treatments-xml/data/D8/38/E0/D838E015FFA0B055FF5DC17396C57FA2.xml

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<document id="36601F5E061BB25269F436ACD74D1DC5" ID-CLB-Dataset="307866" ID-DOI="10.11646/zootaxa.5570.2.1" ID-GBIF-Dataset="185fad02-aee4-4e05-9295-ea36217424d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734877" ID-ZooBank="7177B69B-6132-485D-87C7-4EA807692757" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737737014959" checkinUser="plazi" docAuthor="Rojas-Arias, Laura, Griswold, Terry, Hinojosa-Díaz, Ismael, Morrone, Juan J. &amp; Barajas, Ricardo Ayala" docDate="2025" docId="D838E015FFA0B055FF5DC17396C57FA2" docLanguage="en" docName="zootaxa.5570.2.1.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heriades Spinola 1808" docType="key" docVersion="5" lastPageNumber="207" masterDocId="2401986DFFA4B053FFCAC76393587A50" masterDocTitle="Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species" masterLastPageNumber="259" masterPageNumber="201" pageNumber="205" updateTime="1738007222907" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4C679DCD1DAF9CD47C29957AE964FC7F">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<mods:namePart id="E045793B2B7B30B3B132E9AA850F01F3">Rojas-Arias, Laura</mods:namePart>
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<mods:affiliation id="3EC5435FB6DFA0A458C6040995A7A2CC">Posgrado en Ciencias Biológicas, Universidad Nacional Autónoma de México (Posgrado en Ciencias Biológicas, UNAM. Unidad de</mods:affiliation>
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<mods:namePart id="F13C9C6D0FA0907BF740F4A4A48A2D8C">Griswold, Terry</mods:namePart>
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<mods:namePart id="BE7AD291747CAD6E86B48922B50E801B">Hinojosa-Díaz, Ismael</mods:namePart>
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<mods:namePart id="83B4E07F5081D53DB0DE98008ED7C5E6">Morrone, Juan J.</mods:namePart>
<mods:affiliation id="CA6D5CCD685F37AC4CEE7BD00D62D03A">Museo de Zoología “ Alfonso L. Herrera ”, Departamento de Biología Evolutiva, Facultad de Ciencias, Universidad Nacional Autónoma</mods:affiliation>
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<mods:namePart id="4C157894A24CDAD1603A6972035F2C0C">Barajas, Ricardo Ayala</mods:namePart>
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<paragraph id="502E5103FFA0B057FF5DC17397E07C7A" blockId="4.[151,1208,1551,1578]" box="[151,1208,1551,1578]" pageId="4" pageNumber="205">
<heading id="0B66E66FFFA0B057FF5DC17397E07C7A" bold="true" box="[151,1208,1551,1578]" fontSize="11" level="1" pageId="4" pageNumber="205" reason="1">
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Key to species of
<taxonomicName id="97912A80FFA0B057FEA4C16C92837C7A" ID-CoL="4VSY" authorityName="Spinola" authorityYear="1808" box="[366,475,1551,1578]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="62E58D11FFA0B057FEA4C16C92837C7A" bold="true" box="[366,475,1551,1578]" italics="true" pageId="4" pageNumber="205">Heriades</emphasis>
</taxonomicName>
for the Neotropical region of
<collectingCountry id="28861193FFA0B057FC87C17390F07C7A" box="[845,936,1552,1578]" name="Mexico" pageId="4" pageNumber="205">Mexico</collectingCountry>
and Central America
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<paragraph id="502E5103FFA0B057FF5DC13B93877C22" blockId="4.[151,1437,1624,2031]" box="[151,223,1624,1650]" pageId="4" pageNumber="205">
<emphasis id="62E58D11FFA0B057FF5DC13B93877C22" bold="true" box="[151,223,1624,1650]" pageId="4" pageNumber="205">Males</emphasis>
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<emphasis id="62E58D11FFA0B057FF5DC11A93FB7CDE" bold="true" box="[151,163,1657,1678]" pageId="4" pageNumber="205">1</emphasis>
Posterior margin of S1 on ventral view truncate medially (
<figureCitation id="C8AA4D86FFA0B057FCD2C11A90EA7CDE" box="[792,946,1657,1679]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="4" pageNumber="205">Fig. 2E, J, K, L</figureCitation>
), (with two small, triangular sublateral projections for
<taxonomicName id="97912A80FFA0B057FECDC1F4920C7CFC" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[263,340,1687,1708]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="nora" status="sp. nov.">
<emphasis id="62E58D11FFA0B057FECDC1F4920C7CFC" box="[263,340,1687,1708]" italics="true" pageId="4" pageNumber="205">H. nora</emphasis>
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<taxonomicNameLabel id="79D6306AFFA0B057FE94C1F492F47CFC" box="[350,428,1687,1708]" pageId="4" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="C8AA4D86FFA0B057FE71C1F591577CFB" box="[443,527,1686,1708]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="4" pageNumber="205">Fig. 2M</figureCitation>
), or medially with broad emargination forming two lobes (
<figureCitation id="C8AA4D86FFA0B057FB94C1F597B77CFB" box="[1118,1263,1686,1708]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="4" pageNumber="205">Fig. 2A, D, G</figureCitation>
). (Sometimes the morphological posterior margin is hidden by a large transversal process)......................................... 2
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<keyLead id="EB60F236FFA0B057FF5DC1B296C47D53" pageId="4" pageNumber="205">
- Posterior margin of S1 on ventral view medially produced to a rounded or pointed apex (
<figureCitation id="C8AA4D86FFA0B057FBABC1B296797CB6" box="[1121,1313,1745,1767]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="4" pageNumber="205">Fig. 2C, F, H, I, N</figureCitation>
), sometimes produced with minute emargination at apex (
<figureCitation id="C8AA4D86FFA0B057FD5AC18D91AF7D53" box="[656,759,1774,1796]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="4" pageNumber="205">Fig. 2B, O</figureCitation>
)...................................................... 9
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<keyStep id="EB6549A6FFA0B057FF5DC06896C47D6E" pageId="4" pageNumber="205">
<paragraph id="502E5103FFA0B057FF5DC06896C47D71" blockId="4.[151,1437,1624,2031]" box="[151,1436,1803,1825]" pageId="4" pageNumber="205">
<keyLead id="EB60F236FFA0B057FF5DC06896C47D71" box="[151,1436,1803,1825]" pageId="4" pageNumber="205">
<emphasis id="62E58D11FFA0B057FF5DC068939D7D71" bold="true" box="[151,197,1803,1825]" pageId="4" pageNumber="205">2 (1)</emphasis>
S
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profile with strong ventral process (
<figureCitation id="C8AA4D86FFA0B057FDBAC06890757D71" box="[624,813,1803,1825]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="4" pageNumber="205">Fig. 2A, D, G, J, M</figureCitation>
)................................................. 3
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</paragraph>
<paragraph id="502E5103FFA0B057FF5DC04A96C47D6E" blockId="4.[151,1437,1624,2031]" box="[151,1436,1833,1855]" pageId="4" pageNumber="205">
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- S
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profile with rounded low angle (
<figureCitation id="C8AA4D86FFA0B057FD9AC04A91897D6E" box="[592,721,1833,1855]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="4" pageNumber="205">Fig. 2E, K, L</figureCitation>
)......................................................... 7
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<paragraph id="502E5103FFA0B057FF5DC02596C57DC6" blockId="4.[151,1437,1624,2031]" pageId="4" pageNumber="205">
<keyLead id="EB60F236FFA0B057FF5DC02596C57DC6" pageId="4" pageNumber="205">
<emphasis id="62E58D11FFA0B057FF5DC025939D7D0C" bold="true" box="[151,197,1862,1884]" pageId="4" pageNumber="205">3 (2)</emphasis>
S2 and S3 with long, white, plumose hairs; apex of S5 with posterior margin wavy and with two small tufts of capitate hairs directed laterally, on posterior-lateral angles (
<figureCitation id="C8AA4D86FFA0B057FD54C00091AF7D29" box="[670,759,1891,1913]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734958" httpUri="https://zenodo.org/record/14734958/files/figure.png" pageId="4" pageNumber="205">Fig. 27E</figureCitation>
); apex of S6 with a single central tuft of hooked hairs, increasing its length outwards (
<figureCitation id="C8AA4D86FFA0B057FE43C0E292847DC6" box="[393,476,1921,1943]" captionStart="FIGURE 27" captionStartId="45.[151,250,1943,1967]" captionTargetBox="[159,1419,192,1905]" captionTargetId="figure-18@45.[159,1428,179,1917]" captionTargetPageId="45" captionText="FIGURE 27. Heriades nora sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734958" httpUri="https://zenodo.org/record/14734958/files/figure.png" pageId="4" pageNumber="205">Fig. 27F</figureCitation>
)...........................................................
