treatments-xml/data/03/E0/87/03E087DFBF4CFFFDFF5C5C85FAE0F121.xml

<document id="544A57F69B30947286CC1AEADD1FEC7B" ID-CLB-Dataset="43001" ID-DOI="10.11646/zootaxa.3717.4.5" ID-GBIF-Dataset="16d0cbc8-dd2c-4a41-8ab3-4649778bcee0" ID-ISSN="1175-5326" ID-Zenodo-Dep="249082" ID-ZooBank="7AC6654C-0D33-4921-B89D-807ADE0FF438" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" checkinTime="1460684842358" checkinUser="plazi" docAuthor="Wanat, Marek" docDate="2013" docId="03E087DFBF4CFFFDFF5C5C85FAE0F121" docLanguage="en" docName="zt03717p542.pdf" docOrigin="Zootaxa 3717 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archellerenius giustoi Wanat, 2013, sp. nov." docType="treatment" docVersion="12" lastPageNumber="518" masterDocId="FFD9FFA7BF4FFFFEFFCB5C12FFC6F66C" masterDocTitle="The Apionidae (Coleoptera: Curculionoidea) of New Caledonia. Genera Archellerenius gen. nov. and Hellerenius Wanat" masterLastPageNumber="542" masterPageNumber="515" pageNumber="518" updateTime="1728639270943" updateUser="ExternalLinkService">
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<mods:title id="89C6C190E2B6ABD9A0CC7A8C30444917">The Apionidae (Coleoptera: Curculionoidea) of New Caledonia. Genera Archellerenius gen. nov. and Hellerenius Wanat</mods:title>
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<mods:namePart id="BFBB538D505F75CDAC3E5242D23A267F">Wanat, Marek</mods:namePart>
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<mods:date id="CFC44854F4FA3D91B1F20EF1D21CEBCE">2013</mods:date>
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<mods:identifier id="481939D55975468F19EF5C54003392A2" type="DOI">10.11646/zootaxa.3717.4.5</mods:identifier>
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<treatment id="03E087DFBF4CFFFDFF5C5C85FAE0F121" ID-DOI="http://doi.org/10.5281/zenodo.6153098" ID-GBIF-Taxon="119578258" ID-Zenodo-Dep="6153098" LSID="urn:lsid:plazi:treatment:03E087DFBF4CFFFDFF5C5C85FAE0F121" httpUri="http://treatment.plazi.org/id/03E087DFBF4CFFFDFF5C5C85FAE0F121" lastPageNumber="518" pageId="3" pageNumber="518">
<subSubSection id="C3536542BF4CFFFDFF5C5C85FEF2F6BF" pageId="3" pageNumber="518" type="nomenclature">
<paragraph id="8BF636C9BF4CFFFDFF5C5C85FDCEF6DE" blockId="3.[151,520,151,211]" box="[151,520,151,178]" pageId="3" pageNumber="518">
<heading id="D0BE81A5BF4CFFFDFF5C5C85FDCEF6DE" bold="true" box="[151,520,151,178]" fontSize="11" level="1" pageId="3" pageNumber="518" reason="1">
<emphasis id="B93DEADBBF4CFFFDFF5C5C85FDCEF6DE" bold="true" box="[151,520,151,178]" pageId="3" pageNumber="518">
<taxonomicName id="4C494D4ABF4CFFFDFF5C5C85FE64F6DD" authorityName="Wanat" authorityYear="2013" box="[151,418,151,177]" class="Insecta" family="Brentidae" genus="Archellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="species" species="giustoi" status="sp. nov.">
<emphasis id="B93DEADBBF4CFFFDFF5C5C85FE64F6DD" bold="true" box="[151,418,151,177]" italics="true" pageId="3" pageNumber="518">Archellerenius giustoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57A0BF4CFFFDFE615C8AFDCEF6DE" box="[426,520,152,178]" pageId="3" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF5C5CA8FEF2F6BF" blockId="3.[151,520,151,211]" box="[151,308,186,211]" pageId="3" pageNumber="518">
