treatments-xml/data/03/A1/87/03A187A9292BFFE2FF03C5B59986FDED.xml

<document id="051EFA6FD22F2F64E353E7A7013393CB" ID="10.11646/zootaxa.4157.1.1" ID-CLB-Dataset="38320" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="diego" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Perkovsky, Evgeny" docDate="2016" docId="03A187A9292BFFE2FF03C5B59986FDED" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Vertheia quadrisetosa Jałoszyński &amp; Perkovsky, 2016, sp. n." docType="treatment" docVersion="13" lastPageNumber="20" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="19" updateTime="1738750171177" updateUser="ExternalLinkService">
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<mods:title id="07B153B1CCBD7A9B864200CBAA692741">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:namePart id="DCAF595600A9C675B627200F4E9B7C97">Jałoszyński, Paweł</mods:namePart>
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<treatment id="03A187A9292BFFE2FF03C5B59986FDED" ID-DOI="http://doi.org/10.5281/zenodo.5681737" ID-GBIF-Taxon="124407823" ID-Zenodo-Dep="5681737" LSID="urn:lsid:plazi:treatment:03A187A9292BFFE2FF03C5B59986FDED" httpUri="http://treatment.plazi.org/id/03A187A9292BFFE2FF03C5B59986FDED" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" scope_family="Staphylinidae" scope_order="Coleoptera" scope_tribe="Eutheiini">
<subSubSection id="C3126534292BFFE3FF03C5B59C17FF02" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BB736BF292BFFE3FF03C5B59CA1FF63" blockId="18.[151,501,151,211]" box="[151,501,151,178]" pageId="18" pageNumber="19">
<heading id="D0FF81D3292BFFE3FF03C5B59CA1FF63" bold="true" box="[151,501,151,178]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B97CEAAD292BFFE3FF03C5B59CA1FF63" bold="true" box="[151,501,151,178]" pageId="18" pageNumber="19">
† 
<taxonomicName id="4C084D3C292BFFE3FF31C5BA9CFFFF60" ID-CoL="5B8TR" box="[165,427,151,177]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quadrisetosa" status="sp. nov.">
<emphasis id="B97CEAAD292BFFE3FF31C5BA9CFFFF60" bold="true" box="[165,427,151,177]" italics="true" pageId="18" pageNumber="19">Vertheia quadrisetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D6292BFFE3FE26C5B59CA1FF63" box="[434,501,152,178]" pageId="18" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736BF292BFFE3FF03C5979C17FF02" blockId="18.[151,501,151,211]" box="[151,323,186,211]" pageId="18" pageNumber="19">
(
<figureCitation id="13332A3A292BFFE3FF0BC5979DB3FF02" box="[159,231,186,211]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." figureDoi="http://doi.org/10.5281/zenodo.262414" httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="18" pageNumber="19">Figs 2</figureCitation>
, 
<figureCitation id="13332A3A292BFFE3FF67C5979C6FFF02" box="[243,315,186,211]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">49–55</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3126534292BFFE3FF03C42F9956FEEE" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BB736BF292BFFE3FF03C42F9956FEEE" blockId="18.[151,1436,258,391]" pageId="18" pageNumber="19">
<materialsCitation id="3B603CE2292BFFE3FF03C42F9956FEEE" ID-GBIF-Occurrence="1300093515" collectionCode="SIZK" location="Europe" municipality="Late Eocene" pageId="18" pageNumber="19" specimenCount="2" stateProvince="Rovno" typeStatus="holotype">
<emphasis id="B97CEAAD292BFFE3FF03C42F9EFFFECA" bold="true" box="[151,939,258,283]" pageId="18" pageNumber="19">
<typeStatus id="54B3881D292BFFE3FF03C42F9D86FECA" box="[151,210,258,283]" pageId="18" pageNumber="19">Type</typeStatus>
material. 
