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<mods:title id="A80E17D542B647E53D3D34A0748FDC91">Late Eocene-early Oligocene snakes from the Transylvanian Basin (Romania)</mods:title>
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<mods:namePart id="8099597BBCE331F73227ED5E72A0ECB1">Venczel, Márton</mods:namePart>
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<mods:affiliation id="6E6AAAB52DAB292E0D77B8A2EE6E443E">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & Țării CriȘurilor Museum, Natural History Department, 1 / A Armatei Române, Oradea, RO- 410087 (Romania)</mods:affiliation>
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<mods:namePart id="33BEA354CE3A07D993C8725287C1C43A">Codrea, Vlad A.</mods:namePart>
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<mods:affiliation id="3C74DCA68C79410885AD06364A7757ED">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & Țării CriȘurilor Museum, Natural History Department, 1 / A Armatei Române, Oradea, RO- 410087 (Romania) & MureȘ County Museum, Natural History Department, 24 Horea Street, Târgu MureȘ, RO- 540036 (Romania) & ’ Emil Racoviță’ Speleological Institute, Calea 13 Septembrie, RO- 050711, Sector 5, Bucharest (Romania)</mods:affiliation>
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<mods:namePart id="5474E436DF8E386AA497E90EF61CEFDC">Solomon, Alexandru A.</mods:namePart>
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<mods:affiliation id="85F4AFB4ABDFE904035BABF64D8D2508">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & MureȘ County Museum, Natural History Department, 24 Horea Street, Târgu MureȘ, RO- 540036 (Romania)</mods:affiliation>
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<mods:namePart id="5D75AB6AA442F1E82FA7C6F357033AFB">Fărcaș, Cristina</mods:namePart>
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<mods:affiliation id="077F78CC13BC3EE3EEDA6BC6D0D6BFBF">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania)</mods:affiliation>
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<mods:namePart id="44D685F7067353C589662320A745F344">Bordeianu, Marian</mods:namePart>
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<mods:affiliation id="7B004FBCCDA9247BFE95C629B4EBF8E4">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & Transgex SA, Geological Compartment, 2 Vasile Alecsandri Street, RO- 410072, Oradea (Romania)</mods:affiliation>
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<mods:number id="AD3DDB44C68131C5CF360ED9F1B24478">2025-05-14</mods:number>
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<emphasis id="B96AEABBFFACFFE2FE90FDA8FEC2FDA8" bold="true" box="[280,368,566,593]" italics="true" pageId="5" pageNumber="232">Falseryx</emphasis>
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<bibRefCitation id="EF8F4B58FFACFFE2FEFFFDA8FDC1FDA8" author="SZYNDLAR Z. & RAGE J. - C." box="[375,627,566,593]" pageId="5" pageNumber="232" refId="ref10028" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 110 p." type="book" year="2003">Szyndlar & Rage, 2003</bibRefCitation>
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<emphasis id="B96AEABBFFACFFE2FF0CFDF1FF7CFD7E" box="[132,206,623,647]" italics="true" pageId="5" pageNumber="232">Falseryx</emphasis>
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<bibRefCitation id="EF8F4B58FFACFFE2FF5DFDF1FE51FD7E" author="SZYNDLAR Z. & RAGE J. - C." box="[213,483,623,648]" pageId="5" pageNumber="232" refId="ref10028" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 110 p." type="book" year="2003">Szyndlar & Rage, 2003: 59</bibRefCitation>
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.
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<paragraph id="8BA136A9FFACFFE2FF0CFD48FCBBFB57" blockId="5.[130,777,695,1199]" pageId="5" pageNumber="232">
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The genus
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<emphasis id="B96AEABBFFACFFE2FF66FD48FE8FFD09" box="[238,317,726,752]" italics="true" pageId="5" pageNumber="232">Falseryx</emphasis>
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<bibRefCitation id="EF8F4B58FFACFFE2FECBFD48FD81FD09" author="SZYNDLAR Z. & RAGE J. - C." box="[323,563,726,752]" pageId="5" pageNumber="232" refId="ref10028" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 110 p." type="book" year="2003">Szyndlar & Rage, 2003</bibRefCitation>
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, was originally established as a member of
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Boinae
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<bibRefCitation id="EF8F4B58FFACFFE2FE4EFD68FD8CFCE8" author="GRAY J. E." box="[454,574,758,785]" pageId="5" pageNumber="232" pagination="193 - 217" refId="ref7904" refString="GRAY J. E. 1825. - A synopsis of the genera of Reptiles and Amphibia, with a description of some new species. Annals of Philosophy, Series 2, 10: 193-217. https://www.biodiversitylibrary.org/page/2531387" type="journal article" year="1825">Gray, 1825</bibRefCitation>
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or Tropidophiinae Brongersma, 1951 (
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<bibRefCitation id="EF8F4B58FFACFFE2FEDDFC88FDF0FCC9" author="SZYNDLAR Z. & RAGE J. - C." box="[341,578,790,816]" pageId="5" pageNumber="232" refId="ref10028" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 110 p." type="book" year="2003">Szyndlar & Rage 2003</bibRefCitation>
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), ‘Tropidophiidae’ (
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<bibRefCitation id="EF8F4B58FFACFFE2FF05FCA8FEDDFCA9" author="SZYNDLAR Z. & SMITH R. & RAGE J. - C." box="[141,367,822,848]" pageId="5" pageNumber="232" pagination="393 - 406" refId="ref10114" refString="SZYNDLAR Z., SMITH R. & RAGE J. - C. 2008. - A new dwarf boa (Serpentes, Booidea, ' Tropidophiidae') from the Early Oligocene of Belgium: a case of the isolation of Western European snake faunas. Zoological Journal of the Linnean Society 152 (2): 393-406. https://doi.org/10.1111/j.1096-3642.2007.00357.x" type="journal article" year="2008">
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Szyndlar
|
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<emphasis id="B96AEABBFFACFFE2FF7EFCA8FE9EFCA9" box="[246,300,822,848]" italics="true" pageId="5" pageNumber="232">et al.</emphasis>
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2008
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</bibRefCitation>
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),
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<taxonomicName id="4C1E4D2AFFACFFE2FE0EFCA8FCB5FCA9" authority="Brongersma, 1951" authorityName="Brongersma" authorityYear="1951" box="[390,775,821,848]" class="Reptilia" family="Tropidophiidae" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="232" phylum="Chordata" rank="family">Tropidophiidae Brongersma, 1951</taxonomicName>
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(
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<bibRefCitation id="EF8F4B58FFACFFE2FF05FCC8FEA0FC89" author="IVANOV M." box="[141,274,854,880]" pageId="5" pageNumber="232" pagination="85 - 110" refId="ref8191" refString="IVANOV M. 2022. - Miocene Snakes of Eurasia: A review of the evolution of snake communities. in GOWER D. & ZAHER H. (eds), The Origin and Early Evolution of Snakes. Cambridge University Press, Cambridge: 85-110. https://doi.org/10.1017/9781108938891.007" type="book chapter" year="2022">Ivanov 2022</bibRefCitation>
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), and finally classified as Alethinophidia
|
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<emphasis id="B96AEABBFFACFFE2FD3EFCC8FF03FC76" italics="true" pageId="5" pageNumber="232">incertae sedis</emphasis>
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(
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<bibRefCitation id="EF8F4B58FFACFFE2FF49FCEBFE7FFC69" author="SMITH K. T. & GEORGALIS G. L." box="[193,461,885,912]" pageId="5" pageNumber="232" pagination="55 - 84" refId="ref9597" refString="SMITH K. T. & GEORGALIS G. L. 2022. - The diversity and distribution of Palaeogene snakes: a review with comments on vertebral sufficiency, in GOWER D. & ZAHER H. (eds), The Origin and Early Evolution of Snakes. Cambridge University Press, Cambridge: 55-84." type="book chapter" year="2022">Smith & Georgalis 2022</bibRefCitation>
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||||
). The uncertainties regarding the phylogenetic position of
|
||||
<taxonomicName id="4C1E4D2AFFACFFE2FE42FC0BFDACFC56" authorityName="Szyndlar & Rage" authorityYear="2003" box="[458,542,917,943]" class="Reptilia" family="Tropidophiidae" genus="Falseryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="232" phylum="Chordata" rank="genus">
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<emphasis id="B96AEABBFFACFFE2FE42FC0BFDACFC56" box="[458,542,917,943]" italics="true" pageId="5" pageNumber="232">Falseryx</emphasis>
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</taxonomicName>
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stem primarily from the fact that it is known exclusively from vertebrae. There is a single published vertebral synapomorphy of tropidophiids: the presence of an anteroventral straight corner of the hypapophyses of trunk vertebrae (
|
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<bibRefCitation id="EF8F4B58FFACFFE2FE7CFB8BFD4DFBD6" author="SMITH K. T. & GEORGALIS G. L." box="[500,767,1045,1071]" pageId="5" pageNumber="232" pagination="55 - 84" refId="ref9597" refString="SMITH K. T. & GEORGALIS G. L. 2022. - The diversity and distribution of Palaeogene snakes: a review with comments on vertebral sufficiency, in GOWER D. & ZAHER H. (eds), The Origin and Early Evolution of Snakes. Cambridge University Press, Cambridge: 55-84." type="book chapter" year="2022">Smith & Georgalis 2022</bibRefCitation>
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;
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<bibRefCitation id="EF8F4B58FFACFFE2FF0CFBABFE1AFBB6" author="SZYNDLAR Z. & GEORGALIS G. L." box="[132,424,1077,1103]" pageId="5" pageNumber="232" pagination="717 - 886" refId="ref9968" refString="SZYNDLAR Z. & GEORGALIS G. L. 2023. - An illustrated atlas of the vertebral morphology of extant non-caenophidian snakes, with special emphasis on the cloacal and caudal portions of the column. Vertebrate Zoology 73: 717-886. https://doi.org/10.3897/vz.73.e101372" type="journal article" year="2023">Szyndlar & Georgalis 2023</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EF8F4B58FFACFFE2FE3EFBABFDDDFBB6" author="ZAHER H. & TRUSZ C. & KOCH C. & ENTIAUSPE-NETO O. M. & BATTILANA J. & GRAZZIOTIN F. G." box="[438,623,1077,1103]" pageId="5" pageNumber="232" pagination="2319289" refId="ref10799" refString="ZAHER H., TRUSZ C., KOCH C., ENTIAUSPE-NETO O. M., BATTILANA J. & GRAZZIOTIN F. G. 2024. - Molecular phylogeny and biogeography of the dwarf boas of the family Tropidophiidae (Serpentes: Alethinophidia). Systematics and Biodiversity 22 (1): 2319289. https://doi.org/10.1080/14772000.2024.2319289" type="journal article" year="2024">
|
||||
Zaher
|
||||
<emphasis id="B96AEABBFFACFFE2FE74FBA8FD9DFBB6" box="[508,559,1077,1103]" italics="true" pageId="5" pageNumber="232">et al.</emphasis>
|
||||
2024
|
||||
</bibRefCitation>
|
||||
); notably, this feature is absent in
|
||||
<taxonomicName id="4C1E4D2AFFACFFE2FEDBFBCBFE16FB96" authorityName="Szyndlar & Rage" authorityYear="2003" box="[339,420,1109,1135]" class="Reptilia" family="Tropidophiidae" genus="Falseryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="232" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFACFFE2FEDBFBCBFE16FB96" box="[339,420,1109,1135]" italics="true" pageId="5" pageNumber="232">Falseryx</emphasis>
|
||||
</taxonomicName>
|
||||
, which has a haemal keel instead of a hypapophysis in its mid- and posterior trunk vertebrae, hinting further that this genus does not belong to tropidophiids.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
240
data/03/B7/87/03B787BFFFACFFE2FF2FFB6AFBF0F815.xml
Normal file
240
data/03/B7/87/03B787BFFFACFFE2FF2FFB6AFBF0F815.xml
Normal file
|
|
@ -0,0 +1,240 @@
|
|||
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<treatment id="03B787BFFFACFFE2FF2FFB6AFBF0F815" LSID="urn:lsid:plazi:treatment:03B787BFFFACFFE2FF2FFB6AFBF0F815" httpUri="http://treatment.plazi.org/id/03B787BFFFACFFE2FF2FFB6AFBF0F815" lastPageNumber="232" pageId="5" pageNumber="232">
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<subSubSection id="C3046522FFACFFE2FF2FFB6AFD56FAF7" box="[167,740,1268,1295]" pageId="5" pageNumber="232" type="nomenclature">
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<paragraph id="8BA136A9FFACFFE2FF2FFB6AFD56FAF7" blockId="5.[167,740,1268,1295]" box="[167,740,1268,1295]" pageId="5" pageNumber="232">
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<heading id="D0E981C5FFACFFE2FF2FFB6AFD56FAF7" box="[167,740,1268,1295]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="232" reason="2">
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<taxonomicName id="4C1E4D2AFFACFFE2FF2FFB6AFD56FAF7" authority="Szyndlar, Smith & Rage, 2008" authorityName="Szyndlar, Smith & Rage" authorityYear="2008" box="[167,740,1268,1295]" class="Reptilia" family="Tropidophiidae" genus="Falseryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="232" phylum="Chordata" rank="species" species="neervelpensis">
|
||||
<emphasis id="B96AEABBFFACFFE2FF2FFB6AFE23FAF6" bold="true" box="[167,401,1268,1295]" italics="true" pageId="5" pageNumber="232">Falseryx neervelpensis</emphasis>
|
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<bibRefCitation id="EF8F4B58FFACFFE2FE1FFB6AFD56FAF7" author="SZYNDLAR Z. & SMITH R. & RAGE J. - C." box="[407,740,1268,1295]" pageId="5" pageNumber="232" pagination="393 - 406" refId="ref10114" refString="SZYNDLAR Z., SMITH R. & RAGE J. - C. 2008. - A new dwarf boa (Serpentes, Booidea, ' Tropidophiidae') from the Early Oligocene of Belgium: a case of the isolation of Western European snake faunas. Zoological Journal of the Linnean Society 152 (2): 393-406. https://doi.org/10.1111/j.1096-3642.2007.00357.x" type="journal article" year="2008">Szyndlar, Smith & Rage, 2008</bibRefCitation>
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</taxonomicName>
|
||||
</heading>
|
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</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3046522FFACFFE2FECFFAAAFDF7FAB7" box="[327,581,1332,1358]" pageId="5" pageNumber="232" type="reference_group">
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<paragraph id="8BA136A9FFACFFE2FECFFAAAFDF7FAB7" blockId="5.[327,581,1332,1358]" box="[327,581,1332,1358]" pageId="5" pageNumber="232">
|
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<taxonomicName id="4C1E4D2AFFACFFE2FECFFAAAFDF7FAB7" authorityName="Szyndlar, Smith & Rage" authorityYear="2008" box="[327,581,1332,1358]" class="Reptilia" family="Tropidophiidae" genus="Falseryx" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="232" phylum="Chordata" rank="species" species="neervelpensis">
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<emphasis id="B96AEABBFFACFFE2FECFFAAAFE2BFAB7" box="[327,409,1332,1358]" italics="true" pageId="5" pageNumber="232">Falseryx</emphasis>
|
||||
cf.
|
||||
<emphasis id="B96AEABBFFACFFE2FE4AFAAAFDF7FAB7" box="[450,581,1332,1358]" italics="true" pageId="5" pageNumber="232">neervelpensis</emphasis>
|
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</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3046522FFACFFE2FE16FACAFE5FFA97" box="[414,493,1364,1390]" pageId="5" pageNumber="232" type="description">
|
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<paragraph id="8BA136A9FFACFFE2FE16FACAFE5FFA97" blockId="5.[414,493,1364,1390]" box="[414,493,1364,1390]" pageId="5" pageNumber="232">
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(
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<figureCitation id="13252A2CFFACFFE2FE2FFACAFE56FA97" box="[423,484,1364,1390]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
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<subSubSection id="C3046522FFACFFE2FF0CFA13FEF2FA46" pageId="5" pageNumber="232" type="materials_examined">
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<paragraph id="8BA136A9FFACFFE2FF0CFA13FEF2FA46" blockId="5.[132,774,1421,1471]" pageId="5" pageNumber="232">
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|
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. —
|
||||
<materialsCitation id="3B763CF4FFACFFE2FE92FA13FE8EFA46" collectionCode="UBB" location="Suceag" pageId="5" pageNumber="232" specimenCode="UBB V 1042/1-2, UBB V 1043/1-9" specimenCount="1">
|
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<location id="8EC16072FFACFFE2FE92FA13FEECFA5C" LSID="urn:lsid:plazi:treatment:03B787BFFFACFFE2FF2FFB6AFBF0F815:8EC16072FFACFFE2FE92FA13FEECFA5C" box="[282,350,1421,1445]" name="Suceag" pageId="5" pageNumber="232">Suceag</location>
|
||||
1: 11 trunk vertebrae (
|
||||
<specimenCode id="DBB89ED2FFACFFE2FDC5FA13FCB1FA5C" box="[589,771,1421,1445]" pageId="5" pageNumber="232">
|
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UBB
|
||||
<date id="FFA01069FFACFFE2FD01FA13FD69FA5C" box="[649,731,1421,1445]" pageId="5" pageNumber="232" value="1042-05">V 1042</date>
|
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/1-2
|
||||
</specimenCode>
|
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;
|
||||
<specimenCode id="DBB89ED2FFACFFE2FF0EFA36FE86FA46" box="[134,308,1447,1471]" pageId="5" pageNumber="232">
|
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UBB
|
||||
<date id="FFA01069FFACFFE2FF36FA36FEBCFA46" box="[190,270,1447,1471]" pageId="5" pageNumber="232" value="1043-05">V 1043</date>
|
||||
/1-9
|
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</specimenCode>
|
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)
|
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</materialsCitation>
|
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.
