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<mods:title id="E1E4A802A308C55360439E7898D1F4BA">On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar</mods:title>
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<mods:namePart id="4E1688846A97A975E95C0682BE5CCE3C">Henrard, Arnaud</mods:namePart>
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Family
<taxonomicName id="7EFE2A9FFFF2FFC0FDC20999FBF3DED6" ID-CoL="HN5" ID-ENA="2511328" authority="Griswold &amp; Polotow, 2015" authorityName="Griswold &amp; Polotow" authorityYear="2015" box="[616,1062,445,472]" class="Arachnida" family="Udubidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
<emphasis id="8B8A8D0EFFF2FFC0FDC20999FD37DED9" bold="true" box="[616,738,445,471]" pageId="10" pageNumber="11">Udubidae</emphasis>
Griswold &amp;
<bibRefCitation id="DD6F2CEDFFF2FFC0FCD60999FBF3DED6" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[892,1062,445,472]" pageId="10" pageNumber="11" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">Polotow, 2015</bibRefCitation>
</taxonomicName>
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<paragraph id="B941511CFFF2FFC0FF170A25FC55DD12" blockId="10.[189,896,513,540]" box="[189,896,513,540]" pageId="10" pageNumber="11">
<taxonomicName id="7EFE2A9FFFF2FFC0FF170A25FDFEDD15" authority="Griswold &amp; Polotow" authorityName="Griswold &amp; Polotow" authorityYear="2015" box="[189,555,513,539]" class="Arachnida" family="Udubidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Udubidae Griswold &amp; Polotow</taxonomicName>
in
<bibRefCitation id="DD6F2CEDFFF2FFC0FDFA0A25FCA9DD12" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[592,892,513,540]" pageId="10" pageNumber="11" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow
<emphasis id="8B8A8D0EFFF2FFC0FD130A26FD21DD15" box="[697,756,513,539]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
, 2015: 151
</bibRefCitation>
.
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<typeStatus id="6645EFBEFFF2FFC0FF170A60FF2FDD50" box="[189,250,580,606]" pageId="10" pageNumber="11">Type</typeStatus>
genus
</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFF2FFC0FF170A51FD8CDDBA" blockId="10.[189,1399,628,692]" pageId="10" pageNumber="11">
<taxonomicName id="7EFE2A9FFFF2FFC0FF170A51FE3DDD81" authority="Simon, 1880: 343" authorityName="Simon" authorityPageNumber="343" authorityYear="1880" box="[189,488,629,655]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFF2FFC0FF170A51FED8DD81" box="[189,269,629,655]" italics="true" pageId="10" pageNumber="11">Uduba</emphasis>
<bibRefCitation id="DD6F2CEDFFF2FFC0FEBF0A51FE3DDD81" author="Simon E." box="[277,488,629,655]" pageId="10" pageNumber="11" pagination="223 - 351" refId="ref38313" refString="Simon E. 1880. Revision de la famille des Sparassidae (Arachnides). Actes de la Societe Linneenne de Bordeaux 34 (2 / 3 / 4): 223 - 351." type="journal article" year="1880">Simon, 1880: 343</bibRefCitation>
</taxonomicName>
(
<typeStatus id="6645EFBEFFF2FFC0FE520A51FDE1DD81" box="[504,564,629,655]" pageId="10" pageNumber="11">Type</typeStatus>
species by monotypy:
<taxonomicName id="7EFE2A9FFFF2FFC0FCEC0A51FAD3DD80" authority="Vinson, 1863" authorityName="Vinson" authorityYear="1863" box="[838,1286,628,655]" class="Arachnida" family="Sparassidae" genus="Olios" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="8B8A8D0EFFF2FFC0FCEC0A51FBB7DD81" box="[838,1122,629,655]" italics="true" pageId="10" pageNumber="11">Olios madagascariensis</emphasis>
<bibRefCitation id="DD6F2CEDFFF2FFC0FBC30A51FAD3DD80" author="Vinson A." box="[1129,1286,628,655]" pageId="10" pageNumber="11" refId="ref39036" refString="Vinson A. 1863. Araneides des Iles de La Reunion, Maurice et Madagascar. Librairie Classique Eugene Belin, Paris. https: // doi. org / 10.5962 / bhl. title. 125517" type="book" year="1863">Vinson, 1863</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="7EFE2A9FFFF2FFC0FA8D0A51FD80DDBA" authority="(Vinson, 1863))" authorityName="" baseAuthorityName="Vinson" baseAuthorityYear="1863" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="8B8A8D0EFFF2FFC0FA8D0A51FE44DDBA" italics="true" pageId="10" pageNumber="11">Uduba madagascariensis</emphasis>
(
<bibRefCitation id="DD6F2CEDFFF2FFC0FE0A0ABEFD97DDBA" author="Vinson A." box="[416,578,666,692]" pageId="10" pageNumber="11" refId="ref39036" refString="Vinson A. 1863. Araneides des Iles de La Reunion, Maurice et Madagascar. Librairie Classique Eugene Belin, Paris. https: // doi. org / 10.5962 / bhl. title. 125517" type="book" year="1863">Vinson, 1863</bibRefCitation>
))
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFF2FFC1FF170AF9FB71DE78" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="B941511CFFF2FFC0FF170AF9FEE3DDF9" blockId="10.[189,310,733,759]" box="[189,310,733,759]" pageId="10" pageNumber="11">
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<emphasis id="8B8A8D0EFFF2FFC0FF170AF9FEE3DDF9" bold="true" box="[189,310,733,759]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFF2FFC0FF170B29FAA0DA04" blockId="10.[187,1400,781,1290]" pageId="10" pageNumber="11">
<taxonomicName id="7EFE2A9FFFF2FFC0FF170B29FEFADC29" authorityName="Griswold &amp; Polotow" authorityYear="2015" box="[189,303,781,807]" class="Arachnida" family="Udubidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Udubidae</taxonomicName>
are entelegyne RTA-clade spiders with three tarsal claws (
<figureCitation id="21C54D99FFF2FFC0FC780B29FB8ADC26" box="[978,1119,781,808]" captionStart="Fig" captionStartId="49.[192,236,1327,1353]" captionTargetBox="[189,1398,265,1289]" captionTargetId="figure-118@49.[189,1398,265,1289]" captionTargetPageId="49" captionText="Fig. 21. Zorascar pasunepomme gen. et sp. nov., paratype juvenile (BE_RMCA_ARA.Ara.247347; RMCA). A. Tarsal claws of leg 1. B. Tarsal claws of leg 4. C. Tarsal organ of leg 2. D. Tarsal organ and trichobothrium of leg 4. E. Trichobothrium, tarsus 1. Scale bars: A = 50 μm; B = 20 μm; C, E = 5 μm; D = 10 μm. SEM images by Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001841" httpUri="https://zenodo.org/record/14001841/files/figure.png" pageId="10" pageNumber="11">Fig. 21AB</figureCitation>
), multiple rows of tarsal trichobothria of about equal length (
<figureCitation id="21C54D99FFF2FFC0FDCC0B16FD1ADC43" box="[614,719,818,845]" captionStart="Fig" captionStartId="38.[191,233,1722,1748]" captionTargetBox="[191,1399,265,1682]" captionTargetId="figure-19@38.[191,1399,265,1682]" captionTargetPageId="38" captionText="Fig. 13. Zorascar pasunepipe gen. et sp. nov.; A, C, EF = holotype male (BE_RMCA_ARA.Ara.247343; RMCA); B, D = paratype female (CASENT_9016138; CAS). AB. Habitus, dorsal view.CD. Prosoma, dorsal view. E. Habitus, ventral view. F. Tarsus of leg I, retrolateral view. Scale bars: AB, E = 2 mm; CD = 1 mm; F = 0.5 mm. Stacked microphotographs A, C, EF by Arnaud Henrard and Rudy Jocqué; B, D by Hannah Wood." figureDoi="http://doi.org/10.5281/zenodo.14001823" httpUri="https://zenodo.org/record/14001823/files/figure.png" pageId="10" pageNumber="11">Figs 13F</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FD760B17FCFADC43" box="[732,815,819,845]" captionStart="Fig" captionStartId="48.[189,233,1705,1731]" captionTargetBox="[189,1398,265,1665]" captionTargetId="figure-19@48.[189,1398,265,1665]" captionTargetPageId="48" captionText="Fig. 20. Zorascar pasunepomme gen. et sp. nov., paratype juvenile (BE_RMCA_ARA.Ara.247347; RMCA), SEM pictures. A. ALS, caudal view. B. PLS, posterior view. C. PMS, posterior view. D. Right serrula, ventral view. E. Tarsus of leg 1, arrow pointing to the tarsal organ. F. Tarsus of leg 4. Scale bars: AD = 10 μm; EF = 100 μm. SEM images by Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001839" httpUri="https://zenodo.org/record/14001839/files/figure.png" pageId="10" pageNumber="11">20EF</figureCitation>
), eight eyes in 2 nearly straight rows (
<figureCitation id="21C54D99FFF2FFC0FB580B16FAA4DC43" box="[1266,1393,818,845]" captionStart="Fig" captionStartId="16.[189,233,1892,1918]" captionTargetBox="[189,1398,467,1854]" captionTargetId="figure-79@16.[189,1398,467,1854]" captionTargetPageId="16" captionText="Fig. 3. Tabiboka milleri gen. et sp. nov., AB, G = male paratype (CASENT_9016097; CAS); C = male paratype (CASENT_9016098; CAS); DF, H = female paratype (CASENT_9016094; CAS). A, D. Habitus, dorsal view. B, E. Idem, ventral view. C, F. Carapace, dorsal view. GH. Habitus, lateral view. Scale bars = 2 mm. Stacked microphotographs by Hannah Wood." figureDoi="http://doi.org/10.5281/zenodo.14001794" httpUri="https://zenodo.org/record/14001794/files/figure.png" pageId="10" pageNumber="11">Figs 3C, F</figureCitation>
, 8AB, 13CD, 17A, 22A, C, 24A, C), indirect eyes with canoe-shaped tapeta and the cheliceral fang with interior serrula weak to absent. Males may or may not have a subbasal crack or line of autospasy on the leg tibiae (
<figureCitation id="21C54D99FFF2FFC0FE210B86FE20DCB2" box="[395,501,930,956]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="10" pageNumber="11">Fig. 24B</figureCitation>
, arrow;
<bibRefCitation id="DD6F2CEDFFF2FFC0FDFF0B86FCDCDCB2" author="Griswold C. E." box="[597,777,930,956]" pageId="10" pageNumber="11" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: figs 34). All species have a hyaline conductor (C = the “Sierwald conductor” of
<bibRefCitation id="DD6F2CEDFFF2FFC0FDA60BE3FD21DCEC" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[524,756,967,994]" pageId="10" pageNumber="11" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow
<emphasis id="8B8A8D0EFFF2FFC0FDDE0BECFD7BDCEF" box="[628,686,967,993]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2015
</bibRefCitation>
), which is relatively small, transverse, hyaline and fan-shaped, prolaterally extending from narrow base to wide flattened tip (
<figureCitation id="21C54D99FFF2FFC0FC460BC8FBBADB09" box="[1004,1135,1004,1031]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="10" pageNumber="11">Figs 5AC</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FBD00BC8FB68DB08" box="[1146,1213,1004,1030]" captionStart="Fig" captionStartId="19.[188,230,1787,1813]" captionTargetBox="[188,1399,479,1749]" captionTargetId="figure-89@19.[188,1399,479,1749]" captionTargetPageId="19" captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AC = 0.43 mm; DF = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick." figureDoi="http://doi.org/10.5281/zenodo.14001804" httpUri="https://zenodo.org/record/14001804/files/figure.png" pageId="10" pageNumber="11">6DF</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FB620BC8FAC4DB08" box="[1224,1297,1004,1030]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.AC = male holotype (CASENT_9016063; MNHN). DE = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations AC by Jenny Speckels; DE by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="10" pageNumber="11">9AC</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FAB10BC8FAA4DB08" box="[1307,1393,1004,1030]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; AC= male holotype (CASENT_9016117; CAS); DE = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="10" pageNumber="11">11AC</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FF170C35FF3BDB25" box="[189,238,1041,1067]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; AF = male holotype (CASENT_9016117; CAS); GH= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AD = 0.5 mm; EF = 0.15 mm; GH = 0.1 mm. SEM images AF by Charles Griswold and Darrell Ubick, GH by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="10" pageNumber="11">12E</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FF530C35FE9CDB22" box="[249,329,1041,1068]" captionStart="Fig" captionStartId="39.[189,232,1712,1738]" captionTargetBox="[189,1398,459,1674]" captionTargetId="figure-47@39.[189,1398,459,1674]" captionTargetPageId="39" captionText="Fig. 14. Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ARA.Ara.247343; RMCA). A. Male right palp (mirrored), retrolateral view. B. Idem, ventral view. C. Idem, dorsal view. D. Idem, bulbus, retrolateral view. E. Idem, ventral view. F. Idem, prolateral view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AC = 0.5mm; DF = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001825" httpUri="https://zenodo.org/record/14001825/files/figure.png" pageId="10" pageNumber="11">14D, F</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FEF90C35FE7DDB22" box="[339,424,1041,1068]" captionStart="Fig" captionStartId="40.[189,233,1731,1757]" captionTargetBox="[189,1398,478,1693]" captionTargetId="figure-47@40.[189,1398,478,1693]" captionTargetPageId="40" captionText="Fig. 15. Zorascar pasunepipe gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247343; RMCA), micro-CT scans of right male palp (mirrored). A. Retrolateral view. B. Ventral view. C. Prolateral view. D. Bulbus, retrolateral view.E. Bulbus, ventral view.F. Bulbus, apical view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AC = 0.5 mm; DF = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001827" httpUri="https://zenodo.org/record/14001827/files/figure.png" pageId="10" pageNumber="11">15DF</figureCitation>
