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128 lines
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<document id="54AE299151A1A6623A1AE7C9836B820D" ID-DOI="10.11646/zootaxa.4048.4.1" ID-GBIF-Dataset="bbb51c92-e184-489d-aba8-18d1dc9a25cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="234023" ID-ZooBank="71E2B8DA-825E-429F-BAA0-F582702B4A80" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461348015003" checkinUser="plazi" docAuthor="López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán" docDate="2015" docId="990DA529FF90FF89FF4643D615A4FCAD" docLanguage="en" docName="zt04048p492.pdf" docOrigin="Zootaxa 4048 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hylobothynus Ohaus 1910" docType="treatment" docVersion="11" lastPageNumber="465" masterDocId="6534DD51FF9EFF87FFD1425F1659FFE6" masterDocTitle="The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution" masterLastPageNumber="492" masterPageNumber="451" pageNumber="465" updateTime="1698621022009" updateUser="plazi">
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<mods:title id="8B353841A0FB0F6C348BC56321C6DADD">The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution</mods:title>
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<mods:namePart id="B6B942A651B895FC29C5EE72C9CD2781">López-García, Margarita M.</mods:namePart>
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<mods:name id="A80E3FE35893C28DA57F732D0A6C1880" type="personal">
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<mods:namePart id="58D4D68E84B139594A1EFEC3E14FFA17">Gasca-Álvarez, Héctor J.</mods:namePart>
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<mods:name id="FD53DD76BD809C0D75394ADA106ECC30" type="personal">
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<mods:namePart id="60EAA9B06256B312DEE6E37EC2A54F0D">Amat-García, Germán</mods:namePart>
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<mods:typeOfResource id="F48DCC37D202467F96DEE2D8DA79F328">text</mods:typeOfResource>
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<mods:title id="892D7E6903C1DFDAF9C1D91CAA856DEF">Zootaxa</mods:title>
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<mods:part id="B696387A66A816B5CE904FC8A8C6649D">
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<mods:date id="AC9AA4993754F2DD484741C06B26A876">2015</mods:date>
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<mods:detail id="7466207CCBA9D6D8DD3FEE7AED25E02B" type="volume">
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<mods:number id="644FB11EB16E90B972EF2CEFE8575BB4">4048</mods:number>
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<mods:number id="C195A5039B8103D830830E8D74B3E10E">4</mods:number>
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<mods:identifier id="42982136B5AFAF30BDEED9E3375C5F7A" type="DOI">10.11646/zootaxa.4048.4.1</mods:identifier>
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<mods:identifier id="DE11EC8BE198269F88B0A15ED7001D56" type="ISSN">1175-5326</mods:identifier>
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<treatment id="990DA529FF90FF89FF4643D615A4FCAD" ID-DOI="http://doi.org/10.5281/zenodo.5699356" ID-GBIF-Taxon="119645653" ID-Zenodo-Dep="5699356" LSID="urn:lsid:plazi:treatment:990DA529FF90FF89FF4643D615A4FCAD" httpUri="http://treatment.plazi.org/id/990DA529FF90FF89FF4643D615A4FCAD" lastPageNumber="465" pageId="14" pageNumber="465">
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<subSubSection id="59BE47B4FF90FF89FF4643D61777FE22" pageId="14" pageNumber="465" type="nomenclature">
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<paragraph id="111B143FFF90FF89FF4643D617BBFE45" blockId="14.[151,482,393,453]" box="[151,482,393,419]" pageId="14" pageNumber="465">
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<heading id="4A53A353FF90FF89FF4643D617BBFE45" bold="true" box="[151,482,393,419]" fontSize="11" level="1" pageId="14" pageNumber="465" reason="1">
