<documentid="8D9871389A7B689DDD17BBF1AF571F5A"ID-CLB-Dataset="304902"ID-DOI="10.5852/ejt.2024.966.2697"ID-GBIF-Dataset="0bf71223-cd52-42bc-8bce-edad65412211"ID-ISSN="2118-9773"ID-Zenodo-Dep="14001790"ID-ZooBank="FFE41347-8B55-4CF1-B10B-53EBB4CF0E26"IM.bibliography_approvedBy="carolina"IM.illustrations_approvedBy="carolina"IM.materialsCitations_approvedBy="carolina"IM.metadata_approvedBy="carolina"IM.tables_approvedBy="carolina"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="carolina"IM.treatments_approvedBy="carolina"checkinTime="1730118416961"checkinUser="plazi"docAuthor="Henrard, Arnaud, Griswold, Charles & Jocqué, Rudy"docDate="2024"docId="3157E00AFFF6FFDDFDF10CBEFD20D971"docLanguage="en"docName="EJT.2024.966.1-80.pdf"docOrigin="European Journal of Taxonomy 966 (1)"docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Tabiboka milleri Henrard, Griswold & Jocqué, 2024, gen. et sp. nov."docType="treatment"docUuid="33C8C4F1-A241-41E2-9B38-1D31AFBF4BE4"docUuidSource="ZooBank"docVersion="8"lastPageNumber="24"masterDocId="CD6E9872FFF8FFCAFFAA0824FFD5DF0E"masterDocTitle="On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar"masterLastPageNumber="80"masterPageNumber="1"pageNumber="15"updateTime="1730317898278"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:affiliationid="22E89C955DEDD136DC2F4FC0433AA43C">Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium.</mods:affiliation>
<mods:affiliationid="0150E50687F4C74AE15DC45B5340ED75">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA.</mods:affiliation>
<mods:affiliationid="DB7EA1984416B5877F39D38AF8731FDE">Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium.</mods:affiliation>
<taxonomicNameid="7EFE2A9FFFF6FFC4FDF10CBEFCF7DBBB"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[603,802,1178,1205]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="milleri"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFF6FFC4FC830CBFFC0CDBBB"box="[809,985,1179,1205]"pageId="14"pageNumber="15"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<figureCitationid="21C54D99FFF6FFC4FCEF0CC1FCB7DBF1"box="[837,866,1253,1279]"captionStart="Fig"captionStartId="63.[189,231,1857,1883]"captionTargetBox="[189,1398,1005,1819]"captionTargetId="figure-208@63.[189,1398,1005,1819]"captionTargetPageId="63"captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)."figureDoi="http://doi.org/10.5281/zenodo.14001851"httpUri="https://zenodo.org/record/14001851/files/figure.png"pageId="14"pageNumber="15">26</figureCitation>
<bibRefCitationid="DD6F2CEDFFF6FFC4FD780D0DFC74DA4D"author="Morphbank"box="[722,929,1321,1347]"pageId="14"pageNumber="15"refId="ref37471"refString="Morphbank. 2023. Biological Imaging. Florida State University, Department of Scientific Computing, Tallahassee, FL 32306 - 4026 USA. Available from https: // www. morphbank. net / [accessed 20 Oct. 2023]."type="url"year="2023">Morphbank 2023</bibRefCitation>
<treatmentCitationid="385F770DFFF6FFC4FE0D0D57FD45DA80"author="Polotow D. & Carmichael A. & Griswold C. E."box="[423,656,1395,1422]"pageId="14"pageNumber="15"year="2015">
<bibRefCitationid="DD6F2CEDFFF6FFC4FE0D0D57FD45DA80"author="Polotow D. & Carmichael A. & Griswold C. E."box="[423,656,1395,1422]"pageId="14"pageNumber="15"pagination="124 - 163"refId="ref37852"refString="Polotow D., Carmichael A. & Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041"type="journal article"year="2015">
<treatmentCitationid="385F770DFFF6FFC4FE470DBCFD0DDABC"author="Wheeler W. C. & Coddington J. A. & Crowley L. M. & Dimitrov D. & Goloboff P. A. & Griswold C. E. & Hormiga G. & Prendini L. & Ramirez M. J. & Sierwald P. & Almeida-Silva L. M. & Alvarez-Padilla F. & Arnedo M. A. & Benavides L. R. & Benjamin S. P. & Bond J. E. & Grismado C. J. & Hasan E. & Hedin M. & Izquierdo M. A. & Labarque F. M. & Ledford J. & Lopardo L. & Maddison W. P. & Miller J. A. & Piacentini L. N. & Platnick N. I. & Polotow D. & Silva-Davila D. & Scharff N. & Szuts T. & Ubick D. & Vink C. & Wood H. M. & Zhang J. X."box="[493,728,1432,1458]"pageId="14"pageNumber="15"year="2017">
<bibRefCitationid="DD6F2CEDFFF6FFC4FE470DBCFD0DDABC"author="Wheeler W. C. & Coddington J. A. & Crowley L. M. & Dimitrov D. & Goloboff P. A. & Griswold C. E. & Hormiga G. & Prendini L. & Ramirez M. J. & Sierwald P. & Almeida-Silva L. M. & Alvarez-Padilla F. & Arnedo M. A. & Benavides L. R. & Benjamin S. P. & Bond J. E. & Grismado C. J. & Hasan E. & Hedin M. & Izquierdo M. A. & Labarque F. M. & Ledford J. & Lopardo L. & Maddison W. P. & Miller J. A. & Piacentini L. N. & Platnick N. I. & Polotow D. & Silva-Davila D. & Scharff N. & Szuts T. & Ubick D. & Vink C. & Wood H. M. & Zhang J. X."box="[493,728,1432,1458]"pageId="14"pageNumber="15"pagination="576 - 616"refId="ref39136"refString="Wheeler W. C., Coddington J. A., Crowley L. M., Dimitrov D., Goloboff P. A., Griswold C. E., Hormiga G., Prendini L., Ramirez M. J., Sierwald P., Almeida-Silva L. M., Alvarez-Padilla F., Arnedo M. A., Benavides L. R., Benjamin S. P., Bond J. E., Grismado C. J., Hasan E., Hedin M., Izquierdo M. A., Labarque F. M., Ledford J., Lopardo L., Maddison W. P., Miller J. A., Piacentini L. N., Platnick N. I., Polotow D., Silva-Davila D., Scharff N., Szuts T., Ubick D., Vink C., Wood H. M. & Zhang J. X. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. https: // doi. org / 10.1111 / cla. 12182"type="journal article"year="2017">