). Most have an additional tegular process accompanying the embolus that is a narrow membranous or sclerotized extension inserted between the tegulum and the base of the median apophysis (MA), running backward to MA and ending with fattened fan-shaped tip accompanying the embolus: this is the paraconductor (Pc = the “MTP” and “STP” of
<bibRefCitation id="DD6F2CEDFFF2FFC0FC8A0CA4FC05DB94" author="Griswold C. E." box="[800,976,1152,1178]" pageId="10" pageNumber="11" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
) (
<figureCitation id="21C54D99FFF2FFC0FC420CA4FBBEDB95" box="[1000,1131,1152,1179]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="10" pageNumber="11">Figs 5AC</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FBDD0CA4FB6EDB94" box="[1143,1211,1152,1178]" captionStart="Fig" captionStartId="19.[188,230,1787,1813]" captionTargetBox="[188,1399,479,1749]" captionTargetId="figure-89@19.[188,1399,479,1749]" captionTargetPageId="19" captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AC = 0.43 mm; DF = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick." figureDoi="http://doi.org/10.5281/zenodo.14001804" httpUri="https://zenodo.org/record/14001804/files/figure.png" pageId="10" pageNumber="11">6DF</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FB6C0CA4FADADB94" box="[1222,1295,1152,1178]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.AC = male holotype (CASENT_9016063; MNHN). DE = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations AC by Jenny Speckels; DE by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="10" pageNumber="11">9AC</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FAB00CA4FAA5DB94" box="[1306,1392,1152,1178]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; AC= male holotype (CASENT_9016117; CAS); DE = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="10" pageNumber="11">11AC</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FF160C81FF3BDBB1" box="[188,238,1189,1215]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; AF = male holotype (CASENT_9016117; CAS); GH= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AD = 0.5 mm; EF = 0.15 mm; GH = 0.1 mm. SEM images AF by Charles Griswold and Darrell Ubick, GH by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="10" pageNumber="11">12E</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FF510C81FEE5DBB1" box="[251,304,1189,1215]" captionStart="Fig" captionStartId="39.[189,232,1712,1738]" captionTargetBox="[189,1398,459,1674]" captionTargetId="figure-47@39.[189,1398,459,1674]" captionTargetPageId="39" captionText="Fig. 14. Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ARA.Ara.247343; RMCA). A. Male right palp (mirrored), retrolateral view. B. Idem, ventral view. C. Idem, dorsal view. D. Idem, bulbus, retrolateral view. E. Idem, ventral view. F. Idem, prolateral view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AC = 0.5mm; DF = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001825" httpUri="https://zenodo.org/record/14001825/files/figure.png" pageId="10" pageNumber="11">14D</figureCitation>
,
<figureCitation id="21C54D99FFF2FFC0FE960C81FE47DBCE" box="[316,402,1189,1216]" captionStart="Fig" captionStartId="40.[189,233,1731,1757]" captionTargetBox="[189,1398,478,1693]" captionTargetId="figure-47@40.[189,1398,478,1693]" captionTargetPageId="40" captionText="Fig. 15. Zorascar pasunepipe gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247343; RMCA), micro-CT scans of right male palp (mirrored). A. Retrolateral view. B. Ventral view. C. Prolateral view. D. Bulbus, retrolateral view.E. Bulbus, ventral view.F. Bulbus, apical view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AC = 0.5 mm; DF = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001827" httpUri="https://zenodo.org/record/14001827/files/figure.png" pageId="10" pageNumber="11">15DF</figureCitation>
). The paraconductor is found in other members of the Oval Calamistrum (OC) clade (the “TA” in
<bibRefCitation id="DD6F2CEDFFF2FFC0FEF80CEEFD96DBEB" author="Griswold C. E. &amp; Ramirez M. J. &amp; Coddington J. A. &amp; Platnick N. I." box="[338,579,1226,1253]" pageId="10" pageNumber="11" pagination="1 - 324" refId="ref36236" refString="Griswold C. E., Ramirez M. J., Coddington J. A. &amp; Platnick N. I. 2005. Atlas of phylogenetic data for entelegyne spiders (Araneae: Araneomorphae: Entelegynae) with comments on their phylogeny. Proceedings of the California Academy of Sciences 56 (Suppl. II): 1 - 324." type="journal article" year="2005">
Griswold
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2005
</bibRefCitation>
: figs 186BC, 195BC), and the Sierwald conductor (C) occurs widely among the OC clade and Dionycha
<bibRefCitation id="DD6F2CEDFFF2FFC0FDFD0CCBFC96DA04" author="Petrunkevitch A." box="[599,835,1263,1290]" pageId="10" pageNumber="11" pagination="1 - 270" refId="ref37617" refString="Petrunkevitch A. 1928. Systema Aranearum. Transactions of the Connecticut Academy of Arts and Sciences 29: 1 - 270." type="journal article" year="1928">Petrunkevitch, 1928</bibRefCitation>
(
<bibRefCitation id="DD6F2CEDFFF2FFC0FCFA0CCBFBE3DA04" author="Polotow D. &amp; Carmichael A. &amp; Griswold C. E." box="[848,1078,1263,1290]" pageId="10" pageNumber="11" pagination="124 - 163" refId="ref37852" refString="Polotow D., Carmichael A. &amp; Griswold C. E. 2015. Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae). Invertebrate Systematics 29: 124 - 163. https: // doi. org / 10.1071 / IS 14041" type="journal article" year="2015">
Polotow
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: figs. 13EF, 15AB, EF).
</paragraph>
<paragraph id="B941511CFFF2FFC0FF170D17FD68D8E6" blockId="10.[187,1400,1331,2025]" pageId="10" pageNumber="11">
<taxonomicName id="7EFE2A9FFFF2FFC0FF170D17FEFADA43" authorityName="Griswold &amp; Polotow" authorityYear="2015" box="[189,303,1331,1357]" class="Arachnida" family="Udubidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Udubidae</taxonomicName>
may be distinguished from most other araneomorph spider families found in
<collectingCountry id="C1E9118CFFF2FFC0FB6E0D17FA87DA43" box="[1220,1362,1331,1357]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
by the form of their tarsi. The presence of tarsal trichobothria distinguishes udubids from the
<taxonomicName id="7EFE2A9FFFF2FFC0FB5E0D7CFEA2DA99" authority="Ausserer, 1867" authorityName="Ausserer" authorityYear="1867" class="Arachnida" family="Filistatidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Filistatidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FF140D59FEA2DA99" author="Ausserer A." box="[190,375,1405,1431]" pageId="10" pageNumber="11" pagination="7 - 8" refId="ref35267" refString="Ausserer A. 1867. Die Arachniden Tirols nach ihrer horizontalen und verticalen Verbreitung; I. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 17: 137 - 170 ,, pl. 7 - 8." type="journal article" year="1867">Ausserer, 1867</bibRefCitation>
</taxonomicName>
, Synspermiata
<bibRefCitation id="DD6F2CEDFFF2FFC0FD9B0D59FCAEDA99" author="Michalik P. &amp; Ramirez M." box="[561,891,1405,1431]" pageId="10" pageNumber="11" pagination="291 - 322" refId="ref37310" refString="Michalik P. &amp; Ramirez M. 2014. Evolutionary morphology of the male reproductive system, spermatozoa and seminal fluid of spiders (Araneae, Arachnida) - Current knowledge and future directions. Arthropod Structure &amp; Development 43 (4): 291 - 322. https: // doi. org / 10.1016 / j. asd. 2014.05.005" type="journal article" year="2014">Michalik &amp; Ramírez, 2014</bibRefCitation>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FC230D59FBEEDA99" authorityName="Forster &amp; Platnick" authorityYear="1984" box="[905,1083,1405,1431]" class="Arachnida" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="68" phylum="Arthropoda" rank="superFamily" superFamily="Palpimanoidea">Palpimanoidea</taxonomicName>
<bibRefCitation id="DD6F2CEDFFF2FFC0FBEE0D5AFAA6DA99" author="Forster R. R. &amp; Platnick N. I." box="[1092,1395,1405,1432]" pageId="10" pageNumber="11" pagination="1 - 106" refId="ref35944" refString="Forster R. R. &amp; Platnick N. I. 1984. A review of the archaeid spiders and their relatives, with notes on the limits of the superfamily Palpimanoidea (Arachnida, Araneae). Bulletin of the American Museum of Natural History 178: 1 - 106." type="journal article" year="1984">
Forster &amp;
<taxonomicName id="7EFE2A9FFFF2FFC0FB680D59FAF2DA99" authority="Ubick 2007" authorityName="Marusik &amp; Fomichev" authorityYear="2020" box="[1218,1319,1405,1431]" class="Arachnida" family="Liocranidae" genus="Platnick" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="2" phylum="Arthropoda" rank="genus">Platnick</taxonomicName>
, 1984
</bibRefCitation>
, Araneoidea
<bibRefCitation id="DD6F2CEDFFF2FFC0FEE70D86FE29DAB2" author="Latreille P. A." box="[333,508,1442,1468]" pageId="10" pageNumber="11" pagination="82 - 127" refId="ref37107" refString="Latreille P. A. 1806. Araneae. In: Genera crustaceorum et insectorum. Tome 1: 82 - 127. A. Koenig, Paris." type="book chapter" year="1806">Latreille, 1806</bibRefCitation>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FDA20D86FCECDAB3" authority="C. L. Koch, 1845" authorityName="C. L. Koch" authorityYear="1845" box="[520,825,1442,1469]" class="Arachnida" family="Eresidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Eresidae C.L. Koch, 1845</taxonomicName>
(in
<bibRefCitation id="DD6F2CEDFFF2FFC0FCC20D86FBDFDAB3" author="Berendt G. C." box="[872,1034,1442,1469]" pageId="10" pageNumber="11" pagination="41 - 60" refId="ref35368" refString="Berendt G. C. 1845. Die organischen Bernstein-Einschlusse im Allgemeinen. In: Berendt G. C. (ed.) Die im Bernstein befindlichen organischen Reste der Vorwelt gesammelt in Verbindung mit Mehreren bearbeitet. Erster Band. II. Abtheilung: 41 - 60. Nicolaische Buchhandlung, Berlin. https: // doi. org / 10.5962 / bhl. title. 66910" type="book chapter" year="1845">Berendt 1845</bibRefCitation>
),
<taxonomicName id="7EFE2A9FFFF2FFC0FBB50D86FA92DAB2" authority="Thorell, 1869" authorityName="Thorell" authorityYear="1869" box="[1055,1351,1442,1468]" class="Arachnida" family="Hersiliidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Hersiliidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FB0D0D86FA92DAB2" author="Thorell T." box="[1191,1351,1442,1468]" pageId="10" pageNumber="11" pagination="109 - 242" refId="ref38870" refString="Thorell T. 1869. On European spiders. Review of the European genera of spiders, preceded by some observations on zoological nomenclature [first part]. Nova Acta Regiae Societatis Scientiarum Upsaliensis (3) 7 (I, 5): 1 - 108. [incl. pp. 109 - 242 from 1870]" type="journal article" year="1869">Thorell, 1869</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="7EFE2A9FFFF2FFC0FF170DE3FDC5DAEF" authority="Blackwall, 1862" authorityName="Blackwall" authorityYear="1862" box="[189,528,1479,1505]" class="Arachnida" family="Oecobiidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Oecobiidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FEE10DE3FDC5DAEF" author="Blackwall J." box="[331,528,1479,1505]" pageId="10" pageNumber="11" pagination="370 - 382" refId="ref35673" refString="Blackwall J. 1862. Descriptions of newly-discovered spiders from the island of Madeira. Annals and Magazine of Natural History (3) 9 (53): 370 - 382. https: // doi. org / 10.1080 / 00222936208681249 Bonnet P. 1959. Bibliographia Araneorum. Analyse Methodique de Toute la Litterature Araneologique Jusqu'en 1939. Tome II. Systematique des Raignees (Etude par Ordre Alphabetique) (5 eme Partie: T - Z). Douladoure, Toulouse." type="journal article" year="1862">Blackwall, 1862</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FDB70DE3FCA6DAEF" authority="Lehtinen, 1967" authorityName="Lehtinen" authorityYear="1967" box="[541,883,1479,1505]" class="Arachnida" family="Phyxelididae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Phyxelididae
<bibRefCitation id="DD6F2CEDFFF2FFC0FD150DE3FCA6DAEF" author="Lehtinen P. T." box="[703,883,1479,1505]" pageId="10" pageNumber="11" pagination="199 - 468" refId="ref37164" refString="Lehtinen P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="7EFE2A9FFFF2FFC0FC040DE3FAD1DAEF" authority="Lehtinen, 1967" authorityName="Lehtinen" authorityYear="1967" box="[942,1284,1479,1505]" class="Arachnida" family="Titanoecidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Titanoecidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FBE70DE3FAD1DAEF" author="Lehtinen P. T." box="[1101,1284,1479,1505]" pageId="10" pageNumber="11" pagination="199 - 468" refId="ref37164" refString="Lehtinen P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
</taxonomicName>
. Multiple rows of tarsal trichobothria distinguish
<taxonomicName id="7EFE2A9FFFF2FFC0FD2C0DC8FD2DD908" authorityName="Griswold &amp; Polotow" authorityYear="2015" box="[646,760,1516,1542]" class="Arachnida" family="Udubidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Udubidae</taxonomicName>
from taxa with only a single row of tarsal trichobothria including
<taxonomicName id="7EFE2A9FFFF2FFC0FE990E35FD5BD925" authority="C. L. Koch, 1837" authorityName="C. L. Koch" authorityYear="1837" box="[307,654,1553,1579]" class="Arachnida" family="Agelenidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Agelenidae C.L.