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</paragraph>
|
||||
</subSubSection>
|
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<subSubSection id="C3046522FFACFFE2FF0CFA6AFBFDFA14" pageId="5" pageNumber="232" type="description">
|
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<paragraph id="8BA136A9FFACFFE2FF0CFA6AFEA7F9F5" blockId="5.[131,777,1524,2028]" box="[132,277,1524,1550]" pageId="5" pageNumber="232">
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<smallCapsWord id="8D47A075FFACFFE2FF0CFA6AFEA7F9F5" baselines="1543,1543" box="[132,277,1524,1550]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="5" pageNumber="232">DESCRIPTION</smallCapsWord>
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</paragraph>
|
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<paragraph id="8BA136A9FFACFFE2FF0CF98AFA1DFC69" blockId="5.[131,777,1524,2028]" lastBlockId="5.[811,1457,216,1517]" pageId="5" pageNumber="232">
|
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UBB
|
||||
<date id="FFA01069FFACFFE2FF4AF98AFEAEF9D4" box="[194,284,1555,1582]" pageId="5" pageNumber="232" value="1042-05">V 1042</date>
|
||||
/1, is the largest available specimen representing a middle trunk vertebra, lacking its prezygapophyseal areas and the right posterolateral side of its neural arch. In dorsal view, the vertebra appears slightly longer than wide featuring a relatively deep interzygapophyseal constriction, whereas the posterior notch of the neural arch is also deep, bordered by the nearly straight posteromedian margins of the neural arch (
|
||||
<figureCitation id="13252A2CFFACFFE2FE87F96CFED6F8F5" box="[271,356,1778,1804]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2A</figureCitation>
|
||||
). The neural spine, representing about one third of the length of the neural arch, is extremely low and widened, and without an anterior or posterior overhang; tiny tubercles are present on its dorsal anterior limit. The anterior margin of the zygosphene is slightly convex, with two protruding lateral lobes (the left one is damaged). In ventral view, the centrum is longer than wide (centrum length (CL) ≈
|
||||
<quantity id="4CE69B4CFFACFFE2FEABF84FFE38F815" box="[291,394,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" pageId="5" pageNumber="232" unit="mm" value="3.85">3.85 mm</quantity>
|
||||
; centrum width (CW) ≈
|
||||
<quantity id="4CE69B4CFFACFFE2FD14F84FFCB6F815" box="[668,772,2001,2028]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" pageId="5" pageNumber="232" unit="mm" value="3.26">3.26 mm</quantity>
|
||||
; CL/CW ≈ 1.18), whereas the haemal keel is prominent, somewhat broadened anteriorly and posteriorly, but tapers near the condylar neck; on the right side a distinct subcotylar tubercle is preserved. The subcentral grooves are indistinct, whereas the subcentral foramina are present; on the right side a distinctly large subcentral foramen is preserved. Only the right paradiapophysis is preserved; it strongly projects laterally and features a well-developed tubercle on the anterior margin of the prezygapophyseal buttress (
|
||||
<figureCitation id="13252A2CFFACFFE2FAA8FE49FAC0FE08" box="[1312,1394,471,497]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2B</figureCitation>
|
||||
). The surface of the left postzygapophysis preserves numerous lines of arrested growths (LAGs) (for their interpretation, see
|
||||
<bibRefCitation id="EF8F4B58FFACFFE2FCDCFDA8FB94FDA8" author="VENCZEL M. & VASILE S. & CSIKI-SAVA Z." box="[852,1062,566,593]" pageId="5" pageNumber="232" pagination="152 - 163" refId="ref10392" refString="VENCZEL M., VASILE S. & CSIKI-SAVA Z. 2015. - A Late Cretaceous madtsoiid snake from Romania associated with a megaloolithid egg nest - Paleoecological inferences. Cretaceous Research 55: 152-163. https://doi.org/10.1016/j.cretres.2015.02.009" type="journal article" year="2015">
|
||||
Venczel
|
||||
<emphasis id="B96AEABBFFACFFE2FC27FDA9FC56FDA9" box="[943,996,566,592]" italics="true" pageId="5" pageNumber="232">et al.</emphasis>
|
||||
2015
|
||||
</bibRefCitation>
|
||||
), of which six are more discernible, considered as indication of intermittent seasonal growths (
|
||||
<bibRefCitation id="EF8F4B58FFACFFE2FCBEFDE8FC44FD68" author="VENCZEL M. & VASILE S. & CSIKI-SAVA Z." box="[822,1014,630,657]" pageId="5" pageNumber="232" pagination="152 - 163" refId="ref10392" refString="VENCZEL M., VASILE S. & CSIKI-SAVA Z. 2015. - A Late Cretaceous madtsoiid snake from Romania associated with a megaloolithid egg nest - Paleoecological inferences. Cretaceous Research 55: 152-163. https://doi.org/10.1016/j.cretres.2015.02.009" type="journal article" year="2015">
|
||||
Venczel
|
||||
<emphasis id="B96AEABBFFACFFE2FC02FDE9FC09FD69" box="[906,955,630,656]" italics="true" pageId="5" pageNumber="232">et al.</emphasis>
|
||||
2015
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EF8F4B58FFACFFE2FB8AFDE8FB3DFD69" author="VENCZEL M." box="[1026,1167,630,656]" pageId="5" pageNumber="232" pagination="1217025" refId="ref10192" refString="VENCZEL M. 2023. - Updating the fossil record of the alligatoroid crocodylian Diplocynodon from the late Eocene of Transylvanian Basin. Frontiers in Amphibian and Reptile Science 1: 1217025. https://doi.org/10.3389/famrs.2023.1217025" type="journal article" year="2023">Venczel 2023</bibRefCitation>
|
||||
). In lateral view, the haemal keel is prominent and its ventral margin is relatively straight, whereas the subcentral ridges are weakly defined (
|
||||
<figureCitation id="13252A2CFFACFFE2FAC4FD28FA13FD29" box="[1356,1441,694,720]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2C</figureCitation>
|
||||
). In anterior view, the roof of the zygosphene is straight with the lateral lobes tilting upward; the cotylar rim is damaged (
|
||||
<figureCitation id="13252A2CFFACFFE2FCBEFC88FC22FCC9" box="[822,912,790,816]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2D</figureCitation>
|
||||
). In posterior view, the neural arch is depressed, slightly convex dorsally and preserves on the left dorsal side of the postzygapophysis a low bony ridge extending anteroposteriorly. No parazygantral foramen is preserved (
|
||||
<figureCitation id="13252A2CFFACFFE2FADAFCEBFA11FC69" box="[1362,1443,885,912]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2E</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BA136A9FFACFFE2FCCBFC0BFBFDFA14" blockId="5.[811,1457,216,1517]" pageId="5" pageNumber="232">
|
||||
The remaining trunk vertebrae are smaller and fragmentary, exhibiting some morphological variation compared to the UBB
|
||||
<date id="FFA01069FFACFFE2FCE6FC48FC74FC16" box="[878,966,981,1008]" pageId="5" pageNumber="232" value="1042-05">V 1042</date>
|
||||
/
|
||||
<specimenCount id="9D18FD20FFACFFE2FC58FC4BFBFEFC16" box="[976,1100,981,1007]" count="1" pageId="5" pageNumber="232" type="generic">1 specimen</specimenCount>
|
||||
. The neural spine is consistently low and strongly broadened and with either parallel lateral margins (
|
||||
<figureCitation id="13252A2CFFACFFE2FC1CFB8BFC56FBD6" box="[916,996,1045,1071]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2F</figureCitation>
|
||||
), or distinct posterior widening (
|
||||
<figureCitation id="13252A2CFFACFFE2FAC4FB8BFA10FBD6" box="[1356,1442,1045,1071]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2G</figureCitation>
|
||||
). Some specimens feature a well-developed median lobe on the zygosphene (
|
||||
<figureCitation id="13252A2CFFACFFE2FC6FFBCBFBE8FB96" box="[999,1114,1109,1135]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2F, G</figureCitation>
|
||||
). In the UBB
|
||||
<date id="FFA01069FFACFFE2FB74FBCBFAE3FB96" box="[1276,1361,1109,1135]" pageId="5" pageNumber="232" value="1042-05">V 1042</date>
|
||||
/
|
||||
<specimenCount id="9D18FD20FFACFFE2FAD2FBCBFCD2FB76" count="2" pageId="5" pageNumber="232" type="generic">2 specimen</specimenCount>
|
||||
, the paradiapophyses are well-preserved and somewhat differentiated into diapophyseal and parapophyseal portions. The portions of prezygapophyseal buttresses between the paradiapophyses and prezygapophyseal processes, as seen in
|
||||
<taxonomicName id="4C1E4D2AFFACFFE2FCA4FB6AFBBDFAF7" authorityName="Szyndlar, Smith & Rage" authorityYear="2008" box="[812,1039,1268,1294]" class="Reptilia" family="Tropidophiidae" genus="Falseryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="232" phylum="Chordata" rank="species" species="neervelpensis">
|
||||
<emphasis id="B96AEABBFFACFFE2FCA4FB6AFBBDFAF7" box="[812,1039,1268,1294]" italics="true" pageId="5" pageNumber="232">Falseryx neervelpensis</emphasis>
|
||||
</taxonomicName>
|
||||
(see
|
||||
<bibRefCitation id="EF8F4B58FFACFFE2FBC5FB6AFA9EFAF7" author="SZYNDLAR Z. & SMITH R. & RAGE J. - C." box="[1101,1324,1268,1294]" pageId="5" pageNumber="232" pagination="393 - 406" refId="ref10114" refString="SZYNDLAR Z., SMITH R. & RAGE J. - C. 2008. - A new dwarf boa (Serpentes, Booidea, ' Tropidophiidae') from the Early Oligocene of Belgium: a case of the isolation of Western European snake faunas. Zoological Journal of the Linnean Society 152 (2): 393-406. https://doi.org/10.1111/j.1096-3642.2007.00357.x" type="journal article" year="2008">
|
||||
Szyndlar
|
||||
<emphasis id="B96AEABBFFACFFE2FB3BFB6BFB5BFAF7" box="[1203,1257,1268,1294]" italics="true" pageId="5" pageNumber="232">et al.</emphasis>
|
||||
2008
|
||||
</bibRefCitation>
|
||||
), are developed into prominent, anteriorly facing tubercles (
|
||||
<figureCitation id="13252A2CFFACFFE2FAC2FA8AFA13FAD7" box="[1354,1441,1300,1326]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2H</figureCitation>
|
||||
). In anterior view, the outline of the cotyle appears circular, without paracotylar foramina (
|
||||
<figureCitation id="13252A2CFFACFFE2FBF6FACAFB78FA97" box="[1150,1226,1364,1390]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2I</figureCitation>
|
||||
). In UBB
|
||||
<date id="FFA01069FFACFFE2FAB5FACAFA24FA97" box="[1341,1430,1364,1390]" pageId="5" pageNumber="232" value="1043-05">V 1043</date>
|
||||
/1, representing an anterior trunk vertebra (
|
||||
<figureCitation id="13252A2CFFACFFE2FB53FAEAFAF1FA77" box="[1243,1347,1396,1422]" captionStart="FIG" captionStartId="6.[132,143,1335,1352]" captionTargetBox="[132,1455,218,1284]" captionTargetId="figure-408@6.[748,1392,198,759]" captionTargetPageId="6" captionText="FIG. 2. — Isolated trunk vertebrae of Falseryx cf. neervelpensis: A-E, middle trunk vertebra (UBB V 1042/1) in dorsal (A), ventral (B), left lateral (C), anterior (D) and posterior (E) views; F-I, partial trunk vertebrae (UBB V 1043/2, 1043/3 and 1042/2) in dorsal (F, G), ventral (H) and anterior (I) views; J-K, anterior trunk vertebra (UBB V 1043/1) in left lateral (J) and posterior (K) views. Abbreviations: hk, haemal keel; ns, neural spine; tb, tubercul on the prezygapophyseal buttress. Scale bar: 2 mm." pageId="5" pageNumber="232">Fig. 2J, K</figureCitation>
|
||||
), the neural arch appears more vaulted and less elongated, whereas the haemal keel is more prominent and extends posteriorly beyond the condylar neck.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3046522FFACFFE2FCA5F98AFBF0F815" pageId="5" pageNumber="232" type="discussion">
|
||||
<paragraph id="8BA136A9FFACFFE2FCA5F98AFC23F9D5" blockId="5.[810,1457,1556,2028]" box="[813,913,1556,1582]" pageId="5" pageNumber="232">
|
||||
<smallCapsWord id="8D47A075FFACFFE2FCA5F98AFC23F9D5" baselines="1575,1575" box="[813,913,1556,1582]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="5" pageNumber="232">REMARKS</smallCapsWord>
|
||||
</paragraph>
|
||||
<paragraph id="8BA136A9FFACFFE2FCA5F9ADFBF0F815" blockId="5.[810,1457,1556,2028]" pageId="5" pageNumber="232">
|
||||
The size and morphology of the above-described vertebrae from Suceag 1 are closely resemble those of
|
||||
<taxonomicName id="4C1E4D2AFFACFFE2FB5BF9CDFA03F994" authorityName="Szyndlar, Smith & Rage" authorityYear="2008" box="[1235,1457,1619,1645]" class="Reptilia" family="Tropidophiidae" genus="Falseryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="232" phylum="Chordata" rank="species" species="neervelpensis">
|
||||
<emphasis id="B96AEABBFFACFFE2FB5BF9CDFA03F994" box="[1235,1457,1619,1645]" italics="true" pageId="5" pageNumber="232">Falseryx neervelpensis</emphasis>
|
||||
</taxonomicName>
|
||||
known from the early Oligocene (MP 21) locality of Boutersem TGV,
|
||||
<collectingCountry id="F3097639FFACFFE2FC3CF90DFBA1F954" box="[948,1043,1683,1709]" name="Belgium" pageId="5" pageNumber="232">Belgium</collectingCountry>
|
||||
. In particular, the prominent tubercles protruding anteriorly on the prezygapophyseal buttresses are highly similar to those observed in this species (
|
||||
<bibRefCitation id="EF8F4B58FFACFFE2FA95F94CFCD7F8F5" author="SZYNDLAR Z. & SMITH R. & RAGE J. - C." pageId="5" pageNumber="232" pagination="393 - 406" refId="ref10114" refString="SZYNDLAR Z., SMITH R. & RAGE J. - C. 2008. - A new dwarf boa (Serpentes, Booidea, ' Tropidophiidae') from the Early Oligocene of Belgium: a case of the isolation of Western European snake faunas. Zoological Journal of the Linnean Society 152 (2): 393-406. https://doi.org/10.1111/j.1096-3642.2007.00357.x" type="journal article" year="2008">
|
||||
Szyndlar
|
||||
<emphasis id="B96AEABBFFACFFE2FAF6F94DFA02F915" box="[1406,1456,1746,1772]" italics="true" pageId="5" pageNumber="232">et al.</emphasis>
|
||||
2008
|
||||
</bibRefCitation>
|
||||
: figs 1, 2). The vertebrae of the
|
||||
<typeStatus id="54A5880BFFACFFE2FB26F96DFB6EF8F4" box="[1198,1244,1779,1805]" pageId="5" pageNumber="232">type</typeStatus>
|
||||
species of the genus (i.e.,
|
||||
<taxonomicName id="4C1E4D2AFFACFFE2FCE8F88CFAA9F8D5" authority="Szyndlar & Rage, 2003" authorityName="Szyndlar & Rage" authorityYear="2003" box="[864,1307,1810,1837]" class="Reptilia" family="Tropidophiidae" genus="Falseryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="232" phylum="Chordata" rank="species" species="petersbuchi">
|
||||
<emphasis id="B96AEABBFFACFFE2FCE8F88CFB96F8D5" box="[864,1060,1810,1836]" italics="true" pageId="5" pageNumber="232">Falseryx petersbuchi</emphasis>
|
||||
<bibRefCitation id="EF8F4B58FFACFFE2FBAFF88CFAA9F8D5" author="SZYNDLAR Z. & RAGE J. - C." box="[1063,1307,1810,1837]" pageId="5" pageNumber="232" refId="ref10028" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 110 p." type="book" year="2003">Szyndlar & Rage, 2003</bibRefCitation>
|
||||
</taxonomicName>
|
||||
), known from the Early Miocene (MN 4) localities of Petersbuch 2,
|
||||
<collectingCountry id="F3097639FFACFFE2FAC2F8ACFA1FF8B5" box="[1354,1453,1842,1868]" name="Germany" pageId="5" pageNumber="232">Germany</collectingCountry>
|
||||
and Dolnice,
|
||||
<collectingCountry id="F3097639FFACFFE2FC36F8CCFBDAF895" box="[958,1128,1874,1900]" name="Czech Republic" pageId="5" pageNumber="232">Czech Republic</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="EF8F4B58FFACFFE2FBF2F8CCFAC2F895" author="SZYNDLAR Z. & RAGE J. - C." box="[1146,1392,1874,1901]" pageId="5" pageNumber="232" refId="ref10028" refString="SZYNDLAR Z. & RAGE J. - C. 2003. - Non-erycine Booidea from the Oligocene and Miocene of Europe. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Krakow, 110 p." type="book" year="2003">Szyndlar & Rage 2003</bibRefCitation>
|
||||
), also bear resemblance to the specimens from Suceag 1. However, they lack the tubercles developed on the prezygapophyseal buttresses, which are considered diagnostic of
|
||||
<taxonomicName id="4C1E4D2AFFACFFE2FA9CF82CFA1CF835" authorityName="Szyndlar, Smith & Rage" authorityYear="2008" box="[1300,1454,1970,1996]" class="Reptilia" family="Tropidophiidae" genus="Falseryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="232" phylum="Chordata" rank="species" species="neervelpensis">
|
||||
<emphasis id="B96AEABBFFACFFE2FA9CF82CFA1CF835" box="[1300,1454,1970,1996]" italics="true" pageId="5" pageNumber="232">F. neervelpensis</emphasis>
|
||||
</taxonomicName>
|
||||
(see
|
||||
<bibRefCitation id="EF8F4B58FFACFFE2FCD4F84FFB81F815" author="SZYNDLAR Z. & SMITH R. & RAGE J. - C." box="[860,1075,2001,2028]" pageId="5" pageNumber="232" pagination="393 - 406" refId="ref10114" refString="SZYNDLAR Z., SMITH R. & RAGE J. - C. 2008. - A new dwarf boa (Serpentes, Booidea, ' Tropidophiidae') from the Early Oligocene of Belgium: a case of the isolation of Western European snake faunas. Zoological Journal of the Linnean Society 152 (2): 393-406. https://doi.org/10.1111/j.1096-3642.2007.00357.x" type="journal article" year="2008">
|
||||
Szyndlar
|
||||
<emphasis id="B96AEABBFFACFFE2FC48F84CFC41F812" box="[960,1011,2001,2027]" italics="true" pageId="5" pageNumber="232">et al.</emphasis>
|
||||
2008
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
451
data/03/B7/87/03B787BFFFAEFFEEFF63FA0DFB0AFCA9.xml
Normal file
451
data/03/B7/87/03B787BFFFAEFFEEFF63FA0DFB0AFCA9.xml
Normal file
|
|
@ -0,0 +1,451 @@
|
|||
<document id="1505204C6E2C4AC3A0DDD1A5A64D8069" ID-DOI="10.5852/cr-palevol2025v24a13" ID-ISSN="1777-571X" ID-Zenodo-Dep="15624605" ID-ZooBank="urn:lsid:zoobank.org:pub:E1B31130-3680-402C-987D-F1FF3DB0D8DD" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1749476715322" checkinUser="felipe" docAuthor="Venczel, Márton, Codrea, Vlad A., Solomon, Alexandru A., Fărcaș, Cristina & Bordeianu, Marian" docDate="2025" docId="03B787BFFFAEFFEEFF63FA0DFB0AFCA9" docLanguage="en" docName="CRPalevol.24.13.229-240.pdf" docOrigin="Comptes Rendus Palevol 24 (13)" docSource="https://doi.org/10.5852/cr-palevol2025v24a13" docStyle="DocumentStyle:DACF86F01658A8850E13A63D98C287FD.1:CRPalevol.2020-.journal_article" docStyleId="DACF86F01658A8850E13A63D98C287FD" docStyleName="CRPalevol.2020-.journal_article" docStyleVersion="1" docTitle="Messelophis variatus Baszio 2004" docType="treatment" docVersion="1" lastPageNumber="236" masterDocId="FF8EFFC7FFA9FFE7FF88FF9EFFB2FFF9" masterDocTitle="Late Eocene-early Oligocene snakes from the Transylvanian Basin (Romania)" masterLastPageNumber="240" masterPageNumber="229" pageNumber="234" updateTime="1749562744962" updateUser="jonas" zenodo-license-document="CC-BY-4.0">
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<mods:title id="41E6F63A0A5555666ACFB81387F5C783">Late Eocene-early Oligocene snakes from the Transylvanian Basin (Romania)</mods:title>
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<mods:namePart id="AEF3E309CCB422234806677CB20FB3F6">Venczel, Márton</mods:namePart>
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<mods:affiliation id="CBD4739386E72ABFEACE8E2DCE886068">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & Țării CriȘurilor Museum, Natural History Department, 1 / A Armatei Române, Oradea, RO- 410087 (Romania)</mods:affiliation>
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<mods:namePart id="DEF9565CFB8CECEAA87BC5020C920733">Codrea, Vlad A.</mods:namePart>
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<mods:affiliation id="3B254E0777AA45EBA0A77F27E7DF79BC">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & Țării CriȘurilor Museum, Natural History Department, 1 / A Armatei Române, Oradea, RO- 410087 (Romania) & MureȘ County Museum, Natural History Department, 24 Horea Street, Târgu MureȘ, RO- 540036 (Romania) & ’ Emil Racoviță’ Speleological Institute, Calea 13 Septembrie, RO- 050711, Sector 5, Bucharest (Romania)</mods:affiliation>
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<mods:nameIdentifier id="591BE943497E3C70C795499DA4D9ADB4" type="email">codrea_vlad@yahoo.fr</mods:nameIdentifier>
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<mods:namePart id="F5F958F0781AE80B8C67C0DB62016AC3">Solomon, Alexandru A.</mods:namePart>
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<mods:affiliation id="8CD3873F2B70B8D05D4AABAA666DCF88">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & MureȘ County Museum, Natural History Department, 24 Horea Street, Târgu MureȘ, RO- 540036 (Romania)</mods:affiliation>
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<mods:namePart id="B0B887D332C8DDD9F273F2FF94BF7458">Fărcaș, Cristina</mods:namePart>
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<mods:affiliation id="CCDF1F3DF1BFE7AC2911C4FA665B9EDC">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania)</mods:affiliation>
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<mods:namePart id="57733B626603B7265AD3470B924BC4EB">Bordeianu, Marian</mods:namePart>
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<mods:affiliation id="1307A95DD47CBC913DE69C9C3E3886B1">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & Transgex SA, Geological Compartment, 2 Vasile Alecsandri Street, RO- 410072, Oradea (Romania)</mods:affiliation>
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<mods:identifier id="901F27A74FA4477C89FAF047A935AAC4" type="ZooBank">urn:lsid:zoobank.org:pub:E1B31130-3680-402C-987D-F1FF3DB0D8DD</mods:identifier>
|
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</mods:mods>
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<treatment id="03B787BFFFAEFFEEFF63FA0DFB0AFCA9" LSID="urn:lsid:plazi:treatment:03B787BFFFAEFFEEFF63FA0DFB0AFCA9" httpUri="http://treatment.plazi.org/id/03B787BFFFAEFFEEFF63FA0DFB0AFCA9" lastPageId="9" lastPageNumber="236" pageId="7" pageNumber="234">
|
||||
<subSubSection id="C3046522FFAEFFE0FF63FA0DFD12FA57" box="[235,672,1427,1454]" pageId="7" pageNumber="234" type="nomenclature">
|
||||
<paragraph id="8BA136A9FFAEFFE0FF63FA0DFD12FA57" blockId="7.[235,672,1427,1454]" box="[235,672,1427,1454]" pageId="7" pageNumber="234">
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||||
<heading id="D0E981C5FFAEFFE0FF63FA0DFD12FA57" box="[235,672,1427,1454]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="234" reason="2">
|
||||
Species
|
||||
<taxonomicName id="4C1E4D2AFFAEFFE0FEB7FA0DFD12FA57" authority="Baszio, 2004" authorityName="Baszio" authorityYear="2004" box="[319,672,1427,1454]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="234" phylum="Chordata" rank="species" species="variatus">
|
||||
<emphasis id="B96AEABBFFAEFFE0FEB7FA0DFDBDFA54" box="[319,527,1427,1453]" italics="true" pageId="7" pageNumber="234">Messelophis variatus</emphasis>
|
||||
<bibRefCitation id="EF8F4B58FFAEFFE0FD9EFA0DFD12FA57" author="BASZIO S." box="[534,672,1427,1454]" pageId="7" pageNumber="234" pagination="47 - 66" refId="ref7068" refString="BASZIO S. 2004. - Messelophis variatus n. gen. n. sp., from the Eocene of Messel: a tropidopheine snake with affinities to Erycinae (Boidae). Courier Forschungsinstitut Senckenberg 252: 47-66." type="journal article" year="2004">Baszio, 2004</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3046522FFAEFFE0FEC5FA4DFD8CFA14" box="[333,574,1491,1517]" pageId="7" pageNumber="234" type="reference_group">
|
||||
<paragraph id="8BA136A9FFAEFFE0FEC5FA4DFD8CFA14" blockId="7.[333,574,1491,1517]" box="[333,574,1491,1517]" pageId="7" pageNumber="234">
|
||||
cf.
