<documentid="E9AF779A743593CD088A6E2B0D5A10CA"ID-CLB-Dataset="302934"ID-DOI="10.11646/zootaxa.5506.4.6"ID-GBIF-Dataset="d61b3502-9ddf-4c07-8264-257dccfd0f59"ID-ISSN="1175-5326"ID-Zenodo-Dep="13747461"ID-ZooBank="933199B2-0F15-435D-BFB4-E8C3710281C9"IM.bibliography_approvedBy="valdenar"IM.illustrations_approvedBy="valdenar"IM.materialsCitations_approvedBy="valdenar"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="valdenar"checkinTime="1726055156787"checkinUser="plazi"docAuthor="Naydenov, Artem E., Yakovlev, Roman V. & Penco, Fernando C."docDate="2024"docId="214D180CFFB0F553B4C0FB25FE18DF33"docLanguage="en"docName="zootaxa.5506.4.6.pdf"docOrigin="Zootaxa 5506 (4)"docSource="http://dx.doi.org/10.11646/zootaxa.5506.4.6"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Givira Walker 1856"docType="treatment"docVersion="4"lastPageNumber="569"masterDocId="DD746074FFB1F551B457FFEBFFC7D87F"masterDocTitle="Review of Givira Walker, 1856 (Lepidoptera: Cossidae: Hypoptinae) with description of a new species"masterLastPageNumber="576"masterPageNumber="567"pageNumber="568"updateTime="1726081912127"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="70AC0BAC9F36DB631716EE760E403F13">Review of Givira Walker, 1856 (Lepidoptera: Cossidae: Hypoptinae) with description of a new species</mods:title>
<mods:namePartid="08C4F079D2E5933834D34F8C462C6C83">Yakovlev, Roman V.</mods:namePart>
<mods:affiliationid="CCD9BCFB703C48BDB3A3FFDDF86B72A8">Altai State University, pr. Lenina 61, Barnaul 656049, Russia. & Tomsk State University, Laboratory of Biodiversity and Ecology, Lenin pr. 36, 634050 Tomsk, Russia. & Western Caspian University, 31, Istiglaliyyat Street, Baku, Azerbaijan.</mods:affiliation>
<mods:namePartid="8C902E66FFDFD4F6EB312CA1B17F4E2E">Penco, Fernando C.</mods:namePart>
<mods:affiliationid="92DEE08111EA48A403AD5A73BEDFBB35">Fundación de Historia Natural “ Félix de Azara ”, Departamento de Ciencias Naturales y Antropología, Universidad Maimónides, Hidalgo 775 piso 7 (1405 BDB) Ciudad Autónoma de Buenos Aires, Argentina.</mods:affiliation>
<bibRefCitationid="CD75D4EBFFB0F550B515FB24FE21DC97"author="Walker, F."box="[322,486,1230,1257]"pageId="1"pageNumber="568"pagination="1509 - 1808"refId="ref6274"refString="Walker, F. (1856) n. k. In: List of the specimens of lepidopterous insects in the collection of the British Museum. Part VII. Lepidoptera Heterocera. British Museum (Natural History), London, pp. 1509 - 1808."type="book chapter"year="1856">Walker, 1856</bibRefCitation>
<bibRefCitationid="CD75D4EBFFB0F550B4C0FAF8FE65DD54"author="Walker, F."box="[151,418,1299,1323]"pageId="1"pageNumber="568"pagination="1509 - 1808"refId="ref6274"refString="Walker, F. (1856) n. k. In: List of the specimens of lepidopterous insects in the collection of the British Museum. Part VII. Lepidoptera Heterocera. British Museum (Natural History), London, pp. 1509 - 1808."type="book chapter"year="1856">Walker, 1856: 1526−1527</bibRefCitation>
<bibRefCitationid="CD75D4EBFFB0F550B736FAF8FC2FDD54"author="Walker, F."box="[865,1000,1299,1323]"pageId="1"pageNumber="568"pagination="1509 - 1808"refId="ref6274"refString="Walker, F. (1856) n. k. In: List of the specimens of lepidopterous insects in the collection of the British Museum. Part VII. Lepidoptera Heterocera. British Museum (Natural History), London, pp. 1509 - 1808."type="book chapter"year="1856">Walker, 1856</bibRefCitation>
</taxonomicName>
(
