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<document id="B42B9520F901316891EF99127CF1AB34" ID-CLB-Dataset="304853" ID-DOI="10.1649/0010-065X-77.1.19" ID-GBIF-Dataset="46a65ae6-3993-4152-b3ee-044f1620d6b0" ID-ISSN="1938-4394" ID-Zenodo-Dep="13242666" ID-ZooBank="urn:lsid:zoobank.org:pub:9FDA95A2-78A0-469B-8A8A-EF75A959E49E" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1722966744805" checkinUser="felipe" docAuthor="Ratcliffe, Brett C. &amp; Seidel, Matthias" docDate="2023" docId="03EB5D28FF8306484DA0FB44FCF18DD2" docLanguage="en" docName="TheColeopBull.77.1.19-23.pdf" docOrigin="The Coleopterists Bulletin 77 (1)" docSource="http://dx.doi.org/10.1649/0010-065x-77.1.19" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Cyclocephala ignota Ratcliffe and Seidel 2023, new species" docType="treatment" docUuid="97A9536A-74EC-4334-92EB-3CB936680056" docUuidSource="ZooBank" docVersion="3" lastPageNumber="22" masterDocId="FFD22550FF81064C4D40FFC9FFF08E2B" masterDocTitle="Cyclocephala ignota, a New Species of Cyclocephala Dejean, 1821 from Bolivia (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini)" masterLastPageNumber="23" masterPageNumber="19" pageNumber="20" updateTime="1730119312136" updateUser="ExternalLinkService">
<mods:mods id="E64B7072083DF27BD5E465F5CDA55BC9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F9A1D38ECD483EA7107D0B9CB51160BC">Cyclocephala ignota, a New Species of Cyclocephala Dejean, 1821 from Bolivia (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini)</mods:title>
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<mods:namePart id="ECFAE0093E96B7A9F95DFE8DB16488EB">Ratcliffe, Brett C.</mods:namePart>
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<mods:date id="0F5D3DE12B495196E2BE645058676136">2023</mods:date>
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<subSubSection id="C358BFB5FF83064E4DA0FB44FDF58A95" pageId="2" pageNumber="20" type="nomenclature">
<paragraph id="8BFDEC3EFF83064E4DA0FB44FDF58A95" blockId="2.[171,619,1165,1294]" pageId="2" pageNumber="20">
<emphasis id="B936302CFF83064E4DA0FB44FDF58A95" bold="true" pageId="2" pageNumber="20">
<taxonomicName id="4C4297BDFF83064E4DA0FB44FE758A95" ID-CoL="B7V5Z" authority="Ratcliffe and Seidel" authorityName="Ratcliffe and Seidel" authorityYear="2023" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="ignota" status="sp. nov.">
<emphasis id="B936302CFF83064E4DA0FB44FE5B8A88" bold="true" box="[224,427,1165,1187]" italics="true" pageId="2" pageNumber="20">Cyclocephala ignota</emphasis>
Ratcliffe and Seidel
</taxonomicName>
,
<taxonomicNameLabel id="A2058D57FF83064E4CCFFB61FDF58A95" box="[399,517,1192,1214]" pageId="2" pageNumber="20" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C358BFB5FF83064E4D94FB0AFE218B26" pageId="2" pageNumber="20" type="description">
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zoobank.org/
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urn:lsid:zoobank.org:act:
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</uri>
(
<figureCitation id="1379F0BBFF83064E4C0CFB31FE578B26" box="[332,423,1272,1294]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Figs. 1–7</figureCitation>
,
<figureCitation id="1379F0BBFF83064E4CF2FB31FE398B26" box="[434,457,1272,1293]" captionStart="Fig" captionStartId="4.[176,207,1437,1456]" captionTargetBox="[160,638,893,1420]" captionTargetId="figure-233@4.[152,638,893,1420]" captionTargetPageId="4" captionText="Fig. 11. Location of type locality of Cyclocephala ignota, new species at Caranavi, Caranavi Province in La Paz Department, Bolivia." figureDoi="http://doi.org/10.5281/zenodo.13242674" httpUri="https://zenodo.org/record/13242674/files/figure.png" pageId="2" pageNumber="20">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C358BFB5FF83064E4DF0FAEAFF0B8876" pageId="2" pageNumber="20" type="materials_examined">
<paragraph id="8BFDEC3EFF83064E4DF0FAEAFF0B8876" blockId="2.[152,639,1315,1737]" pageId="2" pageNumber="20">
<emphasis id="B936302CFF83064E4DF0FAEAFEB68B12" bold="true" box="[176,326,1315,1337]" pageId="2" pageNumber="20">Type Material.</emphasis>
<materialsCitation id="3B2AE663FF83064E4C0BFAEAFEFC8BA2" collectingDate="2010-03" collectionCode="NHMW" collectorName="Matthias" country="Bolivia" location="Caranavi" municipality="Caranavi" pageId="2" pageNumber="20" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54F9529CFF83064E4C0BFAEAFE548B12" box="[331,420,1315,1337]" pageId="2" pageNumber="20" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4427B7FF83064E4CE9FAEAFE4B8B13" box="[425,443,1315,1336]" count="1" pageId="2" pageNumber="20" type="male">♂</specimenCount>
labeled “
<collectingCountry id="F355ACAEFF83064E4F52FAEAFD8A8B13" box="[530,634,1315,1336]" name="Bolivia" pageId="2" pageNumber="20">BOLIVIA</collectingCountry>
: Nor Yungas, /
<collectingMunicipality id="6B997644FF83064E4C62FAF7FE8B8B7F" box="[290,379,1342,1364]" pageId="2" pageNumber="20">Caranavi</collectingMunicipality>
,
<date id="FFFCCAFEFF83064E4CC5FAF7FE3F8B7F" box="[389,463,1342,1364]" pageId="2" pageNumber="20" value="2010-03">
<collectingDate id="EFB83316FF83064E4CC5FAF7FE3F8B7F" box="[389,463,1342,1364]" pageId="2" pageNumber="20" value="2010-03">iii.2010</collectingDate>
</date>
/ local collector // Coll.
<collectorName id="26B789E8FF83064E4D8EFA91FED28B45" box="[206,290,1368,1390]" pageId="2" pageNumber="20">Matthias</collectorName>
/ Seidel 2020” and with our red holotype label
</materialsCitation>
.
<materialsCitation id="3B2AE663FF83064E4C54FABAFE038B8F" collectingDate="2010-03" collectionCode="NMPC, MSPC, BCRC" collectorName="Matthias" country="Bolivia" location="Caranavi" municipality="Caranavi" pageId="2" pageNumber="20" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D4427B7FF83064E4C54FABAFE938BA3" box="[276,355,1395,1417]" count="3" pageId="2" pageNumber="20" type="male">Three ♂</specimenCount>
<typeStatus id="54F9529CFF83064E4C29FABAFE358BA3" box="[361,453,1395,1416]" pageId="2" pageNumber="20" type="paratype">paratypes</typeStatus>
with the same data and with our yellow paratype label
</materialsCitation>
.
<typeStatus id="54F9529CFF83064E4CBDFA47FDA68B8F" box="[509,598,1422,1444]" pageId="2" pageNumber="20" type="holotype">Holotype</typeStatus>
deposited in the Naturhistorisches Museum Wien (
<collectionCode id="ED5374FBFF83064E4DDFFA0AFF038BF3" box="[159,243,1475,1496]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" pageId="2" pageNumber="20">NHMW</collectionCode>
), Austria. One paratype deposited in National Museum Prague (
<collectionCode id="ED5374FBFF83064E4CCFFA17FE268BD8" box="[399,470,1502,1523]" collectionName="Czech Republic, Prague, National Museum (Natural History)" pageId="2" pageNumber="20">NMPC</collectionCode>
), Czech Republic; one
<typeStatus id="54F9529CFF83064E4DFEFA30FEFE8825" box="[190,270,1529,1550]" pageId="2" pageNumber="20">paratype</typeStatus>
deposited in Matthias Seidel Collection (
<collectionCode id="ED5374FBFF83064E4DDFF9DAFF138803" box="[159,227,1555,1576]" collectionName="Italy, Varallo, Museo di Storia Naturale &quot;Pietro Calderini&quot;" pageId="2" pageNumber="20">MSPC</collectionCode>
), Wien, Austria; and one paratype deposited in the B. C. Ratcliffe Collection (
<collectionCode id="ED5374FBFF83064E4C97F9E7FDEC8868" box="[471,540,1582,1603]" collectionName="Brett C. Ratcliffe" pageId="2" pageNumber="20">BCRC</collectionCode>
),
<collectorName id="26B789E8FF83064E4F6DF9E7FF4A8875" pageId="2" pageNumber="20">Lincoln, NE</collectorName>
,
<collectingCountry id="F355ACAEFF83064E4D84F981FF078876" box="[196,247,1608,1629]" name="United States of America" pageId="2" pageNumber="20">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C358BFB5FF83064E4DF0F9AAFC0B8D23" pageId="2" pageNumber="20" type="diagnosis">
<paragraph id="8BFDEC3EFF83064E4DF0F9AAFBEE8F6A" blockId="2.[152,639,1315,1737]" lastBlockId="2.[685,1172,194,1736]" pageId="2" pageNumber="20">
<emphasis id="B936302CFF83064E4DF0F9AAFEE78852" bold="true" box="[176,279,1635,1657]" pageId="2" pageNumber="20">Diagnosis.</emphasis>
This species will key to couplet 55 for
<taxonomicName id="4C4297BDFF83064E4DD8F9B7FE2788B8" authority="Martinez" authorityName="Martinez" box="[152,471,1662,1683]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="pereirai">
<emphasis id="B936302CFF83064E4DD8F9B7FE8488B8" box="[152,372,1662,1683]" italics="true" pageId="2" pageNumber="20">Cyclocephala pereirai</emphasis>
Martínez
</taxonomicName>
(
<figureCitation id="1379F0BBFF83064E4CAAF9B7FDA788B8" box="[490,599,1662,1684]" captionStart="Figs" captionStartId="4.[176,215,783,802]" captionTargetBox="[151,1171,193,765]" captionTargetId="figure-211@4.[151,1171,193,765]" captionTargetPageId="4" captionText="Figs. 8–10. Cyclocephala pereirai. 8) Dorsal view; 9) Lateral view (to show body shape only); 10) Parameres, caudal view." figureDoi="http://doi.org/10.5281/zenodo.13242672" httpUri="https://zenodo.org/record/13242672/files/figure.png" pageId="2" pageNumber="20">Figs. 8–10</figureCitation>
) in
<bibRefCitation id="EFD391CFFF83064E4DD8F951FEDE8885" author="Endrodi, S." box="[152,302,1688,1710]" pageId="2" pageNumber="20" refId="ref1883" refString="Endrodi, S. 1985. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, The Netherlands, 800 pp." type="book" year="1985">Endrödi (1985)</bibRefCitation>
where the protarsi are considered scarcely thickened and with a slightly curved inner claw versus strongly thickened and with a strongly curved inner claw.
<taxonomicName id="4C4297BDFF83064E4EFBFF15FB628ED9" box="[955,1170,220,242]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="pereirai">
<emphasis id="B936302CFF83064E4EFBFF15FB628ED9" box="[955,1170,220,242]" italics="true" pageId="2" pageNumber="20">Cyclocephala pereirai</emphasis>
</taxonomicName>
was described from
<collectingCountry id="F355ACAEFF83064E4E31FF3EFC218F26" box="[881,977,247,269]" name="Argentina" pageId="2" pageNumber="20">Argentina</collectingCountry>
and remains rare in collections, and its parameres are similar to those of
<taxonomicName id="4C4297BDFF83064E4F87FEE4FCD08F69" box="[711,800,301,322]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="ignota">
<emphasis id="B936302CFF83064E4F87FEE4FCD08F69" box="[711,800,301,322]" italics="true" pageId="2" pageNumber="20">C. ignota</emphasis>
</taxonomicName>
(compare
<figureCitation id="1379F0BBFF83064E4EC7FEE5FC398F69" box="[903,969,300,322]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Figs. 6</figureCitation>
and
<figureCitation id="1379F0BBFF83064E4EB7FEE5FBE28F6A" box="[1015,1042,300,321]" captionStart="Figs" captionStartId="4.[176,215,783,802]" captionTargetBox="[151,1171,193,765]" captionTargetId="figure-211@4.[151,1171,193,765]" captionTargetPageId="4" captionText="Figs. 8–10. Cyclocephala pereirai. 8) Dorsal view; 9) Lateral view (to show body shape only); 10) Parameres, caudal view." figureDoi="http://doi.org/10.5281/zenodo.13242672" httpUri="https://zenodo.org/record/13242672/files/figure.png" pageId="2" pageNumber="20">10</figureCitation>
).
</paragraph>
<paragraph id="8BFDEC3EFF83064E4F85FE8EFC0B8D23" blockId="2.[685,1172,194,1736]" pageId="2" pageNumber="20">
We characterize the male protarsi of our new species as slender and only slightly curved, thus suggesting
<taxonomicName id="4C4297BDFF83064E4E63FEB4FC748FB9" box="[803,900,381,402]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="ignota">
<emphasis id="B936302CFF83064E4E63FEB4FC748FB9" box="[803,900,381,402]" italics="true" pageId="2" pageNumber="20">C. ignota</emphasis>
</taxonomicName>
is closest to
<taxonomicName id="4C4297BDFF83064E495CFEB4FB7C8FB9" box="[1052,1164,381,402]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="pereirai">
<emphasis id="B936302CFF83064E495CFEB4FB7C8FB9" box="[1052,1164,381,402]" italics="true" pageId="2" pageNumber="20">C. pereirai</emphasis>
</taxonomicName>
. However, in dorsal view the proportions of the pronotum and elytra are different between the two species (compare
<figureCitation id="1379F0BBFF83064E4E28FE05FC5F8FCA" box="[872,943,460,482]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Figs. 1</figureCitation>
and
<figureCitation id="1379F0BBFF83064E4EA5FE05FC028FCA" box="[997,1010,460,481]" captionStart="Figs" captionStartId="4.[176,215,783,802]" captionTargetBox="[151,1171,193,765]" captionTargetId="figure-211@4.[151,1171,193,765]" captionTargetPageId="4" captionText="Figs. 8–10. Cyclocephala pereirai. 8) Dorsal view; 9) Lateral view (to show body shape only); 10) Parameres, caudal view." figureDoi="http://doi.org/10.5281/zenodo.13242672" httpUri="https://zenodo.org/record/13242672/files/figure.png" pageId="2" pageNumber="20">8</figureCitation>
), and in lateral view
<taxonomicName id="4C4297BDFF83064E4FAAFE2EFCBD8FD6" box="[746,845,487,509]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="ignota">
<emphasis id="B936302CFF83064E4FAAFE2EFCBD8FD6" box="[746,845,487,509]" italics="true" pageId="2" pageNumber="20">C. ignota</emphasis>
</taxonomicName>
is dorsoventrally compressed, whereas
<taxonomicName id="4C4297BDFF83064E4E45FDCBFC818C3C" box="[773,881,514,535]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="pereirai">
<emphasis id="B936302CFF83064E4E45FDCBFC818C3C" box="[773,881,514,535]" italics="true" pageId="2" pageNumber="20">C. pereirai</emphasis>
</taxonomicName>
is not (as is typical for most
<taxonomicName id="4C4297BDFF83064E4FEDFDD5FC718C19" box="[685,897,540,562]" pageId="2" pageNumber="20">
<emphasis id="B936302CFF83064E4FEDFDD5FCC48C1A" box="[685,820,540,561]" italics="true" pageId="2" pageNumber="20">Cyclocephala</emphasis>
species
</taxonomicName>
) (compare
<figureCitation id="1379F0BBFF83064E4EAFFDD5FBC18C1A" box="[1007,1073,540,562]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Figs. 2</figureCitation>
and
<figureCitation id="1379F0BBFF83064E491FFDD5FB9D8C1A" box="[1119,1133,540,561]" captionStart="Figs" captionStartId="4.[176,215,783,802]" captionTargetBox="[151,1171,193,765]" captionTargetId="figure-211@4.[151,1171,193,765]" captionTargetPageId="4" captionText="Figs. 8–10. Cyclocephala pereirai. 8) Dorsal view; 9) Lateral view (to show body shape only); 10) Parameres, caudal view." figureDoi="http://doi.org/10.5281/zenodo.13242672" httpUri="https://zenodo.org/record/13242672/files/figure.png" pageId="2" pageNumber="20">9</figureCitation>
). In addition,
<taxonomicName id="4C4297BDFF83064E4E4DFDFEFC9B8C66" box="[781,875,567,589]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="ignota">
<emphasis id="B936302CFF83064E4E4DFDFEFC9B8C66" box="[781,875,567,589]" italics="true" pageId="2" pageNumber="20">C. ignota</emphasis>
</taxonomicName>
has long, dense setae on the ventral surfaces of the metatibiae and metatarsomeres, and
<taxonomicName id="4C4297BDFF83064E4E67FDA4FC798CA9" box="[807,905,621,642]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="pereirai">
<emphasis id="B936302CFF83064E4E67FDA4FC798CA9" box="[807,905,621,642]" italics="true" pageId="2" pageNumber="20">C. pereirai</emphasis>
</taxonomicName>
does not (compare
<figureCitation id="1379F0BBFF83064E4977FDA5FB7E8CAA" box="[1079,1166,620,642]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Figs.1, 3</figureCitation>
, and 5 vs.
<figureCitation id="1379F0BBFF83064E4E4EFD4EFCB98CB7" box="[782,841,647,669]" captionStart="Figs" captionStartId="4.[176,215,783,802]" captionTargetBox="[151,1171,193,765]" captionTargetId="figure-211@4.[151,1171,193,765]" captionTargetPageId="4" captionText="Figs. 8–10. Cyclocephala pereirai. 8) Dorsal view; 9) Lateral view (to show body shape only); 10) Parameres, caudal view." figureDoi="http://doi.org/10.5281/zenodo.13242672" httpUri="https://zenodo.org/record/13242672/files/figure.png" pageId="2" pageNumber="20">Fig. 8</figureCitation>
). Lastly, the color of
<taxonomicName id="4C4297BDFF83064E495FFD4EFB8C8CB6" box="[1055,1148,647,669]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="ignota">
<emphasis id="B936302CFF83064E495FFD4EFB8C8CB6" box="[1055,1148,647,669]" italics="true" pageId="2" pageNumber="20">C. ignota</emphasis>
</taxonomicName>
is light brown, and the color of
<taxonomicName id="4C4297BDFF83064E4E93FD6BFBCE8C9C" box="[979,1086,674,695]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="20" phylum="Arthropoda" rank="species" species="pereirai">
<emphasis id="B936302CFF83064E4E93FD6BFBCE8C9C" box="[979,1086,674,695]" italics="true" pageId="2" pageNumber="20">C. pereirai</emphasis>
</taxonomicName>
is testaceous to reddish brown, although color may be unreliable depending on age of the specimens and method of specimen preservation.
</paragraph>
</subSubSection>
<subSubSection id="C358BFB5FF83064F4F85FCC5FE0D88F9" lastPageId="3" lastPageNumber="21" pageId="2" pageNumber="20" type="description">
<paragraph id="8BFDEC3EFF83064F4F85FCC5FED088AA" blockId="2.[685,1172,194,1736]" lastBlockId="3.[173,659,1617,1746]" lastPageId="3" lastPageNumber="21" pageId="2" pageNumber="20">
<emphasis id="B936302CFF83064E4F85FCC5FC388D0A" bold="true" box="[709,968,780,802]" pageId="2" pageNumber="20">
Description of
<typeStatus id="54F9529CFF83064E4E2BFCC5FC388D0A" box="[875,968,780,801]" pageId="2" pageNumber="20" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="1379F0BBFF83064E4E9EFCC5FBB28D0A" box="[990,1090,780,802]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Figs. 1–7</figureCitation>
). Male. Length 12.0 mm; width 6.5 mm. Head dark brown, reminder of body light brown. Body dorsoventrally compressed (
<figureCitation id="1379F0BBFF83064E4E70FC95FC808D5A" box="[816,880,860,882]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Fig. 2</figureCitation>
).
<emphasis id="B936302CFF83064E4EC6FC95FC358D5A" bold="true" box="[902,965,860,881]" pageId="2" pageNumber="20">Head:</emphasis>
Surface completely, densely rugopunctate with long, yellowish brown setae on frons either side of middle. Frontoclypeal region with a slender, transversely biarcuate carinula not reaching lateral margins. Clypeal apex broadly parabolic, distinctly and narrowly reflexed. Interocular width equals 8.0 transverse eye diameters; ocular canthi short. Antenna with 9 antennomeres, club subequal in length to antennomeres 2–6. Mandibles projecting beyond clypeal apex (
<figureCitation id="1379F0BBFF83064E490DFB85FB748A4A" box="[1101,1156,1100,1122]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Fig. 1</figureCitation>
). Mentum slightly convex, strongly narrowed to apex, and with long, dense, yellowish brown setae.
<emphasis id="B936302CFF83064E4FEDFB55FCD18A9A" bold="true" box="[685,801,1180,1201]" pageId="2" pageNumber="20">Pronotum:</emphasis>
Widest just before middle (
<figureCitation id="1379F0BBFF83064E4906FB55FB748A9A" box="[1094,1156,1180,1202]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Fig. 1</figureCitation>
). Surface with punctures moderate in size and density, setigerous; setae long, dense, yellowish brown (
<figureCitation id="1379F0BBFF83064E4FF4FB25FD1F8B2A" box="[692,751,1260,1282]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Fig. 4</figureCitation>
). Base with marginal bead.
<emphasis id="B936302CFF83064E4EBAFB25FBB38B2A" bold="true" box="[1018,1091,1260,1281]" pageId="2" pageNumber="20">Elytra:</emphasis>
Surface weakly roughened and disc with small, dense, shallow, setigerous punctures; setae long, sparse, yellowish brown. Sides weakly, transversely wrinkled; setae along lateral margins dense (
<figureCitation id="1379F0BBFF83064E497DFA9EFB768B47" box="[1085,1158,1367,1389]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Fig. 3</figureCitation>
).
<emphasis id="B936302CFF83064E4FEDFAB8FCE88BAC" bold="true" box="[685,792,1393,1415]" pageId="2" pageNumber="20">Pygidium:</emphasis>
Disc with minute to small, sparse punctures; basal half and lateral angles more densely punctate and with long, sparse, yellowish brown setae (
<figureCitation id="1379F0BBFF83064E4FABFA0BFCD78BFC" box="[747,807,1473,1496]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Fig. 5</figureCitation>
). In lateral view, surface weakly convex.
<emphasis id="B936302CFF83064E4F9DFA15FCE48BDA" bold="true" box="[733,788,1500,1521]" pageId="2" pageNumber="20">Legs:</emphasis>
Protibia tridentate. Protarsomere 5 and median claw slightly enlarged; claw weakly arcuate, slender, apex narrowly cleft. Metatarsus longer than metatibia, both with long, dense, yellowish brown setae on their ventral surfaces (
<figureCitation id="1379F0BBFF83064E4954F98EFB768876" box="[1044,1158,1607,1629]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Figs. 1, 3, 5</figureCitation>
).
<emphasis id="B936302CFF83064E4FEDF9ABFD0B885C" bold="true" box="[685,763,1634,1655]" pageId="2" pageNumber="20">Venter:</emphasis>
Prosternal process, if present, small and obscured by long, dense, yellowish brown setae. Thoracic ventrites and coxae with long, dense, yellowish brown setae (
<figureCitation id="1379F0BBFF83064E4EE4F97BFC1488EC" box="[932,996,1714,1736]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="2" pageNumber="20">Fig. 3</figureCitation>
).
<emphasis id="B936302CFF83064E4EBBF97BFB8388EC" bold="true" box="[1019,1139,1714,1735]" pageId="2" pageNumber="20">Parameres:</emphasis>
In caudal view, shape subtriangular, apices rounded (
<figureCitation id="1379F0BBFF82064F4DF5F9A5FEE388AA" box="[181,275,1644,1666]" captionStart="Figs" captionStartId="3.[197,236,1537,1556]" captionTargetBox="[177,1182,203,1520]" captionTargetId="figure-7@3.[171,1195,193,1520]" captionTargetPageId="3" captionText="Figs. 1–7. Holotype of Cyclocephala ignota, new species. 1) Dorsal view; 2) Lateral view; 3) Ventral view; 4) Frontal view; 5) Caudal view; 6) Parameres, caudal view; 7) Parameres, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="3" pageNumber="21">Figs. 6, 7</figureCitation>
).
</paragraph>
<caption id="DF3DBCB6FF82064F4D85F9C8FC6A8807" ID-DOI="http://doi.org/10.5281/zenodo.14001856" ID-Zenodo-Dep="14001856" httpUri="https://zenodo.org/record/14001856/files/figure.png" pageId="3" pageNumber="21" startId="3.[197,236,1537,1556]" targetBox="[177,1182,203,1520]" targetPageId="3">
<paragraph id="8BFDEC3EFF82064F4D85F9C8FC6A8807" blockId="3.[173,1193,1537,1581]" pageId="3" pageNumber="21">
<emphasis id="B936302CFF82064F4D85F9C8FEEF883F" bold="true" box="[197,287,1537,1556]" pageId="3" pageNumber="21">Figs. 1–7.</emphasis>
Holotype of
<taxonomicName id="4C4297BDFF82064F4CE8F9C8FDAE883F" authorityName="Ratcliffe &amp; Seidel" authorityYear="2023" box="[424,606,1537,1556]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="ignota" status="sp. nov.">
<emphasis id="B936302CFF82064F4CE8F9C8FDAE883F" box="[424,606,1537,1556]" italics="true" pageId="3" pageNumber="21">Cyclocephala ignota</emphasis>
</taxonomicName>
,
<emphasis id="B936302CFF82064F4F2CF9CBFD29883F" bold="true" box="[620,729,1537,1557]" pageId="3" pageNumber="21">
<taxonomicNameLabel id="A2058D57FF82064F4F2CF9CBFD29883F" box="[620,729,1537,1557]" pageId="3" pageNumber="21" rank="species">new species</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B936302CFF82064F4FA7F9C8FD09883F" bold="true" box="[743,761,1537,1556]" pageId="3" pageNumber="21">1)</emphasis>
Dorsal view;
<emphasis id="B936302CFF82064F4E3DF9C8FC7F883F" bold="true" box="[893,911,1537,1556]" pageId="3" pageNumber="21">2)</emphasis>
Lateral view;
<emphasis id="B936302CFF82064F4957F9C8FBD8883F" bold="true" box="[1047,1064,1537,1556]" pageId="3" pageNumber="21">3)</emphasis>
Ventral view;
<emphasis id="B936302CFF82064F4DEDF9D0FF4F8807" bold="true" box="[173,191,1561,1580]" pageId="3" pageNumber="21">4)</emphasis>
Frontal view;
<emphasis id="B936302CFF82064F4C7CF9D0FEBE8807" bold="true" box="[316,334,1561,1580]" pageId="3" pageNumber="21">5)</emphasis>
Caudal view;
<emphasis id="B936302CFF82064F4C8AF9D0FE2C8807" bold="true" box="[458,476,1561,1580]" pageId="3" pageNumber="21">6)</emphasis>
Parameres, caudal view;
<emphasis id="B936302CFF82064F4FF6F9D0FD378807" bold="true" box="[694,711,1561,1580]" pageId="3" pageNumber="21">7)</emphasis>
Parameres, lateral view.
</paragraph>
</caption>
<paragraph id="8BFDEC3EFF82064F4D85F94FFE0D88F9" blockId="3.[173,659,1617,1746]" pageId="3" pageNumber="21">
<emphasis id="B936302CFF82064F4D85F94FFEDB88B7" bold="true" box="[197,299,1670,1692]" pageId="3" pageNumber="21">Variation.</emphasis>
Male (
<specimenCount id="9D4427B7FF82064F4C2DF94FFE2E88B0" box="[365,478,1670,1691]" count="3" pageId="3" pageNumber="21" type="generic" typeStatus="paratype">3 paratypes</specimenCount>
). Length 12.0–12.5 mm; width 6.2–7.0 mm. The
<typeStatus id="54F9529CFF82064F4C89F968FDD5889D" box="[457,549,1697,1718]" pageId="3" pageNumber="21" type="paratype">paratypes</typeStatus>
do not significantly differ from the
<typeStatus id="54F9529CFF82064F4CE2F975FE0988F9" box="[418,505,1724,1746]" pageId="3" pageNumber="21" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C358BFB5FF82064F4F9BF998FC4288FA" pageId="3" pageNumber="21" type="etymology">
<paragraph id="8BFDEC3EFF82064F4F9BF998FC4288FA" blockId="3.[707,1192,1617,1746]" pageId="3" pageNumber="21">
<emphasis id="B936302CFF82064F4F9BF998FCBD884D" bold="true" box="[731,845,1617,1638]" pageId="3" pageNumber="21">Etymology.</emphasis>
The specific epithet is from the Latin
<emphasis id="B936302CFF82064F4F83F9A5FCFA88AA" box="[707,778,1644,1665]" italics="true" pageId="3" pageNumber="21">ignotus</emphasis>
, an adjective referring to unknown or strange and used here in reference to the atypical body morphology of this species relative to other species of
<taxonomicName id="4C4297BDFF82064F4E68F975FC5C88FA" authorityName="DEJEAN" authorityYear="1821" box="[808,940,1724,1745]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B936302CFF82064F4E68F975FC5C88FA" box="[808,940,1724,1745]" italics="true" pageId="3" pageNumber="21">Cyclocephala</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF3DBCB6FF8506484DF0FCC6FEF48D11" ID-DOI="http://doi.org/10.5281/zenodo.13242672" ID-Zenodo-Dep="13242672" httpUri="https://zenodo.org/record/13242672/files/figure.png" pageId="4" pageNumber="22" startId="4.[176,215,783,802]" targetBox="[151,1171,193,765]" targetPageId="4" targetType="figure">
<paragraph id="8BFDEC3EFF8506484DF0FCC6FEF48D11" blockId="4.[152,1172,783,826]" pageId="4" pageNumber="22">
<emphasis id="B936302CFF8506484DF0FCC6FEE38D09" bold="true" box="[176,275,783,802]" pageId="4" pageNumber="22">Figs. 8–10.</emphasis>
<taxonomicName id="4C4297BDFF8506484C68FCC6FE198D09" box="[296,489,783,802]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="22" phylum="Arthropoda" rank="species" species="pereirai">
<emphasis id="B936302CFF8506484C68FCC6FE198D09" box="[296,489,783,802]" italics="true" pageId="4" pageNumber="22">Cyclocephala pereirai</emphasis>
</taxonomicName>
.
<emphasis id="B936302CFF8506484CB6FCC6FDF98D09" bold="true" box="[502,521,783,802]" pageId="4" pageNumber="22">8)</emphasis>
Dorsal view;
<emphasis id="B936302CFF8506484FC8FCC6FD6A8D09" bold="true" box="[648,666,783,802]" pageId="4" pageNumber="22">9)</emphasis>
Lateral view (to show body shape only);
<emphasis id="B936302CFF8506484950FCC6FBDC8D09" bold="true" box="[1040,1068,783,802]" pageId="4" pageNumber="22">10)</emphasis>
Parameres, caudal view.
</paragraph>
</caption>
<caption id="DF3DBCB6FF8506484DF0FA54FE818BCB" ID-DOI="http://doi.org/10.5281/zenodo.13242674" ID-Zenodo-Dep="13242674" httpUri="https://zenodo.org/record/13242674/files/figure.png" pageId="4" pageNumber="22" startId="4.[176,207,1437,1456]" targetBox="[160,638,893,1420]" targetPageId="4" targetType="figure">
<paragraph id="8BFDEC3EFF8506484DF0FA54FE818BCB" blockId="4.[152,638,1437,1505]" pageId="4" pageNumber="22">
<emphasis id="B936302CFF8506484DF0FA54FF058B9B" bold="true" box="[176,245,1437,1456]" pageId="4" pageNumber="22">Fig. 11.</emphasis>
Location of type locality of
<taxonomicName id="4C4297BDFF8506484F48FA54FF3F8BE3" authorityName="Ratcliffe &amp; Seidel" authorityYear="2023" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="22" phylum="Arthropoda" rank="species" species="ignota" status="sp. nov.">
<emphasis id="B936302CFF8506484F48FA54FF3F8BE3" italics="true" pageId="4" pageNumber="22">Cyclocephala ignota</emphasis>
</taxonomicName>
,
<emphasis id="B936302CFF8506484D98FA7CFEB18BE3" bold="true" box="[216,321,1461,1480]" pageId="4" pageNumber="22">
<taxonomicNameLabel id="A2058D57FF8506484D98FA7CFEB18BE3" box="[216,321,1461,1480]" pageId="4" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
at Caranavi, Caranavi Province in La Paz Department, Bolivia.
</paragraph>
</caption>
<subSubSection id="C358BFB5FF8506484DF0F9D8FF068886" pageId="4" pageNumber="22" type="distribution">
<paragraph id="8BFDEC3EFF8506484DF0F9D8FE188877" blockId="4.[152,639,1553,1709]" pageId="4" pageNumber="22">
<emphasis id="B936302CFF8506484DF0F9D8FEC0880C" bold="true" box="[176,304,1553,1575]" pageId="4" pageNumber="22">Distribution.</emphasis>
<taxonomicName id="4C4297BDFF8506484C75F9D8FE06880C" authorityName="Ratcliffe &amp; Seidel" authorityYear="2023" box="[309,502,1553,1575]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="22" phylum="Arthropoda" rank="species" species="ignota">
<emphasis id="B936302CFF8506484C75F9D8FE06880C" box="[309,502,1553,1575]" italics="true" pageId="4" pageNumber="22">Cyclocephala ignota</emphasis>
</taxonomicName>
is known only from Caranavi in the Department of
<collectingRegion id="498622DCFF8506484CBCF9E4FDB08869" box="[508,576,1581,1602]" country="Bolivia" name="La Paz" pageId="4" pageNumber="22">La Paz</collectingRegion>
in the Yungas region of
<collectingCountry id="F355ACAEFF8506484C02F98EFE798876" box="[322,393,1607,1629]" name="Bolivia" pageId="4" pageNumber="22">Bolivia</collectingCountry>
(
<figureCitation id="1379F0BBFF8506484CD6F98EFE2C8877" box="[406,476,1607,1629]" captionStart="Fig" captionStartId="4.[176,207,1437,1456]" captionTargetBox="[160,638,893,1420]" captionTargetId="figure-233@4.[152,638,893,1420]" captionTargetPageId="4" captionText="Fig. 11. Location of type locality of Cyclocephala ignota, new species at Caranavi, Caranavi Province in La Paz Department, Bolivia." figureDoi="http://doi.org/10.5281/zenodo.13242674" httpUri="https://zenodo.org/record/13242674/files/figure.png" pageId="4" pageNumber="22">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8BFDEC3EFF8506484DF0F9A8FF068886" blockId="4.[152,639,1553,1709]" pageId="4" pageNumber="22">
<emphasis id="B936302CFF8506484DF0F9A8FE98885C" bold="true" box="[176,360,1633,1655]" pageId="4" pageNumber="22">Locality Records.</emphasis>
<specimenCount id="9D4427B7FF8506484C33F9ABFDFF885C" box="[371,527,1633,1655]" count="4" pageId="4" pageNumber="22" type="generic">Four specimens</specimenCount>
examined.
<emphasis id="B936302CFF8506484DD8F9B5FED888B9" bold="true" box="[152,296,1660,1682]" pageId="4" pageNumber="22">
<collectingCountry id="F355ACAEFF8506484DD8F9B5FF0F88B9" box="[152,255,1660,1682]" name="Bolivia" pageId="4" pageNumber="22">BOLIVIA</collectingCountry>
(3):
</emphasis>
<collectingRegion id="498622DCFF8506484C6DF9B5FE8C88BA" box="[301,380,1660,1681]" country="Bolivia" name="La Paz" pageId="4" pageNumber="22">LA PAZ</collectingRegion>
(3):
<collectingMunicipality id="6B997644FF8506484CEAF9B5FD8A88B9" box="[426,634,1660,1682]" pageId="4" pageNumber="22">Caranavi Department</collectingMunicipality>
,
<location id="8E9DBAE5FF8506484DD8F95EFF028886" LSID="urn:lsid:plazi:treatment:03EB5D28FF8306484DA0FB44FCF18DD2:8E9DBAE5FF8506484DD8F95EFF028886" box="[152,242,1687,1709]" country="Bolivia" municipality="Caranavi Department" name="Caranavi" pageId="4" pageNumber="22" stateProvince="La Paz">Caranavi</location>
.
</paragraph>
</subSubSection>
<subSubSection id="C358BFB5FF8506484F85FCB0FCF18DD2" pageId="4" pageNumber="22" type="biology_ecology">
<paragraph id="8BFDEC3EFF8506484F85FCB0FBEF8DA5" blockId="4.[685,1171,888,1017]" box="[709,1055,888,911]" pageId="4" pageNumber="22">
<emphasis id="B936302CFF8506484F85FCB0FC428DA5" bold="true" box="[709,946,888,910]" pageId="4" pageNumber="22">Temporal Distribution.</emphasis>
March (4).
</paragraph>
<paragraph id="8BFDEC3EFF8506484F85FC5DFCF18DD2" blockId="4.[685,1171,888,1017]" pageId="4" pageNumber="22">
<emphasis id="B936302CFF8506484F85FC5DFC808D82" bold="true" box="[709,880,915,937]" pageId="4" pageNumber="22">Natural History.</emphasis>
Nothing is known of the life history of
<taxonomicName id="4C4297BDFF8506484E5AFC67FC898DEF" authorityName="Ratcliffe &amp; Seidel" authorityYear="2023" box="[794,889,942,964]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="22" phylum="Arthropoda" rank="species" species="ignota">
<emphasis id="B936302CFF8506484E5AFC67FC898DEF" box="[794,889,942,964]" italics="true" pageId="4" pageNumber="22">C. ignota</emphasis>
</taxonomicName>
. Adults of other species of
<taxonomicName id="4C4297BDFF8506484FEDFC00FCDC8DF5" authorityName="DEJEAN" authorityYear="1821" box="[685,812,969,990]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B936302CFF8506484FEDFC00FCDC8DF5" box="[685,812,969,990]" italics="true" pageId="4" pageNumber="22">Cyclocephala</emphasis>
</taxonomicName>
are nocturnal and commonly collected at lights.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="B941511CFFC2FFF0FDFF0DF7FC0ADAE1" blockId="58.[366,1221,1491,1590]" box="[597,991,1491,1519]" pageId="58" pageNumber="59">
<heading id="E209E670FFC2FFF0FDFF0DF7FC0ADAE1" box="[597,991,1491,1519]" centered="true" fontSize="11" level="2" pageId="58" pageNumber="59" reason="2">
<taxonomicName id="7EFE2A9FFFC2FFF0FDFF0DF7FCA8DAE0" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[597,893,1491,1518]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="silvadavilae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC2FFF0FDFF0DF7FCA8DAE0" bold="true" box="[597,893,1491,1518]" italics="true" pageId="58" pageNumber="59">Zorodictyna silvadavilae</emphasis>
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<paragraph id="B941511CFFC2FFF0FD720E38FC8ED938" blockId="58.[366,1221,1491,1590]" box="[728,859,1564,1590]" pageId="58" pageNumber="59">
<figureCitation id="21C54D99FFC2FFF0FD720E38FC8ED938" box="[728,859,1564,1590]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="59.[189,234,1894,1920]" captionStartId-1="61.[189,231,1752,1778]" captionStartId-2="63.[189,231,1857,1883]" captionTargetBox-0="[189,1398,1046,1855]" captionTargetBox-1="[211,1360,510,1693]" captionTargetBox-2="[189,1398,1005,1819]" captionTargetId-0="figure-180@59.[189,1398,1046,1855]" captionTargetId-1="figure-97@61.[189,1398,507,1714]" captionTargetId-2="figure-208@63.[189,1398,1005,1819]" captionTargetPageId-0="59" captionTargetPageId-1="61" captionTargetPageId-2="63" captionText-0="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; A–B = male holotype (CASENT_9006231; CAS); C–D = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." captionText-1="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." captionText-2="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi-0="http://doi.org/10.5281/zenodo.14001847" figureDoi-1="http://doi.org/10.5281/zenodo.14001849" figureDoi-2="http://doi.org/10.5281/zenodo.14001851" httpUri-0="https://zenodo.org/record/14001847/files/figure.png" httpUri-1="https://zenodo.org/record/14001849/files/figure.png" httpUri-2="https://zenodo.org/record/14001851/files/figure.png" pageId="58" pageNumber="59">Figs 24–26</figureCitation>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170E7BFD18D977" blockId="58.[189,717,1631,1657]" box="[189,717,1631,1657]" pageId="58" pageNumber="59">
<taxonomicName id="7EFE2A9FFFC2FFF0FF170E7BFEA1D977" box="[189,372,1631,1657]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8B8A8D0EFFC2FFF0FF170E7BFE9ED977" box="[189,331,1631,1657]" italics="true" pageId="58" pageNumber="59">Zorodictyna</emphasis>
sp.
</taxonomicName>
CG47 –
<bibRefCitation id="DD6F2CEDFFC2FFF0FE740E7BFD1CD977" author="Wheeler W. C. &amp; Coddington J. A. &amp; Crowley L. M. &amp; Dimitrov D. &amp; Goloboff P. A. &amp; Griswold C. E. &amp; Hormiga G. &amp; Prendini L. &amp; Ramirez M. J. &amp; Sierwald P. &amp; Almeida-Silva L. M. &amp; Alvarez-Padilla F. &amp; Arnedo M. A. &amp; Benavides L. R. &amp; Benjamin S. P. &amp; Bond J. E. &amp; Grismado C. J. &amp; Hasan E. &amp; Hedin M. &amp; Izquierdo M. A. &amp; Labarque F. M. &amp; Ledford J. &amp; Lopardo L. &amp; Maddison W. P. &amp; Miller J. A. &amp; Piacentini L. N. &amp; Platnick N. I. &amp; Polotow D. &amp; Silva-Davila D. &amp; Scharff N. &amp; Szuts T. &amp; Ubick D. &amp; Vink C. &amp; Wood H. M. &amp; Zhang J. X." box="[478,713,1631,1657]" pageId="58" pageNumber="59" pagination="576 - 616" refId="ref39136" refString="Wheeler W. C., Coddington J. A., Crowley L. M., Dimitrov D., Goloboff P. A., Griswold C. E., Hormiga G., Prendini L., Ramirez M. J., Sierwald P., Almeida-Silva L. M., Alvarez-Padilla F., Arnedo M. A., Benavides L. R., Benjamin S. P., Bond J. E., Grismado C. J., Hasan E., Hedin M., Izquierdo M. A., Labarque F. M., Ledford J., Lopardo L., Maddison W. P., Miller J. A., Piacentini L. N., Platnick N. I., Polotow D., Silva-Davila D., Scharff N., Szuts T., Ubick D., Vink C., Wood H. M. &amp; Zhang J. X. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. https: // doi. org / 10.1111 / cla. 12182" type="journal article" year="2017">
Wheeler
<emphasis id="8B8A8D0EFFC2FFF0FDE30E44FD51D977" box="[585,644,1631,1657]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
2017
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFC2FFF1FF170E85FE5EDE48" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="B941511CFFC2FFF0FF170E85FEE3D9B5" blockId="58.[189,310,1697,1723]" box="[189,310,1697,1723]" pageId="58" pageNumber="59">
<heading id="E209E670FFC2FFF0FF170E85FEE3D9B5" bold="true" box="[189,310,1697,1723]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="8B8A8D0EFFC2FFF0FF170E85FEE3D9B5" bold="true" box="[189,310,1697,1723]" pageId="58" pageNumber="59">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC2FFF1FF170EF5FE5EDE48" blockId="58.[188,1400,1745,2025]" lastBlockId="59.[188,1399,265,326]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
Males of
<taxonomicName id="7EFE2A9FFFC2FFF0FE990EF5FD8AD9E5" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[307,607,1745,1771]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="silvadavilae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC2FFF0FE990EF5FD8AD9E5" box="[307,607,1745,1771]" italics="true" pageId="58" pageNumber="59">Zorodictyna silvadavilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC2FFF0FDC20EF5FD13D9E5" box="[616,710,1745,1771]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
are distinguished from those of
<taxonomicName id="7EFE2A9FFFC2FFF0FBFE0EF6FAA3D9E5" authority="(Lenz, 1891)" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[1108,1398,1745,1771]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFC2FFF0FBFE0EF6FB07D9E5" box="[1108,1234,1745,1771]" italics="true" pageId="58" pageNumber="59">Z. oswaldi</emphasis>
(
<bibRefCitation id="DD6F2CEDFFC2FFF0FB490EF5FABBD9E5" author="Lenz H." box="[1251,1390,1745,1771]" pageId="58" pageNumber="59" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">Lenz, 1891</bibRefCitation>
)
</taxonomicName>
by the retrobasal cymbial extension (CE) that extends at least ¼ the length of papal tibia and thick retrolateral paracymbial process (PP) (vs
<taxonomicName id="7EFE2A9FFFC2FFF0FD170F3EFCEED83D" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[701,827,1817,1843]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFC2FFF0FD170F3EFCEED83D" box="[701,827,1817,1843]" italics="true" pageId="58" pageNumber="59">Z. oswaldi</emphasis>
</taxonomicName>
lacks the PP and has very short CE;
<bibRefCitation id="DD6F2CEDFFC2FFF0FAAD0F3DFF2BD856" author="Griswold C. E." pageId="58" pageNumber="59" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: figs 19–21); males are distinguished from
<taxonomicName id="7EFE2A9FFFC2FFF0FCAE0F1AFCB8D859" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[772,877,1853,1879]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC2FFF0FCAE0F1AFCB8D859" box="[772,877,1853,1879]" italics="true" pageId="58" pageNumber="59">Z. almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC2FFF0FCDC0F1AFC06D856" box="[886,979,1854,1880]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
by having only a small or no crest on embolus, and the retrolateral branch of RTA simple (
<figureCitation id="21C54D99FFC2FFF0FCFD0F46FC30D872" box="[855,997,1889,1916]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="58" pageNumber="59">Fig. 25A–C</figureCitation>
) (vs
<taxonomicName id="7EFE2A9FFFC2FFF0FB880F46FB5FD872" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1058,1162,1890,1916]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="almae">
<emphasis id="8B8A8D0EFFC2FFF0FB880F46FB5FD872" box="[1058,1162,1890,1916]" italics="true" pageId="58" pageNumber="59">Z. almae</emphasis>
</taxonomicName>
has a large crest on the E and the retrolateral branch of RTA is forked apically,
<figureCitation id="21C54D99FFC2FFF0FCDE0FA2FBD6D8AE" box="[884,1027,1926,1952]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="58" pageNumber="59">Fig. 23A–C</figureCitation>
). Females of
<taxonomicName id="7EFE2A9FFFC2FFF0FB350FA3FA99D8AE" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1183,1356,1926,1952]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="silvadavilae">
<emphasis id="8B8A8D0EFFC2FFF0FB350FA3FA99D8AE" box="[1183,1356,1926,1952]" italics="true" pageId="58" pageNumber="59">Z. silvadavilae</emphasis>
</taxonomicName>
are distinguished from
<taxonomicName id="7EFE2A9FFFC2FFF0FE340F8FFDC2D8CA" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[414,535,1962,1988]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFC2FFF0FE340F8FFDC2D8CA" box="[414,535,1962,1988]" italics="true" pageId="58" pageNumber="59">Z. oswaldi</emphasis>
</taxonomicName>
by lacking teeth on lateral lobes (LL) (
<figureCitation id="21C54D99FFC2FFF0FC770F8EFB92D8CB" box="[989,1095,1962,1989]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="58" pageNumber="59">Fig. 25D</figureCitation>
) (vs
<taxonomicName id="7EFE2A9FFFC2FFF0FBD50F8FFB2DD8CA" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[1151,1272,1962,1988]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFC2FFF0FBD50F8FFB2DD8CA" box="[1151,1272,1962,1988]" italics="true" pageId="58" pageNumber="59">Z. oswaldi</emphasis>
</taxonomicName>
have teeth on LL;
<bibRefCitation id="DD6F2CEDFFC2FFF0FEBD0FEAFE19D8E6" author="Griswold C. E." box="[279,460,1998,2024]" pageId="58" pageNumber="59" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: fig. 24). The females are distinguished from
<taxonomicName id="7EFE2A9FFFC2FFF0FC450FEBFB8DD8E6" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1007,1112,1998,2024]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="almae">
<emphasis id="8B8A8D0EFFC2FFF0FC450FEBFB8DD8E6" box="[1007,1112,1998,2024]" italics="true" pageId="58" pageNumber="59">Z. almae</emphasis>
</taxonomicName>
by having an endogyne with copulatory ducts (CD) equal to spermathecae (S) (
<figureCitation id="21C54D99FFC3FFF1FC91092DFC74DE2A" box="[827,929,265,292]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="59" pageNumber="60">Fig. 25E</figureCitation>
) (vs in
<taxonomicName id="7EFE2A9FFFC3FFF1FC5F092EFB8EDE2D" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1013,1115,265,291]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="almae">
<emphasis id="8B8A8D0EFFC3FFF1FC5F092EFB8EDE2D" box="[1013,1115,265,291]" italics="true" pageId="59" pageNumber="60">Z. almae</emphasis>
</taxonomicName>
, the CD are much larger than S,
<figureCitation id="21C54D99FFC3FFF1FEBE0908FEABDE48" box="[276,382,300,326]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="59" pageNumber="60">Fig. 23E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFC3FFF1FF17094AFE8DDEF2" pageId="59" pageNumber="60" type="etymology">
<paragraph id="B941511CFFC3FFF1FF17094AFE91DE86" blockId="59.[189,324,366,392]" box="[189,324,366,392]" pageId="59" pageNumber="60">
<heading id="E209E670FFC3FFF1FF17094AFE91DE86" bold="true" box="[189,324,366,392]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="8B8A8D0EFFC3FFF1FF17094AFE91DE86" bold="true" box="[189,324,366,392]" pageId="59" pageNumber="60">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC3FFF1FF1709B8FE8DDEF2" blockId="59.[189,1399,412,508]" pageId="59" pageNumber="60">
The species name is a matronym in honor of Dr Diana Silva Dávila, who made an extensive expedition to
<collectingCountry id="C1E9118CFFC3FFF1FF71099BFEBCDED7" box="[219,361,447,473]" name="Madagascar" pageId="59" pageNumber="60">Madagascar</collectingCountry>
in 2003, collecting specimens and tissues for the Spiders–AToL (Assembling the Tree of Life) project.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFC3FFF6FF170A00FDFEDD9C" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="materials_examined">
<paragraph id="B941511CFFC3FFF1FF170A00FEB9DD30" blockId="59.[189,364,548,574]" box="[189,364,548,574]" pageId="59" pageNumber="60">
<heading id="E209E670FFC3FFF1FF170A00FEB9DD30" bold="true" box="[189,364,548,574]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="8B8A8D0EFFC3FFF1FF170A00FEB9DD30" bold="true" box="[189,364,548,574]" pageId="59" pageNumber="60">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC3FFF1FF400A77FAA2DDD8" blockId="59.[189,1399,595,761]" pageId="59" pageNumber="60">
<materialsCitation id="09965B41FFC3FFF1FF400A77FAA2DDD8" collectingDate="1993-11-10" collectingDateMax="1993-11-16" collectingDateMin="1993-11-10" collectorName="C. Griswold &amp; J. Coddington &amp; N. Scharff &amp; S. Larcher &amp; R. Andriamasimanana" country="Madagascar" elevation="700" latitude="-14.433333" location="Marojejy Reserve" longLatPrecision="1288" longitude="49.75" pageId="59" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="holotype">
<emphasis id="8B8A8D0EFFC3FFF1FF400A77FE89DD63" bold="true" box="[234,348,595,621]" pageId="59" pageNumber="60">
<typeStatus id="6645EFBEFFC3FFF1FF400A77FE89DD63" box="[234,348,595,621]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="C1E9118CFFC3FFF1FF170A52FE47DD9E" box="[189,402,630,656]" name="Madagascar" pageId="59" pageNumber="60">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFC3FFF1FE1C0A51FD04DD81" bold="true" box="[438,721,629,655]" pageId="59" pageNumber="60">
<collectingRegion id="7B3A9FFEFFC3FFF1FE1C0A51FD04DD81" box="[438,721,629,655]" country="Madagascar" name="Antsiranana" pageId="59" pageNumber="60">Antsiranana Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFC3FFF1FD470A52FCD3DD9E" box="[749,774,630,656]" count="1" pageId="59" pageNumber="60" type="male">♂</specimenCount>
;
<location id="BC2107C7FFC3FFF1FCBF0A52FC24DD9E" LSID="urn:lsid:plazi:treatment:3157E00AFFC2FFF5FDFF0DF7FC8FDCA1:BC2107C7FFC3FFF1FCBF0A52FC24DD9E" box="[789,1009,630,656]" country="Madagascar" latitude="-14.433333" longLatPrecision="1288" longitude="49.75" name="Marojejy Reserve" pageId="59" pageNumber="60" stateProvince="Antsiranana">Marojejy Reserve</location>
,
<locationDeviation id="A8F3483EFFC3FFF1FC550A52FAA7DD9E" box="[1023,1394,630,656]" location="Manantenina" pageId="59" pageNumber="60">
<quantity id="7E06FCF9FFC3FFF1FC550A52FB81DD9E" box="[1023,1108,630,656]" metricMagnitude="3" metricUnit="m" metricValue="8.4" pageId="59" pageNumber="60" unit="km" value="8.4">8.4 km</quantity>
NNW of Manantenina
</locationDeviation>
;
<geoCoordinate id="DCCA37DBFFC3FFF1FF170AB3FEF9DDBD" box="[189,300,663,691]" degrees="14" direction="south" minutes="26" orientation="latitude" pageId="59" pageNumber="60" precision="925" value="-14.433333">14°26′ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFC3FFF1FE970AB3FE7ADDBD" box="[317,431,663,691]" degrees="49" direction="east" minutes="45" orientation="longitude" pageId="59" pageNumber="60" precision="925" value="49.75">49°45′ E</geoCoordinate>
;
<elevation id="32D3B62FFFC3FFF1FE6A0ABDFD8DDDBD" box="[448,600,665,691]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="59" pageNumber="60" unit="m" value="700.0">
<quantity id="7E06FCF9FFC3FFF1FE6A0ABDFDC5DDBD" box="[448,528,665,691]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="59" pageNumber="60" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFC3FFF1FDC10ABDFC9DDDBD" box="[619,840,665,691]" pageId="59" pageNumber="60" value="1993-11-10" valueMax="1993-11-16" valueMin="1993-11-10">
<collectingDate id="DD048E34FFC3FFF1FDC10ABDFC9DDDBD" box="[619,840,665,691]" pageId="59" pageNumber="60" value="1993-11-10" valueMax="1993-11-16" valueMin="1993-11-10">10–16 Nov. 1993</collectingDate>
</date>
;
<collectorName id="140B34CAFFC3FFF1FCF00ABDFC20DDBD" box="[858,1013,665,691]" pageId="59" pageNumber="60">C. Griswold</collectorName>
,
<collectorName id="140B34CAFFC3FFF1FBAC0ABDFB6CDDBD" box="[1030,1209,665,691]" pageId="59" pageNumber="60">J. Coddington</collectorName>
,
<collectorName id="140B34CAFFC3FFF1FB630ABDFA9BDDBD" box="[1225,1358,665,691]" pageId="59" pageNumber="60">N. Scharff</collectorName>
,
<collectorName id="140B34CAFFC3FFF1FAF40ABDFECEDDD8" pageId="59" pageNumber="60">S. Larcher</collectorName>
,
<collectorName id="140B34CAFFC3FFF1FE820A99FDE4DDD8" box="[296,561,700,727]" pageId="59" pageNumber="60">R. Andriamasimanana</collectorName>
leg.; general collecting; CEG008; [habitus and male palp illustrated];
</materialsCitation>
</paragraph>
<paragraph id="B941511CFFC3FFF1FF150AFBFE23DDF7" blockId="59.[189,1399,595,761]" box="[191,502,735,761]" pageId="59" pageNumber="60">
CASENT_9006231;
<collectionCode id="DFEFC9D9FFC3FFF1FE1F0AFBFE27DDF7" box="[437,498,735,761]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="60" type="Museum">CAS</collectionCode>
.
</paragraph>
<paragraph id="B941511CFFC3FFF1FF400B05FEB2DC35" blockId="59.[189,1399,801,968]" box="[234,359,801,827]" pageId="59" pageNumber="60">
<emphasis id="8B8A8D0EFFC3FFF1FF400B05FEB2DC35" bold="true" box="[234,359,801,827]" pageId="59" pageNumber="60">
<typeStatus id="6645EFBEFFC3FFF1FF400B05FEB2DC35" box="[234,359,801,827]" pageId="59" pageNumber="60" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFC3FFF1FF170B60FAA2DCC6" blockId="59.[189,1399,801,968]" pageId="59" pageNumber="60">
<materialsCitation id="09965B41FFC3FFF1FF170B60FAB3DC50" box="[189,1382,836,863]" collectingDate="1993-11-10" collectingDateMax="1993-11-16" collectingDateMin="1993-11-10" collectionCode="CAS" collectorName="C. Griswold &amp; J. Coddington &amp; N. Scharff &amp; S. Larcher &amp; R. Andriamasimanana" country="Madagascar" elevation="700" latitude="-14.433333" location="Marojejy Reserve" longLatPrecision="1288" longitude="49.75" pageId="59" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="paratype">
<collectingCountry id="C1E9118CFFC3FFF1FF170B60FE58DC50" box="[189,397,836,862]" name="Madagascar" pageId="59" pageNumber="60">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFC3FFF1FE0C0B60FD63DC50" bold="true" box="[422,694,836,862]" pageId="59" pageNumber="60">
<collectingRegion id="7B3A9FFEFFC3FFF1FE0C0B60FD63DC50" box="[422,694,836,862]" country="Madagascar" name="Antsiranana" pageId="59" pageNumber="60">Antsiranana Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFC3FFF1FD670B60FD2EDC51" box="[717,763,836,863]" count="1" pageId="59" pageNumber="60" type="male">1 ♂</specimenCount>
; same data as for holotype; CASENT_9006231;
<collectionCode id="DFEFC9D9FFC3FFF1FA870B60FAB3DC50" box="[1325,1382,836,862]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="59" pageNumber="60" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFC3FFF1FF170B43FCA2DC8F" box="[189,887,871,898]" collectingDate="1993-11-10" collectingDateMax="1993-11-16" collectingDateMin="1993-11-10" collectionCode="USNM" collectorName="C. Griswold &amp; J. Coddington &amp; N. Scharff &amp; S. Larcher &amp; R. Andriamasimanana" country="Madagascar" elevation="700" latitude="-14.433333" location="Marojejy Reserve" longLatPrecision="1288" longitude="49.75" pageId="59" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="paratype">
<specimenCount id="AFF89A95FFC3FFF1FF170B43FF3CDC8C" box="[189,233,871,898]" count="1" pageId="59" pageNumber="60" type="male">1 ♂</specimenCount>
; same data as for preceding; CASENT_9082258;
<collectionCode id="DFEFC9D9FFC3FFF1FC890B43FCA2DC8F" box="[803,887,871,897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="59" pageNumber="60" type="Museum">USNM</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFC3FFF1FC260B43FC91DCAA" collectingDate="1993-11-10" collectingDateMax="1993-11-16" collectingDateMin="1993-11-10" collectionCode="USNM" collectorName="S. Larcher" country="Madagascar" elevation="700" latitude="-14.433333" location="Marojejy Reserve" longLatPrecision="1288" longitude="49.75" pageId="59" pageNumber="60" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana" typeStatus="paratype">
<specimenCount id="AFF89A95FFC3FFF1FC260B43FC62DC8C" box="[908,951,871,898]" count="1" pageId="59" pageNumber="60" type="female">1 ♀</specimenCount>
; same data as for preceding;
<collectorName id="140B34CAFFC3FFF1FAAA0B43FAA3DC8F" box="[1280,1398,871,897]" pageId="59" pageNumber="60">S. Larcher</collectorName>
leg.; [endogyne drawn]; CASENT_9006230;
<collectionCode id="DFEFC9D9FFC3FFF1FD400BAEFC91DCAA" box="[746,836,906,932]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="59" pageNumber="60" type="Museum">USNM</collectionCode>
</materialsCitation>
. –
<materialsCitation id="09965B41FFC3FFF1FCC70BAEFAA6DCC6" collectorName="Mar" country="Madagascar" county="Parc National Masoala" elevation="955" latitude="-15.572222" location="Ambanizana" longLatPrecision="21" longitude="50.006943" municipality="Ambohitsitondroina Mt." pageId="59" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
<emphasis id="8B8A8D0EFFC3FFF1FCC70BAEFBA6DCAA" bold="true" box="[877,1139,906,932]" pageId="59" pageNumber="60">
<collectingRegion id="7B3A9FFEFFC3FFF1FCC70BAEFBA6DCAA" box="[877,1139,906,932]" country="Madagascar" name="Toamasina" pageId="59" pageNumber="60">Toamasina Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFC3FFF1FB3B0BAEFB15DCAB" box="[1169,1216,906,933]" count="1" pageId="59" pageNumber="60" type="male">1 ♂</specimenCount>
;
<collectingCounty id="50202990FFC3FFF1FB7A0BAFFEF6DCC9" pageId="59" pageNumber="60">Parc National Masoala</collectingCounty>
,
<collectingMunicipality id="5925CB66FFC3FFF1FE850B89FD85DCC6" box="[303,592,941,968]" pageId="59" pageNumber="60">Ambohitsitondroina Mt.</collectingMunicipality>
,
<location id="BC2107C7FFC3FFF1FDF60B89FD21DCC9" LSID="urn:lsid:plazi:treatment:3157E00AFFC2FFF5FDFF0DF7FC8FDCA1:BC2107C7FFC3FFF1FDF60B89FD21DCC9" box="[604,756,941,967]" country="Madagascar" county="Parc National Masoala" latitude="-15.572222" longLatPrecision="21" longitude="50.006943" municipality="Ambohitsitondroina Mt." name="Ambanizana" pageId="59" pageNumber="60" stateProvince="Toamasina">Ambanizana</location>
;
<geoCoordinate id="DCCA37DBFFC3FFF1FCA80B88FC46DCC6" box="[770,915,940,968]" degrees="15" direction="south" minutes="34" orientation="latitude" pageId="59" pageNumber="60" precision="15" seconds="20" value="-15.572222">15°34′20″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFC3FFF1FC0B0B88FBE0DCC6" box="[929,1077,940,968]" degrees="50" direction="east" minutes="00" orientation="longitude" pageId="59" pageNumber="60" precision="15" seconds="25" value="50.006943">50°00′25″ E</geoCoordinate>
;
<elevation id="32D3B62FFFC3FFF1FBE90B8AFACCDCC9" box="[1091,1305,941,968]" metricMagnitude="2" metricUnit="m" metricValue="9.55" metricValueMax="10.1" metricValueMin="9.0" pageId="59" pageNumber="60" unit="m" value="955.0" valueMax="1010.0" valueMin="900.0">
<quantity id="7E06FCF9FFC3FFF1FBE90B8AFB0DDCC6" box="[1091,1240,942,968]" metricMagnitude="2" metricUnit="m" metricValue="9.55" metricValueMax="10.1" metricValueMin="9.0" pageId="59" pageNumber="60" unit="m" value="955.0" valueMax="1010.0" valueMin="900.0">900–1010 m</quantity>
a.s.l.
