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<document id="D82D76991308C6498FFD5421EFA7017A" ID-CLB-Dataset="308024" ID-DOI="10.11646/zootaxa.5583.3.10" ID-GBIF-Dataset="befbcf80-cb1a-43f4-b120-3b4d4a6b7b1c" ID-ISSN="1175-5326" ID-Zenodo-Dep="14804463" ID-ZooBank="80B79386-3B99-4FF3-B509-E36FB4DB7285" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1738698321192" checkinUser="plazi" docAuthor="Jin, Yuan Yuan &amp; Han, Hui Lin" docDate="2025" docId="03B2879EC87BF354FF07FC48FBD87FA8" docLanguage="en" docName="zootaxa.5583.3.10.pdf" docOrigin="Zootaxa 5583 (3)" docSource="https://doi.org/10.11646/zootaxa.5583.3.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rivula puerensis Jin &amp; Han, 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="584" masterDocId="FF8BFFE6C87FF351FF90FFE6FFF87D2D" masterDocTitle="Descriptions of three new species of Rivula Guenée [1845] 1844 (Lepidoptera, Erebidae, Rivulinae) from China, with the checklist of Chinese species" masterLastPageNumber="588" masterPageNumber="579" pageNumber="583" updateTime="1738869718305" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3DDA3A58DA1703E4C7E7E6A02C4656C5">Descriptions of three new species of Rivula Guenée [1845] 1844 (Lepidoptera, Erebidae, Rivulinae) from China, with the checklist of Chinese species</mods:title>
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<mods:namePart id="9AFE036A7C5AD69511F1267E8D763453">Jin, Yuan Yuan</mods:namePart>
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<mods:affiliation id="9D81AE09A29D877E3017EDC62BEE46FF">Northeast Forestry University, School of Forestry, Harbin 150040, China.</mods:affiliation>
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<mods:namePart id="2FFC2659D47DF6460B67A94EAFDA0594">Han, Hui Lin</mods:namePart>
<mods:affiliation id="8155119FFD149A9FE5397E2941E6AC2E">Northeast Forestry University, School of Forestry, Harbin 150040, China. &amp; Northeast Forestry University, Northeast Asia Biodiversity Research Center, Harbin 150040, China. &amp; Northeast Forestry University, Ministry of Education, Key Laboratory of Sustainable Forest Ecosystem Management, Harbin 150040, China. Corresponding author</mods:affiliation>
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<paragraph id="8BA43688C87BF355FF07FC48FE3F7EE5" blockId="4.[151,952,942,1077]" box="[151,455,942,969]" pageId="4" pageNumber="583">
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<emphasis id="B96FEA9AC87BF355FF07FC48FE3F7EE5" bold="true" box="[151,455,942,969]" pageId="4" pageNumber="583">
<taxonomicName id="4C1B4D0BC87BF355FF07FC48FE9A7EE5" authorityName="Jin &amp; Han" authorityYear="2025" box="[151,354,942,969]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="puerensis" status="sp. nov.">
<emphasis id="B96FEA9AC87BF355FF07FC48FE9A7EE5" bold="true" box="[151,354,942,969]" italics="true" pageId="4" pageNumber="583">Rivula puerensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E1C87BF355FEF9FC48FE3F7EE5" box="[361,455,942,968]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BA43688C87BF355FF07FC34FE1A7EC1" blockId="4.[151,952,942,1077]" box="[151,482,978,1005]" pageId="4" pageNumber="583">
(Chinese name:
<emphasis id="B96FEA9AC87BF355FEDDFC35FE217EC7" bold="true" box="[333,473,979,1002]" pageId="4" pageNumber="583">Ḃ洱涓njď</emphasis>
)
</paragraph>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BA43688C87BF355FF07FBFCFEBE7919" blockId="4.[151,952,942,1077]" box="[151,326,1050,1077]" pageId="4" pageNumber="583">Figures 4, 5, 14</paragraph>
</subSubSection>
<subSubSection id="C3016503C87BF355FF07FB84FC6479E8" pageId="4" pageNumber="583" type="materials_examined">
<paragraph id="8BA43688C87BF355FF07FB84FC6479E8" blockId="4.[151,1437,1122,2013]" pageId="4" pageNumber="583">
<emphasis id="B96FEA9AC87BF355FF07FB84FEC67951" bold="true" box="[151,318,1122,1148]" pageId="4" pageNumber="583">Type material</emphasis>
.
