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<document id="5E0B9AB9680A0EB49019B41E6EC2C3CF" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Perkovsky, Evgeny" docDate="2016" docId="03A187A92929FFE0FF03C7AF984AF971" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Vertheia Jałoszyński &amp; Perkovsky, 2016, gen. n." docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="17" updateTime="1698382676775" updateUser="plazi">
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<mods:title id="510478BECAE48D68CE4A47C81A5E2E94">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
<document id="2A5CFCAB55EA4A3B1FFD1966946C2105" ID="10.11646/zootaxa.4157.1.1" ID-CLB-Dataset="38320" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="diego" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Perkovsky, Evgeny" docDate="2016" docId="03A187A92929FFE0FF03C7AF984AF971" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Vertheia Jałoszyński &amp; Perkovsky, 2016, gen. n." docType="treatment" docVersion="13" lastPageNumber="18" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="17" updateTime="1738750171177" updateUser="ExternalLinkService">
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<mods:title id="C0B4D2534502F558E49F381723A4E834">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:namePart id="83A85A8F0E54F61BF62F759ED422D972">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="B3C0CF74C7B7AF8B5716B97AE1D21680">Perkovsky, Evgeny</mods:namePart>
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<mods:date id="9AA7C2C55B861007D5FED1E4C4708C8B">2016</mods:date>
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<treatment id="03A187A92929FFE0FF03C7AF984AF971" ID-DOI="http://doi.org/10.5281/zenodo.5681735" ID-GBIF-Taxon="124407822" ID-Zenodo-Dep="5681735" LSID="urn:lsid:plazi:treatment:03A187A92929FFE0FF03C7AF984AF971" httpUri="http://treatment.plazi.org/id/03A187A92929FFE0FF03C7AF984AF971" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<treatment id="03A187A92929FFE0FF03C7AF984AF971" ID-DOI="http://doi.org/10.5281/zenodo.5681735" ID-GBIF-Taxon="124407822" ID-Zenodo-Dep="5681735" LSID="urn:lsid:plazi:treatment:03A187A92929FFE0FF03C7AF984AF971" httpUri="http://treatment.plazi.org/id/03A187A92929FFE0FF03C7AF984AF971" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" scope_family="Staphylinidae" scope_order="Coleoptera" scope_tribe="Eutheiini">
<subSubSection id="C31265342929FFE1FF03C7AF9C30FD4D" box="[151,356,642,668]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BB736BF2929FFE1FF03C7AF9C30FD4D" blockId="16.[151,356,642,668]" box="[151,356,642,668]" pageId="16" pageNumber="17">
<heading id="D0FF81D32929FFE1FF03C7AF9C30FD4D" bold="true" box="[151,356,642,668]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B97CEAAD2929FFE1FF03C7AF9C30FD4D" bold="true" box="[151,356,642,668]" pageId="16" pageNumber="17">
†
<taxonomicName id="4C084D3C2929FFE1FF31C7AF9C53FD4D" box="[165,263,642,668]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C2929FFE1FF31C7AF9C53FD4D" ID-CoL="8XQNM" box="[165,263,642,668]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2929FFE1FF31C7AF9C53FD4D" bold="true" box="[165,263,642,668]" italics="true" pageId="16" pageNumber="17">Vertheia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D62929FFE1FE86C7AF9C30FD4D" box="[274,356,642,668]" pageId="16" pageNumber="17" rank="genus">gen. n.</taxonomicNameLabel>
@ -58,7 +59,7 @@
<subSubSection id="C31265342929FFE1FF03C7E79F94FD0E" box="[151,704,713,735]" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BB736BF2929FFE1FF03C7E79F94FD0E" blockId="16.[151,704,713,735]" box="[151,704,713,735]" pageId="16" pageNumber="17">
Type species:
<taxonomicName id="4C084D3C2929FFE1FEB2C7E49F50FD0E" box="[294,516,713,735]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="quadrisetosa">
<taxonomicName id="4C084D3C2929FFE1FEB2C7E49F50FD0E" box="[294,516,713,735]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="quadrisetosa">
<emphasis id="B97CEAAD2929FFE1FEB2C7E49F50FD0E" box="[294,516,713,735]" italics="true" pageId="16" pageNumber="17">Vertheia quadrisetosa</emphasis>
</taxonomicName>
(here designated).
@ -76,180 +77,180 @@ with a pair of pits on posterior portion of frons, each pit connected to oblique
<paragraph id="8BB736BF2929FFE1FF53C6B19E95FC06" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
<emphasis id="B97CEAAD2929FFE1FF53C6B19C0DFC64" bold="true" box="[199,345,924,949]" pageId="16" pageNumber="17">Description.</emphasis>
Body (
<figureCitation id="13332A3A2929FFE1FE3FC6B19F78FC64" box="[427,556,924,949]" captionStart-0="FIGURES 49 – 55" captionStart-1="FIGURES 56 – 60" captionStart-2="FIGURES 61 – 64" captionStartId-0="18.[151,264,1720,1743]" captionStartId-1="19.[151,264,1962,1985]" captionStartId-2="21.[151,264,1846,1869]" captionTargetBox-0="[151,1436,443,1689]" captionTargetBox-1="[151,1436,628,1941]" captionTargetBox-2="[160,1393,591,1818]" captionTargetId-0="figure@18.[151,1436,443,1699]" captionTargetId-1="figure@19.[151,1436,628,1941]" captionTargetId-2="figure@21.[151,1436,591,1824]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="21" captionText-0="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." captionText-1="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." captionText-2="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri-0="https://zenodo.org/record/262420/files/figure.png" httpUri-1="https://zenodo.org/record/262421/files/figure.png" httpUri-2="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Figs 49–64</figureCitation>
<figureCitation id="13332A3A2929FFE1FE3FC6B19F78FC64" box="[427,556,924,949]" captionStart-0="FIGURES 49 – 55" captionStart-1="FIGURES 56 – 60" captionStart-2="FIGURES 61 – 64" captionStartId-0="18.[151,264,1720,1743]" captionStartId-1="19.[151,264,1962,1985]" captionStartId-2="21.[151,264,1846,1869]" captionTargetBox-0="[151,1436,443,1689]" captionTargetBox-1="[151,1436,628,1941]" captionTargetBox-2="[160,1393,591,1818]" captionTargetId-0="figure@18.[151,1436,443,1699]" captionTargetId-1="figure@19.[151,1436,628,1941]" captionTargetId-2="figure@21.[151,1436,591,1824]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="21" captionText-0="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." captionText-1="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." captionText-2="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi-0="http://doi.org/10.5281/zenodo.262420" figureDoi-1="http://doi.org/10.5281/zenodo.262421" figureDoi-2="http://doi.org/10.5281/zenodo.262422" httpUri-0="https://zenodo.org/record/262420/files/figure.png" httpUri-1="https://zenodo.org/record/262421/files/figure.png" httpUri-2="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Figs 49–64</figureCitation>
) elongate and flattened, with distinctly marked constriction between head and pronotum but weakly so between pronotum and elytra, BL below
<quantity id="4CF09B5A2929FFE1FCEDC6ED9EE9FC06" box="[889,957,960,984]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C6C99EA9FB29" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Head capsule (
<figureCitation id="13332A3A2929FFE1FEE7C6C99CADFC2C" box="[371,505,996,1021]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–54</figureCitation>
<figureCitation id="13332A3A2929FFE1FEE7C6C99CADFC2C" box="[371,505,996,1021]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–54</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD93C6C99F1AFC2C" box="[519,590,996,1021]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–63</figureCitation>
