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<taxonomicName id="A9DB1E85FF86FF975AB06D3CF2F1FA88" authorityName="Manchester" authorityYear="2014" box="[1000,1221,1295,1318]" class="Magnoliopsida" family="Cucurbitaceae" genus="Cucurbitaciphyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cucurbitales" pageId="16" pageNumber="169" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="5CAFB914FF86FF975AB06D3CF2F1FA88" bold="true" box="[1000,1221,1295,1318]" italics="true" pageId="16" pageNumber="169">Cucurbitaciphyllum</emphasis>
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<paragraph id="6E646506FF86FF975A296D0DF34DF9F8" blockId="16.[843,1482,1342,1622]" pageId="16" pageNumber="169">D i a g n o s i s. Leaves palmately trilobed, petiole narrow, not swollen at junction with lamina. Lamina base cordate, lobe apices acute; lobal sinuses rounded, margins of the three main lobes with smaller lobes and teeth. Venation actinodromous with a primary vein directed into each major lobe. Secondary veins pinnate, camptodromous to semicraspedodromous, sometimes interconnecting to form an intramarginal vein in lower portion of the lobes. Lower surface of leaf covered by closely spaced trichomes.</paragraph>
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T y p e s p e c i e s.
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<emphasis id="5CAFB914FF86FF975D126E5DF347F92B" box="[1098,1395,1646,1669]" italics="true" pageId="16" pageNumber="169">Cucurbitaciphyllum lobatum</emphasis>
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(
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<paragraph id="6E646506FF86FF975A296E8EF5B7F89A" blockId="16.[843,1482,1725,1844]" pageId="16" pageNumber="169">
E t y m o l o g y. The generic name combines the roots, Cucurbitac-, referring to the family
<taxonomicName id="A9DB1E85FF86FF975D846EEEF34FF95A" box="[1244,1403,1757,1780]" class="Magnoliopsida" family="Cucurbitaceae" kingdom="Plantae" order="Cucurbitales" pageId="16" pageNumber="169" phylum="Tracheophyta" rank="family">Cucurbitaceae</taxonomicName>
, which has similar leaves among extant genera, and -phyllon (Gr. = leaf).
</paragraph>
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</treatment>
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<taxonomicName id="A9DB1E85FF9CFF8D5ACD6EE9F337F95F" authority="(NEWBERRY)" authorityName="Manchester" authorityYear="2014" baseAuthorityName="Manchester" baseAuthorityYear="2014" box="[917,1283,1754,1777]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="10" pageNumber="163" phylum="Tracheophyta" rank="species" species="raynoldsii" status="comb. nov.">
<emphasis id="5CAFB914FF9CFF8D5ACD6EE9F243F95F" bold="true" box="[917,1143,1754,1777]" italics="true" pageId="10" pageNumber="163">Platanites raynoldsii</emphasis>
(
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<figureCitation id="F6E07983FF9CFF8D5D6B6F39F2D5F88E" box="[1075,1249,1802,1824]" captionStart="Text-fig" captionStartId="12.[129,210,1934,1955]" captionTargetBox="[129,1482,152,1901]" captionTargetId="figure-11@12.[156,1454,178,1867]" captionTargetPageId="12" captionText="Text-fig. 6. Platanaceae 13. Platanites raynoldsii (NEWBERRY) comb. nov. 1. Holotype of Platanus raynoldsii NEWBERRY (1868, 1898), Banks of the Yellowstone River, Montana, USNM 7000. 2. Holotype of Platanites canadensis MCIVER et BASINGER (1993), here treated as synonym of P. raynoldsii, showing clearly the position of the lateral leaflets, US 3-66, Ravenscrag Formation, Saskatchewan, Canada. 3. Specimen from Yellowstone Bridge, Miles City, Montana. UF19021-39390. 4. Detail from fig. 2. Scale bars 5 cm." figureDoi="http://doi.org/10.5281/zenodo.13183586" httpUri="https://zenodo.org/record/13183586/files/figure.png" pageId="10" pageNumber="163">Text-figs 6.16.4</figureCitation>
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<paragraph id="6E646506FF9CFF8D5A136F0AF20BF81F" blockId="10.[843,1482,1849,2099]" pageId="10" pageNumber="163">
1868
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<emphasis id="5CAFB914FF9CFF8D5AF36F0AF24DF8FE" box="[939,1145,1849,1872]" italics="true" pageId="10" pageNumber="163">Platanus raynoldsi</emphasis>
<smallCapsWord id="6882F3DAFF9CFF8D5DD96F0AF2C6F8E1" baselines="1866,1866" box="[1153,1266,1849,1872]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Newberry" pageId="10" pageNumber="163">NEWBERRY</smallCapsWord>
, Lyceum Nat. Hist.
</taxonomicName>
<collectingRegion id="AC1FABE4FF9CFF8D5AF36F6AF221F8DE" box="[939,1045,1881,1904]" country="United States of America" name="New York" pageId="10" pageNumber="163">New York</collectingRegion>
Ann., 9, p. 69 (basionym); 1898
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<emphasis id="5CAFB914FF9CFF8D5C346F6AF23AF83F" italics="true" pageId="10" pageNumber="163">Platanus raynoldsi</emphasis>
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, U. S. Geol. Surv. Monogr.
