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<document id="095A4931BC33A547E5CBF3827D6CAC41" ID-DOI="10.5281/zenodo.169943" ID-GBIF-Dataset="a80741a3-d6db-4474-a7e6-79293fb0e80f" ID-ISSN="1175­5326" ID-Zenodo-Dep="169943" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459748760006" checkinUser="plazi" docAuthor="Will, Kipling W." docDate="2005" docId="5D21241C5C7F48693F184A5FFE99FD59" docLanguage="en" docName="zt01049p017.pdf" docOrigin="Zootaxa 1049" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Stolonis yasuni Will, new species" docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="A1185C645C7348673E104913FFD3FFC7" masterDocTitle="The Neotropical genera Oxycrepis Reiche and Stolonis Motschulsky: a taxonomic review, key to the described species and description of new Stolonis species from Ecuador (Coleoptera: Carabidae: Loxandrini)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="13" updateTime="1698211146980" updateUser="plazi">
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<mods:title id="640AA777FE35554593349A7714E672E5">The Neotropical genera Oxycrepis Reiche and Stolonis Motschulsky: a taxonomic review, key to the described species and description of new Stolonis species from Ecuador (Coleoptera: Carabidae: Loxandrini)</mods:title>
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<mods:namePart id="CE4E0251E33383A7BB320E48B8653323">Will, Kipling W.</mods:namePart>
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<mods:date id="3E47B8D089BA63DE3E677A31B71C489B">2005</mods:date>
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<treatment id="5D21241C5C7F48693F184A5FFE99FD59" ID-DOI="http://doi.org/10.5281/zenodo.6266246" ID-GBIF-Taxon="119337162" ID-Zenodo-Dep="6266246" LSID="urn:lsid:plazi:treatment:5D21241C5C7F48693F184A5FFE99FD59" httpUri="http://treatment.plazi.org/id/5D21241C5C7F48693F184A5FFE99FD59" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="9D92C6815C7F486B3F184A5FFDD6FC4E" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="D537950A5C7F486B3F184A5FFD48FCA1" blockId="12.[264,667,844,905]" box="[264,667,844,870]" pageId="12" pageNumber="13">
<heading id="8E7F22665C7F486B3F184A5FFD48FCA1" bold="true" box="[264,667,844,870]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="E7FC49185C7F486B3F184A5FFD48FCA1" bold="true" box="[264,667,844,870]" pageId="12" pageNumber="13">
<taxonomicName id="1288EE895C7F486B3F184A5FFE2DFCA1" ID-CoL="8LNLW" authority="Will" authorityName="Will" box="[264,510,844,870]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="yasuni" status="sp. nov.">
<emphasis id="E7FC49185C7F486B3F184A5FFE13FCA1" bold="true" box="[264,448,844,870]" italics="true" pageId="12" pageNumber="13">Stolonis yasuni</emphasis>
Will
</taxonomicName>
,
<taxonomicNameLabel id="FCCFF4635C7F486B3C1B4A5FFD48FCA1" box="[523,667,844,870]" pageId="12" pageNumber="13" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D537950A5C7F486B3F184A7CFDD6FC4E" blockId="12.[264,667,844,905]" box="[264,517,879,905]" pageId="12" pageNumber="13">
(
<figureCitation id="4DB3898F5C7F486B3F014A7CFEB1FC4E" box="[273,354,879,905]" captionStart="FIGURE 2" captionStartId="8.[264,367,1326,1350]" captionTargetBox="[337,1321,462,1293]" captionTargetId="figure@8.[267,1326,452,1303]" captionTargetPageId="8" captionText="FIGURE 2. Pronota of: A— Stolonis tapiai, B— Stolonis scortensis, C— Stolonis spinosus, D— Stolonis yasuni, E— Stolonis catenarius." httpUri="https://zenodo.org/record/169945/files/figure.png" pageId="12" pageNumber="13">Figs. 2</figureCitation>
D, 3MP, 4B)
</paragraph>
</subSubSection>
<subSubSection id="9D92C6815C7F486B3F184AADFB5BFB0E" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="D537950A5C7F486B3F184AADFB5BFB0E" blockId="12.[264,1323,958,1905]" pageId="12" pageNumber="13">
<emphasis id="E7FC49185C7F486B3F184AADFE13FC1F" bold="true" box="[264,448,958,984]" pageId="12" pageNumber="13">
<typeStatus id="0A332BA85C7F486B3F184AADFE90FC1F" box="[264,323,958,984]" pageId="12" pageNumber="13">Type</typeStatus>
Material
</emphasis>
:
<typeStatus id="0A332BA85C7F486B3FCA4AACFDAAFC1E" box="[474,633,959,985]" pageId="12" pageNumber="13" type="holotype">HOLOTYPE</typeStatus>
