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<mods:title id="FA3348EECEF18D5B6096B38AD60FA0F2">The Apionidae (Coleoptera: Curculionoidea) of New Caledonia. Genera Archellerenius gen. nov. and Hellerenius Wanat</mods:title>
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<taxonomicName id="4C494D4ABF42FFF3FF5C5D7DFE59F7E5" box="[151,415,367,393]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="528" phylum="Tracheophyta" rank="species" species="monteithi" status="sp. nov.">
<taxonomicName id="4C494D4ABF42FFF3FF5C5D7DFE59F7E5" authorityName="Wanat" authorityYear="2013" box="[151,415,367,393]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="528" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
<emphasis id="B93DEADBBF42FFF3FF5C5D7DFE59F7E5" bold="true" box="[151,415,367,393]" italics="true" pageId="13" pageNumber="528">Hellerenius monteithi</emphasis>
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<taxonomicNameLabel id="A20E57A0BF42FFF3FE6C5D62FDC2F7E6" box="[423,516,368,394]" pageId="13" pageNumber="528" rank="species">sp. nov.</taxonomicNameLabel>
@ -137,7 +138,7 @@ Total distribution in
Largest member of the genus, the only one exceeding
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of body length without rostrum, also unique in having dull black elytra with the intervals completely matt due to heavy scale-like microsculpture and the striae shallowly impressed as shiny rows, and long and thin rostrum, particularly in female. The species resembles
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<taxonomicName id="4C494D4ABF42FFF3FF5C5FE5FECCF262" authorityName="Wanat" authorityYear="2001" box="[151,266,1014,1038]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="528" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF42FFF3FF5C5FE5FECCF262" box="[151,266,1014,1038]" italics="true" pageId="13" pageNumber="528">H. lobifer</emphasis>
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in both male and female sexual characters, namely the structure of last abdominal ventrite, male metaventrite, pygidium and aedeagus, but clearly differs in having larger body, alutaceous black elytral intervals and much longer rostrum, in male over 1.7×, in female over 2.7× longer than pronotum.
@ -152,20 +153,20 @@ in both male and female sexual characters, namely the structure of last abdomina
</paragraph>
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Rostrum long and thin (Figs 6466), glabrous, with fine puncturation restricted to metarostrum and better developed on its sides, at base densely microreticulate; underside of prorostrum with distinct subgenal sulci in apical two-thirds (
<figureCitation id="13722A4CBF42FFF3FEA358E0FE68F367" box="[360,430,1266,1291]" captionStart="FIGURES 1 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="13" pageNumber="528">Fig. 5</figureCitation>
<figureCitation id="13722A4CBF42FFF3FEA358E0FE68F367" box="[360,430,1266,1291]" captionStart="FIGURES 1 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." figureDoi="http://doi.org/10.5281/zenodo.249083" httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="13" pageNumber="528">Fig. 5</figureCitation>
); median part of rostrum underside with single shallow median sulcus. Head small and narrow; frons with elongate median pit; inner eye margins distinctly angulate; vertex with convex and distinctly punctured anterior part well separated from polished remainder (Fig. 60); temples shorter than eye diameter; head underside with short U-shaped wrinkles; subocular tooth indistinct, obtuse. Antennae extremely long and thin (Figs 61, 62); scape as long (♂) or longer (♀) than funicle, length/width ratios of funicular segments: 1st 3.54.0, 2nd 4.5 5.0, 7th 1.21.5, club 4.04.2; club 0.5 (♂) to 0.4 (♀) scape length, shorter than combined length of 5 distal funicular segments. Pronotum markedly widened and rounded in apical third (Fig. 60); front margin not scratched; disc densely microreticulate, with relatively dense and superficial microscopic puncturation, subapical constriction traced across pronotal disc; subbasal line reduced; posterior lateral groove narrow but well impressed, with vestigial spur; hypomeron with small median prominence on posterior margin; prosternellum small, strongly prominent between coxae. Elytra with extended caudal part, highly convex but not angled in profile (Figs 58, 59); intervals evenly shagreened and totally alutaceous, with no trace of transverse grooves, all gently convex; striae shallow, with only vestigial punctures, distinctly shiny and thus contrasting with completely matt intervals; sensory setae up to 8 ordered along 9th stria and entering 2nd interval. Wings with largely expanded melanisations; radial window vestigial, narrow; anal 1A as two unequal rudiments (Fig. 73). Legs long; femora moderately clavate, metafemur exceeding elytra by ca. 0.25 length; tibiae straight, with dense and short protruding setae along major part of inner margin (Fig.69); protarsus as in Figs 70, 71; claws short, with low triangular teeth (Fig. 68).
</paragraph>
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Male. Rostrum thicker, with slightly constricted prorostrum (Figs 64, 65); metarostrum shallowly furrowed on sides apart from punctation. Antennal insertion 0.420.45. Metaventrite with round median impression and small pointed median tubercle close to metacoxae. Last abdominal ventrite with two raised, shortly setose lobes bordering median depression (Figs 72, 74). Pygidium flattened, in apical third with shallow transverse sulcus
<emphasis id="B93DEADBBF41FFF0FF5C5B6EFE95F1FE" bold="true" box="[151,339,1916,1939]" pageId="14" pageNumber="529">FIGURES 5872.</emphasis>
<taxonomicName id="4C494D4ABF41FFF0FE935B6FFDF5F1FE" box="[344,563,1917,1938]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="14" pageNumber="529" phylum="Tracheophyta" rank="species" species="monteithi">
<taxonomicName id="4C494D4ABF41FFF0FE935B6FFDF5F1FE" authorityName="Wanat" authorityYear="2013" box="[344,563,1917,1938]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="529" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B93DEADBBF41FFF0FE935B6FFDF5F1FE" box="[344,563,1917,1938]" italics="true" pageId="14" pageNumber="529">Hellerenius monteithi</emphasis>
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, male (59, 61, 6365, 68, 70, 72), female (58, 60, 62, 66, 67, 69, 71): 58) elytra, lateral view; 59) elytra, dorsal view; 60) head and pronotum, dorsal view; 61, 62) antenna; 63) head and pronotum, lateral view; 64 66) head with rostrum in dorsal (64) and lateral (65, 66) view; 67) rostrum base, lateral view; 68) tarsal claws; 69) fore femur and tibia; 70, 71) protarsus; 72) apex of last abdominal ventrite, posterior view.
</paragraph>
<paragraph id="8BF636C9BF40FFF1FF5C5B4CFD4EF183" blockId="15.[151,1436,1886,2031]" pageId="15" pageNumber="530">
<emphasis id="B93DEADBBF40FFF1FF5C5B4CFE92F118" bold="true" box="[151,340,1886,1909]" pageId="15" pageNumber="530">FIGURES 7385.</emphasis>
<taxonomicName id="4C494D4ABF40FFF1FE915B4DFDF3F118" box="[346,565,1887,1908]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="15" pageNumber="530" phylum="Tracheophyta" rank="species" species="monteithi">
<taxonomicName id="4C494D4ABF40FFF1FE915B4DFDF3F118" authorityName="Wanat" authorityYear="2013" box="[346,565,1887,1908]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="530" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B93DEADBBF40FFF1FE915B4DFDF3F118" box="[346,565,1887,1908]" italics="true" pageId="15" pageNumber="530">Hellerenius monteithi</emphasis>
</taxonomicName>
, male (74, 77, 78, 82), female (73, 75, 76,7981, 8385): 73) metathoracic wing; 74 76) all abdominal ventrites in dorsal (74, 75) and lateral (76) view; 77, 78) pygidium in dorsal (77) and lateral (78) view; 79) spermatheca; 8082) membranous pouch between tergites VII and VIII in dorsal (80, 82) and lateral (81) view, arrowed; 83) tergite VII; 84) sternite VIII, basal (posterior) part; 85) ovipositor, enlarged apices of gonocoxites with styli (85A). Abbreviations: d, spermathecal duct; gl, s. gland.
@ -174,7 +175,7 @@ Male. Rostrum thicker, with slightly constricted prorostrum (Figs 64, 65); metar
<paragraph id="8BF636C9BF5FFFEEFF5C5805FC69F241" blockId="16.[151,943,1047,1069]" box="[151,943,1047,1069]" pageId="16" pageNumber="531">
<emphasis id="B93DEADBBF5FFFEEFF5C5805FEE6F240" bold="true" box="[151,288,1047,1069]" pageId="16" pageNumber="531">FIGURE 86.</emphasis>
Localities of
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<taxonomicName id="4C494D4ABF5FFFEEFE605805FD43F240" authorityName="Wanat" authorityYear="2013" box="[427,645,1047,1068]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B93DEADBBF5FFFEEFE605805FD43F240" box="[427,645,1047,1068]" italics="true" pageId="16" pageNumber="531">Hellerenius monteithi</emphasis>
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in New Caledonia mainland.
@ -182,21 +183,21 @@ in New Caledonia mainland.
</caption>
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separating setose exposed part (Figs 77, 78), on anterior margin with pointed median tooth (Fig. 82). Sternite VIII evenly sclerotised, lobes broadly rounded, shorter than in
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<taxonomicName id="4C494D4ABF5FFFEEFC85586DFC79F2FA" authorityName="Wanat" authorityYear="2001" box="[846,959,1150,1174]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF5FFFEEFC85586DFC79F2FA" box="[846,959,1150,1174]" italics="true" pageId="16" pageNumber="531">H. lobifer</emphasis>
</taxonomicName>
. Sternite IX with well open basal fork; apodeme with minute tooth close to fork (
<figureCitation id="13722A4CBF5FFFEEFDB958B0FD00F2D6" box="[626,710,1186,1211]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="16" pageNumber="531">Fig. 91</figureCitation>
<figureCitation id="13722A4CBF5FFFEEFDB958B0FD00F2D6" box="[626,710,1186,1211]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249088" httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="16" pageNumber="531">Fig. 91</figureCitation>
). Tegminal plate (
<figureCitation id="13722A4CBF5FFFEEFC5958B0FBD2F2D6" box="[914,1044,1186,1211]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="16" pageNumber="531">Figs 89, 90</figureCitation>
<figureCitation id="13722A4CBF5FFFEEFC5958B0FBD2F2D6" box="[914,1044,1186,1211]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249088" httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="16" pageNumber="531">Figs 89, 90</figureCitation>
) and penis (
<figureCitation id="13722A4CBF5FFFEEFB5458B0FAE7F2D6" box="[1183,1313,1186,1211]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="16" pageNumber="531">Figs 87, 88</figureCitation>
<figureCitation id="13722A4CBF5FFFEEFB5458B0FAE7F2D6" box="[1183,1313,1186,1211]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249088" httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="16" pageNumber="531">Figs 87, 88</figureCitation>
) similar to those in
<taxonomicName id="4C494D4ABF5FFFEEFF3D58D5FEA2F2B2" box="[246,356,1222,1246]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="16" pageNumber="531" phylum="Tracheophyta" rank="species" species="lobifer">
<taxonomicName id="4C494D4ABF5FFFEEFF3D58D5FEA2F2B2" authorityName="Wanat" authorityYear="2001" box="[246,356,1222,1246]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF5FFFEEFF3D58D5FEA2F2B2" box="[246,356,1222,1246]" italics="true" pageId="16" pageNumber="531">H. lobifer</emphasis>
</taxonomicName>
; tegminal membranous lobes more acute apically; penis apodemes shorter than pedon and more expanded distally; pedon more elongate (almost 7× as long as broad, while ca. 6× in
<taxonomicName id="4C494D4ABF5FFFEEFBA458F9FB19F36E" box="[1135,1247,1258,1282]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="16" pageNumber="531" phylum="Tracheophyta" rank="species" species="lobifer">
<taxonomicName id="4C494D4ABF5FFFEEFBA458F9FB19F36E" authorityName="Wanat" authorityYear="2001" box="[1135,1247,1258,1282]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF5FFFEEFBA458F9FB19F36E" box="[1135,1247,1258,1282]" italics="true" pageId="16" pageNumber="531">H. lobifer</emphasis>
</taxonomicName>
); apex of pedon rather triangular than truncate.
@ -219,7 +220,7 @@ Perkins
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<emphasis id="B93DEADBBF5FFFEEFF0C5B17FE88F172" bold="true" box="[199,334,1797,1822]" pageId="16" pageNumber="531">Etymology.</emphasis>
Named to honor Geoff B. Monteith, a great enthusiast of New Caledonian nature and very efficient explorer of its beetle fauna, who first collected this outstanding species, as well as hundreds of other specimens of the
<taxonomicName id="4C494D4ABF5FFFEEFE915B5FFE15F10A" box="[346,467,1869,1894]" class="Insecta" family="Apionidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="family">Apionidae</taxonomicName>
<taxonomicName id="4C494D4ABF5FFFEEFE915B5FFE15F10A" box="[346,467,1869,1894]" class="Insecta" family="Apionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="531" phylum="Arthropoda" rank="family">Apionidae</taxonomicName>
.
</paragraph>
</subSubSection>

View file

@ -1,45 +1,46 @@
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<taxonomicName id="4C494D4ABF45FFF4FF5C598DFE12F3D6" authority="Wanat" authorityName="Wanat" authorityYear="2001" box="[151,468,1439,1466]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="525" phylum="Arthropoda" rank="species" species="lobifer">
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<emphasis id="B93DEADBBF45FFF4FF5C598DFEBDF3D5" bold="true" box="[151,379,1439,1465]" italics="true" pageId="10" pageNumber="525">Hellerenius lobifer</emphasis>
Wanat
@ -49,13 +50,13 @@ Wanat
</paragraph>
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(
<figureCitation id="13722A4CBF45FFF4FF5459D0FEE6F3B7" box="[159,288,1474,1499]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="10" pageNumber="525">Figs 3845</figureCitation>
<figureCitation id="13722A4CBF45FFF4FF5459D0FEE6F3B7" box="[159,288,1474,1499]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="10" pageNumber="525">Figs 3845</figureCitation>
,
<figureCitation id="13722A4CBF45FFF4FEE059D0FEB5F3B7" box="[299,371,1474,1499]" captionStart-0="FIGURES 47 56" captionStart-1="FIGURE 57" captionStartId-0="11.[151,264,1660,1683]" captionStartId-1="12.[151,250,2001,2023]" captionTargetBox-0="[163,1422,412,1628]" captionTargetId-0="figure@11.[151,1436,412,1640]" captionTargetPageId-0="11" captionText-0="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." captionText-1="FIGURE 57. Localities of Hellerenius lobifer in New Caledonia mainland." httpUri-0="https://zenodo.org/record/249087/files/figure.png" pageId="10" pageNumber="525">4757</figureCitation>
<figureCitation id="13722A4CBF45FFF4FEE059D0FEB5F3B7" box="[299,371,1474,1499]" captionStart-0="FIGURES 47 56" captionStart-1="FIGURE 57" captionStartId-0="11.[151,264,1660,1683]" captionStartId-1="12.[151,250,2001,2023]" captionTargetBox-0="[163,1422,412,1628]" captionTargetId-0="figure@11.[151,1436,412,1640]" captionTargetPageId-0="11" captionText-0="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." captionText-1="FIGURE 57. Localities of Hellerenius lobifer in New Caledonia mainland." figureDoi-0="http://doi.org/10.5281/zenodo.249087" httpUri-0="https://zenodo.org/record/249087/files/figure.png" pageId="10" pageNumber="525">4757</figureCitation>
)
</paragraph>
<paragraph id="8BF636C9BF45FFF4FF5C5A17FDD0F077" blockId="10.[151,534,1541,1563]" box="[151,534,1541,1563]" pageId="10" pageNumber="525">
<taxonomicName id="4C494D4ABF45FFF4FF5C5A17FDD4F077" authority="Wanat, 2001: 139" authorityName="Wanat" authorityPageNumber="139" authorityYear="2001" box="[151,530,1541,1563]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="10" pageNumber="525" phylum="Tracheophyta" rank="species" species="lobifer">
<taxonomicName id="4C494D4ABF45FFF4FF5C5A17FDD4F077" authority="Wanat, 2001: 139" authorityName="Wanat" authorityPageNumber="139" authorityYear="2001" box="[151,530,1541,1563]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="525" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF45FFF4FF5C5A17FE90F076" box="[151,342,1541,1562]" italics="true" pageId="10" pageNumber="525">Hellerenius lobifer</emphasis>
Wanat, 2001: 139
</taxonomicName>
@ -71,19 +72,19 @@ Body length 4.0
</paragraph>
<paragraph id="8BF636C9BF45FFF5FF0C5A7FFEC6F70B" blockId="10.[151,1436,1608,2034]" lastBlockId="11.[151,1436,151,359]" lastPageId="11" lastPageNumber="526" pageId="10" pageNumber="525">
Rostrum as in
<figureCitation id="13722A4CBF45FFF4FEBF5A7EFE31F0E9" box="[372,503,1644,1669]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="10" pageNumber="525">Figs 5153</figureCitation>
<figureCitation id="13722A4CBF45FFF4FEBF5A7EFE31F0E9" box="[372,503,1644,1669]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." figureDoi="http://doi.org/10.5281/zenodo.249087" httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="10" pageNumber="525">Figs 5153</figureCitation>
, glabrous, impunctate, finely microreticulate, at base irregularly crenulate (more distinctly in male); venter of prorostrum polished, with traces of ventral sulci in apical third. Head: frons often with indistinct median pit; vertex weakly convex and finely punctate in front; temples evidently shorter than eye; head underside shagreened, posteriorly with coarse wrinkles and usually with small median tubercle. Antennae (
<figureCitation id="13722A4CBF45FFF4FAA55ACAFF7EF17B" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="10" pageNumber="525">Fig. 50</figureCitation>
<figureCitation id="13722A4CBF45FFF4FAA55ACAFF7EF17B" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." figureDoi="http://doi.org/10.5281/zenodo.249087" httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="10" pageNumber="525">Fig. 50</figureCitation>
): length/width ratios: scape ca. 10.0, funicular segments: 1st 2.63.0, 2nd 3.44.3, 7th 1.21.5, club 4.04.2; scape as long as 56 basal funicular segments combined; club ca. 0.75 scape length. Pronotum: front margin partly with more or less distinct short, oblique scratches; disc densely microreticulate, with microscopic punctulation and subapical constriction traced across; subbasal line often retained; posterior lateral groove shallow; behind procoxae sternellum and hypomeron even. Elytra with narrowly extended caudal part—especially in male, in profile somewhat conical (
<figureCitation id="13722A4CBF45FFF4FEBF5BA1FE3DF1A7" box="[372,507,1971,1996]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="10" pageNumber="525">Figs 47, 48</figureCitation>
<figureCitation id="13722A4CBF45FFF4FEBF5BA1FE3DF1A7" box="[372,507,1971,1996]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." figureDoi="http://doi.org/10.5281/zenodo.249087" httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="10" pageNumber="525">Figs 47, 48</figureCitation>
); intervals with dense scale-like microsculpture and irregular transverse grooves; mid portions of striae progressively broader outwards; sensory setae 34 ordered along 9th stria and 1 closer to junction of striae 3+6. Wing: radial window obsolescent; anal 1A as two rudiments; 3A distinct, with expanded apical melanisation, forming nearly complete anal cell with 2A. Metaventrite only 1.1× longer than large and prominent mesocoxae. Legs robust; coxae and trochanters with sparse, minute, erect setae, in male present also on base of femur; metafemur exceeding elytra by ca. 0.33 length; tibiae straight, with apical tuft extended along distal third of inner margin; protarsus 2.72.9×, its 1st segment 2.32.6× as long as wide, 2nd tarsomere slightly transverse (
<figureCitation id="13722A4CBF44FFF5FF545D5CFF32F70B" box="[159,244,334,359]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="11" pageNumber="526">Fig. 49</figureCitation>
<figureCitation id="13722A4CBF44FFF5FF545D5CFF32F70B" box="[159,244,334,359]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." figureDoi="http://doi.org/10.5281/zenodo.249087" httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="11" pageNumber="526">Fig. 49</figureCitation>
).