<materialsCitation id="3B733CD5C87BF355FEDDFB84FDE0798C" collectingDate="2009-05-15" collectingDateMax="2009-05-19" collectingDateMin="2009-05-15" collectionCode="NEFU" collectorName="HL. Han &amp; MJ. Qi" country="China" county="Puer City" location="Simao" municipality="Simao" pageId="4" pageNumber="583" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B96FEA9AC87BF355FEDDFB84FE427951" bold="true" box="[333,442,1122,1148]" pageId="4" pageNumber="583">
<typeStatus id="54A0882AC87BF355FEDDFB84FE427951" box="[333,442,1122,1148]" pageId="4" pageNumber="583" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="9D1DFD01C87BF355FE58FB84FDF97951" box="[456,513,1122,1148]" pageId="4" pageNumber="583" type="male">male</specimenCount>
,
<collectingCountry id="F30C7618C87BF355FD9DFB84FDAB7951" box="[525,595,1122,1148]" name="China" pageId="4" pageNumber="583">China</collectingCountry>
, Prov.
<collectingRegion id="49DFF86AC87BF355FD0FFB85FD007950" box="[671,760,1123,1149]" country="China" name="Yunnan" pageId="4" pageNumber="583">Yunnan</collectingRegion>
,
<collectingCounty id="62C54E04C87BF355FC93FB85FC977951" box="[771,879,1122,1149]" pageId="4" pageNumber="583">Puer City</collectingCounty>
,
<collectingMunicipality id="6BC0ACF2C87BF355FCEBFB84FC3D7951" box="[891,965,1122,1148]" pageId="4" pageNumber="583">Simao</collectingMunicipality>
,
<date id="FFA51048C87BF355FC41FB84FB627951" box="[977,1178,1122,1149]" pageId="4" pageNumber="583" value="2009-05-15" valueMax="2009-05-19" valueMin="2009-05-15">
<collectingDate id="EFE1E9A0C87BF355FC41FB84FB627951" box="[977,1178,1122,1149]" pageId="4" pageNumber="583" value="2009-05-15" valueMax="2009-05-19" valueMin="2009-05-15">1519 May. 2009</collectingDate>
</date>
, leg.
<collectorName id="26EE535EC87BF355FB46FB85FAC37950" box="[1238,1339,1123,1149]" pageId="4" pageNumber="583">HL. Han</collectorName>
,
<collectorName id="26EE535EC87BF355FAD7FB85FA6F7951" box="[1351,1431,1122,1149]" pageId="4" pageNumber="583">MJ. Qi</collectorName>
, genit. prep. JYY-75-1, coll.
<collectionCode id="ED0AAE4DC87BF355FE5BFB61FDE0798C" box="[459,536,1159,1185]" pageId="4" pageNumber="583">NEFU</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B733CD5C87BF355FDB1FB60FC6079E8" collectingDate="2018-08-01" collectingDateMax="2018-08-02" collectingDateMin="2018-08-01" collectionCode="NEFU" collectorName="HL. Han &amp; MR. Li" country="China" county="Puer City" location="Jiangcheng" municipality="Jiangcheng" pageId="4" pageNumber="583" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B96FEA9AC87BF355FDB1FB60FD61798D" bold="true" box="[545,665,1158,1184]" pageId="4" pageNumber="583">
<typeStatus id="54A0882AC87BF355FDB1FB60FD61798D" box="[545,665,1158,1184]" pageId="4" pageNumber="583" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<specimenCount id="9D1DFD01C87BF355FD36FB60FD06798D" box="[678,766,1158,1184]" count="2" pageId="4" pageNumber="583" type="male">2 males</specimenCount>
,
<collectingCountry id="F30C7618C87BF355FC98FB60FCB6798D" box="[776,846,1158,1184]" name="China" pageId="4" pageNumber="583">China</collectingCountry>
, Prov.
