<documentid="65E15225A51C80E7993C7B6F4B0E465A"ID-DOI="10.25674/SO94iSS3iD303"ID-ISSN="2509-9523"ID-Zenodo-Dep="13193962"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1722631614709"checkinUser="felipe"docAuthor="Frisch, Johannes & Mainda, Tobias"docDate="2022"docId="039B5575FF9FDF7D3ACDF9FFFDE2FDC0"docLanguage="en"docName="SoilOrg.94.3.139-147.pdf"docOrigin="Soil Organisms 94 (3)"docSource="http://dx.doi.org/10.25674/so94iss3id303"docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article"docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2"docStyleName="SoilOrg.2018-.journal_article"docStyleVersion="2"docTitle="Scopaeus kokodanus Cameron 1938"docType="treatment"docVersion="2"lastPageNumber="140"masterDocId="FFA22D0DFF9EDF7E3A19FFECFFA6FFF1"masterDocTitle="The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species"masterLastPageNumber="147"masterPageNumber="139"pageNumber="140"updateTime="1722679412379"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:titleid="91EC868CA51562180C0E2CD1BD1567B6">The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species</mods:title>
<mods:affiliationid="E7862654889ECE70944FD5ACDB37C15C">MUSeUM füR NatURKUnDe BeRLin, Leibniz InStitUte fOR EvOLUtiOn anD BiODiveRSity Science,</mods:affiliation>
<figureCitationid="1309F8E6FF9FDF7F3ADDF955FE8CF920"box="[196,298,1721,1745]"captionStart="Figures 1–4"captionStartId="2.[151,224,1930,1952]"captionTargetBox="[275,1286,249,1823]"captionTargetId="figure-24@2.[151,1398,230,1903]"captionTargetPageId="2"captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)."figureDoi="http://doi.org/10.5281/zenodo.13193964"httpUri="https://zenodo.org/record/13193964/files/figure.png"pageId="1"pageNumber="140">Figs 1–4</figureCitation>
. Head slightly elongate to strongly pyriform with maximum width across eyes and tempora evenly narrowed from eyes to neck constriction, head 1.08 – 1.16 times as long as wide, head across eyes 1.17 – 1.42 times as wide as tempora at half of length from eyes to neck constriction. Median labral denticles acute, lateral labral denticles shorter and stouter, with about right-angled apex. Antenna slender with elongate distal segments, penultimate segment 1.1 – 1.14 times as long as wide. Protarsomeres about twice as wide as long. Legs slender, tibia of mesothoracic leg 6.7 – 11.2 times as long as wide. Elytra with well developed shoulders, about 1.1 – 1.23 times as long as pronotum. Metathoracic wings fully developed and functional. Surface of forebody shiny with very fine and spacious setose punctation and without microreticulation. Abdomen finely microreticulate, matter than forebody.
), lateral lobes strongly projecting, each bearing apical and ventral group of long setae; apical lobes short, almost semicircular at apex in dorsal and ventral view (
<figureCitationid="1309F8E6FF9FDF7F39C2FC48FB33FC4D"box="[987,1173,932,956]"captionStart="Figures 17–24"captionStartId="6.[151,224,1885,1907]"captionTargetBox="[222,1362,228,1856]"captionTargetId="figure-24@6.[151,1398,224,1865]"captionTargetPageId="6"captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts."figureDoi="http://doi.org/10.5281/zenodo.13193972"httpUri="https://zenodo.org/record/13193972/files/figure.png"pageId="1"pageNumber="140">Figs 17, 19, 21</figureCitation>
) slightly and broadly emarginate posteriorly and with strong, asetose, median depression covering more than median half of sternite width and all of sternite length, except for strongly sclerotized posterior margin, and being surrounded by a varying number of long, black, medioposteriad pointing setae. Posterior margin of abdominal sternite VIII (
