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<document ID-CLB-Dataset= "33135" ID-DOI= "http://dx.doi.org/10.3897/jhr.55.11482" ID-GBIF-Dataset= "310c95ad-8bf1-4955-acfb-c1368c0222a7" ID-Pensoft-Pub= "1314-2607-55-139" ID-Pensoft-UUID= "FFF5667EAD5E7A6FFFEC2B6CFFB7D800" ID-ZBK= "0E1B0A14F8714CD89F2910B255677621" ID-Zenodo-Dep= "575140" ID-ZooBank= "0E1B0A14F8714CD89F2910B255677621" ModsDocAuthor= "" ModsDocDate= "2017" ModsDocID= "1314-2607-55-139" ModsDocOrigin= "Journal of Hymenoptera Research 55" ModsDocTitle= "Biting the bullet: revisionary notes on the Oraseminae of the Old World (Hymenoptera, Chalcidoidea, Eucharitidae)" checkinTime= "1493464749952" checkinUser= "pensoft" docAuthor= "Burks, Roger A., Heraty, John M., Mottern, Jason, Dominguez, Chrysalyn & Heacox, Scott" docDate= "2017" docId= "AE1728788E749D5DA71EF101C44C8181" docLanguage= "en" docName= "JourHymenoptRes 55: 139-188" docOrigin= "Journal of Hymenoptera Research 55" docPubDate= "2017-04-28" docSource= "http://dx.doi.org/10.3897/jhr.55.11482" docTitle= "Ivieosema confluens Burks 2017, sp. n." docType= "treatment" docUuid= "DD89153A-F762-43D6-80DE-B823E00EC907" docUuidSource= "ZooBank" docVersion= "6" id= "FFF5667EAD5E7A6FFFEC2B6CFFB7D800" lastPageNumber= "165" masterDocId= "FFF5667EAD5E7A6FFFEC2B6CFFB7D800" masterDocTitle= "Biting the bullet: revisionary notes on the Oraseminae of the Old World (Hymenoptera, Chalcidoidea, Eucharitidae)" masterLastPageNumber= "188" masterPageNumber= "139" pageNumber= "163" updateTime= "1732937005971" updateUser= "ExternalLinkService" >
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<mods:title id= "ACFD154AF33984F477E38824A4D51909" > Biting the bullet: revisionary notes on the Oraseminae of the Old World (Hymenoptera, Chalcidoidea, Eucharitidae)</mods:title>
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<mods:namePart id= "4673C9C389199E61924410C0781A2E14" > Burks, Roger A.</mods:namePart>
<mods:affiliation id= "9BD162D02080646A3E53D10D990F253B" > Department of Entomology, University of California, Riverside, CA 92521</mods:affiliation>
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<mods:namePart id= "9AD7B927C9176361C2FA8EEA1875D420" > Heraty, John M.</mods:namePart>
<mods:affiliation id= "D0B0039A115175532F06F5982F977AB4" > Department of Entomology, University of California, Riverside, CA 92521</mods:affiliation>
<mods:nameIdentifier id= "9C63D94F1E58D129DBF150BB6D74DCFD" type= "email" > john.heraty@ucr.edu</mods:nameIdentifier>
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<mods:namePart id= "4896FF39748E33FC369F7F5DC25F9EF0" > Mottern, Jason</mods:namePart>
<mods:affiliation id= "B5EB335C0013F02F586B872992F2C531" > Department of Entomology, University of California, Riverside, CA 92521 & USDA Systematic Entomology Laboratory, Washington, DC 20013 - 7012</mods:affiliation>
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<mods:namePart id= "527268CA819F118B2B36391C57EB6C81" > Dominguez, Chrysalyn</mods:namePart>
<mods:affiliation id= "4CDE4EB22240655EA01CFBDC648B2B6B" > Department of Entomology, University of California, Riverside, CA 92521</mods:affiliation>
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<mods:namePart id= "89AD12A1EC37E3215599642400242BEF" > Heacox, Scott</mods:namePart>