|
||||
<taxonomicName id="4C1E4D2AFFAEFFE0FEE7FA4DFD8CFA14" authorityName="Baszio" authorityYear="2004" box="[367,574,1491,1517]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="234" phylum="Chordata" rank="species" species="variatus">
|
||||
<emphasis id="B96AEABBFFAEFFE0FEE7FA4DFD8CFA14" box="[367,574,1491,1517]" italics="true" pageId="7" pageNumber="234">Messelophis variatus</emphasis>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3046522FFAEFFE0FEEEFA6DFD97F9F4" box="[358,549,1523,1549]" pageId="7" pageNumber="234" type="description">
|
||||
<paragraph id="8BA136A9FFAEFFE0FEEEFA6DFD97F9F4" blockId="7.[358,549,1523,1549]" box="[358,549,1523,1549]" pageId="7" pageNumber="234">
|
||||
(
|
||||
<figureCitation id="13252A2CFFAEFFE0FEE7FA6DFE1DF9F4" box="[367,431,1523,1549]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="7" pageNumber="234">Figs 3</figureCitation>
|
||||
A-K; 4A-I)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3046522FFAEFFE0FF0CF9B3FCB4F980" pageId="7" pageNumber="234" type="materials_examined">
|
||||
<paragraph id="8BA136A9FFAEFFE0FF0CF9B3FCB4F980" blockId="7.[130,775,1580,1657]" pageId="7" pageNumber="234">
|
||||
<smallCapsWord id="8D47A075FFAEFFE0FF0CF9B3FF55F9BA" baselines="1598,1599" box="[132,231,1581,1604]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Material" pageId="7" pageNumber="234">MATERIAL</smallCapsWord>
|
||||
. —
|
||||
<materialsCitation id="3B763CF4FFAEFFE0FE9EF9B2FD8DF9A7" collectionCode="UBB" location="CetaTuia Hill" pageId="7" pageNumber="234" specimenCode="UBB V 1045, UBB V 1046/1-4" specimenCount="1">
|
||||
<location id="8EC16072FFAEFFE0FE9EF9B2FE28F9BD" LSID="urn:lsid:plazi:treatment:03B787BFFFAEFFEEFF63FA0DFB0AFCA9:8EC16072FFAEFFE0FE9EF9B2FE28F9BD" box="[278,410,1580,1604]" name="CetaTuia Hill" pageId="7" pageNumber="234">CetăŢuia Hill</location>
|
||||
: one anterior trunk vertebra (
|
||||
<specimenCode id="DBB89ED2FFAEFFE0FD35F9B2FF0BF9A7" pageId="7" pageNumber="234">
|
||||
UBB
|
||||
<date id="FFA01069FFAEFFE0FD7EF9B3FF0BF9A7" pageId="7" pageNumber="234" value="1045-05">V 1045</date>
|
||||
</specimenCode>
|
||||
), four trunk vertebrae (
|
||||
<specimenCode id="DBB89ED2FFAEFFE0FE05F9D9FD83F9A6" box="[397,561,1607,1631]" pageId="7" pageNumber="234">
|
||||
UBB
|
||||
<date id="FFA01069FFAEFFE0FE4BF9D9FDBFF9A6" box="[451,525,1607,1631]" pageId="7" pageNumber="234" value="1046-05">V 1046</date>
|
||||
/1-4
|
||||
</specimenCode>
|
||||
);
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B763CF4FFAEFFE0FDCBF9D9FCB3F980" collectionCode="UBB" location="Suceag" pageId="7" pageNumber="234" specimenCode="UBB V 1047/1-8, UBB V 1044" specimenCount="1">
|
||||
<location id="8EC16072FFAEFFE0FDCBF9D9FD31F9A6" LSID="urn:lsid:plazi:treatment:03B787BFFFAEFFEEFF63FA0DFB0AFCA9:8EC16072FFAEFFE0FDCBF9D9FD31F9A6" box="[579,643,1607,1631]" name="Suceag" pageId="7" pageNumber="234">Suceag</location>
|
||||
1: eight trunk vertebrae (
|
||||
<specimenCode id="DBB89ED2FFAEFFE0FF60F9FFFE25F980" box="[232,407,1633,1657]" pageId="7" pageNumber="234">
|
||||
UBB
|
||||
<date id="FFA01069FFAEFFE0FEA9F9FCFEC3F980" box="[289,369,1633,1657]" pageId="7" pageNumber="234" value="1047-05">V 1047</date>
|
||||
/1-8
|
||||
</specimenCode>
|
||||
), one caudal vertebra (
|
||||
<specimenCode id="DBB89ED2FFAEFFE0FDFCF9FFFD48F980" box="[628,762,1633,1657]" pageId="7" pageNumber="234">
|
||||
UBB
|
||||
<date id="FFA01069FFAEFFE0FD24F9FCFD48F980" box="[684,762,1633,1657]" pageId="7" pageNumber="234" value="1044-05">V 1044</date>
|
||||
</specimenCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3046522FFAEFFEFFF0CF92DFA2EF9F4" lastPageId="8" lastPageNumber="235" pageId="7" pageNumber="234" type="description">
|
||||
<paragraph id="8BA136A9FFAEFFE0FF0CF92DFEA7F932" blockId="7.[130,775,1715,2028]" box="[132,277,1715,1741]" pageId="7" pageNumber="234">
|
||||
<smallCapsWord id="8D47A075FFAEFFE0FF0CF92DFEA7F932" baselines="1734,1734" box="[132,277,1715,1741]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="7" pageNumber="234">DESCRIPTION</smallCapsWord>
|
||||
</paragraph>
|
||||
<paragraph id="8BA136A9FFAEFFEFFF0CF94CFD3AF8D5" blockId="7.[130,775,1715,2028]" lastBlockId="8.[130,777,1364,2028]" lastPageId="8" lastPageNumber="235" pageId="7" pageNumber="234">
|
||||
The specimen UBB
|
||||
<date id="FFA01069FFAEFFE0FED0F94DFE15F914" box="[344,423,1746,1773]" pageId="7" pageNumber="234" value="1045-05">V 1045</date>
|
||||
represents a partial anterior trunk vertebra with the roof of its neural arch broken off (
|
||||
<figureCitation id="13252A2CFFAEFFE0FD18F96CFD49F8F5" box="[656,763,1778,1805]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="7" pageNumber="234">Fig. 3A, B</figureCitation>
|
||||
). The centrum is slightly longer than wide and features a salient and posteroventrally projecting hypapophysis.Two small subcentral foramina are present and the paradiapophyses appear robust and undifferentiated, however their surfaces are strongly eroded. One of the best preserved mid-trunk vertebrae from the CetăŢuia Hill locality is represented by UBB
|
||||
<date id="FFA01069FFAEFFE0FD7BF82CFF76F812" pageId="7" pageNumber="234" value="1046-05">V 1046</date>
|
||||
/1(
|
||||
<figureCitation id="13252A2CFFAEFFE0FF6AF84FFE97F815" box="[226,293,2001,2028]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="7" pageNumber="234">Fig. 3</figureCitation>
|
||||
C-F); however, in the specimen the right prezygapophysis and the left postzygapophysis are completely broken off, whereas the lateral margin of the left prezygapophysis, the ventral margin of the cotylar lip and the posterior tip of the condyle are also missing. The estimated centrum length of the UBB
|
||||
<date id="FFA01069FFAEFFE0FC7CFA6AFBFAF9F4" box="[1012,1096,1523,1550]" pageId="7" pageNumber="234" value="1046-05">V 1046</date>
|
||||
/
|
||||
<specimenCount id="9D18FD20FFAEFFE0FBDAFA6DFB76F9F4" box="[1106,1220,1523,1549]" count="1" pageId="7" pageNumber="234" type="generic">1 specimen</specimenCount>
|
||||
approaches
|
||||
<quantity id="4CE69B4CFFAEFFE0FACFFA6DFA1EF9F7" box="[1351,1452,1523,1550]" metricMagnitude="-3" metricUnit="m" metricValue="2.42" pageId="7" pageNumber="234" unit="mm" value="2.42">2.42 mm</quantity>
|
||||
, whereas its centrum width approaches
|
||||
<quantity id="4CE69B4CFFAEFFE0FB4DF98DFA98F9D7" box="[1221,1322,1555,1582]" metricMagnitude="-3" metricUnit="m" metricValue="2.39" pageId="7" pageNumber="234" unit="mm" value="2.39">2.39 mm</quantity>
|
||||
; in the UBB
|
||||
<date id="FFA01069FFAEFFE0FCA3F9AAFC30F9B4" box="[811,898,1587,1614]" pageId="7" pageNumber="234" value="1047-05">V 1047</date>
|
||||
/
|
||||
<specimenCount id="9D18FD20FFAEFFE0FC04F9ADFBB1F9B4" box="[908,1027,1587,1613]" count="1" pageId="7" pageNumber="234" type="generic">1 specimen</specimenCount>
|
||||
from Suceag 1 locality (
|
||||
<figureCitation id="13252A2CFFAEFFE0FA87F9ADFAFCF9B4" box="[1295,1358,1587,1613]" captionStart="FIG" captionStartId="8.[132,143,1214,1231]" captionTargetBox="[135,1455,217,1172]" captionTargetId="figure-585@8.[747,1389,194,740]" captionTargetPageId="8" captionText="FIG. 4. — Isolated trunk vertebrae of cf. Messelophis variatus: (A-I) and Ungaliophiidae indet. (J-N): A-C, middle trunk vertebra (UBB V 1047/1) in dorsal (A), ventral (B) and right lateral (C) views; D, middle trunk vertebra (UBB V 1047/3) in left lateral view; E, F, middle trunk vertebra (UBB V 1046/4) in dorsal (E) and ventral (F) views; G-I, caudal vertebra (UBB V 1044) in dorsal (G), ventral (H) and posterior (I) views; J-N, caudal vertebra (UBB V 1048) in dorsal (J), ventral (K), right lateral (L), anterior (M) and posterior (N) views. Abbreviation: ha, haemapophysis; hk, haemal keel; ns, neural spine; pla, pleurapophysis. Scale bar: 2 mm." pageId="7" pageNumber="234">Fig. 4</figureCitation>
|
||||
A-C) the centrum length reaches
|
||||
<quantity id="4CE69B4CFFAEFFE0FBA1F9CDFB3DF994" box="[1065,1167,1619,1645]" metricMagnitude="-3" metricUnit="m" metricValue="2.64" pageId="7" pageNumber="234" unit="mm" value="2.64">2.64 mm</quantity>
|
||||
, whereas its centrum width equals
|
||||
<quantity id="4CE69B4CFFAEFFE0FCF0F9EDFC51F974" box="[888,995,1651,1677]" metricMagnitude="-3" metricUnit="m" metricValue="2.46" pageId="7" pageNumber="234" unit="mm" value="2.46">2.46 mm</quantity>
|
||||
. In dorsal view, the most striking feature is that the neural spine is extremely short and point-like, positioned on the upraised dorsoposterior border of the neural arch, whereas the area in front of the neural spine is flat (
|
||||
<figureCitation id="13252A2CFFAEFFE0FCD7F96CFC4BF8F4" box="[863,1017,1778,1805]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="7" pageNumber="234">Figs 3C, G, H</figureCitation>
|
||||
;
|
||||
<figureCitation id="13252A2CFFAEFFE0FB8AF96CFBD7F8F5" box="[1026,1125,1778,1805]" captionStart="FIG" captionStartId="8.[132,143,1214,1231]" captionTargetBox="[135,1455,217,1172]" captionTargetId="figure-585@8.[747,1389,194,740]" captionTargetPageId="8" captionText="FIG. 4. — Isolated trunk vertebrae of cf. Messelophis variatus: (A-I) and Ungaliophiidae indet. (J-N): A-C, middle trunk vertebra (UBB V 1047/1) in dorsal (A), ventral (B) and right lateral (C) views; D, middle trunk vertebra (UBB V 1047/3) in left lateral view; E, F, middle trunk vertebra (UBB V 1046/4) in dorsal (E) and ventral (F) views; G-I, caudal vertebra (UBB V 1044) in dorsal (G), ventral (H) and posterior (I) views; J-N, caudal vertebra (UBB V 1048) in dorsal (J), ventral (K), right lateral (L), anterior (M) and posterior (N) views. Abbreviation: ha, haemapophysis; hk, haemal keel; ns, neural spine; pla, pleurapophysis. Scale bar: 2 mm." pageId="7" pageNumber="234">4A, D, E</figureCitation>
|
||||
); however, in all the specimens the distal tip of the neural spine is broken off (
|
||||
<figureCitation id="13252A2CFFAEFFE0FAA5F88CFA18F8D4" box="[1325,1450,1810,1837]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="7" pageNumber="234">Figs 3G, H</figureCitation>
|
||||
;
|
||||
<figureCitation id="13252A2CFFAEFFE0FCA5F8ACFCE6F8B5" box="[813,852,1842,1868]" captionStart="FIG" captionStartId="8.[132,143,1214,1231]" captionTargetBox="[135,1455,217,1172]" captionTargetId="figure-585@8.[747,1389,194,740]" captionTargetPageId="8" captionText="FIG. 4. — Isolated trunk vertebrae of cf. Messelophis variatus: (A-I) and Ungaliophiidae indet. (J-N): A-C, middle trunk vertebra (UBB V 1047/1) in dorsal (A), ventral (B) and right lateral (C) views; D, middle trunk vertebra (UBB V 1047/3) in left lateral view; E, F, middle trunk vertebra (UBB V 1046/4) in dorsal (E) and ventral (F) views; G-I, caudal vertebra (UBB V 1044) in dorsal (G), ventral (H) and posterior (I) views; J-N, caudal vertebra (UBB V 1048) in dorsal (J), ventral (K), right lateral (L), anterior (M) and posterior (N) views. Abbreviation: ha, haemapophysis; hk, haemal keel; ns, neural spine; pla, pleurapophysis. Scale bar: 2 mm." pageId="7" pageNumber="234">4H</figureCitation>
|
||||
). The interzygapophyseal constriction is moderate and a well-defined posterior notch is present on the neural arch. The anterior margin of the zygosphene is provided with two lateral lobes and with a less developed median lobe (
|
||||
<figureCitation id="13252A2CFFAEFFE0FADEF80CFA19F855" box="[1366,1451,1938,1964]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="7" pageNumber="234">Figs 3C</figureCitation>
|
||||
;
|
||||
<figureCitation id="13252A2CFFAEFFE0FCA5F82CFCE3F835" box="[813,849,1970,1996]" captionStart="FIG" captionStartId="8.[132,143,1214,1231]" captionTargetBox="[135,1455,217,1172]" captionTargetId="figure-585@8.[747,1389,194,740]" captionTargetPageId="8" captionText="FIG. 4. — Isolated trunk vertebrae of cf. Messelophis variatus: (A-I) and Ungaliophiidae indet. (J-N): A-C, middle trunk vertebra (UBB V 1047/1) in dorsal (A), ventral (B) and right lateral (C) views; D, middle trunk vertebra (UBB V 1047/3) in left lateral view; E, F, middle trunk vertebra (UBB V 1046/4) in dorsal (E) and ventral (F) views; G-I, caudal vertebra (UBB V 1044) in dorsal (G), ventral (H) and posterior (I) views; J-N, caudal vertebra (UBB V 1048) in dorsal (J), ventral (K), right lateral (L), anterior (M) and posterior (N) views. Abbreviation: ha, haemapophysis; hk, haemal keel; ns, neural spine; pla, pleurapophysis. Scale bar: 2 mm." pageId="7" pageNumber="234">4A</figureCitation>
|
||||
). In ventral view, the haemal keel is well-defined, more salient in the specimens from CetăŢuia Hill (
|
||||
<figureCitation id="13252A2CFFAEFFE0FA81F84FFAC4F815" box="[1289,1398,2001,2028]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="7" pageNumber="234">Figs 3D, I</figureCitation>
|
||||
;
|
||||
<figureCitation id="13252A2CFFAEFFE0FA08F84CFA10F815" box="[1408,1442,2002,2028]" captionStart="FIG" captionStartId="8.[132,143,1214,1231]" captionTargetBox="[135,1455,217,1172]" captionTargetId="figure-585@8.[747,1389,194,740]" captionTargetPageId="8" captionText="FIG. 4. — Isolated trunk vertebrae of cf. Messelophis variatus: (A-I) and Ungaliophiidae indet. (J-N): A-C, middle trunk vertebra (UBB V 1047/1) in dorsal (A), ventral (B) and right lateral (C) views; D, middle trunk vertebra (UBB V 1047/3) in left lateral view; E, F, middle trunk vertebra (UBB V 1046/4) in dorsal (E) and ventral (F) views; G-I, caudal vertebra (UBB V 1044) in dorsal (G), ventral (H) and posterior (I) views; J-N, caudal vertebra (UBB V 1048) in dorsal (J), ventral (K), right lateral (L), anterior (M) and posterior (N) views. Abbreviation: ha, haemapophysis; hk, haemal keel; ns, neural spine; pla, pleurapophysis. Scale bar: 2 mm." pageId="7" pageNumber="234">4F</figureCitation>
|
||||
), whereas in those from Suceag 1, it is somewhat widened and flattened (
|
||||
<figureCitation id="13252A2CFFA1FFEFFF7BFAEAFEF6FA77" box="[243,324,1396,1422]" captionStart="FIG" captionStartId="8.[132,143,1214,1231]" captionTargetBox="[135,1455,217,1172]" captionTargetId="figure-585@8.[747,1389,194,740]" captionTargetPageId="8" captionText="FIG. 4. — Isolated trunk vertebrae of cf. Messelophis variatus: (A-I) and Ungaliophiidae indet. (J-N): A-C, middle trunk vertebra (UBB V 1047/1) in dorsal (A), ventral (B) and right lateral (C) views; D, middle trunk vertebra (UBB V 1047/3) in left lateral view; E, F, middle trunk vertebra (UBB V 1046/4) in dorsal (E) and ventral (F) views; G-I, caudal vertebra (UBB V 1044) in dorsal (G), ventral (H) and posterior (I) views; J-N, caudal vertebra (UBB V 1048) in dorsal (J), ventral (K), right lateral (L), anterior (M) and posterior (N) views. Abbreviation: ha, haemapophysis; hk, haemal keel; ns, neural spine; pla, pleurapophysis. Scale bar: 2 mm." pageId="8" pageNumber="235">Fig. 4B</figureCitation>
|
||||
). The subcentral foramina are present and of variable size, the subcentral grooves are weakly defined, whereas the paradiapophyses project strongly laterally; the prezygapophyseal process is vestigial (
|
||||
<figureCitation id="13252A2CFFA1FFEFFDAEFA4DFD3FFA14" box="[550,653,1491,1518]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="8" pageNumber="235">Fig. 3I, J</figureCitation>
|
||||
). In lateral view, the vertebrae appear moderately flattened with the neural arch elevated near the neural spine (
|
||||
<figureCitation id="13252A2CFFA1FFEFFDDBF98DFD78F9D4" box="[595,714,1555,1581]" captionStart="FIG" captionStartId="8.[132,143,1214,1231]" captionTargetBox="[135,1455,217,1172]" captionTargetId="figure-585@8.[747,1389,194,740]" captionTargetPageId="8" captionText="FIG. 4. — Isolated trunk vertebrae of cf. Messelophis variatus: (A-I) and Ungaliophiidae indet. (J-N): A-C, middle trunk vertebra (UBB V 1047/1) in dorsal (A), ventral (B) and right lateral (C) views; D, middle trunk vertebra (UBB V 1047/3) in left lateral view; E, F, middle trunk vertebra (UBB V 1046/4) in dorsal (E) and ventral (F) views; G-I, caudal vertebra (UBB V 1044) in dorsal (G), ventral (H) and posterior (I) views; J-N, caudal vertebra (UBB V 1048) in dorsal (J), ventral (K), right lateral (L), anterior (M) and posterior (N) views. Abbreviation: ha, haemapophysis; hk, haemal keel; ns, neural spine; pla, pleurapophysis. Scale bar: 2 mm." pageId="8" pageNumber="235">Fig. 4C, D</figureCitation>