: ɗ (
<collectionCode id="B3990DCF5C7F486B3CA84AACFCDEFC1E" LSID="urn:lsid:biocol.org:col:1013" box="[696,781,959,985]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="12" pageNumber="13">EMEC</collectionCode>
) genitalia dissected. Labeled: &quot;00°40'36S&quot;/
<geoCoordinate id="B0BCF3CD5C7F486B3F184AF4FE4BFBC6" box="[264,408,999,1025]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-76.40056">76°24'02&quot;W</geoCoordinate>
,
<collectingCountry id="AD9FD59A5C7F486B3FA04AF4FD92FBC6" box="[432,577,999,1025]" name="Ecuador" pageId="12" pageNumber="13">ECUADOR</collectingCountry>
,Napo Prov.,
<taxonomicName id="1288EE895C7F486B3CFF4AF4FCECFBC6" box="[751,831,999,1025]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="16" phylum="Arthropoda" rank="species" species="yasuni">Yasuni</taxonomicName>
Scientific Station, 19:IV:
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1998,
<specimenCount id="C38E5E835C7F486B3AF04AF4FAF4FBC6" box="[1248,1319,999,1025]" pageId="12" pageNumber="13" type="generic">210m</specimenCount>
</quantity>
. Col.K.Will&quot; /[red label]&quot;
<taxonomicName id="1288EE895C7F486B3C524D1CFCA6FBEE" authority="K.Will" authorityName="K.Will" box="[578,885,1039,1065]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="yasuni">Stolonis yasuni, K.Will</taxonomicName>
&quot;/ &quot;U.C.Berkeley
<collectionCode id="B3990DCF5C7F486B3A4F4D1CFB66FBEE" LSID="urn:lsid:biocol.org:col:1013" box="[1119,1205,1039,1065]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="12" pageNumber="13">EMEC</collectionCode>
1003668&quot;. ALLOTYPE: Ψ (
<collectionCode id="B3990DCF5C7F486B3FC54D24FDFEFB96" LSID="urn:lsid:biocol.org:col:1013" box="[469,557,1079,1105]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="12" pageNumber="13">EMEC</collectionCode>
), same data and deposition as
<typeStatus id="0A332BA85C7F486B3D894D24FBD3FB96" box="[921,1024,1079,1105]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
except database number, &quot;
<collectionCode id="B3990DCF5C7F486B3F014D4CFEBBFBBE" LSID="urn:lsid:biocol.org:col:1013" box="[273,360,1119,1145]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="12" pageNumber="13">EMEC</collectionCode>
1003669&quot;.
<typeStatus id="0A332BA85C7F486B3FF64D4CFD5FFBBE" box="[486,652,1119,1145]" pageId="12" pageNumber="13" type="paratype">PARATYPES</typeStatus>
:
<emphasis id="E7FC49185C7F486B3C864D4DFC01FBBF" bold="true" box="[662,978,1118,1144]" pageId="12" pageNumber="13">
<collectingCountry id="AD9FD59A5C7F486B3C864D4DFCD5FBBF" box="[662,774,1118,1144]" name="Ecuador" pageId="12" pageNumber="13">Ecuador</collectingCountry>
: Orellana Prov.:
</emphasis>
<taxonomicName id="1288EE895C7F486B3DC94D4CFBFAFBBE" box="[985,1065,1119,1145]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="16" phylum="Arthropoda" rank="species" species="yasuni">Yasuni</taxonomicName>
Scientific Station: 1Ψ same data as
<typeStatus id="0A332BA85C7F486B3FB34D94FDDAFB66" box="[419,521,1159,1185]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
except database number
<collectionCode id="B3990DCF5C7F486B3D214D94FC56FB66" LSID="urn:lsid:biocol.org:col:1013" box="[817,901,1159,1185]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="12" pageNumber="13">EMEC</collectionCode>
1003680; 1ɗ and 1Ψ (
<collectionCode id="B3990DCF5C7F486B3A954D94FB08FB66" box="[1157,1243,1159,1185]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="12" pageNumber="13">QCAZ</collectionCode>
), same data as
<typeStatus id="0A332BA85C7F486B3F704DBCFE14FB0E" box="[352,455,1199,1225]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
except database numbers,
<collectionCode id="B3990DCF5C7F486B3D114DBCFC86FB0E" LSID="urn:lsid:biocol.org:col:1013" box="[769,853,1199,1225]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="12" pageNumber="13">EMEC</collectionCode>
1003670,
<collectionCode id="B3990DCF5C7F486B3DD84DBCFBCEFB0E" LSID="urn:lsid:biocol.org:col:1013" box="[968,1053,1199,1225]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="12" pageNumber="13">EMEC</collectionCode>
1003671.