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<taxonomicName id="D6A46FBCFF90FF89FF4643D617BBFE45" authority="Ohaus, 1910" authorityName="Ohaus" authorityYear="1910" box="[151,482,393,419]" class="Insecta" family="Dynastidae" genus="Hylobothynus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF90FF89FF4643D617BBFE45" bold="true" box="[151,482,393,419]" pageId="14" pageNumber="465">
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<emphasis id="23D0C82DFF90FF89FF4643D61719FE45" bold="true" box="[151,320,393,419]" italics="true" pageId="14" pageNumber="465">Hylobothynus</emphasis>
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<bibRefCitation id="753569CEFF90FF89FE9643D617BBFE45" author="Ohaus" box="[327,482,393,419]" pageId="14" pageNumber="465" refString="Ohaus, F. (1910) Neue sudamericanische Dynastiden (Col.) Deutsche Entomologische Zeitschrift, 1910, 671 - 690." type="journal article" year="1910">Ohaus, 1910</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="111B143FFF90FF89FF4643F31777FE22" blockId="14.[151,482,393,453]" box="[151,302,428,453]" pageId="14" pageNumber="465">
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(
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<figureCitation id="899F08BAFF90FF89FF4E43F3177CFE22" box="[159,293,428,453]" captionStart="FIGURES 40 – 48" captionStartId="14.[151,264,1664,1687]" captionTargetBox="[160,1429,889,1640]" captionTargetId="figure@14.[151,1436,878,1643]" captionTargetPageId="14" captionText="FIGURES 40 – 48. Hylobothynus columbianus (holotype): 40) habitus in dorsal view, 41) parameres in frontal view, 42) parameres in lateral view, 43) maxillary apical palpomere, 44) habitus in lateral view. Hylobothynus obesus (holotype): 45) habitus in dorsal view, 46) parameres in frontal view, 47) parameres in lateral view, 48) locality records of Hylobothynus in Colombia." httpUri="https://zenodo.org/record/234028/files/figure.png" pageId="14" pageNumber="465">
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Figs. 40
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<emphasis id="23D0C82DFF90FF89FF2B43F31751FE23" bold="true" box="[250,264,428,453]" pageId="14" pageNumber="465">–</emphasis>
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48
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</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="59BE47B4FF90FF89FF4643AB1248FDEC" box="[151,1041,500,522]" pageId="14" pageNumber="465" type="reference_group">
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<paragraph id="111B143FFF90FF89FF4643AB1248FDEC" blockId="14.[151,1041,500,522]" box="[151,1041,500,522]" pageId="14" pageNumber="465">
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<treatmentCitationGroup id="31B43311FF90FF89FF4643AB1248FDEC" box="[151,1041,500,522]" pageId="14" pageNumber="465">
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<taxonomicName id="D6A46FBCFF90FF89FF4643AB17B9FDEC" authority="Ohaus, 1910: 677" authorityName="Ohaus" authorityPageNumber="677" authorityYear="1910" box="[151,480,500,522]" class="Insecta" family="Dynastidae" genus="Hylobothynus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF90FF89FF4643AB177DFDEC" box="[151,292,500,522]" italics="true" pageId="14" pageNumber="465">Hylobothynus</emphasis>
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<treatmentCitation id="9005322EFF90FF89FEF843AB17B9FDEC" author="Ohaus" box="[297,480,500,522]" page="677" pageId="14" pageNumber="465" year="1910">
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<bibRefCitation id="753569CEFF90FF89FEF843AB17F4FDEC" author="Ohaus" box="[297,429,500,522]" pageId="14" pageNumber="465" refString="Ohaus, F. (1910) Neue sudamericanische Dynastiden (Col.) Deutsche Entomologische Zeitschrift, 1910, 671 - 690." type="journal article" year="1910">Ohaus, 1910</bibRefCitation>
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: 677
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</treatmentCitation>
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</taxonomicName>
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.
