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<mods:namePart id= "9AFE036A7C5AD69511F1267E8D763453" > Jin, Yuan Yuan</mods:namePart>
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<mods:namePart id= "2FFC2659D47DF6460B67A94EAFDA0594" > Han, Hui Lin</mods:namePart>
<mods:affiliation id= "8155119FFD149A9FE5397E2941E6AC2E" > Northeast Forestry University, School of Forestry, Harbin 150040, China. & Northeast Forestry University, Northeast Asia Biodiversity Research Center, Harbin 150040, China. & Northeast Forestry University, Ministry of Education, Key Laboratory of Sustainable Forest Ecosystem Management, Harbin 150040, China. Corresponding author</mods:affiliation>
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<taxonomicName id= "4C1B4D0BC87BF355FF07FC48FE9A7EE5" authorityName= "Jin & Han" authorityYear= "2025" box= "[151,354,942,969]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "puerensis" status= "sp. nov." >
<emphasis id= "B96FEA9AC87BF355FF07FC48FE9A7EE5" bold= "true" box= "[151,354,942,969]" italics= "true" pageId= "4" pageNumber= "583" > Rivula puerensis</emphasis>
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(Chinese name:
<emphasis id= "B96FEA9AC87BF355FEDDFC35FE217EC7" bold= "true" box= "[333,473,979,1002]" pageId= "4" pageNumber= "583" > Ḃ洱涓njď</emphasis>
)
</paragraph>
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<paragraph id= "8BA43688C87BF355FF07FBFCFEBE7919" blockId= "4.[151,952,942,1077]" box= "[151,326,1050,1077]" pageId= "4" pageNumber= "583" > Figures 4, 5, 14</paragraph>
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<paragraph id= "8BA43688C87BF355FF07FB84FC6479E8" blockId= "4.[151,1437,1122,2013]" pageId= "4" pageNumber= "583" >
<emphasis id= "B96FEA9AC87BF355FF07FB84FEC67951" bold= "true" box= "[151,318,1122,1148]" pageId= "4" pageNumber= "583" > Type material</emphasis>
.
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<materialsCitation id= "3B733CD5C87BF355FEDDFB84FDE0798C" collectingDate= "2009-05-15" collectingDateMax= "2009-05-19" collectingDateMin= "2009-05-15" collectionCode= "NEFU" collectorName= "HL. Han & MJ. Qi" country= "China" county= "Puer City" location= "Simao" municipality= "Simao" pageId= "4" pageNumber= "583" specimenCount= "1" stateProvince= "Yunnan" typeStatus= "holotype" >
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.
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,
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, Prov.
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,
<collectingCounty id= "62C54E04C87BF355FC93FB85FC977951" box= "[771,879,1122,1149]" pageId= "4" pageNumber= "583" > Puer City</collectingCounty>
,
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,
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, leg.
<collectorName id= "26EE535EC87BF355FB46FB85FAC37950" box= "[1238,1339,1123,1149]" pageId= "4" pageNumber= "583" > HL. Han</collectorName>
,
<collectorName id= "26EE535EC87BF355FAD7FB85FA6F7951" box= "[1351,1431,1122,1149]" pageId= "4" pageNumber= "583" > MJ. Qi</collectorName>
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, genit. prep. JYY-75-1, coll.
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.
2025-02-06 20:26:23 +01:00
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.
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,
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, Prov.
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,
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,
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,
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leg.
<collectorName id= "26EE535EC87BF355FF57FB4DFED479E8" box= "[199,300,1195,1221]" pageId= "4" pageNumber= "583" > HL. Han</collectorName>
,
<collectorName id= "26EE535EC87BF355FEA7FB4DFE7479E9" box= "[311,396,1194,1221]" pageId= "4" pageNumber= "583" > MR. Li</collectorName>
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, genit. prep. JYY-73-1, JYY-74-1, coll.
<collectionCode id= "ED0AAE4DC87BF355FCDBFB4DFC6079E8" box= "[843,920,1195,1221]" pageId= "4" pageNumber= "583" > NEFU</collectionCode>
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.
