;
<date id="481307E7FF511FF1978B894D3D2FE054" value="2003-03-19">
<collectingDate id="42BE3A9999390E35513E86ADB9E89F25" value="2003-03-19">19. iii. 2003</collectingDate>
</date>
;
<collectorName id="A56061F7B3EAF8314072867F183F771E">J. Heraty. Will</collectorName>
be housed in
<collectionCode id="65E5D98AB1E5225E51AA819F2553EF8E" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" type="University or college">UCRC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3MAC" type="description">
<paragraph id="97ED734BDDBEF62FBEE80D0C20DF5E92">
<heading id="EE8E94BE8638B25202589F15A0D2F0C3" reason="title">Description.</heading>
</paragraph>
<paragraph id="70EB9C71B16EF4A05DC8543618469A61">
<emphasis id="9BCBB890B11AC7F67AAA578914411DEF" bold="true" italics="true">Body length</emphasis>
:
<quantity id="BAAD5B36D0607D163A5D5E20BA9FD367" metricMagnitude="-3" metricUnit="m" metricValue="3.7" unit="mm" value="3.7">3.7 mm</quantity>
.
<emphasis id="04E11861B0965CC2A87592E1C47CEAC4" bold="true" italics="true">Length of forewing</emphasis>
:
<quantity id="12162B96FB42B3920FF864AE49C1AAD4" metricMagnitude="-3" metricUnit="m" metricValue="3.3" unit="mm" value="3.3">3.3 mm</quantity>
.
<emphasis id="52A0F1FD4DC7777B2F98638B778A395A" bold="true" italics="true">Length of hind wing</emphasis>
:
<quantity id="0477B58F45FC4681D46B4923C0E140B7" metricMagnitude="-3" metricUnit="m" metricValue="2.6" unit="mm" value="2.6">2.6 mm</quantity>
.
<emphasis id="2AD32D9D4D50FF5278FCBA33D0C424CC" bold="true">
<emphasis id="534D8537ACE8FAEBFB91F5C2782FC373" bold="true" italics="true">Head</emphasis>
.
</emphasis>
Antenna relatively thick with 24 flagellomeres; first flagellomere 1.5 × as long as second. Median width of eye 0.3 × longer than median width of gena in lateral view (0.3: 0.1). Width of anterior ocellus 0.9 × longer than distance between posterior ocelli (0.08: 0.09). Apex of clypeus convex with a smooth apical margin. Maxillary palpus 5 - segmented; labial palpus 4 - segmented. Occipital carina well developed dorsally, absent ventrally (This may be a pseudo-occipital carina, i. e., secondarily derived, as is found in some Agathidinae, e. g.,
<taxonomicName id="3E30DABB366DEEC1363AE90F8108F8D4" class="Insecta" family="Braconidae" genus="Marjoriella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="FC1938088C55427E3941C2367B3B2004" italics="true">Marjoriella</emphasis>
spp.
</taxonomicName>
). Most of head with weak microsculpture contrasting sharply with the smooth, glabrous occiput. Malar suture present. Interantennal space with a bicarinate ridge. Median ocellus surrounded medially and laterally by a smooth, curved ridge.
<emphasis id="44C7B62A3DAC7EFF080E49C547764DE8" bold="true">
<emphasis id="A2FB02012086603999970D180124906D" bold="true" italics="true">Mesosoma</emphasis>
.
</emphasis>
Pronotum bilobed anteriorly with a transverse plate dorsally. Notauli deeply impressed and entirely costate. Median lobe of mesoscutum bilobed. Scutellar sulcus deep with a median carina; median width of scutellar sulcus
<quantity id="A6A339E301F1423E0D8D85E7DFCA7666" metricMagnitude="-4" metricUnit="m" metricValue="4.0" unit="mm" value="0.4">0.4 mm</quantity>
; median length of scutellar sulcus
<quantity id="1C3CB3A7A8A3AC766194251ED821C19E" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
; median length of scutellar sulcus 0.1 × longer than median length of mesosoma in dorsal view (0.1: 0.9). Scutellum smooth and flat. Postscutellar depression absent. Propleuron lacking a posterolateral lobe. Epicnemial carina strong and complete. Precoxal sulcus well defined with ~ 5 costulae. Propodeum rugose with a large, well-defined median areola.
<emphasis id="EDF0B759881C624EF2FB29AF575BC422" bold="true">
<emphasis id="3601D961ACA154EFA310B78D2F9A0DA1" bold="true" italics="true">Wings</emphasis>
.
</emphasis>
Forewing M + Cu entirely tubular; 1 RS vein long; second submarginal cell large and greatly compressed apically, trapezoid, maximum length of the cell 1.6 × longer than its maximum height (0.46: 0.28); apical abscissa of RS entirely nebulous and almost straight, very slightly curved posteriorly; (RS + M) b about 3 × longer than m-cu vein; 1 M about 3 × longer than m-cu vein; anal crossvein indicated by a slight swelling on vein A. Hind wing unremarkable; lacking distinct claval lobe; veins M + Cu and M about equal in length; r crossvein absent.
<emphasis id="56A7E4FB049549B4D0C813663E378E72" bold="true">
<emphasis id="FA7C57C979BDEF69CEEAD348BB9C326F" bold="true" italics="true">Legs</emphasis>
.
</emphasis>
Midtibia 3.4 × longer than midbasitarsomere (0.64: 0.19). Hind femur 0.9 × longer than hind tibia (0.75: 0.84). Hind basitarsomere swollen. Tarsal claws rather large with pectinate base.
<emphasis id="B888EE9AA3F66F5C69E68E724E0DF096" bold="true">
<emphasis id="E1F85E9AB46550E6938E8EC6926B3CD5" bold="true" italics="true">Metasoma</emphasis>
.
</emphasis>
Metasoma 1.1 × longer than mesosoma (1.70: 1.56). Spiracle of
<collectionCode id="8E824967E7DFD2AA42F0FE201D7F5BFE" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
1 on membranous laterotergite.
<collectionCode id="C418333B0A0A092F313DE96B8DCE44F6" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
1 0.7 × longer than its apical width (0.63: 0.45), with carinate lateral margins; median area of
<collectionCode id="716F6174380FDE7D372152E5EE292CB3" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
1 with an inverse Y-shaped depression. Remaining terga smooth and rather weakly sclerotized.
<collectionCode id="90616A9BEC675023EE040B8060DA05E1" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
2 transverse, much wider than long. Hypopygium acute apically and not nearly reaching apex of metasoma. Ovipositor sheath about half as long as metasoma, strongly compressed laterally, with fine sparse setae. Ovipositor simple, slightly downcurved but otherwise unmodified.
<emphasis id="7FEF2CFC0DF3BD27D9407EBEBF6936C2" bold="true">
<emphasis id="0D04860856245C994840606E28F24DE0" bold="true" italics="true">Color</emphasis>
.
</emphasis>
Head and mesosoma mostly light brown; antenna brown, foreleg and midleg entirely pale, hind tibia and tarsus yellow medially, hind claw brown. Metasoma mostly pale except ovipositor sheath, ovipositor sheath light brown basally, apically dark brown. Wings entirely infuscate.
</paragraph>
<paragraph id="32EAE59F4CFD89EDA5A81C95EEB14E35">
<emphasis id="87C82585ACA6F595E7087FB560CCCF3E" bold="true">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVOAC" type="biology">
<paragraph id="0ABF06679ACE584BC332A4C62BD90533">
<heading id="6B85B07E0A4C5BE135FCA8CD29E979C3" reason="title">Biology.</heading>
</paragraph>
<paragraph id="EB9232BDEC4704AC5548C569D6C34370">Unknown.</paragraph>
</subSubSection>
<subSubSection id="SECID0E1OAC" type="distribution">
<paragraph id="9BF1E7C86B03CE583AB9D175035D1DBD">
<heading id="8B131A966906F8041E40D44F53946BA5" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="BD4F2E0502DB6B76ED801A7A1061C90F">
Neotropics.
<taxonomicName id="DE3101A332684F6B189AFE025234C77D" authority="Kang &amp; Sharkey" authorityName="Kang &amp; Sharkey" authorityYear="2024" class="Insecta" family="Braconidae" genus="Dickyyuella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="argentinensis" status="sp. nov.">
<emphasis id="9DCF1182A0279586DFAD4E61F36D07EF" italics="true">Dickyyuella argentinensis</emphasis>
Kang &amp; Sharkey
</taxonomicName>
,
<taxonomicNameLabel id="EEF585F2BDDA9F066714A2186A2E7B4F" rank="species">sp. nov.</taxonomicNameLabel>
, is known from Tapia,
<collectingRegion id="F0E6F28A76A74BD34B59B878B587E58C" country="Argentina" name="Tucuman">Tucumán</collectingRegion>
Pr.,
<collectingCountry id="81AA9692A0154BBA40B65ACC3BF19F71" name="Argentina">Argentina</collectingCountry>
, near Rio
<collectingCountry id="67B39FA0A049B03B07656B9553373EB1" name="India">India</collectingCountry>
Muerta.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELPAC" type="etymology">
<paragraph id="65B214B6602EABF743B53BC6A7F0AD3F">
<heading id="7F39BC168351829A9C0C1EBC81566E59" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="0BA9DD705C8ED807646163299C233E2C">
The species is named after the collecting country, “
<collectingCountry id="D67FA39973FBF15A18025374A9FEBFFD" name="Argentina">Argentina</collectingCountry>
”.
</paragraph>
</subSubSection>
</treatment>
</document>

410
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<treatment id="507D9749BFBD54018DADAD5538DE287C" ID-DOI="http://doi.org/10.5281/zenodo.13124039" ID-Zenodo-Dep="13124039" ID-arpha="507D9749-BFBD-5401-8DAD-AD5538DE287C" LSID="urn:lsid:plazi:treatment:507D9749BFBD54018DADAD5538DE287C" httpUri="http://treatment.plazi.org/id/507D9749BFBD54018DADAD5538DE287C">
<subSubSection id="C8A7230CE852ED7D1E95ED8BBE46E9C7" type="nomenclature">
<paragraph id="72F3CB5D9BCE20CAB94346B42BE7087F">
<heading id="8A42322A9212B8C13A4A21A2C9870F1D" reason="label">Expanded description of</heading>
<taxonomicName id="C9CE8533B709300C820FD4517F01AB40" ID-CoL="8H73R" authority="Li, 2007" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="926891F2589AF1637DEA5CD2730D25FB" italics="true">Cyrtodactylus cayuensis</emphasis>
<bibRefCitation id="2D910A302259869858BEE2C3DD5603BA" author="Li PP" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="733-737" refId="B22" refString="Li PP (2007) Description of a new subspecies of Cyrtodactylus khasiensis from China. Acta Zootaxonomica Sinica 32: 733 - 737." title="Description of a new subspecies of Cyrtodactylus khasiensis from China." volume="32" year="2007">Li, 2007</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="BA6A2F25E6CB04563CFA677285AD63BF" type="description">
<paragraph id="0FC529ADE614032C5367AD68577BCF25">
<figureCitation id="A5F206DB3B5C978AD34809A3BB205C9C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyrtodactylus cayuensis from India showing the variation in dorsal marking pattern. A WII-ADR 1219 (Balek), B WII-ADR 1681 (Ezengo), C WII-ADR 1199 (Jengging), D WII-ADR 1218 (Balek), E WII-ADR 454 (Jengging), F WII-ADR 1213 (Ramsing), G WII-ADR 698 (Potin), H WII-ADR 3017 (Glaw lake), I precloacal pores in WII-ADR 698, and J precloacal pores in WII-ADR 478. Scale bar: 10 mm. I, J not to scale." figureDoi="10.3897/vz.74.e124752.figure10" httpUri="https://binary.pensoft.net/fig/1102276">Figures 10</figureCitation>
,
<tableCitation id="6D24A2E86AE0C2BB86FADA220227FB43" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">S 3; Tables 2, S 4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMZDK" type="Chresonyms">
<paragraph id="CF2D1ACD8A411EB2D190E43B9602410A">
<heading id="2A92FE860E2C1172119951CA0C4B6445" reason="title">Chresonyms.</heading>
</paragraph>
<paragraph id="CA47BE8FC6EF9515DD067EF8372714AD">
<taxonomicName id="B310044B4BA164D8C5AE41C35E2E3FD5" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="subSpecies" species="khasiensis" subSpecies="cayuensis">
<emphasis id="72092101DE52188070AE7E9ADAB9905B" italics="true">Cyrtodactylus khasiensis cayuensis</emphasis>
</taxonomicName>
–
<bibRefCitation id="249F5CC457CAD413AE0E30ED18C144AA" author="Li PP" firstAuthor="Li" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="733-737" refId="B22" refString="Li PP (2007) Description of a new subspecies of Cyrtodactylus khasiensis from China. Acta Zootaxonomica Sinica 32: 733–737." title="Description of a new subspecies of Cyrtodactylus khasiensis from China." volume="32" year="2007">Li (2007)</bibRefCitation>
</paragraph>
<paragraph id="20CA001A5EAAB3CABCFDAF5B722A59E6">
<taxonomicName id="A3876E8EFFDDAB4EECD9EEC55EEC7309" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="1D4BC9D4D69FF84E72D38A5FEF5BFB16" italics="true">Cyrtodactylus cayuensis</emphasis>
</taxonomicName>
– Agarwal et al. (2018: 337)
</paragraph>
<paragraph id="201EED750B227F91DCED5A733F074AC2">
<taxonomicName id="C0C0A5762A17572546C6FC46359DCCCA" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="arunachalensis">
<emphasis id="26715360AD3BE9CD4CF4792D5E208CD4" italics="true">Cyrtodactylus arunachalensis</emphasis>
</taxonomicName>
–
<bibRefCitation id="5036E5447E8BA7FE0583FF745F171D60" DOI="10.3897/evolsyst.5.61667" author="Mirza ZA &amp; Bhosale H &amp; Ansari F &amp; Phansalkar P &amp; Sawant M &amp; Gowande G &amp; Patel H" etAl="et al." firstAuthor="Mirza" journalOrPublisher="Evolutionary Systematics" pagination="13-23" refId="B28" refString="Mirza ZA, Bhosale H, Ansari F, Phansalkar P, Sawant M, Gowande G, Patel H (2021) A new species of geckos of the genus Cyrtodactylus Gray, 1827 from Arunachal Pradesh, India. Evolutionary Systematics 5: 13–23. https://doi.org/10.3897/evolsyst.5.61667" title="A new species of geckos of the genus Cyrtodactylus Gray, 1827 from Arunachal Pradesh, India." volume="5" year="2021">Mirza et al. (2021)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EC2DK" type="material">
<paragraph id="D8BCC3A27E3FFBCF668E47B2EFD0CFE1">
<heading id="4C3D6B323DC39CB2A574707C23F46EC7" reason="title">Materials examined.</heading>
</paragraph>
<paragraph id="778113877C026504E58CADD12B84CDDE">
(Fig.
<figureCitation id="E6C15AE00D7800CAE646D20BFBA5DCE9" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyrtodactylus cayuensis from India showing the variation in dorsal marking pattern. A WII-ADR 1219 (Balek), B WII-ADR 1681 (Ezengo), C WII-ADR 1199 (Jengging), D WII-ADR 1218 (Balek), E WII-ADR 454 (Jengging), F WII-ADR 1213 (Ramsing), G WII-ADR 698 (Potin), H WII-ADR 3017 (Glaw lake), I precloacal pores in WII-ADR 698, and J precloacal pores in WII-ADR 478. Scale bar: 10 mm. I, J not to scale." figureDoi="10.3897/vz.74.e124752.figure10" httpUri="https://binary.pensoft.net/fig/1102276">10</figureCitation>
).
<collectingRegion id="99A66E4C22AA6BF8E1F495EBA10EFD16" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
:
<specimenCount id="207B96D2CFF269066E33A683CAAD9077" count="1" type="male">one male</specimenCount>
(WII-ADR 1219) and
<specimenCount id="0A778DBEE8C1EFB27BFF071EC46F7DBA" count="1" type="female">one female</specimenCount>
(WII-ADR 1218) collected from Balek village (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP2DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.272100,28.062400]}">
<geoCoordinate id="75F2584ACA713396B799F29F9F69F5DB" degrees="28.0624" direction="north" orientation="latitude" precision="5" value="28.0624">28.0624 ° N</geoCoordinate>
;
<geoCoordinate id="BCA63C1A6B619CF6C8232BCA11A09999" degrees="95.2721" direction="east" orientation="longitude" precision="5" value="95.2721">95.2721 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="C59C3841943AC9D1DFF398EB93A9851E" metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">450 m</quantity>
a. s. l.), East Siang District on
<date id="31BBBA95F5569F63A1A04F48C439847A" value="2021-10-29">29 October 2021</date>
by Bitupan Boruah;
<specimenCount id="29BB93CF6B2F8B184B29200C53119BAC" count="1" type="male">one male</specimenCount>
(WII-ADR 1213) collected from Ramsing (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX2DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[94.979500,28.656300]}">
<geoCoordinate id="F8897D5244653885B97E9D1D28ECD03E" degrees="28.6563" direction="north" orientation="latitude" precision="5" value="28.6563">28.6563 ° N</geoCoordinate>
;
<geoCoordinate id="04D1B3D647C89A53608C01DB4B714064" degrees="94.9795" direction="east" orientation="longitude" precision="5" value="94.9795">94.9795 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="E2579E4E2FCD3554D27DA29809C53FF5" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
a. s. l.), Mouling National Park, Upper Siang District on
<date id="E92BB40106586A5FFB897F30283C97E7" value="2021-10-27">27 October 2021</date>
by Bitupan Boruah;
<specimenCount id="56B7E3F556DB4553B1339FE72622FB02" count="1" type="female">one female</specimenCount>
(WII-ADR 1199) collected from Syrnyup stream (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E62DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.030500,28.534000]}">
<geoCoordinate id="BE2A74361CE825ABA05338B581420200" degrees="28.5340" direction="north" orientation="latitude" precision="5" value="28.534">28.5340 ° N</geoCoordinate>
;
<geoCoordinate id="73979654104F37809B98C863027B4DCC" degrees="95.0305" direction="east" orientation="longitude" precision="5" value="95.0305">95.0305 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="329965E1309EFA79A6BE2D2C4F30306D" metricMagnitude="2" metricUnit="m" metricValue="8.9" unit="m" value="890.0">890 m</quantity>
a. s. l.), Jengging, Mouling National Park, Upper Siang District on
<date id="74002D5F7A7C46E2989AE8955CB16213" value="2021-10-26">26 October 2021</date>
by Bitupan Boruah;
<specimenCount id="EAB5A5201F674947B54FDCDEF0964A98" count="2" type="male">two males</specimenCount>
(WII-ADR 695 and WII-ADR 696) and
<specimenCount id="FB615ABC298C53CF136A47668ADFB78A" count="2" type="female">two females</specimenCount>
(WII-ADR 697 and WII-ADR 698) collected from Potin (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH3DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[93.849700,27.347800]}">
<geoCoordinate id="1BCAEB6108EB205809C1C1F14E4A1871" degrees="27.3478" direction="north" orientation="latitude" precision="5" value="27.3478">27.3478 ° N</geoCoordinate>
,
<geoCoordinate id="6104781A052483D83FE2646A0385D20F" degrees="93.8497" direction="east" orientation="longitude" precision="5" value="93.8497">93.8497 ° E</geoCoordinate>
</named-content>
, elevation
<quantity id="EDCC4D11652435D48665772D276076BE" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</quantity>
a. s. l.), Lower Subansiri District on
<date id="3DB9D5066C035EB365FCE3DDDAD0EF60" value="2019-10-05">5 October 2019</date>
by Bitupan Boruah;
<specimenCount id="CB5AF0605439ECC8A81E9883A036C211" count="1" type="female">one female</specimenCount>
(WII-ADR 3017) collected near Glaw lake (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP3DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.445600,27.696000]}">
<geoCoordinate id="F5115D947AFC324DBEF425E7401A6282" degrees="27.6960" direction="north" orientation="latitude" precision="5" value="27.696">27.6960 ° N</geoCoordinate>
;
<geoCoordinate id="9A314D5B0239E88C73437EAE8BC461A0" degrees="96.4456" direction="east" orientation="longitude" precision="5" value="96.4456">96.4456 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="B883FAB4D2C42BEFB836CC65304E01F5" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
a. s. l.), Kamlang Tiger Reserve, Lohit District on
<date id="0DCBC3E5C73A607C47E838073B1A0244" value="2022-09-02">2 September 2022</date>
by Abhijit Das, Bitupan Boruah and Naitik G. Patel;
<specimenCount id="5983550108D8B5E41FDE6B9164BB818A" count="1" type="male">one male</specimenCount>
(WII-ADR 1682) and
<specimenCount id="1AB4A844CCC6CBF52C5565AD354F2AEE" count="1" type="female">one female</specimenCount>
(WII-ADR 1681) collected from Ezengo (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX3DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.863800,28.156500]}">
<geoCoordinate id="F70328F07D5627C626CF9F702A505183" degrees="28.1565" direction="north" orientation="latitude" precision="5" value="28.1565">28.1565 ° N</geoCoordinate>
;
<geoCoordinate id="D48505D8A2B7956AAB23489B3B2D144F" degrees="95.8638" direction="east" orientation="longitude" precision="5" value="95.8638">95.8638 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="E6094C6516B8B1883D3E9FD0551BCC67" metricMagnitude="2" metricUnit="m" metricValue="5.6" unit="m" value="560.0">560 m</quantity>
a. s. l.), Lower Dibang Valley District on
<date id="7D91172C428B060DC773E804AB60CC11" value="2022-08-03">3 August 2022</date>
By Bitupan Boruah;
<specimenCount id="36E563588AA5A6D34FEA761A5BE7CC3A" count="3" type="female">three females</specimenCount>
(WII-ADR 453, WII-ADR 454 and WII-ADR 459) collected from Jengging (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E63DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.053700,28.549900]}">
<geoCoordinate id="98E6CAB88A3E1B58DEE3628294964A8B" degrees="28.5499" direction="north" orientation="latitude" precision="5" value="28.5499">28.5499 ° N</geoCoordinate>
;
<geoCoordinate id="7D9F321ECC03AE9DDE319284FD073E46" degrees="95.0537" direction="east" orientation="longitude" precision="5" value="95.0537">95.0537 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="C4F79E803110B8E81A1353CFCA55D0E8" metricMagnitude="2" metricUnit="m" metricValue="7.6" unit="m" value="760.0">760 m</quantity>
a. s. l.) by Abhijit Das on
<date id="3B5BC1D537033D4E28C6066DB9C3DBB1" value="2018-10-05">5 October 2018</date>
;
<specimenCount id="A35CB19E2045DBDF80E2832C93610F27" count="1" type="female">one female</specimenCount>
(WII-ADR 478) and
<specimenCount id="BDF452773A67C8CCEE1F3E25336696B3" count="1" type="male">one male</specimenCount>
(WII-ADR 473) collected from
<quantity id="515323FACB6D73565D972F6E03CEE122" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
northwest to Pasighat (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH4DK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[95.268200,28.094500]}">
<geoCoordinate id="7E5A365D295C37F2863527C9CA23DA69" degrees="28.0945" direction="north" orientation="latitude" precision="5" value="28.0945">28.0945 ° N</geoCoordinate>
;
<geoCoordinate id="28D82508028FD3D230C39435D8479B15" degrees="95.2682" direction="east" orientation="longitude" precision="5" value="95.2682">95.2682 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="5B9B9B3D86A396CEDD0E49D29616382A" metricMagnitude="2" metricUnit="m" metricValue="4.1" unit="m" value="410.0">410 m</quantity>
a. s. l.) by Abhijit Das on
<date id="EE44B8C8746D60186591CDB47D6ED123" value="2018-10-02">2 October 2018</date>
.
</paragraph>
<caption id="9A6CE0EB7C8B2849EA5740F64A78FD75" ID-DOI="10.3897/vz.74.e124752.figure10" ID-arpha="85FD849D-6749-530E-B888-2043F84A1248" httpUri="https://binary.pensoft.net/fig/1102276" startId="F10">
<paragraph id="1305AA44A36028E2F2F090602D725D3F">
<label id="549BEF6610A56DC097A34246C80A069A">Figure 10.</label>
</paragraph>
<paragraph id="BB98C6501DFA8FAE3B2EFCBD1E774B18">
<taxonomicName id="B9E732165BD3395A8530084AEEB6EC99" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="2D91098550B316A036CAAE3C1662303C" italics="true">Cyrtodactylus cayuensis</emphasis>
</taxonomicName>
from India showing the variation in dorsal marking pattern.
<emphasis id="9DB13E9CF5B9CE758EB75D173582D238" bold="true">A</emphasis>
WII-ADR 1219 (Balek),
<emphasis id="AC25C0262069E488206AEC361CCC7F6A" bold="true">B</emphasis>
WII-ADR 1681 (Ezengo),
<emphasis id="1735B8C9054955D98F61025001FF9814" bold="true">C</emphasis>
WII-ADR 1199 (Jengging),
<emphasis id="189317A6F9AA7CE8373D7D7E385DCF63" bold="true">D</emphasis>
WII-ADR 1218 (Balek),
<emphasis id="FA0BC65F5AC74B70D4DFC503D9016530" bold="true">E</emphasis>
WII-ADR 454 (Jengging),
<emphasis id="27E3EC9C73D1F0D9A00B6F220C179E30" bold="true">F</emphasis>
WII-ADR 1213 (Ramsing),
<emphasis id="12B99593C367D72F5554E0912229323F" bold="true">G</emphasis>
WII-ADR 698 (Potin),
<emphasis id="E253066F32F753AED84417B3A86F6CA7" bold="true">H</emphasis>
WII-ADR 3017 (Glaw lake),
<emphasis id="F067B8AE1E672EEFCE3D7F33FFECF438" bold="true">I</emphasis>
precloacal pores in WII-ADR 698, and
<emphasis id="ED247180329B79ECB8B428552FEDF8C5" bold="true">J</emphasis>
precloacal pores in WII-ADR 478. Scale bar: 10 mm.
<emphasis id="810B37CBE3573B27FA9688CB82640A3E" bold="true">I</emphasis>
,
<emphasis id="1BD501DA45BE36B39D9EAC222F25D318" bold="true">J</emphasis>
not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E35DK" type="Revised diagnosis">
<paragraph id="279F2D38D7AB03B5EBFD59230388F4A0">
<heading id="F38363A7982DA57E4FB53DF577CA96E2" reason="title">Revised diagnosis.</heading>
</paragraph>
<paragraph id="14EAAF97811352C282538C37C27E9989">
Medium-sized gecko,
<abbrev id="ABBRID0EC6DK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="5429E424F9D710D30115D3E0C71974C0" metricMagnitude="-2" metricUnit="m" metricValue="7.234999999999999" metricValueMax="8.35" metricValueMin="6.12" unit="mm" value="72.35" valueMax="83.5" valueMin="61.2">61.2–83.5 mm</quantity>
in adult males and
<abbrev id="ABBRID0EG6DK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="4A83F868424A4C4599230829F2B38DE2" metricMagnitude="-2" metricUnit="m" metricValue="7.15" metricValueMax="8.36" metricValueMin="5.9399999999999995" unit="mm" value="71.5" valueMax="83.6" valueMin="59.4">59.4–83.6 mm</quantity>
in adult females; 8–13 supralabials and 8–12 infralabials; 18–26 rows of bluntly conical and feebly keeled enlarged tubercles across mid-dorsum; 27–38 paravertebral tubercles between the level of axilla and groin; 28–44 mid-ventral scales; 13–21 subdigital lamellae beneath the fourth digit of manus; 11–23 subdigital lamellae beneath the fourth digit of pes; colouration and marking pattern variable; dorsally greyish-brown, pale-brown or pale-yellowish-brown; five to eight dark-brown bands consisting of irregular shaped and sized enlarged spots on back between axilla and groin, or dorsum with dark-brown reticulation; tail with 9–13 dark and 9–12 light bands alternatively arranged on top.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK6DK" type="material">
<paragraph id="1EB40EEF3C92BD6E2077AA2E5DF2D727">
<heading id="C6284181699E7E95ABE33566BB9C0E01" reason="title">Description based on newly collected materials.</heading>
</paragraph>
<paragraph id="8E7C302500064FB15E38599C6CEC4A2E">
Medium-sized gecko,
<abbrev id="ABBRID0EQ6DK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="D2440B1CEACA9C72A8034E5323493F82" metricMagnitude="-2" metricUnit="m" metricValue="6.815" metricValueMax="7.51" metricValueMin="6.12" unit="mm" value="68.15" valueMax="75.1" valueMin="61.2">61.2–75.1 mm</quantity>
in males (n = 6) and
<abbrev id="ABBRID0EU6DK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="02518CD647FD6FC9B19551A0E35D8F94" metricMagnitude="-2" metricUnit="m" metricValue="7.15" metricValueMax="8.36" metricValueMin="5.9399999999999995" unit="mm" value="71.5" valueMax="83.6" valueMin="59.4">59.4–83.6 mm</quantity>
in females (n = 11); head moderately large (
<abbrev id="ABBRID0EY6DK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
/
<abbrev id="ABBRID0E36DK" xlink_title="snout to vent length">SVL</abbrev>
= 0.25–0.28), dorsoventrally depressed, longer than width (
<abbrev id="ABBRID0EAAEK" xlink_title="maximum head width">HW</abbrev>
/
<abbrev id="ABBRID0EEAEK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.67–0.75), distinct from neck, broader at occipital region; snout rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat, a longitudinal furrow on dorsal surface of the snout, snout short (
<abbrev id="ABBRID0EIAEK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
/
<abbrev id="ABBRID0EMAEK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.38–0.43), longer than orbit (
<abbrev id="ABBRID0EQAEK" xlink_title="orbit diameter, taken between the anterior and posterior bony orbital borders">OD</abbrev>
/
<abbrev id="ABBRID0EUAEK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
= 0.49–0.78); nostril opening directed posterolaterally; ear opening oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous; supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases; rostral wide, a short groove at the middle on top; granular scales at parietal region and occipital region intermixed with slightly large rounded granular tubercles, dense in occipital and temporal region and size increases towards nape; 9–13 supralabials present, size decreases towards angle of jaw; a series of scales, slightly larger than the loreal scales present above the supralabials; 8–12 infralabials, size decreases towards angle of jaw; two or three rows of slightly enlarged and narrow scales along the infralabials, posteriorly size of those decreases; rest of the gular scales are small, granular juxtaposed, nearly homogeneous, scales on throat are imbricate.
</paragraph>
<paragraph id="5EC4FE9B55FCB690B1E39E6948A35B2F">
Habitus slender (
<abbrev id="ABBRID0E1AEK" xlink_title="maximum body width">BW</abbrev>
/
<abbrev id="ABBRID0E5AEK" xlink_title="snout to vent length">SVL</abbrev>
= 0.16–0.22,
<abbrev id="ABBRID0ECBEK" xlink_title="trunk length, between the axilla and groin">TRL</abbrev>
/
<abbrev id="ABBRID0EGBEK" xlink_title="snout to vent length">SVL</abbrev>
= 0.41–0.47), dorsoventrally depressed; dorsal scales granular, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, continuing up to third or fifth segment of the tail, size of the tubercles increases towards posterior body, pronounced at sacrum and base of the tail, posteriormost rows indistinctly keeled; 15–21 rows of dorsal tubercles across mid dorsum; 27–38 paravertebral tubercles; ventrolateral fold present; ventral scales on chest and belly larger than those of dorsal, flat, smooth, cycloid, subimbricate to imbricate; 34–44 mid ventral scales; seven or nine distinct precloacal pores in male and 7–10 small precloacal pores in female;
<abbrev id="ABBRID0EKBEK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
followed by three to six unpored enlarged scales below.
</paragraph>
<paragraph id="FBA8A732B126673E687C88DCC784E8EF">
Forelimbs and hindlimbs slender (
<abbrev id="ABBRID0EQBEK" xlink_title="forearm length, taken from the elbow to the wrist">FL</abbrev>
/
<abbrev id="ABBRID0EUBEK" xlink_title="snout to vent length">SVL</abbrev>
= 0.13–0.15,
<abbrev id="ABBRID0EYBEK" xlink_title="crus length, taken from the knee to the base of the foot">CL</abbrev>
/
<abbrev id="ABBRID0E3BEK" xlink_title="snout to vent length">SVL</abbrev>
= 0.16–0.18); digits strongly inflected at the joints, all bearing large recurved claw and enlarged subdigital lamellae, dorsal scales on forelimbs heterogeneous, granular juxtaposed, smooth and subimbricate at distal end of forearm; forearm intermixed with a few rounded, large tubercles; dorsal scales of hindlimbs granular intermixed with large, rounded, bluntly conical tubercles; thigh scales towards lateral and dorsolateral side are smooth and subimbricate; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous, granular juxtaposed; scales on ventral side of hindlimbs slightly smaller than or nearly equal to those of belly, smooth, cycloid and subimbricate; scales on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate.
</paragraph>
<paragraph id="DAC23334F71077903DA787AFA3CF2B96">
Tail slender (
<abbrev id="ABBRID0ECCEK" xlink_title="tail length, from the cloaca to the tail tip">TL</abbrev>
=
<quantity id="8DDDC19420FDD395A98C56CA4A87ED6C" metricMagnitude="-2" metricUnit="m" metricValue="7.75" metricValueMax="9.7" metricValueMin="5.8" unit="mm" value="77.5" valueMax="97.0" valueMin="58.0">58–97 mm</quantity>
), gradually tapering, segments indistinct; dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size; large feebly keeled scales up to third to fifth segment of the tail, those on basal segment are pronounced and posterior rows with indistinct keel; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; two to four bluntly conical spurs present at the base of the tail. Detailed morphological characteristics of the collected samples are provided in Table S 4.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGCEK" type="Sequence divergence">
<paragraph id="DA9A72FEE8B6FD2E213E3578AA3A395D">
<heading id="2DE3283C476D34AD3E0B9365D77303C8" reason="title">Sequence divergence.</heading>
</paragraph>
<paragraph id="9016B803E4DD2F3E4737426025F8AEDA">
The intraspecific divergence within
<taxonomicName id="1A740EADDDE34B2A45EA2685424B5BD1" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="CCD6C08266E14897DE53936551A9386D" italics="true">C. cayuensis</emphasis>
</taxonomicName>
ranges between 0.1–6.4 %. Among the two clades identified by
<abbrev id="ABBRID0EXCEK" xlink_title="Bayesian Poisson Tree Processes">bPTP</abbrev>
, both clades have an intraspecific divergence between 0.1–4.3 % and 0.2–2.4 %, respectively. The divergence between these two clades range between 3–6.4 %.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2CEK" type="Colouration in life">
<paragraph id="B503F7703BE9108064D81DB916B75B74">
<heading id="ECD89F7288261A73881B22060188292D" reason="title">Colouration in life.</heading>
</paragraph>
<paragraph id="943CF06A3C03576575EA9878FB86BC20">Dorsal colour and marking pattern variable (Fig. S 3). Head and body greyish-brown, pale-brown to pale-yellowish-brown on top; head on top irregularly mottled with dark-brown, slightly enlarged dark-brown irregular patches may be present on occipital area; dark-brown postorbital stripe and loreal stripe present; a light stripe above loreal stripe and postorbital stripe may be visible; a few pale-yellow spots may be present on temporal and occipital region; supraciliary yellowish-brown; neck with short dark-brown streak with irregular edge or with irregular shaped patches; trunk dorsally with five to eight dark-brown bands consisting of irregular shaped and sized enlarged spots and posteriorly pale-yellowish or light-edged, these bands mid dorsally interrupted, or dorsum with dark-brown reticulation; limbs dorsally greyish-brown or pale-yellowish-brown or pale-brown with distinct or indistinct dark-brown reticulation; a dark-brown band on sacrum, band mid dorsally intersected; tail dorsally with 9–13 dark and 9–12 light bands alternatively arranged, bands are with irregular edged, anterior one or two dark bands may be mid dorsally intersected, dark bands are broad and light bands are comparatively narrow, dark bands are posteriorly becomes black or more dark-brown, while light bands become whitish; light bands may have a few dark-brown spots.</paragraph>
</subSubSection>
<subSubSection id="SECID0EADEK" type="distribution">
<paragraph id="C7C5E01291724B4421EF6507310D9001">
<heading id="99BCA824112410B94FBB1510B8F0CB21" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="DE222B6093D082C936AFA56C8F3EF1C1">
The
<typeStatus id="56046C7452172E48D70B41084D1869C4">type</typeStatus>
locality,
<collectingRegion id="CCF01467387DFDCD2590598B4DA75269" country="China" name="Xizang">Xizang</collectingRegion>
(
<collectingRegion id="1640CA3865AE026CCA9C597449F4E094" country="China" name="Xizang">Tibet</collectingRegion>
),
<collectingCountry id="8050427475CEFE288A1D97D1C6BA890B" name="China">China</collectingCountry>
by
<taxonomicName id="23CBA55514CAAB750C404CA1A998EF80" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="24A584D64025CCE57CD9D1141E16793A" italics="true">C. cayuensis</emphasis>
</taxonomicName>
also appears to be widely distributed in
<collectingRegion id="08238081176E002A0DBBC4B2A555A30B" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="3FB691275992F34A6FD5C90062EE3DA0" name="India">India</collectingCountry>
(Fig.
<figureCitation id="F1F9916961C1031B15599488433F545C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 B</figureCitation>
), within an elevational range of
<quantity id="311F248A55EFEF908B0F988579416614" metricMagnitude="2" metricUnit="m" metricValue="8.05" metricValueMax="12.0" metricValueMin="4.1" unit="m" value="805.0" valueMax="1200.0" valueMin="410.0">410–1200 m</quantity>
a. s. l.
<taxonomicName id="21F95172509E7678550E270A19BF4A61" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="2635E1D087874EAD98888F865ECF7601" italics="true">Cyrtodactylus cayuensis</emphasis>
</taxonomicName>
is widespread in the foothills of
<collectingRegion id="1042A0AE8FD505A6BE5D1DECA5C17D17" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
from the western part of the state, at Seijusa (Pakke Tiger Reserve) to the eastern part (Kamlang Tiger Reserve) and it has successfully spread across major tributaries of the Brahmaputra River, i. e., the Subansiri, Siang, Dibang, Lohit up to at least north of Noa-Dihing. Populations of
<taxonomicName id="FEA8D256B6D9349F7821AA5E7C6B6957" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="54A2B80E2B18BC5089AAC0148D4A0312" italics="true">C. cayuensis</emphasis>
</taxonomicName>
have been recorded in degraded evergreen forest as well as in undisturbed semi-evergreen and evergreen forests.
<taxonomicName id="8F899589036A29CFF3BB1F7E4839F34A" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="9F2F435888C14922D82894E7EAC63FD8" italics="true">Cyrtodactylus cayuensis</emphasis>
</taxonomicName>
is particularly abundant in the vegetation close to the picturesque Glaw Lake in the Kamlang Tiger Reserve. We observed individuals at night on shrubs and bedrocks (sedimentary rock) along small forest streams. We also encountered them on small trees up to a 2.0 m inside the forest at Jengging, Kamlang Tiger Reserve and Mehao Wildlife Sanctuary.
</paragraph>
</subSubSection>
</treatment>
</document>

319
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@ -0,0 +1,319 @@
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<mods:title id="A485C9F1507991C7E470EF85C185188C">Four new Dolichopoda species from Greece, one Troglophilus new to Greece and new locality records (Orthoptera, Rhaphidophoridae)</mods:title>
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<mods:affiliation id="772F227ADA8FAE98C36BACBDC0B53618">Dipartimento di Biologia e Biotecnologie “ C. Darwin ”, Università di Roma “ La Sapienza ”, Roma, Italy</mods:affiliation>
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<subSubSection id="61C0EAC9AB3B670EBB4D3FB733B2E584" type="nomenclature">
<paragraph id="C332C2CA5DF5AC5DDCEAC482097ACCD5">
<taxonomicName id="70F6CB2E856CC1825219D3DCB69970E9" authority="Alexiou" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="dirussoi" status="sp. nov.">
<emphasis id="364FB400E6E51A926E8A9150EEAB3B45" italics="true">Dolichopoda dirussoi</emphasis>
Alexiou
</taxonomicName>
<taxonomicNameLabel id="7D191AA1E25776CEAB1D138E2BDA35ED" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="20429359CD2B98D025F6D1FB1D88C7C7" type="description">
<paragraph id="7F07B03DAF82C197CE6D2DCF8C97FFFD">
<figureCitation id="14199DB6059EC739FAB98C56B276D7AE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographic distribution of D. athosensis sp. nov., D. dirussoi sp. nov., D. karoutsosi sp. nov. and D. kotsabasi sp. nov." figureDoi="10.3897/contrib.entomol.74.e121614.figure1" httpUri="https://binary.pensoft.net/fig/1103848">Figs 1</figureCitation>
,
<figureCitation id="42CCD907F4FE7C9BE874DD39DBC61074" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dolichopoda dirussoi sp. nov.: A. X tergum, male, dorsal view; B. Subgenital plate, female, lateral view; C. Epiphallus, lateral view; D. Epiphallus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure3" httpUri="https://binary.pensoft.net/fig/1103850">3</figureCitation>
,
<figureCitation id="31DAFA3A52DA3591B576AA999D9424A4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Geographic distribution of Dolichopoda species at W Sterea Ellas and S Ionian Islands." figureDoi="10.3897/contrib.entomol.74.e121614.figure4" httpUri="https://binary.pensoft.net/fig/1103851">4</figureCitation>
,
<figureCitation id="0C4C0BF96CC0F0439EACF310DDBC37EE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dolichopoda dirussoi sp. nov., in copula, cave Geralexis (photo Bakolitsas)." figureDoi="10.3897/contrib.entomol.74.e121614.figure5" httpUri="https://binary.pensoft.net/fig/1103852">5</figureCitation>
</paragraph>
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<paragraph id="4797BB2CE9436AD967815AEA86C8AAE7">
<heading id="F2B5851DFAD8725164CA5D04DC915050" reason="title">Type material.</heading>
</paragraph>
<paragraph id="AD574EED7F64623903906AD67C86A9A9">
<materialsCitation id="C1C1A236A8FE249C6FE5FAD01F7D73F6" collectingDate="2017-11-25" collectorName="K. Bakolitsas" country="Greece" county="Geralexis cave" elevation="102" latitude="38.635834" location="Sterea Ellas" longLatPrecision="1" longitude="21.210283" municipality="Skourtou" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="8DC3503A601CFA61166F1336EBC8DF14" bold="true" italics="true">
<typeStatus id="59EB731F9431283FD815D8AE45D9A417" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="509A2D8E2DD853039FF43BF98019B0BF" name="Greece">GREECE</collectingCountry>
•
<specimenCount id="6D46F3B47A083AD7D5873DA142C9D228" count="1" type="male">♂</specimenCount>
;
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,
<location id="FBE2E9DB03EF99759712ED3BA4B8AEA4" LSID="urn:lsid:plazi:treatment:57BD0BA9537D57418BC12159A226F739:FBE2E9DB03EF99759712ED3BA4B8AEA4" country="Greece" county="Geralexis cave" latitude="38.635834" longLatPrecision="1" longitude="21.210283" municipality="Skourtou" name="Nomos Etolias-Akarnanias">Nomos Etolias-Akarnanias</location>
,
<collectingMunicipality id="E3031E44251A5358DF3E812EEFE6AEB4">Skourtou</collectingMunicipality>
,
<collectingCounty id="98CF6F5DF5201BC2AD78785D5B3CEBCE">Geralexis cave</collectingCounty>
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<quantity id="E799718CEB3FBC7038D054A3E3D8050C" metricMagnitude="2" metricUnit="m" metricValue="1.02" unit="m" value="102.0">102 m</quantity>
a. s. l.
</elevation>
;
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<geoCoordinate id="6588C02500D141B83A8FE936916F7CFD" degrees="38" direction="north" minutes="38.150" orientation="latitude" precision="1" value="38.635834">38 ° 38.150 ' N</geoCoordinate>
,
<geoCoordinate id="48F927DA4EEC5ED1DB1372DC5B394F6A" degrees="21" direction="east" minutes="12.617" orientation="longitude" precision="1" value="21.210283">21 ° 12.617 ' E</geoCoordinate>
</named-content>
;
<date id="1567710B67488C55B2342B104F3DFF9E" value="2017-11-25">
<collectingDate id="6464AD07D9F804C2531FDFBB40B71FAD" value="2017-11-25">25 Nov. 2017</collectingDate>
</date>
;
<collectorName id="9F375592709EE07A51D64AA291189EB9">K. Bakolitsas</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0EEOAC" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
.
<materialsCitation id="8277534DE5790FBF0753AF1EDC912B27" collectingDate="2017-11-25" collectorName="K. Bakolitsas" country="Greece" county="Geralexis cave" elevation="102" latitude="38.635834" location="Sterea Ellas" longLatPrecision="1" longitude="21.210283" municipality="Skourtou" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<emphasis id="E70AE05244303BED2E20E682A587D605" bold="true" italics="true">
<typeStatus id="6DD5D5B784183AA1CA26944BCE37B772" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="52CE29C14F0230DADB101FA87ACF8445" name="Greece">GREECE</collectingCountry>
•
<specimenCount id="2330EB2E034A2C4AE7D757A9636F1EBA" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="1E498CF3D55642ECE21AD6ED27D1B670" count="3" type="female">3 ♀</specimenCount>
; same data as for holotype
</materialsCitation>
.
<emphasis id="0C659B3D45C0464E221F3BC5CD32FF01" bold="true" italics="true">Other material</emphasis>
:
<materialsCitation id="D11DE64C74AA7D4206628ED8784E0C53" collectingDate="2018-01-07" collectorName="K. Bakolitsas" country="Greece" county="cave Gouria" latitude="38.456383" location="Sterea Ellas" longLatPrecision="1" longitude="21.274633" municipality="Gouria Aitolikou" specimenCount="8" specimenCount-female="6" specimenCount-male="2">
<collectingCountry id="4ECDCA694D616770FDF4FA9181F41203" name="Greece">GREECE</collectingCountry>
•
<specimenCount id="C10D4187F29E1D77E9043072CFD4EF66" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="A54277A05A829957CDB61D9A9C2F48C6" count="6" type="female">6 ♀</specimenCount>
;
<location id="FE4F7BFD70DDE03025F4EBC9A4F09217" LSID="urn:lsid:plazi:treatment:57BD0BA9537D57418BC12159A226F739:FE4F7BFD70DDE03025F4EBC9A4F09217" country="Greece" county="cave Gouria" latitude="38.456383" longLatPrecision="1" longitude="21.274633" municipality="Gouria Aitolikou" name="Sterea Ellas">Sterea Ellas</location>
,
<location id="6209338C29CCD3236D640CC4F2EF4391" LSID="urn:lsid:plazi:treatment:57BD0BA9537D57418BC12159A226F739:6209338C29CCD3236D640CC4F2EF4391" country="Greece" county="cave Gouria" latitude="38.456383" longLatPrecision="1" longitude="21.274633" municipality="Gouria Aitolikou" name="Nomos Etolias-Acarnanias">Nomos Etolias-Acarnanias</location>
,
<collectingMunicipality id="E98B7E4643B027233DF49796C8FB62AF">Gouria Aitolikou</collectingMunicipality>
,
<collectingCounty id="E1CC70F532ACC3D9B0D9BCF07EEA5271">cave Gouria</collectingCounty>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESOAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[21.274633,38.456383]}">
<geoCoordinate id="06AD774C83248275DCD7EBDE760D6A49" degrees="38" direction="north" minutes="27.383" orientation="latitude" precision="1" value="38.456383">38 ° 27.383 ' N</geoCoordinate>
,
<geoCoordinate id="F311632ABB8590D0C33C7D7063063364" degrees="21" direction="east" minutes="16.478" orientation="longitude" precision="1" value="21.274633">21 ° 16.478 ' E</geoCoordinate>
</named-content>
;
<date id="5ED5B6BCEEBB752589621AD9A2098A0B" value="2018-01-07">
<collectingDate id="D994C0110F0299CF76826A75E20EAA7E" value="2018-01-07">7 Jan. 2018</collectingDate>
</date>
;
<collectorName id="4ED53B6982F7B879B404670250809969">K. Bakolitsas</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0EXOAC" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3OAC" type="diagnosis">
<paragraph id="03C900DAF3E89BA96C5DEEDD5BBC988A">
<heading id="CAE0B8C9AD4BE4F191F2CFBFC93D7393" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="3014EC8BDC689434FDB90E7AFD648A6A">
The new taxon shows affinities with the other
<taxonomicName id="66EE6B5D3B8354D1AB5254506328174E">
<emphasis id="26194473BA365A92645C893F090AF72E" italics="true">Dolichopoda</emphasis>
species
</taxonomicName>
of the west Sterea Ellas,
<taxonomicName id="AA76BB77EA4D29A9F1834238EAD7FCCA" authority="Rampini &amp; Di Russo, 2008" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="giachinoi">
<emphasis id="600C9678DF921779DC421AF539DA4491" italics="true">D. giachinoi</emphasis>
<bibRefCitation id="3CA1CDE7CA82D48F86A4C73EA2F5FC63" DOI="10.11646/zootaxa.1923.1.1" author="Rampini M &amp; Di Russo C &amp; Pavesi F &amp; Cobolli M" journalOrPublisher="Zootaxa" pagination="1-17" refId="B12" refString="Rampini M, Di Russo C, Pavesi F, Cobolli M (2008) The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae). Zootaxa 1923: 1 - 17. https: // doi. org / 10.11646 / zootaxa. 1923.1. 1" title="The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae)." volume="1923" year="2008">Rampini &amp; Di Russo, 2008</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="1A565278A0E800F952F6AF1BE02E019A" authority="Rampini &amp; Di Russo, 2017" authorityName="Rampini &amp; Di Russo" authorityYear="2017" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="bakolitsasi">
<emphasis id="8BC1C125B7DA090A97B7719F14A12B11" italics="true">D. bakolitsasi</emphasis>
Rampini &amp;
<bibRefCitation id="515AA7CDAD7BA81F914823F5D5A8EAE2" DOI="10.11646/zootaxa.4290.2.9" author="Di Russo C &amp; Alexiou S &amp; Rampini M" journalOrPublisher="Zootaxa" pagination="380-384" refId="B8" refString="Di Russo C, Alexiou S, Rampini M (2017) A new species of Dolichopoda (Orthoptera: Rhaphidophoridae; Dolichopodainae) from Western Greece. Zootaxa 4290: 380 - 384. https: // doi. org / 10.11646 / zootaxa. 4290.2. 9" title="A new species of Dolichopoda (Orthoptera: Rhaphidophoridae; Dolichopodainae) from Western Greece." volume="4290" year="2017">Di Russo, 2017</bibRefCitation>
</taxonomicName>
as well as the species of the south
<collectingCountry id="8EE23EC7E74E8C8F3D81F68394C7892A" name="Greece">Ionian Islands</collectingCountry>
<taxonomicName id="A0DBC543031D6300799403C8357BEABC" authority="Rampini &amp; Di Russo, 2008" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="gasparoi">
<emphasis id="769CB1A5DD59E52D0A34CDFD6CCB2C00" italics="true">D. gasparoi</emphasis>
<bibRefCitation id="1008AFAD6A287B5A6B87A539450CB559" DOI="10.11646/zootaxa.1923.1.1" author="Rampini M &amp; Di Russo C &amp; Pavesi F &amp; Cobolli M" journalOrPublisher="Zootaxa" pagination="1-17" refId="B12" refString="Rampini M, Di Russo C, Pavesi F, Cobolli M (2008) The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae). Zootaxa 1923: 1 - 17. https: // doi. org / 10.11646 / zootaxa. 1923.1. 1" title="The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae)." volume="1923" year="2008">Rampini &amp; Di Russo, 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7DD589FFAA3F934A09D20FE1731ED2F3" authority="Rampini &amp; Di Russo, 2008" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="ithakii">
<emphasis id="1A40F3D765B9AB195B823EBBE22823C1" italics="true">D. ithakii</emphasis>
<bibRefCitation id="01FCE82F80D823920CBFB7817F5C097E" DOI="10.11646/zootaxa.1923.1.1" author="Rampini M &amp; Di Russo C &amp; Pavesi F &amp; Cobolli M" journalOrPublisher="Zootaxa" pagination="1-17" refId="B12" refString="Rampini M, Di Russo C, Pavesi F, Cobolli M (2008) The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae). Zootaxa 1923: 1 - 17. https: // doi. org / 10.11646 / zootaxa. 1923.1. 1" title="The genus Dolichopoda in Greece. A description of new species from the Ionian Regions and Peloponnisos (Orthoptera, Rhaphidophoridae)." volume="1923" year="2008">Rampini &amp; Di Russo, 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="801D3A88614D5D584C3C76E8F4944FB2" authority="Galvagni, 2002" authorityName="Galvagni" authorityYear="2002" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="pavesii">
<emphasis id="D599096C27BCF4A1227ACD76F79BBB3A" italics="true">D. pavesii</emphasis>
Galvagni, 2002
</taxonomicName>
and
<taxonomicName id="E7F64EE455619CB38408D2F2BC1C6E78" authority="Chopard, 1964" authorityName="Chopard" authorityYear="1964" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="patrizii">
<emphasis id="62DEA65BAD6BAA8EE10D68C7F577AB24" italics="true">D. patrizii</emphasis>
<bibRefCitation id="18AB946214E1B622FD7647245A081C7A" DOI="10.3406/bsef.1964.20741" author="Chopard L" journalOrPublisher="Bulletin de la Société Entomologique de France" pagination="7-20" refId="B5" refString="Chopard L (1964) Descriptions d'orthopteres cavernicoles de Grece. Bulletin de la Societe Entomologique de France 69: 7 - 20. https: // doi. org / 10.3406 / bsef. 1964.20741" title="Descriptions d’orthoptères cavernicoles de Grèce." volume="69" year="1964">Chopard, 1964</bibRefCitation>
</taxonomicName>
. (Fig.
<figureCitation id="775CB0D2E525603FBF8AE4BAB33B9E82" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Geographic distribution of Dolichopoda species at W Sterea Ellas and S Ionian Islands." figureDoi="10.3897/contrib.entomol.74.e121614.figure4" httpUri="https://binary.pensoft.net/fig/1103851">4</figureCitation>
). The combination of the morphological characters of
<taxonomicName id="3DB29EDE2D8B6F34C8A6798B090821C0" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="dirussoi" status="sp. nov.">
<emphasis id="9018D2A2F307B892DD92E9615DE83252" italics="true">D. dirussoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8BD5CF64A933DE87BB5DE4323B183BEA" rank="species">sp. nov.</taxonomicNameLabel>
is unique among the neighbouring species, with the most outstanding difference being the lack of a ridge or tubercles on the X tergite of males. Despite the obvious differences of the new species from
<taxonomicName id="ADAF69E9297506E8FA02B69611E5BD45" authorityName="Rampini &amp; Di Russo" authorityYear="2017" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="bakolitsasi">
<emphasis id="625E7B2B06EC6408D7DE9E6CFF12CE78" italics="true">D. bakolitsasi</emphasis>
</taxonomicName>
, especially in the shape of the lateral lobes of the X tergum, the two species stand out from the others by the strongly curved median process of the epiphallus.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGCAE" type="description">
<paragraph id="C791E25E063B0DBA2C70948F38116060">
<heading id="F7DE0EEB08DB7FD0F6744F130087ED19" reason="title">Description.</heading>
</paragraph>
<paragraph id="4DADB05E1E78160F15D9761F83EB452A">Body pale brownish-yellow, legs elongated, hind femora unarmed. Fore tibiae with 3 spines on both sides of the ventral side, mid tibiae with 4 spines on both sides of the ventral side and hind tibiae with 20 spines on both sides of the dorsal side.</paragraph>
<paragraph id="81AB12DBF69CC7517B5DF22287D63871">
X tergum (last abdominal tergite) (Fig.
<figureCitation id="D860454B1CB621E8E9368B80828D72C5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dolichopoda dirussoi sp. nov.: A. X tergum, male, dorsal view; B. Subgenital plate, female, lateral view; C. Epiphallus, lateral view; D. Epiphallus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure3" httpUri="https://binary.pensoft.net/fig/1103850">3 A</figureCitation>
) with two lateral lobes, lobes shallowly bilobed at apex. Lateral lobes separated by a broad concavity, broader than the lobes, inner side of the concavity slightly more thickened.
</paragraph>
<caption id="B7247537F63800E21208FF517BF66E66" ID-DOI="10.3897/contrib.entomol.74.e121614.figure3" ID-arpha="F13DF463-3386-5047-935B-B4A12AD8CDBD" httpUri="https://binary.pensoft.net/fig/1103850" startId="F3">
<paragraph id="864EAEF81DD01C5B074DD10E5F87666F">
<label id="74684222C1B7424BE318DF38C8183D6F">Figure 3.</label>
</paragraph>
<paragraph id="B0FB72DF91D23D96D0CA212CCA61888F">
<taxonomicName id="A8C342B9DEEBE3BD0DD9FE886D977E62" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="dirussoi" status="sp. nov.">
<emphasis id="921D87CC9204B81B6E5A68A624A46F78" italics="true">Dolichopoda dirussoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D64D3768099DA9743D631903AFD7A04C" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="17577DF316908554B942A75B7E2A9634" bold="true">A.</emphasis>
X tergum, male, dorsal view;
<emphasis id="FF04F15B14D129DE04A8F2877FA84498" bold="true">B.</emphasis>
Subgenital plate, female, lateral view;
<emphasis id="C76BA7DC10455186ADD2A398F1486ABA" bold="true">C.</emphasis>
Epiphallus, lateral view;
<emphasis id="BA8DFF944002B3E42A226BE444363BBC" bold="true">D.</emphasis>
Epiphallus, dorsal view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="CD46320C54992693C36F8CAEC5ED5C10" ID-DOI="10.3897/contrib.entomol.74.e121614.figure4" ID-arpha="859F3FF4-2A61-5822-A352-144917B0AC78" httpUri="https://binary.pensoft.net/fig/1103851" startId="F4">
<paragraph id="BA867CA9B6F76284A47B17C3BFD4323F">
<label id="57579B47D31BDBE8D258F94B9A66A830">Figure 4.</label>
</paragraph>
<paragraph id="65025C02FEBAF285D16911D8ACC7BFB8">
Geographic distribution of
<taxonomicName id="7A55D116EEE75499ECA0DFB494146DF5">
<emphasis id="FBC0E7A9B45C3AAB451D91FF69A446FD" italics="true">Dolichopoda</emphasis>
species
</taxonomicName>
at W Sterea Ellas and S Ionian Islands.
</paragraph>
</caption>
<paragraph id="457296DE102214F1A051DDDF9E561821">Subgenital plate convex, with a wide median incision that runs for half of the total length; lateral lobes rather rounded, styli short.</paragraph>
<paragraph id="CF82885311BAEA0E5643CB89B2C70446">
Epiphallus sclerotized, with the median process thickened to about 2 / 3 of its length, strongly curved and acute (Fig.
<figureCitation id="455B4C50DF903D83C67ABCA8C8968522" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dolichopoda dirussoi sp. nov.: A. X tergum, male, dorsal view; B. Subgenital plate, female, lateral view; C. Epiphallus, lateral view; D. Epiphallus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure3" httpUri="https://binary.pensoft.net/fig/1103850">3 C, D</figureCitation>
).
</paragraph>
<paragraph id="B35DFB6CAC6504B2BCC74EEB6427996A">Measurements (length in mm): body 16, pronotum 3.4, fore femora 17, middle femora 18, hind femora 23.</paragraph>
<paragraph id="AEE5E0B92A74131BC1C1D622FAE736B0">
<emphasis id="CD91FB3635C4692904EB467D249CD3F6" bold="true">Female</emphasis>
: Characters similar to male. Subgenital plate large, triangular with a rounded apex. Sternite VII with a very large protuberance covering the anterior half of the sternite (Fig.
<figureCitation id="7B441C0C88BA464DE664D9A153ADD563" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dolichopoda dirussoi sp. nov.: A. X tergum, male, dorsal view; B. Subgenital plate, female, lateral view; C. Epiphallus, lateral view; D. Epiphallus, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure3" httpUri="https://binary.pensoft.net/fig/1103850">3 B</figureCitation>
), in the shape of flattened triangle with thickened margins and rounded apex. Ovipositor
<quantity id="E8D3AEA672CB84D1042D4C334997CDF2" metricMagnitude="-2" metricUnit="m" metricValue="1.4" unit="mm" value="14.0">14 mm</quantity>
in length, almost straight, the inferior valves with 19 apical denticles.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVEAE" type="etymology">
<paragraph id="8452AD24008FE8C359AA547AB79D205B">
<heading id="E8CDF8DC6C391A4A1CBB5686DB898FA6" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="7846CF059116EB521C8B3C61BA75D3C9">The new species is named after our dear friend and co-author Claudio Di Russo, who introduced S. A. into the study of Greek cave crickets thirteen years ago.</paragraph>
<caption id="5AA5BBA02EBD39DAC6F0E58F339A7DEB" ID-DOI="10.3897/contrib.entomol.74.e121614.figure5" ID-arpha="47A40A97-8367-56C4-953D-4B8E0296BA8D" httpUri="https://binary.pensoft.net/fig/1103852" startId="F5">
<paragraph id="029C0B4175A68A8C020072D1271DD7DC">
<label id="4F38B024AB2EB34860BBD42871435AA5">Figure 5.</label>
</paragraph>
<paragraph id="6268685AC562B3543E60F14563E8CD8F">
<taxonomicName id="A80C55E1ABAB14E67136EA013E58BAA6" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="dirussoi" status="sp. nov.">
<emphasis id="EF16A99CFB99F21496E91B24634468FA" italics="true">Dolichopoda dirussoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="401FA7F9A71E4FCFA95E35D7D5B86E27" rank="species">sp. nov.</taxonomicNameLabel>
, in copula, cave Geralexis (photo Bakolitsas).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ESFAE" type="distribution">
<paragraph id="EA54E0D8355BF7A0B5E212AC38E24939">
<heading id="938FB3B1329BFFE99635E4DBB0576B0E" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="BC3BDE9B69FA5B9E62F79E084862F16B">
Known only from the two caves of Etolia-Akarnania. Cave Geralexis is approximately
<quantity id="9B321ACFF24788A527127EB42A416B1E" metricMagnitude="1" metricUnit="m" metricValue="3.0" unit="m" value="30.0">30 m</quantity>
long, with its entrance facing east. It appears to have been twice as long but has collapsed. The entrance is quite large (4.0 × 8.0 m), although well hidden by lush vegetation. The cave is composed of one main room and a second smaller room to the left, where the cave crickets were found.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:affiliation id="3A03C4AC679CCBD612B3072D139C29F9">Department of Zoology &amp; Entomology, University of Pretoria, Private Bag X 20, Hatfield, 0028, South Africa &amp; Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
</mods:name>
<mods:name id="FA4721FF774950035A6B984BB7C2C480" type="personal">
<mods:role id="D17467D14D109F970E4E7CD5AE40B0EA">
<mods:roleTerm id="FFAD48F26DAF03887576B6D95CEF718F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="644BF72D57ACCD5E609A145FC93CF8FD">Strümpher, Werner P.</mods:namePart>
<mods:nameIdentifier id="A940DF9DC373EB0A85999752553E043F" type="ORCID">0000-0002-7047-2666</mods:nameIdentifier>
<mods:affiliation id="A04ACB1FA66A769CFB1E1920826CA8F9">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa</mods:affiliation>
</mods:name>
<mods:name id="B194C0015FF51C20DB0AEFF26D0E1B39" type="personal">
<mods:role id="DB85BD5EB17C7D0A769A8A7B51959018">
<mods:roleTerm id="2BE94BD164DA6DFA3375CEA6936DEB79">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CAA8EA67CEDD08CE1D2286B56AFA9898">Barclay, Maxwell V. L.</mods:namePart>
<mods:nameIdentifier id="A6B791F7ADAE1230727F4F9496D88625" type="ORCID">0000-0003-4989-2014</mods:nameIdentifier>
<mods:affiliation id="9603E4583D491F34FB494A732A8CC01F">Department of Life Sciences, Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name id="4176200ABE6C3CA0BDCB4ADEDB2B8ABB" type="personal">
<mods:role id="86599C4A169396F8C14159F9F8CB32F4">
<mods:roleTerm id="0EB364BCA1EBC769E5842E8F99A4029A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C26F870FCA63F7287D0BA9125A3FE70A">Vaz-de-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="5B60C8417929FFEF426879C77C1A62D1" type="ORCID">0000-0001-9697-320X</mods:nameIdentifier>
<mods:affiliation id="D669C27902550D9476A7D6B21A9D53E5">Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B471DF9BBC8009FD725E8E0EA44DF14F">text</mods:typeOfResource>
<mods:relatedItem id="FDDD3FFD108449DCB743D14AC48B244C" type="host">
<mods:titleInfo id="C3B45FEA6FBE85F99C89CBEA70600A60">
<mods:title id="69B739E0496C2A04047778CED70DFED0">Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="5CF8B6A8CA1D672025E12F9E8D750380">
<mods:date id="FADA77C8EA3C45AA58801378698EC22E">2024</mods:date>
<mods:detail id="546A6BDA132926EDFF52E31E296BA882" type="pubDate">
<mods:number id="2A8627CE6C83C4C8207EAA2F2F97D279">2024-07-29</mods:number>
</mods:detail>
<mods:detail id="F743B333B578C27BF65D7A3C54C7900B" type="volume">
<mods:number id="198350F4684A4F0BC9B6A1B40D245427">74</mods:number>
</mods:detail>
<mods:detail id="3FB2D130C85D2F0EB13DCE07E5B5C460" type="issue">
<mods:number id="FF1024216588C0BF66E12574AC9C71E2">1</mods:number>
</mods:detail>
<mods:extent id="1D0D3927916A32A83F91AC5C6D3A7D89" unit="page">
<mods:start id="7094B0833E043250766C3918C76F0A3A">81</mods:start>
<mods:end id="69488944BEE4CDAF688AD0AB0F81E9CA">101</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3B17C06FF0911F5C12D485C4125D6223">journal article</mods:classification>
<mods:identifier id="64A8C97F5C13A2AF1F9DDAF450504169" type="DOI">10.3897/contrib.entomol.74.e126799</mods:identifier>
<mods:identifier id="C77AE3A906B7BA5C15EE380BFE56B94B" type="ZooBank">339AB68C-DCB1-435F-B88E-2C42D0A7B05E</mods:identifier>
</mods:mods>
<treatment id="5A39B49D422A51EC89FA77D9D59DB106" ID-DOI="http://doi.org/10.5281/zenodo.13123987" ID-Zenodo-Dep="13123987" ID-arpha="5A39B49D-422A-51EC-89FA-77D9D59DB106" LSID="urn:lsid:plazi:treatment:5A39B49D422A51EC89FA77D9D59DB106" httpUri="http://treatment.plazi.org/id/5A39B49D422A51EC89FA77D9D59DB106">
<subSubSection id="35D16A468F63D6E9F83626960ACD4D5E" type="nomenclature">
<paragraph id="A0BF5169264DB8727424A3F9D5C1CA0F">
<taxonomicName id="05442DF7EB665F6189AB8E01A94916BC" ID-CoL="49DRL" authority="(Vaurie, 1955)" baseAuthorityName="Vaurie" baseAuthorityYear="1955" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="loxus" subGenus="Omorgus">
<emphasis id="2A002517F66BEF01A644D32354167859" italics="true">Omorgus</emphasis>
(
<emphasis id="B8B09F101CD3C3E8A465D619E217C726" italics="true">Omorgus</emphasis>
)
<emphasis id="D0602477102D0910819C894F05A3D84C" italics="true">loxus</emphasis>
(
<bibRefCitation id="BB6D767B16A51F2E77EE4670B4652474" author="Vaurie P" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="1-89" refId="B108" refString="Vaurie P (1955) A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). The Bulletin of the American Museum of Natural History 106: 1 - 89." title="A revision of the genus Trox in North America (Coleoptera: Scarabaeidae)." volume="106" year="1955">Vaurie, 1955</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="150060509A2083A2BDFE812B51F5AF47" type="reference_group">
<paragraph id="A73860144E9FDE6579BD88F98BA55CCE">
<treatmentCitationGroup id="8EF9E105DB7074E7348E3110C8D9F047">
<treatmentCitation id="81FC63DEDEEDCAC5C45BF5B1128E9129" author="Vaurie P" page="58" year="1955">
<taxonomicName id="60D3581A895BFF3C703BD72D1F9309F4" ID-CoL="59DM8" authority="Vaurie, 1955: 58" authorityName="Vaurie" authorityPageNumber="58" authorityYear="1955" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="loxus">
<emphasis id="DA41BE28BBA6B56D42016952C43CCF98" italics="true">Trox loxus</emphasis>
<comment id="1FEB31B17BE69388634E33FB2A0875B4">
<bibRefCitation id="17EF6E029CD8089932E6E6C496BB0E97" author="Vaurie P" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="1-89" refId="B108" refString="Vaurie P (1955) A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). The Bulletin of the American Museum of Natural History 106: 1 - 89." title="A revision of the genus Trox in North America (Coleoptera: Scarabaeidae)." volume="106" year="1955">Vaurie, 1955: 58</bibRefCitation>
(original description);
<treatmentCitation id="1903FC443D99EECB77657C2DE20F54FA" author="Vaurie P" page="147" year="1962">
<bibRefCitation id="E69FECB61F6996BEF29DCA9B2FC58A61" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" page="147" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105–167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962: 147</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="66C6BB43EA80FDBD0255D1B1BDC9B17B" author="Ratcliffe BC" page="301" year="1978">
<bibRefCitation id="B4DD5C3B6CCBC150B26D08146D6DF7B1" DOI="10.1590/1809-43921978082299" author="Ratcliffe BC" firstAuthor="Ratcliffe" issue="2" journalOrPublisher="Acta Amazonica" page="301" pagination="299-302" refId="B88" refString="Ratcliffe BC (1978) A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae). Acta Amazonica 8 (2): 299–302. https://doi.org/10.1590/1809-43921978082299" title="A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae)." volume="8" year="1978">Ratcliffe 1978: 301</bibRefCitation>
</treatmentCitation>
(new distribution records).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="C8CF9578101690BBA525A8CFFF686407">
<treatmentCitationGroup id="9F835307EA5C754F13CD849A927622B9">
<treatmentCitation id="0BEE4DF44AEE9DFE54B90D69026EF90D">
<taxonomicName id="4E6B79AE6A1186D358040A8457395367" ID-CoL="59DM8" authorityName="Vaurie" authorityYear="1955" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="loxus" subGenus="Omorgus">
<emphasis id="CBFABD0FF6AD2D42F69F8B40621BC4EA" italics="true">Trox</emphasis>
(
<emphasis id="503D92BBF34E6C5F23DA3D9A189B1F0B" italics="true">Omorgus</emphasis>
)
<emphasis id="541AE9D19D70E7AB97039DAA1BF76451" italics="true">loxus</emphasis>
</taxonomicName>
<comment id="1D2D6769AAEF51D66989DC46C73AFB0D">
:
<treatmentCitation id="2B0BA2E2E387523821062106AC5BEA7B" author="Scholtz CH" page="11" year="1982">
<bibRefCitation id="C2E74B532153F62ADF73239E18C4F965" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa" page="11" pagination="1-27" refId="B96" refString="Scholtz CH (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa 54: 1–27." title="Catalogue of world Trogidae (Coleoptera: Scarabaeoidea)." volume="54" year="1982">Scholtz 1982: 11</bibRefCitation>
</treatmentCitation>
(catalogue).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="6ABE61D0D074752F235843C711AC6423">
<treatmentCitationGroup id="0A796C59B0424CAA05380D78C546AF79">
<treatmentCitation id="B7E5618013E0929667017FBAD7785ADD">
<taxonomicName id="E15393CCA47A0B326B1402B59B99491F" ID-CoL="49DRL" baseAuthorityName="Vaurie" baseAuthorityYear="1955" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="loxus" subGenus="Omorgus">
<emphasis id="B99D3F977F91866AD188AF1F26602B34" italics="true">Omorgus</emphasis>
(
<emphasis id="17042944F18EC680A8EDCE377BCEB4A8" italics="true">Omorgus</emphasis>
)
<emphasis id="24107353E27BCD95CE73B2D89109B685" italics="true">loxus</emphasis>
</taxonomicName>
<comment id="682DE14DFAF222FE2062DCAF12498977">
:
<treatmentCitation id="F429B65EC5C3D72D49EBD22FF5113AC6" author="Scholtz CH" page="361" year="">
<bibRefCitation id="D5F3D6EC746F40497AA5B8C88BF91C75" DOI="10.1111/j.1365-3113.1986.tb00186.x" author="Scholtz CH" journalOrPublisher="Systematic Entomology" pagination="355-363" refId="B97" refString="Scholtz CH (1986 a) Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology 11: 355–363. https://doi.org/10.1111/j.1365-3113.1986.tb00186.x" title="Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea)." volume="11">Scholtz 1986 a</bibRefCitation>
: 361
</treatmentCitation>
(phylogenetics);
<treatmentCitation id="12423392DFEA30E3CAA72CCE8F1669B1" author="Scholtz CH" page="1416" year="1990">
<bibRefCitation id="855184FE097BB03BFBCE8FD42AA95696" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" page="1416" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990: 1416</bibRefCitation>
</treatmentCitation>
(redescription),
<treatmentCitation id="2169ADBB5AB90CD090A23E4F80939722" author="Moragues G." page="77" year="2010">
<bibRefCitation id="C0EDD6CEDD62BEBCBBBEC159324EA75C" author="Moragues G." firstAuthor="Moragues" page="77" refId="B70" refString="Moragues G. (2010) Aperçu des Trox de Guyane (Coleoptera: Trogidae). Contribution à l’étude des Coléoptères de Guyane, 1: 76–78. [Supplément au Bulletin de liaison de l’ACOREP France “ Le Coléoptériste ”]." year="2010">Moragues 2010: 77</bibRefCitation>
</treatmentCitation>
(checklist – as “
<emphasis id="67AF4737EA138900240A5506C123C5DC" italics="true">lauxus</emphasis>
”);
<treatmentCitation id="273B66A32C24ED63C42E5DCA81EC48F7" author="Zidek J" page="12" year="2013">
<bibRefCitation id="00C6027EB35FE47F9E344AA806C5A0AF" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="12" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1–38." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 12</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="51909CE56669B901988AF9BC548C1406" author="Zidek J" page="103" year="2017">
<bibRefCitation id="CD0C37AB2D44BCAB8486A0CD6E9B8531" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="103" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 103</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="756775F217CC7C99BC10BC7659F4E47B" author="Hielkema AJ &amp; Hielkema MA" page="11" year="2019">
<bibRefCitation id="3FF57A911BFD6CFA42FF9D0575563352" author="Hielkema AJ &amp; Hielkema MA" firstAuthor="Hielkema" journalOrPublisher="Insecta Mundi" page="11" pagination="1-306" refId="B53" refString="Hielkema AJ, Hielkema MA (2019) An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732: 1–306." title="An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas." volume="732" year="2019">Hielkema and Hielkema 2019: 11</bibRefCitation>
</treatmentCitation>
(checklist of the Guianas);
<treatmentCitation id="0D1FF152BA1C4D686350B0BB72F1D81D" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" httpUri="http://treatment.plazi.org/id/03DA2410FFA8FFEDFE4EFDE6FDFA1B0E" page="1999" year="2021">
<bibRefCitation id="CAABEEF8FB95F52C82A713237638B736" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="1999" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021: 1999</bibRefCitation>
</treatmentCitation>
(review of Brazilian species).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="05F155C1B51A83DA09B66C370A73F54D">
<treatmentCitationGroup id="CCD4EC1B9170A2AC8EF9E32FC1F6E1FC">
<treatmentCitation id="BFFF42DEC532A0AC9DE44A7A93BC6854">
<taxonomicName id="2503F88D4FB4098B643F2483A01613D4" ID-CoL="49DRL" baseAuthorityName="Vaurie" baseAuthorityYear="1955" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="loxus">
<emphasis id="28AFCC690D48020F3EDFF508E69A69F1" italics="true">Omorgus loxus</emphasis>
</taxonomicName>
<comment id="738703A9E1B3CDD8D3957AA248D7B216">
:
<treatmentCitation id="8A27B3B3DC75620F293EEACC1A71A555" author="Deloya C" page="67" year="2000">
<bibRefCitation id="6DAB0C4D09A4DDA558063C0E4415CD39" DOI="10.21829/azm.2000.81811871" author="Deloya C" firstAuthor="Deloya" journalOrPublisher="Acta Zoológica Mexicana" page="67" pagination="63-76" refId="B31" refString="Deloya C (2000) Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia). Acta Zoológica Mexicana 81: 63–76. https://doi.org/10.21829/azm.2000.81811871" title="Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia)." volume="81" year="2000">Deloya 2000: 67</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="5211487CCCA50F16A6DD9EE6B01CD443" author="Diéguez VM &amp; Gómez RS" page="94" year="2004">
<bibRefCitation id="6ACA8FE641FEB056898AEC2273E475CE" author="Diéguez VM &amp; Gómez RS" firstAuthor="Dieguez" issue="1-2" journalOrPublisher="Revista de la Sociedad Entomológica de Argentina" page="94" pagination="92-95" refId="B36" refString="Diéguez VM, Gómez RS (2004) Aporte al conocimiento de las Trogidae de la Argentina. Revista de la Sociedad Entomológica de Argentina 63 (1–2): 92–95." title="Aporte al conocimiento de las Trogidae de la Argentina." volume="63" year="2004">Diéguez and Gómez 2004: 94</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="3D44BE7ADDA698FBA727034E70BEC52D" author="Deloya C" page="122" year="2005">
<bibRefCitation id="C45AB900CF3DFAAB06CFE05422B14F57" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121–129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="DE6CD3C895D0A7523A9585EBEEAF2A8E" author="Gómez RS" page="515" year="2008">
<bibRefCitation id="37552E8A7258D7E6F630D9B581D78384" author="Gómez RS" editor="Claps LE &amp; Debandi G &amp; Roig-Juñent S" firstAuthor="Gomez" journalOrPublisher="Sociedad Entomológica Argentina" page="515" pagination="509-518" refId="B46" refString="Gómez RS (2008) Trogidae. In: Claps LE, Debandi G, Roig-Juñent S (Eds) Biodiversidad de Artrópodos Argentinos II. Sociedad Entomológica Argentina, 509–518." title="Trogidae." volumeTitle="Biodiversidad de Artrópodos Argentinos II." year="2008">Gómez 2008: 515</bibRefCitation>
</treatmentCitation>
(key for Argentinean species).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFTBG" type="Type specimen examined">
<paragraph id="49A7A8D0CEE889AEE75A7F9BC8288C26">
<heading id="E2EC941BD7CC501B6800A7FC00C17D88" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="23181946324CD5DF3E8D2D5D48F63912">
<materialsCitation id="2B8915E224F4F1FB018FF5FF83511C29" collectionCode="MCZH" country="Mexico" location="Mexico" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="491D3DEDD8F0F5B266B93A4A9C3E1015" bold="true" italics="true">
<typeStatus id="E8F9AE9600FBE1F858D5BC42F86A2BD1" type="holotype">Holotype</typeStatus>
, by original designation
</emphasis>
(
<specimenCount id="046A6F075419EBE16B7EB9F89419B385" count="1" type="female">♀</specimenCount>
<emphasis id="8DBB820871AA619EFBB1227E0D1CD59E" bold="true">
<collectionCode id="F546DA91192F2A2E80496EC3107D520F">MCZH</collectionCode>
</emphasis>
). See
<bibRefCitation id="A5753A5AFB4FCF40C2D162F74AF93972" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="1999" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021: 1999)</bibRefCitation>
for high-quality images of the type specimen.
<emphasis id="08BB037A801368609E88C451324A114B" bold="true">Type locality</emphasis>
: “ El Palmar, 16 k. W. of Tetzonapa ” [
<collectingCountry id="81ED483CC26481D5409C8F47D47F005E" name="Mexico">Mexico</collectingCountry>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYTBG" type="distribution">
<paragraph id="D5EC77CB19D2875B3E4E64E31CA37F08">
<heading id="ECE7A01F2FB12FBF39A2062B7887706F" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="87C8014F2E77498DDDBC90DC74AC3194">
<collectingCountry id="71C2DF5EF148E0F3C9E02E5233829250" name="Argentina">Argentina</collectingCountry>
,
<collectingCountry id="2FCFF275164B0E1066AFAB3F6CE0322B" name="Brazil">Brazil</collectingCountry>
,
<collectingCountry id="63292CC591C2F30478CCF5E28A7326ED" name="Colombia">Colombia</collectingCountry>
,
<collectingCountry id="7211F211975B89C8CC49BF0594B7923A" name="Costa Rica">Costa Rica</collectingCountry>
, French Guiana,
<collectingCountry id="A92F09F0C0D21C5710B7EA05285342D9" name="Mexico">Mexico</collectingCountry>
,
<collectingCountry id="E56F26D97F5845BB7C852EFFFC6BE22B" name="Panama">Panama</collectingCountry>
, and
<collectingCountry id="5C3A82EB6730EFED815B366A4F0793FE" name="Paraguay">Paraguay</collectingCountry>
(
<bibRefCitation id="898A89771B5FE48D6D3FE3F8774BA2C8" author="Vaurie" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="1-89" refId="B108" refString="Vaurie P (1955) A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). The Bulletin of the American Museum of Natural History 106: 1–89." title="A revision of the genus Trox in North America (Coleoptera: Scarabaeidae)." volume="106" year="1955">Vaurie 1955</bibRefCitation>
;
<bibRefCitation id="E26CA2B03919E236A0C1AFBB8156884D" DOI="10.1590/1809-43921978082299" author="Ratcliffe" firstAuthor="Ratcliffe" issue="2" journalOrPublisher="Acta Amazonica" pagination="299-302" refId="B88" refString="Ratcliffe BC (1978) A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae). Acta Amazonica 8 (2): 299–302. https://doi.org/10.1590/1809-43921978082299" title="A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae)." volume="8" year="1978">Ratcliffe 1978</bibRefCitation>
;
<bibRefCitation id="3BD12F4E2335DDB984C3098E12215059" DOI="10.1080/00222939000770841" author="Scholtz" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990</bibRefCitation>
;
<bibRefCitation id="87051E3FEB3C36EC99215F3BB81598A0" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOUBG" type="type material">
<paragraph id="BE8844FE7871EC2807F997761BEC1F62">
<heading id="18A11338F222CAF51E4F8ABC3BF32E3C" reason="title">Non-type examined material</heading>
</paragraph>
<paragraph id="C6BC3A7B7240225F4986F1E142FE5366">
<emphasis id="0B0D3BE00461F7F8A29D5152F111E821" bold="true">
(
<specimenCount id="A787E021C5872F4B8AAD64400CB89BA4" count="3" type="generic">three specimens</specimenCount>
additionally to
<bibRefCitation id="B4D9FED5D92F12ABEC5C038BE1609D0B" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021</bibRefCitation>
).
<materialsCitation id="557F80FCE7814425266C9F01D44EE1B7" country="Argentina" location="La Rioja" specimenCount="1" stateProvince="La Rioja">
<collectingCountry id="80F122F331AC5D2F72744133431352E1" name="Argentina">ARGENTINA</collectingCountry>
–
<emphasis id="A49E18AD2611174F13F881C5D748D7F6" bold="true">
<collectingRegion id="5119A9D8312CCFE17FB1019475F1C294">La Rioja</collectingRegion>
</emphasis>
</materialsCitation>
</emphasis>
•
<materialsCitation id="C4F22C57818968C39EA5F5C4B64C1B86" collectionCode="NHMUK" collectorName="Patquia &amp; K. J. Hayward" country="Argentina" specimenCount="1" specimenCount-female="1" stateProvince="La Rioja">
<specimenCount id="B50C35402E325CA72D80D3574E344022" count="1" type="female">1 ♀</specimenCount>
;
<collectorName id="12BF69E38120890AA62B2E5B37E741FC">Patquia</collectorName>
; 1933;
<collectorName id="05A31D4C41A21F02EDAAB9919F33D39C">K. J. Hayward</collectorName>
leg.;
<named-content id="A3FDF1608375F14445CA8B59B76630B7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="F539B92C8EB6A2CD1A7F5F76795450AA" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
</materialsCitation>
•
<materialsCitation id="EFC1595557E94B5D87A17A0CCA86BBCA" collectionCode="MNHN" collectorName="Bords du Rio Salado &amp; D'Icano &amp; Mistol &amp; E. R. Wagner" country="Argentina" location="Rio Salado" specimenCount="1" specimenCount-female="1" stateProvince="Santiago del Estero">
<specimenCount id="39C0C7C75A313CD6897B707CAF48CC47" count="1" type="female">1 ♀</specimenCount>
;
<collectingRegion id="8D82A7205117EF5C462021D7193D5F19" country="Argentina" name="Santiago del Estero">Santiago del Estero</collectingRegion>
,
<collectorName id="FF0609118B434B63CE70590A02AE6558">
Bords du
<location id="1325C7DB9231310E479C705B8258A465" LSID="urn:lsid:plazi:treatment:5A39B49D422A51EC89FA77D9D59DB106:1325C7DB9231310E479C705B8258A465" country="Argentina" name="Rio Salado" stateProvince="Santiago del Estero">Rio Salado</location>
</collectorName>
,
<collectorName id="ED313298A14B2214248DD5B385D92056">D’Icano</collectorName>
,
<collectorName id="470788EBA78E48FF9D99D7F0B1BA196E">Mistol</collectorName>
paso; 1909;
<collectorName id="FE74171DE3AD39FD7A006BCF03D2F8EC">E. R. Wagner</collectorName>
leg.;
<named-content id="758CC27BE319EAFB49F80DCDFCC53933" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire naturelle, Paris">
<collectionCode id="89C17B6C168D59C99A8B13BB84EB1630" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<paragraph id="A0455129CBE569633AC3B5F83673032B">
<emphasis id="A3013D352EFBF27D87D30D83EE197615" bold="true">
<collectingCountry id="B4DA4CCA6C4D9B6528B1DA65FA7F7D60" name="Brazil">BRAZIL</collectingCountry>
</emphasis>
• 1; Matto Grosso (sic!); 1886; P. Germain leg.;
<named-content id="98734EE5C3E0F300BF46F6E8AFC1705F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire naturelle, Paris">
<collectionCode id="A698A4B91C30F5AF7D85F38F8ABAAE2E" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
.
</paragraph>
</subSubSection>
</treatment>
</document>