</paragraph>
<caption id="DF366641BF44FFF5FF5C5A6EFCFDF083" httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="11" pageNumber="526" targetBox="[163,1422,412,1628]" targetPageId="11">
<caption id="DF366641BF44FFF5FF5C5A6EFCFDF083" ID-DOI="http://doi.org/10.5281/zenodo.249087" httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="11" pageNumber="526" targetBox="[163,1422,412,1628]" targetPageId="11">
<paragraph id="8BF636C9BF44FFF5FF5C5A6EFCFDF083" blockId="11.[151,1436,1660,1775]" pageId="11" pageNumber="526">
<emphasis id="B93DEADBBF44FFF5FF5C5A6EFE90F0FE" bold="true" box="[151,342,1660,1683]" pageId="11" pageNumber="526">FIGURES 4756.</emphasis>
<taxonomicName id="4C494D4ABF44FFF5FE965A6FFDD8F0FE" box="[349,542,1661,1682]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="11" pageNumber="526" phylum="Tracheophyta" rank="species" species="lobifer">
<taxonomicName id="4C494D4ABF44FFF5FE965A6FFDD8F0FE" authorityName="Wanat" authorityYear="2001" box="[349,542,1661,1682]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="526" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF44FFF5FE965A6FFDD8F0FE" box="[349,542,1661,1682]" italics="true" pageId="11" pageNumber="526">Hellerenius lobifer</emphasis>
</taxonomicName>
, male (49, 51), female (47, 48, 50, 5256): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view.
@ -91,24 +92,24 @@ Rostrum as in
</caption>
<paragraph id="8BF636C9BF44FFF2FF0C5B04FBD7F7E4" blockId="11.[151,1436,1814,2019]" lastBlockId="12.[151,1436,151,1077]" lastPageId="12" lastPageNumber="527" pageId="11" pageNumber="526">
Male. Rostrum bare ventrally. Antennal insertion 0.370.39. Legs unmodified. Metaventrite with round median depression bearing small tubercle on posterior margin. Last abdominal ventrite with pair of highly elevated, protrudingly setose lateral lobes bordering polished and finely punctured median concavity. Pygidium apical third exposed, separated by shallow but complete transverse sulcus; marginal rim obsolete; tongue-like process broadly rounded. Sternite VIII evenly sclerotised, lobes long, triangular. Sternite IX with narrow and short basal fork; apodeme with minute tooth close to fork (
<figureCitation id="13722A4CBF44FFF5FD265BD8FC87F18F" box="[749,833,1994,2019]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="11" pageNumber="526">Fig. 38</figureCitation>
<figureCitation id="13722A4CBF44FFF5FD265BD8FC87F18F" box="[749,833,1994,2019]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="11" pageNumber="526">Fig. 38</figureCitation>
). Tegminal plate (
<figureCitation id="13722A4CBF44FFF5FBC05BD8FB4AF18F" box="[1035,1164,1994,2019]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="11" pageNumber="526">Figs 41, 42</figureCitation>
<figureCitation id="13722A4CBF44FFF5FBC05BD8FB4AF18F" box="[1035,1164,1994,2019]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="11" pageNumber="526">Figs 41, 42</figureCitation>
) twice as long as broad, subarticulated, with lobes only narrowly separated; membranous lobes very short; suprafenestral sclerites narrowing distad, about twice as long as fenestrae; macrochaetae 1215, varying in length but never longer than fenestrae, spread through apical half of each suprafenestral sclerite; sensilla numerous; fenestrae elongate, clearly margined; postfenestral plate largely reduced; prostegium long, uniform, shallowly incised at sides. Penis: apodemes and pedon equally long; pedon parallel sided, slightly constricted subapically, subtruncate and weakly downcurved at apex (
<figureCitation id="13722A4CBF43FFF2FE425D5EFE10F709" box="[393,470,332,357]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="12" pageNumber="527">Fig.39</figureCitation>
<figureCitation id="13722A4CBF43FFF2FE425D5EFE10F709" box="[393,470,332,357]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="12" pageNumber="527">Fig.39</figureCitation>
); tectum membranous, shortened; terminal membranous process of endophallus small (
<figureCitation id="13722A4CBF43FFF2FF545D7DFF2DF7E4" box="[159,235,367,392]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="12" pageNumber="527">Fig.40</figureCitation>
<figureCitation id="13722A4CBF43FFF2FF545D7DFF2DF7E4" box="[159,235,367,392]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="12" pageNumber="527">Fig.40</figureCitation>
), suborificial region armed with minute, isodiametric sclerotized plates.
</paragraph>
<paragraph id="8BF636C9BF43FFF2FF0C5D86FEDDF4A1" blockId="12.[151,1436,151,1077]" pageId="12" pageNumber="527">
Female. Prorostrum narrowest at apex; underside of mesorostrum polished and impunctate. Antennal insertion 0.300.36, segments of funicle usually more elongate. Last abdominal ventrite triangular, ca. 1.6× as wide as long, elevating apicad, with large median depression and distinct, polished apical flange formed of narrow posterior extensions of laterosternites (
<figureCitation id="13722A4CBF43FFF2FE295DEDFDA0F474" box="[482,614,511,536]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="12" pageNumber="527">Figs 5456</figureCitation>
<figureCitation id="13722A4CBF43FFF2FE295DEDFDA0F474" box="[482,614,511,536]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." figureDoi="http://doi.org/10.5281/zenodo.249087" httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="12" pageNumber="527">Figs 5456</figureCitation>
). Tergite VII campaniform in outline (
<figureCitation id="13722A4CBF43FFF2FBD05DEDFBB6F474" box="[1051,1136,511,536]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="12" pageNumber="527">Fig. 44</figureCitation>
<figureCitation id="13722A4CBF43FFF2FBD05DEDFBB6F474" box="[1051,1136,511,536]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="12" pageNumber="527">Fig. 44</figureCitation>
). Terminalia as in
<figureCitation id="13722A4CBF43FFF2FA8F5DEDFA51F474" box="[1348,1431,511,536]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="12" pageNumber="527">Fig. 45</figureCitation>
<figureCitation id="13722A4CBF43FFF2FA8F5DEDFA51F474" box="[1348,1431,511,536]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="12" pageNumber="527">Fig. 45</figureCitation>
; tergite VIII slightly elongate, without evident submedian constriction, with short marginal setae; sternite VIII ca. 2.4× longer than coxite, curved and with elongate apical plate; ovipositor parallelsided; foretube (genital sheath) densely clothed with microtrichia simple or arranged in combs, in part covering gonocoxites replaced with simple triangular plates; spermatheca rather reverse L-shaped (
<figureCitation id="13722A4CBF43FFF2FCCD5E9DFC9CF4C4" box="[774,858,655,680]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="12" pageNumber="527">Fig. 43</figureCitation>
<figureCitation id="13722A4CBF43FFF2FCCD5E9DFC9CF4C4" box="[774,858,655,680]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="12" pageNumber="527">Fig. 43</figureCitation>
); openings of duct and gland well separated, not on projections.
</paragraph>
<paragraph id="8BF636C9BF43FFF2FF0C5ECAFAE8F49C" blockId="12.[151,1436,151,1077]" box="[199,1326,727,752]" pageId="12" pageNumber="527">For more detailed morphological description consult Wanat (2001). Morphological indices in Tab. 1.</paragraph>
@ -117,10 +118,11 @@ Female. Prorostrum narrowest at apex; underside of mesorostrum polished and impu
<paragraph id="8BF636C9BF43FFF2FF0C5EEEFEA1F554" blockId="12.[151,1436,151,1077]" pageId="12" pageNumber="527">
<emphasis id="B93DEADBBF43FFF2FF0C5EEEFEE1F579" bold="true" box="[199,295,764,789]" pageId="12" pageNumber="527">Biology.</emphasis>
Specimens were observed while piercing ripened fruit of
<taxonomicName id="4C494D4ABF43FFF2FC2E5EEEFBA6F578" box="[997,1120,764,788]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="12" pageNumber="527" phylum="Tracheophyta" rank="genus">
<taxonomicName id="4C494D4ABF43FFF2FC2E5EEEFB48F578" box="[997,1166,764,788]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="12" pageNumber="527" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93DEADBBF43FFF2FC2E5EEEFBA6F578" box="[997,1120,764,788]" italics="true" pageId="12" pageNumber="527">Hedycarya</emphasis>
sp.
</taxonomicName>
sp. for feeding (Wanat &amp; Munzinger, 2012).
for feeding (Wanat &amp; Munzinger, 2012).
</paragraph>
</subSubSection>
<subSubSection id="C3536542BF43FFF2FF0C5F56FC28F259" pageId="12" pageNumber="527" type="materials_examined">
@ -135,10 +137,11 @@ Province Nord: Thoven,
,
<date id="FFF71009BF43FFF2FB215F56FA9EF530" box="[1258,1368,836,861]" pageId="12" pageNumber="527" value="2004-02-05">5 II 2004</date>
, from
<taxonomicName id="4C494D4ABF43FFF2FF5C5F7AFED4F5EC" box="[151,274,872,896]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="12" pageNumber="527" phylum="Tracheophyta" rank="genus">
<taxonomicName id="4C494D4ABF43FFF2FF5C5F7AFEFCF513" box="[151,314,872,896]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="12" pageNumber="527" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93DEADBBF43FFF2FF5C5F7AFED4F5EC" box="[151,274,872,896]" italics="true" pageId="12" pageNumber="527">Hedycarya</emphasis>
sp.
</taxonomicName>
sp., 2 ♀, leg. MW (MW).
, 2 ♀, leg. MW (MW).
</paragraph>
<paragraph id="8BF636C9BF43FFF2FF0C5F9EFB28F27C" blockId="12.[151,1436,151,1077]" pageId="12" pageNumber="527">
Province Sud: Mt Mou [-22.06/166.35], 7-12
@ -184,7 +187,7 @@ Remaining material listed in Wanat (2001). Total distribution in
<paragraph id="8BF636C9BF43FFF2FF5C5BC3FC55F18B" blockId="12.[151,915,2001,2023]" box="[151,915,2001,2023]" pageId="12" pageNumber="527">
<emphasis id="B93DEADBBF43FFF2FF5C5BC3FED9F18A" bold="true" box="[151,287,2001,2023]" pageId="12" pageNumber="527">FIGURE 57.</emphasis>
Localities of
<taxonomicName id="4C494D4ABF43FFF2FE605BC3FDADF18A" box="[427,619,2001,2022]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="12" pageNumber="527" phylum="Tracheophyta" rank="species" species="lobifer">
<taxonomicName id="4C494D4ABF43FFF2FE605BC3FDADF18A" authorityName="Wanat" authorityYear="2001" box="[427,619,2001,2022]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="527" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF43FFF2FE605BC3FDADF18A" box="[427,619,2001,2022]" italics="true" pageId="12" pageNumber="527">Hellerenius lobifer</emphasis>
</taxonomicName>
in New Caledonia mainland.
@ -193,33 +196,33 @@ in New Caledonia mainland.
<subSubSection id="C3536542BF42FFF3FF0C5C85FD15F771" pageId="13" pageNumber="528" type="discussion">
<paragraph id="8BF636C9BF42FFF3FF0C5C85FD15F771" blockId="13.[151,1436,151,285]" pageId="13" pageNumber="528">
<emphasis id="B93DEADBBF42FFF3FF0C5C85FE89F6DC" bold="true" box="[199,335,151,176]" pageId="13" pageNumber="528">Comments.</emphasis>
<taxonomicName id="4C494D4ABF42FFF3FE935C8AFDEBF6DC" box="[344,557,152,176]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="528" phylum="Tracheophyta" rank="species" species="lobifer">
<taxonomicName id="4C494D4ABF42FFF3FE935C8AFDEBF6DC" authorityName="Wanat" authorityYear="2001" box="[344,557,152,176]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="528" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF42FFF3FE935C8AFDEBF6DC" box="[344,557,152,176]" italics="true" pageId="13" pageNumber="528">Hellerenius lobifer</emphasis>
</taxonomicName>
turns out to be a much less common species in
<collectingCountry id="F35E7659BF42FFF3FB9D5C8AFAC3F6DC" box="[1110,1285,151,176]" name="New Caledonia" pageId="13" pageNumber="528">New Caledonia</collectingCountry>
than the two other species described below, namely
<taxonomicName id="4C494D4ABF42FFF3FDAF5CAFFD36F6B8" box="[612,752,188,212]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="528" phylum="Tracheophyta" rank="species" species="plumipes" status="sp. nov.">
<taxonomicName id="4C494D4ABF42FFF3FDAF5CAFFD36F6B8" authorityName="Wanat" authorityYear="2013" box="[612,752,188,212]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="528" phylum="Arthropoda" rank="species" species="plumipes" status="sp. nov.">
<emphasis id="B93DEADBBF42FFF3FDAF5CAFFDBEF6B8" box="[612,632,189,212]" italics="true" pageId="13" pageNumber="528">H</emphasis>
.
<emphasis id="B93DEADBBF42FFF3FD405CAEFD36F6B8" box="[651,752,188,212]" italics="true" pageId="13" pageNumber="528">plumipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57A0BF42FFF3FD375CAFFC9EF6B8" box="[764,856,189,212]" pageId="13" pageNumber="528" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C494D4ABF42FFF3FC535CAFFBCBF6B8" box="[920,1037,188,212]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="528" phylum="Tracheophyta" rank="species" species="tibialis" status="sp. nov.">
<taxonomicName id="4C494D4ABF42FFF3FC535CAFFBCBF6B8" authorityName="Wanat" authorityYear="2013" box="[920,1037,188,212]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="528" phylum="Arthropoda" rank="species" species="tibialis" status="sp. nov.">
<emphasis id="B93DEADBBF42FFF3FC535CAFFC6AF6B8" box="[920,940,189,212]" italics="true" pageId="13" pageNumber="528">H</emphasis>
.