<collectingRegion id="49DFF86AC87BF355FC07FB61FC08798C" box="[919,1008,1159,1185]" country="China" name="Yunnan" pageId="4" pageNumber="583">Yunnan</collectingRegion>
,
<collectingCounty id="62C54E04C87BF355FC69FB61FB9C798D" box="[1017,1124,1158,1185]" pageId="4" pageNumber="583">Puer City</collectingCounty>
,
<collectingMunicipality id="6BC0ACF2C87BF355FBFEFB60FB17798D" box="[1134,1263,1158,1184]" pageId="4" pageNumber="583">Jiangcheng</collectingMunicipality>
,
<date id="FFA51048C87BF355FB69FB60FA64798D" box="[1273,1436,1158,1184]" pageId="4" pageNumber="583" value="2018-08-01" valueMax="2018-08-02" valueMin="2018-08-01">
<collectingDate id="EFE1E9A0C87BF355FB69FB60FA64798D" box="[1273,1436,1158,1184]" pageId="4" pageNumber="583" value="2018-08-01" valueMax="2018-08-02" valueMin="2018-08-01">12 Aug. 2018</collectingDate>
</date>
leg.
<collectorName id="26EE535EC87BF355FF57FB4DFED479E8" box="[199,300,1195,1221]" pageId="4" pageNumber="583">HL. Han</collectorName>
,
<collectorName id="26EE535EC87BF355FEA7FB4DFE7479E9" box="[311,396,1194,1221]" pageId="4" pageNumber="583">MR. Li</collectorName>
, genit. prep. JYY-73-1, JYY-74-1, coll.
<collectionCode id="ED0AAE4DC87BF355FCDBFB4DFC6079E8" box="[843,920,1195,1221]" pageId="4" pageNumber="583">NEFU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016503C87BF355FF57FB28FC8D7B7D" pageId="4" pageNumber="583" type="diagnosis">
<paragraph id="8BA43688C87BF355FF57FB28FB177821" blockId="4.[151,1437,1122,2013]" pageId="4" pageNumber="583">
<emphasis id="B96FEA9AC87BF355FF57FB28FEC279C5" bold="true" box="[199,314,1230,1256]" pageId="4" pageNumber="583">Diagnosis</emphasis>
. The new species is similar to
<taxonomicName id="4C1B4D0BC87BF355FD1DFB29FCB079C5" authorityName="Holloway" authorityYear="2008" box="[653,840,1230,1256]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FD1DFB29FCB079C5" box="[653,840,1230,1256]" italics="true" pageId="4" pageNumber="583">R. calcaripuncta</emphasis>
</taxonomicName>
in morphological characters and male genitalia (Figs. 6, 15), but it differs in the following characters (characters for
<taxonomicName id="4C1B4D0BC87BF355FCC3FB15FBF67821" authorityName="Holloway" authorityYear="2008" box="[851,1038,1266,1292]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FCC3FB15FBF67821" box="[851,1038,1266,1292]" italics="true" pageId="4" pageNumber="583">R. calcaripuncta</emphasis>
</taxonomicName>
are in parentheses):
</paragraph>
<paragraph id="8BA43688C87BF355FF57FAF0FAA078B1" blockId="4.[151,1437,1122,2013]" pageId="4" pageNumber="583">
Adult of
<taxonomicName id="4C1B4D0BC87BF355FEB5FAF1FE5F781D" authorityName="Jin &amp; Han" authorityYear="2025" box="[293,423,1303,1328]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="puerensis" status="sp. nov.">
<emphasis id="B96FEA9AC87BF355FEB5FAF1FE5F781D" box="[293,423,1303,1328]" italics="true" pageId="4" pageNumber="583">R. puerensis</emphasis>
</taxonomicName>
<emphasis id="B96FEA9AC87BF355FE3BFAF0FDF8781D" bold="true" box="[427,512,1302,1328]" pageId="4" pageNumber="583">
<taxonomicNameLabel id="A25C57E1C87BF355FE3BFAF0FDF8781D" box="[427,512,1302,1328]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Figs. 4, 5), forewing with few pale spots of various sizes, which is surrounded by black (more pale spots and pale spots even in size in
<taxonomicName id="4C1B4D0BC87BF355FD06FADCFCD47879" authorityName="Holloway" authorityYear="2008" box="[662,812,1338,1364]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FD06FADCFCD47879" box="[662,812,1338,1364]" italics="true" pageId="4" pageNumber="583">calcaripuncta</emphasis>