</paragraph>
</subSubSection>
<subSubSection id= "C3016503C87BF355FF57FB28FC8D7B7D" pageId= "4" pageNumber= "583" type= "diagnosis" >
<paragraph id= "8BA43688C87BF355FF57FB28FB177821" blockId= "4.[151,1437,1122,2013]" pageId= "4" pageNumber= "583" >
<emphasis id= "B96FEA9AC87BF355FF57FB28FEC279C5" bold= "true" box= "[199,314,1230,1256]" pageId= "4" pageNumber= "583" > Diagnosis</emphasis>
. The new species is similar to
<taxonomicName id= "4C1B4D0BC87BF355FD1DFB29FCB079C5" authorityName= "Holloway" authorityYear= "2008" box= "[653,840,1230,1256]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FD1DFB29FCB079C5" box= "[653,840,1230,1256]" italics= "true" pageId= "4" pageNumber= "583" > R. calcaripuncta</emphasis>
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in morphological characters and male genitalia (Figs. 6, 15), but it differs in the following characters (characters for
<taxonomicName id= "4C1B4D0BC87BF355FCC3FB15FBF67821" authorityName= "Holloway" authorityYear= "2008" box= "[851,1038,1266,1292]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FCC3FB15FBF67821" box= "[851,1038,1266,1292]" italics= "true" pageId= "4" pageNumber= "583" > R. calcaripuncta</emphasis>
</taxonomicName>
are in parentheses):
</paragraph>
<paragraph id= "8BA43688C87BF355FF57FAF0FAA078B1" blockId= "4.[151,1437,1122,2013]" pageId= "4" pageNumber= "583" >
Adult of
<taxonomicName id= "4C1B4D0BC87BF355FEB5FAF1FE5F781D" authorityName= "Jin & Han" authorityYear= "2025" box= "[293,423,1303,1328]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "puerensis" status= "sp. nov." >
<emphasis id= "B96FEA9AC87BF355FEB5FAF1FE5F781D" box= "[293,423,1303,1328]" italics= "true" pageId= "4" pageNumber= "583" > R. puerensis</emphasis>
</taxonomicName>
<emphasis id= "B96FEA9AC87BF355FE3BFAF0FDF8781D" bold= "true" box= "[427,512,1302,1328]" pageId= "4" pageNumber= "583" >
<taxonomicNameLabel id= "A25C57E1C87BF355FE3BFAF0FDF8781D" box= "[427,512,1302,1328]" pageId= "4" pageNumber= "583" rank= "species" > sp. nov.</taxonomicNameLabel>
</emphasis>
(Figs. 4, 5), forewing with few pale spots of various sizes, which is surrounded by black (more pale spots and pale spots even in size in
<taxonomicName id= "4C1B4D0BC87BF355FD06FADCFCD47879" authorityName= "Holloway" authorityYear= "2008" box= "[662,812,1338,1364]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FD06FADCFCD47879" box= "[662,812,1338,1364]" italics= "true" pageId= "4" pageNumber= "583" > calcaripuncta</emphasis>
</taxonomicName>
); ground color dark brownish black at costal margin area (uniformly colored in
<taxonomicName id= "4C1B4D0BC87BF355FE12FAB8FDE07855" authorityName= "Holloway" authorityYear= "2008" box= "[386,536,1374,1400]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FE12FAB8FDE07855" box= "[386,536,1374,1400]" italics= "true" pageId= "4" pageNumber= "583" > calcaripuncta</emphasis>
</taxonomicName>
); orbicular spot with large white dot (small white dot in
<taxonomicName id= "4C1B4D0BC87BF355FBFBFAB8FAF97855" authorityName= "Holloway" authorityYear= "2008" box= "[1131,1281,1374,1400]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FBFBFAB8FAF97855" box= "[1131,1281,1374,1400]" italics= "true" pageId= "4" pageNumber= "583" > calcaripuncta</emphasis>
</taxonomicName>
); the reniform spot formed by 2 irregular and small white spots, which is surrounded by black (without black in