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, 35, p. 109, pl. 35.
</paragraph>
<paragraph id="6E646506FF9CFF8D5A136F88F2A8F85C" blockId="10.[843,1482,1849,2099]" pageId="10" pageNumber="163">
1993
<taxonomicName id="A9DB1E85FF9CFF8D5AF36F88F34AF87F" authority="MCIVER et BASINGER" authorityName="MCIVER et BASINGER" box="[939,1406,1979,2002]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="10" pageNumber="163" phylum="Tracheophyta" rank="species" species="canadensis">
<emphasis id="5CAFB914FF9CFF8D5AF36F88F2A2F87C" box="[939,1174,1979,2002]" italics="true" pageId="10" pageNumber="163">Platanites canadensis</emphasis>
<smallCapsWord id="6882F3DAFF9CFF8D5DC56F88F2C7F87F" baselines="1996,1996" box="[1181,1267,1979,2002]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Mc" pageId="10" pageNumber="163">MCIVER</smallCapsWord>
et
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, p. 38, pl. 23, pl. 24, figs 14.
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2008
<taxonomicName id="A9DB1E85FF9CFF8D5AF36FCFF3F7F7BC" authority="(NEWBERRY) MOISEEVA" authorityName="MOISEEVA" baseAuthorityName="NEWBERRY" box="[939,1475,2044,2067]" class="Magnoliopsida" family="Vitaceae" genus="Ettingshausenia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Vitales" pageId="10" pageNumber="163" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9CFF8D5AF36FCFF2F3F7BD" box="[939,1223,2044,2067]" italics="true" pageId="10" pageNumber="163">Ettingshausenia raynoldsii</emphasis>
(
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)
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, p. 324 [excluding all of the Asian figured specimens].
</paragraph>
<caption id="3AA4358EFF9DFF8C59326FBDF331F79D" ID-DOI="http://doi.org/10.5281/zenodo.13183584" ID-Zenodo-Dep="13183584" httpUri="https://zenodo.org/record/13183584/files/figure.png" pageId="11" pageNumber="164" startId="11.[106,187,1934,1955]" targetBox="[106,1459,152,1900]" targetPageId="11" targetType="figure">
<paragraph id="6E646506FF9DFF8C59326FBDF331F79D" blockId="11.[106,1459,1934,2099]" pageId="11" pageNumber="164">
<emphasis id="5CAFB914FF9DFF8C59326FBDF5BDF7BB" bold="true" pageId="11" pageNumber="164">
Text-fig. 5.
<taxonomicName id="A9DB1E85FF9DFF8C59B86FBDF76FF80D" box="[224,347,1934,1955]" class="Magnoliopsida" family="Platanaceae" kingdom="Plantae" order="Proteales" pageId="11" pageNumber="164" phylum="Tracheophyta" rank="family">Platanaceae</taxonomicName>
13.
<taxonomicName id="A9DB1E85FF9DFF8C58CC6FBDF4EFF80D" authority="(NEWBERRY)" authorityName="Manchester" authorityYear="2014" baseAuthorityName="Manchester" baseAuthorityYear="2014" box="[404,731,1934,1955]" class="Magnoliopsida" family="Platanaceae" genus="Macginitiea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="11" pageNumber="164" phylum="Tracheophyta" rank="species" species="nobilis" status="comb. nov.">
<emphasis id="5CAFB914FF9DFF8C58CC6FBDF462F80D" bold="true" box="[404,598,1934,1955]" italics="true" pageId="11" pageNumber="164">Macginitiea nobilis</emphasis>
(
<smallCapsWord id="6882F3DAFF9DFF8C5B3C6FBDF4E0F80C" baselines="1950,1950" box="[612,724,1934,1955]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Newberry" pageId="11" pageNumber="164">NEWBERRY</smallCapsWord>
)
</taxonomicName>
<taxonomicNameLabel id="479C046FFF9DFF8C5BBB6FBDF567F80D" box="[739,851,1934,1955]" pageId="11" pageNumber="164" rank="species">comb. nov.</taxonomicNameLabel>
1. This specimen is labeled as corresponding to
<bibRefCitation id="0A4A18F7FF9DFF8C5C146FBDF6A8F811" author="Newberry, J. S." pageId="11" pageNumber="164" pagination="1 - 295" refId="ref32108" refString="Newberry, J. S. (1898): The later extinct floras of North America. - U. S. Geol. Surv. Monogr., 35: 1 - 295, 68 pls." type="journal article" year="1898">Newberry 1898</bibRefCitation>
, pl. 50, fig. 1 although it does not match the published drawing exactly. From near Fort Clark, North Dakota, USNM 6964. 2. Lectotype from
<bibRefCitation id="0A4A18F7FF9DFF8C58716FF4F7FBF872" author="Newberry, J. S." box="[297,463,1991,2012]" pageId="11" pageNumber="164" pagination="1 - 295" refId="ref32108" refString="Newberry, J. S. (1898): The later extinct floras of North America. - U. S. Geol. Surv. Monogr., 35: 1 - 295, 68 pls." type="journal article" year="1898">Newberry (1898</bibRefCitation>
, pl. 34), from near Fort Clark, North Dakota; composite picture assembled from images of both counterparts. USNM 1070. 3. Trilobed leaf from Seven Mile Creek, Montana (orig. figured as
<taxonomicName id="A9DB1E85FF9DFF8C5D616FD7F364F856" authority="NEWBERRY" authorityName="NEWBERRY. However" authorityYear="1868" box="[1081,1360,2020,2041]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="11" pageNumber="164" phylum="Tracheophyta" rank="species" species="nobilis">
<emphasis id="5CAFB914FF9DFF8C5D616FD7F2E8F857" bold="true" box="[1081,1244,2020,2041]" italics="true" pageId="11" pageNumber="164">Platanus nobilis</emphasis>
<smallCapsWord id="6882F3DAFF9DFF8C5DBB6FD7F364F856" baselines="2036,2036" box="[1251,1360,2020,2041]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Newberry" pageId="11" pageNumber="164">NEWBERRY</smallCapsWord>
</taxonomicName>
by Ward 1886, pl. 41, fig. 1). USNM 4093. 4.