<bibRefCitation id="DD6F2CEDFFF2FFC0FDAA0E35FD5BD925" author="Koch C. L." box="[512,654,1553,1579]" pageId="10" pageNumber="11" refId="ref36844" refString="Koch C. L. 1837. Ubersicht des Arachnidensystems. Heft 1. C. H. Zeh'sche Buchhandlung, Nurnberg. https: // doi. org / 10.5962 / bhl. title. 39561" type="book" year="1837">Koch, 1837</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FD310E35FB54D925" authority="O. Pickard-Cambridge, 1871" authorityName="O. Pickard-Cambridge" authorityYear="1871" box="[667,1153,1553,1579]" class="Arachnida" family="Dictynidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Dictynidae O.
<bibRefCitation id="DD6F2CEDFFF2FFC0FCE10E35FB54D925" author="Pickard-Cambridge O." box="[843,1153,1553,1579]" pageId="10" pageNumber="11" pagination="207 - 224" refId="ref37701" refString="Pickard-Cambridge O. 1871. Arachnida. The Zoological Record 7: 207 - 224." type="journal article" year="1871">Pickard-Cambridge, 1871</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FB250E35FF2FD95F" authority="Bertkau, 1878" authorityName="Bertkau" authorityYear="1878" class="Arachnida" family="Hahniidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Hahniidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FABB0E35FF2FD95F" author="Bertkau P." pageId="10" pageNumber="11" pagination="351 - 410" refId="ref35457" refString="Bertkau P. 1878. Versuch einer naturlichen Anordnung der Spinnen, nebst Bemerkungen zu einzelnen Gattungen. Archiv fur Naturgeschichte 44: 351 - 410." type="journal article" year="1878">Bertkau, 1878</bibRefCitation>
</taxonomicName>
and most
<taxonomicName id="7EFE2A9FFFF2FFC0FEDD0E12FD75D95F" authority="Thorell, 1881" authorityName="Thorell" authorityYear="1881" box="[375,672,1590,1617]" class="Arachnida" family="Zodariidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Zodariidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FE550E12FD75D95F" author="Thorell T." box="[511,672,1590,1617]" pageId="10" pageNumber="11" pagination="1 - 720" refId="ref38930" refString="Thorell T. 1881. Studi sui Ragni Malesi e Papuani. III. Ragni dell'Austro Malesia e del Capo York, conservati nel Museo civico di storia naturale di Genova. Annali del Museo Civico di Storia Naturale di Genova 17: 1 - 720." type="journal article" year="1881">Thorell, 1881</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="7EFE2A9FFFF2FFC0FD710E12FBE7D95F" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" box="[731,1074,1590,1617]" class="Arachnida" family="Thomisidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Thomisidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FCC50E12FBE7D95F" author="Sundevall C. J." box="[879,1074,1590,1617]" pageId="10" pageNumber="11" refId="ref38806" refString="Sundevall C. J. 1833. Conspectus Arachnidum. C. F. Berling, Londini Gothorum [Lund]." type="book" year="1833">Sundevall, 1833</bibRefCitation>
</taxonomicName>
. Among taxa with multiple rows of tarsal trichobothria udubids may be distinguished from
<taxonomicName id="7EFE2A9FFFF2FFC0FC1A0E7FFB24D978" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" box="[944,1265,1627,1654]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Lycosidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FB850E7FFB24D978" author="Sundevall C. J." box="[1071,1265,1627,1654]" pageId="10" pageNumber="11" refId="ref38806" refString="Sundevall C. J. 1833. Conspectus Arachnidum. C. F. Berling, Londini Gothorum [Lund]." type="book" year="1833">Sundevall, 1833</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FB540E7FFE86D995" authority="Simon, 1890" authorityName="Simon" authorityYear="1890" class="Arachnida" family="Pisauridae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Pisauridae
<bibRefCitation id="DD6F2CEDFFF2FFC0FF140EA4FE86D995" author="Simon E." box="[190,339,1664,1691]" pageId="10" pageNumber="11" pagination="77 - 124" refId="ref38391" refString="Simon E. 1890. Etudes arachnologiques. 22 e Memoire. XXXIV. Etude sur les arachnides de l'Yemen. Annales de la Societe entomologique de France (6) 10: 77 - 124." type="journal article" year="1890">Simon, 1890</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="7EFE2A9FFFF2FFC0FE200EA4FD7AD994" authority="Thorell, 1869" authorityName="Thorell" authorityYear="1869" box="[394,687,1664,1690]" class="Arachnida" family="Oxyopidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Oxyopidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FDBA0EA4FD7AD994" author="Thorell T." box="[528,687,1664,1690]" pageId="10" pageNumber="11" pagination="109 - 242" refId="ref38870" refString="Thorell T. 1869. On European spiders. Review of the European genera of spiders, preceded by some observations on zoological nomenclature [first part]. Nova Acta Regiae Societatis Scientiarum Upsaliensis (3) 7 (I, 5): 1 - 108. [incl. pp. 109 - 242 from 1870]" type="journal article" year="1869">
Thorell,
<quantity id="7E06FCF9FFF2FFC0FDDE0EA4FD7AD994" box="[628,687,1664,1690]" metricMagnitude="1" metricUnit="m" metricValue="4.74726" pageId="10" pageNumber="11" unit="in" value="1869.0">1869</quantity>
</bibRefCitation>
</taxonomicName>
in that these latter families have at least one eye row strongly curved, whereas
<taxonomicName id="7EFE2A9FFFF2FFC0FE290E82FE20D9CE" authorityName="Griswold &amp; Polotow" authorityYear="2015" box="[387,501,1702,1728]" class="Arachnida" family="Udubidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Udubidae</taxonomicName>
have both eye rows nearly straight. Udubids have at least a rudiment of the inferior tarsal claw and lack claw tufts, characters that distinguish them from
<taxonomicName id="7EFE2A9FFFF2FFC0FBEB0EEFFAADD9EB" authority="Bertkau, 1872" authorityName="Bertkau" authorityYear="1872" box="[1089,1400,1739,1765]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Sparassidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FB780EEFFAADD9EB" author="Bertkau P." box="[1234,1400,1739,1765]" pageId="10" pageNumber="11" pagination="208 - 233" refId="ref35437" refString="Bertkau P. 1872. Uber die Respirationsorgane der Araneen. Archiv fur Naturgeschichte 38: 208 - 233." type="journal article" year="1872">Bertkau, 1872</bibRefCitation>
</taxonomicName>
and the Dionycha (
<taxonomicName id="7EFE2A9FFFF2FFC0FE3C0ED4FCDDD804" authority="Wagner, 1887" authorityName="Wagner" authorityYear="1887" box="[406,776,1776,1802]" class="Arachnida" family="Cheiracanthiidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Cheiracanthiidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FDCF0ED4FCDDD804" author="Wagner W. A." box="[613,776,1776,1802]" pageId="10" pageNumber="11" pagination="3 - 132" refId="ref39073" refString="Wagner W. A. 1887. Copulationsorgane des Mannchens als Criterium fur die Systematik der Spinnen. Horae Societatis Entomologicae Rossicae 22: 3 - 132." type="journal article" year="1887">Wagner, 1887</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FCB90ED4FB95D804" authority="Simon, 1878" authorityName="Simon" authorityYear="1878" box="[787,1088,1776,1802]" class="Arachnida" family="Clubionidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Clubionidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FC030ED4FB95D804" author="Simon E." box="[937,1088,1776,1802]" pageId="10" pageNumber="11" refId="ref38290" refString="Simon E. 1878. Les Arachnides de France. Tome Quatrieme, Contenant la Famille des Drassidae. Roret, Paris." type="book" year="1878">Simon, 1878</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FBE10ED4FAA4D804" authority="Karsch, 1880" authorityName="Karsch" authorityYear="1880" box="[1099,1393,1776,1802]" class="Arachnida" family="Corinnidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Corinnidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FB790ED4FAA4D804" author="Karsch F." box="[1235,1393,1776,1802]" pageId="10" pageNumber="11" pagination="373 - 409" refId="ref36642" refString="Karsch F. 1880. Arachnologische Blatter (Decas I). Zeitschrift fur die Gesammten Naturwissenschaften, Dritte Folge 5: 373 - 409, pl. 12." type="journal article" year="1880">Karsch, 1880</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FF160F31FE2ED821" authority="Millot, 1947" authorityName="Millot" authorityYear="1947" box="[188,507,1813,1839]" class="Arachnida" family="Gallieniellidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Gallieniellidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FEC10F31FE2ED821" author="Millot J." box="[363,507,1813,1839]" pageId="10" pageNumber="11" pagination="158 - 160" refId="ref37426" refString="Millot J. 1947. Une araignee malgache enigmatique, Gallieniella mygaloides n. g., n. sp. Bulletin du Museum national d'Histoire naturelle de Paris (2) 19 (2): 158 - 160, pl. 1." type="journal article" year="1947">Millot, 1947</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FDAC0F31FCE6D821" authority="Banks, 1892" authorityName="Banks" authorityYear="1892" box="[518,819,1813,1839]" class="Arachnida" family="Gnaphosidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Gnaphosidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FD080F31FCE6D821" author="Banks N." box="[674,819,1813,1839]" pageId="10" pageNumber="11" pagination="88 - 97" refId="ref35303" refString="Banks N. 1892. A classification of the North American spiders. The Canadian Entomologist 24 (4): 88 - 97. https: // doi. org / 10.4039 / Ent 2488 - 4" type="journal article" year="1892">Banks, 1892</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FC970F31FBB5D821" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[829,1120,1813,1839]" class="Arachnida" family="Liocranidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Liocranidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FC660F31FBB5D821" author="Simon E." box="[972,1120,1813,1839]" pageId="10" pageNumber="11" refId="ref38503" refString="Simon E. 1897. Histoire naturelle des Araignees. Deuxieme Edition, Tome second. Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FBC10F31FF2DD85A" authority="Thorell, 1869" authorityName="Thorell" authorityYear="1869" class="Arachnida" family="Philodromidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Philodromidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FAB30F31FF2DD85A" author="Thorell T." pageId="10" pageNumber="11" pagination="109 - 242" refId="ref38870" refString="Thorell T. 1869. On European spiders. Review of the European genera of spiders, preceded by some observations on zoological nomenclature [first part]. Nova Acta Regiae Societatis Scientiarum Upsaliensis (3) 7 (I, 5): 1 - 108. [incl. pp. 109 - 242 from 1870]" type="journal article" year="1869">Thorell, 1869</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FEAE0F1EFD97D85A" authority="Blackwall, 1841" authorityName="Blackwall" authorityYear="1841" box="[260,578,1850,1876]" class="Arachnida" family="Salticidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Salticidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FED50F1EFD97D85A" author="Blackwall J." box="[383,578,1850,1876]" pageId="10" pageNumber="11" pagination="601 - 670" refId="ref35591" refString="Blackwall J. 1841. The difference in the number of eyes with which spiders are provided proposed as the basis of their distribution into tribes; with descriptions of newly discovered species and the characters of a new family and three new genera of spiders. Transactions of the Linnean Society of London 18 (4): 601 - 670. https: // doi. org / 10.1111 / j. 1095 - 8339.1838. tb 00210. x" type="journal article" year="1841">Blackwall, 1841</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FDE40F1EFCA9D85A" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[590,892,1850,1876]" class="Arachnida" family="Selenopidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Selenopidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FD4E0F1EFCA9D85A" author="Simon E." box="[740,892,1850,1876]" pageId="10" pageNumber="11" refId="ref38503" refString="Simon E. 1897. Histoire naturelle des Araignees. Deuxieme Edition, Tome second. Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FC220F1EFB64D85A" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[904,1201,1850,1876]" class="Arachnida" family="Trachelidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Trachelidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FBB20F1EFB64D85A" author="Simon E." box="[1048,1201,1850,1876]" pageId="10" pageNumber="11" refId="ref38503" refString="Simon E. 1897. Histoire naturelle des Araignees. Deuxieme Edition, Tome second. Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FB160F1EFE83D877" authority="Karsch, 1879" authorityName="Karsch" authorityYear="1879" class="Arachnida" family="Trochanteriidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Trochanteriidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FF170F7BFE83D877" author="Karsch F." box="[189,342,1887,1913]" pageId="10" pageNumber="11" pagination="534 - 562" refId="ref36623" refString="Karsch F. 1879. Arachnologische Beitrage. Zeitschrift fur die Gesammten Naturwissenschaften 52: 534 - 562." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="7EFE2A9FFFF2FFC0FE210F7BFD05D877" authority="Lehtinen, 1967" authorityName="Lehtinen" authorityYear="1967" box="[395,720,1887,1913]" class="Arachnida" family="Viridasiidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Viridasiidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FDB70F7BFD05D877" author="Lehtinen P. T." box="[541,720,1887,1913]" pageId="10" pageNumber="11" pagination="199 - 468" refId="ref37164" refString="Lehtinen P. T. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
</taxonomicName>
). Finally, like
<taxonomicName id="7EFE2A9FFFF2FFC0FCDA0F7BFC31D877" authorityName="Griswold &amp; Polotow" authorityYear="2015" box="[880,996,1887,1913]" class="Arachnida" family="Udubidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Udubidae</taxonomicName>
, the lone genus of
<taxonomicName id="7EFE2A9FFFF2FFC0FB1C0F7BFF23D891" authority="Pocock, 1895" authorityName="Pocock" authorityYear="1895" class="Arachnida" family="Desidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
Desidae
<bibRefCitation id="DD6F2CEDFFF2FFC0FAB30F7BFF23D891" author="Pocock R. I." pageId="10" pageNumber="11" pagination="139 - 143" refId="ref37777" refString="Pocock R. I. 1895. Description of two new spiders obtained by Messrs. J. J. Quelch and F. McConnell on the summit of Mount Roraima, in Demerara; with a note upon the systematic position of the genus Desis. Annals and Magazine of Natural History (6) 16 (92): 139 - 143. https: // doi. org / 10.1080 / 00222939508680241" type="journal article" year="1895">Pocock, 1895</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="C1E9118CFFF2FFC0FE900FA0FE1CD890" box="[314,457,1924,1950]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
,
<taxonomicName id="7EFE2A9FFFF2FFC0FE7E0FA1FD3BD890" authority="Walckenaer, 1837" authorityName="Walckenaer" authorityYear="1837" box="[468,750,1924,1950]" class="Arachnida" family="Desidae" genus="Desis" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFF2FFC0FE7E0FA1FDC0D890" box="[468,533,1925,1950]" italics="true" pageId="10" pageNumber="11">Desis</emphasis>
<bibRefCitation id="DD6F2CEDFFF2FFC0FDB60FA0FD3BD890" author="Walckenaer C. A." box="[540,750,1924,1950]" pageId="10" pageNumber="11" refId="ref39101" refString="Walckenaer C. A. 1837. Histoire naturelle des Insectes. Apteres. Tome premier. Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 61095" type="book" year="1837">Walckenaer, 1837</bibRefCitation>
</taxonomicName>
, has numerous tarsal trichobothria and straight eye rows but the latter is quite distinct in having elongate chelicerae, a dense cover of fine hairs and the posterior tracheal spiracle on the abdomen advanced.