659
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View file

@ -0,0 +1,659 @@
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<taxonomicName id="988ED48D112E19989234EB6128439372" ID-CoL="49DRB" authority="(Robinson, 1941)" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus" subGenus="Omorgus">
<emphasis id="D600B45D9B8164D8EBA9CE6C72C5B732" italics="true">Omorgus</emphasis>
(
<emphasis id="B909FE0FA626C7F663C5BC12B5A8C33F" italics="true">Omorgus</emphasis>
)
<emphasis id="B2E13B34926012655C74B1CE51CB3EE1" italics="true">fuliginosus</emphasis>
(
<bibRefCitation id="5FB702A6B7DF92E6F5CEC1D346B063FF" author="Robinson M" journalOrPublisher="Entomological News" pagination="134-135" refId="B94" refString="Robinson M (1941) A new species of Trox from Texas (Coleoptera: Scarabaeidae). Entomological News 52: 134 - 135." title="A new species of Trox from Texas (Coleoptera: Scarabaeidae)." volume="52" year="1941">Robinson, 1941</bibRefCitation>
)
</taxonomicName>
</paragraph>
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<subSubSection id="9F37447530F3CB7BA8EE8668C79276D5" type="description">
<paragraph id="73B11EABFF5D255BE3332BF5CDA7D9DF">
<figureCitation id="886FE705D28190DDE0095B9B42265799" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Trox fuliginosus Robinson, 1941 (now Omorgus fuliginosus). A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Robert Finn (USNM). Length of specimen: 14–15 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure11" httpUri="https://binary.pensoft.net/fig/1103873">Fig. 11</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="D803650B1F3B447DC4FB2BDB7C0F5720">
<treatmentCitationGroup id="F779CA53637A9D33C3AA06759C9878F6">
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<taxonomicName id="6295A037A78A711DBB507382877670CB" ID-CoL="59DFH" authority="Robison, 1941: 134" authorityName="Robinson" authorityPageNumber="134" authorityYear="1941" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus" subGenus="Omorgus">
<emphasis id="9F90AC47089F897E77B20ACAD280AB0F" italics="true">Trox</emphasis>
(
<emphasis id="D6018A49BB028A74DD7F96302B1E0949" italics="true">Omorgus</emphasis>
)
<emphasis id="9E78CA5ACF8463DE29C9D6E9BAAFAD29" italics="true">fuliginosus</emphasis>
<comment id="60DB313E4650289E8FE0AAA6090991F7">
Robison, 1941: 134 (original description);
<treatmentCitation id="B31259DFEC5E691932F807B4DC3DA8ED" author="Blackwelder RE &amp; Blackwelder RM" page="31" year="1948">
<bibRefCitation id="C4C2C8ABBA3E366959A04AFF1B1A0585" author="Blackwelder RE &amp; Blackwelder RM" firstAuthor="Blackwelder" journalOrPublisher="Vernon, New York" page="31" refId="B11" refString="Blackwelder RE, Blackwelder RM (1948) Fifth supplement to the Leng Catalogue of Coleoptera of America, north of Mexico. John D. Sherman; Mt. Vernon, New York, 87 pp." title="Fifth supplement to the Leng Catalogue of Coleoptera of America, north of Mexico. John D. Sherman; Mt." year="1948">Blackwelder and Blackwelder 1948: 31</bibRefCitation>
</treatmentCitation>
(checklist).
</comment>
</taxonomicName>
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</paragraph>
<paragraph id="16341BA662E28D9E9EECDC1EF3E58F1A">
<treatmentCitationGroup id="711200A9D18E7C4E65E0E11F47AC5693">
<treatmentCitation id="CD1CEF09A4C2803C0890C71992452AB8">
<taxonomicName id="3082087830595B1630EA33696DBA5EE5" ID-CoL="59DFH" authorityName="Robinson" authorityYear="1941" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="C820EDD87835DC690F4180D9CF935CF1" italics="true">Trox fuliginosus</emphasis>
</taxonomicName>
<comment id="9BFAE624F018E211FA1DA3A61FCCFAC9">
:
<treatmentCitation id="5B8A4C7C12191ECEA531ED94ADF3830B" author="Vaurie P" page="66" year="1955">
<bibRefCitation id="34CAEE766A063328BAB31F766A2A63BD" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" page="66" pagination="1-89" refId="B108" refString="Vaurie P (1955) A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). The Bulletin of the American Museum of Natural History 106: 1–89." title="A revision of the genus Trox in North America (Coleoptera: Scarabaeidae)." volume="106" year="1955">Vaurie 1955: 66</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="67D3951D3DF2F66B723A864995DB4BEA" author="Vaurie P" page="45" year="1958">
<bibRefCitation id="ADD687B5AEBC86F631156E78113AD368" DOI="10.5962/p.372226" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Coleopterists Bulletin" page="45" pagination="43-46" refId="B109" refString="Vaurie P (1958) New distribution records of North American Trox (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 12: 43–46. https://doi.org/10.5962/p.372226" title="New distribution records of North American Trox (Coleoptera: Scarabaeidae)." volume="12" year="1958">Vaurie 1958: 45</bibRefCitation>
</treatmentCitation>
(distribution); Blackwelder 1973: 35 (checklist);
<treatmentCitation id="144FE2432B8692766EFCC73146F91B58" author="Ratcliffe BC" page="301" year="1978">
<bibRefCitation id="A9262BB4D01D61D41FF45B168B6DBCDB" DOI="10.1590/1809-43921978082299" author="Ratcliffe BC" firstAuthor="Ratcliffe" issue="2" journalOrPublisher="Acta Amazonica" page="301" pagination="299-302" refId="B88" refString="Ratcliffe BC (1978) A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae). Acta Amazonica 8 (2): 299–302. https://doi.org/10.1590/1809-43921978082299" title="A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae)." volume="8" year="1978">Ratcliffe 1978: 301</bibRefCitation>
</treatmentCitation>
(distribution).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="42DCF0CEDE4BC102ECBF4513D3A81910">
<treatmentCitationGroup id="BC579E576EFD90D96E351CF15A9D58A9">
<treatmentCitation id="60B02B20336E481DF5CF4768E664087B">
<taxonomicName id="DDAF698ABD22DE0CD4D8CBCC28F1BE3C" ID-CoL="49DRB" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="C291800C30D667C9B1B7780845A0F5A4" italics="true">Omorgus fuliginosus</emphasis>
</taxonomicName>
<comment id="2D55657D75584DD721AE4A2BBB78450F">
:
<treatmentCitation id="95AAE5D44BAD8A69D10B93CE013FAAE2" author="Baker CW" page="43" year="1968">
<bibRefCitation id="1F00563BA2A9C89E0B56D204C6D623B5" DOI="10.5479/si.03629236.279.1" author="Baker CW" firstAuthor="Baker" issue="279" journalOrPublisher="Bulletin of the United States National Museum" page="43" pagination="1-79" refId="B5" refString="Baker CW (1968) Larval taxonomy of the Troginae in North America with notes on biologies and histories (Coleoptera: Scarabaeidae). Bulletin of the United States National Museum 279 (279): 1–79. https://doi.org/10.5479/si.03629236.279.1" title="Larval taxonomy of the Troginae in North America with notes on biologies and histories (Coleoptera: Scarabaeidae)." volume="279" year="1968">Baker 1968: 43</bibRefCitation>
</treatmentCitation>
(larvae description);
<treatmentCitation id="1BFC92C25DEA721C7D6EB50C31866E4F" author="Deloya C" page="3" year="1992">
<bibRefCitation id="10D061D1DB2FCCB6ED8B4D1B4030331B" DOI="10.21829/azm.1992.49521952" author="Deloya C" firstAuthor="Deloya" journalOrPublisher="Acta Zoologica Mexicana" page="3" pagination="1-13" refId="B29" refString="Deloya C (1992) Necrophilous Scarabaeidae and Trogidae beetles of tropical deciduous forest in Tepexco, Puebla, Mexico. Acta Zoologica Mexicana 52: 1–13. https://doi.org/10.21829/azm.1992.49521952" title="Necrophilous Scarabaeidae and Trogidae beetles of tropical deciduous forest in Tepexco, Puebla, Mexico." volume="52" year="1992">Deloya 1992: 3</bibRefCitation>
</treatmentCitation>
(natural history);
<treatmentCitation id="DA2387C8FC44CA701133FA8D10EEEA67" author="Deloya C" page="43" year="1996">
<bibRefCitation id="0A8F960DFEACC10A9FBF13AE0261645D" author="Deloya C" firstAuthor="Deloya" journalOrPublisher="Folia Entomologica Mexicana" page="43" pagination="39-54" refId="B30" refString="Deloya C (1996) Los Macro-Coleopteros necrofilos de Tepoztlan, Morelos, Mexico (Scarabaeidae, Trogidae, Silphidae). Folia Entomologica Mexicana 97: 39–54." title="Los Macro-Coleopteros necrofilos de Tepoztlan, Morelos, Mexico (Scarabaeidae, Trogidae, Silphidae)." volume="97" year="1996">Deloya 1996: 43</bibRefCitation>
</treatmentCitation>
(natural history);
<treatmentCitation id="F2B314802FB346B660061857B29C7EB9" author="Morón MA" page="91" year="1998">
<bibRefCitation id="7715709B316D87BA4359A15BEC06DACA" DOI="10.21829/azm.1998.75751708" author="Morón MA" journalOrPublisher="Acta Zoologica Mexicana" pagination="73-116" refId="B73" refString="Moron MA (1998) Fauna de Coleoptera Lamellicornia de la region de Tepic, Nayarit, Mexico. Acta Zoologica Mexicana 75: 73 - 116 https: // doi. org / 10.21829 / azm. 1998.75751708." title="Fauna de Coleoptera Lamellicornia de la región de Tepic, Nayarit, Mexico." volume="75" year="1998">Morón et al. 1998: 91</bibRefCitation>
</treatmentCitation>
(distribution);
<treatmentCitation id="1ED4B97531AE12E45AB64776C4E7F7AC" author="Deloya C" page="66" year="2000">
<bibRefCitation id="3E12D0E45BF523708B9828A25C6EF3B5" DOI="10.21829/azm.2000.81811871" author="Deloya C" firstAuthor="Deloya" journalOrPublisher="Acta Zoológica Mexicana" page="66" pagination="63-76" refId="B31" refString="Deloya C (2000) Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia). Acta Zoológica Mexicana 81: 63–76. https://doi.org/10.21829/azm.2000.81811871" title="Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia)." volume="81" year="2000">Deloya 2000: 66</bibRefCitation>
</treatmentCitation>
, 73 (checklist);
<treatmentCitation id="375F27797B09F04F8896DE04BAC44027" author="Smith ABT" page="6" year="2003">
<bibRefCitation id="87F09841D854A6D62ACD1FC95A4541DB" author="Smith ABT" firstAuthor="Smith" journalOrPublisher="Electronically published, Lincoln, Nebraska" page="6" refId="B101" refString="Smith ABT (2003) Checklist of the Scarabaeoidea of the Nearctic Realm. Version 3. Electronically published, Lincoln, Nebraska, 74 pp." title="Checklist of the Scarabaeoidea of the Nearctic Realm. Version 3." year="2003">Smith 2003: 6</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="15EA1F37A69D053F22A302A85D613254" author="Deloya C" page="130" year="2003">
<bibRefCitation id="F920DCD97C60EE16E1C86DC0C79467A4" author="Deloya C" editor="Morón MA" firstAuthor="Deloya" journalOrPublisher="Vol II. Barcelona" page="130" pagination="125-133" refId="B32" refString="Deloya C (2003) Familia Trogidae. In: Morón MA (Ed.) Atlas de los escarabajos de México. Vol II. Barcelona, 125–133." title="Familia Trogidae." volumeTitle="Atlas de los escarabajos de México." year="2003">Deloya 2003: 130</bibRefCitation>
</treatmentCitation>
(diagnosis and distribution);
<treatmentCitation id="68C69A686B377444103D19E248AB0445" author="Deloya C" page="122" year="2005">
<bibRefCitation id="5B9D5A03C97D71493DD59E7143E2A1B8" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121–129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
, 124, 128 (key);
<treatmentCitation id="5248209F2E0B52F9187E162EA1ECAADD" author="Mora-Aguilar EF &amp; Montes de Oca E" page="575" year="2009">
<bibRefCitation id="9B77404E7CDC5568FEB5990BEAA2517D" DOI="10.21829/azm.2009.253662" author="Mora-Aguilar EF &amp; Montes de Oca E" firstAuthor="Mora-Aguilar" issue="3" journalOrPublisher="Acta Zoológica Mexicana (nueva serie)" page="575" pagination="569-588" refId="B69" refString="Mora-Aguilar EF, Montes de Oca E (2009) Escarabajos necrófagos (Coleoptera: Scarabaeidae y Trogidae) de la región central baja de Veracruz, México. Acta Zoológica Mexicana (nueva serie) 25 (3): 569–588. https://doi.org/10.21829/azm.2009.253662" title="Escarabajos necrófagos (Coleoptera: Scarabaeidae y Trogidae) de la región central baja de Veracruz, México." volume="25" year="2009">Mora-Aguilar and Montes de Oca 2009: 575</bibRefCitation>
</treatmentCitation>
(distribution).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="4133E2F45F3789249B3F23D4A57A9454">
<treatmentCitationGroup id="ED1E23F98B6AD5E497F97A815D1E71EC">
<treatmentCitation id="6DC094B5C74761FB1D9FED79197CA6E0">
<taxonomicName id="E358A35A887F67C9D01B5A59B25D0413" ID-CoL="49DRB" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus" subGenus="Omorgus">
<emphasis id="450FA151DE885B01CFCED4379C80D009" italics="true">Omorgus</emphasis>
(
<emphasis id="617A642B0A6F5C60CE97DE84F5A28EE6" italics="true">Omorgus</emphasis>
)
<emphasis id="D85F0FECAD9751403241D8F829E5B9FF" italics="true">fuliginosus</emphasis>
</taxonomicName>
<comment id="5C484BBB0636DA11F34FD986252D7658">
:
<treatmentCitation id="7C05F32AC04B9AFF2EDC4D4A2BE1245F" author="Zidek J" page="10" year="2013">
<bibRefCitation id="559DF89A988B03C25BF66DF53589AFD3" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="10" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1–38." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 10</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="064892EE877308CC8894008E03C8BB44" author="Zidek J" page="101" year="2017">
<bibRefCitation id="F59CF943BDE01694774CCE0582C34A42" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="101" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 101</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="675C4D262F61C53C0E9CC74629754E75" author="Pablo-Cea JD &amp; Cave RD &amp; Serrano-Peraza FA &amp; Alvarado-Larios R &amp; Deloya C &amp; Serrano-Chicas KA &amp; Alfaro E &amp; Chinchilla-Rodríguez AC &amp; Girón-Segovia D &amp; Noriega JA" page="16" year="2023">
<bibRefCitation id="B96EE450E29B16E25ADEE312491B1955" DOI="10.22201/ib.20078706e.2023.94.5117" author="Pablo-Cea JD &amp; Cave RD &amp; Serrano-Peraza FA &amp; Alvarado-Larios R &amp; Deloya C &amp; Serrano-Chicas KA &amp; Alfaro E &amp; Chinchilla-Rodríguez AC &amp; Girón-Segovia D &amp; Noriega JA" etAl="et al." firstAuthor="Pablo-Cea" page="16" refId="B79" refString="Pablo-Cea JD, Cave RD, Serrano-Peraza FA, Alvarado-Larios R, Deloya C, Serrano-Chicas KA, Alfaro E, Chinchilla-Rodríguez AC, Girón-Segovia D, Noriega JA (2023) Catálogo y atlas de distribución de los Scarabaeoidea (Insecta: Coleoptera) de El Salvador. Revista Mexicana de Biodiversidad 94: e 945117. https://doi.org/10.22201/ib.20078706e.2023.94.5117" year="2023">Pablo-Cea et al. 2023: 16</bibRefCitation>
</treatmentCitation>
(catalogue of
<collectingCountry id="6A4BC205F903AA5C60EFD184A91D8947" name="El Salvador">El Salvador</collectingCountry>
).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E91AF09F3847DD5BE5163D1C84035845">
<treatmentCitationGroup id="21CF9227D0E1B6A7CA91A98DBC6A5255">
<treatmentCitation id="1B28C6D80B4FDAB3E31E5E7CF1AEA970" author="Scholtz CH" page="1413" year="1990">
<taxonomicName id="DE6A31F89F7BE066E1B1D81C6535A4C4" ID-CoL="74LV9" authority="Scholtz, 1990: 1413" authorityName="Scholtz" authorityPageNumber="1413" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus" subGenus="Omorgus">
<emphasis id="39A3D2DE75F6081F0711A88F7449A878" italics="true">Omorgus</emphasis>
(
<emphasis id="8E503C37727E1BA340258C4D4B23EC2E" italics="true">Omorgus</emphasis>
)
<emphasis id="F814A9F9F724379623C85AA81F1624FA" italics="true">capillaceus</emphasis>
<comment id="D5410E73A0480168C0665F552B4AAFB7">
<bibRefCitation id="5F45B569DF0D394B1F47C88424281BC4" DOI="10.1080/00222939000770841" author="Scholtz CH" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391 - 1456. https: // doi. org / 10.1080 / 00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz, 1990: 1413</bibRefCitation>
(new synonym):
<treatmentCitation id="59CC80419FD9ADDD97D7F7FC8A61A071" author="Deloya C" page="122" year="2005">
<bibRefCitation id="83DEB05824532C232072F37DA46E3925" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121–129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="CF1EE051A1BBA36187DB3402AFB55692" author="Zidek J" page="8" year="2013">
<bibRefCitation id="CDC5D1E30779CD77B1E82079B04FFC60" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="8" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1–38." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 8</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="7FEBBBE659526E65DA4A756EE0061614" author="Zidek J" page="99" year="2017">
<bibRefCitation id="62340DCCF13B6F2D9443015FA7E39E11" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="99" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 99</bibRefCitation>
</treatmentCitation>
(checklist).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDCDI" type="Type specimen examined">
<paragraph id="A8FA490D5B9E555F91B622B36BEF0E0A">
<heading id="033B5CB0064030A8791A3C8E0E887546" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="C497237C5EB0204EFD923B631FA13309">
<materialsCitation id="F7536AC93FC5802550AEB919961C7766" collectionCode="USNM" collectorName="H. Mittendorf" country="United States of America" location="Sixth" specimenCount="1" specimenCount-male="1" stateProvince="Texas" typeStatus="holotype">
<emphasis id="AE8BA7E92710129A1942006B30966201" bold="true" italics="true">
<typeStatus id="ED4D93327F36D57C28FFA38F509BAB52" type="holotype">Holotype</typeStatus>
, by original designation
</emphasis>
(
<specimenCount id="9697EE6574827F25A98722C847CF97A9" count="1" type="male">♂</specimenCount>
<emphasis id="3A6C7B21D625791EAC89A014587BD693" bold="true">
<collectionCode id="770A20050779BE417C844D4CD756545B" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</emphasis>
– Fig.
<figureCitation id="0CB5A403F195275E5179ED58D75AD4D9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Trox fuliginosus Robinson, 1941 (now Omorgus fuliginosus). A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Robert Finn (USNM). Length of specimen: 14–15 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure11" httpUri="https://binary.pensoft.net/fig/1103873">11</figureCitation>
). First label [white aged, typeset]: “ New Braunfels / IV. 10 ’ 02 Tex ”. Second label [white aged, typeset]: “
<collectorName id="CB46CA0CA779C50668F63E5A53BA6513">H. Mittendorf</collectorName>
/ Collector ”. Third label [red, printed]: “ Type No / 55468 / USNM ”. Fourth label [red, Mark Robinson’s handwriting]: “
<typeStatus id="7AF7928EB9A48762C2C1F9D4E124F885" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="7141587E841E097D7ED6680D9A935E83" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="5B834B6B71D17962336046C48146E863" italics="true">Trox</emphasis>
</taxonomicName>
/
<taxonomicName id="2CF313D43A80EDCC280F437EE44D84BF" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="418426C5A6B6CBF3729B53BD3D1335DC" italics="true">fuliginosus</emphasis>
</taxonomicName>
/ Mark Robinson ”.
<location id="B80FAA717C7E776FCC5081025B0CD742" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:B80FAA717C7E776FCC5081025B0CD742" country="United States of America" name="Sixth" stateProvince="Texas">Sixth</location>
label [white, printed]: “ USNMENT / [QR Code] / 01474112 ” (Fig.
<figureCitation id="48BD05307ED7ED56A60EDB93B75EF080" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Trox fuliginosus Robinson, 1941 (now Omorgus fuliginosus). A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Robert Finn (USNM). Length of specimen: 14–15 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure11" httpUri="https://binary.pensoft.net/fig/1103873">11 C</figureCitation>
).
<emphasis id="07522E0CA1A3A84352DD797D9076F291" bold="true">Type locality</emphasis>
: “ New Braunfels ” [
<collectingRegion id="10506A673E9F14A2CCBBF2B312D54860" country="United States of America" name="Texas">Texas</collectingRegion>
,
<collectingCountry id="6B961C8B2BE6DB7F5CFC2C3844C861C6" name="United States of America">United States</collectingCountry>
].
</materialsCitation>
</paragraph>
<caption id="F373B4943A79520D80A01D58F8EB89D2" ID-DOI="10.3897/contrib.entomol.74.e126799.figure11" ID-arpha="501E6441-5C5E-55B5-ACF7-6A376FA85A20" httpUri="https://binary.pensoft.net/fig/1103873" startId="F11">
<paragraph id="80E5A2120E2B6BDC056EBF1D228F0BBC">
<label id="2EABC1A37259379E41ED521CE091EC3A">Figure 11.</label>
</paragraph>
<paragraph id="D0AD4436B835411D2A1384E652AEAD72">
Holotype of
<taxonomicName id="8D16551ABB2BA8FFBB2E1037501A2B4F" authority="Robinson, 1941" authorityName="Robinson" authorityYear="1941" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="E459BB307AE4A5CFF1D9D3340A558216" italics="true">Trox fuliginosus</emphasis>
<bibRefCitation id="DC11276DD551FA3502609B73553A6EEB" author="Robinson M" journalOrPublisher="Entomological News" pagination="134-135" refId="B94" refString="Robinson M (1941) A new species of Trox from Texas (Coleoptera: Scarabaeidae). Entomological News 52: 134 - 135." title="A new species of Trox from Texas (Coleoptera: Scarabaeidae)." volume="52" year="1941">Robinson, 1941</bibRefCitation>
</taxonomicName>
(now
<taxonomicName id="22024A3DDDABB2F4D4352BF43F86D5F8" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="8FC446D3B77E730C2FC7E11EB189A592" italics="true">Omorgus fuliginosus</emphasis>
</taxonomicName>
).
<emphasis id="32B66BDFA8D2B0A78E19C5481E0EFD8F" bold="true">A.</emphasis>
Habitus in dorsal view;
<emphasis id="C8234EBC7316D0805370C53C3C32D69A" bold="true">B.</emphasis>
Habitus in lateral view;
<emphasis id="924BAAEFD2DF99ADD18F1F4A61431328" bold="true">C.</emphasis>
Labels. Photos by Robert Finn (USNM). Length of specimen: 14–15 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ESEDI" type="Type specimen">
<paragraph id="085CA7F9DF1F5720CC265FD97D92F357">
<heading id="5F1D01B51940F5F0CFE9DEFC3E933BD6" reason="title">Type specimen</heading>
</paragraph>
<paragraph id="8DF07285A4D2BAB5FBAB34EC6465009B">
<emphasis id="CEFF01AD41AA7DA97000656C7C232335" bold="true">
of
<taxonomicName id="F2007180F79436EB420A96F1988E0616" authority="Scholtz, 1990" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus" subGenus="Omorgus">
<emphasis id="E6A52333762217A00E55DDA42BE68F5B" italics="true">Omorgus</emphasis>
(
<emphasis id="0DD6FF2B872951EA744D2B4D8D6E1D43" italics="true">Omorgus</emphasis>
)
<emphasis id="9A8818486178091EAB0482E0464E3627" italics="true">capillaceus</emphasis>
<bibRefCitation id="8F5491FA72DFE58F8DB0A41DC2DFC2D1" DOI="10.1080/00222939000770841" author="Scholtz CH" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391 - 1456. https: // doi. org / 10.1080 / 00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz, 1990</bibRefCitation>
</taxonomicName>
(new synonym) examined.
<materialsCitation id="48DC9176FDD71CF6F7FF7E21F1E18396" collectionCode="CMNCH" collectorName="S. Amer. &amp; C. H. Scholtz" country="Colombia" location="Fourth" specimenCount="1" specimenCount-female="1" stateProvince="Bogota" typeStatus="holotype">
<emphasis id="B8368ECDD369205D131F2EC09FDE9083" bold="true" italics="true">
<typeStatus id="883021CEA95F686A314A23DD8113B966" type="holotype">Holotype</typeStatus>
, by original designation
</emphasis>
(
<specimenCount id="F7BFF917E741CE8881CEDF44C0A7B0CD" count="1" type="female">♀</specimenCount>
<named-content id="E7AFA3FEE5D06D65D24DC045143D33D0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<emphasis id="4B9D5A5FD562D3DEBFED861EF71B728E" bold="true">
<collectionCode id="B4820C5A3ADBCEF65DA7577259B4BA8E">CMNCH</collectionCode>
</emphasis>
</named-content>
– Fig.
<figureCitation id="4C8874A3288CC6B97908EED735477410" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Holotype of Omorgus capillaceus Scholtz, 1990. A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Vanessa Verdecia (CMNCH)." figureDoi="10.3897/contrib.entomol.74.e126799.figure12" httpUri="https://binary.pensoft.net/fig/1103874">12</figureCitation>
). First label [white aged, unknown’s handwriting]: “ Botoga /
<collectingCountry id="6D58672C89B9C3EA4311378C193D8008" name="Colombia">Colombia</collectingCountry>
/
<collectorName id="EDAAB97787EE26D64EA1353760990B10">S. Amer.</collectorName>
”. Second label [white, typeset]: “ Carn. Mus. / Acc. 2275 ”. Third label [white, printed]: “ [QR code] /
<named-content id="73A5D7CBAF0F1877C4E4D3BDA1B0F6ED" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="30BC601B32B894F0CD33ADAABF1F6FBC">CMNCH</collectionCode>
</named-content>
- IZ / 724,596 ”.
<location id="2DA896EAAC6A899D35099C1875FFEA27" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:2DA896EAAC6A899D35099C1875FFEA27" country="Colombia" name="Fourth" stateProvince="Bogota">Fourth</location>
label [white with red border, Clark Scholtz`s handwriting]: “
<taxonomicName id="B8D09DB0365514C37336F871ACDCBE37" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="DF0C1F6D72D3180226A7F721C81C0291" italics="true">Omorgus</emphasis>
</taxonomicName>
/
<taxonomicName id="0F70278FEFCDA2EACAEE63895081BB14" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="2D9C2B86986F1E1A9A845621FF11BE0C" italics="true">capillaceus</emphasis>
</taxonomicName>
/
<collectorName id="D2412C0F3B0FFF255E2F01E915CCCC65">C. H. Scholtz</collectorName>
1988 /
<typeStatus id="0D8031F404283168C7C07BBDCD668B08" type="holotype">HOLOTYPE</typeStatus>
” (Fig.
<figureCitation id="5ACD02FE10938FEBCE598637C1610CAF" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Holotype of Omorgus capillaceus Scholtz, 1990. A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Vanessa Verdecia (CMNCH)." figureDoi="10.3897/contrib.entomol.74.e126799.figure12" httpUri="https://binary.pensoft.net/fig/1103874">12 C</figureCitation>
).
<emphasis id="6DE3B9795723AC4D336A712A6240B8F9" bold="true">Type locality</emphasis>
: “
<collectingRegion id="6B41E2B77482EDB0B33D7C8C48911669" country="Colombia" name="Bogota">Bogota</collectingRegion>
,
<collectingCountry id="C64ED4442F71B1345E8548FB2957A5E7" name="Colombia">Colombia</collectingCountry>
”.
</materialsCitation>
</emphasis>
</paragraph>
<caption id="068F04ADF8231B00E5C3222E7BAD63DA" ID-DOI="10.3897/contrib.entomol.74.e126799.figure12" ID-arpha="C4BA82C9-09BB-5DA5-904B-9D35C6AC0731" httpUri="https://binary.pensoft.net/fig/1103874" startId="F12">
<paragraph id="D1FD68A640EB877329036A85321D5940">
<label id="582B1C0C57DF10D4D7412ACCDC6A407F">Figure 12.</label>
</paragraph>
<paragraph id="40AC9FDE60642CA4B8F1CAB0DCDC5458">
Holotype of
<taxonomicName id="A07ADB579B7FFE6B2B355DDAFEA40CCA" authority="Scholtz, 1990" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="CD4311521E25F548FCB3A553427B483F" italics="true">Omorgus capillaceus</emphasis>
<bibRefCitation id="EB982361ABF609226DA2FAC292BE8ADA" DOI="10.1080/00222939000770841" author="Scholtz CH" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391 - 1456. https: // doi. org / 10.1080 / 00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz, 1990</bibRefCitation>
</taxonomicName>
.
<emphasis id="07E018B19EA5812E8F993C37C331AAAB" bold="true">A.</emphasis>
Habitus in dorsal view;
<emphasis id="C647F0F4146EC6B950A5270E838DE2C5" bold="true">B.</emphasis>
Habitus in lateral view;
<emphasis id="1AD08413B9E1EBFA910371D625C88788" bold="true">C.</emphasis>
Labels. Photos by Vanessa Verdecia (
<named-content id="B3153F2F62DA83887D1B25C7BFAF9233" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">CMNCH</named-content>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EOHDI" type="remarks">
<paragraph id="F599E736ABF2F06559A481CBC2ECF660">
<heading id="A63428702B1081FCEF062A921B32E19B" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="A3696EC91D1EE41DBCFB275617CFA775">
<taxonomicName id="7A21DE2DCF82030BE386BA5BB1F0147D" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus" subGenus="Omorgus">
<emphasis id="23164DF7CA61E242B2D44B05B7AC2301" italics="true">Omorgus</emphasis>
(
<emphasis id="C3AB6A93E3FD3E82E5684421682D5AF8" italics="true">Omorgus</emphasis>
)
<emphasis id="7F9863CADF83C7F829F066769686470B" italics="true">capillaceus</emphasis>
</taxonomicName>
was one of the species described by Scholtz in 1990 during his monographic revision of the
<taxonomicName id="0CDA72476D6DF1759F4EC58A52A9D7CA" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
of South America. This species was identified in the collection of the Carnegie Museum of Natural History (
<named-content id="98595F500B6BE4EEFF59FD9DDDE95B28" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="00AC3575AEEEB6BFAC4329EE496BEABB">CMNCH</collectionCode>
</named-content>
) and described based on a single female specimen, distinguished by its morphological differences from other South American species. Scholtz noted that
<taxonomicName id="E4B7DFB8FB808F99C13AA759D032E1D0" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="2E07CACB1CCF2E531D346A76237696AA" italics="true">O. capillaceus</emphasis>
</taxonomicName>
was similar to
<taxonomicName id="0B516331FED6E2BA05268ABFD759E50D" authority="(Herbst, 1790)" baseAuthorityName="Herbst" baseAuthorityYear="1790" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="monachus">
<emphasis id="660E2A9C35E9D68D17F38D970D961E23" italics="true">O. monachus</emphasis>
(
<bibRefCitation id="A45F1A4157376AFDDBB901C0655BA6A8" author="Herbst JFW" refId="B52" refString="Herbst JFW (1790) [Hefte 1: 1 - 151, 6 pls.]. In: Herbst JFW (Ed.) 1790 - 1791. Natursystem aller bekannten in- und auslandischen Insekten, als eine Forsetzung der von Buffonschen Naturgeschichte. Der Kafer Dritter Theil. Berlin: Joachim Pauli; p. xiv + 325 pp. + pls. 21 - 34. D - E [publication date following Bousquet 2016]." year="1790">Herbst, 1790</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="8B34402EACB0F1D950C9FFEC05112ABA" authority="(Robinson, 1941)" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="3F2BC18FD139F87770D5950293D778E8" italics="true">O. fuliginosus</emphasis>
(
<bibRefCitation id="8AC3596EDCAFC7CF65FD0E9ED53704E1" author="Robinson M" journalOrPublisher="Entomological News" pagination="134-135" refId="B94" refString="Robinson M (1941) A new species of Trox from Texas (Coleoptera: Scarabaeidae). Entomological News 52: 134 - 135." title="A new species of Trox from Texas (Coleoptera: Scarabaeidae)." volume="52" year="1941">Robinson, 1941</bibRefCitation>
)
</taxonomicName>
, both of which are North American species. However, he highlighted a diagnostic feature that distinguishes
<taxonomicName id="2EF4DF044ADBC6B439041B5F6A7D79C4" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="98721404D7FAB6E01F6708DAEAF46132" italics="true">O. capillaceus</emphasis>
</taxonomicName>
from other similar-looking species: the absence of a velutinous covering on the elytral disc, which is present in its relatives.
</paragraph>
<paragraph id="852990D6BF2D38B13B487A94558E2DC6">
Upon reviewing the
<typeStatus id="8CF5EBC2BA6A7DEAA461B7828DDAB522" type="holotype">holotype</typeStatus>
of
<taxonomicName id="9865ADA5272FD4113069D6F90D32CC27" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="D4C2DC64460109E2012692029171815A" italics="true">O. capillaceus</emphasis>
</taxonomicName>
, it became evident that the specimen is merely a worn variation of
<taxonomicName id="986BD6DADD87C05A1E1ADF02FC08D914" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="44421D089AE9D901AFE279088BECF7B7" italics="true">O. fuliginosus</emphasis>
</taxonomicName>
. While we concur with Scholtz’s observation of the glabrous elytral disc, this feature alone is insufficient for distinguishing
<taxonomicName id="0482BBAC516781DE136F3D447287D0A6" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
species without considering male genitalia.
<taxonomicName id="68FB1733E408148D6F2D35DADAF91C2C" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
specimens often exhibit significant intraspecific variation, and it is not uncommon to encounter specimens that are partially or entirely glabrous (or worn) in collections (V. Costa-Silva, personal observation).
</paragraph>
<paragraph id="F87DD5CFE1E26860D82174AE490144CF">
Another factor that may have led Scholtz to describe a new species is the specimen’s locality:
<collectingRegion id="AE2E4BC114DEF1698E36DB9D47342147" country="Colombia" name="Bogota">Bogota</collectingRegion>
,
<collectingCountry id="6871F1A2B6FAA344A3267025946F5229" name="Colombia">Colombia</collectingCountry>
. The
<typeStatus id="7C289DA8D6E29C9C3282EAB806343FDC" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3DC14AC5C9891BD893F5CC2454FBEB5C" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="A9155788D26B558C5EC5689A8E426E1D" italics="true">O. capillaceus</emphasis>
</taxonomicName>
was associated with the label number “ 2275 ” (Fig.
<figureCitation id="A30672AEFDA736B3D80C066624DBBC8C" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Holotype of Omorgus capillaceus Scholtz, 1990. A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Vanessa Verdecia (CMNCH)." figureDoi="10.3897/contrib.entomol.74.e126799.figure12" httpUri="https://binary.pensoft.net/fig/1103874">12 C</figureCitation>
), which corresponds to specimen from the Henry Klages Collection, acquired by the
<named-content id="E0C8FD4A64F7DE0A61DCB1253E035AD8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="3277DBCB682A4F31965C4E2AFBCB6067">CMNCH</collectionCode>
</named-content>
on
<date id="6B2E1182D3507FF83EAE8320D0C46CB9" value="1903-05-28">May 28, 1903</date>
(Robert Androw, personal communication to
<collectionCode id="46750F8B1A7F108F413FA51A793D8869">VCS</collectionCode>
). However, as documented by Nearns and Androw (2013) and
<bibRefCitation id="D8BF995EFE0E8B7EDCE84F8ED2AD0DD8" DOI="10.1080/00222933.2023.2260060" author="Costa-Silva V &amp; Strümpher WP &amp; Thyssen PJ &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="1-4" journalOrPublisher="Journal of Natural History" pagination="14-166" refId="B27" refString="Costa-Silva V, Strümpher WP, Thyssen PJ, Vaz-de-Mello FZ (2024) Taxonomic revision of the South American genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae). Journal of Natural History 58 (1–4): 14–166. https://doi.org/10.1080/00222933.2023.2260060" title="Taxonomic revision of the South American genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae)." volume="58" year="2024">Costa-Silva et al. (2024)</bibRefCitation>
, the Henry Klages collection has been noted for containing numerous mislabelled Nearctic and Neotropical specimens, including some cases of misidentification within the
<taxonomicName id="D544F5A20FAA36C48A93D96848D4FCB2" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
(see
<bibRefCitation id="BA6E8BBF1F9403337CB85D301A69C714" DOI="10.1080/00222933.2023.2260060" author="Costa-Silva" etAl="et al." firstAuthor="Costa-Silva" issue="1-4" journalOrPublisher="Journal of Natural History" pagination="14-166" refId="B27" refString="Costa-Silva V, Strümpher WP, Thyssen PJ, Vaz-de-Mello FZ (2024) Taxonomic revision of the South American genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae). Journal of Natural History 58 (1–4): 14–166. https://doi.org/10.1080/00222933.2023.2260060" title="Taxonomic revision of the South American genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae)." volume="58" year="2024">Costa-Silva et al. 2024</bibRefCitation>
for examples).
<taxonomicName id="72B1051E63F7E3E1C462A88809402D4B" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="F1AD4BDA4664E26A31FD3DB91DC3E0A6" italics="true">Omorgus fuliginosus</emphasis>
</taxonomicName>
is known from the
<collectingCountry id="D8F07AF3D92805EFAF799B79DA0A231B" name="United States of America">United States</collectingCountry>
,
<collectingCountry id="18AEEB233088EA723093943EDB195ABA" name="Mexico">Mexico</collectingCountry>
,
<collectingCountry id="E22403128B46AEF8DF17B0C2D894B5C8" name="Guatemala">Guatemala</collectingCountry>
,
<collectingCountry id="AFBA156B066F24DB051DBC1C22601055" name="El Salvador">El Salvador</collectingCountry>
, and
<collectingCountry id="619C99181F10885E917FD0A3FEDE62D1" name="Costa Rica">Costa Rica</collectingCountry>
(
<bibRefCitation id="8147639BCB7EA952A9D4DA3E99BD4B14" author="Vaurie" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="1-89" refId="B108" refString="Vaurie P (1955) A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). The Bulletin of the American Museum of Natural History 106: 1–89." title="A revision of the genus Trox in North America (Coleoptera: Scarabaeidae)." volume="106" year="1955">Vaurie 1955</bibRefCitation>
;
<bibRefCitation id="0F2205092994A9C33D9148D88B699D48" DOI="10.1590/1809-43921978082299" author="Ratcliffe" firstAuthor="Ratcliffe" issue="2" journalOrPublisher="Acta Amazonica" pagination="299-302" refId="B88" refString="Ratcliffe BC (1978) A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae). Acta Amazonica 8 (2): 299–302. https://doi.org/10.1590/1809-43921978082299" title="A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae)." volume="8" year="1978">Ratcliffe 1978</bibRefCitation>
;
<bibRefCitation id="D85FD5DBE127594B8A7E6595B04A2672" DOI="10.21829/azm.2000.81811871" author="Deloya" firstAuthor="Deloya" journalOrPublisher="Acta Zoológica Mexicana" pagination="63-76" refId="B31" refString="Deloya C (2000) Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia). Acta Zoológica Mexicana 81: 63–76. https://doi.org/10.21829/azm.2000.81811871" title="Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia)." volume="81" year="2000">Deloya 2000</bibRefCitation>
), making its record in
<collectingCountry id="5FD227E750995849612A577B86E5D1CD" name="Colombia">Colombia</collectingCountry>
questionable, but not impossible.
</paragraph>
<paragraph id="65442283DD62A1C46D8BE36FF2D36A5E">
Therefore, we conclude that
<taxonomicName id="26B1F2C929E3EF537CC47F08E9C2AC0E" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="4F4C8D36E8CC41B10317139F4D14EEAD" italics="true">O. capillaceus</emphasis>
</taxonomicName>
should be considered a junior subjective synonym of
<taxonomicName id="7D7D33F1995E459280A8EEF5D3E89011" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="DCE33F6A6DA30BB380C459A4544C4A16" italics="true">O. fuliginosus</emphasis>
</taxonomicName>
, having been described based on a worn and mislabelled specimen from
<collectingRegion id="2089C1A901B7BB8DEA6DB2D7E9EB0F49" country="Colombia" name="Bogota">Bogotá</collectingRegion>
,
<collectingCountry id="E7A495306FD6A7F5019E16B36F2F53A7" name="Colombia">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKNDI" type="distribution">
<paragraph id="004672A8451E0A1D06BBD61127EAD9A0">
<heading id="4C71708D5B20ECCD0C4EE97D38E38333" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="E04EDA7A1D6C1229FF55C303F0AAFBC2">
<collectingCountry id="D916E9F61666CDD870BF0906EADA23B6" name="Costa Rica">Costa Rica</collectingCountry>
,
<collectingCountry id="5D96742922CAFCA2392C86299CCFBDDE" name="El Salvador">El Salvador</collectingCountry>
,
<collectingCountry id="66000913A028F66DB44DB714A50795F9" name="Guatemala">Guatemala</collectingCountry>
,
<collectingCountry id="EAD8CA554E2E89C22934EE5F53B01BAD" name="Mexico">Mexico</collectingCountry>
, and
<collectingCountry id="0BC252DE50D033074241FB82A2C8AAC5" name="United States of America">United States</collectingCountry>
(
<bibRefCitation id="FEE08A355946DBED1BCF3036EF8F68FC" author="Vaurie" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="1-89" refId="B108" refString="Vaurie P (1955) A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). The Bulletin of the American Museum of Natural History 106: 1–89." title="A revision of the genus Trox in North America (Coleoptera: Scarabaeidae)." volume="106" year="1955">Vaurie 1955</bibRefCitation>
;
<bibRefCitation id="91CF2C5CDFD10B0BE1E57DF1C2C9C731" DOI="10.1590/1809-43921978082299" author="Ratcliffe" firstAuthor="Ratcliffe" issue="2" journalOrPublisher="Acta Amazonica" pagination="299-302" refId="B88" refString="Ratcliffe BC (1978) A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae). Acta Amazonica 8 (2): 299–302. https://doi.org/10.1590/1809-43921978082299" title="A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae)." volume="8" year="1978">Ratcliffe 1978</bibRefCitation>
;
<bibRefCitation id="96E2C707C6FF3A869C815FFD64E5B41E" DOI="10.21829/azm.2000.81811871" author="Deloya" firstAuthor="Deloya" journalOrPublisher="Acta Zoológica Mexicana" pagination="63-76" refId="B31" refString="Deloya C (2000) Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia). Acta Zoológica Mexicana 81: 63–76. https://doi.org/10.21829/azm.2000.81811871" title="Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia)." volume="81" year="2000">Deloya 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3NDI" type="type material">
<paragraph id="FA547A1C25E107F9BC1411A48B00C47F">
<heading id="A17C5D3483139778E7DC6AF4A38A6B5C" reason="title">Non-type examined material</heading>
</paragraph>
<paragraph id="1E0964619D34288EEF9A1E7FCE07D272">
<emphasis id="6DC4655556F8296CACDB73B73BCB7994" bold="true">
(
<specimenCount id="1409DE641938E8EA40D593D777DD1B0A" count="9" type="generic">9 specimens</specimenCount>
).
<materialsCitation id="58F0800151F8174B490E7645D00EA025" country="Mexico" location="Veracruz" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<collectingCountry id="18D0C04E1C931C585C0E93AA15DE4776" name="Mexico">MEXICO</collectingCountry>
–
<emphasis id="369E7A3F1D399C31C37870B7F989274F" bold="true">
<collectingRegion id="C3704BD4BC0590F68807EDC13D741799" country="Mexico" name="Veracruz de Ignacio de la Llave">Veracruz</collectingRegion>
</emphasis>
</materialsCitation>
</emphasis>
•
<materialsCitation id="3AC181AF02E4638C9B92B5931660FC71" collectingDate="1994-08-30" collectionCode="CEMT" collectorName="L. Arellano &amp; R. Sanchez" country="Mexico" location="Passando la Desv. Idolos" municipality="Carr. Actopan" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
5;
<collectingMunicipality id="58D12BB5986986F0D15E0987F1990FC3">Carr. Actopan</collectingMunicipality>
,
<location id="23D4E8AA15A901E4B621035136F83345" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:23D4E8AA15A901E4B621035136F83345" country="Mexico" municipality="Carr. Actopan" name="Passando la Desv. Idolos" stateProvince="Veracruz de Ignacio de la Llave">Passando la Desv. Idolos</location>
;
<date id="1E0AA59A72749628AAD00635B5BB656C" value="1994-08-30">
<collectingDate id="81D59ADDADBE3ED0740A86E5C215F030" value="1994-08-30">30 Aug. 1994</collectingDate>
</date>
;
<collectorName id="11F811ACD17F2E87B66DF45AC0F29252">L. Arellano</collectorName>
&amp;
<collectorName id="E7BF0881F9D67C25FAFB06DA37FA0FAD">R. Sanchez</collectorName>
leg.;
<abbrev id="ABBRID0EGODI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="AD4B91A35C9EA48580165375F0272C50">CEMT</collectionCode>
</abbrev>
</materialsCitation>
•
<materialsCitation id="58AC5736FE238FEDD6C20B039249C2E7" collectingDate="1994-10-18" collectionCode="CEMT" collectorName="L. Arellano &amp; R. Sanchez" country="Mexico" county="Carr. Alto Lucero" location="Desviacion la Concepcion" municipality="Km" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
1;
<collectingCounty id="00B17659746547B0EB20736068AC0461">Carr. Alto Lucero</collectingCounty>
,
<collectingMunicipality id="F7ED1E4D25D6E3ACA076F0093408FFDC">Km</collectingMunicipality>
2
<location id="FD24C7D6801129C3E25CEE6CB173E88D" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:FD24C7D6801129C3E25CEE6CB173E88D" country="Mexico" county="Carr. Alto Lucero" municipality="Km" name="Desviacion la Concepcion" stateProvince="Veracruz de Ignacio de la Llave">Desviación la Concepción</location>
;
<date id="1192A72B218FF6018E0CD412FD4F21B1" value="1994-10-18">
<collectingDate id="037387E2C7007F1045F541C995CCEE25" value="1994-10-18">18 Oct. 1994</collectingDate>
</date>
;
<collectorName id="58FF8F026F564A3E1E158BBDF02F1543">L. Arellano</collectorName>
&amp;
<collectorName id="A11A3BA9E2ED84F323E749801FC352CF">R. Sanchez</collectorName>
leg.;
<abbrev id="ABBRID0EKODI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="3F60FE637D7080B64A6390C10D8F9453">CEMT</collectionCode>
</abbrev>
</materialsCitation>
•
<materialsCitation id="150A3AE3444455CEA08238392AE88C6A" collectingDate="1994-08-31" collectionCode="CEMT" collectorName="de Espinal &amp; L. Arellano &amp; R. Sanchez" country="Mexico" location="Carretera Xalapa-Alto Lucera" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
1;
<location id="1499164471E6CD8BE10AD81402EE5325" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:1499164471E6CD8BE10AD81402EE5325" country="Mexico" name="Carretera Xalapa-Alto Lucera" stateProvince="Veracruz de Ignacio de la Llave">Carretera Xalapa-Alto Lucera</location>
,
<quantity id="65904F7204F71D39055F32AC9BADD2E3" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
<collectorName id="61E7AB7D521DFCD1649A7CD7A17F6BDF">de Espinal</collectorName>
;
<date id="606AECAC345B8F02DF3A002AB2629C31" value="1994-08-31">
<collectingDate id="20F9A6A48390D9DE2782F08D61A075CA" value="1994-08-31">31 Aug. 1994</collectingDate>
</date>
;
<collectorName id="7F0C4CF0EC57E972326811F6CBA4B153">L. Arellano</collectorName>
&amp;
<collectorName id="7797435199397EDE2617EC1EB55A0C50">R. Sanchez</collectorName>
leg.;
<abbrev id="ABBRID0EOODI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="9C7CDFD56796489F5352E2E579890912">CEMT</collectionCode>
</abbrev>
</materialsCitation>
•
<materialsCitation id="6811D983EAB73133C0C82F2E6CBE1993" collectingDate="1990-04-29" collectionCode="CEMT" collectorName="Parq. de la Flora &amp; F. Capistran" country="Mexico" location="Fauna Silvestre" municipality="Catemaco" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
1;
<collectingMunicipality id="50764706A0195B9BCBB3493C76B49F46">Catemaco</collectingMunicipality>
,
<collectorName id="495FBF39E2AE710C87CDCD33647E3747">Parq. de la Flora</collectorName>
y
<location id="5B9AE01455920ACBEA699526D751A229" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:5B9AE01455920ACBEA699526D751A229" country="Mexico" municipality="Catemaco" name="Fauna Silvestre" stateProvince="Veracruz de Ignacio de la Llave">Fauna Silvestre</location>
tropical;
<date id="D7FE193A34DA3604C3FE7285BFBF9BFB" value="1990-04-29">
<collectingDate id="F8186E0E1004213FED1581EA3C6FAC85" value="1990-04-29">29 Apr. 1990</collectingDate>
</date>
;
<collectorName id="FD26AC175ABB5C4C701C1B3D943C6AD6">F. Capistran</collectorName>
leg.;
<abbrev id="ABBRID0ESODI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="0DE3F3E8DF0EC632BB16A8B4615660E6">CEMT</collectionCode>
</abbrev>
</materialsCitation>
•
<materialsCitation id="54F57D8F6F3EA2ED2373EF71350B90FD" collectingDate="1997-03-19" collectionCode="CEMT" collectorName="R. Sanchez &amp; M. E. Favila" country="Mexico" location="Mpio de Papantla" municipality="Plan" specimenCount="1" stateProvince="Hidalgo">
1;
<collectingMunicipality id="B43FD220DE7D162AAAB70873032167CB">Plan</collectingMunicipality>
de
<collectingRegion id="C24B441F9CD941C8C66425872AD1C9E6" country="Mexico" name="Hidalgo">Hidalgo</collectingRegion>
,
<location id="4700DF845DDA0AB2DFFA895F6F9781DA" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:4700DF845DDA0AB2DFFA895F6F9781DA" country="Mexico" municipality="Plan" name="Mpio de Papantla" stateProvince="Hidalgo">Mpio de Papantla</location>
;
<date id="6EFBD817BA01D39B263995C53F67EB27" value="1997-03-19">
<collectingDate id="78577906DC5BAEA292EF7F72D17D4AB5" value="1997-03-19">19 Mar. 1997</collectingDate>
</date>
;
<collectorName id="21E798B50B0D8C34F96D6D91941B08B9">R. Sanchéz</collectorName>
&amp;
<collectorName id="5055F68AC08588846A12D6D497108B55">M. E. Favila</collectorName>
leg.;
<abbrev id="ABBRID0EWODI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="CA56FB68DC8D2BA1F75A7A1B0503F406">CEMT</collectionCode>
</abbrev>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