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<typeStatus id="CE1FAA9DFF90FF89FE3843AA1442FDEC" box="[489,539,501,522]" pageId="14" pageNumber="465">Type</typeStatus>
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species:
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<taxonomicName id="D6A46FBCFF90FF89FDA843AB1254FDEC" authority="Ohaus, 1910: 678" authorityName="Ohaus" authorityPageNumber="678" authorityYear="1910" box="[633,1037,500,522]" class="Insecta" family="Dynastidae" genus="Hylobothynus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="obesus">
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<emphasis id="23D0C82DFF90FF89FDA843AB1508FDEC" box="[633,849,500,522]" italics="true" pageId="14" pageNumber="465">Hylobothynus obesus</emphasis>
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<treatmentCitation id="9005322EFF90FF89FC8743AB1254FDEC" author="Ohaus" box="[854,1037,500,522]" page="678" pageId="14" pageNumber="465" year="1910">
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<bibRefCitation id="753569CEFF90FF89FC8743AB1583FDEC" author="Ohaus" box="[854,986,500,522]" pageId="14" pageNumber="465" refString="Ohaus, F. (1910) Neue sudamericanische Dynastiden (Col.) Deutsche Entomologische Zeitschrift, 1910, 671 - 690." type="journal article" year="1910">Ohaus, 1910</bibRefCitation>
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: 678
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="59BE47B4FF90FF89FF4640691528FD5D" pageId="14" pageNumber="465" type="diagnosis">
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<paragraph id="111B143FFF90FF89FF4640691528FD5D" blockId="14.[151,1437,566,843]" pageId="14" pageNumber="465">
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<emphasis id="23D0C82DFF90FF89FF4640691748FDA9" bold="true" box="[151,273,566,591]" pageId="14" pageNumber="465">Diagnosis.</emphasis>
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The apical maxillary palpomere is triangular and expanded towards the apex (Fig. 6E), a unique character among the genera of the tribe. Furthermore, members of the genus can be recognized by the following: clypeus triangular, strongly narrowed towards apex, with 2 apical teeth; frontal carina evident; mandibles tridentate; protibia with 3 teeth; males with protarsi not enlarged.
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</paragraph>
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</subSubSection>
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<subSubSection id="59BE47B4FF90FF89FF16409915A4FCAD" pageId="14" pageNumber="465" type="distribution">
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<paragraph id="111B143FFF90FF89FF16409915A4FCAD" blockId="14.[151,1437,566,843]" pageId="14" pageNumber="465">
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<emphasis id="23D0C82DFF90FF89FF1640991739FD39" bold="true" box="[199,352,710,735]" pageId="14" pageNumber="465">Distribution.</emphasis>
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<taxonomicName id="D6A46FBCFF90FF89FEB84099145CFD39" box="[361,517,710,735]" class="Insecta" family="Dynastidae" genus="Hylobothynus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF90FF89FEB84099145CFD39" box="[361,517,710,735]" italics="true" pageId="14" pageNumber="465">Hylobothynus</emphasis>
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</taxonomicName>
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is mainly an Amazonian genus (
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<bibRefCitation id="753569CEFF90FF89FCAC4099127DFD39" author="Ratcliffe" box="[893,1060,710,735]" pageId="14" pageNumber="465" refString="Ratcliffe, B. C. (1981) The fourth Hylobothynus: spanning the Amazonian gap (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin, 35, 303 - 306." type="journal article" year="1981">Ratcliffe 1981</bibRefCitation>
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), currently containing five South American species; one from
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<collectingCountry id="69B354AFFF90FF89FE2340B514F8FCE5" box="[498,673,746,771]" name="French Guiana" pageId="14" pageNumber="465">French Guiana</collectingCountry>
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; two from
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<collectingCountry id="69B354AFFF90FF89FCFF40B515C6FCE5" box="[814,927,746,771]" name="Colombia" pageId="14" pageNumber="465">Colombia</collectingCountry>
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,
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<collectingCountry id="69B354AFFF90FF89FC6040B51249FCE5" box="[945,1040,746,771]" name="Ecuador" pageId="14" pageNumber="465">Ecuador</collectingCountry>
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, and
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<collectingCountry id="69B354AFFF90FF89FB8640B412D7FCE4" box="[1111,1166,747,770]" name="Peru" pageId="14" pageNumber="465">Peru</collectingCountry>
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; and two from
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<collectingCountry id="69B354AFFF90FF89FA8040B513C0FCE5" box="[1361,1433,746,771]" name="Brazil" pageId="14" pageNumber="465">Brazil</collectingCountry>
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. However, these species are not common in
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<collectingCountry id="69B354AFFF90FF89FD474151155EFCC1" box="[662,775,782,807]" name="Colombia" pageId="14" pageNumber="465">Colombia</collectingCountry>
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, and there have been no recent collections, so only
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<typeStatus id="CE1FAA9DFF90FF89FABD415013C5FCC1" box="[1388,1436,783,807]" pageId="14" pageNumber="465">type</typeStatus>
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material was available. Therefore, only a diagnosis is shown for each species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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