<figureCitationid="31DFB59FFFB0F550B7A2FAF8FBF1DD54"box="[1013,1078,1299,1323]"captionStart="FIGURES 1–7"captionStartId="3.[151,264,1007,1031]"captionTargetBox="[151,1435,176,984]"captionTargetId="figure-17@3.[151,1435,176,984]"captionTargetPageId="3"captionText="FIGURES 1–7. Givira, adults: 1. G. cinnamonea Naydenov, Yakovlev & Penco sp. nov., male, holotype, Brazil, Minas Gerais (MWM); 2. G. difflua Dognin, 1920, male, holotype, Argentina, La Rioja (USNM); 3. G. difflua Dognin, 1920, male, Argentina, Jujuy Prov., Santa Barbara Mts. (RYB); 4. G. difflua Dognin, 1920, female, Argentina, Jujuy Prov., Santa Barbara Mts. (RYB); 5. G. modisma Schaus, 1922, male, holotype, Guatemala, Quirigua (USNM); 6. G. modisma Schaus, 1922, male, Colombia, Huila (RYB); 7. G. tristis Walker, 1856, male, holotype (NHMUK)."figureDoi="http://doi.org/10.5281/zenodo.13747463"httpUri="https://zenodo.org/record/13747463/files/figure.png"pageId="1"pageNumber="568">Figs 7</figureCitation>
,
<figureCitationid="31DFB59FFFB0F550B016FAF8FB9CDD54"box="[1089,1115,1299,1323]"captionStart="FIGURES 8–11"captionStartId="5.[151,264,1837,1861]"captionTargetBox="[153,1433,594,1812]"captionTargetId="figure-148@5.[151,1435,592,1814]"captionTargetPageId="5"captionText="FIGURES 8–11. Givira, male genitalia: 8. G. cinnamonea Naydenov, Yakovlev & Penco sp. nov., male, holotype, Brazil, Minas Gerais, GenPrMWM: 28.562 (MWM); 9. G. difflua Dognin, 1920, male, holotype, Argentina, La Rioja, genitalia slide by P. Gentili №85.229 (USNM); 10. G. modisma Schaus, 1922, male, holotype, Guatemala, Quirigua, genitalia slide by P. Gentili №85.253 (USNM); 11. G. tristis Walker, 1856, male, holotype, genitalia slide №125 (NHMUK)."figureDoi="http://doi.org/10.5281/zenodo.13747465"httpUri="https://zenodo.org/record/13747465/files/figure.png"pageId="1"pageNumber="568">11</figureCitation>
<figureCitationid="31DFB59FFFB0F550B031FAF8FB45DD54"box="[1126,1154,1299,1323]"captionStart="FIGURE 13"captionStartId="7.[151,250,1571,1595]"captionTargetId="figure-181@7.[366,1222,684,1548]"captionTargetPageId="7"captionText="FIGURE 13. Distributional records of Givira Walker, 1856."figureDoi="http://doi.org/10.5281/zenodo.13750667"httpUri="https://zenodo.org/record/13750667/files/figure.png"pageId="1"pageNumber="568">13</figureCitation>
; antennae brown, relatively short (not longer than 1/3 of fore wing in length), bipectinate, with short rami; head, thorax and abdomen covered with brown, dark-grey and light-grey scales; fore wings mostly brown (with brownish or beige tint) with poorly pronounced wavy pattern, usually more contrasting on costal edge. Hind wing relatively short, light colored (rarely, with darkened distal half) with poorly pronounced wavy pattern; darkening at vein A
. Uncus relatively long with small oval or round extension and small incisure at top; arms of gnathos short or completely reduced; gnathos absent; valva with simple harpe; juxta robust, lamellar, with ring-like aperture and massive hood-shaped abdominal process; saccus small; phallus equal to valva in length or shorter, thin, straight or slightly curved, several specimens have small spiky process at distal tip.
. Ovipositor short; papillae anales slightly acute, with small bundles of setae; anterior and posterior apophyses relatively short, approximately equal in length; ostium cup-like, sclerotized, with well-developed lamella postvaginalis; ductus relatively long and thin; smoothly passing into bursa small, oval, has several signa.