</taxonomicName>
); ground color dark brownish black at costal margin area (uniformly colored in
<taxonomicName id="4C1B4D0BC87BF355FE12FAB8FDE07855" authorityName="Holloway" authorityYear="2008" box="[386,536,1374,1400]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FE12FAB8FDE07855" box="[386,536,1374,1400]" italics="true" pageId="4" pageNumber="583">calcaripuncta</emphasis>
</taxonomicName>
); orbicular spot with large white dot (small white dot in
<taxonomicName id="4C1B4D0BC87BF355FBFBFAB8FAF97855" authorityName="Holloway" authorityYear="2008" box="[1131,1281,1374,1400]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FBFBFAB8FAF97855" box="[1131,1281,1374,1400]" italics="true" pageId="4" pageNumber="583">calcaripuncta</emphasis>
</taxonomicName>
); the reniform spot formed by 2 irregular and small white spots, which is surrounded by black (without black in
<taxonomicName id="4C1B4D0BC87BF355FB22FA64FAB078B1" authorityName="Holloway" authorityYear="2008" box="[1202,1352,1410,1436]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FB22FA64FAB078B1" box="[1202,1352,1410,1436]" italics="true" pageId="4" pageNumber="583">calcaripuncta</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BA43688C87BF355FF57FA41FC8D7B7D" blockId="4.[151,1437,1122,2013]" pageId="4" pageNumber="583">
In the male genitalia of
<taxonomicName id="4C1B4D0BC87BF355FE46FA41FDA778ED" authorityName="Jin &amp; Han" authorityYear="2025" box="[470,607,1447,1472]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="puerensis" status="sp. nov.">
<emphasis id="B96FEA9AC87BF355FE46FA41FDA778ED" box="[470,607,1447,1472]" italics="true" pageId="4" pageNumber="583">R. puerensis</emphasis>
</taxonomicName>
<emphasis id="B96FEA9AC87BF355FDF7FA40FD3A78ED" bold="true" box="[615,706,1446,1472]" pageId="4" pageNumber="583">
<taxonomicNameLabel id="A25C57E1C87BF355FDF7FA40FD3A78ED" box="[615,706,1446,1472]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Fig. 14), valva as long as tegumen (3/5 times longer than valva in
<taxonomicName id="4C1B4D0BC87BF355FF24FA2CFEA878C9" authorityName="Holloway" authorityYear="2008" box="[180,336,1482,1508]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FF24FA2CFEA878C9" box="[180,336,1482,1508]" italics="true" pageId="4" pageNumber="583">calcaripuncta</emphasis>
</taxonomicName>
); cucullus rounded and flat (narrow and oval-shaped in
<taxonomicName id="4C1B4D0BC87BF355FC56FA2CFB9A78C9" authorityName="Holloway" authorityYear="2008" box="[966,1122,1482,1508]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FC56FA2CFB9A78C9" box="[966,1122,1482,1508]" italics="true" pageId="4" pageNumber="583">calcaripuncta</emphasis>
</taxonomicName>
); saccus broad and strongly sclerotized and tongue-shaped (short and narrow in
<taxonomicName id="4C1B4D0BC87BF355FD72FA08FC867B25" authorityName="Holloway" authorityYear="2008" box="[738,894,1518,1544]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FD72FA08FC867B25" box="[738,894,1518,1544]" italics="true" pageId="4" pageNumber="583">calcaripuncta</emphasis>
</taxonomicName>
). Aedeagus tube-shaped and of equal thickness, strongly sclerotized (slightly sclerotized in
<taxonomicName id="4C1B4D0BC87BF355FD13F9F4FCE77B01" authorityName="Holloway" authorityYear="2008" box="[643,799,1554,1580]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FD13F9F4FCE77B01" box="[643,799,1554,1580]" italics="true" pageId="4" pageNumber="583">calcaripuncta</emphasis>
</taxonomicName>