</paragraph>
<paragraph id="B941511CFFF3FFC1FF17092CFE88DE2C" blockId="11.[189,349,264,290]" box="[189,349,264,290]" pageId="11" pageNumber="12">
<heading id="E209E670FFF3FFC1FF17092CFE88DE2C" bold="true" box="[189,349,264,290]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="8B8A8D0EFFF3FFC1FF17092CFE88DE2C" bold="true" box="[189,349,264,290]" pageId="11" pageNumber="12">Composition</emphasis>
</heading>
</paragraph>
<paragraph id="B941511CFFF3FFC1FF17091CFB71DE78" blockId="11.[189,1399,312,375]" pageId="11" pageNumber="12">
<taxonomicName id="7EFE2A9FFFF3FFC1FF17091CFDE9DE5C" authority="Karsch, 1892" authorityName="Karsch" authorityYear="1892" box="[189,572,312,338]" class="Arachnida" family="Udubidae" genus="Campostichomma" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFF3FFC1FF17091CFE46DE5C" box="[189,403,312,338]" italics="true" pageId="11" pageNumber="12">Campostichomma</emphasis>
<bibRefCitation id="DD6F2CEDFFF3FFC1FE36091CFDE9DE5C" author="Karsch F." box="[412,572,312,338]" pageId="11" pageNumber="12" pagination="267 - 310" refId="ref36672" refString="Karsch F. 1892. Arachniden von Ceylon und von Minikoy gesammelt von den Herren Doctoren P. und F. Sarasin. Berliner Entomologische Zeitschrift 36 (2, 1891): 267 - 310, pl. 10 - 12. [Published mid March 1892.]" type="journal article" year="1892">Karsch, 1892</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="C1E9118CFFF3FFC1FDE7091CFD1CDE5C" box="[589,713,312,338]" name="Sri Lanka" pageId="11" pageNumber="12">Sri Lanka</collectingCountry>
),
<taxonomicName id="7EFE2A9FFFF3FFC1FD4A091DFC0BDE5C" authority="Simon, 1892" authorityName="Simon" authorityYear="1892" box="[736,990,312,338]" class="Arachnida" family="Udubidae" genus="Raecius" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFF3FFC1FD4A091DFCEBDE5C" box="[736,830,313,338]" italics="true" pageId="11" pageNumber="12">Raecius</emphasis>
<bibRefCitation id="DD6F2CEDFFF3FFC1FCEC091CFC0BDE5C" author="Simon E." box="[838,990,312,338]" pageId="11" pageNumber="12" refId="ref38428" refString="Simon E. 1892. Histoire naturelle des Araignees. Deuxieme Edition, Tome premier. Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1892">Simon, 1892</bibRefCitation>
</taxonomicName>
(Africa), and
<taxonomicName id="7EFE2A9FFFF3FFC1FB27091CFB2FDE5C" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1165,1274,312,338]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8B8A8D0EFFF3FFC1FB27091CFB2FDE5C" box="[1165,1274,312,338]" italics="true" pageId="11" pageNumber="12">Tabiboka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFF3FFC1FAA8091DFAA7DE5D" box="[1282,1394,313,339]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
,
<taxonomicName id="7EFE2A9FFFF3FFC1FF150978FE70DE78" authority="Simon 1880" authorityName="Simon" authorityYear="1880" box="[191,421,348,374]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFF3FFC1FF150978FEDADE78" box="[191,271,348,374]" italics="true" pageId="11" pageNumber="12">Uduba</emphasis>
<bibRefCitation id="DD6F2CEDFFF3FFC1FEBE0978FE70DE78" author="Simon E." box="[276,421,348,374]" pageId="11" pageNumber="12" pagination="223 - 351" refId="ref38313" refString="Simon E. 1880. Revision de la famille des Sparassidae (Arachnides). Actes de la Societe Linneenne de Bordeaux 34 (2 / 3 / 4): 223 - 351." type="journal article" year="1880">Simon 1880</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7EFE2A9FFFF3FFC1FE1B0979FDC8DE78" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[433,541,349,374]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8B8A8D0EFFF3FFC1FE1B0979FDC8DE78" box="[433,541,349,374]" italics="true" pageId="11" pageNumber="12">Zorascar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFF3FFC1FD8E0979FD5ADE79" box="[548,655,349,375]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
and
<taxonomicName id="7EFE2A9FFFF3FFC1FD620978FC21DE78" authority="Strand, 1907" authorityName="Strand" authorityYear="1907" box="[712,1012,348,374]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFF3FFC1FD620978FC83DE78" box="[712,854,348,374]" italics="true" pageId="11" pageNumber="12">Zorodictyna</emphasis>
<bibRefCitation id="DD6F2CEDFFF3FFC1FCF70978FC21DE78" author="Strand E." box="[861,1012,348,374]" pageId="11" pageNumber="12" pagination="725 - 748" refId="ref38669" refString="Strand E. 1907. Diagnosen neuer Spinnen aus Madagaskar und Sansibar. Zoologischer Anzeiger 31: 725 - 748." type="journal article" year="1907">Strand, 1907</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="C1E9118CFFF3FFC1FBA90978FB4DDE78" box="[1027,1176,348,374]" name="Madagascar" pageId="11" pageNumber="12">Madagascar</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="F1E40297FFF3FFC6FF1709BBFAA2DBF1" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="key">
<paragraph id="B941511CFFF3FFC1FF1709BBFD86DEB7" blockId="11.[189,595,415,441]" box="[189,595,415,441]" pageId="11" pageNumber="12">
<heading id="E209E670FFF3FFC1FF1709BBFD86DEB7" bold="true" box="[189,595,415,441]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="8B8A8D0EFFF3FFC1FF1709BBFD86DEB7" bold="true" box="[189,595,415,441]" pageId="11" pageNumber="12">
Key to
<taxonomicName id="7EFE2A9FFFF3FFC1FEBD09BBFE44DEB7" authorityName="Griswold &amp; Polotow" authorityYear="2015" box="[279,401,415,441]" class="Arachnida" family="Udubidae" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Udubidae</taxonomicName>
of
<collectingCountry id="C1E9118CFFF3FFC1FE1209BBFD86DEB7" box="[440,595,415,441]" name="Madagascar" pageId="11" pageNumber="12">Madagascar</collectingCountry>
</emphasis>
</heading>
</paragraph>
<key id="CD6F062DFFF3FFC6FF1709EBFAA2DBF1" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<keyStep id="020A49B9FFF3FFC1FF1709EBFAA3DB23" pageId="11" pageNumber="12">
<paragraph id="B941511CFFF3FFC1FF1709EBFAA2DDCC" blockId="11.[189,1400,462,1070]" pageId="11" pageNumber="12">
<keyLead id="020FF229FFF3FFC1FF1709EBFAA2DDCC" pageId="11" pageNumber="12">
1. Male palp without process between tibia and patella; cymbium with retrobasal extension (CE), short to as long as rest of cymbium (
<figureCitation id="21C54D99FFF3FFC1FDFF09D7FD0DDD00" box="[597,728,499,526]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="11" pageNumber="12">Figs 5BC</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FD4E09D7FCF8DD03" box="[740,813,499,525]" captionStart="Fig" captionStartId="19.[188,230,1787,1813]" captionTargetBox="[188,1399,479,1749]" captionTargetId="figure-89@19.[188,1399,479,1749]" captionTargetPageId="19" captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AC = 0.43 mm; DF = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick." figureDoi="http://doi.org/10.5281/zenodo.14001804" httpUri="https://zenodo.org/record/14001804/files/figure.png" pageId="11" pageNumber="12">6AC</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FC9309D7FC55DD03" box="[825,896,499,525]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.AC = male holotype (CASENT_9016063; MNHN). DE = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations AC by Jenny Speckels; DE by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="11" pageNumber="12">9BC</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FC2709D7FC37DD03" box="[909,994,499,525]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; AC= male holotype (CASENT_9016117; CAS); DE = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="11" pageNumber="12">11BC</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FC4409D7FB90DD03" box="[1006,1093,499,525]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; AF = male holotype (CASENT_9016117; CAS); GH= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AD = 0.5 mm; EF = 0.15 mm; GH = 0.1 mm. SEM images AF by Charles Griswold and Darrell Ubick, GH by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="11" pageNumber="12">12BD</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FBF809D7FB7CDD03" box="[1106,1193,499,525]" captionStart="Fig" captionStartId="39.[189,232,1712,1738]" captionTargetBox="[189,1398,459,1674]" captionTargetId="figure-47@39.[189,1398,459,1674]" captionTargetPageId="39" captionText="Fig. 14. Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ARA.Ara.247343; RMCA). A. Male right palp (mirrored), retrolateral view. B. Idem, ventral view. C. Idem, dorsal view. D. Idem, bulbus, retrolateral view. E. Idem, ventral view. F. Idem, prolateral view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AC = 0.5mm; DF = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001825" httpUri="https://zenodo.org/record/14001825/files/figure.png" pageId="11" pageNumber="12">14AC</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FB1C09D7FAD8DD00" box="[1206,1293,499,526]" captionStart="Fig" captionStartId="40.[189,233,1731,1757]" captionTargetBox="[189,1398,478,1693]" captionTargetId="figure-47@40.[189,1398,478,1693]" captionTargetPageId="40" captionText="Fig. 15. Zorascar pasunepipe gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247343; RMCA), micro-CT scans of right male palp (mirrored). A. Retrolateral view. B. Ventral view. C. Prolateral view. D. Bulbus, retrolateral view.E. Bulbus, ventral view.F. Bulbus, apical view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AC = 0.5 mm; DF = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001827" httpUri="https://zenodo.org/record/14001827/files/figure.png" pageId="11" pageNumber="12">15AB</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FAB309D7FAA4DD03" box="[1305,1393,499,525]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; AB = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); CD = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AB = 2 mm; CE = 0.5 mm. Illustrations AB, E by Alain Reygel; CD by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="11" pageNumber="12">16AB</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FF410A33FE94DD3F" box="[235,321,535,561]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="11" pageNumber="12">23BC</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FEE40A33FE56DD3C" box="[334,387,535,562]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="11" pageNumber="12">25C</figureCitation>
); embolus broad and flat to slender, cylindrical, firmly attached to point of origin on retromargin of tegulum (
<figureCitation id="21C54D99FFF3FFC1FDB20A1FFD74DD58" box="[536,673,571,598]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="11" pageNumber="12">Figs 5AB</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FD050A1FFD22DD5B" box="[687,759,571,597]" captionStart="Fig" captionStartId="19.[188,230,1787,1813]" captionTargetBox="[188,1399,479,1749]" captionTargetId="figure-89@19.[188,1399,479,1749]" captionTargetPageId="19" captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AC = 0.43 mm; DF = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick." figureDoi="http://doi.org/10.5281/zenodo.14001804" httpUri="https://zenodo.org/record/14001804/files/figure.png" pageId="11" pageNumber="12">6DE</figureCitation>
, 8F, 9AB, 11AB, 12E, 14EF, 15B, DF, 23AB, 25AB); tegulum oval apically; conductor hyaline, fan-shaped crest arising from middle to apex of tegulum; female endogyne with spermathecal ducts short, stout, not making loops, at most twisted (