<taxonomicName id="97912A80FFA0B057FB0BC0E296147DC6" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1217,1356,1921,1942]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="nora" status="sp. nov.">
<emphasis id="62E58D11FFA0B057FB0BC0E296147DC6" box="[1217,1356,1921,1942]" italics="true" pageId="4" pageNumber="205">Heriades nora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA0B057FA98C0E296C57DC6" box="[1362,1437,1921,1942]" pageId="4" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
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<paragraph id="502E5103FFA0B057FF5DC0FC96C47DBE" blockId="4.[151,1437,1624,2031]" pageId="4" pageNumber="205">
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- S2 and S3 bare, without long, white, plumose hairs; apex of S5 with two rectangular lobes, subtriangular medially, one transverse line of capitate hairs increasing in length towards sides (
<figureCitation id="C8AA4D86FFA0B057FCCBC0D890137D81" box="[769,843,1979,2001]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734896" httpUri="https://zenodo.org/record/14734896/files/figure.png" pageId="4" pageNumber="205">Fig. 7E</figureCitation>
,
<figureCitation id="C8AA4D86FFA0B057FC9FC0DF90267D81" box="[853,894,1980,2001]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734926" httpUri="https://zenodo.org/record/14734926/files/figure.png" pageId="4" pageNumber="205">17E</figureCitation>
,
<figureCitation id="C8AA4D86FFA0B057FC42C0D890E97D81" box="[904,945,1979,2001]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734952" httpUri="https://zenodo.org/record/14734952/files/figure.png" pageId="4" pageNumber="205">25E</figureCitation>
,
<figureCitation id="C8AA4D86FFA0B057FC71C0DF90BD7D81" box="[955,997,1980,2001]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734974" httpUri="https://zenodo.org/record/14734974/files/figure.png" pageId="4" pageNumber="205">31E</figureCitation>
); apex of S6 with two tufts (
<figureCitation id="C8AA4D86FFA0B057FB32C0D896137D81" box="[1272,1355,1979,2001]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734952" httpUri="https://zenodo.org/record/14734952/files/figure.png" pageId="4" pageNumber="205">Fig. 25F</figureCitation>
,
<figureCitation id="C8AA4D86FFA0B057FA9FC0DF96237D81" box="[1365,1403,1980,2001]" captionStart="FIGURE 30" captionStartId="49.[151,250,1071,1095]" captionTargetBox="[159,1428,157,1045]" captionTargetId="figure-18@49.[151,1436,149,1045]" captionTargetPageId="49" captionText="FIGURE 30. Known distribution records for Heriades viridiana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14734968" httpUri="https://zenodo.org/record/14734968/files/figure.png" pageId="4" pageNumber="205">30F</figureCitation>
) or a line of hairs (
<figureCitation id="C8AA4D86FFA0B057FEB8C0BA92E17DBE" box="[370,441,2009,2031]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734896" httpUri="https://zenodo.org/record/14734896/files/figure.png" pageId="4" pageNumber="205">Fig. 7F</figureCitation>
,
<figureCitation id="C8AA4D86FFA0B057FE09C0BA92B17DBE" box="[451,489,2009,2030]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734926" httpUri="https://zenodo.org/record/14734926/files/figure.png" pageId="4" pageNumber="205">17F</figureCitation>
)............................................................................ 4
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<paragraph id="502E5103FFA1B056FF5DC7F496C47AFD" blockId="5.[151,1437,151,1581]" box="[151,1436,151,173]" pageId="5" pageNumber="206">
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<emphasis id="62E58D11FFA1B056FF5DC7F4939D7AFD" bold="true" box="[151,197,151,173]" pageId="5" pageNumber="206">4 (3)</emphasis>
Process of S
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posterior view with one cusp, rounded, quadrate or slightly acute (
<figureCitation id="C8AA4D86FFA1B056FBCAC7F497607AFC" box="[1024,1080,151,173]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="5" pageNumber="206">Fig. 2</figureCitation>
Ac, Gc, Jc)................... 5
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</paragraph>
<paragraph id="502E5103FFA1B056FF5DC7D696C57AB7" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC7D696C57AB7" pageId="5" pageNumber="206">
- Process of S
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posterior view with two rounded cusps, divided by a soft small emargination (
<figureCitation id="C8AA4D86FFA1B056FBBAC7D797E37A9A" box="[1136,1211,180,202]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="5" pageNumber="206">Fig. 2D</figureCitation>
)................................................................................................
<taxonomicName id="97912A80FFA1B056FB54C7B196147AB7" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1182,1356,210,231]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="briceida" status="sp. nov.">
<emphasis id="62E58D11FFA1B056FB54C7B196147AB7" box="[1182,1356,210,231]" italics="true" pageId="5" pageNumber="206">Heriades briceida</emphasis>
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<taxonomicNameLabel id="79D6306AFFA1B056FA98C7B196C57AB7" box="[1362,1437,210,231]" pageId="5" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="502E5103FFA1B056FF5DC78C96C57B6F" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
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<emphasis id="62E58D11FFA1B056FF5DC78C939D7B55" bold="true" box="[151,197,239,261]" pageId="5" pageNumber="206">5 (4)</emphasis>
Flagellum brown dorsally and yellow ventrally (
<figureCitation id="C8AA4D86FFA1B056FD71C78C904E7B54" box="[699,790,239,261]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734974" httpUri="https://zenodo.org/record/14734974/files/figure.png" pageId="5" pageNumber="206">Fig. 31A</figureCitation>
); pronotal lobe with white, thin, plumose pubescence as in rest of mesosoma; ocelli large, 2.5 times diameter of a puncture (
<figureCitation id="C8AA4D86FFA1B056FCD5C66F90217B72" box="[799,889,268,290]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734974" httpUri="https://zenodo.org/record/14734974/files/figure.png" pageId="5" pageNumber="206">Fig. 31B</figureCitation>
); process on S
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posterior view with acute apex (
<figureCitation id="C8AA4D86FFA1B056FABFC66F93B67B6F" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="5" pageNumber="206">Fig. 2</figureCitation>
Gc)..........................................................................
<taxonomicName id="97912A80FFA1B056FB5CC64996147B6F" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1174,1356,298,319]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="viridiana" status="sp. nov.">
<emphasis id="62E58D11FFA1B056FB5CC64996147B6F" box="[1174,1356,298,319]" italics="true" pageId="5" pageNumber="206">Heriades viridiana</emphasis>
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</paragraph>
<paragraph id="502E5103FFA1B056FF5DC62B96C47BC8" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC62B96C47BC8" pageId="5" pageNumber="206">
- Flagellum light brown (
<figureCitation id="C8AA4D86FFA1B056FE0FC62491457B0C" box="[453,541,327,349]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734926" httpUri="https://zenodo.org/record/14734926/files/figure.png" pageId="5" pageNumber="206">Fig. 17A</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FDECC624910B7B0C" box="[550,595,327,348]" captionStart="FIGURE 24" captionStartId="41.[151,250,1098,1122]" captionTargetBox="[160,1428,187,1063]" captionTargetId="figure-18@41.[151,1436,179,1072]" captionTargetPageId="41" captionText="FIGURE 24. Known distribution records for Heriades micheneri." figureDoi="http://doi.org/10.5281/zenodo.14734948" httpUri="https://zenodo.org/record/14734948/files/figure.png" pageId="5" pageNumber="206">24A</figureCitation>
); pronotal lobe with patch of distinctly thick, yellow, or white plumose hairs contrasting with the rest of mesosoma; ocelli small, 1.5 times diameter of a puncture (
<figureCitation id="C8AA4D86FFA1B056FC54C60797497B2A" box="[926,1041,356,378]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734926" httpUri="https://zenodo.org/record/14734926/files/figure.png" pageId="5" pageNumber="206">Fig. 17B, D</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FBD0C60697397B2A" box="[1050,1121,357,378]" captionStart="FIGURE 24" captionStartId="41.[151,250,1098,1122]" captionTargetBox="[160,1428,187,1063]" captionTargetId="figure-18@41.[151,1436,179,1072]" captionTargetPageId="41" captionText="FIGURE 24. Known distribution records for Heriades micheneri." figureDoi="http://doi.org/10.5281/zenodo.14734948" httpUri="https://zenodo.org/record/14734948/files/figure.png" pageId="5" pageNumber="206">24B, D</figureCitation>
); process on S
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posterior view with subquadrangular apex (
<figureCitation id="C8AA4D86FFA1B056FE39C6E191747BC7" box="[499,556,386,408]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="5" pageNumber="206">Fig. 2</figureCitation>
Ac, Jc).................................................................. 6
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<keyLead id="EB60F236FFA1B056FF5DC6FC96C5785D" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC6FC939D7BE5" bold="true" box="[151,197,415,437]" pageId="5" pageNumber="206">6 (5)</emphasis>
Outer surface of mandible with two strong tubercles, increasing in size from base to apex and abruptly truncate in middle of mandible; clypeus with yellowish plumose hairs, in profile forming a positive diagonal (
<figureCitation id="C8AA4D86FFA1B056FBBBC6DF97957B82" box="[1137,1229,444,466]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734926" httpUri="https://zenodo.org/record/14734926/files/figure.png" pageId="5" pageNumber="206">Fig. 17A</figureCitation>
); apex of gonostylus subquadrangular with preapical, long, plumose hairs (
<figureCitation id="C8AA4D86FFA1B056FD26C6B9903C7BBF" box="[748,868,474,496]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734926" httpUri="https://zenodo.org/record/14734926/files/figure.png" pageId="5" pageNumber="206">Fig. 17G, H</figureCitation>
); apical margin of S6 with transverse line of capitate hairs (
<figureCitation id="C8AA4D86FFA1B056FF23C6949264785C" box="[233,316,503,525]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734926" httpUri="https://zenodo.org/record/14734926/files/figure.png" pageId="5" pageNumber="206">Fig. 17F</figureCitation>
)......................................................................
