The Apionidae (Coleoptera: Curculionoidea) of New Caledonia. Genera Archellerenius gen. nov. and Hellerenius Wanat Author Wanat, Marek text Zootaxa 2013 3717 4 515 542 journal article 43001 10.11646/zootaxa.3717.4.5 16d0cbc8-dd2c-4a41-8ab3-4649778bcee0 1175-5326 249082 7AC6654C-0D33-4921-B89D-807ADE0FF438 Archellerenius giustoi sp. nov. ( Figs 3 , 9–37 ) Type material. Holotype ♂: a) New Caledonia , Col des Roussettes, 500 m , 5 Feb 1963 , G. Kuschel (NZAC). Paratypes ( 5 ♂ 3 ♀): Province Nord: Mt. Aoupinié, W of Ponerihouen [near sawmill, -21.1555/165.3201], 550 m , 30–31 VII 1971 , beating, 1 ♂ , leg. J. Gressitt (BPBM). Mt Panié (refuge hut), 20.34.43S/164.46.04E [-20.5786/ 164.7678], 1330 m , humid montane forest, 2 II 2004 , 1 ♀; 1200–1300 m , 3 II 2004 , 1 ♂ 1 ♀ - all leg. MW (MW). Upper Tchamba Valley: Wâo Uni (refuge), -21.0057/165.2487, 400 m , 15 I 2007 , night beating, 1 ♂ , leg. MW (MW). Col des Roussettes (rainforest near refuge), -21.4074/165.5250, 530 m , 2 XII 2010 , 1 ♀, leg. MW (MW). Province Sud: Pic du Pin base (forest & plantation), -22.2482/166.829, 280 m, 26 XII 2006 , 1 ♂ , leg. MW & RD (MW). Mt Humboldt (S trail), Botanical Reserve, -21.8827/166.3997, 1130–1280 m , moss forest, 10 XI 2008 , 1 ♂ , leg. MW (MW). Total distribution in Fig. 46 . Diagnosis. See generic diagnosis above. Description. Body 2.8–3.2 mm long; antennal scape entirely testaceous. Rostrum shaped as in Figs 13–16 , at base finely punctured, rugose at sides; prorostrum shiny, with microscopic punctures at least basally. Antennal club as long as 6 distal funicular segments combined. Head: frons usually with shallow V-like fovea; vertex evenly flattened throughout; subocular tooth usually weakly expressed in head profile. Antennae ( Fig. 19 ): length/width ratios: scape 6.5–8.0, funicular segments: 1st 1.3–1.5, 2nd ca. 2.5, 6th–7th 1.0, scape as long or longer than 6 basal funicular segments combined. Pronotum weakly rounded at sides ( Fig.12 ); hypomeron acutely prominent at posterior margin. Elytra moderately convex ( Figs 9–11 ); striae superficial; intervals flat, finely microreticulate and shiny. Fore leg as in Figs 18, 20 . Abdomen (ventrites 1–5) 1.2× longer than broad. Male. Antennal club about 2.5× as long as wide. Abdominal ventrites narrower ( Fig. 25 ); 5th ventrite twice as wide as long, broadly truncate, moderately and evenly convex, middle of apical margin with sparse fringe of white setae ( Fig. 26 ). Pygidium: apical third exposed, strongly declivitous, with distinct marginal rim; separating sulcus obsolescent ( Figs 23, 24 ). Sternite VIII with distinct carinae. Sternite IX with untoothed apodeme, ca. 1.6× as long as basal fork ( Fig. 35 ). Tegminal apodeme strongly expanded in apical third; tegminal plate 1.3–1.4× as long as broad, with lateral arms much thinner than adjoining arms of basal piece ( Figs 36, 37 ); membranous lobes short, rounded, apically extended into narrow, acute processes, these microsetose apically; suprafenestral sclerites half as long as fenestrae, acutely produced along internal margins; sensillae indistinct; macrochaetae 6–8, short, hardly exceeding acute sclerotized projections of parameroid lobes; membrane beneath fenestrae sclerotized equally to prostegium; postfenestral sclerite bar-like; prostegium long, deeply incised laterally, divided on its 0.6–0.7 length. Penis: apodemes as long as pedon, strongly expanded apicad; tectum reaching apical fourth of pedon; pedon with rounded, strongly upcurved apex ( Figs 33, 34 ), ventrally membranous in basal half; endophallus with enclosed part densely microspinose in basal third, with sharp and straight distal border-line separating it from big, simply membranous terminal process, the latter with paired basal lobes covering entrance of ejaculatory duct, microplates in sub-orificial region extremely fine and indistinct. Female. Antennal club more obtuse, 2.0–2.2× as long as wide ( Fig. 19 ). Abdominal ventrites shaped as in Figs 27, 28 . Tergite VII short, broadly rounded ( Fig. 30 ). Tergite VIII transverse, largely membranous, with narrow but complete marginal sclerotisation. Sternite VIII (spiculum ventrale) relatively short, slightly widened in middle part, apically with triangular expansion ( Fig. 32 ). Styli stout and short, hemispherical, with several apical setae; coxites subtruncate apically. Foretube darkened, with dense long microtrichia. Bursa copulatrix relatively large, with corrugated longitudinal folds. Spermatheca: corpus with hook-like process receiving relatively thick spermathecal duct ( Fig. 31 ). Morphological indices in Tab. 1. For other characters see generic description above. Etymology. This remarkable and rare species is named to honor Mr. Carlo Giusto, an enthusiastic student of Apionidae , for his passion for these inconspicuous weevils, and shared interest in the genus Hellerenius .