<emphasis id="B93DEADBBF42FFF3FC745CAEFBCBF6B8" box="[959,1037,188,212]" italics="true" pageId="13" pageNumber="528">tibialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57A0BF42FFF3FBD25CAFFBB2F6B8" box="[1049,1140,189,212]" pageId="13" pageNumber="528" rank="species">sp. nov.</taxonomicNameLabel>
It also seems to express different phenology being mostly collected between January and October, while both
<taxonomicName id="4C494D4ABF42FFF3FBA35CF3FB37F694" box="[1128,1265,224,248]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="528" phylum="Tracheophyta" rank="species" species="plumipes">
<taxonomicName id="4C494D4ABF42FFF3FBA35CF3FB37F694" authorityName="Wanat" authorityYear="2013" box="[1128,1265,224,248]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="528" phylum="Arthropoda" rank="species" species="plumipes">
<emphasis id="B93DEADBBF42FFF3FBA35CF3FBBAF694" box="[1128,1148,225,248]" italics="true" pageId="13" pageNumber="528">H</emphasis>
.
<emphasis id="B93DEADBBF42FFF3FB475CF2FB37F694" box="[1164,1265,224,248]" italics="true" pageId="13" pageNumber="528">plumipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D4ABF42FFF3FAE15CF3FA5DF694" box="[1322,1435,224,248]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="13" pageNumber="528" phylum="Tracheophyta" rank="species" species="tibialis">
<taxonomicName id="4C494D4ABF42FFF3FAE15CF3FA5DF694" authorityName="Wanat" authorityYear="2013" box="[1322,1435,224,248]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="528" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B93DEADBBF42FFF3FAE15CF3FAF8F694" box="[1322,1342,225,248]" italics="true" pageId="13" pageNumber="528">H</emphasis>
.
<emphasis id="B93DEADBBF42FFF3FA855CF2FA5DF694" box="[1358,1435,224,248]" italics="true" pageId="13" pageNumber="528">tibialis</emphasis>

View file

@ -1,46 +1,47 @@
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<treatment id="03E087DFBF4CFFFDFF5C5C85FAE0F121" ID-DOI="http://doi.org/10.5281/zenodo.6153098" ID-GBIF-Taxon="119578258" ID-Zenodo-Dep="6153098" LSID="urn:lsid:plazi:treatment:03E087DFBF4CFFFDFF5C5C85FAE0F121" httpUri="http://treatment.plazi.org/id/03E087DFBF4CFFFDFF5C5C85FAE0F121" lastPageNumber="518" pageId="3" pageNumber="518">
<subSubSection id="C3536542BF4CFFFDFF5C5C85FEF2F6BF" pageId="3" pageNumber="518" type="nomenclature">
<paragraph id="8BF636C9BF4CFFFDFF5C5C85FDCEF6DE" blockId="3.[151,520,151,211]" box="[151,520,151,178]" pageId="3" pageNumber="518">
<heading id="D0BE81A5BF4CFFFDFF5C5C85FDCEF6DE" bold="true" box="[151,520,151,178]" fontSize="11" level="1" pageId="3" pageNumber="518" reason="1">
<emphasis id="B93DEADBBF4CFFFDFF5C5C85FDCEF6DE" bold="true" box="[151,520,151,178]" pageId="3" pageNumber="518">
<taxonomicName id="4C494D4ABF4CFFFDFF5C5C85FE64F6DD" box="[151,418,151,177]" class="Insecta" family="Apionidae" genus="Archellerenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="species" species="giustoi" status="sp. nov.">
<taxonomicName id="4C494D4ABF4CFFFDFF5C5C85FE64F6DD" authorityName="Wanat" authorityYear="2013" box="[151,418,151,177]" class="Insecta" family="Brentidae" genus="Archellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="species" species="giustoi" status="sp. nov.">
<emphasis id="B93DEADBBF4CFFFDFF5C5C85FE64F6DD" bold="true" box="[151,418,151,177]" italics="true" pageId="3" pageNumber="518">Archellerenius giustoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57A0BF4CFFFDFE615C8AFDCEF6DE" box="[426,520,152,178]" pageId="3" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
@ -49,9 +50,9 @@
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF5C5CA8FEF2F6BF" blockId="3.[151,520,151,211]" box="[151,308,186,211]" pageId="3" pageNumber="518">
(
<figureCitation id="13722A4CBF4CFFFDFF545CA8FF21F6BF" box="[159,231,186,211]" captionStart="FIGURES 1 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="3" pageNumber="518">Figs 3</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFF545CA8FF21F6BF" box="[159,231,186,211]" captionStart="FIGURES 1 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." figureDoi="http://doi.org/10.5281/zenodo.249083" httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="3" pageNumber="518">Figs 3</figureCitation>
,
<figureCitation id="13722A4CBF4CFFFDFF385CA9FEEAF6BF" box="[243,300,187,211]" captionStart-0="FIGURES 9 20" captionStart-1="FIGURES 21 32" captionStart-2="FIGURES 33 45" captionStartId-0="6.[151,264,1953,1976]" captionStartId-1="7.[151,264,1850,1873]" captionStartId-2="8.[151,264,1923,1946]" captionTargetBox-0="[151,1431,193,1930]" captionTargetBox-1="[191,1396,200,1822]" captionTargetBox-2="[165,1408,193,1864]" captionTargetId-0="figure@6.[151,1436,193,1932]" captionTargetId-1="figure@7.[191,1396,193,1823]" captionTargetId-2="figure@8.[165,1409,193,1876]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." captionText-1="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." captionText-2="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri-0="https://zenodo.org/record/249084/files/figure.png" httpUri-1="https://zenodo.org/record/249085/files/figure.png" httpUri-2="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">937</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFF385CA9FEEAF6BF" box="[243,300,187,211]" captionStart-0="FIGURES 9 20" captionStart-1="FIGURES 21 32" captionStart-2="FIGURES 33 45" captionStartId-0="6.[151,264,1953,1976]" captionStartId-1="7.[151,264,1850,1873]" captionStartId-2="8.[151,264,1923,1946]" captionTargetBox-0="[151,1431,193,1930]" captionTargetBox-1="[191,1396,200,1822]" captionTargetBox-2="[165,1408,193,1864]" captionTargetId-0="figure@6.[151,1436,193,1932]" captionTargetId-1="figure@7.[191,1396,193,1823]" captionTargetId-2="figure@8.[165,1409,193,1876]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." captionText-1="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." captionText-2="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.249084" figureDoi-1="http://doi.org/10.5281/zenodo.249085" figureDoi-2="http://doi.org/10.5281/zenodo.249086" httpUri-0="https://zenodo.org/record/249084/files/figure.png" httpUri-1="https://zenodo.org/record/249085/files/figure.png" httpUri-2="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">937</figureCitation>
)
</paragraph>
</subSubSection>
@ -136,43 +137,43 @@ long; antennal scape entirely testaceous.
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C5EA1FF18F5A6" blockId="3.[151,1437,258,1870]" pageId="3" pageNumber="518">
Rostrum shaped as in
<figureCitation id="13722A4CBF4CFFFDFE775EA0FDFCF4A7" box="[444,570,690,715]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Figs 1316</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFE775EA0FDFCF4A7" box="[444,570,690,715]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Figs 1316</figureCitation>
, at base finely punctured, rugose at sides; prorostrum shiny, with microscopic punctures at least basally. Antennal club as long as 6 distal funicular segments combined. Head: frons usually with shallow V-like fovea; vertex evenly flattened throughout; subocular tooth usually weakly expressed in head profile. Antennae (
<figureCitation id="13722A4CBF4CFFFDFEDA5F33FEA3F556" box="[273,357,801,826]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Fig. 19</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFEDA5F33FEA3F556" box="[273,357,801,826]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Fig. 19</figureCitation>
): length/width ratios: scape 6.58.0, funicular segments: 1st 1.31.5, 2nd ca. 2.5, 6th7th 1.0, scape as long or longer than 6 basal funicular segments combined. Pronotum weakly rounded at sides (
<figureCitation id="13722A4CBF4CFFFDFA895F57FA48F532" box="[1346,1422,837,862]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Fig.12</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFA895F57FA48F532" box="[1346,1422,837,862]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Fig.12</figureCitation>
); hypomeron acutely prominent at posterior margin. Elytra moderately convex (
<figureCitation id="13722A4CBF4CFFFDFBF55F7BFB7FF5EE" box="[1086,1209,873,898]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Figs 911</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFBF55F7BFB7FF5EE" box="[1086,1209,873,898]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Figs 911</figureCitation>
); striae superficial; intervals flat, finely microreticulate and shiny. Fore leg as in
<figureCitation id="13722A4CBF4CFFFDFCF75F9FFC7DF5CA" box="[828,955,909,934]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Figs 18, 20</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFCF75F9FFC7DF5CA" box="[828,955,909,934]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Figs 18, 20</figureCitation>
. Abdomen (ventrites 15) 1.2× longer than broad.
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C5FC7FD1DF38A" blockId="3.[151,1437,258,1870]" pageId="3" pageNumber="518">
Male. Antennal club about 2.5× as long as wide. Abdominal ventrites narrower (
<figureCitation id="13722A4CBF4CFFFDFB9B5FC7FB62F582" box="[1104,1188,981,1006]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 25</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFB9B5FC7FB62F582" box="[1104,1188,981,1006]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 25</figureCitation>
); 5th ventrite twice as wide as long, broadly truncate, moderately and evenly convex, middle of apical margin with sparse fringe of white setae (
<figureCitation id="13722A4CBF4CFFFDFF14580FFEF2F25A" box="[223,308,1053,1078]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 26</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFF14580FFEF2F25A" box="[223,308,1053,1078]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 26</figureCitation>
). Pygidium: apical third exposed, strongly declivitous, with distinct marginal rim; separating sulcus obsolescent (
<figureCitation id="13722A4CBF4CFFFDFEE25853FE6DF236" box="[297,427,1089,1114]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Figs 23, 24</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFEE25853FE6DF236" box="[297,427,1089,1114]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Figs 23, 24</figureCitation>
). Sternite VIII with distinct carinae. Sternite IX with untoothed apodeme, ca. 1.6× as long as basal fork (
<figureCitation id="13722A4CBF4CFFFDFEF15877FE57F212" box="[314,401,1125,1150]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">Fig. 35</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFEF15877FE57F212" box="[314,401,1125,1150]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">Fig. 35</figureCitation>
). Tegminal apodeme strongly expanded in apical third; tegminal plate 1.31.4× as long as broad, with lateral arms much thinner than adjoining arms of basal piece (
<figureCitation id="13722A4CBF4CFFFDFC2F589BFBAFF2CE" box="[996,1129,1161,1186]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">Figs 36, 37</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFC2F589BFBAFF2CE" box="[996,1129,1161,1186]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">Figs 36, 37</figureCitation>
); membranous lobes short, rounded, apically extended into narrow, acute processes, these microsetose apically; suprafenestral sclerites half as long as fenestrae, acutely produced along internal margins; sensillae indistinct; macrochaetae 68, short, hardly exceeding acute sclerotized projections of parameroid lobes; membrane beneath fenestrae sclerotized equally to prostegium; postfenestral sclerite bar-like; prostegium long, deeply incised laterally, divided on its 0.60.7 length. Penis: apodemes as long as pedon, strongly expanded apicad; tectum reaching apical fourth of pedon; pedon with rounded, strongly upcurved apex (
<figureCitation id="13722A4CBF4CFFFDFDDE5973FD50F316" box="[533,662,1377,1402]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">Figs 33, 34</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFDDE5973FD50F316" box="[533,662,1377,1402]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="3" pageNumber="518">Figs 33, 34</figureCitation>
), ventrally membranous in basal half; endophallus with enclosed part densely microspinose in basal third, with sharp and straight distal border-line separating it from big, simply membranous terminal process, the latter with paired basal lobes covering entrance of ejaculatory duct, microplates in sub-orificial region extremely fine and indistinct.
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C59E3FEF1F08E" blockId="3.[151,1437,258,1870]" pageId="3" pageNumber="518">
Female. Antennal club more obtuse, 2.02.2× as long as wide (
<figureCitation id="13722A4CBF4CFFFDFC4259E3FC1AF066" box="[905,988,1521,1546]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Fig. 19</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFC4259E3FC1AF066" box="[905,988,1521,1546]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="3" pageNumber="518">Fig. 19</figureCitation>
). Abdominal ventrites shaped as in
<figureCitation id="13722A4CBF4CFFFDFAA059E3FF26F042" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Figs 27, 28</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFAA059E3FF26F042" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Figs 27, 28</figureCitation>
. Tergite VII short, broadly rounded (
<figureCitation id="13722A4CBF4CFFFDFDB45A07FD12F042" box="[639,724,1557,1582]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 30</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFDB45A07FD12F042" box="[639,724,1557,1582]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 30</figureCitation>
). Tergite VIII transverse, largely membranous, with narrow but complete marginal sclerotisation. Sternite VIII (spiculum ventrale) relatively short, slightly widened in middle part, apically with triangular expansion (
<figureCitation id="13722A4CBF4CFFFDFDED5A4FFDBDF01A" box="[550,635,1629,1654]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 32</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFDED5A4FFDBDF01A" box="[550,635,1629,1654]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 32</figureCitation>
). Styli stout and short, hemispherical, with several apical setae; coxites subtruncate apically. Foretube darkened, with dense long microtrichia. Bursa copulatrix relatively large, with corrugated longitudinal folds. Spermatheca: corpus with hook-like process receiving relatively thick spermathecal duct (
<figureCitation id="13722A4CBF4CFFFDFF1D5ADBFEECF08E" box="[214,298,1737,1762]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 31</figureCitation>
<figureCitation id="13722A4CBF4CFFFDFF1D5ADBFEECF08E" box="[214,298,1737,1762]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="3" pageNumber="518">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="8BF636C9BF4CFFFDFF0C5AFFFBB2F16A" blockId="3.[151,1437,258,1870]" box="[199,1140,1773,1798]" pageId="3" pageNumber="518">Morphological indices in Tab. 1. For other characters see generic description above.</paragraph>
@ -181,9 +182,11 @@ Female. Antennal club more obtuse, 2.02.2× as long as wide (
<paragraph id="8BF636C9BF4CFFFDFF0C5B03FAE0F121" blockId="3.[151,1437,258,1870]" pageId="3" pageNumber="518">
<emphasis id="B93DEADBBF4CFFFDFF0C5B03FE88F146" bold="true" box="[199,334,1809,1834]" pageId="3" pageNumber="518">Etymology.</emphasis>
This remarkable and rare species is named to honor Mr. Carlo Giusto, an enthusiastic student of
<taxonomicName id="4C494D4ABF4CFFFDFF5C5B27FEC9F122" box="[151,271,1845,1870]" class="Insecta" family="Apionidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="family">Apionidae</taxonomicName>
<taxonomicName id="4C494D4ABF4CFFFDFF5C5B27FEC9F122" box="[151,271,1845,1870]" class="Insecta" family="Apionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="family">Apionidae</taxonomicName>
, for his passion for these inconspicuous weevils, and shared interest in the genus
<taxonomicName id="4C494D4ABF4CFFFDFB555B27FAE6F121" box="[1182,1312,1845,1869]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="518" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4CFFFDFB555B27FAE6F121" box="[1182,1312,1845,1869]" italics="true" pageId="3" pageNumber="518">Hellerenius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>

View file

@ -1,39 +1,40 @@
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@ -41,7 +42,7 @@
<heading id="D0BE81A5BF4DFFFCFF5C5C8AFDD0F6DE" bold="true" box="[151,534,151,178]" fontSize="11" level="1" pageId="2" pageNumber="517" reason="1">
<emphasis id="B93DEADBBF4DFFFCFF5C5C8AFDD0F6DE" bold="true" box="[151,534,151,178]" pageId="2" pageNumber="517">
Genus
<taxonomicName id="4C494D4ABF4DFFFCFF265C85FE66F6DD" box="[237,416,151,177]" class="Insecta" family="Apionidae" genus="Archellerenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C494D4ABF4DFFFCFF265C85FE66F6DD" authorityName="Wanat" authorityYear="2013" box="[237,416,151,177]" class="Insecta" family="Brentidae" genus="Archellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93DEADBBF4DFFFCFF265C85FE66F6DD" bold="true" box="[237,416,151,177]" italics="true" pageId="2" pageNumber="517">Archellerenius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57A0BF4DFFFCFE635C8AFDD0F6DE" box="[424,534,152,178]" pageId="2" pageNumber="517" rank="genus">gen. nov.</taxonomicNameLabel>
@ -51,7 +52,7 @@ Genus
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<typeStatus id="54F2886BBF4DFFFCFF5C5CCDFF0FF698" box="[151,201,223,244]" pageId="2" pageNumber="517">Type</typeStatus>
species:
<taxonomicName id="4C494D4ABF4DFFFCFEED5CCDFDC2F698" box="[294,516,223,244]" class="Insecta" family="Apionidae" genus="Archellerenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="species" species="giustoi" status="sp. nov.">
<taxonomicName id="4C494D4ABF4DFFFCFEED5CCDFDC2F698" authorityName="Wanat" authorityYear="2013" box="[294,516,223,244]" class="Insecta" family="Brentidae" genus="Archellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="species" species="giustoi" status="sp. nov.">
<emphasis id="B93DEADBBF4DFFFCFEED5CCDFDC2F698" box="[294,516,223,244]" italics="true" pageId="2" pageNumber="517">Archellerenius giustoi</emphasis>
</taxonomicName>
<emphasis id="B93DEADBBF4DFFFCFDC05CCDFD82F698" bold="true" box="[523,580,223,244]" pageId="2" pageNumber="517">
@ -63,49 +64,67 @@ species:
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<emphasis id="B93DEADBBF4DFFFCFF5C5D33FED7F756" bold="true" box="[151,273,289,314]" pageId="2" pageNumber="517">Diagnosis.</emphasis>
The new, monotypic genus is closely related to
<taxonomicName id="4C494D4ABF4DFFFCFCE05D33FC6BF755" box="[811,941,289,313]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4DFFFCFCE05D33FC6BF755" box="[811,941,289,313]" italics="true" pageId="2" pageNumber="517">Hellerenius</emphasis>
</taxonomicName>
, based on the shared shape of rostrum, head and elytra, isodiametric scutellum, arrangement of elytral striae and sensory setae, femora devoid of pseudotrochanter, general structure of male endophallus and presence of internal lamina in the female rectum. It is distinct from
<taxonomicName id="4C494D4ABF4DFFFCFEFB5D9FFE74F7C9" box="[304,434,397,421]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4DFFFCFEFB5D9FFE74F7C9" box="[304,434,397,421]" italics="true" pageId="2" pageNumber="517">Hellerenius</emphasis>
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primarily in having the rostrum short and not differentiated sexually, smaller body size, polished elytra, well retained radial window of the wing, unmodified last abdominal ventrite, and much different structure of the ovipositor. It resembles the species of
<taxonomicName id="4C494D4ABF4DFFFCFCD35DC4FC08F781" authority="Wanat" authorityName="Wanat" authorityYear="2001" box="[792,974,470,493]" class="Insecta" family="Brentidae" genus="Zimmius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4DFFFCFCD35DC4FCBEF781" box="[792,888,470,493]" italics="true" pageId="2" pageNumber="517">Zimmius</emphasis>
Wanat, from which it can be distinguished by thinner rostrum devoid of extended and emarginate expansion of mesorostrum, legs with no trace of pseudotrochanter and with more robust true trochanters, elytra more regularly oval in dorsal outline, with complete striae, and the endophallus bearing a terminal, simply membranous extension. The superficially similar
Wanat
</taxonomicName>
, from which it can be distinguished by thinner rostrum devoid of extended and emarginate expansion of mesorostrum, legs with no trace of pseudotrochanter and with more robust true trochanters, elytra more regularly oval in dorsal outline, with complete striae, and the endophallus bearing a terminal, simply membranous extension. The superficially similar
<taxonomicName id="4C494D4ABF4DFFFCFF5C5E77FE40F411" authority="Wanat" authorityName="Wanat" authorityYear="2001" box="[151,390,613,637]" class="Insecta" family="Brentidae" genus="Caledonapion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4DFFFCFF5C5E77FEF0F411" box="[151,310,613,637]" italics="true" pageId="2" pageNumber="517">Caledonapion</emphasis>
Wanat is clearly distinct in its short and stout rostrum, coarsely punctured ventrally, and thin legs with a pseudotrochanter.