<taxonomicName id="97912A80FFA1B056FB5FC6949614785C" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1173,1356,503,524]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="danuncia" status="sp. nov.">
<emphasis id="62E58D11FFA1B056FB5FC6949614785C" box="[1173,1356,503,524]" italics="true" pageId="5" pageNumber="206">Heriades danuncia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA1B056FA98C69B96C5785D" box="[1362,1437,504,525]" pageId="5" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA1B056FF5DC57696C47834" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC57696C47834" pageId="5" pageNumber="206">
- Outer surface of mandible smooth, without tubercles; clypeus with white plumose hairs, in profile without a clear pattern (
<figureCitation id="C8AA4D86FFA1B056FABFC57792567818" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734952" httpUri="https://zenodo.org/record/14734952/files/figure.png" pageId="5" pageNumber="206">Fig. 25A</figureCitation>
); gonostylus rounded without hairs (
<figureCitation id="C8AA4D86FFA1B056FDA4C55191BC7817" box="[622,740,562,584]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734952" httpUri="https://zenodo.org/record/14734952/files/figure.png" pageId="5" pageNumber="206">Fig. 25G, H</figureCitation>
); S6 with two tufts of hooked hairs directed anteriorly (
<figureCitation id="C8AA4D86FFA1B056FB30C55196157818" box="[1274,1357,562,584]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734952" httpUri="https://zenodo.org/record/14734952/files/figure.png" pageId="5" pageNumber="206">Fig. 25F</figureCitation>
).........................................................................................
<taxonomicName id="97912A80FFA1B056FB14C52C96C47834" ID-CoL="3KYS3" box="[1246,1436,591,612]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="micheneri">
<emphasis id="62E58D11FFA1B056FB14C52C96C47834" box="[1246,1436,591,612]" italics="true" pageId="5" pageNumber="206">Heriades micheneri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA1B056FF5DC50E96C47962" pageId="5" pageNumber="206">
<paragraph id="502E5103FFA1B056FF5DC50E96C578ED" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC50E96C578ED" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC50E939D78D2" bold="true" box="[151,197,620,642]" pageId="5" pageNumber="206">7 (2)</emphasis>
Large-sized bees (
<quantity id="9769FCE6FFA1B056FE5AC50E915178D2" box="[400,521,621,642]" metricMagnitude="-3" metricUnit="m" metricValue="7.45" metricValueMax="7.7" metricValueMin="7.2" pageId="5" pageNumber="206" unit="mm" value="7.45" valueMax="7.7" valueMin="7.2">7.27.7 mm</quantity>
); Head and mesosoma with plumose, thin, long, white, scattered hairs, evenly distributed on its entire surface (
<figureCitation id="C8AA4D86FFA1B056FEBAC5E9929078CF" box="[368,456,650,672]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734982" httpUri="https://zenodo.org/record/14734982/files/figure.png" pageId="5" pageNumber="206">Fig. 33B</figureCitation>
); S5 with two tufts of capitate hairs directed medially (
<figureCitation id="C8AA4D86FFA1B056FC1FC5E9977478CF" box="[981,1068,650,672]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734982" httpUri="https://zenodo.org/record/14734982/files/figure.png" pageId="5" pageNumber="206">Fig. 33E</figureCitation>
); S6 with two subapical, dense tufts of simple hairs directed downward (
<figureCitation id="C8AA4D86FFA1B056FDE8C5C4912078EC" box="[546,632,679,701]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734982" httpUri="https://zenodo.org/record/14734982/files/figure.png" pageId="5" pageNumber="206">Fig. 33F</figureCitation>
); restricted to
<collectingCountry id="28861193FFA1B056FCCBC5C4901278ED" box="[769,842,679,701]" name="Mexico" pageId="5" pageNumber="206">Mexico</collectingCountry>
(
<figureCitation id="C8AA4D86FFA1B056FC92C5C490C578EC" box="[856,925,679,701]" captionStart="FIGURE 32" captionStartId="54.[151,250,1108,1132]" captionTargetBox="[159,1428,187,1083]" captionTargetId="figure-22@54.[151,1436,179,1083]" captionTargetPageId="54" captionText="FIGURE 32. Known distribution records for Heriades vulcanica sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14734976" httpUri="https://zenodo.org/record/14734976/files/figure.png" pageId="5" pageNumber="206">Fig. 32</figureCitation>
)...................
<taxonomicName id="97912A80FFA1B056FB5BC5C4961478EC" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1169,1356,679,700]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="vulcanica" status="sp. nov.">
<emphasis id="62E58D11FFA1B056FB5BC5C4961478EC" box="[1169,1356,679,700]" italics="true" pageId="5" pageNumber="206">Heriades vulcanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA1B056FA98C5CB96C578ED" box="[1362,1437,680,701]" pageId="5" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA1B056FF5DC5A696C47962" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC5A696C47962" pageId="5" pageNumber="206">
- Small-sized bees (
<quantity id="9769FCE6FFA1B056FE5FC5A792A6788A" box="[405,510,708,730]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="6.0" metricValueMin="4.2" pageId="5" pageNumber="206" unit="mm" value="5.1" valueMax="6.0" valueMin="4.2">4.26 mm</quantity>
); Head and mesosoma with plumose, thin, short, white hairs, densely distributed in tufts, and frons, scutum and mesepisternum with large disc areas with simple, very short hairs on each puncture (
<figureCitation id="C8AA4D86FFA1B056FB66C581965A78A8" box="[1196,1282,738,760]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734920" httpUri="https://zenodo.org/record/14734920/files/figure.png" pageId="5" pageNumber="206">Fig. 15B</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FAC1C581966F78A7" box="[1291,1335,738,759]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734946" httpUri="https://zenodo.org/record/14734946/files/figure.png" pageId="5" pageNumber="206">23B</figureCitation>
); posterior margin of S5 with one line of scattered, capitate hairs and simple hairs medially (
<figureCitation id="C8AA4D86FFA1B056FC2CC59C97637945" box="[998,1083,767,789]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734920" httpUri="https://zenodo.org/record/14734920/files/figure.png" pageId="5" pageNumber="206">Fig. 15E</figureCitation>
), or two lines of dense, capitate hairs (
<figureCitation id="C8AA4D86FFA1B056FF23C47F92187962" box="[233,320,796,818]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734946" httpUri="https://zenodo.org/record/14734946/files/figure.png" pageId="5" pageNumber="206">Fig. 23E</figureCitation>
); posterior margin of S6 with one line of capitated hairs (
<figureCitation id="C8AA4D86FFA1B056FCAFC47F90E07962" box="[869,952,796,818]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734920" httpUri="https://zenodo.org/record/14734920/files/figure.png" pageId="5" pageNumber="206">Fig. 15F</figureCitation>
) or two spikes (
<figureCitation id="C8AA4D86FFA1B056FB98C47F97FE7962" box="[1106,1190,796,818]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734946" httpUri="https://zenodo.org/record/14734946/files/figure.png" pageId="5" pageNumber="206">Fig. 23F</figureCitation>
).................. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA1B056FF5DC45996C479B2" pageId="5" pageNumber="206">
<paragraph id="502E5103FFA1B056FF5DC45996C579DA" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC45996C579DA" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC459939C791F" bold="true" box="[151,196,825,847]" pageId="5" pageNumber="206">8 (7)</emphasis>
Clypeus, frons and paraocular area with golden, plumose hairs (
<figureCitation id="C8AA4D86FFA1B056FCFEC45990D3791F" box="[820,907,826,848]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734946" httpUri="https://zenodo.org/record/14734946/files/figure.png" pageId="5" pageNumber="206">Fig. 23A</figureCitation>
); interantennal distance 1.6 times antennocular distance; posterior margin of S5 with two dense lines of capitate hairs (
<figureCitation id="C8AA4D86FFA1B056FCFFC43490D4793C" box="[821,908,855,877]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734946" httpUri="https://zenodo.org/record/14734946/files/figure.png" pageId="5" pageNumber="206">Fig. 23E</figureCitation>
); posterior margin of S6 with two subapical, long, thin spikes enlarged at base and quadrate al tip (
<figureCitation id="C8AA4D86FFA1B056FDB5C417918C79DA" box="[639,724,884,906]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734946" httpUri="https://zenodo.org/record/14734946/files/figure.png" pageId="5" pageNumber="206">Fig. 23F</figureCitation>
); restricted to
<collectingCountry id="28861193FFA1B056FC96C416909C79DA" box="[860,964,884,906]" name="Costa Rica" pageId="5" pageNumber="206">Costa Rica</collectingCountry>
(
<figureCitation id="C8AA4D86FFA1B056FC1AC417974E79DA" box="[976,1046,884,906]" captionStart="FIGURE 22" captionStartId="38.[151,250,1068,1092]" captionTargetBox="[159,1428,157,1033]" captionTargetId="figure-22@38.[151,1436,149,1042]" captionTargetPageId="38" captionText="FIGURE 22. Known distribution records for Heriades mariana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14734942" httpUri="https://zenodo.org/record/14734942/files/figure.png" pageId="5" pageNumber="206">Fig. 22</figureCitation>
)...........
