Review of Givira Walker, 1856 (Lepidoptera: Cossidae: Hypoptinae) with description of a new species
Author
Naydenov, Artem E.
Altai State University, pr. Lenina 61, Barnaul 656049, Russia.
Author
Yakovlev, Roman V.
Altai State University, pr. Lenina 61, Barnaul 656049, Russia. & Tomsk State University, Laboratory of Biodiversity and Ecology, Lenin pr. 36, 634050 Tomsk, Russia. & Western Caspian University, 31, Istiglaliyyat Street, Baku, Azerbaijan.
Author
Penco, Fernando C.
Fundación de Historia Natural “ Félix de Azara ”, Departamento de Ciencias Naturales y Antropología, Universidad Maimónides, Hidalgo 775 piso 7 (1405 BDB) Ciudad Autónoma de Buenos Aires, Argentina.
text
Zootaxa
2024
2024-09-10
5506
4
567
576
http://dx.doi.org/10.11646/zootaxa.5506.4.6
journal article
302934
10.11646/zootaxa.5506.4.6
d61b3502-9ddf-4c07-8264-257dccfd0f59
1175-5326
13747461
933199B2-0F15-435D-BFB4-E8C3710281C9
Genus
Givira
Walker, 1856
Walker, 1856: 1526−1527
.
Type
species (by monotypy):
Givira tristis
Walker, 1856
(
Figs 7
,
11
,
13
).
Description
. Size medium; length of male fore wing
12‒17 mm
; length of female fore wing
16‒19 mm
; antennae brown, relatively short (not longer than 1/3 of fore wing in length), bipectinate, with short rami; head, thorax and abdomen covered with brown, dark-grey and light-grey scales; fore wings mostly brown (with brownish or beige tint) with poorly pronounced wavy pattern, usually more contrasting on costal edge. Hind wing relatively short, light colored (rarely, with darkened distal half) with poorly pronounced wavy pattern; darkening at vein A
1 in
apical third (in some specimens, darkening noticeable only at wing margin on top of A1).
Male genitalia
. Uncus relatively long with small oval or round extension and small incisure at top; arms of gnathos short or completely reduced; gnathos absent; valva with simple harpe; juxta robust, lamellar, with ring-like aperture and massive hood-shaped abdominal process; saccus small; phallus equal to valva in length or shorter, thin, straight or slightly curved, several specimens have small spiky process at distal tip.
Female genitalia
. Ovipositor short; papillae anales slightly acute, with small bundles of setae; anterior and posterior apophyses relatively short, approximately equal in length; ostium cup-like, sclerotized, with well-developed lamella postvaginalis; ductus relatively long and thin; smoothly passing into bursa small, oval, has several signa.
Diagnosis
.
Givira
differs from the following genera of
Hypoptinae
by:
—absence of processes on valvae, deeply forked uncus, pronounced long saccus, big spiky processes on phallus and sharp transition between ductus and bursa from
Inguromorpha
(
type
species by monotypy:
Inguromorpha slossoni
Edwards, 1888
);
Puseyia
Dyar, 1940
(
type
species by original designation:
Puseyia puseyiae
Dyar, 1940
);
Uretiana
Yakovlev, Naydenov & Penco, 2019
(
type
species by original designation:
Givira vicunensis
Ureta, 1957
);
Dogniniya
Yakovlev, Naydenov & Penco, 2019
(
type
species by original designation:
Hypopta sterila
Dognin, 1910
);
Qhichwaruna
Yakovlev, Naydenov & Penco, 2019
(
type
species by monotypy:
Qhichwaruna pennata
Yakovlev, Naydenov & Penco, 2019
);
Wiraqucha
Yakovlev, Naydenov & Penco, 2019
(
type
species by original designation:
Wiraqucha equadorica
Yakovlev, Naydenov & Penco, 2019
);
Thonyocossus
Yakovlev, Naydenov & Penco, 2019
(
type
species by monotypy:
Thonyocossus brasiliensis
Yakovlev, Naydenov & Penco, 2019
); and