(
<figureCitation id="13722A4CBF4CFFFDFF545CA8FF21F6BF" box="[159,231,186,211]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." figureDoi="http://doi.org/10.5281/zenodo.249083" httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="3" pageNumber="518">Figs 3</figureCitation>
, 
<figureCitation id="13722A4CBF4CFFFDFF385CA9FEEAF6BF" box="[243,300,187,211]" captionStart-0="FIGURES 9 – 20" captionStart-1="FIGURES 21 – 32" captionStart-2="FIGURES 33 – 45" captionStartId-0="6.[151,264,1953,1976]" captionStartId-1="7.[151,264,1850,1873]" captionStartId-2="8.[151,264,1923,1946]" captionTargetBox-0="[151,1431,193,1930]" captionTargetBox-1="[191,1396,200,1822]" captionTargetBox-2="[165,1408,193,1864]" captionTargetId-0="figure@6.[151,1436,193,1932]" captionTargetId-1="figure@7.[191,1396,193,1823]" captionTargetId-2="figure@8.[165,1409,193,1876]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 9 – 20. Archellerenius giustoi, male (13, 15), female (9 – 12, 14, 17 – 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." captionText-1="FIGURES 21 – 32. Archellerenius giustoi, male (23 – 26), female (21, 22, 27 – 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 – 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." captionText-2="FIGURES 33 – 45. Archellerenius giustoi, male (33 – 37), Hellerenius lobifer, male (38 – 42) and female (43 – 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.249084" figureDoi-1="http://doi.org/10.5281/zenodo.249085" figureDoi-2="http://doi.org/10.5281/zenodo.249086" httpUri-0="https://zenodo.org/record/249084/files/figure.png" httpUri-1="https://zenodo.org/record/249085/files/figure.png" httpUri-2="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">9–37</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3536542BF4CFFFDFF5C5D10FDC4F433" pageId="3" pageNumber="518" type="materials_examined">
<paragraph id="8BF636C9BF4CFFFDFF5C5D10FA9CF777" blockId="3.[151,1437,258,1870]" box="[151,1370,258,283]" pageId="3" pageNumber="518">
<emphasis id="B93DEADBBF4CFFFDFF5C5D10FE83F777" bold="true" box="[151,325,258,283]" pageId="3" pageNumber="518">
<typeStatus id="54F2886BBF4CFFFDFF5C5D10FF14F777" box="[151,210,258,283]" pageId="3" pageNumber="518">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54F2886BBF4CFFFDFE875D10FE72F777" box="[332,436,258,283]" pageId="3" pageNumber="518" type="holotype">Holotype</typeStatus>
♂: a) 
<collectingCountry id="F35E7659BF4CFFFDFE395D11FD63F777" box="[498,677,258,283]" name="New Caledonia" pageId="3" pageNumber="518">New Caledonia</collectingCountry>
, Col des Roussettes, 
<quantity id="4CB19B2CBF4CFFFDFC5A5D10FC1CF776" box="[913,986,258,283]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="518" unit="m" value="500.0">500 m</quantity>
, 
<date id="FFF71009BF4CFFFDFC2D5D10FBA1F777" box="[998,1127,258,283]" pageId="3" pageNumber="518" value="1963-02-05">5 Feb 1963</date>
, G. Kuschel (NZAC).
