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<document id="8290645B39363FF056FA3F5E896113D2" ID-DOI="10.3897/contrib.entomol.74.e126799" ID-ZooBank="339AB68C-DCB1-435F-B88E-2C42D0A7B05E" ID-publisher-id="126799" URI-arpha="B2861E35-EA80-53AF-A4AD-1BA2816CBD9C" URI-zoobank="http://zoobank.org/339AB68C-DCB1-435F-B88E-2C42D0A7B05E" article-type="research-article" checkinTime="1722264657988" checkinUser="pensoft" docAuthor="da Costa-Silva, Vinícius, Strümpher, Werner P., Barclay, Maxwell V. L. &amp; Vaz-de-Mello, Fernando Z." docDate="2024" docId="5EABDD6770CF585E881D66A15D4697CB" docLanguage="en" docName="Contributions to Entomology 74(1): 81-101" docOrigin="Contributions to Entomology 74 (1)" docSource="https://contributions-to-entomology.arphahub.com/article/126799/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Omorgus (Omorgus) fuliginosus" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="101" masterDocId="B2861E35EA8053AFA4AD1BA2816CBD9C" masterDocTitle="An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes" masterLastPageNumber="101" masterPageNumber="81" pageNumber="81" updateTime="1722265173734" updateUser="ExternalLinkService">
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<mods:title id="A30D03AB5600541F7B60CFAD56BE37EC">An illustrated catalogue of South American species of Omorgus Erichson, 1847 (Coleoptera, Trogidae, Omorginae) including a neotype designation and taxonomical changes</mods:title>
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<mods:name id="CAF6BA7FE0093CEF35E67A53BEFB27C9" type="personal">
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<mods:namePart id="E11C2A24731FC6FCF382EE4E77339316">da Costa-Silva, Vinícius</mods:namePart>
<mods:nameIdentifier id="6F6FA5E3202EEEEC33A0953E1F707678" type="ORCID">0000-0002-4556-3793</mods:nameIdentifier>
<mods:affiliation id="D0C9E6DE0E08CA86C55DE71960176FF9">Department of Zoology &amp; Entomology, University of Pretoria, Private Bag X 20, Hatfield, 0028, South Africa &amp; Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="DC58BD585527CB7D4B4A705490186366">Strümpher, Werner P.</mods:namePart>
<mods:nameIdentifier id="C3AD5E90969A3193D10B976CEA3E520E" type="ORCID">0000-0002-7047-2666</mods:nameIdentifier>
<mods:affiliation id="8B60D89943B35EB5E4A6E0811A21FF63">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa</mods:affiliation>
</mods:name>
<mods:name id="A917D5743D3DB1BE06700233D08C25BE" type="personal">
<mods:role id="8F4C394BEDF4F3F44A7F68FEFAF65C99">
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</mods:role>
<mods:namePart id="4FB438C0AD4E7BBDCF1000D1E9767D0C">Barclay, Maxwell V. L.</mods:namePart>
<mods:nameIdentifier id="EC198479CAEC55D80D95F8A6A669423A" type="ORCID">0000-0003-4989-2014</mods:nameIdentifier>
<mods:affiliation id="49DCFF649B11F57984FE9BB32EAFCDC2">Department of Life Sciences, Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name id="AF83D5112C2D3174BAB15AB62CC16AC1" type="personal">
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<mods:roleTerm id="E6FD05BFCA397EEDF606207250905DEE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6132056494F3E87EDF6A57F6A53A3BBE">Vaz-de-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="E51D7DD24A773E4092BD6DEB53F7D4D1" type="ORCID">0000-0001-9697-320X</mods:nameIdentifier>
<mods:affiliation id="8BB00671C55D50BFB8DD52659949F4BC">Laboratório de Scarabaeoidologia, Instituto de Biociências, Universidade Federal de Mato Grosso (UFMT), Cuiabá, Mato Grosso, Brazil</mods:affiliation>
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<mods:title id="D7E9EA4BA9B247F0E42461961743C50B">Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="6B02E155D83979B609A7F505347E961C">
<mods:date id="0F7F98E8F9D16509D25B5AFCAB1D5E90">2024</mods:date>
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<mods:number id="94BF658BA1495652EA1182C71B6D1BC7">2024-07-29</mods:number>
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<mods:number id="A6EBE5F8F4122024A2C309A83C9FD35F">74</mods:number>
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<mods:start id="84A857DD2CE53BD951C380CB58CA1C27">81</mods:start>
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<treatment id="5EABDD6770CF585E881D66A15D4697CB" ID-DOI="http://doi.org/10.5281/zenodo.13124003" ID-Zenodo-Dep="13124003" ID-arpha="5EABDD67-70CF-585E-881D-66A15D4697CB" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB" httpUri="http://treatment.plazi.org/id/5EABDD6770CF585E881D66A15D4697CB">
<subSubSection id="AA8677C473929DA600CE679C75446A26" type="nomenclature">
<paragraph id="14292E5D64582FFB9FE1221EE26AF365">
<taxonomicName id="988ED48D112E19989234EB6128439372" ID-CoL="49DRB" authority="(Robinson, 1941)" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus" subGenus="Omorgus">
<emphasis id="D600B45D9B8164D8EBA9CE6C72C5B732" italics="true">Omorgus</emphasis>
(
<emphasis id="B909FE0FA626C7F663C5BC12B5A8C33F" italics="true">Omorgus</emphasis>
)
<emphasis id="B2E13B34926012655C74B1CE51CB3EE1" italics="true">fuliginosus</emphasis>
(
<bibRefCitation id="5FB702A6B7DF92E6F5CEC1D346B063FF" author="Robinson M" journalOrPublisher="Entomological News" pagination="134-135" refId="B94" refString="Robinson M (1941) A new species of Trox from Texas (Coleoptera: Scarabaeidae). Entomological News 52: 134 - 135." title="A new species of Trox from Texas (Coleoptera: Scarabaeidae)." volume="52" year="1941">Robinson, 1941</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="9F37447530F3CB7BA8EE8668C79276D5" type="description">
<paragraph id="73B11EABFF5D255BE3332BF5CDA7D9DF">
<figureCitation id="886FE705D28190DDE0095B9B42265799" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Trox fuliginosus Robinson, 1941 (now Omorgus fuliginosus). A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Robert Finn (USNM). Length of specimen: 1415 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure11" httpUri="https://binary.pensoft.net/fig/1103873">Fig. 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8993F16DB978B9125AFD93BFF045D519" type="reference_group">
<paragraph id="D803650B1F3B447DC4FB2BDB7C0F5720">
<treatmentCitationGroup id="F779CA53637A9D33C3AA06759C9878F6">
<treatmentCitation id="9152A04ED2CD8D247562AEC1DB82D99B">