<figureCitation id="21C54D99FFF3FFC1FF590A8CFE87DDCD" box="[243,338,680,707]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="11" pageNumber="12">Figs 5E</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FEF40A8CFE71DDCC" box="[350,420,680,706]" captionStart="Fig" captionStartId="20.[189,232,1822,1848]" captionTargetBox="[189,1398,461,1784]" captionTargetId="figure-79@20.[189,1398,461,1784]" captionTargetPageId="20" captionText="Fig. 7. Tabiboka milleri gen. nov. et sp. nov. SEM of female genitalia, AB, EF = female paratype (CASENT_9016094; CAS); CD = female paratype (CASENT_9035866; CAS). A. Epigyne, ventral view. B. Idem, ventro-posterior view. C. Endogyne, dorsal view. D. Idem, slightly anterior view. E. Spermatheca, right dorsal view. F. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: AE = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila." figureDoi="http://doi.org/10.5281/zenodo.14001806" httpUri="https://zenodo.org/record/14001806/files/figure.png" pageId="11" pageNumber="12">7CE</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FE1A0A8CFE06DDCC" box="[432,467,680,706]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.AC = male holotype (CASENT_9016063; MNHN). DE = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations AC by Jenny Speckels; DE by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="11" pageNumber="12">9E</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FE750A8CFDC5DDCC" box="[479,528,680,706]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; AC= male holotype (CASENT_9016117; CAS); DE = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="11" pageNumber="12">11E</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FDB60A8CFD85DDCC" box="[540,592,680,706]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; AF = male holotype (CASENT_9016117; CAS); GH= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AD = 0.5 mm; EF = 0.15 mm; GH = 0.1 mm. SEM images AF by Charles Griswold and Darrell Ubick, GH by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="11" pageNumber="12">12H</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FDF70A8CFD47DDCC" box="[605,658,680,706]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; AB = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); CD = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AB = 2 mm; CE = 0.5 mm. Illustrations AB, E by Alain Reygel; CD by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="11" pageNumber="12">16D</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FD340A8CFD23DDCC" box="[670,758,680,706]" captionStart="Fig" captionStartId="46.[189,233,1894,1920]" captionTargetBox="[189,1398,1053,1856]" captionTargetId="figure-159@46.[189,1398,1053,1859]" captionTargetPageId="46" captionText="Fig. 18. Zorascar pasunepomme gen. et sp. nov. holotype female (BE_RMCA_ARA.Ara.247346; RMCA), genitalia. A. Epigyne, ventral view. B. Endogyne, cleared, dorsal view. C. Idem, dorso-anterior view. D. Idem, dorso-posterior view. Scale bars = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001835" httpUri="https://zenodo.org/record/14001835/files/figure.png" pageId="11" pageNumber="12">18BD</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FCA80A8CFC87DDCC" box="[770,850,680,706]" captionStart="Fig" captionStartId="47.[189,232,1570,1596]" captionTargetBox="[189,1398,265,1533]" captionTargetId="figure-11@47.[189,1398,265,1533]" captionTargetPageId="47" captionText="Fig. 19. Zorascar pasunepomme gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247346; RMCA), epigyne, CT-scan images. A. ventral view. B. ventro-anterior view. C. dorsal view. D. dorso-anterior view. E. idem, detail.F. Idem, dorso-lateral view. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; ML = median lobe; MS = median sector; S = spermatheca. Scale bars = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001837" httpUri="https://zenodo.org/record/14001837/files/figure.png" pageId="11" pageNumber="12">19CF</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FCF40A8CFC45DDCC" box="[862,912,680,706]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="11" pageNumber="12">23E</figureCitation>
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<figureCitation id="21C54D99FFF3FFC1FC360A8CFC19DDCD" box="[924,972,680,707]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="11" pageNumber="12">25E</figureCitation>
) ..................................................... 2
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</paragraph>
<paragraph id="B941511CFFF3FFC1FF6B0AE9FAA3DB23" blockId="11.[189,1400,462,1070]" pageId="11" pageNumber="12">
<keyLead id="020FF229FFF3FFC1FF6B0AE9FAA3DB23" pageId="11" pageNumber="12">
Male palp with small ventral process at base of tibia between tibia and patella (“
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” in
<bibRefCitation id="DD6F2CEDFFF3FFC1FAA30AE8FEA5DC05" author="Griswold C. &amp; Ubick D. &amp; Ledford J. &amp; Polotow D." pageId="11" pageNumber="12" pagination="1 - 193" refId="ref36360" refString="Griswold C., Ubick D., Ledford J. &amp; Polotow D. 2022. A revision of the Malagasy crack-leg spiders of the genus Uduba Simon 1880 (Araneae, Udubidae), with description of 35 new species from Madagascar. Proceedings of the California Academy of Sciences (4) 67 (Supplement 2): 1 - 193." type="journal article" year="2022">
Griswold
<emphasis id="8B8A8D0EFFF3FFC1FF410AD6FEF2DC05" box="[235,295,753,779]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2022
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: figs 27AB, 39AC, 40AC, 41B); cymbium rounded retrobasally; embolus flexibly attached to large, swollen base, embolus elongate and extending behind tegulum and subtegulum to emerge from alveolus near bulb apex (
<bibRefCitation id="DD6F2CEDFFF3FFC1FD180B1DFC79DC5D" author="Griswold C. &amp; Ubick D. &amp; Ledford J. &amp; Polotow D." box="[690,940,825,851]" pageId="11" pageNumber="12" pagination="1 - 193" refId="ref36360" refString="Griswold C., Ubick D., Ledford J. &amp; Polotow D. 2022. A revision of the Malagasy crack-leg spiders of the genus Uduba Simon 1880 (Araneae, Udubidae), with description of 35 new species from Madagascar. Proceedings of the California Academy of Sciences (4) 67 (Supplement 2): 1 - 193." type="journal article" year="2022">
Griswold
<emphasis id="8B8A8D0EFFF3FFC1FC800B1EFCB0DC5D" box="[810,869,825,851]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2022
</bibRefCitation>
: figs 27AB, 32C, 41F, 51A); tegulum trilobed apically, with retrolateral (TA1), median (TA2) and prolateral (TA3) processes interacting apically (
<bibRefCitation id="DD6F2CEDFFF3FFC1FEF10BA6FD8FDC92" author="Griswold C. &amp; Ubick D. &amp; Ledford J. &amp; Polotow D." box="[347,602,898,924]" pageId="11" pageNumber="12" pagination="1 - 193" refId="ref36360" refString="Griswold C., Ubick D., Ledford J. &amp; Polotow D. 2022. A revision of the Malagasy crack-leg spiders of the genus Uduba Simon 1880 (Araneae, Udubidae), with description of 35 new species from Madagascar. Proceedings of the California Academy of Sciences (4) 67 (Supplement 2): 1 - 193." type="journal article" year="2022">
Griswold
<emphasis id="8B8A8D0EFFF3FFC1FE7F0BA7FDC4DC92" box="[469,529,898,924]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2022
</bibRefCitation>
: figs 27AB, 29AI, 51AC); conductor rectangular hyaline crest arising along apical margin of prolateral tegular lobe near TA3; female endogyne with spermathecal ducts elongate, making at least one (to as many as five) loops (
<bibRefCitation id="DD6F2CEDFFF3FFC1FC760BEEFB0FDCEB" author="Griswold C. &amp; Ubick D. &amp; Ledford J. &amp; Polotow D." box="[988,1242,970,997]" pageId="11" pageNumber="12" pagination="1 - 193" refId="ref36360" refString="Griswold C., Ubick D., Ledford J. &amp; Polotow D. 2022. A revision of the Malagasy crack-leg spiders of the genus Uduba Simon 1880 (Araneae, Udubidae), with description of 35 new species from Madagascar. Proceedings of the California Academy of Sciences (4) 67 (Supplement 2): 1 - 193." type="journal article" year="2022">
Griswold
<emphasis id="8B8A8D0EFFF3FFC1FBFF0BEFFB47DCEA" box="[1109,1170,970,996]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2022
</bibRefCitation>
: figs 68BD, 80BD, FG, IJ) ............................................................................................................................... .........
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<emphasis id="8B8A8D0EFFF3FFC1FE990C36FE51DB23" bold="true" box="[307,388,1042,1069]" italics="true" pageId="11" pageNumber="12">Uduba</emphasis>
<bibRefCitation id="DD6F2CEDFFF3FFC1FE260C37FDF7DB23" author="Simon E." box="[396,546,1043,1069]" pageId="11" pageNumber="12" pagination="223 - 351" refId="ref38313" refString="Simon E. 1880. Revision de la famille des Sparassidae (Arachnides). Actes de la Societe Linneenne de Bordeaux 34 (2 / 3 / 4): 223 - 351." type="journal article" year="1880">Simon, 1880</bibRefCitation>
</taxonomicName>
(the key to
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<emphasis id="8B8A8D0EFFF3FFC1FD040C37FD2BDB23" box="[686,766,1043,1069]" italics="true" pageId="11" pageNumber="12">Uduba</emphasis>
species
</taxonomicName>
is amply detailed in
<bibRefCitation id="DD6F2CEDFFF3FFC1FBE50C37FAB8DB23" author="Griswold C. &amp; Ubick D. &amp; Ledford J. &amp; Polotow D." box="[1103,1389,1043,1069]" pageId="11" pageNumber="12" pagination="1 - 193" refId="ref36360" refString="Griswold C., Ubick D., Ledford J. &amp; Polotow D. 2022. A revision of the Malagasy crack-leg spiders of the genus Uduba Simon 1880 (Araneae, Udubidae), with description of 35 new species from Madagascar. Proceedings of the California Academy of Sciences (4) 67 (Supplement 2): 1 - 193." type="journal article" year="2022">
Griswold
<emphasis id="8B8A8D0EFFF3FFC1FB690C30FB2BDB23" box="[1219,1278,1043,1069]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2022: 13
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)
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</paragraph>
</keyStep>
<keyStep id="020A49B9FFF3FFC1FF170C72FAA2DA0F" pageId="11" pageNumber="12">
<paragraph id="B941511CFFF3FFC1FF170C72FAA2DBB6" blockId="11.[189,1399,1109,1281]" pageId="11" pageNumber="12">
<keyLead id="020FF229FFF3FFC1FF170C72FAA2DBB6" pageId="11" pageNumber="12">
2. With entire cribellum or vestige; PLS and ALS equal in length; strong scopulae beneath leg tarsi and most of length of metatarsi; males with crack-suture on leg tibiae just anteriad of basalmost pair of ventral spines (
<figureCitation id="21C54D99FFF3FFC1FE370CBAFDD3DBB6" box="[413,518,1182,1208]" captionStart="Fig" captionStartId="59.[189,234,1894,1920]" captionTargetBox="[189,1398,1046,1855]" captionTargetId="figure-180@59.[189,1398,1046,1855]" captionTargetPageId="59" captionText="Fig. 24. Zorodictyna silvadavilae sp. nov., male and female habitus; AB = male holotype (CASENT_9006231; CAS); CD = female paratype (FMNH-0000-056-622; FMNH). A, C. Dorsal view. B, D. Ventral view, the arrow in B points to the tibial crack. Scale bars = 5 mm. Stacked microphotographs by Franklyn Cala Riquelme." figureDoi="http://doi.org/10.5281/zenodo.14001847" httpUri="https://zenodo.org/record/14001847/files/figure.png" pageId="11" pageNumber="12">Fig. 24B</figureCitation>
) ........................................................................3.