<taxonomicName id="A9DB1E85FF9DFF8C58B36032F5BDF7BB" authority="PIGG et STOCKEY" authorityName="PIGG et STOCKEY" box="[491,905,2049,2070]" class="Magnoliopsida" family="Platanaceae" genus="Platananthus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="11" pageNumber="164" phylum="Tracheophyta" rank="species" species="speirsae">
<emphasis id="5CAFB914FF9DFF8C58B36032F4F9F7B8" bold="true" box="[491,717,2049,2070]" italics="true" pageId="11" pageNumber="164">Platananthus speirsae</emphasis>
<smallCapsWord id="6882F3DAFF9DFF8C5B8E6032F53CF7BB" baselines="2065,2065" box="[726,776,2049,2070]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pigg" pageId="11" pageNumber="164">PIGG</smallCapsWord>
et
<smallCapsWord id="6882F3DAFF9DFF8C5A746032F5BDF7BB" baselines="2065,2065" box="[812,905,2049,2070]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Stockey" pageId="11" pageNumber="164">STOCKEY</smallCapsWord>
</taxonomicName>
</emphasis>
axis with at least 9 attached pedunculate staminate inflorescences (arrows), Seven Mile Creek, Montana (orig.
<bibRefCitation id="0A4A18F7FF9DFF8C5B9B602DF576F79D" author="Ward, L. F." box="[707,834,2078,2099]" pageId="11" pageNumber="164" pagination="400 - 570" refId="ref35088" refString="Ward, L. F. (1885 b): Synopsis of the flora of the Laramie group. - U. S. Geol. Surv. Ann. Rep., 6: 400 - 570." type="journal article" year="1885">Ward 1885b</bibRefCitation>
, pl. 32, fig. 7), USNM 4225. Scale bars 5 cm.
</paragraph>
</caption>
<caption id="3AA4358EFF9AFF8B59D96FBDF33EF857" ID-DOI="http://doi.org/10.5281/zenodo.13183586" ID-Zenodo-Dep="13183586" httpUri="https://zenodo.org/record/13183586/files/figure.png" pageId="12" pageNumber="165" startId="12.[129,210,1934,1955]" targetBox="[129,1482,152,1901]" targetPageId="12" targetType="figure">
<paragraph id="6E646506FF9AFF8B59D96FBDF33EF857" blockId="12.[129,1482,1934,2041]" pageId="12" pageNumber="165">
<emphasis id="5CAFB914FF9AFF8B59D96FBDF57CF811" bold="true" pageId="12" pageNumber="165">
Text-fig. 6.
<taxonomicName id="A9DB1E85FF9AFF8B59AD6FBDF744F80D" box="[245,368,1934,1955]" class="Magnoliopsida" family="Platanaceae" kingdom="Plantae" order="Proteales" pageId="12" pageNumber="165" phylum="Tracheophyta" rank="family">Platanaceae</taxonomicName>
13.