<taxonomicNameid="7EFE2A9FFFF6FFC4FE9A0E28FE27D928"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[304,498,1548,1574]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="milleri"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFF6FFC4FE500E29FD7BD929"box="[506,686,1549,1575]"pageId="14"pageNumber="15"rank="species">gen. et sp. nov.</taxonomicNameLabel>
may be distinguished by the morphology of the palpal tibia with the RTA having an elongate curved spur arising retrobasally and extending distally to the apex of the tibia (
<figureCitationid="21C54D99FFF6FFC4FE860E72FE7AD97F"box="[300,431,1622,1649]"captionStart="Fig"captionStartId="18.[189,233,1787,1813]"captionTargetBox="[219,1365,478,1733]"captionTargetId="figure-82@18.[189,1398,445,1749]"captionTargetPageId="18"captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001800"httpUri="https://zenodo.org/record/14001800/files/figure.png"pageId="14"pageNumber="15">Figs 5B–C</figureCitation>
,
<figureCitationid="21C54D99FFF6FFC4FE110E72FDD7D97E"box="[443,514,1622,1648]"captionStart="Fig"captionStartId="19.[188,230,1787,1813]"captionTargetBox="[188,1399,479,1749]"captionTargetId="figure-89@19.[188,1399,479,1749]"captionTargetPageId="19"captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–C = 0.43 mm; D–F = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick."figureDoi="http://doi.org/10.5281/zenodo.14001804"httpUri="https://zenodo.org/record/14001804/files/figure.png"pageId="14"pageNumber="15">6A–C</figureCitation>
) (vs
<taxonomicNameid="7EFE2A9FFFF6FFC4FD960E73FD70D97E"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[572,677,1622,1648]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="milloti"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFF6FFC4FD060E73FC89D97F"box="[684,860,1623,1649]"pageId="14"pageNumber="15"rank="species">gen. et sp. nov.</taxonomicNameLabel>
and
<taxonomicNameid="7EFE2A9FFFF6FFC4FC3E0E73FBFED97E"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[916,1067,1622,1648]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="polotowae"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFF6FFC4FB980E73FB37D97F"box="[1074,1250,1623,1649]"pageId="14"pageNumber="15"rank="species">gen. et sp. nov.</taxonomicNameLabel>
have a short and stout RTA,
<figureCitationid="21C54D99FFF6FFC4FED90E5FFE26D998"box="[371,499,1659,1686]"captionStart="Fig"captionStartId="27.[189,231,1595,1621]"captionTargetBox="[260,1379,284,1517]"captionTargetId="figure-11@27.[189,1398,265,1558]"captionTargetPageId="27"captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.A–C = male holotype (CASENT_9016063; MNHN). D–E = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold."figureDoi="http://doi.org/10.5281/zenodo.14001810"httpUri="https://zenodo.org/record/14001810/files/figure.png"pageId="14"pageNumber="15">Figs 9B–C</figureCitation>
,
<figureCitationid="21C54D99FFF6FFC4FDAA0E58FD80D998"box="[512,597,1660,1686]"captionStart="Fig"captionStartId="31.[189,231,1719,1745]"captionTargetBox="[211,1371,542,1665]"captionTargetId="figure-76@31.[189,1398,527,1680]"captionTargetPageId="31"captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001819"httpUri="https://zenodo.org/record/14001819/files/figure.png"pageId="14"pageNumber="15">11B–C</figureCitation>
), a short, broad embolus (E) and a small, slender paraconductor (Pc) largely hidden behind the embolus (
<figureCitationid="21C54D99FFF6FFC4FDF60E85FD0AD9B5"box="[604,735,1696,1723]"captionStart="Fig"captionStartId="18.[189,233,1787,1813]"captionTargetBox="[219,1365,478,1733]"captionTargetId="figure-82@18.[189,1398,445,1749]"captionTargetPageId="18"captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001800"httpUri="https://zenodo.org/record/14001800/files/figure.png"pageId="14"pageNumber="15">Figs 5A–C</figureCitation>
,
<figureCitationid="21C54D99FFF6FFC4FD430E84FCFBD9B4"box="[745,814,1696,1722]"captionStart="Fig"captionStartId="19.[188,230,1787,1813]"captionTargetBox="[188,1399,479,1749]"captionTargetId="figure-89@19.[188,1399,479,1749]"captionTargetPageId="19"captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–C = 0.43 mm; D–F = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick."figureDoi="http://doi.org/10.5281/zenodo.14001804"httpUri="https://zenodo.org/record/14001804/files/figure.png"pageId="14"pageNumber="15">6D–F</figureCitation>