<taxonomicName id="7EFE2A9FFFF3FFC1FBED0CB9FAA2DBB6" authority="Strand, 1907" authorityName="Strand" authorityYear="1907" box="[1095,1399,1181,1208]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="8B8A8D0EFFF3FFC1FBED0CB9FB0FDBB6" bold="true" box="[1095,1242,1181,1208]" italics="true" pageId="11" pageNumber="12">Zorodictyna</emphasis>
<bibRefCitation id="DD6F2CEDFFF3FFC1FB4B0CBAFAA2DBB6" author="Strand E." box="[1249,1399,1182,1208]" pageId="11" pageNumber="12" pagination="725 - 748" refId="ref38669" refString="Strand E. 1907. Diagnosen neuer Spinnen aus Madagaskar und Sansibar. Zoologischer Anzeiger 31: 725 - 748." type="journal article" year="1907">Strand, 1907</bibRefCitation>
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</paragraph>
<paragraph id="B941511CFFF3FFC1FF6B0CE7FAA2DA0F" blockId="11.[189,1399,1109,1281]" pageId="11" pageNumber="12">
<keyLead id="020FF229FFF3FFC1FF6B0CE7FAA2DA0F" pageId="11" pageNumber="12"> Ecribellate; PLS much smaller than ALS; scopulae absent or only with weak scopulae beneath leg tarsi; most males without crack-suture on leg tibiae ........................................................................ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="020A49B9FFF3FFC1FF170D0DFAA2D913" pageId="11" pageNumber="12">
<paragraph id="B941511CFFF3FFC1FF170D0DFAA2DA82" blockId="11.[189,1400,1321,1565]" pageId="11" pageNumber="12">
<keyLead id="020FF229FFF3FFC1FF170D0DFAA2DA82" pageId="11" pageNumber="12">
3. Male palpal cymbium with rounded retrolateral paracymbial process (PP) (
<figureCitation id="21C54D99FFF3FFC1FBC50D0DFAD7DA4D" box="[1135,1282,1321,1347]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="11" pageNumber="12">Figs 23BC</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FABA0D0DFABCDA4A" box="[1296,1385,1321,1348]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="11" pageNumber="12">25BC</figureCitation>
); cymbial retrobasal extension (CE) extending at least one-third length of tibia (
<figureCitation id="21C54D99FFF3FFC1FBDD0D69FADDDA66" box="[1143,1288,1357,1384]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="11" pageNumber="12">Figs 23BC</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FAB90D69FABFDA66" box="[1299,1386,1357,1384]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="11" pageNumber="12">25BC</figureCitation>
); female epigyne lateral lobes (LL) lacking teeth (
<figureCitation id="21C54D99FFF3FFC1FC8A0D56FC44DA82" box="[800,913,1394,1420]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="11" pageNumber="12">Figs 23D</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FC360D55FC05DA82" box="[924,976,1393,1420]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="11" pageNumber="12">25D</figureCitation>
) .................................................... 4
</keyLead>
</paragraph>
<paragraph id="B941511CFFF3FFC1FF6B0DB2FAA2D913" blockId="11.[189,1400,1321,1565]" pageId="11" pageNumber="12">
<keyLead id="020FF229FFF3FFC1FF6B0DB2FAA2D913" pageId="11" pageNumber="12">
Male palpal cymbium evenly convex retrolaterally, without paracymbial process (
<bibRefCitation id="DD6F2CEDFFF3FFC1FB110DB2FAA7DABE" author="Griswold C. E." box="[1211,1394,1430,1456]" pageId="11" pageNumber="12" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: figs 2021); cymbial retrobasal extension, short, extending only to tibial apex (
<bibRefCitation id="DD6F2CEDFFF3FFC1FB1D0D9EFAA6DADA" author="Griswold C. E." box="[1207,1395,1466,1492]" pageId="11" pageNumber="12" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: figs 2021); female epigyne lateral lobes with short teeth (
<bibRefCitation id="DD6F2CEDFFF3FFC1FC3C0DFAFB9EDAF7" author="Griswold C. E." box="[918,1099,1502,1529]" pageId="11" pageNumber="12" pagination="1 - 39" refId="ref36151" refString="Griswold C. E. 1993. Investigations into the phylogeny of the lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea). Smithsonian Contributions to Zoology 539: 1 - 39. https: // doi. org / 10.5479 / si. 00810282.539" type="journal article" year="1993">Griswold 1993</bibRefCitation>
: fig. 24) ......................... ......................................................................................................
<taxonomicName id="7EFE2A9FFFF3FFC1FC480E26FAA2D913" authority="(Lenz, 1891)" authorityName="" baseAuthorityName="Lenz" baseAuthorityYear="1891" box="[994,1399,1538,1565]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="8B8A8D0EFFF3FFC1FC480E26FB02D913" bold="true" box="[994,1239,1538,1565]" italics="true" pageId="11" pageNumber="12">Zorodictyna oswaldi</emphasis>
(
<bibRefCitation id="DD6F2CEDFFF3FFC1FB4D0E27FABBD913" author="Lenz H." box="[1255,1390,1539,1565]" pageId="11" pageNumber="12" pagination="161 - 182" refId="ref37136" refString="Lenz H. 1891. Spinnen von Madagascar und Nossibe. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten 9: 161 - 182, pl. 1 - 2." type="journal article" year="1891">Lenz, 1891</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="020A49B9FFF3FFC1FF170E61FAA2D88C" pageId="11" pageNumber="12">
<paragraph id="B941511CFFF3FFC1FF170E61FAA2D9C3" blockId="11.[189,1400,1605,1922]" pageId="11" pageNumber="12">
<keyLead id="020FF229FFF3FFC1FF170E61FAA2D9C3" pageId="11" pageNumber="12">
4. Male palp with basal embolic process (EBP) nearly triangular, embolus lacking crest, tegulum with retromedian longitudinal ridge that extends for most of tegulum length (
<figureCitation id="21C54D99FFF3FFC1FBD50E4DFAC4D98A" box="[1151,1297,1641,1668]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="11" pageNumber="12">Fig. 23AC</figureCitation>
); female epigyne lateral lobes (LL) parallel, median sector (MS) rectangular (
<figureCitation id="21C54D99FFF3FFC1FB8E0EAAFB41D9A6" box="[1060,1172,1678,1704]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="11" pageNumber="12">Fig. 23D</figureCitation>
); female endogyne with copulatory ducts (CD) larger than spermathecae (
<figureCitation id="21C54D99FFF3FFC1FCC00E96FC04D9C2" box="[874,977,1714,1740]" captionStart="Fig" captionStartId="57.[189,232,1636,1662]" captionTargetBox="[219,1348,279,1580]" captionTargetId="figure-12@57.[189,1398,266,1598]" captionTargetPageId="57" captionText="Fig. 23. Zorodictyna almae sp. nov., genitalia drawings; AC = male holotype (CASENT_9082245; FMNH); DE= female paratype (CASENT_9002829; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001845" httpUri="https://zenodo.org/record/14001845/files/figure.png" pageId="11" pageNumber="12">Fig. 23E</figureCitation>
) ..........
<taxonomicName id="7EFE2A9FFFF3FFC1FB980E96FAC0D9C3" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1074,1301,1714,1741]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="almae" status="sp. nov.">
<emphasis id="8B8A8D0EFFF3FFC1FB980E96FAC0D9C3" bold="true" box="[1074,1301,1714,1741]" italics="true" pageId="11" pageNumber="12">Zorodictyna almae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFF3FFC1FAB60E97FAA2D9C3" box="[1308,1399,1715,1741]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="B941511CFFF3FFC1FF6B0EF3FAA2D88C" blockId="11.[189,1400,1605,1922]" pageId="11" pageNumber="12">
<keyLead id="020FF229FFF3FFC1FF6B0EF3FAA2D88C" pageId="11" pageNumber="12">
Male palp with basal embolic process (EBP) slender, crescentic, embolus with subapical distal crest, and tegulum retromedian longitudinal ridge weak, short, extending for less than one-half tegulum length (
<figureCitation id="21C54D99FFF3FFC1FEEE0F3BFE0CD834" box="[324,473,1823,1850]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="11" pageNumber="12">Figs 25AC</figureCitation>
); female epigyne lateral lobes (LL) converging posteriorly, median sector (MS) triangular (
<figureCitation id="21C54D99FFF3FFC1FEC60F67FE0FD850" box="[364,474,1859,1886]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="11" pageNumber="12">Fig. 25D</figureCitation>
); female endogyne with copulatory ducts (CD) and spermathecae equal in size (
<figureCitation id="21C54D99FFF3FFC1FF590F43FE8ED88C" box="[243,347,1895,1922]" captionStart="Fig" captionStartId="61.[189,231,1752,1778]" captionTargetBox="[211,1360,510,1693]" captionTargetId="figure-97@61.[189,1398,507,1714]" captionTargetPageId="61" captionText="Fig. 25. Zorodictyna silvadavilae sp. nov., genitalia drawings; AC = male holotype (CASENT_9006231; CAS); DE = female paratype (CASENT_9006230; USNM).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view. E. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; EBP = embolus basal process; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; PP = paracymbial process; RTA= retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001849" httpUri="https://zenodo.org/record/14001849/files/figure.png" pageId="11" pageNumber="12">Fig. 25E</figureCitation>
) .......................................................................................
<taxonomicName id="7EFE2A9FFFF3FFC1FC470F43FAC0D88C" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1005,1301,1895,1922]" class="Arachnida" family="Udubidae" genus="Zorodictyna" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="silvadavilae" status="sp. nov.">
<emphasis id="8B8A8D0EFFF3FFC1FC470F43FAC0D88C" bold="true" box="[1005,1301,1895,1922]" italics="true" pageId="11" pageNumber="12">Zorodictyna silvadavilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFF3FFC1FAB60F4CFAA2D88C" box="[1308,1399,1896,1922]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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<keyStep id="020A49B9FFF3FFC6FF170F8EFAA2DED9" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<paragraph id="B941511CFFF3FFC6FF170F8EFAA2DE49" blockId="11.[189,1399,1962,2025]" lastBlockId="12.[193,1400,265,471]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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5. Male palp with cymbial retrobasal extension (CE) elongate, extending most of length of tibia, palpal tibia with RTA only, lacking VTA (
<figureCitation id="21C54D99FFF3FFC1FD300FEAFCE6D8E6" box="[666,819,1998,2024]" captionStart="Fig" captionStartId="39.[189,232,1712,1738]" captionTargetBox="[189,1398,459,1674]" captionTargetId="figure-47@39.[189,1398,459,1674]" captionTargetPageId="39" captionText="Fig. 14. Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ARA.Ara.247343; RMCA). A. Male right palp (mirrored), retrolateral view. B. Idem, ventral view. C. Idem, dorsal view. D. Idem, bulbus, retrolateral view. E. Idem, ventral view. F. Idem, prolateral view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AC = 0.5mm; DF = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001825" httpUri="https://zenodo.org/record/14001825/files/figure.png" pageId="11" pageNumber="12">Figs 14AB</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FCEB0FEAFC4FD8E7" box="[833,922,1998,2025]" captionStart="Fig" captionStartId="40.[189,233,1731,1757]" captionTargetBox="[189,1398,478,1693]" captionTargetId="figure-47@40.[189,1398,478,1693]" captionTargetPageId="40" captionText="Fig. 15. Zorascar pasunepipe gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247343; RMCA), micro-CT scans of right male palp (mirrored). A. Retrolateral view. B. Ventral view. C. Prolateral view. D. Bulbus, retrolateral view.E. Bulbus, ventral view.F. Bulbus, apical view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AC = 0.5 mm; DF = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001827" httpUri="https://zenodo.org/record/14001827/files/figure.png" pageId="11" pageNumber="12">15AB</figureCitation>
,
<figureCitation id="21C54D99FFF3FFC1FC030FEAFBD1D8E6" box="[937,1028,1998,2024]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; AB = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); CD = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AB = 2 mm; CE = 0.5 mm. Illustrations AB, E by Alain Reygel; CD by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="11" pageNumber="12">16AB</figureCitation>
); female epigyne median lobe (ML) short, concave medially, not extending to posterior margin, copulatory openings (CO) exposed (
<figureCitation id="21C54D99FFF4FFC6FF5B0909FE55DE49" box="[241,384,301,327]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; AB = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); CD = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AB = 2 mm; CE = 0.5 mm. Illustrations AB, E by Alain Reygel; CD by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="12" pageNumber="13">Figs 16C, E</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FE270909FE15DE49" box="[397,448,301,327]" captionStart="Fig" captionStartId="46.[189,233,1894,1920]" captionTargetBox="[189,1398,1053,1856]" captionTargetId="figure-159@46.[189,1398,1053,1859]" captionTargetPageId="46" captionText="Fig. 18. Zorascar pasunepomme gen. et sp. nov. holotype female (BE_RMCA_ARA.Ara.247346; RMCA), genitalia. A. Epigyne, ventral view. B. Endogyne, cleared, dorsal view. C. Idem, dorso-anterior view. D. Idem, dorso-posterior view. Scale bars = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001835" httpUri="https://zenodo.org/record/14001835/files/figure.png" pageId="12" pageNumber="13">18A</figureCitation>
) ............................................................................................6.