<typeStatus id="54B3881D292BFFE3FEC6C42F9C97FECA" box="[338,451,258,283]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
: 
<collectingMunicipality id="6BD3ACC5292BFFE3FE46C42F9F33FECA" box="[466,615,258,283]" pageId="18" pageNumber="19">Late Eocene</collectingMunicipality>
of 
<location id="8ED76064292BFFE3FD07C42F9FB9FECA" LSID="urn:lsid:plazi:treatment:03A187A9292BFFE2FF03C5B59986FDED:8ED76064292BFFE3FD07C42F9FB9FECA" box="[659,749,258,283]" municipality="Late Eocene" name="Europe" pageId="18" pageNumber="19" stateProvince="Rovno">Europe</location>
, 
<collectingRegion id="49CCF85D292BFFE3FD68C42F9E1EFECA" box="[764,842,258,283]" country="Ukraine" name="Rivne" pageId="18" pageNumber="19">Rovno</collectingRegion>
amber:
</emphasis>
sex unknown; inclusion in elongate, dropshaped piece of amber 
<quantity id="4CF09B5A292BFFE3FE0DC40A9CBCFEEF" box="[409,488,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="18" pageNumber="19" unit="mm" value="14.0">14 mm</quantity>
long (
<figureCitation id="13332A3A292BFFE3FDA4C40B9F20FEEE" box="[560,628,294,319]" captionStart="FIGURES 1 – 20" captionStartId="7.[151,264,1794,1817]" captionTargetBox="[157,1423,512,1763]" captionTargetId="figure@7.[151,1436,503,1772]" captionTargetPageId="7" captionText="FIGURES 1 – 20. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 27418, Rovnoleptochromus ableptonoides sp. n. (1); K- 243, Vertheia quadrisetosa sp. n. (2); UA- 442, Vertheia sp. 1 (3); K- 800, Cephennodes sp. 1 (4); K- 6011, Cephennomicrus giganteus sp. n. (5); K- 8289, Cephennomicrus sp. 1 (6); K- 7010, Cephennomicrus sp. 2 (7); K- 27565, Cephennomicrus sp. 3 (8); K- 8912, Cephennomicrus sp. 4 (9); DU- 32, Cephennomicrus sp. 5 (10); K- 5770, Glaesoconnus unicus sp. n. (11); K- 9579, Rovnoscydmus frontalis sp. n. (12); K- 347, Rovnoscydmus microscopicus sp. n. (13); K- 25702, Rovnoscydmus sp. specimen 1 (14); K- 26348, Rovnoscydmus sp. specimen 2 (15); UA- 1556, Rovnoscydmus sp. specimen 3 (16); K- 8970, Rovnoscydmus sp. specimen 4 (17); K- 8968, Rovnoscydmus sp. specimen 5 (18); K- 7350, Rovnoscydmus sp. specimen 6 (19); K- 3037, Rovnoscydmus sp. specimen 7 (20)." figureDoi="http://doi.org/10.5281/zenodo.262414" httpUri="https://zenodo.org/record/262414/files/figure.png" pageId="18" pageNumber="19">Fig. 2</figureCitation>
), collection number K-243 (
<collectionCode id="ED19AE7A292BFFE3FC26C40A9EA2FEEE" box="[946,1014,295,319]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="18" pageNumber="19">SIZK</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126534292BFFE2FF53C4679E4DFDC9" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8BB736BF292BFFE3FF53C4679945FE57" blockId="18.[151,1436,258,391]" pageId="18" pageNumber="19">
<emphasis id="B97CEAAD292BFFE3FF53C4679C15FEB2" bold="true" box="[199,321,330,355]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
As for genus; additionally pronotum with two pairs of long lateral setae and EI nearly 1.3. 
<emphasis id="B97CEAAD292BFFE3FF53C4439C0DFE56" bold="true" box="[199,345,366,391]" pageId="18" pageNumber="19">Description.</emphasis>
Body (
<figureCitation id="13332A3A292BFFE3FE3EC4439F7FFE56" box="[426,555,366,391]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">Figs 49–55</figureCitation>
) elongate and flatten, brown; BL 
<quantity id="4CF09B5A292BFFE3FC32C4429959FE57" box="[934,1037,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="18" pageNumber="19" unit="mm" value="0.98">0.98 mm</quantity>
.