<taxonomicNameid="6EE4D299FFB0F550B765F8C6FC72DF38"authorityName="Neumoegen & Dyar"authorityYear="1894"box="[818,949,1837,1863]"class="Insecta"family="Cossidae"kingdom="Animalia"order="Lepidoptera"pageId="1"pageNumber="568"phylum="Arthropoda"rank="subFamily"subFamily="Hypoptinae">Hypoptinae</taxonomicName>
—absence of processes on valvae, deeply forked uncus, pronounced long saccus, big spiky processes on phallus and sharp transition between ductus and bursa from
<bibRefCitationid="CD75D4EBFFB0F550B4AFF872FE63DFCC"author="Edwards, H."box="[248,420,1945,1971]"pageId="1"pageNumber="568"pagination="181 - 185"refId="ref5293"refString="Edwards, H. (1888) New Genera and Species of North American Moths. Entomologica Americana, 3, 181 - 185."type="journal article"year="1888">Edwards, 1888</bibRefCitation>
<bibRefCitationid="CD75D4EBFFB0F550B64FF871FD52DFCC"author="Dyar, H. G."box="[536,661,1945,1972]"pageId="1"pageNumber="568"pagination="1265 - 1285"refId="ref5250"refString="Dyar, H. G. (1940) Cossidae. In: Seitz, A. (Ed.), Die exotischen Grossschmetterlinge, Die Amerikanischen Spinner und Schwarmer. Alfred Kernen, Stuttgart, pp. 1265 - 1285."type="book chapter"year="1940">Dyar, 1940</bibRefCitation>
<bibRefCitationid="CD75D4EBFFB0F550B159F871FA48DFCC"author="Dyar, H. G."box="[1294,1423,1945,1972]"pageId="1"pageNumber="568"pagination="1265 - 1285"refId="ref5250"refString="Dyar, H. G. (1940) Cossidae. In: Seitz, A. (Ed.), Die exotischen Grossschmetterlinge, Die Amerikanischen Spinner und Schwarmer. Alfred Kernen, Stuttgart, pp. 1265 - 1285."type="book chapter"year="1940">Dyar, 1940</bibRefCitation>
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);
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<bibRefCitationid="CD75D4EBFFB3F553B7E2FEA0FA96D91A"author="Yakovlev, R. V. & Naydenov, A. E. & Penco, F. C."box="[949,1361,331,357]"pageId="2"pageNumber="569"pagination="4 - 13"refId="ref6323"refString="Yakovlev, R. V., Naydenov, A. E. & Penco, F. C. (2019) New data on Neotropical Carpenter-Moths of Subfamily Hypoptinae Neumoegen & Dyar, 1894 (Lepidoptera: Cossidae). I. New taxa with bifurcated uncus. Ecologica Montenegrina, 26, 4 - 13."type="journal article"year="2019">Yakovlev, Naydenov & Penco, 2019</bibRefCitation>
<bibRefCitationid="CD75D4EBFFB3F553B768FE78FC14D9D2"author="Schaus, W."box="[831,979,403,429]"pageId="2"pageNumber="569"pagination="612 - 634"refId="ref6044"refString="Schaus, W. (1911) New Species of Heterocera from Costa Rica. VIII. Annals and Magazine of Natural History, Series 8, 7 (42), 612 - 634. https: // doi. org / 10.1080 / 00222931108692985"type="journal article"year="1911">Schaus, 1911</bibRefCitation>
<bibRefCitationid="CD75D4EBFFB3F553B4AAFE5CFE52D9AE"author="Schaus, W."box="[253,405,439,465]"pageId="2"pageNumber="569"pagination="612 - 634"refId="ref6044"refString="Schaus, W. (1911) New Species of Heterocera from Costa Rica. VIII. Annals and Magazine of Natural History, Series 8, 7 (42), 612 - 634. https: // doi. org / 10.1080 / 00222931108692985"type="journal article"year="1911">Schaus, 1911</bibRefCitation>
—shape of wings and absence of spots in wing pattern, presence of oval extension on tip of uncus, longer valvae and absence of distinctly pronounced saccus from
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<bibRefCitationid="CD75D4EBFFB3F553B06FFD80FA9CDAFA"author="Naydenov, A. E. & Yakovlev, R. V. & Penco, F. C. & Sinyaev, V. V."box="[1080,1371,619,645]"pageId="2"pageNumber="569"pagination="82 - 114"refId="ref5515"refString="Naydenov, A. E., Yakovlev, R. V., Penco, F. C. & Sinyaev, V. V. (2020) New data on Neotropical Carpenter-Moths of Subfamily Hypoptinae Neumoegen & Dyar, 1894 (Lepidoptera: Cossi- dae). II. A review of the genus Dolecta Herrich-Schaffer, [1854], with description of seventeen new species. Ecologica Montenegrina, 35, 82 - 114. https: // doi. org / 10.37828 / em. 2020.35.7"type="book chapter"year="2020">Naydenov & Penco, 2020</bibRefCitation>
—shorter phallus and uncus, poorly developed saccus and small apical incisure on uncus from