<taxonomicName id="7EFE2A9FFFF4FFC6FB3D0909FAD1DE49" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1175,1284,301,327]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8B8A8D0EFFF4FFC6FB3D0909FAD1DE49" bold="true" box="[1175,1284,301,327]" italics="true" pageId="12" pageNumber="13">Zorascar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFF4FFC6FAA60909FAA2DE49" box="[1292,1399,301,327]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="B941511CFFF4FFC6FF6B0975FAA2DED9" blockId="12.[193,1400,265,471]" pageId="12" pageNumber="13">
<keyLead id="020FF229FFF4FFC6FF6B0975FAA2DED9" pageId="12" pageNumber="13">
Male palp with cymbial retrobasal extension (CE) shorter, extending to or just past tibial apex, palpal tibia with both VTA and RTA (
<figureCitation id="21C54D99FFF4FFC6FDFE0950FD0DDE81" box="[596,728,372,399]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="12" pageNumber="13">Figs 5AC</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FD4F0950FCFBDE80" box="[741,814,372,398]" captionStart="Fig" captionStartId="19.[188,230,1787,1813]" captionTargetBox="[188,1399,479,1749]" captionTargetId="figure-89@19.[188,1399,479,1749]" captionTargetPageId="19" captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AC = 0.43 mm; DF = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick." figureDoi="http://doi.org/10.5281/zenodo.14001804" httpUri="https://zenodo.org/record/14001804/files/figure.png" pageId="12" pageNumber="13">6AC</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FC900951FC56DE81" box="[826,899,373,399]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.AC = male holotype (CASENT_9016063; MNHN). DE = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations AC by Jenny Speckels; DE by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="12" pageNumber="13">9AC</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FC250951FC30DE81" box="[911,997,373,399]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; AC= male holotype (CASENT_9016117; CAS); DE = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="12" pageNumber="13">11AC</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FC580951FB98DE81" box="[1010,1101,373,399]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; AF = male holotype (CASENT_9016117; CAS); GH= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AD = 0.5 mm; EF = 0.15 mm; GH = 0.1 mm. SEM images AF by Charles Griswold and Darrell Ubick, GH by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="12" pageNumber="13">12AD</figureCitation>
); female epigyne median lobe (ML) extends to posterior margin, separating lateral lobes (LL) and covering copulatory openings (CO) (
<figureCitation id="21C54D99FFF4FFC6FE9F0998FE41DED8" box="[309,404,444,470]" captionStart="Fig" captionStartId="17.[189,231,1892,1918]" captionTargetBox="[189,1398,962,1853]" captionTargetId="figure-276@17.[189,1398,962,1853]" captionTargetPageId="17" captionText="Fig. 4. Tabiboka milleri gen. et sp. nov.A. Female paratype (CASENT_9029871; CAS) from Ranomafana (Photo © N. Scharff). B. Female paratype (CASENT_9016098; CAS), epigyne, ventral view. C. Female paratype (CASENT_9016094; CAS), trochanter 1, ventral view. D. Idem, trochanter 4. (No scales provided). Stacked microphotographs by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001796" httpUri="https://zenodo.org/record/14001796/files/figure.png" pageId="12" pageNumber="13">Figs 4B</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FE0B0998FE12DED9" box="[417,455,444,471]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="12" pageNumber="13">5D</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FE790999FDC9DED9" box="[467,540,445,471]" captionStart="Fig" captionStartId="20.[189,232,1822,1848]" captionTargetBox="[189,1398,461,1784]" captionTargetId="figure-79@20.[189,1398,461,1784]" captionTargetPageId="20" captionText="Fig. 7. Tabiboka milleri gen. nov. et sp. nov. SEM of female genitalia, AB, EF = female paratype (CASENT_9016094; CAS); CD = female paratype (CASENT_9035866; CAS). A. Epigyne, ventral view. B. Idem, ventro-posterior view. C. Endogyne, dorsal view. D. Idem, slightly anterior view. E. Spermatheca, right dorsal view. F. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: AE = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila." figureDoi="http://doi.org/10.5281/zenodo.14001806" httpUri="https://zenodo.org/record/14001806/files/figure.png" pageId="12" pageNumber="13">7AB</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FD820998FD9BDED8" box="[552,590,444,470]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.AC = male holotype (CASENT_9016063; MNHN). DE = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations AC by Jenny Speckels; DE by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="12" pageNumber="13">9D</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FDF10998FD5ADED8" box="[603,655,444,470]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; AC= male holotype (CASENT_9016117; CAS); DE = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="12" pageNumber="13">11D</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FD310998FD1ADED8" box="[667,719,444,470]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; AF = male holotype (CASENT_9016117; CAS); GH= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AD = 0.5 mm; EF = 0.15 mm; GH = 0.1 mm. SEM images AF by Charles Griswold and Darrell Ubick, GH by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="12" pageNumber="13">12G</figureCitation>
) .......................................................7.
<taxonomicName id="7EFE2A9FFFF4FFC6FB3F0998FAD1DED9" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[1173,1284,444,471]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8B8A8D0EFFF4FFC6FB3F0998FAD1DED9" bold="true" box="[1173,1284,444,471]" italics="true" pageId="12" pageNumber="13">Tabiboka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90B93075FFF4FFC6FAA60999FAA2DED9" box="[1292,1399,445,471]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="020A49B9FFF4FFC6FF1709DAFAA2DDC2" pageId="12" pageNumber="13">
<paragraph id="B941511CFFF4FFC6FF1709DAFAA2DD6F" blockId="12.[189,1399,510,716]" pageId="12" pageNumber="13">
<keyLead id="020FF229FFF4FFC6FF1709DAFAA2DD6F" pageId="12" pageNumber="13">
6. Female epigyne median lobe (ML) broad and shallowly concave medially, copulatory openings (CO) exposed anteromedially (
<figureCitation id="21C54D99FFF4FFC6FDA00A06FDA4DD32" box="[522,625,546,572]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; AB = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); CD = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AB = 2 mm; CE = 0.5 mm. Illustrations AB, E by Alain Reygel; CD by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="12" pageNumber="13">Fig. 16C</figureCitation>
), spermathecae elongate, curved, eggplant-shaped (
<figureCitation id="21C54D99FFF4FFC6FB150A06FAFFDD32" box="[1215,1322,546,572]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; AB = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); CD = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AB = 2 mm; CE = 0.5 mm. Illustrations AB, E by Alain Reygel; CD by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="12" pageNumber="13">Fig. 16D</figureCitation>
); male palp as in figures (
<figureCitation id="21C54D99FFF4FFC6FE6A0A62FD9BDD6E" box="[448,590,582,608]" captionStart="Fig" captionStartId="39.[189,232,1712,1738]" captionTargetBox="[189,1398,459,1674]" captionTargetId="figure-47@39.[189,1398,459,1674]" captionTargetPageId="39" captionText="Fig. 14. Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ARA.Ara.247343; RMCA). A. Male right palp (mirrored), retrolateral view. B. Idem, ventral view. C. Idem, dorsal view. D. Idem, bulbus, retrolateral view. E. Idem, ventral view. F. Idem, prolateral view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AC = 0.5mm; DF = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001825" httpUri="https://zenodo.org/record/14001825/files/figure.png" pageId="12" pageNumber="13">Figs 14AF</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FDF30A62FD7EDD6F" box="[601,683,582,609]" captionStart="Fig" captionStartId="40.[189,233,1731,1757]" captionTargetBox="[189,1398,478,1693]" captionTargetId="figure-47@40.[189,1398,478,1693]" captionTargetPageId="40" captionText="Fig. 15. Zorascar pasunepipe gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247343; RMCA), micro-CT scans of right male palp (mirrored). A. Retrolateral view. B. Ventral view. C. Prolateral view. D. Bulbus, retrolateral view.E. Bulbus, ventral view.F. Bulbus, apical view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AC = 0.5 mm; DF = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001827" httpUri="https://zenodo.org/record/14001827/files/figure.png" pageId="12" pageNumber="13">15AF</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FD1D0A62FCD8DD6E" box="[695,781,582,608]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; AB = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); CD = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AB = 2 mm; CE = 0.5 mm. Illustrations AB, E by Alain Reygel; CD by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="12" pageNumber="13">16AB</figureCitation>
) .....................
<taxonomicName id="7EFE2A9FFFF4FFC6FC680A62FB6ADD6E" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[962,1215,582,608]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pasunepipe" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFF4FFC6FC680A62FB6ADD6E" bold="true" box="[962,1215,582,608]" italics="true" pageId="12" pageNumber="13">Zorascar pasunepipe</emphasis>
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</paragraph>
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Female epigyne median lobe (ML) narrow and deeply concave medially, copulatory openings (CO) exposed laterad of median lobe (
<figureCitation id="21C54D99FFF4FFC6FDDF0AAAFD30DDA6" box="[629,741,654,680]" captionStart="Fig" captionStartId="43.[189,232,1580,1606]" captionTargetBox="[220,1354,460,1507]" captionTargetId="figure-30@43.[189,1398,448,1544]" captionTargetPageId="43" captionText="Fig. 16. Drawings of genitalia; AB = Zorascar pasunepipe gen. et sp. nov., holotype male (BE_RMCA_ ARA.Ara.247343; RMCA); CD = Z. pasunepipe, paratype female (CASENT_9016138; CAS); E = Zorascar pasunepomme gen. et sp. nov., holotype female (BE_RMCA_ARA.Ara.247346; RMCA). A. Male right palp (mirrored), ventral view. B. Idem, retrolateral view. C, E. Epigyne, ventral view. D. Endogyne, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; MA = median apophysis; ML = median lobe; MS = median sector; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe. Scale bars: AB = 2 mm; CE = 0.5 mm. Illustrations AB, E by Alain Reygel; CD by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001829" httpUri="https://zenodo.org/record/14001829/files/figure.png" pageId="12" pageNumber="13">Figs 16E</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FD590AAAFCFDDDA6" box="[755,808,654,680]" captionStart="Fig" captionStartId="46.[189,233,1894,1920]" captionTargetBox="[189,1398,1053,1856]" captionTargetId="figure-159@46.[189,1398,1053,1859]" captionTargetPageId="46" captionText="Fig. 18. Zorascar pasunepomme gen. et sp. nov. holotype female (BE_RMCA_ARA.Ara.247346; RMCA), genitalia. A. Epigyne, ventral view. B. Endogyne, cleared, dorsal view. C. Idem, dorso-anterior view. D. Idem, dorso-posterior view. Scale bars = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001835" httpUri="https://zenodo.org/record/14001835/files/figure.png" pageId="12" pageNumber="13">18A</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FC9D0AAAFC45DDA6" box="[823,912,654,680]" captionStart="Fig" captionStartId="47.[189,232,1570,1596]" captionTargetBox="[189,1398,265,1533]" captionTargetId="figure-11@47.[189,1398,265,1533]" captionTargetPageId="47" captionText="Fig. 19. Zorascar pasunepomme gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247346; RMCA), epigyne, CT-scan images. A. ventral view. B. ventro-anterior view. C. dorsal view. D. dorso-anterior view. E. idem, detail.F. Idem, dorso-lateral view. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; ML = median lobe; MS = median sector; S = spermatheca. Scale bars = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001837" httpUri="https://zenodo.org/record/14001837/files/figure.png" pageId="12" pageNumber="13">19AB</figureCitation>
), spermathecae stout, rounded, reniform (
<figureCitation id="21C54D99FFF4FFC6FF5E0A96FE52DDC2" box="[244,391,690,716]" captionStart="Fig" captionStartId="46.[189,233,1894,1920]" captionTargetBox="[189,1398,1053,1856]" captionTargetId="figure-159@46.[189,1398,1053,1859]" captionTargetPageId="46" captionText="Fig. 18. Zorascar pasunepomme gen. et sp. nov. holotype female (BE_RMCA_ARA.Ara.247346; RMCA), genitalia. A. Epigyne, ventral view. B. Endogyne, cleared, dorsal view. C. Idem, dorso-anterior view. D. Idem, dorso-posterior view. Scale bars = 0.2 mm. Stacked microphotographs by Arnaud Henrard and Rudy Jocqué." figureDoi="http://doi.org/10.5281/zenodo.14001835" httpUri="https://zenodo.org/record/14001835/files/figure.png" pageId="12" pageNumber="13">Figs 18BD</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FE390A96FE32DDC2" box="[403,487,690,716]" captionStart="Fig" captionStartId="47.[189,232,1570,1596]" captionTargetBox="[189,1398,265,1533]" captionTargetId="figure-11@47.[189,1398,265,1533]" captionTargetPageId="47" captionText="Fig. 19. Zorascar pasunepomme gen. et sp. nov., holotype (BE_RMCA_ARA.Ara.247346; RMCA), epigyne, CT-scan images. A. ventral view. B. ventro-anterior view. C. dorsal view. D. dorso-anterior view. E. idem, detail.F. Idem, dorso-lateral view. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; LP = lateral process; ML = median lobe; MS = median sector; S = spermatheca. Scale bars = 0.2 mm. 3D micro-CT scans by Arnaud Henrard." figureDoi="http://doi.org/10.5281/zenodo.14001837" httpUri="https://zenodo.org/record/14001837/files/figure.png" pageId="12" pageNumber="13">19CF</figureCitation>
); male unknown ................................