126
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<subSubSection id="A20F31918C03BB48DD36858B43B222EC" type="nomenclature">
<paragraph id="DE2CB2791FDF757BC28C5F9DB27EAB14">
<heading id="601597A16D95BC2608E1AE262E3A6B8A" reason="label">Genus</heading>
<taxonomicName id="10E6FC8CD4F9E61F33D106D7F9E7983F" ID-CoL="63FX3" authority="Erichson, 1847" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="3C3A582B23442E8A083F11E8FDB00771" italics="true">Omorgus</emphasis>
<bibRefCitation id="095DABD1696A732369AE4723D1B09CCF" author="Erichson WF" issue="1" journalOrPublisher="Archiv für Naturgeschichte" pagination="67-185" refId="B37" refString="Erichson WF (1847) Conspectus coleopterorum quae in Republica Peruana observata sunt. Archiv fur Naturgeschichte 13 (1): 67 - 185." title="Conspectus coleopterorum quae in Republica Peruana observata sunt." volume="13" year="1847">Erichson, 1847</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJAAE" type="notes">
<paragraph id="1C69ED2F5055A77BE64883500D0ED934">
<heading id="1DF4A535DDE6BAF7A0A8D7B34FF5B5F9" reason="title">Note.</heading>
</paragraph>
<paragraph id="3793CF04B0E63CB2F496DFB2876ACC96">
For morphological description and diagnosis, see
<bibRefCitation id="93D03C6AB93D1E339315903FE23CE153" author="Vaurie" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105–167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie (1962</bibRefCitation>
– as “ species group
<emphasis id="66CAE53E1EB18BA005332622349D3E02" italics="true">suberosus</emphasis>
”),
<bibRefCitation id="395B42CD6A43C4002772A0873AAA3F0F" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz (1990)</bibRefCitation>
, and
<bibRefCitation id="DDBA615C96A92C92E1F135581E593E30" DOI="10.1163/1876312X-46052133" author="Strümpher WP &amp; Villet MH &amp; Sole CL &amp; Scholtz CH" etAl="et al." firstAuthor="Strumpher" issue="1" journalOrPublisher="Insect Systematics &amp; Evolution" pagination="53-82" refId="B105" refString="Strümpher WP, Villet MH, Sole CL, Scholtz CH (2016) Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea). Insect Systematics &amp; Evolution 47 (1): 53–82. https://doi.org/10.1163/1876312X-46052133" title="Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea)." volume="47" year="2016">Strümpher et al. (2016)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFBAE" type="type species">
<paragraph id="BD511BD73905E22B54B4D3A9F7F64A51">
<heading id="493050660BC88EB9C38D31D6FD538362" reason="title">
<typeStatus id="0CC3B0462D95C90B6AF99D014A91FBAA">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="3B27ED2D414E45AD97E4D983DDE44324">
<taxonomicName id="CFC94A51DA3E5BECA983A6D415F03FE5" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="suberosus">
<emphasis id="92E0AF3AAF2A99983017FAF173261C25" italics="true">Trox suberosus</emphasis>
<bibRefCitation id="42FB10C6C0BA20F01FEFA6D42D7ED9CD" DOI="10.5962/bhl.title.36510" author="Fabricius JC" refId="B38" refString="Fabricius JC (1775) Systema Entomologiae, sistens insectorvm Classes, Ordines, Genera, Species, adiectis Synonymis, Locis, Descriptionibvs, Observationibus. Kortii, Flensburg and Leipzig [&quot; Flensburgi et Lipsiae &quot;]; Havniae [= Copenhagen], p. [32] + 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510" year="1775">Fabricius, 1775</bibRefCitation>
</taxonomicName>
(by subsequent designation,
<bibRefCitation id="65E4E52772FA4100C13F203FD1CFCACB" author="Lacordaire JT" journalOrPublisher="Tome troisième contenant les familles des pectinicornes et lamellicornes, Roret, Paris" refId="B61" refString="Lacordaire JT (1856) Histoire naturelle des insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme contenant les familles des pectinicornes et lamellicornes, Roret, Paris, 594 pp." title="Histoire naturelle des insectes. Genera des Coléoptères ou exposé méthodique et critique de tous les genres proposés jusqu’ici dans cet ordre d’insectes." year="1856">Lacordaire, 1856</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWBAE" type="Identification key">
<paragraph id="019403250DB8338661ABE8CE85A2CAF8">
<heading id="85DA863AB527ED7E1C2F935CB7EBB090" reason="title">Identification key.</heading>
</paragraph>
<paragraph id="6246484F8FCCF9634503FFC38FE9E6F0">
The identification keys of
<bibRefCitation id="E5E7BDAC6C6C781F39EEE8FEB83BFB03" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105–167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie (1962)</bibRefCitation>
and
<bibRefCitation id="BA6B9658C28BE60D9B39DF7A433F323A" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz (1990)</bibRefCitation>
can be used to identify South American species of
<taxonomicName id="296C5351476ACE8E891683AEE532A985" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="FD463BB10CEC4DB45AA0063010E3F0AB" italics="true">Omorgus</emphasis>
</taxonomicName>
, while
<bibRefCitation id="01B6D9A3E663399824F6E05EBA748D7C" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021)</bibRefCitation>
can be used for Brazilian species.
</paragraph>
</subSubSection>
</treatment>
</document>

197
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<mods:title id="4FB05B1FCCA3ADE86095B9AF3FAE2C6F">Dickyyuella argentinensis a tentative new genus and species of Cardiochilinae (Hymenoptera, Braconidae) from the Neotropical region</mods:title>
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<subSubSection id="8CBC8EE8129BCAD724D0F960878624D6" type="nomenclature">
<paragraph id="74C9890485E8D3FA2105C022CCCF2505">
<taxonomicName id="A3EFCFCB0A75C805783421AD53CCDFA3" authority="Kang &amp; Sharkey" authorityName="Kang &amp; Sharkey" authorityYear="2024" class="Insecta" family="Braconidae" genus="Dickyyuella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F5FD33C7508CE135884E94781FF1EDA5" italics="true">Dickyyuella</emphasis>
Kang &amp; Sharkey
</taxonomicName>
<taxonomicNameLabel id="3CAC58575D15EF30B44D78DE66A84956" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="81B60FCDB792CF511AE01AABC69FA5C0" type="description">
<paragraph id="333BCF0DA5922AD7989D6ABE6CA7CE65">
<figureCitation id="6ECAD2CAC54032EBDDDE6E36C7FAE1DE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">Fig. 1 A – E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESFAC" type="type species">
<paragraph id="C4AD4C0B19E2266B5963B4B1F7932B02">
<heading id="9ABFA559E131C071F72BFCEB9973C7C9" reason="title">
<typeStatus id="8274B7CA325F62F1934A9838EDA1FBA8">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="29D9A4EC270DFC57B86EBB6060C97EDA">
<taxonomicName id="3696347C4C02DBD1E5560E6EC8198F8C" authority="Kang &amp; Sharkey" authorityName="Kang &amp; Sharkey" authorityYear="2024" class="Insecta" family="Braconidae" genus="Dickyyuella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="argentinensis" status="sp. nov.">
<emphasis id="9BBE78127D8F62125D721D62487A21D4" italics="true">Dickyyuella argentinensis</emphasis>
Kang &amp; Sharkey
</taxonomicName>
,
<taxonomicNameLabel id="55CCF508BCA63495EBFAFFB826C01775" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDGAC" type="diagnosis">
<paragraph id="EE6A918D1AE7A5BCD535F2CC151016A5">
<heading id="C4DDA70ABBAD2A03FFE794DE62B44BFF" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="0FF037CB65D6D8817DDEABBC7B14961B">
Body relatively small compared to members of the other cardiochiline genera, with strong sculpture, especially on mesosoma. Antenna thick (Fig.
<figureCitation id="653799D30A618A9A30978BE1E5917409" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 A</figureCitation>
). Eyes bare (Fig.
<figureCitation id="4B4410780F0EC7599124E5A2BD25CCB4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 B</figureCitation>
). Occipital carina well developed dorsally, absent ventrally (Fig.
<figureCitation id="6E590F8EE8B0B7908C10A5BD0E802088" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 E</figureCitation>
). Most of head with weak microsculpture. Median ocellus surrounded medially and laterally by a smooth, curved ridge. Pronotum bilobed anteriorly with a transverse plate dorsally. Notauli deeply impressed and entirely costate (Fig.
<figureCitation id="CE58C2897523FA54EA4C11B818F6B750" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 E</figureCitation>
). Median lobe of mesoscutum bilobed. Scutellar sulcus deep with a median carina (Fig.
<figureCitation id="6FE61C446E52271972E11904F391EA69" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 E</figureCitation>
). Scutellum smooth and flat. Postscutellar depression absent (Fig.
<figureCitation id="7DCE9C00AEFB1F61D737F48D3BA52C08" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 E</figureCitation>
). Epicnemial carina strong and complete (Fig.
<figureCitation id="CE9D23D6ED9A84704A2393009DA7DD76" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 B</figureCitation>
). Precoxal sulcus well defined with ~ 5 costulae (Fig.
<figureCitation id="CC19E0998DCEBBDF1390CF249B9E24B0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 B</figureCitation>
). Propodeum rugose with a large, well-defined median areola. Apical abscissa of RS entirely nebulous and almost straight, very slightly curved posteriorly (Fig.
<figureCitation id="817B1BCCA43A35F2A92D728112FA0B03" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 C</figureCitation>
). (RS + M) b about 3 × longer than m-cu vein (Fig.
<figureCitation id="5C1C249EFC00A31E11F54D2D91770033" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 C</figureCitation>
). 1 M about 3 × longer than m-cu vein; lacking distinct claval lobe. Veins M + Cu and M about equal in length. Hind basitarsomere swollen (Fig.
<figureCitation id="6FF2AE2FDB748FFE2A249ADA4B618FB3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 A</figureCitation>
). Tarsal claws rather large with pectinate base (Fig.
<figureCitation id="D0815885D0A713AE60F9400FFD21B18D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 D</figureCitation>
). Spiracle of
<collectionCode id="BAD10D2F547EEA0C38FD831C55BD0E4C" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
1 on membranous laterotergite (Fig.
<figureCitation id="59F7498E9FEA893E1C729C6AC9F54898" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 B</figureCitation>
).
<collectionCode id="AD294916A7A4EF2416A9E8E4FE02508E" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
1 wide with carinate lateral margins; medial area of
<collectionCode id="4819847763D41B3D71913396CF8303B9" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
1 with an inverse Y-shaped depression (Fig.
<figureCitation id="069EC528EA9021FFF7834AA41A3F67EC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dickyyuella argentinensis Kang &amp; Sharkey, sp. nov. holotype A lateral habitus of the specimen, including antennae and wings B lateral view, zoomed in on head, mesosoma, and metasoma C wings D hind tarsal claw E dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma." figureDoi="10.3897/zookeys.1208.128640.figure1" httpUri="https://binary.pensoft.net/fig/1103660">1 E</figureCitation>
).
</paragraph>
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<paragraph id="4F09CF749F4259E813ED105104A57071">
<label id="7B27E0BF885ACD9AA214472031A85CB8">Figure 1.</label>
</paragraph>
<paragraph id="3D7F6D2F0497E32B43CAEB771D1801A1">
<taxonomicName id="350F3059D0F3723D4B24D950DDC1C99E" authority="Kang &amp; Sharkey" authorityName="Kang &amp; Sharkey" authorityYear="2024" class="Insecta" family="Braconidae" genus="Dickyyuella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="argentinensis" status="sp. nov.">
<emphasis id="3D96DDCA700381CA69848C24E2002BD4" italics="true">Dickyyuella argentinensis</emphasis>
Kang &amp; Sharkey
</taxonomicName>
,
<taxonomicNameLabel id="1CBDBA5ED5F3C4EE00C11239EA563822" rank="species">sp. nov.</taxonomicNameLabel>
holotype
<emphasis id="D58BB26B103FCB9D5D83A179D7AD3550" bold="true">A</emphasis>
lateral habitus of the specimen, including antennae and wings
<emphasis id="C115C06403F83A9FB0D3D1908BE4AAF1" bold="true">B</emphasis>
lateral view, zoomed in on head, mesosoma, and metasoma
<emphasis id="26C53DD7DC3AAE8CD4E916B353376130" bold="true">C</emphasis>
wings
<emphasis id="22592C3B1FB34B3F800A77CF16B5121A" bold="true">D</emphasis>
hind tarsal claw
<emphasis id="E6FC2DE9F937A355DEEDD1761BAB1E60" bold="true">E</emphasis>
dorsal habitus of the specimen, zoomed in on head, mesosoma, and metasoma.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EDJAC" type="biology">
<paragraph id="19140D10152908DEB858433E6761E43D">
<heading id="DF62C738790924B3EE085037B7B0F6F9" reason="title">Biology.</heading>
</paragraph>
<paragraph id="DB9FD8A904F846C977DD55578E75057D">Unknown.</paragraph>
</subSubSection>
<subSubSection id="SECID0EIJAC" type="distribution">
<paragraph id="F13760085085EDE22530CF628FECDB85">
<heading id="5198E584890B215A218AE6615C32688D" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="86EC4C89CA732C089A23EC02AA8F1DE6">Neotropics.</paragraph>
</subSubSection>
<subSubSection id="SECID0ENJAC" type="etymology">
<paragraph id="B2C66B8BFC6C1D30EB7091AD90C09822">
<heading id="2D8E0695D563CD53752EAB2B59407AFC" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="E3CF20F32A67F092CDEEB3890E00B96C">
The genus name is a patronym in honor of Dicky Sick Ki Yu, who developed Taxapad and made significant contributions to
<taxonomicName id="0352B98ED5F5D53C5A5F66FA8D0ED6DD" authorityName="Latreille" authorityYear="1829" class="Insecta" family="Braconidae" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="family">Braconidae</taxonomicName>
and
<taxonomicName id="CB836C48A0993E1DEDE0B00902275010" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="family">Ichneumonidae</taxonomicName>
systematics research. Gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4JAC" type="notes">
<paragraph id="9092F034C09E5FC94BFCD2B56CA806AA">
<heading id="84A9E7EBFCA634CCC2378145FA0E8708" reason="title">Notes.</heading>
</paragraph>
<paragraph id="858454E5E43CF8E4B74988F543F418CE">
The members of
<taxonomicName id="C1637EAF8FEC9CB44E260DC4F33834B5" authority="Kang &amp; Sharkey" authorityName="Kang &amp; Sharkey" authorityYear="2024" class="Insecta" family="Braconidae" genus="Dickyyuella" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CAD5BF3A7D0C035A4895370A4091D76D" italics="true">Dickyyuella</emphasis>
Kang &amp; Sharkey
</taxonomicName>
,
<taxonomicNameLabel id="D2FE8F8E0F7BA36D840DCDC9D74F48D3" rank="genus">gen. nov.</taxonomicNameLabel>
will run to couplet
<quantity id="2DB10BF24327BB04AF73C6174E5233F9" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
the key to the world genera by
<bibRefCitation id="D77AC064726C7EE8C1986968DF772EF5" DOI="10.1071/IT98020" author="Dangerfield PC &amp; Austin AD &amp; Whitfield JB" etAl="et al." firstAuthor="Dangerfield" issue="6" journalOrPublisher="Invertebrate Systematics" pagination="917-976" refId="B3" refString="Dangerfield PC, Austin AD, Whitfield JB (1999) Systematics of the world genera of Cardiochilinae (Hymenoptera: Braconidae). Invertebrate Systematics 13 (6): 917–976. https://doi.org/10.1071/IT98020" title="Systematics of the world genera of Cardiochilinae (Hymenoptera: Braconidae)." volume="13" year="1999">Dangerfield et al. (1999)</bibRefCitation>
, but it can be easily distinguished from members of
<taxonomicName id="BD5CBB94AF0E00059BE79C8B8A1285A8" authorityName="Brullé" authorityYear="1846" class="Insecta" family="Braconidae" genus="Heteropteron" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="F81BF2C3DED78FF7DF27F05A2213465F" italics="true">Heteropteron</emphasis>
</taxonomicName>
and
<taxonomicName id="B7C6059C6CBE52DCC447E99A93DDD5B6" authorityName="Szépligeti" authorityYear="1908" class="Insecta" family="Braconidae" genus="Neocardiochiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="F35448A7BA3A3E53BA746123D9471854" italics="true">Neocardiochiles</emphasis>
</taxonomicName>
by the size, well-developed occipital carina, deep and broad notauli, large median areola on propodeum, and rugose propodeum.
</paragraph>
<paragraph id="705D78D8FB5E3479D2036872A24D5668">
<title id="EA80D14165180F3610C7E8B032A4B16D">Species description</title>
</paragraph>
</subSubSection>
</treatment>
</document>

514
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@ -0,0 +1,514 @@
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<subSubSection id="09ED2136318458953535773DE83E0C40" type="nomenclature">
<paragraph id="F39A703A5952783FEE9D77ADB3FA8510">
<taxonomicName id="ECA509D015A3E35CC4FCD39E6EC9F886" authority="Boruah, Narayanan, Deepak &amp; Das" authorityName="Boruah, Narayanan, Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namdaphaensis" status="sp. nov.">
<emphasis id="62F7D0BCEC35E4DC75EEA83FE92A743E" italics="true">Cyrtodactylus namdaphaensis</emphasis>
Boruah, Narayanan, Deepak &amp; Das
</taxonomicName>
<taxonomicNameLabel id="42178951028679F972E6CB0BCAAF1346" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="F94BED39C9C9C107B144087EE2CD0A63" type="description">
<paragraph id="8A36BF21293AD9908FAA4E4D6FB3F42D">
<figureCitation id="E8B53B4FA7E2B2D6389CC80E5102CF0D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyrtodactylus namdaphaensis sp. nov. A – G holotype (WII-ADR 1416), A dorsal, and B ventral view of the whole body; C dorsal, D ventral, and E lateral view of head; F ventral view of left manus and G left pes; H precloacal pores of paratype WII-ADR 1417; I paratype WII-ADR 1417, and J paratype WII-ADR 1415 in life. Scale bar: 10 mm. I, J not to scale." figureDoi="10.3897/vz.74.e124752.figure8" httpUri="https://binary.pensoft.net/fig/1102274">Figure 8</figureCitation>
;
<tableCitation id="F72CC7A904C88D6391F042C7D73D1965" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">Tables 2, S 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EY5CK" type="Holotype">
<paragraph id="0DD30D386E1F6B21EC86F7288BCF3935">
<heading id="34FF587484DA47BEFC2984318A278286" reason="title">
<typeStatus id="43E21D17D290ED480584DAD6A8E07761" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="97399F86377217FB2F8E8F0D7AF8CDB3">
<materialsCitation id="7E7FE9731E7B89E3D7AAD6FBB3F8D56A" collectingDate="2022-05-18" country="India" county="Bitupan Boruah" elevation="650" latitude="27.4595" location="Kamala Valley" longLatPrecision="6" longitude="96.4279" municipality="Abhijit Das" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
Adult male (WII-ADR 1416), collected from
<location id="016069FAB6E452DB5B75AF67F673C652" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:016069FAB6E452DB5B75AF67F673C652" country="India" county="Bitupan Boruah" latitude="27.4595" longLatPrecision="6" longitude="96.4279" municipality="Abhijit Das" name="Kamala Valley" stateProvince="Arunachal Pradesh">Kamala Valley</location>
(
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<geoCoordinate id="356961E8EF822186FD330ADC4D1E4829" degrees="27.4595" direction="north" orientation="latitude" precision="5" value="27.4595">27.4595 ° N</geoCoordinate>
;
<geoCoordinate id="8F9E22EEED298D2D4D2CEA7C3C0281B6" degrees="96.4279" direction="east" orientation="longitude" precision="5" value="96.4279">96.4279 ° E</geoCoordinate>
</named-content>
;
<elevation id="96793DD8815A9D9C46300B3D6A95DC72" metricMagnitude="2" metricUnit="m" metricValue="6.5" unit="m" value="650.0">
elevation
<quantity id="D79A83D1CBB41264A384D5A381E57B3D" metricMagnitude="2" metricUnit="m" metricValue="6.5" unit="m" value="650.0">650 m</quantity>
a. s. l.
</elevation>
),
<location id="C885B0EF22DD1794A31BF04675E2F406" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:C885B0EF22DD1794A31BF04675E2F406" country="India" county="Bitupan Boruah" latitude="27.4595" longLatPrecision="6" longitude="96.4279" municipality="Abhijit Das" name="Namdapha Tiger Reserve" stateProvince="Arunachal Pradesh">Namdapha Tiger Reserve</location>
,
<location id="E4A3028A8163D85E4C350A92639AAFD3" LSID="urn:lsid:plazi:treatment:6B05318011F35D7C95E621A314732FC0:E4A3028A8163D85E4C350A92639AAFD3" country="India" county="Bitupan Boruah" latitude="27.4595" longLatPrecision="6" longitude="96.4279" municipality="Abhijit Das" name="Changlang District" stateProvince="Arunachal Pradesh">Changlang District</location>
,
<collectingRegion id="20E2ADC38446B988C443265855911E38" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="A6584DC4680CF6BDF07EC84EF1884EA4" name="India">India</collectingCountry>
by
<collectingMunicipality id="77A37FE350048406A86502CF3B73DEEA">Abhijit Das</collectingMunicipality>
and
<collectingCounty id="C85A79F6C94EA4028B5331AF27B836D6">Bitupan Boruah</collectingCounty>
on
<date id="C348B582366DE5FFD64907F3ECF29BEC" value="2022-05-18">
<collectingDate id="AFB9BB98FA99EF9B7CAA20A1813CF2DB" value="2022-05-18">18 May 2022</collectingDate>
</date>
(Fig.
<figureCitation id="9CC475C6FCE56B30D46B9F79E1D7428C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 A</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK6CK" type="Paratypes">
<paragraph id="BCE382CAE81E64506A16DF384631BD20">
<heading id="5E5D34A2A77F5C926422521525134123" reason="title">
<typeStatus id="BFF8E6BE8C85F10FFFCDE5FD66ABF5A6" type="paratype">Paratypes</typeStatus>
.
</heading>
</paragraph>
<paragraph id="ABDEAD43EDE1F01394612366B3D4CDDC">
<specimenCount id="08CA1623AEA3FDD42BC03789E6267BC1" count="2" type="adult">Two adult</specimenCount>
males (WII-ADR 1415, WII-ADR 1417) collected from the same locality as the
<typeStatus id="52FCA7858678FB875BEB952F25F8E1A3" type="holotype">holotype</typeStatus>
on the same date;
<specimenCount id="1D3E531609EE26BDF27A66241AF4724A" count="1" type="adult">one adult</specimenCount>
female (WII-ADR 1404) collected near Deban (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ET6CK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.370100,27.494200]}">
<geoCoordinate id="2E6BF5C260830EFE0F4A91D09D1ACFF1" degrees="27.4942" direction="north" orientation="latitude" precision="5" value="27.4942">27.4942 ° N</geoCoordinate>
;
<geoCoordinate id="9F1DAECC4EE6FF4053191F9FB46DDAF6" degrees="96.3701" direction="east" orientation="longitude" precision="5" value="96.3701">96.3701 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="DF9E7E6654A2636A40D7779BFBA416CC" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</quantity>
a. s. l.), Namdapha Tiger Reserve, Changlang District,
<collectingRegion id="9A92A90AA0264A67C2147FEF7BFDA7B3" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="E511377949896FE8D05F38888D81D3C4" name="India">India</collectingCountry>
by Abhijit Das and Bitupan Boruah on
<date id="A89057C8B6144D5D640508DE0356BA53" value="2022-05-11">11 May 2022</date>
;
<specimenCount id="141D74BF4D005A226356967F18E82507" count="2" type="adult">two adult</specimenCount>
females (WII-ADR 3067, WII-ADR 3068) collected from Motijheel trail (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E26CK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.334800,27.489900]}">
<geoCoordinate id="981C8340D3E57524E03BAC6093CC3627" degrees="27.4899" direction="north" orientation="latitude" precision="5" value="27.4899">27.4899 ° N</geoCoordinate>
,
<geoCoordinate id="D80A7B12AE4559C5427AB52A772B3C8A" degrees="96.3348" direction="east" orientation="longitude" precision="5" value="96.3348">96.3348 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="EDF933E331F4678E124DE2BDA270F9A5" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</quantity>
a. s. l.), Gibbons’ Land, Namdapha Tiger Reserve, Changlang District,
<collectingRegion id="218B4A141E95789A7C4783F22F7D5267" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="ED607637F6B0E11B13AA78BA5DDC488C" name="India">India</collectingCountry>
by Abhijit Das and Bitupan Boruah on
<date id="9318970E16A177EBCF0961655A62ADB6" value="2022-09-11">11 September 2022</date>
;
<specimenCount id="D6A07B524509B399A17720C247AE0F4E" count="1" type="adult">one adult</specimenCount>
female (WII-ADR 1790) collected near
<collectingCountry id="4B71C6FED809D0FEFE8763D962639713" name="Myanmar">Burma</collectingCountry>
Nullah (
<quantity id="E32FD9799914E401015B53E5FF09022A" metricMagnitude="4" metricUnit="m" metricValue="6.437376" unit="mi" value="40.0">40 mile</quantity>
point) (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDADK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.541600,27.487800]}">
<geoCoordinate id="158529EBBBE4070647DCD5E7FF080AB0" degrees="27.4878" direction="north" orientation="latitude" precision="5" value="27.4878">27.4878 ° N</geoCoordinate>
;
<geoCoordinate id="65EFBA1B38799B371864B3C214766B61" degrees="96.5416" direction="east" orientation="longitude" precision="5" value="96.5416">96.5416 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="9D2B9A25B85DA756FB0F933227428D96" metricMagnitude="2" metricUnit="m" metricValue="4.8" unit="m" value="480.0">480 m</quantity>
a. s. l.), Namdapha Tiger Reserve, Changlang District,
<collectingRegion id="24F47448037CA7AA5CB483F4A4BB7FCB" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="6DA73970ADDA3FB26FDC461BC844DD19" name="India">India</collectingCountry>
by Abhijit Das and Bitupan Boruah on
<date id="6AD3105EF9C5983158E2FC2F59CAF407" value="2022-09-18">18 September 2022</date>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIADK" type="material">
<paragraph id="E200D9F0E977C04092CB564C213B6C55">
<heading id="D1ED0B7FAA7E3C93B8AA6226622FCADB" reason="title">Referred materials.</heading>
</paragraph>
<paragraph id="5F3DB67A0F51D5FF026FD5826CE54144">
An adult female (WII-ADR 3060) collected from Sinabrai (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERADK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.387200,27.743400]}">
<geoCoordinate id="568417FD29E6E62FF1C96DBF361BBF61" degrees="27.7434" direction="north" orientation="latitude" precision="5" value="27.7434">27.7434 ° N</geoCoordinate>
;
<geoCoordinate id="454F33EF3373862354F0583E73FC6D81" degrees="96.3872" direction="east" orientation="longitude" precision="5" value="96.3872">96.3872 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="792D1D7B387058FF20F7BD645771916D" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</quantity>
a. s. l.), Kamlang Tiger Reserve, Lohit District,
<collectingRegion id="2F9F785362B33C113032AFBEEA2C2FDA" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="3A4C4FBB6A39EC1615B02F1652B217B8" name="India">India</collectingCountry>
on
<date id="5AF38BDD10C50A76F60DA449B6A79B3A" value="2022-09-05">5 September 2022</date>
by Bitupan Boruah and Abhijit Das; an adult female (WII-ADR 3281) and an adult male (WII-ADR 3282) collected from Hornbill (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZADK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.440300,27.538100]}">
<geoCoordinate id="FBCD3653BAA983934A5DDC1F71FC6D78" degrees="27.5381" direction="north" orientation="latitude" precision="5" value="27.5381">27.5381 ° N</geoCoordinate>
;
<geoCoordinate id="A7F213FD7EB9092A0BABEED401953DF2" degrees="96.4403" direction="east" orientation="longitude" precision="5" value="96.4403">96.4403 ° E</geoCoordinate>
</named-content>
; elevation
<quantity id="E5181AA3D7199B34DEACB46554A16801" metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">670 m</quantity>
a. s. l.), Namdapha Tiger Reserve, Changlang District,
<collectingRegion id="CB00BD2DD73E6FC9F6AE1CDEB4930F37" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="2E5ADDDF27246FD866EFDE3A3C06F01C" name="India">India</collectingCountry>
on
<date id="BFFBF276C5EBDD13DE60D0D944EB4E18" value="2023-05-10">10 May 2023</date>
by Rajiv
<taxonomicNameLabel id="841FB27F9F7F3678962C7C0DEE96B224" rank="variety">
N.
<collectionCode id="01534B89436CB073006DC30D33A7124C" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5ADK" type="diagnosis">
<paragraph id="8F51397702DB3881E735595F932A7E5F">
<heading id="8B53C5433271BA7439219AB20630E53A" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="AD17BB2DB9D91480764AEFA861412A08">
Medium-sized gecko (
<abbrev id="ABBRID0EEBDK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="795CF03282CDD901515B4D16C210D7C2" metricMagnitude="-2" metricUnit="m" metricValue="6.409999999999999" metricValueMax="7.07" metricValueMin="5.75" unit="mm" value="64.1" valueMax="70.7" valueMin="57.5">57.5–70.7 mm</quantity>
in males and
<abbrev id="ABBRID0EIBDK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="6A21ED7ACE9131253BF8DABADDD766C7" metricMagnitude="-2" metricUnit="m" metricValue="6.205" metricValueMax="6.93" metricValueMin="5.4799999999999995" unit="mm" value="62.05" valueMax="69.3" valueMin="54.8">54.8–69.3 mm</quantity>
in females); supralabials and infralabials 8–11; dorsum with weakly keeled and bluntly conical tubercles, 29–36 paravertebral tubercles between the level of axilla and level of groin; 17–19 dorsal tubercle rows at mid body; 33–40 mid ventral scale rows; seven to nine precloacal pores in males and 8–10 small precloacal pores in females in a continuous series,
<abbrev id="ABBRID0EMBDK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
much smaller in females than that of males; 12–14 subdigital lamellae on finger IV and 11–17 subdigital lamellae on toe IV; irregular dark-brown spots or stripes on dorsum.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQBDK" type="description">
<paragraph id="DBB41B8F57BE486F16760F22661AA774">
<heading id="C6549646AB5212FAF5F345AAA1C418AA" reason="title">
Description of
<typeStatus id="FFD454A43977B0890FB0EDF196F9DF6E" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="C25DCCC1997E8FFDDD16C9EBFF322FCE">
<typeStatus id="2B07886253F0AC10FE82B29EEA90EB69" type="holotype">Holotype</typeStatus>
well preserved except a ventrolateral incision below left axilla. Snout-vent length
<quantity id="31553A942A94FD357522118B5346B715" metricMagnitude="-2" metricUnit="m" metricValue="6.529999999999999" unit="mm" value="65.3">65.3 mm</quantity>
. Head moderately large (
<abbrev id="ABBRID0EWBDK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
/
<abbrev id="ABBRID0E1BDK" xlink_title="snout to vent length">SVL</abbrev>
= 0.27), dorsoventrally depressed, ovoid in shape, longer than width (
<abbrev id="ABBRID0E5BDK" xlink_title="maximum head width">HW</abbrev>
/
<abbrev id="ABBRID0ECCDK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.66), distinct from neck, broader at occipital region; snout rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat, a longitudinal furrow on dorsal surface of the snout, snout short (
<abbrev id="ABBRID0EGCDK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
/
<abbrev id="ABBRID0EKCDK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.4), longer than orbit (
<abbrev id="ABBRID0EOCDK" xlink_title="orbit diameter, taken between the anterior and posterior bony orbital borders">OD</abbrev>
/
<abbrev id="ABBRID0ESCDK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
= 0.71); nostril semicircular, opening directed posterolaterally; ear opening oval and oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids nearly homogeneous, granular juxtaposed; supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at the anterodorsal region; rostral wide, a short groove at the middle on top, rostral connected with nasals, supranasals, internasals and first supralabials; granular scales at parietal, occipital and temporal region intermixed with slightly large, rounded and bluntly conical tubercles, dense in occipital and temporal region and size increases towards nape; supralabial 11 on right and 10 on left side, supralabials up to midorbit eight on right and seven on left side, size decreases towards angle of jaw; a series of scales slightly larger than the loreal scales present above the supralabials, posteriorly size decreased; mental as wide as rostral, nearly triangular, connected with first infralabials, inner postmentals; nine infralabials on right and eight on left side, size decreases towards angle of jaw; inner pair of postmentals are larger than the outer postmentals; two rows of enlarged scales along the infralabials starting below the outer postmentals, posteriorly size of those decreases; rest of the gular scales are small, granular juxtaposed, homogeneous, size increases towards throat where they becomes imbricate.
</paragraph>
<paragraph id="8F452A802A91E0788BABBC16018EE6BA">
Habitus slender (
<abbrev id="ABBRID0EYCDK" xlink_title="maximum body width">BW</abbrev>
/
<abbrev id="ABBRID0E3CDK" xlink_title="snout to vent length">SVL</abbrev>
= 0.17,
<abbrev id="ABBRID0EADDK" xlink_title="trunk length, between the axilla and groin">TRL</abbrev>
/
<abbrev id="ABBRID0EEDDK" xlink_title="snout to vent length">SVL</abbrev>
= 0.47), dorsoventrally depressed; dorsal scales granular, rounded, heterogeneous, intermixed with rounded, irregularly arranged weakly keeled and bluntly conical tubercles, these tubercles continues to fourth segment of the tail, size increases towards posterior body and pronounced; 18 dorsal tubercles across mid dorsum; 36 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate, largest towards belly; 40 mid-ventral scales between ventrolateral fold; seven precloacal pores arranged in an inverted “
<collectionCode id="80499BBBB258E799C45077234DD03805" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
” shaped continuous series, followed by five unpored, large scales below it, one enlarged unpored scale present on right end of the
<abbrev id="ABBRID0EIDDK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
series.
</paragraph>
<paragraph id="A9FBE752BD42314C7AA9D4EA344C8FCD">
Forelimbs and hindlimbs slender (
<abbrev id="ABBRID0EODDK" xlink_title="forearm length, taken from the elbow to the wrist">FL</abbrev>
/
<abbrev id="ABBRID0ESDDK" xlink_title="snout to vent length">SVL</abbrev>
= 0.15,
<abbrev id="ABBRID0EWDDK" xlink_title="crus length, taken from the knee to the base of the foot">CL</abbrev>
/
<abbrev id="ABBRID0E1DDK" xlink_title="snout to vent length">SVL</abbrev>
= 0.18); digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae; lamellae beneath digit IV of right and left manus (given as basal + distal) is 5 + 8; lamellae beneath digit IV of right and left pes (given as basal + distal) is 6 + 8 and 5 + 8 respectively; dorsal scales on forelimbs smooth, subimbricate and heterogeneous, small and granular at elbow, scales on forearm nearly rounded while those on hind arm are posteriorly tapering; forearm scales intermixed with enlarged rounded and bluntly conical tubercles; dorsal scales of hindlimbs heterogeneous, intermixed with large, rounded and bluntly conical tubercles, dense than those on forelimbs; horizontally upper half of the thigh scales are smooth, large and subimbricate, those on lower half small granular; scales on tibia are small, granular juxtaposed; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous in shape and size, granular juxtaposed; scales on ventral side of hindlimbs smaller than those of belly, smooth, cycloid and subimbricate, but on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate.
</paragraph>
<paragraph id="DF5872E036CF2F0F3401F0529671BFAA">
Tail regenerated (
<abbrev id="ABBRID0EAEDK" xlink_title="tail length, from the cloaca to the tail tip">TL</abbrev>
=
<quantity id="D70DD07D3CA5A414B6B2B44F881A4719" metricMagnitude="-2" metricUnit="m" metricValue="7.9" unit="mm" value="79.0">79 mm</quantity>
), slender, gradually tapering towards tip, segments indistinct, in the original part of the tail dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size; in the regenerated part of the tail, scales are irregular in shape and size; large feebly keeled scales upto fourth segment of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; two and four bluntly conical spurs on right and left side of the tail base respectively.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEEDK" type="Colouration in preservative">
<paragraph id="A39CE101E135058D187990E4E31AFACA">
<heading id="4C85FCDDE0EFAD31553BD03834AAC2FF" reason="title">Colouration in preservative.</heading>
</paragraph>
<paragraph id="CCCE88DB0D567FFAB27BD3388944A443">Top of head pale-brown, upper eyelids grey; neck and dorsum dorsally greyish-brown; two brown stripes with irregular edge on dorsal side of the neck continuing to the level of forelimb insertion, and another stripe on each lateral side of the neck; seven pairs of slightly dark-brown elongated spots of irregular shape and size on dorsum, these spots are outwardly connected to a narrow, brown dorsolateral stripe; tail dorsally pale-brown with broad dark-brown cross bars of irregular shape and size on the original part, the first bar broken into two elongated spots, regenerated part of the tail is plain pale-brown; ventrally head, trunk and tail pale-cream coloured, a few irregular brown spots on tail.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJEDK" type="Colouration in life">
<paragraph id="4DCEEBA4416797CC5BF03E046144EADE">
<heading id="C982E26317EF19A36F2EB846D2BF9265" reason="title">Colouration in life.</heading>
</paragraph>
<paragraph id="77EF1AC4FE58B2DA9D3F7C1A0CE47016">
(Based on
<typeStatus id="B206B4E6C1BFF7FEB954A9482F733A9B" type="paratype">paratype</typeStatus>
WII-ADR 1417) (Fig.
<figureCitation id="C58C103BD7BF9A93CBA6B7463C767D6F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyrtodactylus namdaphaensis sp. nov. A – G holotype (WII-ADR 1416), A dorsal, and B ventral view of the whole body; C dorsal, D ventral, and E lateral view of head; F ventral view of left manus and G left pes; H precloacal pores of paratype WII-ADR 1417; I paratype WII-ADR 1417, and J paratype WII-ADR 1415 in life. Scale bar: 10 mm. I, J not to scale." figureDoi="10.3897/vz.74.e124752.figure8" httpUri="https://binary.pensoft.net/fig/1102274">8 I</figureCitation>
); head dorsally brown with irregular dark-brown patches, a pair of pale-yellowish spots on loreal in front of anterodorsal corner of eyes, lips slightly paler than dorsal head colour with irregular pale-yellowish spots; a pale-brown postorbital streak; pale-brown irregular spots on dark-brown background on neck; upper eyelids greyish-brown; slightly dark-brown large spots of irregular size and shape on dorsal and lateral side of the trunk, interspaced with pale-brown patches, dark-brown spots on dorsal side of the trunk giving appearance of continuous stripes up to middle of the trunk; anterior two third of dorsal side of the tail with alternative broad dark-brown and narrow pale-brown bands, those dark-brown bands anteriorly diffused and posterior edge zigzagged, posterior one third brown with dark-brown marbling; limbs brown with irregular pale spots and dark-brown reticulation; digits with alternative dark-brown and pale-brown bands.
</paragraph>
<caption id="F3D54A94EE3D9E6FB3C41FC62FC0C6AA" ID-DOI="10.3897/vz.74.e124752.figure8" ID-arpha="07806C9F-284E-51CC-BB20-CC98298333D2" httpUri="https://binary.pensoft.net/fig/1102274" startId="F8">
<paragraph id="BBAA09A2F31EB11C0A7AE8922FA96ED8">
<label id="753EF062A39D6005CF92C56A8F5D05AE">Figure 8.</label>
</paragraph>
<paragraph id="E406844513192CF48518FA45F82B138A">
<taxonomicName id="B279394B54E31E094BD1B249A4ADD69B" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namdaphaensis" status="sp. nov.">
<emphasis id="7F023E3F961C7811DA5EB70DB77E31A8" italics="true">Cyrtodactylus namdaphaensis</emphasis>
</taxonomicName>
<emphasis id="24A12E52E0F7C62158108BFD2F310601" bold="true">
<taxonomicNameLabel id="AF5CAB4320AED4AC837CD332BB0B4C32" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
–
<emphasis id="0936969B886F50CE25738E5B2D7CE0FD" bold="true">G</emphasis>
holotype (WII-ADR 1416),
<emphasis id="896CBAEE5C9BC5B40EE16E68E5ACE118" bold="true">A</emphasis>
dorsal, and
<emphasis id="1FACD4F7828FAC1093812041669319F7" bold="true">B</emphasis>
ventral view of the whole body;
<emphasis id="09080EBE886B37FA8DE32D01AFA736A5" bold="true">C</emphasis>
dorsal,
<emphasis id="F1630067FFC9CC188B89A70BE354E2F8" bold="true">D</emphasis>
ventral, and
<emphasis id="0CD7DA64D2B93D8C1AF1D4FB0C32D4CB" bold="true">E</emphasis>
lateral view of head;
<emphasis id="E76840F531950E8B33729AB699818C0C" bold="true">F</emphasis>
ventral view of left manus and
<emphasis id="1A782C7EEB92A4791E5525853B6C0101" bold="true">G</emphasis>
left pes;
<emphasis id="34AF92A7CF93E051E307F19A5530FE6A" bold="true">H</emphasis>
precloacal pores of paratype WII-ADR 1417;
<emphasis id="3B204674B7882FFD73BD7E1E29D73CFD" bold="true">I</emphasis>
paratype WII-ADR 1417, and
<emphasis id="7A19A74013128DA7309F9AF54EC69E39" bold="true">J</emphasis>
paratype WII-ADR 1415 in life. Scale bar: 10 mm.
<emphasis id="B22C2436E46EB813A0302A4F51932537" bold="true">I, J</emphasis>
not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EFGDK" type="Morphological variation">
<paragraph id="2A7912C654749788ECEFA31FCADE3B5B">
<heading id="05F06E40064E4FE6B3F5B03FE24B401E" reason="title">Morphological variation.</heading>
</paragraph>
<paragraph id="1A9B401CE4780394F8A4D4E915D57F5E">Morphological variations are given in Table S 2. Except for those, the dorsal marking pattern varied among the collected specimens. The dorsal spots of WII-ADR 1417 are in the form of two continuous stripes starting from neck, posteriorly broken; a spot at the middle of occipital region followed by a dark cross bar on the nape; WII-ADR 3067 and WII-ADR 3281 has four dorsal stripes starting from neck to the level of hind limb insertion; WII-ADR 3068 has four stripes on neck, followed by five irregular and zigzag cross bars on dorsum; in WII-ADR 1790, dorsal spots are indistinct and irregular; WII-ADR 1415 has three pairs of dark spots posteriorly bordered with white spots on neck, cross bars on dorsum narrower, irregular and posteriorly with white (Fig. S 2); WII-ADR 3060 has six dark brown cross bands on back between the level of axilla and groin, consists of irregular shaped enlarged individual spots, mid dorsally these spots are remarkably disjunct,. Precloacal pores in females are smaller than that of males.</paragraph>
</subSubSection>
<subSubSection id="SECID0EKGDK" type="comparisons">
<paragraph id="036CE7FD74697C82EEE7B6018792E23D">
<heading id="FE6642BB6AA80520F1F59FCEF7F0C794" reason="title">Comparison.</heading>
</paragraph>
<paragraph id="FF0DFD1370C63A48402700CC91D682FB">
<taxonomicName id="C9E0B3FF6B89DAE41B0C962E570C313D" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namdaphaensis" status="sp. nov.">
<emphasis id="45FF8D4D1F2D8ED65099BF037FDBC0F0" italics="true">Cyrtodactylus namdaphaensis</emphasis>
</taxonomicName>
<emphasis id="D827469D8A3E844C59B66A0AF561EF7A" bold="true">
<taxonomicNameLabel id="0A0DA9476A414EBD2AA246D04A9DE0B1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="7FDA3D37A7A94A37A9F39F4EF1C2A815" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="mombergi">
<emphasis id="3768A790DB01AB1EE1156591BF1E6BA4" italics="true">C. mombergi</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0EIHDK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
17–19 (vs.
<abbrev id="ABBRID0EMHDK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="54E997A248D5EB5DE9B94306A1C17E57">DTR</collectionCode>
</abbrev>
23–27), by fewer precloacal pores,
<abbrev id="ABBRID0EQHDK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7–9 (vs.
<abbrev id="ABBRID0EUHDK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
10–11). Morphological differences with other members of
<taxonomicName id="5D6695EF9CB04A2500013CC0386C3906" baseAuthorityName="Jerdon" baseAuthorityYear="1870" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="3BD5B20B120A97485EB4A6A9C83FA978" italics="true">khasiensis</emphasis>
</taxonomicName>
group is presented in Table
<tableCitation id="8F7C0316243A2C36374D3E4A2B3475F1" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGIDK" type="Sequence divergence">
<paragraph id="2E2A992E0795726730293B168977EDE0">
<heading id="A7746CE31213A5E134C1911E4FDC9AB1" reason="title">Sequence divergence.</heading>
</paragraph>
<paragraph id="F865B98BCF98ED5F2B758F50452F0EEA">
<taxonomicName id="1C1EB2371AED3F8A62F7E7AF52A0D4FD" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namdaphaensis" status="sp. nov.">
<emphasis id="C1705B8AE4ABB2536B099E7EB02DA994" italics="true">Cyrtodactylus namdaphaensis</emphasis>
</taxonomicName>
<emphasis id="013DBB072DCBFFACA1D990FBAD27A957" bold="true">
<taxonomicNameLabel id="D06A5CA1BC0DABACB80F9E26972BF2FC" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a moderate genetic divergence of 6.2–7.9 % from the closely related
<taxonomicName id="5DA1CDB30BDA76BF68F454AFAC5B5148" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="mombergi">
<emphasis id="4A019352EF9FC27957A3755B55BB0BA8" italics="true">C. mombergi</emphasis>
</taxonomicName>
. The intraspecific divergence among the thirteen samples of
<taxonomicName id="61D8BF674D5C1DED76F854AF2DFB172A" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namdaphaensis" status="sp. nov.">
<emphasis id="C8E92813504A18F440CD5B2952FA1AFB" italics="true">C. namdaphaensis</emphasis>
</taxonomicName>
<emphasis id="E173F159A812CEB766FE42B92EE5599C" bold="true">
<taxonomicNameLabel id="1CC2DCC3E701C21838EFD3ED1E499C5F" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
ranges between 0.1–5 %.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERJDK" type="etymology">
<paragraph id="B7145EED3C4321C17D31CAB78F002F42">
<heading id="17E1DAFFB68F9BBB0F73FE8C719750CF" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="396F0A99E917C96A131E6A8ADD72AB84">
The specific epithet is a toponym named after its
<typeStatus id="812E0499C4A546FA8F1D77B4ABB10F1A">type</typeStatus>
locality Namdapha Tiger Reserve in
<collectingRegion id="C44C9587FFDE5F569F840067BC24077D" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="9163649EFEF2A7B9721E9181E4D3180E" name="India">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWJDK" type="Suggested common name">
<paragraph id="59F9983F06742A16F30C1E1EE1211266">
<heading id="78C720DA88AC87977549F855BA7CF0E3" reason="title">Suggested common name.</heading>
</paragraph>
<paragraph id="9AE7C70AA3BFD8A76B2A3D5B1C21EB93">Namdapha bent-toed gecko.</paragraph>
</subSubSection>
<subSubSection id="SECID0E2JDK" type="distribution">
<paragraph id="EDA13AF3F487DDAADD1A44424731C7D5">
<heading id="532E4F246E7A9BFED44AF1604D3B3C7E" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="55F8EF3E62C3F7A18AC68D997A49CD43">
So far,
<taxonomicName id="AEC656B837926D0D0881D6CE96D7F10F" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namdaphaensis" status="sp. nov.">
<emphasis id="028A616A39FB26F797473AFE6B25F11A" italics="true">Cyrtodactylus namdaphaensis</emphasis>
</taxonomicName>
<emphasis id="1F87AD96E4E86FD26E15B48CFD207492" bold="true">
<taxonomicNameLabel id="0155A58E2E2F5F2478EFA618379D36C0" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has been recorded within an elevational range of
<quantity id="9B0B9D333C201C7EF9C18B4DD1281D2F" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="6.5" metricValueMin="4.0" unit="m" value="525.0" valueMax="650.0" valueMin="400.0">400–650 m</quantity>
a. s. l. inside Namdapha Tiger Reserve. All the localities are south of Noa-Dihing River in Changlang District,
<collectingRegion id="2C99A9D761A26D06809A01B538BA06BB" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="B5BDE214604BB32968EA456924D21C69" name="India">India</collectingCountry>
. We recorded this species between
<date id="4D1605E801EED748375053DB395509AF" value="2022-05" valueMax="2022-09-31" valueMin="2022-05-00">May and September 2022</date>
. Individuals were recorded on tree bark, ferns, and riparian vegetation and along forest trails between 18: 00–23: 00 hrs. The forest
<typeStatus id="D522B6BA5D96E5798ED299C2E45338FF">type</typeStatus>
can be classified as
<collectingRegion id="DE458EA2BB1FAC62E08F0A00AA53AD3E" country="India" name="Assam">Assam</collectingRegion>
Valley Tropical Evergreen Forest. The area had a distinct understory with a thick covering of leaf-litter. Currently the species has been recorded from 25 Mile,
<collectingCountry id="B8BF08A4A8C9F94AE33622E3E940024C" name="Myanmar">Burma</collectingCountry>
Nullah (40 Mile), Gibbons Land, Motijheel trail and at Hornbill camp located at the north bank of Noa-Dihing River within the Namdapha Tiger Reserve. We also recorded this species on rocks and vegetation near Kamlang River at Sinabrai, near the Kamlang Tiger Reserve, at an elevation of
<quantity id="B884D89A3A7B72230F56447DFE8F1E1E" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</quantity>
a. s. l. We observed juveniles of this species on the forest floor in the leaf litter during May, 2022. During 2023, adult individuals were seen on the ferns overhanging first order streams. At the slightest disturbance, the lizards would drop into the thick vegetation below. Other arboreal reptile taxa from the area included
<taxonomicName id="EA6874855050572901F2674949B28919" class="Squamata" family="Agamidae" genus="Ptyctolaemus" kingdom="Animalia" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="D51A250B154600D5EDB2B69CD19767C5" italics="true">Ptyctolaemus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="0D7AB285527A6297EF2E26BA25006F39" class="Squamata" family="Agamidae" genus="Japalura" kingdom="Animalia" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="6AAA922653F3C2BE29454A4DCCD842C9" italics="true">Japalura</emphasis>
sp.
</taxonomicName>
and
<taxonomicName id="BFEF5D4593310DFD1F9A84267CAA702F" class="Squamata" family="Pareidae" genus="Pareas" kingdom="Animalia" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="7FEA4DC194A4127EB9113C091AFE21B1" italics="true">Pareas</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

