<documentid="EE7FA24D4CD10063C7090CC8F22F1191"ID-CLB-Dataset="303592"ID-DOI="10.50826/bnmnszool.49.2_57"ID-GBIF-Dataset="90b4626e-63df-4241-a3cd-abe4d57452d2"ID-ISSN="2434-091X"ID-Zenodo-Dep="13826588"IM.bibliography_approvedBy="karina"IM.illustrations_approvedBy="karina"IM.materialsCitations_approvedBy="karina"IM.metadata_approvedBy="karina"IM.taxonomicNames_approvedBy="carolina"IM.treatmentCitations_approvedBy="karina"IM.treatments_approvedBy="karina"checkinTime="1718644530338"checkinUser="felipe"docAuthor="Kumar, Deepthi S & Sankar, Manimuthu M"docDate="2023"docId="37193B620F0FFFB4FF76FB9A992EFD99"docLanguage="en"docName="BullNatlMusNatSciSerA.49.2.57-74.pdf"docOrigin="Bulletin of the National Museum of Nature and Science. Series A, Zoology 49 (2)"docSource="http://dx.doi.org/10.4103/jpbs.jpbs_546_23"docStyle="DocumentStyle:A6FFB80BFE6276686563735C80A89639.2:BullNatlMusNatSciSerA.2021-.journal_article.english.affilindex"docStyleId="A6FFB80BFE6276686563735C80A89639"docStyleName="BullNatlMusNatSciSerA.2021-.journal_article.english.affilindex"docStyleVersion="2"docTitle="Nematus yokohamensis"docType="treatment"docVersion="7"lastPageNumber="67"masterDocId="CB20431A0F0AFFBFFFC8FFF09D60FFF4"masterDocTitle="Taxonomic Notes and New Distribution and Host Plant Records for Sawflies and Woodwasps (Hymenoptera, Symphyta) of Japan VIII"masterLastPageNumber="74"masterPageNumber="57"pageNumber="62"updateTime="1736767391982"updateUser="carolina"zenodo-license-document="CC-BY-4.0">
<mods:titleid="CE0F0914F9DACFBB02DD8B0D8F9B4164">Taxonomic Notes and New Distribution and Host Plant Records for Sawflies and Woodwasps (Hymenoptera, Symphyta) of Japan VIII</mods:title>
<bibRefCitationid="DB21F7850F0FFFBAFE18FB9A9F03FB76"author="Konow, F. W."box="[464,611,1130,1154]"pageId="5"pageNumber="62"pagination="50 - 57"refId="ref10226"refString="Konow, F. W. 1895. Neue oder wenig bekannte Tenthre- diniden und eine analytische Ubersicht der Gattung Holcocneme Knw. Termeszetrajzi Fuzetek 18: 50 - 57."type="journal article"year="1895">Konow, 1895</bibRefCitation>
<figureCitationid="278B96F10F0FFFBAFEA1FB4C9CA0FB26"box="[361,448,1212,1234]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStart-3="Fig"captionStartId-0="6.[163,196,842,864]"captionStartId-1="7.[163,196,1263,1285]"captionStartId-2="8.[163,196,1308,1330]"captionStartId-3="9.[163,196,1002,1024]"captionTargetBox-0="[164,1216,215,801]"captionTargetBox-1="[159,1216,215,1228]"captionTargetBox-2="[159,1213,215,1273]"captionTargetBox-3="[159,1216,215,967]"captionTargetId-0="figure-391@6.[157,1218,213,808]"captionTargetId-1="figure-121@7.[157,1218,213,1230]"captionTargetId-2="figure-189@8.[157,1218,213,1275]"captionTargetId-3="figure-321@9.[157,1218,213,969]"captionTargetPageId-0="6"captionTargetPageId-1="7"captionTargetPageId-2="8"captionTargetPageId-3="9"captionText-0="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."captionText-1="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."captionText-2="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."captionText-3="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi-0="http://doi.org/10.5281/zenodo.13826606"figureDoi-1="http://doi.org/10.5281/zenodo.13826608"figureDoi-2="http://doi.org/10.5281/zenodo.13826612"figureDoi-3="http://doi.org/10.5281/zenodo.13826614"httpUri-0="https://zenodo.org/record/13826606/files/figure.png"httpUri-1="https://zenodo.org/record/13826608/files/figure.png"httpUri-2="https://zenodo.org/record/13826612/files/figure.png"httpUri-3="https://zenodo.org/record/13826614/files/figure.png"pageId="5"pageNumber="62">Figs. 6–9</figureCitation>
<treatmentCitationid="3E11AC650F0FFFBAFE08FB1C9F0FFAF6"author="Konow, F. W."box="[448,623,1260,1282]"page="54"pageId="5"pageNumber="62"year="1895">
<bibRefCitationid="DB21F7850F0FFFBAFE08FB1C9F0FFAF6"author="Konow, F. W."box="[448,623,1260,1282]"pageId="5"pageNumber="62"pagination="50 - 57"refId="ref10226"refString="Konow, F. W. 1895. Neue oder wenig bekannte Tenthre- diniden und eine analytische Ubersicht der Gattung Holcocneme Knw. Termeszetrajzi Fuzetek 18: 50 - 57."type="journal article"year="1895">Konow, 1895: 54</bibRefCitation>
</treatmentCitation>
, 56;
<treatmentCitationid="3E11AC650F0FFFBAFF6FFAFA9C25FAEB"author="Konow, F. W."box="[167,325,1289,1311]"page="62"pageId="5"pageNumber="62"year="1905">
<bibRefCitationid="DB21F7850F0FFFBAFF6FFAFA9C25FAEB"author="Konow, F. W."box="[167,325,1289,1311]"pageId="5"pageNumber="62"pagination="1 - 176"refId="ref10257"refString="Konow, F. W. 1905. Hymenoptera. Fam. Tenthredinidae. Genera Insectorum 29: 1 - 176."type="journal article"year="1905">Konow, 1905: 62</bibRefCitation>
<bibRefCitationid="DB21F7850F0FFFBAFE87FAF99C93FAEB"author="Enslin, E."box="[335,499,1289,1311]"pageId="5"pageNumber="62"pagination="315 - 317"refId="ref9785"refString="Enslin, E. 1910. Eine neue Holcocneme aus Deutschland nebst einer Bestimmungstabelle der bisher bekannten Arten. (Hym.). Deutsche Entomologische Zeitschrift [1910] (3): 315 - 317."type="journal article"year="1910">Enslin, 1910: 316</bibRefCitation>
<bibRefCitationid="DB21F7850F0FFFBAFE59FAD69F31FAC8"author="Takeuchi, K."box="[401,593,1318,1340]"pageId="5"pageNumber="62"pagination="150 - 164"refId="ref11317"refString="Takeuchi, K. 1936. Some sawflies from Sado Island. Ten- thredo 1: 150 - 164."type="journal article"year="1936">Takeuchi, 1936: 162</bibRefCitation>
</treatmentCitation>
;
<treatmentCitationid="3E11AC650F0FFFBAFD94FAD69CFCFAAD"author="Oehlke, J. & J. Wudowenz"page="420"pageId="5"pageNumber="62"year="1984">
<bibRefCitationid="DB21F7850F0FFFBAFD94FAD69CFCFAAD"author="Oehlke, J. & J. Wudowenz"pageId="5"pageNumber="62"pagination="363 - 420"refId="ref10575"refString="Oehlke, J. and J. Wudowenz 1984. Katalog der in den Sammlungen der Abteilung Taxonomie der Insekten des Institutes fur Pflanzenschutzforschung, Bereich Eberswalde (ehemals Deutsches Entomologisches Institut), aufbewahrten Typen - XXII (Hymenoptera: Symphyta). Beitrage zur Entomologie 34 (2): 363 - 420."type="journal article"year="1984">Oehlke and Wudowenz, 1984: 420</bibRefCitation>
<bibRefCitationid="DB21F7850F0FFFBAFC7BFF209913FF12"author="Takeuchi, K."box="[947,1139,208,230]"pageId="5"pageNumber="62"refId="ref11364"refString="Takeuchi, K. 1952. A Generic Classification of the Japa- nese Tenthredinidae (Hymenoptera: Symphyta). 90 pp. Kyoto."type="book"year="1952">Takeuchi, 1952: 68</bibRefCitation>
