treatments-xml/data/03/A6/DA/03A6DA14FFF38D7B289D7A71FEC9F298.xml

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<mods:title id="4F47624E9E47B51DC040BF24D8F02A30">Revision of the Chinese endemic subgenus Tschitscherinea Berg, with descriptions of three new species (Coleoptera, Carabidae)</mods:title>
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<mods:namePart id="D0153C655654405F4261DB92A7519B4F">Yin, Wen-Qi</mods:namePart>
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<mods:namePart id="20DBEB4F60C51D86CDE657AB9C9D59FB">Shi, Hong-Liang</mods:namePart>
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<paragraph id="8BB06B02FFF38D7C289D7A71FB9EF20D" blockId="11.[151,1127,546,645]" box="[151,1127,546,573]" pageId="11" pageNumber="54">
<heading id="D0F8DC6EFFF38D7C289D7A71FB9EF20D" bold="true" box="[151,1127,546,573]" fontSize="11" level="1" pageId="11" pageNumber="54" reason="1">
<emphasis id="B97BB710FFF38D7C289D7A71FB9EF20D" bold="true" box="[151,1127,546,573]" pageId="11" pageNumber="54">
<taxonomicName id="4C0F1081FFF38D7C289D7A71FCE0F20D" ID-CoL="4Q3TW" authority="Jedlicka, 1934" authorityName="Jedlicka" authorityYear="1934" box="[151,793,546,573]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="54" phylum="Arthropoda" rank="species" species="sterbai" status="subgen. nov." subGenus="Tschitscherinea">
<emphasis id="B97BB710FFF38D7C289D7A71FED4F20C" bold="true" box="[151,301,546,573]" italics="true" pageId="11" pageNumber="54">Pterostichus</emphasis>
(
<emphasis id="B97BB710FFF38D7C29347A71FE05F20C" bold="true" box="[318,508,546,573]" italics="true" pageId="11" pageNumber="54">Tschitscherinea</emphasis>
) 
<emphasis id="B97BB710FFF38D7C2A077A71FDA7F20C" bold="true" box="[525,606,546,573]" italics="true" pageId="11" pageNumber="54">sterbai</emphasis>
<bibRefCitation id="EF9E16F3FFF38D7C2A6C7A71FCE0F20D" author="Jedlicka, A." box="[614,793,546,572]" pageId="11" pageNumber="54" pagination="13 - 19" refId="ref15381" refString="Jedlicka, A. (1934) Carabiden aus Ostasien. (5 Teil.). Casopis Ceskoslovenske Spolecnosti Entomologicke, 31, 13 - 19." type="journal article" year="1934">Jedlička, 1934</bibRefCitation>
</taxonomicName>
, 
<taxonomicNameLabel id="A2480A6BFFF38D7C2B2C7A70FC2DF20C" box="[806,980,547,573]" pageId="11" pageNumber="54" rank="subGenus">new subgenus</taxonomicNameLabel>
assignment
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3153889FFF38D7C289D7A14FE4FF250" box="[151,438,582,609]" pageId="11" pageNumber="54" type="description">
<paragraph id="8BB06B02FFF38D7C289D7A14FE4FF250" blockId="11.[151,1127,546,645]" box="[151,438,582,609]" pageId="11" pageNumber="54">
<figureCitation id="13347787FFF38D7C289D7A14FEC2F250" box="[151,315,582,609]" captionStart="FIGURES 13–18. 13–14" captionStartId="15.[152,270,1767,1791]" captionTargetBox="[158,1431,181,1736]" captionTargetId="figure-18@15.[151,1436,181,1743]" captionTargetPageId="15" captionText="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742732" httpUri="https://zenodo.org/record/12742732/files/figure.png" pageId="11" pageNumber="54">Figures 13–15</figureCitation>
, 
<figureCitation id="13347787FFF38D7C294C7A14FE9DF250" box="[326,356,583,609]" captionStart="FIGURES 25–30" captionStartId="17.[152,270,1537,1561]" captionTargetBox="[158,1430,183,1507]" captionTargetId="figure-18@17.[151,1436,181,1513]" captionTargetPageId="17" captionText="FIGURES 25–30. Endophallus of Pterostichus (Tschitscherinea) spp., left lateral view and right lateral view. (25) P.spissipedes sp. nov., Jianchuan, paratype. (26) P. jiuzhaiicus sp. nov., holotype. (27) P. mulensis Sciaky, Muli. (28) P. farkaci Sciaky, Hongshan, Zhongdian. (29) P. farkaci Sciaky, Tianchi, Zhongdian. (30) P. sterbai Jedlička, Pe-Yen-Tsin. Abbreviations: gp gonopore, la left apical lobe, ra right apical lobe, va ventral apical lobe. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742740" httpUri="https://zenodo.org/record/12742740/files/figure.png" pageId="11" pageNumber="54">30</figureCitation>