<figureCitation id="13332A3A2929FFE1FD93C6C99F1AFC2C" box="[519,590,996,1021]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–63</figureCitation>
) with anterior part (in front of occipital constriction) subtrapezoidal and transverse; eyes located posteriorly, large, strongly convex and moderately coarsely faceted; tempora (
<figureCitation id="13332A3A2929FFE1FA81C12A9838FBF1" box="[1301,1388,1031,1056]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
<figureCitation id="13332A3A2929FFE1FA81C12A9838FBF1" box="[1301,1388,1031,1056]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FAECC12A98C3FBF1" box="[1400,1431,1031,1056]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
<figureCitation id="13332A3A2929FFE1FAECC12A98C3FBF1" box="[1400,1431,1031,1056]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FF03C1009DE7FB95" box="[151,179,1069,1092]" italics="true" pageId="16" pageNumber="17">tm</emphasis>
) barely discernible, very short; occipital constriction (
<figureCitation id="13332A3A2929FFE1FCB5C1019E2EFB94" box="[801,890,1068,1093]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
<figureCitation id="13332A3A2929FFE1FCB5C1019E2EFB94" box="[801,890,1068,1093]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FC1CC1019EF3FB94" box="[904,935,1068,1093]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
<figureCitation id="13332A3A2929FFE1FC1CC1019EF3FB94" box="[904,935,1068,1093]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FC20C1009E8FFB95" box="[948,987,1069,1092]" italics="true" pageId="16" pageNumber="17">occ</emphasis>
) as broad as frons between eyes; frons (
<figureCitation id="13332A3A2929FFE1FF0BC1629DA2FBB9" box="[159,246,1103,1128]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
<figureCitation id="13332A3A2929FFE1FF0BC1629DA2FBB9" box="[159,246,1103,1128]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FE97C1629C76FBB9" box="[259,290,1103,1128]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
<figureCitation id="13332A3A2929FFE1FE97C1629C76FBB9" box="[259,290,1103,1128]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FEBAC17D9C15FBB9" box="[302,321,1104,1128]" italics="true" pageId="16" pageNumber="17">fr</emphasis>
) subtrapezoidal, posteriorly confluent with strongly transverse, subrectangular vertex (
<figureCitation id="13332A3A2929FFE1FA80C1629838FBB9" box="[1300,1388,1103,1128]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
<figureCitation id="13332A3A2929FFE1FA80C1629838FBB9" box="[1300,1388,1103,1128]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FAECC16298C3FBB9" box="[1400,1431,1103,1128]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
<figureCitation id="13332A3A2929FFE1FAECC16298C3FBB9" box="[1400,1431,1103,1128]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FF03C1589DFFFB5D" box="[151,171,1141,1164]" italics="true" pageId="16" pageNumber="17">vt</emphasis>
); frons posteriorly with pair of small but distinct pits (
<figureCitation id="13332A3A2929FFE1FCB1C1599E2AFB5C" box="[805,894,1140,1165]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
<figureCitation id="13332A3A2929FFE1FCB1C1599E2AFB5C" box="[805,894,1140,1165]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FC19C1599EF8FB5C" box="[909,940,1140,1165]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
<figureCitation id="13332A3A2929FFE1FC19C1599EF8FB5C" box="[909,940,1140,1165]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FC2EC1599E8FFB5D" box="[954,987,1140,1164]" italics="true" pageId="16" pageNumber="17">frp</emphasis>
), each posteriorly connected to groove extending obliquely to posterolateral margin of vertex; 'neck' region (
<figureCitation id="13332A3A2929FFE1FC27C1BA995FFB61" box="[947,1035,1175,1200]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
<figureCitation id="13332A3A2929FFE1FC27C1BA995FFB61" box="[947,1035,1175,1200]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FB8DC1BA996CFB61" box="[1049,1080,1175,1200]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
<figureCitation id="13332A3A2929FFE1FB8DC1BA996CFB61" box="[1049,1080,1175,1200]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FBD2C1B4990BFB61" box="[1094,1119,1177,1200]" italics="true" pageId="16" pageNumber="17">nr</emphasis>
) as broad as frons between eyes. Submentum (
<figureCitation id="13332A3A2929FFE1FEE3C1919C9BFB04" box="[375,463,1212,1237]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Fig. 63</figureCitation>
<figureCitation id="13332A3A2929FFE1FEE3C1919C9BFB04" box="[375,463,1212,1237]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Fig. 63</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FE74C1909F59FB05" box="[480,525,1213,1236]" italics="true" pageId="16" pageNumber="17">smn</emphasis>
) subhexagonal, about as broad as long; posterior tentorial pits (
<figureCitation id="13332A3A2929FFE1FA92C191980AFB04" box="[1286,1374,1212,1237]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Fig. 61</figureCitation>
<figureCitation id="13332A3A2929FFE1FA92C191980AFB04" box="[1286,1374,1212,1237]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Fig. 61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FAFBC19098C7FB05" box="[1391,1427,1213,1236]" italics="true" pageId="16" pageNumber="17">ptp</emphasis>
) distinct, located in transverse groove demarcating anteriorly the 'neck' region.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C0299F03FA91" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Antennae (
<figureCitation id="13332A3A2929FFE1FED2C0299C99FACC" box="[326,461,1284,1309]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–54</figureCitation>
<figureCitation id="13332A3A2929FFE1FED2C0299C99FACC" box="[326,461,1284,1309]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–54</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FE48C0299F71FACC" box="[476,549,1284,1309]" captionStart="FIGURES 56 – 60" captionStartId="19.[151,264,1962,1985]" captionTargetBox="[151,1436,628,1941]" captionTargetId="figure@19.[151,1436,628,1941]" captionTargetPageId="19" captionText="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." httpUri="https://zenodo.org/record/262421/files/figure.png" pageId="16" pageNumber="17">56–58</figureCitation>
<figureCitation id="13332A3A2929FFE1FE48C0299F71FACC" box="[476,549,1284,1309]" captionStart="FIGURES 56 – 60" captionStartId="19.[151,264,1962,1985]" captionTargetBox="[151,1436,628,1941]" captionTargetId="figure@19.[151,1436,628,1941]" captionTargetPageId="19" captionText="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." figureDoi="http://doi.org/10.5281/zenodo.262421" httpUri="https://zenodo.org/record/262421/files/figure.png" pageId="16" pageNumber="17">56–58</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FDA1C0299F28FACC" box="[565,636,1284,1309]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62</figureCitation>
<figureCitation id="13332A3A2929FFE1FDA1C0299F28FACC" box="[565,636,1284,1309]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62</figureCitation>
) moderately long and slender, with enlarged scape and pedicel and gradually but weakly thickened distally.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C0609942FA25" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Pronotum (
<figureCitation id="13332A3A2929FFE1FEDCC0619F72FAB4" box="[328,550,1356,1381]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–50, 52–54</figureCitation>