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C5D35FA41F7A3" blockId="3.[151,1437,258,1870]" pageId="3" pageNumber="518">
<typeStatus id="54F2886BBF4CFFFDFF0C5D35FEF2F753" box="[199,308,295,319]" pageId="3" pageNumber="518" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D4FFD40BF4CFFFDFE885D34FEABF752" box="[323,365,294,319]" pageId="3" pageNumber="518" type="generic">5 ♂</specimenCount>
3 ♀): Province Nord: Mt. Aoupinié, W of Ponerihouen [near sawmill, -21.1555/165.3201], 
<quantity id="4CB19B2CBF4CFFFDFAB95D34FF69F70E" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="518" unit="m" value="550.0">550 m</quantity>
, 
<date id="FFF71009BF4CFFFDFF715D59FEA9F70F" box="[186,367,331,355]" pageId="3" pageNumber="518" value="1971-07-30" valueMax="1971-07-31" valueMin="1971-07-30">30–31 VII 1971</date>
, beating, 
<specimenCount id="9D4FFD40BF4CFFFDFE105D59FDC0F70E" box="[475,518,331,355]" pageId="3" pageNumber="518" type="generic">1 ♂</specimenCount>
, leg. J. Gressitt (BPBM). Mt Panié (refuge hut), 20.34.43S/164.46.04E [-20.5786/ 164.7678], 
<quantity id="4CB19B2CBF4CFFFDFEDC5D7DFEA9F7EA" box="[279,367,367,391]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="3" pageNumber="518" unit="m" value="1330.0">1330 m</quantity>
, humid montane forest, 
<date id="FFF71009BF4CFFFDFDB45D7DFD2FF7EB" box="[639,745,367,391]" pageId="3" pageNumber="518" value="2004-02-02">2 II 2004</date>
, 1 ♀; 
<quantity id="4CB19B2CBF4CFFFDFCE55D7DFC0DF7EA" box="[814,971,367,391]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="3" pageNumber="518" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200–1300 m</quantity>
, 
<date id="FFF71009BF4CFFFDFC1C5D7DFB84F7EB" box="[983,1090,367,391]" pageId="3" pageNumber="518" value="2004-02-03">3 II 2004</date>
, 
<specimenCount id="9D4FFD40BF4CFFFDFB865D7DFBB1F7EA" box="[1101,1143,367,391]" pageId="3" pageNumber="518" type="generic">1 ♂</specimenCount>
1 ♀ - all leg. MW (MW). Upper Tchamba Valley: Wâo Uni (refuge), -21.0057/165.2487, 
<quantity id="4CB19B2CBF4CFFFDFCBF5D81FC06F7C6" box="[884,960,403,427]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="518" unit="m" value="400.0">400 m</quantity>
, 
<date id="FFF71009BF4CFFFDFC045D80FB85F7C7" box="[975,1091,402,427]" pageId="3" pageNumber="518" value="2007-01-15">15 I 2007</date>
, night beating, 
<specimenCount id="9D4FFD40BF4CFFFDFB335D81FAE0F7C6" box="[1272,1318,403,427]" pageId="3" pageNumber="518" type="generic">1 ♂</specimenCount>
, leg. MW (MW). Col des Roussettes (rainforest near refuge), -21.4074/165.5250, 
<quantity id="4CB19B2CBF4CFFFDFC715DA4FBC5F7A2" box="[954,1027,438,463]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="518" unit="m" value="530.0">530 m</quantity>
, 
<date id="FFF71009BF4CFFFDFBC45DA5FB4BF7A3" box="[1039,1165,439,463]" pageId="3" pageNumber="518" value="2010-12-02">2 XII 2010</date>
, 1 ♀, leg. MW (MW).
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C5DC8FE41F457" blockId="3.[151,1437,258,1870]" pageId="3" pageNumber="518">
Province Sud: Pic du Pin base (forest &amp; plantation), -22.2482/166.829, 280 m, 
<date id="FFF71009BF4CFFFDFB8C5DC8FB12F79F" box="[1095,1236,474,499]" pageId="3" pageNumber="518" value="2006-12-26">26 XII 2006</date>
, 
<specimenCount id="9D4FFD40BF4CFFFDFB2B5DC9FACBF79E" box="[1248,1293,475,499]" pageId="3" pageNumber="518" type="generic">1 ♂</specimenCount>
, leg. MW &amp; RD (MW). Mt Humboldt (S trail), Botanical Reserve, -21.8827/166.3997, 
<quantity id="4CB19B2CBF4CFFFDFC165DEDFBBCF47A" box="[989,1146,511,535]" metricMagnitude="3" metricUnit="m" metricValue="1.205" metricValueMax="1.28" metricValueMin="1.13" pageId="3" pageNumber="518" unit="m" value="1205.0" valueMax="1280.0" valueMin="1130.0">1130–1280 m</quantity>
, moss forest, 
<date id="FFF71009BF4CFFFDFADE5DEDFA5EF47B" box="[1301,1432,511,535]" pageId="3" pageNumber="518" value="2008-11-10">10 XI 2008</date>
, 
<specimenCount id="9D4FFD40BF4CFFFDFF5C5E31FF05F456" box="[151,195,547,571]" pageId="3" pageNumber="518" type="generic">1 ♂</specimenCount>
, leg. MW (MW).