<taxonomicName id="A9DB1E85FF9AFF8B58FF6FBDF4C3F80D" authority="(NEWBERRY)" authorityName="Manchester" authorityYear="2014" baseAuthorityName="Manchester" baseAuthorityYear="2014" box="[423,759,1934,1955]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="12" pageNumber="165" phylum="Tracheophyta" rank="species" species="raynoldsii" status="comb. nov.">
<emphasis id="5CAFB914FF9AFF8B58FF6FBDF447F80D" bold="true" box="[423,627,1934,1955]" italics="true" pageId="12" pageNumber="165">Platanites raynoldsii</emphasis>
(
<smallCapsWord id="6882F3DAFF9AFF8B5BD86FBDF4DBF80C" baselines="1950,1950" box="[640,751,1934,1955]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Newberry" pageId="12" pageNumber="165">NEWBERRY</smallCapsWord>
)
</taxonomicName>
<taxonomicNameLabel id="479C046FFF9AFF8B5BA66FBDF558F80D" box="[766,876,1934,1955]" pageId="12" pageNumber="165" rank="species">comb. nov.</taxonomicNameLabel>
1. Holotype of
<taxonomicName id="A9DB1E85FF9AFF8B5D526FBDF3B4F80D" authority="NEWBERRY (1868" authorityName="NEWBERRY" authorityYear="1868" box="[1034,1408,1934,1955]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="12" pageNumber="165" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9AFF8B5D526FBDF2F8F80D" bold="true" box="[1034,1228,1934,1955]" italics="true" pageId="12" pageNumber="165">Platanus raynoldsii</emphasis>
<bibRefCitation id="0A4A18F7FF9AFF8B5D8A6FBDF374F80C" author="Newberry, J. S." box="[1234,1344,1934,1955]" pageId="12" pageNumber="165" pagination="1 - 76" refId="ref31946" refString="Newberry, J. S. (1868): Notes on the later extinct floras of North America, with descriptions of some new species of fossil plants from the Cretaceous and tertiary strata. - Lyceum Nat. Hist. New York Ann., 9: 1 - 76. http: // dx. doi. org / 10.1111 / j. 1749 - 6632.1870. tb 00175. x" type="journal article" year="1868">
<smallCapsWord id="6882F3DAFF9AFF8B5D8A6FBDF374F80C" baselines="1950,1950" box="[1234,1344,1934,1955]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Newberry" pageId="12" pageNumber="165">NEWBERRY</smallCapsWord>
</bibRefCitation>
(1868
</taxonomicName>
,
<bibRefCitation id="0A4A18F7FF9AFF8B5CD46FBDF38AF80D" author="Newberry, J. S." box="[1420,1470,1934,1955]" pageId="12" pageNumber="165" pagination="1 - 295" refId="ref32108" refString="Newberry, J. S. (1898): The later extinct floras of North America. - U. S. Geol. Surv. Monogr., 35: 1 - 295, 68 pls." type="journal article" year="1898">1898</bibRefCitation>
), Banks of the Yellowstone River, Montana, USNM 7000. 2. Holotype of
</emphasis>
<taxonomicName id="A9DB1E85FF9AFF8B5A166F98F370F811" authority="MCIVER et BASINGER (1993)" authorityName="MCIVER et BASINGER" authorityYear="1993" box="[846,1348,1962,1984]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="12" pageNumber="165" phylum="Tracheophyta" rank="species" species="canadensis">
<emphasis id="5CAFB914FF9AFF8B5A166F98F2C9F810" bold="true" box="[846,1277,1962,1984]" pageId="12" pageNumber="165">
<emphasis id="5CAFB914FF9AFF8B5A166F98F217F86E" bold="true" box="[846,1059,1963,1984]" italics="true" pageId="12" pageNumber="165">Platanites canadensis</emphasis>
<bibRefCitation id="0A4A18F7FF9AFF8B5D716F99F248F810" author="McIver, E. E. &amp; Basinger, J. F." box="[1065,1148,1962,1983]" pageId="12" pageNumber="165" pagination="1 - 167" refId="ref31643" refString="McIver, E. E., Basinger, J. F. (1993): Flora of the Ravenscrag Formation (Paleocene), southwestern Saskatchewan, Canada. - Palaeontogr. Can., 10: 1 - 167." type="journal article" year="1993">
<smallCapsWord id="6882F3DAFF9AFF8B5D716F99F248F810" baselines="1978,1978" box="[1065,1148,1962,1983]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Mc" pageId="12" pageNumber="165">MCIVER</smallCapsWord>
</bibRefCitation>
et
<smallCapsWord id="6882F3DAFF9AFF8B5DC36F99F2C9F810" baselines="1978,1978" box="[1179,1277,1962,1983]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Basinger" pageId="12" pageNumber="165">BASINGER</smallCapsWord>
</emphasis>
(1993)
</taxonomicName>
, here treated as synonym of
<taxonomicName id="A9DB1E85FF9AFF8B584C6FF4F7B9F872" authorityName="NEWBERRY" authorityYear="1868" box="[276,397,1991,2012]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="12" pageNumber="165" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9AFF8B584C6FF4F7B9F872" bold="true" box="[276,397,1991,2012]" italics="true" pageId="12" pageNumber="165">P. raynoldsii</emphasis>
</taxonomicName>
, showing clearly the position of the lateral leaflets, US 3-66, Ravenscrag Formation, Saskatchewan, Canada. 3. Specimen from Yellowstone Bridge, Miles City, Montana. UF19021-39390. 4. Detail from fig. 2. Scale bars 5 cm.