296
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<mods:title id="FA8E47A69E2DF260664191631B7B2C9D">Four new Dolichopoda species from Greece, one Troglophilus new to Greece and new locality records (Orthoptera, Rhaphidophoridae)</mods:title>
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<mods:namePart id="6A7D3EB13035CB30E47F7AFF6727F10D">Alexiou, Sotiris</mods:namePart>
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<mods:affiliation id="4F56E11459AA135D58433DF0AC896CDF">Korinthian Museum of Natural History, Isthmion 201, Korinthos 20100, Greece</mods:affiliation>
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<mods:namePart id="22B464A79D41B17719F0A7F3609FE3E5">Bakolitsas, Kostas</mods:namePart>
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<mods:affiliation id="2AB9B800D7EF4D0D70AC920FC8A77622">Laboratory of Natural Sciences 3 rd General Lyceum, 301 31 Agrinion, Greece</mods:affiliation>
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<mods:affiliation id="E87282B3EAE77581322818951C164A9D">Dipartimento di Biologia e Biotecnologie “ C. Darwin ”, Università di Roma “ La Sapienza ”, Roma, Italy</mods:affiliation>
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<mods:affiliation id="798239D16CD40AB5CC1B4B283E31A08C">Dipartimento di Biologia e Biotecnologie “ C. Darwin ”, Università di Roma “ La Sapienza ”, Roma, Italy</mods:affiliation>
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<mods:date id="7261F21C1224F89F4E5A425343553950">2024</mods:date>
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<subSubSection id="50DC9498A01B7333BB3B4C69EC701D3B" type="nomenclature">
<paragraph id="7F67099C6C126AE96F556C039AD759FF">
<taxonomicName id="EB1CF9D5B96D6034C5A99BF5A615F7D2" authority="Alexiou" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="kotsabasi" status="sp. nov.">
<emphasis id="76E45A7C5B90067C84D5E19E5FC55A8A" italics="true">Dolichopoda kotsabasi</emphasis>
Alexiou
</taxonomicName>
<taxonomicNameLabel id="34DAE73CC8398C4DB9642FADD483B71B" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6ED690087A12E3A27F270321286E0422" type="description">
<paragraph id="25D4AB4EA09211B4995BB5858F6DE3C7">
<figureCitation id="A8AFC432AC41B64C165DA883A058C52B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographic distribution of D. athosensis sp. nov., D. dirussoi sp. nov., D. karoutsosi sp. nov. and D. kotsabasi sp. nov." figureDoi="10.3897/contrib.entomol.74.e121614.figure1" httpUri="https://binary.pensoft.net/fig/1103848">Figs 1</figureCitation>
,
<figureCitation id="BD8BABDCF3719522CE70910E57406D0A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Dolichopoda kotsabasi sp. nov.: A. X tergum, male, dorsal view; B. Subgenital plate, male, ventral view; C. Epiphallus, dorsal view; D. Epiphallus, lateral view; E. Subgenital plate, female, ventral view; F. Ovipositor, lateral view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure7" httpUri="https://binary.pensoft.net/fig/1103854">7</figureCitation>
</paragraph>
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<paragraph id="1D2E7E8730F93A1F71678C39D3831D11">
<heading id="9A5A5A5AAC5642FABB20804F60D27018" reason="title">Type material.</heading>
</paragraph>
<paragraph id="3634DE3BDB0397338CE445C1D871F63D">
<materialsCitation id="862F68F07E443159E33CDEC2D28F521B" collectingDate="2022-09-24" collectorName="S. Alexiou &amp; K. Kotsabas" country="Greece" county="cave Alama" elevation="370" latitude="37.602066" location="Samou" longLatPrecision="1" longitude="26.180267" municipality="Akamatra" specimenCount="371" specimenCount-male="1" typeStatus="holotype">
<emphasis id="D83A41E387FEACCFC6663FC8E0100B7D" bold="true" italics="true">
<typeStatus id="8A2DC42055A2284783DF17F664DF9C71" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="8D16C23ADA2327F38F530A35EC3ABE12" name="Greece">GREECE</collectingCountry>
•
<specimenCount id="EBA4A85685C09471AC2EAC101A8FC4AB" count="1" type="male">♂</specimenCount>
; Νomos
<location id="565F3608C302ADC8CD40A1D192BE7D14" LSID="urn:lsid:plazi:treatment:6B18655057BE5BF7B105FA47E165B89A:565F3608C302ADC8CD40A1D192BE7D14" country="Greece" county="cave Alama" latitude="37.602066" longLatPrecision="1" longitude="26.180267" municipality="Akamatra" name="Samou">Samou</location>
,
<location id="5A77D4D23AB9AAE3B991674A2939D300" LSID="urn:lsid:plazi:treatment:6B18655057BE5BF7B105FA47E165B89A:5A77D4D23AB9AAE3B991674A2939D300" country="Greece" county="cave Alama" latitude="37.602066" longLatPrecision="1" longitude="26.180267" municipality="Akamatra" name="Ikaria Isl.">Ikaria Isl.</location>
, near
<collectingMunicipality id="2E4C00047A2538CC17F8816627FE63E6">Akamatra</collectingMunicipality>
,
<collectingCounty id="21871B71A79B4A183B1C61082E5BE815">cave Alama</collectingCounty>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIOAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[26.180267,37.602067]}">
<geoCoordinate id="4BDFA865BB0B11CF6E1156A894513864" degrees="37" direction="north" minutes="36.124" orientation="latitude" precision="1" value="37.602066">37 ° 36.124 ' N</geoCoordinate>
,
<geoCoordinate id="2A8C15C816FCE681B2C176BC9DCC422C" degrees="26" direction="east" minutes="10.816" orientation="longitude" precision="1" value="26.180267">26 ° 10.816 ' E</geoCoordinate>
</named-content>
;
<specimenCount id="499FA2D21CF949E893738072BF985C02" count="370" type="generic">
<quantity id="81C398620621A0DFAF15970D1614A911" metricMagnitude="2" metricUnit="m" metricValue="3.7" unit="m" value="370.0">
<elevation id="4969635E43D4374DECC49B21C7636AE2" metricMagnitude="2" metricUnit="m" metricValue="3.7" unit="m" value="370.0">370 m</elevation>
</quantity>
.
</specimenCount>
asl. ;
<date id="2616A9618531CDD32CFE3B3D543AC522" value="2022-09-24">
<collectingDate id="C44B3DF8ADCEE81493FABF2E836ECD27" value="2022-09-24">24 Sep. 2022</collectingDate>
</date>
;
<collectorName id="C69619A00E0B0BBA3ACEBF05AE6D1B9C">S. Alexiou</collectorName>
&amp;
<collectorName id="77DC0D713AB9AA4355A0F9EAF04A34E2">K. Kotsabas</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0ENOAE" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
.
<materialsCitation id="04CF36E63B87085D1B711ED330C408F2" collectingDate="2022-09-24" collectorName="S. Alexiou &amp; K. Kotsabas" country="Greece" county="cave Alama" elevation="370" latitude="37.602066" location="Samou" longLatPrecision="1" longitude="26.180267" municipality="Akamatra" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="1A2BCE9A6E1B098CFEFC0AECABB5D165" bold="true" italics="true">
<typeStatus id="720DA5553AD66589E59EA43E39B000F7" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="1CDAA6AEE4E928ABA556BBE8F065E74F" name="Greece">GREECE</collectingCountry>
•
<specimenCount id="7FC5D97409DED9FD7F2DEC061A022AE1" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="6636EB1F671254DA0310FEAA3818AEA8" count="1" type="female">1 ♀</specimenCount>
, same data as for holotype
</materialsCitation>
.
<emphasis id="7AA16D6D34DC0365FCFB48B9625B0174" bold="true" italics="true">Other material</emphasis>
:
<materialsCitation id="95D59FF92FDD23A66235C14C5E40518C" collectingDate="2022-09-24" collectorName="S. Alexiou &amp; K. Kotsabas" country="Greece" county="cave Raos Choutra" location="Samou" municipality="Petropouli" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCountry id="D98D5CBEA3B1081D9706F5FA4374ECE2" name="Greece">GREECE</collectingCountry>
•
<specimenCount id="AF4059E99767D5793CD47338119F2378" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="8A48DB0A7D32DA5F9610566E9619E4B6" count="1" type="female">1 ♀</specimenCount>
; Νomos
<location id="D5FAC5E32F6F3B3B4528DE88294D78AB" LSID="urn:lsid:plazi:treatment:6B18655057BE5BF7B105FA47E165B89A:D5FAC5E32F6F3B3B4528DE88294D78AB" country="Greece" county="cave Raos Choutra" municipality="Petropouli" name="Samou">Samou</location>
,
<location id="05D40E4D39CC205FDFBDBDA271BD50FE" LSID="urn:lsid:plazi:treatment:6B18655057BE5BF7B105FA47E165B89A:05D40E4D39CC205FDFBDBDA271BD50FE" country="Greece" county="cave Raos Choutra" municipality="Petropouli" name="Ikaria Isl.">Ikaria Isl.</location>
,
<collectingMunicipality id="8CA7BFB269B6B67F8BBD6D43451A5339">Petropouli</collectingMunicipality>
,
<collectingCounty id="58C5FDC42140C6354612CC3BA08F330F">cave Raos Choutra</collectingCounty>
;
<date id="3E3EC4BEB4A7AD6DB52E354D7DADA6E7" value="2022-09-24">
<collectingDate id="8E326F044654BA4BC048AE819DF985A4" value="2022-09-24">24 Sep. 2022</collectingDate>
</date>
;
<collectorName id="CBB5AA89E305538F4F4E5764C3470531">S. Alexiou</collectorName>
&amp;
<collectorName id="21BB9853A4DF37469C61E8835018E02F">K. Kotsabas</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0EYOAE" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4OAE" type="diagnosis">
<paragraph id="E6A7846441ADBF201AE2519975B3953B">
<heading id="E3CBDF9456ADC2DAE2C31C16F7741566" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="30296D3A09C647C09B202B4DA8BAE4BE">
The new species shows a strong resemblance to the already known species of the central
<collectingRegion id="A4BDB9937536DCC73DE183AFCA1C9D73" country="Greece" name="Voreio Aigaio">Aegean</collectingRegion>
area. It is mostly similar to the two cave crickets endemic on the nearby island of Samos,
<taxonomicName id="753FAAF95BCD24D3A6BE7322ACCEA115" authority="Rampini &amp; Di Russo, 2012" authorityName="Rampini &amp; Di Russo" authorityYear="2012" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="giulianae">
<emphasis id="7A31CDDCD63A8A535F61ED088CDA29BB" italics="true">D. giulianae</emphasis>
<bibRefCitation id="91580CD4743560CAF579ED206260E9F7" DOI="10.3897/zookeys.201.2609" author="Rampini M &amp; Di Russo C &amp; Taylan MS &amp; Gelosa A &amp; Cobolli M" journalOrPublisher="ZooKeys" pagination="43-58" refId="B13" refString="Rampini M, Di Russo C, Taylan MS, Gelosa A, Cobolli M (2012) Four new species of Dolichopoda Bolivar, 1880 from Southern Sporades and Western Turkey (Orthoptera, Rhaphidophoridae, Dolichopodainae). ZooKeys 201: 43 - 58. https: // doi. org / 10.3897 / zookeys. 201.2609" title="Four new species of Dolichopoda Bolivar, 1880 from Southern Sporades and Western Turkey (Orthoptera, Rhaphidophoridae, Dolichopodainae)." volume="201" year="2012">Rampini &amp; Di Russo, 2012</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="5124D8C85C8020EA7BDDD1ACD205950A" authority="Di Russo &amp; Rampini, 2012" authorityName="Di Russo &amp; Rampini" authorityYear="2012" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="kalithea">
<emphasis id="356F61B99BA25BA152AEDAA19BC6F540" italics="true">D. kalithea</emphasis>
Di Russo &amp;
<bibRefCitation id="6CA6F0CD00FD654992975A487CC1A78E" DOI="10.3897/zookeys.201.2609" author="Rampini M &amp; Di Russo C &amp; Taylan MS &amp; Gelosa A &amp; Cobolli M" journalOrPublisher="ZooKeys" pagination="43-58" refId="B13" refString="Rampini M, Di Russo C, Taylan MS, Gelosa A, Cobolli M (2012) Four new species of Dolichopoda Bolivar, 1880 from Southern Sporades and Western Turkey (Orthoptera, Rhaphidophoridae, Dolichopodainae). ZooKeys 201: 43 - 58. https: // doi. org / 10.3897 / zookeys. 201.2609" title="Four new species of Dolichopoda Bolivar, 1880 from Southern Sporades and Western Turkey (Orthoptera, Rhaphidophoridae, Dolichopodainae)." volume="201" year="2012">Rampini, 2012</bibRefCitation>
</taxonomicName>
, as well as to the Anatolian
<taxonomicName id="170D3D0AA7FBC621A7CF2B82B74BBC49" authority="Rampini &amp; Taylan, 2012" authorityName="Rampini &amp; Taylan" authorityYear="2012" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="sutini">
<emphasis id="83290271931A65F23FD7E102CF803D6A" italics="true">D. sutini</emphasis>
<bibRefCitation id="770BDC3D8E0E601BCAD71837AAD65F24" DOI="10.3897/zookeys.201.2609" author="Rampini M &amp; Di Russo C &amp; Taylan MS &amp; Gelosa A &amp; Cobolli M" journalOrPublisher="ZooKeys" pagination="43-58" refId="B13" refString="Rampini M, Di Russo C, Taylan MS, Gelosa A, Cobolli M (2012) Four new species of Dolichopoda Bolivar, 1880 from Southern Sporades and Western Turkey (Orthoptera, Rhaphidophoridae, Dolichopodainae). ZooKeys 201: 43 - 58. https: // doi. org / 10.3897 / zookeys. 201.2609" title="Four new species of Dolichopoda Bolivar, 1880 from Southern Sporades and Western Turkey (Orthoptera, Rhaphidophoridae, Dolichopodainae)." volume="201" year="2012">Rampini &amp; Taylan, 2012</bibRefCitation>
</taxonomicName>
, the females especially being very similar (
<bibRefCitation id="6E87BF6F46C6F79AC709B7021083830B" DOI="10.3897/zookeys.201.2609" author="Rampini" etAl="et al." firstAuthor="Rampini" journalOrPublisher="ZooKeys" pagination="43-58" refId="B13" refString="Rampini M, Di Russo C, Taylan MS, Gelosa A, Cobolli M (2012) Four new species of Dolichopoda Bolivar, 1880 from Southern Sporades and Western Turkey (Orthoptera, Rhaphidophoridae, Dolichopodainae). ZooKeys 201: 43–58. https://doi.org/10.3897/zookeys.201.2609" title="Four new species of Dolichopoda Bolivar, 1880 from Southern Sporades and Western Turkey (Orthoptera, Rhaphidophoridae, Dolichopodainae)." volume="201" year="2012">Rampini et al. 2012</bibRefCitation>
). The males of both species of Samos have a more robust and arched median process of the epiphallus than
<taxonomicName id="56B4675AE2D73588ED24FD222368813C" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="kotsabasi" status="sp. nov.">
<emphasis id="7BA0C66520360B78BBFDE35693B18ABD" italics="true">D. kotsabasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AA371258E656386CB7CDE77C9FBC944C" rank="species">sp. nov.</taxonomicNameLabel>
Furthermore, the males of
<taxonomicName id="C6FA5D362D2658CEB57CB2BB67101B7C" authorityName="Di Russo &amp; Rampini" authorityYear="2012" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="kalithea">
<emphasis id="30C78263209846E88BAF64A4A975AF37" italics="true">D. kalithea</emphasis>
</taxonomicName>
have trapezoidal, not globular, lobes at the subgenital plate and
<taxonomicName id="E9267FA6F13FB1B78F8796ED8BDE43D6" authorityName="Rampini &amp; Di Russo" authorityYear="2012" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="giulianae">
<emphasis id="1142EFBF06B260D4211940FA30E7584E" italics="true">D. giulianae</emphasis>
</taxonomicName>
have squared lobes at the X tergum.
<taxonomicName id="4F2CEC188B2ABAE94FC26F27228B626C" authorityName="Rampini &amp; Taylan" authorityYear="2012" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="sutini">
<emphasis id="C4A2AD2AB41717469A3359493850B4F5" italics="true">Dolichopoda sutini</emphasis>
</taxonomicName>
also presents a differently shaped median process of the epiphallus as well as differently shaped lateral lobes of the subgenital plate of the males. The new species shows very similar median process of the epiphallus with
<taxonomicName id="98C82BD267F6FD4B3E37710664187CA1" authorityName="Rampini &amp; Di Russo" authorityYear="2012" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="calidnae">
<emphasis id="E9B0FAC2BC2D2A3810052098111ABA12" italics="true">D. calidnae</emphasis>
</taxonomicName>
, endemic to Kalymnos Island,
<taxonomicName id="3134A21D725E4C9A9F36C023519254A7" authorityName="Boudou-Saltet" authorityYear="1973" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="paraskevi">
<emphasis id="AB141E704AE894C989A805A6306BF874" italics="true">D. paraskevi</emphasis>
</taxonomicName>
, endemic to
<collectingRegion id="300B868F29DF898B12DC432001201A5B" country="Greece" name="Kriti">Crete</collectingRegion>
, as well as to
<taxonomicName id="73D0610DA23AC3A800E030BD4DF09AE4" authorityName="Boudou-Saltet" authorityYear="1973" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="naxia">
<emphasis id="D2E818CF6B17729EF4C7C7DD2735FE53" italics="true">D. naxia</emphasis>
</taxonomicName>
, endemic to Naxos Island, being elongated, more slender and less arched. However, it shows significant differences in most other characters for both sexes.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVSAE" type="description">
<paragraph id="C5C849C66857587A99917A2168C3654A">
<heading id="2DD82915898548406A61E27C95947A76" reason="title">Description.</heading>
</paragraph>
<paragraph id="FF507A9FEB91DBDEF9FEAC1000437A4D">Body pale testaceous, posterior margin of the terga darker. Hind femora unarmed, hind tibiae with 24 / 25 spines on both sides.</paragraph>
<paragraph id="3C72EE6EEDAB6397BBC84E7FB370FA6B">Fore tibiae with 4 spines on both sides of ventral side, mid tibiae with 5 spines on both sides of ventral side and hind tibiae with 19 spines on both sides of dorsal side.</paragraph>
<paragraph id="0A41536BD77303C676E995F06AAF3669">
X tergum (last abdominal tergite) (Fig.
<figureCitation id="404A208471A8ED53169BE3E516B24C58" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Dolichopoda kotsabasi sp. nov.: A. X tergum, male, dorsal view; B. Subgenital plate, male, ventral view; C. Epiphallus, dorsal view; D. Epiphallus, lateral view; E. Subgenital plate, female, ventral view; F. Ovipositor, lateral view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure7" httpUri="https://binary.pensoft.net/fig/1103854">7 A</figureCitation>
) slightly incised in the middle, with two lateral triangular lobes, lobes with rounded apex.
</paragraph>
<caption id="8AB934608C2A6D881162F1E0B2E9F9D4" ID-DOI="10.3897/contrib.entomol.74.e121614.figure7" ID-arpha="59540CDC-AFC7-54E5-95F4-537B3949C6FA" httpUri="https://binary.pensoft.net/fig/1103854" startId="F7">
<paragraph id="C2CAF489C1C7BFE22673EBC1D6976A25">
<label id="4D84A6D5C88C8D93575425D71867B217">Figure 7.</label>
</paragraph>
<paragraph id="32737ECFAF2486128D18B36E513F4907">
<taxonomicName id="282DD51C603499DBAD6BD7B5EE49460B" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="kotsabasi" status="sp. nov.">
<emphasis id="16CD4680F79EF4060A621F9AC8958826" italics="true">Dolichopoda kotsabasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C333B022D32A922783F3385678DF5B0C" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="F02EDD479EF6839178DD56CC8A9DB572" bold="true">A.</emphasis>
X tergum, male, dorsal view;
<emphasis id="51B01760FC7CA5EA9477254EFCED51A6" bold="true">B.</emphasis>
Subgenital plate, male, ventral view;
<emphasis id="FB81E4E963BBC7406A436DB052E46F0B" bold="true">C.</emphasis>
Epiphallus, dorsal view;
<emphasis id="46C55145CB7A785DA9E278E5E458E11A" bold="true">D.</emphasis>
Epiphallus, lateral view;
<emphasis id="9881421DA722049FA72F3716557753A1" bold="true">E.</emphasis>
Subgenital plate, female, ventral view;
<emphasis id="9B71E91B0CF25C9048CF721C4F2671A2" bold="true">F.</emphasis>
Ovipositor, lateral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="6E81527A17A0157FF81CCDE80CCD8E8B">
Subgenital plate (Fig.
<figureCitation id="4B5A7A5328DE6C35C0589F87BB1EC16D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Dolichopoda kotsabasi sp. nov.: A. X tergum, male, dorsal view; B. Subgenital plate, male, ventral view; C. Epiphallus, dorsal view; D. Epiphallus, lateral view; E. Subgenital plate, female, ventral view; F. Ovipositor, lateral view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure7" httpUri="https://binary.pensoft.net/fig/1103854">7 B</figureCitation>
) globular, divided by a median triangular incision, with short lateral lobes ; styli short, inserted almost at the apex of the plate, where an incision occurs.
</paragraph>
<paragraph id="C5873DFEEFB0CF76055C0900516A79DF">
Epiphallus sclerotized, with an acute cylindrical median process. Median process slightly curved inwards, laterally robust at the base (Fig.
<figureCitation id="65587A7FB6C2F355EE148FC33F9C851F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Dolichopoda kotsabasi sp. nov.: A. X tergum, male, dorsal view; B. Subgenital plate, male, ventral view; C. Epiphallus, dorsal view; D. Epiphallus, lateral view; E. Subgenital plate, female, ventral view; F. Ovipositor, lateral view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure7" httpUri="https://binary.pensoft.net/fig/1103854">7 C, D</figureCitation>
).
</paragraph>
<paragraph id="AC8BFE3C0C53D1240E8BA062628382A3">Measurements (length in mm): body 17, pronotum 4, fore femora 15, middle femora 16, hind femora 26.</paragraph>
<paragraph id="57628E89F17C0D1CBB7E6C2521B23372">
<emphasis id="64AE90B1B10A48F5596696A3CA166738" bold="true">Female.</emphasis>
As in male. Subgenital plate rounded, slightly incised in the middle (7 E). Ovipositor
<quantity id="CAA6E8CAD9B8E5EA5BC45CDAF1E48872" metricMagnitude="-3" metricUnit="m" metricValue="9.0" unit="mm" value="9.0">9 mm</quantity>
, robust at the base, slightly arched upwards, lower valves with 15 denticles (7 F).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWUAE" type="etymology">
<paragraph id="E6EB4E8D9DE0695FAF9A6A4B5E5D505A">
<heading id="5ACB0D9892A694CBB1267E44A521F100" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="E6554E186D2734071045B87B13E0546E">The new species is named after botanist Konstantinos Kotsabas, a researcher of the flora of Ikaria who was the first to observe the cave crickets on the island.</paragraph>
</subSubSection>
<subSubSection id="SECID0E2UAE" type="distribution">
<paragraph id="770861DFDCD60737520F7AC55D8C14E8">
<heading id="062DB92DD1FBD4E43721277BB142A7A7" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="7DD219277A50C3D94596BB4E5F5FC0ED">Known only from two caves on Ikaria Island. A third population of presumably the same species was observed by Konstantinos Kotsabas at Chalavra Katafigi, inside a narrow cave opening underneath the chapel of Agios Nikolaos.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="2358F8748C4553636A8A4980D538FA7D">Description of six new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from northeastern India</mods:title>
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<mods:affiliation id="4C23DBA0A609E6E7C35264C65BF9F320">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India</mods:affiliation>
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<mods:affiliation id="723457748101333C6A1E2B296ACA0897">SM Sehgal Foundation Center for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka 560064, India &amp; Yenepoya Research Centre, Yenepoya University, Derlakatte, Mangalore, India</mods:affiliation>
</mods:name>
<mods:name id="68511DF08A77F1481C818EF8D7EA93A6" type="personal">
<mods:role id="FF67E8EE9A70D6D6A0F3B0AF38FCE0DA">
<mods:roleTerm id="ED42BA52069CEE88512AE56A30831F19">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5CF51156547EA48FE5EEBB0CF856FDD2">Lalronunga, Samuel</mods:namePart>
<mods:nameIdentifier id="B5F882BAB62E1C7C57B9C00A760E5FBA" type="ORCID">0000-0002-7542-4815</mods:nameIdentifier>
<mods:affiliation id="463612D41B6FB52AC46A3B1A2D46A105">Holy Child Society, Nalkata, Tripura 799263, India</mods:affiliation>
</mods:name>
<mods:name id="74636169D9AE65BEC281B75D5A7E919A" type="personal">
<mods:role id="B8290A77DFE243F4669390C45798FD38">
<mods:roleTerm id="F5FA1A5A655E913463E19BC2C9373C8C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="04A616FA04FE212B1D579A8CFA84D3A6">Deepak, V.</mods:namePart>
<mods:nameIdentifier id="12F3EAE2404BC9E0304F71783B129360" type="ORCID">0000-0002-8826-9367</mods:nameIdentifier>
<mods:affiliation id="5BDCF2797885D27DFED7F34608E80C89">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK &amp; School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne NE 1 7 RU, UK</mods:affiliation>
</mods:name>
<mods:name id="183D86B985A7034B036DAD524EFE64FE" type="personal">
<mods:role id="E506A41835CD0E98445A53080697C7FA">
<mods:roleTerm id="E53FB784299C877F1124471B1851BC48">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BF732EA4B49880D250C948486FDEA423">Das, Abhijit</mods:namePart>
<mods:nameIdentifier id="365A23C7A2733B980BB976B5D3B28BDF" type="ORCID">0000-0002-5851-8457</mods:nameIdentifier>
<mods:affiliation id="84C6612204D601E188ADBE35381F35C6">Wildlife Institute of India, Chandrabani, Dehradun, Uttarakhand 248001, India</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D5310254772667E547F230EDF3AA02AB">text</mods:typeOfResource>
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<mods:titleInfo id="A7B2BF26D45710A56838B8F7B5DC6646">
<mods:title id="38C7302DFAFDA621679A52E1683CF8F8">Vertebrate Zoology</mods:title>
</mods:titleInfo>
<mods:part id="D6E3D6BB88A82C4E57BEDBADB1E19034">
<mods:date id="18853C51FBD60744E0EFED7DD5E4A65B">2024</mods:date>
<mods:detail id="867A6CD3097644709EACA97A12C26AEC" type="pubDate">
<mods:number id="62AE2E5F05B7E81EA4E97994DD56A23C">2024-07-29</mods:number>
</mods:detail>
<mods:detail id="D62D63B360FE4E265850C3F96C1B0BBE" type="volume">
<mods:number id="B8DE4D54AACF26E823121D1D00AFF1BA">74</mods:number>
</mods:detail>
<mods:extent id="25D2F8D026E0C71F545394495BE44684" unit="page">
<mods:start id="D737F0D2A2A6163B84576E0F4ED75C99">453</mods:start>
<mods:end id="D53B2BF2E407643D2BEAFF1869D8D82D">486</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5B7E4B5BBC19D557B9A56129601CF216">journal article</mods:classification>
<mods:identifier id="F8D775CD82F9DEBF541B6B5DD5848519" type="DOI">10.3897/vz.74.e124752</mods:identifier>
<mods:identifier id="D5847E85843C8579A404ADE2F1F3A80B" type="ZooBank">154CE236-EFA8-4411-834B-234A9B45A63F</mods:identifier>
</mods:mods>
<treatment id="739EA53FD4D75FBFB2DB62ECAC41A8D3" ID-DOI="http://doi.org/10.5281/zenodo.13124029" ID-Zenodo-Dep="13124029" ID-arpha="739EA53F-D4D7-5FBF-B2DB-62ECAC41A8D3" ID-zoobank="https://zoobank.org/9AEE0E40-7983-444E-AC19-53AA8CE709D7" LSID="urn:lsid:zoobank.org:act:9AEE0E40-7983-444E-AC19-53AA8CE709D7" LSID-ZBK="urn:lsid:zoobank.org:act:9AEE0E40-7983-444E-AC19-53AA8CE709D7" httpUri="http://treatment.plazi.org/id/739EA53FD4D75FBFB2DB62ECAC41A8D3">
<subSubSection id="0620ACE781A8C5C6502441820332457C" type="nomenclature">
<paragraph id="9AD0A380D7E710BABE78DAA9F73C7314">
<taxonomicName id="A262AFE602D052D548130A495EF28B85" authority="Boruah, Narayanan, Deepak &amp; Das" authorityName="Boruah, Narayanan, Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="manipurensis" status="sp. nov.">
<emphasis id="5F6825074104D3D05E622FB1A649480D" italics="true">Cyrtodactylus manipurensis</emphasis>
Boruah, Narayanan, Deepak &amp; Das
</taxonomicName>
<taxonomicNameLabel id="254E1AFE97EDCE19ECA6EE7F51E2706E" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D2161ED2478E1BB0BF02AFD40429C881" type="description">
<paragraph id="26DA8AA246533D811DAB7CC584707F37">
<figureCitation id="BD8A1F61B3ED25E46954B11637FDF38A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype (WII-ADR 1596) of Cyrtodactylus manipurensis sp. nov. in preserved condition. A dorsal and B ventral view of the whole body; C ventral, D dorsal, and E lateral view of head; F dorsal enlarged tubercles on trunk; G ventral view of right manus and H of right pes. Scale bar: 10 mm." figureDoi="10.3897/vz.74.e124752.figure6" httpUri="https://binary.pensoft.net/fig/1102272">Figure 6</figureCitation>
;
<tableCitation id="BB9F8FA5D98B21FC30C1FD9304EE523D" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">Tables 2, S 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4JBK" type="Holotype">
<paragraph id="3B20C361D030E972EFF1AD88FD7767D1">
<heading id="FD983ACF905F03882FCCFEE8703D4B86" reason="title">
<typeStatus id="68FD44D14155E64934AA7EEFD57140A0" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="7C7FC2740E2792CE77E38D3F0305C7F6">
<materialsCitation id="71FDF20A7AA196EF57454C30D8C25E2E" collectingDate="2022-07-25" collectorName="Bitupan Boruah" country="India" elevation="1240" latitude="24.5954" location="Lamdan Kabui village" longLatPrecision="7" longitude="93.7085" municipality="Churachandpur District" specimenCount="1" stateProvince="Manipur" typeStatus="holotype">
Adult male (WII-ADR 1596), collected near
<location id="8F00AAB846E65BB43BBD8234773AACF6" LSID="urn:lsid:plazi:treatment:739EA53FD4D75FBFB2DB62ECAC41A8D3:8F00AAB846E65BB43BBD8234773AACF6" country="India" latitude="24.5954" longLatPrecision="7" longitude="93.7085" municipality="Churachandpur District" name="Lamdan Kabui village" stateProvince="Manipur">Lamdan Kabui village</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGKBK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[93.708500,24.595400]}">
<geoCoordinate id="FC129D6F8811F998299FAFA55751AAAF" degrees="24.5954" direction="north" orientation="latitude" precision="5" value="24.5954">24.5954 ° N</geoCoordinate>
;
<geoCoordinate id="135C4070C755FF39B31244D5A0A83E7F" degrees="93.7085" direction="east" orientation="longitude" precision="5" value="93.7085">93.7085 ° E</geoCoordinate>
</named-content>
;
<elevation id="F457AF4BC5E3225C609CAE2D90D14A26" metricMagnitude="3" metricUnit="m" metricValue="1.24" unit="m" value="1240.0">
elevation
<quantity id="5EEDD9284A49F4519CDE640A64782588" metricMagnitude="3" metricUnit="m" metricValue="1.24" unit="m" value="1240.0">1240 m</quantity>
a. s. l.
</elevation>
) (Fig.
<figureCitation id="E2D6F7413813B8D1C4E1C4454B925817" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 A</figureCitation>
),
<collectingMunicipality id="1CF99C873034C1D5EA7ED5DA665E0CD6">Churachandpur District</collectingMunicipality>
,
<collectingRegion id="DFFDA889672F15D6C6D1CBB4725D107E" country="India" name="Manipur">Manipur</collectingRegion>
,
<collectingCountry id="CAEC4F48F5B0B8EA15B9E2FE2E56797E" name="India">India</collectingCountry>
collected by
<collectorName id="FD5A199EA7E7D5A89B5F6F8BD100A6D7">Bitupan Boruah</collectorName>
on
<date id="20C7547AA0FDA64DADCCCD6E2DF26872" value="2022-07-25">
<collectingDate id="120B6E55E61EC70876242F49B817DD55" value="2022-07-25">25 July 2022</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption id="F16245DA920B26476EC6EEA9E97726F5" ID-DOI="10.3897/vz.74.e124752.figure6" ID-arpha="6008AF31-8C27-58D6-A030-7C0FDA0091C3" httpUri="https://binary.pensoft.net/fig/1102272" startId="F6">
<paragraph id="D808E1DEB1C90A739CDDB307A90F9EF2">
<label id="7525362A0A7DCABCD6DA88E762F1751B">Figure 6.</label>
</paragraph>
<paragraph id="207D35A611B5CB03FBCCAF9B9E250833">
Holotype (WII-ADR 1596) of
<taxonomicName id="8FEB927D81B4E1AE8772B450D6D5AA61" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="manipurensis" status="sp. nov.">
<emphasis id="CD2B94B8D0A25A362A3004FFAE821B27" italics="true">Cyrtodactylus manipurensis</emphasis>
</taxonomicName>
<emphasis id="291F3E7F758A04A143EF013E70034DBE" bold="true">
<taxonomicNameLabel id="07FC4F52168AA403C160DC265AF56220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in preserved condition.
<emphasis id="B26A7BE1011DAD81B8242043CE20E981" bold="true">A</emphasis>
dorsal and
<emphasis id="20A07F716706B71D89C3E9098E0C1DA5" bold="true">B</emphasis>
ventral view of the whole body;
<emphasis id="94DB13A12AD8E161CDE10A682AF37D9F" bold="true">C</emphasis>
ventral,
<emphasis id="AB2A952086B3F04C4423B0C7C4DC2A91" bold="true">D</emphasis>
dorsal, and
<emphasis id="B0F6A2623A3C113E40409F7E07D8A9CE" bold="true">E</emphasis>
lateral view of head;
<emphasis id="6A8D129ABF12DDFEA063673D88C726BB" bold="true">F</emphasis>
dorsal enlarged tubercles on trunk;
<emphasis id="3A9CE39224AFF71BF911C8667B92054B" bold="true">G</emphasis>
ventral view of right manus and
<emphasis id="08043CFDAFEA7D2A0554FE392FBFF5B2" bold="true">H</emphasis>
of right pes. Scale bar: 10 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EZLBK" type="diagnosis">
<paragraph id="F9598474E104AA50B5DF13F9541C4A6A">
<heading id="ABA52E5444DE6B3B9D5B520164CC8052" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="9C85782A7E52A0A02DBE879347D4A0E6">
Medium-sized gecko,
<abbrev id="ABBRID0E6LBK" xlink_title="snout to vent length">SVL</abbrev>
at least
<quantity id="13A2A17A165DFF12D28FD943ADB36F4C" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
in adult male; 10 supralabials; eight or nine infralabials; 21 bluntly conical and feebly keeled tubercles across midbody; 37 paravertebral tubercles; 36 mid-ventral scales between ventrolateral folds; 11 or 12 subdigital lamellae beneath digit IV of manus; 13–16 subdigital lamellae beneath digit IV of pes; seven precloacal pores arranged in a continuous series; six irregular shaped dark-brown cross bands on back between axilla and groin; tail dorsally with eight dark-brown and seven pale-brown bands arranged alternatively.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDMBK" type="description">
<paragraph id="BBF5D4BCEEEC3221FF515998BD98A122">
<heading id="E748ECAA9B232EF4C991092CF2C5E0EC" reason="title">
Description of the
<typeStatus id="6E492EB195EC776A6973633F66FC3CC6" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="7805DF5F0F726AF71D35CD90E823F0BC">
Specimen well preserved except an incision below left axilla ventrolaterally. Snout-vent length
<quantity id="346813F6713831BC1E8F6B20C29202DD" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
. Head moderately large (
<abbrev id="ABBRID0EJMBK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
/
<abbrev id="ABBRID0ENMBK" xlink_title="snout to vent length">SVL</abbrev>
= 0.26), oval, dorsoventrally depressed, longer than width (
<abbrev id="ABBRID0ERMBK" xlink_title="maximum head width">HW</abbrev>
/
<abbrev id="ABBRID0EVMBK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.71), distinct from neck, broader at occipital region; snout tip rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat; a longitudinal furrow on dorsal surface of the snout; snout short (
<abbrev id="ABBRID0EZMBK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
/
<abbrev id="ABBRID0E4MBK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.39), longer than orbit (
<abbrev id="ABBRID0EBNBK" xlink_title="orbit diameter, taken between the anterior and posterior bony orbital borders">OD</abbrev>
/
<abbrev id="ABBRID0EFNBK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
= 0.68); nostril nearly rounded, opening directed posterolaterally; ear opening rounded; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous; supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at the anterodorsal region; rostral wide, a short groove at the middle on top, rostral connected with nasals, supranasals, an internasal and first supralabials, two scales between the supranasals paced longitudinally, granular scales at parietal, occipital and temporal region intermixed with slightly large rounded granular tubercles starting from the level of posterior margin of the upper eyelids, those on temporal region are slightly larger than that of occipital region, size of the tubercles increases towards nape; 10 supralabial scales on both sides, size decreases towards angle of jaw; 8 supralabials up to midorbit on both sides; mental nearly triangular, connected with first infralabials, inner postmentals; nine infralabials on right side and eight on left side, size decreases towards angle of jaw; first infralabials connected with mental, second infralabial, inner and outer postmentals; inner pair of postmentals are larger than the outer postmentals; two rows of slightly enlarged scales along the infralabials starting below the outer postmentals, posteriorly size of those decreases; rest of the gular scales are small, granular juxtaposed, nearly homogeneous, size increases towards the throat where they become imbricate.
</paragraph>
<paragraph id="30FFD2E2B8F4D3D2DF344688D63DB837">
Habitus slender (
<abbrev id="ABBRID0ELNBK" xlink_title="maximum body width">BW</abbrev>
/
<abbrev id="ABBRID0EPNBK" xlink_title="snout to vent length">SVL</abbrev>
= 0.16,
<abbrev id="ABBRID0ETNBK" xlink_title="trunk length, between the axilla and groin">TRL</abbrev>
/
<abbrev id="ABBRID0EXNBK" xlink_title="snout to vent length">SVL</abbrev>
= 0.49), dorsoventrally depressed, dorsal scales granular, rounded, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, up to third segment of the tail, size increases towards posterior body and pronounced at the base of tail; 21 dorsal tubercles across mid dorsum; 37 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate; 36 mid-ventral scales between ventrolateral fold; seven precloacal pores arranged in an inverted “
<collectionCode id="8C1D06DE5B66CCDEEDFA7078CCC7410A" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
” shaped continuous series, followed by a series of five unpored enlarged scales below it, largest at the apex, an unpored scale equal size to
<abbrev id="ABBRID0E2NBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
present in continuous with the
<abbrev id="ABBRID0E6NBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
in both ends; scales above the
<abbrev id="ABBRID0EDOBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
larger than those of belly scales.
</paragraph>
<paragraph id="05AC5AEBC4111AD726E81BD0C90F7DDB">
Forelimbs and hindlimbs slender (
<abbrev id="ABBRID0EJOBK" xlink_title="forearm length, taken from the elbow to the wrist">FL</abbrev>
/
<abbrev id="ABBRID0ENOBK" xlink_title="snout to vent length">SVL</abbrev>
= 0.14,
<abbrev id="ABBRID0EROBK" xlink_title="crus length, taken from the knee to the base of the foot">CL</abbrev>
/
<abbrev id="ABBRID0EVOBK" xlink_title="snout to vent length">SVL</abbrev>
= 0.17), digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae; lamellae beneath digit IV of right and left manus (given as basal + distal) is 6 + 5 and 6 + 6 respectively; lamellae beneath digit IV of both right and left pes (given as basal + distal) is 6 + 7 and 7 + 9 respectively; dorsal scales on forelimbs heterogeneous is size, mostly granular; proximal scales on upper arm are smaller than that of lower arm; scales near elbow on lower arm smooth and subimbricate; upper arm scales granular at proximal part and at the distal end it is smooth, cycloid and imbricate; dorsal scales of hindlimbs heterogeneous, intermixed with slightly enlarged, feebly keeled and bluntly conical tubercles; scales on inner lateral side of the thighs are smooth, large and subimbricate, those on dorsal side are small granular; scales on tibia are small, granular juxtaposed; ventral scales of forelimbs granular, juxtaposed, heterogeneous; scales on palm heterogeneous in shape and size, granular juxtaposed; ventral scales on hindlimbs heterogeneous; most of the thigh scales are smooth, cycloid and subimbricate, but on the knee and below the level of precloacal pores, scales are smaller and granular; scales above the vent granular; tibia scales smooth, nearly homogeneous, cycloid and subimbricate; scales on soles heterogeneous, granular, juxtaposed to subimbricate.
</paragraph>
<paragraph id="ABAF3048690B3A2467BE741056B7DC7D">
Tail complete (
<abbrev id="ABBRID0E2OBK" xlink_title="tail length, from the cloaca to the tail tip">TL</abbrev>
=
<quantity id="8C62E474DCF3E2E573890F7D39D5431B" metricMagnitude="-2" metricUnit="m" metricValue="6.7" unit="mm" value="67.0">67 mm</quantity>
), slender, gradually tapering towards tip, segments indistinct, dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size, large feebly keeled scales up to third segment of the tail, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; no enlarged plate like series of subcaudal scales; three bluntly conical spurs on both sides of the tail base.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6OBK" type="Colouration in preservative">
<paragraph id="85DE975F549A46DA3873F69FF1F400A7">
<heading id="DF8D82EED3EFA575FB1656D564E8E475" reason="title">Colouration in preservative.</heading>
</paragraph>
<paragraph id="034F666B53845C87330C3623536B8E92">Head on top and laterally brown; upper eyelids grey; dorsal ground colour of neck and back slightly paler than that of head; dark-brown irregular patches on neck up to fore limb insertion level; six irregular shaped dark-brown cross bands on back between axilla and groin, mid-dorsally interrupted; one dark-brown band on sacrum; tail dorsally with eight dark-brown and seven pale-brown bands arranged alternatively; dark bands are comparatively broader than the pale bands; the first dark-brown band on tail base broken mid-dorsally into two enlarged elongated spots. Ventrally head, neck, trunk and limbs light-brown; tail with irregular dark-brown specks.</paragraph>
</subSubSection>
<subSubSection id="SECID0EEPBK" type="comparisons">
<paragraph id="BFF96A897D806F58673E87C3E2CADD37">
<heading id="23B5D12D1F725938AD8863BFE7EEE5AF" reason="title">Comparison.</heading>
</paragraph>
<paragraph id="08A32520CFD2707E32F1986E5A7C653B">
<taxonomicName id="F2C8697266240254000C0F76CF95A23D" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="manipurensis" status="sp. nov.">
<emphasis id="FAAE0E34454F8289D46AE0E4D9AEADA9" italics="true">Cyrtodactylus manipurensis</emphasis>
</taxonomicName>
<emphasis id="9F8D96EAC9F5D6E6C56E47FC59059042" bold="true">
<taxonomicNameLabel id="6A431120FBE5C4BEA78660482502D655" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="D21A32565BBA539E7D61EC8ABDE728D5" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aaronbaueri">
<emphasis id="A4956D154ADE6751699FD8607E485BE6" italics="true">C. aaronbaueri</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0ECQBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0EGQBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="C8BD8C3A8533C2393BD50C7FD47F9A5F">DTR</collectionCode>
</abbrev>
22–28); differs from
<taxonomicName id="BC404615932E38FE5E76DDD3BD6F1436" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aunglini">
<emphasis id="649A3634CE861C7DB67F44D3BF41B02B" italics="true">C. aunglini</emphasis>
</taxonomicName>
by fewer mid-ventral scale rows,
<abbrev id="ABBRID0EVQBK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="73CD4B4C06EFA043009EA6CFE3991F0B">MVSR</collectionCode>
</abbrev>
36 (vs.
<abbrev id="ABBRID0EZQBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="149FB05B122FF4FAEA0D7650AC160502">DTR</collectionCode>
</abbrev>
47–49), by fewer precloacal pores,
<abbrev id="ABBRID0E4QBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0EBRBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
12 or 13); differs from
<taxonomicName id="27B5F42F5D270AAB251A8ADFB4C5047A" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="barailensis" status="sp. nov.">
<emphasis id="A47058AF00962AEF68E4F8CBC82D0854" italics="true">C. barailensis</emphasis>
</taxonomicName>
<emphasis id="DE449E1CD91A90F3DB56E83D3BEAC7C8" bold="true">
<taxonomicNameLabel id="B2348BDB2D072179BABCE6E55EB008FF" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by number of dorsal tubercle rows,
<abbrev id="ABBRID0ESRBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0EWRBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="26CFFF408CA30237CDB7B2F5FC116A68">DTR</collectionCode>
</abbrev>
17), number of paravertebral tubercle rows,
<abbrev id="ABBRID0E1RBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
37 (vs.
<abbrev id="ABBRID0E5RBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="5EC03ABB24555729C29EFE0CF40CF9D2">PVT</collectionCode>
</abbrev>
32); differs from
<taxonomicName id="24F89A370325CC81C6BB9F1E5AB8CD98" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="bengkhuaiai">
<emphasis id="2C11DC776872C5982B45EF267D9251F6" italics="true">C. bengkhuaiai</emphasis>
</taxonomicName>
by fewer dorsal tubercle rows,
<abbrev id="ABBRID0ENSBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0ERSBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="B49923EE72542A772EFC25BF4A2493F5">DTR</collectionCode>
</abbrev>
22–26); differs from
<taxonomicName id="3154D65BD6A875B1FDD3E08580D69309" authorityName="Bauer" authorityYear="2002" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="brevidactylus">
<emphasis id="24B3BF80F3DE1D0FEC8846873975F9BC" italics="true">C. brevidactylus</emphasis>
</taxonomicName>
by having fewer dorsal tubercle rows,
<abbrev id="ABBRID0EATBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0EETBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="9C4A4F6AEB43725F476340579123C3A7">DTR</collectionCode>
</abbrev>
27–30), enlarged chocolate-brown patches on head and back absent (vs. present); differs from
<taxonomicName id="2AC5C3524ACBE6CCB5E4AF5A87DAE35A" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="chrysopylos">
<emphasis id="C52360A5CA627FCA244064F44B58BCAD" italics="true">C. chrysopylos</emphasis>
</taxonomicName>
by much smaller body size,
<abbrev id="ABBRID0ETTBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="E743E589A53ABB4EB0752BACCAE47C3C" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
(vs.
<abbrev id="ABBRID0EXTBK" xlink_title="snout to vent length">
<collectionCode id="D6622687119C1A338A6F27F9B2C6AC43">SVL</collectionCode>
</abbrev>
<quantity id="342C0ECD33F49391AA69E28BA8FB4BE9" metricMagnitude="-2" metricUnit="m" metricValue="7.2" metricValueMax="7.909999999999999" metricValueMin="6.49" unit="mm" value="72.0" valueMax="79.1" valueMin="64.9">64.9–79.1 mm</quantity>
in male), fewer precloacal pores,
<abbrev id="ABBRID0E2TBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs
<abbrev id="ABBRID0E6TBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
8–13); differs from
<taxonomicName id="18C5317C08FB0DE488DAF2AD7722C212" authorityName="Liu &amp; Rao" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="dianxiensis">
<emphasis id="9793AE215740729FC5DCAB227BF3FCF9" italics="true">C. dianxiensis</emphasis>
</taxonomicName>
by smaller body size,
<abbrev id="ABBRID0EOUBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="79EC13178443A19605C780E536412309" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
(vs.
<abbrev id="ABBRID0ESUBK" xlink_title="snout to vent length">
<collectionCode id="780AB5955949C8E03DF8033C5A2CD97A">SVL</collectionCode>
</abbrev>
<quantity id="B94A7FE1669E5BDECF03502894DA6366" metricMagnitude="-2" metricUnit="m" metricValue="7.685" metricValueMax="7.99" metricValueMin="7.38" unit="mm" value="76.85" valueMax="79.9" valueMin="73.8">73.8–79.9 mm</quantity>
), 21 rows of dorsal tubercle rows (vs.
<abbrev id="ABBRID0EWUBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="6B8802CEE14E236A88A9589EFAA65313">DTR</collectionCode>
</abbrev>
18 or 19); differs from
<taxonomicName id="F92AE2BDA49D0E05D1350AB272BED886" authorityName="Bauer" authorityYear="2003" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="gansi">
<emphasis id="445860ABF17DF766B5A41DA44D456CCF" italics="true">C. gansi</emphasis>
</taxonomicName>
by fewer precloacal pores,
<abbrev id="ABBRID0EFVBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0EJVBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
16–29), ventrolateral fold present on trunk (vs. absent); differs from
<taxonomicName id="954E22EF8E3D538C4EE78440EC7B76A4" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="jaintiaensis">
<emphasis id="8FEB6E60965B5C171910AA8B3C0F1985" italics="true">C. jaintiaensis</emphasis>
</taxonomicName>
by much smaller body size,
<abbrev id="ABBRID0EYVBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="FC5101E69C6AD649DCBD3D27FAA4A436" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
(vs.
<abbrev id="ABBRID0E3VBK" xlink_title="snout to vent length">
<collectionCode id="BA5FF1B68466A96765F81818789D0BBE">SVL</collectionCode>
</abbrev>
87.0–
<quantity id="288EF0CD8B9433AF692717A943233F3D" metricMagnitude="-2" metricUnit="m" metricValue="8.83" unit="mm" value="88.3">88.3 mm</quantity>
in male), fewer precloacal pores in male,
<abbrev id="ABBRID0EAWBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0EEWBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
11 or 12); differs from
<taxonomicName id="326C2C53733A585F8E1BE009097240DB" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="20679E38A689B23EB4BBFC20830FE339" italics="true">C. kiphire</emphasis>
</taxonomicName>
<emphasis id="4A496A8F8AA822EA6444F8C6ADC8CC35" bold="true">
<taxonomicNameLabel id="5C46630D0A80FC02E9B4D7D90DBCA073" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by smaller body size,
<abbrev id="ABBRID0EVWBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="179348035F6C676C48C904A2A067700C" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
(
<abbrev id="ABBRID0EZWBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="B67EFF2C9AA3FF7850289E4DAA9EA09F" metricMagnitude="-2" metricUnit="m" metricValue="6.43" metricValueMax="6.47" metricValueMin="6.39" unit="mm" value="64.3" valueMax="64.7" valueMin="63.9">63.9–64.7 mm</quantity>
), number of dorsal tubercle rows,
<abbrev id="ABBRID0E4WBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0EBXBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="BEC4CD02DEF3BE237EE966F9FD08C0A4">DTR</collectionCode>
</abbrev>
16), number of paravertebral tubercle rows,
<abbrev id="ABBRID0EFXBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">PVT</abbrev>
37 (vs.
<abbrev id="ABBRID0EJXBK" xlink_title="paravertebral tubercles on the trunk, counted between the level of the axilla and the level of the groin">
<collectionCode id="6A688FBFD62BB82C8F4F5E85110395A4">PVT</collectionCode>
</abbrev>
26 or 29); differs from
<taxonomicName id="239C2FCBD42D744120D8CA20DDDEC6B4" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="lungleiensis">
<emphasis id="A1CE98F6EE16ECCEAFC75C32A0969F7D" italics="true">C. lungleiensis</emphasis>
</taxonomicName>
by higher number of precloacal pores,
<abbrev id="ABBRID0EYXBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0E3XBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
<quantity id="B0F27D983C18812485F5B65A61B76CE6" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">3–5 in</quantity>
male), fewer dorsal tubercle rows,
<abbrev id="ABBRID0EAYBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs. 24–28); differs from
<taxonomicName id="942C21A573571D22879667502953DC50" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="montanus">
<emphasis id="0417EBB504BD1F9860D5DB9B1D0C6A6A" italics="true">C. montanus</emphasis>
</taxonomicName>
by larger body size,
<abbrev id="ABBRID0EPYBK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="1A803DA34AE1607445548588E5A288DC" metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
(vs. 53.6–55.0 mm in male); differs from
<taxonomicName id="8C4476CEF25977CF4AF9734C3C56C052" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="myaleiktaung">
<emphasis id="0626093DBCA48BBF875CE2023F0B1BC5" italics="true">C. myaleiktaung</emphasis>
</taxonomicName>
by fewer mid-ventral scale rows,
<abbrev id="ABBRID0E5YBK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="64317A82276EA08210050DADB603C5F8">MVSR</collectionCode>
</abbrev>
36 (vs.
<abbrev id="ABBRID0ECZBK" xlink_title="mid-ventral scale rows, counted between ventrolateral folds at the midpoint between axilla and groin">
<collectionCode id="07A2C3AB1BCE576B1335ED5D98C2CFD5">MVSR</collectionCode>
</abbrev>
57), precloacal pores present (vs.
<abbrev id="ABBRID0EGZBK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
absent), broad regular dark bands absent on dorsum (vs. present); differs from
<taxonomicName id="1587C6320E6F9D7205868D8A9B176B0F" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="nagalandensis">
<emphasis id="FE7B3FF8D2ABA32823D88A776217E646" italics="true">C. nagalandensis</emphasis>
</taxonomicName>
by higher number of dorsal tubercle rows,
<abbrev id="ABBRID0EVZBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">DTR</abbrev>
21 (vs.
<abbrev id="ABBRID0EZZBK" xlink_title="dorsal tubercle rows, counted transversely across the body at the midpoint between axilla and groin">
<collectionCode id="DBF5EE27CCAD33CEE116B2D83B3F03D1">DTR</collectionCode>
</abbrev>
16–18); differs from
<taxonomicName id="DD92636A43960E6F56645EB61E3570D1" authorityName="Mahony &amp; Kamei" authorityYear="2021" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="namtiram">
<emphasis id="8105F1FAAE33942DFB5848D2EE578168" italics="true">C. namtiram</emphasis>
</taxonomicName>
by having fewer precloacal pores,
<abbrev id="ABBRID0EI1BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0EM1BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
12); differs from
<taxonomicName id="784386F859640E9085523CBA733884E2" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngopensis">
<emphasis id="8BF7E6901C7535C4ED37B63A305103E1" italics="true">C. ngopensis</emphasis>
</taxonomicName>
by the number of precloacal pores,
<abbrev id="ABBRID0E21BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0E61BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
6); differs from
<taxonomicName id="75D0F1876C856C6B0078E7123B23230B" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siahaensis">
<emphasis id="B623C3E85FADB9FD306808E36F6E93A5" italics="true">C. siahaensis</emphasis>
</taxonomicName>
by be fewer subdigital lamellae, 11 or 12 lamellae under fourth finger (vs. 13 or 14 lamellae under fourth finger), dark-brown broad bands consists of two series of enlarged spots on dorsum (vs. dark-brown blotches in the form of reticulation on dorsum); differs from
<taxonomicName id="5823E40797750B7F8CBEEB74A25DC822" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="vairengtensis">
<emphasis id="C092F38654F54858C9E4CB8AB77E80F8" italics="true">C. vairengtensis</emphasis>
</taxonomicName>
by fewer precloacal pores,
<abbrev id="ABBRID0EZ2BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
7 (vs.
<abbrev id="ABBRID0E42BK" xlink_title="precloacal pores, a continuous series of pore-bearing scales on the precloacal region that does not extend onto the thighs">PcP</abbrev>
9–11). Morphological differences with other members of
<taxonomicName id="12DB307EA7EA1DC6E7E220E0031DC732" baseAuthorityName="Jerdon" baseAuthorityYear="1870" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="C259E8D5EF271F9332EA7E67EBA34B30" italics="true">khasiensis</emphasis>
</taxonomicName>
group is presented in Table
<tableCitation id="1FE329B200BE8F8E2208046872CC71D7" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EP3BK" type="Sequence divergence">
<paragraph id="E11F7423E607E8B3E25EF97290D90A14">
<heading id="CE6B432ACA643337FB4AB0E7AF68D504" reason="title">Sequence divergence.</heading>
</paragraph>
<paragraph id="8E198900797B5288C374D675D9339C43">
<taxonomicName id="C92DFBFCB13F8CDDD1D41EDCA4242DDE" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="manipurensis" status="sp. nov.">
<emphasis id="A96FD26C492C02A422321EA0A4E0CB18" italics="true">Cyrtodactylus manipurensis</emphasis>
</taxonomicName>
<emphasis id="A410B67C0B1420DDC5DCDE889ED63C5C" bold="true">
<taxonomicNameLabel id="1F31FC271704EA45EEF504D74B5905D1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a high genetic divergence of 11.2–11.3 % from its closely related
<taxonomicName id="014FC2C4D1AE0D7D13EBA48090D00858" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="ngopensis">
<emphasis id="817D0220941C0471869BAE3FA598E251" italics="true">C. ngopensis</emphasis>
</taxonomicName>
, 10–10.3 % from
<taxonomicName id="C93CD769FAE042ADD49C62180ED9A53D" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="aaronbaueri">
<emphasis id="B84B98217FF80B4715C85AD0F1DC625C" italics="true">C. aaronbaueri</emphasis>
</taxonomicName>
, 10.2–10.5 % from
<taxonomicName id="B16371B8E905CFDAF6891C10C8B9DDFD" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="vairengtensis">
<emphasis id="189B270AC549CC7D703AD3BE17385383" italics="true">C. vairengtensis</emphasis>
</taxonomicName>
and 9.7–11.1 % from the
<taxonomicName id="0115F927457EE26D5B6BE03FA16AE5A6" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="montanus">
<emphasis id="EE94C4B9A5C45C87EBD78DB462D44195" italics="true">C. montanus</emphasis>
</taxonomicName>
. With other members of the clade,
<taxonomicName id="394A09A3E2E92E46EE95EB84996FA459" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="kiphire" status="sp. nov.">
<emphasis id="26C85D73B2D18FFCDD3830A05C170BDF" italics="true">C. kiphire</emphasis>
</taxonomicName>
<emphasis id="4E4CD84F441E1E9804404055F139113B" bold="true">
<taxonomicNameLabel id="A401388F9AFAF8D6D424E1EEA0130D14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a genetic divergence of 8.6 % and 23 % in the ND 2 gene.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E25BK" type="etymology">
<paragraph id="8EB34D7DE05395680F819129CCFF3559">
<heading id="47DC0361EC668A9B95E3D341D24FB14C" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="9F9D2A627E60E5F35A20C0D598482CE4">
This species is named after
<collectingRegion id="7196F82B0F88285AA3D4B49E13632965" country="India" name="Manipur">Manipur state</collectingRegion>
in
<collectingCountry id="B7D9FD37A9D115EECEAD5244B71CFAA7" name="India">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EA6BK" type="Suggested common name">
<paragraph id="D2D035E27C25B69A3B28C7EF91F8973A">
<heading id="46D344BE69F1B1C03A096AC3D848F597" reason="title">Suggested common name.</heading>
</paragraph>
<paragraph id="9F61F7917C160037EACD18ABE1A2FC52">
<collectingRegion id="DBEB75D324CB5A0DF5E6319CB3C67A07" country="India" name="Manipur">Manipur</collectingRegion>
bent-toed gecko.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EF6BK" type="distribution">
<paragraph id="95A0D7DC0F06A37507DFA1EB28737E9B">
<heading id="C0321E2E9C5BC2E7C462089782050778" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="C7E4C8B5541D29B294BF28958ABBB55A">
<taxonomicName id="F6931D89F9FAF8C5FBB519E2F19D66DD" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="manipurensis" status="sp. nov.">
<emphasis id="789F958F4A278FDB9C2FD4A1B5F1A719" italics="true">Cyrtodactylus manipurensis</emphasis>
</taxonomicName>
<emphasis id="BC311887EA894D357CB38CD93FB57B61" bold="true">
<taxonomicNameLabel id="4659FDB93C760BFACEBA6D278EC578EE" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently only recorded from the
<typeStatus id="933A15DFE99319D2D943B943FA2CA831">type</typeStatus>
locality. We collected a single individual at 18: 00 hrs on
<date id="2A81CCF026A391872CDC8CC0E4C271F2" value="2022-07-25">25 July 2022</date>
near Lamdan Kabui village. It was perched on a shrub at a height of approximately
<quantity id="A35F8395BB9B318B8A37F1FB7E2D4EE7" metricMagnitude="0" metricUnit="m" metricValue="1.5" unit="m" value="1.5">1.5 m</quantity>
, on the road connecting Leimatak and Charoikhullen. The habitat is secondary forest, with
<taxonomicName id="CCAEC143F246A4BBEEF1FD3A43B59A26" class="Liliopsida" family="Zingiberaceae" genus="Zingiber" kingdom="Plantae" order="Zingiberales" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="A8D395E30503CF911D2663EED735AED6" italics="true">Zingiber</emphasis>
sp.
</taxonomicName>
cultivation and settlements.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="6AE026E0AF7AF0649F628D99E0C598D3">New species and a fascinating diversity of Chironomidae (Diptera, Insecta) in and around an overlooked urban vernal pool</mods:title>
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<mods:namePart id="0A788A8037F4447718971156B35A3C39">Namayandeh, Armin</mods:namePart>
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<mods:namePart id="53ABB6993A462ACE66905AFD676EE888">James, Taylor</mods:namePart>
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<mods:affiliation id="257AC5294A711D8BC2348D8FDC876DAE">5597 New Meadow Drive, Ypsilanti, MI 48197, USA</mods:affiliation>
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<mods:namePart id="F0BB33986BF98E5568F0A249B732B1DB">Ghaderi, Edris</mods:namePart>
<mods:nameIdentifier id="07E656B57D5E0705A010C0B3D69DEED5" type="ORCID">0000-0002-9875-7735</mods:nameIdentifier>
<mods:affiliation id="1AB7D51BAC4DBD425D881A484D417044">Department of Fisheries Sciences, Faculty of Natural Resources, University of Kurdistan, Sanandaj, Iran &amp; Department of Fisheries and Aquatic Ecology, Faculty of Fisheries and Environmental Sciences, Gorgan University of Agricultural Sciences and Natural Resources, Gorgan, Iran</mods:affiliation>
</mods:name>
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<mods:namePart id="F9F1107C0D796F30337BE731B767F182">Vasquez, Adrian A.</mods:namePart>
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<mods:affiliation id="9CA607911C4E9A15CDD16882BB8CA13A">Department of Biology, College of Liberal Arts and Sciences, Mercer University, 1501 Mercer University Drive, Macon, GA 31207, USA</mods:affiliation>
</mods:name>
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<mods:namePart id="67D710EE353B3547ED23C92A83F8F3A9">Ram, Jeffrey L.</mods:namePart>
<mods:nameIdentifier id="AD3F70DD21FA2785496ACD285596798C" type="ORCID">0000-0002-1063-546X</mods:nameIdentifier>
<mods:affiliation id="E90F0AFAC2A80091D9BF89DCBB22B4AD">Department of Physiology, School of Medicine, Wayne State University, Detroit, MI 48201, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="2F165AFBC00A8C50EA7D0951FB6FACB6">text</mods:typeOfResource>
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</mods:titleInfo>
<mods:part id="953AB2AAD134BDDD99F76A6E55635D4D">
<mods:date id="888BEDF24F81517F853F32328C760827">2024</mods:date>
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</mods:mods>
<treatment id="877B68D2BDD15340A3E3E1E75489F301" ID-DOI="http://doi.org/10.5281/zenodo.13124009" ID-Zenodo-Dep="13124009" ID-arpha="877B68D2-BDD1-5340-A3E3-E1E75489F301" ID-zoobank="https://zoobank.org/38F6ED6F-9E46-4A00-93EE-0110BF969C6C" LSID="urn:lsid:zoobank.org:act:38F6ED6F-9E46-4A00-93EE-0110BF969C6C" LSID-ZBK="urn:lsid:zoobank.org:act:38F6ED6F-9E46-4A00-93EE-0110BF969C6C" httpUri="http://treatment.plazi.org/id/877B68D2BDD15340A3E3E1E75489F301">
<subSubSection id="DD79D1657617AC643C58531F214D5DEF" type="nomenclature">
<paragraph id="0DD86EA2B4978677C0E1DD9EFCD7BCC8">
<taxonomicName id="4DBCA6DFD4F55153FA7ACD4AF4082409" authority="Namayandeh, Guerra &amp; Ram" authorityName="Namayandeh, Guerra &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="stagnum" status="sp. nov.">
<emphasis id="2901B936F1B9BFA88A5EA32159991E52" italics="true">Limnophyes stagnum</emphasis>
Namayandeh, Guerra &amp; Ram
</taxonomicName>
<taxonomicNameLabel id="D0EF41F1293D57772240DC954E766691" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B7E6D4E65680360042B0C8EDA87D37E4" type="description">
<paragraph id="23F79687ACA982E13670F3510D5DD601">
<figureCitation id="64ADDD03A700B9E9C72E022CAC24C82E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Limnophyes stagnum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure8" httpUri="https://binary.pensoft.net/fig/1103619">Figs 8</figureCitation>
,
<figureCitation id="D7AA66D4BE786A78EB2AE364546572E5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Limnophyes stagnum sp. nov., adult female A head B thorax C genitalia ventral E genitalia dorsal. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure9" httpUri="https://binary.pensoft.net/fig/1103620">9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDPBG" type="type material">
<paragraph id="44213A7C4BEBA7C48CA9EA7A5F6A9406">
<heading id="039B2B983157E6AE760AB17BECBC5C42" reason="title">Type material.</heading>
</paragraph>
<paragraph id="6CA21814AC038BC597FDFD987548BBEC">
<materialsCitation id="6D95A651C7F4FEB103CF5968CD0BD94B" collectingDate="2023-11-29" collectionCode="ARC" collectorName="Pond A &amp; P. L. Hudson" country="United States of America" latitude="42.42766" location="Palmer Park" longLatPrecision="1" longitude="-83.11741" municipality="Detroit" specimenCount="1" specimenCount-male="1" stateProvince="Michigan" typeStatus="holotype">
<emphasis id="7DEAA2D42CEFDCCB32AFA437A7508741" bold="true" italics="true">
<typeStatus id="C9FBD067F8DC71068153AE7514B12E31" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="2CB86DFCA9CCCF5E34EB41135361686D" count="1" type="male">1 male</specimenCount>
;
<collectingCountry id="6C538D140899FB5EBF1292CEEBA7503D" name="United States of America">USA</collectingCountry>
,
<collectingRegion id="212484ED191B5025761DADDE5EA1D164" country="United States of America" name="Michigan">Michigan</collectingRegion>
,
<collectingMunicipality id="BCA306CDC0E4B43B5E18DDE70512D68F">Detroit</collectingMunicipality>
,
<location id="8386ED36C1BC0E3C7DB8F7CB74DAFC26" LSID="urn:lsid:plazi:treatment:877B68D2BDD15340A3E3E1E75489F301:8386ED36C1BC0E3C7DB8F7CB74DAFC26" country="United States of America" latitude="42.42766" longLatPrecision="1" longitude="-83.11741" municipality="Detroit" name="Palmer Park" stateProvince="Michigan">Palmer Park</location>
,
<collectorName id="3C09C024C5F4FC5D8148254AB8FA0D94">Pond A</collectorName>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERPBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.117410,42.427660]}">
<geoCoordinate id="60DA158D839B8D4B49BAA42FD5E40865" degrees="42.42766" direction="north" orientation="latitude" precision="1" value="42.42766">42.42766 ° N</geoCoordinate>
,
<geoCoordinate id="2A105E5C5AFEBD672D18770E2330A7DE" degrees="83.11741" direction="west" orientation="longitude" precision="1" value="-83.11741">83.11741 ° W</geoCoordinate>
</named-content>
; leg.
<collectorName id="73A88A7122D14D374F6DC2B8E89DD7C2">P. L. Hudson</collectorName>
;
<date id="731ECA9F0F3052F83179B933B5AD7FC2" value="2023-11-29">
<collectingDate id="CD58A0D8F405DBCF756B6CD6FD3D5E90" value="2023-11-29">29. xi. 2023</collectingDate>
</date>
, dep.
<abbrev id="ABBRID0EWPBG" xlink_title="Albert J. Cook Arthropod Research Collection">
<collectionCode id="3E61655A833900B0BDAC8BFADCEEF39A" country="Canada" httpUri="http://grbio.org/cool/r82x-h8re" name="Atlantic Reference Centre" type="">ARC</collectionCode>
</abbrev>
</materialsCitation>
.
<emphasis id="7882A3F00211893CBAF07D17DC0A6F1E" bold="true" italics="true">
<typeStatus id="345ACED00CC242033EC9674A4B8852D0" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="5904C70F2A5A583741A7425E9B51E600" count="2" type="male">2 males</specimenCount>
,
<specimenCount id="CF5C9259E96E18F283D7A0F39CF437A7" count="3" type="female">3 females</specimenCount>
; same as
<typeStatus id="2023193342B9BACA8927B82888A40B9A" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4PBG" type="description">
<paragraph id="BCE8451370DE11C627A978C7C96A53E4">
<heading id="EDB5B07F84B0CCD64C12B040AA5B036C" reason="title">Diagnostic characters.</heading>
</paragraph>
<paragraph id="7085DA885AD60064D2CA457B6E688CEA">
Adults of this species can be separated from other
<taxonomicName id="E8890236272BD51F9F2E424D27F1FBB0" authorityName="Eaton" authorityYear="1875" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="E40F51FEEF499747BC6FDE31B8393A13" italics="true">Limnophyes</emphasis>
</taxonomicName>
by the combination of the following characteristics: Adults with no thoracic lanceolate setae and 2 prescutellars; adult male without humeral setae, female with single humeral setae; male with 3 epimerons, 1 posterior anepisternum II, 1–2 preepisternals anteriorly parallel to antepronotum and close to anapleural suture; female with 2 posterior anepisternals II; 6 epimeron II; 11 preepisternals which 9 anteriorly clustered horizontally, separated from 2 vertical; male antenna with 10 flagellomeres and AR 0.86; female antenna with 4 flagellomere and AR 0.5; male anal point extremely short, almost receded, wide and triangular with apex rounded and gonostylus expanded evenly from base to apex; female with apodeme lobe not distinct; cercus pediform.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKQBG" type="description">
<paragraph id="7462B1E581A591380FD99B3D66AE7307">
<heading id="5523B721B3042EF8709289F6ACC3A870" reason="title">Description.</heading>
</paragraph>
<paragraph id="F9BBC99748F69253FA4DBFDE3B025EE0">
<emphasis id="F7F567A09A1E6949E0FFEB3A3568524B" bold="true">Male</emphasis>
(
<emphasis id="2FF1AF567530956FC40B0874DACAFC93" italics="true">n</emphasis>
= 3, unless otherwise indicated). Total length
<quantity id="FACF018E787B43D74DA41C9EE87EDD6A" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 mm</quantity>
. Wing
<quantity id="174AEF6A92502DB304B868ED36B54723" metricMagnitude="-3" metricUnit="m" metricValue="1.005" metricValueMax="1.1" metricValueMin="0.91" unit="mm" value="1.005" valueMax="1.1" valueMin="0.91">0.91–1.1 mm</quantity>
long and
<quantity id="651BFB9DC5F9BE564F6E2AF39BBC7038" metricMagnitude="-4" metricUnit="m" metricValue="3.0" unit="mm" value="0.3">0.3 mm</quantity>
wide.
</paragraph>
<paragraph id="681526A9A910331225E66699ED0E3F48">
<emphasis id="B8A0F2CF915C3B62752320A7EA14575C" bold="true">
<emphasis id="07A46F7376942E766992C9DA425C4529" bold="true" italics="true">Coloration</emphasis>
.
</emphasis>
Head, thorax, legs, tergites, sternites, and hypopygium blackish brown. Wings and halters grey.
</paragraph>
<paragraph id="F10CEC39562B1561E051BFB186D5FA4C">
<emphasis id="DC0067BAF3B68B59A1C64B337CBB0928" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="0FF43FC616879EC45CF71B38BC079D07" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Limnophyes stagnum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure8" httpUri="https://binary.pensoft.net/fig/1103619">8 A, B</figureCitation>
). Antenna with 10 flagellomeres, last flagellomere with 4 sensilla chaetica, groove starts at the apex of the second segment (Fig.
<figureCitation id="9577E0E4485B8532FD9D5EED037664B2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Limnophyes stagnum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure8" httpUri="https://binary.pensoft.net/fig/1103619">8 A</figureCitation>
), AR 0.86 (
<emphasis id="1AB91F62A09573843C2D3FA3E121C28F" italics="true">n</emphasis>
= 1). Eyes bare, without dorsomedial extension. Temporal setae 1 inner vertical (Fig.
<figureCitation id="1234DAA34B12953CD798058288A480CC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Limnophyes stagnum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure8" httpUri="https://binary.pensoft.net/fig/1103619">8 B</figureCitation>
). Tentorium 113–123, 118 μm long. Clypeus triangular, 82–95, 88 μm long and 106–125, 116 μm wide, bearing 8–10, 9 setae, setae 42–71, 56 μm long. Palpal segment lengths (in μm): 38, 36, 58, 54, 83 (
<emphasis id="D4BC6E8610F302158A311069C025EB64" italics="true">n</emphasis>
= 1).
</paragraph>
<caption id="CBD7DB91B6BAC058CC0EE7F28036DFEA" ID-DOI="10.3897/zookeys.1208.124495.figure8" ID-arpha="7102A1B7-BFD5-5ECD-B1BE-F9634DDA52EA" httpUri="https://binary.pensoft.net/fig/1103619" startId="F8">
<paragraph id="E94F8913B5C042B4EDA2232E2CD573F5">
<label id="602BD99F87BC71AA725FE04B66917C03">Figure 8.</label>
</paragraph>
<paragraph id="FE2B4D93C5F7E2B9BFCF23B00DA68F5C">
<taxonomicName id="5C42620A9E9D965744F7DAC1243848EC" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="stagnum" status="sp. nov.">
<emphasis id="235CECE25FE52690308310EB47235AEB" italics="true">Limnophyes stagnum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96CEBA0426CE09383E30788ECC3E0D76" rank="species">sp. nov.</taxonomicNameLabel>
, adult male
<emphasis id="3EA459DA2E4C08491E09C17360ABBF69" bold="true">A</emphasis>
antenna
<emphasis id="767C9BC7A0224E048047660A1F2C036E" bold="true">B</emphasis>
head
<emphasis id="3DDD921AE70AA4FA23F467EC9C205A77" bold="true">C</emphasis>
thorax
<emphasis id="062BD5A98BDEB5DC9FDC1997B35FE68B" bold="true">D</emphasis>
wing
<emphasis id="8FE0E8700232605D9B14B9BDD76655FA" bold="true">E</emphasis>
hypopygium. Scale bars: 100 µm.
</paragraph>
</caption>
<paragraph id="013F14BDF1AD181235263942111C75F3">
<emphasis id="C3CC21C0D19ECC54CE30980CD70B4E56" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="7DBC068BFD0EDAD19543BE3C6F28523D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Limnophyes stagnum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure8" httpUri="https://binary.pensoft.net/fig/1103619">8 C</figureCitation>
). Lanceolate setae absent. Acrostichals 6; dorsocentrals 9–10; prealars 5; scutellars
<quantity id="957C99936DC533DE5E244987E8424A05" metricMagnitude="-1" metricUnit="m" metricValue="1.016" unit="in" value="4.0">4 in</quantity>
single row; antepronotals 2; lanceolate humerals absent; 2 prescutellars; epimeron 3; posterior anepisternum II 1; 1–2 preepisternals anteriorly parallel to antepronotum and close to anapleural suture.
</paragraph>
<paragraph id="EEC021CDD6C5A37181D8209790A938EE">
<emphasis id="EA2468B1A5944FE29D0418DDBB9201D6" bold="true" italics="true">Wing</emphasis>
(Fig.
<figureCitation id="5C2864D77583DCD28D2557D62CDEC914" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Limnophyes stagnum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure8" httpUri="https://binary.pensoft.net/fig/1103619">8 D</figureCitation>
). Brachiolum with 1 seta. Squama with 2 setae. All veins bare. Costa extension 62 μm long (
<emphasis id="862A39E1CEAE44A825EF5F6646FAEE5F" italics="true">n</emphasis>
= 1). Anal lobe well-reduced. Microtrichia visible at 10 ×.
</paragraph>
<paragraph id="38985B11989E3C25879B30B15088887C">
<emphasis id="CD86311FC442CE6C29A1E5F155294339" bold="true">
<emphasis id="53EA6435040538A3450740787C86762D" bold="true" italics="true">Legs</emphasis>
.
</emphasis>
Fore tibia spur 36–45, 40 μm long, mid tibia spurs 16 and 14 μm long, hind tibia spurs 31 and 24 μm long, hind tibia comb with around 10 spines. Lengths and proportions of legs as in Table
<tableCitation id="6D3A52B503DCAD3DBB396F368B9FF45E" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Male leg lengths (μm) and proportions of Limnophyes stagnum sp. nov." httpUri="http://table.plazi.org/id/072D12A45F673371973D8978256799E9" tableUuid="072D12A45F673371973D8978256799E9">3</tableCitation>
.
</paragraph>
<caption id="072D12A45F673371973D8978256799E9" ID-Table-UUID="072D12A45F673371973D8978256799E9" httpUri="http://table.plazi.org/id/072D12A45F673371973D8978256799E9" startId="T3">
<paragraph id="66DAD6339100223CE5FDAFDF0F0C9477">
<label id="B0CD293353B48AD1E70891C0D77391BC">Table 3.</label>
</paragraph>
<paragraph id="730642FF844FEE747234527E4E515AA7">
Male leg lengths (μm) and proportions of
<taxonomicName id="C830E89A0F90E93DF2EA6158643A5A6F" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="stagnum" status="sp. nov.">
<emphasis id="59AD74849CA35927288D3290BE53CD2F" italics="true">Limnophyes stagnum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB4A0CD97D9168976D872961D6949C05" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="2D02514F511C3E1798681B935BB6BC05">
<table id="TID0EEZBI" rules="all">
<tbody id="5CA006C4E503B6CAF0D66E60B5CCF7D0">
<tr id="06E922E03D82226C7A6177AD62365B4B">
<th id="F5211F219B41BDCD2C364C1E8333E5D8" colspan="1" isEmpty="true" rowspan="1">-</th>
<th id="D0847D69DAB87E9C195CFA4A1D11FE18" colspan="1" rowspan="1">fe</th>
<th id="0587439D1C50CAF11551558E584F016F" colspan="1" rowspan="1">ti</th>
<th id="0D447C97519ED0E2B3B18AC9B9236DD1" colspan="1" rowspan="1">
ta
<subScript id="5A699A9A62AFDC4FADAB1BCB11875C9B">1</subScript>
</th>
<th id="41BA74EEC4BC2E34706386E986EA4453" colspan="1" rowspan="1">
ta
<subScript id="9FADE11E3BDD73F04D8C8A09AD6D0A20">2</subScript>
</th>
<th id="005832D3F498808087AF1FF32C253A9D" colspan="1" rowspan="1">
ta
<subScript id="75F62B537767F7FC4188F2A9938B8AE2">3</subScript>
</th>
<th id="C4861C92CA7F20AA3C5D277CF5EA8A8E" colspan="1" rowspan="1">
ta
<subScript id="EE6F627B5B30AFE516DE198DA4CB5E1C">4</subScript>
</th>
<th id="40091DED23CF1A795D6809BCB75F4CCE" colspan="1" rowspan="1">
ta
<subScript id="4C5DB6440AEA9FC258EBDFF3E0B2304D">5</subScript>
</th>
<th id="E096375E5E0EFFB40534652C510D4B27" colspan="1" rowspan="1">LR</th>
<th id="3CA2AF271367338DC99A09C46102FBBC" colspan="1" rowspan="1">BV</th>
<th id="33E5E8B160F5D3B19A0587FF0829D652" colspan="1" rowspan="1">SV</th>
</tr>
<tr id="C03349252D6CE160F6B8B7C881C013AE">
<td id="41350908876B0B38D6919D81865A2C01" colspan="1" rowspan="1">
<emphasis id="3D6139E2A508D52CD21B4328400DE6DB" bold="true">
P
<subScript id="0E57377F1B04867DCF6D9C34F51F333D">1</subScript>
</emphasis>
</td>
<td id="EE4C611121BA4EC0647384D8DA9B34FF" colspan="1" rowspan="1">469</td>
<td id="D8C52E5E3DE5869DE713056FBA4D8351" colspan="1" rowspan="1">541</td>
<td id="DBA90DA4B26658AF1B65D9167F1A819D" colspan="1" rowspan="1">278</td>
<td id="EB5BC5607D66C9AC64A9BD84F2204BD1" colspan="1" rowspan="1">163</td>
<td id="4F00E956575E5823987D6D8924EEDD80" colspan="1" rowspan="1">104</td>
<td id="44A30457453D169575D29FE0092DE652" colspan="1" rowspan="1">60</td>
<td id="4A3D12817B017201F9C11D49AD62C369" colspan="1" rowspan="1">69</td>
<td id="ED0FA4CDA633B244BAB958A5FE74919E" colspan="1" rowspan="1">0.50</td>
<td id="4EE3F394C1A5B5CDEEC3D40AB937B053" colspan="1" rowspan="1">3.3</td>
<td id="8E1B8BBA087888DA035749553BB88B2C" colspan="1" rowspan="1">3.6</td>
</tr>
<tr id="A828FF12AC193C319396604A80186DC8">
<td id="D5046838D68C55C0DD79D7975E731049" colspan="1" rowspan="1">
<emphasis id="38A076D75B8DF2F6BE3BE2EAB57F6924" bold="true">
P
<subScript id="715AAF52C10EC313D185ADCEEA9BD99E">2</subScript>
</emphasis>
</td>
<td id="1FCCAD9ADD4E0446385757AAE3AA9629" colspan="1" rowspan="1">496</td>
<td id="89AA8549205F748235F771B2C8F75A3B" colspan="1" rowspan="1">471</td>
<td id="67985A725945F04F30C2F04E6FA94819" colspan="1" rowspan="1">205</td>
<td id="EE9108806B67CDEF867EAA78279686A2" colspan="1" rowspan="1">116</td>
<td id="AF6F5F2E24C7F1E1887FFA3F45A9A7EC" colspan="1" rowspan="1">69</td>
<td id="128DB70A57699FB1496CCEB7DE101BC2" colspan="1" rowspan="1">47</td>
<td id="8BADEBED22F957731083D74C217B16E4" colspan="1" rowspan="1">63</td>
<td id="F7D4DD786494711226CC9E0D8A1798AD" colspan="1" rowspan="1">0.40</td>
<td id="6CCB05F82D5E050E90B8EE5460F6E7AB" colspan="1" rowspan="1">4.0</td>
<td id="3AB2966F1929539EF5085D19A7A40209" colspan="1" rowspan="1">4.7</td>
</tr>
<tr id="315F7572F72BDE548BF4EAD36BC4F1C8">
<td id="F2AD1DE20D34197693513EE019EA0646" colspan="1" rowspan="1">
<emphasis id="F14AFD50ECFB6B98A2B74F24272E7412" bold="true">
P
<subScript id="709E51E2A288C987CC6DEA022A018D5A">3</subScript>
</emphasis>
</td>
<td id="56024D251A240E92361C4748D0AC930D" colspan="1" rowspan="1">447</td>
<td id="AEA4BB84DCDC45C52D3215F679DFBA51" colspan="1" rowspan="1">515</td>
<td id="B1ACA09A17D9D2147EDE95FBF5AE8D93" colspan="1" rowspan="1">282</td>
<td id="30C3D6A68B915211D93801F4A5BA281B" colspan="1" rowspan="1">150</td>
<td id="0611A8FBDD7E217FF50189F900AAC35B" colspan="1" rowspan="1">118</td>
<td id="39C197A863FF524C5D077A4D075E0BBE" colspan="1" rowspan="1">41</td>
<td id="233A9CA5376A941765C76584574C25AD" colspan="1" rowspan="1">68</td>
<td id="5F4E2A40E60F9BC93DEDFD3D9FCE205E" colspan="1" rowspan="1">0.50</td>
<td id="4B7FE553F6E34BE508A019AC0B3CED84" colspan="1" rowspan="1">3.3</td>
<td id="1B01DCD866AC445288AE3E2C9DDB81B4" colspan="1" rowspan="1">3.4</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="F51A283BE941450282B6C0ED4A8F73AE">
<emphasis id="1A71728D206D7AF57E999B7B8ED8012C" bold="true" italics="true">Hypopygium</emphasis>
(Fig.
<figureCitation id="0C888D58ED70E14CDF6E8BC68F5B3A83" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Limnophyes stagnum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure8" httpUri="https://binary.pensoft.net/fig/1103619">8 E</figureCitation>
). Anal point extremely short, almost receded, wide, and triangular with apex rounded; anal point 6–15, 10 μm long and 14–34, 24 μm wide. Virga bifid and short, 11–12 μm long. Transverse sternapodeme with well-developed oral projections; sternapodeme 75–79, 77 μm long. Phallapodeme 28–40, 34 μm long. Inferior volsella small triangular lobe covered in numerous simple setae. Gonostylus expanded evenly from base to apex, 67–73, 70 μm long; crista
<taxonomicName id="2735D35D43E8F226DEE48B11AE451991" authorityName="Meigen" authorityYear="1818" class="Insecta" family="Chironomidae" genus="Chironomus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="EC55D04573EEB25B6E85100BD0FA38DB" italics="true">dorsalis</emphasis>
</taxonomicName>
very narrow. Gonocoxite 90–116, 103 μm long. HR 1.2–1.7, 1.5, HV 2.5–2.9, 2.7.
</paragraph>
<paragraph id="9AF36E176673049773A023AB874D14BB">
<emphasis id="E1701EC46E277A497F7E7E66D5CEEAAA" bold="true">Female</emphasis>
(
<emphasis id="EE1062A218DE49C297599300C2E44FEF" italics="true">n</emphasis>
= 3). Total length
<quantity id="53BFEDEDA8A8F17874C77A4773617B51" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
. Wing
<quantity id="994DAD0C106D3A06AD28899B91C3DE8D" metricMagnitude="-4" metricUnit="m" metricValue="9.6" unit="mm" value="0.96">0.96 mm</quantity>
long,
<quantity id="8BE71236B3837E6FD8330F09FE7C2F44" metricMagnitude="-4" metricUnit="m" metricValue="3.8" unit="mm" value="0.38">0.38 mm</quantity>
wide.
</paragraph>
<paragraph id="EBA7E06DED4FD7C931214BB6AEAE965C">
<emphasis id="54BC33D6AE42203058F55BDDB8BCCC39" bold="true">
<emphasis id="607D25588860E64BD2DB5B5389B61220" bold="true" italics="true">Coloration</emphasis>
.
</emphasis>
Same as the male.
</paragraph>
<paragraph id="67B1B191010D2DF99161BB3F387076A4">
<emphasis id="CA410913679F6EC7CDFBEFD4E05FDE8E" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="EFF261C3F728B3033F9338928035A316" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Limnophyes stagnum sp. nov., adult female A head B thorax C genitalia ventral E genitalia dorsal. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure9" httpUri="https://binary.pensoft.net/fig/1103620">9 A</figureCitation>
). Antenna with 4 flagellomeres, last flagellomere with 4 sensilla chaetica, Antennal segments 1–4 (in μm): 75–77, 76; 49, 47, 87, AR 0.5. Eyes bare. Temporal setae 2, including 1 inner vertical and 1 frontal. Tentorium 141–145, 143 μm long. Clypeus triangular, 59 μm long and 46 μm wide, bearing 13 setae, setae 37–59, 47 μm long. Palpal segment lengths (in μm): 46, 32, 52, 54, 75.
</paragraph>
<caption id="E1290CED11C0E047DBC255ABCE9C09B7" ID-DOI="10.3897/zookeys.1208.124495.figure9" ID-arpha="DEA6E524-3639-5C9B-AC30-F5FBC8AB2CFC" httpUri="https://binary.pensoft.net/fig/1103620" startId="F9">
<paragraph id="47193424ADE7CFB9615C0FA34B045945">
<label id="98E32378B5DDF448C394EBEB0C44EE08">Figure 9.</label>
</paragraph>
<paragraph id="75DF123C994A25DC6F09D7C48B791E69">
<taxonomicName id="3F16EFFE44CA1141EE373C351E0E910F" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="stagnum" status="sp. nov.">
<emphasis id="59A895FAB77BE64541ADC6B656A0F326" italics="true">Limnophyes stagnum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="245CCF9A90B3C7B40F97F1BB33C4D354" rank="species">sp. nov.</taxonomicNameLabel>
, adult female
<emphasis id="5E5F72FB8D6732CE81AE094E7053E995" bold="true">A</emphasis>
head
<emphasis id="7A631F28D076931435332AE2714016D5" bold="true">B</emphasis>
thorax
<emphasis id="253A7C57375D11B23B6806C5E59384A6" bold="true">C</emphasis>
genitalia ventral
<emphasis id="E08646C85C686F3379F3471B3142DAC1" bold="true">E</emphasis>
genitalia dorsal. Scale bars: 100 µm.
</paragraph>
</caption>
<paragraph id="A3A703F61343C2B1C7EE4DD24160F84C">
<emphasis id="7652C816CEF7ADE3F771CB0B1144830F" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="C5BCDBDF0331E50FF7965AAA44C167AD" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Limnophyes stagnum sp. nov., adult female A head B thorax C genitalia ventral E genitalia dorsal. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure9" httpUri="https://binary.pensoft.net/fig/1103620">9 B</figureCitation>
). Acrostichals 5; dorsocentrals 8–12,
<quantity id="3389D023B935AC9149E181B56C72B932" metricMagnitude="-1" metricUnit="m" metricValue="2.286" unit="in" value="9.0">9 in</quantity>
a single row; prealars 5; scutellars
<quantity id="9E58604536680167FBE8A53D49BE2844" metricMagnitude="-1" metricUnit="m" metricValue="1.016" unit="in" value="4.0">4 in</quantity>
single row; 1 humerals, non-lanceolate; 2 prescutellars non-lanceolate; 7 antepronotals; 2 posterior anepisternals II; 6 epimeron II; 11 preepisternals, 9 clustered horizontally close to epimeron, 4 horizontally arching close to anapleural suture.
</paragraph>
<paragraph id="5F008E50502CE800F98C0A50546B4742">
<emphasis id="70595BE50AAA941D580DA82779A7108D" bold="true" italics="true">Wing</emphasis>
(Fig.
<figureCitation id="87F23D4C1E346D4890CB30C0E925F2DF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Limnophyes stagnum sp. nov., adult female A head B thorax C genitalia ventral E genitalia dorsal. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure9" httpUri="https://binary.pensoft.net/fig/1103620">9 C</figureCitation>
). Brachiolum with 1 seta. Squama bare.
<collectionCode id="6DE826E345A6AA946DA561CA59F5E6B4" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
with 3–8, 5 setae,
<collectionCode id="FA5FED7DCAAA9EE229BD3374B5A53C83" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="AA682C0BBE0C3B0301286E8CC2F67308">1</subScript>
with 4–5 setae;
<collectionCode id="B44E241D847BCE43533D5B60767E0642" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="7AAFF4E2894F6E972CDF20008DAB77AF">4 + 5</subScript>
with 9–10 setae; other veins bare. Costa extension 50–74, 62 μm. Microtrichia visible at 10 × magnification.
</paragraph>
<paragraph id="85F55688FB0F74C8A6F66E0EFA009C1F">
<emphasis id="3713C8424DBBD8CA7C49F568884C1D5F" bold="true">
<emphasis id="6B728B6C164AED29F3B8AC3C71D71C14" bold="true" italics="true">Legs</emphasis>
.
</emphasis>
Hind and mid femur with keel. Fore tibia spur 22–23 μm long, mid tibia spur 21 μm long, second one broken; hind tibia missing. Lengths and proportions of legs as in Table
<tableCitation id="D1B0589330830774B22556ADD5595143" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Female leg lengths (μm) and proportions of Limnophyes stagnum sp. nov." httpUri="http://table.plazi.org/id/8593F9866B640773B3A8BB79F9BFAFEF" tableUuid="8593F9866B640773B3A8BB79F9BFAFEF">4</tableCitation>
.
</paragraph>
<caption id="8593F9866B640773B3A8BB79F9BFAFEF" ID-Table-UUID="8593F9866B640773B3A8BB79F9BFAFEF" httpUri="http://table.plazi.org/id/8593F9866B640773B3A8BB79F9BFAFEF" startId="T4">
<paragraph id="F5B1E70A5372493A505B9913F01F3832">
<label id="BC0EC66ACEF0C275D73DB67409BAA0FB">Table 4.</label>
</paragraph>
<paragraph id="E9DAA37799611991491E328C956B77DC">
Female leg lengths (μm) and proportions of
<taxonomicName id="75418F0F271625CFFCD532D9E8ADB1F4" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="stagnum" status="sp. nov.">
<emphasis id="53468A57FD368949B6FDE8F99634427A" italics="true">Limnophyes stagnum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9ABF3201E71A25761506665E669487D5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="246B96347C244870BF44AE32FF428B11">
<table id="TID0EFBCI" rules="all">
<tbody id="34322DE60755CC75F9D7AD25B2C3B658">
<tr id="899CDE6B40D98877B0B7356B6920AD8B">
<th id="9B215C22DCB84166821E4DB7A8DA25EC" colspan="1" isEmpty="true" rowspan="1">-</th>
<th id="4BF06C3D36A5C1FA41D7B73E510A15B1" colspan="1" rowspan="1">fe</th>
<th id="1A8A047C9F2168FD92FD07722F9AC553" colspan="1" rowspan="1">ti</th>
<th id="42B3B82E03275FBA2597F045304CBED2" colspan="1" rowspan="1">
ta
<subScript id="45BB2F650DFDA5566BB510A7531AF360">1</subScript>
</th>
<th id="543546F6519656894527440999C742AA" colspan="1" rowspan="1">
ta
<subScript id="DD2F6CED5A18CA8509AB5337AFA0EF57">2</subScript>
</th>
<th id="1D09D1FBD363B133462C694D12C2F529" colspan="1" rowspan="1">
ta
<subScript id="C98CC95FDA74F5B2B65CD243EBEC2D16">3</subScript>
</th>
<th id="1B93604E2B9239C23E4309AA33638F43" colspan="1" rowspan="1">
ta
<subScript id="3BFB65AB7336CBF78D688535677917B0">4</subScript>
</th>
<th id="32BF240C1A14A6FF874A265E689F8B0E" colspan="1" rowspan="1">
ta
<subScript id="B3CA78150EB6BB43B33CF9F4263ECB51">5</subScript>
</th>
<th id="4AF8C51B4BB1747CCD8FA5F93CCEFAA5" colspan="1" rowspan="1">LR</th>
<th id="4A5B2EDA954BD6D22A38EB7BA657DBD1" colspan="1" rowspan="1">BV</th>
<th id="E597AAD98E678F735D1A4028979120BE" colspan="1" rowspan="1">SV</th>
</tr>
<tr id="AE11A0A67211038FC6D7A405A44E766F">
<td id="6D0E33C244241BDC0D1832A17E75C99E" colspan="1" rowspan="1">
<emphasis id="FD572CC4ABE5E05F847B7D8131685D93" bold="true">
P
<subScript id="5D394AC0670994636EDF0202F49E1D73">1</subScript>
</emphasis>
</td>
<td id="CE448172ACF957D8994BC6A18D67B5DB" colspan="1" rowspan="1">358</td>
<td id="606BB8548D7D595D1D8BFF84A1C973F7" colspan="1" rowspan="1">440</td>
<td id="16BB69D6EB5A8F00CF33B0583A00FCB9" colspan="1" rowspan="1">224</td>
<td id="73A9C453265A53E8A64B8356CC78F6FE" colspan="1" rowspan="1">135</td>
<td id="A1C5CDF894EFC5B40DFE5E9C528740B4" colspan="1" rowspan="1">86</td>
<td id="22B924C286E56DFADAC227E193EE71BA" colspan="1" rowspan="1">51</td>
<td id="70E9C1E6F9B27380B364040120338BAC" colspan="1" rowspan="1">22</td>
<td id="8C895051223E4F264F9D4A0BFDC683A4" colspan="1" rowspan="1">0.50</td>
<td id="8995349FA32CFC7E73BFF3F19BD58516" colspan="1" rowspan="1">3.0</td>
<td id="8DD0BF3E391299D2B4FBD80F113FA82D" colspan="1" rowspan="1">3.6</td>
</tr>
<tr id="C908034BD10A1C7C1EC820F97F0E7922">
<td id="00F0D3FAAF7B6173518A8C47485B156C" colspan="1" rowspan="1">
<emphasis id="B3C703C4524E56B76B15B9CA655423AB" bold="true">
P
<subScript id="316BD94B601994AF405525ECC2456A1D">2</subScript>
</emphasis>
</td>
<td id="3B22381BBAA1243807433CD5D2AA3B6A" colspan="1" rowspan="1">440</td>
<td id="4AFB7AEBF0019EC397F2969AEB261F54" colspan="1" rowspan="1">427</td>
<td id="FD37AAD40CE8F8F81F91F0F2CA66F267" colspan="1" rowspan="1">190</td>
<td id="0203CA991B88D66821E82D6A002B1B7D" colspan="1" rowspan="1">90</td>
<td id="0B790BFD564F8A6AE2A1B7738A21C2B8" colspan="1" rowspan="1">61</td>
<td id="8F02D20C0C9B160C71CB514BC18F9B3F" colspan="1" rowspan="1">39</td>
<td id="8D9969889C626B8B6E4AE93913883066" colspan="1" rowspan="1">64</td>
<td id="C20DCA96E9AFF132F0823198B9075C8F" colspan="1" rowspan="1">0.40</td>
<td id="6782A5D59F9315C9B7131D7F982437D8" colspan="1" rowspan="1">4.2</td>
<td id="C7B0BED0DFF98DB856E002C216843103" colspan="1" rowspan="1">4.6</td>
</tr>
<tr id="A40AEAF05D54FD11F725D52C538EA416">
<td id="678FC32CCFF3010C8CA42D391C940BA6" colspan="1" rowspan="1">
<emphasis id="D9A7DFD93AFCAB7D1235CABCC9D23A9A" bold="true">
P
<subScript id="32DF68BDB84B74DADFE4845AB9111123">3</subScript>
</emphasis>
</td>
<td id="477F9B9A1421A6046392664A9194EA5F" colspan="1" rowspan="1">–</td>
<td id="C5031C35365561B475BDE16B6A0EA184" colspan="1" rowspan="1">–</td>
<td id="BFD5F1849F337796013B69BB41C33B2F" colspan="1" rowspan="1">–</td>
<td id="ADA980A848A07BFCC147568D92424C0B" colspan="1" rowspan="1">–</td>
<td id="52C7F5DC2B06B9331C8B12A45E042BBE" colspan="1" rowspan="1">–</td>
<td id="F2FC8E350376F18A91272043483DC144" colspan="1" rowspan="1">–</td>
<td id="4AD32EE2AE7FB32735121CBF33436F7C" colspan="1" rowspan="1">–</td>
<td id="BA4139B11E11D1F16672282DD6E3B74A" colspan="1" rowspan="1">–</td>
<td id="7E9513344E219E1380F60227C4198826" colspan="1" rowspan="1">–</td>
<td id="FFC8A54AD0F64C042CF7CAAAF8439576" colspan="1" rowspan="1">–</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="D5D12705DB60E2DBAF3EB0B0B5D39476">
<emphasis id="520E41576B1FD857B88A0A0F21E9DE40" bold="true" italics="true">Genitalia</emphasis>
(Figs
<figureCitation id="2A8D03D2744AA1C6569F8FC51475C869" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Limnophyes stagnum sp. nov., adult female A head B thorax C genitalia ventral E genitalia dorsal. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure9" httpUri="https://binary.pensoft.net/fig/1103620">9 D, E</figureCitation>
). Seminal capsules 62–89, 72 μm long, and 47–90, 71 μm wide, semi-circular; spermathecal very long, with well-developed bulb (Fig.
<figureCitation id="2B25EA91CAA75B1FEEBDE69C916459C9" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Limnophyes stagnum sp. nov., adult female A head B thorax C genitalia ventral E genitalia dorsal. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure9" httpUri="https://binary.pensoft.net/fig/1103620">9 D</figureCitation>
). Notum 76–113, 92 µm long, notum and ramus 143–191, 165 µm long. Gonapophysis VIII divided into ventrolateral and thin dorsomesal lobe (Fig.
<figureCitation id="000392B816A1FDBE7A06D42D59A1A92B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Limnophyes stagnum sp. nov., adult female A head B thorax C genitalia ventral E genitalia dorsal. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure9" httpUri="https://binary.pensoft.net/fig/1103620">9 D</figureCitation>
). Apodeme lobe not distinct. Gonocoxite developed, triangular with around 6–9 setae (Fig.
<figureCitation id="719226CF8425486FA5060624F9D695CB" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Limnophyes stagnum sp. nov., adult female A head B thorax C genitalia ventral E genitalia dorsal. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure9" httpUri="https://binary.pensoft.net/fig/1103620">9 E</figureCitation>
). Tergite IX undivided. Cercus small, pediform, 58–72, 65 µm long, and 42–54, 48 µm wide (Fig.
<figureCitation id="4647FD5331614709C704163E124BA380" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Limnophyes stagnum sp. nov., adult female A head B thorax C genitalia ventral E genitalia dorsal. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure9" httpUri="https://binary.pensoft.net/fig/1103620">9 E</figureCitation>
).
</paragraph>
<paragraph id="B64E2EA298255835D4D68CD5CD55C26D">
<emphasis id="05297DF3830445E87827C1A56EBB31F2" bold="true">Immatures.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1DAI" type="etymology">
<paragraph id="1BA8A154290AD76016762581A058CF6F">
<heading id="7F83B5C44EF7E5ADEE2D6A2AC282215A" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="953FDA45490841D96940335786246038">
The species is named after the habitat where it is found. The word
<taxonomicName id="6C74F4CC88D488BD238671873ABF20AA" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="stagnum">
<emphasis id="8CDB4740CC6855E787BF46529FA97610" italics="true">stagnum</emphasis>
</taxonomicName>
is Latin, meaning pond or pool.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKEAI" type="distribution">
<paragraph id="28FC2A473D6EFCDD8D46B6701C4E901A">
<heading id="4E1CB0E0DFEE380B85F05D8A6F4FBE01" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="9F278BF0B5D3DD5A782EA48FAE774751">
<collectingCountry id="5CA4493D235AC7E86C17A224AAABE947" name="Canada">Canada</collectingCountry>
(
<collectingRegion id="02ACEE0C289314427EA07ADCB95B0E64" country="Canada" name="British Columbia">British Columbia</collectingRegion>
,
<collectingRegion id="AA915BB2C30F66E1239FAE353BA56BE6" country="Canada" name="Ontario">Ontario</collectingRegion>
);
<collectingCountry id="67291309FF552E79EB0B826E122E2A91" name="United States of America">USA</collectingCountry>
(
<collectingRegion id="C917C3E13E0E97ACA85E4188CE961990" country="United States of America" name="Michigan">Michigan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPEAI" type="remarks">
<paragraph id="DD776FB42F59C5F2F44EC8A2924FD81B">
<heading id="AD7288433BE56515C45A1FDFF8527DE8" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="3E11F2681DFB851640FC7DA60A1B825E">
This is a very unusual
<taxonomicName id="911B62C33EE1048846B1E3CC8D912CDA">
<emphasis id="94A42046057D2D7F98577B5A91155C34" italics="true">Limnophyes</emphasis>
species
</taxonomicName>
; lack of lanceolate setae on both sexes, lack of humeral setae in males, and single humeral setae in females are distinguishing characteristics. However, the Neotropical species
<taxonomicName id="39748D08D5245F40FD9FB5FFB38A7C0E" authority="(Edwards, 1931)" baseAuthorityName="Edwards" baseAuthorityYear="1931" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="brachyarthra">
<emphasis id="3AB99D8838836587BB858C8FF153500D" italics="true">Limnophyes brachyarthra</emphasis>
(Edwards, 1931)
</taxonomicName>
described by
<bibRefCitation id="FE44AAEB00FEC554D2D17E6392B692F7" DOI="10.1163/187631290X00229" author="Sæther OA" firstAuthor="Saether" issue="3" journalOrPublisher="Insect Systematics &amp; Evolution" pagination="305-319" refId="B50" refString="Sæther OA (1990) A revision of the Neotropical types described as Spaniotoma (Limnophyes) by Edwards 1931, with the description of Edwardsidia gen. n. (Diptera: Chironomidae). Insect Systematics &amp; Evolution 21 (3): 305–319. https://doi.org/10.1163/187631290X00229" title="A revision of the Neotropical types described as Spaniotoma (Limnophyes) by Edwards 1931, with the description of Edwardsidia gen. n. (Diptera: Chironomidae)." volume="21" year="1990">Sæther (1990)</bibRefCitation>
also lacks lanceolate setae and humerals. The adults of
<taxonomicName id="2EB29967D0FDFABF36BACBD958B7A343" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="stagnum" status="sp. nov.">
<emphasis id="8DD41BDCE2666E39FCFEF4F5C99EDCF0" italics="true">L. stagnum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="97E39ECE5E35184A262F194217A0CB63" rank="species">sp. nov.</taxonomicNameLabel>
can be easily separated from those of
<taxonomicName id="ECE566B76F5D0F5946A6C729B86A384E" baseAuthorityName="Edwards" baseAuthorityYear="1931" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="brachyarthra">
<emphasis id="D478335BAFFA74378CD79CF5FFC2D07F" italics="true">L. brachyarthra</emphasis>
</taxonomicName>
based on the number of antennal flagellomere and AR. Additionally, the chaetotaxy of the thorax varies between the two species, and adult males have gonostylus of different shapes and sizes. According to
<bibRefCitation id="24EB1D0B2B8E6467E00CC4FF477325EB" author="Sæther OA" firstAuthor="Saether" refId="B48" refString="Sæther OA (1985) A review of the genus Limnophyes Eaton from the Holarctic and Afrotropical regions (Diptera: Chironomidae, Orthocladiinae). Entomologica Scandinavica (Supplement 35): 1–139." year="1985">Sæther (1985)</bibRefCitation>
the placement of lanceolate humeral setae in relation to the humeral pit varies among
<taxonomicName id="E4DCF098E7D4180C107C7BAC14E28074" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4AB497085BCD192B9BD9DB853BD34921" italics="true">Limnophyes</emphasis>
species.
</taxonomicName>
The lanceolate setae can be in or on the margin of humeral pit, concentrated around or above the pit, or scattered over the humerals. Although the lanceolate setae are missing in
<taxonomicName id="E57976A80DB92DA95C55D3CE5FA3CC4B" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="stagnum" status="sp. nov.">
<emphasis id="E6A3DA1360D3FDDB82C663ADA84E6B68" italics="true">L. stagnum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BDBF839024006B4904F9F1C8034B668" rank="species">sp. nov.</taxonomicNameLabel>
, the humeral seta of the female is located on the pit, similar to that in
<taxonomicName id="726BC82DE3C5ABA1579FF57BF177632E" authorityName="Kieffer" authorityYear="1914" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="18F9D93045ADB296192FB51C95F844E4" italics="true">L. natalensis</emphasis>
</taxonomicName>
. The shape of the hypopygium of the new species also resembles
<taxonomicName id="7A92E5BF70A4326353831FF478C57AA4" authority="Kieffer, 1914" authorityName="Kieffer" authorityYear="1914" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="6E35FE21F8C20104BEA3A3B963FE508E" italics="true">Limnophyes natalensis</emphasis>
Kieffer, 1914
</taxonomicName>
as well as
<taxonomicName id="400ED54C98DA93AA598EC24C766A9968" authority="Brundin, 1947" authorityName="Brundin" authorityYear="1947" class="Insecta" family="Chironomidae" genus="Limnopyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="difficilis">
<emphasis id="944A295E5AFC4112E9E77ADAC07F2320" italics="true">Limnopyes difficilis</emphasis>
Brundin, 1947
</taxonomicName>
. The adult male of the new species can be separated from the adult males of
<taxonomicName id="FD5730BE1F691B63E001B8A13ADAB45C" authorityName="Kieffer" authorityYear="1914" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="20310321D8E1706943DADC6B3FD5E1F3" italics="true">L. natalensis</emphasis>
</taxonomicName>
and
<taxonomicName id="5528B2A69D873F4120F03F388ACCA58A" authorityName="Brundin" authorityYear="1947" class="Insecta" family="Chironomidae" genus="Limnopyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="difficilis">
<emphasis id="7F89349D9F60E46C05F9A1DA236C0021" italics="true">L. difficilis</emphasis>
</taxonomicName>
based on the number of antennal flagellomeres, antennal ratio, lack of lanceolate and humeral setae, number and formation of thoracic setation, and bare squama. The adult female of the new species can be separated from the adult females of
<taxonomicName id="E371730C84D5358326C8143803324435" authorityName="Kieffer" authorityYear="1914" class="Insecta" family="Chironomidae" genus="Limnophyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="D8EDAFDE7DD5565BD2322FFBAED4E97C" italics="true">L. natalensis</emphasis>
</taxonomicName>
and
<taxonomicName id="26AD906EC865A0CF76E45490C33BB37E" authorityName="Brundin" authorityYear="1947" class="Insecta" family="Chironomidae" genus="Limnopyes" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="difficilis">
<emphasis id="2E2CECB3AAF9ABDB622FA200EA03D843" italics="true">L. difficilis</emphasis>
</taxonomicName>
based on the number of humeral setae, number and formation of thoracic setation, bare squama, and size of the notum.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="ECEDFFBAB334CC73638166D454FA152E" ID-DOI="10.3897/contrib.entomol.74.e126799" ID-ZooBank="339AB68C-DCB1-435F-B88E-2C42D0A7B05E" ID-publisher-id="126799" URI-arpha="B2861E35-EA80-53AF-A4AD-1BA2816CBD9C" URI-zoobank="http://zoobank.org/339AB68C-DCB1-435F-B88E-2C42D0A7B05E" article-type="research-article" checkinTime="1722264657988" checkinUser="pensoft" docAuthor="da Costa-Silva, Vinícius, Strümpher, Werner P., Barclay, Maxwell V. L. &amp; Vaz-de-Mello, Fernando Z." docDate="2024" docId="91E0E2086F9B525B9E6564F2976E63D1" docLanguage="en" docName="Contributions to Entomology 74(1): 81-101" docOrigin="Contributions to Entomology 74 (1)" docSource="https://contributions-to-entomology.arphahub.com/article/126799/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Omorgus (Omorgus) borrei" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="101" masterDocId="B2861E35EA8053AFA4AD1BA2816CBD9C" masterDocTitle="An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes" masterLastPageNumber="101" masterPageNumber="81" pageNumber="81" updateTime="1722265173734" updateUser="ExternalLinkService">
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<mods:title id="4FACF63BC0F0B0354DE6F2F94986D5EE">An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes</mods:title>
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<mods:name id="0D9551E57456BE77D777973D433F1F39" type="personal">
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<mods:namePart id="57C1802A1FD94A95C2CB100ECA626E72">da Costa-Silva, Vinícius</mods:namePart>
<mods:nameIdentifier id="F5B7DA831FB24CCAA7C758F9D52D1499" type="ORCID">0000-0002-4556-3793</mods:nameIdentifier>
<mods:affiliation id="E904F14EFD7CCF23D4F8BDC441B06027">Department of Zoology &amp; Entomology, University of Pretoria, Private Bag X 20, Hatfield, 0028, South Africa &amp; Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
</mods:name>
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<mods:namePart id="4B2BCD7AB11D29EAF2A4BEA65692D039">Strümpher, Werner P.</mods:namePart>
<mods:nameIdentifier id="1BF951C779D4A3A7F1B9F5DA9A68A7BB" type="ORCID">0000-0002-7047-2666</mods:nameIdentifier>
<mods:affiliation id="96D0A9CCD2704EE7890A87694818CEC7">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa</mods:affiliation>
</mods:name>
<mods:name id="645393753D99FA8D7FFF2004C7B7C7AF" type="personal">
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<mods:namePart id="51BE62F82B3A73CA57B6D03E90F68409">Barclay, Maxwell V. L.</mods:namePart>
<mods:nameIdentifier id="F1DD35345F872A9061C53D55AEDC9D3B" type="ORCID">0000-0003-4989-2014</mods:nameIdentifier>
<mods:affiliation id="B8EC832FFA81095A4EC7AA1FF940D7A9">Department of Life Sciences, Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name id="B95460C77CB67F29FDCA74E95685AE93" type="personal">
<mods:role id="C869353B1896909BE8E5D7E7FE6944AA">
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</mods:role>
<mods:namePart id="412B0723FB42C0F8223D5A55CE1DC02E">Vaz-de-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="114AF6D556EE4DC03C5C0C297088D512" type="ORCID">0000-0001-9697-320X</mods:nameIdentifier>
<mods:affiliation id="7483C485FCC3BD05F0509E7EA0EDB253">Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
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<mods:title id="47A1E5E4903AF3BA2AEA1B08188D38EE">Contributions to Entomology</mods:title>
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<mods:part id="2C1464BABB295B0E716C42FA9E9AE4EE">
<mods:date id="9AA5687A3B3FA558E1DC9F076FCC6547">2024</mods:date>
<mods:detail id="DAD36EA7D265C57DD59A56DDBC902841" type="pubDate">
<mods:number id="A6844D6E2A4D8B707D549DF685A2DD42">2024-07-29</mods:number>
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<mods:number id="067B1EC7B78505031522864CB9C2FA50">74</mods:number>
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<treatment id="91E0E2086F9B525B9E6564F2976E63D1" ID-DOI="http://doi.org/10.5281/zenodo.13123978" ID-Zenodo-Dep="13123978" ID-arpha="91E0E208-6F9B-525B-9E65-64F2976E63D1" LSID="urn:lsid:plazi:treatment:91E0E2086F9B525B9E6564F2976E63D1" httpUri="http://treatment.plazi.org/id/91E0E2086F9B525B9E6564F2976E63D1">
<subSubSection id="D3119FFA820158594C62B58B29297299" type="nomenclature">
<paragraph id="E6DF4DA5D4E4455B2ADDFBCBEA815A71">
<taxonomicName id="496870A726D20E5BC5EC342DD9A41057" ID-CoL="49DQQ" authority="(Harold, 1872)" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="borrei" subGenus="Omorgus">
<emphasis id="A3258A6A88FD78C1B17704D21756BC61" italics="true">Omorgus</emphasis>
(
<emphasis id="926E839A39B9A1AA045CA2E0A4D3A994" italics="true">Omorgus</emphasis>
)
<emphasis id="3EE2CBBCED8A539C666A04206F704E05" italics="true">borrei</emphasis>
(
<bibRefCitation id="1FE3D03F6FB5E1A1C35FF79E14789EB4" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="82067B844CDE5C7FA7307591F4993ED3" type="reference_group">
<paragraph id="DCBFF41FAF307F49BC3B27C054B1FBA1">
<treatmentCitationGroup id="FA42275AF603A0F579EB1B83CBFFEF51">
<treatmentCitation id="9731E6CFBE33C02BB9FFC9A136EEDE7F" author="Harold E" page="84" year="1872">
<taxonomicName id="01A57A4D3F4BA3E833F18A86E1699225" ID-CoL="59DHF" authority="Harold, 1872: 84" authorityName="Harold" authorityPageNumber="84" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="borrei">
<emphasis id="961CB0584F06CD963E967B31E3B30343" italics="true">Trox borrei</emphasis>
<comment id="FFAC18175ED4849942A154FB68ABD231">
<bibRefCitation id="8D92D3CF67363941A75A4BFF81A87C47" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872: 84</bibRefCitation>
(original description);
<treatmentCitation id="098BC486B8733C935D48CB045524A60F" author="Bruch C" page="193" year="1911">
<bibRefCitation id="87A4772C5202024916C9934FCD14F374" DOI="10.5962/bhl.title.8799" author="Bruch C" firstAuthor="Bruch" page="193" refId="B16" refString="Bruch C (1911) Catálogo sistemático de los Coleópteros de la República Argentina. Pars IV: Familias Lucanidae, Scarabaeidae (Coproini - Cetonini), Passalidae. Revista del Museo de La Plata, Tomo XVII, 181–225. https://doi.org/10.5962/bhl.title.8799" year="1911">Bruch 1911: 193</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="4C6E21157B95174331DD5F0CEB87F507" author="Arrow GJ" page="54" year="1912">
<bibRefCitation id="95C983CAF9C93F1B161F5588479A01AC" author="Arrow GJ" firstAuthor="Arrow" journalOrPublisher="Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Berlin: W. Junk" page="54" refId="B3" refString="Arrow GJ (1912) Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 43. Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Berlin: W. Junk, 66 pp." title="Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 43." year="1912">Arrow 1912: 54</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0471E35649C9A99256417A5D51A42F0E" author="Blackwelder RE" page="218" year="1944">
<bibRefCitation id="DF2F2F8F0A6297E05D2E6510A50126D8" DOI="10.5479/si.03629236.185.2" author="Blackwelder RE" firstAuthor="Blackwelder" journalOrPublisher="Bulletin of the United States National Museum" page="218" pagination="218-219" refId="B10" refString="Blackwelder RE (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Scarabaeidae, Troginae. Bulletin of the United States National Museum 185: 218–219. https://doi.org/10.5479/si.03629236.185.2" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Scarabaeidae, Troginae." volume="185" year="1944">Blackwelder 1944: 218</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="F6D22BA8C9CC28788CFD40DCD94EB5DF" author="Vaurie P" page="147" year="1962">
<bibRefCitation id="7468E75BFBBD660793C41128EA5C9FF6" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" page="147" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105–167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962: 147</bibRefCitation>
</treatmentCitation>
(redescription).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="371594A0EEF16F16C4B1E470FCC8F66B">
<treatmentCitationGroup id="7CC1A7D844656DA9BA213A835A0C33A8">
<treatmentCitation id="450896D293FCC56BC4FC41CD9E35EA92">
<taxonomicName id="80F934100D0892180C4955C66A5F093B" ID-CoL="59DHF" authorityName="Harold" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="borrei" subGenus="Omorgus">
<emphasis id="E9256053D40F4644F6E470E4C575456E" italics="true">Trox</emphasis>
(
<emphasis id="664E21D080DB1ADC2BEED3B12F9268B5" italics="true">Omorgus</emphasis>
)
<emphasis id="9F4DBBFE0D12B20140F9C662F3859902" italics="true">borrei</emphasis>
</taxonomicName>
<comment id="D81E0CFB4CA6391F62E4E42CED24B9CC">
:
<treatmentCitation id="E9C19379D5907DB1405DCE80A1947805" author="Scholtz CH" page="9" year="1982">
<bibRefCitation id="21AFFD44AF15295B2B8097B09EFDA072" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa" page="9" pagination="1-27" refId="B96" refString="Scholtz CH (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa 54: 1–27." title="Catalogue of world Trogidae (Coleoptera: Scarabaeoidea)." volume="54" year="1982">Scholtz 1982: 9</bibRefCitation>
</treatmentCitation>
(catalogue).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="2FD2BE4F3B4168156CDD4E5119E22553">
<treatmentCitationGroup id="D720F8376D9983BCE8F0C1AB4E9B53B1">
<treatmentCitation id="B81034CB8BF347F669E75F06E624EF3E">
<taxonomicName id="BFA79D40557454E51ED100D1FBC612D1" ID-CoL="49DQQ" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="borrei" subGenus="Omorgus">
<emphasis id="0575276C55815F6301AD8C02DCAC6E65" italics="true">Omorgus</emphasis>
(
<emphasis id="A2045149D374231D998A9A85CB7BEF8A" italics="true">Omorgus</emphasis>
)
<emphasis id="7E819643E753A365C2341175B233EEEA" italics="true">borrei</emphasis>
</taxonomicName>
<comment id="8D37C1B8D278BAD3CAFA6CFA03287D71">
:
<treatmentCitation id="E9CFC62EF67F330931F6A4B72CFC059F" author="Scholtz CH" page="361" year="">
<bibRefCitation id="2D63464D878C4C61CCD92D66C33E5020" DOI="10.1111/j.1365-3113.1986.tb00186.x" author="Scholtz CH" journalOrPublisher="Systematic Entomology" pagination="355-363" refId="B97" refString="Scholtz CH (1986 a) Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology 11: 355–363. https://doi.org/10.1111/j.1365-3113.1986.tb00186.x" title="Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea)." volume="11">Scholtz 1986 a</bibRefCitation>
: 361
</treatmentCitation>
(phylogenetics);
<treatmentCitation id="3E3E16CCAEB74F28BCD00529728AB5A0" author="Scholtz CH" page="1417" year="1990">
<bibRefCitation id="9ACA84F878B761FCAE1D8C3983FD2F2D" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" page="1417" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990: 1417</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="E54E17A75E4DFBA3DD32C97A7DC27BD6" author="Zidek J" page="8" year="2013">
<bibRefCitation id="AB00D9DADB558A4A4AE7822FF0EB99BA" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="8" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1–38." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 8</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="FD0172CB973F4C3DAF875F2BF3977840" author="Zidek J" page="98" year="2017">
<bibRefCitation id="033D2EBED77F2690756E8D3E6472923D" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="98" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 98</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="473AE8D1B11A65978DBA762E0857DB53" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" httpUri="http://treatment.plazi.org/id/03DA2410FFA5FFE9FE48FD5AFEA21ED0" page="2002" year="2021">
<bibRefCitation id="F483AB3C03BAD8E80EA0F028C0F877AA" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="2002" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021: 2002</bibRefCitation>
</treatmentCitation>
(review of Brazilian species,
<typeStatus id="A62C6B3B7D1D3F38BBFB770BFF02CA11" type="lectotype">lectotype</typeStatus>
designation).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="6737EDB2AC9683E01267581DDD217C86">
<treatmentCitationGroup id="6650F6ABCF6C4BF4F6617215A448EAD6">
<treatmentCitation id="431D5A094401AB24BA290007627D9C78">
<taxonomicName id="995AB3BF2A587260C1C10575AE3BB1FC" ID-CoL="49DQQ" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="borrei">
<emphasis id="2F6EE38FEA9945899C6B86917EE8515C" italics="true">Omorgus borrei</emphasis>
</taxonomicName>
<comment id="25EB15649BCE54CEE46C62F91AD42E8A">
:
<treatmentCitation id="B4A17E9E1F9174FB0F712360615DA867" author="Deloya C" page="122" year="2005">
<bibRefCitation id="BA86FA3F840CCE19BFEAF12AF5AEB2BB" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121–129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist, as “
<emphasis id="35280413F7BF9CD2DAA4AE74D1E6DB4B" italics="true">borreri</emphasis>
”);
<treatmentCitation id="1B0EC364FFFCDC666615EAA52977FE82" author="Morrone JJ" page="78" year="2014">
<bibRefCitation id="228BF1F527CED32BCD4FC51E5568DE76" DOI="10.11646/zootaxa.3782.1.1" author="Morrone JJ" firstAuthor="Morrone" issue="1" journalOrPublisher="Zootaxa" page="78" pagination="1-110" refId="B72" refString="Morrone JJ (2014) Biogeographical regionalisation of the Netropical region. Zootaxa 3782 (1): 1–110. https://doi.org/10.11646/zootaxa.3782.1.1" title="Biogeographical regionalisation of the Netropical region." volume="3782" year="2014">Morrone 2014: 78</bibRefCitation>
</treatmentCitation>
(mention, biogeography).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYYAE" type="Type specimen examined">
<paragraph id="8581F3ABD4339F0476BA5A62341FDB30">
<heading id="FD10C783772417E59D60F64781A237F0" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="8911105F494B7DD30CBF2122E64D7E3E">
<materialsCitation id="EB103F4D420DCB75CB55596A8BCF3DC3" collectionCode="RBINS" country="Uruguay" location="See" specimenCount="1" stateProvince="Montevideo" typeStatus="lectotype">
<emphasis id="91778E8A21D4640BBC3F2A0BD3CA0900" bold="true" italics="true">
<typeStatus id="6187E7CAFE79E43A57BC22723C7C9F4B" type="lectotype">Lectotype</typeStatus>
</emphasis>
(
<named-content id="3817AC9DCD5B7B9778CBBAE1FCE5DB3D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/royal-belgian-institute-natural-sciences" xlink_title="Royal Belgian Institute of Natural Sciences">
<emphasis id="607BB2C87C4EF791760E1F987EFADF08" bold="true">
<collectionCode id="C9188D2D8DBBA7FCA280760EE00E627A" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" type="Museum">RBINS</collectionCode>
</emphasis>
</named-content>
).
<location id="0A7948ADFDF6E013163C21831786B56E" LSID="urn:lsid:plazi:treatment:91E0E2086F9B525B9E6564F2976E63D1:0A7948ADFDF6E013163C21831786B56E" country="Uruguay" name="See" stateProvince="Montevideo">See</location>
<bibRefCitation id="01D0B5E756F941591243806DDD52558C" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="2002" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021: 2002)</bibRefCitation>
for high-quality images of the type specimen.
<emphasis id="A97346E9FEE2A3EBB62EF2565E8342B5" bold="true">Type locality</emphasis>
: “
<collectingRegion id="C5C5D410754C6704B32A46B14F50E569" country="Uruguay" name="Montevideo">Montevideo</collectingRegion>
,
<collectingCountry id="E45F77CBD8BFBF776D2B8C3E59ED313C" name="Uruguay">Uruguay</collectingCountry>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOZAE" type="distribution">
<paragraph id="011A14A5B64DEA738E233979304069CE">
<heading id="BD9D0CE6DEFA8A0B95C18172BBC00FFE" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="D6849CBAE8D54A1C7F96090E49FA334F">
<collectingCountry id="EC07239F2BF4ED7E1DFDBE98918AE182" name="Argentina">Argentina</collectingCountry>
,
<collectingCountry id="EECEDA618ACE97B750B60FF5D357ACB0" name="Bolivia">Bolivia</collectingCountry>
,
<collectingCountry id="6AF6F022AB85F84CA4098C035CE19576" name="Brazil">Brazil</collectingCountry>
,
<collectingCountry id="2A5D247C3B823AC0B6E60F7FF230D96F" name="Paraguay">Paraguay</collectingCountry>
and
<collectingCountry id="27FF071877F1D2F755726ADEC918F0BA" name="Uruguay">Uruguay</collectingCountry>
(
<bibRefCitation id="F0B67C1CB5194DCF4820918AE4AB8F96" author="Vaurie" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105–167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962</bibRefCitation>
;
<bibRefCitation id="745DB85848420B14EC1EBB36A91833FD" DOI="10.1080/00222939000770841" author="Scholtz" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990</bibRefCitation>
;
<bibRefCitation id="892E584304E7C8177C43479357E21D65" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EA1AE" type="type material">
<paragraph id="BEFDB051038D3B38B64F1A578A79AA01">
<heading id="401D77679D102E8E91E29D0A18B53D0D" reason="title">Non-type examined material</heading>
</paragraph>
<paragraph id="656FDE4F3B98A885D1117FBB9B8960BA">
<emphasis id="B69DB488E87FC9519D757B2CA03175D3" bold="true">
(
<specimenCount id="0A07EA2917670D5C68CF374A41281FA5" count="23" type="generic">23 specimens</specimenCount>
additionally to
<bibRefCitation id="3CC92ED5A14A5D983B7F47E19627F997" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021</bibRefCitation>
).
</emphasis>
<materialsCitation id="E1DB20573D805C13AFE07B76A7BFF1AF" country="Argentina" location="Chaco" specimenCount="1" stateProvince="Chaco">
<collectingCountry id="3E52974BADBAA9B45DC32EDC9584ED7A" name="Argentina">ARGENTINA</collectingCountry>
–
<emphasis id="E173F5EF21B6A1C96458173E432755BE" bold="true">
<collectingRegion id="D1B8321BA3D49BDFE36A9D1B6AED8771" country="Argentina" name="Chaco">Chaco</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="33DE051CA4205CAF3F4AEDE85B55EEED" collectingDate="1995-12" collectionCode="NHMUK" collectorName="S. Bolle" country="Argentina" location="Charata" specimenCount="1" specimenCount-male="1" stateProvince="Chaco">
<specimenCount id="C75FE41EEA6EF6DCFC603D57E727521C" count="1" type="male">1 ♂</specimenCount>
and 1 unsexed;
<location id="0FC04FE20E18A22C9A1A7B81E747460A" LSID="urn:lsid:plazi:treatment:91E0E2086F9B525B9E6564F2976E63D1:0FC04FE20E18A22C9A1A7B81E747460A" country="Argentina" name="Charata" stateProvince="Chaco">Charata</location>
;
<date id="F1C908E27DE1A013C042EE16D7B0080B" value="1995-12">
<collectingDate id="185A66AE153565820E0F31755F0FE559" value="1995-12">Dec. 1995</collectingDate>
</date>
;
<collectorName id="07A1747B2B0297B28A53447A0C3824C6">S. Bolle</collectorName>
leg.;
<named-content id="DCFAA453A36EE5EAC301D1242BD5D127" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="34126B4F9A4813DDA86A5874465CB9A9" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
</materialsCitation>
•
<materialsCitation id="F34663D29CCA23A896BF03DDA11A1EF3" collectingDate="1982-09" collectionCode="A, CMNC" collectorName="Martinez &amp; Currientes" country="Argentina" location="Pampa del Infierno" specimenCount="1" stateProvince="Chaco">
1;
<location id="A42E560139D56CC3E8C251060BD0FF86" LSID="urn:lsid:plazi:treatment:91E0E2086F9B525B9E6564F2976E63D1:A42E560139D56CC3E8C251060BD0FF86" country="Argentina" name="Pampa del Infierno" stateProvince="Chaco">Pampa del Infierno</location>
;
<date id="BA0EC8109BC0F64FAA99E1A8B8401A69" value="1982-09">
<collectingDate id="E1DED8CA56DA98811B022048E82CCCBD" value="1982-09">Set. 1982</collectingDate>
</date>
,
<collectionCode id="30662C03BD633F0484301FFF8A54DE2B">A</collectionCode>
.
<collectorName id="81B2E166E9E777173B08AEDFB58FD4FC">Martinez</collectorName>
leg.;
<named-content id="35ED5FC80DA5D1F18421998AD4169B13" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">
<collectionCode id="F48AFEADCCE8C44B314E9D0E1C5855A1">CMNC</collectionCode>
</named-content>
. –
<emphasis id="71222C6C1F31F180CF9B3BB13575B07C" bold="true">
<collectorName id="43D7E976C40AD5EE87060FC58C42C2A6">Currientes</collectorName>
</emphasis>
</materialsCitation>
•
<materialsCitation id="B081AEF66D7FE337478AFB0E43FCB0F8" collectingDate="1903-12" collectionCode="MNHN" collectorName="Bords du Parana &amp; E. R. Wagner" country="Argentina" location="Bella Vista" specimenCount="1" specimenCount-female="1" stateProvince="Chaco">
<specimenCount id="25868378B6A0680B4F349ED76D650045" count="1" type="female">1 ♀</specimenCount>
;
<location id="C6C2E107CEDCD6369E1C72A3860C017B" LSID="urn:lsid:plazi:treatment:91E0E2086F9B525B9E6564F2976E63D1:C6C2E107CEDCD6369E1C72A3860C017B" country="Argentina" name="Bella Vista" stateProvince="Chaco">Bella Vista</location>
,
<collectorName id="00E29CA03671DF1D79D86530FDC477BB">Bords du Parana</collectorName>
;
<date id="0547962DBA54802D11927FE9EB7FD889" value="1903-12">
<collectingDate id="BAAAE84A51256209D855B24C97A335AB" value="1903-12">Dec. 1903</collectingDate>
</date>
,
<collectorName id="C01EAA00566676F67A2DCE8EF7C4D2E5">E. R. Wagner</collectorName>
leg.;
<named-content id="A387A5CCF8852E5076FA63BFD37BDBA7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire naturelle, Paris">
<collectionCode id="7D919CB1211AFA7ECCCA80F319F867EF" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
</materialsCitation>
•
<materialsCitation id="997122D7C729ECED7C124DE479A9E003" collectingDate="1827-03-15" collectingDateMax="1828-04-20" collectingDateMin="1827-03-15" collectionCode="MNHN" collectorName="d'Orbigny &amp; Misiones" country="Argentina" specimenCount="1" stateProvince="Chaco">
1;
<date id="E1AFA1C638657B50641B567B8890DF1C" value="1827-03-15">
<collectingDate id="D42E10D88C4A07BFBB8268503067282A" value="1827-03-15">15 Mar. 1827</collectingDate>
</date>
–
<date id="4F687040571D78EE8B72070346DC8A43" value="1828-04-20">
<collectingDate id="5FD6F855DD78D0688B1B2D17BB1CD025" value="1828-04-20">20 Apr. 1828</collectingDate>
</date>
,
<collectorName id="CB12C61E35B7DC9ADE8EA1E269100264">d’Orbigny</collectorName>
leg.;
<named-content id="66B2E6EE6846856CD18A0D80B3D35086" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire naturelle, Paris">
<collectionCode id="99FF46281463298F465E1A1834A64883" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
. –
<emphasis id="DA9D7A8FFAAC4FD995E8602A9C93C0CF" bold="true">
<collectorName id="A420D167FB016EE0BD30377FFC22C439">Misiones</collectorName>
</emphasis>
</materialsCitation>
• 1; 1932;
<collectorName id="76D8245FE9AD26D3AD9E7FBB8DB87F67">K. J. Hayward</collectorName>
leg.
<named-content id="3ECD1330B88D86F933A6AA93366FB295" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="A414B3AC23A9F2935032EE15378B7B21" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
. –
<materialsCitation id="338BAE382792D2DCC8AB4159592B5F54" collectingDate="1925-11" collectionCode="NHMUK" collectorName="K. J. Hayward" country="Argentina" location="Villa Ana" specimenCount="1" stateProvince="Santa Fe">
<emphasis id="C6C9FB6FD833960E5945F2C8A046E8FE" bold="true">
<collectingRegion id="03402BCCDE81BD1FD647BA91BB9A5094" country="Argentina" name="Santa Fe">Santa Fe</collectingRegion>
</emphasis>
• 1;
<location id="4F35D3FF9C9418BD9CA4378055C93043" LSID="urn:lsid:plazi:treatment:91E0E2086F9B525B9E6564F2976E63D1:4F35D3FF9C9418BD9CA4378055C93043" country="Argentina" name="Villa Ana" stateProvince="Santa Fe">Villa Ana</location>
;
<date id="E22544D30D23A8CF925026F47533AEDA" value="1925-11">
<collectingDate id="21B35062F0723A7FB24CA14DA6B6BF9D" value="1925-11">Nov. 1925</collectingDate>
</date>
,
<collectorName id="EDD4291A4D0B0878603EB7A37F06C6ED">K. J. Hayward</collectorName>
leg.;
<named-content id="8DE70AC59222D4C4F5F01F792D89B0E1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="264EFAA2FB964320C4A9D88E87BD863B" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
</materialsCitation>
. –
<materialsCitation id="A926196CB738E5CB3FD027A4F186D12A" collectingDate="1956-12" collectionCode="CNC" collectorName="El Pinto" country="Argentina" location="Santiago del Estero" specimenCount="1" specimenCount-male="1" stateProvince="Santiago del Estero">
<emphasis id="BCDCBCB6EF2F7A4F834FABE24C7E788F" bold="true">
<collectingRegion id="B5CFDF244427FC94B567B32E5851AC32" country="Argentina" name="Santiago del Estero">Santiago del Estero</collectingRegion>
</emphasis>
•
<specimenCount id="67C046B348A7EC4CB242023CB3C53A36" count="1" type="male">1 ♂</specimenCount>
and 7 unsexed;
<collectorName id="A639E323D8B0DC1D8C08242FBBA2C8C1">El Pinto</collectorName>
;
<date id="C395003D74EC0DABA6C191B8D7CBB231" value="1956-12">
<collectingDate id="FE6BF8871D762C54EBE0E7EF13BF014B" value="1956-12">Dec. 1956</collectingDate>
</date>
;
<named-content id="8C7DCFB444C0254A0657C51784659C34" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection of Insects">
<collectionCode id="63E9BC9493D86CE805B072444F8309EC" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
</named-content>
</materialsCitation>
•
<materialsCitation id="5DAA9336E7EF3145714C9C3F436C0C0F" collectingDate="1910-12" collectionCode="MNHN" collectorName="E. R. Wagner" country="Argentina" location="Edges of Rio Salado" municipality="Icano" specimenCount="1" specimenCount-female="1" stateProvince="Santiago del Estero">
<specimenCount id="A046775C8B5AC36C76F18D5494EE7133" count="1" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="08B6B16801436A0A25B0334C182B3938">Icaño</collectingMunicipality>
,
<location id="A4D35890E7F44AB55FF10B14DB389745" LSID="urn:lsid:plazi:treatment:91E0E2086F9B525B9E6564F2976E63D1:A4D35890E7F44AB55FF10B14DB389745" country="Argentina" municipality="Icano" name="Edges of Rio Salado" stateProvince="Santiago del Estero">Edges of Rio Salado</location>
;
<date id="226E10DFE860AF69B4FCDE0382B3C8BA" value="1910-12">
<collectingDate id="F7E7784D7DBAF090FF7B8D8998C4878F" value="1910-12">Dec. 1910</collectingDate>
</date>
,
<collectorName id="CE71F1478D7B89EAEA9A41F5C4E933D7">E. R. Wagner</collectorName>
leg.;
<named-content id="5629CF2ACEAD3CA947F1A5B26A770938" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire naturelle, Paris">
<collectionCode id="80BE475869436BCD62CDB3D1754C8825" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<paragraph id="E1D5D44CC1BF70560450A37810D7E287">
<collectingCountry id="03B9343F9F8302CDFA7D2ED591C98C61" name="Bolivia">BOLIVIA</collectingCountry>
• 1; Jansen leg.;
<named-content id="9B069648E889E8FEFA918F6223A6D1B8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="551FAB59E591BBE59F32A4822F193412" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
.
</paragraph>
<paragraph id="129B8271A212DCD2BA3CA0486126B436">
<collectingCountry id="5C334911E17DCE71208EC4537995777B" name="Brazil">BRAZIL</collectingCountry>
• 1; Matto Grosso (sic!); 1886; P. Germain leg.;
<named-content id="4D5D18515C6481AD0DF983CCA801B6E9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire naturelle, Paris">
<collectionCode id="B2630CF9C3E7FEAE1C4FE415A4FCFE0B" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
.
</paragraph>
<paragraph id="3BEDCF93F3763B162ADFEA1CAEC67820">
<materialsCitation id="277A8C3E0250AFD691C1FC487100EC56" collectingDate="1927-03" collectionCode="NHMUK" collectorName="G. S. Carter" country="Paraguay" location="Nanawa" specimenCount="1" stateProvince="Presidente Hayes">
<collectingCountry id="D412D38ABE8706BFFE92560C39DB8B0F" name="Paraguay">PARAGUAY</collectingCountry>
–
<emphasis id="F3D75B2422049A364A78B94AC556A850" bold="true">
<collectingRegion id="2B707DCED52F1D362F306A556F0B0EE4" country="Paraguay" name="Presidente Hayes">Presidente Hayes</collectingRegion>
</emphasis>
• 2;
<location id="458AD6FC52247F892D6ADD6B89761ECD" LSID="urn:lsid:plazi:treatment:91E0E2086F9B525B9E6564F2976E63D1:458AD6FC52247F892D6ADD6B89761ECD" country="Paraguay" name="Nanawa" stateProvince="Presidente Hayes">Nanawa</location>
, “ Paraguayan Chaco ”;
<date id="9BD5055A2EBB1D7FB0777497D3512035" value="1927-03">
<collectingDate id="7DC528F339A04004B616DB728922D76B" value="1927-03">Mar. 1927</collectingDate>
</date>
;
<collectorName id="887B88878ADCB2A34F8219201AF0159A">G. S. Carter</collectorName>
leg.;
<named-content id="9647AA4E7C061FD7B78A087F0CF11D77" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="A85F9D4970CC824CD2539ACBA19685D3" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<paragraph id="14502DC7D504E683BC967A9627B52480">
<materialsCitation id="7625C1DA33ACD87D4CA2D042C60BDCE2" collectingDate="2002-12-27" collectionCode="CMNC" collectorName="S. &amp; J. Peck" country="Uruguay" latitude="-30.516666" location="Artigas" longLatPrecision="1221" longitude="-56.366665" specimenCount="1" stateProvince="Artigas">
<collectingCountry id="F2A8E38FC749EDFE7879D0B45296865C" name="Uruguay">URUGUAY</collectingCountry>
–
<emphasis id="6B303F3FE74220FB2B153EAE60DEC690" bold="true">
<collectingRegion id="45310E7C64A850F7833E6C29A2153741" country="Uruguay" name="Artigas">Artigas</collectingRegion>
</emphasis>
• 1;
<quantity id="DF2C5D5604659F4791F2754C88708035" metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
SE
<location id="90113E5BB7882F3B5A6D951F1FEC7A33" LSID="urn:lsid:plazi:treatment:91E0E2086F9B525B9E6564F2976E63D1:90113E5BB7882F3B5A6D951F1FEC7A33" country="Uruguay" latitude="-30.516666" longLatPrecision="1221" longitude="-56.366665" name="Artigas" stateProvince="Artigas">Artigas</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ES3AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-56.366667,-30.516667]}">
<geoCoordinate id="A89DB75C5396E52B3EDB768C2DBCDFD5" degrees="30" direction="south" minutes="31" orientation="latitude" precision="925" value="-30.516666">30 ° 31 ' S</geoCoordinate>
,
<geoCoordinate id="F80501D0E620A5878E9594DAEED83E90" degrees="56" direction="west" minutes="22" orientation="longitude" precision="925" value="-56.366665">56 ° 22 ' W</geoCoordinate>
</named-content>
;
<date id="A3FD4585F841E53404BA21751E047EDC" value="2002-12-27">
<collectingDate id="D28CC62CE6D1F01AD7BC2E2EF783738C" value="2002-12-27">27 Dec. 2002</collectingDate>
</date>
,
<collectorName id="3C55F41E5528CEF03B13B2712A2037A0">S. &amp; J. Peck</collectorName>
leg.;
<named-content id="CA303240857054748FDE782CA4C1016F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">
<collectionCode id="30081AB54991C4CAF40032BECA8AF08B">CMNC</collectionCode>
</named-content>
</materialsCitation>
. –
<materialsCitation id="69642A49431FB43635C96F5C2C2B4845" collectionCode="MNHN" country="Uruguay" location="Montevideo" specimenCount="1" stateProvince="Montevideo">
<emphasis id="7885A7CDC74195F3DE361F48811C35BF" bold="true">
<collectingRegion id="922E88A6BAC8FFBCEA450AA65BC47F27" country="Uruguay" name="Montevideo">Montevideo</collectingRegion>
</emphasis>
• 1; Montevideo;
<named-content id="EB28F0B885711186EEEB170E1E7DB17E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire naturelle, Paris">
<collectionCode id="8E3C63F0033D453EF3DBCE8600C0AD38" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
. –
<emphasis id="064DB15B707E833CC0001CBF2CD4EF64" bold="true">Without specific locality</emphasis>
</materialsCitation>
•
<materialsCitation id="8F571E36FBADD2ADB15E8C46C74CB029" collectionCode="MNHN" country="Uruguay" specimenCount="1" specimenCount-female="1" stateProvince="Montevideo">
<specimenCount id="5857660120170DD72686017E2DA022DC" count="1" type="female">1 ♀</specimenCount>
;
<named-content id="0BC74CA499852A4FD87815C1B38D72A9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire naturelle, Paris">
<collectionCode id="51B5A4957EEFDC2235F2310E8301EF6D" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