Wanat
</taxonomicName>
is clearly distinct in its short and stout rostrum, coarsely punctured ventrally, and thin legs with a pseudotrochanter.
</paragraph>
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<emphasis id="B93DEADBBF4DFFFCFF0C5EBFFE9FF4AA" bold="true" box="[199,345,685,710]" pageId="2" pageNumber="517">Description.</emphasis>
Body length about 3.0 mm. Integument black, glabrous, not shagreened, devoid of scale-like microsculpture. Rostrum not distinctly sexually dimorphic, straight, only slightly (1.11.2×) longer than pronotum, narrowed at base of prorostrum, the latter with vestigial ventral sulci in distal half; subgenal and latero-ventral sulci absent; scrobes entering head venter, their septum shallowly foveate; labrum with only lateral setae. Mouthparts as in
<taxonomicName id="4C494D4ABF4DFFFCFF725F29FEFCF53F" box="[185,314,827,851]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4DFFFCFF725F29FEFCF53F" box="[185,314,827,851]" italics="true" pageId="2" pageNumber="517">Hellerenius</emphasis>
</taxonomicName>
(
<figureCitation id="13722A4CBF4DFFFCFE845F29FE5FF53F" box="[335,409,827,852]" captionStart="FIGURES 1 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="2" pageNumber="517">Fig. 3</figureCitation>
<figureCitation id="13722A4CBF4DFFFCFE845F29FE5FF53F" box="[335,409,827,852]" captionStart="FIGURES 1 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." figureDoi="http://doi.org/10.5281/zenodo.249083" httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="2" pageNumber="517">Fig. 3</figureCitation>
). Head subisodiametric; frons gently elevated; vertex impunctate; head underside with completely irregular wrinkles near eyes; subocular setae absent. Antennae shorter than in
<taxonomicName id="4C494D4ABF4DFFFCFB585F4CFAD3F51A" box="[1171,1301,862,886]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4DFFFCFB585F4CFAD3F51A" box="[1171,1301,862,886]" italics="true" pageId="2" pageNumber="517">Hellerenius</emphasis>
</taxonomicName>
, not clearly pseudogeniculate; scape at most 2× as long as mesorostrum breadth; at least two terminal funicular segments isodiametric; club compact, 2.02.5× as long as broad, 0.80.9× as long as scape. Pronotum slightly elongate, with obsolete subapical constriction; disc with subbasal line at most finely traced; posterior lateral groove shallow, with minute spur at upper end; prosternum extremely short. Scutellum minute, isodiametric. Elytra oval, finely microreticulate, shiny, without extended caudal part, in profile regularly convex; humeral calli prominent; no trace of basal rim; striae narrow and weakly impressed, arrangement of inner striae apically indistinct and often irregular, outer striae finely punctured, narrow throughout; intervals all flat; sensory setae
<quantity id="4CB19B2CBF4DFFFCFBE65848FBB0F21E" box="[1069,1142,1113,1138]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="2" pageNumber="517" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">23 in</quantity>
row on 9th interval. Wing with radial window large and distinct; anal vein complex with 1A retained as paired rudiments, 2A without spur, 3A largely reduced (
<figureCitation id="13722A4CBF4DFFFCFEB658B2FE17F2D5" box="[381,465,1184,1209]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="2" pageNumber="517">Fig. 21</figureCitation>
<figureCitation id="13722A4CBF4DFFFCFEB658B2FE17F2D5" box="[381,465,1184,1209]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="2" pageNumber="517">Fig. 21</figureCitation>
). Mesoventrite distinctly angled in profile; mesepimeral sulcus narrow, sharply edged, impunctate. Metaventrite in middle ca. 1.5× longer than mesocoxa; rim of mesocoxae narrow but complete and well raised; intercoxal process elevated. Legs moderately long, unmodified in both sexes; femora weakly inflated, without pseudotrochanter, their bases and trochanters relatively robust; metafemur not exceeding elytral apex; tibiae straight, parallelsided, with extended and indistinct apical tuft of setae (
<figureCitation id="13722A4CBF4DFFFCFBDC593CFBA8F32B" box="[1047,1134,1326,1351]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="2" pageNumber="517">Fig. 20</figureCitation>
<figureCitation id="13722A4CBF4DFFFCFBDC593CFBA8F32B" box="[1047,1134,1326,1351]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="2" pageNumber="517">Fig. 20</figureCitation>
); tarsi slender; claws with short, blunt teeth (
<figureCitation id="13722A4CBF4DFFFCFEA85940FE70F306" box="[355,438,1362,1387]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="2" pageNumber="517">Fig. 17</figureCitation>
<figureCitation id="13722A4CBF4DFFFCFEA85940FE70F306" box="[355,438,1362,1387]" captionStart="FIGURES 9 20" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[151,1431,193,1930]" captionTargetId="figure@6.[151,1436,193,1932]" captionTargetPageId="6" captionText="FIGURES 9 20. Archellerenius giustoi, male (13, 15), female (9 12, 14, 17 20): 9) habitus, lateral view; 10) apex of elytra, postero-lateral view; 11) pronotum and elytra, dorsal view; 12) pronotum, dorsal view; 13, 14) head with rostrum, dorsal view; 15, 16) head with rostrum, lateral view; 17) tarsal claws; 18) protarsus; 19) antenna; 20) fore femur (part) and tibia." figureDoi="http://doi.org/10.5281/zenodo.249084" httpUri="https://zenodo.org/record/249084/files/figure.png" pageId="2" pageNumber="517">Fig. 17</figureCitation>
). Membrane between tergites VII and VIII with pouch-like process in both sexes.
</paragraph>
<paragraph id="8BF636C9BF4DFFFCFF0C5967FE38F0EA" blockId="2.[151,1436,289,2025]" pageId="2" pageNumber="517">
Male: Last abdominal ventrite (V) simply convex, unmodified. Pouch-like process of tergite VII membrane small, triangular, at base 45 × narrower than tergite VIII. Pygidium of exposed aspidapionine
<typeStatus id="54F2886BBF4DFFFCFB1B598BFAC5F3DD" box="[1232,1283,1433,1457]" pageId="2" pageNumber="517">type</typeStatus>
, with distinct marginal rim. Sternite VIII short and shallowly emarginate, Sternite IX with transverse apical plate. Tegmen fused, with distinctly forked basal piece; tegminal plate deeply notched, membranous lobes present; suprafenestral sclerites and postfenestral plate both short; macrochaetae short and fine; fenestrae enlarged, well margined; prostegium long, bifurcate. Penis with shortly dichotomous apodemes; tectum rod-like throughout; pedon asymmetrical; endophallus as in
<taxonomicName id="4C494D4ABF4DFFFCFDD85A58FD53F00E" box="[531,661,1610,1634]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4DFFFCFDD85A58FD53F00E" box="[531,661,1610,1634]" italics="true" pageId="2" pageNumber="517">Hellerenius</emphasis>
</taxonomicName>
with long, simply membranous terminal process sharply separated from microspinulose remainder.
</paragraph>
<paragraph id="8BF636C9BF4DFFFCFF0C5A83FB26F15B" blockId="2.[151,1436,289,2025]" pageId="2" pageNumber="517">
Female. Antennal insertion as in male. Fifth abdominal ventrite flattened, its laterosternites united to form narrow apical flange (
<figureCitation id="13722A4CBF4DFFFCFE6E5AA6FDC4F0A1" box="[421,514,1716,1741]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="2" pageNumber="517">Fig. 22</figureCitation>
<figureCitation id="13722A4CBF4DFFFCFE6E5AA6FDC4F0A1" box="[421,514,1716,1741]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="2" pageNumber="517">Fig. 22</figureCitation>
). Tergite VII regularly rounded. Tergite VIII narrowly but fully divided to hemitergites. Pouch-like process of tergite VII membrane very large, at base almost as broad as tergite VIII. Ovipositor short, with markedly divergent gonocoxites bearing stout, hemispherical styli. Spermathecal duct short, equally wide throughout. Rectum with thin ribbon-like internal lamina as in
<taxonomicName id="4C494D4ABF4DFFFCFC255B0DFBB6F15B" box="[1006,1136,1823,1847]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4DFFFCFC255B0DFBB6F15B" box="[1006,1136,1823,1847]" italics="true" pageId="2" pageNumber="517">Hellerenius</emphasis>
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(
<figureCitation id="13722A4CBF4DFFFCFBB45B0DFB12F15B" box="[1151,1236,1823,1848]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="2" pageNumber="517">Fig. 29</figureCitation>
<figureCitation id="13722A4CBF4DFFFCFBB45B0DFB12F15B" box="[1151,1236,1823,1848]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="2" pageNumber="517">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
@ -130,7 +149,9 @@ The stem
of the proposed new name is derived from the Greek adjective archaios (in Latin
<emphasis id="B93DEADBBF4DFFFCFF5C5BBFFF29F1A9" box="[151,239,1965,1989]" italics="true" pageId="2" pageNumber="517">archeos</emphasis>
)—old, ancient, combined with the name of the closely related genus
<taxonomicName id="4C494D4ABF4DFFFCFBD85BBFFB53F1A9" box="[1043,1173,1965,1989]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF4DFFFCFBD85BBFFB53F1A9" box="[1043,1173,1965,1989]" italics="true" pageId="2" pageNumber="517">Hellerenius</emphasis>
</taxonomicName>
, to underline the more plesiotypic set of characters.
</paragraph>
</subSubSection>

View file

@ -1,39 +1,40 @@
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<treatment id="03E087DFBF55FFE4FF5C5F32FA50F072" ID-DOI="http://doi.org/10.5281/zenodo.6153108" ID-GBIF-Taxon="119578254" ID-Zenodo-Dep="6153108" LSID="urn:lsid:plazi:treatment:03E087DFBF55FFE4FF5C5F32FA50F072" httpUri="http://treatment.plazi.org/id/03E087DFBF55FFE4FF5C5F32FA50F072" lastPageNumber="541" pageId="26" pageNumber="541">
<subSubSection id="C3536542BF55FFE4FF5C5F32FD2AF555" box="[151,748,799,826]" pageId="26" pageNumber="541" type="nomenclature">
@ -41,11 +42,13 @@
<heading id="D0BE81A5BF55FFE4FF5C5F32FD2AF555" bold="true" box="[151,748,799,826]" fontSize="11" level="1" pageId="26" pageNumber="541" reason="1">
<emphasis id="B93DEADBBF55FFE4FF5C5F32FD2AF555" bold="true" box="[151,748,799,826]" pageId="26" pageNumber="541">
Key to species of
<taxonomicName id="4C494D4ABF55FFE4FEA65F0DFDE6F555" box="[365,544,799,825]" class="Insecta" family="Apionidae" genus="Archellerenius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C494D4ABF55FFE4FEA65F0DFDE6F555" authorityName="Wanat" authorityYear="2013" box="[365,544,799,825]" class="Insecta" family="Brentidae" genus="Archellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF55FFE4FEA65F0DFDE6F555" bold="true" box="[365,544,799,825]" italics="true" pageId="26" pageNumber="541">Archellerenius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D4ABF55FFE4FD955F0DFD2AF555" box="[606,748,799,825]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF55FFE4FD955F0DFD2AF555" bold="true" box="[606,748,799,825]" italics="true" pageId="26" pageNumber="541">Hellerenius</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
@ -55,17 +58,17 @@ and
<paragraph id="8BF636C9BF55FFE4FF5C5F77FA5AF5DA" blockId="26.[151,1436,869,1566]" pageId="26" pageNumber="541">
<keyLead id="30B895FCBF55FFE4FF5C5F77FA5AF5DA" pageId="26" pageNumber="541">
1. Body length up to 3.0 mm. Rostrum in both sexes at most 1.30× as long as pronotum, less than 4.5× as long as broad. Elytra polished. Last abdominal ventrite simple, gently convex. Penis evidently asymmetrical (
<figureCitation id="13722A4CBF55FFE4FBE05F91FBB5F5F5" box="[1067,1139,899,921]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="26" pageNumber="541">Fig. 34</figureCitation>
<figureCitation id="13722A4CBF55FFE4FBE05F91FBB5F5F5" box="[1067,1139,899,921]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="26" pageNumber="541">Fig. 34</figureCitation>
). Ovipositor broadly triangular (
<figureCitation id="13722A4CBF55FFE4FF235FB2FEF6F5DA" box="[232,304,928,950]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="26" pageNumber="541">Fig. 32</figureCitation>
<figureCitation id="13722A4CBF55FFE4FF235FB2FEF6F5DA" box="[232,304,928,950]" captionStart="FIGURES 21 32" captionStartId="7.[151,264,1850,1873]" captionTargetBox="[191,1396,200,1822]" captionTargetId="figure@7.[191,1396,193,1823]" captionTargetPageId="7" captionText="FIGURES 21 32. Archellerenius giustoi, male (23 26), female (21, 22, 27 32): 21) metathoracic wing, enlarged radial cell to show distinct radial window (21 A); 22) apical flange of last abdominal ventrite, postero-lateral view; 23, 24) pygidium in dorsal (23) and lateral (24) view; 25 28) abdomina in ventral (25, 27) and lateral (26, 28) view (notice mite nymph attached to female 2 nd ventrite); 29) rectum; 30) tergite VII; 31) spermatheca and bursa copulatrix; 32) terminalia with enlarged ovipositor. Abbreviations: b, bursa; c, coxite, d, spermathecal duct, g, gonads; p, pygidium; rr, rectal ribbon-like internal lamina, s, spermatheca; st, stylus, sv, spiculum ventrale; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249085" httpUri="https://zenodo.org/record/249085/files/figure.png" pageId="26" pageNumber="541">Fig. 32</figureCitation>
). Genus
<taxonomicName id="4C494D4ABF55FFE4FE4F5FB2FDD1F5DA" box="[388,535,928,950]" class="Insecta" family="Apionidae" genus="Archellerenius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="genus" status="gen. nov.">
<taxonomicName id="4C494D4ABF55FFE4FE4F5FB2FDD1F5DA" authorityName="Wanat" authorityYear="2013" box="[388,535,928,950]" class="Insecta" family="Brentidae" genus="Archellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93DEADBBF55FFE4FE4F5FB2FDD1F5DA" bold="true" box="[388,535,928,950]" italics="true" pageId="26" pageNumber="541">Archellerenius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57A0BF55FFE4FDD75FB3FDB0F5DA" box="[540,630,929,950]" pageId="26" pageNumber="541" rank="genus">gen. nov.</taxonomicNameLabel>
....................................................