</paragraph>
<caption id="DF776637292BFFE3FF03C3959CE4F8B9" ID-DOI="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19" targetBox="[151,1436,443,1689]" targetPageId="18">
<paragraph id="8BB736BF292BFFE3FF03C3959CE4F8B9" blockId="18.[151,1436,1720,1896]" pageId="18" pageNumber="19">
<emphasis id="B97CEAAD292BFFE3FF03C3959C01F91F" bold="true" box="[151,341,1720,1743]" pageId="18" pageNumber="19">FIGURES 49–55.</emphasis>
Holotype of 
<taxonomicName id="4C084D3C292BFFE3FE4BC3949FEBF91E" box="[479,703,1721,1743]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="quadrisetosa" status="sp. nov.">
<emphasis id="B97CEAAD292BFFE3FE4BC3949FEBF91E" box="[479,703,1721,1743]" italics="true" pageId="18" pageNumber="19">Vertheia quadrisetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F57D6292BFFE3FD52C3949E54F91F" box="[710,768,1721,1742]" pageId="18" pageNumber="19" rank="species">
<emphasis id="B97CEAAD292BFFE3FD52C3949E54F91F" bold="true" box="[710,768,1721,1742]" pageId="18" pageNumber="19">sp. n.</emphasis>
</taxonomicNameLabel>
(K-27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp3−4, maxillary palpomere III −IV; metaventral intercoxal process; nr, 'neck' region; nss, notosternal suture; occ, occipital constriction; st3−st8; abdominal sternite III −VIII; tm, temple; v3, metaventrite; vt, vertex.
</paragraph>
</caption>
<paragraph id="8BB736BF292BFFE2FF53C2A29F63FF26" blockId="18.[151,1437,1935,2033]" lastBlockId="19.[151,1436,151,573]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Head (
<figureCitation id="13332A3A292BFFE3FE82C2A29CCEF879" box="[278,410,1935,1960]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">Figs 49–54</figureCitation>
) broadest at large eyes, HL 
<quantity id="4CF09B5A292BFFE3FD66C2A29E09F876" box="[754,861,1935,1960]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="mm" value="0.15">0.15 mm</quantity>
, HW 
<quantity id="4CF09B5A292BFFE3FC33C2A29947F876" box="[935,1043,1935,1960]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="mm" value="0.25">0.25 mm</quantity>
; vertex and frons together nearly subquadrate; supraantennal tubercles indistinct. Punctures on head fine and unremarkable, setae only partly visible (
<figureCitation id="13332A3A292BFFE3FF0BC2F59DA2F820" box="[159,246,2008,2033]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="18" pageNumber="19">Fig. 54</figureCitation>
), sparse, short and suberect, except for pair of remarkably long setae, each inserted near posteromesal margin of compound eye (
<figureCitation id="13332A3A292AFFE2FE56C5BA9F4CFF61" box="[450,536,151,176]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 54</figureCitation>
). Antennae (
<figureCitation id="13332A3A292AFFE2FD33C5BA9E7CFF61" box="[679,808,151,176]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Figs 49–54</figureCitation>
) about 0.4 × as long as body, AnL 
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, scape and pedicel strongly elongate, antennomeres III–X slightly to strongly transverse, XI about as long as IX–X together, about as broad as X, with blunt apex.