<bibRefCitationid="DB21F7850F0FFFBAFB4DFF209E53FEF6"author="Okutani, T."pageId="5"pageNumber="62"pagination="75 - 80"refId="ref10631"refString="Okutani, T. 1954. Studies on Symphyta (I). Symphyta of Sasayama with description of a new species. The Sci- ence Reports of the Hyogo University of Agriculture, Agricultural Biology, Sasayama 1 (2): 75 - 80."type="journal article"year="1954">Okutani, 1954: 80</bibRefCitation>
</treatmentCitation>
;
<treatmentCitationid="3E11AC650F0FFFBAFC89FF1D9962FEF7"author="Zinovjev, A. G."box="[833,1026,237,259]"page="627"pageId="5"pageNumber="62"year="1978">
<bibRefCitationid="DB21F7850F0FFFBAFC89FF1D9962FEF7"author="Zinovjev, A. G."box="[833,1026,237,259]"pageId="5"pageNumber="62"pagination="625 - 635"refId="ref11645"refString="Zinovjev, A. G. 1978. Description of a new subgenus Paranematus subgen. n. for the group of species related to Nematus wahlbergi Thomson (Hymenoptera, Tenth- redinidae) with a review of species from the European part of the USSR. Entomologicheskoe Obozrenie 57 (3): 625 - 635 (in Russian, abstract in English)."type="journal article"year="1978">Zinovjev, 1978: 627</bibRefCitation>
</treatmentCitation>
;
<treatmentCitationid="3E11AC650F0FFFBAFBC7FF1D99B1FEF6"author="Zinovjev, A. G."box="[1039,1233,237,259]"page="430"pageId="5"pageNumber="62"year="1979">
<bibRefCitationid="DB21F7850F0FFFBAFBC7FF1D99B1FEF6"author="Zinovjev, A. G."box="[1039,1233,237,259]"pageId="5"pageNumber="62"pagination="429 - 436"refId="ref11711"refString="Zinovjev, A. G. 1979. Erection of a new subgenus Paranematus, subgen. n. (Hymenoptera, Tenthredinidae) for sawflies of the Nematus wahlbergi Thomson group and a review of species from the European regions of the USSR. Entomological review 57: 429 - 436 (English translation of Zinovjev, 1978)."type="journal article"year="1979">Zinovjev, 1979: 430</bibRefCitation>
</treatmentCitation>
;
<treatmentCitationid="3E11AC650F0FFFBAFD15FEFA9E90FED4"author="Abe, M. & I. Togashi"box="[733,1008,266,288]"page="552"pageId="5"pageNumber="62"year="1989">
<bibRefCitationid="DB21F7850F0FFFBAFD15FEFA9E90FED4"author="Abe, M. & I. Togashi"box="[733,1008,266,288]"pageId="5"pageNumber="62"pagination="541 - 560"refId="ref9735"refString="Abe, M. and I. Togashi 1989. Symphyta. In Hirashima, Y. (ed.): A Check List of Japanese Insects, pp. 541 - 560. Entomological Laboratory Faculty of Agriculture, Kyushu University, Fukuoka (in Japanese)."type="book chapter"year="1989">Abe and Togashi, 1989: 552</bibRefCitation>
</treatmentCitation>
;
<treatmentCitationid="3E11AC650F0FFFBAFC35FEFA99B0FED4"author="Naito, T. & H. Yoshida & H. Nakamine & T. Morita & T. Ikeda & H. Suzuki & A. Nakanishi"box="[1021,1232,266,288]"page="27"pageId="5"pageNumber="62"year="2004">
<bibRefCitationid="DB21F7850F0FFFBAFC35FEFA99B0FED4"author="Naito, T. & H. Yoshida & H. Nakamine & T. Morita & T. Ikeda & H. Suzuki & A. Nakanishi"box="[1021,1232,266,288]"pageId="5"pageNumber="62"pagination="1 - 2"refId="ref10496"refString="Naito, T., H. Yoshida, H. Nakamine, T. Morita, T. Ikeda, H. Suzuki and A. Nakanishi 2004. Species diversity of sawflies in Hyogo Prefecture, central Japan. Museum of Nature and Human Activities, Hyogo, Monograph of Natural History and Environmental Science (1): 1 - 2 + pls. 1 - 10 + 1 - 85 (in Japanese)."type="journal article"year="2004">
<bibRefCitationid="DB21F7850F0FFFBAFD15FED89EE6FECA"author="Yoshida, H."box="[733,902,296,318]"pageId="5"pageNumber="62"refId="ref11611"refString="Yoshida, H. 2006. Symphyta (Hymenoptera) of Osaka Prefecture, Japan. 24 pls. + 128 pp. West Japan Hymenopterists` Club, Kakogawa (in Japanese)."type="book"year="2006">Yoshida, 2006: 54</bibRefCitation>
<bibRefCitationid="DB21F7850F0FFFBAFC58FED8992AFECA"author="Sundukov, Yu. N."box="[912,1098,296,318]"pageId="5"pageNumber="62"pagination="20 - 117"refId="ref11055"refString="Sundukov, Yu. N. 2017. Suborder Symphyta - Sawflies and wood wasps. In Lelej, A. S., M. Yu. Proshchalykin and V. M. Loktionov (eds.): Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences, Supple- ment 6, pp. 20 - 117."type="journal article"year="2017">Sundukov, 2017: 69</bibRefCitation>
<bibRefCitationid="DB21F7850F0FFFBAFC46FEB59977FEAE"author="Hara, H."box="[910,1047,325,346]"pageId="5"pageNumber="62"pagination="69 - 87"refId="ref9893"refString="Hara, H. 2019. Subfamily Nematinae. In Editorial Committee of Catalogue of the Insects of Japan (ed.): Cata- logue of the Insects of Japan, Volume 9 Hymenoptera (Part 1 Symphyta), pp. 69 - 87. Entomological Society of Japan, Kyoto (in Japanese)."type="book chapter"year="2019">Hara, 2019: 83</bibRefCitation>
<bibRefCitationid="DB21F7850F0FFFBAFBE9FEB599D7FEAF"author="Hara, H."box="[1057,1207,325,347]"pageId="5"pageNumber="62"pagination="323 - 350"refId="ref9951"refString="Hara, H. 2020. Subfamily Nematinae. In Naito, T., A. Shi- nohara, H. Hara and F. Ito, Sawflies and Woodwasps of Japan, pp. 75 - 89, 323 - 350. Hokkaido University Press, Sapporo (in Japanese)."type="book chapter"year="2020">Hara, 2020: 345</bibRefCitation>
<bibRefCitationid="DB21F7850F0FFFBAFBEFFE9299ADFE83"author="Vikberg, V."box="[1063,1229,354,376]"pageId="5"pageNumber="62"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972: 30</bibRefCitation>
<treatmentCitationid="3E11AC650F0FFFBAFB84FE709E5CFE46"author="Taeger, A. & S. M. Blank & A. D. Liston"page="420"pageId="5"pageNumber="62"year="2010">
<bibRefCitationid="DB21F7850F0FFFBAFB84FE709E5CFE46"author="Taeger, A. & S. M. Blank & A. D. Liston"pageId="5"pageNumber="62"pagination="1 - 1064"refId="ref11283"refString="Taeger, A., S. M. Blank and A. D. Liston 2010. World cat- alog of Symphyta (Hymenoptera). Zootaxa 2580: 1 - 1064."type="journal article"year="2010">
<treatmentCitationid="3E11AC650F0FFFBAFC8DFE6D9939FE47"author="Sundukov, Yu. N. & A. S. Lelej"box="[837,1113,413,435]"page="85"pageId="5"pageNumber="62"year="2012">
<bibRefCitationid="DB21F7850F0FFFBAFC8DFE6D9939FE47"author="Sundukov, Yu. N. & A. S. Lelej"box="[837,1113,413,435]"pageId="5"pageNumber="62"pagination="62 - 119"refId="ref11135"refString="Sundukov, Yu. N. and A. S. Lelej 2012. Podotryad Symphyta - Sidyachebryukhie. In Lelej, A. S. (ed.): Annotated Catalogue of the Insects of Russian Far East, Volume I, Hymenoptera, pp. 62 - 119. Dalnauka, Vladivostok (in Russian)."type="book chapter"year="2012">Sundukov and Lelej, 2012: 85</bibRefCitation>
<figureCitationid="278B96F10F0FFFBAFBE9FDFF9907FDD3"box="[1057,1127,527,551]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="5"pageNumber="62">Fig. 6</figureCitation>