</elevation>
; 5
<collectorName id="140B34CAFFC3FFF1FAEA0B8AFAA6DCC6" box="[1344,1395,942,968]" pageId="59" pageNumber="60">Mar</collectorName>
</materialsCitation>
.
</paragraph>
<caption id="ED810194FFC3FFF1FF170F42FDDDD8E7" ID-DOI="http://doi.org/10.5281/zenodo.14001847" ID-Zenodo-Dep="14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="59" pageNumber="60" startId="59.[189,234,1894,1920]" targetBox="[189,1398,1046,1855]" targetPageId="59">
<paragraph id="B941511CFFC3FFF1FF170F42FDDDD8E7" blockId="59.[187,1399,1894,2025]" pageId="59" pageNumber="60">
<emphasis id="8B8A8D0EFFC3FFF1FF170F42FEF0D88E" bold="true" box="[189,293,1894,1920]" pageId="59" pageNumber="60">Fig. 24.</emphasis>
<taxonomicName id="7EFE2A9FFFC3FFF1FE9E0F42FDA5D88E" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[308,624,1894,1920]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="silvadavilae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC3FFF1FE9E0F42FDA5D88E" box="[308,624,1894,1920]" italics="true" pageId="59" pageNumber="60">Zorodictyna silvadavilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC3FFF1FD2A0F42FD3CD88E" box="[640,745,1894,1920]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
, male and female habitus; A–B = male holotype (CASENT_9006231; CAS); C–D = female paratype (FMNH-0000-056-622; FMNH).
<emphasis id="8B8A8D0EFFC3FFF1FB0A0FADFB02D8AD" bold="true" box="[1184,1239,1929,1955]" pageId="59" pageNumber="60">A, C</emphasis>
. Dorsal view.
<emphasis id="8B8A8D0EFFC3FFF1FF110F88FF24D8C8" bold="true" box="[187,241,1964,1990]" pageId="59" pageNumber="60">B, D</emphasis>
. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme.
</paragraph>
</caption>
<paragraph id="B941511CFFC4FFF6FF17092DFF29DED9" blockId="60.[187,1399,265,471]" pageId="60" pageNumber="61">
2003;
<collectorName id="140B34CAFFC4FFF6FEA2092DFE3DDE2D" box="[264,488,265,291]" pageId="60" pageNumber="61">Diana Silva Dávila</collectorName>
leg.; raking tree trunks; montane rainforest; DNA vouchers ARACG000047; DSD0024; CASENT_9014998;
<collectionCode id="DFEFC9D9FFC4FFF6FD9A0909FDBDDE49" box="[560,616,301,327]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="60" pageNumber="61" type="Museum">CAS</collectionCode>
•
<materialsCitation id="09965B41FFC4FFF6FDD40909FACCDE49" box="[638,1305,301,328]" collectionCode="CAS" pageId="60" pageNumber="61" specimenCount="1" specimenCount-female="1">
<specimenCount id="AFF89A95FFC4FFF6FDD40909FD7DDE46" box="[638,680,301,328]" count="1" pageId="60" pageNumber="61" type="female">1 ♀</specimenCount>
; same data as for preceding; CASENT_9015400;
<collectionCode id="DFEFC9D9FFC4FFF6FB4B0909FACCDE49" box="[1249,1305,301,327]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="60" pageNumber="61" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFC4FFF6FA850909FD89DE9E" collectingDate="2002-02" collectionCode="FMNH" collectorName="S. Goodman" elevation="810" location="Parc National de Marojejy" pageId="60" pageNumber="61" specimenCount="2" specimenCount-female="2">
<specimenCount id="AFF89A95FFC4FFF6FA850909FAA5DE46" box="[1327,1392,301,328]" count="2" pageId="60" pageNumber="61" type="female">2 ♀♀</specimenCount>
;
<location id="BC2107C7FFC4FFF6FF170976FE20DE65" LSID="urn:lsid:plazi:treatment:3157E00AFFC2FFF5FDFF0DF7FC8FDCA1:BC2107C7FFC4FFF6FF170976FE20DE65" box="[189,501,337,364]" name="Parc National de Marojejy" pageId="60" pageNumber="61">Parc National de Marojejy</location>
, NW;
<elevation id="32D3B62FFFC4FFF6FDEA0975FD1EDE65" box="[576,715,337,364]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="60" pageNumber="61" unit="m" value="810.0">
<quantity id="7E06FCF9FFC4FFF6FDEA0975FD5EDE62" box="[576,651,337,364]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="60" pageNumber="61" unit="m" value="810.0">810 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFC4FFF6FD730975FC87DE65" box="[729,850,337,363]" pageId="60" pageNumber="61" value="2002-02">
<collectingDate id="DD048E34FFC4FFF6FD730975FC87DE65" box="[729,850,337,363]" pageId="60" pageNumber="61" value="2002-02">Feb. 2002</collectingDate>
</date>
;
<collectorName id="140B34CAFFC4FFF6FCF40975FC24DE65" box="[862,1009,337,363]" pageId="60" pageNumber="61">S. Goodman</collectorName>
leg.;
<collectingMethod id="60BF290BFFC4FFF6FB990975FBA8DE65" box="[1075,1149,337,363]" pageId="60" pageNumber="61">pitfall</collectingMethod>
; [habitus illustrated]; FMNH-INS 0000-056-622;
<collectionCode id="DFEFC9D9FFC4FFF6FDAD0952FD89DE9E" box="[519,604,374,400]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="60" pageNumber="61" type="Museum">FMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFC4FFF6FDD90951FF2DDED9" collectingDate="2003-03-01" collectionCode="CAS" collectorName="D. Andriamalala &amp; D. Silva Davila" elevation="775" latitude="-15.574972" location="Mt. Ambanizana" longLatPrecision="1" longitude="50.003418" pageId="60" pageNumber="61" specimenCount="1" specimenCount-female="1">
<specimenCount id="AFF89A95FFC4FFF6FDD90951FD4ADE9E" box="[627,671,373,400]" count="1" pageId="60" pageNumber="61" type="female">1 ♀</specimenCount>
;
<location id="BC2107C7FFC4FFF6FD070952FCA1DE81" LSID="urn:lsid:plazi:treatment:3157E00AFFC2FFF5FDFF0DF7FC8FDCA1:BC2107C7FFC4FFF6FD070952FCA1DE81" box="[685,884,373,400]" latitude="-15.574972" longLatPrecision="1" longitude="50.003418" name="Mt. Ambanizana" pageId="60" pageNumber="61">Mt. Ambanizana</location>
;
<geoCoordinate id="DCCA37DBFFC4FFF6FCD50950FBF1DE81" box="[895,1060,372,400]" degrees="15" direction="south" minutes="34" orientation="latitude" pageId="60" pageNumber="61" precision="1" seconds="29.9" value="-15.574972">15°34′29.9″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFC4FFF6FB9A0950FB32DE9E" box="[1072,1255,372,400]" degrees="050" direction="east" minutes="00" orientation="longitude" pageId="60" pageNumber="61" precision="1" seconds="12.3" value="50.003418">050°00′12.3″ E</geoCoordinate>
;
<elevation id="32D3B62FFFC4FFF6FB580951FF20DEBD" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="60" pageNumber="61" unit="m" value="775.0" valueMax="800.0" valueMin="750.0">
<quantity id="7E06FCF9FFC4FFF6FB580951FAA2DE9E" box="[1266,1399,373,400]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="60" pageNumber="61" unit="m" value="775.0" valueMax="800.0" valueMin="750.0">750–800 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFC4FFF6FEAA09BDFE58DEBD" box="[256,397,409,436]" pageId="60" pageNumber="61" value="2003-03-01">
<collectingDate id="DD048E34FFC4FFF6FEAA09BDFE58DEBD" box="[256,397,409,436]" pageId="60" pageNumber="61" value="2003-03-01">1 Mar. 2003</collectingDate>
</date>
;
<collectorName id="140B34CAFFC4FFF6FE3D09BEFD82DEBD" box="[407,599,409,436]" pageId="60" pageNumber="61">D. Andriamalala</collectorName>
,
<collectorName id="140B34CAFFC4FFF6FDCB09BEFCC4DEBD" box="[609,785,409,436]" pageId="60" pageNumber="61">D. Silva Dávila</collectorName>
<emphasis id="8B8A8D0EFFC4FFF6FCBF09BEFC9BDEBD" box="[789,846,409,435]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
leg.; rainforest; DSD0017; CASENT_9015400;
<collectionCode id="DFEFC9D9FFC4FFF6FF110999FF2DDED9" box="[187,248,445,471]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="60" pageNumber="61" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="B941511CFFC4FFF6FF1709DBFE2CDD17" blockId="60.[189,505,511,537]" box="[189,505,511,537]" pageId="60" pageNumber="61">
<heading id="E209E670FFC4FFF6FF1709DBFE2CDD17" bold="true" box="[189,505,511,537]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="8B8A8D0EFFC4FFF6FF1709DBFE2CDD17" bold="true" box="[189,505,511,537]" pageId="60" pageNumber="61">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC4FFF6FF170A0BFDFEDD9C" blockId="60.[189,1398,559,658]" pageId="60" pageNumber="61">
<materialsCitation id="09965B41FFC4FFF6FF170A0BFDF2DD9C" collectingDate="1996-10-24" collectingDateMax="1996-11-02" collectingDateMin="1996-10-24" collectionCode="FMNH" collectorName="R. N. I. Marojejy &amp; S. Goodman &amp; Mad" country="Madagascar" elevation="1225" latitude="-14.4366665" location="Antranohofa" longLatPrecision="128" longitude="49.741665" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1">
<collectingCountry id="C1E9118CFFC4FFF6FF170A0BFE47DD47" box="[189,402,559,585]" name="Madagascar" pageId="60" pageNumber="61">MADAGASCAR</collectingCountry>
•
<specimenCount id="AFF89A95FFC4FFF6FE1A0A0BFE37DD44" box="[432,482,559,586]" count="1" pageId="60" pageNumber="61" type="male">1 ♂</specimenCount>
;
<collectorName id="140B34CAFFC4FFF6FE5B0A14FD63DD47" box="[497,694,559,586]" pageId="60" pageNumber="61">R.N.I. Marojejy</collectorName>
,
<locationDeviation id="A8F3483EFFC4FFF6FD690A0BFBCDDD47" box="[707,1048,559,586]" location="Manantenina" pageId="60" pageNumber="61">
<quantity id="7E06FCF9FFC4FFF6FD690A0BFCC5DD47" box="[707,784,559,585]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="60" pageNumber="61" unit="km" value="11.0">11 km</quantity>
NW of Manantenina
</locationDeviation>
,
<location id="BC2107C7FFC4FFF6FB8F0A0BFB14DD47" LSID="urn:lsid:plazi:treatment:3157E00AFFC2FFF5FDFF0DF7FC8FDCA1:BC2107C7FFC4FFF6FB8F0A0BFB14DD47" box="[1061,1217,559,585]" country="Madagascar" latitude="-14.4366665" longLatPrecision="128" longitude="49.741665" name="Antranohofa" pageId="60" pageNumber="61">Antranohofa</location>
;
<elevation id="32D3B62FFFC4FFF6FB650A0BFAA5DD47" box="[1231,1392,559,586]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="60" pageNumber="61" unit="m" value="1225.0">
<quantity id="7E06FCF9FFC4FFF6FB650A0BFAF9DD44" box="[1231,1324,559,586]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="60" pageNumber="61" unit="m" value="1225.0">1225 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="DCCA37DBFFC4FFF6FF170A75FEE9DD63" box="[189,316,593,621]" degrees="14" direction="south" minutes="26.2" orientation="latitude" pageId="60" pageNumber="61" precision="92" value="-14.4366665">14°26.2′ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFC4FFF6FEE10A75FE18DD60" box="[331,461,593,622]" degrees="49" direction="east" minutes="44.5" orientation="longitude" pageId="60" pageNumber="61" precision="92" value="49.741665">49°44.5′ E</geoCoordinate>
;
<date id="CD4077DCFFC4FFF6FE760A77FD09DD63" box="[476,732,595,622]" pageId="60" pageNumber="61" value="1996-10-24" valueMax="1996-11-02" valueMin="1996-10-24">
<collectingDate id="DD048E34FFC4FFF6FE760A77FD09DD63" box="[476,732,595,622]" pageId="60" pageNumber="61" value="1996-10-24" valueMax="1996-11-02" valueMin="1996-10-24">24 Oct.–2 Nov. 1996</collectingDate>
</date>
;
<collectorName id="140B34CAFFC4FFF6FD400A77FC55DD63" box="[746,896,595,621]" pageId="60" pageNumber="61">S. Goodman</collectorName>
leg.; undisturbed mossy forest;
<collectingMethod id="60BF290BFFC4FFF6FAAF0A77FA8DDD63" box="[1285,1368,595,621]" pageId="60" pageNumber="61">pitfalls</collectingMethod>
at camp #3;
<collectorName id="140B34CAFFC4FFF6FE9B0A53FEBFDD9F" box="[305,362,631,657]" pageId="60" pageNumber="61">Mad</collectorName>
1996#7;
<collectionCode id="DFEFC9D9FFC4FFF6FE640A5CFDF2DD9C" box="[462,551,632,658]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="60" pageNumber="61" type="Museum">FMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFC4FFF5FF170A9DFBE5DC12" lastPageId="63" lastPageNumber="64" pageId="60" pageNumber="61" type="description">
<paragraph id="B941511CFFC4FFF6FF170A9DFE9BDDDD" blockId="60.[189,334,697,723]" box="[189,334,697,723]" pageId="60" pageNumber="61">
<heading id="E209E670FFC4FFF6FF170A9DFE9BDDDD" bold="true" box="[189,334,697,723]" fontSize="11" level="3" pageId="60" pageNumber="61" reason="3">
<emphasis id="8B8A8D0EFFC4FFF6FF170A9DFE9BDDDD" bold="true" box="[189,334,697,723]" pageId="60" pageNumber="61">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC4FFF6FF400ACDFB85DC0D" blockId="60.[189,1189,745,808]" box="[234,1104,745,771]" pageId="60" pageNumber="61">
<emphasis id="8B8A8D0EFFC4FFF6FF400ACDFE48DC0D" bold="true" box="[234,413,745,771]" pageId="60" pageNumber="61">
Male
<typeStatus id="6645EFBEFFC4FFF6FE9B0ACDFE48DC0D" box="[305,413,745,771]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
</emphasis>
(CASENT_9006231; CAS; except as otherwise indicated)
</paragraph>
<paragraph id="B941511CFFC4FFF6FF170B29FB70DC26" blockId="60.[189,1189,745,808]" box="[189,1189,781,808]" pageId="60" pageNumber="61">MEASUREMENTS. Total length: 17.60. Carapace: length 10.50, width 8.30, height 4.50.</paragraph>
<paragraph id="B941511CFFC4FFF6FF170B6BFBFBDC83" blockId="60.[189,1398,847,910]" pageId="60" pageNumber="61">
COLORATION (
<figureCitation id="21C54D99FFC4FFF6FECF0B6BFE22DC67" box="[357,503,847,874]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; A–B = male holotype (CASENT_9006231; CAS); C–D = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="60" pageNumber="61">Fig. 24A–B</figureCitation>
). Dorsal and lateral of cephalothorax, abdomen and legs unicolorous grey-brown, venter of sternum, mouthparts, coxae and trochanters yellow white.
</paragraph>
<paragraph id="B941511CFFC4FFF6FF170B92FB8ADCC1" blockId="60.[189,1119,949,976]" box="[189,1119,949,976]" pageId="60" pageNumber="61">
STERNUM (
<figureCitation id="21C54D99FFC4FFF6FE970B91FE7DDCDE" box="[317,424,949,976]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; A–B = male holotype (CASENT_9006231; CAS); C–D = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="60" pageNumber="61">Fig. 24B</figureCitation>
). Oval with slightly sinuous margins; 3.70 wide, 4.50 long.
</paragraph>
<paragraph id="B941511CFFC4FFF6FF170BDCFDF7DB1C" blockId="60.[189,546,1016,1042]" box="[189,546,1016,1042]" pageId="60" pageNumber="61">CHILUM. 0.18 high, 0.63 wide.</paragraph>
<paragraph id="B941511CFFC4FFF6FF170C1EFCCADB77" blockId="60.[188,1399,1082,1145]" pageId="60" pageNumber="61">EYES. ALE 0.28; AME 0.23; PLE 0.28; PME 0.25; ALE–AME 0.30; AME–AME 0.40; PLE–PME 0.35; PME–PME 0.15; MOQ: AW 0.58, PW 0.73, L 0.65.</paragraph>
<paragraph id="B941511CFFC4FFF6FF170C85FDCDDBB5" blockId="60.[189,536,1184,1211]" box="[189,536,1184,1211]" pageId="60" pageNumber="61">CLYPEUS. With ten long setae.</paragraph>
<paragraph id="B941511CFFC4FFF6FF170CC7FADFDA2F" blockId="60.[189,1398,1251,1313]" pageId="60" pageNumber="61">LEGS. Trochanters I–III shallowly notched, IV entire. Unpaired claws smooth, rudimentary on legs III and IV. Scopulae beneath all tarsi, apical half of metatarsus I and apical third of metatarsus II.</paragraph>
<paragraph id="B941511CFFC4FFF6FF170D6EFC84DA6D" blockId="60.[189,849,1353,1380]" box="[189,849,1353,1380]" pageId="60" pageNumber="61">
Leg measurements (
<typeStatus id="6645EFBEFFC4FFF6FE030D6DFDC5DA6D" box="[425,528,1353,1379]" pageId="60" pageNumber="61" type="paratype">paratype</typeStatus>
CASENT_9014998; CAS)
</paragraph>
<paragraph id="B941511CFFC4FFF6FEA00D50FCC1D939" pageId="60" pageNumber="61">
<table id="CBFEA3BCFFC40035FF610D57FCC1D939" box="[203,788,1395,1591]" gridcols="7" gridrows="5" pageId="60" pageNumber="61" rowsContinueIn="60.[203,946,1665,1850]">
<tr id="07CE535EFFC40035FF610D57FCC1DA80" box="[203,788,1395,1422]" gridrow="0" pageId="60" pageNumber="61" rowspan-0="1">
<th id="441F3A22FFC40035FEA00D57FE99DA80" box="[266,332,1395,1422]" gridcol="1" gridrow="0" pageId="60" pageNumber="61">F</th>
<th id="441F3A22FFC40035FEC50D57FE77DA80" box="[367,418,1395,1422]" gridcol="2" gridrow="0" pageId="60" pageNumber="61">P</th>
<th id="441F3A22FFC40035FE6E0D57FE22DA80" box="[452,503,1395,1422]" gridcol="3" gridrow="0" pageId="60" pageNumber="61">T</th>
<th id="441F3A22FFC40035FDB20D57FD8FDA80" box="[536,602,1395,1422]" gridcol="4" gridrow="0" pageId="60" pageNumber="61">Mt</th>
<th id="441F3A22FFC40035FDD70D57FD65DA80" box="[637,688,1395,1422]" gridcol="5" gridrow="0" pageId="60" pageNumber="61">t</th>
<th id="441F3A22FFC40035FD780D57FCC1DA80" box="[722,788,1395,1422]" gridcol="6" gridrow="0" pageId="60" pageNumber="61">Total</th>
</tr>
<tr id="07CE535EFFC40035FF610DB8FCC1DACE" box="[203,788,1436,1472]" gridrow="1" pageId="60" pageNumber="61">
<th id="441F3A22FFC40035FF610DB8FF3FDACE" box="[203,234,1436,1472]" gridcol="0" gridrow="1" pageId="60" pageNumber="61">I</th>
<td id="441F3A22FFC40035FEA00DB8FE99DACE" box="[266,332,1436,1472]" gridcol="1" gridrow="1" pageId="60" pageNumber="61">9.70</td>
<td id="441F3A22FFC40035FEC50DB8FE77DACE" box="[367,418,1436,1472]" gridcol="2" gridrow="1" pageId="60" pageNumber="61">3.80</td>
<td id="441F3A22FFC40035FE6E0DB8FE22DACE" box="[452,503,1436,1472]" gridcol="3" gridrow="1" pageId="60" pageNumber="61">–</td>
<td id="441F3A22FFC40035FDB20DB8FD8FDACE" box="[536,602,1436,1472]" gridcol="4" gridrow="1" pageId="60" pageNumber="61">–</td>
<td id="441F3A22FFC40035FDD70DB8FD65DACE" box="[637,688,1436,1472]" gridcol="5" gridrow="1" pageId="60" pageNumber="61">–</td>
<td id="441F3A22FFC40035FD780DB8FCC1DACE" box="[722,788,1436,1472]" gridcol="6" gridrow="1" pageId="60" pageNumber="61">–</td>
</tr>
<tr id="07CE535EFFC40035FF610DE7FCC1DAE9" box="[203,788,1475,1511]" gridrow="2" pageId="60" pageNumber="61">
<th id="441F3A22FFC40035FF610DE7FF3FDAE9" box="[203,234,1475,1511]" gridcol="0" gridrow="2" pageId="60" pageNumber="61">II</th>
<td id="441F3A22FFC40035FEA00DE7FE99DAE9" box="[266,332,1475,1511]" gridcol="1" gridrow="2" pageId="60" pageNumber="61">8.50</td>
<td id="441F3A22FFC40035FEC50DE7FE77DAE9" box="[367,418,1475,1511]" gridcol="2" gridrow="2" pageId="60" pageNumber="61">3.70</td>
<td id="441F3A22FFC40035FE6E0DE7FE22DAE9" box="[452,503,1475,1511]" gridcol="3" gridrow="2" pageId="60" pageNumber="61">7.90</td>
<td id="441F3A22FFC40035FDB20DE7FD8FDAE9" box="[536,602,1475,1511]" gridcol="4" gridrow="2" pageId="60" pageNumber="61">8.00</td>
<td id="441F3A22FFC40035FDD70DE7FD65DAE9" box="[637,688,1475,1511]" gridcol="5" gridrow="2" pageId="60" pageNumber="61">3.90</td>
<td id="441F3A22FFC40035FD780DE7FCC1DAE9" box="[722,788,1475,1511]" gridcol="6" gridrow="2" pageId="60" pageNumber="61">32.00</td>
</tr>
<tr id="07CE535EFFC40035FF610DCFFCC1D901" box="[203,788,1515,1551]" gridrow="3" pageId="60" pageNumber="61">
<th id="441F3A22FFC40035FF610DCFFF3FD901" box="[203,234,1515,1551]" gridcol="0" gridrow="3" pageId="60" pageNumber="61">III</th>
<td id="441F3A22FFC40035FEA00DCFFE99D901" box="[266,332,1515,1551]" gridcol="1" gridrow="3" pageId="60" pageNumber="61">7.50</td>
<td id="441F3A22FFC40035FEC50DCFFE77D901" box="[367,418,1515,1551]" gridcol="2" gridrow="3" pageId="60" pageNumber="61">3.40</td>
<td id="441F3A22FFC40035FE6E0DCFFE22D901" box="[452,503,1515,1551]" gridcol="3" gridrow="3" pageId="60" pageNumber="61">6.00</td>
<td id="441F3A22FFC40035FDB20DCFFD8FD901" box="[536,602,1515,1551]" gridcol="4" gridrow="3" pageId="60" pageNumber="61">7.70</td>
<td id="441F3A22FFC40035FDD70DCFFD65D901" box="[637,688,1515,1551]" gridcol="5" gridrow="3" pageId="60" pageNumber="61">3.60</td>
<td id="441F3A22FFC40035FD780DCFFCC1D901" box="[722,788,1515,1551]" gridcol="6" gridrow="3" pageId="60" pageNumber="61">28.20</td>
</tr>
<tr id="07CE535EFFC40035FF610E37FCC1D939" box="[203,788,1555,1591]" gridrow="4" pageId="60" pageNumber="61">
<th id="441F3A22FFC40035FF610E37FF3FD939" box="[203,234,1555,1591]" gridcol="0" gridrow="4" pageId="60" pageNumber="61">IV</th>
<td id="441F3A22FFC40035FEA00E37FE99D939" box="[266,332,1555,1591]" gridcol="1" gridrow="4" pageId="60" pageNumber="61">10.00</td>
<td id="441F3A22FFC40035FEC50E37FE77D939" box="[367,418,1555,1591]" gridcol="2" gridrow="4" pageId="60" pageNumber="61">3.20</td>
<td id="441F3A22FFC40035FE6E0E37FE22D939" box="[452,503,1555,1591]" gridcol="3" gridrow="4" pageId="60" pageNumber="61">8.50</td>
<td id="441F3A22FFC40035FDB20E37FD8FD939" box="[536,602,1555,1591]" gridcol="4" gridrow="4" pageId="60" pageNumber="61">12.60</td>
<td id="441F3A22FFC40035FDD70E37FD65D939" box="[637,688,1555,1591]" gridcol="5" gridrow="4" pageId="60" pageNumber="61">4.70</td>
<td id="441F3A22FFC40035FD780E37FCC1D939" box="[722,788,1555,1591]" gridcol="6" gridrow="4" pageId="60" pageNumber="61">39.00</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFC4FFF6FF170E72FD31D97E" blockId="60.[189,740,1622,1648]" box="[189,740,1622,1648]" pageId="60" pageNumber="61">
Spination (
<typeStatus id="6645EFBEFFC4FFF6FE970E72FE71D97E" box="[317,420,1622,1648]" pageId="60" pageNumber="61" type="paratype">paratype</typeStatus>
CASENT_9014998; CAS)
</paragraph>
<paragraph id="B941511CFFC4FFF6FEA00EA5FC67D834" pageId="60" pageNumber="61">
<table id="CBFEA3BCFFC40035FF610EA5FC67D834" box="[203,946,1665,1850]" gridcols="5" gridrows="5" pageId="60" pageNumber="61" rowsContinueFrom="60.[203,788,1395,1591]">
<tr id="07CE535EFFC40035FF610EA5FC67D995" box="[203,946,1665,1691]" gridrow="0" pageId="60" pageNumber="61" rowspan-0="1">
<th id="441F3A22FFC40035FEA00EA5FEBBD995" box="[266,366,1665,1691]" gridcol="1" gridrow="0" pageId="60" pageNumber="61">F</th>
<th id="441F3A22FFC40035FE390EA5FE0CD995" box="[403,473,1665,1691]" gridcol="2" gridrow="0" pageId="60" pageNumber="61">P</th>
<th id="441F3A22FFC40035FE570EA5FD12D995" box="[509,711,1665,1691]" gridcol="3" gridrow="0" pageId="60" pageNumber="61">T</th>
<th id="441F3A22FFC40035FCAE0EA5FC67D995" box="[772,946,1665,1691]" gridcol="4" gridrow="0" pageId="60" pageNumber="61">Mt</th>
</tr>
<tr id="07CE535EFFC40035FF610E8CFC67D9C2" box="[203,946,1704,1740]" gridrow="1" pageId="60" pageNumber="61">
<th id="441F3A22FFC40035FF610E8CFF3FD9C2" box="[203,234,1704,1740]" gridcol="0" gridrow="1" pageId="60" pageNumber="61">I</th>
<td id="441F3A22FFC40035FEA00E8CFEBBD9C2" box="[266,366,1704,1740]" gridcol="1" gridrow="1" pageId="60" pageNumber="61">pl2d3rl4</td>
<td id="441F3A22FFC40035FE390E8CFE0CD9C2" box="[403,473,1704,1740]" gridcol="2" gridrow="1" pageId="60" pageNumber="61">pl1rl1</td>
<td id="441F3A22FFC40035FE570E8CFD12D9C2" box="[509,711,1704,1740]" gridcol="3" gridrow="1" pageId="60" pageNumber="61">–</td>
<td id="441F3A22FFC40035FCAE0E8CFC67D9C2" box="[772,946,1704,1740]" gridcol="4" gridrow="1" pageId="60" pageNumber="61">–</td>
</tr>
<tr id="07CE535EFFC40035FF610EF4FC67D9E5" box="[203,946,1744,1771]" gridrow="2" pageId="60" pageNumber="61">
<th id="441F3A22FFC40035FF610EF4FF3FD9E5" box="[203,234,1744,1771]" gridcol="0" gridrow="2" pageId="60" pageNumber="61">II</th>
<td id="441F3A22FFC40035FEA00EF4FEBBD9E5" box="[266,366,1744,1771]" gridcol="1" gridrow="2" pageId="60" pageNumber="61">pl3d3rl4</td>
<td id="441F3A22FFC40035FE390EF4FE0CD9E5" box="[403,473,1744,1771]" gridcol="2" gridrow="2" pageId="60" pageNumber="61">pl1rl1</td>
<td id="441F3A22FFC40035FE570EF4FD12D9E5" box="[509,711,1744,1771]" gridcol="3" gridrow="2" pageId="60" pageNumber="61">pl2d2rl3v2-2-2-2</td>
<td id="441F3A22FFC40035FCAE0EF4FC67D9E5" box="[772,946,1744,1771]" gridcol="4" gridrow="2" pageId="60" pageNumber="61">pl4rl3v2-3-2</td>
</tr>
<tr id="07CE535EFFC40035FF610EDCFC67D81D" box="[203,946,1784,1811]" gridrow="3" pageId="60" pageNumber="61">
<th id="441F3A22FFC40035FF610EDCFF3FD81D" box="[203,234,1784,1811]" gridcol="0" gridrow="3" pageId="60" pageNumber="61">III</th>
<td id="441F3A22FFC40035FEA00EDCFEBBD81D" box="[266,366,1784,1811]" gridcol="1" gridrow="3" pageId="60" pageNumber="61">pl3d3rl4</td>
<td id="441F3A22FFC40035FE390EDCFE0CD81D" box="[403,473,1784,1811]" gridcol="2" gridrow="3" pageId="60" pageNumber="61">pl1rl1</td>
<td id="441F3A22FFC40035FE570EDCFD12D81D" box="[509,711,1784,1811]" gridcol="3" gridrow="3" pageId="60" pageNumber="61">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFC40035FCAE0EDCFC67D81D" box="[772,946,1784,1811]" gridcol="4" gridrow="3" pageId="60" pageNumber="61">pl5rl5v2-1-2-2</td>
</tr>
<tr id="07CE535EFFC40035FF610F3BFC67D834" box="[203,946,1823,1850]" gridrow="4" pageId="60" pageNumber="61">
<th id="441F3A22FFC40035FF610F3BFF3FD834" box="[203,234,1823,1850]" gridcol="0" gridrow="4" pageId="60" pageNumber="61">IV</th>
<td id="441F3A22FFC40035FEA00F3BFEBBD834" box="[266,366,1823,1850]" gridcol="1" gridrow="4" pageId="60" pageNumber="61">pl4d3rl3</td>
<td id="441F3A22FFC40035FE390F3BFE0CD834" box="[403,473,1823,1850]" gridcol="2" gridrow="4" pageId="60" pageNumber="61">pl1rl1</td>
<td id="441F3A22FFC40035FE570F3BFD12D834" box="[509,711,1823,1850]" gridcol="3" gridrow="4" pageId="60" pageNumber="61">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFC40035FCAE0F3BFC67D834" box="[772,946,1823,1850]" gridcol="4" gridrow="4" pageId="60" pageNumber="61">pl5rl5v2-1-2-2</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFC4FFF7FF170F46FAA0DEC4" blockId="60.[187,1401,1890,2025]" lastBlockId="61.[187,1399,265,458]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
MALE PALP (
<figureCitation id="21C54D99FFC4FFF6FEFD0F46FE39D873" box="[343,492,1890,1917]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="60" pageNumber="61">Fig. 25A–C</figureCitation>
). Tibia with apical, short pronged VTA connected to conical RTA, behind which dorso-retrolateral excavation accommodates retrobasal cymbial extension; cymbium oval with narrowly-rounded apical extremity, without dorsal patch of chemosensitive setae, having stout retrolateral paracymbial process (PP) and retrobasal cymbial extension (CE) extending at least ¼ length of papal tibia; subtegulum large, filling most of cymbial concavity and extending to prolateral cymbial margin, and with probasal broad ridge (SL) that interlocks with tegulum; tegulum rounded with conical process (TL) on prolateral margin, this process interlocking with subtegular ridge; embolus (E) broad and thick, arising proapically, with pointed apex, small crest subapically and curved basomediam process (EBP) arising from embolus base; median apophysis (MA) strongly sclerotized, short, hooked, curved prolaterad; conductor (C) hyaline fan-shaped, narrow at base; paraconductor (Pc) small, cradling inner margin of E.
</paragraph>
<caption id="ED810194FFC5FFF7FF170EFCFB2DD8E6" ID-DOI="http://doi.org/10.5281/zenodo.14001849" ID-Zenodo-Dep="14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="61" pageNumber="62" startId="61.[189,231,1752,1778]" targetBox="[211,1360,510,1693]" targetPageId="61">
<paragraph id="B941511CFFC5FFF7FF170EFCFB2DD8E6" blockId="61.[187,1400,1752,2025]" pageId="61" pageNumber="62">
<emphasis id="8B8A8D0EFFC5FFF7FF170EFCFEC3D9FC" bold="true" box="[189,278,1752,1778]" pageId="61" pageNumber="62">Fig. 25.</emphasis>
<taxonomicName id="7EFE2A9FFFC5FFF7FEB60EFCFDEFD9FC" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[284,570,1752,1778]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="silvadavilae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC5FFF7FEB60EFCFDEFD9FC" box="[284,570,1752,1778]" italics="true" pageId="61" pageNumber="62">Zorodictyna silvadavilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC5FFF7FDEA0EFCFD4FD9FC" box="[576,666,1752,1778]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
, genitalia drawings; A–C
<emphasis id="8B8A8D0EFFC5FFF7FC150EFCFC05D9FC" bold="true" box="[959,976,1752,1778]" pageId="61" pageNumber="62">=</emphasis>
male holotype (CASENT_9006231; CAS); D–E
<emphasis id="8B8A8D0EFFC5FFF7FEE90EDFFE81D81B" bold="true" box="[323,340,1787,1813]" pageId="61" pageNumber="62">=</emphasis>
female paratype (CASENT_9006230; USNM).
<emphasis id="8B8A8D0EFFC5FFF7FCDC0EDFFC5ED81B" bold="true" box="[886,907,1787,1813]" pageId="61" pageNumber="62">A</emphasis>
. Left palp, prolateral view.
<emphasis id="8B8A8D0EFFC5FFF7FB140EDFFB04D81B" bold="true" box="[1214,1233,1787,1813]" pageId="61" pageNumber="62">B</emphasis>
. Idem, ventral view.
<emphasis id="8B8A8D0EFFC5FFF7FEAF0F3AFECFD836" bold="true" box="[261,282,1822,1848]" pageId="61" pageNumber="62">C</emphasis>
. Idem, retrolateral view.
<emphasis id="8B8A8D0EFFC5FFF7FDE10F3AFD8AD836" bold="true" box="[587,607,1822,1848]" pageId="61" pageNumber="62">D</emphasis>
. Epigyne, ventral view.
<emphasis id="8B8A8D0EFFC5FFF7FC2C0F3AFC4DD836" bold="true" box="[902,920,1822,1848]" pageId="61" pageNumber="62">E</emphasis>
. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels.
</paragraph>
</caption>
<paragraph id="B941511CFFC6FFF4FF40092DFB71DE46" blockId="62.[189,1188,264,328]" pageId="62" pageNumber="63">
<emphasis id="8B8A8D0EFFC6FFF4FF40092DFE68DE2D" bold="true" box="[234,445,265,291]" pageId="62" pageNumber="63">
Female
<typeStatus id="6645EFBEFFC6FFF4FEE7092DFE68DE2D" box="[333,445,265,291]" pageId="62" pageNumber="63" type="paratype">paratype</typeStatus>
</emphasis>
(CASENT_9015400; CAS; except as otherwise indicated) MEASUREMENTS. Total length: 23.00. Carapace: length 11.20, width 8.30, height 5.00.
</paragraph>
<paragraph id="B941511CFFC6FFF4FF17094BFBB8DEA3" blockId="62.[189,1399,367,430]" pageId="62" pageNumber="63">
COLOUR (
<figureCitation id="21C54D99FFC6FFF4FE9B094BFE14DE87" box="[305,449,367,394]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; A–B = male holotype (CASENT_9006231; CAS); C–D = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="62" pageNumber="63">Fig. 24C–D</figureCitation>
). As in male: dorsal and lateral of cephalothorax, abdomen and legs unicolorous grey-brown, venter of sternum, mouthparts, coxae and trochanters yellow white.
</paragraph>
<paragraph id="B941511CFFC6FFF4FF1709F2FB7EDEFE" blockId="62.[189,1195,470,497]" box="[189,1195,470,497]" pageId="62" pageNumber="63">
STERNUM (
<figureCitation id="21C54D99FFC6FFF4FE9709F2FE7FDEFE" box="[317,426,470,497]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; A–B = male holotype (CASENT_9006231; CAS); C–D = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="62" pageNumber="63">Fig. 24D</figureCitation>
). Oval with sides straight next to leg coxae; 4.10 wide, 4.90 long.
</paragraph>
<paragraph id="B941511CFFC6FFF4FF170A3DFDF7DD3C" blockId="62.[189,546,536,563]" box="[189,546,536,563]" pageId="62" pageNumber="63">CHILUM. 0.25 high, 0.75 wide.</paragraph>
<paragraph id="B941511CFFC6FFF4FF170A7FFCCADD97" blockId="62.[188,1399,603,666]" pageId="62" pageNumber="63">EYES. ALE 0.30; AME 0.29; PLE 0.31; PME 0.28; ALE–AME 0.18; AME–AME 0.50; PLE–PME 0.48; PME–PME 0.23; MOQ: AW 0.73, PW 0.83, L 0.23.</paragraph>
<paragraph id="B941511CFFC6FFF4FF170AE6FDC7DDD2" blockId="62.[189,530,706,732]" box="[189,530,706,732]" pageId="62" pageNumber="63">CLYPEUS. With 14 long setae.</paragraph>
<paragraph id="B941511CFFC6FFF4FF170B20FB39DC11" blockId="62.[189,1260,772,799]" box="[189,1260,772,799]" pageId="62" pageNumber="63">LEG. Scopulae beneath all tarsi, apical half of metatarsus I and apical third of metatarsus II.</paragraph>
<paragraph id="B941511CFFC6FFF4FF170B63FE4FDC6F" blockId="62.[189,410,839,865]" box="[189,410,839,865]" pageId="62" pageNumber="63">Leg measurements</paragraph>
<paragraph id="B941511CFFC6FFF4FEA00B56FCD1DB25" pageId="62" pageNumber="63">
<table id="CBFEA3BCFFC60035FF610B55FCD1DB22" box="[203,772,881,1068]" gridcols="7" gridrows="5" pageId="62" pageNumber="63" rowsContinueIn="62.[203,936,1151,1336]">
<tr id="07CE535EFFC60035FF610B55FCD1DC82" box="[203,772,881,908]" gridrow="0" pageId="62" pageNumber="63" rowspan-0="1">
<th id="441F3A22FFC60035FEA00B55FEE8DC82" box="[266,317,881,908]" gridcol="1" gridrow="0" pageId="62" pageNumber="63">F</th>
<th id="441F3A22FFC60035FEF50B55FE47DC82" box="[351,402,881,908]" gridcol="2" gridrow="0" pageId="62" pageNumber="63">P</th>
<th id="441F3A22FFC60035FE1E0B55FE32DC82" box="[436,487,881,908]" gridcol="3" gridrow="0" pageId="62" pageNumber="63">T</th>
<th id="441F3A22FFC60035FDA20B55FD9FDC82" box="[520,586,881,908]" gridcol="4" gridrow="0" pageId="62" pageNumber="63">Mt</th>
<th id="441F3A22FFC60035FDC70B55FD75DC82" box="[621,672,881,908]" gridcol="5" gridrow="0" pageId="62" pageNumber="63">t</th>
<th id="441F3A22FFC60035FD680B55FCD1DC82" box="[706,772,881,908]" gridcol="6" gridrow="0" pageId="62" pageNumber="63">Total</th>
</tr>
<tr id="07CE535EFFC60035FF610BBEFCD1DCBA" box="[203,772,922,948]" gridrow="1" pageId="62" pageNumber="63">
<th id="441F3A22FFC60035FF610BBEFF3FDCBA" box="[203,234,922,948]" gridcol="0" gridrow="1" pageId="62" pageNumber="63">I</th>
<td id="441F3A22FFC60035FEA00BBEFEE8DCBA" box="[266,317,922,948]" gridcol="1" gridrow="1" pageId="62" pageNumber="63">8.30</td>
<td id="441F3A22FFC60035FEF50BBEFE47DCBA" box="[351,402,922,948]" gridcol="2" gridrow="1" pageId="62" pageNumber="63">4.30</td>
<td id="441F3A22FFC60035FE1E0BBEFE32DCBA" box="[436,487,922,948]" gridcol="3" gridrow="1" pageId="62" pageNumber="63">7.00</td>
<td id="441F3A22FFC60035FDA20BBEFD9FDCBA" box="[520,586,922,948]" gridcol="4" gridrow="1" pageId="62" pageNumber="63">6.20</td>
<td id="441F3A22FFC60035FDC70BBEFD75DCBA" box="[621,672,922,948]" gridcol="5" gridrow="1" pageId="62" pageNumber="63">3.10</td>
<td id="441F3A22FFC60035FD680BBEFCD1DCBA" box="[706,772,922,948]" gridcol="6" gridrow="1" pageId="62" pageNumber="63">28.90</td>
</tr>
<tr id="07CE535EFFC60035FF610BE5FCD1DCD2" box="[203,772,961,988]" gridrow="2" pageId="62" pageNumber="63">
<th id="441F3A22FFC60035FF610BE5FF3FDCD2" box="[203,234,961,988]" gridcol="0" gridrow="2" pageId="62" pageNumber="63">II</th>
<td id="441F3A22FFC60035FEA00BE5FEE8DCD2" box="[266,317,961,988]" gridcol="1" gridrow="2" pageId="62" pageNumber="63">7.50</td>
<td id="441F3A22FFC60035FEF50BE5FE47DCD2" box="[351,402,961,988]" gridcol="2" gridrow="2" pageId="62" pageNumber="63">4.00</td>
<td id="441F3A22FFC60035FE1E0BE5FE32DCD2" box="[436,487,961,988]" gridcol="3" gridrow="2" pageId="62" pageNumber="63">5.60</td>
<td id="441F3A22FFC60035FDA20BE5FD9FDCD2" box="[520,586,961,988]" gridcol="4" gridrow="2" pageId="62" pageNumber="63">5.80</td>
<td id="441F3A22FFC60035FDC70BE5FD75DCD2" box="[621,672,961,988]" gridcol="5" gridrow="2" pageId="62" pageNumber="63">2.90</td>
<td id="441F3A22FFC60035FD680BE5FCD1DCD2" box="[706,772,961,988]" gridcol="6" gridrow="2" pageId="62" pageNumber="63">25.80</td>
</tr>
<tr id="07CE535EFFC60035FF610BCDFCD1DB0A" box="[203,772,1001,1028]" gridrow="3" pageId="62" pageNumber="63">
<th id="441F3A22FFC60035FF610BCDFF3FDB0A" box="[203,234,1001,1028]" gridcol="0" gridrow="3" pageId="62" pageNumber="63">III</th>
<td id="441F3A22FFC60035FEA00BCDFEE8DB0A" box="[266,317,1001,1028]" gridcol="1" gridrow="3" pageId="62" pageNumber="63">6.20</td>
<td id="441F3A22FFC60035FEF50BCDFE47DB0A" box="[351,402,1001,1028]" gridcol="2" gridrow="3" pageId="62" pageNumber="63">3.20</td>
<td id="441F3A22FFC60035FE1E0BCDFE32DB0A" box="[436,487,1001,1028]" gridcol="3" gridrow="3" pageId="62" pageNumber="63">4.60</td>
<td id="441F3A22FFC60035FDA20BCDFD9FDB0A" box="[520,586,1001,1028]" gridcol="4" gridrow="3" pageId="62" pageNumber="63">5.00</td>
<td id="441F3A22FFC60035FDC70BCDFD75DB0A" box="[621,672,1001,1028]" gridcol="5" gridrow="3" pageId="62" pageNumber="63">3.00</td>
<td id="441F3A22FFC60035FD680BCDFCD1DB0A" box="[706,772,1001,1028]" gridcol="6" gridrow="3" pageId="62" pageNumber="63">22.00</td>
</tr>
<tr id="07CE535EFFC60035FF610C35FCD1DB22" box="[203,772,1041,1068]" gridrow="4" pageId="62" pageNumber="63">
<th id="441F3A22FFC60035FF610C35FF3FDB22" box="[203,234,1041,1068]" gridcol="0" gridrow="4" pageId="62" pageNumber="63">IV</th>
<td id="441F3A22FFC60035FEA00C35FEE8DB22" box="[266,317,1041,1068]" gridcol="1" gridrow="4" pageId="62" pageNumber="63">8.80</td>
<td id="441F3A22FFC60035FEF50C35FE47DB22" box="[351,402,1041,1068]" gridcol="2" gridrow="4" pageId="62" pageNumber="63">3.60</td>
<td id="441F3A22FFC60035FE1E0C35FE32DB22" box="[436,487,1041,1068]" gridcol="3" gridrow="4" pageId="62" pageNumber="63">7.00</td>
<td id="441F3A22FFC60035FDA20C35FD9FDB22" box="[520,586,1041,1068]" gridcol="4" gridrow="4" pageId="62" pageNumber="63">12.40</td>
<td id="441F3A22FFC60035FDC70C35FD75DB22" box="[621,672,1041,1068]" gridcol="5" gridrow="4" pageId="62" pageNumber="63">3.50</td>
<td id="441F3A22FFC60035FD680C35FCD1DB22" box="[706,772,1041,1068]" gridcol="6" gridrow="4" pageId="62" pageNumber="63">33.30</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFC6FFF4FF170C70FC0DD95A" blockId="62.[189,301,1108,1134]" lastBlockId="62.[187,1400,1376,1621]" pageId="62" pageNumber="63">
Spination GENITALIA. Epigyne (
<typeStatus id="6645EFBEFFC6FFF4FE1C0D45FDC8DA75" box="[438,541,1377,1403]" pageId="62" pageNumber="63" type="paratype">paratype</typeStatus>
CASENT_9006230; USNM), (
<figureCitation id="21C54D99FFC6FFF4FC3A0D44FC2EDA75" box="[912,1019,1376,1403]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="62" pageNumber="63">Fig. 25D</figureCitation>
) with quadrangular median lobe (ML) plate transverse with smooth margin; copulatory openings (CO) hidden beneath anterolateral corners of ML; lateral lobes (LL) convex, without lateral teeth or projections but with faint concavities along margin of ML, converging posteriorly and exposing triangular area of median sector (MS). Endogyne (
<figureCitation id="21C54D99FFC6FFF4FF6D0DD6FEE5D902" box="[199,304,1521,1548]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="62" pageNumber="63">Fig. 25E</figureCitation>
) with broad, semitransparent copulatory ducts (CD) extending from beneath ML to simple, convex, bean-shaped spermathecae (S), spermathecal head with several small pores; spermathecal bases meeting in center; fertilization ducts (FD) short and slightly curved.
</paragraph>
<paragraph id="B941511CFFC6FFF4FEA00C5BFC7DDA36" pageId="62" pageNumber="63">
<table id="CBFEA3BCFFC60035FF610C5BFC7DDA36" box="[203,936,1151,1336]" gridcols="5" gridrows="5" pageId="62" pageNumber="63" rowsContinueFrom="62.[203,772,881,1068]">
<tr id="07CE535EFFC60035FF610C5BFC7DDB97" box="[203,936,1151,1177]" gridrow="0" pageId="62" pageNumber="63" rowspan-0="1">
<th id="441F3A22FFC60035FEA00C5BFEBBDB97" box="[266,366,1151,1177]" gridcol="1" gridrow="0" pageId="62" pageNumber="63">F</th>
<th id="441F3A22FFC60035FE390C5BFE0CDB97" box="[403,473,1151,1177]" gridcol="2" gridrow="0" pageId="62" pageNumber="63">P</th>
<th id="441F3A22FFC60035FE570C5BFD7ADB97" box="[509,687,1151,1177]" gridcol="3" gridrow="0" pageId="62" pageNumber="63">T</th>
<th id="441F3A22FFC60035FD750C5BFC7DDB97" box="[735,936,1151,1177]" gridcol="4" gridrow="0" pageId="62" pageNumber="63">Mt</th>
</tr>
<tr id="07CE535EFFC60035FF610C82FC7DDBCF" box="[203,936,1190,1217]" gridrow="1" pageId="62" pageNumber="63">
<th id="441F3A22FFC60035FF610C82FF3FDBCF" box="[203,234,1190,1217]" gridcol="0" gridrow="1" pageId="62" pageNumber="63">I</th>
<td id="441F3A22FFC60035FEA00C82FEBBDBCF" box="[266,366,1190,1217]" gridcol="1" gridrow="1" pageId="62" pageNumber="63">pl2d1</td>
<td id="441F3A22FFC60035FE390C82FE0CDBCF" box="[403,473,1190,1217]" gridcol="2" gridrow="1" pageId="62" pageNumber="63">–</td>
<td id="441F3A22FFC60035FE570C82FD7ADBCF" box="[509,687,1190,1217]" gridcol="3" gridrow="1" pageId="62" pageNumber="63">v2-2-2-2</td>
<td id="441F3A22FFC60035FD750C82FC7DDBCF" box="[735,936,1190,1217]" gridcol="4" gridrow="1" pageId="62" pageNumber="63">v2-2-2</td>
</tr>
<tr id="07CE535EFFC60035FF610CEAFC7DDBE7" box="[203,936,1230,1257]" gridrow="2" pageId="62" pageNumber="63">
<th id="441F3A22FFC60035FF610CEAFF3FDBE7" box="[203,234,1230,1257]" gridcol="0" gridrow="2" pageId="62" pageNumber="63">II</th>
<td id="441F3A22FFC60035FEA00CEAFEBBDBE7" box="[266,366,1230,1257]" gridcol="1" gridrow="2" pageId="62" pageNumber="63">pl2d1</td>
<td id="441F3A22FFC60035FE390CEAFE0CDBE7" box="[403,473,1230,1257]" gridcol="2" gridrow="2" pageId="62" pageNumber="63">–</td>
<td id="441F3A22FFC60035FE570CEAFD7ADBE7" box="[509,687,1230,1257]" gridcol="3" gridrow="2" pageId="62" pageNumber="63">v2-2-2-2</td>
<td id="441F3A22FFC60035FD750CEAFC7DDBE7" box="[735,936,1230,1257]" gridcol="4" gridrow="2" pageId="62" pageNumber="63">v2-2-2</td>
</tr>
<tr id="07CE535EFFC60035FF610CD2FC7DDA1F" box="[203,936,1270,1297]" gridrow="3" pageId="62" pageNumber="63">
<th id="441F3A22FFC60035FF610CD2FF3FDA1F" box="[203,234,1270,1297]" gridcol="0" gridrow="3" pageId="62" pageNumber="63">III</th>
<td id="441F3A22FFC60035FEA00CD2FEBBDA1F" box="[266,366,1270,1297]" gridcol="1" gridrow="3" pageId="62" pageNumber="63">pl2d3rl3</td>
<td id="441F3A22FFC60035FE390CD2FE0CDA1F" box="[403,473,1270,1297]" gridcol="2" gridrow="3" pageId="62" pageNumber="63">pl1rl1</td>
<td id="441F3A22FFC60035FE570CD2FD7ADA1F" box="[509,687,1270,1297]" gridcol="3" gridrow="3" pageId="62" pageNumber="63">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFC60035FD750CD2FC7DDA1F" box="[735,936,1270,1297]" gridcol="4" gridrow="3" pageId="62" pageNumber="63">pl4rl4v2-2-1-2</td>
</tr>
<tr id="07CE535EFFC60035FF610D3AFC7DDA36" box="[203,936,1310,1336]" gridrow="4" pageId="62" pageNumber="63">
<th id="441F3A22FFC60035FF610D3AFF3FDA36" box="[203,234,1310,1336]" gridcol="0" gridrow="4" pageId="62" pageNumber="63">IV</th>
<td id="441F3A22FFC60035FEA00D3AFEBBDA36" box="[266,366,1310,1336]" gridcol="1" gridrow="4" pageId="62" pageNumber="63">pl3d3rl1</td>
<td id="441F3A22FFC60035FE390D3AFE0CDA36" box="[403,473,1310,1336]" gridcol="2" gridrow="4" pageId="62" pageNumber="63">pl1rl1</td>
<td id="441F3A22FFC60035FE570D3AFD7ADA36" box="[509,687,1310,1336]" gridcol="3" gridrow="4" pageId="62" pageNumber="63">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFC60035FD750D3AFC7DDA36" box="[735,936,1310,1336]" gridcol="4" gridrow="4" pageId="62" pageNumber="63">pl4d2rl3v2-2-1-2</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFC6FFF4FF170E58FEE6D998" blockId="62.[189,307,1660,1686]" box="[189,307,1660,1686]" pageId="62" pageNumber="63">
<heading id="E209E670FFC6FFF4FF170E58FEE6D998" bold="true" box="[189,307,1660,1686]" fontSize="11" level="3" pageId="62" pageNumber="63" reason="3">
<emphasis id="8B8A8D0EFFC6FFF4FF170E58FEE6D998" bold="true" box="[189,307,1660,1686]" pageId="62" pageNumber="63">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC6FFF4FF400E88FE58D9C8" blockId="62.[189,1399,1708,2025]" box="[234,397,1708,1735]" pageId="62" pageNumber="63">
<emphasis id="8B8A8D0EFFC6FFF4FF400E88FEE0D9C8" bold="true" box="[234,309,1708,1734]" pageId="62" pageNumber="63">Males</emphasis>
(n = 4)
</paragraph>
<paragraph id="B941511CFFC6FFF4FF170EF4FC41D8E6" blockId="62.[189,1399,1708,2025]" pageId="62" pageNumber="63">Total length = 17.60–24.00, carapace length/width = 1.12–1.28, carapace height/width = 0.44–0.54, PER/carapace width = 0.20–0.22, PER/OAL = 2.50–2.80, PER/AER = 1.35–1.56, OAL/OQL = 1.00– 1.13, OQP/OQA = 1.22–1.46, sternum length/width = 1.22–1.33, palpal coxa length/width = 1.92–2.90, femur I length/carapace width = 1.04–1.23, metatarsus I length/carapace width = 0.98–1.16, femur IV length/carapace width = 1.10–1.31. Male palp with apical RTA tips acute (Marojejy) to blunt (Masoala), EBP hook at embolic base triangular (Marojejy) to crescentic (Masoala), retromedian tegular lock (TL) blunt (Marojejy) to sharply triangular (Masoala). Males from Marojejy have a small subterminal crest on the embolus, which is absent from the male from Masoala.</paragraph>
<paragraph id="B941511CFFC7FFF5FF40092DFE7CDE2D" blockId="63.[189,1399,265,502]" box="[234,425,265,291]" pageId="63" pageNumber="64">
<emphasis id="8B8A8D0EFFC7FFF5FF40092DFE84DE2D" bold="true" box="[234,337,265,291]" pageId="63" pageNumber="64">Females</emphasis>
(n = 4)
</paragraph>
<paragraph id="B941511CFFC7FFF5FF170908FB9EDEF8" blockId="63.[189,1399,265,502]" pageId="63" pageNumber="64">Total length = 23.00–32.50, carapace length/width = 1.26–1.41, carapace height/width = 0.46–0.60, PER/carapace width = 0.23–0.26, PER/OAL = 2.87–3.41, PER/AER = 1.30–1.43, OAL/OQL = 1.03– 1.06, OQP/OQA = 1.14–1.23, sternum length/width = 1.18–1.24, palpal coxa length/width = 2.32–3.45, femur I length/carapace width = 0.92–1.00, metatarsus I length/carapace width = 0.66–0.75, femur IV length/carapace width = 1.02–1.07. Epigyne with median lobe length 0.70–0.78 times width; distance from median lobe base to epigastric fold 0.29–0.40 times median lobe length.</paragraph>
<paragraph id="B941511CFFC7FFF5FF170A39FE40DD39" blockId="63.[189,405,541,567]" box="[189,405,541,567]" pageId="63" pageNumber="64">
<heading id="E209E670FFC7FFF5FF170A39FE40DD39" bold="true" box="[189,405,541,567]" fontSize="11" level="3" pageId="63" pageNumber="64" reason="3">
<emphasis id="8B8A8D0EFFC7FFF5FF170A39FE40DD39" bold="true" box="[189,405,541,567]" pageId="63" pageNumber="64">Note on synonym</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC7FFF5FF170A68FB10DD87" blockId="63.[189,1399,588,649]" pageId="63" pageNumber="64">
A specimen of this species (CASENT_9014998) was used as an exemplar in a previously published molecular phylogeny: it was listed as “
<taxonomicName id="7EFE2A9FFFC7FFF5FD2F0A4BFCE9DD87" box="[645,828,623,649]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8B8A8D0EFFC7FFF5FD2F0A4BFCC6DD87" box="[645,787,623,649]" italics="true" pageId="63" pageNumber="64">Zorodictyna</emphasis>
sp.
</taxonomicName>
CG47” by
<bibRefCitation id="DD6F2CEDFFC7FFF5FC680A4BFB14DD87" author="Wheeler W. C. &amp; Coddington J. A. &amp; Crowley L. M. &amp; Dimitrov D. &amp; Goloboff P. A. &amp; Griswold C. E. &amp; Hormiga G. &amp; Prendini L. &amp; Ramirez M. J. &amp; Sierwald P. &amp; Almeida-Silva L. M. &amp; Alvarez-Padilla F. &amp; Arnedo M. A. &amp; Benavides L. R. &amp; Benjamin S. P. &amp; Bond J. E. &amp; Grismado C. J. &amp; Hasan E. &amp; Hedin M. &amp; Izquierdo M. A. &amp; Labarque F. M. &amp; Ledford J. &amp; Lopardo L. &amp; Maddison W. P. &amp; Miller J. A. &amp; Piacentini L. N. &amp; Platnick N. I. &amp; Polotow D. &amp; Silva-Davila D. &amp; Scharff N. &amp; Szuts T. &amp; Ubick D. &amp; Vink C. &amp; Wood H. M. &amp; Zhang J. X." box="[962,1217,623,649]" pageId="63" pageNumber="64" pagination="576 - 616" refId="ref39136" refString="Wheeler W. C., Coddington J. A., Crowley L. M., Dimitrov D., Goloboff P. A., Griswold C. E., Hormiga G., Prendini L., Ramirez M. J., Sierwald P., Almeida-Silva L. M., Alvarez-Padilla F., Arnedo M. A., Benavides L. R., Benjamin S. P., Bond J. E., Grismado C. J., Hasan E., Hedin M., Izquierdo M. A., Labarque F. M., Ledford J., Lopardo L., Maddison W. P., Miller J. A., Piacentini L. N., Platnick N. I., Polotow D., Silva-Davila D., Scharff N., Szuts T., Ubick D., Vink C., Wood H. M. &amp; Zhang J. X. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. https: // doi. org / 10.1111 / cla. 12182" type="journal article" year="2017">
Wheeler
<emphasis id="8B8A8D0EFFC7FFF5FB870A54FBBDDD87" box="[1069,1128,623,649]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
(2017)
</bibRefCitation>
.
</paragraph>
<paragraph id="B941511CFFC7FFF5FF170A94FEABDDC4" blockId="63.[189,382,688,714]" box="[189,382,688,714]" pageId="63" pageNumber="64">
<heading id="E209E670FFC7FFF5FF170A94FEABDDC4" bold="true" box="[189,382,688,714]" fontSize="11" level="3" pageId="63" pageNumber="64" reason="3">
<emphasis id="8B8A8D0EFFC7FFF5FF170A94FEABDDC4" bold="true" box="[189,382,688,714]" pageId="63" pageNumber="64">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC7FFF5FF170AFBFBE5DC12" blockId="63.[187,1398,735,797]" pageId="63" pageNumber="64">This species has been collected when running on leaf litter in closed canopy montane rainforest. Individuals have an entire cribellum but we have yet to learn details of their use of silk.</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFC7FFF5FF170B67FC8FDCA1" pageId="63" pageNumber="64" type="distribution">
<paragraph id="B941511CFFC7FFF5FF170B67FE83DC53" blockId="63.[189,342,835,861]" box="[189,342,835,861]" pageId="63" pageNumber="64">
<heading id="E209E670FFC7FFF5FF170B67FE83DC53" bold="true" box="[189,342,835,861]" fontSize="11" level="3" pageId="63" pageNumber="64" reason="3">
<emphasis id="8B8A8D0EFFC7FFF5FF170B67FE83DC53" bold="true" box="[189,342,835,861]" pageId="63" pageNumber="64">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC7FFF5FF170B56FC8FDCA1" blockId="63.[189,1399,882,943]" pageId="63" pageNumber="64">
Individuals of this species are known only from the montane rainforests on the Marojejy Massif and the Masoala Peninsula in northeastern
<collectingCountry id="C1E9118CFFC7FFF5FDFD0BB1FD30DCA1" box="[599,741,917,943]" name="Madagascar" pageId="63" pageNumber="64">Madagascar</collectingCountry>
(
<figureCitation id="21C54D99FFC7FFF5FD5F0BB1FC98DCA1" box="[757,845,917,943]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="63" pageNumber="64">Fig. 26</figureCitation>
).