|
||||
). The paradiapophyses are undifferentiated and robustly built, as seen in specimen UBB
|
||||
<date id="FFA01069FFA1FFEFFE24F9CDFDB6F994" box="[428,516,1619,1645]" pageId="8" pageNumber="235" value="1046-05">V 1046</date>
|
||||
/3 (
|
||||
<figureCitation id="13252A2CFFA1FFEFFDA2F9CDFDCAF994" box="[554,632,1619,1645]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="8" pageNumber="235">Fig. 3J</figureCitation>
|
||||
). In anterior view, the neural arch is slightly flattened, the neural spine is relatively low, whereas the cotyle is circular and the paracotylar foramina are lacking; the zygosphenal roof is almost horizontal (
|
||||
<figureCitation id="13252A2CFFA1FFEFFE8EF94CFECDF914" box="[262,383,1746,1773]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="8" pageNumber="235">Fig. 3E, H</figureCitation>
|
||||
). In posterior view, the neural arch appears slightly convex, the condyle is rounded, whereas the parazygantral foramina are lacking (
|
||||
<figureCitation id="13252A2CFFA1FFEFFD84F88CFDC9F8D5" box="[524,635,1810,1837]" captionStart="FIG" captionStartId="7.[132,143,1248,1265]" captionTargetBox="[134,1453,215,1206]" captionTargetId="figure-517@7.[746,1385,196,758]" captionTargetPageId="7" captionText="FIG. 3. — Isolated trunk vertebrae of cf. Messelophis variatus: A, B, anterior trunk vertebra (UBB V 1045) in left lateral (A) and ventral (B) views; C-F, middle trunk vertebra (UBB V 1046/1) in dorsal (C), ventral (D), anterior (E) and posterior views (F); G, H, partial trunk vertebra (UBB V 1046/2) in dorsal (G) and anterior (H) views; I-J, partial trunk vertebra (UBB V 1046/3) in ventral (I) and left ventrolateral (J) views; K, partial trunk vertebra (UBB V 1047/2) in posterior view. Abbreviations: hy, hypapophysis; ns, neural spine. Scale bar: 2 mm." pageId="8" pageNumber="235">Fig. 3F, K</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF616621FFA1FFEFFF0CFB20FA03FAEA" pageId="8" pageNumber="235" startId="8.[132,143,1214,1231]" targetBox="[135,1455,217,1172]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="8BA136A9FFA1FFEFFF0CFB20FA03FAEA" blockId="8.[132,1457,1214,1299]" pageId="8" pageNumber="235">
|
||||
<smallCapsWord id="8D47A075FFA1FFEFFF0CFB20FF2FFB37" baselines="1227,1227" box="[132,157,1214,1231]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="8" pageNumber="235">FIG</smallCapsWord>
|
||||
. 4. — Isolated trunk vertebrae of cf.
|
||||
<taxonomicName id="4C1E4D2AFFA1FFEFFE51FB20FD35FB36" authorityName="Baszio" authorityYear="2004" box="[473,647,1214,1231]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="235" phylum="Chordata" rank="species" species="variatus">
|
||||
<emphasis id="B96AEABBFFA1FFEFFE51FB20FD35FB36" box="[473,647,1214,1231]" italics="true" pageId="8" pageNumber="235">Messelophis variatus</emphasis>
|
||||
</taxonomicName>
|
||||
: (
|
||||
<emphasis id="B96AEABBFFA1FFEFFD10FB20FD00FB36" bold="true" box="[664,690,1214,1231]" pageId="8" pageNumber="235">A -I</emphasis>
|
||||
) and
|
||||
<taxonomicName id="4C1E4D2AFFA1FFEFFD6BFB20FCD2FB36" authorityName="McDowell" authorityYear="1987" box="[739,864,1214,1231]" class="Reptilia" family="Ungaliophiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="235" phylum="Chordata" rank="family">Ungaliophiidae</taxonomicName>
|
||||
indet. (
|
||||
<emphasis id="B96AEABBFFA1FFEFFC28FB20FC72FB36" bold="true" box="[928,960,1214,1231]" pageId="8" pageNumber="235">J -N</emphasis>
|
||||
):
|
||||
<emphasis id="B96AEABBFFA1FFEFFC59FB20FC41FB36" bold="true" box="[977,1011,1214,1231]" pageId="8" pageNumber="235">A -C</emphasis>
|
||||
, middle trunk vertebra (UBB V 1047/1) in dorsal (
|
||||
<emphasis id="B96AEABBFFA1FFEFFA10FB20FA17FB36" bold="true" box="[1432,1445,1214,1231]" pageId="8" pageNumber="235">A</emphasis>
|
||||
), ventral (
|
||||
<emphasis id="B96AEABBFFA1FFEFFF4FFB4BFF66FB1F" bold="true" box="[199,212,1237,1254]" pageId="8" pageNumber="235">B</emphasis>
|
||||
) and right lateral (
|
||||
<emphasis id="B96AEABBFFA1FFEFFEE5FB4BFEC9FB1F" bold="true" box="[365,379,1237,1254]" pageId="8" pageNumber="235">C</emphasis>
|
||||
) views;
|
||||
<emphasis id="B96AEABBFFA1FFEFFE48FB4BFE7CFB1F" bold="true" box="[448,462,1237,1254]" pageId="8" pageNumber="235">D</emphasis>
|
||||
, middle trunk vertebra (UBB V 1047/3) in left lateral view;
|
||||
<emphasis id="B96AEABBFFA1FFEFFC3FFB4BFC71FB1F" bold="true" box="[951,963,1237,1254]" pageId="8" pageNumber="235">E</emphasis>
|
||||
,
|
||||
<emphasis id="B96AEABBFFA1FFEFFC46FB4BFC6BFB1F" bold="true" box="[974,985,1237,1254]" pageId="8" pageNumber="235">F</emphasis>
|
||||
, middle trunk vertebra (UBB V 1046/4) in dorsal (
|
||||
<emphasis id="B96AEABBFFA1FFEFFAF1FB4BFA37FB1F" bold="true" box="[1401,1413,1237,1254]" pageId="8" pageNumber="235">E</emphasis>
|
||||
) and ventral (
|
||||
<emphasis id="B96AEABBFFA1FFEFFF4EFB75FF63FB05" bold="true" box="[198,209,1259,1276]" pageId="8" pageNumber="235">F</emphasis>
|
||||
) views;
|
||||
<emphasis id="B96AEABBFFA1FFEFFE9DFB75FE83FB05" bold="true" box="[277,305,1259,1276]" pageId="8" pageNumber="235">G -I</emphasis>
|
||||
, caudal vertebra (UBB V 1044) in dorsal (
|
||||
<emphasis id="B96AEABBFFA1FFEFFD01FB75FD25FB05" bold="true" box="[649,663,1259,1276]" pageId="8" pageNumber="235">G</emphasis>
|
||||
), ventral (
|
||||
<emphasis id="B96AEABBFFA1FFEFFD61FB75FD45FB05" bold="true" box="[745,759,1259,1276]" pageId="8" pageNumber="235">H</emphasis>
|
||||
) and posterior (
|
||||
<emphasis id="B96AEABBFFA1FFEFFCF2FB75FC32FB05" bold="true" box="[890,896,1259,1276]" pageId="8" pageNumber="235">I</emphasis>
|
||||
) views;
|
||||
<emphasis id="B96AEABBFFA1FFEFFC4BFB75FC51FB05" bold="true" box="[963,995,1259,1276]" pageId="8" pageNumber="235">J -N</emphasis>
|
||||
, caudal vertebra (UBB V 1048) in dorsal (
|
||||
<emphasis id="B96AEABBFFA1FFEFFAB4FB75FAF4FB05" bold="true" box="[1340,1350,1259,1276]" pageId="8" pageNumber="235">J</emphasis>
|
||||
), ventral (
|
||||
<emphasis id="B96AEABBFFA1FFEFFA10FB75FA17FB05" bold="true" box="[1432,1445,1259,1276]" pageId="8" pageNumber="235">K</emphasis>
|
||||
), right lateral (
|
||||
<emphasis id="B96AEABBFFA1FFEFFF63FA9CFF44FAEA" bold="true" box="[235,246,1282,1299]" pageId="8" pageNumber="235">L</emphasis>
|
||||
), anterior (
|
||||
<emphasis id="B96AEABBFFA1FFEFFEC7FA9CFED2FAEA" bold="true" box="[335,352,1282,1299]" pageId="8" pageNumber="235">M</emphasis>
|
||||
) and posterior (
|
||||
<emphasis id="B96AEABBFFA1FFEFFE6BFA9CFE43FAEA" bold="true" box="[483,497,1282,1299]" pageId="8" pageNumber="235">N</emphasis>
|
||||
) views. Abbreviation:
|
||||
<emphasis id="B96AEABBFFA1FFEFFD2FFA9CFD0FFAEA" bold="true" box="[679,701,1282,1299]" pageId="8" pageNumber="235">ha</emphasis>
|
||||
, haemapophysis;
|
||||
<emphasis id="B96AEABBFFA1FFEFFCDEFA9CFCDEFAEA" bold="true" box="[854,876,1282,1299]" pageId="8" pageNumber="235">hk</emphasis>
|
||||
, haemal keel;
|
||||
<emphasis id="B96AEABBFFA1FFEFFC6CFA9CFC4BFAEA" bold="true" box="[996,1017,1282,1299]" pageId="8" pageNumber="235">ns</emphasis>
|
||||
, neural spine;
|
||||
<emphasis id="B96AEABBFFA1FFEFFBFBFA9CFB3CFAEA" bold="true" box="[1139,1166,1282,1299]" pageId="8" pageNumber="235">pla</emphasis>
|
||||
, pleurapophysis. Scale bar: 2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BA136A9FFA1FFEFFF13F8ACFA2EF9F4" blockId="8.[130,777,1364,2028]" lastBlockId="8.[811,1457,1364,1550]" pageId="8" pageNumber="235">
|
||||
A single caudal vertebra (UBB
|
||||
<date id="FFA01069FFA1FFEFFE5FF8ADFD9BF8B5" box="[471,553,1842,1869]" pageId="8" pageNumber="235" value="1044-05">V 1044</date>
|
||||
), representing a small sized individual (however,about five lines of the arrested growths are observed on its prezygapophysis indicating that it was an adult snake), possesses a low neural spine extending from the base of the zygosphene to the posterior notch of the neural arch (
|
||||
<figureCitation id="13252A2CFFA1FFEFFF49F84FFEA7F815" box="[193,277,2001,2028]" captionStart="FIG" captionStartId="8.[132,143,1214,1231]" captionTargetBox="[135,1455,217,1172]" captionTargetId="figure-585@8.[747,1389,194,740]" captionTargetPageId="8" captionText="FIG. 4. — Isolated trunk vertebrae of cf. Messelophis variatus: (A-I) and Ungaliophiidae indet. (J-N): A-C, middle trunk vertebra (UBB V 1047/1) in dorsal (A), ventral (B) and right lateral (C) views; D, middle trunk vertebra (UBB V 1047/3) in left lateral view; E, F, middle trunk vertebra (UBB V 1046/4) in dorsal (E) and ventral (F) views; G-I, caudal vertebra (UBB V 1044) in dorsal (G), ventral (H) and posterior (I) views; J-N, caudal vertebra (UBB V 1048) in dorsal (J), ventral (K), right lateral (L), anterior (M) and posterior (N) views. Abbreviation: ha, haemapophysis; hk, haemal keel; ns, neural spine; pla, pleurapophysis. Scale bar: 2 mm." pageId="8" pageNumber="235">Fig. 4G</figureCitation>
|
||||
). The centrum is longer than wide, whereas the neural arch is depressed possessing a weak interzygapophyseal constriction (
|
||||
<figureCitation id="13252A2CFFA1FFEFFC61FAEAFB8FFA77" box="[1001,1085,1396,1422]" captionStart="FIG" captionStartId="8.[132,143,1214,1231]" captionTargetBox="[135,1455,217,1172]" captionTargetId="figure-585@8.[747,1389,194,740]" captionTargetPageId="8" captionText="FIG. 4. — Isolated trunk vertebrae of cf. Messelophis variatus: (A-I) and Ungaliophiidae indet. (J-N): A-C, middle trunk vertebra (UBB V 1047/1) in dorsal (A), ventral (B) and right lateral (C) views; D, middle trunk vertebra (UBB V 1047/3) in left lateral view; E, F, middle trunk vertebra (UBB V 1046/4) in dorsal (E) and ventral (F) views; G-I, caudal vertebra (UBB V 1044) in dorsal (G), ventral (H) and posterior (I) views; J-N, caudal vertebra (UBB V 1048) in dorsal (J), ventral (K), right lateral (L), anterior (M) and posterior (N) views. Abbreviation: ha, haemapophysis; hk, haemal keel; ns, neural spine; pla, pleurapophysis. Scale bar: 2 mm." pageId="8" pageNumber="235">Fig. 4H</figureCitation>
|
||||
); below the right prezygapophysis, a tiny prezygapophyseal accessory process is preserved, and the prezygapophysis itself is elongated. In ventral view, the pleurapophyses are broken off; however, the remnants of the paired haemapophyses are partially preserved (
|
||||
<figureCitation id="13252A2CFFA1FFEFFAA9FA6DFA3CF9F4" box="[1313,1422,1523,1550]" captionStart="FIG" captionStartId="8.[132,143,1214,1231]" captionTargetBox="[135,1455,217,1172]" captionTargetId="figure-585@8.[747,1389,194,740]" captionTargetPageId="8" captionText="FIG. 4. — Isolated trunk vertebrae of cf. Messelophis variatus: (A-I) and Ungaliophiidae indet. (J-N): A-C, middle trunk vertebra (UBB V 1047/1) in dorsal (A), ventral (B) and right lateral (C) views; D, middle trunk vertebra (UBB V 1047/3) in left lateral view; E, F, middle trunk vertebra (UBB V 1046/4) in dorsal (E) and ventral (F) views; G-I, caudal vertebra (UBB V 1044) in dorsal (G), ventral (H) and posterior (I) views; J-N, caudal vertebra (UBB V 1048) in dorsal (J), ventral (K), right lateral (L), anterior (M) and posterior (N) views. Abbreviation: ha, haemapophysis; hk, haemal keel; ns, neural spine; pla, pleurapophysis. Scale bar: 2 mm." pageId="8" pageNumber="235">Fig. 4H, I</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3046522FFA1FFEEFCA5F9AAFB0AFCA9" lastPageId="9" lastPageNumber="236" pageId="8" pageNumber="235" type="discussion">
|
||||
<paragraph id="8BA136A9FFA1FFEFFCA5F9AAFC23F9B5" blockId="8.[811,1457,1588,2028]" box="[813,913,1588,1614]" pageId="8" pageNumber="235">
|
||||
<smallCapsWord id="8D47A075FFA1FFEFFCA5F9AAFC23F9B5" baselines="1607,1607" box="[813,913,1588,1614]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="8" pageNumber="235">REMARKS</smallCapsWord>
|
||||
</paragraph>
|
||||
<paragraph id="8BA136A9FFA1FFEEFCA5F9CDFDC5FBF6" blockId="8.[811,1457,1588,2028]" lastBlockId="9.[130,777,694,2027]" lastPageId="9" lastPageNumber="236" pageId="8" pageNumber="235">
|
||||
Some of the phenotypical features of the vertebrae described above closely resemble those of
|
||||
<taxonomicName id="4C1E4D2AFFA1FFEFFB14F9EDFAC5F974" authorityName="Baszio" authorityYear="2004" box="[1180,1399,1651,1677]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="235" phylum="Chordata" rank="species" species="variatus">
|
||||
<emphasis id="B96AEABBFFA1FFEFFB14F9EDFAC5F974" box="[1180,1399,1651,1677]" italics="true" pageId="8" pageNumber="235">Messelophis variatus</emphasis>
|
||||
</taxonomicName>
|
||||
(e.g. presence of an extremely short “point-like” neural spine, developed near the posterior border of the neural arch and the paracotylar foramina absent). Similar to the specimens of our study, the vertebrae of
|
||||
<taxonomicName id="4C1E4D2AFFA1FFEFFBEBF96DFB6EF8F5" authorityName="Baszio" authorityYear="2004" box="[1123,1244,1779,1804]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="235" phylum="Chordata" rank="species" species="variatus">
|
||||
<emphasis id="B96AEABBFFA1FFEFFBEBF96DFB6EF8F5" box="[1123,1244,1779,1804]" italics="true" pageId="8" pageNumber="235">M. variatus</emphasis>
|
||||
</taxonomicName>
|
||||
are elongated, with extremely short neural spines developed at the posterior border of the neural arch, the paracotylar foramina are absent and the prezygapophyseal accessory processes are weakly developed (
|
||||
<bibRefCitation id="EF8F4B58FFA1FFEFFCBEF8ECFC08F875" author="BASZIO S." box="[822,954,1906,1932]" pageId="8" pageNumber="235" pagination="47 - 66" refId="ref7068" refString="BASZIO S. 2004. - Messelophis variatus n. gen. n. sp., from the Eocene of Messel: a tropidopheine snake with affinities to Erycinae (Boidae). Courier Forschungsinstitut Senckenberg 252: 47-66." type="journal article" year="2004">Baszio 2004</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EF8F4B58FFA1FFEFFC42F8ECFB08F875" author="SCHAAL S. & BASZIO S." box="[970,1210,1906,1932]" pageId="8" pageNumber="235" pagination="67 - 77" refId="ref9491" refString="SCHAAL S. & BASZIO S. 2004. - Messelophis ermannorum n. sp., eine neue Zwergboa (Serpentes: Boidae: Tropidopheinae) aus dem Mittel-Eozan von Messel. Courier Forschungsinstitut Senckenberg 252: 67-77." type="journal article" year="2004">Schaal & Baszio 2004</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EF8F4B58FFA1FFEFFB41F8ECFA1AF875" author="SCANFERLA A. & SMITH K. T. & SCHAAL S. F. K." box="[1225,1448,1906,1932]" pageId="8" pageNumber="235" pagination="182 - 206" refId="ref9420" refString="SCANFERLA A., SMITH K. T. & SCHAAL S. F. K. 2016. - Revision of the cranial anatomy and phylogenetic relationships of the Eocene minute boas Messelophis variatus and Messelophis ermannorum (Serpentes, Booidea). Zoological Journal of the Linnean Society 176 (1): 182-206. https://doi.org/10.1111/zoj.12300" type="journal article" year="2016">
|
||||
Scanferla
|
||||
<emphasis id="B96AEABBFFA1FFEFFABCF8EDFADAF875" box="[1332,1384,1906,1932]" italics="true" pageId="8" pageNumber="235">et al.</emphasis>
|
||||
2016
|
||||
</bibRefCitation>
|
||||
: fig. 1B). Moreover, in the
|
||||
<typeStatus id="54A5880BFFA1FFEFFBDDF80CFB04F855" box="[1109,1206,1938,1964]" pageId="8" pageNumber="235" type="holotype">holotype</typeStatus>
|
||||
of
|
||||
<taxonomicName id="4C1E4D2AFFA1FFEFFB69F80DFAEDF855" authorityName="Baszio" authorityYear="2004" box="[1249,1375,1939,1964]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="235" phylum="Chordata" rank="species" species="variatus">
|
||||
<emphasis id="B96AEABBFFA1FFEFFB69F80DFAEDF855" box="[1249,1375,1939,1964]" italics="true" pageId="8" pageNumber="235">M. variatus</emphasis>
|
||||
</taxonomicName>
|
||||