, 
<figureCitation id="13347787FFF38D7C297A7A14FE77F250" box="[368,398,583,609]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="11" pageNumber="54">43</figureCitation>
, 
<figureCitation id="13347787FFF38D7C29907A14FE4FF250" box="[410,438,583,609]" captionStart="FIGURES 48–53. 48–49" captionStartId="20.[152,270,1577,1601]" captionTargetBox="[151,1415,181,1541]" captionTargetId="figure-36@20.[151,1435,181,1553]" captionTargetPageId="20" captionText="FIGURES 48–53. 48–49. Male sternite VII of Pterostichus (Tschitscherinea) spp., (48) P. jiuzhaiicus sp. nov., Jiuzhaigou, paratype. (49) P. sterbai Jedlička, Pe-Yen-Tsin. Scale bars: 0.5 mm. 50–53. Female genitalia of Pterostichus (Tschitscherinea) mulensis Sciaky, Muli. (50) tergite VIII. (51) sternite VIII. (52) female internal reproductive tracts. (53) ventral view of gonocoxites. Abbreviations: g1 gonocoxite 1, g2 gonocoxite 2, es ensiform setae, ns nematiform setae, bc bursa copulatrix, co common oviduct, sc seminal canal, rc receptaculum, sg spermathecal gland, spc spermathecal canal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742750" httpUri="https://zenodo.org/record/12742750/files/figure.png" pageId="11" pageNumber="54">49</figureCitation>
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Chinese vernacular name: 
<emphasis id="B97BB710FFF38D7C29CB7A38FD90F2B4" bold="true" box="[449,617,619,645]" pageId="11" pageNumber="54">什ĸdzā±Ƥ</emphasis>
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<paragraph id="8BB06B02FFF38D7C289D7AFCFC5BF2D7" blockId="11.[151,1436,687,742]" pageId="11" pageNumber="54">
<bibRefCitation id="EF9E16F3FFF38D7C289D7AFCFEB1F2F6" author="Jedlicka, A." box="[151,328,687,711]" pageId="11" pageNumber="54" pagination="13 - 19" refId="ref15381" refString="Jedlicka, A. (1934) Carabiden aus Ostasien. (5 Teil.). Casopis Ceskoslovenske Spolecnosti Entomologicke, 31, 13 - 19." type="journal article" year="1934">Jedlička 1934: 18</bibRefCitation>
(Original: 
<taxonomicName id="4C0F1081FFF38D7C29BD7AFCFDCCF2F6" box="[439,565,687,711]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B97BB710FFF38D7C29BD7AFCFDCCF2F6" box="[439,565,687,711]" italics="true" pageId="11" pageNumber="54">Pterostichus</emphasis>
</taxonomicName>
: subgenus incertae; 
<typeStatus id="54B4D5A0FFF38D7C2B087AFCFCA2F2F6" box="[770,859,687,711]" pageId="11" pageNumber="54" type="holotype">holotype</typeStatus>
in NMPC; type locality: 
<collectingRegion id="49CBA5E0FFF38D7C2C5C7AE3FB5EF2F6" box="[1110,1191,688,711]" country="China" name="Yunnan" pageId="11" pageNumber="54">Yunnan</collectingRegion>
: Pei-Yen-Tsin); 
<bibRefCitation id="EF9E16F3FFF38D7C2D437AFCFED4F2D7" author="Jedlicka, A." pageId="11" pageNumber="54" pagination="177 - 346" refId="ref15409" refString="Jedlicka, A. (1962) Monographie des Tribus Pterostichini aus Osatasien (Pterostichini, Trigonotomi, Myadi) (Coleoptera- Carabidae). Abhandlungen und Berichte aus dem Staatlichen Museum fur Tierkunde in Dresden, 26, 177 - 346." type="journal article" year="1962">Jedlička 1962: 303</bibRefCitation>
(subgenus incertae); Sciaky 1997: 163 (subgenus 
<taxonomicName id="4C0F1081FFF38D7C2B217A9DFC6AF2D7" authorityName="Sciaky" authorityYear="1997" box="[811,915,718,742]" class="Insecta" family="Carabidae" genus="Jedlickaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B97BB710FFF38D7C2B217A9DFC6AF2D7" box="[811,915,718,742]" italics="true" pageId="11" pageNumber="54">Jedlickaia</emphasis>
</taxonomicName>
).