</paragraph>
</caption>
<paragraph id="6E646506FF9BFF8A59C868ABF4B8FDE3" blockId="13.[106,745,152,1829]" pageId="13" pageNumber="166">
<bibRefCitation id="0A4A18F7FF9BFF8A59C868ABF7E0FF01" author="McIver, E. E. &amp; Basinger, J. F." box="[144,468,152,175]" pageId="13" pageNumber="166" pagination="1 - 167" refId="ref31643" refString="McIver, E. E., Basinger, J. F. (1993): Flora of the Ravenscrag Formation (Paleocene), southwestern Saskatchewan, Canada. - Palaeontogr. Can., 10: 1 - 167." type="journal article" year="1993">McIver and Basinger (1993)</bibRefCitation>
illustrated a beautifully complete compound leaf of
<taxonomicName id="A9DB1E85FF9BFF8A58C7688BF439FF61" box="[415,525,184,207]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="genus">
<emphasis id="5CAFB914FF9BFF8A58C7688BF439FF61" box="[415,525,184,207]" italics="true" pageId="13" pageNumber="166">Platanites</emphasis>
</taxonomicName>
from the Paleocene Ravenscrag Formation of southern
<collectingRegion id="AC1FABE4FF9BFF8A58B368EBF4B2FF41" box="[491,646,216,239]" country="Canada" name="Saskatchewan" pageId="13" pageNumber="166">Saskatchewan</collectingRegion>
,
<collectingCountry id="16CC2596FF9BFF8A5BCA68EBF4D1FF41" box="[658,741,216,239]" name="Canada" pageId="13" pageNumber="166">Canada</collectingCountry>
, showing a pair of lateral elliptical leaves attached to the “petiole” of a leaf matching architecturally to the
<typeStatus id="B160DBA4FF9BFF8A5BD36924F4DDFE80" box="[651,745,279,302]" pageId="13" pageNumber="166" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A9DB1E85FF9BFF8A59D46904F75CFEE0" authorityName="NEWBERRY" authorityYear="1868" box="[140,360,311,334]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9BFF8A59D46904F75CFEE0" box="[140,360,311,334]" italics="true" pageId="13" pageNumber="166">Platanus raynoldsii</emphasis>
</taxonomicName>
(reillustrated here,
<figureCitation id="F6E07983FF9BFF8A5B146904F4E9FEE0" box="[588,733,311,334]" captionStart="Text-fig" captionStartId="12.[129,210,1934,1955]" captionTargetBox="[129,1482,152,1901]" captionTargetId="figure-11@12.[156,1454,178,1867]" captionTargetPageId="12" captionText="Text-fig. 6. Platanaceae 13. Platanites raynoldsii (NEWBERRY) comb. nov. 1. Holotype of Platanus raynoldsii NEWBERRY (1868, 1898), Banks of the Yellowstone River, Montana, USNM 7000. 2. Holotype of Platanites canadensis MCIVER et BASINGER (1993), here treated as synonym of P. raynoldsii, showing clearly the position of the lateral leaflets, US 3-66, Ravenscrag Formation, Saskatchewan, Canada. 3. Specimen from Yellowstone Bridge, Miles City, Montana. UF19021-39390. 4. Detail from fig. 2. Scale bars 5 cm." figureDoi="http://doi.org/10.5281/zenodo.13183586" httpUri="https://zenodo.org/record/13183586/files/figure.png" pageId="13" pageNumber="166">Text-fig. 6.2</figureCitation>
). Hence, the petiole of
<taxonomicName id="A9DB1E85FF9BFF8A58086964F7E2FEC0" authorityName="NEWBERRY" authorityYear="1868" box="[336,470,343,366]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9BFF8A58086964F7E2FEC0" box="[336,470,343,366]" italics="true" pageId="13" pageNumber="166">P. raynoldsii</emphasis>
</taxonomicName>
is actually the rachis of a compound leaf. Because Newberrys species has long been considered to be a simple leaf, as expected for extant
<taxonomicName id="A9DB1E85FF9BFF8A59326985F421FE63" authority=", McIver and Basinger (1993)" authorityName=", McIver and Basinger" authorityYear="1993" box="[106,533,438,461]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="genus">
<emphasis id="5CAFB914FF9BFF8A59326985F6FFFE63" box="[106,203,438,461]" italics="true" pageId="13" pageNumber="166">Platanus</emphasis>
,
<bibRefCitation id="0A4A18F7FF9BFF8A59826985F421FE63" author="McIver, E. E. &amp; Basinger, J. F." box="[218,533,438,461]" pageId="13" pageNumber="166" pagination="1 - 167" refId="ref31643" refString="McIver, E. E., Basinger, J. F. (1993): Flora of the Ravenscrag Formation (Paleocene), southwestern Saskatchewan, Canada. - Palaeontogr. Can., 10: 1 - 167." type="journal article" year="1993">McIver and Basinger (1993)</bibRefCitation>
</taxonomicName>
considered the the Ravenscrag material to be distinct and described it as a new species,
<emphasis id="5CAFB914FF9BFF8A599D69C5F782FDA3" box="[197,438,502,525]" italics="true" pageId="13" pageNumber="166">
<taxonomicName id="A9DB1E85FF9BFF8A599D69C5F786FDA3" box="[197,434,502,525]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="canadensis">Platanites canadensis</taxonomicName>
.
</emphasis>
They used the generic name based on similar compound platanaceous leaves from
<collectingCountry id="16CC2596FF9BFF8A59326A05F6F3FDE3" box="[106,199,566,589]" name="United Kingdom" pageId="13" pageNumber="166">Scotland</collectingCountry>
(
<taxonomicName id="A9DB1E85FF9BFF8A598F6A05F783FDE2" authority="FORBES" authorityName="FORBES" box="[215,439,566,589]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="hebridicus">
<emphasis id="5CAFB914FF9BFF8A598F6A05F756FDE3" box="[215,354,566,589]" italics="true" pageId="13" pageNumber="166">P. hebridicus</emphasis>
<smallCapsWord id="6882F3DAFF9BFF8A58306A05F783FDE2" baselines="583,583" box="[360,439,566,589]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Forbes" pageId="13" pageNumber="166">FORBES</smallCapsWord>
</taxonomicName>
;
<bibRefCitation id="0A4A18F7FF9BFF8A589D6A05F449FDE3" author="Crane, P. R. &amp; Manchester, S. R. &amp; Dilcher, D. L." box="[453,637,566,589]" pageId="13" pageNumber="166" pagination="503 - 517" refId="ref25333" refString="Crane, P. R., Manchester, S. R., Dilcher, D. L. (1988): Morphology and Phylogenetic significance of the angiosperm Platanites hebridicus from the Palaeocene of Scotland. - Palaeontology, 31: 503 - 517." type="journal article" year="1988">Crane et al. 1988</bibRefCitation>
).