(SMF-ME 1828 A-B), exposing a short portion of the ventral side of its vertebral column (MV pers. obs.), and in the recently referred specimen SMF-ME 513a from the early middle Eocene of Messel,
|
||||
<collectingCountry id="F3097639FFA0FFEEFEC7FD48FE04FD09" box="[335,438,726,752]" name="Germany" pageId="9" pageNumber="236">Germany</collectingCountry>
|
||||
, exposing the ventral side of its vertebral column (
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFEDAFD68FDC3FCE9" author="SCANFERLA A. & SMITH K. T." box="[338,625,758,784]" pageId="9" pageNumber="236" pagination="615 - 620" refId="ref9359" refString="SCANFERLA A. & SMITH K. T. 2020 b. - Additional anatomical information on the Eocene minute boas Messelophis variatus and Rieppelophis ermannorum (Messel Formation, Germany). Vertebrate Zoology 70 (4): 615-620. https://doi.org/10.26049/VZ70-4-2020-06" type="journal article" year="2020">Scanferla & Smith 2020b</bibRefCitation>
|
||||
: fig. 1B), the trunk vertebrae possess prominent haemal keels of variable shape (i.e., of gladiate or spatulate-shape,
|
||||
<emphasis id="B96AEABBFFA0FFEEFDDAFCA8FD35FCB6" box="[594,647,822,847]" italics="true" pageId="9" pageNumber="236">sensu</emphasis>
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFD06FCA8FF0FFC96" author="AUFFENBERG W." pageId="9" pageNumber="236" pagination="131 - 216" refId="ref6982" refString="AUFFENBERG W. 1963. - The fossil snakes of Florida. Tulane Studies in Zoology 10: 131-216. https://doi.org/10.5962/bhl.part.4641" type="journal article" year="1963">Auffenberg 1963</bibRefCitation>
|
||||
). The caudal vertebra UBB
|
||||
<date id="FFA01069FFA0FFEEFE68FCC8FD80FC89" box="[480,562,854,880]" pageId="9" pageNumber="236" value="1044-05">V 1044</date>
|
||||
, similarly to
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFD3EFCC8FF04FC76" authorityName="Baszio" authorityYear="2004" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="species" species="variatus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFD3EFCC8FF04FC76" italics="true" pageId="9" pageNumber="236">M. variatus</emphasis>
|
||||
</taxonomicName>
|
||||
, possesses a low neural spine and provided with paired haemapophyses (
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFEC9FC0BFD96FC56" author="SCANFERLA A. & SMITH K. T. & SCHAAL S. F. K." box="[321,548,917,943]" pageId="9" pageNumber="236" pagination="182 - 206" refId="ref9420" refString="SCANFERLA A., SMITH K. T. & SCHAAL S. F. K. 2016. - Revision of the cranial anatomy and phylogenetic relationships of the Eocene minute boas Messelophis variatus and Messelophis ermannorum (Serpentes, Booidea). Zoological Journal of the Linnean Society 176 (1): 182-206. https://doi.org/10.1111/zoj.12300" type="journal article" year="2016">
|
||||
Scanferla
|
||||
<emphasis id="B96AEABBFFA0FFEEFE25FC08FE51FC56" box="[429,483,917,943]" italics="true" pageId="9" pageNumber="236">et al.</emphasis>
|
||||
2016
|
||||
</bibRefCitation>
|
||||
: fig. 18A); however, in
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFF2AFC28FEAFFC36" authorityName="Baszio" authorityYear="2004" box="[162,285,950,975]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="species" species="variatus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFF2AFC28FEAFFC36" box="[162,285,950,975]" italics="true" pageId="9" pageNumber="236">M. variatus</emphasis>
|
||||
</taxonomicName>
|
||||
the haemapophyses appear laminar, whereas in UBB
|
||||
<date id="FFA01069FFA0FFEEFF6BFC48FE85FC16" box="[227,311,981,1008]" pageId="9" pageNumber="236" value="1044-05">V 1044</date>
|
||||
the remnants of the latter structure comparatively are less elongated anteroposteriorly.
|
||||
</paragraph>
|
||||
<caption id="DF616621FFA0FFEEFF0BFDD0FD04FD8C" pageId="9" pageNumber="236" startId="9.[131,142,590,607]" targetBox="[132,1452,215,558]" targetPageId="9" targetType="figure">
|
||||
<paragraph id="8BA136A9FFA0FFEEFF0BFDD0FD04FD8C" blockId="9.[131,1456,590,629]" pageId="9" pageNumber="236">
|
||||
<smallCapsWord id="8D47A075FFA0FFEEFF0BFDD0FF2FFDA7" baselines="603,603" box="[131,157,590,607]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="9" pageNumber="236">FIG</smallCapsWord>
|
||||
. 5. — Isolated trunk vertebra of Alethinophidia indet.:
|
||||
<emphasis id="B96AEABBFFA0FFEEFDDFFDD0FDCBFDA6" bold="true" box="[599,633,590,607]" pageId="9" pageNumber="236">A -D</emphasis>
|
||||
, incomplete middle trunk vertebra (UBB V 1049) in dorsal (
|
||||
<emphasis id="B96AEABBFFA0FFEEFBD1FDD0FBD4FDA6" bold="true" box="[1113,1126,590,607]" pageId="9" pageNumber="236">A</emphasis>
|
||||
), ventral (
|
||||
<emphasis id="B96AEABBFFA0FFEEFB3DFDD0FB70FDA6" bold="true" box="[1205,1218,590,607]" pageId="9" pageNumber="236">B</emphasis>
|
||||
), right lateral (
|
||||
<emphasis id="B96AEABBFFA0FFEEFABDFDD0FAF1FDA6" bold="true" box="[1333,1347,590,607]" pageId="9" pageNumber="236">C</emphasis>
|
||||
) and anterior (
|
||||
<emphasis id="B96AEABBFFA0FFEEFF01FDFAFF25FD8C" bold="true" box="[137,151,612,629]" pageId="9" pageNumber="236">D</emphasis>
|
||||
) views. Abbreviation:
|
||||
<emphasis id="B96AEABBFFA0FFEEFEC6FDFAFED8FD8C" bold="true" box="[334,362,612,629]" pageId="9" pageNumber="236">paf</emphasis>
|
||||
, paracotylar foramen. Scale bar: 2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BA136A9FFA0FFEEFF13FB8BFB0AFCA9" blockId="9.[130,777,694,2027]" lastBlockId="9.[814,1457,694,848]" pageId="9" pageNumber="236">
|
||||
A number of comparable attributes of the studied specimens should be noted also in the fossil genus
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFDA5FBABFD58FBB6" authority="Hecht" authorityName="Hecht" authorityYear="1959" box="[557,746,1077,1103]" class="Reptilia" family="Tropidophiidae" genus="Dunnophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFDA5FBABFD12FBB6" box="[557,672,1077,1103]" italics="true" pageId="9" pageNumber="236">Dunnophis</emphasis>
|
||||
Hecht
|
||||
</taxonomicName>
|
||||
<emphasis id="B96AEABBFFA0FFEEFD79FBA8FCB5FBB6" box="[753,775,1078,1103]" italics="true" pageId="9" pageNumber="236">in</emphasis>
|
||||
McGrew
|
||||
<emphasis id="B96AEABBFFA0FFEEFF62FBCBFEAFFB96" box="[234,285,1109,1135]" italics="true" pageId="9" pageNumber="236">et al.</emphasis>
|
||||
, 1959, which was originally designated based on isolated vertebrae as an
|
||||
<emphasis id="B96AEABBFFA0FFEEFE22FBEBFD82FB77" box="[426,560,1140,1166]" italics="true" pageId="9" pageNumber="236">incertae sedis</emphasis>
|
||||
snake from the late early Eocene of North America (
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFE6AFB0AFDD7FB56" author="HECHT M. K." box="[482,613,1172,1199]" pageId="9" pageNumber="236" pagination="130 - 146" refId="ref7955" refString="HECHT M. K. 1959. - Amphibians and Reptiles, in MCGREW P. O., BERMAN J. E., HECHT M. K., HUMMEL J. M., SIMPSON G. G. & WOOD A. E. (eds), The geology and paleontology of the Elk Mountain and Tabernacle Butte area, Wyoming. Bulletin of American Museum of Natural History 117: 130-146." type="journal article" year="1959">Hecht 1959</bibRefCitation>
|
||||
), and recorded later from the Eocene of Western Europe (e.g.
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFD04FB2BFD4DFB37" author="RAGE J. - C." box="[652,767,1204,1231]" pageId="9" pageNumber="236" pagination="76 - 78" refId="ref8844" refString="RAGE J. - C. 1973. - Presence de Dunnophis (Reptilia, Serpentes) dans l'Eocene et l'Oligocene europeens. Bulletin de la Societe Geologique de France 15 (3 - 4): 76-78." type="journal article" year="1973">Rage 1973</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFF0CFB4AFF0EFB17" author="RAGE J. - C." box="[132,188,1236,1262]" pageId="9" pageNumber="236" pagination="274 - 303" refId="ref8885" refString="RAGE J. - C. 1974. - Les Serpents des Phosphorites du Quercy. Palaeovertebrata 6: 274-303." type="journal article" year="1974">1974</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFF42FB4AFEB0FB17" author="RAGE J. - C." box="[202,258,1236,1262]" pageId="9" pageNumber="236" pagination="1 - 80" refId="ref8908" refString="RAGE J. - C. 1984. - Serpentes, in WELLNHOFER P. (ed.), Encyclopedia of Paleoherpetology. Part 11. Gustav Fischer, Stuttgart, New York: 1-80." type="book chapter" year="1984">1984</bibRefCitation>
|
||||
, 2006;
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFEDEFB4BFD90FB17" author="RAGE J. - C. & FORD R. L. E." box="[342,546,1236,1263]" pageId="9" pageNumber="236" pagination="47 - 60" refId="ref9124" refString="RAGE J. - C. & FORD R. L. E. 1980. - Amphibians and Squamates from the Upper Eocene of the Isle of Wight. Tertiary Research 3: 47-60." type="journal article" year="1980">Rage & Ford 1980</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFDB8FB4BFCB2FB17" author="RAGE J. - C. & AUGE M." box="[560,768,1236,1263]" pageId="9" pageNumber="236" pagination="103 - 116" refId="ref9025" refString="RAGE J. - C. & AUGE M. 2003. - Amphibians and squamate reptiles from the lower Eocene of Silveirinha (Portugal). Ciencias da Terra, Universidade Nova de Lisboa 15: 103-116." type="journal article" year="2003">Rage & Augé 2003</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFF0CFB6AFF0EFAF7" author="RAGE J. - C. & AUGE M." box="[132,188,1268,1294]" pageId="9" pageNumber="236" pagination="253 - 268" refId="ref9066" refString="RAGE J. - C. & AUGE M. 2010. - Squamate reptiles from the middle Eocene of Lissieu (France). A landmark in the middle Eocene of Europe. Geobios 43 (2): 253-268. https://doi.org/10.1016/j.geobios.2009.08.002" type="journal article" year="2010">2010</bibRefCitation>
|
||||
; see
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFF79FB6AFE4CFAF7" author="SMITH K. T. & GEORGALIS G. L." box="[241,510,1268,1294]" pageId="9" pageNumber="236" pagination="55 - 84" refId="ref9597" refString="SMITH K. T. & GEORGALIS G. L. 2022. - The diversity and distribution of Palaeogene snakes: a review with comments on vertebral sufficiency, in GOWER D. & ZAHER H. (eds), The Origin and Early Evolution of Snakes. Cambridge University Press, Cambridge: 55-84." type="book chapter" year="2022">Smith & Georgalis 2022</bibRefCitation>
|
||||
), and even from the late Palaeocene of
|
||||
<collectingCountry id="F3097639FFA0FFEEFEABFA8BFE34FAD6" box="[291,390,1301,1327]" name="Morocco" pageId="9" pageNumber="236">Morocco</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFE10FA8AFDC0FAD7" author="AUGE M. & RAGE J. - C." box="[408,626,1300,1327]" pageId="9" pageNumber="236" pagination="235 - 253" refId="ref7018" refString="AUGE M. & RAGE J. - C. 2006. - Herpetofaunas from the upper Paleocene and lower Eocene of Morocco. Annales de Paleontologie 92 (3): 235-253. https://doi.org/10.1016/j.annpal.2005.09.001" type="journal article" year="2006">Augé & Rage 2006</bibRefCitation>
|
||||
). Among the peculiar features of
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFEC5FAAAFE0DFAB7" authorityName="Hecht" authorityYear="1959" box="[333,447,1332,1358]" class="Reptilia" family="Tropidophiidae" genus="Dunnophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFEC5FAAAFE0DFAB7" box="[333,447,1332,1358]" italics="true" pageId="9" pageNumber="236">Dunnophis</emphasis>
|
||||
</taxonomicName>
|
||||
, as noted by the above authors, are the following: the centrum is depressed, narrow and longer than wide; the extremely short neural spine is developed near the margin of the deep posterior notch of the neural arch; the prezygapophyseal processes are absent or vestigial; and the caudal vertebrae, rarely present in the fossil material, likely possessed haemapophyses, as seen in the North American
|
||||
<typeStatus id="54A5880BFFA0FFEEFD0FFA6AFD07F9F7" box="[647,693,1524,1550]" pageId="9" pageNumber="236">type</typeStatus>
|
||||
species of the genus,
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFE85F98DFDFEF9D4" authority="Hecht" authorityName="Hecht" authorityYear="1959" box="[269,588,1555,1581]" class="Reptilia" family="Tropidophiidae" genus="Dunnophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="species" species="microechinis">
|
||||
<emphasis id="B96AEABBFFA0FFEEFE85F98DFDB0F9D4" box="[269,514,1555,1581]" italics="true" pageId="9" pageNumber="236">Dunnophis microechinis</emphasis>
|
||||
Hecht
|
||||
</taxonomicName>
|
||||
<emphasis id="B96AEABBFFA0FFEEFDC7F98AFDD7F9D4" box="[591,613,1556,1581]" italics="true" pageId="9" pageNumber="236">in</emphasis>
|
||||
McGrew
|
||||
<emphasis id="B96AEABBFFA0FFEEFD45F98AFD4DF9D4" box="[717,767,1555,1581]" italics="true" pageId="9" pageNumber="236">et al.</emphasis>
|
||||
, 1959 (see
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFF64F9ADFED8F9B7" author="HECHT M. K." box="[236,362,1587,1614]" pageId="9" pageNumber="236" pagination="130 - 146" refId="ref7955" refString="HECHT M. K. 1959. - Amphibians and Reptiles, in MCGREW P. O., BERMAN J. E., HECHT M. K., HUMMEL J. M., SIMPSON G. G. & WOOD A. E. (eds), The geology and paleontology of the Elk Mountain and Tabernacle Butte area, Wyoming. Bulletin of American Museum of Natural History 117: 130-146." type="journal article" year="1959">Hecht 1959</bibRefCitation>
|
||||
: plate 56: 7-10). Resemblance between
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFF0AF9CDFF47F994" authorityName="Hecht" authorityYear="1959" box="[130,245,1619,1645]" class="Reptilia" family="Tropidophiidae" genus="Dunnophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFF0AF9CDFF47F994" box="[130,245,1619,1645]" italics="true" pageId="9" pageNumber="236">Dunnophis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFEA6F9CDFDF2F994" authority="Muller, 1880" authorityName="Muller" authorityYear="1880" box="[302,576,1619,1645]" class="Reptilia" family="Boidae" genus="Ungaliophis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFEA6F9CDFE18F994" box="[302,426,1619,1645]" italics="true" pageId="9" pageNumber="236">Ungaliophis</emphasis>
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFE3AF9CDFDF2F994" author="MULLER F." box="[434,576,1619,1645]" pageId="9" pageNumber="236" pagination="120 - 165" refId="ref8573" refString="MULLER F. 1880. - Erster Nachtrag zum Katalog der herpetologischen Sammlung des Basler Museums. Verhandlungen der Naturforschenden Gesellschaft in Basel 7: 120-165." type="journal article" year="1880">Müller, 1880</bibRefCitation>
|
||||
</taxonomicName>
|
||||
was already noted by
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFF2FF9EDFEF0F974" author="BOGERT C. M." box="[167,322,1651,1677]" pageId="9" pageNumber="236" pagination="1 - 38" refId="ref7111" refString="BOGERT C. M. 1968. - A new genus and species of dwarf boa from Southern Mexico. American Museum Novitates 2354: 1-38. https://www.biodiversitylibrary.org/page/63004213" type="journal article" year="1968">Bogert (1968)</bibRefCitation>
|
||||
, while similarity between
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFDEDF9EDFD6AF974" authorityName="Hecht" authorityYear="1959" box="[613,728,1651,1677]" class="Reptilia" family="Tropidophiidae" genus="Dunnophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFDEDF9EDFD6AF974" box="[613,728,1651,1677]" italics="true" pageId="9" pageNumber="236">Dunnophis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFF0CF90DFF4BF954" authorityName="Baszio" authorityYear="2004" box="[132,249,1683,1709]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFF0CF90DFF4BF954" box="[132,249,1683,1709]" italics="true" pageId="9" pageNumber="236">Messelophis</emphasis>
|
||||
</taxonomicName>
|
||||
was mentioned by
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFE58F90DFDD4F954" author="BASZIO S." box="[464,614,1683,1709]" pageId="9" pageNumber="236" pagination="47 - 66" refId="ref7068" refString="BASZIO S. 2004. - Messelophis variatus n. gen. n. sp., from the Eocene of Messel: a tropidopheine snake with affinities to Erycinae (Boidae). Courier Forschungsinstitut Senckenberg 252: 47-66." type="journal article" year="2004">Baszio (2004)</bibRefCitation>
|
||||
.