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<bibRefCitationid="CD75D4EBFFB3F553B52BFCA8FDF2DB22"author="Naydenov, A. E. & Yakovlev, R. V. & Penco, F. C. & Sinyaev, V. V."box="[380,565,835,861]"pageId="2"pageNumber="569"pagination="82 - 114"refId="ref5515"refString="Naydenov, A. E., Yakovlev, R. V., Penco, F. C. & Sinyaev, V. V. (2020) New data on Neotropical Carpenter-Moths of Subfamily Hypoptinae Neumoegen & Dyar, 1894 (Lepidoptera: Cossi- dae). II. A review of the genus Dolecta Herrich-Schaffer, [1854], with description of seventeen new species. Ecologica Montenegrina, 35, 82 - 114. https: // doi. org / 10.37828 / em. 2020.35.7"type="book chapter"year="2020">Naydenov, 2020</bibRefCitation>
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<bibRefCitationid="CD75D4EBFFB3F553B4C0FC8CFE94DBFE"author="Naydenov, A. E. & Yakovlev, R. V. & Penco, F. C. & Sinyaev, V. V."box="[151,339,871,897]"pageId="2"pageNumber="569"pagination="82 - 114"refId="ref5515"refString="Naydenov, A. E., Yakovlev, R. V., Penco, F. C. & Sinyaev, V. V. (2020) New data on Neotropical Carpenter-Moths of Subfamily Hypoptinae Neumoegen & Dyar, 1894 (Lepidoptera: Cossi- dae). II. A review of the genus Dolecta Herrich-Schaffer, [1854], with description of seventeen new species. Ecologica Montenegrina, 35, 82 - 114. https: // doi. org / 10.37828 / em. 2020.35.7"type="book chapter"year="2020">Naydenov, 2020</bibRefCitation>
—absence of bright decorative pattern, robust saccus and processes on the valvae from
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<bibRefCitationid="CD75D4EBFFB3F553B57BFC1CFE71DC6E"author="Penco, F. C. & Yakovlev, R. V. & Naydenov, A. E."box="[300,438,1015,1041]"pageId="2"pageNumber="569"pagination="53 - 63"refId="ref5925"refString="Penco, F. C., Yakovlev, R. V. & Naydenov, A. E. (2022) New data on Neotropical Carpenter-Moths of Subfamily Hypoptinae Neumoegen & Dyar, 1894 (Lepidoptera: Cossidae). IV. A review of the genus Hypopta Hubner, 1818. Ecologica Montenegrina, 51, 53 - 63. https: // doi. org / 10.37828 / em. 2022.51.6"type="journal article"year="2022">Penco, 2022</bibRefCitation>
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<bibRefCitationid="CD75D4EBFFB3F553B106FC1CFF13DC4A"author="Penco, F. C. & Yakovlev, R. V. & Naydenov, A. E."pageId="2"pageNumber="569"pagination="53 - 63"refId="ref5925"refString="Penco, F. C., Yakovlev, R. V. & Naydenov, A. E. (2022) New data on Neotropical Carpenter-Moths of Subfamily Hypoptinae Neumoegen & Dyar, 1894 (Lepidoptera: Cossidae). IV. A review of the genus Hypopta Hubner, 1818. Ecologica Montenegrina, 51, 53 - 63. https: // doi. org / 10.37828 / em. 2022.51.6"type="journal article"year="2022">Penco, 2022</bibRefCitation>
—light hind wings, absence of pronounced transverse light stripes, presence of more robust tegumen and uncus, special shape of valvae and phallus and apical small incisure on uncus from
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<bibRefCitationid="CD75D4EBFFB3F553B106FB88FF08DCDE"author="Penco, F. C. & Yakovlev, R. V. & Naydenov, A. E."pageId="2"pageNumber="569"pagination="53 - 63"refId="ref5925"refString="Penco, F. C., Yakovlev, R. V. & Naydenov, A. E. (2022) New data on Neotropical Carpenter-Moths of Subfamily Hypoptinae Neumoegen & Dyar, 1894 (Lepidoptera: Cossidae). IV. A review of the genus Hypopta Hubner, 1818. Ecologica Montenegrina, 51, 53 - 63. https: // doi. org / 10.37828 / em. 2022.51.6"type="journal article"year="2022">Penco, 2022</bibRefCitation>
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<bibRefCitationid="CD75D4EBFFB3F553B784FB6CFBA5DCDE"author="Penco, F. C. & Yakovlev, R. V. & Naydenov, A. E."box="[979,1122,1159,1185]"pageId="2"pageNumber="569"pagination="53 - 63"refId="ref5925"refString="Penco, F. C., Yakovlev, R. V. & Naydenov, A. E. (2022) New data on Neotropical Carpenter-Moths of Subfamily Hypoptinae Neumoegen & Dyar, 1894 (Lepidoptera: Cossidae). IV. A review of the genus Hypopta Hubner, 1818. Ecologica Montenegrina, 51, 53 - 63. https: // doi. org / 10.37828 / em. 2022.51.6"type="journal article"year="2022">Penco, 2022</bibRefCitation>
—special wing pattern without light-colored spots on fore wing basally and in light-colored hind wings; absence of needle-like basal processes on valvae; robust saccus; special shape of juxta and small apical incisure on uncus from