) (vs in
<taxonomicNameid="7EFE2A9FFFF6FFC4FC280E85FC3ED9B5"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[898,1003,1697,1723]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="milloti">
<taxonomicNameid="7EFE2A9FFFF6FFC4FB8B0E85FB62D9B5"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[1057,1207,1697,1723]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="polotowae">
<figureCitationid="21C54D99FFF6FFC4FD580EE2FC99D9EE"box="[754,844,1734,1760]"captionStart="Fig"captionStartId="25.[189,232,1717,1743]"captionTargetId="figure-19@26.[189,1398,304,1979]"captionTargetPageId="26"captionText="Fig. 8(on next page). Tabiboka milloti gen. nov. et sp. nov. male holotype (CASENT_9016063; MNHN). A. Habitus, dorsal view. B. Carapace, dorsal view. C. SEM of right male palp (mirrored), prolateral view. D. Idem, ventral view. E. Idem, retrolateral view. F. Idem, detail of bulb, ventral view. G. Idem, detail of the RTA. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; MA = median apophysis; Pc = paraconductor; dRTA = retrodorsal tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars: A–B = no scales provided (note: carapace length = 4.6 mm); C–E = 0.5 mm; F–G = 0.1 mm. Stacked microphotographs by Hannah Wood; SEM images by Charles Griswold and Darrell Ubick."figureDoi="http://doi.org/10.5281/zenodo.14014722"httpUri="https://zenodo.org/record/14014722/files/figure.png"pageId="14"pageNumber="15">Figs 8F</figureCitation>
<taxonomicNameid="7EFE2A9FFFF6FFC4FB0A0EE3FADED9EE"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[1184,1291,1734,1760]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="milleri">
have the epigyne with a broad, swollen, oval median lobe (ML) that extends posteriorly beyond the epigastric furrow, narrow anteriorly (anterior width less than half the total width,
<figureCitationid="21C54D99FFF6FFC4FBF60F34FB57D825"box="[1116,1154,1808,1835]"captionStart="Fig"captionStartId="18.[189,233,1787,1813]"captionTargetBox="[219,1365,478,1733]"captionTargetId="figure-82@18.[189,1398,445,1749]"captionTargetPageId="18"captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001800"httpUri="https://zenodo.org/record/14001800/files/figure.png"pageId="14"pageNumber="15">5D</figureCitation>
,
<figureCitationid="21C54D99FFF6FFC4FB270F34FB67D824"box="[1165,1202,1808,1834]"captionStart="Fig"captionStartId="20.[189,232,1822,1848]"captionTargetBox="[189,1398,461,1784]"captionTargetId="figure-79@20.[189,1398,461,1784]"captionTargetPageId="20"captionText="Fig. 7. Tabiboka milleri gen. nov. et sp. nov. SEM of female genitalia, A–B, E–F = female paratype (CASENT_9016094; CAS); C–D = female paratype (CASENT_9035866; CAS). A. Epigyne, ventral view. B. Idem, ventro-posterior view. C. Endogyne, dorsal view. D. Idem, slightly anterior view. E. Spermatheca, right dorsal view. F. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: A–E = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila."figureDoi="http://doi.org/10.5281/zenodo.14001806"httpUri="https://zenodo.org/record/14001806/files/figure.png"pageId="14"pageNumber="15">7A</figureCitation>
) (vs in
<taxonomicNameid="7EFE2A9FFFF6FFC4FAAC0F35FABBD824"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[1286,1390,1808,1834]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="milloti">
<figureCitationid="21C54D99FFF6FFC4FE560F11FD8CD841"box="[508,601,1845,1871]"captionStart="Fig"captionStartId="27.[189,231,1595,1621]"captionTargetBox="[260,1379,284,1517]"captionTargetId="figure-11@27.[189,1398,265,1558]"captionTargetPageId="27"captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.A–C = male holotype (CASENT_9016063; MNHN). D–E = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold."figureDoi="http://doi.org/10.5281/zenodo.14001810"httpUri="https://zenodo.org/record/14001810/files/figure.png"pageId="14"pageNumber="15">Fig. 9D</figureCitation>
; and in
<taxonomicNameid="7EFE2A9FFFF6FFC4FD190F12FC9CD841"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[691,841,1845,1871]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="14"pageNumber="15"phylum="Arthropoda"rank="species"species="polotowae">
<figureCitationid="21C54D99FFF6FFC4FB100F11FAF2D841"box="[1210,1319,1845,1871]"captionStart="Fig"captionStartId="31.[189,231,1719,1745]"captionTargetBox="[211,1371,542,1665]"captionTargetId="figure-76@31.[189,1398,527,1680]"captionTargetPageId="31"captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001819"httpUri="https://zenodo.org/record/14001819/files/figure.png"pageId="14"pageNumber="15">Figs 11D</figureCitation>
,