<taxonomicName id= "4C1B4D0BC87BF355FB22FA64FAB078B1" authorityName= "Holloway" authorityYear= "2008" box= "[1202,1352,1410,1436]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FB22FA64FAB078B1" box= "[1202,1352,1410,1436]" italics= "true" pageId= "4" pageNumber= "583" > calcaripuncta</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id= "8BA43688C87BF355FF57FA41FC8D7B7D" blockId= "4.[151,1437,1122,2013]" pageId= "4" pageNumber= "583" >
In the male genitalia of
<taxonomicName id= "4C1B4D0BC87BF355FE46FA41FDA778ED" authorityName= "Jin & Han" authorityYear= "2025" box= "[470,607,1447,1472]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "puerensis" status= "sp. nov." >
<emphasis id= "B96FEA9AC87BF355FE46FA41FDA778ED" box= "[470,607,1447,1472]" italics= "true" pageId= "4" pageNumber= "583" > R. puerensis</emphasis>
</taxonomicName>
<emphasis id= "B96FEA9AC87BF355FDF7FA40FD3A78ED" bold= "true" box= "[615,706,1446,1472]" pageId= "4" pageNumber= "583" >
<taxonomicNameLabel id= "A25C57E1C87BF355FDF7FA40FD3A78ED" box= "[615,706,1446,1472]" pageId= "4" pageNumber= "583" rank= "species" > sp. nov.</taxonomicNameLabel>
</emphasis>
(Fig. 14), valva as long as tegumen (3/5 times longer than valva in
<taxonomicName id= "4C1B4D0BC87BF355FF24FA2CFEA878C9" authorityName= "Holloway" authorityYear= "2008" box= "[180,336,1482,1508]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FF24FA2CFEA878C9" box= "[180,336,1482,1508]" italics= "true" pageId= "4" pageNumber= "583" > calcaripuncta</emphasis>
</taxonomicName>
); cucullus rounded and flat (narrow and oval-shaped in
<taxonomicName id= "4C1B4D0BC87BF355FC56FA2CFB9A78C9" authorityName= "Holloway" authorityYear= "2008" box= "[966,1122,1482,1508]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FC56FA2CFB9A78C9" box= "[966,1122,1482,1508]" italics= "true" pageId= "4" pageNumber= "583" > calcaripuncta</emphasis>
</taxonomicName>
); saccus broad and strongly sclerotized and tongue-shaped (short and narrow in
<taxonomicName id= "4C1B4D0BC87BF355FD72FA08FC867B25" authorityName= "Holloway" authorityYear= "2008" box= "[738,894,1518,1544]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FD72FA08FC867B25" box= "[738,894,1518,1544]" italics= "true" pageId= "4" pageNumber= "583" > calcaripuncta</emphasis>
</taxonomicName>
). Aedeagus tube-shaped and of equal thickness, strongly sclerotized (slightly sclerotized in
<taxonomicName id= "4C1B4D0BC87BF355FD13F9F4FCE77B01" authorityName= "Holloway" authorityYear= "2008" box= "[643,799,1554,1580]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FD13F9F4FCE77B01" box= "[643,799,1554,1580]" italics= "true" pageId= "4" pageNumber= "583" > calcaripuncta</emphasis>
</taxonomicName>
and gradually becoming thicker from cecum to carina); carinal plate sclerotized, with spine bar (absent in
<taxonomicName id= "4C1B4D0BC87BF355FD59F9D0FC9D7B7D" authorityName= "Holloway" authorityYear= "2008" box= "[713,869,1590,1616]" class= "Insecta" family= "Erebidae" genus= "Rivula" kingdom= "Animalia" order= "Lepidoptera" pageId= "4" pageNumber= "583" phylum= "Arthropoda" rank= "species" species= "calcaripuncta" >
<emphasis id= "B96FEA9AC87BF355FD59F9D0FC9D7B7D" box= "[713,869,1590,1616]" italics= "true" pageId= "4" pageNumber= "583" > calcaripuncta</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id= "C3016503C87BF354FF57F9BCFC557F11" lastPageId= "5" lastPageNumber= "584" pageId= "4" pageNumber= "583" type= "description" >
<paragraph id= "8BA43688C87BF355FF57F9BCFCAF7AF1" blockId= "4.[151,1437,1122,2013]" pageId= "4" pageNumber= "583" >
<emphasis id= "B96FEA9AC87BF355FF57F9BCFEB67B59" bold= "true" box= "[199,334,1626,1652]" pageId= "4" pageNumber= "583" > Description</emphasis>
.