<taxonomicName id="97912A80FFA1B056FB6AC417961679DA" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1184,1358,884,906]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="mariana" status="sp. nov.">
<emphasis id="62E58D11FFA1B056FB6AC417961679DA" box="[1184,1358,884,906]" italics="true" pageId="5" pageNumber="206">Heriades mariana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA1B056FA99C41696C579DA" box="[1363,1437,885,906]" pageId="5" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA1B056FF5DC4F196C479B2" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC4F196C479B2" pageId="5" pageNumber="206">
- Clypeus, frons and paraocular area with white, plumose hairs (
<figureCitation id="C8AA4D86FFA1B056FCE4C4F190DE79F7" box="[814,902,914,936]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734946" httpUri="https://zenodo.org/record/14734946/files/figure.png" pageId="5" pageNumber="206">Fig. 23A</figureCitation>
); interantennal distance 1.8 times antennocular distance; posterior margin of S5 with one line of capitate hairs (
<figureCitation id="C8AA4D86FFA1B056FD29C4CC90617995" box="[739,825,943,965]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734920" httpUri="https://zenodo.org/record/14734920/files/figure.png" pageId="5" pageNumber="206">Fig. 15E</figureCitation>
); posterior margin of S6 with one preapical line of capitate hairs (
<figureCitation id="C8AA4D86FFA1B056FF23C4AF926F79B2" box="[233,311,972,994]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734920" httpUri="https://zenodo.org/record/14734920/files/figure.png" pageId="5" pageNumber="206">Fig. 15f</figureCitation>
)..............................................................................
<taxonomicName id="97912A80FFA1B056FB3CC4AF96C479B2" ID-CoL="3KYQL" authorityName="Michener" authorityYear="1943" box="[1270,1436,972,994]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="currani">
<emphasis id="62E58D11FFA1B056FB3CC4AF96C479B2" box="[1270,1436,972,994]" italics="true" pageId="5" pageNumber="206">Heriades currani</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA1B056FF5DC48996C47E4C" pageId="5" pageNumber="206">
<paragraph id="502E5103FFA1B056FF5DC48996C579AF" blockId="5.[151,1437,151,1581]" box="[151,1437,1001,1024]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC48996C579AF" box="[151,1437,1001,1024]" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC489939D79AF" bold="true" box="[151,197,1001,1023]" pageId="5" pageNumber="206">9 (1)</emphasis>
Pedicel and scape yellowish (
<figureCitation id="C8AA4D86FFA1B056FE37C489910B79AF" box="[509,595,1002,1024]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734908" httpUri="https://zenodo.org/record/14734908/files/figure.png" pageId="5" pageNumber="206">Fig. 11A</figureCitation>
).................................................
<taxonomicName id="97912A80FFA1B056FB6FC489961479AF" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1189,1356,1002,1023]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="cristina" status="sp. nov.">
<emphasis id="62E58D11FFA1B056FB6FC489961479AF" box="[1189,1356,1002,1023]" italics="true" pageId="5" pageNumber="206">Heriades cristina</emphasis>
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<taxonomicNameLabel id="79D6306AFFA1B056FA98C48996C579AF" box="[1362,1437,1002,1023]" pageId="5" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA1B056FF5DC36B96C47E4C" blockId="5.[151,1437,151,1581]" box="[151,1436,1031,1053]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC36B96C47E4C" box="[151,1436,1031,1053]" pageId="5" pageNumber="206">- Pedicel and scape black............................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA1B056FF5DC34796C47EC2" pageId="5" pageNumber="206">
<paragraph id="502E5103FFA1B056FF5DC34796C47E07" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC34796C47E07" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC34793897E6A" bold="true" box="[151,209,1060,1082]" pageId="5" pageNumber="206">10 (9)</emphasis>
Posterior margin of S5 with simple hairs medially (
<figureCitation id="C8AA4D86FFA1B056FD04C34790407E6A" box="[718,792,1060,1082]" captionStart="FIGURE 9" captionStartId="18.[151,250,1944,1968]" captionTargetBox="[152,1422,179,1905]" captionTargetId="figure-22@18.[152,1435,179,1918]" captionTargetPageId="18" captionText="FIGURE 9. Heriades bruneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734902" httpUri="https://zenodo.org/record/14734902/files/figure.png" pageId="5" pageNumber="206">Fig. 9E</figureCitation>
); apical terga on metasoma with orange pubescence (
<figureCitation id="C8AA4D86FFA1B056FAD2C34796397E6A" box="[1304,1377,1060,1082]" captionStart="FIGURE 9" captionStartId="18.[151,250,1944,1968]" captionTargetBox="[152,1422,179,1905]" captionTargetId="figure-22@18.[152,1435,179,1918]" captionTargetPageId="18" captionText="FIGURE 9. Heriades bruneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734902" httpUri="https://zenodo.org/record/14734902/files/figure.png" pageId="5" pageNumber="206">Fig. 9B</figureCitation>
).........................................................................................
<taxonomicName id="97912A80FFA1B056FB3CC32196C47E07" ID-CoL="3KYPX" authorityName="Titus" authorityYear="1904" box="[1270,1436,1090,1111]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="bruneri">
<emphasis id="62E58D11FFA1B056FB3CC32196C47E07" box="[1270,1436,1090,1111]" italics="true" pageId="5" pageNumber="206">Heriades bruneri</emphasis>
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</paragraph>
<paragraph id="502E5103FFA1B056FF5DC30396C47EC2" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC30396C47EC2" pageId="5" pageNumber="206">
- Posterior margin of S5 with capitate hairs medially (
<figureCitation id="C8AA4D86FFA1B056FD38C33C90677E25" box="[754,831,1119,1141]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734890" httpUri="https://zenodo.org/record/14734890/files/figure.png" pageId="5" pageNumber="206">Fig. 5E</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FC86C33C902D7E24" box="[844,885,1119,1140]" captionStart="FIGURE 13" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1897]" captionTargetId="figure-22@24.[161,1426,179,1917]" captionTargetPageId="24" captionText="FIGURE 13. Heriades crucifera. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734914" httpUri="https://zenodo.org/record/14734914/files/figure.png" pageId="5" pageNumber="206">13E</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FC48C33C90F37E24" box="[898,939,1119,1140]" captionStart="FIGURE 19" captionStartId="32.[151,250,1943,1967]" captionTargetBox="[163,1412,184,1905]" captionTargetId="figure-22@32.[163,1425,179,1917]" captionTargetPageId="32" captionText="FIGURE 19. Heriades leavitti. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734934" httpUri="https://zenodo.org/record/14734934/files/figure.png" pageId="5" pageNumber="206">19E</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FC72C33C90B87E24" box="[952,992,1119,1140]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="5" pageNumber="206">21E</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FC24C33C974F7E24" box="[1006,1047,1119,1140]" captionStart="FIGURE 29" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[162,1407,188,1896]" captionTargetId="figure-22@48.[162,1426,179,1916]" captionTargetPageId="48" captionText="FIGURE 29. Heriades texana. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734964" httpUri="https://zenodo.org/record/14734964/files/figure.png" pageId="5" pageNumber="206">29E</figureCitation>
); apical terga on metasoma with white pubescence......................................................................................... 11
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<paragraph id="502E5103FFA1B056FF5DC3F996C47EFF" blockId="5.[151,1437,151,1581]" box="[151,1436,1177,1200]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC3F996C47EFF" box="[151,1436,1177,1200]" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC3F993847EFF" bold="true" box="[151,220,1177,1199]" pageId="5" pageNumber="206">11 (10)</emphasis>
Posterior margin of S1 with minute emargination (
<figureCitation id="C8AA4D86FFA1B056FD0CC3F991A77EE0" box="[710,767,1178,1200]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="5" pageNumber="206">Fig. 2</figureCitation>
Bb, Ob)............................................... 12
</keyLead>
</paragraph>
<paragraph id="502E5103FFA1B056FF5DC3DB96C47E9C" blockId="5.[151,1437,151,1581]" box="[151,1436,1207,1229]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC3DB96C47E9C" box="[151,1436,1207,1229]" pageId="5" pageNumber="206">
- Posterior margin of S1 entire, without minute emargination (
<figureCitation id="C8AA4D86FFA1B056FCEDC3D490387E9D" box="[807,864,1207,1229]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="5" pageNumber="206">Fig. 2</figureCitation>
Fb, Hb, Nb)................................... 13
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</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA1B056FF5DC3B796C57F12" pageId="5" pageNumber="206">
<paragraph id="502E5103FFA1B056FF5DC3B796C47F57" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC3B796C47F57" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC3B793847EBA" bold="true" box="[151,220,1236,1258]" pageId="5" pageNumber="206">12 (11)</emphasis>
Apical margin of clypeus entire, without emargination; apex of S6 with two tufts of short, simple hairs (
<figureCitation id="C8AA4D86FFA1B056FB7DC3B796537EBA" box="[1207,1291,1236,1258]" captionStart="FIGURE 13" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1897]" captionTargetId="figure-22@24.[161,1426,179,1917]" captionTargetPageId="24" captionText="FIGURE 13. Heriades crucifera. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734914" httpUri="https://zenodo.org/record/14734914/files/figure.png" pageId="5" pageNumber="206">Fig. 13F</figureCitation>
); S6 with acute apex (
<figureCitation id="C8AA4D86FFA1B056FED6C39192377F57" box="[284,367,1266,1288]" captionStart="FIGURE 13" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1897]" captionTargetId="figure-22@24.[161,1426,179,1917]" captionTargetPageId="24" captionText="FIGURE 13. Heriades crucifera. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734914" httpUri="https://zenodo.org/record/14734914/files/figure.png" pageId="5" pageNumber="206">Fig. 13F</figureCitation>
).........................................................................