<mods:affiliation id= "2EAFD8FEB26D6CB831322FD3AA78E40A" > Department of Entomology, University of California, Riverside, CA 92521</mods:affiliation>
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<mods:title id= "E07FFEDE051AA8B8FF4D7BC90F5414E2" > Journal of Hymenoptera Research</mods:title>
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<mods:number id= "7743A39ED1198A9BE26ABF46FF4DAF81" > 55</mods:number>
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<treatment id= "AE1728788E749D5DA71EF101C44C8181" ID-GBIF-Taxon= "128834697" LSID= "urn:lsid:zoobank.org:act:DD89153A-F762-43D6-80DE-B823E00EC907" httpUri= "http://treatment.plazi.org/id/AE1728788E749D5DA71EF101C44C8181" lastPageId= "26" lastPageNumber= "165" pageId= "24" pageNumber= "163" scope_family= "Eucharitidae" scope_order= "Hymenoptera" scope_superFamily= "Chalcidoidea" >
<subSubSection id= "9F3309B32084F484558C1C80FB466D02" pageId= "24" pageNumber= "163" type= "nomenclature" >
<paragraph id= "9373421A82A8998C9BAFA4F360DDF0E1" pageId= "24" pageNumber= "163" >
<taxonomicName id= "3971A9A0ADB6DF3E7E7ABBDC518178F9" LSID= "http://zoobank.org/DD89153A-F762-43D6-80DE-B823E00EC907" authority= "Burks" authorityName= "Burks" authorityYear= "2017" class= "Insecta" family= "Eucharitidae" genus= "Ivieosema" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Ivieosema confluens" order= "Hymenoptera" pageId= "24" pageNumber= "163" phylum= "Arthropoda" rank= "species" species= "confluens" status= "sp. n." > Ivieosema confluens Burks</taxonomicName>
<taxonomicNameLabel id= "5D5D82AF36FDAE5C7EE8765FF5DA92ED" pageId= "24" pageNumber= "163" > sp. n.</taxonomicNameLabel>
<figureCitation id= "F344174D49EC0DA444AE043086ED7D37" captionStart= "Figures 38– captionStartId= "F9" captionText= "Figures 38 - 43. Ivieosema confluens. Paratype female: 38 head and mesosoma, lateral 39 head, anterior 40 antenna, medial 41 mesosoma, lateral 42 mesosoma, dorsal 43 mesosoma, posterior. dfd = dorsal frontal depression; F 1 / an = anellus; lg = longitudinal groove; sod = subocellar depression; tsa = transscutal articulation." figureDoi= "10.3897/jhr.55.11482.figures38-43" httpUri= "https://binary.pensoft.net/fig/134110" pageId= "24" pageNumber= "163" > Figs 38-43</figureCitation>
<figureCitation id= "D2B0C8330BF5D226FC640BFA422529E8" captionStart= "Figures 44– captionStartId= "F10" captionText= "Figures 44 - 48. Ivieosema confluens. Paratype female: 44 fore wing 45 metasoma, lateral, lateral view of ovipositor in top left corner. Holotype male: 46 habitus, lateral 47 head, anterior 48 antennae, dorsal. dv = dorsal (2 nd) valvula, F 1 / an = anellus; vv = ventral (1 st) valvula." figureDoi= "10.3897/jhr.55.11482.figures44-48" httpUri= "https://binary.pensoft.net/fig/134113" pageId= "24" pageNumber= "163" > , 44-48</figureCitation>
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</paragraph>
</subSubSection>
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<subSubSection id= "FD242A17ABD8EAD6EA5043B0B136F664" pageId= "24" pageNumber= "163" type= "etymology" >
<paragraph id= "07D828FD456DC8803A438464D50B2A66" pageId= "24" pageNumber= "163" > Etymology.</paragraph>