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<figureCitation id="21C54D99FFCEFFFCFD680974FC9DDE65" box="[706,840,336,363]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="56.[189,231,1928,1954]" captionStartId-1="57.[189,232,1636,1662]" captionTargetBox-0="[189,1398,1121,1890]" captionTargetBox-1="[219,1348,279,1580]" captionTargetId-0="figure-176@56.[189,1398,1121,1890]" captionTargetId-1="figure-12@57.[189,1398,266,1598]" captionTargetPageId-0="56" captionTargetPageId-1="57" captionText-0="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." captionText-1="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi-0="http://doi.org/10.5281/zenodo.14001843" figureDoi-1="http://doi.org/10.5281/zenodo.14001845" httpUri-0="https://zenodo.org/record/14001843/files/figure.png" httpUri-1="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Figs 22–23</figureCitation>
,
<figureCitation id="21C54D99FFCEFFFCFCFE0974FCA4DE64" box="[852,881,336,362]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="54" pageNumber="55">26</figureCitation>
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<taxonomicName id="7EFE2A9FFFCEFFFCFF1709B6FE5FDEA3" box="[189,394,402,429]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="8B8A8D0EFFCEFFFCFF1709B6FE9EDEA2" box="[189,331,402,428]" italics="true" pageId="54" pageNumber="55">Zorodictyna</emphasis>
sp. 1
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–
<bibRefCitation id="DD6F2CEDFFCEFFFCFE0D09B6FD45DEA3" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[423,656,402,429]" pageId="54" pageNumber="55" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow
<emphasis id="8B8A8D0EFFCEFFFCFDBB09B7FD9EDEA2" box="[529,587,402,428]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2015
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFCEFFFCFF1709F0FCDEDCE1" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="B941511CFFCEFFFCFF1709F0FEE3DEE0" blockId="54.[189,310,468,494]" box="[189,310,468,494]" pageId="54" pageNumber="55">
<heading id="E209E670FFCEFFFCFF1709F0FEE3DEE0" bold="true" box="[189,310,468,494]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="8B8A8D0EFFCEFFFCFF1709F0FEE3DEE0" bold="true" box="[189,310,468,494]" pageId="54" pageNumber="55">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170A27FCDEDCE1" blockId="54.[187,1400,515,1007]" pageId="54" pageNumber="55">
Males of
<taxonomicName id="7EFE2A9FFFCEFFFCFE8F0A27FE2BDD13" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[293,510,515,541]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFCEFFFCFE8F0A27FE2BDD13" box="[293,510,515,541]" italics="true" pageId="54" pageNumber="55">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCEFFFCFDAE0A20FD89DD10" box="[516,604,516,542]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
are distinguished from
<taxonomicName id="7EFE2A9FFFCEFFFCFCCF0A20FBA1DD10" authority="(Lenz, 1891)" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[869,1140,515,542]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCEFFFCFCCF0A20FC09DD13" box="[869,988,515,541]" italics="true" pageId="54" pageNumber="55">Z. oswaldi</emphasis>
(
<bibRefCitation id="DD6F2CEDFFCEFFFCFC430A20FBB9DD10" author="Lenz H." box="[1001,1132,516,542]" pageId="54" pageNumber="55" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">Lenz, 1891</bibRefCitation>
)
</taxonomicName>
by having a retrobasal cymbial extension (CE) that extends at least ¼ the length of the papal tibia and a thick retrobasal paracymbial process (PP) (
<figureCitation id="21C54D99FFCEFFFCFE510A6FFD52DD6B" box="[507,647,587,613]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Fig. 23B–C</figureCitation>
) (vs
<taxonomicName id="7EFE2A9FFFCEFFFCFD6E0A68FCEADD6B" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[708,831,587,613]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCEFFFCFD6E0A68FCEADD6B" box="[708,831,587,613]" italics="true" pageId="54" pageNumber="55">Z. oswaldi</emphasis>
</taxonomicName>
lacks the PP and has a very short CE;
<bibRefCitation id="DD6F2CEDFFCEFFFCFAA30A6FFE8CDD87" author="Griswold C. E." pageId="54" pageNumber="55" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993: 19–21</bibRefCitation>
). It is distinguished from
<taxonomicName id="7EFE2A9FFFCEFFFCFD390A4BFC91DD87" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[659,836,623,649]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="silvadavilae" status="sp. nov.">
<emphasis id="8B8A8D0EFFCEFFFCFD390A4BFC91DD87" box="[659,836,623,649]" italics="true" pageId="54" pageNumber="55">Z. silvadavilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFCEFFFCFCE10A4BFC7CDD87" box="[843,937,623,649]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
in having the male palp with a nearly triangular basal embolic hook, the embolus lacking a crest, and a retromedian longitudinal ridge that extends for most of the tegulum length (
<figureCitation id="21C54D99FFCEFFFCFD080A92FCE4DDDE" box="[674,817,694,720]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Fig. 23A–C</figureCitation>
) (vs
<taxonomicName id="7EFE2A9FFFCEFFFCFCDA0A93FBCBDDDE" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[880,1054,694,720]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="silvadavilae">
<emphasis id="8B8A8D0EFFCEFFFCFCDA0A93FBCBDDDE" box="[880,1054,694,720]" italics="true" pageId="54" pageNumber="55">Z. silvadavilae</emphasis>
</taxonomicName>
has slender, crescentic basal embolic hook, the embolus with subapical distal crest, and the tegular retromedian longitudinal ridge that is weak, short, extending for less than one-half tegulum length,
<figureCitation id="21C54D99FFCEFFFCFC490AD9FBA6DC16" box="[995,1139,765,792]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="54" pageNumber="55">Fig. 25A–C</figureCitation>
). Females of
<taxonomicName id="7EFE2A9FFFCEFFFCFABA0ADAFAA2DC19" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1296,1399,765,791]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="almae">
<emphasis id="8B8A8D0EFFCEFFFCFABA0ADAFAA2DC19" box="[1296,1399,765,791]" italics="true" pageId="54" pageNumber="55">Z. almae</emphasis>
</taxonomicName>
are distinguished from
<taxonomicName id="7EFE2A9FFFCEFFFCFE780B06FD98DC35" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[466,589,801,827]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCEFFFCFE780B06FD98DC35" box="[466,589,801,827]" italics="true" pageId="54" pageNumber="55">Z. oswaldi</emphasis>
</taxonomicName>
by lacking teeth on the lateral lobes (LL) of the epigyne (
<figureCitation id="21C54D99FFCEFFFCFAA90B05FABADC35" box="[1283,1391,801,827]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Fig. 23D</figureCitation>
) (vs
<taxonomicName id="7EFE2A9FFFCEFFFCFF4D0B62FEB5DC51" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[231,352,837,863]" class="Arachnida" family="Liocranidae" genus="Agroeca" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFCEFFFCFF4D0B62FEB5DC51" box="[231,352,837,863]" italics="true" pageId="54" pageNumber="55">Z. oswaldi</emphasis>
</taxonomicName>
has the LL with teeth;
<bibRefCitation id="DD6F2CEDFFCEFFFCFDC20B61FCCFDC51" author="Griswold C. E." box="[616,794,837,863]" pageId="54" pageNumber="55" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: fig. 24); females of
<taxonomicName id="7EFE2A9FFFCEFFFCFBAF0B62FBBEDC51" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1029,1131,837,863]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="almae">
<emphasis id="8B8A8D0EFFCEFFFCFBAF0B62FBBEDC51" box="[1029,1131,837,863]" italics="true" pageId="54" pageNumber="55">Z. almae</emphasis>
</taxonomicName>
are distinguished from
<taxonomicName id="7EFE2A9FFFCEFFFCFF160B4EFEBFDC8D" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[188,362,873,899]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="silvadavilae">
<emphasis id="8B8A8D0EFFCEFFFCFF160B4EFEBFDC8D" box="[188,362,873,899]" italics="true" pageId="54" pageNumber="55">Z. silvadavilae</emphasis>
</taxonomicName>
by having the lateral lobes of epigyne parallel and the median sector (MS) rectangular (
<figureCitation id="21C54D99FFCEFFFCFF6D0BA8FEE1DCA9" box="[199,308,908,935]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Fig. 23D</figureCitation>
). The female endogyne has copulatory ducts (CD) larger than spermathecae (
<figureCitation id="21C54D99FFCEFFFCFB150BA8FAF3DCA9" box="[1215,1318,908,935]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="54" pageNumber="55">Fig. 23E</figureCitation>
) (vs in
<taxonomicName id="7EFE2A9FFFCEFFFCFF170B95FEBCDCC4" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[189,361,944,970]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="silvadavilae">
<emphasis id="8B8A8D0EFFCEFFFCFF170B95FEBCDCC4" box="[189,361,944,970]" italics="true" pageId="54" pageNumber="55">Z. silvadavilae</emphasis>
</taxonomicName>
, the LL are converging posteriorly, and the MS is triangular,
<figureCitation id="21C54D99FFCEFFFCFB980B94FB48DCC5" box="[1074,1181,944,971]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="54" pageNumber="55">Fig. 25D</figureCitation>
, and the endogyne has CD and spermathecae equal in size,
<figureCitation id="21C54D99FFCEFFFCFD3F0BF0FD2BDCE1" box="[661,766,980,1007]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9006231; CAS); D–E = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="54" pageNumber="55">Fig. 25E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFCEFFFCFF170C32FAA2DBA8" pageId="54" pageNumber="55" type="etymology">
<paragraph id="B941511CFFCEFFFCFF170C32FE91DB3E" blockId="54.[189,324,1046,1072]" box="[189,324,1046,1072]" pageId="54" pageNumber="55">
<heading id="E209E670FFCEFFFCFF170C32FE91DB3E" bold="true" box="[189,324,1046,1072]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="8B8A8D0EFFCEFFFCFF170C32FE91DB3E" bold="true" box="[189,324,1046,1072]" pageId="54" pageNumber="55">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170C61FAA2DBA8" blockId="54.[188,1400,1093,1191]" pageId="54" pageNumber="55">
The species name is a matronym in honor of Alma Saucedo Mejia, participant in the 2011 expedition that produced the female
<typeStatus id="6645EFBEFFCEFFFCFE1D0C4DFDC9DB8D" box="[439,540,1129,1155]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
(CASENT_9031271) that was a DNA voucher in
<bibRefCitation id="DD6F2CEDFFCEFFFCFBDD0C4DFAADDB8D" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[1143,1400,1128,1155]" pageId="54" pageNumber="55" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow
<emphasis id="8B8A8D0EFFCEFFFCFB4F0C4DFAF4DB8D" box="[1253,1313,1129,1155]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
(2015)
</bibRefCitation>
[DNA-DP-Lyc-voucher, Z-1, DP2014d]. She is also the discoverer of many
<collectingCountry id="C1E9118CFFCEFFFCFB9C0CA8FB11DBA8" box="[1078,1220,1164,1190]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
goblin spiders.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFCEFFFCFF170CEBFE20D85A" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="B941511CFFCEFFFCFF170CEBFEB9DBE6" blockId="54.[189,364,1230,1257]" box="[189,364,1230,1257]" pageId="54" pageNumber="55">
<heading id="E209E670FFCEFFFCFF170CEBFEB9DBE6" bold="true" box="[189,364,1230,1257]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="8B8A8D0EFFCEFFFCFF170CEBFEB9DBE6" bold="true" box="[189,364,1230,1257]" pageId="54" pageNumber="55">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF400CDAFE89DA16" blockId="54.[189,1398,1278,1376]" box="[234,348,1278,1304]" pageId="54" pageNumber="55">
<emphasis id="8B8A8D0EFFCEFFFCFF400CDAFE89DA16" bold="true" box="[234,348,1278,1304]" pageId="54" pageNumber="55">
<typeStatus id="6645EFBEFFCEFFFCFF400CDAFE89DA16" box="[234,348,1278,1304]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170D05FA9FDA51" blockId="54.[189,1398,1278,1376]" pageId="54" pageNumber="55">
<materialsCitation id="09965B41FFCEFFFCFF170D05FA9FDA51" collectingDate="1997-02-04" collectingDateMax="1997-02-07" collectingDateMin="1997-02-04" collectionCode="FMNH" collectorName="R. F. l'Ankarafantsika &amp; S. Goodman" country="Madagascar" elevation="160" latitude="-16.338333" location="Mahajunga Province" longLatPrecision="127" longitude="46.793335" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="C1E9118CFFCEFFFCFF170D05FE47DA35" box="[189,402,1313,1339]" name="Madagascar" pageId="54" pageNumber="55">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFCEFFFCFE1D0D05FD1DDA35" bold="true" box="[439,712,1313,1339]" pageId="54" pageNumber="55">
<location id="BC2107C7FFCEFFFCFE1D0D05FD1DDA35" LSID="urn:lsid:plazi:treatment:3157E00AFFCEFFF0FDDD092CFDB7DA9B:BC2107C7FFCEFFFCFE1D0D05FD1DDA35" box="[439,712,1313,1339]" country="Madagascar" latitude="-16.338333" longLatPrecision="127" longitude="46.793335" name="Mahajunga Province" pageId="54" pageNumber="55">Mahajunga Province</location>
</emphasis>
•
<specimenCount id="AFF89A95FFCEFFFCFD430D06FCD7DA32" box="[745,770,1314,1340]" count="1" pageId="54" pageNumber="55" type="male">♂</specimenCount>
;
<collectorName id="140B34CAFFCEFFFCFCB80D06FBFCDA35" box="[786,1065,1313,1340]" pageId="54" pageNumber="55">R. F. l’Ankarafantsika</collectorName>
,
<locationDeviation id="A8F3483EFFCEFFFCFB920D05FAA7DA35" box="[1080,1394,1313,1340]" location="Ampijaroa" pageId="54" pageNumber="55">
<quantity id="7E06FCF9FFCEFFFCFB920D05FBADDA35" box="[1080,1144,1313,1340]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="54" pageNumber="55" unit="km" value="5.0">5 km</quantity>
SSW of Ampijaroa
</locationDeviation>
;
<geoCoordinate id="DCCA37DBFFCEFFFCFF170D67FEECDA51" box="[189,313,1347,1375]" degrees="16" direction="south" minutes="20.3" orientation="latitude" pageId="54" pageNumber="55" precision="92" value="-16.338333">16°20.3′ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFCEFFFCFEEC0D67FE13DA51" box="[326,454,1347,1375]" degrees="46" direction="east" minutes="47.6" orientation="longitude" pageId="54" pageNumber="55" precision="92" value="46.793335">46°47.6′ E</geoCoordinate>
;
<elevation id="32D3B62FFFCEFFFCFE780D61FD89DA51" box="[466,604,1349,1375]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="54" pageNumber="55" unit="m" value="160.0">
<quantity id="7E06FCF9FFCEFFFCFE780D61FDC9DA51" box="[466,540,1349,1375]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="54" pageNumber="55" unit="m" value="160.0">160 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFCEFFFCFDC00D61FCC3DA51" box="[618,790,1349,1375]" pageId="54" pageNumber="55" value="1997-02-04" valueMax="1997-02-07" valueMin="1997-02-04">
<collectingDate id="DD048E34FFCEFFFCFDC00D61FCC3DA51" box="[618,790,1349,1375]" pageId="54" pageNumber="55" value="1997-02-04" valueMax="1997-02-07" valueMin="1997-02-04">4–7 Feb. 1997</collectingDate>
</date>
;
<collectorName id="140B34CAFFCEFFFCFC880D61FC60DA51" box="[802,949,1349,1375]" pageId="54" pageNumber="55">S. Goodman</collectorName>
leg.; CASENT_9082245;
<collectionCode id="DFEFC9D9FFCEFFFCFB470D61FA93DA51" box="[1261,1350,1349,1375]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="54" pageNumber="55" type="Museum">FMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF400DA3FEB2DAAF" blockId="54.[188,1399,1415,1691]" box="[234,359,1415,1441]" pageId="54" pageNumber="55">
<emphasis id="8B8A8D0EFFCEFFFCFF400DA3FEB2DAAF" bold="true" box="[234,359,1415,1441]" pageId="54" pageNumber="55">
<typeStatus id="6645EFBEFFCEFFFCFF400DA3FEB2DAAF" box="[234,359,1415,1441]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170D8FFC02D995" blockId="54.[188,1399,1415,1691]" pageId="54" pageNumber="55">
<materialsCitation id="09965B41FFCEFFFCFF170D8FFAA2DACB" box="[189,1399,1450,1477]" collectingDate="1997-02-04" collectingDateMax="1997-02-07" collectingDateMin="1997-02-04" collectionCode="FMNH" collectorName="R. F. l'Ankarafantsika &amp; S. Goodman" country="Madagascar" elevation="160" latitude="-16.338333" location="Mahajunga Province" longLatPrecision="127" longitude="46.793335" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" stateProvince="Mahajunga Province" typeStatus="paratype">
<collectingCountry id="C1E9118CFFCEFFFCFF170D8FFE58DACB" box="[189,397,1451,1477]" name="Madagascar" pageId="54" pageNumber="55">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFCEFFFCFE0C0D8EFD7FDACA" bold="true" box="[422,682,1450,1476]" pageId="54" pageNumber="55">
<collectingRegion id="7B3A9FFEFFCEFFFCFE0C0D8EFD7FDACA" box="[422,682,1450,1476]" pageId="54" pageNumber="55">Mahajunga Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFCEFFFCFD6A0D8FFD38DACB" box="[704,749,1451,1477]" count="1" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
; same data as for holotype; CASENT_9082245;
<collectionCode id="DFEFC9D9FFCEFFFCFA880D8FFAA2DACB" box="[1314,1399,1451,1477]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="54" pageNumber="55" type="Museum">FMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFCEFFFCFF7A0DEBFAFDD93E" collectingDate="2009-01-26" collectingDateMax="2009-01-28" collectingDateMin="2009-01-26" collectionCode="CAS" collectorName="C. Griswold &amp; A. Saucedo &amp; H. Wood &amp; Polotow D. &amp; Carmichael A. &amp; Griswold C. E." country="Madagascar" county="Ampijoroa" location="Lake Ravelobe" municipality="National Park Ankarafantsika" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" stateProvince="Mahajunga Province" typeStatus="paratype">
<specimenCount id="AFF89A95FFCEFFFCFF7A0DEBFF2ADAE7" box="[208,255,1487,1513]" count="1" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
;
<collectingCounty id="50202990FFCEFFFCFEA70DEAFE45DAE6" box="[269,400,1486,1512]" pageId="54" pageNumber="55">Ampijoroa</collectingCounty>
,
<collectingMunicipality id="5925CB66FFCEFFFCFE340DEAFCD7DAE6" box="[414,770,1486,1512]" pageId="54" pageNumber="55">National Park Ankarafantsika</collectingMunicipality>
, near
<location id="BC2107C7FFCEFFFCFCE10DEAFBD7DAE6" LSID="urn:lsid:plazi:treatment:3157E00AFFCEFFF0FDDD092CFDB7DA9B:BC2107C7FFCEFFFCFCE10DEAFBD7DAE6" box="[843,1026,1486,1512]" country="Madagascar" county="Ampijoroa" municipality="National Park Ankarafantsika" name="Lake Ravelobe" pageId="54" pageNumber="55" stateProvince="Mahajunga Province">Lake Ravelobe</location>
; 16°17.48′7″ S, 46°48.50′7″ E;
<date id="CD4077DCFFCEFFFCFF170DD6FE55D902" box="[189,384,1522,1549]" pageId="54" pageNumber="55" value="2009-01-26" valueMax="2009-01-28" valueMin="2009-01-26">
<collectingDate id="DD048E34FFCEFFFCFF170DD6FE55D902" box="[189,384,1522,1549]" pageId="54" pageNumber="55" value="2009-01-26" valueMax="2009-01-28" valueMin="2009-01-26">26–28 Jan. 2009</collectingDate>
</date>
;
<collectorName id="140B34CAFFCEFFFCFE210DD6FDC9D902" box="[395,540,1522,1548]" pageId="54" pageNumber="55">C. Griswold</collectorName>
,
<collectorName id="140B34CAFFCEFFFCFD8F0DD6FD7ED902" box="[549,683,1522,1548]" pageId="54" pageNumber="55">A. Saucedo</collectorName>
and
<collectorName id="140B34CAFFCEFFFCFD490DD7FC9FD902" box="[739,842,1522,1549]" pageId="54" pageNumber="55">H. Wood</collectorName>
leg.; DNA-DP-Lyc-voucher,
<taxonomicName id="7EFE2A9FFFCEFFFCFB090DD6FABAD902" box="[1187,1391,1522,1549]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="8B8A8D0EFFCEFFFCFB090DD6FAE4D902" box="[1187,1329,1522,1548]" italics="true" pageId="54" pageNumber="55">Zorodictyna</emphasis>
sp. 1
</taxonomicName>
, in
<collectorName id="140B34CAFFCEFFFCFF710E32FEB6D93E" box="[219,355,1558,1584]" pageId="54" pageNumber="55">Polotow D.</collectorName>
,
<collectorName id="140B34CAFFCEFFFCFEDA0E32FDCFD93E" box="[368,538,1558,1584]" pageId="54" pageNumber="55">Carmichael A.</collectorName>
&amp;
<collectorName id="140B34CAFFCEFFFCFDE80E32FD3ED93E" box="[578,747,1558,1584]" pageId="54" pageNumber="55">Griswold C.E.</collectorName>
2015; Z-1; DP2014d; CASENT_9031271;
<collectionCode id="DFEFC9D9FFCEFFFCFB450E32FAFDD93E" box="[1263,1320,1558,1584]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFCEFFFCFAE80E32FC06D995" collectingDate="2001-04-02" collectingDateMax="2001-04-08" collectingDateMin="2001-04-02" collectionCode="CAS" collectorName="de Tsimaloto &amp; J. J. Rafanomezantsoa" country="Madagascar" elevation="135" latitude="-16.061388" location="Foret de Tsimaloto" longLatPrecision="21" longitude="46.143612" municipality="Parc National d'Ankarafantsika" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" stateProvince="Mahajunga Province" typeStatus="paratype">
<specimenCount id="AFF89A95FFCEFFFCFAE80E32FABAD93E" box="[1346,1391,1558,1584]" count="1" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="5925CB66FFCEFFFCFF170E1EFDECD95D" box="[189,569,1593,1620]" pageId="54" pageNumber="55">Parc National d’Ankarafantsika</collectingMunicipality>
,
<location id="BC2107C7FFCEFFFCFDED0E1EFCF9D95D" LSID="urn:lsid:plazi:treatment:3157E00AFFCEFFF0FDDD092CFDB7DA9B:BC2107C7FFCEFFFCFDED0E1EFCF9D95D" box="[583,812,1593,1620]" country="Madagascar" latitude="-16.061388" longLatPrecision="21" longitude="46.143612" municipality="Parc National d'Ankarafantsika" name="Foret de Tsimaloto" pageId="54" pageNumber="55" stateProvince="Mahajunga Province">
Forêt
<collectorName id="140B34CAFFCEFFFCFD240E1DFCF9D95D" box="[654,812,1593,1619]" pageId="54" pageNumber="55">de Tsimaloto</collectorName>
</location>
,
<locationDeviation id="A8F3483EFFCEFFFCFC930E1EFB35D95D" box="[825,1248,1593,1620]" location="Tsaramandroso" pageId="54" pageNumber="55">
<quantity id="7E06FCF9FFCEFFFCFC930E1EFC49D95D" box="[825,924,1593,1620]" metricMagnitude="4" metricUnit="m" metricValue="1.83" pageId="54" pageNumber="55" unit="km" value="18.3">18.3 km</quantity>
46° NE of Tsaramandroso
</locationDeviation>
;
<geoCoordinate id="DCCA37DBFFCEFFFCFB470E1CFAA4D95A" box="[1261,1393,1592,1620]" degrees="16" direction="south" minutes="3" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="41" value="-16.061388">16°3′41″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFCEFFFCFF170E78FE95D976" box="[189,320,1628,1656]" degrees="46" direction="east" minutes="8" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="37" value="46.143612">46°8′37″ E</geoCoordinate>
;
<elevation id="32D3B62FFFCEFFFCFEE10E79FE07D979" box="[331,466,1629,1656]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="54" pageNumber="55" unit="m" value="135.0">
<quantity id="7E06FCF9FFCEFFFCFEE10E79FE46D976" box="[331,403,1629,1656]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="54" pageNumber="55" unit="m" value="135.0">135 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFCEFFFCFE740E79FD50D979" box="[478,645,1629,1656]" pageId="54" pageNumber="55" value="2001-04-02" valueMax="2001-04-08" valueMin="2001-04-02">
<collectingDate id="DD048E34FFCEFFFCFE740E79FD50D979" box="[478,645,1629,1656]" pageId="54" pageNumber="55" value="2001-04-02" valueMax="2001-04-08" valueMin="2001-04-02">2–8 Apr. 2001</collectingDate>
</date>
;
<collectorName id="140B34CAFFCEFFFCFD3A0E7AFC59D979" box="[656,908,1629,1656]" pageId="54" pageNumber="55">J.J. Rafanomezantsoa</collectorName>
<emphasis id="8B8A8D0EFFCEFFFCFC380E7AFC1ED979" box="[914,971,1629,1655]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
leg.; general collecting, day, tropical dry forest; [epigyne illustrated by CEG]; CASENT_9002829;
<collectionCode id="DFEFC9D9FFCEFFFCFC3C0EA5FC06D995" box="[918,979,1665,1691]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170EE7FE2CD9D3" blockId="54.[189,505,1731,1757]" box="[189,505,1731,1757]" pageId="54" pageNumber="55">
<heading id="E209E670FFCEFFFCFF170EE7FE2CD9D3" bold="true" box="[189,505,1731,1757]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="8B8A8D0EFFCEFFFCFF170EE7FE2CD9D3" bold="true" box="[189,505,1731,1757]" pageId="54" pageNumber="55">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170ED6FE20D85A" blockId="54.[189,1399,1778,1876]" pageId="54" pageNumber="55">
<materialsCitation id="09965B41FFCEFFFCFF170ED6FE24D85A" collectingDate="2009-01-26" collectingDateMax="2009-01-28" collectingDateMin="2009-01-26" collectionCode="CAS" collectorName="C. Griswold &amp; A. Saucedo &amp; H. Wood" country="Madagascar" county="Ampijoroa" latitude="-16.29145" location="Lake Ravelobe" longLatPrecision="1" longitude="46.80845" municipality="National Park Ankarafantsika" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" stateProvince="Mahajanga">
<collectingCountry id="C1E9118CFFCEFFFCFF170ED6FE5BD802" box="[189,398,1778,1804]" name="Madagascar" pageId="54" pageNumber="55">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFCEFFFCFE070ED6FD67D802" bold="true" box="[429,690,1778,1804]" pageId="54" pageNumber="55">
<collectingRegion id="7B3A9FFEFFCEFFFCFE070ED6FD67D802" box="[429,690,1778,1804]" country="Madagascar" name="Mahajanga" pageId="54" pageNumber="55">Mahajanga Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFCEFFFCFD670ED6FD2FD803" box="[717,762,1778,1805]" count="1" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
;
<collectingCounty id="50202990FFCEFFFCFCAD0ED6FC5CD802" box="[775,905,1778,1804]" pageId="54" pageNumber="55">Ampijoroa</collectingCounty>
,
<collectingMunicipality id="5925CB66FFCEFFFCFC3C0ED6FB20D802" box="[918,1269,1778,1804]" pageId="54" pageNumber="55">National Park Ankarafantsika</collectingMunicipality>
, near
<location id="BC2107C7FFCEFFFCFA960ED6FEE5D83E" LSID="urn:lsid:plazi:treatment:3157E00AFFCEFFF0FDDD092CFDB7DA9B:BC2107C7FFCEFFFCFA960ED6FEE5D83E" country="Madagascar" county="Ampijoroa" latitude="-16.29145" longLatPrecision="1" longitude="46.80845" municipality="National Park Ankarafantsika" name="Lake Ravelobe" pageId="54" pageNumber="55" stateProvince="Mahajanga">Lake Ravelobe</location>
;
<geoCoordinate id="DCCA37DBFFCEFFFCFE970F30FE0CD83E" box="[317,473,1812,1840]" degrees="16" direction="south" minutes="17.487" orientation="latitude" pageId="54" pageNumber="55" precision="1" value="-16.29145">16°17.487′ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFCEFFFCFE4D0F30FD53D83E" box="[487,646,1812,1840]" degrees="46" direction="east" minutes="48.507" orientation="longitude" pageId="54" pageNumber="55" precision="1" value="46.80845">46°48.507′ E</geoCoordinate>
;
<date id="CD4077DCFFCEFFFCFD3E0F32FC88D83E" box="[660,861,1814,1840]" pageId="54" pageNumber="55" value="2009-01-26" valueMax="2009-01-28" valueMin="2009-01-26">
<collectingDate id="DD048E34FFCEFFFCFD3E0F32FC88D83E" box="[660,861,1814,1840]" pageId="54" pageNumber="55" value="2009-01-26" valueMax="2009-01-28" valueMin="2009-01-26">26–28 Jan. 2009</collectingDate>
</date>
;
<collectorName id="140B34CAFFCEFFFCFCC00F32FC2AD83E" box="[874,1023,1814,1840]" pageId="54" pageNumber="55">C. Griswold</collectorName>
,
<collectorName id="140B34CAFFCEFFFCFBA00F32FB46D83E" box="[1034,1171,1814,1840]" pageId="54" pageNumber="55">A. Saucedo</collectorName>
and
<collectorName id="140B34CAFFCEFFFCFB650F32FAEFD83E" box="[1231,1338,1814,1840]" pageId="54" pageNumber="55">H. Wood</collectorName>
leg.; CASENT_9031270;
<collectionCode id="DFEFC9D9FFCEFFFCFE1E0F1EFE24D85A" box="[436,497,1850,1876]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFCEFFF0FF170F5FFC8DDBF1" lastPageId="58" lastPageNumber="59" pageId="54" pageNumber="55" type="description">
<paragraph id="B941511CFFCEFFFCFF170F5FFE9BD89B" blockId="54.[189,334,1915,1941]" box="[189,334,1915,1941]" pageId="54" pageNumber="55">
<heading id="E209E670FFCEFFFCFF170F5FFE9BD89B" bold="true" box="[189,334,1915,1941]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="8B8A8D0EFFCEFFFCFF170F5FFE9BD89B" bold="true" box="[189,334,1915,1941]" pageId="54" pageNumber="55">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFCEFFFCFF400F8FFCD6D8CB" blockId="54.[189,1189,1962,2025]" box="[234,771,1962,1989]" pageId="54" pageNumber="55">
<emphasis id="8B8A8D0EFFCEFFFCFF400F8FFE48D8CB" bold="true" box="[234,413,1963,1989]" pageId="54" pageNumber="55">
Male
<typeStatus id="6645EFBEFFCEFFFCFE9B0F8FFE48D8CB" box="[305,413,1963,1989]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
</emphasis>
(CASENT_9082245; FMNH)
</paragraph>
<paragraph id="B941511CFFCEFFFCFF170FEAFB70D8E7" blockId="54.[189,1189,1962,2025]" box="[189,1189,1998,2025]" pageId="54" pageNumber="55">MEASUREMENTS. Total length: 21.00. Carapace: length 12.00, width 9.80, height 5.70.</paragraph>
<paragraph id="B941511CFFCFFFFDFF17092DFBBBDE4B" blockId="55.[189,1399,265,326]" pageId="55" pageNumber="56">
COLORATION (
<figureCitation id="21C54D99FFCFFFFDFECC092DFE2DDE2D" box="[358,504,265,291]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="55" pageNumber="56">Fig. 22A–B</figureCitation>
). Dorsal and lateral sides of cephalothorax, abdomen and legs unicolorous grey-brown, venter of sternum, mouthparts, coxae and trochanters yellow white.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170948FB8ADE88" blockId="55.[189,1119,364,391]" box="[189,1119,364,391]" pageId="55" pageNumber="56">
STERNUM (
<figureCitation id="21C54D99FFCFFFFDFE970948FE7DDE88" box="[317,424,364,391]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="55" pageNumber="56">Fig. 22B</figureCitation>
). Oval with slightly sinuous margins; 4.50 wide, 5.60 long.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170989FDF7DEC9" blockId="55.[189,546,428,455]" box="[189,546,428,455]" pageId="55" pageNumber="56">CHILUM. 0.25 high, 0.85 wide.</paragraph>
<paragraph id="B941511CFFCFFFFDFF1709C9FCCADD24" blockId="55.[188,1399,493,554]" pageId="55" pageNumber="56">EYES. ALE 0.28; AME 0.28; PLE 0.35; PME 0.30; ALE–AME 0.13; AME–AME 0.70; PLE–PME 0.45; PME–PME 0.28; MOQ: AW 0.73, PW 0.83, L 0.80.</paragraph>
<paragraph id="B941511CFFCFFFFDFF170A75FDC7DD65" blockId="55.[189,530,592,619]" box="[189,530,592,619]" pageId="55" pageNumber="56">CLYPEUS. With 18 long setae.</paragraph>
<paragraph id="B941511CFFCFFFFDFF170AB5FACFDDC0" blockId="55.[189,1399,657,718]" pageId="55" pageNumber="56">LEGS. Trochanters deeply notched. Unpaired claws smooth, rudimentary on legs III and IV. Strong scopulae beneath all tarsi, apices of metatarsi III and IV, and apical halves of metatarsi I and II.</paragraph>
<paragraph id="B941511CFFCFFFFDFF170AD1FE4FDC01" blockId="55.[189,410,757,783]" box="[189,410,757,783]" pageId="55" pageNumber="56">Leg measurements</paragraph>
<paragraph id="B941511CFFCFFFFDFEA00B3BFCF1DCD7" pageId="55" pageNumber="56">
<table id="CBFEA3BCFFCF0035FF610B3BFCF1DCD7" box="[203,804,799,985]" gridcols="7" gridrows="5" pageId="55" pageNumber="56" rowsContinueIn="55.[203,988,1066,1252]">
<tr id="07CE535EFFCF0035FF610B3BFCF1DC37" box="[203,804,799,825]" gridrow="0" pageId="55" pageNumber="56" rowspan-0="1">
<th id="441F3A22FFCF0035FEA00B3BFE99DC37" box="[266,332,799,825]" gridcol="1" gridrow="0" pageId="55" pageNumber="56">F</th>
<th id="441F3A22FFCF0035FEC50B3BFE77DC37" box="[367,418,799,825]" gridcol="2" gridrow="0" pageId="55" pageNumber="56">P</th>
<th id="441F3A22FFCF0035FE6E0B3BFDD3DC37" box="[452,518,799,825]" gridcol="3" gridrow="0" pageId="55" pageNumber="56">T</th>
<th id="441F3A22FFCF0035FD820B3BFDBFDC37" box="[552,618,799,825]" gridcol="4" gridrow="0" pageId="55" pageNumber="56">Mt</th>
<th id="441F3A22FFCF0035FD270B3BFD15DC37" box="[653,704,799,825]" gridcol="5" gridrow="0" pageId="55" pageNumber="56">t</th>
<th id="441F3A22FFCF0035FD480B3BFCF1DC37" box="[738,804,799,825]" gridcol="6" gridrow="0" pageId="55" pageNumber="56">Total</th>
</tr>
<tr id="07CE535EFFCF0035FF610B63FCF1DC6C" box="[203,804,839,866]" gridrow="1" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610B63FF3FDC6C" box="[203,234,839,866]" gridcol="0" gridrow="1" pageId="55" pageNumber="56">I</th>
<td id="441F3A22FFCF0035FEA00B63FE99DC6C" box="[266,332,839,866]" gridcol="1" gridrow="1" pageId="55" pageNumber="56">10.20</td>
<td id="441F3A22FFCF0035FEC50B63FE77DC6C" box="[367,418,839,866]" gridcol="2" gridrow="1" pageId="55" pageNumber="56">5.10</td>
<td id="441F3A22FFCF0035FE6E0B63FDD3DC6C" box="[452,518,839,866]" gridcol="3" gridrow="1" pageId="55" pageNumber="56">11.50</td>
<td id="441F3A22FFCF0035FD820B63FDBFDC6C" box="[552,618,839,866]" gridcol="4" gridrow="1" pageId="55" pageNumber="56">11.10</td>
<td id="441F3A22FFCF0035FD270B63FD15DC6C" box="[653,704,839,866]" gridcol="5" gridrow="1" pageId="55" pageNumber="56">6.30</td>
<td id="441F3A22FFCF0035FD480B63FCF1DC6C" box="[738,804,839,866]" gridcol="6" gridrow="1" pageId="55" pageNumber="56">44.20</td>
</tr>
<tr id="07CE535EFFCF0035FF610B4BFCF1DC84" box="[203,804,879,906]" gridrow="2" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610B4BFF3FDC84" box="[203,234,879,906]" gridcol="0" gridrow="2" pageId="55" pageNumber="56">II</th>
<td id="441F3A22FFCF0035FEA00B4BFE99DC84" box="[266,332,879,906]" gridcol="1" gridrow="2" pageId="55" pageNumber="56">11.00</td>
<td id="441F3A22FFCF0035FEC50B4BFE77DC84" box="[367,418,879,906]" gridcol="2" gridrow="2" pageId="55" pageNumber="56">5.00</td>
<td id="441F3A22FFCF0035FE6E0B4BFDD3DC84" box="[452,518,879,906]" gridcol="3" gridrow="2" pageId="55" pageNumber="56">9.70</td>
<td id="441F3A22FFCF0035FD820B4BFDBFDC84" box="[552,618,879,906]" gridcol="4" gridrow="2" pageId="55" pageNumber="56">9.80</td>
<td id="441F3A22FFCF0035FD270B4BFD15DC84" box="[653,704,879,906]" gridcol="5" gridrow="2" pageId="55" pageNumber="56">5.00</td>
<td id="441F3A22FFCF0035FD480B4BFCF1DC84" box="[738,804,879,906]" gridcol="6" gridrow="2" pageId="55" pageNumber="56">40.50</td>
</tr>
<tr id="07CE535EFFCF0035FF610BB3FCF1DCBC" box="[203,804,919,946]" gridrow="3" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610BB3FF3FDCBC" box="[203,234,919,946]" gridcol="0" gridrow="3" pageId="55" pageNumber="56">III</th>
<td id="441F3A22FFCF0035FEA00BB3FE99DCBC" box="[266,332,919,946]" gridcol="1" gridrow="3" pageId="55" pageNumber="56">9.30</td>
<td id="441F3A22FFCF0035FEC50BB3FE77DCBC" box="[367,418,919,946]" gridcol="2" gridrow="3" pageId="55" pageNumber="56">4.10</td>
<td id="441F3A22FFCF0035FE6E0BB3FDD3DCBC" box="[452,518,919,946]" gridcol="3" gridrow="3" pageId="55" pageNumber="56">7.60</td>
<td id="441F3A22FFCF0035FD820BB3FDBFDCBC" box="[552,618,919,946]" gridcol="4" gridrow="3" pageId="55" pageNumber="56">9.40</td>
<td id="441F3A22FFCF0035FD270BB3FD15DCBC" box="[653,704,919,946]" gridcol="5" gridrow="3" pageId="55" pageNumber="56">4.60</td>
<td id="441F3A22FFCF0035FD480BB3FCF1DCBC" box="[738,804,919,946]" gridcol="6" gridrow="3" pageId="55" pageNumber="56">35.00</td>
</tr>
<tr id="07CE535EFFCF0035FF610B9AFCF1DCD7" box="[203,804,958,985]" gridrow="4" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610B9AFF3FDCD7" box="[203,234,958,985]" gridcol="0" gridrow="4" pageId="55" pageNumber="56">IV</th>
<td id="441F3A22FFCF0035FEA00B9AFE99DCD7" box="[266,332,958,985]" gridcol="1" gridrow="4" pageId="55" pageNumber="56">11.90</td>
<td id="441F3A22FFCF0035FEC50B9AFE77DCD7" box="[367,418,958,985]" gridcol="2" gridrow="4" pageId="55" pageNumber="56">4.70</td>
<td id="441F3A22FFCF0035FE6E0B9AFDD3DCD7" box="[452,518,958,985]" gridcol="3" gridrow="4" pageId="55" pageNumber="56">10.90</td>
<td id="441F3A22FFCF0035FD820B9AFDBFDCD7" box="[552,618,958,985]" gridcol="4" gridrow="4" pageId="55" pageNumber="56">15.20</td>
<td id="441F3A22FFCF0035FD270B9AFD15DCD7" box="[653,704,958,985]" gridcol="5" gridrow="4" pageId="55" pageNumber="56">6.00</td>
<td id="441F3A22FFCF0035FD480B9AFCF1DCD7" box="[738,804,958,985]" gridcol="6" gridrow="4" pageId="55" pageNumber="56">48.70</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170BDBFC6AD991" blockId="55.[189,301,1023,1049]" lastBlockId="55.[187,1400,1290,1696]" pageId="55" pageNumber="56">
Spination MALE PALP (
<figureCitation id="21C54D99FFCFFFFDFEFA0D2EFE33DA2A" box="[336,486,1290,1317]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="55" pageNumber="56">Fig. 23 A–C</figureCitation>
). Tibia VTA apical, curved and blunt-tipped, RTA triangular, sharply pointed, located behind this dorso-retrolateral excavation to accommodate retrobasal cymbial extension (CE); cymbium oval, apical extremity narrowly rounded, without dorsal patch of chemosensitive setae, with protruding retrolateral paracymbial process (PP), retrobasal cymbial extension (CE) conical, extending at least ¼ the length of papal tibia; subtegulum large, filling most of cymbial concavity and extending to prolateral cymbial margin, with probasal broad ridge (SL) that interlocks with tegulum; tegulum rounded with rectangular process on prolateral margin (TL), which interlocks with subtegular ridge; embolus (E) proapical, broad and thick, with pointed apex, inner margin with twisted longitudinal ridge and triangular basomedian process (EBP) arising from embolus base; median apophysis (MA) strongly sclerotized, short, hooked, curved prolaterad; conductor (C) hyaline, fan shaped, narrow at base; paraconductor (Pc) large, cradling inner margin of E and extending past E apex; retroventral surface of tegulum with deep longitudinal groove that extends for two thirds of tegulum length.
</paragraph>
<paragraph id="B941511CFFCFFFFDFEA00C0EFC09DBED" pageId="55" pageNumber="56">
<table id="CBFEA3BCFFCF0035FF610C0EFC09DBEA" box="[203,988,1066,1252]" gridcols="5" gridrows="5" pageId="55" pageNumber="56" rowsContinueFrom="55.[203,804,799,985]">
<tr id="07CE535EFFCF0035FF610C0EFC09DB4A" box="[203,988,1066,1092]" gridrow="0" pageId="55" pageNumber="56" rowspan-0="1">
<th id="441F3A22FFCF0035FEA00C0EFEBBDB4A" box="[266,366,1066,1092]" gridcol="1" gridrow="0" pageId="55" pageNumber="56">F</th>
<th id="441F3A22FFCF0035FE300C0EFE35DB4A" box="[410,480,1066,1092]" gridcol="2" gridrow="0" pageId="55" pageNumber="56">P</th>
<th id="441F3A22FFCF0035FDBD0C0EFD37DB4A" box="[535,738,1066,1092]" gridcol="3" gridrow="0" pageId="55" pageNumber="56">T</th>
<th id="441F3A22FFCF0035FCBD0C0EFC09DB4A" box="[791,988,1066,1092]" gridcol="4" gridrow="0" pageId="55" pageNumber="56">Mt</th>
</tr>
<tr id="07CE535EFFCF0035FF610C76FC09DB63" box="[203,988,1106,1133]" gridrow="1" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610C76FF3FDB63" box="[203,234,1106,1133]" gridcol="0" gridrow="1" pageId="55" pageNumber="56">I</th>
<td id="441F3A22FFCF0035FEA00C76FEBBDB63" box="[266,366,1106,1133]" gridcol="1" gridrow="1" pageId="55" pageNumber="56">pl3d3rl2</td>
<td id="441F3A22FFCF0035FE300C76FE35DB63" box="[410,480,1106,1133]" gridcol="2" gridrow="1" pageId="55" pageNumber="56">pl1rl1</td>
<td id="441F3A22FFCF0035FDBD0C76FD37DB63" box="[535,738,1106,1133]" gridcol="3" gridrow="1" pageId="55" pageNumber="56">pl3d3rl3v2-2-2-2</td>
<td id="441F3A22FFCF0035FCBD0C76FC09DB63" box="[791,988,1106,1133]" gridcol="4" gridrow="1" pageId="55" pageNumber="56">pl4rl4v2-2-3</td>
</tr>
<tr id="07CE535EFFCF0035FF610C5EFC09DB9A" box="[203,988,1146,1172]" gridrow="2" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610C5EFF3FDB9A" box="[203,234,1146,1172]" gridcol="0" gridrow="2" pageId="55" pageNumber="56">II</th>
<td id="441F3A22FFCF0035FEA00C5EFEBBDB9A" box="[266,366,1146,1172]" gridcol="1" gridrow="2" pageId="55" pageNumber="56">pl3d3rl3</td>
<td id="441F3A22FFCF0035FE300C5EFE35DB9A" box="[410,480,1146,1172]" gridcol="2" gridrow="2" pageId="55" pageNumber="56">–</td>
<td id="441F3A22FFCF0035FDBD0C5EFD37DB9A" box="[535,738,1146,1172]" gridcol="3" gridrow="2" pageId="55" pageNumber="56">pl2d3rl2v2-2-2-2</td>
<td id="441F3A22FFCF0035FCBD0C5EFC09DB9A" box="[791,988,1146,1172]" gridcol="4" gridrow="2" pageId="55" pageNumber="56">pl3rl3v2-2-3</td>
</tr>
<tr id="07CE535EFFCF0035FF610C86FC09DBB2" box="[203,988,1186,1212]" gridrow="3" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610C86FF3FDBB2" box="[203,234,1186,1212]" gridcol="0" gridrow="3" pageId="55" pageNumber="56">III</th>
<td id="441F3A22FFCF0035FEA00C86FEBBDBB2" box="[266,366,1186,1212]" gridcol="1" gridrow="3" pageId="55" pageNumber="56">pl3d3rl3</td>
<td id="441F3A22FFCF0035FE300C86FE35DBB2" box="[410,480,1186,1212]" gridcol="2" gridrow="3" pageId="55" pageNumber="56">pl1rl1</td>
<td id="441F3A22FFCF0035FDBD0C86FD37DBB2" box="[535,738,1186,1212]" gridcol="3" gridrow="3" pageId="55" pageNumber="56">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFCF0035FCBD0C86FC09DBB2" box="[791,988,1186,1212]" gridcol="4" gridrow="3" pageId="55" pageNumber="56">pl4d1rl4v2-2-2</td>
</tr>
<tr id="07CE535EFFCF0035FF610CEDFC09DBEA" box="[203,988,1225,1252]" gridrow="4" pageId="55" pageNumber="56">
<th id="441F3A22FFCF0035FF610CEDFF3FDBEA" box="[203,234,1225,1252]" gridcol="0" gridrow="4" pageId="55" pageNumber="56">IV</th>
<td id="441F3A22FFCF0035FEA00CEDFEBBDBEA" box="[266,366,1225,1252]" gridcol="1" gridrow="4" pageId="55" pageNumber="56">pl3d3rl3</td>
<td id="441F3A22FFCF0035FE300CEDFE35DBEA" box="[410,480,1225,1252]" gridcol="2" gridrow="4" pageId="55" pageNumber="56">pl1rl1</td>
<td id="441F3A22FFCF0035FDBD0CEDFD37DBEA" box="[535,738,1225,1252]" gridcol="3" gridrow="4" pageId="55" pageNumber="56">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFCF0035FCBD0CEDFC09DBEA" box="[791,988,1225,1252]" gridcol="4" gridrow="4" pageId="55" pageNumber="56">pl4rl4v1-1-1-2-3</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFCFFFFDFF400EE2FB66D80C" blockId="55.[189,1203,1733,1794]" pageId="55" pageNumber="56">
<emphasis id="8B8A8D0EFFCFFFFDFF400EE2FE68D9EE" bold="true" box="[234,445,1734,1760]" pageId="55" pageNumber="56">
Female
<typeStatus id="6645EFBEFFCFFFFDFEE70EE2FE68D9EE" box="[333,445,1734,1760]" pageId="55" pageNumber="56" type="paratype">paratype</typeStatus>
</emphasis>
(CASENT_9082245; FMNH; except as otherwise indicated) MEASUREMENTS. Total length: 30.00. Carapace: length 14.00, width 11.30, height 4.40.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170F0DFAD4D86B" blockId="55.[189,1398,1833,1894]" pageId="55" pageNumber="56">
COLOUR (
<figureCitation id="21C54D99FFCFFFFDFE9F0F0DFE1FD84D" box="[309,458,1833,1859]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="55" pageNumber="56">Fig. 22C–D</figureCitation>
). As in male: dorsal and lateral surfaces of cephalothorax, abdomen and legs unicolorous grey-brown; venter of sternum, mouthparts, coxae and trochanters yellow white.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170FA8FBE8D8A8" blockId="55.[189,1085,1932,1958]" box="[189,1085,1932,1958]" pageId="55" pageNumber="56">
STERNUM (
<figureCitation id="21C54D99FFCFFFFDFE970FA8FE7FD8A8" box="[317,426,1932,1958]" captionStart="Fig" captionStartId="56.[189,231,1928,1954]" captionTargetBox="[189,1398,1121,1890]" captionTargetId="figure-176@56.[189,1398,1121,1890]" captionTargetPageId="56" captionText="Fig. 22. Zorodictyna almae sp. nov., male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH). A, C. Dorsal view. B, D. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="55" pageNumber="56">Fig. 22D</figureCitation>
). Oval with slightly sinuous sides; 4.90 wide, 6.40 long.
</paragraph>
<paragraph id="B941511CFFCFFFFDFF170FEAFDF7D8E6" blockId="55.[189,546,1998,2025]" box="[189,546,1998,2025]" pageId="55" pageNumber="56">CHILUM. 0.25 high, 1.08 wide.</paragraph>
<paragraph id="B941511CFFC0FFF2FF17092DFCCADE48" blockId="56.[188,1399,265,327]" pageId="56" pageNumber="57">EYES. ALE 0.33; AME 0.33; PLE 0.31; PME 0.29; ALE–AME 0.20; AME–AME 0.75; PLE–PME 0.55; PME–PME 0.35; MOQ: AW 0.78, PW 0.93, L 0.78.</paragraph>
<paragraph id="B941511CFFC0FFF2FF17094AFDCDDE86" blockId="56.[189,536,366,392]" box="[189,536,366,392]" pageId="56" pageNumber="57">CLYPEUS. With ten long setae.</paragraph>
<paragraph id="B941511CFFC0FFF2FF17098BFA86DEE3" blockId="56.[189,1399,431,493]" pageId="56" pageNumber="57">LEGS. Trochanters deeply notched. Unpaired claws smooth, rudimentary on legs III and IV. Strong scopulae beneath all tarsi, apex of metatarsus III, apical half of metatarsus II and all of metatarsus I.</paragraph>
<paragraph id="B941511CFFC0FFF2FF170A30FE4FDD20" blockId="56.[189,410,532,558]" box="[189,410,532,558]" pageId="56" pageNumber="57">Leg measurements</paragraph>
<paragraph id="B941511CFFC0FFF2FEA00A1AFCF1DDF9" pageId="56" pageNumber="57">
<table id="CBFEA3BCFFC00035FF610A19FCF1DDF9" box="[203,804,573,759]" gridcols="7" gridrows="5" pageId="56" pageNumber="57" rowsContinueIn="56.[203,912,840,1026]">
<tr id="07CE535EFFC00035FF610A19FCF1DD56" box="[203,804,573,600]" gridrow="0" pageId="56" pageNumber="57" rowspan-0="1">
<th id="441F3A22FFC00035FEA00A19FE99DD56" box="[266,332,573,600]" gridcol="1" gridrow="0" pageId="56" pageNumber="57">F</th>
<th id="441F3A22FFC00035FEC50A19FE77DD56" box="[367,418,573,600]" gridcol="2" gridrow="0" pageId="56" pageNumber="57">P</th>
<th id="441F3A22FFC00035FE6E0A19FDD0DD56" box="[452,517,573,600]" gridcol="3" gridrow="0" pageId="56" pageNumber="57">T</th>
<th id="441F3A22FFC00035FD820A19FDBFDD56" box="[552,618,573,600]" gridcol="4" gridrow="0" pageId="56" pageNumber="57">Mt</th>
<th id="441F3A22FFC00035FD270A19FD15DD56" box="[653,704,573,600]" gridcol="5" gridrow="0" pageId="56" pageNumber="57">t</th>
<th id="441F3A22FFC00035FD480A19FCF1DD56" box="[738,804,573,600]" gridcol="6" gridrow="0" pageId="56" pageNumber="57">Total</th>
</tr>
<tr id="07CE535EFFC00035FF610A41FCF1DD8E" box="[203,804,613,640]" gridrow="1" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610A41FF3FDD8E" box="[203,234,613,640]" gridcol="0" gridrow="1" pageId="56" pageNumber="57">I</th>
<td id="441F3A22FFC00035FEA00A41FE99DD8E" box="[266,332,613,640]" gridcol="1" gridrow="1" pageId="56" pageNumber="57">10.30</td>
<td id="441F3A22FFC00035FEC50A41FE77DD8E" box="[367,418,613,640]" gridcol="2" gridrow="1" pageId="56" pageNumber="57">5.20</td>
<td id="441F3A22FFC00035FE6E0A41FDD0DD8E" box="[452,517,613,640]" gridcol="3" gridrow="1" pageId="56" pageNumber="57">11.50</td>
<td id="441F3A22FFC00035FD820A41FDBFDD8E" box="[552,618,613,640]" gridcol="4" gridrow="1" pageId="56" pageNumber="57">11.10</td>
<td id="441F3A22FFC00035FD270A41FD15DD8E" box="[653,704,613,640]" gridcol="5" gridrow="1" pageId="56" pageNumber="57">6.30</td>
<td id="441F3A22FFC00035FD480A41FCF1DD8E" box="[738,804,613,640]" gridcol="6" gridrow="1" pageId="56" pageNumber="57">44.20</td>
</tr>
<tr id="07CE535EFFC00035FF610AA9FCF1DDA6" box="[203,804,653,680]" gridrow="2" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610AA9FF3FDDA6" box="[203,234,653,680]" gridcol="0" gridrow="2" pageId="56" pageNumber="57">II</th>
<td id="441F3A22FFC00035FEA00AA9FE99DDA6" box="[266,332,653,680]" gridcol="1" gridrow="2" pageId="56" pageNumber="57">9.50</td>
<td id="441F3A22FFC00035FEC50AA9FE77DDA6" box="[367,418,653,680]" gridcol="2" gridrow="2" pageId="56" pageNumber="57">4.90</td>
<td id="441F3A22FFC00035FE6E0AA9FDD0DDA6" box="[452,517,653,680]" gridcol="3" gridrow="2" pageId="56" pageNumber="57">7.20</td>
<td id="441F3A22FFC00035FD820AA9FDBFDDA6" box="[552,618,653,680]" gridcol="4" gridrow="2" pageId="56" pageNumber="57">6.80</td>
<td id="441F3A22FFC00035FD270AA9FD15DDA6" box="[653,704,653,680]" gridcol="5" gridrow="2" pageId="56" pageNumber="57">3.80</td>
<td id="441F3A22FFC00035FD480AA9FCF1DDA6" box="[738,804,653,680]" gridcol="6" gridrow="2" pageId="56" pageNumber="57">32.30</td>
</tr>
<tr id="07CE535EFFC00035FF610A91FCF1DDDE" box="[203,804,693,720]" gridrow="3" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610A91FF3FDDDE" box="[203,234,693,720]" gridcol="0" gridrow="3" pageId="56" pageNumber="57">III</th>
<td id="441F3A22FFC00035FEA00A91FE99DDDE" box="[266,332,693,720]" gridcol="1" gridrow="3" pageId="56" pageNumber="57">8.20</td>
<td id="441F3A22FFC00035FEC50A91FE77DDDE" box="[367,418,693,720]" gridcol="2" gridrow="3" pageId="56" pageNumber="57">4.20</td>
<td id="441F3A22FFC00035FE6E0A91FDD0DDDE" box="[452,517,693,720]" gridcol="3" gridrow="3" pageId="56" pageNumber="57">5.80</td>
<td id="441F3A22FFC00035FD820A91FDBFDDDE" box="[552,618,693,720]" gridcol="4" gridrow="3" pageId="56" pageNumber="57">7.20</td>
<td id="441F3A22FFC00035FD270A91FD15DDDE" box="[653,704,693,720]" gridcol="5" gridrow="3" pageId="56" pageNumber="57">3.80</td>
<td id="441F3A22FFC00035FD480A91FCF1DDDE" box="[738,804,693,720]" gridcol="6" gridrow="3" pageId="56" pageNumber="57">29.20</td>
</tr>
<tr id="07CE535EFFC00035FF610AF9FCF1DDF9" box="[203,804,733,759]" gridrow="4" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610AF9FF3FDDF9" box="[203,234,733,759]" gridcol="0" gridrow="4" pageId="56" pageNumber="57">IV</th>
<td id="441F3A22FFC00035FEA00AF9FE99DDF9" box="[266,332,733,759]" gridcol="1" gridrow="4" pageId="56" pageNumber="57">10.90</td>
<td id="441F3A22FFC00035FEC50AF9FE77DDF9" box="[367,418,733,759]" gridcol="2" gridrow="4" pageId="56" pageNumber="57">4.70</td>
<td id="441F3A22FFC00035FE6E0AF9FDD0DDF9" box="[452,517,733,759]" gridcol="3" gridrow="4" pageId="56" pageNumber="57">9.00</td>
<td id="441F3A22FFC00035FD820AF9FDBFDDF9" box="[552,618,733,759]" gridcol="4" gridrow="4" pageId="56" pageNumber="57">12.60</td>
<td id="441F3A22FFC00035FD270AF9FD15DDF9" box="[653,704,733,759]" gridcol="5" gridrow="4" pageId="56" pageNumber="57">4.80</td>
<td id="441F3A22FFC00035FD480AF9FCF1DDF9" box="[738,804,733,759]" gridcol="6" gridrow="4" pageId="56" pageNumber="57">42.00</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFC0FFF0FF170B3AFC47DED6" blockId="56.[189,301,798,824]" lastBlockId="58.[187,1400,265,473]" lastPageId="58" lastPageNumber="59" pageId="56" pageNumber="57">
Spination GENITALIA. Epigyne (
<typeStatus id="6645EFBEFFC2FFF0FE1A092DFDC0DE2D" box="[432,533,265,291]" pageId="58" pageNumber="59" type="paratype">paratype</typeStatus>
CASENT_9002829; CAS) (
<figureCitation id="21C54D99FFC2FFF0FCF7092DFC11DE2D" box="[861,964,265,291]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="58" pageNumber="59">Fig. 23D</figureCitation>
) transverse quadrangular median lobe (ML) plate with smooth margin; copulatory openings (CO) hidden beneath anterolateral corners of ML; lateral lobes (LL) convex, without lateral teeth or projections but with concavities along lateral margins of ML, LL straight and parallel medially, exposing rectangular area of median sector (MS). Endogyne (
<figureCitation id="21C54D99FFC2FFF0FF6D09BEFEFADEBA" box="[199,303,410,436]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; A–C = male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="58" pageNumber="59">Fig. 23E</figureCitation>
) with broad, simple, dark copulatory ducts (CD) larger than spermathecae (S), spermathecae with one half twist; fertilization ducts (FD) short and straight.