<emphasis id="B93DEADBBF55FFE4FB205FB2FA5AF5DA" bold="true" box="[1259,1436,928,950]" pageId="26" pageNumber="541">
<taxonomicName id="4C494D4ABF55FFE4FB205FB2FA8FF5DA" box="[1259,1353,928,950]" class="Insecta" family="Apionidae" genus="Archellerenius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="species" species="giustoi" status="sp. nov.">
<taxonomicName id="4C494D4ABF55FFE4FB205FB2FA8FF5DA" authorityName="Wanat" authorityYear="2013" box="[1259,1353,928,950]" class="Insecta" family="Brentidae" genus="Archellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="species" species="giustoi" status="sp. nov.">
<emphasis id="B93DEADBBF55FFE4FB205FB2FB3DF5DA" bold="true" box="[1259,1275,928,950]" italics="true" pageId="26" pageNumber="541">A</emphasis>
.
<emphasis id="B93DEADBBF55FFE4FACD5FB2FA8FF5DA" bold="true" box="[1286,1353,928,950]" italics="true" pageId="26" pageNumber="541">giustoi</emphasis>
@ -81,8 +84,11 @@ and
. Rostrum in male at least 1.35×, in female 1.8× as long as pronotum, its length/width ratio at least 5.0 in male,
<quantity id="4CB19B2CBF55FFE4FEFF5FC9FEAAF59D" box="[308,364,987,1009]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" pageId="26" pageNumber="541" unit="in" value="6.5">6.5 in</quantity>
female. Elytra weakly shiny to completely matt. Last abdominal ventrite with outstanding modifications. Penis symmetrical. Ovipositor narrow, parallel-sided. Genus
<taxonomicName id="4C494D4ABF55FFE4FD3B5FEAFC6BF262" authority="Wanat." authorityName="Wanat." box="[752,941,1016,1038]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF55FFE4FD3B5FEAFCA2F262" bold="true" box="[752,868,1016,1038]" italics="true" pageId="26" pageNumber="541">Hellerenius</emphasis>
Wanat........................................ 2
Wanat.
</taxonomicName>
....................................... 2
</keyLead>
</paragraph>
</keyStep>
@ -91,7 +97,7 @@ Wanat........................................ 2
<keyLead id="30B895FCBF55FFE4FF5C5807FA5DF225" pageId="26" pageNumber="541">
2. Body length over 6.0 mm. Elytra dull black, with shagreened and completely matt intervals well contrasting with shiny and shallow striae. Rostrum in male over 1.7×, in female over 2.6× as long as pronotum................
<emphasis id="B93DEADBBF55FFE4FB075821FA5DF225" bold="true" box="[1228,1435,1075,1097]" pageId="26" pageNumber="541">
<taxonomicName id="4C494D4ABF55FFE4FB075821FA8CF225" box="[1228,1354,1075,1097]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="26" pageNumber="541" phylum="Tracheophyta" rank="species" species="monteithi" status="sp. nov.">
<taxonomicName id="4C494D4ABF55FFE4FB075821FA8CF225" authorityName="Wanat" authorityYear="2013" box="[1228,1354,1075,1097]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
<emphasis id="B93DEADBBF55FFE4FB075821FB19F225" bold="true" box="[1228,1247,1075,1097]" italics="true" pageId="26" pageNumber="541">H</emphasis>
.
<emphasis id="B93DEADBBF55FFE4FB205821FA8CF225" bold="true" box="[1259,1354,1075,1097]" italics="true" pageId="26" pageNumber="541">monteithi</emphasis>
@ -110,13 +116,13 @@ Wanat........................................ 2
3 Body length 4.0
<quantity id="4CB19B2CBF55FFE4FE435899FE13F2CD" box="[392,469,1163,1185]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="26" pageNumber="541" unit="mm" value="4.6">4.6 mm</quantity>
. Elytra bluish, highly convex, somewhat angulate on top in profile (
<figureCitation id="13722A4CBF55FFE4FBAE5899FB6BF2CD" box="[1125,1197,1163,1185]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="26" pageNumber="541">Fig. 47</figureCitation>
<figureCitation id="13722A4CBF55FFE4FBAE5899FB6BF2CD" box="[1125,1197,1163,1185]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." figureDoi="http://doi.org/10.5281/zenodo.249087" httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="26" pageNumber="541">Fig. 47</figureCitation>
); intervals with irregular transverse furrows or wrinkles. Last abdominal ventrite with paired raised lobes bordering median depression (
<figureCitation id="13722A4CBF55FFE4FAD558BAFA56F2D2" box="[1310,1424,1192,1214]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="26" pageNumber="541">Figs 54, 55</figureCitation>
<figureCitation id="13722A4CBF55FFE4FAD558BAFA56F2D2" box="[1310,1424,1192,1214]" captionStart="FIGURES 47 56" captionStartId="11.[151,264,1660,1683]" captionTargetBox="[163,1422,412,1628]" captionTargetId="figure@11.[151,1436,412,1640]" captionTargetPageId="11" captionText="FIGURES 47 56. Hellerenius lobifer, male (49, 51), female (47, 48, 50, 52 56): 47) pronotum and elytra, lateral view; 48) elytra, dorsal view; 49) protarsus; 50) antenna; 51) head with rostrum and pronotum, dorsal view; 52) head with rostrum, dorsal view; 44) rostrum base, lateral view; 54) apical flange of 5 th abdominal ventrite and corresponding tergite VII (t), posterolateral view; 55, 56) abdomen in ventral (55) and lateral (56) view." figureDoi="http://doi.org/10.5281/zenodo.249087" httpUri="https://zenodo.org/record/249087/files/figure.png" pageId="26" pageNumber="541">Figs 54, 55</figureCitation>
), much higher and setose in male. Penis subtruncate (
<figureCitation id="13722A4CBF55FFE4FD1A58D7FCD1F2B7" box="[721,791,1221,1243]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="26" pageNumber="541">Fig. 39</figureCitation>
<figureCitation id="13722A4CBF55FFE4FD1A58D7FCD1F2B7" box="[721,791,1221,1243]" captionStart="FIGURES 33 45" captionStartId="8.[151,264,1923,1946]" captionTargetBox="[165,1408,193,1864]" captionTargetId="figure@8.[165,1409,193,1876]" captionTargetPageId="8" captionText="FIGURES 33 45. Archellerenius giustoi, male (33 37), Hellerenius lobifer, male (38 42) and female (43 45): 33) penis with apodemes, lateral view; 34, 39) penis, dorsal view; 35, 38) sternites VIII and IX; 36, 41) tegminal plate and basal piece, lateral view; 37, 42) tegminal plate, dorsal view; 43) spermatheca; 44) tergite VII; 45) complete terminalia. Abbreviations: b, bursa; d, spermathecal duct; ed, ejaculatory duct; gl, spermathecal gland; r, rectum; s, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.249086" httpUri="https://zenodo.org/record/249086/files/figure.png" pageId="26" pageNumber="541">Fig. 39</figureCitation>
).......................................
<taxonomicName id="4C494D4ABF55FFE4FB3358D4FA5CF2B7" authority="Wanat" authorityName="Wanat" box="[1272,1434,1222,1244]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="26" pageNumber="541" phylum="Tracheophyta" rank="species" species="lobifer">
<taxonomicName id="4C494D4ABF55FFE4FB3358D4FA5CF2B7" authority="Wanat" authorityName="Wanat" authorityYear="2001" box="[1272,1434,1222,1244]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF55FFE4FB3358D4FA9FF2B0" bold="true" box="[1272,1369,1222,1244]" pageId="26" pageNumber="541">
<emphasis id="B93DEADBBF55FFE4FB3358D4FACDF2B0" bold="true" box="[1272,1291,1222,1244]" italics="true" pageId="26" pageNumber="541">H</emphasis>
.
@ -131,9 +137,9 @@ Wanat
-. Body length
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. Elytra with evident bronze tinge, not much extended apically, in profile more roundly convex (
<figureCitation id="13722A4CBF55FFE4FABA58F1FF3DF37A" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="26" pageNumber="541">Figs 94</figureCitation>
<figureCitation id="13722A4CBF55FFE4FABA58F1FF3DF37A" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249088" httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="26" pageNumber="541">Figs 94</figureCitation>
, 123); intervals evenly flattened. Last abdominal ventrite convex in both sexes. Penis pointed (
<figureCitation id="13722A4CBF55FFE4FB435912FB18F37A" box="[1160,1246,1280,1302]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="26" pageNumber="541">Figs 118</figureCitation>
<figureCitation id="13722A4CBF55FFE4FB435912FB18F37A" box="[1160,1246,1280,1302]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="26" pageNumber="541">Figs 118</figureCitation>
, 148)..........4
</keyLead>
</paragraph>
@ -145,7 +151,7 @@ Wanat
<collectingCountry id="F35E7659BF55FFE4FF2A590FFED7F35F" box="[225,273,1309,1331]" name="India" pageId="26" pageNumber="541">Hind</collectingCountry>
tibiae sinuate, compressed and twisted apically (Figs 132134). Coxae, trochanters and femora asetose. Male: underside of rostrum bare; all tibiae mucronate; last abdominal ventrite with large, rectangular protuberance (Fig.138); penis (excluding apodemes) more than 5× as long as broad (Fig. 148); tegminal parameroid lobes with dense tuft of long and numerous setae (Fig. 145). Female: last abdominal ventrite with triangular elevation..............................
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<taxonomicName id="4C494D4ABF55FFE4FB295964FA85F3E0" authorityName="Wanat" authorityYear="2013" box="[1250,1347,1398,1420]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="species" species="tibialis" status="sp. nov.">
<emphasis id="B93DEADBBF55FFE4FB295964FB33F3E0" bold="true" box="[1250,1269,1398,1420]" italics="true" pageId="26" pageNumber="541">H</emphasis>
.
<emphasis id="B93DEADBBF55FFE4FACB5964FA85F3E0" bold="true" box="[1280,1347,1398,1420]" italics="true" pageId="26" pageNumber="541">tibialis</emphasis>
@ -157,12 +163,12 @@ tibiae sinuate, compressed and twisted apically (Figs 132134). Coxae, trochan
<paragraph id="8BF636C9BF55FFE4FF5C5986FA50F072" blockId="26.[151,1436,869,1566]" pageId="26" pageNumber="541">
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-. All tibiae straight, unmodified (Figs 106108). Underside of coxae, trochanters and bases of femora with dense protruding setae, forming distinct fringe in male. Male: underside of rostrum protrudingly pilose; all tibiae unarmed; last abdominal ventrite roundly convex, with densely setose depression (Fig. 109); penis (excluding apodemes) less than 3× as long as broad (
<figureCitation id="13722A4CBF55FFE4FF2359F9FEF3F06D" box="[232,309,1515,1537]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="26" pageNumber="541">Fig.118</figureCitation>
<figureCitation id="13722A4CBF55FFE4FF2359F9FEF3F06D" box="[232,309,1515,1537]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="26" pageNumber="541">Fig.118</figureCitation>
); parameroid lobes with few inconspicuous setae (
<figureCitation id="13722A4CBF55FFE4FCD259F9FCAAF06D" box="[793,876,1515,1537]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="26" pageNumber="541">Fig. 117</figureCitation>
<figureCitation id="13722A4CBF55FFE4FCD259F9FCAAF06D" box="[793,876,1515,1537]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="26" pageNumber="541">Fig. 117</figureCitation>
). Female: last abdominal ventrite regularly convex.........................................................................................
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<taxonomicName id="4C494D4ABF55FFE4FB005A1AFA85F072" box="[1227,1347,1544,1566]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="26" pageNumber="541" phylum="Tracheophyta" rank="species" species="plumipes" status="sp. nov.">
<taxonomicName id="4C494D4ABF55FFE4FB005A1AFA85F072" authorityName="Wanat" authorityYear="2013" box="[1227,1347,1544,1566]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="species" species="plumipes" status="sp. nov.">
<emphasis id="B93DEADBBF55FFE4FB005A1AFB18F072" bold="true" box="[1227,1246,1544,1566]" italics="true" pageId="26" pageNumber="541">H</emphasis>
.
<emphasis id="B93DEADBBF55FFE4FB215A1AFA85F072" bold="true" box="[1258,1347,1544,1566]" italics="true" pageId="26" pageNumber="541">plumipes</emphasis>

View file

@ -1,46 +1,47 @@
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<emphasis id="B93DEADBBF59FFE8FF5C5EA1FEBBF4A1" bold="true" box="[151,381,691,717]" italics="true" pageId="22" pageNumber="537">Hellerenius tibialis</emphasis>
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@ -67,7 +68,7 @@ material.
c)
<emphasis id="B93DEADBBF59FFE8FED75F51FE27F536" box="[284,481,835,859]" italics="true" pageId="22" pageNumber="537">
Helleristus
<taxonomicName id="4C494D4ABF59FFE8FE555F51FE27F536" box="[414,481,835,858]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="22" pageNumber="537" phylum="Tracheophyta" rank="species" species="novae">novae</taxonomicName>
<taxonomicName id="4C494D4ABF59FFE8FE555F51FE27F536" box="[414,481,835,858]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="537" phylum="Arthropoda" rank="species" species="novae">novae</taxonomicName>
</emphasis>
-
<emphasis id="B93DEADBBF59FFE8FE205F51FDA0F537" box="[491,614,835,859]" italics="true" pageId="22" pageNumber="537">caledoniae</emphasis>
@ -88,7 +89,7 @@ Helleristus
Province Nord: Houadou [=Houaïlou, -21.287/165.634],
<date id="FFF71009BF59FFE8FCAF5F99FC3AF5CF" box="[868,1020,907,931]" pageId="22" pageNumber="537" value="1914-07-28">28 VII 1914</date>
, 1 ♀, leg. P.Montague, labelled as allotype of
<taxonomicName id="4C494D4ABF59FFE8FED25FBDFE46F5AA" box="[281,384,943,966]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="22" pageNumber="537" phylum="Tracheophyta" rank="species" species="novae">
<taxonomicName id="4C494D4ABF59FFE8FED25FBDFE46F5AA" box="[281,384,943,966]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="537" phylum="Arthropoda" rank="species" species="novae">
<emphasis id="B93DEADBBF59FFE8FED25FBDFEEBF5AA" box="[281,301,943,966]" italics="true" pageId="22" pageNumber="537">H</emphasis>
.
<emphasis id="B93DEADBBF59FFE8FEF65FBDFE46F5AA" box="[317,384,943,966]" italics="true" pageId="22" pageNumber="537">novae</emphasis>
@ -153,10 +154,11 @@ XII 1990,
,
<specimenCount id="9D4FFD40BF59FFE8FEFD58E1FEA5F366" box="[310,355,1267,1291]" pageId="22" pageNumber="537" type="generic">1 ♂</specimenCount>
, from
<taxonomicName id="4C494D4ABF59FFE8FE6558E1FDEFF367" box="[430,553,1267,1291]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="22" pageNumber="537" phylum="Tracheophyta" rank="genus">
<taxonomicName id="4C494D4ABF59FFE8FE6558E1FD95F366" box="[430,595,1267,1291]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="22" pageNumber="537" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93DEADBBF59FFE8FE6558E1FDEFF367" box="[430,553,1267,1291]" italics="true" pageId="22" pageNumber="537">Hedycarya</emphasis>
sp.