</paragraph>
<paragraph id="8BB736BF292AFFE2FF53C4289ECAFE7C" blockId="19.[151,1436,151,573]" pageId="19" pageNumber="20">
Pronotum (
<figureCitation id="13332A3A292AFFE2FED3C4299F1AFECC" box="[327,590,260,285]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Figs 49–50, 52, 53–54</figureCitation>
) subrectangular; PL 
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, PW 
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; pronotal disc broadest near anterior third, sides near posterior third indistinctly concave; transverse antebasal groove (
<figureCitation id="13332A3A292AFFE2FB05C40A99B1FE91" box="[1169,1253,295,320]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 53</figureCitation>
; 
<emphasis id="B97CEAAD292AFFE2FB65C40A984FFE91" box="[1265,1307,295,320]" italics="true" pageId="19" pageNumber="20">abg</emphasis>
) seems not connected to lateral antebasal impressions (
<figureCitation id="13332A3A292AFFE2FD15C4619F81FEB4" box="[641,725,332,357]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 53</figureCitation>
; 
<emphasis id="B97CEAAD292AFFE2FD76C4619E52FEB4" box="[738,774,332,357]" italics="true" pageId="19" pageNumber="20">abi</emphasis>
). Each lateral pronotal margin with two long setae located in median region; pronotum with shallow but distinct punctures separated by spaces subequal to two diameters of punctures; setae sparse, moderately long, nearly recumbent (
<figureCitation id="13332A3A292AFFE2FCA9C4B99EC5FE7C" box="[829,913,404,429]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 54</figureCitation>
).
</paragraph>
<paragraph id="8BB736BF292AFFE2FF53C49A9942FE24" blockId="19.[151,1436,151,573]" pageId="19" pageNumber="20">
Elytra (
<figureCitation id="13332A3A292AFFE2FE88C49A9CA1FE01" box="[284,501,439,464]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Figs 49–50, 53–54</figureCitation>
) elongate oval, flattened, EL 
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, EW 
<quantity id="4CF09B5A292AFFE2FC63C495990BFE1E" box="[1015,1119,440,464]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="19" pageNumber="20" unit="mm" value="0.43">0.43 mm</quantity>
, EI 1.29; punctures slightly denser than those on pronotal disc; setae similar to those on pronotum (
<figureCitation id="13332A3A292AFFE2FC21C4F1995EFE24" box="[949,1034,476,501]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 54</figureCitation>
).
</paragraph>
<paragraph id="8BB736BF292AFFE2FF53C72D9E4DFDC9" blockId="19.[151,1436,151,573]" box="[199,793,511,536]" pageId="19" pageNumber="20">
Legs (
<figureCitation id="13332A3A292AFFE2FE99C4D29C0AFDC9" box="[269,350,511,536]" captionStart="FIGURES 49 – 55" captionStartId="18.[151,264,1720,1743]" captionTargetBox="[151,1436,443,1689]" captionTargetId="figure@18.[151,1436,443,1699]" captionTargetPageId="18" captionText="FIGURES 49 – 55. Holotype of Vertheia quadrisetosa sp. n. (K- 27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp 3 − 4, maxillary palpomere III − IV; metaventral intercoxal process; nr, ' neck' region; nss, notosternal suture; occ, occipital constriction; st 3 − st 8; abdominal sternite III − VIII; tm, temple; v 3, metaventrite; vt, vertex." figureDoi="http://doi.org/10.5281/zenodo.262420" httpUri="https://zenodo.org/record/262420/files/figure.png" pageId="19" pageNumber="20">Fig. 55</figureCitation>
) moderately long, slender, unmodified.
</paragraph>
</subSubSection>
<subSubSection id="C3126534292AFFE2FF53C7099986FDED" box="[199,1234,548,573]" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8BB736BF292AFFE2FF53C7099986FDED" blockId="19.[151,1436,151,573]" box="[199,1234,548,573]" pageId="19" pageNumber="20">
<emphasis id="B97CEAAD292AFFE2FF53C7099C1AFDEC" bold="true" box="[199,334,548,573]" pageId="19" pageNumber="20">Etymology.</emphasis>
The name 
<taxonomicName id="4C084D3C292AFFE2FE5FC7099F08FDEC" box="[459,604,548,573]" class="Insecta" family="Staphylinidae" genus="Vertheia" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="quadrisetosa">
<emphasis id="B97CEAAD292AFFE2FE5FC7099F08FDEC" box="[459,604,548,573]" italics="true" pageId="19" pageNumber="20">quadrisetosa</emphasis>
</taxonomicName>
refers to the long paired setae on sides of the pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>