</paragraph>
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<paragraph id="8BB06B02FFF38D7C289D7B47FE8FF3D2" blockId="11.[151,1437,788,1931]" pageId="11" pageNumber="54">
<emphasis id="B97BB710FFF38D7C289D7B47FE79F31F" bold="true" box="[151,384,788,814]" pageId="11" pageNumber="54">Material examined.</emphasis>
<materialsCitation id="3B67615FFFF38D7C298D7B47FB61F363" collectionCode="NMPC" determinerName="Det. Ing. Jedlicka" location="Yunnan" pageId="11" pageNumber="54" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54B4D5A0FFF38D7C298D7B47FE16F31F" box="[391,495,788,814]" pageId="11" pageNumber="54" type="holotype">Holotype</typeStatus>
of 
<taxonomicName id="4C0F1081FFF38D7C2A197B46FD1BF31F" authority="Jedlicka" authorityName="Jedlicka" box="[531,738,788,814]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="54" phylum="Arthropoda" rank="species" species="sterbai">
<emphasis id="B97BB710FFF38D7C2A197B46FD85F31F" box="[531,636,788,814]" italics="true" pageId="11" pageNumber="54">P. sterbai</emphasis>
Jedlička
</taxonomicName>
, female (
<collectionCode id="ED1EF3C7FFF38D7C2B4C7B47FC63F31F" box="[838,922,788,814]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="11" pageNumber="54">NMPC</collectionCode>
, 
<figureCitation id="13347787FFF38D7C2BAF7B47FC0EF31F" box="[933,1015,788,814]" captionStart="FIGURES 13–18. 13–14" captionStartId="15.[152,270,1767,1791]" captionTargetBox="[158,1431,181,1736]" captionTargetId="figure-18@15.[151,1436,181,1743]" captionTargetPageId="15" captionText="FIGURES 13–18. 13–14. Habitus and labels of P. sterbai Jedlička: (13) holotype. (14) a male paratype. Scale bars: 2 mm. 15–18. Male genitalia of Pterostichus (Tschitscherinea) spp., left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (15) P. sterbai Jedlička, Pe-Yen-Tsin. (16) P. spissipedes sp. nov., holotype. (17) P. spissipedes sp. nov., Deqin, paratype. (18) P. spectabilis sp. nov., holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742732" httpUri="https://zenodo.org/record/12742732/files/figure.png" pageId="11" pageNumber="54">Fig. 13</figureCitation>
): “Pe Yen Tsin, 
<collectingRegion id="49CBA5E0FFF38D7C2CA27B47FAFDF31F" box="[1192,1284,788,814]" country="China" name="Yunnan" pageId="11" pageNumber="54">Yunnan</collectingRegion>
”; “Type” [red label], “Mus. Nat. Pragae, Inv. 
<emphasis id="B97BB710FFF38D7C29F87B6BFDC1F363" box="[498,568,824,850]" italics="true" pageId="11" pageNumber="54">22404</emphasis>
”, 
<taxonomicName id="4C0F1081FFF38D7C2A587B6BFD55F363" box="[594,684,824,850]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="54" phylum="Arthropoda" rank="species" species="sterbai">
“ 
<emphasis id="B97BB710FFF38D7C2A547B6BFD55F363" box="[606,684,824,850]" italics="true" pageId="11" pageNumber="54">sterbai</emphasis>
</taxonomicName>
, type, sp.n., 
<determinerName id="7E6C87C6FFF38D7C2B307B6AFBF8F363" box="[826,1025,824,851]" pageId="11" pageNumber="54">Det. Ing. Jedlička</determinerName>
” [pink label]
</materialsCitation>
. 
<typeStatus id="54B4D5A0FFF38D7C2CA87B6AFAF6F362" box="[1186,1295,825,851]" pageId="11" pageNumber="54" type="paratype">Paratypes</typeStatus>
of 
<taxonomicName id="4C0F1081FFF38D7C2D397B6AFF0EF347" authority="Jedlicka" authorityName="Jedlicka" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="54" phylum="Arthropoda" rank="species" species="sterbai">
<emphasis id="B97BB710FFF38D7C2D397B6AFA65F363" box="[1331,1436,824,850]" italics="true" pageId="11" pageNumber="54">P. sterbai</emphasis>
Jedlička
</taxonomicName>
: 
<materialsCitation id="3B67615FFFF38D7C29087B0FFD83F3AB" collectionCode="NMPC" determinerName="Ing. Jedlicka" location="Yunnan" pageId="11" pageNumber="54" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D09A08BFFF38D7C29087B0FFE97F347" box="[258,366,860,886]" count="2" pageId="11" pageNumber="54" type="female">2 females</specimenCount>
(
<collectionCode id="ED1EF3C7FFF38D7C29777B0FFE2DF347" box="[381,468,860,886]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="11" pageNumber="54">NMPC</collectionCode>
): “Pe Yen Tsin, 
<collectingRegion id="49CBA5E0FFF38D7C2A8E7B0FFD26F347" box="[644,735,860,886]" country="China" name="Yunnan" pageId="11" pageNumber="54">Yunnan</collectingRegion>
”, “Cotype” [red label], “Mus. Nat. Pragae, Inv. 
<emphasis id="B97BB710FFF38D7C2CFF7B0FFA70F347" box="[1269,1417,860,886]" italics="true" pageId="11" pageNumber="54">22405/22406</emphasis>
”, 
<taxonomicName id="4C0F1081FFF38D7C289D7BD3FF0BF3AB" box="[151,242,896,922]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="54" phylum="Arthropoda" rank="species" species="sterbai">
“ 
<emphasis id="B97BB710FFF38D7C28AE7BD3FF0BF3AB" box="[164,242,896,922]" italics="true" pageId="11" pageNumber="54">sterbai</emphasis>
</taxonomicName>
, 
<emphasis id="B97BB710FFF38D7C28F47BD2FEE3F3AB" box="[254,282,897,922]" italics="true" pageId="11" pageNumber="54">m.</emphasis>
, det. 