<figureCitationid="278B96F10F0FFFBAFC0AFDC29974FDBE"box="[962,1044,562,586]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="5"pageNumber="62">Fig. 7G</figureCitation>
<figureCitationid="278B96F10F0FFFBAFC11FD87994EFD7B"box="[985,1070,631,655]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="5"pageNumber="62">Fig. 7H</figureCitation>
<figureCitationid="278B96F10F0FFFBAFBBBFD4D99A7FD21"box="[1139,1223,701,725]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="5"pageNumber="62">Fig. 6C</figureCitation>
), rarely yellow. Tegula often yellow or brown laterally. Legs white or pale yellow on apices of coxae, part or most of fore and middle trochanters and trochantelli, hind trochanter and trochantellus, most or wide base of fore tibia, basal third to two thirds of middle tibia, basal third of hind tibia and bases of fore and middle tarsi (
<figureCitationid="278B96F10F0FFFBAFBEDFC5F99A8FC33"box="[1061,1224,943,967]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="5"pageNumber="62">Fig. 6A, B, D</figureCitation>
<figureCitationid="278B96F10F0FFFBAFC53FBE79E8FFBDB"box="[923,1007,1047,1071]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="5"pageNumber="62">Fig. 6A</figureCitation>
<figureCitationid="278B96F10F0FFFBAFD03FBAD9E44FB81"box="[715,804,1117,1141]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="5"pageNumber="62">Fig. 7A</figureCitation>
0.8–1.1: 1.0: 0.8–1.0. Frontal area with lateral ridge low or indistinct and anterior ridge well developed and laterally fused with well-developed transverse ridge above dorsal tentorial pit (
<figureCitationid="278B96F10F0FFFBAFC26FADD9927FAB1"box="[1006,1095,1325,1349]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="5"pageNumber="62">Fig. 7C</figureCitation>
<figureCitationid="278B96F10F0CFFB9FE7CFC059F49FBF8"box="[436,553,1013,1037]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7C, E</figureCitation>
<figureCitationid="278B96F10F0CFFB9FF5CFB529C72FB4E"box="[148,274,1186,1210]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7G–H</figureCitation>
); depth of emargination 0.3–1.2× median height of clypeus. Malar space 0.8– 1.0×as long as median ocellus width. Antenna tapering, with length 2.7–2.8×head width (
<figureCitationid="278B96F10F0CFFB9FDBAFAFA9DD0FAB1"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="6"pageNumber="63">Fig. 6D</figureCitation>
<figureCitationid="278B96F10F0CFFB9FF18FA829C45FA7E"box="[208,293,1394,1418]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7E</figureCitation>
<figureCitationid="278B96F10F0CFFB9FE1AFA479F4EFA3B"box="[466,558,1463,1487]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7H</figureCitation>
<figureCitationid="278B96F10F0CFFB9FE13FA0D9F47F9E1"box="[475,551,1533,1557]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7I</figureCitation>
<figureCitationid="278B96F10F0CFFB9FE1AF9B29F42F9AE"box="[466,546,1602,1626]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7J</figureCitation>
<figureCitationid="278B96F10F0CFFB9FEF9F95A9CF4F936"box="[305,404,1706,1730]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7K</figureCitation>
). Mesepisternum with groove along anterior edge dorsally extending to anterodorsal corner; epicnemium with maximum breadth shorter than mesothoracic spiracle height; epicnemial groove distinct and sharp. Katepimeron glabrous except for one to several setae on posterior edge. Hind tibia dorsally rounded, in lateral view 0.8×as broad as hind femur; posterior tibial spur length 1.5–1.8×apical breadth of tibia in lateral view, 0.6×first tarsomere length. Hind tarsus 0.7–0.8×as long as hind tibia (
<figureCitationid="278B96F10F0CFFB9FCF0FB179EEFFB0B"box="[824,911,1255,1279]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="6"pageNumber="63">Fig. 6D</figureCitation>
<figureCitationid="278B96F10F0CFFB9FC06FAFA9945FAD6"box="[974,1061,1290,1314]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7L</figureCitation>
); depth of concavity between teeth longer than distance between teeth. Fore wing with cell Sc 0.4– 0.8×as wide as vein C at middle of base of vein Rs+M. Hind wing with section of vein 1A between cell 1A and crossvein cu-a 1.5–1.8×as long as crossvein cu-a (
<figureCitationid="278B96F10F0CFFB9FC76FA2A9974FA06"box="[958,1044,1498,1522]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7D</figureCitation>
. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara.
Hypopygium with posterior edge moderately concave beside median projection. Cercus posteriorly extending slightly before near or slightly beyond ovipositor sheath (
<figureCitationid="278B96F10F0CFFB9FC23F995992BF989"box="[1003,1099,1637,1661]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7M</figureCitation>
<figureCitationid="278B96F10F0CFFB9FCB7F93D9EBFF911"box="[895,991,1741,1765]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7M</figureCitation>
<figureCitationid="278B96F10F0CFFB9FC10F91F9952F8F3"box="[984,1074,1775,1799]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="6"pageNumber="63">Fig. 7N</figureCitation>
<figureCitationid="278B96F10F0DFFB8FF29FA0D9C5BF9E1"box="[225,315,1533,1557]"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="7"pageNumber="64">Fig. 8A</figureCitation>
<figureCitationid="278B96F10F0DFFB8FEE5F9EF9CC4F9C3"box="[301,420,1567,1591]"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="7"pageNumber="64">Fig. 8B–C</figureCitation>
) with 20–21 serrulae; sclerite before first annulus wide; basal two annuli without ctenidia; annular sutures sinuate, dorsally curved posteriorly, ventrally slightly curved anteriorly.
. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in anterior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara.
</paragraph>
</caption>
<paragraphid="BF0F8A740F0DFFB8FF6FF93D9EAEF9C3"blockId="7.[140,669,1498,1799]"lastBlockId="7.[707,1236,1498,1799]"pageId="7"pageNumber="64">Body shiny and smooth, with punctures minute or indistinct. Mesopostnotum medially smooth, laterally slightly microsculptured. Metapostnotum smooth. Basal abdominal terga slightly microsculptured.</paragraph>
<figureCitationid="278B96F10F0DFFB8FBF9F99599E9F989"box="[1073,1161,1637,1661]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="7"pageNumber="64">Fig. 6E</figureCitation>
). Legs often pale on apices of fore and middle femora; fore and middle tibiae often entirely pale; fore and middle tarsi often pale except for narrow apices.