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFDFCF90DFF7CF934" author="RAGE J. - C. & AUGE M." pageId="9" pageNumber="236" pagination="253 - 268" refId="ref9066" refString="RAGE J. - C. & AUGE M. 2010. - Squamate reptiles from the middle Eocene of Lissieu (France). A landmark in the middle Eocene of Europe. Geobios 43 (2): 253-268. https://doi.org/10.1016/j.geobios.2009.08.002" type="journal article" year="2010">Rage & Augé (2010)</bibRefCitation>
|
||||
suggested that
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFEEFF92CFE6BF935" authorityName="Baszio" authorityYear="2004" box="[359,473,1714,1740]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFEEFF92CFE6BF935" box="[359,473,1714,1740]" italics="true" pageId="9" pageNumber="236">Messelophis</emphasis>
|
||||
</taxonomicName>
|
||||
might be considered a junior synonym of
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFE87F94CFE33F915" authorityName="Hecht" authorityYear="1959" box="[271,385,1746,1772]" class="Reptilia" family="Tropidophiidae" genus="Dunnophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFE87F94CFE33F915" box="[271,385,1746,1772]" italics="true" pageId="9" pageNumber="236">Dunnophis</emphasis>
|
||||
</taxonomicName>
|
||||
. On the other hand, the specimens from the early Oligocene of
|
||||
<collectingCountry id="F3097639FFA0FFEEFE27F96CFDA3F8F5" box="[431,529,1778,1804]" name="Romania" pageId="9" pageNumber="236">Romania</collectingCountry>
|
||||
described herein, share with the two recent genera of ungaliophiids (i.e.,
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFD39F88CFEA6F8B5" authority="Bogert, 1968" authorityName="Bogert" authorityYear="1968" class="Reptilia" family="Boidae" genus="Exiliboa" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFD39F88CFCBBF8D5" box="[689,777,1810,1836]" italics="true" pageId="9" pageNumber="236">Exiliboa</emphasis>
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFF0CF8ADFEA6F8B5" author="BOGERT C. M." box="[132,276,1842,1869]" pageId="9" pageNumber="236" pagination="1 - 38" refId="ref7111" refString="BOGERT C. M. 1968. - A new genus and species of dwarf boa from Southern Mexico. American Museum Novitates 2354: 1-38. https://www.biodiversitylibrary.org/page/63004213" type="journal article" year="1968">Bogert, 1968</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFEC4F8ACFE7AF8B5" authorityName="Muller" authorityYear="1880" box="[332,456,1842,1868]" class="Reptilia" family="Boidae" genus="Ungaliophis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFEC4F8ACFE7AF8B5" box="[332,456,1842,1868]" italics="true" pageId="9" pageNumber="236">Ungaliophis</emphasis>
|
||||
</taxonomicName>
|
||||
) a number of phenotypic features, as follows: lightly built vertebrae with their centrum longer than wide, the presence of a prominent haemal keel in the trunk vertebrae (according to
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFD9FF80CFF71F835" author="SZYNDLAR Z. & GEORGALIS G. L." pageId="9" pageNumber="236" pagination="717 - 886" refId="ref9968" refString="SZYNDLAR Z. & GEORGALIS G. L. 2023. - An illustrated atlas of the vertebral morphology of extant non-caenophidian snakes, with special emphasis on the cloacal and caudal portions of the column. Vertebrate Zoology 73: 717-886. https://doi.org/10.3897/vz.73.e101372" type="journal article" year="2023">Szyndlar & Georgalis, 2023</bibRefCitation>
|
||||
, the haemal keel is less developed in
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFDFEF82CFD44F835" authorityName="Muller" authorityYear="1880" box="[630,758,1970,1996]" class="Reptilia" family="Boidae" genus="Ungaliophis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFA0FFEEFDFEF82CFD44F835" box="[630,758,1970,1996]" italics="true" pageId="9" pageNumber="236">Ungaliophis</emphasis>
|
||||
</taxonomicName>
|
||||
), the absence of paracotylar foramina, and weakly developed or vestigial prezygapophyseal accessory processes. Some of these characters (e.g. elongation of the trunk vertebrae and presence of haemal keel in the caudal vertebrae) have been considered synapomorphies of
|
||||
<taxonomicName id="4C1E4D2AFFA0FFEEFB02FC88FA86FCC9" authorityName="McDowell" authorityYear="1987" box="[1162,1332,790,816]" class="Reptilia" family="Ungaliophiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="236" phylum="Chordata" rank="family">Ungaliophiidae</taxonomicName>
|
||||
within the clade of Constrictores (
|
||||
<bibRefCitation id="EF8F4B58FFA0FFEEFBA0FCA8FB1BFCA9" author="SMITH K. T." box="[1064,1193,822,848]" pageId="9" pageNumber="236" pagination="157 - 182" refId="ref9533" refString="SMITH K. T. 2013. - New constraints on the evolution of the snake clades Ungaliophiinae, Loxocemidae and Colubridae (Serpentes), with comments on the fossil history of erycine boids in North America. Zoologischer Anzeiger 252 (2): 157-182. https://doi.org/10.1016/j.jcz.2012.05.006" type="journal article" year="2013">Smith 2013</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
191
data/03/B7/87/03B787BFFFAFFFE1FE80F9EDFB37F815.xml
Normal file
191
data/03/B7/87/03B787BFFFAFFFE1FE80F9EDFB37F815.xml
Normal file
|
|
@ -0,0 +1,191 @@
|
|||
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<mods:titleInfo id="2C8DC591843297BA32A0CB90CA976EF7">
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<mods:title id="20CEAE2937969A0D55A10402E222B900">Late Eocene-early Oligocene snakes from the Transylvanian Basin (Romania)</mods:title>
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<mods:namePart id="C262EB9798F940C376BDCBA654F5BA58">Venczel, Márton</mods:namePart>
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<mods:affiliation id="BA551C2B83003B011F7801536BCFA1F9">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & Țării CriȘurilor Museum, Natural History Department, 1 / A Armatei Române, Oradea, RO- 410087 (Romania)</mods:affiliation>
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</mods:name>
|
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<mods:namePart id="783843C4A2908326622AA76EA4BAD4D0">Codrea, Vlad A.</mods:namePart>
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<mods:affiliation id="94C829ADF615EE21A21ABDDA4EB17EFA">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & Țării CriȘurilor Museum, Natural History Department, 1 / A Armatei Române, Oradea, RO- 410087 (Romania) & MureȘ County Museum, Natural History Department, 24 Horea Street, Târgu MureȘ, RO- 540036 (Romania) & ’ Emil Racoviță’ Speleological Institute, Calea 13 Septembrie, RO- 050711, Sector 5, Bucharest (Romania)</mods:affiliation>
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<mods:nameIdentifier id="2EB662E0B8128FBC3D429361D001C8C7" type="email">codrea_vlad@yahoo.fr</mods:nameIdentifier>
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<mods:namePart id="68AF0511B995CA7C0425E7997D612765">Solomon, Alexandru A.</mods:namePart>
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<mods:affiliation id="5EDB10892DC71DA1193720C48FAC0C1A">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & MureȘ County Museum, Natural History Department, 24 Horea Street, Târgu MureȘ, RO- 540036 (Romania)</mods:affiliation>
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<mods:namePart id="60ABF103CB051A2C40594E3C470E224D">Fărcaș, Cristina</mods:namePart>
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<mods:namePart id="9608134536DFF6AEE7D995384EC5A6BB">Bordeianu, Marian</mods:namePart>
|
||||
<mods:affiliation id="F65C92D8A76B7036DF515EF7330E9B8D">Scientific and Technological Advanced Research (STAR) Institute, BabeȘ-Bolyai University, Cluj-Napoca, Romania, 1 Kogălniceanu Street, Cluj-Napoca, RO- 400084 (Romania) & Transgex SA, Geological Compartment, 2 Vasile Alecsandri Street, RO- 410072, Oradea (Romania)</mods:affiliation>
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<mods:titleInfo id="038ED08CB8E7D30091E7E775BDAE7204">
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<mods:title id="16D82A731BD415BAD30FE7B416731FDC">Comptes Rendus Palevol</mods:title>
|
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</mods:titleInfo>
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<mods:part id="E91DC5BAF150F158E963C93207A4F030">
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<mods:date id="48792966AFC3C17C1C124CDDAC7C3AB7">2025</mods:date>
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<mods:number id="9655F481F8312F50117A6221C0689A41">2025-05-14</mods:number>
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<mods:detail id="344415194EC9BCD5805F9CB677041C57" type="volume">
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<mods:number id="FE63A2DB39DB327D0730198BDA9CB364">24</mods:number>
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</mods:detail>
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<mods:detail id="959BC65AB9F427747DAE6743FE083D19" type="issue">
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<mods:number id="524544B383109DB18C090E7E89D888D1">13</mods:number>
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<mods:start id="7FF837BEF9A4C703CC49D166C52B996D">229</mods:start>
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<mods:end id="222F26FC43E73E1A4319144903711BFD">240</mods:end>
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<mods:url id="B6CC89227E70B4A7FBD869A0B402C137">https://doi.org/10.5852/cr-palevol2025v24a13</mods:url>
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</mods:location>
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<mods:classification id="9AACAC6D05239E847440FFE8E6F3B957">journal article</mods:classification>
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<mods:identifier id="6FD4BACB3902A55302B25E4F70BE2BAF" type="DOI">10.5852/cr-palevol2025v24a13</mods:identifier>
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<mods:identifier id="13E605BB48363426E6506E84E57EE513" type="ISSN">1777-571X</mods:identifier>
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<mods:identifier id="16D4A7EAEECE3CA9F74F9C8A71378D56" type="Zenodo-Dep">15624605</mods:identifier>
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<treatment id="03B787BFFFAFFFE1FE80F9EDFB37F815" LSID="urn:lsid:plazi:treatment:03B787BFFFAFFFE1FE80F9EDFB37F815" httpUri="http://treatment.plazi.org/id/03B787BFFFAFFFE1FE80F9EDFB37F815" lastPageNumber="233" pageId="6" pageNumber="233">
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<subSubSection id="C3046522FFAFFFE1FE80F9EDFD31F974" box="[264,643,1651,1677]" pageId="6" pageNumber="233" type="nomenclature">
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<paragraph id="8BA136A9FFAFFFE1FE80F9EDFD31F974" blockId="6.[264,643,1651,1677]" box="[264,643,1651,1677]" pageId="6" pageNumber="233">
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<heading id="D0E981C5FFAFFFE1FE80F9EDFD31F974" box="[264,643,1651,1677]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="233" reason="7">
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<taxonomicName id="4C1E4D2AFFAFFFE1FE80F9EDFD31F974" authority="McDowell, 1987" authorityName="McDowell" authorityYear="1987" box="[264,643,1651,1677]" class="Reptilia" family="Ungaliophiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="family">
|
||||
<smallCapsWord id="8D47A075FFAFFFE1FE80F9EDFE70F972" baselines="1670,1670" box="[264,450,1651,1677]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Ungaliophiidae" pageId="6" pageNumber="233">UNGALIOPHIIDAE</smallCapsWord>
|
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<bibRefCitation id="EF8F4B58FFAFFFE1FE41F9EDFD31F974" author="MCDOWELL S. B." box="[457,643,1651,1677]" pageId="6" pageNumber="233" pagination="3 - 50" refId="ref8484" refString="MCDOWELL S. B. 1987. - Systematics, in SEIGEL R. A., COLLINS J. T. & NOVAK S. S. (eds), Snakes: Ecology and Evolutionary Biology. Macmillan, New York: 3-50. https://doi.org/10.2307/1445695" type="book chapter" year="1987">McDowell, 1987</bibRefCitation>
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</taxonomicName>
|
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</heading>
|
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</paragraph>
|
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</subSubSection>
|
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<subSubSection id="C3046522FFAFFFE1FEB9F90DFDE9F954" box="[305,603,1683,1709]" pageId="6" pageNumber="233" type="description">
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<paragraph id="8BA136A9FFAFFFE1FEB9F90DFDE9F954" blockId="6.[305,603,1683,1709]" box="[305,603,1683,1709]" pageId="6" pageNumber="233">
|
||||
(
|
||||
<emphasis id="B96AEABBFFAFFFE1FEB2F90DFEDCF955" box="[314,366,1683,1708]" italics="true" pageId="6" pageNumber="233">sensu</emphasis>
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FEFDF90DFDE0F954" author="BURBRINK F. T. & GRAZZIOTIN F. G. & PYRON R. A. & CUNDALL D. & DONNELLAN S. & IRISH F. & KEOGH J. S. & KRAUS F. & MURPHY R. W. & NOONAN B. & RAXWORTHY C. J. & RUANE S. & LEMMON A. R. & LEMMON E. M. & ZAHER H." box="[373,594,1683,1709]" pageId="6" pageNumber="233" pagination="502 - 520" refId="ref7152" refString="BURBRINK F. T., GRAZZIOTIN F. G., PYRON R. A., CUNDALL D., DONNELLAN S., IRISH F., KEOGH J. S., KRAUS F., MURPHY R. W., NOONAN B., RAXWORTHY C. J., RUANE S., LEMMON A. R., LEMMON E. M. & ZAHER H. 2020. - Interrogating genomic-scale data for Squamata (lizards, snakes, and amphisbaenians) shows no support for key traditional morphological relationships. Systematic Biology 69 (3): 502-520. https://doi.org/10.1093/sysbio/syz062" type="journal article" year="2020">
|
||||
Burbrink
|
||||
<emphasis id="B96AEABBFFAFFFE1FE57F90DFDA1F954" box="[479,531,1683,1709]" italics="true" pageId="6" pageNumber="233">et al.</emphasis>
|
||||
2020
|
||||
</bibRefCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C3046522FFAFFFE1FF0CF94DFB37F815" pageId="6" pageNumber="233" type="discussion">
|
||||
<paragraph id="8BA136A9FFAFFFE1FF0CF94DFF5AF912" blockId="6.[132,782,1747,2028]" box="[132,232,1747,1773]" pageId="6" pageNumber="233">
|
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<smallCapsWord id="8D47A075FFAFFFE1FF0CF94DFF5AF912" baselines="1766,1766" box="[132,232,1747,1773]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="6" pageNumber="233">REMARKS</smallCapsWord>
|
||||
</paragraph>
|
||||
<paragraph id="8BA136A9FFAFFFE1FF0CF96CFB37F815" blockId="6.[132,782,1747,2028]" lastBlockId="6.[811,1457,1523,2028]" pageId="6" pageNumber="233">
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FF0CF96CFE85F8F5" authorityName="McDowell" authorityYear="1987" box="[132,311,1778,1804]" class="Reptilia" family="Ungaliophiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="family">Ungaliophiidae</taxonomicName>
|
||||
were previously classified as a subfamily within
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FF74F88CFE19F8D5" ID-CoL="6273Q" authorityName="Brongersma" authorityYear="1951" box="[252,427,1810,1836]" class="Reptilia" family="Tropidophiidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="family">Tropidophiidae</taxonomicName>
|
||||
(originally Ungaliopheinae of
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FF0CF8ACFEFAF8B5" author="MCDOWELL S. B." box="[132,328,1842,1868]" pageId="6" pageNumber="233" pagination="3 - 50" refId="ref8484" refString="MCDOWELL S. B. 1987. - Systematics, in SEIGEL R. A., COLLINS J. T. & NOVAK S. S. (eds), Snakes: Ecology and Evolutionary Biology. Macmillan, New York: 3-50. https://doi.org/10.2307/1445695" type="book chapter" year="1987">McDowell 1987</bibRefCitation>
|
||||
). However, based on molecular data, ungaliophiids are considered to be much more distantly related to tropidophiids and instead represent distinct booids. This taxonomic classification is usually regarded as a subfamily within Charinaidae Gray, 1849 (
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FDD3F82CFCBCF835" ID-CoL="87CPR" box="[603,782,1970,1996]" class="Reptilia" family="Boidae" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="subFamily" subFamily="Ungaliophiinae">Ungaliophiinae</taxonomicName>
|
||||
of
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FF2DF84CFEDDF815" author="PYRON R. A. & REYNOLDS G. A. & BURBRINK F. T." box="[165,367,2001,2028]" pageId="6" pageNumber="233" pagination="249 - 260" refId="ref8791" refString="PYRON R. A., REYNOLDS G. A. & BURBRINK F. T. 2014. - A taxonomic revision of boas (Serpentes: Boidae). Zootaxa 3846 (2): 249-260. https://doi.org/10.11646/zootaxa.3846.2.5" type="journal article" year="2014">
|
||||
Pyron
|
||||
<emphasis id="B96AEABBFFAFFFE1FF7BF84CFE99F812" box="[243,299,2001,2027]" italics="true" pageId="6" pageNumber="233">et al.</emphasis>
|
||||
2014
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FE08F84FFD13F815" author="GEORGALIS G. L. & SMITH K. T." box="[384,673,2001,2028]" pageId="6" pageNumber="233" pagination="291 - 304" refId="ref7560" refString="GEORGALIS G. L. & SMITH K. T. 2020. - Constrictores Oppel, 1811 - the available name for the taxonomic group uniting boas and pythons. Vertebrate Zoology 70 (3): 291-304. https://doi.org/10.26049/VZ70-3-2020-03" type="journal article" year="2020">Georgalis & Smith 2020</bibRefCitation>
|
||||
), or as a distinct family of Booidea, the
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FB03FA6DFA8BF9F4" authorityName="McDowell" authorityYear="1987" box="[1163,1337,1523,1549]" class="Reptilia" family="Ungaliophiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="family">Ungaliophiidae</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FACEFA6DFC2DF9D4" author="BURBRINK F. T. & GRAZZIOTIN F. G. & PYRON R. A. & CUNDALL D. & DONNELLAN S. & IRISH F. & KEOGH J. S. & KRAUS F. & MURPHY R. W. & NOONAN B. & RAXWORTHY C. J. & RUANE S. & LEMMON A. R. & LEMMON E. M. & ZAHER H." pageId="6" pageNumber="233" pagination="502 - 520" refId="ref7152" refString="BURBRINK F. T., GRAZZIOTIN F. G., PYRON R. A., CUNDALL D., DONNELLAN S., IRISH F., KEOGH J. S., KRAUS F., MURPHY R. W., NOONAN B., RAXWORTHY C. J., RUANE S., LEMMON A. R., LEMMON E. M. & ZAHER H. 2020. - Interrogating genomic-scale data for Squamata (lizards, snakes, and amphisbaenians) shows no support for key traditional morphological relationships. Systematic Biology 69 (3): 502-520. https://doi.org/10.1093/sysbio/syz062" type="journal article" year="2020">
|
||||
Burbrink
|
||||
<emphasis id="B96AEABBFFAFFFE1FCA5F98AFCD2F9D4" box="[813,864,1555,1581]" italics="true" pageId="6" pageNumber="233">et al.</emphasis>
|
||||
2020
|
||||
</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FC24F98DFB6AF9D4" author="SZYNDLAR Z. & GEORGALIS G. L." box="[940,1240,1555,1581]" pageId="6" pageNumber="233" pagination="717 - 886" refId="ref9968" refString="SZYNDLAR Z. & GEORGALIS G. L. 2023. - An illustrated atlas of the vertebral morphology of extant non-caenophidian snakes, with special emphasis on the cloacal and caudal portions of the column. Vertebrate Zoology 73: 717-886. https://doi.org/10.3897/vz.73.e101372" type="journal article" year="2023">Szyndlar & Georgalis 2023</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FB6CF98DFA2DF9D4" author="ZAHER H. & MOHABEY D. M. & GRAZZIOTIN F. G. & MANTILLA J. A. W." box="[1252,1439,1555,1581]" pageId="6" pageNumber="233" pagination="656 - 697" refId="ref10722" refString="ZAHER H., MOHABEY D. M., GRAZZIOTIN F. G. & MANTILLA J. A. W. 2023. - The skull of Sanajeh indicus, a Cretaceous snake with an upper temporal bar, and the origin of ophidian wide-gaped feeding. Zoological Journal of the Linnean Society 197 (3): 656-697. https://doi.org/10.1093/zoolinnean/zlac001" type="journal article" year="2023">
|
||||
Zaher
|
||||
<emphasis id="B96AEABBFFAFFFE1FAA4F98AFAD2F9D4" box="[1324,1376,1555,1581]" italics="true" pageId="6" pageNumber="233">et al.</emphasis>
|
||||
2023
|
||||
</bibRefCitation>
|
||||
). Parsimony analyses have recovered the latter as the sister clade of Charinaidae (
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FBA5F9CDFAA9F994" author="BURBRINK F. T. & GRAZZIOTIN F. G. & PYRON R. A. & CUNDALL D. & DONNELLAN S. & IRISH F. & KEOGH J. S. & KRAUS F. & MURPHY R. W. & NOONAN B. & RAXWORTHY C. J. & RUANE S. & LEMMON A. R. & LEMMON E. M. & ZAHER H." box="[1069,1307,1619,1645]" pageId="6" pageNumber="233" pagination="502 - 520" refId="ref7152" refString="BURBRINK F. T., GRAZZIOTIN F. G., PYRON R. A., CUNDALL D., DONNELLAN S., IRISH F., KEOGH J. S., KRAUS F., MURPHY R. W., NOONAN B., RAXWORTHY C. J., RUANE S., LEMMON A. R., LEMMON E. M. & ZAHER H. 2020. - Interrogating genomic-scale data for Squamata (lizards, snakes, and amphisbaenians) shows no support for key traditional morphological relationships. Systematic Biology 69 (3): 502-520. https://doi.org/10.1093/sysbio/syz062" type="journal article" year="2020">
|
||||
Burbrink
|
||||
<emphasis id="B96AEABBFFAFFFE1FB16F9CAFB64F994" box="[1182,1238,1619,1645]" italics="true" pageId="6" pageNumber="233">et al.</emphasis>
|
||||
2020
|
||||
</bibRefCitation>
|
||||
) (for further details see also
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FC57F9EDFAA8F974" author="SZYNDLAR Z. & GEORGALIS G. L." box="[991,1306,1651,1677]" pageId="6" pageNumber="233" pagination="717 - 886" refId="ref9968" refString="SZYNDLAR Z. & GEORGALIS G. L. 2023. - An illustrated atlas of the vertebral morphology of extant non-caenophidian snakes, with special emphasis on the cloacal and caudal portions of the column. Vertebrate Zoology 73: 717-886. https://doi.org/10.3897/vz.73.e101372" type="journal article" year="2023">Szyndlar & Georgalis 2023</bibRefCitation>
|
||||
).