<paragraph id= "2D84AA4D75505F05898A60A7E093F752" pageId= "24" pageNumber= "163" > Latin noun meaning " a place where rivers meet," referring to the mesosomal surface sculpture.</paragraph>
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</subSubSection>
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<subSubSection id= "7F19CBD1EA840F15A5408E8B57DAF7DD" pageId= "24" pageNumber= "163" type= "diagnosis" >
<paragraph id= "17FD7CDA41048E3388E29BFAABE53348" pageId= "24" pageNumber= "163" > Diagnosis.</paragraph>
<paragraph id= "7AD08BC1D4C2AB963217E015B2D4F141" pageId= "24" pageNumber= "163" >
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Recognized from other
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<taxonomicName id= "FA866358A1355F38454BB4ECB9147856" authorityName= "Burks & Heraty & Mottern & Dominguez & Heacox" authorityYear= "2017" class= "Insecta" family= "Eucharitidae" genus= "Orasema" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Orasema" order= "Hymenoptera" pageId= "24" pageNumber= "163" phylum= "Arthropoda" rank= "genus" >
<emphasis id= "7C44F2BE021F0804F6456E223717F58D" italics= "true" pageId= "24" pageNumber= "163" > Orasema</emphasis>
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</taxonomicName>
and African
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<taxonomicName id= "8223ADE00C7F5E2D15E624F262C5A006" authorityName= "Heraty & Burks" authorityYear= "2017" class= "Insecta" family= "Eucharitidae" genus= "Ivieosema" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Ivieosema" order= "Hymenoptera" pageId= "24" pageNumber= "163" phylum= "Arthropoda" rank= "genus" >
<emphasis id= "C6CA1C20A635754A1113798EC82BFA7B" italics= "true" pageId= "24" pageNumber= "163" > Ivieosema</emphasis>
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</taxonomicName>
by the fore wing lacking setae and with a medial longitudinal fold. Distinguished from
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<taxonomicName id= "09B9332CF7C7D4B5D30C3A105E8BADE6" lsidName= "I. limulus" pageId= "24" pageNumber= "163" rank= "species" species= "limulus" >
<emphasis id= "02BDF7F46F36216794F3C1E1980CAFD0" italics= "true" pageId= "24" pageNumber= "163" > I. limulus</emphasis>
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</taxonomicName>
by the smooth face, frons with 2 depressions between eye and antennal scrobe (Fig.
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<figureCitation id= "6BDD27B87970DE76FED099EF4CB42525" captionStart= "Figures 38– captionStartId= "F9" captionText= "Figures 38 - 43. Ivieosema confluens. Paratype female: 38 head and mesosoma, lateral 39 head, anterior 40 antenna, medial 41 mesosoma, lateral 42 mesosoma, dorsal 43 mesosoma, posterior. dfd = dorsal frontal depression; F 1 / an = anellus; lg = longitudinal groove; sod = subocellar depression; tsa = transscutal articulation." figureDoi= "10.3897/jhr.55.11482.figures38-43" httpUri= "https://binary.pensoft.net/fig/134110" pageId= "24" pageNumber= "163" > 39</figureCitation>
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), and the basal gastral tergite with a laterally carinate median longitudinal depression extending the equivalent length of the petiole.