<figureCitationid="278B96F10F02FFB7FED1FA659C12FA59"box="[281,370,1429,1453]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="8"pageNumber="65">Fig. 7B</figureCitation>
<figureCitationid="278B96F10F02FFB7FEB4FA2A9CB2FA06"box="[380,466,1498,1522]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="8"pageNumber="65">Fig. 6D</figureCitation>
<figureCitationid="278B96F10F02FFB7FE44F9B29CBEF9AE"box="[396,478,1602,1626]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="8"pageNumber="65">Fig. 7F</figureCitation>
<figureCitationid="278B96F10F02FFB7FE55F93D9C96F911"box="[413,502,1741,1765]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="8"pageNumber="65">Fig. 7O</figureCitation>
<figureCitationid="278B96F10F02FFB7FE7CF91F9F48F8F3"box="[436,552,1775,1799]"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="8"pageNumber="65">Fig. 8D–E</figureCitation>
<figureCitationid="278B96F10F02FFB7FB8EFA6599F6FA59"box="[1094,1174,1429,1453]"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="8"pageNumber="65">Fig. 8F</figureCitation>
<figureCitationid="278B96F10F02FFB7FC0EFA0D997DF9E1"box="[966,1053,1533,1557]"captionStart="Fig"captionStartId="9.[163,196,1002,1024]"captionTargetBox="[159,1216,215,967]"captionTargetId="figure-321@9.[157,1218,213,969]"captionTargetPageId="9"captionText="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826614"httpUri="https://zenodo.org/record/13826614/files/figure.png"pageId="8"pageNumber="65">Fig. 9C</figureCitation>
<figureCitationid="278B96F10F02FFB7FC00F9B29940F9AE"box="[968,1056,1602,1626]"captionStart="Fig"captionStartId="9.[163,196,1002,1024]"captionTargetBox="[159,1216,215,967]"captionTargetId="figure-321@9.[157,1218,213,969]"captionTargetPageId="9"captionText="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826614"httpUri="https://zenodo.org/record/13826614/files/figure.png"pageId="8"pageNumber="65">Fig. 9D</figureCitation>
<figureCitationid="278B96F10F02FFB7FC11F9959953F989"box="[985,1075,1637,1661]"captionStart="Fig"captionStartId="9.[163,196,1002,1024]"captionTargetBox="[159,1216,215,967]"captionTargetId="figure-321@9.[157,1218,213,969]"captionTargetPageId="9"captionText="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826614"httpUri="https://zenodo.org/record/13826614/files/figure.png"pageId="8"pageNumber="65">Fig. 9E</figureCitation>
): Head black; trunk dorsally purplish black, laterally and ventrally pale grayish green; thoracic legs with lateral parts of coxae and claws black; surpedal and subspiracular lobes greenish gray. Final instar (
<figureCitationid="278B96F10F03FFB6FF5CFB8F9C64FB63"box="[148,260,1151,1175]"captionStart="Fig"captionStartId="9.[163,196,1002,1024]"captionTargetBox="[159,1216,215,967]"captionTargetId="figure-321@9.[157,1218,213,969]"captionTargetPageId="9"captionText="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826614"httpUri="https://zenodo.org/record/13826614/files/figure.png"pageId="9"pageNumber="66">Fig. 9F–I</figureCitation>
; color as in middle instar, but dark markings on lateral region reduced; prolegs on abdominal segments 2–7 and 10; caudal protuberances close to each other; clypeus with two pairs of setae; labrum ventrally deeply incised, medially convex with median furrow, with two pairs of setae (
<figureCitationid="278B96F10F03FFB6FE29FABF9F56FA93"box="[481,566,1359,1383]"captionStart="Fig"captionStartId="9.[163,196,1002,1024]"captionTargetBox="[159,1216,215,967]"captionTargetId="figure-321@9.[157,1218,213,969]"captionTargetPageId="9"captionText="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826614"httpUri="https://zenodo.org/record/13826614/files/figure.png"pageId="9"pageNumber="66">Fig. 9H</figureCitation>
<figureCitationid="278B96F10F03FFB6FEFEFA659CE8FA59"box="[310,392,1429,1453]"captionStart="Fig"captionStartId="9.[163,196,1002,1024]"captionTargetBox="[159,1216,215,967]"captionTargetId="figure-321@9.[157,1218,213,969]"captionTargetPageId="9"captionText="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826614"httpUri="https://zenodo.org/record/13826614/files/figure.png"pageId="9"pageNumber="66">Fig. 9I</figureCitation>
. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20 mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon. A–G, Photographed by Ibuki; H–K, photographed by Hara.
<figureCitationid="278B96F10F03FFB6FEF9F9EF9CFFF9C3"box="[305,415,1567,1591]"captionStart="Fig"captionStartId="9.[163,196,1002,1024]"captionTargetBox="[159,1216,215,967]"captionTargetId="figure-321@9.[157,1218,213,969]"captionTargetPageId="9"captionText="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826614"httpUri="https://zenodo.org/record/13826614/files/figure.png"pageId="9"pageNumber="66">Fig. 9J–K</figureCitation>
<taxonomicNameid="78B0F1F70F03FFB6FF11F95A9F4BF936"authority="Knw. O. Conde"box="[217,555,1706,1730]"class="Insecta"family="Tenthredinidae"genus="Nematus"kingdom="Animalia"order="Hymenoptera"pageId="9"pageNumber="66"phylum="Arthropoda"rank="species"species="yokohamensis"subOrder="Symphyta">yokohamensis Knw. O. Conde</taxonomicName>
<collectionCodeid="D9A112B10F03FFB6FF14F9009D8FF8F3"box="[220,239,1776,1799]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="9"pageNumber="66"type="Museum">V</collectionCode>
<figureCitationid="278B96F10F03FFB6FCD3FB529EF9FB4E"box="[795,921,1186,1210]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="9"pageNumber="66">Fig. 6A–C</figureCitation>
<figureCitationid="278B96F10F03FFB6FC66FB529E99FB4E"box="[942,1017,1186,1210]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="9"pageNumber="66">7G, L</figureCitation>
), deposited in the Senckenberg Deutsches Entomologisches Institut, Müncheberg.
<bibRefCitationid="DB21F7850F03FFB6FC42FB18994CFB0B"author="Konow, F. W."box="[906,1068,1255,1279]"pageId="9"pageNumber="66"pagination="50 - 57"refId="ref10226"refString="Konow, F. W. 1895. Neue oder wenig bekannte Tenthre- diniden und eine analytische Ubersicht der Gattung Holcocneme Knw. Termeszetrajzi Fuzetek 18: 50 - 57."type="journal article"year="1895">Konow (1895)</bibRefCitation>
. It is not clear whether the specimen he had was only one or more than one.
<bibRefCitationid="DB21F7850F03FFB6FD0BFABF9E06FA93"author="Vikberg, V."box="[707,870,1359,1383]"pageId="9"pageNumber="66"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg (1972)</bibRefCitation>
referred to the above specimen as lthe holotypez. We regard this as fixation of the
<bibRefCitationid="DB21F7850F03FFB6FCFFFA6599EEFA59"author="Oehlke, J. & J. Wudowenz"box="[823,1166,1429,1453]"pageId="9"pageNumber="66"pagination="363 - 420"refId="ref10575"refString="Oehlke, J. and J. Wudowenz 1984. Katalog der in den Sammlungen der Abteilung Taxonomie der Insekten des Institutes fur Pflanzenschutzforschung, Bereich Eberswalde (ehemals Deutsches Entomologisches Institut), aufbewahrten Typen - XXII (Hymenoptera: Symphyta). Beitrage zur Entomologie 34 (2): 363 - 420."type="journal article"year="1984">Oehlke and Wudowenz (1984)</bibRefCitation>
<figureCitationid="278B96F10F03FFB6FBB8F9EF99A7F9C3"box="[1136,1223,1567,1591]"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="9"pageNumber="66">Fig. 8B</figureCitation>
<collectionCodeid="D9A112B10F03FFB6FB79F97899B0F96B"box="[1201,1232,1672,1695]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="9"pageNumber="66">VI</collectionCode>
. 2020
</date>
, mat. 12, 14.