<taxonomicName id="6295A037A78A711DBB507382877670CB" ID-CoL="59DFH" authority="Robison, 1941: 134" authorityName="Robinson" authorityPageNumber="134" authorityYear="1941" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus" subGenus="Omorgus">
<emphasis id="9F90AC47089F897E77B20ACAD280AB0F" italics="true">Trox</emphasis>
(
<emphasis id="D6018A49BB028A74DD7F96302B1E0949" italics="true">Omorgus</emphasis>
)
<emphasis id="9E78CA5ACF8463DE29C9D6E9BAAFAD29" italics="true">fuliginosus</emphasis>
<comment id="60DB313E4650289E8FE0AAA6090991F7">
Robison, 1941: 134 (original description);
<treatmentCitation id="B31259DFEC5E691932F807B4DC3DA8ED" author="Blackwelder RE &amp; Blackwelder RM" page="31" year="1948">
<bibRefCitation id="C4C2C8ABBA3E366959A04AFF1B1A0585" author="Blackwelder RE &amp; Blackwelder RM" firstAuthor="Blackwelder" journalOrPublisher="Vernon, New York" page="31" refId="B11" refString="Blackwelder RE, Blackwelder RM (1948) Fifth supplement to the Leng Catalogue of Coleoptera of America, north of Mexico. John D. Sherman; Mt. Vernon, New York, 87 pp." title="Fifth supplement to the Leng Catalogue of Coleoptera of America, north of Mexico. John D. Sherman; Mt." year="1948">Blackwelder and Blackwelder 1948: 31</bibRefCitation>
</treatmentCitation>
(checklist).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="16341BA662E28D9E9EECDC1EF3E58F1A">
<treatmentCitationGroup id="711200A9D18E7C4E65E0E11F47AC5693">
<treatmentCitation id="CD1CEF09A4C2803C0890C71992452AB8">
<taxonomicName id="3082087830595B1630EA33696DBA5EE5" ID-CoL="59DFH" authorityName="Robinson" authorityYear="1941" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="C820EDD87835DC690F4180D9CF935CF1" italics="true">Trox fuliginosus</emphasis>
</taxonomicName>
<comment id="9BFAE624F018E211FA1DA3A61FCCFAC9">
:
<treatmentCitation id="5B8A4C7C12191ECEA531ED94ADF3830B" author="Vaurie P" page="66" year="1955">
<bibRefCitation id="34CAEE766A063328BAB31F766A2A63BD" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" page="66" pagination="1-89" refId="B108" refString="Vaurie P (1955) A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). The Bulletin of the American Museum of Natural History 106: 189." title="A revision of the genus Trox in North America (Coleoptera: Scarabaeidae)." volume="106" year="1955">Vaurie 1955: 66</bibRefCitation>
</treatmentCitation>
(redescription);
<treatmentCitation id="67D3951D3DF2F66B723A864995DB4BEA" author="Vaurie P" page="45" year="1958">
<bibRefCitation id="ADD687B5AEBC86F631156E78113AD368" DOI="10.5962/p.372226" author="Vaurie P" firstAuthor="Vaurie" journalOrPublisher="The Coleopterists Bulletin" page="45" pagination="43-46" refId="B109" refString="Vaurie P (1958) New distribution records of North American Trox (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 12: 4346. https://doi.org/10.5962/p.372226" title="New distribution records of North American Trox (Coleoptera: Scarabaeidae)." volume="12" year="1958">Vaurie 1958: 45</bibRefCitation>
</treatmentCitation>
(distribution); Blackwelder 1973: 35 (checklist);
<treatmentCitation id="144FE2432B8692766EFCC73146F91B58" author="Ratcliffe BC" page="301" year="1978">
<bibRefCitation id="A9262BB4D01D61D41FF45B168B6DBCDB" DOI="10.1590/1809-43921978082299" author="Ratcliffe BC" firstAuthor="Ratcliffe" issue="2" journalOrPublisher="Acta Amazonica" page="301" pagination="299-302" refId="B88" refString="Ratcliffe BC (1978) A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae). Acta Amazonica 8 (2): 299302. https://doi.org/10.1590/1809-43921978082299" title="A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae)." volume="8" year="1978">Ratcliffe 1978: 301</bibRefCitation>
</treatmentCitation>
(distribution).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="42DCF0CEDE4BC102ECBF4513D3A81910">
<treatmentCitationGroup id="BC579E576EFD90D96E351CF15A9D58A9">
<treatmentCitation id="60B02B20336E481DF5CF4768E664087B">
<taxonomicName id="DDAF698ABD22DE0CD4D8CBCC28F1BE3C" ID-CoL="49DRB" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="C291800C30D667C9B1B7780845A0F5A4" italics="true">Omorgus fuliginosus</emphasis>
</taxonomicName>
<comment id="2D55657D75584DD721AE4A2BBB78450F">
:
<treatmentCitation id="95AAE5D44BAD8A69D10B93CE013FAAE2" author="Baker CW" page="43" year="1968">
<bibRefCitation id="1F00563BA2A9C89E0B56D204C6D623B5" DOI="10.5479/si.03629236.279.1" author="Baker CW" firstAuthor="Baker" issue="279" journalOrPublisher="Bulletin of the United States National Museum" page="43" pagination="1-79" refId="B5" refString="Baker CW (1968) Larval taxonomy of the Troginae in North America with notes on biologies and histories (Coleoptera: Scarabaeidae). Bulletin of the United States National Museum 279 (279): 179. https://doi.org/10.5479/si.03629236.279.1" title="Larval taxonomy of the Troginae in North America with notes on biologies and histories (Coleoptera: Scarabaeidae)." volume="279" year="1968">Baker 1968: 43</bibRefCitation>
</treatmentCitation>
(larvae description);
<treatmentCitation id="1BFC92C25DEA721C7D6EB50C31866E4F" author="Deloya C" page="3" year="1992">
<bibRefCitation id="10D061D1DB2FCCB6ED8B4D1B4030331B" DOI="10.21829/azm.1992.49521952" author="Deloya C" firstAuthor="Deloya" journalOrPublisher="Acta Zoologica Mexicana" page="3" pagination="1-13" refId="B29" refString="Deloya C (1992) Necrophilous Scarabaeidae and Trogidae beetles of tropical deciduous forest in Tepexco, Puebla, Mexico. Acta Zoologica Mexicana 52: 113. https://doi.org/10.21829/azm.1992.49521952" title="Necrophilous Scarabaeidae and Trogidae beetles of tropical deciduous forest in Tepexco, Puebla, Mexico." volume="52" year="1992">Deloya 1992: 3</bibRefCitation>
</treatmentCitation>
(natural history);
<treatmentCitation id="DA2387C8FC44CA701133FA8D10EEEA67" author="Deloya C" page="43" year="1996">
<bibRefCitation id="0A8F960DFEACC10A9FBF13AE0261645D" author="Deloya C" firstAuthor="Deloya" journalOrPublisher="Folia Entomologica Mexicana" page="43" pagination="39-54" refId="B30" refString="Deloya C (1996) Los Macro-Coleopteros necrofilos de Tepoztlan, Morelos, Mexico (Scarabaeidae, Trogidae, Silphidae). Folia Entomologica Mexicana 97: 3954." title="Los Macro-Coleopteros necrofilos de Tepoztlan, Morelos, Mexico (Scarabaeidae, Trogidae, Silphidae)." volume="97" year="1996">Deloya 1996: 43</bibRefCitation>