<determinerName id="7E6C87C6FFF38D7C295F7BD2FE1CF3AB" box="[341,485,896,923]" pageId="11" pageNumber="54">Ing. Jedlička</determinerName>
” [pink label]
</materialsCitation>
; 
<materialsCitation id="3B67615FFFF38D7C2A8E7BD3FE10F38F" collectionCode="NMPC" determinerName="Ing. Jedlicka" location="Yunnan" pageId="11" pageNumber="54" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D09A08BFFF38D7C2A8E7BD3FD34F3AB" box="[644,717,896,922]" count="1" pageId="11" pageNumber="54" type="male">1 male</specimenCount>
(
<collectionCode id="ED1EF3C7FFF38D7C2AD67BD3FCCAF3AB" box="[732,819,896,922]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="11" pageNumber="54">NMPC</collectionCode>
): “Pe Yen Tsin, 
<collectingRegion id="49CBA5E0FFF38D7C2BD77BD3FBC1F3AB" box="[989,1080,896,922]" country="China" name="Yunnan" pageId="11" pageNumber="54">Yunnan</collectingRegion>
”, “Cotype” [red label], 
<taxonomicName id="4C0F1081FFF38D7C2D317BD3FA6CF3AB" box="[1339,1429,896,922]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="54" phylum="Arthropoda" rank="species" species="sterbai">
“ 
<emphasis id="B97BB710FFF38D7C2D4D7BD3FA6CF3AB" box="[1351,1429,896,922]" italics="true" pageId="11" pageNumber="54">sterbai</emphasis>
</taxonomicName>
, det. 
<determinerName id="7E6C87C6FFF38D7C28C27BF6FEA3F38F" box="[200,346,932,959]" pageId="11" pageNumber="54">Ing. Jedlička</determinerName>
”[pink label]
</materialsCitation>
; 
<materialsCitation id="3B67615FFFF38D7C29FF7BF6FE8BF3D2" collectionCode="MNHN" collectorName="Coll. R. Oberthur" location="Yunnan" pageId="11" pageNumber="54" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
Non-type material: 
<specimenCount id="9D09A08BFFF38D7C2ADA7BF7FCE5F38F" box="[720,796,932,958]" count="1" pageId="11" pageNumber="54" type="male">1 male</specimenCount>
(
<collectionCode id="ED1EF3C7FFF38D7C2B217BF7FC71F38F" box="[811,904,932,958]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="54">MNHN</collectionCode>
): “Pe Yen Tsin, 
<collectingRegion id="49CBA5E0FFF38D7C2C337BF7FB6DF38F" box="[1081,1172,932,958]" country="China" name="Yunnan" pageId="11" pageNumber="54">Yunnan</collectingRegion>
”, “Muséum Paris, 
<collectorName id="26FA0ED4FFF38D7C2D6F7BF7FEE6F3D3" pageId="11" pageNumber="54">Coll. R. Oberthür</collectorName>
, 1952”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3153889FFF38D7C28CD7BBFFDFCF6E7" pageId="11" pageNumber="54" type="diagnosis">
<paragraph id="8BB06B02FFF38D7C28CD7BBFFE5EF4A7" blockId="11.[151,1437,788,1931]" pageId="11" pageNumber="54">
<emphasis id="B97BB710FFF38D7C28CD7BBFFEBBF437" bold="true" box="[199,322,1004,1030]" pageId="11" pageNumber="54">Diagnosis.</emphasis>
This species is distinguishable among the subgenus by the combination of following characters: maxim width of head at outer margin of tempora; pronotum ovate, lateral margins distinctly arched near middle; eyes small, tempora strongly swollen behind eyes; elytra with isodiametric microsculpture; parascutellar pore absent; interval 3 with two setigerous pores. This species can be readily distinguished among this subgenus by its special pronotum shape.