<collectionCodeid="D9A112B10F03FFB6FC5EF95A9ED7F935"box="[918,951,1706,1729]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="9"pageNumber="66">VI</collectionCode>
., em. 24,
<dateid="CB0EACB40F03FFB6FBEDF95A99D1F936"box="[1061,1201,1706,1730]"pageId="9"pageNumber="66"value="2021-04-29">29. IV. 2021</date>
<figureCitationid="278B96F10F03FFB6FCC5F93D9E0CF911"box="[781,876,1741,1765]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="9"pageNumber="66">Figs. 6E</figureCitation>
<figureCitationid="278B96F10F03FFB6FCB2F93D9ED5F910"box="[890,949,1741,1764]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="9"pageNumber="66">7H–J</figureCitation>
<figureCitationid="278B96F10F03FFB6FC0BF93D99EDF911"box="[963,1165,1741,1765]"captionStart="Fig"captionStartId="9.[163,196,1002,1024]"captionTargetBox="[159,1216,215,967]"captionTargetId="figure-321@9.[157,1218,213,969]"captionTargetPageId="9"captionText="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826614"httpUri="https://zenodo.org/record/13826614/files/figure.png"pageId="9"pageNumber="66">9A–B, D–E, G–K</figureCitation>
<collectionCodeid="D9A112B10F00FFB5FE6AFF289CA1FF1B"box="[418,449,216,239]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="10"pageNumber="67">VI</collectionCode>
. 2020
</date>
, mat. 10.
<collectionCodeid="D9A112B10F00FFB5FDBCFF289FF5FF1B"box="[628,661,216,239]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="10"pageNumber="67">VI</collectionCode>
., em.
<dateid="CB0EACB40F00FFB5FF72FF0A9C20FEE6"box="[186,320,250,274]"pageId="10"pageNumber="67"value="2021-04-19">19. IV. 2021</date>
<figureCitationid="278B96F10F00FFB5FE66FF0A9F6DFEE6"box="[430,525,250,274]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="10"pageNumber="67">Figs. 6D</figureCitation>
<figureCitationid="278B96F10F00FFB5FDD0FF0A9DA9FEC1"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="10"pageNumber="67">7A, C, E, K, M–N</figureCitation>
<collectionCodeid="D9A112B10F00FFB5FE23FEED9F6CFEC0"box="[491,524,285,308]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="10"pageNumber="67">VI</collectionCode>
<figureCitationid="278B96F10F00FFB5FF1CFECF9C4AFEA3"box="[212,298,319,343]"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="10"pageNumber="67">Fig. 8C</figureCitation>
<dateid="CB0EACB40F00FFB5FF44FE929C21FE8E"box="[140,321,354,378]"pageId="10"pageNumber="67"value="2021-04-22"valueMax="2021-04-23"valueMin="2021-04-22">22–23. IV. 2021</date>
, larvae hatch. 30. IV., mat. 15.
<collectionCodeid="D9A112B10F00FFB5FF44FE759DC1FE68"box="[140,161,389,412]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
., em. 22,
<dateid="CB0EACB40F00FFB5FECEFE759CE9FE69"box="[262,393,389,413]"pageId="10"pageNumber="67"value="2022-04-24">24. IV. 2022</date>
<figureCitationid="278B96F10F00FFB5FE50FE759F30FE69"box="[408,592,389,413]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="10"pageNumber="67">Figs. 7B, D, F, O</figureCitation>
<figureCitationid="278B96F10F00FFB5FD94FE759FF9FE69"box="[604,665,389,413]"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="10"pageNumber="67">8D–F</figureCitation>
<figureCitationid="278B96F10F00FFB5FF44FE579DAFFE4B"box="[140,207,423,447]"captionStart="Fig"captionStartId="9.[163,196,1002,1024]"captionTargetBox="[159,1216,215,967]"captionTargetId="figure-321@9.[157,1218,213,969]"captionTargetPageId="9"captionText="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826614"httpUri="https://zenodo.org/record/13826614/files/figure.png"pageId="10"pageNumber="67">9C, F</figureCitation>
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<collectionCodeid="D9A112B10F00FFB5FEECFDA59C57FD98"box="[292,311,597,620]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
<collectionCodeid="D9A112B10F00FFB5FEEEFD889C59FD7B"box="[294,313,632,655]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
<collectionCodeid="D9A112B10F00FFB5FF44FD6A9DFFFD45"box="[140,159,666,689]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
<collectionCodeid="D9A112B10F00FFB5FDD3FD6A9F4EFD45"box="[539,558,666,689]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
. 1954
</date>
, K. Sato; 1 $1 Ə, ditto but
<dateid="CB0EACB40F00FFB5FE50FD4D9F43FD20"box="[408,547,700,725]"pageId="10"pageNumber="67"value="1955-04-22">22. IV. 1955</date>
<collectionCodeid="D9A112B10F00FFB5FE10FD109C8BFD03"box="[472,491,736,759]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
<figureCitationid="278B96F10F00FFB5FF5CFCF29D8DFCEE"box="[148,237,770,794]"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="10"pageNumber="67">Fig. 8A</figureCitation>
<collectionCodeid="D9A112B10F00FFB5FF65FCD59DACFCC8"box="[173,204,805,828]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="10"pageNumber="67">VI</collectionCode>
<collectionCodeid="D9A112B10F00FFB5FECCFCB89C43FCAB"box="[260,291,840,863]"country="Norway"name="Mykotektet, National Veterinary Institute"pageId="10"pageNumber="67">VI</collectionCode>
<collectionCodeid="D9A112B10F00FFB5FD4EFCB89FF9FCAB"box="[646,665,840,863]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
<collectionCodeid="D9A112B10F00FFB5FED8FC7D9C43FC50"box="[272,291,909,932]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
. 1932
</date>
, Takeuchi (cited by
<bibRefCitationid="DB21F7850F00FFB5FD88FC7D9DA6FC33"author="Yoshida, H."pageId="10"pageNumber="67"refId="ref11611"refString="Yoshida, H. 2006. Symphyta (Hymenoptera) of Osaka Prefecture, Japan. 24 pls. + 128 pp. West Japan Hymenopterists` Club, Kakogawa (in Japanese)."type="book"year="2006">Yoshida, 2006</bibRefCitation>
<dateid="CB0EACB40F00FFB5FEACFC229C90FC1E"box="[356,496,978,1002]"pageId="10"pageNumber="67"value="1963-04-10">10. IV. 1963</date>
,
<collectionCodeid="D9A112B10F00FFB5FE35FC229F6FFC1D"box="[509,527,978,1001]"name="Tavera, Department of Geology and Geophysics"pageId="10"pageNumber="67">T</collectionCode>
<collectionCodeid="D9A112B10F00FFB5FF70FBE89DABFBDB"box="[184,203,1048,1071]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
. 1963
</date>
, A. Nakanishi; 1 Ə, Mt. Kora-san,
<dateid="CB0EACB40F00FFB5FF44FBCA9C75FBA6"box="[140,277,1082,1106]"pageId="10"pageNumber="67"value="1962-04-17">17. IV. 1962</date>
<collectionCodeid="D9A112B10F00FFB5FEB9FBAD9CE4FB80"box="[369,388,1117,1140]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
. 1962
</date>
, A. Nakanishi. — Locality unknown: 1 $, l[Chikusho-tani]z(in Japanese),
<dateid="CB0EACB40F00FFB5FECAFB529CF6FB4E"box="[258,406,1186,1210]"pageId="10"pageNumber="67"value="1932-07-25">25. VII. 1932</date>
<collectionCodeid="D9A112B10F00FFB5FEC1FB359C7CFB28"box="[265,284,1221,1244]"country="Canada"lsid="urn:lsid:biocol.org:col:13946"name="Royal British Columbia Museum - Herbarium"pageId="10"pageNumber="67"type="Museum">V</collectionCode>
<bibRefCitationid="DB21F7850F00FFB5FDE3FADD9C5FFA93"author="Hara, H. & A. Shinohara"pageId="10"pageNumber="67"pagination="93 - 113"refId="ref10098"refString="Hara, H. and A. Shinohara 2018. Taxonomic notes and new distribution and host plant records for sawflies and woodwasps (Hymenoptera, Symphyta) of Japan IV. Bulletin of the National Museum of Nature and Sci- ence, Series A 44: 93 - 113."type="journal article"year="2018">Hara and Shinohara, 2018</bibRefCitation>
), Honshu (