</treatmentCitation>
(natural history);
<treatmentCitation id="F2B314802FB346B660061857B29C7EB9" author="Morón MA" page="91" year="1998">
<bibRefCitation id="7715709B316D87BA4359A15BEC06DACA" DOI="10.21829/azm.1998.75751708" author="Morón MA" journalOrPublisher="Acta Zoologica Mexicana" pagination="73-116" refId="B73" refString="Moron MA (1998) Fauna de Coleoptera Lamellicornia de la region de Tepic, Nayarit, Mexico. Acta Zoologica Mexicana 75: 73 - 116 https: // doi. org / 10.21829 / azm. 1998.75751708." title="Fauna de Coleoptera Lamellicornia de la región de Tepic, Nayarit, Mexico." volume="75" year="1998">Morón et al. 1998: 91</bibRefCitation>
</treatmentCitation>
(distribution);
<treatmentCitation id="1ED4B97531AE12E45AB64776C4E7F7AC" author="Deloya C" page="66" year="2000">
<bibRefCitation id="3E12D0E45BF523708B9828A25C6EF3B5" DOI="10.21829/azm.2000.81811871" author="Deloya C" firstAuthor="Deloya" journalOrPublisher="Acta Zoológica Mexicana" page="66" pagination="63-76" refId="B31" refString="Deloya C (2000) Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia). Acta Zoológica Mexicana 81: 6376. https://doi.org/10.21829/azm.2000.81811871" title="Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia)." volume="81" year="2000">Deloya 2000: 66</bibRefCitation>
</treatmentCitation>
, 73 (checklist);
<treatmentCitation id="375F27797B09F04F8896DE04BAC44027" author="Smith ABT" page="6" year="2003">
<bibRefCitation id="87F09841D854A6D62ACD1FC95A4541DB" author="Smith ABT" firstAuthor="Smith" journalOrPublisher="Electronically published, Lincoln, Nebraska" page="6" refId="B101" refString="Smith ABT (2003) Checklist of the Scarabaeoidea of the Nearctic Realm. Version 3. Electronically published, Lincoln, Nebraska, 74 pp." title="Checklist of the Scarabaeoidea of the Nearctic Realm. Version 3." year="2003">Smith 2003: 6</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="15EA1F37A69D053F22A302A85D613254" author="Deloya C" page="130" year="2003">
<bibRefCitation id="F920DCD97C60EE16E1C86DC0C79467A4" author="Deloya C" editor="Morón MA" firstAuthor="Deloya" journalOrPublisher="Vol II. Barcelona" page="130" pagination="125-133" refId="B32" refString="Deloya C (2003) Familia Trogidae. In: Morón MA (Ed.) Atlas de los escarabajos de México. Vol II. Barcelona, 125133." title="Familia Trogidae." volumeTitle="Atlas de los escarabajos de México." year="2003">Deloya 2003: 130</bibRefCitation>
</treatmentCitation>
(diagnosis and distribution);
<treatmentCitation id="68C69A686B377444103D19E248AB0445" author="Deloya C" page="122" year="2005">
<bibRefCitation id="5B9D5A03C97D71493DD59E7143E2A1B8" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
, 124, 128 (key);
<treatmentCitation id="5248209F2E0B52F9187E162EA1ECAADD" author="Mora-Aguilar EF &amp; Montes de Oca E" page="575" year="2009">
<bibRefCitation id="9B77404E7CDC5568FEB5990BEAA2517D" DOI="10.21829/azm.2009.253662" author="Mora-Aguilar EF &amp; Montes de Oca E" firstAuthor="Mora-Aguilar" issue="3" journalOrPublisher="Acta Zoológica Mexicana (nueva serie)" page="575" pagination="569-588" refId="B69" refString="Mora-Aguilar EF, Montes de Oca E (2009) Escarabajos necrófagos (Coleoptera: Scarabaeidae y Trogidae) de la región central baja de Veracruz, México. Acta Zoológica Mexicana (nueva serie) 25 (3): 569588. https://doi.org/10.21829/azm.2009.253662" title="Escarabajos necrófagos (Coleoptera: Scarabaeidae y Trogidae) de la región central baja de Veracruz, México." volume="25" year="2009">Mora-Aguilar and Montes de Oca 2009: 575</bibRefCitation>
</treatmentCitation>
(distribution).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="4133E2F45F3789249B3F23D4A57A9454">
<treatmentCitationGroup id="ED1E23F98B6AD5E497F97A815D1E71EC">
<treatmentCitation id="6DC094B5C74761FB1D9FED79197CA6E0">
<taxonomicName id="E358A35A887F67C9D01B5A59B25D0413" ID-CoL="49DRB" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus" subGenus="Omorgus">
<emphasis id="450FA151DE885B01CFCED4379C80D009" italics="true">Omorgus</emphasis>
(
<emphasis id="617A642B0A6F5C60CE97DE84F5A28EE6" italics="true">Omorgus</emphasis>
)
<emphasis id="D85F0FECAD9751403241D8F829E5B9FF" italics="true">fuliginosus</emphasis>
</taxonomicName>
<comment id="5C484BBB0636DA11F34FD986252D7658">
:
<treatmentCitation id="7C05F32AC04B9AFF2EDC4D4A2BE1245F" author="Zidek J" page="10" year="2013">
<bibRefCitation id="559DF89A988B03C25BF66DF53589AFD3" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="10" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 138." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 10</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="064892EE877308CC8894008E03C8BB44" author="Zidek J" page="101" year="2017">
<bibRefCitation id="F59CF943BDE01694774CCE0582C34A42" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="101" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 101</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="675C4D262F61C53C0E9CC74629754E75" author="Pablo-Cea JD &amp; Cave RD &amp; Serrano-Peraza FA &amp; Alvarado-Larios R &amp; Deloya C &amp; Serrano-Chicas KA &amp; Alfaro E &amp; Chinchilla-Rodríguez AC &amp; Girón-Segovia D &amp; Noriega JA" page="16" year="2023">
<bibRefCitation id="B96EE450E29B16E25ADEE312491B1955" DOI="10.22201/ib.20078706e.2023.94.5117" author="Pablo-Cea JD &amp; Cave RD &amp; Serrano-Peraza FA &amp; Alvarado-Larios R &amp; Deloya C &amp; Serrano-Chicas KA &amp; Alfaro E &amp; Chinchilla-Rodríguez AC &amp; Girón-Segovia D &amp; Noriega JA" etAl="et al." firstAuthor="Pablo-Cea" page="16" refId="B79" refString="Pablo-Cea JD, Cave RD, Serrano-Peraza FA, Alvarado-Larios R, Deloya C, Serrano-Chicas KA, Alfaro E, Chinchilla-Rodríguez AC, Girón-Segovia D, Noriega JA (2023) Catálogo y atlas de distribución de los Scarabaeoidea (Insecta: Coleoptera) de El Salvador. Revista Mexicana de Biodiversidad 94: e 945117. https://doi.org/10.22201/ib.20078706e.2023.94.5117" year="2023">Pablo-Cea et al. 2023: 16</bibRefCitation>
</treatmentCitation>
(catalogue of
<collectingCountry id="6A4BC205F903AA5C60EFD184A91D8947" name="El Salvador">El Salvador</collectingCountry>
).