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FABEF9EDFC0DF954" authority="Baszio, 2004" authorityName="Baszio" authorityYear="2004" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFAFFFE1FABEF9EDFA03F974" box="[1334,1457,1651,1677]" italics="true" pageId="6" pageNumber="233">Messelophis</emphasis>
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FCA5F90DFC0DF954" author="BASZIO S." box="[813,959,1683,1709]" pageId="6" pageNumber="233" pagination="47 - 66" refId="ref7068" refString="BASZIO S. 2004. - Messelophis variatus n. gen. n. sp., from the Eocene of Messel: a tropidopheine snake with affinities to Erycinae (Boidae). Courier Forschungsinstitut Senckenberg 252: 47-66." type="journal article" year="2004">Baszio, 2004</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, is a genus of fossil dwarf boas from the middle Eocene of Messel locality,
|
||||
<collectingCountry id="F3097639FFAFFFE1FB0CF92DFB5DF934" box="[1156,1263,1715,1741]" name="Germany" pageId="6" pageNumber="233">Germany</collectingCountry>
|
||||
, described firstly by
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FCDAF94CFC46F914" author="BASZIO S." box="[850,1012,1746,1773]" pageId="6" pageNumber="233" pagination="47 - 66" refId="ref7068" refString="BASZIO S. 2004. - Messelophis variatus n. gen. n. sp., from the Eocene of Messel: a tropidopheine snake with affinities to Erycinae (Boidae). Courier Forschungsinstitut Senckenberg 252: 47-66." type="journal article" year="2004">Baszio (2004)</bibRefCitation>
|
||||
, based mainly on the morphology of its vertebrae. Another species of the genus,
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FABBF96CFB0BF8D5" authority="Schaal & Baszio, 2004" authorityName="Schaal & Baszio" authorityYear="2004" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="species" species="ermannorum">
|
||||
<emphasis id="B96AEABBFFAFFFE1FABBF96CFC0BF8D5" italics="true" pageId="6" pageNumber="233">Messelophis ermannorum</emphasis>
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FC37F88CFB0BF8D5" author="SCHAAL S. & BASZIO S." box="[959,1209,1810,1836]" pageId="6" pageNumber="233" pagination="67 - 77" refId="ref9491" refString="SCHAAL S. & BASZIO S. 2004. - Messelophis ermannorum n. sp., eine neue Zwergboa (Serpentes: Boidae: Tropidopheinae) aus dem Mittel-Eozan von Messel. Courier Forschungsinstitut Senckenberg 252: 67-77." type="journal article" year="2004">Schaal & Baszio, 2004</bibRefCitation>
|
||||
</taxonomicName>
|
||||
, also from Messel, was later reassigned to a new genus by
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FB27F8ACFA1AF8B5" author="SCANFERLA A. & SMITH K. T. & SCHAAL S. F. K." box="[1199,1448,1842,1868]" pageId="6" pageNumber="233" pagination="182 - 206" refId="ref9420" refString="SCANFERLA A., SMITH K. T. & SCHAAL S. F. K. 2016. - Revision of the cranial anatomy and phylogenetic relationships of the Eocene minute boas Messelophis variatus and Messelophis ermannorum (Serpentes, Booidea). Zoological Journal of the Linnean Society 176 (1): 182-206. https://doi.org/10.1111/zoj.12300" type="journal article" year="2016">
|
||||
Scanferla
|
||||
<emphasis id="B96AEABBFFAFFFE1FA96F8ADFAE1F8B5" box="[1310,1363,1842,1868]" italics="true" pageId="6" pageNumber="233">et al.</emphasis>
|
||||
(2016)
|
||||
</bibRefCitation>
|
||||
:
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FCA5F8CCFA87F895" authority="Scanferla, Smith & Schaal, 2016" authorityName="Scanferla, Smith & Schaal" authorityYear="2016" box="[813,1333,1874,1900]" class="Reptilia" family="Boidae" genus="Rieppelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFAFFFE1FCA5F8CCFC06F895" box="[813,948,1874,1900]" italics="true" pageId="6" pageNumber="233">Rieppelophis</emphasis>
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FC33F8CCFA87F895" author="SCANFERLA A. & SMITH K. T. & SCHAAL S. F. K." box="[955,1333,1874,1900]" pageId="6" pageNumber="233" pagination="182 - 206" refId="ref9420" refString="SCANFERLA A., SMITH K. T. & SCHAAL S. F. K. 2016. - Revision of the cranial anatomy and phylogenetic relationships of the Eocene minute boas Messelophis variatus and Messelophis ermannorum (Serpentes, Booidea). Zoological Journal of the Linnean Society 176 (1): 182-206. https://doi.org/10.1111/zoj.12300" type="journal article" year="2016">Scanferla, Smith & Schaal, 2016</bibRefCitation>
|
||||
</taxonomicName>
|
||||
. The latter genus appears in parsimony analyses as the sister taxon of
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FCC5F80CFC7BF855" authorityName="Baszio" authorityYear="2004" box="[845,969,1938,1964]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFAFFFE1FCC5F80CFC7BF855" box="[845,969,1938,1964]" italics="true" pageId="6" pageNumber="233">Messelophis</emphasis>
|
||||
</taxonomicName>
|
||||
, while
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FBA9F80CFB2FF855" authorityName="Baszio" authorityYear="2004" box="[1057,1181,1938,1964]" class="Reptilia" family="Ungaliophiidae" genus="Messelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFAFFFE1FBA9F80CFB2FF855" box="[1057,1181,1938,1964]" italics="true" pageId="6" pageNumber="233">Messelophis</emphasis>
|
||||
</taxonomicName>
|
||||
+
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FB34F80CFAF1F855" authorityName="Scanferla, Smith & Schaal" authorityYear="2016" box="[1212,1347,1938,1964]" class="Reptilia" family="Boidae" genus="Rieppelophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFAFFFE1FB34F80CFAF1F855" box="[1212,1347,1938,1964]" italics="true" pageId="6" pageNumber="233">Rieppelophis</emphasis>
|
||||
</taxonomicName>
|
||||
has been recovered as the sister taxon of
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FB21F82CFAB5F835" ID-CoL="634XB" authorityName="Bogert" authorityYear="1968" box="[1193,1287,1970,1996]" class="Reptilia" family="Boidae" genus="Exiliboa" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFAFFFE1FB21F82CFAB5F835" box="[1193,1287,1970,1996]" italics="true" pageId="6" pageNumber="233">Exiliboa</emphasis>
|
||||
</taxonomicName>
|
||||
+
|
||||
<taxonomicName id="4C1E4D2AFFAFFFE1FAA0F82CFA1CF835" ID-CoL="84VP" authorityName="Muller" authorityYear="1880" box="[1320,1454,1970,1996]" class="Reptilia" family="Boidae" genus="Ungaliophis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="233" phylum="Chordata" rank="genus">
|
||||
<emphasis id="B96AEABBFFAFFFE1FAA0F82CFA1CF835" box="[1320,1454,1970,1996]" italics="true" pageId="6" pageNumber="233">Ungaliophis</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EF8F4B58FFAFFFE1FCBEF84FFBE5F815" author="SCANFERLA A. & SMITH K. T." box="[822,1111,2001,2028]" pageId="6" pageNumber="233" pagination="100" refId="ref9304" refString="SCANFERLA A. & SMITH K. T. 2020 a. - Exquisitely preserved fossil snakes of Messel: insight into the evolution, biogeography, habitat preferences and sensory ecology of Early Boas. Diversity 12 (3): 100. https://doi.org/10.3390/d12030100" type="journal article" year="2020">Scanferla & Smith 2020a</bibRefCitation>
|
||||
, b).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
59
data/03/F0/DD/03F0DD67D304FFBFFCB0021AFACBFEA4.xml
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59
data/03/F0/DD/03F0DD67D304FFBFFCB0021AFACBFEA4.xml
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<treatment id="03F0DD67D304FFBFFCB0021AFACBFEA4" LSID="urn:lsid:plazi:treatment:03F0DD67D304FFBFFCB0021AFACBFEA4" httpUri="http://treatment.plazi.org/id/03F0DD67D304FFBFFCB0021AFACBFEA4" lastPageNumber="29" pageId="14" pageNumber="29">
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Family
|
||||
<taxonomicName id="4C5917F2D304FFBFFC0B021AFA8DFE84" ID-CoL="7NJFV" authority="Bourguignat, 1883" authorityName="Bourguignat" authorityYear="1883" box="[907,1294,310,336]" class="Gastropoda" family="Ferussaciidae" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="29" phylum="Mollusca" rank="family">Ferussaciidae Bourguignat, 1883</taxonomicName>
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<paragraph id="8BE66C71D304FFBFFCB0027AFACBFEA4" blockId="14.[816,1352,310,369]" box="[816,1352,342,369]" pageId="14" pageNumber="29">Common name: Blind awl snails or pin snails</paragraph>
|
||||
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|
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|
||||
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|
||||
59
data/03/F0/DD/03F0DD67D304FFBFFCB0071AFB4EFBA4.xml
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59
data/03/F0/DD/03F0DD67D304FFBFFCB0071AFB4EFBA4.xml
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<treatment id="03F0DD67D304FFBFFCB0071AFB4EFBA4" LSID="urn:lsid:plazi:treatment:03F0DD67D304FFBFFCB0071AFB4EFBA4" httpUri="http://treatment.plazi.org/id/03F0DD67D304FFBFFCB0071AFB4EFBA4" lastPageNumber="29" pageId="14" pageNumber="29">
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Family
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<taxonomicName id="4C5917F2D304FFBFFC0B071AFB5CFB84" ID-CoL="7NJHB" authority="Pilsbry, 1918" authorityName="Pilsbry" authorityYear="1918" box="[907,1247,1078,1104]" class="Gastropoda" family="Gastrocoptidae" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="29" phylum="Mollusca" rank="family">Gastrocoptidae Pilsbry, 1918</taxonomicName>
|
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<paragraph id="8BE66C71D304FFBFFCB0077AFB4EFBA4" blockId="14.[816,1247,1078,1137]" box="[816,1229,1110,1137]" pageId="14" pageNumber="29">Common name: Minute land snails</paragraph>
|
||||
</subSubSection>
|
||||
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|
||||
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|
||||
59
data/03/F0/DD/03F0DD67D305FFBEFCB0075AFB0DFB64.xml
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59
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59
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59
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59
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59
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<taxonomicName id="4C5917F2D306FFBDFF7206DAFDA6F9C4" ID-CoL="7NGVW" authority="Tryon, 1866" authorityName="Tryon" authorityYear="1866" box="[242,549,1526,1553]" class="Gastropoda" family="Cochliopidae" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="27" phylum="Mollusca" rank="family">Cochliopidae Tryon, 1866</taxonomicName>
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|
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<paragraph id="8BE66C71D306FFBDFF17053AFDEEF9E4" blockId="12.[151,621,1526,1585]" box="[151,621,1558,1585]" pageId="12" pageNumber="27">Common name: Small freshwater snails</paragraph>
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||||
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60
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59
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Family
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|
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63
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63
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Family
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Common name:
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snails
|
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|
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63
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63
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Common name: Toothless
|
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mussels
|
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59
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Family
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59
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59
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Family
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59
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Family
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|
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133
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<mods:namePart id="C43D57803B6ED4875EAF9B642D53CBFC">Rapalai, Boikhutso Lerato</mods:namePart>
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<mods:title id="16DEF90897B8683CA0E7F5F979D2B7C0">Indago: Investigating nature and humanity in Africa</mods:title>
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<mods:part id="C4DF746BC0661131CC841BE3FCAB05BA">
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<mods:date id="3F435742A6481ABCF7C5FF363AA009BF">2024</mods:date>
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<mods:number id="F18854988A68AF2E68C10C81625CBD0E">41</mods:number>
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</mods:mods>
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<treatment id="03F0DD67D31AFFA1FCB0005AFB54FA64" LSID="urn:lsid:plazi:treatment:03F0DD67D31AFFA1FCB0005AFB54FA64" httpUri="http://treatment.plazi.org/id/03F0DD67D31AFFA1FCB0005AFB54FA64" lastPageNumber="31" pageId="16" pageNumber="31">
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<subSubSection id="C3433FFAD31AFFA1FCB0005AFA92FC44" box="[816,1297,885,912]" pageId="16" pageNumber="31" type="nomenclature">
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<paragraph id="8BE66C71D31AFFA1FCB0005AFA92FC44" blockId="16.[816,1437,885,1457]" box="[816,1297,885,912]" pageId="16" pageNumber="31">
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<heading id="D0AEDB1DD31AFFA1FCB0005AFA92FC44" box="[816,1297,885,912]" fontSize="11" level="2" pageId="16" pageNumber="31" reason="4">
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<emphasis id="B92DB063D31AFFA1FCB0005AFA92FC44" bold="true" box="[816,1297,885,912]" pageId="16" pageNumber="31">
|
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62.
|
||||
<taxonomicName id="4C5917F2D31AFFA1FCDC0059FB99FC44" ID-CoL="3RVCF" baseAuthorityName="Ferussac" baseAuthorityYear="1822" box="[860,1050,885,912]" class="Gastropoda" family="Veronicellidae" genus="Laevicaulis" kingdom="Animalia" order="Systellommatophora" pageId="16" pageNumber="31" phylum="Mollusca" rank="species" species="alte">
|
||||
<emphasis id="B92DB063D31AFFA1FCDC0059FB99FC44" bold="true" box="[860,1050,885,912]" italics="true" pageId="16" pageNumber="31">Laevicaulis alte</emphasis>
|
||||
</taxonomicName>
|
||||
(A.