</paragraph>
</subSubSection>
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<subSubSection id= "EE4AC6DFF3D6FA7F784DC2B148824C7E" lastPageId= "26" lastPageNumber= "165" pageId= "24" pageNumber= "163" type= "female" >
<paragraph id= "99A449E55C0D13323BF7BFCD5589726C" pageId= "24" pageNumber= "163" > Female.</paragraph>
<paragraph id= "885E4E84442E69754DA753CECB72BC50" pageId= "24" pageNumber= "163" > Length 2.8 mm. Head and mesosoma dark brown. Scape, pedicel, and anellus light brown, rest of flagellum darker brown. Coxae, trochanters, most of femora and last tarsomere dark brown; femoral apices, tibiae and tarsi pale white yellow with last tarsomere brown. Fore wing venation glossy and brown. Gaster brown.</paragraph>
<paragraph id= "607D011AEDD2FA600C3CBEBD6859D2B9" pageId= "24" pageNumber= "163" >
<emphasis id= "B56C5DD3AC51BE4478A7AD7748AA6131" italics= "true" pageId= "24" pageNumber= "163" > Head</emphasis>
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(Figs
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<figureCitation id= "7D556915BF58ED4CE35C9E6E65BCF89B" captionStart= "Figures 38– captionStartId= "F9" captionText= "Figures 38 - 43. Ivieosema confluens. Paratype female: 38 head and mesosoma, lateral 39 head, anterior 40 antenna, medial 41 mesosoma, lateral 42 mesosoma, dorsal 43 mesosoma, posterior. dfd = dorsal frontal depression; F 1 / an = anellus; lg = longitudinal groove; sod = subocellar depression; tsa = transscutal articulation." figureDoi= "10.3897/jhr.55.11482.figures38-43" httpUri= "https://binary.pensoft.net/fig/134110" pageId= "24" pageNumber= "163" > 39-40</figureCitation>
) broadly subtriangular, 1.4
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as broad as long. Face flat and smooth; scrobal depression broad and shallow, extending laterally well beyond toruli and forming a single dorsal frontal depression (Fig.
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<figureCitation id= "CAD7BC7D86B8FBB95C15D0D9A214FD4D" captionStart= "Figures 38– captionStartId= "F9" captionText= "Figures 38 - 43. Ivieosema confluens. Paratype female: 38 head and mesosoma, lateral 39 head, anterior 40 antenna, medial 41 mesosoma, lateral 42 mesosoma, dorsal 43 mesosoma, posterior. dfd = dorsal frontal depression; F 1 / an = anellus; lg = longitudinal groove; sod = subocellar depression; tsa = transscutal articulation." figureDoi= "10.3897/jhr.55.11482.figures38-43" httpUri= "https://binary.pensoft.net/fig/134110" pageId= "24" pageNumber= "163" > 39</figureCitation>
, dfd); lower frons with a vague longitudinal groove; scrobal area with a single medial large pit just below median ocellus; supraclypeal area smooth, clypeus with shallow transverse sculpture, anterior tentorial pits very deep and broad, anteclypeus short laterally and absent medially; scrobal depression shallow and hardly evident; vertex not carinate. Labrum with 4 digits. Maxillary and labial palps absent. Mandibular formula 3:2, mandibles small but falcate. Flagellum 1.0
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head height, 7 funiculars; anellus longer than broad (Fig.
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<figureCitation id= "8D7933353BF8A469DD0CEBAE0B3C9A0A" captionStart= "Figures 38– captionStartId= "F9" captionText= "Figures 38 - 43. Ivieosema confluens. Paratype female: 38 head and mesosoma, lateral 39 head, anterior 40 antenna, medial 41 mesosoma, lateral 42 mesosoma, dorsal 43 mesosoma, posterior. dfd = dorsal frontal depression; F 1 / an = anellus; lg = longitudinal groove; sod = subocellar depression; tsa = transscutal articulation." figureDoi= "10.3897/jhr.55.11482.figures38-43" httpUri= "https://binary.pensoft.net/fig/134110" pageId= "24" pageNumber= "163" > 40</figureCitation>
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,
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<figureCitation id= "D26C40397A10B49A0678A81F2D9B9C21" captionStart= "Figures 1– captionStartId= "F1" captionText= "Figures 1 - 6. Australosema politurae. Paratype female: 1 habitus, lateral 2 head, anterior 3 antennae, ventral 4 mesosoma, dorsal 5 propodeum and petiole, oblique dorsal. Paratype male: 6 antennae, lateral. acl = anteclypeus; acr = acrosternite; acs = antecostal sulcus; apc = anterior petiolar carina; ax = axilla; cal = callus; cly = clypeus; F = flagellomere; fr = frons; Fu = funicular; F 1 / an = anellus; llm = mesoscutal lateral lobe; lpr = lateral propodeal disc; mlm = mesoscutal midlobe; mpr = median propodeal disc; pet = petiole; not = notauli; scd = scutellar disc." figureDoi= "10.3897/jhr.55.11482.figures1-6" httpUri= "https://binary.pensoft.net/fig/134091" pageId= "24" pageNumber= "163" > F1</figureCitation>
); F2 1.7
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as long as broad, 1.1
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as long as F3.