</comment>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E91AF09F3847DD5BE5163D1C84035845">
<treatmentCitationGroup id="21CF9227D0E1B6A7CA91A98DBC6A5255">
<treatmentCitation id="1B28C6D80B4FDAB3E31E5E7CF1AEA970" author="Scholtz CH" page="1413" year="1990">
<taxonomicName id="DE6A31F89F7BE066E1B1D81C6535A4C4" ID-CoL="74LV9" authority="Scholtz, 1990: 1413" authorityName="Scholtz" authorityPageNumber="1413" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus" subGenus="Omorgus">
<emphasis id="39A3D2DE75F6081F0711A88F7449A878" italics="true">Omorgus</emphasis>
(
<emphasis id="8E503C37727E1BA340258C4D4B23EC2E" italics="true">Omorgus</emphasis>
)
<emphasis id="F814A9F9F724379623C85AA81F1624FA" italics="true">capillaceus</emphasis>
<comment id="D5410E73A0480168C0665F552B4AAFB7">
<bibRefCitation id="5F45B569DF0D394B1F47C88424281BC4" DOI="10.1080/00222939000770841" author="Scholtz CH" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391 - 1456. https: // doi. org / 10.1080 / 00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz, 1990: 1413</bibRefCitation>
(new synonym):
<treatmentCitation id="59CC80419FD9ADDD97D7F7FC8A61A071" author="Deloya C" page="122" year="2005">
<bibRefCitation id="83DEB05824532C232072F37DA46E3925" author="Deloya C" firstAuthor="Deloya" page="122" refId="B33" refString="Deloya C (2005) Omorgus rodriguezae a new species from Mexico, and a key for the separation of the Central and North America species of the genus (Coleoptera: Trogidae). Folia Entomologica Mexicana 44 (Supl. 1): 121129." year="2005">Deloya 2005: 122</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="CF1EE051A1BBA36187DB3402AFB55692" author="Zidek J" page="8" year="2013">
<bibRefCitation id="CDC5D1E30779CD77B1E82079B04FFC60" author="Zidek J" firstAuthor="Zidek" journalOrPublisher="Insecta Mundi" page="8" pagination="1-38" refId="B115" refString="Zidek J (2013) Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea). Insecta Mundi 314: 138." title="Checklist and bibliography of the Trogidae (Coleoptera: Scarabaeoidea)." volume="314" year="2013">Zidek 2013: 8</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="7FEBBBE659526E65DA4A756EE0061614" author="Zidek J" page="99" year="2017">
<bibRefCitation id="62340DCCF13B6F2D9443015FA7E39E11" author="Zidek J" firstAuthor="Zidek" issue="1" journalOrPublisher="Folia Heyrovskyana" page="99" pagination="93-127" refId="B116" refString="Zidek J (2017) Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea). Folia Heyrovskyana 25 (1): 93127." title="Updated checklist and bibliography of family Trogidae (Coleoptera: Scarabaeoidea)." volume="25" year="2017">Zidek 2017: 99</bibRefCitation>
</treatmentCitation>
(checklist).
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDCDI" type="Type specimen examined">
<paragraph id="A8FA490D5B9E555F91B622B36BEF0E0A">
<heading id="033B5CB0064030A8791A3C8E0E887546" reason="title">Type specimen examined.</heading>
</paragraph>
<paragraph id="C497237C5EB0204EFD923B631FA13309">
<materialsCitation id="F7536AC93FC5802550AEB919961C7766" collectionCode="USNM" collectorName="H. Mittendorf" country="United States of America" location="Sixth" specimenCount="1" specimenCount-male="1" stateProvince="Texas" typeStatus="holotype">
<emphasis id="AE8BA7E92710129A1942006B30966201" bold="true" italics="true">
<typeStatus id="ED4D93327F36D57C28FFA38F509BAB52" type="holotype">Holotype</typeStatus>
, by original designation
</emphasis>
(
<specimenCount id="9697EE6574827F25A98722C847CF97A9" count="1" type="male"></specimenCount>
<emphasis id="3A6C7B21D625791EAC89A014587BD693" bold="true">
<collectionCode id="770A20050779BE417C844D4CD756545B" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</emphasis>
Fig.
<figureCitation id="0CB5A403F195275E5179ED58D75AD4D9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Trox fuliginosus Robinson, 1941 (now Omorgus fuliginosus). A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Robert Finn (USNM). Length of specimen: 1415 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure11" httpUri="https://binary.pensoft.net/fig/1103873">11</figureCitation>
). First label [white aged, typeset]: “ New Braunfels / IV. 10 02 Tex ”. Second label [white aged, typeset]: “
<collectorName id="CB46CA0CA779C50668F63E5A53BA6513">H. Mittendorf</collectorName>
/ Collector ”. Third label [red, printed]: “ Type No / 55468 / USNM ”. Fourth label [red, Mark Robinsons handwriting]: “
<typeStatus id="7AF7928EB9A48762C2C1F9D4E124F885" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="7141587E841E097D7ED6680D9A935E83" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="5B834B6B71D17962336046C48146E863" italics="true">Trox</emphasis>
</taxonomicName>
/
<taxonomicName id="2CF313D43A80EDCC280F437EE44D84BF" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="418426C5A6B6CBF3729B53BD3D1335DC" italics="true">fuliginosus</emphasis>
</taxonomicName>
/ Mark Robinson ”.
<location id="B80FAA717C7E776FCC5081025B0CD742" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:B80FAA717C7E776FCC5081025B0CD742" country="United States of America" name="Sixth" stateProvince="Texas">Sixth</location>
label [white, printed]: “ USNMENT / [QR Code] / 01474112 ” (Fig.
<figureCitation id="48BD05307ED7ED56A60EDB93B75EF080" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Holotype of Trox fuliginosus Robinson, 1941 (now Omorgus fuliginosus). A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Robert Finn (USNM). Length of specimen: 1415 mm." figureDoi="10.3897/contrib.entomol.74.e126799.figure11" httpUri="https://binary.pensoft.net/fig/1103873">11 C</figureCitation>
).
<emphasis id="07522E0CA1A3A84352DD797D9076F291" bold="true">Type locality</emphasis>
: “ New Braunfels ” [
<collectingRegion id="10506A673E9F14A2CCBBF2B312D54860" country="United States of America" name="Texas">Texas</collectingRegion>
,
<collectingCountry id="6B961C8B2BE6DB7F5CFC2C3844C861C6" name="United States of America">United States</collectingCountry>
].
</materialsCitation>
</paragraph>
<caption id="F373B4943A79520D80A01D58F8EB89D2" ID-DOI="10.3897/contrib.entomol.74.e126799.figure11" ID-arpha="501E6441-5C5E-55B5-ACF7-6A376FA85A20" httpUri="https://binary.pensoft.net/fig/1103873" startId="F11">
<paragraph id="80E5A2120E2B6BDC056EBF1D228F0BBC">
<label id="2EABC1A37259379E41ED521CE091EC3A">Figure 11.</label>
</paragraph>
<paragraph id="D0AD4436B835411D2A1384E652AEAD72">
Holotype of
<taxonomicName id="8D16551ABB2BA8FFBB2E1037501A2B4F" authority="Robinson, 1941" authorityName="Robinson" authorityYear="1941" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="E459BB307AE4A5CFF1D9D3340A558216" italics="true">Trox fuliginosus</emphasis>
<bibRefCitation id="DC11276DD551FA3502609B73553A6EEB" author="Robinson M" journalOrPublisher="Entomological News" pagination="134-135" refId="B94" refString="Robinson M (1941) A new species of Trox from Texas (Coleoptera: Scarabaeidae). Entomological News 52: 134 - 135." title="A new species of Trox from Texas (Coleoptera: Scarabaeidae)." volume="52" year="1941">Robinson, 1941</bibRefCitation>
</taxonomicName>
(now
<taxonomicName id="22024A3DDDABB2F4D4352BF43F86D5F8" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="8FC446D3B77E730C2FC7E11EB189A592" italics="true">Omorgus fuliginosus</emphasis>
</taxonomicName>
).