<figureCitationid="21C54D99FFF6FFC4FA990F11FABCD841"box="[1331,1385,1845,1871]"captionStart="Fig"captionStartId="32.[189,232,1717,1743]"captionTargetBox="[277,1311,265,1679]"captionTargetId="figure-18@32.[277,1311,265,1679]"captionTargetPageId="32"captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; A–F = male holotype (CASENT_9016117; CAS); G–H= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–D = 0.5 mm; E–F = 0.15 mm; G–H = 0.1 mm. SEM images A–F by Charles Griswold and Darrell Ubick, G–H by Daniele Polotow."figureDoi="http://doi.org/10.5281/zenodo.14001821"httpUri="https://zenodo.org/record/14001821/files/figure.png"pageId="14"pageNumber="15">12G</figureCitation>
and led a team that carried out a pioneering study of entelegyne spider phylogeny (
<bibRefCitationid="DD6F2CEDFFF6FFC4FB800FEAFB2AD8E6"author="Miller M. A. & Pfeiffer W. & Schwartz T."box="[1066,1279,1998,2024]"pageId="14"pageNumber="15"pagination="1 - 8"refId="ref37372"refString="Miller M. A., Pfeiffer W. & Schwartz T. 2010. Creating the CIPRES Science Gateway for inference of large phylogenetic trees. In: 2010 Gateway Computing Environments Workshop (GCE): 1 - 8. https: // doi. org / 10.1109 / GCE. 2010.5676129"type="proceedings paper"year="2010">
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; same collection data as for holotype; [Standard view preparations and images for AToL-Spiders project; SEM preparations Box 79-8, 10, 11, 19, Box 81, 1, 2]; CASENT_9016094;
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<materialsCitationid="09965B41FFECFFDEFD9F092DFAA3DE48"collectedFrom="general collecting from foliage"collectingDate="1992-04"collectionCode="CAS"collectorName="V. and B. Roth & S. Kariko"country="Madagascar"latitude="-21.2"location="Ranomafana"longLatPrecision="1264"longitude="47.45"pageId="20"pageNumber="21"specimenCode="CASENT_9016099"specimenCount="2"specimenCount-female="2"stateProvince="Fianarantsoa">
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<materialsCitationid="09965B41FFECFFDFFA840957FC11DE48"collectingDate="1992-05-20"collectingDateMax="1992-05-31"collectingDateMin="1992-05-20"collectingMethod="can traps"collectionCode="MCZ"collectorName="V. and B. Roth"country="Madagascar"lastPageId="21"lastPageNumber="22"latitude="-21.2"location="Ranomafana"longLatPrecision="1264"longitude="47.45"pageId="20"pageNumber="21"specimenCode="CASENT_9016101"specimenCount="11"specimenCount-female="7"specimenCount-male="4"stateProvince="Fianarantsoa">
<collectingDateid="DD048E34FFEDFFDFFB82092DFB2CDE2D"box="[1064,1273,265,291]"pageId="21"pageNumber="22"value="1992-05-20"valueMax="1992-05-31"valueMin="1992-05-20">20–31 May 1992</collectingDate>
</date>
;
<collectorNameid="140B34CAFFEDFFDFFAAC092DFF22DE48"pageId="21"pageNumber="22">V. and B. Roth</collectorName>
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<materialsCitationid="09965B41FFEDFFDFFC480908FE24DE64"collectingDate="1992-05-20"collectingDateMax="1992-05-31"collectingDateMin="1992-05-20"collectionCode="CAS"collectorName="V. and B. Roth"country="Madagascar"latitude="-21.2"location="Ranomafana"longLatPrecision="1264"longitude="47.45"pageId="21"pageNumber="22"specimenCode="CASENT_9016102"specimenCount="1"specimenCount-male="1"stateProvince="Fianarantsoa">
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<taxonomicNameid="7EFE2A9FFFE8FFDAFEBD0F41FE35D871"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[279,480,1893,1919]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="16"pageNumber="17"phylum="Arthropoda"rank="species"species="milleri"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFE8FFDAFE430F41FD73D871"box="[489,678,1893,1919]"pageId="16"pageNumber="17"rank="species">gen. et sp. nov.</taxonomicNameLabel>
, A–B, G = male paratype (CASENT_9016097; CAS); C = male paratype (CASENT_9016098; CAS); D–F, H = female paratype (CASENT_9016094; CAS).
<taxonomicNameid="7EFE2A9FFFE9FFDBFEA70F41FE1FD871"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[269,458,1893,1919]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="17"pageNumber="18"phylum="Arthropoda"rank="species"species="milleri"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFE9FFDBFE7A0F41FDACD871"box="[464,633,1893,1919]"pageId="17"pageNumber="18"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<taxonomicNameid="7EFE2A9FFFEAFFD8FEB50EDFFE38D81B"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[287,493,1787,1813]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="18"pageNumber="19"phylum="Arthropoda"rank="species"species="milleri"status="gen. nov. et sp. nov.">
<taxonomicNameLabelid="90B93075FFEAFFD8FE500ED8FD2AD818"box="[506,767,1788,1814]"pageId="18"pageNumber="19"rank="species">gen. nov. et sp. nov.</taxonomicNameLabel>
genitalia drawings of male and female paratypes (CASENT_9016086; CAS).