<figureCitation id="13332A3A2929FFE1FEDCC0619F72FAB4" box="[328,550,1356,1381]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–50, 52–54</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FDA1C0619F29FAB4" box="[565,637,1356,1381]" captionStart="FIGURES 56 – 60" captionStartId="19.[151,264,1962,1985]" captionTargetBox="[151,1436,628,1941]" captionTargetId="figure@19.[151,1436,628,1941]" captionTargetPageId="19" captionText="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." httpUri="https://zenodo.org/record/262421/files/figure.png" pageId="16" pageNumber="17">56–57</figureCitation>
<figureCitation id="13332A3A2929FFE1FDA1C0619F29FAB4" box="[565,637,1356,1381]" captionStart="FIGURES 56 – 60" captionStartId="19.[151,264,1962,1985]" captionTargetBox="[151,1436,628,1941]" captionTargetId="figure@19.[151,1436,628,1941]" captionTargetPageId="19" captionText="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." figureDoi="http://doi.org/10.5281/zenodo.262421" httpUri="https://zenodo.org/record/262421/files/figure.png" pageId="16" pageNumber="17">56–57</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD19C0619E56FAB4" box="[653,770,1356,1381]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62, 64</figureCitation>
<figureCitation id="13332A3A2929FFE1FD19C0619E56FAB4" box="[653,770,1356,1381]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62, 64</figureCitation>
) in dorsal view subrectangular with distinctly or slightly concave anterior margin, feebly marked and obtuse-angled anterior pronotal corners, rounded sides, distinct obtuse-angled posterior pronotal corners and arcuate posterior margin, slightly flattened in front of scutellum. Sides with distinct lateral pronotal carinae or edges (
<figureCitation id="13332A3A2929FFE1FD7EC09A9E6AFA01" box="[746,830,1463,1488]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Fig. 64</figureCitation>
<figureCitation id="13332A3A2929FFE1FD7EC09A9E6AFA01" box="[746,830,1463,1488]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">Fig. 64</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FCDFC09A9E39FA01" box="[843,877,1463,1488]" italics="true" pageId="16" pageNumber="17">lpc</emphasis>
); base with shallow lateral antebasal impressions (
<figureCitation id="13332A3A2929FFE1FF0BC0F19DA1FA24" box="[159,245,1500,1525]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
<figureCitation id="13332A3A2929FFE1FF0BC0F19DA1FA24" box="[159,245,1500,1525]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FE95C0F19C74FA24" box="[257,288,1500,1525]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
<figureCitation id="13332A3A2929FFE1FE95C0F19C74FA24" box="[257,288,1500,1525]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FEB8C0F19C04FA24" box="[300,336,1500,1525]" italics="true" pageId="16" pageNumber="17">abi</emphasis>
) and long, faint transverse antebasal groove (
<figureCitation id="13332A3A2929FFE1FCDAC0F19EF1FA24" box="[846,933,1500,1525]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
<figureCitation id="13332A3A2929FFE1FCDAC0F19EF1FA24" box="[846,933,1500,1525]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FC25C0F19E84FA24" box="[945,976,1500,1525]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
<figureCitation id="13332A3A2929FFE1FC25C0F19E84FA24" box="[945,976,1500,1525]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FC48C0F19952FA24" box="[988,1030,1500,1525]" italics="true" pageId="16" pageNumber="17">abg</emphasis>
).
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C0D29FEDF976" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Basisternal part of prosternum (
<figureCitation id="13332A3A2929FFE1FDBDC0D29FD4F9C9" box="[553,640,1535,1560]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 55</figureCitation>
<figureCitation id="13332A3A2929FFE1FDBDC0D29FD4F9C9" box="[553,640,1535,1560]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD18C0D29FFFF9C9" box="[652,683,1535,1560]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">63</figureCitation>
<figureCitation id="13332A3A2929FFE1FD18C0D29FFFF9C9" box="[652,683,1535,1560]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">63</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FD23C0D29F83F9C9" box="[695,727,1535,1560]" italics="true" pageId="16" pageNumber="17">bst</emphasis>
) nearly as long as coxal part, laterally demarcated by complete notosternal sutures (
<figureCitation id="13332A3A2929FFE1FEE8C3099C85F9EC" box="[380,465,1572,1597]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
<figureCitation id="13332A3A2929FFE1FEE8C3099C85F9EC" box="[380,465,1572,1597]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FE4AC3089F56F9ED" box="[478,514,1573,1596]" italics="true" pageId="16" pageNumber="17">nss</emphasis>
); procoxae contiguous, indicating that prosternal process is either absent or very narrow and not elevated. Prothoracic hypomeron (
<figureCitation id="13332A3A2929FFE1FD45C36A9E71F9B1" box="[721,805,1607,1632]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
<figureCitation id="13332A3A2929FFE1FD45C36A9E71F9B1" box="[721,805,1607,1632]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FCA7C36A9E19F9B1" box="[819,845,1607,1632]" italics="true" pageId="16" pageNumber="17">hy</emphasis>
) flat and with distinctly demarcated inner (adcoxal) part by complete and recurved hypomeral ridge (
<figureCitation id="13332A3A2929FFE1FD72C3419E6BF954" box="[742,831,1644,1669]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
<figureCitation id="13332A3A2929FFE1FD72C3419E6BF954" box="[742,831,1644,1669]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FCC5C3419E22F954" box="[849,886,1644,1669]" italics="true" pageId="16" pageNumber="17">hyr</emphasis>
). Shape of inner (adcoxal) hypomeral margin indicates that procoxal sockets are broadly open.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C3999CE0F921" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Mesoscutellum (
<figureCitation id="13332A3A2929FFE1FE16C3999F51F91C" box="[386,517,1716,1741]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53–54</figureCitation>
<figureCitation id="13332A3A2929FFE1FE16C3999F51F91C" box="[386,517,1716,1741]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 53–54</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD86C3999F0EF91C" box="[530,602,1716,1741]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62</figureCitation>
<figureCitation id="13332A3A2929FFE1FD86C3999F0EF91C" box="[530,602,1716,1741]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62</figureCitation>
) well-visible between elytral bases, broadly subtriangular, with indistinct median basal impression.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C3D1982DF8C5" blockId="16.[151,1437,780,2029]" box="[199,1401,1788,1813]" pageId="16" pageNumber="17">
Mesoventrite (
<figureCitation id="13332A3A2929FFE1FEFEC3D19CEFF8C4" box="[362,443,1788,1813]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
<figureCitation id="13332A3A2929FFE1FEFEC3D19CEFF8C4" box="[362,443,1788,1813]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
) with broadly subtrapezoidal and flat mesoventral intercoxal process (
<figureCitation id="13332A3A2929FFE1FB44C3D19877F8C4" box="[1232,1315,1788,1813]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
<figureCitation id="13332A3A2929FFE1FB44C3D19877F8C4" box="[1232,1315,1788,1813]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Fig. 55</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FABBC3D0983DF8C5" box="[1327,1385,1789,1812]" italics="true" pageId="16" pageNumber="17">msvp</emphasis>
).