</paragraph>
<paragraph id="8BB06B02FFF38D7C28CD7CF3FDFCF6E7" blockId="11.[151,1437,788,1931]" pageId="11" pageNumber="54">
<emphasis id="B97BB710FFF38D7C28CD7CF3FDEFF48B" bold="true" box="[199,534,1184,1210]" pageId="11" pageNumber="54">Supplementary description.</emphasis>
BL = 
<quantity id="4CF7C6E7FFF38D7C2A637CF3FCF6F48A" box="[617,783,1184,1211]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.16" metricValueMin="1.14" pageId="11" pageNumber="54" unit="mm" value="11.5" valueMax="11.6" valueMin="11.4">11.4–11.6 mm</quantity>
, BW = 4.0– 
<quantity id="4CF7C6E7FFF38D7C2BA87CF3FC07F48A" box="[930,1022,1184,1211]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="11" pageNumber="54" unit="mm" value="4.3">4.3 mm</quantity>
; eyes quite small, tempora strongly swollen behind eyes, longer than eyes, eyL/tpL = 0.63–0.71, maxim width of head at outer margin of tempora, HW/PW = 0.66–0.68. Pronotum ovate, PW/PL = 1.18–1.23, PBW/PW = 0.66–0.69, widest near anterior third; anterior margin slightly wider than posterior margin, PAW/PBW = 1.06–1.10; lateral margins distinctly arched near middle, gradually curved without sinuation to posterior angles; posterior angles rounded-obtuse, without denticle. Basal foveae shallow, distinctly punctate, inner and outer grooves faintly defined, fused to each at base. Elytra with isodiametric microsculpture; parascutellar pore present; humerus with small tooth, sharply projected out; interval 3 with two setigerous pores, both adjacent to stria 2. Male sternite VII with a round depression at middle (
<figureCitation id="13347787FFF38D7C2D317DCFFA76F587" box="[1339,1423,1436,1462]" captionStart="FIGURES 48–53. 48–49" captionStartId="20.[152,270,1577,1601]" captionTargetBox="[151,1415,181,1541]" captionTargetId="figure-36@20.[151,1435,181,1553]" captionTargetPageId="20" captionText="FIGURES 48–53. 48–49. Male sternite VII of Pterostichus (Tschitscherinea) spp., (48) P. jiuzhaiicus sp. nov., Jiuzhaigou, paratype. (49) P. sterbai Jedlička, Pe-Yen-Tsin. Scale bars: 0.5 mm. 50–53. Female genitalia of Pterostichus (Tschitscherinea) mulensis Sciaky, Muli. (50) tergite VIII. (51) sternite VIII. (52) female internal reproductive tracts. (53) ventral view of gonocoxites. Abbreviations: g1 gonocoxite 1, g2 gonocoxite 2, es ensiform setae, ns nematiform setae, bc bursa copulatrix, co common oviduct, sc seminal canal, rc receptaculum, sg spermathecal gland, spc spermathecal canal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742750" httpUri="https://zenodo.org/record/12742750/files/figure.png" pageId="11" pageNumber="54">Fig. 49</figureCitation>
); meso- and metafemora distinctly thickened in males; metatarsus in a stout form in males (mt1 length/width = 2.1, 
<figureCitation id="13347787FFF38D7C289D7DB7FF15F5CF" box="[151,236,1508,1534]" captionStart="FIGURES 35–47. 35–43" captionStartId="19.[152,270,1487,1511]" captionTargetBox="[158,1436,181,1457]" captionTargetId="figure-18@19.[151,1436,181,1463]" captionTargetPageId="19" captionText="FIGURES 35–47. 35–43. Metatarsi of Pterostichus (Tschitscherinea) spp. (35) P. spissipedes sp. nov., male. (36) P. spissipedes sp. nov., female. (37) P. spectabilis sp. nov., male. (38) P. spectabilis sp. nov., female. (39) P. mulensis Sciaky, male. (40) P. mulensis Sciaky, female. (41) P. farkaci Sciaky, male. (42) P. krali Sciaky, male. (43) P. sterbai Jedlička, male. Scale bar: 1 mm. 44–47. Elytral microsculpture of Pterostichus (Tschitscherinea) spp. (44) isodiametric, P. specillatus sp. nov., holotype. (45) transverse-isodiametric, P. mulensis Sciaky, a male from Muli. (46) isodiametric, P. farkaci Sciaky, a male from Tianchi, Zhongdian. (47) transverse-linear, P. krali Sciaky, a male from Haba Snow Mountain, Zhongdian. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742746" httpUri="https://zenodo.org/record/12742746/files/figure.png" pageId="11" pageNumber="54">Fig. 43</figureCitation>
), rather slender in females, segments 1–2 with outer-lateral ridge. Median lobe of aedeagus stout (length about 
<quantity id="4CF7C6E7FFF38D7C28D07E5BFEB9F613" box="[218,320,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="54" unit="mm" value="1.25">1.25 mm</quantity>
), apex distinctly curved ventrally in lateral view; apical lamella short and wide, apex rounded, slightly twisted to right; right paramere nearly straight, apical portion gradually attenuate to apex, apex sharply pointed. Endophallus with three lobes near gonopore all without sclerotized apex: left apical lobe (la) nearly triangular with sharp apex, surface finely scaled with slightly sclerotized apex; right apical lobe (ra) oblate, about equal sized as la, with rounded apex; ventral apical lobe (va) papillary, slightly smaller than other lobes, located on the dorsal apical surface of endophallus. (
<figureCitation id="13347787FFF38D7C29A17EEFFE05F6E7" box="[427,508,1724,1750]" captionStart="FIGURES 25–30" captionStartId="17.[152,270,1537,1561]" captionTargetBox="[158,1430,183,1507]" captionTargetId="figure-18@17.[151,1436,181,1513]" captionTargetPageId="17" captionText="FIGURES 25–30. Endophallus of Pterostichus (Tschitscherinea) spp., left lateral view and right lateral view. (25) P.spissipedes sp. nov., Jianchuan, paratype. (26) P. jiuzhaiicus sp. nov., holotype. (27) P. mulensis Sciaky, Muli. (28) P. farkaci Sciaky, Hongshan, Zhongdian. (29) P. farkaci Sciaky, Tianchi, Zhongdian. (30) P. sterbai Jedlička, Pe-Yen-Tsin. Abbreviations: gp gonopore, la left apical lobe, ra right apical lobe, va ventral apical lobe. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742740" httpUri="https://zenodo.org/record/12742740/files/figure.png" pageId="11" pageNumber="54">Fig. 30</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3153889FFF38D7C28CD7EB3FDFEF72F" pageId="11" pageNumber="54" type="discussion">
<paragraph id="8BB06B02FFF38D7C28CD7EB3FDFEF72F" blockId="11.[151,1437,788,1931]" pageId="11" pageNumber="54">
<emphasis id="B97BB710FFF38D7C28CD7EB3FECDF6CB" bold="true" box="[199,308,1760,1786]" pageId="11" pageNumber="54">Remarks</emphasis>
. This species is newly assigned to the subgenus 
<taxonomicName id="4C0F1081FFF38D7C2B587EB3FBF8F6CB" box="[850,1025,1760,1786]" pageId="11" pageNumber="54" rank="subGenus" subGenus="Tschitscherinea">
<emphasis id="B97BB710FFF38D7C2B587EB3FBF8F6CB" box="[850,1025,1760,1786]" italics="true" pageId="11" pageNumber="54">Tschitscherinea</emphasis>
</taxonomicName>
for the reasons presented above (see discussions under the subgenus).