<bibRefCitationid="DB21F7850F00FFB5FE7DFABF9F2BFA93"author="Konow, F. W."box="[437,587,1359,1383]"pageId="10"pageNumber="67"pagination="50 - 57"refId="ref10226"refString="Konow, F. W. 1895. Neue oder wenig bekannte Tenthre- diniden und eine analytische Ubersicht der Gattung Holcocneme Knw. Termeszetrajzi Fuzetek 18: 50 - 57."type="journal article"year="1895">Konow, 1895</bibRefCitation>
), Sadogashima Is. (
<bibRefCitationid="DB21F7850F00FFB5FEE4FA829C82FA7E"author="Takeuchi, K."box="[300,482,1394,1418]"pageId="10"pageNumber="67"pagination="150 - 164"refId="ref11317"refString="Takeuchi, K. 1936. Some sawflies from Sado Island. Ten- thredo 1: 150 - 164."type="journal article"year="1936">Takeuchi, 1936</bibRefCitation>
<bibRefCitationid="DB21F7850F00FFB5FEF1FA659F3EFA59"author="Sundukov, Yu. N. & A. S. Lelej"box="[313,606,1429,1453]"pageId="10"pageNumber="67"pagination="62 - 119"refId="ref11135"refString="Sundukov, Yu. N. and A. S. Lelej 2012. Podotryad Symphyta - Sidyachebryukhie. In Lelej, A. S. (ed.): Annotated Catalogue of the Insects of Russian Far East, Volume I, Hymenoptera, pp. 62 - 119. Dalnauka, Vladivostok (in Russian)."type="book chapter"year="2012">Sundukov and Lelej, 2012</bibRefCitation>
<bibRefCitationid="DB21F7850F00FFB5FE32FA2A9C69F9E1"author="Sundukov, Yu. N. & A. S. Lelej"pageId="10"pageNumber="67"pagination="62 - 119"refId="ref11135"refString="Sundukov, Yu. N. and A. S. Lelej 2012. Podotryad Symphyta - Sidyachebryukhie. In Lelej, A. S. (ed.): Annotated Catalogue of the Insects of Russian Far East, Volume I, Hymenoptera, pp. 62 - 119. Dalnauka, Vladivostok (in Russian)."type="book chapter"year="2012">Sundukov and Lelej, 2012</bibRefCitation>
;
<bibRefCitationid="DB21F7850F00FFB5FEDBFA0D9CA5F9E1"author="Sundukov, Yu. N."box="[275,453,1533,1557]"pageId="10"pageNumber="67"pagination="20 - 117"refId="ref11055"refString="Sundukov, Yu. N. 2017. Suborder Symphyta - Sawflies and wood wasps. In Lelej, A. S., M. Yu. Proshchalykin and V. M. Loktionov (eds.): Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences, Supple- ment 6, pp. 20 - 117."type="journal article"year="2017">Sundukov, 2017</bibRefCitation>
<bibRefCitationid="DB21F7850F00FFB5FEDAF9EF9F56F9C3"author="Sundukov, Yu. N. & A. S. Lelej"box="[274,566,1567,1591]"pageId="10"pageNumber="67"pagination="62 - 119"refId="ref11135"refString="Sundukov, Yu. N. and A. S. Lelej 2012. Podotryad Symphyta - Sidyachebryukhie. In Lelej, A. S. (ed.): Annotated Catalogue of the Insects of Russian Far East, Volume I, Hymenoptera, pp. 62 - 119. Dalnauka, Vladivostok (in Russian)."type="book chapter"year="2012">Sundukov and Lelej (2012)</bibRefCitation>
in the distribution of this species, the reason or original source is unknown.
<bibRefCitationid="DB21F7850F00FFB5FEC8F9789CDBF96B"author="Lee, J. - W. & J. - K. Choi & B. Park"box="[256,443,1671,1695]"pageId="10"pageNumber="67"pagination="1 - 96"refId="ref10279"refString="Lee, J. - W., J. - K. Choi and B. Park 2019. Synoptic list of Symphyta (Hymenoptera) in Korea. Journal of Species Research 8 (1): 1 - 96."type="journal article"year="2019">
<bibRefCitationid="DB21F7850F00FFB5FBCEFF2799B3FF1B"author="Wei, M. & H. Nie & A. Taeger"box="[1030,1235,215,239]"pageId="10"pageNumber="67"pagination="505 - 574"refId="ref11540"refString="Wei, M., H. Nie and A. Taeger 2006. Sawflies (Hymenop- tera: Symphyta) of China - Checklist and review of research. In Blank, S. M., S. Schmidt and A. Taeger (eds.): Recent Sawfly Research: Synthesis and Pros- pects, pp. 505 - 574. Goecke & Evers, Keltern."type="book chapter"year="2006">
<bibRefCitationid="DB21F7850F00FFB5FBE4FF0A99CEFEE6"author="Kang, L."box="[1068,1198,250,274]"pageId="10"pageNumber="67"refId="ref10193"refString="Kang, L. (ed.) 1992. The Insect Ecosystem of Inner Mon- golia grassland, pp. 239 - 246. Science Press, Beijing. Not seen."type="book"year="1992">Kang, 1992</bibRefCitation>
) as being based on misidentification.
<bibRefCitationid="DB21F7850F00FFB5FBFFFEED9E29FEA3"author="Sundukov, Yu. N. & A. S. Lelej"pageId="10"pageNumber="67"pagination="62 - 119"refId="ref11135"refString="Sundukov, Yu. N. and A. S. Lelej 2012. Podotryad Symphyta - Sidyachebryukhie. In Lelej, A. S. (ed.): Annotated Catalogue of the Insects of Russian Far East, Volume I, Hymenoptera, pp. 62 - 119. Dalnauka, Vladivostok (in Russian)."type="book chapter"year="2012">Sundukov and Lelej (2012)</bibRefCitation>
<bibRefCitationid="DB21F7850F00FFB5FC25FE929E6DFE69"author="Hara, H. & A. Shinohara"pageId="10"pageNumber="67"pagination="93 - 113"refId="ref10098"refString="Hara, H. and A. Shinohara 2018. Taxonomic notes and new distribution and host plant records for sawflies and woodwasps (Hymenoptera, Symphyta) of Japan IV. Bulletin of the National Museum of Nature and Sci- ence, Series A 44: 93 - 113."type="journal article"year="2018">Hara and Shinohara (2018)</bibRefCitation>
. However,
<bibRefCitationid="DB21F7850F00FFB5FC54FE759901FE69"author="Sundukov, Yu. N."box="[924,1121,389,413]"pageId="10"pageNumber="67"pagination="20 - 117"refId="ref11055"refString="Sundukov, Yu. N. 2017. Suborder Symphyta - Sawflies and wood wasps. In Lelej, A. S., M. Yu. Proshchalykin and V. M. Loktionov (eds.): Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences, Supple- ment 6, pp. 20 - 117."type="journal article"year="2017">Sundukov (2017)</bibRefCitation>
<bibRefCitationid="DB21F7850F00FFB5FBB0FE1D9F9BFDD3"author="Vikberg, V."pageId="10"pageNumber="67"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
<bibRefCitationid="DB21F7850F00FFB5FD0BFDC29E3EFDBE"author="Vikberg, V."box="[707,862,562,586]"pageId="10"pageNumber="67"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
</taxonomicName>
, described from Siberia, both originally regarded as the subspecies of
<bibRefCitationid="DB21F7850F00FFB5FBA9FD2F9F9DFCEE"author="Sundukov, Yu. N."pageId="10"pageNumber="67"pagination="20 - 117"refId="ref11055"refString="Sundukov, Yu. N. 2017. Suborder Symphyta - Sawflies and wood wasps. In Lelej, A. S., M. Yu. Proshchalykin and V. M. Loktionov (eds.): Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences, Supple- ment 6, pp. 20 - 117."type="journal article"year="2017">Sundukov, 2017</bibRefCitation>
. In the lowlands of Honshu, adults were collected from late April to early June and larvae in early June. The larvae collected in early June matured in middle June and became adults in the following spring under rearing conditions. This sawfly has one generation per year. Under rearing conditions, a female inserted her eggs singly into leaves beside or at the main or lateral veins (
<figureCitationid="278B96F10F00FFB5FC1BFBCA992AFBA6"box="[979,1098,1082,1106]"captionStart="Fig"captionStartId="9.[163,196,1002,1024]"captionTargetBox="[159,1216,215,967]"captionTargetId="figure-321@9.[157,1218,213,969]"captionTargetPageId="9"captionText="Fig. 9.ɹNematus yokohamensis. A–B, Egg marks (arrowed) and first instar larvae, 2. VI. 2020; C, ditto, 1. V. 2021; D, early instar larva, 4. VI. 2020; E, middle instar larva, 6. VI. 2020; F, final instar larva, 13. V. 2021; G, mature larva, 10. VI. 2020 (about 20mm long); H, exuvia in cocoon, labrum; I, ditto, antenna, above dorsal; J–K, cocoon.A–G, Photographed by Ibuki; H–K, photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826614"httpUri="https://zenodo.org/record/13826614/files/figure.png"pageId="10"pageNumber="67">Fig. 9A–C</figureCitation>
). Larvae solitarily fed on leaves. No extra molt was observed when the larvae reached maturity. Cocoons were made between papers or in the soil in captivity.