</paragraph>
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<paragraph id= "F39104D99D06EC99D027867BDE8908D0" lastPageId= "26" lastPageNumber= "165" pageId= "24" pageNumber= "163" >
<emphasis id= "1BCA7FEB164995BA755950CC30BC175B" italics= "true" pageId= "24" pageNumber= "163" > Mesosoma</emphasis>
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(Figs
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<figureCitation id= "B567A095C7FBA61E48C9E4FC363DA255" captionStart= "Figures 38– captionStartId= "F9" captionText= "Figures 38 - 43. Ivieosema confluens. Paratype female: 38 head and mesosoma, lateral 39 head, anterior 40 antenna, medial 41 mesosoma, lateral 42 mesosoma, dorsal 43 mesosoma, posterior. dfd = dorsal frontal depression; F 1 / an = anellus; lg = longitudinal groove; sod = subocellar depression; tsa = transscutal articulation." figureDoi= "10.3897/jhr.55.11482.figures38-43" httpUri= "https://binary.pensoft.net/fig/134110" pageId= "24" pageNumber= "163" > 41</figureCitation>
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-
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<figureCitation id= "4F9B79F92F461C7A330128C70800848E" captionStart= "Figures 44– captionStartId= "F10" captionText= "Figures 44 - 48. Ivieosema confluens. Paratype female: 44 fore wing 45 metasoma, lateral, lateral view of ovipositor in top left corner. Holotype male: 46 habitus, lateral 47 head, anterior 48 antennae, dorsal. dv = dorsal (2 nd) valvula, F 1 / an = anellus; vv = ventral (1 st) valvula." figureDoi= "10.3897/jhr.55.11482.figures44-48" httpUri= "https://binary.pensoft.net/fig/134113" pageId= "24" pageNumber= "163" > 44</figureCitation>
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). Mesoscutal midlobe finely reticulate, lateral lobe mostly coriaceous, notauli very shallow and with vague margins; transscutal articulation represented only by a vague depression medially (Fig.
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<figureCitation id= "C9C984F5E100DB42ECDF22EDB89E6412" captionStart= "Figures 38– captionStartId= "F9" captionText= "Figures 38 - 43. Ivieosema confluens. Paratype female: 38 head and mesosoma, lateral 39 head, anterior 40 antenna, medial 41 mesosoma, lateral 42 mesosoma, dorsal 43 mesosoma, posterior. dfd = dorsal frontal depression; F 1 / an = anellus; lg = longitudinal groove; sod = subocellar depression; tsa = transscutal articulation." figureDoi= "10.3897/jhr.55.11482.figures38-43" httpUri= "https://binary.pensoft.net/fig/134110" pageId= "24" pageNumber= "163" > 42</figureCitation>
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, tsa). Axilla anteriorly reticulate, posteriorly longitudinally striate; mesoscutellum and axillula finely longitudinally striate; frenal line as a smooth raised strip. Metascutellum protruding, not separated from rest of metanotum. Mesepisternum reticulate dorsally, smooth ventrally, forming a short rounded horizontal area anterior to mesocoxae; mesepimeron reticulate, without
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<pageBreakToken id= "58FBBDE80C0647143730232ECFD90886" pageId= "25" pageNumber= "164" start= "start" > transepimeral</pageBreakToken>
sulcus. Propodeum finely reticulate, callus bare and smooth to shallowly reticulate. Metacoxa smooth. Fore wing 2.3
<normalizedToken id= "6B60DA90371D2543FB9F431B83766486" originalValue= "× > x</normalizedToken>
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as long as broad; slightly infuscate along cubital fold and posterior to marginal and submarginal veins, and between the oblique folds in the speculum; venation with only a few tiny setae, including on submarginal vein; costal cell bare; basal cell bare; speculum present, with two oblique folds (Fig.
