The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution Author López-García, Margarita M. Author Gasca-Álvarez, Héctor J. Author Amat-García, Germán text Zootaxa 2015 4048 4 451 492 journal article 10.11646/zootaxa.4048.4.1 bbb51c92-e184-489d-aba8-18d1dc9a25cd 1175-5326 234023 71E2B8DA-825E-429F-BAA0-F582702B4A80 Hylobothynus Ohaus, 1910 ( Figs. 40 – 48 ) Hylobothynus Ohaus, 1910 : 677 . Type species: Hylobothynus obesus Ohaus, 1910 : 678 . Diagnosis. The apical maxillary palpomere is triangular and expanded towards the apex (Fig. 6E), a unique character among the genera of the tribe. Furthermore, members of the genus can be recognized by the following: clypeus triangular, strongly narrowed towards apex, with 2 apical teeth; frontal carina evident; mandibles tridentate; protibia with 3 teeth; males with protarsi not enlarged. Distribution. Hylobothynus is mainly an Amazonian genus ( Ratcliffe 1981 ), currently containing five South American species; one from French Guiana ; two from Colombia , Ecuador , and Peru ; and two from Brazil . However, these species are not common in Colombia , and there have been no recent collections, so only type material was available. Therefore, only a diagnosis is shown for each species.