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C23299B7F851" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Metaventrite (
<figureCitation id="13332A3A2929FFE1FEF2C2329CEBF8E9" box="[358,447,1823,1848]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 55</figureCitation>
<figureCitation id="13332A3A2929FFE1FEF2C2329CEBF8E9" box="[358,447,1823,1848]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FE58C2329CBFF8E9" box="[460,491,1823,1848]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
<figureCitation id="13332A3A2929FFE1FE58C2329CBFF8E9" box="[460,491,1823,1848]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<date id="FFB6107F2929FFE1FE6CC20D9F46F8E9" box="[504,530,1824,1848]" pageId="16" pageNumber="17">
<emphasis id="B97CEAAD2929FFE1FE6CC20D9F46F8E9" box="[504,530,1824,1848]" italics="true" pageId="16" pageNumber="17">v3</emphasis>
</date>
) subquadrate with weakly rounded sides, anteriorly at middle demarcated from mesoventrite; posteriorly strongly concave laterally in admetacoxal regions and medially forming very short and broad metaventral intercoxal process (
<figureCitation id="13332A3A2929FFE1FDD7C24A9FCDF851" box="[579,665,1895,1920]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 55</figureCitation>
<figureCitation id="13332A3A2929FFE1FDD7C24A9FCDF851" box="[579,665,1895,1920]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD31C24A9F90F851" box="[677,708,1895,1920]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
<figureCitation id="13332A3A2929FFE1FD31C24A9F90F851" box="[677,708,1895,1920]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61</figureCitation>
;
<emphasis id="B97CEAAD2929FFE1FD44C2449E52F851" box="[720,774,1897,1920]" italics="true" pageId="16" pageNumber="17">mtvp</emphasis>
) with distinctly concave posterior margin.
</paragraph>
<paragraph id="8BB736BF2929FFE1FF53C2A19C50F83C" blockId="16.[151,1437,780,2029]" pageId="16" pageNumber="17">
Elytra (
<figureCitation id="13332A3A2929FFE1FE89C2A19CA3F874" box="[285,503,1932,1957]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–50, 53–54</figureCitation>
<figureCitation id="13332A3A2929FFE1FE89C2A19CA3F874" box="[285,503,1932,1957]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="16" pageNumber="17">Figs 49–50, 53–54</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FD92C2A19F1AF874" box="[518,590,1932,1957]" captionStart="FIGURES 56 – 60" captionStartId="19.[151,264,1962,1985]" captionTargetBox="[151,1436,628,1941]" captionTargetId="figure@19.[151,1436,628,1941]" captionTargetPageId="19" captionText="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." httpUri="https://zenodo.org/record/262421/files/figure.png" pageId="16" pageNumber="17">56–58</figureCitation>
<figureCitation id="13332A3A2929FFE1FD92C2A19F1AF874" box="[518,590,1932,1957]" captionStart="FIGURES 56 – 60" captionStartId="19.[151,264,1962,1985]" captionTargetBox="[151,1436,628,1941]" captionTargetId="figure@19.[151,1436,628,1941]" captionTargetPageId="19" captionText="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." figureDoi="http://doi.org/10.5281/zenodo.262421" httpUri="https://zenodo.org/record/262421/files/figure.png" pageId="16" pageNumber="17">56–58</figureCitation>
,
<figureCitation id="13332A3A2929FFE1FDC9C2A19F84F874" box="[605,720,1932,1957]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62, 64</figureCitation>
<figureCitation id="13332A3A2929FFE1FDC9C2A19F84F874" box="[605,720,1932,1957]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="16" pageNumber="17">61–62, 64</figureCitation>
) elongate and nearly oval, except for truncated apices; elytral base with foveae absent or rudimentary (indiscernible in studied specimens, in which the elytral base is wellexposed).
</paragraph>
<paragraph id="8BB736BF2928FFE0FF53C5BA9FE9FF61" blockId="17.[151,1437,151,1544]" box="[199,701,151,176]" pageId="17" pageNumber="18">Hind wings not visible in studied specimens.</paragraph>
<paragraph id="8BB736BF2928FFE0FF53C5909FADFECC" blockId="17.[151,1437,151,1544]" pageId="17" pageNumber="18">
Legs (
<figureCitation id="13332A3A2928FFE0FE99C5919C30FF04" box="[269,356,188,213]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="17" pageNumber="18">Figs 55</figureCitation>
<figureCitation id="13332A3A2928FFE0FE99C5919C30FF04" box="[269,356,188,213]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="17" pageNumber="18">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2928FFE0FEE5C5919CEDFF04" box="[369,441,188,213]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">63–64</figureCitation>
<figureCitation id="13332A3A2928FFE0FEE5C5919CEDFF04" box="[369,441,188,213]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">63–64</figureCitation>
) moderately long, slender; pro- and mesocoxae suboval, metacoxae strongly transverse; all trochanters small and subtriangular; femora weakly clavate; tibiae slender, straight or nearly straight; tarsi long and slender, all tarsomeres elongate, slightly flattened.
</paragraph>
<paragraph id="8BB736BF2928FFE0FF53C40A9E61FE59" blockId="17.[151,1437,151,1544]" pageId="17" pageNumber="18">
Abdomen (
<figureCitation id="13332A3A2928FFE0FED7C40A9CCEFE91" box="[323,410,295,320]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="17" pageNumber="18">Figs 55</figureCitation>
<figureCitation id="13332A3A2928FFE0FED7C40A9CCEFE91" box="[323,410,295,320]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="17" pageNumber="18">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2928FFE0FE31C40A9CB8FE91" box="[421,492,295,320]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">62–64</figureCitation>
<figureCitation id="13332A3A2928FFE0FE31C40A9CB8FE91" box="[421,492,295,320]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">62–64</figureCitation>
) protruding from under truncated elytra, six sternites visible (
<figureCitation id="13332A3A2928FFE0FB0EC40A99A5FE91" box="[1178,1265,295,320]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="17" pageNumber="18">Figs 55</figureCitation>
<figureCitation id="13332A3A2928FFE0FB0EC40A99A5FE91" box="[1178,1265,295,320]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="17" pageNumber="18">Figs 55</figureCitation>
,
<figureCitation id="13332A3A2928FFE0FB6FC40A9810FE91" box="[1275,1348,295,320]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">63–64</figureCitation>
<figureCitation id="13332A3A2928FFE0FB6FC40A9810FE91" box="[1275,1348,295,320]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">63–64</figureCitation>
;
<emphasis id="B97CEAAD2928FFE0FADBC40598DFFE91" box="[1359,1419,296,320]" italics="true" pageId="17" pageNumber="18">st3–8</emphasis>
), sternite III only slightly longer than IV, sternites VII and VIII longest, suture between them partly indistinct. Pygidium (
<figureCitation id="13332A3A2928FFE0FE87C4429C32FE59" box="[275,358,367,392]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">Fig. 62</figureCitation>
<figureCitation id="13332A3A2928FFE0FE87C4429C32FE59" box="[275,358,367,392]" captionStart="FIGURES 61 – 64" captionStartId="21.[151,264,1846,1869]" captionTargetBox="[160,1393,591,1818]" captionTargetId="figure@21.[151,1436,591,1824]" captionTargetPageId="21" captionText="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262422" httpUri="https://zenodo.org/record/262422/files/figure.png" pageId="17" pageNumber="18">Fig. 62</figureCitation>
;
<emphasis id="B97CEAAD2928FFE0FEE6C45C9CDAFE59" box="[370,398,369,392]" italics="true" pageId="17" pageNumber="18">pg</emphasis>
) visible in dorsal view, subtriangular.