and gradually becoming thicker from cecum to carina); carinal plate sclerotized, with spine bar (absent in
<taxonomicName id="4C1B4D0BC87BF355FD59F9D0FC9D7B7D" authorityName="Holloway" authorityYear="2008" box="[713,869,1590,1616]" class="Insecta" family="Erebidae" genus="Rivula" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="calcaripuncta">
<emphasis id="B96FEA9AC87BF355FD59F9D0FC9D7B7D" box="[713,869,1590,1616]" italics="true" pageId="4" pageNumber="583">calcaripuncta</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016503C87BF354FF57F9BCFC557F11" lastPageId="5" lastPageNumber="584" pageId="4" pageNumber="583" type="description">
<paragraph id="8BA43688C87BF355FF57F9BCFCAF7AF1" blockId="4.[151,1437,1122,2013]" pageId="4" pageNumber="583">
<emphasis id="B96FEA9AC87BF355FF57F9BCFEB67B59" bold="true" box="[199,334,1626,1652]" pageId="4" pageNumber="583">Description</emphasis>
.
<emphasis id="B96FEA9AC87BF355FECBF9BCFE667B59" bold="true" box="[347,414,1626,1652]" pageId="4" pageNumber="583">Adult</emphasis>
(Figs. 4, 5). Wingspan
<quantity id="4CE39B6DC87BF355FD33F9BCFCE47B58" box="[675,796,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="4" pageNumber="583" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">1112 mm</quantity>
. Head yellowish brown; labial palpi brown and upcurved, extending forward but not beyond the frontal tuft; antennae brown and filiform and lighter at the base; Thorax dark brown red; patagium and tegula lighter.Abdomen dark brown, and scales scattered throughout, particularly posteriorly. Forewing brownish black to brown, costal margin much darker than ground color; the basal line only with a white dot at costal margin; antemedial line weakly excurved, before vein Cu, after Cu incurved to 2A, then excurved to inner margin, formed by 6 white dots, surrounded by black; median line only with a small white dot at costal margin; postmedial lines roughly similar to antemedial line; subterminal line dark brown to blackish brown, very thin, with white and small spots, indistinct, surrounded entirely by black; fringe long, grayish brown; orbicular spot large with white dot; reniform spot formed by 2 small and white spots, surrounded entirely by black-brown, antemedial and basal line region darker than ground color. Hindwing blackish brown with diffuse smoke gray; discal spot indistinct; fringe more chocolate-brown, but with grayish brown on inner margin.
</paragraph>
<paragraph id="8BA43688C87AF354FF57FF70FB5B7C81" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="584">
<emphasis id="B96FEA9AC87AF354FF57FF70FE947D9D" bold="true" box="[199,364,150,176]" pageId="5" pageNumber="584">Male genitalia</emphasis>
(Fig. 14). Uncus sclerotized, sickle-shaped, basal part broad, approximately 4/5 as long as tegumen. Tegumen broad and slightly sclerotized, inverted funnel-shaped. Vinculum slender and strongly sclerotized. Juxta narrow, slightly sclerotized, and large toast-shaped. Valva more membranous, hammer-shaped, flat on apex, and thinly covered with small trichopore; costa thinner than sacculus, nearly 1/2 width of valva base, and with a large and arc-shape concavity at basal part; cucullus membranous and rounded, flat on apex; saccus bulky and tongue-shaped.Aedeagus of equal thickness, tube-shaped, strongly sclerotized; cecum 1/4 as long as aedeagus, carina sclerotized, serrate-shaped. Vesica membranous, with 1 subbasal diverticulum, which covered shorter conical-shaped cornute plate; and 5 median diverticula, bearing numerous mastoid-shaped cornuti on the 4th and 5th median diverticulum.