</paragraph>
</subSubSection>
<subSubSection id="C3153889FFF38D7B28CD7F7BFEC9F298" lastPageId="12" lastPageNumber="55" pageId="11" pageNumber="54" type="distribution">
<paragraph id="8BB06B02FFF38D7C28CD7F7BFDA3F7BB" blockId="11.[151,1437,788,1931]" pageId="11" pageNumber="54">
<emphasis id="B97BB710FFF38D7C28CD7F7BFE98F773" bold="true" box="[199,353,1832,1858]" pageId="11" pageNumber="54">Distribution.</emphasis>
This species was only known from its 
<typeStatus id="54B4D5A0FFF38D7C2B137F7AFCB0F772" box="[793,841,1833,1859]" pageId="11" pageNumber="54">type</typeStatus>
locality, Pe Yen Tsin (= Shiyang Township 
<geoCoordinate id="EE3B0DC5FFF38D7C2D3D7F7BFA61F772" box="[1335,1432,1832,1859]" degrees="25.85" direction="north" orientation="latitude" pageId="11" pageNumber="54" precision="555" value="25.85">N25.85°</geoCoordinate>
, 
<geoCoordinate id="EE3B0DC5FFF38D7C289D7F1FFEFAF757" box="[151,259,1868,1894]" degrees="101.10" direction="east" orientation="longitude" pageId="11" pageNumber="54" precision="555" value="101.1">E101.10°</geoCoordinate>
, Dayao county, Chuxiong Yi Autonomous Prefecture, today), 
<collectingRegion id="49CBA5E0FFF38D7C2BD47F1EFB5CF757" box="[990,1189,1868,1895]" country="China" name="Yunnan" pageId="11" pageNumber="54">Yunnan province</collectingRegion>
. There is no modern specimen of this species. (
<figureCitation id="13347787FFF38D7C29B77F23FDF6F7BA" box="[445,527,1904,1931]" captionStart="FIGURES 54–57. 54" captionStartId="21.[152,270,1525,1549]" captionTargetBox="[151,1435,181,1494]" captionTargetId="figure-18@21.[151,1435,181,1501]" captionTargetPageId="21" captionText="FIGURES 54–57. 54. Confirmed distribution of the Pterostichus (Tschitscherinea) spp. in China: Red P. spissipedes sp. nov. Blue P. spectabilis sp. nov. Black P. jiuzhaiicus sp. nov. Pink P. filum (Tschitschérine). Purple P. mulensis Sciaky. Green P. fakarci Sciaky. Yellow P. krali Sciaky. Cyan P. sterbai Jedlička. 55–57. Endophallus of Pterostichus spp., right lateral view and left lateral view: (55) Pterostichus (Cryobius) jaechi Krischenhofer, Changbaishan, Jilin. (56) Pterostichus (Morphohaptoderus) janatai Sciaky &amp; Wrase, Pingheliang, Shaanxi. (57) Pterostichus (Morphohaptoderus) dundai Sciaky, Houzhenzi, Shaanxi. Scale bars = 0.5 mm, a (55), b (56–57)." figureDoi="http://doi.org/10.5281/zenodo.12742754" httpUri="https://zenodo.org/record/12742754/files/figure.png" pageId="11" pageNumber="54">Fig. 54</figureCitation>
, cyan)
</paragraph>
<paragraph id="8BB06B02FFF48D7B289D78C4FEC9F298" blockId="12.[151,1437,151,681]" pageId="12" pageNumber="55">
The mysterious species 
<taxonomicName id="4C0F1081FFF48D7B29A278C4FCBFF080" authority="Sciaky &amp; Wrase, 1997" authorityName="Sciaky &amp; Wrase" authorityYear="1997" box="[424,838,151,177]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B97BB710FFF48D7B29A278C4FDC4F080" box="[424,573,151,177]" italics="true" pageId="12" pageNumber="55">P. parvicollis</emphasis>
<bibRefCitation id="EF9E16F3FFF48D7B2A4C78C4FCBFF080" author="Sciaky, R. &amp; Wrase, D. W." box="[582,838,151,177]" pageId="12" pageNumber="55" pagination="1087 - 1139" refId="ref15565" refString="Sciaky, R. &amp; Wrase, D. W. (1997) Twenty-nine new taxa of Pterostichinae from Shaanxi (Coleoptera, Carabidae). Linzer Biologische Beitrage, 29, 1087 - 1139." type="journal article" year="1997">Sciaky &amp; Wrase, 1997</bibRefCitation>
</taxonomicName>
was only known by the female 
<typeStatus id="54B4D5A0FFF48D7B2CBC78C4FAE1F080" box="[1206,1304,151,177]" pageId="12" pageNumber="55" type="holotype">holotype</typeStatus>
previously. It was placed in the subgenus 
<taxonomicName id="4C0F1081FFF48D7B29F678E8FD2DF0E4" box="[508,724,187,213]" pageId="12" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
<emphasis id="B97BB710FFF48D7B29F678E8FD2DF0E4" box="[508,724,187,213]" italics="true" pageId="12" pageNumber="55">Morphohaptoderus</emphasis>