@ -258,11 +259,11 @@ Abdomen (
<subSubSection id="C31265342928FFE0FF53C4B99E9FFE01" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BB736BF2928FFE0FF53C4B99E9FFE01" blockId="17.[151,1437,151,1544]" pageId="17" pageNumber="18">
<emphasis id="B97CEAAD2928FFE0FF53C4B99F79FE7C" bold="true" box="[199,557,404,429]" pageId="17" pageNumber="18">Distribution and composition.</emphasis>
<taxonomicName id="4C084D3C2928FFE0FDA1C4B99FC4FE7C" box="[565,656,404,429]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C2928FFE0FDA1C4B99FC4FE7C" box="[565,656,404,429]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FDA1C4B99FC4FE7C" box="[565,656,404,429]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
is represented by one nominal and one undescribed species from the Eocene of Europe (
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<figureCitation id="13332A3A2928FFE0FEFBC49A9C96FE01" box="[367,450,439,464]" captionStart="FIGURES 35 – 38" captionStartId="10.[151,264,1940,1963]" captionTargetBox="[181,1396,193,1909]" captionTargetId="figure@10.[181,1396,193,1910]" captionTargetPageId="10" captionText="FIGURES 35 – 38. Location of the Rovno Region (35 − 36) and the Klesov, Dubrovitsa and Vyrka amber mines (37) in the present-day Europe; location of the Rovno Region during Eocene (38)." figureDoi="http://doi.org/10.5281/zenodo.262416" httpUri="https://zenodo.org/record/262416/files/figure.png" pageId="17" pageNumber="18">Fig. 38</figureCitation>
), area currently within north-western
<collectingCountry id="F31F762F2928FFE0FCFDC49A9E93FE01" box="[873,967,439,464]" name="Ukraine" pageId="17" pageNumber="18">Ukraine</collectingCountry>
.
@ -333,7 +334,7 @@ and
<bibRefCitation id="EF994B4E2928FFE0FE60C6A19E4BFC74" author="Jaloszynski" box="[500,799,908,933]" pageId="17" pageNumber="18" refString="Jaloszynski, P. &amp; Peris, D. (2016) Cretaceous amber inclusions of Spain and Myanmar demonstrate early diversification and wide dispersal of Cephenniitae (Coleoptera: Staphylinidae: Scydmaeninae). Cretaceous Research, 57, 190 - 198. http: // dx. doi. org / 10.1016 / j. cretres. 2015.09.002" type="journal article" year="2016">Jałoszyński &amp; Peris, 2016</bibRefCitation>
</taxonomicName>
have the maxillary palpomere IV subconical. The new genus is very similar to all these three taxa in the general body form, the truncate elytral apices and the procoxae not separated by prosternal process.
<taxonomicName id="4C084D3C2928FFE0FDB8C6F99FD3FC3C" box="[556,647,980,1005]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C2928FFE0FDB8C6F99FD3FC3C" box="[556,647,980,1005]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FDB8C6F99FD3FC3C" box="[556,647,980,1005]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
has the pair of distinct pits on the posterior portion of the frons, with a groove extending from each pit posterolaterally; previously this character was thought to be a hallmark of
@ -355,7 +356,7 @@ have traces of similar pits or grooves, which are rather faint compared to those
,
<bibRefCitation id="EF994B4E2928FFE0FB4DC11299BDFB89" author="Jaloszynski" box="[1241,1257,1087,1112]" pageId="17" pageNumber="18" refString="Jaloszynski, P. (2008 b) Revision of Eutheia Stephens of Taiwan (Coleoptera, Scydmaenidae). Genus, 19 (2), 177 - 190." type="journal article" year="2008" yearSuffix="b">b</bibRefCitation>
)). Interestingly, although this character makes
<taxonomicName id="4C084D3C2928FFE0FE65C1499F18FBAC" box="[497,588,1124,1149]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C2928FFE0FE65C1499F18FBAC" box="[497,588,1124,1149]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FE65C1499F18FBAC" box="[497,588,1124,1149]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
very similar to
@ -369,7 +370,7 @@ in dorsal view, the broad, flat and subtrapezoidal mesoventral intercoxal proces
. The latter genus has the mesoventral process narrowly carinate and elevated (illustrated in
<bibRefCitation id="EF994B4E2928FFE0FB3DC18198CCFB14" author="Jaloszynski" box="[1193,1432,1196,1221]" pageId="17" pageNumber="18" refString="Jaloszynski, P. (2014 b) Phylogeny of a new supertribe Cephenniitae with generic review of Eutheiini and description of a new tribe Marcepaniini (Coleoptera: Staphylinidae: Scydmaeninae). Systematic Entomology, 39, 159 - 189. http: // dx. doi. org / 10.1111 / syen. 12044" type="journal article" year="2014" yearSuffix="b">Jałoszyński (2014b))</bibRefCitation>
. The intermesocoxal region in
<taxonomicName id="4C084D3C2928FFE0FE7DC1E29F10FB39" box="[489,580,1231,1256]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C2928FFE0FE7DC1E29F10FB39" box="[489,580,1231,1256]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FE7DC1E29F10FB39" box="[489,580,1231,1256]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
is nearly identical to that of
@ -381,7 +382,7 @@ and
<emphasis id="B97CEAAD2928FFE0FB8EC1E299F6FB39" box="[1050,1186,1231,1256]" italics="true" pageId="17" pageNumber="18">Archeutheia</emphasis>
</taxonomicName>
.
<taxonomicName id="4C084D3C2928FFE0FB24C1E2985FFB39" box="[1200,1291,1231,1256]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C2928FFE0FB24C1E2985FFB39" box="[1200,1291,1231,1256]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FB24C1E2985FFB39" box="[1200,1291,1231,1256]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
differs from these two genera in absent or rudimentary basal elytral foveae (the elytral base is well exposed in two studied inclusions, but the foveae are not discernible), and from
@ -393,7 +394,7 @@ additionally in complete hypomeral ridges (absent in the Cretaceous taxon). More
<emphasis id="B97CEAAD2928FFE0FCCDC0119EE4FA84" box="[857,944,1340,1365]" italics="true" pageId="17" pageNumber="18">Eutheia</emphasis>
</taxonomicName>
is short and transverse, whereas in
<taxonomicName id="4C084D3C2928FFE0FAAAC01198CCFA84" box="[1342,1432,1340,1365]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C2928FFE0FAAAC01198CCFA84" box="[1342,1432,1340,1365]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FAAAC01198CCFA84" box="[1342,1432,1340,1365]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
it is similar to that in
@ -436,7 +437,7 @@ with four-segmented maxillary palps and subconical maxillary palpomere IV.