</paragraph>
</subSubSection>
<subSubSection id="9D92C6815C7F486B3F284DC5FCFFFADE" pageId="12" pageNumber="13" type="etymology">
<paragraph id="D537950A5C7F486B3F284DC5FCFFFADE" blockId="12.[264,1323,958,1905]" pageId="12" pageNumber="13">
<emphasis id="E7FC49185C7F486B3F284DC5FE6EFB37" bold="true" box="[312,445,1238,1264]" pageId="12" pageNumber="13">Etymology</emphasis>
. Name is a noun in apposition of the name of the
<typeStatus id="0A332BA85C7F486B3A244DC4FBB4FB36" box="[1076,1127,1239,1265]" pageId="12" pageNumber="13">type</typeStatus>
locality,
<taxonomicName id="1288EE895C7F486B3ACB4DC4FAF8FB36" box="[1243,1323,1239,1265]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="16" phylum="Arthropoda" rank="species" species="yasuni">Yasuni</taxonomicName>
Scientific Station, Orellana Province,
<collectingCountry id="AD9FD59A5C7F486B3CD44DECFCFBFADE" box="[708,808,1279,1305]" name="Ecuador" pageId="12" pageNumber="13">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9D92C6815C7F486B3F284C35FCD8FA56" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="D537950A5C7F486B3F284C35FCD8FA56" blockId="12.[264,1323,958,1905]" pageId="12" pageNumber="13">
<emphasis id="E7FC49185C7F486B3F284C35FE63FA87" bold="true" box="[312,432,1318,1344]" pageId="12" pageNumber="13">Diagnosis</emphasis>
. Differs from other
<taxonomicName id="1288EE895C7F486B3C8F4C34FD2EFA87" box="[671,765,1319,1344]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="E7FC49185C7F486B3C8F4C34FD2EFA87" box="[671,765,1319,1344]" italics="true" pageId="12" pageNumber="13">Stolonis</emphasis>
</taxonomicName>
species by the combination of broad pronotum (
<figureCitation id="4DB3898F5C7F486B3F014C5CFE84FAAE" box="[273,343,1359,1385]" captionStart="FIGURE 2" captionStartId="8.[264,367,1326,1350]" captionTargetBox="[337,1321,462,1293]" captionTargetId="figure@8.[267,1326,452,1303]" captionTargetPageId="8" captionText="FIGURE 2. Pronota of: A— Stolonis tapiai, B— Stolonis scortensis, C— Stolonis spinosus, D— Stolonis yasuni, E— Stolonis catenarius." httpUri="https://zenodo.org/record/169945/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
D) with broadly expanded lateral margins, immaculate elytra and basally crenulate and apically nearly smooth elytral intervals.
</paragraph>
</subSubSection>
<subSubSection id="9D92C6815C7F48693F284C8DFE99FD59" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph id="D537950A5C7F486B3F284C8DFADAF9CE" blockId="12.[264,1323,958,1905]" pageId="12" pageNumber="13">
<emphasis id="E7FC49185C7F486B3F284C8DFE1AFA7F" bold="true" box="[312,457,1438,1464]" pageId="12" pageNumber="13">Description</emphasis>
. Length
<quantity id="127038EF5C7F486B3C294C8CFD58FA7E" box="[569,651,1439,1465]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="12" pageNumber="13" unit="mm" value="7.4">7.4mm</quantity>
(7.0
<quantity id="127038EF5C7F486B3CC54C8CFCF8FA7E" box="[725,811,1439,1465]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="13" unit="mm" value="7.5">7.5mm</quantity>
). Dorsal surface deep black, shiny, lightly iridescent; antennomeres 13 brunneous; 46 and 11 black, 710 white; mouth parts brunneous and legs flavotestaceous with coxae and ventral region of femora infuscated.