<emphasis id="32B66BDFA8D2B0A78E19C5481E0EFD8F" bold="true">A.</emphasis>
Habitus in dorsal view;
<emphasis id="C8234EBC7316D0805370C53C3C32D69A" bold="true">B.</emphasis>
Habitus in lateral view;
<emphasis id="924BAAEFD2DF99ADD18F1F4A61431328" bold="true">C.</emphasis>
Labels. Photos by Robert Finn (USNM). Length of specimen: 1415 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ESEDI" type="Type specimen">
<paragraph id="085CA7F9DF1F5720CC265FD97D92F357">
<heading id="5F1D01B51940F5F0CFE9DEFC3E933BD6" reason="title">Type specimen</heading>
</paragraph>
<paragraph id="8DF07285A4D2BAB5FBAB34EC6465009B">
<emphasis id="CEFF01AD41AA7DA97000656C7C232335" bold="true">
of
<taxonomicName id="F2007180F79436EB420A96F1988E0616" authority="Scholtz, 1990" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus" subGenus="Omorgus">
<emphasis id="E6A52333762217A00E55DDA42BE68F5B" italics="true">Omorgus</emphasis>
(
<emphasis id="0DD6FF2B872951EA744D2B4D8D6E1D43" italics="true">Omorgus</emphasis>
)
<emphasis id="9A8818486178091EAB0482E0464E3627" italics="true">capillaceus</emphasis>
<bibRefCitation id="8F5491FA72DFE58F8DB0A41DC2DFC2D1" DOI="10.1080/00222939000770841" author="Scholtz CH" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391 - 1456. https: // doi. org / 10.1080 / 00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz, 1990</bibRefCitation>
</taxonomicName>
(new synonym) examined.
<materialsCitation id="48DC9176FDD71CF6F7FF7E21F1E18396" collectionCode="CMNCH" collectorName="S. Amer. &amp; C. H. Scholtz" country="Colombia" location="Fourth" specimenCount="1" specimenCount-female="1" stateProvince="Bogota" typeStatus="holotype">
<emphasis id="B8368ECDD369205D131F2EC09FDE9083" bold="true" italics="true">
<typeStatus id="883021CEA95F686A314A23DD8113B966" type="holotype">Holotype</typeStatus>
, by original designation
</emphasis>
(
<specimenCount id="F7BFF917E741CE8881CEDF44C0A7B0CD" count="1" type="female"></specimenCount>
<named-content id="E7AFA3FEE5D06D65D24DC045143D33D0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<emphasis id="4B9D5A5FD562D3DEBFED861EF71B728E" bold="true">
<collectionCode id="B4820C5A3ADBCEF65DA7577259B4BA8E">CMNCH</collectionCode>
</emphasis>
</named-content>
Fig.
<figureCitation id="4C8874A3288CC6B97908EED735477410" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Holotype of Omorgus capillaceus Scholtz, 1990. A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Vanessa Verdecia (CMNCH)." figureDoi="10.3897/contrib.entomol.74.e126799.figure12" httpUri="https://binary.pensoft.net/fig/1103874">12</figureCitation>
). First label [white aged, unknowns handwriting]: “ Botoga /
<collectingCountry id="6D58672C89B9C3EA4311378C193D8008" name="Colombia">Colombia</collectingCountry>
/
<collectorName id="EDAAB97787EE26D64EA1353760990B10">S. Amer.</collectorName>
”. Second label [white, typeset]: “ Carn. Mus. / Acc. 2275 ”. Third label [white, printed]: “ [QR code] /
<named-content id="73A5D7CBAF0F1877C4E4D3BDA1B0F6ED" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="30BC601B32B894F0CD33ADAABF1F6FBC">CMNCH</collectionCode>
</named-content>
- IZ / 724,596 ”.
<location id="2DA896EAAC6A899D35099C1875FFEA27" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:2DA896EAAC6A899D35099C1875FFEA27" country="Colombia" name="Fourth" stateProvince="Bogota">Fourth</location>
label [white with red border, Clark Scholtz`s handwriting]: “
<taxonomicName id="B8D09DB0365514C37336F871ACDCBE37" authorityName="Erichson" authorityYear="1847" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="DF0C1F6D72D3180226A7F721C81C0291" italics="true">Omorgus</emphasis>
</taxonomicName>
/
<taxonomicName id="0F70278FEFCDA2EACAEE63895081BB14" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="2D9C2B86986F1E1A9A845621FF11BE0C" italics="true">capillaceus</emphasis>
</taxonomicName>
/
<collectorName id="D2412C0F3B0FFF255E2F01E915CCCC65">C. H. Scholtz</collectorName>
1988 /
<typeStatus id="0D8031F404283168C7C07BBDCD668B08" type="holotype">HOLOTYPE</typeStatus>
” (Fig.
<figureCitation id="5ACD02FE10938FEBCE598637C1610CAF" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Holotype of Omorgus capillaceus Scholtz, 1990. A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Vanessa Verdecia (CMNCH)." figureDoi="10.3897/contrib.entomol.74.e126799.figure12" httpUri="https://binary.pensoft.net/fig/1103874">12 C</figureCitation>
).
<emphasis id="6DE3B9795723AC4D336A712A6240B8F9" bold="true">Type locality</emphasis>
: “
<collectingRegion id="6B41E2B77482EDB0B33D7C8C48911669" country="Colombia" name="Bogota">Bogota</collectingRegion>
,
<collectingCountry id="C64ED4442F71B1345E8548FB2957A5E7" name="Colombia">Colombia</collectingCountry>
”.
</materialsCitation>
</emphasis>
</paragraph>
<caption id="068F04ADF8231B00E5C3222E7BAD63DA" ID-DOI="10.3897/contrib.entomol.74.e126799.figure12" ID-arpha="C4BA82C9-09BB-5DA5-904B-9D35C6AC0731" httpUri="https://binary.pensoft.net/fig/1103874" startId="F12">
<paragraph id="D1FD68A640EB877329036A85321D5940">
<label id="582B1C0C57DF10D4D7412ACCDC6A407F">Figure 12.</label>
</paragraph>
<paragraph id="40AC9FDE60642CA4B8F1CAB0DCDC5458">
Holotype of
<taxonomicName id="A07ADB579B7FFE6B2B355DDAFEA40CCA" authority="Scholtz, 1990" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="CD4311521E25F548FCB3A553427B483F" italics="true">Omorgus capillaceus</emphasis>
<bibRefCitation id="EB982361ABF609226DA2FAC292BE8ADA" DOI="10.1080/00222939000770841" author="Scholtz CH" journalOrPublisher="Journal of Natural History" pagination="1391-1456" refId="B99" refString="Scholtz CH (1990) Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391 - 1456. https: // doi. org / 10.1080 / 00222939000770841" title="Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea)." volume="24" year="1990">Scholtz, 1990</bibRefCitation>
</taxonomicName>
.