<emphasis id="B97CEAAD2928FFE0FC77C3A7990AF971" box="[995,1118,1674,1696]" italics="true" pageId="17" pageNumber="18">Archeutheia</emphasis>
</taxonomicName>
†
<taxonomicName id="4C084D3C2928FFE0FB5EC3A7984AF971" box="[1226,1310,1674,1696]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C2928FFE0FB5EC3A7984AF971" box="[1226,1310,1674,1696]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD2928FFE0FB5EC3A7984AF971" box="[1226,1310,1674,1696]" italics="true" pageId="17" pageNumber="18">Vertheia</emphasis>
</taxonomicName>
</paragraph>

103
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</mods:mods>
<treatment id="03A187A9292BFFE2FF03C5B59986FDED" ID-DOI="http://doi.org/10.5281/zenodo.5681737" ID-GBIF-Taxon="124407823" ID-Zenodo-Dep="5681737" LSID="urn:lsid:plazi:treatment:03A187A9292BFFE2FF03C5B59986FDED" httpUri="http://treatment.plazi.org/id/03A187A9292BFFE2FF03C5B59986FDED" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<treatment id="03A187A9292BFFE2FF03C5B59986FDED" ID-DOI="http://doi.org/10.5281/zenodo.5681737" ID-GBIF-Taxon="124407823" ID-Zenodo-Dep="5681737" LSID="urn:lsid:plazi:treatment:03A187A9292BFFE2FF03C5B59986FDED" httpUri="http://treatment.plazi.org/id/03A187A9292BFFE2FF03C5B59986FDED" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" scope_family="Staphylinidae" scope_order="Coleoptera" scope_tribe="Eutheiini">
<subSubSection id="C3126534292BFFE3FF03C5B59C17FF02" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BB736BF292BFFE3FF03C5B59CA1FF63" blockId="18.[151,501,151,211]" box="[151,501,151,178]" pageId="18" pageNumber="19">
<heading id="D0FF81D3292BFFE3FF03C5B59CA1FF63" bold="true" box="[151,501,151,178]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B97CEAAD292BFFE3FF03C5B59CA1FF63" bold="true" box="[151,501,151,178]" pageId="18" pageNumber="19">
†
<taxonomicName id="4C084D3C292BFFE3FF31C5BA9CFFFF60" box="[165,427,151,177]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quadrisetosa" status="sp. nov.">
<taxonomicName id="4C084D3C292BFFE3FF31C5BA9CFFFF60" ID-CoL="5B8TR" box="[165,427,151,177]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quadrisetosa" status="sp. nov.">
<emphasis id="B97CEAAD292BFFE3FF31C5BA9CFFFF60" bold="true" box="[165,427,151,177]" italics="true" pageId="18" pageNumber="19">Vertheia quadrisetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D6292BFFE3FE26C5B59CA1FF63" box="[434,501,152,178]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
@ -56,9 +57,9 @@
</paragraph>
<paragraph id="8BB736BF292BFFE3FF03C5979C17FF02" blockId="18.[151,501,151,211]" box="[151,323,186,211]" pageId="18" pageNumber="19">
(
<figureCitation id="13332A3A292BFFE3FF0BC5979DB3FF02" box="[159,231,186,211]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="18" pageNumber="19">Figs 2</figureCitation>
<figureCitation id="13332A3A292BFFE3FF0BC5979DB3FF02" box="[159,231,186,211]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." figureDoi="http://doi.org/10.5281/zenodo.262414" httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="18" pageNumber="19">Figs 2</figureCitation>
,
<figureCitation id="13332A3A292BFFE3FF67C5979C6FFF02" box="[243,315,186,211]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">49–55</figureCitation>
<figureCitation id="13332A3A292BFFE3FF67C5979C6FFF02" box="[243,315,186,211]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">49–55</figureCitation>
)
</paragraph>
</subSubSection>
@ -80,7 +81,7 @@ amber:
sex unknown; inclusion in elongate, dropshaped piece of amber
<quantity id="4CF09B5A292BFFE3FE0DC40A9CBCFEEF" box="[409,488,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="18" pageNumber="19" unit="mm" value="14.0">14 mm</quantity>
long (
<figureCitation id="13332A3A292BFFE3FDA4C40B9F20FEEE" box="[560,628,294,319]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="18" pageNumber="19">Fig. 2</figureCitation>
<figureCitation id="13332A3A292BFFE3FDA4C40B9F20FEEE" box="[560,628,294,319]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." figureDoi="http://doi.org/10.5281/zenodo.262414" httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="18" pageNumber="19">Fig. 2</figureCitation>
), collection number K-243 (
<collectionCode id="ED19AE7A292BFFE3FC26C40A9EA2FEEE" box="[946,1014,295,319]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="18" pageNumber="19">SIZK</collectionCode>
).
@ -93,16 +94,16 @@ long (
As for genus; additionally pronotum with two pairs of long lateral setae and EI nearly 1.3.
<emphasis id="B97CEAAD292BFFE3FF53C4439C0DFE56" bold="true" box="[199,345,366,391]" pageId="18" pageNumber="19">Description.</emphasis>
Body (
<figureCitation id="13332A3A292BFFE3FE3EC4439F7FFE56" box="[426,555,366,391]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">Figs 49–55</figureCitation>
<figureCitation id="13332A3A292BFFE3FE3EC4439F7FFE56" box="[426,555,366,391]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">Figs 49–55</figureCitation>
) elongate and flatten, brown; BL
<quantity id="4CF09B5A292BFFE3FC32C4429959FE57" box="[934,1037,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="18" pageNumber="19" unit="mm" value="0.98">0.98 mm</quantity>
.
</paragraph>
<caption id="DF776637292BFFE3FF03C3959CE4F8B9" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19" targetBox="[151,1436,443,1689]" targetPageId="18">
<caption id="DF776637292BFFE3FF03C3959CE4F8B9" ID-DOI="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19" targetBox="[151,1436,443,1689]" targetPageId="18">
<paragraph id="8BB736BF292BFFE3FF03C3959CE4F8B9" blockId="18.[151,1436,1720,1896]" pageId="18" pageNumber="19">
<emphasis id="B97CEAAD292BFFE3FF03C3959C01F91F" bold="true" box="[151,341,1720,1743]" pageId="18" pageNumber="19">FIGURES 49–55.</emphasis>
Holotype of
<taxonomicName id="4C084D3C292BFFE3FE4BC3949FEBF91E" box="[479,703,1721,1743]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quadrisetosa" status="sp. nov.">
<taxonomicName id="4C084D3C292BFFE3FE4BC3949FEBF91E" box="[479,703,1721,1743]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quadrisetosa" status="sp. nov.">
<emphasis id="B97CEAAD292BFFE3FE4BC3949FEBF91E" box="[479,703,1721,1743]" italics="true" pageId="18" pageNumber="19">Vertheia quadrisetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D6292BFFE3FD52C3949E54F91F" box="[710,768,1721,1742]" pageId="18" pageNumber="19" rank="species">
@ -113,54 +114,54 @@ Holotype of
</caption>
<paragraph id="8BB736BF292BFFE2FF53C2A29F63FF26" blockId="18.[151,1437,1935,2033]" lastBlockId="19.[151,1436,151,573]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Head (
<figureCitation id="13332A3A292BFFE3FE82C2A29CCEF879" box="[278,410,1935,1960]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">Figs 49–54</figureCitation>
<figureCitation id="13332A3A292BFFE3FE82C2A29CCEF879" box="[278,410,1935,1960]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">Figs 49–54</figureCitation>
) broadest at large eyes, HL
<quantity id="4CF09B5A292BFFE3FD66C2A29E09F876" box="[754,861,1935,1960]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="mm" value="0.15">0.15 mm</quantity>
, HW
<quantity id="4CF09B5A292BFFE3FC33C2A29947F876" box="[935,1043,1935,1960]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="mm" value="0.25">0.25 mm</quantity>
; vertex and frons together nearly subquadrate; supraantennal tubercles indistinct. Punctures on head fine and unremarkable, setae only partly visible (
<figureCitation id="13332A3A292BFFE3FF0BC2F59DA2F820" box="[159,246,2008,2033]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">Fig. 54</figureCitation>
<figureCitation id="13332A3A292BFFE3FF0BC2F59DA2F820" box="[159,246,2008,2033]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">Fig. 54</figureCitation>
), sparse, short and suberect, except for pair of remarkably long setae, each inserted near posteromesal margin of compound eye (
<figureCitation id="13332A3A292AFFE2FE56C5BA9F4CFF61" box="[450,536,151,176]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 54</figureCitation>
<figureCitation id="13332A3A292AFFE2FE56C5BA9F4CFF61" box="[450,536,151,176]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 54</figureCitation>
). Antennae (
<figureCitation id="13332A3A292AFFE2FD33C5BA9E7CFF61" box="[679,808,151,176]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Figs 49–54</figureCitation>
<figureCitation id="13332A3A292AFFE2FD33C5BA9E7CFF61" box="[679,808,151,176]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Figs 49–54</figureCitation>
) about 0.4 × as long as body, AnL
<quantity id="4CF09B5A292AFFE2FB2EC5B59875FF7E" box="[1210,1313,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="mm" value="0.4">0.40 mm</quantity>
, scape and pedicel strongly elongate, antennomeres III–X slightly to strongly transverse, XI about as long as IX–X together, about as broad as X, with blunt apex.