237
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<mods:title id="7F97DC4C1307FAD2B1DA2814E54A82FE">An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes</mods:title>
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<mods:affiliation id="59B3FB02B96B44CD6C0F1CF84EF4246D">Department of Zoology &amp; Entomology, University of Pretoria, Private Bag X 20, Hatfield, 0028, South Africa &amp; Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
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<mods:namePart id="740184B96F02073C720538AC8BDA5BA8">Strümpher, Werner P.</mods:namePart>
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<subSubSection id="77B22E18EE96C55FB5EDA63D6CC48F66" type="nomenclature">
<paragraph id="8BF031D2825CFF2F84FE710896CC7E9E">
<taxonomicName id="43EC5F9793CC85783EFB3D836D6B230E" ID-CoL="49DRX" authority="Scholtz, 1990" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nocheles" subGenus="Omorgus">
<emphasis id="FD957989D1E4AB82124BEAEC9C54E64D" italics="true">Omorgus</emphasis>
(
<emphasis id="0BBD64D0887A6528B87632323D07528B" italics="true">Omorgus</emphasis>
)
<emphasis id="A9CCBF55CA3E3DA3E48B8B4C0BE2939B" italics="true">nocheles</emphasis>
<bibRefCitation id="EED348EAEBBF8D76127404CBB33F5EAE" DOI="10.1080/00222939000770841" author="Scholtz CH" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391 - 1456. https: // doi. org / 10.1080 / 00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz, 1990</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="5BFB04C949F7CA673A053FEB9C91B330" type="description">
<paragraph id="0967A1E886F46280158B42A42E163B93">
<figureCitation id="51EC077817B38F60A83EAF7403289EEA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Omorgus nocheles Scholtz, 1990. A. Habitus in dorsal view; B. Habitus in lateral view; C. Aedeagus in dorsal view; D. Aedeagus in lateral view; E. Labels. Photos by Brian Brown (LACM). Length of specimen: 15.5–16.5 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure5" httpUri="https://binary.pensoft.net/fig/1103867">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D2010D0BA6C1616CD7739E50E44D1300" type="reference_group">
<paragraph id="23A0E01A406E4F3D269A630F72084A1C">
<treatmentCitationGroup id="993633A9D95BCD7017CBE07C35D2EA83">
<treatmentCitation id="E7AFED5DC2AE30426C362B458E7D729F" author="Scholtz CH" page="1412" year="1990">
<taxonomicName id="E6F03CB9D69B63833C6FEFACEDA10633" ID-CoL="49DRX" authority="Scholtz, 1990: 1412" authorityName="Scholtz, 1990:" authorityPageNumber="1412" authorityYear="1412" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nocheles" subGenus="Omorgus">
<emphasis id="29989CB8B0F530B4BED6FDA63F6B21B4" italics="true">Omorgus</emphasis>
(
<emphasis id="34465DDFB9BDC6CDC18819440EFB82F3" italics="true">Omorgus</emphasis>
)
<emphasis id="D5DBD0B163FAA6CA3934DA0793F068C7" italics="true">nocheles</emphasis>
<comment id="D16D0152B7DF771789C9CDC4C958A453">
<bibRefCitation id="CCD984D470E8996E95D7EA0AEE951791" DOI="10.1080/00222939000770841" author="Scholtz CH" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391 - 1456. https: // doi. org / 10.1080 / 00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz, 1990: 1412</bibRefCitation>
(original description);
<treatmentCitation id="0FD81CE5383A27ABD5C96C7412860EC8" author="Gómez RS" page="515" year="2008">
<bibRefCitation id="8507CEF44C62A05F4D102F714BDD611A" author="Gómez RS" editor="Claps LE &amp; Debandi G &amp; Roig-Juñent S" firstAuthor="Gomez" journalOrPublisher="Sociedad Entomológica Argentina" page="515" pagination="509-518" refId="B46" refString="Gómez RS (2008) Trogidae. In: Claps LE, Debandi G, Roig-Juñent S (Eds) Biodiversidad de Artrópodos Argentinos II. Sociedad Entomológica Argentina, 509–518." title="Trogidae." volumeTitle="Biodiversidad de Artrópodos Argentinos II." year="2008">Gómez 2008: 515</bibRefCitation>
</treatmentCitation>
(key for Argentinean species);
<treatmentCitation id="87B463AE0DA0ABCCC77F0DDF9D4805AA" author="Zidek J" page="14" year="2013">
<bibRefCitation id="F049A0BDEAF4DBCC62539B9363572861" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="14" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1–38." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 14</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="C4C70961BB8C626A97AE697C6CD91EF8" author="Zidek J" page="105" year="2017">
<bibRefCitation id="3DF17DC05C8720162CD49B75C78C7C14" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="105" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 105</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="0B58DC3CD2B5C6B79D0A7C1F942AC05E" author="Smith ABT" page="86" year="2017">
<bibRefCitation id="FAAADEB69E12744704C923BA84AD028C" author="Smith ABT" firstAuthor="Smith" journalOrPublisher="Revista Chilena de Entomología" page="86" pagination="81-103" refId="B102" refString="Smith ABT (2017) Southern South American Trogidae (Coleoptera): verification of specimens and notes on geographic distribution. Revista Chilena de Entomología 43: 81–103." title="Southern South American Trogidae (Coleoptera): verification of specimens and notes on geographic distribution." volume="43" year="2017">Smith 2017: 86</bibRefCitation>
</treatmentCitation>
(notes).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="C5029AC22C64E744283FA746919E2554">
<treatmentCitationGroup id="4990D9A9DC19D0A12A985970F2AD9A12">
<treatmentCitation id="2E1396F1009F96BAF4D47D9F66150BF5">
<taxonomicName id="297CD42063B579CA7E7E416F48B1C8DF" ID-CoL="49DRX" authorityName="Scholtz, 1990:" authorityYear="1412" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nocheles">
<emphasis id="B02A5F40C251584226707F1949FC9AA2" italics="true">Omorgus nocheles</emphasis>
</taxonomicName>
<comment id="422B1E5666E12FB67EA21C28DFF1D1B7">
:
<treatmentCitation id="26398C8E4500FB53489290072C720403" author="Deloya C" page="122" year="2005">
<bibRefCitation id="019FB8B70E7E0645990AECF49F78E9E8" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121–129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMYBG" type="Type specimen examined">
<paragraph id="582A74AF922C5BBFDDC63A84676E9327">
<heading id="2F319CD9CB9FB7D8760340BEFB3EF481" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="0A5731EBBB8CFA8207A8B52CE655063D">
<materialsCitation id="7291451BD4827CB7DAE4CFA73ACFFF80" collectingDate="1956-10-24" collectingDateMax="1956-10-25" collectingDateMin="1956-10-24" collectionCode="LACM" collectorName="Second &amp; Clarke Scholtz's &amp; Third &amp; Fig." country="Argentina" location="First" specimenCode="ENT 160322" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="0B50EF78F5E8B1E03F46096976EC4B04" bold="true" italics="true">
<typeStatus id="660A141DC886A0C85EC26312BC6644B4" type="holotype">Holotype</typeStatus>
, by original designation
</emphasis>
(
<specimenCount id="6F05732BA8B7DE7D017D3AD6CABDA441" count="1" type="male">♂</specimenCount>
<named-content id="A7EE1B91140C17EFEBFC5E68A0CC7B74" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-los-angeles-county" xlink_title="Los Angeles County Museum of Natural History">
<emphasis id="8D43093AAC27A7A97419DD2F37FEE396" bold="true">
<collectionCode id="B1D213A8D05EF8A8583AB8BA86D061E9" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" type="Museum">LACM</collectionCode>
</emphasis>
</named-content>
).
<location id="C2A6B3AFB6CDB0303D8532C99DA8782B" LSID="urn:lsid:plazi:treatment:A182B442C86E55DAB2E2411DCF6DE621:C2A6B3AFB6CDB0303D8532C99DA8782B" country="Argentina" name="First">First</location>
label [white, typeset]: “ El Bolson, Rio Negro, /
<collectingCountry id="0A69F7ED15E70A0F71E9B86EEFA25CF1" name="Argentina">ARGENTINA</collectingCountry>
/
<date id="5ABB321AEE08BE1668763432AC0827B9" value="1956-10-24" valueMax="1956-10-25" valueMin="1956-10-24">
<collectingDate id="6F5236B17B110BF484B7DF76D8BFEAB7" value="1956-10-24" valueMax="1956-10-25" valueMin="1956-10-24">Oct. 24–25, 1956</collectingDate>
</date>
/ Andor Kovacs ”.
<collectorName id="1DF7F26EC4465A8EFC8C9C67B69A72ED">Second</collectorName>
label [white with red border,
<collectorName id="7D2F02D73F78F71CBA1070B4B41E0803">Clarke Scholtz’s</collectorName>
handwriting]: “
<taxonomicName id="CEB0BCE44FE5BB094405CB6AD243F028" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="2E0CAEC95657233AAED691E54B8A4D2D" italics="true">Omorgus</emphasis>
</taxonomicName>
/
<taxonomicName id="4E8A56E8388BD30EAD000661297D3D36" authorityName="Scholtz, 1990:" authorityYear="1412" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nocheles">
<emphasis id="1DAC39DE40BEC947955EDA1EE51E4557" italics="true">nocheles</emphasis>
</taxonomicName>
/ 1988 / Scholtz /
<typeStatus id="876BBB90B734ABBB03ABB6843C4624E0" type="holotype">HOLOTYPE</typeStatus>
”.
<collectorName id="D91AAF562281C8F03486105022E1F6CA">Third</collectorName>
label [white, printed]: “ [QR Code] /
<named-content id="FEEE47DB5F8AC4E956F4B67397E4C6E8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-los-angeles-county" xlink_title="Los Angeles County Museum of Natural History">
<collectionCode id="371EE81E9F28FDBF33ABD98AAFDA0655" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" type="Museum">LACM</collectionCode>
</named-content>
<specimenCode id="ED39F1A6817E9BD789909C3E25CA69B4" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" type="Museum">ENT 160322</specimenCode>
” (
<collectorName id="6FF2D71C4E8B973D349C2BC441DCEAD1">Fig.</collectorName>
<figureCitation id="E3CFCD420D7512307ED4E5531A788F0D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Omorgus nocheles Scholtz, 1990. A. Habitus in dorsal view; B. Habitus in lateral view; C. Aedeagus in dorsal view; D. Aedeagus in lateral view; E. Labels. Photos by Brian Brown (LACM). Length of specimen: 15.5–16.5 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure5" httpUri="https://binary.pensoft.net/fig/1103867">5 E</figureCitation>
).
<emphasis id="1B6A793ADDCE4BC43968060F57ED6724" bold="true">Type locality</emphasis>
: “ Argentina, El Bolsón, Río Negro ”.
</materialsCitation>
</paragraph>
<caption id="C7A5662C9FF94DB618D54A6100D4E422" ID-DOI="10.3897/contrib.entomol.74.e126799.figure5" ID-arpha="6FFC447E-F115-5C50-B1A4-1C2D0634977F" httpUri="https://binary.pensoft.net/fig/1103867" startId="F5">
<paragraph id="02802699A18DD7E8418691E8E777D97C">
<label id="68A8906B7062B732460637BFD5EF1404">Figure 5.</label>
</paragraph>
<paragraph id="A4B2B874696FC7FD3EA72A5E6107AB66">
Holotype of
<taxonomicName id="7AACD648FE4F2B7414FB509DAD64B490" authority="Scholtz, 1990" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nocheles">
<emphasis id="670463C87C85395C8129E09E9E9E2BDB" italics="true">Omorgus nocheles</emphasis>
<bibRefCitation id="5D4B1A707DA1D9B9A9C43C453D206EC1" DOI="10.1080/00222939000770841" author="Scholtz CH" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391 - 1456. https: // doi. org / 10.1080 / 00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz, 1990</bibRefCitation>
</taxonomicName>
.
<emphasis id="2658C3088CF1DAABB302CB7830B7573C" bold="true">A.</emphasis>
Habitus in dorsal view;
<emphasis id="437D0632DD11C596511EF46DECC77694" bold="true">B.</emphasis>
Habitus in lateral view;
<emphasis id="F30E60D3AA9DAC12BBEC0F4E13FE4D21" bold="true">C.</emphasis>
Aedeagus in dorsal view;
<emphasis id="B688CC9EA9656534556C8C3C8B66B73C" bold="true">D.</emphasis>
Aedeagus in lateral view;
<emphasis id="B0F3F5B9EE2CFDAC863DFF34290F4CBF" bold="true">E.</emphasis>
Labels. Photos by Brian Brown (
<named-content id="4D7240623CF653192E45E1409DEF0ED6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-los-angeles-county" xlink_title="Los Angeles County Museum of Natural History">LACM</named-content>
). Length of specimen: 15.5–16.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E21BG" type="distribution">
<paragraph id="54DD9F4399C5284F118B22C9B8CD346C">
<heading id="D2173972C98B9C077A6DA9A3466455C5" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="A0858A3BFD8A3BAEE6F3BB492D1A96C9">
<taxonomicName id="1E3E739CDCD85A5E296D04B9BD9C3009" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nocheles" subGenus="Omorgus">
<emphasis id="242B414BEE3D047557ECCB38B40FBEA5" italics="true">Omorgus</emphasis>
(
<emphasis id="85CEEA72E6B45D43A839AC0FEADDCBF6" italics="true">O.</emphasis>
)
<emphasis id="F0452638E8F98A620C0114A80AA1A2D4" italics="true">nocheles</emphasis>
</taxonomicName>
is known only from
<specimenCount id="7CF1B7A2B1CCC67F83F0BF76B9B61618" count="2" type="generic">two specimens</specimenCount>
collected in
<quantity id="19235650E596316327752D562715D637" metricMagnitude="1" metricUnit="m" metricValue="4.96824" unit="in" value="1956.0">1956 in</quantity>
El Bolsón,
<collectingCountry id="806BA29FE899FE1C7072CB9897849686" name="Argentina">Argentina</collectingCountry>
(
<bibRefCitation id="C2F71BD760AE9CE2317763ED725A9704" DOI="10.1080/00222939000770841" author="Scholtz" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