<bibRefCitationid="DB21F7850F00FFB5FCE4FB189EB7FB0B"author="Konow, F. W."box="[812,983,1255,1279]"pageId="10"pageNumber="67"pagination="50 - 57"refId="ref10226"refString="Konow, F. W. 1895. Neue oder wenig bekannte Tenthre- diniden und eine analytische Ubersicht der Gattung Holcocneme Knw. Termeszetrajzi Fuzetek 18: 50 - 57."type="journal article"year="1895">Konow (1895)</bibRefCitation>
, containing only the female, is very simple.
<bibRefCitationid="DB21F7850F00FFB5FC07FAFA99E3FAD6"author="Takeuchi, K."box="[975,1155,1290,1314]"pageId="10"pageNumber="67"pagination="150 - 164"refId="ref11317"refString="Takeuchi, K. 1936. Some sawflies from Sado Island. Ten- thredo 1: 150 - 164."type="journal article"year="1936">Takeuchi (1936)</bibRefCitation>
briefly described the male as lUndescribed male agrees very well with the female except the most of four anterior tibiae and their tarsi white. Length of male
<bibRefCitationid="DB21F7850F00FFB5FBF8FA6599B4FA59"author="Vikberg, V."box="[1072,1236,1429,1453]"pageId="10"pageNumber="67"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg (1972)</bibRefCitation>
gave the key characters with an excellent figure of the lancet of the lholotypez, but he examined only the lholotypez. We here redescribe this species based on a good series of specimens listed above, considering the variation and the characters previously not mentioned.
This species is distinguished from other Japanese nematines in having the frons with an anterior ridge well developed and laterally extending into a facial orbit (
<figureCitationid="278B96F10F00FFB5FC5AF91F9967F8F3"box="[914,1031,1775,1799]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="10"pageNumber="67">Fig. 7C–D</figureCitation>
<figureCitationid="278B96F10F01FFB4FDD1FF279FF9FF1B"box="[537,665,215,239]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="11"pageNumber="68">Fig. 7C–F</figureCitation>
<bibRefCitationid="DB21F7850F01FFB4FE92FE3A9C9CFE16"author="Vikberg, V."box="[346,508,458,482]"pageId="11"pageNumber="68"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
<figureCitationid="278B96F10F01FFB4FF05FD6A9C48FD46"box="[205,296,666,690]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="11"pageNumber="68">Fig. 6D</figureCitation>
<bibRefCitationid="DB21F7850F01FFB4FEEBFD4D9CAAFD21"author="Vikberg, V."box="[291,458,701,725]"pageId="11"pageNumber="68"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
) and the second annulus of the lancet without ctenidial teeth (
<figureCitationid="278B96F10F01FFB4FDBAFD2F9DADFCEE"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="11"pageNumber="68">Fig. 8B–C</figureCitation>
<bibRefCitationid="DB21F7850F01FFB4FEACFCF29F67FCEE"author="Vikberg, V."box="[356,519,770,794]"pageId="11"pageNumber="68"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
<bibRefCitationid="DB21F7850F01FFB4FE48FCB79F7EFCAB"author="Vikberg, V."box="[384,542,839,863]"pageId="11"pageNumber="68"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
<bibRefCitationid="DB21F7850F01FFB4FDF0FC9A9DA1FC51"author="Zinovjev, A. G."pageId="11"pageNumber="68"pagination="625 - 635"refId="ref11645"refString="Zinovjev, A. G. 1978. Description of a new subgenus Paranematus subgen. n. for the group of species related to Nematus wahlbergi Thomson (Hymenoptera, Tenth- redinidae) with a review of species from the European part of the USSR. Entomologicheskoe Obozrenie 57 (3): 625 - 635 (in Russian, abstract in English)."type="journal article"year="1978">Zinovjev, 1978</bibRefCitation>
<bibRefCitationid="DB21F7850F01FFB4FDF5FC7D9DADFC33"author="Zinovjev, A. G."pageId="11"pageNumber="68"pagination="625 - 635"refId="ref11645"refString="Zinovjev, A. G. 1978. Description of a new subgenus Paranematus subgen. n. for the group of species related to Nematus wahlbergi Thomson (Hymenoptera, Tenth- redinidae) with a review of species from the European part of the USSR. Entomologicheskoe Obozrenie 57 (3): 625 - 635 (in Russian, abstract in English)."type="journal article"year="1978">Zinovjev (1978</bibRefCitation>
,
<bibRefCitationid="DB21F7850F01FFB4FF12FC5F9C73FC33"author="Zinovjev, A. G."box="[218,275,943,967]"pageId="11"pageNumber="68"pagination="429 - 436"refId="ref11711"refString="Zinovjev, A. G. 1979. Erection of a new subgenus Paranematus, subgen. n. (Hymenoptera, Tenthredinidae) for sawflies of the Nematus wahlbergi Thomson group and a review of species from the European regions of the USSR. Entomological review 57: 429 - 436 (English translation of Zinovjev, 1978)."type="journal article"year="1979">1979</bibRefCitation>
<bibRefCitationid="DB21F7850F01FFB4FD89FC229DA1FBF9"author="Vikberg, V."pageId="11"pageNumber="68"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
<bibRefCitationid="DB21F7850F01FFB4FF44FB529C4BFB4E"author="Vikberg, V."box="[140,299,1186,1210]"pageId="11"pageNumber="68"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
<figureCitationid="278B96F10F01FFB4FDBAFB529DCCFB29"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="11"pageNumber="68">Fig. 7C</figureCitation>
<figureCitationid="278B96F10F01FFB4FF1EFADD9C30FAB1"box="[214,336,1325,1349]"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="11"pageNumber="68">Fig. 8B–C</figureCitation>
<bibRefCitationid="DB21F7850F01FFB4FDC9FA829FFDFA7E"author="Vikberg, V."box="[513,669,1394,1418]"pageId="11"pageNumber="68"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
<bibRefCitationid="DB21F7850F01FFB4FE8EFA659C8BFA59"author="Zinovjev, A. G."box="[326,491,1429,1453]"pageId="11"pageNumber="68"pagination="625 - 635"refId="ref11645"refString="Zinovjev, A. G. 1978. Description of a new subgenus Paranematus subgen. n. for the group of species related to Nematus wahlbergi Thomson (Hymenoptera, Tenth- redinidae) with a review of species from the European part of the USSR. Entomologicheskoe Obozrenie 57 (3): 625 - 635 (in Russian, abstract in English)."type="journal article"year="1978">Zinovjev, 1978</bibRefCitation>
<figureCitationid="278B96F10F01FFB4FDC2FA2A9FF5FA06"box="[522,661,1498,1522]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="11"pageNumber="68">Fig. 6B, D</figureCitation>