</taxonomicName>
for external similarities to 
<taxonomicName id="4C0F1081FFF48D7B2C1C78E8FA61F0E4" authority="Sciaky &amp; Wrase, 1997" authorityName="Sciaky &amp; Wrase" authorityYear="1997" box="[1046,1432,187,213]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="geberti">
<emphasis id="B97BB710FFF48D7B2C1C78E8FB7DF0E4" box="[1046,1156,187,213]" italics="true" pageId="12" pageNumber="55">P. geberti</emphasis>
<bibRefCitation id="EF9E16F3FFF48D7B2C8578E8FA61F0E4" author="Sciaky, R. &amp; Wrase, D. W." box="[1167,1432,187,213]" pageId="12" pageNumber="55" pagination="1087 - 1139" refId="ref15565" refString="Sciaky, R. &amp; Wrase, D. W. (1997) Twenty-nine new taxa of Pterostichinae from Shaanxi (Coleoptera, Carabidae). Linzer Biologische Beitrage, 29, 1087 - 1139." type="journal article" year="1997">Sciaky &amp; Wrase, 1997</bibRefCitation>
</taxonomicName>
. However, this species has some external characteristics that aligned with the subgenus 
<taxonomicName id="4C0F1081FFF48D7B2C7E788CFADAF0C8" box="[1140,1315,223,249]" pageId="12" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
<emphasis id="B97BB710FFF48D7B2C7E788CFADAF0C8" box="[1140,1315,223,249]" italics="true" pageId="12" pageNumber="55">Tschitscherinea</emphasis>
</taxonomicName>
, including the somewhat parallel-sided body shape, thickened male metatarsus, and prominent elytral humeral tooth. In this study, we report the male of 
<taxonomicName id="4C0F1081FFF48D7B29E77974FD7CF170" box="[493,645,295,321]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B97BB710FFF48D7B29E77974FD7CF170" box="[493,645,295,321]" italics="true" pageId="12" pageNumber="55">P. parvicollis</emphasis>
</taxonomicName>
. The morphological study of its male genitalia (
<figureCitation id="13347787FFF48D7B2CB47974FAEAF170" box="[1214,1299,295,321]" captionStart="FIGURES 31–34" captionStartId="18.[152,270,1908,1932]" captionTargetBox="[184,1403,188,1878]" captionTargetId="figure-36@18.[177,1409,181,1885]" captionTargetPageId="18" captionText="FIGURES 31–34. habitus and male genitalia of P. parvicollis Sciaky &amp; Wrase. (31) habitus of holotype. (32–34) a male specimen from Pingheliang, Shaanxi. (32) habitus. (33) male genitalia, left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (34) endophallus. Abbreviations: gp gonopore, gpp gonopore piece, la left apical lobe, ra right apical lobe. Scale bars: 2 mm (31–32), 0.5 mm (33–34)." figureDoi="http://doi.org/10.5281/zenodo.12742744" httpUri="https://zenodo.org/record/12742744/files/figure.png" pageId="12" pageNumber="55">Fig. 33</figureCitation>
) suggests a closer relationship to subgenus 
<taxonomicName id="4C0F1081FFF48D7B29F07918FD2BF154" box="[506,722,331,357]" pageId="12" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
<emphasis id="B97BB710FFF48D7B29F07918FD2BF154" box="[506,722,331,357]" italics="true" pageId="12" pageNumber="55">Morphohaptoderus</emphasis>
</taxonomicName>
: (1) the right paramere of 
<taxonomicName id="4C0F1081FFF48D7B2BF77918FB68F154" box="[1021,1169,331,357]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B97BB710FFF48D7B2BF77918FB68F154" box="[1021,1169,331,357]" italics="true" pageId="12" pageNumber="55">P. parvicollis</emphasis>
</taxonomicName>
is very short, typical as nearly all members of 
<taxonomicName id="4C0F1081FFF48D7B299C793CFD97F1B8" box="[406,622,367,393]" pageId="12" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
<emphasis id="B97BB710FFF48D7B299C793CFD97F1B8" box="[406,622,367,393]" italics="true" pageId="12" pageNumber="55">Morphohaptoderus</emphasis>
</taxonomicName>
; (2) the apical lamella of aedeagus is somewhat similar to other species of 
<taxonomicName id="4C0F1081FFF48D7B28B279C0FE9EF19C" box="[184,359,403,429]" pageId="12" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