</paragraph>
<paragraph id="D537950A5C7F486B3F284F04FCDFF96E" blockId="12.[264,1323,958,1905]" pageId="12" pageNumber="13">Form of head average build; eyes moderately prominent; slightly constricted behind eyes; frontal impressions distinct, shallow, rectangular, short, length less than half distance from base of clypeus to anterior supraorbital setae; frons between eyes shiny, scarcely iridescent near base, microlines not evident.</paragraph>
<paragraph id="D537950A5C7F486A3F284FA4FEB9FD59" blockId="12.[264,1323,958,1905]" lastBlockId="13.[264,1323,284,670]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
Pronotum (
<figureCitation id="4DB3898F5C7F486B3FAD4FA4FDD5F916" box="[445,518,1719,1745]" captionStart="FIGURE 2" captionStartId="8.[264,367,1326,1350]" captionTargetBox="[337,1321,462,1293]" captionTargetId="figure@8.[267,1326,452,1303]" captionTargetPageId="8" captionText="FIGURE 2. Pronota of: A— Stolonis tapiai, B— Stolonis scortensis, C— Stolonis spinosus, D— Stolonis yasuni, E— Stolonis catenarius." httpUri="https://zenodo.org/record/169945/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
D); clearly broader than long; narrowly constricted at base; anterior submarginal sulcus deep and complete; basal impressions of moderate length, 1/3 length of pronotum or less, very shallow apically, divergent apically when evident; lateral margins broadly rounded and widely explanate, constricted just apicad hind setae; not evident basad hind setae; no angular process at hind setae; shallowly punctate over base and in basal impressions. Elytra, elongate oval, length
<quantity id="127038EF5C7E486A3DA9480FFBDCFEF1" box="[953,1039,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="13" pageNumber="14" unit="mm" value="4.3">4.3mm</quantity>
, overall width
<quantity id="127038EF5C7E486A3ADF480FFAF5FEF1" box="[1231,1318,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="14" unit="mm" value="2.8">2.8mm</quantity>
; slightly convex, rounded and notably sinuate near apex; humeri sloped and rounded; striae shallowly and broadly punctate, punctures less evident to impunctate apically; intervals, broad, slightly convex, clearly crenulate in basal half and laterally, not or very superficially crenulate apical of discal setae on interval 3. Legs, slender; meso­ and metatarsi with prominent external sulcus; fifth tarsomeres ventrally glabrous or with few very small setae. Ventral surface shiny, clearly iridescent; mesosternum with 810 deep, broad punctures; metasternum laterally with 68 deep broad punctures; metepisternum with medial sulcus very deeply impressed, slightly punctate, slightly longer than wide (l/ w= 1.3).
</paragraph>
<caption id="81F7C5825C7E486A3F184C48FC71FA7F" httpUri="https://zenodo.org/record/169947/files/figure.png" pageId="13" pageNumber="14" targetBox="[293,1293,740,1348]" targetPageId="13">
<paragraph id="D537950A5C7E486A3F184C48FC71FA7F" blockId="13.[264,1323,1371,1464]" pageId="13" pageNumber="14">
<emphasis id="E7FC49185C7E486A3F184C48FE5FFAB4" bold="true" box="[264,396,1371,1395]" pageId="13" pageNumber="14">FIGURE 4.</emphasis>
Female reproductive tract, ventral views of: A—
<taxonomicName id="1288EE895C7E486A3DB94C4FFB97FAB4" box="[937,1092,1372,1395]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tapiai">
<emphasis id="E7FC49185C7E486A3DB94C4FFB97FAB4" box="[937,1092,1372,1395]" italics="true" pageId="13" pageNumber="14">Stolonis tapiai</emphasis>
</taxonomicName>
, B—
<taxonomicName id="1288EE895C7E486A3A904C4FFAF7FAB4" box="[1152,1316,1372,1395]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="yasuni">
<emphasis id="E7FC49185C7E486A3A904C4FFAF7FAB4" box="[1152,1316,1372,1395]" italics="true" pageId="13" pageNumber="14">Stolonis yasuni</emphasis>
</taxonomicName>
. bc, bursa copulatrix. bs, bursal sclerites. co, common oviduct. gx1, gonocoxite­1. gx2, gonocoxite­ 2. lt, laterotergite IX. sg, spermathecal gland. sp, spermatheca.