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C5E54FDC4F433" blockId="3.[151,1437,258,1870]" box="[199,514,582,607]" pageId="3" pageNumber="518">
Total distribution in 
<figureCitation id="13722A4CBF4CFFFDFE675E54FE38F433" box="[428,510,582,607]" captionStart="FIGURE 46" captionStartId="9.[151,250,1040,1062]" captionText="FIGURE 46. Localities of Archellerenius giustoi in New Caledonia mainland." pageId="3" pageNumber="518">Fig. 46</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536542BF4CFFFDFF0C5E78FD4AF4EF" box="[199,652,618,643]" pageId="3" pageNumber="518" type="diagnosis">
<paragraph id="8BF636C9BF4CFFFDFF0C5E78FD4AF4EF" blockId="3.[151,1437,258,1870]" box="[199,652,618,643]" pageId="3" pageNumber="518">
<emphasis id="B93DEADBBF4CFFFDFF0C5E78FE87F4EF" bold="true" box="[199,321,618,643]" pageId="3" pageNumber="518">Diagnosis.</emphasis>
See generic diagnosis above.
</paragraph>
</subSubSection>
<subSubSection id="C3536542BF4CFFFDFF0C5E9CFBB2F16A" pageId="3" pageNumber="518" type="description">
<paragraph id="8BF636C9BF4CFFFDFF0C5E9CFC35F4CA" blockId="3.[151,1437,258,1870]" box="[199,1011,654,679]" pageId="3" pageNumber="518">
<emphasis id="B93DEADBBF4CFFFDFF0C5E9CFE9FF4CB" bold="true" box="[199,345,654,679]" pageId="3" pageNumber="518">Description.</emphasis>
Body 
<quantity id="4CB19B2CBF4CFFFDFE685E9DFDECF4CA" box="[419,554,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="3" pageNumber="518" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8–3.2 mm</quantity>
long; antennal scape entirely testaceous.
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C5EA1FF18F5A6" blockId="3.[151,1437,258,1870]" pageId="3" pageNumber="518">
Rostrum shaped as in 
<figureCitation id="13722A4CBF4CFFFDFE775EA0FDFCF4A7" box="[444,570,690,715]" captionStart="FIGURES 9 – 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 – 20. Archellerenius giustoi, male (13, 15), female (9 – 12, 14, 17 – 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Figs 13–16</figureCitation>
, at base finely punctured, rugose at sides; prorostrum shiny, with microscopic punctures at least basally. Antennal club as long as 6 distal funicular segments combined. Head: frons usually with shallow V-like fovea; vertex evenly flattened throughout; subocular tooth usually weakly expressed in head profile. Antennae (
<figureCitation id="13722A4CBF4CFFFDFEDA5F33FEA3F556" box="[273,357,801,826]" captionStart="FIGURES 9 – 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 – 20. Archellerenius giustoi, male (13, 15), female (9 – 12, 14, 17 – 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Fig. 19</figureCitation>
): length/width ratios: scape 6.5–8.0, funicular segments: 1st 1.3–1.5, 2nd ca. 2.5, 6th–7th 1.0, scape as long or longer than 6 basal funicular segments combined. Pronotum weakly rounded at sides (
<figureCitation id="13722A4CBF4CFFFDFA895F57FA48F532" box="[1346,1422,837,862]" captionStart="FIGURES 9 – 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 – 20. Archellerenius giustoi, male (13, 15), female (9 – 12, 14, 17 – 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Fig.12</figureCitation>
); hypomeron acutely prominent at posterior margin. Elytra moderately convex (
<figureCitation id="13722A4CBF4CFFFDFBF55F7BFB7FF5EE" box="[1086,1209,873,898]" captionStart="FIGURES 9 – 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 – 20. Archellerenius giustoi, male (13, 15), female (9 – 12, 14, 17 – 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Figs 9–11</figureCitation>
); striae superficial; intervals flat, finely microreticulate and shiny. Fore leg as in 
<figureCitation id="13722A4CBF4CFFFDFCF75F9FFC7DF5CA" box="[828,955,909,934]" captionStart="FIGURES 9 – 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 – 20. Archellerenius giustoi, male (13, 15), female (9 – 12, 14, 17 – 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Figs 18, 20</figureCitation>
. Abdomen (ventrites 1–5) 1.2× longer than broad.