<emphasis id="B97BB710FFF48D7B28B279C0FE9EF19C" box="[184,359,403,429]" italics="true" pageId="12" pageNumber="55">Tschitscherinea</emphasis>
</taxonomicName>
, but slightly longer than all known species of the subgenus; (3) endophallus has a sclerotized piece near gonopore (
<figureCitation id="13347787FFF48D7B298479E4FE1BF1E0" box="[398,482,439,465]" captionStart="FIGURES 31–34" captionStartId="18.[152,270,1908,1932]" captionTargetBox="[184,1403,188,1878]" captionTargetId="figure-36@18.[177,1409,181,1885]" captionTargetPageId="18" captionText="FIGURES 31–34. habitus and male genitalia of P. parvicollis Sciaky &amp; Wrase. (31) habitus of holotype. (32–34) a male specimen from Pingheliang, Shaanxi. (32) habitus. (33) male genitalia, left lateral view, dorsal view of median lobe of aedeagus, and right paramere. (34) endophallus. Abbreviations: gp gonopore, gpp gonopore piece, la left apical lobe, ra right apical lobe. Scale bars: 2 mm (31–32), 0.5 mm (33–34)." figureDoi="http://doi.org/10.5281/zenodo.12742744" httpUri="https://zenodo.org/record/12742744/files/figure.png" pageId="12" pageNumber="55">Fig. 34</figureCitation>
), and such structure is absent in all known species of 
<taxonomicName id="4C0F1081FFF48D7B2C4079E4FB00F1E0" box="[1098,1273,439,465]" pageId="12" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
<emphasis id="B97BB710FFF48D7B2C4079E4FB00F1E0" box="[1098,1273,439,465]" italics="true" pageId="12" pageNumber="55">Tschitscherinea</emphasis>
</taxonomicName>
but present in some 
<taxonomicName id="4C0F1081FFF48D7B28D37988FE48F1C4" box="[217,433,475,501]" pageId="12" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
<emphasis id="B97BB710FFF48D7B28D37988FE48F1C4" box="[217,433,475,501]" italics="true" pageId="12" pageNumber="55">Morphohaptoderus</emphasis>
</taxonomicName>
. Moreover, 
<taxonomicName id="4C0F1081FFF48D7B2A317988FD37F1C4" box="[571,718,475,501]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B97BB710FFF48D7B2A317988FD37F1C4" box="[571,718,475,501]" italics="true" pageId="12" pageNumber="55">P. parvicollis</emphasis>
</taxonomicName>
is also different from all other species of 
<taxonomicName id="4C0F1081FFF48D7B2CAB7988FAA9F1C4" box="[1185,1360,475,501]" pageId="12" pageNumber="55" rank="subGenus" subGenus="Tschitscherinea">
<emphasis id="B97BB710FFF48D7B2CAB7988FAA9F1C4" box="[1185,1360,475,501]" italics="true" pageId="12" pageNumber="55">Tschitscherinea</emphasis>
</taxonomicName>
by the submentum with only one seta on each side, angular base of stria 1 almost absent, and the fifth tarsomere setose ventrally. Based on the above evidence, it is better to keep 
<taxonomicName id="4C0F1081FFF48D7B2B3C7A70FC32F20C" box="[822,971,547,573]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B97BB710FFF48D7B2B3C7A70FC32F20C" box="[822,971,547,573]" italics="true" pageId="12" pageNumber="55">P. parvicollis</emphasis>
</taxonomicName>
within the subgenus 
<taxonomicName id="4C0F1081FFF48D7B2CB77A70FA6CF20C" box="[1213,1429,547,573]" pageId="12" pageNumber="55" rank="subGenus" subGenus="Morphohaptoderus">
<emphasis id="B97BB710FFF48D7B2CB77A70FA6CF20C" box="[1213,1429,547,573]" italics="true" pageId="12" pageNumber="55">Morphohaptoderus</emphasis>
</taxonomicName>
. However, its systematic position still needs future investigation. To facilitate a better morphological recognition and future systematic study for 
<taxonomicName id="4C0F1081FFF48D7B29C07A38FDA4F2B4" box="[458,605,619,645]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="55" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B97BB710FFF48D7B29C07A38FDA4F2B4" box="[458,605,619,645]" italics="true" pageId="12" pageNumber="55">P. parvicollis</emphasis>
</taxonomicName>
, descriptions and illustrations for males of this species are provided in the present work.
</paragraph>
</subSubSection>
</treatment>
</document>