<emphasis id= "B96FEA9AC87BF355FECBF9BCFE667B59" bold= "true" box= "[347,414,1626,1652]" pageId= "4" pageNumber= "583" > Adult</emphasis>
(Figs. 4, 5). Wingspan
<quantity id= "4CE39B6DC87BF355FD33F9BCFCE47B58" box= "[675,796,1626,1653]" metricMagnitude= "-2" metricUnit= "m" metricValue= "1.15" metricValueMax= "1.2" metricValueMin= "1.1" pageId= "4" pageNumber= "583" unit= "mm" value= "11.5" valueMax= "12.0" valueMin= "11.0" > 11– </quantity>
. Head yellowish brown; labial palpi brown and upcurved, extending forward but not beyond the frontal tuft; antennae brown and filiform and lighter at the base; Thorax dark brown red; patagium and tegula lighter.Abdomen dark brown, and scales scattered throughout, particularly posteriorly. Forewing brownish black to brown, costal margin much darker than ground color; the basal line only with a white dot at costal margin; antemedial line weakly excurved, before vein Cu, after Cu incurved to 2A, then excurved to inner margin, formed by 6 white dots, surrounded by black; median line only with a small white dot at costal margin; postmedial lines roughly similar to antemedial line; subterminal line dark brown to blackish brown, very thin, with white and small spots, indistinct, surrounded entirely by black; fringe long, grayish brown; orbicular spot large with white dot; reniform spot formed by 2 small and white spots, surrounded entirely by black-brown, antemedial and basal line region darker than ground color. Hindwing blackish brown with diffuse smoke gray; discal spot indistinct; fringe more chocolate-brown, but with grayish brown on inner margin.
</paragraph>
<paragraph id= "8BA43688C87AF354FF57FF70FB5B7C81" blockId= "5.[151,1437,150,645]" pageId= "5" pageNumber= "584" >
<emphasis id= "B96FEA9AC87AF354FF57FF70FE947D9D" bold= "true" box= "[199,364,150,176]" pageId= "5" pageNumber= "584" > Male genitalia</emphasis>
(Fig. 14). Uncus sclerotized, sickle-shaped, basal part broad, approximately 4/5 as long as tegumen. Tegumen broad and slightly sclerotized, inverted funnel-shaped. Vinculum slender and strongly sclerotized. Juxta narrow, slightly sclerotized, and large toast-shaped. Valva more membranous, hammer-shaped, flat on apex, and thinly covered with small trichopore; costa thinner than sacculus, nearly 1/2 width of valva base, and with a large and arc-shape concavity at basal part; cucullus membranous and rounded, flat on apex; saccus bulky and tongue-shaped.Aedeagus of equal thickness, tube-shaped, strongly sclerotized; cecum 1/4 as long as aedeagus, carina sclerotized, serrate-shaped. Vesica membranous, with 1 subbasal diverticulum, which covered shorter conical-shaped cornute plate; and 5 median diverticula, bearing numerous mastoid-shaped cornuti on the 4th and 5th median diverticulum.
</paragraph>
<paragraph id= "8BA43688C87AF354FF57FE50FDE87CFD" blockId= "5.[151,1437,150,645]" box= "[199,528,438,465]" pageId= "5" pageNumber= "584" >
<emphasis id= "B96FEA9AC87AF354FF57FE50FE747CFD" bold= "true" box= "[199,396,438,464]" pageId= "5" pageNumber= "584" > Female genitalia</emphasis>
. Unknown.