<taxonomicName id="97912A80FFA1B056FB2CC39196C47F57" ID-CoL="8SG9K" authorityName="Cockerell" authorityYear="1897" box="[1254,1436,1266,1287]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="62E58D11FFA1B056FB2CC39196C47F57" box="[1254,1436,1266,1287]" italics="true" pageId="5" pageNumber="206">Heriades crucifera</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA1B056FF5DC27396C57F12" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC27396C57F12" pageId="5" pageNumber="206">
- Apical margin of clypeus with emargination; apex of S6 with two tufts of 3 to 4 thick, long hairs resembling a spike (
<figureCitation id="C8AA4D86FFA1B056FA8DC26C96C87F75" box="[1351,1424,1295,1317]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734890" httpUri="https://zenodo.org/record/14734890/files/figure.png" pageId="5" pageNumber="206">Fig. 5F</figureCitation>
); S6 with rounded apex (
<figureCitation id="C8AA4D86FFA1B056FE0BC24F91507F12" box="[449,520,1324,1346]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734890" httpUri="https://zenodo.org/record/14734890/files/figure.png" pageId="5" pageNumber="206">Fig. 5F</figureCitation>
)......................................................
<taxonomicName id="97912A80FFA1B056FB6BC24F96147F11" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1185,1356,1324,1345]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="amanda" status="sp. nov.">
<emphasis id="62E58D11FFA1B056FB6BC24F96147F11" box="[1185,1356,1324,1345]" italics="true" pageId="5" pageNumber="206">Heriades amanda</emphasis>
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<taxonomicNameLabel id="79D6306AFFA1B056FA98C24E96C57F12" box="[1362,1437,1325,1346]" pageId="5" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<keyStep id="EB6549A6FFA1B056FF5DC22996C57FE7" pageId="5" pageNumber="206">
<paragraph id="502E5103FFA1B056FF5DC22996C47F2C" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC22996C47F2C" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC22993857F0F" bold="true" box="[151,221,1353,1375]" pageId="5" pageNumber="206">13 (10)</emphasis>
Clypeus with plumose, white hairs on its entire surface or at least dense on inferior half; posterior margin of S5 without projections, with two tufts of capitate hairs directed medially, of different lengths (
<figureCitation id="C8AA4D86FFA1B056FC33C20497177F2C" box="[1017,1103,1383,1405]" captionStart="FIGURE 18" captionStartId="31.[151,250,1098,1122]" captionTargetBox="[159,1428,187,1063]" captionTargetId="figure-18@31.[151,1436,179,1072]" captionTargetPageId="31" captionText="FIGURE 18. Known distribution records for Heriades leavitti." figureDoi="http://doi.org/10.5281/zenodo.14734930" httpUri="https://zenodo.org/record/14734930/files/figure.png" pageId="5" pageNumber="206">Fig. 18E</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FB93C20497D97F2C" box="[1113,1153,1383,1404]" captionStart="FIGURE 28" captionStartId="47.[151,250,1117,1141]" captionTargetBox="[160,1428,206,1083]" captionTargetId="figure-18@47.[151,1436,198,1091]" captionTargetPageId="47" captionText="FIGURE 28. Known distribution records for Heriades texana." figureDoi="http://doi.org/10.5281/zenodo.14734962" httpUri="https://zenodo.org/record/14734962/files/figure.png" pageId="5" pageNumber="206">28E</figureCitation>
).................... 14
</keyLead>
</paragraph>
<paragraph id="502E5103FFA1B056FF5DC2E696C57FE7" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC2E696C57FE7" pageId="5" pageNumber="206">
- Clypeus with plumose hairs seemly absent; posterior margin of S5 with two small, sub quadrangular projections, with few short, capitate hairs on top (
<figureCitation id="C8AA4D86FFA1B056FE27C2C191197FE7" box="[493,577,1442,1464]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="5" pageNumber="206">Fig. 21E</figureCitation>
)....................................................
<taxonomicName id="97912A80FFA1B056FB0AC2C196157FE7" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1216,1357,1442,1463]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="luisa" status="sp. nov.">
<emphasis id="62E58D11FFA1B056FB0AC2C196157FE7" box="[1216,1357,1442,1463]" italics="true" pageId="5" pageNumber="206">Heriades luisa</emphasis>
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<taxonomicNameLabel id="79D6306AFFA1B056FA98C2C196C57FE7" box="[1362,1437,1442,1463]" pageId="5" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="502E5103FFA1B056FF5DC2DC96C57FA1" blockId="5.[151,1437,151,1581]" pageId="5" pageNumber="206">
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<emphasis id="62E58D11FFA1B056FF5DC2DC93857F85" bold="true" box="[151,221,1471,1493]" pageId="5" pageNumber="206">14 (13)</emphasis>
Clypeus with white, plumose hairs on apical half; S
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profile, with ventral angle more than one third of distance from apex to base (
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)...........................................................................
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<emphasis id="62E58D11FFA1B056FB36C2BF96C57FA1" box="[1276,1437,1500,1521]" italics="true" pageId="5" pageNumber="206">Heriades leavitti</emphasis>
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- Clypeus with white, plumose hairs on its entire surface; S
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profile, with ventral angle one third or less of distance from apex to base (
<figureCitation id="C8AA4D86FFA1B056FEF8C17492257C7C" box="[306,381,1559,1581]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="5" pageNumber="206">Fig. 2H</figureCitation>
).........................................................................
<taxonomicName id="97912A80FFA1B056FB34C17496C47C7D" ID-CoL="6M45K" box="[1278,1436,1559,1581]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="62E58D11FFA1B056FB34C17496C47C7D" box="[1278,1436,1559,1581]" italics="true" pageId="5" pageNumber="206">Heriades texana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="502E5103FFA1B056FF5DC13F93A17C26" blockId="5.[151,1437,1628,2034]" box="[151,249,1628,1654]" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC13F93A17C26" bold="true" box="[151,249,1628,1654]" pageId="5" pageNumber="206">Females</emphasis>
</paragraph>
<keyStep id="EB6549A6FFA1B056FF5DC11F96C47C9D" pageId="5" pageNumber="206">
<paragraph id="502E5103FFA1B056FF5DC11F96C47CC2" blockId="5.[151,1437,1628,2034]" box="[151,1436,1660,1682]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC11F96C47CC2" box="[151,1436,1660,1682]" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC11F93FB7CC1" bold="true" box="[151,163,1660,1681]" pageId="5" pageNumber="206">1</emphasis>
Outer surface of mandible with one or more tubercles, or a strong elevation at base of outer and/or condylar ridge........ 2
</keyLead>
</paragraph>
<paragraph id="502E5103FFA1B056FF5DC1F996C47C9D" blockId="5.[151,1437,1628,2034]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC1F996C47C9D" pageId="5" pageNumber="206">
- Outer surface of mandible without tubercle or any noticeable elevation at base of outer and condylar ridge (except in a few specimens of
<taxonomicName id="97912A80FFA1B056FEACC1DB92937C9C" ID-CoL="3KYPX" authorityName="Titus" authorityYear="1904" box="[358,459,1719,1741]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="bruneri">
<emphasis id="62E58D11FFA1B056FEACC1DB92937C9C" box="[358,459,1719,1741]" italics="true" pageId="5" pageNumber="206">H. bruneri</emphasis>
</taxonomicName>
, they can be identified by small elevation on outer ridge and orangish hairs on posterior terga).... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA1B056FF5DC1B796C47D2C" pageId="5" pageNumber="206">
<paragraph id="502E5103FFA1B056FF5DC1B796C47D74" blockId="5.[151,1437,1628,2034]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC1B796C47D74" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC1B7939D7CBA" bold="true" box="[151,197,1748,1770]" pageId="5" pageNumber="206">2 (1)</emphasis>
Posterior edge of malar area with large, strong, subtriangular angle, similar-sized to condyle (
<figureCitation id="C8AA4D86FFA1B056FBBCC1B797967CBA" box="[1142,1230,1748,1770]" captionStart="FIGURE 13" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1897]" captionTargetId="figure-22@24.[161,1426,179,1917]" captionTargetPageId="24" captionText="FIGURE 13. Heriades crucifera. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734914" httpUri="https://zenodo.org/record/14734914/files/figure.png" pageId="5" pageNumber="206">Fig. 13C</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FB13C1B6965D7CBA" box="[1241,1285,1749,1770]" captionStart="FIGURE 29" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[162,1407,188,1896]" captionTargetId="figure-22@48.[162,1426,179,1916]" captionTargetPageId="48" captionText="FIGURE 29. Heriades texana. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734964" httpUri="https://zenodo.org/record/14734964/files/figure.png" pageId="5" pageNumber="206">29C</figureCitation>
); distance from posterior ocellus to preoccipital ridge large (2.503.35 times LO) (