</paragraph>
<paragraph id="8BA43688C87AF354FF57FE50FDE87CFD" blockId="5.[151,1437,150,645]" box="[199,528,438,465]" pageId="5" pageNumber="584">
<emphasis id="B96FEA9AC87AF354FF57FE50FE747CFD" bold="true" box="[199,396,438,464]" pageId="5" pageNumber="584">Female genitalia</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BA43688C87AF354FF57FE3CFC557F11" blockId="5.[151,1437,150,645]" pageId="5" pageNumber="584">
<emphasis id="B96FEA9AC87AF354FF57FE3CFEBB7CD9" bold="true" box="[199,323,474,500]" pageId="5" pageNumber="584">Bionomics</emphasis>
. This species was collected between May and early August using a light trap near a coniferous forest. The vegetation around the
<typeStatus id="54A0882AC87AF354FD8FFE18FD797F35" box="[543,641,510,536]" pageId="5" pageNumber="584" type="holotype">holotype</typeStatus>
collection site includes
<taxonomicName id="4C1B4D0BC87AF354FC0BFE19FB367F35" authority="Franch." authorityName="Franch." box="[923,1230,510,536]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="584" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="B96FEA9AC87AF354FC0BFE19FB967F35" box="[923,1134,511,536]" italics="true" pageId="5" pageNumber="584">Pinus yunnanensis</emphasis>
Franch.
</taxonomicName>
(
<taxonomicName id="4C1B4D0BC87AF354FB4FFE18FAB17F35" box="[1247,1353,510,536]" class="Pinopsida" family="Pinaceae" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="584" phylum="Tracheophyta" rank="family">Pinaceae</taxonomicName>
), along with grasslands predominantly featuring
<taxonomicName id="4C1B4D0BC87AF354FDF2FDC5FD467F10" box="[610,702,547,573]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="5" pageNumber="584" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
plants (Figs. 20, 21).
</paragraph>
</subSubSection>
<subSubSection id="C3016503C87AF354FF57FDA0FDD97F4D" box="[199,545,582,609]" pageId="5" pageNumber="584" type="distribution">
<paragraph id="8BA43688C87AF354FF57FDA0FDD97F4D" blockId="5.[151,1437,150,645]" box="[199,545,582,609]" pageId="5" pageNumber="584">
<emphasis id="B96FEA9AC87AF354FF57FDA0FEA17F4D" bold="true" box="[199,345,582,608]" pageId="5" pageNumber="584">Distribution</emphasis>
.
<collectingCountry id="F30C7618C87AF354FEF7FDA0FE527F4D" box="[359,426,582,608]" name="China" pageId="5" pageNumber="584">China</collectingCountry>
(
<collectingRegion id="49DFF86AC87AF354FE29FDA0FDED7F4D" box="[441,533,582,608]" country="China" name="Yunnan" pageId="5" pageNumber="584">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016503C87AF354FF57FD8CFBD87FA8" box="[199,1056,618,645]" pageId="5" pageNumber="584" type="etymology">
<paragraph id="8BA43688C87AF354FF57FD8CFBD87FA8" blockId="5.[151,1437,150,645]" box="[199,1056,618,645]" pageId="5" pageNumber="584">
<emphasis id="B96FEA9AC87AF354FF57FD8CFEB07FA9" bold="true" box="[199,328,618,644]" pageId="5" pageNumber="584">Etymology</emphasis>
. This species name is derived from the locality of the
<typeStatus id="54A0882AC87AF354FC38FD8DFC237FA8" box="[936,987,619,645]" pageId="5" pageNumber="584">type</typeStatus>
, Puer.
</paragraph>
</subSubSection>
</treatment>
</document>