</paragraph>
</caption>
<paragraph id="D537950A5C7E48693F284F07FBEBFE99" blockId="13.[264,1324,1556,1902]" lastBlockId="14.[264,1323,284,670]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Base of abdominal sternum II with row of dense, moderately deeply impressed punctures; base of sterna IVVII with dull band of coarse irregular microsculpture, more extensive on VII; sterna IVVI with one pair paramedial setae; in male one pair paramedial setae on VII, female with two pairs. Aedeagus (
<figureCitation id="4DB3898F5C7E486A3DD34F9FFBC0F961" box="[963,1043,1676,1702]" captionStart="FIGURE 3" captionStartId="9.[264,368,1716,1740]" captionTargetBox="[284,1304,346,1671]" captionTargetId="figure@9.[264,1323,329,1690]" captionTargetPageId="9" captionText="FIGURE 3. Aedeagus dorsal, left lateral, right lateral and ventral views of: A D— Stolonis tapiai, E H— Stolonis scortensis, I L— Stolonis spinosus, M P— Stolonis yasuni, Q T— Stolonis catenarius." httpUri="https://zenodo.org/record/169946/files/figure.png" pageId="13" pageNumber="14">Figs. 3</figureCitation>
MP) with median lobe simply rounded at tip, ostium dorsal and large; ventral surface well sclerotized; in repose endophallus with clearly defined scale field field near apex, left of midline and two spines located at 1/2 length of median lobe. Female reproductive tract (
<figureCitation id="4DB3898F5C7E486A3A034E17FB8FF8D9" box="[1043,1116,1796,1822]" captionStart="FIGURE 4" captionStartId="13.[264,367,1371,1395]" captionTargetBox="[293,1293,740,1348]" captionTargetId="figure@13.[274,1312,729,1359]" captionTargetPageId="13" captionText="FIGURE 4. Female reproductive tract, ventral views of: A— Stolonis tapiai, B— Stolonis yasuni. bc, bursa copulatrix. bs, bursal sclerites. co, common oviduct. gx 1, gonocoxite­ 1. gx 2, gonocoxite­ 2. lt, laterotergite IX. sg, spermathecal gland. sp, spermatheca." httpUri="https://zenodo.org/record/169947/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
B) with elongate, right dorsolateral bursal pouch, smaller expanded region subtending the broadly connected spermatheca and common oviduct; gland connected by a short duct to the subspermathecal pouch; laterotergites IX with scattered setae; gonocoxite­1 with 57 apical setae; gonocoxite 2 with 2 lateral ensiform setae, 1 apical nematiform setae.
</paragraph>
<paragraph id="D537950A5C7D48693F28487FFD5BFE11" blockId="14.[264,1323,284,670]" pageId="14" pageNumber="15">
Pygidial gland form simple, no extra lobes, chemical compounds produced: formic and acetic acids, 2­pentadecanone and C10:0, C11:0 hydrocarbons (as species number EC199802
<collectingCountry id="AD9FD59A5C7D48693F9648AFFE49FE11" box="[390,410,444,470]" name="Iceland" pageId="14" pageNumber="15">O</collectingCountry>
in
<bibRefCitation id="B119E8FB5C7D48693FD048AFFDAFFE11" author="Will" box="[448,636,444,470]" pageId="14" pageNumber="17" refString="Will, K. W., Attagalle, A. B. &amp; Herath, K. (2000) New defensive chemical data for Ground Beetles (Coleoptera: Carabidae): interpretations in a phylogenetic framework. Biological Journal of the Linnean Society, 71, 459 - 481." type="journal article" year="2000">
Will
<emphasis id="E7FC49185C7D48693FEA48AFFDFDFE12" box="[506,558,444,469]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 2000
</bibRefCitation>
).
</paragraph>
<paragraph id="D537950A5C7D48693F2848F0FE99FD59" blockId="14.[264,1323,284,670]" pageId="14" pageNumber="15">
<emphasis id="E7FC49185C7D48693F2848F0FCBBFE3A" bold="true" box="[312,872,483,509]" pageId="14" pageNumber="15">Natural history and collecting information</emphasis>
. Night active in very wet areas of primary forest. Taken treading vegetation in muddy areas and near fallen
<taxonomicName id="1288EE895C7D48693A634B1FFB67FDE2" box="[1139,1204,524,549]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="E7FC49185C7D48693A634B1FFB67FDE2" box="[1139,1204,524,549]" italics="true" pageId="14" pageNumber="15">Ficus</emphasis>
</taxonomicName>
and palm plants. Collected with
<taxonomicName id="1288EE895C7D48693C0F4B27FD6DFD8A" box="[543,702,564,589]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="catenarius">
<emphasis id="E7FC49185C7D48693C0F4B27FD6DFD8A" box="[543,702,564,589]" italics="true" pageId="14" pageNumber="15">S. catenarius</emphasis>
</taxonomicName>
and
<taxonomicName id="1288EE895C7D48693D104B27FC5BFD8A" box="[768,904,564,589]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="E7FC49185C7D48693D104B27FC5BFD8A" box="[768,904,564,589]" italics="true" pageId="14" pageNumber="15">S. spinosus</emphasis>
</taxonomicName>
. The first two larval instars were reared for this species in the laboratory. Six pigmented stemmata are present in the active larva.
</paragraph>
</subSubSection>
</treatment>
</document>