<bibRefCitationid="DB21F7850F01FFB4FEA7F9D09F51F9C3"author="Prous, M. & A. Liston & K. Kramp & H. Savina & H. Vardal & A. Taeger"box="[367,561,1567,1591]"pageId="11"pageNumber="68"pagination="63 - 127"refId="ref10845"refString="Prous, M., A. Liston, K. Kramp, H. Savina, H. Vardal and A. Taeger 2019. The West Palaearctic genera of Nema- tinae (Hymenoptera, Tenthredinidae). ZooKeys 875: 63 - 127."type="journal article"year="2019">
<figureCitationid="278B96F10F01FFB4FD8CF9959FF5F989"box="[580,661,1637,1661]"captionStart="Fig"captionStartId="7.[163,196,1263,1285]"captionTargetBox="[159,1216,215,1228]"captionTargetId="figure-121@7.[157,1218,213,1230]"captionTargetPageId="7"captionText="Fig. 7.ɹNematus yokohamensis. A, Head in dorsal view, female, B, ditto, male; C, frons in anterodorsal view, female (convexity arrowed); D, ditto, male; E, head and base of antenna in lateral view, female (convexity arrowed); F, ditto, male; G, ventral part of head in anterior view (lectotype); H, clypeus and mandibles in ante- rior view, female; I–J, right and left mandibles in outer views, female; K, posterior part of thorax in dorsal view, female; L, claw (lectotype); M, apex of female abdomen in dorsal view; N, apex of female abdomen in lateral view; O, apex of male abdomen in dorsal view. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826608"httpUri="https://zenodo.org/record/13826608/files/figure.png"pageId="11"pageNumber="68">Fig. 7E</figureCitation>
<bibRefCitationid="DB21F7850F01FFB4FF44F93D9C51F911"author="Zinovjev, A. G."box="[140,305,1741,1765]"pageId="11"pageNumber="68"pagination="625 - 635"refId="ref11645"refString="Zinovjev, A. G. 1978. Description of a new subgenus Paranematus subgen. n. for the group of species related to Nematus wahlbergi Thomson (Hymenoptera, Tenth- redinidae) with a review of species from the European part of the USSR. Entomologicheskoe Obozrenie 57 (3): 625 - 635 (in Russian, abstract in English)."type="journal article"year="1978">Zinovjev, 1978</bibRefCitation>
,
<bibRefCitationid="DB21F7850F01FFB4FEF5F93D9C14F911"author="Zinovjev, A. G."box="[317,372,1741,1765]"pageId="11"pageNumber="68"pagination="429 - 436"refId="ref11711"refString="Zinovjev, A. G. 1979. Erection of a new subgenus Paranematus, subgen. n. (Hymenoptera, Tenthredinidae) for sawflies of the Nematus wahlbergi Thomson group and a review of species from the European regions of the USSR. Entomological review 57: 429 - 436 (English translation of Zinovjev, 1978)."type="journal article"year="1979">1979</bibRefCitation>
) and the second annulus of the lancet without ctenidial teeth (
<figureCitationid="278B96F10F01FFB4FE32F91F9F0EF8F3"box="[506,622,1775,1799]"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="11"pageNumber="68">Fig. 8B–C</figureCitation>
<bibRefCitationid="DB21F7850F01FFB4FC5EFF27995AFF1B"author="Vikberg, V."box="[918,1082,215,239]"pageId="11"pageNumber="68"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
<bibRefCitationid="DB21F7850F01FFB4FC40FEED9946FEC1"author="Vikberg, V."box="[904,1062,285,309]"pageId="11"pageNumber="68"pagination="25 - 39"refId="ref11452"refString="Vikberg, V. 1972. A contribution to the taxonomy of the Nematus wahlbergi and lonicerae groups (Hym., Ten- thredinidae) feeding on Lonicera. Annales Entomolog- ici Fennici 38: 25 - 39."type="book chapter"year="1972">Vikberg, 1972</bibRefCitation>
<bibRefCitationid="DB21F7850F01FFB4FD0BFECF9E0DFEA3"author="Zinovjev, A. G."box="[707,877,319,343]"pageId="11"pageNumber="68"pagination="625 - 635"refId="ref11645"refString="Zinovjev, A. G. 1978. Description of a new subgenus Paranematus subgen. n. for the group of species related to Nematus wahlbergi Thomson (Hymenoptera, Tenth- redinidae) with a review of species from the European part of the USSR. Entomologicheskoe Obozrenie 57 (3): 625 - 635 (in Russian, abstract in English)."type="journal article"year="1978">Zinovjev, 1978</bibRefCitation>
<figureCitationid="278B96F10F01FFB4FCA4FE3A9EA7FE16"box="[876,967,458,482]"captionStart="Fig"captionStartId="6.[163,196,842,864]"captionTargetBox="[164,1216,215,801]"captionTargetId="figure-391@6.[157,1218,213,808]"captionTargetPageId="6"captionText="Fig. 6.ɹNematus yokohamensis. A–B, Female in dorsolateral and ventrolateral views (lectotype); C, pronotum in lateral view (lectotype); D, female, head in anterior view, whole in dorsal and ventral views; E, male, head in anterior view, whole in dorsal and ventral views. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826606"httpUri="https://zenodo.org/record/13826606/files/figure.png"pageId="11"pageNumber="68">Fig. 6D</figureCitation>
<bibRefCitationid="DB21F7850F01FFB4FB40FE1D9F9BFDD3"author="Enslin, E."pageId="11"pageNumber="68"pagination="311 - 412"refId="ref9823"refString="Enslin, E. 1915. Die Tenthredinoidea Mitteleuropas IV. Deutsche Entomologische Zeitschrift [1915] (Beiheft 4): 311 - 412."type="journal article"year="1915">Enslin, 1915</bibRefCitation>
;
<bibRefCitationid="DB21F7850F01FFB4FCC3FDE09EB8FDD3"author="Prous, M. & A. Liston & K. Kramp & H. Savina & H. Vardal & A. Taeger"box="[779,984,527,551]"pageId="11"pageNumber="68"pagination="63 - 127"refId="ref10845"refString="Prous, M., A. Liston, K. Kramp, H. Savina, H. Vardal and A. Taeger 2019. The West Palaearctic genera of Nema- tinae (Hymenoptera, Tenthredinidae). ZooKeys 875: 63 - 127."type="journal article"year="2019">
<figureCitationid="278B96F10F01FFB4FB62FDC29FBFFD99"captionStart="Fig"captionStartId="8.[163,196,1308,1330]"captionTargetBox="[159,1213,215,1273]"captionTargetId="figure-189@8.[157,1218,213,1275]"captionTargetPageId="8"captionText="Fig. 8.ɹNematus yokohamensis. A, Lance; B–C, lancet; D–E, male genitalia in dorsal and ventral views; F, penis valve. Photographed by Hara."figureDoi="http://doi.org/10.5281/zenodo.13826612"httpUri="https://zenodo.org/record/13826612/files/figure.png"pageId="11"pageNumber="68">Fig. 8F</figureCitation>
<bibRefCitationid="DB21F7850F01FFB4FC4AFDA59922FD99"author="Prous, M. & A. Liston & K. Kramp & H. Savina & H. Vardal & A. Taeger"box="[898,1090,597,621]"pageId="11"pageNumber="68"pagination="63 - 127"refId="ref10845"refString="Prous, M., A. Liston, K. Kramp, H. Savina, H. Vardal and A. Taeger 2019. The West Palaearctic genera of Nema- tinae (Hymenoptera, Tenthredinidae). ZooKeys 875: 63 - 127."type="journal article"year="2019">