<taxonomicNameid="7EFE2A9FFFEBFFD9FEA20EDFFE15D81B"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[264,448,1787,1813]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="19"pageNumber="20"phylum="Arthropoda"rank="species"species="milleri"status="gen. nov. et sp. nov.">
. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–C = 0.43 mm; D–F = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick.
<taxonomicNameid="7EFE2A9FFFECFFDEFEBC0F3AFE08D836"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[278,477,1822,1848]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="20"pageNumber="21"phylum="Arthropoda"rank="species"species="milleri"status="gen. nov. et sp. nov.">
<taxonomicNameLabelid="90B93075FFECFFDEFE4C0F3BFD0CD837"box="[486,729,1823,1849]"pageId="20"pageNumber="21"rank="species">gen. nov. et sp. nov.</taxonomicNameLabel>
SEM of female genitalia, A–B, E–F = female paratype (CASENT_9016094; CAS); C–D = female paratype (CASENT_9035866; CAS).
<figureCitationid="21C54D99FFEDFFDFFEF50A01FE29DD31"box="[351,508,549,575]"captionStart="Fig"captionStartId="16.[189,233,1892,1918]"captionTargetBox="[189,1398,467,1854]"captionTargetId="figure-79@16.[189,1398,467,1854]"captionTargetPageId="16"captionText="Fig. 3. Tabiboka milleri gen. et sp. nov., A–B, G = male paratype (CASENT_9016097; CAS); C = male paratype (CASENT_9016098; CAS); D–F, H = female paratype (CASENT_9016094; CAS). A, D. Habitus, dorsal view. B, E. Idem, ventral view. C, F. Carapace, dorsal view. G–H. Habitus, lateral view. Scale bars = 2 mm. Stacked microphotographs by Hannah Wood."figureDoi="http://doi.org/10.5281/zenodo.14001794"httpUri="https://zenodo.org/record/14001794/files/figure.png"pageId="21"pageNumber="22">Fig. 3A–C, G</figureCitation>
). Carapace brownish yellow, darker along margin, with two faint, longitudinal dark bands along margin of pars cephalica; black on ocular area and surrounding all eyes; chelicerae brownish yellow; sternum yellow, unmarked; legs brownish yellow, unmarked; abdomen brownish yellow with darker cardiac mark and dorsolateral dark bands that break up into chevrons posteriorly, sides brownish yellow mottled with dark marks, venter yellow except darker around spinneret base.
<figureCitationid="21C54D99FFEDFFDFFE970AD0FE4DDC00"box="[317,408,756,782]"captionStart="Fig"captionStartId="16.[189,233,1892,1918]"captionTargetBox="[189,1398,467,1854]"captionTargetId="figure-79@16.[189,1398,467,1854]"captionTargetPageId="16"captionText="Fig. 3. Tabiboka milleri gen. et sp. nov., A–B, G = male paratype (CASENT_9016097; CAS); C = male paratype (CASENT_9016098; CAS); D–F, H = female paratype (CASENT_9016094; CAS). A, D. Habitus, dorsal view. B, E. Idem, ventral view. C, F. Carapace, dorsal view. G–H. Habitus, lateral view. Scale bars = 2 mm. Stacked microphotographs by Hannah Wood."figureDoi="http://doi.org/10.5281/zenodo.14001794"httpUri="https://zenodo.org/record/14001794/files/figure.png"pageId="21"pageNumber="22">Fig. 3B</figureCitation>
). Oval with slightly sinuous margins; 2.30 wide, 2.60 long.
<paragraphid="B941511CFFEDFFDFFF170B53FCCADCBB"blockId="21.[188,1398,887,949]"pageId="21"pageNumber="22">EYES. ALE 0.18; AME 0.13; PLE 0.18; PME 0.15; ALE–AME 0.06; AME–AME 0.40; PLE–PME 0.20; PME–PME 0.09; MOQ: AW 0.33, PW 0.48, L 0.35.</paragraph>
<paragraphid="B941511CFFEDFFDFFF170BF8FD7DDCF9"blockId="21.[189,680,988,1015]"box="[189,680,988,1015]"pageId="21"pageNumber="22">CLYPEUS. 0.25 high, with eight long setae.</paragraph>
<paragraphid="B941511CFFEDFFDFFF170C3AFDEEDB55"blockId="21.[189,1398,1054,1116]"pageId="21"pageNumber="22">LEGS. Trochanter I notched, trochanter IV shallowly notched. Unpaired claw smooth, rudimentary on legs III and IV. Scopulae absent.</paragraph>
<figureCitationid="21C54D99FFEDFFDFFEFB0E97FE01D9C0"box="[337,468,1715,1742]"captionStart="Fig"captionStartId="18.[189,233,1787,1813]"captionTargetBox="[219,1365,478,1733]"captionTargetId="figure-82@18.[189,1398,445,1749]"captionTargetPageId="18"captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001800"httpUri="https://zenodo.org/record/14001800/files/figure.png"pageId="21"pageNumber="22">Figs 5A–C</figureCitation>
,
<figureCitationid="21C54D99FFEDFFDFFE4B0E97FE27D9C3"box="[481,498,1715,1741]"captionStart="Fig"captionStartId="19.[188,230,1787,1813]"captionTargetBox="[188,1399,479,1749]"captionTargetId="figure-89@19.[188,1399,479,1749]"captionTargetPageId="19"captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–C = 0.43 mm; D–F = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick."figureDoi="http://doi.org/10.5281/zenodo.14001804"httpUri="https://zenodo.org/record/14001804/files/figure.png"pageId="21"pageNumber="22">6</figureCitation>
). Tibia with slender apical VTA, bifid apical RTA continuous with elongate, slender basal RTA; tibia with dorso-retrolateral excavation (BC) bordered dorsad of RTA, accommodating conical retrobasal cymbial extension; cymbium oval with rounded apical extremity, without dorsal patch of chemosensitive setae, with sinuous retrolateral margin, with conical retrobasal cymbial extension (CE) that extends ⅓ of tibial length from tibial apex; subtegulum large, filling most of cymbial concavity, with rounded lobe extending proximad and small, conical lobe (SL) that interacts with the tegulum; tegulum oval with large, rounded retrodistal protuberance and conical promedian lobe (TL) that locks with subtegulum; embolus (E) arises from center of tegulum, short, broad; strongly sclerotized MA forms short hook curved prolaterad; conductor (C) hyaline, translucent, flattened, fan-shaped, narrow at base and strongly widened at extremity; paraconductor (Pc) as finger-like membranous structure originating at base of MA and extending to anterior half of E.