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<figureCitation id= "22A93178449921E5F34CB1190B3DC3A7" captionStart= "Figures 44– captionStartId= "F10" captionText= "Figures 44 - 48. Ivieosema confluens. Paratype female: 44 fore wing 45 metasoma, lateral, lateral view of ovipositor in top left corner. Holotype male: 46 habitus, lateral 47 head, anterior 48 antennae, dorsal. dv = dorsal (2 nd) valvula, F 1 / an = anellus; vv = ventral (1 st) valvula." figureDoi= "10.3897/jhr.55.11482.figures44-48" httpUri= "https://binary.pensoft.net/fig/134113" pageId= "25" pageNumber= "164" > 44</figureCitation>
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, fold) that converge towards stigma and basal fold; marginal vein very thick basally and
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<pageBreakToken id= "596A985E807A0EED2600E114970C2661" pageId= "26" pageNumber= "165" start= "start" > narrowing</pageBreakToken>
apically, not separate from wing margin at base of stigmal vein; stigmal vein 4.5
<normalizedToken id= "A20F598BEEB85EC67211A61533FB86C5" originalValue= "× > x</normalizedToken>
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as long as broad, without an uncus; postmarginal vein absent; cubital and subcubital folds bare; marginal fringe absent. Hind wing costal cell bare.
</paragraph>
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<paragraph id= "04AA4D29EB36DD71786618B768388DFA" pageId= "26" pageNumber= "165" >
<emphasis id= "A0F0CFFC360B4DBCED9C8B511C0C9238" italics= "true" pageId= "26" pageNumber= "165" > Metasoma</emphasis>
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(Fig.
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<figureCitation id= "2D883E9349782377F0C447566AF49E04" captionStart= "Figures 44– captionStartId= "F10" captionText= "Figures 44 - 48. Ivieosema confluens. Paratype female: 44 fore wing 45 metasoma, lateral, lateral view of ovipositor in top left corner. Holotype male: 46 habitus, lateral 47 head, anterior 48 antennae, dorsal. dv = dorsal (2 nd) valvula, F 1 / an = anellus; vv = ventral (1 st) valvula." figureDoi= "10.3897/jhr.55.11482.figures44-48" httpUri= "https://binary.pensoft.net/fig/134113" pageId= "26" pageNumber= "165" > 45</figureCitation>
). Petiole 1.7
<normalizedToken id= "7F15AF4125088BA544B7A2CBFCD09D39" originalValue= "× > x</normalizedToken>
as long as broad, 0.9
<normalizedToken id= "0AE250C5D4CABEDF80BDCD18131B5D1B" originalValue= "× > x</normalizedToken>
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as long as metacoxa, longitudinally shallowly rugose-reticulate. Gt1 with a deep longitudinal median depression with carinate lateral margins equal to length of petiole; shallower smooth depression present on Gt2. Antecostal sulcus of Gs1 smooth, very shallow medially; acrosternite swollen and with shallow irregular sculpture; posterior Gs1 smooth to finely punctate. Cercus with all setae subequal in length. Hypopygium with a few small subapical setae laterally. Ovipositor dorsal valve with 9 annuli, including a basal annulus that is broadly separated from the rest; ventral valve with 5-6 lateral teeth and with transverse carina basal to toothed apex.