425
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@ -0,0 +1,425 @@
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<mods:title id="02665A194A241E7AE298A4413CDBE0AD">An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes</mods:title>
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<mods:affiliation id="B6A6FF1F37E9CD58792414124E984C94">Department of Zoology &amp; Entomology, University of Pretoria, Private Bag X 20, Hatfield, 0028, South Africa &amp; Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
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<treatment id="AAE41682797B55719EB05CFA4076CFA2" ID-DOI="http://doi.org/10.5281/zenodo.13123972" ID-Zenodo-Dep="13123972" ID-arpha="AAE41682-797B-5571-9EB0-5CFA4076CFA2" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2" httpUri="http://treatment.plazi.org/id/AAE41682797B55719EB05CFA4076CFA2">
<subSubSection id="E05724007124FDC08A328DBE4A79B0AD" type="nomenclature">
<paragraph id="28F2B34AC059E70A768251F6F9C0C984">
<taxonomicName id="3F49B37964F30FFDDF2C97376FC19A74" ID-CoL="49DQD" authority="(Harold, 1872)" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi" subGenus="Haroldomorgus">
<emphasis id="18A0905C01B2B97105D042950FB926E4" italics="true">Omorgus</emphasis>
(
<emphasis id="D289EF1589062A0BAFDED198C81CE9D0" italics="true">Haroldomorgus</emphasis>
)
<emphasis id="ADE265465134BF2FD509D4D3514763B2" italics="true">batesi</emphasis>
(
<bibRefCitation id="2179EACC62224DF209BDA66A6DD52BD0" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F098D270F818191E31EEB89FCA860700" type="reference_group">
<paragraph id="2734DCE8896DBF476A3789525C29088D">
<treatmentCitationGroup id="915A9A3F65FDC9CFDEF97AD56673F485">
<treatmentCitation id="4A9C7525FC2A560FECB1ED2BD00A361E" author="Harold E" page="126" year="1872">
<taxonomicName id="213F023B508B350A9B40488159C1E4E4" ID-CoL="7CWTQ" authority="Harold, 1872: 126" authorityName="Harold" authorityPageNumber="126" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="4AF456D29BA5DC8AC5AFA2308027A199" italics="true">Trox batesi</emphasis>
<comment id="2E56592B4811A3B2D7AB9B67207843D4">
<bibRefCitation id="F817089CEA34F4F71DF0BB82AC7E18DB" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872: 126</bibRefCitation>
(original description);
<treatmentCitation id="239DDA19717C9EC90F54CFE751592B1C" author="Arrow GJ" page="54" year="1912">
<bibRefCitation id="45ADC035C5E4F5871F9663711D7CCB68" author="Arrow GJ" firstAuthor="Arrow" journalOrPublisher="Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Berlin: W. Junk" page="54" refId="B3" refString="Arrow GJ (1912) Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 43. Scarabaeidae: Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. Berlin: W. Junk, 66 pp." title="Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 43." year="1912">Arrow 1912: 54</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="CEE4D830B1C1DDAB569A99AEB3E4FFC8" author="Blackwelder RE" page="218" year="1944">
<bibRefCitation id="0766F4392EF3517A57937CA337A4A17A" DOI="10.5479/si.03629236.185.2" author="Blackwelder RE" firstAuthor="Blackwelder" journalOrPublisher="Bulletin of the United States National Museum" page="218" pagination="218-219" refId="B10" refString="Blackwelder RE (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Scarabaeidae, Troginae. Bulletin of the United States National Museum 185: 218–219. https://doi.org/10.5479/si.03629236.185.2" title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Scarabaeidae, Troginae." volume="185" year="1944">Blackwelder 1944: 218</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="C5106E47BB89F04CC7F2BDAC1E83786D" author="Vaurie P" page="154" year="1962">
<bibRefCitation id="CCF4719E9D0B04AC59B2D6EEF60869C2" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" page="154" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105–167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962: 154</bibRefCitation>
</treatmentCitation>
(redescription).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="1FEF7DA64A61FB96DB335678AE25DAA5">
<treatmentCitationGroup id="CDFE8CCDD95705D629AD9AB21AA80484">
<treatmentCitation id="6798AC2A9E9A92A89B68B4C55D45502D">
<taxonomicName id="B79E35AC5192185C0E3D3C24BBC5BC0D" ID-CoL="7CWTQ" authorityName="Harold" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi" subGenus="Omorgus">
<emphasis id="66FF06731FBEA2EBD818E3D985AC730B" italics="true">Trox</emphasis>
(
<emphasis id="354C0FB483F3977EA2EA92F618E9EBDF" italics="true">Omorgus</emphasis>
)
<emphasis id="0A4ED761E89B0B3BDF523A4A3C2F7D6C" italics="true">batesi</emphasis>
</taxonomicName>
<comment id="0B3EC5E1435751E3261F1B655B29FABD">
:
<treatmentCitation id="2C0BDC587ADC88549CD84C990960894F" author="Scholtz CH" page="8" year="1982">
<bibRefCitation id="9D9BF70B015E18B53B35D00A2EE2296E" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa" page="8" pagination="1-27" refId="B96" refString="Scholtz CH (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa 54: 1–27." title="Catalogue of world Trogidae (Coleoptera: Scarabaeoidea)." volume="54" year="1982">Scholtz 1982: 8</bibRefCitation>
</treatmentCitation>
(catalogue).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E552B1F79CB0BF14010A8984E9D59C50">
<treatmentCitationGroup id="489BA9630C7115C7BEA3DAF5719E8637">
<treatmentCitation id="00651084B88D851637C277FCC1BAEBBB">
<taxonomicName id="A11BC77431E4CDC71906759F27053F7F" ID-CoL="49DQD" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi" subGenus="Haroldomorgus">
<emphasis id="0582ACB32A01E8D39BD257ACC3617CDA" italics="true">Omorgus</emphasis>
(
<emphasis id="E6248EB8D433B501A64E226037F68690" italics="true">Haroldomorgus</emphasis>
)
<emphasis id="04BB2877B3F1F142CAFD8056E3F9C8F3" italics="true">batesi</emphasis>
</taxonomicName>
<comment id="39E5D5B0EB51354EF40906981B749332">
:
<treatmentCitation id="71EB29838CC2C4765CB2C54222C4F117" author="Scholtz CH" page="362" year="">
<bibRefCitation id="05D83253B29A3F4904A2799EDE2F1437" DOI="10.1111/j.1365-3113.1986.tb00186.x" author="Scholtz CH" journalOrPublisher="Systematic Entomology" pagination="355-363" refId="B97" refString="Scholtz CH (1986 a) Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology 11: 355–363. https://doi.org/10.1111/j.1365-3113.1986.tb00186.x" title="Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea)." volume="11">Scholtz 1986 a</bibRefCitation>
: 362
</treatmentCitation>
(phylogenetics);
<treatmentCitation id="9B7AD59A87EA40B88C7054D375CB6578" author="Scholtz CH" page="1418" year="1990">
<bibRefCitation id="437066C765781BB89A018AD672A376B4" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" page="1418" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990: 1418</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="C6DC83BBDAB422F88D143BB3D70F129A" author="Zidek J" page="7" year="2013">
<bibRefCitation id="3874947F874FA806A22B77A1293E7289" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="7" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1–38." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 7</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="27D203EE47F7BEF2EBE49402EFC37D0C" author="Strümpher WP &amp; Sole CL &amp; Villet MH &amp; Scholtz CH" page="549" year="2014">
<bibRefCitation id="4820DB48A01CE31F5A152A50AE25F147" DOI="10.1111/syen.12074" author="Strümpher WP &amp; Sole CL &amp; Villet MH &amp; Scholtz CH" etAl="et al." firstAuthor="Strumpher" issue="3" journalOrPublisher="Systematic Entomology" page="549" pagination="548-562" refId="B104" refString="Strümpher WP, Sole CL, Villet MH, Scholtz CH (2014) Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data. Systematic Entomology 39 (3): 548–562. https://doi.org/10.1111/syen.12074" title="Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data." volume="39" year="2014">Strümpher et al. 2014: 549</bibRefCitation>
</treatmentCitation>
(systematics);
<treatmentCitation id="1F3F915DA6484672EF592E924FDFFC78" author="Strümpher WP &amp; Villet MH &amp; Sole CL &amp; Scholtz CH" page="62" year="2016">
<bibRefCitation id="D0F7218971895599127F4F45C6DF0A59" DOI="10.1163/1876312X-46052133" author="Strümpher WP &amp; Villet MH &amp; Sole CL &amp; Scholtz CH" etAl="et al." firstAuthor="Strumpher" issue="1" journalOrPublisher="Insect Systematics &amp; Evolution" page="62" pagination="53-82" refId="B105" refString="Strümpher WP, Villet MH, Sole CL, Scholtz CH (2016) Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea). Insect Systematics &amp; Evolution 47 (1): 53–82. https://doi.org/10.1163/1876312X-46052133" title="Overview and revision of the extant genera and subgenera of Trogidae (Coleoptera: Scarabaeoidea)." volume="47" year="2016">Strümpher et al. 2016: 62</bibRefCitation>
</treatmentCitation>
(systematics);
<treatmentCitation id="3E09C5246716990D974F80A36C0ADCB0" author="Zidek J" page="98" year="2017">
<bibRefCitation id="34DCA27B00B23F688E262119DD6813C7" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="98" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 98</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="E6D16DDCFE33C5301932A7BD18EEDA07" author="Hielkema AJ &amp; Hielkema MA" page="10" year="2019">
<bibRefCitation id="3029D5D0547DDCE5814341F9E3E6C7FB" author="Hielkema AJ &amp; Hielkema MA" firstAuthor="Hielkema" journalOrPublisher="Insecta Mundi" page="10" pagination="1-306" refId="B53" refString="Hielkema AJ, Hielkema MA (2019) An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732: 1–306." title="An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas." volume="732" year="2019">Hielkema and Hielkema 2019: 10</bibRefCitation>
</treatmentCitation>
(checklist for the Guianas);
<treatmentCitation id="11CBBCF171BD5B876F32D002CB857293" ID-DOI="http://doi.org/10.5281/zenodo.5030752" ID-Zenodo-Dep="5030752" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" httpUri="http://treatment.plazi.org/id/03DA2410FFAAFFE3FE95FE32FD431855" page="1997" year="2021">
<bibRefCitation id="E999BF3E46A51F460CBFF75630FE2EEF" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="1997" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021: 1997</bibRefCitation>
</treatmentCitation>
(review of Brazilian species).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="F2BFDCF0C7F933D6BB7FEA4CA30ADDA7">
<treatmentCitationGroup id="A56E8793CB213AFB437E41016FB53726">
<treatmentCitation id="3E555C8DDF9A3EAFE005F9279AECA269">
<taxonomicName id="F8DE3757E6E23C040DAEEEF86F591B70" ID-CoL="49DQD" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="B9B9B003D7E3AE8F0505A735E057B2CD" italics="true">Omorgus batesi</emphasis>
</taxonomicName>
<comment id="1A90622963B1F14EB8952E7715BFE2A9">
:
<treatmentCitation id="D5493AA2EF33D14F4D6A4AD34574CA2D" author="Deloya C" page="122" year="2005">
<bibRefCitation id="A56520A56F70DA122DF40CA0A042076B" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121–129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESHAE" type="Type specimen examined">
<paragraph id="54B66FCCFAFE2500EB8B568E4B97628F">
<heading id="6E59C1732FAA405479C6F3FC4AC02A70" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="1F52F2EA4FF344F7547EA0CE47BC448B">
<materialsCitation id="B0699F9D505DE7A0E4C67E48A369981A" collectionCode="MNHN" location="See" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="holotype">
<emphasis id="97E75DBBE64257BA4D2C087C0063A43B" bold="true" italics="true">
<typeStatus id="B7AE2040C56DB441D98DAD959A2130C3" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="E2B0157FC712C4EA1960F9E51E4E4CC8" count="1" type="male">♂</specimenCount>
<named-content id="A64F786CC72E9580701AE5E670E70EE4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum national d’Histoire naturelle, Paris">
<emphasis id="98E72B59F2BF38459DC2FA93791EBFC3" bold="true">
<collectionCode id="465F18F8DAA9B43E1830EF5B1A96440D" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</emphasis>
</named-content>
).
<location id="FCACA370276EF1D7D07440550412AB42" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2:FCACA370276EF1D7D07440550412AB42" name="See" stateProvince="Amazonas">See</location>
<bibRefCitation id="D0036923992B3EDB9854EED693A3408A" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="1997" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021: 1997)</bibRefCitation>
for high-quality images of the type specimen.
<emphasis id="759151E0DEA0E6D0B2AE25532050756C" bold="true">Type locality</emphasis>
: “ Amazonestrom ” [
<collectingRegion id="19C4BB1046DC8B3BE33002FE690C6C39" country="Peru" name="Amazonas">Amazonas</collectingRegion>
River].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIIAE" type="distribution">
<paragraph id="2FACC83005367B2E2957795391E05EA6">
<heading id="51878925EF05374370DEE0A335A7B196" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="3496C75F20C14C92C47006454221F112">
<collectingCountry id="46A1F7963692B5116DBB2304AE66FDFF" name="Argentina">Argentina</collectingCountry>
,
<collectingCountry id="E07B7FC7676D921699F2600EAD05793D" name="Bolivia">Bolivia</collectingCountry>
,
<collectingCountry id="0866953D3D982EDE9F8FC4A7132E8348" name="Brazil">Brazil</collectingCountry>
and
<collectingCountry id="67722E7B65FF79275F7545766DCFFCBC" name="Paraguay">Paraguay</collectingCountry>
(
<bibRefCitation id="AB40FA841FF9D3CE5B0B05671C8D515E" author="Zidek" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017</bibRefCitation>
;
<bibRefCitation id="F92A5997C1C4994F2091DC64ACC268D9" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWIAE" type="type material">
<paragraph id="2930A1EC1954BF7380153AE6F0FC88FD">
<heading id="60284A0E5C2F7424EA560C28F485250C" reason="title">Non-type examined material</heading>
</paragraph>
<paragraph id="A9ACC2DE8B54796CA65C55D3847AC080">
<emphasis id="B24FDD138C753F035E174AE089122149" bold="true">
(
<specimenCount id="483D60EBC816F1CE3666C997A9D1F38E" count="4" type="generic">four specimens</specimenCount>
additionally to
<bibRefCitation id="49BFD1558FE4E943188AFEC646715D76" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz (1990)</bibRefCitation>
and
<bibRefCitation id="9D77B9BEEE9C0537F841026521CAF9D9" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021)</bibRefCitation>
).
</emphasis>
<materialsCitation id="1C04C9A866F4D95EF706A062EA2257B0" country="Brazil" location="Amazonas" specimenCount="1" stateProvince="Amazonas">
<collectingCountry id="81D96254E8C59CE9512F61C8B34004B9" name="Brazil">BRAZIL</collectingCountry>
–
<emphasis id="604E570B3E1D66EC050A101E5E42502D" bold="true">
<collectingRegion id="691B60F8459693157383967D0F30AC0D">Amazonas</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="C596DD577D8A061E0E358F1B5152210B" collectingDate="1969-11-08" collectionCode="A, CNC" collectorName="Jameson ML &amp; Campbell" country="Brazil" location="Eva" specimenCount="1" stateProvince="Parana">
1;
<collectingRegion id="F61D637CC6346328D480D2686D4BD2B5" country="Brazil" name="Parana">Parana</collectingRegion>
da
<location id="5EBBDF5FC2C89D28ACC225224EFE520B" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2:5EBBDF5FC2C89D28ACC225224EFE520B" country="Brazil" name="Eva" stateProvince="Parana">Eva</location>
;
<date id="1F4AC7AE4F0C1A5AADC40E0D430D96C7" value="1969-11-08">
<collectingDate id="4961A59397E60CF71440315A066CAEFA" value="1969-11-08">8 Nov. 1969</collectingDate>
</date>
;
<collectorName id="F21EF3035214F6938EBC642146FE4A09" name="Jameson ML">J. M.</collectorName>
&amp; B.
<collectionCode id="C7A360541499C4308F575536981BD3B8">A</collectionCode>
.
<collectorName id="DE831129E126E66B8578732E0AAACEAF">Campbell</collectorName>
leg.;
<named-content id="A0FBB658F861BDB889F1265E4758524F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection of Insects">
<collectionCode id="368F226CA005239B79DDC9229A05BDAE" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
</named-content>
. –
<emphasis id="6FF498D99D04E89C46867106D47030F5" bold="true">
<collectingRegion id="A7627F64786029B29F793B78A70CDE61" country="Brazil" name="Mato Grosso">Mato Grosso</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="5F6A384B081D57B69FAF82523A56111F" collectingDate="2022-10-01" collectionCode="CEMT" collectorName="K. R. Ferreira" country="Brazil" location="Juina" specimenCount="1" stateProvince="Amazonas">
1;
<location id="4F819FB0167B2C8450793960ED8477F0" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2:4F819FB0167B2C8450793960ED8477F0" country="Brazil" name="Juina" stateProvince="Amazonas">Juina</location>
;
<date id="EA13D9AE731F9D3BD460A02BB68C9543" value="2022-10-01">
<collectingDate id="493493A2698F5C767EF9145806DD8ADB" value="2022-10-01">1 Oct. 2022</collectingDate>
</date>
;
<collectorName id="2EC9E1B572BDE27BB1ED1FE92107C38A">K. R. Ferreira</collectorName>
leg;
<abbrev id="ABBRID0EPJAE" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="4DB5D43D5F2552A68C934D9952648640">CEMT</collectionCode>
</abbrev>
</materialsCitation>
•
<materialsCitation id="E9DFF68FFEF76BCE6BCAB59BC6AE24F7" collectingDate="2024-01-11" collectingDateMax="2024-01-17" collectingDateMin="2024-01-11" collectionCode="CEMT" collectorName="D. F. Rodrigues &amp; T. I. Vitor-da-Silva" country="Brazil" latitude="-9.761389" location="Faz. da Ilha" longLatPrecision="21" longitude="-52.26139" municipality="Santa Cruz do Xingu" specimenCount="1" stateProvince="Rio de Janeiro">
1;
<collectingMunicipality id="9EA9DA32C7EED12DA116ECE21F406FE4">Santa Cruz do Xingu</collectingMunicipality>
,
<location id="8BCF9027FAB8EF3E2B00AAD941214C59" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2:8BCF9027FAB8EF3E2B00AAD941214C59" country="Brazil" latitude="-9.761389" longLatPrecision="21" longitude="-52.26139" municipality="Santa Cruz do Xingu" name="Faz. da Ilha" stateProvince="Rio de Janeiro">Faz. da Ilha</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWJAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-52.261389,-9.761389]}">
<geoCoordinate id="A87E4344BDBD7ACD0E134A8B63DDB42C" degrees="9" direction="south" minutes="45" orientation="latitude" precision="15" seconds="41" value="-9.761389">9 ° 45 ' 41 &quot; S</geoCoordinate>
,
<geoCoordinate id="81083514922CE537C8ECF11BA81D3F5E" degrees="52" direction="west" minutes="15" orientation="longitude" precision="15" seconds="41" value="-52.26139">52 ° 15 ' 41 &quot; W</geoCoordinate>
</named-content>
;
<collectingMethod id="95B49C19D90A5763E56F3ED60CDA6D91">UV light trap</collectingMethod>
;
<date id="A7C32D1D6CC95E1834676A267FB6266C" value="2024-01-11" valueMax="2024-01-17" valueMin="2024-01-11">
<collectingDate id="00972538014D0C9CA055371648D6446A" value="2024-01-11" valueMax="2024-01-17" valueMin="2024-01-11">11–17 Jan. 2024</collectingDate>
</date>
;
<collectorName id="442F052A9A0FD2B5C70FFC5112DBAEA9">D. F. Rodrigues</collectorName>
&amp;
<collectorName id="09FB234272DF1CFB048E1690EFAF116D">T. I. Vitor-da-Silva</collectorName>
leg.;
<abbrev id="ABBRID0E2JAE" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="B0008BB4687853B58A080868F1241F4A">CEMT</collectionCode>
</abbrev>
. –
<emphasis id="836133B69DA0E4E041A1D82BBDC76C72" bold="true">
<collectingRegion id="76A87E0D415349E688BFC023FC53C646" country="Brazil" name="Rio de Janeiro">Rio de Janeiro</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="01D6DC5AB2E502C72F68CACC7343C1B6" collectingDate="1999-10-15" collectingDateMax="1999-11-10" collectingDateMin="1999-10-15" collectionCode="NHMUK" collectorName="W. Grosser" country="Brazil" location="Mage" specimenCount="1" specimenCount-female="1" stateProvince="Rio de Janeiro">
<specimenCount id="1B8876D3B5554028EBF86FC9FC99AEED" count="1" type="female">1 ♀</specimenCount>
;
<location id="9F97267D65E2569387024A491E0E2B8A" LSID="urn:lsid:plazi:treatment:AAE41682797B55719EB05CFA4076CFA2:9F97267D65E2569387024A491E0E2B8A" country="Brazil" name="Mage" stateProvince="Rio de Janeiro">Magé</location>
,
<quantity id="64542373FFE67E3B1F4F4A9E50F54B55" metricMagnitude="4" metricUnit="m" metricValue="7.0" unit="km" value="70.0">70 km</quantity>
N [north] of
<collectingRegion id="D672974F6221E9DAF74F23D0F57ADF04" country="Brazil" name="Rio de Janeiro">Rio de Janeiro</collectingRegion>
;
<date id="FC3ACC23E81456089A4A0BCF6463DA50" value="1999-10-15" valueMax="1999-11-10" valueMin="1999-10-15">
<collectingDate id="5B0BE7A3D73FEDD51F16B3BFE217542F" value="1999-10-15" valueMax="1999-11-10" valueMin="1999-10-15">15 Oct. – 10 Nov. 1999</collectingDate>
</date>
;
<collectorName id="C2F1E24E107F6A5274508E8535E8C64F">W. Grosser</collectorName>
leg.;
<named-content id="28E757B24B40347A777EC92E5344945E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="9061BB9BA0BE0D7303CF40186BE78BDF" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFKAE" type="notes">
<paragraph id="9C12FF3D4E641624810228FD12090448">
<heading id="5A727C8499411A245A21365875D79F37" reason="title">Note.</heading>
</paragraph>
<paragraph id="FA819C789A590BE073DE666F79E51A6B">
The monotypic subgenus
<taxonomicName id="1191D229115F5DE0D32C73596305B57F" authority="Scholtz, 1986 a" authorityName="Scholtz" authorityYear="1986" rank="subGenus" subGenus="Haroldomorgus">
<emphasis id="030685FA542DF0CBB1862C6F7DC8B1F1" italics="true">Haroldomorgus</emphasis>
Scholtz, 1986 a
</taxonomicName>
was established to accommodate
<taxonomicName id="98F2235B1AC6BB14C93D0F993887CFB4" authority="Harold, 1872" authorityName="Harold" authorityYear="1872" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="70E51C81F4AD5631F3D9F64D25299CAB" italics="true">Trox batesi</emphasis>
<bibRefCitation id="5C5200841EA701E055F503BF8B3E7DDE" author="Harold E" journalOrPublisher="Coleopterologische Hefte" pagination="1-192" refId="B49" refString="Harold E (1872) Monographie der Gattung Trox. Coleopterologische Hefte 9: 1 - 192." title="Monographie der Gattung Trox." volume="9" year="1872">Harold, 1872</bibRefCitation>
</taxonomicName>
, following a morphological phylogenetic analysis aimed at elucidating the suprageneric relationships within
<taxonomicName id="8988907B2F0F6D953645983D047FBE11" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
(refer to Scholtz, 1986 a for details). One of the most distinctive features of
<taxonomicName id="86EC15DBF40EAAF49E49CF9D69B7EDA6" authorityName="Scholtz" authorityYear="1986" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="subGenus" subGenus="Haroldomorgus">
<emphasis id="7D70C348A148E13282BDCDFF06666EEE" italics="true">Omorgus</emphasis>
(
<emphasis id="FBDB75CD4942CE3178DBD2FD8D4D80B7" italics="true">Haroldomorgus</emphasis>
)
</taxonomicName>
species is a complete absence of tubercles or ridges on the pronotum and elytra. Additionally, this species exhibits an oval-shaped scutellum that is not constricted at the base, which distinguishes it from the typical modern concept of
<taxonomicName id="4BA3C87E50059230618065039D1EFB8E" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C6EA6C9FC187D5219D89A6629E13C3CB" italics="true">Omorgus</emphasis>
species.
</taxonomicName>
Following a morphological phylogenetic analysis conducted by some of the authors (
<collectionCode id="5E0C3343E9AE24BE74AB8488A083531F">VCS</collectionCode>
, WPS, and FZVM –
<bibRefCitation id="DD2A7732C2543293748529819F10B38A" author="Costa-Silva V &amp; Strümpher WP &amp; Thyssen PJ &amp; Vaz-de-Mello FZ" refId="B28" refString="Costa-Silva V, Strümpher WP, Thyssen PJ, Vaz-de-Mello FZ (in press) Phylogenetic systematics of the keratin-feeding genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae). Zoological Journal of the Linnean Society.">Costa-Silva et al. in press</bibRefCitation>
), the genus
<taxonomicName id="7C26A7022E01C3DA8EC872C74C6DFE68" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="0A17E395206F26D152CEA9A7DA8D8231" italics="true">Omorgus</emphasis>
</taxonomicName>
was recovered to be paraphyletic, restoring
<taxonomicName id="EA2E9B6DF4C8965868285E5F7004844B" authorityName="Scholtz" authorityYear="1986" rank="subGenus" subGenus="Haroldomorgus">
<emphasis id="6C091A55D3620B9DBB34209FA0B3C5AA" italics="true">Haroldomorgus</emphasis>
</taxonomicName>
as a sister group to
<taxonomicName id="64126C94FFD2526C398778F9E8859AB0" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="55D5EA04CAB9911D7C0FE5DAB8BC5A37" italics="true">Omorgus</emphasis>
</taxonomicName>
+
<taxonomicName id="2E8F67F4D087DEB75EC590E404056D07" authorityName="Burmeister" authorityYear="1876" class="Insecta" family="Trogidae" genus="Polynoncus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="8D52D92AB81C5C5AB439E1FF7CE71F48" italics="true">Polynoncus</emphasis>
</taxonomicName>
. In the first and only molecular phylogeny of
<taxonomicName id="41FE7EDBD778203F8D6D605DA9E2EB0A" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
performed to date, by
<bibRefCitation id="81E2070E34F893CCC367BFC8D0C71B0E" DOI="10.1111/syen.12074" author="Strümpher WP &amp; Sole CL &amp; Villet MH &amp; Scholtz CH" etAl="et al." firstAuthor="Strumpher" issue="3" journalOrPublisher="Systematic Entomology" pagination="548-562" refId="B104" refString="Strümpher WP, Sole CL, Villet MH, Scholtz CH (2014) Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data. Systematic Entomology 39 (3): 548–562. https://doi.org/10.1111/syen.12074" title="Phylogeny of the family Trogidae (Coleoptera: Scarabaeoidea) inferred from mitochondrial and nuclear ribosomal DNA sequence data." volume="39" year="2014">Strümpher et al. (2014)</bibRefCitation>
, no specimens of
<taxonomicName id="A84DF574A01CD9351EED5EB7C8E95C48" baseAuthorityName="Harold" baseAuthorityYear="1872" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="batesi" subGenus="Haroldomorgus">
<emphasis id="8AAF9D706FAD7BBE758A6FC92D638B6A" italics="true">Omorgus</emphasis>
(
<emphasis id="5D679551323FABE55EFFFC0A28EE723E" italics="true">Haroldomorgus</emphasis>
)
<emphasis id="AA23D8D4B43B9864AABC1C2B127CC5CD" italics="true">batesi</emphasis>
</taxonomicName>
were used in the analysis. However, we have now obtained fresh specimens to revisit the molecular analysis and finally test the
<bibRefCitation id="670856A99D9105E9BE9D8CABF78F7581" DOI="10.1111/j.1365-3113.1986.tb00186.x" author="Scholtz CH" journalOrPublisher="Systematic Entomology" pagination="355-363" refId="B97" refString="Scholtz CH (1986 a) Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology 11: 355–363. https://doi.org/10.1111/j.1365-3113.1986.tb00186.x" title="Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea)." volume="11">Scholtz’s (1986 a</bibRefCitation>
) hypothesis and the position of
<taxonomicName id="4A4535816AC1F4140677D82FC8544AB2" authorityName="Scholtz" authorityYear="1986" rank="subGenus" subGenus="Haroldomorgus">
<emphasis id="C0F7A454FF5BFD4A2C47A18FDED47DB8" italics="true">Haroldomorgus</emphasis>
</taxonomicName>
within
<taxonomicName id="8D1B4137ED52B76F8DA9FDA49502A5E2" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

330
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<taxonomicName id="34AE3608F3880DC88CE7C3D372B2DE20" ID-CoL="49DRH" authority="Scholtz, 1990" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="indigenus" subGenus="Omorgus">
<emphasis id="DE7AA231F304ACEE8C62E593BAD74D02" italics="true">Omorgus</emphasis>
(
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)
<emphasis id="D31FD0E59C1F936A2EACF37322AB69D3" italics="true">indigenus</emphasis>
<bibRefCitation id="D19EC2B26EDDFD40313C066A059BEC1F" DOI="10.1080/00222939000770841" author="Scholtz CH" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391 - 1456. https: // doi. org / 10.1080 / 00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz, 1990</bibRefCitation>
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<paragraph id="83EE0FEE56B6AA67647C9F6C0ADA7834">
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</paragraph>
</subSubSection>
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<paragraph id="E9585441FC5061247706F91166720FDF">
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<treatmentCitation id="76CB5D0DD6839A26568827687AFCD6E9" author="Scholtz CH" page="1411" year="1990">
<taxonomicName id="6F4D785E9221EA175BD93EA69607BB66" ID-CoL="49DRH" authority="Scholtz 1990: 1411" authorityName="Scholtz 1990:" authorityPageNumber="1411" authorityYear="1411" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="indigenus" subGenus="Omorgus">
<emphasis id="9A0CF510202397D9A9BFA953F36CEEF0" italics="true">Omorgus</emphasis>
(
<emphasis id="120ABD90D7A963121CC583EDAAAFC07A" italics="true">Omorgus</emphasis>
)
<emphasis id="D0F3ADBE127E043D31D219FD1F68D053" italics="true">indigenus</emphasis>
<comment id="1B6F7203731FFA62CB2AE37549545B0A">
<bibRefCitation id="A9941F7C71C0CDB095856A6A35CF2B03" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" page="1411" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990: 1411</bibRefCitation>
(original description);
<treatmentCitation id="8625EC4FB5418EFFD25AA4F523BCB4F9" author="Zidek J" page="11" year="2013">
<bibRefCitation id="6C26B44EEC727ACC626B54A973DAD3DE" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="11" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1–38." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 11</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="57024DCBC42CEB35907EC71E09B7CBC0" author="Zidek J" page="102" year="2017">
<bibRefCitation id="2AAF6FAB501DE81AFBFD4521845D03FB" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="102" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 102</bibRefCitation>
</treatmentCitation>
(checklist).
</comment>
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</paragraph>
<paragraph id="907FFEBB0011282C2BAB97DABA882EE8">
<treatmentCitationGroup id="F5FE837D3DDF72F44846AED267A8BB83">
<treatmentCitation id="B3525DFD19B6B4C39D5FA853CAF52FA1">
<taxonomicName id="E18CB328D234437901F829293BD7ED23" ID-CoL="49DRH" authorityName="Scholtz 1990:" authorityYear="1411" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="indigenus">
<emphasis id="71C53E8C952551DA15A2A2C8BF629274" italics="true">Omorgus indigenus</emphasis>
</taxonomicName>
<comment id="6757F5DC9BF6F13C3CB63C35FAFC4279">
:
<treatmentCitation id="920A67688DFD082579609BF4C4C1C9F6" author="Morrone JJ" page="63" year="2001">
<bibRefCitation id="14108E38F62E9726345CD44ED0114D5E" author="Morrone JJ" firstAuthor="Morrone" issue="1" journalOrPublisher="Caldasia" page="63" pagination="43-76" refId="B71" refString="Morrone JJ (2001) Toward a cladistic model for the Caribbean subregion: delimitation of areas of endemism. Caldasia 23 (1): 43–76." title="Toward a cladistic model for the Caribbean subregion: delimitation of areas of endemism." volume="23" year="2001">Morrone 2001: 63</bibRefCitation>
</treatmentCitation>
(mention);
<treatmentCitation id="69E5A0D8DBC70E6332AA103C8F7E61C7" author="Deloya C" page="122" year="2005">
<bibRefCitation id="53978BDDBB9B20D070F3BB1C8D8BFEE4" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121–129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="44A73A77A4A4CA02D80739FF87A91569" author="Morrone JJ" page="62" year="2014">
<bibRefCitation id="9AD07F2BDC50F0D24E32BFDC9576E94E" DOI="10.11646/zootaxa.3782.1.1" author="Morrone JJ" firstAuthor="Morrone" issue="1" journalOrPublisher="Zootaxa" page="62" pagination="1-110" refId="B72" refString="Morrone JJ (2014) Biogeographical regionalisation of the Netropical region. Zootaxa 3782 (1): 1–110. https://doi.org/10.11646/zootaxa.3782.1.1" title="Biogeographical regionalisation of the Netropical region." volume="3782" year="2014">Morrone 2014: 62</bibRefCitation>
</treatmentCitation>
(mention, biogeography).
</comment>
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</paragraph>
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<subSubSection id="SECID0EZKBG" type="material">
<paragraph id="6102830EA573F74C84DF08276FEEA770">
<heading id="3DA9EBBB345194AF9D48345D65FA9991" reason="title">Type specimens examined.</heading>
</paragraph>
<paragraph id="F9FE2DCD22A9547407828556EBCB8B64">
<materialsCitation id="88A57FBD0E761A3E8456DD0F416D6241" collectionCode="CMNC" collectorName="C. H. Scholtz" country="Ecuador" location="Bahia Manzanilla" municipality="Bahia Manzanilla" specimenCount="1" specimenCount-male="1" stateProvince="Galapagos" typeStatus="holotype">
<emphasis id="921F2720E1DCB131D4C2138382E5651D" bold="true" italics="true">
<typeStatus id="9F312ABA8FF82EF70B8C9500B866E002" type="holotype">Holotype</typeStatus>
, by original designation
</emphasis>
(
<specimenCount id="CF0378BD09C578B06BEBB92E43093EFD" count="1" type="male">♂</specimenCount>
<named-content id="18078B2D276233909744852823E0A5D3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">
<emphasis id="00EC81ABD256741C7706078DFDA4D01D" bold="true">
<collectionCode id="6B25FC5E5690CF7BA2BBB48257CC2F75">CMNC</collectionCode>
</emphasis>
</named-content>
). First label [white, typeset]: “ ECU: Galap; Espanola /
<location id="6660461765F2935488AD19334C4DDFE9" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:6660461765F2935488AD19334C4DDFE9" country="Ecuador" municipality="Bahia Manzanilla" name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
/ 5–10. VI. 85, S &amp; J Peck / Prospis grove behind / beach,
<collectingMethod id="F980E25BA8E0A7CDEE7CEDF06C7D894B">carrion traps</collectingMethod>
”. Second label [white with black border, printed]: “ [QR code] / Canadian Museum of / Musée canadien de la / NATURE / CMNCEN 00011596 ”. Third label [white with red border, Clarke Scholtz’s handwriting]:
<taxonomicName id="BBB7C4FC3011C49ECC4A8C607AA902EC" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="CA392913041E100B48D5933B1F6507D7" italics="true">Omorgus</emphasis>
</taxonomicName>
/ indigenous /
<collectorName id="D778236E2D5D2118CB6C6778CF6F4770">C. H. Scholtz</collectorName>
1988 /
<typeStatus id="5E5BF3ADF8E49D30F1F91F24F5F78385" type="holotype">HOLOTYPE</typeStatus>
”.
<emphasis id="556357735B7F51055D12AB51B557240D" bold="true">Type locality</emphasis>
: “
<collectingMunicipality id="BC2C37A6666B2D0FC6604C19C0F2FE05">Bahia Manzanilla</collectingMunicipality>
, Española Is.,
<collectingRegion id="3F8B059B424DF3B6200649B6B9D475AD" country="Ecuador" name="Galapagos">Galapagos</collectingRegion>
” [
<collectingCountry id="A22B443ABB804748B77B5B32EEB26877" name="Ecuador">Ecuador</collectingCountry>
].
</materialsCitation>
</paragraph>
<caption id="09D680F8DC4BD196348FCE1F818EAC5F" ID-DOI="10.3897/contrib.entomol.74.e126799.figure4" ID-arpha="22753C35-15FF-5752-8358-36578E5355AB" httpUri="https://binary.pensoft.net/fig/1103866" startId="F4">
<paragraph id="84104B8C9052F7BA645DB5D039630D1E">
<label id="6858AB02C6600966F9DDB85DE4F6BCE6">Figure 4.</label>
</paragraph>
<paragraph id="BBA64780EFC96A11D24DCD67480226CE">
Holotype of
<taxonomicName id="ADA779211081E5336A091D7CF2CBA764" authority="Scholtz 1990" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="indigenus">
<emphasis id="45270672DF9D50C3BFF3A6734A0FF3AF" italics="true">Omorgus indigenus</emphasis>
<bibRefCitation id="78BFF2D7AA43EA7B9C4D506925770432" DOI="10.1080/00222939000770841" author="Scholtz" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990</bibRefCitation>
</taxonomicName>
.
<emphasis id="4F5F7878C0FCB07EA380AB44DA13586C" bold="true">A.</emphasis>
Habitus in dorsal view;
<emphasis id="BDC5BD46BCCA9987DC77EDC48A192E8C" bold="true">B.</emphasis>
Habitus in lateral view;
<emphasis id="8D4F1A958C359AE4DD56F177D3E82BB5" bold="true">C.</emphasis>
Aedeagus in dorsal view;
<emphasis id="323D11CB7ED574D91025C46CDE22C587" bold="true">D.</emphasis>
Aedeagus in lateral view;
<emphasis id="0EF19F521B6FCD9EEE474C0847C10F59" bold="true">E.</emphasis>
Labels. Photos by François Génier (
<named-content id="75DC47F8695132F9025D60739F22A313" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">CMNC</named-content>
).
</paragraph>
</caption>
<paragraph id="5DC3A163E6C012956E454402D0482C71">
<materialsCitation id="8C220CD440F25A5EF6438ADAE0608C8F" country="Ecuador" location="Galapagos" specimenCount="1" stateProvince="Galapagos" typeStatus="paratype">
<emphasis id="6FE4887AD17D8B2B91B5D3F3A6A04C8B" bold="true" italics="true">
<typeStatus id="A1004E3D6BDDBF47E9D38B0A0498B26E" type="paratype">Paratypes</typeStatus>
examined
</emphasis>
:
<collectingCountry id="3F5341C5AB1D6E41EFD0889D62B186FA" name="Ecuador">ECUADOR</collectingCountry>
–
<emphasis id="8A58CEE7BA30B113E7556FB89791BA0A" bold="true">
<collectingRegion id="21D49FDAEFE54C6F7EBC5E3B8B5C768A" country="Ecuador" name="Galapagos">Galapagos</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="0FC5DF0230B647B7E5B1CC027A25228F" collectingDate="1985-06-05" collectingDateMax="1985-06-10" collectingDateMin="1985-06-05" collectionCode="CMNC" collectorName="Espanola Is. &amp; Prosopis &amp; S. &amp; J. Peck" country="Ecuador" location="Bahia Manzanilla" specimenCount="1" stateProvince="Galapagos" typeStatus="paratype">
15;
<collectorName id="CA86002861929AEE1A73DD3A7D4DC637">Española Is.</collectorName>
,
<location id="231F03F65B72598682A0CC00B5670B30" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:231F03F65B72598682A0CC00B5670B30" country="Ecuador" name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
;
<collectorName id="C36DE96F24C1BEFEB57A2D7CA28867DD">Prosopis</collectorName>
grove behind beach;
<collectingMethod id="A22090C2F9A540311667884F14FED301">carrion traps</collectingMethod>
;
<date id="F72110ED87B4E181D1970CF23A377C4A" value="1985-06-05" valueMax="1985-06-10" valueMin="1985-06-05">
<collectingDate id="29DCA970E2DA1B761302E6F610D4F84D" value="1985-06-05" valueMax="1985-06-10" valueMin="1985-06-05">5–10 Jun. 1985</collectingDate>
</date>
;
<collectorName id="A98AEC5FABFAC1E48CFEE654262F8041">S. &amp; J. Peck</collectorName>
leg.;
<named-content id="790685DFBCEC067AB6BE95DCAA98B9ED" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">
<collectionCode id="7D83E59504F11EB42062C2E83A7CF17A">CMNC</collectionCode>
</named-content>
</materialsCitation>
•
<materialsCitation id="DB749C411A534F2B888CE41F004CE320" collectingDate="1985-06-05" collectingDateMax="1985-06-10" collectingDateMin="1985-06-05" collectionCode="NHMUK" collectorName="Espanola Is. &amp; Prosopis &amp; S. &amp; J. Peck" country="Ecuador" location="Bahia Manzanilla" specimenCount="1" stateProvince="Galapagos" typeStatus="paratype">
2; same collection data;
<named-content id="96A8BEA9D5E6BC626A4B77B4B14474EF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="FF12F6FE37C8D1C8AE5F4271B3325695" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMNBG" type="distribution">
<paragraph id="4937172E983DF1229FBF1B7CDA72D898">
<heading id="F33BAA34E8399D3E1708ADBE0CBC5A8E" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="5A00485E1F13B5C2F6BFFC7BF729BBC7">
Endemic to
<collectingRegion id="BCD46A7A89DB0B68960DB8D2DEA24F92" country="Ecuador" name="Galapagos">Galapagos Islands</collectingRegion>
[
<collectingCountry id="966430EF206384988613E7F10AA4EB2A" name="Ecuador">Ecuador</collectingCountry>
].
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERNBG" type="type material">
<paragraph id="BDE944E30923699829AAB2C29F3F9D76">
<heading id="55CBC7378A139098562E7A78452D9430" reason="title">Non-type examined material</heading>
</paragraph>
<paragraph id="1074F02AE012FD92E4C075F112170C5E">
<emphasis id="DBFF6A67DA2800C98B095F3FF8E044C8" bold="true">
(
<specimenCount id="C44DAAC69A8046B97A25CA345B52BCE7" count="24" type="generic">24 specimens</specimenCount>
).
</emphasis>
<materialsCitation id="34B020451435EE41CB728958F03B0515" country="Ecuador" location="Galapagos" specimenCount="1" stateProvince="Galapagos">
<collectingCountry id="34A9A47F13665BCD98FE6EB18B353214" name="Ecuador">ECUADOR</collectingCountry>
–
<emphasis id="B38EA99FBA0B50A24F2B7102F7BEDCB2" bold="true">
<collectingRegion id="C478DDC8F764C50328E3B0C52A64E8FD" country="Ecuador" name="Galapagos">Galapagos</collectingRegion>
</emphasis>
</materialsCitation>
•
<materialsCitation id="0AFCB62069FBB10F6EDA59BE264FF3E4" collectingDate="1985-06-05" collectingDateMax="1985-06-10" collectingDateMin="1985-06-05" collectionCode="CMNC" collectorName="Espanola Is. &amp; S. &amp; J. Peck" country="Ecuador" location="Bahia Manzanilla" specimenCount="1" stateProvince="Galapagos">
3;
<collectorName id="1130970AA8D2C3827D467416FE67288E">Española Is.</collectorName>
,
<location id="D427AFB25F630DD51B4E13CF67D76FE2" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:D427AFB25F630DD51B4E13CF67D76FE2" country="Ecuador" name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
;
<date id="71327C9CFE1BEB8E570C2732E797C412" value="1985-06-05" valueMax="1985-06-10" valueMin="1985-06-05">
<collectingDate id="C84A906A2FC3781AC1F1A5E4C843A407" value="1985-06-05" valueMax="1985-06-10" valueMin="1985-06-05">5–10 Jun. 1985</collectingDate>
</date>
;
<collectorName id="3387F94359B496A4C94A40297524DA0C">S. &amp; J. Peck</collectorName>
leg.;
<named-content id="42314BE7FBA8C273A861EA5D3B91E8C0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">
<collectionCode id="935228EF2A80AD8A2E0759F88AEFD5B8">CMNC</collectionCode>
</named-content>
</materialsCitation>
•
<materialsCitation id="09E966FDB46D10AAA453AE8F195BAFCB" collectingDate="1985-04-25" collectingDateMax="1985-05-02" collectingDateMin="1985-04-25" collectionCode="CMNC" collectorName="Espanola Is. &amp; S. Peck" country="Ecuador" location="Bahia Manzanilla" municipality="Galapagos" specimenCount="1" stateProvince="Galapagos">
15;
<collectingMunicipality id="B9F740F940ED5D7F842C9F79B4EC7112">Galapagos</collectingMunicipality>
,
<collectorName id="5E2FF14EFEEBDCAEEC0E16765E9DB1BF">Española Is.</collectorName>
,
<location id="37B408DB98959D55DFCC78A20DC9067C" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:37B408DB98959D55DFCC78A20DC9067C" country="Ecuador" municipality="Galapagos" name="Bahia Manzanilla" stateProvince="Galapagos">Bahia Manzanilla</location>
;
<date id="CCA2D3C61F56CFF8423F304844D9F9F1" value="1985-04-25" valueMax="1985-05-02" valueMin="1985-04-25">
<collectingDate id="0739CB44711063D86BC86D6BB74E7438" value="1985-04-25" valueMax="1985-05-02" valueMin="1985-04-25">25 Apr. – 2 May 1985</collectingDate>
</date>
;
<collectorName id="4ED76C4786F94297F8256A5189DD7794">S. Peck</collectorName>
leg.;
<named-content id="A8F2AED1553373B0FB08C34F0C04C3C7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">
<collectionCode id="F5D59A423C7B386BC5079E075FAF1525">CMNC</collectionCode>
</named-content>
</materialsCitation>
•
<materialsCitation id="554F6F69D632899ACAFCEA06AB4805A5" collectingDate="1992-06-02" collectionCode="CMNC" collectorName="Espanola Is. &amp; S. Peck" country="Ecuador" location="Pta. Suarez" municipality="Galapagos" specimenCount="1" stateProvince="Galapagos">
6;
<collectingMunicipality id="39C7FD0A8316B68CDBDB9EBEAAA36CA5">Galapagos</collectingMunicipality>
,
<collectorName id="6A7B41509819FB44DF3D5141CAE8BBFF">Española Is.</collectorName>
,
<location id="518CC963CCF79E9A5C46D8C0067692F4" LSID="urn:lsid:plazi:treatment:ADD0D40BE88359D1A6767E32E3629EC7:518CC963CCF79E9A5C46D8C0067692F4" country="Ecuador" municipality="Galapagos" name="Pta. Suarez" stateProvince="Galapagos">Pta. Suarez</location>
;
<date id="D542D485DA0B5E5172803572B11A8A33" value="1992-06-02">
<collectingDate id="E90B840E9A032EC60E36C642E0656B9F" value="1992-06-02">2 Jun. 1992</collectingDate>
</date>
;
<collectorName id="4D3F3F74C2F31E020AE76E51EB4329CB">S. Peck</collectorName>
leg.;
<named-content id="1632F935614F2A378B030B34C7EA93D5" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-museum-nature-3" xlink_title="Canadian Museum of Nature">
<collectionCode id="F7009DD9662AD169D58D65380ACFDD73">CMNC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