<figureCitation id="C8AA4D86FFA1B056FCB9C19190957D57" box="[883,973,1778,1800]" captionStart="FIGURE 13" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1897]" captionTargetId="figure-22@24.[161,1426,179,1917]" captionTargetPageId="24" captionText="FIGURE 13. Heriades crucifera. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734914" httpUri="https://zenodo.org/record/14734914/files/figure.png" pageId="5" pageNumber="206">Fig. 13D</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FC13C191975E7D57" box="[985,1030,1778,1799]" captionStart="FIGURE 29" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[162,1407,188,1896]" captionTargetId="figure-22@48.[162,1426,179,1916]" captionTargetPageId="48" captionText="FIGURE 29. Heriades texana. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734964" httpUri="https://zenodo.org/record/14734964/files/figure.png" pageId="5" pageNumber="206">29D</figureCitation>
); punctures on T3 medium-sized or large (~0.66 DOP of scutum)................................................................................ 3
</keyLead>
</paragraph>
<paragraph id="502E5103FFA1B056FF5DC04E96C47D2C" blockId="5.[151,1437,1628,2034]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC04E96C47D2C" pageId="5" pageNumber="206">
- Posterior edge of malar area without large, strong subtriangular angle (
<figureCitation id="C8AA4D86FFA1B056FC52C04F90A87D12" box="[920,1008,1836,1858]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734920" httpUri="https://zenodo.org/record/14734920/files/figure.png" pageId="5" pageNumber="206">Fig. 15C</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FC36C04E977E7D12" box="[1020,1062,1837,1858]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734946" httpUri="https://zenodo.org/record/14734946/files/figure.png" pageId="5" pageNumber="206">23C</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FBFBC04E97057D12" box="[1073,1117,1837,1858]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734982" httpUri="https://zenodo.org/record/14734982/files/figure.png" pageId="5" pageNumber="206">33C</figureCitation>
); distance from posterior ocellus to preoccipital ridge short (1.25 times LO) (
<figureCitation id="C8AA4D86FFA1B056FD43C02991BA7D30" box="[649,738,1866,1888]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[161,1421,179,1898]" captionTargetId="figure-22@26.[161,1426,179,1917]" captionTargetPageId="26" captionText="FIGURE 15. Heriades currani. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734920" httpUri="https://zenodo.org/record/14734920/files/figure.png" pageId="5" pageNumber="206">Fig. 15D</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FD27C02990407D0F" box="[749,792,1866,1887]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734946" httpUri="https://zenodo.org/record/14734946/files/figure.png" pageId="5" pageNumber="206">23D</figureCitation>
,
<figureCitation id="C8AA4D86FFA1B056FCE8C02990177D0F" box="[802,847,1866,1887]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734982" httpUri="https://zenodo.org/record/14734982/files/figure.png" pageId="5" pageNumber="206">33D</figureCitation>
); punctures on T3 small and sparse (0.400.45 times DOP of scutum)............................................................................................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA1B056FF5DC0E796C47DA1" pageId="5" pageNumber="206">
<paragraph id="502E5103FFA1B056FF5DC0E796C47DE7" blockId="5.[151,1437,1628,2034]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC0E796C47DE7" pageId="5" pageNumber="206">
<emphasis id="62E58D11FFA1B056FF5DC0E7939D7DCA" bold="true" box="[151,197,1924,1946]" pageId="5" pageNumber="206">3 (2)</emphasis>
Ventral margin of mandible with well-defined angle (
<figureCitation id="C8AA4D86FFA1B056FD11C0E790777DCA" box="[731,815,1924,1946]" captionStart="FIGURE 29" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[162,1407,188,1896]" captionTargetId="figure-22@48.[162,1426,179,1916]" captionTargetPageId="48" captionText="FIGURE 29. Heriades texana. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734964" httpUri="https://zenodo.org/record/14734964/files/figure.png" pageId="5" pageNumber="206">Fig. 29C</figureCitation>
arrow); outer surface of mandible without notorious tubercle, the angle is joining point at base of outer and condylar ridges.........................................
<taxonomicName id="97912A80FFA1B056FB34C0C196C47DE7" ID-CoL="6M45K" box="[1278,1436,1954,1975]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="texana">
<emphasis id="62E58D11FFA1B056FB34C0C196C47DE7" box="[1278,1436,1954,1975]" italics="true" pageId="5" pageNumber="206">Heriades texana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA1B056FF5DC0A396C47DA1" blockId="5.[151,1437,1628,2034]" pageId="5" pageNumber="206">
<keyLead id="EB60F236FFA1B056FF5DC0A396C47DA1" pageId="5" pageNumber="206">
- Ventral margin of mandible straight without angle (
<figureCitation id="C8AA4D86FFA1B056FD00C0DC90797D84" box="[714,801,1983,2005]" captionStart="FIGURE 13" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1897]" captionTargetId="figure-22@24.[161,1426,179,1917]" captionTargetPageId="24" captionText="FIGURE 13. Heriades crucifera. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734914" httpUri="https://zenodo.org/record/14734914/files/figure.png" pageId="5" pageNumber="206">Fig. 13C</figureCitation>
); outer surface of mandible with notorious semicircular tubercle at base of outer and condylar ridges, from base of mandible to interior margin........................
<taxonomicName id="97912A80FFA1B056FB2CC0BF96C47DA1" ID-CoL="8SG9K" authorityName="Cockerell" authorityYear="1897" box="[1254,1436,2012,2033]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="crucifera">
<emphasis id="62E58D11FFA1B056FB2CC0BF96C47DA1" box="[1254,1436,2012,2033]" italics="true" pageId="5" pageNumber="206">Heriades crucifera</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC7F496C47B10" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC7F496C47AB7" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC7F496C47AB7" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC7F4939D7AFD" bold="true" box="[151,197,151,173]" pageId="6" pageNumber="207">4 (2)</emphasis>
Outer surface of mandible with notorious tubercle at base of condylar ridge, and a low Y-like tubercle at base of outer ridge; interior margin of mandible with tooth-like elevation; apical margin of clypeus with two medium-sized to prominent tubercles................................................................................
<taxonomicName id="97912A80FFA2B055FB3CC7B196C47AB7" ID-CoL="3KYQL" authorityName="Michener" authorityYear="1943" box="[1270,1436,210,231]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="currani">
<emphasis id="62E58D11FFA2B055FB3CC7B196C47AB7" box="[1270,1436,210,231]" italics="true" pageId="6" pageNumber="207">Heriades currani</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC79396C47B10" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC79396C47B10" pageId="6" pageNumber="207">- Outer surface of mandible with one notorious tubercle at base condylar ridge and outer ridge; interior margin of mandible with carina (without tooth-like elevation); apical margin of clypeus with five or more small tubercles, sometimes as irregular crenulations......................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC62496C57B2A" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC62496C57B0D" blockId="6.[151,1437,151,1522]" box="[151,1437,327,349]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC62496C57B0D" box="[151,1437,327,349]" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC624939D7B0D" bold="true" box="[151,197,327,349]" pageId="6" pageNumber="207">5 (4)</emphasis>
Apical margin of clypeus with emargination; hairs on posterior terga yellowish to orange (
<figureCitation id="C8AA4D86FFA2B055FB8EC62497C37B0C" box="[1092,1179,327,349]" captionStart="FIGURE 33" captionStartId="55.[151,250,1943,1967]" captionTargetBox="[155,1419,189,1905]" captionTargetId="figure-18@55.[155,1432,179,1917]" captionTargetPageId="55" captionText="FIGURE 33. Heriades vulcanica sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734982" httpUri="https://zenodo.org/record/14734982/files/figure.png" pageId="6" pageNumber="207">Fig. 33D</figureCitation>
)...
<taxonomicName id="97912A80FFA2B055FB18C62B96147B0C" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1234,1356,327,349]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="vulcanica" status="sp. nov.">
<emphasis id="62E58D11FFA2B055FB18C62B96147B0C" box="[1234,1356,327,349]" italics="true" pageId="6" pageNumber="207">H. vulcanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA2B055FA98C62B96C57B0D" box="[1362,1437,328,349]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC60696C57B2A" blockId="6.[151,1437,151,1522]" box="[151,1437,356,378]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC60696C57B2A" box="[151,1437,356,378]" pageId="6" pageNumber="207">
- Apical margin of clypeus straight, without emargination; hairs on posterior terga white (
<figureCitation id="C8AA4D86FFA2B055FBD4C607972C7B2A" box="[1054,1140,356,378]" captionStart="FIGURE 23" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[172,1397,192,1905]" captionTargetId="figure-18@39.[172,1416,179,1917]" captionTargetPageId="39" captionText="FIGURE 23. Heriades mariana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734946" httpUri="https://zenodo.org/record/14734946/files/figure.png" pageId="6" pageNumber="207">Fig. 23D</figureCitation>
)...