</paragraph>
<paragraph id="8BB736BF292AFFE2FF53C4289ECAFE7C" blockId="19.[151,1436,151,573]" pageId="19" pageNumber="20">
Pronotum (
<figureCitation id="13332A3A292AFFE2FED3C4299F1AFECC" box="[327,590,260,285]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Figs 49–50, 52, 53–54</figureCitation>
<figureCitation id="13332A3A292AFFE2FED3C4299F1AFECC" box="[327,590,260,285]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Figs 49–50, 52, 53–54</figureCitation>
) subrectangular; PL
<quantity id="4CF09B5A292AFFE2FCD5C4299EFEFECD" box="[833,938,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="20" unit="mm" value="0.28">0.28 mm</quantity>
, PW
<quantity id="4CF09B5A292AFFE2FC79C4299903FECD" box="[1005,1111,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="20" unit="mm" value="0.3">0.30 mm</quantity>
; pronotal disc broadest near anterior third, sides near posterior third indistinctly concave; transverse antebasal groove (
<figureCitation id="13332A3A292AFFE2FB05C40A99B1FE91" box="[1169,1253,295,320]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 53</figureCitation>
<figureCitation id="13332A3A292AFFE2FB05C40A99B1FE91" box="[1169,1253,295,320]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 53</figureCitation>
;
<emphasis id="B97CEAAD292AFFE2FB65C40A984FFE91" box="[1265,1307,295,320]" italics="true" pageId="19" pageNumber="20">abg</emphasis>
) seems not connected to lateral antebasal impressions (
<figureCitation id="13332A3A292AFFE2FD15C4619F81FEB4" box="[641,725,332,357]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 53</figureCitation>
<figureCitation id="13332A3A292AFFE2FD15C4619F81FEB4" box="[641,725,332,357]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 53</figureCitation>
;
<emphasis id="B97CEAAD292AFFE2FD76C4619E52FEB4" box="[738,774,332,357]" italics="true" pageId="19" pageNumber="20">abi</emphasis>
). Each lateral pronotal margin with two long setae located in median region; pronotum with shallow but distinct punctures separated by spaces subequal to two diameters of punctures; setae sparse, moderately long, nearly recumbent (
<figureCitation id="13332A3A292AFFE2FCA9C4B99EC5FE7C" box="[829,913,404,429]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 54</figureCitation>
<figureCitation id="13332A3A292AFFE2FCA9C4B99EC5FE7C" box="[829,913,404,429]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 54</figureCitation>
).
</paragraph>
<paragraph id="8BB736BF292AFFE2FF53C49A9942FE24" blockId="19.[151,1436,151,573]" pageId="19" pageNumber="20">
Elytra (
<figureCitation id="13332A3A292AFFE2FE88C49A9CA1FE01" box="[284,501,439,464]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Figs 49–50, 53–54</figureCitation>
<figureCitation id="13332A3A292AFFE2FE88C49A9CA1FE01" box="[284,501,439,464]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Figs 49–50, 53–54</figureCitation>
) elongate oval, flattened, EL
<quantity id="4CF09B5A292AFFE2FCD8C49A9EE0FE1E" box="[844,948,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="19" pageNumber="20" unit="mm" value="0.55">0.55 mm</quantity>
, EW
<quantity id="4CF09B5A292AFFE2FC63C495990BFE1E" box="[1015,1119,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="19" pageNumber="20" unit="mm" value="0.43">0.43 mm</quantity>
, EI 1.29; punctures slightly denser than those on pronotal disc; setae similar to those on pronotum (
<figureCitation id="13332A3A292AFFE2FC21C4F1995EFE24" box="[949,1034,476,501]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 54</figureCitation>
<figureCitation id="13332A3A292AFFE2FC21C4F1995EFE24" box="[949,1034,476,501]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 54</figureCitation>
).
</paragraph>
<paragraph id="8BB736BF292AFFE2FF53C72D9E4DFDC9" blockId="19.[151,1436,151,573]" box="[199,793,511,536]" pageId="19" pageNumber="20">
Legs (
<figureCitation id="13332A3A292AFFE2FE99C4D29C0AFDC9" box="[269,350,511,536]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 55</figureCitation>
<figureCitation id="13332A3A292AFFE2FE99C4D29C0AFDC9" box="[269,350,511,536]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 55</figureCitation>
) moderately long, slender, unmodified.
</paragraph>
</subSubSection>
@ -168,7 +169,7 @@ Legs (
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<emphasis id="B97CEAAD292AFFE2FF53C7099C1AFDEC" bold="true" box="[199,334,548,573]" pageId="19" pageNumber="20">Etymology.</emphasis>
The name
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<taxonomicName id="4C084D3C292AFFE2FE5FC7099F08FDEC" box="[459,604,548,573]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="quadrisetosa">
<emphasis id="B97CEAAD292AFFE2FE5FC7099F08FDEC" box="[459,604,548,573]" italics="true" pageId="19" pageNumber="20">quadrisetosa</emphasis>
</taxonomicName>
refers to the long paired setae on sides of the pronotum.