498
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<mods:affiliation id="CACB9CB349DC4AA521DC3AB83C9A56DA">Department of Environmental and Life Sciences, Trent University, 1600 West Bank Drive, Peterborough, ON, Canada &amp; Department of Physiology, School of Medicine, Wayne State University, Detroit, MI 48201, USA</mods:affiliation>
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<mods:affiliation id="515C8AE97367F50AD593731BD1D04DBD">Department of Fisheries Sciences, Faculty of Natural Resources, University of Kurdistan, Sanandaj, Iran &amp; Department of Fisheries and Aquatic Ecology, Faculty of Fisheries and Environmental Sciences, Gorgan University of Agricultural Sciences and Natural Resources, Gorgan, Iran</mods:affiliation>
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</mods:role>
<mods:namePart id="F5B7D76C54DEBAAD39CFEA89CBB4186A">Yusuf, Thameena</mods:namePart>
<mods:nameIdentifier id="1A99B40D7BA32055C3F4B25A4C527134" type="ORCID">https://orcid.org/0009-0000-6296-6693</mods:nameIdentifier>
<mods:affiliation id="96E2BB3F6A21D78370CA5FDDF06EBA0C">Department of Physiology, School of Medicine, Wayne State University, Detroit, MI 48201, USA</mods:affiliation>
</mods:name>
<mods:name id="935A7A27987E00615DD613BCA6CED45E" type="personal">
<mods:role id="41EC2212015F34C220DBA09333A65B32">
<mods:roleTerm id="E2C071BCDA8EE1573D002B951A9DCFE4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5E7AC575161A25187C92B3F7F73F475C">Vasquez, Adrian A.</mods:namePart>
<mods:nameIdentifier id="EBB6BFC3CD5179BEB4A9236CD9430C1C" type="ORCID">0000-0002-2434-3388</mods:nameIdentifier>
<mods:affiliation id="2615C77FC13145302C1EC942A8851AC5">Department of Biology, College of Liberal Arts and Sciences, Mercer University, 1501 Mercer University Drive, Macon, GA 31207, USA</mods:affiliation>
</mods:name>
<mods:name id="B5618B822E39B92ADB7B08E58A9AB7DB" type="personal">
<mods:role id="23A8D75B4BC01397FFE193CFC2559EFD">
<mods:roleTerm id="C95947B4F120674F0711BE3587AF74D2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C6B8D84B0571923DAF0A5CFFE4E53650">Ram, Jeffrey L.</mods:namePart>
<mods:nameIdentifier id="DA30D8AD90A2EB244FEE22C0BBBA5940" type="ORCID">0000-0002-1063-546X</mods:nameIdentifier>
<mods:affiliation id="836C90217F217E7CAB28A0DAB4323302">Department of Physiology, School of Medicine, Wayne State University, Detroit, MI 48201, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="4B30BC656B60672046F6E8F2FC3D03FE">text</mods:typeOfResource>
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<mods:titleInfo id="726A997E2F2097884577CE45E94DFFD4">
<mods:title id="D2ED5B89A2D654D9054CE48068EFFBA1">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="ADFBDA4392D51FCE00A92A85A6F6F329">
<mods:date id="6BF8161496385B84F52254E9EC1716AD">2024</mods:date>
<mods:detail id="A5A6388DE088340749BC0B3F00C8590A" type="pubDate">
<mods:number id="54883690A067E7119FE2F56112C2A697">2024-07-29</mods:number>
</mods:detail>
<mods:detail id="519F5FAA102DCB7823605068C888C079" type="volume">
<mods:number id="CCD60B7548254B3356C320EBAAB4F609">1208</mods:number>
</mods:detail>
<mods:extent id="9FAA0933744EC93295DC7F378104CF22" unit="page">
<mods:start id="5DE31117A9AB998F4C5B89A23F7D7B78">133</mods:start>
<mods:end id="CF7539CF39A8C7C2EAA14033A609844F">163</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0ECF604F0C1363DAF757EC37C36CCF28">journal article</mods:classification>
<mods:identifier id="DF3982F83AA474FA3862BD7722827943" type="DOI">10.3897/zookeys.1208.124495</mods:identifier>
<mods:identifier id="EF024C8FE3277C85D237F6368713750F" type="ZooBank">FFB06B11-084A-4C08-A050-AFBA7655DF2F</mods:identifier>
</mods:mods>
<treatment id="AEA57F1722625230AFA0B36F70AD9613" ID-DOI="http://doi.org/10.5281/zenodo.13124007" ID-Zenodo-Dep="13124007" ID-arpha="AEA57F17-2262-5230-AFA0-B36F70AD9613" ID-zoobank="https://zoobank.org/E9668141-118E-4C08-A504-A65A97E08615" LSID="urn:lsid:zoobank.org:act:E9668141-118E-4C08-A504-A65A97E08615" LSID-ZBK="urn:lsid:zoobank.org:act:E9668141-118E-4C08-A504-A65A97E08615" httpUri="http://treatment.plazi.org/id/AEA57F1722625230AFA0B36F70AD9613">
<subSubSection id="5076EBB397B46D3C4B4C2FBC44F45A22" type="nomenclature">
<paragraph id="6FF137697B322F7C551C44DB5286C561">
<taxonomicName id="5325813F39FA2F29E7894C0549FBF51C" authority="Namayandeh &amp; Hudson" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="palmerparcum" status="sp. nov.">
<emphasis id="AF7A0BE2A90EF074961486D22570EEDE" italics="true">Bryophaenocladius palmerparcum</emphasis>
Namayandeh &amp; Hudson
</taxonomicName>
<taxonomicNameLabel id="059442A962D462FCCEFE112F74BEB00E" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="EBBF0C7BECCD7FFAECC8068B2AB2018D" type="description">
<paragraph id="6F15FCACE386505B044EAABBEB07F8EF">
<figureCitation id="F0718D88986BAAC95DF1F0CEC7C7B3DE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bryophaenocladius palmerparcum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure7" httpUri="https://binary.pensoft.net/fig/1103618">Fig. 7 A – D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EU3AG" type="type material">
<paragraph id="43715E34E023A098A42DEA4AAC609F10">
<heading id="B8B7BE5FDC3C7D33373D394630BCC6D2" reason="title">Type material.</heading>
</paragraph>
<paragraph id="85A05A4A291827FCFC413495ED50DE09">
<materialsCitation id="A4684E931A620BC85EFB56E5ADDAB84F" collectingDate="2022-06-30" collectionCode="ARC" collectorName="Pond A &amp; P. L. Hudson" country="United States of America" latitude="42.42766" location="Palmer Park" longLatPrecision="1" longitude="-83.11741" municipality="Detroit" specimenCount="1" specimenCount-male="1" stateProvince="Michigan" typeStatus="holotype">
<emphasis id="B842CBEC7852DBAC1AA45DFD7E1CF3A3" bold="true" italics="true">
<typeStatus id="AFF0E40F91546DD7676C248CF530B586" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="A7BC5AE296F918ABA23A7295485C4E1A" count="1" type="male">1 male</specimenCount>
;
<collectingCountry id="B9DA85CD9F05A4DC4D73473FBC86745F" name="United States of America">USA</collectingCountry>
,
<collectingRegion id="518E0BA2BAF0D3173406D0BA3B886616" country="United States of America" name="Michigan">Michigan</collectingRegion>
,
<collectingMunicipality id="3AB48B527A88E71C2BAE092B3C9B428D">Detroit</collectingMunicipality>
,
<location id="C582BC1D065CF2DDB01EF6EE9261DA07" LSID="urn:lsid:plazi:treatment:AEA57F1722625230AFA0B36F70AD9613:C582BC1D065CF2DDB01EF6EE9261DA07" country="United States of America" latitude="42.42766" longLatPrecision="1" longitude="-83.11741" municipality="Detroit" name="Palmer Park" stateProvince="Michigan">Palmer Park</location>
,
<collectorName id="0C084684E740F043F3B65710C1DA3EEF">Pond A</collectorName>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EC4AG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.117410,42.427660]}">
<geoCoordinate id="2F2C525318C3F76B6376214042E48B21" degrees="42.42766" direction="north" orientation="latitude" precision="1" value="42.42766">42.42766 ° N</geoCoordinate>
,
<geoCoordinate id="CD0EC98E7F35D3869D958BD72726DCF1" degrees="83.11741" direction="west" orientation="longitude" precision="1" value="-83.11741">83.11741 ° W</geoCoordinate>
</named-content>
; leg.
<collectorName id="FF9409DA877F8BD718AC7436FE4903D2">P. L. Hudson</collectorName>
;
<date id="2446E3AF70BFF8D01F506822FAD5FAE2" value="2022-06-30">
<collectingDate id="B9E611FCD64BAC7C9716E8606B455183" value="2022-06-30">30. vi. 2022</collectingDate>
</date>
, dep.
<abbrev id="ABBRID0EH4AG" xlink_title="Albert J. Cook Arthropod Research Collection">
<collectionCode id="F4740A6FD2FDEDBBDE9D941736DBFA95" country="Canada" httpUri="http://grbio.org/cool/r82x-h8re" name="Atlantic Reference Centre" type="">ARC</collectionCode>
</abbrev>
</materialsCitation>
.
<emphasis id="7BE5A541E1F9050C6DD65EBF0F91AC2C" bold="true" italics="true">
<typeStatus id="0D6E8EC03AF83DB1CDCA366146AD7BBA" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="77973C065AF9255E77204900DE32736E" count="2" type="male">2 males</specimenCount>
; same as
<typeStatus id="EB76F6133E546D2514B065476CEEEC76" type="holotype">holotype</typeStatus>
except leg. A. Namayandeh;
<date id="3149BD88816C3D4211620951844500D8" value="2023-06-28">28. vi. 2023</date>
, dep.
<abbrev id="ABBRID0EO4AG" xlink_title="Albert J. Cook Arthropod Research Collection">
<collectionCode id="E5AEDFEE1DE40BFA7809300A599B2800" country="Canada" httpUri="http://grbio.org/cool/r82x-h8re" name="Atlantic Reference Centre" type="">ARC</collectionCode>
</abbrev>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ES4AG" type="description">
<paragraph id="0D1D77246EDD3D20AE8E129173A6F2BF">
<heading id="5A7B041DABEECBBBDF0BCC8E9D29869D" reason="title">Diagnostic characters.</heading>
</paragraph>
<paragraph id="ACCBDE1D6F6062AD3F717A60520AA93F">
The adult male of
<taxonomicName id="96A1C3617050E1BE7EEE8A0D42688B42" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="palmerparcum" status="sp. nov.">
<emphasis id="D852B1CF55C65C789C350F6FE50DFBA6" italics="true">B. palmerparcum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="16C496E563276190E18EC9FDA2879072" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from other
<taxonomicName id="EEC1EE378EC402E6EE86F146C618534F" authority="Thienemann, 1934" authorityName="Thienemann" authorityYear="1934" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="25CB89E64D3144F053ACFB9C683F60D6" italics="true">Bryophaenocladius</emphasis>
Thienemann, 1934
</taxonomicName>
males by the combination of the following characters: AR 1.4–1.5; costa not well-extended; anal point short, wide, and triangular, not surpassing the apex of tergite IX, with 11–15 setae; virga inconspicuous, consists of two long spines, looped; sternapodeme straight without large oral projections; superior volsella collar shaped; inferior volsella absent; gonostylus straight with a mid-section wide, and short collar-shaped crista
<taxonomicName id="70D4954D2E723BD2AB51EEE6EA6896DC" authorityName="Meigen" authorityYear="1818" class="Insecta" family="Chironomidae" genus="Chironomus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="1F68E4010A2C251679AA5F0205A5FE59" italics="true">dorsalis</emphasis>
</taxonomicName>
; HR 1.5–1.9; HV 2.6–2.9.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ET5AG" type="description">
<paragraph id="D1B2AD1C8243DE7F3A8FC52D64C94C62">
<heading id="3386BDA8E5294BB0F3CF03D7B5FDA00B" reason="title">Description.</heading>
</paragraph>
<paragraph id="397F73DFBA134FD117089D788BD41C24">
<emphasis id="B58D64D8CC878A2171DDD65A104C01EB" bold="true">Male</emphasis>
(
<emphasis id="CB740DDA6D1E8ED064357ED50B4963C5" italics="true">n</emphasis>
= 3; unless otherwise stated). Total length
<quantity id="29DB1E529347E4A35C4A2FD931C26B7C" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.4" metricValueMin="2.7" unit="mm" value="3.05" valueMax="3.4" valueMin="2.7">2.7–3.4 mm</quantity>
, Wing
<quantity id="25653A5E66A0BB46D8D0BC3BD657C320" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.4–1.7 mm</quantity>
long and
<quantity id="80B75E839A66B6605FC05BAFCA3E8CFB" metricMagnitude="-4" metricUnit="m" metricValue="4.0" unit="mm" value="0.4">0.4 mm</quantity>
wide.
</paragraph>
<paragraph id="E8801BF7C8663BE8DE9AFF71D7459C4E">
<emphasis id="DD351AF8F89F9E0B2F650F1B4475CD7B" bold="true">
<emphasis id="AD6AB8687755A3D5DF1D04C60B0B659A" bold="true" italics="true">Coloration</emphasis>
.
</emphasis>
Head, abdomen, and halter dark brown. Thorax dark brown with much paler yellowish areas in portion of dorsocentrals, in anteprontal, anepisternum, and apical portion of preepisternum regions. Legs golden brown. Wing greyish brown.
</paragraph>
<paragraph id="9D6DF20ED621E64D499111702C5E0893">
<emphasis id="AEF872E35438DAE6FB5FD35F4B536242" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="AF76C3A509F1FBAD36B58028A952C9DB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bryophaenocladius palmerparcum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure7" httpUri="https://binary.pensoft.net/fig/1103618">7 A, B</figureCitation>
). Antenna with 13 flagellomere, last flagellomere with 10 sensilla chaetica, each of 2
<superScript id="C78E4212460B2324AF31DB425CC8EE16">nd</superScript>
and 3
<superScript id="178B77233B65855470F53B46D8C90370">rd</superScript>
flagellomere with pair of sensilla chaetica; shaft starts at base of 4
<superScript id="09021FEFC2D2359E6B2A807B773DD291">th</superScript>
flagellomere (Fig.
<figureCitation id="A12618ACC6572B6DE0B7C5EFAE3A59AF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bryophaenocladius palmerparcum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure7" httpUri="https://binary.pensoft.net/fig/1103618">7 A</figureCitation>
); AR 1.4–1.5. Temporal setae 9, uniserial. Tentorium 150–190 µm long, tentorial pit close to apex. Clypeus squared, 101 µm long and 125 µm wide, with 20 setae, setae 73 µm long (
<emphasis id="109AC4AF3AB9009144695F10D660E622" italics="true">n</emphasis>
= 1). Palpal segments lengths (in μm): 55–77, 66; 33–57, 45; 157–164, 161; 125–131, 128; 93, third palpomere with 3 sensilla clavata, and without any projection.
</paragraph>
<caption id="6DF037DDA5AC6AF5A78EDE198217E285" ID-DOI="10.3897/zookeys.1208.124495.figure7" ID-arpha="1186822A-2A3D-530F-98E7-94C457B15F8F" httpUri="https://binary.pensoft.net/fig/1103618" startId="F7">
<paragraph id="86AE216DDCAC716E034F4880C304770C">
<label id="D27357080B620D3863BA167CD3463442">Figure 7.</label>
</paragraph>
<paragraph id="AF3BCC1DA9988E590E4A41F9D30EA8B6">
<taxonomicName id="10A4B96A76B38C0A93A2E57D5ECE4E8A" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="palmerparcum" status="sp. nov.">
<emphasis id="FB8D0DF7662203E561FD50E21EDDE14F" italics="true">Bryophaenocladius palmerparcum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFEDD89CE7374216E7607297D3E447F0" rank="species">sp. nov.</taxonomicNameLabel>
, adult male
<emphasis id="10FA3C69107AE112E8818D90CE586C75" bold="true">A</emphasis>
antenna
<emphasis id="B2CF29E1FCCC54704181D55CCF8EFB72" bold="true">B</emphasis>
head
<emphasis id="4E0178C8426293A4260560F01B53FD9C" bold="true">C</emphasis>
thorax
<emphasis id="AB822D4F56DF353A482B83B6E09AF5C1" bold="true">D</emphasis>
wing
<emphasis id="11087A107F5593B37944B9B50453877C" bold="true">E</emphasis>
hypopygium. Scale bars: 100 µm.
</paragraph>
</caption>
<paragraph id="6304224C06457C16D81898317F2A1D69">
<emphasis id="B4CE51BD0182E5C2A4D9347448D0104B" bold="true" italics="true">Thorax</emphasis>
(Fig.
<figureCitation id="C6B522551388AE0EE91A619E88C94B8A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bryophaenocladius palmerparcum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure7" httpUri="https://binary.pensoft.net/fig/1103618">7 C</figureCitation>
): Achrostichals 8–10, starting close to antepronotum; dorsocentrals 12–23,
<quantity id="6BC4596FCB902EFCAD910C8F5A3C8190" metricMagnitude="-1" metricUnit="m" metricValue="4.572" unit="in" value="18.0">18 in</quantity>
two rows; prealars 4–6, 5; scutellars 12; antepronotals 3 basoventrally.
</paragraph>
<paragraph id="3F887CA47DDEFFF9F691697C980EB5FF">
<emphasis id="E924895F7CAC6E731D8655D70B26887A" bold="true" italics="true">Wing</emphasis>
(Fig.
<figureCitation id="D40C6CFDEDA7F07D9596F59FF1694464" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bryophaenocladius palmerparcum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure7" httpUri="https://binary.pensoft.net/fig/1103618">7 D</figureCitation>
): Brachiolum with 2 setae;
<collectionCode id="0BFE47E7E5A18C427F5ECAB8B2C08202" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
with 7–13, 10 setae;
<collectionCode id="028F1A6E48A00472D00125CE8D31BB3A" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="90BB9D004A5AB03FAFFFF6D407E4956F">1</subScript>
with 4 setae;
<collectionCode id="3521C5FBAE7C5C310211A82EAEB25086" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
<subScript id="626B0F358388D38079A10A0A5405D96E">4 + 5</subScript>
with 4 setae; other veins bare. Squama with 7–8 setae. Anal lobe produced and squared. Costa not well-extended, 17–20, 18 µm long. Microtrichia visible at 10 × magnification.
</paragraph>
<paragraph id="E70A5A442DA438ECAA0F2CF10C36FF69">
<emphasis id="1E9820E427B162EBEFB1833299522C44" bold="true">
<emphasis id="8C4E5D9B9C6D8D25EECD2FCF76F5FF2D" bold="true" italics="true">Legs</emphasis>
.
</emphasis>
Foreleg spur 58 μm long (
<emphasis id="AE6986A7B6F5305325F7C45CE06D7C08" italics="true">n</emphasis>
= 1), midleg spur 30 μm long (
<emphasis id="A590817A2125850245DAAA42A34EC190" italics="true">n</emphasis>
= 1), other spur damaged; hindleg spurs damaged, hind tarsus missing, hindleg comb with 12 spines (
<emphasis id="3465ADF5ECA1EBE8292EE2F83E232E2A" italics="true">n</emphasis>
= 1). Lengths and proportions of legs as in Table
<tableCitation id="9533ABE25A3D4544BB4280A00E7DAF54" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Male leg lengths (μm) and proportions of Bryophaenocladius palmerparcum sp. nov." httpUri="http://table.plazi.org/id/B04F8C284B4BFF207C98839762AE96CA" tableUuid="B04F8C284B4BFF207C98839762AE96CA">2</tableCitation>
.
</paragraph>
<caption id="B04F8C284B4BFF207C98839762AE96CA" ID-Table-UUID="B04F8C284B4BFF207C98839762AE96CA" httpUri="http://table.plazi.org/id/B04F8C284B4BFF207C98839762AE96CA" startId="T2">
<paragraph id="A6D455C689BB4853FC69DE7198596843">
<label id="637A399A204785E78E8BC314CD4D036D">Table 2.</label>
</paragraph>
<paragraph id="5B623C302CD1AEB4C95A8C2A7816A6AC">
Male leg lengths (μm) and proportions of
<taxonomicName id="DC02DD66BD8EC597CBCA3606A05C7D99" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="palmerparcum" status="sp. nov.">
<emphasis id="A25DB230840F63ABEEBDE3A96C7D2D2E" italics="true">Bryophaenocladius palmerparcum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="063D87CD610B8992CD8403037AE9F530" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="A4C1951FA02C77EB266EC527AC15E5BB">
<table id="TID0EIOBI" rules="all">
<tbody id="D8DC1FEAFF9E3B493D47D705B1E7B8CA">
<tr id="56D5AB9155109209AC23B268877BCBC4">
<th id="801B97DB25198A59DCFBFC4F08E910E4" colspan="1" isEmpty="true" rowspan="1">-</th>
<th id="4A0F393D42E33C3F6A6F2D53B2630DE0" colspan="1" rowspan="1">fe</th>
<th id="5C2C9F18E8E9FCE67595C4ECCC414529" colspan="1" rowspan="1">ti</th>
<th id="84A86F65F2F1E27E0EC32F25659EA1C4" colspan="1" rowspan="1">
ta
<subScript id="567845D220CA8BD8079D285677FA0AA9">1</subScript>
</th>
<th id="C32F344F1B19F97B7FA651923F8580B8" colspan="1" rowspan="1">
ta
<subScript id="53031AEC232C17DD3770815D40260F7C">2</subScript>
</th>
<th id="091424B6C536415FD37008ACED20D51A" colspan="1" rowspan="1">
ta
<subScript id="77B47B12FA9C230799A2152038A0AE2C">3</subScript>
</th>
<th id="6DCE97D56A6FD4E4A9303CC6332333CE" colspan="1" rowspan="1">
ta
<subScript id="5D922D4067E957219571BC5EA433CC00">4</subScript>
</th>
<th id="2EB0309B2B94A9C5E16FA2E4E216599F" colspan="1" rowspan="1">
ta
<subScript id="92290B81A9618DD31548BFA45B6794D8">5</subScript>
</th>
<th id="C5AC0FAFC0C44449F876461EEF56575E" colspan="1" rowspan="1">LR</th>
<th id="4DCEC15C5D41BF29C3DEE60AB0405EA4" colspan="1" rowspan="1">BV</th>
<th id="6A889F538B8A3E8AD1E7BA0C2A1CCD74" colspan="1" rowspan="1">SV</th>
</tr>
<tr id="ED912E8157F21E3740482AEE766140B6">
<td id="79105D02477E8223CB74BA86509F7553" colspan="1" rowspan="1">
<emphasis id="35BEB84D11250CAFB1C1EADC6D84E208" bold="true">
P
<subScript id="AA7129BC7C52E7F4D97FF740B0DEE29E">1</subScript>
</emphasis>
</td>
<td id="DD434BC9948BDDD9843AD446C5E4FCFD" colspan="1" rowspan="1">627</td>
<td id="FD912E13B3F3153CF27199E01A32A5C5" colspan="1" rowspan="1">748</td>
<td id="FD984336ACD71C29DF146C30CE23B4DA" colspan="1" rowspan="1">729</td>
<td id="F8C8D295B4B5DFF2F82AC6AC6CDDAEA9" colspan="1" rowspan="1">252</td>
<td id="B19B96F4DD043419E0C2F2AD2F567C93" colspan="1" rowspan="1">169</td>
<td id="5D1D90EDB066B7D487585CCA66D89C05" colspan="1" rowspan="1">116</td>
<td id="8B281424CD285FB465F6E413EE0CD05C" colspan="1" rowspan="1">81</td>
<td id="1A98173F07EF1DEC5B92A79951350532" colspan="1" rowspan="1">0.97</td>
<td id="1144807C4C545FA316793B0A6ECD00D3" colspan="1" rowspan="1">3.4</td>
<td id="D24F023496F1EF53B8B730E3C6216DBB" colspan="1" rowspan="1">1.9</td>
</tr>
<tr id="E30C1B1892FAEA94A9A641BAC5A66B75">
<td id="9376CF04D78F601106CA09EAC70D0E8F" colspan="1" rowspan="1">
<emphasis id="E1685511429DFFE122C2D87CB7BFC0D8" bold="true">
P
<subScript id="7D6FCBCE37DCAC867E207FF55F11E7A6">2</subScript>
</emphasis>
</td>
<td id="3DE39495E6BD4AC0176FA1B6948D729C" colspan="1" rowspan="1">718</td>
<td id="E51C743F643771075BDDF55A60E21487" colspan="1" rowspan="1">805</td>
<td id="9E5CB3163D4944D84E850A0FA97278B7" colspan="1" rowspan="1">391</td>
<td id="B4AA257520AC97365213BA72460F60D8" colspan="1" rowspan="1">214</td>
<td id="7FEB577B52FF551D95FDEA4B6CA5C81B" colspan="1" rowspan="1">142</td>
<td id="8159C80C1B7EBCE250B4DC2C8613641C" colspan="1" rowspan="1">104</td>
<td id="CC5E4666D302A3E6B2BE4BCE17048A51" colspan="1" rowspan="1">73</td>
<td id="85F6FC42CBA302AB817E0AC7CF935E08" colspan="1" rowspan="1">0.50</td>
<td id="24EE05AEA8351033A4009CACDC8989DF" colspan="1" rowspan="1">–</td>
<td id="0016442EEF00EBCBD326D59BC8487AED" colspan="1" rowspan="1">0.50</td>
</tr>
<tr id="D266F57DE6428C1A13889E9A491DA775">
<td id="E1B3075C702FCFDF9A7E1D0173A88CC2" colspan="1" rowspan="1">
<emphasis id="72848A8F0111970EDA3609E9C9DA6A46" bold="true">
P
<subScript id="A66F3E7A9860D7C3D0D5C66EAF12B9AB">3</subScript>
</emphasis>
</td>
<td id="88C21EE99F304FB4F795C5F99872230A" colspan="1" rowspan="1">828</td>
<td id="5A625A630BB59864B57023DF089EAFAC" colspan="1" rowspan="1">990</td>
<td id="9E566A38ECBE35D1B525989F8A3B8CCF" colspan="1" rowspan="1">–</td>
<td id="0B59F21AA271E550E1741DABD3B6D2EC" colspan="1" rowspan="1">–</td>
<td id="6A36213D7B234E941D373B31C01E6851" colspan="1" rowspan="1">–</td>
<td id="15399D73BE09A094E2BE87BA09D46D07" colspan="1" rowspan="1">–</td>
<td id="2C8F15950D04FBB0BED96A3CF840F71A" colspan="1" rowspan="1">–</td>
<td id="B57BADFEC5B2C626CD987E27D2B63E20" colspan="1" rowspan="1">–</td>
<td id="7FAF7B13B8F73E9E54CC464A209EBEC0" colspan="1" rowspan="1">–</td>
<td id="E0133D5DCBD814BF6C8FDF11262F2A0D" colspan="1" rowspan="1">–</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="01A6FAD3A6DB96C5B626C8B97292D05C">
<emphasis id="4965541DB857D272D24C6AD5BA4B302C" bold="true" italics="true">Hypopygium</emphasis>
(Fig.
<figureCitation id="3D801029FAA64C33A5389F30256C2667" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Bryophaenocladius palmerparcum sp. nov., adult male A antenna B head C thorax D wing E hypopygium. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1208.124495.figure7" httpUri="https://binary.pensoft.net/fig/1103618">7 E</figureCitation>
): Anal point short, wide, and triangular, not surpassing the apex of tergite IX, 18 μm long and 34 μm wide at the base (
<emphasis id="84091DA50720FC640344F9640E2C42DB" italics="true">n</emphasis>
= 1), surface with 11–15, 12 setae. Virga inconspicuous, consists of two long spines, looped, 58 μm long (
<emphasis id="7A0482AF0AC7823470B7F1F93AC1D578" italics="true">n</emphasis>
= 1). Sternapodeme straight with large oral projections, 84–104, 97 μm long. Phallapodeme 42–50, 45 μm long. Superior volsella collar shaped, inferior volsella absent. Gonocoxite 158–216, 187 μm long. Gonostylus straight with a mid-section wide and short collar-shaped crista
<taxonomicName id="81D5AA1EB6B8835862163AEA4184B642" authorityName="Meigen" authorityYear="1818" class="Insecta" family="Chironomidae" genus="Chironomus" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="6BF1F5176641A524FE58BA0F51A13F8D" italics="true">dorsalis</emphasis>
</taxonomicName>
, gonostylus 102–114, 110 μm long, megaseta 9 μm long. HR 1.5–1.9, 1.7; HV 2.6–2.9, 2.8.
</paragraph>
<paragraph id="FE3BB6D0CFBADFF305E2E207C07CA62A">
<emphasis id="D0683A0142A0377A4DF6E1A5892755B5" bold="true">Female and immatures.</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4IBG" type="etymology">
<paragraph id="0CD01AD38A7EAB9EACCD7D400384EE9E">
<heading id="1D34F6D5DAC4DA45D5BCB04AA243E5EB" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="B41BA77696776553B30FF75552DFB33F">
The species is named after the locality where it is found, Palmer Park. The word
<emphasis id="E913829403F0D3AFFE1B38E1E9E8E9F6" italics="true">parcum</emphasis>
is Latin, meaning park.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFJBG" type="distribution">
<paragraph id="89CD24FB549E8B7E72E29AABD535DCF7">
<heading id="074736AB79A6346442C8997C7A5274CD" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="708B4888299C1A687CC797399BADE1EC">
<collectingCountry id="C77C09F55578099688C4C8C14F0CF040" name="United States of America">USA</collectingCountry>
(
<collectingRegion id="9B5541DF7398BFD20FD8023070C240E7" country="United States of America" name="Michigan">Michigan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKJBG" type="remarks">
<paragraph id="25E6DA036F9C6BA4F2923DB9915A9449">
<heading id="1D52CD4C416858989C3941F15EA009AC" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="A3ED8E1087A012C1F4685C5BEFA5351C">
A combination of strong decumbent achrostichals close to antepronotum; bare wing with strong punctation of microtrichia, and weak lateral spines attached to the shaft of hind and mid legs spines places this species in
<taxonomicName id="990EA05BF6D8F3C426D20BBA6DB59A79" authorityName="Thienemann" authorityYear="1934" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="4E310DEA851DA9D051DB4E3E40F57A32" italics="true">Bryophaenocladius</emphasis>
</taxonomicName>
. Although, in general, the long, prominent, and hyaline anal point defines many known species of
<taxonomicName id="A8A5C380B65AB6387A252241914B9FEC" authorityName="Thienemann" authorityYear="1934" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="101338806F3848B196955EE5D0B260D9" italics="true">Bryophaenocladius</emphasis>
</taxonomicName>
, the character of the short anal point of this species is not uncommon, and it has been observed among other known species of this genus. Previously,
<bibRefCitation id="5B337A980627383A19DE578C82525963" DOI="10.11646/zootaxa.5448.2.8" author="Donato M &amp; Mauad M &amp; Siri A" etAl="et al." firstAuthor="Donato" issue="2" journalOrPublisher="Zootaxa" pagination="283-291" refId="B16" refString="Donato M, Mauad M, Siri A (2024) New species of Bryophaenocladius Thienemann, 1934 (Diptera: Chironomidae) from Argentina. Zootaxa 5448 (2): 283–291. https://doi.org/10.11646/zootaxa.5448.2.8" title="New species of Bryophaenocladius Thienemann, 1934 (Diptera: Chironomidae) from Argentina." volume="5448" year="2024">Donato et al. (2024)</bibRefCitation>
have demonstrated that the species of
<taxonomicName id="17287903D04B4C0A37C96787CA444F0D" authorityName="Thienemann" authorityYear="1934" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="genus">
<emphasis id="0C9CC56616EDA663960706763364412A" italics="true">Bryophaenocladius</emphasis>
</taxonomicName>
show a significant pattern of anal point size and shape differences. For instance, among those species with short anal points,
<taxonomicName id="8AF46E4C28FCD4D5957559FF35E717C8" authority="Wang, Liu &amp; Epler, 2012" authorityName="Wang, Liu &amp; Epler" authorityYear="2012" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="scleras">
<emphasis id="6E23B674EDAB3321A70988984F41F048" italics="true">Bryophaenocladius scleras</emphasis>
Wang, Liu &amp;
<bibRefCitation id="E07B5F208F495F7D4E3AEA16CE231FBD" DOI="10.11646/zootaxa.3355.1.3" author="Epler JH" issue="1" journalOrPublisher="Zootaxa" pagination="51-61" refId="B17" refString="Epler JH (2012) A brachypterous Bryophaenocladius (Diptera: Chironomidae: Orthocladiinae) with hypopygium inversum from Heggie's Rock, Georgia, USA. Zootaxa 3355 (1): 51 - 61. https: // doi. org / 10.11646 / zootaxa. 3355.1. 3" title="A brachypterous Bryophaenocladius (Diptera: Chironomidae: Orthocladiinae) with hypopygium inversum from Heggie’s Rock, Georgia, USA." volume="3355" year="2012">Epler, 2012</bibRefCitation>
</taxonomicName>
from the Nearctic has a short semicircular anal point (see
<bibRefCitation id="4F6D797B4335E006698187D38D916A31" DOI="10.11646/zootaxa.581.1.1" author="Wang" etAl="et al." firstAuthor="Wang" issue="1" journalOrPublisher="Zootaxa" pagination="1-10" refId="B55" refString="Wang XH, Liu Z, Epler JH (2004) New species of Bryophaenocladius Thienemann from the Nearctic Region (Diptera: Chironomidae: Orthocladiinae). Zootaxa 581 (1): 1–10. https://doi.org/10.11646/zootaxa.581.1.1" title="New species of Bryophaenocladius Thienemann from the Nearctic Region (Diptera: Chironomidae: Orthocladiinae)." volume="581" year="2004">Wang et al. 2004</bibRefCitation>
), and
<taxonomicName id="4F0F32CB3865194CD07494453AE77BC5" authority="(Malloch, 1915)" baseAuthorityName="Malloch" baseAuthorityYear="1915" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pleuralis">
<emphasis id="3F24BD4E6BF489A7295C1BB1798B1ABF" italics="true">Bryophaenocladius pleuralis</emphasis>
(Malloch, 1915)
</taxonomicName>
has a dark short anal point that does not extend beyond tergite IX (
<bibRefCitation id="1A265C40DD55865D1046418D2848645E" author="Makarchenko and Makarchenko" firstAuthor="Makarchenko" journalOrPublisher="Eurasian Entomological Journal" pagination="51-63" refId="B32" refString="Makarchenko EA, Makarchenko MA (2009) New records of chironomids (Diptera, Chironomidae, Orthocladiinae) in Far East and bordering territories. VII. Bryophaenocladius Thienemann. Eurasian Entomological Journal 8: 51–63." title="New records of chironomids (Diptera, Chironomidae, Orthocladiinae) in Far East and bordering territories. VII. Bryophaenocladius Thienemann." volume="8" year="2009">Makarchenko and Makarchenko 2009</bibRefCitation>
). Additionally,
<bibRefCitation id="AEF6E08425F41108342BFFA853BFE31C" DOI="10.11646/zootaxa.3355.1.3" author="Epler JH" firstAuthor="Epler" issue="1" journalOrPublisher="Zootaxa" pagination="51-61" refId="B17" refString="Epler JH (2012) A brachypterous Bryophaenocladius (Diptera: Chironomidae: Orthocladiinae) with hypopygium inversum from Heggie’s Rock, Georgia, USA. Zootaxa 3355 (1): 51–61. https://doi.org/10.11646/zootaxa.3355.1.3" title="A brachypterous Bryophaenocladius (Diptera: Chironomidae: Orthocladiinae) with hypopygium inversum from Heggie’s Rock, Georgia, USA." volume="3355" year="2012">Epler (2012)</bibRefCitation>
described brachypterous
<taxonomicName id="07672C9EB838A873910D42F5EF54115D" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="chrissichuckorum">
<emphasis id="FC0EC717CA62E3473E6EBB9E88E3B87E" italics="true">Bryophaenocladius chrissichuckorum</emphasis>
</taxonomicName>
with a wide and short, triangular anal point similar to that of
<taxonomicName id="A6E9AD9A6AC5F5CD208B2F52C67A879A" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="palmerparcum" status="sp. nov.">
<emphasis id="BEFD1379F13EB0939FC5A457CA83A3A6" italics="true">B. palmerparcum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1F4053EB90FF979C5E6D18B6B55240F2" rank="species">sp. nov.</taxonomicNameLabel>
The first author has also observed this character of short anal point in Neotropical species (AN pers. obs. of specimens from
<collectingCountry id="FC9A018191F72D38BF5133653C4D9EC8" name="Costa Rica">Costa Rica</collectingCountry>
). What is also observable is that besides the variation in shape and size of the anal point, as discussed in
<bibRefCitation id="F6BB3ACFA7C4FDF46D4C034C599CF693" DOI="10.11646/zootaxa.5448.2.8" author="Donato M &amp; Mauad M &amp; Siri A" etAl="et al." firstAuthor="Donato" issue="2" journalOrPublisher="Zootaxa" pagination="283-291" refId="B16" refString="Donato M, Mauad M, Siri A (2024) New species of Bryophaenocladius Thienemann, 1934 (Diptera: Chironomidae) from Argentina. Zootaxa 5448 (2): 283–291. https://doi.org/10.11646/zootaxa.5448.2.8" title="New species of Bryophaenocladius Thienemann, 1934 (Diptera: Chironomidae) from Argentina." volume="5448" year="2024">Donato et al. (2024)</bibRefCitation>
, those species with the short anal point can still possess the hyaline section, such as
<taxonomicName id="2F7ADC7EDAC80CBD118D1095EB3C2ED5" authority="Brundin, 1947" authorityName="Brundin" authorityYear="1947" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="inconstans">
<emphasis id="A9DE2A7FB16A74E27DD8497722150E2F" italics="true">Bryophaenocladius inconstans</emphasis>
Brundin, 1947
</taxonomicName>
and some lacking, such as the case of
<taxonomicName id="5955E69CE7C3873EA994E82EE0F0B8FE" authorityName="Namayandeh &amp; Guerra &amp; Islam &amp; James &amp; Hudson &amp; Ghaderi &amp; Yusuf &amp; Vasquez &amp; Ram" authorityYear="2024" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="palmerparcum">
<emphasis id="1C9554C3393187B65C3B3D4023CEF0AF" italics="true">B. palmerparcum</emphasis>
</taxonomicName>
,
<taxonomicName id="FF599240131C101A0998BE8DF20E15A6" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="chrissichuckorum">
<emphasis id="6BBB4801FDF746FB95321EF11F83400C" italics="true">B. chrissichuckorum</emphasis>
</taxonomicName>
and
<taxonomicName id="74971A57517825CE643E18DF1C3D2D9A" baseAuthorityName="Malloch" baseAuthorityYear="1915" class="Insecta" family="Chironomidae" genus="Bryophaenocladius" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pleuralis">
<emphasis id="1F6BAEA7AB7D77427255234A91B3593A" italics="true">B. pleuralis</emphasis>
</taxonomicName>
. Therefore, the presence of an anterior hyaline section of the anal point can also separate species with a reduced anal point.
</paragraph>
</subSubSection>
</treatment>
</document>

354
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<mods:title id="2C71689734E0D1EE7DD5B73CB8E68E2A">An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes</mods:title>
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<mods:name id="2D34594913ED48ED0A8D3EC96ADA0DEF" type="personal">
<mods:role id="9AC7442F09847B5B8C9BFDB80EEA0E7A">
<mods:roleTerm id="065C431521E84EB89BC017AA78C81DE2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2791977D075F2E44C1E9A60CD9959BF8">Strümpher, Werner P.</mods:namePart>
<mods:nameIdentifier id="371ADA43D9D25D4203EFE6F33D8EED3B" type="ORCID">0000-0002-7047-2666</mods:nameIdentifier>
<mods:affiliation id="820B742F37BB39DFEF2763E770310BE8">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa</mods:affiliation>
</mods:name>
<mods:name id="ABBAE540C9A639350DE78EA5CCC3729F" type="personal">
<mods:role id="940B310F576631CC03FBE6ED127593D2">
<mods:roleTerm id="003569F87070FF5E003653A239AA9A45">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="14561FF6BADE67DBE48898C7B626A03F">Barclay, Maxwell V. L.</mods:namePart>
<mods:nameIdentifier id="64686B510F256EE48261BB24881FE447" type="ORCID">0000-0003-4989-2014</mods:nameIdentifier>
<mods:affiliation id="E360669BA077F22F8D3EB197FC7CC389">Department of Life Sciences, Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name id="3F972E9194577EA17199FB6A02B76B39" type="personal">
<mods:role id="4BC5E9D795E4CAFE12B592218D31438A">
<mods:roleTerm id="EC26EDBA23D12A858DF0349F86F51021">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F91D058922396DC868938A58828A311">Vaz-de-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="24A31BE0FB15A60F291CE6ED98AEC8BD" type="ORCID">0000-0001-9697-320X</mods:nameIdentifier>
<mods:affiliation id="EB4161D2A4CEDEA10A36E576FEC4C43C">Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="FB0CA069B3C060DC3FB75AC74D8D4D1C">text</mods:typeOfResource>
<mods:relatedItem id="B793479A5BD4C9E498FF4434A5DC8096" type="host">
<mods:titleInfo id="735DAE4C24D18A47A603F06961B9A190">
<mods:title id="131273BD408BA70051D3AECE3159D410">Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="1A601C3763D3012331348B8A06A339EC">
<mods:date id="5C6733830D34CDBB4A3D5F2A0A6BF7D4">2024</mods:date>
<mods:detail id="31BF62B9A6EFDC80063F90413CF5B31F" type="pubDate">
<mods:number id="8671326D7BD7A974334DA483517709DE">2024-07-29</mods:number>
</mods:detail>
<mods:detail id="4A63A54E08DCFA8C772B2621CAA8601F" type="volume">
<mods:number id="CF2272A7CF8AF7EF599071825C931E75">74</mods:number>
</mods:detail>
<mods:detail id="66086B726FB6428117E6B6871E372723" type="issue">
<mods:number id="22197FE4791C89AEC907D80423B5D875">1</mods:number>
</mods:detail>
<mods:extent id="F925EE9617BC86B2F39EDB1BB0036C3B" unit="page">
<mods:start id="A00861BC782010E29E2347964B80E568">81</mods:start>
<mods:end id="660D221682EC339D0BA13186B2CC0636">101</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="03D91275595BCBF0CCC1A3A7BFB8FA79">journal article</mods:classification>
<mods:identifier id="CA58DA1CCE87391D88BCC96124567ADA" type="DOI">10.3897/contrib.entomol.74.e126799</mods:identifier>
<mods:identifier id="DE997F41F3A01885E63386690E532F39" type="ZooBank">339AB68C-DCB1-435F-B88E-2C42D0A7B05E</mods:identifier>
</mods:mods>
<treatment id="BAC76C222D505D2BA37175E418D5B1F8" ID-DOI="http://doi.org/10.5281/zenodo.13124000" ID-Zenodo-Dep="13124000" ID-arpha="BAC76C22-2D50-5D2B-A371-75E418D5B1F8" LSID="urn:lsid:plazi:treatment:BAC76C222D505D2BA37175E418D5B1F8" httpUri="http://treatment.plazi.org/id/BAC76C222D505D2BA37175E418D5B1F8">
<subSubSection id="C0049E43CAA3F105B5F51C591B29FB8D" type="nomenclature">
<paragraph id="6B3441C1CF95E5E07D5EFE790C7A1BD5">
<taxonomicName id="27B9F4DAFBAAB74A996E86EF724B19B2" ID-CoL="49DSM" authority="(Vaurie, 1962)" baseAuthorityName="Vaurie" baseAuthorityYear="1962" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spatulatus" subGenus="Omorgus">
<emphasis id="7684F3600F478024C2C00ABDFDC2C7F5" italics="true">Omorgus</emphasis>
(
<emphasis id="5124706BDB983BC5559F47DB03451C5B" italics="true">Omorgus</emphasis>
)
<emphasis id="B62870AAF1C657ED6F9C3B70B8D4C5F3" italics="true">spatulatus</emphasis>
(
<bibRefCitation id="B6A0E938CDE408A2DF9B50B2D498501F" author="Vaurie P" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105 - 167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie, 1962</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="4245A2FA19228768DFA85F026A796175" type="description">
<paragraph id="CAA9649CDFB5FA89D01DF025B8A5CD91">
<figureCitation id="267532DBE6CFB111068D098C1C1FD684" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Trox spatulatus Vaurie, 1962 (now Omorgus spatulatus). A. Habitus in dorsal view; B. Habitus in lateral view; C. Spiculum gastrale and aedeagus in dorsal view; D. Labels. Photos by Christoph Germann (NHMB). Length of specimen: 10 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure6" httpUri="https://binary.pensoft.net/fig/1103868">Fig. 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="083A87AC19557700034871ED1E64C87F" type="reference_group">
<paragraph id="EE8BA254F2CFAF185B0C2A4B5FC7323D">
<treatmentCitationGroup id="B9A022B9000D9630827AF65894841053">
<treatmentCitation id="D4E5ED7E0D1892397E3DF7903F4A001D" author="Vaurie P" page="151" year="1962">
<taxonomicName id="D4C647700648C285ADF4EB78BCCBA749" ID-CoL="59DQR" authority="Vaurie, 1962: 151" authorityName="Vaurie" authorityPageNumber="151" authorityYear="1962" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spatulatus">
<emphasis id="F9A067BE0616705D40B227DA43D5F3B8" italics="true">Trox spatulatus</emphasis>
<comment id="49A58739C6A2DA209C00F0F61CE7AD43">
<bibRefCitation id="8F4752DBD210364F83A3EEF504626A37" author="Vaurie P" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105 - 167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie, 1962: 151</bibRefCitation>
(original description).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="72D3F1285B26D57231255316B9DD13F1">
<treatmentCitationGroup id="8CE84675D29534BAA25DB3E6974530C3">
<treatmentCitation id="AC8BC1D625A895724E91CF451C318665">
<taxonomicName id="F3980DA4660A9EACF214ED6888C3D16C" ID-CoL="59DQR" authorityName="Vaurie" authorityYear="1962" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spatulatus" subGenus="Omorgus">
<emphasis id="E2B53AA07B1E8003DE0325A4CB8C80DF" italics="true">Trox</emphasis>
(
<emphasis id="6C544A546121B7936D0BB7EC72431D4E" italics="true">Omorgus</emphasis>
)
<emphasis id="62E8981792BD6572A5C2FC445E5786F2" italics="true">spatulatus</emphasis>
</taxonomicName>
<comment id="BA4DC7B1EA04134CB3E0AF0113724706">
:
<treatmentCitation id="066FD3A6FC0E84EEFC9B092779CB1553" author="Scholtz CH" page="13" year="1982">
<bibRefCitation id="92D368746DE1952581A151E6273731B1" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa" page="13" pagination="1-27" refId="B96" refString="Scholtz CH (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa 54: 1–27." title="Catalogue of world Trogidae (Coleoptera: Scarabaeoidea)." volume="54" year="1982">Scholtz 1982: 13</bibRefCitation>
</treatmentCitation>
(catalogue).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="935B8DE30E6A21CAE8A81C928F8399BB">
<treatmentCitationGroup id="1FE8A8FCC251058C58CE4386F5366496">
<treatmentCitation id="001CD436064D22C2FE36EBADAF232F92">
<taxonomicName id="BF7974E9EEF225B58EA7A359924CDC80" ID-CoL="49DSM" baseAuthorityName="Vaurie" baseAuthorityYear="1962" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spatulatus">
<emphasis id="80352361F062905D592DC7370B0F9C7C" italics="true">Omorgus spatulatus</emphasis>
</taxonomicName>
<comment id="52CEE850F52E46907740BFCB2D8B3364">
:
<treatmentCitation id="CC52D3D5A31029CD79375D43DE998184" author="Deloya C" page="122" year="2005">
<bibRefCitation id="699B2229416615EF00BFB664897528E2" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121–129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="EF102B612043EB0A3A9487EAED8A02FC">
<treatmentCitationGroup id="698AA8C62FB37D7EF876941C8315EC9E">
<treatmentCitation id="E76D8B2909D7404CF5336AFEFA9DE570">
<taxonomicName id="4518E0A63F769D6AB28F254E423DB2FA" ID-CoL="49DSM" baseAuthorityName="Vaurie" baseAuthorityYear="1962" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spatulatus" subGenus="Omorgus">
<emphasis id="522DB51E31CB4F21D3352E6E7A299A54" italics="true">Omorgus</emphasis>
(
<emphasis id="31068CC2ADF6189959D91278E80F4CB5" italics="true">Omorgus</emphasis>
)
<emphasis id="E133627804BE0921EB1BA672531BBD83" italics="true">spatulatus</emphasis>
</taxonomicName>
<comment id="206857EF081B96B07D4151F3ABDA554D">
:
<treatmentCitation id="8595929BA8B1635883795D9C9AF2FEDF" author="Scholtz CH" page="1404" year="1990">
<bibRefCitation id="21153BC41E9EA78CCC87C50087DBB026" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" page="1404" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990: 1404</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="46ED85678292839752B3137A6D83EC07" author="Zidek J" page="16" year="2013">
<bibRefCitation id="B5EF9FFA780370FA9117ED8FF181EF2C" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="16" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1–38." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 16</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="9C62443F38839C871DACBA6FEF84F7BB" author="Zidek J" page="108" year="2017">
<bibRefCitation id="A78D357A9CD15F4F4FB16A4979B5B191" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="108" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 108</bibRefCitation>
</treatmentCitation>
(checklist).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENBBI" type="Type specimen examined">
<paragraph id="87B413AA082D57B9C7AA46CB4231FAE7">
<heading id="4F3F8469F101AF74FFDDE45191185106" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="376891EA848A10A25D4961F2B4E028D7">
<materialsCitation id="EB569102F543C1B4A1B7FA7FCD05063C" collectionCode="NHMB" country="Argentina" county="Fourth" location="First" municipality="Third" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires" typeStatus="holotype">
<emphasis id="D4404BE84AAFC7E51918408F006B3082" bold="true" italics="true">
<typeStatus id="41E37A9805CBBF7EC1654E54772D3947" type="holotype">Holotype</typeStatus>
, by original designation
</emphasis>
(
<specimenCount id="DDF5A45D6FA5A25D492A61BE7C6C9F5D" count="1" type="male">♂</specimenCount>
<named-content id="24121B6849D2F481872631DD26CA5107" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-basel" xlink_title="Naturhistorisches Museum, Basel">
<emphasis id="E15B28522177F91BE1D5152CFCBF627D" bold="true">
<collectionCode id="1F4B756272F99A904476F723B31C5636" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" type="">NHMB</collectionCode>
</emphasis>
</named-content>
).
<location id="71D0FB3EE0F6129F811583E37EA722FE" LSID="urn:lsid:plazi:treatment:BAC76C222D505D2BA37175E418D5B1F8:71D0FB3EE0F6129F811583E37EA722FE" country="Argentina" county="Fourth" municipality="Third" name="First" stateProvince="Ciudad Autonoma de Buenos Aires">First</location>
label [white aged, typeset]: “ Provincia /
<collectingRegion id="D787E91A99D134F93E099098C5C32D39" country="Argentina" name="Ciudad Autonoma de Buenos Aires">Buenos Aires</collectingRegion>
”.
<location id="36465F738B0DF25343140333E726D167" LSID="urn:lsid:plazi:treatment:BAC76C222D505D2BA37175E418D5B1F8:36465F738B0DF25343140333E726D167" country="Argentina" county="Fourth" municipality="Third" name="Second" stateProvince="Ciudad Autonoma de Buenos Aires">Second</location>
label [white aged, unknown handwriting]: “ II [?] ”.
<collectingMunicipality id="0C6FA939FB08688F92BD40A3525D04EB">Third</collectingMunicipality>
label [white, typeset]: “ Museum Frey / Tutzing ”.
<collectingCounty id="EEB6DE563B237973974870A79A3FCEDE">Fourth</collectingCounty>
label [red, Patricia Vaurie’s handwriting]: “ TYPE /
<taxonomicName id="F5C22E66BA0D1CD099D8D8ABF3A23A84" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="26C1777AC013E49BC9967D7942C29851" italics="true">Trox</emphasis>
</taxonomicName>
/
<taxonomicName id="25A313C64FA618DD3B023D9D28389CDB" baseAuthorityName="Vaurie" baseAuthorityYear="1962" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spatulatus">
<emphasis id="9721E450AC25C9AA421F1F99645124BF" italics="true">spatulatus</emphasis>
</taxonomicName>
/ Vaurie ” (Fig.
<figureCitation id="8CCC7CF24AFE5FE8C8277CCED557A6F9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Holotype of Trox spatulatus Vaurie, 1962 (now Omorgus spatulatus). A. Habitus in dorsal view; B. Habitus in lateral view; C. Spiculum gastrale and aedeagus in dorsal view; D. Labels. Photos by Christoph Germann (NHMB). Length of specimen: 10 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure6" httpUri="https://binary.pensoft.net/fig/1103868">6 D</figureCitation>
).
<emphasis id="2A0B03F9646A6A426EE88F9BFE762034" bold="true">Type locality</emphasis>
: “ Provincia
<collectingRegion id="B4FA523C93B5544FDFB5337A971CFE3C" country="Argentina" name="Ciudad Autonoma de Buenos Aires">Buenos Aires</collectingRegion>
” [
<collectingCountry id="73CF9783C85270AA834B3AD853C8E259" name="Argentina">Argentina</collectingCountry>
].
</materialsCitation>
</paragraph>
<caption id="7C7C8881A92B70D8FA1325D9E4F47562" ID-DOI="10.3897/contrib.entomol.74.e126799.figure6" ID-arpha="AF797C0B-73C4-58B5-8838-205A3572B212" httpUri="https://binary.pensoft.net/fig/1103868" startId="F6">
<paragraph id="1E6CF5E7E116DB524AFA98E19EC1A276">
<label id="ABBC99FF73ED40DA747CFCC31250C7DA">Figure 6.</label>
</paragraph>
<paragraph id="1FFDDFA7DE47B7AE2BC15870B71ECB63">
Holotype of
<taxonomicName id="8FD23DCA537187844EF1C5951ACD695B" authority="Vaurie, 1962" authorityName="Vaurie" authorityYear="1962" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spatulatus">
<emphasis id="7878C8DE8207A7EFDDA48B0690A6BB2B" italics="true">Trox spatulatus</emphasis>
<bibRefCitation id="8D7B249803D4633A312C9611FC23A6B6" author="Vaurie P" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105 - 167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie, 1962</bibRefCitation>
</taxonomicName>
(now
<taxonomicName id="27BC0BB880341EACCC21A8654D5EDD77" baseAuthorityName="Vaurie" baseAuthorityYear="1962" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="spatulatus">
<emphasis id="2E7D1585F5D801FD5B022018239F186E" italics="true">Omorgus spatulatus</emphasis>
</taxonomicName>
).
<emphasis id="65C7885D3C8158E96DE043D091819EB4" bold="true">A.</emphasis>
Habitus in dorsal view;
<emphasis id="062C46774890E1D7F0133E884A564C89" bold="true">B.</emphasis>
Habitus in lateral view;
<emphasis id="F4B45D845DFF2C1DEF94DCD8A85F8F5D" bold="true">C.</emphasis>
Spiculum gastrale and aedeagus in dorsal view;
<emphasis id="8789C2940A882C6D35A9DAE7060768F8" bold="true">D.</emphasis>
Labels. Photos by Christoph Germann (
<named-content id="728AAB570C847BCADBC50552E3EACDF7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturhistorisches-museum-basel" xlink_title="Naturhistorisches Museum, Basel">NHMB</named-content>
). Length of specimen: 10 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EBEBI" type="distribution">
<paragraph id="F7F30E324E137237166023782994A341">
<heading id="A3D2D450F85BA3F7D4667D9828303156" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="31FE21DDD9B51ADF022075C09819489B">
<collectingCountry id="DF2F9B85D2D830C4BC608A143D30E57E" name="Argentina">Argentina</collectingCountry>
(
<bibRefCitation id="0C284E308606A03B6B54C63FDEC60D84" author="Vaurie" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105–167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962</bibRefCitation>
;
<bibRefCitation id="DBFD2840A8B7F23748B8F79E0073006D" DOI="10.1080/00222939000770841" author="Scholtz" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPEBI" type="type material">
<paragraph id="E47ACBCA6C831AD79C1F716C1F03E04E">
<heading id="4881134191EF7DE59FB4308497278F51" reason="title">Non-type examined material</heading>
</paragraph>
<paragraph id="258752ECF610E9F0EE3D447C53CC6AB9">
<emphasis id="BAB8D2DB9B5471F0BDA6C53561FDFB3F" bold="true">
(
<specimenCount id="B0845B9D34EA69161795C090C7F834FE" count="215" type="generic">215 specimens</specimenCount>
).
<materialsCitation id="4D7FFA5F87146138F6CF6A212A662BE2" country="Argentina" location="Buenos Aires" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingCountry id="AF07560E1AC2BBC8B858B87E89641B59" name="Argentina">ARGENTINA</collectingCountry>
–
<emphasis id="BF877056E17B60F89EC595B536C16949" bold="true">
<collectingRegion id="CA4A6694FFCC2A353BBCDDA6A73826DB" country="Argentina" name="Ciudad Autonoma de Buenos Aires">Buenos Aires</collectingRegion>
</emphasis>
</materialsCitation>
</emphasis>
• 1; La Plata; MLPA •
<materialsCitation id="3197756E385A13F8F14DC5EB7FD48DF7" collectingDate="1928-03-07" collectionCode="MACN" collectorName="Quequen Puerto &amp; M. D. Jurado" country="Argentina" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
12;
<collectorName id="8AA8374B010F6EBCE177EEC1EDDF69A4">Quéquen Puerto</collectorName>
;
<date id="AF1F4DA0F946BD5CF502E2BCD87D6FD1" value="1928-03-07">
<collectingDate id="7C49E2635312742275E8F49CDD29993F" value="1928-03-07">7 Mar. 1928</collectingDate>
</date>
;
<collectorName id="ED18970398E6370085358A4F8033892D">M. D. Jurado</collectorName>
leg.;
<named-content id="8A3F28C3784E04378221578CEBC8AE43" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’">
<collectionCode id="D820DFFA8802F9882C23CC1AEF93FDEE" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</named-content>
</materialsCitation>
•
<materialsCitation id="376D60BB3B7DD8AE5FF1D688950A2333" collectionCode="MACN" collectorName="Quequen Puerto &amp; G. Pellerano" country="Argentina" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
1;
<collectorName id="8D5DA5C314495FF6A9239F362C416689">Quéquen Puerto</collectorName>
;
<collectorName id="AEF07571855AA2BECFCA6612EB4573CA">G. Pellerano</collectorName>
leg.;
<named-content id="5C3CAE17D236CE4BAE45D905F01D84F7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’">
<collectionCode id="6FCBD8BF8C093569203AB6EBE58E5207" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</named-content>
</materialsCitation>
•
<materialsCitation id="3018C69844CFDBDE18832936DC346C0F" collectingDate="1937-02" collectionCode="CEMT" collectorName="Quequen" country="Argentina" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
1;
<collectorName id="4DADCD8D10784B02A31AF28F6FF11003">Quéquen</collectorName>
;
<date id="ED456800219C0D6BBB417FE25FE585C6" value="1937-02">
<collectingDate id="97881AB062FBC5A7BF25359DED56C5BF" value="1937-02">Fev. 1937</collectingDate>
</date>
;
<abbrev id="ABBRID0EBFBI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="F41A50B4F1DE24EDEB8A5C921175BA0F">CEMT</collectionCode>
</abbrev>
</materialsCitation>
•
<materialsCitation id="C534F9BD1FCDCACE1569D99A1B527582" collectingDate="1937-02" collectionCode="DZUP" collectorName="Quequen" country="Argentina" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
1;
<collectorName id="816CAFDC6659FAA3151B5DBF7E2D749F">Quéquen</collectorName>
;
<date id="38AF355870CF70F2173288A99ED6980F" value="1937-02">
<collectingDate id="8C78A90C41AB8D3473427FBD586C68C4" value="1937-02">Fev. 1937</collectingDate>
</date>
;
<named-content id="287B8924F9562E90D24A289F5FAEF436" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidade-federal-do-parana-colecao-de-entomologia-pe-jesus-santiago-moure" xlink_title="Museu de Entomologia ‘Pe. Jesus Santiago Moure’, Universidade Federal do Paraná">
<collectionCode id="BD6FD10D65D7DE5E31BC71CEEA1C307E" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" type="University or college">DZUP</collectionCode>
</named-content>
</materialsCitation>
• 12;
<collectionCode id="BC07A61BA70B222AA96B21A8F51EA73C">A</collectionCode>
.
<collectorName id="EC6A6B8F1252C1E202841B9EC503D1CE">Breyer</collectorName>
leg.;
<named-content id="6814D3037B6D56F0FF8577791DE0A662" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’">
<collectionCode id="E54362D2F75BD932AB8EDDD0E9C69959" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</named-content>
• 2;
<collectorName id="DA95C2897BC7C0273537C3C4C5B060B9">C. Bruch</collectorName>
leg.
<named-content id="2B1EACF0AD70CD2B4EB9FA242E150D34" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’">
<collectionCode id="903F85DC8973B8D50794D72C46BDCE37" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</named-content>
•
<materialsCitation id="8C42486AB4D9DA8D80B37908959870CA" collectionCode="MACN" country="Argentina" location="Cordoba" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
175;
<named-content id="BF2172DD7A3CCDE4B514071E7406CFA8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-argentino-de-ciencias-naturales-bernardino-rivadavia" xlink_title="Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’">
<collectionCode id="FC9CA41B48FCEB9EFEBDBB449C03142A" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" type="Museum">MACN</collectionCode>
</named-content>
. –
<emphasis id="E3AD6CAD539E53EB679EF07CB18AE20C" bold="true">
<location id="646ADC34A69B78BC5D65BC6ED4261BB4" LSID="urn:lsid:plazi:treatment:BAC76C222D505D2BA37175E418D5B1F8:646ADC34A69B78BC5D65BC6ED4261BB4" country="Argentina" name="Cordoba" stateProvince="Ciudad Autonoma de Buenos Aires">Córdoba</location>
</emphasis>
</materialsCitation>
•
<materialsCitation id="A84CB7B9BB207C845303E2C5777F5407" country="Argentina" location="Miramar" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
10;
<location id="055F475442329D1EC5E8B6B99A747D7C" LSID="urn:lsid:plazi:treatment:BAC76C222D505D2BA37175E418D5B1F8:055F475442329D1EC5E8B6B99A747D7C" country="Argentina" name="Miramar" stateProvince="Ciudad Autonoma de Buenos Aires">Miramar</location>
; MLPA
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

399
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<subSubSection id="3269139D434245584C4351BA11B30001" type="nomenclature">
<paragraph id="8F573506AD4B03AC2038436AD399719D">
<taxonomicName id="F632F4A6AB76308188DF3A05B4B962AE" authority="Boruah, Narayanan, Aravind, Deepak &amp; Das" authorityName="Boruah, Narayanan, Aravind, Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siangensis" status="sp. nov.">
<emphasis id="4D4966C1A7BDAA5A945BADBB7E3B910E" italics="true">Cyrtodactylus siangensis</emphasis>
Boruah, Narayanan, Aravind, Deepak &amp; Das
</taxonomicName>
<taxonomicNameLabel id="3CFA17787918FC8AF0163E8B3513986B" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D4012049354A69CAF9DF534DD4B9899D" type="description">
<paragraph id="0931CE847F375A25F1CCC16E26D9E629">
<figureCitation id="252EB77CD49ACE5FA717FE50EF85102A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyrtodactylus siangensis sp. nov. A, B, E – I holotype (WII-ADR 1177), A dorsal, and B ventral view of whole body; C paratype (WII-ADR 1582); D paratype (WII-ADR 1581); E dorsal, F ventral, and G lateral view of the head (WII-ADR 1177); H ventral view of left manus, and I left pes (WII-ADR 1177); J an uncollected male in natural habitat from Jorsing, East Siang, Arunachal Pradesh. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure9" httpUri="https://binary.pensoft.net/fig/1102275">Figure 9</figureCitation>
;
<tableCitation id="39653474B3789BB3070AEE1EC514643B" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">Tables 2, S 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1NDK" type="Holotype">
<paragraph id="05EC52921DF1AC9C33E03A7192B04DE4">
<heading id="B2EDE1EED6B9E2F58E4CE20DDC398E0C" reason="title">
<typeStatus id="B02333E393AFCF1D9C3C757BE0E91F01" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="E095E7B39059F4D06C45837085032524">
<materialsCitation id="BEFE5CDC72BEC4D6B7A02F12E50D8A4A" collectingDate="2021-10-23" collectorName="Bitupan Boruah" country="India" elevation="210" latitude="28.1711" location="Bodak village" longLatPrecision="6" longitude="95.242" municipality="East Siang District" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
Adult female (WII-ADR 1177; Fig.
<figureCitation id="B8FEFDE37A633B6AA2534EF4F53E2FA7" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyrtodactylus siangensis sp. nov. A, B, E – I holotype (WII-ADR 1177), A dorsal, and B ventral view of whole body; C paratype (WII-ADR 1582); D paratype (WII-ADR 1581); E dorsal, F ventral, and G lateral view of the head (WII-ADR 1177); H ventral view of left manus, and I left pes (WII-ADR 1177); J an uncollected male in natural habitat from Jorsing, East Siang, Arunachal Pradesh. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure9" httpUri="https://binary.pensoft.net/fig/1102275">9 A, B, E – I</figureCitation>
), from Kalek stream (
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<geoCoordinate id="83221B6AEE17EF4303F4E764E599A5AD" degrees="28.1711" direction="north" orientation="latitude" precision="5" value="28.1711">28.1711 ° N</geoCoordinate>
;
<geoCoordinate id="80673C93C2CD920C69BD092E076F9322" degrees="95.2420" direction="east" orientation="longitude" precision="5" value="95.242">95.2420 ° E</geoCoordinate>
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,
<elevation id="362525FE987F4046CCE6AAB3162BC7F2" metricMagnitude="2" metricUnit="m" metricValue="2.1" unit="m" value="210.0">
elevation
<quantity id="79DE382069BECAE05D62583CAAFF383C" metricMagnitude="2" metricUnit="m" metricValue="2.1" unit="m" value="210.0">210 m</quantity>
a. s. l.
</elevation>
; Fig.
<figureCitation id="680FCDC4B106DC954816012E160DC97F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 A</figureCitation>
);
<quantity id="B3B09A6A7400F3070D27C0ED37B09E74" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="km" value="1.8">1.8 km</quantity>
(aerial distance) northwest from
<location id="4AD28F54663BA6854CE155690CFD0A8F" LSID="urn:lsid:plazi:treatment:CB20AD95DD3E53A58D7FC3B88F1BCC17:4AD28F54663BA6854CE155690CFD0A8F" country="India" latitude="28.1711" longLatPrecision="6" longitude="95.242" municipality="East Siang District" name="Bodak village" stateProvince="Arunachal Pradesh">Bodak village</location>
,
<collectingMunicipality id="6672C9623048A4DCDE134EF5815545A5">East Siang District</collectingMunicipality>
,
<collectingRegion id="73FC755FE57E30B1127535F5ADEA98D0" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="420F45A9E0428423E0CC53A49EE57C68" name="India">India</collectingCountry>
collected by
<collectorName id="09E50D72218E3C554E422AC8BCCD8C09">Bitupan Boruah</collectorName>
on
<date id="C58EB889EECABA6024C3DC5AC54746B5" value="2021-10-23">
<collectingDate id="0D6764AEBB13D690FC34B463024CE852" value="2021-10-23">23 October 2021</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<caption id="DA22507DAFB3BEBF784ACB2D33B55EB6" ID-DOI="10.3897/vz.74.e124752.figure9" ID-arpha="B3C96A6C-1CAB-5164-A8E5-C88081CF74D0" httpUri="https://binary.pensoft.net/fig/1102275" startId="F9">
<paragraph id="E9D6619F779FC826C5F928E3F0A9B46C">
<label id="8878DA47661BFAD303FF19B90171169E">Figure 9.</label>
</paragraph>
<paragraph id="28A6F92B0AFCB6C9065D5C16417771FB">
<taxonomicName id="763BC4653F0E8CB428CF44B10C9AA736" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siangensis" status="sp. nov.">
<emphasis id="D7176B19C1DD59FB3AA343A8EDF0045A" italics="true">Cyrtodactylus siangensis</emphasis>
</taxonomicName>
<emphasis id="5956CBC9334FD6D39B1E15DB0C3DD207" bold="true">
<taxonomicNameLabel id="DEED78164E2B63BE8E2DFB5DD6BD7BA5" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
,
<emphasis id="A68A8739DB265A5B97990727BFDAF3F7" bold="true">B</emphasis>
,
<emphasis id="1E8D5621599E17D5F187579C3339166B" bold="true">E</emphasis>
–
<emphasis id="814F943CC876D417723FAB59D3DCB73A" bold="true">I</emphasis>
holotype (WII-ADR 1177),
<emphasis id="69631AD9EE12C8396A8DCA38406CC9B9" bold="true">A</emphasis>
dorsal, and
<emphasis id="303C83588077C86070CE3B2EF9AE4C3C" bold="true">B</emphasis>
ventral view of whole body;
<emphasis id="DC82E60793E6506D85F2B6339824CEA9" bold="true">C</emphasis>
paratype (WII-ADR 1582);
<emphasis id="D51E513EDEB2CEE35D85927F7BED29F8" bold="true">D</emphasis>
paratype (WII-ADR 1581);
<emphasis id="3D722352F7AB86280F219A2375B2461E" bold="true">E</emphasis>
dorsal,
<emphasis id="5665FCDBE382F0F3E6C1E91DBA75A9F0" bold="true">F</emphasis>
ventral, and
<emphasis id="E2B7906F4B7B8D44E8368DE66198D696" bold="true">G</emphasis>
lateral view of the head (WII-ADR 1177);
<emphasis id="AE63576E38317412672F3FD2828E6657" bold="true">H</emphasis>
ventral view of left manus, and
<emphasis id="616F70A1A7445AE4EB39D135628DC8BA" bold="true">I</emphasis>
left pes (WII-ADR 1177);
<emphasis id="0AA545FE0820680272CEF63241944B10" bold="true">J</emphasis>
an uncollected male in natural habitat from Jorsing, East Siang, Arunachal Pradesh. Scale bar: 10 mm.
<emphasis id="DECBFBEEC8BB3D7BAAFECA957F12EA71" bold="true">J</emphasis>
not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EGQDK" type="Paratypes">
<paragraph id="5F69B82D1ECCF49A0327DF499C892301">
<heading id="DDDC5B5928FCF39D0CDCEC623AB999EB" reason="title">
<typeStatus id="3586B8CB4C144F250AD7E4FCAD8298B7" type="paratype">Paratypes</typeStatus>
.
</heading>
</paragraph>
<paragraph id="E1D7ECE38496E2DAFDE251A840CADF3D">
<specimenCount id="821D08F5249F402AA0B97869E0B30C96" count="2" type="adult">Two adult</specimenCount>
females (WII-ADR 1581 and WII-ADR 1582; Fig.
<figureCitation id="C23D8FCF326552F39CD8ECEDAD3869B8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyrtodactylus siangensis sp. nov. A, B, E – I holotype (WII-ADR 1177), A dorsal, and B ventral view of whole body; C paratype (WII-ADR 1582); D paratype (WII-ADR 1581); E dorsal, F ventral, and G lateral view of the head (WII-ADR 1177); H ventral view of left manus, and I left pes (WII-ADR 1177); J an uncollected male in natural habitat from Jorsing, East Siang, Arunachal Pradesh. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure9" httpUri="https://binary.pensoft.net/fig/1102275">9 C, D</figureCitation>
) from a hill slope near Kalek stream, East Siang District,
<collectingRegion id="18536F4F9E01708FCFC64337D1E8289C" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="0DF3FA7A8DCA8039D8D4B25E17A1FB47" name="India">India</collectingCountry>
collected by Bitupan Boruah on
<date id="1BC5B8AB66F8E7F627F99424C7D0919A" value="2022-06-15">15 June 2022</date>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQQDK" type="diagnosis">
<paragraph id="1C688C7ACDEDACFD426DDE5735F75550">
<heading id="58D351ECFD841A5B56D2885B0FF0F15B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="8B187A93FA82CB80BA8E5CF49331DEE5">
Medium-sized gecko (
<abbrev id="ABBRID0EWQDK" xlink_title="snout to vent length">SVL</abbrev>
<quantity id="59A664DDD316EED88D1E0FA6E7DBC122" metricMagnitude="-2" metricUnit="m" metricValue="7.109999999999999" metricValueMax="7.209999999999999" metricValueMin="7.01" unit="mm" value="71.1" valueMax="72.1" valueMin="70.1">70.1–72.1 mm</quantity>
in females); males unknown; supralabials 8–12 and infralabials 9–12; tubercles on dorsum weakly keeled and bluntly conical, 26–32 paravertebral tubercles between the level of axilla and level of groin; 15 or 16 dorsal tubercle rows at mid body; 40–45 mid ventral scale rows; 8–10 small precloacal pores in a continuous series; 13–17 subdigital lamellae on finger IV and 14–19 subdigital lamellae on toe IV; six or seven irregular and broken dark-brown bands on dorsum between the level of axilla and level of groin, or irregular dark-brown reticulation on dorsum.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1QDK" type="description">
<paragraph id="070706A309647F9B62EC3847A0232827">
<heading id="36DACFDDBFCD4030489FFADF06918ECF" reason="title">
Description of the
<typeStatus id="8D0646964F681C2B18296D718D8A5F5E" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="7B8CBE59E50571A2719944B317B1793E">
<typeStatus id="EE96083136EE3E240E149CF222DABC8B" type="holotype">Holotype</typeStatus>
well preserved except an incision below left axilla ventrolaterally. Snout-vent length
<quantity id="FA4839C4381A1B0F6E5B33AB9058A223" metricMagnitude="-2" metricUnit="m" metricValue="7.209999999999999" unit="mm" value="72.1">72.1 mm</quantity>
. Head moderately large (
<abbrev id="ABBRID0EARDK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
/
<abbrev id="ABBRID0EERDK" xlink_title="snout to vent length">SVL</abbrev>
= 0.26), dorsoventrally depressed, longer than width (
<abbrev id="ABBRID0EIRDK" xlink_title="maximum head width">HW</abbrev>
/
<abbrev id="ABBRID0EMRDK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.72), distinct from neck, broader at occipital region; snout rounded in both dorsal and lateral view; loreal region convex; canthus rostralis rounded, indistinct; interorbital space flat, a longitudinal furrow on dorsal surface of the snout, snout short (
<abbrev id="ABBRID0EQRDK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
/
<abbrev id="ABBRID0EURDK" xlink_title="head length from the retroarticular process of the jaw to the snout tip">HL</abbrev>
= 0.39), longer than orbit (
<abbrev id="ABBRID0EYRDK" xlink_title="orbit diameter, taken between the anterior and posterior bony orbital borders">OD</abbrev>
/
<abbrev id="ABBRID0E3RDK" xlink_title="distance between the snout tip and the anterior bony orbital border">SO</abbrev>
= 0.73); nostril nearly rounded, opening directed posterolaterally; ear opening oval and oblique; scales on head heterogeneous, largest on snout and loreal region, posteriorly smaller in upper eyelid, interorbital space and occipital region, granular juxtaposed; scales on upper eyelids heterogeneous; supraciliaries outwardly sharp giving serrated appearance in dorsal view, size anteriorly and posteriorly decreases, largest at approximately anterior one third of it; rostral wide, a short groove at the middle on top, rostral connected with nasals, supranasals, an internasal and first supralabials; a single scale between the supranasals, larger than the rest of the granular snout scales; granular scales at parietal region and occipital region intermixed with slightly large rounded granular tubercles, dense in occipital and temporal region and size increases towards nape; supralabial eight on right and 11 on left side, supralabials upto midorbit seven on right and eight on left side, size decreases towards angle of jaw; a series of scales nearly rounded, granular and slightly larger than the loreal scales present above the supralabials; mental as wide as rostral, triangular, connected with first infralabials, inner postmentals; 10 infralabials on right and nine on left side, size decreases towards angle of jaw, first infralabials connected with mental, second infralabial, inner and outer postmentals; inner pair of postmentals are larger than the outer postmentals, posterior margin of the inner postmentals bordered by eight granular scales of different size; one or two rows of enlarged scales along the infralabials starting below the outer postmentals, posteriorly size of those decreases, elongated and narrow; rest of the gular scales are small, granular juxtaposed, homogeneous, size increases towards throat where they become imbricate.
</paragraph>
<paragraph id="B3E44B157905B255E989FFC28CC268CA">
Habitus slender (
<abbrev id="ABBRID0ECSDK" xlink_title="maximum body width">BW</abbrev>
/
<abbrev id="ABBRID0EGSDK" xlink_title="snout to vent length">SVL</abbrev>
= 0.18,
<abbrev id="ABBRID0EKSDK" xlink_title="trunk length, between the axilla and groin">TRL</abbrev>
/
<abbrev id="ABBRID0EOSDK" xlink_title="snout to vent length">SVL</abbrev>
= 0.45), dorsoventrally depressed; dorsal scales granular, rounded, heterogeneous, intermixed with rounded, weakly keeled and bluntly conical tubercles irregularly arranged, starting from occipital region to fourth segment of the tail, size increases towards posterior body, pronounced at sacrum and base of the tail; 15 dorsal tubercles across mid dorsum; 26 paravertebral tubercles; ventrolateral fold weak; ventral scales larger than those of dorsal, flat, smooth, cycloid subimbricate to imbricate, largest on belly; 40 mid-ventral scales between ventrolateral fold; very small, 10 precloacal pores arranged in an inverted “
<collectionCode id="0F2145D42EA894EA0D11249394B6CCE4" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
” shaped continuous series, followed by 7 unpored, large scales below it, largest at the apex.
</paragraph>
<paragraph id="013497E41EA44203F59FB9DA1ABD37F1">
Forelimbs and hindlimbs slender (
<abbrev id="ABBRID0EUSDK" xlink_title="forearm length, taken from the elbow to the wrist">FL</abbrev>
/
<abbrev id="ABBRID0EYSDK" xlink_title="snout to vent length">SVL</abbrev>
= 0.14,
<abbrev id="ABBRID0E3SDK" xlink_title="crus length, taken from the knee to the base of the foot">CL</abbrev>
/
<abbrev id="ABBRID0EATDK" xlink_title="snout to vent length">SVL</abbrev>
= 0.17); digits strongly inflected at the joints, all bearing large recurved claw, enlarged subdigital lamellae, lamellae beneath digit IV of right and left manus (given as basal + distal) is 6 + 11 and 6 + 10 respectively, the second most lower lamellae of the basal series of digit IV of both sides are divided (not included in lamellar count); lamellae beneath digit IV of right and left pes (given as basal + distal) is 7 + 11 and 6 + 10 respectively; dorsal scales on forelimbs heterogeneous, granular juxtaposed, smooth and subimbricate at distal end of forearm; forearm intermixed with a few rounded, large tubercles; dorsal scales of hindlimbs granular intermixed with large, rounded, bluntly conical tubercles; scales on thigh towards knee smooth subimbricate; ventral scales of forelimbs granular, juxtaposed, mostly homogeneous; scales on palm heterogeneous, granular juxtaposed; scales on hindlimbs smaller than those of belly, smooth, cycloid and subimbricate; scales on the knee, above cloaca and on thigh below the level of precloacal pores are smaller and granular; scales on soles heterogeneous, granular, juxtaposed to subimbricate.
</paragraph>
<paragraph id="8EA61F4C90E7148ED1E6BD98153FDAEF">
Tail complete, regenerated (
<abbrev id="ABBRID0EGTDK" xlink_title="tail length, from the cloaca to the tail tip">TL</abbrev>
=
<quantity id="AF6F4BFB4BF492B4886A915E77FB49B1" metricMagnitude="-2" metricUnit="m" metricValue="8.0" unit="mm" value="80.0">80 mm</quantity>
), slender, gradually tapering, segments indistinct; dorsal scales small, granular, juxtaposed at the base, posteriorly size increases, flat, smooth, subimbricate, heterogeneous in shape and size; large feebly keeled scales up to fourth segment of the tail at the distal end of each segment, those on basal segment are pronounced; subcaudal scales smooth, subimbricate, wider than that of dorsal, heterogeneous in shape and size; below cloaca scales are granular, smaller than the rest of the tail scales; no enlarged plate like series of subcaudal scales; three bluntly conical spurs on each side of the tail base present.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKTDK" type="Colouration in preservative">
<paragraph id="CB6ED7DF5010F3920A9A28D40893D236">
<heading id="D34CB7BF4258C372462F0AA213CCEE27" reason="title">Colouration in preservative.</heading>
</paragraph>
<paragraph id="525E5EE33D636FCB1618BB92D78457D5">Top of head, limbs and tail pale-brown; dorsum pale-greyish-brown; a slightly dark-brown stripe radiating from posterior orbital border to occipital region, medially “ W ” shaped; an another short dark-brown stripe at the middle of the head; a short, posteriorly diffused dark-brown stripe starting from posterior corner of the upper eyelids; a faint brown stripe on the loreal region; rostral and mental scale, labial scales dark-greyish-brown with white patch posteriorly; short and irregular shaped, longitudinal dark-brown stripe on dorsal side of neck; six irregular shaped, zig-zagged dark-brown bands on dorsum; irregular shaped, large dark-brown spots on sacrum and tail base; zig-zagged dark-brown bands on anterior part of the tail, posterior half of the tail (regenerated part) with brown speculation; indistinct brown reticulation on dorsal side of the limbs; ventrally head, body and limbs creamy-white with brown speculation; chin scales and scales below infralabials heavily speckled; lamellae, palm and feet heavily speckled with brown; indistinct brown patches on ventral side of the tail.</paragraph>
</subSubSection>
<subSubSection id="SECID0EPTDK" type="Morphological variation">
<paragraph id="9C86E3F81F5B1F0DC1742BC3E54B1D2E">
<heading id="77F76E3D25BF0B75EA099D17DCBDAE78" reason="title">Morphological variation.</heading>
</paragraph>
<paragraph id="02BBE62FA49B1ADC61149D264A3D0734">
Detailed morphological variations among the collected individuals of
<taxonomicName id="48FE8FE5B868A7A4E765F99A4B357BCC" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siangensis">
<emphasis id="924F6D8A1DC5A8BC094B6F0D7E79AC1D" italics="true">Cyrtodactylus siangensis</emphasis>
</taxonomicName>
are provided in Table S 2. Dorsal dark brown markings in the
<specimenCount id="518019E06E96980358CA8EE81341592B" count="2" type="generic" typeStatus="paratype">two paratypes</specimenCount>
are variable in the form of irregular zig-zagged bands to reticulation; reticulation on limbs of the
<specimenCount id="A1B9E6929F0D9E0190F860205A87D672" count="2" type="generic" typeStatus="paratype">two paratypes</specimenCount>
distinct than the
<typeStatus id="82BE053DCEFAE93EDCB7E8966D1C87A9" type="holotype">holotype</typeStatus>
; cross bands on tail of WII-ADR 1581 darker posteriorly (Fig.
<figureCitation id="F9E77C93362C422507286D4086577A14" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyrtodactylus siangensis sp. nov. A, B, E – I holotype (WII-ADR 1177), A dorsal, and B ventral view of whole body; C paratype (WII-ADR 1582); D paratype (WII-ADR 1581); E dorsal, F ventral, and G lateral view of the head (WII-ADR 1177); H ventral view of left manus, and I left pes (WII-ADR 1177); J an uncollected male in natural habitat from Jorsing, East Siang, Arunachal Pradesh. Scale bar: 10 mm. J not to scale." figureDoi="10.3897/vz.74.e124752.figure9" httpUri="https://binary.pensoft.net/fig/1102275">9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEUDK" type="comparisons">
<paragraph id="99CF3EC260CB23ED29C21CF90AA03BEC">
<heading id="6A79B4F80FE9C7B4F79A1D6AC21535A0" reason="title">Comparison.</heading>
</paragraph>
<paragraph id="53114BF7C7E9FB0A57AD389D5C144E2B">
Morphologically
<taxonomicName id="CFEC5EA97E8BF44F977ABA5BF7767B88" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siangensis" status="sp. nov.">
<emphasis id="B6139A7B9588E33C83B751699E46C836" italics="true">Cyrtodactylus siangensis</emphasis>
</taxonomicName>
<emphasis id="CD8BE0418374F3099CB683CF6B7B2DEC" bold="true">
<taxonomicNameLabel id="8541F0A9234CA3D95AB9AC648E6AFC06" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is close to
<taxonomicName id="DDCF719A2032282E4DB05E08C4D71297" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="30378C067B4D3CF3EC192959B1071B8C" italics="true">C. cayuensis</emphasis>
</taxonomicName>
, however, phylogenetically it is distinct from the later (Fig.
<figureCitation id="E493ADCCD8DC4BDD209B5132B74295CE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. ML phylogeny showing the relationship of the khasiensis group. New species described here are marked in blue, other samples generated in this study are in bold. Values in the node represent PP and UFB on the left and right side, respectively. See Fig. S 1 for a full tree with outgroups." figureDoi="10.3897/vz.74.e124752.figure2" httpUri="https://binary.pensoft.net/fig/1102268">2</figureCitation>
; Table S 3), also the
<typeStatus id="DBD0E909526F0BFEB24AD2015A63052B">type</typeStatus>
locality of the new species is separated by Siang River from the
<taxonomicName id="A71D22EDDB0C8781C5FE30901522709A" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="79B4D84F12B7C4F376A7C3A3ED7C05EC" italics="true">C. cayuensis</emphasis>
</taxonomicName>
localities recorded in this study (Fig.
<figureCitation id="6B147B0FD36A22074F3D59A5756D89AF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Map showing geographic distribution of the clades within the khasiensis group. A type localities of the members of the four clades in the Indo-Burma region: 1 Cyrtodactylus septentrionalis, 2 C. guwahatiensis, 3 C. urbanus, 4 C. kazirangaensis, 5 C. siangensis sp. nov., 6 C. cayuensis, 7 C. namdaphaensis sp. nov., 8 C. bapme, 9 C. karsticola, 10 C. agarwali, 11 C. exercitus, 12 C. jaintiaensis, 13 C. khasiensis, 14 C. barailensis sp. nov., 15 C. nagalandensis, 16 C. kiphire sp. nov., 17 C. namtiram, 18 C. vairengtensis, 19 C. manipurensis sp. nov., 20 C. mombergi, 21 C. dianxiensis, 22 C. tripuraensis, 23 C. montanus, 24 C. aaronbaueri, 25 C. bengkhuaiai, 26 C. ngopensis, 27 C. lungleiensis, 28 C. ngengpuiensis sp. nov., 29 C. siahaensis, 30 C. gansi, 31 C. myaleiktaung, 32 C. brevidactylus, 33 C. aunglini, 34 C. chrysopylos, 35 C. ayeyarwadyensis; B cayuensis clade, green star (type locality of C. siangensis sp. nov.), pink star (type locality of C. cayuensis), pink triangle (C. “ arunachalensis ”), pink circle (C. cayuensis recorded in this study) (numbers correspond to the locality given in Table 1)." figureDoi="10.3897/vz.74.e124752.figure3" httpUri="https://binary.pensoft.net/fig/1102269">3 B</figureCitation>
). Morphological differences with other members of
<taxonomicName id="1DB55C061D78287B4E451ABFCF20D9C5" baseAuthorityName="Jerdon" baseAuthorityYear="1870" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="011327739C2C333590BBBDCEB925CB02" italics="true">khasiensis</emphasis>
</taxonomicName>
group is presented in Table
<tableCitation id="A8ACF4716B0A744877B5648369693E25" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparative table of summary morphological characteristics among the members of the khasiensis group including the seven new species. NA- data not available; “ * ” data from the literature is not comparable with our data as the count method is different." httpUri="http://table.plazi.org/id/4AAE94DC4D30D8EB1A84D336CC270F8B" tableUuid="4AAE94DC4D30D8EB1A84D336CC270F8B">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDWDK" type="Sequence divergence">
<paragraph id="0BAFF74AC94EC9F18E2AAE89807B6A70">
<heading id="D09661282070E1A935C427AACCF6E9E9" reason="title">Sequence divergence.</heading>
</paragraph>
<paragraph id="7B7A643A528EE22D3D1721F8F4C18C60">
<taxonomicName id="4D860457593EEBAD8F893F5F175D4FAB" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siangensis" status="sp. nov.">
<emphasis id="39098434338B6140DD4C129DE140FB2B" italics="true">Cyrtodactylus siangensis</emphasis>
</taxonomicName>
<emphasis id="91D6671D0B73219B0422ADC4AEC874D5" bold="true">
<taxonomicNameLabel id="B36E1A6D1C569A62D06D3FF494ED621E" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a high genetic divergence of 11.7–17.0 % from its closely related and the only other species in the clade,
<taxonomicName id="1A1290AD89CF8882DD608E16D92AE57A" authorityName="Li" authorityYear="2007" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="cayuensis">
<emphasis id="AD4B376D7E7B75835F82505AC01FFA4C" italics="true">C. cayuensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBXDK" type="etymology">
<paragraph id="A6074BB200BCF578BACABEB66725C42C">
<heading id="7E9DEA0816DA5C12EFD99A8E0B4267CC" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="ADF92A7891E5B7337EBF73F0C33EAC49">
The specific epithet is a toponym derived from the name of the river “ Siang ”. The
<typeStatus id="FCEAF392A1B25D5387698503822F96CD">type</typeStatus>
locality of this species lies in the Siang valley of
<collectingRegion id="0F215FDB60C2462F59AAB4544F16D66E" country="India" name="Arunachal Pradesh">Arunachal Pradesh</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGXDK" type="Suggested common name">
<paragraph id="BC9A6767D2D78A19A395C32C02F25011">
<heading id="FE057BD1A4D6B3FDA07F1EB40861F808" reason="title">Suggested common name.</heading>
</paragraph>
<paragraph id="9627250E832273465F1D9898FDAE9B06">Siang Valley bent-toed gecko.</paragraph>
</subSubSection>
<subSubSection id="SECID0ELXDK" type="distribution">
<paragraph id="F867C425F5F7E9858DF5D2B3E923F8D1">
<heading id="E986A22FB89F9FB90F58B8BB40F7AAC2" reason="title">Distribution and natural history.</heading>
</paragraph>
<paragraph id="3D6992AD4A3927F6CD2E68E3426ADDBC">
<taxonomicName id="49B253F393A3B97FB6F7E89940223F06" authorityName="Boruah &amp; Narayanan &amp; Aravind &amp; Lalronunga &amp; Deepak &amp; Das" authorityYear="2024" class="Squamata" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="siangensis" status="sp. nov.">
<emphasis id="079763C31E537A4DE8193FE6AFA402E6" italics="true">Cyrtodactylus siangensis</emphasis>
</taxonomicName>
<emphasis id="C753C286A09A8ABA20BE75C2C7C6708D" bold="true">
<taxonomicNameLabel id="39BBBF59C2DBD3B2CC4EEBFBE0FAC98D" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently only known from the type locality, Bodak, East Siang District. It appears to be uncommon, and likely a habitat generalist. The
<typeStatus id="2117758CA18B561AE75087D629C5E577" type="holotype">holotype</typeStatus>
was encountered on a shrub at the edge of Kalek stream (a tributary of Siang River) at a height of approximately one meter above the ground, at 18: 15 hrs. The stream at the time of survey was flowing at a moderately speed, was approximately
<quantity id="EBD3810FEDEDA1E97BF9C024B4A174A7" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10 m</quantity>
in width and contained boulders (sedimentary rock) and fallen logs. The
<specimenCount id="DDD8EC2706169E1FFBC469587A020F17" count="2" type="generic" typeStatus="paratype">two paratypes</specimenCount>
were recorded on a small tree at a height of 2.0 m on the slope of a hill close to the collection site for the
<typeStatus id="DCAC4CA0E441CDC556170FE8E9627C59" type="holotype">holotype</typeStatus>
. The type locality is ca.
<quantity id="7601505E58A4B7E5A3AFCB40C359429E" metricMagnitude="2" metricUnit="m" metricValue="3.0" unit="m" value="300.0">300 m</quantity>
from Siang River and approximately
<quantity id="1E705747F034622C96F4EED0CC04B8FC" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</quantity>
from the road connecting Pasighat and Yinkiong.
</paragraph>
</subSubSection>
</treatment>
</document>