</paragraph>
<paragraph id="B941511CFFC0FFF2FEA00B6CFC5EDB0C" pageId="56" pageNumber="57">
<table id="CBFEA3BCFFC00035FF610B6CFC45DB0C" box="[203,912,840,1026]" gridcols="5" gridrows="5" pageId="56" pageNumber="57" rowsContinueFrom="56.[203,804,573,759]">
<tr id="07CE535EFFC00035FF610B6CFC45DC6C" box="[203,912,840,866]" gridrow="0" pageId="56" pageNumber="57" rowspan-0="1">
<th id="441F3A22FFC00035FEA00B6CFEBBDC6C" box="[266,366,840,866]" gridcol="1" gridrow="0" pageId="56" pageNumber="57">F</th>
<th id="441F3A22FFC00035FE390B6CFE0CDC6C" box="[403,473,840,866]" gridcol="2" gridrow="0" pageId="56" pageNumber="57">P</th>
<th id="441F3A22FFC00035FE570B6CFD7ADC6C" box="[509,687,840,866]" gridcol="3" gridrow="0" pageId="56" pageNumber="57">T</th>
<th id="441F3A22FFC00035FD750B6CFC45DC6C" box="[735,912,840,866]" gridcol="4" gridrow="0" pageId="56" pageNumber="57">Mt</th>
</tr>
<tr id="07CE535EFFC00035FF610B54FC45DC85" box="[203,912,880,907]" gridrow="1" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610B54FF3FDC85" box="[203,234,880,907]" gridcol="0" gridrow="1" pageId="56" pageNumber="57">I</th>
<td id="441F3A22FFC00035FEA00B54FEBBDC85" box="[266,366,880,907]" gridcol="1" gridrow="1" pageId="56" pageNumber="57">pl1d2</td>
<td id="441F3A22FFC00035FE390B54FE0CDC85" box="[403,473,880,907]" gridcol="2" gridrow="1" pageId="56" pageNumber="57">–</td>
<td id="441F3A22FFC00035FE570B54FD7ADC85" box="[509,687,880,907]" gridcol="3" gridrow="1" pageId="56" pageNumber="57">v2-2-2-2</td>
<td id="441F3A22FFC00035FD750B54FC45DC85" box="[735,912,880,907]" gridcol="4" gridrow="1" pageId="56" pageNumber="57">v2-2-2</td>
</tr>
<tr id="07CE535EFFC00035FF610BBCFC45DCBC" box="[203,912,920,946]" gridrow="2" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610BBCFF3FDCBC" box="[203,234,920,946]" gridcol="0" gridrow="2" pageId="56" pageNumber="57">II</th>
<td id="441F3A22FFC00035FEA00BBCFEBBDCBC" box="[266,366,920,946]" gridcol="1" gridrow="2" pageId="56" pageNumber="57">pl2d2</td>
<td id="441F3A22FFC00035FE390BBCFE0CDCBC" box="[403,473,920,946]" gridcol="2" gridrow="2" pageId="56" pageNumber="57">–</td>
<td id="441F3A22FFC00035FE570BBCFD7ADCBC" box="[509,687,920,946]" gridcol="3" gridrow="2" pageId="56" pageNumber="57">v2-2-2-2</td>
<td id="441F3A22FFC00035FD750BBCFC45DCBC" box="[735,912,920,946]" gridcol="4" gridrow="2" pageId="56" pageNumber="57">v2-2-2</td>
</tr>
<tr id="07CE535EFFC00035FF610BE4FC45DCD4" box="[203,912,960,986]" gridrow="3" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610BE4FF3FDCD4" box="[203,234,960,986]" gridcol="0" gridrow="3" pageId="56" pageNumber="57">III</th>
<td id="441F3A22FFC00035FEA00BE4FEBBDCD4" box="[266,366,960,986]" gridcol="1" gridrow="3" pageId="56" pageNumber="57">pl3d3rl4</td>
<td id="441F3A22FFC00035FE390BE4FE0CDCD4" box="[403,473,960,986]" gridcol="2" gridrow="3" pageId="56" pageNumber="57">pl1rl1</td>
<td id="441F3A22FFC00035FE570BE4FD7ADCD4" box="[509,687,960,986]" gridcol="3" gridrow="3" pageId="56" pageNumber="57">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFC00035FD750BE4FC45DCD4" box="[735,912,960,986]" gridcol="4" gridrow="3" pageId="56" pageNumber="57">pl4d1rl4v2-2-2</td>
</tr>
<tr id="07CE535EFFC00035FF610BC3FC45DB0C" box="[203,912,999,1026]" gridrow="4" pageId="56" pageNumber="57">
<th id="441F3A22FFC00035FF610BC3FF3FDB0C" box="[203,234,999,1026]" gridcol="0" gridrow="4" pageId="56" pageNumber="57">IV</th>
<td id="441F3A22FFC00035FEA00BC3FEBBDB0C" box="[266,366,999,1026]" gridcol="1" gridrow="4" pageId="56" pageNumber="57">pl1d3rl3</td>
<td id="441F3A22FFC00035FE390BC3FE0CDB0C" box="[403,473,999,1026]" gridcol="2" gridrow="4" pageId="56" pageNumber="57">pl1rl1</td>
<td id="441F3A22FFC00035FE570BC3FD7ADB0C" box="[509,687,999,1026]" gridcol="3" gridrow="4" pageId="56" pageNumber="57">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFC00035FD750BC3FC45DB0C" box="[735,912,999,1026]" gridcol="4" gridrow="4" pageId="56" pageNumber="57">pl5rl4v2-1-2-2</td>
</tr>
</table>
</paragraph>
<caption id="ED810194FFC0FFF2FF170FACFBE1D8E6" ID-DOI="http://doi.org/10.5281/zenodo.14001843" ID-Zenodo-Dep="14001843" httpUri="https://zenodo.org/record/14001843/files/figure.png" pageId="56" pageNumber="57" startId="56.[189,231,1928,1954]" targetBox="[189,1398,1121,1890]" targetPageId="56">
<paragraph id="B941511CFFC0FFF2FF170FACFBE1D8E6" blockId="56.[187,1399,1928,2025]" pageId="56" pageNumber="57">
<emphasis id="8B8A8D0EFFC0FFF2FF170FACFEC3D8AC" bold="true" box="[189,278,1928,1954]" pageId="56" pageNumber="57">Fig. 22.</emphasis>
<taxonomicName id="7EFE2A9FFFC0FFF2FEB10FACFE21D8AC" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[283,500,1928,1954]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC0FFF2FEB10FACFE21D8AC" box="[283,500,1928,1954]" italics="true" pageId="56" pageNumber="57">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC0FFF2FE560FADFD83D8AD" box="[508,598,1929,1955]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
, male and female habitus; A–B = male holotype (CASENT_9082245; FMNH); C–D = female paratype (CASENT_9082245; FMNH).
<emphasis id="8B8A8D0EFFC0FFF2FC190F8FFC39D8CB" bold="true" box="[947,1004,1963,1989]" pageId="56" pageNumber="57">A, C</emphasis>
. Dorsal view.
<emphasis id="8B8A8D0EFFC0FFF2FB3C0F8FFB1BD8CB" bold="true" box="[1174,1230,1963,1989]" pageId="56" pageNumber="57">B, D</emphasis>
. Ventral view. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme.
</paragraph>
</caption>
<caption id="ED810194FFC1FFF3FF170E40FAC7D87A" ID-DOI="http://doi.org/10.5281/zenodo.14001845" ID-Zenodo-Dep="14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="57" pageNumber="58" startId="57.[189,232,1636,1662]" targetBox="[219,1348,279,1580]" targetPageId="57">
<paragraph id="B941511CFFC1FFF3FF170E40FAC7D87A" blockId="57.[187,1400,1636,1909]" pageId="57" pageNumber="58">
<emphasis id="8B8A8D0EFFC1FFF3FF170E40FEC9D970" bold="true" box="[189,284,1636,1662]" pageId="57" pageNumber="58">Fig. 23.</emphasis>
<taxonomicName id="7EFE2A9FFFC1FFF3FE8F0E40FDDDD970" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[293,520,1636,1662]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC1FFF3FE8F0E40FDDDD970" box="[293,520,1636,1662]" italics="true" pageId="57" pageNumber="58">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC1FFF3FDBA0E40FDA5D970" box="[528,624,1636,1662]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
, genitalia drawings; A–C
<emphasis id="8B8A8D0EFFC1FFF3FC060E40FC68D970" bold="true" box="[940,957,1636,1662]" pageId="57" pageNumber="58">=</emphasis>
male holotype (CASENT_9082245; FMNH); D–E= female paratype (CASENT_9002829; CAS).
<emphasis id="8B8A8D0EFFC1FFF3FCDC0EA3FC5ED9AF" bold="true" box="[886,907,1671,1697]" pageId="57" pageNumber="58">A</emphasis>
. Left palp, prolateral view.
<emphasis id="8B8A8D0EFFC1FFF3FB140EA3FB04D9AF" bold="true" box="[1214,1233,1671,1697]" pageId="57" pageNumber="58">B</emphasis>
. Idem, ventral view.
<emphasis id="8B8A8D0EFFC1FFF3FEAF0E8EFECFD9CA" bold="true" box="[261,282,1706,1732]" pageId="57" pageNumber="58">C</emphasis>
. Idem, retrolateral view.
<emphasis id="8B8A8D0EFFC1FFF3FDE10E8EFD8AD9CA" bold="true" box="[587,607,1706,1732]" pageId="57" pageNumber="58">D</emphasis>
. Epigyne, ventral view.
<emphasis id="8B8A8D0EFFC1FFF3FC2C0E8EFC4DD9CA" bold="true" box="[902,920,1706,1732]" pageId="57" pageNumber="58">E</emphasis>
. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold.
</paragraph>
</caption>
<paragraph id="B941511CFFC2FFF0FF170A24FEE6DD14" blockId="58.[189,307,512,538]" box="[189,307,512,538]" pageId="58" pageNumber="59">
<heading id="E209E670FFC2FFF0FF170A24FEE6DD14" bold="true" box="[189,307,512,538]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="8B8A8D0EFFC2FFF0FF170A24FEE6DD14" bold="true" box="[189,307,512,538]" pageId="58" pageNumber="59">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF400A14FEFFDD44" blockId="58.[189,920,560,622]" box="[234,298,560,586]" pageId="58" pageNumber="59">
<emphasis id="8B8A8D0EFFC2FFF0FF400A14FEFFDD44" bold="true" box="[234,298,560,586]" pageId="58" pageNumber="59">Male</emphasis>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170A70FC4DDD60" blockId="58.[189,920,560,622]" box="[189,920,596,622]" pageId="58" pageNumber="59">
Variation unknown: only the single
<typeStatus id="6645EFBEFFC2FFF0FDCB0A70FD1DDD60" box="[609,712,596,622]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
has been studied.
</paragraph>
<paragraph id="B941511CFFC2FFF0FF400AB3FE48DDBF" blockId="58.[187,1400,663,979]" box="[234,413,663,689]" pageId="58" pageNumber="59">
<emphasis id="8B8A8D0EFFC2FFF0FF400AB3FE90DDBF" bold="true" box="[234,325,663,689]" pageId="58" pageNumber="59">Female</emphasis>
(n = 3)
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170A9FFCFDDCDC" blockId="58.[187,1400,663,979]" pageId="58" pageNumber="59">[Female variation from CASENT_9082245, CASENT_9031270 and CASENT_9031271]. Total length = 20.00–30.00, carapace length/width = 1.24–1.30, carapace height/width = 0.39–0.48, PER/carapace width = 0.21–0.24, PER/OAL = 2.90–3.13, PER/AER = 1.35–1.36, OQP/OQA = 1.18–1.23, sternum length/width = 1.20–1.31, palpal coxa length/width = 2.24–2.44, femur I length/carapace width = 0.87– 1.00, metatarsus I length/carapace width = 0.64–0.74, femur IV length/carapace width = 0.96–1.06, palpal tarsus length/carapace width = 0.34–0.41, palpal tarsus length/palpal patella length = 1.46–2.00, palpal tarsus length/palpal tibia length = 1.09–1.57, palpal tarsus length/palpal femur length = 0.76–0.95, palpal tibia length/palpal patella length = 1.08–1.40.</paragraph>
<paragraph id="B941511CFFC2FFF0FF170BDFFE40DB1B" blockId="58.[189,405,1019,1045]" box="[189,405,1019,1045]" pageId="58" pageNumber="59">
<heading id="E209E670FFC2FFF0FF170BDFFE40DB1B" bold="true" box="[189,405,1019,1045]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="8B8A8D0EFFC2FFF0FF170BDFFE40DB1B" bold="true" box="[189,405,1019,1045]" pageId="58" pageNumber="59">Note on synonym</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170C0EFBE8DB67" blockId="58.[189,1399,1066,1129]" pageId="58" pageNumber="59">
A specimen of this species (CASENT_9031271) was used as an exemplar in a previously published molecular phylogeny, listed as “
<taxonomicName id="7EFE2A9FFFC2FFF0FD9C0C6AFCD4DB67" box="[566,769,1102,1129]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="8B8A8D0EFFC2FFF0FD9C0C6AFD11DB66" box="[566,708,1102,1128]" italics="true" pageId="58" pageNumber="59">Zorodictyna</emphasis>
sp. 1
</taxonomicName>
” by
<bibRefCitation id="DD6F2CEDFFC2FFF0FC960C6AFBECDB67" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[828,1081,1102,1129]" pageId="58" pageNumber="59" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow
<emphasis id="8B8A8D0EFFC2FFF0FC0F0C6BFC35DB66" box="[933,992,1102,1128]" italics="true" pageId="58" pageNumber="59">et al.</emphasis>
(2015)
</bibRefCitation>
.
</paragraph>
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<emphasis id="8B8A8D0EFFC2FFF0FF170CB5FEABDBA5" bold="true" box="[189,382,1169,1195]" pageId="58" pageNumber="59">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170CE4FC8DDBF1" blockId="58.[187,1398,1216,1279]" pageId="58" pageNumber="59">
The species
<taxonomicName id="7EFE2A9FFFC2FFF0FEE30CE4FDF0DBD4" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[329,549,1216,1242]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC2FFF0FEE30CE4FDF0DBD4" box="[329,549,1216,1242]" italics="true" pageId="58" pageNumber="59">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC2FFF0FD860CE5FD50DBD5" box="[556,645,1217,1243]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
is recorded from tropical dry forest.Although they have an entire cribellum we know nothing of their use of cribellate silk.
</paragraph>
</subSubSection>
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<heading id="E209E670FFC2FFF0FF170D03FE83DA4F" bold="true" box="[189,342,1319,1345]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="8B8A8D0EFFC2FFF0FF170D03FE83DA4F" bold="true" box="[189,342,1319,1345]" pageId="58" pageNumber="59">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFC2FFF0FF170D73FDB7DA9B" blockId="58.[189,1399,1367,1429]" pageId="58" pageNumber="59">
<taxonomicName id="7EFE2A9FFFC2FFF0FF170D73FE75DA7F" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[189,416,1367,1393]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFC2FFF0FF170D73FE75DA7F" box="[189,416,1367,1393]" italics="true" pageId="58" pageNumber="59">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFC2FFF0FE060D73FDDFDA7F" box="[428,522,1367,1393]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
is known from National Park Ankarafantsika in Mahajunga Province in northwestern
<collectingCountry id="C1E9118CFFC2FFF0FEF50D5FFE38DA9B" box="[351,493,1403,1429]" name="Madagascar" pageId="58" pageNumber="59">Madagascar</collectingCountry>
(
<figureCitation id="21C54D99FFC2FFF0FE570D5FFD80DA9B" box="[509,597,1403,1429]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="58" pageNumber="59">Fig. 26</figureCitation>
).
</paragraph>
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<emphasis id="8B8A8D0EFFD4FFE6FD87092DFC9ADE2D" bold="true" box="[557,847,265,291]" italics="true" pageId="44" pageNumber="45">Zorascar pasunepomme</emphasis>
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<figureCitation id="21C54D99FFD4FFE6FD09096BFCDADE67" box="[675,783,335,361]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; A–B = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); C–D = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–B = 2 mm; C–E = 0.5 mm. Illustrations A–B, E by Alain Reygel; C–D by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="44" pageNumber="45">Figs 16E</figureCitation>
,
<figureCitation id="21C54D99FFD4FFE6FCB1096BFCB2DE67" box="[795,871,335,361]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="45.[189,231,1928,1954]" captionStartId-1="46.[189,233,1894,1920]" captionStartId-2="47.[189,232,1570,1596]" captionStartId-3="48.[189,233,1705,1731]" captionStartId-4="49.[192,236,1327,1353]" captionTargetBox-0="[217,1398,556,1882]" captionTargetBox-1="[189,1398,1053,1856]" captionTargetBox-2="[189,1398,265,1533]" captionTargetBox-3="[189,1398,265,1665]" captionTargetBox-4="[189,1398,265,1289]" captionTargetId-0="figure-69@45.[189,1398,556,1891]" captionTargetId-1="figure-159@46.[189,1398,1053,1859]" captionTargetId-2="figure-11@47.[189,1398,265,1533]" captionTargetId-3="figure-19@48.[189,1398,265,1665]" captionTargetId-4="figure-118@49.[189,1398,265,1289]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionTargetPageId-2="47" captionTargetPageId-3="48" captionTargetPageId-4="49" captionText-0="Fig. 17. Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA), habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." captionText-1="Fig. 18. Zorascar pasunepomme gen. et sp. nov. holotype female (BE_RMCA_ARA.Ara.247346; RMCA), genitalia. A. Epigyne, ventral view. B. Endogyne, cleared, dorsal view. C. Idem, dorso-anterior view. D. Idem, dorso-posterior view. Scale bars = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." captionText-2="Fig. 19. Zorascar pasunepomme gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247346; RMCA), epigyne, CT-scan images. A. ventral view. B. ventro-anterior view. C. dorsal view. D. dorso-anterior view. E. idem, detail.F. Idem, dorso-lateral view. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; ML = median lobe; MS = median sector; S = spermatheca. Scale bars = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." captionText-3="Fig. 20. Zorascar pasunepomme gen. et sp. nov., paratype juvenile (BE_RMCA_ARA.Ara.247347; RMCA), SEM pictures. A. ALS, caudal view. B. PLS, posterior view. C. PMS, posterior view. D. Right serrula, ventral view. E. Tarsus of leg 1, arrow pointing to the tarsal organ. F. Tarsus of leg 4. Scale bars: A–D = 10 μm; E–F = 100 μm. SEM images by Rudy Jocqué." captionText-4="Fig. 21. Zorascar pasunepomme gen. et sp. nov., paratype juvenile (BE_RMCA_ARA.Ara.247347; RMCA). A. Tarsal claws of leg 1. B. Tarsal claws of leg 4. C. Tarsal organ of leg 2. D. Tarsal organ and trichobothrium of leg 4. E. Trichobothrium, tarsus 1. Scale bars: A = 50 μm; B = 20 μm; C, E = 5 μm; D = 10 μm. SEM images by Rudy Jocqué." figureDoi-0="http://doi.org/10.5281/zenodo.14001833" figureDoi-1="http://doi.org/10.5281/zenodo.14001835" figureDoi-2="http://doi.org/10.5281/zenodo.14001837" figureDoi-3="http://doi.org/10.5281/zenodo.14001839" figureDoi-4="http://doi.org/10.5281/zenodo.14001841" httpUri-0="https://zenodo.org/record/14001833/files/figure.png" httpUri-1="https://zenodo.org/record/14001835/files/figure.png" httpUri-2="https://zenodo.org/record/14001837/files/figure.png" httpUri-3="https://zenodo.org/record/14001839/files/figure.png" httpUri-4="https://zenodo.org/record/14001841/files/figure.png" pageId="44" pageNumber="45">17–21</figureCitation>
,
<figureCitation id="21C54D99FFD4FFE6FCD9096BFC45DE67" box="[883,912,335,361]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="44" pageNumber="45">26</figureCitation>
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<subSubSection id="F1E40297FFD4FFE6FF1709B4FD8FDD11" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="B941511CFFD4FFE6FF1709B4FEE3DEA4" blockId="44.[189,310,400,426]" box="[189,310,400,426]" pageId="44" pageNumber="45">
<heading id="E209E670FFD4FFE6FF1709B4FEE3DEA4" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="8B8A8D0EFFD4FFE6FF1709B4FEE3DEA4" bold="true" box="[189,310,400,426]" pageId="44" pageNumber="45">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFD4FFE6FF17099BFD8FDD11" blockId="44.[188,1401,447,543]" pageId="44" pageNumber="45">
The female of
<taxonomicName id="7EFE2A9FFFD4FFE6FEC509E4FD59DED7" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[367,652,448,473]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="pasunepomme" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFD4FFE6FEC509E4FD59DED7" box="[367,652,448,473]" italics="true" pageId="44" pageNumber="45">Zorascar pasunepomme</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFD4FFE6FD3F099BFC9EDED7" box="[661,843,447,473]" pageId="44" pageNumber="45" rank="species">gen. et sp. nov.</taxonomicNameLabel>
is characterized by the epigyne with a median lobe (ML) that is narrow and deeply concave medially and copulatory openings (CO) exposed laterad of median lobe (
<figureCitation id="21C54D99FFD4FFE6FEF60A21FE1CDD11" box="[348,457,517,543]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; A–B = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); C–D = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–B = 2 mm; C–E = 0.5 mm. Illustrations A–B, E by Alain Reygel; C–D by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="44" pageNumber="45">Figs 16E</figureCitation>
,
<figureCitation id="21C54D99FFD4FFE6FE7F0A21FDDFDD11" box="[469,522,517,543]" captionStart="Fig" captionStartId="46.[189,233,1894,1920]" captionTargetBox="[189,1398,1053,1856]" captionTargetId="figure-159@46.[189,1398,1053,1859]" captionTargetPageId="46" captionText="Fig. 18. Zorascar pasunepomme gen. et sp. nov. holotype female (BE_RMCA_ARA.Ara.247346; RMCA), genitalia. A. Epigyne, ventral view. B. Endogyne, cleared, dorsal view. C. Idem, dorso-anterior view. D. Idem, dorso-posterior view. Scale bars = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001835" httpUri="https://zenodo.org/record/14001835/files/figure.png" pageId="44" pageNumber="45">18A</figureCitation>
,
<figureCitation id="21C54D99FFD4FFE6FDBC0A21FD98DD11" box="[534,589,517,543]" captionStart="Fig" captionStartId="47.[189,232,1570,1596]" captionTargetBox="[189,1398,265,1533]" captionTargetId="figure-11@47.[189,1398,265,1533]" captionTargetPageId="47" captionText="Fig. 19. Zorascar pasunepomme gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247346; RMCA), epigyne, CT-scan images. A. ventral view. B. ventro-anterior view. C. dorsal view. D. dorso-anterior view. E. idem, detail.F. Idem, dorso-lateral view. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; ML = median lobe; MS = median sector; S = spermatheca. Scale bars = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001837" httpUri="https://zenodo.org/record/14001837/files/figure.png" pageId="44" pageNumber="45">19A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFD4FFE6FF170A63FB40DDBC" pageId="44" pageNumber="45" type="etymology">
<paragraph id="B941511CFFD4FFE6FF170A63FE91DD6F" blockId="44.[189,324,583,609]" box="[189,324,583,609]" pageId="44" pageNumber="45">
<heading id="E209E670FFD4FFE6FF170A63FE91DD6F" bold="true" box="[189,324,583,609]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="8B8A8D0EFFD4FFE6FF170A63FE91DD6F" bold="true" box="[189,324,583,609]" pageId="44" pageNumber="45">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFD4FFE6FF170A51FB40DDBC" blockId="44.[187,1399,629,691]" pageId="44" pageNumber="45">The species name, a noun in apposition, refers to the famous painting of René Magritte depicting an apple, named ‘Ceci n’est pas une pomme’, to which the genitalia appear to be reminiscent.</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFD4FFE6FF170AFEFBF1DB70" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="B941511CFFD4FFE6FF170AFEFEB9DDFA" blockId="44.[189,364,730,756]" box="[189,364,730,756]" pageId="44" pageNumber="45">
<heading id="E209E670FFD4FFE6FF170AFEFEB9DDFA" bold="true" box="[189,364,730,756]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="8B8A8D0EFFD4FFE6FF170AFEFEB9DDFA" bold="true" box="[189,364,730,756]" pageId="44" pageNumber="45">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFD4FFE6FF400B2DFE89DC2D" blockId="44.[189,1406,777,908]" box="[234,348,777,803]" pageId="44" pageNumber="45">
<emphasis id="8B8A8D0EFFD4FFE6FF400B2DFE89DC2D" bold="true" box="[234,348,777,803]" pageId="44" pageNumber="45">
<typeStatus id="6645EFBEFFD4FFE6FF400B2DFE89DC2D" box="[234,348,777,803]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFD4FFE6FF170B08FD40DC82" blockId="44.[189,1406,777,908]" pageId="44" pageNumber="45">
<materialsCitation id="09965B41FFD4FFE6FF170B08FD40DC82" collectingDate="2019-05-02" collectionCode="RMCA" collectorName="L. Kemp &amp; Ara." country="Madagascar" elevation="1581" latitude="-14.31592" location="Bamevika" longLatPrecision="1" longitude="48.723938" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana" typeStatus="holotype">
<collectingCountry id="C1E9118CFFD4FFE6FF170B08FE5BDC48" box="[189,398,812,838]" name="Madagascar" pageId="44" pageNumber="45">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFD4FFE6FE030B0FFD6EDC4B" bold="true" box="[425,699,811,837]" pageId="44" pageNumber="45">
<collectingRegion id="7B3A9FFEFFD4FFE6FE030B0FFD6EDC4B" box="[425,699,811,837]" country="Madagascar" name="Antsiranana" pageId="44" pageNumber="45">Antsiranana Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFD4FFE6FD790B08FD3FDC48" box="[723,746,812,838]" count="1" pageId="44" pageNumber="45" type="female">♀</specimenCount>
;
<location id="BC2107C7FFD4FFE6FD520B08FCA6DC48" LSID="urn:lsid:plazi:treatment:3157E00AFFD4FFE4FD87092DFA8BDCCE:BC2107C7FFD4FFE6FD520B08FCA6DC48" box="[760,883,812,838]" country="Madagascar" latitude="-14.31592" longLatPrecision="1" longitude="48.723938" name="Bamevika" pageId="44" pageNumber="45" stateProvince="Antsiranana">Bamevika</location>
;
<geoCoordinate id="DCCA37DBFFD4FFE6FCD40B0EFB95DC48" box="[894,1088,810,838]" degrees="14" direction="south" minutes="18" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="57.312" value="-14.31592">14°18′57.312″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFD4FFE6FBE70B0EFAC7DC48" box="[1101,1298,810,838]" degrees="48" direction="east" minutes="43" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="26.180" value="48.723938">48°43′26.180″ E</geoCoordinate>
;
<elevation id="32D3B62FFFD4FFE6FAB40B0FFF22DC67" metricMagnitude="3" metricUnit="m" metricValue="1.581" pageId="44" pageNumber="45" unit="m" value="1581.0">
<quantity id="7E06FCF9FFD4FFE6FAB40B0FFAA2DC48" box="[1310,1399,811,838]" metricMagnitude="3" metricUnit="m" metricValue="1.581" pageId="44" pageNumber="45" unit="m" value="1581.0">1581 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFD4FFE6FEAC0B6BFE48DC67" box="[262,413,847,873]" pageId="44" pageNumber="45" value="2019-05-02">
<collectingDate id="DD048E34FFD4FFE6FEAC0B6BFE48DC67" box="[262,413,847,873]" pageId="44" pageNumber="45" value="2019-05-02">2 May 2019</collectingDate>
</date>
;
<collectorName id="140B34CAFFD4FFE6FE010B6BFDC2DC67" box="[427,535,847,873]" pageId="44" pageNumber="45">L. Kemp</collectorName>
leg.; hand collected, grassland; DNA RJ4; BINCO_MAD_19_0079_2; BE_
<collectionCode id="DFEFC9D9FFD4FFE6FF580B56FE99DC82" box="[242,332,882,908]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="44" pageNumber="45" type="Museum">RMCA</collectionCode>
_ARA.
<collectorName id="140B34CAFFD4FFE6FE350B56FE05DC82" box="[415,464,882,908]" pageId="44" pageNumber="45">Ara.</collectorName>
247346;
<collectionCode id="DFEFC9D9FFD4FFE6FD9C0B56FD45DC82" box="[566,656,882,908]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="44" pageNumber="45" type="Museum">RMCA</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B941511CFFD4FFE6FF400B90FEB2DCC0" blockId="44.[189,1399,948,1150]" box="[234,359,948,974]" pageId="44" pageNumber="45">
<emphasis id="8B8A8D0EFFD4FFE6FF400B90FEB2DCC0" bold="true" box="[234,359,948,974]" pageId="44" pageNumber="45">
<typeStatus id="6645EFBEFFD4FFE6FF400B90FEB2DCC0" box="[234,359,948,974]" pageId="44" pageNumber="45" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFD4FFE6FF170BF3FBF1DB70" blockId="44.[189,1399,948,1150]" pageId="44" pageNumber="45">
<materialsCitation id="09965B41FFD4FFE6FF170BF3FC09DB1A" collectingDate="2019-05-02" collectionCode="RMCA" collectorName="Ara." country="Madagascar" elevation="1581" latitude="-14.31592" location="Bamevika" longLatPrecision="1" longitude="48.723938" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<collectingCountry id="C1E9118CFFD4FFE6FF170BF3FE46DCFF" box="[189,403,983,1009]" name="Madagascar" pageId="44" pageNumber="45">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFD4FFE6FE100BF2FD0FDCFE" bold="true" box="[442,730,982,1008]" pageId="44" pageNumber="45">
<collectingRegion id="7B3A9FFEFFD4FFE6FE100BF2FD0FDCFE" box="[442,730,982,1008]" country="Madagascar" name="Antsiranana" pageId="44" pageNumber="45">Antsiranana Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFD4FFE6FD570BF3FC9CDCFF" box="[765,841,983,1009]" count="1" pageId="44" pageNumber="45" type="generic">1 juv.</specimenCount>
; same data as for holotype; [SEM imaged]; BINCO_MAD_19_0079_2; BE_
<collectionCode id="DFEFC9D9FFD4FFE6FDE90BDEFD48DB1A" box="[579,669,1018,1044]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="44" pageNumber="45" type="Museum">RMCA</collectionCode>
_ARA.
<collectorName id="140B34CAFFD4FFE6FD5A0BDEFCF4DB1A" box="[752,801,1018,1044]" pageId="44" pageNumber="45">Ara.</collectorName>
247347;
<collectionCode id="DFEFC9D9FFD4FFE6FC2D0BDEFC09DB1A" box="[903,988,1018,1044]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="44" pageNumber="45" type="Museum">RMCA</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFD4FFE6FC5E0BDEFCA2DB54" collectionCode="RMCA" collectorName="M. Jocque &amp; Ara." country="Madagascar" elevation="1557" latitude="-14.316361" longLatPrecision="1" longitude="48.72303" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<specimenCount id="AFF89A95FFD4FFE6FC5E0BDEFBECDB1A" box="[1012,1081,1018,1044]" count="1" pageId="44" pageNumber="45" type="generic">1 juv.</specimenCount>
; Bamevika;
<geoCoordinate id="DCCA37DBFFD4FFE6FB610BDCFAA4DB1A" box="[1227,1393,1016,1044]" degrees="14" direction="south" minutes="18" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="58.9" value="-14.316361">14°18′58.9″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFD4FFE6FF170C3FFEBEDB36" box="[189,363,1051,1080]" degrees="48" direction="east" minutes="43" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="22.9" value="48.72303">48°43′22.9″ E</geoCoordinate>
;
<elevation id="32D3B62FFFD4FFE6FED00C39FDCFDB39" box="[378,538,1053,1080]" metricMagnitude="3" metricUnit="m" metricValue="1.557" pageId="44" pageNumber="45" unit="m" value="1557.0">
<quantity id="7E06FCF9FFD4FFE6FED00C39FE02DB36" box="[378,471,1053,1080]" metricMagnitude="3" metricUnit="m" metricValue="1.557" pageId="44" pageNumber="45" unit="m" value="1557.0">1557 m</quantity>
a.s.l.
</elevation>
;
<collectorName id="140B34CAFFD4FFE6FD800C3AFD7CDB36" box="[554,681,1054,1080]" pageId="44" pageNumber="45">M. Jocque</collectorName>
leg.; hand collected, forest leaf litter; DNA RJ2; BINCO_ MAD_19_0079_2; BE_
<collectionCode id="DFEFC9D9FFD4FFE6FE710C64FDE0DB54" box="[475,565,1088,1114]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="44" pageNumber="45" type="Museum">RMCA</collectionCode>
_ARA.
<collectorName id="140B34CAFFD4FFE6FD230C64FD6EDB54" box="[649,699,1088,1114]" pageId="44" pageNumber="45">Ara.</collectorName>
247349;
<collectionCode id="DFEFC9D9FFD4FFE6FC880C64FCA2DB54" box="[802,887,1088,1114]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="44" pageNumber="45" type="Museum">RMCA</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFD4FFE6FC390C64FBCADB70" collectionCode="RMCA" collectorName="Ara." country="Madagascar" elevation="1557" latitude="-14.316361" longLatPrecision="1" longitude="48.72303" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<specimenCount id="AFF89A95FFD4FFE6FC390C64FC0FDB54" box="[915,986,1088,1114]" count="1" pageId="44" pageNumber="45" type="generic">1 juv.</specimenCount>
; same data as for preceding; DNA RJ3; BINCO_MAD_19_0079_2; BE_
<collectionCode id="DFEFC9D9FFD4FFE6FD2B0C40FD0EDB70" box="[641,731,1124,1150]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="44" pageNumber="45" type="Museum">RMCA</collectionCode>
_ARA.
<collectorName id="140B34CAFFD4FFE6FC840C40FCB5DB70" box="[814,864,1124,1150]" pageId="44" pageNumber="45">Ara.</collectorName>
247348;
<collectionCode id="DFEFC9D9FFD4FFE6FC6F0C40FBCADB70" box="[965,1055,1124,1150]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="44" pageNumber="45" type="Museum">RMCA</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFD4FFE4FF170C81FEECDC5E" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45" type="description">
<paragraph id="B941511CFFD4FFE6FF170C81FE9BDBB1" blockId="44.[189,334,1189,1215]" box="[189,334,1189,1215]" pageId="44" pageNumber="45">
<heading id="E209E670FFD4FFE6FF170C81FE9BDBB1" bold="true" box="[189,334,1189,1215]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="8B8A8D0EFFD4FFE6FF170C81FE9BDBB1" bold="true" box="[189,334,1189,1215]" pageId="44" pageNumber="45">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFD4FFE6FF400CF0FC76DBE0" blockId="44.[189,1160,1235,1297]" box="[234,931,1235,1262]" pageId="44" pageNumber="45">
<emphasis id="8B8A8D0EFFD4FFE6FF400CF0FE6CDBE0" bold="true" box="[234,441,1236,1262]" pageId="44" pageNumber="45">
Female
<typeStatus id="6645EFBEFFD4FFE6FEE70CF0FE6CDBE0" box="[333,441,1236,1262]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
</emphasis>
(BE_RMCA_ARA.Ara.247346; RMCA)
</paragraph>
<paragraph id="B941511CFFD4FFE6FF170CD3FB5DDA1F" blockId="44.[189,1160,1235,1297]" box="[189,1160,1270,1297]" pageId="44" pageNumber="45">MEASUREMENTS. Total length: 6.82. Carapace: length 2.91, width 2.30, height 1.42.</paragraph>
<paragraph id="B941511CFFD4FFE6FF170D1CFD91DAD1" blockId="44.[187,1401,1336,1503]" pageId="44" pageNumber="45">
COLOUR (
<figureCitation id="21C54D99FFD4FFE6FE9E0D1CFE44DA5C" box="[308,401,1336,1362]" captionStart="Fig" captionStartId="45.[189,231,1928,1954]" captionTargetBox="[217,1398,556,1882]" captionTargetId="figure-69@45.[189,1398,556,1891]" captionTargetPageId="45" captionText="Fig. 17. Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA), habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001833" httpUri="https://zenodo.org/record/14001833/files/figure.png" pageId="44" pageNumber="45">Figs 17</figureCitation>
). Carapace brownish yellow, with two contrasting dark bands of short setae along margin and faint median band between PME and fovea; chelicerae brownish, anteriorly with two thin longitudinal darker stripes; sternum yellowish brown; legs brownish yellow with three darker rings on femora, two on tibiae, distally and proximally; abdomen brownish yellow with spotted pattern on dorsum and sides; spinnerets pale yellow.
</paragraph>
<paragraph id="B941511CFFD4FFE6FF170E22FBEED92E" blockId="44.[189,1083,1542,1569]" box="[189,1083,1542,1569]" pageId="44" pageNumber="45">
STERNUM (
<figureCitation id="21C54D99FFD4FFE6FE970E22FE72D92E" box="[317,423,1542,1569]" captionStart="Fig" captionStartId="45.[189,231,1928,1954]" captionTargetBox="[217,1398,556,1882]" captionTargetId="figure-69@45.[189,1398,556,1891]" captionTargetPageId="45" captionText="Fig. 17. Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA), habitus. A. Dorsal view. B. Ventral view. C. Lateral view. Scale bars = 1 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001833" httpUri="https://zenodo.org/record/14001833/files/figure.png" pageId="44" pageNumber="45">Fig. 17B</figureCitation>
). Oval with slightly sinuous sides; 1.21 wide, 1.42 long.
</paragraph>
<paragraph id="B941511CFFD4FFE6FF170E6CFDF7D96C" blockId="44.[189,546,1608,1634]" box="[189,546,1608,1634]" pageId="44" pageNumber="45">CHILUM. 0.12 high, 0.20 wide.</paragraph>
<paragraph id="B941511CFFD4FFE6FF170EADFCCAD9C8" blockId="44.[188,1399,1673,1735]" pageId="44" pageNumber="45">EYES. ALE 0.16; AME 0.12; PLE 0.16; PME 0.16; ALE–AME 0.05; AME–AME 0.07; PLE–PME 0.16; PME–PME 0.10; MOQ: AW 0.28, PW 0.39, L 0.36.</paragraph>
<paragraph id="B941511CFFD4FFE6FF170ECAFDF4D806" blockId="44.[189,545,1774,1800]" box="[189,545,1774,1800]" pageId="44" pageNumber="45">CLYPEUS. With five long setae.</paragraph>
<paragraph id="B941511CFFD5FFE7FF17092DFE4FDE2D" blockId="45.[189,410,265,291]" box="[189,410,265,291]" pageId="45" pageNumber="46">Leg measurements</paragraph>
<paragraph id="B941511CFFD5FFE7FEA00917FD31DEE2" pageId="45" pageNumber="46">
<table id="CBFEA3BCFFD50035FF610916FD3BDEE3" box="[203,750,306,493]" gridcols="7" gridrows="5" pageId="45" pageNumber="46" rowsContinueIn="46.[203,912,308,505]">
<tr id="07CE535EFFD50035FF610916FD3BDE43" box="[203,750,306,333]" gridrow="0" pageId="45" pageNumber="46" rowspan-0="1">
<th id="441F3A22FFD50035FEA00916FEE8DE43" box="[266,317,306,333]" gridcol="1" gridrow="0" pageId="45" pageNumber="46">F</th>
<th id="441F3A22FFD50035FEF50916FE47DE43" box="[351,402,306,333]" gridcol="2" gridrow="0" pageId="45" pageNumber="46">P</th>
<th id="441F3A22FFD50035FE1E0916FE32DE43" box="[436,487,306,333]" gridcol="3" gridrow="0" pageId="45" pageNumber="46">T</th>
<th id="441F3A22FFD50035FDA20916FDEEDE43" box="[520,571,306,333]" gridcol="4" gridrow="0" pageId="45" pageNumber="46">Mt</th>
<th id="441F3A22FFD50035FDF70916FD45DE43" box="[605,656,306,333]" gridcol="5" gridrow="0" pageId="45" pageNumber="46">t</th>
<th id="441F3A22FFD50035FD180916FD3BDE43" box="[690,750,306,333]" gridcol="6" gridrow="0" pageId="45" pageNumber="46">Total</th>
</tr>
<tr id="07CE535EFFD50035FF61097FFD3BDE78" box="[203,750,347,374]" gridrow="1" pageId="45" pageNumber="46">
<th id="441F3A22FFD50035FF61097FFF3FDE78" box="[203,234,347,374]" gridcol="0" gridrow="1" pageId="45" pageNumber="46">I</th>
<td id="441F3A22FFD50035FEA0097FFEE8DE78" box="[266,317,347,374]" gridcol="1" gridrow="1" pageId="45" pageNumber="46">2.10</td>
<td id="441F3A22FFD50035FEF5097FFE47DE78" box="[351,402,347,374]" gridcol="2" gridrow="1" pageId="45" pageNumber="46">1.05</td>
<td id="441F3A22FFD50035FE1E097FFE32DE78" box="[436,487,347,374]" gridcol="3" gridrow="1" pageId="45" pageNumber="46">1.82</td>
<td id="441F3A22FFD50035FDA2097FFDEEDE78" box="[520,571,347,374]" gridcol="4" gridrow="1" pageId="45" pageNumber="46">1.61</td>
<td id="441F3A22FFD50035FDF7097FFD45DE78" box="[605,656,347,374]" gridcol="5" gridrow="1" pageId="45" pageNumber="46">1.00</td>
<td id="441F3A22FFD50035FD18097FFD3BDE78" box="[690,750,347,374]" gridcol="6" gridrow="1" pageId="45" pageNumber="46">7.58</td>
</tr>
<tr id="07CE535EFFD50035FF6109A6FD3BDE93" box="[203,750,386,413]" gridrow="2" pageId="45" pageNumber="46">
<th id="441F3A22FFD50035FF6109A6FF3FDE93" box="[203,234,386,413]" gridcol="0" gridrow="2" pageId="45" pageNumber="46">II</th>
<td id="441F3A22FFD50035FEA009A6FEE8DE93" box="[266,317,386,413]" gridcol="1" gridrow="2" pageId="45" pageNumber="46">1.68</td>
<td id="441F3A22FFD50035FEF509A6FE47DE93" box="[351,402,386,413]" gridcol="2" gridrow="2" pageId="45" pageNumber="46">0.91</td>
<td id="441F3A22FFD50035FE1E09A6FE32DE93" box="[436,487,386,413]" gridcol="3" gridrow="2" pageId="45" pageNumber="46">1.47</td>
<td id="441F3A22FFD50035FDA209A6FDEEDE93" box="[520,571,386,413]" gridcol="4" gridrow="2" pageId="45" pageNumber="46">1.40</td>
<td id="441F3A22FFD50035FDF709A6FD45DE93" box="[605,656,386,413]" gridcol="5" gridrow="2" pageId="45" pageNumber="46">0.80</td>
<td id="441F3A22FFD50035FD1809A6FD3BDE93" box="[690,750,386,413]" gridcol="6" gridrow="2" pageId="45" pageNumber="46">6.26</td>
</tr>
<tr id="07CE535EFFD50035FF61098EFD3BDECB" box="[203,750,426,453]" gridrow="3" pageId="45" pageNumber="46">
<th id="441F3A22FFD50035FF61098EFF3FDECB" box="[203,234,426,453]" gridcol="0" gridrow="3" pageId="45" pageNumber="46">III</th>
<td id="441F3A22FFD50035FEA0098EFEE8DECB" box="[266,317,426,453]" gridcol="1" gridrow="3" pageId="45" pageNumber="46">1.75</td>
<td id="441F3A22FFD50035FEF5098EFE47DECB" box="[351,402,426,453]" gridcol="2" gridrow="3" pageId="45" pageNumber="46">0.84</td>
<td id="441F3A22FFD50035FE1E098EFE32DECB" box="[436,487,426,453]" gridcol="3" gridrow="3" pageId="45" pageNumber="46">1.33</td>
<td id="441F3A22FFD50035FDA2098EFDEEDECB" box="[520,571,426,453]" gridcol="4" gridrow="3" pageId="45" pageNumber="46">1.61</td>
<td id="441F3A22FFD50035FDF7098EFD45DECB" box="[605,656,426,453]" gridcol="5" gridrow="3" pageId="45" pageNumber="46">0.70</td>
<td id="441F3A22FFD50035FD18098EFD3BDECB" box="[690,750,426,453]" gridcol="6" gridrow="3" pageId="45" pageNumber="46">6.23</td>
</tr>
<tr id="07CE535EFFD50035FF6109F6FD3BDEE3" box="[203,750,466,493]" gridrow="4" pageId="45" pageNumber="46">
<th id="441F3A22FFD50035FF6109F6FF3FDEE3" box="[203,234,466,493]" gridcol="0" gridrow="4" pageId="45" pageNumber="46">IV</th>
<td id="441F3A22FFD50035FEA009F6FEE8DEE3" box="[266,317,466,493]" gridcol="1" gridrow="4" pageId="45" pageNumber="46">2.31</td>
<td id="441F3A22FFD50035FEF509F6FE47DEE3" box="[351,402,466,493]" gridcol="2" gridrow="4" pageId="45" pageNumber="46">0.91</td>
<td id="441F3A22FFD50035FE1E09F6FE32DEE3" box="[436,487,466,493]" gridcol="3" gridrow="4" pageId="45" pageNumber="46">2.03</td>
<td id="441F3A22FFD50035FDA209F6FDEEDEE3" box="[520,571,466,493]" gridcol="4" gridrow="4" pageId="45" pageNumber="46">2.66</td>
<td id="441F3A22FFD50035FDF709F6FD45DEE3" box="[605,656,466,493]" gridcol="5" gridrow="4" pageId="45" pageNumber="46">1.20</td>
<td id="441F3A22FFD50035FD1809F6FD3BDEE3" box="[690,750,466,493]" gridcol="6" gridrow="4" pageId="45" pageNumber="46">9.11</td>
</tr>
</table>
</paragraph>
<caption id="ED810194FFD5FFE7FF170FACFDA2D8E6" ID-DOI="http://doi.org/10.5281/zenodo.14001833" ID-Zenodo-Dep="14001833" httpUri="https://zenodo.org/record/14001833/files/figure.png" pageId="45" pageNumber="46" startId="45.[189,231,1928,1954]" targetBox="[217,1398,556,1882]" targetPageId="45">
<paragraph id="B941511CFFD5FFE7FF170FACFDA2D8E6" blockId="45.[187,1398,1928,2024]" pageId="45" pageNumber="46">
<emphasis id="8B8A8D0EFFD5FFE7FF170FACFEC0D8AC" bold="true" box="[189,277,1928,1954]" pageId="45" pageNumber="46">Fig. 17.</emphasis>
<taxonomicName id="7EFE2A9FFFD5FFE7FEB30FADFDFFD8AC" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[281,554,1929,1954]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pasunepomme" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFD5FFE7FEB30FADFDFFD8AC" box="[281,554,1929,1954]" italics="true" pageId="45" pageNumber="46">Zorascar pasunepomme</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFD5FFE7FD850FACFD03D8AC" box="[559,726,1928,1954]" pageId="45" pageNumber="46" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype female (BE_RMCA_ARA.Ara.247346; RMCA), habitus.
<emphasis id="8B8A8D0EFFD5FFE7FEB50F8FFEE1D8CB" bold="true" box="[287,308,1963,1989]" pageId="45" pageNumber="46">A</emphasis>
. Dorsal view.
<emphasis id="8B8A8D0EFFD5FFE7FE730F8FFE38D8CB" bold="true" box="[473,493,1963,1989]" pageId="45" pageNumber="46">B</emphasis>
. Ventral view.
<emphasis id="8B8A8D0EFFD5FFE7FD330F8FFD7BD8CB" bold="true" box="[665,686,1963,1989]" pageId="45" pageNumber="46">C</emphasis>
. Lateral view. Scale bars = 1 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué.
</paragraph>
</caption>
<paragraph id="B941511CFFD6FFE4FF17092DFD49DDE5" blockId="46.[189,301,265,291]" lastBlockId="46.[187,1400,545,747]" pageId="46" pageNumber="47">
Spination GENITALIA. Epigyne (
<figureCitation id="21C54D99FFD6FFE4FE1A0A05FDCFDD35" box="[432,538,545,571]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; A–B = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); C–D = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–B = 2 mm; C–E = 0.5 mm. Illustrations A–B, E by Alain Reygel; C–D by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="46" pageNumber="47">Figs 16E</figureCitation>
,
<figureCitation id="21C54D99FFD6FFE4FD8E0A05FD8CDD35" box="[548,601,545,571]" captionStart="Fig" captionStartId="46.[189,233,1894,1920]" captionTargetBox="[189,1398,1053,1856]" captionTargetId="figure-159@46.[189,1398,1053,1859]" captionTargetPageId="46" captionText="Fig. 18. Zorascar pasunepomme gen. et sp. nov. holotype female (BE_RMCA_ARA.Ara.247346; RMCA), genitalia. A. Epigyne, ventral view. B. Endogyne, cleared, dorsal view. C. Idem, dorso-anterior view. D. Idem, dorso-posterior view. Scale bars = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001835" httpUri="https://zenodo.org/record/14001835/files/figure.png" pageId="46" pageNumber="47">18A</figureCitation>
,
<figureCitation id="21C54D99FFD6FFE4FDC90A05FD6DDD35" box="[611,696,545,571]" captionStart="Fig" captionStartId="47.[189,232,1570,1596]" captionTargetBox="[189,1398,265,1533]" captionTargetId="figure-11@47.[189,1398,265,1533]" captionTargetPageId="47" captionText="Fig. 19. Zorascar pasunepomme gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247346; RMCA), epigyne, CT-scan images. A. ventral view. B. ventro-anterior view. C. dorsal view. D. dorso-anterior view. E. idem, detail.F. Idem, dorso-lateral view. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; ML = median lobe; MS = median sector; S = spermatheca. Scale bars = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001837" httpUri="https://zenodo.org/record/14001837/files/figure.png" pageId="46" pageNumber="47">19A–B</figureCitation>
) with median lobe (ML) narrow and deeply concave medially (MS), copulatory openings (CO) exposed laterad of ML; lateral lobe (LL) with pair of stout basal, blunt, conical projections (LP); spermathecae dark, comma shaped as in transparency. Endogyne (
<figureCitation id="21C54D99FFD6FFE4FB750A43FAA4DD8C" box="[1247,1393,615,642]" captionStart="Fig" captionStartId="46.[189,233,1894,1920]" captionTargetBox="[189,1398,1053,1856]" captionTargetId="figure-159@46.[189,1398,1053,1859]" captionTargetPageId="46" captionText="Fig. 18. Zorascar pasunepomme gen. et sp. nov. holotype female (BE_RMCA_ARA.Ara.247346; RMCA), genitalia. A. Epigyne, ventral view. B. Endogyne, cleared, dorsal view. C. Idem, dorso-anterior view. D. Idem, dorso-posterior view. Scale bars = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001835" httpUri="https://zenodo.org/record/14001835/files/figure.png" pageId="46" pageNumber="47">Figs 18B–D</figureCitation>
,
<figureCitation id="21C54D99FFD6FFE4FF160AAFFEC4DDAB" box="[188,273,651,677]" captionStart="Fig" captionStartId="47.[189,232,1570,1596]" captionTargetBox="[189,1398,265,1533]" captionTargetId="figure-11@47.[189,1398,265,1533]" captionTargetPageId="47" captionText="Fig. 19. Zorascar pasunepomme gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247346; RMCA), epigyne, CT-scan images. A. ventral view. B. ventro-anterior view. C. dorsal view. D. dorso-anterior view. E. idem, detail.F. Idem, dorso-lateral view. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; ML = median lobe; MS = median sector; S = spermatheca. Scale bars = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001837" httpUri="https://zenodo.org/record/14001837/files/figure.png" pageId="46" pageNumber="47">19C–F</figureCitation>
): sperm ducts (CD) short, cylindrical, with three twists; spermathecae (S) reniform, with fairly narrow ventral base, and dorsal, roughly semicircular part, provided with few pores; fertilization ducts (FD) short, flattened and slightly curved.
</paragraph>
<paragraph id="B941511CFFD6FFE4FEA00910FCF9DEF6" pageId="46" pageNumber="47">
<table id="CBFEA3BCFFD60035FF610910FC45DEF7" box="[203,912,308,505]" gridcols="5" gridrows="5" pageId="46" pageNumber="47" rowsContinueFrom="45.[203,750,306,493]">
<tr id="07CE535EFFD60035FF610910FC45DE40" box="[203,912,308,334]" gridrow="0" pageId="46" pageNumber="47" rowspan-0="1">
<th id="441F3A22FFD60035FEA00910FEBBDE40" box="[266,366,308,334]" gridcol="1" gridrow="0" pageId="46" pageNumber="47">F</th>
<th id="441F3A22FFD60035FE390910FE0CDE40" box="[403,473,308,334]" gridcol="2" gridrow="0" pageId="46" pageNumber="47">P</th>
<th id="441F3A22FFD60035FE570910FD7ADE40" box="[509,687,308,334]" gridcol="3" gridrow="0" pageId="46" pageNumber="47">T</th>
<th id="441F3A22FFD60035FD750910FC45DE40" box="[735,912,308,334]" gridcol="4" gridrow="0" pageId="46" pageNumber="47">Mt</th>
</tr>
<tr id="07CE535EFFD60035FF61097BFC45DE77" box="[203,912,351,377]" gridrow="1" pageId="46" pageNumber="47">
<th id="441F3A22FFD60035FF61097BFF3FDE77" box="[203,234,351,377]" gridcol="0" gridrow="1" pageId="46" pageNumber="47">I</th>
<td id="441F3A22FFD60035FEA0097BFEBBDE77" box="[266,366,351,377]" gridcol="1" gridrow="1" pageId="46" pageNumber="47">pl1d1v1</td>
<td id="441F3A22FFD60035FE39097BFE0CDE77" box="[403,473,351,377]" gridcol="2" gridrow="1" pageId="46" pageNumber="47">–</td>
<td id="441F3A22FFD60035FE57097BFD7ADE77" box="[509,687,351,377]" gridcol="3" gridrow="1" pageId="46" pageNumber="47">v2-2-2-2</td>
<td id="441F3A22FFD60035FD75097BFC45DE77" box="[735,912,351,377]" gridcol="4" gridrow="1" pageId="46" pageNumber="47">v2-2-2</td>
</tr>
<tr id="07CE535EFFD60035FF6109ADFC45DEAA" box="[203,912,393,420]" gridrow="2" pageId="46" pageNumber="47">
<th id="441F3A22FFD60035FF6109ADFF3FDEAA" box="[203,234,393,420]" gridcol="0" gridrow="2" pageId="46" pageNumber="47">II</th>
<td id="441F3A22FFD60035FEA009ADFEBBDEAA" box="[266,366,393,420]" gridcol="1" gridrow="2" pageId="46" pageNumber="47">pl2d1</td>
<td id="441F3A22FFD60035FE3909ADFE0CDEAA" box="[403,473,393,420]" gridcol="2" gridrow="2" pageId="46" pageNumber="47">–</td>
<td id="441F3A22FFD60035FE5709ADFD7ADEAA" box="[509,687,393,420]" gridcol="3" gridrow="2" pageId="46" pageNumber="47">pl3v2-2-2-2</td>
<td id="441F3A22FFD60035FD7509ADFC45DEAA" box="[735,912,393,420]" gridcol="4" gridrow="2" pageId="46" pageNumber="47">v2-2-2</td>
</tr>
<tr id="07CE535EFFD60035FF610990FC45DEC0" box="[203,912,436,462]" gridrow="3" pageId="46" pageNumber="47">
<th id="441F3A22FFD60035FF610990FF3FDEC0" box="[203,234,436,462]" gridcol="0" gridrow="3" pageId="46" pageNumber="47">III</th>
<td id="441F3A22FFD60035FEA00990FEBBDEC0" box="[266,366,436,462]" gridcol="1" gridrow="3" pageId="46" pageNumber="47">pl3d3rl2</td>
<td id="441F3A22FFD60035FE390990FE0CDEC0" box="[403,473,436,462]" gridcol="2" gridrow="3" pageId="46" pageNumber="47">pl1rl1</td>
<td id="441F3A22FFD60035FE570990FD7ADEC0" box="[509,687,436,462]" gridcol="3" gridrow="3" pageId="46" pageNumber="47">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFD60035FD750990FC45DEC0" box="[735,912,436,462]" gridcol="4" gridrow="3" pageId="46" pageNumber="47">pl3d1rl3v2-2-2</td>
</tr>
<tr id="07CE535EFFD60035FF6109FAFC45DEF7" box="[203,912,478,505]" gridrow="4" pageId="46" pageNumber="47">
<th id="441F3A22FFD60035FF6109FAFF3FDEF7" box="[203,234,478,505]" gridcol="0" gridrow="4" pageId="46" pageNumber="47">IV</th>
<td id="441F3A22FFD60035FEA009FAFEBBDEF7" box="[266,366,478,505]" gridcol="1" gridrow="4" pageId="46" pageNumber="47">pl3d2rl1</td>
<td id="441F3A22FFD60035FE3909FAFE0CDEF7" box="[403,473,478,505]" gridcol="2" gridrow="4" pageId="46" pageNumber="47">pl1rl1</td>
<td id="441F3A22FFD60035FE5709FAFD7ADEF7" box="[509,687,478,505]" gridcol="3" gridrow="4" pageId="46" pageNumber="47">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFD60035FD7509FAFC45DEF7" box="[735,912,478,505]" gridcol="4" gridrow="4" pageId="46" pageNumber="47">disp14</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFD6FFE4FF400B37FEFFDC23" blockId="46.[189,313,787,848]" box="[234,298,787,813]" pageId="46" pageNumber="47">
<emphasis id="8B8A8D0EFFD6FFE4FF400B37FEFFDC23" bold="true" box="[234,298,787,813]" pageId="46" pageNumber="47">Male</emphasis>
</paragraph>
<paragraph id="B941511CFFD6FFE4FF170B12FEECDC5E" blockId="46.[189,313,787,848]" box="[189,313,822,848]" pageId="46" pageNumber="47">Unknown.</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFD6FFE4FF170B53FA8BDCCE" box="[189,1374,887,960]" pageId="46" pageNumber="47" type="distribution">
<paragraph id="B941511CFFD6FFE4FF170B53FE83DC9F" blockId="46.[189,342,887,913]" box="[189,342,887,913]" pageId="46" pageNumber="47">
<heading id="E209E670FFD6FFE4FF170B53FE83DC9F" bold="true" box="[189,342,887,913]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="8B8A8D0EFFD6FFE4FF170B53FE83DC9F" bold="true" box="[189,342,887,913]" pageId="46" pageNumber="47">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFD6FFE4FF170B82FA8BDCCE" blockId="46.[189,1374,933,960]" box="[189,1374,933,960]" pageId="46" pageNumber="47">
Known only from the
<typeStatus id="6645EFBEFFD6FFE4FE680B82FE20DCCE" box="[450,501,934,960]" pageId="46" pageNumber="47">type</typeStatus>
locality in northern
<collectingCountry id="C1E9118CFFD6FFE4FD4E0B82FCA7DCCE" box="[740,882,934,960]" name="Madagascar" pageId="46" pageNumber="47">Madagascar</collectingCountry>
at an altitude of around
<quantity id="7E06FCF9FFD6FFE4FB3A0B81FB3CDCCE" box="[1168,1257,933,960]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="46" pageNumber="47" unit="m" value="1500.0">1500 m</quantity>
(
<figureCitation id="21C54D99FFD6FFE4FB530B82FA84DCCE" box="[1273,1361,934,960]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="46" pageNumber="47">Fig. 26</figureCitation>
).