</paragraph>
<paragraph id="6E646506FF9BFF8A59C86A66F716FB07" blockId="13.[106,745,152,1829]" pageId="13" pageNumber="166">
Close comparison between Newberrys
<typeStatus id="B160DBA4FF9BFF8A5B7F6A66F4B6FDC2" box="[551,642,597,620]" pageId="13" pageNumber="166" type="holotype">holotype</typeStatus>
specimen of
<taxonomicName id="A9DB1E85FF9BFF8A59DC6A46F737FD22" authorityName="NEWBERRY" authorityYear="1868" box="[132,259,629,652]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9BFF8A59DC6A46F737FD22" box="[132,259,629,652]" italics="true" pageId="13" pageNumber="166">P. raynoldsii</emphasis>
</taxonomicName>
(
<figureCitation id="F6E07983FF9BFF8A58566A46F7B9FD22" box="[270,397,629,652]" captionStart="Text-fig" captionStartId="12.[129,210,1934,1955]" captionTargetBox="[129,1482,152,1901]" captionTargetId="figure-11@12.[156,1454,178,1867]" captionTargetPageId="12" captionText="Text-fig. 6. Platanaceae 13. Platanites raynoldsii (NEWBERRY) comb. nov. 1. Holotype of Platanus raynoldsii NEWBERRY (1868, 1898), Banks of the Yellowstone River, Montana, USNM 7000. 2. Holotype of Platanites canadensis MCIVER et BASINGER (1993), here treated as synonym of P. raynoldsii, showing clearly the position of the lateral leaflets, US 3-66, Ravenscrag Formation, Saskatchewan, Canada. 3. Specimen from Yellowstone Bridge, Miles City, Montana. UF19021-39390. 4. Detail from fig. 2. Scale bars 5 cm." figureDoi="http://doi.org/10.5281/zenodo.13183586" httpUri="https://zenodo.org/record/13183586/files/figure.png" pageId="13" pageNumber="166">Text-fig. 6.1</figureCitation>
) and the
<typeStatus id="B160DBA4FF9BFF8A58BF6A46F474FD22" box="[487,576,629,652]" pageId="13" pageNumber="166" type="holotype">holotype</typeStatus>
of
<taxonomicName id="A9DB1E85FF9BFF8A5B386A46F4DCFD22" box="[608,744,629,652]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="canadensis">
<emphasis id="5CAFB914FF9BFF8A5B386A46F4DCFD22" box="[608,744,629,652]" italics="true" pageId="13" pageNumber="166">P. canadensis</emphasis>
</taxonomicName>
(
<figureCitation id="F6E07983FF9BFF8A59296AA6F6C0FD02" box="[113,244,661,684]" captionStart="Text-fig" captionStartId="12.[129,210,1934,1955]" captionTargetBox="[129,1482,152,1901]" captionTargetId="figure-11@12.[156,1454,178,1867]" captionTargetPageId="12" captionText="Text-fig. 6. Platanaceae 13. Platanites raynoldsii (NEWBERRY) comb. nov. 1. Holotype of Platanus raynoldsii NEWBERRY (1868, 1898), Banks of the Yellowstone River, Montana, USNM 7000. 2. Holotype of Platanites canadensis MCIVER et BASINGER (1993), here treated as synonym of P. raynoldsii, showing clearly the position of the lateral leaflets, US 3-66, Ravenscrag Formation, Saskatchewan, Canada. 3. Specimen from Yellowstone Bridge, Miles City, Montana. UF19021-39390. 4. Detail from fig. 2. Scale bars 5 cm." figureDoi="http://doi.org/10.5281/zenodo.13183586" httpUri="https://zenodo.org/record/13183586/files/figure.png" pageId="13" pageNumber="166">Text-fig. 6.2</figureCitation>
) indicates that they are architecturally identical, leading to the conclusion that they are conspecific. Other compound leaves of
<taxonomicName id="A9DB1E85FF9BFF8A58386AE6F7DAFD42" authorityName="NEWBERRY" authorityYear="1868" box="[352,494,725,748]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9BFF8A58386AE6F7DAFD42" box="[352,494,725,748]" italics="true" pageId="13" pageNumber="166">P. raynoldsii</emphasis>
</taxonomicName>
have been recovered occasionally in the Late Cretaceous and Paleocene of North America (e.g., specimens figured by
<bibRefCitation id="0A4A18F7FF9BFF8A58856B27F44BFC85" author="Knowlton, F. H." box="[477,639,788,811]" pageId="13" pageNumber="166" pagination="1 - 142" refId="ref29059" refString="Knowlton, F. H. (1930): The flora of the Denver and associated formations of Colorado: - U. S. Geol. Surv. Prof. Paper, 155: 1 - 142." type="journal article" year="1930">Knowlton 1930</bibRefCitation>
, pl 45, fig. 10 as
<taxonomicName id="A9DB1E85FF9BFF8A59FB6B07F7C5FCE5" authority="LESQUEREUX" authorityName="LESQUEREUX" box="[163,497,820,843]" class="Magnoliopsida" family="Vitaceae" genus="Negundo" kingdom="Plantae" order="Vitales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="decurrens">
<emphasis id="5CAFB914FF9BFF8A59FB6B07F75FFCE5" box="[163,363,820,843]" italics="true" pageId="13" pageNumber="166">Negundo decurrens</emphasis>
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</taxonomicName>
, and by