<figureCitationid="21C54D99FFEEFFDCFE9F09F7FE0BDEE0"box="[309,478,467,494]"captionStart="Fig"captionStartId="16.[189,233,1892,1918]"captionTargetBox="[189,1398,467,1854]"captionTargetId="figure-79@16.[189,1398,467,1854]"captionTargetPageId="16"captionText="Fig. 3. Tabiboka milleri gen. et sp. nov., A–B, G = male paratype (CASENT_9016097; CAS); C = male paratype (CASENT_9016098; CAS); D–F, H = female paratype (CASENT_9016094; CAS). A, D. Habitus, dorsal view. B, E. Idem, ventral view. C, F. Carapace, dorsal view. G–H. Habitus, lateral view. Scale bars = 2 mm. Stacked microphotographs by Hannah Wood."figureDoi="http://doi.org/10.5281/zenodo.14001794"httpUri="https://zenodo.org/record/14001794/files/figure.png"pageId="22"pageNumber="23">Figs 3D–F, H</figureCitation>
). As in male except abdomen much darker, mostly brown with lighter paracardiac marks and faint posterodorsal chevrons, sides dark, venter yellow brown with transverse dark mottling.
<figureCitationid="21C54D99FFEEFFDCFE970A78FE42DD79"box="[317,407,604,631]"captionStart="Fig"captionStartId="16.[189,233,1892,1918]"captionTargetBox="[189,1398,467,1854]"captionTargetId="figure-79@16.[189,1398,467,1854]"captionTargetPageId="16"captionText="Fig. 3. Tabiboka milleri gen. et sp. nov., A–B, G = male paratype (CASENT_9016097; CAS); C = male paratype (CASENT_9016098; CAS); D–F, H = female paratype (CASENT_9016094; CAS). A, D. Habitus, dorsal view. B, E. Idem, ventral view. C, F. Carapace, dorsal view. G–H. Habitus, lateral view. Scale bars = 2 mm. Stacked microphotographs by Hannah Wood."figureDoi="http://doi.org/10.5281/zenodo.14001794"httpUri="https://zenodo.org/record/14001794/files/figure.png"pageId="22"pageNumber="23">Fig. 3E</figureCitation>
). Oval with slightly sinuous margins; 2.60 wide, 3.00 long.
<paragraphid="B941511CFFEEFFDCFF170AC4FCCBDC10"blockId="22.[188,1399,736,798]"pageId="22"pageNumber="23">EYES. ALE 0.18; AME 0.13; PLE 0.18; PME 0.15; ALE–AME 0.06; AME–AME 0.40; PLE–PME 0.25; PME–PME 0.11; MOQ: AW 0.40, PW 0.55, L 0.48.</paragraph>
<paragraphid="B941511CFFEEFFDCFF170B61FD7DDC6E"blockId="22.[189,680,837,864]"box="[189,680,837,864]"pageId="22"pageNumber="23">CLYPEUS. 0.30 high, with eight long setae.</paragraph>
<figureCitationid="21C54D99FFEEFFDCFE6B0E3AFDF2D937"box="[449,551,1566,1593]"captionStart="Fig"captionStartId="18.[189,233,1787,1813]"captionTargetBox="[219,1365,478,1733]"captionTargetId="figure-82@18.[189,1398,445,1749]"captionTargetPageId="18"captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001800"httpUri="https://zenodo.org/record/14001800/files/figure.png"pageId="22"pageNumber="23">Figs 5D</figureCitation>
,
<figureCitationid="21C54D99FFEEFFDCFD9F0E3AFDABD936"box="[565,638,1566,1592]"captionStart="Fig"captionStartId="20.[189,232,1822,1848]"captionTargetBox="[189,1398,461,1784]"captionTargetId="figure-79@20.[189,1398,461,1784]"captionTargetPageId="20"captionText="Fig. 7. Tabiboka milleri gen. nov. et sp. nov. SEM of female genitalia, A–B, E–F = female paratype (CASENT_9016094; CAS); C–D = female paratype (CASENT_9035866; CAS). A. Epigyne, ventral view. B. Idem, ventro-posterior view. C. Endogyne, dorsal view. D. Idem, slightly anterior view. E. Spermatheca, right dorsal view. F. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: A–E = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila."figureDoi="http://doi.org/10.5281/zenodo.14001806"httpUri="https://zenodo.org/record/14001806/files/figure.png"pageId="22"pageNumber="23">7A–B</figureCitation>
) with broad, oval median lobe (ML) that extends posteriad of epigastric furrow; lateral lobes (LL) convex, lacking teeth or concavities; copulatory openings (CO) located beneath anteromedian margin of median lobe and, in many specimens, containing simple plugs. Endogyne (
<figureCitationid="21C54D99FFEEFFDCFEEA0EACFE49D9AD"box="[320,412,1672,1699]"captionStart="Fig"captionStartId="18.[189,233,1787,1813]"captionTargetBox="[219,1365,478,1733]"captionTargetId="figure-82@18.[189,1398,445,1749]"captionTargetPageId="18"captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001800"httpUri="https://zenodo.org/record/14001800/files/figure.png"pageId="22"pageNumber="23">Figs 5E</figureCitation>
,