<taxonomicName id="7EFE2A9FFFF4FFC6FC360A96FB6ADDC2" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[924,1215,690,716]" class="Arachnida" family="Udubidae" genus="Zorascar" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pasunepomme" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFF4FFC6FC360A96FB6ADDC2" bold="true" box="[924,1215,690,716]" italics="true" pageId="12" pageNumber="13">Zorascar pasunepomme</emphasis>
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</paragraph>
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<keyStep id="020A49B9FFF4FFC6FF170AD0FAA2DCCC" pageId="12" pageNumber="13">
<paragraph id="B941511CFFF4FFC6FF170AD0FAA2DC58" blockId="12.[189,1399,756,962]" pageId="12" pageNumber="13">
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7. Male palp with conductor base originating anteriorly, hidden behind embolus or paraconductor (Pc), embolus short, arising retrolaterally near 270° (
<figureCitation id="21C54D99FFF4FFC6FCAD0B3CFC5EDC3D" box="[775,907,792,819]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="12" pageNumber="13">Figs 5AC</figureCitation>
;
<figureCitation id="21C54D99FFF4FFC6FC3C0B3CFC0CDC3C" box="[918,985,792,818]" captionStart="Fig" captionStartId="19.[188,230,1787,1813]" captionTargetBox="[188,1399,479,1749]" captionTargetId="figure-89@19.[188,1399,479,1749]" captionTargetPageId="19" captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AC = 0.43 mm; DF = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick." figureDoi="http://doi.org/10.5281/zenodo.14001804" httpUri="https://zenodo.org/record/14001804/files/figure.png" pageId="12" pageNumber="13">6DF</figureCitation>
, 8F, 9AC); female epigyne median lobe (ML) heart-shaped, sides rounded (
<figureCitation id="21C54D99FFF4FFC6FD6A0B18FCCADC58" box="[704,799,828,854]" captionStart="Fig" captionStartId="17.[189,231,1892,1918]" captionTargetBox="[189,1398,962,1853]" captionTargetId="figure-276@17.[189,1398,962,1853]" captionTargetPageId="17" captionText="Fig. 4. Tabiboka milleri gen. et sp. nov.A. Female paratype (CASENT_9029871; CAS) from Ranomafana (Photo © N. Scharff). B. Female paratype (CASENT_9016098; CAS), epigyne, ventral view. C. Female paratype (CASENT_9016094; CAS), trochanter 1, ventral view. D. Idem, trochanter 4. (No scales provided). Stacked microphotographs by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001796" httpUri="https://zenodo.org/record/14001796/files/figure.png" pageId="12" pageNumber="13">Figs 4B</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FC860B1FFC87DC58" box="[812,850,827,854]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="12" pageNumber="13">5D</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FCF50B18FC50DC58" box="[863,901,828,854]" captionStart="Fig" captionStartId="20.[189,232,1822,1848]" captionTargetBox="[189,1398,461,1784]" captionTargetId="figure-79@20.[189,1398,461,1784]" captionTargetPageId="20" captionText="Fig. 7. Tabiboka milleri gen. nov. et sp. nov. SEM of female genitalia, AB, EF = female paratype (CASENT_9016094; CAS); CD = female paratype (CASENT_9035866; CAS). A. Epigyne, ventral view. B. Idem, ventro-posterior view. C. Endogyne, dorsal view. D. Idem, slightly anterior view. E. Spermatheca, right dorsal view. F. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: AE = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila." figureDoi="http://doi.org/10.5281/zenodo.14001806" httpUri="https://zenodo.org/record/14001806/files/figure.png" pageId="12" pageNumber="13">7A</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FC3B0B18FC62DC58" box="[913,951,828,854]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.AC = male holotype (CASENT_9016063; MNHN). DE = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations AC by Jenny Speckels; DE by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="12" pageNumber="13">9D</figureCitation>
) ....................................................... 8
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<paragraph id="B941511CFFF4FFC6FF6B0B44FAA2DCCC" blockId="12.[189,1399,756,962]" pageId="12" pageNumber="13">
<keyLead id="020FF229FFF4FFC6FF6B0B44FAA2DCCC" pageId="12" pageNumber="13">
Male palp conductor originating medially, near embolus base, not hidden, embolus elongate, arising retrobasally near 200° (
<figureCitation id="21C54D99FFF4FFC6FDAD0BA7FD48DC93" box="[519,669,899,925]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; AC= male holotype (CASENT_9016117; CAS); DE = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="12" pageNumber="13">Figs 11AC</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FD010BA7FCFDDC93" box="[683,808,899,925]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; AF = male holotype (CASENT_9016117; CAS); GH= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AD = 0.5 mm; EF = 0.15 mm; GH = 0.1 mm. SEM images AF by Charles Griswold and Darrell Ubick, GH by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="12" pageNumber="13">12AC, E</figureCitation>
); female epigyne median lobe (ML) rectangular, narrowest in middle or with sides straight (
<figureCitation id="21C54D99FFF4FFC6FD490B83FC87DCCF" box="[739,850,935,961]" captionStart="Fig" captionStartId="31.[189,231,1719,1745]" captionTargetBox="[211,1371,542,1665]" captionTargetId="figure-76@31.[189,1398,527,1680]" captionTargetPageId="31" captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; AC= male holotype (CASENT_9016117; CAS); DE = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001819" httpUri="https://zenodo.org/record/14001819/files/figure.png" pageId="12" pageNumber="13">Figs 11D</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FCF40B83FC47DCCF" box="[862,914,935,961]" captionStart="Fig" captionStartId="32.[189,232,1717,1743]" captionTargetBox="[277,1311,265,1679]" captionTargetId="figure-18@32.[277,1311,265,1679]" captionTargetPageId="32" captionText="Fig. 12. Tabiboka polotowae gen. et sp. nov., SEM of male and female genitalia; AF = male holotype (CASENT_9016117; CAS); GH= female paratype (CASENT_9029889; CAS).A. Right palp (mirrored), prolateral, slightly ventral view. B. Idem, ventral view. C. Idem, retrolateral view. D. Idem, dorsal view. E. Bulb, ventral view. F. RTA, ventral view. G. Epigyne, ventral view. H. Endogyne, dorsal, slightly anterior view.Abbreviations: C = conductor; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AD = 0.5 mm; EF = 0.15 mm; GH = 0.1 mm. SEM images AF by Charles Griswold and Darrell Ubick, GH by Daniele Polotow." figureDoi="http://doi.org/10.5281/zenodo.14001821" httpUri="https://zenodo.org/record/14001821/files/figure.png" pageId="12" pageNumber="13">12G</figureCitation>
) ......
<taxonomicName id="7EFE2A9FFFF4FFC6FC640B83FB6ADCCC" authorityName="Henrard &amp; Griswold &amp; Jocqué" authorityYear="2024" box="[974,1215,935,962]" class="Arachnida" family="Udubidae" genus="Tabiboka" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="polotowae" status="gen. et sp. nov.">
<emphasis id="8B8A8D0EFFF4FFC6FC640B83FB6ADCCC" bold="true" box="[974,1215,935,962]" italics="true" pageId="12" pageNumber="13">Tabiboka polotowae</emphasis>
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8. Male palpal tibia with elongate retrobasal process (RTA), embolus broad, median apophysis (MA) short, apex far proximad of embolus apex (
<figureCitation id="21C54D99FFF4FFC6FD450C29FCA1DB26" box="[751,884,1037,1064]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="12" pageNumber="13">Figs 5AC</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FC280C29FC1CDB29" box="[898,969,1037,1063]" captionStart="Fig" captionStartId="19.[188,230,1787,1813]" captionTargetBox="[188,1399,479,1749]" captionTargetId="figure-89@19.[188,1399,479,1749]" captionTargetPageId="19" captionText="Fig. 6. Tabibokamillerigen.nov.etsp. nov.SEM ofmale rightpalp(mirrored), paratype(CASENT_9016094; CAS). A. Palp, ventral, slightly retrolateral view.B. Idem, retrolateral view.C. Idem, dorsal view.D. Bulb, prolateral view. E. Idem, ventral view. F. Idem, retrolateral view. Abbreviations: BC = basal concavity of the RTA; C = conductor; CE = cymbial retrobasal extension; E = embolus; MA = median apophysis; Pc = paraconductor; RTA = retrolateral tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars: AC = 0.43 mm; DF = 0.2 mm. SEM images by Charles Griswold and Darrell Ubick." figureDoi="http://doi.org/10.5281/zenodo.14001804" httpUri="https://zenodo.org/record/14001804/files/figure.png" pageId="12" pageNumber="13">6DF</figureCitation>
); female epigyne median lobe (ML) narrow anteriorly, anterior width less than one-half total width (
<figureCitation id="21C54D99FFF4FFC6FC4C0C15FB9DDB45" box="[998,1096,1073,1099]" captionStart="Fig" captionStartId="17.[189,231,1892,1918]" captionTargetBox="[189,1398,962,1853]" captionTargetId="figure-276@17.[189,1398,962,1853]" captionTargetPageId="17" captionText="Fig. 4. Tabiboka milleri gen. et sp. nov.A. Female paratype (CASENT_9029871; CAS) from Ranomafana (Photo © N. Scharff). B. Female paratype (CASENT_9016098; CAS), epigyne, ventral view. C. Female paratype (CASENT_9016094; CAS), trochanter 1, ventral view. D. Idem, trochanter 4. (No scales provided). Stacked microphotographs by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001796" httpUri="https://zenodo.org/record/14001796/files/figure.png" pageId="12" pageNumber="13">Figs 4B</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FBFC0C15FBA8DB42" box="[1110,1149,1073,1100]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="12" pageNumber="13">5D</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FB200C15FB64DB45" box="[1162,1201,1073,1099]" captionStart="Fig" captionStartId="20.[189,232,1822,1848]" captionTargetBox="[189,1398,461,1784]" captionTargetId="figure-79@20.[189,1398,461,1784]" captionTargetPageId="20" captionText="Fig. 7. Tabiboka milleri gen. nov. et sp. nov. SEM of female genitalia, AB, EF = female paratype (CASENT_9016094; CAS); CD = female paratype (CASENT_9035866; CAS). A. Epigyne, ventral view. B. Idem, ventro-posterior view. C. Endogyne, dorsal view. D. Idem, slightly anterior view. E. Spermatheca, right dorsal view. F. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: AE = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila." figureDoi="http://doi.org/10.5281/zenodo.14001806" httpUri="https://zenodo.org/record/14001806/files/figure.png" pageId="12" pageNumber="13">7A</figureCitation>
), endogyne with each spermatheca as broad as long (
<figureCitation id="21C54D99FFF4FFC6FD3A0C71FD3ADB7E" box="[656,751,1109,1136]" captionStart="Fig" captionStartId="18.[189,233,1787,1813]" captionTargetBox="[219,1365,478,1733]" captionTargetId="figure-82@18.[189,1398,445,1749]" captionTargetPageId="18" captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels." figureDoi="http://doi.org/10.5281/zenodo.14001800" httpUri="https://zenodo.org/record/14001800/files/figure.png" pageId="12" pageNumber="13">Figs 5E</figureCitation>
,
<figureCitation id="21C54D99FFF4FFC6FD510C71FCEADB61" box="[763,831,1109,1135]" captionStart="Fig" captionStartId="20.[189,232,1822,1848]" captionTargetBox="[189,1398,461,1784]" captionTargetId="figure-79@20.[189,1398,461,1784]" captionTargetPageId="20" captionText="Fig. 7. Tabiboka milleri gen. nov. et sp. nov. SEM of female genitalia, AB, EF = female paratype (CASENT_9016094; CAS); CD = female paratype (CASENT_9035866; CAS). A. Epigyne, ventral view. B. Idem, ventro-posterior view. C. Endogyne, dorsal view. D. Idem, slightly anterior view. E. Spermatheca, right dorsal view. F. Idem, detail. Abbreviations: CD = copulatory duct; CO = copulatory opening; FD = fertilisation duct; LL = lateral lobe; ML = median lobe; S = spermatheca. Scale bars: AE = 0.1 mm; F = 20 µm. SEM images by Diana Silva Dávila." figureDoi="http://doi.org/10.5281/zenodo.14001806" httpUri="https://zenodo.org/record/14001806/files/figure.png" pageId="12" pageNumber="13">7CE</figureCitation>
) ......................
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<emphasis id="8B8A8D0EFFF4FFC6FC530C70FB15DB61" bold="true" box="[1017,1216,1108,1135]" italics="true" pageId="12" pageNumber="13">Tabiboka milleri</emphasis>
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Male palpal tibia with short, blunt retromedian process (RTA), embolus slender; median apophysis (MA) elongate, cruciform, apex extends to embolus apex (Figs 8CF, 9AC); female epigyne median lobe (ML) broad anteriorly, anterior width greater than one-half total width (
<figureCitation id="21C54D99FFF4FFC6FBF30CE4FB63DBD5" box="[1113,1206,1216,1243]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.AC = male holotype (CASENT_9016063; MNHN). DE = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations AC by Jenny Speckels; DE by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="12" pageNumber="13">Fig. 9D</figureCitation>
), endogyne with each spermatheca narrow, longer than broad (
<figureCitation id="21C54D99FFF4FFC6FD550CC0FC82DBF1" box="[767,855,1252,1279]" captionStart="Fig" captionStartId="27.[189,231,1595,1621]" captionTargetBox="[260,1379,284,1517]" captionTargetId="figure-11@27.[189,1398,265,1558]" captionTargetPageId="27" captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.AC = male holotype (CASENT_9016063; MNHN). DE = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations AC by Jenny Speckels; DE by Charles Griswold." figureDoi="http://doi.org/10.5281/zenodo.14001810" httpUri="https://zenodo.org/record/14001810/files/figure.png" pageId="12" pageNumber="13">Fig. 9E</figureCitation>
) ...................
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<emphasis id="8B8A8D0EFFF4FFC6FC500CC0FB6ADBF1" bold="true" box="[1018,1215,1252,1279]" italics="true" pageId="12" pageNumber="13">Tabiboka milloti</emphasis>
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