<emphasis id="07E018B19EA5812E8F993C37C331AAAB" bold="true">A.</emphasis>
Habitus in dorsal view;
<emphasis id="C647F0F4146EC6B950A5270E838DE2C5" bold="true">B.</emphasis>
Habitus in lateral view;
<emphasis id="1AD08413B9E1EBFA910371D625C88788" bold="true">C.</emphasis>
Labels. Photos by Vanessa Verdecia (
<named-content id="B3153F2F62DA83887D1B25C7BFAF9233" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">CMNCH</named-content>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EOHDI" type="remarks">
<paragraph id="F599E736ABF2F06559A481CBC2ECF660">
<heading id="A63428702B1081FCEF062A921B32E19B" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="A3696EC91D1EE41DBCFB275617CFA775">
<taxonomicName id="7A21DE2DCF82030BE386BA5BB1F0147D" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus" subGenus="Omorgus">
<emphasis id="23164DF7CA61E242B2D44B05B7AC2301" italics="true">Omorgus</emphasis>
(
<emphasis id="C3AB6A93E3FD3E82E5684421682D5AF8" italics="true">Omorgus</emphasis>
)
<emphasis id="7F9863CADF83C7F829F066769686470B" italics="true">capillaceus</emphasis>
</taxonomicName>
was one of the species described by Scholtz in 1990 during his monographic revision of the
<taxonomicName id="0CDA72476D6DF1759F4EC58A52A9D7CA" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
of South America. This species was identified in the collection of the Carnegie Museum of Natural History (
<named-content id="98595F500B6BE4EEFF59FD9DDDE95B28" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="00AC3575AEEEB6BFAC4329EE496BEABB">CMNCH</collectionCode>
</named-content>
) and described based on a single female specimen, distinguished by its morphological differences from other South American species. Scholtz noted that
<taxonomicName id="E4B7DFB8FB808F99C13AA759D032E1D0" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="2E07CACB1CCF2E531D346A76237696AA" italics="true">O. capillaceus</emphasis>
</taxonomicName>
was similar to
<taxonomicName id="0B516331FED6E2BA05268ABFD759E50D" authority="(Herbst, 1790)" baseAuthorityName="Herbst" baseAuthorityYear="1790" class="Insecta" family="Trogidae" genus="Trox" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="monachus">
<emphasis id="660E2A9C35E9D68D17F38D970D961E23" italics="true">O. monachus</emphasis>
(
<bibRefCitation id="A45F1A4157376AFDDBB901C0655BA6A8" author="Herbst JFW" refId="B52" refString="Herbst JFW (1790) [Hefte 1: 1 - 151, 6 pls.]. In: Herbst JFW (Ed.) 1790 - 1791. Natursystem aller bekannten in- und auslandischen Insekten, als eine Forsetzung der von Buffonschen Naturgeschichte. Der Kafer Dritter Theil. Berlin: Joachim Pauli; p. xiv + 325 pp. + pls. 21 - 34. D - E [publication date following Bousquet 2016]." year="1790">Herbst, 1790</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="8B34402EACB0F1D950C9FFEC05112ABA" authority="(Robinson, 1941)" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="3F2BC18FD139F87770D5950293D778E8" italics="true">O. fuliginosus</emphasis>
(
<bibRefCitation id="8AC3596EDCAFC7CF65FD0E9ED53704E1" author="Robinson M" journalOrPublisher="Entomological News" pagination="134-135" refId="B94" refString="Robinson M (1941) A new species of Trox from Texas (Coleoptera: Scarabaeidae). Entomological News 52: 134 - 135." title="A new species of Trox from Texas (Coleoptera: Scarabaeidae)." volume="52" year="1941">Robinson, 1941</bibRefCitation>
)
</taxonomicName>
, both of which are North American species. However, he highlighted a diagnostic feature that distinguishes
<taxonomicName id="2EF4DF044ADBC6B439041B5F6A7D79C4" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="98721404D7FAB6E01F6708DAEAF46132" italics="true">O. capillaceus</emphasis>
</taxonomicName>
from other similar-looking species: the absence of a velutinous covering on the elytral disc, which is present in its relatives.
</paragraph>
<paragraph id="852990D6BF2D38B13B487A94558E2DC6">
Upon reviewing the
<typeStatus id="8CF5EBC2BA6A7DEAA461B7828DDAB522" type="holotype">holotype</typeStatus>
of
<taxonomicName id="9865ADA5272FD4113069D6F90D32CC27" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="D4C2DC64460109E2012692029171815A" italics="true">O. capillaceus</emphasis>
</taxonomicName>
, it became evident that the specimen is merely a worn variation of
<taxonomicName id="986BD6DADD87C05A1E1ADF02FC08D914" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="44421D089AE9D901AFE279088BECF7B7" italics="true">O. fuliginosus</emphasis>
</taxonomicName>
. While we concur with Scholtzs observation of the glabrous elytral disc, this feature alone is insufficient for distinguishing
<taxonomicName id="0482BBAC516781DE136F3D447287D0A6" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
species without considering male genitalia.
<taxonomicName id="68FB1733E408148D6F2D35DADAF91C2C" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
specimens often exhibit significant intraspecific variation, and it is not uncommon to encounter specimens that are partially or entirely glabrous (or worn) in collections (V. Costa-Silva, personal observation).
</paragraph>
<paragraph id="F87DD5CFE1E26860D82174AE490144CF">
Another factor that may have led Scholtz to describe a new species is the specimens locality:
<collectingRegion id="AE2E4BC114DEF1698E36DB9D47342147" country="Colombia" name="Bogota">Bogota</collectingRegion>
,
<collectingCountry id="6871F1A2B6FAA344A3267025946F5229" name="Colombia">Colombia</collectingCountry>
. The
<typeStatus id="7C289DA8D6E29C9C3282EAB806343FDC" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3DC14AC5C9891BD893F5CC2454FBEB5C" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="A9155788D26B558C5EC5689A8E426E1D" italics="true">O. capillaceus</emphasis>
</taxonomicName>
was associated with the label number “ 2275 ” (Fig.
<figureCitation id="A30672AEFDA736B3D80C066624DBBC8C" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Holotype of Omorgus capillaceus Scholtz, 1990. A. Habitus in dorsal view; B. Habitus in lateral view; C. Labels. Photos by Vanessa Verdecia (CMNCH)." figureDoi="10.3897/contrib.entomol.74.e126799.figure12" httpUri="https://binary.pensoft.net/fig/1103874">12 C</figureCitation>
), which corresponds to specimen from the Henry Klages Collection, acquired by the
<named-content id="E0C8FD4A64F7DE0A61DCB1253E035AD8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="3277DBCB682A4F31965C4E2AFBCB6067">CMNCH</collectionCode>
</named-content>
on
<date id="6B2E1182D3507FF83EAE8320D0C46CB9" value="1903-05-28">May 28, 1903</date>
(Robert Androw, personal communication to
<collectionCode id="46750F8B1A7F108F413FA51A793D8869">VCS</collectionCode>
). However, as documented by Nearns and Androw (2013) and
<bibRefCitation id="D8BF995EFE0E8B7EDCE84F8ED2AD0DD8" DOI="10.1080/00222933.2023.2260060" author="Costa-Silva V &amp; Strümpher WP &amp; Thyssen PJ &amp; Vaz-de-Mello FZ" etAl="et al." firstAuthor="Costa-Silva" issue="1-4" journalOrPublisher="Journal of Natural History" pagination="14-166" refId="B27" refString="Costa-Silva V, Strümpher WP, Thyssen PJ, Vaz-de-Mello FZ (2024) Taxonomic revision of the South American genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae). Journal of Natural History 58 (14): 14166. https://doi.org/10.1080/00222933.2023.2260060" title="Taxonomic revision of the South American genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae)." volume="58" year="2024">Costa-Silva et al. (2024)</bibRefCitation>
, the Henry Klages collection has been noted for containing numerous mislabelled Nearctic and Neotropical specimens, including some cases of misidentification within the
<taxonomicName id="D544F5A20FAA36C48A93D96848D4FCB2" authorityName="MacLeay" authorityYear="1819" class="Insecta" family="Trogidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="family">Trogidae</taxonomicName>
(see
<bibRefCitation id="BA6E8BBF1F9403337CB85D301A69C714" DOI="10.1080/00222933.2023.2260060" author="Costa-Silva" etAl="et al." firstAuthor="Costa-Silva" issue="1-4" journalOrPublisher="Journal of Natural History" pagination="14-166" refId="B27" refString="Costa-Silva V, Strümpher WP, Thyssen PJ, Vaz-de-Mello FZ (2024) Taxonomic revision of the South American genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae). Journal of Natural History 58 (14): 14166. https://doi.org/10.1080/00222933.2023.2260060" title="Taxonomic revision of the South American genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae)." volume="58" year="2024">Costa-Silva et al. 2024</bibRefCitation>
for examples).