<taxonomicName id="97912A80FFA2B055FB68C60796167B2A" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1186,1358,356,378]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="mariana" status="sp. nov.">
<emphasis id="62E58D11FFA2B055FB68C60796167B2A" box="[1186,1358,356,378]" italics="true" pageId="6" pageNumber="207">Heriades mariana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA2B055FA9EC60696C57B2A" box="[1364,1437,357,378]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC6E196C47B82" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC6E196C47BE5" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC6E196C47BE5" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC6E1939D7BC7" bold="true" box="[151,197,385,407]" pageId="6" pageNumber="207">6 (1)</emphasis>
Punctures on T3 small (0.330.40 DOP as on scutum [except on
<taxonomicName id="97912A80FFA2B055FC80C6E190CE7BC7" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[842,918,386,407]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="luisa" status="sp. nov.">
<emphasis id="62E58D11FFA2B055FC80C6E190CE7BC7" box="[842,918,386,407]" italics="true" pageId="6" pageNumber="207">H. luisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA2B055FC51C6E190B07BC7" box="[923,1000,386,407]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
: 0.60 DOP as on scutum, which punctures are small) (
<figureCitation id="C8AA4D86FFA2B055FEE1C6FC922E7BE4" box="[299,374,415,437]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3K</figureCitation>
)...................................................................................... 7
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC6DE96C47B82" blockId="6.[151,1437,151,1522]" box="[151,1436,444,466]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC6DE96C47B82" box="[151,1436,444,466]" pageId="6" pageNumber="207">
- Punctures on T3 medium-sized or large (~1 DOP as on scutum) (
<figureCitation id="C8AA4D86FFA2B055FC9BC6DF90CC7B82" box="[849,916,444,466]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3J</figureCitation>
)......................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC6B996C47817" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC6B996C4785C" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC6B996C4785C" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC6B9939D7BBF" bold="true" box="[151,197,473,495]" pageId="6" pageNumber="207">7 (6)</emphasis>
Apical margin of clypeus with two prominent or small tubercles on frontal view (
<figureCitation id="C8AA4D86FFA2B055FBD9C6B9973A7BBF" box="[1043,1122,474,496]" captionStart="FIGURE 9" captionStartId="18.[151,250,1944,1968]" captionTargetBox="[152,1422,179,1905]" captionTargetId="figure-22@18.[152,1435,179,1918]" captionTargetPageId="18" captionText="FIGURE 9. Heriades bruneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734902" httpUri="https://zenodo.org/record/14734902/files/figure.png" pageId="6" pageNumber="207">Fig. 9C</figureCitation>
); pubescence on posterior terga orangish (
<figureCitation id="C8AA4D86FFA2B055FE88C69492D4785C" box="[322,396,503,525]" captionStart="FIGURE 9" captionStartId="18.[151,250,1944,1968]" captionTargetBox="[152,1422,179,1905]" captionTargetId="figure-22@18.[152,1435,179,1918]" captionTargetPageId="18" captionText="FIGURE 9. Heriades bruneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734902" httpUri="https://zenodo.org/record/14734902/files/figure.png" pageId="6" pageNumber="207">Fig. 9D</figureCitation>
).......................................................................
<taxonomicName id="97912A80FFA2B055FB3CC69496C4785C" ID-CoL="3KYPX" authorityName="Titus" authorityYear="1904" box="[1270,1436,503,524]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="bruneri">
<emphasis id="62E58D11FFA2B055FB3CC69496C4785C" box="[1270,1436,503,524]" italics="true" pageId="6" pageNumber="207">Heriades bruneri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC57696C47817" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC57696C47817" pageId="6" pageNumber="207">
- Apical margin of clypeus with five or more small tubercles on frontal view (
<figureCitation id="C8AA4D86FFA2B055FC7CC5779759787A" box="[950,1025,532,554]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734890" httpUri="https://zenodo.org/record/14734890/files/figure.png" pageId="6" pageNumber="207">Fig. 5C</figureCitation>
); pubescence on posterior terga white (
<figureCitation id="C8AA4D86FFA2B055FABFC57792597818" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734890" httpUri="https://zenodo.org/record/14734890/files/figure.png" pageId="6" pageNumber="207">Fig. 5D</figureCitation>
,
<figureCitation id="C8AA4D86FFA2B055FEC1C551926E7817" box="[267,310,562,583]" captionStart="FIGURE 20" captionStartId="34.[151,250,1098,1122]" captionTargetBox="[159,1428,187,1064]" captionTargetId="figure-22@34.[151,1436,179,1072]" captionTargetPageId="34" captionText="FIGURE 20. Known distribution records for Heriades luisa sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14734936" httpUri="https://zenodo.org/record/14734936/files/figure.png" pageId="6" pageNumber="207">20D</figureCitation>
)............................................................................................ 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC52C96C578A7" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC52C96C578CF" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC52C96C578CF" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC52C939D7835" bold="true" box="[151,197,591,613]" pageId="6" pageNumber="207">8 (7)</emphasis>
Frons with hooked, bright, golden hairs; punctures on base of clypeus large (~0.33 times OD) (
<figureCitation id="C8AA4D86FFA2B055FBBDC52C979A7835" box="[1143,1218,591,613]" captionStart="FIGURE 5" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[163,1409,179,1905]" captionTargetId="figure-18@11.[163,1423,179,1918]" captionTargetPageId="11" captionText="FIGURE 5. Heriades amanda sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734890" httpUri="https://zenodo.org/record/14734890/files/figure.png" pageId="6" pageNumber="207">Fig. 5C</figureCitation>
); punctures on scutum medium-sized (0.38 times OD) (
<figureCitation id="C8AA4D86FFA2B055FDE8C50F913778D2" box="[546,623,620,642]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3E</figureCitation>
); metepisternum with few plumose, white, erected, thin hairs and disc with tuft of plumose, bright white, short hairs (
<figureCitation id="C8AA4D86FFA2B055FDF8C5E9912578CF" box="[562,637,650,672]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3D</figureCitation>
)............................................
<taxonomicName id="97912A80FFA2B055FB6BC5E9961478CF" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1185,1356,650,671]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="amanda" status="sp. nov.">
<emphasis id="62E58D11FFA2B055FB6BC5E9961478CF" box="[1185,1356,650,671]" italics="true" pageId="6" pageNumber="207">Heriades amanda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA2B055FA98C5E996C578CF" box="[1362,1437,650,671]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC5CB96C578A7" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC5CB96C578A7" pageId="6" pageNumber="207">
- Frons with simple, white hairs; punctures on base of clypeus small (~0.25 times OD) (
<figureCitation id="C8AA4D86FFA2B055FBE7C5C497DE78EC" box="[1069,1158,679,701]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="6" pageNumber="207">Fig. 21C</figureCitation>
); punctures on scutum small (~0.20 times OD) (
<figureCitation id="C8AA4D86FFA2B055FE53C5A792BD788A" box="[409,485,708,730]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3D</figureCitation>
); metepisternum with few plumose, white, erected, thin hairs without tuft (
<figureCitation id="C8AA4D86FFA2B055FB7FC5A79653788A" box="[1205,1291,708,730]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="6" pageNumber="207">Fig. 21D</figureCitation>
)...........................................................................................
<taxonomicName id="97912A80FFA2B055FB0AC581961578A7" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1216,1357,738,759]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="luisa" status="sp. nov.">
<emphasis id="62E58D11FFA2B055FB0AC581961578A7" box="[1216,1357,738,759]" italics="true" pageId="6" pageNumber="207">Heriades luisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA2B055FA98C58196C578A7" box="[1362,1437,738,759]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC59C96C4793C" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC59C96C47962" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC59C96C47962" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC59C939D7945" bold="true" box="[151,197,767,789]" pageId="6" pageNumber="207">9 (6)</emphasis>
Apical margin of clypeus with two prominent tubercles (
<figureCitation id="C8AA4D86FFA2B055FD36C59C90097944" box="[764,849,767,789]" captionStart="FIGURE 11" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[179,1395,190,1905]" captionTargetId="figure-18@21.[179,1408,179,1917]" captionTargetPageId="21" captionText="FIGURE 11. Heriades cristina sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734908" httpUri="https://zenodo.org/record/14734908/files/figure.png" pageId="6" pageNumber="207">Fig. 11C</figureCitation>
,
<figureCitation id="C8AA4D86FFA2B055FC91C59C90DE7944" box="[859,902,767,788]" captionStart="FIGURE 19" captionStartId="32.[151,250,1943,1967]" captionTargetBox="[163,1412,184,1905]" captionTargetId="figure-22@32.[163,1425,179,1917]" captionTargetPageId="32" captionText="FIGURE 19. Heriades leavitti. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734934" httpUri="https://zenodo.org/record/14734934/files/figure.png" pageId="6" pageNumber="207">19C</figureCitation>
); apical terga fasciae with rows of thin, white hairs (
<figureCitation id="C8AA4D86FFA2B055FABFC59C93A57962" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3L</figureCitation>
)............................................................................................... 10
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC45996C4793C" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC45996C4793C" pageId="6" pageNumber="207">
- Apical margin of clypeus with five or more small tubercles (
<figureCitation id="C8AA4D86FFA2B055FCE8C4599021791F" box="[802,889,826,848]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734926" httpUri="https://zenodo.org/record/14734926/files/figure.png" pageId="6" pageNumber="207">Fig. 17C</figureCitation>
); apical terga fasciae with a tidy row of thick, white hairs (
<figureCitation id="C8AA4D86FFA2B055FF23C4349274793C" box="[233,300,855,877]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3J</figureCitation>
)........................................................................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC41796C579B2" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC41796C579F7" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC41796C579F7" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC417938979DA" bold="true" box="[151,209,884,906]" pageId="6" pageNumber="207">10 (9)</emphasis>
Frons in front of ocelli with small punctures (
<quantity id="9769FCE6FFA2B055FD57C41791AE79DA" box="[669,758,884,906]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="207" unit="mm" value="0.05">0.05 mm</quantity>
) and on T3 medium-sized (0.19 times OD); surface of T3 dull; wings usually dusky............................................................................
<taxonomicName id="97912A80FFA2B055FB36C4F196C579F7" ID-CoL="3KYRS" authorityName="Crawford" authorityYear="1913" box="[1276,1437,914,935]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="leavitti">
<emphasis id="62E58D11FFA2B055FB36C4F196C579F7" box="[1276,1437,914,935]" italics="true" pageId="6" pageNumber="207">Heriades leavitti</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC4D396C579B2" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC4D396C579B2" pageId="6" pageNumber="207">
- Frons in front of ocelli and T3 with large punctures (
<quantity id="9769FCE6FFA2B055FD10C4CC906C7995" box="[730,820,943,965]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="207" unit="mm" value="0.06">0.06 mm</quantity>
, 0.30 times OD); surface of T3 bright; wings clear.........................................................................................