</taxonomicName>
sp.; (Povila), -20.9528/165.29135,
; (Povila), -20.9528/165.29135,
<quantity id="4CB19B2CBF59FFE8FC7058E1FBC3F366" box="[955,1029,1267,1291]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="22" pageNumber="537" unit="m" value="400.0">400 m</quantity>
, rainforest,
<date id="FFF71009BF59FFE8FB4058E1FAC9F367" box="[1163,1295,1267,1291]" pageId="22" pageNumber="537" value="2008-11-22">22 XI 2008</date>
@ -247,10 +249,11 @@ S/165.19.12E [-21.1556/165.32],
, at light, 1 ♀,
<date id="FFF71009BF59FFE8FF5C5AB1FED8F0D7" box="[151,286,1699,1723]" pageId="22" pageNumber="537" value="2008-11-29">29 XI 2008</date>
, beating fruiting
<taxonomicName id="4C494D4ABF59FFE8FE2E5AB1FDA6F0D7" box="[485,608,1699,1723]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="22" pageNumber="537" phylum="Tracheophyta" rank="genus">
<taxonomicName id="4C494D4ABF59FFE8FE2E5AB1FD4CF0D6" box="[485,650,1699,1723]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="22" pageNumber="537" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93DEADBBF59FFE8FE2E5AB1FDA6F0D7" box="[485,608,1699,1723]" italics="true" pageId="22" pageNumber="537">Hedycarya</emphasis>
sp.
</taxonomicName>
sp.,
,
<specimenCount id="9D4FFD40BF59FFE8FD525AB1FD00F0D6" box="[665,710,1699,1723]" pageId="22" pageNumber="537" type="generic">2 ♂</specimenCount>
1 ♀ - all leg. MW (MW). Thoven,
<geoCoordinate id="EE7D500EBF59FFE8FBA25AB0FB70F0D7" box="[1129,1206,1698,1723]" direction="south" orientation="latitude" pageId="22" pageNumber="537" precision="555" value="-20.47">20.47S</geoCoordinate>
@ -261,10 +264,11 @@ sp.,
,
<date id="FFF71009BF59FFE8FEA05AD4FE12F0B3" box="[363,468,1734,1759]" pageId="22" pageNumber="537" value="2004-02-05">5 II 2004</date>
, from
<taxonomicName id="4C494D4ABF59FFE8FDD75AD4FD51F0B2" box="[540,663,1734,1758]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="22" pageNumber="537" phylum="Tracheophyta" rank="genus">
<taxonomicName id="4C494D4ABF59FFE8FDD75AD4FD79F0B2" box="[540,703,1734,1758]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="22" pageNumber="537" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93DEADBBF59FFE8FDD75AD4FD51F0B2" box="[540,663,1734,1758]" italics="true" pageId="22" pageNumber="537">Hedycarya</emphasis>
sp.
</taxonomicName>
sp.,
,
<specimenCount id="9D4FFD40BF59FFE8FD005AD5FCC4F0B2" box="[715,770,1735,1759]" pageId="22" pageNumber="537" type="generic">11 ♂</specimenCount>
3 ♀. Cascade de Tao,
<geoCoordinate id="EE7D500EBF59FFE8FC375AD4FB8FF0B3" box="[1020,1097,1734,1759]" direction="south" orientation="latitude" pageId="22" pageNumber="537" precision="555" value="-20.33">20.33S</geoCoordinate>
@ -322,14 +326,14 @@ from gate), -21.58383/165.79667,
<specimenCount id="9D4FFD40BF59FFE8FAF45BD1FAAFF1B6" box="[1343,1385,1987,2011]" pageId="22" pageNumber="537" type="generic">1 ♂</specimenCount>
- all
<emphasis id="B93DEADBBF58FFE9FF5C5B4CFEB7F118" bold="true" box="[151,369,1886,1909]" pageId="23" pageNumber="538">FIGURES 123137.</emphasis>
<taxonomicName id="4C494D4ABF58FFE9FEB05B4DFD87F118" box="[379,577,1887,1908]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="23" pageNumber="538" phylum="Tracheophyta" rank="species" species="tibialis">
<taxonomicName id="4C494D4ABF58FFE9FEB05B4DFD87F118" authorityName="Wanat" authorityYear="2013" box="[379,577,1887,1908]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="538" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B93DEADBBF58FFE9FEB05B4DFD87F118" box="[379,577,1887,1908]" italics="true" pageId="23" pageNumber="538">Hellerenius tibialis</emphasis>
</taxonomicName>
, male (123125, 130132, 135137), female (126129, 133, 134): 123) habitus, lateral view; 124) elytra, dorsal view; 125, 126) antenna; 127) apex of elytra; 128) head with rostrum, lateral view; 129) head with rostrum and pronotum, dorsal view; 130) head with rostrum, dorsal view; 131) rostrum (part), ventral view; 132134) metatibia in lateral (132, 133) and dorsal (134) view; 135) fore femur (part) and tibia, enlarged apex of protibia (135A); 136) protarsus; 137) tarsal claws.
</paragraph>
<paragraph id="8BF636C9BF57FFE6FF5C5B31FD46F1D8" blockId="24.[151,1436,1827,1972]" pageId="24" pageNumber="539">
<emphasis id="B93DEADBBF57FFE6FF5C5B31FEABF155" bold="true" box="[151,365,1827,1850]" pageId="24" pageNumber="539">FIGURES 138148.</emphasis>
<taxonomicName id="4C494D4ABF57FFE6FEB85B36FDF2F155" box="[371,564,1828,1849]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="24" pageNumber="539" phylum="Tracheophyta" rank="species" species="tibialis">
<taxonomicName id="4C494D4ABF57FFE6FEB85B36FDF2F155" authorityName="Wanat" authorityYear="2013" box="[371,564,1828,1849]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="539" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B93DEADBBF57FFE6FEB85B36FDF2F155" box="[371,564,1828,1849]" italics="true" pageId="24" pageNumber="539">Hellerenius tibialis</emphasis>
</taxonomicName>
, male (138, 142, 144148), female (139141, 143): 138, 139) all abdominal ventrites, ventral view; 140) rectum; 141) tergite VII; 142) shaped endophallus in distilled water; 143)spermatheca; 144, 145) tegminal plate and basal piece in dorsal (144) and lateral (145) view; 146) sternite VIII and IX; 147) entire penis, lateral view; 148) pedo-tectal part of penis, dorsal view. Abbreviations: ed, ejaculatory duct; pn, pennons on ventral fold of posttegminal membrane, rr, rectal ribbon-like internal lamina.
@ -338,7 +342,7 @@ from gate), -21.58383/165.79667,
<paragraph id="8BF636C9BF56FFE7FF5C580AFC65F242" blockId="25.[151,931,1048,1070]" box="[151,931,1048,1070]" pageId="25" pageNumber="540">
<emphasis id="B93DEADBBF56FFE7FF5C580AFEEAF241" bold="true" box="[151,300,1048,1070]" pageId="25" pageNumber="540">FIGURE 149.</emphasis>
Localities of
<taxonomicName id="4C494D4ABF56FFE7FE73580BFDBFF242" box="[440,633,1049,1070]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="25" pageNumber="540" phylum="Tracheophyta" rank="species" species="tibialis">
<taxonomicName id="4C494D4ABF56FFE7FE73580BFDBFF242" authorityName="Wanat" authorityYear="2013" box="[440,633,1049,1070]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B93DEADBBF56FFE7FE73580BFDBFF242" box="[440,633,1049,1070]" italics="true" pageId="25" pageNumber="540">Hellerenius tibialis</emphasis>
</taxonomicName>
in New Caledonia mainland.
@ -382,7 +386,7 @@ Occurs mainly in northern
(
<figureCitation id="13722A4CBF56FFE7FD0B591DFCE5F34B" box="[704,803,1295,1320]" captionStart="FIGURE 149" captionStartId="25.[151,250,1048,1070]" captionText="FIGURE 149. Localities of Hellerenius tibialis in New Caledonia mainland." pageId="25" pageNumber="540">Fig. 149</figureCitation>
), where it can be frequently collected together with
<taxonomicName id="4C494D4ABF56FFE7FA4A5902FF3AF327" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="25" pageNumber="540" phylum="Tracheophyta" rank="species" species="plumipes">
<taxonomicName id="4C494D4ABF56FFE7FA4A5902FF3AF327" authorityName="Wanat" authorityYear="2013" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="plumipes">
<emphasis id="B93DEADBBF56FFE7FA4A5902FA53F34B" box="[1409,1429,1296,1319]" italics="true" pageId="25" pageNumber="540">H</emphasis>
.
<emphasis id="B93DEADBBF56FFE7FF5C5921FF3AF327" box="[151,252,1331,1355]" italics="true" pageId="25" pageNumber="540">plumipes</emphasis>
@ -394,9 +398,11 @@ from the same plant. In Southern Province only in Col dAmieu, Plateau de Dogn
<paragraph id="8BF636C9BF56FFE7FF0C5969FCE3F06C" blockId="25.[151,1436,1115,2006]" pageId="25" pageNumber="540">
<emphasis id="B93DEADBBF56FFE7FF0C5969FE87F3F8" bold="true" box="[199,321,1403,1428]" pageId="25" pageNumber="540">Diagnosis.</emphasis>
The only species of
<taxonomicName id="4C494D4ABF56FFE7FD8B5969FD04F3FF" box="[576,706,1403,1427]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF56FFE7FD8B5969FD04F3FF" box="[576,706,1403,1427]" italics="true" pageId="25" pageNumber="540">Hellerenius</emphasis>
</taxonomicName>
with sinuate, twisted and compressed hind tibiae (weaker but expressed also in female), all mucronate in male. Most similar to
<taxonomicName id="4C494D4ABF56FFE7FC4F59B2FBCAF3DB" box="[900,1036,1439,1463]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="25" pageNumber="540" phylum="Tracheophyta" rank="species" species="plumipes">
<taxonomicName id="4C494D4ABF56FFE7FC4F59B2FBCAF3DB" authorityName="Wanat" authorityYear="2013" box="[900,1036,1439,1463]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="plumipes">
<emphasis id="B93DEADBBF56FFE7FC4F59B2FC5EF3DB" box="[900,920,1440,1463]" italics="true" pageId="25" pageNumber="540">H</emphasis>
.
<emphasis id="B93DEADBBF56FFE7FC6C598DFBCAF3DB" box="[935,1036,1439,1463]" italics="true" pageId="25" pageNumber="540">plumipes</emphasis>
@ -410,7 +416,7 @@ in body size and color, but easy to distinguish by totally bare coxae, trochante
Body length
<quantity id="4CB19B2CBF56FFE7FE335A1EFD43F04F" box="[504,645,1548,1572]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.7" metricValueMin="3.4" pageId="25" pageNumber="540" unit="mm" value="3.55" valueMax="3.7" valueMin="3.4">3.43.7 mm</quantity>
. Coloration and most external characters as in
<taxonomicName id="4C494D4ABF56FFE7FB635A1EFAF4F04F" box="[1192,1330,1547,1571]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="25" pageNumber="540" phylum="Tracheophyta" rank="species" species="plumipes" status="sp. nov.">
<taxonomicName id="4C494D4ABF56FFE7FB635A1EFAF4F04F" authorityName="Wanat" authorityYear="2013" box="[1192,1330,1547,1571]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="540" phylum="Arthropoda" rank="species" species="plumipes" status="sp. nov.">
<emphasis id="B93DEADBBF56FFE7FB635A1EFB7AF04F" box="[1192,1212,1548,1571]" italics="true" pageId="25" pageNumber="540">H</emphasis>
.
<emphasis id="B93DEADBBF56FFE7FB065A19FAF4F04F" box="[1229,1330,1547,1571]" italics="true" pageId="25" pageNumber="540">plumipes</emphasis>
@ -420,21 +426,21 @@ Body length
</paragraph>
<paragraph id="8BF636C9BF56FFE7FF0C5A46FBB9F092" blockId="25.[151,1436,1115,2006]" pageId="25" pageNumber="540">
Rostrum in both sexes slightly shorter than in previous species; underside of prorostrum polished, with subgenal sulci largely vanished (
<figureCitation id="13722A4CBF56FFE7FDC55A68FD8AF0FF" box="[526,588,1658,1683]" captionStart="FIGURES 1 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="25" pageNumber="540">Fig.7</figureCitation>
<figureCitation id="13722A4CBF56FFE7FDC55A68FD8AF0FF" box="[526,588,1658,1683]" captionStart="FIGURES 1 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." figureDoi="http://doi.org/10.5281/zenodo.249083" httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="25" pageNumber="540">Fig.7</figureCitation>
). Distal antennal segments more elongate, funicular 7th segment 1.31.4×, club 4.44.5× as long as wide (Figs 125, 126). Elytra less widened (Fig. 124). Legs shorter, ventrally not protrudingly pilose; hind tibiae sinuate and twisted (Figs 132134); protarsus as in Fig. 136; claws roundly expanded at base (Fig. 137). Wing: anal 1A retained as single rudiment; 2A without spur.
</paragraph>
<paragraph id="8BF636C9BF56FFE4FF0C5B18FD31F708" blockId="25.[151,1436,1115,2006]" lastBlockId="26.[151,1436,151,717]" lastPageId="26" lastPageNumber="541" pageId="25" pageNumber="540">
Male. Rostrum bare ventrally. All tibiae with minute, acute mucro (Fig. 135); mid-tibia irregularly curved; hind tibia strongly sinuate, its distal half compressed and slightly twisted, separated by oblique carina (Fig. 132). Last abdominal ventrite with large, rectangular prominence, lacking setose depression (Fig. 138). Pygidium with third part exposed and separated by complete sulcus; marginal rim present, polished. Sternite VIII entire evenly sclerotised. Sternite IX with untoothed apodeme 1.61.7× as long as large fork (Fig. 146). Tegminal plate peculiar, almost 3× as long as broad; membranous lobes about twice as long as parameroid ones, clearly separated; suprafenestral sclerites each with more than 50 long macrochaetae forming dense tuft completely covering membranous lobe; fenestrae large, elongate, well margined; postfenestral plate membranous; prostegium with incomplete median suture (Figs 144, 145).
<collectingCountry id="F35E7659BF55FFE4FD4F5CCDFD71F694" box="[644,695,223,248]" name="Iceland" pageId="26" pageNumber="541">Fold</collectingCountry>
of post tegminal membrane with long pennons (Fig. 147). Penis apodemes slightly shorter than pedon; the latter 5.86.0× as long as broad, tapering, with long, narrow tip, ventrally notched from base to nearly apical fourth (Fig. 148); tectum only slightly shorter than pedon, pointed apically, well sclerotised throughout; endophallus as in
<taxonomicName id="4C494D4ABF55FFE4FDA25D5FFD36F708" box="[617,752,332,356]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="26" pageNumber="541" phylum="Tracheophyta" rank="species" species="plumipes">
<taxonomicName id="4C494D4ABF55FFE4FDA25D5FFD36F708" authorityName="Wanat" authorityYear="2013" box="[617,752,332,356]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="species" species="plumipes">
<emphasis id="B93DEADBBF55FFE4FDA25D5FFD36F708" box="[617,752,332,356]" italics="true" pageId="26" pageNumber="541">H. plumipes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF636C9BF55FFE4FF0C5D7DFA94F450" blockId="26.[151,1436,151,717]" pageId="26" pageNumber="541">
Female. Mid-tibiae straight; hind tibiae slightly sinuate and compressed in distal part (Figs 133, 134), without separating carina; their outer edge sharp. Last abdominal ventrite unevenly convex, with elevation triangular in outline (Fig. 139). Tergite VII strongly transverse, almost twice as wide as long (Fig. 141). Pouch-like median process of membrane small, much longer than wide. Tergite VIII slightly longer than broad, with clear median line as in
<taxonomicName id="4C494D4ABF55FFE4FF1F5E13FE9BF474" box="[212,349,512,536]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="26" pageNumber="541" phylum="Tracheophyta" rank="species" species="plumipes">
<taxonomicName id="4C494D4ABF55FFE4FF1F5E13FE9BF474" authorityName="Wanat" authorityYear="2013" box="[212,349,512,536]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="541" phylum="Arthropoda" rank="species" species="plumipes">
<emphasis id="B93DEADBBF55FFE4FF1F5E13FF2EF474" box="[212,232,513,536]" italics="true" pageId="26" pageNumber="541">H</emphasis>
.