<figureCitationid="1309F8E6FF9FDF7F3F74FB85FC04FB53"captionStart="Figures 17–24"captionStartId="6.[151,224,1885,1907]"captionTargetBox="[222,1362,228,1856]"captionTargetId="figure-24@6.[151,1398,224,1865]"captionTargetPageId="6"captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts."figureDoi="http://doi.org/10.5281/zenodo.13193972"httpUri="https://zenodo.org/record/13193972/files/figure.png"pageId="1"pageNumber="140">Figs 18, 20, 22</figureCitation>
<figureCitationid="1309F8E6FF9FDF7F3E4CFB01FB0AFAF4"box="[1109,1196,1261,1285]"captionStart="Figures 17–24"captionStartId="6.[151,224,1885,1907]"captionTargetBox="[222,1362,228,1856]"captionTargetId="figure-24@6.[151,1398,224,1865]"captionTargetPageId="6"captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts."figureDoi="http://doi.org/10.5281/zenodo.13193972"httpUri="https://zenodo.org/record/13193972/files/figure.png"pageId="1"pageNumber="140">Figs 23</figureCitation>
<bibRefCitationid="EFA39992FF9FDF7F3E75FAE2FACAFAD7"author="Frisch, J. & D. Burckhardt & V. Wolters"box="[1132,1388,1294,1318]"pageId="1"pageNumber="140"pagination="27 - 53"refId="ref4585"refString="Frisch, J., D. Burckhardt & V. Wolters (2002): Rove beetles of the subtribe Scopaeina Coiffait (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity & Evolution 2 (1): 27 - 53. - Electronic Supplement: Synonymic catalog of West Palaearctic Scopaeina including biogeographic information: 1 - 13 [http: // senckenberg. uni-frankfurt. de / odes / 02 - 02. htm]."type="journal article"year="2002">Frisch et al. 2002: 30</bibRefCitation>
<figureCitationid="1309F8E6FF9FDF7F397AFAC2FC10FAB6"box="[867,950,1326,1351]"captionStart="Figures 17–24"captionStartId="6.[151,224,1885,1907]"captionTargetBox="[222,1362,228,1856]"captionTargetId="figure-24@6.[151,1398,224,1865]"captionTargetPageId="6"captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts."figureDoi="http://doi.org/10.5281/zenodo.13193972"httpUri="https://zenodo.org/record/13193972/files/figure.png"pageId="1"pageNumber="140">Figs 24</figureCitation>
–29), but without species group and species specific features. Bursa copulatrix (see
<bibRefCitationid="EFA39992FF9FDF7F3EC7FAA3FCF3FA79"author="Frisch, J. & D. Burckhardt & V. Wolters"pageId="1"pageNumber="140"pagination="27 - 53"refId="ref4585"refString="Frisch, J., D. Burckhardt & V. Wolters (2002): Rove beetles of the subtribe Scopaeina Coiffait (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity & Evolution 2 (1): 27 - 53. - Electronic Supplement: Synonymic catalog of West Palaearctic Scopaeina including biogeographic information: 1 - 13 [http: // senckenberg. uni-frankfurt. de / odes / 02 - 02. htm]."type="journal article"year="2002">Frisch et al. 2002: 30</bibRefCitation>
<bibRefCitationid="EFA39992FF9FDF7F39DBF974FB17F941"author="Frisch, J. & D. Burckhardt & V. Wolters"box="[962,1201,1688,1712]"pageId="1"pageNumber="140"pagination="27 - 53"refId="ref4585"refString="Frisch, J., D. Burckhardt & V. Wolters (2002): Rove beetles of the subtribe Scopaeina Coiffait (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity & Evolution 2 (1): 27 - 53. - Electronic Supplement: Synonymic catalog of West Palaearctic Scopaeina including biogeographic information: 1 - 13 [http: // senckenberg. uni-frankfurt. de / odes / 02 - 02. htm]."type="journal article"year="2002">Frisch et al. (2002: 28)</bibRefCitation>