<taxonomicName id="97912A80FFA2B055FB6FC4AF961479B2" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1189,1356,972,994]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="cristina" status="sp. nov.">
<emphasis id="62E58D11FFA2B055FB6FC4AF961479B2" box="[1189,1356,972,994]" italics="true" pageId="6" pageNumber="207">Heriades cristina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA2B055FA98C4AE96C579B2" box="[1362,1437,973,994]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC48996C47E07" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC48996C57E4D" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC48996C57E4D" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC489939779AF" bold="true" box="[151,207,1001,1023]" pageId="6" pageNumber="207">11 (9)</emphasis>
Base of propodeum with large pits (~
<quantity id="9769FCE6FFA2B055FD9CC48991EA79AF" box="[598,690,1002,1023]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="6" pageNumber="207" unit="mm" value="0.07">0.07 mm</quantity>
) (
<figureCitation id="C8AA4D86FFA2B055FD02C489905679AF" box="[712,782,1002,1024]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3I</figureCitation>
); punctures on posterior surface of propodeum abundant reaching superior edge or carina...............................................................
<taxonomicName id="97912A80FFA2B055FB0BC36496147E4D" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1217,1356,1031,1053]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="nora" status="sp. nov.">
<emphasis id="62E58D11FFA2B055FB0BC36496147E4D" box="[1217,1356,1031,1053]" italics="true" pageId="6" pageNumber="207">Heriades nora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA2B055FA98C36B96C57E4D" box="[1362,1437,1032,1053]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC34696C47E07" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC34696C47E07" pageId="6" pageNumber="207">
- Base of propodeum with small to middle pits (~
<quantity id="9769FCE6FFA2B055FD62C34690597E6A" box="[680,769,1061,1082]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="207" unit="mm" value="0.04">0.04 mm</quantity>
) (
<figureCitation id="C8AA4D86FFA2B055FCDEC34790267E6A" box="[788,894,1060,1082]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3G, H</figureCitation>
); punctures on posterior surface of propodeum scarce on superior edge or carina............................................................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC33C96C47F57" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC33C96C47EFF" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC33C96C47EFF" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC33C93847E25" bold="true" box="[151,220,1119,1141]" pageId="6" pageNumber="207">12 (11)</emphasis>
Punctures on frons in front of ocelli and vertex large (0.70.9 times OD); middle ocelli smaller than laterals; dorsolateral angles of pronotum with rounded tuft of plumose, orangish to white-yellowish, thick hairs (
<figureCitation id="C8AA4D86FFA2B055FBEBC31F97227EC2" box="[1057,1146,1148,1170]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734952" httpUri="https://zenodo.org/record/14734952/files/figure.png" pageId="6" pageNumber="207">Fig. 25D</figureCitation>
,
<figureCitation id="C8AA4D86FFA2B055FB4CC31E97EB7EC2" box="[1158,1203,1149,1170]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734926" httpUri="https://zenodo.org/record/14734926/files/figure.png" pageId="6" pageNumber="207">17D</figureCitation>
); punctures on T3 large (~0.5 times OD)..................................................................................... 13
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC3DB96C47F57" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC3DB96C47F57" pageId="6" pageNumber="207">
- Punctures on frons in front of ocelli and vertex medium-sized (0.20.4 times OD); middle ocelli as large as laterals; dorsolateral angles of pronotum with few plumose, white, thin hairs joining hairs on pronotum but not forming a rounded tuft (
<figureCitation id="C8AA4D86FFA2B055FA80C3B796C07EBA" box="[1354,1432,1236,1258]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[163,1410,183,1910]" captionTargetId="figure-18@15.[163,1425,179,1917]" captionTargetPageId="15" captionText="FIGURE 7. Heriades briceida sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734896" httpUri="https://zenodo.org/record/14734896/files/figure.png" pageId="6" pageNumber="207">Fig. 7D</figureCitation>
,
<figureCitation id="C8AA4D86FFA2B055FF28C39192577F57" box="[226,271,1266,1287]" captionStart="FIGURE 31" captionStartId="51.[151,250,1943,1967]" captionTargetBox="[182,1392,192,1905]" captionTargetId="figure-18@51.[182,1405,179,1917]" captionTargetPageId="51" captionText="FIGURE 31. Heriades viridiana sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734974" httpUri="https://zenodo.org/record/14734974/files/figure.png" pageId="6" pageNumber="207">31D</figureCitation>
); punctures on T3 medium-sized (~0.4 times OD)...................................................... 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC26C96C57F2D" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC26C96C47F11" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC26C96C47F11" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC26C93857F75" bold="true" box="[151,221,1295,1317]" pageId="6" pageNumber="207">13 (12)</emphasis>
Scape thin, 3.1 times longer than wide (
<figureCitation id="C8AA4D86FFA2B055FDA5C26C91917F75" box="[623,713,1295,1317]" captionStart="FIGURE 25" captionStartId="42.[151,250,1943,1967]" captionTargetBox="[160,1414,187,1905]" captionTargetId="figure-22@42.[160,1427,179,1917]" captionTargetPageId="42" captionText="FIGURE 25. Heriades micheneri. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734952" httpUri="https://zenodo.org/record/14734952/files/figure.png" pageId="6" pageNumber="207">Fig. 25C</figureCitation>
); punctures on mesepisternum medium-sized (0.4 times OD) and slightly sparse (0.3 times DOP).................................................................
<taxonomicName id="97912A80FFA2B055FB14C24F96C47F11" ID-CoL="3KYS3" box="[1246,1436,1324,1345]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="micheneri">
<emphasis id="62E58D11FFA2B055FB14C24F96C47F11" box="[1246,1436,1324,1345]" italics="true" pageId="6" pageNumber="207">Heriades micheneri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC22996C57F2D" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC22996C57F2D" pageId="6" pageNumber="207">
- Scape thick, 2.7 times longer than wide (
<figureCitation id="C8AA4D86FFA2B055FDBFC22991967F0F" box="[629,718,1354,1376]" captionStart="FIGURE 17" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[169,1398,185,1905]" captionTargetId="figure-18@29.[168,1420,179,1917]" captionTargetPageId="29" captionText="FIGURE 17. Heriades danuncia sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734926" httpUri="https://zenodo.org/record/14734926/files/figure.png" pageId="6" pageNumber="207">Fig. 17C</figureCitation>
); punctures on mesepisternum large (0.6 times OD) and dense (0.2 times DOP).........................................................................
<taxonomicName id="97912A80FFA2B055FB5FC20496147F2C" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1173,1356,1383,1404]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="danuncia" status="sp. nov.">
<emphasis id="62E58D11FFA2B055FB5FC20496147F2C" box="[1173,1356,1383,1404]" italics="true" pageId="6" pageNumber="207">Heriades danuncia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA2B055FA98C20B96C57F2D" box="[1362,1437,1384,1405]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="EB6549A6FFA2B055FF5DC2E796C57FA2" pageId="6" pageNumber="207">
<paragraph id="502E5103FFA2B055FF5DC2E796C57FE7" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC2E796C57FE7" pageId="6" pageNumber="207">
<emphasis id="62E58D11FFA2B055FF5DC2E793857FCA" bold="true" box="[151,221,1412,1434]" pageId="6" pageNumber="207">14 (12)</emphasis>
Apical margin of clypeus with tubercles in the middle; ocelli medium-sized (
<figureCitation id="C8AA4D86FFA2B055FC0AC2E797537FCA" box="[960,1035,1412,1434]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3B</figureCitation>
); punctures on scutum medium-sized (0.4 times OD) (
<figureCitation id="C8AA4D86FFA2B055FE9EC2C192C37FE7" box="[340,411,1442,1464]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3E</figureCitation>
)..............................................................
<taxonomicName id="97912A80FFA2B055FB54C2C196147FE7" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1182,1356,1442,1463]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="briceida" status="sp. nov.">
<emphasis id="62E58D11FFA2B055FB54C2C196147FE7" box="[1182,1356,1442,1463]" italics="true" pageId="6" pageNumber="207">Heriades briceida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA2B055FA98C2C196C57FE7" box="[1362,1437,1442,1463]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="502E5103FFA2B055FF5DC2A396C57FA2" blockId="6.[151,1437,151,1522]" pageId="6" pageNumber="207">
<keyLead id="EB60F236FFA2B055FF5DC2A396C57FA2" pageId="6" pageNumber="207">
- Apical margin of clypeus without tubercles in the middle; ocelli large (
<figureCitation id="C8AA4D86FFA2B055FC4AC2DC90937F84" box="[896,971,1471,1493]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3C</figureCitation>
); punctures on scutum large (0.5 times OD) (
<figureCitation id="C8AA4D86FFA2B055FABFC2DC93A47FA2" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.AC ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). DF scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). GI propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). JL Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="6" pageNumber="207">Fig. 3F</figureCitation>
)...........................................................................
<taxonomicName id="97912A80FFA2B055FB5CC2BF96147FA1" authorityName="Rojas-Arias &amp; Griswold &amp; Hinojosa-Díaz &amp; Morrone &amp; Barajas" authorityYear="2025" box="[1174,1356,1500,1521]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="viridiana" status="sp. nov.">
<emphasis id="62E58D11FFA2B055FB5CC2BF96147FA1" box="[1174,1356,1500,1521]" italics="true" pageId="6" pageNumber="207">Heriades viridiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79D6306AFFA2B055FA98C2BE96C57FA2" box="[1362,1437,1501,1522]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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