<bibRefCitation id="0A4A18F7FF9BFF8A5B1E6B07F4D1FCE5" author="Ellis, B. &amp; Johnson, K. R. &amp; Dunn, R. E." box="[582,741,820,843]" pageId="13" pageNumber="166" pagination="73 - 100" refId="ref26012" refString="Ellis, B., Johnson, K. R., Dunn, R. E. (2003): Evidence for an in situ early Paleocene rainforest from Castle Rock, Colorado. - Rocky Mt. Geol., 38 (1): 73 - 100. http: // dx. doi. org / 10.2113 / gsrocky. 38.1.173" type="journal article" year="2003">Ellis et al. 2003</bibRefCitation>
, fig. 9a as
<taxonomicName id="A9DB1E85FF9BFF8A59806B67F4D6FCC4" authority="(LESQUEREUX) K. JOHNSON" authorityName="K. JOHNSON" baseAuthorityName="LESQUEREUX" box="[216,738,852,875]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="marginata">
<emphasis id="5CAFB914FF9BFF8A59806B67F78DFCC5" box="[216,441,852,875]" italics="true" pageId="13" pageNumber="166">Platanites marginata</emphasis>
(
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) K.
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, but usually the petiole/rachis lacks lateral leaflets, giving appearance that “they leaves” were simple. We do not know whether this is due to deciduousness and/or damage prior to deposition, or if
<taxonomicName id="A9DB1E85FF9BFF8A58746BE0F783FC44" authorityName="NEWBERRY" authorityYear="1868" box="[300,439,979,1002]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9BFF8A58746BE0F783FC44" box="[300,439,979,1002]" italics="true" pageId="13" pageNumber="166">P. raynoldsii</emphasis>
</taxonomicName>
might have been variable, producing both simple and compound leaves on the same individual (perhaps a difference between seedling and mature leaves, for example). Nevertheless, the tendency to produce compound leaves distinguishes this species from all extant species of
<taxonomicName id="A9DB1E85FF9BFF8A59806C41F700FB27" authorityName=", McIver and Basinger" authorityYear="1993" box="[216,308,1138,1161]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="genus">
<emphasis id="5CAFB914FF9BFF8A59806C41F700FB27" box="[216,308,1138,1161]" italics="true" pageId="13" pageNumber="166">Platanus</emphasis>
</taxonomicName>
, and justifies its assignment to the extinct genus,
<taxonomicName id="A9DB1E85FF9BFF8A59EB6CA1F72FFB07" box="[179,283,1170,1193]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="genus">
<emphasis id="5CAFB914FF9BFF8A59EB6CA1F72FFB07" box="[179,283,1170,1193]" italics="true" pageId="13" pageNumber="166">Platanites</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="6E646506FF9BFF8A59C86C81F489F88B" blockId="13.[106,745,152,1829]" pageId="13" pageNumber="166">
Prior to the realization that this species represents a compound-leaved plant,
<bibRefCitation id="0A4A18F7FF9BFF8A58216CE1F421FB47" author="Brown, R. W." box="[377,533,1234,1257]" pageId="13" pageNumber="166" pagination="1 - 119" refId="ref24721" refString="Brown, R. W. (1962): Paleocene flora of the Rocky Mountains and Great Plains. - U. S. Geol. Surv. Prof. Pap., 375: 1 - 119." type="journal article" year="1962">Brown (1962)</bibRefCitation>
, adopted the name
<taxonomicName id="A9DB1E85FF9BFF8A59326CC1F70EFAA7" authorityName="NEWBERRY" authorityYear="1868" box="[106,314,1266,1289]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9BFF8A59326CC1F70EFAA7" box="[106,314,1266,1289]" italics="true" pageId="13" pageNumber="166">Platanus raynoldsii</emphasis>
</taxonomicName>
(and included Newberrys
<taxonomicName id="A9DB1E85FF9BFF8A5B066CC1F4F8FAA7" authorityName="Newberry" authorityYear="1868" box="[606,716,1266,1289]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="haydeni">
<emphasis id="5CAFB914FF9BFF8A5B066CC1F4F8FAA7" box="[606,716,1266,1289]" italics="true" pageId="13" pageNumber="166">P. haydeni</emphasis>
</taxonomicName>
in synonymy), for most weakly trilobate platanaceous leaves from the Paleocene of the Rocky Mountains and Great Plains region and provided a thorough synonymy of earlier published specimens from the works of Newberry, Lesquereux, Ward and Knowlton. The same epithet was applied to similar laminae from the late Cretaceous and Paleocene of Far Eastern
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(
<bibRefCitation id="0A4A18F7FF9BFF8A59296DE3F726FA49" author="Krassilov, V." box="[113,274,1488,1511]" pageId="13" pageNumber="166" refId="ref29130" refString="Krassilov, V. (1976): Tsagayanskaya flora Amurskoy oblasti. Nauka, Moskva (in Russian)." type="book" year="1976">Krassilov 1976</bibRefCitation>
). Based mainly on her investigations of the far eastern Russian material,