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<emphasis id="8B8A8D0EFFDDFFEFFD9509B5FCE8DEA4" bold="true" box="[575,829,400,427]" italics="true" pageId="37" pageNumber="38">Zorascar pasunepipe</emphasis>
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<subSubSection id="F1E40297FFDDFFEFFD3A09FCFC76DEFC" box="[656,931,472,499]" pageId="37" pageNumber="38" type="description">
<paragraph id="B941511CFFDDFFEFFD3A09FCFC76DEFC" blockId="37.[364,1224,400,499]" box="[656,931,472,499]" pageId="37" pageNumber="38">
<figureCitation id="21C54D99FFDDFFEFFD3A09FCFCC0DEFD" box="[656,789,472,499]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="38.[191,233,1722,1748]" captionStartId-1="39.[189,232,1712,1738]" captionStartId-2="40.[189,233,1731,1757]" captionTargetBox-0="[191,1399,265,1682]" captionTargetBox-1="[189,1398,459,1674]" captionTargetBox-2="[189,1398,478,1693]" captionTargetId-0="figure-19@38.[191,1399,265,1682]" captionTargetId-1="figure-47@39.[189,1398,459,1674]" captionTargetId-2="figure-47@40.[189,1398,478,1693]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionTargetPageId-2="40" captionText-0="Fig. 13. Zorascar pasunepipe gen. et sp. nov.; A, C, E–F = holotype male (BE_RMCA_ARA.Ara.247343; RMCA); B, D = paratype female (CASENT_9016138; CAS). A–B. Habitus, dorsal view.C–D. Prosoma, dorsal view. E. Habitus, ventral view. F. Tarsus of leg I, retrolateral view. Scale bars: A–B, E = 2 mm; C–D = 1 mm; F = 0.5 mm. Stacked microphotographs A, C, E–F by Arnaud Henrard and Rudy Jocqué; B, D by Hannah Wood." captionText-1="Fig. 14. Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ARA.Ara.247343; RMCA). A. Male right palp (mirrored), retrolateral view. B. Idem, ventral view. C. Idem, dorsal view. D. Idem, bulbus, retrolateral view. E. Idem, ventral view. F. Idem, prolateral view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–C = 0.5mm; D–F = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." captionText-2="Fig. 15. Zorascar pasunepipe gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247343; RMCA), micro-CT scans of right male palp (mirrored). A. Retrolateral view. B. Ventral view. C. Prolateral view. D. Bulbus, retrolateral view.E. Bulbus, ventral view.F. Bulbus, apical view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–C = 0.5 mm; D–F = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi-0="http://doi.org/10.5281/zenodo.14001823" figureDoi-1="http://doi.org/10.5281/zenodo.14001825" figureDoi-2="http://doi.org/10.5281/zenodo.14001827" httpUri-0="https://zenodo.org/record/14001823/files/figure.png" httpUri-1="https://zenodo.org/record/14001825/files/figure.png" httpUri-2="https://zenodo.org/record/14001827/files/figure.png" pageId="37" pageNumber="38">Figs 13–15</figureCitation>
,
<figureCitation id="21C54D99FFDDFFEFFC8B09FCFCAEDEFC" box="[801,891,472,498]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; A–B = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); C–D = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–B = 2 mm; C–E = 0.5 mm. Illustrations A–B, E by Alain Reygel; C–D by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="37" pageNumber="38">16A–D</figureCitation>
,
<figureCitation id="21C54D99FFDDFFEFFC2C09FCFC76DEFC" box="[902,931,472,498]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="37" pageNumber="38">26</figureCitation>
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<emphasis id="8B8A8D0EFFDDFFEFFF170A3EFEE3DD3A" bold="true" box="[189,310,538,564]" pageId="37" pageNumber="38">Diagnosis</emphasis>
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The male of
<taxonomicName id="7EFE2A9FFFDDFFEFFEFF0A6FFD9BDD6A" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[341,590,587,612]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pasunepipe" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFDDFFEFFEFF0A6FFD9BDD6A" box="[341,590,587,612]" italics="true" pageId="37" pageNumber="38">Zorascar pasunepipe</emphasis>
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is easily recognized by the presence of an enormous and tortuous cymbial retrobasal extension (CE), extending most of the tibial length (
<figureCitation id="21C54D99FFDDFFEFFB2A0A4AFAC4DD86" box="[1152,1297,622,648]" captionStart="Fig" captionStartId="39.[189,232,1712,1738]" captionTargetBox="[189,1398,459,1674]" captionTargetId="figure-47@39.[189,1398,459,1674]" captionTargetPageId="39" captionText="Fig. 14. Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ARA.Ara.247343; RMCA). A. Male right palp (mirrored), retrolateral view. B. Idem, ventral view. C. Idem, dorsal view. D. Idem, bulbus, retrolateral view. E. Idem, ventral view. F. Idem, prolateral view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–C = 0.5mm; D–F = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001825" httpUri="https://zenodo.org/record/14001825/files/figure.png" pageId="37" pageNumber="38">Figs 14A–C</figureCitation>
,
<figureCitation id="21C54D99FFDDFFEFFAB10A4AFAA4DD87" box="[1307,1393,622,649]" captionStart="Fig" captionStartId="40.[189,233,1731,1757]" captionTargetBox="[189,1398,478,1693]" captionTargetId="figure-47@40.[189,1398,478,1693]" captionTargetPageId="40" captionText="Fig. 15. Zorascar pasunepipe gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247343; RMCA), micro-CT scans of right male palp (mirrored). A. Retrolateral view. B. Ventral view. C. Prolateral view. D. Bulbus, retrolateral view.E. Bulbus, ventral view.F. Bulbus, apical view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–C = 0.5 mm; D–F = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001827" httpUri="https://zenodo.org/record/14001827/files/figure.png" pageId="37" pageNumber="38">15A–B</figureCitation>
,
<figureCitation id="21C54D99FFDDFFEFFF110AB6FEC0DDA2" box="[187,277,658,684]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; A–B = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); C–D = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–B = 2 mm; C–E = 0.5 mm. Illustrations A–B, E by Alain Reygel; C–D by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="37" pageNumber="38">16A–B</figureCitation>
). The female is provided with an epigyne (
<figureCitation id="21C54D99FFDDFFEFFCAA0AB6FCBDDDA2" box="[768,872,658,684]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; A–B = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); C–D = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–B = 2 mm; C–E = 0.5 mm. Illustrations A–B, E by Alain Reygel; C–D by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="37" pageNumber="38">Fig. 16C</figureCitation>
) with the median lobe (ML) that is broad and shallowly concave medially, copulatory openings (CO) exposed anteromedially (vs the female epigyne of
<taxonomicName id="7EFE2A9FFFDDFFEFFF770AFEFE71DDFD" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[221,420,730,755]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pasunepomme" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFDDFFEFFF770AFEFE71DDFD" box="[221,420,730,755]" italics="true" pageId="37" pageNumber="38">Z. pasunepomme</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFDDFFEFFE010AFEFD8EDDFA" box="[427,603,730,756]" pageId="37" pageNumber="38" rank="species">gen. et sp. nov.</taxonomicNameLabel>
has a ML that is narrow and deeply concave medially, and with CO exposed laterad of the ML,
<figureCitation id="21C54D99FFDDFFEFFDAB0AD9FDBFDC19" box="[513,618,765,791]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; A–B = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); C–D = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–B = 2 mm; C–E = 0.5 mm. Illustrations A–B, E by Alain Reygel; C–D by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="37" pageNumber="38">Fig. 16E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFDDFFEFFF170B64FBCFDCDF" pageId="37" pageNumber="38" type="etymology">
<paragraph id="B941511CFFDDFFEFFF170B64FE91DC54" blockId="37.[189,324,832,858]" box="[189,324,832,858]" pageId="37" pageNumber="38">
<heading id="E209E670FFDDFFEFFF170B64FE91DC54" bold="true" box="[189,324,832,858]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="8B8A8D0EFFDDFFEFFF170B64FE91DC54" bold="true" box="[189,324,832,858]" pageId="37" pageNumber="38">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFDDFFEFFF170B4BFBCFDCDF" blockId="37.[188,1398,879,977]" pageId="37" pageNumber="38">
The species name, a noun in apposition, refers to the famous painting of René Magritte depicting a pipe, named ‘Ceci n’est pas une pipe’, to which the cymbial extension is reminiscent, at least according the artist who made the drawings of
<taxonomicName id="7EFE2A9FFFDDFFEFFD970B9CFD7CDCDF" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[573,681,952,977]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8B8A8D0EFFDDFFEFFD970B9CFD7CDCDF" box="[573,681,952,977]" italics="true" pageId="37" pageNumber="38">Zorascar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFDDFFEFFD1A0B93FCCEDCDF" box="[688,795,951,977]" pageId="37" pageNumber="38" rank="genus">gen. nov.</taxonomicNameLabel>
for the present paper.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFDDFFEFFF170BDDFD2CD9C7" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="B941511CFFDDFFEFFF170BDDFEB9DB1D" blockId="37.[189,364,1017,1043]" box="[189,364,1017,1043]" pageId="37" pageNumber="38">
<heading id="E209E670FFDDFFEFFF170BDDFEB9DB1D" bold="true" box="[189,364,1017,1043]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="8B8A8D0EFFDDFFEFFF170BDDFEB9DB1D" bold="true" box="[189,364,1017,1043]" pageId="37" pageNumber="38">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFDDFFEFFF400C0CFE89DB4C" blockId="37.[189,1399,1064,1199]" box="[234,348,1064,1090]" pageId="37" pageNumber="38">
<emphasis id="8B8A8D0EFFDDFFEFFF400C0CFE89DB4C" bold="true" box="[234,348,1064,1090]" pageId="37" pageNumber="38">
<typeStatus id="6645EFBEFFDDFFEFFF400C0CFE89DB4C" box="[234,348,1064,1090]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFDDFFEFFF170C68FBF7DBA0" blockId="37.[189,1399,1064,1199]" pageId="37" pageNumber="38">
<materialsCitation id="09965B41FFDDFFEFFF170C68FBF7DBA0" collectingDate="2019-02-06" collectingDateMax="2019-02-11" collectingDateMin="2019-02-06" collectionCode="RMCA" collectorName="M. Jocque &amp; Ara." country="Madagascar" elevation="1586" latitude="-14.31637" location="Andriakanala Forest" longLatPrecision="1" longitude="48.723026" municipality="Bamevika" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="holotype">
<collectingCountry id="C1E9118CFFDDFFEFFF170C68FE47DB68" box="[189,402,1100,1126]" name="Madagascar" pageId="37" pageNumber="38">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFDDFFEFFE1C0C68FD04DB68" bold="true" box="[438,721,1100,1126]" pageId="37" pageNumber="38">
<collectingRegion id="7B3A9FFEFFDDFFEFFE1C0C68FD04DB68" box="[438,721,1100,1126]" country="Madagascar" name="Antsiranana" pageId="37" pageNumber="38">Antsiranana Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFDDFFEFFD450C69FCDDDB69" box="[751,776,1101,1127]" count="1" pageId="37" pageNumber="38" type="male">♂</specimenCount>
;
<collectingMunicipality id="5925CB66FFDDFFEFFCBD0C68FC41DB68" box="[791,916,1100,1126]" pageId="37" pageNumber="38">Bamevika</collectingMunicipality>
,
<location id="BC2107C7FFDDFFEFFC0A0C68FB4EDB69" LSID="urn:lsid:plazi:treatment:3157E00AFFDDFFE1FD9509B5FEFCDE7A:BC2107C7FFDDFFEFFC0A0C68FB4EDB69" box="[928,1179,1100,1127]" country="Madagascar" latitude="-14.31637" longLatPrecision="1" longitude="48.723026" municipality="Bamevika" name="Andriakanala Forest" pageId="37" pageNumber="38" stateProvince="Antsiranana">Andriakanala Forest</location>
;
<geoCoordinate id="DCCA37DBFFDDFFEFFB030C6FFAA5DB68" box="[1193,1392,1099,1127]" degrees="14" direction="south" minutes="18" orientation="latitude" pageId="37" pageNumber="38" precision="1" seconds="58.932" value="-14.31637">14°18′58.932″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFDDFFEFFF170C4BFE52DB85" box="[189,391,1135,1163]" degrees="48" direction="east" minutes="43" orientation="longitude" pageId="37" pageNumber="38" precision="1" seconds="22.886" value="48.723026">48°43′22.886″ E</geoCoordinate>
;
<elevation id="32D3B62FFFDDFFEFFE3C0C54FDE6DB84" box="[406,563,1136,1163]" metricMagnitude="3" metricUnit="m" metricValue="1.5859999999999999" pageId="37" pageNumber="38" unit="m" value="1586.0">
<quantity id="7E06FCF9FFDDFFEFFE3C0C54FE24DB85" box="[406,497,1136,1163]" metricMagnitude="3" metricUnit="m" metricValue="1.5859999999999999" pageId="37" pageNumber="38" unit="m" value="1586.0">1586 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFDDFFEFFDE90C54FCD7DB84" box="[579,770,1136,1162]" pageId="37" pageNumber="38" value="2019-02-06" valueMax="2019-02-11" valueMin="2019-02-06">
<collectingDate id="DD048E34FFDDFFEFFDE90C54FCD7DB84" box="[579,770,1136,1162]" pageId="37" pageNumber="38" value="2019-02-06" valueMax="2019-02-11" valueMin="2019-02-06">6–11 Feb. 2019</collectingDate>
</date>
;
<collectorName id="140B34CAFFDDFFEFFCBA0C55FC59DB85" box="[784,908,1137,1163]" pageId="37" pageNumber="38">M. Jocque</collectorName>
leg.; hand collected around camp; DNA RJ5; BINCO_MAD_19_0079_2; BE_
<collectionCode id="DFEFC9D9FFDDFFEFFD2A0CB0FD0FDBA0" box="[640,730,1172,1198]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="37" pageNumber="38" type="Museum">RMCA</collectionCode>
_ARA.
<collectorName id="140B34CAFFDDFFEFFC870CB0FC8ADBA0" box="[813,863,1172,1198]" pageId="37" pageNumber="38">Ara.</collectorName>
247343;
<collectionCode id="DFEFC9D9FFDDFFEFFC6E0CB0FBCBDBA0" box="[964,1054,1172,1198]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="37" pageNumber="38" type="Museum">RMCA</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B941511CFFDDFFEFFF400CF2FEB2DBFE" blockId="37.[189,1398,1238,1408]" box="[234,359,1238,1264]" pageId="37" pageNumber="38">
<emphasis id="8B8A8D0EFFDDFFEFFF400CF2FEB2DBFE" bold="true" box="[234,359,1238,1264]" pageId="37" pageNumber="38">
<typeStatus id="6645EFBEFFDDFFEFFF400CF2FEB2DBFE" box="[234,359,1238,1264]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFDDFFEFFF170CDEFBF4DA8E" blockId="37.[189,1398,1238,1408]" pageId="37" pageNumber="38">
<materialsCitation id="09965B41FFDDFFEFFF170CDEFCA7DA36" collectingDate="2019-02-06" collectingDateMax="2019-02-11" collectingDateMin="2019-02-06" collectionCode="RMCA" collectorName="Ara." country="Madagascar" elevation="1586" latitude="-14.31637" location="Andriakanala Forest" longLatPrecision="1" longitude="48.723026" municipality="Bamevika" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<collectingCountry id="C1E9118CFFDDFFEFFF170CDEFE45DA1A" box="[189,400,1274,1300]" name="Madagascar" pageId="37" pageNumber="38">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFDDFFEFFE190CDEFD1EDA1A" bold="true" box="[435,715,1274,1300]" pageId="37" pageNumber="38">
<collectingRegion id="7B3A9FFEFFDDFFEFFE190CDEFD1EDA1A" box="[435,715,1274,1300]" country="Madagascar" name="Antsiranana" pageId="37" pageNumber="38">Antsiranana Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFDDFFEFFD4C0CDEFCFBDA1A" box="[742,814,1274,1300]" count="1" pageId="37" pageNumber="38" type="generic">1 juv.</specimenCount>
; same data as for holotype; DNA RJ1; BINCO_ MAD_19_0079_2; BE_
<collectionCode id="DFEFC9D9FFDDFFEFFE720D3AFDE7DA36" box="[472,562,1310,1336]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="37" pageNumber="38" type="Museum">RMCA</collectionCode>
_ARA.
<collectorName id="140B34CAFFDDFFEFFD2F0D3AFD62DA36" box="[645,695,1310,1336]" pageId="37" pageNumber="38">Ara.</collectorName>
247344;
<collectionCode id="DFEFC9D9FFDDFFEFFCB70D3AFCA7DA36" box="[797,882,1310,1336]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="37" pageNumber="38" type="Museum">RMCA</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFDDFFEFFC210D3AFBCBDA8E" collectingDate="1998-11-08" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="400" latitude="-13.966666" location="Reserve Speciale Manongarivo" longLatPrecision="1289" longitude="48.433334" pageId="37" pageNumber="38" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Antsiranana" typeStatus="paratype">
<specimenCount id="AFF89A95FFDDFFEFFC210D3AFC6DDA37" box="[907,952,1310,1337]" count="1" pageId="37" pageNumber="38" type="male">1 ♂</specimenCount>
,
<specimenCount id="AFF89A95FFDDFFEFFC6F0D3AFC27DA37" box="[965,1010,1310,1337]" count="1" pageId="37" pageNumber="38" type="female">1 ♀</specimenCount>
;
<location id="BC2107C7FFDDFFEFFBAB0D3AFAA7DA36" LSID="urn:lsid:plazi:treatment:3157E00AFFDDFFE1FD9509B5FEFCDE7A:BC2107C7FFDDFFEFFBAB0D3AFAA7DA36" box="[1025,1394,1310,1336]" country="Madagascar" latitude="-13.966666" longLatPrecision="1289" longitude="48.433334" name="Reserve Speciale Manongarivo" pageId="37" pageNumber="38" stateProvince="Antsiranana">Réserve Spéciale Manongarivo</location>
,
<locationDeviation id="A8F3483EFFDDFFEFFF170D66FDB5DA52" box="[189,608,1346,1372]" location="Antanambao" pageId="37" pageNumber="38">
<quantity id="7E06FCF9FFDDFFEFFF170D66FEF4DA52" box="[189,289,1346,1372]" metricMagnitude="4" metricUnit="m" metricValue="1.08" pageId="37" pageNumber="38" unit="km" value="10.8">10.8 km</quantity>
229° SW of Antanambao
</locationDeviation>
;
<geoCoordinate id="DCCA37DBFFDDFFEFFDC40D64FD0CDA52" box="[622,729,1344,1372]" degrees="13" direction="south" minutes="58" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-13.966666">13°58′ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFDDFFEFFD420D64FC80DA53" box="[744,853,1344,1373]" degrees="48" direction="east" minutes="26" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="48.433334">48°26′ E</geoCoordinate>
;
<elevation id="32D3B62FFFDDFFEFFCCF0D66FC23DA52" box="[869,1014,1346,1373]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="37" pageNumber="38" unit="m" value="400.0">
<quantity id="7E06FCF9FFDDFFEFFCCF0D66FC67DA53" box="[869,946,1346,1373]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="37" pageNumber="38" unit="m" value="400.0">400 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFDDFFEFFBAC0D66FB75DA52" box="[1030,1184,1346,1373]" pageId="37" pageNumber="38" value="1998-11-08">
<collectingDate id="DD048E34FFDDFFEFFBAC0D66FB75DA52" box="[1030,1184,1346,1373]" pageId="37" pageNumber="38" value="1998-11-08">8 Nov. 1998</collectingDate>
</date>
;
<collectorName id="140B34CAFFDDFFEFFB040D67FAEDDA52" box="[1198,1336,1346,1373]" pageId="37" pageNumber="38">B.L. Fisher</collectorName>
leg.; rainforest; EC19 sifted litter; [SEM stub 30:4]; CASENT_9016138;
<collectionCode id="DFEFC9D9FFDDFFEFFC4B0D42FBCBDA8E" box="[993,1054,1382,1408]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="38" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="B941511CFFDDFFEFFF170D8CFE2CDACC" blockId="37.[189,505,1448,1474]" box="[189,505,1448,1474]" pageId="37" pageNumber="38">
<heading id="E209E670FFDDFFEFFF170D8CFE2CDACC" bold="true" box="[189,505,1448,1474]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="8B8A8D0EFFDDFFEFFF170D8CFE2CDACC" bold="true" box="[189,505,1448,1474]" pageId="37" pageNumber="38">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFDDFFEFFF170DF3FD2CD9C7" blockId="37.[189,1399,1495,1737]" pageId="37" pageNumber="38">
<materialsCitation id="09965B41FFDDFFEFFF170DF3FC0ED937" collectingDate="1998-11-08" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="400" latitude="-13.966666" location="Reserve Speciale Manongarivo" longLatPrecision="1289" longitude="48.433334" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana">
<collectingCountry id="C1E9118CFFDDFFEFFF170DF3FE5BDAFF" box="[189,398,1495,1521]" name="Madagascar" pageId="37" pageNumber="38">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFDDFFEFFE030DF3FD6FDAFF" bold="true" box="[425,698,1495,1521]" pageId="37" pageNumber="38">
<collectingRegion id="7B3A9FFEFFDDFFEFFE030DF3FD6FDAFF" box="[425,698,1495,1521]" country="Madagascar" name="Antsiranana" pageId="37" pageNumber="38">Antsiranana Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFDDFFEFFD780DF3FCD4DAFC" box="[722,769,1495,1522]" count="1" pageId="37" pageNumber="38" type="male">1 ♂</specimenCount>
;
<location id="BC2107C7FFDDFFEFFCA60DF3FBA8DAFF" LSID="urn:lsid:plazi:treatment:3157E00AFFDDFFE1FD9509B5FEFCDE7A:BC2107C7FFDDFFEFFCA60DF3FBA8DAFF" box="[780,1149,1495,1521]" country="Madagascar" latitude="-13.966666" longLatPrecision="1289" longitude="48.433334" name="Reserve Speciale Manongarivo" pageId="37" pageNumber="38" stateProvince="Antsiranana">Réserve Spéciale Manongarivo</location>
,
<locationDeviation id="A8F3483EFFDDFFEFFB2D0DF3FE80D91B" location="Antanambao" pageId="37" pageNumber="38">
<quantity id="7E06FCF9FFDDFFEFFB2D0DF3FB33DAFF" box="[1159,1254,1495,1521]" metricMagnitude="4" metricUnit="m" metricValue="1.08" pageId="37" pageNumber="38" unit="km" value="10.8">10.8 km</quantity>
229° SW of Antanambao
</locationDeviation>
,
<geoCoordinate id="DCCA37DBFFDDFFEFFEF50DDEFE16D91B" box="[351,451,1530,1558]" degrees="13" direction="south" minutes="58" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-13.966666">13°58′ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFDDFFEFFE640DDEFDE0D918" box="[462,565,1530,1558]" degrees="48" direction="east" minutes="26" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="48.433334">48°26′ E</geoCoordinate>
;
<elevation id="32D3B62FFFDDFFEFFDEA0DDFFD13D91B" box="[576,710,1531,1558]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="37" pageNumber="38" unit="m" value="400.0">
<quantity id="7E06FCF9FFDDFFEFFDEA0DDFFD5DD918" box="[576,648,1531,1558]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="37" pageNumber="38" unit="m" value="400.0">400 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFDDFFEFFD7B0DDFFCB4D91B" box="[721,865,1531,1558]" pageId="37" pageNumber="38" value="1998-11-08">
<collectingDate id="DD048E34FFDDFFEFFD7B0DDFFCB4D91B" box="[721,865,1531,1558]" pageId="37" pageNumber="38" value="1998-11-08">8 Nov. 1998</collectingDate>
</date>
;
<collectorName id="140B34CAFFDDFFEFFCC10DD8FC3BD91B" box="[875,1006,1531,1558]" pageId="37" pageNumber="38">B.L. Fisher</collectorName>
leg.; rainforest; EC19 sifted litter; [male palp drawn; male stacked images]; CASENT_9016135;
<collectionCode id="DFEFC9D9FFDDFFEFFC080E3BFC0ED937" box="[930,987,1567,1593]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="38" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFDDFFEFFC5F0E3BFE3BD953" collectingDate="1998-11-08" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="400" latitude="-13.966666" location="Reserve Speciale Manongarivo" longLatPrecision="1289" longitude="48.433334" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana">
<specimenCount id="AFF89A95FFDDFFEFFC5F0E3BFBF0D934" box="[1013,1061,1567,1594]" count="1" pageId="37" pageNumber="38" type="male">1 ♂</specimenCount>
; same data as for preceding; CASENT_9016136;
<collectionCode id="DFEFC9D9FFDDFFEFFE1F0E67FE3BD953" box="[437,494,1603,1629]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="38" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFDDFFEFFDA20E67FB14D953" box="[520,1217,1603,1630]" collectingDate="1998-11-08" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="400" latitude="-13.966666" location="Reserve Speciale Manongarivo" longLatPrecision="1289" longitude="48.433334" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana">
<specimenCount id="AFF89A95FFDDFFEFFDA20E67FDEDD950" box="[520,568,1603,1630]" count="1" pageId="37" pageNumber="38" type="male">1 ♂</specimenCount>
; same data as for preceding; CASENT_9016137;
<collectionCode id="DFEFC9D9FFDDFFEFFB220E67FB14D953" box="[1160,1217,1603,1629]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="38" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFDDFFEFFB710E67FB37D9AB" collectingDate="1998-10-20" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="1175" latitude="-14.0" location="Reserve Speciale Manongarivo" longLatPrecision="1289" longitude="48.433334" pageId="37" pageNumber="38" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana">
<specimenCount id="AFF89A95FFDDFFEFFB710E67FADCD950" box="[1243,1289,1603,1630]" count="1" pageId="37" pageNumber="38" type="female">1 ♀</specimenCount>
;
<location id="BC2107C7FFDDFFEFFAB20E67FE19D98F" LSID="urn:lsid:plazi:treatment:3157E00AFFDDFFE1FD9509B5FEFCDE7A:BC2107C7FFDDFFEFFAB20E67FE19D98F" country="Madagascar" latitude="-14.0" longLatPrecision="1289" longitude="48.433334" name="Reserve Speciale Manongarivo" pageId="37" pageNumber="38" stateProvince="Antsiranana">Réserve Spéciale Manongarivo</location>
,
<locationDeviation id="A8F3483EFFDDFFEFFE700E43FCA0D98F" box="[474,885,1639,1666]" location="Antanambao" pageId="37" pageNumber="38">
<quantity id="7E06FCF9FFDDFFEFFE700E43FDE9D98F" box="[474,572,1639,1666]" metricMagnitude="4" metricUnit="m" metricValue="1.45" pageId="37" pageNumber="38" unit="km" value="14.5">14.5 km</quantity>
220° SW of Antanambao
</locationDeviation>
;
<geoCoordinate id="DCCA37DBFFDDFFEFFC290E41FC3ED98F" box="[899,1003,1637,1665]" degrees="13" direction="south" minutes="60" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-14.0">13°60′ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFDDFFEFFC500E41FBB0D98C" box="[1018,1125,1637,1666]" degrees="48" direction="east" minutes="26" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="48.433334">48°26′ E</geoCoordinate>
;
<elevation id="32D3B62FFFDDFFEFFBD90E43FADAD98F" box="[1139,1295,1639,1666]" metricMagnitude="3" metricUnit="m" metricValue="1.175" pageId="37" pageNumber="38" unit="m" value="1175.0">
<quantity id="7E06FCF9FFDDFFEFFBD90E43FB18D98C" box="[1139,1229,1639,1666]" metricMagnitude="3" metricUnit="m" metricValue="1.175" pageId="37" pageNumber="38" unit="m" value="1175.0">1175 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFDDFFEFFAB40E43FF2ED9AB" pageId="37" pageNumber="38" value="1998-10-20">
<collectingDate id="DD048E34FFDDFFEFFAB40E43FF2ED9AB" pageId="37" pageNumber="38" value="1998-10-20">20 Oct. 1998</collectingDate>
</date>
;
<collectorName id="140B34CAFFDDFFEFFEA20EA8FE5BD9AB" box="[264,398,1675,1702]" pageId="37" pageNumber="38">B.L. Fisher</collectorName>
leg.; rainforest; EB09 sifted litter; BLF1938; CASENT_9016139;
<collectionCode id="DFEFC9D9FFDDFFEFFB030EAFFB37D9AB" box="[1193,1250,1675,1701]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="38" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFDDFFEFFB570EAFFD23D9C7" collectingDate="1998-10-20" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="1175" latitude="-14.0" location="Reserve Speciale Manongarivo" longLatPrecision="1289" longitude="48.433334" pageId="37" pageNumber="38" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana">
<specimenCount id="AFF89A95FFDDFFEFFB570EAFFAFED9A8" box="[1277,1323,1675,1702]" count="1" pageId="37" pageNumber="38" type="female">1 ♀</specimenCount>
; same data as for preceding; CASENT_9016141;
<collectionCode id="DFEFC9D9FFDDFFEFFD130E8BFD23D9C7" box="[697,758,1711,1737]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="38" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFDDFFE0FF170ED5FCA4D8E6" lastPageId="42" lastPageNumber="43" pageId="37" pageNumber="38" type="description">
<paragraph id="B941511CFFDDFFEFFF170ED5FE9BD805" blockId="37.[189,334,1777,1803]" box="[189,334,1777,1803]" pageId="37" pageNumber="38">
<heading id="E209E670FFDDFFEFFF170ED5FE9BD805" bold="true" box="[189,334,1777,1803]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="8B8A8D0EFFDDFFEFFF170ED5FE9BD805" bold="true" box="[189,334,1777,1803]" pageId="37" pageNumber="38">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFDDFFEFFF400F04FC52D834" blockId="37.[189,1160,1824,1887]" box="[234,903,1824,1850]" pageId="37" pageNumber="38">
<emphasis id="8B8A8D0EFFDDFFEFFF400F04FE48D834" bold="true" box="[234,413,1824,1850]" pageId="37" pageNumber="38">
Male
<typeStatus id="6645EFBEFFDDFFEFFE9B0F04FE48D834" box="[305,413,1824,1850]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
</emphasis>
(BE_RMCA_ARA.Ara.247343; RMCA)
</paragraph>
<paragraph id="B941511CFFDDFFEFFF170F60FB5DD851" blockId="37.[189,1160,1824,1887]" box="[189,1160,1860,1887]" pageId="37" pageNumber="38">MEASUREMENTS. Total length: 7.81. Carapace: length 3.91, width 2.91, height 1.85.</paragraph>
<paragraph id="B941511CFFDDFFEDFF170FA3FD1CDE48" blockId="37.[188,1400,1926,2025]" lastBlockId="39.[189,1398,265,327]" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
COLORATION (
<figureCitation id="21C54D99FFDDFFEFFEC80FA2FDC6D8AF" box="[354,531,1926,1953]" captionStart="Fig" captionStartId="38.[191,233,1722,1748]" captionTargetBox="[191,1399,265,1682]" captionTargetId="figure-19@38.[191,1399,265,1682]" captionTargetPageId="38" captionText="Fig. 13. Zorascar pasunepipe gen. et sp. nov.; A, C, E–F = holotype male (BE_RMCA_ARA.Ara.247343; RMCA); B, D = paratype female (CASENT_9016138; CAS). A–B. Habitus, dorsal view.C–D. Prosoma, dorsal view. E. Habitus, ventral view. F. Tarsus of leg I, retrolateral view. Scale bars: A–B, E = 2 mm; C–D = 1 mm; F = 0.5 mm. Stacked microphotographs A, C, E–F by Arnaud Henrard and Rudy Jocqué; B, D by Hannah Wood." figureDoi="http://doi.org/10.5281/zenodo.14001823" httpUri="https://zenodo.org/record/14001823/files/figure.png" pageId="37" pageNumber="38">Figs 13A, C, E</figureCitation>
). Carapace brownish yellow, with two faint, longitudinal dark bands along margin and between PME and fovea; chelicerae brownish with two faint longitudinal darker stripes; sternum brownish yellow, with faint radiating stripes; legs brownish yellow with three faint darker rings on femora and two on tibias, distally and proximally; abdomen brownish yellow with ill-defined pattern on dorsum and sides; spinnerets pale yellow.
</paragraph>
<caption id="ED810194FFDEFFECFF150E9EFE04D86F" ID-DOI="http://doi.org/10.5281/zenodo.14001823" ID-Zenodo-Dep="14001823" httpUri="https://zenodo.org/record/14001823/files/figure.png" pageId="38" pageNumber="39" startId="38.[191,233,1722,1748]" targetBox="[191,1399,265,1682]" targetPageId="38">
<paragraph id="B941511CFFDEFFECFF150E9EFE04D86F" blockId="38.[189,1399,1722,1889]" pageId="38" pageNumber="39">
<emphasis id="8B8A8D0EFFDEFFECFF150E9EFECDD9DA" bold="true" box="[191,280,1722,1748]" pageId="38" pageNumber="39">Fig. 13.</emphasis>
<taxonomicName id="7EFE2A9FFFDEFFECFEB70E9FFDC4D9DA" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[285,529,1723,1748]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pasunepipe" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFDEFFECFEB70E9FFDC4D9DA" box="[285,529,1723,1748]" italics="true" pageId="38" pageNumber="39">Zorascar pasunepipe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFDEFFECFDBC0E9EFD17D9DA" box="[534,706,1722,1748]" pageId="38" pageNumber="39" rank="species">gen. et sp. nov.</taxonomicNameLabel>
; A, C, E–F
<emphasis id="8B8A8D0EFFDEFFECFCE80E9EFC86D9DA" bold="true" box="[834,851,1722,1748]" pageId="38" pageNumber="39">=</emphasis>
holotype male (BE_RMCA_ARA.Ara.247343; RMCA); B, D
<emphasis id="8B8A8D0EFFDEFFECFECF0EF9FEA3D9F9" bold="true" box="[357,374,1757,1783]" pageId="38" pageNumber="39">=</emphasis>
paratype female (CASENT_9016138; CAS).
<emphasis id="8B8A8D0EFFDEFFECFC2D0EF9FC6BD9F9" bold="true" box="[903,958,1757,1783]" pageId="38" pageNumber="39">A–B</emphasis>
. Habitus, dorsal view.
<emphasis id="8B8A8D0EFFDEFFECFB680EF9FB2ED9F9" bold="true" box="[1218,1275,1757,1783]" pageId="38" pageNumber="39">C–D</emphasis>
. Prosoma, dorsal view.
<emphasis id="8B8A8D0EFFDEFFECFEFF0F24FEBDD814" bold="true" box="[341,360,1792,1818]" pageId="38" pageNumber="39">E</emphasis>
. Habitus, ventral view.
<emphasis id="8B8A8D0EFFDEFFECFD280F24FD46D814" bold="true" box="[642,659,1792,1818]" pageId="38" pageNumber="39">F</emphasis>
. Tarsus of leg I, retrolateral view. Scale bars: A–B, E = 2 mm; C–D = 1 mm; F = 0.5 mm. Stacked microphotographs A, C, E–F by Arnaud Henrard and Rudy Jocqué; B, D by Hannah Wood.
</paragraph>
</caption>
<paragraph id="B941511CFFDFFFEDFF17094AFB9ADE86" blockId="39.[189,1103,366,392]" box="[189,1103,366,392]" pageId="39" pageNumber="40">
STERNUM (
<figureCitation id="21C54D99FFDFFFEDFE97094AFE76DE86" box="[317,419,366,392]" captionStart="Fig" captionStartId="38.[191,233,1722,1748]" captionTargetBox="[191,1399,265,1682]" captionTargetId="figure-19@38.[191,1399,265,1682]" captionTargetPageId="38" captionText="Fig. 13. Zorascar pasunepipe gen. et sp. nov.; A, C, E–F = holotype male (BE_RMCA_ARA.Ara.247343; RMCA); B, D = paratype female (CASENT_9016138; CAS). A–B. Habitus, dorsal view.C–D. Prosoma, dorsal view. E. Habitus, ventral view. F. Tarsus of leg I, retrolateral view. Scale bars: A–B, E = 2 mm; C–D = 1 mm; F = 0.5 mm. Stacked microphotographs A, C, E–F by Arnaud Henrard and Rudy Jocqué; B, D by Hannah Wood." figureDoi="http://doi.org/10.5281/zenodo.14001823" httpUri="https://zenodo.org/record/14001823/files/figure.png" pageId="39" pageNumber="40">Fig. 13E</figureCitation>
) oval with slightly sinuous margins; 1.49 wide, 1.70 long.
</paragraph>
<caption id="ED810194FFDFFFEDFF170E94FB03D875" ID-DOI="http://doi.org/10.5281/zenodo.14001825" ID-Zenodo-Dep="14001825" httpUri="https://zenodo.org/record/14001825/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,232,1712,1738]" targetBox="[189,1398,459,1674]" targetPageId="39">
<paragraph id="B941511CFFDFFFEDFF170E94FB03D875" blockId="39.[188,1399,1712,1915]" pageId="39" pageNumber="40">
<emphasis id="8B8A8D0EFFDFFFEDFF170E94FECCD9C4" bold="true" box="[189,281,1712,1738]" pageId="39" pageNumber="40">Fig. 14.</emphasis>
<taxonomicName id="7EFE2A9FFFDFFFEDFE8A0E95FDCCD9C4" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[288,537,1713,1738]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="pasunepipe" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFDFFFEDFE8A0E95FDCCD9C4" box="[288,537,1713,1738]" italics="true" pageId="39" pageNumber="40">Zorascar pasunepipe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFDFFFEDFD8A0E95FD07D9C5" box="[544,722,1713,1739]" pageId="39" pageNumber="40" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype male (BE_RMCA_ARA.Ara.247343; RMCA).
<emphasis id="8B8A8D0EFFDFFFEDFF170EF0FF07D9E0" bold="true" box="[189,210,1748,1774]" pageId="39" pageNumber="40">A</emphasis>
. Male right palp (mirrored), retrolateral view.
<emphasis id="8B8A8D0EFFDFFFEDFD500EF0FCD8D9E0" bold="true" box="[762,781,1748,1774]" pageId="39" pageNumber="40">B</emphasis>
. Idem, ventral view.
<emphasis id="8B8A8D0EFFDFFFEDFBA20EF0FBC9D9E0" bold="true" box="[1032,1052,1748,1774]" pageId="39" pageNumber="40">C</emphasis>
. Idem, dorsal view.
<emphasis id="8B8A8D0EFFDFFFEDFAA50EF0FAF6D9E0" bold="true" box="[1295,1315,1748,1774]" pageId="39" pageNumber="40">D</emphasis>
. Idem, bulbus, retrolateral view.
<emphasis id="8B8A8D0EFFDFFFEDFE4F0ED3FE2CD81F" bold="true" box="[485,505,1783,1809]" pageId="39" pageNumber="40">E</emphasis>
. Idem, ventral view.
<emphasis id="8B8A8D0EFFDFFFEDFD440ED3FCD5D81F" bold="true" box="[750,768,1783,1809]" pageId="39" pageNumber="40">F</emphasis>
. Idem, prolateral view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–C = 0.5mm; D–F = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué.
</paragraph>
</caption>
<paragraph id="B941511CFFD0FFE2FF17092DFDF7DE2D" blockId="40.[189,546,265,291]" box="[189,546,265,291]" pageId="40" pageNumber="41">CHILUM. 0.12 high, 0.20 wide.</paragraph>
<paragraph id="B941511CFFD0FFE2FF17096FFCCADE86" blockId="40.[188,1399,330,392]" pageId="40" pageNumber="41">EYES. ALE 0.18; AME 0.15; PLE 0.18; PME 0.16; ALE–AME 0.05; AME–AME 0.08; PLE–PME 0.18; PME–PME 0.10; MOQ: AW 0.33, PW 0.44, L 0.44.</paragraph>
<caption id="ED810194FFD0FFE2FF170EE7FC6DD880" ID-DOI="http://doi.org/10.5281/zenodo.14001827" ID-Zenodo-Dep="14001827" httpUri="https://zenodo.org/record/14001827/files/figure.png" pageId="40" pageNumber="41" startId="40.[189,233,1731,1757]" targetBox="[189,1398,478,1693]" targetPageId="40">
<paragraph id="B941511CFFD0FFE2FF170EE7FC6DD880" blockId="40.[187,1399,1731,1934]" pageId="40" pageNumber="41">
<emphasis id="8B8A8D0EFFD0FFE2FF170EE7FECAD9D3" bold="true" box="[189,287,1731,1757]" pageId="40" pageNumber="41">Fig. 15.</emphasis>
<taxonomicName id="7EFE2A9FFFD0FFE2FE830EE0FDF9D9D3" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[297,556,1732,1757]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pasunepipe" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFD0FFE2FE830EE0FDF9D9D3" box="[297,556,1732,1757]" italics="true" pageId="40" pageNumber="41">Zorascar pasunepipe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFD0FFE2FD9D0EE0FD2DD9D0" box="[567,760,1732,1758]" pageId="40" pageNumber="41" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype (BE_RMCA_ARA.Ara.247343; RMCA), micro-CT scans of right male palp (mirrored).
<emphasis id="8B8A8D0EFFD0FFE2FD740EC3FD26D80F" bold="true" box="[734,755,1767,1793]" pageId="40" pageNumber="41">A</emphasis>
. Retrolateral view.
<emphasis id="8B8A8D0EFFD0FFE2FC720EC3FC39D80F" bold="true" box="[984,1004,1767,1793]" pageId="40" pageNumber="41">B</emphasis>
. Ventral view.
<emphasis id="8B8A8D0EFFD0FFE2FB310EC3FB65D80F" bold="true" box="[1179,1200,1767,1793]" pageId="40" pageNumber="41">C</emphasis>
. Prolateral view.
<emphasis id="8B8A8D0EFFD0FFE2FF170F2EFF07D82A" bold="true" box="[189,210,1802,1828]" pageId="40" pageNumber="41">D</emphasis>
. Bulbus, retrolateral view.
<emphasis id="8B8A8D0EFFD0FFE2FDAA0F2EFDC6D82A" bold="true" box="[512,531,1802,1828]" pageId="40" pageNumber="41">E</emphasis>
. Bulbus, ventral view.
<emphasis id="8B8A8D0EFFD0FFE2FCBF0F2EFCF2D82A" bold="true" box="[789,807,1802,1828]" pageId="40" pageNumber="41">F</emphasis>
. Bulbus, apical view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–C = 0.5 mm; D–F = 0.2 mm. 3D micro-CT scans by Arnaud Henrard.
</paragraph>
</caption>
<paragraph id="B941511CFFD1FFE3FF17092DFDC7DE2D" blockId="41.[189,530,265,291]" box="[189,530,265,291]" pageId="41" pageNumber="42">CLYPEUS. With 14 long setae.</paragraph>
<paragraph id="B941511CFFD1FFE3FF17096FFAD9DE68" blockId="41.[189,1292,331,358]" box="[189,1292,331,358]" pageId="41" pageNumber="42">LEGS. Trochanters shallowly notched. Unpaired claws smooth, rudimentary on legs III and IV.</paragraph>
<paragraph id="B941511CFFD1FFE3FF1709AAFE4FDEA6" blockId="41.[189,410,398,424]" box="[189,410,398,424]" pageId="41" pageNumber="42">Leg measurements</paragraph>
<paragraph id="B941511CFFD1FFE3FEA0099DFD21DD72" pageId="41" pageNumber="42">
<table id="CBFEA3BCFFD10035FF61099CFD21DD72" box="[203,756,440,636]" gridcols="7" gridrows="5" pageId="41" pageNumber="42" rowsContinueIn="41.[203,950,711,907]">
<tr id="07CE535EFFD10035FF61099CFD21DEDD" box="[203,756,440,467]" gridrow="0" pageId="41" pageNumber="42" rowspan-0="1">
<th id="441F3A22FFD10035FEA0099CFEE8DEDD" box="[266,317,440,467]" gridcol="1" gridrow="0" pageId="41" pageNumber="42">F</th>
<th id="441F3A22FFD10035FEF5099CFE47DEDD" box="[351,402,440,467]" gridcol="2" gridrow="0" pageId="41" pageNumber="42">P</th>
<th id="441F3A22FFD10035FE1E099CFE32DEDD" box="[436,487,440,467]" gridcol="3" gridrow="0" pageId="41" pageNumber="42">T</th>
<th id="441F3A22FFD10035FDA2099CFDEEDEDD" box="[520,571,440,467]" gridcol="4" gridrow="0" pageId="41" pageNumber="42">Mt</th>
<th id="441F3A22FFD10035FDF7099CFD45DEDD" box="[605,656,440,467]" gridcol="5" gridrow="0" pageId="41" pageNumber="42">t</th>
<th id="441F3A22FFD10035FD18099CFD21DEDD" box="[690,756,440,467]" gridcol="6" gridrow="0" pageId="41" pageNumber="42">Total</th>
</tr>
<tr id="07CE535EFFD10035FF6109C4FD21DD0A" box="[203,756,480,516]" gridrow="1" pageId="41" pageNumber="42">
<th id="441F3A22FFD10035FF6109C4FF3FDD0A" box="[203,234,480,516]" gridcol="0" gridrow="1" pageId="41" pageNumber="42">I</th>
<td id="441F3A22FFD10035FEA009C4FEE8DD0A" box="[266,317,480,516]" gridcol="1" gridrow="1" pageId="41" pageNumber="42">3.36</td>
<td id="441F3A22FFD10035FEF509C4FE47DD0A" box="[351,402,480,516]" gridcol="2" gridrow="1" pageId="41" pageNumber="42">1.33</td>
<td id="441F3A22FFD10035FE1E09C4FE32DD0A" box="[436,487,480,516]" gridcol="3" gridrow="1" pageId="41" pageNumber="42">3.15</td>
<td id="441F3A22FFD10035FDA209C4FDEEDD0A" box="[520,571,480,516]" gridcol="4" gridrow="1" pageId="41" pageNumber="42">2.73</td>
<td id="441F3A22FFD10035FDF709C4FD45DD0A" box="[605,656,480,516]" gridcol="5" gridrow="1" pageId="41" pageNumber="42">1.05</td>
<td id="441F3A22FFD10035FD1809C4FD21DD0A" box="[690,756,480,516]" gridcol="6" gridrow="1" pageId="41" pageNumber="42">11.62</td>
</tr>
<tr id="07CE535EFFD10035FF610A2CFD21DD23" box="[203,756,520,557]" gridrow="2" pageId="41" pageNumber="42">
<th id="441F3A22FFD10035FF610A2CFF3FDD23" box="[203,234,520,557]" gridcol="0" gridrow="2" pageId="41" pageNumber="42">II</th>
<td id="441F3A22FFD10035FEA00A2CFEE8DD23" box="[266,317,520,557]" gridcol="1" gridrow="2" pageId="41" pageNumber="42">3.15</td>
<td id="441F3A22FFD10035FEF50A2CFE47DD23" box="[351,402,520,557]" gridcol="2" gridrow="2" pageId="41" pageNumber="42">1.26</td>
<td id="441F3A22FFD10035FE1E0A2CFE32DD23" box="[436,487,520,557]" gridcol="3" gridrow="2" pageId="41" pageNumber="42">2.10</td>
<td id="441F3A22FFD10035FDA20A2CFDEEDD23" box="[520,571,520,557]" gridcol="4" gridrow="2" pageId="41" pageNumber="42">2.73</td>
<td id="441F3A22FFD10035FDF70A2CFD45DD23" box="[605,656,520,557]" gridcol="5" gridrow="2" pageId="41" pageNumber="42">1.33</td>
<td id="441F3A22FFD10035FD180A2CFD21DD23" box="[690,756,520,557]" gridcol="6" gridrow="2" pageId="41" pageNumber="42">10.57</td>
</tr>
<tr id="07CE535EFFD10035FF610A14FD21DD5A" box="[203,756,560,596]" gridrow="3" pageId="41" pageNumber="42">
<th id="441F3A22FFD10035FF610A14FF3FDD5A" box="[203,234,560,596]" gridcol="0" gridrow="3" pageId="41" pageNumber="42">III</th>
<td id="441F3A22FFD10035FEA00A14FEE8DD5A" box="[266,317,560,596]" gridcol="1" gridrow="3" pageId="41" pageNumber="42">2.45</td>
<td id="441F3A22FFD10035FEF50A14FE47DD5A" box="[351,402,560,596]" gridcol="2" gridrow="3" pageId="41" pageNumber="42">0.84</td>
<td id="441F3A22FFD10035FE1E0A14FE32DD5A" box="[436,487,560,596]" gridcol="3" gridrow="3" pageId="41" pageNumber="42">1.96</td>
<td id="441F3A22FFD10035FDA20A14FDEEDD5A" box="[520,571,560,596]" gridcol="4" gridrow="3" pageId="41" pageNumber="42">2.73</td>
<td id="441F3A22FFD10035FDF70A14FD45DD5A" box="[605,656,560,596]" gridcol="5" gridrow="3" pageId="41" pageNumber="42">1.05</td>
<td id="441F3A22FFD10035FD180A14FD21DD5A" box="[690,756,560,596]" gridcol="6" gridrow="3" pageId="41" pageNumber="42">9.03</td>
</tr>
<tr id="07CE535EFFD10035FF610A7CFD21DD72" box="[203,756,600,636]" gridrow="4" pageId="41" pageNumber="42">
<th id="441F3A22FFD10035FF610A7CFF3FDD72" box="[203,234,600,636]" gridcol="0" gridrow="4" pageId="41" pageNumber="42">IV</th>
<td id="441F3A22FFD10035FEA00A7CFEE8DD72" box="[266,317,600,636]" gridcol="1" gridrow="4" pageId="41" pageNumber="42">3.50</td>
<td id="441F3A22FFD10035FEF50A7CFE47DD72" box="[351,402,600,636]" gridcol="2" gridrow="4" pageId="41" pageNumber="42">1.26</td>
<td id="441F3A22FFD10035FE1E0A7CFE32DD72" box="[436,487,600,636]" gridcol="3" gridrow="4" pageId="41" pageNumber="42">3.08</td>
<td id="441F3A22FFD10035FDA20A7CFDEEDD72" box="[520,571,600,636]" gridcol="4" gridrow="4" pageId="41" pageNumber="42">4.27</td>
<td id="441F3A22FFD10035FDF70A7CFD45DD72" box="[605,656,600,636]" gridcol="5" gridrow="4" pageId="41" pageNumber="42">1.40</td>
<td id="441F3A22FFD10035FD180A7CFD21DD72" box="[690,756,600,636]" gridcol="6" gridrow="4" pageId="41" pageNumber="42">13.51</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFD1FFE3FF170ABEFCDDDA37" blockId="41.[189,301,666,692]" lastBlockId="41.[187,1400,948,1337]" pageId="41" pageNumber="42">
Spination MALE PALP (
<figureCitation id="21C54D99FFD1FFE3FEE70B90FE07DCC1" box="[333,466,948,975]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="39.[189,232,1712,1738]" captionStartId-1="40.[189,233,1731,1757]" captionTargetBox-0="[189,1398,459,1674]" captionTargetBox-1="[189,1398,478,1693]" captionTargetId-0="figure-47@39.[189,1398,459,1674]" captionTargetId-1="figure-47@40.[189,1398,478,1693]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionText-0="Fig. 14. Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ARA.Ara.247343; RMCA). A. Male right palp (mirrored), retrolateral view. B. Idem, ventral view. C. Idem, dorsal view. D. Idem, bulbus, retrolateral view. E. Idem, ventral view. F. Idem, prolateral view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–C = 0.5mm; D–F = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." captionText-1="Fig. 15. Zorascar pasunepipe gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247343; RMCA), micro-CT scans of right male palp (mirrored). A. Retrolateral view. B. Ventral view. C. Prolateral view. D. Bulbus, retrolateral view.E. Bulbus, ventral view.F. Bulbus, apical view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–C = 0.5 mm; D–F = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi-0="http://doi.org/10.5281/zenodo.14001825" figureDoi-1="http://doi.org/10.5281/zenodo.14001827" httpUri-0="https://zenodo.org/record/14001825/files/figure.png" httpUri-1="https://zenodo.org/record/14001827/files/figure.png" pageId="41" pageNumber="42">Figs 14–15</figureCitation>
,
<figureCitation id="21C54D99FFD1FFE3FE770B90FDE2DCC0" box="[477,567,948,974]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; A–B = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); C–D = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–B = 2 mm; C–E = 0.5 mm. Illustrations A–B, E by Alain Reygel; C–D by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="41" pageNumber="42">16A–B</figureCitation>
). RTA short, emerging in center of tibia, with cylindrical base and sharp triangular apex; tibia with dorso-retrolateral excavation bordered by median row of long setae and dorsal margin of RTA, accommodating very large, pipe-shaped retrobasal cymbial extension (CE); cymbium oval with rounded apical extremity, without dorsal patch of chemosensitive setaeor retrobasal paracymbial process; subtegulum large, filling posterior third of cymbial concavity, with basal ridge (SL) interacting with the tegulum; tegulum rounded with massive, pointed, prolateral protuberance (TL); embolus (E) inserted on center of tegulum, short, stout at base, with medial groove, lamellar apically and curved retrolaterad; median apophysis (MA) strongly sclerotized, short, hooked, curved prolaterad; conductor (C) translucent, flattened, fan shaped, narrow at base and strongly widened at extremity; paraconductor (Pc) pedicel-like membranous extension originating at base of MA, running backward along MA and ending with flattened fan-shaped tip embracing E.
</paragraph>
<paragraph id="B941511CFFD1FFE3FEA00AE3FC87DC85" pageId="41" pageNumber="42">
<table id="CBFEA3BCFFD10035FF610AE3FC63DC85" box="[203,950,711,907]" gridcols="5" gridrows="5" pageId="41" pageNumber="42" rowsContinueFrom="41.[203,756,440,636]">
<tr id="07CE535EFFD10035FF610AE3FC63DDEF" box="[203,950,711,737]" gridrow="0" pageId="41" pageNumber="42" rowspan-0="1">
<th id="441F3A22FFD10035FEA00AE3FEBBDDEF" box="[266,366,711,737]" gridcol="1" gridrow="0" pageId="41" pageNumber="42">F</th>
<th id="441F3A22FFD10035FE390AE3FE0CDDEF" box="[403,473,711,737]" gridcol="2" gridrow="0" pageId="41" pageNumber="42">P</th>
<th id="441F3A22FFD10035FE570AE3FD12DDEF" box="[509,711,711,737]" gridcol="3" gridrow="0" pageId="41" pageNumber="42">T</th>
<th id="441F3A22FFD10035FCAE0AE3FC63DDEF" box="[772,950,711,737]" gridcol="4" gridrow="0" pageId="41" pageNumber="42">Mt</th>
</tr>
<tr id="07CE535EFFD10035FF610AD5FC63DC02" box="[203,950,753,780]" gridrow="1" pageId="41" pageNumber="42">
<th id="441F3A22FFD10035FF610AD5FF3FDC02" box="[203,234,753,780]" gridcol="0" gridrow="1" pageId="41" pageNumber="42">I</th>
<td id="441F3A22FFD10035FEA00AD5FEBBDC02" box="[266,366,753,780]" gridcol="1" gridrow="1" pageId="41" pageNumber="42">pl2d3rl3</td>
<td id="441F3A22FFD10035FE390AD5FE0CDC02" box="[403,473,753,780]" gridcol="2" gridrow="1" pageId="41" pageNumber="42">pl1</td>
<td id="441F3A22FFD10035FE570AD5FD12DC02" box="[509,711,753,780]" gridcol="3" gridrow="1" pageId="41" pageNumber="42">pl2d2rl2v2-2-2-2</td>
<td id="441F3A22FFD10035FCAE0AD5FC63DC02" box="[772,950,753,780]" gridcol="4" gridrow="1" pageId="41" pageNumber="42">pl2rl2v2-2</td>
</tr>
<tr id="07CE535EFFD10035FF610B38FC63DC38" box="[203,950,796,822]" gridrow="2" pageId="41" pageNumber="42">
<th id="441F3A22FFD10035FF610B38FF3FDC38" box="[203,234,796,822]" gridcol="0" gridrow="2" pageId="41" pageNumber="42">II</th>
<td id="441F3A22FFD10035FEA00B38FEBBDC38" box="[266,366,796,822]" gridcol="1" gridrow="2" pageId="41" pageNumber="42">pl3d3rl4</td>
<td id="441F3A22FFD10035FE390B38FE0CDC38" box="[403,473,796,822]" gridcol="2" gridrow="2" pageId="41" pageNumber="42">pl1</td>
<td id="441F3A22FFD10035FE570B38FD12DC38" box="[509,711,796,822]" gridcol="3" gridrow="2" pageId="41" pageNumber="42">pl2d3rl3v2-2-2-2</td>
<td id="441F3A22FFD10035FCAE0B38FC63DC38" box="[772,950,796,822]" gridcol="4" gridrow="2" pageId="41" pageNumber="42">pl2rl2v2-2-1</td>
</tr>
<tr id="07CE535EFFD10035FF610B62FC63DC6F" box="[203,950,838,865]" gridrow="3" pageId="41" pageNumber="42">
<th id="441F3A22FFD10035FF610B62FF3FDC6F" box="[203,234,838,865]" gridcol="0" gridrow="3" pageId="41" pageNumber="42">III</th>
<td id="441F3A22FFD10035FEA00B62FEBBDC6F" box="[266,366,838,865]" gridcol="1" gridrow="3" pageId="41" pageNumber="42">pl3d3rl4</td>
<td id="441F3A22FFD10035FE390B62FE0CDC6F" box="[403,473,838,865]" gridcol="2" gridrow="3" pageId="41" pageNumber="42">pl1rl1</td>
<td id="441F3A22FFD10035FE570B62FD12DC6F" box="[509,711,838,865]" gridcol="3" gridrow="3" pageId="41" pageNumber="42">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFD10035FCAE0B62FC63DC6F" box="[772,950,838,865]" gridcol="4" gridrow="3" pageId="41" pageNumber="42">pl3d1rl3v2-2-2</td>
</tr>
<tr id="07CE535EFFD10035FF610B55FC63DC85" box="[203,950,881,907]" gridrow="4" pageId="41" pageNumber="42">
<th id="441F3A22FFD10035FF610B55FF3FDC85" box="[203,234,881,907]" gridcol="0" gridrow="4" pageId="41" pageNumber="42">IV</th>
<td id="441F3A22FFD10035FEA00B55FEBBDC85" box="[266,366,881,907]" gridcol="1" gridrow="4" pageId="41" pageNumber="42">pl4d3rl1</td>
<td id="441F3A22FFD10035FE390B55FE0CDC85" box="[403,473,881,907]" gridcol="2" gridrow="4" pageId="41" pageNumber="42">pl1rl1</td>
<td id="441F3A22FFD10035FE570B55FD12DC85" box="[509,711,881,907]" gridcol="3" gridrow="4" pageId="41" pageNumber="42">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFD10035FCAE0B55FC63DC85" box="[772,950,881,907]" gridcol="4" gridrow="4" pageId="41" pageNumber="42">disp14</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFD1FFE3FF400D45FD45DA75" blockId="41.[189,1160,1377,1440]" box="[234,656,1377,1403]" pageId="41" pageNumber="42">
<emphasis id="8B8A8D0EFFD1FFE3FF400D45FE90DA75" bold="true" box="[234,325,1377,1403]" pageId="41" pageNumber="42">Female</emphasis>
(CASENT_9016138; CAS)
</paragraph>
<paragraph id="B941511CFFD1FFE3FF170DA1FB5DDA91" blockId="41.[189,1160,1377,1440]" box="[189,1160,1413,1440]" pageId="41" pageNumber="42">MEASUREMENTS. Total length: 8.50. Carapace: length 4.80, width 3.60, height 1.70.</paragraph>
<paragraph id="B941511CFFD1FFE3FF170DECFC85D9B4" blockId="41.[188,1400,1479,1722]" pageId="41" pageNumber="42">
COLOUR (
<figureCitation id="21C54D99FFD1FFE3FE990DE3FE1CDAEC" box="[307,457,1479,1506]" captionStart="Fig" captionStartId="38.[191,233,1722,1748]" captionTargetBox="[191,1399,265,1682]" captionTargetId="figure-19@38.[191,1399,265,1682]" captionTargetPageId="38" captionText="Fig. 13. Zorascar pasunepipe gen. et sp. nov.; A, C, E–F = holotype male (BE_RMCA_ARA.Ara.247343; RMCA); B, D = paratype female (CASENT_9016138; CAS). A–B. Habitus, dorsal view.C–D. Prosoma, dorsal view. E. Habitus, ventral view. F. Tarsus of leg I, retrolateral view. Scale bars: A–B, E = 2 mm; C–D = 1 mm; F = 0.5 mm. Stacked microphotographs A, C, E–F by Arnaud Henrard and Rudy Jocqué; B, D by Hannah Wood." figureDoi="http://doi.org/10.5281/zenodo.14001823" httpUri="https://zenodo.org/record/14001823/files/figure.png" pageId="41" pageNumber="42">Figs 13B, D</figureCitation>
). Carapace brownish yellow, dark along margin and with two contrasting dark bands along margin of pars cephalica, short longitudinal line behind each PME and longer faint median band between PME and fovea; chelicerae brownish-yellow, anteriorly with two thin longitudinal darker stripes; sternum yellowish white with faint median dusky oval mark; legs brownish yellow with three darker rings basally, medially and apically on femur and two basally and apically on tibia; abdomen yellow-white with spotted pattern on dorsum and sides, venter with two broken dusky longitudinal bands and dark surrounding spinnerets, spinnerets pale yellow.
</paragraph>
<paragraph id="B941511CFFD1FFE3FF170EC7FC6FD9F3" blockId="41.[189,954,1763,1789]" box="[189,954,1763,1789]" pageId="41" pageNumber="42">STERNUM. Oval with slightly sinuous sides; 2.10 wide, 2.20 long.</paragraph>
<paragraph id="B941511CFFD1FFE3FF170F01FDF7D831" blockId="41.[189,546,1829,1856]" box="[189,546,1829,1856]" pageId="41" pageNumber="42">CHILUM. 0.15 high, 0.20 wide.</paragraph>
<paragraph id="B941511CFFD1FFE3FF170F4CFCCAD8A9" blockId="41.[188,1399,1896,1959]" pageId="41" pageNumber="42">EYES. ALE 0.13; AME 0.09; PLE 0.16; PME 0.15; ALE–AME 0.08; AME–AME 0.25; PLE–PME 0.18; PME–PME 0.10; MOQ: AW 0.26, PW 0.43, L 0.35.</paragraph>
<paragraph id="B941511CFFD1FFE3FF170FEAFDFAD8E7" blockId="41.[189,559,1998,2025]" box="[189,559,1998,2025]" pageId="41" pageNumber="42">CLYPEUS. With eight long setae.</paragraph>
<paragraph id="B941511CFFD2FFE0FF17092DFE4FDE2D" blockId="42.[189,410,265,291]" box="[189,410,265,291]" pageId="42" pageNumber="43">Leg measurements</paragraph>
<paragraph id="B941511CFFD2FFE0FEA00912FD21DEF4" pageId="42" pageNumber="43">
<table id="CBFEA3BCFFD20035FF610911FD21DEF4" box="[203,756,309,506]" gridcols="7" gridrows="5" pageId="42" pageNumber="43" rowsContinueIn="42.[203,888,592,788]">
<tr id="07CE535EFFD20035FF610911FD21DE5E" box="[203,756,309,336]" gridrow="0" pageId="42" pageNumber="43" rowspan-0="1">
<th id="441F3A22FFD20035FEA00911FEE8DE5E" box="[266,317,309,336]" gridcol="1" gridrow="0" pageId="42" pageNumber="43">F</th>
<th id="441F3A22FFD20035FEF50911FE47DE5E" box="[351,402,309,336]" gridcol="2" gridrow="0" pageId="42" pageNumber="43">P</th>
<th id="441F3A22FFD20035FE1E0911FE32DE5E" box="[436,487,309,336]" gridcol="3" gridrow="0" pageId="42" pageNumber="43">T</th>
<th id="441F3A22FFD20035FDA20911FDEEDE5E" box="[520,571,309,336]" gridcol="4" gridrow="0" pageId="42" pageNumber="43">Mt</th>
<th id="441F3A22FFD20035FDF70911FD45DE5E" box="[605,656,309,336]" gridcol="5" gridrow="0" pageId="42" pageNumber="43">t</th>
<th id="441F3A22FFD20035FD180911FD21DE5E" box="[690,756,309,336]" gridcol="6" gridrow="0" pageId="42" pageNumber="43">Total</th>
</tr>
<tr id="07CE535EFFD20035FF610944FD21DE75" box="[203,756,352,379]" gridrow="1" pageId="42" pageNumber="43">
<th id="441F3A22FFD20035FF610944FF3FDE75" box="[203,234,352,379]" gridcol="0" gridrow="1" pageId="42" pageNumber="43">I</th>
<td id="441F3A22FFD20035FEA00944FEE8DE75" box="[266,317,352,379]" gridcol="1" gridrow="1" pageId="42" pageNumber="43">3.40</td>
<td id="441F3A22FFD20035FEF50944FE47DE75" box="[351,402,352,379]" gridcol="2" gridrow="1" pageId="42" pageNumber="43">1.60</td>
<td id="441F3A22FFD20035FE1E0944FE32DE75" box="[436,487,352,379]" gridcol="3" gridrow="1" pageId="42" pageNumber="43">3.30</td>
<td id="441F3A22FFD20035FDA20944FDEEDE75" box="[520,571,352,379]" gridcol="4" gridrow="1" pageId="42" pageNumber="43">2.60</td>
<td id="441F3A22FFD20035FDF70944FD45DE75" box="[605,656,352,379]" gridcol="5" gridrow="1" pageId="42" pageNumber="43">1.80</td>
<td id="441F3A22FFD20035FD180944FD21DE75" box="[690,756,352,379]" gridcol="6" gridrow="1" pageId="42" pageNumber="43">12.70</td>
</tr>
<tr id="07CE535EFFD20035FF6109AEFD21DEAB" box="[203,756,394,421]" gridrow="2" pageId="42" pageNumber="43">
<th id="441F3A22FFD20035FF6109AEFF3FDEAB" box="[203,234,394,421]" gridcol="0" gridrow="2" pageId="42" pageNumber="43">II</th>
<td id="441F3A22FFD20035FEA009AEFEE8DEAB" box="[266,317,394,421]" gridcol="1" gridrow="2" pageId="42" pageNumber="43">3.10</td>
<td id="441F3A22FFD20035FEF509AEFE47DEAB" box="[351,402,394,421]" gridcol="2" gridrow="2" pageId="42" pageNumber="43">1.40</td>
<td id="441F3A22FFD20035FE1E09AEFE32DEAB" box="[436,487,394,421]" gridcol="3" gridrow="2" pageId="42" pageNumber="43">2.60</td>
<td id="441F3A22FFD20035FDA209AEFDEEDEAB" box="[520,571,394,421]" gridcol="4" gridrow="2" pageId="42" pageNumber="43">2.50</td>
<td id="441F3A22FFD20035FDF709AEFD45DEAB" box="[605,656,394,421]" gridcol="5" gridrow="2" pageId="42" pageNumber="43">1.60</td>
<td id="441F3A22FFD20035FD1809AEFD21DEAB" box="[690,756,394,421]" gridcol="6" gridrow="2" pageId="42" pageNumber="43">11.20</td>
</tr>
<tr id="07CE535EFFD20035FF610991FD21DEDE" box="[203,756,437,464]" gridrow="3" pageId="42" pageNumber="43">
<th id="441F3A22FFD20035FF610991FF3FDEDE" box="[203,234,437,464]" gridcol="0" gridrow="3" pageId="42" pageNumber="43">III</th>
<td id="441F3A22FFD20035FEA00991FEE8DEDE" box="[266,317,437,464]" gridcol="1" gridrow="3" pageId="42" pageNumber="43">3.00</td>
<td id="441F3A22FFD20035FEF50991FE47DEDE" box="[351,402,437,464]" gridcol="2" gridrow="3" pageId="42" pageNumber="43">1.30</td>
<td id="441F3A22FFD20035FE1E0991FE32DEDE" box="[436,487,437,464]" gridcol="3" gridrow="3" pageId="42" pageNumber="43">2.10</td>
<td id="441F3A22FFD20035FDA20991FDEEDEDE" box="[520,571,437,464]" gridcol="4" gridrow="3" pageId="42" pageNumber="43">2.70</td>
<td id="441F3A22FFD20035FDF70991FD45DEDE" box="[605,656,437,464]" gridcol="5" gridrow="3" pageId="42" pageNumber="43">1.60</td>
<td id="441F3A22FFD20035FD180991FD21DEDE" box="[690,756,437,464]" gridcol="6" gridrow="3" pageId="42" pageNumber="43">10.70</td>
</tr>
<tr id="07CE535EFFD20035FF6109FBFD21DEF4" box="[203,756,479,506]" gridrow="4" pageId="42" pageNumber="43">
<th id="441F3A22FFD20035FF6109FBFF3FDEF4" box="[203,234,479,506]" gridcol="0" gridrow="4" pageId="42" pageNumber="43">IV</th>
<td id="441F3A22FFD20035FEA009FBFEE8DEF4" box="[266,317,479,506]" gridcol="1" gridrow="4" pageId="42" pageNumber="43">4.00</td>
<td id="441F3A22FFD20035FEF509FBFE47DEF4" box="[351,402,479,506]" gridcol="2" gridrow="4" pageId="42" pageNumber="43">1.40</td>
<td id="441F3A22FFD20035FE1E09FBFE32DEF4" box="[436,487,479,506]" gridcol="3" gridrow="4" pageId="42" pageNumber="43">3.60</td>
<td id="441F3A22FFD20035FDA209FBFDEEDEF4" box="[520,571,479,506]" gridcol="4" gridrow="4" pageId="42" pageNumber="43">2.90</td>
<td id="441F3A22FFD20035FDF709FBFD45DEF4" box="[605,656,479,506]" gridcol="5" gridrow="4" pageId="42" pageNumber="43">1.60</td>
<td id="441F3A22FFD20035FD1809FBFD21DEF4" box="[690,756,479,506]" gridcol="6" gridrow="4" pageId="42" pageNumber="43">13.50</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFD2FFE0FF170A07FE66DCE7" blockId="42.[189,301,547,573]" lastBlockId="42.[187,1399,830,1002]" pageId="42" pageNumber="43">
Spination GENITALIA. Epigyne (
<figureCitation id="21C54D99FFD2FFE0FE180B1AFDCDDC56" box="[434,536,830,856]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; A–B = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); C–D = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–B = 2 mm; C–E = 0.5 mm. Illustrations A–B, E by Alain Reygel; C–D by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="42" pageNumber="43">Fig. 16C</figureCitation>
) with median lobe (ML) broad and shallowly concave medially, copulatory openings (CO) exposed anteromedially; lateral lobes (LL) rounded and becoming narrowed to a blunt point medially over median sector (MS); lateral processes (LP) small, blunt. Endogyne (
<figureCitation id="21C54D99FFD2FFE0FB630BA3FAE1DCAF" box="[1225,1332,903,929]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; A–B = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); C–D = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: A–B = 2 mm; C–E = 0.5 mm. Illustrations A–B, E by Alain Reygel; C–D by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="42" pageNumber="43">Fig. 16D</figureCitation>
) with narrow, curved copulatory ducts (CD), these hidden beneath the spermathecal heads (S); fertilization ducts (FD) posterior.