</paragraph>
<paragraph id= "8BA43688C87AF354FF57FE3CFC557F11" blockId= "5.[151,1437,150,645]" pageId= "5" pageNumber= "584" >
<emphasis id= "B96FEA9AC87AF354FF57FE3CFEBB7CD9" bold= "true" box= "[199,323,474,500]" pageId= "5" pageNumber= "584" > Bionomics</emphasis>
. This species was collected between May and early August using a light trap near a coniferous forest. The vegetation around the
<typeStatus id= "54A0882AC87AF354FD8FFE18FD797F35" box= "[543,641,510,536]" pageId= "5" pageNumber= "584" type= "holotype" > holotype</typeStatus>
collection site includes
<taxonomicName id= "4C1B4D0BC87AF354FC0BFE19FB367F35" authority= "Franch." authorityName= "Franch." box= "[923,1230,510,536]" class= "Pinopsida" family= "Pinaceae" genus= "Pinus" kingdom= "Plantae" order= "Pinales" pageId= "5" pageNumber= "584" phylum= "Tracheophyta" rank= "species" species= "yunnanensis" >
<emphasis id= "B96FEA9AC87AF354FC0BFE19FB967F35" box= "[923,1134,511,536]" italics= "true" pageId= "5" pageNumber= "584" > Pinus yunnanensis</emphasis>
Franch.
</taxonomicName>
(
<taxonomicName id= "4C1B4D0BC87AF354FB4FFE18FAB17F35" box= "[1247,1353,510,536]" class= "Pinopsida" family= "Pinaceae" kingdom= "Plantae" order= "Pinales" pageId= "5" pageNumber= "584" phylum= "Tracheophyta" rank= "family" > Pinaceae</taxonomicName>
), along with grasslands predominantly featuring
<taxonomicName id= "4C1B4D0BC87AF354FDF2FDC5FD467F10" box= "[610,702,547,573]" class= "Liliopsida" family= "Poaceae" kingdom= "Plantae" order= "Poales" pageId= "5" pageNumber= "584" phylum= "Tracheophyta" rank= "family" > Poaceae</taxonomicName>
plants (Figs. 20, 21).
</paragraph>
</subSubSection>
<subSubSection id= "C3016503C87AF354FF57FDA0FDD97F4D" box= "[199,545,582,609]" pageId= "5" pageNumber= "584" type= "distribution" >
<paragraph id= "8BA43688C87AF354FF57FDA0FDD97F4D" blockId= "5.[151,1437,150,645]" box= "[199,545,582,609]" pageId= "5" pageNumber= "584" >
<emphasis id= "B96FEA9AC87AF354FF57FDA0FEA17F4D" bold= "true" box= "[199,345,582,608]" pageId= "5" pageNumber= "584" > Distribution</emphasis>
.
<collectingCountry id= "F30C7618C87AF354FEF7FDA0FE527F4D" box= "[359,426,582,608]" name= "China" pageId= "5" pageNumber= "584" > China</collectingCountry>
(
<collectingRegion id= "49DFF86AC87AF354FE29FDA0FDED7F4D" box= "[441,533,582,608]" country= "China" name= "Yunnan" pageId= "5" pageNumber= "584" > Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
2025-02-06 20:26:23 +01:00
<subSubSection id= "C3016503C87AF354FF57FD8CFBD87FA8" box= "[199,1056,618,645]" pageId= "5" pageNumber= "584" type= "etymology" >
2025-02-04 21:03:19 +01:00
<paragraph id= "8BA43688C87AF354FF57FD8CFBD87FA8" blockId= "5.[151,1437,150,645]" box= "[199,1056,618,645]" pageId= "5" pageNumber= "584" >
<emphasis id= "B96FEA9AC87AF354FF57FD8CFEB07FA9" bold= "true" box= "[199,328,618,644]" pageId= "5" pageNumber= "584" > Etymology</emphasis>
. This species name is derived from the locality of the
<typeStatus id= "54A0882AC87AF354FC38FD8DFC237FA8" box= "[936,987,619,645]" pageId= "5" pageNumber= "584" > type</typeStatus>
, Puer.
</paragraph>
</subSubSection>
</treatment>
</document>