79
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View file

@ -1,53 +1,54 @@
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<subSubSection id="C3126534292DFFE5FF03C5B59C17FF02" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BB736BF292DFFE5FF03C5B59C1FFF63" blockId="20.[151,331,151,211]" box="[151,331,151,178]" pageId="20" pageNumber="21">
<heading id="D0FF81D3292DFFE5FF03C5B59C1FFF63" bold="true" box="[151,331,151,178]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B97CEAAD292DFFE5FF03C5B59C1FFF63" bold="true" box="[151,331,151,178]" pageId="20" pageNumber="21">
†
<taxonomicName id="4C084D3C292DFFE5FF31C5BA9C53FF60" box="[165,263,151,177]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C292DFFE5FF31C5BA9C53FF60" ID-CoL="8XQNM" box="[165,263,151,177]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD292DFFE5FF31C5BA9C53FF60" bold="true" box="[165,263,151,177]" italics="true" pageId="20" pageNumber="21">Vertheia</emphasis>
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sp. 1
@ -56,9 +57,9 @@ sp. 1
</paragraph>
<paragraph id="8BB736BF292DFFE5FF03C5979C17FF02" blockId="20.[151,331,151,211]" box="[151,323,186,211]" pageId="20" pageNumber="21">
(
<figureCitation id="13332A3A292DFFE5FF0BC5979DB3FF02" box="[159,231,186,211]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="20" pageNumber="21">Figs 3</figureCitation>
<figureCitation id="13332A3A292DFFE5FF0BC5979DB3FF02" box="[159,231,186,211]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." figureDoi="http://doi.org/10.5281/zenodo.262414" httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="20" pageNumber="21">Figs 3</figureCitation>
,
<figureCitation id="13332A3A292DFFE5FF67C5979C6FFF02" box="[243,315,186,211]" captionStart-0="FIGURES 56 – 60" captionStart-1="FIGURES 61 – 64" captionStartId-0="19.[151,264,1962,1985]" captionStartId-1="21.[151,264,1846,1869]" captionTargetBox-0="[151,1436,628,1941]" captionTargetBox-1="[160,1393,591,1818]" captionTargetId-0="figure@19.[151,1436,628,1941]" captionTargetId-1="figure@21.[151,1436,591,1824]" captionTargetPageId-0="19" captionTargetPageId-1="21" captionText-0="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." captionText-1="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." httpUri-0="https://zenodo.org/record/262421/files/figure.png" httpUri-1="https://zenodo.org/record/262422/files/figure.png" pageId="20" pageNumber="21">56–64</figureCitation>
<figureCitation id="13332A3A292DFFE5FF67C5979C6FFF02" box="[243,315,186,211]" captionStart-0="FIGURES 56 – 60" captionStart-1="FIGURES 61 – 64" captionStartId-0="19.[151,264,1962,1985]" captionStartId-1="21.[151,264,1846,1869]" captionTargetBox-0="[151,1436,628,1941]" captionTargetBox-1="[160,1393,591,1818]" captionTargetId-0="figure@19.[151,1436,628,1941]" captionTargetId-1="figure@21.[151,1436,591,1824]" captionTargetPageId-0="19" captionTargetPageId-1="21" captionText-0="FIGURES 56 – 60. Vertheia sp. 1 (UA- 442). Approximately anterodorsal view (56, 57), lateral view (58), ventral view of head and posteroventral view of thorax and abdomen (59, 60)." captionText-1="FIGURES 61 – 64. Vertheia sp. 1 (UA- 442). Anterodorsal (61), dorsal (62), ventral (63) and lateral (64) view. Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; bsti, basistipes; fr, frons; frp, frontal pits; gp, gular plate; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 2 – 4, maxillary palpomere II – IV; nr, ' neck' region; occ, occipital constriction; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum; st 3 – 8, abdominal sternite III – VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi-0="http://doi.org/10.5281/zenodo.262421" figureDoi-1="http://doi.org/10.5281/zenodo.262422" httpUri-0="https://zenodo.org/record/262421/files/figure.png" httpUri-1="https://zenodo.org/record/262422/files/figure.png" pageId="20" pageNumber="21">56–64</figureCitation>
)
</paragraph>
</subSubSection>
@ -78,7 +79,7 @@ amber:
sex unknown; inclusion in an elongate prism of amber
<quantity id="4CF09B5A292DFFE5FF03C40A9DB2FEEF" box="[151,230,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="mm" value="20.0">20 mm</quantity>
long (
<figureCitation id="13332A3A292DFFE5FEBAC40B9C25FEEE" box="[302,369,294,319]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="20" pageNumber="21">Fig. 3</figureCitation>
<figureCitation id="13332A3A292DFFE5FEBAC40B9C25FEEE" box="[302,369,294,319]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." figureDoi="http://doi.org/10.5281/zenodo.262414" httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="20" pageNumber="21">Fig. 3</figureCitation>
), collection number UA-442 (
<collectionCode id="ED19AE7A292DFFE5FD50C40A9E5CFEEE" box="[708,776,295,319]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="20" pageNumber="21">SIZK</collectionCode>
).
@ -89,11 +90,11 @@ long (
<paragraph id="8BB736BF292DFFE5FF53C4679EFFFD8E" blockId="20.[151,1436,258,679]" pageId="20" pageNumber="21">
<emphasis id="B97CEAAD292DFFE5FF53C4679C6FFEB2" bold="true" box="[199,315,330,355]" pageId="20" pageNumber="21">Remarks.</emphasis>
This specimen is very similar to
<taxonomicName id="4C084D3C292DFFE5FD77C4619EC2FEB2" box="[739,918,330,355]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="quadrisetosa">
<taxonomicName id="4C084D3C292DFFE5FD77C4619EC2FEB2" box="[739,918,330,355]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="quadrisetosa">
<emphasis id="B97CEAAD292DFFE5FD77C4619EC2FEB2" box="[739,918,330,355]" italics="true" pageId="20" pageNumber="21">V. quadrisetosa</emphasis>
</taxonomicName>
, but it lacks any remarkable characters to unambiguously define a new species. The differences include significantly lower EI (i.e., stouter elytra), slightly smaller BL, shorter antennae, lack of the long setae near posteromesal margins of eyes and lateral pronotal setae, and the presence of complete transverse antebasal groove, i.e., connecting lateral antebasal impressions. However, the setae may be broken off, the pronotal groove may show some variation, and proportions of the body parts and the body length may be different in males and females of the same species. For this reason, the specimen of
<taxonomicName id="4C084D3C292DFFE5FF03C70F9DA6FDEA" box="[151,242,546,571]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C292DFFE5FF03C70F9DA6FDEA" box="[151,242,546,571]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD292DFFE5FF03C70F9DA6FDEA" box="[151,242,546,571]" italics="true" pageId="20" pageNumber="21">Vertheia</emphasis>
</taxonomicName>
that shows more unique, remarkable characters was selected as a
@ -102,7 +103,7 @@ of a new species, whereas the other one, slightly different and less remarkable,
</paragraph>
<paragraph id="8BB736BF292DFFE5FF53C7469E59FD76" blockId="20.[151,1436,258,679]" pageId="20" pageNumber="21">
Measurements of
<taxonomicName id="4C084D3C292DFFE5FE07C7479CBAFD52" box="[403,494,618,643]" class="Insecta" family="Chironomidae" genus="Vertheia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C084D3C292DFFE5FE07C7479CBAFD52" box="[403,494,618,643]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B97CEAAD292DFFE5FE07C7479CBAFD52" box="[403,494,618,643]" italics="true" pageId="20" pageNumber="21">Vertheia</emphasis>
</taxonomicName>
sp. (UA-442): BL