</paragraph>
</subSubSection>
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<subSubSection id= "8AAF65D0BB823F4499671E859126B810" pageId= "26" pageNumber= "165" type= "male" >
<paragraph id= "B0E480244EE46DAC7C158ACFE2C4DB48" pageId= "26" pageNumber= "165" > Male</paragraph>
<paragraph id= "7CBE12CA26693305AE1FBB288C934893" pageId= "26" pageNumber= "165" >
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(Figs
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<figureCitation id= "607AB5D762D0C4FBA02B436E486DDDEF" captionStart= "Figures 44– captionStartId= "F10" captionText= "Figures 44 - 48. Ivieosema confluens. Paratype female: 44 fore wing 45 metasoma, lateral, lateral view of ovipositor in top left corner. Holotype male: 46 habitus, lateral 47 head, anterior 48 antennae, dorsal. dv = dorsal (2 nd) valvula, F 1 / an = anellus; vv = ventral (1 st) valvula." figureDoi= "10.3897/jhr.55.11482.figures44-48" httpUri= "https://binary.pensoft.net/fig/134113" pageId= "26" pageNumber= "165" > 46-48</figureCitation>
). Length 2.5 mm. Antennal flagellum densely covered with long decumbent setae, with only a few longitudinal sensilla extending beyond funicular apices; F7 shorter than F6 and F8; entire antenna brown. Petiole 1.3
<normalizedToken id= "67099C4DE87D99748BF8EE1715630B0F" originalValue= "× > x</normalizedToken>
as long as broad, 2.5
<normalizedToken id= "F1D5B1C62B44F0DD363C1695A655EC61" originalValue= "× > x</normalizedToken>
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as long as metacoxa. Gaster with broad, striate longitudinal depression on Gt1. Digitus with 6 spines; parameres present.
</paragraph>
</subSubSection>
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<subSubSection id= "EC9BB3B14A7E0B507D31AE2BF319E42F" pageId= "26" pageNumber= "165" type= "holotype" >
<paragraph id= "D7D82166863CD1EE680F9AA64A592C53" pageId= "26" pageNumber= "165" > Holotype.</paragraph>
<paragraph id= "4F359055634C8D7C5479E322FD507E7E" pageId= "26" pageNumber= "165" >
<emphasis id= "61EBEE0A85F2FD2A2F2518BD94B6832C" bold= "true" pageId= "26" pageNumber= "165" > Madagascar: Toliara Prov.</emphasis>
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: 18 km NNW Betroke, 825m,
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<geoCoordinate id= "80A7FF70FC235E005C46065F5261E064" degrees= "23" direction= "south" minutes= "09" orientation= "latitude" precision= "15" seconds= "48" value= "-23.163334" > 23°09'48" S</geoCoordinate>
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,
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<geoCoordinate id= "1E59AB34744FCF363C19E94FCAC888B9" degrees= "45" direction= "east" minutes= "58" orientation= "longitude" precision= "15" seconds= "07" value= "45.968613" > 45°58'07" E</geoCoordinate>
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, 24.x-4.xii.1994, M.A.Ivie & D.A.Pollock, FIT [1♂, UNMO: UCRCENT00435300], deposited in TMP.
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<emphasis id= "21358C5656657C1551699B82E16CF4FE" bold= "true" pageId= "26" pageNumber= "165" > Paratype: Madagascar: Toliara Prov.</emphasis>
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: Tsimanampetsotsa National Park, Mitoho Forest, 120m,
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<geoCoordinate id= "59D38A82980AB054CEA62DEA8CB3FF9F" degrees= "24" direction= "south" minutes= "02" orientation= "latitude" precision= "15" seconds= "55" value= "-24.04861" > 24°02'55" S</geoCoordinate>
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,
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<geoCoordinate id= "74D29F6041BEB1DD7C7246A698626112" degrees= "43" direction= "east" minutes= "45" orientation= "longitude" precision= "15" seconds= "08" value= "43.752224" > 43°45'08" E</geoCoordinate>
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, 24-29.x.2008, M. Irwin & R.
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<normalizedToken id= "50552F8A40AD0C7BC20B0F06CE9F9008" originalValue= "Harin’ > Harin'Hala</normalizedToken>
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, transitional forest, Malaise trap [1♀, CASC: CASENT2022276].
</paragraph>
</subSubSection>
</treatment>
</document>