<taxonomicNameid="4C329FE0FF9FDF7F3EDCF955FA87F920"authorityName="Frisch & Mainda"authorityYear="2022"box="[1221,1313,1721,1745]"class="Insecta"family="Staphylinidae"genus="Scopaeus"kingdom="Animalia"order="Coleoptera"pageId="1"pageNumber="140"phylum="Arthropoda"rank="species"species="balkei">
<figureCitationid="1309F8E6FF9FDF7F39D8F936FBB0F903"box="[961,1046,1754,1778]"captionStart="Figure 25"captionStartId="7.[189,253,1069,1091]"captionTargetBox="[189,801,239,1046]"captionTargetId="figure-632@7.[189,804,236,1046]"captionTargetPageId="7"captionText="Figure 25. Habitat of Scopaeus balkei in the Foja Mountains, Indonesia, Papua Province."figureDoi="http://doi.org/10.5281/zenodo.13193978"httpUri="https://zenodo.org/record/13193978/files/figure.png"pageId="1"pageNumber="140">Fig. 25</figureCitation>
<taxonomicNameid="4C329FE0FF9FDF7F3921F8F0FC32F8C5"authorityName="Frisch & Mainda"authorityYear="2022"box="[824,916,1820,1844]"class="Insecta"family="Staphylinidae"genus="Scopaeus"kingdom="Animalia"order="Coleoptera"pageId="1"pageNumber="140"phylum="Arthropoda"rank="species"species="balkei">
species group is well defined according to the apomorphic characters of the primary and secondary sexual characters described above. The slender body shape of
<taxonomicNameid="4C329FE0FF9DDF7D3B64FED8FD89FEBA"authorityName="Frisch & Mainda"authorityYear="2022"box="[381,559,307,331]"class="Insecta"family="Staphylinidae"genus="Scopaeus"kingdom="Animalia"order="Coleoptera"pageId="3"pageNumber="142"phylum="Arthropoda"rank="species"species="arfakmontium">
with the characteristic pyriform head is, however, widespread in the genus, found in various species groups and thus a convergency. An example is the Australian
<bibRefCitationid="EFA39992FF9DDF7D3B1FFE5BFE29FE3E"author="Frisch, J."box="[262,399,439,463]"pageId="3"pageNumber="142"pagination="55 - 88"refId="ref4544"refString="Frisch, J. (2016): On the Scopaeina Mulsant & Rey of Australasia (Staphylinidae, Paederinae): type revisions and new biogeographic data. - Soil Organisms 88 (1): 55 - 88."type="journal article"year="2016">Frisch 2016</bibRefCitation>
<figureCitationid="1309F8E6FF9DDF7D3BB9FE5BFE40FE3E"box="[416,486,439,463]"captionStart="Figures 1–4"captionStartId="2.[151,224,1930,1952]"captionTargetBox="[275,1286,249,1823]"captionTargetId="figure-24@2.[151,1398,230,1903]"captionTargetPageId="2"captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)."figureDoi="http://doi.org/10.5281/zenodo.13193964"httpUri="https://zenodo.org/record/13193964/files/figure.png"pageId="3"pageNumber="142">Fig. 4</figureCitation>
<bibRefCitationid="EFA39992FF9DDF7D3B13FDF5FE2AFDC0"author="Frisch, J."box="[266,396,537,561]"pageId="3"pageNumber="142"pagination="55 - 88"refId="ref4544"refString="Frisch, J. (2016): On the Scopaeina Mulsant & Rey of Australasia (Staphylinidae, Paederinae): type revisions and new biogeographic data. - Soil Organisms 88 (1): 55 - 88."type="journal article"year="2016">Frisch 2016</bibRefCitation>
<figureCitationid="1309F8E6FF9DDF7D3B8CFDF5FDB6FDC3"box="[405,528,537,562]"captionStart="Figures 17–24"captionStartId="6.[151,224,1885,1907]"captionTargetBox="[222,1362,228,1856]"captionTargetId="figure-24@6.[151,1398,224,1865]"captionTargetPageId="6"captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts."figureDoi="http://doi.org/10.5281/zenodo.13193972"httpUri="https://zenodo.org/record/13193972/files/figure.png"pageId="3"pageNumber="142">Figs 20–22</figureCitation>