<taxonomicName id="72B1051E63F7E3E1C462A88809402D4B" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="F1AD4BDA4664E26A31FD3DB91DC3E0A6" italics="true">Omorgus fuliginosus</emphasis>
</taxonomicName>
is known from the
<collectingCountry id="D8F07AF3D92805EFAF799B79DA0A231B" name="United States of America">United States</collectingCountry>
,
<collectingCountry id="18AEEB233088EA723093943EDB195ABA" name="Mexico">Mexico</collectingCountry>
,
<collectingCountry id="E22403128B46AEF8DF17B0C2D894B5C8" name="Guatemala">Guatemala</collectingCountry>
,
<collectingCountry id="AFBA156B066F24DB051DBC1C22601055" name="El Salvador">El Salvador</collectingCountry>
, and
<collectingCountry id="619C99181F10885E917FD0A3FEDE62D1" name="Costa Rica">Costa Rica</collectingCountry>
(
<bibRefCitation id="8147639BCB7EA952A9D4DA3E99BD4B14" author="Vaurie" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="1-89" refId="B108" refString="Vaurie P (1955) A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). The Bulletin of the American Museum of Natural History 106: 189." title="A revision of the genus Trox in North America (Coleoptera: Scarabaeidae)." volume="106" year="1955">Vaurie 1955</bibRefCitation>
;
<bibRefCitation id="0F2205092994A9C33D9148D88B699D48" DOI="10.1590/1809-43921978082299" author="Ratcliffe" firstAuthor="Ratcliffe" issue="2" journalOrPublisher="Acta Amazonica" pagination="299-302" refId="B88" refString="Ratcliffe BC (1978) A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae). Acta Amazonica 8 (2): 299302. https://doi.org/10.1590/1809-43921978082299" title="A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae)." volume="8" year="1978">Ratcliffe 1978</bibRefCitation>
;
<bibRefCitation id="D85FD5DBE127594B8A7E6595B04A2672" DOI="10.21829/azm.2000.81811871" author="Deloya" firstAuthor="Deloya" journalOrPublisher="Acta Zoológica Mexicana" pagination="63-76" refId="B31" refString="Deloya C (2000) Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia). Acta Zoológica Mexicana 81: 6376. https://doi.org/10.21829/azm.2000.81811871" title="Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia)." volume="81" year="2000">Deloya 2000</bibRefCitation>
), making its record in
<collectingCountry id="5FD227E750995849612A577B86E5D1CD" name="Colombia">Colombia</collectingCountry>
questionable, but not impossible.
</paragraph>
<paragraph id="65442283DD62A1C46D8BE36FF2D36A5E">
Therefore, we conclude that
<taxonomicName id="26B1F2C929E3EF537CC47F08E9C2AC0E" authorityName="Scholtz" authorityYear="1990" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="capillaceus">
<emphasis id="4F4C8D36E8CC41B10317139F4D14EEAD" italics="true">O. capillaceus</emphasis>
</taxonomicName>
should be considered a junior subjective synonym of
<taxonomicName id="7D7D33F1995E459280A8EEF5D3E89011" baseAuthorityName="Robinson" baseAuthorityYear="1941" class="Insecta" family="Trogidae" genus="Omorgus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="DCE33F6A6DA30BB380C459A4544C4A16" italics="true">O. fuliginosus</emphasis>
</taxonomicName>
, having been described based on a worn and mislabelled specimen from
<collectingRegion id="2089C1A901B7BB8DEA6DB2D7E9EB0F49" country="Colombia" name="Bogota">Bogotá</collectingRegion>
,
<collectingCountry id="E7A495306FD6A7F5019E16B36F2F53A7" name="Colombia">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKNDI" type="distribution">
<paragraph id="004672A8451E0A1D06BBD61127EAD9A0">
<heading id="4C71708D5B20ECCD0C4EE97D38E38333" reason="title">Geographic distribution.</heading>
</paragraph>
<paragraph id="E04EDA7A1D6C1229FF55C303F0AAFBC2">
<collectingCountry id="D916E9F61666CDD870BF0906EADA23B6" name="Costa Rica">Costa Rica</collectingCountry>
,
<collectingCountry id="5D96742922CAFCA2392C86299CCFBDDE" name="El Salvador">El Salvador</collectingCountry>
,
<collectingCountry id="66000913A028F66DB44DB714A50795F9" name="Guatemala">Guatemala</collectingCountry>
,
<collectingCountry id="EAD8CA554E2E89C22934EE5F53B01BAD" name="Mexico">Mexico</collectingCountry>
, and
<collectingCountry id="0BC252DE50D033074241FB82A2C8AAC5" name="United States of America">United States</collectingCountry>
(
<bibRefCitation id="FEE08A355946DBED1BCF3036EF8F68FC" author="Vaurie" firstAuthor="Vaurie" journalOrPublisher="The Bulletin of the American Museum of Natural History" pagination="1-89" refId="B108" refString="Vaurie P (1955) A revision of the genus Trox in North America (Coleoptera: Scarabaeidae). The Bulletin of the American Museum of Natural History 106: 189." title="A revision of the genus Trox in North America (Coleoptera: Scarabaeidae)." volume="106" year="1955">Vaurie 1955</bibRefCitation>
;
<bibRefCitation id="91CF2C5CDFD10B0BE1E57DF1C2C9C731" DOI="10.1590/1809-43921978082299" author="Ratcliffe" firstAuthor="Ratcliffe" issue="2" journalOrPublisher="Acta Amazonica" pagination="299-302" refId="B88" refString="Ratcliffe BC (1978) A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae). Acta Amazonica 8 (2): 299302. https://doi.org/10.1590/1809-43921978082299" title="A new species of Trox from the Amazon Basin with new distributional records for Central American Trox (Coleoptera: Scarabaeidae)." volume="8" year="1978">Ratcliffe 1978</bibRefCitation>
;
<bibRefCitation id="96E2C707C6FF3A869C815FFD64E5B41E" DOI="10.21829/azm.2000.81811871" author="Deloya" firstAuthor="Deloya" journalOrPublisher="Acta Zoológica Mexicana" pagination="63-76" refId="B31" refString="Deloya C (2000) Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia). Acta Zoológica Mexicana 81: 6376. https://doi.org/10.21829/azm.2000.81811871" title="Distribucion de la familia Trogidae en México (Coleoptera Lamellicornia)." volume="81" year="2000">Deloya 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3NDI" type="type material">
<paragraph id="FA547A1C25E107F9BC1411A48B00C47F">
<heading id="A17C5D3483139778E7DC6AF4A38A6B5C" reason="title">Non-type examined material</heading>
</paragraph>
<paragraph id="1E0964619D34288EEF9A1E7FCE07D272">
<emphasis id="6DC4655556F8296CACDB73B73BCB7994" bold="true">
(
<specimenCount id="1409DE641938E8EA40D593D777DD1B0A" count="9" type="generic">9 specimens</specimenCount>
).