Hastam
Yakovlev, Naydenov & Penco, 2019
(
type
species by monotypy:
Hastam gracilis
Yakovlev, Naydenov & Penco, 2019
);
—absence of deeply forked uncus and pronounced long saccus from
Dolecta
(
type
species by monotypy:
Dolecta scariosa
Herrich-Schäffer, [1854]
) and
Psychogena
Schaus, 1911
(
type
species by monotypy:
Psychogena miranda
Schaus, 1911
);
—shape of wings and absence of spots in wing pattern, presence of oval extension on tip of uncus, longer valvae and absence of distinctly pronounced saccus from
Clenchgivira
Heppner, 2023 (
type
species by original designation:
Clenchgivira nigrihyalina
Heppner, 2023);
—absence of long basal processes on valvae and presence of small apical incisure on uncus from
Hypopta
(
type
species by monotypy:
Hypopta ambigua
Hübner, 1818
);
Laberlia
Yakovlev,
Naydenov & Penco, 2020
(
type
species by original designation:
Langsdorfia bellaria
Dognin, 1911
); and
Philanglaus
Butler, 1882
(
type
species by monotypy:
Philanglaus ornatus
Butler, 1882
);
—thinner uncus with a small apical incisure, poorly visible saccus and special shape of juxta from
Schausisca
Gentili, 1989
(
type
species by original designation:
Schausisca desantisi
Gentili, 1989
);
—shorter phallus and uncus, poorly developed saccus and small apical incisure on uncus from
Schajovskoia
Penco, Yakovlev &
Naydenov, 2020
(
type
species by monotypy:
Schajovskoia albomaculata
Penco, Yakovlev &
Naydenov, 2020
);
—absence of heavily sclerotized semicircular portion of juxta and presence of small apical incisure on uncus from
Philiodoron
Clench, 1957
(
type
species by original designation:
Philiodoron cinereum
Clench, 1957
);
—absence of bright decorative pattern, robust saccus and processes on the valvae from
Gentiliocossus
Yakovlev, Naydenov &
Penco, 2022
(
type
species by original designation:
Gentiliocossus alexi
Yakovlev, Naydenov &
Penco, 2022
);
—light hind wings, absence of pronounced transverse light stripes, presence of more robust tegumen and uncus, special shape of valvae and phallus and apical small incisure on uncus from
Givirella
Naydenov, Yakovlev &
Penco, 2022
(
type
species by monotypy:
Givirella batesi
Naydenov, Yakovlev &
Penco, 2022
);
—special wing pattern without spots in apical half of fore wing, rod-like uncus, shape of valvae and small apical incisure on uncus from
Givarbela
Clench, [1957]
(
type
species by monotypy:
Givarbela steinbachi
Clench, [1957]
);
—color and wing pattern without longitudinal stripes parallel to veins, special shape of valvae and juxta, and small apical incisure on uncus from
Breyeriana
Orfila, 1957
(
type
species by monotypy:
Breyeriana cistransandina
Orfila, 1957
);
—bigger size, absence of large setae on antennae, thinner uncus with small incisure on apex and special shape of valvae and juxta from
Acousmaticus
Butler, 1882
(
type
species by monotypy:
Acousmaticus magnicornis
Butler, 1882
);
—special wing pattern without light-colored spots on fore wing basally and in light-colored hind wings; absence of needle-like basal processes on valvae; robust saccus; special shape of juxta and small apical incisure on uncus from
Langsdorfia
Hübner, [1821]
(
type
species by monotypy:
Langsdorfia franckii
Hübner, [1821]
).
Distribution
. Central America (
Guatemala
) and South America (
Colombia
,
Venezuela
,
Brazil
,
Argentina
,
Paraguay
).
Taxonomic note
. Below we review only four species that are surely belong to
Givira
, others should be considered as
Givira auct
.
until reattributed.
Description of new species