<figureCitationid="21C54D99FFEEFFDCFE0D0EADFE3CD9AD"box="[423,489,1673,1699]"captionStart="Fig"captionStartId="20.[189,232,1822,1848]"captionTargetBox="[189,1398,461,1784]"captionTargetId="figure-79@20.[189,1398,461,1784]"captionTargetPageId="20"captionText="Fig. 7. Tabiboka milleri gen. nov. et sp. nov. SEM of female genitalia, A–B, E–F = female paratype (CASENT_9016094; CAS); C–D = female paratype (CASENT_9035866; CAS). A. Epigyne, ventral view. B. Idem, ventro-posterior view. C. Endogyne, dorsal view. D. Idem, slightly anterior view. E. Spermatheca, right dorsal view. F. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: A–E = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila."figureDoi="http://doi.org/10.5281/zenodo.14001806"httpUri="https://zenodo.org/record/14001806/files/figure.png"pageId="22"pageNumber="23">7C–F</figureCitation>
) with spermathecae (S) dark, smoothly convex, simple or with slight transverse indentations; sperm ducts (CD) anteromedian, short, joining seamlessly with spermathecae; spermathecal heads with 20–30 small pores; fertilization ducts (FD) short, flattened and slightly curved (
<figureCitationid="21C54D99FFEEFFDCFB700EEBFAEFD9E4"box="[1242,1338,1743,1770]"captionStart="Fig"captionStartId="18.[189,233,1787,1813]"captionTargetBox="[219,1365,478,1733]"captionTargetId="figure-82@18.[189,1398,445,1749]"captionTargetPageId="18"captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001800"httpUri="https://zenodo.org/record/14001800/files/figure.png"pageId="22"pageNumber="23">Figs 5E</figureCitation>
,
<figureCitationid="21C54D99FFEEFFDCFAEF0EF4FABCD9E4"box="[1349,1385,1744,1770]"captionStart="Fig"captionStartId="20.[189,232,1822,1848]"captionTargetBox="[189,1398,461,1784]"captionTargetId="figure-79@20.[189,1398,461,1784]"captionTargetPageId="20"captionText="Fig. 7. Tabiboka milleri gen. nov. et sp. nov. SEM of female genitalia, A–B, E–F = female paratype (CASENT_9016094; CAS); C–D = female paratype (CASENT_9035866; CAS). A. Epigyne, ventral view. B. Idem, ventro-posterior view. C. Endogyne, dorsal view. D. Idem, slightly anterior view. E. Spermatheca, right dorsal view. F. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: A–E = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila."figureDoi="http://doi.org/10.5281/zenodo.14001806"httpUri="https://zenodo.org/record/14001806/files/figure.png"pageId="22"pageNumber="23">7E</figureCitation>
<paragraphid="B941511CFFEEFFDDFF170F40FB25DD5A"blockId="22.[188,1400,1856,2025]"lastBlockId="23.[189,1399,265,596]"lastPageId="23"lastPageNumber="24"pageId="22"pageNumber="23">Male genitalia display slight geographic variation. Males from Ranomafana have very slender retrobasal tibial processes, with length greater than 20 times width, whereas in males from Ankazomivay and Tsinjoarivo these are thick, with length less than 15 times width. The distal RTA may be deeply cleft in males from Ranomafana or only slightly cleft as in males from Ankazomivady. Most individuals have a crack across the subbasal part of the leg tibia just distad of the basalmost ventral spines, but this varies. Within the Ranomafana population some have the tibial leg crack clearly marked whereas in others the crack cannot be distinguished. Total length = 9.60–12.00, carapace length/width = 1.28–1.41, carapace height/width = 0.44–0.59, PER/carapace width = 0.24–0.26, PER/OAL = 2.33–2.65, PER/AER = 1.38– 1.46, OAL/OQL = 1.24–1.42, OQP/OQA = 1.46–1.57. Clypeus height/AME = 1.20–2.86, cheliceral length/clypeus height = 8.00–13.33, sternum length/width = 1.11–1.19, palpal coxa length/width = 2.14–2.50, femur I length/carapace width = 1.04–1.22, metatarsus I length/carapace width = 1.02–1.11, femur IV length/carapace width = 1.09–1.30, cymbium length/carapace width = 0.36–0.42, cymbium length/palpal patella length = 1.45–2.43, cymbium length/palpal tibia length = 1.23–1.90, cymbium length/palpal femur length = 0.76–0.92, palpal tibia length/palpal patella length 1.00–1.57.</paragraph>
<figureCitationid="21C54D99FFEFFFDDFB6E0BACFAF1DCAC"box="[1220,1316,903,930]"captionStart="Fig"captionStartId="18.[189,233,1787,1813]"captionTargetBox="[219,1365,478,1733]"captionTargetId="figure-82@18.[189,1398,445,1749]"captionTargetPageId="18"captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001800"httpUri="https://zenodo.org/record/14001800/files/figure.png"pageId="23"pageNumber="24">Fig. 5D</figureCitation>
). Median lobe length 0.93–1.38 times width, with median lobe extending posteriad of epigastric furrow by 0.11–0.29 of lobe length. Spermathecae may be evenly rounded or have as many as two shallow dorsal constrictions. Spermathecae separated by 0.27–0.35 the total width of the spermathecae.