<emphasis id="B93DEADBBF55FFE4FF335E12FE9BF474" box="[248,349,512,536]" italics="true" pageId="26" pageNumber="541">plumipes</emphasis>
@ -447,10 +453,10 @@ Female. Mid-tibiae straight; hind tibiae slightly sinuate and compressed in dist
<paragraph id="8BF636C9BF55FFE4FF0C5E7EFBAAF4E8" blockId="26.[151,1436,151,717]" box="[199,1132,620,645]" pageId="26" pageNumber="541">
<emphasis id="B93DEADBBF55FFE4FF0C5E7EFEE1F4E9" bold="true" box="[199,295,620,645]" pageId="26" pageNumber="541">Biology.</emphasis>
Collected in rainforest from leaves and flowers of various
<taxonomicName id="4C494D4ABF55FFE4FC705E7EFBF0F4E8" box="[955,1078,620,644]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="26" pageNumber="541" phylum="Tracheophyta" rank="genus">
<taxonomicName id="4C494D4ABF55FFE4FC705E7EFBAAF4E8" box="[955,1132,620,644]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="26" pageNumber="541" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93DEADBBF55FFE4FC705E7EFBF0F4E8" box="[955,1078,620,644]" italics="true" pageId="26" pageNumber="541">Hedycarya</emphasis>
</taxonomicName>
spp.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3536542BF55FFE4FF0C5E9DFF29F4A1" pageId="26" pageNumber="541" type="etymology">

View file

@ -1,46 +1,47 @@
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<subSubSection id="C3536542BF5EFFEFFF5C5C85FEF3F6BF" pageId="17" pageNumber="532" type="nomenclature">
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<emphasis id="B93DEADBBF5EFFEFFF5C5C85FE38F6DE" bold="true" box="[151,510,151,178]" pageId="17" pageNumber="532">
<taxonomicName id="4C494D4ABF5EFFEFFF5C5C85FE5EF6DD" box="[151,408,151,177]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="17" pageNumber="532" phylum="Tracheophyta" rank="species" species="plumipes" status="sp. nov.">
<taxonomicName id="4C494D4ABF5EFFEFFF5C5C85FE5EF6DD" authorityName="Wanat" authorityYear="2013" box="[151,408,151,177]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="plumipes" status="sp. nov.">
<emphasis id="B93DEADBBF5EFFEFFF5C5C85FE5EF6DD" bold="true" box="[151,408,151,177]" italics="true" pageId="17" pageNumber="532">Hellerenius plumipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57A0BF5EFFEFFE6B5C8AFE38F6DE" box="[416,510,152,178]" pageId="17" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
@ -49,7 +50,7 @@
</paragraph>
<paragraph id="8BF636C9BF5EFFEFFF5C5CA8FEF3F6BF" blockId="17.[151,510,151,211]" box="[151,309,186,211]" pageId="17" pageNumber="532">
(
<figureCitation id="13722A4CBF5EFFEFFF545CA8FEEAF6BF" box="[159,300,186,211]" captionStart-0="FIGURES 87 95" captionStart-1="FIGURES 111 121" captionStart-2="FIGURE 122" captionStartId-0="18.[151,264,1736,1759]" captionStartId-1="20.[151,264,1886,1909]" captionStartId-2="21.[151,250,1991,2013]" captionTargetBox-0="[151,1436,205,1706]" captionTargetBox-1="[182,1411,193,1851]" captionTargetId-0="figure@18.[151,1436,193,1715]" captionTargetId-1="figure@20.[175,1411,193,1865]" captionTargetPageId-0="18" captionTargetPageId-1="20" captionText-0="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." captionText-1="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." captionText-2="FIGURE 122. Localities of Hellerenius plumipes in New Caledonia mainland." httpUri-0="https://zenodo.org/record/249088/files/figure.png" httpUri-1="https://zenodo.org/record/249089/files/figure.png" pageId="17" pageNumber="532">Figs 92122</figureCitation>
<figureCitation id="13722A4CBF5EFFEFFF545CA8FEEAF6BF" box="[159,300,186,211]" captionStart-0="FIGURES 87 95" captionStart-1="FIGURES 111 121" captionStart-2="FIGURE 122" captionStartId-0="18.[151,264,1736,1759]" captionStartId-1="20.[151,264,1886,1909]" captionStartId-2="21.[151,250,1991,2013]" captionTargetBox-0="[151,1436,205,1706]" captionTargetBox-1="[182,1411,193,1851]" captionTargetId-0="figure@18.[151,1436,193,1715]" captionTargetId-1="figure@20.[175,1411,193,1865]" captionTargetPageId-0="18" captionTargetPageId-1="20" captionText-0="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." captionText-1="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." captionText-2="FIGURE 122. Localities of Hellerenius plumipes in New Caledonia mainland." figureDoi-0="http://doi.org/10.5281/zenodo.249088" figureDoi-1="http://doi.org/10.5281/zenodo.249089" httpUri-0="https://zenodo.org/record/249088/files/figure.png" httpUri-1="https://zenodo.org/record/249089/files/figure.png" pageId="17" pageNumber="532">Figs 92122</figureCitation>
)
</paragraph>
</subSubSection>
@ -105,10 +106,11 @@ II 1963,
,
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, from
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<emphasis id="B93DEADBBF5EFFEFFD005DC9FC80F79F" box="[715,838,475,499]" italics="true" pageId="17" pageNumber="532">Hedycarya</emphasis>
sp.
</taxonomicName>
sp.;
;
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,
<date id="FFF71009BF5EFFEFFC115DC9FBA4F79F" box="[986,1122,475,499]" pageId="17" pageNumber="532" value="2008-11-22">22 XI 2008</date>
@ -153,10 +155,11 @@ sp.;
,
<date id="FFF71009BF5EFFEFFBA15EA0FB10F4A7" box="[1130,1238,690,715]" pageId="17" pageNumber="532" value="2004-02-05">5 II 2004</date>
, from
<taxonomicName id="4C494D4ABF5EFFEFFAEA5EA1FA5AF4A7" box="[1313,1436,691,715]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="17" pageNumber="532" phylum="Tracheophyta" rank="genus">
<taxonomicName id="4C494D4ABF5EFFEFFAEA5EA1FF7FF482" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="17" pageNumber="532" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93DEADBBF5EFFEFFAEA5EA1FA5AF4A7" box="[1313,1436,691,715]" italics="true" pageId="17" pageNumber="532">Hedycarya</emphasis>
sp.
</taxonomicName>
sp.,
,
<specimenCount id="9D4FFD40BF5EFFEFFF0F5EC5FF28F482" box="[196,238,727,751]" pageId="17" pageNumber="532" type="generic">3 ♂</specimenCount>
2 ♀, leg. MW (MW). Upper Tchamba valley: Wâo Uni, refuge, -21.0057/165.2487,
<quantity id="4CB19B2CBF5EFFEFFB545EC5FB2EF482" box="[1183,1256,727,751]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="532" unit="m" value="400.0">400 m</quantity>
@ -186,10 +189,11 @@ Province Sud: same locality as the
,
<specimenCount id="9D4FFD40BF5EFFEFFC505F51FC12F536" box="[923,980,835,859]" pageId="17" pageNumber="532" type="generic">33 ♂</specimenCount>
53 ♀ (a number of exs from
<taxonomicName id="4C494D4ABF5EFFEFFAEA5F51FA5AF537" box="[1313,1436,835,859]" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="17" pageNumber="532" phylum="Tracheophyta" rank="genus">
<taxonomicName id="4C494D4ABF5EFFEFFAEA5F51FF7FF513" class="Magnoliopsida" family="Monimiaceae" genus="Hedycarya" kingdom="Plantae" order="Laurales" pageId="17" pageNumber="532" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93DEADBBF5EFFEFFAEA5F51FA5AF537" box="[1313,1436,835,859]" italics="true" pageId="17" pageNumber="532">Hedycarya</emphasis>
sp.
</taxonomicName>
sp.);
);
<date id="FFF71009BF5EFFEFFF1B5F75FE8FF513" box="[208,329,871,895]" pageId="17" pageNumber="532" value="2004-02-11">11 II 2004</date>
, at light,
<specimenCount id="9D4FFD40BF5EFFEFFE7F5F75FE24F512" box="[436,482,871,895]" pageId="17" pageNumber="532" type="generic">1 ♂</specimenCount>
@ -304,10 +308,11 @@ X 1978,
1 ♀ (MNHN). Pic Ningua Reserve (Camps des sapins), [-21.74/166.15],
<date id="FFF71009BF5EFFEFFC54594DFBCBF31B" box="[927,1037,1375,1399]" pageId="17" pageNumber="532" value="2008-10-03">3 X 2008</date>
, 1 ♀, beating
<taxonomicName id="4C494D4ABF5EFFEFFB66594CFAECF31A" box="[1197,1322,1374,1398]" class="Magnoliopsida" family="Myrtaceae" genus="Carpolepis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="17" pageNumber="532" phylum="Magnoliophyta" rank="genus">
<taxonomicName id="4C494D4ABF5EFFEFFB66594CFA97F31A" box="[1197,1361,1374,1398]" class="Magnoliopsida" family="Myrtaceae" genus="Carpolepis" kingdom="Plantae" order="Myrtales" pageId="17" pageNumber="532" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93DEADBBF5EFFEFFB66594CFAECF31A" box="[1197,1322,1374,1398]" italics="true" pageId="17" pageNumber="532">Carpolepis</emphasis>
sp.
</taxonomicName>
sp. (“voir fiche COL/ins/78-08”), leg. C. Mille (IACP). Mt Humboldt (S trail), Botanical Reserve, moss forest,
(“voir fiche COL/ins/78-08”), leg. C. Mille (IACP). Mt Humboldt (S trail), Botanical Reserve, moss forest,
<quantity id="4CB19B2CBF5EFFEFFB375991FA51F3F6" box="[1276,1431,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.205" metricValueMax="1.28" metricValueMin="1.13" pageId="17" pageNumber="532" unit="m" value="1205.0" valueMax="1280.0" valueMin="1130.0">11301280 m</quantity>
,
<date id="FFF71009BF5EFFEFFF5C59B5FEDFF3D3" box="[151,281,1447,1471]" pageId="17" pageNumber="532" value="2008-11-10">10 XI 2008</date>
@ -347,17 +352,19 @@ Distributed throughout the entire mainland of
<paragraph id="8BF636C9BF5EFFEFFF0C5A6CFC6CF127" blockId="17.[151,1437,258,2011]" pageId="17" pageNumber="532">
<emphasis id="B93DEADBBF5EFFEFFF0C5A6CFE87F0FB" bold="true" box="[199,321,1662,1687]" pageId="17" pageNumber="532">Diagnosis.</emphasis>
The most frequently collected species of
<taxonomicName id="4C494D4ABF5EFFEFFCEC5A6CFC6FF0FA" box="[807,937,1662,1686]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEADBBF5EFFEFFCEC5A6CFC6FF0FA" box="[807,937,1662,1686]" italics="true" pageId="17" pageNumber="532">Hellerenius</emphasis>
</taxonomicName>
in
<collectingCountry id="F35E7659BF5EFFEFFC185A6DFB40F0FB" box="[979,1158,1662,1687]" name="New Caledonia" pageId="17" pageNumber="532">New Caledonia</collectingCountry>
, unique in this genus in having protrudingly pilose coxae, trochanters and femora (more distinctly in male). It differs from
<taxonomicName id="4C494D4ABF5EFFEFFAC65AB1FABBF0D7" box="[1293,1405,1699,1723]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="17" pageNumber="532" phylum="Tracheophyta" rank="species" species="lobifer">
<taxonomicName id="4C494D4ABF5EFFEFFAC65AB1FABBF0D7" authorityName="Wanat" authorityYear="2001" box="[1293,1405,1699,1723]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF5EFFEFFAC65AB1FAE7F0D6" box="[1293,1313,1699,1722]" italics="true" pageId="17" pageNumber="532">H</emphasis>
.
<emphasis id="B93DEADBBF5EFFEFFAF95AB1FABBF0D7" box="[1330,1405,1699,1723]" italics="true" pageId="17" pageNumber="532">lobifer</emphasis>
</taxonomicName>
in smaller body size, elytra with more or less distinct bronze tinge, and in the structure of the last abdominal ventrite in both sexes. It resembles
<taxonomicName id="4C494D4ABF5EFFEFFE175AF9FD97F16F" box="[476,593,1771,1795]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="17" pageNumber="532" phylum="Tracheophyta" rank="species" species="tibialis" status="sp. nov.">
<taxonomicName id="4C494D4ABF5EFFEFFE175AF9FD97F16F" authorityName="Wanat" authorityYear="2013" box="[476,593,1771,1795]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="532" phylum="Arthropoda" rank="species" species="tibialis" status="n. sp.">
<emphasis id="B93DEADBBF5EFFEFFE175AF9FD97F16F" box="[476,593,1771,1795]" italics="true" pageId="17" pageNumber="532">H. tibialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20E57A0BF5EFFEFFD975AF8FD59F16F" box="[604,671,1770,1795]" pageId="17" pageNumber="532" rank="species">
@ -375,31 +382,31 @@ long, black with evident bronze sheen; elytra matt; base of antennal scape testa
</paragraph>
<paragraph id="8BF636C9BF5EFFEDFF0C5B8DFC28F19C" blockId="17.[151,1437,258,2011]" lastBlockId="19.[151,1436,1887,2032]" lastPageId="19" lastPageNumber="534" pageId="17" pageNumber="532">
Rostrum shiny; prorostrum slightly thinner than metarostrum, on underside with fine, shagreened sulci throughout entire length (
<figureCitation id="13722A4CBF5EFFEFFE7E5BD0FE3FF1B7" box="[437,505,1986,2011]" captionStart="FIGURES 1 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="17" pageNumber="532">Fig. 8</figureCitation>
<figureCitation id="13722A4CBF5EFFEFFE7E5BD0FE3FF1B7" box="[437,505,1986,2011]" captionStart="FIGURES 1 8" captionStartId="5.[151,264,1860,1883]" captionTargetBox="[191,1381,195,1805]" captionTargetId="figure@5.[191,1382,195,1805]" captionTargetPageId="5" captionText="FIGURES 1 8. Mouthparts and rostrum underside, SEM photographs: 1) Hellerenius monteithi, male mouthparts; 2) H. plumipes, male mouthparts (arrow points on U-shaped stripes on outer side of terminal maxillary palp segment); 3) Archellerenius giustoi, female mouthparts; 4) H. plumipes, sensilla on apex of female maxillary palp, magnification 9600 ×; 5) H. monteithi, female prorostrum, ventral view; 6) H. plumipes, male rostrum, ventral view; 7) H. tibialis, female prorostrum, ventral view; 8) H. plumipes, female prorostrum, ventral view. Abbreviations: ls, mixed setae of labial ligula and maxillary malae; m, mandible; mp, maxillary palp (2 - segmented); pf, palpifer; po, postmentum; pr, prementum; sg, subgenal sulcus." figureDoi="http://doi.org/10.5281/zenodo.249083" httpUri="https://zenodo.org/record/249083/files/figure.png" pageId="17" pageNumber="532">Fig. 8</figureCitation>
). Antenna as in Fig. 99; length/width ratios: scape 10.011.2, funicular segments:
<emphasis id="B93DEADBBF5CFFEDFF5C5B4DFEA6F118" bold="true" box="[151,352,1887,1910]" pageId="19" pageNumber="534">FIGURES 96110.</emphasis>
<taxonomicName id="4C494D4ABF5CFFEDFEA25B72FD84F119" box="[361,578,1888,1909]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="19" pageNumber="534" phylum="Tracheophyta" rank="species" species="plumipes">
<taxonomicName id="4C494D4ABF5CFFEDFEA25B72FD84F119" authorityName="Wanat" authorityYear="2013" box="[361,578,1888,1909]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="534" phylum="Arthropoda" rank="species" species="plumipes">
<emphasis id="B93DEADBBF5CFFEDFEA25B72FD84F119" box="[361,578,1888,1909]" italics="true" pageId="19" pageNumber="534">Hellerenius plumipes</emphasis>
</taxonomicName>
, male (97102, 104106, 109, 110), female (96, 103, 107, 108): 58) elytra, lateral view; 96) elytra, dorsal view; 97) pronotum, dorsal view; 98) head with rostrum, dorsal view; 99) antenna; 100) head and rostrum base, ventral view; 101) rostrum (part), ventral view; 102) rostrum, lateral view; 103) head with rostrum, lateral view; 104) fore femur (part) and tibia; 105) protarsus; 106108) metatibia in lateral (106, 107) and dorsal (108) view; 109) all abdominal ventrites, ventral view; 110) last abdominal ventrite, postero-lateral view.