</paragraph>
<paragraph id="B941511CFFD2FFE0FEA00A74FCADDC1A" pageId="42" pageNumber="43">
<table id="CBFEA3BCFFD20035FF610A74FCADDC1A" box="[203,888,592,788]" gridcols="5" gridrows="5" pageId="42" pageNumber="43" rowsContinueFrom="42.[203,756,309,506]">
<tr id="07CE535EFFD20035FF610A74FCADDD64" box="[203,888,592,618]" gridrow="0" pageId="42" pageNumber="43" rowspan-0="1">
<th id="441F3A22FFD20035FEA00A74FEB8DD64" box="[266,365,592,618]" gridcol="1" gridrow="0" pageId="42" pageNumber="43">F</th>
<th id="441F3A22FFD20035FE390A74FE76DD64" box="[403,419,592,618]" gridcol="2" gridrow="0" pageId="42" pageNumber="43">P</th>
<th id="441F3A22FFD20035FE4B0A74FD46DD64" box="[481,659,592,618]" gridcol="3" gridrow="0" pageId="42" pageNumber="43">T</th>
<th id="441F3A22FFD20035FD660A74FCADDD64" box="[716,888,592,618]" gridcol="4" gridrow="0" pageId="42" pageNumber="43">Mt</th>
</tr>
<tr id="07CE535EFFD20035FF610A5EFCADDD9B" box="[203,888,634,661]" gridrow="1" pageId="42" pageNumber="43">
<th id="441F3A22FFD20035FF610A5EFF3FDD9B" box="[203,234,634,661]" gridcol="0" gridrow="1" pageId="42" pageNumber="43">I</th>
<td id="441F3A22FFD20035FEA00A5EFEB8DD9B" box="[266,365,634,661]" gridcol="1" gridrow="1" pageId="42" pageNumber="43">pl1d12</td>
<td id="441F3A22FFD20035FE390A5EFE76DD9B" box="[403,419,634,661]" gridcol="2" gridrow="1" pageId="42" pageNumber="43">–</td>
<td id="441F3A22FFD20035FE4B0A5EFD46DD9B" box="[481,659,634,661]" gridcol="3" gridrow="1" pageId="42" pageNumber="43">v2-2-2-2</td>
<td id="441F3A22FFD20035FD660A5EFCADDD9B" box="[716,888,634,661]" gridcol="4" gridrow="1" pageId="42" pageNumber="43">v2-2-2</td>
</tr>
<tr id="07CE535EFFD20035FF610A81FCADDDB1" box="[203,888,677,703]" gridrow="2" pageId="42" pageNumber="43">
<th id="441F3A22FFD20035FF610A81FF3FDDB1" box="[203,234,677,703]" gridcol="0" gridrow="2" pageId="42" pageNumber="43">II</th>
<td id="441F3A22FFD20035FEA00A81FEB8DDB1" box="[266,365,677,703]" gridcol="1" gridrow="2" pageId="42" pageNumber="43">d1</td>
<td id="441F3A22FFD20035FE390A81FE76DDB1" box="[403,419,677,703]" gridcol="2" gridrow="2" pageId="42" pageNumber="43">–</td>
<td id="441F3A22FFD20035FE4B0A81FD46DDB1" box="[481,659,677,703]" gridcol="3" gridrow="2" pageId="42" pageNumber="43">v2-2-2-2</td>
<td id="441F3A22FFD20035FD660A81FCADDDB1" box="[716,888,677,703]" gridcol="4" gridrow="2" pageId="42" pageNumber="43">v2-2-2</td>
</tr>
<tr id="07CE535EFFD20035FF610AEBFCADDDE4" box="[203,888,719,746]" gridrow="3" pageId="42" pageNumber="43">
<th id="441F3A22FFD20035FF610AEBFF3FDDE4" box="[203,234,719,746]" gridcol="0" gridrow="3" pageId="42" pageNumber="43">III</th>
<td id="441F3A22FFD20035FEA00AEBFEB8DDE4" box="[266,365,719,746]" gridcol="1" gridrow="3" pageId="42" pageNumber="43">pl1d2rl1</td>
<td id="441F3A22FFD20035FE390AEBFE76DDE4" box="[403,419,719,746]" gridcol="2" gridrow="3" pageId="42" pageNumber="43">–</td>
<td id="441F3A22FFD20035FE4B0AEBFD46DDE4" box="[481,659,719,746]" gridcol="3" gridrow="3" pageId="42" pageNumber="43">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFD20035FD660AEBFCADDDE4" box="[716,888,719,746]" gridcol="4" gridrow="3" pageId="42" pageNumber="43">pl4rl4v2-2-2</td>
</tr>
<tr id="07CE535EFFD20035FF610ADEFCADDC1A" box="[203,888,762,788]" gridrow="4" pageId="42" pageNumber="43">
<th id="441F3A22FFD20035FF610ADEFF3FDC1A" box="[203,234,762,788]" gridcol="0" gridrow="4" pageId="42" pageNumber="43">IV</th>
<td id="441F3A22FFD20035FEA00ADEFEB8DC1A" box="[266,365,762,788]" gridcol="1" gridrow="4" pageId="42" pageNumber="43">pl1d2</td>
<td id="441F3A22FFD20035FE390ADEFE76DC1A" box="[403,419,762,788]" gridcol="2" gridrow="4" pageId="42" pageNumber="43">–</td>
<td id="441F3A22FFD20035FE4B0ADEFD46DC1A" box="[481,659,762,788]" gridcol="3" gridrow="4" pageId="42" pageNumber="43">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFD20035FD660ADEFCADDC1A" box="[716,888,762,788]" gridcol="4" gridrow="4" pageId="42" pageNumber="43">pl4rl4v2-1-2-2</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFD2FFE0FF170C36FEE6DB22" blockId="42.[189,307,1042,1068]" box="[189,307,1042,1068]" pageId="42" pageNumber="43">
<heading id="E209E670FFD2FFE0FF170C36FEE6DB22" bold="true" box="[189,307,1042,1068]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="8B8A8D0EFFD2FFE0FF170C36FEE6DB22" bold="true" box="[189,307,1042,1068]" pageId="42" pageNumber="43">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFD2FFE0FF400C66FE54DB53" blockId="42.[187,1399,1090,1552]" box="[234,385,1090,1117]" pageId="42" pageNumber="43">
<emphasis id="8B8A8D0EFFD2FFE0FF400C66FEFFDB52" bold="true" box="[234,298,1090,1116]" pageId="42" pageNumber="43">Male</emphasis>
(n = 5)
</paragraph>
<paragraph id="B941511CFFD2FFE0FF170C42FE71D91E" blockId="42.[187,1399,1090,1552]" pageId="42" pageNumber="43">
Total length = 7.81–8.60, carapace length/width = 1.21–1.34, carapace height/width = 0.46–0.56, PER/carapace width = 0.23–0.24, PER/OAL = 2.28–2.44, PER/AER = 1.40–1.44, OAL/ OQL = 1.14–1.25, OQP/OQA = 1.00–1.52, sternum length/width = 1.07–1.16, palpal coxa length/width = 2.00–2.50, femur I length/carapace width = 1.05–1.13, metatarsus I length/carapace width = 0.87–1.12, femur IV length/carapace width = 1.17–1.24, cymbium length/carapace width = 0.39–0.46, cymbium length/palpal patella length = 2.00–2.57, cymbium length/palpal tibia length = 1.60–2.29, cymbium length/palpal femur length = 0.84–1.00, palpal tibia length/palpal patella length 1.00–1.43. Male palp of the
<typeStatus id="6645EFBEFFD2FFE0FF4C0D41FE9EDA71" box="[230,331,1381,1407]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
from Andriakanala Forest and of specimens from Manongarivo. i.e.; CASENT_9016135–138 and CASENT_9016141 are very similar but differ in a few characteristics. The the
<typeStatus id="6645EFBEFFD2FFE0FBD60DADFB36DAAD" box="[1148,1251,1417,1443]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
has the RTA more narrowly pointed, the cymbial retrobasal extension longer and more twisted, the tegular prolateral locking process more narrowly triangular and the apex of the MA sharper than those features in males from Manongarivo.
</paragraph>
<paragraph id="B941511CFFD2FFE0FF400E1DFE48D95D" blockId="42.[189,1399,1593,1874]" box="[234,413,1593,1619]" pageId="42" pageNumber="43">
<emphasis id="8B8A8D0EFFD2FFE0FF400E1DFE90D95D" bold="true" box="[234,325,1593,1619]" pageId="42" pageNumber="43">Female</emphasis>
(n = 2)
</paragraph>
<paragraph id="B941511CFFD2FFE0FF170E79FCD1D85C" blockId="42.[189,1399,1593,1874]" pageId="42" pageNumber="43">Total length = 8.50–10.10, carapace length/width = 1.33–1.34, carapace height/ width = 0.47–0.58, PER/AER = 1.37–1.39, OQP/OQA = 1.48–1.62, sternum length/width = 1.05–1.12, palpal coxa length/width = 2.50–2.55, femur I length/carapace width = 0.94–0.97, metatarsus I length/carapace width = 0.72–0.76, palpal tarsus length/carapace width = 0.34–0.36, palpal tarsus length/palpal tibia length = 1.44–2.17, palpal tarsus length/palpal tibia length = 1.30–1.44, palpal tibia length/palpal patella length = 1.11–1.50. Epigyne with median lobe length 1.80–3.33 times width; distance from atrium base to epigastric fold 0.19–0.26 times epigyne length.</paragraph>
<paragraph id="B941511CFFD2FFE0FF170F5EFEABD89A" blockId="42.[189,382,1914,1940]" box="[189,382,1914,1940]" pageId="42" pageNumber="43">
<heading id="E209E670FFD2FFE0FF170F5EFEABD89A" bold="true" box="[189,382,1914,1940]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="8B8A8D0EFFD2FFE0FF170F5EFEABD89A" bold="true" box="[189,382,1914,1940]" pageId="42" pageNumber="43">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFD2FFE0FF170F8EFCA4D8E6" blockId="42.[189,1397,1962,2025]" pageId="42" pageNumber="43">
This species has been collected in forest from
<quantity id="7E06FCF9FFD2FFE0FD400F8EFCE3D8CA" box="[746,822,1962,1988]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="42" pageNumber="43" unit="m" value="400.0">400 m</quantity>
to
<quantity id="7E06FCF9FFD2FFE0FCF50F8EFC6CD8CA" box="[863,953,1962,1988]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="42" pageNumber="43" unit="m" value="1600.0">1600 m</quantity>
a.s.l. The function of the remarkable retrobasal cymbial extension on the male palp is unknown.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFD3FFE1FF17092CFEFCDE7A" pageId="43" pageNumber="44" type="distribution">
<paragraph id="B941511CFFD3FFE1FF17092CFE83DE2C" blockId="43.[189,342,264,290]" box="[189,342,264,290]" pageId="43" pageNumber="44">
<heading id="E209E670FFD3FFE1FF17092CFE83DE2C" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="8B8A8D0EFFD3FFE1FF17092CFE83DE2C" bold="true" box="[189,342,264,290]" pageId="43" pageNumber="44">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFD3FFE1FF17091CFEFCDE7A" blockId="43.[187,1398,311,372]" pageId="43" pageNumber="44">
<taxonomicName id="7EFE2A9FFFD3FFE1FF17091CFE78DE5F" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[189,429,312,337]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pasunepipe" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFD3FFE1FF17091CFE78DE5F" box="[189,429,312,337]" italics="true" pageId="43" pageNumber="44">Zorascar pasunepipe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFD3FFE1FE1B091CFD83DE5C" box="[433,598,312,338]" pageId="43" pageNumber="44" rank="species">gen. et sp. nov.</taxonomicNameLabel>
is known only from
<collectingRegion id="7B3A9FFEFFD3FFE1FC960913FBE4DE5F" box="[828,1073,311,337]" country="Madagascar" name="Antsiranana" pageId="43" pageNumber="44">Antsiranana Province</collectingRegion>
in northwestern
<collectingCountry id="C1E9118CFFD3FFE1FB410913FAA3DE5F" box="[1259,1398,311,337]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
(
<figureCitation id="21C54D99FFD3FFE1FF6E097EFECEDE7A" box="[196,283,346,372]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="43" pageNumber="44">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

830
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<mods:title id="6309CDBFCC01E2242073E84048B24CBE">On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar</mods:title>
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<mods:namePart id="50659DF25FAB1B6FEFF0DC28410BA7C7">Henrard, Arnaud</mods:namePart>
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<mods:namePart id="795B091944C86AEC655BB68A7BBBBDF5">Griswold, Charles</mods:namePart>
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<mods:namePart id="0E4CD813E6A0D9A4CAD1CEAE4DD3262B">Jocqué, Rudy</mods:namePart>
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<taxonomicName id="7EFE2A9FFFE5FFD7FDEC092CFCE3DE2D" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[582,822,264,291]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="polotowae" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFE5FFD7FDEC092CFCE3DE2D" bold="true" box="[582,822,264,291]" italics="true" pageId="29" pageNumber="30">Tabiboka polotowae</emphasis>
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<paragraph id="B941511CFFE5FFD7FD680976FCA4DE62" blockId="29.[376,1212,264,365]" box="[706,881,338,365]" pageId="29" pageNumber="30">
<figureCitation id="21C54D99FFE5FFD7FD680976FC92DE63" box="[706,839,338,365]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="30.[191,235,1928,1954]" captionStartId-1="31.[189,231,1719,1745]" captionStartId-2="32.[189,232,1717,1743]" captionTargetBox-0="[189,1398,529,1888]" captionTargetBox-1="[211,1371,542,1665]" captionTargetBox-2="[277,1311,265,1679]" captionTargetId-0="figure-67@30.[189,1398,529,1888]" captionTargetId-1="figure-76@31.[189,1398,527,1680]" captionTargetId-2="figure-18@32.[277,1311,265,1679]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="32" captionText-0="Fig. 10. Tabiboka polotowae gen. et sp. nov., male and female habitus; A, C = male paratype (CASENT_9016118; CAS); B, D = female paratype (CASENT_9016116; CAS). A–B. Dorsal view. C–D. Ventral view. Scale bars = 2.5 mm. Stacked microphotographs by Franklyn Cala Riquelme." captionText-1="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." captionText-2="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; A–F = male holotype (CASENT_9016117; CAS); G–H= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–D = 0.5 mm; E–F = 0.15 mm; G–H = 0.1 mm. SEM images A–F by Charles Griswold and Darrell Ubick, G–H by Daniele Polotow." figureDoi-0="http://doi.org/10.5281/zenodo.14001815" figureDoi-1="http://doi.org/10.5281/zenodo.14001819" figureDoi-2="http://doi.org/10.5281/zenodo.14001821" httpUri-0="https://zenodo.org/record/14001815/files/figure.png" httpUri-1="https://zenodo.org/record/14001819/files/figure.png" httpUri-2="https://zenodo.org/record/14001821/files/figure.png" pageId="29" pageNumber="30">Figs 10–12</figureCitation>
,
<figureCitation id="21C54D99FFE5FFD7FCFE0976FCA4DE62" box="[852,881,338,364]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="29" pageNumber="30">26</figureCitation>
</paragraph>
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<taxonomicName id="7EFE2A9FFFE5FFD7FF1709B1FE4CDEBE" box="[189,409,405,432]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="undefined-3">
<emphasis id="8B8A8D0EFFE5FFD7FF1709B1FE9EDEA1" box="[189,331,405,431]" italics="true" pageId="29" pageNumber="30">Zorodictyna</emphasis>
spp. 3
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and 4 –
<bibRefCitation id="DD6F2CEDFFE5FFD7FE5409B1FD32DEBE" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[510,743,405,432]" pageId="29" pageNumber="30" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow
<emphasis id="8B8A8D0EFFE5FFD7FDCD09B2FD77DEA1" box="[615,674,405,431]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2015
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFE5FFD7FF1709FCFBD5DC2A" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="B941511CFFE5FFD7FF1709FCFEE3DEFC" blockId="29.[189,310,472,498]" box="[189,310,472,498]" pageId="29" pageNumber="30">
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<emphasis id="8B8A8D0EFFE5FFD7FF1709FCFEE3DEFC" bold="true" box="[189,310,472,498]" pageId="29" pageNumber="30">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE5FFD7FF170A2CFBD5DC2A" blockId="29.[187,1399,520,805]" pageId="29" pageNumber="30">
The males of
<taxonomicName id="7EFE2A9FFFE5FFD7FEFF0A2CFDE8DD2C" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[341,573,520,546]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="polotowae" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFE5FFD7FEFF0A2CFDE8DD2C" box="[341,573,520,546]" italics="true" pageId="29" pageNumber="30">Tabiboka polotowae</emphasis>
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<taxonomicNameLabel id="90B93075FFE5FFD7FDE90A2DFD39DD2D" box="[579,748,521,547]" pageId="29" pageNumber="30" rank="species">gen. et sp. nov.</taxonomicNameLabel>
may be distinguished by an elongate embolus (E) arising retrobasally near 200°, the paraconductor (Pc) visible between embolus and the conductor (C) base, the palpal tibia having blunt retromedian process (RTA) arising near basally and narrow, finger-like ventral apical process (VTA) (
<figureCitation id="21C54D99FFE5FFD7FE640A53FDB6DD9F" box="[462,611,631,657]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="29" pageNumber="30">Figs 11A–C</figureCitation>
,
<figureCitation id="21C54D99FFE5FFD7FDDB0A53FD10DD9F" box="[625,709,631,657]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; A–F = male holotype (CASENT_9016117; CAS); G–H= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–D = 0.5 mm; E–F = 0.15 mm; G–H = 0.1 mm. SEM images A–F by Charles Griswold and Darrell Ubick, G–H by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="29" pageNumber="30">12A–F</figureCitation>
) (vs in
<taxonomicName id="7EFE2A9FFFE5FFD7FC8F0A5CFC46DD9F" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[805,915,631,657]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="milleri" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFE5FFD7FC8F0A5CFC46DD9F" box="[805,915,631,657]" italics="true" pageId="29" pageNumber="30">T. milleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFE5FFD7FC300A53FB81DD9F" box="[922,1108,631,657]" pageId="29" pageNumber="30" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, males have shorter and broader E and MA, a small VTA and greatly elongated RTA,
<figureCitation id="21C54D99FFE5FFD7FCD50ABFFC28DDB8" box="[895,1021,667,694]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="29" pageNumber="30">Fig. 5A–C</figureCitation>
; and in
<taxonomicName id="7EFE2A9FFFE5FFD7FBF90AB8FB69DDBB" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1107,1212,667,693]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="milloti" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFE5FFD7FBF90AB8FB69DDBB" box="[1107,1212,667,693]" italics="true" pageId="29" pageNumber="30">T. milloti</emphasis>
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<taxonomicNameLabel id="90B93075FFE5FFD7FB690AB8FAA7DDB8" box="[1219,1394,668,694]" pageId="29" pageNumber="30" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, the MA is elongate, cruciform, and the VTA is thumb-like,
<figureCitation id="21C54D99FFE5FFD7FCD60AE4FC28DDD4" box="[892,1021,704,730]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.A–C = male holotype (CASENT_9016063; MNHN). D–E = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="29" pageNumber="30">Fig. 9A–C</figureCitation>
). Females of
<taxonomicName id="7EFE2A9FFFE5FFD7FB0A0AE5FAE2DDD4" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1184,1335,704,730]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="polotowae">
<emphasis id="8B8A8D0EFFE5FFD7FB0A0AE5FAE2DDD4" box="[1184,1335,704,730]" italics="true" pageId="29" pageNumber="30">T. polotowae</emphasis>
</taxonomicName>
have epigyne median lobe (ML) rectangular, narrowest in middle or with sides straight (
<figureCitation id="21C54D99FFE5FFD7FB380AC1FAD7DDF1" box="[1170,1282,741,767]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="29" pageNumber="30">Figs 11D</figureCitation>
,
<figureCitation id="21C54D99FFE5FFD7FAA50AC1FA96DDF1" box="[1295,1347,741,767]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; A–F = male holotype (CASENT_9016117; CAS); G–H= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–D = 0.5 mm; E–F = 0.15 mm; G–H = 0.1 mm. SEM images A–F by Charles Griswold and Darrell Ubick, G–H by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="29" pageNumber="30">12G</figureCitation>
) (vs heart-shaped ML in
<taxonomicName id="7EFE2A9FFFE5FFD7FE010B2FFDC0DC2A" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[427,533,778,804]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="8B8A8D0EFFE5FFD7FE010B2FFDC0DC2A" box="[427,533,778,804]" italics="true" pageId="29" pageNumber="30">T. milleri</emphasis>
</taxonomicName>
,
<figureCitation id="21C54D99FFE5FFD7FD8E0B2EFD57DC2A" box="[548,642,778,804]" captionStart="Fig" captionStartId="17.[189,231,1892,1918]" captionTargetBox="[189,1398,962,1853]" captionTargetId="figure-276@17.[189,1398,962,1853]" captionTargetPageId="17" captionText="Fig. 4. Tabiboka milleri gen. et sp. nov.A. Female paratype (CASENT_9029871; CAS) from Ranomafana (Photo © N. Scharff). B. Female paratype (CASENT_9016098; CAS), epigyne, ventral view. C. Female paratype (CASENT_9016094; CAS), trochanter 1, ventral view. D. Idem, trochanter 4. (No scales provided). Stacked microphotographs by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001796" httpUri="https://zenodo.org/record/14001796/files/figure.png" pageId="29" pageNumber="30">Figs 4B</figureCitation>
,
<figureCitation id="21C54D99FFE5FFD7FD250B2EFD60DC2B" box="[655,693,778,805]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="29" pageNumber="30">5D</figureCitation>
,
<figureCitation id="21C54D99FFE5FFD7FD680B2EFCFADC2A" box="[706,815,778,804]" captionStart="Fig" captionStartId="20.[189,232,1822,1848]" captionTargetBox="[189,1398,461,1784]" captionTargetId="figure-79@20.[189,1398,461,1784]" captionTargetPageId="20" captionText="Fig. 7. Tabiboka milleri gen. nov. et sp. nov. SEM of female genitalia, A–B, E–F = female paratype (CASENT_9016094; CAS); C–D = female paratype (CASENT_9035866; CAS). A. Epigyne, ventral view. B. Idem, ventro-posterior view. C. Endogyne, dorsal view. D. Idem, slightly anterior view. E. Spermatheca, right dorsal view. F. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: A–E = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila." figureDoi="http://doi.org/10.5281/zenodo.14001806" httpUri="https://zenodo.org/record/14001806/files/figure.png" pageId="29" pageNumber="30">
7A and
<taxonomicName id="7EFE2A9FFFE5FFD7FCB70B2FFCFADC2A" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[797,815,779,804]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="milloti">
<emphasis id="8B8A8D0EFFE5FFD7FCB70B2FFCFADC2A" box="[797,815,779,804]" italics="true" pageId="29" pageNumber="30">T</emphasis>
</taxonomicName>
</figureCitation>
. milloti,
<figureCitation id="21C54D99FFE5FFD7FC3F0B2EFC27DC2A" box="[917,1010,778,804]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.A–C = male holotype (CASENT_9016063; MNHN). D–E = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="29" pageNumber="30">Fig. 9D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFE5FFD7FF170B69FBDFDB08" pageId="29" pageNumber="30" type="etymology">
<paragraph id="B941511CFFE5FFD7FF170B69FE91DC69" blockId="29.[189,324,845,871]" box="[189,324,845,871]" pageId="29" pageNumber="30">
<heading id="E209E670FFE5FFD7FF170B69FE91DC69" bold="true" box="[189,324,845,871]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="8B8A8D0EFFE5FFD7FF170B69FE91DC69" bold="true" box="[189,324,845,871]" pageId="29" pageNumber="30">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE5FFD7FF170B59FBDFDB08" blockId="29.[189,1400,893,1030]" pageId="29" pageNumber="30">
The species name is a matronym in honor of Dr Daniele Polotow, who studies the phylogenetics and taxonomy of spiders of the OC clade including Lycosoidea. She led the research that in 2015 led to the proposal and diagnosis of the
<taxonomicName id="7EFE2A9FFFE5FFD7FDF30BE2FD18DCEE" authorityName="Griswold &amp; Polotow" authorityYear="2015" box="[601,717,966,992]" class="Arachnida" family="Udubidae" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="family">Udubidae</taxonomicName>
and the current phylogenetic framework for Lycosoidea interrelationships, and she collected many of the data used in this paper.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFE5FFD7FF170C0AFB57D8E6" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="B941511CFFE5FFD7FF170C0AFEB9DB46" blockId="29.[189,364,1070,1096]" box="[189,364,1070,1096]" pageId="29" pageNumber="30">
<heading id="E209E670FFE5FFD7FF170C0AFEB9DB46" bold="true" box="[189,364,1070,1096]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="8B8A8D0EFFE5FFD7FF170C0AFEB9DB46" bold="true" box="[189,364,1070,1096]" pageId="29" pageNumber="30">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE5FFD7FF400C7AFE89DB76" blockId="29.[189,1399,1118,1254]" box="[234,348,1118,1144]" pageId="29" pageNumber="30">
<emphasis id="8B8A8D0EFFE5FFD7FF400C7AFE89DB76" bold="true" box="[234,348,1118,1144]" pageId="29" pageNumber="30">
<typeStatus id="6645EFBEFFE5FFD7FF400C7AFE89DB76" box="[234,348,1118,1144]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFE5FFD7FF170CA7FC80DBE8" blockId="29.[189,1399,1118,1254]" pageId="29" pageNumber="30">
<materialsCitation id="09965B41FFE5FFD7FF170CA7FC80DBE8" collectingDate="1992-07-27" collectionCode="CAS" collectorName="V. and B. Roth" country="Madagascar" latitude="-18.733334" location="Ambohimanga Village" longLatPrecision="1274" longitude="47.566666" pageId="29" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo" typeStatus="holotype">
<collectingCountry id="C1E9118CFFE5FFD7FF170CA7FE5BDB93" box="[189,398,1155,1181]" name="Madagascar" pageId="29" pageNumber="30">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFE5FFD7FE0D0CA7FD1CDB93" bold="true" box="[423,713,1155,1181]" pageId="29" pageNumber="30">
<collectingRegion id="7B3A9FFEFFE5FFD7FE0D0CA7FD1CDB93" box="[423,713,1155,1181]" country="Madagascar" name="Antananarivo" pageId="29" pageNumber="30">Antananarivo Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFE5FFD7FD4A0CA7FD2CDB93" box="[736,761,1155,1181]" count="1" pageId="29" pageNumber="30" type="male">♂</specimenCount>
; NE outskirts of
<collectingRegion id="7B3A9FFEFFE5FFD7FC130CA7FB89DB93" box="[953,1116,1155,1181]" country="Madagascar" name="Antananarivo" pageId="29" pageNumber="30">Antananarivo</collectingRegion>
,
<location id="BC2107C7FFE5FFD7FBCF0CA7FAA5DB93" LSID="urn:lsid:plazi:treatment:3157E00AFFE5FFE9FDEC092CFEAEDD1E:BC2107C7FFE5FFD7FBCF0CA7FAA5DB93" box="[1125,1392,1155,1181]" country="Madagascar" latitude="-18.733334" longLatPrecision="1274" longitude="47.566666" name="Ambohimanga Village" pageId="29" pageNumber="30" stateProvince="Antananarivo">Ambohimanga Village</location>
;
<geoCoordinate id="DCCA37DBFFE5FFD7FF170C82FEF1DBCC" box="[189,292,1190,1218]" degrees="18" direction="south" minutes="44" orientation="latitude" pageId="29" pageNumber="30" precision="925" value="-18.733334">18°44′ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFE5FFD7FE980C82FE4CDBCC" box="[306,409,1190,1218]" degrees="47" direction="east" minutes="34" orientation="longitude" pageId="29" pageNumber="30" precision="925" value="47.566666">47°34′ E</geoCoordinate>
(coordinates approximate);
<date id="CD4077DCFFE5FFD7FD4D0C8CFCA8DBCC" box="[743,893,1191,1218]" pageId="29" pageNumber="30" value="1992-07-27">
<collectingDate id="DD048E34FFE5FFD7FD4D0C8CFCA8DBCC" box="[743,893,1191,1218]" pageId="29" pageNumber="30" value="1992-07-27">27 Jul. 1992</collectingDate>
</date>
;
<collectorName id="140B34CAFFE5FFD7FC230C8CFBECDBCF" box="[905,1081,1191,1218]" pageId="29" pageNumber="30">V. and B. Roth</collectorName>
leg.; VBRoth002; [palp &amp; endogyne drawn; SEM palp]; CASENT_9016117;
<collectionCode id="DFEFC9D9FFE5FFD7FCBE0CE8FC84DBE8" box="[788,849,1228,1254]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B941511CFFE5FFD7FF400D2BFEB2DA27" blockId="29.[187,1399,1295,1689]" box="[234,359,1295,1321]" pageId="29" pageNumber="30">
<emphasis id="8B8A8D0EFFE5FFD7FF400D2BFEB2DA27" bold="true" box="[234,359,1295,1321]" pageId="29" pageNumber="30">
<typeStatus id="6645EFBEFFE5FFD7FF400D2BFEB2DA27" box="[234,359,1295,1321]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFE5FFD7FF170D10FCD5D997" blockId="29.[187,1399,1295,1689]" pageId="29" pageNumber="30">
<materialsCitation id="09965B41FFE5FFD7FF170D10FF23DA7D" collectingDate="1992-07-27" collectionCode="CAS" collectorName="V. and B. Roth" country="Madagascar" latitude="-18.733334" location="Ambohimanga Village" longLatPrecision="1274" longitude="47.566666" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo" typeStatus="paratype">
<collectingCountry id="C1E9118CFFE5FFD7FF170D10FE44DA40" box="[189,401,1332,1358]" name="Madagascar" pageId="29" pageNumber="30">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFE5FFD7FE1E0D10FD0ADA40" bold="true" box="[436,735,1332,1358]" pageId="29" pageNumber="30">
<collectingRegion id="7B3A9FFEFFE5FFD7FE1E0D10FD0ADA40" box="[436,735,1332,1358]" country="Madagascar" name="Antananarivo" pageId="29" pageNumber="30">Antananarivo Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFE5FFD7FD560D10FCFCDA41" box="[764,809,1332,1359]" count="1" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
; same data as for holotype; CASENT_9016117;
<collectionCode id="DFEFC9D9FFE5FFD7FF170D7DFF23DA7D" box="[189,246,1369,1395]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFE5FFD7FEA70D7DFB60DA96" collectingDate="1993-11-01" collectionCode="CAS" collectorName="J. Coddington &amp; N. Scharff &amp; S. Larcher &amp; C. Griswold &amp; R. Andriamasimanana &amp; Berlese" country="Madagascar" latitude="-18.733334" location="Ambohimanga Village" longLatPrecision="1274" longitude="47.566666" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo" typeStatus="paratype">
<specimenCount id="AFF89A95FFE5FFD7FEA70D7DFEECDA7D" box="[269,313,1369,1395]" count="1" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
; same data as for preceding;
<date id="CD4077DCFFE5FFD7FD230D7DFCCFDA7D" box="[649,794,1369,1395]" pageId="29" pageNumber="30" value="1993-11-01">
<collectingDate id="DD048E34FFE5FFD7FD230D7DFCCFDA7D" box="[649,794,1369,1395]" pageId="29" pageNumber="30" value="1993-11-01">1 Nov. 1993</collectingDate>
</date>
;
<collectorName id="140B34CAFFE5FFD7FC8F0D7DFC19DA7D" box="[805,972,1369,1395]" pageId="29" pageNumber="30">J. Coddington</collectorName>
,
<collectorName id="140B34CAFFE5FFD7FC7C0D7DFB84DA7D" box="[982,1105,1369,1395]" pageId="29" pageNumber="30">N. Scharff</collectorName>
,
<collectorName id="140B34CAFFE5FFD7FBF10D7DFB03DA7D" box="[1115,1238,1369,1395]" pageId="29" pageNumber="30">S. Larcher</collectorName>
,
<collectorName id="140B34CAFFE5FFD7FB4B0D7DFAA7DA7D" box="[1249,1394,1369,1395]" pageId="29" pageNumber="30">C. Griswold</collectorName>
,
<collectorName id="140B34CAFFE5FFD7FF170D5AFE68DA99" box="[189,445,1405,1432]" pageId="29" pageNumber="30">R. Andriamasimanana</collectorName>
leg.;
<collectorName id="140B34CAFFE5FFD7FE500D59FD84DA99" box="[506,593,1405,1431]" pageId="29" pageNumber="30">Berlese</collectorName>
of fine leaf litter; CEG010; CASENT_9016116;
<collectionCode id="DFEFC9D9FFE5FFD7FBD70D5AFB60DA96" box="[1149,1205,1406,1432]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFE5FFD7FB610D5AFDB6DAEF" collectingDate="1992-08-05" collectionCode="CAS" collectorName="V. Roth &amp; Roth" country="Madagascar" elevation="1300" latitude="-18.758888" location="10 km NE of Tana" longLatPrecision="21" longitude="48.5625" pageId="29" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo" typeStatus="paratype">
<specimenCount id="AFF89A95FFE5FFD7FB610D5AFB22DA96" box="[1227,1271,1406,1432]" count="1" pageId="29" pageNumber="30" type="male">1 ♂</specimenCount>
;
<location id="BC2107C7FFE5FFD7FAA80D5AFEC2DAB3" LSID="urn:lsid:plazi:treatment:3157E00AFFE5FFE9FDEC092CFEAEDD1E:BC2107C7FFE5FFD7FAA80D5AFEC2DAB3" country="Madagascar" latitude="-18.758888" longLatPrecision="21" longitude="48.5625" name="10 km NE of Tana" pageId="29" pageNumber="30" stateProvince="Antananarivo">
<quantity id="7E06FCF9FFE5FFD7FAA80D5AFA9FDA99" box="[1282,1354,1405,1432]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="30" unit="km" value="10.0">10 km</quantity>
NE of Tana
</location>
;
<geoCoordinate id="DCCA37DBFFE5FFD7FE880D85FE64DAB2" box="[290,433,1441,1469]" degrees="18" direction="south" minutes="45" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="32" value="-18.758888">18°45′32″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFE5FFD7FE170D85FD9ADAB3" box="[445,591,1441,1469]" degrees="48" direction="east" minutes="33" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="45" value="48.5625">48°33′45″ E</geoCoordinate>
;
<elevation id="32D3B62FFFE5FFD7FDF00D86FD24DAB2" box="[602,753,1442,1469]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="30" unit="m" value="1300.0">
<quantity id="7E06FCF9FFE5FFD7FDF00D86FD67DAB3" box="[602,690,1442,1469]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="30" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFE5FFD7FD570D86FC7CDAB2" box="[765,937,1442,1469]" pageId="29" pageNumber="30" value="1992-08-05">
<collectingDate id="DD048E34FFE5FFD7FD570D86FC7CDAB2" box="[765,937,1442,1469]" pageId="29" pageNumber="30" value="1992-08-05">5 August 1992</collectingDate>
</date>
;
<collectorName id="140B34CAFFE5FFD7FC1E0D87FBDADAB2" box="[948,1039,1442,1469]" pageId="29" pageNumber="30">V. Roth</collectorName>
, leg.; [habitus photos of male];
<collectorName id="140B34CAFFE5FFD7FF170DE3FF2FDAEF" box="[189,250,1479,1505]" pageId="29" pageNumber="30">Roth</collectorName>
001; CASENT_9016118;
<collectionCode id="DFEFC9D9FFE5FFD7FD8C0DE3FDB6DAEF" box="[550,611,1479,1505]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<emphasis id="8B8A8D0EFFE5FFD7FDC70DE3FDA9DAEF" bold="true" box="[621,636,1479,1505]" pageId="29" pageNumber="30">–</emphasis>
<materialsCitation id="09965B41FFE5FFD7FD280DE3FAF3D941" collectingDate="2009-01-07" collectionCode="CAS" collectorName="H. Wood &amp; Voucher &amp; Zorodictynidae &amp; D. Polotow Lycosoidea Total Evidence Phylogeny" country="Madagascar" elevation="1580" latitude="-22.146917" location="Parc National Andringitra" longLatPrecision="1" longitude="45.950947" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<emphasis id="8B8A8D0EFFE5FFD7FD280DE3FC48DAEF" bold="true" box="[642,925,1479,1505]" pageId="29" pageNumber="30">
<collectingRegion id="7B3A9FFEFFE5FFD7FD280DE3FC48DAEF" box="[642,925,1479,1505]" country="Madagascar" name="Fianarantsoa" pageId="29" pageNumber="30">Fianarantsoa Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFE5FFD7FC1C0DE3FC36DAEC" box="[950,995,1479,1506]" count="1" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
;
<location id="BC2107C7FFE5FFD7FC5B0DECFAF7DAEF" LSID="urn:lsid:plazi:treatment:3157E00AFFE5FFE9FDEC092CFEAEDD1E:BC2107C7FFE5FFD7FC5B0DECFAF7DAEF" box="[1009,1314,1479,1506]" country="Madagascar" latitude="-22.146917" longLatPrecision="1" longitude="45.950947" name="Parc National Andringitra" pageId="29" pageNumber="30" stateProvince="Fianarantsoa">Parc National Andringitra</location>
,
<locationDeviation id="A8F3483EFFE5FFD7FA870DE3FEA8D908" location="Ambalavao" pageId="29" pageNumber="30">
<quantity id="7E06FCF9FFE5FFD7FA870DE3FAA3DAEF" box="[1325,1398,1479,1505]" metricMagnitude="4" metricUnit="m" metricValue="3.4" pageId="29" pageNumber="30" unit="km" value="34.0">34 km</quantity>
S of Ambalavao
</locationDeviation>
;
<geoCoordinate id="DCCA37DBFFE5FFD7FE220DCEFDF8D908" box="[392,557,1514,1542]" degrees="22" direction="south" minutes="08" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="48.9" value="-22.146917">22°08′48.9″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFE5FFD7FD900DCEFD37D908" box="[570,738,1514,1542]" degrees="45" direction="east" minutes="57" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="03.4" value="45.950947">45°57′03.4″ E</geoCoordinate>
;
<elevation id="32D3B62FFFE5FFD7FD440DC8FC50D908" box="[750,901,1516,1543]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="29" pageNumber="30" unit="m" value="1580.0">
<quantity id="7E06FCF9FFE5FFD7FD440DC8FC93D908" box="[750,838,1516,1543]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="29" pageNumber="30" unit="m" value="1580.0">1580 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFE5FFD7FC380DC8FBCED908" box="[914,1051,1516,1542]" pageId="29" pageNumber="30" value="2009-01-07">
<collectingDate id="DD048E34FFE5FFD7FC380DC8FBCED908" box="[914,1051,1516,1542]" pageId="29" pageNumber="30" value="2009-01-07">7 Jan. 2009</collectingDate>
</date>
;
<collectorName id="140B34CAFFE5FFD7FB8D0DC8FB5BD908" box="[1063,1166,1516,1542]" pageId="29" pageNumber="30">H. Wood</collectorName>
leg.; sifting litter in the day; primary montane rainforest; [SEM stubs 221, 9, 10, 222, 4] [
<collectorName id="140B34CAFFE5FFD7FC600E34FBF8D924" box="[970,1069,1552,1578]" pageId="29" pageNumber="30">Voucher</collectorName>
DP-2014b -“
<collectorName id="140B34CAFFE5FFD7FB6B0E34FAA2D924" box="[1217,1399,1552,1578]" pageId="29" pageNumber="30">Zorodictynidae</collectorName>
sp. 3”,
<collectorName id="140B34CAFFE5FFD7FEA50E12FCB2D941" box="[271,871,1589,1616]" pageId="29" pageNumber="30">D. Polotow Lycosoidea Total Evidence Phylogeny</collectorName>
]; HW0813; CASENT_9029890;
<collectionCode id="DFEFC9D9FFE5FFD7FB470E11FAF3D941" box="[1261,1318,1589,1615]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFE5FFD7FAEA0E11FD28D997" collectingDate="2009-01-07" collectionCode="CAS" collectorName="Zorodictynidae &amp; D. Polotow Lycosoidea Total Evidence Phylogeny" country="Madagascar" elevation="1580" latitude="-22.146917" location="Voucher" longLatPrecision="1" longitude="45.950947" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="AFF89A95FFE5FFD7FAEA0E11FABAD95E" box="[1344,1391,1589,1616]" count="1" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
; same data as for preceding; [
<location id="BC2107C7FFE5FFD7FDA40E7EFDA6D97A" LSID="urn:lsid:plazi:treatment:3157E00AFFE5FFE9FDEC092CFEAEDD1E:BC2107C7FFE5FFD7FDA40E7EFDA6D97A" box="[526,627,1626,1652]" country="Madagascar" name="Voucher" pageId="29" pageNumber="30" stateProvince="Fianarantsoa">Voucher</location>
DP-2014a -“
<collectorName id="140B34CAFFE5FFD7FCA10E7EFC16D97A" box="[779,963,1626,1652]" pageId="29" pageNumber="30">Zorodictynidae</collectorName>
sp. 4”,
<collectorName id="140B34CAFFE5FFD7FB8A0E7FFE63D997" pageId="29" pageNumber="30">D. Polotow Lycosoidea Total Evidence Phylogeny</collectorName>
]; CASENT_9029889;
<collectionCode id="DFEFC9D9FFE5FFD7FD6A0E5BFD28D997" box="[704,765,1663,1689]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="B941511CFFE5FFD7FF170EE5FE2CD9D5" blockId="29.[189,505,1729,1755]" box="[189,505,1729,1755]" pageId="29" pageNumber="30">
<heading id="E209E670FFE5FFD7FF170EE5FE2CD9D5" bold="true" box="[189,505,1729,1755]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="8B8A8D0EFFE5FFD7FF170EE5FE2CD9D5" bold="true" box="[189,505,1729,1755]" pageId="29" pageNumber="30">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE5FFD7FF170ED6FB57D8E6" blockId="29.[187,1399,1778,2025]" pageId="29" pageNumber="30">
<materialsCitation id="09965B41FFE5FFD7FF170ED6FF21D874" collectingDate="2001-09-08" collectionCode="CAS" collectorName="D. Ubick &amp; D. Andriamalala &amp; T. Andriambinintsoa &amp; J. J. Rafanomezantsoa" country="Madagascar" elevation="1300" latitude="-18.933332" location="Station Forestiere Angavokely" longLatPrecision="1273" longitude="47.75" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Antananarivo">
<collectingCountry id="C1E9118CFFE5FFD7FF170ED6FE46D802" box="[189,403,1778,1804]" name="Madagascar" pageId="29" pageNumber="30">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFE5FFD7FE100ED6FD38D802" bold="true" box="[442,749,1778,1804]" pageId="29" pageNumber="30">
<collectingRegion id="7B3A9FFEFFE5FFD7FE100ED6FD38D802" box="[442,749,1778,1804]" country="Madagascar" name="Antananarivo" pageId="29" pageNumber="30">Antananarivo Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFE5FFD7FCBA0ED6FC94D802" box="[784,833,1778,1804]" count="1" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
;
<location id="BC2107C7FFE5FFD7FCFF0ED6FB07D802" LSID="urn:lsid:plazi:treatment:3157E00AFFE5FFE9FDEC092CFEAEDD1E:BC2107C7FFE5FFD7FCFF0ED6FB07D802" box="[853,1234,1778,1804]" country="Madagascar" latitude="-18.933332" longLatPrecision="1273" longitude="47.75" name="Station Forestiere Angavokely" pageId="29" pageNumber="30" stateProvince="Antananarivo">Station Forestière Angavokely</location>
,
<locationDeviation id="A8F3483EFFE5FFD7FB480ED6FE88D83E" location="Antananarivo" pageId="29" pageNumber="30">
<quantity id="7E06FCF9FFE5FFD7FB480ED6FAE6D802" box="[1250,1331,1778,1804]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="km" value="22.0">22 km</quantity>
E of
<collectingRegion id="7B3A9FFEFFE5FFD7FF170F32FE88D83E" box="[189,349,1814,1840]" country="Madagascar" name="Antananarivo" pageId="29" pageNumber="30">Antananarivo</collectingRegion>
</locationDeviation>
;
<geoCoordinate id="DCCA37DBFFE5FFD7FECC0F31FE1DD83F" box="[358,456,1813,1841]" degrees="18" direction="south" minutes="56" orientation="latitude" pageId="29" pageNumber="30" precision="925" value="-18.933332">18°56′ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFE5FFD7FE780F31FDE2D83F" box="[466,567,1813,1841]" degrees="47" direction="east" minutes="45" orientation="longitude" pageId="29" pageNumber="30" precision="925" value="47.75">47°45′ E</geoCoordinate>
;
<elevation id="32D3B62FFFE5FFD7FDEA0F33FD04D83E" box="[576,721,1814,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="30" unit="m" value="1300.0">
<quantity id="7E06FCF9FFE5FFD7FDEA0F33FD43D83F" box="[576,662,1815,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="30" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</elevation>
; low canopy remnant forest, general collecting;
<date id="CD4077DCFFE5FFD7FB400F33FAA7D83F" box="[1258,1394,1815,1841]" pageId="29" pageNumber="30" value="2001-09-08">
<collectingDate id="DD048E34FFE5FFD7FB400F33FAA7D83F" box="[1258,1394,1815,1841]" pageId="29" pageNumber="30" value="2001-09-08">8 Sep. 2001</collectingDate>
</date>
;
<collectorName id="140B34CAFFE5FFD7FF170F18FEF2D85B" box="[189,295,1851,1878]" pageId="29" pageNumber="30">D. Ubick</collectorName>
,
<collectorName id="140B34CAFFE5FFD7FE9B0F18FE25D85B" box="[305,496,1851,1878]" pageId="29" pageNumber="30">D. Andriamalala</collectorName>
,
<collectorName id="140B34CAFFE5FFD7FE500F18FD32D85B" box="[506,743,1851,1878]" pageId="29" pageNumber="30">T.Andriambinintsoa</collectorName>
,
<collectorName id="140B34CAFFE5FFD7FD5B0F18FC32D85B" box="[753,999,1851,1878]" pageId="29" pageNumber="30">J.J. Rafanomezantsoa</collectorName>
leg.; DU-01-2; CASENT_901371;
<collectionCode id="DFEFC9D9FFE5FFD7FF110F44FF21D874" box="[187,244,1888,1914]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFE5FFD7FEA50F44FC23D874" box="[271,1014,1888,1915]" collectingDate="2001-09-08" collectionCode="CAS" collectorName="D. Ubick &amp; D. Andriamalala &amp; T. Andriambinintsoa &amp; J. J. Rafanomezantsoa" country="Madagascar" elevation="1300" latitude="-18.933332" location="Station Forestiere Angavokely" longLatPrecision="1273" longitude="47.75" pageId="29" pageNumber="30" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount id="AFF89A95FFE5FFD7FEA50F44FEEAD875" box="[271,319,1888,1915]" count="1" pageId="29" pageNumber="30" type="male">1 ♂</specimenCount>
,
<specimenCount id="AFF89A95FFE5FFD7FEE60F44FEAFD875" box="[332,378,1888,1915]" count="1" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
; same data as for preceding; CASENT_901317;
<collectionCode id="DFEFC9D9FFE5FFD7FC170F44FC23D874" box="[957,1014,1888,1914]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFE5FFD7FBBA0F44FDDDD8CA" collectingDate="1993-11-02" collectionCode="USNM" collectorName="C. Griswold &amp; S. Larcher &amp; N. Scharff" country="Madagascar" elevation="1400" latitude="-18.733334" location="Ambohimanga" longLatPrecision="1274" longitude="47.566666" pageId="29" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount id="AFF89A95FFE5FFD7FBBA0F44FB95D875" box="[1040,1088,1888,1915]" count="1" pageId="29" pageNumber="30" type="male">1 ♂</specimenCount>
;
<location id="BC2107C7FFE5FFD7FBE60F44FB2BD874" LSID="urn:lsid:plazi:treatment:3157E00AFFE5FFE9FDEC092CFEAEDD1E:BC2107C7FFE5FFD7FBE60F44FB2BD874" box="[1100,1278,1888,1914]" country="Madagascar" latitude="-18.733334" longLatPrecision="1274" longitude="47.566666" name="Ambohimanga" pageId="29" pageNumber="30" stateProvince="Antananarivo">Ambohimanga</location>
;
<geoCoordinate id="DCCA37DBFFE5FFD7FAA00F7AFAA4D874" box="[1290,1393,1886,1914]" degrees="18" direction="south" minutes="44" orientation="latitude" pageId="29" pageNumber="30" precision="925" value="-18.733334">18°44′ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFE5FFD7FF170FA7FEF0D891" box="[189,293,1923,1951]" degrees="47" direction="east" minutes="34" orientation="longitude" pageId="29" pageNumber="30" precision="925" value="47.566666">47°34′ E</geoCoordinate>
;
<elevation id="32D3B62FFFE5FFD7FE9A0FA1FE13D891" box="[304,454,1925,1951]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="30" unit="m" value="1400.0">
<quantity id="7E06FCF9FFE5FFD7FE9A0FA1FE52D891" box="[304,391,1925,1951]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="30" unit="m" value="1400.0">1400 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFE5FFD7FE780FA1FDB7D891" box="[466,610,1925,1951]" pageId="29" pageNumber="30" value="1993-11-02">
<collectingDate id="DD048E34FFE5FFD7FE780FA1FDB7D891" box="[466,610,1925,1951]" pageId="29" pageNumber="30" value="1993-11-02">2 Nov. 1993</collectingDate>
</date>
;
<collectorName id="140B34CAFFE5FFD7FDC70FA1FD2BD891" box="[621,766,1925,1951]" pageId="29" pageNumber="30">C. Griswold</collectorName>
,
<collectorName id="140B34CAFFE5FFD7FCA20FA1FC56D891" box="[776,899,1925,1951]" pageId="29" pageNumber="30">S. Larcher</collectorName>
,
<collectorName id="140B34CAFFE5FFD7FC270FA1FBD0D891" box="[909,1029,1925,1951]" pageId="29" pageNumber="30">N. Scharff</collectorName>
leg.; night collecting; CEG009; CASENT_9016115;
<collectionCode id="DFEFC9D9FFE5FFD7FE190F8EFDDDD8CA" box="[435,520,1962,1988]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="30" type="Museum">USNM</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFE5FFD7FD8B0F8EFBABD8E6" collectingDate="1992-08-05" collectionCode="CAS" collectorName="V. Roth &amp; Roth" country="Madagascar" elevation="1300" latitude="-18.758888" location="10 km NE of Tana" longLatPrecision="21" longitude="48.5625" pageId="29" pageNumber="30" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount id="AFF89A95FFE5FFD7FD8B0F8EFD85D8CA" box="[545,592,1962,1988]" count="1" pageId="29" pageNumber="30" type="male">1 ♂</specimenCount>
;
<location id="BC2107C7FFE5FFD7FDF70F8EFCECD8CA" LSID="urn:lsid:plazi:treatment:3157E00AFFE5FFE9FDEC092CFEAEDD1E:BC2107C7FFE5FFD7FDF70F8EFCECD8CA" box="[605,825,1961,1988]" country="Madagascar" latitude="-18.758888" longLatPrecision="21" longitude="48.5625" name="10 km NE of Tana" pageId="29" pageNumber="30" stateProvince="Antananarivo">
<quantity id="7E06FCF9FFE5FFD7FDF70F8EFD72D8CD" box="[605,679,1961,1988]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="30" unit="km" value="10.0">10 km</quantity>
NE of Tana
</location>
;
<geoCoordinate id="DCCA37DBFFE5FFD7FCEF0F8CFC00D8CA" box="[837,981,1960,1988]" degrees="18" direction="south" minutes="45" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="32" value="-18.758888">18°45′32″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFE5FFD7FC480F8CFBA0D8CA" box="[994,1141,1960,1988]" degrees="48" direction="east" minutes="33" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="45" value="48.5625">48°33′45″ E</geoCoordinate>
;
<elevation id="32D3B62FFFE5FFD7FB2B0F8EFACFD8CD" box="[1153,1306,1961,1988]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="30" unit="m" value="1300.0">
<quantity id="7E06FCF9FFE5FFD7FB2B0F8EFB0FD8CA" box="[1153,1242,1962,1988]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="30" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFE5FFD7FA820F8DFF2ED8E6" pageId="29" pageNumber="30" value="1992-08-05">
<collectingDate id="DD048E34FFE5FFD7FA820F8DFF2ED8E6" pageId="29" pageNumber="30" value="1992-08-05">5 Aug. 1992</collectingDate>
</date>
;
<collectorName id="140B34CAFFE5FFD7FEAD0FEBFEB6D8E6" box="[263,355,1998,2025]" pageId="29" pageNumber="30">V. Roth</collectorName>
, leg.; [habitus photos of male];
<collectorName id="140B34CAFFE5FFD7FD7D0FEAFCC1D8E6" box="[727,788,1998,2024]" pageId="29" pageNumber="30">Roth</collectorName>
001; CASENT_9016118;
<collectionCode id="DFEFC9D9FFE5FFD7FBEB0FEAFBABD8E6" box="[1089,1150,1998,2024]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFE6FFE9FF17092CFBE3DE79" lastPageId="35" lastPageNumber="36" pageId="30" pageNumber="31" type="description">
<paragraph id="B941511CFFE6FFD4FF17092CFE9BDE2C" blockId="30.[189,334,264,290]" box="[189,334,264,290]" pageId="30" pageNumber="31">
<heading id="E209E670FFE6FFD4FF17092CFE9BDE2C" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="8B8A8D0EFFE6FFD4FF17092CFE9BDE2C" bold="true" box="[189,334,264,290]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE6FFD4FF400913FD33DE5F" blockId="30.[189,1160,311,373]" box="[234,742,311,337]" pageId="30" pageNumber="31">
<emphasis id="8B8A8D0EFFE6FFD4FF400913FE48DE5F" bold="true" box="[234,413,311,337]" pageId="30" pageNumber="31">
Male
<typeStatus id="6645EFBEFFE6FFD4FE9B0913FE48DE5F" box="[305,413,311,337]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
</emphasis>
(CASENT_9016117; CAS)
</paragraph>
<paragraph id="B941511CFFE6FFD4FF17097EFB5DDE7A" blockId="30.[189,1160,311,373]" box="[189,1160,346,373]" pageId="30" pageNumber="31">MEASUREMENTS. Total length: 8.10. Carapace: length 4.50, width 3.40, height 1.90.</paragraph>
<paragraph id="B941511CFFE6FFD5FF1709B8FC71DE82" blockId="30.[188,1399,412,473]" lastBlockId="31.[189,1400,265,397]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
COLORATION (
<figureCitation id="21C54D99FFE6FFD4FECB09B8FE38DEB8" box="[353,493,412,438]" captionStart="Fig" captionStartId="30.[191,235,1928,1954]" captionTargetBox="[189,1398,529,1888]" captionTargetId="figure-67@30.[189,1398,529,1888]" captionTargetPageId="30" captionText="Fig. 10. Tabiboka polotowae gen. et sp. nov., male and female habitus; A, C = male paratype (CASENT_9016118; CAS); B, D = female paratype (CASENT_9016116; CAS). A–B. Dorsal view. C–D. Ventral view. Scale bars = 2.5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001815" httpUri="https://zenodo.org/record/14001815/files/figure.png" pageId="30" pageNumber="31">Fig. 10A, C</figureCitation>
). Carapace brownish yellow, darker along margin, with two faint, longitudinal dark bands along margin of pars cephalica; black on ocular area and surrounding all eyes; chelicerae brownish yellow; sternum yellow, unmarked; legs brownish yellow, with subbasal and subapical dusky annuli on all femora, other segments unmarked; abdomen brownish yellow with darker cardiac mark and dorsolateral dark bands that break up into chevrons posteriorly, sides brownish yellow mottled with dark marks, venter yellow except darker around spinneret base.
</paragraph>
<caption id="ED810194FFE6FFD4FF150FACFAE6D8E6" ID-DOI="http://doi.org/10.5281/zenodo.14001815" ID-Zenodo-Dep="14001815" httpUri="https://zenodo.org/record/14001815/files/figure.png" pageId="30" pageNumber="31" startId="30.[191,235,1928,1954]" targetBox="[189,1398,529,1888]" targetPageId="30">
<paragraph id="B941511CFFE6FFD4FF150FACFAE6D8E6" blockId="30.[189,1399,1928,2025]" pageId="30" pageNumber="31">
<emphasis id="8B8A8D0EFFE6FFD4FF150FACFEF6D8AC" bold="true" box="[191,291,1928,1954]" pageId="30" pageNumber="31">Fig. 10.</emphasis>
<taxonomicName id="7EFE2A9FFFE6FFD4FE840FACFDFED8AC" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[302,555,1928,1954]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="polotowae" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFE6FFD4FE840FACFDFED8AC" box="[302,555,1928,1954]" italics="true" pageId="30" pageNumber="31">Tabiboka polotowae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFE6FFD4FD9D0FACFCD5D8AC" box="[567,768,1928,1954]" pageId="30" pageNumber="31" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, male and female habitus; A, C = male paratype (CASENT_9016118; CAS); B, D = female paratype (CASENT_9016116; CAS).
<emphasis id="8B8A8D0EFFE6FFD4FB320F8FFB1BD8CB" bold="true" box="[1176,1230,1963,1989]" pageId="30" pageNumber="31">A–B</emphasis>
. Dorsal view.
<emphasis id="8B8A8D0EFFE6FFD4FF150FEAFF23D8E6" bold="true" box="[191,246,1998,2024]" pageId="30" pageNumber="31">C–D</emphasis>
. Ventral view. Scale bars = 2.5 mm. Stacked microphotographs by Franklyn Cala Riquelme.
</paragraph>
</caption>
<paragraph id="B941511CFFE7FFD5FF170990FB93DEC0" blockId="31.[189,1094,436,463]" box="[189,1094,436,463]" pageId="31" pageNumber="32">
STERNUM (
<figureCitation id="21C54D99FFE7FFD5FE970990FE7DDEC0" box="[317,424,436,463]" captionStart="Fig" captionStartId="30.[191,235,1928,1954]" captionTargetBox="[189,1398,529,1888]" captionTargetId="figure-67@30.[189,1398,529,1888]" captionTargetPageId="30" captionText="Fig. 10. Tabiboka polotowae gen. et sp. nov., male and female habitus; A, C = male paratype (CASENT_9016118; CAS); B, D = female paratype (CASENT_9016116; CAS). A–B. Dorsal view. C–D. Ventral view. Scale bars = 2.5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001815" httpUri="https://zenodo.org/record/14001815/files/figure.png" pageId="31" pageNumber="32">Fig. 10C</figureCitation>
). Oval with gently curved margins; 1.70 wide, 2.20 long.
</paragraph>
<caption id="ED810194FFE7FFD5FF170E93FC95D8AA" ID-DOI="http://doi.org/10.5281/zenodo.14001819" ID-Zenodo-Dep="14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,231,1719,1745]" targetBox="[211,1371,542,1665]" targetPageId="31">
<paragraph id="B941511CFFE7FFD5FF170E93FC95D8AA" blockId="31.[187,1399,1719,1957]" pageId="31" pageNumber="32">
<emphasis id="8B8A8D0EFFE7FFD5FF170E93FEC6D9DF" bold="true" box="[189,275,1719,1745]" pageId="31" pageNumber="32">Fig. 11.</emphasis>
<taxonomicName id="7EFE2A9FFFE7FFD5FEB20E93FE28D9DF" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[280,509,1719,1745]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="polotowae" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFE7FFD5FEB20E93FE28D9DF" box="[280,509,1719,1745]" italics="true" pageId="31" pageNumber="32">Tabiboka polotowae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFE7FFD5FDA80E9CFD7FD9DC" box="[514,682,1720,1746]" pageId="31" pageNumber="32" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, genitalia drawings; A–C
<emphasis id="8B8A8D0EFFE7FFD5FC6D0E93FC0DD9DF" bold="true" box="[967,984,1719,1745]" pageId="31" pageNumber="32">=</emphasis>
male holotype (CASENT_9016117; CAS); D–E
<emphasis id="8B8A8D0EFFE7FFD5FEED0EFEFE8DD9FA" bold="true" box="[327,344,1754,1780]" pageId="31" pageNumber="32">=</emphasis>
female paratype (CASENT_9016117; CAS).