|
||||
<bibRefCitation id="EFC81180D31AFFA1FBCF0059FA8AFC44" author="FERUSSAC, A. E. J. P. F. & D'AUDEBARD, DE" box="[1103,1289,885,912]" pageId="16" pageNumber="31" refId="ref23751" refString="FERUSSAC, A. E. J. P. F., D'AUDEBARD, DE. 1822. Tableaux systematiques des animaux mollusques classes en familles naturelles, dans lesquels on a etabli la concordance de tous les systemes; suivis d'un Prodrome general pour tous les mollusques ou fluviatiles, vivantes ou fossiles. Paris: A. Bertrand. https://biodiversitylibrary.org/page/11057234" type="url" year="1822">Férussac, 1822</bibRefCitation>
|
||||
)
|
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</emphasis>
|
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</heading>
|
||||
</paragraph>
|
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</subSubSection>
|
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<subSubSection id="C3433FFAD31AFFA1FCB000BAFB54FA64" pageId="16" pageNumber="31" type="distribution">
|
||||
<paragraph id="8BE66C71D31AFFA1FCB000BAFC09FC05" blockId="16.[816,1437,885,1457]" pageId="16" pageNumber="31">
|
||||
Distribution and year:
|
||||
<collectingRegion id="499DA293D31AFFA1FBB300BAFB40FC64" box="[1075,1219,918,944]" country="Botswana" name="Francistown" pageId="16" pageNumber="31">Francistown</collectingRegion>
|
||||
2018, 2019 (
|
||||
<bibRefCitation id="EFC81180D31AFFA1FAD800BAFCFAFC04" author="GBIF" pageId="16" pageNumber="31" refId="ref24043" refString="GBIF. 2023 b. Occurrence download. The Global Biodiversity Information Facility. https://www.gbif.org/occurrence/download/0257952-" type="url" year="2023">
|
||||
GBIF 2023
|
||||
<emphasis id="B92DB063D31AFFA1FCEA009AFCFAFC04" box="[874,889,950,976]" italics="true" pageId="16" pageNumber="31">b</emphasis>
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE66C71D31AFFA1FCB000FAFC6BFBE4" blockId="16.[816,1437,885,1457]" pageId="16" pageNumber="31">
|
||||
Geographic range:
|
||||
<collectingCountry id="F34E2CE1D31AFFA1FB9800FBFB0CFC25" box="[1048,1167,983,1009]" name="Botswana" pageId="16" pageNumber="31">Botswana</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F34E2CE1D31AFFA1FB1F00FAFC4EFBC4" name="Democratic Republic of the Congo" pageId="16" pageNumber="31">Democratic Republic of the Congo</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F34E2CE1D31AFFA1FC5900DAFBB5FBC4" box="[985,1078,1014,1040]" name="Malawi" pageId="16" pageNumber="31">Malawi</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F34E2CE1D31AFFA1FBC100DAFB5AFBC4" box="[1089,1241,1014,1040]" name="South Africa" pageId="16" pageNumber="31">South Africa</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F34E2CE1D31AFFA1FB6400DAFAD2FBC4" box="[1252,1361,1014,1040]" name="Tanzania" pageId="16" pageNumber="31">Tanzania</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F34E2CE1D31AFFA1FADD00DBFCD4FBE4" name="Zambia" pageId="16" pageNumber="31">Zambia</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F34E2CE1D31AFFA1FCE3073AFC67FBE4" box="[867,996,1046,1072]" name="Zimbabwe" pageId="16" pageNumber="31">Zimbabwe</collectingCountry>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE66C71D31AFFA1FCB0071AFB95FB84" blockId="16.[816,1437,885,1457]" box="[816,1046,1078,1104]" pageId="16" pageNumber="31">Habitat: Terrestrial.</paragraph>
|
||||
<paragraph id="8BE66C71D31AFFA1FCB0077BFAFDFA84" blockId="16.[816,1437,885,1457]" pageId="16" pageNumber="31">
|
||||
Notes: The species is widely distributed across Southern Africa, occurring in several
|
||||
<typeStatus id="54E2D2D3D31AFFA1FB21075BFB5CFB45" box="[1185,1247,1143,1169]" pageId="16" pageNumber="31">types</typeStatus>
|
||||
of habitats such as the savanna and the equatorial forest (van Bruggen & Appleton 1977), unknown to occur at very high altitudes, and has been introduced in several parts of the world (
|
||||
<bibRefCitation id="EFC81180D31AFFA1FC3407DAFB5DFAC4" author="HERBERT, D. & KILBURN, D." box="[948,1246,1270,1296]" pageId="16" pageNumber="31" refId="ref24573" refString="HERBERT, D. & KILBURN, D. 2004. Field guide to the land snails and slugs of eastern South Africa. Pietermaritzburg: Natal Museum." type="book" year="2004">Herbert & Kilburn 2004</bibRefCitation>
|
||||
). The species is known as an intermediate host of larval nematode parasites affecting vertebrates (
|
||||
<bibRefCitation id="EFC81180D31AFFA1FB00061AFAF2FA84" author="MAHAJAN, R. K. & ALMEIDA, A. J. & SENGUPTA, S. R. & RENAPURKAR, D. M." box="[1152,1393,1334,1360]" pageId="16" pageNumber="31" pagination="669 - 671" refId="ref26282" refString="MAHAJAN, R. K., ALMEIDA, A. J., SENGUPTA, S. R. & RENAPURKAR, D. M. 1992. Seasonal intensity of Angiostrongylus cantonensis in the intermediate host, Laevicaulis alte. International Journal of Parasitology 22 (5): 669-671. https://doi.org/10.1016/0020-7519(92)90017-F" type="journal article" year="1992">
|
||||
Mahajan
|
||||
<emphasis id="B92DB063D31AFFA1FB70061BFAA8FA84" box="[1264,1323,1334,1360]" italics="true" pageId="16" pageNumber="31">et al.</emphasis>
|
||||
1992
|
||||
</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BE66C71D31AFFA1FCB0067BFAFEFAA4" blockId="16.[816,1437,885,1457]" box="[816,1405,1366,1393]" pageId="16" pageNumber="31">
|
||||
<materialsCitation id="3B31662CD31AFFA1FCB0067BFAFAFAA4" box="[816,1401,1366,1393]" country="India" location="Pondicherry" pageId="16" pageNumber="31" specimenCount="1" typeStatus="holotype">
|
||||
<typeStatus id="54E2D2D3D31AFFA1FCB0067BFCE9FAA5" box="[816,874,1367,1393]" pageId="16" pageNumber="31">Type</typeStatus>
|
||||
locality:
|
||||
<collectingCountry id="F34E2CE1D31AFFA1FC59067AFBE9FAA4" box="[985,1130,1366,1392]" name="India" pageId="16" pageNumber="31">Pondicherry</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F34E2CE1D31AFFA1FBF6067AFB31FAA4" box="[1142,1202,1366,1392]" name="India" pageId="16" pageNumber="31">India</collectingCountry>
|
||||
(
|
||||
<bibRefCitation id="EFC81180D31AFFA1FB41067AFAF3FAA4" author="FERUSSAC, A. E. J. P. F. & D'AUDEBARD, DE" box="[1217,1392,1366,1392]" pageId="16" pageNumber="31" refId="ref23751" refString="FERUSSAC, A. E. J. P. F., D'AUDEBARD, DE. 1822. Tableaux systematiques des animaux mollusques classes en familles naturelles, dans lesquels on a etabli la concordance de tous les systemes; suivis d'un Prodrome general pour tous les mollusques ou fluviatiles, vivantes ou fossiles. Paris: A. Bertrand. https://biodiversitylibrary.org/page/11057234" type="url" year="1822">Férussac 1822</bibRefCitation>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE66C71D31AFFA1FCB0065AFBC3FA45" blockId="16.[816,1437,885,1457]" box="[816,1088,1398,1425]" pageId="16" pageNumber="31">
|
||||
Source:
|
||||
<bibRefCitation id="EFC81180D31AFFA1FC11065AFBB8FA45" author="GBIF" box="[913,1083,1398,1425]" pageId="16" pageNumber="31" refId="ref24043" refString="GBIF. 2023 b. Occurrence download. The Global Biodiversity Information Facility. https://www.gbif.org/occurrence/download/0257952-" type="url" year="2023">
|
||||
GBIF (2023
|
||||
<emphasis id="B92DB063D31AFFA1FBA0065AFBACFA44" box="[1056,1071,1398,1424]" italics="true" pageId="16" pageNumber="31">b</emphasis>
|
||||
)
|
||||
</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BE66C71D31AFFA1FCB006BAFB54FA64" blockId="16.[816,1437,885,1457]" box="[816,1239,1430,1457]" pageId="16" pageNumber="31">
|
||||
Conservation status:
|
||||
<collectingRegion id="499DA293D31AFFA1FBA706BBFBD0FA65" box="[1063,1107,1431,1457]" country="Cameroon" name="North" pageId="16" pageNumber="31">Not</collectingRegion>
|
||||
Evaluated.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
138
data/B3/48/87/B34887E9B705D55CFF59F67FB6AFFC81.xml
Normal file
138
data/B3/48/87/B34887E9B705D55CFF59F67FB6AFFC81.xml
Normal file
|
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@ -0,0 +1,138 @@
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<mods:namePart id="7F2CE4A848684D3CE71E62C6012567A3">Channing, Alan</mods:namePart>
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<mods:affiliation id="AE0B7E9564D8BA90421913F90210D155">Unit for Environmental Sciences and Management, North-West University, Potchefstroom, South Africa;</mods:affiliation>
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<mods:number id="E366C41DB0D46E815AC36A985346E07A">73</mods:number>
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<mods:detail id="E09F0C68D3C0EB712DBD38F9B0955A9F" type="issue">
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||||
<mods:location id="9A72CD2036F0F531ED6F597EF90AB495">
|
||||
<mods:url id="149B00DA91E9ED9466B67E934598C47B">https://doi.org/10.1080/21564574.2023.2296654</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="AAD8EFDA142C724C7865CB8F9CA865A1">journal article</mods:classification>
|
||||
<mods:identifier id="9612252FD732AB83035BBA2A13A3AC7B" type="CLB-Dataset">310359</mods:identifier>
|
||||
<mods:identifier id="730A9B9AB8C28364C9A62B4DACFAD1A8" type="DOI">10.1080/21564574.2023.2296654</mods:identifier>
|
||||
<mods:identifier id="00A144D084BCB529B2BBE9E25BA2B839" type="GBIF-Dataset">ef787402-b1fd-44d4-853d-1680553ebb50</mods:identifier>
|
||||
<mods:identifier id="A796779C0DEC60F4509F0ECCB660A457" type="ISSN">2153-3660</mods:identifier>
|
||||
<mods:identifier id="2FA771575B007FDD34E08F1970297B44" type="Zenodo-Dep">15555879</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="B34887E9B705D55CFF59F67FB6AFFC81" ID-DOI="http://doi.org/10.5281/zenodo.15632231" ID-Zenodo-Dep="15632231" LSID="urn:lsid:plazi:treatment:B34887E9B705D55CFF59F67FB6AFFC81" httpUri="http://treatment.plazi.org/id/B34887E9B705D55CFF59F67FB6AFFC81" lastPageNumber="85" pageId="25" pageNumber="85" scope_family="Pyxicephalidae" scope_genus="Pyxicephalus">
|
||||
<subSubSection id="73FB6574B705D55CFF59F67FB62DFEF4" box="[160,1025,314,339]" pageId="25" pageNumber="85" type="nomenclature">
|
||||
<paragraph id="3B5E36FFB705D55CFF59F67FB62DFEF4" blockId="25.[160,1025,314,339]" box="[160,1025,314,339]" pageId="25" pageNumber="85">
|
||||
<emphasis id="0995EAEDB705D55CFF59F67FB62DFEF4" bold="true" box="[160,1025,314,339]" italics="true" pageId="25" pageNumber="85">
|
||||
Key to the described species of southern and eastern
|
||||
<taxonomicName id="FCE14D7CB705D55CFCCDF67FB62DFEF4" ID-CoL="7697" authority="Tschudi, 1838" authorityName="Tschudi" authorityYear="1838" box="[820,1025,314,339]" class="Amphibia" family="Pyxicephalidae" genus="Pyxicephalus" kingdom="Animalia" order="Anura" pageId="25" pageNumber="85" phylum="Chordata" rank="genus">African bullfrogs</taxonomicName>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="73FB6574B705D55CFF59F62BB6AFFC81" pageId="25" pageNumber="85" type="key">
|
||||
<key id="4F7061CEB705D55CFF59F62BB6AFFC81" pageId="25" pageNumber="85">
|
||||
<keyStep id="80152E5AB705D55CFF59F62BB6A8FE0F" pageId="25" pageNumber="85">
|
||||
<paragraph id="3B5E36FFB705D55CFF59F62BB6AFFE21" blockId="25.[160,1156,366,806]" box="[160,1155,366,390]" pageId="25" pageNumber="85">
|
||||
<keyLead id="801095CAB705D55CFF59F62BB6AFFE21" box="[160,1155,366,390]" pageId="25" pageNumber="85">
|
||||
1a. Tympanum larger than eye, known from
|
||||
<collectingCountry id="43F6766FB705D55CFD62F62BB0DFFE21" box="[667,755,366,390]" name="Somalia" pageId="25" pageNumber="85">Somalia</collectingCountry>
|
||||
.........................
|
||||
<taxonomicName id="FCE14D7CB705D55CFC75F62BB6AFFE21" baseAuthorityName="Scortecci" baseAuthorityYear="1933" box="[908,1155,366,390]" class="Amphibia" family="Pyxicephalidae" genus="Pyxicephalus" kingdom="Animalia" order="Anura" pageId="25" pageNumber="85" phylum="Chordata" rank="species" species="obbianus">
|
||||
<emphasis id="0995EAEDB705D55CFC75F62BB6AFFE21" box="[908,1155,366,390]" italics="true" pageId="25" pageNumber="85">Pyxicephalus obbianus</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="3B5E36FFB705D55CFF59F6D5B6A8FE0F" blockId="25.[160,1156,366,806]" box="[160,1156,400,424]" pageId="25" pageNumber="85">
|
||||
<keyLead id="801095CAB705D55CFF59F6D5B6A8FE0F" box="[160,1156,400,424]" pageId="25" pageNumber="85">1b. Tympanum equal or smaller than eye ............................................................................................ 2</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="80152E5AB705D55CFF59F6F6B6AFFDB7" pageId="25" pageNumber="85">
|
||||
<paragraph id="3B5E36FFB705D55CFF59F6F6B6A8FE49" blockId="25.[160,1156,366,806]" pageId="25" pageNumber="85">
|
||||
<keyLead id="801095CAB705D55CFF59F6F6B6A8FE49" pageId="25" pageNumber="85">2a. White mark on tympanum (sometimes faint), dorsal vertebral stripe usually present ......................................................................................................................................................................... 3</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="3B5E36FFB705D55CFF59F6BDB6AFFDB7" blockId="25.[160,1156,366,806]" box="[160,1155,504,528]" pageId="25" pageNumber="85">
|
||||
<keyLead id="801095CAB705D55CFF59F6BDB6AFFDB7" box="[160,1155,504,528]" pageId="25" pageNumber="85">
|
||||
2b. No white mark on tympanum, no dorsal vertebral stripe .........
|
||||
<taxonomicName id="FCE14D7CB705D55CFC7FF6BDB6AFFDB7" authorityName="Tschudi" authorityYear="1838" box="[902,1155,504,528]" class="Amphibia" family="Pyxicephalidae" genus="Pyxicephalus" kingdom="Animalia" order="Anura" pageId="25" pageNumber="85" phylum="Chordata" rank="species" species="adspersus">
|
||||
<emphasis id="0995EAEDB705D55CFC7FF6BDB6AFFDB7" box="[902,1155,504,528]" italics="true" pageId="25" pageNumber="85">Pyxicephalus adspersus</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="80152E5AB705D55CFF59F55EB6AFFDF1" pageId="25" pageNumber="85">
|
||||
<paragraph id="3B5E36FFB705D55CFF59F55EB6A8FD94" blockId="25.[160,1156,366,806]" box="[160,1156,539,563]" pageId="25" pageNumber="85">
|
||||
<keyLead id="801095CAB705D55CFF59F55EB6A8FD94" box="[160,1156,539,563]" pageId="25" pageNumber="85">3a. Longitudinal dorsal ridges present, with speckling between dorsal mottles ................. 4</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="3B5E36FFB705D55CFF59F57BB6AFFDF1" blockId="25.[160,1156,366,806]" box="[160,1155,574,598]" pageId="25" pageNumber="85">
|
||||
<keyLead id="801095CAB705D55CFF59F57BB6AFFDF1" box="[160,1155,574,598]" pageId="25" pageNumber="85">
|
||||
3b. Oval warts on dorsum, no speckles between dorsal mottles ......
|
||||
<taxonomicName id="FCE14D7CB705D55CFC85F57BB6AFFDF1" authorityName="Parry" authorityYear="1982" box="[892,1155,574,598]" class="Amphibia" family="Pyxicephalidae" genus="Pyxicephalus" kingdom="Animalia" order="Anura" pageId="25" pageNumber="85" phylum="Chordata" rank="species" species="angusticeps">
|
||||
<emphasis id="0995EAEDB705D55CFC85F57BB6AFFDF1" box="[892,1155,574,598]" italics="true" pageId="25" pageNumber="85">Pyxicephalus angusticeps</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="80152E5AB705D55CFF59F525B6AFFC81" pageId="25" pageNumber="85">
|
||||
<paragraph id="3B5E36FFB705D55CFF59F525B6A8FD19" blockId="25.[160,1156,366,806]" pageId="25" pageNumber="85">
|
||||
<keyLead id="801095CAB705D55CFF59F525B6A8FD19" pageId="25" pageNumber="85">
|
||||
4a. No pale crossbar between eyes, upper jaw barring absent (or faint), no cream-coloured lateral stripes or mottles, tympanum smaller than eye-tympanum distance ...........................................................................................................................................
|
||||
<taxonomicName id="FCE14D7CB705D55CFC5DF5E3B6A8FD19" box="[932,1156,678,702]" class="Amphibia" family="Pyxicephalidae" genus="Pyxicephalus" kingdom="Animalia" order="Anura" pageId="25" pageNumber="85" phylum="Chordata" rank="species" species="beytelli">
|
||||
<emphasis id="0995EAEDB705D55CFC5DF5E3B6A8FD19" box="[932,1156,678,702]" italics="true" pageId="25" pageNumber="85">Pyxicephalus beytelli</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="3B5E36FFB705D55CFF59F58DB6AFFC81" blockId="25.[160,1156,366,806]" pageId="25" pageNumber="85">
|
||||
<keyLead id="801095CAB705D55CFF59F58DB6AFFC81" pageId="25" pageNumber="85">
|
||||
4b. Pale crossbar between eyes, sometimes incomplete; upper jaw barring present, cream-coloured lateral stripes or mottles present, tympanum larger than eye-tympanum distance ...........................................................................................................................
|
||||
<taxonomicName id="FCE14D7CB705D55CFC4BF44BB6AFFC81" authorityName="Peters" authorityYear="1854" box="[946,1155,782,806]" class="Amphibia" family="Pyxicephalidae" genus="Pyxicephalus" kingdom="Animalia" order="Anura" pageId="25" pageNumber="85" phylum="Chordata" rank="species" species="edulis">
|
||||
<emphasis id="0995EAEDB705D55CFC4BF44BB6AFFC81" box="[946,1155,782,806]" italics="true" pageId="25" pageNumber="85">Pyxicephalus edulis</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</key>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
1524
data/B3/48/87/B34887E9B711D550FF59F390B030FCEB.xml
Normal file
1524
data/B3/48/87/B34887E9B711D550FF59F390B030FCEB.xml
Normal file
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