217
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<mods:title id="AE05694B3A3A3C2A6676EF3F93BEE4A5">Four new Dolichopoda species from Greece, one Troglophilus new to Greece and new locality records (Orthoptera, Rhaphidophoridae)</mods:title>
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<mods:namePart id="568EC55F4BB8493E49EADDCB3E6A9A56">Alexiou, Sotiris</mods:namePart>
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<mods:affiliation id="CE6B4FD88730940DFC6AD24BD37C676F">Korinthian Museum of Natural History, Isthmion 201, Korinthos 20100, Greece</mods:affiliation>
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<mods:namePart id="2CF32F6BC65522A8F81CD67124A9FC35">Bakolitsas, Kostas</mods:namePart>
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<mods:affiliation id="3C2A0360C548FDF2920243464A79D20D">Laboratory of Natural Sciences 3 rd General Lyceum, 301 31 Agrinion, Greece</mods:affiliation>
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<mods:namePart id="C95FE06C3D71B3151611377B71C1A4AC">Rampini, Mauro</mods:namePart>
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<subSubSection id="1F3A796F4C1D761D10D42E7B039E1745" type="nomenclature">
<paragraph id="8D65CA0DF030069E1AFFEA43D7E9A9A3">
<taxonomicName id="4B5A8D024D0280F03AC77863D20A6FB2" authority="Alexiou" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="athosensis" status="sp. nov.">
<emphasis id="96CC16163D20D9C4D6397D7C327213C1" italics="true">Dolichopoda athosensis</emphasis>
Alexiou
</taxonomicName>
<taxonomicNameLabel id="043B5C25A4C5D40008578D3D070935E7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="700DFDC9CC0B71C7E23959005DF855D4" type="description">
<paragraph id="CA9DF881F39261A914A8670FA068CDC2">
<figureCitation id="8B2A51D86BDCDAC77215BE5AED609FEE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographic distribution of D. athosensis sp. nov., D. dirussoi sp. nov., D. karoutsosi sp. nov. and D. kotsabasi sp. nov." figureDoi="10.3897/contrib.entomol.74.e121614.figure1" httpUri="https://binary.pensoft.net/fig/1103848">Figs 1</figureCitation>
,
<figureCitation id="12890FDF264CE1C4A5F4881DBE804418" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dolichopoda athosensis sp. nov. male: A. Subgenital plate, ventral view; B. Epiphallus, lateral view; C. X tergum, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure2" httpUri="https://binary.pensoft.net/fig/1103849">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDIAC" type="type material">
<paragraph id="1C8D9EEC9B9B6CAEE9F543E6B8ACABD9">
<heading id="8D8288E292D10D347F7EB3C6AED19930" reason="title">Type material.</heading>
</paragraph>
<paragraph id="C7BE4369B254A623E965C9968E3DE05B">
<materialsCitation id="5CA4B1046D5EF94DC1ECB99C371EA30F" collectingDate="2017-07-15" collectorName="K. Bakolitsas" country="Greece" county="Chalkidikis" latitude="40.308132" location="cave Skiti" longLatPrecision="1" longitude="24.163933" municipality="Zographou" specimenCount="1" specimenCount-male="1" stateProvince="Dytiki Makedonia" typeStatus="holotype">
<emphasis id="D6D295ED50382C6E6893F84C8841CE88" bold="true" italics="true">
<typeStatus id="8537F9037D015CF91AED81EDB17AB291" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="64B8ADC976A4D9AEF643541022604916" name="Greece">GREECE</collectingCountry>
•
<specimenCount id="CA3391654C891E4579F6B8F78977EB80" count="1" type="male">♂</specimenCount>
;
<collectingRegion id="E75B88605B9F58A674828BE1608C4BDD" country="Greece" name="Dytiki Makedonia">Macedonia</collectingRegion>
, Νomos
<collectingCounty id="83E36C424775DF2563DB4F46E3A5BCFD">Chalkidikis</collectingCounty>
,
<collectingRegion id="3197673A095D077A032015E1B99C6B2F" country="Greece" name="Agion Oros">Agion Oros</collectingRegion>
peninsula, Mt.
<collectingRegion id="5954FEE28398D286C1A2D358CFB5F745" country="Greece" name="Agion Oros">Athos</collectingRegion>
,
<collectingMunicipality id="1034BCFA26C03B7C51AAAF80F70D3F49">Zographou</collectingMunicipality>
monastery,
<location id="5D18AB323A09C8BE4F4FAA58A6D8CF23" LSID="urn:lsid:plazi:treatment:D800469F5A535AD69C71745FB63330DF:5D18AB323A09C8BE4F4FAA58A6D8CF23" country="Greece" county="Chalkidikis" latitude="40.308132" longLatPrecision="1" longitude="24.163933" municipality="Zographou" name="cave Skiti" stateProvince="Dytiki Makedonia">cave Skiti</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERIAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[24.163933,40.308133]}">
<geoCoordinate id="FC18BB79D4CCFAED33AFF101394B7474" degrees="40" direction="north" minutes="18.488" orientation="latitude" precision="1" value="40.308132">40 ° 18.488 ' N</geoCoordinate>
,
<geoCoordinate id="A8DC9D39114E24BD6BD43A55844FECD2" degrees="24" direction="east" minutes="9.836" orientation="longitude" precision="1" value="24.163933">24 ° 9.836 ' E</geoCoordinate>
</named-content>
;
<date id="A795C90D0B7A015E28834E9A7F0D2874" value="2017-07-15">
<collectingDate id="0410AE9EEB134944FF422A16ACE84F2F" value="2017-07-15">15 Jul. 2017</collectingDate>
</date>
;
<collectorName id="82B0D792B818D16F7287F44311EA5BC9">K. Bakolitsas</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0EWIAC" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
.
<materialsCitation id="873ADFB757F81D274CA76BA19E29C836" collectingDate="2017-07-15" collectorName="K. Bakolitsas" country="Greece" county="Chalkidikis" latitude="40.308132" location="cave Skiti" longLatPrecision="1" longitude="24.163933" municipality="Zographou" specimenCount="1" stateProvince="Dytiki Makedonia" typeStatus="paratype">
<emphasis id="13A322E97D7740104768133E151F3E2C" bold="true" italics="true">
<typeStatus id="15CBEAE3163C3D97D6F518D8139EA7FC" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="46B3D38ECB11ECD5EFE64459063DFDB1" name="Greece">GREECE</collectingCountry>
• 5 nymphs, same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5IAC" type="diagnosis">
<paragraph id="7A8E6A91069CD9748A14C37E3CA376E1">
<heading id="EB872E0F31509315FEC2AB9D9E43B2CD" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="A843CAAE5168A795955514E911B6C0BC">
The new species is related to
<taxonomicName id="24120C9EF5308815353F9875445F3267" authority="Chopard, 1964" authorityName="Chopard" authorityYear="1964" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="thasosensis">
<emphasis id="D8968CE5FE2E57C3C94DBBC9C2D32048" italics="true">D. thasosensis</emphasis>
<bibRefCitation id="7E5BC74EB23B587480A0EB9B8E443AAF" DOI="10.3406/bsef.1964.20741" author="Chopard L" journalOrPublisher="Bulletin de la Société Entomologique de France" pagination="7-20" refId="B5" refString="Chopard L (1964) Descriptions d'orthopteres cavernicoles de Grece. Bulletin de la Societe Entomologique de France 69: 7 - 20. https: // doi. org / 10.3406 / bsef. 1964.20741" title="Descriptions d’orthoptères cavernicoles de Grèce." volume="69" year="1964">Chopard, 1964</bibRefCitation>
</taxonomicName>
, endemic to the island of Thasos (
<bibRefCitation id="B2A81D9B92258B7FCA37E7827C13B489" DOI="10.3406/bsef.1964.20741" author="Chopard" firstAuthor="Chopard" journalOrPublisher="Bulletin de la Société Entomologique de France" pagination="7-20" refId="B5" refString="Chopard L (1964) Descriptions d’orthoptères cavernicoles de Grèce. Bulletin de la Société Entomologique de France 69: 7–20. https://doi.org/10.3406/bsef.1964.20741" title="Descriptions d’orthoptères cavernicoles de Grèce." volume="69" year="1964">Chopard 1964</bibRefCitation>
). It differs in the less pronounced lateral lobes of the X tergite, the dramatically curved median process of the epiphallus and the subgenital plate, which has broader incision with larger styli.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETJAC" type="description">
<paragraph id="25312ACBC65DDD1AB7ECB382A5256A33">
<heading id="4FF696C5B842AB21553EF3FCE12319DA" reason="title">Description.</heading>
</paragraph>
<paragraph id="F83F71E84C110D1E39CB4C36D5B32AD1">Body pale testaceous, posterior margin of the terga dark. Legs elongated, hind femora unarmed. Fore tibiae with 4 spines on both sides of the ventral side, mid tibiae with 5 spines on both sides of the ventral side and hind tibiae with 18 spines on both sides of the dorsal side.</paragraph>
<paragraph id="7FABC61908DB57A34DB8B50EB9A4237B">
X tergum (last abdominal tergite) (Fig.
<figureCitation id="2E2E68F04D09F52BF00269B5014C4BFF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dolichopoda athosensis sp. nov. male: A. Subgenital plate, ventral view; B. Epiphallus, lateral view; C. X tergum, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure2" httpUri="https://binary.pensoft.net/fig/1103849">2 C</figureCitation>
) square, with a shallow section in the middle and with two lateral lobes. Lobes not very pronounced, with rounded apex.
</paragraph>
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<paragraph id="2F78453E41B60CD3928E2F3759F69811">
<label id="FA54DB6273D3A6B02EFDC54EBC498EF8">Figure 2.</label>
</paragraph>
<paragraph id="DE86C3B8EAD9C910DFB62A56F6596F28">
<taxonomicName id="1290E0798ED8573BB57CA6E16F18516C" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="athosensis" status="sp. nov.">
<emphasis id="F3A758A7654135F09CA08426F1D425DF" italics="true">Dolichopoda athosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E357535C8005024838002FA880C83E68" rank="species">sp. nov.</taxonomicNameLabel>
male:
<emphasis id="17FB3EF0B112B0552B7615117816BCF6" bold="true">A.</emphasis>
Subgenital plate, ventral view;
<emphasis id="AFA8867572E31C27BE97FF83EF4CA842" bold="true">B.</emphasis>
Epiphallus, lateral view;
<emphasis id="D41924BCCCD4B8B1EE8C4E7041A1DD91" bold="true">C.</emphasis>
X tergum, dorsal view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="60074B68292AC16D84640AE2C59DB216">
Subgenital plate (Fig.
<figureCitation id="D692A37A9D45DF78B518F72C0AE4A25C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dolichopoda athosensis sp. nov. male: A. Subgenital plate, ventral view; B. Epiphallus, lateral view; C. X tergum, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure2" httpUri="https://binary.pensoft.net/fig/1103849">2 A</figureCitation>
) divided by a broad median incision, with triangular lateral lobes ; styli short.
</paragraph>
<paragraph id="F8DDD41528B0F44D1D8DFC39C8E958A0">
Epiphallus sclerotized (Fig.
<figureCitation id="5B2F84772671E74587CF71D234FA929B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dolichopoda athosensis sp. nov. male: A. Subgenital plate, ventral view; B. Epiphallus, lateral view; C. X tergum, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure2" httpUri="https://binary.pensoft.net/fig/1103849">2 B</figureCitation>
). Median process of epiphallus with a broad base, strongly curved in lateral view, evenly leading to a rounded tip.
</paragraph>
<paragraph id="51FEC5808C52DDE172B0B93C32E99376">Measurements (length in mm): body 17, pronotum 4, fore femora 16, middle femora 17, hind femora 17.</paragraph>
<paragraph id="241ACA8AC75D88A2F6D2A0847F5E8DF7">
<emphasis id="C948B11917957529DD994BCAF5864C49" bold="true">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENLAC" type="etymology">
<paragraph id="FAE7681FDC4A34A48EC1B422FB5108D0">
<heading id="285AEE230EC25C5F73673C755AD2D889" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A4BA0284224B5399D39E56D657DF1247">
The new species is named after
<collectingRegion id="AEA5C3F3E5A1C787133FF1A04CEEAF80" country="Greece" name="Agion Oros">Athos</collectingRegion>
, a mountain dominating the
<collectingRegion id="EB0A421E57028EF369689A8E09186DDE" country="Greece" name="Agion Oros">Agion Oros</collectingRegion>
peninsula.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESLAC" type="distribution">
<paragraph id="C9B22298B7FC76EB425108E43F565B97">
<heading id="D8D14487E866EE0A5BBDFD2B8AA06FBA" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="FFC00824E87B69FF7F033A09773ECA8B">
Known only from the
<typeStatus id="0DF2D3062F2643BB4970B5324EE5A93F">type</typeStatus>
locality. Cave Skiti is located a few meters away from the Monastery of Zographou. The cave is about
<quantity id="88BBED58C16B181AA405F55165F5E380" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10 m</quantity>
long, with the entrance facing south-east. The entrance is small (around 0.5 × 1.0 m), and well hidden by lush vegetation.
</paragraph>
</subSubSection>
</treatment>
</document>

131
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<taxonomicName id="5E79E726F54A242248BCDED589AE4E7E" ID-CoL="8P972" authority="Chopard, 1964" authorityName="Chopard" authorityYear="1964" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="96561C0B50FF87D9883D9BC84890BC87" italics="true">Dolichopoda graeca</emphasis>
<bibRefCitation id="0E2EA3BFB0E1A6490E9DFF480D0E042C" DOI="10.3406/bsef.1964.20741" author="Chopard L" journalOrPublisher="Bulletin de la Société Entomologique de France" pagination="7-20" refId="B5" refString="Chopard L (1964) Descriptions d'orthopteres cavernicoles de Grece. Bulletin de la Societe Entomologique de France 69: 7 - 20. https: // doi. org / 10.3406 / bsef. 1964.20741" title="Descriptions d’orthoptères cavernicoles de Grèce." volume="69" year="1964">Chopard, 1964</bibRefCitation>
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<paragraph id="ABF1A18CDFA7D09904622483A35AB9F1">
<heading id="6EA5ACE5DA8E60E05527828E3093FDDA" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="C48C01A22F52F2ADC2DFF67B70E1777A">
<materialsCitation id="629DBC22253994800A0A2531165D8767" collectingDate="2021-10-17" collectorName="S. Alexiou &amp; S. Zacharias &amp; O. Tzortzakaki" country="Greece" county="Nomos Ioanninon" location="Aslan Tzami" municipality="Ioannina" specimenCount="2" specimenCount-male="2" stateProvince="Ipeiros">
<collectingCountry id="4302ECC10A9DADB227A6F0B746BE02B0" name="Greece">GREECE</collectingCountry>
•
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;
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,
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,
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(ʻKatakomvesʼ) ;
<date id="8CC595E91203F9126A381045315E717C" value="2021-10-17">
<collectingDate id="3767EE7CBF522C46CF2038C47B4F90F0" value="2021-10-17">17 Oct. 2021</collectingDate>
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;
<collectorName id="204E34BADCC78394D127C626D7E0EB56">S. Alexiou</collectorName>
,
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&amp;
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leg. (
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)
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.
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<paragraph id="1421DDACC4F32298920DCF4610832F2D">
<heading id="B0B9394D7FC26B3A0A628E8C340D0179" reason="title">Comment.</heading>
</paragraph>
<paragraph id="5F3D1A497B16C73E54A031E3FA16FCE2">
Anna Petrochilou, the prominent Greek speleologist, was the first to observe and report (
<bibRefCitation id="B762A172149DD8760680610B6E469468" author="Petrochilou A" refId="B11" refString="Petrochilou A (1975–1976) Cave ‘ Katakomvon’ Ioanninon. Bulletin la Société Spéléologique de Gréce XIII: 106–113. [in Greek]">Petrochilou 1975–1976</bibRefCitation>
) cave crickets at her inspection of ‘ Katakomves’ which are the caves situated underneath the mosque of Aslan Tzami. The mοsque was built by the Ottoman pasha Aslan in
<quantity id="50AE964B68FF0BAAB7681F7C50D91875" metricMagnitude="1" metricUnit="m" metricValue="4.10972" unit="in" value="1618.0">1618 in</quantity>
the castle of the town of Ioannina and is now operating as the Municipal Museum of Ioannina. After almost half a century, a name can finally be assigned to the population; they belong to
<taxonomicName id="DA95BCCE69BCF7C5AA0343EBA0002C01" authorityName="Chopard" authorityYear="1964" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="graeca">
<emphasis id="4B58D4C155D7A7EE25C308943997F9C5" italics="true">Dolichopoda graeca</emphasis>
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, a species known only from the
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locality, the cave of Perama,
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from the town of Ioannina (
<bibRefCitation id="CCB300E09252F9A2AD48FA0B1CFE558E" DOI="10.3406/bsef.1964.20741" author="Chopard" firstAuthor="Chopard" journalOrPublisher="Bulletin de la Société Entomologique de France" pagination="7-20" refId="B5" refString="Chopard L (1964) Descriptions d’orthoptères cavernicoles de Grèce. Bulletin de la Société Entomologique de France 69: 7–20. https://doi.org/10.3406/bsef.1964.20741" title="Descriptions d’orthoptères cavernicoles de Grèce." volume="69" year="1964">Chopard 1964</bibRefCitation>
;
<bibRefCitation id="2029AA599B3FEB0904C833C0BE1B9806" author="Di Russo" etAl="et al." firstAuthor="Di Russo" journalOrPublisher="Biodiversity Journal" pagination="397-420" refId="B7" refString="Di Russo C, Rampini M, Cobolli M (2014) The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss, 1879. Biodiversity Journal 5: 397–420." title="The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss, 1879." volume="5" year="2014">Di Russo et al. 2014</bibRefCitation>
).
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<mods:affiliation id="13293D15FA76BA6B4849824BA4D06A4A">Dipartimento di Biologia e Biotecnologie “ C. Darwin ”, Università di Roma “ La Sapienza ”, Roma, Italy</mods:affiliation>
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<paragraph id="3BE5D90FE91688E8EA5A81AAD4ECBCAF">
<taxonomicName id="2C9ABD1F374078790768CFB10CB0EEEA" authority="Alexiou" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="karoutsosi" status="sp. nov.">
<emphasis id="908E0FADF17315D0BE52B7EE5FB2EE9B" italics="true">Dolichopoda karoutsosi</emphasis>
Alexiou
</taxonomicName>
<taxonomicNameLabel id="E45977E6CFB82A874C1AC838E30E8FFE" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="1ED8D8F36D10D7097A2C595DE04B88D3" type="description">
<paragraph id="A9A86F1C3993357D9FFC1CC8F80F7D4C">
<figureCitation id="95A0C24E0F2A2B0011883031C64F0507" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographic distribution of D. athosensis sp. nov., D. dirussoi sp. nov., D. karoutsosi sp. nov. and D. kotsabasi sp. nov." figureDoi="10.3897/contrib.entomol.74.e121614.figure1" httpUri="https://binary.pensoft.net/fig/1103848">Figs 1</figureCitation>
,
<figureCitation id="09D4F5D8730758140C4EA3AEEAA30C83" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Geographic distribution of Dolichopoda species at W Sterea Ellas and S Ionian Islands." figureDoi="10.3897/contrib.entomol.74.e121614.figure4" httpUri="https://binary.pensoft.net/fig/1103851">4</figureCitation>
,
<figureCitation id="FF801DA002D381EC148B888AD0B5766E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dolichopoda karoutsosi sp. nov., male: A. X tergum, dorsal view; B. Epiphallus, dorsal view; C. Epiphallus, lateral view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure6" httpUri="https://binary.pensoft.net/fig/1103853">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENHAE" type="type material">
<paragraph id="82DFDD2020CFAD06916EFBCD990DC966">
<heading id="66B92AD8830FEF402BCAE2CE556536F0" reason="title">Type material.</heading>
</paragraph>
<paragraph id="95FCFE650F75EBA74CBC2E171D9CE649">
<materialsCitation id="369975D24C4C76ECD4ECBF6EE1D3B6C0" collectingDate="2017-08-07" collectorName="K. Bakolitsas" country="Greece" county="cave Siko" elevation="344" latitude="38.723915" location="Sterea Ellas" longLatPrecision="1" longitude="21.116283" municipality="Agios Nikolaos" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="0556FCEC0D5F404B577CFC6EF366C7B1" bold="true" italics="true">
<typeStatus id="893FF442153798D6A2C30EDB1A8DDBC6" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="3B004BA50D78D4968DD68B8432969BD7" name="Greece">GREECE</collectingCountry>
•
<specimenCount id="3CA40C1D25DAD8298CDE70ABA82C4EFD" count="1" type="male">♂</specimenCount>
;
<location id="D55B6DAE1A4E9EE2309E863C935324E2" LSID="urn:lsid:plazi:treatment:DA4A247D2C9955F28788813AF6E97F17:D55B6DAE1A4E9EE2309E863C935324E2" country="Greece" county="cave Siko" latitude="38.723915" longLatPrecision="1" longitude="21.116283" municipality="Agios Nikolaos" name="Sterea Ellas">Sterea Ellas</location>
,
<location id="4529271D92BBA4B83DE48F420C43C5BB" LSID="urn:lsid:plazi:treatment:DA4A247D2C9955F28788813AF6E97F17:4529271D92BBA4B83DE48F420C43C5BB" country="Greece" county="cave Siko" latitude="38.723915" longLatPrecision="1" longitude="21.116283" municipality="Agios Nikolaos" name="Nomos Etolias-Akarnanias">Nomos Etolias-Akarnanias</location>
,
<location id="64834A829A9AF17ED20307E3333F7FD1" LSID="urn:lsid:plazi:treatment:DA4A247D2C9955F28788813AF6E97F17:64834A829A9AF17ED20307E3333F7FD1" country="Greece" county="cave Siko" latitude="38.723915" longLatPrecision="1" longitude="21.116283" municipality="Agios Nikolaos" name="Aetos">Aetos</location>
,
<collectingMunicipality id="8789517FEB56149FA68BDB8C71DE8DA4">Agios Nikolaos</collectingMunicipality>
,
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;
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<geoCoordinate id="C9C644DD94CC19FABC5DF9CB41756E13" degrees="38" direction="north" minutes="43.435" orientation="latitude" precision="1" value="38.723915">38 ° 43.435 ' N</geoCoordinate>
,
<geoCoordinate id="4FE27D71F945F8A6E14C4C60712C32B6" degrees="21" direction="east" minutes="6.977" orientation="longitude" precision="1" value="21.116283">21 ° 6.977 ' E</geoCoordinate>
</named-content>
;
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<quantity id="AD808AB674E33B15FA6CF24D4BBF608A" metricMagnitude="2" metricUnit="m" metricValue="3.44" unit="m" value="344.0">344 m</quantity>
a. s. l.
</elevation>
;
<date id="F8ECD91AC11946DBC555C6937AE2869C" value="2017-08-07">
<collectingDate id="99E28897AAB8A241C2E71A40DD79C4CE" value="2017-08-07">7 Aug. 2017</collectingDate>
</date>
;
<collectorName id="15DD81089F1FBA4025E1FAAB0A1E4AAD">K. Bakolitsas</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0EAIAE" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
.
<materialsCitation id="1C70BC713693215661D2F7912313407B" collectingDate="2017-08-07" collectorName="K. Bakolitsas" country="Greece" county="cave Siko" elevation="344" latitude="38.723915" location="Sterea Ellas" longLatPrecision="1" longitude="21.116283" municipality="Agios Nikolaos" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="5E44DDEC821E658EEC970CBE0E57B1F1" bold="true" italics="true">
<typeStatus id="6FEAD42ED3C585A52FF5F3FCE3F43D78" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="3B19610AA3A394E4E515237EECD2BA04" name="Greece">GREECE</collectingCountry>
•
<specimenCount id="2F7E491092B338FEC210AAE4ADCD5DCC" count="2" type="male">2 ♂</specimenCount>
, 1 nymph, same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIIAE" type="diagnosis">
<paragraph id="7929F7B8BDBA58BFAB7F0375109727C4">
<heading id="F2F247A79E623D44260DD88132ABBD27" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="19E60C23F57347B1C6F66EB6FDA54206">
The new taxon presents one morphological character unique for the genus: the presence of a notch at the median process of the epiphallus (present at both dissected males). The shape of the lateral lobes of X tergum of the males shows resemblance to
<taxonomicName id="C6321ADCAB20CBACE363FC912D8E03BE" authorityName="Rampini &amp; Di Russo" authorityYear="2017" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="bakolitsasi">
<emphasis id="86C218635926A1BA17D7AABCDA4D866B" italics="true">D. bakolitsasi</emphasis>
</taxonomicName>
, a species known from a cave a few km away. The elevated ridge at the posterior margin of the concavity of the X tergite and the rather slender median process of epiphallus connects the new species with the species of the south Ionian region, e. g.
<taxonomicName id="89632443E8116AD7A9C53E052C6781F1" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="garparoi">
<emphasis id="2855861BE773481155D5C2D6FB66B30D" italics="true">D. garparoi</emphasis>
</taxonomicName>
,
<taxonomicName id="28FFF187A42F9D293BAC3E0C8D1BAEAF" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="ithakii">
<emphasis id="AE292D36795FEFEFA0E2582A7FC1C669" italics="true">D. ithakii</emphasis>
</taxonomicName>
,
<taxonomicName id="6CB1AA62D6C7C338209D1940AE2F0CC3" authorityName="Rampini &amp; Di Russo" authorityYear="2008" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="giachinoi">
<emphasis id="EB6F2DAD611FE5DAC20DE7F7833E8B34" italics="true">D. giachinoi</emphasis>
</taxonomicName>
and
<taxonomicName id="783C9A02BC00746CAC92B8EE8A4F567C" authorityName="Chopard" authorityYear="1964" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="patrizii">
<emphasis id="EED2CE83640F8A463F48A65B5F27C215" italics="true">D. patrizii</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="062460714ACC276F95FC209A4846385A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Geographic distribution of Dolichopoda species at W Sterea Ellas and S Ionian Islands." figureDoi="10.3897/contrib.entomol.74.e121614.figure4" httpUri="https://binary.pensoft.net/fig/1103851">4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJKAE" type="description">
<paragraph id="9B7FC9EB5C7418D6FF164F0B0C6E3DC7">
<heading id="80C77066FF2141F8080D114D125D67B6" reason="title">Description.</heading>
</paragraph>
<paragraph id="535EECA276741B8337383AB79D0B271B">Body pale brownish-yellow, legs elongated, hind femora unarmed. Fore tibiae with 4 spines on both sides of the ventral side, mid tibiae with 5 spines on both sides of the ventral side and hind tibiae with 19 spines on both sides of the dorsal side.</paragraph>
<paragraph id="556CFB0BB3A2B23C067CCCC52699EE7D">
X tergum (last abdominal tergite) (Fig.
<figureCitation id="127BE2985E4253BF638543BCB2B39D60" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dolichopoda karoutsosi sp. nov., male: A. X tergum, dorsal view; B. Epiphallus, dorsal view; C. Epiphallus, lateral view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure6" httpUri="https://binary.pensoft.net/fig/1103853">6 A</figureCitation>
) with two lateral lobes, shallowly bilobed at apex. Bilobed apex very unequal, the external side elongated, cylindrical, pointing dorso-posteriorly. The two lobes are separated by a large concavity showing an elevated ridge on the posterior margin.
</paragraph>
<caption id="9B5E093D36AFF30F28F7CD5AC6689DBE" ID-DOI="10.3897/contrib.entomol.74.e121614.figure6" ID-arpha="289EE310-3C14-5E94-99DC-2262648604E4" httpUri="https://binary.pensoft.net/fig/1103853" startId="F6">
<paragraph id="3055F48FDB05D1F3110D557C91E59563">
<label id="AB9D84C9FB1D80CAA55682B81815DDA6">Figure 6.</label>
</paragraph>
<paragraph id="7F63E763078A297B48B1A718A7399421">
<taxonomicName id="653D0029885598CFE0597F128B902F7E" authorityName="Alexiou &amp; Bakolitsas &amp; Di Russo &amp; Rampini" authorityYear="2024" class="Insecta" family="Rhaphidophoridae" genus="Dolichopoda" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="karoutsosi" status="sp. nov.">
<emphasis id="4499BA87D5419E25AC342F9FAD8E8499" italics="true">Dolichopoda karoutsosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC3666B6ACF3FC68FFBE67A3B81FE768" rank="species">sp. nov.</taxonomicNameLabel>
, male:
<emphasis id="C71ED4297E88A1D655C667CE1BCCBF06" bold="true">A.</emphasis>
X tergum, dorsal view;
<emphasis id="B71C829295572A737712F9C496E09FDC" bold="true">B.</emphasis>
Epiphallus, dorsal view;
<emphasis id="3DDC2E10B81A61D6A2DC931B9D388191" bold="true">C.</emphasis>
Epiphallus, lateral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="D709175F5BD25F7BF2D1456CE1DBC5ED">Subgenital plate convex, with a wide median incision that runs for 1 / 3 of the total length; lateral lobes rounded, styli short, inserted at the side of the lateral lobes.</paragraph>
<paragraph id="2E124FB89F6DC31615F65DD2197BB911">
Epiphallus sclerotized, median process rather slender and acute, the last ¼ curved inwards, an obvious notch is present at the beginning of the curve (Fig.
<figureCitation id="E95660A046B620CFEC856A554D1F6F40" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dolichopoda karoutsosi sp. nov., male: A. X tergum, dorsal view; B. Epiphallus, dorsal view; C. Epiphallus, lateral view. Scale bars: 1 mm." figureDoi="10.3897/contrib.entomol.74.e121614.figure6" httpUri="https://binary.pensoft.net/fig/1103853">6 B, C</figureCitation>
).
</paragraph>
<paragraph id="8F93F48FEEC81930FB94B624E912F8CE">Measurements (length in mm): pronotum 3.6, fore femora 17, middle femora 18, hind femora 26, body 16.</paragraph>
<paragraph id="0E44C3601BF79B3964FEE5A34ADE5D18">
<emphasis id="A1564ABFA6B4AA4032BC3A337A0B31FB" bold="true">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5LAE" type="etymology">
<paragraph id="DB71086C5E40D411DF7CA65AB6B18F3E">
<heading id="C89F960FFD9FDB16BDE261AF5F87439E" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="5BCC2DC7EBC46606523B2D575CA00B8B">
The new species is named after our friend, geologist Panos Karoutsos, a dear companion of the second author who visited the
<typeStatus id="81AEB76D0CD0AEEE4BF9B53C2628AC79">type</typeStatus>
locality and many other caves together.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDMAE" type="distribution">
<paragraph id="FDBD532FD03B5E3394BBFF42F248E3F1">
<heading id="8497B8A4D295DBE434CF1520EF808B47" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="27993E720DDC949B65BCB76761C63F8A">
Known only from the
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locality. Siko is a pit fall
<quantity id="EFC11DAEF9D15B9256C33C7BFD562456" metricMagnitude="0" metricUnit="m" metricValue="7.0" unit="m" value="7.0">7 m</quantity>
deep, with a spacious 8 ×
<quantity id="0B0E3CC2BBF206BB5EF48543D0DA84BF" metricMagnitude="0" metricUnit="m" metricValue="7.0" unit="m" value="7.0">7 m</quantity>
room at its base, without decoration and with high humidity. The entrance is about 0.5 ×
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, facing east and on the floor there are many bones belonging to animals that have fallen from the cave entrance.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:role id="0A576CC6427E18338050F056FA989F1A">
<mods:roleTerm id="967AB5030062097D779CDF91AA821F7D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D4FAD87666B0F6E5D6CD3E781E01098D">Strümpher, Werner P.</mods:namePart>
<mods:nameIdentifier id="DFCA87D6AFC033AE5D60D580991AC482" type="ORCID">0000-0002-7047-2666</mods:nameIdentifier>
<mods:affiliation id="F1BFCEA5EE76644F54A264E92DFBF25E">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa</mods:affiliation>
</mods:name>
<mods:name id="DD2EA6818C63F86B8D4BBA1F73605161" type="personal">
<mods:role id="FF0074A86681E3CCD1CDC91036412673">
<mods:roleTerm id="41C972E7C37AB49CC811732AEDDBB86D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E4BD404B1891F2552180550DD735C540">Barclay, Maxwell V. L.</mods:namePart>
<mods:nameIdentifier id="A28C394E549EE62724AA4B4943CF2374" type="ORCID">0000-0003-4989-2014</mods:nameIdentifier>
<mods:affiliation id="8481C26032A78AC50FDC5DA4D512F379">Department of Life Sciences, Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name id="26646300E81766DE906F67B714A2DDBD" type="personal">
<mods:role id="700D6ABE5C9782B18EB824EA17C5DB96">
<mods:roleTerm id="D19DDCE8FC027E80DAC53D20A262F255">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DFC68AB21D060CD2DA6A8E948C63192D">Vaz-de-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="C59F971CE95EB8CC353F38B402713BB3" type="ORCID">0000-0001-9697-320X</mods:nameIdentifier>
<mods:affiliation id="61EA1AE8AEC3EE079F8EDB0A994DEC20">Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0DD5426181F7ACB1515312701156E1AC">text</mods:typeOfResource>
<mods:relatedItem id="FB8209C7278A31E76AD87E2615EBD098" type="host">
<mods:titleInfo id="D2F3B0A0483D218784857D913AB382C3">
<mods:title id="B422174011276BECEBE866A5F71758E2">Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="0D667D4C81F93F619C222C4C665985C2">
<mods:date id="C562830ED605D22AD4224B5421256612">2024</mods:date>
<mods:detail id="1020E2C6A6EAB77649765BA99B8550B1" type="pubDate">
<mods:number id="E7251272A210BEA35A88D2E810911C7F">2024-07-29</mods:number>
</mods:detail>
<mods:detail id="8492E1A5EC4111984F8E5C9A2B45621B" type="volume">
<mods:number id="719E325E99146429AD99AB934A54311B">74</mods:number>
</mods:detail>
<mods:detail id="B0FE1F06EE078BC2DB71CF4B72F389AA" type="issue">
<mods:number id="4E193B0F8365C3D5DC36E741C4D66C90">1</mods:number>
</mods:detail>
<mods:extent id="21FE1EBDB76138BCAFF1859A87E8BD13" unit="page">
<mods:start id="EF8D1A2752013015FE8C319BBA88813C">81</mods:start>
<mods:end id="1053BE84AAC04ACC0DE941CA5D3D465D">101</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CBECE8301B09EA4AA7F9B4A9972890FE">journal article</mods:classification>
<mods:identifier id="1C01171F0AEAB7888215C66E34C34875" type="DOI">10.3897/contrib.entomol.74.e126799</mods:identifier>
<mods:identifier id="0F382E996FA4A7FDEB03C137F09CFFD9" type="ZooBank">339AB68C-DCB1-435F-B88E-2C42D0A7B05E</mods:identifier>
</mods:mods>
<treatment id="F29208555F635C0DB1C55BEC367E3C94" ID-DOI="http://doi.org/10.5281/zenodo.13123997" ID-Zenodo-Dep="13123997" ID-arpha="F2920855-5F63-5C0D-B1C5-5BEC367E3C94" LSID="urn:lsid:plazi:treatment:F29208555F635C0DB1C55BEC367E3C94" httpUri="http://treatment.plazi.org/id/F29208555F635C0DB1C55BEC367E3C94">
<subSubSection id="B4324F27AF22E3474575D786C7D81EB4" type="nomenclature">
<paragraph id="28248B587AF3CA49FDEE1659F3334A1C">
<taxonomicName id="022067FCCBFD560100B3601C96059EE0" ID-CoL="49DS8" authority="(Vaurie, 1962)" baseAuthorityName="Vaurie" baseAuthorityYear="1962" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="persuberosus" subGenus="Omorgus">
<emphasis id="2EF841978F7F155B9CACA6132F84B7AE" italics="true">Omorgus</emphasis>
(
<emphasis id="D2E2767FB1D6E0A2187E48BEB9584C35" italics="true">Omorgus</emphasis>
)
<emphasis id="A9B3D0589D86207FF5A3AC7AEE4C7DA0" italics="true">persuberosus</emphasis>
(
<bibRefCitation id="43FF4117061F553B43B6DCD85C097EF2" author="Vaurie P" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105 - 167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie, 1962</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D444AA0CE714D3487D31D90E2BFED2BC" type="reference_group">
<paragraph id="7F3C402E8951F7A542F4FD0EB230D57A">
<treatmentCitationGroup id="2EE28A590B383BC26133FE98C3100CA4">
<treatmentCitation id="B9EC976864DCC750D20C9BBAA181D909" author="Vaurie P" page="145" year="1962">
<taxonomicName id="BE4FB7AE3B293DA3BB379B8ED1D7907F" authority="Vaurie, 1962: 145" authorityName="Vaurie" authorityPageNumber="145" authorityYear="1962" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="persuberorus">
<emphasis id="62A92DF268F667D6EF3B11EE4421D7FD" italics="true">Trox persuberorus</emphasis>
<comment id="19B336D88DA7B15832D777BFC027AE58">
<bibRefCitation id="338B32CACFA7C52BF7027F73B1D9C694" author="Vaurie P" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105 - 167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie, 1962: 145</bibRefCitation>
(original description).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="49BD5B1B3BBE567D86DB5C2D0A153C9E">
<treatmentCitationGroup id="4B0E174BE131D0A1D342050288662224">
<treatmentCitation id="71D7D24E0CABEC62209D7E2D0609538B">
<taxonomicName id="12FF43D871B88E2D87AC1B6D66C49AC3" ID-CoL="7D8TZ" authorityName="Vaurie" authorityYear="1962" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="persuberosus" subGenus="Omorgus">
<emphasis id="75D45DA876C60BF644911F91DC31FD1C" italics="true">Trox</emphasis>
(
<emphasis id="7F84246B44736F25507ACAB3060285A9" italics="true">Omorgus</emphasis>
)
<emphasis id="1DE337B7B7B53F87192CEA84EB58890F" italics="true">persuberosus</emphasis>
</taxonomicName>
<comment id="566A8E31F963281FF78266379A4790AF">
:
<treatmentCitation id="4349D1DE269C37B49D98CF9478564A96" author="Scholtz CH" page="12" year="1982">
<bibRefCitation id="518D46186C91A0FBCA7E8FF9C6066242" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa" page="12" pagination="1-27" refId="B96" refString="Scholtz CH (1982) Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries Republic of South Africa 54: 1–27." title="Catalogue of world Trogidae (Coleoptera: Scarabaeoidea)." volume="54" year="1982">Scholtz 1982: 12</bibRefCitation>
</treatmentCitation>
(catalogue).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="576C958FAE9ACEB393F99B6A8A3D3F23">
<treatmentCitationGroup id="2C78165F91174E6086FCC723A7C5E748">
<treatmentCitation id="36714F9A231D7F46E8053063BB3F8AD0">
<taxonomicName id="478739D6C674D04C693B91B17F69DEA8" ID-CoL="49DS8" baseAuthorityName="Vaurie" baseAuthorityYear="1962" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="persuberosus">
<emphasis id="AD2CAB761012783583E3D1600F9768AF" italics="true">Omorgus persuberosus</emphasis>
</taxonomicName>
<comment id="E0845DBF84504BFF85A54FFD8179205B">
:
<treatmentCitation id="42DF8D0F7EE18DC3A5509055BDD4BE1D" author="Deloya C" page="122" year="2005">
<bibRefCitation id="2E8DF63E5515B6590929E19D2E58E9C7" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121–129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="A2E5932FEEDE3E3DF8C9FFB24FD87235" author="Ratcliffe BC &amp; Jameson ML &amp; Figueroa L &amp; Cave RD &amp; Paulsen MJ &amp; Cano EB &amp; Beza-Beza C &amp; Jimenez-Ferbans L &amp; Reyes-Castillo P" page="189" year="2015">
<bibRefCitation id="75B718911ECAFC8D5B509D415F94CAE4" DOI="10.2317/kent-88-02-186-207.1" author="Ratcliffe BC &amp; Jameson ML &amp; Figueroa L &amp; Cave RD &amp; Paulsen MJ &amp; Cano EB &amp; Beza-Beza C &amp; Jimenez-Ferbans L &amp; Reyes-Castillo P" issue="2" journalOrPublisher="Journal of the Kansas Entomological Society" pagination="186-207" refId="B91" refString="Ratcliffe BC, Jameson ML, Figueroa L, Cave RD, Paulsen MJ, Cano EB, Beza-Beza C, Jimenez-Ferbans L, Reyes-Castillo P (2015) Beetles (Coleoptera) of Peru: a survey of the families. Scarabaeoidea. Journal of the Kansas Entomological Society 88 (2): 186 - 207. https: // doi. org / 10.2317 / kent- 88 - 02 - 186 - 207.1" title="Beetles (Coleoptera) of Peru: a survey of the families. Scarabaeoidea." volume="88" year="2015">Ratcliffe 2015: 189</bibRefCitation>
</treatmentCitation>
(checklist from
<collectingCountry id="276D48482178269DF526E4CF734FAFE1" name="Peru">Peru</collectingCountry>
);
<treatmentCitation id="430F7429E2541C8BBE1D268EAB05CC79" author="Casari SA &amp; Biffi G &amp; Ide S" httpUri="http://treatment.plazi.org/id/F29208555F635C0DB1C55BEC367E3C94" page="629" year="2024">
<bibRefCitation id="DB80AA6A892948ABAFD8E363BD0453E2" DOI="10.61818/56330464c31" author="Casari SA &amp; Biffi G &amp; Ide S" editor="Rafael JA &amp; Melo GAR &amp; Carvalho CJB &amp; Casari S &amp; Constantino R" etAl="et al." firstAuthor="Casari" journalOrPublisher="2 ª ed. Instituto Nacional de Pesquisas da Amazônia, Manaus" page="629" pagination="575-698" refId="B20" refString="Casari SA, Biffi G, Ide S (2024) Coleoptera Linnaeus, 1758. In: Rafael JA, Melo GAR, Carvalho CJB, Casari S, Constantino R (Eds) Insetos do Brasil: Diversidade e Taxonomia. 2 ª ed. Instituto Nacional de Pesquisas da Amazônia, Manaus, 575–698. https://doi.org/10.61818/56330464c31" title="Coleoptera Linnaeus, 1758." volumeTitle="Insetos do Brasil: Diversidade e Taxonomia." year="2024">Casari et al. 2024: 629</bibRefCitation>
</treatmentCitation>
(image).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="50C34391535F5C3180986A5673006C46">
<treatmentCitationGroup id="9818AE29C20E06EA4C5FE20060FC965D">
<treatmentCitation id="25C993DA006711D9B268E4F53DA0D6DB">
<taxonomicName id="CE292D5B3396F2F17093FBC699EC1045" ID-CoL="49DS8" baseAuthorityName="Vaurie" baseAuthorityYear="1962" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="persuberosus" subGenus="Omorgus">
<emphasis id="3625A38F72DB99634C058A7F4E41AFAF" italics="true">Omorgus</emphasis>
(
<emphasis id="E1E5B65037AD82FD79FF74A371636A47" italics="true">Omorgus</emphasis>
)
<emphasis id="14FA6C5DD97C540ADE1EA02AF19483B7" italics="true">persuberosus</emphasis>
</taxonomicName>
<comment id="2797768BA59C47041EA07181F8E764BF">
:
<treatmentCitation id="73C9153844F2854C5ED884C7FB228F14" author="Scholtz CH" page="361" year="">
<bibRefCitation id="216A986AF5AD89C8DD8DE09ED38A0A74" DOI="10.1111/j.1365-3113.1986.tb00186.x" author="Scholtz CH" journalOrPublisher="Systematic Entomology" pagination="355-363" refId="B97" refString="Scholtz CH (1986 a) Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology 11: 355–363. https://doi.org/10.1111/j.1365-3113.1986.tb00186.x" title="Phylogeny and systematics of the Trogidae (Coleoptera: Scarabaeoidea)." volume="11">Scholtz 1986 a</bibRefCitation>
: 361
</treatmentCitation>
(phylogenetics);
<treatmentCitation id="FFA0404DD46CDB07422C8D604555FD79" author="Scholtz CH" page="1411" year="1990">
<bibRefCitation id="D18E1F65E1C1AB9365C795E09295C50C" DOI="10.1080/00222939000770841" author="Scholtz CH" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" page="1411" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990: 1411</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="37D9717358635DA4A3A589943E3FE23C" author="Zidek J" page="15" year="2013">
<bibRefCitation id="8BDEE51623EE4285E2008EBDA369B018" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="15" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 1–38." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 15</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="1AD57ED94662125E29D3BD0C7C660AA5" author="Zidek J" page="106" year="2017">
<bibRefCitation id="04F806469AC5D3B24A447C0DEE8C63DB" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="106" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93–127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 106</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="CC225822CFE9B15C92FEDF8C4C043422" author="Smith ABT" page="87" year="2017">
<bibRefCitation id="19E3D0F31435C06ACAA8E9711DB1DEC0" author="Smith ABT" firstAuthor="Smith" journalOrPublisher="Revista Chilena de Entomología" page="87" pagination="81-103" refId="B102" refString="Smith ABT (2017) Southern South American Trogidae (Coleoptera): verification of specimens and notes on geographic distribution. Revista Chilena de Entomología 43: 81–103." title="Southern South American Trogidae (Coleoptera): verification of specimens and notes on geographic distribution." volume="43" year="2017">Smith 2017: 87</bibRefCitation>
</treatmentCitation>
(distribution data);
<treatmentCitation id="03BD6E1CB65AAC774A87E2D842AD02CD" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" httpUri="http://treatment.plazi.org/id/03DA2410FFA2FFF4FE92FB65FE791E84" page="2005" year="2021">
<bibRefCitation id="76C79EFB693E763A25E2099E9526BF79" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="2005" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021: 2005</bibRefCitation>
</treatmentCitation>
(review of Brazilian species).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E52AI" type="material">
<paragraph id="2C44FB3E9FE8679B280E841555BC23D5">
<heading id="EDB4ECD962DCBAFAC545A53DD00AA78E" reason="title">Type specimens examined.</heading>
</paragraph>
<paragraph id="E3E039699A578536CCE5B545FD6C79E9">
<materialsCitation id="6B66BE4532874381FCA93D70E1DE180F" collectionCode="MZSP" country="Brazil" location="See" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="C01DBACC3DBA24A0D318F5FE6F6E0F1D" bold="true" italics="true">
<typeStatus id="72A0FF564BF23693C695DF0D7E7A4816" type="holotype">Holotype</typeStatus>
, by original designation
</emphasis>
(
<specimenCount id="6465AC026C2F80282ED208B3EAFD5105" count="1" type="male">♂</specimenCount>
<named-content id="1B78520AA4D5175D8EDFADCE39E31F9F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/sao-paulo-museu-de-zoologia-da-universidade-de-sao-paulo" xlink_title="Museu de Zoologia da Universidade de São Paulo">
<emphasis id="74F77042F0A11853EC38D658B3BA943C" bold="true">
<collectionCode id="FA92D5A547D39D08B116A96BA07E6444" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" type="Museum">MZSP</collectionCode>
</emphasis>
</named-content>
).
<location id="C691E3ADDBB529DCE48D8E7AC8C069AE" LSID="urn:lsid:plazi:treatment:F29208555F635C0DB1C55BEC367E3C94:C691E3ADDBB529DCE48D8E7AC8C069AE" country="Brazil" name="See" stateProvince="Sao Paulo">See</location>
<bibRefCitation id="E42ABA82EC8349713E4030771FB52A2A" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva V &amp; Strümpher WP &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" page="2003" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. (2021: 2003)</bibRefCitation>
for high-quality images of the type specimen.
<emphasis id="2279864EEE8105000992B08CC6E5B9BB" bold="true">Type locality</emphasis>
: “ Ypiranga,
<collectingRegion id="9EE9CDA7D87A5AB2B0F56AC962DA65F4" country="Brazil" name="Sao Paulo">São Paulo</collectingRegion>
” [
<collectingCountry id="CCFC260782390E258A3107DB8A03635A" name="Brazil">Brazil</collectingCountry>
].
</materialsCitation>
</paragraph>
<paragraph id="F98377FAE2BD07B45964CE04D9B62AB5">
<emphasis id="8FB3E4EFFC8A44238B6B886A7EF29696" bold="true" italics="true">
<typeStatus id="D487AA3733440901B6706313534C838D" type="paratype">Paratypes</typeStatus>
</emphasis>
. Forty specimens from several localities in different institutions (see
<bibRefCitation id="F2C9B9327FA89CEF56EBF9723B3E343B" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" page="146" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105–167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962: 146</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E63AI" type="distribution">
<paragraph id="EC9A6498B3A5CDDB0625971E555539F3">
<heading id="B1F44B7377372182363E0198398A0B7E" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="AF2248D7EF08A397C4F18EDFD29BFB6F">
<collectingCountry id="DF65B5334354979B1C7025D15B6545C9" name="Argentina">Argentina</collectingCountry>
,
<collectingCountry id="5218A82EDBBB02675E6A0814597EFB21" name="Bolivia">Bolivia</collectingCountry>
,
<collectingCountry id="E21D76C68AA4A5A2F961FA309C532AB2" name="Brazil">Brazil</collectingCountry>
,
<collectingCountry id="8558A298CC83741FBB72D11C80106238" name="Paraguay">Paraguay</collectingCountry>
,
<collectingCountry id="02C5D9A8143CD00FFD65A1AF53BD1759" name="Peru">Peru</collectingCountry>
, and
<collectingCountry id="39360342EA907AF1EE09D30F880C2E72" name="Uruguay">Uruguay</collectingCountry>
(
<bibRefCitation id="8A56A6B45A213A50672F558099B08170" author="Vaurie" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="105-167" refId="B110" refString="Vaurie P (1962) A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea). The Bulletin of the American Museum of Natural History 124: 105–167." title="A revision of the genus Trox in South America (Coleoptera: Scarabaeoidea)." volume="124" year="1962">Vaurie 1962</bibRefCitation>
;
<bibRefCitation id="612F8A4613F78003873EBDAFBEF28096" DOI="10.1080/00222939000770841" author="Scholtz" firstAuthor="Scholtz" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391–1456. https://doi.org/10.1080/00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz 1990</bibRefCitation>
;
<bibRefCitation id="3193F703931A50C4F515871361C0E421" DOI="10.1080/00222933.2020.1833999" author="Costa-Silva" etAl="et al." firstAuthor="Costa-Silva" issue="31-32" journalOrPublisher="Journal of Natural History" pagination="1993-2024" refId="B26" refString="Costa-Silva V, Strümpher WP, Vaz-de-Mello FZ (2021) Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae). Journal of Natural History 54 (31–32): 1993–2024. https://doi.org/10.1080/00222933.2020.1833999" title="Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae)." volume="54" year="2021">Costa-Silva et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

144
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@ -0,0 +1,144 @@
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<mods:mods id="A2AA40EBF9B510E0BD115125630795FA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0A698BD94143D166975422477DF7B206">Four new Dolichopoda species from Greece, one Troglophilus new to Greece and new locality records (Orthoptera, Rhaphidophoridae)</mods:title>
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<mods:namePart id="E55DB071A40BAB5AE8DB81F56B88178A">Alexiou, Sotiris</mods:namePart>
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<mods:namePart id="BF4FF836AF41F0339782B9507881D50A">Bakolitsas, Kostas</mods:namePart>
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<mods:affiliation id="035613D4498A2CBCD318929374C6610D">Laboratory of Natural Sciences 3 rd General Lyceum, 301 31 Agrinion, Greece</mods:affiliation>
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<mods:affiliation id="0D8C15FD619A71FA8C04FFFFDE70BE8B">Dipartimento di Biologia e Biotecnologie “ C. Darwin ”, Università di Roma “ La Sapienza ”, Roma, Italy</mods:affiliation>
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<treatment id="FE2DEEC326735818B399EBF4EE2ABCCD" ID-DOI="http://doi.org/10.5281/zenodo.13123944" ID-Zenodo-Dep="13123944" ID-arpha="FE2DEEC3-2673-5818-B399-EBF4EE2ABCCD" LSID="urn:lsid:plazi:treatment:FE2DEEC326735818B399EBF4EE2ABCCD" httpUri="http://treatment.plazi.org/id/FE2DEEC326735818B399EBF4EE2ABCCD">
<subSubSection id="B03A545D3B07EC8EF886D19D098C1ECF" type="nomenclature">
<paragraph id="C964CE9E717FA872EFF6770E6A548D9C">
<emphasis id="CEA66878FDBC3E7ECCE77951540266EB" italics="true">
Τroglophilus
<taxonomicName id="05C0927809543CF16C6C902AC0B87DDA" ID-CoL="59948" authority="Di Russo, Rampini &amp; Cobolli, 2014" authorityName="Di Russo, Rampini &amp; Cobolli" authorityYear="2014" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="zoiai">
zoiai
<bibRefCitation id="C7CD42DB342EF3ECA87F868A81193A45" author="Di Russo C &amp; Rampini M &amp; Cobolli M" journalOrPublisher="Biodiversity Journal" pagination="397-420" refId="B7" refString="Di Russo C, Rampini M, Cobolli M (2014) The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss, 1879. Biodiversity Journal 5: 397 - 420." title="The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss, 1879." volume="5" year="2014">Di Russo, Rampini &amp; Cobolli, 2014</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4ZAE" type="material">
<paragraph id="56F53E140980CC08A2F213FCF0E5D369">
<heading id="96CE24422747C59720C298BBBFA0C230" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="F9DEFDF2C0120954101C9A7912D3DBFE">
<materialsCitation id="87EBB40C0654E35854397A3541E17819" collectingDate="2015-09-06" collectorName="K. Bakolitsas" country="Greece" county="Nomos Phthiotidos" elevation="1260" location="Mt. Iti" municipality="Nomos Phokidos" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sterea Ellas">
<collectingCountry id="E6DAF60F3C4D359CBA3FD975B8066DF1" name="Greece">GREECE</collectingCountry>
-
<emphasis id="D465ACE3098BA62F536411F79C810BAD" bold="true">
<collectingRegion id="A2EE76317FF853311E2C45D2F472F43B">Sterea Ellas</collectingRegion>
</emphasis>
•
<specimenCount id="5410CA42947C2691BD296B4E5EA251E4" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="26A08B485FE9F657529E2D4C69C9F03F" count="1" type="female">1 ♀</specimenCount>
;
<collectingCounty id="1D9883BC1BA02C227791C32E22070CEB">Nomos Phthiotidos</collectingCounty>
/
<collectingMunicipality id="BAADE57DEEB33E0ED2BF49C24C850B9A">Nomos Phokidos</collectingMunicipality>
,
<location id="5DFBC976DB69EA93BF5A375B57E11C4E" LSID="urn:lsid:plazi:treatment:FE2DEEC326735818B399EBF4EE2ABCCD:5DFBC976DB69EA93BF5A375B57E11C4E" country="Greece" county="Nomos Phthiotidos" municipality="Nomos Phokidos" name="Mt. Iti" stateProvince="Sterea Ellas">Mt. Iti</location>
,
<location id="2A203974AAD34DD7C900C9812482541B" LSID="urn:lsid:plazi:treatment:FE2DEEC326735818B399EBF4EE2ABCCD:2A203974AAD34DD7C900C9812482541B" country="Greece" county="Nomos Phthiotidos" municipality="Nomos Phokidos" name="cave Agio Pneuma" stateProvince="Sterea Ellas">cave Agio Pneuma</location>
;
<quantity id="FECAA328E027274ACC859920DC50256B" metricMagnitude="3" metricUnit="m" metricValue="1.26" unit="m" value="1260.0">
<elevation id="5441F8C076EAD9104A0BA1853A832131" metricMagnitude="3" metricUnit="m" metricValue="1.26" unit="m" value="1260.0">1260 m</elevation>
</quantity>
. a. s. l ;
<date id="6DA876E98552F8B07904CEC8E5666E64" value="2015-09-06">
<collectingDate id="7972E5099C9C1FC89D74C6369932B166" value="2015-09-06">6 Sep. 2015</collectingDate>
</date>
;
<collectorName id="091D12EA27F0C6810BEB54629DD83EDB">K. Bakolitsas</collectorName>
leg. (
<abbrev content-type="institution" id="ABBRID0EF1AE" xlink_title="Korinthian Museum of Natural History">KMNH</abbrev>
)
</materialsCitation>
•
<materialsCitation id="886A6DF957D6D327FEFCD1E6C64D2F1F" collectingDate="2014-12-27" collectorName="Kofinas" country="Greece" county="Nomos Phokidas" location="cave Baroutospilia" municipality="Mt. Parnassos" specimenCount="1" stateProvince="Sterea Ellas">
<collectingCounty id="8B22F9D26D0263C17CA5B50889ABAADE">Nomos Phokidas</collectingCounty>
,
<collectingMunicipality id="6618B058FFFAF9E3058633B7557D6BE8">Mt. Parnassos</collectingMunicipality>
,
<location id="1ABADB03227E2ACE77A2B98B20760D33" LSID="urn:lsid:plazi:treatment:FE2DEEC326735818B399EBF4EE2ABCCD:1ABADB03227E2ACE77A2B98B20760D33" country="Greece" county="Nomos Phokidas" municipality="Mt. Parnassos" name="cave Baroutospilia" stateProvince="Sterea Ellas">cave Baroutospilia</location>
;
<date id="61C8783A7CDAC8DB1D8127D9BF9D4354" value="2014-12-27">
<collectingDate id="0C6F84298296288ED8270C4DF03C55C9" value="2014-12-27">27 Dec. 2014</collectingDate>
</date>
; obs. &amp; ph.
<collectorName id="E20B2AE3D882A77E5891BE0B0932CA0E">Kofinas</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK1AE" type="comments">
<paragraph id="0C5F86EA80035C38DF760C774F88DDBB">
<heading id="881C437DDBC8D277E6E38538985DF681" reason="title">Comment.</heading>
</paragraph>
<paragraph id="7301C467250EE1A292C0738C9FB661CA">
A species recently described from Mt. Parnassos,
<collectingRegion id="76CF35EE75F9AB8EC71251D82139CF52" country="Greece" name="Sterea Ellada">Central Greece</collectingRegion>
, present also at Mt. Vardousia and Mt. Giona (
<bibRefCitation id="E73764E16B1E291018F6E9CF526D0819" author="Alexiou" etAl="et al." firstAuthor="Alexiou" journalOrPublisher="Parnassiana Archives" pagination="71-81" refId="B2" refString="Alexiou S, Bakolitsas K, Di Russo C, Latella L, Rampini M, Zacharias S (2014) Biospeleological notes on the caves of the Parnassos, Giona and Vardousia Mountains (Central Greece). Parnassiana Archives 2: 71–81." title="Biospeleological notes on the caves of the Parnassos, Giona and Vardousia Mountains (Central Greece)." volume="2" year="2014">Alexiou et al. 2014</bibRefCitation>
;
<bibRefCitation id="D0672583D208949E0D98FD1186959314" author="Di Russo" etAl="et al." firstAuthor="Di Russo" journalOrPublisher="Biodiversity Journal" pagination="397-420" refId="B7" refString="Di Russo C, Rampini M, Cobolli M (2014) The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss, 1879. Biodiversity Journal 5: 397–420." title="The cave crickets of Greece: a contribution to the study of Southern Balkan Rhaphidophoridae diversity (Orthoptera), with the description of a new species of Troglophilus Krauss, 1879." volume="5" year="2014">Di Russo et al. 2014</bibRefCitation>
).
</paragraph>
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</treatment>
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