<emphasis id="8B8A8D0EFFE7FFD5FCC30EFEFCABD9FA" bold="true" box="[873,894,1754,1780]" pageId="31" pageNumber="32">A</emphasis>
. Left palp, prolateral view.
<emphasis id="8B8A8D0EFFE7FFD5FB100EFEFB1BD9FA" bold="true" box="[1210,1230,1754,1780]" pageId="31" pageNumber="32">B</emphasis>
. Idem, ventral view.
<emphasis id="8B8A8D0EFFE7FFD5FEAB0ED9FEC3D819" bold="true" box="[257,278,1789,1815]" pageId="31" pageNumber="32">C</emphasis>
. Idem, retrolateral view.
<emphasis id="8B8A8D0EFFE7FFD5FD9C0EDAFD9ED816" bold="true" box="[566,587,1790,1816]" pageId="31" pageNumber="32">D</emphasis>
. Epigyne, ventral view.
<emphasis id="8B8A8D0EFFE7FFD5FCC80EDAFCA3D816" bold="true" box="[866,886,1790,1816]" pageId="31" pageNumber="32">E</emphasis>
. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels.
</paragraph>
</caption>
<caption id="ED810194FFD8FFEAFF170E91FBD6D8E7" ID-DOI="http://doi.org/10.5281/zenodo.14001821" ID-Zenodo-Dep="14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="32" pageNumber="33" startId="32.[189,232,1717,1743]" targetBox="[277,1311,265,1679]" targetPageId="32">
<paragraph id="B941511CFFD8FFEAFF170E91FBD6D8E7" blockId="32.[187,1399,1717,2025]" pageId="32" pageNumber="33">
<emphasis id="8B8A8D0EFFD8FFEAFF170E91FECFD9C1" bold="true" box="[189,282,1717,1743]" pageId="32" pageNumber="33">Fig. 12.</emphasis>
<taxonomicName id="7EFE2A9FFFD8FFEAFE8B0E91FDDAD9C1" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[289,527,1717,1743]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="polotowae" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFD8FFEAFE8B0E91FDDAD9C1" box="[289,527,1717,1743]" italics="true" pageId="32" pageNumber="33">Tabiboka polotowae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFD8FFEAFDBD0E92FD19D9DE" box="[535,716,1718,1744]" pageId="32" pageNumber="33" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, SEM of male and female genitalia; A–F
<emphasis id="8B8A8D0EFFD8FFEAFB1D0E91FB1DD9C1" bold="true" box="[1207,1224,1717,1743]" pageId="32" pageNumber="33">=</emphasis>
male holotype (CASENT_9016117; CAS); G–H
<emphasis id="8B8A8D0EFFD8FFEAFDEA0EFCFD84D9FC" bold="true" box="[576,593,1752,1778]" pageId="32" pageNumber="33">=</emphasis>
female paratype (CASENT_9029889; CAS).
<emphasis id="8B8A8D0EFFD8FFEAFBF30EFCFBBBD9FC" bold="true" box="[1113,1134,1752,1778]" pageId="32" pageNumber="33">A</emphasis>
. Right palp (mirrored), prolateral, slightly ventral view.
<emphasis id="8B8A8D0EFFD8FFEAFD9D0ED8FD9ED818" bold="true" box="[567,587,1788,1814]" pageId="32" pageNumber="33">B</emphasis>
. Idem, ventral view.
<emphasis id="8B8A8D0EFFD8FFEAFCEB0EDFFC83D81B" bold="true" box="[833,854,1787,1813]" pageId="32" pageNumber="33">C</emphasis>
. Idem, retrolateral view.
<emphasis id="8B8A8D0EFFD8FFEAFBD10ED8FB45D818" bold="true" box="[1147,1168,1788,1814]" pageId="32" pageNumber="33">D</emphasis>
. Idem, dorsal view.
<emphasis id="8B8A8D0EFFD8FFEAFF160F3BFF05D837" bold="true" box="[188,208,1823,1849]" pageId="32" pageNumber="33">E</emphasis>
. Bulb, ventral view.
<emphasis id="8B8A8D0EFFD8FFEAFE650F3BFE0AD837" bold="true" box="[463,479,1823,1849]" pageId="32" pageNumber="33">F</emphasis>
. RTA, ventral view.
<emphasis id="8B8A8D0EFFD8FFEAFD710F3AFD25D836" bold="true" box="[731,752,1822,1848]" pageId="32" pageNumber="33">G</emphasis>
. Epigyne, ventral view.
<emphasis id="8B8A8D0EFFD8FFEAFBB20F3BFBF8D837" bold="true" box="[1048,1069,1823,1849]" pageId="32" pageNumber="33">H</emphasis>
. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–D = 0.5 mm; E–F = 0.15 mm; G–H = 0.1 mm. SEM images A–F by Charles Griswold and Darrell Ubick, G–H by Daniele Polotow.
</paragraph>
</caption>
<paragraph id="B941511CFFD9FFEBFF17092DFDF7DE2D" blockId="33.[189,546,265,291]" box="[189,546,265,291]" pageId="33" pageNumber="34">CHILUM. 0.08 high, 0.28 wide.</paragraph>
<paragraph id="B941511CFFD9FFEBFF170969FCCADE82" blockId="33.[188,1399,332,396]" pageId="33" pageNumber="34">EYES. ALE 0.11; AME 0.09; PLE 0.13; PME 0.13; ALE–AME 0.05; AME–AME 0.20; PLE–PME 0.33; PME–PME 0.13; MOQ: AW 0.23, PW 0.33, L 0.28.</paragraph>
<paragraph id="B941511CFFD9FFEBFF170992FD4BDEDE" blockId="33.[189,670,437,464]" box="[189,670,437,464]" pageId="33" pageNumber="34">CLYPEUS. 0.18 high, with four long setae.</paragraph>
<paragraph id="B941511CFFD9FFEBFF1709DDFDE9DD36" blockId="33.[187,1398,505,568]" pageId="33" pageNumber="34">LEGS. Trochanters shallowly notched. Unpaired claw smooth, well developed on all legs. Metatarsi lacking subbasal crack. Scopulae absent.</paragraph>
<paragraph id="B941511CFFD9FFEBFF170A46FE4FDD72" blockId="33.[189,410,610,636]" box="[189,410,610,636]" pageId="33" pageNumber="34">Leg measurements</paragraph>
<paragraph id="B941511CFFD9FFEBFEA00AAAFD21DC49" pageId="33" pageNumber="34">
<table id="CBFEA3BCFFD90035FF610AA9FD21DC46" box="[203,756,653,840]" gridcols="7" gridrows="5" pageId="33" pageNumber="34" rowsContinueIn="33.[203,932,925,1111]">
<tr id="07CE535EFFD90035FF610AA9FD21DDA6" box="[203,756,653,680]" gridrow="0" pageId="33" pageNumber="34" rowspan-0="1">
<th id="441F3A22FFD90035FEA00AA9FEE8DDA6" box="[266,317,653,680]" gridcol="1" gridrow="0" pageId="33" pageNumber="34">F</th>
<th id="441F3A22FFD90035FEF50AA9FE47DDA6" box="[351,402,653,680]" gridcol="2" gridrow="0" pageId="33" pageNumber="34">P</th>
<th id="441F3A22FFD90035FE1E0AA9FE32DDA6" box="[436,487,653,680]" gridcol="3" gridrow="0" pageId="33" pageNumber="34">T</th>
<th id="441F3A22FFD90035FDA20AA9FDEEDDA6" box="[520,571,653,680]" gridcol="4" gridrow="0" pageId="33" pageNumber="34">Mt</th>
<th id="441F3A22FFD90035FDF70AA9FD45DDA6" box="[605,656,653,680]" gridcol="5" gridrow="0" pageId="33" pageNumber="34">t</th>
<th id="441F3A22FFD90035FD180AA9FD21DDA6" box="[690,756,653,680]" gridcol="6" gridrow="0" pageId="33" pageNumber="34">Total</th>
</tr>
<tr id="07CE535EFFD90035FF610A92FD21DDDF" box="[203,756,694,721]" gridrow="1" pageId="33" pageNumber="34">
<th id="441F3A22FFD90035FF610A92FF3FDDDF" box="[203,234,694,721]" gridcol="0" gridrow="1" pageId="33" pageNumber="34">I</th>
<td id="441F3A22FFD90035FEA00A92FEE8DDDF" box="[266,317,694,721]" gridcol="1" gridrow="1" pageId="33" pageNumber="34">3.05</td>
<td id="441F3A22FFD90035FEF50A92FE47DDDF" box="[351,402,694,721]" gridcol="2" gridrow="1" pageId="33" pageNumber="34">1.50</td>
<td id="441F3A22FFD90035FE1E0A92FE32DDDF" box="[436,487,694,721]" gridcol="3" gridrow="1" pageId="33" pageNumber="34">3.20</td>
<td id="441F3A22FFD90035FDA20A92FDEEDDDF" box="[520,571,694,721]" gridcol="4" gridrow="1" pageId="33" pageNumber="34">2.80</td>
<td id="441F3A22FFD90035FDF70A92FD45DDDF" box="[605,656,694,721]" gridcol="5" gridrow="1" pageId="33" pageNumber="34">1.80</td>
<td id="441F3A22FFD90035FD180A92FD21DDDF" box="[690,756,694,721]" gridcol="6" gridrow="1" pageId="33" pageNumber="34">12.35</td>
</tr>
<tr id="07CE535EFFD90035FF610AF9FD21DDF6" box="[203,756,733,760]" gridrow="2" pageId="33" pageNumber="34">
<th id="441F3A22FFD90035FF610AF9FF3FDDF6" box="[203,234,733,760]" gridcol="0" gridrow="2" pageId="33" pageNumber="34">II</th>
<td id="441F3A22FFD90035FEA00AF9FEE8DDF6" box="[266,317,733,760]" gridcol="1" gridrow="2" pageId="33" pageNumber="34">3.20</td>
<td id="441F3A22FFD90035FEF50AF9FE47DDF6" box="[351,402,733,760]" gridcol="2" gridrow="2" pageId="33" pageNumber="34">1.50</td>
<td id="441F3A22FFD90035FE1E0AF9FE32DDF6" box="[436,487,733,760]" gridcol="3" gridrow="2" pageId="33" pageNumber="34">2.50</td>
<td id="441F3A22FFD90035FDA20AF9FDEEDDF6" box="[520,571,733,760]" gridcol="4" gridrow="2" pageId="33" pageNumber="34">2.30</td>
<td id="441F3A22FFD90035FDF70AF9FD45DDF6" box="[605,656,733,760]" gridcol="5" gridrow="2" pageId="33" pageNumber="34">1.60</td>
<td id="441F3A22FFD90035FD180AF9FD21DDF6" box="[690,756,733,760]" gridcol="6" gridrow="2" pageId="33" pageNumber="34">11.10</td>
</tr>
<tr id="07CE535EFFD90035FF610B21FD21DC2E" box="[203,756,773,800]" gridrow="3" pageId="33" pageNumber="34">
<th id="441F3A22FFD90035FF610B21FF3FDC2E" box="[203,234,773,800]" gridcol="0" gridrow="3" pageId="33" pageNumber="34">III</th>
<td id="441F3A22FFD90035FEA00B21FEE8DC2E" box="[266,317,773,800]" gridcol="1" gridrow="3" pageId="33" pageNumber="34">3.00</td>
<td id="441F3A22FFD90035FEF50B21FE47DC2E" box="[351,402,773,800]" gridcol="2" gridrow="3" pageId="33" pageNumber="34">1.30</td>
<td id="441F3A22FFD90035FE1E0B21FE32DC2E" box="[436,487,773,800]" gridcol="3" gridrow="3" pageId="33" pageNumber="34">2.10</td>
<td id="441F3A22FFD90035FDA20B21FDEEDC2E" box="[520,571,773,800]" gridcol="4" gridrow="3" pageId="33" pageNumber="34">2.60</td>
<td id="441F3A22FFD90035FDF70B21FD45DC2E" box="[605,656,773,800]" gridcol="5" gridrow="3" pageId="33" pageNumber="34">1.50</td>
<td id="441F3A22FFD90035FD180B21FD21DC2E" box="[690,756,773,800]" gridcol="6" gridrow="3" pageId="33" pageNumber="34">10.50</td>
</tr>
<tr id="07CE535EFFD90035FF610B09FD21DC46" box="[203,756,813,840]" gridrow="4" pageId="33" pageNumber="34">
<th id="441F3A22FFD90035FF610B09FF3FDC46" box="[203,234,813,840]" gridcol="0" gridrow="4" pageId="33" pageNumber="34">IV</th>
<td id="441F3A22FFD90035FEA00B09FEE8DC46" box="[266,317,813,840]" gridcol="1" gridrow="4" pageId="33" pageNumber="34">3.90</td>
<td id="441F3A22FFD90035FEF50B09FE47DC46" box="[351,402,813,840]" gridcol="2" gridrow="4" pageId="33" pageNumber="34">1.40</td>
<td id="441F3A22FFD90035FE1E0B09FE32DC46" box="[436,487,813,840]" gridcol="3" gridrow="4" pageId="33" pageNumber="34">2.30</td>
<td id="441F3A22FFD90035FDA20B09FDEEDC46" box="[520,571,813,840]" gridcol="4" gridrow="4" pageId="33" pageNumber="34">4.30</td>
<td id="441F3A22FFD90035FDF70B09FD45DC46" box="[605,656,813,840]" gridcol="5" gridrow="4" pageId="33" pageNumber="34">2.00</td>
<td id="441F3A22FFD90035FD180B09FD21DC46" box="[690,756,813,840]" gridcol="6" gridrow="4" pageId="33" pageNumber="34">13.90</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFD9FFEBFF170B55FAB5DACB" blockId="33.[189,301,881,907]" lastBlockId="33.[188,1400,1152,1477]" pageId="33" pageNumber="34">
Spination MALE PALP (
<figureCitation id="21C54D99FFD9FFEBFEE60CA4FE0EDB94" box="[332,475,1152,1178]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="33" pageNumber="34">Figs 11A–C</figureCitation>
,
<figureCitation id="21C54D99FFD9FFEBFE4F0CA4FDE9DB94" box="[485,572,1152,1178]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; A–F = male holotype (CASENT_9016117; CAS); G–H= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–D = 0.5 mm; E–F = 0.15 mm; G–H = 0.1 mm. SEM images A–F by Charles Griswold and Darrell Ubick, G–H by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="33" pageNumber="34">12A–F</figureCitation>
). Tibia with narrow finger-shaped VTA at retroapical margin and broad, median RTA arising near tibia base; cymbium oval, without dorsal patch of chemosensitive setae, with broad, sharp, conical retrobasal cymbial extension (CE) that extends to tibial apex; subtegulum large, filling most of cymbial concavity, with basal ridge (SL) at 180° that interacts with tegulum; tegulum oval with broad, basal notch, beneath this, conical lobe (TL) at 180° that locks with subtegulum; embolus (E) arises from retromargin of tegulum, simple, narrow, elongate, extending past apex of tegulum; median apophysis (MA) as sclerotized hook arising from soft base; conductor (C) hyaline, translucent, with narrow base arising near E base and greatly widened at extremity; paraconductor (Pc) as finger-like membranous structure originating between bases of E and C and extending less than half length of E.
</paragraph>
<paragraph id="B941511CFFD9FFEBFEA00BB9FC71DB58" pageId="33" pageNumber="34">
<table id="CBFEA3BCFFD90035FF610BB9FC71DB59" box="[203,932,925,1111]" gridcols="5" gridrows="5" pageId="33" pageNumber="34" rowsContinueFrom="33.[203,756,653,840]">
<tr id="07CE535EFFD90035FF610BB9FC71DCB9" box="[203,932,925,951]" gridrow="0" pageId="33" pageNumber="34" rowspan-0="1">
<th id="441F3A22FFD90035FEA00BB9FEBBDCB9" box="[266,366,925,951]" gridcol="1" gridrow="0" pageId="33" pageNumber="34">F</th>
<th id="441F3A22FFD90035FE390BB9FE0CDCB9" box="[403,473,925,951]" gridcol="2" gridrow="0" pageId="33" pageNumber="34">P</th>
<th id="441F3A22FFD90035FE570BB9FD7ADCB9" box="[509,687,925,951]" gridcol="3" gridrow="0" pageId="33" pageNumber="34">T</th>
<th id="441F3A22FFD90035FD750BB9FC71DCB9" box="[735,932,925,951]" gridcol="4" gridrow="0" pageId="33" pageNumber="34">Mt</th>
</tr>
<tr id="07CE535EFFD90035FF610BE1FC71DCD1" box="[203,932,965,991]" gridrow="1" pageId="33" pageNumber="34">
<th id="441F3A22FFD90035FF610BE1FF3FDCD1" box="[203,234,965,991]" gridcol="0" gridrow="1" pageId="33" pageNumber="34">I</th>
<td id="441F3A22FFD90035FEA00BE1FEBBDCD1" box="[266,366,965,991]" gridcol="1" gridrow="1" pageId="33" pageNumber="34">pl1d3</td>
<td id="441F3A22FFD90035FE390BE1FE0CDCD1" box="[403,473,965,991]" gridcol="2" gridrow="1" pageId="33" pageNumber="34">–</td>
<td id="441F3A22FFD90035FE570BE1FD7ADCD1" box="[509,687,965,991]" gridcol="3" gridrow="1" pageId="33" pageNumber="34">pl2d1v2-2-2-2</td>
<td id="441F3A22FFD90035FD750BE1FC71DCD1" box="[735,932,965,991]" gridcol="4" gridrow="1" pageId="33" pageNumber="34">pl2rl2v2-2-2</td>
</tr>
<tr id="07CE535EFFD90035FF610BC8FC71DB09" box="[203,932,1004,1031]" gridrow="2" pageId="33" pageNumber="34">
<th id="441F3A22FFD90035FF610BC8FF3FDB09" box="[203,234,1004,1031]" gridcol="0" gridrow="2" pageId="33" pageNumber="34">II</th>
<td id="441F3A22FFD90035FEA00BC8FEBBDB09" box="[266,366,1004,1031]" gridcol="1" gridrow="2" pageId="33" pageNumber="34">pl1d3rl1</td>
<td id="441F3A22FFD90035FE390BC8FE0CDB09" box="[403,473,1004,1031]" gridcol="2" gridrow="2" pageId="33" pageNumber="34">–</td>
<td id="441F3A22FFD90035FE570BC8FD7ADB09" box="[509,687,1004,1031]" gridcol="3" gridrow="2" pageId="33" pageNumber="34">pl2v2-2-2-2</td>
<td id="441F3A22FFD90035FD750BC8FC71DB09" box="[735,932,1004,1031]" gridcol="4" gridrow="2" pageId="33" pageNumber="34">pl3d1rl2v2-2-2</td>
</tr>
<tr id="07CE535EFFD90035FF610C30FC71DB21" box="[203,932,1044,1071]" gridrow="3" pageId="33" pageNumber="34">
<th id="441F3A22FFD90035FF610C30FF3FDB21" box="[203,234,1044,1071]" gridcol="0" gridrow="3" pageId="33" pageNumber="34">III</th>
<td id="441F3A22FFD90035FEA00C30FEBBDB21" box="[266,366,1044,1071]" gridcol="1" gridrow="3" pageId="33" pageNumber="34">pl2d2rl3</td>
<td id="441F3A22FFD90035FE390C30FE0CDB21" box="[403,473,1044,1071]" gridcol="2" gridrow="3" pageId="33" pageNumber="34">pl1rl1</td>
<td id="441F3A22FFD90035FE570C30FD7ADB21" box="[509,687,1044,1071]" gridcol="3" gridrow="3" pageId="33" pageNumber="34">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFD90035FD750C30FC71DB21" box="[735,932,1044,1071]" gridcol="4" gridrow="3" pageId="33" pageNumber="34">pl4rl3v2-2-2</td>
</tr>
<tr id="07CE535EFFD90035FF610C18FC71DB59" box="[203,932,1084,1111]" gridrow="4" pageId="33" pageNumber="34">
<th id="441F3A22FFD90035FF610C18FF3FDB59" box="[203,234,1084,1111]" gridcol="0" gridrow="4" pageId="33" pageNumber="34">IV</th>
<td id="441F3A22FFD90035FEA00C18FEBBDB59" box="[266,366,1084,1111]" gridcol="1" gridrow="4" pageId="33" pageNumber="34">pl2d3rl1</td>
<td id="441F3A22FFD90035FE390C18FE0CDB59" box="[403,473,1084,1111]" gridcol="2" gridrow="4" pageId="33" pageNumber="34">rl1</td>
<td id="441F3A22FFD90035FE570C18FD7ADB59" box="[509,687,1084,1111]" gridcol="3" gridrow="4" pageId="33" pageNumber="34">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFD90035FD750C18FC71DB59" box="[735,932,1084,1111]" gridcol="4" gridrow="4" pageId="33" pageNumber="34">pl3rl4v1-1-2-2-2</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFD9FFEBFF400DCAFCD2D906" blockId="33.[189,1174,1518,1582]" box="[234,775,1518,1544]" pageId="33" pageNumber="34">
<emphasis id="8B8A8D0EFFD9FFEBFF400DCAFE68D906" bold="true" box="[234,445,1518,1544]" pageId="33" pageNumber="34">
Female
<typeStatus id="6645EFBEFFD9FFEBFEE70DCAFE68D906" box="[333,445,1518,1544]" pageId="33" pageNumber="34" type="paratype">paratype</typeStatus>
</emphasis>
(CASENT_9016116; CAS)
</paragraph>
<paragraph id="B941511CFFD9FFEBFF170E30FB43D920" blockId="33.[189,1174,1518,1582]" box="[189,1174,1555,1582]" pageId="33" pageNumber="34">MEASUREMENTS. Total length: 11.50. Carapace: length 5.20, width 3.60, height 2.00.</paragraph>
<paragraph id="B941511CFFD9FFEBFF170E73FAF0D97F" blockId="33.[189,1317,1623,1650]" box="[189,1317,1623,1650]" pageId="33" pageNumber="34">
COLOUR (
<figureCitation id="21C54D99FFD9FFEBFE9A0E73FE6AD97F" box="[304,447,1623,1650]" captionStart="Fig" captionStartId="30.[191,235,1928,1954]" captionTargetBox="[189,1398,529,1888]" captionTargetId="figure-67@30.[189,1398,529,1888]" captionTargetPageId="30" captionText="Fig. 10. Tabiboka polotowae gen. et sp. nov., male and female habitus; A, C = male paratype (CASENT_9016118; CAS); B, D = female paratype (CASENT_9016116; CAS). A–B. Dorsal view. C–D. Ventral view. Scale bars = 2.5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001815" httpUri="https://zenodo.org/record/14001815/files/figure.png" pageId="33" pageNumber="34">Fig. 10B, D</figureCitation>
). As in male except ocular area not all black and legs only faintly marked.
</paragraph>
<paragraph id="B941511CFFD9FFEBFF170EBFFBB5D9BB" blockId="33.[189,1120,1691,1717]" box="[189,1120,1691,1717]" pageId="33" pageNumber="34">
STERNUM (
<figureCitation id="21C54D99FFD9FFEBFE970EBFFE7FD9BB" box="[317,426,1691,1717]" captionStart="Fig" captionStartId="30.[191,235,1928,1954]" captionTargetBox="[189,1398,529,1888]" captionTargetId="figure-67@30.[189,1398,529,1888]" captionTargetPageId="30" captionText="Fig. 10. Tabiboka polotowae gen. et sp. nov., male and female habitus; A, C = male paratype (CASENT_9016118; CAS); B, D = female paratype (CASENT_9016116; CAS). A–B. Dorsal view. C–D. Ventral view. Scale bars = 2.5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001815" httpUri="https://zenodo.org/record/14001815/files/figure.png" pageId="33" pageNumber="34">Fig. 10D</figureCitation>
). Oval with slightly sinuous margins; 2.10 wide, 2.40 long.
</paragraph>
<paragraph id="B941511CFFD9FFEBFF170EFAFDF7D9F6" blockId="33.[189,546,1758,1784]" box="[189,546,1758,1784]" pageId="33" pageNumber="34">CHILUM. 0.09 high, 0.33 wide.</paragraph>
<paragraph id="B941511CFFD9FFEBFF170F06FCCAD86F" blockId="33.[188,1398,1826,1890]" pageId="33" pageNumber="34">EYES. ALE 0.14; AME 0.08; PLE 0.15; PME 0.13; ALE–AME 0.06; AME–AME 0.25; PLE–PME 0.20; PME–PME 0.09; MOQ: AW 0.28, PW 0.38, L 0.28.</paragraph>
<paragraph id="B941511CFFD9FFEBFF170FAFFD4BD8AB" blockId="33.[189,670,1931,1957]" box="[189,670,1931,1957]" pageId="33" pageNumber="34">CLYPEUS. 0.10 high, with four long setae.</paragraph>
<paragraph id="B941511CFFD9FFEBFF170FEAFC39D8E6" blockId="33.[189,1004,1998,2025]" box="[189,1004,1998,2025]" pageId="33" pageNumber="34">LEGS. Very weak scopula beneath tarsus I, scopulae otherwise absent.</paragraph>
<paragraph id="B941511CFFDAFFE8FF17092DFE4FDE2D" blockId="34.[189,410,265,291]" box="[189,410,265,291]" pageId="34" pageNumber="35">Leg measurements</paragraph>
<paragraph id="B941511CFFDAFFE8FEA00911FD21DEE1" pageId="34" pageNumber="35">
<table id="CBFEA3BCFFDA0035FF610910FD21DEE1" box="[203,756,308,495]" gridcols="7" gridrows="5" pageId="34" pageNumber="35" rowsContinueIn="34.[203,923,580,765]">
<tr id="07CE535EFFDA0035FF610910FD21DE41" box="[203,756,308,335]" gridrow="0" pageId="34" pageNumber="35" rowspan-0="1">
<th id="441F3A22FFDA0035FEA00910FEE8DE41" box="[266,317,308,335]" gridcol="1" gridrow="0" pageId="34" pageNumber="35">F</th>
<th id="441F3A22FFDA0035FEF50910FE47DE41" box="[351,402,308,335]" gridcol="2" gridrow="0" pageId="34" pageNumber="35">P</th>
<th id="441F3A22FFDA0035FE1E0910FE32DE41" box="[436,487,308,335]" gridcol="3" gridrow="0" pageId="34" pageNumber="35">T</th>
<th id="441F3A22FFDA0035FDA20910FDEEDE41" box="[520,571,308,335]" gridcol="4" gridrow="0" pageId="34" pageNumber="35">Mt</th>
<th id="441F3A22FFDA0035FDF70910FD45DE41" box="[605,656,308,335]" gridcol="5" gridrow="0" pageId="34" pageNumber="35">t</th>
<th id="441F3A22FFDA0035FD180910FD21DE41" box="[690,756,308,335]" gridcol="6" gridrow="0" pageId="34" pageNumber="35">Total</th>
</tr>
<tr id="07CE535EFFDA0035FF610979FD21DE79" box="[203,756,349,375]" gridrow="1" pageId="34" pageNumber="35">
<th id="441F3A22FFDA0035FF610979FF3FDE79" box="[203,234,349,375]" gridcol="0" gridrow="1" pageId="34" pageNumber="35">I</th>
<td id="441F3A22FFDA0035FEA00979FEE8DE79" box="[266,317,349,375]" gridcol="1" gridrow="1" pageId="34" pageNumber="35">3.30</td>
<td id="441F3A22FFDA0035FEF50979FE47DE79" box="[351,402,349,375]" gridcol="2" gridrow="1" pageId="34" pageNumber="35">1.60</td>
<td id="441F3A22FFDA0035FE1E0979FE32DE79" box="[436,487,349,375]" gridcol="3" gridrow="1" pageId="34" pageNumber="35">2.70</td>
<td id="441F3A22FFDA0035FDA20979FDEEDE79" box="[520,571,349,375]" gridcol="4" gridrow="1" pageId="34" pageNumber="35">2.20</td>
<td id="441F3A22FFDA0035FDF70979FD45DE79" box="[605,656,349,375]" gridcol="5" gridrow="1" pageId="34" pageNumber="35">1.60</td>
<td id="441F3A22FFDA0035FD180979FD21DE79" box="[690,756,349,375]" gridcol="6" gridrow="1" pageId="34" pageNumber="35">11.40</td>
</tr>
<tr id="07CE535EFFDA0035FF6109A0FD21DE91" box="[203,756,388,415]" gridrow="2" pageId="34" pageNumber="35">
<th id="441F3A22FFDA0035FF6109A0FF3FDE91" box="[203,234,388,415]" gridcol="0" gridrow="2" pageId="34" pageNumber="35">II</th>
<td id="441F3A22FFDA0035FEA009A0FEE8DE91" box="[266,317,388,415]" gridcol="1" gridrow="2" pageId="34" pageNumber="35">3.05</td>
<td id="441F3A22FFDA0035FEF509A0FE47DE91" box="[351,402,388,415]" gridcol="2" gridrow="2" pageId="34" pageNumber="35">1.70</td>
<td id="441F3A22FFDA0035FE1E09A0FE32DE91" box="[436,487,388,415]" gridcol="3" gridrow="2" pageId="34" pageNumber="35">2.30</td>
<td id="441F3A22FFDA0035FDA209A0FDEEDE91" box="[520,571,388,415]" gridcol="4" gridrow="2" pageId="34" pageNumber="35">2.25</td>
<td id="441F3A22FFDA0035FDF709A0FD45DE91" box="[605,656,388,415]" gridcol="5" gridrow="2" pageId="34" pageNumber="35">1.40</td>
<td id="441F3A22FFDA0035FD1809A0FD21DE91" box="[690,756,388,415]" gridcol="6" gridrow="2" pageId="34" pageNumber="35">10.70</td>
</tr>
<tr id="07CE535EFFDA0035FF610988FD21DEC9" box="[203,756,428,455]" gridrow="3" pageId="34" pageNumber="35">
<th id="441F3A22FFDA0035FF610988FF3FDEC9" box="[203,234,428,455]" gridcol="0" gridrow="3" pageId="34" pageNumber="35">III</th>
<td id="441F3A22FFDA0035FEA00988FEE8DEC9" box="[266,317,428,455]" gridcol="1" gridrow="3" pageId="34" pageNumber="35">2.80</td>
<td id="441F3A22FFDA0035FEF50988FE47DEC9" box="[351,402,428,455]" gridcol="2" gridrow="3" pageId="34" pageNumber="35">1.30</td>
<td id="441F3A22FFDA0035FE1E0988FE32DEC9" box="[436,487,428,455]" gridcol="3" gridrow="3" pageId="34" pageNumber="35">2.00</td>
<td id="441F3A22FFDA0035FDA20988FDEEDEC9" box="[520,571,428,455]" gridcol="4" gridrow="3" pageId="34" pageNumber="35">2.30</td>
<td id="441F3A22FFDA0035FDF70988FD45DEC9" box="[605,656,428,455]" gridcol="5" gridrow="3" pageId="34" pageNumber="35">1.50</td>
<td id="441F3A22FFDA0035FD180988FD21DEC9" box="[690,756,428,455]" gridcol="6" gridrow="3" pageId="34" pageNumber="35">9.90</td>
</tr>
<tr id="07CE535EFFDA0035FF6109F0FD21DEE1" box="[203,756,468,495]" gridrow="4" pageId="34" pageNumber="35">
<th id="441F3A22FFDA0035FF6109F0FF3FDEE1" box="[203,234,468,495]" gridcol="0" gridrow="4" pageId="34" pageNumber="35">IV</th>
<td id="441F3A22FFDA0035FEA009F0FEE8DEE1" box="[266,317,468,495]" gridcol="1" gridrow="4" pageId="34" pageNumber="35">3.90</td>
<td id="441F3A22FFDA0035FEF509F0FE47DEE1" box="[351,402,468,495]" gridcol="2" gridrow="4" pageId="34" pageNumber="35">1.60</td>
<td id="441F3A22FFDA0035FE1E09F0FE32DEE1" box="[436,487,468,495]" gridcol="3" gridrow="4" pageId="34" pageNumber="35">3.00</td>
<td id="441F3A22FFDA0035FDA209F0FDEEDEE1" box="[520,571,468,495]" gridcol="4" gridrow="4" pageId="34" pageNumber="35">4.00</td>
<td id="441F3A22FFDA0035FDF709F0FD45DEE1" box="[605,656,468,495]" gridcol="5" gridrow="4" pageId="34" pageNumber="35">2.00</td>
<td id="441F3A22FFDA0035FD1809F0FD21DEE1" box="[690,756,468,495]" gridcol="6" gridrow="4" pageId="34" pageNumber="35">14.50</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFDAFFE8FF170A3CFBD4DCDB" blockId="34.[189,301,536,562]" lastBlockId="34.[188,1401,806,982]" pageId="34" pageNumber="35">
Spination GENITALIA. Epigyne (
<figureCitation id="21C54D99FFDAFFE8FE6B0B02FDE0DC4F" box="[449,565,806,833]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="34" pageNumber="35">Figs 11D</figureCitation>
,
<figureCitation id="21C54D99FFDAFFE8FDEE0B03FDADDC4F" box="[580,632,807,833]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; A–F = male holotype (CASENT_9016117; CAS); G–H= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–D = 0.5 mm; E–F = 0.15 mm; G–H = 0.1 mm. SEM images A–F by Charles Griswold and Darrell Ubick, G–H by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="34" pageNumber="35">12G</figureCitation>
) median lobe (ML) rectangular, elongate, narrowest in middle with apex of ML extending beyond epigastric furrow; lateral lobe (LL) convex, with transverse curved concavity posteriorly, narrow, only 0.25 maximum width of ML, copulatory openings (CO) hidden beneath anterolateral margins of ML. Endogyne (
<figureCitation id="21C54D99FFDAFFE8FD540BB2FCBFDCBE" box="[766,874,918,944]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="34" pageNumber="35">Figs 11E</figureCitation>
,
<figureCitation id="21C54D99FFDAFFE8FCDC0BB2FC78DCBE" box="[886,941,918,944]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; A–F = male holotype (CASENT_9016117; CAS); G–H= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–D = 0.5 mm; E–F = 0.15 mm; G–H = 0.1 mm. SEM images A–F by Charles Griswold and Darrell Ubick, G–H by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="34" pageNumber="35">12H</figureCitation>
): spermathecae (S) simple, oval, length 1.06 times width, with scattered pores anteriorly on spermathecal head.
</paragraph>
<paragraph id="B941511CFFDAFFE8FEA00A60FC4EDDF3" pageId="34" pageNumber="35">
<table id="CBFEA3BCFFDA0035FF610A60FC4EDDF3" box="[203,923,580,765]" gridcols="5" gridrows="5" pageId="34" pageNumber="35" rowsContinueFrom="34.[203,756,308,495]">
<tr id="07CE535EFFDA0035FF610A60FC4EDD50" box="[203,923,580,606]" gridrow="0" pageId="34" pageNumber="35" rowspan-0="1">
<th id="441F3A22FFDA0035FEA00A60FEBBDD50" box="[266,366,580,606]" gridcol="1" gridrow="0" pageId="34" pageNumber="35">F</th>
<th id="441F3A22FFDA0035FE350A60FE7ADD50" box="[415,431,580,606]" gridcol="2" gridrow="0" pageId="34" pageNumber="35">P</th>
<th id="441F3A22FFDA0035FE640A60FD55DD50" box="[462,640,580,606]" gridcol="3" gridrow="0" pageId="34" pageNumber="35">T</th>
<th id="441F3A22FFDA0035FD130A60FC4EDD50" box="[697,923,580,606]" gridcol="4" gridrow="0" pageId="34" pageNumber="35">Mt</th>
</tr>
<tr id="07CE535EFFDA0035FF610A4FFC4EDD88" box="[203,923,619,646]" gridrow="1" pageId="34" pageNumber="35">
<th id="441F3A22FFDA0035FF610A4FFF3FDD88" box="[203,234,619,646]" gridcol="0" gridrow="1" pageId="34" pageNumber="35">I</th>
<td id="441F3A22FFDA0035FEA00A4FFEBBDD88" box="[266,366,619,646]" gridcol="1" gridrow="1" pageId="34" pageNumber="35">pl1d1</td>
<td id="441F3A22FFDA0035FE350A4FFE7ADD88" box="[415,431,619,646]" gridcol="2" gridrow="1" pageId="34" pageNumber="35">–</td>
<td id="441F3A22FFDA0035FE640A4FFD55DD88" box="[462,640,619,646]" gridcol="3" gridrow="1" pageId="34" pageNumber="35">v2-2-2-2</td>
<td id="441F3A22FFDA0035FD130A4FFC4EDD88" box="[697,923,619,646]" gridcol="4" gridrow="1" pageId="34" pageNumber="35">v2-2-2</td>
</tr>
<tr id="07CE535EFFDA0035FF610AB7FC4EDDA0" box="[203,923,659,686]" gridrow="2" pageId="34" pageNumber="35">
<th id="441F3A22FFDA0035FF610AB7FF3FDDA0" box="[203,234,659,686]" gridcol="0" gridrow="2" pageId="34" pageNumber="35">II</th>
<td id="441F3A22FFDA0035FEA00AB7FEBBDDA0" box="[266,366,659,686]" gridcol="1" gridrow="2" pageId="34" pageNumber="35">pl1d1</td>
<td id="441F3A22FFDA0035FE350AB7FE7ADDA0" box="[415,431,659,686]" gridcol="2" gridrow="2" pageId="34" pageNumber="35">–</td>
<td id="441F3A22FFDA0035FE640AB7FD55DDA0" box="[462,640,659,686]" gridcol="3" gridrow="2" pageId="34" pageNumber="35">v2-2-2</td>
<td id="441F3A22FFDA0035FD130AB7FC4EDDA0" box="[697,923,659,686]" gridcol="4" gridrow="2" pageId="34" pageNumber="35">v2-2-2</td>
</tr>
<tr id="07CE535EFFDA0035FF610A9FFC4EDDD8" box="[203,923,699,726]" gridrow="3" pageId="34" pageNumber="35">
<th id="441F3A22FFDA0035FF610A9FFF3FDDD8" box="[203,234,699,726]" gridcol="0" gridrow="3" pageId="34" pageNumber="35">III</th>
<td id="441F3A22FFDA0035FEA00A9FFEBBDDD8" box="[266,366,699,726]" gridcol="1" gridrow="3" pageId="34" pageNumber="35">pl2d3rl1</td>
<td id="441F3A22FFDA0035FE350A9FFE7ADDD8" box="[415,431,699,726]" gridcol="2" gridrow="3" pageId="34" pageNumber="35">–</td>
<td id="441F3A22FFDA0035FE640A9FFD55DDD8" box="[462,640,699,726]" gridcol="3" gridrow="3" pageId="34" pageNumber="35">pl2d1rl2v2-2-2</td>
<td id="441F3A22FFDA0035FD130A9FFC4EDDD8" box="[697,923,699,726]" gridcol="4" gridrow="3" pageId="34" pageNumber="35">pl5rl3v2-2-3</td>
</tr>
<tr id="07CE535EFFDA0035FF610AC7FC4EDDF3" box="[203,923,739,765]" gridrow="4" pageId="34" pageNumber="35">
<th id="441F3A22FFDA0035FF610AC7FF3FDDF3" box="[203,234,739,765]" gridcol="0" gridrow="4" pageId="34" pageNumber="35">IV</th>
<td id="441F3A22FFDA0035FEA00AC7FEBBDDF3" box="[266,366,739,765]" gridcol="1" gridrow="4" pageId="34" pageNumber="35">pl1d2rl1</td>
<td id="441F3A22FFDA0035FE350AC7FE7ADDF3" box="[415,431,739,765]" gridcol="2" gridrow="4" pageId="34" pageNumber="35">–</td>
<td id="441F3A22FFDA0035FE640AC7FD55DDF3" box="[462,640,739,765]" gridcol="3" gridrow="4" pageId="34" pageNumber="35">pl1d2v1-2-2</td>
<td id="441F3A22FFDA0035FD130AC7FC4EDDF3" box="[697,923,739,765]" gridcol="4" gridrow="4" pageId="34" pageNumber="35">pl4d2rl1v1-1-2-2-3</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFDAFFE8FF170BDAFEE6DB16" blockId="34.[189,307,1022,1048]" box="[189,307,1022,1048]" pageId="34" pageNumber="35">
<heading id="E209E670FFDAFFE8FF170BDAFEE6DB16" bold="true" box="[189,307,1022,1048]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="8B8A8D0EFFDAFFE8FF170BDAFEE6DB16" bold="true" box="[189,307,1022,1048]" pageId="34" pageNumber="35">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFDAFFE8FF400C0BFE54DB47" blockId="34.[189,1399,1071,1358]" box="[234,385,1071,1098]" pageId="34" pageNumber="35">
<emphasis id="8B8A8D0EFFDAFFE8FF400C0BFEFFDB47" bold="true" box="[234,298,1071,1097]" pageId="34" pageNumber="35">Male</emphasis>
(n = 2)
</paragraph>
<paragraph id="B941511CFFDAFFE8FF170C70FBB7DA40" blockId="34.[189,1399,1071,1358]" pageId="34" pageNumber="35">Total length = 8.10–8.50, carapace length/width = 1.32–1.34, carapace height/width = 0.40–0.56, PER/OAL = 2.21–2.22, PER/AER = 1.33–1.38, OAL/OQL = 1.22–1.23, OQP/OQA = 1.44–2.00, clypeus height/AME = 1.00–2.33, cheliceral length/clypeus height = 8.57–17.00, sternum length/width = 1.05–1.29, palpal coxa length/width = 2.08–2.17, femur I length/carapace width = 0.90–1.06, metatarsus I length/carapace width = 0.82–0.86, cymbium length/carapace width = 0.37–0.40, cymbium length/palpal patella length = 2.15–2.27, cymbium length/palpal tibia length = 1.75–2.27, cymbium length/palpal femur length = 0.78–1.00, palpal tibia length/palpal patella length = 1.00–1.23.</paragraph>
<paragraph id="B941511CFFDAFFE8FF400D53FE48DA9F" blockId="34.[189,1400,1399,1834]" box="[234,413,1399,1425]" pageId="34" pageNumber="35">
<emphasis id="8B8A8D0EFFDAFFE8FF400D53FE90DA9F" bold="true" box="[234,325,1399,1425]" pageId="34" pageNumber="35">Female</emphasis>
(n = 3)
</paragraph>
<paragraph id="B941511CFFDAFFE8FF170DB8FCF2D824" blockId="34.[189,1400,1399,1834]" pageId="34" pageNumber="35">
Total length = 8.40–17.00, carapace length/width = 1.31–1.46, carapace height/ width = 0.44–0.75, PER/carapace width = 0.25–0.27, PER/OAL = 2.40–2.88, PER/AER = 1.31–1.45, OAL/OQL = 1.20– 1.36, OQP/OQA= 1.36–1.56, clypeus height/AME = 1.33–2.33, cheliceral length/clypeus height = 10.67– 20.00, sternum length/width = 1.14–1.22, palpal coxa length/width = 1.57–2.26, femur I length/carapace width = 0.89–1.32, metatarsus I length/carapace width = 0.61–0.64, femur IV length/carapace width = 1.00–1.18, palpal tarsus length/carapace width = 0.33–0.38, palpal tarsus length/palpal tibia length = 1.91–2.15, palpal tarsus length/palpal tibia length = 1.33–1.75, palpal tarsus length/palpal femur length = 0.88–0.89, palpal tibia length/palpal patella length = 1.09–1.23. Epigyne with median lobe length 1.29– 1.38 times width; median lobe maximum width 1.34–2.10 times minimum width; median lobe width 2.00–5.00 times lateral lobe width; median lobe extends to (
<figureCitation id="21C54D99FFDAFFE8FC2C0ECFFC27D80B" box="[902,1010,1771,1797]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; A–F = male holotype (CASENT_9016117; CAS); G–H= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: A–D = 0.5 mm; E–F = 0.15 mm; G–H = 0.1 mm. SEM images A–F by Charles Griswold and Darrell Ubick, G–H by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="34" pageNumber="35">Fig. 12G</figureCitation>
) or beyond (
<figureCitation id="21C54D99FFDAFFE8FB210ECFFB20D80B" box="[1163,1269,1771,1797]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="34" pageNumber="35">Fig. 11D</figureCitation>
) epigastric furrow. Spermatheca (each) length 1.06 times width.
</paragraph>
<paragraph id="B941511CFFDAFFE8FF170F70FE74D860" blockId="34.[189,417,1876,1902]" box="[189,417,1876,1902]" pageId="34" pageNumber="35">
<heading id="E209E670FFDAFFE8FF170F70FE74D860" bold="true" box="[189,417,1876,1902]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="8B8A8D0EFFDAFFE8FF170F70FE74D860" bold="true" box="[189,417,1876,1902]" pageId="34" pageNumber="35">Note on synonyms</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFDAFFE8FF170FA0FD23D8E6" blockId="34.[189,1399,1924,2025]" pageId="34" pageNumber="35">
Specimens of this species have been used as exemplars in a previously published molecular phylogeny: they were listed by
<bibRefCitation id="DD6F2CEDFFDAFFE8FE1D0F8DFD17D8CA" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[439,706,1961,1988]" pageId="34" pageNumber="35" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow
<emphasis id="8B8A8D0EFFDAFFE8FD8C0F8EFDB3D8CD" box="[550,614,1961,1987]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
(2015)
</bibRefCitation>
as “
<taxonomicName id="7EFE2A9FFFDAFFE8FD540F8DFC0DD8CD" box="[766,984,1961,1988]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="undefined-3">
<emphasis id="8B8A8D0EFFDAFFE8FD540F8DFC44D8CD" box="[766,913,1961,1987]" italics="true" pageId="34" pageNumber="35">Zorodictyna</emphasis>
sp. 3
</taxonomicName>
” (CASENT_9029890; CAS) and “
<taxonomicName id="7EFE2A9FFFDAFFE8FF600FEAFE4DD8E6" box="[202,408,1998,2025]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="undefined-4">
<emphasis id="8B8A8D0EFFDAFFE8FF600FEAFE8DD8E6" box="[202,344,1998,2024]" italics="true" pageId="34" pageNumber="35">Zorodictyna</emphasis>
sp. 4
</taxonomicName>
” (CASENT_9029889; CAS).
</paragraph>
<paragraph id="B941511CFFDBFFE9FF17092DFEABDE2C" blockId="35.[189,382,264,291]" box="[189,382,264,291]" pageId="35" pageNumber="36">
<heading id="E209E670FFDBFFE9FF17092DFEABDE2C" bold="true" box="[189,382,264,291]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="8B8A8D0EFFDBFFE9FF17092DFEABDE2C" bold="true" box="[189,382,264,291]" pageId="35" pageNumber="36">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFDBFFE9FF17091DFBE3DE79" blockId="35.[188,1399,313,376]" pageId="35" pageNumber="36">
The species
<taxonomicName id="7EFE2A9FFFDBFFE9FEE6091DFDEDDE5D" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[332,568,313,339]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="polotowae" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFDBFFE9FEE6091DFDEDDE5D" box="[332,568,313,339]" italics="true" pageId="35" pageNumber="36">Tabiboka polotowae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFDBFFE9FD95091DFD3BDE5D" box="[575,750,313,339]" pageId="35" pageNumber="36" rank="species">gen. et sp. nov.</taxonomicNameLabel>
is recorded from both highly disturbed and intact forest habitats. Many specimens have been collected while concealed in leaf litter.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFDBFFE9FF170984FEAEDD1E" pageId="35" pageNumber="36" type="distribution">
<paragraph id="B941511CFFDBFFE9FF170984FE83DEB4" blockId="35.[189,342,416,442]" box="[189,342,416,442]" pageId="35" pageNumber="36">
<heading id="E209E670FFDBFFE9FF170984FE83DEB4" bold="true" box="[189,342,416,442]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="8B8A8D0EFFDBFFE9FF170984FE83DEB4" bold="true" box="[189,342,416,442]" pageId="35" pageNumber="36">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFDBFFE9FF1709F5FEAEDD1E" blockId="35.[189,1399,465,528]" pageId="35" pageNumber="36">
<taxonomicName id="7EFE2A9FFFDBFFE9FF1709F5FE7EDEE5" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[189,427,465,491]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="polotowae" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFDBFFE9FF1709F5FE7EDEE5" box="[189,427,465,491]" italics="true" pageId="35" pageNumber="36">Tabiboka polotowae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFDBFFE9FE1E09F5FDB3DEE5" box="[436,614,465,491]" pageId="35" pageNumber="36" rank="species">gen. et sp. nov.</taxonomicNameLabel>
occurs in the eastern uplands of
<collectingCountry id="C1E9118CFFDBFFE9FC4109F5FBACDEE5" box="[1003,1145,465,491]" name="Madagascar" pageId="35" pageNumber="36">Madagascar</collectingCountry>
from the south to the center (
<figureCitation id="21C54D99FFDBFFE9FEBF09D2FEB8DD1E" box="[277,365,502,528]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="35" pageNumber="36">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="AEB00F9D7D6791A3A06D0698CCE3CDA1">On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar</mods:title>
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<mods:namePart id="D5230FB178FAD456951673654F6909ED">Henrard, Arnaud</mods:namePart>
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<mods:namePart id="A0AF158EA275710AE93CEDCFEBFCC749">Griswold, Charles</mods:namePart>
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<mods:affiliation id="7EE1078138C9E0448387429271E6E74E">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA.</mods:affiliation>
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<mods:namePart id="AAEA78D23D8700EA1A0B1315380AECFC">Jocqué, Rudy</mods:namePart>
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<mods:affiliation id="FDE67A7434FB1A868E5F398CE44B7836">Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium.</mods:affiliation>
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<paragraph id="B941511CFFEFFFDDFDF60E9FFC0DD9D8" blockId="23.[364,1223,1723,1817]" box="[604,984,1723,1750]" pageId="23" pageNumber="24">
<heading id="E209E670FFEFFFDDFDF60E9FFC0DD9D8" box="[604,984,1723,1750]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<taxonomicName id="7EFE2A9FFFEFFFDDFDF60E9FFCF5D9D8" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[604,800,1723,1750]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="milloti" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFEFFFDDFDF60E9FFCF5D9D8" bold="true" box="[604,800,1723,1750]" italics="true" pageId="23" pageNumber="24">Tabiboka milloti</emphasis>
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<paragraph id="B941511CFFEFFFDDFD7B0EDBFCB7D817" blockId="23.[364,1223,1723,1817]" box="[721,866,1791,1817]" pageId="23" pageNumber="24">Figs 8–9, 26</paragraph>
<paragraph id="B941511CFFEFFFDDFF170F1BFD18D857" blockId="23.[189,717,1855,1882]" box="[189,717,1855,1882]" pageId="23" pageNumber="24">
<taxonomicName id="7EFE2A9FFFEFFFDDFF170F1BFEA1D854" box="[189,372,1855,1882]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8B8A8D0EFFEFFFDDFF170F1BFE9ED857" box="[189,331,1855,1881]" italics="true" pageId="23" pageNumber="24">Zorodictyna</emphasis>
sp.
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CG45 –
<bibRefCitation id="DD6F2CEDFFEFFFDDFE740F1BFD1CD857" author="Wheeler W. C. &amp; Coddington J. A. &amp; Crowley L. M. &amp; Dimitrov D. &amp; Goloboff P. A. &amp; Griswold C. E. &amp; Hormiga G. &amp; Prendini L. &amp; Ramirez M. J. &amp; Sierwald P. &amp; Almeida-Silva L. M. &amp; Alvarez-Padilla F. &amp; Arnedo M. A. &amp; Benavides L. R. &amp; Benjamin S. P. &amp; Bond J. E. &amp; Grismado C. J. &amp; Hasan E. &amp; Hedin M. &amp; Izquierdo M. A. &amp; Labarque F. M. &amp; Ledford J. &amp; Lopardo L. &amp; Maddison W. P. &amp; Miller J. A. &amp; Piacentini L. N. &amp; Platnick N. I. &amp; Polotow D. &amp; Silva-Davila D. &amp; Scharff N. &amp; Szuts T. &amp; Ubick D. &amp; Vink C. &amp; Wood H. M. &amp; Zhang J. X." box="[478,713,1855,1881]" pageId="23" pageNumber="24" pagination="576 - 616" refId="ref39136" refString="Wheeler W. C., Coddington J. A., Crowley L. M., Dimitrov D., Goloboff P. A., Griswold C. E., Hormiga G., Prendini L., Ramirez M. J., Sierwald P., Almeida-Silva L. M., Alvarez-Padilla F., Arnedo M. A., Benavides L. R., Benjamin S. P., Bond J. E., Grismado C. J., Hasan E., Hedin M., Izquierdo M. A., Labarque F. M., Ledford J., Lopardo L., Maddison W. P., Miller J. A., Piacentini L. N., Platnick N. I., Polotow D., Silva-Davila D., Scharff N., Szuts T., Ubick D., Vink C., Wood H. M. &amp; Zhang J. X. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. https: // doi. org / 10.1111 / cla. 12182" type="journal article" year="2017">
Wheeler
<emphasis id="8B8A8D0EFFEFFFDDFDE30F64FD51D857" box="[585,644,1855,1881]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2017
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.