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C5FC7FD1DF38A" blockId="3.[151,1437,258,1870]" pageId="3" pageNumber="518">
Male. Antennal club about 2.5× as long as wide. Abdominal ventrites narrower (
<figureCitation id="13722A4CBF4CFFFDFB9B5FC7FB62F582" box="[1104,1188,981,1006]" captionStart="FIGURES 21 – 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 – 32. Archellerenius giustoi, male (23 – 26), female (21, 22, 27 – 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 – 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 25</figureCitation>
); 5th ventrite twice as wide as long, broadly truncate, moderately and evenly convex, middle of apical margin with sparse fringe of white setae (
<figureCitation id="13722A4CBF4CFFFDFF14580FFEF2F25A" box="[223,308,1053,1078]" captionStart="FIGURES 21 – 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 – 32. Archellerenius giustoi, male (23 – 26), female (21, 22, 27 – 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 – 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 26</figureCitation>
). Pygidium: apical third exposed, strongly declivitous, with distinct marginal rim; separating sulcus obsolescent (
<figureCitation id="13722A4CBF4CFFFDFEE25853FE6DF236" box="[297,427,1089,1114]" captionStart="FIGURES 21 – 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 – 32. Archellerenius giustoi, male (23 – 26), female (21, 22, 27 – 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 – 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Figs 23, 24</figureCitation>
). Sternite VIII with distinct carinae. Sternite IX with untoothed apodeme, ca. 1.6× as long as basal fork (
<figureCitation id="13722A4CBF4CFFFDFEF15877FE57F212" box="[314,401,1125,1150]" captionStart="FIGURES 33 – 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 – 45. Archellerenius giustoi, male (33 – 37), Hellerenius lobifer, male (38 – 42) and female (43 – 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">Fig. 35</figureCitation>
). Tegminal apodeme strongly expanded in apical third; tegminal plate 1.3–1.4× as long as broad, with lateral arms much thinner than adjoining arms of basal piece (
<figureCitation id="13722A4CBF4CFFFDFC2F589BFBAFF2CE" box="[996,1129,1161,1186]" captionStart="FIGURES 33 – 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 – 45. Archellerenius giustoi, male (33 – 37), Hellerenius lobifer, male (38 – 42) and female (43 – 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">Figs 36, 37</figureCitation>
); membranous lobes short, rounded, apically extended into narrow, acute processes, these microsetose apically; suprafenestral sclerites half as long as fenestrae, acutely produced along internal margins; sensillae indistinct; macrochaetae 6–8, short, hardly exceeding acute sclerotized projections of parameroid lobes; membrane beneath fenestrae sclerotized equally to prostegium; postfenestral sclerite bar-like; prostegium long, deeply incised laterally, divided on its 0.6–0.7 length. Penis: apodemes as long as pedon, strongly expanded apicad; tectum reaching apical fourth of pedon; pedon with rounded, strongly upcurved apex (
<figureCitation id="13722A4CBF4CFFFDFDDE5973FD50F316" box="[533,662,1377,1402]" captionStart="FIGURES 33 – 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 – 45. Archellerenius giustoi, male (33 – 37), Hellerenius lobifer, male (38 – 42) and female (43 – 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">Figs 33, 34</figureCitation>
), ventrally membranous in basal half; endophallus with enclosed part densely microspinose in basal third, with sharp and straight distal border-line separating it from big, simply membranous terminal process, the latter with paired basal lobes covering entrance of ejaculatory duct, microplates in sub-orificial region extremely fine and indistinct.