<bibRefCitation id="0A4A18F7FF9BFF8A582E6DC3F41CF9A9" author="Moiseeva, M. G." box="[374,552,1520,1543]" pageId="13" pageNumber="166" pagination="92 - 105" refId="ref31804" refString="Moiseeva, M. G. (2008): New Angiosperms from the Maastrichtian of the Amaam Lagoon Area (Northeastern Russia). - Paleont. J., 42 (3): 313 - 327. (orig in Russian: Paleont. Zhurn., 2008 (3): 92 - 105.)" type="journal article" year="2008">Moiseeva (2008)</bibRefCitation>
established a new combination,
<taxonomicName id="A9DB1E85FF9BFF8A585B6E23F412F989" box="[259,550,1552,1575]" class="Magnoliopsida" family="Vitaceae" genus="Ettingshausenia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Vitales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9BFF8A585B6E23F412F989" box="[259,550,1552,1575]" italics="true" pageId="13" pageNumber="166">Ettingshausenia raynoldsii</emphasis>
</taxonomicName>
, for the species, applying the fossil-genus name
<taxonomicName id="A9DB1E85FF9BFF8A58896E03F686F9C8" authority="STIEHLER (1857)" authorityName="STIEHLER" authorityYear="1857" class="Magnoliopsida" family="Vitaceae" genus="Ettingshausenia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Vitales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="genus">
<emphasis id="5CAFB914FF9BFF8A58896E03F44FF9E9" box="[465,635,1584,1607]" italics="true" pageId="13" pageNumber="166">Ettingshausenia</emphasis>
<bibRefCitation id="0A4A18F7FF9BFF8A5BDF6E03F686F9C8" author="Stiehler, A. W." pageId="13" pageNumber="166" pagination="71 - 80" refId="ref33689" refString="Stiehler, A. W. (1857): Beitrage zur Kenntniss der vorweltlichen Flora des Kreidegebirges im Harze. II. Die Flora des Langeberges bei Quedlinburg. - Palaeontographica, 5: 71 - 80." type="journal article" year="1857">
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<emphasis id="5CAFB914FF9BFF8A59326E7CF646F9C8" bold="true" box="[106,114,1615,1638]" pageId="13" pageNumber="166">(</emphasis>
1857)
</bibRefCitation>
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. However,
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<emphasis id="5CAFB914FF9BFF8A58706E7CF7FAF9C8" box="[296,462,1615,1638]" italics="true" pageId="13" pageNumber="166">Ettingshausenia</emphasis>
</taxonomicName>
is believed to be a simpleleaved genus. It is possible that the genus
<taxonomicName id="A9DB1E85FF9BFF8A5B706E5CF4FBF928" authorityName="STIEHLER" authorityYear="1857" box="[552,719,1647,1670]" class="Magnoliopsida" family="Vitaceae" genus="Ettingshausenia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Vitales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="genus">
<emphasis id="5CAFB914FF9BFF8A5B706E5CF4FBF928" box="[552,719,1647,1670]" italics="true" pageId="13" pageNumber="166">Ettingshausenia</emphasis>
</taxonomicName>
is appropriately applied to theAsian fossils, but a different specific epithet is needed from that applied to the North American species considered here. It is not appropriate to use the name
<taxonomicName id="A9DB1E85FF9BFF8A59326EDCF724F8A8" authorityName="STIEHLER" authorityYear="1857" box="[106,272,1775,1798]" class="Magnoliopsida" family="Vitaceae" genus="Ettingshausenia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Vitales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="genus">
<emphasis id="5CAFB914FF9BFF8A59326EDCF724F8A8" box="[106,272,1775,1798]" italics="true" pageId="13" pageNumber="166">Ettingshausenia</emphasis>
</taxonomicName>
in the case of the
<typeStatus id="B160DBA4FF9BFF8A589B6EDCF7C4F8A8" box="[451,496,1775,1798]" pageId="13" pageNumber="166">type</typeStatus>
material of
<taxonomicName id="A9DB1E85FF9BFF8A5B306EDCF6ECF88B" authority="NEWBERRY" authorityName="NEWBERRY" class="Magnoliopsida" family="Vitaceae" genus="Ettingshausenia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Vitales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="species" species="raynoldsii">
<emphasis id="5CAFB914FF9BFF8A5B306EDCF4DCF8A8" box="[616,744,1775,1798]" italics="true" pageId="13" pageNumber="166">P. raynoldsii</emphasis>
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, which conforms instead to
<taxonomicName id="A9DB1E85FF9BFF8A58A36F3DF489F88B" authority="FORBES." authorityName="FORBES." box="[507,701,1806,1829]" class="Magnoliopsida" family="Platanaceae" genus="Platanites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Proteales" pageId="13" pageNumber="166" phylum="Tracheophyta" rank="genus">
<emphasis id="5CAFB914FF9BFF8A58A36F3DF450F88B" box="[507,612,1806,1829]" italics="true" pageId="13" pageNumber="166">Platanites</emphasis>
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.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>