<materialsCitation id="58F0800151F8174B490E7645D00EA025" country="Mexico" location="Veracruz" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<collectingCountry id="18D0C04E1C931C585C0E93AA15DE4776" name="Mexico">MEXICO</collectingCountry>
<emphasis id="369E7A3F1D399C31C37870B7F989274F" bold="true">
<collectingRegion id="C3704BD4BC0590F68807EDC13D741799" country="Mexico" name="Veracruz de Ignacio de la Llave">Veracruz</collectingRegion>
</emphasis>
</materialsCitation>
</emphasis>
<materialsCitation id="3AC181AF02E4638C9B92B5931660FC71" collectingDate="1994-08-30" collectionCode="CEMT" collectorName="L. Arellano &amp; R. Sanchez" country="Mexico" location="Passando la Desv. Idolos" municipality="Carr. Actopan" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
5;
<collectingMunicipality id="58D12BB5986986F0D15E0987F1990FC3">Carr. Actopan</collectingMunicipality>
,
<location id="23D4E8AA15A901E4B621035136F83345" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:23D4E8AA15A901E4B621035136F83345" country="Mexico" municipality="Carr. Actopan" name="Passando la Desv. Idolos" stateProvince="Veracruz de Ignacio de la Llave">Passando la Desv. Idolos</location>
;
<date id="1E0AA59A72749628AAD00635B5BB656C" value="1994-08-30">
<collectingDate id="81D59ADDADBE3ED0740A86E5C215F030" value="1994-08-30">30 Aug. 1994</collectingDate>
</date>
;
<collectorName id="11F811ACD17F2E87B66DF45AC0F29252">L. Arellano</collectorName>
&amp;
<collectorName id="E7BF0881F9D67C25FAFB06DA37FA0FAD">R. Sanchez</collectorName>
leg.;
<abbrev id="ABBRID0EGODI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="AD4B91A35C9EA48580165375F0272C50">CEMT</collectionCode>
</abbrev>
</materialsCitation>
<materialsCitation id="58AC5736FE238FEDD6C20B039249C2E7" collectingDate="1994-10-18" collectionCode="CEMT" collectorName="L. Arellano &amp; R. Sanchez" country="Mexico" county="Carr. Alto Lucero" location="Desviacion la Concepcion" municipality="Km" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
1;
<collectingCounty id="00B17659746547B0EB20736068AC0461">Carr. Alto Lucero</collectingCounty>
,
<collectingMunicipality id="F7ED1E4D25D6E3ACA076F0093408FFDC">Km</collectingMunicipality>
2
<location id="FD24C7D6801129C3E25CEE6CB173E88D" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:FD24C7D6801129C3E25CEE6CB173E88D" country="Mexico" county="Carr. Alto Lucero" municipality="Km" name="Desviacion la Concepcion" stateProvince="Veracruz de Ignacio de la Llave">Desviación la Concepción</location>
;
<date id="1192A72B218FF6018E0CD412FD4F21B1" value="1994-10-18">
<collectingDate id="037387E2C7007F1045F541C995CCEE25" value="1994-10-18">18 Oct. 1994</collectingDate>
</date>
;
<collectorName id="58FF8F026F564A3E1E158BBDF02F1543">L. Arellano</collectorName>
&amp;
<collectorName id="A11A3BA9E2ED84F323E749801FC352CF">R. Sanchez</collectorName>
leg.;
<abbrev id="ABBRID0EKODI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="3F60FE637D7080B64A6390C10D8F9453">CEMT</collectionCode>
</abbrev>
</materialsCitation>
<materialsCitation id="150A3AE3444455CEA08238392AE88C6A" collectingDate="1994-08-31" collectionCode="CEMT" collectorName="de Espinal &amp; L. Arellano &amp; R. Sanchez" country="Mexico" location="Carretera Xalapa-Alto Lucera" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
1;
<location id="1499164471E6CD8BE10AD81402EE5325" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:1499164471E6CD8BE10AD81402EE5325" country="Mexico" name="Carretera Xalapa-Alto Lucera" stateProvince="Veracruz de Ignacio de la Llave">Carretera Xalapa-Alto Lucera</location>
,
<quantity id="65904F7204F71D39055F32AC9BADD2E3" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
<collectorName id="61E7AB7D521DFCD1649A7CD7A17F6BDF">de Espinal</collectorName>
;
<date id="606AECAC345B8F02DF3A002AB2629C31" value="1994-08-31">
<collectingDate id="20F9A6A48390D9DE2782F08D61A075CA" value="1994-08-31">31 Aug. 1994</collectingDate>
</date>
;
<collectorName id="7F0C4CF0EC57E972326811F6CBA4B153">L. Arellano</collectorName>
&amp;
<collectorName id="7797435199397EDE2617EC1EB55A0C50">R. Sanchez</collectorName>
leg.;
<abbrev id="ABBRID0EOODI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="9C7CDFD56796489F5352E2E579890912">CEMT</collectionCode>
</abbrev>
</materialsCitation>
<materialsCitation id="6811D983EAB73133C0C82F2E6CBE1993" collectingDate="1990-04-29" collectionCode="CEMT" collectorName="Parq. de la Flora &amp; F. Capistran" country="Mexico" location="Fauna Silvestre" municipality="Catemaco" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
1;
<collectingMunicipality id="50764706A0195B9BCBB3493C76B49F46">Catemaco</collectingMunicipality>
,
<collectorName id="495FBF39E2AE710C87CDCD33647E3747">Parq. de la Flora</collectorName>
y
<location id="5B9AE01455920ACBEA699526D751A229" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:5B9AE01455920ACBEA699526D751A229" country="Mexico" municipality="Catemaco" name="Fauna Silvestre" stateProvince="Veracruz de Ignacio de la Llave">Fauna Silvestre</location>
tropical;
<date id="D7FE193A34DA3604C3FE7285BFBF9BFB" value="1990-04-29">
<collectingDate id="F8186E0E1004213FED1581EA3C6FAC85" value="1990-04-29">29 Apr. 1990</collectingDate>
</date>
;
<collectorName id="FD26AC175ABB5C4C701C1B3D943C6AD6">F. Capistran</collectorName>
leg.;
<abbrev id="ABBRID0ESODI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="0DE3F3E8DF0EC632BB16A8B4615660E6">CEMT</collectionCode>
</abbrev>
</materialsCitation>
<materialsCitation id="54F57D8F6F3EA2ED2373EF71350B90FD" collectingDate="1997-03-19" collectionCode="CEMT" collectorName="R. Sanchez &amp; M. E. Favila" country="Mexico" location="Mpio de Papantla" municipality="Plan" specimenCount="1" stateProvince="Hidalgo">
1;
<collectingMunicipality id="B43FD220DE7D162AAAB70873032167CB">Plan</collectingMunicipality>
de
<collectingRegion id="C24B441F9CD941C8C66425872AD1C9E6" country="Mexico" name="Hidalgo">Hidalgo</collectingRegion>
,
<location id="4700DF845DDA0AB2DFFA895F6F9781DA" LSID="urn:lsid:plazi:treatment:5EABDD6770CF585E881D66A15D4697CB:4700DF845DDA0AB2DFFA895F6F9781DA" country="Mexico" municipality="Plan" name="Mpio de Papantla" stateProvince="Hidalgo">Mpio de Papantla</location>
;
<date id="6EFBD817BA01D39B263995C53F67EB27" value="1997-03-19">
<collectingDate id="78577906DC5BAEA292EF7F72D17D4AB5" value="1997-03-19">19 Mar. 1997</collectingDate>
</date>
;
<collectorName id="21E798B50B0D8C34F96D6D91941B08B9">R. Sanchéz</collectorName>
&amp;
<collectorName id="5055F68AC08588846A12D6D497108B55">M. E. Favila</collectorName>
leg.;
<abbrev id="ABBRID0EWODI" xlink_title="Coleção Entomológica de Mato Grosso Eurides Furtado">
<collectionCode id="CA56FB68DC8D2BA1F75A7A1B0503F406">CEMT</collectionCode>
</abbrev>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>