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C59E3FEF1F08E" blockId="3.[151,1437,258,1870]" pageId="3" pageNumber="518">
Female. Antennal club more obtuse, 2.0–2.2× as long as wide (
<figureCitation id="13722A4CBF4CFFFDFC4259E3FC1AF066" box="[905,988,1521,1546]" captionStart="FIGURES 9 – 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 – 20. Archellerenius giustoi, male (13, 15), female (9 – 12, 14, 17 – 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Fig. 19</figureCitation>
). Abdominal ventrites shaped as in 
<figureCitation id="13722A4CBF4CFFFDFAA059E3FF26F042" captionStart="FIGURES 21 – 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 – 32. Archellerenius giustoi, male (23 – 26), female (21, 22, 27 – 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 – 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Figs 27, 28</figureCitation>
. Tergite VII short, broadly rounded (
<figureCitation id="13722A4CBF4CFFFDFDB45A07FD12F042" box="[639,724,1557,1582]" captionStart="FIGURES 21 – 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 – 32. Archellerenius giustoi, male (23 – 26), female (21, 22, 27 – 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 – 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 30</figureCitation>
). Tergite VIII transverse, largely membranous, with narrow but complete marginal sclerotisation. Sternite VIII (spiculum ventrale) relatively short, slightly widened in middle part, apically with triangular expansion (
<figureCitation id="13722A4CBF4CFFFDFDED5A4FFDBDF01A" box="[550,635,1629,1654]" captionStart="FIGURES 21 – 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 – 32. Archellerenius giustoi, male (23 – 26), female (21, 22, 27 – 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 – 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 32</figureCitation>
). Styli stout and short, hemispherical, with several apical setae; coxites subtruncate apically. Foretube darkened, with dense long microtrichia. Bursa copulatrix relatively large, with corrugated longitudinal folds. Spermatheca: corpus with hook-like process receiving relatively thick spermathecal duct (
<figureCitation id="13722A4CBF4CFFFDFF1D5ADBFEECF08E" box="[214,298,1737,1762]" captionStart="FIGURES 21 – 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 – 32. Archellerenius giustoi, male (23 – 26), female (21, 22, 27 – 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 – 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C5AFFFBB2F16A" blockId="3.[151,1437,258,1870]" box="[199,1140,1773,1798]" pageId="3" pageNumber="518">Morphological indices in Tab. 1. For other characters see generic description above.</paragraph>
</subSubSection>
<subSubSection id="C3536542BF4CFFFDFF0C5B03FAE0F121" pageId="3" pageNumber="518" type="etymology">
<paragraph id="8BF636C9BF4CFFFDFF0C5B03FAE0F121" blockId="3.[151,1437,258,1870]" pageId="3" pageNumber="518">
<emphasis id="B93DEADBBF4CFFFDFF0C5B03FE88F146" bold="true" box="[199,334,1809,1834]" pageId="3" pageNumber="518">Etymology.</emphasis>
This remarkable and rare species is named to honor Mr. Carlo Giusto, an enthusiastic student of 
<taxonomicName id="4C494D4ABF4CFFFDFF5C5B27FEC9F122" box="[151,271,1845,1870]" class="Insecta" family="Apionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="family">Apionidae</taxonomicName>
, for his passion for these inconspicuous weevils, and shared interest in the genus 
<taxonomicName id="4C494D4ABF4CFFFDFB555B27FAE6F121" box="[1182,1312,1845,1869]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4CFFFDFB555B27FAE6F121" box="[1182,1312,1845,1869]" italics="true" pageId="3" pageNumber="518">Hellerenius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>