Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber Author Jałoszyński, Paweł Author Perkovsky, Evgeny text Zootaxa 2016 4157 1 1 85 journal article 38320 10.11646/zootaxa.4157.1.1 2389f078-1812-4b4e-826a-561cd7e46654 1175-5326 262413 6BF4514A-892F-499F-BC1E-B7920C7A00B0 † Vertheia quadrisetosa sp. n. ( Figs 2 , 49–55 ) Type material. Holotype : Late Eocene of Europe , Rovno amber: sex unknown; inclusion in elongate, dropshaped piece of amber 14 mm long ( Fig. 2 ), collection number K-243 ( SIZK ). Diagnosis. As for genus; additionally pronotum with two pairs of long lateral setae and EI nearly 1.3. Description. Body ( Figs 49–55 ) elongate and flatten, brown; BL 0.98 mm . FIGURES 49–55. Holotype of Vertheia quadrisetosa sp. n. (K-27418). Habitus in anterodorsal (49), dorsal (50, 53), ventral (51) and anterodorsal (54) views, head and prothorax in dorsal view (52), pterothorax in ventral view (55). Abbreviations: abg, antebasal groove; abi, antebasal impression; bst, basisternal part of prosternum; fr, frons; frp, frontal pit; hy, hypomeron; hyr, hypomeral ridge; msvp, mesoventral intercoxal process; mtvp, mxp3−4, maxillary palpomere III −IV; metaventral intercoxal process; nr, 'neck' region; nss, notosternal suture; occ, occipital constriction; st3−st8; abdominal sternite III −VIII; tm, temple; v3, metaventrite; vt, vertex. Head ( Figs 49–54 ) broadest at large eyes, HL 0.15 mm , HW 0.25 mm ; vertex and frons together nearly subquadrate; supraantennal tubercles indistinct. Punctures on head fine and unremarkable, setae only partly visible ( Fig. 54 ), sparse, short and suberect, except for pair of remarkably long setae, each inserted near posteromesal margin of compound eye ( Fig. 54 ). Antennae ( Figs 49–54 ) about 0.4 × as long as body, AnL 0.40 mm , scape and pedicel strongly elongate, antennomeres III–X slightly to strongly transverse, XI about as long as IX–X together, about as broad as X, with blunt apex. Pronotum ( Figs 49–50, 52, 53–54 ) subrectangular; PL 0.28 mm , PW 0.30 mm ; pronotal disc broadest near anterior third, sides near posterior third indistinctly concave; transverse antebasal groove ( Fig. 53 ; abg ) seems not connected to lateral antebasal impressions ( Fig. 53 ; abi ). Each lateral pronotal margin with two long setae located in median region; pronotum with shallow but distinct punctures separated by spaces subequal to two diameters of punctures; setae sparse, moderately long, nearly recumbent ( Fig. 54 ). Elytra ( Figs 49–50, 53–54 ) elongate oval, flattened, EL 0.55 mm , EW 0.43 mm , EI 1.29; punctures slightly denser than those on pronotal disc; setae similar to those on pronotum ( Fig. 54 ). Legs ( Fig. 55 ) moderately long, slender, unmodified. Etymology. The name quadrisetosa refers to the long paired setae on sides of the pronotum.