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<subSubSection id="F1E40297FFEFFFD2FF170F5BFAA3DEF0" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="B941511CFFEFFFDDFF170F5BFEE3D897" blockId="23.[189,310,1919,1945]" box="[189,310,1919,1945]" pageId="23" pageNumber="24">
<heading id="E209E670FFEFFFDDFF170F5BFEE3D897" bold="true" box="[189,310,1919,1945]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="8B8A8D0EFFEFFFDDFF170F5BFEE3D897" bold="true" box="[189,310,1919,1945]" pageId="23" pageNumber="24">Diagnosis</emphasis>
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<paragraph id="B941511CFFEFFFD2FF170F88FAA3DEF0" blockId="23.[188,1398,1964,2025]" lastBlockId="24.[187,1400,265,511]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
The males of
<taxonomicName id="7EFE2A9FFFEFFFDDFEF30F88FDCDD8C8" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[345,536,1964,1990]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="milloti" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFEFFFDDFEF30F88FDCDD8C8" box="[345,536,1964,1990]" italics="true" pageId="23" pageNumber="24">Tabiboka milloti</emphasis>
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may be distinguished by the palpal tibia with a short, blunt retromedian process (RTA) and broad, thumb-like ventral apical process (VTA) (Figs 8E, G, 9B–C), the slender embolus (E) and the elongate, cruciform median apophysis (MA) with the apex extending to embolus tip (Figs 8F, 9A–B) (vs in
<taxonomicName id="7EFE2A9FFFE0FFD2FD28090AFD38DE49" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[642,749,301,327]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="milleri" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFE0FFD2FD28090AFD38DE49" box="[642,749,301,327]" italics="true" pageId="24" pageNumber="25">T. milleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFE0FFD2FD5C090AFC7FDE46" box="[758,938,302,328]" pageId="24" pageNumber="25" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, the the VTA is smaller and the RTA is greatly elongated,
<figureCitation id="21C54D99FFE0FFD2FE320976FDC2DE63" box="[408,535,338,365]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="24" pageNumber="25">Fig. 5A–C</figureCitation>
, and the E and the MA are shorted and broader; and in
<taxonomicName id="7EFE2A9FFFE0FFD2FB0C0977FAE8DE62" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1190,1341,338,364]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="polotowae" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFE0FFD2FB0C0977FAE8DE62" box="[1190,1341,338,364]" italics="true" pageId="24" pageNumber="25">T. polotowae</emphasis>
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<taxonomicNameLabel id="90B93075FFE0FFD2FAEF0976FEEFDE9F" pageId="24" pageNumber="25" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, the VTA is finger-like and the MA is shorter,
<figureCitation id="21C54D99FFE0FFD2FCC10952FC2EDE9E" box="[875,1019,374,400]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="24" pageNumber="25">Fig. 11A–C</figureCitation>
). Females of
<taxonomicName id="7EFE2A9FFFE0FFD2FB370953FADCDE9E" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1181,1289,374,400]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="milloti">
<emphasis id="8B8A8D0EFFE0FFD2FB370953FADCDE9E" box="[1181,1289,374,400]" italics="true" pageId="24" pageNumber="25">T. milloti</emphasis>
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have the epigyne with a heart-shaped median lobe (ML), broad anteriorly (anterior width greater than one-half total width,
<figureCitation id="21C54D99FFE0FFD2FEA1099BFEB1DED7" box="[267,356,447,473]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.A–C = male holotype (CASENT_9016063; MNHN). D–E = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="24" pageNumber="25">Fig. 9D</figureCitation>
) (vs in
<taxonomicName id="7EFE2A9FFFE0FFD2FE1309E4FDF7DED7" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[441,546,447,473]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="8B8A8D0EFFE0FFD2FE1309E4FDF7DED7" box="[441,546,447,473]" italics="true" pageId="24" pageNumber="25">T. milleri</emphasis>
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, the ML is narrow anteriorly, anterior width less than one-half total width,
<figureCitation id="21C54D99FFE0FFD2FF1109C0FECDDEF1" box="[187,280,484,511]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="24" pageNumber="25">Fig. 5D</figureCitation>
; and in
<taxonomicName id="7EFE2A9FFFE0FFD2FEC409C1FDD6DEF0" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[366,515,484,510]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="polotowae">
<emphasis id="8B8A8D0EFFE0FFD2FEC409C1FDD6DEF0" box="[366,515,484,510]" italics="true" pageId="24" pageNumber="25">T. polotowae</emphasis>
</taxonomicName>
, the ML is rectangular, narrowest in middle or with sides straight,
<figureCitation id="21C54D99FFE0FFD2FAAA09C0FABCDEF0" box="[1280,1385,484,510]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="24" pageNumber="25">Fig. 11D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFE0FFD2FF170A03FAE7DD9B" pageId="24" pageNumber="25" type="etymology">
<paragraph id="B941511CFFE0FFD2FF170A03FE91DD4F" blockId="24.[189,324,551,577]" box="[189,324,551,577]" pageId="24" pageNumber="25">
<heading id="E209E670FFE0FFD2FF170A03FE91DD4F" bold="true" box="[189,324,551,577]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="8B8A8D0EFFE0FFD2FF170A03FE91DD4F" bold="true" box="[189,324,551,577]" pageId="24" pageNumber="25">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE0FFD2FF170A72FAE7DD9B" blockId="24.[189,1399,598,661]" pageId="24" pageNumber="25">
The species name is a patronym in honor of Jacques Millot, who collected the
<typeStatus id="6645EFBEFFE0FFD2FBCA0A72FB12DD7E" box="[1120,1223,598,624]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
of this species and many other interesting arthropods in
<collectingCountry id="C1E9118CFFE0FFD2FD080A5FFCE5DD9B" box="[674,816,635,661]" name="Madagascar" pageId="24" pageNumber="25">Madagascar</collectingCountry>
during his missions in the mid-20
<superScript id="4E8BFC54FFE0FFD2FB150A5EFB19DD86" attach="left" box="[1215,1228,634,648]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
century.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFE0FFD2FF170A9AFB66DA27" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="B941511CFFE0FFD2FF170A9AFEB9DDD9" blockId="24.[189,364,701,728]" box="[189,364,701,728]" pageId="24" pageNumber="25">
<heading id="E209E670FFE0FFD2FF170A9AFEB9DDD9" bold="true" box="[189,364,701,728]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="8B8A8D0EFFE0FFD2FF170A9AFEB9DDD9" bold="true" box="[189,364,701,728]" pageId="24" pageNumber="25">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE0FFD2FF400AC9FE89DC09" blockId="24.[189,1398,749,849]" box="[234,348,749,775]" pageId="24" pageNumber="25">
<emphasis id="8B8A8D0EFFE0FFD2FF400AC9FE89DC09" bold="true" box="[234,348,749,775]" pageId="24" pageNumber="25">
<typeStatus id="6645EFBEFFE0FFD2FF400AC9FE89DC09" box="[234,348,749,775]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFE0FFD2FF170B36FC16DC5F" blockId="24.[189,1398,749,849]" pageId="24" pageNumber="25">
<materialsCitation id="09965B41FFE0FFD2FF170B36FC16DC5F" collectingDate="1946-08" collectionCode="MNHN" collectorName="J. Millot" country="Madagascar" location="Toamasina Province" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="holotype">
<collectingCountry id="C1E9118CFFE0FFD2FF170B36FE5BDC22" box="[189,398,786,812]" name="Madagascar" pageId="24" pageNumber="25">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFE0FFD2FE020B35FD71DC25" bold="true" box="[424,676,785,811]" pageId="24" pageNumber="25">
<collectingRegion id="7B3A9FFEFFE0FFD2FE020B35FD71DC25" box="[424,676,785,811]" country="Madagascar" name="Toamasina" pageId="24" pageNumber="25">Toamasina Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFE0FFD2FD160B36FD00DC22" box="[700,725,786,812]" count="1" pageId="24" pageNumber="25" type="male">♂</specimenCount>
; Périnet (= Andasibe), Res. Analamazaotra, Parc National Andasibe;
<date id="CD4077DCFFE0FFD2FE900B12FE6CDC5E" box="[314,441,822,848]" pageId="24" pageNumber="25" value="1946-08">
<collectingDate id="DD048E34FFE0FFD2FE900B12FE6CDC5E" box="[314,441,822,848]" pageId="24" pageNumber="25" value="1946-08">Aug. 1946</collectingDate>
</date>
;
<collectorName id="140B34CAFFE0FFD2FE6F0B13FDFDDC5E" box="[453,552,822,849]" pageId="24" pageNumber="25">J. Millot</collectorName>
leg.; CASENT_9016063;
<collectionCode id="DFEFC9D9FFE0FFD2FCCB0B13FC6ADC5F" box="[865,959,823,849]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="25">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B941511CFFE0FFD2FF400B5DFEB2DC9D" blockId="24.[189,1399,889,1098]" box="[234,359,889,915]" pageId="24" pageNumber="25">
<emphasis id="8B8A8D0EFFE0FFD2FF400B5DFEB2DC9D" bold="true" box="[234,359,889,915]" pageId="24" pageNumber="25">
<typeStatus id="6645EFBEFFE0FFD2FF400B5DFEB2DC9D" box="[234,359,889,915]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B941511CFFE0FFD2FF170BB9FA97DB47" blockId="24.[189,1399,889,1098]" pageId="24" pageNumber="25">
<materialsCitation id="09965B41FFE0FFD2FF170BB9FAF3DB0E" collectingDate="2003-01-16" collectingDateMax="2003-01-18" collectingDateMin="2003-01-16" collectionCode="CAS" collectorName="C. Griswold &amp; D. Silva &amp; D. Andriamalala" country="Madagascar" elevation="960" latitude="-18.943888" location="Parc National Andasibe" longLatPrecision="21" longitude="48.4175" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" stateProvince="Toamasina" typeStatus="paratype">
<collectingCountry id="C1E9118CFFE0FFD2FF170BB9FE58DCB9" box="[189,397,925,951]" name="Madagascar" pageId="24" pageNumber="25">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFE0FFD2FE0F0BB9FD76DCB9" bold="true" box="[421,675,925,951]" pageId="24" pageNumber="25">
<collectingRegion id="7B3A9FFEFFE0FFD2FE0F0BB9FD76DCB9" box="[421,675,925,951]" country="Madagascar" name="Toamasina" pageId="24" pageNumber="25">Toamasina Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFE0FFD2FD140BB9FD39DCB6" box="[702,748,925,952]" count="1" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
; Res. Analamazaotra,
<location id="BC2107C7FFE0FFD2FC5C0BBAFAC5DCB9" LSID="urn:lsid:plazi:treatment:3157E00AFFEFFFD6FDF60E9FFA8ED8E6:BC2107C7FFE0FFD2FC5C0BBAFAC5DCB9" box="[1014,1296,925,952]" country="Madagascar" latitude="-18.943888" longLatPrecision="21" longitude="48.4175" name="Parc National Andasibe" pageId="24" pageNumber="25" stateProvince="Toamasina">Parc National Andasibe</location>
, road
<locationDeviation id="A8F3483EFFE0FFD2FAF30BB9FE6DDCD2" location="Moramanga" pageId="24" pageNumber="25">
<quantity id="7E06FCF9FFE0FFD2FAF30BB9FF37DCD2" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="24" pageNumber="25" unit="km" value="23.0">23 km</quantity>
E of Moramanga
</locationDeviation>
;
<elevation id="32D3B62FFFE0FFD2FE6F0BE6FD84DCD2" box="[453,593,962,988]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="24" pageNumber="25" unit="m" value="960.0">
<quantity id="7E06FCF9FFE0FFD2FE6F0BE6FDC5DCD2" box="[453,528,962,988]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="24" pageNumber="25" unit="m" value="960.0">960 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="DCCA37DBFFE0FFD2FDCA0BE4FD24DCD2" box="[608,753,960,988]" degrees="18" direction="south" minutes="56" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="38" value="-18.943888">18°56′38″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFE0FFD2FD550BE4FC46DCD2" box="[767,915,960,988]" degrees="48" direction="east" minutes="25" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="03" value="48.4175">48°25′03″ E</geoCoordinate>
;
<date id="CD4077DCFFE0FFD2FC0A0BE6FBB2DCD2" box="[928,1127,962,988]" pageId="24" pageNumber="25" value="2003-01-16" valueMax="2003-01-18" valueMin="2003-01-16">
<collectingDate id="DD048E34FFE0FFD2FC0A0BE6FBB2DCD2" box="[928,1127,962,988]" pageId="24" pageNumber="25" value="2003-01-16" valueMax="2003-01-18" valueMin="2003-01-16">16–18 Jan. 2003</collectingDate>
</date>
;
<collectorName id="140B34CAFFE0FFD2FBDF0BE6FADDDCD2" box="[1141,1288,962,988]" pageId="24" pageNumber="25">C. Griswold</collectorName>
,
<collectorName id="140B34CAFFE0FFD2FABF0BE6FAA3DCD2" box="[1301,1398,962,988]" pageId="24" pageNumber="25">D. Silva</collectorName>
and
<collectorName id="140B34CAFFE0FFD2FF5A0BC3FE66DB0E" box="[240,435,998,1025]" pageId="24" pageNumber="25">D. Andriamalala</collectorName>
leg.; general collecting in rainforest; BLF7993; CASENT_9018308;
<collectionCode id="DFEFC9D9FFE0FFD2FB470BC2FAF3DB0E" box="[1261,1318,998,1024]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="24" pageNumber="25" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFE0FFD2FAEB0BC2FDB8DB47" collectingDate="2003-01-16" collectingDateMax="2003-01-18" collectingDateMin="2003-01-16" collectionCode="CAS" collectorName="C. Griswold &amp; D. Silva &amp; D. Andriamalala" country="Madagascar" elevation="960" latitude="-18.943888" location="Parc National Andasibe" longLatPrecision="21" longitude="48.4175" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
<specimenCount id="AFF89A95FFE0FFD2FAEB0BC2FAA4DB0F" box="[1345,1393,998,1025]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
; same data as for preceding; general collecting at night in rainforest; AToL DNA voucher: ARACG045; BLF7994; CASENT_9005657;
<collectionCode id="DFEFC9D9FFE0FFD2FD9E0C0BFDB8DB47" box="[564,621,1071,1097]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="24" pageNumber="25" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFE0FFD2FD2C0C0BFAEBDB47" box="[646,1342,1071,1098]" collectingDate="2003-01-16" collectingDateMax="2003-01-18" collectingDateMin="2003-01-16" collectionCode="CAS" collectorName="C. Griswold &amp; D. Silva &amp; D. Andriamalala" country="Madagascar" elevation="960" latitude="-18.943888" location="Parc National Andasibe" longLatPrecision="21" longitude="48.4175" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
<specimenCount id="AFF89A95FFE0FFD2FD2C0C0BFD60DB44" box="[646,693,1071,1098]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
; same data as for preceding; CASENT_9005261;
<collectionCode id="DFEFC9D9FFE0FFD2FAAB0C0BFAEBDB47" box="[1281,1342,1071,1097]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="24" pageNumber="25" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="B941511CFFE0FFD2FF170C56FE2CDB82" blockId="24.[189,505,1138,1164]" box="[189,505,1138,1164]" pageId="24" pageNumber="25">
<heading id="E209E670FFE0FFD2FF170C56FE2CDB82" bold="true" box="[189,505,1138,1164]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="8B8A8D0EFFE0FFD2FF170C56FE2CDB82" bold="true" box="[189,505,1138,1164]" pageId="24" pageNumber="25">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE0FFD2FF170C86FB66DA27" blockId="24.[189,1399,1184,1322]" pageId="24" pageNumber="25">
<materialsCitation id="09965B41FFE0FFD2FF170C86FCCEDA0B" collectingDate="2015-01-11" collectingDateMax="2015-01-14" collectingDateMin="2015-01-11" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="1385" latitude="-19.715721" location="Ankadivory" longLatPrecision="1" longitude="47.82084" municipality="Tsinjoarivo forest" pageId="24" pageNumber="25" specimenCount="2" specimenCount-male="2" stateProvince="Antananarivo">
<collectingCountry id="C1E9118CFFE0FFD2FF170C86FE5BDBB2" box="[189,398,1186,1212]" name="Madagascar" pageId="24" pageNumber="25">MADAGASCAR</collectingCountry>
–
<emphasis id="8B8A8D0EFFE0FFD2FE000C85FD1BDBB5" bold="true" box="[426,718,1185,1211]" pageId="24" pageNumber="25">
<collectingRegion id="7B3A9FFEFFE0FFD2FE000C85FD1BDBB5" box="[426,718,1185,1211]" country="Madagascar" name="Antananarivo" pageId="24" pageNumber="25">Antananarivo Province</collectingRegion>
</emphasis>
•
<specimenCount id="AFF89A95FFE0FFD2FD4C0C86FCFEDBB2" box="[742,811,1186,1212]" count="2" pageId="24" pageNumber="25" type="male">2 ♂♂</specimenCount>
;
<collectingMunicipality id="5925CB66FFE0FFD2FC9D0C86FBDCDBB2" box="[823,1033,1186,1212]" pageId="24" pageNumber="25">Tsinjoarivo forest</collectingMunicipality>
,
<location id="BC2107C7FFE0FFD2FBB90C86FB76DBB2" LSID="urn:lsid:plazi:treatment:3157E00AFFEFFFD6FDF60E9FFA8ED8E6:BC2107C7FFE0FFD2FBB90C86FB76DBB2" box="[1043,1187,1186,1212]" country="Madagascar" latitude="-19.715721" longLatPrecision="1" longitude="47.82084" municipality="Tsinjoarivo forest" name="Ankadivory" pageId="24" pageNumber="25" stateProvince="Antananarivo">Ankadivory</location>
;
<geoCoordinate id="DCCA37DBFFE0FFD2FB040C84FAA4DBB2" box="[1198,1393,1184,1212]" degrees="19" direction="south" minutes="42" orientation="latitude" pageId="24" pageNumber="25" precision="1" seconds="56.592" value="-19.715721">19°42′56.592″ S</geoCoordinate>
,
<geoCoordinate id="DCCA37DBFFE0FFD2FF170CE1FE4FDBEF" box="[189,410,1221,1249]" degrees="47" direction="east" minutes="49" orientation="longitude" pageId="24" pageNumber="25" precision="1" seconds="15.0234" value="47.82084">47°49′15.0234″ E</geoCoordinate>
;
<elevation id="32D3B62FFFE0FFD2FE030CE2FD9EDBEE" box="[425,587,1222,1249]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="24" pageNumber="25" unit="m" value="1385.0">
<quantity id="7E06FCF9FFE0FFD2FE030CE2FDD2DBEF" box="[425,519,1222,1249]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="24" pageNumber="25" unit="m" value="1385.0">1385 m</quantity>
a.s.l.
</elevation>
;
<date id="CD4077DCFFE0FFD2FDF60CE2FCFFDBEF" box="[604,810,1222,1249]" pageId="24" pageNumber="25" value="2015-01-11" valueMax="2015-01-14" valueMin="2015-01-11">
<collectingDate id="DD048E34FFE0FFD2FDF60CE2FCFFDBEF" box="[604,810,1222,1249]" pageId="24" pageNumber="25" value="2015-01-11" valueMax="2015-01-14" valueMin="2015-01-11">11–14 Jan. 2015</collectingDate>
</date>
;
<collectorName id="140B34CAFFE0FFD2FC930CE3FC16DBEE" box="[825,963,1222,1249]" pageId="24" pageNumber="25">B.L. Fisher</collectorName>
<emphasis id="8B8A8D0EFFE0FFD2FC640CE3FBD8DBEE" box="[974,1037,1222,1248]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
leg.;
<collectingMethod id="60BF290BFFE0FFD2FBFC0CE2FB31DBEF" box="[1110,1252,1222,1249]" pageId="24" pageNumber="25">pitfall traps</collectingMethod>
in montane rainforest; BLF35447-5; CASENT_9065578;
<collectionCode id="DFEFC9D9FFE0FFD2FD480CCFFCCEDA0B" box="[738,795,1259,1285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="24" pageNumber="25" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFE0FFD2FC9D0CCFFE38DA27" collectingDate="2015-01-11" collectingDateMax="2015-01-14" collectingDateMin="2015-01-11" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="1385" latitude="-19.715721" location="Ankadivory" longLatPrecision="1" longitude="47.82084" municipality="Tsinjoarivo forest" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount id="AFF89A95FFE0FFD2FC9D0CCFFCBDDA0B" box="[823,872,1259,1285]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
; same data as for preceding; BLF35447-20; CASENT_9065583;
<collectionCode id="DFEFC9D9FFE0FFD2FE1E0D2BFE38DA27" box="[436,493,1295,1321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="24" pageNumber="25" type="Museum">CAS</collectionCode>
</materialsCitation>
•
<materialsCitation id="09965B41FFE0FFD2FDAC0D2BFB7ADA27" box="[518,1199,1295,1322]" collectingDate="2015-01-11" collectingDateMax="2015-01-14" collectingDateMin="2015-01-11" collectionCode="CAS" collectorName="B. L. Fisher" country="Madagascar" elevation="1385" latitude="-19.715721" location="Ankadivory" longLatPrecision="1" longitude="47.82084" municipality="Tsinjoarivo forest" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo">
<specimenCount id="AFF89A95FFE0FFD2FDAC0D2BFDE0DA24" box="[518,565,1295,1322]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
; same data as for preceding; CASENT_906558;
<collectionCode id="DFEFC9D9FFE0FFD2FBD80D2BFB7ADA27" box="[1138,1199,1295,1321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="24" pageNumber="25" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFE0FFD6FF170D75FBE6D87B" lastPageId="28" lastPageNumber="29" pageId="24" pageNumber="25" type="description">
<paragraph id="B941511CFFE0FFD2FF170D75FE9BDA65" blockId="24.[189,334,1361,1387]" box="[189,334,1361,1387]" pageId="24" pageNumber="25">
<heading id="E209E670FFE0FFD2FF170D75FE9BDA65" bold="true" box="[189,334,1361,1387]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="8B8A8D0EFFE0FFD2FF170D75FE9BDA65" bold="true" box="[189,334,1361,1387]" pageId="24" pageNumber="25">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE0FFD2FF400DA6FCDDDA92" blockId="24.[189,1160,1409,1473]" box="[234,776,1409,1436]" pageId="24" pageNumber="25">
<emphasis id="8B8A8D0EFFE0FFD2FF400DA6FE48DA92" bold="true" box="[234,413,1410,1436]" pageId="24" pageNumber="25">
Male
<typeStatus id="6645EFBEFFE0FFD2FE9B0DA6FE48DA92" box="[305,413,1410,1436]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
</emphasis>
(CASENT_9016063; MNHN)
</paragraph>
<paragraph id="B941511CFFE0FFD2FF170D82FB5DDACE" blockId="24.[189,1160,1409,1473]" box="[189,1160,1446,1473]" pageId="24" pageNumber="25">MEASUREMENTS. Total length: 7.50. Carapace: length 4.60, width 3.70, height 1.80.</paragraph>
<paragraph id="B941511CFFE0FFD2FF170DCDFD54D9B7" blockId="24.[187,1398,1513,1721]" pageId="24" pageNumber="25">COLORATION (Fig. 8A–B). Carapace pale brownish yellow, darker along margin, with two faint, longitudinal dark bands along margin of pars cephalica; black surrounding all eyes; chelicerae brownish yellow; sternum yellow-white, unmarked; legs brownish yellow, femora, tibiae and metatarsi with faint median and subapical dusky annuli; abdomen pale yellow with darker cardiac mark and dorsolateral dark bands that break up into chevrons posteriorly, sides mottled with dark marks, venter yellow except with darker mottling around spinneret base.</paragraph>
<paragraph id="B941511CFFE0FFD2FF170EC6FC08D9F2" blockId="24.[189,989,1762,1789]" box="[189,989,1762,1789]" pageId="24" pageNumber="25">STERNUM. Oval with slightly sinuous margins; 2.00 wide, 2.50 long.</paragraph>
<paragraph id="B941511CFFE0FFD2FF170F01FDF7D830" blockId="24.[189,546,1828,1855]" box="[189,546,1828,1855]" pageId="24" pageNumber="25">CHILUM. 0.13 high, 0.35 wide.</paragraph>
<paragraph id="B941511CFFE0FFD2FF170F43FCCAD8A8" blockId="24.[188,1399,1895,1958]" pageId="24" pageNumber="25">EYES. ALE 0.33; AME 0.13; PLE 0.15; PME 0.14; ALE–AME 0.05; AME–AME 0.20; PLE–PME 0.13; PME–PME 0.06; MOQ: AW 0.26, PW 0.38, L 0.30.</paragraph>
<paragraph id="B941511CFFE0FFD2FF170FEAFD5AD8E7" blockId="24.[189,655,1998,2025]" box="[189,655,1998,2025]" pageId="24" pageNumber="25">CLYPEUS. 0.10 high, with six long setae.</paragraph>
<paragraph id="B941511CFFE1FFD3FF17092DFD49DE4B" blockId="25.[189,1398,265,325]" pageId="25" pageNumber="26">LEGS. Trochanters deeply notched. Unpaired claw smooth, rudimentary on legs III and IV. Metatarsi lacking subbasal crack. Scopulae absent.</paragraph>
<paragraph id="B941511CFFE1FFD3FF170949FE4FDE89" blockId="25.[189,410,365,391]" box="[189,410,365,391]" pageId="25" pageNumber="26">Leg measurements</paragraph>
<paragraph id="B941511CFFE1FFD3FEA009BDFD21DD53" pageId="25" pageNumber="26">
<table id="CBFEA3BCFFE10035FF6109BCFD21DD50" box="[203,756,408,606]" gridcols="7" gridrows="5" pageId="25" pageNumber="26" rowsContinueIn="25.[203,961,688,874]">
<tr id="07CE535EFFE10035FF6109BCFD21DEBD" box="[203,756,408,435]" gridrow="0" pageId="25" pageNumber="26" rowspan-0="1">
<th id="441F3A22FFE10035FEA009BCFEE8DEBD" box="[266,317,408,435]" gridcol="1" gridrow="0" pageId="25" pageNumber="26">F</th>
<th id="441F3A22FFE10035FEF509BCFE47DEBD" box="[351,402,408,435]" gridcol="2" gridrow="0" pageId="25" pageNumber="26">P</th>
<th id="441F3A22FFE10035FE1E09BCFE32DEBD" box="[436,487,408,435]" gridcol="3" gridrow="0" pageId="25" pageNumber="26">T</th>
<th id="441F3A22FFE10035FDA209BCFDEEDEBD" box="[520,571,408,435]" gridcol="4" gridrow="0" pageId="25" pageNumber="26">Mt</th>
<th id="441F3A22FFE10035FDF709BCFD45DEBD" box="[605,656,408,435]" gridcol="5" gridrow="0" pageId="25" pageNumber="26">t</th>
<th id="441F3A22FFE10035FD1809BCFD21DEBD" box="[690,756,408,435]" gridcol="6" gridrow="0" pageId="25" pageNumber="26">Total</th>
</tr>
<tr id="07CE535EFFE10035FF6109E7FD21DED0" box="[203,756,451,478]" gridrow="1" pageId="25" pageNumber="26">
<th id="441F3A22FFE10035FF6109E7FF3FDED0" box="[203,234,451,478]" gridcol="0" gridrow="1" pageId="25" pageNumber="26">I</th>
<td id="441F3A22FFE10035FEA009E7FEE8DED0" box="[266,317,451,478]" gridcol="1" gridrow="1" pageId="25" pageNumber="26">4.30</td>
<td id="441F3A22FFE10035FEF509E7FE47DED0" box="[351,402,451,478]" gridcol="2" gridrow="1" pageId="25" pageNumber="26">1.80</td>
<td id="441F3A22FFE10035FE1E09E7FE32DED0" box="[436,487,451,478]" gridcol="3" gridrow="1" pageId="25" pageNumber="26">3.90</td>
<td id="441F3A22FFE10035FDA209E7FDEEDED0" box="[520,571,451,478]" gridcol="4" gridrow="1" pageId="25" pageNumber="26">3.50</td>
<td id="441F3A22FFE10035FDF709E7FD45DED0" box="[605,656,451,478]" gridcol="5" gridrow="1" pageId="25" pageNumber="26">2.20</td>
<td id="441F3A22FFE10035FD1809E7FD21DED0" box="[690,756,451,478]" gridcol="6" gridrow="1" pageId="25" pageNumber="26">15.50</td>
</tr>
<tr id="07CE535EFFE10035FF6109CAFD21DD06" box="[203,756,494,520]" gridrow="2" pageId="25" pageNumber="26">
<th id="441F3A22FFE10035FF6109CAFF3FDD06" box="[203,234,494,520]" gridcol="0" gridrow="2" pageId="25" pageNumber="26">II</th>
<td id="441F3A22FFE10035FEA009CAFEE8DD06" box="[266,317,494,520]" gridcol="1" gridrow="2" pageId="25" pageNumber="26">3.80</td>
<td id="441F3A22FFE10035FEF509CAFE47DD06" box="[351,402,494,520]" gridcol="2" gridrow="2" pageId="25" pageNumber="26">1.80</td>
<td id="441F3A22FFE10035FE1E09CAFE32DD06" box="[436,487,494,520]" gridcol="3" gridrow="2" pageId="25" pageNumber="26">3.10</td>
<td id="441F3A22FFE10035FDA209CAFDEEDD06" box="[520,571,494,520]" gridcol="4" gridrow="2" pageId="25" pageNumber="26">3.00</td>
<td id="441F3A22FFE10035FDF709CAFD45DD06" box="[605,656,494,520]" gridcol="5" gridrow="2" pageId="25" pageNumber="26">1.90</td>
<td id="441F3A22FFE10035FD1809CAFD21DD06" box="[690,756,494,520]" gridcol="6" gridrow="2" pageId="25" pageNumber="26">13.60</td>
</tr>
<tr id="07CE535EFFE10035FF610A3CFD21DD3D" box="[203,756,536,563]" gridrow="3" pageId="25" pageNumber="26">
<th id="441F3A22FFE10035FF610A3CFF3FDD3D" box="[203,234,536,563]" gridcol="0" gridrow="3" pageId="25" pageNumber="26">III</th>
<td id="441F3A22FFE10035FEA00A3CFEE8DD3D" box="[266,317,536,563]" gridcol="1" gridrow="3" pageId="25" pageNumber="26">3.50</td>
<td id="441F3A22FFE10035FEF50A3CFE47DD3D" box="[351,402,536,563]" gridcol="2" gridrow="3" pageId="25" pageNumber="26">1.50</td>
<td id="441F3A22FFE10035FE1E0A3CFE32DD3D" box="[436,487,536,563]" gridcol="3" gridrow="3" pageId="25" pageNumber="26">2.60</td>
<td id="441F3A22FFE10035FDA20A3CFDEEDD3D" box="[520,571,536,563]" gridcol="4" gridrow="3" pageId="25" pageNumber="26">3.30</td>
<td id="441F3A22FFE10035FDF70A3CFD45DD3D" box="[605,656,536,563]" gridcol="5" gridrow="3" pageId="25" pageNumber="26">1.80</td>
<td id="441F3A22FFE10035FD180A3CFD21DD3D" box="[690,756,536,563]" gridcol="6" gridrow="3" pageId="25" pageNumber="26">12.70</td>
</tr>
<tr id="07CE535EFFE10035FF610A67FD21DD50" box="[203,756,579,606]" gridrow="4" pageId="25" pageNumber="26">
<th id="441F3A22FFE10035FF610A67FF3FDD50" box="[203,234,579,606]" gridcol="0" gridrow="4" pageId="25" pageNumber="26">IV</th>
<td id="441F3A22FFE10035FEA00A67FEE8DD50" box="[266,317,579,606]" gridcol="1" gridrow="4" pageId="25" pageNumber="26">5.30</td>
<td id="441F3A22FFE10035FEF50A67FE47DD50" box="[351,402,579,606]" gridcol="2" gridrow="4" pageId="25" pageNumber="26">1.80</td>
<td id="441F3A22FFE10035FE1E0A67FE32DD50" box="[436,487,579,606]" gridcol="3" gridrow="4" pageId="25" pageNumber="26">4.00</td>
<td id="441F3A22FFE10035FDA20A67FDEEDD50" box="[520,571,579,606]" gridcol="4" gridrow="4" pageId="25" pageNumber="26">5.40</td>
<td id="441F3A22FFE10035FDF70A67FD45DD50" box="[605,656,579,606]" gridcol="5" gridrow="4" pageId="25" pageNumber="26">2.40</td>
<td id="441F3A22FFE10035FD180A67FD21DD50" box="[690,756,579,606]" gridcol="6" gridrow="4" pageId="25" pageNumber="26">18.90</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFE1FFD3FF170AA1FB57DBB1" blockId="25.[189,301,645,671]" lastBlockId="25.[187,1401,912,1215]" pageId="25" pageNumber="26">Spination MALE PALP (Figs 8C–G, 9A–C). Tibia with VTA thumb-shaped, RTA broad, median with sharp, ventroapical point (vRTA), blunt retrodorsal projection (dRTA); cymbium oval with rounded apical extremity, without dorsal patch of chemosensitive setae, with small, sharp, conical retrobasal cymbial extension (CE) extending to tibial apex; subtegulum large, filling most of cymbial concavity, with transverse ridge (SL) that interacts with tegulum; tegulum oval with large, with rounded retrodistal protuberance and conical promedian lobe (TL) that locks with subtegulum; embolus (E) arises from retromargin of tegulum, short, simple, narrow; median apophysis (MA) strongly sclerotized elongate hook curved 270° prolaterad; conductor (C) hyaline, translucent, flattened, fan-shaped, widened at extremity; paraconductor (Pc) finger-like membranous, originating at base of MA and extending two-thirds length of E.</paragraph>
<paragraph id="B941511CFFE1FFD3FEA00A94FC14DC67" pageId="25" pageNumber="26">
<table id="CBFEA3BCFFE10035FF610A94FC14DC64" box="[203,961,688,874]" gridcols="5" gridrows="5" pageId="25" pageNumber="26" rowsContinueFrom="25.[203,756,408,606]">
<tr id="07CE535EFFE10035FF610A94FC14DDC4" box="[203,961,688,714]" gridrow="0" pageId="25" pageNumber="26" rowspan-0="1">
<th id="441F3A22FFE10035FEA00A94FEBBDDC4" box="[266,366,688,714]" gridcol="1" gridrow="0" pageId="25" pageNumber="26">F</th>
<th id="441F3A22FFE10035FE390A94FE0CDDC4" box="[403,473,688,714]" gridcol="2" gridrow="0" pageId="25" pageNumber="26">P</th>
<th id="441F3A22FFE10035FE570A94FD7BDDC4" box="[509,686,688,714]" gridcol="3" gridrow="0" pageId="25" pageNumber="26">T</th>
<th id="441F3A22FFE10035FD750A94FC14DDC4" box="[735,961,688,714]" gridcol="4" gridrow="0" pageId="25" pageNumber="26">Mt</th>
</tr>
<tr id="07CE535EFFE10035FF610AFCFC14DDFD" box="[203,961,728,755]" gridrow="1" pageId="25" pageNumber="26">
<th id="441F3A22FFE10035FF610AFCFF3FDDFD" box="[203,234,728,755]" gridcol="0" gridrow="1" pageId="25" pageNumber="26">I</th>
<td id="441F3A22FFE10035FEA00AFCFEBBDDFD" box="[266,366,728,755]" gridcol="1" gridrow="1" pageId="25" pageNumber="26">pl1d3rl1</td>
<td id="441F3A22FFE10035FE390AFCFE0CDDFD" box="[403,473,728,755]" gridcol="2" gridrow="1" pageId="25" pageNumber="26">–</td>
<td id="441F3A22FFE10035FE570AFCFD7BDDFD" box="[509,686,728,755]" gridcol="3" gridrow="1" pageId="25" pageNumber="26">pl2rl1v2-2-2-2</td>
<td id="441F3A22FFE10035FD750AFCFC14DDFD" box="[735,961,728,755]" gridcol="4" gridrow="1" pageId="25" pageNumber="26">pl1-2v2-2-2</td>
</tr>
<tr id="07CE535EFFE10035FF610B24FC14DC14" box="[203,961,768,794]" gridrow="2" pageId="25" pageNumber="26">
<th id="441F3A22FFE10035FF610B24FF3FDC14" box="[203,234,768,794]" gridcol="0" gridrow="2" pageId="25" pageNumber="26">II</th>
<td id="441F3A22FFE10035FEA00B24FEBBDC14" box="[266,366,768,794]" gridcol="1" gridrow="2" pageId="25" pageNumber="26">pl2d3rl2</td>
<td id="441F3A22FFE10035FE390B24FE0CDC14" box="[403,473,768,794]" gridcol="2" gridrow="2" pageId="25" pageNumber="26">–</td>
<td id="441F3A22FFE10035FE570B24FD7BDC14" box="[509,686,768,794]" gridcol="3" gridrow="2" pageId="25" pageNumber="26">pl2rl2v2-2-2-2</td>
<td id="441F3A22FFE10035FD750B24FC14DC14" box="[735,961,768,794]" gridcol="4" gridrow="2" pageId="25" pageNumber="26">pl1-2v2-2-2</td>
</tr>
<tr id="07CE535EFFE10035FF610B0CFC14DC4C" box="[203,961,808,834]" gridrow="3" pageId="25" pageNumber="26">
<th id="441F3A22FFE10035FF610B0CFF3FDC4C" box="[203,234,808,834]" gridcol="0" gridrow="3" pageId="25" pageNumber="26">III</th>
<td id="441F3A22FFE10035FEA00B0CFEBBDC4C" box="[266,366,808,834]" gridcol="1" gridrow="3" pageId="25" pageNumber="26">pl3d3rl2</td>
<td id="441F3A22FFE10035FE390B0CFE0CDC4C" box="[403,473,808,834]" gridcol="2" gridrow="3" pageId="25" pageNumber="26">pl1rl1</td>
<td id="441F3A22FFE10035FE570B0CFD7BDC4C" box="[509,686,808,834]" gridcol="3" gridrow="3" pageId="25" pageNumber="26">pl2rl2v2-2-2</td>
<td id="441F3A22FFE10035FD750B0CFC14DC4C" box="[735,961,808,834]" gridcol="4" gridrow="3" pageId="25" pageNumber="26">pl2rl2v2-2-2</td>
</tr>
<tr id="07CE535EFFE10035FF610B6BFC14DC64" box="[203,961,847,874]" gridrow="4" pageId="25" pageNumber="26">
<th id="441F3A22FFE10035FF610B6BFF3FDC64" box="[203,234,847,874]" gridcol="0" gridrow="4" pageId="25" pageNumber="26">IV</th>
<td id="441F3A22FFE10035FEA00B6BFEBBDC64" box="[266,366,847,874]" gridcol="1" gridrow="4" pageId="25" pageNumber="26">pl3d3rl2</td>
<td id="441F3A22FFE10035FE390B6BFE0CDC64" box="[403,473,847,874]" gridcol="2" gridrow="4" pageId="25" pageNumber="26">rl1</td>
<td id="441F3A22FFE10035FE570B6BFD7BDC64" box="[509,686,847,874]" gridcol="3" gridrow="4" pageId="25" pageNumber="26">pl2d1rl2v2-2-2</td>
<td id="441F3A22FFE10035FD750B6BFC14DC64" box="[735,961,847,874]" gridcol="4" gridrow="4" pageId="25" pageNumber="26">pl4d3rl2v1-2-1-2-2</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFE1FFD3FF400CC2FCDDDA0E" blockId="25.[189,1175,1254,1315]" box="[234,776,1254,1280]" pageId="25" pageNumber="26">
<emphasis id="8B8A8D0EFFE1FFD3FF400CC2FE68DA0E" bold="true" box="[234,445,1254,1280]" pageId="25" pageNumber="26">
Female
<typeStatus id="6645EFBEFFE1FFD3FEE70CC2FE68DA0E" box="[333,445,1254,1280]" pageId="25" pageNumber="26" type="paratype">paratype</typeStatus>
</emphasis>
(CASENT_9018308; CAS)
</paragraph>
<paragraph id="B941511CFFE1FFD3FF170D2CFB42DA2C" blockId="25.[189,1175,1254,1315]" box="[189,1175,1288,1315]" pageId="25" pageNumber="26">MEASUREMENTS. Total length: 10.20. Carapace: length 5.00, width 3.60, height 1.80.</paragraph>
<paragraph id="B941511CFFE1FFD3FF170D6DFCB9DA88" blockId="25.[187,1399,1353,1414]" pageId="25" pageNumber="26">COLOUR.As in male except contrast between yellow-white background and dusky markings much bolder, especially annuli on posterior femora, tibiae and metatarsi.</paragraph>
<paragraph id="B941511CFFE1FFD3FF170D89FC6FDAC8" blockId="25.[189,954,1452,1479]" box="[189,954,1452,1479]" pageId="25" pageNumber="26">STERNUM. Oval with slightly sinuous sides; 2.20 wide, 2.40 long.</paragraph>
<paragraph id="B941511CFFE1FFD3FF170DC9FDF7D909" blockId="25.[189,546,1517,1544]" box="[189,546,1517,1544]" pageId="25" pageNumber="26">CHILUM. 0.13 high, 0.35 wide.</paragraph>
<paragraph id="B941511CFFE1FFD3FF170E0AFCCAD963" blockId="25.[188,1399,1582,1645]" pageId="25" pageNumber="26">EYES. ALE 0.18; AME 0.13; PLE 0.18; PME 0.18; ALE–AME 0.08; AME–AME 0.15; PLE–PME 0.18; PME–PME 0.16; MOQ: AW 0.25, PW 0.38, L 0.38.</paragraph>
<paragraph id="B941511CFFE1FFD3FF170E91FDDBD8E7" blockId="25.[187,1399,1717,2025]" pageId="25" pageNumber="26">
<emphasis id="8B8A8D0EFFE1FFD3FF170E91FED7D9C1" bold="true" box="[189,258,1717,1743]" pageId="25" pageNumber="26">Fig. 8</emphasis>
(on next page).
<taxonomicName id="7EFE2A9FFFE1FFD3FE120E91FDA0D9C1" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[440,629,1717,1743]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="milloti" status="gen. nov. et sp. nov.">
<emphasis id="8B8A8D0EFFE1FFD3FE120E91FDA0D9C1" box="[440,629,1717,1743]" italics="true" pageId="25" pageNumber="26">Tabiboka milloti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFE1FFD3FDD70E92FC8AD9DE" box="[637,863,1718,1744]" pageId="25" pageNumber="26" rank="species">gen. nov. et sp. nov.</taxonomicNameLabel>
male
<typeStatus id="6645EFBEFFE1FFD3FC080E91FBDDD9C1" box="[930,1032,1717,1743]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
(CASENT_9016063; MNHN).
<emphasis id="8B8A8D0EFFE1FFD3FF110EFCFF05D9FC" bold="true" box="[187,208,1752,1778]" pageId="25" pageNumber="26">A</emphasis>
. Habitus, dorsal view.
<emphasis id="8B8A8D0EFFE1FFD3FE460EFCFE2BD9FC" bold="true" box="[492,510,1752,1778]" pageId="25" pageNumber="26">B</emphasis>
. Carapace, dorsal view.
<emphasis id="8B8A8D0EFFE1FFD3FC800EFCFCEBD9FC" bold="true" box="[810,830,1752,1778]" pageId="25" pageNumber="26">C</emphasis>
. SEM of right male palp (mirrored), prolateral view.
<emphasis id="8B8A8D0EFFE1FFD3FEAE0ED8FECCD818" bold="true" box="[260,281,1788,1814]" pageId="25" pageNumber="26">D</emphasis>
. Idem, ventral view.
<emphasis id="8B8A8D0EFFE1FFD3FDBF0ED8FDFDD818" bold="true" box="[533,552,1788,1814]" pageId="25" pageNumber="26">E</emphasis>
. Idem, retrolateral view.
<emphasis id="8B8A8D0EFFE1FFD3FCF90ED8FCB1D818" bold="true" box="[851,868,1788,1814]" pageId="25" pageNumber="26">F</emphasis>
. Idem, detail of bulb, ventral view.
<emphasis id="8B8A8D0EFFE1FFD3FAA70EDFFAF1D81B" bold="true" box="[1293,1316,1787,1813]" pageId="25" pageNumber="26">G</emphasis>
. Idem, detail of the RTA. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; MA = median apophysis; Pc = paraconductor; dRTA = retrodorsal tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars: A–B = no scales provided (note: carapace length =
<quantity id="7E06FCF9FFE1FFD3FAA10FACFABFD8AC" box="[1291,1386,1928,1954]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="25" pageNumber="26" unit="mm" value="4.6">4.6 mm</quantity>
); C–E =
<quantity id="7E06FCF9FFE1FFD3FEBB0F8FFEBBD8C8" box="[273,366,1963,1990]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="0.5">0.5 mm</quantity>
; F–G =
<quantity id="7E06FCF9FFE1FFD3FE650F88FDF8D8C8" box="[463,557,1964,1990]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="mm" value="0.1">0.1 mm</quantity>
. Stacked microphotographs by Hannah Wood; SEM images by Charles Griswold and Darrell Ubick.
</paragraph>
<caption id="ED810194FFE3FFD1FF170E1FFE20D842" ID-DOI="http://doi.org/10.5281/zenodo.14001810" ID-Zenodo-Dep="14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,231,1595,1621]" targetBox="[260,1379,284,1517]" targetPageId="27">
<paragraph id="B941511CFFE3FFD1FF170E1FFE20D842" blockId="27.[187,1399,1595,1869]" pageId="27" pageNumber="28">
<emphasis id="8B8A8D0EFFE3FFD1FF170E1FFED2D95B" bold="true" box="[189,263,1595,1621]" pageId="27" pageNumber="28">Fig. 9.</emphasis>
<taxonomicName id="7EFE2A9FFFE3FFD1FEA60E18FE13D958" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[268,454,1596,1622]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="milloti" status="gen. nov. et sp. nov.">
<emphasis id="8B8A8D0EFFE3FFD1FEA60E18FE13D958" box="[268,454,1596,1622]" italics="true" pageId="27" pageNumber="28">Tabiboka milloti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFE3FFD1FE610E18FD72D958" box="[459,679,1596,1622]" pageId="27" pageNumber="28" rank="species">gen. nov. et sp. nov.</taxonomicNameLabel>
genitalia drawings.
<emphasis id="8B8A8D0EFFE3FFD1FC2D0E1FFC00D958" bold="true" box="[903,981,1595,1622]" pageId="27" pageNumber="28">A–C =</emphasis>
male holotype (CASENT_9016063; MNHN).
<emphasis id="8B8A8D0EFFE3FFD1FE800E7BFEA2D977" bold="true" box="[298,375,1631,1657]" pageId="27" pageNumber="28">D–E =</emphasis>
female paratype (CASENT_9018308; CAS).
<emphasis id="8B8A8D0EFFE3FFD1FCD20E7BFC58D977" bold="true" box="[888,909,1631,1657]" pageId="27" pageNumber="28">A</emphasis>
. Left palp, prolateral view.
<emphasis id="8B8A8D0EFFE3FFD1FB150E7BFB07D977" bold="true" box="[1215,1234,1631,1657]" pageId="27" pageNumber="28">B</emphasis>
. Idem, ventral view.
<emphasis id="8B8A8D0EFFE3FFD1FEAB0EA6FEC3D992" bold="true" box="[257,278,1666,1692]" pageId="27" pageNumber="28">C</emphasis>
. Idem, retrolateral view.
<emphasis id="8B8A8D0EFFE3FFD1FD9C0EA6FD9ED992" bold="true" box="[566,587,1666,1692]" pageId="27" pageNumber="28">D</emphasis>
. Epigyne, ventral view.
<emphasis id="8B8A8D0EFFE3FFD1FCC80EA6FCA3D992" bold="true" box="[866,886,1666,1692]" pageId="27" pageNumber="28">E</emphasis>
. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold.
</paragraph>
</caption>
<paragraph id="B941511CFFE4FFD6FF17092DFD4CDE2D" blockId="28.[189,665,265,291]" box="[189,665,265,291]" pageId="28" pageNumber="29">CLYPEUS. 0.20 high, with five long setae.</paragraph>
<paragraph id="B941511CFFE4FFD6FF170968FE19DE68" blockId="28.[189,460,332,358]" box="[189,460,332,358]" pageId="28" pageNumber="29">LEGS. Scopulae absent.</paragraph>
<paragraph id="B941511CFFE4FFD6FF1709B4FE4FDEA4" blockId="28.[189,410,400,426]" box="[189,410,400,426]" pageId="28" pageNumber="29">Leg measurements</paragraph>
<paragraph id="B941511CFFE4FFD6FEA0099AFD21DD8C" pageId="28" pageNumber="29">
<table id="CBFEA3BCFFE40035FF610999FD21DD8C" box="[203,756,445,642]" gridcols="7" gridrows="5" pageId="28" pageNumber="29" rowsContinueIn="28.[203,880,730,916]">
<tr id="07CE535EFFE40035FF610999FD21DED6" box="[203,756,445,472]" gridrow="0" pageId="28" pageNumber="29" rowspan-0="1">
<th id="441F3A22FFE40035FEA00999FEE8DED6" box="[266,317,445,472]" gridcol="1" gridrow="0" pageId="28" pageNumber="29">F</th>
<th id="441F3A22FFE40035FEF50999FE47DED6" box="[351,402,445,472]" gridcol="2" gridrow="0" pageId="28" pageNumber="29">P</th>
<th id="441F3A22FFE40035FE1E0999FE32DED6" box="[436,487,445,472]" gridcol="3" gridrow="0" pageId="28" pageNumber="29">T</th>
<th id="441F3A22FFE40035FDA20999FDEEDED6" box="[520,571,445,472]" gridcol="4" gridrow="0" pageId="28" pageNumber="29">Mt</th>
<th id="441F3A22FFE40035FDF70999FD45DED6" box="[605,656,445,472]" gridcol="5" gridrow="0" pageId="28" pageNumber="29">t</th>
<th id="441F3A22FFE40035FD180999FD21DED6" box="[690,756,445,472]" gridcol="6" gridrow="0" pageId="28" pageNumber="29">Total</th>
</tr>
<tr id="07CE535EFFE40035FF6109CCFD21DD0D" box="[203,756,488,515]" gridrow="1" pageId="28" pageNumber="29">
<th id="441F3A22FFE40035FF6109CCFF3FDD0D" box="[203,234,488,515]" gridcol="0" gridrow="1" pageId="28" pageNumber="29">I</th>
<td id="441F3A22FFE40035FEA009CCFEE8DD0D" box="[266,317,488,515]" gridcol="1" gridrow="1" pageId="28" pageNumber="29">3.50</td>
<td id="441F3A22FFE40035FEF509CCFE47DD0D" box="[351,402,488,515]" gridcol="2" gridrow="1" pageId="28" pageNumber="29">1.70</td>
<td id="441F3A22FFE40035FE1E09CCFE32DD0D" box="[436,487,488,515]" gridcol="3" gridrow="1" pageId="28" pageNumber="29">3.00</td>
<td id="441F3A22FFE40035FDA209CCFDEEDD0D" box="[520,571,488,515]" gridcol="4" gridrow="1" pageId="28" pageNumber="29">2.50</td>
<td id="441F3A22FFE40035FDF709CCFD45DD0D" box="[605,656,488,515]" gridcol="5" gridrow="1" pageId="28" pageNumber="29">1.40</td>
<td id="441F3A22FFE40035FD1809CCFD21DD0D" box="[690,756,488,515]" gridcol="6" gridrow="1" pageId="28" pageNumber="29">12.10</td>
</tr>
<tr id="07CE535EFFE40035FF610A36FD21DD23" box="[203,756,530,557]" gridrow="2" pageId="28" pageNumber="29">
<th id="441F3A22FFE40035FF610A36FF3FDD23" box="[203,234,530,557]" gridcol="0" gridrow="2" pageId="28" pageNumber="29">II</th>
<td id="441F3A22FFE40035FEA00A36FEE8DD23" box="[266,317,530,557]" gridcol="1" gridrow="2" pageId="28" pageNumber="29">3.20</td>
<td id="441F3A22FFE40035FEF50A36FE47DD23" box="[351,402,530,557]" gridcol="2" gridrow="2" pageId="28" pageNumber="29">1.70</td>
<td id="441F3A22FFE40035FE1E0A36FE32DD23" box="[436,487,530,557]" gridcol="3" gridrow="2" pageId="28" pageNumber="29">2.50</td>
<td id="441F3A22FFE40035FDA20A36FDEEDD23" box="[520,571,530,557]" gridcol="4" gridrow="2" pageId="28" pageNumber="29">2.30</td>
<td id="441F3A22FFE40035FDF70A36FD45DD23" box="[605,656,530,557]" gridcol="5" gridrow="2" pageId="28" pageNumber="29">1.50</td>
<td id="441F3A22FFE40035FD180A36FD21DD23" box="[690,756,530,557]" gridcol="6" gridrow="2" pageId="28" pageNumber="29">11.20</td>
</tr>
<tr id="07CE535EFFE40035FF610A19FD21DD56" box="[203,756,573,600]" gridrow="3" pageId="28" pageNumber="29">
<th id="441F3A22FFE40035FF610A19FF3FDD56" box="[203,234,573,600]" gridcol="0" gridrow="3" pageId="28" pageNumber="29">III</th>
<td id="441F3A22FFE40035FEA00A19FEE8DD56" box="[266,317,573,600]" gridcol="1" gridrow="3" pageId="28" pageNumber="29">3.20</td>
<td id="441F3A22FFE40035FEF50A19FE47DD56" box="[351,402,573,600]" gridcol="2" gridrow="3" pageId="28" pageNumber="29">1.30</td>
<td id="441F3A22FFE40035FE1E0A19FE32DD56" box="[436,487,573,600]" gridcol="3" gridrow="3" pageId="28" pageNumber="29">2.20</td>
<td id="441F3A22FFE40035FDA20A19FDEEDD56" box="[520,571,573,600]" gridcol="4" gridrow="3" pageId="28" pageNumber="29">2.70</td>
<td id="441F3A22FFE40035FDF70A19FD45DD56" box="[605,656,573,600]" gridcol="5" gridrow="3" pageId="28" pageNumber="29">1.40</td>
<td id="441F3A22FFE40035FD180A19FD21DD56" box="[690,756,573,600]" gridcol="6" gridrow="3" pageId="28" pageNumber="29">10.80</td>
</tr>
<tr id="07CE535EFFE40035FF610A43FD21DD8C" box="[203,756,615,642]" gridrow="4" pageId="28" pageNumber="29">
<th id="441F3A22FFE40035FF610A43FF3FDD8C" box="[203,234,615,642]" gridcol="0" gridrow="4" pageId="28" pageNumber="29">IV</th>
<td id="441F3A22FFE40035FEA00A43FEE8DD8C" box="[266,317,615,642]" gridcol="1" gridrow="4" pageId="28" pageNumber="29">4.50</td>
<td id="441F3A22FFE40035FEF50A43FE47DD8C" box="[351,402,615,642]" gridcol="2" gridrow="4" pageId="28" pageNumber="29">1.60</td>
<td id="441F3A22FFE40035FE1E0A43FE32DD8C" box="[436,487,615,642]" gridcol="3" gridrow="4" pageId="28" pageNumber="29">3.40</td>
<td id="441F3A22FFE40035FDA20A43FDEEDD8C" box="[520,571,615,642]" gridcol="4" gridrow="4" pageId="28" pageNumber="29">4.50</td>
<td id="441F3A22FFE40035FDF70A43FD45DD8C" box="[605,656,615,642]" gridcol="5" gridrow="4" pageId="28" pageNumber="29">1.90</td>
<td id="441F3A22FFE40035FD180A43FD21DD8C" box="[690,756,615,642]" gridcol="6" gridrow="4" pageId="28" pageNumber="29">15.90</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFE4FFD6FF170A88FEDDDB11" blockId="28.[189,301,684,710]" lastBlockId="28.[189,1398,956,1055]" pageId="28" pageNumber="29">
Spination GENITALIA. Epigyne (
<figureCitation id="21C54D99FFE4FFD6FE1D0B98FDC6DCD8" box="[439,531,956,982]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.A–C = male holotype (CASENT_9016063; MNHN). D–E = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="28" pageNumber="29">Fig. 9D</figureCitation>
) median lobe (ML) heart-shaped, sides rounded, broad anteriorly, anterior width greater than one-half total width. Endogyne (
<figureCitation id="21C54D99FFE4FFD6FCBC0BC5FCBADCF5" box="[790,879,993,1019]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.A–C = male holotype (CASENT_9016063; MNHN). D–E = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="28" pageNumber="29">Fig. 9E</figureCitation>
) with each spermatheca narrow, longer than broad.
</paragraph>
<paragraph id="B941511CFFE4FFD6FEA00AFEFCA5DC9A" pageId="28" pageNumber="29">
<table id="CBFEA3BCFFE40035FF610AFEFCA5DC9A" box="[203,880,730,916]" gridcols="5" gridrows="5" pageId="28" pageNumber="29" rowsContinueFrom="28.[203,756,445,642]">
<tr id="07CE535EFFE40035FF610AFEFCA5DDFA" box="[203,880,730,756]" gridrow="0" pageId="28" pageNumber="29" rowspan-0="1">
<th id="441F3A22FFE40035FEA00AFEFEB8DDFA" box="[266,365,730,756]" gridcol="1" gridrow="0" pageId="28" pageNumber="29">F</th>
<th id="441F3A22FFE40035FE390AFEFE76DDFA" box="[403,419,730,756]" gridcol="2" gridrow="0" pageId="28" pageNumber="29">P</th>
<th id="441F3A22FFE40035FE680AFEFDA1DDFA" box="[450,628,730,756]" gridcol="3" gridrow="0" pageId="28" pageNumber="29">T</th>
<th id="441F3A22FFE40035FD0C0AFEFCA5DDFA" box="[678,880,730,756]" gridcol="4" gridrow="0" pageId="28" pageNumber="29">Mt</th>
</tr>
<tr id="07CE535EFFE40035FF610B25FCA5DC12" box="[203,880,769,796]" gridrow="1" pageId="28" pageNumber="29">
<th id="441F3A22FFE40035FF610B25FF3FDC12" box="[203,234,769,796]" gridcol="0" gridrow="1" pageId="28" pageNumber="29">I</th>
<td id="441F3A22FFE40035FEA00B25FEB8DC12" box="[266,365,769,796]" gridcol="1" gridrow="1" pageId="28" pageNumber="29">pl1d1</td>
<td id="441F3A22FFE40035FE390B25FE76DC12" box="[403,419,769,796]" gridcol="2" gridrow="1" pageId="28" pageNumber="29">–</td>
<td id="441F3A22FFE40035FE680B25FDA1DC12" box="[450,628,769,796]" gridcol="3" gridrow="1" pageId="28" pageNumber="29">v2-2-2-2</td>
<td id="441F3A22FFE40035FD0C0B25FCA5DC12" box="[678,880,769,796]" gridcol="4" gridrow="1" pageId="28" pageNumber="29">v2-2-2</td>
</tr>
<tr id="07CE535EFFE40035FF610B0DFCA5DC4A" box="[203,880,809,836]" gridrow="2" pageId="28" pageNumber="29">
<th id="441F3A22FFE40035FF610B0DFF3FDC4A" box="[203,234,809,836]" gridcol="0" gridrow="2" pageId="28" pageNumber="29">II</th>
<td id="441F3A22FFE40035FEA00B0DFEB8DC4A" box="[266,365,809,836]" gridcol="1" gridrow="2" pageId="28" pageNumber="29">pl1d1</td>
<td id="441F3A22FFE40035FE390B0DFE76DC4A" box="[403,419,809,836]" gridcol="2" gridrow="2" pageId="28" pageNumber="29">–</td>
<td id="441F3A22FFE40035FE680B0DFDA1DC4A" box="[450,628,809,836]" gridcol="3" gridrow="2" pageId="28" pageNumber="29">v2-2-2-2</td>
<td id="441F3A22FFE40035FD0C0B0DFCA5DC4A" box="[678,880,809,836]" gridcol="4" gridrow="2" pageId="28" pageNumber="29">v2-2-2</td>
</tr>
<tr id="07CE535EFFE40035FF610B75FCA5DC62" box="[203,880,849,876]" gridrow="3" pageId="28" pageNumber="29">
<th id="441F3A22FFE40035FF610B75FF3FDC62" box="[203,234,849,876]" gridcol="0" gridrow="3" pageId="28" pageNumber="29">III</th>
<td id="441F3A22FFE40035FEA00B75FEB8DC62" box="[266,365,849,876]" gridcol="1" gridrow="3" pageId="28" pageNumber="29">pl1d3rl1</td>
<td id="441F3A22FFE40035FE390B75FE76DC62" box="[403,419,849,876]" gridcol="2" gridrow="3" pageId="28" pageNumber="29">–</td>
<td id="441F3A22FFE40035FE680B75FDA1DC62" box="[450,628,849,876]" gridcol="3" gridrow="3" pageId="28" pageNumber="29">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFE40035FD0C0B75FCA5DC62" box="[678,880,849,876]" gridcol="4" gridrow="3" pageId="28" pageNumber="29">pl4d2rl3v2-2-2</td>
</tr>
<tr id="07CE535EFFE40035FF610B5DFCA5DC9A" box="[203,880,889,916]" gridrow="4" pageId="28" pageNumber="29">
<th id="441F3A22FFE40035FF610B5DFF3FDC9A" box="[203,234,889,916]" gridcol="0" gridrow="4" pageId="28" pageNumber="29">IV</th>
<td id="441F3A22FFE40035FEA00B5DFEB8DC9A" box="[266,365,889,916]" gridcol="1" gridrow="4" pageId="28" pageNumber="29">pl1d3rl1</td>
<td id="441F3A22FFE40035FE390B5DFE76DC9A" box="[403,419,889,916]" gridcol="2" gridrow="4" pageId="28" pageNumber="29">–</td>
<td id="441F3A22FFE40035FE680B5DFDA1DC9A" box="[450,628,889,916]" gridcol="3" gridrow="4" pageId="28" pageNumber="29">pl2d2rl2v2-2-2</td>
<td id="441F3A22FFE40035FD0C0B5DFCA5DC9A" box="[678,880,889,916]" gridcol="4" gridrow="4" pageId="28" pageNumber="29">pl2d5rl3v2-1-2-2</td>
</tr>
</table>
</paragraph>
<paragraph id="B941511CFFE4FFD6FF170C6CFEE6DB6C" blockId="28.[189,307,1096,1122]" box="[189,307,1096,1122]" pageId="28" pageNumber="29">
<heading id="E209E670FFE4FFD6FF170C6CFEE6DB6C" bold="true" box="[189,307,1096,1122]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="8B8A8D0EFFE4FFD6FF170C6CFEE6DB6C" bold="true" box="[189,307,1096,1122]" pageId="28" pageNumber="29">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE4FFD6FF400C5DFE57DB9D" blockId="28.[189,1399,1145,1539]" box="[234,386,1145,1171]" pageId="28" pageNumber="29">
<emphasis id="8B8A8D0EFFE4FFD6FF400C5DFEFFDB9D" bold="true" box="[234,298,1145,1171]" pageId="28" pageNumber="29">Male</emphasis>
(n = 3)
</paragraph>
<paragraph id="B941511CFFE4FFD6FF170CB9FCDFD90D" blockId="28.[189,1399,1145,1539]" pageId="28" pageNumber="29">
Total length = 6.90–7.50, carapace length/width = 1.21–1.29, carapace height/width = 0.43–0.59, PER/carapace width = 0.23–0.24, PER/OAL = 2.38–2.44, PER/AER = 1.38–1.39, OAL/OQL = 1.13– 1.23, OQP/OQA= 1.38–1.43, clypeus height/AME = 1.14–1.56, cheliceral length/clypeus height = 9.14– 18.00, sternum length/width = 1.05–1.25, palpal coxa length/width = 2.00–2.80, femur I length/carapace width = 1.11–1.12, metatarsus I length/carapace width = 0.95–1.00, femur IV length/carapace width = 1.23–1.43, cymbium length/carapace width = 0.32–0.35, cymbium length/palpal patella length = 1.20– 1.50, cymbium length/palpal tibia length = 1.09–1.26, cymbium length/palpal femur length = 0.63–0.75, palpal tibia length/palpal patella length 1.10–1.38. Males from Tsinjoarivo and Analamazaotra are marked like the
<typeStatus id="6645EFBEFFE4FFD6FE8A0DE7FE5CDAD3" box="[288,393,1475,1501]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
but the contrast between the yellow-white background and dusky markings on the cephalothorax, legs and abdomen is much greater.
</paragraph>
<paragraph id="B941511CFFE4FFD6FF400E0FFE90D94B" blockId="28.[189,869,1579,1643]" box="[234,325,1579,1605]" pageId="28" pageNumber="29">
<emphasis id="8B8A8D0EFFE4FFD6FF400E0FFE90D94B" bold="true" box="[234,325,1579,1605]" pageId="28" pageNumber="29">Female</emphasis>
</paragraph>
<paragraph id="B941511CFFE4FFD6FF170E74FCB0D964" blockId="28.[189,869,1579,1643]" box="[189,869,1616,1643]" pageId="28" pageNumber="29">
Variation unknown; only
<specimenCount id="AFF89A95FFE4FFD6FE420E75FDBFD964" box="[488,618,1616,1643]" count="1" pageId="28" pageNumber="29" type="female">one female</specimenCount>
has been discovered.
</paragraph>
<paragraph id="B941511CFFE4FFD6FF170EB7FE40D9A3" blockId="28.[189,405,1683,1709]" box="[189,405,1683,1709]" pageId="28" pageNumber="29">
<heading id="E209E670FFE4FFD6FF170EB7FE40D9A3" bold="true" box="[189,405,1683,1709]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="8B8A8D0EFFE4FFD6FF170EB7FE40D9A3" bold="true" box="[189,405,1683,1709]" pageId="28" pageNumber="29">Note on synonym</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE4FFD6FF170EE7FB10D80C" blockId="28.[189,1399,1731,1795]" pageId="28" pageNumber="29">
A specimen of this species (CASENT_9005657) was used as an exemplar in a previously published molecular phylogeny: it was listed as “
<taxonomicName id="7EFE2A9FFFE4FFD6FD2F0ECCFCE9D80D" box="[645,828,1768,1795]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8B8A8D0EFFE4FFD6FD2F0ECCFCC6D80C" box="[645,787,1768,1794]" italics="true" pageId="28" pageNumber="29">Zorodictyna</emphasis>
sp.
</taxonomicName>
CG45” by
<bibRefCitation id="DD6F2CEDFFE4FFD6FC680ECCFB14D80C" author="Wheeler W. C. &amp; Coddington J. A. &amp; Crowley L. M. &amp; Dimitrov D. &amp; Goloboff P. A. &amp; Griswold C. E. &amp; Hormiga G. &amp; Prendini L. &amp; Ramirez M. J. &amp; Sierwald P. &amp; Almeida-Silva L. M. &amp; Alvarez-Padilla F. &amp; Arnedo M. A. &amp; Benavides L. R. &amp; Benjamin S. P. &amp; Bond J. E. &amp; Grismado C. J. &amp; Hasan E. &amp; Hedin M. &amp; Izquierdo M. A. &amp; Labarque F. M. &amp; Ledford J. &amp; Lopardo L. &amp; Maddison W. P. &amp; Miller J. A. &amp; Piacentini L. N. &amp; Platnick N. I. &amp; Polotow D. &amp; Silva-Davila D. &amp; Scharff N. &amp; Szuts T. &amp; Ubick D. &amp; Vink C. &amp; Wood H. M. &amp; Zhang J. X." box="[962,1217,1768,1794]" pageId="28" pageNumber="29" pagination="576 - 616" refId="ref39136" refString="Wheeler W. C., Coddington J. A., Crowley L. M., Dimitrov D., Goloboff P. A., Griswold C. E., Hormiga G., Prendini L., Ramirez M. J., Sierwald P., Almeida-Silva L. M., Alvarez-Padilla F., Arnedo M. A., Benavides L. R., Benjamin S. P., Bond J. E., Grismado C. J., Hasan E., Hedin M., Izquierdo M. A., Labarque F. M., Ledford J., Lopardo L., Maddison W. P., Miller J. A., Piacentini L. N., Platnick N. I., Polotow D., Silva-Davila D., Scharff N., Szuts T., Ubick D., Vink C., Wood H. M. &amp; Zhang J. X. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. https: // doi. org / 10.1111 / cla. 12182" type="journal article" year="2017">
Wheeler
<emphasis id="8B8A8D0EFFE4FFD6FB870ECDFBBDD80C" box="[1069,1128,1768,1794]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
(2017)
</bibRefCitation>
.
</paragraph>
<paragraph id="B941511CFFE4FFD6FF170F0FFEABD84B" blockId="28.[189,382,1835,1861]" box="[189,382,1835,1861]" pageId="28" pageNumber="29">
<heading id="E209E670FFE4FFD6FF170F0FFEABD84B" bold="true" box="[189,382,1835,1861]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="8B8A8D0EFFE4FFD6FF170F0FFEABD84B" bold="true" box="[189,382,1835,1861]" pageId="28" pageNumber="29">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE4FFD6FF170F7FFBE6D87B" blockId="28.[189,1075,1883,1910]" box="[189,1075,1883,1910]" pageId="28" pageNumber="29">
The species
<taxonomicName id="7EFE2A9FFFE4FFD6FEE40F7FFDDBD87B" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[334,526,1883,1909]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="milloti" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFE4FFD6FEE40F7FFDDBD87B" box="[334,526,1883,1909]" italics="true" pageId="28" pageNumber="29">Tabiboka milloti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFE4FFD6FDBF0F78FD10D878" box="[533,709,1884,1910]" pageId="28" pageNumber="29" rank="species">gen. et sp. nov.</taxonomicNameLabel>
is only known from rainforest.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFE4FFD6FF170FBAFA8ED8E6" box="[189,1371,1950,2025]" pageId="28" pageNumber="29" type="distribution">
<paragraph id="B941511CFFE4FFD6FF170FBAFE83D8B6" blockId="28.[189,342,1950,1976]" box="[189,342,1950,1976]" pageId="28" pageNumber="29">
<heading id="E209E670FFE4FFD6FF170FBAFE83D8B6" bold="true" box="[189,342,1950,1976]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="8B8A8D0EFFE4FFD6FF170FBAFE83D8B6" bold="true" box="[189,342,1950,1976]" pageId="28" pageNumber="29">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFE4FFD6FF170FEAFA8ED8E6" blockId="28.[189,1371,1998,2025]" box="[189,1371,1998,2025]" pageId="28" pageNumber="29">
<taxonomicName id="7EFE2A9FFFE4FFD6FF170FEAFEA8D8E6" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[189,381,1998,2024]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="milloti" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFE4FFD6FF170FEAFEA8D8E6" box="[189,381,1998,2024]" italics="true" pageId="28" pageNumber="29">Tabiboka milloti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFE4FFD6FE2E0FEBFDE0D8E7" box="[388,565,1999,2025]" pageId="28" pageNumber="29" rank="species">gen. et sp. nov.</taxonomicNameLabel>
is known only from mountains in east-central
<collectingCountry id="C1E9118CFFE4FFD6FBF20FEAFB33D8E6" box="[1112,1254,1998,2024]" name="Madagascar" pageId="28" pageNumber="29">Madagascar</collectingCountry>
(
<figureCitation id="21C54D99FFE4FFD6FB5F0FEAFA98D8E6" box="[1269,1357,1998,2024]" captionStart="Fig" captionStartId="63.[189,231,1857,1883]" captionTargetBox="[189,1398,1005,1819]" captionTargetId="figure-208@63.[189,1398,1005,1819]" captionTargetPageId="63" captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)." figureDoi="http://doi.org/10.5281/zenodo.14001851" httpUri="https://zenodo.org/record/14001851/files/figure.png" pageId="28" pageNumber="29">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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