Specimens of this species have been used as exemplars in previous studies including morphological documentation and in published molecular phylogenies. The morphology of the species was documented for the Spiders Assembling the Tree of Life (AToL) study (2001–2009) as the exemplar “
<bibRefCitationid="DD6F2CEDFFEFFFDDFCAF0D27FC0ADA13"author="Morphbank"box="[773,991,1283,1309]"pageId="23"pageNumber="24"refId="ref37471"refString="Morphbank. 2023. Biological Imaging. Florida State University, Department of Scientific Computing, Tallahassee, FL 32306 - 4026 USA. Available from https: // www. morphbank. net / [accessed 20 Oct. 2023]."type="url"year="2023">Morphbank 2023</bibRefCitation>
): CASENT_9016094. Specimens were used as the exemplars “
<bibRefCitationid="DD6F2CEDFFEFFFDDFC890D01FBCADA4E"author="Polotow D. & Carmichael A. & Griswold C. E."box="[803,1055,1317,1344]"pageId="23"pageNumber="24"pagination="124 - 163"refId="ref37852"refString="Polotow D., Carmichael A. & Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041"type="journal article"year="2015">
<bibRefCitationid="DD6F2CEDFFEFFFDDFDBC0D63FCC5DA6F"author="Wheeler W. C. & Coddington J. A. & Crowley L. M. & Dimitrov D. & Goloboff P. A. & Griswold C. E. & Hormiga G. & Prendini L. & Ramirez M. J. & Sierwald P. & Almeida-Silva L. M. & Alvarez-Padilla F. & Arnedo M. A. & Benavides L. R. & Benjamin S. P. & Bond J. E. & Grismado C. J. & Hasan E. & Hedin M. & Izquierdo M. A. & Labarque F. M. & Ledford J. & Lopardo L. & Maddison W. P. & Miller J. A. & Piacentini L. N. & Platnick N. I. & Polotow D. & Silva-Davila D. & Scharff N. & Szuts T. & Ubick D. & Vink C. & Wood H. M. & Zhang J. X."box="[534,784,1351,1377]"pageId="23"pageNumber="24"pagination="576 - 616"refId="ref39136"refString="Wheeler W. C., Coddington J. A., Crowley L. M., Dimitrov D., Goloboff P. A., Griswold C. E., Hormiga G., Prendini L., Ramirez M. J., Sierwald P., Almeida-Silva L. M., Alvarez-Padilla F., Arnedo M. A., Benavides L. R., Benjamin S. P., Bond J. E., Grismado C. J., Hasan E., Hedin M., Izquierdo M. A., Labarque F. M., Ledford J., Lopardo L., Maddison W. P., Miller J. A., Piacentini L. N., Platnick N. I., Polotow D., Silva-Davila D., Scharff N., Szuts T., Ubick D., Vink C., Wood H. M. & Zhang J. X. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. https: // doi. org / 10.1111 / cla. 12182"type="journal article"year="2017">
<paragraphid="B941511CFFEFFFDDFF170D90FCC3DAFE"blockId="23.[189,1398,1460,1520]"pageId="23"pageNumber="24">This species is commonly found running on leaf litter in closed-canopy montane rainforest. Individuals were also collected in pitfall traps or under objects.</paragraph>
<taxonomicNameid="7EFE2A9FFFEFFFDDFF170E67FE51D953"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[189,388,1603,1629]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="23"pageNumber="24"phylum="Arthropoda"rank="species"species="milleri"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFEFFFDDFE270E60FD90D950"box="[397,581,1604,1630]"pageId="23"pageNumber="24"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<figureCitationid="21C54D99FFEFFFDDFD250E41FD32D971"box="[655,743,1637,1663]"captionStart="Fig"captionStartId="63.[189,231,1857,1883]"captionTargetBox="[189,1398,1005,1819]"captionTargetId="figure-208@63.[189,1398,1005,1819]"captionTargetPageId="63"captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)."figureDoi="http://doi.org/10.5281/zenodo.14001851"httpUri="https://zenodo.org/record/14001851/files/figure.png"pageId="23"pageNumber="24">Fig. 26</figureCitation>