</paragraph>
<caption id="DF366641BF5DFFECFF5C5ADAFF28F114" httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="18" pageNumber="533" targetBox="[151,1436,205,1706]" targetPageId="18">
<caption id="DF366641BF5DFFECFF5C5ADAFF28F114" ID-DOI="http://doi.org/10.5281/zenodo.249088" httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="18" pageNumber="533" targetBox="[151,1436,205,1706]" targetPageId="18">
<paragraph id="8BF636C9BF5DFFECFF5C5ADAFF28F114" blockId="18.[151,1436,1736,1912]" pageId="18" pageNumber="533">
<emphasis id="B93DEADBBF5DFFECFF5C5ADAFE92F0B1" bold="true" box="[151,340,1736,1759]" pageId="18" pageNumber="533">FIGURES 8795.</emphasis>
<taxonomicName id="4C494D4ABF5DFFECFE925ADBFDF2F0B2" box="[345,564,1737,1758]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="18" pageNumber="533" phylum="Tracheophyta" rank="species" species="monteithi">
<taxonomicName id="4C494D4ABF5DFFECFE925ADBFDF2F0B2" authorityName="Wanat" authorityYear="2013" box="[345,564,1737,1758]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B93DEADBBF5DFFECFE925ADBFDF2F0B2" box="[345,564,1737,1758]" italics="true" pageId="18" pageNumber="533">Hellerenius monteithi</emphasis>
</taxonomicName>
, male (8791),
<taxonomicName id="4C494D4ABF5DFFECFD1E5ADBFC6AF0B2" box="[725,940,1737,1758]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="18" pageNumber="533" phylum="Tracheophyta" rank="species" species="plumipes">
<taxonomicName id="4C494D4ABF5DFFECFD1E5ADBFC6AF0B2" authorityName="Wanat" authorityYear="2013" box="[725,940,1737,1758]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="533" phylum="Arthropoda" rank="species" species="plumipes">
<emphasis id="B93DEADBBF5DFFECFD1E5ADBFC6AF0B2" box="[725,940,1737,1758]" italics="true" pageId="18" pageNumber="533">Hellerenius plumipes</emphasis>
</taxonomicName>
, male (95) and female (9294): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88A) and endophallus (88B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window.
</paragraph>
</caption>
<caption id="DF366641BF5BFFEAFF5C5B4CFC99F183" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="20" pageNumber="535" targetBox="[182,1411,193,1851]" targetPageId="20">
<caption id="DF366641BF5BFFEAFF5C5B4CFC99F183" ID-DOI="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="20" pageNumber="535" targetBox="[182,1411,193,1851]" targetPageId="20">
<paragraph id="8BF636C9BF5BFFEAFF5C5B4CFC99F183" blockId="20.[151,1436,1886,2031]" pageId="20" pageNumber="535">
<emphasis id="B93DEADBBF5BFFEAFF5C5B4CFEA8F118" bold="true" box="[151,366,1886,1909]" pageId="20" pageNumber="535">FIGURES 111121.</emphasis>
<taxonomicName id="4C494D4ABF5BFFEAFEBC5B4DFD94F118" box="[375,594,1887,1908]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="20" pageNumber="535" phylum="Tracheophyta" rank="species" species="plumipes">
<taxonomicName id="4C494D4ABF5BFFEAFEBC5B4DFD94F118" authorityName="Wanat" authorityYear="2013" box="[375,594,1887,1908]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="535" phylum="Arthropoda" rank="species" species="plumipes">
<emphasis id="B93DEADBBF5BFFEAFEBC5B4DFD94F118" box="[375,594,1887,1908]" italics="true" pageId="20" pageNumber="535">Hellerenius plumipes</emphasis>
</taxonomicName>
, male (117120), female (111116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX.
@ -407,35 +414,35 @@ Rostrum shiny; prorostrum slightly thinner than metarostrum, on underside with f
</caption>
<paragraph id="8BF636C9BF5AFFEBFF5C5C89FEEFF42E" blockId="21.[151,1436,151,1082]" pageId="21" pageNumber="536">
1st 2.02.6, 2nd 3.13.8, 7th 1.01.1, club 3.23.4; scape as long as entire funicle. Head: frons with shallow median depression; vertex weakly convex and finely punctured in front; temple 0.50.7 as long as eye; underside weakly convex, behind eyes with indistinct concentric wrinkles, rarely with median tubercle (Fig. 100). Pronotum: front margin not scratched; disc microreticulate and punctate, punctures less than an ommatidium diameter, subapical constriction scarcely visible on dorsum, subbasal line never traced; sternellum weakly prominent; posterior margin of hypomeron with pointed tubercle. Scutellum often slightly elongate. Elytra highly convex, compared to
<taxonomicName id="4C494D4ABF5AFFEBFA4A5D42FF24F7E7" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="21" pageNumber="536" phylum="Tracheophyta" rank="species" species="lobifer">
<taxonomicName id="4C494D4ABF5AFFEBFA4A5D42FF24F7E7" authorityName="Wanat" authorityYear="2001" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF5AFFEBFA4A5D42FA53F70B" box="[1409,1429,336,359]" italics="true" pageId="21" pageNumber="536">H</emphasis>
.
<emphasis id="B93DEADBBF5AFFEBFF5C5D61FF24F7E7" box="[151,226,371,395]" italics="true" pageId="21" pageNumber="536">lobifer</emphasis>
</taxonomicName>
with less prominent caudal part and declivities gently arched in profile (
<figureCitation id="13722A4CBF5AFFEBFBD95D60FBADF7E7" box="[1042,1131,370,395]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="21" pageNumber="536">Figs 94</figureCitation>
<figureCitation id="13722A4CBF5AFFEBFBD95D60FBADF7E7" box="[1042,1131,370,395]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249088" httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="21" pageNumber="536">Figs 94</figureCitation>
, 96); intervals all flat, with dense scale-like microsculpture; sensory setae 23, ordered. Metaventrite about 1.5× as long as diameter of mesocoxa. Wing (
<figureCitation id="13722A4CBF5AFFEBFEAD5DA8FE7BF7BF" box="[358,445,442,467]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="21" pageNumber="536">Fig. 93</figureCitation>
<figureCitation id="13722A4CBF5AFFEBFEAD5DA8FE7BF7BF" box="[358,445,442,467]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249088" httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="21" pageNumber="536">Fig. 93</figureCitation>
): radial window almost completely reduced; anal 1A as single rudiment (occasionally much shorter second vein retained); 2A sometimes with minute spur; 3A relatively long. Legs moderately long and robust; tibiae all straight, with blunt outer edges (Figs 104, 106108); protarsus as in Fig. 105; 2nd tarsomere isodiametric.
</paragraph>
<paragraph id="8BF636C9BF5AFFEBFF0C5E5FFB1DF256" blockId="21.[151,1436,151,1082]" pageId="21" pageNumber="536">
Male. Underside of rostrum with short, white, erect setae (Fig. 102); similar but longer setae form distinct fringe on undersides of coxae, trochanters and basal halves of femora on fore and mid-legs (
<figureCitation id="13722A4CBF5AFFEBFB2F5E63FAFAF4E6" box="[1252,1340,625,650]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="21" pageNumber="536">Fig. 95</figureCitation>
<figureCitation id="13722A4CBF5AFFEBFB2F5E63FAFAF4E6" box="[1252,1340,625,650]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249088" httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="21" pageNumber="536">Fig. 95</figureCitation>
). Tibiae straight, unarmed, metatibia with long apical fringe (Fig. 106). Fifth abdominal ventrite V ca. 1.81.9× broader than long, strongly convex, at apical declivity with round, setose impression (Figs 109, 110). Pygidium: exposed part distinctly flattened, 0.200.25 total pygidium length, with no trace of marginal rim; transverse sulcus incomplete, in middle as semicircular depression. Sternite VIII with short lobes. Sternite IX forked, apodeme untoothed (
<figureCitation id="13722A4CBF5AFFEBFEDC5F37FEBFF552" box="[279,377,805,830]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="21" pageNumber="536">Fig. 121</figureCitation>
<figureCitation id="13722A4CBF5AFFEBFEDC5F37FEBFF552" box="[279,377,805,830]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="21" pageNumber="536">Fig. 121</figureCitation>
). Tegmen connate; basal piece largely sclerotized between thin arms; tegminal plate 1.31.4× longer than broad, widened apicad; membranous lobes and fenestrae absent; suprafenestral sclerites expanded ventrad, with one row of short macrochaetae along inner margins, outer margins transparent and microsetose; postfenestral plate membranous; prostegium divided into pair of widely separated tails (
<figureCitation id="13722A4CBF5AFFEBFBB25F83FB1DF5C6" box="[1145,1243,913,938]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="21" pageNumber="536">Fig. 117</figureCitation>
<figureCitation id="13722A4CBF5AFFEBFBB25F83FB1DF5C6" box="[1145,1243,913,938]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="21" pageNumber="536">Fig. 117</figureCitation>
). Penis short and broad, flat (
<figureCitation id="13722A4CBF5AFFEBFED15FA7FE7FF5A2" box="[282,441,949,974]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="21" pageNumber="536">Figs 118, 119</figureCitation>
<figureCitation id="13722A4CBF5AFFEBFED15FA7FE7FF5A2" box="[282,441,949,974]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="21" pageNumber="536">Figs 118, 119</figureCitation>
); pedon 2.42.7 × as long as broad, much shorter than its apodemes, rhombic, distinctly narrowing basad and apicad from basal 0.4 length, ventrally membranous except margins and triangular apex; base of tectum expanded, with longitudinal folds, apex of tectum thin and long; endophallus with microspinose area close to apodemal base and transverse microplates in orificial region, terminal part as in
<figureCitation id="13722A4CBF5AFFEBFBBD5833FB11F256" box="[1142,1239,1057,1082]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="21" pageNumber="536">Fig. 120</figureCitation>
<figureCitation id="13722A4CBF5AFFEBFBBD5833FB11F256" box="[1142,1239,1057,1082]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="21" pageNumber="536">Fig. 120</figureCitation>
.
</paragraph>
<caption id="DF366641BF5AFFEBFF5C5BD5FC71F1B1" box="[151,951,1991,2013]" pageId="21" pageNumber="536">
<paragraph id="8BF636C9BF5AFFEBFF5C5BD5FC71F1B1" blockId="21.[151,951,1991,2013]" box="[151,951,1991,2013]" pageId="21" pageNumber="536">
<emphasis id="B93DEADBBF5AFFEBFF5C5BD5FEEAF1B0" bold="true" box="[151,300,1991,2013]" pageId="21" pageNumber="536">FIGURE 122.</emphasis>
Localities of
<taxonomicName id="4C494D4ABF5AFFEBFE735BD5FD49F1B0" box="[440,655,1991,2012]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="21" pageNumber="536" phylum="Tracheophyta" rank="species" species="plumipes">
<taxonomicName id="4C494D4ABF5AFFEBFE735BD5FD49F1B0" authorityName="Wanat" authorityYear="2013" box="[440,655,1991,2012]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="536" phylum="Arthropoda" rank="species" species="plumipes">
<emphasis id="B93DEADBBF5AFFEBFE735BD5FD49F1B0" box="[440,655,1991,2012]" italics="true" pageId="21" pageNumber="536">Hellerenius plumipes</emphasis>
</taxonomicName>
in New Caledonia mainland.
@ -443,21 +450,21 @@ in New Caledonia mainland.
</caption>
<paragraph id="8BF636C9BF59FFE8FF0C5C85FC7DF7C0" blockId="22.[151,1436,151,608]" pageId="22" pageNumber="537">
Female. Rostrum as in Fig. 103, asetose ventrally; prorostrum slightly wider at apex than in middle. Apices of coxae, often also underside of trochanters with fine, protruding setae much shorter than in male; underside of femora bare. Fifth abdominal ventrite twice as wide as long, weakly and regularly convex, finely rimmed around (
<figureCitation id="13722A4CBF59FFE8FF545D16FEFBF770" box="[159,317,260,285]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="22" pageNumber="537">Figs 111113</figureCitation>
<figureCitation id="13722A4CBF59FFE8FF545D16FEFBF770" box="[159,317,260,285]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="22" pageNumber="537">Figs 111113</figureCitation>
). Tergite VII semicircular, at most 1.4× wider than long (
<figureCitation id="13722A4CBF59FFE8FC1A5D16FBF2F771" box="[977,1076,260,285]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="22" pageNumber="537">Fig. 116</figureCitation>
<figureCitation id="13722A4CBF59FFE8FC1A5D16FBF2F771" box="[977,1076,260,285]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="22" pageNumber="537">Fig. 116</figureCitation>
). Pouch-like median process of membrane barely longer than wide, rounded apically, smaller than in
<taxonomicName id="4C494D4ABF59FFE8FC0C5D3BFBF1F72C" box="[967,1079,296,320]" class="Magnoliopsida" family="Monimiaceae" genus="Hellerenius" kingdom="Plantae" order="Laurales" pageId="22" pageNumber="537" phylum="Tracheophyta" rank="species" species="lobifer">
<taxonomicName id="4C494D4ABF59FFE8FC0C5D3BFBF1F72C" authorityName="Wanat" authorityYear="2001" box="[967,1079,296,320]" class="Insecta" family="Brentidae" genus="Hellerenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="537" phylum="Arthropoda" rank="species" species="lobifer">
<emphasis id="B93DEADBBF59FFE8FC0C5D3BFC1DF72C" box="[967,987,297,320]" italics="true" pageId="22" pageNumber="537">H</emphasis>
.
<emphasis id="B93DEADBBF59FFE8FC255D3AFBF1F72C" box="[1006,1079,296,320]" italics="true" pageId="22" pageNumber="537">lobifer</emphasis>
</taxonomicName>
. Tergite VIII wider than long, round, weakly and evenly sclerotised except for cleared median line, with sparse, short setae (
<figureCitation id="13722A4CBF59FFE8FB065D5EFAF6F709" box="[1229,1328,332,357]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="22" pageNumber="537">Fig. 115</figureCitation>
<figureCitation id="13722A4CBF59FFE8FB065D5EFAF6F709" box="[1229,1328,332,357]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="22" pageNumber="537">Fig. 115</figureCitation>
). Sternite VIII (spiculum ventrale) 2.0× as long as gonocoxite, with large apical spatula (
<figureCitation id="13722A4CBF59FFE8FBE85D7DFB40F7E4" box="[1059,1158,367,392]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="22" pageNumber="537">Fig. 114</figureCitation>
<figureCitation id="13722A4CBF59FFE8FBE85D7DFB40F7E4" box="[1059,1158,367,392]" captionStart="FIGURES 111 121" captionStartId="20.[151,264,1886,1909]" captionTargetBox="[182,1411,193,1851]" captionTargetId="figure@20.[175,1411,193,1865]" captionTargetPageId="20" captionText="FIGURES 111 121. Hellerenius plumipes, male (117 120), female (111 116): 111, 112) all abdominal ventrites in dorsal (111) and lateral (112) view; 113) fifth abdominal ventrite, posterior view; 114) entire ovipositor with spiculum ventrale, lateral view; 115) tergite VIII and ovipositor, enlarged stylus; 116) tergite VII; 117) tegminal plate and basal piece, dorsal view; 118, 119) penis with partial apodemes and endophallus in dorsal (118) and lateral (119) view; 120) terminal parts of endophallus, tegminal and penial apodemes, lateral view; 121) sternite VIII and IX." figureDoi="http://doi.org/10.5281/zenodo.249089" httpUri="https://zenodo.org/record/249089/files/figure.png" pageId="22" pageNumber="537">Fig. 114</figureCitation>
). Styli ca. 2× as long as wide. Spermatheca having straight and relatively thick corpus (
<figureCitation id="13722A4CBF59FFE8FC915D86FC68F7C0" box="[858,942,404,429]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="22" pageNumber="537">Fig. 92</figureCitation>
<figureCitation id="13722A4CBF59FFE8FC915D86FC68F7C0" box="[858,942,404,429]" captionStart="FIGURES 87 95" captionStartId="18.[151,264,1736,1759]" captionTargetBox="[151,1436,205,1706]" captionTargetId="figure@18.[151,1436,193,1715]" captionTargetPageId="18" captionText="FIGURES 87 95. Hellerenius monteithi, male (87 91), Hellerenius plumipes, male (95) and female (92 94): 87) penis, lateral view; 88) entire penis, enlarged pedo-tectal part in dorsal view (88 A) and endophallus (88 B) in lateral view; 89) tegminal plate and basal piece, lateral view; 90) entire tegmen, enlarged tegminal plate (90 A), dorsal view; 91) sternites VIII and IX, enlarged fork of s. IX (91 A); 92) spermatheca; 93) metathoracic wing, enlarged radial cell to show vestigial radial window (93 A); 94) elytra, lateral view; 95) pilose mesocoxae, trochanters and femoral bases. Abbreviations: ed, ejaculatory duct; w, radial window." figureDoi="http://doi.org/10.5281/zenodo.249088" httpUri="https://zenodo.org/record/249088/files/figure.png" pageId="22" pageNumber="537">Fig. 92</figureCitation>
).
</paragraph>
<paragraph id="8BF636C9BF59FFE8FF0C5DA5FDF2F7BC" blockId="22.[151,1436,151,608]" box="[199,564,439,464]" pageId="22" pageNumber="537">Morphological indices in Tab. 1.</paragraph>